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Sample records for surrounding yellowstone national

  1. New challenges for grizzly bear management in Yellowstone National Park

    Science.gov (United States)

    van Manen, Frank T.; Gunther, Kerry A.

    2016-01-01

    A key factor contributing to the success of grizzly bear Ursus arctos conservation in the Greater Yellowstone Ecosystem has been the existence of a large protected area, Yellowstone National Park. We provide an overview of recovery efforts, how demographic parameters changed as the population increased, and how the bear management program in Yellowstone National Park has evolved to address new management challenges over time. Finally, using the management experiences in Yellowstone National Park, we present comparisons and perspectives regarding brown bear management in Shiretoko National Park.

  2. Atlas of Yellowstone

    Science.gov (United States)

    Pierce, Kenneth L.; Marcus, A. W.; Meachan, J. E.; Rodman, A. W.; Steingisser, A. Y.; Allan, Stuart; West, Ross

    2012-01-01

    Established in 1872, Yellowstone National Park was the world’s first national park. In a fitting tribute to this diverse and beautiful region, the Atlas of Yellowstone is a compelling visual guide to this unique national park and its surrounding area. Ranging from art to wolves, from American Indians to the Yellowstone Volcano, and from geysers to population, each page explains something new about the dynamic forces shaping Yellowstone. Equal parts reference and travel guide, the Atlas of Yellowstone is an unsurpassed resource.

  3. Exterior sound level measurements of snowcoaches at Yellowstone National Park

    Science.gov (United States)

    2010-04-01

    Sounds associated with oversnow vehicles, such as snowmobiles and snowcoaches, are an important management concern at Yellowstone and Grand Teton National Parks. The John A. Volpe National Transportation Systems Centers Environmental Measurement a...

  4. 75 FR 27579 - Bison Brucellosis Remote Vaccination, Draft Environmental Impact Statement, Yellowstone National...

    Science.gov (United States)

    2010-05-17

    ... DEPARTMENT OF THE INTERIOR National Park Service Bison Brucellosis Remote Vaccination, Draft... Brucellosis Remote Vaccination Program, Yellowstone National Park. SUMMARY: Pursuant to the National... the Bison Brucellosis Remote Vaccination Draft Environmental Impact Statement (EIS) for Yellowstone...

  5. 78 FR 12353 - Winter Use Plan, Supplemental Environmental Impact Statement, Yellowstone National Park

    Science.gov (United States)

    2013-02-22

    ...] Winter Use Plan, Supplemental Environmental Impact Statement, Yellowstone National Park AGENCY: National... Final Supplemental Environmental Impact Statement (SEIS) for a Winter Use Plan for Yellowstone National... link to the 2012 Supplemental Winter Use Plan EIS), and at Yellowstone National Park headquarters...

  6. 76 FR 77131 - Special Regulations; Areas of the National Park System, Yellowstone National Park

    Science.gov (United States)

    2011-12-12

    ... FURTHER INFORMATION CONTACT: Steve Iobst, Deputy Superintendent, Yellowstone National Park, (307) 344-2002... material way the economy, productivity, competition, jobs, the environment, public health or safety, or...

  7. Examining winter visitor use in Yellowstone National Park

    Science.gov (United States)

    Mae A. Davenport; Wayne A. Freimund; William T. Borrie; Robert E. Manning; William A. Valliere; Benjamin Wang

    2000-01-01

    This research was designed to assist the managers of Yellowstone National Park (YNP) in their decision making about winter visitation. The focus of this report is on winter use patterns and winter visitor preferences. It is the author’s hope that this information will benefit both the quality of winter experiences and the stewardship of the park resources. This report...

  8. Wildlife health initiatives in Yellowstone National Park

    Science.gov (United States)

    Cross, Paul C.; Plumb, G.

    2007-01-01

    Yellowstone Science 15(2) • 2007 and conservation organizations ( see inset page 7, The Yellowstone Wildlife Health Program ). Wildlife and Human Health are Linked Much of the interest in disease ecology and wildlife health has been prompted by the emergence, or resurgence, of many parasites that move between livestock, wildlife, and/or humans. Wildlife diseases are important because of their impact on both the natural ecosystem and human health. Many human dis - eases arise from animal reservoirs (WHO 2002). Hantaviruses, West Nile virus, avian influenza, and severe acute respiratory syndrome (SARS) are examples of disease issues that have arisen over the last decade. Indeed, nearly 75% of all emerg - ing human infectious diseases are zoonotic (a disease that has spread to humans from another animal species). Many of these diseases have spilled over from natural wildlife reservoirs either directly into humans or via domestic animals (WHO/FAO/ OIE 2004). Unprecedented human population abundance and distribution, combined with anthropogenic environmental change, has resulted in dramatic increases in human–animal contact, thus increasing the intimate linkages between animal and human health (Figure 1). Linkage of human and animal health is not a new phenomenon, but the scope, scale, and worldwide impacts of contemporary zoonoses have no historical precedent (OIE 2004a). Zoonotic infectious diseases can have major impacts on wild and domestic animals and human health, resulting in

  9. 77 FR 6581 - Winter Use Plan, Supplemental Environmental Impact Statement, Yellowstone National Park, Idaho...

    Science.gov (United States)

    2012-02-08

    ... DEPARTMENT OF THE INTERIOR National Park Service [2310-0070-422] Winter Use Plan, Supplemental... the Winter Use Plan, Yellowstone National Park. SUMMARY: Pursuant to the National Environmental Policy... Statement (SEIS) for a Winter Use Plan for Yellowstone National Park, located in Idaho, Montana and Wyoming...

  10. 76 FR 77249 - Winter Use Plan, Final Environmental Impact Statement Record of Decision, Yellowstone National...

    Science.gov (United States)

    2011-12-12

    ... DEPARTMENT OF THE INTERIOR National Park Service Winter Use Plan, Final Environmental Impact... Impact Statement for a Winter Use Plan, Yellowstone National Park. SUMMARY: Pursuant to Sec. 102(2)(C) of... Winter Use Plan for Yellowstone National Park, located in Idaho, Montana, and Wyoming. On December 5...

  11. Grizzly bear management in Yellowstone National Park: The heart of recovery in the Yellowstone Ecosystem

    Science.gov (United States)

    Schwartz, C.C.; Gunther, K.; McCullough, Dale R.; Kaji, Koichi; Yamanaka, Masami

    2006-01-01

    Grizzly bear (Ursus arctos) management in the Greater Yellowstone Ecosystem (GYE) in the past quarter century has resulted in more than doubling of the population from around 200 to more than 500, expansion of range back into habitats where the bear has extirpated more than a century ago, and a move toward removal from the U.S. Endangered Species list. At the center of this success story are the management programs in Yellowstone National Park (YNP). Regulations that restrict human activity, camping, and food storage, elimination of human food and garbage as attractants, and ranger attendance of roadside bears have all resulted in the population of grizzlies in YNP approaching carrying capacity. Recent studies suggest, however, that YNP alone is too small to support the current population, making management beyond the park boundary important and necessary to the demographics of the population as a whole. Demographic analyses suggest a source-sink dynamic exists within the GYE, with YNP and lands outside the park within the Grizzly Bear Recovery Zone (RZ) representing source habitats, whereas lands beyond the RZ constitute sinks. The source-sink demography in the GYE is indicative of carnivore conservation issues worldwide where many national parks or preserves designed to protect out natural resources are inadequate in size or shape to provide all necessary life history requirements for these wide-ranging species. Additionally, wide-ranging behavior and long-distance dispersal seem inherent to large carnivores, so mortality around the edges is virtually inevitable, and conservation in the GYE is inextricably linked to management regimes not only within YNP, but within the GYE as a whole. We discuss those needs here.

  12. The 2017 Maple Creek Seismic Swarm in Yellowstone National Park

    Science.gov (United States)

    Pang, G.; Hale, J. M.; Farrell, J.; Burlacu, R.; Koper, K. D.; Smith, R. B.

    2017-12-01

    The University of Utah Seismograph Stations (UUSS) performs near-real-time monitoring of seismicity in the region around Yellowstone National Park in partnership with the United States Geological Survey and the National Park Service. UUSS operates and maintains 29 seismic stations with network code WY (short-period, strong-motion, and broadband) and records data from five other seismic networks—IW, MB, PB, TA, and US—to enhance the location capabilities in the Yellowstone region. A seismic catalog is produced using a conventional STA/LTA detector and single-event location techniques (Hypoinverse). On June 12, 2017, a seismic swarm began in Yellowstone National Park about 5 km east of Hebgen Lake. The swarm is adjacent to the source region of the 1959 MW 7.3 Hebgen Lake earthquake, in an area corresponding to positive Coulumb stress change from that event. As of Aug. 1, 2017, the swarm consists of 1481 earthquakes with 1 earthquake above magnitude 4, 8 earthquakes in the magnitude 3 range, 115 earthquakes in the magnitude 2 range, 469 earthquakes in the magnitude 1 range, 856 earthquakes in the magnitude 0 range, 22 earthquakes with negative magnitudes, and 10 earthquakes with no magnitude. Earthquake depths are mostly between 3 and 10 km and earthquake depth increases toward the northwest. Moment tensors for the 2 largest events (3.6 MW and 4.4. MW) show strike-slip faulting with T axes oriented NE-SW, consistent with the regional stress field. We are currently using waveform cross-correlation methods to measure differential travel times that are being used with the GrowClust program to generate high-accuracy relative relocations. Those locations will be used to identify structures in the seismicity and make inferences about the tectonic and magmatic processes causing the swarm.

  13. Exterior sound level measurements of over-snow vehicles at Yellowstone National Park.

    Science.gov (United States)

    2008-09-30

    Sounds associated with oversnow vehicles, such as snowmobiles and snowcoaches, are an : important management concern at Yellowstone and Grand Teton National Parks. The John A. : Volpe National Transportation Systems Centers Environmental Measureme...

  14. 75 FR 53979 - Bison Brucellosis Remote Vaccination, Draft Environmental Impact Statement, Yellowstone National...

    Science.gov (United States)

    2010-09-02

    ... DEPARTMENT OF THE INTERIOR National Park Service Bison Brucellosis Remote Vaccination, Draft Environmental Impact Statement, Yellowstone National Park, Wyoming ACTION: Reopening of public comment period... Brucellosis Remote Vaccination Draft Environmental Impact Statement. The original comment period was from 28...

  15. Serological survey for diseases in free-ranging coyotes (Canis latrans) in Yellowstone National Park, Wyoming.

    Science.gov (United States)

    Gese, E M; Schultz, R D; Johnson, M R; Williams, E S; Crabtree, R L; Ruff, R L

    1997-01-01

    From October 1989 to June 1993, we captured and sampled 110 coyotes (Canis latrans) for various diseases in Yellowstone National Park, Wyoming (USA). Prevalence of antibodies against canine parvovirus (CPV) was 100% for adults (> 24 months old), 100% for yearlings (12 to 24 months old), and 100% for old pups (4 to 12 months old); 0% of the young pups (Yellowstone National Park, with CPV influencing coyote pup survival during the first 3 months of life; eight of 21 transmitted pups died of CPV infection in 1992. The potential impact of these canine pathogens on wolves (C. lupus) reintroduced to Yellowstone National Park remains to be documented.

  16. Fungi from geothermal soils in Yellowstone National Park

    Science.gov (United States)

    Redman, R.S.; Litvintseva, A.; Sheehan, K.B.; Henson, J.M.; Rodriguez, R.J.

    1999-01-01

    Geothermal soils near Amphitheater Springs in Yellowstone National Park were characterized by high temperatures (up to 70??C), high heavy metal content, low pH values (down to pH 2.7), sparse vegetation, and limited organic carbon. From these soils we cultured 16 fungal species. Two of these species were thermophilic, and six were thermotolerant. We cultured only three of these species from nearby cool (0 to 22??C) soils. Transect studies revealed that higher numbers of CFUs occurred in and below the root zone of the perennial plant Dichanthelium lanuginosum (hot springs panic grass). The dynamics of fungal CFUs in geothermal soil and nearby nongeothermal soil were investigated for 12 months by examining soil cores and in situ mesocosms. For all of the fungal species studied, the temperature of the soil from which the organisms were cultured corresponded with their optimum axenic growth temperature.

  17. On the origin of brucellosis in bison of Yellowstone National Park: a review

    Science.gov (United States)

    Meagher, Mary; Meyer, Margaret E.

    1994-01-01

    Brucellosis caused by Brucella abortus occurs in the free-ranging bison (Bison bison) of Yellowstone and Wood Buffalo National Parks and in elk (Cervus elaphus) of the Greater Yellowstone Area. As a result of nationwide bovine brucellosis eradication programs, states and provinces proximate to the national parks are considered free of bovine brucellosis. Thus, increased attention has been focused on the wildlife within these areas as potential reservoirs for transmission to cattle. Because the national parks are mandated as natural areas, the question has been raised as to whether Brucella abortus is endogenous or exogenous to bison, particularly for Yellowstone National Park. We synthesized diverse lines of inquiry, including the evolutionary history of both bison and Brucella, wild animals as Brucella hosts, biochemical and genetic information, behavioral characteristics of host and organism, and area history to develop an evaluation of the question for the National Park Service. All lines of inquiry indicated that the organism was introduced to North America with cattle, and that the introduction into the Yellowstone bison probably was directly from cattle shortly before 1917. Fistulous withers of horses was a less likely possibility. Elk on winter feedgrounds south of Yellowstone National Park apparently acquired the disease directly from cattle. Bison presently using Grand Teton National Park probably acquired brucellosis from feedground elk.

  18. Wolf-bison interactions in Yellowstone National Park

    Science.gov (United States)

    Smith, Douglas W.; Mech, L. David; Meagher, Mary; Clark, Wendy E.; Jaffe, Rosemary; Phillips, Michael K.; Mack, John A.

    2000-01-01

    We studied interactions of reintroduced wolves (Canis lupus) with bison (Bison bison) in Yellowstone National Park. Only 2 of 41 wolves in this study had been exposed to bison before their translocation. Wolves were more successful killing elk (Cervus elaphus) than bison, and elk were more abundant than bison, so elk were the primary prey of wolves. Except for a lone emaciated bison calf killed by 8 1-year-old wolves 21 days after their release, the 1st documented kill occurred 25 months after wolves were released. Fourteen bison kills were documented from April 1995 through March 1999. All kills were made in late winter when bison were vulnerable because of poor condition or of bison that were injured or young. Wolves learned to kill bison and killed more bison where elk were absent or scarce. We predict that wolves that have learned to kill bison will kill them more regularly, at least in spring. The results of this study indicate how adaptable wolves are at killing prey species new to them.

  19. Brucellosis in Yellowstone National Park bison: Quantitative serology and infection

    Science.gov (United States)

    Roffe, T.J.; Rhyan, Jack C.; Aune, K.; Philo, L.M.; Ewalt, D.R.; Gidlewski, T.; Hennager, S.G.

    1999-01-01

    We collected complete sets of tissues, fluids, and swabs (approx 30) from 37 Yellowstone National Park (YNP) female bison (Bison bison) killed as a result of management actions by the Montana Department of Livestock and YNP personnel. Our goal was to establish the relation between blood tests demonstrating an animal has antibody to Brucella and the potential of that animal to be infected during the second trimester of pregnancy, the time when most management actions are taken. Twenty-eight of the 37 bison were seropositive adults (27) or a seropositive calf (1). We cultured samples using macerated whole tissues plated onto 4 Brucella-selective media and incubated with added CO2 for 1 week. Specimens from 2 adult seropositive females were contaminated, thus eliminating them from our data. Twelve of the remaining 26 seropositive adult and calf female bison (46%) were culture positive for Brucella abortus from 1 or more tissues. Culture positive adult females had high serologic titers. All 11 adults measured 3+ at 1:40 for 10 of 11 (91%) animals. All culture positive female adults had either a PCFIA ???0.080 or a CF reaction ???4+ at 1:80. However 5 (36%) bison with high titers were culture negative for B. abortus. Our findings on the relation between Brucella serology and culture are similar to those reported from studies of chronically infected cattle herds.

  20. Pathology of brucellosis in bison from Yellowstone National Park

    Science.gov (United States)

    Rhyan, Jack C.; Gidlewski, T.; Roffe, T.J.; Aune, K.; Philo, L.M.; Ewalt, D.R.

    2001-01-01

    Between February 1995 and June 1999, specimens from seven aborted bison (Bison bison) fetuses or stillborn calves and their placentas, two additional placentas, three dead neonates, one 2-wk-old calf, and 35 juvenile and adult female bison from Yellowstone National Park (USA) were submitted for bacteriologic and histopathologic examination. One adult animal with a retained placenta had recently aborted. Serum samples from the 35 juvenile and adult bison were tested for Brucella spp. antibodies. Twenty-six bison, including the cow with the retained placenta, were seropositive, one was suspect, and eight were seronegative. Brucella abortus biovar 1 was isolated from three aborted fetuses and associated placentas, an additional placenta, the 2-wk-old calf, and 11 of the seropositive female bison including the animal that had recently aborted. Brucella abortus biovar 2 was isolated from one additional seropositive adult female bison. Brucella abortus was recovered from numerous tissue sites from the aborted fetuses, placentas and 2-wk-old calf. In the juvenile and adult bison, the organism was more frequently isolated from supramammary (83%), retropharyngeal (67%), and iliac (58%) lymph nodes than from other tissues cultured. Cultures from the seronegative and suspect bison were negative for B. abortus. Lesions in the B. abortus-infected, aborted placentas and fetuses consisted of necropurulent placentitis and mild bronchointerstitial pneumonia. The infected 2-wk-old calf had bronchointerstitial pneumonia, focal splenic infarction, and purulent nephritis. The recently-aborting bison cow had purulent endometritis and necropurulent placentitis. Immunohistochemical staining of tissues from the culture-positive aborted fetuses, placentas, 2-wk-old calf, and recently-aborting cow disclosed large numbers of B. abortus in placental trophoblasts and exudate, and fetal and calf lung. A similar study with the same tissue collection and culture protocol was done using six

  1. Community analysis of plant biomass-degrading microorganisms from Obsidian Pool, Yellowstone National Park.

    Science.gov (United States)

    Vishnivetskaya, Tatiana A; Hamilton-Brehm, Scott D; Podar, Mircea; Mosher, Jennifer J; Palumbo, Anthony V; Phelps, Tommy J; Keller, Martin; Elkins, James G

    2015-02-01

    The conversion of lignocellulosic biomass into biofuels can potentially be improved by employing robust microorganisms and enzymes that efficiently deconstruct plant polysaccharides at elevated temperatures. Many of the geothermal features of Yellowstone National Park (YNP) are surrounded by vegetation providing a source of allochthonic material to support heterotrophic microbial communities adapted to utilize plant biomass as a primary carbon and energy source. In this study, a well-known hot spring environment, Obsidian Pool (OBP), was examined for potential biomass-active microorganisms using cultivation-independent and enrichment techniques. Analysis of 33,684 archaeal and 43,784 bacterial quality-filtered 16S rRNA gene pyrosequences revealed that archaeal diversity in the main pool was higher than bacterial; however, in the vegetated area, overall bacterial diversity was significantly higher. Of notable interest was a flooded depression adjacent to OBP supporting a stand of Juncus tweedyi, a heat-tolerant rush commonly found growing near geothermal features in YNP. The microbial community from heated sediments surrounding the plants was enriched in members of the Firmicutes including potentially (hemi)cellulolytic bacteria from the genera Clostridium, Anaerobacter, Caloramator, Caldicellulosiruptor, and Thermoanaerobacter. Enrichment cultures containing model and real biomass substrates were established at a wide range of temperatures (55-85 °C). Microbial activity was observed up to 80 °C on all substrates including Avicel, xylan, switchgrass, and Populus sp. Independent of substrate, Caloramator was enriched at lower (65 °C) temperatures.

  2. Magnetic susceptibility measurements in Yellowstone National Park, USA; Beikoku Yellowstone kokuritsu koen ni okeru genchi jikaritsu sokutei

    Energy Technology Data Exchange (ETDEWEB)

    Okuma, S [Geological Survey of Japan, Tsukuba (Japan)

    1997-05-27

    For the purpose of interpreting data of the aeromagnetic anomalies in Yellowstone National Park in the U.S.A., in-situ magnetization intensity measurements have been carried out in 1994 and 1995 on geological outcrops of rocks in that area. Comparisons and discussions were given on the measurement results, and existing rock magnetic data and aeromagnetic anomaly data available for the area. Outside the Yellowstone caldera, part of granitic gneisses among the Precambrian granitic gneisses and crystalline schists distributed to the north has an abnormally high magnetization intensity of 1 {times} 10 {sup -2} SI. This could be a powerful anomaly source for the high magnetic anomaly in this area. Paleogene volcanic rocks distributed widely in the eastern part of the park also have magnetization intensity as high as 1 {times} 10 {sup -2} SI or higher, which are also thought a powerful anomaly source in this area. Part of Pleistocene basalts which are exposed partially in the western part of the park has also very high magnetization intensity at 1 {times} 10 {sup -2} SI or higher. This suggests correlation with the magnetic anomaly in the east-west direction distributed in this area. Quaternary rhyolites are more magnetic than Quaternary welded tuffs, which should give greater effects to the magnetic anomaly. 10 refs., 5 figs.

  3. 77 FR 38824 - Winter Use Plan, Supplemental Draft Environmental Impact Statement, Yellowstone National Park

    Science.gov (United States)

    2012-06-29

    ... DEPARTMENT OF THE INTERIOR National Park Service [2310-0070-422] Winter Use Plan, Supplemental.... ACTION: Notice of Availability of the Draft Supplemental Environmental Impact Statement for the Winter... Supplemental Environmental Impact Statement (Draft SEIS) for a Winter Use Plan for Yellowstone National Park...

  4. 76 FR 68503 - Winter Use Plan, Final Environmental Impact Statement, Yellowstone National Park, Idaho, Montana...

    Science.gov (United States)

    2011-11-04

    ... DEPARTMENT OF THE INTERIOR National Park Service Winter Use Plan, Final Environmental Impact.... ACTION: Notice of availability of the Final Environmental Impact Statement for the Winter Use Plan... Winter Use Plan for Yellowstone National Park, located in Idaho, Montana, and [[Page 68504

  5. 75 FR 4842 - Winter Use Plan, Environmental Impact Statement, Yellowstone National Park

    Science.gov (United States)

    2010-01-29

    ... DEPARTMENT OF THE INTERIOR National Park Service Winter Use Plan, Environmental Impact Statement... to prepare an Environmental Impact Statement for a Winter Use Plan, Yellowstone National Park... Park Service (NPS) is preparing an Environmental Impact Statement (EIS) for a Winter Use Plan for...

  6. The U S national parks in international perspective: The Yellowstone model or conservation syncretism?

    Science.gov (United States)

    John Schelhas

    2010-01-01

    In recent years, international conservation scholars and practitioners have largely dismissed the U.S. national park experience, often termed the “Yellowstone model,” as being too protectionist and exclusionary, and therefore irrelevant and even detrimental to park management and policy in lesser developed countries. A review of the U.S. national park experience finds...

  7. Winter visitor use planning in Yellowstone and Grand Teton National Parks

    Science.gov (United States)

    John A. Sacklin; Kristin L. Legg; M. Sarah Creachbaum; Clifford L. Hawkes; George Helfrich

    2000-01-01

    Winter use in Yellowstone and Grand Teton National Parks increased dramatically in the 1980s and early 1990s. That increase and the emphasis on snowmobiles as the primary mode of transportation brought into focus a host of winter-related issues, including air pollution, unwanted sound, wildlife impacts and the adequacy of agency budgets, staff and infrastructure to...

  8. Denali Park wolf studies: Implications for Yellowstone

    Science.gov (United States)

    Mech, L. David; Meier, Thomas J.; Burch, John W.

    1991-01-01

    The Northern Rocky Mountain Wolf Recovery Plan approved by the U.S. Fish and Wildlife Service (1987) recommends re-establishment of wolves (Canis lupus) in Yellowstone National Park. Bills proposing wolf re-establishment in the Park have been introduced into the U.S. House and Senate. However, several questions have been raised about the possible effects of wolf re-establishment on other Yellowstone Park fauna, on human use of the Park and on human use of surrounding areas. Thus the proposed wolf re-establishment remains controversial.Information pertinent to some of the above questions is available from a current study of wolf ecology in Denali National Park and Preserve, Alaska, which we began in 1986. Although Denali Park differs from Yellowstone in several ways, it is also similar enough in important respects to provide insight into questions raised about wolf re-establishment in Yellowstone.

  9. Creating Conditions for Policy Change in National Parks: Contrasting Cases in Yellowstone and Yosemite

    Science.gov (United States)

    Yochim, Michael J.; Lowry, William R.

    2016-05-01

    Public agencies face significant political obstacles when they try to change long-standing policies. This paper examines efforts by the U.S. National Park Service to change long-term policies in Yellowstone and Yosemite national parks. We argue that, to be successful, the agency and pro-change allies must expand the sphere of conflict to engage the support of the broader American public through positive framing, supportive science, compelling economic arguments, consistent goals, and the commitment of other institutional actors. We show that the agency is capable of creating these conditions, as in the reintroduction of wolves to Yellowstone, but we argue that this is not always the outcome, as in reducing automobile congestion in Yosemite Valley.

  10. A Serological Survey of Infectious Disease in Yellowstone National Park?s Canid Community

    OpenAIRE

    Almberg, Emily S.; Mech, L. David; Smith, Douglas W.; Sheldon, Jennifer W.; Crabtree, Robert L.

    2009-01-01

    BACKGROUND: Gray wolves (Canis lupus) were reintroduced into Yellowstone National Park (YNP) after a >70 year absence, and as part of recovery efforts, the population has been closely monitored. In 1999 and 2005, pup survival was significantly reduced, suggestive of disease outbreaks. METHODOLOGY/PRINCIPAL FINDINGS: We analyzed sympatric wolf, coyote (Canis latrans), and red fox (Vulpes vulpes) serologic data from YNP, spanning 1991-2007, to identify long-term patterns of pathogen exposure, i...

  11. Ferdinand Vandiveer Hayden and the founding of the Yellowstone National Park

    Science.gov (United States)

    ,

    1973-01-01

    Following the Civil War, the United States intensified the exploration of her western frontiers to gain a measure of the vast lands and natural resources in the region now occupied by our Rocky Mountain States. As part of this effort, the U.S. Geological and Geographical Survey of the Territories was formed and staffed under the leadership of geologist Ferdinand Vandiveer Hayden. Originally organized under the U.S. Public Land Office in 1861, the Hayden Survey (as it was most often identified) was placed under the Secretary of the Interior in 1869 and later, under the newly created U.S. Geological Survey. Its records, maps, and photographs were then transferred to the latter agency. In commemorating the centennial of Yellowstone National Park, the U.S. Geological Survey drew upon those items deposited by Hayden to describe the early exploration of the Yellowstone area and to recount events that led to the establishment of Yellowstone as the Nation's first national park.

  12. Predatory behavior of grizzly bears feeding on elk calves in Yellowstone National Park

    Science.gov (United States)

    French, Steven P.; French, Marilynn G.

    1990-01-01

    Grizzly bears (Ursus arctos horribilis) were observed preying on elk calves (Cervus elaphus) on 60 occasions in Yellowstone National Park, with 29 confirmed kills. Some bears were deliberate predators and effectively preyed on elk calves for short periods each spring, killing up to 1 calf daily. Primary hunting techniques were searching and chasing although some bears used a variety of techniques during a single hunt. They hunted both day and night and preyed on calves in the open and in the woods. Excess killing occurred when circumstances permitted. One bear caught 5 calves in a 15-minute interval. Elk used a variety of antipredator defenses and occasionally attacked predacious bears. The current level of this feeding behavior appears to be greater than previously reported. This is probably related to the increased availability of calves providing a greater opportunity for learning, and the adaptation of a more predatory behavior by some grizzly bears in Yellowstone.

  13. Climatic change and wetland desiccation cause amphibian decline in Yellowstone National Park.

    Science.gov (United States)

    McMenamin, Sarah K; Hadly, Elizabeth A; Wright, Christopher K

    2008-11-04

    Amphibians are a bellwether for environmental degradation, even in natural ecosystems such as Yellowstone National Park in the western United States, where species have been actively protected longer than anywhere else on Earth. We document that recent climatic warming and resultant wetland desiccation are causing severe declines in 4 once-common amphibian species native to Yellowstone. Climate monitoring over 6 decades, remote sensing, and repeated surveys of 49 ponds indicate that decreasing annual precipitation and increasing temperatures during the warmest months of the year have significantly altered the landscape and the local biological communities. Drought is now more common and more severe than at any time in the past century. Compared with 16 years ago, the number of permanently dry ponds in northern Yellowstone has increased 4-fold. Of the ponds that remain, the proportion supporting amphibians has declined significantly, as has the number of species found in each location. Our results indicate that climatic warming already has disrupted one of the best-protected ecosystems on our planet and that current assessments of species' vulnerability do not adequately consider such impacts.

  14. Protecting the Geyser Basins of Yellowstone National Park: Toward a New National Policy for a Vulnerable Environmental Resource

    Science.gov (United States)

    Barrick, Kenneth A.

    2010-01-01

    Geyser basins provide high value recreation, scientific, economic and national heritage benefits. Geysers are globally rare, in part, because development activities have quenched about 260 of the natural endowment. Today, more than half of the world’s remaining geysers are located in Yellowstone National Park, northwest Wyoming, USA. However, the hydrothermal reservoirs that supply Yellowstone’s geysers extend well beyond the Park borders, and onto two “Known Geothermal Resource Areas”—Island Park to the west and Corwin Springs on the north. Geysers are sensitive geologic features that are easily quenched by nearby geothermal wells. Therefore, the potential for geothermal energy development adjacent to Yellowstone poses a threat to the sustainability of about 500 geysers and 10,000 hydrothermal features. The purpose here is to propose that Yellowstone be protected by a “Geyser Protection Area” (GPA) extending in a 120-km radius from Old Faithful Geyser. The GPA concept would prohibit geothermal and large-scale groundwater wells, and thereby protect the water and heat supply of the hydrothermal reservoirs that support Yellowstone’s geyser basins and important hot springs. Proactive federal leadership, including buyouts of private groundwater development rights, can assist in navigating the GPA through the greater Yellowstone area’s “wicked” public policy environment. Moreover, the potential impacts on geyser basins from intrusive research sampling techniques are considered in order to facilitate the updating of national park research regulations to a precautionary standard. The GPA model can provide the basis for protecting the world’s few remaining geyser basins.

  15. History of pronghorn population monitoring, research, and management in Yellowstone National Park

    Science.gov (United States)

    Keating, Kim A.

    2002-01-01

    Pronghorn antelope in Yellowstone National Park (YNP) persist in a small population that historically has experienced recurrent, sometimes dramatic declines. They apparently are isolated from other pronghorns, depend partly on private lands for winter range, experience heavy predation of fawns, and concentrate during winter in a relatively small area, thereby increasing their vulnerability to factors like disease or locally extreme weather. Overall, the situation raises serious concerns about the long-term viability of this population. Although such concerns are not new, evidence of a dramatic population decline since 1991 and continued poor recruitment has created a renewed sense of urgency.

  16. Predicting breeding habitat for amphibians: a spatiotemporal analysis across Yellowstone National Park.

    Science.gov (United States)

    Bartelt, Paul E; Gallant, Alisa L; Klaver, Robert W; Wright, Chris K; Patla, Debra A; Peterson, Charles R

    2011-10-01

    The ability to predict amphibian breeding across landscapes is important for informing land management decisions and helping biologists better understand and remediate factors contributing to declines in amphibian populations. We built geospatial models of likely breeding habitats for each of four amphibian species that breed in Yellowstone National Park (YNP). We used field data collected in 2000-2002 from 497 sites among 16 basins and predictor variables from geospatial models produced from remotely sensed data (e.g., digital elevation model, complex topographic index, landform data, wetland probability, and vegetative cover). Except for 31 sites in one basin that were surveyed in both 2000 and 2002, all sites were surveyed once. We used polytomous regression to build statistical models for each species of amphibian from (1) field survey site data only, (2) field data combined with data from geospatial models, and (3) data from geospatial models only. Based on measures of receiver operating characteristic (ROC) scores, models of the second type best explained likely breeding habitat because they contained the most information (ROC values ranged from 0.70 to 0.88). However, models of the third type could be applied to the entire YNP landscape and produced maps that could be verified with reserve field data. Accuracy rates for models built for single years were highly variable, ranging from 0.30 to 0.78. Accuracy rates for models built with data combined from multiple years were higher and less variable, ranging from 0.60 to 0.80. Combining results from the geospatial multiyear models yielded maps of "core" breeding areas (areas with high probability values for all three years) surrounded by areas that scored high for only one or two years, providing an estimate of variability among years. Such information can highlight landscape options for amphibian conservation. For example, our models identify alternative areas that could be protected for each species

  17. Population viability of Arctic grayling in the Gibbon River, Yellowstone National Park

    Science.gov (United States)

    Steed, Amber C.; Zale, Alexander V.; Koel, Todd M.; Kalinowski, Steven T.

    2010-01-01

    The fluvial Arctic grayling Thymallus arcticus is restricted to less than 5% of its native range in the contiguous United States and was relisted as a category 3 candidate species under the U.S. Endangered Species Act in 2010. Although fluvial Arctic grayling of the lower Gibbon River, Yellowstone National Park, Wyoming, were considered to have been extirpated by 1935, anglers and biologists have continued to report catching low numbers of Arctic grayling in the river. Our goal was to determine whether a viable population of fluvial Arctic grayling persisted in the Gibbon River or whether the fish caught in the river were downstream emigrants from lacustrine populations in headwater lakes. We addressed this goal by determining relative abundances, sources, and evidence for successful spawning of Arctic grayling in the Gibbon River. During 2005 and 2006, Arctic grayling comprised between 0% and 3% of the salmonid catch in riverwide electrofishing (mean Back-calculated lengths at most ages were similar among all fish, and successful spawning within the Gibbon River below the headwater lakes was not documented. Few Arctic grayling adults and no fry were detected in the Gibbon River, implying that a reproducing fluvial population does not exist there. These findings have implications for future Endangered Species Act considerations and management of fluvial Arctic grayling within and outside of Yellowstone National Park. Our comprehensive approach is broadly applicable to the management of sparsely detected aquatic species worldwide.

