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Sample records for wing venation patterns

  1. Variable wing venation in Agathiphaga (Lepidoptera: Agathiphagidae) is key to understanding the evolution of basal moths

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    Gibbs, George W.

    2016-01-01

    Details of the ancestral groundplan of wing venation in moths remain uncertain, despite approximately a century of study. Here, we describe a 3-branched subcostal vein, a 5-branched medial vein and a 2-branched cubitus posterior vein on the forewing of Agathiphaga vitiensis Dumbleton 1952 from Vanuatu. Such veins had not previously been described in any Lepidoptera. Because wing veins are typically lost during lepidopteran evolutionary history, rarely—if ever—to be regained, the venation of A. vitiensis probably represents the ancestral character state for moths. Wing venation is often used to identify fossil insects as moths, because wing scales are not always preserved; the presence of a supposedly trichopteran 3-branched subcostal vein in crown Lepidoptera may decrease the certainty with which certain amphiesmenopteran fossils from the Mesozoic can be classified. And because plesiomorphic veins can influence the development of lepidopteran wing patterns even if not expressed in the adult wing, the veins described here may determine the location of wing pattern elements in many lepidopteran taxa. PMID:27853559

  2. Basal Complex and Basal Venation of Odonata Wings: Structural Diversity and Potential Role in the Wing Deformation.

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    H Rajabi

    Full Text Available Dragonflies and damselflies, belonging to the order Odonata, are known to be excellent fliers with versatile flight capabilities. The ability to fly over a wide range of speeds, high manoeuvrability and great agility are a few characteristics of their flight. The architecture of the wings and their structural elements have been found to play a major role in this regard. However, the precise influence of individual wing components on the flight performance of these insects remains unknown. The design of the wing basis (so called basal complex and the venation of this part are responsible for particular deformability and specific shape of the wing blade. However, the wing bases are rather different in representatives of different odonate groups. This presumably reflects the dimensions of the wings on one hand, and different flight characteristics on the other hand. In this article, we develop the first three-dimensional (3D finite element (FE models of the proximal part of the wings of typical representatives of five dragonflies and damselflies families. Using a combination of the basic material properties of insect cuticle, a linear elastic material model and a nonlinear geometric analysis, we simulate the mechanical behaviour of the wing bases. The results reveal that although both the basal venation and the basal complex influence the structural stiffness of the wings, it is only the latter which significantly affects their deformation patterns. The use of numerical simulations enabled us to address the role of various wing components such as the arculus, discoidal cell and triangle on the camber formation in flight. Our study further provides a detailed representation of the stress concentration in the models. The numerical analysis presented in this study is not only of importance for understanding structure-function relationship of insect wings, but also might help to improve the design of the wings for biomimetic micro-air vehicles (MAVs.

  3. Leaf venation pattern to recognize austral South American medicinal species of "cow's hoof" (Bauhinia L., Fabaceae

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    Renée H. Fortunato

    Full Text Available ABSTRACT The leaves extracts of some species of Bauhinia L. s.l. are consumed to treat diabetes, inflammation, pains and several disorders in traditional medicine in austral South America. Despite its wide use and commercialization, sale is not controlled, and botanical quality of samples is not always adequate because of plant misidentification and adulteration. Here, we characterized leaf vein pattern in nineteen taxa to contribute to the recognition and commercial quality control of plant material commercially available. The vein characters intercostal tertiary and quinternary vein fabric, areole development and shape, free ending veinlet branching and marginal ultimate venation allowed to distinguish the main medicinal species in the region.

  4. A Remarkable New Family of Jurassic Insects (Neuroptera) with Primitive Wing Venation and Its Phylogenetic Position in Neuropterida

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    Yang, Qiang; Makarkin, Vladimir N.; Winterton, Shaun L.; Khramov, Alexander V.; Ren, Dong

    2012-01-01

    Background Lacewings (insect order Neuroptera), known in the fossil record since the Early Permian, were most diverse in the Mesozoic. A dramatic variety of forms ranged in that time from large butterfly-like Kalligrammatidae to minute two-winged Dipteromantispidae. Principal Findings We describe the intriguing new neuropteran family Parakseneuridae fam. nov. with three new genera and 15 new species from the Middle Jurassic of Daohugou (Inner Mongolia, China) and the Early/Middle Jurassic of Sai-Sagul (Kyrgyzstan): Parakseneura undula gen. et sp. nov., P. albomacula gen. et sp. nov., P. curvivenis gen. et sp. nov., P. nigromacula gen. et sp. nov., P. nigrolinea gen. et sp. nov., P. albadelta gen. et sp. nov., P. cavomaculata gen. et sp. nov., P. inflata gen. et sp. nov., P. metallica gen. et sp. nov., P. emarginata gen. et sp. nov., P. directa gen. et sp. nov., Pseudorapisma jurassicum gen. et sp. nov., P. angustipenne gen. et sp. nov., P. maculatum gen. et sp. nov. (Daohugou); Shuraboneura ovata gen. et sp. nov. (Sai-Sagul). The family comprises large neuropterans with most primitive wing venation in the order indicated by the presence of ScA and AA1+2, and the dichotomous branching of MP, CuA, CuP, AA3+4, AP1+2. The phylogenetic position of Parakseneuridae was investigated using a phylogenetic analysis of morphological scoring for 33 families of extinct and extant Neuropterida combined with DNA sequence data for representatives of all extant families. Parakseneuridae were recovered in a clade with Osmylopsychopidae, Prohemerobiidae, and Ithonidae. Conclusions/Significance The presence of the presumed AA1+2 in wings of Parakseneuridae is a unique plesiomorphic condition hitherto unknown in Neuropterida, the clade comprising Neuroptera, Megaloptera, Raphidioptera. The relative uncertainty of phylogenetic position of Parakseneuridae and the majority of other families of Neuroptera reflects deficient paleontological data, especially from critical important periods

  5. Stable structural color patterns displayed on transparent insect wings.

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    Shevtsova, Ekaterina; Hansson, Christer; Janzen, Daniel H; Kjærandsen, Jostein

    2011-01-11

    Color patterns play central roles in the behavior of insects, and are important traits for taxonomic studies. Here we report striking and stable structural color patterns--wing interference patterns (WIPs)--in the transparent wings of small Hymenoptera and Diptera, patterns that have been largely overlooked by biologists. These extremely thin wings reflect vivid color patterns caused by thin film interference. The visibility of these patterns is affected by the way the insects display their wings against various backgrounds with different light properties. The specific color sequence displayed lacks pure red and matches the color vision of most insects, strongly suggesting that the biological significance of WIPs lies in visual signaling. Taxon-specific color patterns are formed by uneven membrane thickness, pigmentation, venation, and hair placement. The optically refracted pattern is also stabilized by microstructures of the wing such as membrane corrugations and spherical cell structures that reinforce the pattern and make it essentially noniridescent over a large range of light incidences. WIPs can be applied to map the micromorphology of wings through direct observation and are useful in several fields of biology. We demonstrate their usefulness as identification patterns to solve cases of cryptic species complexes in tiny parasitic wasps, and indicate their potentials for research on the genetic control of wing development through direct links between the transregulatory wing landscape and interference patterns we observe in Drosophila model species. Some species display sexually dimorphic WIPs, suggesting sexual selection as one of the driving forces for their evolution.

  6. An automated or semi-automated identification system using venation pattern to delimit Indian leaf drugs: A proposal

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    Selvam, A.B.D.

    2010-01-01

    About 7,200 medicinal plants are known to occur in India, of which, the leaves of a few hundred plants have medicinal properties. Identification of leaf drugs using venation is considered as one of the most reliable and convenient methods. Leaf identification by mechanical means may often lead to wrong identification. Due to the growing volume of illegal trade/malpractice in the crude drug industry on the one hand and lack of sufficient experts on the other, a much faster, convenient and reliable method is mandatory for the identification of Indian leaf drugs. Therefore, a new automated or semi-automated identification system based on venation pattern is inevitable for the present day condition to identify and authenticate the leaves of Indian medicinal plants. PMID:21713143

  7. Expression of Genes Involved in Drosophila Wing Morphogenesis and Vein Patterning Are Altered by Spaceflight

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    Parsons-Wingerter, Patricia A.; Hosamani, Ravikumar; Bhattacharya, Sharmila

    2015-01-01

    Imaginal wing discs of Drosophila melanogaster (fruit fly) defined during embryogenesis ultimately result in mature wings of stereotyped (specific) venation patterning. Major regulators of wing disc development are the epidermal growth factor receptor (EGF), Notch, Hedgehog (Hh), Wingless (Wg), and Dpp signaling pathways. Highly stereotyped vascular patterning is also characteristic of tissues in other organisms flown in space such as the mouse retina and leaves of Arabidopsis thaliana. Genetic and other adaptations of vascular patterning to space environmental factors have not yet been systematically quantified, despite widespread recognition of their critical importance for terrestrial and microgravity applications. Here we report changes in gene expression with space flight related to Drosophila wing morphogenesis and vein patterning. In addition, genetically modified phenotypes of increasingly abnormal ectopic wing venation in the Drosophila wing1 were analyzed by NASA's VESsel GENeration Analysis (VESGEN) software2. Our goal is to further develop insightful vascular mappings associated with bioinformatic dimensions of genetic or other molecular phenotypes for correlation with genetic and other molecular profiling relevant to NASA's GeneLab and other Space Biology exploration initiatives.

  8. Gradual and contingent evolutionary emergence of leaf mimicry in butterfly wing patterns.

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    Suzuki, Takao K; Tomita, Shuichiro; Sezutsu, Hideki

    2014-11-25

    Special resemblance of animals to natural objects such as leaves provides a representative example of evolutionary adaptation. The existence of such sophisticated features challenges our understanding of how complex adaptive phenotypes evolved. Leaf mimicry typically consists of several pattern elements, the spatial arrangement of which generates the leaf venation-like appearance. However, the process by which leaf patterns evolved remains unclear. In this study we show the evolutionary origin and process for the leaf pattern in Kallima (Nymphalidae) butterflies. Using comparative morphological analyses, we reveal that the wing patterns of Kallima and 45 closely related species share the same ground plan, suggesting that the pattern elements of leaf mimicry have been inherited across species with lineage-specific changes of their character states. On the basis of these analyses, phylogenetic comparative methods estimated past states of the pattern elements and enabled reconstruction of the wing patterns of the most recent common ancestor. This analysis shows that the leaf pattern has evolved through several intermediate patterns. Further, we use Bayesian statistical methods to estimate the temporal order of character-state changes in the pattern elements by which leaf mimesis evolved, and show that the pattern elements changed their spatial arrangement (e.g., from a curved line to a straight line) in a stepwise manner and finally establish a close resemblance to a leaf venation-like appearance. Our study provides the first evidence for stepwise and contingent evolution of leaf mimicry.  Leaf mimicry patterns evolved in a gradual, rather than a sudden, manner from a non-mimetic ancestor. Through a lineage of Kallima butterflies, the leaf patterns evolutionarily originated through temporal accumulation of orchestrated changes in multiple pattern elements.

  9. Stamen venation in the Winteraceae

    NARCIS (Netherlands)

    Sampson, F.B.

    1987-01-01

    Stamen venation is described for all species of Pseudowintera and an interspecific hybrid; the single species of Exospermum, several species of Belliolum, Drimys, Tasmannia (syn. Drimys sect. Tasmannia) and Zygogynum, as well as one species of Bubbia. One, two and three-veined stamens were found in

  10. Artificial insect wings of diverse morphology for flapping-wing micro air vehicles

    International Nuclear Information System (INIS)

    Shang, J K; Finio, B M; Wood, R J; Combes, S A

    2009-01-01

    The development of flapping-wing micro air vehicles (MAVs) demands a systematic exploration of the available design space to identify ways in which the unsteady mechanisms governing flapping-wing flight can best be utilized for producing optimal thrust or maneuverability. Mimicking the wing kinematics of biological flight requires examining the potential effects of wing morphology on flight performance, as wings may be specially adapted for flapping flight. For example, insect wings passively deform during flight, leading to instantaneous and potentially unpredictable changes in aerodynamic behavior. Previous studies have postulated various explanations for insect wing complexity, but there lacks a systematic approach for experimentally examining the functional significance of components of wing morphology, and for determining whether or not natural design principles can or should be used for MAVs. In this work, a novel fabrication process to create centimeter-scale wings of great complexity is introduced; via this process, a wing can be fabricated with a large range of desired mechanical and geometric characteristics. We demonstrate the versatility of the process through the creation of planar, insect-like wings with biomimetic venation patterns that approximate the mechanical properties of their natural counterparts under static loads. This process will provide a platform for studies investigating the effects of wing morphology on flight dynamics, which may lead to the design of highly maneuverable and efficient MAVs and insight into the functional morphology of natural wings.

  11. Sexual selection on wing interference patterns in Drosophila melanogaster.

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    Katayama, Natsu; Abbott, Jessica K; Kjærandsen, Jostein; Takahashi, Yuma; Svensson, Erik I

    2014-10-21

    Animals with color vision use color information in intra- and interspecific communication, which in turn may drive the evolution of conspicuous colored body traits via natural and sexual selection. A recent study found that the transparent wings of small flies and wasps in lower-reflectance light environments display vivid and stable structural color patterns, called "wing interference patterns" (WIPs). Such WIPs were hypothesized to function in sexual selection among small insects with wing displays, but this has not been experimentally verified. Here, to our knowledge we present the first experimental evidence that WIPs in males of Drosophila melanogaster are targets of mate choice from females, and that two different color traits--saturation and hue--experience directional and stabilizing sexual selection, respectively. Using isogenic lines from the D. melanogaster Genetic Reference Panel, we compare attractiveness of different male WIPs against black and white visual backgrounds. We show that males with more vivid wings are more attractive to females than are males with dull wings. Wings with a large magenta area (i.e., intermediate trait values) were also preferred over those with a large blue or yellow area. These experimental results add a visual element to the Drosophila mating array, integrating sexual selection with elements of genetics and evo-devo, potentially applicable to a wide array of small insects with hyaline wings. Our results further underscore that the mode of sexual selection on such visual signals can differ profoundly between different color components, in this case hue and saturation.

  12. Studies on Leaf Venation in Selected Taxa of the Genus Ficus L. (Moraceae) in Peninsular Malaysia.

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    Badron, Ummu Hani; Talip, Noraini; Mohamad, Abdul Latiff; Affenddi, Affina Eliya Aznal; Juhari, Amirul Aiman Ahmad

    2014-12-01

    A study on the variation of leaf venation patterns was conducted on 21 taxa of the genus Ficus in Peninsular Malaysia. The results showed the existence of eight leaf venation patterns based on veinlets, the ultimate marginal and areolar venation. The majority of species, such as F. annulata, F. benghalensis, F. benjamina, F. deltoidea var. angustifolia, F. deltoidea var. kunstleri, F. depressa, F. elastica, F. hispida, F. microcarpa, F. religiosa, F. tinctoria, F. ucinata and F. vasculosa, show tri-veinlets. The others exhibit the following: bi-veinlets in F. aurata and F. heteropleura; uni-veinlets in F. lepicarpa, F. schwarzii and F. superba; and simple veinlets in F. aurantiacea and F. fulva. F. sagittata presents no veinlets for areolar venation. The presence of tracheid or swollen veins at the centre of the lamina and the presence of cystolith cells and trichomes are common anatomical characteristics that could assist in group classification of the studied species. Variations in leaf venation patterns are not only valuable in identifying a taxon group, but can also be used to differentiate between species in the genus Ficus.

  13. Wake patterns of the wings and tail of hovering hummingbirds

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    Altshuler, Douglas L.; Princevac, Marko; Pan, Hansheng; Lozano, Jesse

    The flow fields of slowly flying bats and fasterflying birds differ in that bats produce two vortex loops during each stroke, one per wing, and birds produce a single vortex loop per stroke. In addition, the circulation at stroke transition approaches zero in bats but remains strong in birds. It is unknown if these difference derive from fundamental differences in wing morphology or are a consequence of flight speed. Here, we present an analysis of the horizontal flow field underneath hovering Anna's hummingbirds (Calypte anna) to describe the wake of a bird flying at zero forward velocity. We also consider how the hummingbird tail interacts with the wake generated by the wings. High-speed image recording and analysis from three orthogonal perspectives revealed that the wing tips reach peak velocities in the middle of each stroke and approach zero velocity at stroke transition. Hummingbirds use complex tail kinematic patterns ranging from in phase to antiphase cycling with respect to the wings, covering several phase shifted patterns. We employed particle image velocimetry to attain detailed horizontal flow measurements at three levels with respect to the tail: in the tail, at the tail tip, and just below the tail. The velocity patterns underneath the wings indicate that flow oscillates along the ventral-dorsal axis in response to the down- and up-strokes and that the sideways flows with respect to the bird are consistently from the lateral to medial. The region around the tail is dominated by axial flows in dorsal to ventral direction. We propose that these flows are generated by interaction between the wakes of the two wings at the end of the upstroke, and that the tail actively defects flows to generate moments that contribute to pitch stability. The flow fields images also revealed distinct vortex loops underneath each wing, which were generated during each stroke. From these data, we propose a model for the primary flow structures of hummingbirds that more

  14. Waiting in the wings: what can we learn about gene co-option from the diversification of butterfly wing patterns?

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    Jiggins, Chris D; Wallbank, Richard W R; Hanly, Joseph J

    2017-02-05

    A major challenge is to understand how conserved gene regulatory networks control the wonderful diversity of form that we see among animals and plants. Butterfly wing patterns are an excellent example of this diversity. Butterfly wings form as imaginal discs in the caterpillar and are constructed by a gene regulatory network, much of which is conserved across the holometabolous insects. Recent work in Heliconius butterflies takes advantage of genomic approaches and offers insights into how the diversification of wing patterns is overlaid onto this conserved network. WntA is a patterning morphogen that alters spatial information in the wing. Optix is a transcription factor that acts later in development to paint specific wing regions red. Both of these loci fit the paradigm of conserved protein-coding loci with diverse regulatory elements and developmental roles that have taken on novel derived functions in patterning wings. These discoveries offer insights into the 'Nymphalid Ground Plan', which offers a unifying hypothesis for pattern formation across nymphalid butterflies. These loci also represent 'hotspots' for morphological change that have been targeted repeatedly during evolution. Both convergent and divergent evolution of a great diversity of patterns is controlled by complex alleles at just a few genes. We suggest that evolutionary change has become focused on one or a few genetic loci for two reasons. First, pre-existing complex cis-regulatory loci that already interact with potentially relevant transcription factors are more likely to acquire novel functions in wing patterning. Second, the shape of wing regulatory networks may constrain evolutionary change to one or a few loci. Overall, genomic approaches that have identified wing patterning loci in these butterflies offer broad insight into how gene regulatory networks evolve to produce diversity.This article is part of the themed issue 'Evo-devo in the genomics era, and the origins of morphological

  15. Leaf venations in some Ficus L. (Moraceae) species

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    Siti-Khaulah, A. R.; Noraini, T.

    2016-11-01

    A study on the leaf venation anatomical characteristics was conducted on eight chosen Ficus L. (Moraceae) species in Peninsular Malaysia, namely F. callosa, F. apiocarpa, F. caulocarpa, F. pubigera, F. pendens, F. celebensis, F. mollissima, and F. drupacea. The objective of this study was to observe the significant leaf venation anatomical characteristics that can be used in species classification and identification. Leaf clearing, staining, mounting and observation under a light microscope were techniques used for the study. Veinlets, the ultimate marginal and areolar venation were the main leaf venation anatomical characteristics observed in this study. The presence of complex veinlets in the areolar venation were shown in most species studied (F. callosa, F. pubigera, F. celebensis, F. pendens), whilst simple or uni-veinlets were shown in F. apiocarpa, F. caulocarpa and F. mollissima. Free ending veinlet was absence in F. drupacea. Complete ultimate marginal venation was shown in most species studied, such as in F. callosa, F. caulocarpa, F. pendends, F. mollissima and F. drupacea. Opened areolar venation was observed in most species studied, such as in F. callosa, F. pubigera, F. pendens, F. mollissima, and F. celebensis. Diagnostic leaf venation anatomical characteristic was present in cystolith cells in F. pubigera and trichomes were present on venation in F. mollissima only. As a conclusion, the results of this study have shown that leaf venation anatomical characteristics have taxonomic significance that can be used in differentiation and identification in selected Ficus species studied.

  16. Comparative insights into questions of lepidopteran wing pattern homology

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    Stockslager Steven

    2006-11-01

    Full Text Available Abstract Background Butterfly and moth eyespots can share a similar appearance, involving multiple concentric rings of colored scales, but usually occuring in non-homologous positions on the wing. Within the butterflies, on the other hand, spots that share the same homologous position may not share the concentric ring structure; and, in butterfly species that have eyespots with concentric rings, ectopic eyespots with a similar ring structure can be induced by means of a simple epidermal wound. The extent to which all these eyespots, natural or induced, share similar genes and developmental mechanisms is investigated here by means of protein in-situ localizations in selected butterfly and moth species. In addition to looking at some of the transcription factors previously identified as being involved in eyespot formation, we also tested the involvement of candidate genes from the Wingless and TGF-β signaling pathways as putative morphogens for eyespot development. Results Saturniid moth and nymphalid butterfly eyespots with concentric rings of color express at least two transcription factors, Distal-less and Engrailed, in the center of the future pattern. Nymphalid eyespots centers also express the ligand Wingless and an activated signal transducer, a phosphorylated Smad protein, but neither these proteins nor the previous two proteins are found in pierid spot centers, which consist of a single patch of color. Both butterfly wing patterns, however, express a third transcription factor, Spalt, a portion of whose expression domain maps to the black scales on the adult wing. Wounding a nymphalid wing, on the other hand, leads to upregulation of Distal-less, engrailed and spalt in subsets of cells around the wounding site, mimicking concentric eyespot development. Conclusion Wingless and TGF-β ligands are both candidate morphogens involved in nymphalid butterfly eyespot formation. These eyespots, as well as saturniid moth eyespots with concentric

  17. Resilin-based rubber-like elastic elements in the insect wing: an overview

    Energy Technology Data Exchange (ETDEWEB)

    Gorb, S.N. [Max-Planck-Institut fuer Entwicklungsbiologie, Tuebingen (Germany)

    2000-07-01

    This report summarises data on the presence of resilin, a rubber-like protein in the flexibly-joined junctions of wing veins. This protein is a substance which is responsible for elastic energy storage. Previously this protein has been described in jumping systems of beetles, fleas and leafhoppers abdominal cuticle of workers of the honey-ants and queen termites; eye lens cuticle of dragonflies and the food-pump of reduviid bugs. It is also known in tendons of the dragonfly flight muscles. This protein has not been previously described in the distal regions of damselfly wings. The pattern of resilin distribution in the insect wings is one of several mechanisms which are responsible for automatic performance of passive wing movements. Any folding of the distal wing parts can not be achieved by local muscles, but must be done by remote (thoracic) muscles or local elasticity. Many insects, such as earwigs and beetles, have developed complex mechanisms of wing folding, which are advantagenous for insects living in soil or other narrow substrata. The folding pattern depends on the wing geometry, venation pattern, and material properties of structures involved. Thus design of wings with folding function has an additional complexity: their design is a kind of compromise between flight and folding. (orig.)

  18. Cryptically patterned moths perceive bark structure when choosing body orientations that match wing color pattern to the bark pattern.

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    Chang-Ku Kang

    Full Text Available Many moths have wing patterns that resemble bark of trees on which they rest. The wing patterns help moths to become camouflaged and to avoid predation because the moths are able to assume specific body orientations that produce a very good match between the pattern on the bark and the pattern on the wings. Furthermore, after landing on a bark moths are able to perceive stimuli that correlate with their crypticity and are able to re-position their bodies to new more cryptic locations and body orientations. However, the proximate mechanisms, i.e. how a moth finds an appropriate resting position and orientation, are poorly studied. Here, we used a geometrid moth Jankowskia fuscaria to examine i whether a choice of resting orientation by moths depends on the properties of natural background, and ii what sensory cues moths use. We studied moths' behavior on natural (a tree log and artificial backgrounds, each of which was designed to mimic one of the hypothetical cues that moths may perceive on a tree trunk (visual pattern, directional furrow structure, and curvature. We found that moths mainly used structural cues from the background when choosing their resting position and orientation. Our findings highlight the possibility that moths use information from one type of sensory modality (structure of furrows is probably detected through tactile channel to achieve crypticity in another sensory modality (visual. This study extends our knowledge of how behavior, sensory systems and morphology of animals interact to produce crypsis.

  19. On Pattern Formation Mechanisms for Lepidopteran Wing Patterns and Mammalian Coat Markings

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    Murray, J. D.

    1981-10-01

    The patterns on wings of Lepidoptera can be generated with a few pattern elements, but no mechanism has been suggested for producing them. I consider two of the basic patterns, namely, central symmetry and dependent patterns. A biochemically plausible model mechanism is proposed for generating major aspects of these patterns, based on a diffusing morphogen that activates a gene or colour-specific enzyme in a threshold manner to generate a stable heterogeneous spatial pattern. The model is applied to the determination stream hypothesis of Kuhn & von Engelhardt (Wilhelm Roux Arch. Entw Mech. Org. 130, 660 (1933)), and results from the model compared with their microcautery experiments on the pupal wing of Ephestia kuhniella. In the case of dependent patterns, results are compared with patterns on specific Papilionidae. For the same mechanism and a fixed set of parameters I demonstrate the important roles of geometry and scale on the spatial patterns obtained. The results and evidence presented here suggest the existence of diffusion fields of the order of several millimetres, which are very much larger than most embryonic fields. The existence of zones of polarizing activity is also indicated. Colour patterns on animals are considered to be genetically determined, but the mechanism is not known. I have previously suggested that a single mechanism that can exhibit an infinite variety of patterns is a candidate for that mechanism, and proposed that a reaction-diffusion system that can be diffusively driven unstable could be responsible for the laying down of the spacing patterns that generates the prepattern for animal coat markings. For illustrative purposes I consider a practical reaction mechanism, which exhibits substrate inhibition, and show that the geometry and scale of the domain (part of the epidermis) play a crucial role in the structural patterns that result. Patterns are obtained for a selection of geometries, and general features are related to the coat

  20. Wnt signaling underlies evolution and development of the butterfly wing pattern symmetry systems.

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    Martin, Arnaud; Reed, Robert D

    2014-11-15

    Most butterfly wing patterns are proposed to be derived from a set of conserved pattern elements known as symmetry systems. Symmetry systems are so-named because they are often associated with parallel color stripes mirrored around linear organizing centers that run between the anterior and posterior wing margins. Even though the symmetry systems are the most prominent and diverse wing pattern elements, their study has been confounded by a lack of knowledge regarding the molecular basis of their development, as well as the difficulty of drawing pattern homologies across species with highly derived wing patterns. Here we present the first molecular characterization of symmetry system development by showing that WntA expression is consistently associated with the major basal, discal, central, and external symmetry system patterns of nymphalid butterflies. Pharmacological manipulations of signaling gradients using heparin and dextran sulfate showed that pattern organizing centers correspond precisely with WntA, wingless, Wnt6, and Wnt10 expression patterns, thus suggesting a role for Wnt signaling in color pattern induction. Importantly, this model is supported by recent genetic and population genomic work identifying WntA as the causative locus underlying wing pattern variation within several butterfly species. By comparing the expression of WntA between nymphalid butterflies representing a range of prototypical symmetry systems, slightly deviated symmetry systems, and highly derived wing patterns, we were able to infer symmetry system homologies in several challenging cases. Our work illustrates how highly divergent morphologies can be derived from modifications to a common ground plan across both micro- and macro-evolutionary time scales. Copyright © 2014 Elsevier Inc. All rights reserved.

  1. Wing pattern variation in the Patagonian biting midge, Forcipomyia (Forcipomyia multipicta Ingram & Macfie (Diptera, Ceratopogonidae

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    Gustavo R. SPINELLI

    2012-01-01

    Full Text Available Examination of the type-series and non-type specimens of the Patagonian biting midge, Forcipomyia (Forcipomyia multipicta Ingram & Macfie (Diptera: Ceratopogonidae, revealed considerable variation in wing patterns of both sexes. One pattern includes several distinct light spot areas, whereas another pattern (e.g, in the holotype only features marginal light spots in cell r3, while other light spots are barely perceptible or absent. The cause(s of the differential lack of dark macrotrichia in certain areas of the wing membrane in specimens of some series could not be attributed either to their age, sex, or method of preservation.

  2. Color pattern analysis of nymphalid butterfly wings: revision of the nymphalid groundplan.

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    Otaki, Joji M

    2012-09-01

    To better understand the developmental mechanisms of color pattern variation in butterfly wings, it is important to construct an accurate representation of pattern elements, known as the "nymphalid groundplan". However, some aspects of the current groundplan remain elusive. Here, I examined wing-wide elemental patterns of various nymphalid butterflies and confirmed that wing-wide color patterns are composed of the border, central, and basal symmetry systems. The central and basal symmetry systems can express circular patterns resembling eyespots, indicating that these systems have developmental mechanisms similar to those of the border symmetry system. The wing root band commonly occurs as a distinct symmetry system independent from the basal symmetry system. In addition, the marginal and submarginal bands are likely generated as a single system, referred to as the "marginal band system". Background spaces between two symmetry systems are sometimes light in coloration and can produce white bands, contributing significantly to color pattern diversity. When an element is enlarged with a pale central area, a visually similar (yet developmentally distinct) white band is produced. Based on the symmetric relationships of elements, I propose that both the central and border symmetry systems are comprised of "core elements" (the discal spot and the border ocelli, respectively) and a pair of "paracore elements" (the distal and proximal bands and the parafocal elements, respectively). Both core and paracore elements can be doubled, or outlined. Developmentally, this system configuration is consistent with the induction model, but not with the concentration gradient model for positional information.

  3. Altitudinal Variability in Wing Patterns of Phyllonorycter salictella (Zeller, 1846 (Lepidoptera: Gracillariidae

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    František Gregor

    2017-01-01

    Full Text Available Individual variability of wing patterns is not frequent in European species of the genus Phyllonorycter Hübner, 1822. It is unusually distinct in Phyllonorycter salictella (Zeller, 1846 which causes some taxonomic ambiguities and oversights. We found on numerous material of adults reared from overwintering pupae collected in the Czech and Slovak Republics since 1950 that this variability is related with altitude (temperature. Adults with the distinct generotypic (plesiomorphic wing patterns dominated in submountain populations while individuals with the strongly reduced (apomorphic wing drawings were represented quite negligibly. This ratio was reversed in lowland populations. The observed variability is continuous between extreme variants, it is regionally and trophic unaffected, and more pronounced in males than in females. The results support the view that the questionable taxa such as Phyllonorycter viminiella (Sircom, 1848 and P. heringiella (Grønlien, 1932 represent only individual (local or regional forms of the above species.

  4. Giving eyespots a shiner: Pharmacologic manipulation of the Io moth wing pattern.

    Science.gov (United States)

    Sourakov, Andrei

    2017-01-01

    Our knowledge of wing pattern formation in Lepidoptera has advanced significantly in recent years due to the careful examination of several groups of butterflies. The eyespot is a prominent feature of Lepidoptera wing pattern, especially in the family Saturniidae. The present study examined how sulfated polysaccharides affected the wing pattern formation of the Io moth, Automeris io (Saturniidae).  Prepupae and pupae of this species were subjected to injections of heparin and cold shock. While the cold shock had little to no effect on wing pattern, the aberrations resulting from heparin injections were moderate to profound and depended on the dose and the stage at which injection was made. The changes consisted of expansion of the black ring around the dorsal hindwing eyespots and distortion of discal spots on both dorsal and ventral sides of forewings, suggesting a possible link between genetic controls of these elements. Several different types of scales form the normal color pattern of Automeris io , and heparin-induced changes correspond to changes in shape of scales. The resulting aberrations are dubbed 'Black Eye' and 'Comet Eye.' Other known aberrations of Automeris io eyespots are summarized, illustrated, and named.

  5. The relationship between asymmetry, size and unusual venation in honey bees (Apis mellifera).

    Science.gov (United States)

    Łopuch, S; Tofilski, A

    2016-06-01

    Despite the fact that symmetry is common in nature, it is rarely perfect. Because there is a wide range of phenotypes which differs from the average one, the asymmetry should increase along with deviation. Therefore, the aim of this study was to assess the level of asymmetry in normal individuals as well as in phenodeviants categorized as minor or major based on abnormalities in forewing venation in honey bees. Shape fluctuating asymmetry (FA) was lower in normal individuals and minor phenodeviants compared with major phenodeviants, whereas the former two categories were comparable in drones. In workers and queens, there were not significant differences in FA shape between categories. FA size was significantly lower in normal individuals compared with major phenodeviant drones and higher compared with minor phenodeviant workers. In queens, there were no significant differences between categories. The correlation between FA shape and FA size was significantly positive in drones, and insignificant in workers and queens. Moreover, a considerable level of directional asymmetry was found as the right wing was constantly bigger than the left one. Surprisingly, normal individuals were significantly smaller than minor phenodeviants in queens and drones, and they were comparable with major phenodeviants in all castes. The correlation between wing size and wing asymmetry was negative, indicating that smaller individuals were more asymmetrical. The high proportion of phenodeviants in drones compared with workers and queens confirmed their large variability. Thus, the results of the present study showed that minor phenodeviants were not always intermediate as might have been expected.

  6. The functional basis of wing patterning in Heliconius butterflies: the molecules behind mimicry.

    Science.gov (United States)

    Kronforst, Marcus R; Papa, Riccardo

    2015-05-01

    Wing-pattern mimicry in butterflies has provided an important example of adaptation since Charles Darwin and Alfred Russell Wallace proposed evolution by natural selection >150 years ago. The neotropical butterfly genus Heliconius played a central role in the development of mimicry theory and has since been studied extensively in the context of ecology and population biology, behavior, and mimicry genetics. Heliconius species are notable for their diverse color patterns, and previous crossing experiments revealed that much of this variation is controlled by a small number of large-effect, Mendelian switch loci. Recent comparative analyses have shown that the same switch loci control wing-pattern diversity throughout the genus, and a number of these have now been positionally cloned. Using a combination of comparative genetic mapping, association tests, and gene expression analyses, variation in red wing patterning throughout Heliconius has been traced back to the action of the transcription factor optix. Similarly, the signaling ligand WntA has been shown to control variation in melanin patterning across Heliconius and other butterflies. Our understanding of the molecular basis of Heliconius mimicry is now providing important insights into a variety of additional evolutionary phenomena, including the origin of supergenes, the interplay between constraint and evolvability, the genetic basis of convergence, the potential for introgression to facilitate adaptation, the mechanisms of hybrid speciation in animals, and the process of ecological speciation. Copyright © 2015 by the Genetics Society of America.

  7. The Functional Basis of Wing Patterning in Heliconius Butterflies: The Molecules Behind Mimicry

    Science.gov (United States)

    Kronforst, Marcus R.; Papa, Riccardo

    2015-01-01

    Wing-pattern mimicry in butterflies has provided an important example of adaptation since Charles Darwin and Alfred Russell Wallace proposed evolution by natural selection >150 years ago. The neotropical butterfly genus Heliconius played a central role in the development of mimicry theory and has since been studied extensively in the context of ecology and population biology, behavior, and mimicry genetics. Heliconius species are notable for their diverse color patterns, and previous crossing experiments revealed that much of this variation is controlled by a small number of large-effect, Mendelian switch loci. Recent comparative analyses have shown that the same switch loci control wing-pattern diversity throughout the genus, and a number of these have now been positionally cloned. Using a combination of comparative genetic mapping, association tests, and gene expression analyses, variation in red wing patterning throughout Heliconius has been traced back to the action of the transcription factor optix. Similarly, the signaling ligand WntA has been shown to control variation in melanin patterning across Heliconius and other butterflies. Our understanding of the molecular basis of Heliconius mimicry is now providing important insights into a variety of additional evolutionary phenomena, including the origin of supergenes, the interplay between constraint and evolvability, the genetic basis of convergence, the potential for introgression to facilitate adaptation, the mechanisms of hybrid speciation in animals, and the process of ecological speciation. PMID:25953905

  8. Review of the Australian wolf spider genus Venator (Araneae, Lycosidae).

    Science.gov (United States)

    Framenau, Volker W

    2015-09-11

    Species of the Australian wolf spider genus Venator are reviewed including the type species, V. spenceri Hogg, 1900, from south-eastern Australia and V. immansuetus (Simon, 1909) comb. nov., a common species in south-west Western Australia. Venator marginatus Hogg, 1900 is only known from two female specimens and the genital morphology of this species does not conform to the diagnosis of genus as presented here. Therefore V. marginatus is considered incerta sedis. Venator includes medium-sized (9.0-22 mm body length) wolf spiders of overall brownish colouration, and with a black patch covering the anterior three quarters of the venter. They differ from all other wolf spiders in particular by genitalic characters, namely an elevated atrium of the female epigyne that forms a raised edged against the inverted T-shaped median septum. This edge often corresponds to a retrolateral incision on the tegular apophysis of the male pedipalp. The genus is mainly a representative of the Bassian fauna of the Australian continent where it occurs predominantly in dry sclerophyll forests.

  9. Reading the Leaves: A Comparison of Leaf Rank and Automated Areole Measurement for Quantifying Aspects of Leaf Venation

    Directory of Open Access Journals (Sweden)

    Walton A. Green

    2014-08-01

    Full Text Available The reticulate venation that is characteristic of a dicot leaf has excited interest from systematists for more than a century, and from physiological and developmental botanists for decades. The tools of digital image acquisition and computer image analysis, however, are only now approaching the sophistication needed to quantify aspects of the venation network found in real leaves quickly, easily, accurately, and reliably enough to produce biologically meaningful data. In this paper, we examine 120 leaves distributed across vascular plants (representing 118 genera and 80 families using two approaches: a semiquantitative scoring system called “leaf ranking,” devised by the late Leo Hickey, and an automated image-analysis protocol. In the process of comparing these approaches, we review some methodological issues that arise in trying to quantify a vein network, and discuss the strengths and weaknesses of automatic data collection and human pattern recognition. We conclude that subjective leaf rank provides a relatively consistent, semiquantitative measure of areole size among other variables; that modal areole size is generally consistent across large sections of a leaf lamina; and that both approaches—semiquantitative, subjective scoring; and fully quantitative, automated measurement—have appropriate places in the study of leaf venation.

  10. Antibacterial titanium nano-patterned arrays inspired by dragonfly wings

    Science.gov (United States)

    Bhadra, Chris M.; Khanh Truong, Vi; Pham, Vy T. H.; Al Kobaisi, Mohammad; Seniutinas, Gediminas; Wang, James Y.; Juodkazis, Saulius; Crawford, Russell J.; Ivanova, Elena P.

    2015-11-01

    Titanium and its alloys remain the most popular choice as a medical implant material because of its desirable properties. The successful osseointegration of titanium implants is, however, adversely affected by the presence of bacterial biofilms that can form on the surface, and hence methods for preventing the formation of surface biofilms have been the subject of intensive research over the past few years. In this study, we report the response of bacteria and primary human fibroblasts to the antibacterial nanoarrays fabricated on titanium surfaces using a simple hydrothermal etching process. These fabricated titanium surfaces were shown to possess selective bactericidal activity, eliminating almost 50% of Pseudomonas aeruginosa cells and about 20% of the Staphylococcus aureus cells coming into contact with the surface. These nano-patterned surfaces were also shown to enhance the aligned attachment behavior and proliferation of primary human fibroblasts over 10 days of growth. These antibacterial surfaces, which are capable of exhibiting differential responses to bacterial and eukaryotic cells, represent surfaces that have excellent prospects for biomedical applications.

  11. Antibacterial titanium nano-patterned arrays inspired by dragonfly wings

    Science.gov (United States)

    Bhadra, Chris M.; Khanh Truong, Vi; Pham, Vy T. H.; Al Kobaisi, Mohammad; Seniutinas, Gediminas; Wang, James Y.; Juodkazis, Saulius; Crawford, Russell J.; Ivanova, Elena P.

    2015-01-01

    Titanium and its alloys remain the most popular choice as a medical implant material because of its desirable properties. The successful osseointegration of titanium implants is, however, adversely affected by the presence of bacterial biofilms that can form on the surface, and hence methods for preventing the formation of surface biofilms have been the subject of intensive research over the past few years. In this study, we report the response of bacteria and primary human fibroblasts to the antibacterial nanoarrays fabricated on titanium surfaces using a simple hydrothermal etching process. These fabricated titanium surfaces were shown to possess selective bactericidal activity, eliminating almost 50% of Pseudomonas aeruginosa cells and about 20% of the Staphylococcus aureus cells coming into contact with the surface. These nano-patterned surfaces were also shown to enhance the aligned attachment behavior and proliferation of primary human fibroblasts over 10 days of growth. These antibacterial surfaces, which are capable of exhibiting differential responses to bacterial and eukaryotic cells, represent surfaces that have excellent prospects for biomedical applications. PMID:26576662

  12. Sex Differences in 20-Hydroxyecdysone Hormone Levels Control Sexual Dimorphism in Bicyclus anynana Wing Patterns.

    Science.gov (United States)

    Bhardwaj, Shivam; Prudic, Kathleen L; Bear, Ashley; Dasgupta, Mainak; Wasik, Bethany R; Tong, Xiaoling; Cheong, Wei Fun; Wenk, Markus R; Monteiro, Antónia

    2018-02-01

    In contrast to the important role of hormones in the development of sexual traits in vertebrates (Cox RM, Stenquist DS, Calsbeek R. 2009. Testosterone, growth and the evolution of sexual size dimorphism. J Evol Biol. 22(8):1586-1598.), the differentiation of these traits in insects is attributed almost exclusively to cell-autonomous mechanisms controlled by members of the sex determination pathway (Verhulst EC, van de Zande L. 2015. Double nexus - doublesex is the connecting element in sex determination. Brief Funct Genomics 14(6):396-406.), such as doublesex. Although hormones can shape the development of sexual traits in insects, variation in hormone levels are not conclusively known to cause dimorphism in these traits (Prakash A, Monteiro A. 2016. Molecular mechanisms of secondary sexual trait development in insects. Curr Opin Insect Sci. 17:40-48.). Here, we show that butterflies use sex-specific differences in 20-hydroxyecdysone hormone titers to create sexually dimorphic wing ornaments. Females of the dry season (DS) form of Bicyclus anynana display a larger sexual ornament on their wings than males, whereas in the wet season form both sexes have similarly sized ornaments (Prudic KL, Jeon C, Cao H, Monteiro A. 2011. Developmental plasticity in sexual roles of butterfly species drives mutual sexual ornamentation. Science 331(6013):73-75.). High levels of circulating 20-hydroxyecdysone during larval development in DS females and wet season forms cause proliferation of the cells fated to give rise to this wing ornament, and results in sexual dimorphism in the DS forms. This study advances our understanding of how the environment regulates sex-specific patterns of plasticity of sexual ornaments and conclusively shows that hormones can play a role in the development of secondary sexual traits in insects, just like they do in vertebrates. © The Author 2017. Published by Oxford University Press on behalf of the Society for Molecular Biology and Evolution.

  13. Complex dynamics underlie the evolution of imperfect wing pattern convergence in butterflies.

    Science.gov (United States)

    Finkbeiner, Susan D; Briscoe, Adriana D; Mullen, Sean P

    2017-04-01

    Adaptive radiation is characterized by rapid diversification that is strongly associated with ecological specialization. However, understanding the evolutionary mechanisms fueling adaptive diversification requires a detailed knowledge of how natural selection acts at multiple life-history stages. Butterflies within the genus Adelpha represent one of the largest and most diverse butterfly lineages in the Neotropics. Although Adelpha species feed on an extraordinary diversity of larval hosts, convergent evolution is widespread in this group, suggesting that selection for mimicry may contribute to adaptive divergence among species. To investigate this hypothesis, we conducted predation studies in Costa Rica using artificial butterfly facsimiles. Specifically, we predicted that nontoxic, palatable Adelpha species that do not feed on host plants in the family Rubiaceae would benefit from sharing a locally convergent wing pattern with the presumably toxic Rubiaceae-feeding species via reduced predation. Contrary to expectations, we found that the presumed mimic was attacked significantly more than its locally convergent model at a frequency paralleling attack rates on both novel and palatable prey. Although these data reveal the first evidence for protection from avian predators by the supposed toxic, Rubiaceae-feeding Adelpha species, we conclude that imprecise mimetic patterns have high costs for Batesian mimics in the tropics. © 2017 The Author(s). Evolution © 2017 The Society for the Study of Evolution.

  14. Resilin-based rubber-like elements in the insect wing: an overview

    Energy Technology Data Exchange (ETDEWEB)

    Gorb, S.N. [Max-Planck-Institut fuer Entwicklungsbiologie, Tuebingen (Germany). Abt. Biochemie

    2001-06-01

    Wings of insects are complex structures which are designed to enable controlled torsion essential to flapping flight. Although wing morphology and flight types diversified enormously among a huge number of insect species, the general principle of insect flight remained the same. All muscles are located in the thorax proper, the wing itself is complete without any muscles. Data on the nature and distribution of materials involved in wing design are essential for understanding both the aerodynamics and evolution of insect flight. An insect wing consists of a three-dimensional skeletal network of relatively hard veins which are interconnected through thin membranous areas called cells. These structures consists of cuticle, a biological fibrous composite material, the mechanical properties of which may range from very stiff to flexible, depending on its chemical composition. This report summarises data on the presence of resilin, a rubber-like protein in the flexibly-joined junctions of wing veins. This protein is a substance which is responsible for elastic energy storage. Any folding of the distal wing parts can not be achieved by local muscles, but must be done by remote (thoracic) muscles or local elasticity. Many insects, such as earwigs and beetles, have developed complex mechanisms of wing folding, which are advantageous for insects living in soil or other narrow substrata. The folding pattern depends on the wing geometry, venation pattern, and material properties of structures involved. Thus, design of wings with folding functions has an additional complexity: their design is a kind of compromise between flight and folding. (orig.) [German] Biomechanische Systeme, in denen gummiartige Gelenke vorkommen, reichen von den Sprungsystemen von Kaefern, Floehen, Schaumzikaden und Heuschrecken bis Saugpumpen von Wanzen und Fliegen. Resilin ist ein elastisches Strukturprotein, dessen Name sich vom lateinischen Wort fuer 'zurueckschnellen' herleitet. Sein

  15. Measurement of shape and deformation of insect wing

    Science.gov (United States)

    Yin, Duo; Wei, Zhen; Wang, Zeyu; Zhou, Changqiu

    2018-01-01

    To measure the shape and deformation of an insect wing, a scanning setup adopting laser triangulation and image matching was developed. Only one industry camera with two light sources was employed to scan the transparent insect wings. 3D shape and point to point full field deformation of the wings could be obtained even when the wingspan is less than 3 mm. The venation and corrugation could be significantly identified from the results. The deformation of the wing under pin loading could be seen clearly from the results as well. Calibration shows that the shape and deformation measurement accuracies are no lower than 0.01 mm. Laser triangulation and image matching were combined dexterously to adapt wings' complex shape, size, and transparency. It is suitable for insect flight research or flapping wing micro-air vehicle development.

  16. Wing patterning genes and coevolution of Müllerian mimicry in Heliconius butterflies: Support from phylogeography, cophylogeny, and divergence times.

    Science.gov (United States)

    Hoyal Cuthill, Jennifer F; Charleston, Michael

    2015-12-01

    Examples of long-term coevolution are rare among free-living organisms. Müllerian mimicry in Heliconius butterflies had been suggested as a key example of coevolution by early genetic studies. However, research over the last two decades has been dominated by the idea that the best-studied comimics, H. erato and H. melpomene, did not coevolve at all. Recently sequenced genes associated with wing color pattern phenotype offer a new opportunity to resolve this controversy. Here, we test the hypothesis of coevolution between H. erato and H. melpomene using Bayesian multilocus analysis of five color pattern genes and five neutral genetic markers. We first explore the extent of phylogenetic agreement versus conflict between the different genes. Coevolution is then tested against three aspects of the mimicry diversifications: phylogenetic branching patterns, divergence times, and, for the first time, phylogeographic histories. We show that all three lines of evidence are compatible with strict coevolution of the diverse mimicry wing patterns, contrary to some recent suggestions. Instead, these findings tally with a coevolutionary diversification driven primarily by the ecological force of Müllerian mimicry. © 2015 The Author(s). Evolution © 2015 The Society for the Study of Evolution.

  17. Leaf venation: structure, function, development, evolution, ecology and applications in the past, present and future.

    Science.gov (United States)

    Sack, Lawren; Scoffoni, Christine

    2013-06-01

    The design and function of leaf venation are important to plant performance, with key implications for the distribution and productivity of ecosystems, and applications in paleobiology, agriculture and technology. We synthesize classical concepts and the recent literature on a wide range of aspects of leaf venation. We describe 10 major structural features that contribute to multiple key functions, and scale up to leaf and plant performance. We describe the development and plasticity of leaf venation and its adaptation across environments globally, and a new global data compilation indicating trends relating vein length per unit area to climate, growth form and habitat worldwide. We synthesize the evolution of vein traits in the major plant lineages throughout paleohistory, highlighting the multiple origins of individual traits. We summarize the strikingly diverse current applications of leaf vein research in multiple fields of science and industry. A unified core understanding will enable an increasing range of plant biologists to incorporate leaf venation into their research. © 2013 The Authors New Phytologist © 2013 New Phytologist Trust.

  18. Independently Controlled Wing Stroke Patterns in the Fruit Fly Drosophila melanogaster

    Czech Academy of Sciences Publication Activity Database

    Chakraborty, Soma; Bartussek, Jan; Fry, S.N.; Zápotocký, Martin

    2015-01-01

    Roč. 10, č. 2 (2015), e0116813 E-ISSN 1932-6203 R&D Projects: GA ČR(CZ) GBP304/12/G069 Institutional support: RVO:67985823 Keywords : motor control * wing kinematics * independent component analysis Subject RIV: ED - Physiology Impact factor: 3.057, year: 2015

  19. Nonbreeding isolation and population-specific migration patterns among three populations of Golden-winged Warblers

    Science.gov (United States)

    Kramer, Gunnar R.; Streby, Henry M.; Peterson, Sean M.; Lehman, Justin A.; Buehler, David A.; Wood, Petra; McNeil, Darin J.; Larkin, Jeffrey L.; Andersen, David E.

    2017-01-01

    Golden-winged Warblers (Vermivora chrysoptera) are Nearctic–Neotropical migrants experiencing varied regional population trends not fully explained by breeding-grounds factors such as nest success. A lack of detailed information on the nonbreeding distributions, migration routes, or timing of migration among populations hampers our ability to identify population processes outside the breeding period. We used geolocators to track annual movements of 21 Golden-winged Warblers from 3 North American breeding locations experiencing varying population trends to investigate the potential for nonbreeding site factors to influence breeding populations. We used the template-fit method to estimate locations of individual warblers throughout the year. Geolocator-marked warblers exhibited significant isolation among populations during migration and the nonbreeding period. During the nonbreeding period, Golden-winged Warblers from Minnesota, USA (n = 12) occurred in Central America from southern Mexico to central Nicaragua; warblers from Tennessee, USA (n = 7) occurred along the border of northern Colombia and Venezuela; and warblers from Pennsylvania, USA (n = 2) occurred in north-central Venezuela. Warblers travelled at slower rates over more days in fall migration than spring migration. Fall migration routes at the Gulf of Mexico were population-specific, whereas spring routes were more varied and overlapped among populations. Golden-winged Warblers from Pennsylvania migrated 4,000 and 5,000 km yr−1 farther than Tennessee and Minnesota warblers, respectively, and spent almost twice as long migrating in the fall compared to Minnesota warblers. Our results reveal nearly complete temporal and geographic isolation among 3 populations of Golden-winged Warblers throughout the annual cycle, resulting in opportunities for population- and site-specific factors to differentially influence populations outside the breeding period. Our findings highlight the need for monitoring

  20. Quantitative neutron imaging of water distribution, venation network and sap flow in leaves.

    Science.gov (United States)

    Defraeye, Thijs; Derome, Dominique; Aregawi, Wondwosen; Cantré, Dennis; Hartmann, Stefan; Lehmann, Eberhard; Carmeliet, Jan; Voisard, Frédéric; Verboven, Pieter; Nicolai, Bart

    2014-08-01

    Quantitative neutron imaging is a promising technique to investigate leaf water flow and transpiration in real time and has perspectives towards studies of plant response to environmental conditions and plant water stress. The leaf hydraulic architecture is a key determinant of plant sap transport and plant-atmosphere exchange processes. Non-destructive imaging with neutrons shows large potential for unveiling the complex internal features of the venation network and the transport therein. However, it was only used for two-dimensional imaging without addressing flow dynamics and was still unsuccessful in accurate quantification of the amount of water. Quantitative neutron imaging was used to investigate, for the first time, the water distribution in veins and lamina, the three-dimensional venation architecture and sap flow dynamics in leaves. The latter was visualised using D2O as a contrast liquid. A high dynamic resolution was obtained by using cold neutrons and imaging relied on radiography (2D) as well as tomography (3D). The principle of the technique was shown for detached leaves, but can be applied to in vivo leaves as well. The venation network architecture and the water distribution in the veins and lamina unveiled clear differences between plant species. The leaf water content could be successfully quantified, though still included the contribution of the leaf dry matter. The flow measurements exposed the hierarchical structure of the water transport pathways, and an accurate quantification of the absolute amount of water uptake in the leaf was possible. Particular advantages of neutron imaging, as compared to X-ray imaging, were identified. Quantitative neutron imaging is a promising technique to investigate leaf water flow and transpiration in real time and has perspectives towards studies of plant response to environmental conditions and plant water stress.

  1. Aridity induces super-optimal investment in leaf venation by Eucalyptus and Corymbia

    Science.gov (United States)

    Drake, Paul L.; de Boer, Hugo J.; Price, Charles A.; Veneklaas, Erik J.

    2016-04-01

    The close relationship between leaf water status and stomatal conductance implies that the hydraulic architecture of leaves poses an important constraint on carbon uptake, specifically in arid environments with high evaporative demands. However, it remains uncertain how morphological, hydraulic and photosynthetic traits are coordinated to achieve optimal leaf functioning in arid environments. Zwieniecki and Boyce (2014) proposed a generic framework on the hydraulic architecture of leaves based on the argument that water is optimally distributed when the lateral distance between neighboring water transport veins (dx) is approximately equal to the distance from these veins to the epidermis (dy), expressed as dx:dy ≈1. Many derived angiosperms realize this optimal hydraulic architecture by closely coordinating leaf vein density with leaf thickness and the lateral position of veins inside the leaf. Zwieniecki and Boyce (2014) further suggested that over-investment in veins (dx:dy angiosperms adapted to temperate and moist (sub)tropical environments, we hypothesize that super-investment in leaf venation (resulting in dx:dyEvolution of a unique anatomical precision in angiosperm leaf venation lifts constraints on vascular plant ecology, Proc. R. Soc. B Biol. Sci., 281(1779), 2014.

  2. Drag Performance of Twist Morphing MAV Wing

    Directory of Open Access Journals (Sweden)

    Ismail N.I.

    2016-01-01

    Full Text Available Morphing wing is one of latest evolution found on MAV wing. However, due to few design problems such as limited MAV wing size and complicated morphing mechanism, the understanding of its aerodynamic behaviour was not fully explored. In fact, the basic drag distribution induced by a morphing MAV wing is still remained unknown. Thus, present work is carried out to compare the drag performance between a twist morphing wing with membrane and rigid MAV wing design. A quasi-static aeroelastic analysis by using the Ansys-Fluid Structure Interaction (FSI method is utilized in current works to predict the drag performance a twist morphing MAV wing design. Based on the drag pattern study, the results exhibits that the morphing wing has a partial similarities in overall drag pattern with the baseline (membrane and rigid wing. However, based CD analysis, it shows that TM wing induced higher CD magnitude (between 25% to 82% higher than to the baseline wing. In fact, TM wing also induced the largest CD increment (about 20% to 27% among the wings. The visualization on vortex structure revealed that TM wing also produce larger tip vortex structure (compared to baseline wings which presume to promote higher induce drag component and subsequently induce its higher CD performance.

  3. Experimental studies of vertical mixing patterns in open channel flow generated by two delta wings side-by-side

    Science.gov (United States)

    Vaughan, Garrett

    Open channel raceway bioreactors are a low-cost system used to grow algae for biofuel production. Microalgae have many promises when it comes to renewable energy applications, but many economic hurdles must be overcome to achieve an economic fuel source that is competitive with petroleum-based fuels. One way to make algae more competitive is to improve vertical mixing in algae raceway bioreactors. Previous studies show that mixing may be increased by the addition of mechanisms such as airfoils. The circulation created helps move the algae from the bottom to top surface for necessary photosynthetic exchange. This improvement in light utilization allowed a certain study to achieve 2.2-2.4 times the amount of biomass relative to bioreactors without airfoils. This idea of increasing mixing in open channel raceways has been the focus of the Utah State University (USU) raceway hydraulics group. Computational Fluid Dynamics (CFD), Acoustic Doppler Velocimetry (ADV), and Particle Image Velocimetry (PIV) are all methods used at USU to computationally and experimentally quantify mixing in an open channel raceway. They have also been used to observe the effects of using delta wings (DW) in increasing vertical mixing in the raceway. These efforts showed great potential in the DW in increasing vertical mixing in the open channel bioreactor. However, this research begged the question, does the DW help increase algae growth? Three algae growth experiments comparing growth in a raceway with and without DW were completed. These experiments were successful, yielding an average 27.1% increase in the biomass. The DW appears to be a promising method of increasing algae biomass production. The next important step was to quantify vertical mixing and understand flow patterns due to two DWs side-by-side. Raceway channels are wider as they increase in size; and arrays of DWs will need to be installed to achieve quality mixing throughout the bioreactor. Quality mixing was attained for

  4. Spatial reflection patterns of iridescent wings of male pierid butterflies : Curved scales reflect at a wider angle than flat scales

    NARCIS (Netherlands)

    Pirih, Primož; Wilts, Bodo D.; Stavenga, Doekele G.

    2011-01-01

    The males of many pierid butterflies have iridescent wings, which presumably function in intraspecific communication. The iridescence is due to nanostructured ridges of the cover scales. We have studied the iridescence in the males of a few members of Coliadinae, Gonepteryx aspasia, G. cleopatra, G.

  5. Effects of wing locations on wing rock induced by forebody vortices

    Directory of Open Access Journals (Sweden)

    Ma Baofeng

    2016-10-01

    Full Text Available Previous studies have shown that asymmetric vortex wakes over slender bodies exhibit a multi-vortex structure with an alternate arrangement along a body axis at high angle of attack. In this investigation, the effects of wing locations along a body axis on wing rock induced by forebody vortices was studied experimentally at a subcritical Reynolds number based on a body diameter. An artificial perturbation was added onto the nose tip to fix the orientations of forebody vortices. Particle image velocimetry was used to identify flow patterns of forebody vortices in static situations, and time histories of wing rock were obtained using a free-to-roll rig. The results show that the wing locations can affect significantly the motion patterns of wing rock owing to the variation of multi-vortex patterns of forebody vortices. As the wing locations make the forebody vortices a two-vortex pattern, the wing body exhibits regularly divergence and fixed-point motion with azimuthal variations of the tip perturbation. If a three-vortex pattern exists over the wing, however, the wing-rock patterns depend on the impact of the highest vortex and newborn vortex. As the three vortices together influence the wing flow, wing-rock patterns exhibit regularly fixed-points and limit-cycled oscillations. With the wing moving backwards, the newborn vortex becomes stronger, and wing-rock patterns become fixed-points, chaotic oscillations, and limit-cycled oscillations. With further backward movement of wings, the vortices are far away from the upper surface of wings, and the motions exhibit divergence, limit-cycled oscillations and fixed-points. For the rearmost location of the wing, the wing body exhibits stochastic oscillations and fixed-points.

  6. Uncorrelated evolution of leaf and petal venation patterns across the angiosperm phylogeny.

    Science.gov (United States)

    Roddy, Adam B; Guilliams, C Matt; Lilittham, Terapan; Farmer, Jessica; Wormser, Vanessa; Pham, Trang; Fine, Paul V A; Feild, Taylor S; Dawson, Todd E

    2013-10-01

    Early angiosperm evolution, beginning approximately 140 million years ago, saw many innovations that enabled flowering plants to alter ecosystems globally. These included the development of novel, flower-based pollinator attraction mechanisms and the development of increased water transport capacity in stems and leaves. Vein length per area (VLA) of leaves increased nearly threefold in the first 30-40 million years of angiosperm evolution, increasing the capacity for transpiration and photosynthesis. In contrast to leaves, high water transport capacities in flowers may not be an advantage because flowers do not typically contribute to plant carbon gain. Although flowers of extant basal angiosperms are hydrated by the xylem, flowers of more recently derived lineages may be hydrated predominantly by the phloem. In the present study, we measured leaf and flower VLA for a phylogenetically diverse sample of 132 species from 52 angiosperm families to ask (i) whether flowers have lower VLA than leaves, (ii) whether flowers of basal angiosperm lineages have higher VLA than more recently derived lineages because of differences between xylem and phloem hydration, and (iii) whether flower and leaf VLA evolved independently. It was found that floral structures had lower VLA than leaves, but basal angiosperm flowers did not have higher VLA than more derived lineages. Furthermore, the independent evolution of leaf and petal VLA suggested that these organs may be developmentally modular. Unlike leaves, which have experienced strong selection for increased water transport capacity, flowers may have been shielded from such selective pressures by different developmental processes controlling VLA throughout the plant bauplan.

  7. Improved non-destructive 2D and 3D X-ray imaging of leaf venation.

    Science.gov (United States)

    Schneider, Julio V; Rabenstein, Renate; Wesenberg, Jens; Wesche, Karsten; Zizka, Georg; Habersetzer, Jörg

    2018-01-01

    Leaf venation traits are important for many research fields such as systematics and evolutionary biology, plant physiology, climate change, and paleoecology. In spite of an increasing demand for vein trait data, studies are often still data-limited because the development of methods that allow rapid generation of large sets of vein data has lagged behind. Recently, non-destructive X-ray technology has proven useful as an alternative to traditional slow and destructive chemical-based methods. Non-destructive techniques more readily allow the use of herbarium specimens, which provide an invaluable but underexploited resource of vein data and related environmental information. The utility of 2D X-ray technology and microfocus X-ray computed tomography, however, has been compromised by insufficient image resolution. Here, we advanced X-ray technology by increasing image resolution and throughput without the application of contrast agents. For 2D contact microradiography, we developed a method which allowed us to achieve image resolutions of up to 7 µm, i.e. a 3.6-fold increase compared to the industrial standard (25 µm resolution). Vein tracing was further optimized with our image processing standards that were specifically adjusted for different types of leaf structure and the needs of higher imaging throughput. Based on a test dataset, in 91% of the samples the 7 µm approach led to a significant improvement in estimations of minor vein density compared to the industrial standard. Using microfocus X-ray computed tomography, very high-resolution images were obtained from a virtual 3D-2D transformation process, which was superior to that of 3D images. Our 2D X-ray method with a significantly improved resolution advances rapid non-destructive bulk scanning at a quality that in many cases is sufficient to determine key venation traits. Together with our high-resolution microfocus X-ray computed tomography method, both non-destructive approaches will help in vein

  8. Evolution of a unique anatomical precision in angiosperm leaf venation lifts constraints on vascular plant ecology.

    Science.gov (United States)

    Zwieniecki, Maciej A; Boyce, Charles K

    2014-03-22

    The main role of leaf venation is to supply water across the photosynthetic surface to keep stomata open and allow access to atmospheric CO2 despite evaporative demand. The optimal uniform delivery of water occurs when the distance between veins equals the depth of vein placement within the leaf away from the evaporative surface. As presented here, only angiosperms maintain this anatomical optimum across all leaf thicknesses and different habitats, including sheltered environments where this optimization need not be required. Intriguingly, basal angiosperm lineages tend to be underinvested hydraulically; uniformly high optimization is derived independently in the magnoliids, monocots and core eudicots. Gymnosperms and ferns, including available fossils, are limited by their inability to produce high vein densities. The common association of ferns with shaded humid environments may, in part, be a direct evolutionary consequence of their inability to produce hydraulically optimized leaves. Some gymnosperms do approach optimal vein placement, but only by virtue of their ability to produce thick leaves most appropriate in environments requiring water conservation. Thus, this simple anatomical metric presents an important perspective on the evolution and phylogenetic distribution of plant ecologies and further evidence that the vegetative biology of flowering plants-not just their reproductive biology-is unique.

  9. Wing geometry of Anopheles darlingi Root (Diptera: Culicidae) in five major Brazilian ecoregions.

    Science.gov (United States)

    Motoki, Maysa Tiemi; Suesdek, Lincoln; Bergo, Eduardo Sterlino; Sallum, Maria Anice Mureb

    2012-08-01

    We undertook geometric morphometric analysis of wing venation to assess this character's ability to distinguish Anopheles darlingi Root populations and to test the hypothesis that populations from coastal areas of the Brazilian Atlantic Forest differ from those of the interior Atlantic Forest, Cerrado, and the regions South and North of the Amazon River. Results suggest that populations from the coastal and interior Atlantic Forest are more similar to each other than to any of the other regional populations. Notably, the Cerrado population was more similar to that from north of the Amazon River than to that collected of south of the River, thus showing no correlation with geographical distances. We hypothesize that environmental and ecological factors may affect wing evolution in An. darlingi. Although it is premature to associate environmental and ecological determinants with wing features and evolution of the species, investigations on this field are promising. Copyright © 2012 Elsevier B.V. All rights reserved.

