WorldWideScience

Sample records for temperate heathland plants

  1. Plant nutrient mobilization in temperate heathland responds to elevated CO2, temperature and drought

    DEFF Research Database (Denmark)

    Andresen, Louise C.; Michelsen, Anders; Jonasson, Sven

    2010-01-01

    Temperate terrestrial ecosystems are currently exposed to increased atmospheric CO2 and progressive climatic changes with increased temperature and periodical drought. We here present results from a field experiment, where the effects of these three main climate change related factors...... decreased in response to drought. These complex changes in availability and release of nutrients from soil organic matter turnover and mineralization in response to elevated CO2 and climate change may influence the future plant carbon sequestration and species composition at temperate heathlands....... in Deschampsia soil, and microbial immobilization of N and P decreased in warmed Calluna soil. Warming tended to increase microbial N and P in Calluna but not in Deschampsia soil in fall, and more microbial C was accumulated under drought in Calluna soil. The effects of warming were often counteracted or erased...

  2. Seasonal variations and effects of nutrient applications on N and P and microbial biomass under two temperate heathland plants

    DEFF Research Database (Denmark)

    Nielsen, Pia Lund; Andresen, Louise Christoffersen; Michelsen, Anders

    2009-01-01

    . The microbial biomass on the other hand was positively related to soil water content in fertilized plots indicating that this was due to an indirect effect of enhanced nutrient availability. Microbial N and P pools were respectively 1000 and 100 times higher than the pool of inorganic N and P, and microbes...... this process. In this study the soil properties under two dominant heathland plants, the dwarf shrub Calluna vulgaris and the grass Deschampsia flexuosa, were investigated, with focus on nutrient content in the organic top soil and soil microbes during the main growing season and effects of nutrient amendments...... therefore may play an important role in regulating plant nutrient supply. Judged from responses of inorganic and microbial N and P concentrations to added N and P, N seemed to limit C. vulgaris and soil microbes below while P seemed to limit D. flexuosa and soil microbes below this species. There were lower...

  3. Can changes in soil biochemistry and plant stoichiometry explain loss of animal diversity of heathlands?

    NARCIS (Netherlands)

    Vogels, J.J.; Verberk, W.C.E.P.; Lamers, L.P.M.; Siepel, H.

    2017-01-01

    Increased atmospheric deposition rates of nitrogen (N) and sulphur (S) are known to affect soil biogeochemistry and cause a decline in plant biodiversity of heathlands. Concomitant declines of heathland invertebrates are mainly attributed to changes in vegetation composition and altered habitat

  4. Do competitive interactions in dry heathlands explain plant abundance patterns and species coexistence?

    DEFF Research Database (Denmark)

    Ransijn, Johannes; Damgaard, Christian; Schmidt, Inger K

    2015-01-01

    Plant community patterns in space and time may be explained by the interactions between competing plant species. The presented study investigates this in a nutrient and species poor ecosystem. The study presents a methodology for inferring competitive interactions from yearly vegetation inventories...... to predict the community dynamics of C. vulgaris and D. flexuosa. This was compared with the observed plant community structure at 198 Danish dry heathland sites. Interspecific competition will most likely lead to competitive exclusion of D. flexuosa at the observed temporal and spatial scale...... and uses this to assess the outcome of competitive interactions and to predict community patterns and dynamics in a Northwest-European dry heathland. Inferred competitive interactions from five consecutive years of measurements in permanent vegetation frames at a single dry heathland site were used...

  5. Effects of elevated atmospheric CO2, prolonged summer drought and temperature increase on N2O and CH4 fluxes in a temperate heathland

    DEFF Research Database (Denmark)

    Carter, Mette Sustmann; Ambus, Per; Albert, Kristian Rost

    2011-01-01

    In temperate regions, climate change is predicted to increase annual mean temperature and intensify the duration and frequency of summer droughts, which together with elevated atmospheric carbon dioxide (CO2) concentrations, may affect the exchange of nitrous oxide (N2O) and methane (CH4) between...... terrestrial ecosystems and the atmosphere. We report results from the CLIMAITE experiment, where the effects of these three climate change parameters were investigated solely and in all combinations in a temperate heathland. Field measurements of N2O and CH4 fluxes took place 1–2 years after the climate...... change manipulations were initiated. The soil was generally a net sink for atmospheric CH4. Elevated temperature (T) increased the CH4 uptake by on average 10 μg C m−2 h−1, corresponding to a rise in the uptake rate of about 20%. However, during winter elevated CO2 (CO2) reduced the CH4 uptake, which...

  6. Ecological modules and roles of species in heathland plant-insect flower visitor networks

    DEFF Research Database (Denmark)

    Dupont, Yoko; Olesen, Jens Mogens

    2009-01-01

    1.  Co-existing plants and flower-visiting animals often form complex interaction networks. A long-standing question in ecology and evolutionary biology is how to detect nonrandom subsets (compartments, blocks, modules) of strongly interacting species within such networks. Here we use a network...... analytical approach to (i) detect modularity in pollination networks, (ii) investigate species composition of modules, and (iii) assess the stability of modules across sites. 2.  Interactions between entomophilous plants and their flower-visitors were recorded throughout the flowering season at three...... heathland sites in Denmark, separated by ≥ 10 km. Among sites, plant communities were similar, but composition of flower-visiting insect faunas differed. Visitation frequencies of visitor species were recorded as a measure of insect abundance. 3.  Qualitative (presence-absence) interaction networks were...

  7. Changing heathlands in a changing climate

    DEFF Research Database (Denmark)

    Ransijn, Johannes

    Atmospheric CO2 concentrations and temperatures are rising and precipitation regimes are changing at global scale. How ecosystem will be affected by global climatic change is dependent on the responses of plants and plant communities. This thesis focuses on how climate change affects heathland...... plant communities. Many heathlands have shifted from dwarf shrub dominance to grass dominance and climatic change might affect the competitive balance between dwarf shrubs and grasses. We looked at heathland vegetation dynamics and heathland plant responses to climatic change at different spatial...... between C. vulgaris and D. flexuosa in the same climate change experiment and 5) a study where we compared the responses of shrubland plant communities to experimental warming and recurrent experimental droughts in seven climate change experiments across Europe. Heathland vegetation dynamics are slow...

  8. Life after fire: the impact of fire on species composition and diversity in coastal heathlands

    OpenAIRE

    Bargmann, Tessa

    2015-01-01

    Fire prone coastal heathlands are widespread, and range from naturally fire prone Mediterranean-type heathlands, to the anthropogenic heathlands of north-west Europe. Regardless of its source, fire is an important disturbance factor in these systems. This thesis investigates the effects of fire on the species diversity and composition of two heathland systems; one in south-eastern Tasmania and one in western Norway. I ask whether two important species groups in heathland, plants and carabids ...

  9. High Resilience in Heathland Plants to Changes in Temperature, Drought, and CO2 in Combination: Results from the CLIMAITE Experiment

    DEFF Research Database (Denmark)

    Kongstad, J.; Schmidt, Inger K.; Riis-Nielsen, Torben

    2012-01-01

    of in situexposure to elevated atmospheric CO2 concentration,increased temperature and prolonged droughtperiods on the plant biomass in a dry heathland(Brandbjerg, Denmark). Results after 3 yearsshowed that drought reduced the growth of thetwo dominant species Deschampsia flexuosa and Callunavulgaris. However, both...... the standingbiomass for either D. flexuosa or the ecosystem asmore litter was produced. Treatment combinationsshowed little interactions on the measuredparameters and in particular elevated CO2 did notcounterbalance the drought effect on plant growth,as we had anticipated. The plant community didnot show any......Climate change scenarios predict simultaneouslyincrease in temperature, altered precipitation patternsand elevated atmospheric CO2 concentration,which will affect key ecosystem processes and plantgrowth and species interactions. In a large-scaleexperiment, we investigated the effects...

  10. Survival of Dutch heathlands

    NARCIS (Netherlands)

    Diemont, W.H.

    1996-01-01

    OBJECTIVES OF THE THESIS

    Heathlands in The Netherlands are vanishing due to the invasion of trees. The transition from heathland to woodland in Dutch heathlands may either proceed directly or is preceded by the development of an intermediate grass heath. These changes are due to natural

  11. Short-term utilization of carbon by the soil microbial community under future climatic conditions in a temperate heathland

    DEFF Research Database (Denmark)

    Reinsch, Sabine; Michelsen, Anders; Sárossy, Zsuzsa

    2014-01-01

    An in-situ13C pulse-labeling experiment was carried out in a temperate heath/grassland to study the impacts of elevated CO2 concentration (510ppm), prolonged summer droughts (annual exclusion of 7.6±0.8%) and increased temperature (~1°C) on belowground carbon (C) utilization. Recently assimilated C...... (13C from the pulse-label) was traced into roots, soil and microbial biomass 1, 2 and 8 days after pulse-labeling. The importance of the microbial community in C utilization was investigated using 13C enrichment patterns in different microbial functional groups on the basis of phospholipid fatty acid...... (PLFA) biomarker profiles. Climate treatments did not affect microbial abundance in soil or rhizosphere fractions in terms of total PLFA-C concentration. Elevated CO2 significantly reduced the abundance of gram-negative bacteria (17:0cy), but did not affect the abundance of decomposers (fungi...

  12. Bacteria and Fungi Respond Differently to Multifactorial Climate Change in a Temperate Heathland, Traced with 13C-Glycine and FACE CO2

    Science.gov (United States)

    Andresen, Louise C.; Dungait, Jennifer A. J.; Bol, Roland; Selsted, Merete B.; Ambus, Per; Michelsen, Anders

    2014-01-01

    It is vital to understand responses of soil microorganisms to predicted climate changes, as these directly control soil carbon (C) dynamics. The rate of turnover of soil organic carbon is mediated by soil microorganisms whose activity may be affected by climate change. After one year of multifactorial climate change treatments, at an undisturbed temperate heathland, soil microbial community dynamics were investigated by injection of a very small concentration (5.12 µg C g−1 soil) of 13C-labeled glycine (13C2, 99 atom %) to soils in situ. Plots were treated with elevated temperature (+1°C, T), summer drought (D) and elevated atmospheric carbon dioxide (510 ppm [CO2]), as well as combined treatments (TD, TCO2, DCO2 and TDCO2). The 13C enrichment of respired CO2 and of phospholipid fatty acids (PLFAs) was determined after 24 h. 13C-glycine incorporation into the biomarker PLFAs for specific microbial groups (Gram positive bacteria, Gram negative bacteria, actinobacteria and fungi) was quantified using gas chromatography-combustion-stable isotope ratio mass spectrometry (GC-C-IRMS). Gram positive bacteria opportunistically utilized the freshly added glycine substrate, i.e. incorporated 13C in all treatments, whereas fungi had minor or no glycine derived 13C-enrichment, hence slowly reacting to a new substrate. The effects of elevated CO2 did suggest increased direct incorporation of glycine in microbial biomass, in particular in G+ bacteria, in an ecosystem subjected to elevated CO2. Warming decreased the concentration of PLFAs in general. The FACE CO2 was 13C-depleted (δ13C = 12.2‰) compared to ambient (δ13C = ∼−8‰), and this enabled observation of the integrated longer term responses of soil microorganisms to the FACE over one year. All together, the bacterial (and not fungal) utilization of glycine indicates substrate preference and resource partitioning in the microbial community, and therefore suggests a diversified response pattern to future

  13. Bacteria and fungi respond differently to multifactorial climate change in a temperate heathland, traced with 13C-glycine and FACE CO2.

    Directory of Open Access Journals (Sweden)

    Louise C Andresen

    Full Text Available It is vital to understand responses of soil microorganisms to predicted climate changes, as these directly control soil carbon (C dynamics. The rate of turnover of soil organic carbon is mediated by soil microorganisms whose activity may be affected by climate change. After one year of multifactorial climate change treatments, at an undisturbed temperate heathland, soil microbial community dynamics were investigated by injection of a very small concentration (5.12 µg C g(-1 soil of (13C-labeled glycine ((13C2, 99 atom % to soils in situ. Plots were treated with elevated temperature (+1°C, T, summer drought (D and elevated atmospheric carbon dioxide (510 ppm [CO2], as well as combined treatments (TD, TCO2, DCO2 and TDCO2. The (13C enrichment of respired CO2 and of phospholipid fatty acids (PLFAs was determined after 24 h. (13C-glycine incorporation into the biomarker PLFAs for specific microbial groups (Gram positive bacteria, Gram negative bacteria, actinobacteria and fungi was quantified using gas chromatography-combustion-stable isotope ratio mass spectrometry (GC-C-IRMS. Gram positive bacteria opportunistically utilized the freshly added glycine substrate, i.e. incorporated (13C in all treatments, whereas fungi had minor or no glycine derived (13C-enrichment, hence slowly reacting to a new substrate. The effects of elevated CO2 did suggest increased direct incorporation of glycine in microbial biomass, in particular in G(+ bacteria, in an ecosystem subjected to elevated CO2. Warming decreased the concentration of PLFAs in general. The FACE CO2 was (13C-depleted (δ(13C = 12.2‰ compared to ambient (δ(13C = ∼-8‰, and this enabled observation of the integrated longer term responses of soil microorganisms to the FACE over one year. All together, the bacterial (and not fungal utilization of glycine indicates substrate preference and resource partitioning in the microbial community, and therefore suggests a diversified response pattern to

  14. Competition along productivity gradients: news from heathlands.

    Science.gov (United States)

    Delerue, Florian; Gonzalez, Maya; Achat, David L; Puzos, Luc; Augusto, Laurent

    2018-05-01

    The importance of competition in low productive habitats is still debated. Studies which simultaneously evaluate preemption of resources and consequences for population dynamics are needed for a comprehensive view of competitive outcomes. We cultivated two emblematic species of European heathlands (Calluna vulgaris and Molinia caerulea) in a nursery for 2 years at two fertility levels, reproducing the productivity gradient found in phosphorus (P)-depleted heathlands in southwest France. The second year, we planted Ulex europaeus seedlings, a ubiquitous heathland species, under the cover of the two species to evaluate its ability to regenerate. Half of the seedlings were placed in tubes for exclusion of competitor roots. We measured the development of the competitors aboveground and belowground and their interception of resources (light, water, inorganic P). Ulex seedlings' growth and survival were also measured. Our results on resources interception were consistent with species distribution in heathlands. Molinia, which dominates rich heathlands, was the strongest competitor for light and water in the rich soil. Calluna, which dominates poor heathlands, increased its root allocation in the poor soil, decreasing water and inorganic P availability. However, the impact of total competition and root competition on Ulex seedlings decreased in the poor soil. Other mechanisms, especially decrease of water stress under neighbouring plant cover, appeared to have more influence on the seedlings' response. We found no formal contradiction between Tilman and Grime's theories. Root competition has a primary role in acquisition of soil resources in poor habitats. However, the importance of competition decreases with decreasing fertility.

  15. Temperate and Tropical Plant Collections

    DEFF Research Database (Denmark)

    Friis, Ib

    2015-01-01

    The first botanical gardens and collections of preserved plants in the 16th century served didactic purposes and should ensure correct identification of medicinal, ornamental and other useful plants. Collections of preserved plants were nearly all book-herbaria, emulating illustrated books...... and owned by individual botanists. Curiosity cabinets of nobles and prominent scholars were larger collections, in which all kinds of objects of natural history from remote regions could be incorporated. The Linnaean revolution favoured loose-leaf herbaria over the old book-herbaria: herbaria with loose...... sheets could be reorganised in agreement with new knowledge or theories and newly accessed specimens could be placed next to earlier ones of the same species. However, the Linnaean collections reflected the essentialist species concept, according to which all species consisted of individuals with similar...

  16. Soil respiration is stimulated by elevated CO2 and reduced by summer drought: three years of measurements in a multifactor ecosystem manipulation experiment in a temperate heathland (CLIMAITE)

    DEFF Research Database (Denmark)

    Selsted, Merete Bang; van der Linden, Leon; Ibrom, Andreas

    2012-01-01

    This study investigated the impact of predicted future climatic and atmospheric conditions on soil respiration (RS) in a Danish Calluna‐Deschampsia‐heathland. A fully factorial in situ experiment with treatments of elevated atmospheric CO2 (+130 ppm), raised soil temperature (+0.4 °C) and extended...... summer drought (5–8% precipitation exclusion) was established in 2005. The average RS, observed in the control over 3 years of measurements (1.7 μmol CO2 m−2 sec−1), increased 38% under elevated CO2, irrespective of combination with the drought or temperature treatments. In contrast, extended summer...... due to reduced plant growth or changes in soil water holding capacity. An empirical model that predicts RS from soil temperature, soil moisture and plant biomass was developed and accounted for 55% of the observed variability in RS. The model predicted annual sums of RS in 2006 and 2007...

  17. Nitrogen-Utilization by Plant-Species from Acid Heathland Soils .2. Growth and Shoot/Root Partitioning of No3- Assimilation at Constant Low Ph and Varying No3-/Nh4+ Ratio

    NARCIS (Netherlands)

    Troelstra, S.R.; Wagenaar, R.; Smant, W.

    1995-01-01

    The growth of four heathland species, two grasses (D. flexuosa, M. caerulea) and two dwarf shrubs (C. vulgaris, E. tetralix), was tested in solution culture at pH 4.0 with 2 mol m(-3) N, varying the NO3-/NH4+ ratio up to 40% nitrate. In addition, measurements of NRA, plant chemical composition, and

  18. Experimental manipulation of succession in heathland ecosystems.

    NARCIS (Netherlands)

    Berendse, F.; Schmitz, M.; Visser, de W.

    1994-01-01

    An experiment was carried out in two heathland ecosystems, one dominated by Calluna vulgaris and the other by Molinia caerulea, to analyse the effects of soil organic matter accumulation and nutrient mineralization on plant species dynamics during succession. The experiment included one treatment

  19. Comparisons of invasive plants in southern Africa originating from southern temperate, northern temperate and tropical regions

    Directory of Open Access Journals (Sweden)

    L. Henderson

    2006-08-01

    Full Text Available A subset of invasive alien plant species in southern Africa was analysed in terms of their history of introduction, rate of spread, countries/region of origin, taxonomy, growth forms, cultivated uses, weed status and current distribution in southern Africa, and comparisons made of those originating from south of the tropic of Capricorn, north of the tropic of Cancer and from the tropics. The subset of 233 species, belonging to 58 families, includes all important declared species and some potentially important species. Almost as many species originate from temperate regions (112 as from the tropics (121. Most southern temperate species came from Australia (28/36, most tropical species from tropical America (92/121 and most northern temperate species from Europe (including the Mediterranean and Asia (58/76. Transformers account for 33% of  all species. More transformers are of tropical origin (36 than of northern temperate (24 and southern temperate origin (18. However. 50% of southern temperate species are transformers, compared to 32% of northern temperate and 29% of tropical species. Southern temperate transformer species are mainly woody trees and shrubs that were established on a grand scale as silvicultural crops, barriers (hedges, windbreaks and screens and cover/binders. Most aquatics, herbs, climbers and succulent shrubs an. trom the tropics. Ornamentals are the single largest category of plants from all three regions, the tropics having contributed twice as many species as temperate regions.

  20. Reptile habitat preference in heathland: implications for heathland management

    NARCIS (Netherlands)

    Stumpel, A.H.P.; Werf, van der D.C.

    2012-01-01

    A two-year reptile survey was conducted in a heathland in the north of the Netherlands, using artificial refuges placed in different habitats. The studied habitats differed in their botanical composition and physical structure. Five reptile species were recorded: slow worm (Anguis fragilis),

  1. Nitrogen acquisition by plants and microorganisms in a temperate grassland.

    Science.gov (United States)

    Liu, Qianyuan; Qiao, Na; Xu, Xingliang; Xin, Xiaoping; Han, Jessie Yc; Tian, Yuqiang; Ouyang, Hua; Kuzyakov, Yakov

    2016-03-10

    Nitrogen (N) limitation is common in most terrestrial ecosystems, often leading to strong competition between microorganisms and plants. The mechanisms of niche differentiation to reduce this competition remain unclear. Short-term (15)N experiments with NH4(+), NO3(-), and glycine were conducted in July, August and September in a temperate grassland to evaluate the chemical, spatial and temporal niche differentiation by competition between plants and microorganisms for N. Microorganisms preferred NH4(+) and NO3(-), while plants preferred NO3(-). Both plants and microorganisms acquired more N in August and September than in July. The soil depth had no significant effects on microbial uptake, but significantly affected plant N uptake. Plants acquired 67% of their N from the 0-5 cm soil layer and 33% from the 5-15 cm layer. The amount of N taken up by microorganisms was at least seven times than plants. Although microorganisms efficiently compete for N with plants, the competition is alleviated through chemical partitioning mainly in deeper soil layer. In the upper soil layer, neither chemical nor temporal niche separation is realized leading to strong competition between plants and microorganisms that modifies N dynamics in grasslands.

  2. Organic matter flow in the food web at a temperate heath under multifactorial climate change

    DEFF Research Database (Denmark)

    Andresen, Louise C.; Konestabo, Heidi S.; Maraldo, Kristine

    2011-01-01

    climate change factors (elevated CO2, increased temperature and drought) were investigated in a full factorial field experiment at a temperate heathland location. The combined effect of biotic and abiotic factors on nitrogen and carbon flows was traced in plant root → litter → microbe → detritivore....../omnivore → predator food‐web for one year after amendment with 15N13C2‐glycine. Isotope ratio mass spectrometry (IRMS) measurement of 15N/14N and 13C/12C in soil extracts and functional ecosystem compartments revealed that the recovery of 15N sometimes decreased through the chain of consumption, with the largest...

  3. Communities of terrestrial nematodes after different approaches to heathland restoration

    Science.gov (United States)

    Radochova, Petra; Frouz, Jan

    2016-04-01

    Since the 20th century, the distribution of European heathlands rapidly decreased due to agricultural intensification, heavy use of artificial fertilizers or acidification (Aerts & Heil, 1993). Therefore, various attempts of heathland restoration are under way in these days. Analysis of nematode community composition can be one of the tools suitable for succession evaluation (Ferris et al., 2001). In 2011, 2013 and 2014, soil samples were collected from heathland restoration experiment (launched in 2011) where different restoration methods were applied in a 3 × 3 factorial experiment; existing heathlands were also sampled to identify the target community both in dry and wet heathland. A total of 60 samples of extracted nematodes were analysed for absolute abundance, trophic groups, and genera dominance. Various indices were calculated to describe the nematode community. We were able to prove faster development of wet heathlands towards the target community. However, because of large data variability, there was no significant difference between treatments. Development of wet and dry heathlands differed also in increased proportion of omniphagous nematodes in 2013 and predators in 2014 in dry heathlands. After three years of heathland restoration, nematode community has not yet reached parameters of the target community. References Aerts, R., Heil, G. W., 1993. Heathlands: patterns and processes in a changing environment, 1st ed, Geobotany: 20. Springer Netherlands, Dordrecht, p. 229. Ferris, H., Bongers, T., De Goede, R. G. M., 2001. A framework for soil food web diagnostics: Extension of the nematode faunal analysis oncept. Appl. Soil Ecol. 18, 13-29.

  4. Danish heathland manipulation experiment data in Model-Data-Fusion

    Science.gov (United States)

    Thum, Tea; Peylin, Philippe; Ibrom, Andreas; Van Der Linden, Leon; Beier, Claus; Bacour, Cédric; Santaren, Diego; Ciais, Philippe

    2013-04-01

    In ecosystem manipulation experiments (EMEs) the ecosystem is artificially exposed to different environmental conditions that aim to simulate circumstances in future climate. At Danish EME site Brandbjerg the responses of a heathland to drought, warming and increased atmospheric CO2 concentration are studied. The warming manipulation is realized by passive nighttime warming. The measurements include control plots as well as replicates for each three treatment separately and in combination. The Brandbjerg heathland ecosystem is dominated by heather and wavy hairgrass. These experiments provide excellent data for validation and development of ecosystem models. In this work we used a generic vegetation model ORCHIDEE with Model-Data-Fusion (MDF) approach. ORCHIDEE model is a process-based model that describes the exchanges of carbon, water and energy between the atmosphere and the vegetation. It can be run at different spatial scales from global to site level. Different vegetation types are described in ORCHIDEE as plant functional types. In MDF we are using observations from the site to optimize the model parameters. This enables us to assess the modelling errors and the performance of the model for different manipulation treatments. This insight will inform us whether the different processes are adequately modelled or if the model is missing some important processes. We used a genetic algorithm in the MDF. The data available from the site included measurements of aboveground biomass, heterotrophic soil respiration and total ecosystem respiration from years 2006-2008. The biomass was measured six times doing this period. The respiration measurements were done with manual chamber measurements. For the soil respiration we used results from an empirical model that has been developed for the site. This enabled us to have more data for the MDF. Before the MDF we performed a sensitivity analysis of the model parameters to different data streams. Fifteen most influential

  5. Amino acid and N mineralization dynamics in heathland soil after long-term warming and repetitive drought

    NARCIS (Netherlands)

    Andresen, L.C.; Bode, S.; Tietema, A.; Boeckx, P.; Rütting, T.

    2015-01-01

    Monomeric organic nitrogen (N) compounds such as free amino acids (FAAs) are an important resource for both plants and soil microorganisms and a source of ammonium (NH4+) via microbial FAA mineralization. We compared gross FAA dynamics with gross N mineralization in a Dutch heathland soil using a

  6. A Comparative-Study on Nutrient Cycling in Wet Heathland Ecosystems.2.Litter Decomposition and Nutrient Mineralization

    NARCIS (Netherlands)

    Berendse, F.; Bobbink, R.; Rouwenhorst, G.

    1989-01-01

    The concept of the relative nutrient requirement (L n) that was introduced in the first paper of this series is used to analyse the effects of the dominant plant population on nutrient cycling and nutrient mineralization in wet heathland ecosystems. A distinction is made between the effect that the

  7. Plants for passive cooling. A preliminary investigation of the use of plants for passive cooling in temperate humid climates

    Energy Technology Data Exchange (ETDEWEB)

    Spirn, A W; Santos, A N; Johnson, D A; Harder, L B; Rios, M W

    1981-04-01

    The potential of vegetation for cooling small, detached residential and commercial structures in temperate, humid climates is discussed. The results of the research are documented, a critical review of the literature is given, and a brief review of energy transfer processes is presented. A checklist of design objectives for passive cooling, a demonstration of design applications, and a palette of selected plant species suitable for passive cooling are included.

  8. No evidence of complementary water use along a plant species richness gradient in temperate experimental grasslands.

    Directory of Open Access Journals (Sweden)

    Dörte Bachmann

    Full Text Available Niche complementarity in resource use has been proposed as a key mechanism to explain the positive effects of increasing plant species richness on ecosystem processes, in particular on primary productivity. Since hardly any information is available for niche complementarity in water use, we tested the effects of plant diversity on spatial and temporal complementarity in water uptake in experimental grasslands by using stable water isotopes. We hypothesized that water uptake from deeper soil depths increases in more diverse compared to low diverse plant species mixtures. We labeled soil water in 8 cm (with 18O and 28 cm depth (with ²H three times during the 2011 growing season in 40 temperate grassland communities of varying species richness (2, 4, 8 and 16 species and functional group number and composition (legumes, grasses, tall herbs, small herbs. Stable isotope analyses of xylem and soil water allowed identifying the preferential depth of water uptake. Higher enrichment in 18O of xylem water than in ²H suggested that the main water uptake was in the upper soil layer. Furthermore, our results revealed no differences in root water uptake among communities with different species richness, different number of functional groups or with time. Thus, our results do not support the hypothesis of increased complementarity in water use in more diverse than in less diverse communities of temperate grassland species.

  9. Facilitation between woody and herbaceous plants that associate with arbuscular mycorrhizal fungi in temperate European forests.

    Science.gov (United States)

    Veresoglou, Stavros D; Wulf, Monika; Rillig, Matthias C

    2017-02-01

    In late-successional environments, low in available nutrient such as the forest understory, herbaceous plant individuals depend strongly on their mycorrhizal associates for survival. We tested whether in temperate European forests arbuscular mycorrhizal (AM) woody plants might facilitate the establishment of AM herbaceous plants in agreement with the mycorrhizal mediation hypothesis. We used a dataset spanning over 400 vegetation plots in the Weser-Elbe region (northwest Germany). Mycorrhizal status information was obtained from published resources, and Ellenberg indicator values were used to infer environmental data. We carried out tests for both relative richness and relative abundance of herbaceous plants. We found that the subset of herbaceous individuals that associated with AM profited when there was a high cover of AM woody plants. These relationships were retained when we accounted for environmental filtering effects using path analysis. Our findings build on the existing literature highlighting the prominent role of mycorrhiza as a coexistence mechanism in plant communities. From a nature conservation point of view, it may be possible to promote functional diversity in the forest understory through introducing AM woody trees in stands when absent.

  10. Effects of nitrogen addition and fire on plant nitrogen use in a temperate steppe.

    Directory of Open Access Journals (Sweden)

    Hai-Wei Wei

    Full Text Available Plant nitrogen (N use strategies have great implications for primary production and ecosystem nutrient cycling. Given the increasing atmospheric N deposition received by most of the terrestrial ecosystems, understanding the responses of plant N use would facilitate the projection of plant-mediated N cycling under global change scenarios. The effects of N deposition on plant N use would be affected by both natural and anthropogenic disturbances, such as prescribed fire in the grassland. We examined the effects of N addition (5.25 g N m(-2 yr(-1 and prescribed fire (annual burning on plant N concentrations and N use characters at both species and community levels in a temperate steppe of northern China. We found that N addition and fire independently affected soil N availability and plant N use traits. Nitrogen addition increased aboveground net primary productivity (ANPP, inorganic N, and N uptake, decreased N response efficiency (NRE, but did not affect biomass-weighed N concentrations at community level. Prescribed fire did not change the community level N concentrations, but largely decreased N uptake efficiency and NRE. At the species level, the effects of N addition and fire on plant N use were species-specific. The divergent responses of plant N use at community and species levels to N addition and fire highlight the importance of the hierarchical responses of plant N use at diverse biological organization levels to the alteration of soil N availability. This study will improve our understanding of the responses of plant-mediated N cycling to global change factors and ecosystem management strategies in the semiarid grasslands.

  11. Effects of human proximity and nomadic grazing on the diversity of medicinal plants in temperate hindukush

    International Nuclear Information System (INIS)

    Adnan, M.; Tariq, A.

    2015-01-01

    Over exploitation and overgrazing are considered major factors for decreasing plant species diversity. However, we do need some more exploration on the types of anthropogenic disturbances responsible for variation in species decline among different regions. Therefore, the current study was conducted in a temperate Hindukush region of Pakistan with the aim to assess human settlements pressure on the diversity of medicinal plants. Field data was collected from 15 plots, of which 5 plots each were randomly selected at three distant locations in derived woodland forest. Near to community plots were heavily disturbed due to natives proximity, 2 kilometer away plots from the native community were moderately disturbed, while 4 kilometer away plots were under the disturbance regime from temporary settlers (Nomads). We have found all 10 studied medicinal plant species at 2 Km distance followed by 8 species found at 4 km distance. Economically important species such as Bergenia ciliata, Paeonia emodi, Podophyllum emodi, Valeriana jatamansi and Viola canescens were completely absent near village. Similarly, density (8.72 n m-2) and species richness (8.2 n 40 m-2) were found significantly highest at 2 km distance than other locations. Detrended Correspondence Analysis (DCA) showed that among forest stand structural variables, slope was the influencing variable related to the density of only Bergenia ciliata. In conclusions, moderate disturbance is associated to increase in the diversity of medicinal plants, while high anthropogenic pressures the otherwise. Hence, effective management strategies may be adopted to conserve such precious eroding flora. (author)

  12. Does prescribed burning result in biotic homogenization of coastal heathlands?

    Science.gov (United States)

    Velle, Liv Guri; Nilsen, Liv Sigrid; Norderhaug, Ann; Vandvik, Vigdis

    2014-05-01

    Biotic homogenization due to replacement of native biodiversity by widespread generalist species has been demonstrated in a number of ecosystems and taxonomic groups worldwide, causing growing conservation concern. Human disturbance is a key driver of biotic homogenization, suggesting potential conservation challenges in seminatural ecosystems, where anthropogenic disturbances such as grazing and burning are necessary for maintaining ecological dynamics and functioning. We test whether prescribed burning results in biotic homogenization in the coastal heathlands of north-western Europe, a seminatural landscape where extensive grazing and burning has constituted the traditional land-use practice over the past 6000 years. We compare the beta-diversity before and after fire at three ecological scales: within local vegetation patches, between wet and dry heathland patches within landscapes, and along a 470 km bioclimatic gradient. Within local patches, we found no evidence of homogenization after fire; species richness increased, and the species that entered the burnt Calluna stands were not widespread specialists but native grasses and herbs characteristic of the heathland system. At the landscapes scale, we saw a weak homogenization as wet and dry heathland patches become more compositionally similar after fire. This was because of a decrease in habitat-specific species unique to either wet or dry habitats and postfire colonization by a set of heathland specialists that established in both habitat types. Along the bioclimatic gradient, species that increased after fire generally had more specific environmental requirements and narrower geographical distributions than the prefire flora, resulting in a biotic 'heterogenisation' after fire. Our study demonstrates that human disturbance does not necessarily cause biotic homogenization, but that continuation of traditional land-use practices can instead be crucial for the maintenance of the diversity and ecological

  13. Herbivores modify selection on plant functional traits in a temperate rainforest understory.

    Science.gov (United States)

    Salgado-Luarte, Cristian; Gianoli, Ernesto

    2012-08-01

    There is limited evidence regarding the adaptive value of plant functional traits in contrasting light environments. It has been suggested that changes in these traits in response to light availability can increase herbivore susceptibility. We tested the adaptive value of plant functional traits linked with carbon gain in contrasting light environments and also evaluated whether herbivores can modify selection on these traits in each light environment. In a temperate rainforest, we examined phenotypic selection on functional traits in seedlings of the pioneer tree Aristotelia chilensis growing in sun (canopy gap) and shade (forest understory) and subjected to either natural herbivory or herbivore exclusion. We found differential selection on functional traits depending on light environment. In sun, there was positive directional selection on photosynthetic rate and relative growth rate (RGR), indicating that selection favors competitive ability in a high-resource environment. Seedlings with high specific leaf area (SLA) and intermediate RGR were selected in shade, suggesting that light capture and conservative resource use are favored in the understory. Herbivores reduced the strength of positive directional selection acting on SLA in shade. We provide the first demonstration that natural herbivory rates can change the strength of selection on plant ecophysiological traits, that is, attributes whose main function is resource uptake. Research addressing the evolution of shade tolerance should incorporate the selective role of herbivores.

  14. WITHIN-POPULATION GENETIC DIVERSITY OF CLIMBING PLANTS AND TREES IN A TEMPERATE FOREST IN CENTRAL CHILE

    OpenAIRE

    Torres-Díaz, Cristian; Ruiz, Eduardo; Salgado-Luarte, Cristian; Molina-Montenegro, Marco A; Gianoli, Ernesto

    2013-01-01

    The climbing habit is a key innovation in angiosperm evolution: climbing plant taxa have greater species richness than their non-climbing sister groups. It is considered that highly diversified clades should show increased among-population genetic differentiation. Less clear is the expected pattern regarding within-population genetic diversity in speciose lineages. We tested the hypothesis of greater within-population genetic diversity in climbing plants compared to trees in a temperate fores...

  15. The Spatiotemporal Dynamics of Forest–Heathland Communities over 60 Years in Fontainebleau, France

    Directory of Open Access Journals (Sweden)

    Samira Mobaied

    2015-06-01

    Full Text Available According to the EU Habitats Directive, heathlands are “natural habitats of community interest”. Heathland management aims at conserving these habitats threatened by various changes, including successional processes leading to forest vegetation. We investigate the dynamics of woody species to the detriment of heathland over a period of 60 years in the Fontainebleau forest and we examine the effects of soil types, soil depth and topography parameters on heathland stability. We assess changes in forest cover between 1946 and 2003 by comparing vegetation maps derived from aerial photographs coupled to GIS analyses. The results show the loss of more than 75% of heathland during 1946–2003 due to tree colonisation of abandoned heathland. We detected differences in the dynamics of colonisation between coniferous and deciduous trees. The colonisation of heathland by coniferous species was faster over the last 20 years of our study period. Tree encroachment was faster in north-facing areas and in areas of acidic luvisols. While this dynamic was very slow in acid sandstone soils, heathland stability was more important in shallow soils on flat and south facing areas. Our study has the potential to assist land managers in selecting those heathland areas that will be easier to conserve and/or to restore by focusing on areas and spatial conditions that prevent forest colonisation and hence favour the long-term stability of heathland.

  16. Climbing plants in a temperate rainforest understorey: searching for high light or coping with deep shade?

    Science.gov (United States)

    Valladares, Fernando; Gianoli, Ernesto; Saldaña, Alfredo

    2011-08-01

    While the climbing habit allows vines to reach well-lit canopy areas with a minimum investment in support biomass, many of them have to survive under the dim understorey light during certain stages of their life cycle. But, if the growth/survival trade-off widely reported for trees hold for climbing plants, they cannot maximize both light-interception efficiency and shade avoidance (i.e. escaping from the understorey). The seven most important woody climbers occurring in a Chilean temperate evergreen rainforest were studied with the hypothesis that light-capture efficiency of climbers would be positively associated with their abundance in the understorey. Species abundance in the understorey was quantified from their relative frequency and density in field plots, the light environment was quantified by hemispherical photography, the photosynthetic response to light was measured with portable gas-exchange analyser, and the whole shoot light-interception efficiency and carbon gain was estimated with the 3-D computer model Y-plant. Species differed in specific leaf area, leaf mass fraction, above ground leaf area ratio, light-interception efficiency and potential carbon gain. Abundance of species in the understorey was related to whole shoot features but not to leaf level features such as specific leaf area. Potential carbon gain was inversely related to light-interception efficiency. Mutual shading among leaves within a shoot was very low (<20 %). The abundance of climbing plants in this southern rainforest understorey was directly related to their capacity to intercept light efficiently but not to their potential carbon gain. The most abundant climbers in this ecosystem match well with a shade-tolerance syndrome in contrast to the pioneer-like nature of climbers observed in tropical studies. The climbers studied seem to sacrifice high-light searching for coping with the dim understorey light.

  17. Genome-Wide Association Study for Traits Related to Plant and Grain Morphology, and Root Architecture in Temperate Rice Accessions.

    Science.gov (United States)

    Biscarini, Filippo; Cozzi, Paolo; Casella, Laura; Riccardi, Paolo; Vattari, Alessandra; Orasen, Gabriele; Perrini, Rosaria; Tacconi, Gianni; Tondelli, Alessandro; Biselli, Chiara; Cattivelli, Luigi; Spindel, Jennifer; McCouch, Susan; Abbruscato, Pamela; Valé, Giampiero; Piffanelli, Pietro; Greco, Raffaella

    2016-01-01

    In this study we carried out a genome-wide association analysis for plant and grain morphology and root architecture in a unique panel of temperate rice accessions adapted to European pedo-climatic conditions. This is the first study to assess the association of selected phenotypic traits to specific genomic regions in the narrow genetic pool of temperate japonica. A set of 391 rice accessions were GBS-genotyped yielding-after data editing-57000 polymorphic and informative SNPS, among which 54% were in genic regions. In total, 42 significant genotype-phenotype associations were detected: 21 for plant morphology traits, 11 for grain quality traits, 10 for root architecture traits. The FDR of detected associations ranged from 3 · 10-7 to 0.92 (median: 0.25). In most cases, the significant detected associations co-localised with QTLs and candidate genes controlling the phenotypic variation of single or multiple traits. The most significant associations were those for flag leaf width on chromosome 4 (FDR = 3 · 10-7) and for plant height on chromosome 6 (FDR = 0.011). We demonstrate the effectiveness and resolution of the developed platform for high-throughput phenotyping, genotyping and GWAS in detecting major QTLs for relevant traits in rice. We identified strong associations that may be used for selection in temperate irrigated rice breeding: e.g. associations for flag leaf width, plant height, root volume and length, grain length, grain width and their ratio. Our findings pave the way to successfully exploit the narrow genetic pool of European temperate rice and to pinpoint the most relevant genetic components contributing to the adaptability and high yield of this germplasm. The generated data could be of direct use in genomic-assisted breeding strategies.

  18. Genome-Wide Association Study for Traits Related to Plant and Grain Morphology, and Root Architecture in Temperate Rice Accessions.

    Directory of Open Access Journals (Sweden)

    Filippo Biscarini

    Full Text Available In this study we carried out a genome-wide association analysis for plant and grain morphology and root architecture in a unique panel of temperate rice accessions adapted to European pedo-climatic conditions. This is the first study to assess the association of selected phenotypic traits to specific genomic regions in the narrow genetic pool of temperate japonica. A set of 391 rice accessions were GBS-genotyped yielding-after data editing-57000 polymorphic and informative SNPS, among which 54% were in genic regions.In total, 42 significant genotype-phenotype associations were detected: 21 for plant morphology traits, 11 for grain quality traits, 10 for root architecture traits. The FDR of detected associations ranged from 3 · 10-7 to 0.92 (median: 0.25. In most cases, the significant detected associations co-localised with QTLs and candidate genes controlling the phenotypic variation of single or multiple traits. The most significant associations were those for flag leaf width on chromosome 4 (FDR = 3 · 10-7 and for plant height on chromosome 6 (FDR = 0.011.We demonstrate the effectiveness and resolution of the developed platform for high-throughput phenotyping, genotyping and GWAS in detecting major QTLs for relevant traits in rice. We identified strong associations that may be used for selection in temperate irrigated rice breeding: e.g. associations for flag leaf width, plant height, root volume and length, grain length, grain width and their ratio. Our findings pave the way to successfully exploit the narrow genetic pool of European temperate rice and to pinpoint the most relevant genetic components contributing to the adaptability and high yield of this germplasm. The generated data could be of direct use in genomic-assisted breeding strategies.

  19. Patterns of plant diversity in seven temperate forest types of Western Himalaya, India

    Directory of Open Access Journals (Sweden)

    Javid Ahmad Dar

    2016-09-01

    Full Text Available Plant biodiversity patterns were analyzed in seven temperate forest types [Populus deltoides (PD, Juglans regia, Cedrus deodara, Pinus wallichiana, mixed coniferous, Abies pindrow (AP and Betula utilis (BU] of Kashmir Himalaya. A total of 177 plant species (158 genera, 66 families were recorded. Most of the species are herbs (82.5%, while shrubs account for 9.6% and trees represent 7.9%. Species richness ranged from 24 (PD to 96 (AP. Shannon, Simpson, and Fisher α indices varied: 0.17–1.06, 0.46–1.22, and 2.01–2.82 for trees; 0.36–0.94, 0.43–0.75, and 0.08–0.35 for shrubs; and 0.35–1.41, 0.27–0.95, and 5.61–39.98 for herbs, respectively. A total of five species were endemic. The total stems and basal area of trees were 35,794 stems (stand mean 330 stems/ha and 481.1 m2 (stand mean 40.2 m2/ha, respectively. The mean density and basal area ranged from 103 stems/ha (BU to 1,201 stems/ha (PD, and from 19.4 m2/ha (BU to 51.9 m2/ha (AP, respectively. Tree density decreased with increase in diameter class. A positive relationship was obtained between elevation and species richness and between elevation and evenness (R2 = 0.37 and 0.19, respectively. Tree and shrub communities were homogenous in nature across the seven forest types, while herbs showed heterogeneous distribution pattern.

  20. The effect of prescribed burning on plant rarity in a temperate forest.

    Science.gov (United States)

    Patykowski, John; Holland, Greg J; Dell, Matt; Wevill, Tricia; Callister, Kate; Bennett, Andrew F; Gibson, Maria

    2018-02-01

    Rare species can play important functional roles, but human-induced changes to disturbance regimes, such as fire, can inadvertently affect these species. We examined the influence of prescribed burns on the recruitment and diversity of plant species within a temperate forest in southeastern Australia, with a focus on species that were rare prior to burning. Floristic composition was compared among plots in landscapes before and after treatment with prescribed burns differing in the extent of area burnt and season of burn (before-after, control-impact design). Floristic surveys were conducted before burns, at the end of a decade of drought, and 3 years postburn. We quantified the effect of prescribed burns on species grouped by their frequency within the landscape before burning (common, less common, and rare) and their life-form attributes (woody perennials, perennial herbs or geophytes, and annual herbs). Burn treatment influenced the response of rare species. In spring-burn plots, the recruitment of rare annual herbs was promoted, differentiating this treatment from both autumn-burn and unburnt plots. In autumn-burn plots, richness of rare species increased across all life-form groups, although composition remained statistically similar to control plots. Richness of rare woody perennials increased in control plots. For all other life-form and frequency groups, the floristic composition of landscapes changed between survey years, but there was no effect of burn treatment, suggesting a likely effect of rainfall on species recruitment. A prescribed burn can increase the occurrence of rare species in a landscape, but burn characteristics can affect the promotion of different life-form groups and thus affect functional diversity. Drought-breaking rain likely had an overarching effect on floristic composition during our study, highlighting that weather can play a greater role in influencing recruitment and diversity in plant communities than a prescribed burn.

  1. Responses of plant community composition and biomass production to warming and nitrogen deposition in a temperate meadow ecosystem.

    Science.gov (United States)

    Zhang, Tao; Guo, Rui; Gao, Song; Guo, Jixun; Sun, Wei

    2015-01-01

    Climate change has profound influences on plant community composition and ecosystem functions. However, its effects on plant community composition and biomass production are not well understood. A four-year field experiment was conducted to examine the effects of warming, nitrogen (N) addition, and their interactions on plant community composition and biomass production in a temperate meadow ecosystem in northeast China. Experimental warming had no significant effect on plant species richness, evenness, and diversity, while N addition highly reduced the species richness and diversity. Warming tended to reduce the importance value of graminoid species but increased the value of forbs, while N addition had the opposite effect. Warming tended to increase the belowground biomass, but had an opposite tendency to decrease the aboveground biomass. The influences of warming on aboveground production were dependent upon precipitation. Experimental warming had little effect on aboveground biomass in the years with higher precipitation, but significantly suppressed aboveground biomass in dry years. Our results suggest that warming had indirect effects on plant production via its effect on the water availability. Nitrogen addition significantly increased above- and below-ground production, suggesting that N is one of the most important limiting factors determining plant productivity in the studied meadow steppe. Significant interactive effects of warming plus N addition on belowground biomass were also detected. Our observations revealed that environmental changes (warming and N deposition) play significant roles in regulating plant community composition and biomass production in temperate meadow steppe ecosystem in northeast China.

  2. Impact of prescribed burning on a heathland inhabiting spider community

    Directory of Open Access Journals (Sweden)

    Krause, Rolf Harald

    2016-04-01

    Full Text Available Heathlands can provide refuge for many stenotopic and endangered arthropods, if habitat management practices are applied. A management measure that is rarely being used today, but which has the potential to support diversity of arthropod communities, is prescribed burning. In this study we investigated the effects of prescribed burning on spider assemblages on a burned site with Calluna vulgaris in the nature reserve Lueneburg Heath, northwest Germany. We used pitfall trapping with a sampling design of 39 traps over a period of one year and 17 sampling intervals on a burned and a control site. We compared overall species richness, activity abundance patterns and community composition of the two sites, with a particular focus on stenotopic and endangered species. We collected 5116 adult spiders and 99 species altogether in a relatively small sampling area. This number of species represents nearly one third of the regional species pool of heathland spider species. Twelve species occurred exclusively on the burned site in contrast to 28 species exclusively found on the unburned site. Although we found more than twice as many spider individuals and higher mean species richness on the control site than on the burned site, the species richness of red-listed spiders was higher on the burned site. Especially the fact that we found 24 endangered species on the burned site and only 20 on the control site indicates that the applied measure of prescribed burning can foster certain endangered spider species and contribute to preserving the overall biodiversity of heathland ecosystems.

  3. Ecosystem-atmosphere exchange of carbon in a heathland under future climatic conditions

    Energy Technology Data Exchange (ETDEWEB)

    Bang Selsted, M

    2010-07-15

    Global change is a reality. Atmospheric CO{sub 2} levels are rising as well as mean global temperature and precipitation patterns are changing. These three environmental factors have separately and in combination effect on ecosystem processes. Terrestrial ecosystems hold large amounts of carbon, why understanding plant and soil responses to such changes are necessary, as ecosystems potentially can ameliorate or accelerate global change. To predict the feedback of ecosystems to the atmospheric CO{sub 2} concentrations experiments imitating global change effects are therefore an important tool. This work on ecosystem-atmosphere exchange of carbon in a heathland under future climatic conditions, shows that extended summer drought in combination with elevated temperature will ensure permanent dryer soil conditions, which decreases carbon turnover, while elevated atmospheric CO{sub 2} concentrations will increase carbon turnover. In the full future climate scenario, carbon turnover is over all expected to increase and the heathland to become a source of atmospheric CO{sub 2}. The methodology of static chamber CO{sub 2} flux measurements and applying the technology in a FACE (free air CO{sub 2} enrichment) facility is a challenge. Fluxes of CO{sub 2} from soil to atmosphere depend on a physical equilibrium between those two medias, why it is important to keep the CO{sub 2} gradient between soil and atmosphere unchanged during measurement. Uptake to plants via photosynthesis depends on a physiological process, which depends strongly on the atmospheric CO{sub 2} concentration. Photosynthesis and respiration run in parallel during measurements of net ecosystem exchange, and these measurements should therefore be performed with care to both the atmospheric CO{sub 2} concentration and the CO{sub 2} soil-atmosphere gradient. (author)

  4. Drivers of temporal changes in temperate forest plant diversity vary across spatial scales

    Czech Academy of Sciences Publication Activity Database

    Bernhardt-Römermann, M.; Baeten, L.; Craven, D.; De Frenne, P.; Hédl, Radim; Lenoir, J.; Bert, D.; Brunet, J.; Chudomelová, Markéta; Decocq, G.; Dierschke, H.; Dirnböck, T.; Dörfler, I.; Heinken, T.; Hermy, M.; Hommel, P.; Jaroszewicz, B.; Keczynski, A.; Kelly, D. L.; Kirkby, K.J.; Kopecký, Martin; Macek, Martin; Máliš, F.; Mirtl, M.; Mitchell, F. J. G.; Naaf, T.; Newman, M.; Peterken, G.; Petřík, Petr; Schmidt, W.; Standovár, T.; Tóth, Z.; Van Calster, H.; Verstraeten, G.; Vladovič, J.; Vild, Ondřej; Wulf, M.; Verheyen, K.

    2015-01-01

    Roč. 21, č. 10 (2015), s. 3726-3737 ISSN 1354-1013 R&D Projects: GA MŠk(CZ) EE2.3.20.0267 EU Projects: European Commission(XE) 278065 - LONGWOOD Institutional support: RVO:67985939 Keywords : temperate forest * long-term change * herbaceous layer Subject RIV: EF - Botanics Impact factor: 8.444, year: 2015

  5. Plant Traits Demonstrate That Temperate and Tropical Giant Eucalypt Forests Are Ecologically Convergent with Rainforest Not Savanna

    Science.gov (United States)

    Tng, David Y. P.; Jordan, Greg J.; Bowman, David M. J. S.

    2013-01-01

    Ecological theory differentiates rainforest and open vegetation in many regions as functionally divergent alternative stable states with transitional (ecotonal) vegetation between the two forming transient unstable states. This transitional vegetation is of considerable significance, not only as a test case for theories of vegetation dynamics, but also because this type of vegetation is of major economic importance, and is home to a suite of species of conservation significance, including the world’s tallest flowering plants. We therefore created predictions of patterns in plant functional traits that would test the alternative stable states model of these systems. We measured functional traits of 128 trees and shrubs across tropical and temperate rainforest – open vegetation transitions in Australia, with giant eucalypt forests situated between these vegetation types. We analysed a set of functional traits: leaf carbon isotopes, leaf area, leaf mass per area, leaf slenderness, wood density, maximum height and bark thickness, using univariate and multivariate methods. For most traits, giant eucalypt forest was similar to rainforest, while rainforest, particularly tropical rainforest, was significantly different from the open vegetation. In multivariate analyses, tropical and temperate rainforest diverged functionally, and both segregated from open vegetation. Furthermore, the giant eucalypt forests overlapped in function with their respective rainforests. The two types of giant eucalypt forests also exhibited greater overall functional similarity to each other than to any of the open vegetation types. We conclude that tropical and temperate giant eucalypt forests are ecologically and functionally convergent. The lack of clear functional differentiation from rainforest suggests that giant eucalypt forests are unstable states within the basin of attraction of rainforest. Our results have important implications for giant eucalypt forest management. PMID:24358359

  6. Testing the plant pneumatic method to estimate xylem embolism resistance in stems of temperate trees

    OpenAIRE

    Zhang, Ya; Lamarque, Laurent J.; Torres-Ruiz, José Manuel; Schuldt, Bernhard; Karimi, Zohreh; Li, Shan; Qin, De-Wen; Bittencourt, Paulo; Burlett, Régis; Cao, Kun-Fang; Delzon, Sylvain; Oliveira, Rafael; Pereira, Luciano; Jansen, Steven

    2018-01-01

    Methods to estimate xylem embolism resistance generally rely on hydraulic measurements, which can be far from straightforward. Recently, a pneumatic method based on air flow measurements of terminal branch ends was proposed to construct vulnerability curves by linking the amount of air extracted from a branch with the degree of embolism. We applied this novel technique for 10 temperate tree species, including six diffuse, two ring-porous and two gymnosperm species, and compared the pneumatic ...

  7. R/D and implement of temper bead welding as newly developed maintenance technique in nuclear power plant

    International Nuclear Information System (INIS)

    Hirano, Shinro; Sera, Takehiko; Chigusa, Naoki; Okimura, Koji; Nishimoto, Kazutoshi

    2011-01-01

    Japanese government has recently addressed a policy to increase capacity factor of existing nuclear PPs to achieve the goal to decrease the emission of CO 2 . Numerous preventive measures have taken in nuclear power plants to minimize the risk of unexpected long shutdown. Newly developed mitigation measures or repair methods need to be qualified to satisfy regulatory standards, before it is implemented to nuclear power plants. The qualification process needs to comply regulatory standards though it may consume time to go through each of the required steps. This paper describes such cases namely ambient temper-bead welding and clarifies the issues that need to be resolved regarding qualification process. The qualification process for new methods that has not been prescribed in regulatory standards temporarily completed by go through confirm testing by JAPEIC, RNP and issuance of no action letter in rush. Currently, the qualification process can only be applied on limited area so generalized qualification process needs to be established. (author)

  8. Growth responses of five desert plants as influenced by biological soil crusts from a temperate desert, China

    Science.gov (United States)

    Zhang, Yuanming; Belnap, Jayne

    2015-01-01

    In almost all dryland systems, biological soil crusts (biocrusts) coexist alongside herbaceous and woody vegetation, creating landscape mosaics of vegetated and biocrusted patches. Results from past studies on the interaction between biocrusts and vascular plants have been contradictory. In the Gurbantunggut desert, a large temperate desert in northwestern China, well-developed lichen-dominated crusts dominate the areas at the base and between the sand dunes. We examined the influence of these lichen-dominated biocrusts on the germination, growth, biomass accumulation, and elemental content of five common plants in this desert: two shrubs (Haloxylon persicum, Ephedra distachya) and three herbaceous plants (Ceratocarpus arenarius, Malcolmia africana and Lappula semiglabra) under greenhouse conditions. The influence of biocrusts on seed germination was species-specific. Biocrusts did not affect percent germination in plants with smooth seeds, but inhibited germination of seeds with appendages that reduced or eliminated contact with the soil surface or prevented seeds from slipping into soil cracks. Once seeds had germinated, biocrusts had different influences on growth of shrub and herbaceous plants. The presence of biocrusts increased concentrations of nitrogen but did not affect phosphorus or potassium in tissue of all tested species, while the uptake of the other tested nutrients was species-specific. Our study showed that biocrusts can serve as a biological filter during seed germination and also can influence growth and elemental uptake. Therefore, they may be an important trigger for determining desert plant diversity and community composition in deserts.

  9. The Species Richness of Vascular Plants and Amphibia in Major Plant Communities in Temperate to Tropical Australia: Relationship with Annual Biomass Production

    International Nuclear Information System (INIS)

    Specht, R. L.; Tyler, M.J.

    2010-01-01

    Aerodynamic fluxes (frictional, thermal, evaporative) in the atmosphere as it flows over and through a plant community determine the Foliage Projective Covers and eco-morphological attributes of new leaves developed annually in overstorey and understorey strata. The number of leaves produced on vertical foliage shoots depends on available soil water and nutrients, also ambient temperature, during this short growth season. Stem density (number of stems per hectare) and species richness (number of species per hectare) in the overstorey of major Floristic Groups are correlated with annual shoot growth (ASG, t ha-1) in that stratum. Species richness in the overstorey increases in the climatic gradient from the arid to the humid zone as well as with increasing air temperatures (about 10 degree C) from temperate to tropical Australia. Species richness in the understorey is highest in plant communities in temperate Australia, decreasing in the temperature gradient towards the tropics. As with other major plant and animal groups within an ecosystem, the species richness of Amphibia is correlated with the amount of solar energy fixed (per annum) by the major plant formation in the regional photosynthetic potential determined by the foliage shoots (ASG, t ha-1) produced annually in the overstorey.

  10. The Species Richness of Vascular Plants and Amphibia in Major Plant Communities in Temperate to Tropical Australia: Relationship with Annual Biomass Production

    Directory of Open Access Journals (Sweden)

    R. L. Specht

    2010-01-01

    Full Text Available Aerodynamic fluxes (frictional, thermal, evaporative in the atmosphere as it flows over and through a plant community determine the Foliage Projective Covers and eco-morphological attributes of new leaves developed annually in overstorey and understorey strata. The number of leaves produced on vertical foliage shoots depends on available soil water and nutrients, also ambient temperature, during this short growth season. Stem density (number of stems per hectare and species richness (number of species per hectare in the overstorey of major Floristic Groups are correlated with annual shoot growth (ASG, t ha−1 in that stratum. Species richness in the overstorey increases in the climatic gradient from the arid to the humid zone as well as with increasing air temperatures (about 10oC from temperate to tropical Australia. Species richness in the understorey is highest in plant communities in temperate Australia, decreasing in the temperature gradient towards the tropics. As with other major plant and animal groups within an ecosystem, the species richness of Amphibia is correlated with the amount of solar energy fixed (per annum by the major plant formation in the region—a photosynthetic potential determined by the foliage shoots (ASG, t ha−1 produced annually in the overstorey.

  11. Effects of soil depth and plant-soil interaction on microbial community in temperate grasslands of northern China.

    Science.gov (United States)

    Yao, Xiaodong; Zhang, Naili; Zeng, Hui; Wang, Wei

    2018-07-15

    Although the patterns and drivers of soil microbial community composition are well studied, little is known about the effects of plant-soil interactions and soil depth on soil microbial distribution at a regional scale. We examined 195 soil samples from 13 sites along a climatic transect in the temperate grasslands of northern China to measure the composition of and factors influencing soil microbial communities within a 1-m soil profile. Soil microbial community composition was measured using phospholipid fatty acids (PLFA) analysis. Fungi predominated in topsoil (0-10 cm) and bacteria and actinomycetes in deep soils (40-100 cm), independent of steppe types. This variation was explained by contemporary environmental factors (including above- and below-ground plant biomass, soil physicochemical and climatic factors) >58% in the 0-40 cm of soil depth, but soils. Interestingly, when we considered the interactive effects between plant traits (above ground biomass and root biomass) and soil factors (pH, clay content, and soil total carbon, nitrogen, phosphorous), we observed a significant interaction effect occurring at depths of 10-20 cm soil layer, due to different internal and external factors of the plant-soil system along the soil profile. These results improve understanding of the drivers of soil microbial community composition at regional scales. Copyright © 2018 Elsevier B.V. All rights reserved.

  12. Temperate heath plant response to dry conditions depends on growth strategy and less on physiology

    DEFF Research Database (Denmark)

    Albert, Kristian Rost; Kongstad, J.; Schmidt, I. K.

    2012-01-01

    of these differences in response in dry versus rewetting conditions can be used to highlight the limitations coherent in different strategies adopted by, for example, evergreen shrubs and grasses. We investigated the leaf-level photosynthetic performance, leaf C, N and d13C along with vegetation cover and biomass...... in the evergreen dwarf shrub Calluna vulgaris and the grass species Deschampsia flexuosa in a temperate heath during seasonal changes in soil moisture. Higher photosynthetic capacity compensated for lower stomatal conductance and sustained higher rates of photosynthesis in the grass compared to the dwarf shrub....... In combination with dieback of aboveground biomass and reduction of stomatal conductance reduction during dry conditions, the grass continued to have high carbon uptake in the remaining leaves. The dwarf shrub endured the dry conditions by preserving shoot biomass and reducing stomatal conductance. Soil...

  13. Plant litter chemistry and mycorrhizal roots promote a nitrogen feedback in a temperate forest.

    Science.gov (United States)

    Nina Wurzburger; Ronald L. Hendrick

    2009-01-01

    1. Relationships between mycorrhizal plants and soil nitrogen (N) have led to the speculation that the chemistry of plant litter and the saprotrophy of mycorrhizal symbionts can function together to...

  14. Vascular plant species richness along environmental gradients in a cool temperate to sub-alpine mountainous zone in central Japan.

    Science.gov (United States)

    Tsujino, Riyou; Yumoto, Takakazu

    2013-03-01

    In order to clarify how vegetation types change along the environmental gradients in a cool temperate to sub-alpine mountainous zone and the determinant factors that define plant species richness, we established 360 plots (each 4 × 10 m) within which the vegetation type, species richness, elevation, topographic position index (TPI), slope inclination, and ground light index (GLI) of the natural vegetation were surveyed. Mean elevation, TPI, slope inclination, and GLI differed across vegetation types. Tree species richness was negatively correlated with elevation, whereas fern and herb species richness were positively correlated. Tree species richness was greater in the upper slope area than the lower slope area, whereas fern and herb species richness were greater in the lower slope area. Ferns and trees species richness were smaller in the open canopy, whereas herb species richness was greater in the open canopy. Vegetation types were determined firstly by elevation and secondary by topographic configurations, such as topographic position, and slope inclination. Elevation and topography were the most important factors affecting plant richness, but the most influential variables differed among plant life-form groups. Moreover, the species richness responses to these environmental gradients greatly differed among ferns, herbs, and trees.

  15. Diet selection and performance of horses grazing on different heathland types.

    Science.gov (United States)

    López López, C; Ferreira, L M M; García, U; Moreno-Gonzalo, J; Rodrigues, M A M; Osoro, K; Ferre, I; Celaya, R

    2017-10-01

    The number of horses in northern Spanish mountains has increased in recent decades, but little is known about their grazing behaviour, performance and potential for foal meat production. This research aimed to study the diet selection, liveweight (LW) changes and parasitic status of dry and lactating mares, and foals' LW gains, grazing on heathlands with different botanical composition. The experimental design consisted of three vegetation types: dominated by heather (Ericaceae) species (H), dominated by gorse (Ulex gallii; G) and co-dominated by gorse and heath-grasses (G-G), with four replicates per treatment (12 paddocks of 1.2 ha). The study lasted three grazing seasons (2010-12). Each year, 24 crossbred mature mares (310±52 kg LW) were used, managing one lactating mare with her foal plus one non-lactating mare per paddock from May to late summer or early autumn. In the case of H paddocks, animals had to be removed before (late August to early September) because of apparent loss of body condition. Animals were periodically weighed. Mares' diet composition was estimated using alkane markers, analysing the discrepancies in alkane concentrations between dietary plant components and faeces. Faecal samples were also analysed for gastrointestinal nematodes ova. Chemical composition of the main plant components (i.e. heather, gorse and grasses) revealed a low nutritive value, averaging 79, 115 and 113 g CP/kg dry matter (DM), respectively, that could restrict livestock performance. Mares initially selected gorse and grasses (0.47 and 0.40, respectively, in 2010), increasing heather consumption over time (from 0.13 in 2010 to 0.29 in 2012) as gorse availability decreased. The performance of both mares and foals was lower in H compared with G and G-G paddocks (-216 v. 347 g/day for mares, Phorses on gorse- and grass-gorse-dominated shrublands could be sustainable at least during part of the year (4 to 6 months). However, heather-dominated heathlands are not able to

  16. Vegetation Dynamics and Community Assembly in Post-Agricultural Heathland

    DEFF Research Database (Denmark)

    Kepfer Rojas, Sebastian

    that land-use legacies are still present in the soil and were important determinants of vegetation dynamics and community assembly. However, the effects of land-use legacies were mostly mediated by the understory vegetation and differed according to the functional groups. The distance to the edge, a proxy...... for the proximity to external seed sources, was an important factor affecting different components of the structure of the vegetation, demonstrating the importance of dispersal in the development of the community. My results indicate that the effect of the biotic interactions varies along abiotic gradients (e......Summary This PhD study aims at understanding how biotic, abiotic and stochastic factors interact to structure a heathland vegetation community managed under different traditional land-use practices for centuries prior to abandonment ca. 120 years ago. This study is part of one of the longest...

  17. A novel soil manganese mechanism drives plant species loss with increased nitrogen deposition in a temperate steppe.

    Science.gov (United States)

    Tian, Qiuying; Liu, Nana; Bai, Wenming; Li, Linghao; Chen, Jiquan; Reich, Peter B; Yu, Qiang; Guo, Dali; Smith, Melinda D; Knapp, Alan K; Cheng, Weixin; Lu, Peng; Gao, Yan; Yang, An; Wang, Tianzuo; Li, Xin; Wang, Zhengwen; Ma, Yibing; Han, Xingguo; Zhang, Wen-Hao

    2016-01-01

    Loss of plant diversity with increased anthropogenic nitrogen (N) deposition in grasslands has occurred globally. In most cases, competitive exclusion driven by preemption of light or space is invoked as a key mechanism. Here, we provide evidence from a 9-yr N-addition experiment for an alternative mechanism: differential sensitivity of forbs and grasses to increased soil manganese (Mn) levels. In Inner Mongolia steppes, increasing the N supply shifted plant community composition from grass-forb codominance (primarily Stipa krylovii and Artemisia frigida, respectively) to exclusive dominance by grass, with associated declines in overall species richness. Reduced abundance of forbs was linked to soil acidification that increased mobilization of soil Mn, with a 10-fold greater accumulation of Mn in forbs than in grasses. The enhanced accumulation of Mn in forbs was correlated with reduced photosynthetic rates and growth, and is consistent with the loss of forb species. Differential accumulation of Mn between forbs and grasses can be linked to fundamental differences between dicots and monocots in the biochemical pathways regulating metal transport. These findings provide a mechanistic explanation for N-induced species loss in temperate grasslands by linking metal mobilization in soil to differential metal acquisition and impacts on key functional groups in these ecosystems.

  18. Nematode community shifts in response to experimental warming and canopy conditions are associated with plant community changes in the temperate-boreal forest ecotone.

    Science.gov (United States)

    Thakur, Madhav Prakash; Reich, Peter B; Fisichelli, Nicholas A; Stefanski, Artur; Cesarz, Simone; Dobies, Tomasz; Rich, Roy L; Hobbie, Sarah E; Eisenhauer, Nico

    2014-06-01

    Global climate warming is one of the key forces driving plant community shifts, such as range shifts of temperate species into boreal forests. As plant community shifts are slow to observe, ecotones, boundaries between two ecosystems, are target areas for providing early evidence of ecological responses to warming. The role of soil fauna is poorly explored in ecotones, although their positive and negative effects on plant species can influence plant community structure. We studied nematode communities in response to experimental warming (ambient, +1.7, +3.4 °C) in soils of closed and open canopy forest in the temperate-boreal ecotone of Minnesota, USA and calculated various established nematode indices. We estimated species-specific coverage of understory herbaceous and shrub plant species from the same experimental plots and tested if changes in the nematode community are associated with plant cover and composition. Individual nematode trophic groups did not differ among warming treatments, but the ratio between microbial-feeding and plant-feeding nematodes increased significantly and consistently with warming in both closed and open canopy areas and at both experimental field sites. The increase in this ratio was positively correlated with total cover of understory plant species, perhaps due to increased predation pressure on soil microorganisms causing higher nutrient availability for plants. Multivariate analyses revealed that temperature treatment, canopy conditions and nematode density consistently shaped understory plant communities across experimental sites. Our findings suggest that warming-induced changes in nematode community structure are associated with shifts in plant community composition and productivity in the temperate-boreal forest ecotones.

  19. The impact of forest roads on understory plant diversity in temperate hornbeam-beech forests of Northern Iran.

    Science.gov (United States)

    Deljouei, Azade; Abdi, Ehsan; Marcantonio, Matteo; Majnounian, Baris; Amici, Valerio; Sohrabi, Hormoz

    2017-08-01

    Forest roads alter the biotic and abiotic components of ecosystems, modifying temperature, humidity, wind speed, and light availability that, in turn, cause changes in plant community composition and diversity. We aim at investigating and comparing the diversity of herbaceous species along main and secondary forest roads in a temperate-managed hornbeam-beech forest, north of Iran. Sixteen transects along main and secondary forest roads were established (eight transects along main roads and eight along secondary roads). To eliminate the effect of forest type, all transects were located in Carpinetum-Fagetum forests, the dominant forest type in the study area. The total length of each transect was 200 m (100 m toward up slope and 100 m toward down slope), and plots were established along it at different distances from road edge. The diversity of herbaceous plant species was calculated in each plot using Shannon-Wiener index, species richness, and Pielou's index. The results showed that diversity index decreased when distance from road edge increases. This decreasing trend continued up to 60 m from forest road margin, and after this threshold, the index slightly increased. Depending on the type of road (main or secondary) as well as cut or fill slopes, the area showing a statistical different plant composition and diversity measured through Shannon-Wiener, species richness, and Pielou's index is up to 10 m. The length depth of the road edge effect found in main and secondary forest roads was small, but it could have cumulative effects on forest microclimate and forest-associated biota at the island scale. Forest managers should account for the effect of road buildings on plant communities.

  20. Drag forces of common plant species in temperate streams: consequences of morphology, velocity and biomass

    DEFF Research Database (Denmark)

    Jensen, Kaj Sand

    2008-01-01

    Swift flow in streams may physically influence the morphology and distribution of plants. I quantified drag as a function of velocity, biomass and their interaction on the trailing canopy of seven European stream species in an experimental flume and evaluated its importance for species distributi...... than an uneven distribution with the same biomass confined to dense patches surrounded by open flow channels. Thus, management strategies to ensure a patchy plants distribution should be suitable for combining agricultural drainage and ecological stream quality....

  1. Effects of experimental drought on microbial processes in two temperate heathlands at contrasting water conditions

    DEFF Research Database (Denmark)

    Jensen, K.D.; Beier, C.; Michelsen, A.

    2003-01-01

    by a 27% reduced below ground CO(2) emission, and reduced microbial and soil solution carbon (C) and nitrogen (N) levels. In contrast, microbial activity at the wetter UK site seemed to benefit from the drought as indicated by a 22% increase in below ground CO(2) emission caused by the drought treatment...

  2. The influence of canopy-layer composition on understory plant diversity in southern temperate forests

    Directory of Open Access Journals (Sweden)

    Luciana Mestre

    2017-05-01

    Full Text Available Background Understory plants represents the largest component of biodiversity in most forest ecosystems and plays a key role in forest functioning. Despite their importance, the influence of overstory-layer composition on understory plant diversity is relatively poorly understood within deciduous-evergreen broadleaved mixed forests. The aim of this work was to evaluate how tree overstory-layer composition influences on understory-layer diversity in three forest types (monospecific deciduous Nothofagus pumilio (Np, monospecific evergreen Nothofagus betuloides (Nb, and mixed N. pumilio-N. betuloides (M forests, comparing also between two geographical locations (coast and mountain to estimate differences at landscape level. Results We recorded 46 plant species: 4 ferns, 12 monocots, and 30 dicots. Canopy-layer composition influences the herb-layer structure and diversity in two different ways: while mixed forests have greater similarity to evergreen forests in the understory structural features, deciduous and mixed were similar in terms of the specific composition of plant assemblage. Deciduous pure stands were the most diverse, meanwhile evergreen stands were least diverse. Lack of exclusive species of mixed forest could represent a transition where evergreen and deciduous communities meet and integrate. Moreover, landscape has a major influence on the structure, diversity and richness of understory vegetation of pure and mixed forests likely associated to the magnitude and frequency of natural disturbances, where mountain forest not only had highest herb-layer diversity but also more exclusive species. Conclusions Our study suggests that mixed Nothofagus forest supports coexistence of both pure deciduous and pure evergreen understory plant species and different assemblages in coastal and mountain sites. Maintaining the mixture of canopy patch types within mixed stands will be important for conserving the natural patterns of understory plant

  3. Effect of fuel treatments and backfiring on the recovery of an obligate seeder-dominated heathland

    Energy Technology Data Exchange (ETDEWEB)

    Fernández Filgueira, C.; Vega Hidalgo, J.A.

    2016-07-01

    Aim of the study: To evaluate how a plant community responded to a backfire that occurred four years after application of different types of fuel-reduction treatments. Area of study: Erica umbellata Loefl. (L.)-dominated heathland in Galicia (NW Spain). Materials and Methods: Shrub cover surveys in 16 experimental plots from 2006 to 2014. Fuel reduction treatments (prescribed burning, clearing and mastication) were applied in the spring of 2006 and the area was burned by a wildfire in the summer of 2010. Main results: Shrub total cover recovered quickly after the backfire in both the treated and untreated areas, and the pre-treatment values were reached four years after the fire. Post-wildfire resprouting species cover recovery was not affected by fuel treatments. As a contrast, Erica umbellata cover reached levels similar to those in the untreated plots only in the areas treated by prescribed burning. After the wildfire, grasses cover recovery was greater in the treated than in the untreated areas and the effect lasted until the end of the study. Research highlights: Prescribed fire and backfire was favourable for Erica umbellata regeneration compared to clearing and mastication. (Author)

  4. Effect of fuel treatments and backfiring on the recovery of an obligate seeder-dominated heathland

    Directory of Open Access Journals (Sweden)

    Cristina Fernández Filgueira

    2016-12-01

    Full Text Available Aim of study: To evaluate how a plant community responded to a backfire that occurred four years after application of different types of fuel-reduction treatments. Area of study: Erica umbellata Loefl. (L.-dominated heathland in Galicia (NW Spain. Materials and Methods: Shrub cover surveys in 16 experimental plots from 2006 to 2014. Fuel reduction treatments (prescribed burning, clearing and mastication were applied in the spring of 2006 and the area was burned by a wildfire in the summer of 2010. Main results: Shrub total cover recovered quickly after the backfire in both the treated and untreated areas, and the pre-treatment values were reached four years after the fire. Post-wildfire resprouting species cover recovery was not affected by fuel treatments. As a contrast,  Erica umbellata cover reached levels similar to those in the untreated plots only in the areas treated by prescribed burning. After the wildfire, grasses cover recovery was greater in the treated than in the untreated areas and the effect lasted until the end of the study. Research highlights: Prescribed fire and backfire was favourable for Erica umbellata regeneration compared to clearing and mastication. Keywords: prescribed burning; clearing; mechanical shredding; Erica; wildfire.

