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Sample records for beewolves philanthus triangulum

  1. Refining the Roots of the Beewolf-Streptomyces Symbiosis: Antennal Symbionts in the Rare Genus Philanthinus (Hymenoptera, Crabronidae)

    Science.gov (United States)

    Yildirim, Erol; Gürbüz, M. Faruk; Herzner, Gudrun; Strohm, Erhard

    2012-01-01

    Insects engage in symbiotic associations with a large diversity of beneficial microorganisms. While the majority of well-studied symbioses have a nutritional basis, several cases are known in which bacteria protect their host from pathogen infestation. Solitary wasps of the genera Philanthus and Trachypus (beewolves; Hymenoptera, Crabronidae) cultivate the actinomycete “Candidatus Streptomyces philanthi” in specialized antennal gland reservoirs. The symbionts are transferred to the larval cocoon, where they provide protection against pathogenic fungi by producing at least nine different antibiotics. Here we investigated the closest relatives of Philanthus and Trachypus, the rare genus Philanthinus, for the presence of antennal gland reservoirs and symbiotic streptomycetes. Molecular analyses identified “Ca. Streptomyces philanthi” in reservoirs of Philanthinus quattuordecimpunctatus. Phylogenies based on the 16S rRNA gene suggest that P. quattuordecimpunctatus may have acquired “Ca. Streptomyces philanthi” by horizontal transfer from other beewolf species. In histological sections and three-dimensional reconstructions, the antennal gland reservoirs were found to occupy six antennal segments (as opposed to only five in Philanthus and Trachypus) and to be structurally less complex than those of the evolutionarily more derived genera of beewolves. The presence of “Ca. Streptomyces philanthi” in antennal glands of Philanthinus indicates that the symbiosis between beewolves and Streptomyces bacteria is much older than previously thought. It probably evolved along the branch leading to the monophyletic tribe Philanthini, as it seems to be confined to the genera Philanthus, Trachypus, and Philanthinus, which together comprise 172 described species of solitary wasps. PMID:22113914

  2. Morphology, chemistry and function of the postpharyngeal gland in the South American digger wasps Trachypus boharti and Trachypus elongatus.

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    Gudrun Herzner

    Full Text Available Microbes pose severe threats to animals as competitors or pathogens and strongly affect the evolution of life history traits like parental care. Females of the European beewolf Philanthus triangulum, a solitary digger wasp, provision their offspring with paralyzed honeybees and embalm them with the secretion from large postpharyngeal glands (PPG that contain mainly unsaturated hydrocarbons. This coating changes the physico-chemical properties of the prey surface, causes a reduction of water condensation and retards growth of mold fungi. Here we examined the closely related South American genus Trachypus, which shows a life-history similar to Philanthus. We investigated whether Trachypus spp. also possess PPGs and embalm larval provisions. Using histological methods and 3D reconstructions we show that Trachypus boharti and T. elongatus possess PPGs that are similar to P. triangulum but somewhat smaller. The ultrastructure of the gland epithelium suggests that the gland content is at least partly sequestered from the hemolymph. Chemical analyses using gas chromatography / mass spectrometry revealed that both the cuticle and PPGs of Trachypus contain mainly unsaturated long-chain hydrocarbons. The gland of T. boharti additionally contains long-chain ketones. The hydrocarbons from the PPG of T. elongatus occurred on prey bees excavated from nests in the field but not on conspecific control bees. While the embalming only slightly elevated the amount of hydrocarbons on prey bees, the proportion of unsaturated hydrocarbons, which is crucial for the antifungal effect, was significantly increased. The Trachypus species under study possess PPGs that are very similar to the PPG of P. triangulum with regard to morphology, ultrastructure and chemistry. Moreover, we provide clear evidence that T. elongatus females embalm their prey, presumably as a means of prey preservation. The observed differences among Trachypus and Philanthus in gland size and prey

  3. A cuckoo in wolves' clothing? Chemical mimicry in a specialized cuckoo wasp of the European beewolf (Hymenoptera, Chrysididae and Crabronidae

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    Herzner Gudrun

    2008-01-01

    Full Text Available Abstract Background Host-parasite interactions are among the most important biotic relationships. Host species should evolve mechanisms to detect their enemies and employ appropriate counterstrategies. Parasites, in turn, should evolve mechanisms to evade detection and thus maximize their success. Females of the European beewolf (Philanthus triangulum, Hymenoptera, Crabronidae hunt exclusively honeybee workers as food for their progeny. The brood cells containing the paralyzed bees are severely threatened by a highly specialized cuckoo wasp (Hedychrum rutilans, Hymenoptera, Chrysididae. Female cuckoo wasps enter beewolf nests to oviposit on paralyzed bees that are temporarily couched in the nest burrow. The cuckoo wasp larva kills the beewolf larva and feeds on it and the bees. Here, we investigated whether H. rutilans evades detection by its host. Since chemical senses are most important in the dark nest, we hypothesized that the cuckoo wasp might employ chemical camouflage. Results Field observations suggest that cuckoo wasps are attacked by beewolves in front of their nest, most probably after being recognized visually. In contrast, beewolves seem not to detect signs of the presence of these parasitoids neither when these had visited the nest nor when directly encountered in the dark nest burrow. In a recognition bioassay in observation cages, beewolf females responded significantly less frequently to filter paper discs treated with a cuticular extract from H. rutilans females, than to filter paper discs treated with an extract from another cuckoo wasp species (Chrysis viridula. The behavior to paper discs treated with a cuticular extract from H. rutilans females did not differ significantly from the behavior towards filter paper discs treated with the solvent only. We hypothesized that cuckoo wasps either mimic the chemistry of their beewolf host or their host's prey. We tested this hypothesis using GC-MS analyses of the cuticles of male and

  4. Cartography of Triangulum-Andromeda using SDSS stars

    Science.gov (United States)

    Perottoni, H. D.; Rocha-Pinto, H. J.; Girardi, L.; Balbinot, E.; Santiago, B. X.; Majewski, S. R.; Anders, F.; Da Costa, L.; Maia, M. A. G.

    2018-01-01

    The outer Galactic halo is home to a number of substructures which still have an uncertain origin, but most likely are remnants of former interactions between the Galaxy and its former satellites. Triangulum-Andromeda (TriAnd) is one of these halo substructures, found as an overdensity of Two Micron All Sky Survey (2MASS) M giants. We analysed the region of TriAnd using photometric data from the Ninth Data Release of Sloan Digital Sky Survey. By comparing the observations with simulations from the TRILEGAL Galactic model, we were able to identify and map several scattered overdensities of main-sequence stars that seem to be associated with TriAnd over a large area covering ∼500 deg2. One of these excesses may represent a new stellar overdensity. We also briefly discuss an alternative hypothesis, according to which TriAnd is one of the troughs of oscillation rings in the Galactic disc.

  5. Triangulum II: Possibly a Very Dense Ultra-faint Dwarf Galaxy

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    Kirby, Evan N.; Cohen, Judith G.; Simon, Joshua D.; Guhathakurta, Puragra

    2015-11-01

    Laevens et al. recently discovered Triangulum II (Tri II), a satellite of the Milky Way. Its Galactocentric distance is 36 kpc, and its luminosity is only 450 {L}⊙ . Using Keck/DEIMOS, we measured the radial velocities of six member stars within 1.‧2 of the center of Tri II, and we found a velocity dispersion of {σ }v={5.1}-1.4+4.0 {km} {{{s}}}-1. We also measured the metallicities of three stars and found a range of 0.8 dex in [Fe/H]. The velocity and metallicity dispersions identify Tri II as a dark matter-dominated galaxy. The galaxy is moving very quickly toward the Galactic center ({v}{{GSR}}=-262 {km} {{{s}}}-1). Although it might be in the process of being tidally disrupted as it approaches pericenter, there is no strong evidence for disruption in our data set. The ellipticity is low, and the mean velocity, =-382.1+/- 2.9 {km} {{{s}}}-1, rules out an association with the Triangulum-Andromeda substructure or the Pan-Andromeda Archaeological Survey stellar stream. If Tri II is in dynamical equilibrium, then it would have a mass-to-light ratio of {3600}-2100+3500 {M}⊙ {L}⊙ -1, the highest of any non-disrupting galaxy (those for which dynamical mass estimates are reliable). The density within the 3D half-light radius would be {4.8}-3.5+8.1 {M}⊙ {{{pc}}}-3, even higher than Segue 1. Hence, Tri II is an excellent candidate for the indirect detection of dark matter annihilation. The data presented herein were obtained at the W. M. Keck Observatory, which is operated as a scientific partnership among the California Institute of Technology, the University of California and the National Aeronautics and Space Administration. The Observatory was made possible by the generous financial support of the W. M. Keck Foundation.