  10. Apparent Overinvestment in Leaf Venation Relaxes Leaf Morphological Constraints on Photosynthesis in Arid Habitats1[OPEN

    Science.gov (United States)

    de Boer, Hugo J.; Drake, Paul L.; Wendt, Erin; Price, Charles A.; Schulze, Ernst-Detlef; Turner, Neil C.; Nicolle, Dean

    2016-01-01

    Leaf veins supply the mesophyll with water that evaporates when stomata are open to allow CO2 uptake for photosynthesis. Theoretical analyses suggest that water is optimally distributed in the mesophyll when the lateral distance between veins (dx) is equal to the distance from these veins to the epidermis (dy), expressed as dx:dy ≈ 1. Although this theory is supported by observations of many derived angiosperms, we hypothesize that plants in arid environments may reduce dx:dy below unity owing to climate-specific functional adaptations of increased leaf thickness and increased vein density. To test our hypothesis, we assembled leaf hydraulic, morphological, and photosynthetic traits of 68 species from the Eucalyptus and Corymbia genera (termed eucalypts) along an aridity gradient in southwestern Australia. We inferred the potential gas-exchange advantage of reducing dx beyond dy using a model that links leaf morphology and hydraulics to photosynthesis. Our observations reveal that eucalypts in arid environments have thick amphistomatous leaves with high vein densities, resulting in dx:dy ratios that range from 1.6 to 0.15 along the aridity gradient. Our model suggests that, as leaves become thicker, the effect of reducing dx beyond dy is to offset the reduction in leaf gas exchange that would result from maintaining dx:dy at unity. This apparent overinvestment in leaf venation may be explained from the selective pressure of aridity, under which traits associated with long leaf life span, high hydraulic and thermal capacitances, and high potential rates of leaf water transport confer a competitive advantage. PMID:27784769

  11. Apparent over-investment in leaf venation relaxes leaf morphological constraints on photosynthesis in arid habitats

    Science.gov (United States)

    de Boer, Hugo; Drake, Paul; Veneklaas, Erik

    2017-04-01

    The close relationship between leaf water status and stomatal conductance implies that the hydraulic architecture of leaves poses an important constraint on transpiration, specifically in arid environments with high evaporative demands. However, it remains uncertain how morphological, hydraulic and photosynthetic traits are coordinated to achieve optimal leaf functioning in arid environments. Critical is that leaf veins supply the mesophyll with water that evaporates when stomata are open to allow CO2 uptake for photosynthesis. Theoretical analyses suggest that water is optimally distributed in the mesophyll when the lateral distance between veins (dx) is equal to the distance from these veins to the epidermis (dy), expressed as dx:dy≈1. Although this theory is supported by observations on many derived angiosperms, we hypothesize that plants in arid environments may reduce dx:dy below unity owing to climate-specific functional adaptations of increased leaf thickness and increased vein density. To test our hypothesis we assembled leaf hydraulic, morphological and photosynthetic traits of 68 species from the Eucalyptus and Corymbia genera (termed eucalypts) along an aridity gradient in southwestern Australia. We inferred the potential gas exchange advantage of reducing dx beyond dy using a model that links leaf morphology and hydraulics to photosynthesis. Our observations reveal that eucalypts in arid environments have thick amphistomatous leaves with high vein densities, resulting in dx:dy ratios that range from 1.6 to 0.15 along the aridity gradient. Our model suggests that as leaves become thicker, the effect of reducing dx beyond dy is to offset the reduction in leaf gas exchange that would result from maintaining dx:dy at unity. This apparent over-investment in leaf venation may be explained from the selective pressure of aridity, under which traits associated with long leaf lifespan, high hydraulic and thermal capacitances, and high potential rates of leaf

  12. Veins improve fracture toughness of insect wings.

    Directory of Open Access Journals (Sweden)

    Jan-Henning Dirks

    Full Text Available During the lifetime of a flying insect, its wings are subjected to mechanical forces and deformations for millions of cycles. Defects in the micrometre thin membranes or veins may reduce the insect's flight performance. How do insects prevent crack related material failure in their wings and what role does the characteristic vein pattern play? Fracture toughness is a parameter, which characterises a material's resistance to crack propagation. Our results show that, compared to other body parts, the hind wing membrane of the migratory locust S. gregaria itself is not exceptionally tough (1.04±0.25 MPa√m. However, the cross veins increase the wing's toughness by 50% by acting as barriers to crack propagation. Using fracture mechanics, we show that the morphological spacing of most wing veins matches the critical crack length of the material (1132 µm. This finding directly demonstrates how the biomechanical properties and the morphology of locust wings are functionally correlated in locusts, providing a mechanically 'optimal' solution with high toughness and low weight. The vein pattern found in insect wings thus might inspire the design of more durable and lightweight artificial 'venous' wings for micro-air-vehicles. Using the vein spacing as indicator, our approach might also provide a basis to estimate the wing properties of endangered or extinct insect species.

  13. FijiWings: an open source toolkit for semiautomated morphometric analysis of insect wings.

    Science.gov (United States)

    Dobens, Alexander C; Dobens, Leonard L

    2013-08-07

    Development requires coordination between cell proliferation and cell growth to pattern the proper size of tissues, organs, and whole organisms. The Drosophila wing has landmark features, such as the location of veins patterned by cell groups and trichome structures produced by individual cells, that are useful to examine the genetic contributions to both tissue and cell size. Wing size and trichome density have been measured manually, which is tedious and error prone, and although image processing and pattern-recognition software can quantify features in micrographs, this approach has not been applied to insect wings. Here we present FijiWings, a set of macros designed to perform semiautomated morphophometric analysis of a wing photomicrograph. FijiWings uses plug-ins installed in the Fiji version of ImageJ to detect and count trichomes and measure wing area either to calculate trichome density of a defined region selected by the user or generate a heat map of overall trichome densities. For high-throughput screens we have developed a macro that directs a trainable segmentation plug-in to detect wing vein locations either to measure trichome density in specific intervein regions or produce a heat map of relative intervein areas. We use wing GAL4 drivers and UAS-regulated transgenes to confirm the ability of these tools to detect changes in overall tissue growth and individual cell size. FijiWings is freely available and will be of interest to a broad community of fly geneticists studying both the effect of gene function on wing patterning and the evolution of wing morphology.

  14. Veins Improve Fracture Toughness of Insect Wings

    Science.gov (United States)

    Dirks, Jan-Henning; Taylor, David

    2012-01-01

    During the lifetime of a flying insect, its wings are subjected to mechanical forces and deformations for millions of cycles. Defects in the micrometre thin membranes or veins may reduce the insect’s flight performance. How do insects prevent crack related material failure in their wings and what role does the characteristic vein pattern play? Fracture toughness is a parameter, which characterises a material’s resistance to crack propagation. Our results show that, compared to other body parts, the hind wing membrane of the migratory locust S. gregaria itself is not exceptionally tough (1.04±0.25 MPa√m). However, the cross veins increase the wing’s toughness by 50% by acting as barriers to crack propagation. Using fracture mechanics, we show that the morphological spacing of most wing veins matches the critical crack length of the material (1132 µm). This finding directly demonstrates how the biomechanical properties and the morphology of locust wings are functionally correlated in locusts, providing a mechanically ‘optimal’ solution with high toughness and low weight. The vein pattern found in insect wings thus might inspire the design of more durable and lightweight artificial ‘venous’ wings for micro-air-vehicles. Using the vein spacing as indicator, our approach might also provide a basis to estimate the wing properties of endangered or extinct insect species. PMID:22927966

  15. Geometric morphometrics analysis of the hind wing of leaf beetles: proximal and distal parts are separate modules

    Directory of Open Access Journals (Sweden)

    Jing Ren

    2017-07-01

    Full Text Available The success of beetles is mainly attributed to the possibility to hide the hindwings under the sclerotised elytra. The acquisition of the transverse folding function of the hind wing is an important event in the evolutionary history of beetles. In this study, the morphological and functional variances in the hind wings of 94 leaf beetle species (Coleoptera: Chrysomelinae is explored using geometric morphometrics based on 36 landmarks. Principal component analysis and Canonical variate analysis indicate that changes of apical area, anal area, and middle area are three useful phylogenetic features at a subtribe level of leaf beetles. Variances of the apical area are the most obvious, which strongly influence the entire venation variance. Partial least squares analysis indicates that the proximal and distal parts of hind wings are weakly associated. Modularity tests confirm that the proximal and distal compartments of hind wings are separate modules. It is deduced that for leaf beetles, or even other beetles, the hind wing possibly exhibits significant functional divergences that occurred during the evolution of transverse folding that resulted in the proximal and distal compartments of hind wings evolving into separate functional modules.

  16. A key, based on wing patterns of biting midges (genus Culicoides Latreille - Diptera: Ceratopogonidae in the Iberian Peninsula, for use in epidemiological studies

    Directory of Open Access Journals (Sweden)

    Rawlings, Peter

    1996-12-01

    Full Text Available The identity of vectors of disease are often required speedily in epidemiological studies but with a precision which excludes as many other species as possible. Identification keys usually require the examination of many different parts of the suspected vector to pinpoint the species. This consumes considerable time and resources, so epidemiologists tend to ignore them. Asimplified approach to identification is proposed, using the characteristics of a single part of the body (the wings of biting midges of the genus Culicoides. The level of differentiation was epidemiologically valuable. The monoclave could not differentiate all the species from each other but more than one third (20/58 of identifications were for single species, and a further 12/58 identifications gave only two possibilities, making 55.2% of identifications to an accuracy of at most one of two species. The diagnosis of vector species was reached in a maximum of six decision points. The only notable exception to valuable differentiation was the four species in the Culicoides obsoletus group which had almost identical female wing patterns. The ready availability of simple keys, which can be used by anyone without formal training in taxonomy, for all the species of a group in a region should encourage greater standardisation of identifications in all studies, including those not primarily aimed at systematics. These monoclaves can also serve as the primary tools to build computerised image-recognition systems for genera, families and orders of insects.Con frequencia en los estudios epidemiológicos hace falta conocer con rapidez, pero también con precisión, la identidad de los vectores. Por lo general los procedimientos de identificación y las claves exigen el examen de un elevado número de partes diferentes del vector sospechoso. Este enfoque consume mucho tiempo y recursos por lo que tiende a ser evitado por los epidemiólogos. Se propone un sistema simplificado para la

  17. Shapes of leaves with parallel venation. Modelling of the Epipactis sp. (Orchidaceae) leaves with the help of a system of coupled elastic beams

    OpenAIRE

    Jakubska-Busse, Anna; Janowicz, Maciej; Ochnio, Luiza; Jackowska-Zduniak, Beata

    2016-01-01

    Static properties of leaves with parallel venation, with particular emphasis on the genus EpipactisZinn, 1757 (Orchidaceae, Neottieae) have been modelled with coupled quasi-parallel elastic “beams.” The non-linear theory of strongly bended beams have been employed. The resulting boundary-value problem has been solved numerically with the help of the finite-difference method. Possible dislocations resulting in additional Dirac-delta like forces have been take into account. Morphological simila...

  18. The influence of pigmentation patterning on bumblebee foraging from flowers of Antirrhinum majus.

    Science.gov (United States)

    Whitney, Heather M; Milne, Georgina; Rands, Sean A; Vignolini, Silvia; Martin, Cathie; Glover, Beverley J

    2013-03-01

    Patterns of pigmentation overlying the petal vasculature are common in flowering plants and have been postulated to play a role in pollinator attraction. Previous studies report that such venation patterning is significantly more attractive to bee foragers in the field than ivory or white flowers without veins. To dissect the ways in which venation patterning of pigment can influence bumblebee behaviour, we investigated the response of flower-naïve individuals of Bombus terrestris to veined, ivory and red near-isogenic lines of Antirrhinum majus. We find that red venation shifts flower colour slightly, although the ivory background is the dominant colour. Bees were readily able to discriminate between ivory and veined flowers under differential conditioning but showed no innate preference when presented with a free choice of rewarding ivory and veined flowers. In contrast, both ivory and veined flowers were selected significantly more often than were red flowers. We conclude that advantages conferred by venation patterning might stem from bees learning of their use as nectar guides, rather than from any innate preference for striped flowers.

  19. A new genus of Rhysipoline wasp (Hymenoptera: Braconidae) with modified wing venation from Africa and Papua New Guinea, parasitoid on Choreutidae (Lepidoptera)

    Czech Academy of Sciences Publication Activity Database

    Quicke, D. L. J.; Belokobylskij, S. A.; Smith, M. A.; Rota, J.; Hrček, Jan; Butcher, B. A.

    2016-01-01

    Roč. 66, č. 2 (2016), s. 173-192 ISSN 0003-4541 R&D Projects: GA ČR GD206/08/H044; GA ČR GA206/09/0115; GA ČR GAP505/10/0673; GA MŠk LC06073; GA MŠk ME09082 Grant - others:GA JU(CZ) 136/2010/P Institutional support: RVO:60077344 Keywords : cyclostomes * barcording * Clinocentrus Subject RIV: EG - Zoology Impact factor: 0.699, year: 2016 http://www.bioone.org/doi/abs/10.3161/00034541ANZ2016.66.2.003

  20. Leaf venation, as a resistor, to optimize a switchable IR absorber.

    Science.gov (United States)

    Alston, M E; Barber, R

    2016-08-24

    Leaf vascular patterns are the mechanisms and mechanical support for the transportation of fluidics for photosynthesis and leaf development properties. Vascular hierarchical networks in leaves have far-reaching functions in optimal transport efficiency of functional fluidics. Embedding leaf morphogenesis as a resistor network is significant in the optimization of a translucent thermally functional material. This will enable regulation through pressure equalization by diminishing flow pressure variation. This paper investigates nature's vasculature networks that exhibit hierarchical branching scaling applied to microfluidics. To enable optimum potential for pressure drop regulation by algorithm design. This code analysis of circuit conduit optimization for transport fluidic flow resistance is validated against CFD simulation, within a closed loop network. The paper will propose this self-optimization, characterization by resistance seeking targeting to determine a microfluidic network as a resistor. To advance a thermally function material as a switchable IR absorber.

  1. Shapes of leaves with parallel venation. Modelling of the Epipactis sp. (Orchidaceae leaves with the help of a system of coupled elastic beams

    Directory of Open Access Journals (Sweden)

    Anna Jakubska-Busse

    2016-06-01

    Full Text Available Static properties of leaves with parallel venation, with particular emphasis on the genus EpipactisZinn, 1757 (Orchidaceae, Neottieae have been modelled with coupled quasi-parallel elastic “beams.” The non-linear theory of strongly bended beams have been employed. The resulting boundary-value problem has been solved numerically with the help of the finite-difference method. Possible dislocations resulting in additional Dirac-delta like forces have been take into account. Morphological similarity of the model and real leaves has been obtained. In particular, the concentrated forces have been shown to cause undulation in the leaves.

  2. Flow structure of vortex-wing interaction

    Science.gov (United States)

    McKenna, Christopher K.

    Impingement of a streamwise-oriented vortex upon a fin, tail, blade or wing represents a fundamental class of flow-structure interaction that extends across a range of applications. This interaction can give rise to time-averaged loading, as well as unsteady loading known as buffeting. The loading is sensitive to parameters of the incident vortex as well as the location of vortex impingement on the downstream aerodynamic surface, generically designated as a wing. Particle image velocimetry is employed to determine patterns of velocity, vorticity, swirl ratio, and streamlines on successive cross-flow planes upstream of and along the wing, which lead to volume representations and thereby characterization of the interaction. At locations upstream of the leading edge of the wing, the evolution of the incident vortex is affected by the presence of the wing, and is highly dependent on the spanwise location of vortex impingement. Even at spanwise locations of impingement well outboard of the wing tip, a substantial influence on the structure of the incident vortex at locations significantly upstream of the leading edge of the wing was observed. For spanwise locations close to or intersecting the vortex core, the effects of upstream influence of the wing on the vortex are to: decrease the swirl ratio; increase the streamwise velocity deficit; decrease the streamwise vorticity; increase the azimuthal vorticity; increase the upwash; decrease the downwash; and increase the root-mean-square fluctuations of both streamwise velocity and vorticity. The interrelationship between these effects is addressed, including the rapid attenuation of axial vorticity in presence of an enhanced defect of axial velocity in the central region of the vortex. Moreover, when the incident vortex is aligned with, or inboard of, the tip of the wing, the swirl ratio decreases to values associated with instability of the vortex, giving rise to enhanced values of azimuthal vorticity relative to the

  3. The role of wing kinematics of freely flying birds downstream the wake of flapping wings

    Science.gov (United States)

    Krishnan, Krishnamoorthy; Gurka, Roi

    2016-11-01

    Avian aerodynamics has been a topic of research for centuries. Avian flight features such as flapping, morphing and maneuvering make bird aerodynamics a complex system to study, analyze and understand. Aerodynamic performance of the flapping wings can be quantified by measuring the vortex structures present in the downstream wake. Still, the direct correlation between the flapping wing kinematics and the evolution of wake features need to be established. In this present study, near wake of three bird species (western sandpiper, European starling and American robin) have been measured experimentally. Long duration, time-resolved, particle image velocimetry technique has been used to capture the wake properties. Simultaneously, the bird kinematics have been captured using high speed camera. Wake structures are reconstructed from the collected PIV images for long chord distances downstream. Wake vorticities and circulation are expressed in the wake composites. Comparison of the wake features of the three birds shows similarities and some key differences are also found. Wing tip motions of the birds are extracted for four continuous wing beat cycle to analyze the wing kinematics. Kinematic parameters of all the three birds are compared to each other and similar trends exhibited by all the birds have been observed. A correlation between the wake evolutions with the wing motion is presented. It was found that the wings' motion generates unique flow patterns at the near wake, especially at the transition phases. At these locations, a drastic change in the circulation was observed.

  4. Effects of Canard on the Flowfield over a Wing

    Science.gov (United States)

    Nayebzadeh, Arash

    2015-11-01

    Surface and flowfield pressure measurements have been done over delta wing/canard configuration in a variety of canard vertical and horizontal locations and angles of attack. The experimental model consisted of wing, canard and a body to accommodate pressure tubing and canard rotation mechanism. All the tests have been performed at subsonic velocities and the effect of canard were analyzed through comparison between surface and flowfield pressure distributions. It was found that vortex flow pattern over the wing is dominated mainly by canard vertical position and in some cases, by merging of canard and wing vortices. In addition, the pressure loss induced by canard vortex on the wing surface moves the wing vortex toward the leading edge. In the mid canard configuration, canard and wing vortices merge at x/c greater than 0.5 and as a result of this phenomenon, abrupt pressure loss induces more stable vortex flow over the wing. It is also shown that canard plays a vital role in vortex break down over the wing.

  5. Folding wings like a cockroach: a review of transverse wing folding ensign wasps (Hymenoptera: Evaniidae: Afrevania and Trissevania.

    Directory of Open Access Journals (Sweden)

    István Mikó

    Full Text Available We revise two relatively rare ensign wasp genera, whose species are restricted to Sub-Saharan Africa: Afrevania and Trissevania. Afrevania longipetiolata sp. nov., Trissevania heatherae sp. nov., T. hugoi sp. nov., T. mrimaensis sp. nov. and T. slideri sp. nov. are described, males and females of T. anemotis and Afrevania leroyi are redescribed, and an identification key for Trissevaniini is provided. We argue that Trissevania mrimaensis sp. nov. and T. heatherae sp. nov. populations are vulnerable, given their limited distributions and threats from mining activities in Kenya. We hypothesize that these taxa together comprise a monophyletic lineage, Trissevaniini, tr. nov., the members of which share the ability to fold their fore wings along two intersecting fold lines. Although wing folding of this type has been described for the hind wing of some insects four-plane wing folding of the fore wing has never been documented. The wing folding mechanism and the pattern of wing folds of Trissevaniini is shared only with some cockroach species (Blattodea. It is an interesting coincidence that all evaniids are predators of cockroach eggs. The major wing fold lines of Trissevaniini likely are not homologous to any known longitudinal anatomical structures on the wings of other Evaniidae. Members of the new tribe share the presence of a coupling mechanism between the fore wing and the mesosoma that is composed of a setal patch on the mesosoma and the retinaculum of the fore wing. While the setal patch is an evolutionary novelty, the retinaculum, which originally evolved to facilitate fore and hind wing coupling in Hymenoptera, exemplifies morphological exaptation. We also refine and clarify the Semantic Phenotype approach used in previous taxonomic revisions and explore the consequences of merging new with existing data. The way that semantic statements are formulated can evolve in parallel, alongside improvements to the ontologies themselves.

  6. Temporal pattern of feeding activity in the firebug Pyrrhocoris apterus and its relation to sex, wing dimorphism and physiological state of adults

    Czech Academy of Sciences Publication Activity Database

    Socha, Radomír; Zemek, Rostislav

    2007-01-01

    Roč. 32, č. 1 (2007), s. 16-25 ISSN 0307-6962 R&D Projects: GA ČR GA206/03/0016 Institutional research plan: CEZ:AV0Z50070508 Keywords : feeding pattern * females * males Subject RIV: ED - Physiology Impact factor: 1.410, year: 2007

  7. Patriline Differences Reveal Genetic Influence on Forewing Size and Shape in a Yellowjacket Wasp (Hymenoptera: Vespidae: Vespula flavopilosa Jacobson, 1978.

    Directory of Open Access Journals (Sweden)

    Adrien Perrard

    Full Text Available The wing venation is frequently used as a morphological marker to distinguish biological groups among insects. With geometric morphometrics, minute shape differences can be detected between closely related species or populations, making this technique useful for taxonomy. However, the direct influence of genetic differences on wing morphology has not been explored within colonies of social insects. Here, we show that the father's genotype has a direct effect on wing morphology in colonies of social wasps. Using geometric morphometrics on the venation pattern, we found significant differences in wing size and shape between patrilines of yellowjackets, taking allometry and measurement error into account. The genetic influence on wing size accounted for a small part of the overall size variation, but venation shape was highly structured by the differences between patrilines. Overall, our results showed a strong genetic influence on wing morphology likely acting at multiple levels of venation pattern development. This confirmed the pertinence of this marker for taxonomic purposes and suggests this phenotype as a potentially useful marker for phylogenies. This also raises doubts about the strength of selective pressures on this phenotype, which highlights the need to understand better the role of wing venation shape in insect flight.

  8. WINGS Data Release

    DEFF Research Database (Denmark)

    Moretti, A.; Poggianti, B. M.; Fasano, G.

    2014-01-01

    Context. To effectively investigate galaxy formation and evolution, it is of paramount importance to exploit homogeneous data for large samples of galaxies in different environments. Aims. The WIde-field Nearby Galaxy-cluster Survey (WINGS) project aim is to evaluate physical properties of galaxies...... in a complete sample of low redshift clusters to be used as reference sample for evolutionary studies. The WINGS survey is still ongoing and the original dataset will be enlarged with new observations. This paper presents the entire collection of WINGS measurements obtained so far. Methods. We decided to make......, and on the cluster redshift, reaching on average 90% at V ≲ 21.7. Near-infrared photometric catalogs for 26 (in K) and 19 (in J) clusters are part of the database and the number of sources is 962 344 in K and 628 813 in J. Here again the completeness depends on the data quality, but it is on average higher than 90...

  9. Temporal variation of wing geometry in Aedes albopictus

    Directory of Open Access Journals (Sweden)

    Paloma Oliveira Vidal

    2012-12-01

    Full Text Available Although native to the tropical and subtropical areas of Southeast Asia, Aedes albopictus is now found on five continents, primarily due to its great capacity to adapt to different environments. This species is considered a secondary vector of dengue virus in several countries. Wing geometric morphometrics is widely used to furnish morphological markers for the characterisation and identification of species of medical importance and for the assessment of population dynamics. In this work, we investigated the metric differentiation of the wings of Ae. albopictus samples collected over a four-year period (2007-2010 in São Paulo, Brazil. Wing size significantly decreased during this period for both sexes and the wing shape also changed over time, with the wing shapes of males showing greater differences after 2008 and those of females differing more after 2009. Given that the wings play sex-specific roles, these findings suggest that the males and females could be affected by differential evolutionary pressures. Consistent with this hypothesis, a sexually dimorphic pattern was detected and quantified: the females were larger than the males (with respect to the mean and had a distinct wing shape, regardless of allometric effects. In conclusion, wing alterations, particularly those involving shape, are a sensitive indicator of microevolutionary processes in this species.

  10. Temporal variation of wing geometry in Aedes albopictus.

    Science.gov (United States)

    Vidal, Paloma Oliveira; Carvalho, Eneas; Suesdek, Lincoln

    2012-12-01

    Although native to the tropical and subtropical areas of Southeast Asia, Aedes albopictus is now found on five continents, primarily due to its great capacity to adapt to different environments. This species is considered a secondary vector of dengue virus in several countries. Wing geometric morphometrics is widely used to furnish morphological markers for the characterisation and identification of species of medical importance and for the assessment of population dynamics. In this work, we investigated the metric differentiation of the wings of Ae. albopictus samples collected over a four-year period (2007-2010) in São Paulo, Brazil. Wing size significantly decreased during this period for both sexes and the wing shape also changed over time, with the wing shapes of males showing greater differences after 2008 and those of females differing more after 2009. Given that the wings play sex-specific roles, these findings suggest that the males and females could be affected by differential evolutionary pressures. Consistent with this hypothesis, a sexually dimorphic pattern was detected and quantified: the females were larger than the males (with respect to the mean) and had a distinct wing shape, regardless of allometric effects. In conclusion, wing alterations, particularly those involving shape, are a sensitive indicator of microevolutionary processes in this species.

  11. Wing shape allometry and aerodynamics in calopterygid damselflies: a comparative approach.

    Science.gov (United States)

    Outomuro, David; Adams, Dean C; Johansson, Frank

    2013-06-07

    Wing size and shape have important aerodynamic implications on flight performance. We explored how wing size was related to wing shape in territorial males of 37 taxa of the damselfly family Calopterygidae. Wing coloration was also included in the analyses because it is sexually and naturally selected and has been shown to be related to wing shape. We studied wing shape using both the non-dimensional radius of the second moment of wing area (RSM) and geometric morphometrics. Lower values of the RSM result in less energetically demanding flight and wider ranges of flight speed. We also re-analyzed previously published data on other damselflies and dragonflies. The RSM showed a hump-shaped relationship with wing size. However, after correcting for phylogeny using independent contrast, this pattern changed to a negative linear relationship. The basal genus of the study family, Hetaerina, was mainly driving that change. The obtained patterns were specific for the study family and differed from other damselflies and dragonflies. The relationship between the RSM and wing shape measured by geometric morphometrics was linear, but relatively small changes along the RSM axis can result in large changes in wing shape. Our results also showed that wing coloration may have some effect on RSM. We found that RSM showed a complex relationship with size in calopterygid damselflies, probably as a result of other selection pressures besides wing size per se. Wing coloration and specific behavior (e.g. courtship) are potential candidates for explaining the complexity. Univariate measures of wing shape such as RSM are more intuitive but lack the high resolution of other multivariate techniques such as geometric morphometrics. We suggest that the relationship between wing shape and size are taxa-specific and differ among closely-related insect groups.

  12. WHEN COMPASSION GROWS WINGS

    African Journals Online (AJOL)

    Nicky

    antiretroviral roll-out in full swing, the. WHEN COMPASSION GROWS WINGS. The free time and expertise given by its deeply committed core of professional volunteers. (including pilots) is the lifeblood of the operation. Red Cross Air Mercy Service volunteer, German national Dr Florian Funk, at the AMS Durban base.

  13. Twisted Winged Endoparasitoids

    Indian Academy of Sciences (India)

    Home; Journals; Resonance – Journal of Science Education; Volume 9; Issue 10. Twisted Winged Endoparasitoids - An Enigma for Entomologists. Alpana Mazumdar. General Article Volume 9 Issue 10 October 2004 pp 19-24. Fulltext. Click here to view fulltext PDF. Permanent link:

  14. Logistics Supply of the Distributed Air Wing

    Science.gov (United States)

    2014-09-01

    Event Graph The Consumption Process first instantiates the variables . The model follows a conveyor belt pattern, whereby after processing an event...to any part of the world. A capstone project, conducted by the system engineering curriculum, proposed to distribute the air assets from the aircraft...SUBJECT TERMS distributed air wing, logistics, supply, unmanned air systems , cargo UAS, unmanned systems , discrete event simulation, vehicle routing

  15. Design optimization of deployable wings

    Science.gov (United States)

    Gaddam, Pradeep

    Morphing technology is an important aspect of UAV design, particularly in regards to deployable systems. The design of such system has an important impact on the vehicle's performance. The primary focus of the present research work was to determine the most optimum deployable wing design from 3 competing designs and develop one of the deployable wing designs to test in the research facility. A Matlab code was developed to optimize 3 deployable wing concepts inflatable, inflatable telescopic and rigid-folding wings based on a sequential optimization strategy. The constraints that were part of the code include the packaging constraints during its stowed state, fixed length of the deployed section and the minimum L/D constraint. This code resulted in determining the optimum weight of all the 3 designs, the most optimum weight design is the inflatable wing design. This is a result of the flexible skin material and also due to no rigid parts in the deployed wing section. Another goal of the research involved developing an inflatable telescopic wing. The prototype was tested in a wind tunnel, while the actual wing was tested in the altitude chamber to determine the deployment speed, input pressure, analyze and predict the deployment sequence and behavior of the wing at such high wind speeds and altitudes ranging from 60,000 ft to 90,000 ft. Results from these tests allowed us to conclude the deployment sequence of the telescopic wing followed from the root to the tip section. The results were used to analyze the deployment time of the wing. As expected the deployment time decreased with an increase in input pressure. The results also show us that as the altitude increases, the deployment speed of the wing also increased. This was demonstrated when the wing was tested at a maximum altitude pressure of 90,000ft, well above the design altitude of 60,000ft.

  16. Low Aspect-Ratio Wings for Wing-Ships

    DEFF Research Database (Denmark)

    Filippone, Antonino; Selig, M.

    1998-01-01

    Flying on ground poses technical and aerodynamical challenges. The requirements for compactness, efficiency, manouverability, off-design operation,open new areas of investigations in the fieldof aerodynamic analysis and design. A review ofthe characteristics of low-aspect ratio wings, in- and out...... of ground, is presented. It is shownthat the performance of such wings is generally inferior to that of slender wings, although in ground placement can yield substantial improvements in the aerodynamic efficiency....

  17. Wing sexual dimorphism of pathogen-vector culicids.

    Science.gov (United States)

    Virginio, Flávia; Oliveira Vidal, Paloma; Suesdek, Lincoln

    2015-03-14

    Sexual dimorphism in animals has been studied from different perspectives for decades. In 1874 Darwin hypothesized that it was related to sexual selection, and even after nearly 140 years, when additional empirical data has become available and the subject has been investigated from a contemporary viewpoint, this idea is still supported. Although mosquito (Culicidae) wings are of great importance as they play a sex-specific role, little is known about wing sexual dimorphism in these pathogen-vector insects. Detection and characterization of wing sexual dimorphism in culicids may indirectly enhance our knowledge of their epidemiology or reveal sex-linked genes, aspects that have been discussed by vector control initiatives and developers of genetically modified mosquitoes. Using geometric morphometrics, we carried out a comparative assessment of wing sexual dimorphism in ten culicid species of medical/veterinary importance from genera Culex, Aedes, Anopheles and Ochlerotatus collected in Brazil. Discriminant analysis revealed significant sexual dimorphism in all the species studied, indicating that phenotypic expression of wing shape in mosquitoes is indeed sex-specific. A cross-validated test performed to reclassify the sexes with and without allometry yielded very similar results. Mahalanobis distances among the ten species showed that the species had different patterns of shape sexual dimorphism and that females are larger than males in some species. Wing morphology differed significantly between species. The finding of sexual dimorphism in all the species would suggest that the wing geometry of Culicidae is canalized. Although sexual dimorphism is prevalent, species-specific patterns occur. Allometry was not the main determinant of sexual dimorphism, which suggests that sexual selection or other evolutionary mechanisms underlie wing sexual dimorphism in these insects.

  18. Wing defects in Drosophila xenicid mutant clones are caused by C-terminal deletion of additional sex combs (Asx).

    Science.gov (United States)

    Bischoff, Kara; Ballew, Anna C; Simon, Michael A; O'Reilly, Alana M

    2009-12-01

    The coordinated action of genes that control patterning, cell fate determination, cell size, and cell adhesion is required for proper wing formation in Drosophila. Defects in any of these basic processes can lead to wing aberrations, including blisters. The xenicid mutation was originally identified in a screen designed to uncover regulators of adhesion between wing surfaces [1]. Here, we demonstrate that expression of the betaPS integrin or the patterning protein Engrailed are not affected in developing wing imaginal discs in xenicid mutants. Instead, expression of the homeotic protein Ultrabithorax (Ubx) is strongly increased in xenicid mutant cells. Our results suggest that upregulation of Ubx transforms cells from a wing blade fate to a haltere fate, and that the presence of haltere cells within the wing blade is the primary defect leading to the adult wing phenotypes observed.

  19. Wing defects in Drosophila xenicid mutant clones are caused by C-terminal deletion of additional sex combs (Asx.

    Directory of Open Access Journals (Sweden)

    Kara Bischoff

    2009-12-01

    Full Text Available The coordinated action of genes that control patterning, cell fate determination, cell size, and cell adhesion is required for proper wing formation in Drosophila. Defects in any of these basic processes can lead to wing aberrations, including blisters. The xenicid mutation was originally identified in a screen designed to uncover regulators of adhesion between wing surfaces [1].Here, we demonstrate that expression of the betaPS integrin or the patterning protein Engrailed are not affected in developing wing imaginal discs in xenicid mutants. Instead, expression of the homeotic protein Ultrabithorax (Ubx is strongly increased in xenicid mutant cells.Our results suggest that upregulation of Ubx transforms cells from a wing blade fate to a haltere fate, and that the presence of haltere cells within the wing blade is the primary defect leading to the adult wing phenotypes observed.

  20. Real-time in vivo imaging of butterfly wing development: revealing the cellular dynamics of the pupal wing tissue.

    Directory of Open Access Journals (Sweden)

    Masaki Iwata

    Full Text Available Butterfly wings are covered with regularly arranged single-colored scales that are formed at the pupal stage. Understanding pupal wing development is therefore crucial to understand wing color pattern formation. Here, we successfully employed real-time in vivo imaging techniques to observe pupal hindwing development over time in the blue pansy butterfly, Junonia orithya. A transparent sheet of epithelial cells that were not yet regularly arranged was observed immediately after pupation. Bright-field imaging and autofluorescent imaging revealed free-moving hemocytes and tracheal branches of a crinoid-like structure underneath the epithelium. The wing tissue gradually became gray-white, epithelial cells were arranged regularly, and hemocytes disappeared, except in the bordering lacuna, after which scales grew. The dynamics of the epithelial cells and scale growth were also confirmed by fluorescent imaging. Fluorescent in vivo staining further revealed that these cells harbored many mitochondria at the surface of the epithelium. Organizing centers for the border symmetry system were apparent immediately after pupation, exhibiting a relatively dark optical character following treatment with fluorescent dyes, as well as in autofluorescent images. The wing tissue exhibited slow and low-frequency contraction pulses with a cycle of approximately 10 to 20 minutes, mainly occurring at 2 to 3 days postpupation. The pulses gradually became slower and weaker and eventually stopped. The wing tissue area became larger after contraction, which also coincided with an increase in the autofluorescence intensity that might have been caused by scale growth. Examination of the pattern of color development revealed that the black pigment was first deposited in patches in the central areas of an eyespot black ring and a parafocal element. These results of live in vivo imaging that covered wide wing area for a long time can serve as a foundation for studying the

  1. Aeroelastic Wing Shaping Using Distributed Propulsion

    Science.gov (United States)

    Nguyen, Nhan T. (Inventor); Reynolds, Kevin Wayne (Inventor); Ting, Eric B. (Inventor)

    2017-01-01

    An aircraft has wings configured to twist during flight. Inboard and outboard propulsion devices, such as turbofans or other propulsors, are connected to each wing, and are spaced along the wing span. A flight controller independently controls thrust of the inboard and outboard propulsion devices to significantly change flight dynamics, including changing thrust of outboard propulsion devices to twist the wing, and to differentially apply thrust on each wing to change yaw and other aspects of the aircraft during various stages of a flight mission. One or more generators can be positioned upon the wing to provide power for propulsion devices on the same wing, and on an opposite wing.

  2. Butterflies regulate wing temperatures using radiative cooling

    Science.gov (United States)

    Tsai, Cheng-Chia; Shi, Norman Nan; Ren, Crystal; Pelaez, Julianne; Bernard, Gary D.; Yu, Nanfang; Pierce, Naomi

    2017-09-01

    Butterfly wings are live organs embedded with multiple sensory neurons and, in some species, with pheromoneproducing cells. The proper function of butterfly wings demands a suitable temperature range, but the wings can overheat quickly in the sun due to their small thermal capacity. We developed an infrared technique to map butterfly wing temperatures and discovered that despite the wings' diverse visible colors, regions of wings that contain live cells are the coolest, resulting from the thickness of the wings and scale nanostructures. We also demonstrated that butterflies use behavioral traits to prevent overheating of their wings.

  3. Stiffness of desiccating insect wings

    Energy Technology Data Exchange (ETDEWEB)

    Mengesha, T E; Vallance, R R [Department of Mechanical Engineering, The George Washington University, 738 Phillips Hall, 801 22nd St NW, Washington, DC 20052 (United States); Mittal, R, E-mail: vallance@gwu.edu [Department of Mechanical Engineering, Johns Hopkins University, 126 Latrobe Hall, 3400 N Charles Street, Baltimore, MD 21218 (United States)

    2011-03-15

    The stiffness of insect wings is typically determined through experimental measurements. Such experiments are performed on wings removed from insects. However, the wings are subject to desiccation which typically leads to an increase in their stiffness. Although this effect of desiccation is well known, a comprehensive study of the rate of change in stiffness of desiccating insect wings would be a significant aid in planning experiments as well as interpreting data from such experiments. This communication presents a comprehensive experimental analysis of the change in mass and stiffness of gradually desiccating forewings of Painted Lady butterflies (Vanessa cardui). Mass and stiffness of the forewings of five butterflies were simultaneously measured every 10 min over a 24 h period. The averaged results show that wing mass declined exponentially by 21.1% over this time period with a time constant of 9.8 h, while wing stiffness increased linearly by 46.2% at a rate of 23.4 {mu}N mm{sup -1} h{sup -1}. For the forewings of a single butterfly, the experiment was performed over a period of 1 week, and the results show that wing mass declined exponentially by 52.2% with a time constant of 30.2 h until it reached a steady-state level of 2.00 mg, while wing stiffness increased exponentially by 90.7% until it reached a steady-state level of 1.70 mN mm{sup -1}. (communication)

  4. Morfometria e nervação foliar em procedências de erva-mate (Ilex paraguariensis A. St. Hill. (Aquifoliaceae = Morphometry and foliar venation in origins of maté (Ilex paraguariensis A. St. Hill. (Aquifoliaceae

    Directory of Open Access Journals (Sweden)

    Vânia Helena Techio

    2009-10-01

    Full Text Available O presente trabalho objetivou analisar a morfologia foliar morfometria e nervação foliar de procedências de erva-mate coletadas no Sul do Brasil e Argentina. Analisaram-se 30 folhas de cada procedência, obtendo-se cinco medidas (comprimento total da folha, largura máxima, comprimento desde a base até a largura máxima, comprimento do pecíolo e área e calculando-se dois índices (comprimento total sobre a largura máxima e comprimento desde a base até a largura máxima sobre o comprimento total da folha. Para anervação foliar, as folhas foram clarificadas por meio da técnica de diafanização. Os maiores e menores valores morfométricos foram, respectivamente, 118 e 67 mm para comprimento da folha, 51 e 34 mm para largura máxima, 78 e 42 mm para comprimento desde a base até alargura máxima, 16 e 10 mm para comprimento do pecíolo, 2,3 e 2 mm para comprimento sobre largura máxima e 0,65 e 0,6 para comprimento desde a base até a largura máxima sobre o comprimento da folha. Quanto à nervação foliar, não foram observadas diferenças entre as folhas das distintas procedências, as quais apresentaram nervação primária penada, nervação secundária semicraspedódroma, nervuras terciárias com percurso alternado e sinuoso, nervação quaternária reticulada, constituída por polígonos regulares e nervação de 5ª ordem dicotomizante.The present research aimed to evaluate the foliar morphology(morphometry and leaf venation of origins of maté collected in southern Brazil and Argentina. For the study of the morphometry, thirty leaves/plant were analyzed, and five measurements (leaf total length, maximum width, length from the base until maximum width, area and petiole length and two indices were obtained (total leaf length on maximum width and length from basis until maximum width on total leaf length. The leaves were clarified by the clarificationtechnique and examined in stereomicroscope. The highest and lowest values of leaf

  5. AERODYNAMICS OF WING TIP SAILS

    Directory of Open Access Journals (Sweden)

    MUSHTAK AL-ATABI

    2006-06-01

    Full Text Available Observers have always been fascinated by soaring birds. An interesting feature of these birds is the existence of few feathers extending from the tip of the wing. In this paper, small lifting surfaces were fitted to the tip of a NACA0012 wing in a fashion similar to that of wing tip feathers. Experimental measurements of induced drag, longitudinal static stability and trailing vortex structure were obtained.The tests showed that adding wing tip surfaces (sails decreased the induced drag factor and increased the longitudinal static stability. Results identified two discrete appositely rotated tip vortices and showed the ability of wing tip surfaces to break them down and to diffuse them.

  6. The marginal band system in nymphalid butterfly wings.

    Science.gov (United States)

    Taira, Wataru; Kinjo, Seira; Otaki, Joji M

    2015-01-01

    Butterfly wing color patterns are highly complex and diverse, but they are believed to be derived from the nymphalid groundplan, which is composed of several color pattern systems. Among these pattern systems, the marginal band system, including marginal and submarginal bands, has rarely been studied. Here, we examined the color pattern diversity of the marginal band system among nymphalid butterflies. Marginal and submarginal bands are usually expressed as a pair of linear bands aligned with the wing margin. However, a submarginal band can be expressed as a broken band, an elongated oval, or a single dot. The marginal focus, usually a white dot at the middle of a wing compartment along the wing edge, corresponds to the pupal edge spot, one of the pupal cuticle spots that signify the locations of color pattern organizing centers. A marginal band can be expressed as a semicircle, an elongated oval, or a pair of eyespot-like structures, which suggest the organizing activity of the marginal focus. Physical damage at the pupal edge spot leads to distal dislocation of the submarginal band in Junonia almana and in Vanessa indica, suggesting that the marginal focus functions as an organizing center for the marginal band system. Taken together, we conclude that the marginal band system is developmentally equivalent to other symmetry systems. Additionally, the marginal band is likely a core element and the submarginal band a paracore element of the marginal band system, and both bands are primarily specified by the marginal focus organizing center.

  7. Structural dynamics and aerodynamics measurements of biologically inspired flexible flapping wings.

    Science.gov (United States)

    Wu, P; Stanford, B K; Sällström, E; Ukeiley, L; Ifju, P G

    2011-03-01

    Flapping wing flight as seen in hummingbirds and insects poses an interesting unsteady aerodynamic problem: coupling of wing kinematics, structural dynamics and aerodynamics. There have been numerous studies on the kinematics and aerodynamics in both experimental and computational cases with both natural and artificial wings. These studies tend to ignore wing flexibility; however, observation in nature affirms that passive wing deformation is predominant and may be crucial to the aerodynamic performance. This paper presents a multidisciplinary experimental endeavor in correlating a flapping micro air vehicle wing's aeroelasticity and thrust production, by quantifying and comparing overall thrust, structural deformation and airflow of six pairs of hummingbird-shaped membrane wings of different properties. The results show that for a specific spatial distribution of flexibility, there is an effective frequency range in thrust production. The wing deformation at the thrust-productive frequencies indicates the importance of flexibility: both bending and twisting motion can interact with aerodynamic loads to enhance wing performance under certain conditions, such as the deformation phase and amplitude. By measuring structural deformations under the same aerodynamic conditions, beneficial effects of passive wing deformation can be observed from the visualized airflow and averaged thrust. The measurements and their presentation enable observation and understanding of the required structural properties for a thrust effective flapping wing. The intended passive responses of the different wings follow a particular pattern in correlation to their aerodynamic performance. Consequently, both the experimental technique and data analysis method can lead to further studies to determine the design principles for micro air vehicle flapping wings.

  8. Structural dynamics and aerodynamics measurements of biologically inspired flexible flapping wings

    International Nuclear Information System (INIS)

    Wu, P; Stanford, B K; Ifju, P G; Saellstroem, E; Ukeiley, L

    2011-01-01

    Flapping wing flight as seen in hummingbirds and insects poses an interesting unsteady aerodynamic problem: coupling of wing kinematics, structural dynamics and aerodynamics. There have been numerous studies on the kinematics and aerodynamics in both experimental and computational cases with both natural and artificial wings. These studies tend to ignore wing flexibility; however, observation in nature affirms that passive wing deformation is predominant and may be crucial to the aerodynamic performance. This paper presents a multidisciplinary experimental endeavor in correlating a flapping micro air vehicle wing's aeroelasticity and thrust production, by quantifying and comparing overall thrust, structural deformation and airflow of six pairs of hummingbird-shaped membrane wings of different properties. The results show that for a specific spatial distribution of flexibility, there is an effective frequency range in thrust production. The wing deformation at the thrust-productive frequencies indicates the importance of flexibility: both bending and twisting motion can interact with aerodynamic loads to enhance wing performance under certain conditions, such as the deformation phase and amplitude. By measuring structural deformations under the same aerodynamic conditions, beneficial effects of passive wing deformation can be observed from the visualized airflow and averaged thrust. The measurements and their presentation enable observation and understanding of the required structural properties for a thrust effective flapping wing. The intended passive responses of the different wings follow a particular pattern in correlation to their aerodynamic performance. Consequently, both the experimental technique and data analysis method can lead to further studies to determine the design principles for micro air vehicle flapping wings.

  9. Structural dynamics and aerodynamics measurements of biologically inspired flexible flapping wings

    Energy Technology Data Exchange (ETDEWEB)

    Wu, P; Stanford, B K; Ifju, P G [Department of Mechanical and Aerospace Engineering, MAE-A 231, University of Florida, Gainesville, FL 32611 (United States); Saellstroem, E; Ukeiley, L, E-mail: diccidwp@ufl.edu [Department of Mechanical and Aerospace Engineering, University of Florida, Shalimar, FL 32579 (United States)

    2011-03-15

    Flapping wing flight as seen in hummingbirds and insects poses an interesting unsteady aerodynamic problem: coupling of wing kinematics, structural dynamics and aerodynamics. There have been numerous studies on the kinematics and aerodynamics in both experimental and computational cases with both natural and artificial wings. These studies tend to ignore wing flexibility; however, observation in nature affirms that passive wing deformation is predominant and may be crucial to the aerodynamic performance. This paper presents a multidisciplinary experimental endeavor in correlating a flapping micro air vehicle wing's aeroelasticity and thrust production, by quantifying and comparing overall thrust, structural deformation and airflow of six pairs of hummingbird-shaped membrane wings of different properties. The results show that for a specific spatial distribution of flexibility, there is an effective frequency range in thrust production. The wing deformation at the thrust-productive frequencies indicates the importance of flexibility: both bending and twisting motion can interact with aerodynamic loads to enhance wing performance under certain conditions, such as the deformation phase and amplitude. By measuring structural deformations under the same aerodynamic conditions, beneficial effects of passive wing deformation can be observed from the visualized airflow and averaged thrust. The measurements and their presentation enable observation and understanding of the required structural properties for a thrust effective flapping wing. The intended passive responses of the different wings follow a particular pattern in correlation to their aerodynamic performance. Consequently, both the experimental technique and data analysis method can lead to further studies to determine the design principles for micro air vehicle flapping wings.

  10. Structural Analysis of a Dragonfly Wing

    NARCIS (Netherlands)

    Jongerius, S.R.; Lentink, D.

    2010-01-01

    Dragonfly wings are highly corrugated, which increases the stiffness and strength of the wing significantly, and results in a lightweight structure with good aerodynamic performance. How insect wings carry aerodynamic and inertial loads, and how the resonant frequency of the flapping wings is tuned

  11. Effect of outer wing separation on lift and thrust generation in a flapping wing system

    International Nuclear Information System (INIS)

    Mahardika, Nanang; Viet, Nguyen Quoc; Park, Hoon Cheol

    2011-01-01

    We explore the implementation of wing feather separation and lead-lagging motion to a flapping wing. A biomimetic flapping wing system with separated outer wings is designed and demonstrated. The artificial wing feather separation is implemented in the biomimetic wing by dividing the wing into inner and outer wings. The features of flapping, lead-lagging, and outer wing separation of the flapping wing system are captured by a high-speed camera for evaluation. The performance of the flapping wing system with separated outer wings is compared to that of a flapping wing system with closed outer wings in terms of forward force and downward force production. For a low flapping frequency ranging from 2.47 to 3.90 Hz, the proposed biomimetic flapping wing system shows a higher thrust and lift generation capability as demonstrated by a series of experiments. For 1.6 V application (lower frequency operation), the flapping wing system with separated wings could generate about 56% higher forward force and about 61% less downward force compared to that with closed wings, which is enough to demonstrate larger thrust and lift production capability of the separated outer wings. The experiments show that the outer parts of the separated wings are able to deform, resulting in a smaller amount of drag production during the upstroke, while still producing relatively greater lift and thrust during the downstroke.

  12. Flapping Wing Micro Air Vehicle Wing Manufacture and Force Testing

    Science.gov (United States)

    2011-03-03

    manufacturing techniques have been developed by various universities for research on Flapping Wing Micro Air Vehicles. Minimal attention though is given...collected at 2kHz (www.polytec.com/psv3d). A 0.25V band-limited white noise input signal is input to a Bogen HTA -125 High Performance Amplifier, which...manufacturing techniques have been developed by various universities for research on Flapping Wing Micro Air Vehicles. Minimal attention though is given

  13. Research of Morphing Wing Efficiency

    National Research Council Canada - National Science Library

    Komarov, Valery

    2004-01-01

    This report results from a contract tasking Samara State Aerospace University (SSAU) as follows: The contractor will develop and investigate aerodynamic and structural weight theories associated with morphing wing technology...

  14. Extensive Rangewide Mitochondrial Introgression Indicates Substantial Cryptic Hybridization in the Golden-winged Warbler (Vermivora chrysoptera

    Directory of Open Access Journals (Sweden)

    Rachel Vallender

    2009-12-01

    Full Text Available Widespread population declines of the Golden-winged Warbler (Vermivora chrysoptera are thought to be due in part to hybridization with the expanding Blue-winged Warbler (V. pinus, which predictably replaces Golden-winged Warblers at breeding sites in which the two species come into contact. However, the mechanism by which this replacement occurs remains unresolved. Recent genetic work has indicated that, even in areas where the two species have been in contact for a short period, introgression of Blue-winged mitochondrial (mtDNA and nuclear genes into Golden-winged individuals is common. To explore this process on a broader scale, we screened more than 750 individuals from nine U.S. states and three provinces to examine geographic patterns of mtDNA introgression. The only population in which all phenotypic Golden-winged Warblers had Golden-winged mtDNA haplotypes, and in which there are no breeding Blue-winged or hybrid individuals, was in the province of Manitoba, near the northwestern edge of the species' breeding distribution. The near ubiquity of mitochondrial introgression suggests that there are far fewer genetically pure populations of Golden-winged Warblers than previously believed, a finding with important implications for this threatened species.

  15. Experimental and computational study of transonic flow about swept wings

    Science.gov (United States)

    Bertelrud, A.; Bergmann, M. Y.; Coakley, T. J.

    1980-01-01

    An experimental investigation of NACA 0010 and 10% circular arc wing models, swept at 45 deg, spanning a channel, and at zero angle of attack is described. Measurements include chordwise and spanwise surface pressure distributions and oil-flow patterns for a range of transonic Mach numbers and Reynolds numbers. Calculations using a new three-dimensional Navier-Stokes code and a two-equation turbulence model are included for the circular-arc wing flow. Reasonable agreement between measurements and computations is obtained.

  16. Anopheles darlingi (Diptera: Culicidae Rood 1926: Morphometric variations in wings and legs of populations from Colombia

    Directory of Open Access Journals (Sweden)

    Miguel Alfonso Pacheco

    2017-03-01

    Conclusions. We registered 11 new wing spot patterns in the costal vein and the dominance of the patterns I and VI for populations of An. darlingi from Colombia. We confirmed DSIII2/TaIII2 ratio as a robust diagnostic character for the taxonomy of this species. We found differences between the size and shape of the wings of An. darlingi populations in accordance to their geographical distribution, which constitute important bionomic aspects for this malaria vector.

  17. Transcriptome profiling with focus on potential key genes for wing development and evolution in Megaloprepus caerulatus, the damselfly species with the world's largest wings.

    Science.gov (United States)

    Feindt, Wiebke; Oppenheim, Sara J; DeSalle, Robert; Goldstein, Paul Z; Hadrys, Heike

    2018-01-01

    The evolution, development and coloration of insect wings remains a puzzling subject in evolutionary research. In basal flying insects such as Odonata, genomic research regarding bauplan evolution is still rare. Here we focus on the world's largest odonate species-the "forest giant" Megaloprepus caerulatus, to explore its potential for looking deeper into the development and evolution of wings. A recently discovered cryptic species complex in this genus previously considered monotypic is characterized by morphological differences in wing shape and color patterns. As a first step toward understanding wing pattern divergence and pathways involved in adaptation and speciation at the genomic level, we present a transcriptome profiling of M. caerulatus using RNA-Seq and compare these data with two other odonate species. The de novo transcriptome assembly consists of 61,560 high quality transcripts and is approximately 93% complete. For almost 75% of the identified transcripts a possible function could be assigned: 48,104 transcripts had a hit to an InterPro protein family or domain, and 28,653 were mapped to a Gene Ontology term. In particular, we focused on genes related to wing development and coloration. The comparison with two other species revealed larva-specific genes and a conserved 'core' set of over 8,000 genes forming orthologous clusters with Ischnura elegans and Ladona fulva. This transcriptome may provide a first point of reference for future research in odonates addressing questions surrounding the evolution of wing development, wing coloration and their role in speciation.

  18. Wing shape variation associated with mimicry in butterflies.

    Science.gov (United States)

    Jones, Robert T; Le Poul, Yann; Whibley, Annabel C; Mérot, Claire; ffrench-Constant, Richard H; Joron, Mathieu

    2013-08-01

    Mimetic resemblance in unpalatable butterflies has been studied by evolutionary biologists for over a century, but has largely focused on the convergence in wing color patterns. In Heliconius numata, discrete color-pattern morphs closely resemble comimics in the distantly related genus Melinaea. We examine the possibility that the shape of the butterfly wing also shows adaptive convergence. First, simple measures of forewing dimensions were taken of individuals in a cross between H. numata morphs, and showed quantitative differences between two of the segregating morphs, f. elegans and f. silvana. Second, landmark-based geometric morphometric and elliptical Fourier outline analyses were used to more fully characterize these shape differences. Extension of these techniques to specimens from natural populations suggested that, although many of the coexisting morphs could not be discriminated by shape, the differences we identified between f. elegans and f. silvana hold in the wild. Interestingly, despite extensive overlap, the shape variation between these two morphs is paralleled in their respective Melinaea comimics. Our study therefore suggests that wing-shape variation is associated with mimetic resemblance, and raises the intriguing possibility that the supergene responsible for controlling the major switch in color pattern between morphs also contributes to wing shape differences in H. numata. © 2013 The Author(s). Evolution © 2013 The Society for the Study of Evolution.

  19. Effects of external influences in subsonic delta wing vortices

    Science.gov (United States)

    Washburn, Anthony E.

    1992-01-01

    An experimental investigation was conducted to examine inconsistencies in reported studies for the vortical flow over highly-swept delta wings. A 76-deg swept delta wing was tested in three facilities with open and closed test sections and different model-support systems. The results obtained include surface oil-flow patterns, off-body laser-light-sheet flow visualization, and aerodynamic load measurements. Parameters such as the wall boundaries and model-support systems can drastically alter the loads. The effect of a high level of free-stream turbulence on the delta-wing flowfield was also examined and found to be significant. The increase in free-stream turbulence caused boundary-layer transition, unsteadiness in the vortex core positions, and altered the loads and moments.

  20. Shape matters: improved flight in tapered auto-rotating wings

    Science.gov (United States)

    Liu, Yucen; Vincent, Lionel; Kanso, Eva

    2017-11-01

    Many plants use gravity and wind to disperse their seeds. The shape of seed pods influence their aerodynamics. For example, Liana seeds form aerodynamic gliders and Sycamore trees release airborne ``helicopters.'' Here, we use carefully-controlled experiments and high-speed photography to examine dispersion by tumbling (auto-rotation) and we focus on the effect of geometry on flight characteristics. We consider four families of shapes: rectangular, elliptic, tapered, and sharp-tip wings, and we vary the span-to-chord ratio. We find that tapered wings exhibit extended flight time and range, that is, better performance. A quasi-steady two-dimensional model is used to highlight the mechanisms by which shape affects flight performance. These findings could have significant implications on linking seedpod designs to seed dispersion patterns as well as on optimizing wing design in active flight problems.

  1. Comparación de la morfología alar de Tadarida brasiliensis (Chiroptera: Molossidae y Myotis chiloensis (Chiroptera: Vespertilionidae, representantes de dos diferentes patrones de vuelo Comparison of the wing morphology of Tadarida brasiliensis (Chiroptera: Molossidae and Myotis chiloensis (Chiroptera: Vespertilionidae as representatives of two flight patterns

    Directory of Open Access Journals (Sweden)

    MAURICIO CANALS

    2001-09-01

    Full Text Available La morfología alar de los quirópteros se encuentra relacionada por una parte con la biomecánica y energética del vuelo y por otra parte con aspectos ecológicos y conductuales (i.e., patrón de vuelo, conducta de forrajeo y selección de hábitat y de presas. En este trabajo se compara la morfometría alar de Tadarida brasiliensis (Molossidae y Myotis chiloensis (Vespertilionidae, representantes de diferentes patrones de vuelo, buscando compromisos entre la morfometría alar y aspectos ecológicos y conductuales. Nuestros resultados muestran que T. brasiliensis es un murciélago más robusto, de mayor envergadura, pero con un área alar similar a la de M. chiloensis. Esta última especie tiene una menor variabilidad en su masa y área cortical del húmero, que probablemente se encuentre relacionada con restricciones mecánicas y energéticas impuestas por su diseño. Descontando el efecto de la masa se detectaron diferencias en el diámetro externo y diámetro medular del húmero con una similar área cortical. El húmero de T. brasiliensis es un hueso de similar longitud, más ancho y con un menor grosor cortical que el de M. chiloensis, lo que está relacionado con una mayor resistencia a las fuerzas de flexión y torsión. Las características alares son consistentes con los modos de vida de cada murciélago: vuelos lentos, cortos y maniobrables en zonas arbustivas de M. chiloensis y vuelo veloz y de grandes distancias en espacios abiertos de T. brasiliensisWing morphology is related by one hand to biomechanical properties and energetics of flying, and on the other hand to ecological and behavioral aspects of flying, such as flight pattern, foraging behavior, habitat selection and size of prey. In this work we compare the wing morphology of Tadarida brasiliensis (Molossidae and Myotis chiloensis (Vespertilionidae, as representatives of two flight patterns, and looking for trade-offs between wing morphology, ecology and behavior. Our

  2. Imaging optical scattering of butterfly wing scales with a microscope.

    Science.gov (United States)

    Fu, Jinxin; Yoon, Beom-Jin; Park, Jung Ok; Srinivasarao, Mohan

    2017-08-06

    A new optical method is proposed to investigate the reflectance of structurally coloured objects, such as Morpho butterfly wing scales and cholesteric liquid crystals. Using a reflected-light microscope and a digital single-lens reflex (DSLR) camera, we have successfully measured the two-dimensional reflection pattern of individual wing scales of Morpho butterflies. We demonstrate that this method enables us to measure the bidirectional reflectance distribution function (BRDF). The scattering image observed in the back focal plane of the objective is projected onto the camera sensor by inserting a Bertrand lens in the optical path of the microscope. With monochromatic light illumination, we quantify the angle-dependent reflectance spectra from the wing scales of Morpho rhetenor by retrieving the raw signal from the digital camera sensor. We also demonstrate that the polarization-dependent reflection of individual wing scales is readily observed using this method, using the individual wing scales of Morpho cypris . In an effort to show the generality of the method, we used a chiral nematic fluid to illustrate the angle-dependent reflectance as seen by this method.

  3. Waterproof and translucent wings at the same time: problems and solutions in butterflies

    Science.gov (United States)

    Perez Goodwyn, Pablo; Maezono, Yasunori; Hosoda, Naoe; Fujisaki, Kenji

    2009-07-01

    Although the colour of butterflies attracts the most attention, the waterproofing properties of their wings are also extremely interesting. Most butterfly wings are considered “super-hydrophobic” because the contact angle (CA) with a water drop exceeds 150°. Usually, butterfly wings are covered with strongly overlapping scales; however, in the case of transparent or translucent wings, scale cover is reduced; thus, the hydrophobicity could be affected. Here, we present a comparative analysis of wing hydrophobicity and its dependence on morphology for two species with translucent wings Parantica sita (Nymphalidae) and Parnassius glacialis (Papilionidae). These species have very different life histories: P. sita lives for up to 6 months as an adult and migrates over long distance, whereas P. glacialis lives for less than 1 month and does not migrate. We measured the water CA and analysed wing morphology with scanning electron microscopy and atomic force microscopy. P. sita has super-hydrophobic wing surfaces, with CA > 160°, whereas P. glacialis did not (CA = 100-135°). Specialised scales were found on the translucent portions of P. sita wings. These scales were ovoid and much thinner than common scales, erect at about 30°, and leaving up to 80% of the wing surface uncovered. The underlying bare wing surface had a remarkable pattern of ridges and knobs. P. glacialis also had over 80% of the wing surface uncovered, but the scales were either setae-like or spade-like. The bare surface of the wing had an irregular wavy smooth pattern. We suggest a mode of action that allows this super-hydrophobic effect with an incompletely covered wing surface. The scales bend, but do not collapse, under the pressure of a water droplet, and the elastic recovery of the structure at the borders of the droplet allows a high apparent CA. Thus, P. sita can be translucent without losing its waterproof properties. This characteristic is likely necessary for the long life and migration

  4. The Eastern Arc Mountains and coastal forests of East Africa-an archive to understand large-scale biogeographical patterns: Pseudotomias, a new genus of African Pseudophyllinae (Orthoptera: Tettigoniidae).

    Science.gov (United States)

    Hemp, Claudia

    2016-06-21

    A new genus of Pseudophyllinae restricted to East Africa is described. Data on the ecology, and the habitat are provided. The biogeographical pattern and morphology suggests an old radiation since Tomias from Central and West Africa is the closest relative to Pseudotomias. The old forests of East Africa could hereby be the source of representatives of this old radiation since venation is less reduced in East African taxa of Phyllomimini.

  5. The Efficiency of a Hybrid Flapping Wing Structure—A Theoretical Model Experimentally Verified

    Directory of Open Access Journals (Sweden)

    Yuval Keren

    2016-07-01

    Full Text Available To propel a lightweight structure, a hybrid wing structure was designed; the wing’s geometry resembled a rotor blade, and its flexibility resembled an insect’s flapping wing. The wing was designed to be flexible in twist and spanwise rigid, thus maintaining the aeroelastic advantages of a flexible wing. The use of a relatively “thick” airfoil enabled the achievement of higher strength to weight ratio by increasing the wing’s moment of inertia. The optimal design was based on a simplified quasi-steady inviscid mathematical model that approximately resembles the aerodynamic and inertial behavior of the flapping wing. A flapping mechanism that imitates the insects’ flapping pattern was designed and manufactured, and a set of experiments for various parameters was performed. The simplified analytical model was updated according to the tests results, compensating for the viscid increase of drag and decrease of lift, that were neglected in the simplified calculations. The propelling efficiency of the hovering wing at various design parameters was calculated using the updated model. It was further validated by testing a smaller wing flapping at a higher frequency. Good and consistent test results were obtained in line with the updated model, yielding a simple, yet accurate tool, for flapping wings design.

  6. Conceptual Study of Rotary-Wing Microrobotics

    National Research Council Canada - National Science Library

    Chabak, Kelson D

    2008-01-01

    This thesis presents a novel rotary-wing micro-electro-mechanical systems (MEMS) robot design. Two MEMS wing designs were designed, fabricated and tested including one that possesses features conducive to insect level aerodynamics...

  7. Aerodynamic control with passively pitching wings

    Science.gov (United States)

    Gravish, Nick; Wood, Robert

    Flapping wings may pitch passively under aerodynamic and inertial loads. Such passive pitching is observed in flapping wing insect and robot flight. The effect of passive wing pitch on the control dynamics of flapping wing flight are unexplored. Here we demonstrate in simulation and experiment the critical role wing pitching plays in yaw control of a flapping wing robot. We study yaw torque generation by a flapping wing allowed to passively rotate in the pitch axis through a rotational spring. Yaw torque is generated through alternating fast and slow upstroke and and downstroke. Yaw torque sensitively depends on both the rotational spring force law and spring stiffness, and at a critical spring stiffness a bifurcation in the yaw torque control relationship occurs. Simulation and experiment reveal the dynamics of this bifurcation and demonstrate that anomalous yaw torque from passively pitching wings is the result of aerodynamic and inertial coupling between the pitching and stroke-plane dynamics.

  8. Wind tunnel investigation of the interaction and breakdown characteristics of slender wing vortices at subsonic, transonic, and supersonic speeds

    Science.gov (United States)

    Erickson, Gary E.

    1991-01-01

    The vortex dominated aerodynamic characteristics of a generic 65 degree cropped delta wing model were studied in a wind tunnel at subsonic through supersonic speeds. The lee-side flow fields over the wing-alone configuration and the wing with leading edge extension (LEX) added were observed at M (infinity) equals 0.40 to 1.60 using a laser vapor screen technique. These results were correlated with surface streamline patterns, upper surface static pressure distributions, and six-component forces and moments. The wing-alone exhibited vortex breakdown and asymmetry of the breakdown location at the subsonic and transonic speeds. An earlier onset of vortex breakdown over the wing occurred at transonic speeds due to the interaction of the leading edge vortex with the normal shock wave. The development of a shock wave between the vortex and wing surface caused an early separation of the secondary boundary layer. With the LEX installed, wing vortex breakdown asymmetry did not occur up to the maximum angle of attack in the present test of 24 degrees. The favorable interaction of the LEX vortex with the wing flow field reduced the effects of shock waves on the wing primary and secondary vortical flows. The direct interaction of the wing and LEX vortex cores diminished with increasing Mach number. The maximum attainable vortex-induced pressure signatures were constrained by the vacuum pressure limit at the transonic and supersonic speeds.

  9. Waving Wing Aerodynamics at Low Reynolds Numbers

    Science.gov (United States)

    2010-07-01

    wing. An attached leading edge vortex has been observed by multiple research groups on both mechanical wing flappers (8; 22; 21; 4) and revolving wing...observed by Ellington et al. (8) in their earlier experiments on the mechanical hawkmoth flapper at Re ≈ 10,000. In these experiments the spanwise flow...on mechanical wing flappers at similar Reynolds numbers, Re ≈ 1,000 and 1,400 respectively. Both sets of experiments revealed a stable attached

  10. Cellular basis of morphological variation and temperature-related plasticity in Drosophila melanogaster strains with divergent wing shapes.

    Science.gov (United States)

    Torquato, Libéria Souza; Mattos, Daniel; Matta, Bruna Palma; Bitner-Mathé, Blanche Christine

    2014-12-01

    Organ shape evolves through cross-generational changes in developmental patterns at cellular and/or tissue levels that ultimately alter tissue dimensions and final adult proportions. Here, we investigated the cellular basis of an artificially selected divergence in the outline shape of Drosophila melanogaster wings, by comparing flies with elongated or rounded wing shapes but with remarkably similar wing sizes. We also tested whether cellular plasticity in response to developmental temperature was altered by such selection. Results show that variation in cellular traits is associated with wing shape differences, and that cell number may play an important role in wing shape response to selection. Regarding the effects of developmental temperature, a size-related plastic response was observed, in that flies reared at 16 °C developed larger wings with larger and more numerous cells across all intervein regions relative to flies reared at 25 °C. Nevertheless, no conclusive indication of altered phenotypic plasticity was found between selection strains for any wing or cellular trait. We also described how cell area is distributed across different intervein regions. It follows that cell area tends to decrease along the anterior wing compartment and increase along the posterior one. Remarkably, such pattern was observed not only in the selected strains but also in the natural baseline population, suggesting that it might be canalized during development and was not altered by the intense program of artificial selection for divergent wing shapes.