  5. Temperate Interior West community tree guide: benefits, costs, and strategic planting

    Science.gov (United States)

    Kelaine E. Vargas; E. Gregory McPherson; James R. Simpson; Paula J. Peper; Shelley L. Gardner; Qingfu Xiao

    2007-01-01

    Even as they increase the beauty of our surroundings, trees provide us with a great many ecosystem services, including air quality improvement, energy conservation, stormwater interception, and atmospheric carbon dioxide reduction. These benefits must be weighed against the costs of maintaining trees, including planting, pruning, irrigation, administration, pest...

  6. Alien plants in temperate weed communities: Prehistoric and recent invaders occupy different habitats

    Czech Academy of Sciences Publication Activity Database

    Pyšek, Petr; Jarošík, Vojtěch; Chytrý, M.; Kropáč, Z.; Tichý, L.; Wild, Jan

    2005-01-01

    Roč. 86, č. 3 (2005), s. 772-785 ISSN 0012-9658 R&D Projects: GA ČR(CZ) GA526/00/1443 Institutional research plan: CEZ:AV0Z60050516 Keywords : plant invasions * agricultural weeds * residence time Subject RIV: EF - Botanics Impact factor: 4.506, year: 2005

  7. The effect of topsoil removal in restored heathland on soil fauna, topsoil microstructure, and cellulose decomposition : implications for ecosystem restoration

    NARCIS (Netherlands)

    Frouz, Jan; Van Diggelen, Rudy; Pizl, Vaclav; Stary, Josef; Hanel, Ladislav; Tajovsky, Karel; Kalcik, Jiri

    2009-01-01

    Communities of soil macrofauna, oribatid mites, and nematodes as well as vegetation and soil chemistry were studied on twelve plots representing three replicates of the following treatments: agricultural meadow, heathland, and heathland restored either by partial or complete topsoil removal 15 years

  8. Temper Tantrums

    Science.gov (United States)

    ... Nine Steps to More Effective Parenting How Can Parents Discipline Without Spanking? Delayed Speech or Language Development Talking to Your Child's Preschool Teacher Your Child's Habits Separation Anxiety Breath-Holding Spells Train Your Temper View ...

  9. Extensive Management Promotes Plant and Microbial Nitrogen Retention in Temperate Grassland

    Science.gov (United States)

    de Vries, Franciska T.; Bloem, Jaap; Quirk, Helen; Stevens, Carly J.; Bol, Roland; Bardgett, Richard D.

    2012-01-01

    Leaching losses of nitrogen (N) from soil and atmospheric N deposition have led to widespread changes in plant community and microbial community composition, but our knowledge of the factors that determine ecosystem N retention is limited. A common feature of extensively managed, species-rich grasslands is that they have fungal-dominated microbial communities, which might reduce soil N losses and increase ecosystem N retention, which is pivotal for pollution mitigation and sustainable food production. However, the mechanisms that underpin improved N retention in extensively managed, species-rich grasslands are unclear. We combined a landscape-scale field study and glasshouse experiment to test how grassland management affects plant and soil N retention. Specifically, we hypothesised that extensively managed, species-rich grasslands of high conservation value would have lower N loss and greater N retention than intensively managed, species-poor grasslands, and that this would be due to a greater immobilisation of N by a more fungal-dominated microbial community. In the field study, we found that extensively managed, species-rich grasslands had lower N leaching losses. Soil inorganic N availability decreased with increasing abundance of fungi relative to bacteria, although the best predictor of soil N leaching was the C/N ratio of aboveground plant biomass. In the associated glasshouse experiment we found that retention of added 15N was greater in extensively than in intensively managed grasslands, which was attributed to a combination of greater root uptake and microbial immobilisation of 15N in the former, and that microbial immobilisation increased with increasing biomass and abundance of fungi. These findings show that grassland management affects mechanisms of N retention in soil through changes in root and microbial uptake of N. Moreover, they support the notion that microbial communities might be the key to improved N retention through tightening linkages

  10. Environmental Drivers of Patterns of Plant Diversity Along a Wide Environmental Gradient in Korean Temperate Forests

    Directory of Open Access Journals (Sweden)

    Chang-Bae Lee

    2016-01-01

    Full Text Available Understanding patterns of biodiversity and their drivers along environmental gradients is one of the central topics in ecology. However, whether diversity patterns along environmental gradients differ among diversity components as well as life forms and what kind of variables control or interact to shape the diversity patterns are poorly known. This study scrutinized the distribution patterns of three plant groups with four diversity indices and evaluated the effects of regional area, topography, topographic heterogeneity, climate, primary productivity, vegetation structure diversity and vegetation type diversity along an extensive elevational gradient on the Baekdudaegan Mountains in South Korea. Different elevational patterns, including hump-shaped, reversed hump-shaped, increasing, multimodal and no relationship, were observed among both the diversity indices and the plant groups. Regional area, habitat heterogeneity and climate were included to explain most of the elevational diversity patterns. In particular, habitat heterogeneity was the most important variable for explaining the patterns of diversity. The results suggest that patterns of elevational diversity may differ not only among plant groups but also among diversity indices and that such patterns are primarily caused by habitat heterogeneity in the Baekdudaegan Mountains because more heterogeneous and diverse habitats can support more coexisting species.

  11. Plant community and litter composition in temperate deciduous woodlots along two field gradients of soil Ni, Cu and Co concentrations

    International Nuclear Information System (INIS)

    Hale, Beverley; Robertson, Paul

    2016-01-01

    Perennial plant communities in the proximity of metal smelters and refineries may receive substantial inputs of base metal particulate as well as sulphate from the co-emission of sulphur dioxide. The Ni refinery at Port Colborne (Canada) operated by Inco (now Vale Canada Ltd.) emitted Ni, Co and Cu, along with sulphur dioxide, between 1918 and 1984. The objectives were to determine if vascular plant community composition, including standing litter, in twenty-one woodlots on clay or organic soil, were related to soil Ni concentration which decreased in concentration with distance from the Ni refinery. The soil Ni concentration in the clay woodlots ranged from 16 to 4130 mg Ni/kg, and in the organic woodlots, ranged from 98 to 22,700 mg Ni/kg. The concentrations of Co and Cu in the soils were also elevated, and highly correlated with soil Ni concentration. In consequence, each series of woodlots constituted a ‘fixed ratio ray’ of metal mixture exposure. For each of the woodlots, there were 16 independent measurements of ‘woodlot status’ which were correlated with elevated soil Ni concentration. Of the 32 combinations, there were eight linear correlations with soil Ni concentration, considerably more than would be expected by chance alone at a p-value of 0.05. With the exception of mean crown rating for shrubs at the clay sites, the correlations were consistent with the hypothesis that increased soil metal concentrations would be correlated with decreased diversity, plant community health or fitness, and increased accumulation of litter. Only five of the eight linear correlations were from the organic woodlots, suggesting that the observations were not confounded with soil type nor range in soil metal concentrations. - Highlights: • Temperate woodlots on organic or clay soils with gradient of soil Ni were studied. • Soil Ni ranged up to 4100 mg/kg on clay and up to 22,700 mg/kg on organic. • Most indices of plant community status were not correlated

  12. Plant molecular phylogeography in China and adjacent regions: Tracing the genetic imprints of Quaternary climate and environmental change in the world's most diverse temperate flora.

    Science.gov (United States)

    Qiu, Ying-Xiong; Fu, Cheng-Xing; Comes, Hans Peter

    2011-04-01

    The Sino-Japanese Floristic Region (SJFR) of East Asia harbors the most diverse of the world's temperate flora, and was the most important glacial refuge for its Tertiary representatives ('relics') throughout Quaternary ice-age cycles. A steadily increasing number of phylogeographic studies in the SJFR of mainland China and adjacent areas, including the Qinghai-Tibetan-Plateau (QTP) and Sino-Himalayan region, have documented the population histories of temperate plant species in these regions. Here we review this current literature that challenges the oft-stated view of the SJFR as a glacial sanctuary for temperate plants, instead revealing profound effects of Quaternary changes in climate, topography, and/or sea level on the current genetic structure of such organisms. There are three recurrent phylogeographic scenarios identified by different case studies that broadly agree with longstanding biogeographic or palaeo-ecological hypotheses: (i) postglacial re-colonization of the QTP from (south-)eastern glacial refugia; (ii) population isolation and endemic species formation in Southwest China due to tectonic shifts and river course dynamics; and (iii) long-term isolation and species survival in multiple localized refugia of (warm-)temperate deciduous forest habitats in subtropical (Central/East/South) China. However, in four additional instances, phylogeographic findings seem to conflict with a priori predictions raised by palaeo-data, suggesting instead: (iv) glacial in situ survival of some hardy alpine herbs and forest trees on the QTP platform itself; (v) long-term refugial isolation of (warm-)temperate evergreen taxa in subtropical China; (vi) 'cryptic' glacial survival of (cool-)temperate deciduous forest trees in North China; and (vii) unexpectedly deep (Late Tertiary/early-to-mid Pleistocene) allopatric-vicariant differentiation of disjunct lineages in the East China-Japan-Korea region due to past sea transgressions. We discuss these and other consequences

  13. Tending for Cattle: Traditional Fire Management in Ethiopian Montane Heathlands

    Directory of Open Access Journals (Sweden)

    Maria U. Johansson

    2012-09-01

    Full Text Available Fire has long been a principal tool for manipulating ecosystems, notably for pastoralist cultures, but in modern times, fire use has often been a source of conflicts with state bureaucracies. Despite this, traditional fire management practices have rarely been examined from a perspective of fire behavior and fire effects, which hampers dialogue on management options. In order to analyze the rationale for fire use, its practical handling, and ecological effects in high-elevation ericaceous heathlands in Ethiopia, we used three different information sources: interviews with pastoralists, field observations of fires, and analysis of vegetation age structure at the landscape level. The interviews revealed three primary reasons for burning: increasing the grazing value, controlling a toxic caterpillar, and reducing predator attacks. Informants were well aware of critical factors governing fire behavior, such as slope, wind, vertical and horizontal fuel structure, and fuel moisture. Recent burns (1-4 years since fire were used as firebreaks to control the size of individual burns, which resulted in a mosaic of vegetation of different ages. The age structure indicated an average fire return interval of ~10 years. At these elevations (> 3500 m, the dry period is unreliable, with occasional rains. Of all observed fires, 83% were ignited during very high Fire Weather Index levels, reached during only 11% of all days of the year. Burning is illegal, but if this ban was respected, our data suggest that the Erica shrubs would grow out of reach of cattle within a few years only, creating a dense and continuous canopy. This would also create a risk of large high-intensity wildfires since the landscape is virtually devoid of natural fuel breaks. Under the present management regime, this heathland ecosystem should be quite resilient to degradation by fire due to a relatively slow fuel buildup (limiting fire intervals and an effective regrowth of Erica shoots

  14. Sustained by First Nations: European newcomers' use of Indigenous plant foods in temperate North America

    Directory of Open Access Journals (Sweden)

    Nancy J. Turner

    2012-12-01

    Full Text Available Indigenous Peoples of North America have collectively used approximately 1800 different native species of plants, algae, lichens and fungi as food. When European explorers, traders and settlers arrived on the continent, these native foods, often identified and offered by Indigenous hosts, gave them sustenance and in some cases saved them from starvation. Over the years, some of these species – particularly various types of berries, such as blueberries and cranberries (Vaccinium spp., wild raspberries and blackberries (Rubus spp., and wild strawberries (Fragaria spp., and various types of nuts (Corylus spp., Carya spp., Juglans spp., Pinus spp., along with wild-rice (Zizania spp. and maple syrup (from Acer saccharum – became more widely adopted and remain in use to the present day. Some of these and some other species were used in plant breeding programs, as germplasm for hybridization programs, or to strengthen a crop's resistance to disease. At the same time, many nutritious Indigenous foods fell out of use among Indigenous Peoples themselves, and along with their lessened use came a loss of associated knowledge and cultural identity. Today, for a variety of reasons, from improving people's health and regaining their cultural heritage, to enhancing dietary diversity and enjoyment of diverse foods, some of the species that have dwindled in their use have been “rediscovered” by Indigenous and non-Indigenous Peoples, and indications are that their benefits to humanity will continue into the future.

  15. Lichen ecology of the Scunthorpe heathlands. I. Mineral accumulation

    Energy Technology Data Exchange (ETDEWEB)

    Seaward, M R.D.

    1973-01-01

    An investigation into the distributional and ecological characteristics of the terricolous lichen flora of the Scunthorpe heathlands is described. The Risby Warren area was especially studied, because there is a noticeable terricolous lichen desert and distinctive zonation of lichen communities, each of which incorporates Peltigera rufescens. Peltigera rufescens appeared to be quite tolerant of air pollution in this region, and occurred in sufficient quantities for periodic sampling; this proved valuable for determination of the sulfur, chromium, copper, iron, lead, manganese, and nickel content of the thalli along a 780 m transect. Frequency assessments, together with these element determinations, revealed sharp qualitative and quantitative improvements in the lichen flora with increasing distance from the pollution sources. The stable lichen pattern, together with the mineral accumulation measurements, was indicative of a pollution dome effect operating to a distance of approximately 3 km. Within 100 to 200 m of this critical distance, lichen material had a pronounced metallic accumulation; the latter was compared with urban and herbarium material. The possible use of lichens for the sampling of ambient environmental conditions is discussed.

  16. Prospects for the development of disease-resistant temperate fruit plants by mutation induction

    International Nuclear Information System (INIS)

    Campbell, A.I.; Wilson, D.

    1977-01-01

    In most of the present conventional fruit breeding programmes disease resistance has become an important objective. Progress is slow because of the long generation time and the genetic complexity of most tree fruit species. The complexity is such that cultivars can only be maintained as clones and it is unlikely that identical genotypes could ever be sexually produced. Hence, the prospect of changing a few characters in an otherwise unchanged genetic background, as might be done by somatic mutation, is attractive. The occurence of natural mutations in some fruit cultivars and the induction of mutations in others demonstrates that such an approach is possible for some characters at least and these may include disease resistance. The yet limited success of mutation breeding in fruit crops may be due in part to the innate difficulties with this group of plants but may also be a consequence of the faulty methods that have been used in the past. New techniques of inducing and selecting mutants in fruit trees are reported, with particular reference to disease resistance and some basic guidelines for success are suggested. The type of disease resistance required will undoubtedly affect the approach used. In theory, monogenic resistance seems more likely to respond to change by mutation induction than polygenic resistance. However, the multiple effects seen in the natural spur-type apple mutants and in the preliminary results with induced apple mutations at Long Ashton suggest that field resistance to some major diseases may not be an unreasonable target

  17. A hyperspectral approach to estimating biomass and plant production in a heterogeneous restored temperate peatland

    Science.gov (United States)

    Byrd, K. B.; Schile, L. M.; Windham-Myers, L.; Kelly, M.; Hatala, J.; Baldocchi, D. D.

    2012-12-01

    Restoration of drained peatlands that are managed to reverse subsidence through organic accretion holds significant potential for large-scale carbon storage and sequestration. This potential has been demonstrated in an experimental wetland restoration site established by the U.S. Geological Survey in 1997 on Twitchell Island in the Sacramento-San Joaquin River Delta, where soil carbon storage is up to 1 kg C m-2 and root and rhizome production can reach over 7 kg m-2 annually. Remote sensing-based estimation of biomass and productivity over a large spatial extent helps to monitor carbon storage potential of these restored peatlands. Extensive field measurements of plant biophysical characteristics such as biomass, leaf area index, and the fraction of absorbed photosynthetically active radiation (fAPAR) [an important variable in light-use efficiency (LUE) models] have been collected for agricultural systems and forests. However the small size and local spatial variability of U.S. Pacific Coast wetlands pose new challenges for measuring these variables in the field and generating estimates through remote sensing. In particular background effects of non-photosynthetic vegetation (NPV), floating aquatic vegetation, and inundation of wetland vegetation influence the relationship between field measurements and multispectral or hyperspectral indices. Working at the USGS experimental wetland site, characterized by variable water depth and substantial NPV, or thatch, we collected field data on hardstem bulrush (Schoenoplectus acutus) and cattail (Typha spp.) coupled with reflectance data from a field spectrometer (350-2500 nm) every two to three weeks during the summers of 2011 and 2012. We calculated aboveground biomass with existing allometric relationships, and fAPAR was measured with line and point quantum sensors. We analyzed reflectance data to develop hyperspectral and multispectral indices that predict biomass and fAPAR and account for background effects of water

  18. Soil Respiration on an Aging Managed Heathland: Identifying an Appropriate Empirical Model for Predictive Purposes

    NARCIS (Netherlands)

    Kopittke, G.R.; van Loon, E.E.; Tietema, A.; Asscheman, D.

    2013-01-01

    Heathlands are cultural landscapes which are managed through cyclical cutting, burning or grazing practices. Understanding the carbon (C) fluxes from these ecosystems provides information on the optimal management cycle time to maximise C uptake and minimise C output. The interpretation of field

  19. A comparative study on nutrient cycling in wet heathland ecosystems : II. Litter decomposition and nutrient mineralization.

    Science.gov (United States)

    Berendse, Frank; Bobbink, Roland; Rouwenhorst, Gerrit

    1989-03-01

    The concept of the relative nutrient requirement (L n ) that was introduced in the first paper of this series is used to analyse the effects of the dominant plant population on nutrient cycling and nutrient mineralization in wet heathland ecosystems. A distinction is made between the effect that the dominant plant species has on (1) the distribution of nutrients over the plant biomass and the soil compartment of the ecosystem and (2) the recirculation rate of nutrients. The first effect of the dominant plant species can be calculated on the basis of the δ/k ratio (which is the ratio of the relative mortality to the decomposition constant). The second effect can be analysed using the relative nutrient requirement (L n ). The mass loss and the changes in the amounts of N and P in decomposing above-ground and below-ground litter produced by Erica tetralix and Molinia caerulea were measured over three years. The rates of mass loss from both above-ground and below-ground litter of Molinia were higher than those from Erica litter. After an initial leaching phase, litter showed either a net release or a net immobilization of nitrogen or phosphorus that depended on the initial concentrations of these nutrients. At the same sites, mineralization of nitrogen and phosphorus were measured for two years both in communities dominated by Molinia and in communities dominated by Erica. There were no clear differences in the nitrogen mineralization, but in one of the two years, phosphate mineralization in the Molinia-community was significantly higher. On the basis of the theory that was developed, mineralization rates and ratios between amounts of nutrients in plant biomass and in the soil were calculated on the basis of parameters that were independently measured. There was a reasonable agreement between predicted and measured values in the Erica-communities. In the Molinia-communities there were large differences between calculated and measured values, which was explained by the

  20. Nitrogen cycling in heathland ecosystems and effects of climate change

    DEFF Research Database (Denmark)

    Andresen, Louise Christoffersen

    Terrestrial ecosystems are currently exposed to climatic and air quality changes with increased atmospheric CO2, increased temperature and periodical droughts. At a temperate heath site this was investigated in a unique full factorial in situ experiment (CLIMAITE). The climate change treatments...

  1. Radiation preservation of foods of plant origin. Part V. Temperate fruits: pome fruits, stone fruits, and berries

    International Nuclear Information System (INIS)

    Thomas, P.

    1986-01-01

    The current status of research on the application of ionizing radiation for improving the storage of temperate fruits, i.e., apple, pear, peach, nectarine, apricot, cherry, plum, strawberry, bilberry, cranberry, raspberry, and black currant, is reviewed. Changes in fruit metabolism, chemical composition, texture, and organoleptic quality attributes are discussed with reference to the irradiation dose. The feasibility of using radiation either alone or in conjunction with heat treatment, refrigeration, and controlled atmospheres (CA) for the control of storage decay caused by fungal pathogens is considered. Areas of further research are suggested before irradiation could be considered for practical application in some of these temperate fruits. The recent trends in the possible use of irradiation for disinfestation of certain pome and stone fruits and the prospects for the commercial utilization of irradiation for improving the market life of strawberries are discussed. 156 references

  2. Long-Term Effects of White-Tailed Deer Exclusion on the Invasion of Exotic Plants: A Case Study in a Mid-Atlantic Temperate Forest.

    Directory of Open Access Journals (Sweden)

    Xiaoli Shen

    Full Text Available Exotic plant invasions and chronic high levels of herbivory are two of the major biotic stressors impacting temperate forest ecosystems in eastern North America, and the two problems are often linked. We used a 4-ha deer exclosure maintained since 1991 to examine the influence of a generalist herbivore, white-tailed deer (Odocoileus virginianus, on the abundance of four exotic invasive (Rosa multiflora, Berberis thunbergii, Rubus phoenicolasius and Microstegium vimineum and one native (Cynoglossum virginianum plant species, within a 25.6-ha mature temperate forest dynamics plot in Virginia, USA. We identified significant predictors of the abundance of each focal species using generalized linear models incorporating 10 environmental and landscape variables. After controlling for those predictors, we applied our models to a 4-ha deer exclusion site and a 4-ha reference site, both embedded within the larger plot, to test the role of deer on the abundance of the focal species. Slope, edge effects and soil pH were the most frequent predictors of the abundance of the focal species on the larger plot. The abundance of C. virginianum, known to be deer-dispersed, was significantly lower in the exclosure. Similar patterns were detected for B. thunbergii, R. phoenicolasius and M. vimineum, whereas R. multiflora was more abundant within the exclosure. Our results indicate that chronic high deer density facilitates increased abundances of several exotic invasive plant species, with the notable exception of R. multiflora. We infer that the invasion of many exotic plant species that are browse-tolerant to white-tailed deer could be limited by reducing deer populations.

  3. Tempered fractional calculus

    Energy Technology Data Exchange (ETDEWEB)

    Sabzikar, Farzad, E-mail: sabzika2@stt.msu.edu [Department of Statistics and Probability, Michigan State University, East Lansing, MI 48823 (United States); Meerschaert, Mark M., E-mail: mcubed@stt.msu.edu [Department of Statistics and Probability, Michigan State University, East Lansing, MI 48823 (United States); Chen, Jinghua, E-mail: cjhdzdz@163.com [School of Sciences, Jimei University, Xiamen, Fujian, 361021 (China)

    2015-07-15

    Fractional derivatives and integrals are convolutions with a power law. Multiplying by an exponential factor leads to tempered fractional derivatives and integrals. Tempered fractional diffusion equations, where the usual second derivative in space is replaced by a tempered fractional derivative, govern the limits of random walk models with an exponentially tempered power law jump distribution. The limiting tempered stable probability densities exhibit semi-heavy tails, which are commonly observed in finance. Tempered power law waiting times lead to tempered fractional time derivatives, which have proven useful in geophysics. The tempered fractional derivative or integral of a Brownian motion, called a tempered fractional Brownian motion, can exhibit semi-long range dependence. The increments of this process, called tempered fractional Gaussian noise, provide a useful new stochastic model for wind speed data. A tempered fractional difference forms the basis for numerical methods to solve tempered fractional diffusion equations, and it also provides a useful new correlation model in time series.

  4. Tempered fractional calculus

    Science.gov (United States)

    Sabzikar, Farzad; Meerschaert, Mark M.; Chen, Jinghua

    2015-07-01

    Fractional derivatives and integrals are convolutions with a power law. Multiplying by an exponential factor leads to tempered fractional derivatives and integrals. Tempered fractional diffusion equations, where the usual second derivative in space is replaced by a tempered fractional derivative, govern the limits of random walk models with an exponentially tempered power law jump distribution. The limiting tempered stable probability densities exhibit semi-heavy tails, which are commonly observed in finance. Tempered power law waiting times lead to tempered fractional time derivatives, which have proven useful in geophysics. The tempered fractional derivative or integral of a Brownian motion, called a tempered fractional Brownian motion, can exhibit semi-long range dependence. The increments of this process, called tempered fractional Gaussian noise, provide a useful new stochastic model for wind speed data. A tempered fractional difference forms the basis for numerical methods to solve tempered fractional diffusion equations, and it also provides a useful new correlation model in time series.

  5. Tempered fractional calculus

    International Nuclear Information System (INIS)

    Sabzikar, Farzad; Meerschaert, Mark M.; Chen, Jinghua

    2015-01-01

    Fractional derivatives and integrals are convolutions with a power law. Multiplying by an exponential factor leads to tempered fractional derivatives and integrals. Tempered fractional diffusion equations, where the usual second derivative in space is replaced by a tempered fractional derivative, govern the limits of random walk models with an exponentially tempered power law jump distribution. The limiting tempered stable probability densities exhibit semi-heavy tails, which are commonly observed in finance. Tempered power law waiting times lead to tempered fractional time derivatives, which have proven useful in geophysics. The tempered fractional derivative or integral of a Brownian motion, called a tempered fractional Brownian motion, can exhibit semi-long range dependence. The increments of this process, called tempered fractional Gaussian noise, provide a useful new stochastic model for wind speed data. A tempered fractional difference forms the basis for numerical methods to solve tempered fractional diffusion equations, and it also provides a useful new correlation model in time series

  6. Indirect effects of land-use legacies determine tree colonization patterns in abandoned heathland

    DEFF Research Database (Denmark)

    Kepfer Rojas, Sebastian; Verheyen, Kris; Johannsen, Vivian Kvist

    2015-01-01

    of tree/shrubs in the heathland. Further, we used high-resolution LiDAR data to classify the vegetation and identify forest patches. In the analysis, we first used a logistic mixed model to test whether colonization of tree and shrub species differed between areas with different land-use history......Questions How do land-use legacies and distance to forest patches influence tree colonization at a post-agricultural heathland? Are colonizing species with different life-history traits affected differently by these factors? Is the effect of increased nutrient availability from land-use legacies...... and whether it was influenced by the distance to forest patches and life-history traits (seed mass) of colonizing species. Then, to determine how different factors influence colonization, we explored the direct and indirect relationships among nutrient availability, density of adult trees, canopy cover, cover...

  7. Ecosystem-atmosphere exchange of carbon in a heathland under future climatic conditions

    DEFF Research Database (Denmark)

    Selsted, Merete Bang

    on ecosystem-atmosphere exchange of carbon in a heathland under future climatic conditions, shows that extended summer drought in combination with elevated temperature will ensure permanent dryer soil conditions, which decreases carbon turnover, while elevated atmospheric CO2 concentrations will increase...... carbon turnover. In the full future climate scenario, carbon turnover is over all expected to increase and the heathland to become a source of atmospheric CO2. The methodology of static chamber CO2 flux measurements and applying the technology in a FACE (free air CO2 enrichment) facility is a challenge...... on the atmospheric CO2 concentration. Photosynthesis and respiration run in parallel during measurements of net ecosystem exchange, and these measurements should therefore be performed with care to both the atmospheric CO2 concentration and the CO2 soil-atmosphere gradient....

  8. Analysis of yield and plant traits of oilseed rape (Brassica napus L. cultivated in temperate region in light of the possibilities of sowing in arid areas

    Directory of Open Access Journals (Sweden)

    Tadeusz Zając

    2016-12-01

    Full Text Available This work is a review of selected literature on the species of Brassica with the greatest economic significance. Oilseed rape (Brassica napus ssp. oleifera currently ranks third worldwide among oilseed crops used for oil production and is the most important in the temperate zone. The manifold uses of rape include not only human consumption of oil, but also the use of post-extraction meal to feed livestock as well as industrial applications as a source of bioenergy or cellulose. The improvement in the economic position of rape among crop plants is also due to the doubling of its yield between 1970 and 2009; the average annual increase in seed yield worldwide was 27 kg ha−1 yr−1. The yield level in Europe exceeds the average yields achieved in the world, particularly in Asia. Recently, the cultivation of oilseed rape was started on a relatively large acreage in Iran where the yield amounted 2.1 t ha−1, exceeding the yields of China and India. In Poland, the acreage of oilseed rape cultivation between 1965 and 2013 increased 3–4 times, and during this period the annual increase in seed yield was 29 kg ha−1 yr−1. Under the field conditions of the temperate climate zone, winter oilseed rape yield is mainly determined by agro-climatic conditions during the growing period, the level of nitrogen fertilization, and the production potential of varieties, which is currently highest in hybrids. There is a noticeable tendency of hybrids towards formation of more siliques by individual oilseed plants. Different production categories of plants appear in a rape crop. Semi-dwarf varieties of winter rapeseed are distinguished by greater silique density, particularly on the main shoot. Moreover, these hybrids are characterized by faster growth of the root system, which enables them to take up nitrogen from the soil more efficiently.

  9. Temporal and spatial influences incur reconfiguration of Arctic heathland soil bacterial community structure.

    Science.gov (United States)

    Hill, Richard; Saetnan, Eli R; Scullion, John; Gwynn-Jones, Dylan; Ostle, Nick; Edwards, Arwyn

    2016-06-01

    Microbial responses to Arctic climate change could radically alter the stability of major stores of soil carbon. However, the sensitivity of plot-scale experiments simulating climate change effects on Arctic heathland soils to potential confounding effects of spatial and temporal changes in soil microbial communities is unknown. Here, the variation in heathland soil bacterial communities at two survey sites in Sweden between spring and summer 2013 and at scales between 0-1 m and, 1-100 m and between sites (> 100 m) were investigated in parallel using 16S rRNA gene T-RFLP and amplicon sequencing. T-RFLP did not reveal spatial structuring of communities at scales structuring effects may not confound comparison between plot-scale treatments, temporal change is a significant influence. Moreover, the prominence of two temporally exclusive keystone taxa suggests that the stability of Arctic heathland soil bacterial communities could be disproportionally influenced by seasonal perturbations affecting individual taxa. © 2015 Society for Applied Microbiology and John Wiley & Sons Ltd.

  10. Changes of Species Richness in Heathland Communities over 15 Years following Disturbances

    Directory of Open Access Journals (Sweden)

    L. Calvo

    2012-01-01

    Full Text Available The aim of this study was to define the species richness patterns over a period of 15 years during the vegetation recovery process after disturbances (burning, cutting and ploughing in heathlands. Three communities were selected: two dominated by Erica australis and one dominated by Calluna vulgaris. The alpha and gamma diversity patterns were site specific and influenced by the ecological traits of dominant shrub species. The shrubland dominated by Erica australis, typical resprouters with a fast regeneration, showed the highest values of alpha and gamma diversity during the first 7 years of regeneration. The heathland dominated by Calluna vulgaris, an obligate seeder, had a contrasting pattern of alpha and gamma diversity, as the highest values appeared from year 7 until year 14. Thus, the speed of regeneration of the dominant shrub species could be the main factor affecting structural parameters in these communities. Species richness patterns did not vary in relation to the different types of perturbation. Cutting and burning would be the most suitable forestry management strategies to conserve Erica australis heathlands, but burning is more appropriate in Calluna vulgaris ones because cutting modified this community.

  11. Effects of elevated CO2, warming and drought episodes on plant carbon uptake in a temperate heath ecosystem are controlled by soil water status

    DEFF Research Database (Denmark)

    Albert, Kristian Rost; Ro-Poulsen, H.; Mikkelsen, Teis Nørgaard

    2011-01-01

    The impact of elevated CO2, periodic drought and warming on photosynthesis and leaf characteristics of the evergreen dwarf shrub Calluna vulgaris in a temperate heath ecosystem was investigated. Photosynthesis was reduced by drought in midsummer and increased by elevated CO2 throughout the growing...... season, whereas warming only stimulated photosynthesis early in the year. At the beginning and end of the growing season, a T × CO2 interaction synergistically stimulated plant carbon uptake in the combination of warming and elevated CO2. At peak drought, the D × CO2 interaction antagonistically down......-regulated photosynthesis, suggesting a limited ability of elevated CO2 to counteract the negative effect of drought. The response of photosynthesis in the full factorial combination (TDCO2) could be explained by the main effect of experimental treatments (T, D, CO2) and the two-factor interactions (D × CO2, T × CO2...

  12. Water level, vegetation composition, and plant productivity explain greenhouse gas fluxes in temperate cutover fens after inundation

    Science.gov (United States)

    Minke, Merten; Augustin, Jürgen; Burlo, Andrei; Yarmashuk, Tatsiana; Chuvashova, Hanna; Thiele, Annett; Freibauer, Annette; Tikhonov, Vitalij; Hoffmann, Mathias

    2016-07-01

    Peat extraction leaves a land surface with a strong relief of deep cutover areas and higher ridges. Rewetting inundates the deep parts, while less deeply extracted zones remain at or above the water level. In temperate fens the flooded areas are colonized by helophytes such as Eriophorum angustifolium, Carex spp., Typha latifolia or Phragmites australis dependent on water depth. Reeds of Typha and Phragmites are reported as large sources of methane, but data on net CO2 uptake are contradictory for Typha and rare for Phragmites. Here, we analyze the effect of vegetation, water level and nutrient conditions on greenhouse gas (GHG) emissions for representative vegetation types along water level gradients at two rewetted cutover fens (mesotrophic and eutrophic) in Belarus. Greenhouse gas emissions were measured campaign-wise with manual chambers every 2 to 4 weeks for 2 years and interpolated by modelling. All sites had negligible nitrous oxide exchange rates. Most sites were carbon sinks and small GHG sources. Methane emissions generally increased with net ecosystem CO2 uptake. Mesotrophic small sedge reeds with water table around the land surface were small GHG sources in the range of 2.3 to 4.2 t CO2 eq. ha-1 yr-1. Eutrophic tall sedge - Typha latifolia reeds on newly formed floating mats were substantial net GHG emitters in the range of 25.1 to 39.1 t CO2 eq. ha-1 yr. They represent transient vegetation stages. Phragmites reeds ranged between -1.7 to 4.2 t CO2 eq. ha-1 yr-1 with an overall mean GHG emission of 1.3 t CO2 eq. ha-1 yr-1. The annual CO2 balance was best explained by vegetation biomass, which includes the role of vegetation composition and species. Methane emissions were obviously driven by biological activity of vegetation and soil organisms. Shallow flooding of cutover temperate fens is a suitable measure to arrive at low GHG emissions. Phragmites australis establishment should be promoted in deeper flooded areas and will lead to moderate, but

  13. Temporal Changes in Socio-Ecological Systems and Their Impact on Ecosystem Services at Different Governance Scales: A Case Study of Heathlands

    NARCIS (Netherlands)

    Moran-Ordonez, A.; Bugter, R.J.F.; Suarez-Seoane, S.; Luis, de E.; Calvo, L.

    2013-01-01

    Heathlands are considered biodiversity hotspots of high conservation interest. However, they are at risk of degradation and disappearance in most parts of Europe mainly due to land abandonment, degradation, and conversion to other land uses. Heathlands are semi-natural systems: their maintenance and

  14. Succession influences wild bees in a temperate forest landscape: the value of early successional stages in naturally regenerated and planted forests.

    Science.gov (United States)

    Taki, Hisatomo; Okochi, Isamu; Okabe, Kimiko; Inoue, Takenari; Goto, Hideaki; Matsumura, Takeshi; Makino, Shun'ichi

    2013-01-01

    In many temperate terrestrial forest ecosystems, both natural human disturbances drive the reestablishment of forests. Succession in plant communities, in addition to reforestation following the creation of open sites through harvesting or natural disturbances, can affect forest faunal assemblages. Wild bees perform an important ecosystem function in human-altered and natural or seminatural ecosystems, as they are essential pollinators for both crops and wild flowering plants. To maintain high abundance and species richness for pollination services, it is important to conserve and create seminatural and natural land cover with optimal successional stages for wild bees. We examined the effects of forest succession on wild bees. In particular, we evaluated the importance of early successional stages for bees, which has been suspected but not previously demonstrated. A range of successional stages, between 1 and 178 years old, were examined in naturally regenerated and planted forests. In total 4465 wild bee individuals, representing 113 species, were captured. Results for total bees, solitary bees, and cleptoparasitic bees in both naturally regenerated and planted conifer forests indicated a higher abundance and species richness in the early successional stages. However, higher abundance and species richness of social bees in naturally regenerated forest were observed as the successional stages progressed, whereas the abundance of social bees in conifer planted forest showed a concave-shaped relationship when plotted. The results suggest that early successional stages of both naturally regenerated and conifer planted forest maintain a high abundance and species richness of solitary bees and their cleptoparasitic bees, although social bees respond differently in the early successional stages. This may imply that, in some cases, active forest stand management policies, such as the clear-cutting of planted forests for timber production, would create early successional

  15. Succession influences wild bees in a temperate forest landscape: the value of early successional stages in naturally regenerated and planted forests.

    Directory of Open Access Journals (Sweden)

    Hisatomo Taki

    Full Text Available In many temperate terrestrial forest ecosystems, both natural human disturbances drive the reestablishment of forests. Succession in plant communities, in addition to reforestation following the creation of open sites through harvesting or natural disturbances, can affect forest faunal assemblages. Wild bees perform an important ecosystem function in human-altered and natural or seminatural ecosystems, as they are essential pollinators for both crops and wild flowering plants. To maintain high abundance and species richness for pollination services, it is important to conserve and create seminatural and natural land cover with optimal successional stages for wild bees. We examined the effects of forest succession on wild bees. In particular, we evaluated the importance of early successional stages for bees, which has been suspected but not previously demonstrated. A range of successional stages, between 1 and 178 years old, were examined in naturally regenerated and planted forests. In total 4465 wild bee individuals, representing 113 species, were captured. Results for total bees, solitary bees, and cleptoparasitic bees in both naturally regenerated and planted conifer forests indicated a higher abundance and species richness in the early successional stages. However, higher abundance and species richness of social bees in naturally regenerated forest were observed as the successional stages progressed, whereas the abundance of social bees in conifer planted forest showed a concave-shaped relationship when plotted. The results suggest that early successional stages of both naturally regenerated and conifer planted forest maintain a high abundance and species richness of solitary bees and their cleptoparasitic bees, although social bees respond differently in the early successional stages. This may imply that, in some cases, active forest stand management policies, such as the clear-cutting of planted forests for timber production, would create

  16. The Spatial Pattern and Interactions of Woody Plants on the Temperate Savanna of Inner Mongolia, China: The Effects of Alternating Seasonal Grazing-Mowing Regimes.

    Directory of Open Access Journals (Sweden)

    Xiao Wang

    Full Text Available Ulmus pumila tree-dominated temperate savanna, which is distributed widely throughout the forest-steppe ecotone on the Mongolian Plateau, is a relatively stable woody-herbaceous complex ecosystem in northern China. Relatively more attention has been paid to the degradation of typical steppe areas, whereas less focus has been placed on the succession of this typical temperate savanna under the present management regime. In this study, we established 3 sample plots 100 m×100 m in size along a gradient of fixed distances from one herder's stationary site and then surveyed all the woody plants in these plots. A spatial point pattern analysis was employed to clarify the spatial distribution and interaction of these woody plants. The results indicated that old U. pumila trees (DBH ≥ 20 cm showed a random distribution and that medium U. pumila trees (5 cm ≤ DBH < 20 cm showed an aggregated distribution at a smaller scale and a random distribution at a larger scale; few or no juvenile trees (DBH < 5 cm were present, and seedlings (without DBH formed aggregations in all 3 plots. These findings can be explained by an alternate seasonal grazing-mowing regime (exclosure in summer, mowing in autumn and grazing in winter and spring; the shrubs in all 3 plots exist along a grazing gradient that harbors xerophytic and mesophytic shrubs. Of these shrubs, xerophytic shrubs show significant aggregation at a smaller scale (0-5.5 m, whereas mesophytic shrubs show significant aggregation at a larger scale (0-25 m, which may be the result of the dual effects of grazing pressure and climate change. Medium trees and seedlings significantly facilitate the distributions of xerophytic shrubs and compete significantly with mesophytic shrubs due to differences in water use strategies. We conclude that the implementation of an alternative grazing-mowing regime results in xerophytic shrub encroachment or existence, breaking the chain of normal succession in a U. pumila

  17. Environment vs. Plant Ontogeny: Arthropod Herbivory Patterns on European Beech Leaves along the Vertical Gradient of Temperate Forests in Central Germany

    Directory of Open Access Journals (Sweden)

    Stephanie Stiegel

    2018-01-01

    Full Text Available Environmental and leaf trait effects on herbivory are supposed to vary among different feeding guilds. Herbivores also show variability in their preferences for plant ontogenetic stages. Along the vertical forest gradient, environmental conditions change, and trees represent juvenile and adult individuals in the understorey and canopy, respectively. This study was conducted in ten forests sites in Central Germany for the enrichment of canopy research in temperate forests. Arthropod herbivory of different feeding traces was surveyed on leaves of Fagus sylvatica Linnaeus (European beech; Fagaceae in three strata. Effects of microclimate, leaf traits, and plant ontogenetic stage were analyzed as determining parameters for herbivory. The highest herbivory was caused by exophagous feeding traces. Herbivore attack levels varied along the vertical forest gradient for most feeding traces with distinct patterns. If differences of herbivory levels were present, they only occurred between juvenile and adult F. sylvatica individuals, but not between the lower and upper canopy. In contrast, differences of microclimate and important leaf traits were present between the lower and upper canopy. In conclusion, the plant ontogenetic stage had a stronger effect on herbivory than microclimate or leaf traits along the vertical forest gradient.

  18. Structural and functional responses of plant communities to climate change-mediated alterations in the hydrology of riparian areas in temperate Europe.

    Science.gov (United States)

    Baattrup-Pedersen, Annette; Garssen, Annemarie; Göthe, Emma; Hoffmann, Carl Christian; Oddershede, Andrea; Riis, Tenna; van Bodegom, Peter M; Larsen, Søren E; Soons, Merel

    2018-04-01

    The hydrology of riparian areas changes rapidly these years because of climate change-mediated alterations in precipitation patterns. In this study, we used a large-scale in situ experimental approach to explore effects of drought and flooding on plant taxonomic diversity and functional trait composition in riparian areas in temperate Europe. We found significant effects of flooding and drought in all study areas, the effects being most pronounced under flooded conditions. In near-stream areas, taxonomic diversity initially declined in response to both drought and flooding (although not significantly so in all years) and remained stable under drought conditions, whereas the decline continued under flooded conditions. For most traits, we found clear indications that the functional diversity also declined under flooded conditions, particularly in near-stream areas, indicating that fewer strategies succeeded under flooded conditions. Consistent changes in community mean trait values were also identified, but fewer than expected. This can have several, not mutually exclusive, explanations. First, different adaptive strategies may coexist in a community. Second, intraspecific variability was not considered for any of the traits. For example, many species can elongate shoots and petioles that enable them to survive shallow, prolonged flooding but such abilities will not be captured when applying mean trait values. Third, we only followed the communities for 3 years. Flooding excludes species intolerant of the altered hydrology, whereas the establishment of new species relies on time-dependent processes, for instance the dispersal and establishment of species within the areas. We expect that altered precipitation patterns will have profound consequences for riparian vegetation in temperate Europe. Riparian areas will experience loss of taxonomic and functional diversity and, over time, possibly also alterations in community trait responses that may have cascading effects

  19. Leaf out times of temperate woody plants are related to phylogeny, deciduousness, growth habit and wood anatomy.

    Science.gov (United States)

    Panchen, Zoe A; Primack, Richard B; Nordt, Birgit; Ellwood, Elizabeth R; Stevens, Albert-Dieter; Renner, Susanne S; Willis, Charles G; Fahey, Robert; Whittemore, Alan; Du, Yanjun; Davis, Charles C

    2014-09-01

    Leaf out phenology affects a wide variety of ecosystem processes and ecological interactions and will take on added significance as leaf out times increasingly shift in response to warming temperatures associated with climate change. There is, however, relatively little information available on the factors affecting species differences in leaf out phenology. An international team of researchers from eight Northern Hemisphere temperate botanical gardens recorded leaf out dates of c. 1600 woody species in 2011 and 2012. Leaf out dates in woody species differed by as much as 3 months at a single site and exhibited strong phylogenetic and anatomical relationships. On average, angiosperms leafed out earlier than gymnosperms, deciduous species earlier than evergreen species, shrubs earlier than trees, diffuse and semi-ring porous species earlier than ring porous species, and species with smaller diameter xylem vessels earlier than species with larger diameter vessels. The order of species leaf out was generally consistent between years and among sites. As species distribution and abundance shift due to climate change, interspecific differences in leaf out phenology may affect ecosystem processes such as carbon, water, and nutrient cycling. Our open access leaf out data provide a critical framework for monitoring and modelling such changes going forward. © 2014 The Authors. New Phytologist © 2014 New Phytologist Trust.

  20. Relationships between Plant Diversity and the Abundance and α-Diversity of Predatory Ground Beetles (Coleoptera: Carabidae) in a Mature Asian Temperate Forest Ecosystem

    Science.gov (United States)

    Zou, Yi; Sang, Weiguo; Bai, Fan; Axmacher, Jan Christoph

    2013-01-01

    A positive relationship between plant diversity and both abundance and diversity of predatory arthropods is postulated by the Enemies Hypothesis, a central ecological top-down control hypothesis. It has been supported by experimental studies and investigations of agricultural and grassland ecosystems, while evidence from more complex mature forest ecosystems is limited. Our study was conducted on Changbai Mountain in one of the last remaining large pristine temperate forest environments in China. We used predatory ground beetles (Coleoptera: Carabidae) as target taxon to establish the relationship between phytodiversity and their activity abundance and diversity. Results showed that elevation was the only variable included in both models predicting carabid activity abundance and α-diversity. Shrub diversity was negatively and herb diversity positively correlated with beetle abundance, while shrub diversity was positively correlated with beetle α-diversity. Within the different forest types, a negative relationship between plant diversity and carabid activity abundance was observed, which stands in direct contrast to the Enemies Hypothesis. Furthermore, plant species density did not predict carabid α-diversity. In addition, the density of herbs, which is commonly believed to influence carabid movement, had little impact on the beetle activity abundance recorded on Changbai Mountain. Our study indicates that in a relatively large and heterogeneous mature forest area, relationships between plant and carabid diversity are driven by variations in environmental factors linked with altitudinal change. In addition, traditional top-down control theories that are suitable in explaining diversity patterns in ecosystems of low diversity appear to play a much less pronounced role in highly complex forest ecosystems. PMID:24376582

  1. Relationships between plant diversity and the abundance and α-diversity of predatory ground beetles (Coleoptera: Carabidae) in a mature Asian temperate forest ecosystem.

    Science.gov (United States)

    Zou, Yi; Sang, Weiguo; Bai, Fan; Axmacher, Jan Christoph

    2013-01-01

    A positive relationship between plant diversity and both abundance and diversity of predatory arthropods is postulated by the Enemies Hypothesis, a central ecological top-down control hypothesis. It has been supported by experimental studies and investigations of agricultural and grassland ecosystems, while evidence from more complex mature forest ecosystems is limited. Our study was conducted on Changbai Mountain in one of the last remaining large pristine temperate forest environments in China. We used predatory ground beetles (Coleoptera: Carabidae) as target taxon to establish the relationship between phytodiversity and their activity abundance and diversity. Results showed that elevation was the only variable included in both models predicting carabid activity abundance and α-diversity. Shrub diversity was negatively and herb diversity positively correlated with beetle abundance, while shrub diversity was positively correlated with beetle α-diversity. Within the different forest types, a negative relationship between plant diversity and carabid activity abundance was observed, which stands in direct contrast to the Enemies Hypothesis. Furthermore, plant species density did not predict carabid α-diversity. In addition, the density of herbs, which is commonly believed to influence carabid movement, had little impact on the beetle activity abundance recorded on Changbai Mountain. Our study indicates that in a relatively large and heterogeneous mature forest area, relationships between plant and carabid diversity are driven by variations in environmental factors linked with altitudinal change. In addition, traditional top-down control theories that are suitable in explaining diversity patterns in ecosystems of low diversity appear to play a much less pronounced role in highly complex forest ecosystems.

  2. Relationships between plant diversity and the abundance and α-diversity of predatory ground beetles (Coleoptera: Carabidae in a mature Asian temperate forest ecosystem.

    Directory of Open Access Journals (Sweden)

    Yi Zou

    Full Text Available A positive relationship between plant diversity and both abundance and diversity of predatory arthropods is postulated by the Enemies Hypothesis, a central ecological top-down control hypothesis. It has been supported by experimental studies and investigations of agricultural and grassland ecosystems, while evidence from more complex mature forest ecosystems is limited. Our study was conducted on Changbai Mountain in one of the last remaining large pristine temperate forest environments in China. We used predatory ground beetles (Coleoptera: Carabidae as target taxon to establish the relationship between phytodiversity and their activity abundance and diversity. Results showed that elevation was the only variable included in both models predicting carabid activity abundance and α-diversity. Shrub diversity was negatively and herb diversity positively correlated with beetle abundance, while shrub diversity was positively correlated with beetle α-diversity. Within the different forest types, a negative relationship between plant diversity and carabid activity abundance was observed, which stands in direct contrast to the Enemies Hypothesis. Furthermore, plant species density did not predict carabid α-diversity. In addition, the density of herbs, which is commonly believed to influence carabid movement, had little impact on the beetle activity abundance recorded on Changbai Mountain. Our study indicates that in a relatively large and heterogeneous mature forest area, relationships between plant and carabid diversity are driven by variations in environmental factors linked with altitudinal change. In addition, traditional top-down control theories that are suitable in explaining diversity patterns in ecosystems of low diversity appear to play a much less pronounced role in highly complex forest ecosystems.

  3. Phylogeography of a Tertiary relict plant, Meconopsis cambrica (Papaveraceae), implies the existence of northern refugia for a temperate herb.

    Science.gov (United States)

    Valtueña, Francisco J; Preston, Chris D; Kadereit, Joachim W

    2012-03-01

    The perennial herb Meconopsis cambrica, a western European endemic, is the only European species of the otherwise Himalayan genus Meconopsis and has been interpreted as a Tertiary relict species. Using rbcL and ITS sequence variation, we date the split between M. cambrica and its sister clade Papaver s.str. to the Middle to Upper Miocene (12.8 Myr, 6.4-19.2 Myr HPD). Within M. cambrica, cpDNA sequence variation reveals the existence of two groups of populations with a comparable level of genetic variation: a northern group from Great Britain, the Massif Central, the western Pyrenees and the Iberian System, and a southern group from the central and eastern Pyrenees. Populations from the Cantabrian Mountains were placed in both groups. Based on ITS sequence variation, the divergence between these two groups can be dated to 1.5 Myr (0.4-2.8 Myr HPD), and the age of the British populations is estimated as 0.37 Myr (0.0-0.9 Myr HPD). Amplified fragment length polymorphism results confirm the distinctive nature of the populations from Britain, the Massif Central and the central and eastern Pyrenees. These patterns of latitudinal variation of M. cambrica differ from patterns of longitudinal differentiation found in many other temperate species and imply glacial survival of the northern populations in northerly refugia. The primary differentiation into northern and southern cpDNA groups dates to near the onset of the Quaternary and suggests that an ancient phylogeographic pattern has survived through several glacial periods. Our data provide evidence that the species has persisted for a long period with a highly fragmented and probably very localized distribution. © 2012 Blackwell Publishing Ltd.

  4. Litter decomposition rate and soil organic matter quality in a patchwork heathland of Southern Norway

    Science.gov (United States)

    Certini, G.; Vestgarden, L. S.; Forte, C.; Tau Strand, L.

    2014-07-01

    Norwegian heathland soils, although scant and shallow, are major reservoirs of carbon (C). We aimed at assessing whether vegetation cover and, indirectly, its driving factor soil drainage are good proxies for soil organic matter (SOM) composition and dynamics in a typical heathland area of Southern Norway consisting in a patchwork of three different types of vegetation, dominated by Calluna, Molinia, or Sphagnum. Such vegetation covers were clearly associated to microtopographic differences, which in turn dictated differences in soil moisture regime, Calluna growing in the driest sites, Sphagnum in the wettest, and Molinia in sites with intermediate moisture. Litter decomposition was followed over a period of 1 year, by placing litterbags filled with biomass from each dominant species under each type of vegetation cover. The composition of the living biomass, the bulk SOM and some extractable fractions of SOM were investigated by chemical methods and solid-state 13C nuclear magnetic resonance (NMR) spectroscopy. Litter decomposition was faster for Molinia and Calluna, irrespective of the vegetation cover of the site where they were placed. Sphagnum litter decomposed very slowly, especially under Calluna, where the soil environment is by far more oxidising than under itself. In terms of SOM quality, Calluna covered areas showed the greatest differences from the others, in particular a much higher contribution from lipids and aliphatic biopolymers, apparently related to biomass composition. Our findings showed that in the studied environment litter decomposition rate and SOM composition are actually dependent on vegetation cover and/or soil drainage. On this basis, monitoring changes in the patchwork of vegetation types in boreal heathlands could be a reliable cost-effective way to account for modifications in the SOM potential to last induced by climate change.

  5. Contrasting effects of repeated summer drought on soil carbon efflux in hydric and mesic heathland soils

    DEFF Research Database (Denmark)

    Sowerby, Alwyn; Emmett, Bridget A.; Tietema, Albert

    2008-01-01

    storage. To test the importance of drought, and more importantly repeated drought year-on-year, we used automated retractable curtains to exclude rain and produce repeated summer drought in three heathlands at varying moisture conditions. This included a hydric system limited by water-excess (in the UK...... and DK sites during the winter re-wetting period that indicates any change in soil C storage due to changes in soil C efflux may be short lived in these mesic systems. In contrast, in the hydric UK site after 2 years of drought treatment, the persistent reduction in soil moisture throughout the year...

  6. The influence of the annual invasive plant, Impatiens glandulifera, on the sediment dynamics of inland watercourses in temperate regions

    Science.gov (United States)

    Dalvi, Shrutika; Greenwood, Philip

    2016-04-01

    Impatiens glandulifera (Common English Name - Himalayan Balsam) is a non native annual and highly invasive plant that was introduced into parts of Europe from the Himalaya during the nineteenth century as a colourful adornment to parks and gardens. This Plant colonises areas along the river banks, preferably wet, depositional sites, and displaces natural vegetation. The plant is killed by cold weather. The leaves area of riverbank previously occupied by the plant extremely vulnerable to soil erosion until new plant germinates in the following spring. Research work undertaken in the northwest Switzerland and the soutwestern United Kingdom established s link between accelerated soil erosion caused by Impatiens glandulifera and its detrimental impact on native biodiversity of riparian zone of river catchment area. This study focueses on the potential impact of such erosion on sediment quality. A priory reasoning suggests that the preference of Impatiens glandulifera on young depsotional sites near watercourses affects sediment quality. In this study, the results of a soil quality analysis along Impatiens glandulifera-contaminated river banks is presented. Soil physical and chemical properties are compared to non-affected sites to assess the potential impact of preferential erosion on water quality. In addtiion, soil surface profile (SSP) measuring based on by erosion pins, a micro profile bridge and a digital calliper at different selected locations along the riparian zone of river catchment area is used to determine erosion rates and determine sediment transfer from the riparian zone into the rivers.

  7. Long-term and realistic global change manipulations had low impact on diversity of soil biota in temperate heathland

    DEFF Research Database (Denmark)

    Holmstrup, Martin; Damgaard, Christian; Schmidt, Inger K

    2017-01-01

    C:N ratio. After eight years of treatment, however, the CO2-increased litter C:N ratio did not influence the diversity in any of the four fauna groups. The number of significant correlations between treatments, food source quality, and soil biota diversities was reduced from six to three after two...... and eight years, respectively. These results suggest a remarkable resilience within the soil biota against global climate change treatments in the long term....

  8. Long-term and realistic global change manipulations had low impact on diversity of soil biota in temperate heathland

    DEFF Research Database (Denmark)

    Holmstrup, Martin; Damgaard, Christian; Schmidt, Inger K

    2017-01-01

    C:N ratio. After eight years of treatment, however, the CO2-increased litter C:N ratio did not influence the diversity in any of the four fauna groups. The number of significant correlations between treatments, food source quality, and soil biota diversities was reduced from six to three after two...

  9. The paradox of long-term ungulate impact: increase of plant species richness in a temperate forest

    Czech Academy of Sciences Publication Activity Database

    Vild, Ondřej; Hédl, Radim; Kopecký, Martin; Szabó, Péter; Suchánková, Silvie; Zouhar, V.

    2017-01-01

    Roč. 20, č. 2 (2017), s. 282-292 ISSN 1402-2001 R&D Projects: GA AV ČR IAA600050812 EU Projects: European Commission(XE) 278065 - LONGWOOD Institutional support: RVO:67985939 Keywords : plant-herbivore interactions * vegetation resurvey * species richness Subject RIV: EH - Ecology, Behaviour OBOR OECD: Ecology Impact factor: 2.474, year: 2016

  10. Soil solution chemistry and element fluxes in three European heathlands and their responses to warming and drought

    DEFF Research Database (Denmark)

    Schmidt, I.K.; Tietema, A.; Williams, D.

    2004-01-01

    Soil water chemistry and element budgets were studied at three northwestern European Calluna vulgaris heathland sites in Denmark (DK), The Netherlands (NL), and Wales (UK). Responses to experimental nighttime warming and early summer drought were followed during a two-year period. Soil solution...

  11. Investigation into the role of canopy structure traits and plant functional types in modulating the correlation between canopy nitrogen and reflectance in a temperate forest in northeast China

    Science.gov (United States)

    Yu, Quanzhou; Wang, Shaoqiang; Zhou, Lei

    2017-10-01

    A precise estimate of canopy leaf nitrogen concentration (CNC, based on dry mass) is important for researching the carbon assimilation capability of forest ecosystems. Hyperspectral remote sensing technology has been applied to estimate regional CNC, which can adjust forest photosynthetic capacity and carbon uptake. However, the relationship between forest CNC and canopy spectral reflectance as well as its mechanism is still poorly understood. Using measured CNC, canopy structure and species composition data, four vegetation indices (VIs), and near-infrared reflectance (NIR) derived from EO-1 Hyperion imagery, we investigated the role of canopy structure traits and plant functional types (PFTs) in modulating the correlation between CNC and canopy reflectance in a temperate forest in northeast China. A plot-scale forest structure indicator, named broad foliar dominance index (BFDI), was introduced to provide forest canopy structure and coniferous and broadleaf species composition. Then, we revealed the response of forest canopy reflectance spectrum to BFDI and CNC. Our results showed that leaf area index had no significant effect on NIR (P>0.05) but indicated that there was a significant correlation (R2=0.76, P0.05). On the contrary, removing the CNC effect, the partial correlation between BFDI and NIR was positively significant (R=0.69, Pforest types. Nevertheless, the relationship cannot be considered as a feasible approach of CNC estimation for a single PFT.

  12. Modelling nitrogen saturation and carbon accumulation in heathland soils under elevated nitrogen deposition

    International Nuclear Information System (INIS)

    Evans, C.D.; Caporn, S.J.M.; Carroll, J.A.; Pilkington, M.G.; Wilson, D.B.; Ray, N.; Cresswell, N.

    2006-01-01

    A simple model of nitrogen (N) saturation, based on an extension of the biogeochemical model MAGIC, has been tested at two long-running heathland N manipulation experiments. The model simulates N immobilisation as a function of organic soil C/N ratio, but permits a proportion of immobilised N to be accompanied by accumulation of soil carbon (C), slowing the rate of C/N ratio change and subsequent N saturation. The model successfully reproduced observed treatment effects on soil C and N, and inorganic N leaching, for both sites. At the C-rich upland site, N addition led to relatively small reductions in soil C/N, low inorganic N leaching, and a substantial increase in organic soil C. At the C-poor lowland site, soil C/N ratio decreases and N leaching increases were much more dramatic, and soil C accumulation predicted to be smaller. The study suggests that (i) a simple model can effectively simulate observed changes in soil and leachate N; (ii) previous model predictions based on a constant soil C pool may overpredict future N leaching; (iii) N saturation may develop most rapidly in dry, organic-poor, high-decomposition systems; and (iv) N deposition may lead to significantly enhanced soil C sequestration, particularly in wet, nutrient-poor, organic-rich systems. - Enhanced carbon sequestration may slow the rate of nitrogen saturation in heathlands

  13. In Vitro Propagation and Conservation of Inula racemosa Hook. F. an Endangered Medicinal Plant of Temperate Origin

    Directory of Open Access Journals (Sweden)

    R. Kaur

    2010-07-01

    Full Text Available Inula racemosa is an endangered medicinal plant. It is commonly known as Pushkarmool, Pushkar and Manu. The great sage Charaka has characterized it as Hikka magrahana (stops hiccups and Savasahara (helpful in asthma. Also, he has cited it as the best medicament for pleurisy along with cough and asthma (http://enwikipedia.org/wiki/charaka-samhita. Due to the fragile nature of its habitat and exploitation due to its commercial medicinal properties, the species are facing the onslaught of indiscriminate over-exploitation. So far, this plant has not got the required attention from researchers, hence, except for a few efforts, not much work has been done for its cultivation and conservation. Plant tissue culture offers an attractive and quick method for its multiplication and further conservation. In the present investigation, effective procedures for micropropagation and in vitro conservation by vitrification were developed. In vitro propagation using aseptically grown seedlings and in vitro conservation via vitrification were standardized. The in vitro conserved material could be retrieved and multiplied normally on MS (Murashige and Skoog, 1962 medium fortified with 1.00 mg l-1BA (benzyl adenine which has been recorded as the best performing medium for in vitro shoot multiplication. The conserved shoots showed normal in vitro propagation and after retrieval from vitrification, platelets were hardened and successfully established in the experimental fields under Nauni (Solan, HP conditions at an elevation of around 1275 meters above mean sea level.

  14. Salicornia as a crop plant in temperate regions: selection of genetically characterized ecotypes and optimization of their cultivation conditions.

    Science.gov (United States)

    Singh, Devesh; Buhmann, Anne K; Flowers, Tim J; Seal, Charlotte E; Papenbrock, Jutta

    2014-11-10

    Rising sea levels and salinization of groundwater due to global climate change result in fast-dwindling sources of freshwater. Therefore, it is important to find alternatives to grow food crops and vegetables. Halophytes are naturally evolved salt-tolerant plants that are adapted to grow in environments that inhibit the growth of most glycophytic crop plants substantially. Members of the Salicornioideae are promising candidates for saline agriculture due to their high tolerance to salinity. Our aim was to develop genetically characterized lines of Salicornia and Sarcocornia for further breeding and to determine optimal cultivation conditions. To obtain a large and diverse genetic pool, seeds were collected from different countries and ecological conditions. The external transcribed spacer (ETS) sequence of 62 Salicornia and Sarcocornia accessions was analysed: ETS sequence data showed a clear distinction between the two genera and between different Salicornia taxa. However, in some cases the ETS was not sufficiently variable to resolve morphologically distinct species. For the determination of optimal cultivation conditions, experiments on germination, seedling establishment and growth to a harvestable size were performed using different accessions of Salicornia spp. Experiments revealed that the percentage germination was greatest at lower salinities and with temperatures of 20/10 °C (day/night). Salicornia spp. produced more harvestable biomass in hydroponic culture than in sand culture, but the nutrient concentration requires optimization as hydroponically grown plants showed symptoms of stress. Salicornia ramosissima produced more harvestable biomass than Salicornia dolichostachya in artificial sea water containing 257 mM NaCl. Based on preliminary tests on ease of cultivation, gain in biomass, morphology and taste, S. dolichostachya was investigated in more detail, and the optimal salinity for seedling establishment was found to be 100 mM. Harvesting of S

  15. Land-use history affects understorey plant species distributions in a large temperate-forest complex, Denmark

    DEFF Research Database (Denmark)

    Svenning, J.-C.; Baktoft, Karen H.; Balslev, Henrik

    2009-01-01

    In Europe, forests have been strongly influenced by human land-use for millennia. Here, we studied the importance of anthropogenic historical factors as determinants of understorey species distributions in a 967 ha Danish forest complex using 156 randomly placed 100-m2 plots, 15 environmental, 9...... dispersal and a strong literature record as ancient-forest species, were still concentrated in areas that were high forest in 1805. Among the younger forests, there were clear floristic differences between those on reclaimed bogs and those not. Apparently remnant populations of wet-soil plants were still...

  16. Trace element concentrations in leachates and mustard plant tissue (Sinapis alba L.) after biochar application to temperate soils.

    Science.gov (United States)

    Kloss, Stefanie; Zehetner, Franz; Oburger, Eva; Buecker, Jannis; Kitzler, Barbara; Wenzel, Walter W; Wimmer, Bernhard; Soja, Gerhard

    2014-05-15

    Biochar application to agricultural soils has been increasingly promoted worldwide. However, this may be accompanied by unexpected side effects in terms of trace element (TE) behavior. We used a greenhouse pot experiment to study the influence of woodchip-derived biochar (wcBC) on leaching and plant concentration of various TEs (Al, Cd, Cu, Pb, Mn, As, B, Mo, Se). Three different agricultural soils from Austria (Planosol, Cambisol, Chernozem) were treated with wcBC at application rates of 1 and 3% (w/w) and subsequently planted with mustard (Sinapis alba L.). Soil samples were taken 0 and 7 months after the start of the pot experiment, and leachate water was collected twice (days 0 and 54). The extractability (with NH4NO3) of cationic TEs was decreased in the (acidic) Planosol and Cambisol after wcBC application, whereas in the (neutral) Chernozem it hardly changed. In contrast, anionic TEs were mobilized in all three soils, which resulted in higher anion concentrations in the leachates. The application of wcBC had no effect on Al and Pb in the mustard plants, but increased their B and Mo concentrations and decreased their Cd, Cu and Mn concentrations. A two-way analysis of variance showed significant interactions between wcBC application rate and soil type for most TEs, which indicates that different soil types may react differently upon wcBC application. Correlation and partial correlation analyses revealed that TE behavior was primarily related to soil pH, whereas the involvement of other factors such as electrical conductivity (EC), organic carbon (OC) content and dissolved organic carbon (DOC) was found to be more soil and TE-specific. The application of wcBC may be a useful strategy for the remediation of soils with elevated levels of cationic TEs, but could lead to deficiencies of cationic micronutrients and enhance short-term translocation of anionic TEs towards the groundwater at high leaching rates. Copyright © 2014 Elsevier B.V. All rights reserved.

  17. The Need for Temperance

    Directory of Open Access Journals (Sweden)

    Karl Inge Tangen

    2015-11-01

    Full Text Available This article explores how temperance as a virtue relates to organizational leadership. The study begins with a short survey of classical Greek and Christian notions of temperance before proceeding to ex-plore temperance in relation to self-leadership, visionary and strategic leadership, and relational lead-ership. The final part of the article offers reflections on how temperance might be cultivated from a theological perspective. Temperance is understood not only as sound thinking but also as embodied self-control and active patience. On the level of self-leadership, it is argued that temperance enables the leader to establish forms of integrity that protect the leader’s self from chaos and destruction. Moreover, temperance may also nurture focused visionary leadership that accepts ethical limits and has an eye to the common good. The study also suggests that organizations should cultivate a culture of strategic discipline that is capable of realizing such visions. On the interpersonal level, temperance is viewed as critical in terms of enabling leaders to treat co-workers with respect and wisdom and han-dle conflict with consideration. Finally, is argued that that the cultivation of temperance is not a one-way street from the inside to the outside or a subordination of feelings to reason but rather a very complex process that includes interpersonal humility, finds vision in an encounter with the good, and yet remains a personal responsibility.

  18. Combined climate factors alleviate changes in gross soil nitrogen dynamics in heathlands

    DEFF Research Database (Denmark)

    Bjorsne, Anna-Karin; Rutting, Tobias; Ambus, Per

    2014-01-01

    of exposure to three climate change factors, i.e. warming, elevated CO2 (eCO(2)) and summer drought, applied both in isolation and in combination. By conducting laboratory N-15 tracing experiments we show that warming increased both gross N mineralization and nitrification rates. In contrast, gross......The ongoing climate change affects biogeochemical cycling in terrestrial ecosystems, but the magnitude and direction of this impact is yet unclear. To shed further light on the climate change impact, we investigated alterations in the soil nitrogen (N) cycling in a Danish heathland after 5 years......CO(2). In the full treatment combination, simulating the predicted climate for the year 2075, gross N transformations were only moderately affected compared to control, suggesting a minor alteration of the N cycle due to climate change. Overall, our study confirms the importance of multifactorial field...

  19. Habitat Fragmentation Intensifies Trade-Offs between Biodiversity and Ecosystem Services in a Heathland Ecosystem in Southern England.

    Science.gov (United States)

    Cordingley, Justine E; Newton, Adrian C; Rose, Robert J; Clarke, Ralph T; Bullock, James M

    2015-01-01

    While habitat fragmentation represents a major threat to global biodiversity, its impacts on provision of ecosystem services are largely unknown. We analysed biodiversity value and provision of multiple ecosystem services in 110 fragments of lowland heathland ecosystems in southern England, in which vegetation dynamics have been monitored for over 30 years. Decreasing fragment size was found to be associated with a decrease in biodiversity and recreational values, but an increase in relative carbon storage, aesthetic value and timber value. The trade-off between either biodiversity or recreational values with the other ecosystem services therefore became more pronounced as heathland size decreased. This was attributed to a higher rate of woody succession in smaller heathland fragments over the past three decades, and contrasting values of different successional vegetation types for biodiversity and provision of ecosystem services. These results suggest that habitat fragmentation can reduce the potential for developing "win win" solutions that contribute to biodiversity conservation while also supporting socio-economic development. Approaches to multi-purpose management of fragmented landscapes should therefore consider the potential trade-offs in ecosystem services and biodiversity associated with fragmentation, in order to make an effective contribution to sustainable development.

  20. Effects of restoration measures on plant communities of wet heathland ecosystems

    NARCIS (Netherlands)

    Jansen, AJM; Fresco, LFM; Grootjans, AP; Jalink, Mark H.; Rapson, G.

    2004-01-01

    Question: Which are the success and failure of restoration measures, particularly sod-cutting and hydrological measures, in small wetlands on mineral soils in The Netherlands. Location: Twente. in the eastern part of The Netherlands. Methods: Success or failure of restoration measures has been

  1. Effects of restoration measures on plant communities of wet heathland ecosystems

    NARCIS (Netherlands)

    Jansen, AJM; Fresco, LFM; Grootjans, AP; Jalink, Mark H.; Rapson, G.

    Question: Which are the success and failure of restoration measures, particularly sod-cutting and hydrological measures, in small wetlands on mineral soils in The Netherlands. Location: Twente. in the eastern part of The Netherlands. Methods: Success or failure of restoration measures has been

  2. Effects of restoration measures on plant communities of wet heathland ecosystems

    NARCIS (Netherlands)

    Jansen, A.J.M.; Fresco, L.F.M.; Grootjans, A.P.; Jalink, M.H.

    2004-01-01

    Question: Which are the success and failure of restoration measures, particularly sod-cutting and hydrological measures, in small wetlands on mineral soils in The Netherlands. Location: Twente, in the eastern part of The Netherlands. Methods: Success or failure of restoration measures has been

  3. Combining remote sensing and on-site monitoring methods to investigate footpath erosion within a popular recreational heathland environment.

    Science.gov (United States)

    Rodway-Dyer, Sue; Ellis, Nicola

    2018-06-01

    Footpaths are a prominent consequence of natural area tourism and reflect damage caused to valuable, sensitive habitats by people pressure. Degradation impacts on vegetation, wildlife, on and off-site soil movement and loss, creation of additional informal off-path footpaths (desire lines), and visual destruction of landscapes. Impacts need to be measured and monitored on a large temporal and spatial scale to aid in land management to maintain access and preserve natural environments. This study combined remote sensing (Light Detection and Ranging [LiDAR] and aerial photography) with on-site measurement of footpaths within a sensitive heathland habitat (Land's End, Cornwall, UK). Soil loss, slope angle change, vegetation damage and a hydrology model were combined to comprehensively study the site. Results showed 0.09 m mean soil loss over five years, footpath widening, increasing grass cover into heathland, and water channelling on the footpaths exacerbating erosion. The environments surrounding the footpaths were affected with visitors walking off path, requiring further management and monitoring. Multiple remote sensing techniques were highly successful in comprehensively assessing the area, particularly the hydrology model, demonstrating the potential of providing a valuable objective and quantitative monitoring and management tool. Copyright © 2018 Elsevier Ltd. All rights reserved.

  4. The effect of lichen-dominated biological soil crusts on growth and physiological characteristics of three plant species in a temperate desert of northwest China.

    Science.gov (United States)

    Zhuang, W W; Serpe, M; Zhang, Y M

    2015-11-01

    Biocrusts (biological soil crusts) cover open spaces between vascular plants in most arid and semi-arid areas. Information on effects of biocrusts on seedling growth is controversial, and there is little information on their effects on plant growth and physiology. We examined impacts of biocrusts on growth and physiological characteristics of three habitat-typical plants, Erodium oxyrhynchum, Alyssum linifolium and Hyalea pulchella, growing in the Gurbantunggut Desert, northwest China. The influence of biocrusts on plant biomass, leaf area, leaf relative water content, photosynthesis, maximum quantum efficiency of PSII (F(v)/F(m)), chlorophyll, osmotic solutes (soluble sugars, protein, proline) and antioxidant enzymes (superoxide dismutase, catalase, peroxidase) was investigated on sites with or without biocrust cover. Biomass, leaf area, leaf water content, photosynthesis, F(v)/F(m) and chlorophyll content in crusted soils were higher than in uncrusted soils during early growth and lower later in the growth period. Soluble sugars, proline and antioxidant enzyme activity were always higher in crusted than in uncrusted soils, while soluble protein content was always lower. These findings indicate that biocrusts have different effects on these three ephemeral species during growth in this desert, primarily via effects on soil moisture, and possibly on soil nutrients. The influence of biocrusts changes during plant development: in early plant growth, biocrusts had either positive or no effect on growth and physiological parameters. However, biocrusts tended to negatively influence plants during later growth. Our results provide insights to explain why previous studies have found different effects of biocrusts on vascular plant growth. © 2015 German Botanical Society and The Royal Botanical Society of the Netherlands.