  6. PAndAS' CUBS: DISCOVERY OF TWO NEW DWARF GALAXIES IN THE SURROUNDINGS OF THE ANDROMEDA AND TRIANGULUM GALAXIES

    International Nuclear Information System (INIS)

    Martin, Nicolas F.; McConnachie, Alan W.; Irwin, Mike; Chapman, Scott; Widrow, Lawrence M.; Ferguson, Annette M. N.; Ibata, Rodrigo A.; Dubinski, John; Babul, Arif; Navarro, Julio; Fardal, Mark; Lewis, Geraint F.; Rich, R. Michael

    2009-01-01

    We present the discovery of two new dwarf galaxies, Andromeda XXI and Andromeda XXII, located in the surroundings of the Andromeda and Triangulum galaxies (M31 and M33). These discoveries stem from the first year data of the Pan-Andromeda Archaeological Survey, a photometric survey of the M31/M33 group conducted with the Megaprime/MegaCam Wide-Field Camera mounted on the Canada-France-Hawaii Telescope. Both satellites appear as spatial overdensities of stars which, when plotted in a color-magnitude diagram, follow metal-poor, [Fe/H] = -1.8, red giant branches at the distance of M31/M33. Andromeda XXI is a moderately bright dwarf galaxy (M V = -9.9 ± 0.6), albeit with low surface brightness, emphasizing again that many relatively luminous M31 satellites still remain to be discovered. It is also a large satellite, with a half-light radius close to 1 kpc, making it the fourth largest Local Group dwarf spheroidal galaxy after the recently discovered Andromeda XIX, Andromeda II, and Sagittarius around the Milky Way, and supports the trend that M31 satellites are larger than their Milky Way counterparts. Andromeda XXII is much fainter (M V = -6.5 ± 0.8) and lies a lot closer in projection to M33 than it does to M31 (42 versus 224 kpc), suggesting that it could be the first Triangulum satellite to be discovered. Although this is a very exciting possibility in the context of a past interaction of M33 with M31 and the fate of its satellite system, a confirmation will have to await a good distance estimate to confirm its physical proximity to M33. Along with the dwarf galaxies found in previous surveys of the M31 surroundings, these two new satellites bring the number of dwarf spheroidal galaxies in this region to 20.

  7. Astrometry with Hubble Space Telescope Fine Guidance Sensor 3: The Parallax of the Cataclysmic Variable RW Triangulum

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    McArthur, B. E.; Benedict, G. F.; Lee, J.; Lu, C.-L.; van Altena, W. F.; Deliyannis, C. P.; Girard, T.; Fredrick, L. W.; Nelan, E.; Duncombe, R. L.; Hemenway, P. D.; Jefferys, W. H.; Shelus, P. J.; Franz, O. G.; Wasserman, L. H.

    1999-07-01

    RW Triangulum (RW Tri) is a 13th magnitude nova-like cataclysmic variable star with an orbital period of 0.2319 days (5.56 hr). Infrared observations of RW Tri indicate that its secondary is most likely a late-K dwarf (Dhillon). Past analyses predicted a distance of 270 pc, derived from a blackbody fit to the spectrum of the central part of the disk (Rutten, van Paradijs, & Tinbergen). Recently completed Hubble Space Telescope Fine Guidance Sensor interferometric observations allow us to determine the first trigonometric parallax to RW Tri. This determination puts the distance of RW Tri at 341-31+38, one of the most distant objects with a direct parallax measurement. We compare our result with methods previously employed to estimate distances to cataclysmic variables.

  8. Chemistry of the Triangulum-Andromeda Overdensity as Seen by APOGEE

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    Rochford Hayes, Christian; Majewski, Steven R.; Hasselquist, Sten; Beaton, Rachael; Cunha, Katia M. L.; Smith, Verne V.; Price-Whelan, Adrian M.; APOGEE Team

    2018-06-01

    The nature of the Triangulum-Andromeda (TriAnd) system has been debated since the discovery of this distant, low-latitude Milky Way (MW) overdensity more than a decade ago. Explanations for its origin are either as a halo substructure from the disruption of a dwarf galaxy or a distant extension of the Galactic disk. We test these hypotheses using chemical abundances of a dozen TriAnd members from the Sloan Digital Sky Survey’s 14th Data Release of Apache Point Observatory Galactic Evolution Experiment (APOGEE) data to compare to APOGEE abundances of stars with similar metallicity from both the Sagittarius (Sgr) dSph, and the outer MW disk. We find that TriAnd stars are chemically distinct from Sgr across a variety of elements, (C+N), Mg, K, Ca, Mn, and Ni, with a separation in [X/Fe] of about 0.1 to 0.4 dex depending on the element. Instead, the TriAnd stars, with a median metallicity of about -0.8, exhibit chemical abundance ratios similar to those of the lowest metallicity ([Fe/H] ~ -0.7) stars in the outer Galactic disk, and are consistent with expectations of extrapolated chemical gradients in the outer disk of the MW. These results suggest that TriAnd is associated with the MW disk, and, therefore, that the disk extends to this overdensity - i.e., past a Galactocentric radius of 24 kpc - albeit vertically perturbed about 7 kpc below the nominal disk midplane in this region of the Galaxy.

  9. DETECTION OF OPHIDIOMYCES OPHIODIICOLA IN TWO CAPTIVE BOCOURT WATER SNAKES ( SUBSESSOR BOCOURTI) AND ONE CAPTIVE PUEBLAN MILK SNAKE ( LAMPROPELTIS TRIANGULUM CAMPBELLI).

    Science.gov (United States)

    Picquet, Pierre; Heckers, Kim O; Kolesnik, Ekaterina; Heusinger, Anton; Marschang, Rachel E

    2018-03-01

    Two captive Bocourt water snakes ( Subsessor bocourti) presented with chronic white skin lesions on their heads; Ophidiomyces ophiodiicola was identified by culture and polymerase chain reaction (PCR) in skin scrapings from both snakes. Histopathology performed in one Bocourt water snake revealed fungal hyphae in epidermal structures of lesions. One Pueblan milk snake ( Lampropeltis triangulum campbelli) from the same zoologic institution presented with yellow crusts and white blisters on its body, from which O. ophiodiicola was identified by culture and PCR. Two of the three snakes apparently recovered from lesions after multiple natural sheds, whereas the third snake died. This is the first report of O. ophiodiicola infection in Bocourt water snakes and in a Pueblan milk snake, as well as the first report of O. ophiodiicola in France.

  10. Accelerated evolution of mitochondrial but not nuclear genomes of Hymenoptera: new evidence from crabronid wasps.

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    Martin Kaltenpoth

    Full Text Available Mitochondrial genes in animals are especially useful as molecular markers for the reconstruction of phylogenies among closely related taxa, due to the generally high substitution rates. Several insect orders, notably Hymenoptera and Phthiraptera, show exceptionally high rates of mitochondrial molecular evolution, which has been attributed to the parasitic lifestyle of current or ancestral members of these taxa. Parasitism has been hypothesized to entail frequent population bottlenecks that increase rates of molecular evolution by reducing the efficiency of purifying selection. This effect should result in elevated substitution rates of both nuclear and mitochondrial genes, but to date no extensive comparative study has tested this hypothesis in insects. Here we report the mitochondrial genome of a crabronid wasp, the European beewolf (Philanthus triangulum, Hymenoptera, Crabronidae, and we use it to compare evolutionary rates among the four largest holometabolous insect orders (Coleoptera, Diptera, Hymenoptera, Lepidoptera based on phylogenies reconstructed with whole mitochondrial genomes as well as four single-copy nuclear genes (18S rRNA, arginine kinase, wingless, phosphoenolpyruvate carboxykinase. The mt-genome of P. triangulum is 16,029 bp in size with a mean A+T content of 83.6%, and it encodes the 37 genes typically found in arthropod mt genomes (13 protein-coding, 22 tRNA, and two rRNA genes. Five translocations of tRNA genes were discovered relative to the putative ancestral genome arrangement in insects, and the unusual start codon TTG was predicted for cox2. Phylogenetic analyses revealed significantly longer branches leading to the apocritan Hymenoptera as well as the Orussoidea, to a lesser extent the Cephoidea, and, possibly, the Tenthredinoidea than any of the other holometabolous insect orders for all mitochondrial but none of the four nuclear genes tested. Thus, our results suggest that the ancestral parasitic lifestyle of