  11. Aerodynamic forces and flow structures of the leading edge vortex on a flapping wing considering ground effect

    International Nuclear Information System (INIS)

    Truong, Tien Van; Yoon, Kwang Joon; Byun, Doyoung; Kim, Min Jun; Park, Hoon Cheol

    2013-01-01

    The aim of this work is to provide an insight into the aerodynamic performance of the beetle during takeoff, which has been estimated in previous investigations. We employed a scaled-up electromechanical model flapping wing to measure the aerodynamic forces and the three-dimensional flow structures on the flapping wing. The ground effect on the unsteady forces and flow structures were also characterized. The dynamically scaled wing model could replicate the general stroke pattern of the beetle's hind wing kinematics during takeoff flight. Two wing kinematic models have been studied to examine the influences of wing kinematics on unsteady aerodynamic forces. In the first model, the angle of attack is asymmetric and varies during the translational motion, which is the flapping motion of the beetle's hind wing. In the second model, the angle of attack is constant during the translational motion. The instantaneous aerodynamic forces were measured for four strokes during the beetle's takeoff by the force sensor attached at the wing base. Flow visualization provided a general picture of the evolution of the three-dimensional leading edge vortex (LEV) on the beetle hind wing model. The LEV is stable during each stroke, and increases radically from the root to the tip, forming a leading-edge spiral vortex. The force measurement results show that the vertical force generated by the hind wing is large enough to lift the beetle. For the beetle hind wing kinematics, the total vertical force production increases 18.4% and 8.6% for the first and second strokes, respectively, due to the ground effect. However, for the model with a constant angle of attack during translation, the vertical force is reduced during the first stroke. During the third and fourth strokes, the ground effect is negligible for both wing kinematic patterns. This finding suggests that the beetle's flapping mechanism induces a ground effect that can efficiently lift its body from the ground during takeoff

  12. Aerodynamic forces and flow structures of the leading edge vortex on a flapping wing considering ground effect.

    Science.gov (United States)

    Van Truong, Tien; Byun, Doyoung; Kim, Min Jun; Yoon, Kwang Joon; Park, Hoon Cheol

    2013-09-01

    The aim of this work is to provide an insight into the aerodynamic performance of the beetle during takeoff, which has been estimated in previous investigations. We employed a scaled-up electromechanical model flapping wing to measure the aerodynamic forces and the three-dimensional flow structures on the flapping wing. The ground effect on the unsteady forces and flow structures were also characterized. The dynamically scaled wing model could replicate the general stroke pattern of the beetle's hind wing kinematics during takeoff flight. Two wing kinematic models have been studied to examine the influences of wing kinematics on unsteady aerodynamic forces. In the first model, the angle of attack is asymmetric and varies during the translational motion, which is the flapping motion of the beetle's hind wing. In the second model, the angle of attack is constant during the translational motion. The instantaneous aerodynamic forces were measured for four strokes during the beetle's takeoff by the force sensor attached at the wing base. Flow visualization provided a general picture of the evolution of the three-dimensional leading edge vortex (LEV) on the beetle hind wing model. The LEV is stable during each stroke, and increases radically from the root to the tip, forming a leading-edge spiral vortex. The force measurement results show that the vertical force generated by the hind wing is large enough to lift the beetle. For the beetle hind wing kinematics, the total vertical force production increases 18.4% and 8.6% for the first and second strokes, respectively, due to the ground effect. However, for the model with a constant angle of attack during translation, the vertical force is reduced during the first stroke. During the third and fourth strokes, the ground effect is negligible for both wing kinematic patterns. This finding suggests that the beetle's flapping mechanism induces a ground effect that can efficiently lift its body from the ground during takeoff.

  13. Wing area, wing growth and wing loading of common sandpipers Actitis hypoleucos

    OpenAIRE

    Yalden, Derek; Yalden, D. W.

    2012-01-01

    This study investigates the changes in wing length, area and loading in Common Sandpipers as chicks grow, and as adults add extra mass (during egg-laying or before migration). Common Sandpiper chicks weigh about 17 g and have "hands" that are about 35 mm long at one week old, when the primaries are just emerging from their sheaths. They grow steadily to reach about 40 g, with hands about 85 mm long, at 19 days, when they are just about fledging. Their wings have roughly adult chord width at t...

  14. How Do Wings Generate Lift?

    Indian Academy of Sciences (India)

    Newton's second law of motion. Hence if a wing can generate lift equal to its weight (total weight of the vehicle) it can balance the gravitational pull and can maintain level flight. The equations for fluid flow that are equivalent to the second law are the well- known Navier–Stokes (N–S) equations [1]. These equations have.

  15. Werner helicase wings DNA binding

    OpenAIRE

    Hoadley, Kelly A.; Keck, James L.

    2010-01-01

    In this issue of Structure, Kitano et al. describe the structure of the DNA-bound winged-helix domain from the Werner helicase. This structure of a RecQ/DNA complex offers insights into the DNA unwinding mechanisms of RecQ family helicases.

  16. On Wings: Aerodynamics of Eagles.

    Science.gov (United States)

    Millson, David

    2000-01-01

    The Aerodynamics Wing Curriculum is a high school program that combines basic physics, aerodynamics, pre-engineering, 3D visualization, computer-assisted drafting, computer-assisted manufacturing, production, reengineering, and success in a 15-hour, 3-week classroom module. (JOW)

  17. Qualitative skeletal correlates of wing shape in extant birds (Aves: Neoaves)

    OpenAIRE

    Hieronymus, Tobin L

    2015-01-01

    Background Among living fliers (birds, bats, and insects), birds display relatively high aspect ratios, a dimensionless shape variable that distinguishes long and narrow vs. short and broad wings. Increasing aspect ratio results in a functional tradeoff between low induced drag (efficient cruise) and increased wing inertia (difficult takeoff). Given the wide scope of its functional effects, the pattern of aspect ratio evolution is an important factor that contributes to the substantial ecolog...

  18. Airplane wing deformation and flight flutter detection method by using three-dimensional speckle image correlation technology.

    Science.gov (United States)

    Wu, Jun; Yu, Zhijing; Wang, Tao; Zhuge, Jingchang; Ji, Yue; Xue, Bin

    2017-06-01

    Airplane wing deformation is an important element of aerodynamic characteristics, structure design, and fatigue analysis for aircraft manufacturing, as well as a main test content of certification regarding flutter for airplanes. This paper presents a novel real-time detection method for wing deformation and flight flutter detection by using three-dimensional speckle image correlation technology. Speckle patterns whose positions are determined through the vibration characteristic of the aircraft are coated on the wing; then the speckle patterns are imaged by CCD cameras which are mounted inside the aircraft cabin. In order to reduce the computation, a matching technique based on Geodetic Systems Incorporated coded points combined with the classical epipolar constraint is proposed, and a displacement vector map for the aircraft wing can be obtained through comparing the coordinates of speckle points before and after deformation. Finally, verification experiments containing static and dynamic tests by using an aircraft wing model demonstrate the accuracy and effectiveness of the proposed method.

  19. Review Results on Wing-Body Interference

    Directory of Open Access Journals (Sweden)

    Frolov Vladimir

    2016-01-01

    Full Text Available The paper presents an overview of results for wing-body interference, obtained by the author for varied wing-body combinations. The lift-curve slopes of the wing-body combinations are considered. In this paper a discrete vortices method (DVM and 2D potential model for cross-flow around fuselage are used. The circular and elliptical cross-sections of the fuselage and flat wings of various forms are considered. Calculations showed that the value of the lift-curve slopes of the wing-body combinations may exceed the same value for an isolated wing. This result confirms an experimental data obtained by other authors earlier. Within a framework of the used mathematical models the investigations to optimize the wing-body combination were carried. The present results of the optimization problem for the wing-body combination allowed to select the optimal geometric characteristics for configuration to maximize the values of the lift-curve slopes of the wing-body combination. It was revealed that maximums of the lift-curve slopes for the optimal mid-wing configuration with elliptical cross-section body had a sufficiently large relative width of the body (more than 30% of the span wing.

  20. A single basis for developmental buffering of Drosophila wing shape.

    Directory of Open Access Journals (Sweden)

    Casper J Breuker

    2006-12-01

    Full Text Available The nature of developmental buffering processes has been debated extensively, based on both theoretical reasoning and empirical studies. In particular, controversy has focused on the question of whether distinct processes are responsible for canalization, the buffering against environmental or genetic variation, and for developmental stability, the buffering against random variation intrinsic in developmental processes. Here, we address this question for the size and shape of Drosophila melanogaster wings in an experimental design with extensively replicated and fully controlled genotypes. The amounts of variation among individuals and of fluctuating asymmetry differ markedly among genotypes, demonstrating a clear genetic basis for size and shape variability. For wing shape, there is a high correlation between the amounts of variation among individuals and fluctuating asymmetry, which indicates a correspondence between the two types of buffering. Likewise, the multivariate patterns of shape variation among individuals and of fluctuating asymmetry show a close association. For wing size, however, the amounts of individual variation and fluctuating asymmetry are not correlated. There was a significant link between the amounts of variation between wing size and shape, more so for fluctuating asymmetry than for variation among individuals. Overall, these experiments indicate a considerable degree of shared control of individual variation and fluctuating asymmetry, although it appears to differ between traits.

  1. Winged messengers of disaster

    International Nuclear Information System (INIS)

    Medvedev, Z.

    1977-01-01

    The work of the Soviet ecologists, led by A.I. Il'enko, on birds in the southern Urals area, site of the nuclear disaster in 1958, is discussed. The distribution of 90 Sr and 137 Cs in birds, food chains in a large running-water lake, bird migration patterns, and nest conservatism of ducks have been studied. It is pointed out that the existence of migratory species among contaminated species of the southern Urals provides an opportunity for observers in the West to test the truth about the 1958 nuclear disaster in the southern Urals. It is felt that the reports discussed here corroborate the author's original statement that the Urals nuclear disaster involved nuclear waste rather than a major reactor accident. (U.K.)

  2. A Drosophila wing spot test

    International Nuclear Information System (INIS)

    Ayaki, Toshikazu; Yoshikawa, Isao; Niikawa, Norio; Hoshi, Masaharu.

    1986-01-01

    A Drosophila wing spot test system was used to investigate the effects of low doses of X-rays, gamma rays, and both 2.3 and 14.1 MeV neutrons on somatic chromosome mutation (SCM) induction. The incidence of SCM was significantly increased with any type of radiation, with evident linear dose-response relationship within the range of 3 to 20 cGy. It was estimated that relative biological effectiveness value for SCM induction of 2.3 MeV neutrons to X-rays and gamma rays is much higher than that of 14.1 MeV neutrons to those photons (2.4 vs 8.0). The Drosophila wing spot test system seems to become a promising in vivo experimental method for higher animals in terms of the lack of necessity for a marvelously large number of materials required in conventional test system. (Namekawa, K.)

  3. Aerodynamic effects of flexibility in flapping wings

    Science.gov (United States)

    Zhao, Liang; Huang, Qingfeng; Deng, Xinyan; Sane, Sanjay P.

    2010-01-01

    Recent work on the aerodynamics of flapping flight reveals fundamental differences in the mechanisms of aerodynamic force generation between fixed and flapping wings. When fixed wings translate at high angles of attack, they periodically generate and shed leading and trailing edge vortices as reflected in their fluctuating aerodynamic force traces and associated flow visualization. In contrast, wings flapping at high angles of attack generate stable leading edge vorticity, which persists throughout the duration of the stroke and enhances mean aerodynamic forces. Here, we show that aerodynamic forces can be controlled by altering the trailing edge flexibility of a flapping wing. We used a dynamically scaled mechanical model of flapping flight (Re ≈ 2000) to measure the aerodynamic forces on flapping wings of variable flexural stiffness (EI). For low to medium angles of attack, as flexibility of the wing increases, its ability to generate aerodynamic forces decreases monotonically but its lift-to-drag ratios remain approximately constant. The instantaneous force traces reveal no major differences in the underlying modes of force generation for flexible and rigid wings, but the magnitude of force, the angle of net force vector and centre of pressure all vary systematically with wing flexibility. Even a rudimentary framework of wing veins is sufficient to restore the ability of flexible wings to generate forces at near-rigid values. Thus, the magnitude of force generation can be controlled by modulating the trailing edge flexibility and thereby controlling the magnitude of the leading edge vorticity. To characterize this, we have generated a detailed database of aerodynamic forces as a function of several variables including material properties, kinematics, aerodynamic forces and centre of pressure, which can also be used to help validate computational models of aeroelastic flapping wings. These experiments will also be useful for wing design for small robotic

  4. Aerodynamic effects of flexibility in flapping wings.

    Science.gov (United States)

    Zhao, Liang; Huang, Qingfeng; Deng, Xinyan; Sane, Sanjay P

    2010-03-06

    Recent work on the aerodynamics of flapping flight reveals fundamental differences in the mechanisms of aerodynamic force generation between fixed and flapping wings. When fixed wings translate at high angles of attack, they periodically generate and shed leading and trailing edge vortices as reflected in their fluctuating aerodynamic force traces and associated flow visualization. In contrast, wings flapping at high angles of attack generate stable leading edge vorticity, which persists throughout the duration of the stroke and enhances mean aerodynamic forces. Here, we show that aerodynamic forces can be controlled by altering the trailing edge flexibility of a flapping wing. We used a dynamically scaled mechanical model of flapping flight (Re approximately 2000) to measure the aerodynamic forces on flapping wings of variable flexural stiffness (EI). For low to medium angles of attack, as flexibility of the wing increases, its ability to generate aerodynamic forces decreases monotonically but its lift-to-drag ratios remain approximately constant. The instantaneous force traces reveal no major differences in the underlying modes of force generation for flexible and rigid wings, but the magnitude of force, the angle of net force vector and centre of pressure all vary systematically with wing flexibility. Even a rudimentary framework of wing veins is sufficient to restore the ability of flexible wings to generate forces at near-rigid values. Thus, the magnitude of force generation can be controlled by modulating the trailing edge flexibility and thereby controlling the magnitude of the leading edge vorticity. To characterize this, we have generated a detailed database of aerodynamic forces as a function of several variables including material properties, kinematics, aerodynamic forces and centre of pressure, which can also be used to help validate computational models of aeroelastic flapping wings. These experiments will also be useful for wing design for small

  5. Shape optimisation and performance analysis of flapping wings

    KAUST Repository

    Ghommem, Mehdi

    2012-09-04

    -averaged thrust, while the average aerodynamic power is increased. Furthermore, increasing the number of variables (i.e., providing the wing shape with greater degrees of spatial freedom) is observed to enable superior designs. To gain a better understanding of the reasons for which the obtained optimised shapes produce efficient flapping flights, the wake pattern and its vorticity strength are examined. This work described in this paper should facilitate better guidance for shape design of engineered flying systems.

  6. Melanin pigmentation gives rise to black spots on the wings of the silkworm Bombyx mori.

    Science.gov (United States)

    Ito, Katsuhiko; Yoshikawa, Manabu; Fujii, Takeshi; Tabunoki, Hiroko; Yokoyama, Takeshi

    2016-01-01

    Several mutants of the silkworm Bombyx mori show body color variation at the larval and adult stages. The Wild wing spot (Ws) mutant exhibits a phenotype in which the moth has a spot on the apex of the forewing. In this study, we investigated this trait to elucidate the molecular mechanism underlying the color pattern. Microscopy of the black spot of Ws mutants showed that the pigment emerges in the scales of the wing, and accumulation of the pigment becomes strong just before eclosion. We next examined the relationship between the black spot of the Ws mutant and melanin. The spectrophotometry using alkaline extracts from the black spot in the wing showed the highest absorption intensity at 405nm, which is the absorbance wavelength of melanin. Moreover, inhibition assays for enzymes implicated in melanin synthesis using 3-iodo-l-tyrosine (a tyrosine hydroxylase inhibitor) and L-α-methyl-DOPA (a dopa decarboxylase inhibitor) revealed that treatment with each inhibitor disrupted the pigmentation of the wing of the Ws mutant. On the basis of these results, we analyzed the expression pattern of five genes involved in melanin formation, and found that the expression levels of yellow and laccase2 were increased just before pigmentation, whereas those of DDC, tan, and TH were increased when the apex of the wing turned black. These results showed that melanin pigmentation gives rise to the black spot on the wing. Copyright © 2016 Elsevier Ltd. All rights reserved.

  7. The costae presenting in high-temperature-induced vestigial wings ...

    Indian Academy of Sciences (India)

    Most of the wings have defects in the wing blade and partially formed wing margin, which are the result of autonomous cell death in the ... [Yang D. 2007 The costae presenting in high-temperature-induced vestigial wings of Drosophila: implications for anterior wing margin formation. J. Genet. .... The relevant gene(s) may be.

  8. Role of wing morphing in thrust generation

    Directory of Open Access Journals (Sweden)

    Mehdi Ghommem

    2014-01-01

    Full Text Available In this paper, we investigate the role of morphing on flight dynamics of two birds by simulating the flow over rigid and morphing wings that have the characteristics of two different birds, namely the Giant Petrel and Dove Prion. The simulation of a flapping rigid wing shows that the root of the wing should be placed at a specific angle of attack in order to generate enough lift to balance the weight of the bird. However, in this case the generated thrust is either very small, or even negative, depending on the wing shape. Further, results show that morphing of the wing enables a significant increase in the thrust and propulsive efficiency. This indicates that the birds actually utilize some sort of active wing twisting and bending to produce enough thrust. This study should facilitate better guidance for the design of flapping air vehicles.

  9. Winged scapula incidence and upper limb morbidity after surgery for breast cancer with axillary dissection.

    Science.gov (United States)

    Rizzi, Samantha Karlla Lopes de Almeida; Haddad, Cinira Assad Simão; Giron, Patricia Santolia; Pinheiro, Thaís Lúcia; Nazário, Afonso Celso Pinto; Facina, Gil

    2016-06-01

    The aim of this study was to determine the incidence of winged scapula after breast cancer surgery, its impact on shoulder morbidity and difference in incidence according to surgery type. Patients with breast cancer and surgical indication for axillary dissection were included. A total of 112 patients were surveyed with one physical evaluation before the surgery and others 15, 30, 90, and 180 days after. Winged scapula was assessed with test proposed by Hoppenfeld. Shoulder range of motion (ROM) was assessed with goniometer for flexion, extension, adduction, abduction, internal rotation, and external rotation. A verbal scale from 0 to 10 was used to assess pain. Winged scapula incidence was 8.0 % 15 days after surgery. Two patients recovered from winged scapula 90 days after surgery and four more 180 days after surgery, while three patients still had winged scapula at this time. The incidence after 15 days from surgery was 20.9 and 22.6 % among patients submitted to sentinel node biopsy or axillary lymphadenectomy (AL), respectively (p < 0.01). There was no statistical difference of incidence according to breast surgery type. Operated side shoulder flexion, adduction, and abduction ROM changes were statistically different in patients with or without winged scapula. The mean reduction was higher in patients with winged scapula. Both groups showed the same pattern over time in pain. Winged scapula incidence was 8.0 % and was higher in AL, and prevalence decreased during 6 months after surgery. Patients who developed winged scapula had more shoulder flexion, adduction, and abduction limitation.

  10. Differences in wing melanization and pigmentation pattern in Drosophila buzzatii (Diptera: Drosophilidae under chemical stress Diferencias en la melanización y el patrón de pigmentación alar en Drosophila buzzatii (Diptera: Drosophilidae bajo estrés químico

    Directory of Open Access Journals (Sweden)

    Nicolás Mongiardino Koch

    2012-06-01

    Full Text Available Recently, the genetic basis and developmental mechanisms underlying the process of melanization have become progressively elucidated, allowing us to study the evolutionary processes that govern the huge variability of pigmentation observed in nature. However, environmental factors contributing to such variability have received little attention, even though they might have profound ecological consequences. Here we describe a method for analyzing the wing melanization patterns of drosophilids in both qualitative and quantitative ways. We test this method on wings of Drosophila buzzatii Patterson & Wheeler individuals, reared in control and alkaloid-enriched mediums. The alkaloids were extracted from the secondary host of these flies, Echinopsis terscheckii (Parm. ex Pfeiff. Friedrich & Rowley, and their effect on wing pigmentation is analyzed, bearing in mind the adverse effects that these compounds have on the development of D. buzzatii. Alkaloid-reared flies were shown to attain a higher degree of wing melanization, accompanied with differences in the spatial distribution of the pigment. Modifications at both metabolic and gene regulatory levels are proposed to explain the changes that chemical stressful conditions are shown to induce in this character. We propose the utilization of this procedure in studies of environmental dependence of pigmentation.Recientemente, hemos avanzado en nuestra comprensión de las bases genéticas y los mecanismos subyacentes al proceso de melanización durante el desarrollo, lo que permite el estudio de los procesos evolutivos que gobiernan la variabilidad de pigmentación presente en la naturaleza. Sin embargo, los factores ambientales que contribuyen a dicha variabilidad han recibido poca atención, aún cuando pueden conllevar profundas consecuencias a nivel ecológico. En este trabajo, presentamos un método para analizar los patrones de melanización alar de drosofílidos, de forma cualitativa y cuantitativa

  11. Notch inhibits Yorkie activity in Drosophila wing discs.

    Directory of Open Access Journals (Sweden)

    Alexandre Djiane

    Full Text Available During development, tissues and organs must coordinate growth and patterning so they reach the right size and shape. During larval stages, a dramatic increase in size and cell number of Drosophila wing imaginal discs is controlled by the action of several signaling pathways. Complex cross-talk between these pathways also pattern these discs to specify different regions with different fates and growth potentials. We show that the Notch signaling pathway is both required and sufficient to inhibit the activity of Yorkie (Yki, the Salvador/Warts/Hippo (SWH pathway terminal transcription activator, but only in the central regions of the wing disc, where the TEAD factor and Yki partner Scalloped (Sd is expressed. We show that this cross-talk between the Notch and SWH pathways is mediated, at least in part, by the Notch target and Sd partner Vestigial (Vg. We propose that, by altering the ratios between Yki, Sd and Vg, Notch pathway activation restricts the effects of Yki mediated transcription, therefore contributing to define a zone of low proliferation in the central wing discs.

  12. Qualitative skeletal correlates of wing shape in extant birds (Aves: Neoaves).

    Science.gov (United States)

    Hieronymus, Tobin L

    2015-02-27

    Among living fliers (birds, bats, and insects), birds display relatively high aspect ratios, a dimensionless shape variable that distinguishes long and narrow vs. short and broad wings. Increasing aspect ratio results in a functional tradeoff between low induced drag (efficient cruise) and increased wing inertia (difficult takeoff). Given the wide scope of its functional effects, the pattern of aspect ratio evolution is an important factor that contributes to the substantial ecological and phylogenetic diversity of living birds. However, because the feathers that define the wingtip (and hence wingspan and aspect ratio) often do not fossilize, resolution in the pattern of avian wing shape evolution is obscured by missing information. Here I use a comparative approach to investigate the relationship between skeletal proxies of flight feather attachment and wing shape. An accessory lobe of the internal index process of digit II-1, a bony correlate of distal primary attachment, shows weak but statistically significant relationships to aspect ratio and mass independent of other skeletal morphology. The dorsal phalangeal fossae of digit II-1, which house distal primaries VIII and IX, also show a trend of increased prominence with higher aspect ratio. Quill knobs on the ulna are examined concurrently, but do not show consistent signal with respect to wing shape. Although quill knobs are cited as skeletal correlates of flight performance in birds, their relationship to wing shape is inconsistent among extant taxa, and may reflect diverging selection pressures acting on a conserved architecture. In contrast, correlates of distal primary feather attachment on the major digit show convergent responses to increasing aspect ratio. In light of the diversity of musculoskeletal and integumentary mophology that underlies wing shape in different avian clades, it is unlikely that a single skeletal feature will show consistent predictive power across Neoaves. Confident inference of

  13. Analysis of bat wings for morphing

    Science.gov (United States)

    Leylek, Emily A.; Manzo, Justin E.; Garcia, Ephrahim

    2008-03-01

    The morphing of wings from three different bat species is studied using an extension of the Weissinger method. To understand how camber affects performance factors such as lift and lift to drag ratio, XFOIL is used to study thin (3% thickness to chord ratio) airfoils at a low Reynolds number of 100,000. The maximum camber of 9% yielded the largest lift coefficient, and a mid-range camber of 7% yielded the largest lift to drag ratio. Correlations between bat wing morphology and flight characteristics are covered, and the three bat wing planforms chosen represent various combinations of morphological components and different flight modes. The wings are studied using the extended Weissinger method in an "unmorphed" configuration using a thin, symmetric airfoil across the span of the wing through angles of attack of 0°-15°. The wings are then run in the Weissinger method at angles of attack of -2° to 12° in a "morphed" configuration modeled after bat wings seen in flight, where the camber of the airfoils comprising the wings is varied along the span and a twist distribution along the span is introduced. The morphed wing configurations increase the lift coefficient over 1000% from the unmorphed configuration and increase the lift to drag ratio over 175%. The results of the three different species correlate well with their flight in nature.

  14. Quad-thopter: Tailless Flapping Wing Robot with 4 Pairs of Wings

    NARCIS (Netherlands)

    de Wagter, C.; Karasek, M.; de Croon, G.C.H.E.; J.-M. Moschetta G. Hattenberger, H. de Plinval

    2017-01-01

    We present a novel design of a tailless flapping wing Micro Air Vehicle (MAV), which uses four independently driven pairs of flapping wings in order to fly and perform agile maneuvers. The wing pairs are arranged such that differential thrust generates the desired roll and pitch moments, similar to

  15. Adaptive wing : Investigations of passive wing technologies for loads reduction in the cleansky smart fixed wing aircraft (SFWA) project

    NARCIS (Netherlands)

    Kruger, W.R.; Dillinger, J; De Breuker, R.; Reyes, M.; Haydn, K.

    2016-01-01

    In the work package “Adaptive Wing” in the Clean-Sky “Smart Fixed Wing Aircraft” (SFWA) project, design processes and solutions for aircraft wings have been created, giving optimal response with respect to loads, comfort and performance by the introduction of passive and active concepts. Central

  16. The Realization and Study of Optical Wings

    Science.gov (United States)

    Artusio-Glimpse, Alexandra Brae

    Consider the airfoil: a carefully designed structure capable of stable lift in a uniform air flow. It so happens that air pressure and radiation (light) pressure are similar phenomena because each transfer momentum to flow-disturbing objects. This, then, begs the question: does an optical analogue to the airfoil exist? Though an exceedingly small effect, scientists harness radiation pressure in a wide gamut of applications from micromanipulation of single biological particles to the propulsion of large spacecrafts called solar sails. We introduce a cambered, refractive rod that is subjected to optical forces analogous to those seen in aerodynamics, and I call this analogue the optical wing. Flight characteristics of optical wings are determined by wing shape and material in a uniform radiation field. Theory predicts the lift force and axial torque are functions of the wing's angle of attack with stable and unstable orientations. These structures can operate as intensity-dependent, parametrically driven oscillators. In two-dimensions, the wings exhibit bistability when analyzed in an accelerating frame. In three-dimensions, the motion of axially symmetric spinning hemispherical wings is analogous to a spinning top. Experiments on semi-buoyant wings in water found semicylindrically shaped, refractive microparticles traversed a laser beam and rotated to an illumination-dependent stable orientation. Preliminary tests aid in the development of a calibrated force measurement experiment to directly evaluate the optical forces and torque on these samples. A foundational study of the optical wing, this work contributes to future advancements of flight-by-light.

  17. The Crest Wing Wave Energy Device

    DEFF Research Database (Denmark)

    Kofoed, Jens Peter; Antonishen, Michael Patrick

    This report presents the results of a continuation of an experimental study of the wave energy converting abilities of the Crest Wing wave energy converter (WEC), in the following referred to as ‘Phase 2'. The Crest Wing is a WEC that uses its movement in matching the shape of an oncoming wave...

  18. Effect of leading edge roundness on a delta wing in wing-rock motion

    Science.gov (United States)

    Ng, T. Terry; Malcolm, Gerald N.

    1990-01-01

    The effect of wing leading-edge roundness on wing rock was investigated using flow visualization in a water tunnel. Eighty degree delta wing models were tested on free-to-roll and forced oscillation rigs. The onset of wing rock was delayed by increasing the roundness of the leading edges. The wing rock amplitude and frequency results suggested that damping was increased at lower angles of attack but reduced at higher angles of attack. Vortex lift-off and vortex breakdown, especially during dynamic situations, were strongly affected by the leading edge roundness. Different forms of wing rock motion could be sustained by combinations of vortex breakdown and vortex lift-off. Behaviors of the wing and vortex motions were explained by the influence of leading edge roundness on the separation location, vortex trajectory, and vortex breakdown.

  19. Latitudinal gradient effect on the wing geometry of Auca coctei (Guérin(Lepidoptera, Nymphalidae

    Directory of Open Access Journals (Sweden)

    María-José Sanzana

    2013-12-01

    Full Text Available Latitudinal gradient effect on the wing geometry of Auca coctei (Guérin (Lepidoptera, Nymphalidae. When the environmental conditions change locally, the organisms and populations may also change in response to the selection pressure, so that the development of individuals may become affected in different degrees. There have been only a few studies in which the patterns of wing morphology variation have been looked into along a latitudinal gradient by means of geometric morphometrics. The aim of this work was to assess the morphologic differentiation of wing among butterfly populations of the species Auca coctei. For this purpose, 9 sampling locations were used which are representative of the distribution range of the butterfly and cover a wide latitudinal range in Chile. The wing morphology was studied in a total of 202 specimens of A. coctei (150 males and 52 females, based on digitization of 17 morphologic landmarks. The results show variation of wing shape in both sexes; however, for the centroid size there was significant variation only in females. Females show smaller centroid size at higher latitudes, therefore in this study the Bergmann reverse rule is confirmed for females of A. coctei. Our study extends morphologic projections with latitude, suggesting that wing variation is an environmental response from diverse origins and may influence different characteristics of the life history of a butterfly.

  20. Development of Bird-like Micro Aerial Vehicle with Flapping and Feathering Wing Motions

    Science.gov (United States)

    Maglasang, Jonathan; Goto, Norihiro; Isogai, Koji

    To investigate the feasibility of a highly efficient flapping system capable of avian maneuvers, such as rapid takeoff, hover and gliding, a full scale bird-like (ornithopter) flapping-wing micro aerial vehicle (MAV) shaped and patterned after a typical pigeon (Columba livia) has been designed and constructed. Both numerical and experimental methods have been used in the development of this vehicle. This flapping-wing micro aerial vehicle utilizes both the flapping and feathering motions of an avian wing by employing a novel flapping-feathering mechanism, which has been synthesized and constructed so as to best describe the properly coordinated flapping and feathering wing motions at phase angle difference of 90° in a horizontal steady level flight condition. This design allows high flapping and feathering amplitudes and is configurable for asymmetric wing motions which are desirable in high-speed flapping flight and maneuvering. The preliminary results indicate its viability as a practical and an efficient flapping-wing micro aerial vehicle.

  1. Novel method for measuring a dense 3D strain map of robotic flapping wings

    Science.gov (United States)

    Li, Beiwen; Zhang, Song

    2018-04-01

    Measuring dense 3D strain maps of the inextensible membranous flapping wings of robots is of vital importance to the field of bio-inspired engineering. Conventional high-speed 3D videography methods typically reconstruct the wing geometries through measuring sparse points with fiducial markers, and thus cannot obtain the full-field mechanics of the wings in detail. In this research, we propose a novel system to measure a dense strain map of inextensible membranous flapping wings by developing a superfast 3D imaging system and a computational framework for strain analysis. Specifically, first we developed a 5000 Hz 3D imaging system based on the digital fringe projection technique using the defocused binary patterns to precisely measure the dynamic 3D geometries of rapidly flapping wings. Then, we developed a geometry-based algorithm to perform point tracking on the precisely measured 3D surface data. Finally, we developed a dense strain computational method using the Kirchhoff-Love shell theory. Experiments demonstrate that our method can effectively perform point tracking and measure a highly dense strain map of the wings without many fiducial markers.

  2. Wing geometry of Culex coronator (Diptera: Culicidae) from South and Southeast Brazil.

    Science.gov (United States)

    Demari-Silva, Bruna; Suesdek, Lincoln; Sallum, Maria Anice Mureb; Marrelli, Mauro Toledo

    2014-04-09

    The Coronator Group encompasses Culex coronator Dyar & Knab, Culex camposi Dyar, Culex covagarciai Forattini, Culex ousqua Dyar, Culex usquatissimus Dyar, Culex usquatus Dyar and Culex yojoae Strickman. Culex coronator has the largest geographic distribution, occurring in North, Central and South America. Moreover, it is a potential vector-borne mosquito species because females have been found naturally infected with several arboviruses, i.e., Saint Louis Encephalitis Virus, Venezuelan Equine Encephalitis Virus and West Nile Virus. Considering the epidemiological importance of Cx. coronator, we investigated the wing shape diversity of Cx. coronator from South and Southeast Brazil, a method to preliminarily estimate population diversity. Field-collected immature stages of seven populations from a large geographical area in Brazil were maintained in the laboratory to obtain both females and males linked with pupal and/or larval exuviae. For each individual female, 18 landmarks of left wings were marked and digitalized. After Procrustes superimposition, discriminant analysis of shape was employed to quantify wing shape variation among populations. The isometric estimator centroid size was calculated to assess the overall wing size and allometry. Wing shape was polymorphic among populations of Cx. coronator. However, dissimilarities among populations were higher than those observed within each population, suggesting populational differentiation in Cx. coronator. Morphological distances between populations were not correlated to geographical distances, indicating that other factors may act on wing shape and thus, determining microevolutionary patterns in Cx. coronator. Despite the population differentiation, intrapopulational wing shape variability was equivalent among all seven populations. The wing variability found in Cx. coronator populations brings to light a new biological problem to be investigated: the population genetics of Cx. coronator. Because of differences

  3. Hovering hummingbird wing aerodynamics during the annual cycle. II. Implications of wing feather moult

    Science.gov (United States)

    Sapir, Nir; Elimelech, Yossef

    2018-01-01

    Birds usually moult their feathers in a particular sequence which may incur aerodynamic, physiological and behavioural implications. Among birds, hummingbirds are unique species in their sustained hovering flight. Because hummingbirds frequently hover-feed, they must maintain sufficiently high flight capacities even when moulting their flight feathers. A hummingbird wing consists of 10 primary flight feathers whose absence during moult may strongly affect wing performance. Using dynamic similarity rules, we compared time-accurate aerodynamic loads and flow field measurements over several wing geometries that follow the natural feather moult sequence of Calypte anna, a common hummingbird species in western North America. Our results suggest a drop of more than 20% in lift production during the early stages of the moult sequence in which mid-wing flight feathers are moulted. We also found that the wing's ability to generate lift strongly depended on the morphological integrity of the outer primaries and leading-edge. These findings may explain the evolution of wing morphology and moult attributes. Specifically, the high overlap between adjacent wing feathers, especially at the wing tip, and the slow sequential replacement of the wing feathers result in a relatively small reduction in wing surface area during moult with limited aerodynamic implications. We present power and efficiency analyses for hover flight during moult under several plausible scenarios, suggesting that body mass reduction could be a compensatory mechanism that preserves the energetic costs of hover flight. PMID:29515884

  4. Spectrally tuned structural and pigmentary coloration of birdwing butterfly wing scales

    Science.gov (United States)

    Wilts, Bodo D.; Matsushita, Atsuko; Arikawa, Kentaro; Stavenga, Doekele G.

    2015-01-01

    The colourful wing patterns of butterflies play an important role for enhancing fitness; for instance, by providing camouflage, for interspecific mate recognition, or for aposematic display. Closely related butterfly species can have dramatically different wing patterns. The phenomenon is assumed to be caused by ecological processes with changing conditions, e.g. in the environment, and also by sexual selection. Here, we investigate the birdwing butterflies, Ornithoptera, the largest butterflies of the world, together forming a small genus in the butterfly family Papilionidae. The wings of these butterflies are marked by strongly coloured patches. The colours are caused by specially structured wing scales, which act as a chirped multilayer reflector, but the scales also contain papiliochrome pigments, which act as a spectral filter. The combined structural and pigmentary effects tune the coloration of the wing scales. The tuned colours are presumably important for mate recognition and signalling. By applying electron microscopy, (micro-)spectrophotometry and scatterometry we found that the various mechanisms of scale coloration of the different birdwing species strongly correlate with the taxonomical distribution of Ornithoptera species. PMID:26446560

  5. Beneficial aerodynamic effect of wing scales on the climbing flight of butterflies.

    Science.gov (United States)

    Slegers, Nathan; Heilman, Michael; Cranford, Jacob; Lang, Amy; Yoder, John; Habegger, Maria Laura

    2017-01-30

    It is hypothesized that butterfly wing scale geometry and surface patterning may function to improve aerodynamic efficiency. In order to investigate this hypothesis, a method to measure butterfly flapping kinematics optically over long uninhibited flapping sequences was developed. Statistical results for the climbing flight flapping kinematics of 11 butterflies, based on a total of 236 individual flights, both with and without their wing scales, are presented. Results show, that for each of the 11 butterflies, the mean climbing efficiency decreased after scales were removed. Data was reduced to a single set of differences of climbing efficiency using are paired t-test. Results show a mean decrease in climbing efficiency of 32.2% occurred with a 95% confidence interval of 45.6%-18.8%. Similar analysis showed that the flapping amplitude decreased by 7% while the flapping frequency did not show a significant difference. Results provide strong evidence that butterfly wing scale geometry and surface patterning improve butterfly climbing efficiency. The authors hypothesize that the wing scale's effect in measured climbing efficiency may be due to an improved aerodynamic efficiency of the butterfly and could similarly be used on flapping wing micro air vehicles to potentially achieve similar gains in efficiency.

  6. Semi-automated quantitative Drosophila wings measurements.

    Science.gov (United States)

    Loh, Sheng Yang Michael; Ogawa, Yoshitaka; Kawana, Sara; Tamura, Koichiro; Lee, Hwee Kuan

    2017-06-28

    Drosophila melanogaster is an important organism used in many fields of biological research such as genetics and developmental biology. Drosophila wings have been widely used to study the genetics of development, morphometrics and evolution. Therefore there is much interest in quantifying wing structures of Drosophila. Advancement in technology has increased the ease in which images of Drosophila can be acquired. However such studies have been limited by the slow and tedious process of acquiring phenotypic data. We have developed a system that automatically detects and measures key points and vein segments on a Drosophila wing. Key points are detected by performing image transformations and template matching on Drosophila wing images while vein segments are detected using an Active Contour algorithm. The accuracy of our key point detection was compared against key point annotations of users. We also performed key point detection using different training data sets of Drosophila wing images. We compared our software with an existing automated image analysis system for Drosophila wings and showed that our system performs better than the state of the art. Vein segments were manually measured and compared against the measurements obtained from our system. Our system was able to detect specific key points and vein segments from Drosophila wing images with high accuracy.

  7. Novel Control Effectors for Truss Braced Wing

    Science.gov (United States)

    White, Edward V.; Kapania, Rakesh K.; Joshi, Shiv

    2015-01-01

    At cruise flight conditions very high aspect ratio/low sweep truss braced wings (TBW) may be subject to design requirements that distinguish them from more highly swept cantilevered wings. High aspect ratio, short chord length and relative thinness of the airfoil sections all contribute to relatively low wing torsional stiffness. This may lead to aeroelastic issues such as aileron reversal and low flutter margins. In order to counteract these issues, high aspect ratio/low sweep wings may need to carry additional high speed control effectors to operate when outboard ailerons are in reversal and/or must carry additional structural weight to enhance torsional stiffness. The novel control effector evaluated in this study is a variable sweep raked wing tip with an aileron control surface. Forward sweep of the tip allows the aileron to align closely with the torsional axis of the wing and operate in a conventional fashion. Aft sweep of the tip creates a large moment arm from the aileron to the wing torsional axis greatly enhancing aileron reversal. The novelty comes from using this enhanced and controllable aileron reversal effect to provide roll control authority by acting as a servo tab and providing roll control through intentional twist of the wing. In this case the reduced torsional stiffness of the wing becomes an advantage to be exploited. The study results show that the novel control effector concept does provide roll control as described, but only for a restricted class of TBW aircraft configurations. For the configuration studied (long range, dual aisle, Mach 0.85 cruise) the novel control effector provides significant benefits including up to 12% reduction in fuel burn.

  8. Nonlinear Structures Optimization for Flexible Flapping Wing MAVs

    Science.gov (United States)

    2009-02-01

    nonlinear optimization, flapping wing, fluid structure interaction, micro -air vehicles, flexible wing, flapping mechanism 16. SECURITY... Structures Optimization for Flexible Flapping Wing Micro -Air Vehicles” was funded with Chief Scientist Innovative Research funds. This project was divided...predict a 10% resisting load to the model, and Python Scripting to wrap around everything. 2 Building the Model in Abaqus CAE The flapping wing

  9. Generic Wing-Body Aerodynamics Data Base

    Science.gov (United States)

    Holst, Terry L.; Olsen, Thomas H.; Kwak, Dochan (Technical Monitor)

    2001-01-01

    The wing-body aerodynamics data base consists of a series of CFD (Computational Fluid Dynamics) simulations about a generic wing body configuration consisting of a ogive-circular-cylinder fuselage and a simple symmetric wing mid-mounted on the fuselage. Solutions have been obtained for Nonlinear Potential (P), Euler (E) and Navier-Stokes (N) solvers over a range of subsonic and transonic Mach numbers and angles of attack. In addition, each solution has been computed on a series of grids, coarse, medium and fine to permit an assessment of grid refinement errors.

  10. Unemployment and Right-Wing Extremist Crime

    OpenAIRE

    Falk, Armin; Zweimüller, Josef

    2005-01-01

    Right-wing extremism is a serious problem in many societies. A prominent hypothesis states that unemployment plays a crucial role for the occurrence of right-wing extremist crime. In this paper we empirically test this hypothesis. We use a previously not used data set which includes all officially recorded right-wing criminal acts in Germany. These data are recorded by the German Federal Criminal Police Office on a monthly and state level basis. Our main finding is that there is in fact a sig...

  11. The wings of Bombyx mori develop from larval discs exhibiting an ...

    Indian Academy of Sciences (India)

    Unknown

    field and French 1999). We present here the expression pattern of Nubbin (Nub), Wingless (Wg) and Distal-less. (Dll) in B. mori wing discs using antibodies generated against the corresponding proteins from Drosophila or the butterfly P. coenia. The expression of Wg and Dll as overlapping domains in the distal region in B.

  12. The leading-edge vortex of swift-wing shaped delta wings

    Science.gov (United States)

    Muir, Rowan; Arredondo-Galeana, Abel; Viola, Ignazio Maria

    2017-11-01

    Recent investigations on the aerodynamics of natural fliers have illuminated the significance of the Leading-Edge Vortex (LEV) for lift generation in a variety of flight conditions. In this investigation, a model non-slender delta shaped wing with a sharp leading-edge is tested at low Reynolds Number, along with a delta wing of the same design, but with a modified trailing edge inspired by the wing of a common swift Apus apus. The effect of the tapering swift wing on LEV development and stability is compared with the flow structure over the un-modified delta wing model through particle image velocimetry. For the first time, a leading-edge vortex system consisting of a dual or triple LEV is recorded on a swift-wing shaped delta wing, where such a system is found across all tested conditions. It is shown that the spanwise location of LEV breakdown is governed by the local chord rather than Reynolds Number or angle of attack. These findings suggest that the trailing-edge geometry of the swift wing alone does not prevent the common swift from generating an LEV system comparable with that of a delta shaped wing. This work received funding from the Engineering and Physical Sciences Research Council [EP/M506515/1] and the Consejo Nacional de Ciencia y Tecnología (CONACYT).

  13. Fruit fly scale robots can hover longer with flapping wings than with spinning wings.

    Science.gov (United States)

    Hawkes, Elliot W; Lentink, David

    2016-10-01

    Hovering flies generate exceptionally high lift, because their wings generate a stable leading edge vortex. Micro flying robots with a similar wing design can generate similar high lift by either flapping or spinning their wings. While it requires less power to spin a wing, the overall efficiency depends also on the actuator system driving the wing. Here, we present the first holistic analysis to calculate how long a fly-inspired micro robot can hover with flapping versus spinning wings across scales. We integrate aerodynamic data with data-driven scaling laws for actuator, electronics and mechanism performance from fruit fly to hummingbird scales. Our analysis finds that spinning wings driven by rotary actuators are superior for robots with wingspans similar to hummingbirds, yet flapping wings driven by oscillatory actuators are superior at fruit fly scale. This crossover is driven by the reduction in performance of rotary compared with oscillatory actuators at smaller scale. Our calculations emphasize that a systems-level analysis is essential for trading-off flapping versus spinning wings for micro flying robots. © 2016 The Author(s).

  14. Flow Modulation and Force Control of Flapping Wings

    Science.gov (United States)

    2014-10-29

    tested on a flapping wing model in the oil tank. Robotic flapper equipped with DC motors drove the wing model, and the imbedded servo motor could flap...the overall wake structure on the hovering wings. Totally, two volumetric flow measurements were performed on two mechanical flappers with different...wing kinematics but similar wing geometry. On the flappers with small stroke angle and passive rotation, the general vortex wake structure

  15. Left-Wing Extremism: The Current Threat

    Energy Technology Data Exchange (ETDEWEB)

    Karl A. Seger

    2001-04-30

    Left-wing extremism is ''alive and well'' both in the US and internationally. Although the current domestic terrorist threat within the U. S. is focused on right-wing extremists, left-wing extremists are also active and have several objectives. Leftist extremists also pose an espionage threat to U.S. interests. While the threat to the U.S. government from leftist extremists has decreased in the past decade, it has not disappeared. There are individuals and organizations within the U.S. who maintain the same ideology that resulted in the growth of left-wing terrorism in this country in the 1970s and 1980s. Some of the leaders from that era are still communicating from Cuba with their followers in the U.S., and new leaders and groups are emerging.

  16. A galactic microquasar mimicking winged radio galaxies.

    Science.gov (United States)

    Martí, Josep; Luque-Escamilla, Pedro L; Bosch-Ramon, Valentí; Paredes, Josep M

    2017-11-24

    A subclass of extragalactic radio sources known as winged radio galaxies has puzzled astronomers for many years. The wing features are detected at radio wavelengths as low-surface-brightness radio lobes that are clearly misaligned with respect to the main lobe axis. Different models compete to account for these peculiar structures. Here, we report observational evidence that the parsec-scale radio jets in the Galactic microquasar GRS 1758-258 give rise to a Z-shaped radio emission strongly reminiscent of the X and Z-shaped morphologies found in winged radio galaxies. This is the first time that such extended emission features are observed in a microquasar, providing a new analogy for its extragalactic relatives. From our observations, we can clearly favour the hydrodynamic backflow interpretation against other possible wing formation scenarios. Assuming that physical processes are similar, we can extrapolate this conclusion and suggest that this mechanism could also be at work in many extragalactic cases.

  17. Iridescence and spectral filtering of the gyroid-type photonic crystals in Parides sesostris wing scales

    Science.gov (United States)

    Wilts, Bodo D.; Michielsen, Kristel; De Raedt, Hans; Stavenga, Doekele G.

    2012-01-01

    The cover scales on the wing of the Emerald-patched Cattleheart butterfly, Parides sesostris, contain gyroid-type biological photonic crystals that brightly reflect green light. A pigment, which absorbs maximally at approximately 395 nm, is immersed predominantly throughout the elaborate upper lamina. This pigment acts as a long-pass filter shaping the reflectance spectrum of the underlying photonic crystals. The additional effect of the filtering is that the spatial distribution of the scale reflectance is approximately angle-independent, leading to a stable wing pattern contrast. The spectral tuning of the original reflectance is verified by photonic band structure modelling. PMID:24098853

  18. Does skipping a meal matter to a butterfly's appearance? Effects of larval food stress on wing morphology and color in monarch butterflies.

    Directory of Open Access Journals (Sweden)

    Haley Johnson

    Full Text Available In animals with complex life cycles, all resources needed to form adult tissues are procured at the larval stage. For butterflies, the proper development of wings involves synthesizing tissue during metamorphosis based on the raw materials obtained by larvae. Similarly, manufacture of pigment for wing scales also requires resources acquired by larvae. We conducted an experiment to test the effects of food deprivation in the larval stage on multiple measures of adult wing morphology and coloration of monarch butterflies (Danaus plexippus, a species in which long-distance migration makes flight efficiency critical. In a captive setting, we restricted food (milkweed from late-stage larvae for either 24 hrs or 48 hrs, then after metamorphosis we used image analysis methods to measure forewing surface area and elongation (length/width, which are both important for migration. We also measured the brightness of orange pigment and the intensity of black on the wing. There were correlations between several wing features, including an unexpected association between wing elongation and melanism, which will require further study to fully understand. The clearest effect of food restriction was a reduction in adult wing size in the high stress group (by approximately 2%. Patterns observed for other wing traits were ambiguous: monarchs in the low stress group (but not the high had less elongated and paler orange pigmentation. There was no effect on wing melanism. Although some patterns obtained in this study were unclear, our results concerning wing size have direct bearing on the monarch migration. We show that if milkweed is limited for monarch larvae, their wings become stunted, which could ultimately result in lower migration success.

  19. The leading-edge vortex of swift wing-shaped delta wings.

    Science.gov (United States)

    Muir, Rowan Eveline; Arredondo-Galeana, Abel; Viola, Ignazio Maria

    2017-08-01

    Recent investigations on the aerodynamics of natural fliers have illuminated the significance of the leading-edge vortex (LEV) for lift generation in a variety of flight conditions. A well-documented example of an LEV is that generated by aircraft with highly swept, delta-shaped wings. While the wing aerodynamics of a manoeuvring aircraft, a bird gliding and a bird in flapping flight vary significantly, it is believed that this existing knowledge can serve to add understanding to the complex aerodynamics of natural fliers. In this investigation, a model non-slender delta-shaped wing with a sharp leading edge is tested at low Reynolds number, along with a delta wing of the same design, but with a modified trailing edge inspired by the wing of a common swift Apus apus . The effect of the tapering swift wing on LEV development and stability is compared with the flow structure over the unmodified delta wing model through particle image velocimetry. For the first time, a leading-edge vortex system consisting of a dual or triple LEV is recorded on a swift wing-shaped delta wing, where such a system is found across all tested conditions. It is shown that the spanwise location of LEV breakdown is governed by the local chord rather than Reynolds number or angle of attack. These findings suggest that the trailing-edge geometry of the swift wing alone does not prevent the common swift from generating an LEV system comparable with that of a delta-shaped wing.

  20. Numerical and experimental investigations on unsteady aerodynamics of flapping wings

    Science.gov (United States)

    Yu, Meilin

    suggestions to the design of micro-air-vehicles (MAVs), 3D simulations of the flapping wings are carried out in this work. Both the rectangular and bio-inspired wings with different kinematics are investigated. The formation process of two-jet-like wake patterns behind the finite-span flapping wing is found to be closely related to the interaction between trailing edge vortices and tip vortices. Then the effects of the wing planforms on the aerodynamics performance of the finite-span flapping wings are elucidated in terms of the evolution and dynamic interaction of unsteady vortex structures.

  1. Dpp spreading is required for medial but not for lateral wing disc growth.

    Science.gov (United States)

    Harmansa, Stefan; Hamaratoglu, Fisun; Affolter, Markus; Caussinus, Emmanuel

    2015-11-19

    Drosophila Decapentaplegic (Dpp) has served as a paradigm to study morphogen-dependent growth control. However, the role of a Dpp gradient in tissue growth remains highly controversial. Two fundamentally different models have been proposed: the 'temporal rule' model suggests that all cells of the wing imaginal disc divide upon a 50% increase in Dpp signalling, whereas the 'growth equalization model' suggests that Dpp is only essential for proliferation control of the central cells. Here, to discriminate between these two models, we generated and used morphotrap, a membrane-tethered anti-green fluorescent protein (GFP) nanobody, which enables immobilization of enhanced (e)GFP::Dpp on the cell surface, thereby abolishing Dpp gradient formation. We find that in the absence of Dpp spreading, wing disc patterning is lost; however, lateral cells still divide at normal rates. These data are consistent with the growth equalization model, but do not fit a global temporal rule model in the wing imaginal disc.

  2. Morpho morphometrics: Shared ancestry and selection drive the evolution of wing size and shape in Morpho butterflies.

    Science.gov (United States)

    Chazot, Nicolas; Panara, Stephen; Zilbermann, Nicolas; Blandin, Patrick; Le Poul, Yann; Cornette, Raphaël; Elias, Marianne; Debat, Vincent

    2016-01-01

    Butterfly wings harbor highly diverse phenotypes and are involved in many functions. Wing size and shape result from interactions between adaptive processes, phylogenetic history, and developmental constraints, which are complex to disentangle. Here, we focus on the genus Morpho (Nymphalidae: Satyrinae, 30 species), which presents a high diversity of sizes, shapes, and color patterns. First, we generate a comprehensive molecular phylogeny of these 30 species. Next, using 911 collection specimens, we quantify the variation of wing size and shape across species, to assess the importance of shared ancestry, microhabitat use, and sexual selection in the evolution of the wings. While accounting for phylogenetic and allometric effects, we detect a significant difference in wing shape but not size among microhabitats. Fore and hindwings covary at the individual and species levels, and the covariation differs among microhabitats. However, the microhabitat structure in covariation disappears when phylogenetic relationships are taken into account. Our results demonstrate that microhabitat has driven wing shape evolution, although it has not strongly affected forewing and hindwing integration. We also found that sexual dimorphism of forewing shape and color pattern are coupled, suggesting a common selective force. © 2015 The Author(s). Evolution © 2015 The Society for the Study of Evolution.

  3. Aeroelasticity of morphing wings using neural networks

    Science.gov (United States)

    Natarajan, Anand

    In this dissertation, neural networks are designed to effectively model static non-linear aeroelastic problems in adaptive structures and linear dynamic aeroelastic systems with time varying stiffness. The use of adaptive materials in aircraft wings allows for the change of the contour or the configuration of a wing (morphing) in flight. The use of smart materials, to accomplish these deformations, can imply that the stiffness of the wing with a morphing contour changes as the contour changes. For a rapidly oscillating body in a fluid field, continuously adapting structural parameters may render the wing to behave as a time variant system. Even the internal spars/ribs of the aircraft wing which define the wing stiffness can be made adaptive, that is, their stiffness can be made to vary with time. The immediate effect on the structural dynamics of the wing, is that, the wing motion is governed by a differential equation with time varying coefficients. The study of this concept of a time varying torsional stiffness, made possible by the use of active materials and adaptive spars, in the dynamic aeroelastic behavior of an adaptable airfoil is performed here. Another type of aeroelastic problem of an adaptive structure that is investigated here, is the shape control of an adaptive bump situated on the leading edge of an airfoil. Such a bump is useful in achieving flow separation control for lateral directional maneuverability of the aircraft. Since actuators are being used to create this bump on the wing surface, the energy required to do so needs to be minimized. The adverse pressure drag as a result of this bump needs to be controlled so that the loss in lift over the wing is made minimal. The design of such a "spoiler bump" on the surface of the airfoil is an optimization problem of maximizing pressure drag due to flow separation while minimizing the loss in lift and energy required to deform the bump. One neural network is trained using the CFD code FLUENT to

  4. Hovering hummingbird wing aerodynamics during the annual cycle. I. Complete wing.

    Science.gov (United States)

    Achache, Yonathan; Sapir, Nir; Elimelech, Yossef

    2017-08-01

    The diverse hummingbird family (Trochilidae) has unique adaptations for nectarivory, among which is the ability to sustain hover-feeding. As hummingbirds mainly feed while hovering, it is crucial to maintain this ability throughout the annual cycle-especially during flight-feather moult, in which wing area is reduced. To quantify the aerodynamic characteristics and flow mechanisms of a hummingbird wing throughout the annual cycle, time-accurate aerodynamic loads and flow field measurements were correlated over a dynamically scaled wing model of Anna's hummingbird ( Calypte anna ). We present measurements recorded over a model of a complete wing to evaluate the baseline aerodynamic characteristics and flow mechanisms. We found that the vorticity concentration that had developed from the wing's leading-edge differs from the attached vorticity structure that was typically found over insects' wings; firstly, it is more elongated along the wing chord, and secondly, it encounters high levels of fluctuations rather than a steady vortex. Lift characteristics resemble those of insects; however, a 20% increase in the lift-to-torque ratio was obtained for the hummingbird wing model. Time-accurate aerodynamic loads were also used to evaluate the time-evolution of the specific power required from the flight muscles, and the overall wingbeat power requirements nicely matched previous studies.

  5. The effects of wing flexibility on the flight performance and stability of flapping wing micro air vehicles

    Science.gov (United States)

    Bluman, James Edward

    Insect wings are flexible. However, the influence of wing flexibility on the flight dynamics of insects and flapping wing micro air vehicles is unknown. Most studies in the literature consider rigid wings and conclude that the hover equilibrium is unstable. This dissertation shows that a flapping wing flyer with flexible wings exhibits stable natural modes of the open loop system in hover, never reported before. The free-flight insect flight dynamics is modeled for both flexible and rigid wings. Wing mass and inertia are included in the nonlinear equations of motion. The flapping wing aerodynamics are modeled using a quasi-steady model, a well-validated two dimensional Navier Stokes model, and a coupled, two dimensional Navier Stokes - Euler Bernoulli beam model that accurately models the fluid-structure interaction of flexible wings. Hover equilibrium is systematically and efficiently determined with a coupled quasi-steady and Navier-Stokes equation trimmer. The power and stability are reported at hover while parametrically varying the pitch axis location for rigid wings and the structural stiffness for flexible wings. The results indicate that the rigid wings possess an unstable oscillatory mode mainly due to their pitch sensitivity to horizontal velocity perturbations. The flexible wings stabilize this mode primarily by adjusting their wing shape in the presence of perturbations. The wing's response to perturbations generates significantly more horizontal velocity damping and pitch rate damping than in rigid wings. Furthermore, the flexible wings experience substantially less wing wake interaction, which, for rigid wings, is destabilizing. The power required to hover a fruit fly with actively rotating rigid wings varies between 16.9 and 34.2 W/kg. The optimal power occurs when the pitch axis is located at 30% chord, similar to some biological observations. Flexible wings require 23.1 to 38.5 W/kg. However, flexible wings exhibit more stable system dynamics and

  6. [Wing 1 radiation survey and contamination report

    International Nuclear Information System (INIS)

    Olsen, K.

    1991-01-01

    We have completed the 5480.11 survey for Wing 1. All area(s)/item(s) requested by the 5480.11 committee have been thoroughly surveyed and documented. Decontamination/disposal of contaminated items has been accomplished. The wing 1 survey was started on 8/13/90 and completed 9/18/90. However, the follow-up surveys were not completed until 2/18/91. We received the final set of smear samples for wing 1 on 1/13/91. A total of 5,495 smears were taken from wing 1 and total of 465 smears were taken during the follow-up surveys. There were a total 122 items found to have fixed contamination and 4 items with smearable contamination in excess of the limits specified in DOE ORDER 5480.11 (AR 3-7). The following area(s)/item(s) were not included in the 5480.11 survey: Hallways, Access panels, Men's and women's change rooms, Janitor closets, Wall lockers and item(s) stored in wing 1 hallways and room 1116. If our contract is renewed, we will include those areas in our survey according to your request of April 15, 1991

  7. Flapping wing aerodynamics: from insects to vertebrates.

    Science.gov (United States)

    Chin, Diana D; Lentink, David

    2016-04-01

    More than a million insects and approximately 11,000 vertebrates utilize flapping wings to fly. However, flapping flight has only been studied in a few of these species, so many challenges remain in understanding this form of locomotion. Five key aerodynamic mechanisms have been identified for insect flight. Among these is the leading edge vortex, which is a convergent solution to avoid stall for insects, bats and birds. The roles of the other mechanisms - added mass, clap and fling, rotational circulation and wing-wake interactions - have not yet been thoroughly studied in the context of vertebrate flight. Further challenges to understanding bat and bird flight are posed by the complex, dynamic wing morphologies of these species and the more turbulent airflow generated by their wings compared with that observed during insect flight. Nevertheless, three dimensionless numbers that combine key flow, morphological and kinematic parameters - the Reynolds number, Rossby number and advance ratio - govern flapping wing aerodynamics for both insects and vertebrates. These numbers can thus be used to organize an integrative framework for studying and comparing animal flapping flight. Here, we provide a roadmap for developing such a framework, highlighting the aerodynamic mechanisms that remain to be quantified and compared across species. Ultimately, incorporating complex flight maneuvers, environmental effects and developmental stages into this framework will also be essential to advancing our understanding of the biomechanics, movement ecology and evolution of animal flight. © 2016. Published by The Company of Biologists Ltd.

  8. Principle of bio-inspired insect wing rotational hinge design

    Science.gov (United States)

    Fei, Fan

    A principle for designing and fabricating bio-inspired miniature artificial insect flapping wing using flexure rotational hinge design is presented. A systematic approach of selecting rotational hinge stiffness value is proposed. Based on the understanding of flapping wing aerodynamics, a dynamic simulation is constructed using the established quasi-steady model and the wing design. Simulations were performed to gain insight on how different parameters affect the wing rotational response. Based on system resonance a model to predict the optimal rotational hinge stiffness based on given wing parameter and flapping wing kinematic is proposed. By varying different wing parameters, the proposed method is shown to be applicable to a wide range of wing designs with different sizes and shapes. With the selected hinge stiffness value, aspects of the rotational joint design is discussed and an integrated wing-hinge structure design using laminated carbon fiber and polymer film is presented. Manufacturing process of such composite structure is developed to achieve high accuracy and repeatability. The yielded hinge stiffness is verified by measurements. To validate the proposed model, flapping wing experiments were conducted. A flapping actuation set up is built using DC motor and a controller is implemented on a microcontroller to track desired wing stroke kinematic. Wing stroke and rotation kinematic were extracted using a high speed camera and the lift generation is evaluated. A total of 49 flapping experiments were presented, experimental data shows good correlation with the model's prediction. With the wing rotational hinge stiffness designed so that the rotational resonant frequency is twice as the stroke frequency, the resulting wing rotation generates near optimal lift. With further simulation, the proposed model shows low sensitivity to wing parameter variation. As a result, giving a design parameter of a flapping wing robot platform, the proposed principle can

  9. Gliding swifts attain laminar flow over rough wings.

    Directory of Open Access Journals (Sweden)

    David Lentink

    Full Text Available Swifts are among the most aerodynamically refined gliding birds. However, the overlapping vanes and protruding shafts of their primary feathers make swift wings remarkably rough for their size. Wing roughness height is 1-2% of chord length on the upper surface--10,000 times rougher than sailplane wings. Sailplanes depend on extreme wing smoothness to increase the area of laminar flow on the wing surface and minimize drag for extended glides. To understand why the swift does not rely on smooth wings, we used a stethoscope to map laminar flow over preserved wings in a low-turbulence wind tunnel. By combining laminar area, lift, and drag measurements, we show that average area of laminar flow on swift wings is 69% (n = 3; std 13% of their total area during glides that maximize flight distance and duration--similar to high-performance sailplanes. Our aerodynamic analysis indicates that swifts attain laminar flow over their rough wings because their wing size is comparable to the distance the air travels (after a roughness-induced perturbation before it transitions from laminar to turbulent. To interpret the function of swift wing roughness, we simulated its effect on smooth model wings using physical models. This manipulation shows that laminar flow is reduced and drag increased at high speeds. At the speeds at which swifts cruise, however, swift-like roughness prolongs laminar flow and reduces drag. This feature gives small birds with rudimentary wings an edge during the evolution of glide performance.

  10. Investigating the Force Production of Functionally-Graded Flexible Wings in Flapping Wing Flight

    Science.gov (United States)

    Mudbhari, Durlav; Erdogan, Malcolm; He, Kai; Bateman, Daniel; Lipkis, Rory; Moored, Keith

    2015-11-01

    Birds, insects and bats oscillate their wings to propel themselves over long distances and to maneuver with unprecedented agility. A key element to achieve their impressive aerodynamic performance is the flexibility of their wings. Numerous studies have shown that homogeneously flexible wings can enhance force production, propulsive efficiency and lift efficiency. Yet, animal wings are not homogenously flexible, but instead have varying material properties. The aim of this study is to characterize the force production and energetics of functionally-graded flexible wings. A partially-flexible wing composed of a rigid section and a flexible section is used as a first-order model of functionally-graded materials. The flexion occurs in the spanwise direction and it is affected by the spanwise flexion ratio, that is, the ratio of the length of the rigid section compared to the total span length. By varying the flexion ratio as well as the material properties of the flexible section, the study aims to examine the force production and energetics of flapping flight with functionally-graded flexible wings. Supported by the Office of Naval Research under Program Director Dr. Bob Brizzolara, MURI grant number N00014-14-1-0533.

  11. Aeroelastic Analysis of Modern Complex Wings

    Science.gov (United States)

    Kapania, Rakesh K.; Bhardwaj, Manoj K.; Reichenbach, Eric; Guruswamy, Guru P.

    1996-01-01

    A process is presented by which aeroelastic analysis is performed by using an advanced computational fluid dynamics (CFD) code coupled with an advanced computational structural dynamics (CSD) code. The process is demonstrated on an F/A-18 Stabilator using NASTD (an in-house McDonnell Douglas Aerospace East CFD code) coupled with NASTRAN. The process is also demonstrated on an aeroelastic research wing (ARW-2) using ENSAERO (an in-house NASA Ames Research Center CFD code) coupled with a finite element wing-box structures code. Good results have been obtained for the F/A-18 Stabilator while results for the ARW-2 supercritical wing are still being obtained.

  12. Active Twist Control for a Compliant Wing Structure, Phase I

    Data.gov (United States)

    National Aeronautics and Space Administration — Blended wing body (BWB) aircraft provide an aerodynamically superior solution over traditional tube-and-wing designs for a number of mission profiles. These...

  13. Fixed-Wing Micro Air Vehicles with Hovering Capabilities

    National Research Council Canada - National Science Library

    Bataille, Boris; Poinsot, Damien; Thipyopas, Chinnapat; Moschetta, Jean-Marc

    2007-01-01

    Fixed-wing micro air vehicles (MAV) are very attractive for outdoor surveillance missions since they generally offer better payload and endurance capabilities than rotorcraft or flapping-wing vehicles of equal size...

  14. Nonlinear Dynamics of Wind Turbine Wings

    DEFF Research Database (Denmark)

    Larsen, Jesper Winther

    Wind turbines with a nominal effect of 5MW with a rotor diameter of up to 126m are produced today. With the increasing size wind turbines also become more and more optimized with respect to structural dimensions and material usage, without increasing the stiffness proportionally. Consequently......, large wind turbines become increasingly flexible and dynamically sensitive. This project focuses on the structural analysis of highly flexible wind turbine wings, and the aerodynamic loading of wind turbine wings under large changes in flow field due to elastic deformations and changing wind conditions....