  18. Phenotypic and genetic differentiation among yellow monkeyflower populations from thermal and non-thermal soils in Yellowstone National Park.

    Science.gov (United States)

    Lekberg, Ylva; Roskilly, Beth; Hendrick, Margaret F; Zabinski, Catherine A; Barr, Camille M; Fishman, Lila

    2012-09-01

    In flowering plants, soil heterogeneity can generate divergent natural selection over fine spatial scales, and thus promote local adaptation in the absence of geographic barriers to gene flow. Here, we investigate phenotypic and genetic differentiation in one of the few flowering plants that thrives in both geothermal and non-thermal soils in Yellowstone National Park (YNP). Yellow monkeyflowers (Mimulus guttatus) growing at two geothermal ("thermal") sites in YNP were distinct in growth form and phenology from paired populations growing nearby ( 0.34), which were only weakly differentiated from each other (all F (ST) geothermal gradient in Yellowstone.

  19. Vegetation monitoring to detect and predict vegetation change: Connecting historical and future shrub/steppe data in Yellowstone National Park

    Science.gov (United States)

    Geneva Chong; David Barnett; Benjamin Chemel; Roy Renkin; Pamela Sikkink

    2011-01-01

    A 2002 National Research Council (NRC) evaluation of ungulate management practices in Yellowstone specifically concluded that previous (1957 to present) vegetation monitoring efforts were insufficient to determine whether climate or ungulates were more influential on shrub/steppe dynamics on the northern ungulate winter range. The NRC further recommended that the...

  20. Chemical analyses of waters from geysers, hot springs, and pools in Yellowstone National Park, Wyoming from 1974 to 1978

    Energy Technology Data Exchange (ETDEWEB)

    Thompson, J.M.; Yadav, S.

    1979-01-01

    Waters from geysers, hot springs, and pools of Yellowstone National Park have been analyzed. We report 422 complete major ion analyses from 330 different locations of geysers, hot springs, and pools, collected from 1974 to 1978. Many of the analyses from Upper, Midway, Lower, and Norris Geyser Basin are recollections of features previously reported.

  1. Multi-method, multi-scale geophysical observations in the Obsidian Pool Thermal Area, Yellowstone National Park

    Science.gov (United States)

    Holbrook, W. S.; Carr, B.; Pasquet, S.; Sims, K. W. W.; Dickey, K.

    2016-12-01

    Despite the prominence of Yellowstone as the world's most active hydrothermal province, relatively little is known about the plumbing systems that link deeper hydrothermal fluids to the charismatic hot springs, geysers and mud pots at the surface. We present the results of a multi-method, multi-scale geophysical investigation of the Obsidian Pool Thermal Area (OPTA) in Yellowstone National Park. OPTA hosts acid-sulfate hot springs and mud pots with relatively low pH. We present the results of seismic refraction, electrical resistivity, time-domain EM (TEM), soil conductivity meter (EMI), and GPR data acquired in July 2016. There is a strong contrast in physical properties in the upper 50 m of the subsurface between the low-lying hydrothermal area and surrounding hills: the hydrothermal area has much lower seismic velocities ( 1 km/s vs 3 km/s) and electrical resistivity ( 20 ohm-m vs 300 ohm-m). A prominent zone of very low resistivity (<10 ohm-m) exists at about 20 m depth beneath all hydrothermal features. Poisson's ratio, calculated from P-wave refraction tomography and surface wave inversions, shows low values beneath the "frying pan," where gas is emerging in small fumaroles, suggesting that Poisson's ratio is an effective "gas detector" in hydrothermal areas. Near-surface resistivity mapped from EMI shows a strong correlation with hydrothermal areas previously mapped by heat flow, with areas of high heat flow generally having low resistivity near the surface. Two exceptions are (1) the "frying pan," which shows a central area of high resistivity (corresponding to escaping gas) surrounding by a halo of low resistivity, and (2) a broad area of low resistivity connecting the hydrothermal centers to the lake, which may be clay deposits. TEM data penetrate up to 200 m in depth and suggest that a reservoir of hydrothermal fluids may underlie the entire area, including beneath the forested hills, at depths greater than 100 m, but that they rise toward the surface in

  2. Carbon fluxes in ecosystems of Yellowstone National Park predicted from remote sensing data and simulation modeling.

    Science.gov (United States)

    Potter, Christopher; Klooster, Steven; Crabtree, Robert; Huang, Shengli; Gross, Peggy; Genovese, Vanessa

    2011-08-11

    A simulation model based on remote sensing data for spatial vegetation properties has been used to estimate ecosystem carbon fluxes across Yellowstone National Park (YNP). The CASA (Carnegie Ames Stanford Approach) model was applied at a regional scale to estimate seasonal and annual carbon fluxes as net primary production (NPP) and soil respiration components. Predicted net ecosystem production (NEP) flux of CO2 is estimated from the model for carbon sinks and sources over multi-year periods that varied in climate and (wildfire) disturbance histories. Monthly Enhanced Vegetation Index (EVI) image coverages from the NASA Moderate Resolution Imaging Spectroradiometer (MODIS) instrument (from 2000 to 2006) were direct inputs to the model. New map products have been added to CASA from airborne remote sensing of coarse woody debris (CWD) in areas burned by wildfires over the past two decades. Model results indicated that relatively cooler and wetter summer growing seasons were the most favorable for annual plant production and net ecosystem carbon gains in representative landscapes of YNP. When summed across vegetation class areas, the predominance of evergreen forest and shrubland (sagebrush) cover was evident, with these two classes together accounting for 88% of the total annual NPP flux of 2.5 Tg C yr-1 (1 Tg = 1012 g) for the entire Yellowstone study area from 2000-2006. Most vegetation classes were estimated as net ecosystem sinks of atmospheric CO2 on annual basis, making the entire study area a moderate net sink of about +0.13 Tg C yr-1. This average sink value for forested lands nonetheless masks the contribution of areas burned during the 1988 wildfires, which were estimated as net sources of CO2 to the atmosphere, totaling to a NEP flux of -0.04 Tg C yr-1 for the entire burned area. Several areas burned in the 1988 wildfires were estimated to be among the lowest in overall yearly NPP, namely the Hellroaring Fire, Mink Fire, and Falls Fire areas. Rates of

  3. Eruptions at Lone Star Geyser, Yellowstone National Park, USA, part 1: energetics and eruption dynamics

    Science.gov (United States)

    Karlstrom, Leif; Hurwitz, Shaul; Sohn, Robert; Vandemeulebrouck, Jean; Murphy, Fred; Rudolph, Maxwell L.; Johnston, Malcolm J.S.; Manga, Michael; McCleskey, R. Blaine

    2013-01-01

    Geysers provide a natural laboratory to study multiphase eruptive processes. We present results from a four–day experiment at Lone Star Geyser in Yellowstone National Park, USA. We simultaneously measured water discharge, acoustic emissions, infraredintensity, and visible and infrared video to quantify the energetics and dynamics of eruptions, occurring approximately every three hours. We define four phases in the eruption cycle: 1) a 28 ± 3 minute phase with liquid and steam fountaining, with maximum jet velocities of 16–28 m s− 1, steam mass fraction of less than ∼ 0.01. Intermittently choked flow and flow oscillations with periods increasing from 20 to 40 s are coincident with a decrease in jet velocity and an increase of steam fraction; 2) a 26 ± 8 minute post–eruption relaxation phase with no discharge from the vent, infrared (IR) and acoustic power oscillations gliding between 30 and 40 s; 3) a 59 ± 13 minute recharge period during which the geyser is quiescent and progressively refills, and 4) a 69 ± 14 minute pre–play period characterized by a series of 5–10 minute–long pulses of steam, small volumes of liquid water discharge and 50–70 s flow oscillations. The erupted waters ascend froma 160 − 170° C reservoir and the volume discharged during the entire eruptive cycle is 20.8 ± 4.1 m3. Assuming isentropic expansion, we calculate a heat output from the geyser of 1.4–1.5 MW, which is < 0.1% of the total heat output from Yellowstone Caldera.

  4. Population Dynamics of Wolves and Coyotes at Yellowstone National Park: Modeling Interference Competition with an Infectious Disease

    OpenAIRE

    Blanco, Krystal; Barley, Kamal; Mubayi, Anuj

    2014-01-01

    Gray wolves were reintroduced to Yellowstone National Park (YNP) in 1995. The population initially flourished, but since 2003 the population has experience significant reductions due to factors that may include disease-induced mortality, illegal hunting, park control pro- grams, vehicle induced deaths and intra-species aggression. Despite facing similar conditions, and interference competition with the wolves, the coyote population at YNP has persisted. In this paper we introduce an epidemiol...

  5. Geoarchaeota: a new candidate phylum in the Archaea from high-temperature acidic iron mats in Yellowstone National Park

    OpenAIRE

    Kozubal, Mark A; Romine, Margaret; Jennings, Ryan deM; Jay, Zack J; Tringe, Susannah G; Rusch, Doug B; Beam, Jacob P; McCue, Lee Ann; Inskeep, William P

    2012-01-01

    Geothermal systems in Yellowstone National Park (YNP) provide an outstanding opportunity to understand the origin and evolution of metabolic processes necessary for life in extreme environments including low pH, high temperature, low oxygen and elevated concentrations of reduced iron. Previous phylogenetic studies of acidic ferric iron mats from YNP have revealed considerable diversity of uncultivated and undescribed archaea. The goal of this study was to obtain replicate de novo genome assem...

  6. 76 FR 27087 - Draft Environmental Impact Statement for the Winter Use Plan, Yellowstone National Park

    Science.gov (United States)

    2011-05-10

    ... one of several methods. Internet: We encourage you to comment via the Internet at http://parkplanning... regarding Yellowstone in the winter, including educational materials and a detailed history of winter use in...

  7. Molecular Studies of Filamentous and Biofilm-Forming Hyperthermophilic Communities in Yellowstone National Park

    Science.gov (United States)

    Summons, R. E.; Meyer-Dombard, D. R.; Bradley, A. S.; Dibbell, A. K.; Fredricks, H. F.; Hinrichs, K.; Jahnke, L. L.; Shock, E.; Amend, J. P.

    2005-12-01

    The Aquificales, the most deeply-branching order of Bacteria in the phylogenetic tree of life, comprises eight recognized thermophilic genera, including Aquifex, Hydrogenobacter, and Thermocrinis. The common metabolism for these Bacteria, when grown in culture, is the oxidation of hydrogen with molecular oxygen (Knallgas reaction). Aquificales have been identified by molecular techniques (16S rRNA gene surveys, fluorescent in situ hybridization) in Yellowstone National Park (YNP), sea vent chimneys and fluids, and many other terrestrial and marine locations. In situ, Aquificales can reside as biofilms on vent sinters but they also commonly form filamentous communities, otherwise known as pink streamers, which attach to solid substrates. Initial 16S rRNA gene surveys conducted on streamer communities from Octopus Spring YNP indicated that these were low diversity ecosystems dominated by a few phylotypes including Thermocrinis sp., Thermotoga sp. and one other bacterial clade (Reysenbach et al 1994). Archaea were notable for their absence. In one of the first geobiological studies of pink streamers and vent biofilms in Yellowstone National Park, Jahnke and coworkers (2001) used classical lipidological techniques to compare Aquificales cultures with environmental samples to show that YNP pink filaments were more phylogenetically diverse and physiologically more complex than the early genomic studies indicated. The presence of archaeol, the range and structures of other lipids and a wide dispersion in the carbon isotopic signatures of biomass and individual lipids (-15 to -27%) showed that Archaea were present in pink filament communities and that there was, at least, one additional bacterial group besides the dominant Aquificales component. New molecular studies that comprise analyses of 16S rRNA genes and total lipid extracts by liquid chromatography and mass spectrometry and chemical degradation with gas chromatography and mass spectrometry now show that Crenarchaea

  8. Sulfur geochemistry of hydrothermal waters in Yellowstone National Park: IV Acid-sulfate waters

    Science.gov (United States)

    Nordstrom, D. Kirk; McCleskey, R. Blaine; Ball, J.W.

    2009-01-01

    Many waters sampled in Yellowstone National Park, both high-temperature (30-94 ??C) and low-temperature (0-30 ??C), are acid-sulfate type with pH values of 1-5. Sulfuric acid is the dominant component, especially as pH values decrease below 3, and it forms from the oxidation of elemental S whose origin is H2S in hot gases derived from boiling of hydrothermal waters at depth. Four determinations of pH were obtained: (1) field pH at field temperature, (2) laboratory pH at laboratory temperature, (3) pH based on acidity titration, and (4) pH based on charge imbalance (at both laboratory and field temperatures). Laboratory pH, charge imbalance pH (at laboratory temperature), and acidity pH were in close agreement for pH ??10%, a selection process was used to compare acidity, laboratory, and charge balance pH to arrive at the best estimate. Differences between laboratory and field pH can be explained based on Fe oxidation, H2S or S2O3 oxidation, CO2 degassing, and the temperature-dependence of pK2 for H2SO4. Charge imbalances are shown to be dependent on a speciation model for pH values 350 mg/L Cl) decrease as the Cl- concentration increases from boiling which appears inconsistent with the hypothesis of H2S oxidation as a source of hydrothermal SO4. This trend is consistent with the alternate hypothesis of anhydrite solubility equilibrium. Acid-sulfate water analyses are occasionally high in As, Hg, and NH3 concentrations but in contrast to acid mine waters they are low to below detection in Cu, Zn, Cd, and Pb concentrations. Even concentrations of SO4, Fe, and Al are much lower in thermal waters than acid mine waters of the same pH. This difference in water chemistry may explain why certain species of fly larvae live comfortably in Yellowstone's acid waters but have not been observed in acid rock drainage of the same pH.

  9. Group composition effects on aggressive interpack interactions of gray wolves in Yellowstone National Park

    Science.gov (United States)

    Cassidy, Kira A.; MacNulty, Daniel R.; Stahler, Daniel R.; Smith, Douglas W.; Mech, L. David

    2015-01-01

    Knowledge of characteristics that promote group success during intraspecific encounters is key to understanding the adaptive advantages of sociality for many group-living species. In addition, some individuals in a group may be more likely than others to influence intergroup conflicts, a relatively neglected idea in research on social animals. Here we use observations of aggressive interactions between wolf (Canis lupus) packs over an extended period and use pack characteristics to determine which groups had an advantage over their opponents. During 16 years of observation in Yellowstone National Park from 1995 to 2010, we documented 121 interpack aggressive interactions. We recorded pack sizes, compositions, and spatial orientation related to residency to determine their effects on the outcomes of interactions between packs. Relative pack size (RPS) improved the odds of a pack displacing its opponent. However, pack composition moderated the effect of RPS as packs with relatively more old members (>6.0 years old) or adult males had higher odds of winning despite a numerical disadvantage. The location of the interaction with respect to pack territories had no effect on the outcome of interpack interactions. Although the importance of RPS in successful territorial defense suggests the evolution and maintenance of group living may be at least partly due to larger packs’ success during interpack interactions, group composition is also an important factor, highlighting that some individuals are more valuable than others during interpack conflicts.

  10. Interactions between wolves and female grizzly bears with cubs in Yellowstone National Park

    Science.gov (United States)

    Gunther, Kerry A.; Smith, Douglas W.

    2004-01-01

    Gray wolves (Canis lupus) were extirpated from Yellowstone National Park (YNP) by the 1920s through predator control actions (Murie 1940,Young and Goldman 1944, Weaver 1978), then reintroduced into the park from 1995 to 1996 to restore ecological integrity and adhere to legal mandates (Bangs and Fritts 1996, Phillips and Smith 1996, Smith et al. 2000). Prior to reintroduction, the potential effects of wolves on the region’s threatened grizzly bear (Ursus arctos) population were evaluated (Servheen and Knight 1993). In areas where wolves and grizzly bears are sympatric, interspecific killing by both species occasionally occurs (Ballard 1980, 1982; Hayes and Baer 1992). Most agonistic interactions between wolves and grizzly bears involve defense of young or competition for carcasses (Murie 1944, 1981; Ballard 1982; Hornbeck and Horejsi 1986; Hayes and Mossop 1987; Kehoe 1995; McNulty et al. 2001). Servheen and Knight (1993) predicted that reintroduced wolves could reduce the frequency of winter-killed and disease-killed ungulates available for bears to scavenge, and that grizzly bears would occasionally usurp wolf-killed ungulate carcasses. Servheen and Knight (1993) hypothesized that interspecific killing and competition for carcasses would have little or no population level effect on either species.

  11. Bottom-up factors influencing riparian willow recovery in Yellowstone National Park

    Science.gov (United States)

    Tercek, M.T.; Stottlemyer, R.; Renkin, R.

    2010-01-01

    After the elimination of wolves (Canis lupis L.) in the 1920s, woody riparian plant communities on the northern range of Yellowstone National Park (YNP) declined an estimated 50%. After the reintroduction of wolves in 19951996, riparian willows (Salix spp.) on YNP's northern range showed significant growth for the first time since the 1920s. However, the pace of willow recovery has not been uniform. Some communities have exceeded 400 cm, while others are still at pre-1995 levels of 250 cm max. height) willow sites where willows had escaped elk (Cervus elaphus L.) browsing with "short" willow sites that could still be browsed. Unlike studies that manipulated willow height with fences and artificial dams, we examined sites that had natural growth differences in height since the reintroduction of wolves. Tall willow sites had greater water availability, more-rapid net soil nitrogen mineralization, greater snow depth, lower soil respiration rates, and cooler summer soil temperatures than nearby short willow sites. Most of these differences were measured both in herbaceous areas adjacent to the willow patches and in the willow patches themselves, suggesting that they were not effects of varying willow height recovery but were instead preexisting site differences that may have contributed to increased plant productivity. Our results agree with earlier studies in experimental plots which suggest that the varying pace of willow recovery has been influenced by abiotic limiting factors that interact with top-down reductions in willow browsing by elk. ?? 2010 Western North American Naturalist.

  12. Archaeal and bacterial community analysis of several Yellowstone National Park hot springs

    Science.gov (United States)

    Colman, D. R.; Takacs-Vesbach, C. D.

    2012-12-01

    The hot springs of Yellowstone National Park (YNP) are home to a diverse assemblage of microorganisms. Culture-independent studies have significantly expanded our understanding of the diversity of both Bacteria and Archaea present in YNP springs as well as the geochemical and ecological controls on communities. While the ecological analysis of Bacteria among the physicochemically heterogenous springs of YNP has been previously conducted, less is known about the extent of diversity of Archaeal communities and the chemical and ecological controls on their populations. Here we report a culture-independent analysis of 31 hot spring archaeal and bacterial communities of YNP springs using next generation sequencing. We found the phylogenetic diversity of Archaea to be generally comparable to that of co-occurring bacterial communities although overall, in the springs we investigated, diversity was higher for Bacteria than Archaea. Chemical and physical controls were similar for both domains with pH correlating most strongly with community composition. Community differences reflected the partitioning of taxonomic groups in low or high pH springs for both domains. Results will be discussed in a geochemical and ecological context.

  13. Research Coordination Network: Geothermal Biology and Geochemistry in Yellowstone National Park

    Science.gov (United States)

    Inskeep, W. P.; Young, M. J.; Jay, Z.

    2006-12-01

    The number and diversity of geothermal features in Yellowstone National Park (YNP) represent a fascinating array of high temperature geochemical environments that host a corresponding number of unique and potentially novel organisms in all of the three recognized domains of life: Bacteria, Archaea and Eukarya. The geothermal features of YNP have long been the subject of scientific inquiry, especially in the fields of microbiology, geochemistry, geothermal hydrology, microbial ecology, and population biology. However, there are no organized forums for scientists working in YNP geothermal areas to present research results, exchange ideas, discuss research priorities, and enhance synergism among research groups. The primary goal of the YNP Research Coordination Network (GEOTHERM) is to develop a more unified effort among scientists and resource agencies to characterize, describe, understand and inventory the diverse biota associated with geothermal habitats in YNP. The YNP RCN commenced in January 2005 as a collaborative effort among numerous university scientists, governmental agencies and private industry. The YNP RCN hosted a workshop in February 2006 to discuss research results and to form three working groups focused on (i) web-site and digital library content, (ii) metagenomics of thermophilic microbial communities and (iii) development of geochemical methods appropriate for geomicrobiological studies. The working groups represent one strategy for enhancing communication, collaboration and most importantly, productivity among the RCN participants. If you have an interest in the geomicrobiology of geothermal systems, please feel welcome to join and or participate in the YNP RCN.

  14. Heat‐tolerant Flowering Plants of Active Geothermal Areas in Yellowstone National Park

    Science.gov (United States)

    STOUT, RICHARD G.; AL‐NIEMI, THAMIR S.

    2002-01-01

    A broad survey of most of the major geyser basins within Yellowstone National Park (Wyoming, USA) was conducted to identify the flowering plants which tolerate high rhizosphere temperatures (≥40 °C) in geothermally heated environments. Under such conditions, five species of monocots and four species of dicots were repeatedly found. The predominant flowering plants in hot soils (>40 °C at 2–5 cm depth) were grasses, primarily Dichanthelium lanuginosum. Long‐term (weeks to months) rhizosphere temperatures of individual D. lanuginosum above 40 °C were recorded at several different locations, both in the summer and winter. The potential role of heat shock proteins (HSPs) in the apparent adaptation of these plants to chronically high rhizosphere temperatures was examined. Antibodies to cytoplasmic class I small heat shock proteins (sHSPs) and to HSP101 were used in Western immunoblot analyses of protein extracts from plants collected from geothermally heated soils. Relatively high levels of proteins reacting with anti‐sHSP antibodies were consistently detected in root extracts from plants experiencing rhizosphere temperatures above 40 °C, though these proteins were usually not highly expressed in leaf extracts from the same plants. Proteins reacting with antibodies to HSP101 were also present both in leaf and root extracts from plants collected from geothermal soils, but their levels of expression were not as closely related to the degree of heat exposure as those of sHSPs. PMID:12197524

  15. Methylmercury enters an aquatic food web through acidophilic microbial mats in Yellowstone National Park, Wyoming.

    Science.gov (United States)

    Boyd, Eric S; King, Susan; Tomberlin, Jeffery K; Nordstrom, D Kirk; Krabbenhoft, David P; Barkay, Tamar; Geesey, Gill G

    2009-04-01

    Microbial mats are a visible and abundant life form inhabiting the extreme environments in Yellowstone National Park (YNP), WY, USA. Little is known of their role in food webs that exist in the Park's geothermal habitats. Eukaryotic green algae associated with a phototrophic green/purple Zygogonium microbial mat community that inhabits low-temperature regions of acidic (pH approximately 3.0) thermal springs were found to serve as a food source for stratiomyid (Diptera: Stratiomyidae) larvae. Mercury in spring source water was taken up and concentrated by the mat biomass. Monomethylmercury compounds (MeHg(+)), while undetectable or near the detection limit (0.025 ng l(-1)) in the source water of the springs, was present at concentrations of 4-7 ng g(-1) dry weight of mat biomass. Detection of MeHg(+) in tracheal tissue of larvae grazing the mat suggests that MeHg(+) enters this geothermal food web through the phototrophic microbial mat community. The concentration of MeHg(+) was two to five times higher in larval tissue than mat biomass indicating MeHg(+) biomagnification occurred between primary producer and primary consumer trophic levels. The Zygogonium mat community and stratiomyid larvae may also play a role in the transfer of MeHg(+) to species in the food web whose range extends beyond a particular geothermal feature of YNP.

  16. Eruptions at Lone Star geyser, Yellowstone National Park, USA: 2. Constraints on subsurface dynamics

    Science.gov (United States)

    Vandemeulebrouck, Jean; Sohn, Robert A.; Rudolph, Maxwell L.; Hurwitz, Shaul; Manga, Michael; Johnston, Malcolm J.S.; Soule, S. Adam; McPhee, Darcy K.; Glen, Jonathan M.G.; Karlstrom, Leif; Murphy, Fred

    2014-01-01

    We use seismic, tilt, lidar, thermal, and gravity data from 32 consecutive eruption cycles of Lone Star geyser in Yellowstone National Park to identify key subsurface processes throughout the geyser's eruption cycle. Previously, we described measurements and analyses associated with the geyser's erupting jet dynamics. Here we show that seismicity is dominated by hydrothermal tremor (~5–40 Hz) attributed to the nucleation and/or collapse of vapor bubbles. Water discharge during eruption preplay triggers high-amplitude tremor pulses from a back azimuth aligned with the geyser cone, but during the rest of the eruption cycle it is shifted to the east-northeast. Moreover, ~4 min period ground surface displacements recur every 26 ± 8 min and are uncorrelated with the eruption cycle. Based on these observations, we conclude that (1) the dynamical behavior of the geyser is controlled by the thermo-mechanical coupling between the geyser conduit and a laterally offset reservoir periodically filled with a highly compressible two-phase mixture, (2) liquid and steam slugs periodically ascend into the shallow crust near the geyser system inducing detectable deformation, (3) eruptions occur when the pressure decrease associated with overflow from geyser conduit during preplay triggers an unstable feedback between vapor generation (cavitation) and mass discharge, and (4) flow choking at a constriction in the conduit arrests the runaway process and increases the saturated vapor pressure in the reservoir by a factor of ~10 during eruptions.

  17. Chemical analyses of hot springs, pools, geysers, and surface waters from Yellowstone National Park, Wyoming, and vicinity, 1974-1975

    Science.gov (United States)

    Ball, James W.; Nordstrom, D. Kirk; Jenne, Everett A.; Vivit, Davison V.

    1998-01-01

    This report presents all analytical determinations for samples collected from Yellowstone National Park and vicinity during 1974 and 1975. Water temperature, pH, Eh, and dissolved O2 were determined on-site. Total alkalinity and F were determined on the day of sample collection. Flame atomic-absorption spectrometry was used to determine concentrations of Li, Na, K, Ca, and Mg. Ultraviolet/visible spectrophotometry was used to determine concentrations of Fe(II), Fe(III), As(III), and As(V). Direct-current plasma-optical-emission spectrometry was used to determine the concentrations of B, Ba, Cd, Cs, Cu, Mn, Ni, Pb, Rb, Sr, and Zn. Two samples collected from Yellowstone Park in June 1974 were used as reference samples for testing the plasma analytical method. Results of these tests demonstrate acceptable precision for all detectable elements. Charge imbalance calculations revealed a small number of samples that may have been subject to measurement errors in pH or alkalinity. These data represent some of the most complete analyses of Yellowstone waters available.

  18. Direct Measurement of the Volume of Liquid Water Emitted During Eruptions of Lone Star Geyser, Yellowstone National Park, Wyoming

    Science.gov (United States)

    Murphy, F.; Hurwitz, S.; Johnston, M. J.; Vandemeulebrouck, J.; Pontbriand, C.; Sohn, R. A.; Karlstrom, L.; Rudolph, M. L.

    2011-12-01

    In September, 2010 a comprehensive series of instrumental observations was carried out at Lone Star Geyser in Yellowstone National Park to measure changes in the geyser and its surroundings during eruptions. That project included measurements of flow in the streams that drain the geyser area. Three small streams convey liquid water from the geyser and many of the surrounding hot springs to the Firehole River, about 75 m south of the geyser cone. We developed rating curves for two of these streams by measuring channel cross-sections and timing floating markers (using stopwatches and video recordings) while simultaneously recording stream depth at two-second intervals at two locations using pressure transducers and dataloggers. We estimated the flow in the third (ungaged) stream to be 0.15 of the flow in the easternmost stream, with which it shares a source area and part of its channel. The eruption cycle takes about 3 hours, and a total of nine eruption cycles were observed. During these 3-hour cycles the geyser and the nearby hot springs deliver a total of between 15 and 28 m3 of water to the Firehole River. During the 10-20 minutes of the main phase of an eruption, the geyser delivered between 8 and 11 m3 of water to the three streams. The volume of water emitted during eruptions appears to display a significant diurnal variation which strongly correlates with air temperature, with significantly more flow during early afternoon hours. There were also significant variations in the distribution of flow between the different channels. Our calculations suggest that losses due to evaporation along the flow channels are negligible, and losses due to infiltration appear to be small. The calculated volumes of water discharge do not account for the volume of erupted steam or evaporation of liquid water from the jet. Steam discharge will be assessed using image analysis of high speed video. The calculated volumes provide accurate and important constraint for models of

  19. Hydrogeology of the Old Faithful area, Yellowstone National Park, Wyoming, and its relevance to natural resources and infrastructure

    Science.gov (United States)

    ,; Foley, Duncan; Fournier, Robert O.; Heasler, Henry P.; Hinckley, Bern; Ingebritsen, Steven E.; Lowenstern, Jacob B.; Susong, David D.

    2014-01-01

    A panel of leading experts (The Old Faithful Science Review Panel) was convened by Yellowstone National Park (YNP) to review and summarize the geological and hydrological understanding that can inform National Park Service management of the Upper Geyser Basin area. We give an overview of present geological and hydrological knowledge of the Old Faithful hydrothermal (hot water) system and related thermal areas in the Upper Geyser Basin. We prioritize avenues for improving our understanding of key knowledge gaps that limit informed decision-making regarding human use in this fragile natural landscape. Lastly, we offer guidelines to minimize impacts to the hydrothermal system that could be used to aid decisions by park management.

  20. Use of ASTER and MODIS thermal infrared data to quantify heat flow and hydrothermal change at Yellowstone National Park

    Science.gov (United States)

    Vaughan, R. Greg; Keszthelyi, Laszlo P.; Lowenstern, Jacob B.; Jaworowski, Cheryl; Heasler, Henry

    2012-01-01

    The overarching aim of this study was to use satellite thermal infrared (TIR) remote sensing to monitor geothermal activity within the Yellowstone geothermal area to meet the missions of both the U.S. Geological Survey and the Yellowstone National Park Geology Program. Specific goals were to: 1) address the challenges of monitoring the surface thermal characteristics of the > 10,000 spatially and temporally dynamic thermal features in the Park (including hot springs, pools, geysers, fumaroles, and mud pots) that are spread out over ~ 5000 km2, by using satellite TIR remote sensing tools (e.g., ASTER and MODIS), 2) to estimate the radiant geothermal heat flux (GHF) for Yellowstone's thermal areas, and 3) to identify normal, background thermal changes so that significant, abnormal changes can be recognized, should they ever occur (e.g., changes related to tectonic, hydrothermal, impending volcanic processes, or human activities, such as nearby geothermal development). ASTER TIR data (90-m pixels) were used to estimate the radiant GHF from all of Yellowstone's thermal features and update maps of thermal areas. MODIS TIR data (1-km pixels) were used to record background thermal radiance variations from March 2000 through December 2010 and establish thermal change detection limits. A lower limit for the radiant GHF estimated from ASTER TIR temperature data was established at ~ 2.0 GW, which is ~ 30–45% of the heat flux estimated through geochemical thermometry. Also, about 5 km2 of thermal areas was added to the geodatabase of mapped thermal areas. A decade-long time-series of MODIS TIR radiance data was dominated by seasonal cycles. A background subtraction technique was used in an attempt to isolate variations due to geothermal changes. Several statistically significant perturbations were noted in the time-series from Norris Geyser Basin, however many of these did not correspond to documented thermal disturbances. This study provides concrete examples of the

  1. Wolverine in Greater Yellowstone

    Science.gov (United States)

    Kerry Murphy; Jason Wilmot; Jeff Copeland; Dan Tyers; John. Squires

    2011-01-01

    The wolverine is one of the least studied carnivores in North America, particularly in the contiguous United States where it occurs at the southern extent of its range. This project documented the distribution of wolverines in the eastern portion of Yellowstone National Park and adjoining areas of national forest and their population characteristics, habitat...

  2. Triggering and modulation of geyser eruptions in Yellowstone National Park by earthquakes, earth tides, and weather

    Science.gov (United States)

    Hurwitz, Shaul; Sohn, Robert A.; Luttrell, Karen; Manga, Michael

    2014-01-01

    We analyze intervals between eruptions (IBEs) data acquired between 2001 and 2011 at Daisy and Old Faithful geysers in Yellowstone National Park. We focus our statistical analysis on the response of these geysers to stress perturbations from within the solid earth (earthquakes and earth tides) and from weather (air pressure and temperature, precipitation, and wind). We conclude that (1) the IBEs of these geysers are insensitive to periodic stresses induced by solid earth tides and barometric pressure variations; (2) Daisy (pool geyser) IBEs lengthen by evaporation and heat loss in response to large wind storms and cold air; and (3) Old Faithful (cone geyser) IBEs are not modulated by air temperature and pressure variations, wind, and precipitation, suggesting that the subsurface water column is decoupled from the atmosphere. Dynamic stress changes of 0.1−0.2 MPa resulting from the 2002 M-7.9 Denali, Alaska, earthquake surface waves caused a statistically significant shortening of Daisy geyser's IBEs. Stresses induced by other large global earthquakes during the study period were at least an order of magnitude smaller. In contrast, dynamic stresses of >0.5 MPa from three large regional earthquakes in 1959, 1975, and 1983 caused lengthening of Old Faithful's IBEs. We infer that most subannual geyser IBE variability is dominated by internal processes and interaction with other geysers. The results of this study provide quantitative bounds on the sensitivity of hydrothermal systems to external stress perturbations and have implications for studying the triggering and modulation of volcanic eruptions by external forces.