  11. Exploring halo substructure with giant stars. XIV. The nature of the Triangulum-Andromeda stellar features

    Energy Technology Data Exchange (ETDEWEB)

    Sheffield, Allyson A.; Johnston, Kathryn V. [Department of Astronomy, Columbia University, Mail Code 5246, New York, NY 10027 (United States); Majewski, Steven R.; Damke, Guillermo; Richardson, Whitney; Beaton, Rachael [Department of Astronomy, University of Virginia, P.O. Box 400325, Charlottesville, VA 22904 (United States); Rocha-Pinto, Helio J., E-mail: asheffield@astro.columbia.edu, E-mail: kvj@astro.columbia.edu, E-mail: srm4n@virginia.edu, E-mail: gjd3r@virginia.edu, E-mail: wwr2u@virginia.edu, E-mail: rlb9n@virginia.edu, E-mail: helio@astro.ufrj.br [Observatório do Valongo, Universidade Federal do Rio de Janeiro, Rio de Janeiro (Brazil)

    2014-09-20

    As large-scale stellar surveys have become available over the past decade, the ability to detect and characterize substructures in the Galaxy has increased dramatically. These surveys have revealed the Triangulum-Andromeda (TriAnd) region to be rich with substructures in the distance range 20-30 kpc, and the relation of these features to each other, if any, remains unclear. An exploration using Two Micron All Sky Survey (2MASS) photometry reveals not only the faint sequence in M giants detected by Rocha-Pinto et al. spanning the range 100° < l < 160° and –50° < b < –15°, but, in addition, a second, brighter and more densely populated sequence. These sequences are likely associated with the distinct main sequences (MSs) discovered (and labeled TriAnd1 and TriAnd2) by Martin et al. in an optical survey in the direction of M31, where TriAnd2 is the optical counterpart of the fainter red giant branch (RGB)/asymptotic giant branch sequence of Rocha-Pinto et al. Here, the age, distance, and metallicity ranges for TriAnd1 and TriAnd2 are estimated by simultaneously fitting isochrones to the 2MASS RGB tracks and the optical MS/MS turn-off features. The two populations are clearly distinct in age and distance: the brighter sequence (TriAnd1) is younger (6-10 Gyr) and closer (distance of ∼15-21 kpc), whereas the fainter sequence (TriAnd2) is older (10-12 Gyr) and at an estimated distance of ∼24-32 kpc. A comparison with simulations demonstrates that the differences and similarities between TriAnd1 and TriAnd2 can simultaneously be explained if they represent debris originating from the disruption of the same dwarf galaxy, but torn off during two distinct pericentric passages.

  12. Systematics of the Dendropsophus leucophyllatus species complex (Anura: Hylidae): Cryptic diversity and the description of two new species

    Science.gov (United States)

    Caminer, Marcel A.; Milá, Borja; Jansen, Martin; Fouquet, Antoine; Venegas, Pablo J.; Chávez, Germán; Lougheed, Stephen C.

    2017-01-01

    Genetic data in studies of systematics of Amazonian amphibians frequently reveal that purportedly widespread single species in reality comprise species complexes. This means that real species richness may be significantly higher than current estimates. Here we combine genetic, morphological, and bioacoustic data to assess the phylogenetic relationships and species boundaries of two Amazonian species of the Dendropsophus leucophyllatus species group: D. leucophyllatus and D. triangulum. Our results uncovered the existence of five confirmed and four unconfirmed candidate species. Among the confirmed candidate species, three have available names: Dendropsophus leucophyllatus, Dendropsophus triangulum, and Dendropsophus reticulatus, this last being removed from the synonymy of D. triangulum. A neotype of D. leucophyllatus is designated. We describe the remaining two confirmed candidate species, one from Bolivia and another from Peru. All confirmed candidate species are morphologically distinct and have much smaller geographic ranges than those previously reported for D. leucophyllatus and D. triangulum sensu lato. Dendropsophus leucophyllatus sensu stricto occurs in the Guianan region. Dendropsophus reticulatus comb. nov. corresponds to populations in the Amazon basin of Brazil, Ecuador, and Peru previously referred to as D. triangulum. Dendropsophus triangulum sensu stricto is the most widely distributed species; it occurs in Amazonian Ecuador, Peru and Brazil, reaching the state of Pará. We provide accounts for all described species including an assessment of their conservation status. PMID:28248998

  13. The LAMOST survey of background quasars in the vicinity of the Andromeda and Triangulum galaxies. II. Results from the commissioning observations and the pilot surveys

    International Nuclear Information System (INIS)

    Huo, Zhi-Ying; Bai, Zhong-Rui; Chen, Jian-Jun; Chen, Xiao-Yan; Du, Bing; Jia, Lei; Lei, Ya-Juan; Liu, Xiao-Wei; Yuan, Hai-Bo; Xiang, Mao-Sheng; Huang, Yang; Zhang, Hui-Hua; Yan, Lin; Chu, Jia-Ru; Chu, Yao-Quan; Hu, Hong-Zhuan; Cui, Xiang-Qun; Hou, Yong-Hui; Hu, Zhong-Wen; Jiang, Fang-Hua

    2013-01-01

    We present new quasars discovered in the vicinity of the Andromeda and Triangulum galaxies with the Large Sky Area Multi-Object Fiber Spectroscopic Telescope, also named the Guoshoujing Telescope, during the 2010 and 2011 observational seasons. Quasar candidates are selected based on the available Sloan Digital Sky Survey, Kitt Peak National Observatory 4 m telescope, Xuyi Schmidt Telescope Photometric Survey optical, and Wide-field Infrared Survey Explorer near-infrared photometric data. We present 509 new quasars discovered in a stripe of ∼135 deg 2 from M31 to M33 along the Giant Stellar Stream in the 2011 pilot survey data sets, and also 17 new quasars discovered in an area of ∼100 deg 2 that covers the central region and the southeastern halo of M31 in the 2010 commissioning data sets. These 526 new quasars have i magnitudes ranging from 15.5 to 20.0, redshifts from 0.1 to 3.2. They represent a significant increase of the number of identified quasars in the vicinity of M31 and M33. There are now 26, 62, and 139 known quasars in this region of the sky with i magnitudes brighter than 17.0, 17.5, and 18.0, respectively, of which 5, 20, and 75 are newly discovered. These bright quasars provide an invaluable collection with which to probe the kinematics and chemistry of the interstellar/intergalactic medium in the Local Group of galaxies. A total of 93 quasars are now known with locations within 2.°5 of M31, of which 73 are newly discovered. Tens of quasars are now known to be located behind the Giant Stellar Stream, and hundreds are behind the extended halo and its associated substructures of M31. The much enlarged sample of known quasars in the vicinity of M31 and M33 can potentially be utilized to construct a perfect astrometric reference frame to measure the minute proper motions (PMs) of M31 and M33, along with the PMs of substructures associated with the Local Group of galaxies. Those PMs are some of the most fundamental properties of the Local Group.