  5. Soil microarthropods are only weakly impacted after 13 years of repeated drought treatment in wet and dry heathland soils

    DEFF Research Database (Denmark)

    Holmstrup, Martin; Sørensen, Jesper G.; Schmidt, Inger Kappel

    2013-01-01

    Studies of biological responses in the terrestrial environment to rapid changes in climate have mostly been concerned with aboveground biota, whereas less is known of belowground organisms. The present study focuses on mites and springtails of heathland ecosystems and how the microarthropod...... and temperature. This approach provided an opportunity to study biological responses on a local (within sites) and regional scale. Warming treatments increasing night time temperature (0.3–1 °C higher than ambient at 5 cm soil depth) had no detectable effects on the microarthropod communities. Increased intensity...... and frequency of drought had only weak persistent effects on springtail species composition, but practically no effect on major mite groups (Oribatida, Prostigmata or Mesostigmata) suggesting that ecosystem functions of microarthropods may only be transiently impacted by repeated spring or summer drought....

  6. The effect of wildfire on population dynamics for two native small mammal species in a coastal heathland in Queensland, Australia

    Science.gov (United States)

    Liedloff, Adam C.; Wilson, John C.; Engeman, Richard M.

    2018-04-01

    The influences of wildfire through population dynamics and life history for two species of small mammals in a south-east Queensland heathland on Bribie Island are presented. Trapping results provided information on breeding, immigration and movement of Melomys burtoni (Grassland melomys) and Rattus lutreolus (Swamp rat). We first investigated and optimized the design of trapping methodology for producing mark-recapture population estimates to compare two adjacent populations, one of which was subjected to an extensive wildfire during the two year study. We consider how well rodents survive wildfire and whether the immediate impacts of fire or altered habitat have the greatest impact on each species. We found the R. lutreolus population was far more influenced by the fire than the M. burtoni population both immediately after the fire and over 18 months of vegetation recovery.

  7. Soil environmental conditions and microbial build-up mediate the effect of plant diversity on soil nitrifying and denitrifying enzyme activities in temperate grasslands.

    Directory of Open Access Journals (Sweden)

    Xavier Le Roux

    Full Text Available Random reductions in plant diversity can affect ecosystem functioning, but it is still unclear which components of plant diversity (species number - namely richness, presence of particular plant functional groups, or particular combinations of these and associated biotic and abiotic drivers explain the observed relationships, particularly for soil processes. We assembled grassland communities including 1 to 16 plant species with a factorial separation of the effects of richness and functional group composition to analyze how plant diversity components influence soil nitrifying and denitrifying enzyme activities (NEA and DEA, respectively, the abundance of nitrifiers (bacterial and archaeal amoA gene number and denitrifiers (nirK, nirS and nosZ gene number, and key soil environmental conditions. Plant diversity effects were largely due to differences in functional group composition between communities of identical richness (number of sown species, though richness also had an effect per se. NEA was positively related to the percentage of legumes in terms of sown species number, the additional effect of richness at any given legume percentage being negative. DEA was higher in plots with legumes, decreased with increasing percentage of grasses, and increased with richness. No correlation was observed between DEA and denitrifier abundance. NEA increased with the abundance of ammonia oxidizing bacteria. The effect of richness on NEA was entirely due to the build-up of nitrifying organisms, while legume effect was partly linked to modified ammonium availability and nitrifier abundance. Richness effect on DEA was entirely due to changes in soil moisture, while the effects of legumes and grasses were partly due to modified nitrate availability, which influenced the specific activity of denitrifiers. These results suggest that plant diversity-induced changes in microbial specific activity are important for facultative activities such as denitrification

  8. From Leaf Synthesis to Senescence: n-Alkyl Lipid Abundance and D/H Composition Among Plant Species in a Temperate Deciduous Forest at Brown's Lake Bog, Ohio, USA

    Science.gov (United States)

    Freimuth, E. J.; Diefendorf, A. F.; Lowell, T. V.

    2014-12-01

    The hydrogen isotope composition (D/H, δD) of terrestrial plant leaf waxes is a promising paleohydrology proxy because meteoric water (e.g., precipitation) is the primary hydrogen source for wax synthesis. However, secondary environmental and biological factors modify the net apparent fractionation between precipitation δD and leaf wax δD, limiting quantitative reconstruction of paleohydrology. These secondary factors include soil evaporation, leaf transpiration, biosynthetic fractionation, and the seasonal timing of lipid synthesis. Here, we investigate the influence of each of these factors on n-alkyl lipid δD in five dominant deciduous angiosperm tree species as well as shrubs, ferns and grasses in the watershed surrounding Brown's Lake Bog, Ohio, USA. We quantified n-alkane and n-alkanoic acid concentrations and δD in replicate individuals of each species at weekly to monthly intervals from March to October 2014 to assess inter- and intraspecific isotope variability throughout the growing season. We present soil, xylem and leaf water δD from each individual, and precipitation and atmospheric water vapor δD throughout the season to directly examine the relationship between source water and lipid isotope composition. These data allow us to assess the relative influence of soil evaporation and leaf transpiration among plant types, within species, and along a soil moisture gradient throughout the catchment. We use leaf water δD to approximate biosynthetic fractionation for each individual and test whether this is a species-specific and seasonal constant, and to evaluate variation among plant types with identical growth conditions. Our high frequency sampling approach provides new insights into the seasonal timing of n-alkane and n-alkanoic acid synthesis and subsequent fluctuations in concentration and δD in a temperate deciduous forest. These results will advance understanding of the magnitude and timing of secondary influences on the modern leaf wax

  9. Potential climatic refugia in semi-arid, temperate mountains: plant and arthropod assemblages associated with rock glaciers, talus slopes, and their forefield wetlands, Sierra Nevada, California, USA

    Science.gov (United States)

    Constance I. Millar; Robert D. Westfall; Angela Evenden; Jeffrey G. Holmquist; Jutta Schmidt-Gengenbach; Rebecca S. Franklin; Jan Nachlinger; Diane L. Delany

    2015-01-01

    Unique thermal and hydrologic regimes of rock-glacier and periglacial talus environments support little-studied mountain ecosystems. We report the first studies of vascular plant and arthropod diversity for these habitats in the central Sierra Nevada, California, USA. Surfaces of active rock glaciers develop scattered islands of soil that provide habitat for vegetation...

  10. In situ 13CO2 pulse‐labeling in a temperate heathland – development of a mobile multi‐plot field setup

    DEFF Research Database (Denmark)

    Reinsch, Sabine; Ambus, Per

    2013-01-01

    Pulse‐labeling with 13CO2 and the subsequent analysis of 13C‐carbon via isotope ratio mass spectrometry (IRMS) have been shown to be an excellent method to investigate the terrestrial carbon cycle. Improving 13CO2 manipulation experiments will facilitate our understanding of carbon cycling...

  11. Bacteria and fungi respond differently to multifactorial climate change in a temperate heathland, traced with 13C-Glycine and FACE CO2

    DEFF Research Database (Denmark)

    Andresen, Louise C.; Dungait, Jennifer A.J.; Bol, Roland

    2014-01-01

    PLFAs for specific microbial groups (Gram positive bacteria, Gram negative bacteria, actinobacteria and fungi) was quantified using gas chromatography-combustion-stable isotope ratio mass spectrometry (GC-C-IRMS). Gram positive bacteria opportunistically utilized the freshly added glycine substrate, i...

  12. Short- and medium-term effects of experimental nitrogen fertilization on arthropods associated with Calluna vulgaris heathlands in north-west Spain

    International Nuclear Information System (INIS)

    Cuesta, D.; Taboada, A.; Calvo, L.; Salgado, J.M.

    2008-01-01

    We studied the short- and medium-term effects of experimental nitrogen fertilization (3 and 15 months after the treatment) on the arthropods of Calluna vulgaris heathlands in NW Spain. Three heathland sites were selected with two permanent plots per site: control and fertilized. Ammonium nitrate fertilizer (56 kg N ha -1 yr -1 ) was applied monthly and insects were caught using pitfall traps. We found mainly species-level responses to nitrogen addition. Seven species (e.g. Lochmaea suturalis) showed a consistent trend (benefited or harmed) in both periods and were proposed as possible reliable indicators of the effects of nitrogen deposition in these ecosystems. We also found variable arthropod trophic-group responses: (a) herbivores (leaf beetles, true bugs) increased in abundance on a short-term scale; (b) predators (carabid beetles, true bugs) showed opposite and less clear responses in both periods. Further long-term studies are needed to determine the mechanisms underlying the observed arthropod responses. - We observed consistent species-level and variable trophic-group responses to nitrogen addition in one of the southern-most locations for Calluna vulgaris heathlands within Europe

  13. World temperate fruit production: characteristics and challenges

    Directory of Open Access Journals (Sweden)

    Jorge B. Retamales

    2011-10-01

    Full Text Available In the last 30 years world population has increased 70% but per capita global fruit consumption is only 20% higher. Even though tropical and temperate fruit have similar contributions to the 50 kg/person/year of US consumption of fresh fruit, in the last 30 years this has been slightly greater for temperate fruit. Within fruit consumption, the largest expansion has been for organic fruit which increased more than 50% in the 2002-2006 period. The largest expansion of area planted in the 1996-2006 has been for kiwi (29% and blueberries (20%, while apples (-24% and sour cherries (-13% have had the largest reductions. Nearly 50% of the total global volume of fruit is produced by 5 countries: China, USA, Brazil, Italy and Spain. The main producer (China accounts for 23% of the total. While the main exporters are Spain, USA and Italy, the main importers are Germany, Russia and UK. Demands for the industry have evolved towards quality, food safety and traceability. The industry faces higher productions costs (labor, energy, agrichemicals. The retailers are moving towards consolidation while the customers are changing preferences (food for health. In this context there is greater pressure on growers, processors and retailers. Emerging issues are labor supply, climate change, water availability and sustainability. Recent developments in precision agriculture, molecular biology, phenomics, crop modelling and post harvest physiology should increase yields and quality, and reduce costs for temperate fruit production around the world.

  14. Historia natural y conservación de los mutualismos planta-animal del bosque templado de Sudamérica austral Natural history and conservation of plant-animal mutualisms in the temperate forest of southern South America

    Directory of Open Access Journals (Sweden)

    MARCELO A. AIZEN

    2002-03-01

    Full Text Available El bosque templado de Sudamérica austral (BTSA se caracteriza por poseer un biota altamente endémica y una flora que exhibe una de las más altas incidencias de polinización biótica (particularmente ornitofilia y dispersión animal (particularmente endozoocoria registradas para cualquier bioma templado. A la luz de importantes avances realizados en los últimos años en el conocimiento de estas interacciones, en este trabajo realizamos una revisión sobre (1 los grupos animales que intervienen en estos mutualismos y su importancia relativa, (2 la estructura comunitaria de estas interacciones, y (3 la existencia de variación geográfica en estas relaciones en base a estudios similares realizados en dos o más localidades a ambos lados de los Andes. También comparamos los mutualismos entre plantas y animales del BTSA con los presentes en otros biomas. Por último, reseñamos la evidencia existente sobre los efectos que la deforestación, la fragmentación, y otras formas de perturbación antrópica están teniendo sobre estos mutualismos. Este trabajo avala la hipótesis de que la alta riqueza de plantas que son polinizadas o dispersadas por vertebrados depende de un ensamble extremadamente pobre de animales mutualistas. Por el contrario, en el caso de la polinización por invertebrados este trabajo revela que las flores entomófilas del BTSA interactúan con un ensamble insospechadamente diverso de insectos, rico en grupos taxonómicos exclusivos de este bioma. Aunque muchos de estos mutualismos parecen ser resilientes, otros aparentan tener un alto grado de sensibilidad a distintas formas de perturbación antrópicaThe temperate forest of southern South America (TFSSA has a highly endemic biota with a flora that exhibits one of the highest incidences of biotic pollination (particularly ornithophily and animal dispersal (particularly endozoochory found in any temperate biome worldwide. Much knowledge has been gained in the last few years on

  15. Emissions of biogenic volatile organic compounds and subsequent photochemical production of secondary organic aerosol in mesocosm studies of temperate and tropical plant species

    Science.gov (United States)

    Wyche, K. P.; Ryan, A. C.; Hewitt, C. N.; Alfarra, M. R.; McFiggans, G.; Carr, T.; Monks, P. S.; Smallbone, K. L.; Capes, G.; Hamilton, J. F.; Pugh, T. A. M.; MacKenzie, A. R.

    2014-12-01

    Silver birch (Betula pendula) and three Southeast Asian tropical plant species (Ficus cyathistipula, Ficus benjamina and Caryota millis) from the pantropical fig and palm genera were grown in a purpose-built and environment-controlled whole-tree chamber. The volatile organic compounds emitted from these trees were characterised and fed into a linked photochemical reaction chamber where they underwent photo-oxidation under a range of controlled conditions (relative humidity or RH ~65-89%, volatile organic compound-to-NOx or VOC / NOx ~3-9 and NOx ~2 ppbV). Both the gas phase and the aerosol phase of the reaction chamber were monitored in detail using a comprehensive suite of on-line and off-line chemical and physical measurement techniques. Silver birch was found to be a high monoterpene and sesquiterpene but low isoprene emitter, and its emissions were observed to produce measurable amounts of secondary organic aerosol (SOA) via both nucleation and condensation onto pre-existing seed aerosol (YSOA 26-39%). In contrast, all three tropical species were found to be high isoprene emitters with trace emissions of monoterpenes and sesquiterpenes. In tropical plant experiments without seed aerosol there was no measurable SOA nucleation, but aerosol mass was shown to increase when seed aerosol was present. Although principally isoprene emitting, the aerosol mass produced from tropical fig was mostly consistent (i.e. in 78 out of 120 aerosol mass calculations using plausible parameter sets of various precursor specific yields) with condensation of photo-oxidation products of the minor volatile organic compounds (VOCs) co-emitted; no significant aerosol yield from condensation of isoprene oxidation products was required in the interpretations of the experimental results. This finding is in line with previous reports of organic aerosol loadings consistent with production from minor biogenic VOCs co-emitted with isoprene in principally isoprene-emitting landscapes in Southeast

  16. Burned Area Detection and Burn Severity Assessment of a Heathland Fire in Belgium Using Airborne Imaging Spectroscopy (APEX

    Directory of Open Access Journals (Sweden)

    Lennert Schepers

    2014-02-01

    Full Text Available Uncontrolled, large fires are a major threat to the biodiversity of protected heath landscapes. The severity of the fire is an important factor influencing vegetation recovery. We used airborne imaging spectroscopy data from the Airborne Prism Experiment (APEX sensor to: (1 investigate which spectral regions and spectral indices perform best in discriminating burned from unburned areas; and (2 assess the burn severity of a recent fire in the Kalmthoutse Heide, a heathland area in Belgium. A separability index was used to estimate the effectiveness of individual bands and spectral indices to discriminate between burned and unburned land. For the burn severity analysis, a modified version of the Geometrically structured Composite Burn Index (GeoCBI was developed for the field data collection. The field data were collected in four different vegetation types: Calluna vulgaris-dominated heath (dry heath, Erica tetralix-dominated heath (wet heath, Molinia caerulea (grass-encroached heath, and coniferous woodland. Discrimination between burned and unburned areas differed among vegetation types. For the pooled dataset, bands in the near infrared (NIR spectral region demonstrated the highest discriminatory power, followed by short wave infrared (SWIR bands. Visible wavelengths performed considerably poorer. The Normalized Burn Ratio (NBR outperformed the other spectral indices and the individual spectral bands in discriminating between burned and unburned areas. For the burn severity assessment, all spectral bands and indices showed low correlations with the field data GeoCBI, when data of all pre-fire vegetation types were pooled (R2 maximum 0.41. Analysis per vegetation type, however, revealed considerably higher correlations (R2 up to 0.78. The Mid Infrared Burn Index (MIRBI had the highest correlations for Molinia and Erica (R2 = 0.78 and 0.42, respectively. In Calluna stands, the Char Soil Index (CSI achieved the highest correlations, with R2 = 0

  17. The counteracting effects of elevated atmospheric CO2 concentrations and drought episodes: Studies of enchytraeid communities in a dry heathland

    DEFF Research Database (Denmark)

    Maraldo, Kristine; Krogh, Paul Henning; Linden, Leon

    2010-01-01

    The potential impacts of interactions of multiple climate change factors in soil ecosystems have received little attention. Most studies have addressed effects of single factors such as increased temperature or atmospheric CO2 but little is known about how such environmental factors will interact...... impact of drought on the enchytraeids, compared to the year with a wet summer and autumn (2007). Our study emphasises the importance of multi-factorial experimental design as a means to investigate effects of climatic changes.......The potential impacts of interactions of multiple climate change factors in soil ecosystems have received little attention. Most studies have addressed effects of single factors such as increased temperature or atmospheric CO2 but little is known about how such environmental factors will interact....... In the present study we investigate the effects of in situ exposure to elevated atmospheric CO2 concentration, increased temperatures and prolonged drought episodes on field communities of Enchytraeidae (Oligochaeta) in a dry heathland (Brandbjerg, Denmark). Increased CO2 had a positive effect on enchytraeid...

  18. Heathlands, fire and grazing. A palaeoenvironmental view of Las Hurdes (Cáceres, Spain history during the last 1200 years

    Directory of Open Access Journals (Sweden)

    Daniel Abel-Schaad

    2014-08-01

    Full Text Available Aim of study. The diachronic study of vegetation change through palynological analysis of sedimentary deposits is an essential tool both to design sound strategies on landscape  management and to understand its anthropogenic dynamics.Area of study. La Meseguera mire (Ladrillar, Cáceres, Spain is located in the Hurdes region in the western part of Iberian Central System and started to develop at the beginning of the Islamic period (ca. 770 cal AD, in an area widely dominated by heathland.  Material and methods. Pollen, non-pollen palynomorphs and charcoal accumulation rate (CHAR combined with historical data are useful indicators to assess the increasing role of human influence on vegetation.Main results. The use of fire and livestock husbandry represents the main drivers of landscape change in the course of the history. The establishment of forest afforestation plans, from the middle of 20th century, changed substantially the regional features. The sporadic presence of beech pollen is detected until 16th century, which implies the most western location in the Iberian Central Mountain System.Research highlights. The integration of pollen analysis and historical data is an essential tool when studying the changes in Holocene vegetation. These changes have been mainly driven by anthropogenic disturbances, more specifically by fire and livestock husbandry.Key Words: Anthropogenic dynamics; Central Mountain System; microcharcoals; non-pollen palynomorphs.  

  19. Time-temperature equivalence in Martensite tempering

    Energy Technology Data Exchange (ETDEWEB)

    Hackenberg, Robert E. [Los Alamos National Laboratory; Thomas, Grant A. [CSM; Speer, John G. [CSM; Matlock, David K. [CSM; Krauss, George [CSM

    2008-06-16

    The relationship between time and temperature is of great consequence in many materials-related processes including the tempering of martensite. In 1945, Hollomon and Jaffe quantified the 'degree of tempering' as a function of both tempering time, t, and tempering temperature, T, using the expression, T(log t + c). Here, c is thought to be a material constant and appears to decrease linearly with increasing carbon content. The Hollomon-Jaffe tempering parameter is frequently cited in the literature. This work reviews the original derivation of the tempering parameter concept, and presents the use of the characteristics diffusion distance as an alternative time-temperature relationship during martensite tempering. During the tempering of martensite, interstitial carbon atoms diffuse to form carbides. In addition, austenite decomposes, dislocations and grain boundaries rearrange, associated with iron self diffusion. Since these are all diffusional processes, it is reasonable to expect the degree of tempering to relate to the extent of diffusion.

  20. Tempered stable laws as random walk limits

    OpenAIRE

    Chakrabarty, Arijit; Meerschaert, Mark M.

    2010-01-01

    Stable laws can be tempered by modifying the L\\'evy measure to cool the probability of large jumps. Tempered stable laws retain their signature power law behavior at infinity, and infinite divisibility. This paper develops random walk models that converge to a tempered stable law under a triangular array scheme. Since tempered stable laws and processes are useful in statistical physics, these random walk models can provide a basic physical model for the underlying physical phenomena.

  1. Belowground heathland responses after 2 years of combined warming, elevated CO2 and summer drought

    DEFF Research Database (Denmark)

    Andresen, Louise C.; Michelsen, Anders; Ambus, Per

    2010-01-01

    significantly to the treatments. In the combined temperature and CO2 treatment the dissolved organic nitrogen concentration decreased and the ammonium concentration increased, but this release of nutrients was not mirrored by plant parameters. Microbial biomass carbon and microbial enrichment with 13C and 15N...

  2. Dams release methane even in temperate zoned

    International Nuclear Information System (INIS)

    Lemarchand, F.

    2010-01-01

    The Wohlen lake (near Bern) is a retaining dam built 90 years ago that has undergone a campaign to measure the quantity of methane released. The campaign lasted 1 year and the result was unexpected: 0.15 g/m 2 *day which one of the highest release rates in temperate zones. This result is all the more stunning since water stays only 2 days in average in the reservoir and that the drowned area is not important. In fact the river Aar that feeds the lake is loaded with organic matter coming from humane activities: agriculture and 3 sewage plants. This organic matter decays in the lake releasing methane. (A.C.)

  3. Ecosystem-atmosphere exchange of carbon in a heathland under future climatic conditions

    DEFF Research Database (Denmark)

    Selsted, Merete Bang

    understanding plant and soil responses to such changes are necessary, as ecosystems potentially can ameliorate or accelerate global change. To predict the feedback of ecosystems to the atmospheric CO2 concentrations experiments imitating global change effects are therefore an important tool. This work....... Fluxes of CO2 from soil to atmosphere depend on a physical equilibrium between those two medias, why it is important to keep the CO2 gradient between soil and atmosphere unchanged during measurement. Uptake to plants via photosynthesis depends on a physiological process, which depends strongly...... on the atmospheric CO2 concentration. Photosynthesis and respiration run in parallel during measurements of net ecosystem exchange, and these measurements should therefore be performed with care to both the atmospheric CO2 concentration and the CO2 soil-atmosphere gradient....

  4. Fossil evidence for open, Proteaceae-dominated heathlands and fire in the Late Cretaceous of Australia.

    Science.gov (United States)

    Carpenter, Raymond J; Macphail, Michael K; Jordan, Gregory J; Hill, Robert S

    2015-12-01

    The origin of biomes is of great interest globally. Molecular phylogenetic and pollen evidence suggest that several plant lineages that now characterize open, burnt habitats of the sclerophyll biome, became established during the Late Cretaceous of Australia. However, whether this biome itself dates to that time is problematic, fundamentally because of the near-absence of relevant, appropriately aged, terrestrial plant macro- or mesofossils. We recovered, identified, and interpreted the ecological significance of fossil pollen, foliar and other remains from a section of core drilled in central Australia, which we dated as Late Campanian-Maastrichtian. The sediments contain plant fossils that indicate nutrient-limited, open, sclerophyllous vegetation and abundant charcoal as evidence of fire. Most interestingly, >30 pollen taxa and at least 12 foliage taxa are attributable to the important Gondwanan family Proteaceae, including several minute, amphistomatic, and sclerophyllous foliage forms consistent with subfamily Proteoideae. Microfossils, including an abundance of Sphagnales and other wetland taxa, provided strong evidence of a fenland setting. The local vegetation also included diverse Ericaceae and Liliales, as well as a range of ferns and gymnosperms. The fossils provide strong evidence in support of hypotheses of great antiquity for fire and open vegetation in Australia, point to extraordinary persistence of Proteaceae that are now emblematic of the Mediterranean-type climate southwestern Australian biodiversity hotspot and raise the profile of open habitats as centers of ancient lineages. © 2015 Botanical Society of America.

  5. Adapting a Natura 2000 field guideline for a remote sensing-based assessment of heathland conservation status

    Science.gov (United States)

    Schmidt, Johannes; Fassnacht, Fabian Ewald; Neff, Christophe; Lausch, Angela; Kleinschmit, Birgit; Förster, Michael; Schmidtlein, Sebastian

    2017-08-01

    Remote sensing can be a valuable tool for supporting nature conservation monitoring systems. However, for many areas of conservation interest, there is still a considerable gap between field-based operational monitoring guidelines and the current remote sensing-based approaches. This hampers application in practice of the latter. Here, we propose a remote sensing approach for mapping the conservation status of Calluna-dominated Natura 2000 dwarf shrub habitats that is closely related to field mapping schemes. We transferred the evaluation criteria of the field guidelines to three related variables that can be captured by remote sensing: (1) coverage of the key species, (2) stand structural diversity, and (3) co-occurring species. Continuous information on these variables was obtained by regressing ground reference data from field surveys and UAV flights against airborne hyperspectral imagery. Merging the three resulting quality layers in an RGB representation allowed for illustrating the habitat quality in a continuous way. User-defined thresholds can be applied to this stack of quality layers to derive an overall assessment of habitat quality in terms of nature conservation, i.e. the conservation status. In our study, we found good accordance of the remotely sensed data with field-based information for the three variables key species, stand structural diversity and co-occurring vegetation (R2 of 0.79, 0.69, and 0.71, respectively) and it was possible to derive meaningful habitat quality maps. The conservation status could be derived with an accuracy of 65%. In interpreting these results it should be considered that the remote sensing based layers are independent estimates of habitat quality in their own right and not a mere replacement of the criteria used in the field guidelines. The approach is thought to be transferable to similar regions with minor adaptions. Our results refer to Calluna heathland which we consider a comparably easy target for remote sensing

  6. Ecological anachronisms in the recruitment of temperate light-demanding tree species in wooded pastures

    NARCIS (Netherlands)

    Bakker, ES; Olff, H; Vandenberghe, C; De Maeyer, K; Smit, R; Gleichman, JM; Vera, FWM

    1. Light-demanding trees and thorny shrubs in temperate plant communities may reflect adaptations to now-extinct large grazers, such as aurochs and tarpans, rendering these adaptations ecological anachronisms. 2. We explored the ecological functions of plant traits of Quercus robur and Prunus

  7. The effects of UV-B radiation on European heathland species

    International Nuclear Information System (INIS)

    Björn, L.O.; Callaghan, T.V.; Johnsen, I.; Lee, J.A.; Manetas, Y.; Paul, N.D.; Sonesson, M.; Wellburn, A.R.; Coop, D.; Heide-Jørgensen, H.S.; Gehrke, C.; Gwynn-Jones, D.; Johanson, U.; Kyparissis, A.; Levizou, E.; Nikolopoulos, D.; Petropoulou, Y.; Stephanou, M.

    1997-01-01

    The effects of enhanced UV-B radiation on three examples of European shrub-dominated vegetation were studied in situ. The experiments were in High Arctic Greenland, northern Sweden and Greece, and at all sites investigated the interaction of enhanced UV-B radiation (simulating a 15% reduction in the ozone layer) with artificially increased precipitation. The Swedish experiment also involved a study of the interaction between enhanced UV-B radiation and elevated CO 2 (600 ppm). These field studies were supported by an outdoor controlled environment study in the United Kingdom involving modulated enhancement of UV-B radiation in combination with elevated CO 2 (700 ppm). Effects of the treatments on plant growth, morphology, phenology and physiology were measured. The effects observed were species specific, and included both positive and negative responses to the treatments. In general the negative responses to UV-B treatments of up to three growing seasons were small, but included reductions in shoot growth and premature leaf senescence. Positive responses included a marked increase in flowering in some species and a stimulation of some photosynthetic processes. UV-B treatment enhanced the drought tolerance of Pinus pinea and Pinus halepensis by increasing leaf cuticle thickness. In general, there were few interactions between the elevated CO 2 and enhanced UV-B treatments. There was evidence to suggest that although the negative responses to the treatments were small, damage may be increasing with time in some long-lived woody perennials. There was also evidence in the third year of treatments for effects of UV-B on insect herbivory in Vaccinium species. The experiments point to the necessity for long-term field investigations to predict the likely ecological consequences of increasing UV-B radiation. (author)

  8. Responses of enchytraeids to increased temperature, drought and atmospheric CO2: Results of an eight-year field experiment in dry heathland

    DEFF Research Database (Denmark)

    Holmstrup, Martin; Schmelz, Rüdiger M.; Carrera, Noela

    2015-01-01

    In a long-term field trial we investigated the responses of enchytraeids to simulated future climatic conditions predicted for Denmark. At a semi-natural Danish heathland site we exposed 9.1 m2 plots to elevated atmospheric CO2 concentration (510 ppm), extended summer drought and passive night...... in spring 2013, perhaps indicating that warming stimulates fragmentation (reproduction) rates at this time of the year. Increased drought in MayeJune 2012 did not have lasting effects on abundance or biomass 3 months after the termination of drought treatment. However, comparison with earlier assessments...... of enchytraeids in the CLIMAITE experiment shows that the severity of drought and the time elapsed since the last drought is the best predictor of the biovolume (or biomass) of enchytraeids. Moreover, species richness was significantly impacted by the average soil water content experienced by enchytraeids during...

  9. On choice of tempered steels

    International Nuclear Information System (INIS)

    Govorov, A.A.; Pan'shin, I.F.; Rakhmanov, V.I.

    1978-01-01

    For the purpose of developing a graphical method for choosing structural steels, a change in the propagation work of a crack and in the critical temperature of brittleness of 40, 40Kh, 40KhN, and 40KhNM steels, was examined depending on the hardness after hardening and tempering. A diagram enabling to choose the grade of steel for making an article of known dimensions according to the preset values of its mechanical properties has been plotted. The developed selection scheme takes into account the hardenability of steels and the influence of the hardness after thermal treatment on the cold-shortness of steel

  10. Plant community responses to climate change

    Energy Technology Data Exchange (ETDEWEB)

    Kongstad, J.

    2012-07-01

    Climate change is expected to affect terrestrial ecosystems across the globe with increased atmospheric CO{sub 2} concentration, higher temperatures and changes in precipitation patterns. These environmental factors are drivers of many important ecosystem processes, and changes in ecosystem function are therefore expected in the future. The aim of this PhD-thesis was to examine the effects of climate change on aboveground plant growth, plant composition and plant phenology in Danish heathland ecosystems. Two sites were investigated in large-scale field experiments: 1) the CLIMAITE site, 'Brandbjerg' and 2) the INCREASE site at Mols. Field manipulations lasted years and included: Warming, summer drought and (CLIMAITE only) elevated CO{sub 2} concentrations. The treatments were applied individually and in all possible combinations. Further, at Brandbjerg, but outside the treatment plots, a study was performed on the effects nitrogen and phosphorus addition on phenology, chemistry and growth of the dominant grass Deschampsia flexuosa (Wavy Hairgrass). In general, the aboveground vegetation responded less than expected to changing climatic conditions; even though Calluna vulgaris (Heather) increased in biomass over the study period, the biomass was not affected by the manipulations, indicating that C. vulgaris, has a strong resistance to changes in climate. Also, the grass biomass (primarily D. flexuosa) was not affected and was relatively constant over the period. I argue that the resilience of D. flexuosa towards the climatic treatments came from the plants ability to let the tissue die back, and then quickly recover once conditions again became favourable. That gave the plant a high resilience to changes in climatic factors. Calluna vulgaris, on the other hand, showed a resistance to changes by constantly maintaining the growth during the whole season, probably because of its evergreen status. Together, the two different strategies made the heathland

  11. New temperable solar coatings: Tempsol

    Science.gov (United States)

    Demiryont, Hulya

    2001-11-01

    This paper deals with the large area deposition and coating properties of the thermo-stable (temperable/bendable) solar coating material, CuO, and some new optical coating systems comprising CuO films for architectural and automotive/transportation applications. The CuO solar coating is combined with other coating layers, for example, an anti-reflection film, a reflection film, a coloration coating layer, etc., which are also thermo-stable. The film systems are developed at the research laboratory by D.C. Magnetron reactive sputtering process. The new developed technologies then transferred to the production line. Product performances are compared before and after heat treatment of the coating systems. Performance tables and other physical properties, including optical parameters, mechanical and environmental stability, storage properties, etc., are also presented for this new product series.

  12. The Effect of Tempering on Strength Properties and Seed Coat ...

    African Journals Online (AJOL)

    The effect of tempering on seed coat adhesion strength and mechanical strength of sorghum and millet grain kernels was investigated at different tempering durations. Tempering reduced the kernel breaking strength and had significant effect on seed coat adhesion strength. Tempering the grain for 60 minutes at ambient ...

  13. Elevated carbon dioxide and ozone alter productivity and ecosystem carbon content in northern temperate forests

    Science.gov (United States)

    Alan F. Talhelm; Kurt S. Pregitzer; Mark E. Kubiske; Donald R. Zak; Courtney E. Campany; Andrew J. Burton; Richard E. Dickson; George R. Hendrey; J. G. Isebrands; Keith F. Lewin; John Nagy; David F. Karnosky

    2014-01-01

    Three young northern temperate forest communities in the north-central United States were exposed to factorial combinations of elevated carbon dioxide (CO2) and tropospheric ozone (O3) for 11 years. Here, we report results from an extensive sampling of plant biomass and soil conducted at the conclusion of the experiment...

  14. Arthropod vertical stratification in temperate deciduous forests: Implications for conservation oriented management

    Science.gov (United States)

    Ulyshen Michael

    2011-01-01

    Studies on the vertical distribution patterns of arthropods in temperate deciduous forests reveal highly stratified (i.e., unevenly vertically distributed) communities. These patterns are determined by multiple factors acting simultaneously, including: (1) time (forest age, season, time of day); (2) forest structure (height, vertical foliage complexity, plant surface...

  15. Drought enhances symbiotic dinitrogen fixation and competitive ability of a temperate forest tree

    Science.gov (United States)

    Nina Wurzburger; Chelcy Ford Miniat

    2013-01-01

    General circulation models project more intense and frequent droughts over the next century, but many questions remain about how terrestrial ecosystems will respond. Of particular importance, is to understand how drought will alter the species composition of regenerating temperate forests wherein symbiotic dinitrogen (N2)- fixing plants play a...

  16. Gender-related traits in the dioecious shrub Empetrum rubrum in two plant communities in the Magellanic steppe

    Science.gov (United States)

    Díaz-Barradas, Mari Cruz; Zunzunegui, María; Collantes, Marta; Álvarez-Cansino, Leonor; García Novo, Francisco

    2014-10-01

    Following the theory on costs of reproduction, sexually dimorphic plants may exhibit several trade-offs in energy and resources that can determine gender dimorphism in morphological or physiological traits, especially during the reproductive period. In this study we assess whether the sexes of the dioecious species Empetrum rubrum differ in morphological and ecophysiological traits related to water economy and photochemical efficiency and whether these differences change in nearby populations with contrasting plant communities. We conducted physiological, morphological, sex ratio, and cover measurements in E. rubrum plants in the Magellanic steppe, North-Eastern part of Tierra del Fuego (Argentina), from two types of heathlands with differing community composition. We found differences between sites in soil pH and wind speed at the canopy level. E. rubrum plants exhibited lower photosynthetic height and higher LAI (leaf area index), lower RWC (relative water content) and higher water-use efficiency (lower Δ13C) in the heathland with harsher environmental conditions. Gender dimorphism in the physiological response was patent for photochemical efficiency and water use (RWC and Δ13C discrimination), with males showing a more conservative strategy in relation to females. Accordingly, male-biased sex ratio in the stress-prone community suggested a better performance of male plants under stressful environmental conditions. The integrated analysis of all variables (photochemical efficiency, RWC, leaf dry matter content (LDMC), pigments, and Δ13C) indicated an interaction between gender and heathland community effects in the physiological response. We suggest that female plants may exhibit compensatory mechanisms to face their higher reproductive costs.

  17. Effect of temperate climate tree species on gross ammonification, gross nitrification and N2O formation

    Science.gov (United States)

    Brüggemann, N.; Rosenkranz, P.; Papen, H.; Butterbach-Bahl, K.

    2003-04-01

    Microbial nitrogen turnover processes in the soil, like ammonification, nitrification and denitrification, play an important role in the formation of nitrous oxide (N2O): (i) ammonification, because it releases nitrogen from organic material in the form of ammonium (NH4+), which in turn can serve as substrate for nitrification; (ii) nitrification itself (i.e. the turnover of NH4+ to nitrate, NO3-), during which nitric oxide (NO) and N2O can be released as by-products at varying ratios; (iii) denitrification, in which NO3- serves as electron acceptor and is converted to molecular nitrogen (N2) via NO and N2O as intermediates, that can also be partially lost to the atmosphere. Temperate forest soils are a substantial source of atmospheric N2O contributing up to 10% to the total atmospheric N2O budget. However, this figure is afflicted with a huge uncertainty due to a number of factors governing the soil N2O formation, consumption, release and uptake, which are not fully understood at present. To one of these factors belongs the influence of the tree species on nitrogen turnover processes in the soil and the formation of N trace gases related with them. The aim of the present work was to analyse this tree species effect for the temperate climate region. For this purpose the effect of five different temperate tree species, having the same age and growing on the same soil in direct vicinity to each other, on gross ammonification and gross nitrification as well as on N2O formation was investigated. The trees (common beech, Fagus sylvatica; pedunculate oak, Quercus robur; Norway spruce, Picea abies; Japanese larch, Larix leptolepis; mountain pine, Pinus mugo) were part of a species trial in Western Jutland, Denmark, established in 1965 on a former sandy heathland. Samples from the soil under these five tree species were taken in spring and in summer 2002, respectively, differentiating between organic layer and mineral soil. The gross rates of ammonification as well of

  18. Tracing in situ amino acid uptake in plants and microbes with15N13C labelled compounds

    DEFF Research Database (Denmark)

    Andresen, Louise Christoffersen; Michelsen, Anders; Jonasson, Sven Evert

    amino acids. Furthermore, tannin addition tended to reduce plant uptake of label. By combining data on 15N recovery after 1 day in shoots and roots (fine and coarse) of the dominant heathland plants: the evergreen dwarf shrub Calluna vulgaris and the graminoid Deschampsia flexuosa, in soil...... microorganisms (chloroform fumigation extraction) and in soil water, we discuss the relative importance of free amino acids and ammonium as plant nutrients and microbial substrates in natural N-limited ecosystems with a high proportion of soil N held in tannin-N complexes. ...

  19. Net root growth and nutrient acquisition in response to predicted climate change in two contrasting heathland species

    DEFF Research Database (Denmark)

    Arndal, M.F.; Merrild, M.P.; Michelsen, A.

    2013-01-01

    Accurate predictions of nutrient acquisition by plant roots and mycorrhizas are critical in modelling plant responses to climate change.We conducted a field experiment with the aim to investigate root nutrient uptake in a future climate and studied root production by ingrowth cores, mycorrhizal...... to elevated CO2. The species-specific response to the treatments suggests different sensitivity to global change factors, which could result in changed plant competitive interactions and belowground nutrient pool sizes in response to future climate change....

  20. Microstructural characteristics of steel rebar submitted to the process of self - tempering

    International Nuclear Information System (INIS)

    Barreto, M.F.O.

    2014-01-01

    Every day the construction industry seeks new alternatives to maximize use of the materials provided new tools for structural design of the new techniques in welding rods. In this sense, this work proposes to study the mechanical properties of steel rebar self-tempering for civil construction, choosing the bars from 6.3 to 25.4 mm in diameter for the study. The cooling process of rebar, after rolling, known as self-tempering produces a rebar with excellent mechanical properties and low cost to the plant, they put the rebar shall be composed of concentric layers with different mechanical properties among themselves. To study the influence of these layers on the mechanical behavior of rebar microstructural characterization was made by means of electron microscopy analysis of transmitted light and scanning of all samples. From the results it was found that only the bars 20 and 25mm diameter had a heat treatment for self-tempering relevant. (author)

  1. Elevation alters ecosystem properties across temperate treelines globally

    Science.gov (United States)

    Mayor, Jordan R.; Sanders, Nathan J.; Classen, Aimée T.; Bardgett, Richard D.; Clément, Jean-Christophe; Fajardo, Alex; Lavorel, Sandra; Sundqvist, Maja K.; Bahn, Michael; Chisholm, Chelsea; Cieraad, Ellen; Gedalof, Ze'Ev; Grigulis, Karl; Kudo, Gaku; Oberski, Daniel L.; Wardle, David A.

    2017-01-01

    Temperature is a primary driver of the distribution of biodiversity as well as of ecosystem boundaries. Declining temperature with increasing elevation in montane systems has long been recognized as a major factor shaping plant community biodiversity, metabolic processes, and ecosystem dynamics. Elevational gradients, as thermoclines, also enable prediction of long-term ecological responses to climate warming. One of the most striking manifestations of increasing elevation is the abrupt transitions from forest to treeless alpine tundra. However, whether there are globally consistent above- and belowground responses to these transitions remains an open question. To disentangle the direct and indirect effects of temperature on ecosystem properties, here we evaluate replicate treeline ecotones in seven temperate regions of the world. We find that declining temperatures with increasing elevation did not affect tree leaf nutrient concentrations, but did reduce ground-layer community-weighted plant nitrogen, leading to the strong stoichiometric convergence of ground-layer plant community nitrogen to phosphorus ratios across all regions. Further, elevation-driven changes in plant nutrients were associated with changes in soil organic matter content and quality (carbon to nitrogen ratios) and microbial properties. Combined, our identification of direct and indirect temperature controls over plant communities and soil properties in seven contrasting regions suggests that future warming may disrupt the functional properties of montane ecosystems, particularly where plant community reorganization outpaces treeline advance.

  2. First Temperate Exoplanet Sized Up

    Science.gov (United States)

    2010-03-01

    Combining observations from the CoRoT satellite and the ESO HARPS instrument, astronomers have discovered the first "normal" exoplanet that can be studied in great detail. Designated Corot-9b, the planet regularly passes in front of a star similar to the Sun located 1500 light-years away from Earth towards the constellation of Serpens (the Snake). "This is a normal, temperate exoplanet just like dozens we already know, but this is the first whose properties we can study in depth," says Claire Moutou, who is part of the international team of 60 astronomers that made the discovery. "It is bound to become a Rosetta stone in exoplanet research." "Corot-9b is the first exoplanet that really does resemble planets in our solar system," adds lead author Hans Deeg. "It has the size of Jupiter and an orbit similar to that of Mercury." "Like our own giant planets, Jupiter and Saturn, the planet is mostly made of hydrogen and helium," says team member Tristan Guillot, "and it may contain up to 20 Earth masses of other elements, including water and rock at high temperatures and pressures." Corot-9b passes in front of its host star every 95 days, as seen from Earth [1]. This "transit" lasts for about 8 hours, and provides astronomers with much additional information on the planet. This is fortunate as the gas giant shares many features with the majority of exoplanets discovered so far [2]. "Our analysis has provided more information on Corot-9b than for other exoplanets of the same type," says co-author Didier Queloz. "It may open up a new field of research to understand the atmospheres of moderate- and low-temperature planets, and in particular a completely new window in our understanding of low-temperature chemistry." More than 400 exoplanets have been discovered so far, 70 of them through the transit method. Corot-9b is special in that its distance from its host star is about ten times larger than that of any planet previously discovered by this method. And unlike all such

  3. Impacts of short-term heatwaves on sun-induced chlorophyll fluorescence(SiF) in temperate tree species

    Science.gov (United States)

    Wang, F.; Gu, L.; Guha, A.; Han, J.; Warren, J.

    2017-12-01

    The current projections for global climate change forecast an increase in the intensity and frequency of extreme climatic events, such as droughts and short-term heat waves. Understanding the effects of short-term heat wave on photosynthesis process is of critical importance to predict global impacts of extreme weather event on vegetation. The diurnal and seasonal characteristics of SIF emitted from natural vegetation, e.g., forest and crop, have been studied at the ecosystem-scale, regional-scale and global-scale. However, the detailed response of SIF from different plant species under extremely weather event, especially short-term heat wave, have not been reported. The purpose of this study was to study the response of solar-induced chlorophyll fluorescence, gas exchange and continuous fluorescence at leaf scale for different temperate tree species. The short-term heatwave experiment was conducted using plant growth chamber (CMP6050, Conviron Inc., Canada). We developed an advanced spectral fitting method to obtain the plant SIF in the plant growth chamber. We compared SIF variation among different wavelength and chlorophyll difference among four temperate tree species. The diurnal variation of SIF signals at leaf-scales for temperate tree species are different under heat stress. The SIF response at leaf-scales and their difference for four temperate tree species are different during a cycle of short-term heatwave stress. We infer that SIF be used as a measure of heat tolerance for temperate tree species.

  4. Nematode spatial and ecological patterns from tropical and temperate rainforests.

    Directory of Open Access Journals (Sweden)

    Dorota L Porazinska

    Full Text Available Large scale diversity patterns are well established for terrestrial macrobiota (e.g. plants and vertebrates, but not for microscopic organisms (e.g. nematodes. Due to small size, high abundance, and extensive dispersal, microbiota are assumed to exhibit cosmopolitan distributions with no biogeographical patterns. This assumption has been extrapolated from local spatial scale studies of a few taxonomic groups utilizing morphological approaches. Recent molecularly-based studies, however, suggest something quite opposite. Nematodes are the most abundant metazoans on earth, but their diversity patterns are largely unknown. We conducted a survey of nematode diversity within three vertical strata (soil, litter, and canopy of rainforests at two contrasting latitudes in the North American meridian (temperate: the Olympic National Forest, WA, U.S.A and tropical: La Selva Biological Station, Costa Rica using standardized sampling designs and sample processing protocols. To describe nematode diversity, we applied an ecometagenetic approach using 454 pyrosequencing. We observed that: 1 nematode communities were unique without even a single common species between the two rainforests, 2 nematode communities were unique among habitats in both rainforests, 3 total species richness was 300% more in the tropical than in the temperate rainforest, 4 80% of the species in the temperate rainforest resided in the soil, whereas only 20% in the tropics, 5 more than 90% of identified species were novel. Overall, our data provided no support for cosmopolitanism at both local (habitats and large (rainforests spatial scales. In addition, our data indicated that biogeographical patterns typical of macrobiota also exist for microbiota.

  5. Nematode Spatial and Ecological Patterns from Tropical and Temperate Rainforests

    Science.gov (United States)

    Porazinska, Dorota L.; Giblin-Davis, Robin M.; Powers, Thomas O.; Thomas, W. Kelley

    2012-01-01

    Large scale diversity patterns are well established for terrestrial macrobiota (e.g. plants and vertebrates), but not for microscopic organisms (e.g. nematodes). Due to small size, high abundance, and extensive dispersal, microbiota are assumed to exhibit cosmopolitan distributions with no biogeographical patterns. This assumption has been extrapolated from local spatial scale studies of a few taxonomic groups utilizing morphological approaches. Recent molecularly-based studies, however, suggest something quite opposite. Nematodes are the most abundant metazoans on earth, but their diversity patterns are largely unknown. We conducted a survey of nematode diversity within three vertical strata (soil, litter, and canopy) of rainforests at two contrasting latitudes in the North American meridian (temperate: the Olympic National Forest, WA, U.S.A and tropical: La Selva Biological Station, Costa Rica) using standardized sampling designs and sample processing protocols. To describe nematode diversity, we applied an ecometagenetic approach using 454 pyrosequencing. We observed that: 1) nematode communities were unique without even a single common species between the two rainforests, 2) nematode communities were unique among habitats in both rainforests, 3) total species richness was 300% more in the tropical than in the temperate rainforest, 4) 80% of the species in the temperate rainforest resided in the soil, whereas only 20% in the tropics, 5) more than 90% of identified species were novel. Overall, our data provided no support for cosmopolitanism at both local (habitats) and large (rainforests) spatial scales. In addition, our data indicated that biogeographical patterns typical of macrobiota also exist for microbiota. PMID:22984536

  6. Tempering of Low-Temperature Bainite

    Science.gov (United States)

    Peet, Mathew J.; Babu, Sudarsanam Suresh; Miller, Mike K.; Bhadeshia, H. K. D. H.

    2017-07-01

    Electron microscopy, X-ray diffraction, and atom probe tomography have been used to identify the changes which occur during the tempering of a carbide-free bainitic steel transformed at 473 K (200 °C). Partitioning of solute between ferrite and thin-films of retained austenite was observed on tempering at 673 K (400 °C) for 30 minutes. After tempering at 673 K (400 °C) and 773 K (500 °C) for 30 minutes, cementite was observed in the form of nanometre scale precipitates. Proximity histograms showed that the partitioning of solutes other than silicon from the cementite was slight at 673 K (400 °C) and more obvious at 773 K (500 °C). In both cases, the nanometre scale carbides are greatly depleted in silicon.

  7. Well-Tempered Metadynamics Converges Asymptotically

    Science.gov (United States)

    Dama, James F.; Parrinello, Michele; Voth, Gregory A.

    2014-06-01

    Metadynamics is a versatile and capable enhanced sampling method for the computational study of soft matter materials and biomolecular systems. However, over a decade of application and several attempts to give this adaptive umbrella sampling method a firm theoretical grounding prove that a rigorous convergence analysis is elusive. This Letter describes such an analysis, demonstrating that well-tempered metadynamics converges to the final state it was designed to reach and, therefore, that the simple formulas currently used to interpret the final converged state of tempered metadynamics are correct and exact. The results do not rely on any assumption that the collective variable dynamics are effectively Brownian or any idealizations of the hill deposition function; instead, they suggest new, more permissive criteria for the method to be well behaved. The results apply to tempered metadynamics with or without adaptive Gaussians or boundary corrections and whether the bias is stored approximately on a grid or exactly.

  8. Beneficial Effects of Temperate Forage Legumes that Contain Condensed Tannins

    Directory of Open Access Journals (Sweden)

    Jennifer W. MacAdam

    2015-07-01

    Full Text Available The two temperate forage legumes containing condensed tannins (CT that promote ruminant production are birdsfoot trefoil (Lotus corniculatus L.; BFT and sainfoin (Onobrychis viciifolia Scop.; SF. Both are well-adapted to the cool-temperate climate and alkaline soils of the Mountain West USA. Condensed tannins comprise a diverse family of bioactive chemicals with multiple beneficial functions for ruminants, including suppression of internal parasites and enteric methane. Birdsfoot trefoil contains 10 to 40 g·CT·kg−1 dry matter (DM, while SF contains 30 to 80 g·CT·kg−1 DM. Our studies have focused on these two plant species and have demonstrated consistently elevated rates of gain for beef calves grazing both BFT and SF. Novel results from our BFT research include carcass dressing percentages and consumer sensory evaluations equivalent to feedlot-finished steers and significantly greater than grass-finished steers, but with omega-3 fatty acid concentrations equal to grass-finished beef. We have further demonstrated that ruminants fed BFT or SF will consume more endophyte-infected tall fescue (Schedonorus arundinaceus (Schreb. Dumort. forage or seed than ruminants fed a non-CT forage legume. There is great potential value for sustainable livestock production in the use of highly digestible, nitrogen-fixing legumes containing tannins demonstrated to improve ruminant productivity.

  9. Gas-particle interactions above a Dutch heathland: I. Surface exchange fluxes of NH3, SO2, HNO3 and HCl

    Directory of Open Access Journals (Sweden)

    E. Nemitz

    2004-01-01

    Full Text Available A field measurement campaign was carried out over a Dutch heathland to investigate the effect of gas-to-particle conversion and ammonium aerosol evaporation on surface/atmosphere fluxes of ammonia and related species. Continuous micrometeorological measurements of the surface exchange of NH3, SO2, HNO3 and HCl were made and are analyzed here with regard to average fluxes, deposition velocities (Vd, canopy resistances (Rc and canopy compensation point for NH3. Gradients of SO2, HNO3 and HCl were measured with a novel wet-denuder system with online anion chromatography. Measurements of HNO3 and HCl indicate an Rc of 100 to 200 s m-1 during warm daytime periods, probably at least partly due to non-zero acid partial pressures above NH4NO3 and NH4Cl on the leaf surfaces. Although it is likely that this observation is exacerbated by the effect of the evaporation of airborne NH4+ on the gradient measurements, the findings nevertheless add to the growing evidence that HNO3 and HCl are not always deposited at the maximum rate. Ammonia (NH3 fluxes show mainly deposition, with some periods of significant daytime emission. The net exchange could be reproduced both with an Rc model (deposition fluxes only using resistance parameterizations from former measurements, as well as with the canopy compensation point model, using parameterizations derived from the measurements. The apoplastic ratio of ammonium and hydrogen concentration (Γs=[NH4+]/[H+] of 1200 estimated from the measurements is large for semi-natural vegetation, but smaller than indicated by previous measurements at this site.

  10. Gas-particle interactions above a Dutch heathland: I. Surface exchange fluxes of NH3, SO2, HNO3 and HCl

    Science.gov (United States)

    Nemitz, E.; Sutton, M. A.; Wyers, G. P.; Jongejan, P. A. C.

    2004-07-01

    A field measurement campaign was carried out over a Dutch heathland to investigate the effect of gas-to-particle conversion and ammonium aerosol evaporation on surface/atmosphere fluxes of ammonia and related species. Continuous micrometeorological measurements of the surface exchange of NH3, SO2, HNO3 and HCl were made and are analyzed here with regard to average fluxes, deposition velocities (Vd), canopy resistances (Rc) and canopy compensation point for NH3. Gradients of SO2, HNO3 and HCl were measured with a novel wet-denuder system with online anion chromatography. Measurements of HNO3 and HCl indicate an Rc of 100 to 200 s m-1 during warm daytime periods, probably at least partly due to non-zero acid partial pressures above NH4NO3 and NH4Cl on the leaf surfaces. Although it is likely that this observation is exacerbated by the effect of the evaporation of airborne NH4+ on the gradient measurements, the findings nevertheless add to the growing evidence that HNO3 and HCl are not always deposited at the maximum rate. Ammonia (NH3) fluxes show mainly deposition, with some periods of significant daytime emission. The net exchange could be reproduced both with an Rc model (deposition fluxes only) using resistance parameterizations from former measurements, as well as with the canopy compensation point model, using parameterizations derived from the measurements. The apoplastic ratio of ammonium and hydrogen concentration (Γs=[NH4+]/[H+]) of 1200 estimated from the measurements is large for semi-natural vegetation, but smaller than indicated by previous measurements at this site.

  11. A Facies Model for Temperate Continental Glaciers.

    Science.gov (United States)

    Ashley, Gail Mowry

    1987-01-01

    Discusses the presence and dynamics of continental glaciers in the domination of the physical processes of erosion and deposition in the mid-latitudes during the Pleistocene period. Describes the use of a sedimentary facies model as a guide to recognizing ancient temperate continental glacial deposits. (TW)

  12. The Temperance Movement and Social Work

    Science.gov (United States)

    Murdach, Allison D.

    2009-01-01

    This article examines a forgotten episode in social work history: the involvement of the profession in the temperance movement in the late 19th and early 20th centuries. Though some notable social workers such as Jane Addams, Robert A. Woods, and Representative Jeannette Rankin (the first woman elected to the U.S. Congress), championed the…

  13. Trichinella in arctic, subarctic and temperate regions

    DEFF Research Database (Denmark)

    Kapel, C. M O

    1997-01-01

    The transmission and occurrence of Trichinella spp according to the zoogeography of different climatic conditions, socioeconomy and human activity are discussed. Comparing arctic, subarctic and temperate regions, it appears that the species of Trichinella present, the composition of the fauna...... and the human activity are all very important interacting factors affecting epidemiology. In Greenland, where only sylvatic trichinellosis is present, the high prevalence in wildlife appears closely connected with polar bear hunting. In the Scandinavian countries, the prevalence of both sylvatic and domestic...

  14. Stability of modularity and structural keystone species in temporal cumulative plant- flower-visitor networks

    DEFF Research Database (Denmark)

    Dupont, Yoko; Olesen, Jens Mogens

    2012-01-01

    Modularity is a structural property of ecological networks, which has received much interest, but has been poorly explored. Modules are distinct subsets of species interacting strongly with each other, but sparsely with species outside the subset. Using a series of temporal cumulative networks, we...... all flowering plants and flower-visiting insect species throughout the flowering season at three dry heathland sites in Denmark. For each site, we constructed cumulative networks every 0.5 months, resulting in series of 10–12 networks per site. Numbers of interactions, and plant and insect species...... around one or two hubs. These hub species encompassed a small number of plant species, many of which acted as hubs at several study sites and throughout most of their flowering season. Thus, these plants become of key importance in maintaining the structure of their pollination network. We conclude...

  15. PENGARUH VARIASI SUHU PADA PROSES SELF TEMPERING DAN VARIASI WAKTU TAHAN PADA PROSES TEMPERING TERHADAP SIFAT MEKANIS BAJA AISI 4140

    Directory of Open Access Journals (Sweden)

    Sunardi Sunardi

    2016-07-01

    Full Text Available Tujuan penelitian ini adalah untuk mengetahui sifat mekanis pada baja AISI 4140 dengan proses tempering dengan variasi waktu tahan dan variasi suhu pada proses self tempering. Material terlebih dahulu dipanaskan pada suhu 850°C, di tahan dengan waktu 14 dan 28 menit, media pendinginan adalah Oli SAE 20. Pada proses tempering baja dipanaskan kembali dengan suhu 200°C di tahan dengan waktu 30 dan 120 menit. Sedangkan untuk proses self tempering, baja di panaskan pada suhu 850°C di tahan dengan waktu 14 dan 28 menit kemudian didinginkan, suhu yang harus dicapai pada pendinginan adalah 200°C, 400°C dan 600°C. Proses tempering dengan variasi waktu tahan mempunyai nilai kekerasan terbesar 50,1 HRC dengan waktu tahan 120 menit, sedangkan nilai kekerasan terbesar pada proses self tempering dengan variasi suhu adalah 29,68 HRC pada suhu 200°C. Nilai ketangguhan terbesar pada saat proses tempering adalah 0,341 (J/mm2 dengan waktu tahan 120 menit, sedangkan pada saat proses self tempering ketangguhan terbesar pada suhu 600°C dengan nilai 0,375 (J/mm2. Laju korosi terbesar pada saat tempering adalah 0,055 (mpy dengan waktu tahan 30 menit, sedangkan pada saat proses self tempering laju korosi terbesar pada suhu 400°C dengan nilai 0,0388 (mpy. 

  16. Temperate and virulent Lactobacillus delbrueckii bacteriophages: comparison of their thermal and chemical resistance.

    Science.gov (United States)

    Ebrecht, Ana C; Guglielmotti, Daniela M; Tremmel, Gustavo; Reinheimer, Jorge A; Suárez, Viviana B

    2010-06-01

    The aim of this work was to study the efficiency of diverse chemical and thermal treatments usually used in dairy industries to control the number of virulent and temperate Lactobacillus delbrueckii bacteriophages. Two temperate (Cb1/204 and Cb1/342) and three virulent (BYM, YAB and Ib3) phages were studied. The thermal treatments applied were: 63 degrees C for 30 min (low temperature--long time, LTLT), 72 degrees C for 15 s (high temperature--short time, HTST), 82 degrees C for 5 min (milk destined to yogurt elaboration) and 90 degrees C for 15 min (FIL-IDF). The chemical agents studied were: sodium hypochlorite, ethanol, isopropanol, peracetic acid, biocides A (quaternary ammonium chloride), B (hydrogen peroxide, peracetic acid and peroctanoic acid), C (alkaline chloride foam), D (p-toluensulfonchloroamide, sodium salt) and E (ethoxylated nonylphenol and phosphoric acid). The kinetics of inactivation were drew and T(99) (time necessary to eliminate the 99% of phage particles) calculated. Results obtained showed that temperate phages revealed lower resistance than the virulent ones to the treatment temperatures. Biocides A, C, E and peracetic acid showed a notable efficiency to inactivate high concentrations of temperate and virulent L. delbrueckii phages. Biocide B evidenced, in general, a good capacity to eliminate the phage particles. Particularly for this biocide virulent phage Ib3 showed the highest resistance in comparison to the rest of temperate and virulent ones. On the contrary, biocide D and isopropanol presented a very low capacity to inactivate all phages studied. The efficiency of ethanol and hypochlorite was variable depending to the phages considered. These results allow a better knowledge and give useful information to outline more effective treatments to reduce the phage infections in dairy plants. 2009 Elsevier Ltd. All rights reserved.

  17. Clonal and bud bank traits: patterns across temperate plant communities

    Czech Academy of Sciences Publication Activity Database

    Klimešová, Jitka; Herben, Tomáš

    2015-01-01

    Roč. 26, č. 2 (2015), s. 243-253 ISSN 1100-9233 R&D Projects: GA ČR GB14-36079G; GA ČR GA13-17118S; GA ČR GAP505/12/1007 Institutional support: RVO:67985939 Keywords : clonal and bud bank traits * vegetation * central Europe Subject RIV: EH - Ecology, Behaviour Impact factor: 3.151, year: 2015

  18. Parallel tempering in full QCD with Wilson fermions

    International Nuclear Information System (INIS)

    Ilgenfritz, E.-M.; Kerler, W.; Mueller-Preussker, M.; Stueben, H.

    2002-01-01

    We study the performance of QCD simulations with dynamical Wilson fermions by combining the hybrid Monte Carlo algorithm with parallel tempering on 10 4 and 12 4 lattices. In order to compare tempered with standard simulations, covariance matrices between subensembles have to be formulated and evaluated using the general properties of autocorrelations of the parallel tempering algorithm. We find that rendering the hopping parameter κ dynamical does not lead to an essential improvement. We point out possible reasons for this observation and discuss more suitable ways of applying parallel tempering to QCD

  19. Incidence of invasive macrophytes on methylmercury budget in temperate lakes: Central role of bacterial periphytic communities

    International Nuclear Information System (INIS)

    Gentès, Sophie; Monperrus, Mathilde; Legeay, Alexia; Maury-Brachet, Régine; Davail, Stephane; André, Jean-Marc; Guyoneaud, Rémy

    2013-01-01

    Several studies demonstrated high mercury (Hg) methylation and demethylation in the periphyton associated with floating roots in tropical ecosystems. The importance of aquatic plants on methylmercury production in three temperate ecosystems from south-western France was evaluated through Hg species concentrations, and Hg methylation/demethylation activities by using stable isotopic tracers ( 199 Hg(II), Me 201 Hg). Hg accumulation and high methylation and demethylation yields were detected in plant roots and periphyton, whereas results for sediment and water were low to insignificant. The presence of sulfate reducing prokaryotes was detected in all compartments (T-RFLP based on dsrAB amplified through nested PCR) and their main role in Hg methylation could be demonstrated. In turn, sulfate reduction inhibition did not affect demethylation activities. The estimation of net MeHg budgets in these ecosystems suggested that aquatic rhizosphere is the principal location for methylmercury production and may represent an important source for the contamination of the aquatic food chain. - Highlights: ► Both Hg methylation and demethylation occur in the periphyton of temperate ecosystems. ► Aquatic rhizosphere is the main compartment for net methylmercury production. ► Sulfate reducers are detected in all ecosystem compartments (water, sediment, periphyton). ► Sulfate reducers are responsible for methylmercury production in aquatic roots. - The incidence of periphytic microbial communities on net methylmercury production is highlighted in temperate aquatic ecosystems.

  20. Genome-Wide Association Mapping for Cell Wall Composition and Properties in Temperate Grasses

    DEFF Research Database (Denmark)

    Bellucci, Andrea

    with a wide range of chemical bounds. At present the interest in plant cell wall is growing due to the possibility to convert ligno-cellulosic biomass (e.g. agricultural residues) into bioethanol but also for the benefits to human health of some cell wall constituents found in cereals, in particular beta......-glucans. Plant cell wall biosynthesis is regulated by a large number of genes and regulatory factors but very few of these are known and characterized. This PhD project aimed to the identification of putative candidate genes involved in plant cell wall composition and properties using a genome wide (GWAS......) approach. The species investigate were wheat, barley and B. distachyon, considered a model plant for temperate cereals. Agronomical traits as yield and plant height were also included in the analysis along with cell wall composition and saccharification properties. Several marker-trait associations were...

  1. Oceanic temperate forest versus warm temperate rainforest: a reply to Grubb et al. (2017)

    DEFF Research Database (Denmark)

    McGlone, Matt S.; Buitenwerf, Robert; Richardson, Sarah J.

    2017-01-01

    Grubb et al. (2017) point out that we (McGlone et al. 2016) erroneously stated that the definition of warm temperate rain forest (WTRF; Grubb et al. 2013) was based in part on climatic criteria. We apologise: their text made clear that this was not the case. However, they go on to say that they ‘...

  2. Microstructural characterisation of a P91 steel normalised and tempered at different temperatures

    International Nuclear Information System (INIS)

    Hurtado-Norena, C.; Danon, C.A.; Luppo, M.I.; Bruzzoni, P.

    2015-01-01

    9%Cr-1%Mo martensitic-ferritic steels are used in power plant components with operating temperatures of around 600 deg. C because of their good mechanical properties at high temperature as well as good oxidation resistance. These steels are generally used in the normalised and tempered condition. This treatment results in a structure of tempered lath martensite where the precipitates are distributed along the lath interfaces and within the martensite laths. The characterisation of these precipitates is of fundamental importance because of their relationship with the creep behaviour of these steels in service. In the present work, the different types of precipitates found in these steels have been studied on specimens in different metallurgical conditions. The techniques used in this investigation were X-ray diffraction with synchrotron light, scanning electron microscopy, energy dispersive microanalysis and transmission electron microscopy. (authors)

  3. Adhesion and wear properties of boro-tempered ductile iron

    International Nuclear Information System (INIS)

    Kayali, Yusuf; Yalcin, Yilmaz; Taktak, Suekrue

    2011-01-01

    Highlights: → In this study, the wear and adhesion properties of BDI were investigated. → Boro-tempering process under several heat treatment conditions was examined. → Optical microscope, SEM and XRD analysis were carried out to investigate the microstructure. → It was observed that boro-tempering process improves micro-hardness and wear properties of ductile irons. -- Abstract: In this study, adhesion and wear properties of boro-tempered ductile iron (BDI) were investigated. Boro-tempering was carried out on two stage processes i.e. boronizing and tempering. At the first stage, ductile iron samples were boronized by using pack process at 900 o C for 1, 3, and 5 h and then, secondly tempered at 250, 300, 350, and 400 o C for 1 h. X-ray diffraction (XRD) analysis of boro-tempered samples showed that FeB and Fe 2 B phases were found on the surface of the samples. The Daimler-Benz Rockwell-C adhesion test was used to assess the adhesion of boride layer. Test result showed that adhesion decreased with increasing boriding time and increased with increasing tempering temperature. Dry sliding wear tests of these samples were performed against Al 2 O 3 ball at a constant sliding speed and loads of 5 and 10 N. Wear tests indicated that boro-tempering heat treatment increased wear resistance of ductile iron. In addition, it was found that while wear rate of boro-tempered samples decreased with increasing boriding time, there is no significant affect of tempering temperature on wear rate.

  4. Hardness of H13 Tool Steel After Non-isothermal Tempering

    Science.gov (United States)

    Nelson, E.; Kohli, A.; Poirier, D. R.

    2018-04-01

    A direct method to calculate the tempering response of a tool steel (H13) that exhibits secondary hardening is presented. Based on the traditional method of presenting tempering response in terms of isothermal tempering, we show that the tempering response for a steel undergoing a non-isothermal tempering schedule can be predicted. Experiments comprised (1) isothermal tempering, (2) non-isothermal tempering pertaining to a relatively slow heating to process-temperature and (3) fast-heating cycles that are relevant to tempering by induction heating. After establishing the tempering response of the steel under simple isothermal conditions, the tempering response can be applied to non-isothermal tempering by using a numerical method to calculate the tempering parameter. Calculated results are verified by the experiments.

  5. Dry matter intake and digestibility of temperate pastures ...

    African Journals Online (AJOL)

    The aim of this study was to evaluate the effect of sorghum grain supplementation on total and forage dry matter (DM) intake and digestibility of wethers and heifers consuming temperate pasture. Twenty four Corriedale x Milchschaf wethers and 24 crossbred heifers fed temperate pasture were non-supplemented or ...

  6. Simulation of Residual Stresses at Holes in Tempered Glass

    DEFF Research Database (Denmark)

    Nielsen, Jens Henrik; Olesen, John Forbes; Poulsen, Peter Noe

    2010-01-01

    This work presents a full 3D numerical study of the residual stresses in tempered (toughened) glass near holes using Narayanaswamy’s model for the tempering process. It is the objective of the paper to elucidate the influence on the minimal residual compressive stresses at holes from variations in...

  7. Transfer parameter values in temperate forest ecosystems: a review

    International Nuclear Information System (INIS)

    Calmon, Philippe; Thiry, Yves; Zibold, Gregor; Rantavaara, Aino; Fesenko, Sergei

    2009-01-01

    activity reduction, quantified as an ecological half-life, reflect the soil and pasture conditions at individual locations. Forests in temperate and boreal regions differ with respect to soil type and vegetation, and a faster decline of muscle activity concentrations in deer occurs in the temperate zone. However, in wild boar the caesium activity concentration shows no decline because of its special feeding habits. In the late phase, i.e. at least a few months since the external radionuclide contamination on feed plants has been removed, a T ag value of 0.01 m 2 kg -1 (fresh weight) is common for 137 Cs in the muscles of adult moose and terrestrial birds living in boreal forests, and 0.03 m 2 kg -1 (fresh weight) for arctic hare. Radiocaesium concentrations in reindeer muscle in winter may exceed the summer content by a factor of more than two, the mean T ag values for winter ranging from 0.02 to 0.8 m 2 kg -1 (fresh weight), and in summer from 0.04 to 0.4 m 2 kg -1 . The highest values are found in the year of initial contamination, followed by a gradual reduction. In waterfowl a relatively fast decline in uptake of 137 Cs has been found, with T ag values changing from 0.01 to 0.002 m 2 kg -1 (fresh weight) in the three years after the contaminating event, the rate being determined by the dynamics of 137 Cs in aquatic ecosystems.

  8. Transfer parameter values in temperate forest ecosystems: a review

    Energy Technology Data Exchange (ETDEWEB)

    Calmon, Philippe [Department of Radioecology, Institute of Radioprotection and Nuclear Safety, CE Cadarache, BP 3, 13115 Saint Paul-les-Durance Cedex (France)], E-mail: philippe.calmon@irsn.fr; Thiry, Yves [Biosphere Impact Studies, Belgian Nuclear Research Center (SCK.CEN, Foundation of Public Utility), 2400 Mol (Belgium); Zibold, Gregor [Hochschule Ravensburg-Weingarten, University of Applied Sciences, 88250 Weingarten (Germany); Rantavaara, Aino [Research and Environmental Surveillance, Radiation and Nuclear Safety Authority (STUK), BP 14, FIN-00881 Helsinki (Finland); Fesenko, Sergei [International Atomic Energy Agency (IAEA), 1400 Vienna (Austria)

    2009-09-15

    of radioactive caesium to game animals and reindeer and the rate of activity reduction, quantified as an ecological half-life, reflect the soil and pasture conditions at individual locations. Forests in temperate and boreal regions differ with respect to soil type and vegetation, and a faster decline of muscle activity concentrations in deer occurs in the temperate zone. However, in wild boar the caesium activity concentration shows no decline because of its special feeding habits. In the late phase, i.e. at least a few months since the external radionuclide contamination on feed plants has been removed, a T{sub ag} value of 0.01 m{sup 2} kg{sup -1} (fresh weight) is common for {sup 137}Cs in the muscles of adult moose and terrestrial birds living in boreal forests, and 0.03 m{sup 2} kg{sup -1} (fresh weight) for arctic hare. Radiocaesium concentrations in reindeer muscle in winter may exceed the summer content by a factor of more than two, the mean T{sub ag} values for winter ranging from 0.02 to 0.8 m{sup 2} kg{sup -1} (fresh weight), and in summer from 0.04 to 0.4 m{sup 2} kg{sup -1}. The highest values are found in the year of initial contamination, followed by a gradual reduction. In waterfowl a relatively fast decline in uptake of {sup 137}Cs has been found, with T{sub ag} values changing from 0.01 to 0.002 m{sup 2} kg{sup -1} (fresh weight) in the three years after the contaminating event, the rate being determined by the dynamics of {sup 137}Cs in aquatic ecosystems.

  9. Moist temperate forest butterflies of western Bhutan

    Directory of Open Access Journals (Sweden)

    Arun P. Singh

    2016-03-01

    Full Text Available Random surveys were carried out in moist temperate forests (1,860–3,116 m around Bunakha Village and Dochula Pass, near Thimphu in western Bhutan, recording 65 species of butterflies.  Of these, 11 species, viz., Straightwing Blue Orthomiella pontis pontis Elwes, Slate Royal Maneca bhotea bhotea Moore, Dull Green Hairstreak Esakiozephyrus icana Moore, Yellow Woodbrown Lethe nicetas Hewitson, Small Silverfork Zophoessa jalaurida elwesi Moore, Scarce Labyrinth, Neope pulahina (Evans, Chumbi Wall Chonala masoni Elwes, Pale Hockeystick Sailer Neptis manasa manasa Moore and White Commodore Parasarpa dudu dudu Westwood, are restricted to the eastern Himalaya, northeastern India and Myanmar.  Two other species, Tawny Mime Chiasa agestor agestor (Gray and Himalayan Spotted Flat Celaenorrhinus munda Moore have been only rarely recorded from Bhutan and a few individuals of the rare Bhutan Glory Bhutanitis lidderdalei Atkinson were also recorded near Bunakha.  