  15. Stability and transition on swept wings

    Science.gov (United States)

    Stuckert, Greg; Herbert, Thorwald; Esfahanian, Vahid

    1993-01-01

    This paper describes the extension and application of the Parabolized Stability Equations (PSE) to the stability and transition of the supersonic three-dimensional laminar boundary layer on a swept wing. The problem formulation uses a general coordinate transformation for arbitrary curvilinear body-fitted computational grids. Some testing using these coordinates is briefly described to help validate the software used for the investigation. The disturbance amplitude ratios as a function of chord position for supersonic (Mach 1.5) boundary layers on untapered, untwisted wings of different sweep angles are then presented and compared with those obtained from local parallel analyses.

  16. Damage Considerations of a Flexible Micro Air Vehicle Wing Using 3-D Laser Vibrometry

    National Research Council Canada - National Science Library

    Mendoza, Jr, Leo L

    2007-01-01

    .... The flexible micro air vehicle wing studied was based on a University of Florida micro air vehicle wing design and was examined using measurements from the Polytec 400-3D Scanning Vibrometer. Comparisons of the wing?s natural frequencies and displacements were made between the wing?s undamaged and damaged states.

  17. Integrated multi-disciplinary design of a sailplane wing

    OpenAIRE

    Strauch, Gregory J.

    1985-01-01

    The objective of this research is to investigate the techniques and payoffs of integrated aircraft design. Lifting line theory and beam theory are used for the analysis of the aerodynamics and the structures of a composite sailplane wing. The wing is described by 33 - 34 design variables which involve the planform geometry, the twist distribution, and thicknesses of the spar caps, spar webs, and the skin at various stations along the wing. The wing design must satisfy 30 â ...

  18. Speed-dependent modulation of wing muscle recruitment intensity and kinematics in two bat species.

    Science.gov (United States)

    Konow, Nicolai; Cheney, Jorn A; Roberts, Thomas J; Iriarte-Díaz, Jose; Breuer, Kenneth S; Waldman, J Rhea S; Swartz, Sharon M

    2017-05-15

    Animals respond to changes in power requirements during locomotion by modulating the intensity of recruitment of their propulsive musculature, but many questions concerning how muscle recruitment varies with speed across modes of locomotion remain unanswered. We measured normalized average burst EMG (aEMG) for pectoralis major and biceps brachii at different flight speeds in two relatively distantly related bat species: the aerial insectivore Eptesicus fuscus , and the primarily fruit-eating Carollia perspicillata These ecologically distinct species employ different flight behaviors but possess similar wing aspect ratio, wing loading and body mass. Because propulsive requirements usually correlate with body size, and aEMG likely reflects force, we hypothesized that these species would deploy similar speed-dependent aEMG modulation. Instead, we found that aEMG was speed independent in E. fuscus and modulated in a U-shaped or linearly increasing relationship with speed in C. perspicillata This interspecific difference may be related to differences in muscle fiber type composition and/or overall patterns of recruitment of the large ensemble of muscles that participate in actuating the highly articulated bat wing. We also found interspecific differences in the speed dependence of 3D wing kinematics: E. fuscus modulates wing flexion during upstroke significantly more than C. perspicillata Overall, we observed two different strategies to increase flight speed: C. perspicillata tends to modulate aEMG, and E. fuscus tends to modulate wing kinematics. These strategies may reflect different requirements for avoiding negative lift and overcoming drag during slow and fast flight, respectively, a subject we suggest merits further study. © 2017. Published by The Company of Biologists Ltd.

  19. FijiWingsPolarity: An open source toolkit for semi-automated detection of cell polarity.

    Science.gov (United States)

    Dobens, Leonard L; Shipman, Anna; Axelrod, Jeffrey D

    2017-12-22

    Epithelial cells are defined by apical-basal and planar cell polarity (PCP) signaling, the latter of which establishes an orthogonal plane of polarity in the epithelial sheet. PCP signaling is required for normal cell migration, differentiation, stem cell generation and tissue repair, and defects in PCP have been associated with developmental abnormalities, neuropathologies and cancers. While the molecular mechanism of PCP is incompletely understood, the deepest insights have come from Drosophila, where PCP is manifest in hairs and bristles across the adult cuticle and organization of the ommatidia in the eye. Fly wing cells are marked by actin-rich trichome structures produced at the distal edge of each cell in the developing wing epithelium and in a mature wing the trichomes orient collectively in the distal direction. Genetic screens have identified key PCP signaling pathway components that disrupt trichome orientation, which has been measured manually in a tedious and error prone process. Here we describe a set of image processing and pattern-recognition macros that can quantify trichome arrangements in micrographs and mark these directly by color, arrow or colored arrow to indicate trichome location, length and orientation. Nearest neighbor calculations are made to exploit local differences in orientation to better and more reliably detect and highlight local defects in trichome polarity. We demonstrate the use of these tools on trichomes in adult wing preps and on actin-rich developing trichomes in pupal wing epithelia stained with phalloidin. FijiWingsPolarity is freely available and will be of interest to a broad community of fly geneticists studying the effect of gene function on PCP.

  20. Wind-tunnel investigation of aerodynamic efficiency of three planar elliptical wings with curvature of quarter-chord line

    Science.gov (United States)

    Mineck, Raymond E.; Vijgen, Paul M. H. W.

    1993-01-01

    Three planar, untwisted wings with the same elliptical chord distribution but with different curvatures of the quarter-chord line were tested in the Langley 8-Foot Transonic Pressure Tunnel (8-ft TPT) and the Langley 7- by 10-Foot High-Speed Tunnel (7 x 10 HST). A fourth wing with a rectangular planform and the same projected area and span was also tested. Force and moment measurements from the 8-ft TPT tests are presented for Mach numbers from 0.3 to 0.5 and angles of attack from -4 degrees to 7 degrees. Sketches of the oil-flow patterns on the upper surfaces of the wings and some force and moment measurements from the 7 x 10 HST tests are presented at a Mach number of 0.5. Increasing the curvature of the quarter-chord line makes the angle of zero lift more negative but has little effect on the drag coefficient at zero lift. The changes in lift-curve slope and in the Oswald efficiency factor with the change in curvature of the quarter-chord line (wingtip location) indicate that the elliptical wing with the unswept quarter-chord line has the lowest lifting efficiency and the elliptical wing with the unswept trailing edge has the highest lifting efficiency; the crescent-shaped planform wing has an efficiency in between.

  1. Use of wing morphometry for the discrimination of some Cerceris ...

    African Journals Online (AJOL)

    use

    2011-12-14

    Dec 14, 2011 ... Figure 12. Thin spline plate graphics for the species belonging to the genus Cerceris. Figure 13. Fore wing landmarks of the significant wing characteristics in the honeybee Apis mellifera. Linnaeus. stated as the traditional wing morphometry that enables the practical discrimination of the honeybee (Apis sp ...

  2. How swifts control their glide performance with morphing wings

    NARCIS (Netherlands)

    Lentink, D.; Muller, U. K.; Stamhuis, E. J.; de Kat, R.; van Gestel, W.; Veldhuis, L. L. M.; Henningsson, P.; Hedenstrom, A.; Videler, J. J.

    2007-01-01

    Gliding birds continually change the shape and size of their wings(1-6), presumably to exploit the profound effect of wing morphology on aerodynamic performance(7-9). That birds should adjust wing sweep to suit glide speed has been predicted qualitatively by analytical glide models(2,10), which

  3. Study of design parameters of flapping-wings

    NARCIS (Netherlands)

    Wang, Q.; Goosen, J.F.L.; Van Keulen, F.

    2014-01-01

    As one of the most important components of a flapping-wing micro air vehicle (FWMAV), the design of an energy-efficient flapping-wing has been a research interest recently. Research on insect flight from different perspectives has been carried out, mainly with regard to wing morphology, flapping

  4. Low Reynolds Number Wing Transients in Rotation and Translation

    Science.gov (United States)

    Jones, Anya; Schlueter, Kristy

    2012-11-01

    The unsteady aerodynamic forces and flow fields generated by a wing undergoing transient motions in both rotation and translation were investigated. An aspect ratio 2 flat plate wing at a 45 deg angle of attack was driven over 84 deg of rotation (3 chord-lengths of travel at 3/4 span) and 3 and 10 chord-lengths of translation in quiescent water at Reynolds numbers between 2,500 and 15,000. Flow visualization on the rotating wing revealed a leading edge vortex that lifted off of the wing surface, but remained in the vicinity of the wing for the duration of the wing stroke. A second spanwise vortex with strong axial flow was also observed. As the tip vortex grew, the leading edge vortex joined the tip vortex in a loop-like structure over the aft half of the wing. Near the leading edge, spanwise flow in the second vortex became entrained in the tip vortex near the corner of the wing. Unsteady force measurements revealed that lift coefficient increased through the constant-velocity portion of the wing stroke. Forces were compared for variations in wing acceleration and Reynolds number for both rotational and translational motions. The effect of tank blockage was investigated by repeating the experiments on multiple wings, varying the distance between the wing tip and tank wall. U.S. Air Force Research Laboratory, Summer Faculty Fellowship Program.

  5. Spatially resolved chemical analysis of cicada wings using laser-ablation electrospray ionization (LAESI) imaging mass spectrometry (IMS).

    Science.gov (United States)

    Román, Jessica K; Walsh, Callee M; Oh, Junho; Dana, Catherine E; Hong, Sungmin; Jo, Kyoo D; Alleyne, Marianne; Miljkovic, Nenad; Cropek, Donald M

    2018-03-01

    Laser-ablation electrospray ionization (LAESI) imaging mass spectrometry (IMS) is an emerging bioanalytical tool for direct imaging and analysis of biological tissues. Performing ionization in an ambient environment, this technique requires little sample preparation and no additional matrix, and can be performed on natural, uneven surfaces. When combined with optical microscopy, the investigation of biological samples by LAESI allows for spatially resolved compositional analysis. We demonstrate here the applicability of LAESI-IMS for the chemical analysis of thin, desiccated biological samples, specifically Neotibicen pruinosus cicada wings. Positive-ion LAESI-IMS accurate ion-map data was acquired from several wing cells and superimposed onto optical images allowing for compositional comparisons across areas of the wing. Various putative chemical identifications were made indicating the presence of hydrocarbons, lipids/esters, amines/amides, and sulfonated/phosphorylated compounds. With the spatial resolution capability, surprising chemical distribution patterns were observed across the cicada wing, which may assist in correlating trends in surface properties with chemical distribution. Observed ions were either (1) equally dispersed across the wing, (2) more concentrated closer to the body of the insect (proximal end), or (3) more concentrated toward the tip of the wing (distal end). These findings demonstrate LAESI-IMS as a tool for the acquisition of spatially resolved chemical information from fragile, dried insect wings. This LAESI-IMS technique has important implications for the study of functional biomaterials, where understanding the correlation between chemical composition, physical structure, and biological function is critical. Graphical abstract Positive-ion laser-ablation electrospray ionization mass spectrometry coupled with optical imaging provides a powerful tool for the spatially resolved chemical analysis of cicada wings.

  6. Air Base Wing and Air Mobility Wing Consolidating on AMC-LED Joint Bases: A Delphi Study

    Science.gov (United States)

    2014-06-13

    AIR BASE WING AND AIR MOBILITY WING CONSOLIDATION ON AMC-LED JOINT BASES: A DELPHI STUDY GRADUATE RESEARCH PAPER Mason E. MacGarvey... DELPHI STUDY GRADUATE RESEARCH PAPER Presented to the Faculty Graduate School of Engineering Management Air Force Institute of Technology...iv AIR BASE WING AND AIR MOBILITY WING CONSOLIDATION ON AMC-LED JOINT BASES: A DELPHI STUDY Mason E. MacGarvey, BS, MBA

  7. Morphing Wing: Experimental Boundary Layer Transition Determination and Wing Vibrations Measurements and Analysis =

    Science.gov (United States)

    Tondji Chendjou, Yvan Wilfried

    This Master's thesis is written within the framework of the multidisciplinary international research project CRIAQ MDO-505. This global project consists of the design, manufacture and testing of a morphing wing box capable of changing the shape of the flexible upper skin of a wing using an actuator system installed inside the wing. This changing of the shape generates a delay in the occurrence of the laminar to turbulent transition area, which results in an improvement of the aerodynamic performances of the morphed wing. This thesis is focused on the technologies used to gather the pressure data during the wind tunnel tests, as well as on the post processing methodologies used to characterize the wing airflow. The vibration measurements of the wing and their real-time graphical representation are also presented. The vibration data acquisition system is detailed, and the vibration data analysis confirms the predictions of the flutter analysis performed on the wing prior to wind tunnel testing at the IAR-NRC. The pressure data was collected using 32 highly-sensitive piezoelectric sensors for sensing the pressure fluctuations up to 10 KHz. These sensors were installed along two wing chords, and were further connected to a National Instrument PXI real-time acquisition system. The acquired pressure data was high-pass filtered, analyzed and visualized using Fast Fourier Transform (FFT) and Standard Deviation (SD) approaches to quantify the pressure fluctuations in the wing airflow, as these allow the detection of the laminar to turbulent transition area. Around 30% of the cases tested in the IAR-NRC wind tunnel were optimized for drag reduction by the morphing wing procedure. The obtained pressure measurements results were compared with results obtained by infrared thermography visualization, and were used to validate the numerical simulations. Two analog accelerometers able to sense dynamic accelerations up to +/-16g were installed in both the wing and the aileron boxes

  8. Machine Learning for Flapping Wing Flight Control

    NARCIS (Netherlands)

    Goedhart, Menno; van Kampen, E.; Armanini, S.F.; de Visser, C.C.; Chu, Q.

    2018-01-01

    Flight control of Flapping Wing Micro Air Vehicles is challenging, because of their complex dynamics and variability due to manufacturing inconsistencies. Machine Learning algorithms can be used to tackle these challenges. A Policy Gradient algorithm is used to tune the gains of a

  9. Migration on Wings Aerodynamics and Energetics

    CERN Document Server

    Kantha, Lakshmi

    2012-01-01

    This book is an effort to explore the technical aspects associated with bird flight and migration on wings. After a short introduction on the birds migration, the book reviews the aerodynamics and Energetics of Flight and presents the calculation of the Migration Range. In addition, the authors explains aerodynamics of the formation flight and finally introduces great flight diagrams.

  10. Oblique-Flying-Wing Supersonic Transport Airplane

    Science.gov (United States)

    Van Der Velden, Alexander J. M.

    1992-01-01

    Oblique-flying-wing supersonic airplane proposed as possible alternative to B747B (or equivalent). Tranports passengers and cargo as fast as twice speed of sound at same cost as current subsonic transports. Flies at same holding speeds as present supersonic transports but requires only half takeoff distance.

  11. Chemical evaluation of winged beans ( Psophocarpus ...

    African Journals Online (AJOL)

    Chemical evaluation of winged beans ( Psophocarpus Tetragonolobus ), Pitanga cherries ( Eugenia uniflora) and orchid fruit ( Orchid fruit myristic a) ... The acid value ranged between 0.71 and 2.82 mg/KOH/g while iodine value ranged between 91.15 and 144.57. The refractive index ranged between 1.465 and 1.474 in all ...

  12. ``Schooling'' of wing pairs in flapping flight

    Science.gov (United States)

    Ramananarivo, Sophie; Zhang, Jun; Ristroph, Leif; AML, Courant Collaboration; Physics NYU Collaboration

    2015-11-01

    The experimental setup implements two independent flapping wings swimming in tandem. Both are driven with the same prescribed vertical heaving motion, but the horizontal motion is free, which means that the swimmers can take up any relative position and forward speed. Experiments show however clearly coordinated motions, where the pair of wings `crystallize' into specific stable arrangements. The follower wing locks into the path of the leader, adopting its speed, and with a separation distance that takes on one of several discrete values. By systematically varying the kinematics and wing size, we show that the set of stable spacings is dictated by the wavelength of the periodic wake structure. The forces maintaining the pair cohesion are characterized by applying an external force to the follower to perturb it away from the `stable wells'. These results show that hydrodynamics alone is sufficient to induce cohesive and coordinated collective locomotion through a fluid, and we discuss the hypothesis that fish schools and bird flocks also represent stable modes of motion.

  13. Wind Tunnel Test of a Risk-Reduction Wing/Fuselage Model to Examine Juncture-Flow Phenomena

    Science.gov (United States)

    Kegerise, Michael A.; Neuhart, Dan H.

    2016-01-01

    A wing/fuselage wind-tunnel model was tested in the Langley 14- by 22-foot Subsonic Wind Tunnel in preparation for a highly-instrumented Juncture Flow Experiment to be conducted in the same facility. This test, which was sponsored by the NASA Transformational Tool and Technologies Project, is part of a comprehensive set of experimental and computational research activities to develop revolutionary, physics-based aeronautics analysis and design capability. The objectives of this particular test were to examine the surface and off-body flow on a generic wing/body combination to: 1) choose a final wing for a future, highly instrumented model, 2) use the results to facilitate unsteady pressure sensor placement on the model, 3) determine the area to be surveyed with an embedded laser-doppler velocimetry (LDV) system, 4) investigate the primary juncture corner- flow separation region using particle image velocimetry (PIV) to see if the particle seeding is adequately entrained and to examine the structure in the separated region, and 5) to determine the similarity of observed flow features with those predicted by computational fluid dynamics (CFD). This report documents the results of the above experiment that specifically address the first three goals. Multiple wing configurations were tested at a chord Reynolds number of 2.4 million. Flow patterns on the surface of the wings and in the region of the wing/fuselage juncture were examined using oil- flow visualization and infrared thermography. A limited number of unsteady pressure sensors on the fuselage around the wing leading and trailing edges were used to identify any dynamic effects of the horseshoe vortex on the flow field. The area of separated flow in the wing/fuselage juncture near the wing trailing edge was observed for all wing configurations at various angles of attack. All of the test objectives were met. The staff of the 14- by 22-foot Subsonic Wind Tunnel provided outstanding support and delivered

  14. Aerodynamic comparison of a butterfly-like flapping wing-body model and a revolving-wing model

    Science.gov (United States)

    Suzuki, Kosuke; Yoshino, Masato

    2017-06-01

    The aerodynamic performance of flapping- and revolving-wing models is investigated by numerical simulations based on an immersed boundary-lattice Boltzmann method. As wing models, we use (i) a butterfly-like model with a body and flapping-rectangular wings and (ii) a revolving-wing model with the same wings as the flapping case. Firstly, we calculate aerodynamic performance factors such as the lift force, the power, and the power loading of the two models for Reynolds numbers in the range of 50-1000. For the flapping-wing model, the power loading is maximal for the maximum angle of attack of 90°, a flapping amplitude of roughly 45°, and a phase shift between the flapping angle and the angle of attack of roughly 90°. For the revolving-wing model, the power loading peaks for an angle of attack of roughly 45°. In addition, we examine the ground effect on the aerodynamic performance of the revolving-wing model. Secondly, we compare the aerodynamic performance of the flapping- and revolving-wing models at their respective maximal power loadings. It is found that the revolving-wing model is more efficient than the flapping-wing model both when the body of the latter is fixed and where it can move freely. Finally, we discuss the relative agilities of the flapping- and revolving-wing models.

  15. Paleozoic Nymphal Wing Pads Support Dual Model of Insect Wing Origins.

    Science.gov (United States)

    Prokop, Jakub; Pecharová, Martina; Nel, André; Hörnschemeyer, Thomas; Krzemińska, Ewa; Krzemiński, Wiesław; Engel, Michael S

    2017-01-23

    The appearance of wings in insects, early in their evolution [1], has been one of the more critical innovations contributing to their extraordinary diversity. Despite the conspicuousness and importance of wings, the origin of these structures has been difficult to resolve and represented one of the "abominable mysteries" in evolutionary biology [2]. More than a century of debate has boiled the matter down to two competing alternatives-one of wings representing an extension of the thoracic notum, the other stating that they are appendicular derivations from the lateral body wall. Recently, a dual model has been supported by genomic and developmental data [3-6], representing an amalgamation of elements from both the notal and pleural hypotheses. Here, we reveal crucial information from the wing pad joints of Carboniferous palaeodictyopteran insect nymphs using classical and high-tech techniques. These nymphs had three pairs of wing pads that were medially articulated to the thorax but also broadly contiguous with the notum anteriorly and posteriorly (details unobservable in modern insects), supporting their overall origin from the thoracic notum as well as the expected medial, pleural series of axillary sclerites. Our study provides support for the formation of the insect wing from the thoracic notum as well as the already known pleural elements of the arthropodan leg. These results support the unique, dual model for insect wing origins and the convergent reduction of notal fusion in more derived clades, presumably due to wing rotation during development, and they help to bring resolution to this long-standing debate. Copyright © 2017 Elsevier Ltd. All rights reserved.

  16. Distribution and predictors of wing shape and size variability in three sister species of solitary bees.

    Directory of Open Access Journals (Sweden)

    Simon Dellicour

    Full Text Available Morphological traits can be highly variable over time in a particular geographical area. Different selective pressures shape those traits, which is crucial in evolutionary biology. Among these traits, insect wing morphometry has already been widely used to describe phenotypic variability at the inter-specific level. On the contrary, fewer studies have focused on intra-specific wing morphometric variability. Yet, such investigations are relevant to study potential convergences of variation that could highlight micro-evolutionary processes. The recent sampling and sequencing of three solitary bees of the genus Melitta across their entire species range provides an excellent opportunity to jointly analyse genetic and morphometric variability. In the present study, we first aim to analyse the spatial distribution of the wing shape and centroid size (used as a proxy for body size variability. Secondly, we aim to test different potential predictors of this variability at both the intra- and inter-population levels, which includes genetic variability, but also geographic locations and distances, elevation, annual mean temperature and precipitation. The comparison of spatial distribution of intra-population morphometric diversity does not reveal any convergent pattern between species, thus undermining the assumption of a potential local and selective adaptation at the population level. Regarding intra-specific wing shape differentiation, our results reveal that some tested predictors, such as geographic and genetic distances, are associated with a significant correlation for some species. However, none of these predictors are systematically identified for the three species as an important factor that could explain the intra-specific morphometric variability. As a conclusion, for the three solitary bee species and at the scale of this study, our results clearly tend to discard the assumption of the existence of a common pattern of intra-specific signal

  17. Distribution and predictors of wing shape and size variability in three sister species of solitary bees.

    Science.gov (United States)

    Dellicour, Simon; Gerard, Maxence; Prunier, Jérôme G; Dewulf, Alexandre; Kuhlmann, Michael; Michez, Denis

    2017-01-01

    Morphological traits can be highly variable over time in a particular geographical area. Different selective pressures shape those traits, which is crucial in evolutionary biology. Among these traits, insect wing morphometry has already been widely used to describe phenotypic variability at the inter-specific level. On the contrary, fewer studies have focused on intra-specific wing morphometric variability. Yet, such investigations are relevant to study potential convergences of variation that could highlight micro-evolutionary processes. The recent sampling and sequencing of three solitary bees of the genus Melitta across their entire species range provides an excellent opportunity to jointly analyse genetic and morphometric variability. In the present study, we first aim to analyse the spatial distribution of the wing shape and centroid size (used as a proxy for body size) variability. Secondly, we aim to test different potential predictors of this variability at both the intra- and inter-population levels, which includes genetic variability, but also geographic locations and distances, elevation, annual mean temperature and precipitation. The comparison of spatial distribution of intra-population morphometric diversity does not reveal any convergent pattern between species, thus undermining the assumption of a potential local and selective adaptation at the population level. Regarding intra-specific wing shape differentiation, our results reveal that some tested predictors, such as geographic and genetic distances, are associated with a significant correlation for some species. However, none of these predictors are systematically identified for the three species as an important factor that could explain the intra-specific morphometric variability. As a conclusion, for the three solitary bee species and at the scale of this study, our results clearly tend to discard the assumption of the existence of a common pattern of intra-specific signal/structure within the

  18. A study on forces acting on a flapping wing

    Directory of Open Access Journals (Sweden)

    Cetiner O.

    2013-04-01

    Full Text Available In order to study the forces acting on a flapping wing, an experimental investigation is performed in steady water flow. In this study, a SD7003 airfoil undergoes combined pitching and plunging motion which simulates the forward flight of small birds. The frequency of pitching motion is equal to the frequency of plunging motion and pitch leads the plunge by a phase angle of 90 degrees. The experiments are conducted at Reynolds numbers of 2500 ≤ Re ≤ 13700 and the vortex formation is recorded using the digital particle image velocimetry (DPIV technique. A prediction of thrust force and efficiency is calculated from the average wake deficit of DPIV data, the near-wake vorticity patterns and time dependent velocity vectors are determined to comment on the thrust and drag indication. Direct force measurements are attempted using a Force/Torque sensor which is capable of measuring forces and moments in three axial directions.

  19. Multiple cues for winged morph production in an aphid metacommunity.

    Directory of Open Access Journals (Sweden)

    Mohsen Mehrparvar

    Full Text Available Environmental factors can lead individuals down different developmental pathways giving rise to distinct phenotypes (phenotypic plasticity. The production of winged or unwinged morphs in aphids is an example of two alternative developmental pathways. Dispersal is paramount in aphids that often have a metapopulation structure, where local subpopulations frequently go extinct, such as the specialized aphids on tansy (Tanacetum vulgare. We conducted various experiments to further understand the cues involved in the production of winged dispersal morphs by the two dominant species of the tansy aphid metacommunity, Metopeurum fuscoviride and Macrosiphoniella tanacetaria. We found that the ant-tended M. fuscoviride produced winged individuals predominantly at the beginning of the season while the untended M. tanacetaria produced winged individuals throughout the season. Winged mothers of both species produced winged offspring, although in both species winged offspring were mainly produced by unwinged females. Crowding and the presence of predators, effects already known to influence wing production in other aphid species, increased the percentage of winged offspring in M. tanacetaria, but not in M. fuscoviride. We find there are also other factors (i.e. temporal effects inducing the production of winged offspring for natural aphid populations. Our results show that the responses of each aphid species are due to multiple wing induction cues.

  20. The biomechanical origin of extreme wing allometry in hummingbirds.

    Science.gov (United States)

    Skandalis, Dimitri A; Segre, Paolo S; Bahlman, Joseph W; Groom, Derrick J E; Welch, Kenneth C; Witt, Christopher C; McGuire, Jimmy A; Dudley, Robert; Lentink, David; Altshuler, Douglas L

    2017-10-19

    Flying animals of different masses vary widely in body proportions, but the functional implications of this variation are often unclear. We address this ambiguity by developing an integrative allometric approach, which we apply here to hummingbirds to examine how the physical environment, wing morphology and stroke kinematics have contributed to the evolution of their highly specialised flight. Surprisingly, hummingbirds maintain constant wing velocity despite an order of magnitude variation in body weight; increased weight is supported solely through disproportionate increases in wing area. Conversely, wing velocity increases with body weight within species, compensating for lower relative wing area in larger individuals. By comparing inter- and intraspecific allometries, we find that the extreme wing area allometry of hummingbirds is likely an adaptation to maintain constant burst flight capacity and induced power requirements with increasing weight. Selection for relatively large wings simultaneously maximises aerial performance and minimises flight costs, which are essential elements of humming bird life history.

  1. Association between antennal phenotype, wing polymorphism and sex in the genus Mepraia (Reduviidae: Triatominae).

    Science.gov (United States)

    Moreno, Mariana Laura; Gorla, David; Catalá, Silvia

    2006-05-01

    Mepraia spinolai and Mepraia gajardoi (Hemiptera, Reduviidae) are only found in the arid regions of northern Chile. Mepraia is the only genus of Triatominae with marked wing polymorphism. Females of both species are micropterous, males of M. spinolai may be micropterous, brachypterous or macropterous, while males of M. gajardoi are always brachypterous. Because of this wing polymorphism, Mepraia can be used as a model to analyze morphological adaptations related to the flying activity and evolutionary relationships in the Triatominae. The study presented here analyses the antennal phenotype of the Mepraia species and carries out a comparison with the two species of Triatoma included in the spinolai complex (Triatoma eratyrusiformis and Triatoma breyeri). The analysis of the antennal phenotype of Mepraia showed a marked intraspecific phenotypic variability related with sex and wing condition. The number and length of multiporous trichoid sensilla (TH) on the pedicel are significantly higher and longer in winged males and are strongly reduced in females. The great length of the TH sensillum is an infrequent characteristic within Triatominae. The results show that phenetic distances between sexes are greater than between the two species. Similarity between the Mepraia species and T. eratyrusiformis was found as they all show long bristles (BR) and TH and two sized basiconic receptors (BA), a pattern that have not been observed in other species of Triatominae. These characters are not present in T. breyeri.

  2. Phenotypic differentiation and phylogenetic signal of wing shape in western European biting midges, Culicoides spp., of the subgenus Avaritia

    DEFF Research Database (Denmark)

    Muñoz-Muñoz, F.; Talavera, S.; Carpenter, S.

    2014-01-01

    of the subgenus Avaritia were considered in the study (C. obsoletus (Meigen); C. scoticus Kettle and Lawson; C. chiopterus (Meigen); C. dewulfi Goetghebuer and C. imicola (Kieffer)). The study demonstrated that over 90% of individuals could be separated correctly into species by their wing shape and that patterns...

  3. Identification of the species of the Cheilosia variabilis group (Diptera, Syrphidae) from the Balkan Peninsula using wing geometric morphometrics, with the revision of status of C. melanopa redi Vujic, 1996

    NARCIS (Netherlands)

    Francuski, Lj.; Vujic, A.; Kovacevic, A.; Ludoski, J.; Milankov, V.

    2009-01-01

    The present study investigates phenotypic differentiation patterns among four species of the Cheilosia variabilis group (Diptera, Syrphidae) using a landmark-based geometric morphometric approach. Herein, wing geometric morphometrics established species boundaries that confirm C. melanopa and C.

  4. Is pushing the wall, the best known method for scapular winging, really the best? A Comparative analysis of various methods in neuromuscular disorders.

    Science.gov (United States)

    Khadilkar, Satish V; Chaudhari, Chetan R; Soni, Girish; Bhutada, Ashish

    2015-04-15

    'Pushing the wall' has found acceptance in medical teachings. Other methods of scapular winging are less known. Comparative evaluation of the five available methods has not been undertaken. This study focuses on evaluation of the available methods in groups of neuromuscular disorders to select the most sensitive method and to characterize patterns of scapular winging. A survey of methods practiced by clinicians also forms a part of the study. Prospective study. Part A: questionnaire based survey of clinicians to explore the preferred method of examination for scapular winging. Part B: comparative analysis of five methods of scapular winging in four categories of neuromuscular disorders [facioscapulohumeral muscular dystrophy (FSHD), limb girdle muscular dystrophy, dystrophinopathies and neurogenic disorders]. Forward lowering of arms was the most sensitive method [100%]. The use of this method in clinical teachings and routine bedside examination should be promoted. Pushing the wall was the most popular method, but was fourth in the sensitivity [60.41%]. Arm maneuvers can bring out winging, when it is not apparent at rest. FSHD patients had a unique combination of winging at rest, persistence of winging throughout the range of motion and elevation of scapulae. Copyright © 2015 Elsevier B.V. All rights reserved.

  5. The Banded-wing Moselia infuscata (Claassen) Phenotype from California and Oregon, U.S.A. (Plecoptera: Leuctridae).

    Science.gov (United States)

    Gill, Brian A; Kondratieff, Boris C; Stark, Bill P; Sandberg, John B

    2015-01-21

    Moselia specimens from California and Oregon with a banded-wing phenotype were found to be indistinguishable morphologically from those of M. infuscata (Claassen) with typical wing pigment pattern. Preliminary DNA barcode data (Cytochrome c Oxidase subunit I [COI]), however, show significant genetic variation among four populations including three from northern California sites and one from southern Oregon. Although this genetic variation exceeded standard divergence thresholds often used to recognize distinct stream insect species, no new taxa are proposed at this time due to the preliminary nature of the data. 

  6. The optimal design of UAV wing structure

    Science.gov (United States)

    Długosz, Adam; Klimek, Wiktor

    2018-01-01

    The paper presents an optimal design of UAV wing, made of composite materials. The aim of the optimization is to improve strength and stiffness together with reduction of the weight of the structure. Three different types of functionals, which depend on stress, stiffness and the total mass are defined. The paper presents an application of the in-house implementation of the evolutionary multi-objective algorithm in optimization of the UAV wing structure. Values of the functionals are calculated on the basis of results obtained from numerical simulations. Numerical FEM model, consisting of different composite materials is created. Adequacy of the numerical model is verified by results obtained from the experiment, performed on a tensile testing machine. Examples of multi-objective optimization by means of Pareto-optimal set of solutions are presented.

  7. Adjoint-based optimization for flapping wings

    Science.gov (United States)

    Xu, Min; Wei, Mingjun

    2012-11-01

    Adjoint-based methods show great potential in flow control and optimization of complex problems with high- or infinite-dimensional control space. It is attractive to solve an adjoint problem to understand the complex effects from multiple control parameters to a few performance indicators of the flight of birds or insects. However, the traditional approach to formulate the adjoint problem becomes either impossible or too complex when arbitrary moving boundary (e.g. flapping wings) and its perturbation is considered. Here, we use non-cylindrical calculus to define the perturbation. So that, a simple adjoint system can be derived directly in the inertial coordinate. The approach is first applied to the optimization of cylinder oscillation and later to flapping wings. Supported by AFOSR.

  8. Quantifying the dynamic wing morphing of hovering hummingbird.

    Science.gov (United States)

    Maeda, Masateru; Nakata, Toshiyuki; Kitamura, Ikuo; Tanaka, Hiroto; Liu, Hao

    2017-09-01

    Animal wings are lightweight and flexible; hence, during flapping flight their shapes change. It has been known that such dynamic wing morphing reduces aerodynamic cost in insects, but the consequences in vertebrate flyers, particularly birds, are not well understood. We have developed a method to reconstruct a three-dimensional wing model of a bird from the wing outline and the feather shafts (rachides). The morphological and kinematic parameters can be obtained using the wing model, and the numerical or mechanical simulations may also be carried out. To test the effectiveness of the method, we recorded the hovering flight of a hummingbird ( Amazilia amazilia ) using high-speed cameras and reconstructed the right wing. The wing shape varied substantially within a stroke cycle. Specifically, the maximum and minimum wing areas differed by 18%, presumably due to feather sliding; the wing was bent near the wrist joint, towards the upward direction and opposite to the stroke direction; positive upward camber and the 'washout' twist (monotonic decrease in the angle of incidence from the proximal to distal wing) were observed during both half-strokes; the spanwise distribution of the twist was uniform during downstroke, but an abrupt increase near the wrist joint was found during upstroke.

  9. Optimization of aerodynamic efficiency for twist morphing MAV wing

    Directory of Open Access Journals (Sweden)

    N.I. Ismail

    2014-06-01

    Full Text Available Twist morphing (TM is a practical control technique in micro air vehicle (MAV flight. However, TM wing has a lower aerodynamic efficiency (CL/CD compared to membrane and rigid wing. This is due to massive drag penalty created on TM wing, which had overwhelmed the successive increase in its lift generation. Therefore, further CL/CDmax optimization on TM wing is needed to obtain the optimal condition for the morphing wing configuration. In this paper, two-way fluid–structure interaction (FSI simulation and wind tunnel testing method are used to solve and study the basic wing aerodynamic performance over (non-optimal TM, membrane and rigid wings. Then, a multifidelity data metamodel based design optimization (MBDO process is adopted based on the Ansys-DesignXplorer frameworks. In the adaptive MBDO process, Kriging metamodel is used to construct the final multifidelity CL/CD responses by utilizing 23 multi-fidelity sample points from the FSI simulation and experimental data. The optimization results show that the optimal TM wing configuration is able to produce better CL/CDmax magnitude by at least 2% than the non-optimal TM wings. The flow structure formation reveals that low TV strength on the optimal TM wing induces low CD generation which in turn improves its overall CL/CDmax performance.

  10. CFD Analysis of UAV Flying Wing

    Directory of Open Access Journals (Sweden)

    Vasile PRISACARIU

    2016-09-01

    Full Text Available Numerical methods for solving equations describing the evolution of 3D fluid experienced a significant development closely related to the progress of information systems. Today, especially in the field of fluid mechanics, numerical simulations allow the study of gas-thermodynamic confirmed by experimental techniques in wind tunnel conditions and actual flight tests for modeling complex aircraft. The article shows a case of numerical analysis of the lifting surface on the UAV type flying wing.

  11. Topology Optimization of an Aircraft Wing

    Science.gov (United States)

    2015-06-11

    can combine the advantages of a variable stiffness design with- out the use of actuators. Curved beams, which couple torsion and bending , counteract... torsional deflection, control natural frequency, exploit coupling of bending and tor- sion to control flutter, reduce thickness to chord ratios due to...disregarded any bending or torsional effects caused by displacement of the wing, and was thus not considered. Therefore, the initial design analysis

  12. Tracing the evolution of avian wing digits.

    Science.gov (United States)

    Xu, Xing; Mackem, Susan

    2013-06-17

    It is widely accepted that birds are a subgroup of dinosaurs, but there is an apparent conflict: modern birds have been thought to possess only the middle three fingers (digits II-III-IV) of an idealized five-digit tetrapod hand based on embryological data, but their Mesozoic tetanuran dinosaur ancestors are considered to have the first three digits (I-II-III) based on fossil evidence. How could such an evolutionary quirk arise? Various hypotheses have been proposed to resolve this paradox. Adding to the confusion, some recent developmental studies support a I-II-III designation for avian wing digits whereas some recent paleontological data are consistent with a II-III-IV identification of the Mesozoic tetanuran digits. A comprehensive analysis of both paleontological and developmental data suggests that the evolution of the avian wing digits may have been driven by homeotic transformations of digit identity, which are more likely to have occurred in a partial and piecemeal manner. Additionally, recent genetic studies in mouse models showing plausible mechanisms for central digit loss invite consideration of new alternative possibilities (I-II-IV or I-III-IV) for the homologies of avian wing digits. While much progress has been made, some advances point to the complexity of the problem and a final resolution to this ongoing debate demands additional work from both paleontological and developmental perspectives, which will surely yield new insights on mechanisms of evolutionary adaptation. Copyright © 2013 Elsevier Ltd. All rights reserved.

  13. Unsteady fluid dynamics around a hovering wing

    Science.gov (United States)

    Krishna, Swathi; Green, Melissa; Mulleners, Karen

    2017-11-01

    The unsteady flow around a hovering flat plate wing has been investigated experimentally using particle image velocimetry and direct force measurements. The measurements are conducted on a wing that rotates symmetrically about the stroke reversal at a reduced frequency of k = 0.32 and Reynolds number of Re = 220 . The Lagrangian finite-time Lyapunov exponent method is used to analyse the unsteady flow fields by identifying dynamically relevant flow features such as the primary leading edge vortex (LEV), secondary vortices, and topological saddles, and their evolution within a flapping cycle. The flow evolution is divided into four stages that are characterised by the LEV (a)emergence, (b)growth, (c)lift-off, and (d)breakdown and decay. Tracking saddle points is shown to be helpful in defining the LEV lift-off which occurs at the maximum stroke velocity. The flow fields are correlated with the aerodynamic forces revealing that the maximum lift and drag are observed just before LEV lift-off. The end of wing rotation in the beginning of the stroke stimulates a change in the direction of the LEV growth and the start of rotation at the end of the stroke triggers the breakdown of the LEV.

  14. A new genus and species of tettigarctid cicada from the early Miocene of New Zealand: Paratettigarcta zealandica (Hemiptera, Auchenorrhyncha, Tettigarctidae

    Directory of Open Access Journals (Sweden)

    Uwe Kaulfuss

    2015-03-01

    Full Text Available A new genus and species of primitive cicada (Hemiptera: Tettigarctidae is described from the early Miocene of southern New Zealand. Paratettigarcta zealandica gen. et sp. n. is the first cicada (Cicadoidea fossil from New Zealand and exhibits wing venation patterns typical for the subfamily Tettigarctinae. It differs from other fossil taxa and the extant genus Tettigarcta in the early divergence of CuA2 from the nodal line in the forewing, its parallel-sided subcostal cell, the early bifurcation of vein M and long apical cells of the hindwing, and in wing pigmentation patterns.

  15. Wing and body kinematics of takeoff and landing flight in the pigeon (Columba livia).

    Science.gov (United States)

    Berg, Angela M; Biewener, Andrew A

    2010-05-01

    Takeoff and landing are critical phases in a flight. To better understand the functional importance of the kinematic adjustments birds use to execute these flight modes, we studied the wing and body movements of pigeons (Columba livia) during short-distance free-flights between two perches. The greatest accelerations were observed during the second wingbeat of takeoff. The wings were responsible for the majority of acceleration during takeoff and landing, with the legs contributing only one-quarter of the acceleration. Parameters relating to aerodynamic power output such as downstroke amplitude, wingbeat frequency and downstroke velocity were all greatest during takeoff flight and decreased with each successive takeoff wingbeat. This pattern indicates that downstroke velocity must be greater for accelerating flight to increase the amount of air accelerated by the wings. Pigeons used multiple mechanisms to adjust thrust and drag to accelerate during takeoff and decelerate during landing. Body angle, tail angle and wing plane angles all shifted from more horizontal orientations during takeoff to near-vertical orientations during landing, thereby reducing drag during takeoff and increasing drag during landing. The stroke plane was tilted steeply downward throughout takeoff (increasing from -60+/-5 deg. to -47+/-1 deg.), supporting our hypothesis that a downward-tilted stroke plane pushes more air rearward to accelerate the bird forward. Similarly, the stroke plane tilted upward during landing (increasing from -1+/-2 deg. to 17+/-7 deg.), implying that an upward-tilted stroke plane pushes more air forward to slow the bird down. Rotations of the stroke plane, wing planes and tail were all strongly correlated with rotation of the body angle, suggesting that pigeons are able to redirect aerodynamic force and shift between flight modes through modulation of body angle alone.

  16. PHOTOGRAMMETRIC MEASUREMENTS IN FIXED WING UAV IMAGERY

    Directory of Open Access Journals (Sweden)

    E. Gülch

    2012-07-01

    Full Text Available Several flights have been undertaken with PAMS (Photogrammetric Aerial Mapping System by Germap, Germany, which is briefly introduced. This system is based on the SmartPlane fixed-wing UAV and a CANON IXUS camera system. The plane is equipped with GPS and has an infrared sensor system to estimate attitude values. A software has been developed to link the PAMS output to a standard photogrammetric processing chain built on Trimble INPHO. The linking of the image files and image IDs and the handling of different cases with partly corrupted output have to be solved to generate an INPHO project file. Based on this project file the software packages MATCH-AT, MATCH-T DSM, OrthoMaster and OrthoVista for digital aerial triangulation, DTM/DSM generation and finally digital orthomosaik generation are applied. The focus has been on investigations on how to adapt the "usual" parameters for the digital aerial triangulation and other software to the UAV flight conditions, which are showing high overlaps, large kappa angles and a certain image blur in case of turbulences. It was found, that the selected parameter setup shows a quite stable behaviour and can be applied to other flights. A comparison is made to results from other open source multi-ray matching software to handle the issue of the described flight conditions. Flights over the same area at different times have been compared to each other. The major objective was here to see, on how far differences occur relative to each other, without having access to ground control data, which would have a potential for applications with low requirements on the absolute accuracy. The results show, that there are influences of weather and illumination visible. The "unusual" flight pattern, which shows big time differences for neighbouring strips has an influence on the AT and DTM/DSM generation. The results obtained so far do indicate problems in the stability of the camera calibration. This clearly requests a

  17. First discovery of fossil winged seeds of Pinus L. (family Pinaceae) from the Indian Cenozoic and its palaeobiogeographic significance

    Science.gov (United States)

    Khan, Mahasin Ali; Bera, Subir

    2017-07-01

    The occurrences of Pinus L. (family Pinaceae) megafossils (cones and leaf remains) have been abundantly documented from the Cenozoic sediments of eastern Asia (Japan and China), but none has been confirmed from the Indian Cenozoic till date. Here, we describe Pinus arunachalensis Khan and Bera, sp. nov. on the basis of seed remains from the middle to late Miocene Siwalik sediments of the Dafla Formation exposed around West Kameng district in Arunachal Pradesh, eastern Himalaya. Seeds are winged, broadly oblong to oval in outline, 1.3-1.5 cm long and 0.4-0.6 cm broad (in the middle part), located basipetally and symmetrically to wing, cellular pattern of wing is seemingly undulatory and parallel with the long axis of the wing. So far, this report provides the first ever fossil record of Pinus winged seeds from India. This record suggests that Pinus was an important component of tropical-subtropical evergreen forest in the area during the Miocene and this group subsequently declined from the local vegetation probably because of the gradual intensification of MSI (monsoon index) from the Miocene to the present. We also review the historical phytogeography and highlight the phytogeographic implication of this genus.

  18. Wing geometry of Triatoma sordida (Hemiptera: Reduviidae) populations from Brazil.

    Science.gov (United States)

    Vendrami, Daniel Pagotto; Obara, Marcos Takashi; Gurgel-Gonçalves, Rodrigo; Ceretti-Junior, Walter; Marrelli, Mauro Toledo

    2017-04-01

    Triatoma sordida has a widespread distribution in Argentina, Bolivia, Brazil, Paraguay, and Uruguay and is frequently found in peridomestic environments. We investigated size and shape variability of T. sordida wings across Brazil. Field-collected adults from twelve populations were studied. For each individual female, seven landmarks on the right wing were digitalized. Shape variables derived from Procrustes superimposition were used in Principal Component Analysis (PCA). Wing size and shape variations among populations was explored by means of ANOVA. Wing centroid size was significantly different among T. sordida populations; specimens from Bahia (East) were larger than those of Mato Grosso do Sul (West). PCA based on wing shape variables showed low wing shape variability. These results reinforce previous data showing low genetic variability among T. sordida populations from Brazil. Copyright © 2017 Elsevier B.V. All rights reserved.

  19. Butterfly Wings Are Three-Dimensional: Pupal Cuticle Focal Spots and Their Associated Structures in Junonia Butterflies.

    Science.gov (United States)

    Taira, Wataru; Otaki, Joji M

    2016-01-01

    Butterfly wing color patterns often contain eyespots, which are developmentally determined at the late larval and early pupal stages by organizing activities of focal cells that can later form eyespot foci. In the pupal stage, the focal position of a future eyespot is often marked by a focal spot, one of the pupal cuticle spots, on the pupal surface. Here, we examined the possible relationships of the pupal focal spots with the underneath pupal wing tissues and with the adult wing eyespots using Junonia butterflies. Large pupal focal spots were found in two species with large adult eyespots, J. orithya and J. almana, whereas only small pupal focal spots were found in a species with small adult eyespots, J. hedonia. The size of five pupal focal spots on a single wing was correlated with the size of the corresponding adult eyespots in J. orithya. A pupal focal spot was a three-dimensional bulge of cuticle surface, and the underside of the major pupal focal spot exhibited a hollowed cuticle in a pupal case. Cross sections of a pupal wing revealed that the cuticle layer shows a curvature at a focal spot, and a positional correlation was observed between the cuticle layer thickness and its corresponding cell layer thickness. Adult major eyespots of J. orithya and J. almana exhibited surface elevations and depressions that approximately correspond to the coloration within an eyespot. Our results suggest that a pupal focal spot is produced by the organizing activity of focal cells underneath the focal spot. Probably because the focal cell layer immediately underneath a focal spot is thicker than that of its surrounding areas, eyespots of adult butterfly wings are three-dimensionally constructed. The color-height relationship in adult eyespots might have an implication in the developmental signaling for determining the eyespot color patterns.

  20. Scaling the Drosophila Wing: TOR-Dependent Target Gene Access by the Hippo Pathway Transducer Yorkie.

    Science.gov (United States)

    Parker, Joseph; Struhl, Gary

    2015-10-01

    Organ growth is controlled by patterning signals that operate locally (e.g., Wingless/Ints [Wnts], Bone Morphogenetic Proteins [BMPs], and Hedgehogs [Hhs]) and scaled by nutrient-dependent signals that act systemically (e.g., Insulin-like peptides [ILPs] transduced by the Target of Rapamycin [TOR] pathway). How cells integrate these distinct inputs to generate organs of the appropriate size and shape is largely unknown. The transcriptional coactivator Yorkie (Yki, a YES-Associated Protein, or YAP) acts downstream of patterning morphogens and other tissue-intrinsic signals to promote organ growth. Yki activity is regulated primarily by the Warts/Hippo (Wts/Hpo) tumour suppressor pathway, which impedes nuclear access of Yki by a cytoplasmic tethering mechanism. Here, we show that the TOR pathway regulates Yki by a separate and novel mechanism in the Drosophila wing. Instead of controlling Yki nuclear access, TOR signaling governs Yki action after it reaches the nucleus by allowing it to gain access to its target genes. When TOR activity is inhibited, Yki accumulates in the nucleus but is sequestered from its normal growth-promoting target genes--a phenomenon we term "nuclear seclusion." Hence, we posit that in addition to its well-known role in stimulating cellular metabolism in response to nutrients, TOR also promotes wing growth by liberating Yki from nuclear seclusion, a parallel pathway that we propose contributes to the scaling of wing size with nutrient availability.

  1. Free vibration analysis of dragonfly wings using finite element method

    OpenAIRE

    M Darvizeh; A Darvizeh; H Rajabi; A Rezaei

    2016-01-01

    In the present work, investigations on the microstructure and mechanicalproperties of the dragonfly wing are carried out and numerical modelingbased on Finite Element Method (FEM) is developed to predict Flightcharacteristics of dragonfly wings. Vibrational behavior of wings typestructures is immensely important in analysis, design and manufacturing ofsimilar engineering structures. For this purpose natural frequencies andmode shapes are calculated. In addition, the kind of deformation in eac...

  2. Heat stress and age induced maternal effects on wing size and shape in parthenogenetic Drosophila mercatorum

    DEFF Research Database (Denmark)

    Andersen, DH; Pertoldi, C; Scali, V

    2005-01-01

    and progeny from heat stressed flies in both size and shape with increased maternal heat stress temperature. The effects of maternal age, however, led to different responses in size and shape between the different progeny groups. The observed variation in landmark displacements was similar, and in both cases......Maternal effects on progeny wing size and shape in a homozygous parthenogenetic strain of Drosophila mercatorum were investigated. The impact of external maternal factors (heat stress) and the impact of internal maternal factors (different maternal and grand maternal age) were studied...... in landmark displacement was visualized by principal component analysis. Both kinds of maternal effects had a significant impact on progeny wing size and shape. Maternal heat stress led to the same pattern of response in size and shape among the progeny, with increased difference between the control group...

  3. Investigation and design of a C-Wing passenger aircraft

    Directory of Open Access Journals (Sweden)

    Karan BIKKANNAVAR

    2016-06-01

    Full Text Available A novel nonplanar wing concept called C-Wing is studied and implemented on a commercial aircraft to reduce induced drag which has a significant effect on fuel consumption. A preliminary sizing method which employs an optimization algorithm is utilized. The Airbus A320 aircraft is used as a reference aircraft to evaluate design parameters and to investigate the C-Wing design potential beyond current wing tip designs. An increase in aspect ratio due to wing area reduction at 36m span results in a reduction of required fuel mass by 16%. Also take-off mass savings were obtained for the aircraft with C-Wing configuration. The effect of a variations of height to span ratio (h/b of C-Wings on induced drag factor k, is formulated from a vortex lattice method and literature based equations. Finally the DOC costing methods used by the Association of European Airlines (AEA was applied to the existing A320 aircraft and to the C-Wing configuration obtaining a reduction of 6% in Direct Operating Costs (DOC for the novel concept resulted. From overall outcomes, the C-Wing concept suggests interesting aerodynamic efficiency and stability benefits.

  4. Improvement of the aerodynamic performance by wing flexibility and elytra–hind wing interaction of a beetle during forward flight

    Science.gov (United States)

    Le, Tuyen Quang; Truong, Tien Van; Park, Soo Hyung; Quang Truong, Tri; Ko, Jin Hwan; Park, Hoon Cheol; Byun, Doyoung

    2013-01-01

    In this work, the aerodynamic performance of beetle wing in free-forward flight was explored by a three-dimensional computational fluid dynamics (CFDs) simulation with measured wing kinematics. It is shown from the CFD results that twist and camber variation, which represent the wing flexibility, are most important when determining the aerodynamic performance. Twisting wing significantly increased the mean lift and camber variation enhanced the mean thrust while the required power was lower than the case when neither was considered. Thus, in a comparison of the power economy among rigid, twisting and flexible models, the flexible model showed the best performance. When the positive effect of wing interaction was added to that of wing flexibility, we found that the elytron created enough lift to support its weight, and the total lift (48.4 mN) generated from the simulation exceeded the gravity force of the beetle (47.5 mN) during forward flight. PMID:23740486

  5. Open Rotor Noise Shielding by Blended-Wing-Body Aircraft

    Science.gov (United States)

    Guo, Yueping; Czech, Michael J.; Thomas, Russell H.

    2015-01-01

    This paper presents an analysis of open rotor noise shielding by Blended Wing Body (BWB) aircraft by using model scale test data acquired in the Boeing Low Speed Aeroacoustic Facility (LSAF) with a legacy F7/A7 rotor model and a simplified BWB platform. The objective of the analysis is the understanding of the shielding features of the BWB and the method of application of the shielding data for noise studies of BWB aircraft with open rotor propulsion. By studying the directivity patterns of individual tones, it is shown that though the tonal energy distribution and the spectral content of the wind tunnel test model, and thus its total noise, may differ from those of more advanced rotor designs, the individual tones follow directivity patterns that characterize far field radiations of modern open rotors, ensuring the validity of the use of this shielding data. Thus, open rotor tonal noise shielding should be categorized into front rotor tones, aft rotor tones and interaction tones, not only because of the different directivities of the three groups of tones, but also due to the differences in their source locations and coherence features, which make the respective shielding characteristics of the three groups of tones distinctly different from each other. To reveal the parametric trends of the BWB shielding effects, results are presented with variations in frequency, far field emission angle, rotor operational condition, engine installation geometry, and local airframe features. These results prepare the way for the development of parametric models for the shielding effects in prediction tools.

  6. Cantilever Wings for Modern Aircraft: Some Aspects of Cantilever Wing Construction with Special Reference to Weight and Torsional Stiffness

    Science.gov (United States)

    Stieger, H J

    1929-01-01

    In the foregoing remarks I have made an attempt to touch on some of the structural problems met with in cantilever wings, and dealt rather fully with a certain type of single-spar construction. The experimental test wing was a first attempt to demonstrate the principles of this departure from orthodox methods. The result was a wing both torsionally stiff and of light weight - lighter than a corresponding biplane construction.

  7. Functional dissection of a neuronal network required for cuticle tanning and wing expansion in Drosophila.

    Science.gov (United States)

    Luan, Haojiang; Lemon, William C; Peabody, Nathan C; Pohl, Jascha B; Zelensky, Paul K; Wang, Ding; Nitabach, Michael N; Holmes, Todd C; White, Benjamin H

    2006-01-11

    A subset of Drosophila neurons that expresses crustacean cardioactive peptide (CCAP) has been shown previously to make the hormone bursicon, which is required for cuticle tanning and wing expansion after eclosion. Here we present evidence that CCAP-expressing neurons (NCCAP) consist of two functionally distinct groups, one of which releases bursicon into the hemolymph and the other of which regulates its release. The first group, which we call NCCAP-c929, includes 14 bursicon-expressing neurons of the abdominal ganglion that lie within the expression pattern of the enhancer-trap line c929-Gal4. We show that suppression of activity within this group blocks bursicon release into the hemolymph together with tanning and wing expansion. The second group, which we call NCCAP-R, consists of NCCAP neurons outside the c929-Gal4 pattern. Because suppression of synaptic transmission and protein kinase A (PKA) activity throughout NCCAP, but not in NCCAP-c929, also blocks tanning and wing expansion, we conclude that neurotransmission and PKA are required in NCCAP-R to regulate bursicon secretion from NCCAP-c929. Enhancement of electrical activity in NCCAP-R by expression of the bacterial sodium channel NaChBac also blocks tanning and wing expansion and leads to depletion of bursicon from central processes. NaChBac expression in NCCAP-c929 is without effect, suggesting that the abdominal bursicon-secreting neurons are likely to be silent until stimulated to release the hormone. Our results suggest that NCCAP form an interacting neuronal network responsible for the regulation and release of bursicon and suggest a model in which PKA-mediated stimulation of inputs to normally quiescent bursicon-expressing neurons activates release of the hormone.

  8. High-resolution two-dimensional gel analysis of proteins in wing imaginal discs: A data base of Drosophila

    International Nuclear Information System (INIS)

    Santaren, J.F.; Garcia-Bellido, A.

    1990-01-01

    An improved method of high-resolution two-dimensional gel electrophoresis has been used to study the patterns of protein synthesis in wing imaginal discs of late instar larvae of Drosophila melanogaster. A small number of discs were radiolabeled with a mixture of 14 C-labeled amino acids or with [ 35 S]methionine and the pattern of labeled proteins was analyzed. One thousand and twenty-five polypeptides (787 acidic (IEF) and 238 basic (NEPHGE)) from wing discs of several wild-type strains have so far been separated and cataloged. All these polypeptides have been numbered and presented in a reference map for further studies. When comparing patterns of label we have found small quantitative differences in rate of synthesis between individuals of the same strain, not due to sexual differences, and very few quantitative and qualitative differences between groups of individuals of different strains

  9. Demonstration of an in situ morphing hyperelliptical cambered span wing mechanism

    International Nuclear Information System (INIS)

    Manzo, Justin; Garcia, Ephrahim

    2010-01-01

    Research on efficient shore bird morphology inspired the hyperelliptical cambered span (HECS) wing, a crescent-shaped, aft-swept wing with vertically oriented wingtips. The wing reduces vorticity-induced circulation loss and outperforms an elliptical baseline when planar. Designed initially as a rigid wing, the HECS wing makes use of morphing to transition from a planar to a furled configuration, similar to that of a continuously curved winglet, in flight. A morphing wing concept mechanism is presented, employing shape memory alloy actuators to create a discretized curvature approximation. The aerodynamics for continuous wing shapes is validated quasi-statically through wind tunnel testing, showing enhanced planar HECS wing lift-to-drag performance over an elliptical wing, with the furled HECS wing showing minimal enhancements beyond this point. Wind tunnel tests of the active morphing wing prove the mechanism capable of overcoming realistic loading, while further testing may be required to establish aerodynamic merits of the HECS wing morphing maneuver

  10. Biomechanics of smart wings in a bat robot: morphing wings using SMA actuators

    International Nuclear Information System (INIS)

    Colorado, J; Barrientos, A; Rossi, C; Breuer, K S

    2012-01-01

    This paper presents the design of a bat-like micro aerial vehicle with actuated morphing wings. NiTi shape memory alloys (SMAs) acting as artificial biceps and triceps muscles are used for mimicking the morphing wing mechanism of the bat flight apparatus. Our objective is twofold. Firstly, we have implemented a control architecture that allows an accurate and fast SMA actuation. This control makes use of the electrical resistance measurements of SMAs to adjust morphing wing motions. Secondly, the feasibility of using SMA actuation technology is evaluated for the application at hand. To this purpose, experiments are conducted to analyze the control performance in terms of nominal and overloaded operation modes of the SMAs. This analysis includes: (i) inertial forces regarding the stretchable wing membrane and aerodynamic loads, and (ii) uncertainties due to impact of airflow conditions over the resistance–motion relationship of SMAs. With the proposed control, morphing actuation speed can be increased up to 2.5 Hz, being sufficient to generate lift forces at a cruising speed of 5 m s −1 . (paper)

  11. Biomechanics of smart wings in a bat robot: morphing wings using SMA actuators.

    Science.gov (United States)

    Colorado, J; Barrientos, A; Rossi, C; Bahlman, J W; Breuer, K S

    2012-09-01

    This paper presents the design of a bat-like micro aerial vehicle with actuated morphing wings. NiTi shape memory alloys (SMAs) acting as artificial biceps and triceps muscles are used for mimicking the morphing wing mechanism of the bat flight apparatus. Our objective is twofold. Firstly, we have implemented a control architecture that allows an accurate and fast SMA actuation. This control makes use of the electrical resistance measurements of SMAs to adjust morphing wing motions. Secondly, the feasibility of using SMA actuation technology is evaluated for the application at hand. To this purpose, experiments are conducted to analyze the control performance in terms of nominal and overloaded operation modes of the SMAs. This analysis includes: (i) inertial forces regarding the stretchable wing membrane and aerodynamic loads, and (ii) uncertainties due to impact of airflow conditions over the resistance-motion relationship of SMAs. With the proposed control, morphing actuation speed can be increased up to 2.5 Hz, being sufficient to generate lift forces at a cruising speed of 5 m s(-1).

  12. A parametric wing design study for a modern laminar flow wing

    Science.gov (United States)

    Koegler, J. A., Jr.

    1979-01-01

    The results of a parametric wing design study using a modern laminar flow airfoil designed to exhibit desirable stall characteristics while maintaining high cruise performance are presented. It was found that little is sacrificed in cruise performance when satisfying the stall margin requirements if a taper ratio of 0.65 or greater is used.

  13. SUPERDENSE MASSIVE GALAXIES IN WINGS LOCAL CLUSTERS

    International Nuclear Information System (INIS)

    Valentinuzzi, T.; D'Onofrio, M.; Fritz, J.; Poggianti, B. M.; Bettoni, D.; Fasano, G.; Moretti, A.; Omizzolo, A.; Varela, J.; Cava, A.; Couch, W. J.; Dressler, A.; Moles, M.; Kjaergaard, P.; Vanzella, E.

    2010-01-01

    Massive quiescent galaxies at z > 1 have been found to have small physical sizes, and hence to be superdense. Several mechanisms, including minor mergers, have been proposed for increasing galaxy sizes from high- to low-z. We search for superdense massive galaxies in the WIde-field Nearby Galaxy-cluster Survey (WINGS) of X-ray selected galaxy clusters at 0.04 10 M sun , are mostly S0 galaxies, have a median effective radius (R e ) = 1.61 ± 0.29 kpc, a median Sersic index (n) = 3.0 ± 0.6, and very old stellar populations with a median mass-weighted age of 12.1 ± 1.3 Gyr. We calculate a number density of 2.9 x 10 -2 Mpc -3 for superdense galaxies in local clusters, and a hard lower limit of 1.3 x 10 -5 Mpc -3 in the whole comoving volume between z = 0.04 and z = 0.07. We find a relation between mass, effective radius, and luminosity-weighted age in our cluster galaxies, which can mimic the claimed evolution of the radius with redshift, if not properly taken into account. We compare our data with spectroscopic high-z surveys and find that-when stellar masses are considered-there is consistency with the local WINGS galaxy sizes out to z ∼ 2, while a discrepancy of a factor of 3 exists with the only spectroscopic z > 2 study. In contrast, there is strong evidence for a large evolution in radius for the most massive galaxies with M * > 4 x 10 11 M sun compared to similarly massive galaxies in WINGS, i.e., the brightest cluster galaxies.