  14. Neurotransmitter Receptors and Their Ionic Channels as Targets for Drugs and Toxins

    Science.gov (United States)

    1985-01-06

    over a period of three months, the glands and venom sacs were removed, frozen in liquid nitrogen then lyophilized. This honey bee wolf (Philanthus...uses a paralytic venom to immobilize honey bee workers and steal their honey . Water extractU oF lvophilized venom sacs and glands injected into honey bee ...specifically with any of these regulatory proteins and to under- stand mechanisms by which these receptors regulate the selective transloca- tion of

  15. Noteworthy reptiles records from Guanajuato, Mexico Registros notables de reptiles para Guanajuato, México

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    José Ismael Campos-Rodríguez

    2010-04-01

    Full Text Available Five new reptile records for the Mexican state of Guanajuato are reported: Lampropeltis triangulum dixoni, Leptodeira annulata cusiliris, Oxybelis aeneus, Tropidodipsas sartorii, and Scincella silvicola caudaequinae. These records are within western versant of the Sierra Gorda at Sierra Gorda de Guanajuato. Evidence is given that supports the idea that the area supports a herpetofaunal assemblage of tropical affinities.Se presentan 5 nuevos registros de reptiles para el estado de Guanajuato; Lampropeltis triangulum dixoni, Leptodeira annulata cusiliris, Oxybelis aeneus, Tropidodipsas sartorii y Scincella silvicola caudaequinae. Todos localizados en la vertiente oeste de la sierra Madre Oriental, en la sierra Gorda de Guanajuato. Se proporciona evidencia de un ensamble herpetofaunístico de afinidad tropical a subtropical presente en la región.

  16. Edzard Schaper - mit Massen : Vortrag zum neunzigsten Geburtstag des Dichters in Brig / Claus Sommerhage

    Index Scriptorium Estoniae

    Sommerhage, Claus, 1950-2003

    2005-01-01

    Ettekanne kirjanik Edzard Schaperi 90. sünniaastapäevaks. Ümbertrükk raamatust: Triangulum : Germanistisches Jahrbuch für Estland, Lettland und Litauen. Tartu, 1998. Lehekülgedel numbrid: 192-206

  17. [Konverentsi "Goethe Tartus" ettekandekogumikest] / Kairit Kaur

    Index Scriptorium Estoniae

    Kaur, Kairit

    2001-01-01

    Arvustus: Triangulum. Germanistisches Jahrbuch '99 für Estland, Lettland und Litauen. Sonderheft: Goethe. Sechste Folge / hrsg. Claus Sommerhage. Tartu, 1999 ; Eesti Goethe Seltsi aastaraamat II: Goethe Tartus : konverentsi "Goethe Tartus (1999) ettekanded. Tartu : Eesti Goethe Selts, 2000

  18. From the Cluster Temperature Function to the Mass Function at Low Z

    Science.gov (United States)

    Mushotzky, Richard (Technical Monitor); Markevitch, Maxim

    2004-01-01

    This XMM project consisted of three observations of the nearby, hot galaxy cluster Triangulum Australis, one of the cluster center and two offsets. The goal was to measure the radial gas temperature profile out to large radii and derive the total gravitating mass within the radius of average mass overdensity 500. The central pointing also provides data for a detailed two-dimensional gas temperature map of this interesting cluster. We have analyzed all three observations. The derivation of the temperature map using the central pointing is complete, and the paper is soon to be submitted. During the course of this study and of the analysis of archival XMM cluster observations, it became apparent that the commonly used XMM background flare screening techniques are often not accurate enough for studies of the cluster outer regions. The information on the cluster's total masses is contained at large off-center distances, and it is precisely the temperatures for those low-brightness regions that are most affected by the detector background anomalies. In particular, our two offset observations of the Triangulum have been contaminated by the background flares ("bad cosmic weather") to a degree where they could not be used for accurate spectral analysis. This forced us to expand the scope of our project. We needed to devise a more accurate method of screening and modeling the background flares, and to evaluate the uncertainty of the XMM background modeling. To do this, we have analyzed a large number of archival EPIC blank-field and closed-cover observations. As a result, we have derived stricter background screening criteria. It also turned out that mild flares affecting EPIC-pn can be modeled with an adequate accuracy. Such modeling has been used to derive our Triangulum temperature map. The results of our XMM background analysis, including the modeling recipes, are presented in a paper which is in final preparation and will be submitted soon. It will be useful not only

  19. Noteworthy reptiles records from Guanajuato, Mexico Registros notables de reptiles para Guanajuato, México

    OpenAIRE

    José Ismael Campos-Rodríguez; Berenice Pérez-Valera; Leonor E. Evaristo-Aguilar; Cynthia Elizalde-Arellano; Juan Carlos López-Vidal; Raúl Hernández-Arciga

    2010-01-01

    Five new reptile records for the Mexican state of Guanajuato are reported: Lampropeltis triangulum dixoni, Leptodeira annulata cusiliris, Oxybelis aeneus, Tropidodipsas sartorii, and Scincella silvicola caudaequinae. These records are within western versant of the Sierra Gorda at Sierra Gorda de Guanajuato. Evidence is given that supports the idea that the area supports a herpetofaunal assemblage of tropical affinities.Se presentan 5 nuevos registros de reptiles para el estado de Guanajuato; ...

  20. Molecular Identification of Cryptosporidium Species from Pet Snakes in Thailand

    OpenAIRE

    Yimming, Benjarat; Pattanatanang, Khampee; Sanyathitiseree, Pornchai; Inpankaew, Tawin; Kamyingkird, Ketsarin; Pinyopanuwat, Nongnuch; Chimnoi, Wissanuwat; Phasuk, Jumnongjit

    2016-01-01

    Cryptosporidium is an important pathogen causing gastrointestinal disease in snakes and is distributed worldwide. The main objectives of this study were to detect and identify Cryptosporidium species in captive snakes from exotic pet shops and snake farms in Thailand. In total, 165 fecal samples were examined from 8 snake species, boa constrictor (Boa constrictor constrictor), corn snake (Elaphe guttata), ball python (Python regius), milk snake (Lampropeltis triangulum), king snake (Lampropel...

  1. Exploring M33 Through RR Lyrae Stars

    OpenAIRE

    Pritzl, Barton J.

    2013-01-01

    Recent surveys have detected RR Lyrae stars in M33, the Triangulum Galaxy. These variable stars are excellent tracers of ancient stellar populations. The RR Lyrae stars have been used to estimate metallicities at various locations within M33, as well as determining the distance to the galaxy. A summary of the M33 RR Lyrae stars is presented here as well as an analysis on what their properties imply for the unique M33 galaxy

  2. STUDI EFFEKTIVITAS KEFIR BENING DAN MENIRAN (Philanthus niruri TERHADAP PENURUNAN KADAR GLUKOSA DARAH PADA HEWAN TIKUS WISTAR HIPERGLIKEMIA DIINDUKSI STREPTOZOTOCIN

    Directory of Open Access Journals (Sweden)

    Judiono .

    2014-09-01

    were given feed refers to the AIN 93. Fasting glucose levels before and after treatment were measured by Super (Glucocard II enzimatic method. The results showed that body weight increased in all treatments, except meniran, however the increaments were not differ significantly. Changes in body weight of insulin group were 13.800 + 16.104 g, kefir treatment amounted to 13.812 + 21.294, and meniran increased about 18.394 + 16.225. Changes in blood glucose in the insulin group about -118.571+55.815 mg/dL, kefir treatment at -102.875+60.454, while meniran group -66,625+37,784. In contrast, the positive control group tended to increase glucose levels. Conclusion In vitro, plain kefir and meniran potentially lowered blood glucose levels. However, kefir showed lowered glucose level better than meniran. Futher research needs to study bio- molecular mechanisms of the decline in the future. Key words: diabetes mellitus, meniran, plain kefir, streptozotocin (STZ, hyperglycemic rat

  3. The star-formation law at GMC scales in M33, the Triangulum Galaxy

    Science.gov (United States)

    Williams, Thomas G.; Gear, Walter K.; Smith, Matthew W. L.

    2018-06-01

    We present a high spatial resolution study, on scales of ˜100pc, of the relationship between star-formation rate (SFR) and gas content within Local Group galaxy M33. Combining deep SCUBA-2 observations with archival GALEX, SDSS, WISE, Spitzer and submillimetre Herschel data, we are able to model the entire SED from UV to sub-mm wavelengths. We calculate the SFR on a pixel-by-pixel basis using the total infrared luminosity, and find a total SFR of 0.17 ± 0.06 {M}_⊙/yr, somewhat lower than our other two measures of SFR - combined FUV and 24μ SFR (0.25^{+0.10}_{-0.07} {M}_⊙/yr) and SED-fitting tool MAGPHYS (0.33^{+0.05}_{-0.06} {M}_⊙/yr). We trace the total gas using a combination of the 21cm HI line for atomic hydrogen, and CO(J=2-1) data for molecular hydrogen. We have also traced the total gas using dust masses. We study the star-formation law in terms of molecular gas, total gas, and gas from dust. We perform an analysis of the star-formation law on a variety of pixel scales, from 25" to 500" (100pc to 2kpc). At kpc scales, we find that a linear Schmidt-type power law index is suitable for molecular gas, but the index appears to be much higher with total gas, and gas from dust. Whilst we find a strong scale dependence on the Schmidt index, the gas depletion timescale is invariant with pixel scale.