  3. A serological survey of infectious disease in Yellowstone National Park's canid community.

    Science.gov (United States)

    Almberg, Emily S; Mech, L David; Smith, Douglas W; Sheldon, Jennifer W; Crabtree, Robert L

    2009-09-16

    Gray wolves (Canis lupus) were reintroduced into Yellowstone National Park (YNP) after a >70 year absence, and as part of recovery efforts, the population has been closely monitored. In 1999 and 2005, pup survival was significantly reduced, suggestive of disease outbreaks. We analyzed sympatric wolf, coyote (Canis latrans), and red fox (Vulpes vulpes) serologic data from YNP, spanning 1991-2007, to identify long-term patterns of pathogen exposure, identify associated risk factors, and examine evidence for disease-induced mortality among wolves for which there were survival data. We found high, constant exposure to canine parvovirus (wolf seroprevalence: 100%; coyote: 94%), canine adenovirus-1 (wolf pups [0.5-0.9 yr]: 91%, adults [>or=1 yr]: 96%; coyote juveniles [0.5-1.5 yrs]: 18%, adults [>or=1.6 yrs]: 83%), and canine herpesvirus (wolf: 87%; coyote juveniles: 23%, young adults [1.6-4.9 yrs]: 51%, old adults [>or=5 yrs]: 87%) suggesting that these pathogens were enzootic within YNP wolves and coyotes. An average of 50% of wolves exhibited exposure to the protozoan parasite, Neospora caninum, although individuals' odds of exposure tended to increase with age and was temporally variable. Wolf, coyote, and fox exposure to canine distemper virus (CDV) was temporally variable, with evidence for distinct multi-host outbreaks in 1999 and 2005, and perhaps a smaller, isolated outbreak among wolves in the interior of YNP in 2002. The years of high wolf-pup mortality in 1999 and 2005 in the northern region of the park were correlated with peaks in CDV seroprevalence, suggesting that CDV contributed to the observed mortality. Of the pathogens we examined, none appear to jeopardize the long-term population of canids in YNP. However, CDV appears capable of causing short-term population declines. Additional information on how and where CDV is maintained and the frequency with which future epizootics might be expected might be useful for future management of the Northern Rocky

  4. Importance of dispersal and thermal environment for mycorrhizal communities: lessons from Yellowstone National Park.

    Science.gov (United States)

    Lekberg, Ylva; Meadow, James; Rohr, Jason R; Redecker, Dirk; Zabinski, Catherine A

    2011-06-01

    The relative importance of dispersal and niche restrictions remains a controversial topic in community ecology, especially for microorganisms that are often assumed to be ubiquitous. We investigated the impact of these factors for the community assembly of the root-symbiont arbuscular mycorrhizal fungi (AMF) by sampling roots from geothermal and nonthermal grasslands in Yellowstone National Park (YNP), followed by sequencing and RFLP of AMF ribosomal DNA. With the exception of an apparent generalist RFLP type closely related to Glomus intraradices, a distance-based redundancy analysis indicated that the AMF community composition correlated with soil pH or pH-driven changes in soil chemistry. This was unexpected, given the large differences in soil temperature and plant community composition between the geothermal and nonthermal grasslands. RFLP types were found in either the acidic geothermal grasslands or in the neutral to alkaline grasslands, one of which was geothermal. The direct effect of the soil chemical environment on the distribution of two AMF morphospecies isolated from acidic geothermal grasslands was supported in a controlled greenhouse experiment. Paraglomus occultum and Scutellospora pellucida were more beneficial to plants and formed significantly more spores when grown in acidic than in alkaline soil. Distance among grasslands, used as an estimate of dispersal limitations, was not a significant predictor of AMF community similarity within YNP, and most fungal taxa may be part of a metacommunity. The isolation of several viable AMF taxa from bison feces indicates that wide-ranging bison could be a vector for at least some RFLP types among grasslands within YNP. In support of classical niche theory and the Baas-Becking hypothesis, our results suggest that AMF are not limited by dispersal at the scale of YNP, but that the soil environment appears to be the primary factor affecting community composition and distribution.

  5. A Multi-Method Experiment to Investigate Geyser Dynamics: Lone Star Geyser, Yellowstone National Park

    Science.gov (United States)

    Hurwitz, S.; Vandemeulebrouck, J.; Johnston, M. J.; Sohn, R. A.; Karlstrom, L.; Rudolph, M. L.; Murphy, F.; McPhee, D. K.; Glen, J. M.; Soule, S. A.; Pontbriand, C.; Meertens, C. M.

    2011-12-01

    Geysers are intermittently discharging hot springs that are driven by steam and non-condensable gas. They provide unique opportunities to study multiphase eruption processes and the geophysical signals they induce. In September 2010 we carried out a four-day experiment at Lone Star Geyser in Yellowstone National Park. The geyser is located about 5 km SSE of Old Faithful Geyser and 75 m north of the Upper Firehole River. Lone Star is a cone geyser that was selected for the experiment because it is isolated from other geysers, its eruptions are vigorous and voluminous, and its eruption intervals are relatively constant and predictable, occurring approximately every 3 hours. We made measurements during 32 eruption cycles using a suite of instruments including a broadband seismometer, 2 microphones, 5 platform tiltmeters, 3 collimating InfraRed sensors, 2 gravimeters, 2 self-potential sensors, 2 Light Detection And Ranging (LiDAR) scanners, a Forward Looking InfraRed (FLIR) camera, high-speed video cameras, and stream gauging. We also integrated meteorological data from nearby weather stations. The large dataset acquired during the experiment allows for the detection of a myriad of processes in the subsurface and in the erupting column at many different frequencies. The analyzed data yield new insights on multiphase eruptive processes that have implications for understanding self-organized, intermittent processes in nature that result from phase separation and localized input of energy and mass. The geophysical signals recorded during the experiment allow comparison with signals recorded in more complex volcanic systems where gas-driven and magma-driven processes are often hard to distinguish.

  6. Seeking a scientific approach to backcountry management in Yellowstone National Park

    Science.gov (United States)

    S. Thomas Olliff; Sue Consolo Murphy

    2000-01-01

    Three criteria are used to assess how Yellowstone’s wilderness managers incorporate science into management: preciousness, vulnerability and responsiveness to management. Four observations are proposed. First, where scientists lead, managers will follow. Scientists that leave the best trail will be followed most closely. Second, managers need to refocus efforts on...

  7. Public acceptance of management actions and judgments of responsibility for the wolves of the southern Greater Yellowstone Area: Report to Grand Teton National Park

    Science.gov (United States)

    Taylor, Jonathan G.; Johnson, S. Shea; Shelby, Lori B.

    2005-01-01

    Introduction Wolves of Grand Teton National Park and the Greater Yellowstone Area Gray wolves (Canis lupus) appeared in Grand Teton National Park (GRTE) in October of 1998, two years after being reintroduced to Yellowstone National Park (YNP). Since that time, five packs have been within the GRTE borders - Gros Ventre Pack, Nez Perce Pack, Yellowstone Delta Pack, Teton Pack, and Green River Pack (Table 1). Wolves in the Greater Yellowstone Area are increasing and spreading out geographically (USFWS and others, 2004). This dispersion was demonstrated recently by the death of a 2-year-old female wolf from the Swan Lake pack on I-70 in Colorado (June 7, 2004; http://mountain-prairie.USFWS.gov/pressrel /04-43.htm). The organization of wolf packs in the GYA is dynamic and highly structured. In 2003, for example, a wolf from the Teton Pack joined with the Green River Pack, and several young wolves left the Teton Pack and moved south (USFWS and others, 2004). Pack size (averaging five to ten members) is dependent on hunting efficiency, which depends on prey size, type, and density. Each pack defends home ranges of several hundred square miles. The social structure of the pack is based on a breeding pair (an alpha male and female). Other wolves in the pack can be categorized as betas (males and/or females second in rank to the alphas), subordinates, pups, and occasional omegas (outcasts). Because generally only the alpha pair breeds, subordinate wolves of reproductive age must disperse from their packs and form new associations in order to breed. (http://www.nps.gov/grte/wolf/biolo.htm). The reintroduced wolves are classified by the U.S. Fish and Wildlife Service (USFWS) as "nonessential experimental" under section 10(j) of the Endangered Species Act. The recovery criteria for the GYA wolves were met in 2002 for removing the wolves from the Endangered Species List (30 or more breeding pairs). Currently, the USFWS manages wolf populations in the GYA until delisting occurs

  8. Characterization of novel bacteriochlorophyll-a-containing red filaments from alkaline hot springs in Yellowstone National Park.

    Science.gov (United States)

    Boomer, S M; Pierson, B K; Austinhirst, R; Castenholz, R W

    2000-09-01

    Novel red, filamentous, gliding bacteria formed deep red layers in several alkaline hot springs in Yellowstone National Park. Filaments contained densely layered intracellular membranes and bacteriochlorophyll a. The in vivo absorption spectrum of the red layer filaments was distinct from other phototrophs, with unusual bacteriochlorophyll a signature peaks in the near-infrared (IR) region (807 nm and 911 nm). These absorption peaks were similar to the wavelengths penetrating to the red layer of the mats as measured with in situ spectroradiometry. The filaments also demonstrated maximal photosynthetic uptake of radiolabeled carbon sources at these wavelengths. The red layer filaments displayed anoxygenic photoheterotrophy, as evidenced by the specific incorporation of acetate, not bicarbonate, and by the absence of oxygen production. Photoheterotrophy was unaffected by sulfide and oxygen, but was diminished by high-intensity visible light. Near-IR radiation supported photoheterotrophy. Morphologically and spectrally similar filaments were observed in several springs in Yellowstone National Park, including Octopus Spring. Taken together, these data suggest that the red layer filaments are most similar to the photoheterotroph, Heliothrix oregonensis. Notable differences include mat position and coloration, absorption spectra, and prominent intracellular membranes.

  9. A Serological Survey of Infectious Disease in Yellowstone National Park’s Canid Community

    Science.gov (United States)

    Almberg, Emily S.; Mech, L. David; Smith, Douglas W.; Sheldon, Jennifer W.; Crabtree, Robert L.

    2009-01-01

    Background Gray wolves (Canis lupus) were reintroduced into Yellowstone National Park (YNP) after a >70 year absence, and as part of recovery efforts, the population has been closely monitored. In 1999 and 2005, pup survival was significantly reduced, suggestive of disease outbreaks. Methodology/Principal Findings We analyzed sympatric wolf, coyote (Canis latrans), and red fox (Vulpes vulpes) serologic data from YNP, spanning 1991–2007, to identify long-term patterns of pathogen exposure, identify associated risk factors, and examine evidence for disease-induced mortality among wolves for which there were survival data. We found high, constant exposure to canine parvovirus (wolf seroprevalence: 100%; coyote: 94%), canine adenovirus-1 (wolf pups [0.5–0.9 yr]: 91%, adults [≥1 yr]: 96%; coyote juveniles [0.5–1.5 yrs]: 18%, adults [≥1.6 yrs]: 83%), and canine herpesvirus (wolf: 87%; coyote juveniles: 23%, young adults [1.6–4.9 yrs]: 51%, old adults [≥5 yrs]: 87%) suggesting that these pathogens were enzootic within YNP wolves and coyotes. An average of 50% of wolves exhibited exposure to the protozoan parasite, Neospora caninum, although individuals’ odds of exposure tended to increase with age and was temporally variable. Wolf, coyote, and fox exposure to canine distemper virus (CDV) was temporally variable, with evidence for distinct multi-host outbreaks in 1999 and 2005, and perhaps a smaller, isolated outbreak among wolves in the interior of YNP in 2002. The years of high wolf-pup mortality in 1999 and 2005 in the northern region of the park were correlated with peaks in CDV seroprevalence, suggesting that CDV contributed to the observed mortality. Conclusions/Significance Of the pathogens we examined, none appear to jeopardize the long-term population of canids in YNP. However, CDV appears capable of causing short-term population declines. Additional information on how and where CDV is maintained and the frequency with which future epizootics might

  10. A serological survey of infectious disease in Yellowstone National Park's canid community.

    Directory of Open Access Journals (Sweden)

    Emily S Almberg

    Full Text Available BACKGROUND: Gray wolves (Canis lupus were reintroduced into Yellowstone National Park (YNP after a >70 year absence, and as part of recovery efforts, the population has been closely monitored. In 1999 and 2005, pup survival was significantly reduced, suggestive of disease outbreaks. METHODOLOGY/PRINCIPAL FINDINGS: We analyzed sympatric wolf, coyote (Canis latrans, and red fox (Vulpes vulpes serologic data from YNP, spanning 1991-2007, to identify long-term patterns of pathogen exposure, identify associated risk factors, and examine evidence for disease-induced mortality among wolves for which there were survival data. We found high, constant exposure to canine parvovirus (wolf seroprevalence: 100%; coyote: 94%, canine adenovirus-1 (wolf pups [0.5-0.9 yr]: 91%, adults [>or=1 yr]: 96%; coyote juveniles [0.5-1.5 yrs]: 18%, adults [>or=1.6 yrs]: 83%, and canine herpesvirus (wolf: 87%; coyote juveniles: 23%, young adults [1.6-4.9 yrs]: 51%, old adults [>or=5 yrs]: 87% suggesting that these pathogens were enzootic within YNP wolves and coyotes. An average of 50% of wolves exhibited exposure to the protozoan parasite, Neospora caninum, although individuals' odds of exposure tended to increase with age and was temporally variable. Wolf, coyote, and fox exposure to canine distemper virus (CDV was temporally variable, with evidence for distinct multi-host outbreaks in 1999 and 2005, and perhaps a smaller, isolated outbreak among wolves in the interior of YNP in 2002. The years of high wolf-pup mortality in 1999 and 2005 in the northern region of the park were correlated with peaks in CDV seroprevalence, suggesting that CDV contributed to the observed mortality. CONCLUSIONS/SIGNIFICANCE: Of the pathogens we examined, none appear to jeopardize the long-term population of canids in YNP. However, CDV appears capable of causing short-term population declines. Additional information on how and where CDV is maintained and the frequency with which future

  11. A serological survey of infectious disease in Yellowstone National Park's canid community

    Science.gov (United States)

    Almberg, E.S.; Mech, L.D.; Smith, D.W.; Sheldon, J.W.; Crabtree, R.L.

    2009-01-01

    Background: Gray wolves (Canis lupus) were reintroduced into Yellowstone National Park (YNP) after a >70 year absence, and as part of recovery efforts, the population has been closely monitored. In 1999 and 2005, pup survival was significantly reduced, suggestive of disease outbreaks. Methodology/Principal Findings: We analyzed sympatric wolf, coyote (Canis latrans), and red fox (Vulpes vulpes) serologic data from YNP, spanning 1991–2007, to identify long-term patterns of pathogen exposure, identify associated risk factors, and examine evidence for disease-induced mortality among wolves for which there were survival data. We found high, constant exposure to canine parvovirus (wolf seroprevalence: 100%; coyote: 94%), canine adenovirus-1 (wolf pups [0.5–0.9 yr]: 91%, adults [≥1 yr]: 96%; coyote juveniles [0.5–1.5 yrs]: 18%, adults [≥1.6 yrs]: 83%), and canine herpesvirus (wolf: 87%; coyote juveniles: 23%, young adults [1.6–4.9 yrs]: 51%, old adults [≥5 yrs]: 87%) suggesting that these pathogens were enzootic within YNP wolves and coyotes. An average of 50% of wolves exhibited exposure to the protozoan parasite, Neospora caninum, although individuals’ odds of exposure tended to increase with age and was temporally variable. Wolf, coyote, and fox exposure to canine distemper virus (CDV) was temporally variable, with evidence for distinct multi-host outbreaks in 1999 and 2005, and perhaps a smaller, isolated outbreak among wolves in the interior of YNP in 2002. The years of high wolf-pup mortality in 1999 and 2005 in the northern region of the park were correlated with peaks in CDV seroprevalence, suggesting that CDV contributed to the observed mortality. Conclusions/Significance: Of the pathogens we examined, none appear to jeopardize the long-term population of canids in YNP. However, CDV appears capable of causing short-term population declines. Additional information on how and where CDV is maintained and the frequency with which future epizootics

  12. Holocene seasonal variability inferred from multiple proxy records from Crevice Lake, Yellowstone National Park, USA

    Science.gov (United States)

    Whitlock, Cathy; Dean, Walter E.; Fritz, Sherilyn C.; Stevens, Lora R.; Stone, Jeffery R.; Power, Mitchell J.; Rosenbaum, Joseph R.; Pierce, Kenneth L.; Bracht-Flyr, Brandi B.

    2012-01-01

    A 9400-yr-old record from Crevice Lake, a semi-closed alkaline lake in northern Yellowstone National Park, was analyzed for pollen, charcoal, geochemistry, mineralogy, diatoms, and stable isotopes to develop a nuanced understanding of Holocene environmental history in a region of northern Rocky Mountains that receives both summer and winter precipitation. The limited surface area, conical bathymetry, and deep water (> 31 m) of Crevice Lake create oxygen-deficient conditions in the hypolimnion and preserve annually laminated sediment (varves) for much of the record. Pollen data indicate that the watershed supported a closed Pinus-dominated forest and low fire frequency prior to 8200 cal yr BP, followed by open parkland until 2600 cal yr BP, and open mixed-conifer forest thereafter. Fire activity shifted from infrequent stand-replacing fires initially to frequent surface fires in the middle Holocene and stand-replacing events in recent centuries. Low values of δ18O suggest high winter precipitation in the early Holocene, followed by steadily drier conditions after 8500 cal yr BP. Carbonate-rich sediments before 5000 cal yr BP imply warmer summer conditions than after 5000 cal yr BP. High values of molybdenum (Mo), uranium (U), and sulfur (S) indicate anoxic bottom-waters before 8000 cal yr BP, between 4400 and 3900 cal yr BP, and after 2400 cal yr BP. The diatom record indicates extensive water-column mixing in spring and early summer through much of the Holocene, but a period between 2200 and 800 cal yr BP had strong summer stratification, phosphate limitation, and oxygen-deficient bottom waters. Together, the proxy data suggest wet winters, protracted springs, and warm effectively wet summers in the early Holocene and less snowpack, cool springs, warm dry summers in the middle Holocene. In the late Holocene, the region and lake experienced extreme changes in winter, spring, and summer conditions, with particularly short springs and dry summers and winters during

  13. Microbial Fe(III) Oxide Reduction in Chocolate Pots Hot Springs, Yellowstone National Park

    Science.gov (United States)

    Fortney, N. W.; Roden, E. E.; Boyd, E. S.; Converse, B. J.

    2014-12-01

    Previous work on dissimilatory iron reduction (DIR) in Yellowstone National Park (YNP) has focused on high temperature, low pH environments where soluble Fe(III) is utilized as an electron acceptor for respiration. Much less attention has been paid to DIR in lower temperature, circumneutral pH environments, where solid phase Fe(III) oxides are the dominant forms of Fe(III). This study explored the potential for DIR in the warm (ca. 40-50°C), circumneutral pH Chocolate Pots hot springs (CP) in YNP. Most probable number (MPN) enumerations and enrichment culture studies confirmed the presence of endogenous microbial communities that reduced native CP Fe(III) oxides. Enrichment cultures demonstrated sustained DIR coupled to acetate and lactate oxidation through repeated transfers over ca. 450 days. Pyrosequencing of 16S rRNA genes indicated that the dominant organisms in the enrichments were closely affiliated with the well known Fe(III) reducer Geobacter metallireducens. Additional taxa included relatives of sulfate reducing bacterial genera Desulfohalobium and Thermodesulfovibrio; however, amendment of enrichments with molybdate, an inhibitor of sulfate reduction, suggested that sulfate reduction was not a primary metabolic pathway involved in DIR in the cultures. A metagenomic analysis of enrichment cultures is underway in anticipation of identifying genes involved in DIR in the less well-characterized dominant organisms. Current studies are aimed at interrogating the in situ microbial community at CP. Core samples were collected along the flow path (Fig. 1) and subdivided into 1 cm depth intervals for geochemical and microbiological analysis. The presence of significant quantities of Fe(II) in the solids indicated that DIR is active in situ. A parallel study investigated in vitro microbial DIR in sediments collected from three of the coring sites. DNA was extracted from samples from both studies for 16S rRNA gene and metagenomic sequencing in order to obtain a

  14. An REU Project on the Precambrian Rocks of Yellowstone National Park: Some lessons learned

    Science.gov (United States)

    Henry, D.; Mogk, D. W.; Mueller, P. A.; Foster, D. A.

    2014-12-01

    An NSF-funded REU project (2011-2013), based in Yellowstone National Park (YNP), was designed to characterize the geology, geochemistry and geochronology of Precambrian rocks in northern YNP. Over two field seasons two cadres of 12 students (12 women and 12 men) were chosen from small-to-large state universities and private colleges. REU students participated in three major activities constituting a complete research experience: Field studies involved geologic mapping and sampling of Precambrian basement; formulation of testable research questions by smaller working groups; and mapping and sampling projects to address research questions; Analytical studies, sample preparation immediately followed field work with petrographic analysis at students' home institutions and a week-long visit to analytical laboratories to conduct follow-up studies by small research groups during the academic year (Univ. Florida - geochemistry and geochronology; Univ. Minnesota - EMPA analysis); Communicating results, each working group submitted an abstract and collectively presented 13 posters at the 2011 and 2012 GSA Rocky Mountain sectional meetings. We used directed discovery to engage students in a community of practice in the field and found that a long apprenticeship (2-3 weeks) is optimal for novice-master interactions in exploring natural setting. Initial group hikes were used to normalize methods and language of the discipline. Students developed a sense of ownership of the overall project and assumed personal responsibility for directed research projects. Training was provided to: guide students in selection and appropriate use of tools; develop sampling strategies; discuss communal ethics, values, and expectations; develop efficient work habits; stimulate independent thinking; and engage decision-making. It was important to scaffold the field experience to students' level of development to lead to mastery. Analytical activities were designed from rock to analysis so that each

  15. The natural food habits of grizzly bears in Yellowstone National Park, 1973-74

    Science.gov (United States)

    Mealey, Stephen Patrick

    1980-01-01

     The natural food habits of grizzly bears (Ursus arctos horribilis Ord) in Yellowstone National Park were investigated in 1973-74 to identify the grizzly's energy sources and trophic level(s), nutrient use, and distribution. Food consumption was determined by scat analysis and field observations. Food quality and digestibility were estimated by chemical analysis. Grizzlies were distributed in 3 distinctive feeding economies: valley/plateau, a grass/rodent economy where grizzlies were intensive diggers; mountain, primarily a grass/springbeauty/root economy where grizzlies were casual diggers; and lake, primarily a fish/grass economy where grizzlies were fishers. The economies occured in areas with fertile soils; distribution of bears within each was related to the occurrence of succulent plants. The feeding cycle in the valley/plateau and mountain economies followed plant phenology. Grizzlies fed primarily on meat before green-up and on succulent herbs afterwards; meat, corms, berries, and nuts became important during the postgrowing season. Succulent grasses and sedges with an importance value percentage of 78.5 were the most important food items consumed. Protein from animal tissue was more digestible than protein from plant tissue. Storage fats were more digestible than structural fats. Food energy and digestibility were directly related. Five principle nutrient materials (listed with their percentage digestibilities) contributed to total energy intake: protein from succulent herbs, 42.8; protein and fat from animal material, 78.1; fat and protein from pine nuts, 73.6; starch, 78.8; and sugar from berries and fruits, digestibility undetermined. Protein from succulent herbs, with a nutritive value percentage of 77.3, was the grizzlies' primary energy source. Because succulent, preflowering herbs had higher protein levels than dry, mature herbs, grizzly use of succulent herbs guaranteed them the highest source of herbaceous protein. Low protein digestibility of

  16. Characterization of organic matter in lake sediments from Minnesota and Yellowstone National Park

    Science.gov (United States)

    Dean, Walter E.

    2006-01-01

    Samples of sediment from lakes in Minnesota and Yellowstone National Park (YNP) were analyzed for organic carbon (OC), hydrogen richness by Rock-Eval pyrolysis, and stable carbon- and nitrogen-isotope composition of bulk organic matter. Values of delta 13C of lake plankton tend to be around -28 to -32 parts per thousand (0/00). Organic matter with values of delta 13C in the high negative 20s overlap with those of organic matter derived from C3 higher terrestrial plants but are at least 10 0/00 more depleted in 13C than organic matter derived from C4 terrestrial plants. If the organic matter is produced mainly by photosynthetic plankton and is not oxidized in the water column, there may be a negative correlation between H-richness (Rock-Eval pyrolysis H-index) and delta 13C, with more H-rich, algal organic matter having lower values of delta 13C. However, if aquatic organic matter is oxidized in the water column, or if the organic matter is a mixture of terrestrial and aquatic organic matter, then there may be no correlation between H-richness and carbon-isotopic composition. Values of delta 13C lower than about -28 0/00 probably indicate a contribution of bacterial biomass produced in the hypolimnion by chemoautotrophy or methanotrophy. In highly eutrophic lakes in which large amounts of 13C-depleted organic matter is continually removed from the epilimnion by photosynthesis throughout the growing season, the entire carbon reservoir in the epilimnion may become severely 13C-enriched so that 13C-enriched photosynthetic organic matter may overprint 13C-depleted chemosynthetic bacterial organic matter produced in the hypolimnon. Most processes involved with the nitrogen cycle in lakes, such as production of ammonia and nitrate, tend to produce 15N-enriched values of delta 15N. Most Minnesota lake sediments are 15N-enriched. However, some of the more OC-rich sediments have delta 15N values close to zero (delta 15N of air), suggesting that organic matter production is

  17. Outcomes of the 'Data Curation for Geobiology at Yellowstone National Park' Workshop

    Science.gov (United States)

    Thomer, A.; Palmer, C. L.; Fouke, B. W.; Rodman, A.; Choudhury, G. S.; Baker, K. S.; Asangba, A. E.; Wickett, K.; DiLauro, T.; Varvel, V.

    2013-12-01

    The continuing proliferation of geological and biological data generated at scientifically significant sites (such as hot springs, coral reefs, volcanic fields and other unique, data-rich locales) has created a clear need for the curation and active management of these data. However, there has been little exploration of what these curation processes and policies would entail. To that end, the Site-Based Data Curation (SBDC) project is developing a framework of guidelines and processes for the curation of research data generated at scientifically significant sites. A workshop was held in April 2013 at Yellowstone National Park (YNP) to gather input from scientists and stakeholders. Workshop participants included nine researchers actively conducting geobiology research at YNP, and seven YNP representatives, including permitting staff and information professionals from the YNP research library and archive. Researchers came from a range of research areas -- geology, molecular and microbial biology, ecology, environmental engineering, and science education. Through group discussions, breakout sessions and hands-on activities, we sought to generate policy recommendations and curation guidelines for the collection, representation, sharing and quality control of geobiological datasets. We report on key themes that emerged from workshop discussions, including: - participants' broad conceptions of the long-term usefulness, reusability and value of data. - the benefits of aggregating site-specific data in general, and geobiological data in particular. - the importance of capturing a dataset's originating context, and the potential usefulness of photographs as a reliable and easy way of documenting context. - researchers' and resource managers' overlapping priorities with regards to 'big picture' data collection and management in the long-term. Overall, we found that workshop participants were enthusiastic and optimistic about future collaboration and development of community

  18. Using monitoring data to map amphibian breeding hotspots and describe wetland vulnerability in Yellowstone and Grand Teton National Parks

    Science.gov (United States)

    Ray, Andrew M.; Legg, Kristin; Sepulveda, Adam; Hossack, Blake R.; Patla, Debra

    2015-01-01

    Amphibians have been selected as a “vital sign” by several National Park Service (NPS) Inventory and Monitoring (I&M) networks. An eight-year amphibian monitoring data set provided opportunities to examine spatial and temporal patterns in amphibian breeding richness and wetland desiccation across Yellowstone and Grand Teton National Parks. Amphibian breeding richness was variable across both parks and only four of 31 permanent monitoring catchments contained all four widely distributed species. Annual breeding richness was also variable through time and fluctuated by as much as 75% in some years and catchments. Wetland desiccation was also documented across the region, but alone did not explain variations in amphibian richness. High annual variability across the region emphasizes the need for multiple years of monitoring to accurately describe amphibian richness and wetland desiccation dynamics.

  19. Beaver damming, fluvial geomorphology, and climate in Yellowstone National Park, Wyoming

    Science.gov (United States)

    Persico, L.; Meyer, G.

    2008-12-01

    Beaver habitation is an important component of many fluvial landscapes that can impact a variety of hydrologic, geomorphic, and ecologic processes. Beaver damming, via long term valley aggradation, is thought to be important to the postglacial geomorphic evolution of many smaller mountain stream networks in the western United States. Loss of beaver dams can also cause rapid channel incision. Although several studies have documented rapid short-term aggradation of channels behind single beaver dams, there is little actual data on the long-term cumulative effect of beaver damming. In Yellowstone''s Northern Range, field surveys and stratigraphic section along six streams in the Northern Range reveal net thickness of mostly beaver-pond deposits. We estimate that reaches with clear morphologic and stratigraphic evidence for beaver-related aggradation constitute about 19% of the total stream network length. Reaches with probable and possible beaver-related aggradation make up an additional 8% and 2% of the network, respectively. The remaining 71% of the network has no clear evidence for beaver-related aggradation. Thirty-nine radiocarbon ages on beaver-pond deposits in northern Yellowstone fall primarily within the last 4000 yr, but gaps in dated beaver occupation from 2200-1800 and 950-750 cal yr BP correspond with severe and persistent droughts that likely caused low to ephemeral discharges in smaller streams. In the last two decades, severe drought has also caused streams that were occupied by beaver in the 1920s to become ephemeral. Beaver have been largely absent from the Northern Range since the mid-20th century, probably due to multiple ecological and climatic factors. This loss of beaver is thought to have led to widespread degradation of stream and riparian habitat via channel incision. Although 20th-century beaver loss has caused significant channel incision at some former dam sites, downcutting elsewhere in northern Yellowstone is unrelated to beaver dams or

  20. Large carnivores response to recreational big game hunting along the Yellowstone National Park and Absaroka-Beartooth Wilderness boundary

    Science.gov (United States)

    Ruth, T.E.; Smith, D.W.; Haroldson, M.A.; Buotte, P.C.; Schwartz, C.C.; Quigley, H.B.; Cherry, S.; Tyres, D.; Frey, K.

    2003-01-01

    The Greater Yellowstone Ecosystem contains the rare combination of an intact guild of native large carnivores, their prey, and differing land management policies (National Park versus National Forest; no hunting versus hunting). Concurrent field studies on large carnivores allowed us to investigate activities of humans and carnivores on Yellowstone National Park's (YNP) northern boundary. Prior to and during the backcountry big-game hunting season, we monitored movements of grizzly bears (Ursus arctos), wolves (Canis lupus), and cougars (Puma concolor) on the northern boundary of YNP. Daily aerial telemetry locations (September 1999), augmented with weekly telemetry locations (August and October 1999), were obtained for 3 grizzly bears, 7 wolves in 2 groups of 1 pack, and 3 cougars in 1 family group. Grizzly bears were more likely located inside the YNP boundary during the pre-hunt period and north of the boundary once hunting began. The cougar family tended to be found outside YNP during the pre-hunt period and moved inside YNP when hunting began. Wolves did not significantly change their movement patterns during the pre-hunt and hunting periods. Qualitative information on elk (Cervus elaphus) indicated they moved into YNP once hunting started, suggesting that cougars followed living prey or responded to hunting activity, grizzly bears focused on dead prey (e.g., gut piles, crippled elk), and wolves may have taken advantage of both. Measures of association (Jacob's Index) were positive within carnivore species but inconclusive among species. Further collaborative research and the use of new technologies such as Global Positioning System (GPS) telemetry collars will advance our ability to understand these species, the carnivore community and its interactions, and human influences on carnivores.

  1. Small Scale Biodiversity of an Alkaline Hot Spring in Yellowstone National Park

    Science.gov (United States)

    Walther, K.; Oiler, J.; Meyer-Dombard, D. R.

    2012-12-01

    To date, many phylogenetic diversity studies have been conducted in Yellowstone National Park (YNP) [1-7] focusing on the amplification of the 16S rRNA gene and "metagenomic" datasets. However, few reports focus on diversity at small scales. Here, we report on a small scale biodiversity study of sediment and biofilm communities within a confined area of a YNP hot spring, compare and contrast these communities to other sediment and biofilm communities from previous studies [1-7], and with other sediment and biofilm communities in the same system. Sediment and biofilm samples were collected, using a 30 x 50 cm sampling grid divided in 5 x 5 cm squares, which was placed in the outflow channel of "Bat Pool", an alkaline (pH 7.9) hot spring in YNP. Accompanying geochemical data included a full range of spectrophotometry measurements along with major ions, trace elements, and DIC/DOC. In addition, in situ temperature and conductivity arrays were placed within the grid location. The temperature array closest to the source varied between 83-88°C, while the temperature array 40 cm downstream varied between ~83.5-86.5°C. The two conductivity arrays yielded measurements of 5632 μS and 5710 μS showing little variation within the sampling area. Within the grid space, DO ranged from 0.5-1.33 mg/L, with relatively similar, but slightly lower values down the outflow channel. Sulfide values within the grid ranged from 1020-1671 μg/L, while sulfide values outside of the grid region fluctuated, but generally followed the trend of decreasing from source down the outflow. Despite the relative heterogeneity of chemical and physical parameters in the grid space, there was biological diversity in sediments and biofilms at the 5 cm scale. Small scale biodiversity was analyzed by selecting a representative number of samples from within the grid. DNA was extracted and variable regions V3 and V6 (Archaea and Bacteria, respectively) were sequenced with 454 pyrosequencing. The datasets

  2. The Distribution, Diversity, and Geobiology of Thermoproteales Populations in Yellowstone National Park

    Science.gov (United States)

    Jay, Z.; Beam, J.; Bailey, C.; Dohnalkova, A.; Planer-Friedrich, B.; Romine, M.; Inskeep, W. P.