  14. Effects of boundary layer forcing on wing-tip vortices

    Science.gov (United States)

    Shaw-Ward, Samantha

    The nature of turbulence within wing-tip vortices has been a topic of research for decades, yet accurate measurements of Reynolds stresses within the core are inherently difficult due to the bulk motion wandering caused by initial and boundary conditions in wind tunnels. As a result, characterization of a vortex as laminar or turbulent is inconclusive and highly contradicting. This research uses several experimental techniques to study the effects of broadband turbulence, introduced within the wing boundary layer, on the development of wing-tip vortices. Two rectangular wings with a NACA 0012 profile were fabricated for the use of this research. One wing had a smooth finish and the other rough, introduced by P80 grade sandpaper. Force balance measurements showed a small reduction in wing performance due to surface roughness for both 2D and 3D configurations, although stall characteristics remained relatively unchanged. Seven-hole probes were purpose-built and used to assess the mean velocity profiles of the vortices five chord lengths downstream of the wing at multiple angles of attack. Above an incidence of 4 degrees, the vortices were nearly axisymmetric, and the wing roughness reduced both velocity gradients and peak velocity magnitudes within the vortex. Laser Doppler velocimetry was used to further assess the time-resolved vortex at an incidence of 5 degrees. Evidence of wake shedding frequencies and wing shear layer instabilities at higher frequencies were seen in power spectra within the vortex. Unlike the introduction of freestream turbulence, wing surface roughness did not appear to increase wandering amplitude. A new method for removing the effects of vortex wandering is proposed with the use of carefully selected high-pass filters. The filtered data revealed that the Reynolds stress profiles of the vortex produced by the smooth and rough wing were similar in shape, with a peak occurring away from the vortex centre but inside of the core. Single hot

  15. Folding in and out: passive morphing in flapping wings.

    Science.gov (United States)

    Stowers, Amanda K; Lentink, David

    2015-03-25

    We present a new mechanism for passive wing morphing of flapping wings inspired by bat and bird wing morphology. The mechanism consists of an unactuated hand wing connected to the arm wing with a wrist joint. Flapping motion generates centrifugal accelerations in the hand wing, forcing it to unfold passively. Using a robotic model in hover, we made kinematic measurements of unfolding kinematics as functions of the non-dimensional wingspan fold ratio (2-2.5) and flapping frequency (5-17 Hz) using stereo high-speed cameras. We find that the wings unfold passively within one to two flaps and remain unfolded with only small amplitude oscillations. To better understand the passive dynamics, we constructed a computer model of the unfolding process based on rigid body dynamics, contact models, and aerodynamic correlations. This model predicts the measured passive unfolding within about one flap and shows that unfolding is driven by centrifugal acceleration induced by flapping. The simulations also predict that relative unfolding time only weakly depends on flapping frequency and can be reduced to less than half a wingbeat by increasing flapping amplitude. Subsequent dimensional analysis shows that the time required to unfold passively is of the same order of magnitude as the flapping period. This suggests that centrifugal acceleration can drive passive unfolding within approximately one wingbeat in small and large wings. Finally, we show experimentally that passive unfolding wings can withstand impact with a branch, by first folding and then unfolding passively. This mechanism enables flapping robots to squeeze through clutter without sophisticated control. Passive unfolding also provides a new avenue in morphing wing design that makes future flapping morphing wings possibly more energy efficient and light-weight. Simultaneously these results point to possible inertia driven, and therefore metabolically efficient, control strategies in bats and birds to morph or recover

  16. Video change detection for fixed wing UAVs

    Science.gov (United States)

    Bartelsen, Jan; Müller, Thomas; Ring, Jochen; Mück, Klaus; Brüstle, Stefan; Erdnüß, Bastian; Lutz, Bastian; Herbst, Theresa

    2017-10-01

    In this paper we proceed the work of Bartelsen et al.1 We present the draft of a process chain for an image based change detection which is designed for videos acquired by fixed wing unmanned aerial vehicles (UAVs). From our point of view, automatic video change detection for aerial images can be useful to recognize functional activities which are typically caused by the deployment of improvised explosive devices (IEDs), e.g. excavations, skid marks, footprints, left-behind tooling equipment, and marker stones. Furthermore, in case of natural disasters, like flooding, imminent danger can be recognized quickly. Due to the necessary flight range, we concentrate on fixed wing UAVs. Automatic change detection can be reduced to a comparatively simple photogrammetric problem when the perspective change between the "before" and "after" image sets is kept as small as possible. Therefore, the aerial image acquisition demands a mission planning with a clear purpose including flight path and sensor configuration. While the latter can be enabled simply by a fixed and meaningful adjustment of the camera, ensuring a small perspective change for "before" and "after" videos acquired by fixed wing UAVs is a challenging problem. Concerning this matter, we have performed tests with an advanced commercial off the shelf (COTS) system which comprises a differential GPS and autopilot system estimating the repetition accuracy of its trajectory. Although several similar approaches have been presented,23 as far as we are able to judge, the limits for this important issue are not estimated so far. Furthermore, we design a process chain to enable the practical utilization of video change detection. It consists of a front-end of a database to handle large amounts of video data, an image processing and change detection implementation, and the visualization of the results. We apply our process chain on the real video data acquired by the advanced COTS fixed wing UAV and synthetic data. For the

  17. How the pterosaur got its wings.

    Science.gov (United States)

    Tokita, Masayoshi

    2015-11-01

    Throughout the evolutionary history of life, only three vertebrate lineages took to the air by acquiring a body plan suitable for powered flight: birds, bats, and pterosaurs. Because pterosaurs were the earliest vertebrate lineage capable of powered flight and included the largest volant animal in the history of the earth, understanding how they evolved their flight apparatus, the wing, is an important issue in evolutionary biology. Herein, I speculate on the potential basis of pterosaur wing evolution using recent advances in the developmental biology of flying and non-flying vertebrates. The most significant morphological features of pterosaur wings are: (i) a disproportionately elongated fourth finger, and (ii) a wing membrane called the brachiopatagium, which stretches from the posterior surface of the arm and elongated fourth finger to the anterior surface of the leg. At limb-forming stages of pterosaur embryos, the zone of polarizing activity (ZPA) cells, from which the fourth finger eventually differentiates, could up-regulate, restrict, and prolong expression of 5'-located Homeobox D (Hoxd) genes (e.g. Hoxd11, Hoxd12, and Hoxd13) around the ZPA through pterosaur-specific exploitation of sonic hedgehog (SHH) signalling. 5'Hoxd genes could then influence downstream bone morphogenetic protein (BMP) signalling to facilitate chondrocyte proliferation in long bones. Potential expression of Fgf10 and Tbx3 in the primordium of the brachiopatagium formed posterior to the forelimb bud might also facilitate elongation of the phalanges of the fourth finger. To establish the flight-adapted musculoskeletal morphology shared by all volant vertebrates, pterosaurs probably underwent regulatory changes in the expression of genes controlling forelimb and pectoral girdle musculoskeletal development (e.g. Tbx5), as well as certain changes in the mode of cell-cell interactions between muscular and connective tissues in the early phase of their evolution. Developmental data now

  18. Review Essay: Extremist Right-Wing Orientation and Youth Violence in the Context of Psychologically based Biography Research

    Directory of Open Access Journals (Sweden)

    Silke Baer

    2008-05-01

    Full Text Available Michaela KÖTTIG's book closes two gaps in research on right-wing political extremism. First, she deals with young women in the right-wing scene. This is a group that has been subject to little research, having generally been taken to be on the fringe of activity and with little political activity of its own. Second, KÖTTIG works exclusively with qualitative research methods, using narrative biographical interviews and reconstructive case studies which include three-generational family histories. KÖTTIG also draws on studies of clinical psychological trauma studies and attempts to reconstruct early childhood experiences. The comparison of case studies shows numerous correspondences and variations in the biographical experience of the interviewees as well as in the consequences for individual coping mechanisms. Recurrent issues include experiences of alienation from parents, the eminent importance of one grandparent, and the effects of unconscious trans-generational transmission of affect and thought patterns caused by parents' and grandparents' suppression of the family's history during the Third Reich. These and other factors can contribute to the build-up of extreme right-wing patterns of thought and behavior. KÖTTIG's insightful study shows the importance of this kind of qualitative biographical research in examining the complexity of the biographical conditions for right-wing extremism in girls and young women. URN: urn:nbn:de:0114-fqs080281

  19. How wing kinematics affect power requirements and aerodynamic force production in a robotic bat wing

    International Nuclear Information System (INIS)

    Bahlman, Joseph W; Swartz, Sharon M; Breuer, Kenneth S

    2014-01-01

    Bats display a wide variety of behaviors that require different amounts of aerodynamic force. To control and modulate aerodynamic force, bats change wing kinematics, which, in turn, may change the power required for wing motion. There are many kinematic mechanisms that bats, and other flapping animals, can use to increase aerodynamic force, e.g. increasing wingbeat frequency or amplitude. However, we do not know if there is a difference in energetic cost between these different kinematic mechanisms. To assess the relationship between mechanical power input and aerodynamic force output across different isolated kinematic parameters, we programmed a robotic bat wing to flap over a range of kinematic parameters and measured aerodynamic force and mechanical power. We systematically varied five kinematic parameters: wingbeat frequency, wingbeat amplitude, stroke plane angle, downstroke ratio, and wing folding. Kinematic values were based on observed values from free flying Cynopterus brachyotis, the species on which the robot was based. We describe how lift, thrust, and power change with increases in each kinematic variable. We compare the power costs associated with generating additional force through the four kinematic mechanisms controlled at the shoulder, and show that all four mechanisms require approximately the same power to generate a given force. This result suggests that no single parameter offers an energetic advantage over the others. Finally, we show that retracting the wing during upstroke reduces power requirements for flapping and increases net lift production, but decreases net thrust production. These results compare well with studies performed on C. brachyotis, offering insight into natural flight kinematics. (paper)

  20. Morphing Wing Design with an Innovative Three-Dimensional Warping Actuation Project

    Data.gov (United States)

    National Aeronautics and Space Administration — Advanced wing configurations where traditional control surfaces are replaced by dynamically controlled distribution of wing twist and/or camber can provide...

  1. Variable Geometry Aircraft Wing Supported by Struts And/Or Trusses

    Science.gov (United States)

    Melton, John E. (Inventor); Dudley, Michael R. (Inventor)

    2016-01-01

    The present invention provides an aircraft having variable airframe geometry for accommodating efficient flight. The aircraft includes an elongated fuselage, an oblique wing pivotally connected with said fuselage, a wing pivoting mechanism connected with said oblique wing and said fuselage, and a brace operably connected between said oblique wing and said fuselage. The present invention also provides an aircraft having an elongated fuselage, an oblique wing pivotally connected with said fuselage, a wing pivoting mechanism connected with said oblique wing and said fuselage, a propulsion system pivotally connected with said oblique wing, and a brace operably connected between said propulsion system and said fuselage.

  2. Numerical simulation of X-wing type biplane flapping wings in 3D using the immersed boundary method

    International Nuclear Information System (INIS)

    Tay, W B; Van Oudheusden, B W; Bijl, H

    2014-01-01

    The numerical simulation of an insect-sized ‘X-wing’ type biplane flapping wing configuration is performed in 3D using an immersed boundary method solver at Reynolds numbers equal to 1000 (1 k) and 5 k, based on the wing's root chord length. This X-wing type flapping configuration draws its inspiration from Delfly, a bio-inspired ornithopter MAV which has two pairs of wings flapping in anti-phase in a biplane configuration. The objective of the present investigation is to assess the aerodynamic performance when the original Delfly flapping wing micro-aerial vehicle (FMAV) is reduced to the size of an insect. Results show that the X-wing configuration gives more than twice the average thrust compared with only flapping the upper pair of wings of the X-wing. However, the X-wing's average thrust is only 40% that of the upper wing flapping at twice the stroke angle. Despite this, the increased stability which results from the smaller lift and moment variation of the X-wing configuration makes it more suited for sharp image capture and recognition. These advantages make the X-wing configuration an attractive alternative design for insect-sized FMAVS compared to the single wing configuration. In the Reynolds number comparison, the vorticity iso-surface plot at a Reynolds number of 5 k revealed smaller, finer vortical structures compared to the simulation at 1 k, due to vortices’ breakup. In comparison, the force output difference is much smaller between Re = 1 k and 5 k. Increasing the body inclination angle generates a uniform leading edge vortex instead of a conical one along the wingspan, giving higher lift. Understanding the force variation as the body inclination angle increases will allow FMAV designers to optimize the thrust and lift ratio for higher efficiency under different operational requirements. Lastly, increasing the spanwise flexibility of the wings increases the thrust slightly but decreases the efficiency. The thrust result is similar

  3. Wing design for light transport aircraft with improved fuel economy

    Energy Technology Data Exchange (ETDEWEB)

    Welte, D.; Birrenbach, R.; Haberland, W.

    An advanced technology wing has been designed for a light utility and commuter service aircraft with the requirements for economy, safety and flexibility. Trade-off studies give optimum area and aspect-ratio of the wing. A new airfoil was developed to fulfill the performance requirements. Wing planform and twist were chosen to give high maximum lift, low drag and good stall characteristics. Preset ailerons were optimized for wheel forces and lateral control. The applied aerodynamic methods, including two- and three-dimensional wind tunnel tests are shown. Various structural configurations of the wing and various flap systems are evaluated. The cantilever tapered wing and a Fowler-flap with a two-lever mechanism were found to be the most economic ones. The wing was constructed and flight-tested with a modified Dornier Do 28 Skyservant as a test bed. The new wing is being applied to a family of light transport aircraft. Finally, aircraft with the new wing are compared performancewise with contemporary aircraft.

  4. Design and Testing of Aeroelastically Tailored Wings Under Maneuver Loading

    NARCIS (Netherlands)

    Werter, N.P.M.; Sodja, J.; De Breuker, R.

    2016-01-01

    The goal of the present paper is to provide experimental validation data for the aeroelastic analysis of composite aeroelastically tailored wings with a closed-cell cross-sectional structure. Several rectangular wings with differ- ent skin thicknesses and composite layups are designed in order to

  5. Optimisation of the Sekwa blended-wing-Body research UAV

    CSIR Research Space (South Africa)

    Broughton, BA

    2008-10-01

    Full Text Available A variable stability, blended-wing-body research mini-UAV was developed at the CSIR in South Africa. The purpose of the UAV was to study some of the aerodynamic design and control issues associated with flying wing geometries and to develop a...

  6. Flapping-wing mechanical butterfly on a wheel

    Science.gov (United States)

    Godoy-Diana, Ramiro; Thiria, Benjamin; Pradal, Daniel

    2009-11-01

    We examine the propulsive performance of a flapping-wing device turning on a ``merry-go-round'' type base. The two-wing flapper is attached to a mast that is ball-bearing mounted to a central shaft in such a way that the thrust force produced by the wings makes the flapper turn around this shaft. The oscillating lift force produced by the flapping wings is aligned with the mast to avoid vibration of the system. A turning contact allows to power the motor that drives the wings. We measure power consumption and cruising speed as a function of flapping frequency and amplitude as well as wing flexibility. The design of the wings permits to change independently their flexibility in the span-wise and chord-wise directions and PIV measurements in various planes let us examine the vorticity field around the device. A complete study of the effect of wing flexibility on the propulsive performance of the system will be presented at the conference.

  7. Use of wing morphometry for the discrimination of some Cerceris ...

    African Journals Online (AJOL)

    The outline analysis, in which geometric and traditional morphometry potentials are insufficient, was performed by using the Fourier transformation. As a result of the comprehensive wing morphometry study, it was found that both Cerceris species can be distinguished according to their wing structures and the metric ...

  8. Energy-based Aeroelastic Analysis and Optimisation of Morphing Wings

    NARCIS (Netherlands)

    De Breuker, R.

    2011-01-01

    Morphing aircraft can change their shape radically when confronted with a variety of conflicting flight conditions throughout their mission. For instance the F-14 Tomcat fighter aircraft, known from the movie Top Gun, was able to sweep its wings from a straight wing configuration to a highly swept

  9. COLIBRI : A hovering flapping twin-wing robot

    NARCIS (Netherlands)

    Roshanbin, A.; Altartouri, H.; Karasek, M.; Preumont, André

    2017-01-01

    This paper describes the results of a six-year project aiming at designing and constructing a flapping twin-wing robot of the size of hummingbird (Colibri in French) capable of hovering. Our prototype has a total mass of 22 g, a wing span of 21 cm and a flapping frequency of 22 Hz; it is actively

  10. Design, Development and Testing of Shape Shifting Wing Model

    Directory of Open Access Journals (Sweden)

    Dean Ninian

    2017-11-01

    Full Text Available The design and development of morphing (shape shifting aircraft wings—an innovative technology that has the potential to increase the aerodynamic efficiency and reduce noise signatures of aircrafts—was carried out. This research was focused on reducing lift-induced drag at the flaps of the aerofoil and to improve the design to achieve the optimum aerodynamic efficiency. Simulation revealed a 10.8% coefficient of lift increase for the initial morphing wing and 15.4% for the optimized morphing wing as compared to conventional wing design. At angles of attack of 0, 5, 10 and 15 degrees, the optimized wing has an increase in lift-to-drag ratio of 18.3%, 10.5%, 10.6% and 4% respectively when compared with the conventional wing. Simulations also showed that there is a significant improvement on pressure distribution over the lower surface of the morphing wing aerofoil. The increase in flow smoothness and reduction in vortex size reduced pressure drag along the trailing edge of the wing as a result an increase in pressure on the lower surface was experienced. A morphing wing reduced the size of the vortices and therefore the noise levels measured were reduced by up to 50%.

  11. Hydraulic Evaluation of the Crest Wing Wave Energy Converter

    DEFF Research Database (Denmark)

    Kofoed, Jens Peter; Antonishen, Michael Patrick

    This report presents the results of an experimental study of the wave energy converting abilities of the Crest Wing wave energy converter (WEC). The Crest Wing is a WEC that uses its movement in matching the shape of an oncoming wave to generate power. Model tests have been performed using a scal...

  12. Unsteady Aerodynamics of Flapping Wing of a Bird

    Directory of Open Access Journals (Sweden)

    M. Agoes Moelyadi

    2013-04-01

    Full Text Available The unsteady flow behavior and time-dependent aerodynamic characteristics of the flapping motion of a bird’s wing were investigated using a computational method. During flapping, aerodynamic interactions between bird wing surfaces and surrounding flow may occur, generating local time-dependent flow changes in the flow field and aerodynamic load of birds. To study the effect of flapping speed on unsteady aerodynamic load, two kinds of computational simulations were carried out, namely a quasi-steady and an unsteady simulation. To mimic the movement of the down-stroke and the upstroke of a bird, the flapping path accorded to a sinus function, with the wing attitude changing in dihedral angle and time. The computations of time-dependent viscous flow were based on the solution of the Reynolds Averaged Navier-Stokes equations by applying the k-e turbulence model. In addition, the discretization for the computational domain around the model used multi-block structured grid to provide more accuracy in capturing viscous flow, especially in the vicinity of the wing and body surfaces, to obtain a proper wing-body geometry model. For this research, the seagull bird was chosen, which has high aspect ratio wings with pointed wing-tips and a high camber wing section. The results include mesh movement, velocity contours as well as aerodynamic coefficients of the flapping motion of the bird at various flapping frequencies.

  13. Vortex coupling in trailing vortex-wing interactions

    Science.gov (United States)

    Chen, C.; Wang, Z.; Gursul, I.

    2018-03-01

    The interaction of trailing vortices of an upstream wing with rigid and flexible downstream wings has been investigated experimentally in a wind tunnel, using particle image velocimetry, hot-wire, force, and deformation measurements. Counter-rotating upstream vortices exhibit increased meandering when they are close to the tip of the downstream wing. The upstream vortex forms a pair with the vortex shed from the downstream wing and then exhibits large displacements around the wing tip. This coupled motion of the pair has been found to cause large lift fluctuations on the downstream wing. The meandering of the vortex pair occurs at the natural meandering frequency of the isolated vortex, with a low Strouhal number, and is not affected by the frequency of the large-amplitude wing oscillations if the downstream wing is flexible. The displacement of the leading vortex is larger than that of the trailing vortex; however, it causes highly correlated variations of the core radius, core vorticity, and circulation of the trailing vortex with the coupled meandering motion. In contrast, co-rotating vortices do not exhibit any increased meandering.

  14. Closed-type wing for drones: positive and negative characteristics

    Directory of Open Access Journals (Sweden)

    Leonid I. Gretchihin

    2014-02-01

    Full Text Available The paper presents the aerodynamics of a wing of a closed oval ellipsoidal shape, designed with the use of the molecular-kinetic theory. The positive and negative characteristics of aircraft - drones with an oval wing are described. The theoretical calculations have been experimentally checked.

  15. DETERMINATION OF COMMERCIAL AIRCRAFT WING GEOMETRY DURING THE FLIGHT

    Directory of Open Access Journals (Sweden)

    V. I. Shevyakov

    2015-01-01

    Full Text Available The article deals with the task of determination of wing shape for sub-sonic commercial aircraft by photogrammetric method. It provides the procedure for measurements taken on ground and in flight. It also provides the outcome of wing twist for commercial aircraft at cruise.

  16. Global Local Structural Optimization of Transportation Aircraft Wings

    NARCIS (Netherlands)

    Ciampa, P.D.; Nagel, B.; Van Tooren, M.J.L.

    2010-01-01

    The study presents a multilevel optimization methodology for the preliminary structural design of transportation aircraft wings. A global level is defined by taking into account the primary wing structural components (i.e., ribs, spars and skin) which are explicitly modeled by shell layered finite

  17. Wing flexibility effects in clap-and-fling

    NARCIS (Netherlands)

    Percin, M.; Hu, Y.; Van Oudheusden, B.W.; Remes, B.; Scarano, F.

    2011-01-01

    The work explores the use of time-resolved tomographic PIV measurements to study a flapping-wing model, the related vortex generation mechanisms and the effect of wing flexibility on the clap-and-fling movement in particular. An experimental setup is designed and realized in a water tank by use of a

  18. 14 CFR 23.302 - Canard or tandem wing configurations.

    Science.gov (United States)

    2010-01-01

    ... 14 Aeronautics and Space 1 2010-01-01 2010-01-01 false Canard or tandem wing configurations. 23.302 Section 23.302 Aeronautics and Space FEDERAL AVIATION ADMINISTRATION, DEPARTMENT OF TRANSPORTATION AIRCRAFT AIRWORTHINESS STANDARDS: NORMAL, UTILITY, ACROBATIC, AND COMMUTER CATEGORY AIRPLANES Structure General § 23.302 Canard or tandem wing...

  19. Quantitative-genetic analysis of wing form and bilateral asymmetry ...

    Indian Academy of Sciences (India)

    Unknown

    Overall wing size was analysed here using centroid size. (defined as the square root of the sum .... For those isochromosomal lines that were common to both experimental temperatures .... subobscura reared at 18ºC. CS refers to centroid size (values in pixels2; 1 mm = 144 pixels), and WS to wing shape (all values ×. 104).

  20. Mixed ice accretion on aircraft wings

    Science.gov (United States)

    Janjua, Zaid A.; Turnbull, Barbara; Hibberd, Stephen; Choi, Kwing-So

    2018-02-01

    Ice accretion is a problematic natural phenomenon that affects a wide range of engineering applications including power cables, radio masts, and wind turbines. Accretion on aircraft wings occurs when supercooled water droplets freeze instantaneously on impact to form rime ice or runback as water along the wing to form glaze ice. Most models to date have ignored the accretion of mixed ice, which is a combination of rime and glaze. A parameter we term the "freezing fraction" is defined as the fraction of a supercooled droplet that freezes on impact with the top surface of the accretion ice to explore the concept of mixed ice accretion. Additionally we consider different "packing densities" of rime ice, mimicking the different bulk rime densities observed in nature. Ice accretion is considered in four stages: rime, primary mixed, secondary mixed, and glaze ice. Predictions match with existing models and experimental data in the limiting rime and glaze cases. The mixed ice formulation however provides additional insight into the composition of the overall ice structure, which ultimately influences adhesion and ice thickness, and shows that for similar atmospheric parameter ranges, this simple mixed ice description leads to very different accretion rates. A simple one-dimensional energy balance was solved to show how this freezing fraction parameter increases with decrease in atmospheric temperature, with lower freezing fraction promoting glaze ice accretion.

  1. Computation of Lifting Wing-Flap Configurations

    Science.gov (United States)

    Cantwell, Brian; Kwak, Dochan

    1996-01-01

    Research has been carried out on the computation of lifting wing-flap configurations. The long term goal of the research is to develop improved computational tools for the analysis and design of high lift systems. Results show that state-of-the-art computational methods are sufficient to predict time-averaged lift and overall flow field characteristics on simple high-lift configurations. Recently there has been an increased interest in the problem of airframe generated noise and experiments carried out in the 7 x 10 wind tunnel at NASA Ames have identified the flap edge as an important source of noise. A follow-on set of experiments will be conducted toward the end of 1995. The computations being carried out under this project are coordinated with these experiments. In particular, the model geometry being used in the computations is the same as that in the experiments. The geometry consists of a NACA 63-215 Mod B airfoil section which spans the 7 x lO tunnel. The wing is unswept and has an aspect ratio of two. A 30% chord Fowler flap is deployed modifications of the flap edge geometry have been shown to be effective in reducing noise and the existing code is currently being used to compute the effect of a modified geometry on the edge flow.

  2. New aeroelastic studies for a morphing wing

    Directory of Open Access Journals (Sweden)

    Ruxandra Mihaela BOTEZ*

    2012-06-01

    Full Text Available For this study, the upper surface of a rectangular finite aspect ratio wing, with a laminar airfoil cross-section, was made of a carbon-Kevlar composite material flexible skin. This flexible skin was morphed by use of Shape Memory Alloy actuators for 35 test cases characterized by combinations of Mach numbers, Reynolds numbers and angles of attack. The Mach numbers varied from 0.2 to 0.3 and the angles of attack ranged between -1° and 2°. The optimized airfoils were determined by use of the CFD XFoil code. The purpose of this aeroelastic study was to determine the flutter conditions to be avoided during wind tunnel tests. These studies show that aeroelastic instabilities for the morphing configurations considered appeared at Mach number 0.55, which was higher than the wind tunnel Mach number limit speed of 0.3. The wind tunnel tests could thus be performed safely in the 6’×9’ wind tunnel at the Institute for Aerospace Research at the National Research Council Canada (IAR/NRC, where the new aeroelastic studies, applied on morphing wings, were validated.

  3. Effects of Wing-Cuff on NACA 23015 Aerodynamic Performances

    Directory of Open Access Journals (Sweden)

    Meftah S.M.A

    2014-03-01

    Full Text Available The main subject of this work is the numerical study control of flow separation on a NACA 23015 airfoil by using wing cuff. This last is a leading edge modification done to the wing. The modification consists of a slight extension of the chord on the outboard section of the wings. Different numerical cases are considered for the baseline and modified airfoil NACA 23015 according at different angle of incidence. The turbulence is modeled by two equations k-epsilon model. The results of this numerical investigation showed several benefits of the wing cuff compared with a conventional airfoil and an agreement is observed between the experimental data and the present study. The most intriguing result of this research is the capability for wing cuff to perform short take-offs and landings.

  4. Experimental Investigation of a Wing-in-Ground Effect Craft

    Directory of Open Access Journals (Sweden)

    M. Mobassher Tofa

    2014-01-01

    Full Text Available The aerodynamic characteristics of the wing-in-ground effect (WIG craft model that has a noble configuration of a compound wing was experimentally investigated and Universiti Teknologi Malaysia (UTM wind tunnel with and without endplates. Lift and drag forces, pitching moment coefficients, and the centre of pressure were measured with respect to the ground clearance and the wing angle of attack. The ground effect and the existence of the endplates increase the wing lift-to-drag ratio at low ground clearance. The results of this research work show new proposed design of the WIG craft with compound wing and endplates, which can clearly increase the aerodynamic efficiency without compromising the longitudinal stability. The use of WIG craft is representing an ambitious technology that will help in reducing time, effort, and money of the conventional marine transportation in the future.

  5. Computational wing design studies relating to natural laminar flow

    Science.gov (United States)

    Waggoner, Edgar G.

    1986-01-01

    Two research studies are described which directly relate to the application of natural laminar flow (NLF) technology to transonic transport-type wing planforms. Each involved using state-of-the-art computational methods to design three-dimensional wing contours which generate significant runs of favorable pressure gradients. The first study supported the Variable Sweep Transition Flight Experiment and involves design of a full-span glove which extends from the leading edge to the spoiler hinge line on the upper surface of an F-14 outer wing panel. A wing was designed computationally for a corporate transport aircraft in the second study. The resulting wing design generated favorable pressure gradients from the leading edge aft to the mid-chord on both upper and lower surfaces at the cruise design point. Detailed descriptions of the computational design approach are presented along with the various constraints imposed on each of the designs.

  6. Experimental investigation of a wing-in-ground effect craft.

    Science.gov (United States)

    Tofa, M Mobassher; Maimun, Adi; Ahmed, Yasser M; Jamei, Saeed; Priyanto, Agoes; Rahimuddin

    2014-01-01

    The aerodynamic characteristics of the wing-in-ground effect (WIG) craft model that has a noble configuration of a compound wing was experimentally investigated and Universiti Teknologi Malaysia (UTM) wind tunnel with and without endplates. Lift and drag forces, pitching moment coefficients, and the centre of pressure were measured with respect to the ground clearance and the wing angle of attack. The ground effect and the existence of the endplates increase the wing lift-to-drag ratio at low ground clearance. The results of this research work show new proposed design of the WIG craft with compound wing and endplates, which can clearly increase the aerodynamic efficiency without compromising the longitudinal stability. The use of WIG craft is representing an ambitious technology that will help in reducing time, effort, and money of the conventional marine transportation in the future.

  7. Experimental Investigation of a Wing-in-Ground Effect Craft

    Science.gov (United States)

    Tofa, M. Mobassher; Ahmed, Yasser M.; Jamei, Saeed; Priyanto, Agoes; Rahimuddin

    2014-01-01

    The aerodynamic characteristics of the wing-in-ground effect (WIG) craft model that has a noble configuration of a compound wing was experimentally investigated and Universiti Teknologi Malaysia (UTM) wind tunnel with and without endplates. Lift and drag forces, pitching moment coefficients, and the centre of pressure were measured with respect to the ground clearance and the wing angle of attack. The ground effect and the existence of the endplates increase the wing lift-to-drag ratio at low ground clearance. The results of this research work show new proposed design of the WIG craft with compound wing and endplates, which can clearly increase the aerodynamic efficiency without compromising the longitudinal stability. The use of WIG craft is representing an ambitious technology that will help in reducing time, effort, and money of the conventional marine transportation in the future. PMID:24701170

  8. Osteological histology of the Pan-Alcidae (Aves, Charadriiformes): correlates of wing-propelled diving and flightlessness.

    Science.gov (United States)

    Smith, N Adam; Clarke, Julia A

    2014-02-01

    Although studies of osteological morphology, gross myology, myological histology, neuroanatomy, and wing-scaling have all documented anatomical modifications associated with wing-propelled diving, the osteohistological study of this highly derived method of locomotion has been limited to penguins. Herein we present the first osteohistological study of the derived forelimbs and hind limbs of wing-propelled diving Pan-Alcidae (Aves, Charadriiformes). In addition to detailing differences between wing-propelled diving charadriiforms and nondiving charadriiforms, microstructural modifications to the humeri, ulnae and femora of extinct flightless pan-alcids are contrasted with those of volant alcids. Histological thin-sections of four species of pan-alcids (Alca torda, †Alca grandis, †Pinguinus impennis, †Mancalla cedrosensis) and one outgroup charadriiform (Stercorarius longicaudus) were compared. The forelimb bones of wing-propelled diving charadriiforms were found to have significantly thicker (∼22%) cortical bone walls. Additionally, as in penguins, the forelimbs of flightless pan-alcids are found to be osteosclerotic. However, unlike the pattern documented in penguins that display thickened cortices in both forelimbs and hind limbs, the forelimb and hind limb elements of pan-alcids display contrasting microstructural morphologies with thickened forelimb cortices and relatively thinner femoral cortices. Additionally, the identification of medullary bone in the sampled †Pinguinus impennis specimen suggests that further osteohistological investigation could provide an answer to longstanding questions regarding sexual dimorphism of Great Auks. Finally, these results suggest that it is possible to discern volant from flightless wing-propelled divers from fragmentary fossil remains. Copyright © 2013 Wiley Periodicals, Inc.

  9. Magneto-optical Effects in the Scattering Polarization Wings of the Ca I 4227 Å Resonance Line

    Science.gov (United States)

    Alsina Ballester, E.; Belluzzi, L.; Trujillo Bueno, J.

    2018-02-01

    The linear polarization pattern produced by scattering processes in the Ca I 4227 Å resonance line is a valuable observable for probing the solar atmosphere. Via the Hanle effect, the very significant Q/I and U/I line-center signals are sensitive to the presence of magnetic fields in the lower chromosphere with strengths between 5 and 125 G, approximately. On the other hand, partial frequency redistribution (PRD) produces sizable signals in the wings of the Q/I profile, which have always been thought to be insensitive to the presence of magnetic fields. Interestingly, novel observations of this line revealed a surprising behavior: fully unexpected signals in the wings of the U/I profile and spatial variability in the wings of both Q/I and U/I. We show that the magneto-optical (MO) terms of the Stokes-vector transfer equation produce sizable signals in the wings of U/I and a clear sensitivity of the Q/I and U/I wings to the presence of photospheric magnetic fields with strengths similar to those that produce the Hanle effect in the line core. This radiative transfer investigation on the joint action of scattering processes and the Hanle and Zeeman effects in the Ca I 4227 Å line should facilitate the development of more reliable techniques for exploring the magnetism of stellar atmospheres. To this end, we can now exploit the circular polarization produced by the Zeeman effect, the magnetic sensitivity caused by the above-mentioned MO effects in the Q/I and U/I wings, and the Hanle effect in the line core.

  10. Preliminary development of a wing in ground effect vehicle

    Science.gov (United States)

    Abidin, Razali; Ahamat, Mohamad Asmidzam; Ahmad, Tarmizi; Saad, Mohd Rasdan; Hafizi, Ezzat

    2018-02-01

    Wing in ground vehicle is one of the mode of transportation that allows high speed movement over water by travelling few meters above the water level. Through this manouver strategy, a cushion of compressed air exists between the wing in ground vehicle wings and water. This significantly increase the lift force, thus reducing the necessity in having a long wing span. Our project deals with the development of wing in ground vehicle with the capability of transporting four people. The total weight of this wing in ground vehicle was estimated at 5.4 kN to enable the prediction on required wing area, minimum takeoff velocity, drag force and engine power requirement. The required takeoff velocity is decreases as the lift coefficient increases, and our current mathematical model shows the takeoff velocity at 50 m/s avoid the significant increase in lift coefficient for the wing area of 5 m2. At the velocity of 50 m/s, the drag force created by this wing in ground vehicle is well below 1 kN, which required a 100-120 kW of engine power if the propeller has the efficiency of 0.7. Assessment on the stresses and deflection of the hull structural indicate the capability of plywood to withstand the expected load. However, excessive deflection was expected in the rear section which requires a minor structural modification. In the near future, we expect that the wind tunnel tests of this wing in ground vehicle model would enable more definite prediction on the important parameters related to its performance.

  11. Inwardly rectifying potassium channels influence Drosophila wing morphogenesis by regulating Dpp release.

    Science.gov (United States)

    Dahal, Giri Raj; Pradhan, Sarala Joshi; Bates, Emily Anne

    2017-08-01

    Loss of embryonic ion channel function leads to morphological defects, but the underlying reason for these defects remains elusive. Here, we show that inwardly rectifying potassium (Irk) channels regulate release of the Drosophila bone morphogenetic protein Dpp in the developing fly wing and that this is necessary for developmental signaling. Inhibition of Irk channels decreases the incidence of distinct Dpp-GFP release events above baseline fluorescence while leading to a broader distribution of Dpp-GFP. Work by others in different cell types has shown that Irk channels regulate peptide release by modulating membrane potential and calcium levels. We found calcium transients in the developing wing, and inhibition of Irk channels reduces the duration and amplitude of calcium transients. Depolarization with high extracellular potassium evokes Dpp release. Taken together, our data implicate Irk channels as a requirement for regulated release of Dpp, highlighting the importance of the temporal pattern of Dpp presentation for morphogenesis of the wing. © 2017. Published by The Company of Biologists Ltd.

  12. Vibration receptive sensilla on the wing margins of the silkworm moth Bombyx mori.

    Science.gov (United States)

    Ai, Hiroyuki; Yoshida, Akihiro; Yokohari, Fumio

    2010-03-01

    Bristles along the wing margins (wm-bristles) of the silkworm moth, Bombyx mori, were studied morphologically and electrophysiologically. The male moth has ca. 50 wm-bristles on each forewing and hindwing. Scanning electron microscopy revealed that these wm-bristles are typical mechanosensilla. Leuco-methylene blue staining demonstrated that each wm-bristle has a single receptor neuron, which is also characteristic of the mechanosensillum. The receptor neuron responded to vibrating air currents but did not respond to a constant air current. The wm-bristles showed clear directional sensitivity to vibrating air currents. The wm-bristles were classified into two types, type I and type II, by their response patterns to sinusoidal movements of the bristle. The neuron in type I discharged bursting spikes immediately following stimulation onset and also discharged a single spike for each sinusoidal cycle for frequencies less than ca. 60 Hz. The neuron in type II only responded to vibrations over 40 Hz and, specifically at 75 Hz, discharged a single spike for each sinusoidal cycle throughout the stimulation period. These results suggest that the two types of wm-bristles are highly tuned in different ways to detect vibrations due to the wing beat. The roles of the wm-bristles in the wing beat are discussed. Copyright (c) 2009 Elsevier Ltd. All rights reserved.

  13. Population studies of Glossina pallidipes in Ethiopia: emphasis on cuticular hydrocarbons and wing morphometric analysis.

    Science.gov (United States)

    Getahun, M N; Cecchi, G; Seyoum, E

    2014-10-01

    Tsetse flies, like many insects, use pheromones for inter- and intra-specific communication. Several of their pheromones are cuticular hydrocarbons (CHCs) that are perceived by contact at close range. We hypothesized that for a successful implementation of the Sterile Insect Technique (SIT), along with proper identification of target area and target species, the target tsetse populations and the sterile flies must chemically communicate with each other. To study the population structuring of Glossina pallidipes in Ethiopia, CHCs were extracted and analyzed from three tsetse belts. As a comparative approach, wing morphometric analysis was performed. The analysis of the relative abundance of CHCs revealed that populations of G. pallidipes from the Rift Valley tsetse belt showed a distinct clustering compared to populations from the other two belts. The spatial pattern of CHC differences was complemented by the wing morphometric analysis. Our data suggest that CHCs of known biological and ecological role, when combined with wing morphometric data, will provide an alternative means for the study of population structuring of Glossina populations. This could aid the planning of area wide control strategies using SIT, which is dependent on sexual competence. Copyright © 2014 International Atomic Energy Agency 2014. Published by Elsevier B.V. All rights reserved.

  14. Patterned Anchorage to the Apical Extracellular Matrix Defines Tissue Shape in the Developing Appendages of Drosophila

    Science.gov (United States)

    Ray, Robert P.; Matamoro-Vidal, Alexis; Ribeiro, Paulo S.; Tapon, Nic; Houle, David; Salazar-Ciudad, Isaac; Thompson, Barry J.

    2015-01-01

    Summary How tissues acquire their characteristic shape is a fundamental unresolved question in biology. While genes have been characterized that control local mechanical forces to elongate epithelial tissues, genes controlling global forces in epithelia have yet to be identified. Here, we describe a genetic pathway that shapes appendages in Drosophila by defining the pattern of global tensile forces in the tissue. In the appendages, shape arises from tension generated by cell constriction and localized anchorage of the epithelium to the cuticle via the apical extracellular-matrix protein Dumpy (Dp). Altering Dp expression in the developing wing results in predictable changes in wing shape that can be simulated by a computational model that incorporates only tissue contraction and localized anchorage. Three other wing shape genes, narrow, tapered, and lanceolate, encode components of a pathway that modulates Dp distribution in the wing to refine the global force pattern and thus wing shape. PMID:26190146

  15. Aerodynamic performance of a hovering hawkmoth with flexible wings: a computational approach.

    Science.gov (United States)

    Nakata, Toshiyuki; Liu, Hao

    2012-02-22

    Insect wings are deformable structures that change shape passively and dynamically owing to inertial and aerodynamic forces during flight. It is still unclear how the three-dimensional and passive change of wing kinematics owing to inherent wing flexibility contributes to unsteady aerodynamics and energetics in insect flapping flight. Here, we perform a systematic fluid-structure interaction based analysis on the aerodynamic performance of a hovering hawkmoth, Manduca, with an integrated computational model of a hovering insect with rigid and flexible wings. Aerodynamic performance of flapping wings with passive deformation or prescribed deformation is evaluated in terms of aerodynamic force, power and efficiency. Our results reveal that wing flexibility can increase downwash in wake and hence aerodynamic force: first, a dynamic wing bending is observed, which delays the breakdown of leading edge vortex near the wing tip, responsible for augmenting the aerodynamic force-production; second, a combination of the dynamic change of wing bending and twist favourably modifies the wing kinematics in the distal area, which leads to the aerodynamic force enhancement immediately before stroke reversal. Moreover, an increase in hovering efficiency of the flexible wing is achieved as a result of the wing twist. An extensive study of wing stiffness effect on aerodynamic performance is further conducted through a tuning of Young's modulus and thickness, indicating that insect wing structures may be optimized not only in terms of aerodynamic performance but also dependent on many factors, such as the wing strength, the circulation capability of wing veins and the control of wing movements.

  16. Coloration principles of nymphaline butterflies - thin films, melanin, ommochromes and wing scale stacking.

    Science.gov (United States)

    Stavenga, Doekele G; Leertouwer, Hein L; Wilts, Bodo D

    2014-06-15

    The coloration of the common butterflies Aglais urticae (small tortoiseshell), Aglais io (peacock) and Vanessa atalanta (red admiral), belonging to the butterfly subfamily Nymphalinae, is due to the species-specific patterning of differently coloured scales on their wings. We investigated the scales' structural and pigmentary properties by applying scanning electron microscopy, (micro)spectrophotometry and imaging scatterometry. The anatomy of the wing scales appears to be basically identical, with an approximately flat lower lamina connected by trabeculae to a highly structured upper lamina, which consists of an array of longitudinal, parallel ridges and transversal crossribs. Isolated scales observed at the abwing (upper) side are blue, yellow, orange, red, brown or black, depending on their pigmentation. The yellow, orange and red scales contain various amounts of 3-OH-kynurenine and ommochrome pigment, black scales contain a high density of melanin, and blue scales have a minor amount of melanin pigment. Observing the scales from their adwing (lower) side always revealed a structural colour, which is blue in the case of blue, red and black scales, but orange for orange scales. The structural colours are created by the lower lamina, which acts as an optical thin film. Its reflectance spectrum, crucially determined by the lamina thickness, appears to be well tuned to the scales' pigmentary spectrum. The colours observed locally on the wing are also due to the degree of scale stacking. Thin films, tuned pigments and combinations of stacked scales together determine the wing coloration of nymphaline butterflies. © 2014. Published by The Company of Biologists Ltd.

  17. Mutation predicts 40 million years of fly wing evolution.

    Science.gov (United States)

    Houle, David; Bolstad, Geir H; van der Linde, Kim; Hansen, Thomas F

    2017-08-24

    Mutation enables evolution, but the idea that adaptation is also shaped by mutational variation is controversial. Simple evolutionary hypotheses predict such a relationship if the supply of mutations constrains evolution, but it is not clear that constraints exist, and, even if they do, they may be overcome by long-term natural selection. Quantification of the relationship between mutation and phenotypic divergence among species will help to resolve these issues. Here we use precise data on over 50,000 Drosophilid fly wings to demonstrate unexpectedly strong positive relationships between variation produced by mutation, standing genetic variation, and the rate of evolution over the last 40 million years. Our results are inconsistent with simple constraint hypotheses because the rate of evolution is very low relative to what both mutational and standing variation could allow. In principle, the constraint hypothesis could be rescued if the vast majority of mutations are so deleterious that they cannot contribute to evolution, but this also requires the implausible assumption that deleterious mutations have the same pattern of effects as potentially advantageous ones. Our evidence for a strong relationship between mutation and divergence in a slowly evolving structure challenges the existing models of mutation in evolution.

  18. Genomic sequence around butterfly wing development genes: annotation and comparative analysis.

    Directory of Open Access Journals (Sweden)

    Inês C Conceição

    Full Text Available BACKGROUND: Analysis of genomic sequence allows characterization of genome content and organization, and access beyond gene-coding regions for identification of functional elements. BAC libraries, where relatively large genomic regions are made readily available, are especially useful for species without a fully sequenced genome and can increase genomic coverage of phylogenetic and biological diversity. For example, no butterfly genome is yet available despite the unique genetic and biological properties of this group, such as diversified wing color patterns. The evolution and development of these patterns is being studied in a few target species, including Bicyclus anynana, where a whole-genome BAC library allows targeted access to large genomic regions. METHODOLOGY/PRINCIPAL FINDINGS: We characterize ∼1.3 Mb of genomic sequence around 11 selected genes expressed in B. anynana developing wings. Extensive manual curation of in silico predictions, also making use of a large dataset of expressed genes for this species, identified repetitive elements and protein coding sequence, and highlighted an expansion of Alcohol dehydrogenase genes. Comparative analysis with orthologous regions of the lepidopteran reference genome allowed assessment of conservation of fine-scale synteny (with detection of new inversions and translocations and of DNA sequence (with detection of high levels of conservation of non-coding regions around some, but not all, developmental genes. CONCLUSIONS: The general properties and organization of the available B. anynana genomic sequence are similar to the lepidopteran reference, despite the more than 140 MY divergence. Our results lay the groundwork for further studies of new interesting findings in relation to both coding and non-coding sequence: 1 the Alcohol dehydrogenase expansion with higher similarity between the five tandemly-repeated B. anynana paralogs than with the corresponding B. mori orthologs, and 2 the high

  19. Digital Morphing Wing: Active Wing Shaping Concept Using Composite Lattice-Based Cellular Structures.

    Science.gov (United States)

    Jenett, Benjamin; Calisch, Sam; Cellucci, Daniel; Cramer, Nick; Gershenfeld, Neil; Swei, Sean; Cheung, Kenneth C

    2017-03-01

    We describe an approach for the discrete and reversible assembly of tunable and actively deformable structures using modular building block parts for robotic applications. The primary technical challenge addressed by this work is the use of this method to design and fabricate low density, highly compliant robotic structures with spatially tuned stiffness. This approach offers a number of potential advantages over more conventional methods for constructing compliant robots. The discrete assembly reduces manufacturing complexity, as relatively simple parts can be batch-produced and joined to make complex structures. Global mechanical properties can be tuned based on sub-part ordering and geometry, because local stiffness and density can be independently set to a wide range of values and varied spatially. The structure's intrinsic modularity can significantly simplify analysis and simulation. Simple analytical models for the behavior of each building block type can be calibrated with empirical testing and synthesized into a highly accurate and computationally efficient model of the full compliant system. As a case study, we describe a modular and reversibly assembled wing that performs continuous span-wise twist deformation. It exhibits high performance aerodynamic characteristics, is lightweight and simple to fabricate and repair. The wing is constructed from discrete lattice elements, wherein the geometric and mechanical attributes of the building blocks determine the global mechanical properties of the wing. We describe the mechanical design and structural performance of the digital morphing wing, including their relationship to wind tunnel tests that suggest the ability to increase roll efficiency compared to a conventional rigid aileron system. We focus here on describing the approach to design, modeling, and construction as a generalizable approach for robotics that require very lightweight, tunable, and actively deformable structures.

  20. Repeatable Manufacture of Wings for Flapping Wing Micro Air Vehicles Using Microelectromechanical System (MEMS) Fabrication Techniques

    Science.gov (United States)

    2011-03-01

    life span, and must be cared for and used expeditiously. Once a hawkmoth hatches from its cocoon, its wing is liberated, taking care to cut the...more controlled fashion than the butterfly, but is not sufficiently so for a Micro- MAV (courtesy of http://www.science-store.com/ life /specimens/la460...50. Michelson, Robert C. and Naqvi, Messam A. Extraterrestrial Flight. s.l. : RTO- AVT von Karman Institute for Fluid Dynamics Lecture Series, 2003

  1. Control Power Optimization using Artificial Intelligence for Forward Swept Wing and Hybrid Wing Body Aircraft

    OpenAIRE

    Adegbindin, Moustaine Kolawole Agnide

    2017-01-01

    Many futuristic aircraft such as the Hybrid Wing Body have numerous control surfaces that can result in large hinge moments, high actuation power demands, and large actuator forces/moments. Also, there is no unique relationship between control inputs and the aircraft response. Distinct sets of control surface deflections may result in the same aircraft response, but with large differences in actuation power. An Artificial Neural Network and a Genetic Algorithm were used here for the control a...

  2. Ornithopter Type Flapping Wings for Autonomous Micro Air Vehicles

    Directory of Open Access Journals (Sweden)

    Sutthiphong Srigrarom

    2015-05-01

    Full Text Available In this paper, an ornithopter prototype that mimics the flapping motion of bird flight is developed, and the lift and thrust generation characteristics of different wing designs are evaluated. This project focused on the spar arrangement and material used for the wings that could achieves improved performance. Various lift and thrust measurement techniques are explored and evaluated. Various wings of insects and birds were evaluated to understand how these natural flyers with flapping wings are able to produce sufficient lift to fly. The differences in the flapping aerodynamics were also detailed. Experiments on different wing designs and materials were conducted and a paramount wing was built for a test flight. The first prototype has a length of 46.5 cm, wing span of 88 cm, and weighs 161 g. A mechanism which produced a flapping motion was fabricated and designed to create flapping flight. The flapping flight was produced by using a single motor and a flexible and light wing structure. A force balance made of load cell was then designed to measure the thrust and lift force of the ornithopter. Three sets of wings varying flexibility were fabricated, therefore lift and thrust measurements were acquired from each different set of wings. The lift will be measured in ten cycles computing the average lift and frequency in three different speeds or frequencies (slow, medium and fast. The thrust measurement was measure likewise but in two cycles only. Several observations were made regarding the behavior of flexible flapping wings that should aid in the design of future flexible flapping wing vehicles. The wings angle or phase characteristic were analyze too and studied. The final ornithopter prototype weighs only 160 g, has a wing span of 88.5 cm, that could flap at a maximum flapping frequency of 3.869 Hz, and produce a maximum thrust and lift of about 0.719 and 0.264 N respectively. Next, we proposed resonance type flapping wing utilizes the near

  3. Prediction of Wing Downwash Using CFD

    Directory of Open Access Journals (Sweden)

    Mohammed MAHDI

    2015-06-01

    Full Text Available Wing downwash study and estimation of downwash effect on the tail plane is an important task during the aircraft design process, although a lot of papers and works has been done, but the experimental work is the most important, the progress in CFD simulation has reached to the point it is able to reduce the number of runs in the wind tunnel. In this work CFD has been utilized to calculate the downwash angle and downwash gradient with respect to the angle of attack over a high aspect ratio of a typical UAV. The results of the simulation shall be used in the estimation and calculation of the longitudinal static stability analysis of the UAV.

  4. Static Aeroelastic Effects of Formation Flight for Slender Unswept Wings

    Science.gov (United States)

    Hanson, Curtis E.

    2009-01-01

    The static aeroelastic equilibrium equations for slender, straight wings are modified to incorporate the effects of aerodynamically-coupled formation flight. A system of equations is developed by applying trim constraints and is solved for component lift distribution, trim angle-of-attack, and trim aileron deflection. The trim values are then used to calculate the elastic twist distribution of the wing box. This system of equations is applied to a formation of two gliders in trimmed flight. Structural and aerodynamic properties are assumed for the gliders, and solutions are calculated for flexible and rigid wings in solo and formation flight. It is shown for a sample application of two gliders in formation flight, that formation disturbances produce greater twist in the wingtip immersed in the vortex than for either the opposing wingtip or the wings of a similar airplane in solo flight. Changes in the lift distribution, resulting from wing twist, increase the performance benefits of formation flight. A flexible wing in formation flight will require greater aileron deflection to achieve roll trim than a rigid wing.

  5. Modeling the Motion of a Flapping Wing Aerial Vehicle

    Directory of Open Access Journals (Sweden)

    Vorochaeva L.Y.

    2017-01-01

    Full Text Available The article discusses the vertical flight of a flapping wing aerial vehicle, which is also called an ornithopter. The robot is a chain of five links connected in series by active cylindrical hinges with the central link being the body and the remainder forming folding wings in pairs. The distinctive feature of this device is that the flaps of its wings imitate those of a seagull i.e. the device has a biological prototype. We construct a mathematical model of this device; much attention is given to the model of the interaction of the wings with the air environment and we determine the positions and velocities of points of application of the reduced aerodynamic forces to each of the links. Based on the results of numerical modelling of the vertical flight of the robot three modes of flight were established: ascent, hovering at a certain height and descent. The device can operate in these modes based on the oscillation parameters of the wings in particular flapping frequency and amplitude, the ratio of the amplitudes of two links and one wing and the shift of the equilibrium oscillation position of the wings relative to zero.

  6. Antibiotics, primary symbionts and wing polyphenism in three aphid species.

    Science.gov (United States)

    Hardie, Jim; Leckstein, Peter

    2007-08-01

    The possible role of the primary Buchnera symbionts in wing polyphenism is examined in three aphid species. Presumptive winged aphids were fed on antibiotic-treated beans to destroy these symbionts. As previously reported, this leads to inhibited growth and low/zero fecundity. When such treatment is applied to the short-day-induced gynoparae (the winged autumn migrant) of the black bean aphid, Aphis fabae, it also causes many insects to develop as wingless or winged/wingless intermediate adult forms (apterisation). However, whilst antibiotic treatment of crowd-induced, long-day winged forms of the pea aphid, Acyrthosiphon pisum (a green and a pink clone) and the vetch aphid, Megoura viciae has similar effects on size and fecundity, it does not affect wing development. Food deprivation also promotes apterisation in A. fabae gynoparae but not in the crowd-induced winged morphs of the other two species. Thus, it appears that apterisation in A. fabae is not a direct effect of antibiotic treatment or a novel role for symbionts but is most likely related to impaired nutrition induced by the loss of the symbiont population.

  7. Computational Optimization of a Natural Laminar Flow Experimental Wing Glove

    Science.gov (United States)

    Hartshom, Fletcher

    2012-01-01

    Computational optimization of a natural laminar flow experimental wing glove that is mounted on a business jet is presented and discussed. The process of designing a laminar flow wing glove starts with creating a two-dimensional optimized airfoil and then lofting it into a three-dimensional wing glove section. The airfoil design process does not consider the three dimensional flow effects such as cross flow due wing sweep as well as engine and body interference. Therefore, once an initial glove geometry is created from the airfoil, the three dimensional wing glove has to be optimized to ensure that the desired extent of laminar flow is maintained over the entire glove. TRANAIR, a non-linear full potential solver with a coupled boundary layer code was used as the main tool in the design and optimization process of the three-dimensional glove shape. The optimization process uses the Class-Shape-Transformation method to perturb the geometry with geometric constraints that allow for a 2-in clearance from the main wing. The three-dimensional glove shape was optimized with the objective of having a spanwise uniform pressure distribution that matches the optimized two-dimensional pressure distribution as closely as possible. Results show that with the appropriate inputs, the optimizer is able to match the two dimensional pressure distributions practically across the entire span of the wing glove. This allows for the experiment to have a much higher probability of having a large extent of natural laminar flow in flight.

  8. Flapping and flexible wings for biological and micro air vehicles

    Science.gov (United States)

    Shyy, Wei; Berg, Mats; Ljungqvist, Daniel

    1999-07-01

    Micro air vehicles (MAVs) with wing spans of 15 cm or less, and flight speed of 30-60 kph are of interest for military and civilian applications. There are two prominent features of MAV flight: (i) low Reynolds number (10 4-10 5), resulting in unfavorable aerodynamic conditions to support controlled flight, and (ii) small physical dimensions, resulting in certain favorable scaling characteristics including structural strength, reduced stall speed, and low inertia. Based on observations of biological flight vehicles, it appears that wing motion and flexible airfoils are two key attributes for flight at low Reynolds number. The small size of MAVs corresponds in nature to small birds, which do not glide like large birds, but instead flap with considerable change of wing shape during a single flapping cycle. With flapping and flexible wings, birds overcome the deteriorating aerodynamic performance under steady flow conditions by employing unsteady mechanisms. In this article, we review both biological and aeronautical literatures to present salient features relevant to MAVs. We first summarize scaling laws of biological and micro air vehicles involving wing span, wing loading, vehicle mass, cruising speed, flapping frequency, and power. Next we discuss kinematics of flapping wings and aerodynamic models for analyzing lift, drag and power. Then we present issues related to low Reynolds number flows and airfoil shape selection. Recent work on flexible structures capable of adjusting the airfoil shape in response to freestream variations is also discussed.

  9. Fabrication of corrugated artificial insect wings using laser micromachined molds

    International Nuclear Information System (INIS)

    Tanaka, Hiroto; Wood, Robert J

    2010-01-01

    This paper describes the fabrication of an artificial insect wing with a rich set of topological features by micromolding a thermosetting resin. An example 12 mm long hoverfly-like wing is fabricated with 50–125 µm vein heights and 100 µm corrugation heights. The solid veins and membrane were simultaneously formed and integrated by a single molding process. Employing a layered laser ablation technique, three-dimensional molds were created with 5 µm resolution in height. Safe demolding of the wing was achieved with a water-soluble sacrificial layer on the mold. Measured surface profiles of the wing matched those of the molds, demonstrating the high replication accuracy of this molding process. Using this process, the morphological features of insect wings can be replicated at-scale with high precision, enabling parametric experiments of the functional morphology of insect wings. This fabrication capability also makes it possible to create a variety of wing types for micro air vehicles on scales similar to insects.

  10. The design and testing of subscale smart aircraft wing bolts

    International Nuclear Information System (INIS)

    Vugampore, J M V; Bemont, C

    2012-01-01

    Presently costly periodic inspection is vital in guaranteeing the structural integrity of aircraft. This investigation assesses the potential for significantly reducing aircraft maintenance costs without modification of aircraft structures by implementing smart wing bolts, manufactured from TRIP steel, which can be monitored for damage in situ. TRIP steels undergo a transformation from paramagnetic austenite to ferromagnetic martensite during deformation. Subscale smart aircraft wing bolts were manufactured from hot rolled TRIP steel. These wing bolts were used to demonstrate that washers incorporating embedded inductance coils can be utilized to measure the martensitic transformation occurring in the TRIP steel during bolt deformation. Early in situ warning of a critical bolt stress level was thereby facilitated, potentially reducing the costly requirement for periodic wing bolt removal and inspection. The hot rolled TRIP steels that were utilized in these subscale bolts do not however exhibit the mechanical properties required of wing bolt material. Thus warm rolled TRIP steel alloys were also investigated. The mechanical properties of the best warm rolled TRIP steel alloy tested almost matched those of AISI 4340. The warm rolled alloys were also shown to exhibit transformation before yield, allowing for earlier warning when overload occurs. Further work will be required relating to fatigue crack detection, environmental temperature fluctuation and more thorough material characterization. However, present results show that in situ early detection of wing bolt overload is feasible via the use of high alloy warm rolled TRIP steel wing bolts in combination with inductive sensor embedded washers. (paper)

  11. Free vibration analysis of dragonfly wings using finite element method

    Directory of Open Access Journals (Sweden)

    M Darvizeh

    2016-04-01

    Full Text Available In the present work, investigations on the microstructure and mechanicalproperties of the dragonfly wing are carried out and numerical modelingbased on Finite Element Method (FEM is developed to predict Flightcharacteristics of dragonfly wings. Vibrational behavior of wings typestructures is immensely important in analysis, design and manufacturing ofsimilar engineering structures. For this purpose natural frequencies andmode shapes are calculated. In addition, the kind of deformation in eachmode shape evaluated and the ratio between numerical natural frequencyand experimental natural frequency presented as damping ratio. Theresults obtain from present method are in good agreement with sameexperimental methods.

  12. A Video-Based Experimental Investigation of Wing Rock

    Science.gov (United States)

    1989-08-01

    maintained a negative damping in roll (Fig. 6b). Ross concluded that wing tanks act like an aerodynamic fence, controlling flow separation over the wing...to Ross that wing rock was initiated by a nonlinear yawing moment due to sideslip, which caused a divergent Dutch roll oscillation to grow into the...20 30 40 50 e) (p (D EG) * Fig. 128 Continued S * 158 (Fig. 128a) and cycle B (Fig. 128b ) both occurred early in the 0 build-up and consisted of

  13. Aeroelastic Tailoring of Transport Wings Including Transonic Flutter Constraints

    Science.gov (United States)

    Stanford, Bret K.; Wieseman, Carol D.; Jutte, Christine V.

    2015-01-01

    Several minimum-mass optimization problems are solved to evaluate the effectiveness of a variety of novel tailoring schemes for subsonic transport wings. Aeroelastic stress and panel buckling constraints are imposed across several trimmed static maneuver loads, in addition to a transonic flutter margin constraint, captured with aerodynamic influence coefficient-based tools. Tailoring with metallic thickness variations, functionally graded materials, balanced or unbalanced composite laminates, curvilinear tow steering, and distributed trailing edge control effectors are all found to provide reductions in structural wing mass with varying degrees of success. The question as to whether this wing mass reduction will offset the increased manufacturing cost is left unresolved for each case.

  14. Winging of scapula due to serratus anterior tear

    Directory of Open Access Journals (Sweden)

    Varun Singh Kumar

    2014-10-01

    Full Text Available 【Abstract】Winging of scapula occurs most commonly due to injury to long thoracic nerve supplying serratus anterior muscle. Traumatic injury to serratus anterior muscle itself is very rare. We reported a case of traumatic winging of scapula due to tear of serratus anterior muscle in a 19-year-old male. Winging was present in neutral position and in extension of right shoulder joint but not on "push on wall" test. Patient was managed conservatively and achieved satisfactory result. Key words: Serratus anterior tear; Scapula; Wounds and injuries

  15. Dynamics and control of robotic aircraft with articulated wings

    Science.gov (United States)

    Paranjape, Aditya Avinash

    There is a considerable interest in developing robotic aircraft, inspired by birds, for a variety of missions covering reconnaissance and surveillance. Flapping wing aircraft concepts have been put forth in light of the efficiency of flapping flight at small scales. These aircraft are naturally equipped with the ability to rotate their wings about the root, a form of wing articulation. This thesis covers some problems concerning the performance, stability and control of robotic aircraft with articulated wings in gliding flight. Specifically, we are interested in aircraft without a vertical tail, which would then use wing articulation for longitudinal as well as lateral-directional control. Although the dynamics and control of articulated wing aircraft share several common features with conventional fixed wing aircraft, the presence of wing articulation presents several unique benefits as well as limitations from the perspective of performance and control. One of the objective of this thesis is to understand these features using a combination of theoretical and numerical tools. The aircraft concept envisioned in this thesis uses the wing dihedral angles for longitudinal and lateral-directional control. Aircraft with flexible articulated wings are also investigated. We derive a complete nonlinear model of the flight dynamics incorporating dynamic CG location and the changing moment of inertia. We show that symmetric dihedral configuration, along with a conventional horizontal tail, can be used to control flight speed and flight path angle independently of each other. This characteristic is very useful for initiating an efficient perching maneuver. It is shown that wing dihedral angles alone can effectively regulate sideslip during rapid turns and generate a wide range of equilibrium turn rates while maintaining a constant flight speed and regulating sideslip. We compute the turning performance limitations that arise due to the use of wing dihedral for yaw control

  16. The morphology of mouthparts, wings and genitalia of Paleozoic insect families Protohymenidae and Scytohymenidae reveals new details and supposed function.

    Science.gov (United States)

    Pecharová, Martina; Prokop, Jakub

    2018-01-01

    Megasecoptera is an extinct group of insects with specialized rostrum-like mouthparts, which is a synapomorphy shared with all members of the Late Paleozoic Palaeodictyopterida, and markedly slender wings that are unable to flex backwards. Here we describe the close up morphology of Protohymenidae and Scytohymenidae and uncover new aspects of the endoskeleton (tentorium) of the head, structure of the mouthparts with discernible proximal part of stylets controlled by muscles, surface of compound eyes that consist of a hexagonal pattern of large facets, structure and microstructures on the wings and reconstruct male and female external genitalia using ESEM and light stereomicroscopy. Furthermore, we describe Protohymen novokshonovi sp. n. based on an exceptionally well preserved fossil from the early Permian at Tshekarda in Russia, which shows crucial details, and the earliest species of Protohymenidae, Carbohymen testai gen. et sp. n. from a late Carboniferous siderite nodule at Mazon Creek in Illinois, USA. Our comparative study confirmed a set of structural and microstructural details on their wings, such as the composite anterior wing margin, development of an apical cell and the previously unknown external genitalia. Based on the results and comparison of homologous structures known primarily for extant relatives, such as mayflies and dragonflies, we outline for the first time the function of the mouthparts, in particular, the stylets, structure of the tentorium, vision provided by large hexagonal ommatidia and male copulatory structures bearing curved claspers for holding a female during copulation and penial lobes with seminal grooves. Copyright © 2017 Elsevier Ltd. All rights reserved.

  17. Sexual Dimorphism and Allometric Effects Associated With the Wing Shape of Seven Moth Species of Sphingidae (Lepidoptera: Bombycoidea).

    Science.gov (United States)

    de Camargo, Willian Rogers Ferreira; de Camargo, Nícholas Ferreira; Corrêa, Danilo do Carmo Vieira; de Camargo, Amabílio J Aires; Diniz, Ivone Rezende

    2015-01-01

    Sexual dimorphism is a pronounced pattern of intraspecific variation in Lepidoptera. However, moths of the family Sphingidae (Lepidoptera: Bombycoidea) are considered exceptions to this rule. We used geometric morphometric techniques to detect shape and size sexual dimorphism in the fore and hindwings of seven hawkmoth species. The shape variables produced were then subjected to a discriminant analysis. The allometric effects were measured with a simple regression between the canonical variables and the centroid size. We also used the normalized residuals to assess the nonallometric component of shape variation with a t-test. The deformations in wing shape between sexes per species were assessed with a regression between the nonreduced shape variables and the residuals. We found sexual dimorphism in both wings in all analyzed species, and that the allometric effects were responsible for much of the wing shape variation between the sexes. However, when we removed the size effects, we observed shape sexual dimorphism. It is very common for females to be larger than males in Lepidoptera, so it is expected that the shape of structures such as wings suffers deformations in order to preserve their function. However, sources of variation other than allometry could be a reflection of different reproductive flight behavior (long flights in search for sexual mates in males, and flight in search for host plants in females). © The Author 2015. Published by Oxford University Press on behalf of the Entomological Society of America.

  18. Manufacturing and Evaluation of a Biologically Inspired Engineered MAV Wing Compared to the Manduca Sexta Wing Under Simulated Flapping Conditions

    Science.gov (United States)

    2011-03-24

    thorax to the wings will continue for a short period of time, preserving the integrity of these wings. This small window was considered in order to...PromasterTM Digital XR EDO Aspherical LD (IF) 17-50 mm 1:2.8 Macro φ 67. Photomodeler provides the means to calibrate a camera via subroutine within...36. 20. DeLeón, N., O’Hara, R., and Palazotto, A., “Manufacturing of Engineering Bio- logically Inspired Flapping Wings,” 25th Annual US- Japan

  19. Silent and Efficient Supersonic Bi-Directional Flying Wing

    Data.gov (United States)

    National Aeronautics and Space Administration — We propose a Phase I study for a novel concept of a supersonic bi-directional (SBiDir) flying wing (FW) that has the potential to revolutionize supersonic flight...

  20. Application of SMP composite in designing a morphing wing

    Science.gov (United States)

    Yu, Kai; Yin, Weilong; Liu, Yanju; Leng, Jinsong

    2008-11-01

    A new concept of a morphing wing based on shape memory polymer (SMP) and its reinforced composite is proposed in this paper. SMP used in this study is a thermoset styrene-based resin in contrast to normal thermoplastic SMP. In our design, the wing winded on the airframe can be deployed during heating, which provides main lift for a morphing aircraft to realize stable flight. Aerodynamic characteristics of the deployed morphing wing are calculated by using CFD software. The static deformation of the wing under the air loads is also analyzed by using the finite element method. The results show that the used SMP material can provide enough strength and stiffness for the application.

  1. Toward Wing Aerostructural Optimization Using Simultaneous Analysis and Design Strategy

    NARCIS (Netherlands)

    Elham, A.; van Tooren, M.J.L.

    2017-01-01

    The application and computational efficiency of wing aerostructural optimization us- ing simultaneous analysis and design (SAND) strategy is investigated. A coupled adjoint aerostructural analysis method based on quasi-three-dimensional aerodynamic analysis is used for this research. Two different

  2. Pitching stability analysis of half-rotating wing air vehicle

    Science.gov (United States)

    Wang, Xiaoyi; Wu, Yang; Li, Qian; Li, Congmin; Qiu, Zhizhen

    2017-06-01

    Half-Rotating Wing (HRW) is a new power wing which had been developed by our work team using rotating-type flapping instead of oscillating-type flapping. Half-Rotating Wing Air Vehicle (HRWAV) is similar as Bionic Flapping Wing Air Vehicle (BFWAV). It is necessary to guarantee pitching stability of HRWAV to maintain flight stability. The working principle of HRW was firstly introduced in this paper. The rule of motion indicated that the fuselage of HRWAV without empennage would overturn forward as it generated increased pitching movement. Therefore, the empennage was added on the tail of HRWAV to balance the additional moment generated by aerodynamic force during flight. The stability analysis further shows that empennage could weaken rapidly the pitching disturbance on HRWAV and a new balance of fuselage could be achieved in a short time. Case study using numerical analysis verified correctness and validity of research results mentioned above, which could provide theoretical guidance to design and control HRWAV.

  3. Variable camber wing based on pneumatic artificial muscles

    Science.gov (United States)

    Yin, Weilong; Liu, Libo; Chen, Yijin; Leng, Jinsong

    2009-07-01

    As a novel bionic actuator, pneumatic artificial muscle has high power to weight ratio. In this paper, a variable camber wing with the pneumatic artificial muscle is developed. Firstly, the experimental setup to measure the static output force of pneumatic artificial muscle is designed. The relationship between the static output force and the air pressure is investigated. Experimental result shows the static output force of pneumatic artificial muscle decreases nonlinearly with increasing contraction ratio. Secondly, the finite element model of the variable camber wing is developed. Numerical results show that the tip displacement of the trailing-edge increases linearly with increasing external load and limited with the maximum static output force of pneumatic artificial muscles. Finally, the variable camber wing model is manufactured to validate the variable camber concept. Experimental result shows that the wing camber increases with increasing air pressure and that it compare very well with the FEM result.

  4. Comparative Analysis of Uninhibited and Constrained Avian Wing Aerodynamics

    Science.gov (United States)

    Cox, Jordan A.