  4. Partner choice and fidelity stabilize coevolution in a Cretaceous-age defensive symbiosis

    Science.gov (United States)

    Kaltenpoth, Martin; Roeser-Mueller, Kerstin; Koehler, Sabrina; Peterson, Ashley; Nechitaylo, Taras Y.; Stubblefield, J. William; Herzner, Gudrun; Seger, Jon; Strohm, Erhard

    2014-01-01

    Many insects rely on symbiotic microbes for survival, growth, or reproduction. Over evolutionary timescales, the association with intracellular symbionts is stabilized by partner fidelity through strictly vertical symbiont transmission, resulting in congruent host and symbiont phylogenies. However, little is known about how symbioses with extracellular symbionts, representing the majority of insect-associated microorganisms, evolve and remain stable despite opportunities for horizontal exchange and de novo acquisition of symbionts from the environment. Here we demonstrate that host control over symbiont transmission (partner choice) reinforces partner fidelity between solitary wasps and antibiotic-producing bacteria and thereby stabilizes this Cretaceous-age defensive mutualism. Phylogenetic analyses show that three genera of beewolf wasps (Philanthus, Trachypus, and Philanthinus) cultivate a distinct clade of Streptomyces bacteria for protection against pathogenic fungi. The symbionts were acquired from a soil-dwelling ancestor at least 68 million years ago, and vertical transmission via the brood cell and the cocoon surface resulted in host–symbiont codiversification. However, the external mode of transmission also provides opportunities for horizontal transfer, and beewolf species have indeed exchanged symbiont strains, possibly through predation or nest reuse. Experimental infection with nonnative bacteria reveals that—despite successful colonization of the antennal gland reservoirs—transmission to the cocoon is selectively blocked. Thus, partner choice can play an important role even in predominantly vertically transmitted symbioses by stabilizing the cooperative association over evolutionary timescales. PMID:24733936

  5. UNVEILING THE NEW GENERATION OF STARS IN NGC 604 WITH GEMINI-NIRI

    Energy Technology Data Exchange (ETDEWEB)

    Farina, Cecilia; Bosch, Guillermo L. [Facultad de Ciencias Astronomicas y Geofisicas, Universidad Nacional de La Plata, Paseo de Bosque S/N (B1900FWA), La Plata (Argentina); Barba, Rodolfo H., E-mail: ceciliaf@fcaglp.unlp.edu.ar [Instituto de Ciencias Astronomicas, de la Tierra y del Espacio (ICATE-CONICET), Av. Espana Sur 1512 (J5402DSP), San Juan (Argentina)

    2012-02-15

    We present a near-infrared study focused on the detection and characterization of the youngest stellar component of the NGC 604 giant star-forming region in the Triangulum galaxy (M 33). By means of color-color diagrams derived from the photometry of JHK{sub s} images taken with the Gemini Near Infrared Imaging and Spectrometer (NIRI), we have found 68 candidate massive young stellar objects. The spatial distribution of these sources matches the areas where previous studies suggested that star formation might be taking place, and the high spatial resolution of our deep NIRI imaging allows us to pinpoint the star-forming knots. An analysis of the fraction of objects that show infrared excess suggests that the star formation is still active, supporting the presence of a second generation of stars being born, although the evidence for or against sequential star formation does not seem to be conclusive.

  6. Pulsating red giants and supergiants as probes of galaxy formation and evolution

    Science.gov (United States)

    Theodorus van Loon, Jacco; Javadi, Atefeh; Khosroshahi, Habib; Rezaei, Sara; Golshan, Roya; Saberi, Maryam

    2015-08-01

    We have developed new techniques to use pulsating red giant and supergiants stars to reconstruct the star formation history of galaxies over cosmological time, as well as using them to map the dust production across their host galaxies. We describe the large programme on the Local Group spiral galaxy Triangulum (M33), which we have monitored at near-infrared wavelengths for several years using the United Kingdom InfraRed Telescope in Hawai'i. We outline the methodology and present the results for the central square kiloparsec (Javadi et al. 2011a,b, 2013) and - fresh from the press - the disc of M33 (Javadi et al. 2015, and in preparation). We also describe the results from our application of this new technique to other nearby galaxies: the Magellanic Clouds (published in Rezaei et al. 2014), the dwarf galaxies NGC 147 and 185 (Golshan et al. in preparation), and Centaurus A.

  7. UNVEILING THE NEW GENERATION OF STARS IN NGC 604 WITH GEMINI-NIRI

    International Nuclear Information System (INIS)

    Fariña, Cecilia; Bosch, Guillermo L.; Barbá, Rodolfo H.

    2012-01-01

    We present a near-infrared study focused on the detection and characterization of the youngest stellar component of the NGC 604 giant star-forming region in the Triangulum galaxy (M 33). By means of color-color diagrams derived from the photometry of JHK s images taken with the Gemini Near Infrared Imaging and Spectrometer (NIRI), we have found 68 candidate massive young stellar objects. The spatial distribution of these sources matches the areas where previous studies suggested that star formation might be taking place, and the high spatial resolution of our deep NIRI imaging allows us to pinpoint the star-forming knots. An analysis of the fraction of objects that show infrared excess suggests that the star formation is still active, supporting the presence of a second generation of stars being born, although the evidence for or against sequential star formation does not seem to be conclusive.

  8. The Block-Elmegreen conference

    CERN Document Server

    Elmegreen, Bruce; Block, David; Woolway, Matthew

    2015-01-01

     Our understanding of galaxy formation comes mostly from two sources: sensitive observations at high angular resolution of the high-redshift Universe, where galaxies are observed to be forming, and detailed observations of individual stars and clouds in the Local Group, where telltale remnants from its formative time remain and similar processes operate at a low level today. The current conference focusses on key aspects of the Local Group, composed of the Milky Way, Andromeda and Triangulum Spiral Galaxies, the Large and Small Magellanic Cloud galaxies, numerous dwarf and irregular galaxies, and intergalactic gas. Topics include the halo and thick disk of the Milky Way with its first stars and stellar streams; the Milky Way bar, bulge and outer edge; interstellar dust and turbulence; star formation processes and stellar scattering in spiral arms; views through the infrared Eyes of the Spitzer Space Telescope; globular clusters; the Local Gould Belt; stellar metallicities and elemental abundances; the enviro...

  9. Constraint on the velocity dependent dark matter annihilation cross section from gamma-ray and kinematic observations of ultrafaint dwarf galaxies

    Science.gov (United States)

    Zhao, Yi; Bi, Xiao-Jun; Yin, Peng-Fei; Zhang, Xinmin

    2018-03-01

    Searching for γ rays from dwarf spheroidal galaxies (dSphs) is a promising approach to detect dark matter (DM) due to the high DM densities and low baryon components in dSphs. The Fermi-LAT observations from dSphs have set stringent constraints on the velocity independent annihilation cross section. However, the constraints from dSphs may change in velocity dependent annihilation scenarios because of the different velocity dispersions in galaxies. In this work, we study how to set constraints on the velocity dependent annihilation cross section from the combined Fermi-LAT observations of dSphs with the kinematic data. In order to calculate the γ ray flux from the dSph, the correlation between the DM density profile and velocity dispersion at each position should be taken into account. We study such correlation and the relevant uncertainty from kinematic observations by performing a Jeans analysis. Using the observational results of three ultrafaint dSphs with large J-factors, including Willman 1, Reticulum II, and Triangulum II, we set constraints on the p-wave annihilation cross section in the Galaxy as an example.