  10. Study on tempering behaviour of AISI 410 stainless steel

    International Nuclear Information System (INIS)

    Chakraborty, Gopa; Das, C.R.; Albert, S.K.; Bhaduri, A.K.; Thomas Paul, V.; Panneerselvam, G.; Dasgupta, Arup

    2015-01-01

    Martensitic stainless steels find extensive applications due to their optimum combination of strength, hardness and wear-resistance in tempered condition. However, this class of steels is susceptible to embrittlement during tempering if it is carried out in a specific temperature range resulting in significant reduction in toughness. Embrittlement of as-normalised AISI 410 martensitic stainless steel, subjected to tempering treatment in the temperature range of 673–923 K was studied using Charpy impact tests followed by metallurgical investigations using field emission scanning electron and transmission electron microscopes. Carbides precipitated during tempering were extracted by electrochemical dissolution of the matrix and identified by X-ray diffraction. Studies indicated that temper embrittlement is highest when the steel is tempered at 823 K. Mostly iron rich carbides are present in the steel subjected to tempering at low temperatures of around 723 K, whereas chromium rich carbides (M 23 C 6 ) dominate precipitation at high temperature tempering. The range 773–823 K is the transition temperature range for the precipitates, with both Fe 2 C and M 23 C 6 types of carbides coexisting in the material. The nucleation of Fe 2 C within the martensite lath, during low temperature tempering, has a definite role in the embrittlement of this steel. Embrittlement is not observed at high temperature tempering because of precipitation of M 23 C 6 carbides, instead of Fe 2 C, preferentially along the lath and prior austenite boundaries. Segregation of S and P, which is widely reported as one of the causes for temper embrittlement, could not be detected in the material even through Auger electron spectroscopy studies. - Highlights: • Tempering behaviour of AISI 410 steel is studied within 673–923 K temperature range. • Temperature regime of maximum embrittlement is identified as 773–848 K. • Results show that type of carbide precipitation varies with

  11. Study on tempering behaviour of AISI 410 stainless steel

    Energy Technology Data Exchange (ETDEWEB)

    Chakraborty, Gopa, E-mail: gopa_mjs@igcar.gov.in [Metallurgy & Materials Group, Indira Gandhi Center for Atomic Research, Kalpakkam 603102 (India); Das, C.R.; Albert, S.K.; Bhaduri, A.K.; Thomas Paul, V. [Metallurgy & Materials Group, Indira Gandhi Center for Atomic Research, Kalpakkam 603102 (India); Panneerselvam, G. [Chemistry Group, Indira Gandhi Center for Atomic Research, Kalpakkam 603102 (India); Dasgupta, Arup [Metallurgy & Materials Group, Indira Gandhi Center for Atomic Research, Kalpakkam 603102 (India)

    2015-02-15

    Martensitic stainless steels find extensive applications due to their optimum combination of strength, hardness and wear-resistance in tempered condition. However, this class of steels is susceptible to embrittlement during tempering if it is carried out in a specific temperature range resulting in significant reduction in toughness. Embrittlement of as-normalised AISI 410 martensitic stainless steel, subjected to tempering treatment in the temperature range of 673–923 K was studied using Charpy impact tests followed by metallurgical investigations using field emission scanning electron and transmission electron microscopes. Carbides precipitated during tempering were extracted by electrochemical dissolution of the matrix and identified by X-ray diffraction. Studies indicated that temper embrittlement is highest when the steel is tempered at 823 K. Mostly iron rich carbides are present in the steel subjected to tempering at low temperatures of around 723 K, whereas chromium rich carbides (M{sub 23}C{sub 6}) dominate precipitation at high temperature tempering. The range 773–823 K is the transition temperature range for the precipitates, with both Fe{sub 2}C and M{sub 23}C{sub 6} types of carbides coexisting in the material. The nucleation of Fe{sub 2}C within the martensite lath, during low temperature tempering, has a definite role in the embrittlement of this steel. Embrittlement is not observed at high temperature tempering because of precipitation of M{sub 23}C{sub 6} carbides, instead of Fe{sub 2}C, preferentially along the lath and prior austenite boundaries. Segregation of S and P, which is widely reported as one of the causes for temper embrittlement, could not be detected in the material even through Auger electron spectroscopy studies. - Highlights: • Tempering behaviour of AISI 410 steel is studied within 673–923 K temperature range. • Temperature regime of maximum embrittlement is identified as 773–848 K. • Results show that type of

  12. Forecasting cyanobacteria dominance in Canadian temperate lakes.

    Science.gov (United States)

    Persaud, Anurani D; Paterson, Andrew M; Dillon, Peter J; Winter, Jennifer G; Palmer, Michelle; Somers, Keith M

    2015-03-15

    Predictive models based on broad scale, spatial surveys typically identify nutrients and climate as the most important predictors of cyanobacteria abundance; however these models generally have low predictive power because at smaller geographic scales numerous other factors may be equally or more important. At the lake level, for example, the ability to forecast cyanobacteria dominance is of tremendous value to lake managers as they can use such models to communicate exposure risks associated with recreational and drinking water use, and possible exposure to algal toxins, in advance of bloom occurrence. We used detailed algal, limnological and meteorological data from two temperate lakes in south-central Ontario, Canada to determine the factors that are closely linked to cyanobacteria dominance, and to develop easy to use models to forecast cyanobacteria biovolume. For Brandy Lake (BL), the strongest and most parsimonious model for forecasting % cyanobacteria biovolume (% CB) included water column stability, hypolimnetic TP, and % cyanobacteria biovolume two weeks prior. For Three Mile Lake (TML), the best model for forecasting % CB included water column stability, hypolimnetic TP concentration, and 7-d mean wind speed. The models for forecasting % CB in BL and TML are fundamentally different in their lag periods (BL = lag 1 model and TML = lag 2 model) and in some predictor variables despite the close proximity of the study lakes. We speculate that three main factors (nutrient concentrations, water transparency and lake morphometry) may have contributed to differences in the models developed, and may account for variation observed in models derived from large spatial surveys. Our results illustrate that while forecast models can be developed to determine when cyanobacteria will dominate within two temperate lakes, the models require detailed, lake-specific calibration to be effective as risk-management tools. Copyright © 2015 Elsevier Ltd. All rights reserved.

  13. Staying cool: preadaptation to temperate climates required for colonising tropical alpine-like environments

    Directory of Open Access Journals (Sweden)

    Berit Gehrke

    2018-04-01

    Full Text Available Plant species tend to retain their ancestral ecology, responding to temporal, geographic and climatic changes by tracking suitable habitats rather than adapting to novel conditions. Nevertheless, transitions into different environments or biomes still seem to be common. Especially intriguing are the tropical alpine-like areas found on only the highest mountainous regions surrounded by tropical environments. Tropical mountains are hotspots of biodiversity, often with striking degrees of endemism at higher elevations. On these mountains, steep environmental gradients and high habitat heterogeneity within small spaces coincide with astounding species diversity of great conservation value. The analysis presented here shows that the importance of in situ speciation in tropical alpine-like areas has been underestimated. Additionally and contrary to widely held opinion, the impact of dispersal from other regions with alpine-like environments is relatively minor compared to that of immigration from other biomes with a temperate (but not alpine-like climate. This suggests that establishment in tropical alpine-like regions is favoured by preadaptation to a temperate, especially aseasonal, freezing regime such as the cool temperate climate regions in the Tropics. Furthermore, emigration out of an alpine-like environment is generally rare, suggesting that alpine-like environments – at least tropical ones – are species sinks.

  14. Ecological legacies of Indigenous fire management in high-latitude coastal temperate rainforests, Canada

    Science.gov (United States)

    Hoffman, K.; Lertzman, K. P.; Starzomski, B. M.

    2016-12-01

    Anthropogenic burning is considered to have little impact on coastal temperate rainforest fire regimes in the Pacific Northwest (PNW) of North America, yet few long-term fire histories have been reconstructed in these forests. We use a multidisciplinary approach to reconstruct the ecological impact, scale, and legacies of historic fire regime variability in high-latitude coastal temperate rainforests located in British Columbia, Canada. We map seven centuries of fire activity with fire scars and records of stand establishment, and examine patterns in the distribution and composition of vegetation to assess whether fire was historically used as a tool for resource management. We conduct a paired study of 20 former Indigenous habitation and control sites across a 100 km2 island group to relate historic fire activity with long-term patterns of human land use and contemporary lightning strike densities. Fires were significantly associated with the locations of former Indigenous habitation sites, low and mixed in severity, and likely intentionally used to influence the composition and structure of vegetation, thus increasing the productivity of culturally important plants such as western redcedar, berry-producing shrubs, and bracken fern. Centuries of repeated anthropogenic burning have resulted in a mosaic of vegetation types in different stages of succession. These data are directly relevant to the management of contemporary forests as they do not support the widespread contention that old growth coastal temperate rainforests in this region are pristine landscapes where fire is rare, but more likely the result of long-term human land use practices.

  15. Responses of Winter Wheat Yields to Warming-Mediated Vernalization Variations Across Temperate Europe

    Directory of Open Access Journals (Sweden)

    Xiuchen Wu

    2017-10-01

    Full Text Available Rapid climate warming, with much higher warming rates in winter and spring, could affect the vernalization fulfillment, a critical process for induction of crop reproductive growth and consequent grain filling in temperate winter crops. However, regional observational evidence of the effects of historical warming-mediated vernalization variations on temperate winter crop yields is lacking. Here, we statistically quantified the interannual sensitivity of winter wheat yields to vernalization degree days (VDD during 1975–2009 and its spatial relationship with multi-year mean VDD over temperate Europe (TE, using EUROSTAT crop yield statistics, observed and simulated crop phenology data and gridded daily climate data. Our results revealed a pervasively positive interannual sensitivity of winter wheat yields to variations in VDD (γVDD over TE, with a mean γVDD of 2.8 ± 1.5 kg ha−1 VDD−1. We revealed a significant (p < 0.05 negative exponential relationship between γVDD and multi-year mean VDD for winter wheat across TE, with higher γVDD in winter wheat planting areas with lower multi-year mean VDD. Our findings shed light on potential vulnerability of winter wheat yields to warming-mediated vernalization variations over TE, particularly considering a likely future warmer climate.

  16. Temperate radiations and dying embers of a tropical past: the diversification of Viburnum.

    Science.gov (United States)

    Spriggs, Elizabeth L; Clement, Wendy L; Sweeney, Patrick W; Madriñán, Santiago; Edwards, Erika J; Donoghue, Michael J

    2015-07-01

    We used a near-complete phylogeny for the angiosperm clade Viburnum to assess lineage diversification rates, and to examine possible morphological and ecological factors driving radiations. Maximum-likelihood and Bayesian approaches identified shifts in diversification rate and possible links to character evolution. We inferred the ancestral environment for Viburnum and changes in diversification dynamics associated with subsequent biome shifts. Viburnum probably diversified in tropical forests of Southeast Asia in the Eocene, with three subsequent radiations in temperate clades during the Miocene. Four traits (purple fruits, extrafloral nectaries, bud scales and toothed leaves) were statistically associated with higher rates of diversification. However, we argue that these traits are unlikely to be driving diversification directly. Instead, two radiations were associated with the occupation of mountainous regions and a third with repeated shifts between colder and warmer temperate forests. Early-branching depauperate lineages imply that the rare lowland tropical species are 'dying embers' of once more diverse lineages; net diversification rates in Viburnum likely decreased in these tropical environments after the Oligocene. We suggest that 'taxon pulse' dynamics might characterize other temperate plant lineages. © 2015 The Authors. New Phytologist © 2015 New Phytologist Trust.

  17. Comparison of seasonal soil microbial process in snow-covered temperate ecosystems of northern China.

    Directory of Open Access Journals (Sweden)

    Xinyue Zhang

    Full Text Available More than half of the earth's terrestrial surface currently experiences seasonal snow cover and soil frost. Winter compositional and functional investigations in soil microbial community are frequently conducted in alpine tundra and boreal forest ecosystems. However, little information on winter microbial biogeochemistry is known from seasonally snow-covered temperate ecosystems. As decomposer microbes may differ in their ability/strategy to efficiently use soil organic carbon (SOC within different phases of the year, understanding seasonal microbial process will increase our knowledge of biogeochemical cycling from the aspect of decomposition rates and corresponding nutrient dynamics. In this study, we measured soil microbial biomass, community composition and potential SOC mineralization rates in winter and summer, from six temperate ecosystems in northern China. Our results showed a clear pattern of increased microbial biomass C to nitrogen (N ratio in most winter soils. Concurrently, a shift in soil microbial community composition occurred with higher fungal to bacterial biomass ratio and gram negative (G- to gram positive (G+ bacterial biomass ratio in winter than in summer. Furthermore, potential SOC mineralization rate was higher in winter than in summer. Our study demonstrated a distinct transition of microbial community structure and function from winter to summer in temperate snow-covered ecosystems. Microbial N immobilization in winter may not be the major contributor for plant growth in the following spring.

  18. [Factors affecting the vegetation restoration after fires in cold temperate wetlands: A review].

    Science.gov (United States)

    Zhao, Feng-Jun; Wang, Li-Zhong; Shu, Li-Fu; Chen, Peng-Yu; Chen, Li-guang

    2013-03-01

    Cold temperate wetland plays an important role in maintaining regional ecological balance. Fire is an important disturbance factor in wetland ecosystem. Severe burning can induce the marked degradation of the ecological functions of wetland ecosystem. The vegetation restoration, especially the early vegetation restoration, after fires, is the premise and basis for the recovery of the ecological functions of the ecosystem. This paper reviewed the research progress on the factors affecting the vegetation restoration after fires in wetlands. The vegetation restoration after fires in cold temperate wetlands was controlled by the fire intensity, fire size, vegetation types before fires, regeneration characteristics of plant species, and site conditions. It was considered that the long-term monitoring on the post-fire vegetation restoration in cold temperate wetland, the key factors affecting the vegetation restoration, the roles of frozen soil layer on the post-fire vegetation restoration, and the theories and technologies on the vegetation restoration would be the main research directions in the future.

  19. Stress relaxation in tempered glass caused by heat soak testing

    DEFF Research Database (Denmark)

    Schneider, Jens; Hilcken, Jonas; Aronen, Antti

    2016-01-01

    Heat soak testing of tempered glass is a thermal process required after the tempering process itself to bring glasses of commercial soda-lime-silica-glass to failure that are contaminated with nickel sulphide inclusions, diameter 50 mm to 500 mm typically. Thus, the tests avoid a so-called "spont...... of commercial soda-lime-silica glass, it causes stress relaxation in tempered glass and the fracture pattern of the glass changes accordingly, especially thin glasses are affected. Based on the Tool-Narayanaswamy-Model, this paper comprises the theoretical background of the stress...

  20. Adaptive root foraging strategies along a boreal–temperate forest gradient

    Czech Academy of Sciences Publication Activity Database

    Ostonen, I.; Truu, M.; Helmisaari, H.-S.; Lukač, M.; Borken, W.; Vanguelova, H.; Godbold, Douglas; Löhmus, K.; Zang, U.; Tedersoo, L.; Preem, J.-K.; Rosenvald, K.; Aosaar, J.; Armolaitis, K.; Frey, J.; Kabral, N.; Kukumägi, M.; Leppälammi-Kujansuu, J.; Lindroos, A.-J.; Merila, P.; Napa, Ü.; Nöjd, P.; Parts, K.; Uri, V.; Varik, M.; Truu, J.

    2017-01-01

    Roč. 215, č. 3 (2017), s. 977-991 ISSN 0028-646X EU Projects: European Commission(XE) 315982; European Commission(XE) 315982; European Commission(XE) 90/E38 Institutional support: RVO:67179843 Keywords : boreal and temperate forests * climate gradient * ectomycorrhizal (EcM) mycelium * fine and ectomycorrhizal root biomass * root foraging * root morphology * soil and rhizosphere bacteria * soil C * N ratio Subject RIV: EF - Botanics OBOR OECD: Plant sciences, botany Impact factor: 7.330, year: 2016

  1. Evaluation of hyper-tempering and machining residual stresses in a pipe and a cast elbow

    International Nuclear Information System (INIS)

    Dupas, P.; Le Delliou, P.; Sussen, L.

    1995-01-01

    Cast elbows in austeno-ferritic stainless steel from the primary circuit of nuclear power plants suffer from important residual stresses initiated during their manufacturing (hyper-tempering followed by machining). Measurements and calculations were performed to determine these stresses. Measurements show a difference between circumferential stresses in the depth of a pipe and of an elbow. On the contrary, calculations indicate similar profiles. Thus, the experimental differences cannot be explained by a geometrical effect of the elbow. (J.S.). 4 refs., 5 figs

  2. Using Ecophysiology to Improve Farm Efficiency: Application in Temperate Dairy Grazing Systems

    Directory of Open Access Journals (Sweden)

    David F. Chapman

    2016-04-01

    Full Text Available Information on the physiological ecology of grass-dominant pastures has made a substantial contribution to the development of practices that optimise the amount of feed harvested by grazing animals in temperate livestock systems. However, the contribution of ecophysiology is often under-stated, and the need for further research in this field is sometimes questioned. The challenge for ecophysiolgists, therefore, is to demonstrate how ecophysiological knowledge can help solve significant problems looming for grassland farming in temperate regions while also removing constraints to improved productivity from grazed pastures. To do this, ecophysiological research needs to align more closely with related disciplines, particularly genetics/genomics, agronomy, and farming systems, including systems modelling. This review considers how ecophysiological information has contributed to the development of grazing management practices in the New Zealand dairy industry, an industry that is generally regarded as a world leader in the efficiency with which pasture is grown and utilised for animal production. Even so, there are clear opportunities for further gains in pasture utilisation through the refinement of grazing management practices and the harnessing of those practices to improved pasture plant cultivars with phenotypes that facilitate greater grazing efficiency. Meanwhile, sub-optimal persistence of new pastures continues to constrain productivity in some environments. The underlying plant and population processes associated with this have not been clearly defined. Ecophysiological information, placed in the context of trait identification, grounded in well-designed agronomic studies and linked to plant improvements programmes, is required to address this.

  3. Drilling in tempered glass – modelling and experiments

    DEFF Research Database (Denmark)

    Nielsen, Jens Henrik

    The present paper reports experimentally and numerically obtained results for the process of drilling in tempered glass. The experimental results are drilling depths on the edge in 19mm tempered glass with a known residual stress state measured by a scattered light polariscope. The experiments have...... been modelled using a state-of-the-art model and compared with satisfying result to the performed experiments. The numerical model has been used for a parametric study, investigating the redistribution of residual stresses during the process of drilling. This is done for investigating the possibility...... of applying forces in such holes and thereby being able to mechanically assemble tempered glass without the need of drilling holes before the tempering process. The paper is the result of currently ongoing research and the results should be treated as so....

  4. The effects of warming and nitrogen addition on soil nitrogen cycling in a temperate grassland, northeastern China.

    Directory of Open Access Journals (Sweden)

    Lin-Na Ma

    Full Text Available Both climate warming and atmospheric nitrogen (N deposition are predicted to affect soil N cycling in terrestrial biomes over the next century. However, the interactive effects of warming and N deposition on soil N mineralization in temperate grasslands are poorly understood.A field manipulation experiment was conducted to examine the effects of warming and N addition on soil N cycling in a temperate grassland of northeastern China from 2007 to 2009. Soil samples were incubated at a constant temperature and moisture, from samples collected in the field. The results showed that both warming and N addition significantly stimulated soil net N mineralization rate and net nitrification rate. Combined warming and N addition caused an interactive effect on N mineralization, which could be explained by the relative shift of soil microbial community structure because of fungal biomass increase and strong plant uptake of added N due to warming. Irrespective of strong intra- and inter-annual variations in soil N mineralization, the responses of N mineralization to warming and N addition did not change during the three growing seasons, suggesting independence of warming and N responses of N mineralization from precipitation variations in the temperate grassland.Interactions between climate warming and N deposition on soil N cycling were significant. These findings will improve our understanding on the response of soil N cycling to the simultaneous climate change drivers in temperate grassland ecosystem.

  5. Hot wire TIG temper bead welding for nuclear repairs

    International Nuclear Information System (INIS)

    Lambert, J.A.; Gilston, P.F.

    1989-08-01

    A preliminary assessment has been carried out to determine the suitability of the hot wire tungsten inert gas (TIG) welding process for the repair of thick section, ferritic steel nuclear pressure vessels. The objective has been to identify a hot wire TIG temper bead procedure, suitable for repairs without post weld heat treatment. This procedure involves depositing two weld layers with carefully selected welding parameters such that overlapping thermal cycles produce a refined and tempered heat affected zone, HAZ, microstructure. (author)

  6. The tempered polymerization of human neuroserpin.

    Directory of Open Access Journals (Sweden)

    Rosina Noto

    Full Text Available Neuroserpin, a member of the serpin protein superfamily, is an inhibitor of proteolytic activity that is involved in pathologies such as ischemia, Alzheimer's disease, and Familial Encephalopathy with Neuroserpin Inclusion Bodies (FENIB. The latter belongs to a class of conformational diseases, known as serpinopathies, which are related to the aberrant polymerization of serpin mutants. Neuroserpin is known to polymerize, even in its wild type form, under thermal stress. Here, we study the mechanism of neuroserpin polymerization over a wide range of temperatures by different techniques. Our experiments show how the onset of polymerization is dependent on the formation of an intermediate monomeric conformer, which then associates with a native monomer to yield a dimeric species. After the formation of small polymers, the aggregation proceeds via monomer addition as well as polymer-polymer association. No further secondary mechanism takes place up to very high temperatures, thus resulting in the formation of neuroserpin linear polymeric chains. Most interesting, the overall aggregation is tuned by the co-occurrence of monomer inactivation (i.e. the formation of latent neuroserpin and by a mechanism of fragmentation. The polymerization kinetics exhibit a unique modulation of the average mass and size of polymers, which might suggest synchronization among the different processes involved. Thus, fragmentation would control and temper the aggregation process, instead of enhancing it, as typically observed (e.g. for amyloid fibrillation.

  7. Influence of tempering temperature on mechanical properties of cast steels

    Directory of Open Access Journals (Sweden)

    G. Golański

    2008-12-01

    Full Text Available The paper presents results of research on the influence of tempering temperature on structure and mechanical properties of bainite hardened cast steel: G21CrMoV4 – 6 (L21HMF and G17CrMoV5 – 10 (L17HMF. Investigated cast steels were taken out from internal frames of steam turbines serviced for long time at elevated temperatures. Tempering of the investigated cast steel was carried out within the temperature range of 690 ÷ 730 C (G21CrMoV4 – 6 and 700 ÷ 740 C (G17CrMoV5 – 10. After tempering the cast steels were characterized by a structure of tempered lower bainite with numerous precipitations of carbides. Performed research of mechanical properties has shown that high temperatures of tempering of bainitic structure do not cause decrease of mechanical properties beneath the required minimum.oo It has also been proved that high-temperature tempering (>720 oC ensures high impact energy at the 20% decrease of mechanical properties.

  8. Soil respiration and organic carbon dynamics with grassland conversions to woodlands in temperate china.

    Directory of Open Access Journals (Sweden)

    Wei Wang

    Full Text Available Soils are the largest terrestrial carbon store and soil respiration is the second-largest flux in ecosystem carbon cycling. Across China's temperate region, climatic changes and human activities have frequently caused the transformation of grasslands to woodlands. However, the effect of this transition on soil respiration and soil organic carbon (SOC dynamics remains uncertain in this area. In this study, we measured in situ soil respiration and SOC storage over a two-year period (Jan. 2007-Dec. 2008 from five characteristic vegetation types in a forest-steppe ecotone of temperate China, including grassland (GR, shrubland (SH, as well as in evergreen coniferous (EC, deciduous coniferous (DC and deciduous broadleaved forest (DB, to evaluate the changes of soil respiration and SOC storage with grassland conversions to diverse types of woodlands. Annual soil respiration increased by 3%, 6%, 14%, and 22% after the conversion from GR to EC, SH, DC, and DB, respectively. The variation in soil respiration among different vegetation types could be well explained by SOC and soil total nitrogen content. Despite higher soil respiration in woodlands, SOC storage and residence time increased in the upper 20 cm of soil. Our results suggest that the differences in soil environmental conditions, especially soil substrate availability, influenced the level of annual soil respiration produced by different vegetation types. Moreover, shifts from grassland to woody plant dominance resulted in increased SOC storage. Given the widespread increase in woody plant abundance caused by climate change and large-scale afforestation programs, the soils are expected to accumulate and store increased amounts of organic carbon in temperate areas of China.

  9. [Odocoileus virginianus diet (Artiodactyla: Cervidae) in a temperate forest of Northern Oaxaca, Mexico].

    Science.gov (United States)

    González, Graciela; Briones-Salas, Miguel

    2012-03-01

    The Sierra Madre de Oaxaca region, located in the Northern state of Oaxaca, Mexico, is an area of forest ecosystems subject to high exploitation rates, although in some areas its temperate forests are conserved by indigenous community initiatives that live there. We analyzed the diet of white tailed-deer (Odocoileus virginianus) in the localities of Santa Catarina Lachatao and San Miguel Amatlán from June 1998 to August 1999. Sampling was done during both the wet and dry seasons, and included the observation of browsing traces (238 observations), microhistological analysis of deer feces (28 deer pellet-groups), and two stomach content analysis. The annual diet of white-tailed deer was composed of 42 species from 23 botanical families. The most represented families in the diet of this deer were Fagaceae, Asteraceae, Ericaceae and Fabaceae. There were significant differences in the alpha diversity of the diet during the wet and dry seasons (H'=2.957 and H'=1.832, respectively). The similarity percentage between seasons was 56%. Differences in plant species frequency were significantly higher during the wet season. Herbaceous plants made up the greatest percentage of all the species consumed. The preferred species throughout the year were Senecio sp. (shrub), Sedum dendroideum (herbaceous), Arctostaphylos pungens (shrub) and Satureja macrostema (shrub). Diet species richness was found to be lower than that observed in a tropical forest (Venezuela), tropical dry forest (Mexico) and temperate deciduous and mixed forest (Mexico), but similar to the diet species richness observed in a tropical dry forest (Costa Rica) and temperate coniferous and deciduous forests (USA).

  10. Using flowering and heat-loss models for improving greenhouse energy-use efficiency in annual bedding plant production

    Science.gov (United States)

    In temperate climates, annual bedding plants are typically produced in heated greenhouses from late winter through early summer. Temperature, photoperiod, light intensity, and transplant date are commonly manipulated during commercial production so that plants are in flower for predetermined market ...

  11. Evolution of Microsatellite Loci of Tropical and Temperate Anguilla Eels

    Directory of Open Access Journals (Sweden)

    Mei-Chen Tseng

    2012-04-01

    Full Text Available Anguilla eels are divided into temperate and tropical eels, based on their major distributions. The present study collected two temperate eels, Anguilla japonica and Anguilla anguilla, and two tropical eels, Anguilla marmorata and Anguilla bicolor pacifica, to examine two questions: do temperate and tropical Anguilla eels have different genetic polymorphic patterns?; and do temperate Anguilla japonica and Anguilla anguilla have a closer relationship to each other than to tropical eels? In total, 274 sequences were cloned and sequenced from six conserved microsatellite loci to examine polymorphic patterns of these four catadromous eels. Different mutational events, including substitutions, and repeat-unit deletions and insertions, appeared in major regions, while different point mutations were observed in flanking regions. The results implied that parallel patterns of microsatellite sequences occurred within both tropical and temperate freshwater eels. Consensus flanking sequences of six homologous loci from each of the four species were constructed. Genetic distances ranged from 0.044 (Anguilla bicolor pacifica vs. Anguilla marmorata to 0.061 (Anguilla marmorata vs. Anguilla anguilla. The tree topology suggests the hypothesis of Anguilla japonica and Anguilla anguilla being a sister group must be rejected.

  12. Effects of tempering on internal friction of carbon steels

    International Nuclear Information System (INIS)

    Hoyos, J.J.; Ghilarducci, A.A.; Salva, H.R.; Chaves, C.A.; Velez, J.M.

    2011-01-01

    Research highlights: → Time tempering dependent microstructure of two steels is studied by internal friction. → Internal friction indicates the interactions of dislocations with carbon and carbides. → Internal friction detects the first stage of tempering. → Precipitation hardening is detected by the decrease in the background. - Abstract: Two steels containing 0.626 and 0.71 wt.% carbon have been studied to determine the effects of tempering on the microstructure and the internal friction. The steels were annealed at 1093 K, quenched into water and tempered for 60 min at 423 K, 573 K and 723 K. The increase of the tempering time diminishes the martensite tetragonality due to the redistribution of carbon atoms from octahedrical interstitial sites to dislocations. Internal friction spectrum is decomposed into five peaks and an exponential background, which are attributed to the carbide precipitation and the dislocation relaxation process. Simultaneous presence of peaks P1 and P2 indicates the interaction of dislocations with the segregated carbon and carbide precipitate.

  13. Radiation processing of temperate fruits of Kashmir valley

    International Nuclear Information System (INIS)

    Hussain, Peerzada R.; Meena, Raghuveer S.; Dar, Mohd A.; Wani, Ali M.

    2011-01-01

    Kashmir valley is famous for its temperate horticulture. Main temperate fruits grown commercially in the valley include apple, pear, peach, plum, cherry, strawberry and apricot. These fruits being perishable and susceptible to microbial spoilage, have a short shelf-life. The short shelf-life in an impediment in their transportation and marketing and results in huge losses. Study was carried out at NRL, Srinagar to investigate the effect of gamma irradiation on the keeping quality of most of these fruits. The effect of gamma irradiation alone and in combination with other techniques like controlled low temperature storage, edible polysaccharide coating and calcium chloride treatment was studied in detail. The results revealed that there is a great potential for the use of radiation in extending the storage life of most of the temperate fruits produced in the valley of Kashmir. (author)

  14. Tempered stable distributions stochastic models for multiscale processes

    CERN Document Server

    Grabchak, Michael

    2015-01-01

    This brief is concerned with tempered stable distributions and their associated Levy processes. It is a good text for researchers interested in learning about tempered stable distributions.  A tempered stable distribution is one which takes a stable distribution and modifies its tails to make them lighter. The motivation for this class comes from the fact that infinite variance stable distributions appear to provide a good fit to data in a variety of situations, but the extremely heavy tails of these models are not realistic for most real world applications. The idea of using distributions that modify the tails of stable models to make them lighter seems to have originated in the influential paper of Mantegna and Stanley (1994). Since then, these distributions have been extended and generalized in a variety of ways. They have been applied to a wide variety of areas including mathematical finance, biostatistics,computer science, and physics.

  15. Resistance to small plastic strains during martensite tempering under tension

    Energy Technology Data Exchange (ETDEWEB)

    Zabil' skij, V.V.; Sarrak, V.I. (AN SSSR, Sverdlovsk. Inst. Fiziki Metallov)

    1982-11-01

    The mechanism of plastic deformation of martensite of a series of hardened steels (N18, 20KhG, 50KhFA and others) during tempering under tension and the role of residual internal microstresses and phase transformations are studied. It is shown that martensite low resistance to small plastic deformations during tempering under tension which is usually associated with phase transformations depends as well on the level of residual internal microstresses in the martensite structure. The decrease of resistance to deformation in the course of the decomposition of a solid solution is due to weakening of martensitic matrix because of carbon departure from the solid solution and carbide coarsening. An assumption is made that martensite plastic deformation during tempering under tension is realized at the expense of the directed microplastic deformation in the regions of higher concentration of internal stresses.

  16. Diversity and abundance of photosynthetic sponges in temperate Western Australia

    Directory of Open Access Journals (Sweden)

    Brümmer Franz

    2009-02-01

    Full Text Available Abstract Background Photosynthetic sponges are important components of reef ecosystems around the world, but are poorly understood. It is often assumed that temperate regions have low diversity and abundance of photosynthetic sponges, but to date no studies have investigated this question. The aim of this study was to compare the percentages of photosynthetic sponges in temperate Western Australia (WA with previously published data on tropical regions, and to determine the abundance and diversity of these associations in a range of temperate environments. Results We sampled sponges on 5 m belt transects to determine the percentage of photosynthetic sponges and identified at least one representative of each group of symbionts using 16S rDNA sequencing together with microscopy techniques. Our results demonstrate that photosynthetic sponges are abundant in temperate WA, with an average of 63% of sponge individuals hosting high levels of photosynthetic symbionts and 11% with low to medium levels. These percentages of photosynthetic sponges are comparable to those found on tropical reefs and may have important implications for ecosystem function on temperate reefs in other areas of the world. A diverse range of symbionts sometimes occurred within a small geographic area, including the three "big" cyanobacterial clades, Oscillatoria spongeliae, "Candidatus Synechococcus spongiarum" and Synechocystis species, and it appears that these clades all occur in a wide range of sponges. Additionally, spongin-permeating red algae occurred in at least 7 sponge species. This study provides the first investigation of the molecular phylogeny of rhodophyte symbionts in sponges. Conclusion Photosynthetic sponges are abundant and diverse in temperate WA, with comparable percentages of photosynthetic to non-photosynthetic sponges to tropical zones. It appears that there are three common generalist clades of cyanobacterial symbionts of sponges which occur in a wide

  17. Ultra low carbon bainitic (ULCB) steels after quenching and tempering

    International Nuclear Information System (INIS)

    Lis, A.K.; Lis, J.; Kolan, C.; Jeziorski, L.

    1998-01-01

    The mechanical and Charpy V impact strength properties of new advanced ultra low carbon bainitic (ULBC) steels after water quenching and tempering (WQT) have been investigated. Their chemical compositions are given. The nine continuous cooling transformation diagrams (CCT) of the new ULCB steel grades have been established. The CCT diagrams for ULCB N i steels containing 9% Ni - grade 10N9 and 5% Ni - grade HN5MVNb are given. The comparison between CCT diagrams of 3.5%Ni + 1.5%Cu containing steels grade HSLA 100 and HN3MCu is shown. The effect of the increase in carbon and titanium contents in the chemical composition of ULCB M n steels 04G3Ti, 06G3Ti and 09G3Ti on the kinetics of phase transformations during continuous cooling is presented by the shifting CCT diagrams. The Charpy V impact strength and brittle fracture occurence curves are shown. The effect of tempering temperature on tensile properties of WQT HN3MCu steel is shown and Charpy V impact strength curves after different tempering conditions are shown. The optimum tempering temperatures region of HN3MCu steel for high Charpy V impact toughness at law temperatures - 80 o C(193 K) and -120 o C(153 K) is estimated. The effect of tempering temperature on mechanical properties of HN5MVNb steel is given. The low temperature impact Charpy V toughness of HN5MVNb steel is shown. The optimum range of tempering temperature during 1 hour for high toughness of WQT HN5MVNb steel is given. HN3MCu and HN5MVNb steels after WQT have high yield strength YS≥690 MPa and high Charpy V impact toughness KV≥80 J at -100 o C (173K) and KCV≥50 J/cm 2 at - 120 o C (153K) so they may be used for cryogenic applications

  18. Detection of phytoplasmas of temperate fruit trees.

    Science.gov (United States)

    Laimer, Margit

    2009-01-01

    Phytoplasmas are associated with hundreds of plant diseases globally. Many fruit tree phytoplasmas are transmitted by insect vectors or grafting, are considered quarantine organisms and a major economic threat to orchards. Diagnosis can be difficult, but immunochemical and molecular methods have been developed.

  19. Decoupling of unpolluted temperate forests from rock nutrient sources revealed by natural 87Sr/86Sr and 84Sr tracer addition

    Science.gov (United States)

    Kennedy, Martin J.; Hedin, Lars O.; Derry, Louis A.

    2002-01-01

    An experimental tracer addition of 84Sr to an unpolluted temperate forest site in southern Chile, as well as the natural variation of 87Sr/86Sr within plants and soils, indicates that mechanisms in shallow soil organic horizons are of key importance for retaining and recycling atmospheric cation inputs at scales of decades or less. The dominant tree species Nothofagus nitida feeds nearly exclusively (>90%) on cations of atmospheric origin, despite strong variations in tree size and location in the forest landscape. Our results illustrate that (i) unpolluted temperate forests can become nutritionally decoupled from deeper weathering processes, virtually functioning as atmospherically fed ecosystems, and (ii) base cation turnover times are considerably more rapid than previously recognized in the plant available pool of soil. These results challenge the prevalent paradigm that plants largely feed on rock-derived cations and have important implications for understanding sensitivity of forests to air pollution. PMID:12119394

  20. Heat shock and plant leachates regulate seed germination of the endangered carnivorous plant Drosophyllum lusitanicum

    Directory of Open Access Journals (Sweden)

    S. Gómez-González

    2018-01-01

    Full Text Available In fire-prone ecosystems, many plant species have specialized mechanisms of seed dormancy that ensure a successful recruitment after fire. A well-documented mechanism is the germination stimulated by fire-related cues, such as heat shock and smoke. However, less is known about the role of inhibitory germination signals (e.g. allelopathy in regulating post-fire recruitment. Plant leachates derived from the unburned vegetation can enforce dormancy by means of allelopathic compounds, acting as a signal of unfavourable (highly competitive niche for germination in pyrophyte species. Here, we assessed the separate effects of heat shock and plant leachates on seed germination of Drosophyllum lusitanicum, an endangered carnivorous plant endemic to Mediterranean fire-prone heathlands. We performed a germination experiment in which seeds were subjected to three treatments: (1 5 min at 100 °C, (2 watering with plant leachate, and (3 control. Germination rate and seed viability was determined after 63 days. Heat shock stimulated seed germination in D. lusitanicum while plant leachates had inhibitory germination effects without reducing seed viability. Thus, both positive and negative signals could be involved in its successful post-fire recruitment. Fire would break seed dormancy and stimulate seed germination of D. lusitanicum through high temperatures, but also by eliminating allelochemical compounds from the soil. These results help to understand the population dynamics patterns found for D. lusitanicum in natural populations, and highlight the role of fire in the ecology and conservation of this endangered species. Seed dormancy imposed by plant-derived leachates as an adaptive mechanism should be considered more in fire ecology theory.

  1. SWFSC/MMTD/CCE: Leatherback Use of Temperate Habitat (LUTH) 2008

    Data.gov (United States)

    National Oceanic and Atmospheric Administration, Department of Commerce — The Leatherback Use of Temperate Habitat (LUTH) survey is an ecosystem assessment of temperate foraging habitats of endangered leatherback turtles off the coast of...

  2. Direct dating and identity of fiber temper in pre-contact Bushman (Masarwa) pottery

    CSIR Research Space (South Africa)

    Bollong, CA

    1993-01-01

    Full Text Available , ashed grass, or absorbed blood. Visual characteristics rule out the possibility of animal dung temper. Blood and fat residues absorbed by grass temper could be the combined cause of anomalous Q (13) C and high P values....

  3. Root-Associated Fungi Shared Between Arbuscular Mycorrhizal and Ectomycorrhizal Conifers in a Temperate Forest.

    Science.gov (United States)

    Toju, Hirokazu; Sato, Hirotoshi

    2018-01-01

    Arbuscular mycorrhizal and ectomycorrhizal symbioses are among the most important drivers of terrestrial ecosystem dynamics. Historically, the two types of symbioses have been investigated separately because arbuscular mycorrhizal and ectomycorrhizal plant species are considered to host discrete sets of fungal symbionts (i.e., arbuscular mycorrhizal and ectomycorrhizal fungi, respectively). Nonetheless, recent studies based on high-throughput DNA sequencing technologies have suggested that diverse non-mycorrhizal fungi (e.g., endophytic fungi) with broad host ranges play roles in relationships between arbuscular mycorrhizal and ectomycorrhizal plant species in forest ecosystems. By analyzing an Illumina sequencing dataset of root-associated fungi in a temperate forest in Japan, we statistically examined whether co-occurring arbuscular mycorrhizal ( Chamaecyparis obtusa ) and ectomycorrhizal ( Pinus densiflora ) plant species could share non-mycorrhizal fungal communities. Among the 919 fungal operational taxonomic units (OTUs) detected, OTUs in various taxonomic lineages were statistically designated as "generalists," which associated commonly with both coniferous species. The list of the generalists included fungi in the genera Meliniomyces, Oidiodendron, Cladophialophora, Rhizodermea, Penicillium , and Mortierella . Meanwhile, our statistical analysis also detected fungi preferentially associated with Chamaecyparis (e.g., Pezicula ) or Pinus (e.g., Neolecta ). Overall, this study provides a basis for future studies on how arbuscular mycorrhizal and ectomycorrhizal plant species interactively drive community- or ecosystem-scale processes. The physiological functions of the fungi highlighted in our host-preference analysis deserve intensive investigations for understanding their roles in plant endosphere and rhizosphere.

  4. Factors affecting the ozone sensitivity of temperate European grasslands: An overview

    Energy Technology Data Exchange (ETDEWEB)

    Bassin, S. [Agroscope FAL Reckenholz, Swiss Federal Research Station for Agroecology and Agriculture, Air Pollution/Climate Group, Reckenholzstrasse 191, CH-8046 Zurich (Switzerland)]. E-mail: seraina.bassin@fal.admin.ch; Volk, M. [Agroscope FAL Reckenholz, Swiss Federal Research Station for Agroecology and Agriculture, Air Pollution/Climate Group, Reckenholzstrasse 191, CH-8046 Zurich (Switzerland); Fuhrer, J. [Agroscope FAL Reckenholz, Swiss Federal Research Station for Agroecology and Agriculture, Air Pollution/Climate Group, Reckenholzstrasse 191, CH-8046 Zurich (Switzerland)

    2007-04-15

    This overview of experimentally induced effects of ozone aims to identify physiological and ecological principles, which can be used to classify the sensitivity to ozone of temperate grassland communities in Europe. The analysis of data from experiments with single plants, binary mixtures and multi-species communities illustrates the difficulties to relate individual responses to communities, and thus to identify grassland communities most at risk. Although there is increasing evidence that communities can be separated into broad classes of ozone sensitivity, the database from experiments under realistic conditions with representative systems is too small to draw firm conclusions. But it appears that risk assessments, based on results from individuals or immature mixtures exposed in chambers, are only applicable to intensively managed, productive grasslands, and that the risk of ozone damage for most of perennial grasslands with lower productivity tends to be less than previously expected. - An overview of experimentally induced ozone effects suggests that temperate grasslands could be separated into broad classes of ozone sensitivity based on physiological and ecological principles.

  5. Factors affecting the ozone sensitivity of temperate European grasslands: An overview

    International Nuclear Information System (INIS)

    Bassin, S.; Volk, M.; Fuhrer, J.

    2007-01-01

    This overview of experimentally induced effects of ozone aims to identify physiological and ecological principles, which can be used to classify the sensitivity to ozone of temperate grassland communities in Europe. The analysis of data from experiments with single plants, binary mixtures and multi-species communities illustrates the difficulties to relate individual responses to communities, and thus to identify grassland communities most at risk. Although there is increasing evidence that communities can be separated into broad classes of ozone sensitivity, the database from experiments under realistic conditions with representative systems is too small to draw firm conclusions. But it appears that risk assessments, based on results from individuals or immature mixtures exposed in chambers, are only applicable to intensively managed, productive grasslands, and that the risk of ozone damage for most of perennial grasslands with lower productivity tends to be less than previously expected. - An overview of experimentally induced ozone effects suggests that temperate grasslands could be separated into broad classes of ozone sensitivity based on physiological and ecological principles

  6. Biotechnology of temperate fruit trees and grapevines.

    Science.gov (United States)

    Laimer, Margit; Mendonça, Duarte; Maghuly, Fatemeh; Marzban, Gorji; Leopold, Stephan; Khan, Mahmood; Balla, Ildiko; Katinger, Hermann

    2005-01-01

    Challenges concerning fruit trees and grapevines as long lived woody perennial crops require adapted biotechnological approaches, if solutions are to be found within a reasonable time frame. These challenges are represented by the need for correct identification of genetic resources, with the foreseen use either in conservation or in breeding programmes. Molecular markers provide most accurate information and will be the major solution for questions about plant breeders rights. Providing healthy planting material and rapid detection of newly introduced pathogens by reliable methods involving serological and molecular biological tools will be a future challenge of increases importance, given the fact that plant material travels freely in the entire European Union. But also new breeding goals and transgenic solutions are part of the biotechnological benefits, e.g. resistance against biotic and abiotic stress factors, modified growth habits, modified nutritional properties and altered processing and storage qualities. The successful characterization of transgenic grapevines and stone fruit trees carrying genes of viral origin in different vectors constructed under ecological consideration, will be presented. Beyond technical feasibility, efficiency of resistance, environmental safety and Intellectual Property Rights, also public acceptance needs consideration and has been addressed in a specific project. The molecular determination of internal quality parameters of food can also be addressed by the use of biotechnological tools. Patient independent detection tools for apple allergens have been developed and should allow to compare fruits from different production systems, sites, and genotypes for their content of health threatening compounds.

  7. Multiyear greenhouse gas balances at a rewetted temperate peatland.

    Science.gov (United States)

    Wilson, David; Farrell, Catherine A; Fallon, David; Moser, Gerald; Müller, Christoph; Renou-Wilson, Florence

    2016-12-01

    Drained peat soils are a significant source of greenhouse gas (GHG) emissions to the atmosphere. Rewetting these soils is considered an important climate change mitigation tool to reduce emissions and create suitable conditions for carbon sequestration. Long-term monitoring is essential to capture interannual variations in GHG emissions and associated environmental variables and to reduce the uncertainty linked with GHG emission factor calculations. In this study, we present GHG balances: carbon dioxide (CO 2 ), methane (CH 4 ) and nitrous oxide (N 2 O) calculated for a 5-year period at a rewetted industrial cutaway peatland in Ireland (rewetted 7 years prior to the start of the study); and compare the results with an adjacent drained area (2-year data set), and with ten long-term data sets from intact (i.e. undrained) peatlands in temperate and boreal regions. In the rewetted site, CO 2 exchange (or net ecosystem exchange (NEE)) was strongly influenced by ecosystem respiration (R eco ) rather than gross primary production (GPP). CH 4 emissions were related to soil temperature and either water table level or plant biomass. N 2 O emissions were not detected in either drained or rewetted sites. Rewetting reduced CO 2 emissions in unvegetated areas by approximately 50%. When upscaled to the ecosystem level, the emission factors (calculated as 5-year mean of annual balances) for the rewetted site were (±SD) -104 ± 80 g CO 2 -C m -2  yr -1 (i.e. CO 2 sink) and 9 ± 2 g CH 4 -C m -2  yr -1 (i.e. CH 4 source). Nearly a decade after rewetting, the GHG balance (100-year global warming potential) had reduced noticeably (i.e. less warming) in comparison with the drained site but was still higher than comparative intact sites. Our results indicate that rewetted sites may be more sensitive to interannual changes in weather conditions than their more resilient intact counterparts and may switch from an annual CO 2 sink to a source if triggered by slightly drier

  8. Tempered fractional time series model for turbulence in geophysical flows

    Science.gov (United States)

    Meerschaert, Mark M.; Sabzikar, Farzad; Phanikumar, Mantha S.; Zeleke, Aklilu

    2014-09-01

    We propose a new time series model for velocity data in turbulent flows. The new model employs tempered fractional calculus to extend the classical 5/3 spectral model of Kolmogorov. Application to wind speed and water velocity in a large lake are presented, to demonstrate the practical utility of the model.

  9. Tempered fractional time series model for turbulence in geophysical flows

    International Nuclear Information System (INIS)

    Meerschaert, Mark M; Sabzikar, Farzad; Phanikumar, Mantha S; Zeleke, Aklilu

    2014-01-01

    We propose a new time series model for velocity data in turbulent flows. The new model employs tempered fractional calculus to extend the classical 5/3 spectral model of Kolmogorov. Application to wind speed and water velocity in a large lake are presented, to demonstrate the practical utility of the model. (paper)

  10. Joint measurement of risk aversion, prudence, and temperance

    NARCIS (Netherlands)

    Ebert, S.; Wiesen, D.

    Risk aversion—but also the higher-order risk preferences of prudence and temperance—are fundamental concepts in the study of economic decision making. We propose a method to jointly measure the intensity of risk aversion, prudence, and temperance. Our theoretical approach is to define risk

  11. The Fracture Process of Tempered Soda-Lime-Silica Glass

    DEFF Research Database (Denmark)

    Nielsen, Jens Henrik; Olesen, John Forbes; Stang, Henrik

    2009-01-01

    This work presents experimental observations of the characteristic fracture process of tempered glass. Square specimens with a side length of 300 mm, various thicknesses and a residual stress state characterized by photoelastic measurements were used. Fracture was initiated using a 2.5 mm diamond...

  12. Tempered orbital energies and the potential curve within a molecule

    International Nuclear Information System (INIS)

    Anno, T.; Sakai, Y.

    1979-01-01

    It is shown that the sum of the tempered orbital energies behaves much the same way as does the some of the Hartree--Fock orbital energies. The H 2 moluecule in its minimum basis set description is chosen as an example and comparison of orbital energies is carried out

  13. Significance of rate of work hardening in tempered martensite embrittlement

    International Nuclear Information System (INIS)

    Pietikainen, J.

    1995-01-01

    The main explanations for tempered martensite embrittlement are based on the effects of impurities and cementite precipitation on the prior austenite grain boundaries. There are some studies where the rate of work hardening is proposed as a potential reason for the brittleness. One steel was studied by means of a specially developed precision torsional testing device. The test steel had a high Si and Ni content so ε carbide and Fe 3 C appear in quite different tempering temperature ranges. The M S temperature is low enough so that self tempering does not occur. With the testing device it was possible to obtain the true stress - true strain curves to very high deformations. The minimum toughness was always associated with the minimum of rate of work hardening. The change of deformed steel volume before the loss of mechanical stability is proposed as at least one reason for tempered martensite embrittlement. The reasons for the minimum of the rate of work hardening are considered. (orig.)

  14. Tempered Water Lower Port Connector Structural Analysis Verification

    International Nuclear Information System (INIS)

    CREA, B.A.

    2000-01-01

    Structural analysis of the lower port connection of the Tempered Water System of the Cold Vacuum Drying Facility was performed. Subsequent detailed design changes to enhance operability resulted in the need to re-evaluate the bases of the original analysis to verify its continued validity. This evaluation is contained in Appendix A of this report. The original evaluation is contained in Appendix B

  15. Effects of temperate agriculture on neotropical migrant landbirds

    Science.gov (United States)

    Nicholas L. Rodenhouse; Louis B. Best; Raymond J. O' Connor; Eric K. Bollinger

    1993-01-01

    The ecology of Neotropical migrant landbirds in temperate farmland is reviewed to develop management recommendations for the conservation of migrants. Migrants constitute about 71% of bird species using farmland and 86% of bird species nesting there. The number and abundances of Neotropical migrants using farmland are greatest in uncultivated edges with trees and...

  16. Effect of climate change on temperate forest ecosystems

    NARCIS (Netherlands)

    Brolsma, R.J.

    2010-01-01

    In temperate climates groundwater can have a strong effect on vegetation, because it can influence the spatio-temporal distribution of soil moisture and therefore water and oxygen stress of vegetation. Current IPCC climate projections based on CO2 emission scenarios show a global temperature rise

  17. Solid-wood production from temperate eucalypt plantations: a ...

    African Journals Online (AJOL)

    Since 1988, there has been a major focus in Tasmania on research for the management of temperate eucalypt plantations for solid wood. This coincided with the formal transfer of large areas of native forest that had previously been part of the production forest estate into reserves, a decision that triggered the establishment ...

  18. Changes in soil organic matter and net nitrogen mineralization in heathland soils, after removal, addition or replacement of litter from Erica tetralix or Molinia caerulea.

    NARCIS (Netherlands)

    Vuuren, van M.M.I.; Berendse, F.

    1993-01-01

    The effects of different litter input rates and of different types of litter on soil organic matter accumulation and net N mineralization were investigated in plant communities dominated by Erica tetralix L. or Molinia caerulea (L.) Moench. Plots in which the litter on the soil had repeatedly been

  19. Groundwater flow modelling of periods with temperate climate conditions - Forsmark

    Energy Technology Data Exchange (ETDEWEB)

    Joyce, Steven; Simpson, Trevor; Hartley, Lee; Applegate, David; Hoek, Jaap; Jackson, Peter; Swan, David (Serco Technical Consulting Services (United Kingdom)); Marsic, Niko (Kemakta Konsult AB (Sweden)); Follin, Sven (SF GeoLogic AB (Sweden))

    2010-11-15

    As a part of the license application for a final repository for spent nuclear fuel at Forsmark, the Swedish Nuclear Fuel and Waste Management Company (SKB) has undertaken a series of groundwater flow modelling studies. These represent time periods with different climate conditions and the simulations carried out contribute to the overall evaluation of the repository design and long-term radiological safety. This report concerns the modelling of a repository at the Forsmark site during temperate conditions; i.e. from post-closure and throughout the temperate period up until the receding shoreline leaves the modelling domain at around 12,000 AD. The collation and implementation of onsite hydrogeological and hydrogeochemical data from previous reports are used in the construction of a hydrogeological base case (reference case conceptualisation) and then in an examination of various areas of uncertainty within the current understanding by a series of model variants. The hydrogeological base case models at three different scales, 'repository', 'site' and 'regional', make use of continuous porous medium (CPM), equivalent continuous porous medium (ECPM) and discrete fracture network (DFN) models. The use of hydrogeological models allow for the investigation of the groundwater flow from a deep disposal facility to the biosphere and for the calculation of performance measures that will provide an input to the site performance assessment. The focus of the study described in this report has been to perform numerical simulations of the hydrogeological system from post-closure and throughout the temperate period. Besides providing quantitative results for the immediate temperate period following post-closure, these results are also intended to give a qualitative indication of the evolution of the groundwater system during future temperate periods within an ongoing cycle of glacial/inter-glacial events

  20. Root growth and N dynamics in response to multi-year experimental warming, summer drought and elevated CO2 in a mixed heathland-grass ecosystem

    DEFF Research Database (Denmark)

    Arndal, M. F.; Schmidt, I. K.; Kongstad, J.

    2013-01-01

    growth would be matched by an increase in root nutrient uptake of NH4+-N and NO3- -N. Root growth was significantly increased by elevated CO2. The roots, however, did not fully compensate for the higher growth with a similar increase in nitrogen uptake per unit of root mass. Hence the nitrogen...... concentration in roots was decreased in elevated CO2, whereas the biomass N pool was unchanged or even increased. The higher net root production in elevated CO2 might be a strategy for the plants to cope with increased nutrient demand leading to a long-term increase in N uptake on a whole-plant basis. Drought...... reduced grass root biomass and N uptake, especially when combined with warming, but CO2 was the most pronounced main factor effect. Several significant interactions of the treatments were found, which indicates that the responses were nonadditive and that changes to multiple environmental changes cannot...

  1. Root morphology and mycorrhizal symbioses together shape nutrient foraging strategies of temperate trees.

    Science.gov (United States)

    Chen, Weile; Koide, Roger T; Adams, Thomas S; DeForest, Jared L; Cheng, Lei; Eissenstat, David M

    2016-08-02

    Photosynthesis by leaves and acquisition of water and minerals by roots are required for plant growth, which is a key component of many ecosystem functions. Although the role of leaf functional traits in photosynthesis is generally well understood, the relationship of root functional traits to nutrient uptake is not. In particular, predictions of nutrient acquisition strategies from specific root traits are often vague. Roots of nearly all plants cooperate with mycorrhizal fungi in nutrient acquisition. Most tree species form symbioses with either arbuscular mycorrhizal (AM) or ectomycorrhizal (EM) fungi. Nutrients are distributed heterogeneously in the soil, and nutrient-rich "hotspots" can be a key source for plants. Thus, predicting the foraging strategies that enable mycorrhizal root systems to exploit these hotspots can be critical to the understanding of plant nutrition and ecosystem carbon and nutrient cycling. Here, we show that in 13 sympatric temperate tree species, when nutrient availability is patchy, thinner root species alter their foraging to exploit patches, whereas thicker root species do not. Moreover, there appear to be two distinct pathways by which thinner root tree species enhance foraging in nutrient-rich patches: AM trees produce more roots, whereas EM trees produce more mycorrhizal fungal hyphae. Our results indicate that strategies of nutrient foraging are complementary among tree species with contrasting mycorrhiza types and root morphologies, and that predictable relationships between below-ground traits and nutrient acquisition emerge only when both roots and mycorrhizal fungi are considered together.

  2. Scientists' declaration to conserve Canada's coastal temperate rainforests

    Energy Technology Data Exchange (ETDEWEB)

    NONE

    2000-06-01

    Immediate cessation of large-scale clearcut logging in the watersheds of the northwest temperate rainforests of Canada has been called for by a group of 400+ scientists, citing the threat of destruction to a variety of plant and animal life and the scientific importance of the area. The declaration affirms the ecological importance of these areas, the disruption and destruction of vulnerable ecosystems of global significance that has already taken place, and urges the government to put an end to clearcut logging in these areas, to be replaced by ecosystem-based forestry which includes areas of biological refuge, maintains the ecological characteristics of the original forests, respects and considers the traditional knowledge of First Nations, and provides for the long-term sustainability of coastal communities who depend on fishing and forestry for their economic and cultural well-being. The declaration includes the names of the 400+ signatories.

  3. Solar Radiation Determines Site Occupancy of Coexisting Tropical and Temperate Deer Species Introduced to New Zealand Forests.

    Directory of Open Access Journals (Sweden)

    Robert B Allen

    Full Text Available Assemblages of introduced taxa provide an opportunity to understand how abiotic and biotic factors shape habitat use by coexisting species. We tested hypotheses about habitat selection by two deer species recently introduced to New Zealand's temperate rainforests. We hypothesised that, due to different thermoregulatory abilities, rusa deer (Cervus timorensis; a tropical species would prefer warmer locations in winter than red deer (Cervus elaphus scoticus; a temperate species. Since adult male rusa deer are aggressive in winter (the rut, we also hypothesised that rusa deer and red deer would not use the same winter locations. Finally, we hypothesised that in summer both species would prefer locations with fertile soils that supported more plant species preferred as food. We used a 250 × 250 m grid of 25 remote cameras to collect images in a 100-ha montane study area over two winters and summers. Plant composition, solar radiation, and soil fertility were also determined for each camera location. Multiseason occupancy models revealed that direct solar radiation was the best predictor of occupancy and detection probabilities for rusa deer in winter. Multistate, multiseason occupancy models provided strong evidence that the detection probability of adult male rusa deer was greater in winter and when other rusa deer were present at a location. Red deer mostly vacated the study area in winter. For the one season that had sufficient camera images of both species (summer 2011 to allow two-species occupancy models to be fitted, the detection probability of rusa deer also increased with solar radiation. Detection probability also varied with plant composition for both deer species. We conclude that habitat use by coexisting tropical and temperate deer species in New Zealand likely depends on the interplay between the thermoregulatory and behavioural traits of the deer and the abiotic and biotic features of the habitat.

  4. Diversity patterns of ground beetles and understory vegetation in mature, secondary, and plantation forest regions of temperate northern China.

    Science.gov (United States)

    Zou, Yi; Sang, Weiguo; Wang, Shunzhong; Warren-Thomas, Eleanor; Liu, Yunhui; Yu, Zhenrong; Wang, Changliu; Axmacher, Jan Christoph

    2015-02-01

    Plantation and secondary forests form increasingly important components of the global forest cover, but our current knowledge about their potential contribution to biodiversity conservation is limited. We surveyed understory plant and carabid species assemblages at three distinct regions in temperate northeastern China, dominated by mature forest (Changbaishan Nature Reserve, sampled in 2011 and 2012), secondary forest (Dongling Mountain, sampled in 2011 and 2012), and forest plantation habitats (Bashang Plateau, sampled in 2006 and 2007), respectively. The α-diversity of both taxonomic groups was highest in plantation forests of the Bashang Plateau. Beetle α-diversity was lowest, but plant and beetle species turnover peaked in the secondary forests of Dongling Mountain, while habitats in the Changbaishan Nature Reserve showed the lowest turnover rates for both taxa. Changbaishan Nature Reserve harbored the highest proportion of forest specialists. Our results suggest that in temperate regions of northern China, the protected larch plantation forest established over extensive areas might play a considerable role in maintaining a high biodiversity in relation to understory herbaceous plant species and carabid assemblages, which can be seen as indicators of forest disturbance. The high proportion of phytophagous carabids and the rarity of forest specialists reflect the relatively homogenous, immature status of the forest ecosystems on the Bashang Plateau. China's last remaining large old-growth forests like the ones on Changbaishan represent stable, mature ecosystems which require particular conservation attention.

  5. Possibilities for using higher-tensile, water quenched and tempered AlSiMn fine-grained structural steel for reactor containments

    International Nuclear Information System (INIS)

    Wallner, F.

    1976-01-01

    On water quenching and tempering of weldable AlSiMn structural steels, particularly the grain refining process is made use of, i.e. that measure with the poorest influence on the weldability of steel. Precipitation hardening due to water quenching is, on subsequent tempering, set off to a large extent by means of precipitation resp. coagulation of iron carbides. Minimum yield points up to 580 N/mm 2 and, simultaneously, good viscosity can be obtained by means of water quenching from austeritic temperature and tempering between 550 0 C and 650 0 C, depending on tempering temperatures and sheet thickness. In the paper at hand, results are given, obtained from tests and experience with the steel Aldur 50/65 (the first figure indicates minimum yield points, the second one minimum tensile strength on sheet thickness up to 30 mm). These results are assumed to be essential, also in connection with the construction and working conditions of nuclear power plants. (orig./RW) [de

  6. Bifurcation dynamics of the tempered fractional Langevin equation

    Energy Technology Data Exchange (ETDEWEB)

    Zeng, Caibin, E-mail: macbzeng@scut.edu.cn; Yang, Qigui, E-mail: qgyang@scut.edu.cn [School of Mathematics, South China University of Technology, Guangzhou 510640 (China); Chen, YangQuan, E-mail: ychen53@ucmerced.edu [MESA LAB, School of Engineering, University of California, Merced, 5200 N. Lake Road, Merced, California 95343 (United States)

    2016-08-15

    Tempered fractional processes offer a useful extension for turbulence to include low frequencies. In this paper, we investigate the stochastic phenomenological bifurcation, or stochastic P-bifurcation, of the Langevin equation perturbed by tempered fractional Brownian motion. However, most standard tools from the well-studied framework of random dynamical systems cannot be applied to systems driven by non-Markovian noise, so it is desirable to construct possible approaches in a non-Markovian framework. We first derive the spectral density function of the considered system based on the generalized Parseval's formula and the Wiener-Khinchin theorem. Then we show that it enjoys interesting and diverse bifurcation phenomena exchanging between or among explosive-like, unimodal, and bimodal kurtosis. Therefore, our procedures in this paper are not merely comparable in scope to the existing theory of Markovian systems but also provide a possible approach to discern P-bifurcation dynamics in the non-Markovian settings.

  7. Linking Well-Tempered Metadynamics Simulations with Experiments

    Science.gov (United States)

    Barducci, Alessandro; Bonomi, Massimiliano; Parrinello, Michele

    2010-01-01

    Abstract Linking experiments with the atomistic resolution provided by molecular dynamics simulations can shed light on the structure and dynamics of protein-disordered states. The sampling limitations of classical molecular dynamics can be overcome using metadynamics, which is based on the introduction of a history-dependent bias on a small number of suitably chosen collective variables. Even if such bias distorts the probability distribution of the other degrees of freedom, the equilibrium Boltzmann distribution can be reconstructed using a recently developed reweighting algorithm. Quantitative comparison with experimental data is thus possible. Here we show the potential of this combined approach by characterizing the conformational ensemble explored by a 13-residue helix-forming peptide by means of a well-tempered metadynamics/parallel tempering approach and comparing the reconstructed nuclear magnetic resonance scalar couplings with experimental data. PMID:20441734

  8. Reversible temper brittleness on tensile tests at room temperature

    International Nuclear Information System (INIS)

    Quadros, N.F. de; Cabral, U.Q.

    1976-01-01

    Tensile tests were carried out on unnotched test pieces at room temperature and three strain rates: 2,5x10 -4 , 2,5x10 -3 and 1,0x10 -2 s -1 in a low alloy No-Cr-Mo steel to observe the variation in its mechanical properties with the occurrence of reversible temper brittleness. The brittle samples showed a sensitivity of 50 0 C in a 48 hour heat treatment at 500 0 C. The tests showed that at the strain rate of 2,5x10 -4 s -1 there are statistically significant differences between the elongations of the material in the brittle and the nonbrittle and regenerated states. A short review of reversible temper brittleness is given and a theory suggested for the mechanism [pt

  9. The tempered stable process with infinitely divisible inverse subordinators

    International Nuclear Information System (INIS)

    Wyłomańska, Agnieszka

    2013-01-01

    In the last decade processes driven by inverse subordinators have become extremely popular. They have been used in many different applications, especially for data with observable constant time periods. However, the classical model, i.e. the subordinated Brownian motion, can be inappropriate for the description of observed phenomena that exhibit behavior not adequate for Gaussian systems. Therefore, in this paper we extend the classical approach and replace the Brownian motion by the tempered stable process. Moreover, on the other hand, as an extension of the classical model, we analyze the general class of inverse subordinators. We examine the main properties of the tempered stable process driven by inverse subordinators from the infinitely divisible class of distributions. We show the fractional Fokker–Planck equation of the examined process and the asymptotic behavior of the mean square displacement for two cases of subordinators. Additionally, we examine how an external force can influence the examined characteristics. (paper)

  10. Effect of tempering on corrosion resistance of cast aluminium bronzes

    International Nuclear Information System (INIS)

    Aaltonen, P.; Klemetti, K.; Haenninen, H.

    1985-01-01

    The subject of this study is corrosion resistance of aluminium bronzes, which are copper base alloys containing aluminium up to 12% with additions of nickel, iron and manganese. The main conclutions that can be drawn are: (1) The dealloying corrosion resistance of nickel-aluminium bronze is much better than that of aluminium bronze with iron and manganese additions, but it is not immune; (2) The dealloying corrosion resistance of aluminium bronzes can be improved by appropiate heat treatments. The best properties were obtained by temperering between 600 and 800 deg C, depending on the initial microstructure; (3) In crevice conditions, where local acidification can occur, dealloying of aluminium bronzes is a consequence of the preferential attack of aluminium-rich phases. By appropriate tempering, a uniform distribution of aluminium-rich phases is obtained and the continous path for selective corrosion is not formed

  11. Interspecific variation in total phenolic content in temperate brown algae

    Directory of Open Access Journals (Sweden)

    Anna Maria Mannino

    2017-09-01

    Full Text Available Marine algae synthesize secondary metabolites such as polyphenols that function as defense and protection mechanisms. Among brown algae, Fucales and Dictyotales (Phaeophyceae contain the highest levels of phenolic compounds, mainly phlorotannins, that play multiple roles. Four temperate brown algae (Cystoseira amentacea, Cystoseira compressa, Dictyopteris polypodioides and Padina pavonica were studied for total phenolic contents. Total phenolic content was determined colorimetrically with the Folin-Ciocalteu reagent. Significant differences in total phenolic content were observed between leathery and sheetlike algae and also within each morphological group. Among the four species, the sheet-like alga D. polypodioides, living in the upper infralittoral zone, showed the highest concentration of phenolic compounds. These results are in agreement with the hypothesis that total phenolic content in temperate brown algae is influenced by a combination of several factors, such as growth form, depth, and exposition to solar radiation.

  12. Hurricane Impacts to Tropical and Temperate Forest Landscapes

    OpenAIRE

    Boose, Emery Robert; Foster, David Russell; Fluet, Marcheterre

    1994-01-01

    Hurricanes represent an important natural disturbance process to tropical and temperate forests in many coastal areas of the world. The complex patterns of damage created in forests by hurricane winds result from the interaction of meteorological, physiographic, and biotic factors on a range of spatial scales. To improve our understanding of these factors and of the role of catastrophic hurricane wind as a disturbance process, we take an integrative approach. A simple meteorological model (HU...

  13. First description of underwater acoustic diversity in three temperate ponds

    Directory of Open Access Journals (Sweden)

    Camille Desjonquères

    2015-11-01

    Full Text Available The past decade has produced an increased ecological interest in sonic environments, or soundscapes. However, despite this rise in interest and technological improvements that allow for long-term acoustic surveys in various environments, some habitats’ soundscapes remain to be explored. Ponds, and more generally freshwater habitats, are one of these acoustically unexplored environments. Here we undertook the first long term acoustic monitoring of three temperate ponds in France. By aural and visual inspection of a selection of recordings, we identified 48 different sound types, and according to the rarefaction curves we calculated, more sound types are likely present in one of the three ponds. The richness of sound types varied significantly across ponds. Surprisingly, there was no pond-to-pond daily consistency of sound type richness variation; each pond had its own daily patterns of activity. We also explored the possibility of using six acoustic diversity indices to conduct rapid biodiversity assessments in temperate ponds. We found that all indices were sensitive to the background noise as estimated through correlations with the signal-to-noise ratio (SNR. However, we determined that the AR index could be a good candidate to measure acoustic diversities using partial correlations with the SNR as a control variable. Yet, research is still required to automatically compute the SNR in order to apply this index on a large data set of recordings. The results showed that these three temperate ponds host a high level of acoustic diversity in which the soundscapes were variable not only between but also within the ponds. The sources producing this diversity of sounds and the drivers of difference in daily song type richness variation both require further investigation. Such research would yield insights into the biodiversity and ecology of temperate ponds.

  14. Groundwater flow modelling of periods with temperate climate conditions - Laxemar

    International Nuclear Information System (INIS)

    Joyce, Steven; Simpson, Trevor; Hartley, Lee; Applegate, David; Hoek, Jaap; Jackson, Peter; Roberts, David; Swan, David; Gylling, Bjoern; Marsic, Niko; Rhen, Ingvar

    2010-12-01

    As a part of the license application for a final repository for spent nuclear fuel at Forsmark, the Swedish Nuclear Fuel and Waste Management Company (SKB) has undertaken a series of groundwater flow modelling studies. These represent time periods with different hydraulic conditions and the simulations carried out contribute to the overall evaluation of the repository design and long-term radiological safety. This report concerns the modelling of a repository at the Laxemar-Simpevarp site during temperate climate conditions as a comparison to corresponding modelling carried out for Forsmark /Joyce et al. 2010/. The collation and implementation of onsite hydrogeological and hydrogeochemical data from previous reports are used in the construction of a Hydrogeological base case (reference case conceptualisation) and then an examination of various areas of uncertainty within the current understanding by a series of model variants. The Hydrogeological base case models at three different scales, 'repository', 'site' and 'regional' make use of a discrete fracture network (DFN) and equivalent continuous porous medium (ECPM) models. The use of hydrogeological models allow for the investigation of the groundwater flow from a deep disposal facility to the biosphere and for the calculation of performance measures that will provide an input to the site performance assessment. The focus of the study described in this report has been to perform numerical simulations of the hydrogeological system from post-closure and throughout the temperate period up until the receding shoreline leaves the modelling domain at around 15,000 AD. Besides providing quantitative results for the immediate temperate period following post-closure, these results are also intended to give a qualitative indication of the evolution of the groundwater system during future temperate periods within an ongoing cycle of glacial/inter-glacial events

  15. Groundwater flow modelling of periods with temperate climate conditions - Laxemar

    Energy Technology Data Exchange (ETDEWEB)

    Joyce, Steven; Simpson, Trevor; Hartley, Lee; Applegate, David; Hoek, Jaap; Jackson, Peter; Roberts, David; Swan, David (Serco Technical Consulting Services (United Kingdom)); Gylling, Bjoern; Marsic, Niko (Kemakta Konsult AB, Stockholm (Sweden)); Rhen, Ingvar (SWECO Environment AB, Falun (Sweden))

    2010-12-15

    As a part of the license application for a final repository for spent nuclear fuel at Forsmark, the Swedish Nuclear Fuel and Waste Management Company (SKB) has undertaken a series of groundwater flow modelling studies. These represent time periods with different hydraulic conditions and the simulations carried out contribute to the overall evaluation of the repository design and long-term radiological safety. This report concerns the modelling of a repository at the Laxemar-Simpevarp site during temperate climate conditions as a comparison to corresponding modelling carried out for Forsmark /Joyce et al. 2010/. The collation and implementation of onsite hydrogeological and hydrogeochemical data from previous reports are used in the construction of a Hydrogeological base case (reference case conceptualisation) and then an examination of various areas of uncertainty within the current understanding by a series of model variants. The Hydrogeological base case models at three different scales, 'repository', 'site' and 'regional' make use of a discrete fracture network (DFN) and equivalent continuous porous medium (ECPM) models. The use of hydrogeological models allow for the investigation of the groundwater flow from a deep disposal facility to the biosphere and for the calculation of performance measures that will provide an input to the site performance assessment. The focus of the study described in this report has been to perform numerical simulations of the hydrogeological system from post-closure and throughout the temperate period up until the receding shoreline leaves the modelling domain at around 15,000 AD. Besides providing quantitative results for the immediate temperate period following post-closure, these results are also intended to give a qualitative indication of the evolution of the groundwater system during future temperate periods within an ongoing cycle of glacial/inter-glacial events

  16. Efficient assignment of the temperature set for Parallel Tempering

    International Nuclear Information System (INIS)

    Guidetti, M.; Rolando, V.; Tripiccione, R.

    2012-01-01

    We propose a simple algorithm able to identify a set of temperatures for a Parallel Tempering Monte Carlo simulation, that maximizes the probability that the configurations drift across all temperature values, from the coldest to the hottest ones, and vice versa. The proposed algorithm starts from data gathered from relatively short Monte Carlo simulations and is straightforward to implement. We assess its effectiveness on a test case simulation of an Edwards–Anderson spin glass on a lattice of 12 3 sites.

  17. Grain boundary diffusion in terms of the tempered fractional calculus

    Energy Technology Data Exchange (ETDEWEB)

    Sibatov, R.T., E-mail: ren_sib@bk.ru [Ulyanovsk State University, 432017, 42 Leo Tolstoy str., Ulyanovsk (Russian Federation); Svetukhin, V.V. [Ulyanovsk State University, 432017, 42 Leo Tolstoy str., Ulyanovsk (Russian Federation); Institute of Nanotechnology and Microelectronics of the Russian Academy of Sciences, 115487, 18 Nagatinskaya str., Moscow (Russian Federation)

    2017-06-28

    Mathematical treatment of grain-boundary diffusion based on the model first proposed by Fisher is usually formulated in terms of normal diffusion equations in a two-component nonhomogeneous medium. On the other hand, fractional equations of anomalous diffusion proved themselves to be useful in description of grain-boundary diffusion phenomena. Moreover, the most important propagation regime predicted by Fisher's model demonstrates subdiffusive behavior. However, the direct link between fractional approach and the Fisher model and its modifications has not found yet. Here, we fill this gap and show that solution of fractional subdiffusion equation offers general properties of classical solutions obtained by Whipple and Suzuoka. The tempered fractional approach is a convenient tool for studying precipitation in granular materials as the tempered subdiffusion limited process. - Highlights: • The link connected fractional diffusion approach and Fisher's model of grain-boundary diffusion is derived. • The subdiffusion exponent of grain-boundary diffusion can differ from 1/2. • Nucleation in granular materials is modeled by the process limited by tempered subdiffusion.