    2012-12-01

    The order Thermoproteales (phylum Crenarchaeota) consists of thermophilic, rod-shaped organisms that are found globally in geothermal habitats ranging in pH from ~3-9. Nearly all isolated Thermoproteales couple the respiration of inorganic sulfur species (e.g. elemental sulfur, thiosulfate, sulfate) to the oxidation of hydrogen or complex organic carbon. Prior 16S rRNA and metagenome analysis revealed four prominent Thermoproteales-like populations in hypoxic, sulfidic hot springs In Yellowstone National Park (YNP), WY, USA (Monarch Geyser [80° C, pH 4], Cistern Spring [76° C, pH 5] and Joseph's Coat Hot Spring [JCHS; 80° C, pH 6]). The objectives of this study were to 1) characterize and compare the indigenous Thermoproteales-like de novo assemblies identified from metagenomic sequence data available for geothermal systems across YNP, 2) determine the metabolic potential of the Thermoproteales-like populations and evaluate their role in the geochemical cycling of organic and inorganic constituents, and 3) contrast both the sequenced genome and growth physiology of the first Thermoproteales isolated from YNP ("Pyrobaculum yellowstonensis" strain WP30), to the indigenous Thermoproteales-like de novo assemblies. Sequences related to either Caldivirga or Vulcanisaeta spp. (Type I Thermoproteales) were identified in both aerobic and anaerobic habitats ranging in pH ~3 - 6. Thermoproteus or Pyrobaculum spp. (Type-II Thermoproteales) were identified in anoxic habitats, but were constrained to pH values >4. Annotation of the de novo assemblies indicate that both Type-I and Type-II Thermoproteales populations are primarily heterotrophic, although key proteins of the autotrophic dicarboxylate/4-hydroxybutyrate cycle were also identified. Caldivirga/Vulcanisaeta-like populations appear to respire on elemental sulfur, sulfate, or molecular oxygen, while the Thermoproteus/Pyrobaculum-like population may also oxidize hydrogen and respire on elemental sulfur, thiosulfate

  3. Caldicellulosiruptor obsidiansis sp. nov., an anaerobic, extremely thermophilic, cellulolytic bacterium isolated from Obsidian Pool, Yellowstone National Park.

    Science.gov (United States)

    Hamilton-Brehm, Scott D; Mosher, Jennifer J; Vishnivetskaya, Tatiana; Podar, Mircea; Carroll, Sue; Allman, Steve; Phelps, Tommy J; Keller, Martin; Elkins, James G

    2010-02-01

    A novel, obligately anaerobic, extremely thermophilic, cellulolytic bacterium, designated OB47(T), was isolated from Obsidian Pool, Yellowstone National Park, WY. The isolate was a nonmotile, non-spore-forming, Gram-positive rod approximately 2 microm long by 0.2 microm wide and grew at temperatures between 55 and 85 degrees C, with the optimum at 78 degrees C. The pH range for growth was 6.0 to 8.0, with values of near 7.0 being optimal. Growth on cellobiose produced the fastest specific growth rate at 0.75 h(-1). The organism also displayed fermentative growth on glucose, maltose, arabinose, fructose, starch, lactose, mannose, sucrose, galactose, xylose, arabinogalactan, Avicel, xylan, filter paper, processed cardboard, pectin, dilute acid-pretreated switchgrass, and Populus. OB47(T) was unable to grow on mannitol, fucose, lignin, Gelrite, acetate, glycerol, ribose, sorbitol, carboxymethylcellulose, and casein. Yeast extract stimulated growth, and thiosulfate, sulfate, nitrate, and sulfur were not reduced. Fermentation end products were mainly acetate, H2, and CO2, although lactate and ethanol were produced in 5-liter batch fermentations. The G+C content of the DNA was 35 mol%, and sequence analysis of the small subunit rRNA gene placed OB47(T) within the genus Caldicellulosiruptor. Based on its phylogenetic and phenotypic properties, the isolate is proposed to be designated Caldicellulosiruptor obsidiansis sp. nov. and OB47 is the type strain (ATCC BAA-2073).

  4. Thermodesulfobacterium geofontis sp. nov., a hyperthermophilic, sulfate-reducing bacterium isolated from Obsidian Pool, Yellowstone National Park.

    Science.gov (United States)

    Hamilton-Brehm, Scott D; Gibson, Robert A; Green, Stefan J; Hopmans, Ellen C; Schouten, Stefan; van der Meer, Marcel T J; Shields, John P; Damsté, Jaap S S; Elkins, James G

    2013-03-01

    A novel sulfate-reducing bacterium designated OPF15(T) was isolated from Obsidian Pool, Yellowstone National Park, Wyoming. The phylogeny of 16S rRNA and functional genes (dsrAB) placed the organism within the family Thermodesulfobacteriaceae. The organism displayed hyperthermophilic temperature requirements for growth with a range of 70-90 °C and an optimum of 83 °C. Optimal pH was around 6.5-7.0 and the organism required the presence of H2 or formate as an electron donor and CO2 as a carbon source. Electron acceptors supporting growth included sulfate, thiosulfate, and elemental sulfur. Lactate, acetate, pyruvate, benzoate, oleic acid, and ethanol did not serve as electron donors. Membrane lipid analysis revealed diacyl glycerols and acyl/ether glycerols which ranged from C14:0 to C20:0. Alkyl chains present in acyl/ether and diether glycerol lipids ranged from C16:0 to C18:0. Straight, iso- and anteiso-configurations were found for all lipid types. The presence of OPF15(T) was also shown to increase cellulose consumption during co-cultivation with Caldicellulosiruptor obsidiansis, a fermentative, cellulolytic extreme thermophile isolated from the same environment. On the basis of phylogenetic, phenotypic, and structural analyses, Thermodesulfobacterium geofontis sp. nov. is proposed as a new species with OPF15(T) representing the type strain.

  5. Caldicellulosiruptor obsidiansis sp. nov., an Anaerobic, Extremely Thermophilic, Cellulolytic Bacterium Isolated from Obsidian Pool, Yellowstone National Park▿

    Science.gov (United States)

    Hamilton-Brehm, Scott D.; Mosher, Jennifer J.; Vishnivetskaya, Tatiana; Podar, Mircea; Carroll, Sue; Allman, Steve; Phelps, Tommy J.; Keller, Martin; Elkins, James G.

    2010-01-01

    A novel, obligately anaerobic, extremely thermophilic, cellulolytic bacterium, designated OB47T, was isolated from Obsidian Pool, Yellowstone National Park, WY. The isolate was a nonmotile, non-spore-forming, Gram-positive rod approximately 2 μm long by 0.2 μm wide and grew at temperatures between 55 and 85°C, with the optimum at 78°C. The pH range for growth was 6.0 to 8.0, with values of near 7.0 being optimal. Growth on cellobiose produced the fastest specific growth rate at 0.75 h−1. The organism also displayed fermentative growth on glucose, maltose, arabinose, fructose, starch, lactose, mannose, sucrose, galactose, xylose, arabinogalactan, Avicel, xylan, filter paper, processed cardboard, pectin, dilute acid-pretreated switchgrass, and Populus. OB47T was unable to grow on mannitol, fucose, lignin, Gelrite, acetate, glycerol, ribose, sorbitol, carboxymethylcellulose, and casein. Yeast extract stimulated growth, and thiosulfate, sulfate, nitrate, and sulfur were not reduced. Fermentation end products were mainly acetate, H2, and CO2, although lactate and ethanol were produced in 5-liter batch fermentations. The G+C content of the DNA was 35 mol%, and sequence analysis of the small subunit rRNA gene placed OB47T within the genus Caldicellulosiruptor. Based on its phylogenetic and phenotypic properties, the isolate is proposed to be designated Caldicellulosiruptor obsidiansis sp. nov. and OB47 is the type strain (ATCC BAA-2073). PMID:20023107

  6. Time scale of hydrothermal water-rock reactions in Yellowstone National Park based on radium isotopes and radon

    International Nuclear Information System (INIS)

    Clark, J.F.; Turekian, K.K.

    1990-01-01

    We have measured 224 Ra (3.4 d), 228 Ra (5.7 yr), and 226 Ra (1620 yr) and chloride in hot spring waters from the Norris-Mammoth Corridor, Yellowstone National Park. Two characteristic cold-water components mix with the primary hydrothermal water: one for the travertine-depositing water related to the Mammoth Hot Springs and the other for the sinter-depositing Norris Geyser Basin springs. The Mammoth Hot Springs water is a mixture of the primary hydrothermal fluid with meteoric waters flowing through the Madison Limestone, as shown by the systematic decrease of the ( 228 Ra/ 226 Ra) activity ratio proceeding northward. The Norris Geyser Basin springs are mixtures of primary hydrothermal water with different amounts of cold meteoric water with no modification of the primary hydrothermal ( 228 Ra/ 226 Ra) activity ratio. Using a solution and recoil model for radium isotope supply to the primary hydrothermal water, a mean water-rock reaction time prior to expansion at 350degC and supply to the surface is 540 years assuming that 250 g of water are involved in the release of the radium from one gram of rock. The maximum reaction time allowed by our model is 1150 years. (orig.)

  7. Geoarchaeota: a new candidate phylum in the Archaea from high-temperature acidic iron mats in Yellowstone National Park.

    Science.gov (United States)

    Kozubal, Mark A; Romine, Margaret; Jennings, Ryan deM; Jay, Zack J; Tringe, Susannah G; Rusch, Doug B; Beam, Jacob P; McCue, Lee Ann; Inskeep, William P

    2013-03-01

    Geothermal systems in Yellowstone National Park (YNP) provide an outstanding opportunity to understand the origin and evolution of metabolic processes necessary for life in extreme environments including low pH, high temperature, low oxygen and elevated concentrations of reduced iron. Previous phylogenetic studies of acidic ferric iron mats from YNP have revealed considerable diversity of uncultivated and undescribed archaea. The goal of this study was to obtain replicate de novo genome assemblies for a dominant archaeal population inhabiting acidic iron-oxide mats in YNP. Detailed analysis of conserved ribosomal and informational processing genes indicates that the replicate assemblies represent a new candidate phylum within the domain Archaea referred to here as 'Geoarchaeota' or 'novel archaeal group 1 (NAG1)'. The NAG1 organisms contain pathways necessary for the catabolism of peptides and complex carbohydrates as well as a bacterial-like Form I carbon monoxide dehydrogenase complex likely used for energy conservation. Moreover, this novel population contains genes involved in the metabolism of oxygen including a Type A heme copper oxidase, a bd-type terminal oxidase and a putative oxygen-sensing protoglobin. NAG1 has a variety of unique bacterial-like cofactor biosynthesis and transport genes and a Type3-like CRISPR system. Discovery of NAG1 is critical to our understanding of microbial community structure and function in extant thermophilic iron-oxide mats of YNP, and will provide insight regarding the evolution of Archaea in early Earth environments that may have important analogs active in YNP today.

  8. Radiocarbon dating of silica sinter deposits in shallow drill cores from the Upper Geyser Basin, Yellowstone National Park

    Science.gov (United States)

    Lowenstern, Jacob B.; Hurwitz, Shaul; McGeehin, John P.

    2016-01-01

    To explore the timing of hydrothermal activity at the Upper Geyser Basin (UGB) in Yellowstone National Park, we obtained seven new accelerator mass spectrometry (AMS) radiocarbon 14C ages of carbonaceous material trapped within siliceous sinter. Five samples came from depths of 15-152 cm within the Y-1 well, and two samples were from well Y-7 (depths of 24 cm and 122 cm). These two wells, at Black Sand and Biscuit Basins, respectively, were drilled in 1967 as part of a scientific drilling program by the U.S. Geological Survey (White et al., 1975). Even with samples as small as 15 g, we obtained sufficient carbonaceous material (a mixture of thermophilic mats, pollen, and charcoal) for the 14C analyses. Apparent time of deposition ranged from 3775 ± 25 and 2910 ± 30 14C years BP at the top of the cores to about 8000 years BP at the bottom. The dates are consistent with variable rates of sinter formation at individual sites within the UGB over the Holocene. On a basin-wide scale, though, these and other existing 14C dates hint that hydrothermal activity at the UGB may have been continuous throughout the Holocene.

  9. Geoarchaeota: a new candidate phylum in the Archaea from high-temperature acidic iron mats in Yellowstone National Park

    Energy Technology Data Exchange (ETDEWEB)

    Kozubal, Mark; Romine, Margaret F.; Jennings, Ryan; Jay, Z.; Tringe, Susannah G.; Rusch, Douglas B.; Beam, Jake; McCue, Lee Ann; Inskeep, William P.

    2013-03-01

    Geothermal systems in Yellowstone National Park (YNP) provide an outstanding opportunity to understand the origin and evolution of metabolic processes necessary for life in extreme environments including low pH, high temperature, low oxygen and elevated concentrations of reduced iron. Previous phylogenetic studies of acidic ferric iron mats from YNP have revealed considerable diversity of uncultivated and undescribed archaea. The goal of this study was to obtain replicate de novo genome assemblies for a dominant archaeal population inhabiting acidic iron oxide mats in YNP. Detailed analysis of conserved ribosomal and informational processing genes indicate that the replicate assemblies represent a new phylum-level lineage referred to here as 'novel archaeal group 1 (NAG1)'. The NAG1 organisms contain pathways necessary for the catabolism of peptides and complex carbohydrates as well as a bacterial-like Form I CO dehydrogenase complex likely used for energy conservation. Moreover, this novel population contains genes involved in metabolism of oxygen including a Type A heme copper oxidase, a bd-type terminal oxidase and a putative oxygen sensing protoglobin. NAG1 has a variety of unique bacterial-like cofactor biosynthesis and transport genes and a Type3-like CRISPR system. Discovery of NAG1 is critical to our understanding of microbial community structure and function in extant thermophilic iron mats of YNP, and will provide insight regarding the evolution of Archaea in early Earth environments that may have important analogues active in YNP today.

  10. Multireaction equilibrium geothermometry: A sensitivity analysis using data from the Lower Geyser Basin, Yellowstone National Park, USA

    Science.gov (United States)

    King, Jonathan M.; Hurwitz, Shaul; Lowenstern, Jacob B.; Nordstrom, D. Kirk; McCleskey, R. Blaine

    2016-01-01

    A multireaction chemical equilibria geothermometry (MEG) model applicable to high-temperature geothermal systems has been developed over the past three decades. Given sufficient data, this model provides more constraint on calculated reservoir temperatures than classical chemical geothermometers that are based on either the concentration of silica (SiO2), or the ratios of cation concentrations. A set of 23 chemical analyses from Ojo Caliente Spring and 22 analyses from other thermal features in the Lower Geyser Basin of Yellowstone National Park are used to examine the sensitivity of calculated reservoir temperatures using the GeoT MEG code (Spycher et al. 2013, 2014) to quantify the effects of solute concentrations, degassing, and mineral assemblages on calculated reservoir temperatures. Results of our analysis demonstrate that the MEG model can resolve reservoir temperatures within approximately ±15°C, and that natural variation in fluid compositions represents a greater source of variance in calculated reservoir temperatures than variations caused by analytical uncertainty (assuming ~5% for major elements). The analysis also suggests that MEG calculations are particularly sensitive to variations in silica concentration, the concentrations of the redox species Fe(II) and H2S, and that the parameters defining steam separation and CO2 degassing from the liquid may be adequately determined by numerical optimization. Results from this study can provide guidance for future applications of MEG models, and thus provide more reliable information on geothermal energy resources during exploration.

  11. Insights into archaeal evolution and symbiosis from the genomes of a Nanoarchaeon and its crenarchaeal host from Yellowstone National Park

    Energy Technology Data Exchange (ETDEWEB)

    Podar, Mircea [ORNL; Graham, David E [ORNL; Reysenbach, Anna-Louise [Portland State University; Koonin, Eugene [National Center for Biotechnology Information; Wolf, Yuri [National Center for Biotechnology Information; Makarova, Kira S. [National Center for Biotechnology Information

    2013-01-01

    A hyperthemophilic member of the Nanoarchaeota from Obsidian Pool, a thermal feature in Yellowstone National Park was characterized using single cell isolation and sequencing, together with its putative host, a Sulfolobales archaeon. This first representative of a non-marine Nanoarchaeota (Nst1) resembles Nanoarchaeum equitans by lacking most biosynthetic capabilities, the two forming a deep-branching archaeal lineage. However, the Nst1 genome is over 20% larger, encodes a complete gluconeogenesis pathway and a full complement of archaeal flagellum proteins. Comparison of the two genomes suggests that the marine and terrestrial Nanoarchaeota lineages share a common ancestor that was already a symbiont of another archaeon. With a larger genome, a smaller repertoire of split protein encoding genes and no split non-contiguous tRNAs, Nst1 appears to have experienced less severe genome reduction than N. equitans. The inferred host of Nst1 is potentially autotrophic, with a streamlined genome and simplified central and energetic metabolism as compared to other Sulfolobales. The two distinct Nanoarchaeota-host genomic data sets offer insights into the evolution of archaeal symbiosis and parasitism and will further enable studies of the cellular and molecular mechanisms of these relationships.

  12. Provisional maps of thermal areas in Yellowstone National Park, based on satellite thermal infrared imaging and field observations

    Science.gov (United States)

    Vaughan, R. Greg; Heasler, Henry; Jaworowski, Cheryl; Lowenstern, Jacob B.; Keszthelyi, Laszlo P.

    2014-01-01

    Maps that define the current distribution of geothermally heated ground are useful toward setting a baseline for thermal activity to better detect and understand future anomalous hydrothermal and (or) volcanic activity. Monitoring changes in the dynamic thermal areas also supports decisions regarding the development of Yellowstone National Park infrastructure, preservation and protection of park resources, and ensuring visitor safety. Because of the challenges associated with field-based monitoring of a large, complex geothermal system that is spread out over a large and remote area, satellite-based thermal infrared images from the Advanced Spaceborne Thermal Emission and Reflection Radiometer (ASTER) were used to map the location and spatial extent of active thermal areas, to generate thermal anomaly maps, and to quantify the radiative component of the total geothermal heat flux. ASTER thermal infrared data acquired during winter nights were used to minimize the contribution of solar heating of the surface. The ASTER thermal infrared mapping results were compared to maps of thermal areas based on field investigations and high-resolution aerial photos. Field validation of the ASTER thermal mapping is an ongoing task. The purpose of this report is to make available ASTER-based maps of Yellowstone’s thermal areas. We include an appendix containing the names and characteristics of Yellowstone’s thermal areas, georeferenced TIFF files containing ASTER thermal imagery, and several spatial data sets in Esri shapefile format.

  13. Pullman transportation plan : a transportation and access plan for Pullman National Monument and the surrounding neighborhood

    Science.gov (United States)

    2017-09-01

    The Pullman Transportation Plan provides a holistic set of recommendations to improve access to and from Pullman National Monument and its surrounding neighborhoods for both visitors and residents. In this plan, we identify short-, medium-, and long-...

  14. Foraging and feeding ecology of the gray wolf (Canis lupus): lessons from Yellowstone National Park, Wyoming, USA.

    Science.gov (United States)

    Stahler, Daniel R; Smith, Douglas W; Guernsey, Debra S

    2006-07-01

    The foraging and feeding ecology of gray wolves is an essential component to understanding the role that top carnivores play in shaping the structure and function of terrestrial ecosystems. In Yellowstone National Park (YNP), predation studies on a highly visible, reintroduced population of wolves are increasing our understanding of this aspect of wolf ecology. Wolves in YNP feed primarily on elk, despite the presence of other ungulate species. Patterns of prey selection and kill rates in winter have varied seasonally each year from 1995 to 2004 and changed in recent years as the wolf population has become established. Wolves select elk based on their vulnerability as a result of age, sex, and season and therefore kill primarily calves, old cows, and bulls that have been weakened by winter. Summer scat analysis reveals an increased variety in diet compared with observed winter diets, including other ungulate species, rodents, and vegetation. Wolves in YNP hunt in packs and, upon a successful kill, share in the evisceration and consumption of highly nutritious organs first, followed by major muscle tissue, and eventually bone and hide. Wolves are adapted to a feast-or-famine foraging pattern, and YNP packs typically kill and consume an elk every 2-3 d. However, wolves in YNP have gone without fresh meat for several weeks by scavenging off old carcasses that consist mostly of bone and hide. As patterns of wolf density, prey density, weather, and vulnerability of prey change, in comparison with the conditions of the study period described here, we predict that there will also be significant changes in wolf predation patterns and feeding behavior.

  15. Modeling the Habitat Range of Phototrophic Microorganisms in Yellowstone National Park: Toward the Development of a Comprehensive Fitness Landscape

    Directory of Open Access Journals (Sweden)

    Eric eBoyd

    2012-06-01

    Full Text Available The extent to which geochemical variation constrains the distribution of phototrophic metabolisms was modeled based on 439 observations in geothermal springs in Yellowstone National Park (YNP, Wyoming. Generalized additive models (GAMs were developed to predict the distribution of photosynthesis as a function of spring temperature, pH, and total sulfide. GAMs comprised of temperature explained 42.7% of the variation in the distribution of phototrophic metabolisms whereas GAMs comprised of sulfide and pH explained 20.7% and 11.7% of the variation, respectively. These results suggest that of the measured variables, temperature is the primary constraint on the distribution of phototrophic metabolism in YNP. GAMs comprised of multiple variables explained a larger percentage of the variation in the distribution of phototrophic metabolism, indicating additive interactions among variables. A GAM that combined temperature and sulfide explained the greatest variation in the dataset (54.8% while minimizing the introduction of degrees of freedom. In an effort to verify the extent to which phototroph distribution reflects constraints on activity, we examined the influence of sulfide and temperature on dissolved inorganic carbon (DIC uptake rates under both light and dark conditions. Light-driven DIC uptake decreased systematically with increasing concentrations of sulfide in acidic, algal-dominated systems, but was unaffected in alkaline, bacterial-dominated systems. In both alkaline and acidic systems, light-driven DIC uptake was suppressed in cultures incubated at temperatures 10°C greater than their in situ temperature. Collectively, these results suggest that the habitat range of phototrophs in YNP springs, specifically that of cyanobacteria and algae, largely results from constraints imposed by temperature and sulfide on the activity and fitness of these populations, a finding that is consistent with the predictions from GAMs.

  16. Complete Genome Sequence of Paenibacillus strain Y4.12MC10, a Novel Paenibacillus lautus strain Isolated from Obsidian Hot Spring in Yellowstone National Park.

    Science.gov (United States)

    Mead, David A; Lucas, Susan; Copeland, Alex; Lapidus, Alla; Cheng, Jan-Feng; Bruce, David C; Goodwin, Lynne A; Pitluck, Sam; Chertkov, Olga; Zhang, Xiaojing; Detter, John C; Han, Cliff S; Tapia, Roxanne; Land, Miriam; Hauser, Loren J; Chang, Yun-Juan; Kyrpides, Nikos C; Ivanova, Natalia N; Ovchinnikova, Galina; Woyke, Tanja; Brumm, Catherine; Hochstein, Rebecca; Schoenfeld, Thomas; Brumm, Phillip

    2012-07-30

    Paenibacillus sp.Y412MC10 was one of a number of organisms isolated from Obsidian Hot Spring, Yellowstone National Park, Montana, USA under permit from the National Park Service. The isolate was initially classified as a Geobacillus sp. Y412MC10 based on its isolation conditions and similarity to other organisms isolated from hot springs at Yellowstone National Park. Comparison of 16 S rRNA sequences within the Bacillales indicated that Geobacillus sp.Y412MC10 clustered with Paenibacillus species, and the organism was most closely related to Paenibacillus lautus. Lucigen Corp. prepared genomic DNA and the genome was sequenced, assembled, and annotated by the DOE Joint Genome Institute. The genome sequence was deposited at the NCBI in October 2009 (NC_013406). The genome of Paenibacillus sp. Y412MC10 consists of one circular chromosome of 7,121,665 bp with an average G+C content of 51.2%. Comparison to other Paenibacillus species shows the organism lacks nitrogen fixation, antibiotic production and social interaction genes reported in other paenibacilli. The Y412MC10 genome shows a high level of synteny and homology to the draft sequence of Paenibacillus sp. HGF5, an organism from the Human Microbiome Project (HMP) Reference Genomes. This, combined with genomic CAZyme analysis, suggests an intestinal, rather than environmental origin for Y412MC10.

  17. 77 FR 53908 - Winter Use Plan, Supplemental Draft Environmental Impact Statement, Yellowstone National Park

    Science.gov (United States)

    2012-09-04

    ... DEPARTMENT OF THE INTERIOR National Park Service [NPS-IMRO-YELL-11188; 2310-0070-422] Winter Use... comment period on the Draft Supplemental Environmental Impact Statement (Draft SEIS) for a Winter Use Plan... online at http://parkplanning.nps.gov/YELL (click on the link to the 2012 Supplemental Winter Use Plan...

  18. Effects of potential geothermal development in the Corwin Springs Known Geothermal Resources Area, Montana, on the thermal features of Yellowstone National Park. Water Resources Investigation

    International Nuclear Information System (INIS)

    Sorey, M.L.

    1991-01-01

    A two-year study by the U.S. Geological Survey, in collaboration with the National Park Service, Argonne National Laboratory, and Los Alamos National Laboratory was initiated in 1988 to determine the effects of potential geothermal development in the Corwin Springs Known Geothermal Resources Area (KGRA), Montana, on the thermal features of Yellowstone National Park. The study addressed three principal issues: (1) the sources of thermal water in the hot springs at Mammoth, La Duke, and Bear Creek; (2) the degree of subsurface connection between these areas; and (3) the effects of geothermal development in the Corwin Springs KGRA on the Park's thermal features. The authors investigations included, but were not limited to, geologic mapping, electrical geophysical surveys, chemical sampling and analyses of waters and rocks, determinations of the rates of discharge of various thermal springs, and hydrologic tracer tests

  19. Yellowstone Lake Nanoarchaeota

    Directory of Open Access Journals (Sweden)

    Scott eClingenpeel

    2013-09-01

    Full Text Available Considerable Nanoarchaeota novelty and diversity were encountered in Yellowstone Lake, Yellowstone National Park, where sampling targeted lake floor hydrothermal vent fluids, streamers and sediments associated with these vents, and in planktonic photic zones in three different regions of the lake. Significant homonucleotide repeats (HR were observed in pyrosequence reads and in near full-length Sanger sequences, averaging 112 HR per 1,349 bp clone and could confound diversity estimates derived from pyrosequencing, resulting in false nucleotide insertions or deletions (indels. However, Sanger sequencing of two different sets of PCR clones (110 bp, 1349 bp demonstrated that at least some of these indels are real. The majority of the Nanoarchaeota PCR amplicons were vent associated; however, curiously, one relatively small Nanoarchaeota OTU (70 pyrosequencing reads was only found in photic zone water samples obtained from a region of the lake furthest removed from the hydrothermal regions of the lake. Extensive pyrosequencing failed to demonstrate the presence of an Ignicoccus lineage in this lake, suggesting the Nanoarchaeota in this environment are associated with novel Archaea hosts. Defined phylogroups based on near full-length PCR clones document the significant Nanoarchaeota 16S rRNA gene diversity in this lake and firmly establish a terrestrial clade distinct from the marine Nanoarcheota as well as from other geographical locations.

  20. Metal loading in Soda Butte Creek upstream of Yellowstone National Park, Montana and Wyoming; a retrospective analysis of previous research; and quantification of metal loading, August 1999

    Science.gov (United States)

    Boughton, G.K.

    2001-01-01

    Acid drainage from historic mining activities has affected the water quality and aquatic biota of Soda Butte Creek upstream of Yellowstone National Park. Numerous investigations focusing on metals contamination have been conducted in the Soda Butte Creek basin, but interpretations of how metals contamination is currently impacting Soda Butte Creek differ greatly. A retrospective analysis of previous research on metal loading in Soda Butte Creek was completed to provide summaries of studies pertinent to metal loading in Soda Butte Creek and to identify data gaps warranting further investigation. Identification and quantification of the sources of metal loading to Soda Butte Creek was recognized as a significant data gap. The McLaren Mine tailings impoundment and mill site has long been identified as a source of metals but its contribution relative to the total metal load entering Yellowstone National Park was unknown. A tracer-injection and synoptic-sampling study was designed to determine metal loads upstream of Yellowstone National Park.A tracer-injection and synoptic-sampling study was conducted on an 8,511-meter reach of Soda Butte Creek from upstream of the McLaren Mine tailings impoundment and mill site downstream to the Yellowstone National Park boundary in August 1999. Synoptic-sampling sites were selected to divide the creek into discrete segments. A lithium bromide tracer was injected continuously into Soda Butte Creek for 24.5 hours. Downstream dilution of the tracer and current-meter measurements were used to calculate the stream discharge. Stream discharge values, combined with constituent concentrations obtained by synoptic sampling, were used to quantify constituent loading in each segment of Soda Butte Creek.Loads were calculated for dissolved calcium, silica, and sulfate, as well as for dissolved and total-recoverable iron, aluminum, and manganese. Loads were not calculated for cadmium, copper, lead, and zinc because these elements were infrequently

  1. Seasonal patterns of predation for gray wolves in the multi-prey system of Yellowstone National Park.

    Science.gov (United States)

    Metz, Matthew C; Smith, Douglas W; Vucetich, John A; Stahler, Daniel R; Peterson, Rolf O

    2012-05-01

    1. For large predators living in seasonal environments, patterns of predation are likely to vary among seasons because of related changes in prey vulnerability. Variation in prey vulnerability underlies the influence of predators on prey populations and the response of predators to seasonal variation in rates of biomass acquisition. Despite its importance, seasonal variation in predation is poorly understood. 2. We assessed seasonal variation in prey composition and kill rate for wolves Canis lupus living on the Northern Range (NR) of Yellowstone National Park. Our assessment was based on data collected over 14 winters (1995-2009) and five spring-summers between 2004 and 2009. 3. The species composition of wolf-killed prey and the age and sex composition of wolf-killed elk Cervus elaphus (the primary prey for NR wolves) varied among seasons. 4. One's understanding of predation depends critically on the metric used to quantify kill rate. For example, kill rate was greatest in summer when quantified as the number of ungulates acquired per wolf per day, and least during summer when kill rate was quantified as the biomass acquired per wolf per day. This finding contradicts previous research that suggests that rates of biomass acquisition for large terrestrial carnivores tend not to vary among seasons. 5. Kill rates were not well correlated among seasons. For example, knowing that early-winter kill rate is higher than average (compared with other early winters) provides little basis for anticipating whether kill rates a few months later during late winter will be higher or lower than average (compared with other late winters). This observation indicates how observing, for example, higher-than-average kill rates throughout any particular season is an unreliable basis for inferring that the year-round average kill rate would be higher than average. 6. Our work shows how a large carnivore living in a seasonal environment displays marked seasonal variation in

  2. Insights into Near-Surface Structural Control of Hydrothermal Fluid Movement at Rabbit Creek Thermal Area, Yellowstone National Park

    Science.gov (United States)

    Carr, B.; Elliot, M.; Sims, K. W. W.

    2017-12-01

    Recent geophysical imaging efforts at Yellowstone National Park have generated questions about the geologic controls of hydrothermal fluid movement within the parks thermal areas. Currently, faults and lava flow contacts are assumed to be the primary permeability pathways for deeper fluid migration to the surface. Although intuition dictates that these structures are responsible, few studies have definitively shown that this is true. Earlier geophysical imaging efforts of phase separation in Norris Geyser Basin have shown strong evidence for fractures and faulting conducting hydrothermal waters. However, no geologically mapped faults are at the surface to confirm these interpretations. Therefore, during the summer of 2017, UW surface geophysical data acquisition focused on understanding the geologic controls for a thermal area within the well-mapped Rabbit Creek Fault Zone (RCFZ). The RCFZ strikes N-S along the eastern edge of Midway Geyser Basin (i.e. the western edge of the Mallard Lake Dome) about 2.8 Km SE of Grand Prismatic spring. The section of the fault zone within the Rabbit Creek thermal area is exposed on the eastern valley wall and dips steeply to the west. Regardless at our site, this puts the two of the plateau rhyolites (i.e. the Biscuit Basin Flow and Mallard Lake flow) next to each other ( 100 m apart) with a small amount of overlying alluvial, glacial and hydrothermal deposits covering the actual fault trace. Interestingly, at least two mapped reverse faults from the Mallard Lake Dome trend NW-SE into the site and are interpreted to intersect to the RCFZ. At RCFZ, DC resistivity and seismic refraction profiling combined with Self-Potential, Magnetics, and Transient Electromagnetic soundings were acquired to provide images and in situ geophysical properties. These data highlight the variable fracturing and surface expressions of the hydrothermal fluids associated with the RCFZ and the NW trending fault zone associated with the Mallard Lake Dome

  3. Isolation and Distribution of a Novel Iron-Oxidizing Crenarchaeon from Acidic Geothermal Springs in Yellowstone National Park▿ †

    Science.gov (United States)

    Kozubal, M.; Macur, R. E.; Korf, S.; Taylor, W. P.; Ackerman, G. G.; Nagy, A.; Inskeep, W. P.

    2008-01-01

    Novel thermophilic crenarchaea have been observed in Fe(III) oxide microbial mats of Yellowstone National Park (YNP); however, no definitive work has identified specific microorganisms responsible for the oxidation of Fe(II). The objectives of the current study were to isolate and characterize an Fe(II)-oxidizing member of the Sulfolobales observed in previous 16S rRNA gene surveys and to determine the abundance and distribution of close relatives of this organism in acidic geothermal springs containing high concentrations of dissolved Fe(II). Here we report the isolation and characterization of the novel, Fe(II)-oxidizing, thermophilic, acidophilic organism Metallosphaera sp. strain MK1 obtained from a well-characterized acid-sulfate-chloride geothermal spring in Norris Geyser Basin, YNP. Full-length 16S rRNA gene sequence analysis revealed that strain MK1 exhibits only 94.9 to 96.1% sequence similarity to other known Metallosphaera spp. and less than 89.1% similarity to known Sulfolobus spp. Strain MK1 is a facultative chemolithoautotroph with an optimum pH range of 2.0 to 3.0 and an optimum temperature range of 65 to 75°C. Strain MK1 grows optimally on pyrite or Fe(II) sorbed onto ferrihydrite, exhibiting doubling times between 10 and 11 h under aerobic conditions (65°C). The distribution and relative abundance of MK1-like 16S rRNA gene sequences in 14 acidic geothermal springs containing Fe(III) oxide microbial mats were evaluated. Highly related MK1-like 16S rRNA gene sequences (>99% sequence similarity) were consistently observed in Fe(III) oxide mats at temperatures ranging from 55 to 80°C. Quantitative PCR using Metallosphaera-specific primers confirmed that organisms highly similar to strain MK1 comprised up to 40% of the total archaeal community at selected sites. The broad distribution of highly related MK1-like 16S rRNA gene sequences in acidic Fe(III) oxide microbial mats is consistent with the observed characteristics and growth optima of

  4. Utilizing Structure-from-Motion Photogrammetry with Airborne Visual and Thermal Images to Monitor Thermal Areas in Yellowstone National Park

    Science.gov (United States)

    Carr, B. B.; Vaughan, R. G.