  10. Monitoring pulsating giant stars in M33: star formation history and chemical enrichment

    Science.gov (United States)

    Javadi, A.; van Loon, J. Th

    2017-06-01

    We have conducted a near-infrared monitoring campaign at the UK InfraRed Telescope (UKIRT), of the Local Group spiral galaxy M33 (Triangulum). A new method has been developed by us to use pulsating giant stars to reconstruct the star formation history of galaxies over cosmological time as well as using them to map the dust production across their host galaxies. In first Instance the central square kiloparsec of M33 was monitored and long period variable stars (LPVs) were identified. We give evidence of two epochs of a star formation rate enhanced by a factor of a few. These stars are also important dust factories, we measure their dust production rates from a combination of our data with Spitzer Space Telescope mid-IR photometry. Then the monitoring survey was expanded to cover a much larger part of M33 including spiral arms. Here we present our methodology and describe results for the central square kiloparsec of M33 [1-4] and disc of M33 [5-8].

  11. Monitoring pulsating giant stars in M33: star formation history and chemical enrichment

    International Nuclear Information System (INIS)

    Javadi, A; Van Loon, J Th

    2017-01-01

    We have conducted a near-infrared monitoring campaign at the UK InfraRed Telescope (UKIRT), of the Local Group spiral galaxy M33 (Triangulum). A new method has been developed by us to use pulsating giant stars to reconstruct the star formation history of galaxies over cosmological time as well as using them to map the dust production across their host galaxies. In first Instance the central square kiloparsec of M33 was monitored and long period variable stars (LPVs) were identified. We give evidence of two epochs of a star formation rate enhanced by a factor of a few. These stars are also important dust factories, we measure their dust production rates from a combination of our data with Spitzer Space Telescope mid-IR photometry. Then the monitoring survey was expanded to cover a much larger part of M33 including spiral arms. Here we present our methodology and describe results for the central square kiloparsec of M33 [1–4] and disc of M33 [5–8]. (paper)

  12. Dactylogyrid monogeneans parasitising Cyphocharax voga (Hensel) (Teleostei: Curimatidae) from the Pampas region, Argentina: new and previously described species.

    Science.gov (United States)

    Rossin, María Alejandra; Timi, Juan Tomás

    2016-09-01

    Most studies on dactylogyrid monogeneans in Argentina have been carried out during 1980s and 1990s. Many of these species have been later synonymised and other remain under a confusing taxonomic status, particularly those parasitising Cyphocharax voga (Hensel) (Teleostei: Curimatidae). In order to clarify the identity of dactylogyrids, new material was collected from fishes in Lake Chascomús, Buenos Aires Province, Argentina. A total of four species was found in the gills of C. voga. Two known species, Curvianchoratus singularis (Suriano, 1980) Suriano, 1986 and Palombitrema triangulum (Suriano, 1981) Suriano, 1997, are redescribed and their generic and specific status discussed, and two new species are described. Urocleidoides surianoae n. sp. can be distinguished from its congeners by having an anterior medial projection in the ventral bar and a laminar ligament connecting the base of the male copulatory organ and accessory piece. Annulotrematoides bonaerensis n. sp. differs from its congeners principally by having a ventral bar with an anterior medial projection. The diversity of dactylogyrids harboured by C. voga indicates the need of further studies in the Pampas region, which will provide interesting and valuable sources of evidence for future zoogeographical and evolutionary research on dactylogyrids in the Neotropics.

  13. Molecular Identification of Cryptosporidium Species from Pet Snakes in Thailand.

    Science.gov (United States)

    Yimming, Benjarat; Pattanatanang, Khampee; Sanyathitiseree, Pornchai; Inpankaew, Tawin; Kamyingkird, Ketsarin; Pinyopanuwat, Nongnuch; Chimnoi, Wissanuwat; Phasuk, Jumnongjit

    2016-08-01

    Cryptosporidium is an important pathogen causing gastrointestinal disease in snakes and is distributed worldwide. The main objectives of this study were to detect and identify Cryptosporidium species in captive snakes from exotic pet shops and snake farms in Thailand. In total, 165 fecal samples were examined from 8 snake species, boa constrictor (Boa constrictor constrictor), corn snake (Elaphe guttata), ball python (Python regius), milk snake (Lampropeltis triangulum), king snake (Lampropeltis getula), rock python (Python sebae), rainbow boa (Epicrates cenchria), and carpet python (Morelia spilota). Cryptosporidium oocysts were examined using the dimethyl sulfoxide (DMSO)-modified acid-fast staining and a molecular method based on nested-PCR, PCR-RFLP analysis, and sequencing amplification of the SSU rRNA gene. DMSO-modified acid-fast staining revealed the presence of Cryptosporidium oocysts in 12 out of 165 (7.3%) samples, whereas PCR produced positive results in 40 (24.2%) samples. Molecular characterization indicated the presence of Cryptosporidium parvum (mouse genotype) as the most common species in 24 samples (60%) from 5 species of snake followed by Cryptosporidium serpentis in 9 samples (22.5%) from 2 species of snake and Cryptosporidium muris in 3 samples (7.5%) from P. regius.

  14. A VLA Search for Radio Signals from M31 and M33

    Science.gov (United States)

    Gray, Robert H.; Mooley, Kunal

    2017-03-01

    Observing nearby galaxies would facilitate the search for artificial radio signals by sampling several billions of stars simultaneously, but few efforts have been made to exploit this opportunity. An added attraction is that the Milky Way is the second largest member of the Local Group, so our galaxy might be a probable target for hypothetical broadcasters in nearby galaxies. We present the first relatively high spectral resolution (intelligent radio signals of complete galaxies in the Local Group with the Jansky VLA, observing the galaxies M31 (Andromeda) and M33 (Triangulum)—the first and third largest members of the group, respectively—sampling more stars than any prior search of this kind. We used 122 Hz channels over a 1 MHz spectral window in the target galaxy velocity frame of reference, and 15 Hz channels over a 125 kHz window in our local standard of rest. No narrowband signals were detected above a signal-to-noise ratio of 7, suggesting the absence of continuous narrowband flux greater than approximately 0.24 and 1.33 Jy in the respective spectral windows illuminating our part of the Milky Way during our observations in 2014 December and 2015 January. This is also the first study in which the upgraded VLA has been used for SETI.

  15. Explaining formation of Astronomical Jets using Dynamic Universe Model

    Science.gov (United States)

    Naga Parameswara Gupta, Satyavarapu

    2016-07-01

    Astronomical jets are observed from the centres of many Galaxies including our own Milkyway. The formation of such jet is explained using SITA simulations of Dynamic Universe Model. For this purpose the path traced by a test neutron is calculated and depicted using a set up of one densemass of the mass equivalent to mass of Galaxy center, 90 stars with similar masses of stars near Galaxy center, mass equivalents of 23 Globular Cluster groups, 16 Milkyway parts, Andromeda and Triangulum Galaxies at appropriate distances. Five different kinds of theoretical simulations gave positive results The path travelled by this test neutron was found to be an astronomical jet emerging from Galaxy center. This is another result from Dynamic Universe Model. It solves new problems like a. Variable Mass Rocket Trajectory Problem b. Explaining Very long baseline interferometry (VLBI) observations c. Astronomical jets observed from Milkyway Center d. Prediction of Blue shifted Galaxies e. Explaining Pioneer Anomaly f. Prediction of New Horizons satellite trajectory etc. Dynamic Universe Model never reduces to General relativity on any condition. It uses a different type of mathematics based on Newtonian physics. This mathematics used here is simple and straightforward. As there are no differential equations present in Dynamic Universe Model, the set of equations give single solution in x y z Cartesian coordinates for every point mass for every time step

  16. The UK Infrared Telescope M33 monitoring project - I. Variable red giant stars in the central square kiloparsec

    Science.gov (United States)

    Javadi, Atefeh; van Loon, Jacco Th.; Mirtorabi, Mohammad Taghi

    2011-02-01

    We have conducted a near-infrared monitoring campaign at the UK Infrared Telescope (UKIRT), of the Local Group spiral galaxy M33 (Triangulum). The main aim was to identify stars in the very final stage of their evolution, and for which the luminosity is more directly related to the birth mass than the more numerous less-evolved giant stars that continue to increase in luminosity. The most extensive data set was obtained in the K band with the UIST instrument for the central 4 × 4 arcmin2 (1 kpc2) - this contains the nuclear star cluster and inner disc. These data, taken during the period 2003-2007, were complemented by J- and H-band images. Photometry was obtained for 18 398 stars in this region; of these, 812 stars were found to be variable, most of which are asymptotic giant branch (AGB) stars. Our data were matched to optical catalogues of variable stars and carbon stars and to mid-infrared photometry from the Spitzer Space Telescope. In this first of a series of papers, we present the methodology of the variability survey and the photometric catalogue - which is made publicly available at the Centre de Données astronomiques de Strasbourg - and discuss the properties of the variable stars. The most dusty AGB stars had not been previously identified in optical variability surveys, and our survey is also more complete for these types of stars than the Spitzer survey.