  18. Biochar boosts tropical but not temperate crop yields

    Science.gov (United States)

    Jeffery, Simon; Abalos, Diego; Prodana, Marija; Catarina Bastos, Ana; van Groenigen, Jan Willem; Hungate, Bruce A.; Verheijen, Frank

    2017-05-01

    Applying biochar to soil is thought to have multiple benefits, from helping mitigate climate change [1, 2], to managing waste [3] to conserving soil [4]. Biochar is also widely assumed to boost crop yield [5, 6], but there is controversy regarding the extent and cause of any yield benefit [7]. Here we use a global-scale meta-analysis to show that biochar has, on average, no effect on crop yield in temperate latitudes, yet elicits a 25% average increase in yield in the tropics. In the tropics, biochar increased yield through liming and fertilization, consistent with the low soil pH, low fertility, and low fertilizer inputs typical of arable tropical soils. We also found that, in tropical soils, high-nutrient biochar inputs stimulated yield substantially more than low-nutrient biochar, further supporting the role of nutrient fertilization in the observed yield stimulation. In contrast, arable soils in temperate regions are moderate in pH, higher in fertility, and generally receive higher fertilizer inputs, leaving little room for additional benefits from biochar. Our findings demonstrate that the yield-stimulating effects of biochar are not universal, but may especially benefit agriculture in low-nutrient, acidic soils in the tropics. Biochar management in temperate zones should focus on potential non-yield benefits such as lime and fertilizer cost savings, greenhouse gas emissions control, and other ecosystem services.

  19. Grain boundary diffusion in terms of the tempered fractional calculus

    International Nuclear Information System (INIS)

    Sibatov, R.T.; Svetukhin, V.V.

    2017-01-01

    Mathematical treatment of grain-boundary diffusion based on the model first proposed by Fisher is usually formulated in terms of normal diffusion equations in a two-component nonhomogeneous medium. On the other hand, fractional equations of anomalous diffusion proved themselves to be useful in description of grain-boundary diffusion phenomena. Moreover, the most important propagation regime predicted by Fisher's model demonstrates subdiffusive behavior. However, the direct link between fractional approach and the Fisher model and its modifications has not found yet. Here, we fill this gap and show that solution of fractional subdiffusion equation offers general properties of classical solutions obtained by Whipple and Suzuoka. The tempered fractional approach is a convenient tool for studying precipitation in granular materials as the tempered subdiffusion limited process. - Highlights: • The link connected fractional diffusion approach and Fisher's model of grain-boundary diffusion is derived. • The subdiffusion exponent of grain-boundary diffusion can differ from 1/2. • Nucleation in granular materials is modeled by the process limited by tempered subdiffusion.

  20. Comparative visual ecophysiology of mid-Atlantic temperate reef fishes

    Directory of Open Access Journals (Sweden)

    Andrij Z. Horodysky

    2013-11-01

    The absolute light sensitivities, temporal properties, and spectral sensitivities of the visual systems of three mid-Atlantic temperate reef fishes (Atlantic spadefish [Ephippidae: Chaetodipterus faber], tautog [Labridae: Tautoga onitis], and black sea bass [Serranidae: Centropristis striata] were studied via electroretinography (ERG. Pelagic Atlantic spadefish exhibited higher temporal resolution but a narrower dynamic range than the two more demersal foragers. The higher luminous sensitivities of tautog and black sea bass were similar to other benthic and demersal coastal mid-Atlantic fishes. Flicker fusion frequency experiments revealed significant interspecific differences at maximum intensities that correlated with lifestyle and habitat. Spectral responses of the three species spanned 400–610 nm, with high likelihood of cone dichromacy providing the basis for color and contrast discrimination. Significant day-night differences in spectral responses were evident in spadefish and black sea bass but not tautog, a labrid with characteristic structure-associated nocturnal torpor. Atlantic spadefish responded to a wider range of wavelengths than did deeper-dwelling tautog or black sea bass. Collectively, these results suggest that temperate reef-associated fishes are well-adapted to their gradient of brighter to dimmer photoclimates, representative of their unique ecologies and life histories. Continuing anthropogenic degradation of water quality in coastal environments, at a pace faster than the evolution of visual systems, may however impede visual foraging and reproductive signaling in temperate reef fishes.

  1. Carbon sequestration in managed temperate coniferous forests under climate change

    Science.gov (United States)

    Dymond, Caren C.; Beukema, Sarah; Nitschke, Craig R.; Coates, K. David; Scheller, Robert M.

    2016-03-01

    Management of temperate forests has the potential to increase carbon sinks and mitigate climate change. However, those opportunities may be confounded by negative climate change impacts. We therefore need a better understanding of climate change alterations to temperate forest carbon dynamics before developing mitigation strategies. The purpose of this project was to investigate the interactions of species composition, fire, management, and climate change in the Copper-Pine Creek valley, a temperate coniferous forest with a wide range of growing conditions. To do so, we used the LANDIS-II modelling framework including the new Forest Carbon Succession extension to simulate forest ecosystems under four different productivity scenarios, with and without climate change effects, until 2050. Significantly, the new extension allowed us to calculate the net sector productivity, a carbon accounting metric that integrates aboveground and belowground carbon dynamics, disturbances, and the eventual fate of forest products. The model output was validated against literature values. The results implied that the species optimum growing conditions relative to current and future conditions strongly influenced future carbon dynamics. Warmer growing conditions led to increased carbon sinks and storage in the colder and wetter ecoregions but not necessarily in the others. Climate change impacts varied among species and site conditions, and this indicates that both of these components need to be taken into account when considering climate change mitigation activities and adaptive management. The introduction of a new carbon indicator, net sector productivity, promises to be useful in assessing management effectiveness and mitigation activities.

  2. Phytoremediation of metal-contaminated soil in temperate humid regions of British Columbia, Canada.

    Science.gov (United States)

    Padmavathiamma, Prabha K; Li, Loretta Y

    2009-08-01

    The suitability of five plant species was studied for phytoextraction and phytostabilisation in a region with temperate maritime climate of coastal British Columbia, Canada. Pot experiments were conducted using Lolium perenne L (perennial rye grass), Festuca rubra L (creeping red fescue), Helianthus annuus L (sunflower), Poa pratensis L (Kentucky bluegrass) and Brassica napus L (rape) in soils treated with three different metal (Cu, Pb, Mn, and Zn) concentrations. The bio-metric characters of plants in soils with multiple-metal contaminations, their metal accumulation characteristics, translocation properties and metal removal were assessed at different stages of plant growth, 90 and 120 DAS (days after sowing). Lolium was found to be suitable for the phytostabilisation of Cu and Pb, Festuca for Mn and Poa for Zn. Metal removal was higher at 120 than at 90 days after sowing, and metals concentrated more in the underground tissues with less translocation to the aboveground parts. Bioconcentration factors indicate that Festuca had the highest accumulation for Cu, Helianthus for Pb and Zn and Poa for Mn.

  3. Melvin Calvin: Fuels from Plants

    Energy Technology Data Exchange (ETDEWEB)

    Taylor, S.E.; Otvos, J.W.

    1998-11-24

    A logical extension of his early work on the path of carbon during photosynthesis, Calvin's studies on the production of hydrocarbons by plants introduced many in the scientific and agricultural worlds to the potential of renewable fuel and chemical feedstocks. He and his co-workers identified numerous candidate compounds from plants found in tropical and temperate climates from around the world. His travels and lectures concerning the development of alternative fuel supplies inspired laboratories worldwide to take up the investigation of plant-derived energy sources as an alternative to fossil fuels.

  4. The possibility of tribopair lifetime extending by welding of quenched and tempered stainless steel with quenched and tempered carbon steel

    Directory of Open Access Journals (Sweden)

    V. Marušić

    2015-04-01

    Full Text Available In the conditions of tribocorrosion wear, extending of parts lifetime could be achieved by using stainless steel,which is hardened to sufficiently high hardness. In the tribosystem bolt/ bushing shell/link plate of the bucket elevator transporter conveyor machine, the previously quenched and tempered martensitic stainless steel for bolts is hardened at ≈47 HRC and welded with the quenched and tempered high yield carbon steel for bolts. Additional material, based on Cr-Ni-Mo (18/8/6 is used. The microstructure and hardness of welded samples are tested. On the tensile tester, resistance of the welded joint is tested with a simulated experiment. Dimensional control of worn tribosystem elements was performed after six months of service.

  5. Spent nuclear fuel project cold vacuum drying facility tempered water and tempered water cooling system design description

    International Nuclear Information System (INIS)

    IRWIN, J.J.

    1998-01-01

    This document provides the System Design Description (SDD) for the Cold Vacuum Drying Facility (CVDF) Tempered Water (TW) and Tempered Water Cooling (TWC) System . The SDD was developed in conjunction with HNF-SD-SNF-SAR-002, Safety Analysis Report for the Cold Vacuum Drying Facility, Phase 2, Supporting Installation of Processing Systems (Garvin 1998), The HNF-SD-SNF-DRD-O02, 1998, Cold Vacuum Drying Facility Design Requirements, and the CVDF Design Summary Report. The SDD contains general descriptions of the TW and TWC equipment, the system functions, requirements and interfaces. The SDD provides references for design and fabrication details, operation sequences and maintenance. This SOD has been developed for the SNFP Operations Organization and shall be updated, expanded, and revised in accordance with future design, construction and startup phases of the CVDF until the CVDF final ORR is approved

  6. Prediction of hardness in pieces of quenched and tempered steel

    International Nuclear Information System (INIS)

    Yanzon, Rodolfo Carlos; Rodriguez, Augusto; Sanchez, Arlington Ricardo

    2006-01-01

    This presentation describes the first stage of a work plan to obtain a simple software, for predicting properties at certain points of a quenched and tempered piece. In this first stage, this prediction is limited to cylindrical pieces made with steels whose chemical composition is within a certain range. The methodology basically consists of obtaining , from experimental data, a mathematical tool able to predict the hardness value, for Jominy test piece ends made with this type of steel. This meant beginning with the analysis of the usual forms of theoretical calculation of Jominy curves of quenched samples, which resulted in a proposal to modify the Just equation. Two different mathematical methods were then developed, that could predict hardness values in tempered Jominy test pieces. One, based on the determination of polynomic equations, that reproduces the loss of hardness at points along the test piece, base on the quenching value and as a function of the tempering temperature. The other one, which uses the lineal multidimensional interpolation method, because of its ease of application, has been selected as the mathematical tool to use in the software under development. At this stage of the work, the relationship between the points on the piece and those on the Jominy test pieces is carried out by the Lamont method and the representative variable of the temperature/time combination for the tempering process itself, is obtained with software based on the Hollomon and Jaffe expression. Data is needed to define: a) chemical composition and grain size of the steel used, b) diameter of the piece, c) 'H G ' severity of the quenching medium d) temperature and time of the tempering. The work's second stage continued with the addition of hardness values measured in Jominy test pieces made with other steels. The chemical composition and grain size data of each steel introduced are converted by the software into one more variable, using the concept of ideal critical

  7. Defense Signaling in Plants -R-ES-ONAN

    Indian Academy of Sciences (India)

    which feed on the leaves of these plants develop the problem of improper digestion .... bacteria also leads to damage and disintegration of the host plant cells in the ... tion) and oligosaccharides (products of cellulose degradation). It appears that .... compared to the temperate countries where most research on this topic has ...

  8. A model of plant strategies in fluvial hydrosystems

    NARCIS (Netherlands)

    Bornette, G.; Tabacchi, E.; Hupp, C.; Puijalon, S.; Rostan, J.C.

    2008-01-01

    1. We propose a model of plant strategies in temperate fluvial hydrosystems that considers the hydraulic and geomorphic features that control plant recruitment, establishment and growth in river floodplains. 2. The model describes first how the disturbance gradient and the grain-size of the river

  9. Linking xylem water storage with anatomical parameters in five temperate tree species.

    Science.gov (United States)

    Jupa, Radek; Plavcová, Lenka; Gloser, Vít; Jansen, Steven

    2016-06-01

    The release of water from storage compartments to the transpiration stream is an important functional mechanism that provides the buffering of sudden fluctuations in water potential. The ability of tissues to release water per change in water potential, referred to as hydraulic capacitance, is assumed to be associated with the anatomy of storage tissues. However, information about how specific anatomical parameters determine capacitance is limited. In this study, we measured sapwood capacitance (C) in terminal branches and roots of five temperate tree species (Fagus sylvatica L., Picea abies L., Quercus robur L., Robinia pseudoacacia L., Tilia cordata Mill.). Capacitance was calculated separately for water released mainly from capillary (CI; open vessels, tracheids, fibres, intercellular spaces and cracks) and elastic storage compartments (CII; living parenchyma cells), corresponding to two distinct phases of the moisture release curve. We found that C was generally higher in roots than branches, with CI being 3-11 times higher than CII Sapwood density and the ratio of dead to living xylem cells were most closely correlated with C In addition, the magnitude of CI was strongly correlated with fibre/tracheid lumen area, whereas CII was highly dependent on the thickness of axial parenchyma cell walls. Our results indicate that water released from capillary compartments predominates over water released from elastic storage in both branches and roots, suggesting the limited importance of parenchyma cells for water storage in juvenile xylem of temperate tree species. Contrary to intact organs, water released from open conduits in our small wood samples significantly increased CI at relatively high water potentials. Linking anatomical parameters with the hydraulic capacitance of a tissue contributes to a better understanding of water release mechanisms and their implications for plant hydraulics. © The Author 2016. Published by Oxford University Press. All rights

  10. Reduced germination success of temperate grassland seeds sown in dung: consequences for post-dispersal seed fate.

    Science.gov (United States)

    Milotić, T; Hoffmann, M

    2016-11-01

    Endozoochory is one of the main drivers shaping temperate grassland communities by maintaining plant populations of its constituents and enabling plants to colonize new habitats. Successful endozoochorous dispersal implies that seeds not only get consumed and survive the digestive tract but are also able to develop into viable seedlings in a dung environment. We experimentally assessed the germination probability and timing of 15 annual and perennial temperate European grassland species in cattle and horse dung and in different climatic conditions (greenhouse and outdoor conditions). Interspecific variation in germinability and germination timing are found, while life strategy had only an effect on germination timing. We found adverse effects of both cattle and horse dung on the germination characteristics of all tested grassland species, but the effects of cattle dung were more pronounced. In comparison with the control treatment, fewer seeds emerged in dung and more time was needed to germinate. Also, germination metrics clearly differed between the artificial greenhouse and outdoor conditions, with generally a lower germinability in outdoor conditions. According to our results, a large cost seems to be associated with endozoochorous dispersal in this stage of the life cycle, as seed dispersal effectiveness strongly depends on the quality of the deposition site with a lowered survival and germination probability when seeds are deposited in dung. © 2016 German Botanical Society and The Royal Botanical Society of the Netherlands.

  11. The tropicalization of temperate marine ecosystems: climate-mediated changes in herbivory and community phase shifts

    Science.gov (United States)

    Vergés, Adriana; Steinberg, Peter D.; Hay, Mark E.; Poore, Alistair G. B.; Campbell, Alexandra H.; Ballesteros, Enric; Heck, Kenneth L.; Booth, David J.; Coleman, Melinda A.; Feary, David A.; Figueira, Will; Langlois, Tim; Marzinelli, Ezequiel M.; Mizerek, Toni; Mumby, Peter J.; Nakamura, Yohei; Roughan, Moninya; van Sebille, Erik; Gupta, Alex Sen; Smale, Dan A.; Tomas, Fiona; Wernberg, Thomas; Wilson, Shaun K.

    2014-01-01

    Climate-driven changes in biotic interactions can profoundly alter ecological communities, particularly when they impact foundation species. In marine systems, changes in herbivory and the consequent loss of dominant habitat forming species can result in dramatic community phase shifts, such as from coral to macroalgal dominance when tropical fish herbivory decreases, and from algal forests to ‘barrens’ when temperate urchin grazing increases. Here, we propose a novel phase-shift away from macroalgal dominance caused by tropical herbivores extending their range into temperate regions. We argue that this phase shift is facilitated by poleward-flowing boundary currents that are creating ocean warming hotspots around the globe, enabling the range expansion of tropical species and increasing their grazing rates in temperate areas. Overgrazing of temperate macroalgae by tropical herbivorous fishes has already occurred in Japan and the Mediterranean. Emerging evidence suggests similar phenomena are occurring in other temperate regions, with increasing occurrence of tropical fishes on temperate reefs. PMID:25009065

  12. An investigation on high temperature fatigue properties of tempered nuclear-grade deposited weld metals

    Science.gov (United States)

    Cao, X. Y.; Zhu, P.; Yong, Q.; Liu, T. G.; Lu, Y. H.; Zhao, J. C.; Jiang, Y.; Shoji, T.

    2018-02-01

    Effect of tempering on low cycle fatigue (LCF) behaviors of nuclear-grade deposited weld metal was investigated, and The LCF tests were performed at 350 °C with strain amplitudes ranging from 0.2% to 0.6%. The results showed that at a low strain amplitude, deposited weld metal tempered for 1 h had a high fatigue resistance due to high yield strength, while at a high strain amplitude, the one tempered for 24 h had a superior fatigue resistance due to high ductility. Deposited weld metal tempered for 1 h exhibited cyclic hardening at the tested strain amplitudes. Deposited weld metal tempered for 24 h exhibited cyclic hardening at a low strain amplitude but cyclic softening at a high strain amplitude. Existence and decomposition of martensite-austenite (M-A) islands as well as dislocations activities contributed to fatigue property discrepancy among the two tempered deposited weld metal.

  13. Effect of Tempering and Baking on the Charpy Impact Energy of Hydrogen-Charged 4340 Steel

    Science.gov (United States)

    Mori, K.; Lee, E. W.; Frazier, W. E.; Niji, K.; Battel, G.; Tran, A.; Iriarte, E.; Perez, O.; Ruiz, H.; Choi, T.; Stoyanov, P.; Ogren, J.; Alrashaid, J.; Es-Said, O. S.

    2015-01-01

    Tempered AISI 4340 steel was hydrogen charged and tested for impact energy. It was found that samples tempered above 468 °C (875 °F) and subjected to hydrogen charging exhibited lower impact energy values when compared to uncharged samples. No significant difference between charged and uncharged samples tempered below 468 °C (875 °F) was observed. Neither exposure nor bake time had any significant effect on impact energy within the tested ranges.

  14. On the evolution of Cu-Ni-rich bridges of Alnico alloys with tempering

    Energy Technology Data Exchange (ETDEWEB)

    Fan, M. [Department of Materials Science and Engineering, North Carolina State University, Campus Box 7907, Raleigh, NC 27695-7907 (United States); Liu, Y. [Department of Materials Science and Engineering, North Carolina State University, Campus Box 7907, Raleigh, NC 27695-7907 (United States); Analytical Instrumentation Facility, North Carolina State University, Raleigh, NC 27695 (United States); Jha, Rajesh; Dulikravich, George S. [Departments of Mechanical and Materials Engineering, MAIDROC, Florida International University, EC3462, 10555 West Flagler Street, Miami, FL 33174 (United States); Schwartz, J.; Koch, C.C. [Department of Materials Science and Engineering, North Carolina State University, Campus Box 7907, Raleigh, NC 27695-7907 (United States)

    2016-12-15

    Tempering is a critical step in Alnico alloy processing, yet the effects of tempering on microstructure have not been well studied. Here we report these effects, and in particular the effects on the Cu-Ni bridges. Energy-dispersive X-ray spectroscopy (EDS) maps and line scans show that tempering changes the elemental distribution in the Cu-Ni bridges, but not the morphology and distribution of Cu-bridges. The Cu concentration in the Cu-Ni bridges increases after tempering while other element concentrations decrease, especially Ni and Al. Furthermore, tempering sharpens the Cu bridge boundaries. These effects are primarily related to the large 2C{sub 44}/(C{sub 11}−C{sub 12}) ratio for Cu, largest of all elements in Alnico. In addition, the Ni-Cu loops around the α{sub 1} phases become inconspicuous with tempering. The diffusion of Fe and Co to the α{sub 1} phase during tempering, which increases the difference of saturation magnetization between the α{sub 1} and α{sub 2} phases, is observed by EDS. In summary, α{sub 1}, α{sub 2} and Cu-bridges are concentrated with their major elements during tempering which improves the magnetic properties. The formation of these features formed through elemental diffusion is discussed via energy theories. - Highlights: • Tempering changes the elemental distribution in the Cu-Ni bridges, but not morphology. • Cu concentration in the Cu-Ni bridges increases after tempering while others decrease. • These effects are related to the large 2C{sub 44}/(C{sub 11}−C{sub 12}) ratio for Cu. • The Ni-Cu loops around the α{sub 1} phases become inconspicuous with tempering. • The diffusion of Fe and Co to the α{sub 1} phase during tempering is observed by EDS.

  15. Method to Predict Tempering of Steels Under Non-isothermal Conditions

    Science.gov (United States)

    Poirier, D. R.; Kohli, A.

    2017-05-01

    A common way of representing the tempering responses of steels is with a "tempering parameter" that includes the effect of temperature and time on hardness after hardening. Such functions, usually in graphical form, are available for many steels and have been applied for isothermal tempering. In this article, we demonstrate that the method can be extended to non-isothermal conditions. Controlled heating experiments were done on three grades in order to verify the method.

  16. Complementary models of tree species-soil relationships in old-growth temperate forests

    Science.gov (United States)

    Cross, Alison; Perakis, Steven S.

    2011-01-01

    Ecosystem level studies identify plant soil feed backs as important controls on soil nutrient availability,particularly for nitrogen and phosphorus. Although site and species specific studies of tree species soil relationships are relatively common,comparatively fewer studies consider multiple coexisting speciesin old-growth forests across a range of sites that vary underlying soil fertility. We characterized patterns in forest floor and mineral soil nutrients associated with four common tree species across eight undisturbed old-growth forests in Oregon, USA, and used two complementary conceptual models to assess tree species soil relationships. Plant soil feedbacks that could reinforce sitelevel differences in nutrient availability were assessed using the context dependent relationships model, where by relative species based differences in each soil nutrient divergedorconvergedas nutrient status changed across sites. Tree species soil relationships that did not reflect strong feedbacks were evaluated using a site independent relationships model, where by forest floor and surface mineral soil nutrient tools differed consistently by tree species across sites,without variation in deeper mineral soils. We found that theorganically cycled elements carbon, nitrogen, and phosphorus exhibited context-dependent differences among species in both forest floor and mineral soil, and most of ten followed adivergence model,where by species differences were greatest at high-nutrient sites. These patterns are consistent with the oryemphasizing biotic control of these elements through plant soil feedback mechanisms. Site independent species differences were strongest for pool so if the weather able cations calcium, magnesium, potassium,as well as phosphorus, in mineral soils. Site independent species differences in forest floor nutrients we reattributable too nespecies that displayed significant greater forest floor mass accumulation. Our finding confirmed that site-independent and

  17. Importance of biotic and abiotic components in feedback between plants and soil

    OpenAIRE

    Hanzelková, Věra

    2017-01-01

    The plant-soil feedback affects the forming of a plant community. Plants affect their own species as well as other species. The plant-soil feedback can be both positive and negative. Plants affect soil, change its properties, and the soil affects the plants reciprocally. Soil components can be divided into biotic and abiotic ones. The abiotic component is represented by physical and chemical properties of the soil. The main properties are the soil structure, the soil moisture, the soil temper...

  18. Microstructural characteristics of steel rebar submitted to the process of self - tempering; Caracterizacao microestrutural de superficies e interfaces de vergalhoes de aco auto-revenidos utilizando Microscopia Eletronica de Varredura (MEV) e analise quimica

    Energy Technology Data Exchange (ETDEWEB)

    Barreto, M.F.O., E-mail: matheusfob@yahoo.com.br [Universidade Federal de Minas Gerais (UFMG), MG (Brazil)

    2014-07-01

    Every day the construction industry seeks new alternatives to maximize use of the materials provided new tools for structural design of the new techniques in welding rods. In this sense, this work proposes to study the mechanical properties of steel rebar self-tempering for civil construction, choosing the bars from 6.3 to 25.4 mm in diameter for the study. The cooling process of rebar, after rolling, known as self-tempering produces a rebar with excellent mechanical properties and low cost to the plant, they put the rebar shall be composed of concentric layers with different mechanical properties among themselves. To study the influence of these layers on the mechanical behavior of rebar microstructural characterization was made by means of electron microscopy analysis of transmitted light and scanning of all samples. From the results it was found that only the bars 20 and 25mm diameter had a heat treatment for self-tempering relevant. (author)

  19. Time-dependent temper embrittlement of reactor pressure vessel steel: Correlation between microstructural evolution and mechanical properties during tempering at 650 °C

    Energy Technology Data Exchange (ETDEWEB)

    Li, Chuanwei; Han, Lizhan; Yan, Guanghua; Liu, Qingdong; Luo, Xiaomeng; Gu, Jianfeng, E-mail: gujf@sjtu.edu.cn

    2016-11-15

    The microstructural evolution of reactor pressure vessel (RPV) steel and its effect on the mechanical properties during tempering at 650 °C were studied to reveal the time-dependent toughness and temper embrittlement. The results show that the toughening of the material should be attributed to the decomposition of the martensite/austenite constituents and uniform distribution of carbides. When the tempering duration was 5 h, the strength of the investigated steel decreased to strike a balance with the material impact toughness that reached a plateau. As the tempering duration was further increased, the material strength was slightly reduced but the material impact toughness deteriorated drastically. This time-dependent temper embrittlement is different from traditional temper embrittlement, and it can be partly attributed to the softening of the matrix and the broadening of the ferrite laths. Moreover, the dimensions and distribution of the grain carbides are the most important factors of the impact toughness. - Highlights: • The fracture mechanism of reactor pressure vessel (RPV) steels under impact load was investigated. • The Charpy V-notch impact test and the hinge model were employed for the study. • Grain boundary carbides play a key role in the impact toughness and fracture toughness. • The dependence of the deterioration of impact toughness on tempering time was analyzed for the first time.

  20. Temporal-spatial segregation among hummingbirds foraging on honeydew in a temperate forest in Mexico

    Directory of Open Access Journals (Sweden)

    Carlos LARA, Vanessa MARTÍNEZ-GARCÍA, Raúl ORTIZ-PULIDO, Jessica BRAVO-CADENA, Salvador LORANCA, Alex CÓRDOBA-AGUILAR

    2011-02-01

    Full Text Available Spatial and temporal variation in interactions between hummingbirds and plants have often been examined, and hummingbirds and insects are known to indirectly interact in networks of nectar plants. In a highland temperate forest in Hidalgo, Mexico some oak trees were heavily infested by honeydew-producing insects (family Margarodidae, tribe Xylococcini, genus Strigmacoccus and the honeydew was consumed by hummingbirds. Here using survival analysis we investigate how the honeydew produced by dense populations of these margarodids is temporally and spatially partitioned by hummingbirds. We also measured the availability and quality of honeydew exudates, and then we recorded the time until a bird visited and used such resources. Four hummingbird species consumed this resource (Atthis eloisa, Hylocharis leucotis, Colibri thalassinus and Eugenes fulgens. Data from 294 hours of observation on seven focal trees suggested temporal and spatial segregation among visiting birds according to body size and territorial behavior during the most honeydew-limited time. Hummingbird species differed in the daily times they foraged, as well as in the location where honeydew-producing insects were visited on the trees. Temporal and spatial segregation among hummingbird species is interpreted as an adaptation to reduce the risk of aggressive encounters. This may facilitate multispecies coexistence and allow these birds to exploit honeydew more effectively [Current Zoology 57 (1: 56–62, 2011].

  1. Linking root traits to nutrient foraging in arbuscular mycorrhizal trees in a temperate forest.

    Science.gov (United States)

    Eissenstat, David M; Kucharski, Joshua M; Zadworny, Marcin; Adams, Thomas S; Koide, Roger T

    2015-10-01

    The identification of plant functional traits that can be linked to ecosystem processes is of wide interest, especially for predicting vegetational responses to climate change. Root diameter of the finest absorptive roots may be one plant trait that has wide significance. Do species with relatively thick absorptive roots forage in nutrient-rich patches differently from species with relatively fine absorptive roots? We measured traits related to nutrient foraging (root morphology and architecture, root proliferation, and mycorrhizal colonization) across six coexisting arbuscular mycorrhizal (AM) temperate tree species with and without nutrient addition. Root traits such as root diameter and specific root length were highly correlated with root branching intensity, with thin-root species having higher branching intensity than thick-root species. In both fertilized and unfertilized soil, species with thin absorptive roots and high branching intensity showed much greater root length and mass proliferation but lower mycorrhizal colonization than species with thick absorptive roots. Across all species, fertilization led to increased root proliferation and reduced mycorrhizal colonization. These results suggest that thin-root species forage more by root proliferation, whereas thick-root species forage more by mycorrhizal fungi. In mineral nutrient-rich patches, AM trees seem to forage more by proliferating roots than by mycorrhizal fungi. © 2015 The Authors. New Phytologist © 2015 New Phytologist Trust.

  2. Joints in Tempered Glass Using Glass Dowel Discs

    DEFF Research Database (Denmark)

    Nielsen, Jens Henrik; Poulsen, Peter Noe

    One of the major reasons for using glass in structures is its transparency; however, traditional mechanical joints such as friction joints and steel dowel pinned connections are compromising the transparency. The present paper describes a novel joint which is practically maintaining the complete...... transparency of the glass. This is achieved by using a dowel disc made entirely of tempered glass. The concept of the joint is proved by pilot tests and numerical models. From the work it is seen that the load-carrying capacity of such a connection is similar to what is found for traditionally in-plane loaded...

  3. Impact of soil nematodes on salt-marsh plants : a pilot experiment

    NARCIS (Netherlands)

    Dormann, CF; van der Wal, R

    2001-01-01

    We tested whether the removal of nematodes by means of nematicide application changed plant performance or influenced plant competition. The study involved the two common plant species Artemisia maritima and Festuca rubra growing in intact sods collected from a temperate salt marsh. Half of the sods

  4. Effects of competition and herbivory over woody seedling growth in a temperate woodland trump the effects of elevated CO2.

    Science.gov (United States)

    Collins, L; Boer, M M; de Dios, V Resco; Power, S A; Bendall, E R; Hasegawa, S; Hueso, R Ochoa; Nevado, J Piñeiro; Bradstock, R A

    2018-04-27

    A trend of increasing woody plant density, or woody thickening, has been observed across grassland and woodland ecosystems globally. It has been proposed that increasing atmospheric [CO 2 ] is a major driver of broad scale woody thickening, though few field-based experiments have tested this hypothesis. Our study utilises a Free Air CO 2 Enrichment experiment to examine the effect of elevated [CO 2 ] (eCO 2 ) on three mechanisms that can cause woody thickening, namely (i) woody plant recruitment, (ii) seedling growth, and (iii) post-disturbance resprouting. The study took place in a eucalypt-dominated temperate grassy woodland. Annual assessments show that juvenile woody plant recruitment occurred over the first 3 years of CO 2 fumigation, though eCO 2 did not affect rates of recruitment. Manipulative experiments were established to examine the effect of eCO 2 on above-ground seedling growth using transplanted Eucalyptus tereticornis (Myrtaceae) and Hakea sericea (Proteaceae) seedlings. There was no positive effect of eCO 2 on biomass of either species following 12 months of exposure to treatments. Lignotubers (i.e., resprouting organs) of harvested E. tereticornis seedlings that were retained in situ for an additional year were used to examine resprouting response. The likelihood of resprouting and biomass of resprouts increased with lignotuber volume, which was not itself affected by eCO 2 . The presence of herbaceous competitors and defoliation by invertebrates and pathogens were found to greatly reduce growth and/or resprouting response of seedlings. Our findings do not support the hypothesis that future increases in atmospheric [CO 2 ] will, by itself, promote woody plant recruitment in eucalypt-dominated temperate grassy woodlands.

  5. Monoterpene emissions in response to long-term night-time warming, elevated CO2 and extended summer drought in a temperate heath ecosystem

    DEFF Research Database (Denmark)

    Tiiva, Päivi; Tang, Jing; Michelsen, Anders

    2017-01-01

    Monoterpenes emitted from plants have an important role in atmospheric chemistry through changing atmospheric oxidative capacity, forming new particles and secondary organic aerosols. The emission rates and patterns can be affected by changing climate. In this study, emission responses to six years...... of climatic manipulations (elevated CO2, extended summer drought and night-time warming) were investigated in a temperate semi-natural heath ecosystem. Samples for monoterpene analysis were collected in seven campaigns during an entire growing season (April-November, 2011). The results showed...... that the temperate heath ecosystem was a considerable source of monoterpenes to the atmosphere, with the emission averaged over the 8month measurement period of 21.7±6.8μgm(-2)groundareah(-1) for the untreated heath. Altogether, 16 monoterpenes were detected, of which the most abundant were α-pinene, δ-3-carene...

  6. Negative correlation between rates of molecular evolution and flowering cycles in temperate woody bamboos revealed by plastid phylogenomics.

    Science.gov (United States)

    Ma, Peng-Fei; Vorontsova, Maria S; Nanjarisoa, Olinirina Prisca; Razanatsoa, Jacqueline; Guo, Zhen-Hua; Haevermans, Thomas; Li, De-Zhu

    2017-12-21

    Heterogeneous rates of molecular evolution are universal across the tree of life, posing challenges for phylogenetic inference. The temperate woody bamboos (tribe Arundinarieae, Poaceae) are noted for their extremely slow molecular evolutionary rates, supposedly caused by their mysterious monocarpic reproduction. However, the correlation between substitution rates and flowering cycles has not been formally tested. Here we present 15 newly sequenced plastid genomes of temperate woody bamboos, including the first genomes ever sequenced from Madagascar representatives. A data matrix of 46 plastid genomes representing all 12 lineages of Arundinarieae was assembled for phylogenetic and molecular evolutionary analyses. We conducted phylogenetic analyses using different sequences (e.g., coding and noncoding) combined with different data partitioning schemes, revealing conflicting relationships involving internodes among several lineages. A great difference in branch lengths were observed among the major lineages, and topological inconsistency could be attributed to long-branch attraction (LBA). Using clock model-fitting by maximum likelihood and Bayesian approaches, we furthermore demonstrated extensive rate variation among these major lineages. Rate accelerations mainly occurred for the isolated lineages with limited species diversification, totaling 11 rate shifts during the tribe's evolution. Using linear regression analysis, we found a negative correlation between rates of molecular evolution and flowering cycles for Arundinarieae, notwithstanding that the correlation maybe insignificant when taking the phylogenetic structure into account. Using the temperate woody bamboos as an example, we found further evidence that rate heterogeneity is universal in plants, suggesting that this will pose a challenge for phylogenetic reconstruction of bamboos. The bamboos with longer flowering cycles tend to evolve more slowly than those with shorter flowering cycles, in accordance

  7. Increased drought impacts on temperate rainforests from southern South America: results of a process-based, dynamic forest model.

    Directory of Open Access Journals (Sweden)

    Alvaro G Gutiérrez

    Full Text Available Increased droughts due to regional shifts in temperature and rainfall regimes are likely to affect forests in temperate regions in the coming decades. To assess their consequences for forest dynamics, we need predictive tools that couple hydrologic processes, soil moisture dynamics and plant productivity. Here, we developed and tested a dynamic forest model that predicts the hydrologic balance of North Patagonian rainforests on Chiloé Island, in temperate South America (42°S. The model incorporates the dynamic linkages between changing rainfall regimes, soil moisture and individual tree growth. Declining rainfall, as predicted for the study area, should mean up to 50% less summer rain by year 2100. We analysed forest responses to increased drought using the model proposed focusing on changes in evapotranspiration, soil moisture and forest structure (above-ground biomass and basal area. We compared the responses of a young stand (YS, ca. 60 years-old and an old-growth forest (OG, >500 years-old in the same area. Based on detailed field measurements of water fluxes, the model provides a reliable account of the hydrologic balance of these evergreen, broad-leaved rainforests. We found higher evapotranspiration in OG than YS under current climate. Increasing drought predicted for this century can reduce evapotranspiration by 15% in the OG compared to current values. Drier climate will alter forest structure, leading to decreases in above ground biomass by 27% of the current value in OG. The model presented here can be used to assess the potential impacts of climate change on forest hydrology and other threats of global change on future forests such as fragmentation, introduction of exotic tree species, and changes in fire regimes. Our study expands the applicability of forest dynamics models in remote and hitherto overlooked regions of the world, such as southern temperate rainforests.

  8. Increased drought impacts on temperate rainforests from southern South America: results of a process-based, dynamic forest model.

    Science.gov (United States)

    Gutiérrez, Alvaro G; Armesto, Juan J; Díaz, M Francisca; Huth, Andreas

    2014-01-01

    Increased droughts due to regional shifts in temperature and rainfall regimes are likely to affect forests in temperate regions in the coming decades. To assess their consequences for forest dynamics, we need predictive tools that couple hydrologic processes, soil moisture dynamics and plant productivity. Here, we developed and tested a dynamic forest model that predicts the hydrologic balance of North Patagonian rainforests on Chiloé Island, in temperate South America (42°S). The model incorporates the dynamic linkages between changing rainfall regimes, soil moisture and individual tree growth. Declining rainfall, as predicted for the study area, should mean up to 50% less summer rain by year 2100. We analysed forest responses to increased drought using the model proposed focusing on changes in evapotranspiration, soil moisture and forest structure (above-ground biomass and basal area). We compared the responses of a young stand (YS, ca. 60 years-old) and an old-growth forest (OG, >500 years-old) in the same area. Based on detailed field measurements of water fluxes, the model provides a reliable account of the hydrologic balance of these evergreen, broad-leaved rainforests. We found higher evapotranspiration in OG than YS under current climate. Increasing drought predicted for this century can reduce evapotranspiration by 15% in the OG compared to current values. Drier climate will alter forest structure, leading to decreases in above ground biomass by 27% of the current value in OG. The model presented here can be used to assess the potential impacts of climate change on forest hydrology and other threats of global change on future forests such as fragmentation, introduction of exotic tree species, and changes in fire regimes. Our study expands the applicability of forest dynamics models in remote and hitherto overlooked regions of the world, such as southern temperate rainforests.

  9. Endemic shrubs in temperate arid and semiarid regions of northern China and their potentials for rangeland restoration.

    Science.gov (United States)

    Chu, Jianmin; Yang, Hongxiao; Lu, Qi; Zhang, Xiaoyan

    2015-06-03

    Some endemic shrubs in arid and semiarid ecosystems are in danger of extinction, and yet they can play useful roles in maintaining or restoring these ecosystems, thus practical efforts are needed to conserve them. The shrubs Amygdalus pedunculata Pall., Amygdalus mongolica (Maxim.) Ricker and Ammopiptanthus mongolicus (Maxim. ex Kom.) Cheng f. are endemic species in arid and semiarid regions of northern China, where rangeland desertification is pronounced due to chronic overgrazing. In this study, we tested the hypothesis that these endemic shrubs have developed adaptations to arid and semiarid environments and could play critical roles as nurse species to initiate the process of rangeland recovery. Based on careful vegetation surveys, we analysed the niches of these species in relation to precipitation, temperature and habitats. All sampling plots were categorized by these endemics and sorted by the non-metric multidimensional scaling method. Species ratios of each life form and species co-occurrence rates with the endemics were also evaluated. Annual average temperature and annual precipitation were found to be the key factors determining vegetation diversity and distributions. Amygdalus pedunculata prefers low hills and sandy land in temperate semiarid regions. Amygdalus mongolica prefers gravel deserts of temperate semiarid regions. Ammopiptanthus mongolicus prefers sandy land of temperate arid regions. Communities of A. pedunculata have the highest diversity and the largest ratios of long-lived grass species, whereas those of A. mongolicus have the lowest diversity but the largest ratios of shrub species. Communities of A. mongolica are a transition between the first two community types. These findings demonstrate that our focal endemic shrubs have evolved adaptations to arid and semiarid conditions, thus they can be nurse plants to stabilize sand ground for vegetation restoration. We suggest that land managers begin using these shrub species to restore

  10. Comparative life history of the south temperate Cape Penduline Tit (Anthoscopus minutus) and north temperate Remizidae species

    Science.gov (United States)

    Lloyd, Penn; Frauenknecht, Bernhard D.; du Plessis, Morné A.; Martin, Thomas E.

    2017-01-01

    We studied the breeding biology of the south temperate Cape Penduline Tit (Anthoscopus minutus) in order to compare its life history traits with those of related north temperate members of the family Remizidae, namely the Eurasian Penduline Tit (Remiz pendulinus) and the Verdin (Auriparus flaviceps). We used this comparison to test key predictions of three hypotheses thought to explain latitudinal variation in life histories among bird species—the seasonality and food limitation hypothesis, nest predation hypothesis and adult mortality hypothesis. Contrary to the general pattern of smaller clutch size and lower adult mortality among south-temperate birds living in less seasonal environments, the Cape Penduline Tit has a clutch size larger than that of the Verdin and similar to that of the Eurasian Penduline Tit, and higher adult mortality than both of the other two species. The most notable difference between the Cape Penduline Tit and the two other species is in parental behavioural strategy, with the former exhibiting bi-parental care at all stages of nesting together with facultative cooperative breeding, whereas the Eurasian Penduline Tit has uni-parental care and the Verdin has a combination of female-only incubation but bi-parental nestling care. Consequently, in comparison to the other two species, the Cape Penduline Tit exhibits greater nest attentiveness during incubation, a similar per-nestling feeding rate and greater post-fledging survival. Its relatively large clutch size, high parental investment and associated high adult mortality in a less seasonal environment are consistent with key predictions of the adult mortality hypothesis but not with key predictions of the seasonality and food limitation hypothesis in explaining life history variation among Remizidae species. These results add to a growing body of evidence of the importance of age-specific mortality in shaping life history evolution.

  11. Chilling and heat requirements for flowering in temperate fruit trees.

    Science.gov (United States)

    Guo, Liang; Dai, Junhu; Ranjitkar, Sailesh; Yu, Haiying; Xu, Jianchu; Luedeling, Eike

    2014-08-01

    Climate change has affected the rates of chilling and heat accumulation, which are vital for flowering and production, in temperate fruit trees, but few studies have been conducted in the cold-winter climates of East Asia. To evaluate tree responses to variation in chill and heat accumulation rates, partial least squares regression was used to correlate first flowering dates of chestnut (Castanea mollissima Blume) and jujube (Zizyphus jujube Mill.) in Beijing, China, with daily chill and heat accumulation between 1963 and 2008. The Dynamic Model and the Growing Degree Hour Model were used to convert daily records of minimum and maximum temperature into horticulturally meaningful metrics. Regression analyses identified the chilling and forcing periods for chestnut and jujube. The forcing periods started when half the chilling requirements were fulfilled. Over the past 50 years, heat accumulation during tree dormancy increased significantly, while chill accumulation remained relatively stable for both species. Heat accumulation was the main driver of bloom timing, with effects of variation in chill accumulation negligible in Beijing’s cold-winter climate. It does not seem likely that reductions in chill will have a major effect on the studied species in Beijing in the near future. Such problems are much more likely for trees grown in locations that are substantially warmer than their native habitats, such as temperate species in the subtropics and tropics.

  12. Chilling and heat requirements for flowering in temperate fruit trees

    Science.gov (United States)

    Guo, Liang; Dai, Junhu; Ranjitkar, Sailesh; Yu, Haiying; Xu, Jianchu; Luedeling, Eike

    2014-08-01

    Climate change has affected the rates of chilling and heat accumulation, which are vital for flowering and production, in temperate fruit trees, but few studies have been conducted in the cold-winter climates of East Asia. To evaluate tree responses to variation in chill and heat accumulation rates, partial least squares regression was used to correlate first flowering dates of chestnut ( Castanea mollissima Blume) and jujube ( Zizyphus jujube Mill.) in Beijing, China, with daily chill and heat accumulation between 1963 and 2008. The Dynamic Model and the Growing Degree Hour Model were used to convert daily records of minimum and maximum temperature into horticulturally meaningful metrics. Regression analyses identified the chilling and forcing periods for chestnut and jujube. The forcing periods started when half the chilling requirements were fulfilled. Over the past 50 years, heat accumulation during tree dormancy increased significantly, while chill accumulation remained relatively stable for both species. Heat accumulation was the main driver of bloom timing, with effects of variation in chill accumulation negligible in Beijing's cold-winter climate. It does not seem likely that reductions in chill will have a major effect on the studied species in Beijing in the near future. Such problems are much more likely for trees grown in locations that are substantially warmer than their native habitats, such as temperate species in the subtropics and tropics.

  13. Nitrous oxide emission reduction in temperate biochar-amended soils

    Science.gov (United States)

    Felber, R.; Hüppi, R.; Leifeld, J.; Neftel, A.

    2012-01-01

    Biochar, a pyrolysis product of organic residues, is an amendment for agricultural soils to improve soil fertility, sequester CO2 and reduce greenhouse gas (GHG) emissions. In highly weathered tropical soils laboratory incubations of soil-biochar mixtures revealed substantial reductions for nitrous oxide (N2O) and carbon dioxide (CO2). In contrast, evidence is scarce for temperate soils. In a three-factorial laboratory incubation experiment two different temperate agricultural soils were amended with green waste and coffee grounds biochar. N2O and CO2 emissions were measured at the beginning and end of a three month incubation. The experiments were conducted under three different conditions (no additional nutrients, glucose addition, and nitrate and glucose addition) representing different field conditions. We found mean N2O emission reductions of 60 % compared to soils without addition of biochar. The reduction depended on biochar type and soil type as well as on the age of the samples. CO2 emissions were slightly reduced, too. NO3- but not NH4+ concentrations were significantly reduced shortly after biochar incorporation. Despite the highly significant suppression of N2O emissions biochar effects should not be transferred one-to-one to field conditions but need to be tested accordingly.

  14. Temperate marine protected area provides recruitment subsidies to local fisheries.

    Science.gov (United States)

    Le Port, A; Montgomery, J C; Smith, A N H; Croucher, A E; McLeod, I M; Lavery, S D

    2017-10-25

    The utility of marine protected areas (MPAs) as a means of protecting exploited species and conserving biodiversity within MPA boundaries is supported by strong empirical evidence. However, the potential contribution of MPAs to fished populations beyond their boundaries is still highly controversial; empirical measures are scarce and modelling studies have produced a range of predictions, including both positive and negative effects. Using a combination of genetic parentage and relatedness analysis, we measured larval subsidies to local fisheries replenishment for Australasian snapper ( Chrysophrys auratus : Sparidae) from a small (5.2 km 2 ), well-established, temperate, coastal MPA in northern New Zealand. Adult snapper within the MPA contributed an estimated 10.6% (95% CI: 5.5-18.1%) of newly settled juveniles to surrounding areas (approx. 400 km 2 ), with no decreasing trend in contributions up to 40 km away. Biophysical modelling of larval dispersal matched experimental data, showing larvae produced inside the MPA dispersed over a comparable distance. These results demonstrate that temperate MPAs have the potential to provide recruitment subsidies at magnitudes and spatial scales relevant to fisheries management. The validated biophysical model provides a cost-efficient opportunity to generalize these findings to other locations and climate conditions, and potentially informs the design of MPA networks for enhancing fisheries management. © 2017 The Author(s).

  15. Intensity of Urban Heat Islands in Tropical and Temperate Climates

    Directory of Open Access Journals (Sweden)

    Margarete Cristiane de Costa Trindade Amorim

    2017-12-01

    Full Text Available Nowadays, most of the Earth’s population lives in urban areas. The replacement of vegetation by buildings and the general soil sealing, associated with human activity, lead to a rise in cities temperature, resulting in the formation of urban heat islands. This article aims to evaluate the intensity and the hourly maintenance of the atmospheric heat islands in two climates: one tropical (Presidente Prudente, Brazil and one temperate (Rennes, France throughout 2016. For this, air temperature and hourly averages were measured and calculated using both a HOBO datalogger (U23-002—protected under the same RS3 brand and weather stations Davis Vantage PRO 2. The daily evolution of the heat islands presented characteristics that varied according to the hours and seasons of the year. For both Rennes and Presidente Prudente, the largest magnitudes occurred overnight, being more greatly expressed in the tropical environment and during the driest months (winter in the tropical city and summer in the temperate one. The variability of synoptic conditions from one month to another also leads to a great heterogeneity of UHI intensity throughout the year.

  16. Carbon and Water Exchanges in a Chronosequence of Temperate White Pine Forest

    Science.gov (United States)

    Arain, M.; Restrepo, N.; Pejam, M.; Khomik, M.

    2003-12-01

    Quantification of carbon sink or source strengths of temperate forest ecosystems, growing in northern mid-latitudes, is essential to resolve uncertainties in carbon balance of the world's terrestrial ecosystems. Long-term flux measurements are needed to quantify seasonal and annual variability of carbon and water exchanges from these ecosystems and to relate the variability to environmental and physiological factors. Such long-term measurements are of particular interest for different stand developmental stages. An understanding of environmental control factors is necessary to improve predictive capabilities of terrestrial carbon and water cycles. A long-term year-round measurement program has been initiated to observe energy, water vapour, and carbon dioxide fluxes in a chronosequence of white pine (Pinus Strobus) forests in southeastern Canada. White pine is an important species in the North American landscape because of its ability to adapt to dry environments. White pine efficiently grows on coarse and sandy soils, where other deciduous and conifer species cannot survive. Generally, it is the first woody species to flourish after disturbances such as fire and clearing. The climate at the study site is temperate, with a mean annual temperature of 8 degree C and a mean annual precipitation of about 800 mm. The growing season is one of the longest in Canada, with at least 150 frost-free days. Measurements at the site began in June 2002 and are continuing at present. Flux measurements at the 60 year old stand are being made using a close-path eddy covariance (EC) system, while fluxes at the three younger stands (30, 15 and 1 year old) are being measured over 10 to 20 day periods using a roving open-path EC system Soil respiration is being measured every 2-weeks across 50-m transects at all four sites using a mobile chamber system (LI-COR 6400). The mature stand was a sink of carbon with annual NEP value of 140 g C m-2 from June 2002 to May 2003. Gross ecosystem

  17. Response of Submerged Macrophyte Communities to External and Internal Restoration Measures in North Temperate Shallow Lakes

    Science.gov (United States)

    Hilt, Sabine; Alirangues Nuñez, Marta M.; Bakker, Elisabeth S.; Blindow, Irmgard; Davidson, Thomas A.; Gillefalk, Mikael; Hansson, Lars-Anders; Janse, Jan H.; Janssen, Annette B. G.; Jeppesen, Erik; Kabus, Timm; Kelly, Andrea; Köhler, Jan; Lauridsen, Torben L.; Mooij, Wolf M.; Noordhuis, Ruurd; Phillips, Geoff; Rücker, Jacqueline; Schuster, Hans-Heinrich; Søndergaard, Martin; Teurlincx, Sven; van de Weyer, Klaus; van Donk, Ellen; Waterstraat, Arno; Willby, Nigel; Sayer, Carl D.

    2018-01-01

    Submerged macrophytes play a key role in north temperate shallow lakes by stabilizing clear-water conditions. Eutrophication has resulted in macrophyte loss and shifts to turbid conditions in many lakes. Considerable efforts have been devoted to shallow lake restoration in many countries, but long-term success depends on a stable recovery of submerged macrophytes. However, recovery patterns vary widely and remain to be fully understood. We hypothesize that reduced external nutrient loading leads to an intermediate recovery state with clear spring and turbid summer conditions similar to the pattern described for eutrophication. In contrast, lake internal restoration measures can result in transient clear-water conditions both in spring and summer and reversals to turbid conditions. Furthermore, we hypothesize that these contrasting restoration measures result in different macrophyte species composition, with added implications for seasonal dynamics due to differences in plant traits. To test these hypotheses, we analyzed data on water quality and submerged macrophytes from 49 north temperate shallow lakes that were in a turbid state and subjected to restoration measures. To study the dynamics of macrophytes during nutrient load reduction, we adapted the ecosystem model PCLake. Our survey and model simulations revealed the existence of an intermediate recovery state upon reduced external nutrient loading, characterized by spring clear-water phases and turbid summers, whereas internal lake restoration measures often resulted in clear-water conditions in spring and summer with returns to turbid conditions after some years. External and internal lake restoration measures resulted in different macrophyte communities. The intermediate recovery state following reduced nutrient loading is characterized by a few macrophyte species (mainly pondweeds) that can resist wave action allowing survival in shallow areas, germinate early in spring, have energy-rich vegetative

  18. Response of Submerged Macrophyte Communities to External and Internal Restoration Measures in North Temperate Shallow Lakes.

    Science.gov (United States)

    Hilt, Sabine; Alirangues Nuñez, Marta M; Bakker, Elisabeth S; Blindow, Irmgard; Davidson, Thomas A; Gillefalk, Mikael; Hansson, Lars-Anders; Janse, Jan H; Janssen, Annette B G; Jeppesen, Erik; Kabus, Timm; Kelly, Andrea; Köhler, Jan; Lauridsen, Torben L; Mooij, Wolf M; Noordhuis, Ruurd; Phillips, Geoff; Rücker, Jacqueline; Schuster, Hans-Heinrich; Søndergaard, Martin; Teurlincx, Sven; van de Weyer, Klaus; van Donk, Ellen; Waterstraat, Arno; Willby, Nigel; Sayer, Carl D

    2018-01-01

    Submerged macrophytes play a key role in north temperate shallow lakes by stabilizing clear-water conditions. Eutrophication has resulted in macrophyte loss and shifts to turbid conditions in many lakes. Considerable efforts have been devoted to shallow lake restoration in many countries, but long-term success depends on a stable recovery of submerged macrophytes. However, recovery patterns vary widely and remain to be fully understood. We hypothesize that reduced external nutrient loading leads to an intermediate recovery state with clear spring and turbid summer conditions similar to the pattern described for eutrophication. In contrast, lake internal restoration measures can result in transient clear-water conditions both in spring and summer and reversals to turbid conditions. Furthermore, we hypothesize that these contrasting restoration measures result in different macrophyte species composition, with added implications for seasonal dynamics due to differences in plant traits. To test these hypotheses, we analyzed data on water quality and submerged macrophytes from 49 north temperate shallow lakes that were in a turbid state and subjected to restoration measures. To study the dynamics of macrophytes during nutrient load reduction, we adapted the ecosystem model PCLake. Our survey and model simulations revealed the existence of an intermediate recovery state upon reduced external nutrient loading, characterized by spring clear-water phases and turbid summers, whereas internal lake restoration measures often resulted in clear-water conditions in spring and summer with returns to turbid conditions after some years. External and internal lake restoration measures resulted in different macrophyte communities. The intermediate recovery state following reduced nutrient loading is characterized by a few macrophyte species (mainly pondweeds) that can resist wave action allowing survival in shallow areas, germinate early in spring, have energy-rich vegetative

  19. 46 CFR 54.25-25 - Welding of quenched and tempered steels (modifies UHT-82).

    Science.gov (United States)

    2010-10-01

    ... 46 Shipping 2 2010-10-01 2010-10-01 false Welding of quenched and tempered steels (modifies UHT-82... ENGINEERING PRESSURE VESSELS Construction With Carbon, Alloy, and Heat Treated Steels § 54.25-25 Welding of quenched and tempered steels (modifies UHT-82). (a) The qualification of welding procedures, welders, and...

  20. Rumen pH and NH3-N concentration of sheep fed temperate ...

    African Journals Online (AJOL)

    The aim of this study was to evaluate the effect of sorghum grain supplementation on ruminal pH and NH. 3-N concentration of wethers consuming a fresh temperate pasture (Lotus corniculatus) in metabolism cages. Sixteen Corriedale x Milchschaf wethers were fed temperate pastures ad libitum and were ...

  1. Effects of contrasting omnivorous fish on submerged macrophyte biomass in temperate lakes: a mesocosm experiment

    NARCIS (Netherlands)

    Dorenbosch, M.; Bakker, E.S.

    2012-01-01

    1.Freshwater fish can affect aquatic vegetation directly by consuming macrophytes or indirectly by changing water quality. However, most fish in the temperate climate zone have an omnivorous diet. The impact of fish as aquatic herbivores in temperate climates therefore remains unclear and depends on

  2. The effects of boro-tempering heat treatment on microstructural properties of ductile iron

    International Nuclear Information System (INIS)

    Kayali, Yusuf; Yalcin, Yilmaz

    2011-01-01

    In this study, the effects of boro-tempering heat treatment on microstructural properties of ductile iron were investigated. Test samples with dimensions of 10 x 10 x 55 mm were boronized at 900 o C for 1, 3 and 5 h and then tempered at four different temperatures (250, 300, 350 and 450 o C) for 1 h. Both optical microscopy and scanning electron microscopy were used to reveal the microstructural details of coating and matrix of boro-tempered ductile iron. X-ray diffraction was used to determine the constituents of the coating layer. The boride layer formed on the surface of boro-tempered ductile cast iron is tooth shape form and consisted of FeB and Fe 2 B phases. The thickness of boride layer increases as the boronizing time increases and tempering temperature decreases. Tempering temperature is more effective than boronizing time on the matrix structure. Boro-tempering heat treatment reduces the formation of lower and upper ausferritic matrix temperature according to classical austempering. This causes formation of upper ausferritic matrix in the sample when tempered at 300 o C. This is in contrast to general case which is the formation of lower ausferritic matrix via austempering at this temperature.

  3. Investigation on tempering of granular bainite in an offshore platform steel

    Energy Technology Data Exchange (ETDEWEB)

    Zhou, Yanlei; Jia, Tao; Zhang, Xiangjun [The State Key Laboratory of Rolling and Automation, Northeastern University, P.O. Box 105, No. 11, Lane 3, Wenhua Road, HePing District, Shenyang 110819 (China); Liu, Zhenyu, E-mail: zyliu@mail.neu.edu.cn [The State Key Laboratory of Rolling and Automation, Northeastern University, P.O. Box 105, No. 11, Lane 3, Wenhua Road, HePing District, Shenyang 110819 (China); Misra, R.D.K. [Department of Metallurgical and Materials Engineering, University of Texas at El Paso, 500 W. University Avenue, El Paso, TX 79968-0521 (United States)

    2015-02-25

    Granular bainite, where M-A constituents dispersed in bainitic ferrite matrix usually presents at the half thickness region in thermo-mechanically processed heavy gauge offshore platform steel. In the present work, the decomposition of M-A constituents during tempering at 600 °C was firstly revealed by transmission electron microscopy (TEM) analysis, which primarily involves the precipitation of cementite, recovery and recrystallization of highly dislocated ferrite matrix. Then, the effect of tempering on mechanical properties was investigated by tempering at different temperature for 60 min. Results indicated that, at tempering temperature of 500–600 °C, large quantity of micro-alloying carbides precipitated and partially compensated the loss of strength mainly due to the decomposition of M-A constituents. Compared with the as-rolled state, the decomposition of M-A constituents and softening of bainitic ferrite matrix after tempering have resulted in higher density of microvoids and substantial plastic deformation before impact failure.

  4. Wear behavior of tempered and borided tool steels under various conditions

    International Nuclear Information System (INIS)

    Al-Haidary, T. J.; Faleh, M. N.

    2000-01-01

    . Tool steel 61CrV5, 50 NiCr13 and X1000Cr MoV51 were used in the first stage of this investigation. They have been treated as follows: boriding, boriding and tempering and hardening and tempering. The wear tests were conducted under fixed conditions (150 N/mm 2 , 0.48m/sec) with and without lubricant. The wear rate and coefficient of friction of 61Cr Si V5 steel have been studied in the second stage hoping to find the influence of working conditions on these parameters and then to compare these results with the case of hardening and tempering which is the usual case in the actual working field. The study gives a good indication about the improvement achieved in boriding and tempering cases (∼ 30%) as compared with hardening tempering cases in dry sliding conditions -∼5% with lubricating ones. (authors). 13 refs., 19 figs., 1 table

  5. Tree fern trunks facilitate seedling regeneration in a productive lowland temperate rain forest.

    Science.gov (United States)

    Gaxiola, Aurora; Burrows, Larry E; Coomes, David A

    2008-03-01

    Seedling regeneration on forest floors is often impaired by competition with established plants. In some lowland temperate rain forests, tree fern trunks provide safe sites on which tree species establish, and grow large enough to take root in the ground and persist. Here we explore the competitive and facilitative effects of two tree fern species, Cyathea smithii and Dicksonia squarrosa, on the epiphytic regeneration of tree species in nutrient-rich alluvial forests in New Zealand. The difficulties that seedlings have in establishing on vertical tree fern trunks were indicated by the following observations. First, seedling abundance was greatest on the oldest sections of tree fern trunks, near the base, suggesting that trunks gradually recruited more and more seedlings over time, but many sections of trunk were devoid of seedlings, indicating the difficulty of establishment on a vertical surface. Second, most seedlings were from small-seeded species, presumably because smaller seeds can easily lodge on tree fern trunks. Deer browsing damage was observed on 73% of epiphytic seedlings growing within 2 m of the ground, whereas few seedlings above that height were browsed. This suggests that tree ferns provide refugia from introduced deer, and may slow the decline in population size of deer-preferred species. We reasoned that tree ferns would compete with epiphytic seedlings for light, because below the tree fern canopy photosynthetically active radiation (PAR) was about 1% of above-canopy PAR. Frond removal almost tripled %PAR on the forest floor, leading to a significant increase in the height growth rate (HGR) of seedlings planted on the forest floor, but having no effects on the HGRs of epiphytic seedlings. Our study shows evidence of direct facilitative interactions by tree ferns during seedling establishment in plant communities associated with nutrient-rich soils.

  6. Long-term litter manipulation alters soil organic matter turnover in a temperate deciduous forest.

    Science.gov (United States)

    Wang, Jun-Jian; Pisani, Oliva; Lin, Lisa H; Lun, Olivia O Y; Bowden, Richard D; Lajtha, Kate; Simpson, André J; Simpson, Myrna J

    2017-12-31

    Understanding soil organic matter (OM) biogeochemistry at the molecular-level is essential for assessing potential impacts from management practices and climate change on shifts in soil carbon storage. Biomarker analyses and nuclear magnetic resonance (NMR) spectroscopy were used in an ongoing detrital input and removal treatment experiment in a temperate deciduous forest in Pennsylvania, USA, to examine how above- and below-ground plant inputs control soil OM quantity and quality at the molecular-level. From plant material to surface soils, the free acyclic lipids and cutin, suberin, and lignin biomarkers were preferentially retained over free sugars and free cyclic lipids. After 20years of above-ground litter addition (Double Litter) or exclusion (No Litter) treatments, soil OM composition was relatively more degraded, as revealed by solid-state 13 C NMR spectroscopy. Under Doubled Litter inputs, soil carbon and phospholipid fatty acid (PLFA) concentrations were unchanged, suggesting that the current OM degradation status is a reflection of microbial-mediated degradation that occurred prior to the 20-year sampling campaign. Soil OM degradation was higher in the No Litter treatments, likely due to the decline in fresh, above-ground litter inputs over time. Furthermore, root and root and litter exclusion treatments (No Roots and No Inputs, respectively) both significantly reduced free sugars and PLFAs and increased preservation of suberin-derived compounds. PLFA stress ratios and the low N-acetyl resonances from diffusion edited 1 H NMR also indicate substrate limitations and reduced microbial biomass with these treatments. Overall, we highlight that storage of soil carbon and its biochemical composition do not linearly increase with plant inputs because the microbial processing of soil OM is also likely altered in the studied forest. Copyright © 2017 Elsevier B.V. All rights reserved.

  7. Understorey productivity in temperate grassy woodland responds to soil water availability but not to elevated [CO2 ].

    Science.gov (United States)

    Collins, Luke; Bradstock, Ross A; Resco de Dios, Victor; Duursma, Remko A; Velasco, Sabrina; Boer, Matthias M

    2018-06-01

    Rising atmospheric [CO 2 ] and associated climate change are expected to modify primary productivity across a range of ecosystems globally. Increasing aridity is predicted to reduce grassland productivity, although rising [CO 2 ] and associated increases in plant water use efficiency may partially offset the effect of drying on growth. Difficulties arise in predicting the direction and magnitude of future changes in ecosystem productivity, due to limited field experimentation investigating climate and CO 2 interactions. We use repeat near-surface digital photography to quantify the effects of water availability and experimentally manipulated elevated [CO 2 ] (eCO 2 ) on understorey live foliage cover and biomass over three growing seasons in a temperate grassy woodland in south-eastern Australia. We hypothesised that (i) understorey herbaceous productivity is dependent upon soil water availability, and (ii) that eCO 2 will increase productivity, with greatest stimulation occurring under conditions of low water availability. Soil volumetric water content (VWC) determined foliage cover and growth rates over the length of the growing season (August to March), with low VWC (productivity. However, eCO 2 did not increase herbaceous cover and biomass over the duration of the experiment, or mitigate the effects of low water availability on understorey growth rates and cover. Our findings suggest that projected increases in aridity in temperate woodlands are likely to lead to reduced understorey productivity, with little scope for eCO 2 to offset these changes. © 2018 John Wiley & Sons Ltd.

  8. Effect of climate warming on the annual terrestrial net ecosystem CO2 exchange globally in the boreal and temperate regions.

    Science.gov (United States)

    Zhang, Zhiyuan; Zhang, Renduo; Cescatti, Alessandro; Wohlfahrt, Georg; Buchmann, Nina; Zhu, Juan; Chen, Guanhong; Moyano, Fernando; Pumpanen, Jukka; Hirano, Takashi; Takagi, Kentaro; Merbold, Lutz

    2017-06-08

    The net ecosystem CO 2 exchange is the result of the imbalance between the assimilation process (gross primary production, GPP) and ecosystem respiration (RE). The aim of this study was to investigate temperature sensitivities of these processes and the effect of climate warming on the annual terrestrial net ecosystem CO 2 exchange globally in the boreal and temperate regions. A database of 403 site-years of ecosystem flux data at 101 sites in the world was collected and analyzed. Temperature sensitivities of rates of RE and GPP were quantified with Q 10 , defined as the increase of RE (or GPP) rates with a temperature rise of 10 °C. Results showed that on the annual time scale, the intrinsic temperature sensitivity of GPP (Q 10sG ) was higher than or equivalent to the intrinsic temperature sensitivity of RE (Q 10sR ). Q 10sG was negatively correlated to the mean annual temperature (MAT), whereas Q 10sR was independent of MAT. The analysis of the current temperature sensitivities and net ecosystem production suggested that temperature rise might enhance the CO 2 sink of terrestrial ecosystems both in the boreal and temperate regions. In addition, ecosystems in these regions with different plant functional types should sequester more CO 2 with climate warming.

  9. Quantitative consideration for the tempering effect during multi-pass thermal cycle in HAZ of low-alloy steel

    International Nuclear Information System (INIS)

    Yu, Lina; Nakabayashi, Yuma; Saida, Kazuyoshi; Mochizuki, Masahito; Nishimoto, Kazutoshi; Kameyama, Masashi; Hirano, Shinro; Chigusa, Naoki

    2011-01-01

    A new Thermal Cycle Tempering Parameter (TCTP) to deal with the tempering effect during multi-pass thermal cycles has been proposed by extending Larson-Miller parameter (LMP). Experimental result revealed that the hardness in synthetic HAZ of the low alloy steel subjected to multi tempering thermal cycles has a good linear relationship with TCTP. By using this relationship, the hardness of the low-alloy steel reheated with tempering thermal cycles can be predicted when the original hardness is known. (author)

  10. Crown-To-Rhizosphere Carbon Transfer In A Temperate Mixed Forest

    Science.gov (United States)

    Siegwolf, R. T.; Steinmann, K.; Saurer, M.; Koerner, C.

    2005-12-01

    Flux measurements across a range of (managed) European forests showed that ecosystem respiration amounts up to 80 percent of gross primary production (Janssens et al. 2001), the rest is in large sequestered into biomass. According to Malhi et al. (1999) soil respiration accounts for 60-70 percent of total forest ecosystem respiration. A considerable part is released as CO2 via belowground plant component (autotrophic) and soil micro-organism (heterotrophic) respiration. Recent studies on the autotrophic and heterotrophic respiratory fluxes indicate that the proportion of the autotrophic respiration was most likely underestimated (Hoegberg et al, 2001). Furthermore, highly diverging lengths of time have been estimated between the synthesis of carbohydrates and their availability in the rhizosphere. The goal of the presented study was to i) estimate the transport time for new photosynthates from the leaves to the rhizosphere, ii) determine the spatial distribution of these products, and iii) detect a seasonal course in the autotrophic and heterotrophic respiration of freshly formed assimilates. This study was carried out in a temperate mixed forest (The Swiss Canopy Crane Project in Hofstetten near Basel, Switzerland, cf. Pepin and Koerner 2002, Koerner et al, 2005), exposed to an elevated mean CO2 concentration of 530 ppm. The added CO2 originated from fossil fuel combustion and was depleted in 13C, thus serving as an ideal tracer. Based on the isotopic signature of the soil CO2 it was shown that freshly assimilated carbohydrates were transferred to the rhizosphere within ca. 5 days. The spatial variability was considerable and could mostly be explained with the varying tree population, whereas, the broad-leafed area revealed a more negative d13C value than the conifers. A distinct seasonal course in soil ?13C of the CO2 concentration indicated a seasonal variation in the crown-to-rhizosphere carbon transfer Steinmann et al (2004). Hoegberg P, et al. (2001) Large

  11. Corrosion fatigue in nitrocarburized quenched and tempered steels

    Science.gov (United States)

    Khani, M. Karim; Dengel, D.

    1996-05-01

    In order to investigate the fatigue strength and fracture mechanism of salt bath nitrocarburized steels, specimens of the steels SAE 4135 and SAE 4140, in a quenched and tempered state, and additionally in a salt bath nitrocarburized and oxidizing cooled state as well as in a polished (after the oxidizing cooling) and renewed oxidized state, were subjected to comparative rotating bending fatigue tests in inert oil and 5 pct NaCl solution. In addition, some of the quenched and tempered specimens of SAE 4135 material were provided with an approximately 50-μm-thick electroless Ni-P layer, in order to compare corrosion fatigue behavior between the Ni-P layer and the nitride layers. Long-life corrosion fatigue tests of SAE 4135 material were carried out under small stresses in the long-life range up to 108 cycles with a test frequency of 100 Hz. Fatigue tests of SAE 4140 material were carried out in the range of finite life (low-cycle range) with a test frequency of 13 Hz. The results show that the 5 pct NaCl environment drastically reduced fatigue life, but nitrocarburizing plus oxidation treatment was found to improve the corrosion fatigue life over that of untreated and Ni-P coated specimens. The beneficial effect of nitrocarburizing followed by oxidation treatment on cor-rosion fatigue life results from the protection rendered by the compound layer by means of a well-sealed oxide layer, whereby the pores present in the compound layer fill up with oxides. The role of inclusions in initiating fatigue cracks was investigated. It was found that under corrosion fatigue conditions, the fatigue cracks started at cavities along the interfaces of MnS inclusions and matrix in the case of quenched and tempered specimens. The nitrocarburized specimens, however, showed a superposition of pitting corrosion and corrosion fatigue in which pores and nonmetallic inclusions in the compound layer play a predominant role concerning the formation of pits in the substrate.

  12. Soil phosphorus fractionation as a tool for monitoring dust phosphorus signature underneath a Blue Pine (Pinus wallichiana canopy in a Temperate Forest

    Directory of Open Access Journals (Sweden)

    Mustafa-Nawaz Shafqat

    2016-12-01

    Full Text Available Aims of the study: This study aims (i to monitor the amount of dust deposition during dry season in the moist temperate forest; (ii to study nature of P fractions in the dust samples falling on the trees in the region; (iii to study soil P fractions as influenced by the processes of throughfall and stemflow of a Blue Pine (Pinus wallichiana canopy and to finger print the contribution of dust towards P input in the temperate forest ecosystem. Area of study: The site used for the collection of soil samples was situated at an elevation of 6900 feet above sea levels (temperate forest in Himalaya region in the Thandani area national forest located in the north west of Pakistan. Material and methods:  For soil sampling and processing, three forest sites with three old tree plants per site were selected at approximately leveled plain for surface soil sampling. Two dust samples were collected and analyzed for different physicochemical properties along with different P fractions. First dust sample was collected from a site situated at an elevation of 4000 feet and second one was collected from an elevation of 6500 feet above sea levels. Modified Hedley procedure for the fractionation of P in the dust and soil samples were used. Main results: The input of dust was 43 and 20 kg ha-1 during drier months of the year (September-June at lower and higher elevation sites respectively, and the dust from lower elevation site had relative more all P fractions than the other dust sample. However, HCl-Pi fraction was dominant in both samples. Both labile (water plus NaHCO3 and non-labile (NaOH plus HCl inorganic P (Pi fractions were significantly increased in the surface soil by both stemflow and throughfall compared to the open field soil. The buildup of NaOH and HCl-Pi pools in soils underneath the canopy might prove useful in fingerprinting the contribution of atmospheric dust towards P cycling in this temperate forest. Research highlights: The role of dust in

  13. Microevolution of European temperate oaks in response to environmental changes.

    Science.gov (United States)

    Kremer, Antoine

    2016-01-01

    This review reconstructs microevolutionary processes that allowed long-lived species as temperate oaks (Quercus petraea and Q. robur) to cope with climate change since the last glacial maximum, by assembling insights from complementary synchronic and allochronic approaches. Paleobotanical and genetic investigations show that oaks migrated at larger velocities than expected, thanks to long-distance rare events and most likely human interferences. Hybridization was a key mechanism accelerating migration and enhancing species succession. Common garden experiments and genome wide association studies demonstrated that diversifying selection across large environmental gradients contributed to rapid local adaptation. Finally the review explores how lessons taken from past evolutionary scenarios may help to predict future responses of oaks to ongoing climate change. Copyright © 2016. Published by Elsevier SAS.

  14. Nitrogen Deposition Effects on Soil Carbon Dynamics in Temperate Forests

    DEFF Research Database (Denmark)

    Ginzburg Ozeri, Shimon

    Soils contain the largest fraction of terrestrial carbon (C). Understanding the factors regulating the decomposition and storage of soil organic matter (SOM) is essential for predictions of the C sink strength of the terrestrial environment in the light of global change. Elevated long-term nitrog...... implications for modelling the carbon sink-strength of temperate forests under global change.......Soils contain the largest fraction of terrestrial carbon (C). Understanding the factors regulating the decomposition and storage of soil organic matter (SOM) is essential for predictions of the C sink strength of the terrestrial environment in the light of global change. Elevated long-term nitrogen...... (N) deposition into forest ecosystems has been increasing globally and was hypothesized to raise soil organic C (SOC) stocks by increasing forest productivity and by reducing SOM decomposition. Yet, these effects of N deposition on forest SOC stocks are uncertain and largely based on observations...

  15. Temperance internationalism: Guy Hayler and the World Prohibition Federation.

    Science.gov (United States)

    Fahey, David M

    2006-01-01

    The World Prohibition Federation, organized in 1909, and the International Record, published from 1917 to 1968, sought to internationalize the temperance movement by collecting and disseminating anti-drink news from around the globe. The Federation was based in London, and most of its activities were English-speakers. A British reformer named Guy Hayler served as its honorary president for thirty years and edited the "International Record" until his death in 1943. The Federation emphasized that prohibitionists comprised a moral community, united regardless of race, religion, nationality, or politics. Poorly funded, the Federation had difficulty competing with the World League against Alcoholism after the Anti-Saloon League of America organized this rival propaganda society in 1919.