    2017-12-01

    The thermal areas in Yellowstone National Park (Wyoming, USA) are constantly changing. Persistent monitoring of these areas is necessary to better understand the behavior and potential hazards of both the thermal features and the deeper hydrothermal system driving the observed surface activity. As part of the Park's monitoring program, thousands of visual and thermal infrared (TIR) images have been acquired from a variety of airborne platforms over the past decade. We have used structure-from-motion (SfM) photogrammetry techniques to generate a variety of data products from these images, including orthomosaics, temperature maps, and digital elevation models (DEMs). Temperature maps were generated for Upper Geyser Basin and Norris Geyser Basin for the years 2009-2015, by applying SfM to nighttime TIR images collected from an aircraft-mounted forward-looking infrared (FLIR) camera. Temperature data were preserved through the SfM processing by applying a uniform linear stretch over the entire image set to convert between temperature and a 16-bit digital number. Mosaicked temperature maps were compared to the original FLIR image frames and to ground-based temperature data to constrain the accuracy of the method. Due to pixel averaging and resampling, among other issues, the derived temperature values are typically within 5-10 ° of the values of the un-resampled image frame. We also created sub-meter resolution DEMs from airborne daytime visual images of individual thermal areas. These DEMs can be used for resource and hazard management, and in cases where multiple DEMs exist from different times, for measuring topographic change, including change due to thermal activity. For example, we examined the sensitivity of the DEMs to topographic change by comparing DEMs of the travertine terraces at Mammoth Hot Springs, which can grow at > 1 m per year. These methods are generally applicable to images from airborne platforms, including planes, helicopters, and unmanned aerial

  5. Water-chemistry data for selected springs, geysers, and streams in Yellowstone National Park, Wyoming, 1999-2000

    Science.gov (United States)

    Ball, James W.; McCleskey, R. Blaine; Nordstrom, D. Kirk; Holloway, JoAnn M.; Verplanck, Philip L.; Sturtevant, Sabin A.

    2002-01-01

    Sixty-seven water analyses are reported for samples collected from 44 hot springs and their overflow drainages and two ambient-temperature acid streams in Yellowstone National Park (YNP) during 1990-2000. Thirty-seven analyses are reported for 1999, 18 for June of 2000, and 12 for September of 2000. These water samples were collected and analyzed as part of research investigations in YNP on microbially mediated sulfur oxidation in stream water, arsenic and sulfur redox speciation in hot springs, and chemical changes in overflow drainages that affect major ions, redox species, and trace elements. Most samples were collected from sources in the Norris Geyser Basin. Two ambient-temperature acidic stream systems, Alluvium and Columbine Creeks and their tributaries in Brimstone Basin, were studied in detail. Analyses were performed at or near the sampling site, in an on-site mobile laboratory truck, or later in a USGS laboratory, depending on stability of the constituent and whether or not it could be preserved effectively. Water temperature, specific conductance, pH, Eh, dissolved oxygen (D.O.), and dissolved H2S were determined on-site at the time of sampling. Alkalinity, acidity, and F were determined within a few days of sample collection by titration with acid, titration with base, and ion-selective electrode or ion chromatography (IC), respectively. Concentrations of S2O3 and SxO6 were determined as soon as possible (minutes to hours later) by IC. Concentrations of Br, Cl, NH4, NO2, NO3, SO4, Fe(II), and Fe(total) were determined within a few days of sample collection. Densities were determined later in the USGS laboratory. Concentrations of Li and K were determined by flame atomic absorption spectrometry. Concentrations of Al, As(total), B, Ba, Be, Ca, Cd, Co, Cr, Cu, Fe(total), K, Li, Mg, Mn, Na, Ni, Pb, Se, Si, Sr, V, and Zn were determined by inductively-coupled plasma-optical emission spectrometry. Trace concentrations of Cd, Cr, Cu, Pb, and Sb were

  6. Water-chemistry data for selected springs, geysers, and streams in Yellowstone National Park, Wyoming, 2006-2008

    Science.gov (United States)

    Ball, James W.; McMleskey, R. Blaine; Nordstrom, D. Kirk

    2010-01-01

    Water analyses are reported for 104 samples collected from numerous thermal and non-thermal features in Yellowstone National Park (YNP) during 2006-2008. Water samples were collected and analyzed for major and trace constituents from 10 areas of YNP including Apollinaris Spring and Nymphy Creek along the Norris-Mammoth corridor, Beryl Spring in Gibbon Canyon, Norris Geyser Basin, Lower Geyser Basin, Crater Hills, the Geyser Springs Group, Nez Perce Creek, Rabbit Creek, the Mud Volcano area, and Washburn Hot Springs. These water samples were collected and analyzed as part of research investigations in YNP on arsenic, antimony, iron, nitrogen, and sulfur redox species in hot springs and overflow drainages, and the occurrence and distribution of dissolved mercury. Most samples were analyzed for major cations and anions, trace metals, redox species of antimony, arsenic, iron, nitrogen, and sulfur, and isotopes of hydrogen and oxygen. Analyses were performed at the sampling site, in an on-site mobile laboratory vehicle, or later in a U.S. Geological Survey laboratory, depending on stability of the constituent and whether it could be preserved effectively. Water samples were filtered and preserved on-site. Water temperature, specific conductance, pH, emf (electromotive force or electrical potential), and dissolved hydrogen sulfide were measured on-site at the time of sampling. Dissolved hydrogen sulfide was measured a few to several hours after sample collection by ion-specific electrode on samples preserved on-site. Acidity was determined by titration, usually within a few days of sample collection. Alkalinity was determined by titration within 1 to 2 weeks of sample collection. Concentrations of thiosulfate and polythionate were determined as soon as possible (generally a few to several hours after sample collection) by ion chromatography in an on-site mobile laboratory vehicle. Total dissolved iron and ferrous iron concentrations often were measured on-site in the

  7. Cognitive Factors that Impact Learning in the Field: Observations from an REU Project on Precambrian Rocks of Yellowstone National Park

    Science.gov (United States)

    Henry, D.; Mogk, D. W.; Goodwin, C.

    2011-12-01

    Field work requires cognitive processing on many different levels, and constitutes a powerful and important learning environment. To be effective and meaningful, the context of field work must be fully understood in terms of key research questions, earlier published work, regional geology, geologic history, and geologic processes. Scale(s) of observation and sample selection methods and strategies must be defined. Logistical decisions must be made about equipment needed, points of access, and navigation in the field. Professional skills such as field note-taking, measuring structural data, and rock descriptions must be employed, including appropriate use of field tools. Interpretations of geologic features in the field must be interpreted through recall of concepts from the geologic knowledge base (e.g. crystallization history of igneous rocks interpreted through phase diagrams). Field workers need to be able to self-monitor and self-regulate their actions (metacognitively), and make adjustments to daily plans as needed. The results of field work must be accurately and effectively communicated to other geoscientists. Personal and professional ethics and values are brought to bear as decisions are made about whether or not the work has been satisfactorily completed at a field site. And, all of this must be done against a back drop of environmental factors that affect the ability to do this work (e.g. inclement weather, bears, impassable landscapes). The simultaneous relevance of all these factors creates a challenging, but rewarding environment for learning on many different scales. During our REU project to study the Precambrian rocks in the back country of Yellowstone National Park (YNP), we considered these cognitive factors in designing our project curriculum. To reduce the "novelty space" of the project a website was developed that described the project goals and expected outcomes, introduced primary literature, and alerted students about the physical demands

  8. Sulfur geochemistry of hydrothermal waters in Yellowstone National Park, Wyoming, USA. III. An anion-exchange resin technique for sampling and preservation of sulfoxyanions in natural waters

    Directory of Open Access Journals (Sweden)

    Ball James W

    2003-06-01

    Full Text Available A sampling protocol for the retention, extraction, and analysis of sulfoxyanions in hydrothermal waters has been developed in the laboratory and tested at Yellowstone National Park and Green Lake, NY. Initial laboratory testing of the anion-exchange resin Bio-Rad™ AG1-X8 indicated that the resin was well suited for the sampling, preservation, and extraction of sulfate and thiosulfate. Synthetic solutions containing sulfate and thiosulfate were passed through AG1-X8 resin columns and eluted with 1 and 3 M KCl, respectively. Recovery ranged from 89 to 100%. Comparison of results for water samples collected from five pools in Yellowstone National Park between on-site IC analysis (U.S. Geological Survey mobile lab and IC analysis of resin-stored sample at SUNY-Stony Brook indicates 96 to 100% agreement for three pools (Cinder, Cistern, and an unnamed pool near Cistern and 76 and 63% agreement for two pools (Sulfur Dust and Frying Pan. Attempts to extract polythionates from the AG1-X8 resin were made using HCl solutions, but were unsuccessful. Bio-Rad™ AG2-X8, an anion-exchange resin with weaker binding sites than the AG1-X8 resin, is better suited for polythionate extraction. Sulfate and thiosulfate extraction with this resin has been accomplished with KCl solutions of 0.1 and 0.5 M, respectively. Trithionate and tetrathionate can be extracted with 4 M KCl. Higher polythionates can be extracted with 9 M hydrochloric acid. Polythionate concentrations can then be determined directly using ion chromatographic methods, and laboratory results indicate recovery of up to 90% for synthetic polythionate solutions using AG2-X8 resin columns.

  9. Sulfur geochemistry of hydrothermal waters in Yellowstone National Park, Wyoming, USA. III. An anion-exchange resin technique for sampling and preservation of sulfoxyanions in natural waters

    Science.gov (United States)

    Druschel, G.K.; Schoonen, M.A.A.; Nordstorm, D.K.; Ball, J.W.; Xu, Y.; Cohn, C.A.

    2003-01-01

    A sampling protocol for the retention, extraction, and analysis of sulfoxyanions in hydrothermal waters has been developed in the laboratory and tested at Yellowstone National Park and Green Lake, NY. Initial laboratory testing of the anion-exchange resin Bio-Rad??? AG1-X8 indicated that the resin was well suited for the sampling, preservation, and extraction of sulfate and thiosulfate. Synthetic solutions containing sulfate and thiosulfate were passed through AG1-X8 resin columns and eluted with 1 and 3 M KCl, respectively. Recovery ranged from 89 to 100%. Comparison of results for water samples collected from five pools in Yellowstone National Park between on-site IC analysis (U.S. Geological Survey mobile lab) and IC analysis of resin-stored sample at SUNY-Stony Brook indicates 96 to 100% agreement for three pools (Cinder, Cistern, and an unnamed pool near Cistern) and 76 and 63% agreement for two pools (Sulfur Dust and Frying Pan). Attempts to extract polythionates from the AG1-X8 resin were made using HCl solutions, but were unsuccessful. Bio-Rad??? AG2-X8, an anion-exchange resin with weaker binding sites than the AG1-X8 resin, is better suited for polythionate extraction. Sulfate and thiosulfate extraction with this resin has been accomplished with KCl solutions of 0.1 and 0.5 M, respectively. Trithionate and tetrathionate can be extracted with 4 M KCl. Higher polythionates can be extracted with 9 M hydrochloric acid. Polythionate concentrations can then be determined directly using ion chromatographic methods, and laboratory results indicate recovery of up to 90% for synthetic polythionate solutions using AG2-X8 resin columns. ?? The Royal Society of Chemistry and the Division of Geochemistry of the American Chemical Society 2003.

  10. Integrated Geoscience Studies in the Greater Yellowstone Area - Volcanic, Tectonic, and Hydrothermal Processes in the Yellowstone Geoecosystem

    Science.gov (United States)

    Morgan, Lisa A.

    2007-01-01

    Yellowstone National Park, rimmed by a crescent of older mountainous terrain, has at its core the Quaternary Yellowstone Plateau, an undulating landscape shaped by forces of volcanism, tectonism, and later glaciation. Its spectacular hydrothermal systems cap this landscape. From 1997 through 2003, the United States Geological Survey Mineral Resources Program conducted a multidisciplinary project of Yellowstone National Park entitled Integrated Geoscience Studies of the Greater Yellowstone Area, building on a 130-year foundation of extensive field studies (including the Hayden survey of 1871, the Hague surveys of the 1880s through 1896, the studies of Iddings, Allen, and Day during the 1920s, and NASA-supported studies starting in the 1970s - now summarized in USGS Professional Paper 729 A through G) in this geologically dynamic terrain. The project applied a broad range of scientific disciplines and state-of-the-art technologies targeted to improve stewardship of the unique natural resources of Yellowstone and enable the National Park Service to effectively manage resources, protect park visitors from geologic hazards, and better educate the public on geologic processes and resources. This project combined a variety of data sets in characterizing the surficial and subsurface chemistry, mineralogy, geology, geophysics, and hydrothermal systems in various parts of the park. The sixteen chapters presented herein in USGS Professional Paper 1717, Integrated Geoscience Studies in the Greater Yellowstone Area - Volcanic, Tectonic, and Hydrothermal Processes in the Yellowstone Geoecosystem, can be divided into four major topical areas: (1) geologic studies, (2) Yellowstone Lake studies, (3) geochemical studies, and (4) geophysical studies. The geologic studies include a paper by Ken Pierce and others on the influence of the Yellowstone hotspot on landscape formation, the ecological effects of the hotspot, and the human experience and human geography of the greater

  11. The question of recharge to the deep thermal reservoir underlying the geysers and hot springs of Yellowstone National Park: Chapter H in Integrated geoscience studies in Integrated geoscience studies in the Greater Yellowstone Area—Volcanic, tectonic, and hydrothermal processes in the Yellowstone geoecosystem

    Science.gov (United States)

    Rye, Robert O.; Truesdell, Alfred Hemingway; Morgan, Lisa A.

    2007-01-01

    The extraordinary number, size, and unspoiled beauty of the geysers and hot springs of Yellowstone National Park (the Park) make them a national treasure. The hydrology of these special features and their relation to cold waters of the Yellowstone area are poorly known. In the absence of deep drill holes, such information is available only indirectly from isotope studies. The δD-δ18O values of precipitation and cold surface-water and ground-water samples are close to the global meteoric water line (Craig, 1961). δD values of monthly samples of rain and snow collected from 1978 to 1981 at two stations in the Park show strong seasonal variations, with average values for winter months close to those for cold waters near the collection sites. δD values of more than 300 samples from cold springs, cold streams, and rivers collected during the fall from 1967 to 1992 show consistent north-south and east-west patterns throughout and outside of the Park, although values at a given site vary by as much as 8 ‰ from year to year. These data, along with hot-spring data (Truesdell and others, 1977; Pearson and Truesdell, 1978), show that ascending Yellowstone thermal waters are modified isotopically and chemically by a variety of boiling and mixing processes in shallow reservoirs. Near geyser basins, shallow recharge waters from nearby rhyolite plateaus dilute the ascending deep thermal waters, particularly at basin margins, and mix and boil in reservoirs that commonly are interconnected. Deep recharge appears to derive from a major deep thermal-reservoir fluid that supplies steam and hot water to all geyser basins on the west side of the Park and perhaps in the entire Yellowstone caldera. This water (T ≥350°C; δD = –149±1 ‰) is isotopically lighter than all but the farthest north, highest altitude cold springs and streams and a sinter-producing warm spring (δD = –153 ‰) north of the Park. Derivation of this deep fluid solely from present-day recharge is

  12. Carbon dioxide and hydrogen sulfide degassing and cryptic thermal input to Brimstone Basin, Yellowstone National Park, Wyoming

    Science.gov (United States)

    Bergfeld, D.; Evans, William C.; Lowenstern, J. B.; Hurwitz, S.

    2012-01-01

    Brimstone Basin, a remote area of intense hydrothermal alteration a few km east of the Yellowstone Caldera, is rarely studied and has long been considered to be a cold remnant of an ancient hydrothermal system. A field campaign in 2008 confirmed that gas emissions from the few small vents were cold and that soil temperatures in the altered area were at background levels. Geochemical and isotopic evidence from gas samples (3He/4He ~ 3RA, δ13C-CO2 ~ − 3‰) however, indicate continuing magmatic gas input to the system. Accumulation chamber measurements revealed a surprisingly large diffuse flux of CO2 (~ 277 t d-1) and H2S (0.6 t d-1). The flux of CO2 reduces the 18O content of the overlying cold groundwater and related stream waters relative to normal meteoric waters. Simple isotopic modeling reveals that the CO2 likely originates from geothermal water at a temperature of 93 ± 19 °C. These results and the presence of thermogenic hydrocarbons (C1:C2 ~ 100 and δ13C-CH4 = − 46.4 to − 42.8‰) in gases require some heat source at depth and refute the assumption that this is a “fossil” hydrothermal system.

  13. Age-specific survival of reintroduced swift fox in Badlands National Park and surrounding lands

    Science.gov (United States)

    Sasmal, Indrani; Klaver, Robert W.; Jenks, Jonathan A.; Schroeder, Greg M.

    2016-01-01

    In 2003, a reintroduction program was initiated at Badlands National Park (BNP), South Dakota, USA, with swift foxes (Vulpes velox) translocated from Colorado and Wyoming, USA, as part of a restoration effort to recover declining swift fox populations throughout its historical range. Estimates of age-specific survival are necessary to evaluate the potential for population growth of reintroduced populations. We used 7 years (2003–2009) of capture–recapture data of 243 pups, 29 yearlings, and 69 adult swift foxes at BNP and the surrounding area to construct Cormack–Jolly–Seber model estimates of apparent survival within a capture–mark–recapture framework using Program MARK. The best model for estimating recapture probabilities included no differences among age classes, greater recapture probabilities during early years of the monitoring effort than later years, and variation among spring, winter, and summer. Our top ranked survival model indicated pup survival differed from that of yearlings and adults and varied by month and year. The apparent annual survival probability of pups (0.47, SE = 0.10) in our study area was greater than the apparent annual survival probability of yearlings and adults (0.27, SE = 0.08). Our results indicate low survival probabilities for a reintroduced population of swift foxes in the BNP and surrounding areas. Management of reintroduced populations and future reintroductions of swift foxes should consider the effects of relative low annual survival on population demography.

  14. Complete genome sequences of Geobacillus sp. Y412MC52, a xylan-degrading strain isolated from obsidian hot spring in Yellowstone National Park.

    Science.gov (United States)

    Brumm, Phillip; Land, Miriam L; Hauser, Loren J; Jeffries, Cynthia D; Chang, Yun-Juan; Mead, David A

    2015-01-01

    Geobacillus sp. Y412MC52 was isolated from Obsidian Hot Spring, Yellowstone National Park, Montana, USA under permit from the National Park Service. The genome was sequenced, assembled, and annotated by the DOE Joint Genome Institute and deposited at the NCBI in December 2011 (CP002835). Based on 16S rRNA genes and average nucleotide identity, Geobacillus sp. Y412MC52 and the related Geobacillus sp. Y412MC61 appear to be members of a new species of Geobacillus. The genome of Geobacillus sp. Y412MC52 consists of one circular chromosome of 3,628,883 bp, an average G + C content of 52 % and one circular plasmid of 45,057 bp and an average G + C content of 45 %. Y412MC52 possesses arabinan, arabinoglucuronoxylan, and aromatic acid degradation clusters for degradation of hemicellulose from biomass. Transport and utilization clusters are also present for other carbohydrates including starch, cellobiose, and α- and β-galactooligosaccharides.

  15. Complete genome sequence of Geobacillus thermoglucosidasius C56-YS93, a novel biomass degrader isolated from obsidian hot spring in Yellowstone National Park.

    Science.gov (United States)

    Brumm, Phillip J; Land, Miriam L; Mead, David A

    2015-01-01

    Geobacillus thermoglucosidasius C56-YS93 was one of several thermophilic organisms isolated from Obsidian Hot Spring, Yellowstone National Park, Montana, USA under permit from the National Park Service. Comparison of 16 S rRNA sequences confirmed the classification of the strain as a G. thermoglucosidasius species. The genome was sequenced, assembled, and annotated by the DOE Joint Genome Institute and deposited at the NCBI in December 2011 (CP002835). The genome of G. thermoglucosidasius C56-YS93 consists of one circular chromosome of 3,893,306 bp and two circular plasmids of 80,849 and 19,638 bp and an average G + C content of 43.93 %. G. thermoglucosidasius C56-YS93 possesses a xylan degradation cluster not found in the other G. thermoglucosidasius sequenced strains. This cluster appears to be related to the xylan degradation cluster found in G. stearothermophilus. G. thermoglucosidasius C56-YS93 possesses two plasmids not found in the other two strains. One plasmid contains a novel gene cluster coding for proteins involved in proline degradation and metabolism, the other contains a collection of mostly hypothetical proteins.

  16. Analysing land cover and land use change in the Matobo National Park and surroundings in Zimbabwe

    Science.gov (United States)

    Scharsich, Valeska; Mtata, Kupakwashe; Hauhs, Michael; Lange, Holger; Bogner, Christina

    2016-04-01

    Natural forests are threatened worldwide, therefore their protection in National Parks is essential. Here, we investigate how this protection status affects the land cover. To answer this question, we analyse the surface reflectance of three Landsat images of Matobo National Park and surrounding in Zimbabwe from 1989, 1998 and 2014 to detect changes in land cover in this region. To account for the rolling countryside and the resulting prominent shadows, a topographical correction of the surface reflectance was required. To infer land cover changes it is not only necessary to have some ground data for the current satellite images but also for the old ones. In particular for the older images no recent field study could help to reconstruct these data reliably. In our study we follow the idea that land cover classes of pixels in current images can be transferred to the equivalent pixels of older ones if no changes occurred meanwhile. Therefore we combine unsupervised clustering with supervised classification as follows. At first, we produce a land cover map for 2014. Secondly, we cluster the images with clara, which is similar to k-means, but suitable for large data sets. Whereby the best number of classes were determined to be 4. Thirdly, we locate unchanged pixels with change vector analysis in the images of 1989 and 1998. For these pixels we transfer the corresponding cluster label from 2014 to 1989 and 1998. Subsequently, the classified pixels serve as training data for supervised classification with random forest, which is carried out for each image separately. Finally, we derive land cover classes from the Landsat image in 2014, photographs and Google Earth and transfer them to the other two images. The resulting classes are shrub land; forest/shallow waters; bare soils/fields with some trees/shrubs; and bare light soils/rocks, fields and settlements. Subsequently the three different classifications are compared and land changes are mapped. The main changes are

  17. Aerial radiological survey of the Brookhaven National Laboratory and surrounding area, May 1980

    International Nuclear Information System (INIS)

    Hobaugh, J.L.; Steiner, P.A.

    1984-01-01

    An aerial radiological survey was performed from 15 to 18 May 1980 over approximately a 21-square-kilometer (8-square-mile) area surrounding the Brookhaven National Laboratory (BNL). BNL is located in the center of Suffolk County, Long Island, New York. All gamma radiation data were collected by flying north-south lines spaced 91 meters (300 feet) apart at an altitude of 61 meters (200 feet) above ground level. A total of 17 anomalous areas were identified. The dominant isotopes found over these areas were cesium-137, manganese-54, and cobalt-60. All anomalies identified by the aerial measurements were correlated to site activities and storage facilities. 4 references, 18 figures, 1 table

  18. A checklist of plant and animal species at Los Alamos National Laboratory and surrounding areas

    Energy Technology Data Exchange (ETDEWEB)

    Hinojosa, H. [comp.

    1998-02-01

    Past and current members of the Biology Team (BT) of the Ecology Group have completed biological assessments (BAs) for all of the land that comprises Los Alamos National Laboratory (LANL). Within these assessments are lists of plant and animal species with the potential to exist on LANL lands and the surrounding areas. To compile these lists, BT members examined earlier published and unpublished reports, surveys, and data bases that pertained to the biota of this area or to areas that are similar. The species lists that are contained herein are compilations of the lists from these BAs, other lists that were a part of the initial research for the performance of these BAs, and more recent surveys.

  19. Geologic field-trip guide to the volcanic and hydrothermal landscape of the Yellowstone Plateau

    Science.gov (United States)

    Morgan Morzel, Lisa Ann; Shanks, W. C. Pat; Lowenstern, Jacob B.; Farrell, Jamie M.; Robinson, Joel E.

    2017-11-20

    Yellowstone National Park, a nearly 9,000 km2 (~3,468 mi2) area, was preserved in 1872 as the world’s first national park for its unique, extraordinary, and magnificent natural features. Rimmed by a crescent of older mountainous terrain, Yellowstone National Park has at its core the Quaternary Yellowstone Plateau, an undulating landscape shaped by forces of late Cenozoic explosive and effusive volcanism, on-going tectonism, glaciation, and hydrothermal activity. The Yellowstone Caldera is the centerpiece of the Yellowstone Plateau. The Yellowstone Plateau lies at the most northeastern front of the 17-Ma Yellowstone hot spot track, one of the few places on Earth where time-transgressive processes on continental crust can be observed in the volcanic and tectonic (faulting and uplift) record at the rate and direction predicted by plate motion. Over six days, this field trip presents an intensive overview into volcanism, tectonism, and hydrothermal activity on the Yellowstone Plateau (fig. 1). Field stops are linked directly to conceptual models related to monitoring of the various volcanic, geochemical, hydrothermal, and tectonic aspects of the greater Yellowstone system. Recent interest in young and possible future volcanism at Yellowstone as well as new discoveries and synthesis of previous studies, (for example, tomographic, deformation, gas, aeromagnetic, bathymetric, and seismic surveys), provide a framework in which to discuss volcanic, hydrothermal, and seismic activity in this dynamic region.

  20. Water-chemistry and on-site sulfur-speciation data for selected springs in Yellowstone National Park, Wyoming, 1994-1995

    Science.gov (United States)

    Ball, James W.; Nordstrom, D. Kirk; Cunningham, Kirk M.; Schoonen, Martin A.; Xu, Yong; DeMonge, Jennifer M.

    1998-01-01

    Forty-two water analyses are reported for samples collected at 8 hot springs and their overflow drainages, two geysers, and two ambient-temperature acid streams in Yellowstone National Park during 1994-95. These water samples were collected and analyzed as part of the initial research investigations on sulfur redox speciation in the hot springs of Yellowstone and to document chemical changes in overflows that affect major ions, redox species, and trace elements. The sulfur redox speciation research is a collaboration between the State University of New York (SUNY) at Stony Brook and the U.S. Geological Survey (USGS). Four hot springs, Ojo Caliente, Azure, Frying Pan, and Angel Terrace, were studied in detail. Analyses were performed adjacent to the sampling site or in an on-site mobile lab truck constructed by the USGS, or later in a USGS laboratory. Water temperature, specific conductance, pH, Eh, D.O., and dissolved H2S were determined adjacent to the sample source at the time of sampling. Alkalinity and F- were determined on-site on the day of sample collection. Thiosulfate and polythionates were determined as soon as possible (minutes to hours later) by ion chromatography (IC). Other major anions (Cl-, SO4 2-, Br-) also were determined on-site by IC within two days of sample collection. Ammonium, Fe(II), and Fe(total) were determined on-site by ultraviolet/visible spectrophotometry within two days of sample collection. Later in the USGS laboratory, densities were determined. Concentrations of Ca, Mg, Li, Na, and K were determined by flame atomic absorption and emission (Na, K) spectrometry. Concentrations of Al, As, B, Ba, Be, Ca, Cd, Co, Cr, Cu, Fe(total), K, Mg, Mn, Na, Ni, Pb, Si, Sr, V, and Zn were determined by inductively-coupled plasma optical emission spectrometry. Trace concentrations of Al and Mg were determined by Zeeman-corrected graphite furnace atomic absorption spectrometry. Three important conclusions from the sampling and analyses are: (1

  1. Precipitation Depth-Duration-Frequency Analysis for the Nevada National Security Site and Surrounding Areas

    Energy Technology Data Exchange (ETDEWEB)

    Chen, Li [Desert Research Inst. (DRI), Las Vegas, NV (United States). Division of Hydrologic Sciences; Miller, Julianne J. [Desert Research Inst. (DRI), Las Vegas, NV (United States). Division of Hydrologic Sciences

    2016-08-01

    Accurate precipitation frequency data are important for Environmental Management Soils Activities on the Nevada National Security Site (NNSS). These data are important for environmental assessments performed for regulatory closure of Soils Corrective Action Unit (CAU) Sites, as well as engineering mitigation designs and post-closure monitoring strategies to assess and minimize potential contaminant migration from Soils CAU Sites. Although the National Oceanic and Atmospheric Administration (NOAA) Atlas 14 (Bonnin et al., 2011) provides precipitation frequency data for the NNSS area, the NNSS-specific observed precipitation data were not consistent with the NOAA Atlas 14 predicted data. This is primarily due to the NOAA Atlas 14 products being produced from analyses without including the approximately 30 NNSS precipitation gage records, several of which approach or exceed 50 year of record. Therefore, a study of precipitation frequency that incorporated the NNSS precipitation gage records into the NOAA Atlas 14 dataset, was performed specifically for the NNSS to derive more accurate site-specific precipitation data products. Precipitation frequency information, such as the depth-duration-frequency (DDF) relationships, are required to generate synthetic standard design storm hydrographs and assess actual precipitation events. In this study, the actual long-term NNSS precipitation gage records, some of which are the longest gage records in southern and central Nevada, were analyzed to allow for more accurate precipitation DDF estimates to be developed for the NNSS. Gridded maps of precipitation frequency for the NNSS and surrounding areas were then produced.

  2. Pyrobaculum Yellowstonensis Strain WP30 Respires On Elemental Sulfur And/or Arsenate in Circumneutral Sulfidic Sediments of Yellowstone National Park

    Energy Technology Data Exchange (ETDEWEB)

    Jay, Z.; Beam, Jake; Dohnalkova, Alice; Lohmayer, R.; Bodle, B.; Planer-Friedrich, B.; Romine, Margaret F.; Inskeep, William

    2015-09-15

    Thermoproteales populations (phylum Crenarchaeota) are abundant in high-25 temperature (>70° C) environments of Yellowstone National Park (YNP) and are important in mediating biogeochemical cycles of sulfur, arsenic and carbon. The objectives of this study were to determine specific physiological attributes of the isolate Pyrobaculum yellowstonensis strain WP30, which was obtained from an elemental sulfur sediment (Joseph’s Coat Hot Spring [JCHS]; 80 °C; pH 6.1), and relate this organism to geochemical processes occurring in situ. Strain WP30 is a chemoheterotroph that utilizes organic carbon as a source of carbon and electrons and requires elemental sulfur and/or arsenic as electron acceptors. Growth in the presence of elemental sulfur and arsenate resulted in the production of thioarsenates and polysulfides relative to sterile controls. The complete genome of this organism was sequenced (1.99 Mb, 58 % G+C) and revealed numerous metabolic pathways for the degradation of carbohydrates, amino acids and lipids, multiple dimethylsulfoxide molybdopterin (DMSO-MPT) oxidoreductase genes, which are implicated in the reduction of sulfur and arsenic, and pathways for the de novo synthesis of nearly all required cofactors and metabolites. Comparative genomics of P. yellowstonensis versus assembled metagenome sequence from JCHS showed that this organisms is highly-related (~95% average nucleotide identity) to in situ populations. The physiological attributes and metabolic capabilities of P. yellowstonensis provide importanat information towards understanding the distribution and function of these populations in YNP.

  3. Carbon and oxygen isotope analysis of leaf biomass reveals contrasting photosynthetic responses to elevated CO2 near geologic vents in Yellowstone National Park

    Directory of Open Access Journals (Sweden)

    D. G. Williams

    2009-01-01

    Full Text Available In this study we explore the use of natural CO2 emissions in Yellowstone National Park (YNP in Wyoming, USA to study responses of natural vegetation to elevated CO2 levels. Radiocarbon (14C analysis of leaf biomass from a conifer (Pinus contortus; lodgepole pine and an invasive, non-native herb (Linaria dalmatica; Dalmation toadflax was used to trace the inputs of vent CO2 and quantify assimilation-weighted CO2 concentrations experienced by individual plants near vents and in comparable locations with no geologic CO2 exposure. The carbon and oxygen isotopic composition and nitrogen percent of leaf biomass from the same plants was used to investigate photosynthetic responses of these plants to naturally elevated atmospheric CO2 concentrations. The coupled shifts in carbon and oxygen isotope values suggest that dalmation toadflax responded to elevated CO2 exposure by increasing stomatal conductance with no change in photosynthetic capacity and lodgepole pine apparently responded by decreasing stomatal conductance and photosynthetic capacity. Lodgepole pine saplings exposed to elevated levels of CO2 likewise had reduced leaf nitrogen concentrations compared to plants with no enhanced CO2 exposure, further suggesting widespread and dominant conifer down-regulated photosynthetic capacity under elevated CO2 levels near geologic vents.

  4. Formaldehyde as a carbon and electron shuttle between autotroph and heterotroph populations in acidic hydrothermal vents of Norris Geyser Basin, Yellowstone National Park

    Energy Technology Data Exchange (ETDEWEB)

    Moran, James J.; Whitmore, Laura M.; Isern, Nancy G.; Romine, Margaret F.; Riha, Krystin M.; Inskeep, William P.; Kreuzer, Helen W.