  17. A Novel Pattern of Yolk Processing in Developing Snake Eggs (Colubridae: Lampropeltini) and its Functional and Evolutionary Implications.

    Science.gov (United States)

    Powers, Kathryn G; Blackburn, Daniel G

    2017-07-01

    Early amniotic vertebrates evolved large-yolked eggs that permitted production of well-developed, terrestrial hatchlings. This reproductive pattern required new mechanisms for cellularizing the yolk and mobilizing it for embryonic use. In birds, cells that line the yolk sac cavity phagocytose and digest the yolk material, a pattern that is commonly assumed to be universal among oviparous amniotes. However, recent evidence challenges the assumption that all squamate reptiles conform to the avian developmental pattern. In this paper, scanning electron microscopy and histology were used to study mechanisms of yolk processing in two colubrid snakes, the kingsnake Lampropeltis getula and the milksnake L. triangulum. Endodermal cells from the yolk sac splanchnopleure proliferate massively as they invade the yolk sac cavity, forming elaborate chains of interlinked cells. These cells grow in size as they phagocytose yolk material. Subsequently, vitelline capillaries invade the masses of yolk-laden cells and become coated with the endodermal cells, forming an elaborate meshwork of cell-coated strands. The close association of cells, yolk, and blood vessels allows yolk material to be cellularized, digested, and transported for embryonic use. The overall pattern is like that of the corn snake Pantherophis guttatus, but contrasts markedly with that of birds. Given recent evidence that this developmental pattern may also occur in certain lizards, we postulate that it is ancestral for squamates. Studies of lizards, crocodilians, and turtles are needed to clarify the evolutionary history of this pattern and its implications for the evolution of the amniotic (terrestrial) vertebrate egg. © 2017 Wiley Periodicals, Inc.

  18. The LAMOST survey of background quasars in the vicinity of M31 and M33 – III. results from the 2013 regular survey

    International Nuclear Information System (INIS)

    Huo, Zhi-Ying; Shi, Jian-Rong; Yang, Ming; Liu, Xiao-Wei; Xiang, Mao-Sheng; Huang, Yang; Yuan, Hai-Bo; Zhang, Yong; Hou, Yong-Hui; Wang, Yue-Fei

    2015-01-01

    In this work, we report new quasars discovered in fields in the vicinity of the Andromeda (M31) and Triangulum (M33) galaxies with the Large Sky Area Multi-Object Fiber Spectroscopic Telescope (LAMOST, also called the Guo Shou Jing Telescope) during the 2013 observational season, the second year of the regular survey. In total, 1330 new quasars are discovered in an area of ∼133 deg 2 around M31 and M33. With i magnitudes ranging from 14.79 to 20.0 and redshifts from 0.08 to 4.85, the 1330 new quasars represent a significant increase in the number of identified quasars in fields in the vicinity of M31 and M33. Up to now, there have been a total of 1870 quasars discovered by LAMOST in this area. The much enlarged sample of known quasars in this area can potentially be utilized to construct a precise astrometric reference frame for the measurement of minute proper motions of M31, M33 and their associated substructures, which are vital for understanding the formation and evolution of M31, M33 and the Local Group of galaxies. Moreover, in the sample, there are a total of 45, 98 and 225 quasars with i magnitudes brighter than 17.0, 17.5 and 18.0 respectively. In the aforementioned brightness bins, 15, 35 and 84 quasars are reported here for the first time, and 6, 21 and 81 are reported in our pervious work. In addition, 0, 1 and 6 are from the Sloan Digital Sky Survey and 24, 41 and 54 are from the NED database. These bright quasars provide an invaluable sample to study the kinematics and chemistry of the interstellar/intergalactic medium of the Local Group. (paper)

  19. A deep narrowband survey for planetary nebulae at the outskirts of M 33

    Science.gov (United States)

    Galera-Rosillo, R.; Corradi, R. L. M.; Mampaso, A.

    2018-04-01

    Context. Planetary nebulae (PNe) are excellent tracers of stellar populations with low surface brightness, and therefore provide a powerful method to detect and explore the rich system of substructures discovered around the main spiral galaxies of the local group. Aim. We searched the outskirts of the local group spiral galaxy M 33 (the Triangulum) for PNe to gain new insights into the extended stellar substructure on the northern side of the disc and to study the existence of a faint classical halo. Methods: The search is based on wide field imaging covering a 4.5 square degree area out to a maximum projected distance of about 40 kpc from the centre of the galaxy. The PN candidates are detected by the combination of images obtained in narrowband filters selecting the [OIII]λ5007 Å and Hα + [NII] nebular lines and in the continuum g' and r' broadband filters. Results: Inside the bright optical disc of M 33, eight new PN candidates were identified, three of which were spectroscopically confirmed. No PN candidates were found outside the limits of the disc. Fourteen additional sources showing [OIII] excess were also discovered. Conclusions: The absence of bright PN candidates in the area outside the galaxy disc covered by this survey sets an upper limit to the luminosity of the underlying population of 1.6 × 107 L⊙, suggesting the lack of a massive classical halo, which is in agreement with the results obtained using the red giant branch population. Based on observations made with the Isaac Newton Telescope and service observations made with the William Herschel Telescope operated on the island of La Palma by the Isaac Newton Group of Telescopes in the Spanish Observatorio del Roque de los Muchachos of the Instituto de Astrofísica de Canarias.

  20. The UK Infrared Telescope M 33 monitoring project - V. The star formation history across the galactic disc

    Science.gov (United States)

    Javadi, Atefeh; van Loon, Jacco Th.; Khosroshahi, Habib G.; Tabatabaei, Fatemeh; Hamedani Golshan, Roya; Rashidi, Maryam

    2017-01-01

    We have conducted a near-infrared monitoring campaign at the UK Infrared Telescope of the Local Group spiral galaxy M 33 (Triangulum). On the basis of their variability, we have identified stars in the very final stage of their evolution, and for which the luminosity is more directly related to the birth mass than the more numerous less-evolved giant stars that continue to increase in luminosity. In this fifth paper of the series, we construct the birth mass function and hence derive the star formation history across the galactic disc of M 33. The star formation rate has varied between ˜0.010 ± 0.001 (˜0.012 ± 0.007) and 0.060±0.005 (0.052±0.009) M⊙ yr-1 kpc-2 statistically (systematically) in the central square kiloparsec of M 33, comparable with the values derived previously with another camera. The total star formation rate in M 33 within a galactocentric radius of 14 kpc has varied between ˜0.110 ± 0.005 (˜0.174 ± 0.060) and ˜0.560 ± 0.028 (˜0.503 ± 0.100) M⊙ yr-1 statistically (systematically). We find evidence of two epochs during which the star formation rate was enhanced by a factor of a few - one that started ˜6 Gyr ago and lasted ˜3 Gyr and produced ≥71 per cent of the total mass in stars, and one ˜250 Myr ago that lasted ˜200 Myr and formed ≤13 per cent of the mass in stars. Radial star formation history profiles suggest that the inner disc of M 33 was formed in an inside-out formation scenario. The outskirts of the disc are dominated by the old population, which may be the result of dynamical effects over many Gyr. We find correspondence to spiral structure for all stars, but enhanced only for stars younger than ˜100 Myr; this suggests that the spiral arms are transient features and not a part of a global density wave potential.