    The flight of birds has intrigued and motivated man for many years. Bird flight served as the primary inspiration of flying machines developed by Leonardo Da Vinci, Otto Lilienthal, and even the Wright brothers. Avian flight has once again drawn the attention of the scientific community as unmanned aerial vehicles (UAV) are not only becoming more popular, but smaller. Birds are once again influencing the designs of aircraft. Small UAVs operating within flight conditions and low Reynolds numbers common to birds are not yet capable of the high levels of control and agility that birds display with ease. Many researchers believe the potential to improve small UAV performance can be obtained by applying features common to birds such as feathers and flapping flight to small UAVs. Although the effects of feathers on a wing have received some attention, the effects of localized transient feather motion and surface geometry on the flight performance of a wing have been largely overlooked. In this research, the effects of freely moving feathers on a preserved red tailed hawk wing were studied. A series of experiments were conducted to measure the aerodynamic forces on a hawk wing with varying levels of feather movement permitted. Angle of attack and air speed were varied within the natural flight envelope of the hawk. Subsequent identical tests were performed with the feather motion constrained through the use of externally-applied surface treatments. Additional tests involved the study of an absolutely fixed geometry mold-and-cast wing model of the original bird wing. Final tests were also performed after applying surface coatings to the cast wing. High speed videos taken during tests revealed the extent of the feather movement between wing models. Images of the microscopic surface structure of each wing model were analyzed to establish variations in surface geometry between models. Recorded aerodynamic forces were then compared to the known feather motion and surface

  5. The influence of wing morphology on the three-dimensional flow patterns of a flapping wing at bird scale

    NARCIS (Netherlands)

    Thielicke, William; Stamhuis, Eize J.

    The effect of airfoil design parameters, such as airfoil thickness and camber, are well understood in steady-state aerodynamics. But this knowledge cannot be readily applied to the flapping flight in insects and birds: flow visualizations and computational analyses of flapping flight have identified

  6. Supercritical Wing Technology: A Progress Report on Flight Evaluations

    Science.gov (United States)

    1972-01-01

    The papers in this compilation were presented at the NASA Symposium on "Supercritical Wing Technology: A Progress Report on Flight Evaluation" held at the NASA Flight Research Center, Edwards, Calif., on February 29, 1972. The purpose of the symposium was to present timely information on flight results obtained with the F-8 and T-2C supercritical wing configurations, discuss comparisons with wind-tunnel predictions, and project [ ] flight programs planned for the F-8 and F-III (TACT) airplanes.

  7. Simulating Bird Strike on Aircraft Composite Wing Leading Edge.

    OpenAIRE

    Ericsson, Max

    2012-01-01

    In this master thesis project the possibility to model the response of a wing when subjected to bird strike using finite elements is analyzed. Since this transient event lasts only a few milliseconds the used solution method is explicit time integration. The wing is manufactured using carbon fiber laminate. Carbon fiber laminates have orthotropic material properties with different stiffness in different directions. Accordingly, there are damage mechanisms not considered when using metal that ...

  8. Mathematical modeling of planar cell polarity signaling in the Drosophila melanogaster wing

    Science.gov (United States)

    Amonlirdviman, Keith

    Planar cell polarity (PCP) signaling refers to the coordinated polarization of cells within the plane of various epithelial tissues to generate sub-cellular asymmetry along an axis orthogonal to their apical-basal axes. For example, in the Drosophila wing, PCP is seen in the parallel orientation of hairs that protrude from each of the approximately 30,000 epithelial cells to robustly point toward the wing tip. Through a poorly understood mechanism, cell clones mutant for some PCP signaling components, including some, but not all alleles of the receptor frizzled, cause polarity disruptions of neighboring, wild-type cells, a phenomenon referred to as domineering nonautonomy. Previous models have proposed diffusible factors to explain nonautonomy, but no such factors have yet been found. This dissertation describes the mathematical modeling of PCP in the Drosophila wing, based on a contact dependent signaling hypothesis derived from experimental results. Intuition alone is insufficient to deduce that this hypothesis, which relies on a local feedback loop acting at the cell membrane, underlies the complex patterns observed in large fields of cells containing mutant clones, and others have argued that it cannot account for observed phenotypes. Through reaction-diffusion, partial differential equation modeling and simulation, the feedback loop is shown to fully reproduce PCP phenotypes, including domineering nonautonomy. The sufficiency of this model and the experimental validation of model predictions argue that previously proposed diffusible factors need not be invoked to explain PCP signaling and reveal how specific protein-protein interactions lead to autonomy or domineering nonautonomy. Based on these results, an ordinary differential equation model is derived to study the relationship of the feedback loop with upstream signaling components. The cadherin Fat transduces a cue to the local feedback loop, biasing the polarity direction of each cell toward the wing tip

  9. Shock/shock interactions between bodies and wings

    Directory of Open Access Journals (Sweden)

    Gaoxiang XIANG

    2018-02-01

    Full Text Available This paper examines the Shock/Shock Interactions (SSI between the body and wing of aircraft in supersonic flows. The body is simplified to a flat wedge and the wing is assumed to be a sharp wing. The theoretical spatial dimension reduction method, which transforms the 3D problem into a 2D one, is used to analyze the SSI between the body and wing. The temperature and pressure behind the Mach stem induced by the wing and body are obtained, and the wave configurations in the corner are determined. Numerical validations are conducted by solving the inviscid Euler equations in 3D with a Non-oscillatory and Non-free-parameters Dissipative (NND finite difference scheme. Good agreements between the theoretical and numerical results are obtained. Additionally, the effects of the wedge angle and sweep angle on wave configurations and flow field are considered numerically and theoretically. The influences of wedge angle are significant, whereas the effects of sweep angle on wave configurations are negligible. This paper provides useful information for the design and thermal protection of aircraft in supersonic and hypersonic flows. Keywords: Body and wing, Flow field, Hypersonic flow, Shock/shock interaction, Wave configurations

  10. Unsteady flow over flexible wings at different low Reynolds numbers

    Directory of Open Access Journals (Sweden)

    Genç Mustafa Serdar

    2016-01-01

    Full Text Available In this study, unsteady flow around flexible membrane wing which had aspect ratio of 1 (AR=1 was investigated experimentally at various Reynolds numbers (Re = 25000 and Re = 50000. Smoke-wire technique for flow visualization over the flexible membrane wing was utilized in the experiments. Digital Image Correlation system (DIC was used for measuring deformation of AR = 1 flexible membrane wing. Instantaneous deformation measurements of membrane wing were combined with the flow field measurements. In low aspect ratio flexible membrane wings, unsteadiness includes tip vortices and vortex shedding, and the combination of tip vortices. In these types of wings, complex unsteady deformations occurred due to vortex shedding. The results showed that the increasing angle of attack results in increase of membrane deformation. Moreover, it was concluded that analysis of the instantaneous deformation revealed chordwise and spanwise, modes which were due to the shedding of leading-edge vortices as well as tip vortices. Consequently, vibrational mode decreased and maximum standard deviation location approached to the trailing edge by reason of increasing angle of attack.

  11. Effect of flexibility on flapping wing characteristics under forward flight

    International Nuclear Information System (INIS)

    Zhu, Jianyang; Jiang, Lin; Zhou, Chaoying; Wang, Chao

    2014-01-01

    Through two-dimensional numerical simulation and by solving the unsteady incompressible Navier–Stokes (NS) equations, coupled with the structural dynamic equation for the motion of the wing, the effect of flexibility on flapping wing characteristics during forward flight is systematically studied. The flapping wing is considered as a cantilever, which performs the translational and rotational motion at its leading edge, and the other part is passively deformed by the aerodynamic force. The frequency ratio ω* and mass ratio m* are defined and used to characterize the flexibility of the flapping wing. It has been found that an optimal range of the frequency ratio exists in which the flexible wing possesses both a larger propulsive efficiency and lifting efficiency than their rigid counterpart. Also, the flexible wing with the smaller mass ratio may be of benefit to generate thrust, while the larger mass ratio may be of benefit to generate lift. In addition, a stronger leading edge vortex and reattachment vortex are observed around the appropriate flexibility wing’s surface, which therefore leads to better aerodynamic characteristics. (paper)

  12. Application of Piezoelectrics to Flapping-Wing MAVs

    Science.gov (United States)

    Widstrand, Alex; Hubner, J. Paul

    2015-11-01

    Micro air vehicles (MAVs) are a class of unmanned aerial vehicles that are size-restricted and operate at low velocities and low Reynolds numbers. An ongoing challenge with MAVs is that their flight-related operations are highly constrained by their size and weight, which limits battery size and, therefore, available power. One type of MAV called an ornithopter flies using flapping wings to create both lift and thrust, much like birds and insects do. Further bio-inspiration from bats led to the design of membrane wings for these vehicles, which provide aerodynamic benefits through passive vibration. In an attempt to capitalize on this vibration, a piezoelectric film, which generates a voltage when stressed, was investigated as the wing surface. Two wing planforms with constant area were designed and fabricated. The goal was to measure the wings' flight characteristics and output energy in freestream conditions. Complications with the flapper arose which prevented wind tunnel tests from being performed; however, energy data was obtained from table-top shaker tests. Preliminary results indicate that wing shape affects the magnitude of the charge generated, with a quarter-elliptic planform outperforming a rectangular planform. Funding provided by NSF REU Site Award number 1358991.

  13. Flow structure on a rotating wing undergoing deceleration to rest

    Science.gov (United States)

    Tudball Smith, Daniel; Rockwell, Donald; Sheridan, John

    2013-11-01

    Inspired by the behavior of small biological flyers and micro aerial Vehicles, this study experimentally addresses the flow structure on a low aspect ratio rotating wing at low Reynolds number. The study focuses on a wing decelerating to rest after rotating at constant velocity. The wing was set to a constant 45° angle of attack and, during the initial phase of the motion, accelerated to a constant velocity at its radius of gyration, which resulted in a Reynolds number of 1400 based on the chord length. Stereoscopic PIV was used to construct phase-averaged three-dimensional (volumetric) velocity fields that develop and relax throughout the deceleration and cessation of the wing motion. During gradual deceleration, the flow structure is maintained when normalised by the instantaneous velocity; the distinguishing feature is shedding of a trailing edge vortex that develops due to the deceleration. At higher deceleration rates to rest, the flow structure quickly degrades. Induced flow in the upstream direction along the surface of the wing causes detachment of the previously stable leading edge vortex; simultaneously, a trailing-edge vortex and the reoriented tip vortex form a co-rotating vortex pair, drawing flow downward away from the wing.

  14. Structure design of an innovative adaptive variable camber wing

    Directory of Open Access Journals (Sweden)

    Zhao An-Min

    2018-01-01

    Full Text Available In this paper, an innovative double rib sheet structure is proposed, which can replace the traditional rigid hinge joint with the surface contact. On the one hand, the variable camber wing structural design not only can improve the capacity to sustain more load but also will not increase the overall weight of the wing. On the other hand, it is a simple mechanical structure design to achieve the total wing camber change. Then the numerical simulation results show that the maximum stress at the connect of the wing rib is 88.2MPa, and the double ribs sheet engineering design meet the structural strength requirements. In addition, to make a fair comparison, the parameters of variable camber are fully referenced to the Talon Unmanned Aerial Vehicle (UAV. The results reveal that the total variable camber wing can further enhance aircraft flight efficiency by 29.4%. The design of the whole variable camber wing structure proposed in this paper has high engineering value and feasibility.

  15. Flying Wings. A New Paradigm for Civil Aviation?

    Directory of Open Access Journals (Sweden)

    R. Martinez-Val

    2007-01-01

    Full Text Available Over the last 50 years, commercial aviation has been mainly based what is currently called the conventional layout, characterized by a slender fuselage mated to a high aspect ratio wing, with aft-tail planes and pod-mounted engines under the wing. However, it seems that this primary configuration is approaching an asymptote in its productivity and performance characteristics. One of the most promising configurations for the future is the flying wing in its distinct arrangements: blended-wing-body, C-wing, tail-less aircraft, etc. These layouts might provide significant fuel savings and, hence, a decrease in pollution. This configuration would also reduce noise in take-off and landing. All this explains the great deal of activity carried out by the aircraft industry and by numerous investigators to perform feasibility and conceptual design studies of this aircraft layout to gain better knowledge of its main characteristics: productivity, airport compatibility, passenger acceptance, internal architecture, emergency evacuation, etc. The present paper discusses the main features of flying wings, their advantages over conventional competitors, and some key operational issues, such as evacuation and vortex wake intensity. 

  16. Wing-pitching mechanism of hovering Ruby-throated hummingbirds.

    Science.gov (United States)

    Song, Jialei; Luo, Haoxiang; Hedrick, Tyson L

    2015-01-19

    In hovering flight, hummingbirds reverse the angle of attack of their wings through pitch reversal in order to generate aerodynamic lift during both downstroke and upstroke. In addition, the wings may pitch during translation to further enhance lift production. It is not yet clear whether these pitching motions are caused by the wing inertia or actuated through the musculoskeletal system. Here we perform a computational analysis of the pitching dynamics by incorporating the realistic wing kinematics to determine the inertial effects. The aerodynamic effect is also included using the pressure data from a previous three-dimensional computational fluid dynamics simulation of a hovering hummingbird. The results show that like many insects, pitch reversal of the hummingbird is, to a large degree, caused by the wing inertia. However, actuation power input at the root is needed in the beginning of pronation to initiate a fast pitch reversal and also in mid-downstroke to enable a nose-up pitching motion for lift enhancement. The muscles on the wing may not necessarily be activated for pitching of the distal section. Finally, power analysis of the flapping motion shows that there is no requirement for substantial elastic energy storage or energy absorption at the shoulder joint.

  17. Wing-pitching mechanism of hovering Ruby-throated hummingbirds

    International Nuclear Information System (INIS)

    Song, Jialei; Luo, Haoxiang; Hedrick, Tyson L

    2015-01-01

    In hovering flight, hummingbirds reverse the angle of attack of their wings through pitch reversal in order to generate aerodynamic lift during both downstroke and upstroke. In addition, the wings may pitch during translation to further enhance lift production. It is not yet clear whether these pitching motions are caused by the wing inertia or actuated through the musculoskeletal system. Here we perform a computational analysis of the pitching dynamics by incorporating the realistic wing kinematics to determine the inertial effects. The aerodynamic effect is also included using the pressure data from a previous three-dimensional computational fluid dynamics simulation of a hovering hummingbird. The results show that like many insects, pitch reversal of the hummingbird is, to a large degree, caused by the wing inertia. However, actuation power input at the root is needed in the beginning of pronation to initiate a fast pitch reversal and also in mid-downstroke to enable a nose-up pitching motion for lift enhancement. The muscles on the wing may not necessarily be activated for pitching of the distal section. Finally, power analysis of the flapping motion shows that there is no requirement for substantial elastic energy storage or energy absorption at the shoulder joint. (paper)

  18. Reassessment of the wing feathers of Archaeopteryx lithographica suggests no robust evidence for the presence of elongated dorsal wing coverts.

    Directory of Open Access Journals (Sweden)

    Robert L Nudds

    Full Text Available Recently it was proposed that the primary feathers of Archaeopteryx lithographica (HMN1880 were overlaid by long covert feathers, and that a multilayered feathered wing was a feature of early fossils with feathered forelimbs. The proposed long covert feathers of Archaeopteryx were previously interpreted as dorsally displaced remiges or a second set of impressions made by the wing. The following study shows that the qualitative arguments forwarded in support of the elongated covert hypothesis are neither robust nor supported quantitatively. The idea that the extant bird wing with its single layer of overlapping primaries evolved from an earlier multilayered heavily coveted feathered forelimb as seen in Anchiornis huxleyi is reasonable. At this juncture, however, it is premature to conclude unequivocally that the wing of Archaeopteryx consisted of primary feathers overlaid with elongated coverts.

  19. Mercury concentrations in eggs of red-winged blackbirds and tree swallows breeding in Voyageurs National Park, Minnesota

    Science.gov (United States)

    Tyser, Robin W.; Rolfhus, Kristofer R.; Wiener, James G.; Windels, Steve K.; Custer, Thomas W.; Dummer, Paul

    2016-01-01

    Most investigations of the environmental effects of mercury (Hg) have focused on aquatic food webs that include piscivorous fish or wildlife. However, recent investigations have shown that other species, including passerine songbirds, may also be at risk from exposure to methylmercury (MeHg). We quantified Hg concentrations in eggs of two species of songbirds, red-winged blackbirds (Agelaius phoeniceus) and tree swallows (Tachycineta bicolor), nesting in Voyageurs National Park, Minnesota, USA. Geometric mean concentrations of total Hg (THg) were lower in red-winged blackbird eggs [218 and 107 ng/g dry weight (dw) for 2012 and 2013, respectively] than in tree swallow eggs (228 and 300 ng/g dw for 2012 and 2013, respectively), presumably reflecting differences in the trophic positions of these two species. Concentrations of MeHg averaged 98.4 % of THg in red-winged blackbird eggs. Levels of THg observed in this study were well below critical toxicological benchmarks commonly applied to eggs of avian species, suggesting these breeding populations were not adversely affected by exposure to MeHg. In red-winged blackbirds, concentrations of THg in eggs collected in 2012 were twice those in eggs collected in 2013. Hg levels in eggs of both species increased with date of clutch initiation. In red-winged blackbirds, for example, temporal patterns showed that a 3-week delay in clutch initiation increased egg THg by 60 %. These observations indicate that in ovo exposure of wetland birds to MeHg can vary significantly within nesting season as well as between years.

  20. Fuzzy Model-based Pitch Stabilization and Wing Vibration Suppression of Flexible Wing Aircraft.

    Science.gov (United States)

    Ayoubi, Mohammad A.; Swei, Sean Shan-Min; Nguyen, Nhan T.

    2014-01-01

    This paper presents a fuzzy nonlinear controller to regulate the longitudinal dynamics of an aircraft and suppress the bending and torsional vibrations of its flexible wings. The fuzzy controller utilizes full-state feedback with input constraint. First, the Takagi-Sugeno fuzzy linear model is developed which approximates the coupled aeroelastic aircraft model. Then, based on the fuzzy linear model, a fuzzy controller is developed to utilize a full-state feedback and stabilize the system while it satisfies the control input constraint. Linear matrix inequality (LMI) techniques are employed to solve the fuzzy control problem. Finally, the performance of the proposed controller is demonstrated on the NASA Generic Transport Model (GTM).

  1. The Effect of Height, Wing Length, and Wing Symmetry on Tabebuia rosea Seed Dispersal

    Directory of Open Access Journals (Sweden)

    Yasmeen Moussa

    2014-12-01

    Full Text Available The relationship between the vertical drop height and the horizontal distance traveled (dispersal ratio was investigated for a sample of fifty Tabebuia rosea seeds by dropping the seeds from five heights ranging from 1.00 to 2.00 meters. The dispersal ratio was found to be a constant 0.16 m/m for these heights. The effects of total seed length and asymmetry of seed wings on dispersal ratio were also measured using separate samples of fifty Tabebuia rosea seeds. It was found that neither seed length nor asymmetry had a significant effect on the dispersal ratio.

  2. Aerodynamic effects of corrugation and deformation in flapping wings of hovering hoverflies.

    Science.gov (United States)

    Du, Gang; Sun, Mao

    2012-05-07

    We investigated the aerodynamic effects of wing deformation and corrugation of a three-dimensional model hoverfly wing at a hovering condition by solving the Navier-Stokes equations on a dynamically deforming grid. Various corrugated wing models were tested. Insight into whether or not there existed significant aerodynamic coupling between wing deformation (camber and twist) and wing corrugation was obtained by comparing aerodynamic forces of four cases: a smooth-plate wing in flapping motion without deformation (i.e. a rigid flat-plate wing in flapping motion); a smooth-plate wing in flapping motion with deformation; a corrugated wing in flapping motion without deformation (i.e. a rigid corrugated wing in flapping motion); a corrugated wing in flapping motion with deformation. There was little aerodynamic coupling between wing deformation and corrugation: the aerodynamic effect of wing deformation and corrugation acting together was approximately a superposition of those of deformation and corrugation acting separately. When acting alone, the effect of wing deformation was to increase the lift by 9.7% and decrease the torque (or aerodynamic power) by 5.2%, and that of wing corrugation was to decrease the lift by 6.5% and increase the torque by 2.2%. But when acting together, the wing deformation and corrugation only increased the lift by ~3% and decreased the torque by ~3%. That is, the combined aerodynamic effect of deformation and corrugation is rather small. Thus, wing corrugation is mainly for structural, not aerodynamic, purpose, and in computing or measuring the aerodynamic forces, using a rigid flat-plate wing to model the corrugated deforming wing at hovering condition can be a good approximation. Copyright © 2012 Elsevier Ltd. All rights reserved.

  3. Butterfly wing colors : glass scales of Graphium sarpedon cause polarized iridescence and enhance blue/green pigment coloration of the wing membrane

    NARCIS (Netherlands)

    Stavenga, Doekele G.; Giraldo, Marco A.; Leertouwer, Hein L.

    2010-01-01

    The wings of the swordtail butterfly Graphium sarpedon nipponum contain the bile pigment sarpedobilin, which causes blue/green colored wing patches. Locally the bile pigment is combined with the strongly blue-absorbing carotenoid lutein, resulting in green wing patches and thus improving camouflage.

  4. Observations and Measurements of Wing Parameters of the Selected Beetle Species and the Design of a Mechanism Structure Implementing a Complex Wing Movement

    Science.gov (United States)

    Geisler, T.

    2016-12-01

    Beetle wings perform a flapping movement, consisting of the rotation relative to the two axes. This paper presents the results of observations and measurements of wings operating parameters in different planes of some beetle species. High speed photos and videos were used. The concept of the mechanism performing a complex wing movement was proposed and developed.

  5. Observations and Measurements of Wing Parameters of the Selected Beetle Species and the Design of a Mechanism Structure Implementing a Complex Wing Movement

    Directory of Open Access Journals (Sweden)

    Geisler T.

    2016-12-01

    Full Text Available Beetle wings perform a flapping movement, consisting of the rotation relative to the two axes. This paper presents the results of observations and measurements of wings operating parameters in different planes of some beetle species. High speed photos and videos were used. The concept of the mechanism performing a complex wing movement was proposed and developed.

  6. Induced mutations of winged bean in Ghana

    International Nuclear Information System (INIS)

    Klu, G.Y.P.; Quaynor-Addy, M.; Dinku, E.; Dikumwin, E.

    1989-01-01

    Winged bean (Psophocarpus tetragonolobus (L.) D.C.) was introduced into Ghana about two decades ago and not long after a high quality baby food was compounded from it. Germplasm collections are established at the Kade Agricultural Research Station of the University of Ghana and the University of Cape Coast. In 1980 a mutation breeding project was initiated at the University of Cape Coast under FAO/IAEA research contract and among various mutants a single erect stem mutant, a multiple branched bush type and a mutant with extra long pods were obtained. A similar programme was started at the National Nuclear Research Centre Kwabenya in 1982. Seeds of accessions UPS 122 and Kade 6/16 were gamma irradiated (100-400 Gy). In M 2 a mutant was obtained that did not flower throughout a growing period of five months. This mutant had very few leaves but developed an underground tuber weighing ca. 100 g. The parent, UPS 122, although normally tuber producing did not form tubers at Kwabenya within the period studied. In M 3 , mutants with variations in seed size and seed coat colour have been detected

  7. A new technique for investigating the induced and profile drag coefficients of a smooth wing and a tubercled wing

    Directory of Open Access Journals (Sweden)

    Bolzon Michael

    2016-01-01

    Full Text Available The induced and profile drag coefficients of a wing are typically determined through a complex experimental technique, such as wake surveying. Such a technique requires measurement of all three orthogonal components of the downstream velocity to find the components of drag, which results in the necessary usage of a sophisticated and costly measurement device, such as multi-hole pressure probe. However, in this paper data is presented which demonstrate that the relative changes in the induced and profile drag coefficients can largely be determined through the sole measurement of the downstream, streamwise velocity. To demonstrate this, the induced and profile drags of two NACA 0021 wings, one with a smooth leading edge and the other wing a tubercled leading edge for comparison, are determined through the measurement of the three orthogonal velocities. The downstream, streamwise velocity distribution of each wing is then constructed and relationships can be determined. The wings were surveyed at 3°, 9°, and 12°. It has been found that the relative magnitude of the profile drag coefficient can be found for all considered angles of attack, while the relative magnitude of the induced drag coefficient can be found at 9° and 12°. These findings produce an innovative, simpler, and more cost effective experimental technique in determining the components of drag of a wing, and reduces the burdensome requirement of a sophisticated measurement device for such an experiment. Further investigation is required to determine the induced drag at 3°.

  8. Assessing genotoxicity of diuron on Drosophila melanogaster by the wing-spot test and the wing imaginal disk comet assay.

    Science.gov (United States)

    Peraza-Vega, Ricardo I; Castañeda-Sortibrán, América N; Valverde, Mahara; Rojas, Emilio; Rodríguez-Arnaiz, Rosario

    2017-05-01

    The aim of this study was to evaluate the genotoxicity of the herbicide diuron in the wing-spot test and a novel wing imaginal disk comet assay in Drosophila melanogaster. The wing-spot test was performed with standard (ST) and high-bioactivation (HB) crosses after providing chronic 48 h treatment to third instar larvae. A positive dose-response effect was observed in both crosses, but statistically reduced spot frequencies were registered for the HB cross compared with the ST. This latter finding suggests that metabolism differences play an important role in the genotoxic effect of diuron. To verify diuron's ability to produce DNA damage, a wing imaginal disk comet assay was performed after providing 24 h diuron treatment to ST and HB third instar larvae. DNA damage induced by the herbicide had a significantly positive dose-response effect even at very low concentrations in both strains. However, as noted for the wing-spot test, a significant difference between strains was not observed that could be related to the duration of exposure between both assays. A positive correlation between the comet assay and the wing-spot test was found with regard to diuron genotoxicity.

  9. MicroRNAs of the mesothorax in Qinlingacris elaeodes, an alpine grasshopper showing a wing polymorphism with unilateral wing form.

    Science.gov (United States)

    Li, R; Jiang, G F; Ren, Q P; Wang, Y T; Zhou, X M; Zhou, C F; Qin, D Z

    2016-04-01

    MicroRNAs (miRNAs) are now recognized as key post-transcriptional regulators in regulation of phenotypic diversity. Qinlingacris elaeodes is a species of the alpine grasshopper, which is endemic to China. Adult individuals have three wing forms: wingless, unilateral-winged and short-winged. This is an ideal species to investigate the phenotypic plasticity, development and evolution of insect wings because of its case of unilateral wing form in both the sexes. We sequenced a small RNA library prepared from mesothoraxes of the adult grasshoppers using the Illumina deep sequencing technology. Approximately 12,792,458 raw reads were generated, of which the 854,580 high-quality reads were used only for miRNA identification. In this study, we identified 49 conserved miRNAs belonging to 41 families and 69 species-specific miRNAs. Moreover, seven miRNA*s were detected both for conserved miRNAs and species-specific miRNAs, which were supported by hairpin forming precursors based on polymerase chain reaction. This is the first description of miRNAs in alpine grasshoppers. The results provide a useful resource for further studies on molecular regulation and evolution of miRNAs in grasshoppers. These findings not only enrich the miRNAs for insects but also lay the groundwork for the study of post-transcriptional regulation of wing forms.

  10. Physiological trade-off between cellular immunity and flight capability in the wing-dimorphic cricket, Gryllus firmus

    Science.gov (United States)

    The sand cricket, Gryllus firmus, is a wing-dimorphic species with long-wing (LW) and short wing (LW) morphs. The LW forms have very well developed wings and flight muscles and their SW counterparts have reduced wings and flight muscles, coupled with greater resource allocations to reproduction. Thi...

  11. Pulsed eddy current inspection of CF-188 inner wing spar

    Science.gov (United States)

    Horan, Peter Francis

    Royal Canadian Air Force (RCAF) CF-188 Hornet aircraft engineering authorities have stated a requirement for a Non-Destructive Evaluation (NDE) technique to detect Stress Corrosion Cracking (SCC) in the inner wing spars without fastener or composite wing skin removal. Current radiographic inspections involve significant aircraft downtime, and Pulsed Eddy Current (PEC) inspection is proposed as a solution. The aluminum inner wing spars of CF-188 Hornet aircraft may undergo stress corrosion cracking (SCC) along the spar between the fasteners that secure carbon-fiber/ epoxy composite skin to the wing. Inspection of the spar through the wing skin is required to avoid wing disassembly. The thickness of the wing skin varies between 8 and 20 mm (0.3 to 0.8 inch) and fasteners may be either titanium or ferrous. PEC generated by a probe centered over a fastener, demonstrates capability of detecting simulated cracks within spars with the wing skin present. Comparison of signals from separate sensors, mounted to either side of the excitation coil, is used to detect differences in induced eddy current fields, which arise in the presence of cracks. To overcome variability in PEC signal response due to variation in 1) skin thickness, 2) fastener material and size, and 3) centering over fasteners, a large calibration data set is acquired. Multi-dimensional scores from a Modified Principal Components Analysis (PCA) of the data are reduced to one dimension (1D) using a Discriminant Analysis method. Under inspection conditions, calibrated PCA scores combined with discriminant analysis permit rapid real time go/no-go PEC detection of cracks in CF-188 inner wing spar. Probe designs using both pickup coils and Giant Magnetoresistive (GMR) sensors were tested on samples with the same ferrous and titanium fasteners found on the CF-188. Flaws were correctly detected at lift-offs of up to 21mm utilizing a variety of insulating skin materials simulating the carbon-fibre reinforced polymer

  12. Investigation of asymmetry of vortex flow over slender delta wings

    Science.gov (United States)

    Atashbaz, Ghasem

    Vortex flow, a major area of interest in fluid mechanics, is widespread in nature and in many man-made fluid mechanical devices. It can create havoc as cyclones or tornadoes or have significant implications in the performance of turbo-fluid machines or supersonic vehicles and so forth. Asymmetric vortices can cause a loss of lift and increase in rolling moment which can significantly affect wing stability and control. Up until the early nineties, it was generally believed that vortex asymmetry was the result of vortex interactions due to the close proximity of vortices over slender delta wings. However, some recent studies have thrown considerable doubt on the validity of this hypothesis. As a result, wind tunnel investigations were conducted on a series of nine delta wing planforms with sharp and round leading edges to examine the occurrence of vortex asymmetry at different angles of attack and sideslip. The study included surface oil and laser light sheet flow visualization in addition to surface pressure and hot-wire velocity measurements under static conditions. The effects of incidence, sideslip and sweep angles as well as Reynolds number variations were investigated. In this study, it was found that the effect of apex and leading edge shape played an important role in vortex asymmetry generation at high angle of attack. Vortex asymmetry was not observed over slender sharp leading edge delta wings due to the separation point being fixed at the sharp leading edge. Experimental results for these wings showed that the vortices do not impinge on one another because they do not get any closer beyond a certain value of angle of attack. Thus vortex asymmetry was not generated. However, significant vortex asymmetry was observed for round leading-edged delta wings. Asymmetric separation positions over the round leading edge was the result of laminar/turbulent transition which caused vortex asymmetry on these delta wing configurations. Sideslip angle and vortex

  13. Performance Assessment in a Heat Exchanger Tube with Opposite/Parallel Wing Twisted Tapes

    Directory of Open Access Journals (Sweden)

    S. Eiamsa-ard

    2015-02-01

    Full Text Available The thermohydraulic performance in a tube containing a modified twisted tape with alternate-axes and wing arrangements is reported. This work aims to investigate the effects of wing arrangements (opposite (O and parallel (P wings at different wing shapes (triangle (Tri, rectangular (Rec, and trapezoidal (Tra wings and on the thermohydraulic performance characteristics. The obtained results show that wing twisted tapes with all wing shape arrangements (O-Tri/O-Rec/O-Tra/P-Tri/P-Rec/P-Tra give superior thermohydraulic performance and heat transfer rate to the typical twisted tape. In addition, the tapes with opposite wing arrangement of O-Tra, O-Rec, and O-Tri give superior thermohydraulic performances to those with parallel wing arrangement of P-Tra, P-Rec, and P-Tri around 2.7%, 3.5%, and 3.2%, respectively.

  14. Heat transfer and oil flow studies on a single-stage-to-orbit control-configured winged entry vehicle

    Science.gov (United States)

    Helms, V. T., III; Bradley, P. F.

    1984-01-01

    Results are presented for oil flow and phase change paint heat transfer tests conducted on a 0.006 scale model of a proposed single stage to orbit control configured vehicle. The data were taken at angles of attack up to 40 deg at a free stream Mach number of 10 for Reynolds numbers based on model length of 0.5 x 10 to the 6th power, 1.0 x 10 to the 6th power and 2.0 x 10 to the 6th power. The magnitude and distribution of heating are characterized in terms of angle of attack and Reynolds number aided by an analysis of the flow data which are used to suggest the presence of various three dimensional flow structures that produce the observed heating patterns. Of particular interest are streak heating patterns that result in high localized heat transfer rates on the wing windward surface at low to moderate angles of attack. These streaks are caused by the bow-shock/wing-shock interaction and formation of the wing-shock. Embedded vorticity was found to be associated with these interactions.

  15. Analysis of Low Speed Stall Aerodynamics of a Swept Wing with Laminar Flow Glove

    Science.gov (United States)

    Bui, Trong T.

    2014-01-01

    Reynolds-Averaged Navier-Stokes (RANS) computational fluid dynamics (CFD) analysis was conducted to study the low-speed stall aerodynamics of a GIII aircraft's swept wing modified with a laminar-flow wing glove. The stall aerodynamics of the gloved wing were analyzed and compared with the unmodified wing for the flight speed of 120 knots and altitude of 2300 ft above mean sea level (MSL). The Star-CCM+ polyhedral unstructured CFD code was first validated for wing stall predictions using the wing-body geometry from the First American Institute of Aeronautics and Astronautics (AIAA) CFD High-Lift Prediction Workshop. It was found that the Star-CCM+ CFD code can produce results that are within the scattering of other CFD codes considered at the workshop. In particular, the Star-CCM+ CFD code was able to predict wing stall for the AIAA wing-body geometry to within 1 degree of angle of attack as compared to benchmark wind-tunnel test data. Current results show that the addition of the laminar-flow wing glove causes the gloved wing to stall much earlier than the unmodified wing. Furthermore, the gloved wing has a different stall characteristic than the clean wing, with no sharp lift drop-off at stall for the gloved wing.

  16. Analysis of Low-Speed Stall Aerodynamics of a Swept Wing with Laminar-Flow Glove

    Science.gov (United States)

    Bui, Trong T.

    2014-01-01

    Reynolds-Averaged Navier-Stokes (RANS) computational fluid dynamics (CFD) analysis was conducted to study the low-speed stall aerodynamics of a GIII aircraft's swept wing modified with a laminar-flow wing glove. The stall aerodynamics of the gloved wing were analyzed and compared with the unmodified wing for the flight speed of 120 knots and altitude of 2300 ft above mean sea level (MSL). The Star-CCM+ polyhedral unstructured CFD code was first validated for wing stall predictions using the wing-body geometry from the First American Institute of Aeronautics and Astronautics (AIAA) CFD High-Lift Prediction Workshop. It was found that the Star-CCM+ CFD code can produce results that are within the scattering of other CFD codes considered at the workshop. In particular, the Star-CCM+ CFD code was able to predict wing stall for the AIAA wing-body geometry to within 1 degree of angle of attack as compared to benchmark wind-tunnel test data. Current results show that the addition of the laminar-flow wing glove causes the gloved wing to stall much earlier than the unmodified wing. Furthermore, the gloved wing has a different stall characteristic than the clean wing, with no sharp lift drop-off at stall for the gloved wing.

  17. Effects of flexibility and aspect ratio on the aerodynamic performance of flapping wings.

    Science.gov (United States)

    Fu, Junjiang; Liu, Xiaohui; Shyy, Wei; Qiu, Huihe

    2018-01-26

    In the current study, we experimentally investigated the flexibility effects on the aerodynamic performance of flapping wings and the correlation with aspect ratio at angle of attack α = 45o. The Reynolds number based on the chord length and the wing tip velocity is maintained at Re = 5.3x103. Our result for compliant wings with an aspect ratio of 4 shows that wing flexibility can offer improved aerodynamic performance compared to that of a rigid wing. Flexible wings are found to offer higher lift-to-drag ratios; in particular, there is significant reduction in drag with little compromise in lift. The mechanism of the flexibility effects on the aerodynamic performance is addressed by quantifying the aerodynamic lift and drag forces, the transverse displacement on the wings and the flow field around the wings. The regime of the effective stiffness that offers improved aerodynamic performance is quantified in a range of about 0.5~10 and it matches the stiffness of insect wings with similar aspect ratios. Furthermore, we find that the aspect ratio of the wing is the predominant parameter determining the flexibility effects of compliant wings. Compliant wings with an aspect ratio of two do not demonstrate improved performance compared to their rigid counterparts throughout the entire stiffness regime investigated. The correlation between wing flexibility effects and the aspect ratio is supported by the stiffness of real insect wings. © 2018 IOP Publishing Ltd.

  18. Microarray Data Analysis of Space Grown Arabidopsis Leaves for Genes Important in Vascular Patterning

    Science.gov (United States)

    Weitzeal, A. J.; Wyatt, S. E.; Parsons-Wingerter, P.

    2016-01-01

    Venation patterning in leaves is a major determinant of photosynthesis efficiency because of its dependency on vascular transport of photoassimilates, water, and minerals. Arabidopsis thaliana grown in microgravity show delayed growth and leaf maturation. Gene expression data from the roots, hypocotyl, and leaves of A. thaliana grown during spaceflight vs. ground control analyzed by Affymetrix microarray are available through NASAs GeneLab (GLDS-7). We analyzed the data for differential expression of genes in leaves resulting from the effects of spaceflight on vascular patterning. Two genes were found by preliminary analysis to be upregulated during spaceflight that may be related to vascular formation. The genes are responsible for coding an ARGOS like protein (potentially affecting cell elongation in the leaves), and an F-boxkelch-repeat protein (possibly contributing to protoxylem specification). Further analysis that will focus on raw data quality assessment and a moderated t-test may further confirm upregulation of the two genes and/or identify other gene candidates. Plants defective in these genes will then be assessed for phenotype by the mapping and quantification of leaf vascular patterning by NASAs VESsel GENeration (VESGEN) software to model specific vascular differences of plants grown in spaceflight.

  19. Classification of Camellia (Theaceae) species using leaf architecture variations and pattern recognition techniques.

    Science.gov (United States)

    Lu, Hongfei; Jiang, Wu; Ghiassi, M; Lee, Sean; Nitin, Mantri

    2012-01-01

    Leaf characters have been successfully utilized to classify Camellia (Theaceae) species; however, leaf characters combined with supervised pattern recognition techniques have not been previously explored. We present results of using leaf morphological and venation characters of 93 species from five sections of genus Camellia to assess the effectiveness of several supervised pattern recognition techniques for classifications and compare their accuracy. Clustering approach, Learning Vector Quantization neural network (LVQ-ANN), Dynamic Architecture for Artificial Neural Networks (DAN2), and C-support vector machines (SVM) are used to discriminate 93 species from five sections of genus Camellia (11 in sect. Furfuracea, 16 in sect. Paracamellia, 12 in sect. Tuberculata, 34 in sect. Camellia, and 20 in sect. Theopsis). DAN2 and SVM show excellent classification results for genus Camellia with DAN2's accuracy of 97.92% and 91.11% for training and testing data sets respectively. The RBF-SVM results of 97.92% and 97.78% for training and testing offer the best classification accuracy. A hierarchical dendrogram based on leaf architecture data has confirmed the morphological classification of the five sections as previously proposed. The overall results suggest that leaf architecture-based data analysis using supervised pattern recognition techniques, especially DAN2 and SVM discrimination methods, is excellent for identification of Camellia species.

  20. Aerodynamics and Ecomorphology of Flexible Feathers and Morphing Bird Wings

    Science.gov (United States)

    Klaassen van Oorschot, Brett

    Birds are talented fliers capable of vertical take-off and landing, navigating turbulent air, and flying thousands of miles without rest. How is this possible? What allows birds to exploit the aerial environment with such ease? In part, it may be because bird wings are unlike any engineered wing. They are flexible, strong, lightweight, and dynamically capable of changes in shape on a nearly instantaneous basis (Rayner, 1988; Tobalske, 2007). Moreover, much of this change is passive, modulated only by changes in airflow angle and velocity. Birds actively morph their wings and their feathers morph passively in response to airflow to meet aerodynamic demands. Wings are highly adapted to myriad aeroecological factors and aerodynamic conditions (e.g. Lockwood et al., 1998; Bowlin and Winkler, 2004). This dissertation contains the results of my research on the complexities of morphing avian wings and feathers. I chose to study three related-but-discrete aspects of the avian wing: 1) the aerodynamics of morphing wings during take-off and gliding flight, 2) the presence and significance of wing tip slots across the avian clade, and 3) the aerodynamic role of the emarginate primary feathers that form these wing tip slots. These experiments ask fundamental questions that have intrigued me since childhood: Why do birds have different wing shapes? And why do some birds have slotted wing tips? It's fair to say that you will not find definitive answers here--rather, you will find the methodical, incremental addition of new hypotheses and empirical evidence which will serve future researchers in their own pursuits of these questions. The first chapter explores active wing morphing in two disparate aerodynamic regimes: low-advance ratio flapping (such as during takeoff) and high-advance ratio gliding. This chapter was published in the Journal of Experimental Biology (Klaassen van Oorschot et al., 2016) with the help of an undergraduate researcher, Emily Mistick. We found that wing

  1. Assessment of the correlation between wing size and body weight in captive Culex quinquefasciatus

    Directory of Open Access Journals (Sweden)

    Vivian Petersen

    Full Text Available Abstract: INTRODUCTION: Mass production of mosquitoes under laboratory conditions allows implementing methods to control vector mosquitoes. Colony development depends on mosquito size and weight. Body size can be estimated from its correlation with wing size, whereas weight is more difficult to determine. Our goal was to test whether wing size can predict the weight. METHODS: We compared dry weight and wing centroid size of Culex quinquefasciatus reared at different temperatures and four diets. RESULTS: Weight and wing size were strongly correlated. The diets did not influence wing size. CONCLUSIONS: Wing centroid size is a good predictor of Cx. quinquefasciatus body weight.

  2. Analysis of Low-Speed Stall Aerodynamics of a Business Jets Wing Using STAR-CCM+

    Science.gov (United States)

    Bui, Trong

    2016-01-01

    Reynolds-Averaged Navier-Stokes (RANS) computational fluid dynamics (CFD) analysis was conducted: to study the low-speed stall aerodynamics of a GIII aircrafts swept wing modified with (1) a laminar-flow wing glove, or (2) a seamless flap. The stall aerodynamics of these two different wing configurations were analyzed and compared with the unmodified baseline wing for low-speed flight. The Star-CCM+ polyhedral unstructured CFD code was first validated for wing stall predictions using the wing-body geometry from the First AIAA CFD High-Lift Prediction Workshop.

  3. Assessment of the correlation between wing size and body weight in captive Culex quinquefasciatus.

    Science.gov (United States)

    Petersen, Vivian; Marchi, Marco Jacometto; Natal, Delsio; Marrelli, Mauro Toledo; Barbosa, Admilson Clayton; Suesdek, Lincoln

    2016-01-01

    Mass production of mosquitoes under laboratory conditions allows implementing methods to control vector mosquitoes. Colony development depends on mosquito size and weight. Body size can be estimated from its correlation with wing size, whereas weight is more difficult to determine. Our goal was to test whether wing size can predict the weight. We compared dry weight and wing centroid size of Culex quinquefasciatus reared at different temperatures and four diets. Weight and wing size were strongly correlated. The diets did not influence wing size. Wing centroid size is a good predictor of Cx. quinquefasciatus body weight.

  4. Aeroelasticity Benchmark Assessment: Subsonic Fixed Wing Program

    Science.gov (United States)

    Florance, Jennifer P.; Chwalowski, Pawel; Wieseman, Carol D.

    2010-01-01

    Aeroelasticity Branch will examine other experimental efforts within the Subsonic Fixed Wing (SFW) program (such as testing of the NASA Common Research Model (CRM)) and other NASA programs and assess aeroelasticity issues and research topics.

  5. Aerodynamics of wing-assisted incline running in birds.

    Science.gov (United States)

    Tobalske, Bret W; Dial, Kenneth P

    2007-05-01

    Wing-assisted incline running (WAIR) is a form of locomotion in which a bird flaps its wings to aid its hindlimbs in climbing a slope. WAIR is used for escape in ground birds, and the ontogeny of this behavior in precocial birds has been suggested to represent a model analogous to transitional adaptive states during the evolution of powered avian flight. To begin to reveal the aerodynamics of flap-running, we used digital particle image velocimetry (DPIV) and measured air velocity, vorticity, circulation and added mass in the wake of chukar partridge Alectoris chukar as they engaged in WAIR (incline 65-85 degrees; N=7 birds) and ascending flight (85 degrees, N=2). To estimate lift and impulse, we coupled our DPIV data with three-dimensional wing kinematics from a companion study. The ontogeny of lift production was evaluated using three age classes: baby birds incapable of flight [6-8 days post hatching (d.p.h.)] and volant juveniles (25-28 days) and adults (45+ days). All three age classes of birds, including baby birds with partially emerged, symmetrical wing feathers, generated circulation with their wings and exhibited a wake structure that consisted of discrete vortex rings shed once per downstroke. Impulse of the vortex rings during WAIR was directed 45+/-5 degrees relative to horizontal and 21+/-4 degrees relative to the substrate. Absolute values of circulation in vortex cores and induced velocity increased with increasing age. Normalized circulation was similar among all ages in WAIR but 67% greater in adults during flight compared with flap-running. Estimated lift during WAIR was 6.6% of body weight in babies and between 63 and 86% of body weight in juveniles and adults. During flight, average lift was 110% of body weight. Our results reveal for the first time that lift from the wings, rather than wing inertia or profile drag, is primarily responsible for accelerating the body toward the substrate during WAIR, and that partially developed wings, not yet

  6. Age-class separation of blue-winged ducks

    Science.gov (United States)

    Hohman, W.L.; Moore, J.L.; Twedt, D.J.; Mensik, John G.; Logerwell, E.

    1995-01-01

    Accurate determination of age is of fundamental importance to population and life history studies of waterfowl and their management. Therefore, we developed quantitative methods that separate adult and immature blue-winged teal (Anas discors), cinnamon teal (A. cyanoptera), and northern shovelers (A. clypeata) during spring and summer. To assess suitability of discriminant models using 9 remigial measurements, we compared model performance (% agreement between predicted age and age assigned to birds on the basis of definitive cloacal or rectral feather characteristics) in different flyways (Mississippi and Pacific) and between years (1990-91 and 1991-92). We also applied age-classification models to wings obtained from U.S. Fish and Wildlife Service harvest surveys in the Mississippi and Central-Pacific flyways (wing-bees) for which age had been determined using qualitative characteristics (i.e., remigial markings, shape, or wear). Except for male northern shovelers, models correctly aged lt 90% (range 70-86%) of blue-winged ducks. Model performance varied among species and differed between sexes and years. Proportions of individuals that were correctly aged were greater for males (range 63-86%) than females (range 39-69%). Models for northern shovelers performed better in flyway comparisons within year (1991-92, La. model applied to Calif. birds, and Calif. model applied to La. birds: 90 and 94% for M, and 89 and 76% for F, respectively) than in annual comparisons within the Mississippi Flyway (1991-92 model applied to 1990-91 data: 79% for M, 50% for F). Exclusion of measurements that varied by flyway or year did not improve model performance. Quantitative methods appear to be of limited value for age separation of female blue-winged ducks. Close agreement between predicted age and age assigned to wings from the wing-bees suggests that qualitative and quantitative methods may be equally accurate for age separation of male blue-winged ducks. We interpret annual

  7. Towards long term cultivation of Drosophila wing imaginal discs in vitro.

    Science.gov (United States)

    Handke, Björn; Szabad, János; Lidsky, Peter V; Hafen, Ernst; Lehner, Christian F

    2014-01-01

    The wing imaginal disc of Drosophila melanogaster is a prominent experimental system for research on control of cell growth, proliferation and death, as well as on pattern formation and morphogenesis during organogenesis. The precise genetic methodology applicable in this system has facilitated conceptual advances of fundamental importance for developmental biology. Experimental accessibility and versatility would gain further if long term development of wing imaginal discs could be studied also in vitro. For example, culture systems would allow live imaging with maximal temporal and spatial resolution. However, as clearly demonstrated here, standard culture methods result in a rapid cell proliferation arrest within hours of cultivation of dissected wing imaginal discs. Analysis with established markers for cells in S- and M phase, as well as with RGB cell cycle tracker, a novel reporter transgene, revealed that in vitro cultivation interferes with cell cycle progression throughout interphase and not just exclusively during G1. Moreover, quantification of EGFP expression from an inducible transgene revealed rapid adverse effects of disc culture on basic cellular functions beyond cell cycle progression. Disc transplantation experiments confirmed that these detrimental consequences do not reflect fatal damage of imaginal discs during isolation, arguing clearly for a medium insufficiency. Alternative culture media were evaluated, including hemolymph, which surrounds imaginal discs during growth in situ. But isolated larval hemolymph was found to be even less adequate than current culture media, presumably as a result of conversion processes during hemolymph isolation or disc culture. The significance of prominent growth-regulating pathways during disc culture was analyzed, as well as effects of insulin and disc co-culture with larval tissues as potential sources of endocrine factors. Based on our analyses, we developed a culture protocol that prolongs cell

  8. Towards long term cultivation of Drosophila wing imaginal discs in vitro.

    Directory of Open Access Journals (Sweden)

    Björn Handke

    Full Text Available The wing imaginal disc of Drosophila melanogaster is a prominent experimental system for research on control of cell growth, proliferation and death, as well as on pattern formation and morphogenesis during organogenesis. The precise genetic methodology applicable in this system has facilitated conceptual advances of fundamental importance for developmental biology. Experimental accessibility and versatility would gain further if long term development of wing imaginal discs could be studied also in vitro. For example, culture systems would allow live imaging with maximal temporal and spatial resolution. However, as clearly demonstrated here, standard culture methods result in a rapid cell proliferation arrest within hours of cultivation of dissected wing imaginal discs. Analysis with established markers for cells in S- and M phase, as well as with RGB cell cycle tracker, a novel reporter transgene, revealed that in vitro cultivation interferes with cell cycle progression throughout interphase and not just exclusively during G1. Moreover, quantification of EGFP expression from an inducible transgene revealed rapid adverse effects of disc culture on basic cellular functions beyond cell cycle progression. Disc transplantation experiments confirmed that these detrimental consequences do not reflect fatal damage of imaginal discs during isolation, arguing clearly for a medium insufficiency. Alternative culture media were evaluated, including hemolymph, which surrounds imaginal discs during growth in situ. But isolated larval hemolymph was found to be even less adequate than current culture media, presumably as a result of conversion processes during hemolymph isolation or disc culture. The significance of prominent growth-regulating pathways during disc culture was analyzed, as well as effects of insulin and disc co-culture with larval tissues as potential sources of endocrine factors. Based on our analyses, we developed a culture protocol that

  9. Unsteady Flow Interactions Between Pitching Wings In Schooling Arrangements

    Science.gov (United States)

    Kurt, Melike; Moored, Keith

    2017-11-01

    In nature, many fish aggregate into large groups or schools for protection against predators, for social interactions and to save energy during migrations. Regardless of their prime motivation, fish experience three-dimensional flow interactions amongst themselves that can improve or hamper swimming performance and give rise to fluid-mediated forces between individuals. To date, the unsteady, three-dimensional flow interactions among schooling fish remains relatively unexplored. In order to study these interactions, the caudal fins of two interacting fish are idealized as two finite span pitching wings arranged in mixtures of canonical in-line and side-by-side arrangements. The forces and moments acting on the wings in the streamwise and cross-stream directions are quantified as the arrangement and the phase delay between the wings is altered. Particle image velocimetry is employed to characterize the flow physics during high efficiency locomotion. Finally, the forces and flowfields of two-dimensional pitching wings are compared with three-dimensional wings to distinguish how three-dimensionality alters the flow interactions in schools of fish.

  10. Multidisciplinary Shape Optimization of a Composite Blended Wing Body Aircraft

    Science.gov (United States)

    Boozer, Charles Maxwell

    A multidisciplinary shape optimization tool coupling aerodynamics, structure, and performance was developed for battery powered aircraft. Utilizing high-fidelity computational fluid dynamics analysis tools and a structural wing weight tool, coupled based on the multidisciplinary feasible optimization architecture; aircraft geometry is modified in the optimization of the aircraft's range or endurance. The developed tool is applied to three geometries: a hybrid blended wing body, delta wing UAS, the ONERA M6 wing, and a modified ONERA M6 wing. First, the optimization problem is presented with the objective function, constraints, and design vector. Next, the tool's architecture and the analysis tools that are utilized are described. Finally, various optimizations are described and their results analyzed for all test subjects. Results show that less computationally expensive inviscid optimizations yield positive performance improvements using planform, airfoil, and three-dimensional degrees of freedom. From the results obtained through a series of optimizations, it is concluded that the newly developed tool is both effective at improving performance and serves as a platform ready to receive additional performance modules, further improving its computational design support potential.

  11. Evaluation of Aircraft Wing-Tip Vortex Using PIV

    Science.gov (United States)

    Alsayed, Omer A.; Asrar, Waqar; Omar, Ashraf A.

    2010-06-01

    The formation and development of a wing-tip vortex in a near and extended near filed were studied experimentally. Particle image velocimetry was used in a wind tunnel to measure the tip vortex velocity field and hence investigate the flow structure in a wake of aircraft half-wing model. The purpose of this investigation is to evaluate the main features of the lift generated vortices in order to find ways to alleviate hazardous wake vortex encounters for follower airplanes during start and approach such that the increase in airport capacity can be achieved. First the wake structure at successive downstream planes crosswise to the axis of the wake vortices was investigated by measuring parameters such as core radius, maximum tangential velocities, vorticities and circulation distributions. The effect of different angles of attack setting on vortex parameters was examined at one downstream location. In very early stages the vortex sheet evolution makes the tip vortex to move inward and to the suction side of the wing. While the core radius and circulation distributions hardly vary with the downstream distance, noticeable differences for the same vortex parameters at different angles of attack settings were observed. The center of the wing tip vortices scatter in a circle of radius nearly equal to 1% of the mean wing chord and wandering amplitudes shows no direct dependence on the vortex strength but linearly increase with the downstream distance.

  12. Aerodynamic performance of a hovering hawkmoth with flexible wings: a computational approach

    Science.gov (United States)

    Nakata, Toshiyuki; Liu, Hao

    2012-01-01

    Insect wings are deformable structures that change shape passively and dynamically owing to inertial and aerodynamic forces during flight. It is still unclear how the three-dimensional and passive change of wing kinematics owing to inherent wing flexibility contributes to unsteady aerodynamics and energetics in insect flapping flight. Here, we perform a systematic fluid-structure interaction based analysis on the aerodynamic performance of a hovering hawkmoth, Manduca, with an integrated computational model of a hovering insect with rigid and flexible wings. Aerodynamic performance of flapping wings with passive deformation or prescribed deformation is evaluated in terms of aerodynamic force, power and efficiency. Our results reveal that wing flexibility can increase downwash in wake and hence aerodynamic force: first, a dynamic wing bending is observed, which delays the breakdown of leading edge vortex near the wing tip, responsible for augmenting the aerodynamic force-production; second, a combination of the dynamic change of wing bending and twist favourably modifies the wing kinematics in the distal area, which leads to the aerodynamic force enhancement immediately before stroke reversal. Moreover, an increase in hovering efficiency of the flexible wing is achieved as a result of the wing twist. An extensive study of wing stiffness effect on aerodynamic performance is further conducted through a tuning of Young's modulus and thickness, indicating that insect wing structures may be optimized not only in terms of aerodynamic performance but also dependent on many factors, such as the wing strength, the circulation capability of wing veins and the control of wing movements. PMID:21831896

  13. Do hummingbirds use a different mechanism than insects to flip and twist their wings?

    Science.gov (United States)

    Song, Jialei; Luo, Haoxiang; Hedrick, Tyson

    2014-11-01

    Hovering hummingbirds flap their wings in an almost horizontal stroke plane and flip the wings to invert the angle of attack after stroke reversal, a strategy also utilized by many hovering insects such as fruit flies. However, unlike insects whose wing actuation mechanism is only located at the base, hummingbirds have a vertebrate musculoskeletal system and their wings contain bones and muscles and thus, they may be capable of both actively flipping and twisting their wings. To investigate this issue, we constructed a hummingbird wing model and study its pitching dynamics. The wing kinematics are reconstructed from high-speed imaging data, and the inertial torques are calculated in a rotating frame of reference using mass distribution data measured from dissections of hummingbird wings. Pressure data from a previous CFD study of the same wing kinematics are used to calculate the aerodynamic torque. The results show that like insect wings, the hummingbird wing pitching is driven by its own inertia during reversal, and the aerodynamic torque is responsible for wing twist during mid-stroke. In conclusion, our study suggests that their wing dynamics are very similar even though their actuation systems are entirely different. This research was supported by the NSF.

  14. Short revolving wings enable hovering animals to avoid stall and reduce drag

    Science.gov (United States)

    Lentink, David; Kruyt, Jan W.; Heijst, Gertjan F.; Altshuler, Douglas L.

    2014-11-01

    Long and slender wings reduce the drag of airplanes, helicopters, and gliding animals, which operate at low angle of attack (incidence). Remarkably, there is no evidence for such influence of wing aspect ratio on the energetics of hovering animals that operate their wings at much higher incidence. High incidence causes aircraft wings to stall, hovering animals avoid stall by generating an attached vortex along the leading edge of their wings that elevates lift. Hypotheses that explain this capability include the necessity for a short radial distance between the shoulder joint and wing tip, measured in chord lengths, instead of the long tip-to-tip distance that elevates aircraft performance. This stems from how hovering animals revolve their wings around a joint, a condition for which the precise effect of aspect ratio on stall performance is unknown. Here we show that the attachment of the leading edge vortex is determined by wing aspect ratio with respect to the center of rotation-for a suite of aspect ratios that represent both animal and aircraft wings. The vortex remains attached when the local radius is shorter than 4 chord lengths, and separates outboard on more slender wings. Like most other hovering animals, hummingbirds have wing aspect ratios between 3 and 4, much stubbier than helicopters. Our results show this makes their wings robust against flow separation, which reduces drag below values obtained with more slender wings. This revises our understanding of how aspect ratio improves performance at low Reynolds numbers.

  15. The bHLH factors Dpn and members of the E(spl complex mediate the function of Notch signalling regulating cell proliferation during wing disc development

    Directory of Open Access Journals (Sweden)

    Beatriz P. San Juan

    2012-05-01

    The Notch signalling pathway plays an essential role in the intricate control of cell proliferation and pattern formation in many organs during animal development. In addition, mutations in most members of this pathway are well characterized and frequently lead to tumour formation. The Drosophila imaginal wing discs have provided a suitable model system for the genetic and molecular analysis of the different pathway functions. During disc development, Notch signalling at the presumptive wing margin is necessary for the restricted activation of genes required for pattern formation control and disc proliferation. Interestingly, in different cellular contexts within the wing disc, Notch can either promote cell proliferation or can block the G1-S transition by negatively regulating the expression of dmyc and bantam micro RNA. The target genes of Notch signalling that are required for these functions have not been identified. Here, we show that the Hes vertebrate homolog, deadpan (dpn, and the Enhancer-of-split complex (E(splC genes act redundantly and cooperatively to mediate the Notch signalling function regulating cell proliferation during wing disc development.

  16. Effect of leading- and trailing-edge flaps on clipped delta wings with and without wing camber at supersonic speeds

    Science.gov (United States)

    Hernandez, Gloria; Wood, Richard M.; Covell, Peter F.

    1994-01-01

    An experimental investigation of the aerodynamic characteristics of thin, moderately swept fighter wings has been conducted to evaluate the effect of camber and twist on the effectiveness of leading- and trailing-edge flaps at supersonic speeds in the Langley Unitary Plan Wind Tunnel. The study geometry consisted of a generic fuselage with camber typical of advanced fighter designs without inlets, canopy, or vertical tail. The model was tested with two wing configurations an uncambered (flat) wing and a cambered and twisted wing. Each wing had an identical clipped delta planform with an inboard leading edge swept back 65 deg and an outboard leading edge swept back 50 deg. The trailing edge was swept forward 25 deg. The leading-edge flaps were deflected 4 deg to 15 deg, and the trailing-edge flaps were deflected from -30 deg to 10 deg. Longitudinal force and moment data were obtained at Mach numbers of 1.60, 1.80, 2.00, and 2.16 for an angle-of-attack range 4 deg to 20 deg at a Reynolds number of 2.16 x 10(exp 6) per foot and for an angle-of-attack range 4 deg to 20 deg at a Reynolds number of 2.0 x 10(exp 6) per foot. Vapor screen, tuft, and oil flow visualization data are also included.

  17. Longer wings for faster springs - wing length relates to spring phenology in a long-distance migrant across its range.

    Science.gov (United States)

    Hahn, Steffen; Korner-Nievergelt, Fränzi; Emmenegger, Tamara; Amrhein, Valentin; Csörgő, Tibor; Gursoy, Arzu; Ilieva, Mihaela; Kverek, Pavel; Pérez-Tris, Javier; Pirrello, Simone; Zehtindjiev, Pavel; Salewski, Volker

    2016-01-01

    In migratory birds, morphological adaptations for efficient migratory flight often oppose morphological adaptations for efficient behavior during resident periods. This includes adaptations in wing shape for either flying long distances or foraging in the vegetation and in climate-driven variation of body size. In addition, the timing of migratory flights and particularly the timely arrival at local breeding sites is crucial because fitness prospects depend on site-specific phenology. Thus, adaptations for efficient long-distance flights might be also related to conditions at destination areas. For an obligatory long-distance migrant, the common nightingale, we verified that wing length as the aerodynamically important trait, but not structural body size increased from the western to the eastern parts of the species range. In contrast with expectation from aerodynamic theory, however, wing length did not increase with increasing migration distances. Instead, wing length was associated with the phenology at breeding destinations, namely the speed of local spring green-up. We argue that longer wings are beneficial for adjusting migration speed to local conditions for birds breeding in habitats with fast spring green-up and thus short optimal arrival periods. We suggest that the speed of spring green-up at breeding sites is a fundamental variable determining the timing of migration that fine tune phenotypes in migrants across their range.

  18. Leading-edge vortex shedding from rotating wings

    Energy Technology Data Exchange (ETDEWEB)

    Kolomenskiy, Dmitry [Centre de Recherches Mathématiques (CRM), Department of Mathematics and Statistics, McGill University, 805 Sherbrooke W., Montreal, QC H3A 0B9 (Canada); Elimelech, Yossef [Faculty of Aerospace Engineering, Technion-Israel Institute of Technology, Haifa 32000 (Israel); Schneider, Kai, E-mail: dkolom@gmail.com [M2P2–CNRS, Université d' Aix-Marseille, 39, rue Frédéric Joliot-Curie, F-13453 Marseille Cedex 13 (France)

    2014-06-01

    This paper presents a numerical investigation of the leading-edge vortices generated by rotating triangular wings at Reynolds number Re = 250. A series of three-dimensional numerical simulations have been carried out using a Fourier pseudo-spectral method with volume penalization. The transition from stable attachment of the leading-edge vortex to periodic vortex shedding is explored, as a function of the wing aspect ratio and the angle of attack. It is found that, in a stable configuration, the spanwise flow in the recirculation bubble past the wing is due to the centrifugal force, incompressibility and viscous stresses. For the flow outside of the bubble, an inviscid model of spanwise flow is presented. (papers)

  19. An overview of two nonlinear supersonic wing design studies

    Science.gov (United States)

    Miller, D. S.; Pittman, J. L.; Wood, R. M.

    1983-01-01

    The progress of two studies which apply nonlinear aerodynamics to supersonic wing design is reviewed. The first study employed a nonlinear potential flow code to design wings for high lift and low drag due to lift by employing a controlled leading-edge expansion in which the crossflow accelerates to supercritical conditions and decelerates through a weak shock. The second study utilized a modified linearized theory code to explore the concept of using 'attainable' leading-edge thrust as a guide for selecting a wing leading-edge shape (planform and radius) for maintaining attached flow and maximizing leading-edge thrust. Experimental and theoretical results obtained during the course of these two studies are discussed.

  20. A novel mechanism for emulating insect wing kinematics

    International Nuclear Information System (INIS)

    Seshadri, Pranay; Benedict, Moble; Chopra, Inderjit

    2012-01-01

    A novel dual-differential four-bar flapping mechanism that can accurately emulate insect wing kinematics in all three degrees of freedom (translation, rotation and stroke plane deviation) is developed. The mechanism is specifically designed to be simple and scalable such that it can be utilized on an insect-based flapping wing micro air vehicle. Kinematic formulations for the wing stroke position, pitch angle and coning angle for this model are derived from first principles and compared with a 3D simulation. A benchtop flapping mechanism based on this model was designed and built, which was also equipped with a balance for force measurements. 3D motion capture tests were conducted on this setup to demonstrate the capability of generating complex figure-of-eight flapping motions along with dynamic pitching. The dual-differential four-bar mechanism was implemented on a light-weight vehicle that demonstrated tethered hover. (paper)

  1. AERODYNAMIC LOAD OF AN AIRCRAFT WITH A HIGHLY ELASTIC WING

    Directory of Open Access Journals (Sweden)

    Pavel Schoř

    2017-09-01

    Full Text Available In this article, a method for calculation of air loads of an aircraft with an elastic wing is presented. The method can predict a redistribution of air loads when the elastic wing deforms. Unlike the traditional Euler or Navier-Stokes CFD to FEM coupling, the method uses 3D panel method as a source of aerodynamic data. This makes the calculation feasible on a typical recent workstation. Due to a short computational time and low hardware demands this method is suitable for both the preliminary design stage and the load evaluation stage. A case study is presented. The study compares a glider wing performing a pull maneuver at both rigid and and elastic state. The study indicates a significant redistribution of air load at the elastic case.

  2. Aeroelastic Flutter of Subsonic Aircraft Wing Section with Control Surface

    Directory of Open Access Journals (Sweden)

    Aeroelastic Flutter of Subsonic Aircraft Wing Section with Control Surface

    2015-12-01

    Full Text Available Aeroelastic flutter in aircraft mechanisms is unavoidable, essentially in the wing and control surface. In this work a three degree-of-freedom aeroelastic wing section with trailing edge flap is modeled numerically and theoretically. FLUENT code based on the steady finite volume is used for the prediction of the steady aerodynamic characteristics (lift, drag, pitching moment, velocity, and pressure distribution as well as the Duhamel formulation is used to model the aerodynamic loads theoretically. The system response (pitch, flap pitch and plunge was determined by integration the governing equations using MATLAB with a standard Runge–Kutta algorithm in conjunction with Henon’s method. The results are compared with previous experimental data. The results show that the aerodynamic loads and wing-flap system response are increased when increasing the flow speed. On the other hand the aeroelastic response led up to limit cycle oscillation when the flow equals or more than flutter speed.