  16. Influence of titanium on the tempering structure of austenitic steels

    International Nuclear Information System (INIS)

    Ghuezaiel, M.J.

    1985-10-01

    The microstructure of titanium-stabilized and initially deformed (approximately 20%) austenitic stainless steels used in structures of fast neutrons reactors has been studied after one hour duration annealings (500 0 C) by X-ray diffraction, optical microscopy, microhardness and transmission electron microscopy. The studied alloys were either of industrial type CND 17-13 (0.23 to 0.45 wt% Ti) or pure steels (18% Cr, 14% Ni, 0 or 0.3 wt% Ti). During tempering, the pure steels presented some restauration before recristallization. In the industrial steels, only recristallization occurred, and this only in the most deformed steel. Precipitation does not occur in the titanium-free pure steel. In industrial steels, many intermetallic phases are formed when recristallization starts [fr

  17. Temporal flexibility of reproduction in temperate-breeding dabbling ducks

    Science.gov (United States)

    Krapu, Gary L.

    2000-01-01

    I compared nesting intervals during three consecutive years in five species of temperate-nesting dabbling ducks (Mallard [Anas platyrhynchos], Northern Pintail [Anas acuta], Northern Shoveler [Anas clypeata], Blue-winged Teal [Anas discors], Gadwall [Anas strepera]) and assessed whether differences existed in timing of refractoriness. Most nesting by females of all five species ended by the summer solstice. Nesting ended earliest for Northern Shovelers and Northern Pintails and latest for Gadwalls. Some Mallards, Blue-winged Teal, and Gadwalls continued to nest into mid- and late summer, whereas Northern Shovelers and Northern pintails did not. Mallards, Blue-winged Teal, and Gadwalls accounted for 99% (81 of 82) of flightless broods resulting from nests initiated during mid- or late summer in North Dakota and 98% (58 of 59) of flightless juveniles shot on or after 1 October by a random sample of duck hunters from across the United States. Early cessation of breeding by Northern Shovelers may have evolved in response to the species' limited flexibility in diet. Photorefractory mechanisms that limit most breeding to spring presumably evolved in response to severe constraints on reproductive success when nesting continued through summer(e.g. mortality of late-hatched young and molting females due to low temperatures). Interspecific differences in photosensitivity may account for variation in timing of cessation of nesting in late spring, but controlled experiments are needed to assess the possible role of non-photic influences. My results suggest that the refractory mechanisms controlling length of the breeding in temperate-nesting dabbling ducks are more varied and complex than previously thought, with non-photic influences (e.g. water conditions, food availability, food quality) having a larger role than indicated by earlier research.

  18. Influence of tempering on mechanical properties of ferritic martensitic steels

    International Nuclear Information System (INIS)

    Chun, Y. B.; Han, C. H.; Choi, B. K.; Lee, D. W.; Kim, T. K.; Jeong, Y. H.; Cho, S.

    2012-01-01

    In the mid-1980s research programs for development of low activation materials began. This is based on the US Nuclear Regulatory Commission Guidelines (10CFR part 61) that were developed to reduce long-lived radioactive isotopes, which allows nuclear reactor waste to be disposed of by shallow land burial when removed from service. Development of low activation materials is also key issue in nuclear fusion systems, as the structural components can became radioactive due to nuclear transmutation caused by exposure to high dose neutron irradiation. Reduced-activation ferritic martensitic (RAFM) steels have been developed in the leading countries in nuclear fusion technology, and are now being considered as primary candidate material for the test blanket module (TBM) in the international thermonuclear experiment reactor (ITER). RAFM steels developed so far (e.g., EUROFER 97 and F82H) meet the requirement for structural application in the ITER. However, if such alloys are used in the DEMO or commercial fusion reactor is still unclear, as the reactors are designed to operate under much severe conditions (i.e., higher outlet coolant temperature and neutron fluences). Such harsh operating conditions lead to development of RAFM steels with better creep and irradiation resistances. Mechanical properties of RAFM steels are strongly affected by microstructural features including the distribution, size and type of precipitates, dislocation density and grain size. For a given composition, such microstructural characteristics are determined mainly by thermo-mechanical process employed to fabricate the final product, and accordingly a final heat treatment, i.e., tempering is the key step to control the microstructure and mechanical properties. In the present work, we investigated mechanical properties of the RAFM steels with a particular attention being paid to effects of tempering on impact and creep properties

  19. Seasonality of temperate forest photosynthesis and daytime respiration.

    Science.gov (United States)

    Wehr, R; Munger, J W; McManus, J B; Nelson, D D; Zahniser, M S; Davidson, E A; Wofsy, S C; Saleska, S R

    2016-06-30

    Terrestrial ecosystems currently offset one-quarter of anthropogenic carbon dioxide (CO2) emissions because of a slight imbalance between global terrestrial photosynthesis and respiration. Understanding what controls these two biological fluxes is therefore crucial to predicting climate change. Yet there is no way of directly measuring the photosynthesis or daytime respiration of a whole ecosystem of interacting organisms; instead, these fluxes are generally inferred from measurements of net ecosystem-atmosphere CO2 exchange (NEE), in a way that is based on assumed ecosystem-scale responses to the environment. The consequent view of temperate deciduous forests (an important CO2 sink) is that, first, ecosystem respiration is greater during the day than at night; and second, ecosystem photosynthetic light-use efficiency peaks after leaf expansion in spring and then declines, presumably because of leaf ageing or water stress. This view has underlain the development of terrestrial biosphere models used in climate prediction and of remote sensing indices of global biosphere productivity. Here, we use new isotopic instrumentation to determine ecosystem photosynthesis and daytime respiration in a temperate deciduous forest over a three-year period. We find that ecosystem respiration is lower during the day than at night-the first robust evidence of the inhibition of leaf respiration by light at the ecosystem scale. Because they do not capture this effect, standard approaches overestimate ecosystem photosynthesis and daytime respiration in the first half of the growing season at our site, and inaccurately portray ecosystem photosynthetic light-use efficiency. These findings revise our understanding of forest-atmosphere carbon exchange, and provide a basis for investigating how leaf-level physiological dynamics manifest at the canopy scale in other ecosystems.

  20. Social implications of residential demand response in cool temperate climates

    International Nuclear Information System (INIS)

    Darby, Sarah J.; McKenna, Eoghan

    2012-01-01

    Residential electrical demand response (DR) offers the prospect of reducing the environmental impact of electricity use, and also the supply costs. However, the relatively small loads and numerous actors imply a large effort: response ratio. Residential DR may be an essential part of future smart grids, but how viable is it in the short to medium term? This paper reviews some DR concepts, then evaluates the propositions that households in cool temperate climates will be in a position to contribute to grid flexibility within the next decade, and that that they will allow some automated load control. Examples of demand response from around the world are discussed in order to assess the main considerations for cool climates. Different tariff types and forms of control are assessed in terms of what is being asked of electricity users, with a focus on real-time pricing and direct load control in energy systems with increasingly distributed resources. The literature points to the significance of thermal loads, supply mix, demand-side infrastructure, market regulation, and the framing of risks and opportunities associated with DR. In concentrating on social aspects of residential demand response, the paper complements the body of work on technical and economic potential. - Highlights: ► Demand response implies major change in governance of electricity systems. ► Households in cool temperate climates can be flexible, mainly with thermal loads. ► DR requires simple tariffs, appropriate enabling technology, education, and feedback. ► Need to test consumer acceptance of DR in specific conditions. ► Introduce tariffs with technologies e.g., TOU tariff plus DLC with electric vehicles.

  1. Differentiated dynamics of bud dormancy and growth in temperate fruit trees relating to bud phenology adaptation, the case of apple and almond trees

    Science.gov (United States)

    El Yaacoubi, Adnane; Malagi, Gustavo; Oukabli, Ahmed; Citadin, Idemir; Hafidi, Majida; Bonhomme, Marc; Legave, Jean-Michel

    2016-11-01

    Few studies have focused on the characterization of bud dormancy and growth dynamics for temperate fruit species in temperate and mild cropping areas, although this is an appropriate framework to anticipate phenology adaptation facing future warming contexts which would potentially combine chill declines and heat increases. To examine this issue, two experimental approaches and field observations were used for high- and low-chill apple cultivars in temperate climate of southern France and in mild climates of northern Morocco and southern Brazil. Low-chill almond cultivars offered an additional relevant plant material for comparison with apple in northern Morocco. Divergent patterns of dormancy and growth dynamics were clearly found in apple tree between southern France and southern Brazil. Divergences were less pronounced between France and Morocco. A global view outlined main differences in the dormancy chronology and intensity, the transition between endordormancy and ecodormancy and the duration of ecodormancy. A key role of bud rehydration in the transition period was shown. High-chill cultivars would be submitted in mild conditions to heterogeneous rehydration capacities linked to insufficient chill fulfillment and excessive forcing linked to high temperatures. This would favor bud competitions and consequently excessive flowering durations and weak flowering. Low chilling requirements in apple and almond would conversely confer biological capacities to tolerate superficial dormancy and abrupt transition from endordormancy to ecodormancy without important heterogeneous rehydration states within buds. It may also assume that low-chill cultivars can also tolerate high temperatures during ecodormancy as well as extended flowering durations.

  2. Differentiated dynamics of bud dormancy and growth in temperate fruit trees relating to bud phenology adaptation, the case of apple and almond trees.

    Science.gov (United States)

    El Yaacoubi, Adnane; Malagi, Gustavo; Oukabli, Ahmed; Citadin, Idemir; Hafidi, Majida; Bonhomme, Marc; Legave, Jean-Michel

    2016-11-01

    Few studies have focused on the characterization of bud dormancy and growth dynamics for temperate fruit species in temperate and mild cropping areas, although this is an appropriate framework to anticipate phenology adaptation facing future warming contexts which would potentially combine chill declines and heat increases. To examine this issue, two experimental approaches and field observations were used for high- and low-chill apple cultivars in temperate climate of southern France and in mild climates of northern Morocco and southern Brazil. Low-chill almond cultivars offered an additional relevant plant material for comparison with apple in northern Morocco. Divergent patterns of dormancy and growth dynamics were clearly found in apple tree between southern France and southern Brazil. Divergences were less pronounced between France and Morocco. A global view outlined main differences in the dormancy chronology and intensity, the transition between endordormancy and ecodormancy and the duration of ecodormancy. A key role of bud rehydration in the transition period was shown. High-chill cultivars would be submitted in mild conditions to heterogeneous rehydration capacities linked to insufficient chill fulfillment and excessive forcing linked to high temperatures. This would favor bud competitions and consequently excessive flowering durations and weak flowering. Low chilling requirements in apple and almond would conversely confer biological capacities to tolerate superficial dormancy and abrupt transition from endordormancy to ecodormancy without important heterogeneous rehydration states within buds. It may also assume that low-chill cultivars can also tolerate high temperatures during ecodormancy as well as extended flowering durations.

  3. Monitoring Spring Recovery of Photosynthesis and Spectral Reflectance in Temperate Evergreen and Mixed Deciduous Forests

    Science.gov (United States)

    Wong, C. Y.; Arain, M. A.; Ensminger, I.

    2015-12-01

    Evergreen conifers in boreal and temperate regions undergo strong seasonal changes in photoperiod and temperatures, which characterizes their photosynthetic activity with high activity in the growing season and downregulation during the winter season. Monitoring the timing of the transitions in evergreens is difficult since it's a largely invisible process, unlike deciduous trees that have a visible budding and senescence sequence. Spectral reflectance and the photochemical reflectance index (PRI), often used as a proxy for photosynthetic light-use efficiency, provides a promising tool to track the transition of evergreens between inactive and active photosynthetic states. To better understand the relationship between PRI and photosynthetic activity and to contrast this relationship between plant functional types, the spring recovery of an evergreen forest and mixed deciduous forest was monitored using spectral reflectance, chlorophyll fluorescence and gas exchange. All metrics indicate photosynthetic recovery during the spring season. These findings indicate that PRI can be used to observe the spring recovery of photosynthesis in evergreen conifers but may not be best suited for deciduous trees. These findings have implications for remote sensing, which provides a promising long-term monitoring system of whole ecosystems, which is important since their roles in the carbon cycle may shift in response to climate change.

  4. A comparative analysis of colour preferences in temperate and tropical social bees

    Science.gov (United States)

    Balamurali, G. S.; Nicholls, Elizabeth; Somanathan, Hema; Hempel de Ibarra, Natalie

    2018-02-01

    The spontaneous occurrence of colour preferences without learning has been demonstrated in several insect species; however, the underlying mechanisms are still not understood. Here, we use a comparative approach to investigate spontaneous and learned colour preferences in foraging bees of two tropical and one temperate species. We hypothesised that tropical bees utilise different sets of plants and therefore might differ in their spontaneous colour preferences. We tested colour-naive bees and foragers from colonies that had been enclosed in large flight cages for a long time. Bees were shortly trained with triplets of neutral, UV-grey stimuli placed randomly at eight locations on a black training disk to induce foraging motivation. During unrewarded tests, the bees' responses to eight colours were video-recorded. Bees explored all colours and displayed an overall preference for colours dominated by long or short wavelengths, rather than a single colour stimulus. Naive Apis cerana and Bombus terrestris showed similar choices. Both inspected long-wavelength stimuli more than short-wavelength stimuli, whilst responses of the tropical stingless bee Tetragonula iridipennis differed, suggesting that resource partitioning could be a determinant of spontaneous colour preferences. Reward on an unsaturated yellow colour shifted the bees' preference curves as predicted, which is in line with previous findings that brief colour experience overrides the expression of spontaneous preferences. We conclude that rather than determining foraging behaviour in inflexible ways, spontaneous colour preferences vary depending on experimental settings and reflect potential biases in mechanisms of learning and decision-making in pollinating insects.

  5. Flow Regime Changes: From Impounding a Temperate Lowland River to Small Hydropower Operations

    Directory of Open Access Journals (Sweden)

    Petras Punys

    2015-07-01

    Full Text Available This article discusses the environmental issues facing small hydropower plants (SHPs operating in temperate lowland rivers of Lithuania. The research subjects are two medium head reservoir type hydro schemes considered within a context of the global fleet of SHPs in the country. This research considers general abiotic indicators (flow, level, water retention time in the reservoirs of the stream that may affect the aquatic systems. The main idea was to test whether the hydrologic regime has been altered by small hydropower dams. The analysis of changes in abiotic indicators is a complex process, including both pre- and post-reservoir construction and post commissioning of the SHPs under operation. Downstream hydrograph (flow and stage ramping is also an issue for operating SHPs that can result in temporary rapid changes in flow and consequently negatively impact aquatic resources. This ramping has been quantitatively evaluated. To avoid the risk of excessive flow ramping, the types of turbines available were evaluated and the most suitable types for the natural river flow regime were identified. The results of this study are to allow for new hydro schemes or upgrades to use water resources in a more sustainable way.

  6. Litter type control on soil C and N stabilization dynamics in a temperate forest.

    Science.gov (United States)

    Hatton, Pierre-Joseph; Castanha, Cristina; Torn, Margaret S; Bird, Jeffrey A

    2015-03-01

    While plant litters are the main source of soil organic matter (SOM) in forests, the controllers and pathways to stable SOM formation remain unclear. Here, we address how litter type ((13) C/(15) N-labeled needles vs. fine roots) and placement-depth (O vs. A horizon) affect in situ C and N dynamics in a temperate forest soil after 5 years. Litter type rather than placement-depth controlled soil C and N retention after 5 years in situ, with belowground fine root inputs greatly enhancing soil C (x1.4) and N (x1.2) retention compared with aboveground needles. While the proportions of added needle and fine root-derived C and N recovered into stable SOM fractions were similar, they followed different transformation pathways into stable SOM fractions: fine root transfer was slower than for needles, but proportionally more of the remaining needle-derived C and N was transferred into stable SOM fractions. The stoichiometry of litter-derived C vs. N within individual SOM fractions revealed the presence at least two pools of different turnover times (per SOM fraction) and emphasized the role of N-rich compounds for long-term persistence. Finally, a regression approach suggested that models may underestimate soil C retention from litter with fast decomposition rates. © 2014 John Wiley & Sons Ltd.

  7. A comparative analysis of colour preferences in temperate and tropical social bees.

    Science.gov (United States)

    Balamurali, G S; Nicholls, Elizabeth; Somanathan, Hema; Hempel de Ibarra, Natalie

    2018-01-02

    The spontaneous occurrence of colour preferences without learning has been demonstrated in several insect species; however, the underlying mechanisms are still not understood. Here, we use a comparative approach to investigate spontaneous and learned colour preferences in foraging bees of two tropical and one temperate species. We hypothesised that tropical bees utilise different sets of plants and therefore might differ in their spontaneous colour preferences. We tested colour-naive bees and foragers from colonies that had been enclosed in large flight cages for a long time. Bees were shortly trained with triplets of neutral, UV-grey stimuli placed randomly at eight locations on a black training disk to induce foraging motivation. During unrewarded tests, the bees' responses to eight colours were video-recorded. Bees explored all colours and displayed an overall preference for colours dominated by long or short wavelengths, rather than a single colour stimulus. Naive Apis cerana and Bombus terrestris showed similar choices. Both inspected long-wavelength stimuli more than short-wavelength stimuli, whilst responses of the tropical stingless bee Tetragonula iridipennis differed, suggesting that resource partitioning could be a determinant of spontaneous colour preferences. Reward on an unsaturated yellow colour shifted the bees' preference curves as predicted, which is in line with previous findings that brief colour experience overrides the expression of spontaneous preferences. We conclude that rather than determining foraging behaviour in inflexible ways, spontaneous colour preferences vary depending on experimental settings and reflect potential biases in mechanisms of learning and decision-making in pollinating insects.

  8. Thermotolerance and responses to short duration heat stress in tropical and temperate species

    Science.gov (United States)

    Marias, D.; Meinzer, F. C.; Still, C. J.

    2017-12-01

    Temperature and heat waves are predicted to increase throughout the 21st century in both tropical and temperate regions. Tropical species are vulnerable to heat stress because of the higher radiation load and the narrower distribution of temperatures typically experienced compared to extratropical species. Germinant seedlings are also vulnerable to heat stress because they inhabit the boundary layer close to the soil surface where intense heating occurs. We quantified the effect of leaf age and heat stress duration (45 min, 90 min) on leaf thermotolerance and whole plant physiological responses to heat stress in Coffea arabica (COAR) saplings. We also evaluated leaf thermotolerance and whole plant responses to heat stress of seedlings in two populations each of Pinus ponderosa (PIPO) and Pseudotsuga menziesii (PSME) from contrasting climates. Thermotolerance of detached leaves/needles was evaluated using chlorophyll fluorescence (FV/FM, FO) and electrolyte leakage. After exposure of whole plants to a simulated heat wave in a growth chamber, we monitored FV/FM, photosynthesis (A), stomatal conductance (gs), non-structural carbohydrates (NSCs), and carbon isotope ratios (δ13C). In COAR, thermotolerance and rate of recovery increased with leaf age. Following heat treatment, reductions in A and gs led to reduced intrinsic water use efficiency (iWUE) and increased leaf temperatures. NSC results suggested that starch was converted to sugars for recovery from heat stress and phloem transport was inhibited. Plants failed to flower in both heat stress duration treatments. In PIPO and PSME, heat treatment induced significant reductions in FV/FM and A. NSC results suggested that starch was converted to glucose + fructose to aid recovery from heat-induced damage. Populations from drier sites had greater δ13C values than those from wetter sites, consistent with higher iWUE of populations from drier climates. Thermotolerance and heat stress responses appeared to be

  9. Non-Linear Nitrogen Cycling and Ecosystem Calcium Depletion Along a Temperate Forest Soil Nitrogen Gradient

    Science.gov (United States)

    Sinkhorn, E. R.; Perakis, S. S.; Compton, J. E.; Cromack, K.; Bullen, T. D.

    2007-12-01

    Understanding how N availability influences base cation stores is critical for assessing long-term ecosystem sustainability. Indices of nitrogen (N) availability and the distribution of nutrients in plant biomass, soil, and soil water were examined across ten Douglas-fir (Pseudotsuga menziesii) stands spanning a three-fold soil N gradient (0-10 cm: 0.21 - 0.69% N, 0-100 cm: 9.2 - 28.8 Mg N ha-1) in the Oregon Coast Range. This gradient is largely the consequence of historical inputs from N2-fixing red alder stands that can add 100-200 kg N ha-1 yr-1 to the ecosystem for decades. Annual net N mineralization and litterfall N return displayed non-linear relationships with soil N, increasing initially, and then decreasing as N-richness increased. In contrast, nitrate leaching from deep soils increased linearly across the soil N gradient and ranged from 0.074 to 30 kg N ha-1 yr-1. Soil exchangeable Ca, Mg, and K pools to 1 m depth were negatively related to nitrate losses across sites. Ca was the only base cation exhibiting concentration decreases in both plant and soil pools across the soil N gradient, and a greater proportion of total available ecosystem Ca was sequestered in aboveground plant biomass at high N, low Ca sites. Our work supports a hierarchical model of coupled N-Ca cycles across gradients of soil N enrichment, with microbial production of mobile nitrate anions leading to depletion of readily available Ca at the ecosystem scale, and plant sequestration promoting Ca conservation as Ca supply diminishes. The preferential storage of Ca in aboveground biomass at high N and low Ca sites, while critical for sustaining plant productivity, may also predispose forests to Ca depletion in areas managed for intensive biomass removal. Long-term N enrichment of temperate forest soils appears capable of sustaining an open N cycle and key symptoms of N-saturation for multiple decades after the cessation of elevated N inputs.

  10. Effect of tempering upon the tensile properties of a nanostructured bainitic steel

    Energy Technology Data Exchange (ETDEWEB)

    Hasan, H.S. [University of Technology, Baghdad (Iraq); Peet, M.J., E-mail: mjp54@cam.ac.uk [Department of Materials Science and Metallurgy, 27 Charles Babbage Road, Cambridge CB3 0FS (United Kingdom); Avettand-Fènoël, M-N. [Unité Matériaux Et Transformations (UMET) UMR CNRS 8207, Université, Lille 1, 59655 Villeneuve D' ASCQ (France); Bhadeshia, H.K.D.H. [Department of Materials Science and Metallurgy, 27 Charles Babbage Road, Cambridge CB3 0FS (United Kingdom)

    2014-10-06

    The tensile properties of a nanostructured carbide-free bainitic steel formed at 200–250 °C are compared against those after tempering sufficiently to remove the retained austenite. Although significant ductility is observed following tempering, a comparison of tempered and untempered samples shows that it is in fact reduced when a comparison is made at identical strength. The shape of the stress–strain curves shows clear evidence that the capacity for work hardening is reduced with the loss of austenite. The nanostructure of the steel transformed at 250 °C is examined by transmission electron microscopy, to compare the as-transformed to the tempered structure. In this case after tempering at 500 °C the energy absorbed during the tensile test is lower, due to the lower strength. Reduction of strength is caused by the slight coarsening of the bainite plates, and lower dislocation density after tempering. Considering the formation of carbide particles in high strength steel, impressive ductility is exhibited even in the tempered condition.

  11. Effect of tempering upon the tensile properties of a nanostructured bainitic steel

    International Nuclear Information System (INIS)

    Hasan, H.S.; Peet, M.J.; Avettand-Fènoël, M-N.; Bhadeshia, H.K.D.H.

    2014-01-01

    The tensile properties of a nanostructured carbide-free bainitic steel formed at 200–250 °C are compared against those after tempering sufficiently to remove the retained austenite. Although significant ductility is observed following tempering, a comparison of tempered and untempered samples shows that it is in fact reduced when a comparison is made at identical strength. The shape of the stress–strain curves shows clear evidence that the capacity for work hardening is reduced with the loss of austenite. The nanostructure of the steel transformed at 250 °C is examined by transmission electron microscopy, to compare the as-transformed to the tempered structure. In this case after tempering at 500 °C the energy absorbed during the tensile test is lower, due to the lower strength. Reduction of strength is caused by the slight coarsening of the bainite plates, and lower dislocation density after tempering. Considering the formation of carbide particles in high strength steel, impressive ductility is exhibited even in the tempered condition

  12. DETERMINATION OF THE OPTIMAL TEMPERING TEMPERATURE IN HARD FACING OF THE FORGING DIES

    Directory of Open Access Journals (Sweden)

    Milan Mutavdžić

    2012-06-01

    Full Text Available Here is analyzed selection of the optimal technology for heat treatment during the reparation of the damaged forging dies. Those tools are manufactured from alloyed tool steels for operation at elevated temperatures. Those steels are prone to self-hardening, so in reparatory hard-facing they must be preheated, additionally heated and tempered. During the tempering, in temperature interval 500-600°C, a secondary increase of hardness and decrease of impact toughness occurs, the so-called reversible tempering brittleness. Here is shown that it can be avoided by application of metallurgical and technological measures. Metallurgical measures assume adequate selection of steels. Since the considered steels are per se prone to tempering brittleness, we conducted experimental investigations to define the technological measures to avoid it. Tests on models were conducted: tempering from different temperatures, slow heating and cooling in still air. Hardness measurements showed that at 520°C, the secondary increase of hardness occurs, with drop of the impact toughness. Additional hard-facing tests included samples tempered at various regimes. Samples were prepared for mechanical and metallographic investigations. Results presented illustrate influence of additional heat treatment on structure, hardness and mechanical properties of the hard-faced layers. This enabled establishing the possibility of avoiding the tempering brittleness through technological measures.

  13. Determination of the optimal tempering temperature in hard facing of the forging dies

    Directory of Open Access Journals (Sweden)

    Milan Mutavdžić

    2012-05-01

    Full Text Available Here is analyzed selection of the optimal technology for heat treatment during the reparation of the damaged forging dies. Those tools are manufactured from alloyed tool steels for operation at elevated temperatures. Those steels are prone to self-hardening, so in reparatory hard-facing they must be preheated, additionally heated and tempered. During the tempering, in temperature interval 500-600°C, a secondary increase of hardness and decrease of impact toughness occurs, the so-called reversible tempering brittleness. Here is shown that it can be avoided by application of metallurgical and technological measures. Metallurgical measures assume adequate selection of steels. Since the considered steels are per se prone to tempering brittleness, we conducted experimental investigations to define the technological measures to avoid it. Tests on models were conducted: tempering from different temperatures, slow heating and cooling in still air. Hardness measurements showed that at 520°C, the secondary increase of hardness occurs, with drop of the impact toughness. Additional hard-facing tests included samples tempered at various regimes. Samples were prepared for mechanical and metallographic investigations. Results presented illustrate influence of additional heat treatment on structure, hardness and mechanical properties of the hard-faced layers. This enabled establishing the possibility of avoiding the tempering brittleness through technological measures. 

  14. Effect of tempering on the microstructure and mechanical properties of a medium carbon bainitic steel

    Energy Technology Data Exchange (ETDEWEB)

    Kang, J. [State Key Laboratory of Metastable Materials Science and Technology, Yanshan University, Qinhuangdao 066004 (China); Zhang, F.C., E-mail: zfc@ysu.edu.cn [State Key Laboratory of Metastable Materials Science and Technology, Yanshan University, Qinhuangdao 066004 (China); National Engineering Research Center for Equipment and Technology of Cold Strip Rolling, Yanshan University, Qinhuangdao 066004 (China); Yang, X.W. [State Key Laboratory of Metastable Materials Science and Technology, Yanshan University, Qinhuangdao 066004 (China); Lv, B. [College of Environmental and Chemical Engineering, Yanshan University, Qinhuangdao 066004 (China); Wu, K.M. [International Research Institute for Steel Technology, Wuhan University of Science and Technology, Wuhan 430081 (China); Materials Science and Metallurgy, University of Cambridge, Cambridge CB2 3QZ (United Kingdom)

    2017-02-16

    The effect of tempering on the microstructure and mechanical properties of a medium carbon bainitic steel has been investigated through optical microscopy, electron back-scattered diffraction, transmission electron microscopy and X-ray diffraction analyses. A nano-level microstructure containing plate-like bainitic ferrite and film-like retained austenite is obtained by isothermal transformation at Ms+10 °C followed by tempering within 240–450 °C. Results show that the sample tempered at 340 °C occupies the optimal balance of strength and toughness by maintaining a certain level of plasticity; samples tempered at 320 °C and 360 °C with low and high yield ratio come second. The microstructure of the steel is not sensitive to tempering temperatures before 360 °C. When the temperature is increased to 450 °C, the significantly coarsened bainitic ferrite plate and the occurrence of a small quantity of carbide precipitation account for its low toughness. The amount of retained austenite increases with the tempering temperature before 400 °C, followed by decreasing with further increase in the temperature. This behavior is related to the competition between retained austenite further transforming into bainite and decomposing into carbide during tempering.

  15. Reconstructing Fire Disturbances in Coastal Temperate Rainforests on the Central Coast of British Columbia, Canada

    Science.gov (United States)

    Hoffman, Kira; Smith, Dan; Lertzman, Ken; Starzomski, Brian

    2015-04-01

    The coastal temperate rainforests of British Columbia's Central Coast are comprised of old growth, mixed-age stands and a mosaic of non-forested bogs. This region receives approximately 4000 mm of annual rainfall, and fire disturbances caused by lightning are thought to be very rare. Because of the late successional characteristics of these forests and the presumed lack of visible fire evidence, fires have been estimated to occur at up to 6000-year return intervals. We attempt to distinguish the roles of natural and cultural (First Nations) fires using multiple lines of evidence from tree ring records, fire-scarred trees, soil charcoal and archaeological evidence from First Nations settlement areas. To reconstruct the Holocene fire history of the study area located on Hecate Island (N 51 38 W -128 05), thirty 400m2 forest mensuration plots were systematically established in a 287-hectare area burned in 1893. Analyses focused on the relationship between fire events and climate recorded in tree rings and instrumental records, as well as nutrient concentrations and pH of soils and plant community characteristics. Four fire events (1893, 1776, 1525, 1372) were recorded in forty-five living, fire-scarred western redcedar (Thuja plicata), yellow cedar (Xanthocyparis nootkatensis) and shore pine (Pinus contorta var. contorta) trees. Five additional fire events (1785 Cal BP, 2760 Cal BP, 3355 Cal BP, 4735 Cal BP, 7740 Cal BP) were dated with accelerated mass spectrometry radiocarbon dating of in situ macro charcoal (> 5mm) buried in stratigraphy in both organic and mineral soils. The short intervals between fire events, coupled with the long history of First Nations settlement and land use in the study area, suggest purposeful and repeated low-intensity ground fires. Our research demonstrates that fires are more widespread and common than previously recorded on the very wet Central Coast of British Columbia. It is important to incorporate cultural fires into fire history

  16. Mowing exacerbates the loss of ecosystem stability under nitrogen enrichment in a temperate grassland.

    Science.gov (United States)

    Zhang, Yunhai; Loreau, Michel; He, Nianpeng; Zhang, Guangming; Han, Xingguo

    2017-08-04

    1. Global reactive nitrogen (N) is projected to further increase in the coming years. Previous studies have demonstrated that N enrichment weakens the temporal stability of the ecosystem and the primary productivity through decreased biodiversity and species asynchrony. Mowing is a globally common practise in grasslands; and infrequent mowing can maintain or increase plant diversity under N enrichment conditions. However, it is unclear how infrequent mowing affects ecosystem stability in the face of N enrichment. 2. By independently manipulating the frequency (twice vs. monthly additions per year) and rate (i.e. 0, 1, 2, 3, 5, 10, 15, 20, and 50 g N m -2 year -1 ) of NH 4 NO 3 inputs and mowing (unmown vs. mown) over 3 years (2011-2013) in a temperate grassland of northern China, we aimed to examine the interactive effects of N enrichment and mowing on ecosystem stability. 3. The results show that mowing maintained a positive relationship between species richness and ecosystem stability despite N addition, but that it exacerbated the negative effects of N addition on ecosystem stability. Mowing increased mean primary productivity and plant species richness, but it also increased the synchrony of population fluctuations and the variability of primary productivity under N enrichment, thereby contributing to a decline in the ecosystem stability. 4. Thus, our study reveals that infrequent mowing can buffer the negative effects of N enrichment on biodiversity to some extent and further increase the primary productivity, but it exacerbates the loss of ecosystem stability with N enrichment, thereby threatening local and/or semiarid regional food security.

  17. Effects of water and nitrogen addition on species turnover in temperate grasslands in northern China.

    Directory of Open Access Journals (Sweden)

    Zhuwen Xu

    Full Text Available Global nitrogen (N deposition and climate change have been identified as two of the most important causes of current plant diversity loss. However, temporal patterns of species turnover underlying diversity changes in response to changing precipitation regimes and atmospheric N deposition have received inadequate attention. We carried out a manipulation experiment in a steppe and an old-field in North China from 2005 to 2009, to test the hypothesis that water addition enhances plant species richness through increase in the rate of species gain and decrease in the rate of species loss, while N addition has opposite effects on species changes. Our results showed that water addition increased the rate of species gain in both the steppe and the old field but decreased the rates of species loss and turnover in the old field. In contrast, N addition increased the rates of species loss and turnover in the steppe but decreased the rate of species gain in the old field. The rate of species change was greater in the old field than in the steppe. Water interacted with N to affect species richness and species turnover, indicating that the impacts of N on semi-arid grasslands were largely mediated by water availability. The temporal stability of communities was negatively correlated with rates of species loss and turnover, suggesting that water addition might enhance, but N addition would reduce the compositional stability of grasslands. Experimental results support our initial hypothesis and demonstrate that water and N availabilities differed in the effects on rate of species change in the temperate grasslands, and these effects also depend on grassland types and/or land-use history. Species gain and loss together contribute to the dynamic change of species richness in semi-arid grasslands under future climate change.

  18. Root water uptake and lateral interactions among root systems in a temperate forest

    Science.gov (United States)

    Agee, E.; He, L.; Bisht, G.; Gough, C. M.; Couvreur, V.; Matheny, A. M.; Bohrer, G.; Ivanov, V. Y.

    2016-12-01

    A growing body of research has highlighted the importance of root architecture and hydraulic properties to the maintenance of the transpiration stream under water limitation and drought. Detailed studies of single plant systems have shown the ability of root systems to adjust zones of uptake due to the redistribution of local water potential gradients, thereby delaying the onset of stress under drying conditions. An open question is how lateral interactions and competition among neighboring plants impact individual and community resilience to water stress. While computational complexity has previously hindered the implementation of microscopic root system structure and function in larger scale hydrological models, newer hybrid approaches allow for the resolution of these properties at the plot scale. Using a modified version of the PFLOTRAN model, which represents the 3-D physics of variably saturated soil, we model root water uptake in a one-hectare temperate forest plot under natural and synthetic forcings. Two characteristic hydraulic architectures, tap roots and laterally sprawling roots, are implemented in an ensemble of simulations. Variations of root architecture, their hydraulic properties, and degree of system interactions produce variable local response to water limitation and provide insights on individual and community response to changing meteorological conditions. Results demonstrate the ability of interacting systems to shift areas of active uptake based on local gradients, allowing individuals to meet water demands despite competition from their peers. These results further illustrate how inter- and intra-species variations in root properties may influence not only individual response to water stress, but also help quantify the margins of resilience for forest ecosystems under changing climate.

  19. Effect of microalloying elements on microstructure and properties of quenched and tempered constructional steel

    Science.gov (United States)

    Ma, Qingshen; Huang, Leqing; Di, Guobiao; Wang, Yanfeng; Yang, Yongda; Ma, Changwen

    2017-09-01

    The effects of microalloying elements Nb, V and Ti on microstructure and properties of quenched and tempered steel were studied. Results showed that the addition of microalloying elements led to the formation of bainite and increased strength, while the austenization and ferrite transformation temperature was barely affected, i.e. 10°C. Microalloying elements shortened the incubation time for bainite transformation by refinement of austenite grain, and decreased the hardenability by forming carbides and therefore reducing the carbon content of super-cooled austenite. Either of them promoted the bainite transformation. The better tempering stability was ascribed to the as hot-rolled bainite microstructure and secondary carbide precipitation during tempering.

  20. Internal strains after recovery of hardness in tempered martensitic steels for fusion reactors

    Science.gov (United States)

    Brunelli, L.; Gondi, P.; Montanari, R.; Coppola, R.

    1991-03-01

    After tempering, with recovery of hardness, MANET steels present internal strains; these residual strains increase with quenching rate prior to tempering, and they remain after prolonged tempering times. On account of their persistence, after thermal treatments which lead to low dislocation and sub-boundary densities, the possibility has been considered that the high swelling resistance of MANET is connected with these centres of strain, probably connected with the formation, in ferrite, of Cr-enriched and contiguous Cr-depleted zones which may act as sinks for interstitials. Comparative observations on the internal strain behaviour of cold worked 316L stainless steel appear consistent with this possibility.

  1. On the tempered martensite embrittlement in AISI 4140 low alloy steel

    Energy Technology Data Exchange (ETDEWEB)

    Darwish, F.A. (Dept. of Materials Science and Metallurgy, Catholic Univ., Rio de Janeiro, RJ (Brazil)); Pereira, L.C.; Gatts, C. (Dept. of Metallurgy and Materials Engineering, Federal Univ., Rio de Janeiro, RJ (Brazil)); Graca, M.L. (Materials Div., Technical Aerospace Center, Sao Jose dos Campos, SP (Brazil))

    1991-02-01

    In the present investigation the Auger electron spectroscopy (AES) technique was used to determine local carbon and phosphorus concentrations on the fracture surfaces of as-quenched and quenched-and-tempered (at 350deg C) AISI 4140 steel specimens austenitized at low and high temperatures. The AES results were rationalized to conclude that, although carbide growth as well as phosphorus segregation are expected to contribute to tempered martensite embrittlement, carbide precipitation on prior austenite grain boundaries during tempering is seen to be the microstructural change directly responsible for the occurrence of the referred embrittlement phenomenon. (orig.).

  2. Effect of tempering time on the ballistic performance of a high strength armour steel

    OpenAIRE

    Jena, Pradipta Kumar; Senthil P., Ponguru; K., Siva Kumar

    2016-01-01

    The investigation describes and analyses the effect of tempering time on the mechanical and ballistic performance of a high strength armour steel. The steel is subjected to tempering at 300 °C for 2, 24 and 48 h. A marginal variation in strength and hardness is observed with increase in tempering time, whereas ductility and Charpy impact values are found to be decreasing. Ballistic performance of the samples are evaluated by impacting 7.62 mm and 12.7 mm armour piercing projectiles at 0° angl...

  3. Synchrotron micro-diffraction analysis of the microstructure of cryogenically treated high performance tool steels prior to and after tempering

    Energy Technology Data Exchange (ETDEWEB)

    Xu, N.; Cavallaro, G.P. [Applied Centre for Structural and Synchrotron Studies, Mawson Lakes Blvd, University of South Australia, Mawson Lakes, South Australia 5095 (Australia); Gerson, A.R., E-mail: Andrea.Gerson@unisa.edu.au [Applied Centre for Structural and Synchrotron Studies, Mawson Lakes Blvd, University of South Australia, Mawson Lakes, South Australia 5095 (Australia)

    2010-10-15

    The phase transformation and strain changes within cryogenically (-196 deg. C) treated high performance tool steels (AISI H13) before and after tempering have been examined using both laboratory XRD and synchrotron micro-diffraction. The martensitic unit cell was found to have very low tetragonality as expected for low carbon steel. Tempering resulted in the diffusion of excess carbon out of the martensite phase and consequent unit cell shrinkage. In addition on tempering the martensite became more homogeneous as compared to the same samples prior to tempering. For cryogenically treated samples, the effect was most pronounced for the rapidly cooled sample which was the least homogenous sample prior to tempering but was the most homogenous sample after tempering. This suggests that the considerable degree of disorder resulting from rapid cryogenic cooling results in the beneficial release of micro-stresses on tempering thus possibly resulting in the improved wear resistance and durability observed for cryogenically treated tool steels.

  4. Synchrotron micro-diffraction analysis of the microstructure of cryogenically treated high performance tool steels prior to and after tempering

    International Nuclear Information System (INIS)

    Xu, N.; Cavallaro, G.P.; Gerson, A.R.

    2010-01-01

    The phase transformation and strain changes within cryogenically (-196 deg. C) treated high performance tool steels (AISI H13) before and after tempering have been examined using both laboratory XRD and synchrotron micro-diffraction. The martensitic unit cell was found to have very low tetragonality as expected for low carbon steel. Tempering resulted in the diffusion of excess carbon out of the martensite phase and consequent unit cell shrinkage. In addition on tempering the martensite became more homogeneous as compared to the same samples prior to tempering. For cryogenically treated samples, the effect was most pronounced for the rapidly cooled sample which was the least homogenous sample prior to tempering but was the most homogenous sample after tempering. This suggests that the considerable degree of disorder resulting from rapid cryogenic cooling results in the beneficial release of micro-stresses on tempering thus possibly resulting in the improved wear resistance and durability observed for cryogenically treated tool steels.

  5. Seasonal pattern of anthropogenic salinization in temperate forested headwater streams.

    Science.gov (United States)

    Timpano, Anthony J; Zipper, Carl E; Soucek, David J; Schoenholtz, Stephen H

    2018-04-15

    Salinization of freshwaters by human activities is of growing concern globally. Consequences of salt pollution include adverse effects to aquatic biodiversity, ecosystem function, human health, and ecosystem services. In headwater streams of the temperate forests of eastern USA, elevated specific conductance (SC), a surrogate measurement for the major dissolved ions composing salinity, has been linked to decreased diversity of aquatic insects. However, such linkages have typically been based on limited numbers of SC measurements that do not quantify intra-annual variation. Effective management of salinization requires tools to accurately monitor and predict salinity while accounting for temporal variability. Toward that end, high-frequency SC data were collected within the central Appalachian coalfield over 4 years at 25 forested headwater streams spanning a gradient of salinity. A sinusoidal periodic function was used to model the annual cycle of SC, averaged across years and streams. The resultant model revealed that, on average, salinity deviated approximately ±20% from annual mean levels across all years and streams, with minimum SC occurring in late winter and peak SC occurring in late summer. The pattern was evident in headwater streams influenced by surface coal mining, unmined headwater reference streams with low salinity, and larger-order salinized rivers draining the study area. The pattern was strongly responsive to varying seasonal dilution as driven by catchment evapotranspiration, an effect that was amplified slightly in unmined catchments with greater relative forest cover. Evaluation of alternative sampling intervals indicated that discrete sampling can approximate the model performance afforded by high-frequency data but model error increases rapidly as discrete sampling intervals exceed 30 days. This study demonstrates that intra-annual variation of salinity in temperate forested headwater streams of Appalachia USA follows a natural seasonal

  6. Fructan biosynthesis in crop plants : the molecular regulation of fructan biosynthesis in chicory (Cichorium intybus L.)

    NARCIS (Netherlands)

    Arkel, van J.

    2013-01-01

    Fructan is a polymer of fructose produced by plants and microorganisms. Within the plant kingdom about 45.000 species accumulate fructan as storage carbohydrate in addition to, or instead of, starch. Fructan accumulating species are mainly found in temperate and sub-tropical regions with

  7. The geology, botany and chemistry of selected peat-forming environments from temperate and tropical latitudes

    Science.gov (United States)

    Cameron, C.C.; Esterle, J.S.; Palmer, C.A.

    1989-01-01

    Peat has been studied in several geologic settings: (1) glaciated terrain in cold temperate Maine and Minnesota, U.S.A.; (2) an island in the Atlantic Ocean off the coast of Maine, where sea level is rising; (3) the warm temperate U.S. Atlantic and Gulf Coastal Plains, where sea level has changed often; and (4) the tropical coast of Sarawak, Malaysia, and the tropical delta of the Batang Hari River, Sumatra, Indonesia. Most of these deposits are domed (ombrotrophic or partly ombrotrophic) bogs in which peat accumulation continued above the surface of the surrounding soil. However, the bogs of the U.S. Atlantic and Gulf Coastal Plains are comparatively not as domed, and many have almost level surfaces. In some bogs, aquatic or semi-aquatic plant materials accumulated, replaced water in the depressions, and formed a surface on which marsh or swamp vegetation could subsequently live, die, and accumulate. In others, the plant materials accumulated initially on level silt or sand surfaces supporting marshes or swamps. As the peat dome formed, plants growing on it changed from luxuriant ones near the base of the dome, where nutrients were brought into the bog by surface and ground water, to stunted ones at the top of the dome, where the raised bogs are fed by nutrient-poor precipitation. The physical and chemical changes that take place in the sequence of environments from the pond stage of deposit development, through the grassy marsh stage, through the forested swamp stage, and finally through the heath dome stage can be measured in terms of acidity and ash, volatile matter, carbon, hydrogen, nitrogen, sulfur and oxygen contents, as well as in the kind and distribution of trace elements. The organic and inorganic contents of the deposits relate to geomorphology, and geomorphology relates to their settings. As models of coal formation, some domed peat deposits may help in solving problems of distribution and character of ancient coal beds. But clearly not all peat

  8. In Situ Study of Phase Transformations during Non-Isothermal Tempering of Bainitic and Martensitic Microstructures

    Directory of Open Access Journals (Sweden)

    S. Hesamodin Talebi

    2017-09-01

    Full Text Available Phase transformations during non-isothermal tempering of bainitic or martensitic microstructures obtained after quenching of a medium-carbon low-alloy steel was studied. The microstructures correspond to different locations of an as-quenched large-sized forged ingot used as a die material in the automotive industry. High-resolution dilatometry experiments were conducted to simulate the heat treatment process, as well as to investigate different phenomena occurring during non-isothermal tempering. The microstructures were characterized using optical and scanning electron microscopy. Dilatometry analyses demonstrated that tempering behavior varied significantly from bainitic to martensitic microstructures. Retained austenite, which exists between bainitic ferrite sheaves, decomposes to lower bainite causing a remarkable volume increase. It was found that this decomposition finishes below 386 °C. By contrast, martensite tempering was accompanied with a volume decrease due to the decomposition of medium-carbon martensite to low carbon martensite and carbides.

  9. A common framework for greenhouse gas assessment protocols in temperate agroforestry systems: Connecting via GRACEnet

    Science.gov (United States)

    Agroforestry systems offer many ecosystem benefits, but such systems have previously been marginalized in temperate environments due to overriding economic goals and perceived management complexity. In view of adaptation to a changing climate, agroforestry systems offer advantages that require quan...

  10. Understanding seagrass resilience in temperate systems: the importance of timing of the disturbance

    NARCIS (Netherlands)

    Soissons, L.M.; Li, B.; Han, Q.; van Katwijk, M.M.; Ysebaert, T.; Herman, P.M.J.; Bouma, T.J.

    2016-01-01

    Temperate seagrass meadows form valuable ecosystems in coastal environments and present a distinctseasonal growth. They are threatened by an increasing amount of stressors, potentially affecting theircapacity to recover from disturbances. We hypothesized that their resilience to disturbances is

  11. Understanding seagrass resilience in temperate systems: the importance of timing of the disturbance

    NARCIS (Netherlands)

    Soissons, L.M.; Li, B.Q.; Han, Q.Y.; Katwijk, M.M. van; Ysebaert, T.; Herman, P.M.J.; Bouma, T.J.

    2016-01-01

    Temperate seagrass meadows form valuable ecosystems in coastal environments and present a distinct seasonal growth. They are threatened by an increasing amount of stressors, potentially affecting their capacity to recover from disturbances. We hypothesized that their resilience to disturbances is

  12. Studies on growth and age of bivalves from temperate and tropical estuarine ecosystems

    Digital Repository Service at National Institute of Oceanography (India)

    Parulekar, A.H.

    Comparison of growth progression and age composition of Abra alba and Nuculana minuta from temperate estuarine ecosystem with Meretrix casta and Paphia malabarica from tropical estuarine environment, revealed that the annual growth rate in tropical...

  13. Analysis of mechanical properties of steel 1045 plasma nitriding: with and without tempering

    International Nuclear Information System (INIS)

    Machado, N.T.B.; Passos, M.L.M. dos; Riani, J.C.; Recco, A.A.C.

    2014-01-01

    The purpose of this study was to evaluate the possibility of tempering during the nitriding of AISI 1045 steel. The objective was to evaluate the possibility of eliminating this phase, with the nitriding properties remaining unaltered. For this, three parameter samples were compared: quenched, tempered and nitrided for 2h; quenching and nitrided for 2h and quenching and nitrided for 4h. The analysis techniques used for characterizing the samples before and after nitriding were optical microscopy, hardness Rockwell C (HRC), scanning electron microscopy (SEM), X-ray diffraction (XRD). Results showed that phase γ is the most favorable of all parameters tested. The hardness assays showed that samples with different initial hardness (with and without tempering) and even nitriding time showed similar mechanical properties. This fact suggests that the tempering process occurred parallel to the nitriding process. (author)

  14. Evolutionary and ecological differentiation in the pantropical to warm-temperate seaweed Digenea simplex (Rhodophyta)

    NARCIS (Netherlands)

    Pakker, H; Klerk, H; vanCampen, JH; Olsen, JL; Breeman, AM

    Genetic differentiation among geographic isolates of the pantropical to warm-temperate red alga Digenea simplex (Wulfen) C. Agardh was investigated using random amplified polymorphic DNA (RAPD) markers, crossing studies, and temperature tolerances experiments. Eleven isolates representing

  15. Can Detectability Analysis Improve the Utility of Point Counts for Temperate Forest Raptors?

    Science.gov (United States)

    Temperate forest breeding raptors are poorly represented in typical point count surveys because these birds are cryptic and typically breed at low densities. In recent years, many new methods for estimating detectability during point counts have been developed, including distanc...

  16. Comparison of the segregation behavior between tempered martensite and tempered bainite in Ni-Cr-Mo high strength low alloy RPV steel

    Energy Technology Data Exchange (ETDEWEB)

    Park, Sang Gyu; Kim, Min Chul; Kim, Hyung Jun; Lee, Bong Sang [Korea Atomic Energy Research Institute, Daejeon (Korea, Republic of)

    2012-05-15

    SA508 Gr.4N Ni-Cr-Mo low alloy steel has an superior fracture toughness and strength, compared to commercial Mn-Mo-Ni low alloy RPV steel SA508 Gr.3. Higher strength and fracture toughness of low alloy steels could be obtained by adding Ni and Cr. So several were performed on researches on SA508 Gr.4N low alloy steel for a RPV application. The operation temperature and term of a reactor pressure vessel is more than 300 .deg. C and over 40 years. Therefore, in order to apply the SA508 Gr.4N low alloy steel for a reactor pressure vessel, the resistance of thermal embrittlement in the high temperature range including temper embrittlement is required. S. Raoul reported that the susceptibility to temper embrittlement was increasing a function of the cooling rate in SA533 steel, which suggests the martensitic microstructures resulting from increased cooling rates are more susceptible to temper embrittlement. However, this result has not been proved yet. So the comparison of temper embrittlement behavior was made between martensitic microstructure and bainitic microstructure with a viewpoint of boundary features in SA508 Gr.4N, which have mixture of tempered bainite/martensite. We have compared temper embrittlement behaviors of SA508 Gr.4N low alloy steel with changing volume fraction of martensite. The mechanical properties of these low alloy steels were evaluated after a long-term heat treatment. Then, the the segregated boundaries were observed and segregation behavior was analyzed by AES. In order to compare the misorientation distributions of model alloys, grain boundary structures were measured with EBSD

  17. Sorption of pesticides in tropical and temperate soils from Australia and the Philippines.

    Science.gov (United States)

    Oliver, Danielle P; Kookana, Rai S; Quintana, Belen

    2005-08-10

    The sorption behavior of diuron, imidacloprid, and thiacloprid was investigated using 22 soils collected in triplicate from temperate environments in Australia and tropical environments in Australia and the Philippines. Within the temperate environment in Australia, the soils were selected from a range of land uses. The average KOC values (L/kg) for imidacloprid were 326, 322, and 336; for thiacloprid, the values were 915, 743, and 842; and for diuron, the values were 579, 536, and 618 for the Ord (tropical), Mt. Lofty (temperate), and Philippines (tropical) soils, respectively. For all soils, the sorption coefficients decreased in the following order: thiacloprid > diuron > imidacloprid. There were no significant differences in sorption behavior between the tropical soils from the Philippines and the temperate soils from Australia. Sorption was also not significantly related with soil characteristics, namely, organic carbon (OC) content, clay content, and pH, for any of the three chemicals studied. When the data were sorted into separate land uses, the sorption of all three chemicals was highly correlated (P soils from the Philippines. Sorption coefficients for all three chemicals were highly correlated with OC in temperate, native soils only when one extreme value was removed. The relationships between sorption of all three chemicals and OC in temperate, pasture soils were best described by a polynomial. Sorption coefficients for imidacloprid and thiacloprid determined in the temperate pasture soils remained fairly consistent as the OC content increased from 3.3 to 5.3%, indicating that, although the total OC in the pasture soils was increasing, the component of OC involved with sorption of these two compounds may have been remaining constant. This study demonstrated that the origin of the soils (i.e., temperate vs tropical) had no significant effect on the sorption behavior, but in some cases, land use significantly affected the sorption behavior of the three

  18. Synchronous dynamics of zooplankton competitors prevail in temperate lake ecosystems.

    Science.gov (United States)

    Vasseur, David A; Fox, Jeremy W; Gonzalez, Andrew; Adrian, Rita; Beisner, Beatrix E; Helmus, Matthew R; Johnson, Catherine; Kratina, Pavel; Kremer, Colin; de Mazancourt, Claire; Miller, Elizabeth; Nelson, William A; Paterson, Michael; Rusak, James A; Shurin, Jonathan B; Steiner, Christopher F

    2014-08-07

    Although competing species are expected to exhibit compensatory dynamics (negative temporal covariation), empirical work has demonstrated that competitive communities often exhibit synchronous dynamics (positive temporal covariation). This has led to the suggestion that environmental forcing dominates species dynamics; however, synchronous and compensatory dynamics may appear at different length scales and/or at different times, making it challenging to identify their relative importance. We compiled 58 long-term datasets of zooplankton abundance in north-temperate and sub-tropical lakes and used wavelet analysis to quantify general patterns in the times and scales at which synchronous/compensatory dynamics dominated zooplankton communities in different regions and across the entire dataset. Synchronous dynamics were far more prevalent at all scales and times and were ubiquitous at the annual scale. Although we found compensatory dynamics in approximately 14% of all combinations of time period/scale/lake, there were no consistent scales or time periods during which compensatory dynamics were apparent across different regions. Our results suggest that the processes driving compensatory dynamics may be local in their extent, while those generating synchronous dynamics operate at much larger scales. This highlights an important gap in our understanding of the interaction between environmental and biotic forces that structure communities. © 2014 The Author(s) Published by the Royal Society. All rights reserved.

  19. Soil Respiration in Semiarid Temperate Grasslands under Various Land Management.

    Directory of Open Access Journals (Sweden)

    Zhen Wang

    Full Text Available Soil respiration, a major component of the global carbon cycle, is significantly influenced by land management practices. Grasslands are potentially a major sink for carbon, but can also be a source. Here, we investigated the potential effect of land management (grazing, clipping, and ungrazed enclosures on soil respiration in the semiarid grassland of northern China. Our results showed the mean soil respiration was significantly higher under enclosures (2.17 μmol.m(-2.s(-1 and clipping (2.06 μmol.m(-2.s(-1 than under grazing (1.65 μmol.m-(2.s(-1 over the three growing seasons. The high rates of soil respiration under enclosure and clipping were associated with the higher belowground net primary productivity (BNPP. Our analyses indicated that soil respiration was primarily related to BNPP under grazing, to soil water content under clipping. Using structural equation models, we found that soil water content, aboveground net primary productivity (ANPP and BNPP regulated soil respiration, with soil water content as the predominant factor. Our findings highlight that management-induced changes in abiotic (soil temperature and soil water content and biotic (ANPP and BNPP factors regulate soil respiration in the semiarid temperate grassland of northern China.

  20. Seasonal sediment dynamics shape temperate bedrock reef communities

    Science.gov (United States)

    Figurski, Jared D.; Freiwald, Jan; Lonhart, Steve I.; Storlazzi, Curt

    2016-01-01

    Mobilized seafloor sediment can impact benthic reef communities through burial, scour, and turbidity. These processes are ubiquitous in coastal oceans and, through their influence on the survival, fitness, and interactions of species, can alter the structure and function of benthic communities. In northern Monterey Bay, California, USA, as much as 30% of the seafloor is buried or exposed seasonally, making this an ideal location to test how subtidal temperate rocky reef communities vary in the presence and absence of chronic sediment-based disturbances. Designated dynamic plots were naturally inundated by sediment in summer (50 to 100% cover) and swept clean in winter, whereas designated stable plots remained free of sediment during our study. Multivariate analyses indicated significant differences in the structure of sessile and mobile communities between dynamic and stable reef habitats. For sessile species, community structure in disturbed plots was less variable in space and time than in stable plots due to the maintenance of an early successional state. In contrast, community structure of mobile species varied more in disturbed plots than in stable plots, reflecting how mobile species distribute in response to sediment dynamics. Some species were found only in these disturbed areas, suggesting that the spatial mosaic of disturbance could increase regional diversity. We discuss how the relative ability of species to tolerate disturbance at different life history stages and their ability to colonize habitat translate into community-level differences among habitats, and how this response varies between mobile and sessile communities.

  1. Winter habitat occurrence patterns of temperate migrant birds in Belize

    Science.gov (United States)

    Dawson, D.K.; Robbins, C.S.; Sauer, J.R.

    1992-01-01

    We used mist nets and point counts to sample bird populations in 61 sites in Belize during January-March of 1987-1991. Sites were classified as forest, second growth, woody agricultural crops (citrus, mango, cacao, and cashew), or non-woody agricultural crops (rice and sugar cane). We evaluated patterns of occurence of wintering temperate migrant bird species in these habitats. Mist net captures of 22 of 31 migrant species differed significantly among habitats. Of these, 13 species were captured more frequently in the agricultural habitats. American Redstart (Setophaga ruticilla), Black-and-white Warbler (Mniotilta varia), and Magnolia Warbler (Dendroica magnolia) were among the species captured most frequently in woody agricultural habitats; captures of Common Yellowthroat (Geothlypis trichas), Indigo Bunting (Passerina cyanea), and Northern (lcterus galbula) and Orchard orioles (I. spur/anus) were highest in the non-woody agricultural sites. We relate these occurrence patterns to trends in breeding populations in North America. While count data provide a wide picture of winter habitat distribution of migrants, more intensive work is necessary to assess temporal and geographic variation of migrant bird use of agricultural habitats.

  2. Ecosystem respiration depends strongly on photosynthesis in a temperate heath

    DEFF Research Database (Denmark)

    Larsen, Klaus Steenberg; Ibrom, Andreas; Beier, Claus

    2007-01-01

    We measured net ecosystem CO2 flux (F-n) and ecosystem respiration (R-E), and estimated gross ecosystem photosynthesis (P-g) by difference, for two years in a temperate heath ecosystem using a chamber method. The exchange rates of carbon were high and of similar magnitude as for productive forest...... ecosystems with a net ecosystem carbon gain during the second year of 293 +/- 11 g C m(-2) year(-1) showing that the carbon sink strength of heather-dominated ecosystems may be considerable when C. vulgaris is in the building phase of its life cycle. The estimated gross ecosystem photosynthesis and ecosystem.......65) was improved when the P-g rate was incorporated into the model (second year; R-2 = 0.79), suggesting that daytime R-E increased with increasing photosynthesis. Furthermore, the temperature sensitivity of R-E decreased from apparent Q(10) values of 3.3 to 3.9 by the classic equation to a more realistic Q(10...

  3. Reasons to temper enthusiasm about open access nursing journals.

    Science.gov (United States)

    de Jong, Gideon

    2017-04-01

    Open access is a relatively new phenomenon within nursing science. Several papers from various nursing journals have been published recently on the disadvantages of the traditional model of purchasing proprietary fee-based databases to access scholarly information. Just few nursing scholars are less optimistic about the possible benefits of open access nursing journals. A critical reflection on the merits and pitfalls of open access journals along insights from the literature and personal opinion. Two arguments are discussed, providing justification for tempering enthusiasm about open access journals. First, only research groups with sufficient financial resources can publish in open access journals. Second, open access has conflicting incentives, where the aim is to expand production at the expense of publishing quality articles; a business model that fits well into a neoliberal discourse. There are valid reasons to criticise the traditional publishers for the excessive costs of a single article, therefore preventing the dissemination of scholarly nursing information. On the contrary, the business model of open access publishers is no less imbued with the neoliberal tendency of lining the pockets.

  4. Soil Respiration in Semiarid Temperate Grasslands under Various Land Management.

    Science.gov (United States)

    Wang, Zhen; Ji, Lei; Hou, Xiangyang; Schellenberg, Michael P

    2016-01-01

    Soil respiration, a major component of the global carbon cycle, is significantly influenced by land management practices. Grasslands are potentially a major sink for carbon, but can also be a source. Here, we investigated the potential effect of land management (grazing, clipping, and ungrazed enclosures) on soil respiration in the semiarid grassland of northern China. Our results showed the mean soil respiration was significantly higher under enclosures (2.17 μmol.m(-2).s(-1)) and clipping (2.06 μmol.m(-2).s(-1)) than under grazing (1.65 μmol.m-(2).s(-1)) over the three growing seasons. The high rates of soil respiration under enclosure and clipping were associated with the higher belowground net primary productivity (BNPP). Our analyses indicated that soil respiration was primarily related to BNPP under grazing, to soil water content under clipping. Using structural equation models, we found that soil water content, aboveground net primary productivity (ANPP) and BNPP regulated soil respiration, with soil water content as the predominant factor. Our findings highlight that management-induced changes in abiotic (soil temperature and soil water content) and biotic (ANPP and BNPP) factors regulate soil respiration in the semiarid temperate grassland of northern China.

  5. Mercury emission from a temperate lake during autumn turnover

    International Nuclear Information System (INIS)

    Wollenberg, Jennifer L.; Peters, Stephen C.

    2009-01-01

    Lakes in temperate regions stratify during summer and winter months, creating distinct layers of water differentiated by their physical and chemical characteristics. When lakes mix in autumn and spring, mercury cycling may be affected by the chemical changes that occur during mixing. Sampling was conducted in Lake Lacawac, Eastern Pennsylvania, USA, throughout the autumn of 2007 to characterize changes in emission of gaseous elemental mercury (Hg 0 ) from the lake surface and dissolved mercury profiles in the water column during mixing. Water chemistry and weather parameters were also measured, including dissolved organic carbon (DOC), iron, and solar radiation which have been shown to interact with mercury species. Results indicate that emission of Hg 0 from the lake to the atmosphere during turnover was controlled both by solar radiation and by surface water mercury concentration. As autumn turnover progressed through the months of October and November, higher mercury concentration water from the hypolimnion mixed with epilimnetic water, increasing mercury concentration in epilimnetic waters. Dissolved absorbance was significantly correlated with mercury concentrations and with iron, but DOC concentrations were essentially constant throughout the study period and did not exhibit a relationship with either dissolved mercury concentrations or emission rates. Positive correlations between dissolved mercury and iron and manganese also suggest a role for these elements in mercury transport within the lake, but iron and manganese did not demonstrate a relationship with emission rates. This research indicates that consideration of seasonal processes in lakes is important when evaluating mercury cycling in aquatic systems

  6. Growth mindset tempers the effects of poverty on academic achievement.

    Science.gov (United States)

    Claro, Susana; Paunesku, David; Dweck, Carol S

    2016-08-02

    Two largely separate bodies of empirical research have shown that academic achievement is influenced by structural factors, such as socioeconomic background, and psychological factors, such as students' beliefs about their abilities. In this research, we use a nationwide sample of high school students from Chile to investigate how these factors interact on a systemic level. Confirming prior research, we find that family income is a strong predictor of achievement. Extending prior research, we find that a growth mindset (the belief that intelligence is not fixed and can be developed) is a comparably strong predictor of achievement and that it exhibits a positive relationship with achievement across all of the socioeconomic strata in the country. Furthermore, we find that students from lower-income families were less likely to hold a growth mindset than their wealthier peers, but those who did hold a growth mindset were appreciably buffered against the deleterious effects of poverty on achievement: students in the lowest 10th percentile of family income who exhibited a growth mindset showed academic performance as high as that of fixed mindset students from the 80th income percentile. These results suggest that students' mindsets may temper or exacerbate the effects of economic disadvantage on a systemic level.

  7. Quantifying the effects of tempering on individual phase properties of DP980 steel with nanoindentation

    Energy Technology Data Exchange (ETDEWEB)

    Cheng, G.; Zhang, F.; Ruimi, A.; Field, D. P.; Sun, X.

    2016-06-01

    We conduct a series of thermal and mechanical testing on a commercial dual phase (DP) 980 steel in order to quantify the effects of tempering on its individual phase properties. Tempering treatment is conducted at 250 °C and 400 °C for 60 minutes each. Ferrite and martensite grains are distinguished using electron backscatter diffraction (EBSD) and scanning probe microscopy (SPM), and the martensite volume fractions (MVF) are determined based on the image quality (IQ) map. Multi-scale indentation tests combined with a newly developed inverse method are used to obtain the individual phase flow properties in each tempered DP980 sample. The results show that, i) tempering significantly reduces martensite yield strength, while it only slightly reduces the ferrite yield strength; ii) tempering temperature has a more significant influence on the work hardening exponent of ferrite than that of martensite; iii) the elastic modulus of martensite is consistently higher than that of ferrite. As a validation, a simple rule of mixtures is used to verify the above-predicted individual phase flow stresses with the experimentally obtained overall true stress vs. true strain curves. The methodology and the corresponding results shown in this study can help guide the selection of tempering parameters in optimizing the mechanical properties of DP steels for their intended applications.

  8. Influence of Mo addition on the tempered properties of 13Cr martensitic stainless steel

    International Nuclear Information System (INIS)

    Jung, Byong Ho; Ahn, Yong Sik

    1998-01-01

    In order to investigate the effect of Mo addition on the mechanical properties of 13Cr-0.2C martensitic stainless steel, tensile test and Charpy V-notch test were performed after tempering at the temperature range of 200∼700 .deg. C following austenitizing at 1100 .deg. C. The yield strength and hardness of the steel were increased with the increase of Mo content at all tempering conditions, because Mo causes retardation of precipitation and coarsening of carbides and solid solution strengthening of matrix. Except 500 .deg. C of tempering temperature, the Charpy impact energy was significantly increased with Mo content and showed the highest value at 1.5 wt% addition. The increase of impact energy of the steel containing Mo is thought to be caused by δ-ferrite formed in the tempered martensitic matrix. At 500 .deg. C tempering, Charpy impact energy was decreased drastically due to temper embrittlement and it was not possible to prevent it even though Mo was added up to 1.5 wt%

  9. A Novel Methods for Fracture Toughness Evaluation of Tool Steels with Post-Tempering Cryogenic Treatment

    Directory of Open Access Journals (Sweden)

    Ramona Sola

    2017-02-01

    Full Text Available Cryogenic treatments are usually carried out immediately after quenching, but their use can be extended to post tempering in order to improve their fracture toughness. This research paper focuses on the influence of post-tempering cryogenic treatment on the microstructure and mechanical properties of tempered AISI M2, AISI D2, and X105CrCoMo18 steels. The aforementioned steels have been analysed after tempering and tempering + cryogenic treatment with scanning electron microscopy, X-ray diffraction for residual stress measurements, and micro- and nano-indentation to determine Young’s modulus and plasticity factor measurement. Besides the improvement of toughness, a further aim of the present work is the investigation of the pertinence of a novel technique for characterizing the fracture toughness via scratch experiments on cryogenically-treated steels. Results show that the application of post-tempering cryogenic treatment on AISI M2, AISI D2, and X105CrCoMo18 steels induce precipitation of fine and homogeneously dispersed sub-micrometric carbides which do not alter hardness and Young’s modulus values, but reduce residual stresses and increase fracture toughness. Finally, scratch test proved to be an alternative simple technique to determine the fracture toughness of cryogenically treated steels.

  10. Mechanism of Secondary Hardening in Rapid Tempering of Dual-Phase Steel

    Science.gov (United States)

    Saha, Dulal Chandra; Nayak, Sashank S.; Biro, Elliot; Gerlich, Adrian P.; Zhou, Y.

    2014-12-01

    Dual-phase steel with ferrite-martensite-bainite microstructure exhibited secondary hardening in the subcritical heat affected zone during fiber laser welding. Rapid isothermal tempering conducted in a Gleeble simulator also indicated occurrence of secondary hardening at 773 K (500 °C), as confirmed by plotting the tempered hardness against the Holloman-Jaffe parameter. Isothermally tempered specimens were characterized by analytic transmission electron microscopy and high-angle annular dark-field imaging. The cementite (Fe3C) and TiC located in the bainite phase of DP steel decomposed upon rapid tempering to form needle-shaped Mo2C (aspect ratio ranging from 10 to 25) and plate-shaped M4C3 carbides giving rise to secondary hardening. Precipitation of these thermodynamically stable and coherent carbides promoted the hardening phenomenon. However, complex carbides were only seen in the tempered bainite and were not detected in the tempered martensite. The martensite phase decomposed into ferrite and spherical Fe3C, and interlath-retained austenite decomposed into ferrite and elongated carbide.

  11. Tempering of Mn and Mn-Si-V dual-phase steels

    Science.gov (United States)

    Speich, G. R.; Schwoeble, A. J.; Huffman, G. P.

    1983-06-01

    Changes in the yield behavior, strength, and ductility of a Mn and a Mn-Si-V d11Al-phase (ferrite-martensite) steel were investigated after tempering one hour at 200 to 600 °C. The change in yield behavior was complex in both steels with the yield strength first increasing and then decreasing as the tempering temperature was increased. This complex behavior is attributed to a combination of factors including carbon segregation to dislocations, a return of discontinuous yielding, and the relief of resid11Al stresses. In contrast, the tensile strength decreased continuously as the tempering temperature was increased in a manner that could be predicted from the change in hardness of the martensite phase using a simple composite strengthening model. The initial tensile ductility (total elongation) of the Mn-Si-V steel was much greater than that of the Mn steel. However, upon tempering up to 400 °C, the ductility of the Mn-Si-V decreased whereas that of the Mn steel increased. As a result, both steels had similar ductilities after tempering at 400 °C or higher temperatures. These results are attributed to the larger amounts of retained austenite in the Mn-Si-V steel (9 pct) compared to the Mn steel (3 pct) and its contribution to tensile ductility by transforming to martensite during plastic straining. Upon tempering at 400 °C, the retained austenite decomposes to bainite and its contribution to tensile ductility is eliminated.

  12. Fledgling survival increases with development time and adult survival across north and south temperate zones

    Science.gov (United States)

    Lloyd, Penn; Martin, Thomas E.

    2016-01-01

    Slow life histories are characterized by high adult survival and few offspring, which are thought to allow increased investment per offspring to increase juvenile survival. Consistent with this pattern, south temperate zone birds are commonly longer-lived and have fewer young than north temperate zone species. However, comparative analyses of juvenile survival, including during the first few weeks of the post-fledging period when most juvenile mortality occurs, are largely lacking. We combined our measurements of fledgling survival for eight passerines in South Africa with estimates from published studies of 57 north and south temperate zone songbird species to test three predictions: (1) fledgling survival increases with length of development time in the nest; (2) fledgling survival increases with adult survival and reduced brood size controlled for development time; and (3) south temperate zone species, with their higher adult survival and smaller brood sizes, exhibit higher fledgling survival than north temperate zone species controlled for development time. We found that fledgling survival was higher among south temperate zone species and generally increased with development time and adult survival within and between latitudinal regions. Clutch size did not explain additional variation, but was confounded with adult survival. Given the importance of age-specific mortality to life history evolution, understanding the causes of these geographical patterns of mortality is important.

  13. Impacts of extreme winter warming events on plant physiology in a sub-Arctic heath community.

    Science.gov (United States)

    Bokhorst, Stef; Bjerke, Jarle W; Davey, Matthew P; Taulavuori, Kari; Taulavuori, Erja; Laine, Kari; Callaghan, Terry V; Phoenix, Gareth K

    2010-10-01

    Insulation provided by snow cover and tolerance of freezing by physiological acclimation allows Arctic plants to survive cold winter temperatures. However, both the protection mechanisms may be lost with winter climate change, especially during extreme winter warming events where loss of snow cover from snow melt results in exposure of plants to warm temperatures and then returning extreme cold in the absence of insulating snow. These events cause considerable damage to Arctic plants, but physiological responses behind such damage remain unknown. Here, we report simulations of extreme winter warming events using infrared heating lamps and soil warming cables in a sub-Arctic heathland. During these events, we measured maximum quantum yield of photosystem II (PSII), photosynthesis, respiration, bud swelling and associated bud carbohydrate changes and lipid peroxidation to identify physiological responses during and after the winter warming events in three dwarf shrub species: Empetrum hermaphroditum, Vaccinium vitis-idaea and Vaccinium myrtillus. Winter warming increased maximum quantum yield of PSII, and photosynthesis was initiated for E. hermaphroditum and V. vitis-idaea. Bud swelling, bud carbohydrate decreases and lipid peroxidation were largest for E. hermaphroditum, whereas V. myrtillus and V. vitis-idaea showed no or less strong responses. Increased physiological activity and bud swelling suggest that sub-Arctic plants can initiate spring-like development in response to a short winter warming event. Lipid peroxidation suggests that plants experience increased winter stress. The observed differences between species in physiological responses are broadly consistent with interspecific differences in damage seen in previous studies, with E. hermaphroditum and V. myrtillus tending to be most sensitive. This suggests that initiation of spring-like development may be a major driver in the damage caused by winter warming events that are predicted to become more

  14. Soil phosphorus fractionation as a tool for monitoring dust phosphorus signature underneath a Blue Pine (Pinus wallichiana) canopy in a Temperate Forest

    Energy Technology Data Exchange (ETDEWEB)

    Shafqat, M.N.; Shahid, S.; Eqani, S.A.M.A.S.; Shah, S.H.; Waseem, A.