    2016-03-19

    The Norris Geyser Basin in Yellowstone National Park contains a large number of hydrothermal systems, which host microbial populations supported by primary productivity associated with a suite of chemolithotrophic metabolisms. We demonstrate that Metallosphaera yellowstonesis MK1, a facultative autotrophic archaeon isolated from a hyperthermal acidic hydrous ferric oxide (HFO) spring in Norris Geyser Basin, excretes formaldehyde during autotrophic growth. To determine the fate of formaldehyde in this low organic carbon environment, we incubated native microbial mat (containing M. yellowstonensis) from a HFO spring with 13C-formaldehyde. Isotopic analysis of incubation-derived CO2 and biomass showed that formaldehyde was both oxidized and assimilated by members of the community. Autotrophy, formaldehyde oxidation, and formaldehyde assimilation displayed different sensitivities to chemical inhibitors, suggesting that distinct sub-populations in the mat selectively perform these functions. Our results demonstrate that electrons originally resulting from iron oxidation can energetically fuel autotrophic carbon fixation and associated formaldehyde excretion, and that formaldehyde is both oxidized and assimilated by different organisms within the native microbial community. Thus, formaldehyde can effectively act as a carbon and electron shuttle connecting the autotrophic, iron oxidizing members with associated heterotrophic members in the HFO community.

  5. Gray Wolves as Climate Change Buffers in Yellowstone

    OpenAIRE

    Wilmers Christopher C; Getz Wayne M; Getz Wayne M

    2005-01-01

    Understanding the mechanisms by which climate and predation patterns by top predators co-vary to affect community structure accrues added importance as humans exert growing influence over both climate and regional predator assemblages. In Yellowstone National Park, winter conditions and reintroduced gray wolves (Canis lupus) together determine the availability of winter carrion on which numerous scavenger species depend for survival and reproduction. As climate changes in Yellowstone, therefo...

  6. Gray wolves as climate change buffers in Yellowstone.

    OpenAIRE

    Christopher C Wilmers; Wayne M Getz

    2005-01-01

    Understanding the mechanisms by which climate and predation patterns by top predators co-vary to affect community structure accrues added importance as humans exert growing influence over both climate and regional predator assemblages. In Yellowstone National Park, winter conditions and reintroduced gray wolves (Canis lupus) together determine the availability of winter carrion on which numerous scavenger species depend for survival and reproduction. As climate changes in Yellowstone, therefo...

  7. 76 FR 18040 - Amendment of Class E Airspace; West Yellowstone, MT

    Science.gov (United States)

    2011-04-01

    ... airspace at Yellowstone Airport, West Yellowstone, MT, to accommodate aircraft using Instrument Landing... the earth. * * * * * ANM MT E5 West Yellowstone, MT [Amended] West Yellowstone, Yellowstone Airport...

  8. Animals and their products utilized as medicines by the inhabitants surrounding the Ranthambhore National Park, India

    Directory of Open Access Journals (Sweden)

    Jaroli DP

    2006-11-01

    Full Text Available Abstract The present ethnozoological study describes the traditional knowledge related to the use of different animals and animal-derived products as medicines by the inhabitants of villages surrounding the Ranthambhore National Park of India (Bawaria, Mogya, Meena, which is well known for its very rich biodiversity. The field survey was conducted from May to July 2005 by performing interviews through structured questionnaires with 24 informants (16 men and 8 women, who provided information regarding therapeutic uses of animals. A total of 15 animals and animal products were recorded and they are used for different ethnomedical purposes, including tuberculosis, asthma, paralysis, jaundice, earache, constipation, weakness, snake poisoning. The zootherapeutic knowledge was mostly based on domestic animals, but some protected species like the collared dove (Streptopelia sp., hard shelled turtle (Kachuga tentoria, sambhar (Cervus unicolor were also mentioned as important medicinal resources. We would suggest that this kind of neglected traditional knowledge should be included into the strategies of conservation and management of faunistic resources in the investigated area.

  9. An aerial radiological survey of the Sandia National Laboratories and surrounding area

    International Nuclear Information System (INIS)

    Riedhauser, S.R.

    1994-06-01

    A team from the Remote Sensing Laboratory conducted an aerial radiological survey of the area surrounding the Sandia National Laboratories and Kirtland Air Force Base in Albuquerque, New Mexico, during March and April 1993. The survey team measured the terrestrial gamma radiation at the site to determine the levels of natural and man-made radiation. This survey includes the areas covered by a previous survey in 1981. The results of the aerial survey show a background exposure rate which varies between 5 and 18 μR/h plus an approximate 6 μR/h contribution from cosmic rays. The major radioactive isotopes found in this survey were: potassium-40, thallium-208, bismuth-214, and actinium-228, which are all naturally-occurring isotopes, and cobalt-60, cesium-137, and excess amounts of thallium-208 and actinium-228, which are due to human actions in the survey area. In regions away from man-made activity, the exposure rates inferred from this survey's gamma ray measurements agree almost exactly with the exposure rates inferred from the 1981 survey. In addition to the aerial measurements, another survey team conducted in situ and soil sample radiation measurements at three sites within the survey perimeter. These ground-based measurements agree with the aerial measurements within ± 5%

  10. 77 FR 74027 - Winter Use Plan, Final Environmental Impact Statement Amended Record of Decision, Yellowstone...

    Science.gov (United States)

    2012-12-12

    ...] Winter Use Plan, Final Environmental Impact Statement Amended Record of Decision, Yellowstone National... Availability of Amended Record of Decision for the Final Environmental Impact Statement for a Winter Use Plan... Record of Decision for the Winter Use Plan for Yellowstone National Park, located in Idaho, Montana, and...

  11. Potential application of radiogenic isotopes and geophysical methods to understand the hydrothermal dystem of the Upper Geyser Basin, Yellowstone National Park

    Science.gov (United States)

    Paces, James B.; Long, Andrew J.; Koth, Karl R.

    2015-01-01

    Numerous geochemical and geophysical studies have been conducted at Yellowstone National Park to better understand the hydrogeologic processes supporting the thermal features of the Park. This report provides the first 87Sr/86Sr and 234U/238U data for thermal water from the Upper Geyser Basin (UGB) intended to evaluate whether heavy radiogenic isotopes might provide insight to sources of groundwater supply and how they interact over time and space. In addition, this report summarizes previous geophysical studies made at Yellowstone National Park and provides suggestions for applying non-invasive ground and airborne studies to better understand groundwater flow in the subsurface of the UGB. Multiple samples from Old Faithful, Aurum, Grand, Oblong, and Daisy geysers characterized previously for major-ion concentrations and isotopes of water (δ2H, δ18O, and 3H) were analyzed for Sr and U isotopes. Concentrations of dissolved Sr and U are low (4.3–128 ng g-1 Sr and 0.026–0.0008 ng g-1 U); consequently only 87Sr/86Sr data are reported for most samples. Values of 87Sr/86Sr for most geysers remained uniform between April and September 2007, but show large increases in all five geysers between late October 2007 and early April, 2008. By late summer of 2008, 87Sr/86Sr values returned to values similar to those observed a year earlier. Similar patterns are not present in major-ion data measured on the same samples. Furthermore, large geochemical differences documented between geysers are not observed in 87Sr/86Sr data, although smaller differences between sites may be present. Sr-isotope data are consistent with a stratified hydrologic system where water erupted in spring and summer of 2007 and summer of 2008 equilibrated with local intracaldera rhyolite flows at shallower depths. Water erupted between October 2007 and April 2008 includes greater amounts of groundwater that circulated deep enough to acquire a radiogenic 87Sr/86Sr, most likely from Archean basement

  12. Insights into archaeal evolution and symbiosis from the genomes of a nanoarchaeon and its inferred crenarchaeal host from Obsidian Pool, Yellowstone National Park.

    Science.gov (United States)

    Podar, Mircea; Makarova, Kira S; Graham, David E; Wolf, Yuri I; Koonin, Eugene V; Reysenbach, Anna-Louise

    2013-04-22

    A single cultured marine organism, Nanoarchaeum equitans, represents the Nanoarchaeota branch of symbiotic Archaea, with a highly reduced genome and unusual features such as multiple split genes. The first terrestrial hyperthermophilic member of the Nanoarchaeota was collected from Obsidian Pool, a thermal feature in Yellowstone National Park, separated by single cell isolation, and sequenced together with its putative host, a Sulfolobales archaeon. Both the new Nanoarchaeota (Nst1) and N. equitans lack most biosynthetic capabilities, and phylogenetic analysis of ribosomal RNA and protein sequences indicates that the two form a deep-branching archaeal lineage. However, the Nst1 genome is more than 20% larger, and encodes a complete gluconeogenesis pathway as well as the full complement of archaeal flagellum proteins. With a larger genome, a smaller repertoire of split protein encoding genes and no split non-contiguous tRNAs, Nst1 appears to have experienced less severe genome reduction than N. equitans. These findings imply that, rather than representing ancestral characters, the extremely compact genomes and multiple split genes of Nanoarchaeota are derived characters associated with their symbiotic or parasitic lifestyle. The inferred host of Nst1 is potentially autotrophic, with a streamlined genome and simplified central and energetic metabolism as compared to other Sulfolobales. Comparison of the N. equitans and Nst1 genomes suggests that the marine and terrestrial lineages of Nanoarchaeota share a common ancestor that was already a symbiont of another archaeon. The two distinct Nanoarchaeota-host genomic data sets offer novel insights into the evolution of archaeal symbiosis and parasitism, enabling further studies of the cellular and molecular mechanisms of these relationships. This article was reviewed by Patrick Forterre, Bettina Siebers (nominated by Michael Galperin) and Purification Lopez-Garcia.

  13. Evidence for high-temperature in situ nifH transcription in an alkaline hot spring of Lower Geyser Basin, Yellowstone National Park.

    Science.gov (United States)

    Loiacono, Sara T; Meyer-Dombard, D'Arcy R; Havig, Jeff R; Poret-Peterson, Amisha T; Hartnett, Hilairy E; Shock, Everett L

    2012-05-01

    Genes encoding nitrogenase (nifH) were amplified from sediment and photosynthetic mat samples collected in the outflow channel of Mound Spring, an alkaline thermal feature in Yellowstone National Park. Results indicate the genetic capacity for nitrogen fixation over the entire range of temperatures sampled (57.2°C to 80.2°C). Amplification of environmental nifH transcripts revealed in situ expression of nifH genes at temperatures up to 72.7°C. However, we were unable to amplify transcripts of nifH at the higher-temperature locations (> 72.7°C). These results indicate that microbes at the highest temperature sites contain the genetic capacity to fix nitrogen, yet either do not express nifH or do so only transiently. Field measurements of nitrate and ammonium show fixed nitrogen limitation as temperature decreases along the outflow channel, suggesting nifH expression in response to the downstream decrease in bioavailable nitrogen. Nitrogen stable isotope values of Mound Spring sediment communities further support geochemical and genetic data. DNA and cDNA nifH amplicons form several unique phylogenetic clades, some of which appear to represent novel nifH sequences in both photosynthetic and chemosynthetic microbial communities. This is the first report of in situ nifH expression in strictly chemosynthetic zones of terrestrial (non-marine) hydrothermal systems, and sets a new upper temperature limit (72.7°C) for nitrogen fixation in alkaline, terrestrial hydrothermal environments. © 2012 Society for Applied Microbiology and Blackwell Publishing Ltd.

  14. Paleomagnetic results from Tertiary volcanic strata and intrusions, Absaroka Volcanic Supergroup, Yellowstone National Park and vicinity: Contributions to the North American apparent polar wander path

    Science.gov (United States)

    Harlan, S.S.; Morgan, L.A.

    2010-01-01

    We report paleomagnetic and rock magnetic data from volcanic, volcaniclastic, and intrusive rocks of the 55-44Ma Absaroka Volcanic Supergroup (AVS) exposed along the northeastern margin of Yellowstone National Park and adjacent areas. Demagnetization behavior and rock magnetic experiments indicate that the remanence in most samples is carried by low-Ti titanomagnetite, although high-coercivity phases are present in oxidized basalt flows. Paleomagnetic demagnetization and rock magnetic characteristics, the presence of normal and reverse polarity sites, consistency with previous results, and positive conglomerate tests suggest that the observed remanences are primary thermoremanent magnetizations of Eocene age (c. 50Ma). An in situ grand-mean for 22 individual site- or cooling-unit means from this study that yield acceptable data combined with published data from Independence volcano yields a declination of 347.6?? and inclination of 59.2?? (k=21.8, ??95=6.8??) and a positive reversal test. Averaging 21 virtual geomagnetic poles (VGPs) that are well-grouped yields a mean at 137.1??E, 82.5??N (K=17.6, A95=7.8??), similar to results previously obtained from published studies from the AVS. Combining the VGPs from our study with published data yields a combined AVS pole at 146.3??E, 83.1??N (K=13.5, A95=6.2??, N=42 VGPs). Both poles are indistinguishable from c. 50Ma cratonic and synthetic reference poles for North America, and demonstrate the relative stability of this part of the Cordillera with respect to the craton. ?? 2009 Elsevier B.V.

  15. Ectomycorrhizal Community Structure and Soil Characteristics of Mature Lodgepole Pine (Pinus Contorta) and Adjacent Stands of Old Growth Mixed Conifer in Yellowstone National Park, Wyoming USA

    Science.gov (United States)

    Douglas, Robert B.; Parker, V. Thomas; Cullings, Kenneth W.; Sun, Sidney (Technical Monitor)

    2003-01-01

    Forest development patterns following disturbance are known to influence the physical and chemical attributes of soils at different points in time. Changes in soil resources are thought to have a corresponding effect on ectomycorrhizal (ECM) community structure. We used molecular methods to compare below-ground ECM species richness, composition, and abundance between adjacent stands of homogenous lodgepole pine and old growth mixed conifer in Yellowstone National Park (YNP). In each stand-type we collected soil cores to both identify mycorrhizae and assess soil chemistry. Although no statistical difference was observed in the mean number of ECM root tips per core between stand types, the total number of species identified (85 versus 35) and the mean number of species per core (8.8 +/- 0.6 versus 2.5 +/- 0.3) were significantly higher in lodgepole pine. Differences between the actual and estimated species richness levels indicated that these forest types support a high number of ECM species and that undersampling was severe. Species compositions were widely disparate between stands where only four species were shared out of a total of 116. Soil analysis also revealed that mixed conifer was significantly lower in pH, but higher in organic matter, potassium, phosphorus, and ammonium when compared to lodgepole pine stands. Species richness per core was correlated with these chemical data, however, analysis of covariance indicated that stand type was the only statistically significant factor in the observed difference in species richness. Our data suggest that ECM fungal richness increases as homogenous lodgepole pine stands grow and mature, but declines after Engelmann spruce and subalpine fir colonize. Despite difficulties linking species composition with soil chemistry, there are a variety of physical and chemical factors that could be influencing ECM community structure. Future field experiments are necessary to test some of the mechanisms potentially operating

  16. Insights into the biological source and environmental gradients shaping the distribution of H-shaped glycerol dialkyl glycerol tetraethers in Yellowstone National Park geothermal springs

    Science.gov (United States)

    Jia, C.; Xie, W.; Wang, J.; Boyd, E. S.; Zhang, C.

    2013-12-01

    Archaea are ubiquitous in natural environments. The unique tetraether lipids in archaeal membranes enable the maintenance of ion permeability across broad environmental gradients. H-shaped isoprenoid glycerol dialkyl glycerol tetraethers (H-GDGTs), in which the two biphytanyl carbon skeletons are covalently bound by a carbon-carbon bond, have been recently identified in both marine and geothermal environments. Here we report the core H-GDGTs (C-H-GDGTs) and polar H-GDGTs (P-H-GDGTs) associated with sediments sampled from geothermal springs in Yellowstone National Park and investigate their abundance in relation to environmental gradients. The abundance of C- and P-H-GDGTs exhibit strong and negative correlation with pH (P = 0.007), suggesting that H-shaped GDGTs help to maintain cell membrane fluidity in acidic environments. Reanalysis of archaeal 16S rRNA gene pyrotags published previously from (Boyd E. Hamilton T. L., Wang J., He L., Zhang C. L. 2013. The role of tetraether lipid composition in the adaptation of thermophilic archaea to acidity. Frontiers in Terrestrial Microbiology. 4: doi: 10.3389/fmicb.2013.00062) indicates that these H-GDGTs are associated with environments dominanted by Thermoplasmatales, which are thermoacidiphiles. Two equations were established to define the relationships between the abundance of H-GDGTs, the abundance of archaeal taxa based on 16S rRNA gene phylogenetic affiliations, and pH. Both equations have high predictive capacity in predicting the distribution of archaeal lipids in the geothermal system. These observations provide new insight into the biological source of H-GDGTs and suggest a prominent role for these lipids in the diversification of archaea into or out of acidic high temperature environments.

  17. Microbial iron cycling in acidic geothermal springs of Yellowstone National Park: Integrating molecular surveys, geochemical processes and isolation of novel Fe-active microorganisms

    Directory of Open Access Journals (Sweden)

    Mark A Kozubal

    2012-03-01

    Full Text Available Geochemical, molecular, and physiological analyses of microbial isolates were combined to study the geomicrobiology of acidic iron oxide mats in Yellowstone National Park (YNP. Nineteen sampling locations from 11 geothermal springs were studied ranging in temperature from 53 to 84 °C and pH 2.4 to 3.6. All iron-oxide mats exhibited high diversity of crenarchaeal sequences from the Sulfolobales, Thermoproteales, and Desulfurococcales. The predominant Sulfolobales sequences were highly similar to Metallosphaera yellowstonensis str. MK1, previously isolated from one of these sites. Other groups of archaea were consistently associated with different types of iron oxide mats, including undescribed members of the phyla Thaumarchaeota and Euryarchaeota. Bacterial sequences were dominated by relatives of Hydrogenobaculum spp. above 65-70 °C, but increased in diversity below 60 °C. Cultivation of relevant iron-oxidizing and iron-reducing microbial isolates included Sulfolobus str. MK3, Sulfobacillus str. MK2, Acidicaldus str. MK6, and a new candidate genus in the Sulfolobales referred to as Sulfolobales str. MK5. Strains MK3 and MK5 are capable of oxidizing ferrous iron autotrophically, while strain MK2 oxidizes iron mixotrophically. Similar rates of iron oxidation were observed for M. yellowstonensis str. MK1 and Sulfolobales str. MK5 cultures, and these rates are close to those measured in situ. Biomineralized phases of ferric iron varied among cultures and field sites, and included ferric oxyhydroxides, K-jarosite, goethite, hematite, and scorodite depending on geochemical conditions. Strains MK5 and MK6 are capable of reducing ferric iron under anaerobic conditions with complex carbon sources. The combination of geochemical and molecular data as well as physiological observations of isolates suggests that the community structure of acidic Fe mats is linked with Fe cycling across temperatures ranging from 53 to 88 oC.

  18. Environmental Conditions Constrain the Distribution and Diversity of Archaeal merA in Yellowstone National Park, Wyoming, U.S.A.

    Science.gov (United States)

    Wang, Y.; Boyd, E.; Crane, S.; Lu-Irving, P.; Krabbenhoft, D.; King, S.; Dighton, J.; Geesey, G.; Barkay, T.

    2011-01-01

    The distribution and phylogeny of extant protein-encoding genes recovered from geochemically diverse environments can provide insight into the physical and chemical parameters that led to the origin and which constrained the evolution of a functional process. Mercuric reductase (MerA) plays an integral role in mercury (Hg) biogeochemistry by catalyzing the transformation of Hg(II) to Hg(0). Putative merA sequences were amplified from DNA extracts of microbial communities associated with mats and sulfur precipitates from physicochemically diverse Hg-containing springs in Yellowstone National Park, Wyoming, using four PCR primer sets that were designed to capture the known diversity of merA. The recovery of novel and deeply rooted MerA lineages from these habitats supports previous evidence that indicates merA originated in a thermophilic environment. Generalized linear models indicate that the distribution of putative archaeal merA lineages was constrained by a combination of pH, dissolved organic carbon, dissolved total mercury and sulfide. The models failed to identify statistically well supported trends for the distribution of putative bacterial merA lineages as a function of these or other measured environmental variables, suggesting that these lineages were either influenced by environmental parameters not considered in the present study, or the bacterial primer sets were designed to target too broad of a class of genes which may have responded differently to environmental stimuli. The widespread occurrence of merA in the geothermal environments implies a prominent role for Hg detoxification in these environments. Moreover, the differences in the distribution of the merA genes amplified with the four merA primer sets suggests that the organisms putatively engaged in this activity have evolved to occupy different ecological niches within the geothermal gradient. ?? 2011 Springer Science+Business Media, LLC.

  19. Environmental conditions constrain the distribution and diversity of archaeal merA in Yellowstone National Park, Wyoming, U.S.A.

    Science.gov (United States)

    Wang, Yanping; Boyd, Eric; Crane, Sharron; Lu-Irving, Patricia; Krabbenhoft, David; King, Susan; Dighton, John; Geesey, Gill; Barkay, Tamar

    2011-11-01

    The distribution and phylogeny of extant protein-encoding genes recovered from geochemically diverse environments can provide insight into the physical and chemical parameters that led to the origin and which constrained the evolution of a functional process. Mercuric reductase (MerA) plays an integral role in mercury (Hg) biogeochemistry by catalyzing the transformation of Hg(II) to Hg(0). Putative merA sequences were amplified from DNA extracts of microbial communities associated with mats and sulfur precipitates from physicochemically diverse Hg-containing springs in Yellowstone National Park, Wyoming, using four PCR primer sets that were designed to capture the known diversity of merA. The recovery of novel and deeply rooted MerA lineages from these habitats supports previous evidence that indicates merA originated in a thermophilic environment. Generalized linear models indicate that the distribution of putative archaeal merA lineages was constrained by a combination of pH, dissolved organic carbon, dissolved total mercury and sulfide. The models failed to identify statistically well supported trends for the distribution of putative bacterial merA lineages as a function of these or other measured environmental variables, suggesting that these lineages were either influenced by environmental parameters not considered in the present study, or the bacterial primer sets were designed to target too broad of a class of genes which may have responded differently to environmental stimuli. The widespread occurrence of merA in the geothermal environments implies a prominent role for Hg detoxification in these environments. Moreover, the differences in the distribution of the merA genes amplified with the four merA primer sets suggests that the organisms putatively engaged in this activity have evolved to occupy different ecological niches within the geothermal gradient.

  20. Optimal wildlife management in the greater Yellowstone ecosystem: A spatiotemporal model of disease risk

    Science.gov (United States)

    South of Yellowstone National Park there are twenty-three sites where elk herds are provided supplementary feeding during the winter and spring months. Supplementary feeding of elk in the Greater Yellowstone Ecosystem (GYE) has been practiced since the early twentieth century, but the practice has b...

  1. Investigation of a Modern Incipient Stromatolite from Obsidian Pool Prime, Yellowstone National Park: Implications for Early Lithification in the Formation of Light-Dark Stromatolite Laminae

    Science.gov (United States)

    Corsetti, F. A.; Berelson, W.; Pepe-Ranney, C. P.; Mata, S. A.; Spear, J. R.

    2016-12-01

    Stromatolites have been defined multiple ways, but the presence of lamination is common to all definitions. Despite this commonality, the origin of the lamination in many ancient stromatolites remains vague. Lamination styles vary, but sub-mm light-dark couplets are common in many ancient stromatolites. Here, we investigate an actively forming incipient stromatolite from Obsidian Pool Prime (OPP), a hot spring in Yellowstone National Park, to better understand the formation of light-dark couplets similar to many ancient stromatolites in texture and structure. In the OPP stromatolites, a dense network of layer-parallel bundles of cyanobacterial filaments (a dark layer) is followed by an open network of layer-perpendicular or random filaments (a light layer) that reflect a diurnal cycle in the leading edge of the microbial mat that coats the stromatolite's surface. Silica crust encases the cyanobacterial filaments maintaining the integrity of the lamination. Bubbles formed via oxygenic photosynthesis are commonly trapped within the light layers, indicating that lithification occurs rapidly before the bubbles can collapse. The filamentous, non-heterocystous stromatoite-building cyanobacterium from OPP is most closely related to a stromatolite-building cyanobacterium from a hot spring in Japan. Once built, "tenants" from multiple microbial phyla move into the structure, mixing and mingling to produce a complicated integrated biogeochemical signal that may be difficult to untangle in ancient examples. While the cyanobacterial response to the diurnal cycle has been previously implicated in the formation of light-dark couplets, the OPP example highlights the importance of early lithification in maintaining the fabric. Thus, the presence of light-dark couplets and bubble structures may indicate very early lithification and therefore a certain degree of mineral saturation in the ancient ocean or other aquatic system, and that bubble structures, if present, may be evidence

  2. Water-Chemistry and On-Site Sulfur-Speciation Data for Selected Springs in Yellowstone National Park, Wyoming, 1996-1998

    Science.gov (United States)

    Ball, James W.; Nordstrom, D. Kirk; McCleskey, R. Blaine; Schoonen, Martin A.A.; Xu, Yong

    2001-01-01

    Fifty-eight water analyses are reported for samples collected from 19 hot springs and their overflow drainages and one ambient-temperature acid stream in Yellowstone National Park (YNP) during 1996-98. These water samples were collected and analyzed as part of research investigations on microbially mediated sulfur oxidation in stream waters and sulfur redox speciation in hot springs in YNP and chemical changes in overflow drainages that affect major ions, redox species, and trace elements. The research on sulfur redox speciation in hot springs is a collaboration with the State University of New York at Stony Brook, Northern Arizona University, and the U.S. Geological Survey (USGS). One ambient-temperature acidic stream system, Alluvium Creek and its tributaries in Brimstone Basin, was studied in detail. Analyses were performed adjacent to the sampling site, in an on-site mobile laboratory truck, or later in a USGS laboratory, depending on stability and preservability of the constituent. Water temperature, specific conductance, pH, Eh, dissolved oxygen (D.O.), and dissolved H2S were determined on-site at the time of sampling. Alkalinity and F were determined within a few days of sample collection by titration and by ion-selective electrode, respectively. Concentrations of S2O3 and SxO6 were determined as soon as possible (minutes to hours later) by ion chromatography (IC). Concentrations of Cl, SO4, and Br were determined by IC within a few days of sample collection. Concentrations of Fe(II) and Fe(total) were determined by ultraviolet/visible spectrophotometry within a few days of sample collection. Densities were determined later in the USGS laboratory. Concentrations of Li, Na, and K were determined by flame atomic absorption (Li) and emission (Na, K) spectrometry. Concentrations of Al, As(total), B, Ba, Be, Ca, Cd, Co, Cr, Cu, Fe(total), Mg, Mn, Ni, Pb, Si, Sr, V, and Zn were determined by inductively-coupled plasma optical emission spectrometry. Trace

  3. Depositional facies and aqueous-solid geochemistry of travertine-depositing hot springs (Angel Terrace, Mammoth Hot Springs, Yellowstone National Park, USA)

    Energy Technology Data Exchange (ETDEWEB)

    Fouke, B.W.; Farmer, J.D.; Des Marais, D.J.; Pratt, L.; Sturchio, N.C.; Burns, P.C.; Discipulo, M.K.

    2000-05-01

    Petrographic and geochemical analyses of travertine-depositing hot springs at Angel Terrace, Mammoth Hot Springs, Yellowstone National Park, have been used to define five depositional facies along the spring drainage system. Spring waters are expelled in the vent facies at 71 to 73 C and precipitate mounded travertine composed of aragonite needle botryoids. The apron and channel facies (43--72 C) is floored by hollow tubes composed of aragonite needle botryoids that encrust sulfide-oxidizing Aquificales bacteria. The travertine of the pond facies (30--62 C) varies in composition from aragonite needle shrubs formed at higher temperatures to ridged networks of calcite and aragonite at lower temperatures. Calcite ice sheets, calcified bubbles, and aggregates of aragonite needles (fuzzy dumbbells) precipitate at the air-water interface and settle to pond floors. The proximal-slope facies (28--54 C), which forms the margins of terracette pools, is composed of arcuate aragonite needle shrubs that create small microterracettes on the steep slope face. Finally, the distal-slope facies (28--30 C) is composed of calcite spherules and calcite feather crystals. Despite the presence of abundant microbial mat communities and their observed role in providing substrates for mineralization, the compositions of spring-water and travertine predominantly reflect abiotic physical and chemical processes. Vigorous CO{sub 2} degassing causes a +2 unit increase in spring water pH, as well as Rayleigh-type covariations between the concentration of dissolved inorganic carbon and corresponding {delta}{sup 13}C. Travertine {delta}{sup 13}C and {delta}{sup 18}O are nearly equivalent to aragonite and calcite equilibrium values calculated from spring water in the higher-temperature ({approximately}50--73 C) depositional facies. Conversely, travertine precipitating in the lower-temperature (<{approximately}50 C) depositional facies exhibits {delta}{sup 13}C and {delta}{sup 18}O values that are as

  4. Cross-boundary management between national parks and surrounding lands: A review and discussion

    Science.gov (United States)

    Schonewald-Cox, Christine; Buechner, Marybeth; Sauvajot, Raymond; Wilcox, Bruce A.

    1992-03-01

    Protecting biodiversity on public lands is difficult, requiring the management of a complex array of factors. This is especially true when the ecosystems in question are affected by, or extend onto, lands outside the boundaries of the protected area. In this article we review recent developments in the cross-boundary management of protected natural resources, such as parks, wildlife reserves, and designated wilderness areas. Five ecological and 11 anthropic techniques have been suggested for use in cross-boundary management. The categories are not mutually exclusive, but each is a distinct and representative approach, suggested by various authors from academic, managerial, and legal professions. The ecological strategies stress the collection of basic data and documentation of trends. The anthropic techniques stress the usefulness of cooperative guidelines and the need to develop a local constituency which supports park goals. However, the situation is complex and the needed strategies are often difficult to implement. Diverse park resources are influenced by events in surrounding lands. The complexity and variability of sources, the ecological systems under protection, and the uncertainty of the effects combine to produce situations for which there are no simple answers. The solution to coexistence of the park and surrounding land depends upon creative techniques and recommendations, many still forthcoming. Ecological, sociological, legal, and economic disciplines as well as the managing agency should all contribute to these recommendations. Platforms for change include legislation, institutional policies, communication, education, management techniques, and ethics.

  5. Pregnancy rates in central Yellowstone bison

    Science.gov (United States)

    Gogan, Peter J.; Russell, Robin E.; Olexa, Edward M.; Podruzny, Kevin M.

    2013-01-01

    Plains bison (Bison b. bison) centered on Yellowstone National Park are chronically infected with brucellosis (Brucella abortus) and culled along the park boundaries to reduce the probability of disease transmission to domestic livestock. We evaluated the relationship between pregnancy rates and age, dressed carcass weight, and serological status for brucellosis among bison culled from the central Yellowstone subpopulation during the winters of 1996–1997, 2001–2002, and 2002–2003. A model with only dressed carcass weight was the best predictor of pregnancy status for all ages with the odds of pregnancy increasing by 1.03 (95% CI = 1.02–1.04) for every 1-kg increase in weight. We found no effect of age or the serological status for brucellosis on pregnancy rates across age classes; however, we did find a positive association between age and pregnancy rates for bison ≥2 years old. Bison ≥2 years old had an overall pregnancy rate of 65% with markedly different rates in alternate ages for animals between 3 and 7 years old. Pregnancy rates were 0.50 (95% CI = 0.31–0.69) for brucellosis positive and 0.57 (95% CI = 0.34–0.78) for brucellosis negative 2- and 3-year-olds and 0.74 (95% CI = 0.60–0.85) in brucellosis positive and 0.69 (95% CI = 0.49–0.85) in brucellosis negative bison ≥4 years old. Only 1 of 21 bison <2 years old was pregnant. Our findings are important to accurately predict the effects of brucellosis on Yellowstone bison population dynamics. We review our results relative to other studies of Yellowstone bison that concluded serological status for brucellosis influences pregnancy rates.

  6. An aerial radiological survey of the Fermi National Accelerator Laboratory and surrounding area, Batavia, Illinois

    International Nuclear Information System (INIS)

    Fritzsche, A.E.

    1990-11-01

    An aerial radiological gamma survey was conducted over the Fermi National Accelerator Laboratory during 1 through 6 June 1989. Flight lines at 150-foot altitude and 250-foot line spacings assured nearly 100% coverage. The terrestrial exposure at about 6 μR/h was nearly the same as that measured by the previous survey of this area (May 1977). Ten anomalous areas, mostly Na-22 and Mn-54, were detected within buildings and laboratories in the area. Although these locations have changed somewhat from the 1977 survey, the aerial data shows good agreement with the ground-based ion chamber and soil sample data. 7 refs., 15 figs., 1 tab

  7. References to Africa in Adam Smith's Wealth of Nations and Some Key Propositions Surrounding Them

    OpenAIRE

    Amavilah, Voxi Heinrich

    2010-01-01

    Adam Smith sought to illustrate some of his propositions in The Wealth of Nations (WON) with examples from Africa. However, the examples were few, and many were neither profound nor instructive from a principles viewpoint. I find that Africa figures very little in the WON, and nearly every time it appears cursory. With 20/20 hindsight, one may conclude that opinions about Africa have remained invariant with respect to time. Final value-judgment about whether that is a good or bad thing rests...

  8. Anatomy of Old Faithful from subsurface seismic imaging of the Yellowstone Upper Geyser Basin

    KAUST Repository

    Wu, Sin-Mei; Ward, Kevin M.; Farrell, Jamie; Lin, Fan-Chi; Karplus, Marianne; Smith, Robert B.

    2017-01-01

    The Upper Geyser Basin in Yellowstone National Park contains one of the highest concentrations of hydrothermal features on Earth including the iconic Old Faithful geyser. Although this system has been the focus of many geological, geochemical

  9. Study of Plant Species Richness in Habitats with Different Grazing Intensities at Golestan National Park and Surrounding Area

    Directory of Open Access Journals (Sweden)

    A. Bagheri

    2016-12-01

    Full Text Available Considering the importance of plant diversity and to evaluate the effect of grazing pressure on species richness and structure of plant communities, this experiment was conducted at Golestan National Park and its surrounding areas in the north east of Iran. Sampling was conducted in intact and abandoned habitats and habitats under seasonal and heavy grazing, using Modified Whitaker Plot in 1, 10,100 and 1000 m2 spatial scales. Results showed that the composition of plant species from different habitats was different. In addition the increasing intensity of grazing increased the importance of therophytes and decreased the role of hemicryptophytes and phanerophytes and also decreasd the amount of species richness. Mean species richness of studied habitat showed a significant difference in all four sampling spatial scales. The results showed that plant species richness decreased in the areas affected by heavy grazing and conservation against grazing plays an important role in maintaining species richness.