  3. Surface Roughness Measurement on a Wing Aircraft by Speckle Correlation

    Directory of Open Access Journals (Sweden)

    Alberto Barrientos

    2013-09-01

    Full Text Available The study of the damage of aeronautical materials is important because it may change the microscopic surface structure profiles. The modification of geometrical surface properties can cause small instabilities and then a displacement of the boundary layer. One of the irregularities we can often find is surface roughness. Due to an increase of roughness and other effects, there may be extra momentum losses in the boundary layer and a modification in the parasite drag. In this paper we present a speckle method for measuring the surface roughness on an actual unmanned aircraft wing. The results show an inhomogeneous roughness distribution on the wing, as expected according to the anisotropic influence of the winds over the entire wing geometry. A calculation of the uncertainty of the technique is given.

  4. Surface roughness measurement on a wing aircraft by speckle correlation.

    Science.gov (United States)

    Salazar, Félix; Barrientos, Alberto

    2013-09-05

    The study of the damage of aeronautical materials is important because it may change the microscopic surface structure profiles. The modification of geometrical surface properties can cause small instabilities and then a displacement of the boundary layer. One of the irregularities we can often find is surface roughness. Due to an increase of roughness and other effects, there may be extra momentum losses in the boundary layer and a modification in the parasite drag. In this paper we present a speckle method for measuring the surface roughness on an actual unmanned aircraft wing. The results show an inhomogeneous roughness distribution on the wing, as expected according to the anisotropic influence of the winds over the entire wing geometry. A calculation of the uncertainty of the technique is given.

  5. Development and experiments of the Sea-Wing underwater glider

    Science.gov (United States)

    Yu, Jian-Cheng; Zhang, Ai-Qun; Jin, Wen-Ming; Chen, Qi; Tian, Yu; Liu, Chong-Jie

    2011-12-01

    Underwater gliders, which glide through water columns by use of a pair of wings, are efficient long-distance, long-duration marine environment observatory platforms. The Sea-Wing underwater glider, developed by the Shenyang Institute of Automation, CAS, is designed for the application of deep-sea environment variables observation. The system components, the mechanical design, and the control system design of the Sea-Wing underwater glider are described in this paper. The pitch and roll adjusting models are derived based on the mechanical design, and the adjusting capabilities for the pitch and roll are analyzed according to the models. Field experiments have been carried out for validating the gliding motion and the ability of measuring ocean environment variables. Experimental results of the motion performances of the glider are presented.

  6. HYDRODYNAMICS OF OSCILLATING WING ON THE PITCH ANGLE

    Directory of Open Access Journals (Sweden)

    Vitalii Korobov

    2017-07-01

    Full Text Available Purpose: research of the hydrodynamic characteristics of a wing in a nonstationary stream. Methods: The experimental studies of the hydrodynamic load acting on the wing of 1.5 elongation, wich harmonically oscillated respect to the transversal axis in the frequency range of 0.2-2.5 Hz. The flow speed in the hydrodynamic tunnel ranged of 0.2-1.5 m/s. Results: The instantaneous values of the coefficients of lift and drag / thrust on the pitch angle at unsteady flow depends on the Strouhal number.Discussion: with increasing oscillation frequency coefficients of hydrodynamic force components significantly higher than the data for the stationary blowing out of the wing.

  7. Spanwise drag variation on low Re wings -- revisited

    Science.gov (United States)

    Yang, Shanling; Spedding, Geoffrey

    2011-11-01

    Aerodynamic performance measurement and prediction of airfoils and wings at chord Reynolds numbers below 105 is both difficult and increasingly important in application to small-scale aircraft. Not only are the aerodynamics strongly affected by the dynamics of the unstable laminar boundary layer but the flow is decreasingly likely to be two-dimensional as Re decreases. The spanwise variation of the flow along a two-dimensional geometry is often held to be responsible for the large variations in measured profile drag coefficient. Here we measure local two-dimensional drag coefficients along a finite wing using non-intrusive PIV methods. Variations in Cd (y) can be related to local flow variations on the wing itself. Integrated values can be compared with force balance data, and the proper description of drag components at low Re will be discussed.

  8. Flow Measurements of a Plunging Wing in Unsteady Environment

    Science.gov (United States)

    Wengel, Jesse; Nathan, Rungun; Cheng, Bo; Eslam-Panah, Azar

    2017-11-01

    Despite the great progress in their design and control, Unmanned Aerial Vehicles (UAVs) are tremendously troubled while flying in turbulent environments, which are common in the lower atmospheric boundary layer (ABL). A nominally 2D plunging wing was developed and tested in the presence of unsteady wake to investigate the effect of the flow disturbances on vorticity fields. The experiments were conducted in a water channel facility with test section width of 0.76 m, and a water depth of 0.6 m. The unsteady wake in the form of von Kármán Vortex Street was generated by a cylinder located upstream of the plunging wing. The plunge amplitude and frequency of the oscillation were adjusted to bracket the range of Strouhal numbers relevant to the biological locomotion (0.25PIV) was employed to quantitatively study the effect of unsteady wake on the flow measurements of the plunging wing.

  9. Wing bone geometry reveals active flight in Archaeopteryx.

    Science.gov (United States)

    Voeten, Dennis F A E; Cubo, Jorge; de Margerie, Emmanuel; Röper, Martin; Beyrand, Vincent; Bureš, Stanislav; Tafforeau, Paul; Sanchez, Sophie

    2018-03-13

    Archaeopteryx is an iconic fossil taxon with feathered wings from the Late Jurassic of Germany that occupies a crucial position for understanding the early evolution of avian flight. After over 150 years of study, its mosaic anatomy unifying characters of both non-flying dinosaurs and flying birds has remained challenging to interpret in a locomotory context. Here, we compare new data from three Archaeopteryx specimens obtained through phase-contrast synchrotron microtomography to a representative sample of archosaurs employing a diverse array of locomotory strategies. Our analyses reveal that the architecture of Archaeopteryx's wing bones consistently exhibits a combination of cross-sectional geometric properties uniquely shared with volant birds, particularly those occasionally utilising short-distance flapping. We therefore interpret that Archaeopteryx actively employed wing flapping to take to the air through a more anterodorsally posteroventrally oriented flight stroke than used by modern birds. This unexpected outcome implies that avian powered flight must have originated before the latest Jurassic.

  10. Design, realization and structural testing of a compliant adaptable wing

    International Nuclear Information System (INIS)

    Molinari, G; Arrieta, A F; Ermanni, P; Quack, M; Morari, M

    2015-01-01

    This paper presents the design, optimization, realization and testing of a novel wing morphing concept, based on distributed compliance structures, and actuated by piezoelectric elements. The adaptive wing features ribs with a selectively compliant inner structure, numerically optimized to achieve aerodynamically efficient shape changes while simultaneously withstanding aeroelastic loads. The static and dynamic aeroelastic behavior of the wing, and the effect of activating the actuators, is assessed by means of coupled 3D aerodynamic and structural simulations. To demonstrate the capabilities of the proposed morphing concept and optimization procedure, the wings of a model airplane are designed and manufactured according to the presented approach. The goal is to replace conventional ailerons, thus to achieve controllability in roll purely by morphing. The mechanical properties of the manufactured components are characterized experimentally, and used to create a refined and correlated finite element model. The overall stiffness, strength, and actuation capabilities are experimentally tested and successfully compared with the numerical prediction. To counteract the nonlinear hysteretic behavior of the piezoelectric actuators, a closed-loop controller is implemented, and its capability of accurately achieving the desired shape adaptation is evaluated experimentally. Using the correlated finite element model, the aeroelastic behavior of the manufactured wing is simulated, showing that the morphing concept can provide sufficient roll authority to allow controllability of the flight. The additional degrees of freedom offered by morphing can be also used to vary the plane lift coefficient, similarly to conventional flaps. The efficiency improvements offered by this technique are evaluated numerically, and compared to the performance of a rigid wing. (paper)

  11. Design of a hybrid rocket / inflatable wing UAV

    Science.gov (United States)

    Sudduth, Cory

    This paper discusses the design challenges and development of a UAV that transitions from a rocket, which allows the aircraft to reach a target altitude rapidly, and then deploys an inflatable wing from an enclosed shell in midflight to allow for loitering and surveillance. The wing deployment and transition is tested in static and dynamic environments, while the performance and stability of both the aircraft mode and rocket mode are examined analytically. An in-depth discussion of key components, including the design, analysis and testing, is also included. Designing an UAV that transitions from a high velocity rocket, to a slow velocity UAV provides many difficult and unique design challenges. For example, the incorporation of deployable wing technology into a full UAV system results in many design constraints. In this particular design inflatable wings are used to generate lift during aircraft mode, and the stabilizing fins for the main wing also acted as the fins for the vehicle during its rocket phase. This required the balancing of the two different vehicle configurations to ensure that the aircraft would be able to fly stably in both modes, and transition between them without catastrophic failure. Significant research, and testing went into the finding the best method of storing the inflatable wing, as well as finding the required inflation rate to minimize unsteady aerodynamic affects. Design work was also invested in the development of an inflation system, as it had to be highly reliable, and yet very light weight for use in this small UAV. This paper discusses how these design challenges were overcome, the development and testing of individual sub-components and how they are incorporated into the overall vehicle. The analysis that went into this UAV, as well as methods used to optimize the design in order to minimize weight and maximize the aircraft performance and loitering time is also discussed.

  12. Cabin-fuselage-wing structural design concept with engine installation

    Science.gov (United States)

    Ariotti, Scott; Garner, M.; Cepeda, A.; Vieira, J.; Bolton, D.

    1993-01-01

    The purpose of this project is to provide a fuselage structural assembly and wing structural design that will be able to withstand the given operational parameters and loads provided by Federal Aviation Regulation Part 23 (FAR 23) and the Statement of Work (SOW). The goal is to provide a durable lightweight structure that will transfer the applied loads through the most efficient load path. Areas of producibility and maintainability of the structure will also be addressed. All of the structural members will also meet or exceed the desired loading criteria, along with providing adequate stiffness, reliability, and fatigue life as stated in the SOW. Considerations need to be made for control system routing and cabin heating/ventilation. The goal of the wing structure and carry through structure is also to provide a simple, lightweight structure that will transfer the aerodynamic forces produced by the wing, tailboom, and landing gear. These forces will be channeled through various internal structures sized for the pre-determined loading criteria. Other considerations were to include space for flaps, ailerons, fuel tanks, and electrical and control system routing. The difficulties encountered in the fuselage design include expanding the fuselage cabin to accept a third occupant in a staggered configuration and providing ample volume for their safety. By adding a third person the CG of aircraft will move forward so the engine needs to be moved aft to compensate for the difference in the moment. This required the provisions of a ring frame structure for the new position of the engine mount. The difficulties encountered in the wing structural design include resizing the wing for the increased capacity and weight, and compensating for a large torsion produced by the tail boom by placing a great number of stiffeners inside the boom, which will result in the relocation of the fuel tank. Finally, an adequate carry through structure for the wing and fuselage interface will be

  13. Experimental Investigation of Pitch Control Enhancement to the Flapping Wing Micro Air Vehicle

    National Research Council Canada - National Science Library

    Kian, Chin C

    2006-01-01

    .... The MAV without the main fixed-wing is placed in a laminar flow field within a low speed wind tunnel with the wake after the flapping wings characterized with a constant temperature anemometer...

  14. Reflection characterization of nano-sized dielectric structure in Morpho butterfly wings

    Science.gov (United States)

    Zhu, Dong

    2017-10-01

    Morpho butterflies living in Central and South America are well-known for their structural-colored blue wings. The blue coloring originates from the interaction of light with nano-sized dielectric structures that are equipped on the external surface of scales covering over their wings. The high-accuracy nonstandard finite-difference time domain (NS-FDTD) method is used to investigate the reflection characterization from the nanostructures. In the NS-FDTD calculation, a computational model is built to mimic the actual tree-like multilayered structures wherever possible using the hyperbolic tangent functions. It is generally known that both multilayer interference and diffraction grating phenomena can occur when light enters the nano-sized multilayered structure. To answer the question that which phenomenon is mainly responsible for the blue coloring, the NS-FDTD calculation is performed under various incidence angles at wavelengths from 360 to 500 nm. The calculated results at one incident wavelength under different incidence angles are visualized in a two-dimensional mapping image, where horizontal and vertical axes are incidence and reflection angles, respectively. The images demonstrate a remarkable transition from a ring-like pattern at shorter wavelengths to a retro-reflection pattern at longer wavelengths. To clarify the origin of the pattern transition, the model is separated into several simpler parts and compared their mapping images with the theoretical diffraction calculations. It can be concluded that the blue coloring at longer wavelengths is mainly caused by the cooperation of multilayer interference and retro-reflection while the effect of diffraction grating is predominant at shorter wavelengths.

  15. Crossflow-Vortex Breakdown on Swept Wings: Correlation of Nonlinear Physics

    Science.gov (United States)

    Joslin, R. D.; Streett, C. L.

    1994-01-01

    The spatial evolution of cross flow-vortex packets in a laminar boundary layer on a swept wing are computed by the direct numerical simulation of the incompressible Navier- Stokes equations. A wall-normal velocity distribution of steady suction and blowing at the wing surface is used to generate a strip of equally spaced and periodic disturbances along the span. Three simulations are conducted to study the effect of initial amplitude on the disturbance evolution, to determine the role of traveling cross ow modes in transition, and to devise a correlation function to guide theories of transition prediction. In each simulation, the vortex packets first enter a chordwise region of linear independent growth, then, the individual packets coalesce downstream and interact with adjacent packets, and, finally, the vortex packets nonlinearly interact to generate inflectional velocity profiles. As the initial amplitude of the disturbance is increased, the length of the evolution to breakdown decreases. For this pressure gradient, stationary modes dominate the disturbance evolution. A two-coeffcient function was devised to correlate the simulation results. The coefficients, combined with a single simulation result, provide sufficient information to generate the evolution pattern for disturbances of any initial amplitude.

  16. Numerical investigation of unsteady vortex breakdown past 80°/65° double-delta wing

    Directory of Open Access Journals (Sweden)

    Liu Jian

    2014-06-01

    Full Text Available An improved delayed detached eddy simulation (IDDES method based on the k-ω-SST (shear stress transport turbulence model was applied to predict the unsteady vortex breakdown past an 80°/65° double-delta wing (DDW, where the angles of attack (AOAs range from 30° to 40°. Firstly, the IDDES model and the relative numerical methods were validated by simulating the massively separated flow around an NACA0021 straight wing at the AOA of 60°. The fluctuation properties of the lift and pressure coefficients were analyzed and compared with the available measurements. For the DDW case, the computations were compared with such measurements as the mean lift, drag, pitching moment, pressure coefficients and breakdown locations. Furthermore, the unsteady properties were investigated in detail, such as the frequencies of force and moments, pressure fluctuation on the upper surface, typical vortex breakdown patterns at three moments, and the distributions of kinetic turbulence energy at a stream wise section. Two dominated modes are observed, in which their Strouhal numbers are 1.0 at the AOAs of 30°, 32° and 34° and 0.7 at the AOAs of 36°, 38° and 40°. The breakdown vortex always moves upstream and downstream and its types change alternatively. Furthermore, the vortex can be identified as breakdown or not through the mean pressure, root mean square of pressure, or even through correlation analysis.

  17. Molecular organization of the nanoscale surface structures of the dragonfly Hemianax papuensis wing epicuticle.

    Directory of Open Access Journals (Sweden)

    Elena P Ivanova

    Full Text Available The molecular organization of the epicuticle (the outermost layer of insect wings is vital in the formation of the nanoscale surface patterns that are responsible for bestowing remarkable functional properties. Using a combination of spectroscopic and chromatographic techniques, including Synchrotron-sourced Fourier-transform infrared microspectroscopy (FTIR, x-ray photoelectron spectroscopy (XPS depth profiling and gas chromatography-mass spectrometry (GCMS, we have identified the chemical components that constitute the nanoscale structures on the surface of the wings of the dragonfly, Hemianax papuensis. The major components were identified to be fatty acids, predominantly hexadecanoic acid and octadecanoic acid, and n-alkanes with even numbered carbon chains ranging from C14 to C30. The data obtained from XPS depth profiling, in conjunction with that obtained from GCMS analyses, enabled the location of particular classes of compounds to different regions within the epicuticle. Hexadecanoic acid was found to be a major component of the outer region of the epicuticle, which forms the surface nanostructures, and was also detected in deeper layers along with octadecanoic acid. Aliphatic compounds were detected throughout the epicuticle, and these appeared to form a third discrete layer that was separate from both the inner and outer epicuticles, which has never previously been reported.

  18. The Drosophila wings apart gene anchors a novel, evolutionarily conserved pathway of neuromuscular development.

    Science.gov (United States)

    Morriss, Ginny R; Jaramillo, Carmelita T; Mikolajczak, Crystal M; Duong, Sandy; Jaramillo, Maryann S; Cripps, Richard M

    2013-11-01

    wings apart (wap) is a recessive, semilethal gene located on the X chromosome in Drosophila melanogaster, which is required for normal wing-vein patterning. We show that the wap mutation also results in loss of the adult jump muscle. We use complementation mapping and gene-specific RNA interference to localize the wap locus to the proximal X chromosome. We identify the annotated gene CG14614 as the gene affected by the wap mutation, since one wap allele contains a non-sense mutation in CG14614, and a genomic fragment containing only CG14614 rescues the jump-muscle phenotypes of two wap mutant alleles. The wap gene lies centromere-proximal to touch-insensitive larva B and centromere-distal to CG14619, which is tentatively assigned as the gene affected in introverted mutants. In mutant wap animals, founder cell precursors for the jump muscle are specified early in development, but are later lost. Through tissue-specific knockdowns, we demonstrate that wap function is required in both the musculature and the nervous system for normal jump-muscle formation. wap/CG14614 is homologous to vertebrate wdr68, DDB1 and CUL4 associated factor 7, which also are expressed in neuromuscular tissues. Thus, our findings provide insight into mechanisms of neuromuscular development in higher animals and facilitate the understanding of neuromuscular diseases that may result from mis-expression of muscle-specific or neuron-specific genes.

  19. The effective compliance of spatially evolving planar wing-cracks

    Science.gov (United States)

    Ayyagari, R. S.; Daphalapurkar, N. P.; Ramesh, K. T.

    2018-02-01

    We present an analytic closed form solution for anisotropic change in compliance due to the spatial evolution of planar wing-cracks in a material subjected to largely compressive loading. A fully three-dimensional anisotropic compliance tensor is defined and evaluated considering the wing-crack mechanism, using a mixed-approach based on kinematic and energetic arguments to derive the coefficients in incremental compliance. Material, kinematic and kinetic parametric influences on the increments in compliance are studied in order to understand their physical implications on material failure. Model verification is carried out through comparisons to experimental uniaxial compression results to showcase the predictive capabilities of the current study.

  20. Artificial Bird Feathers: An Adaptive Wing with High Lift Capability.

    Science.gov (United States)

    Hage, W.; Meyer, R.; Bechert, D. W.

    1997-11-01

    In Wind tunnel experiments, the operation of the covering feathers of bird wings has been investigated. At incipient flow separation, local flow reversal lifts the feathers and inhibits the spreading of the separation regime towards the leading edge. This mechanism can be utilized by movable flaps on airfoils. The operation of quasi-steady and of vibrating movable flaps is outlined. These devices are self-actuated, require no energy and do not produce parasitic drag. They are compatible with laminar and turbulent airfoils as well as with various conventional flaps on aircraft wings. Laboratory and flight experiments are shown. Ref: AIAA-Paper 97-1960.

  1. Aerodynamic Optimization of an Over-the-Wing-Nacelle-Mount Configuration

    OpenAIRE

    Sasaki, Daisuke; Nakahashi, Kazuhiro

    2011-01-01

    An over-the-wing-nacelle-mount airplane configuration is known to prevent the noise propagation from jet engines toward ground. However, the configuration is assumed to have low aerodynamic efficiency due to the aerodynamic interference effect between a wing and a nacelle. In this paper, aerodynamic design optimization is conducted to improve aerodynamic efficiency to be equivalent to conventional under-the-wing-nacelle-mount configuration. The nacelle and wing geometry are modified to achiev...

  2. A new genus of long-legged flies displaying remarkable wing directional asymmetry.

    OpenAIRE

    Runyon, Justin B; Hurley, Richard L

    2004-01-01

    A previously unknown group of flies is described whose males exhibit directional asymmetry, in that the left wing is larger than, and of a different shape from, the right wing. To our knowledge, wing asymmetry of this degree has not previously been reported in an animal capable of flight. Such consistent asymmetry must result from a left-right axis during development, a level of differentiation whose existence has been questioned for insects. Wing asymmetry of this magnitude has implications ...

  3. Nutritional physiology of life-history trade-offs: how food protein-carbohydrate content influences life-history traits in the wing-polymorphic cricket Gryllus firmus.

    Science.gov (United States)

    Clark, Rebecca M; Zera, Anthony J; Behmer, Spencer T

    2015-01-15

    Although life-history trade-offs result from the differential acquisition and allocation of nutritional resources to competing physiological functions, many aspects of this topic remain poorly understood. Wing-polymorphic insects, which possess alternative morphs that trade off allocation to flight capability versus early reproduction, provide a good model system for exploring this topic. In this study, we used the wing-polymorphic cricket Gryllus firmus to test how expression of the flight capability versus reproduction trade-off was modified across a heterogeneous protein-carbohydrate nutritional landscape. Newly molted adult female long- and short-winged crickets were given one of 13 diets with different concentrations and ratios of protein and digestible carbohydrate; for each cricket, we measured consumption patterns, growth and allocation to reproduction (ovary mass) versus flight muscle maintenance (flight muscle mass and somatic lipid stores). Feeding responses in both morphs were influenced more by total macronutrient concentration than by protein-carbohydrate ratio, except at high-macronutrient concentration, where protein-carbohydrate balance was important. Mass gain tended to be greatest on protein-biased diets for both morphs, but was consistently lower across all diets for long-winged females. When long-winged females were fed high-carbohydrate foods, they accumulated greater somatic lipid stores; on high-protein foods, they accumulated greater somatic protein stores. Food protein-carbohydrate content also affected short-winged females (selected for early reproductive onset), which showed dramatic increases in ovary size, including ovarian stores of lipid and protein, on protein-biased foods. This is the first study to show how the concentration and ratio of dietary protein and carbohydrate affects consumption and allocation to key physiological features associated with the reproduction-dispersal life-history trade-off. © 2015. Published by The

  4. Exploring and exploiting natural variation in the wings of a predatory ladybird beetle for biological control

    NARCIS (Netherlands)

    Lommen, S.T.E.

    2013-01-01

    The central theme of this PhD thesis is natural variation in the wing length of the predatory two-spot ladybird beetle, Adalia bipunctata. ‘Wingless’ individuals of this species occur occasionally. They possess truncated wing covers and flight wings and cannot fly, but the extent of the reduction is

  5. A mechanical model of wing and theoretical estimate of taper factor ...

    Indian Academy of Sciences (India)

    PRAKASH

    β angle between humerus and body βmin minimum angle between humerus and body βe empirical span ratio (bobs/ bmax) βmin,sr minimum span ratio b wing span. bB body width bmax maximum wing span bobs observed wingspan c wing chord. Cl,max maximum lift coefficient e taper factor g acceleration due to gravity.

  6. Development and Testing of an Unconventional Morphing Wing Concept with Variable Chord and Camber

    NARCIS (Netherlands)

    Keidel, D.H.K.; Sodja, J.; Werter, N.P.M.; De Breuker, R.; Ermanni, P.; Monajjemi, M.; Liang, W.

    2015-01-01

    Driven by the need to improve the performance and energy-efficiency of aircraft, current research in the field of morphing wings is growing in significance. The most recently developed concepts typically adjust only one characteristic of the wing. Within this paper a new concept for morphing wings

  7. Power reduction and the radial limit of stall delay in revolving wings of different aspect ratio

    NARCIS (Netherlands)

    Kruyt, J.W.; Heijst, Van G.F.; Altshuler, D.L.; Lentink, David

    2015-01-01

    Airplanes and helicopters use high aspect ratio wings to reduce the power required to fly, but must operate at low angle of attack to prevent flow separation and stall. Animals capable of slow sustained flight, such as hummingbirds, have low aspect ratio wings and flap their wings at high angle

  8. Flapping wing actuation using resonant compliant mechanisms : An insect-inspired design

    NARCIS (Netherlands)

    Bolsman, C.T.

    2010-01-01

    The realization of a wing actuation mechanism for a flapping wing micro air vehicle requires a move away from traditional designs based on gears and links. An approach inspired by nature’s flyers is better suited. For flapping flight two wing motions are important: the sweeping and the pitching

  9. A predictive quasi-steady model of aerodynamic loads on flapping wings

    NARCIS (Netherlands)

    Wang, Q.; Goosen, J.F.L.; van Keulen, A.

    2016-01-01

    Quasi-steady aerodynamic models play an important role in evaluating aerodynamic performance and conducting design and optimization of flapping wings. The kinematics of flapping wings is generally a resultant motion of wing translation (yaw) and rotation (pitch and roll). Most quasi-steady models

  10. Note on the glide of a bird with wings bent downwards

    NARCIS (Netherlands)

    Sparenberg, J. A.

    This note considers the influence of the bending down of the wings of a bird on the performance of its glide. The induced drag of bent wings is compared with the induced drag of a corresponding straight wing. Numerical results are given.

  11. 78 FR 3356 - Airworthiness Directives; Various Aircraft Equipped With Wing Lift Struts

    Science.gov (United States)

    2013-01-16

    ...) of this AD, inspect the wing lift strut forks for cracks using magnetic particle procedures, such as... Airworthiness Directives; Various Aircraft Equipped With Wing Lift Struts AGENCY: Federal Aviation... existing airworthiness directive (AD) that applies to certain aircraft equipped with wing lift struts. The...

  12. Image Processing Method for Automatic Discrimination of Hoverfly Species

    Directory of Open Access Journals (Sweden)

    Vladimir Crnojević

    2014-01-01

    Full Text Available An approach to automatic hoverfly species discrimination based on detection and extraction of vein junctions in wing venation patterns of insects is presented in the paper. The dataset used in our experiments consists of high resolution microscopic wing images of several hoverfly species collected over a relatively long period of time at different geographic locations. Junctions are detected using the combination of the well known HOG (histograms of oriented gradients and the robust version of recently proposed CLBP (complete local binary pattern. These features are used to train an SVM classifier to detect junctions in wing images. Once the junctions are identified they are used to extract statistics characterizing the constellations of these points. Such simple features can be used to automatically discriminate four selected hoverfly species with polynomial kernel SVM and achieve high classification accuracy.

  13. PHOTOGRAMMETRIC EVALUATION OF MULTI-TEMPORAL FIXED WING UAV IMAGERY

    Directory of Open Access Journals (Sweden)

    E. Gülch

    2012-09-01

    Full Text Available Several flights have been undertaken with PAMS (Photogrammetric Aerial Mapping System by Germatics, Germany, which is briefly introduced. This system is based on the SmartPlane fixed-wing UAV and a CANON IXUS camera system. The plane is equipped with GPS and has an infrared sensor system to estimate attitude values. A software has been developed to link the PAMS output to a standard photogrammetric processing chain built on Trimble INPHO. The linking of the image files and image IDs and the handling of different cases with partly corrupted output have to be solved to generate an INPHO project file. Based on this project file the software packages MATCH-AT, MATCH-T DSM, OrthoMaster and OrthoVista for digital aerial triangulation, DTM/DSM generation and finally digital orthomosaik generation are applied. The focus has been on investigations on how to adapt the "usual" parameters for the digital aerial triangulation and other software to the UAV flight conditions, which are showing high overlaps, large kappa angles and a certain image blur in case of turbulences. It was found, that the selected parameter setup shows a quite stable behaviour and can be applied to other flights. Investigations have been performed to improve the image quality estimates by the PAMS software and extend it to whole images. This gives the user a reliable basis when deciding on rejecting images with low quality for the follow-up process. Flights over the same area at different times have been compared to each other. The major objective was first to see, on how far differences occur relative to each other, without having access to ground control data, which would have a potential for applications with low requirements on the absolute accuracy. In a second stage the results are compared to GPS measurements on the ground. The results show, that there are influences of weather and illumination visible. The "unusual" flight pattern, which shows big time differences for

  14. Study on flow over finite wing with respect to F-22 raptor, Supermarine Spitfire, F-7 BG aircraft wing and analyze its stability performance and experimental values

    Science.gov (United States)

    Ali, Md. Nesar; Alam, Mahbubul

    2017-06-01

    A finite wing is a three-dimensional body, and consequently the flow over the finite wing is three-dimensional; that is, there is a component of flow in the span wise direction. The physical mechanism for generating lift on the wing is the existence of a high pressure on the bottom surface and a low pressure on the top surface. The net imbalance of the pressure distribution creates the lift. As a by-product of this pressure imbalance, the flow near the wing tips tends to curl around the tips, being forced from the high-pressure region just underneath the tips to the low-pressure region on top. This flow around the wing tips is shown in the front view of the wing. As a result, on the top surface of the wing, there is generally a span wise component of flow from the tip toward the wing root, causing the streamlines over the top surface to bend toward the root. On the bottom surface of the wing, there is generally a span wise component of flow from the root toward the tip, causing the streamlines over the bottom surface to bend toward the tip. Clearly, the flow over the finite wing is three-dimensional, and therefore we would expect the overall aerodynamic properties of such a wing to differ from those of its airfoil sections. The tendency for the flow to "leak" around the wing tips has another important effect on the aerodynamics of the wing. This flow establishes a circulatory motion that trails downstream of the wing; that is, a trailing vortex is created at each wing tip. The aerodynamics of finite wings is analyzed using the classical lifting line model. This simple model allows a closed-form solution that captures most of the physical effects applicable to finite wings. The model is based on the horseshoe-shaped vortex that introduces the concept of a vortex wake and wing tip vortices. The downwash induced by the wake creates an induced drag that did not exist in the two-dimensional analysis. Furthermore, as wingspan is reduced, the wing lift slope decreases

  15. Aerodynamic performance and particle image velocimetery of piezo actuated biomimetic manduca sexta engineered wings towards the design and application of a flapping wing flight vehicle

    Science.gov (United States)

    DeLuca, Anthony M.

    Considerable research and investigation has been conducted on the aerodynamic performance, and the predominate flow physics of the Manduca Sexta size of biomimetically designed and fabricated wings as part of the AFIT FWMAV design project. Despite a burgeoning interest and research into the diverse field of flapping wing flight and biomimicry, the aerodynamics of flapping wing flight remains a nebulous field of science with considerable variance into the theoretical abstractions surrounding aerodynamic mechanisms responsible for aerial performance. Traditional FWMAV flight models assume a form of a quasi-steady approximation of wing aerodynamics based on an infinite wing blade element model (BEM). An accurate estimation of the lift, drag, and side force coefficients is a critical component of autonomous stability and control models. This research focused on two separate experimental avenues into the aerodynamics of AFIT's engineered hawkmoth wings|forces and flow visualization. 1. Six degree of freedom force balance testing, and high speed video analysis was conducted on 30°, 45°, and 60° angle stop wings. A novel, non-intrusive optical tracking algorithm was developed utilizing a combination of a Gaussian Mixture Model (GMM) and ComputerVision (OpenCV) tools to track the wing in motion from multiple cameras. A complete mapping of the wing's kinematic angles as a function of driving amplitude was performed. The stroke angle, elevation angle, and angle of attack were tabulated for all three wings at driving amplitudes ranging from A=0.3 to A=0.6. The wing kinematics together with the force balance data was used to develop several aerodynamic force coefficient models. A combined translational and rotational aerodynamic model predicted lift forces within 10%, and vertical forces within 6%. The total power consumption was calculated for each of the three wings, and a Figure of Merit was calculated for each wing as a general expression of the overall efficiency of

  16. Effect of wing planform and canard location and geometry on the longitudinal aerodynamic characteristics of a close-coupled canard wing model at subsonic speeds

    Science.gov (United States)

    Gloss, B. B.

    1975-01-01

    A generalized wind-tunnel model with canard and wing planforms typical of highly maneuverable aircraft was tested in the Langley 7- by 10-foot high-speed tunnel at a Mach number of 0.30 to determine the effect of canard location, canard size, wing sweep, and canard strake on canard-wing interference to high angles of attack. The major results of this investigation may be summarized as follows: the high-canard configuration (excluding the canard strake and canard flap), for both the 60 deg and 44 deg swept leading-edge wings, produced the highest maximum lift coefficient and the most linear pitching-moment curves; substantially larger gains in the canard lift and total lift were obtained by adding a strake to the canard located below the wing chord plane rather than by adding a strake to the canard located above the wing chord plane.

  17. Multidimensional analysis of Drosophila wing variation in Evolution ...

    Indian Academy of Sciences (India)

    In this study, using Drosophila melanogaster isofemale lines derived from wild flies collected on both slopes of the canyon, we investigated the effect of developmental temperature upon the different components of phenotypic variation of a complex trait: the wing. Combining geometric and traditional morphometrics, we find ...

  18. Investigating Biological Controls to Suppress Spotted Wing Drosophila Populations

    Science.gov (United States)

    The spotted wing drosophila has become a major cherry pest in California. To develop sustainable management options for this highly mobile pest, we worked with cooperators at Oregon State University and the USDA to discover and import natural enemies of the fly from its native range in South Korea ...

  19. Monitoring of a Full-Scale Wing Fatigue Test

    NARCIS (Netherlands)

    Heida, Jaap; Hwang, Joong Sun

    2014-01-01

    A wing of a decommissioned aircraft of the Royal Netherlands Air Force (RNLAF) was fatigue tested to more than two times the design life. Part of the test was the evaluation of load monitoring and Structural Health Monitoring (SHM) techniques. For load monitoring the data of conventional resistance

  20. Electric Propulsion Concepts for an Inverted Joined Wing Airplane Demonstrator

    Directory of Open Access Journals (Sweden)

    Cezary Galinski

    2017-05-01

    Full Text Available One of the airplane design concepts that potentially allows for significantly increased efficiency, but has not yet been investigated thoroughly, is the inverted joined wing configuration, where the upper wing is positioned in front of the lower one. We performed wind tunnel and flight testing of a demonstrator of this concept, first by applying electrical propulsion to simplify wind tunnel testing, and then the same electrical-propulsion demonstrator performed several flights. As the chosen propulsion method proved to be too cumbersome for an intensive flight campaign and significant loss of battery performance was also observed, the electrical propulsion was then replaced by internal combustion propulsion in the second phase, involving longer-duration flight testing. Next we identified and analyzed two potentially beneficial modifications to the design tested: one involved shifting the center of gravity towards the aft, the other involved modifying the thrust vector position, both with the assumption that electric motors can be applied for propulsion. On this basis, the paper finishes with some conclusions concerning a new concept of electrical propulsion for an inverted joined wing design, combining two ideas: hybridization and distribution along the aft wing leading edge.

  1. Modeling Aircraft Wing Loads from Flight Data Using Neural Networks

    Science.gov (United States)

    Allen, Michael J.; Dibley, Ryan P.

    2003-01-01

    Neural networks were used to model wing bending-moment loads, torsion loads, and control surface hinge-moments of the Active Aeroelastic Wing (AAW) aircraft. Accurate loads models are required for the development of control laws designed to increase roll performance through wing twist while not exceeding load limits. Inputs to the model include aircraft rates, accelerations, and control surface positions. Neural networks were chosen to model aircraft loads because they can account for uncharacterized nonlinear effects while retaining the capability to generalize. The accuracy of the neural network models was improved by first developing linear loads models to use as starting points for network training. Neural networks were then trained with flight data for rolls, loaded reversals, wind-up-turns, and individual control surface doublets for load excitation. Generalization was improved by using gain weighting and early stopping. Results are presented for neural network loads models of four wing loads and four control surface hinge moments at Mach 0.90 and an altitude of 15,000 ft. An average model prediction error reduction of 18.6 percent was calculated for the neural network models when compared to the linear models. This paper documents the input data conditioning, input parameter selection, structure, training, and validation of the neural network models.

  2. Wings of the Night: The Natural History of Bats

    Indian Academy of Sciences (India)

    Home; Journals; Resonance – Journal of Science Education; Volume 12; Issue 11. Wings of the Night: The Natural History of Bats. Uttam Saikia. General Article Volume 12 Issue 11 November 2007 pp 63-76. Fulltext. Click here to view fulltext PDF. Permanent link: http://www.ias.ac.in/article/fulltext/reso/012/11/0063-0076 ...

  3. Robust Backstepping Control of Wing Rock Using Disturbance Observer

    Directory of Open Access Journals (Sweden)

    Dawei Wu

    2017-02-01

    Full Text Available Wing rock is a highly nonlinear phenomenon when the aircraft suffers undesired roll-dominated oscillatory at high angle of attack (AOA. Considering the strong nonlinear and unsteady aerodynamic characteristics, an uncertain multi-input and multi-output (MIMO nonlinear wing rock model is studied, and system uncertainties, unsteady aerodynamic disturbances and externaldisturbancesareconsideredinthedesignofwingrockcontrollaw. Tohandletheproblemof multipledisturbances,arobustcontrolschemeisproposedbasedontheextendedstateobserver(ESO and the radial basis function neural network (RBFNN technique. Considering that the effectiveness of actuators are greatly decreased at high AOA, the input saturation problem is also handled by constructing a corresponding auxiliary system. Based on the improved ESO and the auxiliary system, a robust backstepping control law is proposed for the wing rock control. In addition, the dynamic surface control (DSC technique is introduced to avoid the tedious computations of time derivatives for the virtual control laws in the backstepping method. The stability of the closed-loop system is guaranteed via rigorously Lyapunov analysis. Finally, simulation results are presented to illustrate the effectiveness of the ESO and the proposed wing rock control approach.

  4. Male secondary sexual characters in Aphnaeinae wings (Lepidoptera: Lycaenidae)

    OpenAIRE

    Bálint, Zsolt; Heath, Alan; Katona, Gergely; Kertész, Krisztián; Sáfián, Szabolcs

    2017-01-01

    Male secondary sexual characters have been discovered on the hindwing verso of genera Aphnaeus Hübner, [1819], Cigaritis Donzel, 1847, Lipaphnaeus Aurivillius, 1916 and Pseudaletis Druce, 1888 representing the Palaeotropical subfamily Aphnaeinae Lycaenidae: Lepidoptera). Relevant wing parts are illustrated, described, and some observations on the organs are briefly annotated. With an appendix and 14 figures.

  5. Dynamic control of a bistable wing under aerodynamic loading

    International Nuclear Information System (INIS)

    Bilgen, Onur; Arrieta, Andres F; Friswell, Michael I; Hagedorn, Peter

    2013-01-01

    The aerodynamic evaluation of a dynamic control technique applied to a bistable unsymmetrical cross-ply composite plate with surface bonded piezoelectric actuators is presented. The plate is clamped on one end to form a low-aspect-ratio wing. A previously proposed dynamic control method, utilizing bending resonance in different stable equilibrium positions, is used to induce snap-through between the two equilibrium states. Compared to quasi-static actuation, driving the bistable plate near resonance using surface bonded piezoelectric materials requires, theoretically, a lower peak excitation voltage to achieve snap-through. First, a set of extensive wind tunnel experiments are conducted on the passive bistable wing to understand the change in the dynamic behavior under various aerodynamic conditions. The passive wing demonstrated sufficient bending stiffness to sustain its shape under aerodynamic loading while preserving the desired bistable behavior. Next, by the use of the resonant control technique, the plate is turned into an effectively monostable structure, or alternatively, both stable equilibrium positions can be reached actively from the other stable equilibrium. Dynamic forward and reverse snap-through is demonstrated in the wind tunnel which shows both the effectiveness of the piezoelectric actuation as well as the load carrying capability of both states of the bistable wing. (paper)

  6. Aeroelastic Modelling and Design of Aeroelastically Tailored and Morphing Wings

    NARCIS (Netherlands)

    Werter, N.P.M.

    2017-01-01

    In order to accommodate the growth in air traffic whilst reducing the impact on the environment, operational efficiency is becoming more and more important in the design of the aircraft of the future. A possible approach to increase the operational efficiency of aircraft wings is the use of

  7. Rotational accelerations stabilize leading edge vortices on revolving fly wings

    NARCIS (Netherlands)

    Lentink, D.; Dickinson, M.H.

    2009-01-01

    The aerodynamic performance of hovering insects is largely explained by the presence of a stably attached leading edge vortex (LEV) on top of their wings. Although LEVs have been visualized on real, physically modeled, and simulated insects, the physical mechanisms responsible for their stability

  8. Effect of Winged Subsoiler and Traditional Tillage Integrated with ...

    African Journals Online (AJOL)

    Effect of Winged Subsoiler and Traditional Tillage Integrated with Fanya Juu on Selected Soil Physico-Chemical and Soil Water Properties in the Northwestern ... Soil evaporation was estimated by a conceptual model whereby leaf area index, canopy cover, crop root length, moisture at saturation and field capacity were ...

  9. 14 CFR 23.201 - Wings level stall.

    Science.gov (United States)

    2010-01-01

    ... 14 Aeronautics and Space 1 2010-01-01 2010-01-01 false Wings level stall. 23.201 Section 23.201 Aeronautics and Space FEDERAL AVIATION ADMINISTRATION, DEPARTMENT OF TRANSPORTATION AIRCRAFT AIRWORTHINESS.... Starting from a speed at least 10 knots above the stall speed, the elevator control must be pulled back so...

  10. Consequences of outbreeding on phenotypic plasticity in Drosophila mercatorum wings

    DEFF Research Database (Denmark)

    Krag, Kristian; Thomsen, Hans Paarup; Faurby, Søren

    2009-01-01

    for the wing shape. For all the other treatments higher DI was found in the F1 when the difference was significant, which is suggestive of outbreeding depression. These findings are difficult to interpret since an apparent heterotic effect of size at 25°C is accompanied by higher DI (though not significant...

  11. Calculation of Airloads for a Flexible Wing via NASTRAN.

    Science.gov (United States)

    1980-12-01

    IV. C oiiutiuULtiozial 1rcdu’.......... . .. .. . ..... interfaced NASTRAN -USSAERO Sequence .... 9 The DMAP Sequence.................11...not considered. A new instruction sequence known as a Direct Matrix Abstraction Program ( DMAP ) was developed for use within NASTRAN to ualculate...since the inclusion of aerodynamic theories in the NASTRAN program. The calculation of flexible wing airloads by this DMAP sequence is an iterative

  12. Flight mechanics of a tailless articulated wing aircraft.

    Science.gov (United States)

    Paranjape, Aditya A; Chung, Soon-Jo; Selig, Michael S

    2011-06-01

    This paper investigates the flight mechanics of a micro aerial vehicle without a vertical tail in an effort to reverse-engineer the agility of avian flight. The key to stability and control of such a tailless aircraft lies in the ability to control the incidence angles and dihedral angles of both wings independently. The dihedral angles can be varied symmetrically on both wings to control aircraft speed independently of the angle of attack and flight path angle, while asymmetric dihedral can be used to control yaw in the absence of a vertical stabilizer. It is shown that wing dihedral angles alone can effectively regulate sideslip during rapid turns and generate a wide range of equilibrium turn rates while maintaining a constant flight speed and regulating sideslip. Numerical continuation and bifurcation analysis are used to compute trim states and assess their stability. This paper lays the foundation for design and stability analysis of a flapping wing aircraft that can switch rapidly from flapping to gliding flight for agile manoeuvring in a constrained environment.

  13. Model identification of a flapping wing micro aerial vehicle

    NARCIS (Netherlands)

    Aguiar Vieira Caetano, J.V.

    2016-01-01

    Different flapping wing micro aerial vehicles (FWMAV) have been developed for academic (Harvard’s RoboBee), military (Israel Aerospace Industries’ Butterfly) and technology demonstration (Aerovironment’s NanoHummingBird) purposes. Among these, theDelFly II is recognized as one of themost successful

  14. Male secondary sexu al characters in Aphnaeinae wings (Lepidoptera: Lycaenidae

    Directory of Open Access Journals (Sweden)

    Bálint, Zsolt

    2017-04-01

    Full Text Available Male secondary sexual characters have been discovered on the hindwing verso of genera Aphnaeus Hübner, [1819], Cigaritis Donzel, 1847, Lipaphnaeus Aurivillius, 1916 and Pseudaletis Druce, 1888 representing the Palaeotropical subfamily Aphnaeinae Lycaenidae: Lepidoptera. Relevant wing parts are illustrated, described, and some observations on the organs are briefly annotated. With an appendix and 14 figures.

  15. Media coverage of right-wing populist leaders

    NARCIS (Netherlands)

    Bos, L.; van der Brug, W.; de Vreese, C.

    2010-01-01

    This article focuses on how leaders of new right-wing populist parties are portrayed in the mass media. More so than their established counterparts, new parties depend on the media for their electoral breakthrough. From a theoretical perspective, we expect prominence, populism, and authoritativeness

  16. Flow cytometry determination of ploidy level in winged bean ...

    African Journals Online (AJOL)

    Ploidy determination and mutation breeding of crop plants are inseparable twins given that mutation breeding is hinged majorly on polyploidization of crop's chromosome number. The present research was aimed at determining the ploidy level of 20 accessions of winged bean (Psophoscarpus tetragonolobus) using known ...

  17. Reinforcements in avian wing bones: Experiments, analysis, and modeling.

    Science.gov (United States)

    Novitskaya, E; Ruestes, C J; Porter, M M; Lubarda, V A; Meyers, M A; McKittrick, J

    2017-12-01

    Almost all species of modern birds are capable of flight; the mechanical competency of their wings and the rigidity of their skeletal system evolved to enable this outstanding feat. One of the most interesting examples of structural adaptation in birds is the internal structure of their wing bones. In flying birds, bones need to be sufficiently strong and stiff to withstand forces during takeoff, flight, and landing, with a minimum of weight. The cross-sectional morphology and presence of reinforcing structures (struts and ridges) found within bird wing bones vary from species to species, depending on how the wings are utilized. It is shown that both morphology and internal features increases the resistance to flexure and torsion with a minimum weight penalty. Prototypes of reinforcing struts fabricated by 3D printing were tested in diametral compression and torsion to validate the concept. In compression, the ovalization decreased through the insertion of struts, while they had no effect on torsional resistance. An elastic model of a circular ring reinforced by horizontal and vertical struts is developed to explain the compressive stiffening response of the ring caused by differently oriented struts. Copyright © 2017 Elsevier Ltd. All rights reserved.

  18. Sensory biology: echolocation from click to call, mouth to wing.

    Science.gov (United States)

    Fenton, M Brock; Ratcliffe, John M

    2014-12-15

    Echolocators use echoes of sounds they produce, clicks or calls, to detect objects. Usually, these signals originate from the head. New work reveals that three species of bats use their wings to generate echolocation signals. Copyright © 2014 Elsevier Ltd. All rights reserved.

  19. Experimental investigation into the feasibility of an extruded wing

    Science.gov (United States)

    Morelli, P.; Romeo, G.

    1979-01-01

    Fabrications of extruded aluminum alloy structures are reviewed. The design criteria and the fabrication of the main structure of a sailplane wing made of a few extruded profiles longitudinally connected one to the other are illustrated. Structural tests recently carried out are reported upon.

  20. Flight in slow motion: aerodynamics of the pterosaur wing.

    Science.gov (United States)

    Palmer, Colin

    2011-06-22

    The flight of pterosaurs and the extreme sizes of some taxa have long perplexed evolutionary biologists. Past reconstructions of flight capability were handicapped by the available aerodynamic data, which was unrepresentative of possible pterosaur wing profiles. I report wind tunnel tests on a range of possible pterosaur wing sections and quantify the likely performance for the first time. These sections have substantially higher profile drag and maximum lift coefficients than those assumed before, suggesting that large pterosaurs were aerodynamically less efficient and could fly more slowly than previously estimated. In order to achieve higher efficiency, the wing bones must be faired, which implies extensive regions of pneumatized tissue. Whether faired or not, the pterosaur wings were adapted to low-speed flight, unsuited to marine style dynamic soaring but adapted for thermal/slope soaring and controlled, low-speed landing. Because their thin-walled bones were susceptible to impact damage, slow flight would have helped to avoid injury and may have contributed to their attaining much larger sizes than fossil or extant birds. The trade-off would have been an extreme vulnerability to strong or turbulent winds both in flight and on the ground, akin to modern-day paragliders.

  1. Playback interference of glassy-winged sharp shooter communication

    Science.gov (United States)

    Animal communication is vital to reproduction, particularly for securing a mate. Insects commonly communicate by exchanging vibrational signals that are transmitted through host plants. The glassy-winged sharpshooter (GWSS), Homalodisca vitripennis, is an important vector of Xylella fastidiosa, a pl...

  2. Velocity and turbulence at a wing-wall abutment

    Indian Academy of Sciences (India)

    Experimental investigation of the 3D turbulent flow field around a 45° wing-wall abutment, resting on a rough rigid bed, is reported. The experiment was conducted ... The shear stresses acting on the bed around the abutment are estimated from the Reynolds stresses and velocity gradients. The data presented in this study ...

  3. Surfzone monitoring using rotary wing unmanned aerial vehicles

    NARCIS (Netherlands)

    Brouwer, R.L.; De Schipper, M.A.; Rynne, P.F.; Graham, F.J.; Reniers, A.J.H.M.; Macmahan, J.H.

    2015-01-01

    This study investigates the potential of rotary wing unmanned aerial vehicles (UAVs) to monitor the surfzone. This paper shows that these UAVs are extremely flexible surveying platforms that can gather nearcontinuous moderate spatial resolution and high temporal resolution imagery from a fixed

  4. The costae presenting in high-temperature-induced vestigial wings ...

    Indian Academy of Sciences (India)

    Judging by the costae presenting in the anterior wing margin, I propose that the proximal and distal costal sections are independent growth units. ... The distal costal section seems to be an extension of the radius vein. vestigial, one of the most intensely researched temperature-sensitive mutations, is a good candidate for ...

  5. Innovative Wing Structures for Improved Aerodynamic and Aeroelastic Performance

    Science.gov (United States)

    2016-06-09

    directions in the printer. The colour represent the volume of defects found by VG Studio Max holes/defects detection tool. 5.3 Conclusions on material... visual inspection of the skin surface of the different wings was performed to assess any noticeable differences. Skin surface finish is paramount for

  6. Interactive flutter analysis and parametric study for conceptual wing design

    Science.gov (United States)

    Mukhopadhyay, Vivek

    1995-01-01

    An interactive computer program was developed for wing flutter analysis in the conceptual design stage. The objective was to estimate the flutter instability boundary of a flexible cantilever wing, when well defined structural and aerodynamic data are not available, and then study the effect of change in Mach number, dynamic pressure, torsional frequency, sweep, mass ratio, aspect ratio, taper ratio, center of gravity, and pitch inertia, to guide the development of the concept. The software was developed on MathCad (trademark) platform for Macintosh, with integrated documentation, graphics, database and symbolic mathematics. The analysis method was based on nondimensional parametric plots of two primary flutter parameters, namely Regier number and Flutter number, with normalization factors based on torsional stiffness, sweep, mass ratio, aspect ratio, center of gravity location and pitch inertia radius of gyration. The plots were compiled in a Vaught Corporation report from a vast database of past experiments and wind tunnel tests. The computer program was utilized for flutter analysis of the outer wing of a Blended Wing Body concept, proposed by McDonnell Douglas Corporation. Using a set of assumed data, preliminary flutter boundary and flutter dynamic pressure variation with altitude, Mach number and torsional stiffness were determined.

  7. Velocity and turbulence at a wing-wall abutment

    Indian Academy of Sciences (India)

    R. Narasimhan (Krishtel eMaging) 1461 1996 Oct 15 13:05:22

    When a wing-wall abutment is placed vertically on a rigid-bed rectangular channel by attach- ing it to one of the vertical sidewalls of the channel, the approaching turbulent boundary layer undergoes separation and rolles up to form the well-known primary vortex, which swept out by the side of the abutment. Limited research ...

  8. A bio-inspired study on tidal energy extraction with flexible flapping wings.

    Science.gov (United States)

    Liu, Wendi; Xiao, Qing; Cheng, Fai

    2013-09-01

    Previous research on the flexible structure of flapping wings has shown an improved propulsion performance in comparison to rigid wings. However, not much is known about this function in terms of power efficiency modification for flapping wing energy devices. In order to study the role of the flexible wing deformation in the hydrodynamics of flapping wing energy devices, we computationally model the two-dimensional flexible single and twin flapping wings in operation under the energy extraction conditions with a large Reynolds number of 106. The flexible motion for the present study is predetermined based on a priori structural result which is different from a passive flexibility solution. Four different models are investigated with additional potential local distortions near the leading and trailing edges. Our simulation results show that the flexible structure of a wing is beneficial to enhance power efficiency by increasing the peaks of lift force over a flapping cycle, and tuning the phase shift between force and velocity to a favourable trend. Moreover, the impact of wing flexibility on efficiency is more profound at a low nominal effective angle of attack (AoA). At a typical flapping frequency f * = 0.15 and nominal effective AoA of 10°, a flexible integrated wing generates 7.68% higher efficiency than a rigid wing. An even higher increase, around six times that of a rigid wing, is achievable if the nominal effective AoA is reduced to zero degrees at feathering condition. This is very attractive for a semi-actuated flapping energy system, where energy input is needed to activate the pitching motion. The results from our dual-wing study found that a parallel twin-wing device can produce more power compared to a single wing due to the strong flow interaction between the two wings.

  9. Power reduction and the radial limit of stall delay in revolving wings of different aspect ratio.

    Science.gov (United States)

    Kruyt, Jan W; van Heijst, GertJan F; Altshuler, Douglas L; Lentink, David

    2015-04-06

    Airplanes and helicopters use high aspect ratio wings to reduce the power required to fly, but must operate at low angle of attack to prevent flow separation and stall. Animals capable of slow sustained flight, such as hummingbirds, have low aspect ratio wings and flap their wings at high angle of attack without stalling. Instead, they generate an attached vortex along the leading edge of the wing that elevates lift. Previous studies have demonstrated that this vortex and high lift can be reproduced by revolving the animal wing at the same angle of attack. How do flapping and revolving animal wings delay stall and reduce power? It has been hypothesized that stall delay derives from having a short radial distance between the shoulder joint and wing tip, measured in chord lengths. This non-dimensional measure of wing length represents the relative magnitude of inertial forces versus rotational accelerations operating in the boundary layer of revolving and flapping wings. Here we show for a suite of aspect ratios, which represent both animal and aircraft wings, that the attachment of the leading edge vortex on a revolving wing is determined by wing aspect ratio, defined with respect to the centre of revolution. At high angle of attack, the vortex remains attached when the local radius is shorter than four chord lengths and separates outboard on higher aspect ratio wings. This radial stall limit explains why revolving high aspect ratio wings (of helicopters) require less power compared with low aspect ratio wings (of hummingbirds) at low angle of attack and vice versa at high angle of attack. © 2015 The Author(s) Published by the Royal Society. All rights reserved.

  10. Petiolate wings: effects on the leading-edge vortex in flapping flight.

    Science.gov (United States)

    Phillips, Nathan; Knowles, Kevin; Bomphrey, Richard J

    2017-02-06

    The wings of many insect species including crane flies and damselflies are petiolate (on stalks), with the wing planform beginning some distance away from the wing hinge, rather than at the hinge. The aerodynamic impact of flapping petiolate wings is relatively unknown, particularly on the formation of the lift-augmenting leading-edge vortex (LEV): a key flow structure exploited by many insects, birds and bats to enhance their lift coefficient. We investigated the aerodynamic implications of petiolation P using particle image velocimetry flow field measurements on an array of rectangular wings of aspect ratio 3 and petiolation values of P = 1-3. The wings were driven using a mechanical device, the 'Flapperatus', to produce highly repeatable insect-like kinematics. The wings maintained a constant Reynolds number of 1400 and dimensionless stroke amplitude Λ * (number of chords traversed by the wingtip) of 6.5 across all test cases. Our results showed that for more petiolate wings the LEV is generally larger, stronger in circulation, and covers a greater area of the wing surface, particularly at the mid-span and inboard locations early in the wing stroke cycle. In each case, the LEV was initially arch-like in form with its outboard end terminating in a focus-sink on the wing surface, before transitioning to become continuous with the tip vortex thereafter. In the second half of the wing stroke, more petiolate wings exhibit a more detached LEV, with detachment initiating at approximately 70% and 50% span for P = 1 and 3, respectively. As a consequence, lift coefficients based on the LEV are higher in the first half of the wing stroke for petiolate wings, but more comparable in the second half. Time-averaged LEV lift coefficients show a general rise with petiolation over the range tested.

  11. Microarray Data Analysis of Space Grown Arabidopsis Leaves for Genes Important in Vascular Patterning. Analysis of Space Grown Arabidopsis with Microarray Data from GeneLab: Identification of Genes Important in Vascular Patterning

    Science.gov (United States)

    Weitzel, A. J.; Wyatt, S. E.; Parsons-Wingerter, P.

    2016-01-01

    Venation patterning in leaves is a major determinant of photosynthesis efficiency because of its dependency on vascular transport of photo-assimilates, water, and minerals. Arabidopsis thaliana grown in microgravity show delayed growth and leaf maturation. Gene expression data from the roots, hypocotyl, and leaves of A. thaliana grown during spaceflight vs. ground control analyzed by Affymetrix microarray are available through NASA's GeneLab (GLDS-7). We analyzed the data for differential expression of genes in leaves resulting from the effects of spaceflight on vascular patterning. Two genes were found by preliminary analysis to be up-regulated during spaceflight that may be related to vascular formation. The genes are responsible for coding an ARGOS (Auxin-Regulated Gene Involved in Organ Size)-like protein (potentially affecting cell elongation in the leaves), and an F-box/kelch-repeat protein (possibly contributing to protoxylem specification). Further analysis that will focus on raw data quality assessment and a moderated t-test may further confirm up-regulation of the two genes and/or identify other gene candidates. Plants defective in these genes will then be assessed for phenotype by the mapping and quantification of leaf vascular patterning by NASA's VESsel GENeration (VESGEN) software to model specific vascular differences of plants grown in spaceflight.

  12. Aerodynamic Design of Integrated Propulsion-Airframe Configuration of the Hybrid Wing-Body Aircraft

    Science.gov (United States)

    Liou, May-Fun; Kim, Hyoungjin; Lee, B. J.; Liou, Meng-Sing

    2017-01-01

    Hybrid Wing Body (HWB) aircraft is characterized by a flattened and airfoil-shaped body, which produces a substantial portion of the total lift. The body form is composed of distinct and separate wing structures, though the wings are smoothly blended into the body. This concept has been studied widely and results suggest remarkable performance improvements over the conventional tube and wing transport1,2. HWB incorporates design features from both a futuristic fuselage and flying wing design, which houses most of the crew, payload and equipment inside the main centerbody structure.

  13. Nanomechanical properties of wing membrane layers in the house cricket (Acheta domesticus Linnaeus).

    Science.gov (United States)

    Sample, Caitlin S; Xu, Alan K; Swartz, Sharon M; Gibson, Lorna J

    2015-03-01

    Many insect wings change shape dynamically during the wingbeat cycle, and these deformations have the potential to confer energetic and aerodynamic benefits during flight. Due to the lack of musculature within the wing itself, the changing form of the wing is determined primarily by its passive response to inertial and aerodynamic forces. This response is in part controlled by the wing's mechanical properties, which vary across the membrane to produce regions of differing stiffness. Previous studies of wing mechanical properties have largely focused on surface or bulk measurements, but this ignores the layered nature of the wing. In our work, we investigated the mechanical properties of the wings of the house cricket (Acheta domesticus) with the aim of determining differences between layers within the wing. Nanoindentation was performed on both the surface and the interior layers of cross-sectioned samples of the wing to measure the Young's modulus and hardness of the outer- and innermost layers. The results demonstrate that the interior of the wing is stiffer than the surface, and both properties vary across the wing. Copyright © 2015 Elsevier Ltd. All rights reserved.

  14. A Single-Wing Removal Method to Assess Correspondence Between Gene Expression and Phenotype in Butterflies: The Case of Distal-less.

    Science.gov (United States)

    Adhikari, Kiran; Otaki, Joji M

    2016-02-01

    It is often desirable but difficult to retrieve information on the mature phenotype of an immature tissue sample that has been subjected to gene expression analysis. This problem cannot be ignored when individual variation within a species is large. To circumvent this problem in the butterfly wing system, we developed a new surgical method for removing a single forewing from a pupa using Junonia orithya; the operated pupa was left to develop to an adult without eclosion. The removed right forewing was subjected to gene expression analysis, whereas the non-removed left forewing was examined for color patterns. As a test case, we focused on Distal-less (Dll), which likely plays an active role in inducing elemental patterns, including eyespots. The Dll expression level in forewings was paired with eyespot size data from the same individual. One third of the operated pupae survived and developed wing color patterns. Dll expression levels were significantly higher in males than in females, although male eyespots were smaller in size than female eyespots. Eyespot size data showed weak but significant correlations with the Dll expression level in females. These results demonstrate that a single-wing removal method was successfully applied to the butterfly wing system and suggest the weak and non-exclusive contribution of Dll to eyespot size determination in this butterfly. Our novel methodology for establishing correspondence between gene expression and phenotype can be applied to other candidate genes for color pattern development in butterflies. Conceptually similar methods may also be applicable in other developmental systems.

  15. Effect of delta wing on the particle flow in a novel gas supersonic separator

    DEFF Research Database (Denmark)

    Wen, Chuang; Yang, Yan; Walther, Jens Honore

    2016-01-01

    The present work presents numerical simulations of the complex particle motion in a supersonic separator with a delta wing located in the supersonic flow. The effect of the delta wing on the strong swirling flow is analysed using the Discrete Particle Method. The results show that the delta wings...... re-compress the upstream flow and the gas Mach number decreases correspondingly. However, the Mach number does not vary significantly from the small, medium and large delta wing configurations. The small delta wing generates a swirl near its surface, but has minor influences on the flow above it....... On the contrary, the use of the large delta wing produces a strong swirling flow in the whole downstream region. For the large delta wing, the collection efficiency reaches 70% with 2 μm particles, indicating a good separation performance of the proposed supersonic separator....

  16. Piezoelectric energy harvesting from morphing wing motions for micro air vehicles

    KAUST Repository

    Abdelkefi, Abdessattar

    2013-09-10

    Wing flapping and morphing can be very beneficial to managing the weight of micro air vehicles through coupling the aerodynamic forces with stability and control. In this letter, harvesting energy from the wing morphing is studied to power cameras, sensors, or communication devices of micro air vehicles and to aid in the management of their power. The aerodynamic loads on flapping wings are simulated using a three-dimensional unsteady vortex lattice method. Active wing shape morphing is considered to enhance the performance of the flapping motion. A gradient-based optimization algorithm is used to pinpoint the optimal kinematics maximizing the propellent efficiency. To benefit from the wing deformation, we place piezoelectric layers near the wing roots. Gauss law is used to estimate the electrical harvested power. We demonstrate that enough power can be generated to operate a camera. Numerical analysis shows the feasibility of exploiting wing morphing to harvest energy and improving the design and performance of micro air vehicles.

  17. Force measurements of flexible tandem wings in hovering and forward flights

    International Nuclear Information System (INIS)

    Zheng, Yingying; Wu, Yanhua; Tang, Hui

    2015-01-01

    Aerodynamic forces, power consumptions and efficiencies of flexible and rigid tandem wings undergoing combined plunging/pitching motion were measured in a hovering flight and two forward flights with Strouhal numbers of 0.6 and 0.3. Three flexible dragonfly-like tandem wing models termed Wing I, Wing II, and Wing III which are progressively less flexible, as well as a pair of rigid wings as the reference were operated at three phase differences of 0°, 90° and 180°. The results showed that both the flexibility and phase difference have significant effects on the aerodynamic performances. In both hovering and forward flights at a higher oscillation frequency of 1 Hz (St = 0.6), the Wing III model outperformed the other wing models with larger total horizontal force coefficient and efficiency. In forward flight at the lower frequency of 0.5 Hz (St = 0.3), Wing III, rigid wings and Wing II models performed best at 0°, 90° and 180° phase difference, respectively. From the time histories of force coefficients of fore- and hind-wings, different peak values, phase lags, and secondary peaks were found to be the important reasons to cause the differences in the average horizontal force coefficients. Particle image velocimetry and deformation measurements were performed to provide the insights into how the flexibility affects the aerodynamic performance of the tandem wings. The spanwise bending deformation was found to contribute to the horizontal force, by offering a more beneficial position to make LEV more attached to the wing model in both hovering and forward flights, and inducing a higher-velocity region in forward flight. (paper)

  18. DAST in Flight just after Structural Failure of Right Wing

    Science.gov (United States)

    1980-01-01

    Two BQM-34 Firebee II drones were modified with supercritical airfoils, called the Aeroelastic Research Wing (ARW), for the Drones for Aerodynamic and Structural Testing (DAST) program, which ran from 1977 to 1983. This photo, taken 12 June 1980, shows the DAST-1 (Serial #72-1557) immediately after it lost its right wing after suffering severe wing flutter. The vehicle crashed near Cuddeback Dry Lake. The Firebee II was selected for the DAST program because its standard wing could be removed and replaced by a supercritical wing. The project's digital flutter suppression system was intended to allow lighter wing structures, which would translate into better fuel economy for airliners. Because the DAST vehicles were flown intentionally at speeds and altitudes that would cause flutter, the program anticipated that crashes might occur. These are the image contact sheets for each image resolution of the NASA Dryden Drones for Aerodynamic and Structural Testing (DAST) Photo Gallery. From 1977 to 1983, the Dryden Flight Research Center, Edwards, California, (under two different names) conducted the DAST Program as a high-risk flight experiment using a ground-controlled, pilotless aircraft. Described by NASA engineers as a 'wind tunnel in the sky,' the DAST was a specially modified Teledyne-Ryan BQM-34E/F Firebee II supersonic target drone that was flown to validate theoretical predictions under actual flight conditions in a joint project with the Langley Research Center, Hampton, Virginia. The DAST Program merged advances in electronic remote control systems with advances in airplane design. Drones (remotely controlled, missile-like vehicles initially developed to serve as gunnery targets) had been deployed successfully during the Vietnamese conflict as reconnaissance aircraft. After the war, the energy crisis of the 1970s led NASA to seek new ways to cut fuel use and improve airplane efficiency. The DAST Program's drones provided an economical, fuel-conscious method for

  19. Myrmeleon almohadarum sp. nov., from Spain and North Africa, with description of the larva (Neuroptera Myrmeleontidae).

    Science.gov (United States)

    Badano, Davide; Acevedo, Fernando; Pantaleoni, Roberto A; Monserrat, Víctor J

    2016-11-22

    A new antlion (Neuroptera Myrmeleontidae Myrmeleontini), Myrmeleon almohadarum  sp. nov., is described from southern Spain and Tunisia. The new taxon is closely related  to the mostly sympatric M. inconspicuus Rambur and M. mariaemathildae Pantaleoni, Cesaroni & Nicoli Aldini but differing in body pattern, wing venation and larval chaetotaxy. The validity of the new species is also supported by a phylogenetic analysis based on COI sequences. The larva of this new species is described and compared with congeners. M. almohadarum appears to be associated with sandy environments.