    2016-07-01

    Aim of the study: This study aims (i) to monitor the amount of dust deposition during dry season in the moist temperate forest; (ii) to study nature of P fractions in the dust samples falling on the trees in the region; (iii) to study soil P fractions as influenced by the processes of throughfall and stemflow of a Blue Pine (Pinus wallichiana) canopy and to finger print the contribution of dust towards P input in the temperate forest ecosystem. Area of study: The site used for the collection of soil samples was situated at an elevation of 6900 feet above sea levels (temperate forest in Himalaya region) in the Thandani area national forest located in the north west of Pakistan. Material and methods: For soil sampling and processing, three forest sites with three old tree plants per site were selected at approximately leveled plain for surface soil sampling. Two dust samples were collected and analyzed for different physicochemical properties along with different P fractions. First dust sample was collected from a site situated at an elevation of 4000 feet and second one was collected from an elevation of 6500 feet above sea levels. Modified Hedley procedure for the fractionation of P in the dust and soil samples were used. Main results: The input of dust was 43 and 20 kg ha-1 during drier months of the year (September-June) at lower and higher elevation sites respectively, and the dust from lower elevation site had relative more all P fractions than the other dust sample. However, HCl-Pi fraction was dominant in both samples. Both labile (water plus NaHCO3) and non-labile (NaOH plus HCl) inorganic P (Pi) fractions were significantly increased in the surface soil by both stemflow and throughfall compared to the open field soil. The buildup of NaOH and HCl-Pi pools in soils underneath the canopy might prove useful in fingerprinting the contribution of atmospheric dust towards P cycling in this temperate forest. Research highlights: The role of dust in the cycling of P

  15. Planting history and propagule pressure as predictors of invasions by woody species in a temperate region

    Czech Academy of Sciences Publication Activity Database

    Křivánek, Martin; Pyšek, Petr; Jarošík, Vojtěch

    2006-01-01

    Roč. 20, č. 5 (2006), s. 1487-1498 ISSN 0888-8892 R&D Projects: GA ČR GA206/03/1216 Institutional research plan: CEZ:AV0Z60050516 Keywords : alien forestry trees * invasive trees * naturalization Subject RIV: EF - Botanics Impact factor: 3.762, year: 2006

  16. Driving factors behind the eutrophication signal in understorey plant communities of deciduous temperate forests

    Czech Academy of Sciences Publication Activity Database

    Verheyen, E.; Baeten, L.; De Frenne, P.; Brnhardt-Römermann, M.; Brunet, J.; Cornelis, J.; Decocq, G.; Dierschke, H.; Eriksson, O.; Hédl, Radim; Heinken, T.; Hermy, M.; Hommel, P.; Kirby, K.; Naaf, T.; Peterken, G.; Petřík, Petr; Pfadenhauer, J.; Van Calster, H.; Walther, G.-R.; Wulf, M.; Verstraeten, G.

    2012-01-01

    Roč. 100, č. 2 (2012), s. 352-365 ISSN 0022-0477 R&D Projects: GA AV ČR IAA600050812 Institutional research plan: CEZ:AV0Z60050516 Keywords : Ellenberg indicator values * forest management * large herbivores Subject RIV: EF - Botanics Impact factor: 5.431, year: 2012

  17. Emission of climate relevant volatile organochlorines by plants occurring in temperate forests

    Czech Academy of Sciences Publication Activity Database

    Forczek, Sándor; Laturnus, F.; Doležalová, Jana; Holík, Josef; Wimmer, Zdeněk

    2015-01-01

    Roč. 61, č. 3 (2015), s. 103-108 ISSN 1214-1178 R&D Projects: GA ČR GA13-11101S Institutional support: RVO:61389030 Keywords : volatile chlorinated hydrocarbons * biodegradation * Sphagnum moss Subject RIV: EF - Botanics Impact factor: 1.039, year: 2015

  18. Distinct responses of soil respiration to experimental litter manipulation in temperate woodland and tropical forest.

    Science.gov (United States)

    Bréchet, Laëtitia M; Lopez-Sangil, Luis; George, Charles; Birkett, Ali J; Baxendale, Catherine; Castro Trujillo, Biancolini; Sayer, Emma J

    2018-04-01

    Global change is affecting primary productivity in forests worldwide, and this, in turn, will alter long-term carbon (C) sequestration in wooded ecosystems. On one hand, increased primary productivity, for example, in response to elevated atmospheric carbon dioxide (CO 2 ), can result in greater inputs of organic matter to the soil, which could increase C sequestration belowground. On other hand, many of the interactions between plants and microorganisms that determine soil C dynamics are poorly characterized, and additional inputs of plant material, such as leaf litter, can result in the mineralization of soil organic matter, and the release of soil C as CO 2 during so-called "priming effects". Until now, very few studies made direct comparison of changes in soil C dynamics in response to altered plant inputs in different wooded ecosystems. We addressed this with a cross-continental study with litter removal and addition treatments in a temperate woodland (Wytham Woods) and lowland tropical forest (Gigante forest) to compare the consequences of increased litterfall on soil respiration in two distinct wooded ecosystems. Mean soil respiration was almost twice as high at Gigante (5.0 μmol CO 2  m -2  s -1 ) than at Wytham (2.7 μmol CO 2  m -2  s -1 ) but surprisingly, litter manipulation treatments had a greater and more immediate effect on soil respiration at Wytham. We measured a 30% increase in soil respiration in response to litter addition treatments at Wytham, compared to a 10% increase at Gigante. Importantly, despite higher soil respiration rates at Gigante, priming effects were stronger and more consistent at Wytham. Our results suggest that in situ priming effects in wooded ecosystems track seasonality in litterfall and soil respiration but the amount of soil C released by priming is not proportional to rates of soil respiration. Instead, priming effects may be promoted by larger inputs of organic matter combined with slower turnover rates.

  19. Strong thermal acclimation of photosynthesis in tropical and temperate wet-forest tree species: the importance of altered Rubisco content.

    Science.gov (United States)

    Scafaro, Andrew P; Xiang, Shuang; Long, Benedict M; Bahar, Nur H A; Weerasinghe, Lasantha K; Creek, Danielle; Evans, John R; Reich, Peter B; Atkin, Owen K

    2017-07-01

    Understanding of the extent of acclimation of light-saturated net photosynthesis (A n ) to temperature (T), and associated underlying mechanisms, remains limited. This is a key knowledge gap given the importance of thermal acclimation for plant functioning, both under current and future higher temperatures, limiting the accuracy and realism of Earth system model (ESM) predictions. Given this, we analysed and modelled T-dependent changes in photosynthetic capacity in 10 wet-forest tree species: six from temperate forests and four from tropical forests. Temperate and tropical species were each acclimated to three daytime growth temperatures (T growth ): temperate - 15, 20 and 25 °C; tropical - 25, 30 and 35 °C. CO 2 response curves of A n were used to model maximal rates of RuBP (ribulose-1,5-bisphosphate) carboxylation (V cmax ) and electron transport (J max ) at each treatment's respective T growth and at a common measurement T (25 °C). SDS-PAGE gels were used to determine abundance of the CO 2 -fixing enzyme, Rubisco. Leaf chlorophyll, nitrogen (N) and mass per unit leaf area (LMA) were also determined. For all species and T growth , A n at current atmospheric CO 2 partial pressure was Rubisco-limited. Across all species, LMA decreased with increasing T growth . Similarly, area-based rates of V cmax at a measurement T of 25 °C (V cmax 25 ) linearly declined with increasing T growth , linked to a concomitant decline in total leaf protein per unit leaf area and Rubisco as a percentage of leaf N. The decline in Rubisco constrained V cmax and A n for leaves developed at higher T growth and resulted in poor predictions of photosynthesis by currently widely used models that do not account for T growth -mediated changes in Rubisco abundance that underpin the thermal acclimation response of photosynthesis in wet-forest tree species. A new model is proposed that accounts for the effect of T growth -mediated declines in V cmax 25 on A n , complementing current

  20. Effects of climate variability and functional changes on carbon cycling in a temperate deciduous forest

    Energy Technology Data Exchange (ETDEWEB)

    Wu, Jian

    2013-03-15

    Temperate forests are globally important carbon (C) stocks and sinks. A decadal (1997-2009) trend of increasing C uptake has been observed in an intensively studied temperate deciduous forest, Soroe (Zealand, Denmark). This gave the impetus to investigate the factors controlling the C cycling and the fundamental processes at work in this type of ecosystem. The major objectives of this study were to (1) evaluate to what extent and at what temporal scales, direct climatic variability and functional changes (e.g. changes in the structure or physiological properties) regulate the interannual variability (IAV) in the ecosystem C balance; (2) provide a synthesis of the ecosystem C budget at this site and (3) investigate whether terrestrial ecosystem models can dynamically simulate the trend of increasing C uptake. Data driven analysis, semi-empirical and process-based modelling experiments were performed in a series of studies in order to provide a complete assessment of the carbon storage and allocation within the ecosystem and clarify the mechanisms responsible for the observed variability and trend in the ecosystem C fluxes. Combining all independently estimated ecosystem carbon budget (ECB) datasets and other calculated ECB components based on mass balance equations, a synthesis of the carbon cycling was performed. The results showed that this temperature deciduous forest was moderately productive with both high rates of gross primary production and ecosystem respiration. Approximately 62% of the gross assimilated carbon was respired by the living plants, while 21% was contributed to the soil as litter production, the latter balancing the total heterotrophic respiration. The remaining 17% was either stored in the plants (mainly as aboveground biomass) or removed from the system as wood production. In general, the ECB component datasets were consistent after the cross-checking. This, together with their characterized uncertainties, can be used in model data fusion

  1. Biodiversity of Terrestrial Ecosystems in Tropical to Temperate Australia

    Directory of Open Access Journals (Sweden)

    Raymond L. Specht

    2012-01-01

    Full Text Available During the short period of annual foliage growth in evergreen plant communities, aerodynamic fluxes (frictional, thermal, evaporative in the atmosphere as it flows over and through a plant community determine the Foliage Projective Covers and leaf attributes in overstorey and understorey strata. The number of leaves produced on each vertical foliage shoot depends on available soil water and nutrients during this growth period. The area of all leaves exposed to solar radiation determines net photosynthetic fixation of the plant community throughout the year. In turn, the species richness (number of species per hectare of both plants and resident vertebrates is determined. The species richness of unicellular algae and small multicellular isopods in permanent freshwater lagoons in Northern Australia may possibly have been increased by radiation released from nearby uranium deposits. Evolution of new angiosperms probably occurred in refugia during periods of extreme drought. When favourable climates were restored, the vegetation expanded to result in high Gamma Biodiversity (number of plant species per region but with each major plant community having essentially the same species richness (number of plant species per hectare. The probable effects of pollution and Global Warming on biodiversity in Australian ecosystems, that experience seasonal drought, are discussed.

  2. Biochar decelerates soil organic nitrogen cycling but stimulates soil nitrification in a temperate arable field trial.

    Directory of Open Access Journals (Sweden)

    Judith Prommer

    Full Text Available Biochar production and subsequent soil incorporation could provide carbon farming solutions to global climate change and escalating food demand. There is evidence that biochar amendment causes fundamental changes in soil nutrient cycles, often resulting in marked increases in crop production, particularly in acidic and in infertile soils with low soil organic matter contents, although comparable outcomes in temperate soils are variable. We offer insight into the mechanisms underlying these findings by focusing attention on the soil nitrogen (N cycle, specifically on hitherto unmeasured processes of organic N cycling in arable soils. We here investigated the impacts of biochar addition on soil organic and inorganic N pools and on gross transformation rates of both pools in a biochar field trial on arable land (Chernozem in Traismauer, Lower Austria. We found that biochar increased total soil organic carbon but decreased the extractable organic C pool and soil nitrate. While gross rates of organic N transformation processes were reduced by 50-80%, gross N mineralization of organic N was not affected. In contrast, biochar promoted soil ammonia-oxidizer populations (bacterial and archaeal nitrifiers and accelerated gross nitrification rates more than two-fold. Our findings indicate a de-coupling of the soil organic and inorganic N cycles, with a build-up of organic N, and deceleration of inorganic N release from this pool. The results therefore suggest that addition of inorganic fertilizer-N in combination with biochar could compensate for the reduction in organic N mineralization, with plants and microbes drawing on fertilizer-N for growth, in turn fuelling the belowground build-up of organic N. We conclude that combined addition of biochar with fertilizer-N may increase soil organic N in turn enhancing soil carbon sequestration and thereby could play a fundamental role in future soil management strategies.

  3. Biochar decelerates soil organic nitrogen cycling but stimulates soil nitrification in a temperate arable field trial.

    Science.gov (United States)

    Prommer, Judith; Wanek, Wolfgang; Hofhansl, Florian; Trojan, Daniela; Offre, Pierre; Urich, Tim; Schleper, Christa; Sassmann, Stefan; Kitzler, Barbara; Soja, Gerhard; Hood-Nowotny, Rebecca Clare

    2014-01-01

    Biochar production and subsequent soil incorporation could provide carbon farming solutions to global climate change and escalating food demand. There is evidence that biochar amendment causes fundamental changes in soil nutrient cycles, often resulting in marked increases in crop production, particularly in acidic and in infertile soils with low soil organic matter contents, although comparable outcomes in temperate soils are variable. We offer insight into the mechanisms underlying these findings by focusing attention on the soil nitrogen (N) cycle, specifically on hitherto unmeasured processes of organic N cycling in arable soils. We here investigated the impacts of biochar addition on soil organic and inorganic N pools and on gross transformation rates of both pools in a biochar field trial on arable land (Chernozem) in Traismauer, Lower Austria. We found that biochar increased total soil organic carbon but decreased the extractable organic C pool and soil nitrate. While gross rates of organic N transformation processes were reduced by 50-80%, gross N mineralization of organic N was not affected. In contrast, biochar promoted soil ammonia-oxidizer populations (bacterial and archaeal nitrifiers) and accelerated gross nitrification rates more than two-fold. Our findings indicate a de-coupling of the soil organic and inorganic N cycles, with a build-up of organic N, and deceleration of inorganic N release from this pool. The results therefore suggest that addition of inorganic fertilizer-N in combination with biochar could compensate for the reduction in organic N mineralization, with plants and microbes drawing on fertilizer-N for growth, in turn fuelling the belowground build-up of organic N. We conclude that combined addition of biochar with fertilizer-N may increase soil organic N in turn enhancing soil carbon sequestration and thereby could play a fundamental role in future soil management strategies.

  4. Effect of biochar and liming on soil nitrous oxide emissions from a temperate maize cropping system

    Science.gov (United States)

    Hüppi, R.; Felber, R.; Neftel, A.; Six, J.; Leifeld, J.

    2015-12-01

    Biochar, a carbon-rich, porous pyrolysis product of organic residues may positively affect plant yield and can, owing to its inherent stability, promote soil carbon sequestration when amended to agricultural soils. Another possible effect of biochar is the reduction in emissions of nitrous oxide (N2O). A number of laboratory incubations have shown significantly reduced N2O emissions from soil when mixed with biochar. Emission measurements under field conditions however are more scarce and show weaker or no reductions, or even increases in N2O emissions. One of the hypothesised mechanisms for reduced N2O emissions from soil is owing to the increase in soil pH following the application of alkaline biochar. To test the effect of biochar on N2O emissions in a temperate maize cropping system, we set up a field trial with a 20t ha-1 biochar treatment, a limestone treatment adjusted to the same pH as the biochar treatment (pH 6.5), and a control treatment without any addition (pH 6.1). An automated static chamber system measured N2O emissions for each replicate plot (n = 3) every 3.6 h over the course of 8 months. The field was conventionally fertilised at a rate of 160 kg N ha-1 in three applications of 40, 80 and 40 kg N ha-1 as ammonium nitrate. Cumulative N2O emissions were 52 % smaller in the biochar compared to the control treatment. However, the effect of the treatments overall was not statistically significant (p = 0.27) because of the large variability in the data set. Limed soils emitted similar mean cumulative amounts of N2O as the control. There is no evidence that reduced N2O emissions with biochar relative to the control is solely caused by a higher soil pH.

  5. Biochar Decelerates Soil Organic Nitrogen Cycling but Stimulates Soil Nitrification in a Temperate Arable Field Trial

    Science.gov (United States)

    Prommer, Judith; Wanek, Wolfgang; Hofhansl, Florian; Trojan, Daniela; Offre, Pierre; Urich, Tim; Schleper, Christa; Sassmann, Stefan; Kitzler, Barbara; Soja, Gerhard; Hood-Nowotny, Rebecca Clare

    2014-01-01

    Biochar production and subsequent soil incorporation could provide carbon farming solutions to global climate change and escalating food demand. There is evidence that biochar amendment causes fundamental changes in soil nutrient cycles, often resulting in marked increases in crop production, particularly in acidic and in infertile soils with low soil organic matter contents, although comparable outcomes in temperate soils are variable. We offer insight into the mechanisms underlying these findings by focusing attention on the soil nitrogen (N) cycle, specifically on hitherto unmeasured processes of organic N cycling in arable soils. We here investigated the impacts of biochar addition on soil organic and inorganic N pools and on gross transformation rates of both pools in a biochar field trial on arable land (Chernozem) in Traismauer, Lower Austria. We found that biochar increased total soil organic carbon but decreased the extractable organic C pool and soil nitrate. While gross rates of organic N transformation processes were reduced by 50–80%, gross N mineralization of organic N was not affected. In contrast, biochar promoted soil ammonia-oxidizer populations (bacterial and archaeal nitrifiers) and accelerated gross nitrification rates more than two-fold. Our findings indicate a de-coupling of the soil organic and inorganic N cycles, with a build-up of organic N, and deceleration of inorganic N release from this pool. The results therefore suggest that addition of inorganic fertilizer-N in combination with biochar could compensate for the reduction in organic N mineralization, with plants and microbes drawing on fertilizer-N for growth, in turn fuelling the belowground build-up of organic N. We conclude that combined addition of biochar with fertilizer-N may increase soil organic N in turn enhancing soil carbon sequestration and thereby could play a fundamental role in future soil management strategies. PMID:24497947

  6. Impacts of climate change on paddy rice yield in a temperate climate.

    Science.gov (United States)

    Kim, Han-Yong; Ko, Jonghan; Kang, Suchel; Tenhunen, John

    2013-02-01

    The crop simulation model is a suitable tool for evaluating the potential impacts of climate change on crop production and on the environment. This study investigates the effects of climate change on paddy rice production in the temperate climate regions under the East Asian monsoon system using the CERES-Rice 4.0 crop simulation model. This model was first calibrated and validated for crop production under elevated CO2 and various temperature conditions. Data were obtained from experiments performed using a temperature gradient field chamber (TGFC) with a CO2 enrichment system installed at Chonnam National University in Gwangju, Korea in 2009 and 2010. Based on the empirical calibration and validation, the model was applied to deliver a simulated forecast of paddy rice production for the region, as well as for the other Japonica rice growing regions in East Asia, projecting for years 2050 and 2100. In these climate change projection simulations in Gwangju, Korea, the yield increases (+12.6 and + 22.0%) due to CO2 elevation were adjusted according to temperature increases showing variation dependent upon the cultivars, which resulted in significant yield decreases (-22.1% and -35.0%). The projected yields were determined to increase as latitude increases due to reduced temperature effects, showing the highest increase for any of the study locations (+24%) in Harbin, China. It appears that the potential negative impact on crop production may be mediated by appropriate cultivar selection and cultivation changes such as alteration of the planting date. Results reported in this study using the CERES-Rice 4.0 model demonstrate the promising potential for its further application in simulating the impacts of climate change on rice production from a local to a regional scale under the monsoon climate system. © 2012 Blackwell Publishing Ltd.

  7. Spatial and temporal soil moisture resource partitioning by trees and grasses in a temperate savanna, Arizona, USA.

    Science.gov (United States)

    Weltzin, Jake F; McPherson, Guy R

    1997-10-01

    Stable isotope analysis was used to determine sources of water used by coexisting trees and grasses in a temperate savanna dominated by Quercus emoryi Torr. We predicted that (1) tree seedlings and bunchgrasses utilize shallow sources of soil water, (2) mature savanna trees use deeper sources of water, and (3) trees switch from shallow to deep water sources within 1 year of germination. We found that Q. emoryi trees, saplings, and seedlings (about 2 months, 1 year, and 2 years old), and the dominant bunchgrass [Trachypogon montufari (H.B.K.) Nees.] utilized seasonally available moisture from different depths within the soil profile depending on size/age relationships. Sapling and mature Q. emoryi acquired water from >50 cm deep, 2-month-old seedlings utilized water from emoryi within extant stands of native grasses. The potential for subsequent interaction between Q. emoryi and native grasses was evidenced by similar patterns of soil water use by 1- and 2-year-old seedlings and grasses. Q. emoryi seedlings did not switch from shallow to deep sources of soil water within 2 years of germination: water use by these seedlings apparently becomes independent of water use by grasses after 2 years of age. Finally, older trees (saplings, mature trees) use water from deeper soil layers than grasses, which may facilitate the stable coexistence of mature trees and grasses. Potential shifts in the seasonality of precipitation may alter interactions between woody plants and grasses within temperate savannas characterized by bimodal precipitation regimes: reductions in summer precipitation or soil moisture may be particularly detrimental to warm-season grasses and seedlings of Q. emoryi.

  8. Combined Use of Airborne Lidar and DBInSAR Data to Estimate LAI in Temperate Mixed Forests

    Directory of Open Access Journals (Sweden)

    Ross F. Nelson

    2012-06-01

    Full Text Available The objective of this study was to determine whether leaf area index (LAI in temperate mixed forests is best estimated using multiple-return airborne laser scanning (lidar data or dual-band, single-pass interferometric synthetic aperture radar data (from GeoSAR alone, or both in combination. In situ measurements of LAI were made using the LiCor LAI-2000 Plant Canopy Analyzer on 61 plots (21 hardwood, 36 pine, 4 mixed pine hardwood; stand age ranging from 12-164 years; mean height ranging from 0.4 to 41.2 m in the Appomattox-Buckingham State Forest, Virginia, USA. Lidar distributional metrics were calculated for all returns and for ten one meter deep crown density slices (a new metric, five above and five below the mode of the vegetation returns for each plot. GeoSAR metrics were calculated from the X-band backscatter coefficients (four looks as well as both X- and P-band interferometric heights and magnitudes for each plot. Lidar metrics alone explained 69% of the variability in LAI, while GeoSAR metrics alone explained 52%. However, combining the lidar and GeoSAR metrics increased the R2 to 0.77 with a CV-RMSE of 0.42. This study indicates the clear potential for X-band backscatter and interferometric height (both now available from spaceborne sensors, when combined with small-footprint lidar data, to improve LAI estimation in temperate mixed forests.

  9. Tropical Agroecosystems: These habitats are misunderstood by the temperate zones, mismanaged by the tropics.

    Science.gov (United States)

    Janzen, D H

    1973-12-21

    I have listed some of the ways in which the lowland tropics are not such a warm and wonderful place for the farmer, some of the reasons why it may be unreasonable to expect him to cope with the problems, and some of the ways in which the temperate zones make his task more difficult. The tropics are very close to being a tragedy of the commons on a global scale (69, 103), and it is the temperate zone's shepherds and sheep who are among the greatest offenders (31). Given that the temperate zones have some limited amount of resources with which they are willing to repay the tropics, how can these resources best be spent? The first answer, without doubt, is education, and the incorporation of what is already known about the tropics into that education. Second should be the generation of secure psychological and physical resources for governments that show they are enthusiastic about the development of an SYTA. Third should be support of intensive research needed to generate the set of site-specific rules for specific, clearly identified SYTA's. The subject matter of youths' cultural programming is presumably determined by what they will need during the rest of their lives. A major component of this programming should be the teaching of the socioeconomic rules of a sustained-yield, nonexpanding economy, tuned to the concept of living within the carrying capacity of the country's or region's resources. Incorporating such a process into tropical school systems will cause a major upheaval, if for no other reason than that it will involve an evaluation of the country's resources, what standard of living is to be accepted by those living on them, and who is presently harvesting them. Of even greater impact, it will have to evaluate resources in terms of their ability to raise the standard of living by Y amount for X proportion of the people in the region, rather than in terms of their cash value on the world market. For such a change to be technologically successful, it will

  10. Anthropogenic impact on amorphous silica pools in temperate soils

    Directory of Open Access Journals (Sweden)

    W. Clymans

    2011-08-01

    Full Text Available Human land use changes perturb biogeochemical silica (Si cycling in terrestrial ecosystems. This directly affects Si mobilisation and Si storage and influences Si export from the continents, although the magnitude of the impact is unknown. A major reason for our lack of understanding is that very little information exists on how land use affects amorphous silica (ASi storage in soils. We have quantified and compared total alkali-extracted (PSia and easily soluble (PSie Si pools at four sites along a gradient of anthropogenic disturbance in southern Sweden. Land use clearly affects ASi pools and their distribution. Total PSia and PSie for a continuous forested site at Siggaboda Nature Reserve (66 900 ± 22 800 kg SiO2 ha−1 and 952 ± 16 kg SiO2 ha−1 are significantly higher than disturbed land use types from the Råshult Culture Reserve including arable land (28 800 ± 7200 kg SiO2 ha−1 and 239 ± 91 kg SiO2 ha−1, pasture sites (27 300 ± 5980 kg SiO2 ha−1 and 370 ± 129 kg SiO2 ha−1 and grazed forest (23 600 ± 6370 kg SiO2 ha−1 and 346 ± 123 kg SiO2 ha−1. Vertical PSia and PSie profiles show significant (p < 0.05 variation among the sites. These differences in size and distribution are interpreted as the long-term effect of reduced ASi replenishment, as well as changes in ecosystem specific pedogenic processes and increased mobilisation of the PSia in disturbed soils. We have also made a first, though rough, estimate of the magnitude of change in temperate continental ASi pools due to human disturbance. Assuming that our data are representative, we estimate that total ASi storage in soils has declined by ca. 10 % since the onset of agricultural development (3000 BCE

  11. Carbon density and distribution of six Chinese temperate forests

    Institute of Scientific and Technical Information of China (English)

    2010-01-01

    Quantifying forest carbon(C) storage and distribution is important for forest C cycling studies and terrestrial ecosystem modeling.Forest inventory and allometric approaches were used to measure C density and allocation in six representative temperate forests of similar stand age(42-59 years old) and growing under the same climate in northeastern China.The forests were an aspen-birch forest,a hardwood forest,a Korean pine plantation,a Dahurian larch plantation,a mixed deciduous forest,and a Mongolian oak forest.There were no significant differences in the C densities of ecosystem components(except for detritus) although the six forests had varying vegetation compositions and site conditions.However,the differences were significant when the C pools were normalized against stand basal area.The total ecosystem C density varied from 186.9 tC hm-2 to 349.2 tC hm-2 across the forests.The C densities of vegetation,detritus,and soil ranged from 86.3-122.7 tC hm-2,6.5-10.5 tC hm-2,and 93.7-220.1 tC hm-2,respectively,which accounted for 39.7% ± 7.1%(mean ± SD),3.3% ± 1.1%,and 57.0% ± 7.9% of the total C densities,respectively.The overstory C pool accounted for > 99% of the total vegetation C pool.The foliage biomass,small root(diameter < 5mm) biomass,root-shoot ratio,and small root to foliage biomass ratio varied from 2.08-4.72 tC hm-2,0.95-3.24 tC hm-2,22.0%-28.3%,and 34.5%-122.2%,respectively.The Korean pine plantation had the lowest foliage production efficiency(total biomass/foliage biomass:22.6 g g-1) among the six forests,while the Dahurian larch plantation had the highest small root production efficiency(total biomass/small root biomass:124.7 g g-1).The small root C density decreased with soil depth for all forests except for the Mongolian oak forest,in which the small roots tended to be vertically distributed downwards.The C density of coarse woody debris was significantly less in the two plantations than in the four naturally regenerated forests.The variability

  12. Community composition of root-associated fungi in a Quercus-dominated temperate forest: “codominance” of mycorrhizal and root-endophytic fungi

    Science.gov (United States)

    Toju, Hirokazu; Yamamoto, Satoshi; Sato, Hirotoshi; Tanabe, Akifumi S; Gilbert, Gregory S; Kadowaki, Kohmei

    2013-01-01

    In terrestrial ecosystems, plant roots are colonized by various clades of mycorrhizal and endophytic fungi. Focused on the root systems of an oak-dominated temperate forest in Japan, we used 454 pyrosequencing to explore how phylogenetically diverse fungi constitute an ecological community of multiple ecotypes. In total, 345 operational taxonomic units (OTUs) of fungi were found from 159 terminal-root samples from 12 plant species occurring in the forest. Due to the dominance of an oak species (Quercus serrata), diverse ectomycorrhizal clades such as Russula, Lactarius, Cortinarius, Tomentella, Amanita, Boletus, and Cenococcum were observed. Unexpectedly, the root-associated fungal community was dominated by root-endophytic ascomycetes in Helotiales, Chaetothyriales, and Rhytismatales. Overall, 55.3% of root samples were colonized by both the commonly observed ascomycetes and ectomycorrhizal fungi; 75.0% of the root samples of the dominant Q. serrata were so cocolonized. Overall, this study revealed that root-associated fungal communities of oak-dominated temperate forests were dominated not only by ectomycorrhizal fungi but also by diverse root endophytes and that potential ecological interactions between the two ecotypes may be important to understand the complex assembly processes of belowground fungal communities. PMID:23762515

  13. Are all temperate lakes eutrophying in a warmer world?

    Science.gov (United States)

    Paltsev, A.; Creed, I. F.

    2017-12-01

    Freshwater lakes are at risk of eutrophication due to climate change and intensification of human activities on the planet. In relatively undisturbed areas of the temperate forest biome, lakes are "sentinels" of the effects of rising temperatures. We hypothesise that rising temperatures are driving a shift from nutrient-poor oligotrophic states to nutrient-rich eutrophic states. To test this hypothesis, we examined a time series of satellite based chlorophyll-a (a proxy of algal biomass) of 12,000+ lakes over 30 years in the Canadian portion of the Laurentian Great Lakes basin. From the time series, non-stationary trends (detected by Mann-Kendall analysis) and stationary cycles (revealed through Morlet wavelet analysis) were removed, and the standard deviation (SD) of the remaining residuals was used as an indicator of lake stability. Four classes of lake stability were identified: (1) stable (SD is consistently low); (2) destabilizing (SD increases over time); (3) unstable (SD is consistently high); and (4) stabilizing lakes (SD decreases over time). Stable lakes were either oligotrophic or eutrophic indicating the presence of two stable states in the region. Destabilizing lakes were shifting from oligotrophic to lakes with a higher trophic status (indicating eutrophication), unstable lakes were mostly mesotrophic, and stabilizing lakes were shifting from eutrophic to the lakes with lower trophic status (indicating oligotrophication). In contrast to common expectations, while many lakes (2142) were shifting from oligotrophic to eutrophic states, more lakes (3199) were showing the opposite trend and shifting from eutrophic to oligotrophic states. This finding reveals a complexity of lake responses to rising temperatures and the need to improve understanding of why some lakes shift while others do not. Future work is focused on exploring the interactive effects of global, regional, and local drivers of lake trophic states.

  14. Production of fermentables and biomass by six temperate fuelcrops

    Energy Technology Data Exchange (ETDEWEB)

    Parrish, D.J.; Gammon, T.C.; Graves, B.

    1985-12-01

    Several potential fuelcrops have been studied individually, but relatively little work has been done to compare the various temperate species in side-by-side trials. The production has been examined of readily fermentable carbohydrates and biomass by six fuelcrop candidates: grain sorghum (Sorghum bicolor), Jerusalem articoke (Helianthus tuberosus), maize (Zea Mays), sugarbeet (Beta vulgaris), sweet potato (Ipomoea batatas) and sweet sorghum (Sorghum bicolor). A randomized complete block design with four replicates was employed at each of three locations that were somewhat diverse in soil type, elevation, growing season length, and 1980 rainfall distribution. Fermentables in the harvestable dry matter were determined colorimetrically following dilute acid plus enzymatic hydrolysis. Overall, sugarbeet was the most prolific producer of fermentables (7.4 Mg/ha); Jerusalem artichoke (5.8 Mg/ha), maize (4.8 Mg/ha) and sweet sorghum stems (5.8 Mg/ha) were statistically equivalent, while sweet potato (4.0 Mg/ha) and grain sorghum (3.8 Mg/ha) were less productive than the other candidates. The crops performed somewhat differently at each location, but the most striking site-specific differences were seen at the site with the coarsest textured soil and driest season. At that location, maize produced the least fermentables (0.6 Mg/ha). Biomass production generally reflected either the amount of time each species was actively growing or limiations to growth associated with drought. No general recommendations are made concerning a preferred temperature fuelcrop. Based on the studies, however, maize may not always be the fuelcrop of choice; others, especially sugarbeet and sweet sorghum (when harvested for grain also), may be superior to maize in productivity of fermentable substrates. 6 tabs., 13 refs.

  15. Analysis of ecological thresholds in a temperate forest undergoing dieback.

    Directory of Open Access Journals (Sweden)

    Philip Martin

    Full Text Available Positive feedbacks in drivers of degradation can cause threshold responses in natural ecosystems. Though threshold responses have received much attention in studies of aquatic ecosystems, they have been neglected in terrestrial systems, such as forests, where the long time-scales required for monitoring have impeded research. In this study we explored the role of positive feedbacks in a temperate forest that has been monitored for 50 years and is undergoing dieback, largely as a result of death of the canopy dominant species (Fagus sylvatica, beech. Statistical analyses showed strong non-linear losses in basal area for some plots, while others showed relatively gradual change. Beech seedling density was positively related to canopy openness, but a similar relationship was not observed for saplings, suggesting a feedback whereby mortality in areas with high canopy openness was elevated. We combined this observation with empirical data on size- and growth-mediated mortality of trees to produce an individual-based model of forest dynamics. We used this model to simulate changes in the structure of the forest over 100 years under scenarios with different juvenile and mature mortality probabilities, as well as a positive feedback between seedling and mature tree mortality. This model produced declines in forest basal area when critical juvenile and mature mortality probabilities were exceeded. Feedbacks in juvenile mortality caused a greater reduction in basal area relative to scenarios with no feedback. Non-linear, concave declines of basal area occurred only when mature tree mortality was 3-5 times higher than rates observed in the field. Our results indicate that the longevity of trees may help to buffer forests against environmental change and that the maintenance of old, large trees may aid the resilience of forest stands. In addition, our work suggests that dieback of forests may be avoidable providing pressures on mature and juvenile trees do

  16. Radiation budget changes with dry forest clearing in temperate Argentina.

    Science.gov (United States)

    Houspanossian, Javier; Nosetto, Marcelo; Jobbágy, Esteban G

    2013-04-01

    Land cover changes may affect climate and the energy balance of the Earth through their influence on the greenhouse gas composition of the atmosphere (biogeochemical effects) but also through shifts in the physical properties of the land surface (biophysical effects). We explored how the radiation budget changes following the replacement of temperate dry forests by crops in central semiarid Argentina and quantified the biophysical radiative forcing of this transformation. For this purpose, we computed the albedo and surface temperature for a 7-year period (2003-2009) from MODIS imagery at 70 paired sites occupied by native forests and crops and calculated the radiation budget at the tropopause and surface levels using a columnar radiation model parameterized with satellite data. Mean annual black-sky albedo and diurnal surface temperature were 50% and 2.5 °C higher in croplands than in dry forests. These contrasts increased the outgoing shortwave energy flux at the top of the atmosphere in croplands by a quarter (58.4 vs. 45.9 W m(-2) ) which, together with a slight increase in the outgoing longwave flux, yielded a net cooling of -14 W m(-2) . This biophysical cooling effect would be equivalent to a reduction in atmospheric CO2 of 22 Mg C ha(-1) , which involves approximately a quarter to a half of the typical carbon emissions that accompany deforestation in these ecosystems. We showed that the replacement of dry forests by crops in central Argentina has strong biophysical effects on the energy budget which could counterbalance the biogeochemical effects of deforestation. Underestimating or ignoring these biophysical consequences of land-use changes on climate will certainly curtail the effectiveness of many warming mitigation actions, particularly in semiarid regions where high radiation load and smaller active carbon pools would increase the relative importance of biophysical forcing. © 2012 Blackwell Publishing Ltd.

  17. Microstructural, mechanical and electrochemical behaviour of a 7017 Al–Zn–Mg alloy of different tempers

    Energy Technology Data Exchange (ETDEWEB)

    Rout, Prasanta Kumar, E-mail: prasantonnet55@yahoo.com; Ghosh, M.M.; Ghosh, K.S., E-mail: ksghosh2001@yahoo.co.uk

    2015-06-15

    The aim of the investigation is to assess the microstructural features and associated physical, mechanical and electrochemical properties of a 7017 Al–Zn–Mg alloy of various tempers. A 7017 Al–Zn–Mg alloy was subjected to different ageing schedules to produce under-(T4), peak-(T6), over-(T7) and highly over-aged tempers. Optical microscopy, hardness measurement, electrical conductivity measurement, tensile testing and SEM fractographs, differential scanning calorimetry (DSC), transmission electron microscopy (TEM), and electrochemical polarization studies have been used to characterize the alloy tempers. Hardness measurement and tensile testing showed the characteristic age hardening phenomenon of aluminium alloys. Optical and TEM micrographs have revealed the variation in size of matrix strengthening η′ (MgZn{sub 2}) and also the size and distribution of grain boundary η (MgZn{sub 2}) precipitate with ageing time. DSC thermograms exhibiting exothermic and endothermic peaks indicated the characteristic solid state reaction sequence of the 7017 alloy. Potentiodynamic polarization study of the 7017 alloy of various tempers in 3.5 wt.% NaCl solution at near neutral pH showed typical active metal dissolution behaviour, but at pH 12 an active–passive–transpassive transition behaviour has been observed. - Graphical abstract: TEM micrograph of the 7017 aluminium alloy of various tempers (a, b) under aged (T4), (c, d) peak aged (T6), (e, f) over aged (T7) and (g, h) highly over-aged. Display Omitted - Highlights: • 7017 Al-Zn-Mg alloy was subjected to different artificial ageing treatments. • Characterization of 7017 alloy tempers by hardness, tensile, DSC, TEM and electrochemical behaviour. • Structure-properties relationship of the 7017 Al-Zn-Mg alloy of various tempers.

  18. Cellular metabolic rate is influenced by life-history traits in tropical and temperate birds.

    Science.gov (United States)

    Jimenez, Ana Gabriela; Van Brocklyn, James; Wortman, Matthew; Williams, Joseph B

    2014-01-01

    In general, tropical birds have a "slow pace of life," lower rates of whole-animal metabolism and higher survival rates, than temperate species. A fundamental challenge facing physiological ecologists is the understanding of how variation in life-history at the whole-organism level might be linked to cellular function. Because tropical birds have lower rates of whole-animal metabolism, we hypothesized that cells from tropical species would also have lower rates of cellular metabolism than cells from temperate species of similar body size and common phylogenetic history. We cultured primary dermal fibroblasts from 17 tropical and 17 temperate phylogenetically-paired species of birds in a common nutritive and thermal environment and then examined basal, uncoupled, and non-mitochondrial cellular O2 consumption (OCR), proton leak, and anaerobic glycolysis (extracellular acidification rates [ECAR]), using an XF24 Seahorse Analyzer. We found that multiple measures of metabolism in cells from tropical birds were significantly lower than their temperate counterparts. Basal and uncoupled cellular metabolism were 29% and 35% lower in cells from tropical birds, respectively, a decrease closely aligned with differences in whole-animal metabolism between tropical and temperate birds. Proton leak was significantly lower in cells from tropical birds compared with cells from temperate birds. Our results offer compelling evidence that whole-animal metabolism is linked to cellular respiration as a function of an animal's life-history evolution. These findings are consistent with the idea that natural selection has uniquely fashioned cells of long-lived tropical bird species to have lower rates of metabolism than cells from shorter-lived temperate species.

  19. Cellular metabolic rate is influenced by life-history traits in tropical and temperate birds.

    Directory of Open Access Journals (Sweden)

    Ana Gabriela Jimenez

    Full Text Available In general, tropical birds have a "slow pace of life," lower rates of whole-animal metabolism and higher survival rates, than temperate species. A fundamental challenge facing physiological ecologists is the understanding of how variation in life-history at the whole-organism level might be linked to cellular function. Because tropical birds have lower rates of whole-animal metabolism, we hypothesized that cells from tropical species would also have lower rates of cellular metabolism than cells from temperate species of similar body size and common phylogenetic history. We cultured primary dermal fibroblasts from 17 tropical and 17 temperate phylogenetically-paired species of birds in a common nutritive and thermal environment and then examined basal, uncoupled, and non-mitochondrial cellular O2 consumption (OCR, proton leak, and anaerobic glycolysis (extracellular acidification rates [ECAR], using an XF24 Seahorse Analyzer. We found that multiple measures of metabolism in cells from tropical birds were significantly lower than their temperate counterparts. Basal and uncoupled cellular metabolism were 29% and 35% lower in cells from tropical birds, respectively, a decrease closely aligned with differences in whole-animal metabolism between tropical and temperate birds. Proton leak was significantly lower in cells from tropical birds compared with cells from temperate birds. Our results offer compelling evidence that whole-animal metabolism is linked to cellular respiration as a function of an animal's life-history evolution. These findings are consistent with the idea that natural selection has uniquely fashioned cells of long-lived tropical bird species to have lower rates of metabolism than cells from shorter-lived temperate species.

  20. Cellular Metabolic Rate Is Influenced by Life-History Traits in Tropical and Temperate Birds

    Science.gov (United States)

    Jimenez, Ana Gabriela; Van Brocklyn, James; Wortman, Matthew; Williams, Joseph B.

    2014-01-01

    In general, tropical birds have a “slow pace of life,” lower rates of whole-animal metabolism and higher survival rates, than temperate species. A fundamental challenge facing physiological ecologists is the understanding of how variation in life-history at the whole-organism level might be linked to cellular function. Because tropical birds have lower rates of whole-animal metabolism, we hypothesized that cells from tropical species would also have lower rates of cellular metabolism than cells from temperate species of similar body size and common phylogenetic history. We cultured primary dermal fibroblasts from 17 tropical and 17 temperate phylogenetically-paired species of birds in a common nutritive and thermal environment and then examined basal, uncoupled, and non-mitochondrial cellular O2 consumption (OCR), proton leak, and anaerobic glycolysis (extracellular acidification rates [ECAR]), using an XF24 Seahorse Analyzer. We found that multiple measures of metabolism in cells from tropical birds were significantly lower than their temperate counterparts. Basal and uncoupled cellular metabolism were 29% and 35% lower in cells from tropical birds, respectively, a decrease closely aligned with differences in whole-animal metabolism between tropical and temperate birds. Proton leak was significantly lower in cells from tropical birds compared with cells from temperate birds. Our results offer compelling evidence that whole-animal metabolism is linked to cellular respiration as a function of an animal’s life-history evolution. These findings are consistent with the idea that natural selection has uniquely fashioned cells of long-lived tropical bird species to have lower rates of metabolism than cells from shorter-lived temperate species. PMID:24498080

  1. Specialization to Extremely Low-Nutrient Soils Limits the Nutritional Adaptability of Plant Lineages.

    Science.gov (United States)

    Verboom, G Anthony; Stock, William D; Cramer, Michael D

    2017-06-01

    Specialization to extreme selective situations promotes the acquisition of traits whose coadaptive integration may compromise evolutionary flexibility and adaptability. We test this idea in the context of the foliar stoichiometry of plants native to the South African Cape. Whereas foliar concentrations of nitrogen, phosphorus (P), potassium (K), calcium, magnesium, and sodium showed strong phylogenetic signal, as did the foliar ratios of these nutrients to P, the same was not true of the corresponding soil values. In addition, although foliar traits were often related to soil values, the coefficients of determination were consistently low. These results identify foliar stoichiometry as having a strong genetic component, with variation in foliar nutrient concentrations, especially [P] and [K], being identified as potentially adaptive. Comparison of stoichiometric variation across 11 similarly aged clades revealed consistently low foliar nutrient concentrations in lineages showing specialization to extremely low-nutrient fynbos heathlands. These lineages also display lower rates of evolution of these traits as well as a reduced tendency for foliar [P] to track soil [P]. Reduced evolutionary lability and adaptability in the nutritional traits of fynbos-specialist lineages may explain the floristic distinctness of the fynbos flora and implies a reduced scope for edaphically driven ecological speciation.

  2. Diverse plant and animal genetic records from Holocene and Pleistocene sediments

    DEFF Research Database (Denmark)

    Willerslev, E.; Hansen, Anders J.; Binladen, J.

    2003-01-01

    Genetic analyses of permafrost and temperate sediments reveal that plant and animal DNA may be preserved for long periods, even in the absence of obvious macrofossils. In Siberia, five permafrost cores ranging from 400,000 to 10,000 years old contained at least 19 different plant taxa, including...... the oldest authenticated ancient DNA sequences known, and megafaunal sequences including mammoth, bison, and horse. The genetic data record a number of dramatic changes in the taxonomic diversity and composition of Beringian vegetation and fauna. Temperate cave sediments in New Zealand also yielded DNA...

  3. Detection of phytohormones in temperate forest fungi predicts consistent abscisic acid production and a common pathway for cytokinin biosynthesis.

    Science.gov (United States)

    Morrison, Erin N; Knowles, Sarah; Hayward, Allison; Thorn, R Greg; Saville, Barry J; Emery, R J N

    2015-01-01

    The phytohormones, abscisic acid and cytokinin, once were thought to be present uniquely in plants, but increasing evidence suggests that these hormones are present in a wide variety of organisms. Few studies have examined fungi for the presence of these "plant" hormones or addressed whether their levels differ based on the nutrition mode of the fungus. This study examined 20 temperate forest fungi of differing nutritional modes (ectomycorrhizal, wood-rotting, saprotrophic). Abscisic acid and cytokinin were present in all fungi sampled; this indicated that the sampled fungi have the capacity to synthesize these two classes of phytohormones. Of the 27 cytokinins analyzed by HPLC-ESI MS/MS, seven were present in all fungi sampled. This suggested the existence of a common cytokinin metabolic pathway in fungi that does not vary among different nutritional modes. Predictions regarding the source of isopentenyl, cis-zeatin and methylthiol CK production stemming from the tRNA degradation pathway among fungi are discussed. © 2015 by The Mycological Society of America.

  4. Partitioning the regional and local drivers of phylogenetic and functional diversity along temperate elevational gradients on an East Asian peninsula.

    Science.gov (United States)

    Chun, Jung-Hwa; Lee, Chang-Bae

    2018-02-12

    Species-centric approaches to biodiversity in ecological research are limited in their ability to reflect the evolutionary history and functional diversity of community assembly. Recently, the introduction of alternative facets of biodiversity, such as phylogenetic and functional diversity, has shed light on this problem and improved our understanding of the processes underlying biodiversity patterns. Here, we investigated the phylogenetic and functional diversity patterns of α, β and γ components in woody plant assemblages along regional and local elevational gradients in South Korea. Although the patterns of phylogenetic and functional diversity varied along regional and local elevational transects, the main drivers were partitioned into two categories: regional area or climate for phylogenetic diversity, depending on whether the transect was at a regional or local scale; and habitat heterogeneity for functional diversity, which was derived in elevational bands. Moreover, environmental distance was more important than was geographic distance for phylogenetic and functional β diversity between paired elevational bands. These results support the hypothesis that niche-based deterministic processes such as environmental filtering and competitive exclusion are fundamental in structuring woody plant assemblages along temperate elevational gradients regardless of scale (regional vs. local) in our study areas.

  5. Reference of Temperature and Time during tempering process for non-stoichiometric FTO films

    Science.gov (United States)

    Yang, J. K.; Liang, B.; Zhao, M. J.; Gao, Y.; Zhang, F. C.; Zhao, H. L.

    2015-10-01

    In order to enhance the mechanical strength of Low-E glass, Fluorine-doped tin oxide (FTO) films have to be tempered at high temperatures together with glass substrates. The effects of tempering temperature (600 °C ~ 720 °C) and time (150 s ~ 300 s) on the structural and electrical properties of FTO films were investigated. The results show all the films consist of non-stoichiometric, polycrystalline SnO2 without detectable amounts of fluoride. 700 °C and 260 s may be the critical tempering temperature and time, respectively. FTO films tempered at 700 °C for 260 s possesses the resistivity of 7.54 × 10-4 Ω•cm, the average transmittance in 400 ~ 800 nm of ~80%, and the calculated emissivity of 0.38. Hall mobility of FTO films tempered in this proper condition is mainly limited by the ionized impurity scattering. The value of [O]/[Sn] at the film surface is much higher than the stoichiometric value of 2.0 of pure crystalline SnO2.

  6. Effect of land use change on methane oxidation in temperate forest and grassland soils

    Energy Technology Data Exchange (ETDEWEB)

    Ojima, D.S.; Valentine, D.W.; Mosier, A.R.; Parton, W.J.; Schimel, D.S. (Colorado State University, Fort Collins, CO (USA). Natural Resources Ecology Lab.)

    Evidence is accumulating that land use changes and other human activity during the past 100 to 200 years have contributed to decreased CH[sub 4] oxidation in the soil. Increased N additions to temperate forest soils in the northeastern United States decreased CH[sub 4] uptake by 30 to 60%, and increased N fertilization and conversion to cropland in temperate grasslands decreased CH[sub 4] uptake by 30 to 75%. Using these data, a series of calculations were made to estimate the impact of land use and management changes which have altered soil, the CH[sub 4] sink in temperate forest and grassland ecosystems. As the atmospheric mixing ratio of CH[sub 4] has increased during the past 150 y, the temperate CH[sub 4] sink has risen from approximately 8 Tg y[sup -1] to 27 Tg y[sup -1], assuming no loss of land cover to cropland conversion. The net effect of intensive land cover changes and extensive chronic disturbance (i.e., increased atmospheric N deposition) to these ecosystems have resulted in about 30% reduction in the CH[sub 4] budget even more as atmospheric CH[sub 4] concentrations increase and as a result of further disturbance to other biomes. Without accounting for this approximately 20 Tg y[sup -1] temperate soil sink, the atmospheric CH[sub 4] concentration would be increasing about 1.5 times the current rate. 39 refs., 2 figs., 1 tab.

  7. Methane Ebullition in Temperate Hydropower Reservoirs and Implications for US Policy on Greenhouse Gas Emissions.

    Science.gov (United States)

    Miller, Benjamin L; Arntzen, Evan V; Goldman, Amy E; Richmond, Marshall C

    2017-10-01

    The United States is home to 2198 dams actively used for hydropower production. With the December 2015 consensus adoption of the United Nations Framework Convention on Climate Change Paris Agreement, it is important to accurately quantify anthropogenic greenhouse gas emissions. Methane ebullition, or methane bubbles originating from river or lake sediments, has been shown to account for nearly all methane emissions from tropical hydropower reservoirs to the atmosphere. However, distinct ebullitive methane fluxes have been studied in comparatively few temperate hydropower reservoirs globally. This study measures ebullitive and diffusive methane fluxes from two eastern Washington reservoirs, and synthesizes existing studies of methane ebullition in temperate, boreal, and tropical hydropower reservoirs. Ebullition comprises nearly all methane emissions (>97%) from this study's two eastern Washington hydropower reservoirs to the atmosphere. Summer methane ebullition from these reservoirs was higher than ebullition in six southeastern U.S. hydropower reservoirs, however it was similar to temperate reservoirs in other parts of the world. Our literature synthesis suggests that methane ebullition from temperate hydropower reservoirs can be seasonally elevated compared to tropical climates, however annual emissions are likely to be higher within tropical climates, emphasizing the possible range of methane ebullition fluxes and the need for the further study of temperate reservoirs. Possible future changes to the Intergovernmental Panel on Climate Change and UNFCCC guidelines for national greenhouse gas inventories highlights the need for accurate assessment of reservoir emissions.

  8. Methane Ebullition in Temperate Hydropower Reservoirs and Implications for US Policy on Greenhouse Gas Emissions

    Science.gov (United States)

    Miller, Benjamin L.; Arntzen, Evan V.; Goldman, Amy E.; Richmond, Marshall C.

    2017-10-01

    The United States is home to 2198 dams actively used for hydropower production. With the December 2015 consensus adoption of the United Nations Framework Convention on Climate Change Paris Agreement, it is important to accurately quantify anthropogenic greenhouse gas emissions. Methane ebullition, or methane bubbles originating from river or lake sediments, has been shown to account for nearly all methane emissions from tropical hydropower reservoirs to the atmosphere. However, distinct ebullitive methane fluxes have been studied in comparatively few temperate hydropower reservoirs globally. This study measures ebullitive and diffusive methane fluxes from two eastern Washington reservoirs, and synthesizes existing studies of methane ebullition in temperate, boreal, and tropical hydropower reservoirs. Ebullition comprises nearly all methane emissions (>97%) from this study's two eastern Washington hydropower reservoirs to the atmosphere. Summer methane ebullition from these reservoirs was higher than ebullition in six southeastern U.S. hydropower reservoirs, however it was similar to temperate reservoirs in other parts of the world. Our literature synthesis suggests that methane ebullition from temperate hydropower reservoirs can be seasonally elevated compared to tropical climates, however annual emissions are likely to be higher within tropical climates, emphasizing the possible range of methane ebullition fluxes and the need for the further study of temperate reservoirs. Possible future changes to the Intergovernmental Panel on Climate Change and UNFCCC guidelines for national greenhouse gas inventories highlights the need for accurate assessment of reservoir emissions.

  9. Appropriate welding conditions of temper bead weld repair for SQV2A pressure vessel steel

    International Nuclear Information System (INIS)

    Mizuno, R.; Matsuda, F.; Brziak, P.; Lomozik, M.

    2004-01-01

    Temper bead welding technique is one of the most important repair welding methods for large structures for which it is difficult to perform the specified post weld heat treatment. In this study, appropriate temper bead welding conditions to improve the characteristics of heat affected zone (HAZ) are studied using pressure vessel steel SQV2A corresponding to ASTM A533 Type B Class 1. Thermal/mechanical simulator is employed to give specimens welding thermal cycles from single to quadruple cycle. Charpy absorbed energy and hardness of simulated CGHAZ by first cycle were degraded as compared with base metal. Improvability of these degradations by subsequent cycles is discussed and appropriate temper bead thermal cycles are clarified. When the peak temperature lower than Ac1 and near Ac1 in the second thermal cycle is applied to CGAHZ by first thermal cycle, the characteristics of CGHAZ improve enough. When the other peak temperatures (that is, higher than Ac1) in the second thermal cycle are applied to the CGHAZ, third or more thermal cycle temper bead process should be applied to improve the properties. Appropriate weld condition ranges are selected based on the above results. The validity of the selected ranges is verified by the temper bead welding test. (orig.)

  10. The Kinetics of Phase Transformations During Tempering in Laser Melted High Chromium Cast Steel

    Science.gov (United States)

    Li, M. Y.; Wang, Y.; Han, B.

    2012-06-01

    The precipitation of secondary carbides in the laser melted high chromium cast steels during tempering at 300-650 °C for 2 h in air furnace was characterized and the present phases was identified, by using transmission electron microscopy. Laser melted high chromium cast steel consists of austenitic dendrites and interdendritic M23C6 carbides. The austenite has such a strong tempering stability that it remains unchanged at temperature below 400 °C and the secondary hardening phenomenon starts from 450 °C to the maximum value of 672 HV at 560 °C. After tempering at 450 °C fine M23C6 carbides precipitate from the supersaturated austenite preferentially. In addition, the dislocation lines and slip bands still exist inside the austenite. While tempering at temperature below 560 °C, the secondary hardening simultaneously results from the martensite phase transformation and the precipitation of carbides as well as dislocation strengthening within a refined microstructure. Moreover, the formation of the ferrite matrix and large quality of coarse lamellar M3C carbides when the samples were tempered at 650 °C contributes to the decrease of hardness.

  11. Experimental study and simulation of cyclic softening of tempered martensite ferritic steels

    International Nuclear Information System (INIS)

    Giroux, P.-F.

    2011-01-01

    The present work focuses on the high temperature mechanical behaviour of 9% Cr tempered martensite steels, considered as potential candidates for structural components in the next Generation IV nuclear power plants. Already used for energy production in fossil power plants, they are sensitive to softening during high-temperature cycling and creep-fatigue. This phenomenon is coupled to a pronounced microstructural degradation: mainly vanishing of subgrain boundaries and decrease in dislocation density. This study aims at (i) linking the macroscopic cyclic softening of 9% Cr steels and their microstructural evolution during cycling and (ii) proposing a physically-based modelling of deformation mechanisms in order to predict the macroscopic mechanical behaviour of these steels during cycling. Mechanical study includes uniaxial tensile and cyclic test at 550 C performed on a Grade 92 steel (9Cr-0,5Mo-1,8W-V-Nb). The effect of both strain amplitude and rate on mechanical behaviour is studied. Examination of tensile specimens suggests that the physical mechanism responsible for slight measured softening is mainly the necking phenomenon and the evolution of mean subgrain size, which increases by more than 15 % compared to the as-received state. The evolution of the macroscopic stress during cycling shows that cyclic softening is due to the decrease in kinematic stress. TEM observations highlights that the mean subgrain size increases by 60 to 100 % while the dislocation density decreases by more than 50 % during cycling, compared to the as-received state. A self-consistent homogenization model based on crystalline elasto-visco-plasticity and dislocation densities, predicting the mechanical behaviour of the material and its microstructural evolution during deformation is proposed. This model takes some of the main physical deformation mechanisms into account and only the two parameters of crystalline visco-plasticity should be adjusted (the effective activation energy and

  12. Vegetation Indices for Mapping Canopy Foliar Nitrogen in a Mixed Temperate Forest

    Directory of Open Access Journals (Sweden)

    Zhihui Wang

    2016-06-01

    Full Text Available Hyperspectral remote sensing serves as an effective tool for estimating foliar nitrogen using a variety of techniques. Vegetation indices (VIs are a simple means of retrieving foliar nitrogen. Despite their popularity, few studies have been conducted to examine the utility of VIs for mapping canopy foliar nitrogen in a mixed forest context. In this study, we assessed the performance of 32 vegetation indices derived from HySpex airborne hyperspectral images for estimating canopy mass-based foliar nitrogen concentration (%N in the Bavarian Forest National Park. The partial least squares regression (PLSR was performed for comparison. These vegetation indices were classified into three categories that are mostly correlated to nitrogen, chlorophyll, and structural properties such as leaf area index (LAI. %N was destructively measured in 26 broadleaf, needle leaf, and mixed stand plots to represent the different species and canopy structure. The canopy foliar %N is defined as the plot-level mean foliar %N of all species weighted by species canopy foliar mass fraction. Our results showed that the variance of canopy foliar %N is mainly explained by functional type and species composition. The normalized difference nitrogen index (NDNI produced the most accurate estimation of %N (R2CV = 0.79, RMSECV = 0.26. A comparable estimation of %N was obtained by the chlorophyll index Boochs2 (R2CV = 0.76, RMSECV = 0.27. In addition, the mean NIR reflectance (800–850 nm, representing canopy structural properties, also achieved a good accuracy in %N estimation (R2CV = 0.73, RMSECV = 0.30. The PLSR model provided a less accurate estimation of %N (R2CV = 0.69, RMSECV = 0.32. We argue that the good performance of all three categories of vegetation indices in %N estimation can be attributed to the synergy among plant traits (i.e., canopy structure, leaf chemical and optical properties while these traits may converge across plant species for evolutionary reasons. Our

  13. Field observations of climbing behavior and seed predation by adult ground beetles (Coleoptera: Carabidae) in a lowland area of the temperate zone.

    Science.gov (United States)

    Sasakawa, Kôji

    2010-10-01

    Granivory is a specialized food habit in the predominantly carnivorous beetle family Carabidae. Most studies of carabid granivory have been conducted under laboratory conditions; thus, our knowledge of the feeding ecology of granivorous carabids in the field is insufficient. I conducted field observations of climbing behavior and seed predation by adult carabids in a lowland area of eastern Japan, from early October to late November in 2008. This is the first systematic field observation of the feeding ecology of granivorous carabids in the temperate zone. In total, 176 carabid individuals of 11 species were observed, with 108 individuals feeding on plant seeds/flowers. Each carabid species was primarily observed feeding on a particular plant species. Frequently observed combinations were: Amara gigantea Motschulsky on Humulus scandens (Loureiro) Merrill (Moraceae) seed, Amara lucens Baliani on Artemisia indica Willdenow (Asteraceae) flower, and Amara macronota (Solsky) and Harpalus (Pseudoophonus) spp. on Digitaria ciliaris (Retzius) Koeler (Poaceae) seed. In all but one species, the sex ratio of individuals observed feeding was female-biased. In Am. gigantea and Am. macronota, a larger proportion of females than males ate seeds. In the three Amara species, copulations on plants, with the female feeding on its seeds/flowers, were often observed. These observations may indicate that, whereas females climb onto plants to feed on seeds, males climb to seek females for copulation rather than forage. Because granivorous carabids play important roles as weed-control agents in temperate agro-ecosystems, the present results would provide valuable basic information for future studies on this subject.

  14. Advanced remote sensing to quantify temperate peatland capacity for belowground carbon capture

    Science.gov (United States)

    Byrd, K. B.; Blanchard, S.; Schile, L. M.; Kolding, S.; Kelly, M.; Windham-Myers, L.; Miller, R.

    2011-12-01

    Temperate peatlands typically dominated by grasses and sedges generate among the greatest annual rates of net primary productivity (NPP, up to 4 kg C m-2) and soil carbon storage (up to 1 kg C m-2) for natural ecosystems. Belowground tissues represent 20-80% of total NPP, thus understanding the controls on belowground NPP (BNPP) in these wetland ecosystems is particularly important to quantifying peatland carbon balances. In addition, there is a growing need to quantify large-scale belowground C sequestration rates in wetlands to better understand marsh resilience to sea level rise and to help define eligibility for carbon offset credits. Since plant productivity influences wetland C budgets, combining field and remote sensing techniques for estimating above and belowground productivity of wetland vegetation over a large spatial extent will help to address these needs. We are working in a USGS long-term experimental wetland restoration site on drained peatland in the Sacramento-San Joaquin River Delta. Using the spatial variability in water depth and residence time across the 7 ha wetland, our goal is to develop practical methods to quantify and map BNPP of emergent marsh vegetation from remotely sensed estimates of aboveground plant characteristics and aboveground NPP. Field data collected on wetland plants hardstem bulrush (Schoenoplectus acutus) and cattail (Typha spp.) were coupled with reflectance data from a field spectrometer (range 350-2500 nm) every two to three weeks during the summer of 2011. We are analyzing reflectance data to develop hyperspectral indices that predict the biophysical characteristics of wetland vegetation - biomass, leaf area index (LAI), and the fraction of absorbed photosynthetically active radiation (fAPAR) - which may be used to infer belowground biomass and productivity. Soil cores and root in-growth bags were used to calculate root biomass and productivity rates. Existing allometric relationships were used to calculate

  15. Plant diversity predicts beta but not alpha diversity of soil microbes across grasslands worldwide

    Science.gov (United States)

    Prober, Suzanne M.; Leff, Jonathan W.; Bates, Scott T.; Borer, Elizabeth T.; Firn, Jennifer; Harpole, W. Stanley; Lind, Eric M.; Seabloom, Eric W.; Adler, Peter B.; Bakker, Jonathan D.; Cleland, Elsa E.; DeCrappeo, Nicole; DeLorenze, Elizabeth; Hagenah, Nicole; Hautier, Yann; Hofmockel, Kirsten S.; Kirkman, Kevin P.; Knops, Johannes M. H.; La Pierre, Kimberly J.; MacDougall, Andrew S.; McCulley, Rebecca L.; Mitchell, Charles E.; Risch, Anita C.; Schuetz, Martin; Stevens, Carly J.; Williams, Ryan J.; Fierer, Noah

    2015-01-01

    Aboveground–belowground interactions exert critical controls on the composition and function of terrestrial ecosystems, yet the fundamental relationships between plant diversity and soil microbial diversity remain elusive. Theory predicts predominantly positive associations but tests within single sites have shown variable relationships, and associations between plant and microbial diversity across broad spatial scales remain largely unexplored. We compared the diversity of plant, bacterial, archaeal and fungal communities in one hundred and forty-five 1 m2 plots across 25 temperate grassland sites from four continents. Across sites, the plant alpha diversity patterns were poorly related to those observed for any soil microbial group. However, plant beta diversity (compositional dissimilarity between sites) was significantly correlated with the beta diversity of bacterial and fungal communities, even after controlling for environmental factors. Thus, across a global range of temperate grasslands, plant diversity can predict patterns in the composition of soil microbial communities, but not patterns in alpha diversity.

  16. Bed-Deformation Experiments Beneath a Temperate Glacier

    Science.gov (United States)

    Iverson, N. R.; Hooyer, T. S.; Fischer, U. H.; Cohen, D.; Jackson, M.; Moore, P. L.; Lappegard, G.; Kohler, J.

    2002-12-01

    Fast flow of glaciers and genesis of glacial landforms are commonly attributed to shear deformation of subglacial sediment. Although models of this process abound, data gathered subglacially on the kinematics and mechanics of such deformation are difficult to interpret. Major difficulties stem from the necessity of either measuring deformation near glacier margins, where conditions may be abnormal, or at the bottoms of boreholes, where the scope of instrumentation is limited, drilling disturbs sediment, and local boundary conditions are poorly known. A different approach is possible at the Svartisen Subglacial Laboratory, where tunnels melted in the ice provide temporary human access to the bed of Engabreen, a temperate outlet glacier of the Svartisen Ice Cap in Norway. A trough (2 m x 1.5 m x 0.5 m deep) was blasted in the rock bed, where the glacier is 220 m thick and sliding at 0.1-0.2 m/d. During two spring field seasons, this trough was filled with 2.5 tons of simulated till. Instruments in the till recorded shear (tiltmeters), volume change, total normal stress, and pore-water pressure as ice moved across the till surface. Pore pressure was brought to near the total normal stress by feeding water to the base of the till with a high-pressure pump, operated in a rock tunnel 4 m below the bed surface. Results illustrate some fundamental aspects of bed deformation. Permanent shear deformation requires low effective normal stress and hence high pore-water pressure, owing to the frictional nature of till. Shear strain generally increases upward in the bed toward the glacier sole, consistent with previous measurements beneath thinner ice at glacier margins. At low effective normal stresses, ice sometimes decouples from underlying till. Overall, bed deformation accounts for 10-35 % of basal motion, although this range excludes shear in the uppermost 0.05 m of till where shear was not measured. Pump tests with durations ranging from seconds to hours highlight the need

  17. The effects of the insecticide lambda-Cyhalothrin on the earthworm Eisenia fetida under experimental conditions of tropical and temperate regions

    International Nuclear Information System (INIS)

    Garcia, Marcos; Scheffczyk, Adam; Garcia, Terezinha; Roembke, Joerg

    2011-01-01

    Plant Protection Products can affect soil organisms and thus might have negative impacts on soil functions. Little research has been performed on their impact on tropical soils. Therefore, the effects of the insecticide lambda-Cyhalothrin on earthworms were evaluated in acute and chronic laboratory tests modified for tropical conditions, i.e. at selected temperatures (20 and 28 o C) and with two strains (temperate and tropical) of the compost worm Eisenia fetida. The insecticide was spiked in two natural soils, in OECD artificial soil and a newly developed tropical artificial soil. The effects of lambda-Cyhalothrin did rarely vary in the same soil at tropical (LC50: 68.5-229 mg a.i./kg dry weight (DW); EC50: 54.2-60.2 mg a.i./kg DW) and temperate (LC50: 99.8-140 mg a.i./kg DW; EC50: 37.4-44.5 mg a.i./kg DW) temperatures. In tests with tropical soils and high temperature, effect values differed by up to a factor of ten. - Research highlights: → In one soil, effects of lambda-Cyhalothrin did not vary much at two temperatures. → In tropical soils at high temperature, effects differed by up to a factor of ten. → In the tropics, effects of pesticides can be higher or lower as in temperate regions. - The effects of the insecticide lambda-cyhalothrin on earthworms did not differ considerably when performed in the same soil under different temperatures, but LC/EC 50 values varied by a factor of ten between OECD and tropical artificial soil.

  18. The effects of the insecticide lambda-Cyhalothrin on the earthworm Eisenia fetida under experimental conditions of tropical and temperate regions

    Energy Technology Data Exchange (ETDEWEB)

    Garcia, Marcos [Embrapa Amazonia Ocidental, Rod. AM-10, Km 28, 69.011-970 Manaus, AM (Brazil); Scheffczyk, Adam [ECT Oekotoxikologie, Boettgerstr. 2-14, D-65439 Floersheim (Germany); Garcia, Terezinha [Embrapa Amazonia Ocidental, Rod. AM-10, Km 28, 69.011-970 Manaus, AM (Brazil); Roembke, Joerg, E-mail: j-roembke@ect.d [ECT Oekotoxikologie, Boettgerstr. 2-14, D-65439 Floersheim (Germany)

    2011-02-15

    Plant Protection Products can affect soil organisms and thus might have negative impacts on soil functions. Little research has been performed on their impact on tropical soils. Therefore, the effects of the insecticide lambda-Cyhalothrin on earthworms were evaluated in acute and chronic laboratory tests modified for tropical conditions, i.e. at selected temperatures (20 and 28 {sup o}C) and with two strains (temperate and tropical) of the compost worm Eisenia fetida. The insecticide was spiked in two natural soils, in OECD artificial soil and a newly developed tropical artificial soil. The effects of lambda-Cyhalothrin did rarely vary in the same soil at tropical (LC50: 68.5-229 mg a.i./kg dry weight (DW); EC50: 54.2-60.2 mg a.i./kg DW) and temperate (LC50: 99.8-140 mg a.i./kg DW; EC50: 37.4-44.5 mg a.i./kg DW) temperatures. In tests with tropical soils and high temperature, effect values differed by up to a factor of ten. - Research highlights: In one soil, effects of lambda-Cyhalothrin did not vary much at two temperatures. In tropical soils at high temperature, effects differed by up to a factor of ten. In the tropics, effects of pesticides can be higher or lower as in temperate regions. - The effects of the insecticide lambda-cyhalothrin on earthworms did not differ considerably when performed in the same soil under different temperatures, but LC/EC{sub 50} values varied by a factor of ten between OECD and tropical artificial soil.

  19. Survival rates of birds of tropical and temperate forests: will the dogma survive?

    Science.gov (United States)

    Karr, J.R.; Nichols, J.D.; Klimkiewicz, M.K.; Brawn, J.D.

    1990-01-01

    Survival rates of tropical forest birds are widely assumed to be high relative to the survival rates of temperate forest birds. Much life-history theory is based on this assumption despite the lack of empirical data to support it. We provide the first detailed comparison of survival rates of tropical and temperate forest birds based on extensive data bases and modern capture-recapture models. We find no support for the conventional wisdom. Because clutch size is only one component of reproductive rate, the frequently assumed, simple association between clutch size and adult survival rates should not necessarily be expected. Our results emphasize the need to consider components of fecundity in addition to clutch size when comparing the life histories of tropical and temperate birds and suggest similar considerations in the development of vertebrate life-history theory.

  20. Evaluation of temper embrittlement of martensitic and ferritic-martensitic steels by acoustic emission

    International Nuclear Information System (INIS)

    Lu, Yusho; Takahashi, Hideaki; Shoji, Tetsuo

    1987-01-01

    Martensitic (HT-9) and ferritic-martensitic steels (9Cr-2Mo) are considered as fusion first wall materials. In this investigation in order to understand the sensitivity of temper embrittlement in these steels under actual service condition, fracture toughness testing was made by use of acoustic emission technique. The temper embrittlement was characterized in terms of fracture toughness. The fracture toughness of these steels under 500 deg C, 100 hrs, and 1000 hrs heat treatment was decreased and their changes in micro-fracture process have been observed. The fracture toughness changes by temper embrittlement was discussed by the characteristic of AE, AE spectrum analysis and fractographic investigation. The relation between micro-fracture processes and AE has been clarified. (author)

  1. Quantifying the effects of tempering on individual phase properties of DP980 steel with nanoindentation

    Energy Technology Data Exchange (ETDEWEB)

    Cheng, G. [Physical and Computational Sciences Directorate, Pacific Northwest National Laboratory, P.O. Box 999, Richland, WA (United States); Zhang, F. [School of Mechanical and Materials Engineering, Washington State University, Pullman, WA (United States); Ruimi, A. [Department of Mechanical Engineering, Texas A& M University, Doha (Qatar); Field, D.P. [School of Mechanical and Materials Engineering, Washington State University, Pullman, WA (United States); Sun, X., E-mail: xin.sun@pnnl.gov [Physical and Computational Sciences Directorate, Pacific Northwest National Laboratory, P.O. Box 999, Richland, WA (United States)

    2016-06-14

    Tempering treatment is conducted on a commercial dual phase (DP) 980 steel at 250 °C and 400 °C for 60 min each. Ferrite and martensite grains are distinguished using electron backscatter diffraction (EBSD) and scanning probe microscopy (SPM), and the martensite volume fractions (MVF) are determined based on the image quality (IQ) map. Indentation tests combined with a newly developed inverse method are used to obtain the individual phase flow properties in each sample. The results show that, i) tempering significantly reduces martensite yield strength, while it slightly reduces the ferrite yield strength; ii) tempering temperature has a more significant influence on the work hardening exponent of ferrite than that of martensite. As a validation, a simple rule-of-mixtures is used to verify the above-predicted individual phase flow stresses with the experimentally obtained overall true stress vs. true strain curves.

  2. Testing the correlation of fragmented pollen records of the middle and late Pleistocene temperate stages

    DEFF Research Database (Denmark)

    Kuneš, Petr; Odgaard, Bent Vad

    Quaternary temperate stages have long been described based on changing pollen abundances of various tree taxa in lacustrine sediments. Later, attempts have been made to assign such biostratigraphic units to distinct marine isotope stages (MIS). Existing continuous chronosequences from Southern...... records depends on site-to-site correlations. This comparison has often been performed on a visual basis, lacking clearly defined protocols and statements of underlying assumptions. Here I test the correlation of well and poorly known pollen records of the middle- and late-Pleistocene temperate stages...... from Northern-Central Europe and evaluate the usefulness of several numerical techniques. TWINSPAN analysis identifies groups of temperate stages based on presence/absence of their indicative taxa and may be useful for distinguishing between older and younger interglacials. Site-to-site sequence...