  1. Grassland bird productivity in warm season grass fields in southwest Wisconsin

    Science.gov (United States)

    Byers, Carolyn M.; Ribic, Christine; Sample, David W.; Dadisman, John D.; Guttery, Michael

    2017-01-01

    Surrogate grasslands established through federal set-aside programs, such as U.S. Department of Agriculture's Conservation Reserve Program (CRP), provide important habitat for grassland birds. Warm season grass CRP fields as a group have the potential for providing a continuum of habitat structure for breeding birds, depending on how the fields are managed and their floristic composition. We studied the nesting activity of four obligate grassland bird species, Bobolink (Dolichonyx oryzivorus), Eastern Meadowlark (Sturnella magna), Grasshopper Sparrow (Ammodramus savannarum), and Henslow's Sparrow (A. henslowii), in relation to vegetative composition and fire management in warm season CRP fields in southwest Wisconsin during 2009–2011. Intraspecific variation in apparent nest density was related to the number of years since the field was burned. Apparent Grasshopper Sparrow nest density was highest in the breeding season immediately following spring burns, apparent Henslow's Sparrow nest density was highest 1 y post burn, and apparent Bobolink and Eastern Meadowlark nest densities were higher in post fire years one to three. Grasshopper Sparrow nest density was highest on sites with more diverse vegetation, specifically prairie forbs, and on sites with shorter less dense vegetation. Bobolink, Eastern Meadowlark, and Henslow's Sparrow apparent nest densities were higher on sites with deeper litter; litter was the vegetative component that was most affected by spring burns. Overall nest success was 0.487 for Bobolink (22 d nesting period), 0.478 for Eastern Meadowlark (25 d nesting period), 0.507 for Grasshopper Sparrow (22 d nesting period), and 0.151 for Henslow's Sparrow (21 d nesting period). The major nest predators were grassland-associated species: thirteen-lined ground squirrel (Ictidomys tridecemlineatus), striped skunk (Mephitis mephitis), milk snake (Lampropeltis triangulum), American badger (Taxidea taxus), and western fox snake (Elaphe vulpina). Overall

  2. Orbits of massive satellite galaxies - II. Bayesian estimates of the Milky Way and Andromeda masses using high-precision astrometry and cosmological simulations

    Science.gov (United States)

    Patel, Ekta; Besla, Gurtina; Mandel, Kaisey

    2017-07-01

    In the era of high-precision astrometry, space observatories like the Hubble Space Telescope (HST) and Gaia are providing unprecedented 6D phase-space information of satellite galaxies. Such measurements can shed light on the structure and assembly history of the Local Group, but improved statistical methods are needed to use them efficiently. Here we illustrate such a method using analogues of the Local Group's two most massive satellite galaxies, the Large Magellanic Cloud (LMC) and Triangulum (M33), from the Illustris dark-matter-only cosmological simulation. We use a Bayesian inference scheme combining measurements of positions, velocities and specific orbital angular momenta (j) of the LMC/M33 with importance sampling of their simulated analogues to compute posterior estimates of the Milky Way (MW) and Andromeda's (M31) halo masses. We conclude that the resulting host halo mass is more susceptible to bias when using measurements of the current position and velocity of satellites, especially when satellites are at short-lived phases of their orbits (I.e. at pericentre). Instead, the j value of a satellite is well conserved over time and provides a more reliable constraint on host mass. The inferred virial mass of the MW (M31) using j of the LMC (M33) is {{M}}_{vir, MW} = 1.02^{+0.77}_{-0.55} × 10^{12} M⊙ ({{M}}_{vir, M31} = 1.37^{+1.39}_{-0.75} × 10^{12} M⊙). Choosing simulated analogues whose j values are consistent with the conventional picture of a previous (<3 Gyr ago), close encounter (<100 kpc) of M33 about M31 results in a very low virial mass for M31 (˜1012 M⊙). This supports the new scenario put forth in Patel, Besla & Sohn, wherein M33 is on its first passage about M31 or on a long-period orbit. We conclude that this Bayesian inference scheme, utilizing satellite j, is a promising method to reduce the current factor of 2 spread in the mass range of the MW and M31. This method is easily adaptable to include additional satellites as new 6D

  3. MSU Contributes to New Research on Star Formation

    Science.gov (United States)

    2010-01-01

    constellation called Triangulum Australe, in a giant cluster of galaxies called Abell 3627. It is associated with a galaxy known as ESO 137-001 which is about 219 million light years from our own Milky Way Galaxy. Star formation is a continuous process throughout the universe, where there are estimated to be billions of galaxies, each of which contains billions of stars. Stars are formed from clouds of dusty, cool, dense molecular gas. Molecular gas clouds prefer to inhabit galaxies, particularly the disks of galaxies like the Milky Way. Our sun, a star located within the Milky Way Galaxy, is an average-size star estimated to be about 4.6 billion years old. Michigan State University has been advancing knowledge and transforming lives through innovative teaching, research and outreach for more than 150 years. MSU is known internationally as a major public university with global reach and extraordinary impact. Its 17 degree-granting colleges attract scholars worldwide who are interested in combining education with practical problem solving. More information, including images and other multimedia, can be found at: http://chandra.harvard.edu and http://chandra.nasa.gov

  4. The Milky Way's Tiny but Tough Galactic Neighbour

    Science.gov (United States)

    2009-10-01

    Today ESO announces the release of a stunning new image of one of our nearest galactic neighbours, Barnard's Galaxy, also known as NGC 6822. The galaxy contains regions of rich star formation and curious nebulae, such as the bubble clearly visible in the upper left of this remarkable vista. Astronomers classify NGC 6822 as an irregular dwarf galaxy because of its odd shape and relatively diminutive size by galactic standards. The strange shapes of these cosmic misfits help researchers understand how galaxies interact, evolve and occasionally "cannibalise" each other, leaving behind radiant, star-filled scraps. In the new ESO image, Barnard's Galaxy glows beneath a sea of foreground stars in the direction of the constellation of Sagittarius (the Archer). At the relatively close distance of about 1.6 million light-years, Barnard's Galaxy is a member of the Local Group, the archipelago of galaxies that includes our home, the Milky Way. The nickname of NGC 6822 comes from its discoverer, the American astronomer Edward Emerson Barnard, who first spied this visually elusive cosmic islet using a 125-millimetre aperture refractor in 1884. Astronomers obtained this latest portrait using the Wide Field Imager (WFI) attached to the 2.2-metre MPG/ESO telescope at ESO's La Silla Observatory in northern Chile. Even though Barnard's Galaxy lacks the majestic spiral arms and glowing, central bulge that grace its big galactic neighbours, the Milky Way, the Andromeda and the Triangulum galaxies, this dwarf galaxy has no shortage of stellar splendour and pyrotechnics. Reddish nebulae in this image reveal regions of active star formation, where young, hot stars heat up nearby gas clouds. Also prominent in the upper left of this new image is a striking bubble-shaped nebula. At the nebula's centre, a clutch of massive, scorching stars send waves of matter smashing into the surrounding interstellar material, generating a glowing structure that appears ring-like from our perspective

  5. The Unexpected Past of a Dwarf Galaxy

    Science.gov (United States)

    1996-08-01

    in this way effectively `eats' the smaller one. Thus the Milky Way may contain the remains of many smaller galaxies it has met and consumed in the past. A natural consequence of this theory is that the Milky Way halo may at least partially consist of stars which originally belonged to these smaller galaxies. However, it is also possible that some of the halo stars formed during the early collapse of the gas cloud from which the Milky Way formed. Like the Milky Way, the two nearest, large spiral galaxies (the Andromeda nebula and M33 in the neighbouring Triangulum constellation) are also surrounded by halos of old stars. Contrarily, investigations of the smaller galaxies in the Local Group have until now not shown that they possess such halos. These dwarf galaxies greatly outnumber the large spiral galaxies - to date about two dozen are known - and they are considered to be the last survivors of the earlier cannibalism phase. The nearest are the well-known Magellanic Clouds, about 170,000 (Large Cloud) and 250,000 light years distant (Small Cloud). They can be seen with the unaided eye from the Southern hemisphere. Recent studies indicate that they orbit the Milky Way and that they may eventually fall prey to our galaxy in a future round of cannibalism. So far, no evidence has been found of an old halo around the Magellanic Clouds. This does not necessarily imply that all dwarf galaxies must likewise lack halos: it is also possible that the halos of the Magellanic Clouds were stripped away when they came too close to the Milky Way sometime in the past. The isolated WLM dwarf galaxy Down in the southern sky, in the constellation of Cetus (the Whale or the Sea Monster), lies a relative faint and distant, small galaxy which astronomers normally refer to as the WLM dwarf galaxy . It was first seen in 1909 by the famous astrophotographer Max Wolf on photographic plates obtained at the Heidelberg Observatory (Germany), but it was only in 1926 that its true nature was