  20. How lizards fly: A novel type of wing in animals.

    Directory of Open Access Journals (Sweden)

    J Maximilian Dehling

    Full Text Available Flying lizards of the genus Draco are renowned for their gliding ability, using an aerofoil formed by winglike patagial membranes and supported by elongated thoracic ribs. It remains unknown, however, how these lizards manoeuvre during flight. Here, I present the results of a study on the aerial behaviour of Dussumier's Flying Lizard (Draco dussumieri and show that Draco attaches the forelimbs to the leading edge of the patagium while airborne, forming a hitherto unknown type of composite wing. The attachment of the forelimbs to the patagium suggests that that aerofoil is controlled through movements of the forelimbs. One major advantage for the lizards is that the forelimbs retain their complete range of movement and functionality for climbing and running when not used as a part of the wing. These findings not only shed a new light on the flight of Draco but also have implications for the interpretation of gliding performance in fossil species.

  1. Structural colors from Morpho peleides butterfly wing scales

    KAUST Repository

    Ding, Yong

    2009-01-01

    A male Morpho peleides butterfly wing is decorated by two types of scales, cover and ground scales. We have studied the optical properties of each type of scales in conjunction with the structural information provided by cross-sectional transmission electron microscopy and computer simulation. The shining blue color is mainly from the Bragg reflection of the one-dimensional photonic structure, e.g., the shelf structure packed regularly in each ridges on cover scales. A thin-film-like interference effect from the base plate of the cover scale enhances such blue color and further gives extra reflection peaks in the infrared and ultraviolet regions. The analogy in the spectra acquired from the original wing and that from the cover scales suggests that the cover scales take a dominant role in its structural color. This study provides insight of using the biotemplates for fabricating smart photonic structures. © 2009 American Institute of Physics.

  2. Analysis and optimization of a camber morphing wing model

    Directory of Open Access Journals (Sweden)

    Bing Li

    2016-09-01

    Full Text Available This article proposes a camber morphing wing model that can continuously change its camber. A mathematical model is proposed and a kinematic simulation is performed to verify the wing’s ability to change camber. An aerodynamic model is used to test its aerodynamic characteristics. Some important aerodynamic analyses are performed. A comparative analysis is conducted to explore the relationships between aerodynamic parameters, the rotation angle of the trailing edge, and the angle of attack. An improved artificial fish swarm optimization algorithm is proposed, referred to as the weighted adaptive artificial fish-swarm with embedded Hooke–Jeeves search method. Some comparison tests are used to test the performance of the improved optimization algorithm. Finally, the proposed optimization algorithm is used to optimize the proposed camber morphing wing model.

  3. Comments on prospects of fully adaptive aircraft wings

    Science.gov (United States)

    Inman, Daniel J.; Gern, Frank H.; Robertshaw, Harry H.; Kapania, Rakesh K.; Pettit, Greg; Natarajan, Anand; Sulaeman, Erwin

    2001-06-01

    New generations of highly maneuverable aircraft, such as Uninhabited Combat Air Vehicles (UCAV) or Micro Air Vehicles (MAV) are likely to feature very flexible lifting surfaces. To enhance stealth properties and performance, the replacement of hinged control surfaces by smart wings and morphing airfoils is investigated. This requires a fundamental understanding of the interaction between aerodynamics, structures, and control systems. The goal is to build a model consistent with distributed control and to exercise this model to determine the progress possible in terms of flight control (lift, drag and maneuver performance) with an adaptive wing. Different modeling levels are examined and combined with a variety of distributed control approaches to determine what types of maneuvers and flight regimes may be possible. This paper describes the current progress of the project and highlights some recent findings.

  4. Structural design studies of a supersonic cruise arrow wing configuration

    Science.gov (United States)

    Sobieszczanski, J.; Mccullers, L. A.; Ricketts, R. H.; Santoro, N. J.; Beskenis, S. D.; Kurtze, W. L.

    1976-01-01

    Structural member cross sections were sized with a system of integrated computer programs to satisfy strength and flutter design requirements for several variants of the arrow wing supersonic cruise vehicle. The resulting structural weights provide a measure of the structural efficiency of the planform geometry, structural layout, type of construction, and type of material including composites. The material distribution was determined for a baseline metallic structure and the results indicate that an approximate fatigue constraint has an important effect on the structural weight required for strength but, in all cases, additional material had to be added to satisfy flutter requirements with lighter mass engines with minimum fuel onboard. The use of composite materials on the baseline configuration was explored and indicated increased structural efficiency. In the strength sizing, the all-composite construction provided a lower weight design than the hybrid construction which contained composites only in the wing cover skins. Subsequent flutter analyses indicated a corresponding lower flutter speed.

  5. Gust Load Alleviation with Robust Control for a Flexible Wing

    Directory of Open Access Journals (Sweden)

    Xiang Liu

    2016-01-01

    Full Text Available Traditional methods for gust alleviation of aircraft are mostly proposed based on a specific flight condition. In this paper, robust control laws are designed for a large flexible wing with uncertainty in Mach number and dynamic pressure. To accurately describe the aeroelastic model over a large flight envelope, a nonlinear parameter-varying model is developed which is a function of both Mach number and dynamic pressure. Then a linear fractional transformation is established accordingly and a modified model order reduction technique is applied to reduce the size of the uncertainty block. The developed model, in which the statistic nature of the gust is considered by using the Dryden power spectral density function, enables the use of μ-synthesis procedures for controller design. The simulations show that the μ controller can always effectively reduce the wing root shear force and bending moment at a given range of Mach number and dynamic pressure.

  6. Probabilistic Structural Health Monitoring of the Orbiter Wing Leading Edge

    Science.gov (United States)

    Yap, Keng C.; Macias, Jesus; Kaouk, Mohamed; Gafka, Tammy L.; Kerr, Justin H.

    2011-01-01

    A structural health monitoring (SHM) system can contribute to the risk management of a structure operating under hazardous conditions. An example is the Wing Leading Edge Impact Detection System (WLEIDS) that monitors the debris hazards to the Space Shuttle Orbiter s Reinforced Carbon-Carbon (RCC) panels. Since Return-to-Flight (RTF) after the Columbia accident, WLEIDS was developed and subsequently deployed on board the Orbiter to detect ascent and on-orbit debris impacts, so as to support the assessment of wing leading edge structural integrity prior to Orbiter re-entry. As SHM is inherently an inverse problem, the analyses involved, including those performed for WLEIDS, tend to be associated with significant uncertainty. The use of probabilistic approaches to handle the uncertainty has resulted in the successful implementation of many development and application milestones.

  7. Pretreated Butterfly Wings for Tuning the Selective Vapor Sensing

    Science.gov (United States)

    Piszter, Gábor; Kertész, Krisztián; Bálint, Zsolt; Biró, László Péter

    2016-01-01

    Photonic nanoarchitectures occurring in the scales of Blue butterflies are responsible for their vivid blue wing coloration. These nanoarchitectures are quasi-ordered nanocomposites which are constituted from a chitin matrix with embedded air holes. Therefore, they can act as chemically selective sensors due to their color changes when mixing volatile vapors in the surrounding atmosphere which condensate into the nanoarchitecture through capillary condensation. Using a home-built vapor-mixing setup, the spectral changes caused by the different air + vapor mixtures were efficiently characterized. It was found that the spectral shift is vapor-specific and proportional with the vapor concentration. We showed that the conformal modification of the scale surface by atomic layer deposition and by ethanol pretreatment can significantly alter the optical response and chemical selectivity, which points the way to the efficient production of sensor arrays based on the knowledge obtained through the investigation of modified butterfly wings. PMID:27618045

  8. Waste Assessment Baseline for the IPOC Second Floor, West Wing

    Energy Technology Data Exchange (ETDEWEB)

    McCord, Samuel A [Sandia National Lab. (SNL-NM), Albuquerque, NM (United States). Waste Management and Pollution Prevention

    2015-04-01

    Following a building-wide waste assessment in September, 2014, and subsequent presentation to Sandia leadership regarding the goal of Zero Waste by 2025, the occupants of the IPOC Second Floor, West Wing contacted the Materials Sustainability and Pollution Prevention (MSP2) team to guide them to Zero Waste in advance of the rest of the site. The occupants are from Center 3600, Public Relations and Communications , and Center 800, Independent Audit, Ethics and Business Conduct . To accomplish this, MSP2 conducted a new limited waste assessment from March 2-6, 2015 to compare the second floor, west wing to the building as a whole. The assessment also serves as a baseline with which to mark improvements in diversion in approximately 6 months.

  9. Analysis of Kinematics of Flapping Wing UAV Using OptiTrack Systems

    Directory of Open Access Journals (Sweden)

    Matthew Ng Rongfa

    2016-07-01

    Full Text Available An analysis of the kinematics of a flapping membrane wing using experimental kinematic data is presented. This motion capture technique tracks the positon of the retroreflective marker(s placed on the left wing of a 1.3-m-wingspan ornithopter. The time-varying three-dimensional data of the wing kinematics were recorded for a single frequency. The wing shape data was then plotted on a two-dimensional plane to understand the wing dynamic behaviour of an ornithopter. Specifically, the wing tip path, leading edge bending, wing membrane shape, local twist, stroke angle and wing velocity were analyzed. As the three characteristic angles can be expressed in the Fourier series as a function of time, the kinematics of the wing can be computationally generated for the aerodynamic study of flapping flight through the Fourier coefficients presented. Analysis of the ornithopter wing showed how the ornithopter closely mimics the flight motions of birds despite several physical limitations.

  10. Flow structure and aerodynamic performance of a hovering bristled wing in low Re

    Science.gov (United States)

    Lee, Seunghun; Lahooti, Mohsen; Kim, Daegyoum

    2017-11-01

    Previous studies on a bristled wing have mainly focused on simple kinematics of the wing such as translation or rotation. The aerodynamic performance of a bristled wing in a quasi-steady phase is known to be comparable to that of a smooth wing without a gap because shear layers in the gaps of the bristled wing are sufficiently developed to block the gaps. However, we point out that, in the starting transient phase where the shear layers are not fully developed, the force generation of a bristled wing is not as efficient as that of a quasi-steady state. The performance in the transient phase is important to understand the aerodynamics of a bristled wing in an unsteady motion. In the hovering motion, due to repeated stroke reversals, the formation and development of shear layers inside the gaps is repeated in each stroke. In this study, a bristled wing in hovering is numerically investigated in the low Reynolds number of O(10). We especially focus on the development of shear layers during a stroke reversal and its effect on the overall propulsive performance. Although the aerodynamic force generation is slightly reduced due to the gap vortices, the asymmetric behavior of vortices in a gap between bristles during a stroke reversal makes the bristled wing show higher lift to drag ratio than a smooth wing.

  11. Phasing of dragonfly wings can improve aerodynamic efficiency by removing swirl.

    Science.gov (United States)

    Usherwood, James R; Lehmann, Fritz-Olaf

    2008-11-06

    Dragonflies are dramatic, successful aerial predators, notable for their flight agility and endurance. Further, they are highly capable of low-speed, hovering and even backwards flight. While insects have repeatedly modified or reduced one pair of wings, or mechanically coupled their fore and hind wings, dragonflies and damselflies have maintained their distinctive, independently controllable, four-winged form for over 300Myr. Despite efforts at understanding the implications of flapping flight with two pairs of wings, previous studies have generally painted a rather disappointing picture: interaction between fore and hind wings reduces the lift compared with two pairs of wings operating in isolation. Here, we demonstrate with a mechanical model dragonfly that, despite presenting no advantage in terms of lift, flying with two pairs of wings can be highly effective at improving aerodynamic efficiency. This is achieved by recovering energy from the wake wasted as swirl in a manner analogous to coaxial contra-rotating helicopter rotors. With the appropriate fore-hind wing phasing, aerodynamic power requirements can be reduced up to 22 per cent compared with a single pair of wings, indicating one advantage of four-winged flying that may apply to both dragonflies and, in the future, biomimetic micro air vehicles.

  12. Computational Aerodynamic Analysis of a Micro-CT Based Bio-Realistic Fruit Fly Wing

    Science.gov (United States)

    Brandt, Joshua; Doig, Graham; Tsafnat, Naomi

    2015-01-01

    The aerodynamic features of a bio-realistic 3D fruit fly wing in steady state (snapshot) flight conditions were analyzed numerically. The wing geometry was created from high resolution micro-computed tomography (micro-CT) of the fruit fly Drosophila virilis. Computational fluid dynamics (CFD) analyses of the wing were conducted at ultra-low Reynolds numbers ranging from 71 to 200, and at angles of attack ranging from -10° to +30°. It was found that in the 3D bio-realistc model, the corrugations of the wing created localized circulation regions in the flow field, most notably at higher angles of attack near the wing tip. Analyses of a simplified flat wing geometry showed higher lift to drag performance values for any given angle of attack at these Reynolds numbers, though very similar performance is noted at -10°. Results have indicated that the simplified flat wing can successfully be used to approximate high-level properties such as aerodynamic coefficients and overall performance trends as well as large flow-field structures. However, local pressure peaks and near-wing flow features induced by the corrugations are unable to be replicated by the simple wing. We therefore recommend that accurate 3D bio-realistic geometries be used when modelling insect wings where such information is useful. PMID:25954946

  13. Effect of tip vortices on membrane vibration of flexible wings with different aspect ratios

    Directory of Open Access Journals (Sweden)

    Genç Mustafa Serdar

    2016-01-01

    Full Text Available In this study, the effect of the aspect ratio on the aerodynamics characteristic of flexible membrane wings with different aspect ratios (AR = 1 and AR = 3 is experimentally investigated at Reynolds number of 25000. Time accurate measurements of membrane deformation using Digital Image Correlation system (DIC is carried out while normal forces of the wing will be measured by helping a load-cell system and flow on the wing was visualized by means of smoke wire technic. The characteristics of high aspect ratio wings are shown to be affected by leading edge separation bubbles at low Reynolds number. It is concluded that the camber of membrane wing excites the separated shear layer and this situation increases the lift coefficient relatively more as compared to rigid wings. In membrane wings with low aspect ratio, unsteadiness included tip vortices and vortex shedding, and the combination of tip vortices and vortex shedding causes complex unsteady deformations of these membrane wings. The characteristic of high aspect ratio wings was shown to be affected by leading edge separation bubbles at low Reynolds numbers whereas the deformations of flexible wing with low aspect ratio affected by tip vortices and leading edge separation bubbles.

  14. Time-varying wing-twist improves aerodynamic efficiency of forward flight in butterflies.

    Science.gov (United States)

    Zheng, Lingxiao; Hedrick, Tyson L; Mittal, Rajat

    2013-01-01

    Insect wings can undergo significant chordwise (camber) as well as spanwise (twist) deformation during flapping flight but the effect of these deformations is not well understood. The shape and size of butterfly wings leads to particularly large wing deformations, making them an ideal test case for investigation of these effects. Here we use computational models derived from experiments on free-flying butterflies to understand the effect of time-varying twist and camber on the aerodynamic performance of these insects. High-speed videogrammetry is used to capture the wing kinematics, including deformation, of a Painted Lady butterfly (Vanessa cardui) in untethered, forward flight. These experimental results are then analyzed computationally using a high-fidelity, three-dimensional, unsteady Navier-Stokes flow solver. For comparison to this case, a set of non-deforming, flat-plate wing (FPW) models of wing motion are synthesized and subjected to the same analysis along with a wing model that matches the time-varying wing-twist observed for the butterfly, but has no deformation in camber. The simulations show that the observed butterfly wing (OBW) outperforms all the flat-plate wings in terms of usable force production as well as the ratio of lift to power by at least 29% and 46%, respectively. This increase in efficiency of lift production is at least three-fold greater than reported for other insects. Interestingly, we also find that the twist-only-wing (TOW) model recovers much of the performance of the OBW, demonstrating that wing-twist, and not camber is key to forward flight in these insects. The implications of this on the design of flapping wing micro-aerial vehicles are discussed.

  15. Time-varying wing-twist improves aerodynamic efficiency of forward flight in butterflies.

    Directory of Open Access Journals (Sweden)

    Lingxiao Zheng

    Full Text Available Insect wings can undergo significant chordwise (camber as well as spanwise (twist deformation during flapping flight but the effect of these deformations is not well understood. The shape and size of butterfly wings leads to particularly large wing deformations, making them an ideal test case for investigation of these effects. Here we use computational models derived from experiments on free-flying butterflies to understand the effect of time-varying twist and camber on the aerodynamic performance of these insects. High-speed videogrammetry is used to capture the wing kinematics, including deformation, of a Painted Lady butterfly (Vanessa cardui in untethered, forward flight. These experimental results are then analyzed computationally using a high-fidelity, three-dimensional, unsteady Navier-Stokes flow solver. For comparison to this case, a set of non-deforming, flat-plate wing (FPW models of wing motion are synthesized and subjected to the same analysis along with a wing model that matches the time-varying wing-twist observed for the butterfly, but has no deformation in camber. The simulations show that the observed butterfly wing (OBW outperforms all the flat-plate wings in terms of usable force production as well as the ratio of lift to power by at least 29% and 46%, respectively. This increase in efficiency of lift production is at least three-fold greater than reported for other insects. Interestingly, we also find that the twist-only-wing (TOW model recovers much of the performance of the OBW, demonstrating that wing-twist, and not camber is key to forward flight in these insects. The implications of this on the design of flapping wing micro-aerial vehicles are discussed.

  16. A computational study on the influence of insect wing geometry on bee flight mechanics

    Directory of Open Access Journals (Sweden)

    Jeffrey Feaster

    2017-12-01

    Full Text Available Two-dimensional computational fluid dynamics (CFD is applied to better understand the effects of wing cross-sectional morphology on flow field and force production. This study investigates the influence of wing cross-section on insect scale flapping flight performance, for the first time, using a morphologically representative model of a bee (Bombus pensylvanicus wing. The bee wing cross-section was determined using a micro-computed tomography scanner. The results of the bee wing are compared with flat and elliptical cross-sections, representative of those used in modern literature, to determine the impact of profile variation on aerodynamic performance. The flow field surrounding each cross-section and the resulting forces are resolved using CFD for a flight speed range of 1 to 5 m/s. A significant variation in vortex formation is found when comparing the ellipse and flat plate with the true bee wing. During the upstroke, the bee and approximate wing cross-sections have a much shorter wake structure than the flat plate or ellipse. During the downstroke, the flat plate and elliptical cross-sections generate a single leading edge vortex, while the approximate and bee wings generate numerous, smaller structures that are shed throughout the stroke. Comparing the instantaneous aerodynamic forces on the wing, the ellipse and flat plate sections deviate progressively with velocity from the true bee wing. Based on the present findings, a simplified cross-section of an insect wing can misrepresent the flow field and force production. We present the first aerodynamic study using a true insect wing cross-section and show that the wing corrugation increases the leading edge vortex formation frequency for a given set of kinematics.

  17. Aerodynamic consequences of wing morphing during emulated take-off and gliding in birds.

    Science.gov (United States)

    Klaassen van Oorschot, Brett; Mistick, Emily A; Tobalske, Bret W

    2016-10-01

    Birds morph their wings during a single wingbeat, across flight speeds and among flight modes. Such morphing may allow them to maximize aerodynamic performance, but this assumption remains largely untested. We tested the aerodynamic performance of swept and extended wing postures of 13 raptor species in three families (Accipitridae, Falconidae and Strigidae) using a propeller model to emulate mid-downstroke of flapping during take-off and a wind tunnel to emulate gliding. Based on previous research, we hypothesized that (1) during flapping, wing posture would not affect maximum ratios of vertical and horizontal force coefficients (C V :C H ), and that (2) extended wings would have higher maximum C V :C H when gliding. Contrary to each hypothesis, during flapping, extended wings had, on average, 31% higher maximum C V :C H ratios and 23% higher C V than swept wings across all biologically relevant attack angles (α), and, during gliding, maximum C V :C H ratios were similar for the two postures. Swept wings had 11% higher C V than extended wings in gliding flight, suggesting flow conditions around these flexed raptor wings may be different from those in previous studies of swifts (Apodidae). Phylogenetic affiliation was a poor predictor of wing performance, due in part to high intrafamilial variation. Mass was only significantly correlated with extended wing performance during gliding. We conclude that wing shape has a greater effect on force per unit wing area during flapping at low advance ratio, such as take-off, than during gliding. © 2016. Published by The Company of Biologists Ltd.

  18. Utilization of Optimization for Design of Morphing Wing Structures for Enhanced Flight

    Science.gov (United States)

    Detrick, Matthew Scott

    Conventional aircraft control surfaces constrain maneuverability. This work is a comprehensive study that looks at both smart material and conventional actuation methods to achieve wing twist to potentially improve flight capability using minimal actuation energy while allowing minimal wing deformation under aerodynamic loading. A continuous wing is used in order to reduce drag while allowing the aircraft to more closely approximate the wing deformation used by birds while loitering. The morphing wing for this work consists of a skin supported by an underlying truss structure whose goal is to achieve a given roll moment using less actuation energy than conventional control surfaces. A structural optimization code has been written in order to achieve minimal wing deformation under aerodynamic loading while allowing wing twist under actuation. The multi-objective cost function for the optimization consists of terms that ensure small deformation under aerodynamic loading, small change in airfoil shape during wing twist, a linear variation of wing twist along the length of the wing, small deviation from the desired wing twist, minimal number of truss members, minimal wing weight, and minimal actuation energy. Hydraulic cylinders and a two member linkage driven by a DC motor are tested separately to provide actuation. Since the goal of the current work is simply to provide a roll moment, only one actuator is implemented along the wing span. Optimization is also used to find the best location within the truss structure for the actuator. The active structure produced by optimization is then compared to simulated and experimental results from other researchers as well as characteristics of conventional aircraft.

  19. A computational study on the influence of insect wing geometry on bee flight mechanics.

    Science.gov (United States)

    Feaster, Jeffrey; Battaglia, Francine; Bayandor, Javid

    2017-12-15

    Two-dimensional computational fluid dynamics (CFD) is applied to better understand the effects of wing cross-sectional morphology on flow field and force production. This study investigates the influence of wing cross-section on insect scale flapping flight performance, for the first time, using a morphologically representative model of a bee ( Bombus pensylvanicus ) wing. The bee wing cross-section was determined using a micro-computed tomography scanner. The results of the bee wing are compared with flat and elliptical cross-sections, representative of those used in modern literature, to determine the impact of profile variation on aerodynamic performance. The flow field surrounding each cross-section and the resulting forces are resolved using CFD for a flight speed range of 1 to 5 m/s. A significant variation in vortex formation is found when comparing the ellipse and flat plate with the true bee wing. During the upstroke, the bee and approximate wing cross-sections have a much shorter wake structure than the flat plate or ellipse. During the downstroke, the flat plate and elliptical cross-sections generate a single leading edge vortex, while the approximate and bee wings generate numerous, smaller structures that are shed throughout the stroke. Comparing the instantaneous aerodynamic forces on the wing, the ellipse and flat plate sections deviate progressively with velocity from the true bee wing. Based on the present findings, a simplified cross-section of an insect wing can misrepresent the flow field and force production. We present the first aerodynamic study using a true insect wing cross-section and show that the wing corrugation increases the leading edge vortex formation frequency for a given set of kinematics. © 2017. Published by The Company of Biologists Ltd.

  20. Study on airflow characteristics of rear wing of F1 car

    Science.gov (United States)

    Azmi, A. R. S.; Sapit, A.; Mohammed, A. N.; Razali, M. A.; Sadikin, A.; Nordin, N.

    2017-09-01

    The paper aims to investigate CFD simulation is carried out to investigate the airflow along the rear wing of F1 car with Reynold number of 3 × 106 and velocity, u = 43.82204 m/s. The analysis was done using 2-D model consists of main plane and flap wing, combined together to form rear wing module. Both of the aerofoil is placed inside a box of 350mm long and 220mm height according to regulation set up by FIA. The parameters for this study is the thickness and the chord length of the flap wing aerofoil. The simulations were performed by using FLUENT solver and k-kl-omega model. The wind speed is set up to 43 m/s that is the average speed of F1 car when cornering. This study uses NACA 2408, 2412, and 2415 for the flap wing and BE50 for the main plane. Each cases being simulated with a gap between the aerofoil of 10mm and 50mm when the DRS is activated. Grid independence test and validation was conduct to make sure the result obtained is acceptable. The goal of this study is to investigate aerodynamic behavior of airflow around the rear wing as well as to see how the thickness and the chord length of flap wing influence the airflow at the rear wing. The results show that increasing in thickness of the flap wing aerofoil will decreases the downforce. The results also show that although the short flap wing generate lower downforce than the big flap wing, but the drag force can be significantly reduced as the short flap wing has more change in angle of attack when it is activated. Therefore, the type of aerofoil for the rear wing should be decided according to the circuit track so that it can be fully optimized.

  1. Time-Varying Wing-Twist Improves Aerodynamic Efficiency of Forward Flight in Butterflies

    Science.gov (United States)

    Zheng, Lingxiao; Hedrick, Tyson L.; Mittal, Rajat

    2013-01-01

    Insect wings can undergo significant chordwise (camber) as well as spanwise (twist) deformation during flapping flight but the effect of these deformations is not well understood. The shape and size of butterfly wings leads to particularly large wing deformations, making them an ideal test case for investigation of these effects. Here we use computational models derived from experiments on free-flying butterflies to understand the effect of time-varying twist and camber on the aerodynamic performance of these insects. High-speed videogrammetry is used to capture the wing kinematics, including deformation, of a Painted Lady butterfly (Vanessa cardui) in untethered, forward flight. These experimental results are then analyzed computationally using a high-fidelity, three-dimensional, unsteady Navier-Stokes flow solver. For comparison to this case, a set of non-deforming, flat-plate wing (FPW) models of wing motion are synthesized and subjected to the same analysis along with a wing model that matches the time-varying wing-twist observed for the butterfly, but has no deformation in camber. The simulations show that the observed butterfly wing (OBW) outperforms all the flat-plate wings in terms of usable force production as well as the ratio of lift to power by at least 29% and 46%, respectively. This increase in efficiency of lift production is at least three-fold greater than reported for other insects. Interestingly, we also find that the twist-only-wing (TOW) model recovers much of the performance of the OBW, demonstrating that wing-twist, and not camber is key to forward flight in these insects. The implications of this on the design of flapping wing micro-aerial vehicles are discussed. PMID:23341923

  2. THE AERODYNAMIC ANALYSIS OF THE PROFILES FOR FLYING WINGS

    Directory of Open Access Journals (Sweden)

    Vasile PRISACARIU

    2013-01-01

    Full Text Available The possibility of using an un-piloted aerial vector is determined by the aerodynamic characteristics and performances. The design for a tailless unmanned aerial vehicles starts from defining the aerial vector mission and implies o series of geometrical and aerodynamic aspects for stability. This article proposes to remark the aerodynamic characteristics of three profiles used at flying wing airship through 2D software analysis.

  3. Transonic Wing Shape Optimization Using a Genetic Algorithm

    Science.gov (United States)

    Holst, Terry L.; Pulliam, Thomas H.; Kwak, Dochan (Technical Monitor)

    2002-01-01

    A method for aerodynamic shape optimization based on a genetic algorithm approach is demonstrated. The algorithm is coupled with a transonic full potential flow solver and is used to optimize the flow about transonic wings including multi-objective solutions that lead to the generation of pareto fronts. The results indicate that the genetic algorithm is easy to implement, flexible in application and extremely reliable.

  4. A program for scheduling a Patrol Air Wing training plan.

    OpenAIRE

    Hutson, David V.

    1988-01-01

    Approved for public release; distribution in unlimited. This research examined the feasibility of a computerized scheduling system to assist the development of an annual training plan for a Patrol Air Wing. A prototype is proposed incorporating a modified A* search control structure to handle the combinatorial part of the problem. The system uses a pre-existing file for its database and is implemented on an ISI workstation using the Prolog computer language. Comparisons with...

  5. A Numerical Study of Vortex Dynamics of Flexible Wing Propulsors

    Science.gov (United States)

    2011-03-30

    Final Report Title: A numerical study of vortex dynamics of flexible wing propulsors AFOSR/AOARD Reference Number: AOARD-09-4077 AFOSR/AOARD Program ...NUMBER FA23860914077 5b. GRANT NUMBER 5c. PROGRAM ELEMENT NUMBER 6. AUTHOR(S) Kartik Venkatraman 5d. PROJECT NUMBER 5e. TASK NUMBER 5f. WORK...trailing-edge amplitude showed variations with change in filament length though the frequency of flapping was almost constant. Fitt & Pope (2001) showed the

  6. Insecticides sought to control adult glassy-winged sharpshooter

    OpenAIRE

    Akey, David H.; Henneberry, Thomas J.; Toscano, Nick

    2001-01-01

    The bacterium that causes Pierce's disease (Xylella fastidiosa) is transmitted to grapevines by the glassy-winged sharpshooter (GWSS). Insecticides were evaluated for efficacy and residual activity against adult GWSS on grapevines. Ten insecticides were tested in the cyclo-chlorinated, carbamate, organic phosphate, pyrethroid and neonicotinoid chemical classes. Results from field trials indicate that the pyrethroids and neonicotinoids are promising control agents. Information on efficacious a...

  7. Wing Infrastructure and Development Outlook (WINDO) Final Environmental Assessment

    Science.gov (United States)

    2006-06-01

    divert airfield for military aircraft training in the NTTR and support the flying operations of the 57th Wing, other Air Force units, Navy, Marine ...includes desert iguana (Dipsosaurus dorsalis), zebra-tailed lizard (Callosaurus draconoides), side-blotched lizard, horned lizards (Phrynosoma spp...be stored or used in work areas. Marine mammal —any mammal that is morphologically adapted to the marine environment, or primarily inhabits the

  8. Spanwise morphing trailing edge on a finite wing

    Science.gov (United States)

    Pankonien, Alexander M.; Inman, Daniel J.

    2015-04-01

    Unmanned Aerial Vehicles are prime targets for morphing implementation as they must adapt to large changes in flight conditions associated with locally varying wind or large changes in mass associated with payload delivery. The Spanwise Morphing Trailing Edge concept locally varies the trailing edge camber of a wing or control surface, functioning as a modular replacement for conventional ailerons without altering the spar box. Utilizing alternating active sections of Macro Fiber Composites (MFCs) driving internal compliant mechanisms and inactive sections of elastomeric honeycombs, the SMTE concept eliminates geometric discontinuities associated with shape change, increasing aerodynamic performance. Previous work investigated a representative section of the SMTE concept and investigated the effect of various skin designs on actuation authority. The current work experimentally evaluates the aerodynamic gains for the SMTE concept for a representative finite wing as compared with a conventional, articulated wing. The comparative performance for both wings is evaluated by measuring the drag penalty associated with achieving a design lift coefficient from an off-design angle of attack. To reduce experimental complexity, optimal control configurations are predicted with lifting line theory and experimentally measured control derivatives. Evaluated over a range of off-design flight conditions, this metric captures the comparative capability of both concepts to adapt or "morph" to changes in flight conditions. Even with this simplistic model, the SMTE concept is shown to reduce the drag penalty due to adaptation up to 20% at off-design conditions, justifying the increase in mass and complexity and motivating concepts capable of larger displacement ranges, higher fidelity modelling, and condition-sensing control.

  9. Features wear nodes mechanization wing aircraft operating under dynamic loads

    Directory of Open Access Journals (Sweden)

    А.М. Хімко

    2009-03-01

    Full Text Available  The conducted researches of titanic alloy ВТ-22 at dynamic loading with cycled sliding and dynamic loading in conditions of rolling with slipping. It is established that roller jamming in the carriage increases wear of rod of mechanization of a wing to twenty times. The optimum covering for strengthening wearied sites and restoration of working surfaces of wing’s mechanization rod is defined.

  10. Contribution of a winged phlebotomy device design to blood splatter.

    Science.gov (United States)

    Haiduven, Donna J; McGuire-Wolfe, Christine; Applegarth, Shawn P

    2012-11-01

    Despite a proliferation of phlebotomy devices with engineered sharps injury protection (ESIP), the impact of various winged device designs on blood splatter occurring during venipuncture procedures has not been explored. To evaluate the potential for blood splatter of 6 designs of winged phlebotomy devices. A laboratory-based device evaluation without human subjects, using a simulated patient venous system. We evaluated 18 winged phlebotomy devices of 6 device designs by Terumo, BD Vacutainer (2 designs), Greiner, Smith Medical, and Kendall (designated A-F, respectively). Scientific filters were positioned around the devices and weighed before and after venipuncture was performed. Visible blood on filters, exam gloves, and devices and measurable blood splatter were the primary units of analysis. The percentages of devices and gloves with visible blood on them and filters with measurable blood splatter ranged from 0% to 20%. There was a statistically significant association between device design and visible blood on devices ([Formula: see text]) and between device design and filters with measurable blood splatter ([Formula: see text]), but not between device design and visible blood on gloves. A wide range of associations were demonstrated between device design and visible blood on gloves or devices and incidence of blood splatter. The results of this evaluation suggest that winged phlebotomy devices with ESIP may produce blood splatter during venipuncture. Reinforcing the importance of eye protection and developing a methodology to assess ocular exposure to blood splatter are major implications for healthcare personnel who use these devices. Future studies should focus on evaluating different designs of intravascular devices (intravenous catheters, other phlebotomy devices) for blood splatter.

  11. Ubiquitous Supercritical Wing Design Cuts Billions in Fuel Costs

    Science.gov (United States)

    2015-01-01

    A Langley Research Center engineer’s work in the 1960s and ’70s to develop a wing with better performance near the speed of sound resulted in a significant increase in subsonic efficiency. The design was shared with industry. Today, Renton, Washington-based Boeing Commercial Airplanes, as well as most other plane manufacturers, apply it to all their aircraft, saving the airline industry billions of dollars in fuel every year.

  12. Wing motion measurement and aerodynamics of hovering true hoverflies.

    Science.gov (United States)

    Mou, Xiao Lei; Liu, Yan Peng; Sun, Mao

    2011-09-01

    Most hovering insects flap their wings in a horizontal plane (body having a large angle from the horizontal), called `normal hovering'. But some of the best hoverers, e.g. true hoverflies, hover with an inclined stroke plane (body being approximately horizontal). In the present paper, wing and body kinematics of four freely hovering true hoverflies were measured using three-dimensional high-speed video. The measured wing kinematics was used in a Navier-Stokes solver to compute the aerodynamic forces of the insects. The stroke amplitude of the hoverflies was relatively small, ranging from 65 to 85 deg, compared with that of normal hovering. The angle of attack in the downstroke (∼50 deg) was much larger that in the upstroke (∼20 deg), unlike normal-hovering insects, whose downstroke and upstroke angles of attack are not very different. The major part of the weight-supporting force (approximately 86%) was produced in the downstroke and it was contributed by both the lift and the drag of the wing, unlike the normal-hovering case in which the weight-supporting force is approximately equally contributed by the two half-strokes and the lift principle is mainly used to produce the force. The mass-specific power was 38.59-46.3 and 27.5-35.4 W kg(-1) in the cases of 0 and 100% elastic energy storage, respectively. Comparisons with previously published results of a normal-hovering true hoverfly and with results obtained by artificially making the insects' stroke planes horizontal show that for the true hoverflies, the power requirement for inclined stroke-plane hover is only a little (<10%) larger than that of normal hovering.

  13. High Performance Piezoelectric Actuators and Wings for Nano Air Vehicles

    Science.gov (United States)

    2012-08-26

    process to monolithically fabricate flying robotic insects at the pico air ve- hicle ( PAV ) scale from SUEX dry film, an epoxy based negative photoresist...cost. It simul- taneously defines the PAV airframe, compliant flapping mechanism, and artificial insect wing using photolithography. Using this process...81 4.4.3 Simulated Average Lift Versus Frequency of the Redesigned LionFly 82 5.1.1 Potential Fabrication Process for PAV Flapping

  14. Finite Element Analysis of a Highly Flexible Flapping Wing

    Science.gov (United States)

    2013-03-01

    arm and could account for some difference in tip deflection as well as the wing kinematics. The low stiffness of the membrane made the model...S. Zhu, Z. Su and H. Zhang, "Added Mass Effect and an Extended Unsteady Blade  Element Mode of Insect Hovering," Journal of  Bionic  Engineering, vol

  15. Prediction of vortex breakdown on a delta wing

    Science.gov (United States)

    Agrawal, S.; Robinson, B. A.; Barnett, R. M.

    1992-01-01

    Recent studies of leading-edge vortex flows with computational fluid dynamics codes using Euler or Navier-Stokes formulations have shown fair agreement with experimental data. These studies have concentrated on simulating the flowfields associated with a sharp-edged flat plate 70 deg delta wing at angles of attack where vortex breakdown or burst is observed over the wing. There are, however, a number of discrepancies between the experimental data and the computed flowfields. The location of vortex breakdown in the computational solutions is seen to differ from the experimental data and to vary with changes in the computational grid and freestream Mach number. There also remain issues as to the validity of steady-state computations for cases which contain regions of unsteady flow, such as in the post-breakdown regions. As a partial response to these questions, a number of laminar Navier-Stokes solutions were examined for the 70 deg delta wing. The computed solutions are compared with an experimental database obtained at low subsonic speeds. The convergence of forces, moments and vortex breakdown locations are also analyzed to determine if the computed flowfields actually reach steady-state conditions.

  16. Lifting Wing in Constructing Tall Buildings —Aerodynamic Testing

    Directory of Open Access Journals (Sweden)

    Ian Skelton

    2014-05-01

    Full Text Available This paper builds on previous research by the authors which determined the global state-of-the-art of constructing tall buildings by surveying the most active specialist tall building professionals around the globe. That research identified the effect of wind on tower cranes as a highly ranked, common critical issue in tall building construction. The research reported here presents a design for a “Lifting Wing,” a uniquely designed shroud which potentially allows the lifting of building materials by a tower crane in higher and more unstable wind conditions, thereby reducing delay on the programmed critical path of a tall building. Wind tunnel tests were undertaken to compare the aerodynamic performance of a scale model of a typical “brick-shaped” construction load (replicating a load profile most commonly lifted via a tower crane against the aerodynamic performance of the scale model of the Lifting Wing in a range of wind conditions. The data indicate that the Lifting Wing improves the aerodynamic performance by a factor of up to 50%.

  17. Vorticity Transport on a Flexible Wing in Stall Flutter

    Science.gov (United States)

    Akkala, James; Buchholz, James; Farnsworth, John; McLaughlin, Thomas

    2014-11-01

    The circulation budget within dynamic stall vortices was investigated on a flexible NACA 0018 wing model of aspect ratio 6 undergoing stall flutter. The wing had an initial angle of attack of 6 degrees, Reynolds number of 1 . 5 ×105 and large-amplitude, primarily torsional, limit cycle oscillations were observed at a reduced frequency of k = πfc / U = 0 . 1 . Phase-locked stereo PIV measurements were obtained at multiple chordwise planes around the 62.5% and 75% spanwise locations to characterize the flow field within thin volumetric regions over the suction surface. Transient surface pressure measurements were used to estimate boundary vorticity flux. Recent analyses on plunging and rotating wings indicates that the magnitude of the pressure-gradient-driven boundary flux of secondary vorticity is a significant fraction of the magnitude of the convective flux from the separated leading-edge shear layer, suggesting that the secondary vorticity plays a significant role in regulating the strength of the primary vortex. This phenomenon is examined in the present case, and the physical mechanisms governing the growth and evolution of the dynamic stall vortices are explored. This work was supported by the Air Force Office of Scientific Research through the Flow Interactions and Control Program monitored by Dr. Douglas Smith, and through the 2014 AFOSR/ASEE Summer Faculty Fellowship Program (JA and JB).

  18. The analysis of the flying wing in morphing concept

    Directory of Open Access Journals (Sweden)

    Ionică CÎRCIU

    2013-06-01

    Full Text Available The combination between the flying wing morphing concept and the use of modern command and control system offers exponential advantages having a leverage effect in the economy and research. The flying wing architecture has the advantage of low cost against efficiency, the morphing of this concept defining the new characteristic frontiers and aerodynamic performances which derive immediately. On designing an unmanned aerial vehicle for a various range of missions, its lifting surface needs to display optimal geometrical features, so that the UAV may maintain the induced drag and the moment coefficient at reasonable levels. The command and control of the lifting surfaces in morphing concept offer characteristics and in-flight performances at a superior level. The limits of the system depend on the reliability of the execution elements and the grade of accuracy for the control laws which are implemented in the calculation module. The paper aims at presenting an analysis regarding the robotic air systems of flying wing type through the aerodynamic analysis and with the help of specific software instruments. The performances and flight qualities depend directly on the geometry of the lifting surface of the aerial vehicle.

  19. Wing kinematics and flexibility for optimal manoeuvring and escape

    Science.gov (United States)

    Wong, Jaime Gustav

    Understanding how animals control the dynamic stall vortices in their wake is critical to developing micro-aerial vehicles and autonomous underwater vehicles, not to mention wind turbines, delta wings, and rotor craft that undergo similar dynamic stall processes. Applying this knowledge to biomimetic engineering problems requires progress in three areas: (i) understanding the flow physics of natural swimmers and flyers; (ii) developing flow measurement techniques to resolve this physics; and (iii) deriving low-cost models suitable for studying the vast parameter space observed in nature. This body of work, which consists of five research chapters, focuses on the leading-edge vortex (LEV) that forms on profiles undergoing rapid manoeuvres, delta wings, and similar devices. Lagrangian particle tracking is used throughout this thesis to track the mass and circulation transport in the LEV on manoeuvring profiles. The growth and development of the LEV is studied in relation to: flapping and plunging profile kinematics; spanwise flow from profile sweep and spanwise profile bending; and varying the angle-of-attack gradient along the profile span. Finally, scaling relationships derived from the observations above are used to develop a low-cost model for LEV growth, that is validated on a flat-plate delta wing. Together these results contribute to each of the three topics identified above, as a step towards developing robust, agile biomimetic swimmers and flyers.

  20. Analysis of a Hybrid Wing Body Center Section Test Article

    Science.gov (United States)

    Wu, Hsi-Yung T.; Shaw, Peter; Przekop, Adam

    2013-01-01

    The hybrid wing body center section test article is an all-composite structure made of crown, floor, keel, bulkhead, and rib panels utilizing the Pultruded Rod Stitched Efficient Unitized Structure (PRSEUS) design concept. The primary goal of this test article is to prove that PRSEUS components are capable of carrying combined loads that are representative of a hybrid wing body pressure cabin design regime. This paper summarizes the analytical approach, analysis results, and failure predictions of the test article. A global finite element model of composite panels, metallic fittings, mechanical fasteners, and the Combined Loads Test System (COLTS) test fixture was used to conduct linear structural strength and stability analyses to validate the specimen under the most critical combination of bending and pressure loading conditions found in the hybrid wing body pressure cabin. Local detail analyses were also performed at locations with high stress concentrations, at Tee-cap noodle interfaces with surrounding laminates, and at fastener locations with high bearing/bypass loads. Failure predictions for different composite and metallic failure modes were made, and nonlinear analyses were also performed to study the structural response of the test article under combined bending and pressure loading. This large-scale specimen test will be conducted at the COLTS facility at the NASA Langley Research Center.

  1. Monostatic radar cross section of flying wing delta planforms

    Directory of Open Access Journals (Sweden)

    Sevoor Meenakshisundaram Vaitheeswaran

    2017-04-01

    Full Text Available The design of the flying wing and its variants shapes continues to have a profound influence in the design of the current and future use of military aircraft. There is very little in the open literature available to the understanding and by way of comparison of the radar cross section of the different wing planforms, for obvious reasons of security and sensitivity. This paper aims to provide an insight about the radar cross section of the various flying wing planforms that would aid the need and amount of radar cross section suppression to escape detection from surveillance radars. Towards this, the shooting and bouncing ray method is used for analysis. In this, the geometric optics theory is first used for launching and tracing the electromagnetic rays to calculate the electromagnetic field values as the waves bounce around the target. The physical optics theory is next used to calculate the final scattered electric field using the far field integration along the observation direction. For the purpose of comparison, all the planform shapes are assumed to be having the same area, and only the aspect ratio and taper ratio are varied to feature representative airplanes.

  2. Aerodynamic Design of Wing based on Humpback Whale Flipper

    Science.gov (United States)

    Akram, Saif; Baig, Faisal

    2013-11-01

    The tubercles provide a bio-inspired design that has commercial viability for wing-like structures. Wind tunnel tests at low speeds of model humpback flippers with leading-edge tubercles have demonstrated improvements tubercles make, such as a staggering 32% reduction in drag, 8% improvement in lift, and a 40% increase in angle of attack over smooth flippers before stalling. The tubercles on the leading edge act as a passive-flow control device that improves the performance and maneuverability of the flipper. Possible fluid-dynamic mechanisms for improved performance include delay of stall through generation of a vortex and modification of the boundary layer, and increase in effective span by reduction of both spanwise flow and strength of the tip vortex. In the present work, numerical investigation of a 3D wing with scalloped leading edge inspired by the humpback whale flipper is carried out at high subsonic speeds with variation in angle of attack from 0 to 25 degrees. The effect of using different turbulence models is also investigated in order to attain a better understanding of mechanism(s) responsible for improved aerodynamic performance. This new understanding of humpback whale flipper aerodynamics has strong implications for wing design.

  3. On the leading edge vortex of thin wings

    Science.gov (United States)

    Arredondo, Abel; Viola, Ignazio Maria

    2016-11-01

    On thin wings, the sharp leading edge triggers laminar separation followed by reattachment, forming a Leading Edge Vortex (LEV). This flow feature is of paramount importance because, if periodically shed, it leads to large amplitude load fluctuations, while if stably attached to the wing, it can provide lift augmentation. We found that on asymmetric-spinnaker-type yacht sails, the LEV can be stable despite the relatively low sweep (30°). This finding, which was recently predicted numerically by Viola et al., has been confirmed through current flume tests on a 1:115th model scale sail. Forces were measured and Particle Image Velocimetry was performed on four horizontal sail sections at a Reynolds number of 1.7x104. Vortex detection revealed that the LEV becomes progressively larger and more stable towards the highest sections, where its axis has a smaller angle with respect to the freestream velocity. Mapping the sail section on a rotating cylinder through a Joukowski transformation, we quantified the lift augmentation provided by the LEV on each sail section. These results open up new sail design strategies based on the manipulation of the LEV and can be applicable to the wings of unmanned aerial vehicles and underwater vehicles. Project funded by Conacyt.

  4. Cap-n-Collar Promotes Tissue Regeneration by Regulating ROS and JNK Signaling in the Drosophila melanogaster Wing Imaginal Disc.

    Science.gov (United States)

    Brock, Amanda R; Seto, Mabel; Smith-Bolton, Rachel K

    2017-07-01

    Regeneration is a complex process that requires an organism to recognize and repair tissue damage, as well as grow and pattern new tissue. Here, we describe a genetic screen to identify novel regulators of regeneration. We ablated the Drosophila melanogaster larval wing primordium by inducing apoptosis in a spatially and temporally controlled manner and allowed the tissue to regenerate and repattern. To identify genes that regulate regeneration, we carried out a dominant-modifier screen by assessing the amount and quality of regeneration in adult wings heterozygous for isogenic deficiencies. We have identified 31 regions on the right arm of the third chromosome that modify the regenerative response. Interestingly, we observed several distinct phenotypes: mutants that regenerated poorly, mutants that regenerated faster or better than wild-type, and mutants that regenerated imperfectly and had patterning defects. We mapped one deficiency region to cap-n-collar ( cnc ), the Drosophila Nrf2 ortholog, which is required for regeneration. Cnc regulates reactive oxygen species levels in the regenerating epithelium, and affects c-Jun N-terminal protein kinase (JNK) signaling, growth, debris localization, and pupariation timing. Here, we present the results of our screen and propose a model wherein Cnc regulates regeneration by maintaining an optimal level of reactive oxygen species to promote JNK signaling. Copyright © 2017 by the Genetics Society of America.

  5. Expanding signaling-molecule wavefront model of cell polarization in the Drosophila wing primordium.

    Science.gov (United States)

    Wortman, Juliana C; Nahmad, Marcos; Zhang, Peng Cheng; Lander, Arthur D; Yu, Clare C

    2017-07-01

    In developing tissues, cell polarization and proliferation are regulated by morphogens and signaling pathways. Cells throughout the Drosophila wing primordium typically show subcellular localization of the unconventional myosin Dachs on the distal side of cells (nearest the center of the disc). Dachs localization depends on the spatial distribution of bonds between the protocadherins Fat (Ft) and Dachsous (Ds), which form heterodimers between adjacent cells; and the Golgi kinase Four-jointed (Fj), which affects the binding affinities of Ft and Ds. The Fj concentration forms a linear gradient while the Ds concentration is roughly uniform throughout most of the wing pouch with a steep transition region that propagates from the center to the edge of the pouch during the third larval instar. Although the Fj gradient is an important cue for polarization, it is unclear how the polarization is affected by cell division and the expanding Ds transition region, both of which can alter the distribution of Ft-Ds heterodimers around the cell periphery. We have developed a computational model to address these questions. In our model, the binding affinity of Ft and Ds depends on phosphorylation by Fj. We assume that the asymmetry of the Ft-Ds bond distribution around the cell periphery defines the polarization, with greater asymmetry promoting cell proliferation. Our model predicts that this asymmetry is greatest in the radially-expanding transition region that leaves polarized cells in its wake. These cells naturally retain their bond distribution asymmetry after division by rapidly replenishing Ft-Ds bonds at new cell-cell interfaces. Thus we predict that the distal localization of Dachs in cells throughout the pouch requires the movement of the Ds transition region and the simple presence, rather than any specific spatial pattern, of Fj.

  6. Wind, waves, and wing loading: Morphological specialization may limit range expansion of endangered albatrosses

    Science.gov (United States)

    Suryan, R.M.; Anderson, D.J.; Shaffer, S.A.; Roby, D.D.; Tremblay, Y.; Costa, D.P.; Sievert, P.R.; Sato, F.; Ozaki, K.; Balogh, G.R.; Nakamura, N.

    2008-01-01

    Among the varied adaptations for avian flight, the morphological traits allowing large-bodied albatrosses to capitalize on wind and wave energy for efficient long-distance flight are unparalleled. Consequently, the biogeographic distribution of most albatrosses is limited to the windiest oceanic regions on earth; however, exceptions exist. Species breeding in the North and Central Pacific Ocean (Phoebastria spp.) inhabit regions of lower wind speed and wave height than southern hemisphere genera, and have large intrageneric variation in body size and aerodynamic performance. Here, we test the hypothesis that regional wind and wave regimes explain observed differences in Phoebastria albatross morphology and we compare their aerodynamic performance to representatives from the other three genera of this globally distributed avian family. In the North and Central Pacific, two species (short-tailed P. albatrus and waved P. irrorata) are markedly larger, yet have the smallest breeding ranges near highly productive coastal upwelling systems. Short-tailed albatrosses, however, have 60% higher wing loading (weight per area of lift) compared to waved albatrosses. Indeed, calculated aerodynamic performance of waved albatrosses, the only tropical albatross species, is more similar to those of their smaller congeners (black-footed P. nigripes and Laysan P. immutabilis), which have relatively low wing loading and much larger foraging ranges that include central oceanic gyres of relatively low productivity. Globally, the aerodynamic performance of short-tailed and waved albatrosses are most anomalous for their body sizes, yet consistent with wind regimes within their breeding season foraging ranges. Our results are the first to integrate global wind and wave patterns with albatross aerodynamics, thereby identifying morphological specialization that may explain limited breeding ranges of two endangered albatross species. These results are further relevant to understanding past and

  7. Wind, waves, and wing loading: morphological specialization may limit range expansion of endangered albatrosses.

    Directory of Open Access Journals (Sweden)

    Robert M Suryan

    Full Text Available Among the varied adaptations for avian flight, the morphological traits allowing large-bodied albatrosses to capitalize on wind and wave energy for efficient long-distance flight are unparalleled. Consequently, the biogeographic distribution of most albatrosses is limited to the windiest oceanic regions on earth; however, exceptions exist. Species breeding in the North and Central Pacific Ocean (Phoebastria spp. inhabit regions of lower wind speed and wave height than southern hemisphere genera, and have large intrageneric variation in body size and aerodynamic performance. Here, we test the hypothesis that regional wind and wave regimes explain observed differences in Phoebastria albatross morphology and we compare their aerodynamic performance to representatives from the other three genera of this globally distributed avian family. In the North and Central Pacific, two species (short-tailed P. albatrus and waved P. irrorata are markedly larger, yet have the smallest breeding ranges near highly productive coastal upwelling systems. Short-tailed albatrosses, however, have 60% higher wing loading (weight per area of lift compared to waved albatrosses. Indeed, calculated aerodynamic performance of waved albatrosses, the only tropical albatross species, is more similar to those of their smaller congeners (black-footed P. nigripes and Laysan P. immutabilis, which have relatively low wing loading and much larger foraging ranges that include central oceanic gyres of relatively low productivity. Globally, the aerodynamic performance of short-tailed and waved albatrosses are most anomalous for their body sizes, yet consistent with wind regimes within their breeding season foraging ranges. Our results are the first to integrate global wind and wave patterns with albatross aerodynamics, thereby identifying morphological specialization that may explain limited breeding ranges of two endangered albatross species. These results are further relevant to

  8. Variation in wing characteristics of monarch butterflies during migration: Earlier migrants have redder and more elongated wings

    Directory of Open Access Journals (Sweden)

    Satterfield Dara A.

    2014-01-01

    Full Text Available The migration of monarch butterflies (Danaus plexippus in North America has a number of parallels with long-distance bird migration, including the fact that migratory populations of monarchs have larger and more elongated forewings than residents. These characteristics likely serve to optimize flight performance in monarchs, as they also do with birds. A question that has rarely been addressed thus far in birds or monarchs is if and how wing characteristics vary within a migration season. Individuals with superior flight performance should migrate quickly, and/or with minimal stopovers, and these individuals should be at the forefront of the migratory cohort. Conversely, individuals with poor flight performance and/or low endurance would be more likely to fall behind, and these would comprise the latest migrants. Here we examined how the wing morphology of migrating monarchs varies to determine if wing characteristics of early migrants differ from late migrants. We measured forewing area, elongation (length/width, and redness, which has been shown to predict flight endurance in monarchs. Based on a collection of 75 monarchs made one entire season (fall 2010, results showed that the earliest migrants (n = 20 in this cohort had significantly redder and more elongated forewings than the latest migrants (n = 17. There was also a non-significant tendency for early migrants to have larger forewing areas. These results suggest that the pace of migration in monarchs is at least partly dependent on the properties of their wings. Moreover, these data also raise a number of questions about the ultimate fate of monarchs that fall behind

  9. AEROELASTIC FLUTTER ANALYSIS OF SUPERSONIC WING WITH MULTIPLE EXTERNAL STORES

    Directory of Open Access Journals (Sweden)

    Nur Azam

    2014-12-01

    Full Text Available ABSTRACT: Flutter may be considered to be one of the most dangerous aeroelastic failure phenomenon. The flutter characteristic differs for each aircraft type, and depends on the wing geometry as well as its operational region of subsonic, transonic or supersonic speeds. Prior to performing a flight flutter test, extensive numerical simulations and Ground Vibration Test should be conducted where the structural finite element modes and the experimentation results should be matched, otherwise the numerical simulation model must be rejected. In this paper, the analysis of simulation of a supersonic wing equipped with external missiles loaded on the wing is presented. The structural mode shapes at each generated frequency are also visually presented. The analysis is carried out using MSC Nastran FEM software. The wing flutter with the external stores was simulated at different altitudes. The result shows that the flutter velocity is sensitive to the flight altitude. For this reason, the flutter analysis is conducted also for a negative altitude. The negative altitude is obtained by considering the constant equivalent speed-Mach number rule at the flutter speed boundary as a requirement in standard regulation of transport aircraft. ABSTRAK: Salah satu fenomena kegagalan aeroelastik yang paling membahayakan adalah kipasan (flutter. Ciri-ciri kegagalan kipasan (flutter adalah berbeza untuk setiap jenis pesawat bergantung pada geometri sayap dan regim operasi sama ada subsonik, transonik atau supersonik. Sebelum melakukan ujian penerbangan kipasan , simulasi berangka luas dan ujian getaran peringkat bawahan (darat perlu dijalankan di mana struktur mod unsur terhingga dan keputusan eksperimen harus dipadankan, sebaliknya model simulasi berangka boleh ditolak. Dalam kertas kerja ini, simulasi sayap supersonik dilengkapi dengan beban luaran peluru berpandu di sayap telah dianalisis di daerah supersonik tinggi. Bentuk mod struktur pada setiap mod frekuensi

  10. Development of Delta Wing Aerodynamics Research in Universiti Teknologi Malaysia Low Speed Wind Tunnel

    Directory of Open Access Journals (Sweden)

    Shabudin Mat

    2014-07-01

    Full Text Available This paper presents wind tunnel experiment on two delta wing configurations which are differentiated by their leading edge profiles: sharp and round-edged wings. The experiments were performed as a part of the delta wing aerodynamics research development in Universiti Teknologi Malaysia, low speed tunnel (UTM-LST. Steady load balance and flow visualization tests were conducted at Reynolds numbers of 0.5, 1, and 1.5 × 106, respectively. The flow measurement at low Reynolds number was also performed at as low as speed of 5 m/s. During the experiments, laser with smoke flow visualizations test was performed on both wings. The study has identified interesting features of the interrelationship between the conventional leading edge primary vortex and the occurrence and development of the vortex breakdown above the delta wings. The results conclude the vortex characteristics are largely dependent on the Reynolds number, angle of attack, and leading-edge radii of the wing.

  11. Experimental Wing Damage Affects Foraging Effort and Foraging Distance in Honeybees Apis mellifera

    Directory of Open Access Journals (Sweden)

    Andrew D. Higginson

    2011-01-01

    Full Text Available Bees acquire wing damage as they age, and loss of wing area affects longevity and behaviour. This may influence colony performance via effects on worker behaviour. The effects of experimental wing damage were studied in worker honeybees in observation hives by recording survivorship, how often and for how long bees foraged, and by decoding waggle dances. Mortality rate increased with both age and wing damage. Damaged bees carried out shorter and/or less frequent foraging trips, foraged closer to the hive, and reported the profitability of flower patches to be lower than did controls. These results suggest that wing damage caused a reduction in foraging ability, and that damaged bees adjusted their foraging behaviour accordingly. Furthermore, the results suggest that wing damage affects the profitability of nectar sources. These results have implications for the colony dynamics and foraging efficiency in honeybees.

  12. Dynamic Model and Analysis of Asymmetric Telescopic Wing for Morphing Aircraft

    Directory of Open Access Journals (Sweden)

    Chen Lili

    2016-01-01

    Full Text Available Morphing aircraft has been the research hot topics of new concept aircrafts in aerospace engineering. Telescopic wing is an important morphing technology for morphing aircraft. This paper describes the dynamic equations and kinematic equations based on theorem of momentum and theorem of moment of momentum, which are available for all morphing aircrafts. Meanwhile,as simplified , dynamic equations for rectangular telescopic wing are presented. In order to avoid the complexity using aileron to generate rolling moment , an new idea that asymmetry of wings can generate roll moment is introduced. Finally, roll performance comparison of asymmetric wing and aileron deflection shows that asymmetric telescopic wing can provide the required roll control moment as aileron, and in some cases, telescopic wing has the superior roll performance.

  13. Aircraft wing weight build-up methodology with modification for materials and construction techniques

    Science.gov (United States)

    York, P.; Labell, R. W.

    1980-01-01

    An aircraft wing weight estimating method based on a component buildup technique is described. A simplified analytically derived beam model, modified by a regression analysis, is used to estimate the wing box weight, utilizing a data base of 50 actual airplane wing weights. Factors representing materials and methods of construction were derived and incorporated into the basic wing box equations. Weight penalties to the wing box for fuel, engines, landing gear, stores and fold or pivot are also included. Methods for estimating the weight of additional items (secondary structure, control surfaces) have the option of using details available at the design stage (i.e., wing box area, flap area) or default values based on actual aircraft from the data base.

  14. Aerodynamic Optimization of an Over-the-Wing-Nacelle-Mount Configuration

    Directory of Open Access Journals (Sweden)

    Daisuke Sasaki

    2011-01-01

    Full Text Available An over-the-wing-nacelle-mount airplane configuration is known to prevent the noise propagation from jet engines toward ground. However, the configuration is assumed to have low aerodynamic efficiency due to the aerodynamic interference effect between a wing and a nacelle. In this paper, aerodynamic design optimization is conducted to improve aerodynamic efficiency to be equivalent to conventional under-the-wing-nacelle-mount configuration. The nacelle and wing geometry are modified to achieve high lift-to-drag ratio, and the optimal geometry is compared with a conventional configuration. Pylon shape is also modified to reduce aerodynamic interference effect. The final wing-fuselage-nacelle model is compared with the DLR F6 model to discuss the potential of Over-the-Wing-Nacelle-Mount geometry for an environmental-friendly future aircraft.

  15. Wake Characteristics of a Flapping Wing Optimized for both Aerial and Aquatic Flight

    Science.gov (United States)

    Izraelevitz, Jacob; Kotidis, Miranda; Triantafyllou, Michael

    2017-11-01

    Multiple aquatic bird species (including murres, puffins, and other auks) employ a single actuator to propel themselves in two different fluid media: both flying and swimming using primarily their flapping wings. This impressive design compromise could be adopted by engineered implementations of dual aerial/aquatic robotic platforms, as it offers an existence proof for favorable flow physics. We discuss one realization of a 3D flapping wing actuation system for use in both air and water. The wing oscillates by the root and employs an active in-line motion degree-of-freedom. An experiment-coupled optimization routine generates the wing trajectories, controlling the unsteady forces throughout each flapping cycle. We elucidate the wakes of these wing trajectories using dye visualization, correlating the wake vortex structures with simultaneous force measurements. After optimization, the wing generates the large force envelope necessary for propulsion in both fluid media, and furthermore, demonstrate improved control over the unsteady wake.

  16. Design and flight performance of hybrid underwater glider with controllable wings

    Directory of Open Access Journals (Sweden)

    Yanhui Wang

    2017-05-01

    Full Text Available Hybrid underwater glider combines motion modes of traditional autonomous underwater glider and those of autonomous underwater vehicles. Different motion modes need different flight performance, including flight efficiency, static stability, and maneuverability. Conventional hybrid underwater glider with fixed wings can’t achieve optimal flight performance in one flight mission demanding various motion modes. In this article, controllable wings for hybrid underwater glider Petrel II are designed. Angle of attack, sweep angle, and aspect ratio of controllable wings can be changed to adapt to different motion modes. Kinematics and dynamics models of Petrel II are established based on multibody theory. Motion simulations of Petrel II with different wing configurations are conducted in three motion modes, including glide motion, spiral motion, and horizontal turning motion. The simulation results show the impact of wing parameters on flight performance. Field trials demonstrate that the controllable wings can improve the flight performance.

  17. Pegasus Rocket Wing and PHYSX Glove Undergoes Stress Loads Testing

    Science.gov (United States)

    1997-01-01

    The Pegasus Hypersonic Experiment (PHYSX) Project's Pegasus rocket wing with attached PHYSX glove rests after load-tests at Scaled Composites, Inc., in Mojave, California, in January 1997. Technicians slowly filled water bags beneath the wing, to create the pressure, or 'wing-loading,' required to determine whether the wing could withstand its design limit for stress. The wing sits in a wooden triangular frame which serves as the test-rig, mounted to the floor atop the waterbags. Pegasus is an air-launched space booster produced by Orbital Sciences Corporation and Hercules Aerospace Company (initially; later, Alliant Tech Systems) to provide small satellite users with a cost-effective, flexible, and reliable method for placing payloads into low earth orbit. Pegasus has been used to launch a number of satellites and the PHYSX experiment. That experiment consisted of a smooth glove installed on the first-stage delta wing of the Pegasus. The glove was used to gather data at speeds of up to Mach 8 and at altitudes approaching 200,000 feet. The flight took place on October 22, 1998. The PHYSX experiment focused on determining where boundary-layer transition occurs on the glove and on identifying the flow mechanism causing transition over the glove. Data from this flight-research effort included temperature, heat transfer, pressure measurements, airflow, and trajectory reconstruction. Hypersonic flight-research programs are an approach to validate design methods for hypersonic vehicles (those that fly more than five times the speed of sound, or Mach 5). Dryden Flight Research Center, Edwards, California, provided overall management of the glove experiment, glove design, and buildup. Dryden also was responsible for conducting the flight tests. Langley Research Center, Hampton, Virginia, was responsible for the design of the aerodynamic glove as well as development of sensor and instrumentation systems for the glove. Other participating NASA centers included Ames Research

  18. Effects of the essential oil constituent thymol and other neuroactive chemicals on flight motor activity and wing beat frequency in the blowfly Phaenicia sericata.

    Science.gov (United States)

    Waliwitiya, Ranil; Belton, Peter; Nicholson, Russell A; Lowenberger, Carl A

    2010-03-01

    The effects were evaluated of the plant terpenoid thymol and eight other neuroactive compounds on flight muscle impulses (FMIs) and wing beat frequency (WBF) of tethered blowflies (Phaenicia sericata Meig.). The electrical activity of the dorsolongitudinal flight muscles was closely linked to the WBF of control insects. Topically applied thymol inhibited WBF within 15-30 min and reduced FMI frequency. Octopamine and chlordimeform caused a similar, early-onset bursting pattern that decreased in amplitude with time. Desmethylchlordimeform blocked wing beating within 60 min and generated a profile of continuous but lower-frequency FMIs. Fipronil suppressed wing beating and induced a pattern of continuous, variable-frequency spiking that diminished gradually over 6 h. Cypermethrin- and rotenone-treated flies had initial strong FMIs that declined with time. In flies injected with GABA, the FMIs were generally unidirectional and frequency was reduced, as was seen with thymol. Thymol readily penetrates the cuticle and interferes with flight muscle and central nervous function in the blowfly. The similarity of the action of thymol and GABA suggests that this terpenoid acts centrally in blowflies by mimicking or facilitating GABA action.

  19. A new genus of long-legged flies displaying remarkable wing directional asymmetry

    Science.gov (United States)

    Justin B. Runyon; Richard L. Hurley

    2004-01-01

    A previously unknown group of flies is described whose males exhibit directional asymmetry, in that the left wing is larger than, and of a different shape from, the right wing. To our knowledge, wing asymmetry of this degree has not previously been reported in an animal capable of flight. Such consistent asymmetry must result from a left­right axis during development...

  20. Aerodynamic performance of a hovering hawkmoth with flexible wings: a computational approach

    OpenAIRE

    Nakata, Toshiyuki; Liu, Hao

    2011-01-01

    Insect wings are deformable structures that change shape passively and dynamically owing to inertial and aerodynamic forces during flight. It is still unclear how the three-dimensional and passive change of wing kinematics owing to inherent wing flexibility contributes to unsteady aerodynamics and energetics in insect flapping flight. Here, we perform a systematic fluid-structure interaction based analysis on the aerodynamic performance of a hovering hawkmoth, Manduca, with an integrated comp...