  10. Geomicrobiology of Hydrothermal Vents in Yellowstone Lake: Phylogenetic and Functional Analysis suggest Importance of Geochemistry (Invited)

    Science.gov (United States)

    Inskeep, W. P.; Macur, R.; Jay, Z.; Clingenpeel, S.; Tenney, A.; Lavalvo, D.; Shanks, W. C.; McDermott, T.; Kan, J.; Gorby, Y.; Morgan, L. A.; Yooseph, S.; Varley, J.; Nealson, K.

    2010-12-01

    Yellowstone Lake (Yellowstone National Park, WY, USA) is a large, high-altitude, fresh-water lake that straddles the most recent Yellowstone caldera, and is situated on top of significant hydrothermal activity. An interdisciplinary study is underway to evaluate the geochemical and geomicrobiological characteristics of several hydrothermal vent environments sampled using a remotely operated vehicle, and to determine the degree to which these vents may influence the biology of this young freshwater ecosystem. Approximately six different vent systems (locations) were sampled during 2007 and 2008, and included water obtained directly from the hydrothermal vents as well as biomass and sediment associated with these high-temperature environments. Thorough geochemical analysis of these hydrothermal environments reveals variation in pH, sulfide, hydrogen and other potential electron donors that may drive primary productivity. The concentrations of dissolved hydrogen and sulfide were extremely high in numerous vents sampled, especially the deeper (30-50 m) vents located in the Inflated Plain, West Thumb, and Mary Bay. Significant dilution of hydrothermal fluids occurs due to mixing with surrounding lake water. Despite this, the temperatures observed in many of these hydrothermal vents range from 50-90 C, and elevated concentrations of constituents typically associated with geothermal activity in Yellowstone are observed in waters sampled directly from vent discharge. Microorganisms associated with elemental sulfur mats and filamentous ‘streamer’ communities of Inflated Plain and West Thumb (pH range 5-6) were dominated by members of the deeply-rooted bacterial Order Aquificales, but also contain thermophilic members of the domain Archaea. Assembly of metagenome sequence from the Inflated Plain vent biomass and to a lesser extent, West Thumb vent biomass reveal the importance of Sulfurihydrogenibium-like organisms, also important in numerous terrestrial geothermal

  11. SOILS AND GEOENVIRONMENTS OF THE NATIONAL PARK OF VIRUÁ AND SURROUNDING, RORAIMA: INTEGRATED VISION OF THE LANDSCAPE AND ENVIRONMENTAL SERVICE

    Directory of Open Access Journals (Sweden)

    Bruno Araujo Furtado de Mendonça

    2013-06-01

    Full Text Available http://dx.doi.org/10.5902/198050989287The Viruá National Park (PARNA Viruá occupies 227.011ha, in the region of the low ‘Branco’ river, in Roraima state. This area includes an extensive mosaic of complex seasonally flooded forested and non-forested environments. The present work had as general objective to characterize the pedology aspects and the geo-environmental units of the Park and surroundings, in an integrated vision of the landscape and, additionally, estimate the carbon stocks in the soils and geo-environments. We described and collected 29 soil profiles in the main vegetation types of Campinaranas and Forests of PARNA Viruá and surroundings. The main soil classes are: Espodossolo Humilúvico, Neossolo Quartzarênico, Neossolo Flúvico, Neossolo Litólicos, Latossolo Vermelho-Amarelo, Latossolo Vermelho, Cambissolo Háplico, Cambissolo Flúvico, Gleissolo Háplico and Plintossolo Háplico. The soils present spatial distribution marked by abrupt limits and close association with the vegetation type. We identified three pedo-environments: (1 sandy soils of the Campinaranas; (2 soils associated with the inselbergs and adjacencies; and (3 alluvial soils. We mapped and described 18 geoenvironmental units in PARNA Viruá National Park. The main geo-environments are: i Sandy plains and Paleodunes with grassy and arborous Campinarana on ‘Neossolos Quartzarênicos hidromórficos’ and ‘Espodossolos’; and Floodplains and; ii Terraces with Igapó Forest on sandy hydromorphic soils, occupying 24.6% and 20.1% of the studied area, respectively. In terms of total soil carbon stocks, the geo-environments of the sandy complexes of Campinaranas and associations stand out, with 9450.9 Gg C. The great extension and representativeness of the sandy areas of Campinaranas characterize PARNA Viruá PArk as an important conservation unit for protection Amazonian sandy soil systems. The areas under the domain of ‘Espodossolos’ possess the

  12. Aerial radiological survey of the Brookhaven National Laboratory and surrounding area, Upton, New York. Date of survey: June 1983

    International Nuclear Information System (INIS)

    Hobaugh, J.L.

    1985-02-01

    An aerial radiological survey was performed from 11 to 13 June 1983, over approximately a 64-square-kilometer (25-square-mile) area surrounding the Brookhaven National Laboratory (BNL). BNL is located in the center of Suffolk County, Long Island, New York. All gamma radiation data were collected by flying east-west lines spaced 76 meters (250 feet) apart at an altitude of 46 meters (150 feet) above ground level. Count rates obtained from the aerial platform were converted to exposure rates at 1 meter above the ground. The average background exposure rate in the survey area ranged from 5 to 10 microroentgens per hour (μR/h). The reported exposure rate values include an estimated cosmic ray contribution of 4.0 μR/h. Ground-based measurements made during the same time period were compared to the aerial survey results. Pressurized ion chamber readings and soil samples were taken from two locations within the aerial survey boundaries. Exposure rate values obtained from these measurement techniques were in agreement with those obtained from the aerial data. A total of 23 areas of man-made radioactivity were identified. The dominant isotopes found over these areas were cesium-137, sodium-22, manganese-54, and cobalt-60. A similar survey was conducted in May 1980. The 1983 survey results were similar to the 1980 results. Three areas of low level man-made activity were not reproduced by the 1983 data. Ten new areas were detected. The major difference occurred because of the increased sensitivity and spatial reduction brought on by lowering the altitude and decreasing the line spacing. 8 refs., 28 figs., 4 tabs

  13. Smart Surroundings

    NARCIS (Netherlands)

    Havinga, Paul J.M.; Jansen, P.G.; Lijding, M.E.M.; Scholten, Johan

    2004-01-01

    Ambient systems are networked embedded systems integrated with everyday environments and supporting people in their activities. These systems will create a Smart Surrounding for people to facilitate and enrich daily life and increase productivity at work. Such systems will be quite different from

  14. Human dynamics and forest management: a baseline assessment of the socioeconomic characteristics of the region surrounding the El Yunque National Forest

    Science.gov (United States)

    Kathleen McGinley

    2016-01-01

    In this paper, I examine the socioeconomic dynamics and human–environment interactions in the region surrounding the El Yunque National Forest (EYNF) in northeastern Puerto Rico and their implications for policy development and sustainable resource use. As part of a larger, comprehensive assessment of the conditions and trends of the EYNF and broader region, I...

  15. Evaluating estimators for numbers of females with cubs-of-the-year in the Yellowstone grizzly bear population

    Science.gov (United States)

    Cherry, S.; White, G.C.; Keating, K.A.; Haroldson, Mark A.; Schwartz, Charles C.

    2007-01-01

    Current management of the grizzly bear (Ursus arctos) population in Yellowstone National Park and surrounding areas requires annual estimation of the number of adult female bears with cubs-of-the-year. We examined the performance of nine estimators of population size via simulation. Data were simulated using two methods for different combinations of population size, sample size, and coefficient of variation of individual sighting probabilities. We show that the coefficient of variation does not, by itself, adequately describe the effects of capture heterogeneity, because two different distributions of capture probabilities can have the same coefficient of variation. All estimators produced biased estimates of population size with bias decreasing as effort increased. Based on the simulation results we recommend the Chao estimator for model M h be used to estimate the number of female bears with cubs of the year; however, the estimator of Chao and Shen may also be useful depending on the goals of the research.

  16. 76 FR 61266 - Special Regulations; Areas of the National Park System, Grand Teton National Park, Bicycle Routes...

    Science.gov (United States)

    2011-10-04

    ... Yellowstone National Park, Grand Teton is at the heart of the Greater Yellowstone Ecosystem, and includes the... elk, moose, bison, pronghorn, grizzly and black bears, grey wolves, and coyotes. Other species such as...

  17. Monitoring gas and heat emissions at Norris Geyser Basin, Yellowstone National Park, USA based on a combined eddy covariance and Multi-GAS approach

    Science.gov (United States)

    Lewicki, J. L.; Kelly, P. J.; Bergfeld, D.; Vaughan, R. G.; Lowenstern, J. B.

    2017-11-01

    We quantified gas and heat emissions in an acid-sulfate, vapor-dominated area (0.04-km2) of Norris Geyser Basin, located just north of the 0.63 Ma Yellowstone Caldera and near an area of anomalous uplift. From 14 May to 3 October 2016, an eddy covariance system measured half-hourly CO2, H2O and sensible (H) and latent (LE) heat fluxes and a Multi-GAS instrument measured (1 Hz frequency) atmospheric H2O, CO2 and H2S volumetric mixing ratios. We also measured soil CO2 fluxes using the accumulation chamber method and temperature profiles on a grid and collected fumarole gas samples for geochemical analysis. Eddy covariance CO2 fluxes ranged from - 56 to 885 g m- 2 d- 1. Using wavelet analysis, average daily eddy covariance CO2 fluxes were locally correlated with average daily environmental parameters on several-day to monthly time scales. Estimates of CO2 emission rate from the study area ranged from 8.6 t d- 1 based on eddy covariance measurements to 9.8 t d- 1 based on accumulation chamber measurements. Eddy covariance water vapor fluxes ranged from 1178 to 24,600 g m- 2 d- 1. Nighttime H and LE were considered representative of hydrothermal heat fluxes and ranged from 4 to 183 and 38 to 504 W m- 2, respectively. The total hydrothermal heat emission rate (H + LE + radiant) estimated for the study area was 11.6 MW and LE contributed 69% of the output. The mean ± standard deviation of H2O, CO2 and H2S mixing ratios measured by the Multi-GAS system were 9.3 ± 3.1 parts per thousand, 467 ± 61 ppmv, and 0.5 ± 0.6 ppmv, respectively, and variations in the gas compositions were strongly correlated with diurnal variations in environmental parameters (wind speed and direction, atmospheric temperature). After removing ambient H2O and CO2, the observed variations in the Multi-GAS data could be explained by the mixing of relatively H2O-CO2-H2S-rich fumarole gases with CO2-rich and H2O-H2S-poor soil gases. The fumarole H2O/CO2 and CO2/H2S end member ratios (101.7 and 27

  18. Monitoring gas and heat emissions at Norris Geyser Basin, Yellowstone National Park, USA based on a combined eddy covariance and Multi-GAS approach

    Science.gov (United States)

    Lewicki, Jennifer L.; Kelly, Peter; Bergfeld, Deborah; Vaughan, R. Greg; Lowenstern, Jacob B.

    2017-01-01

    We quantified gas and heat emissions in an acid-sulfate, vapor-dominated area (0.04-km2) of Norris Geyser Basin, located just north of the 0.63 Ma Yellowstone Caldera and near an area of anomalous uplift. From 14 May to 3 October 2016, an eddy covariance system measured half-hourly CO2, H2O and sensible (H) and latent (LE) heat fluxes and a Multi-GAS instrument measured (1 Hz frequency) atmospheric H2O, CO2 and H2S volumetric mixing ratios. We also measured soil CO2 fluxes using the accumulation chamber method and temperature profiles on a grid and collected fumarole gas samples for geochemical analysis. Eddy covariance CO2 fluxes ranged from − 56 to 885 g m− 2 d− 1. Using wavelet analysis, average daily eddy covariance CO2 fluxes were locally correlated with average daily environmental parameters on several-day to monthly time scales. Estimates of CO2emission rate from the study area ranged from 8.6 t d− 1 based on eddy covariance measurements to 9.8 t d− 1 based on accumulation chamber measurements. Eddy covariance water vapor fluxes ranged from 1178 to 24,600 g m− 2 d− 1. Nighttime H and LEwere considered representative of hydrothermal heat fluxes and ranged from 4 to 183 and 38 to 504 W m− 2, respectively. The total hydrothermal heat emission rate (H + LE + radiant) estimated for the study area was 11.6 MW and LE contributed 69% of the output. The mean ± standard deviation of H2O, CO2 and H2S mixing ratios measured by the Multi-GAS system were 9.3 ± 3.1 parts per thousand, 467 ± 61 ppmv, and 0.5 ± 0.6 ppmv, respectively, and variations in the gas compositions were strongly correlated with diurnal variations in environmental parameters (wind speed and direction, atmospheric temperature). After removing ambient H2O and CO2, the observed variations in the Multi-GAS data could be explained by the mixing of relatively H2O-CO2-H2S-rich fumarole gases with CO2-rich and H2O-H2S-poor soil gases. The

  19. Willow on Yellowstone's northern range: evidence for a trophic cascade?

    Science.gov (United States)

    Beyer, Hawthorne L; Merrill, Evelyn H; Varley, Nathan; Boyce, Mark S

    2007-09-01

    Reintroduction of wolves (Canis lupus) to Yellowstone National Park in 1995-1996 has been argued to promote a trophic cascade by altering elk (Cervus elaphus) density, habitat-selection patterns, and behavior that, in turn, could lead to changes within the plant communities used by elk. We sampled two species of willow (Salix boothii and S. geyeriana) on the northern winter range to determine whether (1) there was quantitative evidence of increased willow growth following wolf reintroduction, (2) browsing by elk affected willow growth, and (3) any increase in growth observed was greater than that expected by climatic and hydrological factors alone, thereby indicating a trophic cascade caused by wolves. Using stem sectioning techniques to quantify historical growth patterns we found an approximately twofold increase in stem growth-ring area following wolf reintroduction for both species of willow. This increase could not be explained by climate and hydrological factors alone; the presence of wolves on the landscape was a significant predictor of stem growth above and beyond these abiotic factors. Growth-ring area was positively correlated with the previous year's ring area and negatively correlated with the percentage of twigs browsed from the stem during the winter preceding growth, indicating that elk browse impeded stem growth. Our results are consistent with the hypothesis of a behaviorally mediated trophic cascade on Yellowstone's northern winter range following wolf reintroduction. We suggest that the community-altering effects of wolf restoration are an endorsement of ecological-process management in Yellowstone National Park.

  20. Pregnancy rates in central Yellowstone bison

    Science.gov (United States)

    Gogan, Peter J.; Russell, Robin E.; Olexa, Edward M.; Podruzny, Kevin M.

    2013-01-01

    Plains bison (Bison b. bison) centered on Yellowstone National Park are chronically infected with brucellosis (Brucella abortus) and culled along the park boundaries to reduce the probability of disease transmission to domestic livestock. We evaluated the relationship between pregnancy rates and age, dressed carcass weight, and serological status for brucellosis among bison culled from the central Yellowstone subpopulation during the winters of 1996–1997, 2001–2002, and 2002–2003. A model with only dressed carcass weight was the best predictor of pregnancy status for all ages with the odds of pregnancy increasing by 1.03 (95% CI = 1.02–1.04) for every 1-kg increase in weight. We found no effect of age or the serological status for brucellosis on pregnancy rates across age classes; however, we did find a positive association between age and pregnancy rates for bison ≥2 years old. Bison ≥2 years old had an overall pregnancy rate of 65% with markedly different rates in alternate ages for animals between 3 and 7 years old. Pregnancy rates were 0.50 (95% CI = 0.31–0.69) for brucellosis positive and 0.57 (95% CI = 0.34–0.78) for brucellosis negative 2- and 3-year-olds and 0.74 (95% CI = 0.60–0.85) in brucellosis positive and 0.69 (95% CI = 0.49–0.85) in brucellosis negative bison ≥4 years old. Only 1 of 21 bison pregnancy rates.

  1. Cyanobacterial Community Structure In Lithifying Mats of A Yellowstone Hotspring-Implications for Precambrian Stromatolite Biocomplexity

    Science.gov (United States)

    Lau, Evan; Nash, C. Z.; Vogler, D. R.; Cullings, K.; DeVincenzi, Donald (Technical Monitor)

    2002-01-01

    Denaturing Gradient Gel Electrophoresis (DGGE) of partial 16S rRNA gene sequences was used to investigate the molecular biodiversity of cyanobacterial communities inhabiting various lithified morpho-structures in two hotsprings of Yellowstone National Park. These morpho-structures - flat-topped columns, columnar cones, and ridged cones - resemble ancient stromatolites, which are possibly biogenic in origin. The top, middle and bottom sections of these lithified morpho-structures, as well as surrounding non-lithified mats were analyzed to determine the vertical and spatial distribution of cyanobacterial communities. Results from DGGE indicate that the cyanobacterial community composition of lithified morpho-structures (flat-topped columns, columnar cones, and ridged cones) were largely similar in vertical distribution as well as among the morpho-structures being studied. Preliminary results indicate that the cyanobacterial communities in these lithified morpho-structures were significantly different from communities in surrounding non-lithified mats. These results provide additional support to the theory that certain Phormidium/Leptolyngbya species are involved in the morphogenesis of lithifying morpho-structures in hotsprings and may have played a role in the formation of ancient stromatolites.

  2. Issues Surrounding English, the Internationalisation of Higher Education and National Cultural Identity in Asia: A Focus on Japan

    Science.gov (United States)

    Le Phan, Ha

    2013-01-01

    The English language is significant to the internationalisation of higher education worldwide. Countries in Asia are proactive in appropriating English for their national interests, while paying attention to associated national cultural identity issues. This article examines the ways in which the role of English is interpreted and justified in…

  3. Prodigious degassing of a billion years of accumulated radiogenic helium at Yellowstone

    Science.gov (United States)

    Lowenstern, Jacob B.; Evans, William C.; Bergfeld, D.; Hunt, Andrew G.

    2014-01-01

    Helium is used as a critical tracer throughout the Earth sciences, where its relatively simple isotopic systematics is used to trace degassing from the mantle, to date groundwater and to time the rise of continents1. The hydrothermal system at Yellowstone National Park is famous for its high helium-3/helium-4 isotope ratio, commonly cited as evidence for a deep mantle source for the Yellowstone hotspot2. However, much of the helium emitted from this region is actually radiogenic helium-4 produced within the crust by α-decay of uranium and thorium. Here we show, by combining gas emission rates with chemistry and isotopic analyses, that crustal helium-4 emission rates from Yellowstone exceed (by orders of magnitude) any conceivable rate of generation within the crust. It seems that helium has accumulated for (at least) many hundreds of millions of years in Archaean (more than 2.5 billion years old) cratonic rocks beneath Yellowstone, only to be liberated over the past two million years by intense crustal metamorphism induced by the Yellowstone hotspot. Our results demonstrate the extremes in variability of crustal helium efflux on geologic timescales and imply crustal-scale open-system behaviour of helium in tectonically and magmatically active regions.

  4. The influence of herd size, conspecific risk, and predation risk on the vigilance of elk (Cervus elaphus) in Yellowstone National Park, and, Interest, learning, and a thematic biology course

    Science.gov (United States)

    Lung, Mark A.

    This dissertation is a composite of biological and educational research. The biological research concerns Rocky Mountain elk (Cervus elaphus ) behavior. The educational research presents ideas and findings on the influence of a thematic general biology course on student interest and perception of learning. The dissertation begins with a Preface that attempts to bring the ideas presented in later chapters together. Chapter One is a review of the literature concerning sociality, social behaviors, and elk biology. It summarizes current research literature as a means of introduction to Chapter Two. Chapter Two presents findings concerning the effects of herd size, predation risk, and the risk of being near conspecifics on two behaviors commonly associated with social animals---vigilance and aggression. Vigilance and aggression were measured in elk in Yellowstone National Park in two regions that varied in their presence of elk predators (wolves---Canis lupus, and grizzly bears---Ursus arctos) and in two seasons (spring and fall) that varied in the risks of being near conspecifics. Overall, male and female elk responded very differently. Male elk adjust their vigilance and aggression in response to changes in conspecific risk, but not to changes in predation risk. Female elk adjust their vigilance in response to changes in predation risk, but not to changes in conspecific risk. Males show no response in vigilance to changes in herd size. Non-reproductive females, however, adjust their levels of vigilance with changes in herd size in high risk regions. Interestingly, in the spring, vigilance decreases with increasing herd size, but in the fall, vigilance increases with increasing herd size. Chapter Three presents findings concerning the influence of a thematic course design on student perceptions of interest and teaming in a non-major's biology course (Bins 100: Concepts of Biology). I compared responses on student evaluations from two sections of Bios 100 taught in a

  5. The New Departure Debate Surrounding Congressional Efforts to Create a National System of Education, 1871-1889

    Science.gov (United States)

    Groen, Mark

    2007-01-01

    Congressman George Frisbie Hoar of Massachusetts introduced a bill "to establish a system of national education" on February 25, 1870. This bill, and others that followed, opened an acrimonious political debate that lasted for twenty years. The opening salvos of that debate, and the regional issues of ethnicity and religion that framed…

  6. Aerial radiological survey of the Argonne National Laboratory and surrounding area, Argonne, Illinois. Date of survey: May 1977

    International Nuclear Information System (INIS)

    1982-12-01

    An aerial radiological survey was conducted over the facilities of the Argonne National Laboratory in Argonne, Illinois, on 2 to 13 May 1977. The survey was flown at an altitude of 46 m by a helicopter containing 20 sodium iodide detectors. The line spacing was also 46 m. Enhanced gamma exposure rate levels, which could be attributed to Argonne operations, were observed at many locations

  7. Topogrid Derived 10 Meter Resolution Digital Elevation Model of the Shenandoah National Park and Surrounding Region, Virginia

    Science.gov (United States)

    Chirico, Peter G.; Tanner, Seth D.

    2004-01-01

    Explanation The purpose of developing a new 10m resolution DEM of the Shenandoah National Park Region was to more accurately depict geologic structure, surfical geology, and landforms of the Shenandoah National Park Region in preparation for automated landform classification. Previously, only a 30m resolution DEM was available through the National Elevation Dataset (NED). During production of the Shenandoah10m DEM of the Park the Geography Discipline of the USGS completed a revised 10m DEM to be included into the NED. However, different methodologies were used to produce the two similar DEMs. The ANUDEM algorithm was used to develop the Shenadoah DEM data. This algorithm allows for the inclusion of contours, streams, rivers, lake and water body polygons as well as spot height data to control the elevation model. A statistical analysis using over 800 National Geodetic Survey (NGS) first and second order vertical control points reveals that the Shenandoah10m DEM, produced as a part of the Appalachian Blue Ridge Landscape project, has a vertical accuracy of ?4.87 meters. The metadata for the 10m NED data reports a vertical accuracy of ?7m. A table listing the NGS control points, the elevation comparison, and the RMSE for the Shenandoah10m DEM is provided. The process of automated terrain classification involves developing statistical signatures from the DEM for each type of surficial deposit and landform type. The signature will be a measure of several characteristics derived from the elevation data including slope, aspect, planform curvature, and profile curvature. The quality of the DEM is of critical importance when extracting terrain signatures. The highest possible horizontal and vertical accuracy is required. The more accurate Shenandoah 10m DEM can now be analyzed and integrated with the geologic observations to yield statistical correlations between the two in the development of landform and surface geology mapping projects.

  8. Confusion surrounding the concept of nuclear 'security'. 'Preventing Japan from going nuclear contributes to Japan's national security'?

    International Nuclear Information System (INIS)

    Kubota, Masafumi

    2012-01-01

    A law enacted on June 20 to establish a new Nuclear Regulatory Authority (NRA) fully separated from the nuclear promotional authorities. It added the provision, which says nuclear safety should be guaranteed not only to defend lives, people's health and the environment but also to 'contribute to Japan's national security', to Article 2 of the Atomic Energy Basic Law. NRA integrated the existing regulatory authorities for safety, security and safeguards, into one. Supporters of an amendment quietly slipped into the law were denying it could provide cover for military use of nuclear technology, but arouse international concern about recycling program of extracting plutonium from spent fuels. Nuclear policy minister said: 'The safeguards are in place to prevent nuclear proliferation. The world 'security' precisely means the prevention of nuclear proliferation.' If not used explicitly about safeguards, they left room for stretched interpretation. The author recommended the world' contribute to Japan's national security' should be deleted instead of explaining appropriately, both at home and abroad, the use of nuclear power in Japan limited to peaceful purposes. (T. Tanaka)

  9. Alternative Fuels Data Center: Yellowstone Park Recycles Vehicle Batteries

    Science.gov (United States)

    for Solar Power Yellowstone Park Recycles Vehicle Batteries for Solar Power to someone by E -mail Share Alternative Fuels Data Center: Yellowstone Park Recycles Vehicle Batteries for Solar Power on Facebook Tweet about Alternative Fuels Data Center: Yellowstone Park Recycles Vehicle Batteries

  10. Glacial and Quaternary geology of the northern Yellowstone area, Montana and Wyoming

    Science.gov (United States)

    Pierce, Kenneth L.; Licciardi, Joseph M.; Krause, Teresa R.; Whitlock, Cathy

    2014-01-01

    This field guide focuses on the glacial geology and paleoecology beginning in the Paradise Valley and progressing southward into northern Yellowstone National Park. During the last (Pinedale) glaciation, the northern Yellowstone outlet glacier flowed out of Yellowstone Park and down the Yellowstone River Valley into the Paradise Valley. The field trip will traverse the following Pinedale glacial sequence: (1) deposition of the Eightmile terminal moraines and outwash 16.5 ± 1.4 10Be ka in the Paradise Valley; (2) glacial recession of ~8 km and deposition of the Chico moraines and outwash 16.1 ± 1.7 10Be ka; (3) glacial recession of 45 km to near the northern Yellowstone boundary and moraine deposition during the Deckard Flats readjustment 14.2 ± 1.2 10Be ka; and (4) glacial recession of ~37 km and deposition of the Junction Butte moraines 15.2 ± 1.3 10Be ka (this age is a little too old based on the stratigraphic sequence). Yellowstone's northern range of sagebrush-grasslands and bison, elk, wolf, and bear inhabitants is founded on glacial moraines, sub-glacial till, and outwash deposited during the last glaciation. Floods released from glacially dammed lakes and a landslide-dammed lake punctuate this record. The glacial geologic reconstruction was evaluated by calculation of basal shear stress, and yielded the following values for flow pattern in plan view: strongly converging—1.21 ± 0.12 bars (n = 15); nearly uniform—1.04 ± 0.16 bars (n = 11); and strongly diverging—0.84 ± 0.14 bars (n = 16). Reconstructed mass balance yielded accumulation and ablation each of ~3 km3/yr, with glacial movement near the equilibrium line altitude dominated by basal sliding. Pollen and charcoal records from three lakes in northern Yellowstone provide information on the postglacial vegetation and fire history. Following glacial retreat, sparsely vegetated landscapes were colonized first by spruce parkland and then by closed subalpine forests. Regional fire activity

  11. Exploring Yellowstone National Park with Mathematical Modeling

    Science.gov (United States)

    Wickstrom, Megan H.; Carr, Ruth; Lackey, Dacia

    2017-01-01

    Mathematical modeling, a practice standard in the Common Core State Standards for Mathematics (CCSSM) (CCSSI 2010), is a process by which students develop and use mathematics as a tool to make sense of the world around them. Students investigate a real-world situation by asking mathematical questions; along the way, they need to decide how to use…

  12. Gray wolves as climate change buffers in Yellowstone.

    Science.gov (United States)

    Wilmers, Christopher C; Getz, Wayne M

    2005-04-01

    Understanding the mechanisms by which climate and predation patterns by top predators co-vary to affect community structure accrues added importance as humans exert growing influence over both climate and regional predator assemblages. In Yellowstone National Park, winter conditions and reintroduced gray wolves (Canis lupus) together determine the availability of winter carrion on which numerous scavenger species depend for survival and reproduction. As climate changes in Yellowstone, therefore, scavenger species may experience a dramatic reshuffling of food resources. As such, we analyzed 55 y of weather data from Yellowstone in order to determine trends in winter conditions. We found that winters are getting shorter, as measured by the number of days with snow on the ground, due to decreased snowfall and increased number of days with temperatures above freezing. To investigate synergistic effects of human and climatic alterations of species interactions, we used an empirically derived model to show that in the absence of wolves, early snow thaw leads to a substantial reduction in late-winter carrion, causing potential food bottlenecks for scavengers. In addition, by narrowing the window of time over which carrion is available and thereby creating a resource pulse, climate change likely favors scavengers that can quickly track food sources over great distances. Wolves, however, largely mitigate late-winter reduction in carrion due to earlier snow thaws. By buffering the effects of climate change on carrion availability, wolves allow scavengers to adapt to a changing environment over a longer time scale more commensurate with natural processes. This study illustrates the importance of restoring and maintaining intact food chains in the face of large-scale environmental perturbations such as climate change.

  13. Gray Wolves as Climate Change Buffers in Yellowstone

    Directory of Open Access Journals (Sweden)

    Wilmers Christopher C

    2005-01-01

    Full Text Available Understanding the mechanisms by which climate and predation patterns by top predators co-vary to affect community structure accrues added importance as humans exert growing influence over both climate and regional predator assemblages. In Yellowstone National Park, winter conditions and reintroduced gray wolves (Canis lupus together determine the availability of winter carrion on which numerous scavenger species depend for survival and reproduction. As climate changes in Yellowstone, therefore, scavenger species may experience a dramatic reshuffling of food resources. As such, we analyzed 55 y of weather data from Yellowstone in order to determine trends in winter conditions. We found that winters are getting shorter, as measured by the number of days with snow on the ground, due to decreased snowfall and increased number of days with temperatures above freezing. To investigate synergistic effects of human and climatic alterations of species interactions, we used an empirically derived model to show that in the absence of wolves, early snow thaw leads to a substantial reduction in late-winter carrion, causing potential food bottlenecks for scavengers. In addition, by narrowing the window of time over which carrion is available and thereby creating a resource pulse, climate change likely favors scavengers that can quickly track food sources over great distances. Wolves, however, largely mitigate late-winter reduction in carrion due to earlier snow thaws. By buffering the effects of climate change on carrion availability, wolves allow scavengers to adapt to a changing environment over a longer time scale more commensurate with natural processes. This study illustrates the importance of restoring and maintaining intact food chains in the face of large-scale environmental perturbations such as climate change.

  14. Gray wolves as climate change buffers in Yellowstone.

    Directory of Open Access Journals (Sweden)

    Christopher C Wilmers

    2005-04-01

    Full Text Available Understanding the mechanisms by which climate and predation patterns by top predators co-vary to affect community structure accrues added importance as humans exert growing influence over both climate and regional predator assemblages. In Yellowstone National Park, winter conditions and reintroduced gray wolves (Canis lupus together determine the availability of winter carrion on which numerous scavenger species depend for survival and reproduction. As climate changes in Yellowstone, therefore, scavenger species may experience a dramatic reshuffling of food resources. As such, we analyzed 55 y of weather data from Yellowstone in order to determine trends in winter conditions. We found that winters are getting shorter, as measured by the number of days with snow on the ground, due to decreased snowfall and increased number of days with temperatures above freezing. To investigate synergistic effects of human and climatic alterations of species interactions, we used an empirically derived model to show that in the absence of wolves, early snow thaw leads to a substantial reduction in late-winter carrion, causing potential food bottlenecks for scavengers. In addition, by narrowing the window of time over which carrion is available and thereby creating a resource pulse, climate change likely favors scavengers that can quickly track food sources over great distances. Wolves, however, largely mitigate late-winter reduction in carrion due to earlier snow thaws. By buffering the effects of climate change on carrion availability, wolves allow scavengers to adapt to a changing environment over a longer time scale more commensurate with natural processes. This study illustrates the importance of restoring and maintaining intact food chains in the face of large-scale environmental perturbations such as climate change.

  15. Distribution of grizzly bears in the Greater Yellowstone Ecosystem, 2004

    Science.gov (United States)

    Schwartz, C.C.; Haroldson, M.A.; Gunther, K.; Moody, D.

    2006-01-01

    The US Fish and Wildlife Service (USFWS) proposed delisting the Yellowstone grizzly bear (Ursus arctos horribilis) in November 2005. Part of that process required knowledge of the most current distribution of the species. Here, we update an earlier estimate of occupied range (1990–2000) with data through 2004. We used kernel estimators to develop distribution maps of occupied habitats based on initial sightings of unduplicated females (n = 481) with cubs of the year, locations of radiomarked bears (n = 170), and spatially unique locations of conflicts, confrontations, and mortalities (n = 1,075). Although each data set was constrained by potential sampling bias, together they provided insight into areas in the Greater Yellowstone Ecosystem (GYE) currently occupied by grizzly bears. The current distribution of 37,258 km2 (1990–2004) extends beyond the distribution map generated with data from 1990–2000 (34,416 km2 ). Range expansion is particularly evident in parts of the Caribou–Targhee National Forest in Idaho and north of Spanish Peaks on the Gallatin National Forest in Montana.