  3. Electromagnetic nondestructive evaluation of tempering process in AISI D2 tool steel

    Science.gov (United States)

    Kahrobaee, Saeed; Kashefi, Mehrdad

    2015-05-01

    The present paper investigates the potential of using eddy current technique as a reliable nondestructive tool to detect microstructural changes during the different stages of tempering treatment in AISI D2 tool steel. Five stages occur in tempering of the steel: precipitation of ɛ carbides, formation of cementite, retained austenite decomposition, secondary hardening effect and spheroidization of carbides. These stages were characterized by destructive methods, including dilatometry, differential scanning calorimetry, X-ray diffraction, scanning electron microscopic observations, and hardness measurements. The microstructural changes alter the electrical resistivity/magnetic saturation, which, in turn, influence the eddy current signals. Two EC parameters, induced voltage sensed by pickup coil and impedance point detected by excitation coil, were evaluated as a function of tempering temperature to characterize the microstructural features, nondestructively. The study revealed that a good correlation exists between the EC parameters and the microstructural changes.

  4. Investigation of the self tempering effect of martensite by means of atom probe tomography

    International Nuclear Information System (INIS)

    Sackl, Stephanie; Clemens, Helmut; Primig, Sophie

    2015-01-01

    Self-tempering effects can be observed in steels with relatively high martensite start temperatures. After the formation of the first martensitic laths, carbon is able to diffuse in these laths during cooling, which can be attributed to sufficiently high temperatures. This effect cannot be observed in laths formed at lower temperatures. In steels containing up to 0.2 m.-% carbon, up to 90 % of the carbon atoms in the martensite segregate to dislocations during quenching. Due to its atomic resolution and sensitivity with respect to light elements, atom probe tomography is very well suited for the investigation of this phenomenon. In this study, the self-tempering effect in a quenched and tempered steel 42CrMo4 with a martensite start temperature of 310 C is investigated by means of atom probe tomography.

  5. Resonant behavior of the generalized Langevin system with tempered Mittag–Leffler memory kernel

    Science.gov (United States)

    Chen, Yao; Wang, Xudong; Deng, Weihua

    2018-05-01

    The generalized Langevin equation describes anomalous dynamics. Noise is not only the origin of uncertainty but also plays a positive role in helping to detect signals with information, termed stochastic resonance (SR). This paper analyzes the anomalous resonant behaviors of the generalized Langevin system with a multiplicative dichotomous noise and an internal tempered Mittag–Leffler noise. For a system with a fluctuating harmonic potential, we obtain the exact expressions of several types of SR such as the first moment, the amplitude and autocorrelation function for the output signal as well as the signal–noise ratio. We analyze the influence of the tempering parameter and memory exponent on the bona fide SR and the general SR. Moreover, it is detected that the critical memory exponent changes regularly with the increase of the tempering parameter. Almost all the theoretical results are validated by numerical simulations.

  6. Remaining stress-state and strain-energy in tempered glass fragments

    DEFF Research Database (Denmark)

    Nielsen, Jens Henrik

    2016-01-01

    to the fragmentation process and some authors e.g. Barsom (J Am Ceram Soc 51(2):75, 1968), Gulati (Glass processing days, Tamglass Engineering Oy, Tampere, 1997), Warren (Fractography of glasses and ceramics IV, Alfred University, Alfred, 2001) and Tandon and Glass (Fracture mechanics of ceramics—active materials......When tempered glass breaks, it shatters into relatively small pieces depending on the residual stress state in the glass. This has been known for centuries and is currently used in standards for classifying whether a piece of glass is tempered or not. However, the process of fragmentation...... is complex and only a few, relatively simple, models have been suggested for predicting the fragment size. The full theoretical explanation is still to be found and this work aims at providing another brick to the puzzle. The strain-energy present in tempered glass is obviously contributing...

  7. Tempering response to different morphologies of martensite in tensile deformation of dual-phase steel

    International Nuclear Information System (INIS)

    Ahmad, E.; Manzoor, T.; Sarwar, M.; Arif, M.; Hussain, N.

    2011-01-01

    A low alloy steel containing 0.2% C was heat treated with three cycles of heat treatments with the aim to acquire different morphologies of martensite in dual phase microstructure. Microscopic examination revealed that the morphologies consisting of grain boundary growth, scattered laths and bulk form of martensite were obtained. These morphologies have their distinct patterns of distribution in the matrix (ferrite). In tensile properties observations the dual phase steel with bulk morphology of martensite showed minimum of ductility but high tensile strength as compared to other two morphologies. This may be due to poor alignments of bulk martensite particles along tensile axes during deformation. Tempering was employed with various holding times at 550 deg. C to induce ductility in the heat treated material. The tempering progressively increased the ductility by increasing holding time. However, tempering response to strengths and ductilities was different to all three morphologies of martensite. (author)

  8. Localization and Ballistic Diffusion for the Tempered Fractional Brownian-Langevin Motion

    Science.gov (United States)

    Chen, Yao; Wang, Xudong; Deng, Weihua

    2017-10-01

    This paper discusses the tempered fractional Brownian motion (tfBm), its ergodicity, and the derivation of the corresponding Fokker-Planck equation. Then we introduce the generalized Langevin equation with the tempered fractional Gaussian noise for a free particle, called tempered fractional Langevin equation (tfLe). While the tfBm displays localization diffusion for the long time limit and for the short time its mean squared displacement (MSD) has the asymptotic form t^{2H}, we show that the asymptotic form of the MSD of the tfLe transits from t^2 (ballistic diffusion for short time) to t^{2-2H}, and then to t^2 (again ballistic diffusion for long time). On the other hand, the overdamped tfLe has the transition of the diffusion type from t^{2-2H} to t^2 (ballistic diffusion). The tfLe with harmonic potential is also considered.

  9. T6 and T78 tempers of AA6065 alloy: a quantitative TEM study

    Energy Technology Data Exchange (ETDEWEB)

    Tanaka, M.; Warner, T. [Compagnie de Produits Chimiques et Electrometallurgiques Pechiney, Centre de Recherches de Voreppe, 38 (France)

    2000-07-01

    Quantitative transmission electron microscopy is used to characterize the microstructure of an industrial Al-Mg-Si-Cu alloy (AA6056) in a state (T78 temper) desensitized to intergranular corrosion in comparison with the peak aged state (T6 temper). Analysis of dark-field images and of zero-loss filtered selected-area-diffraction patterns, along with EDX spectroscopy, indicates an advanced precipitation of the quaternary phase containing Si and Cu in the T78-tempered alloy compared to the T6 one. This supports the current theory of the desensitization of AA6056 alloy. However, the application of EDX analysis to provide direct evidence of the chemical composition variation in the solid solution across the grain boundary appears less satisfactory. (orig.)

  10. Plasmodium vivax populations revisited: mitochondrial genomes of temperate strains in Asia suggest ancient population expansion

    Directory of Open Access Journals (Sweden)

    Miao Miao

    2012-02-01

    Full Text Available Abstract Background Plasmodium vivax is the most widely distributed human malaria parasite outside of Africa, and its range extends well into the temperate zones. Previous studies provided evidence for vivax population differentiation, but temperate vivax parasites were not well represented in these analyses. Here we address this deficit by using complete mitochondrial (mt genome sequences to elucidate the broad genetic diversity and population structure of P. vivax from temperate regions in East and Southeast Asia. Results From the complete mtDNA sequences of 99 clinical samples collected in China, Myanmar and Korea, a total of 30 different haplotypes were identified from 26 polymorphic sites. Significant differentiation between different East and Southeast Asian parasite populations was observed except for the comparison between populations from Korea and southern China. Haplotype patterns and structure diversity analysis showed coexistence of two different groups in East Asia, which were genetically related to the Southeast Asian population and Myanmar population, respectively. The demographic history of P. vivax, examined using neutrality tests and mismatch distribution analyses, revealed population expansion events across the entire P. vivax range and the Myanmar population. Bayesian skyline analysis further supported the occurrence of ancient P. vivax population expansion. Conclusions This study provided further resolution of the population structure and evolution of P. vivax, especially in temperate/warm-temperate endemic areas of Asia. The results revealed divergence of the P. vivax populations in temperate regions of China and Korea from other populations. Multiple analyses confirmed ancient population expansion of this parasite. The extensive genetic diversity of the P. vivax populations is consistent with phenotypic plasticity of the parasites, which has implications for malaria control.

  11. Tempering Behavior of TiC-Reinforced SKD11 Steel Matrix Composite

    Science.gov (United States)

    Hwang, Ji-In; Kim, Seong Hoon; Heo, Yoon-Uk; Kim, Dae Ha; Hwang, Keum-Cheol; Suh, Dong-Woo

    2018-03-01

    TiC-reinforced SKD11 steel matrix composite, fabricated by a pressure infiltration casting, undergoes monotonic decrease in hardness as tempering temperature increases. Element mappings by TEM-EDS and thermodynamic calculation indicate that remarkable redistribution of V between the reinforcement and the steel matrix occurs by partial dissolution and re-precipitation of MC carbides upon casting process. The absence of secondary hardening is led by the enrichment of V in the reinforcement that reduces the V content in the steel matrix; this reduction in V content makes the precipitation of fine VC sluggish during the tempering.

  12. X-ray residual stress analysis on machined and tempered HPSN-ceramics

    Energy Technology Data Exchange (ETDEWEB)

    Immelmann, S.; Welle, E.; Reimers, W. [Hahn-Meitner-Institut Berlin GmbH (Germany)

    1997-11-15

    The residual stress state induced by grinding and tempering of hot pressed silicon nitride (HPSN) samples is studied by X-ray diffraction. The results reveal that the residual stress values at the surface of the samples as well as their gradient within the penetration depth of the X-rays depend on the sintering aid and thus, on the glassy phase content of the HPSN. Tempering of the ground HPSN reduces the residual stress values due to microplastic deformation, whereas an oxidation of the glassy phase leads to the formation of compressive residual stresses. (orig.) 35 refs.

  13. Use of the integration elements encoded by the temperate lactococcal bacteriophage TP901-1

    DEFF Research Database (Denmark)

    Brøndsted, Lone; Hammer, Karin

    1999-01-01

    Previously we showed that only one phage-expressed protein (Orf1), a 425-bp region upstream of the orf1 gene (presumably encoding a promoter), and the attP region are necessary and also sufficient for integration of the bacteriophage TP901-1 genome into the chromosome of Lactococcus lactis subsp......P region seem to be necessary for site-specific integration of the temperate bacteriophage TP901-1. By use of the integrative elements (attP and orf1) expressed by the temperate lactococcal bacteriophage TP901-1, a system for obtaining stable chromosomal single-copy transcriptional fusions in L. lactis...

  14. Effect of tempering on microstructure and tensile properties of niobium modified martensitic 9Cr heat resistant steel

    Energy Technology Data Exchange (ETDEWEB)

    Mandal, A., E-mail: anupmetal@gmail.com; Bandyopadhay, T.K.

    2015-01-03

    The effect of tempering on the microstructure of niobium modified 8.8 wt% chromium steel has been evaluated. Steel has been prepared using the conventional melting and casting route. Homogenization and forging is done at 1100 °C. Dilatometric study shows that the Ac{sub 1}, Ac{sub 3} and M{sub s} temperatures are 800, 855, and 131 °C, respectively. Initial cast and forged microstructures consist of martensite/ferrite. The samples are subsequently tempered at 500–800 °C for various intervals of time (1–5 h). The microstructure of the tempered sample is analyzed using optical microscopy, scanning electron microscopy, and X-ray diffraction. High Resolution Transmission Electron Microscopy (HRTEM) is used to identify the precipitate. Nanometer-sized precipitates (50–200 nm) are observed after tempering at 700 °C for 1 h. Niobium rich MC type carbide precipitates and chromium rich M{sub 23}C{sub 6} type precipitates are observed after tempering at 700 °C. Tensile strength decreases with increasing tempering temperature. Maximum tensile strength of 920 MPa is observed after tempering at 700 °C and maximum elongation of ∼11% is observed after tempering at 750 °C.

  15. Effect of tempering temperature on microstructure and sliding wear property of laser quenched 4Cr13 steel

    NARCIS (Netherlands)

    Ouyang, J.H.; Pei, Y.T.; Li, X.D.; Lei, T.C.

    1994-01-01

    4Cr13 martensite stainless steel was quenched by a CO2 laser and tempered for 2 h at different temperatures in the range 200 °C to 550 °C. The microstructure of treated layer was observed by SEM, XRD and TEM. Tempering leads to the decomposition of a large number of retained austenites in laser

  16. Molecular Investigation of the Short-term Sequestration of Natural Abundance 13C -labelled Cow Dung in the Surface Horizons of a Temperate Grassland Soil

    Science.gov (United States)

    Dungait, J.; Bol, R.; Evershed, R. P.

    2004-12-01

    An adequate understanding of the carbon (C) sequestration potential of grasslands requires that the quantity and residence times of C inputs be measured. Herbivore dung is largely comprised of plant cell wall material, a significant source of stable C in intensively grazed temperate grassland ecosystems that contributes to the soil carbon budget. Our work uses compound-specific isotope analysis to identify the pattern of input of dung-derived compounds from natural abundance 13C/-labelled cow dung into the surface horizons of a temperate grassland soil over one year. C4 dung (δ 13C \\-12.6 ‰ ) from maize fed cows was applied to a temperate grassland surface (δ 13C \\-29.95 ‰ ) at IGER-North Wyke (Devon, UK), and dung remains and soil cores beneath the treatments collected at ŧ = 7, 14, 28, 56, 112, 224 and 372 days. Bulk dung carbon present in the 0\\-1 cm and 1\\-5 cm surface horizons of a grassland soil over one year was estimated using Δ 13C between C4 dung and C3 dung, after Bol {\\et al.} (2000). The major biochemical components of dung were quantified using proximate forage fibre analyses, after Goering and Van Soest (1970) and identified using `wet' chemical and GC-MS methods. Plant cell wall polysaccharides and lignin were found to account for up to 67 {%} of dung dry matter. Hydrolysed polysaccharides were prepared as alditol acetates for analyses (after Docherty {\\et al.}, 2001), and a novel application of an off-line pyrolysis method applied to measure lignin-derived phenolic compounds (after Poole & van Bergen, 2002). This paper focuses on major events in the incorporation of dung carbon, estimated using natural abundance 13C&-slash;labelling technique. This revealed a major bulk input of dung carbon after a period of significant rainfall with a consequent decline in bulk soil δ 13C values until the end of the experiment (Dungait {\\et al.}, submitted). Findings will be presented revealing contribution of plant cell wall polysaccharides and

  17. Effect of quenching and tempering process on sulfide stress cracking susceptibility in API-5CT-C110 casing steel

    Energy Technology Data Exchange (ETDEWEB)

    Liu, M.; Wang, C.H.; Dai, Y.C.; Li, X.; Cao, G.H. [State Key Laboratory of Advanced Special Steel & Shanghai Key Laboratory of Advanced Ferrometallurgy & School of Materials Science and Engineering, Shanghai University, 149 Yanchang Road, Shanghai 200072 (China); Russell, A.M. [Division of Materials Science and Engineering, Ames Laboratory of the U.S.D.O.E., and Department of Materials Science and Engineering, Iowa State University, Ames, IA 50011-2300 (United States); Liu, Y.H.; Dong, X.M. [Tube & Pipe Department, Baosteel Research Institute, Baoshan Iron & Steel Co., Ltd, Shanghai 201900 (China); Zhang, Z.H., E-mail: zhzhang@baosteel.com [Tube & Pipe Department, Baosteel Research Institute, Baoshan Iron & Steel Co., Ltd, Shanghai 201900 (China)

    2017-03-14

    Three quenching and tempering processes performed on API-5CT-C110 casing steel produced tempered martensite structures and similar mechanical properties but distinct sulfide stress cracking (SSC) behavior as evaluated by Double Cantilever Beam (DCB) testing. An as-quenched specimen tempered at 690 °C for two hours showed superior SSC behavior compared to another specimen tempered at 715 °C for one hour. The latter contained a larger fraction of low-angle boundaries (LABs) and higher values of kernel average misorientation (KAM) than those in the former. Moreover, one more quenching and tempering on the former specimen would produce better SSC resistance with a decrease in the fraction of LABs and the values of KAM. Since dislocations trap hydrogen more strongly than grain boundaries, the specimen with higher KAM values, as well as higher dislocation density, would trap more hydrogen atoms and lead to greater SSC.

  18. Comparative Mapping of Seed Dormancy Loci Between Tropical and Temperate Ecotypes of Weedy Rice (Oryza sativa L.

    Directory of Open Access Journals (Sweden)

    Lihua Zhang

    2017-08-01

    Full Text Available Genotypic variation at multiple loci for seed dormancy (SD contributes to plant adaptation to diverse ecosystems. Weedy rice (Oryza sativa was used as a model to address the similarity of SD genes between distinct ecotypes. A total of 12 quantitative trait loci (QTL for SD were identified in one primary and two advanced backcross (BC populations derived from a temperate ecotype of weedy rice (34.3°N Lat.. Nine (75% of the 12 loci were mapped to the same positions as those identified from a tropical ecotype of weedy rice (7.1°N Lat.. The high similarity suggested that the majority of SD genes were conserved during the ecotype differentiation. These common loci are largely those collocated/linked with the awn, hull color, pericarp color, or plant height loci. Phenotypic correlations observed in the populations support the notion that indirect selections for the wild-type morphological characteristics, together with direct selections for germination time, were major factors influencing allelic distributions of SD genes across ecotypes. Indirect selections for crop-mimic traits (e.g., plant height and flowering time could also alter allelic frequencies for some SD genes in agroecosystems. In addition, 3 of the 12 loci were collocated with segregation distortion loci, indicating that some gametophyte development genes could also influence the genetic equilibria of SD loci in hybrid populations. The SD genes with a major effect on germination across ecotypes could be used as silencing targets to develop transgene mitigation (TM strategies to reduce the risk of gene flow from genetically modified crops into weed/wild relatives.

  19. Phytoextraction and phytostabilisation of metal-contaminated soil in temperate maritime climate of coastal British Columbia, Canada

    Science.gov (United States)

    Padmavathiamma, P. K.; Li, L. Y.

    2009-04-01

    This research addressed the phytoremediation of roadside soils subjected to multi-component metal solutions. A typical right of way for roads in Canada is around 30 m, and at least 33% of that land in the right of way is unpaved and can support animal life. Thus, land associated with 12,000 km of roads in the province of British Columbia and millions of kilometres around the world represent a substantial quantity of wildlife habitat where metal contamination needs to be remediated. Phytostabilisation, requires least maintenance among different phytoremediation techniques, and it could be a feasible and practical method of remediating in roadside soils along highways and for improving highway runoff drainage. The suitability of five plant species was studied for phytoextraction and phytostabilisation in a region with temperate maritime climate of coastal British Columbia, Canada. Pot experiments were conducted using Lolium perenne L (perennial rye grass), Festuca rubra L (creeping red fescue), Helianthus annuus L (sunflower), Poa pratensis L (Kentucky bluegrass) and Brassica napus L (rape) in soils treated with three different metal (Cu, Pb, Mn and Zn) concentrations. The bio-metric characters of plants in soils with multiple-metal contaminations, their metal accumulation characteristics, translocation properties and metal removal were assessed at different stages of plant growth, 90 and 120 DAS (days after sowing). Lolium was found to be suitable for the phytostabilisation of Cu and Pb, Festuca for Mn and Poa for Zn. Metal removal was higher at 120 than at 90 days after sowing, and metals concentrated more in the underground tissues with less translocation to the above-ground parts. Bioconcentration factors indicate that Festuca had the highest accumulation for Cu, Helianthus for Pb and Zn and Poa for Mn.

  20. The effects of the insecticide lambda-Cyhalothrin on the earthworm Eisenia fetida under experimental conditions of tropical and temperate regions.

    Science.gov (United States)

    Garcia, Marcos; Scheffczyk, Adam; Garcia, Terezinha; Römbke, Jörg

    2011-02-01

    Plant Protection Products can affect soil organisms and thus might have negative impacts on soil functions. Little research has been performed on their impact on tropical soils. Therefore, the effects of the insecticide lambda-Cyhalothrin on earthworms were evaluated in acute and chronic laboratory tests modified for tropical conditions, i.e. at selected temperatures (20 and 28°C) and with two strains (temperate and tropical) of the compost worm Eisenia fetida. The insecticide was spiked in two natural soils, in OECD artificial soil and a newly developed tropical artificial soil. The effects of lambda-Cyhalothrin did rarely vary in the same soil at tropical (LC50: 68.5-229 mg a.i./kg dry weight (DW); EC50: 54.2-60.2 mg a.i./kg DW) and temperate (LC50: 99.8-140 mg a.i./kg DW; EC50: 37.4-44.5 mg a.i./kg DW) temperatures. In tests with tropical soils and high temperature, effect values differed by up to a factor of ten. Copyright © 2010 Elsevier Ltd. All rights reserved.

  1. Species composition and forest structure explain the temperature sensitivity patterns of productivity in temperate forests

    Directory of Open Access Journals (Sweden)

    F. J. Bohn

    2018-03-01

    Full Text Available Rising temperatures due to climate change influence the wood production of forests. Observations show that some temperate forests increase their productivity, whereas others reduce their productivity. This study focuses on how species composition and forest structure properties influence the temperature sensitivity of aboveground wood production (AWP. It further investigates which forests will increase their productivity the most with rising temperatures. We described forest structure by leaf area index, forest height and tree height heterogeneity. Species composition was described by a functional diversity index (Rao's Q and a species distribution index (ΩAWP. ΩAWP quantified how well species are distributed over the different forest layers with regard to AWP. We analysed 370 170 forest stands generated with a forest gap model. These forest stands covered a wide range of possible forest types. For each stand, we estimated annual aboveground wood production and performed a climate sensitivity analysis based on 320 different climate time series (of 1-year length. The scenarios differed in mean annual temperature and annual temperature amplitude. Temperature sensitivity of wood production was quantified as the relative change in productivity resulting from a 1 °C rise in mean annual temperature or annual temperature amplitude. Increasing ΩAWP positively influenced both temperature sensitivity indices of forest, whereas forest height showed a bell-shaped relationship with both indices. Further, we found forests in each successional stage that are positively affected by temperature rise. For such forests, large ΩAWP values were important. In the case of young forests, low functional diversity and small tree height heterogeneity were associated with a positive effect of temperature on wood production. During later successional stages, higher species diversity and larger tree height heterogeneity were an advantage. To achieve such a

  2. Species composition and forest structure explain the temperature sensitivity patterns of productivity in temperate forests

    Science.gov (United States)

    Bohn, Friedrich J.; May, Felix; Huth, Andreas

    2018-03-01

    Rising temperatures due to climate change influence the wood production of forests. Observations show that some temperate forests increase their productivity, whereas others reduce their productivity. This study focuses on how species composition and forest structure properties influence the temperature sensitivity of aboveground wood production (AWP). It further investigates which forests will increase their productivity the most with rising temperatures. We described forest structure by leaf area index, forest height and tree height heterogeneity. Species composition was described by a functional diversity index (Rao's Q) and a species distribution index (ΩAWP). ΩAWP quantified how well species are distributed over the different forest layers with regard to AWP. We analysed 370 170 forest stands generated with a forest gap model. These forest stands covered a wide range of possible forest types. For each stand, we estimated annual aboveground wood production and performed a climate sensitivity analysis based on 320 different climate time series (of 1-year length). The scenarios differed in mean annual temperature and annual temperature amplitude. Temperature sensitivity of wood production was quantified as the relative change in productivity resulting from a 1 °C rise in mean annual temperature or annual temperature amplitude. Increasing ΩAWP positively influenced both temperature sensitivity indices of forest, whereas forest height showed a bell-shaped relationship with both indices. Further, we found forests in each successional stage that are positively affected by temperature rise. For such forests, large ΩAWP values were important. In the case of young forests, low functional diversity and small tree height heterogeneity were associated with a positive effect of temperature on wood production. During later successional stages, higher species diversity and larger tree height heterogeneity were an advantage. To achieve such a development, one could plant

  3. Elevated carbon dioxide and ozone alter productivity and ecosystem carbon content in northern temperate forests

    Science.gov (United States)

    Talhelm, Alan F; Pregitzer, Kurt S; Kubiske, Mark E; Zak, Donald R; Campany, Courtney E; Burton, Andrew J; Dickson, Richard E; Hendrey, George R; Isebrands, J G; Lewin, Keith F; Nagy, John; Karnosky, David F

    2014-01-01

    Three young northern temperate forest communities in the north-central United States were exposed to factorial combinations of elevated carbon dioxide (CO2) and tropospheric ozone (O3) for 11 years. Here, we report results from an extensive sampling of plant biomass and soil conducted at the conclusion of the experiment that enabled us to estimate ecosystem carbon (C) content and cumulative net primary productivity (NPP). Elevated CO2 enhanced ecosystem C content by 11%, whereas elevated O3 decreased ecosystem C content by 9%. There was little variation in treatment effects on C content across communities and no meaningful interactions between CO2 and O3. Treatment effects on ecosystem C content resulted primarily from changes in the near-surface mineral soil and tree C, particularly differences in woody tissues. Excluding the mineral soil, cumulative NPP was a strong predictor of ecosystem C content (r2 = 0.96). Elevated CO2 enhanced cumulative NPP by 39%, a consequence of a 28% increase in canopy nitrogen (N) content (g N m−2) and a 28% increase in N productivity (NPP/canopy N). In contrast, elevated O3 lowered NPP by 10% because of a 21% decrease in canopy N, but did not impact N productivity. Consequently, as the marginal impact of canopy N on NPP (ΔNPP/ΔN) decreased through time with further canopy development, the O3 effect on NPP dissipated. Within the mineral soil, there was less C in the top 0.1 m of soil under elevated O3 and less soil C from 0.1 to 0.2 m in depth under elevated CO2. Overall, these results suggest that elevated CO2 may create a sustained increase in NPP, whereas the long-term effect of elevated O3 on NPP will be smaller than expected. However, changes in soil C are not well-understood and limit our ability to predict changes in ecosystem C content. PMID:24604779

  4. Forest management type influences diversity and community composition of soil fungi across temperate forest ecosystems

    Directory of Open Access Journals (Sweden)

    Kezia eGoldmann

    2015-11-01

    Full Text Available Fungal communities have been shown to be highly sensitive towards shifts in plant diversity and species composition in forest ecosystems. However, little is known about the impact of forest management on fungal diversity and community composition of geographically separated sites. This study examined the effects of four different forest management types on soil fungal communities. These forest management types include age class forests of young managed beech (Fagus sylvatica L., with beech stands age of approximately 30 years, age class beech stands with an age of approximately 70 years, unmanaged beech stands, and coniferous stands dominated by either pine (Pinus sylvestris L. or spruce (Picea abies Karst. which are located in three study sites across Germany. Soil were sampled from 48 study plots and we employed fungal ITS rDNA pyrotag sequencing to assess the soil fungal diversity and community structure.We found that forest management type significantly affects the Shannon diversity of soil fungi and a significant interaction effect of study site and forest management on the fungal OTU richness. Consequently distinct fungal communities were detected in the three study sites and within the four forest management types, which were mainly related to the main tree species. Further analysis of the contribution of soil properties revealed that C/N ratio being the most important factor in all the three study sites whereas soil pH was significantly related to the fungal community in two study sites. Functional assignment of the fungal communities indicated that 38% of the observed communities were Ectomycorrhizal fungi (ECM and their distribution is significantly influenced by the forest management. Soil pH and C/N ratio were found to be the main drivers of the ECM fungal community composition. Additional fungal community similarity analysis revealed the presence of study site and management type specific ECM genera.This study extends our knowledge

  5. Forest Management Type Influences Diversity and Community Composition of Soil Fungi across Temperate Forest Ecosystems.

    Science.gov (United States)

    Goldmann, Kezia; Schöning, Ingo; Buscot, François; Wubet, Tesfaye

    2015-01-01

    Fungal communities have been shown to be highly sensitive toward shifts in plant diversity and species composition in forest ecosystems. However, little is known about the impact of forest management on fungal diversity and community composition of geographically separated sites. This study examined the effects of four different forest management types on soil fungal communities. These forest management types include age class forests of young managed beech (Fagus sylvatica L.), with beech stands age of approximately 30 years, age class beech stands with an age of approximately 70 years, unmanaged beech stands, and coniferous stands dominated by either pine (Pinus sylvestris L.) or spruce (Picea abies Karst.) which are located in three study sites across Germany. Soil were sampled from 48 study plots and we employed fungal ITS rDNA pyrotag sequencing to assess the soil fungal diversity and community structure. We found that forest management type significantly affects the Shannon diversity of soil fungi and a significant interaction effect of study site and forest management on the fungal operational taxonomic units richness. Consequently distinct fungal communities were detected in the three study sites and within the four forest management types, which were mainly related to the main tree species. Further analysis of the contribution of soil properties revealed that C/N ratio being the most important factor in all the three study sites whereas soil pH was significantly related to the fungal community in two study sites. Functional assignment of the fungal communities indicated that 38% of the observed communities were Ectomycorrhizal fungi (ECM) and their distribution is significantly influenced by the forest management. Soil pH and C/N ratio were found to be the main drivers of the ECM fungal community composition. Additional fungal community similarity analysis revealed the presence of study site and management type specific ECM genera. This study extends our

  6. Elevated carbon dioxide and ozone alter productivity and ecosystem carbon content in northern temperate forests.

    Science.gov (United States)

    Talhelm, Alan F; Pregitzer, Kurt S; Kubiske, Mark E; Zak, Donald R; Campany, Courtney E; Burton, Andrew J; Dickson, Richard E; Hendrey, George R; Isebrands, J G; Lewin, Keith F; Nagy, John; Karnosky, David F

    2014-08-01

    Three young northern temperate forest communities in the north-central United States were exposed to factorial combinations of elevated carbon dioxide (CO2 ) and tropospheric ozone (O3 ) for 11 years. Here, we report results from an extensive sampling of plant biomass and soil conducted at the conclusion of the experiment that enabled us to estimate ecosystem carbon (C) content and cumulative net primary productivity (NPP). Elevated CO2 enhanced ecosystem C content by 11%, whereas elevated O3 decreased ecosystem C content by 9%. There was little variation in treatment effects on C content across communities and no meaningful interactions between CO2 and O3 . Treatment effects on ecosystem C content resulted primarily from changes in the near-surface mineral soil and tree C, particularly differences in woody tissues. Excluding the mineral soil, cumulative NPP was a strong predictor of ecosystem C content (r(2) = 0.96). Elevated CO2 enhanced cumulative NPP by 39%, a consequence of a 28% increase in canopy nitrogen (N) content (g N m(-2) ) and a 28% increase in N productivity (NPP/canopy N). In contrast, elevated O3 lowered NPP by 10% because of a 21% decrease in canopy N, but did not impact N productivity. Consequently, as the marginal impact of canopy N on NPP (∆NPP/∆N) decreased through time with further canopy development, the O3 effect on NPP dissipated. Within the mineral soil, there was less C in the top 0.1 m of soil under elevated O3 and less soil C from 0.1 to 0.2 m in depth under elevated CO2 . Overall, these results suggest that elevated CO2 may create a sustained increase in NPP, whereas the long-term effect of elevated O3 on NPP will be smaller than expected. However, changes in soil C are not well-understood and limit our ability to predict changes in ecosystem C content. © 2014 The Authors Global Change Biology Published by John Wiley & Sons Ltd.

  7. Uncertainty analysis of a coupled ecosystem response model simulating greenhouse gas fluxes from a temperate grassland

    Science.gov (United States)

    Liebermann, Ralf; Kraft, Philipp; Houska, Tobias; Breuer, Lutz; Müller, Christoph; Kraus, David; Haas, Edwin; Klatt, Steffen

    2015-04-01

    Among anthropogenic greenhouse gas emissions, CO2 is the dominant driver of global climate change. Next to its direct impact on the radiation budget, it also affects the climate system by triggering feedback mechanisms in terrestrial ecosystems. Such mechanisms - like stimulated photosynthesis, increased root exudations and reduced stomatal transpiration - influence both the input and the turnover of carbon and nitrogen compounds in the soil. The stabilization and decomposition of these compounds determines how increasing CO2 concentrations change the terrestrial trace gas emissions, especially CO2, N2O and CH4. To assess the potential reaction of terrestrial greenhouse gas emissions to rising tropospheric CO2 concentration, we make use of a comprehensive ecosystem model integrating known processes and fluxes of the carbon-nitrogen cycle in soil, vegetation and water. We apply a state-of-the-art ecosystem model with measurements from a long term field experiment of CO2 enrichment. The model - a grassland realization of LandscapeDNDC - simulates soil chemistry coupled with plant physiology, microclimate and hydrology. The data - comprising biomass, greenhouse gas emissions, management practices and soil properties - has been attained from a FACE (Free Air Carbon dioxide Enrichment) experiment running since 1997 on a temperate grassland in Giessen, Germany. Management and soil data, together with weather records, are used to drive the model, while cut biomass as well as CO2 and N2O emissions are used for calibration and validation. Starting with control data from installations without CO2 enhancement, we begin with a GLUE (General Likelihood Uncertainty Estimation) assessment using Latin Hypercube to reduce the range of the model parameters. This is followed by a detailed sensitivity analysis, the application of DREAM-ZS for model calibration, and an estimation of the effect of input uncertainty on the simulation results. Since first results indicate problems with

  8. Plant functional traits in relation to fire in crown-fire ecosystems

    Science.gov (United States)

    Pausas, Juli G.; Bradstock, Ross A.; Keith, David A.; Keeley, Jon E.

    2004-01-01

    Disturbance is a dominant factor in many ecosystems, and the disturbance regime is likely to change over the next decades in response to land-use changes and global warming. We assume that predictions of vegetation dynamics can be made on the basis of a set of life-history traits that characterize the response of a species to disturbance. For crown-fire ecosystems, the main plant traits related to postfire persistence are the ability to resprout (persistence of individuals) and the ability to retain a persistent seed bank (persistence of populations). In this context, we asked (1) to what extent do different life-history traits co-occur with the ability to resprout and/or the ability to retain a persistent seed bank among differing ecosystems and (2) to what extent do combinations of fire-related traits (fire syndromes) change in a fire regime gradient? We explored these questions by reviewing the literature and analyzing databases compiled from different crown-fire ecosystems (mainly eastern Australia, California, and the Mediterranean basin). The review suggests that the pattern of correlation between the two basic postfire persistent traits and other plant traits varies between continents and ecosystems. From these results we predict, for instance, that not all resprouters respond in a similar way everywhere because the associated plant traits of resprouter species vary in different places. Thus, attempts to generalize predictions on the basis of the resprouting capacity may have limited power at a global scale. An example is presented for Australian heathlands. Considering the combination of persistence at individual (resprouting) and at population (seed bank) level, the predictive power at local scale was significantly increased.

  9. The northern flying squirrel as an indicator species of temperate rain forest: test of an hypothesis.

    Science.gov (United States)

    Winston P. Smith; Scott M. Gende; Jeffrey V. Nichols

    2005-01-01

    Management indicator species (MIS) often are selected because their life history and demographics are thought to reflect a suite of ecosystem conditions that are too difficult or costly to measure directly. The northern flying squirrel (Glaucomys sabrinus) has been proposed as an MIS of temperate rain forest of southeastern Alaska based on previous...

  10. Restoration of a boulder reef in temperate waters: Strategy, methodology and lessons learnt

    DEFF Research Database (Denmark)

    Støttrup, Josianne Gatt; Dahl, Karsten; Niemann, Sanne

    2017-01-01

    Anthropogenic impacts on marine habitats are a global problem, particularly in coastal areas. While boulder reefs in temperate waters hold high biomass and biodiversity, and may be unable to recover from anthropogenic stressors without restoration efforts, little is known about how to restore and...

  11. Characterization of Nanometric-Sized Carbides Formed During Tempering of Carbide-Steel Cermets

    Directory of Open Access Journals (Sweden)

    Matus K.

    2016-06-01

    Full Text Available The aim of this article of this paper is to present issues related to characterization of nanometric-sized carbides, nitrides and/or carbonitrides formed during tempering of carbide-steel cermets. Closer examination of those materials is important because of hardness growth of carbide-steel cermet after tempering. The results obtained during research show that the upswing of hardness is significantly higher than for high-speed steels. Another interesting fact is the displacement of secondary hardness effect observed for this material to a higher tempering temperature range. Determined influence of the atmosphere in the sintering process on precipitations formed during tempering of carbide-steel cermets. So far examination of carbidesteel cermet produced by powder injection moulding was carried out mainly in the scanning electron microscope. A proper description of nanosized particles is both important and difficult as achievements of nanoscience and nanotechnology confirm the significant influence of nanocrystalline particles on material properties even if its mass fraction is undetectable by standard methods. The following research studies have been carried out using transmission electron microscopy, mainly selected area electron diffraction and energy dispersive spectroscopy. The obtained results and computer simulations comparison were made.

  12. Modification of Low-Alloy Steel Surface by High-Temperature Gas Nitriding Plus Tempering

    Science.gov (United States)

    Jiao, Dongling; Li, Minsong; Ding, Hongzhen; Qiu, Wanqi; Luo, Chengping

    2018-02-01

    The low-alloy steel was nitrided in a pure NH3 gas atmosphere at 640 660 °C for 2 h, i.e., high-temperature gas nitriding (HTGN), followed by tempering at 225 °C, which can produce a high property surface coating without brittle compound (white) layer. The steel was also plasma nitriding for comparison. The composition, microstructure and microhardness of the nitrided and tempered specimens were examined, and their tribological behavior investigated. The results showed that the as-gas-nitrided layer consisted of a white layer composed of FeN0.095 phase (nitrided austenite) and a diffusional zone underneath the white layer. After tempering, the white layer was decomposed to a nano-sized (α-Fe + γ'-Fe4N + retained austenite) bainitic microstructure with a high hardness of 1150HV/25 g. Wear test results showed that the wear resistance and wear coefficient yielded by the complex HTGN plus tempering were considerably higher and lower, respectively, than those produced by the conventional plasma nitriding.

  13. Effect of microstructure on the susceptibility of a 533 steel to temper embrittlement

    International Nuclear Information System (INIS)

    Raoul, S.; Marini, B.; Pineau, A.

    1998-01-01

    In ferritic steels, brittle fracture usually occurs at low temperature by cleavage. However the segregation of impurities (P, As, Sn etc..) along prior γ grain boundaries can change the brittle fracture mode from transgranular to intergranular. In quenched and tempered steels, this segregation is associated with what is called the temper-embrittlement phenomenon. The main objective of the present study is to investigate the influence of the as-quenched microstructure (lower bainite or martensite) on the susceptibility of a low alloy steel (A533 cl.1) to temper-embrittlement. Dilatometric tests were performed to determine the continous-cooling-transformation (CCT) diagram of the material and to measure the critical cooling rate (V c ) for a martensitic quench. Then subsized Charpy V-notched specimens were given various cooling rates from the austenitization temperature to obtain a wide range of as-quenched microstructures, including martensite and bainite. These specimens were subsequently given a heat treatment to develop temper embrittlement and tested to measure the V-notch fracture toughness at -50 C. The fracture surfaces were examined by SEM. It is shown that martensitic microstructures are more susceptible to intergranular embrittlement than bainitic microstructures. These observed microstructural influences are briefly discussed. (orig.)

  14. Finite Element Implementation of a Glass Tempering Model in Three Dimensions

    DEFF Research Database (Denmark)

    Nielsen, Jens Henrik; Olesen, John Forbes; Poulsen, Peter Noe

    2010-01-01

    The present paper develops and validates a 3D model for the simulation of glass tempering. It is assembled from well-known models of temperature dependent viscoelasticity and structural relaxation and predicts both transient and steady-state stresses in complex 3D glass geometries. The theory and...

  15. Effect of microstructure on the susceptibility of a 533 steel to temper embrittlement

    Energy Technology Data Exchange (ETDEWEB)

    Raoul, S.; Marini, B. [CEA Centre d`Etudes Nucleaires de Saclay, 91 - Gif-sur-Yvette (France). Service de Recherches Metallurgiques Appliquees; Pineau, A. [CNRS, Evry (France). Centre de Materiaux

    1998-11-01

    In ferritic steels, brittle fracture usually occurs at low temperature by cleavage. However the segregation of impurities (P, As, Sn etc..) along prior {gamma} grain boundaries can change the brittle fracture mode from transgranular to intergranular. In quenched and tempered steels, this segregation is associated with what is called the temper-embrittlement phenomenon. The main objective of the present study is to investigate the influence of the as-quenched microstructure (lower bainite or martensite) on the susceptibility of a low alloy steel (A533 cl.1) to temper-embrittlement. Dilatometric tests were performed to determine the continous-cooling-transformation (CCT) diagram of the material and to measure the critical cooling rate (V{sub c}) for a martensitic quench. Then subsized Charpy V-notched specimens were given various cooling rates from the austenitization temperature to obtain a wide range of as-quenched microstructures, including martensite and bainite. These specimens were subsequently given a heat treatment to develop temper embrittlement and tested to measure the V-notch fracture toughness at -50 C. The fracture surfaces were examined by SEM. It is shown that martensitic microstructures are more susceptible to intergranular embrittlement than bainitic microstructures. These observed microstructural influences are briefly discussed. (orig.) 11 refs.

  16. Forest dynamics in the temperate rainforests of Alaska: from individual tree to regional scales

    Science.gov (United States)

    Tara M. Barrett

    2015-01-01

    Analysis of remeasurement data from 1079 Forest Inventory and Analysis (FIA) plots revealed multi-scale change occurring in the temperate rainforests of southeast Alaska. In the western half of the region, including Prince William Sound, aboveground live tree biomass and carbon are increasing at a rate of 8 ( ± 2 ) percent per decade, driven by an increase in Sitka...

  17. Ecological correlates of flying squirrel microhabitat use and density in temperate rainforests of southeastern Alaska

    Science.gov (United States)

    Winston P. Smith; Scott M. Gende; Jeffrey V. Nichols

    2004-01-01

    We studied habitat relations of the Prince of Wales flying squirrel (Glaucomys sabrinus griseifrons), an endemic of the temperate, coniferous rainforest of southeastern Alaska, because of concerns over population viability from extensive clear-cut logging in the region. We used stepwise logistic regression to examine relationships between...

  18. Preparation and characterization of tempered tungsten layers on single crystalline silicon

    International Nuclear Information System (INIS)

    Nitzsche, K.; Knedlik, C.; Tippmann, H.; Spiess, L.; Harman, R.; Vanek, O.; Tvarozek, V.

    1984-01-01

    Tungsten layers have been deposited on single crystalline silicon by sputtering and characterized by measurements of the sheet resistance by a linear four point method and the van der Pauw method. The influence of tempering under argon on the resistance has been studied. By means of the RBS spectroscopy it was found that the increase in the specific resistance is caused by interdiffusion

  19. The effect of prior tempering on cryogenic treatment to reduce retained austenite

    International Nuclear Information System (INIS)

    Stratton, Paul

    2010-01-01

    The consensus view is that a high carbon case gives gears the best overall properties provided that there is no carbide network and that the retained austenite has been reduced below 20% by cryogenic treatment. This view is effectively enshrined in the SAE AMS 2759/7 standard. The cryogenic treatment usually takes place immediately after the quench to avoid austenite stabilisation. However, for some parts with complex geometries that might crack during the treatment, a short low temperature temper is carried out first. Little is known on how this temper affects the subsequent cryogenic treatment. Three carburizing steels used extensively in the aerospace industry were carburized to produce high retained austenite levels in the case using two different, but typical carburizing cycles. The retained austenite was determined by XRD before and after cryogenic treatment carried out in accordance with the standard and compared with that obtained when an intermediate temper was used. This study shows that for three typical carburizing steels, carburized using typical cycles, the efficacy of the cryogenic treatment is reduced only slightly after the temper, and not enough to be industrially significant. (author)

  20. Testing for functional convergence of temperate rainforest tree assemblages in Chile and New Zealand

    NARCIS (Netherlands)

    Lusk, C.H.; Jimenez-Castillo, M.; Aragón, R.; Easdale, T.A.; Poorter, L.; Hinojosa, L.F.; Mason, N.W.H.W.H.

    2016-01-01

    An important tenet of biogeography and comparative ecology is that disjunct assemblages in similar physical environments are functionally more similar to each other than to assemblages from other environments. Temperate rainforests in South America, New Zealand and Australia share certain

  1. Griswold Tempered Water Flow Regulator Valves Used as Anti-Siphon Valves

    International Nuclear Information System (INIS)

    MISKA, C.

    2000-01-01

    FCV-1*22 and 1*23 are Griswold constant flow regulators used as anti-siphon valves in the tempered water system, they fail closed but valve cartridge orifice allows minimum flow to prevent loss of water from the MCO/CASK annulus

  2. Feed efficiency of tropically adapted cattle when fed in winter or spring in a temperate location

    Science.gov (United States)

    Earlier work has shown that young, tropically adapted cattle do not gain as rapidly as temperately adapted cattle during the winter in OK. The objective for this study was to determine if efficiency of gains was also impacted in tropically adapted cattle and if efficiency is consistent in different...

  3. Reinforced glass beams composed of annealed, heat-strengthened and fully tempered glass

    NARCIS (Netherlands)

    Louter, P.C.; Belis, J.L.I.F.; Bos, F.P.; Veer, F.A.

    Annealed, heat-strengthened and fully tempered SG-laminated reinforced glass beam specimens were subjected to four-point bending tests to investigate the effects of glass type on their structural response. During the test the beams showed linear elastic response until initial glass failure, followed

  4. Assessment of Soil Organic Carbon Stock of Temperate Coniferous Forests in Northern Kashmir

    Directory of Open Access Journals (Sweden)

    Davood A. Dar

    2015-02-01

    Full Text Available  Soil organic carbon (SOC estimation in temperate forests of the Himalaya is important to estimate their contribution to regional, national and global carbon stocks. Physico chemical properties of soil were quantified to assess soil organic carbon density (SOC and SOC CO2 mitigation density at two soil depths (0-10 and 10-20 cms under temperate forest in the Northern region of Kashmir Himalayas India. The results indicate that conductance, moisture content, organic carbon and organic matter were significantly higher while as pH and bulk density were lower at Gulmarg forest site. SOC % was ranging from 2.31± 0.96 at Gulmarg meadow site to 2.31 ± 0.26 in Gulmarg forest site. SOC stocks in these temperate forests were from 36.39 ±15.40 to 50.09 ± 15.51 Mg C ha-1. The present study reveals that natural vegetation is the main contributor of soil quality as it maintained the soil organic carbon stock. In addition, organic matter is an important indicator of soil quality and environmental parameters such as soil moisture and soil biological activity change soil carbon sequestration potential in temperate forest ecosystems.DOI: http://dx.doi.org/10.3126/ije.v4i1.12186International Journal of Environment Volume-4, Issue-1, Dec-Feb 2014/15; page: 161-178

  5. Thermal biology of the sub-polar-temperate estuarine crab Hemigrapsus crenulatus (Crustacea: Decapoda: Varunidae).

    Science.gov (United States)

    Cumillaf, Juan P; Blanc, Johnny; Paschke, Kurt; Gebauer, Paulina; Díaz, Fernando; Re, Denisse; Chimal, María E; Vásquez, Jorge; Rosas, Carlos

    2016-02-15

    Optimum temperatures can be measured through aerobic scope, preferred temperatures or growth. A complete thermal window, including optimum, transition (Pejus) and critical temperatures (CT), can be described if preferred temperatures and CT are defined. The crustacean Hemigrapsus crenulatus was used as a model species to evaluate the effect of acclimation temperature on: (i) thermal preference and width of thermal window, (ii) respiratory metabolism, and (iii) haemolymph proteins. Dependant on acclimation temperature, preferred temperature was between 11.8°C and 25.2°C while CT was found between a minimum of 2.7°C (CTmin) and a maximum of 35.9°C (CTmax). These data and data from tropical and temperate crustaceans were compared to examine the association between environmental temperature and thermal tolerance. Temperate species have a CTmax limit around 35°C that corresponded with the low CTmax limit of tropical species (34-36°C). Tropical species showed a CTmin limit around 9°C similar to the maximum CTmin of temperate species (5-6°C). The maximum CTmin of deep sea species that occur in cold environments (2.5°C) matched the low CTmin values (3.2°C) of temperate species. Results also indicate that the energy required to activate the enzyme complex (Ei) involved in respiratory metabolism of ectotherms changes along the latitudinal gradient of temperature. © 2016. Published by The Company of Biologists Ltd.

  6. Thermal biology of the sub-polar–temperate estuarine crab Hemigrapsus crenulatus (Crustacea: Decapoda: Varunidae)

    Science.gov (United States)

    Cumillaf, Juan P.; Blanc, Johnny; Paschke, Kurt; Gebauer, Paulina; Díaz, Fernando; Re, Denisse; Chimal, María E.; Vásquez, Jorge; Rosas, Carlos

    2016-01-01

    ABSTRACT Optimum temperatures can be measured through aerobic scope, preferred temperatures or growth. A complete thermal window, including optimum, transition (Pejus) and critical temperatures (CT), can be described if preferred temperatures and CT are defined. The crustacean Hemigrapsus crenulatus was used as a model species to evaluate the effect of acclimation temperature on: (i) thermal preference and width of thermal window, (ii) respiratory metabolism, and (iii) haemolymph proteins. Dependant on acclimation temperature, preferred temperature was between 11.8°C and 25.2°C while CT was found between a minimum of 2.7°C (CTmin) and a maximum of 35.9°C (CTmax). These data and data from tropical and temperate crustaceans were compared to examine the association between environmental temperature and thermal tolerance. Temperate species have a CTmax limit around 35°C that corresponded with the low CTmax limit of tropical species (34–36°C). Tropical species showed a CTmin limit around 9°C similar to the maximum CTmin of temperate species (5–6°C). The maximum CTmin of deep sea species that occur in cold environments (2.5°C) matched the low CTmin values (3.2°C) of temperate species. Results also indicate that the energy required to activate the enzyme complex (Ei) involved in respiratory metabolism of ectotherms changes along the latitudinal gradient of temperature. PMID:26879464

  7. Human impact on open temperate woodlands during the middle Holocene in Central Europe

    Czech Academy of Sciences Publication Activity Database

    Jamrichová, Eva; Hédl, Radim; Kolář, Jan; Tóth, P.; Bobek, Přemysl; Hajnalová, M.; Procházka, J.; Kadlec, Jaroslav; Szabó, Péter

    2017-01-01

    Roč. 245, OCT 2017 (2017), s. 55-68 ISSN 0034-6667 EU Projects: European Commission(XE) 278065 - LONGWOOD Institutional support: RVO:67985939 ; RVO:67985530 Keywords : temperate oakwoods * Quercus * human impact Subject RIV: EH - Ecology, Behaviour; DE - Earth Magnetism, Geodesy, Geography (GFU-E) OBOR OECD: Ecology; Physical geography (GFU-E) Impact factor: 1.817, year: 2016

  8. Detrital carbon pools in temperate forests: magnitude and potential for landscape-scale assessment

    Science.gov (United States)

    John Bradford; Peter Weishampel; Marie-Louise Smith; Randall Kolka; Richard A. Birdsey; Scott V. Ollinger; Michael G. Ryan

    2009-01-01

    Reliably estimating carbon storage and cycling in detrital biomass is an obstacle to carbon accounting. We examined carbon pools and fluxes in three small temperate forest landscapes to assess the magnitude of carbon stored in detrital biomass and determine whether detrital carbon storage is related to stand structural properties (leaf area, aboveground biomass,...

  9. Soil quality indicators of a mature alley-cropping agroforestry system in temperate North America

    Science.gov (United States)

    Although agroforestry practices are believed to improve soil quality, reports on long-term effects of alley cropping on soils within agroforestry in the temperate zone are limited. The objective of this study was to examine effects of management, landscape, and soil depth of an established agrofores...

  10. Soil quality differences in a mature alley cropping system in temperate North America

    Science.gov (United States)

    Alley cropping in agroforestry practices has been shown to improve soil quality, however information on long-term effects (>10 years) of alley cropping on soils in the temperate zone is very limited. The objective of this study was to examine effects of management, landscape, and soil depth on soil...

  11. Detrimental Influence of Invasive Earthworms on North American Cold-Temperate Forest Soils

    Science.gov (United States)

    Enerson, Isabel

    2012-01-01

    The topic of invasive earthworms is a timely concern that goes against many preconceived notions regarding the positive benefits of all worms. In the cold-temperate forests of North America invasive worms are threatening forest ecosystems, due to the changes they create in the soil, including decreases in C:N ratios and leaf litter, disruption of…

  12. Interannual, seasonal, and retrospective analysis of the methane and carbon dioxide budgets of a temperate peatland

    Science.gov (United States)

    D.M. Olson; T.J. Griffis; A. Noormets; R. Kolka; J. Chen

    2013-01-01

    Three years (2009-2011) of near-continuous methane (CH4) and carbon dioxide (CO2) fluxes were measured with the eddy covariance (EC) technique at a temperate peatland located within the Marcell Experimental Forest, in northern Minnesota, USA. The peatland was a net source of CH4 and a net sink of CO...

  13. Hardness prediction of HAZ in temper bead welding by non-consistent layer technique

    International Nuclear Information System (INIS)

    Yu, Lina; Saida, Kazuyoshi; Mochizuki, Masahito; Kameyama, Masashi; Chigusa, Naoki; Nishimoto, Kazutoshi

    2014-01-01

    Based on the experimentally obtained hardness database, the neural network-based hardness prediction system of heat affect zone (HAZ) in temper bead welding by Consistent Layer (CSL) technique has been constructed by the authors. However in practical operation, CSL technique is sometimes difficult to perform because of difficulty of the precise heat input controlling, and in such case non-CSL techniques are mainly used in the actual repair process. Therefore in the present study, the neural network-based hardness prediction system of HAZ in temper bead welding by non-CSL techniques has been constructed through thermal cycle simplification, from the view of engineering. The hardness distribution in HAZ with non-CSL techniques was calculated based on the thermal cycles numerically obtained by finite element method. The experimental result has shown that the predicted hardness is in good accordance with the measured ones. It follows that the currently proposed method is effective for estimating the tempering effect during temper bead welding by non-CSL techniques. (author)

  14. Electromagnetic nondestructive evaluation of tempering process in AISI D2 tool steel

    International Nuclear Information System (INIS)

    Kahrobaee, Saeed; Kashefi, Mehrdad

    2015-01-01

    The present paper investigates the potential of using eddy current technique as a reliable nondestructive tool to detect microstructural changes during the different stages of tempering treatment in AISI D2 tool steel. Five stages occur in tempering of the steel: precipitation of ε carbides, formation of cementite, retained austenite decomposition, secondary hardening effect and spheroidization of carbides. These stages were characterized by destructive methods, including dilatometry, differential scanning calorimetry, X-ray diffraction, scanning electron microscopic observations, and hardness measurements. The microstructural changes alter the electrical resistivity/magnetic saturation, which, in turn, influence the eddy current signals. Two EC parameters, induced voltage sensed by pickup coil and impedance point detected by excitation coil, were evaluated as a function of tempering temperature to characterize the microstructural features, nondestructively. The study revealed that a good correlation exists between the EC parameters and the microstructural changes. - Highlights: • D2 steel parts were tempered at 200-650 °C to produce various microstructures. • Precipitation of ε and Fe 3 C carbides and spheroidization of carbides were detected. • Retained austenite decomposition and secondary hardening effect were determined. • Variations of electrical resistivity (ρ) and magnetic saturation (Bs) were studied. • Combined effects of ρ and Bs on the EC outputs were evaluated

  15. Electromagnetic nondestructive evaluation of tempering process in AISI D2 tool steel

    Energy Technology Data Exchange (ETDEWEB)

    Kahrobaee, Saeed, E-mail: saeed.kahrobaee@yahoo.com; Kashefi, Mehrdad, E-mail: m-kashefi@um.ac.ir

    2015-05-15

    The present paper investigates the potential of using eddy current technique as a reliable nondestructive tool to detect microstructural changes during the different stages of tempering treatment in AISI D2 tool steel. Five stages occur in tempering of the steel: precipitation of ε carbides, formation of cementite, retained austenite decomposition, secondary hardening effect and spheroidization of carbides. These stages were characterized by destructive methods, including dilatometry, differential scanning calorimetry, X-ray diffraction, scanning electron microscopic observations, and hardness measurements. The microstructural changes alter the electrical resistivity/magnetic saturation, which, in turn, influence the eddy current signals. Two EC parameters, induced voltage sensed by pickup coil and impedance point detected by excitation coil, were evaluated as a function of tempering temperature to characterize the microstructural features, nondestructively. The study revealed that a good correlation exists between the EC parameters and the microstructural changes. - Highlights: • D2 steel parts were tempered at 200-650 °C to produce various microstructures. • Precipitation of ε and Fe{sub 3}C carbides and spheroidization of carbides were detected. • Retained austenite decomposition and secondary hardening effect were determined. • Variations of electrical resistivity (ρ) and magnetic saturation (Bs) were studied. • Combined effects of ρ and Bs on the EC outputs were evaluated.

  16. Evapotranspiration and soil moisture dynamics in a temperate grassland ecosystem in Inner Mongolia China

    Science.gov (United States)

    L. Hao; Ge Sun; Yongqiang Liu; G. S. Zhou; J. H.   Wan;  L. B. Zhang; J. L. Niu; Y. H. Sang;  J. J He

    2015-01-01

    Precipitation, evapotranspiration (ET), and soil moisture are the key controls for the productivity and functioning of temperate grassland ecosystems in Inner Mongolia, northern China. Quantifying the soil moisture dynamics and water balances in the grasslands is essential to sustainable grassland management under global climate change. We...

  17. Cold in the common garden: comparative low-temperature tolerance of boreal and temperate conifer foliage

    Science.gov (United States)

    G. Richard Strimbeck; Trygve D. Kjellsen; Paul G. Schaberg; Paula F. Murakami

    2007-01-01

    Because they maintain green foliage throughout the winter season, evergreen conifers may face special physiological challenges in a warming world. We assessed the midwinter low-temperature (LT) tolerance of foliage from eight temperate and boreal species in each of the genera Abies, Picea, and Pinus growing in an arboretum in...

  18. Effects of seasonal variation of photosynthetic capacity on the carbon fluxes of a temperate deciduous forest

    Science.gov (United States)

    David Medvigy; Su-Jong Jeong; Kenneth L. Clark; Nicholas S. Skowronski; Karina V. R. Schäfer

    2013-01-01

    Seasonal variation in photosynthetic capacity is an important part of the overall seasonal variability of temperate deciduous forests. However, it has only recently been introduced in a few terrestrial biosphere models, and many models still do not include it. The biases that result from this omission are not well understood. In this study, we use the Ecosystem...

  19. Thermal biology of the sub-polar–temperate estuarine crab Hemigrapsus crenulatus (Crustacea: Decapoda: Varunidae

    Directory of Open Access Journals (Sweden)

    Juan P. Cumillaf

    2016-03-01

    Full Text Available Optimum temperatures can be measured through aerobic scope, preferred temperatures or growth. A complete thermal window, including optimum, transition (Pejus and critical temperatures (CT, can be described if preferred temperatures and CT are defined. The crustacean Hemigrapsus crenulatus was used as a model species to evaluate the effect of acclimation temperature on: (i thermal preference and width of thermal window, (ii respiratory metabolism, and (iii haemolymph proteins. Dependant on acclimation temperature, preferred temperature was between 11.8°C and 25.2°C while CT was found between a minimum of 2.7°C (CTmin and a maximum of 35.9°C (CTmax. These data and data from tropical and temperate crustaceans were compared to examine the association between environmental temperature and thermal tolerance. Temperate species have a CTmax limit around 35°C that corresponded with the low CTmax limit of tropical species (34–36°C. Tropical species showed a CTmin limit around 9°C similar to the maximum CTmin of temperate species (5–6°C. The maximum CTmin of deep sea species that occur in cold environments (2.5°C matched the low CTmin values (3.2°C of temperate species. Results also indicate that the energy required to activate the enzyme complex (Ei involved in respiratory metabolism of ectotherms changes along the latitudinal gradient of temperature.

  20. Temperature dependence of UV radiation effects in Arctic and temperate isolates of three red macrophytes

    NARCIS (Netherlands)

    van de Poll, W.H.; Eggert, A.; Buma, A.G.J.; Breeman, Arno

    The temperature dependence of UV effects was studied for Arctic and temperate isolates of the red macrophytes Palmaria palmata, Coccotylus truncatus and Phycodrys rubens. The effects of daily repeated artificial ultraviolet B and A radiation (UVBR: 280-320 nm, UVAR: 320-400 nm) treatments were

  1. Strain-tempering of low carbon martensite steel wire by rapid heating

    International Nuclear Information System (INIS)

    Torisaka, Yasunori; Kihara, Junji

    1978-01-01

    In the production of prestressed concrete steel wires, a series of the cold drawing-patenting process are performed to improve the strength. In order to reduce cyclic process, the low carbon martensite steel wire which can be produced only by the process of hot rolling and direct quench has been investigated as strain-tempering material. When strain-tempering is performed on the low carbon martensite steel wire, stress relaxation (Re%) increases and mechanical properties such as total elongation, reduction of area, ultimate tensile strength and proof stress decrease remarkably by annealing. In order to shorten the heating time, the authors performed on the steel wire the strain-tempering with a heating time of 1.0 s using direct electrical resistance heating and examined the effects of rapid heating on the stress relaxation and the mechanical properties. Stress relaxation decreases without impairment of the mechanical properties up to a strain-tempering temperature of 573 K. Re(%) after 10.8 ks is 0% at the testing temperature 301 K, 0.49% at 363 K and 1.39% at 433 K. (auth.)

  2. On the Nature of Internal Interfaces in Tempered Martensite Ferritic Steels

    Czech Academy of Sciences Publication Activity Database

    Dronhofer, A.; Pešička, J.; Dlouhý, Antonín; Eggeler, G.

    2003-01-01

    Roč. 94, č. 5 (2003), s. 511-520 ISSN 0044-3093 R&D Projects: GA ČR GA106/99/1172 Institutional research plan: CEZ:AV0Z2041904 Keywords : Tempered martensite ferritic steels * martensite variants * orientation imaging Subject RIV: JG - Metallurgy Impact factor: 0.637, year: 2003

  3. Parallel continuous simulated tempering and its applications in large-scale molecular simulations

    Energy Technology Data Exchange (ETDEWEB)

    Zang, Tianwu; Yu, Linglin; Zhang, Chong [Applied Physics Program and Department of Bioengineering, Rice University, Houston, Texas 77005 (United States); Ma, Jianpeng, E-mail: jpma@bcm.tmc.edu [Applied Physics Program and Department of Bioengineering, Rice University, Houston, Texas 77005 (United States); Verna and Marrs McLean Department of Biochemistry and Molecular Biology, Baylor College of Medicine, One Baylor Plaza, BCM-125, Houston, Texas 77030 (United States)

    2014-07-28

    In this paper, we introduce a parallel continuous simulated tempering (PCST) method for enhanced sampling in studying large complex systems. It mainly inherits the continuous simulated tempering (CST) method in our previous studies [C. Zhang and J. Ma, J. Chem. Phys. 130, 194112 (2009); C. Zhang and J. Ma, J. Chem. Phys. 132, 244101 (2010)], while adopts the spirit of parallel tempering (PT), or replica exchange method, by employing multiple copies with different temperature distributions. Differing from conventional PT methods, despite the large stride of total temperature range, the PCST method requires very few copies of simulations, typically 2–3 copies, yet it is still capable of maintaining a high rate of exchange between neighboring copies. Furthermore, in PCST method, the size of the system does not dramatically affect the number of copy needed because the exchange rate is independent of total potential energy, thus providing an enormous advantage over conventional PT methods in studying very large systems. The sampling efficiency of PCST was tested in two-dimensional Ising model, Lennard-Jones liquid and all-atom folding simulation of a small globular protein trp-cage in explicit solvent. The results demonstrate that the PCST method significantly improves sampling efficiency compared with other methods and it is particularly effective in simulating systems with long relaxation time or correlation time. We expect the PCST method to be a good alternative to parallel tempering methods in simulating large systems such as phase transition and dynamics of macromolecules in explicit solvent.

  4. Solar Radiation Disinfection of Drinking Water at Temperate Latitudes: Inactivation rates for an optimized reactor configuration

    Science.gov (United States)

    Solar radiation-driven inactivation of bacteria, virus and protozoan pathogen models was quantified in simulated drinking water at a temperate latitude (34°S). The water was seeded with Enterococcus faecalis, Clostridium sporogenes spores, and P22 bacteriophage, each at ca 1 x 10...

  5. Numerical simulation of tropical-temperate troughs over Southern Africa using the CSU RAMS model

    CSIR Research Space (South Africa)

    Van den Heever, SC

    1997-08-01

    Full Text Available ) and the wet (1981) late summer case studies has been examined. Model simulations reveal that the tropical-temperate troughs form when an upper westerly wave coincides with an easterly, wave or depression in lower levels. These systems occur preferentially over...

  6. Mechanisms of nitrogen deposition effects on temperate forest lichens and trees

    Science.gov (United States)

    Therese S. Carter; Christopher M. Clark; Mark E. Fenn; Sarah Jovan; Steven S. Perakis; Jennifer Riddell; Paul G. Schaberg; Tara L. Greaver; Meredith G. Hastings

    2017-01-01

    We review the mechanisms of deleterious nitrogen (N) deposition impacts on temperate forests, with a particular focus on trees and lichens. Elevated anthropogenic N deposition to forests has varied effects on individual organisms depending on characteristics both of the N inputs (form, timing, amount) and of the organisms (ecology, physiology) involved. Improved...

  7. A conceptual framework for predicting temperate ecosystem sensitivity to human impacts on fire regimes

    Science.gov (United States)

    D. B. McWethy; P. E. Higuera; C. Whitlock; T. T. Veblen; D. M. J. S. Bowman; G. J. Cary; S. G. Haberle; R. E. Keane; B. D. Maxwell; M. S. McGlone; G. L. W. Perry; J. M. Wilmshurst

    2013-01-01

    The increased incidence of large fires around much of the world in recent decades raises questions about human and non-human drivers of fire and the likelihood of increased fire activity in the future. The purpose of this paper is to outline a conceptual framework for examining where human-set fires and feedbacks are likely to be most pronounced in temperate forests...

  8. Influence of tempering on mechanical strains in Mo2Si films

    International Nuclear Information System (INIS)

    Zscheile, H.D.

    1984-01-01

    Amorphous or crystalline MoSi 2 films on (111) silicon wafers, deposited by DC triode sputtering, showed compressive stress. Tensile stress was found in the same films formed by dual electron beam evaporation. By isochronous tempering in the temperature range of 300 to 1270 K the compressive stress of the sputtered films was converted into tensile stress

  9. Estimating Janka hardness from specific gravity for tropical and temperate species

    Science.gov (United States)

    Michael C. Wiemann; David W. Green

    2007-01-01

    Using mean values for basic (green) specific gravity and Janka side hardness for individual species obtained from the world literature, regression equations were developed to predict side hardness from specific gravity. Statistical and graphical methods showed that the hardness–specific gravity relationship is the same for tropical and temperate hardwoods, but that the...

  10. Suitability of close-to-nature silviculture for adapting temperate European forests to climate change

    NARCIS (Netherlands)

    Brang, P.; Spathelf, P.; Larsen, J.B.; Bauhus, J.; Boncina, A.; Mohren, G.M.J.

    2014-01-01

    In many parts of Europe, close-to-nature silviculture (CNS) has been widely advocated as being the best approach for managing forests to cope with future climate change. In this review, we identify and evaluate six principles for enhancing the adaptive capacity of European temperate forests in a

  11. RECONSTRUCTING THE EVOLUTIONARY HISTORY OF THE FOREST FUNGAL PATHOGEN, ARMILLARIA MELLEA, IN A TEMPERATE WORLDWIDE POPULATIONS

    Science.gov (United States)

    The forest pathogen Armillaria mellea s.s. (Basidiomycota, Physalacriaceae) is among the most significant forest pathogens causing root rot in northern temperate forest trees worldwide. Phylogenetic reconstructions for A. mellea show distinct European, Asian and North American lineages. The North Am...

  12. Seed dispersal by fishes in tropical and temperate fresh waters: The growing evidence

    NARCIS (Netherlands)

    Horn, M.H.; Correa, S.B.; Parolin, P.; Pollux, B.J.A.; Anderson, J.T.; Lucas, C.; Widmann, P.; Tjiu, A.; Galetti, M.; Goulding, M.

    2011-01-01

    Fruit-eating by fishes represents an ancient (perhaps Paleozoic) interaction increasingly regarded as important for seed dispersal (ichthyochory) in tropical and temperate ecosystems. Most of the more than 275 known frugivorous species belong to the mainly Neotropical Characiformes (pacus, piranhas)

  13. Substrate and nutrient limitation of ammonia-oxidizing bacteria and archaea in temperate forest soil

    Science.gov (United States)

    J.S. Norman; J.E. Barrett

    2014-01-01

    Ammonia-oxidizing microbes control the rate-limiting step of nitrification, a critical ecosystem process, which affects retention and mobility of nitrogen in soil ecosystems. This study investigated substrate (NH4þ) and nutrient (K and P) limitation of ammonia-oxidizing bacteria (AOB) and ammonia-oxidizing archaea (AOA) in temperate forest soils at Coweeta Hydrologic...

  14. Planetesimal Formation in the Warm, Inner Disk: Experiments with Tempered Dust

    Energy Technology Data Exchange (ETDEWEB)

    De Beule, Caroline; Landers, Joachim; Salamon, Soma; Wende, Heiko; Wurm, Gerhard, E-mail: gerhard.wurm@uni-due.de [Faculty of Physics, University of Duisburg-Essen, Lotharstr. 1, D-47057 Duisburg (Germany)

    2017-03-01

    It is an open question how elevated temperatures in the inner parts of protoplanetary disks influence the formation of planetesimals. We approach this problem here by studying the tensile strength of granular beds with dust samples tempered at different temperatures. We find via laboratory experiments that tempering at increasing temperatures is correlated with an increase in cohesive forces. We studied dust samples of palagonite (JSC Mars-1a) which were tempered for up to 200 hr at temperatures between 600 and 1200 K, and measured the relative tensile strengths of highly porous dust layers once the samples cooled to room temperature. Tempering increases the tensile strength from 800 K upwards. This change is accompanied by mineral transformations, the formation of iron oxide crystallites as analyzed by Mössbauer spectroscopy, changes in the number size distribution, and the morphology of the surface visible as cracks in larger grains. These results suggest a difference in the collisional evolution toward larger bodies with increasing temperature as collisional growth is fundamentally based on cohesion. While high temperatures might also increase sticking (not studied here), compositional evolution will already enhance the cohesion and the possibility of growing larger aggregates on the way toward planetesimals. This might lead to a preferred in situ formation of inner planets and explain the observed presence of dense inner planetary systems.

  15. Mechanisms promoting higher growth rate in arctic than in temperate shorebirds

    NARCIS (Netherlands)

    Schekkerman, H.; Tulp, I.Y.M.; Piersma, T.; Visser, G.H.

    2003-01-01

    We compared prefledging growth, energy expenditure, and time budgets in the arctic-breeding red knot (Calidris canutus) to those in temperate shorebirds, to investigate how arctic chicks achieve a high growth rate despite energetic difficulties associated with precocial development in a cold

  16. Simple model for daily evaporation from fallow tilled soil under spring conditions in a temperate climate.

    NARCIS (Netherlands)

    Boesten, J.J.T.I.; Stroosnijder, L.

    1986-01-01

    A simple parametric model is presented to estimate daily evaporation from fallow tilled soil under spring conditions in a temperate climate. In this model, cumulative actual evaporation during a drying cycle is directly proportional to the square root of cumulative potential evaporation. The model

  17. Mechanisms promoting higher growth rate in arctic than in temperate shorebirds

    NARCIS (Netherlands)

    Schekkerman, H; Tulp, Ingrid; Piersma, T.; Visser, G.H.

    We compared prefledging growth, energy expenditure, and time budgets in the arctic-breeding red knot (Calidris canutus) to those in temperate shorebirds, to investigate how arctic chicks achieve a high growth rate despite energetic difficulties associated with precocial development in a cold

  18. Heat stress in cows at pasture and benefit of shade in a temperate climate region

    Science.gov (United States)

    Veissier, Isabelle; Van laer, Eva; Palme, Rupert; Moons, Christel P. H.; Ampe, Bart; Sonck, Bart; Andanson, Stéphane; Tuyttens, Frank A. M.

    2017-11-01

    Under temperate climates, cattle are often at pasture in summer and are not necessarily provided with shade. We aimed at evaluating in a temperate region (Belgium) to what extent cattle may suffer from heat stress (measured through body temperature, respiration rate and panting score, cortisol or its metabolites in milk, and feces on hot days) and at assessing the potential benefits of shade. During the summer of 2012, 20 cows were kept on pasture without access to shade. During the summer of 2011, ten cows had access to shade (young trees with shade cloth hung between them), whereas ten cows had no access. Climatic conditions were quantified by the Heat Load Index (HLI). In animals without access to shade respiration rates, panting scores, rectal temperatures, and milk cortisol concentrations increased as HLI increased in both 2011 and 2012. Fecal cortisol metabolites varied with HLI in 2011 only. When cattle had access to shade, their use of shade increased as the HLI increased. This effect was more pronounced during the last part of the summer, possibly due to better acquaintance with the shade construction. In this case, shade use increased to 65% at the highest HLI (79). Shade tempered the effects on respiration, rectal temperature, and fecal cortisol metabolites. Milk cortisol was not influenced by HLI for cows using shade for > 10% of the day. Therefore, even in temperate areas, cattle may suffer from heat when they are at pasture in summer and providing shade can reduce such stress.

  19. The marine radiocarbon bomb pulse across the temperate North Atlantic: a compilation of Delta

    NARCIS (Netherlands)

    Scourse, J.D.; Wanamaker jr., A.D.; Weidman, C.; Heinemeier, J.; Reimer, P.J.; Butler, P.G.; Witbaard, R.; Richardson, C.A.

    2012-01-01

    Marine radiocarbon bomb-pulse time histories of annually resolved archives from temperate regions have been underexploited. We present here series of Delta C-14 excess from known-age annual increments of the long-lived bivalve mollusk Arctica islandica from 4 sites across the coastal North Atlantic

  20. Modeling complex effects of multiple environmental stresses on carbon dynamics of Mid-Atlantic temperate forests

    Science.gov (United States)

    Yude Pan; Richard Birdsey; John Hom; Kevin McCullough

    2007-01-01

    We used our GIS variant of the PnET-CN model to investigate changes of forest carbon stocks and fluxes in Mid-Atlantic temperate forests over the last century (1900-2000). Forests in this region are affected by multiple environmental changes including climate, atmospheric CO2 concentration, N deposition and tropospheric ozone, and extensive land disturbances. Our...