  16. The genealogy and genetic viability of reintroduced Yellowstone grey wolves.

    Science.gov (United States)

    Vonholdt, Bridgett M; Stahler, Daniel R; Smith, Douglas W; Earl, Dent A; Pollinger, John P; Wayne, Robert K

    2008-01-01

    The recovery of the grey wolf in Yellowstone National Park is an outstanding example of a successful reintroduction. A general question concerning reintroduction is the degree to which genetic variation has been preserved and the specific behavioural mechanisms that enhance the preservation of genetic diversity and reduce inbreeding. We have analysed 200 Yellowstone wolves, including all 31 founders, for variation in 26 microsatellite loci over the 10-year reintroduction period (1995-2004). The population maintained high levels of variation (1995 H(0) = 0.69; 2004 H(0) = 0.73) with low levels of inbreeding (1995 F(IS) = -0.063; 2004 F(IS) = -0.051) and throughout, the population expanded rapidly (N(1995) = 21; N(2004) = 169). Pedigree-based effective population size ratios did not vary appreciably over the duration of population expansion (1995 N(e)/N(g) = 0.29; 2000 N(e)/N(g) = 0.26; 2004 N(e)/N(g) = 0.33). We estimated kinship and found only two of 30 natural breeding pairs showed evidence of being related (average r = -0.026, SE = 0.03). We reconstructed the genealogy of 200 wolves based on genetic and field data and discovered that they avoid inbreeding through a wide variety of behavioural mechanisms including absolute avoidance of breeding with related pack members, male-biased dispersal to packs where they breed with nonrelatives, and female-biased subordinate breeding. We documented a greater diversity of such population assembly patterns in Yellowstone than previously observed in any other natural wolf population. Inbreeding avoidance is nearly absolute despite the high probability of within-pack inbreeding opportunities and extensive interpack kinship ties between adjacent packs. Simulations showed that the Yellowstone population has levels of genetic variation similar to that of a population managed for high variation and low inbreeding, and greater than that expected for random breeding within packs or across the entire breeding pool. Although short

  17. Siliceous Shrubs in Yellowstone's Hot Springs: Implications for Exobiological Investigations

    Science.gov (United States)

    Guidry, S. A.; Chafetz, H. S.

    2003-01-01

    Potential relict hot springs have been identified on Mars and, using the Earth as an analog, Martian hot springs are postulated to be an optimal locality for recognizing preserved evidence of extraterrestrial life. Distinctive organic and inorganic biomarkers are necessary to recognize preserved evidence of life in terrestrial and extraterrestrial hot spring accumulations. Hot springs in Yellowstone National Park, Wyoming, U.S.A., contain a wealth of information about primitive microbial life and associated biosignatures that may be useful for future exobiological investigations. Numerous siliceous hot springs in Yellowstone contain abundant, centimeter-scale, spinose precipitates of opaline silica (opal-A). Although areally extensive in siliceous hot spring discharge channel facies, these spinose forms have largely escaped attention. These precipitates referred to as shrubs, consist of porous aggregates of spinose opaline silica that superficially resemble miniature woody plants, i.e., the term shrubs. Shrubs in carbonate precipitating systems have received considerable attention, and represent naturally occurring biotically induced precipitates. As such, shrubs have great potential as hot spring environmental indicators and, more importantly, proxies for pre-existing microbial life.

  18. Seasonal gravity change at Yellowstone caldera

    Science.gov (United States)

    Poland, M. P.; de Zeeuw-van Dalfsen, E.

    2017-12-01

    The driving forces behind Yellowstone's dynamic deformation, vigorous hydrothermal system, and abundant seismicity are usually ascribed to "magmatic fluids," which could refer to magma, water, volatiles, or some combination. Deformation data alone cannot distinguish the relative importance of these fluids. Gravity measurements, however, provide an indication of mass change over time and, when combined with surface displacements, can constrain the density of subsurface fluids. Unfortunately, several decades of gravity surveys at Yellowstone have yielded ambiguous results. We suspect that the difficulty in interpreting Yellowstone gravity data is due to seasonal variations in environmental conditions—especially surface and ground water. Yellowstone gravity surveys are usually carried out at the same time of year (generally late summer) to minimize the impact of seasonality. Nevertheless, surface and subsurface water levels are not likely to be constant from year to year, given annual differences in precipitation. To assess the overall magnitude of seasonal gravity changes, we conducted gravity surveys of benchmarks in and around Yellowstone caldera in May, July, August, and October 2017. Our goal was to characterize seasonal variations due to snow melt/accumulation, changes in river and lake levels, changes in groundwater levels, and changes in hydrothermal activity. We also hope to identify sites that show little variation in gravity over the course of the 2017 surveys, as these locations may be less prone to seasonal changes and more likely to detect small variations due to magmatic processes. Preliminary examination of data collected in May and July 2017 emphasizes the importance of site location relative to sources of water. For example, a site on the banks of the Yellowstone River showed a gravity increase of several hundred microgals associated with a 50 cm increase in the river level. A high-altitude site far from rivers and lakes, in contrast, showed a

  19. Geomicrobiology of sublacustrine thermal vents in Yellowstone Lake: Geochemical controls on microbial community structure and function

    Directory of Open Access Journals (Sweden)

    William P. Inskeep

    2015-10-01

    Full Text Available Yellowstone Lake (Yellowstone National Park, WY, USA is a large high-altitude (2200 m, fresh-water lake, which straddles an extensive caldera and is the center of significant geothermal activity. The primary goal of this interdisciplinary study was to evaluate the microbial populations inhabiting thermal vent communities in Yellowstone Lake (Yellowstone Lake using 16S rRNA gene and random metagenome sequencing, and to determine how geochemical attributes of vent waters influence the distribution of specific microorganisms and their metabolic potential. Thermal vent waters and associated microbial biomass were sampled during two field seasons (2007 - 2008 using a remotely operated vehicle (ROV. Sublacustrine thermal vent waters (circa 50 - 90 oC contained elevated concentrations of numerous constituents associated with geothermal activity including dissolved hydrogen, sulfide, methane and carbon dioxide. Microorganisms associated with sulfur-rich filamentous ‘streamer’ communities of Inflated Plain and West Thumb (pH range 5 - 6 were dominated by bacteria from the Aquificales, but also contained thermophilic archaea from the Crenarchaeota and Euryarchaeota. Novel groups of methanogens and members of the Korarchaeota were observed in vents from West Thumb and Elliot’s Crater (pH 5 - 6. Conversely, metagenome sequence from Mary Bay vent sediments did not yield large assemblies, and contained diverse thermophilic and nonthermophilic bacterial relatives. Analysis of functional genes associated with the major vent populations indicated a direct linkage to high concentrations of carbon dioxide, reduced sulfur (sulfide and/or elemental S, hydrogen and methane in the deep thermal ecosystems. Our observations show that sublacustrine thermal vents in Yellowstone Lake support novel thermophilic communities, which contain microorganisms with functional attributes not found to date in terrestrial geothermal systems of YNP.

  20. Lake Generated Microseisms at Yellowstone Lake as a Record of Ice Phenology

    Science.gov (United States)

    Mohd Mokhdhari, A. A.; Koper, K. D.; Burlacu, R.

    2017-12-01

    It has recently been shown that wave action in lakes produces microseisms, which generate noise peaks in the period range of 0.8-1.2 s as recorded by nearby seismic stations. Such noise peaks have been observed at seven seismic stations (H17A, LKWY, B208, B944, YTP, YLA, and YLT) located within 2 km of the Yellowstone Lake shoreline. Initial work using 2016 data shows that the variations in the microseism signals at Yellowstone Lake correspond with the freezing and thawing of lake ice: the seismic noise occurs more frequently in the spring, summer, and fall, and less commonly in the winter. If this can be confirmed, then lake-generated microseisms could provide a consistent measure of the freezing and melting dates of high-latitude lakes in remote areas. The seismic data would then be useful in assessing the effects of climate change on the ice phenology of those lakes. In this work, we analyze continuous seismic data recorded by the seven seismic stations around Yellowstone Lake for the years of 1995 to 2016. We generate probability distribution functions of power spectral density for each station to observe the broad elevation of energy near a period of 1 s. The time dependence of this 1-s seismic noise energy is analyzed by extracting the power spectral density at 1 s from every processed hour. The seismic observations are compared to direct measurements of the dates of ice-out and freeze-up as reported by rangers at Yellowstone National Park. We examine how accurate the seismic data are in recording the freezing and melting of Yellowstone Lake, and how the accuracy changes as a function of the number of stations used. We also examine how sensitive the results are to the particular range of periods that are analyzed.

  1. The Endemic Plant Taxa of the Köprülü Kanyon National Park and Its Surroundings (Antalya-Isparta

    Directory of Open Access Journals (Sweden)

    Hasan ÖZÇELİK

    2014-07-01

    Full Text Available This study has been conducted in 2003-2004 in order to identify the endemic plants of the Köprülü Kanyon National Park (Antalya-Isparta and its surroundings. A total of 230 endemic taxa belonging to 29 families were determined in the national park and its surroundings. There are 229 taxa belonging to Angiospermae subdivision and 1 taxon belonging to Gymnospermae subdivision in these collected and identified endemic taxa from the research area. There is no endemic taxon in the Bryophyta and Pteridophyta divisions of the park. 218 of the 229 taxa belonging to the Angiospermae subdivision are in the Magnoliopsida (Dicotyledoneae class and other 11 are in the Liliopsida (Monocotyledoneae class. 18 taxa of the vascular plants are specific to the research area. 44 of endemic taxa are included in the endangered category. The number of priority conservation requiring taxa is 21. Endemic taxon number is almost 25% of total flora of the area. The top five families with the highest number of taxa in the study area are Lamiaceae (38, Caryophyllaceae (37, Asteraceae (26, Scrophulariaceae (20, Fabaceae (16 (Table 2. The 10 largest genera with the highest number of taxa are as follows: Silene (15, Astragalus (9, Sideritis (8, Verbascum (7, Centaurea (7, Stachys (6, Helichrysum (6, Alkanna (6, Veronica (5 and Minuartia (5. The distributions according to the phytogeographical regions of the endemic plants identified from the area is as follows: 59.565% Mediterranean elements (137 taxa, 23.478% Irano-Turanian elements (54 taxa, 0.304% Euro-Siberian elements (7 taxa and 13.913% with unknown phytogeographical region (32 taxa. The distributions of these taxa according to the conservation status is as follows: CR (Critically Endangered: 3, EN (Endangered: 22, VU (Vulnerable: 34, LR (Low Risk: 164, (cd (Conservation Dependent: 29, (lc (Least Concern: 106, (nt (Near Threatened: 29. In this study, menacing factors on the flora and vegetation of the area and

  2. Hydrothermal processes above the Yellowstone magma chamber: Large hydrothermal systems and large hydrothermal explosions

    Science.gov (United States)

    Morgan, L.A.; Shanks, W.C. Pat; Pierce, K.L.

    2009-01-01

    Hydrothermal explosions are violent and dramatic events resulting in the rapid ejection of boiling water, steam, mud, and rock fragments from source craters that range from a few meters up to more than 2 km in diameter; associated breccia can be emplaced as much as 3 to 4 km from the largest craters. Hydrothermal explosions occur where shallow interconnected reservoirs of steam- and liquid-saturated fluids with temperatures at or near the boiling curve underlie thermal fields. Sudden reduction in confi ning pressure causes fluids to fl ash to steam, resulting in signifi cant expansion, rock fragmentation, and debris ejection. In Yellowstone, hydrothermal explosions are a potentially signifi cant hazard for visitors and facilities and can damage or even destroy thermal features. The breccia deposits and associated craters formed from hydrothermal explosions are mapped as mostly Holocene (the Mary Bay deposit is older) units throughout Yellowstone National Park (YNP) and are spatially related to within the 0.64-Ma Yellowstone caldera and along the active Norris-Mammoth tectonic corridor. In Yellowstone, at least 20 large (>100 m in diameter) hydrothermal explosion craters have been identifi ed; the scale of the individual associated events dwarfs similar features in geothermal areas elsewhere in the world. Large hydrothermal explosions in Yellowstone have occurred over the past 16 ka averaging ??1 every 700 yr; similar events are likely in the future. Our studies of large hydrothermal explosion events indicate: (1) none are directly associated with eruptive volcanic or shallow intrusive events; (2) several historical explosions have been triggered by seismic events; (3) lithic clasts and comingled matrix material that form hydrothermal explosion deposits are extensively altered, indicating that explosions occur in areas subjected to intense hydrothermal processes; (4) many lithic clasts contained in explosion breccia deposits preserve evidence of repeated fracturing

  3. Origins of geothermal gases at Yellowstone

    Science.gov (United States)

    Lowenstern, Jacob B.; Bergfeld, Deborah; Evans, William C.; Hunt, Andrew G.

    2015-01-01

    Gas emissions at the Yellowstone Plateau Volcanic Field (YPVF) reflect open-system mixing of gas species originating from diverse rock types, magmas, and crustal fluids, all combined in varying proportions at different thermal areas. Gases are not necessarily in chemical equilibrium with the waters through which they vent, especially in acid sulfate terrain where bubbles stream through stagnant acid water. Gases in adjacent thermal areas often can be differentiated by isotopic and gas ratios, and cannot be tied to one another solely by shallow processes such as boiling-induced fractionation of a parent liquid. Instead, they inherit unique gas ratios (e.g., CH4/He) from the dominant rock reservoirs where they originate, some of which underlie the Quaternary volcanic rocks. Steam/gas ratios (essentially H2O/CO2) of Yellowstone fumaroles correlate with Ar/He and N2/CO2, strongly suggesting that H2O/CO2 is controlled by addition of steam boiled from water rich in atmospheric gases. Moreover, H2O/CO2 varies systematically with geographic location, such that boiling is more enhanced in some areas than others. The δ13C and 3He/CO2 of gases reflect a dominant mantle origin for CO2 in Yellowstone gas. The mantle signature is most evident at Mud Volcano, which hosts gases with the lowest H2O/CO2, lowest CH4 concentrations and highest He isotope ratios (~16Ra), consistent with either a young subsurface intrusion or less input of crustal and meteoric gas than any other location at Yellowstone. Across the YPVF, He isotope ratios (3He/4He) inversely vary with He concentrations, and reflect varied amounts of long- stored, radiogenic He added to the magmatic endmember within the crust. Similarly, addition of CH4 from organic-rich sediments is common in the eastern thermal areas at Yellowstone. Overall, Yellowstone gases reflect addition of deep, high-temperature magmatic gas (CO2-rich), lower-temperatures crustal gases (4He- and CH4-bearing), and those gases (N2, Ne, Ar) added

  4. Sexual predators, energy development, and conservation in greater Yellowstone.

    Science.gov (United States)

    Berger, Joel; Beckmann, Jon P

    2010-06-01

    In the United States, as elsewhere, a growing debate pits national energy policy and homeland security against biological conservation. In rural communities the extraction of fossil fuels is often encouraged because of the employment opportunities it offers, although the concomitant itinerant workforce is often associated with increased wildlife poaching. We explored possible positive and negative factors associated with energy extraction in the Greater Yellowstone Ecosystem (GYE), an area known for its national parks, intact biological diversity, and some of the New World's longest terrestrial migrations. Specifically, we asked whether counties with different economies-recreation (ski), agrarian (ranching or farming), and energy extractive (petroleum)-differed in healthcare (gauged by the abundance of hospital beds) and in the frequency of sexual predators. The absolute and relative frequency of registered sex offenders grew approximately two to three times faster in areas reliant on energy extraction. Healthcare among counties did not differ. The strong conflation of community dishevel, as reflected by in-migrant sexual predators, and ecological decay in Greater Yellowstone is consistent with patterns seen in similar systems from Ecuador to northern Canada, where social and environmental disarray exist around energy boomtowns. In our case, that groups (albeit with different aims) mobilized campaigns to help maintain the quality of rural livelihoods by protecting open space is a positive sign that conservation can matter, especially in the face of rampant and poorly executed energy extraction projects. Our findings further suggest that the public and industry need stronger regulatory action to instill greater vigilance when and where social factors and land conversion impact biological systems.

  5. Infectious diseases of wolves in Yellowstone

    Science.gov (United States)

    Almberg, Emily S.; Cross, Paul C.; Hudson, Peter J.; Dobson, Andrew P.; Smith, Douglas W.; Stahler, Daniel R.

    2016-01-01

    The summer of 2005 began with such promise for wolves in Yellowstone.  The population had been at an all-time high the last few years, and the wolves appeared to be in good condition.  Several packs had been particularly busy during the breeding season, and early summer pup counts suggested another healthy crop of new wolves rising through the ranks.

  6. Lessons from geothermal gases at Yellowstone

    Science.gov (United States)

    Lowenstern, J. B.; Bergfeld, D.; Evans, W.; Hurwitz, S.

    2015-12-01

    The magma-hydrothermal system of the Yellowstone Plateau Volcanic Field encompasses over ten thousand individual springs, seeps, and fumaroles spread out over >9000 square kilometers, and produces a range of acid, neutral and alkaline waters. A prominent model (Fournier, 1989 and related papers) concludes that many neutral and alkaline fluids found in hot springs and geysers are derived from a uniform, high-enthalpy parent fluid through processes such as deep boiling and mixing with dilute meteoric groundwater. Acid waters are generally condensates of gas-bearing steam that boils off of subsurface geothermal waters. Our recent studies of gases at Yellowstone (Lowenstern et al., 2015 and references therein) are compatible with such a model, but also reveal that gases are largely decoupled from thermal waters due to open-system addition of abundant deep gas to (comparatively) shallow circulating thermal waters. Fumarole emissions at Yellowstone range from gas-rich (up to 15 mol%) composed of deeply derived CO2, He and CH4, to steam-rich emissions (16 RA) and low CH4 and He concentrations and 2) mantle-derived CO2 with much higher CH4 and/or He concentrations and abundant radiogenic He picked up from crustal degassing. Individual thermal areas have distinct CH4/He. It remains unclear whether some gas ratios mainly reflect subsurface geothermal temperatures. Instead, they may simply reflect signatures imparted by local rock types and mixing on timescales too fast for reequilibration. Overall, the gas chemistry reflects a broader view of mantle-crust dynamics than can be appreciated by studies of only dissolved solutes in the neutral and alkaline waters from Yellowstone geysers. Fournier (1989) Ann. Rev. Earth Planet. Sci. v. 17, p. 13-53. Lowenstern et al. (2015) JVGR, v. 302, 87-101.

  7. Dynamics of the Yellowstone hydrothermal system

    Science.gov (United States)

    Hurwitz, Shaul; Lowenstern, Jacob B.

    2014-01-01

    The Yellowstone Plateau Volcanic Field is characterized by extensive seismicity, episodes of uplift and subsidence, and a hydrothermal system that comprises more than 10,000 thermal features, including geysers, fumaroles, mud pots, thermal springs, and hydrothermal explosion craters. The diverse chemical and isotopic compositions of waters and gases derive from mantle, crustal, and meteoric sources and extensive water-gas-rock interaction at variable pressures and temperatures. The thermal features are host to all domains of life that utilize diverse inorganic sources of energy for metabolism. The unique and exceptional features of the hydrothermal system have attracted numerous researchers to Yellowstone beginning with the Washburn and Hayden expeditions in the 1870s. Since a seminal review published a quarter of a century ago, research in many fields has greatly advanced our understanding of the many coupled processes operating in and on the hydrothermal system. Specific advances include more refined geophysical images of the magmatic system, better constraints on the time scale of magmatic processes, characterization of fluid sources and water-rock interactions, quantitative estimates of heat and magmatic volatile fluxes, discovering and quantifying the role of thermophile microorganisms in the geochemical cycle, defining the chronology of hydrothermal explosions and their relation to glacial cycles, defining possible links between hydrothermal activity, deformation, and seismicity; quantifying geyser dynamics; and the discovery of extensive hydrothermal activity in Yellowstone Lake. Discussion of these many advances forms the basis of this review.

  8. Monitoring Greater Yellowstone Ecosystem wetlands: Can long-term monitoring help us understand their future?

    Science.gov (United States)

    Ray, Andrew M.; Sepulveda, Adam; Hossack, Blake R.; Patla, Debra; Thoma, David; Al-Chokhachy, Robert K.; Litt, Andrea R.

    2015-01-01

    In the Greater Yellowstone Ecosystem (GYE), changes in the drying cycles of wetlands have been documented. Wetlands are areas where the water table is at or near the land surface and standing shallow water is present for much or all of the growing season. We discuss how monitoring data can be used to document variation in annual flooding and drying patterns of wetlands monitored across Yellowstone and Grand Teton national parks, investigate how these patterns are related to a changing climate, and explore how drying of wetlands may impact amphibians. The documented declines of some amphibian species are of growing concern to scientists and land managers alike, in part because disappearances have occurred in some of the most protected places. These disappearances are a recognized component of what is being described as Earth’s sixth mass extinction.

  9. Migration of northern yellowstone elk: Implications of spatial structuring

    Science.gov (United States)

    White, P.J.; Proffitt, K.M.; Mech, L.D.; Evans, S.B.; Cunningham, J.A.; Hamlin, K.L.

    2010-01-01

    Migration can enhance survival and recruitment of mammals by increasing access to higher-quality forage or reducing predation risk, or both. We used telemetry locations collected from 140 adult female elk during 20002003 and 20072008 to identify factors influencing the migration of northern Yellowstone elk. Elk wintered in 2 semidistinct herd segments and migrated 10140 km to at least 12 summer areas in Yellowstone National Park (YNP) and nearby areas of Montana. Spring migrations were delayed after winters with increased snow pack, with earlier migration in years with earlier vegetation green-up. Elk wintering at lower elevations outside YNP migrated an average of 13 days earlier than elk at higher elevations. The timing of autumn migrations varied annually, but elk left their summer ranges at about the same time regardless of elevation, wolf numbers, or distance to their wintering areas. Elk monitored for multiple years typically returned to the same summer (96 fidelity, n 52) and winter (61 fidelity, n 41) ranges. Elk that wintered at lower elevations in or near the northwestern portion of the park tended to summer in the western part of YNP (56), and elk that wintered at higher elevations spent summer primarily in the eastern and northern parts of the park (82). Elk did not grossly modify their migration timing, routes, or use areas after wolf restoration. Elk mortality was low during summer and migration (8 of 225 elk-summers). However, spatial segregation and differential mortality and recruitment between herd segments on the northern winter range apparently contributed to a higher proportion of the elk population wintering outside the northwestern portion of YNP and summering in the western portion of the park. This change could shift wolf spatial dynamics more outside YNP and increase the risk of transmission of brucellosis from elk to cattle north of the park. ?? 2010 American Society of Mammalogists.

  10. Modeling survival: application of the Andersen-Gill model to Yellowstone grizzly bears

    Science.gov (United States)

    Johnson, Christopher J.; Boyce, Mark S.; Schwartz, Charles C.; Haroldson, Mark A.

    2004-01-01

     Wildlife ecologists often use the Kaplan-Meier procedure or Cox proportional hazards model to estimate survival rates, distributions, and magnitude of risk factors. The Andersen-Gill formulation (A-G) of the Cox proportional hazards model has seen limited application to mark-resight data but has a number of advantages, including the ability to accommodate left-censored data, time-varying covariates, multiple events, and discontinuous intervals of risks. We introduce the A-G model including structure of data, interpretation of results, and assessment of assumptions. We then apply the model to 22 years of radiotelemetry data for grizzly bears (Ursus arctos) of the Greater Yellowstone Grizzly Bear Recovery Zone in Montana, Idaho, and Wyoming, USA. We used Akaike's Information Criterion (AICc) and multi-model inference to assess a number of potentially useful predictive models relative to explanatory covariates for demography, human disturbance, and habitat. Using the most parsimonious models, we generated risk ratios, hypothetical survival curves, and a map of the spatial distribution of high-risk areas across the recovery zone. Our results were in agreement with past studies of mortality factors for Yellowstone grizzly bears. Holding other covariates constant, mortality was highest for bears that were subjected to repeated management actions and inhabited areas with high road densities outside Yellowstone National Park. Hazard models developed with covariates descriptive of foraging habitats were not the most parsimonious, but they suggested that high-elevation areas offered lower risks of mortality when compared to agricultural areas.

  11. Population growth of Yellowstone grizzly bears: Uncertainty and future monitoring

    Science.gov (United States)

    Harris, R.B.; White, Gary C.; Schwartz, C.C.; Haroldson, M.A.

    2007-01-01

    Grizzly bears (Ursus arctos) in the Greater Yellowstone Ecosystem of the US Rocky Mountains have recently increased in numbers, but remain vulnerable due to isolation from other populations and predicted reductions in favored food resources. Harris et al. (2006) projected how this population might fare in the future under alternative survival rates, and in doing so estimated the rate of population growth, 1983–2002. We address issues that remain from that earlier work: (1) the degree of uncertainty surrounding our estimates of the rate of population change (λ); (2) the effect of correlation among demographic parameters on these estimates; and (3) how a future monitoring system using counts of females accompanied by cubs might usefully differentiate between short-term, expected, and inconsequential fluctuations versus a true change in system state. We used Monte Carlo re-sampling of beta distributions derived from the demographic parameters used by Harris et al. (2006) to derive distributions of λ during 1983–2002 given our sampling uncertainty. Approximate 95% confidence intervals were 0.972–1.096 (assuming females with unresolved fates died) and 1.008–1.115 (with unresolved females censored at last contact). We used well-supported models of Haroldson et al. (2006) and Schwartz et al. (2006a,b,c) to assess the strength of correlations among demographic processes and the effect of omitting them in projection models. Incorporating correlations among demographic parameters yielded point estimates of λ that were nearly identical to those from the earlier model that omitted correlations, but yielded wider confidence intervals surrounding λ. Finally, we suggest that fitting linear and quadratic curves to the trend suggested by the estimated number of females with cubs in the ecosystem, and using AICc model weights to infer population sizes and λ provides an objective means to monitoring approximate population trajectories in addition to demographic

  12. Bighorn sheep (Ovis canadensis) survivorship and habitat studies in Bighorn Canyon National Recreation Area and surrounding lands, Wyoming and Montana, 2000–2003

    Science.gov (United States)

    Schoenecker, Kathryn A.; Singer, Francis J.; Grams, Kayla A.; Roelle, James E.

    2004-01-01

    In the 1850s, bighorn sheep (Ovis canadensis) were numerous and distributed throughout the Bighorn and Pryor Mountains of Montana and Wyoming. After European settlement, bighorn sheep populations declined, and local extinctions occurred in much of their historic range in the western United States. The current bighorn sheep population of Bighorn Canyon National Recreation Area (BICA) is the product of several reintroductions into BICA and surrounding lands. Following a release in 1973 and growth rates near maximum potential of 19.8% per year, the population grew to an estimated peak population of about 211 animals in 1993 and 1994 (Kissell and others, 1996). Recent counts indicate the bighorn sheep population has declined. Kissell and others (1996) reported that the population began to decline rapidly in 1995 and 1996. He noted low ewe:lamb ratios during the decline phase. Bighorn sheep numbers declined to the lowest minimum viable population size of 100 animals recommended by several bighorn sheep experts (Bailey, 1990; Berger, 1990; Smith and others, 1991). National Park Service (NPS) and Bureau of Land Management (BLM) managers were concerned about the decline and requested a study of its causes. In 2000, the U.S. Geological Survey- Biological Resources Division (USGS-BRD) received funding to start a 3-year study of survivorship, condition, and population growth rate of the BICA bighorn sheep population.Several possibilities exist for the bighorn sheep decline. The herd may have experienced a rapid population expansion, followed by a decline to stability at a lower long-term carrying capacity. This pattern of apparently overshooting carrying capacity following an initial release has been reported for a number of ungulates (Caughley, 1976). Disease may have caused the decline; predation and/or competition with wild horses (Equus caballus) may also have been factors. A spatial model of wild horse carrying capacity (Coughenour, 1999) was developed to assist managers

  13. Mapping temperature and radiant geothermal heat flux anomalies in the Yellowstone geothermal system using ASTER thermal infrared data

    Science.gov (United States)

    Vaughan, R. Greg; Lowenstern, Jacob B.; Keszthelyi, Laszlo P.; Jaworowski, Cheryl; Heasler, Henry

    2012-01-01

    The purpose of this work was to use satellite-based thermal infrared (TIR) remote sensing data to measure, map, and monitor geothermal activity within the Yellowstone geothermal area to help meet the missions of both the U.S. Geological Survey Yellowstone Volcano Observatory and the Yellowstone National Park Geology Program. Specifically, the goals were to: 1) address the challenges of remotely characterizing the spatially and temporally dynamic thermal features in Yellowstone by using nighttime TIR data from the Advanced Spaceborne Thermal Emission and Reflection Radiometer (ASTER) and 2) estimate the temperature, geothermal radiant emittance, and radiant geothermal heat flux (GHF) for Yellowstone’s thermal areas (both Park wide and for individual thermal areas). ASTER TIR data (90-m pixels) acquired at night during January and February, 2010, were used to estimate surface temperature, radiant emittance, and radiant GHF from all of Yellowstone’s thermal features, produce thermal anomaly maps, and update field-based maps of thermal areas. A background subtraction technique was used to isolate the geothermal component of TIR radiance from thermal radiance due to insolation. A lower limit for the Yellowstone’s total radiant GHF was established at ~2.0 GW, which is ~30-45% of the heat flux estimated through geochemical (Cl-flux) methods. Additionally, about 5 km2 was added to the geodatabase of mapped thermal areas. This work provides a framework for future satellite-based thermal monitoring at Yellowstone as well as exploration of other volcanic / geothermal systems on a global scale.

  14. Yellowstone wolves and the forces that structure natural systems.

    OpenAIRE

    Andy P Dobson

    2014-01-01

    Since their introduction in 1995 and 1996, wolves have had effects on Yellowstone that ripple across the entire structure of the food web that defines biodiversity in the Northern Rockies ecosystem. Ecological interpretations of the wolves have generated a significant amount of debate about the relative strength of top-down versus bottom-up forces in determining herbivore and vegetation abundance in Yellowstone. Debates such as this are central to the resolution of broader debates about the r...

  15. Effects of exotic species on Yellowstone's grizzly bears

    Science.gov (United States)

    Reinhart, Daniel P.; Haroldson, Mark A.; Mattson, D.J.; Gunther, Kerry A.

    2001-01-01

    Humans have affected grizzly bears (Ursus arctos horribilis) by direct mortality, competition for space and resources, and introduction of exotic species. Exotic organisms that have affected grizzly bears in the Greater Yellowstone Area include common dandelion (Taraxacum officinale), nonnative clovers (Trifolium spp.), domesticated livestock, bovine brucellosis (Brucella abortus), lake trout (Salvelinus namaycush), and white pine blister rust (Cronartium ribicola). Some bears consume substantial amounts of dandelion and clover. However, these exotic foods provide little digested energy compared to higher-quality bear foods. Domestic livestock are of greater energetic value, but use of this food by bears often leads to conflicts with humans and subsequent increases in bear mortality. Lake trout, blister rust, and brucellosis diminish grizzly bears foods. Lake trout prey on native cutthroat trout (Oncorhynchus clarkii) in Yellowstone Lake; white pine blister rust has the potential to destroy native whitebark pine (Pinus albicaulis) stands; and management response to bovine brucellosis, a disease found in the Yellowstone bison (Bison bison) and elk (Cervus elaphus), could reduce populations of these 2 species. Exotic species will likely cause more harm than good for Yellowstone grizzly bears. Managers have few options to mitigate or contain the impacts of exotics on Yellowstone's grizzly bears. Moreover, their potential negative impacts have only begun to unfold. Exotic species may lead to the loss of substantial highquality grizzly bear foods, including much of the bison, trout, and pine seeds that Yellowstone grizzly bears currently depend upon.

  16. Thermomechanical Modeling of the Formation of a Multilevel, Crustal-Scale Magmatic System by the Yellowstone Plume

    Science.gov (United States)

    Colón, D. P.; Bindeman, I. N.; Gerya, T. V.

    2018-05-01

    Geophysical imaging of the Yellowstone supervolcano shows a broad zone of partial melt interrupted by an amagmatic gap at depths of 15-20 km. We reproduce this structure through a series of regional-scale magmatic-thermomechanical forward models which assume that magmatic dikes stall at rheologic discontinuities in the crust. We find that basaltic magmas accumulate at the Moho and at the brittle-ductile transition, which naturally forms at depths of 5-10 km. This leads to the development of a 10- to 15-km thick midcrustal sill complex with a top at a depth of approximately 10 km, consistent with geophysical observations of the pre-Yellowstone hot spot track. We show a linear relationship between melting rates in the mantle and rhyolite eruption rates along the hot spot track. Finally, melt production rates from our models suggest that the Yellowstone plume is 175°C hotter than the surrounding mantle and that the thickness of the overlying lithosphere is 80 km.

  17. Trend of the Yellowstone Grizzly Bear Population

    International Nuclear Information System (INIS)

    Eberhardt, L.L.; Breiwick, J.M.

    2010-01-01

    Yellowstone's grizzlies (Ursus arctos) have been studied for more than 40 years. Radio telemetry has been used to obtain estimates of the rate of increase of the population, with results reported by Schwartz et al. (2006). Counts of females with cubs-of-the-year unduplicated also provide an index of abundance and are the primary subject of this report. An exponential model was fitted to n=24 such counts, using nonlinear least squares. Estimates of the rate of increase, r, were about 0.053. 95% confidence intervals, were obtained by several different methods, and all had lower limits substantially above zero, indicating that the population has been increasing steadily, in contrast to the results of Schwartz et al. (2006), which could not exclude a decreasing population. The grizzly data have been repeatedly mis-used in current literature for reasons explained here.

  18. Trend of the Yellowstone Grizzly Bear Population

    Directory of Open Access Journals (Sweden)

    L. L. Eberhardt

    2010-01-01

    Full Text Available Yellowstone's grizzlies (Ursus arctos have been studied for more than 40 years. Radiotelemetry has been used to obtain estimates of the rate of increase of the population, with results reported by Schwartz et al. (2006. Counts of females with cubs-of-the-year “unduplicated” also provide an index of abundance and are the primary subject of this report. An exponential model was fitted to n=24 such counts, using nonlinear leastsquares. Estimates of the rate of increase, r, were about 0.053. 95% confidence intervals, were obtained by several different methods, and all had lower limits substantially above zero, indicating that the population has been increasing steadily, in contrast to the results of Schwartz et al. (2006, which could not exclude a decreasing population. The grizzly data have been repeatedly mis-used in current literature for reasons explained here.

  19. Influence of whitebark pine decline on fall habitat use and movements of grizzly bears in the Greater Yellowstone Ecosystem.

    Science.gov (United States)

    Costello, Cecily M; van Manen, Frank T; Haroldson, Mark A; Ebinger, Michael R; Cain, Steven L; Gunther, Kerry A; Bjornlie, Daniel D

    2014-05-01

    When abundant, seeds of the high-elevation whitebark pine (WBP; Pinus albicaulis) are an important fall food for grizzly bears (Ursus arctos) in the Greater Yellowstone Ecosystem. Rates of bear mortality and bear/human conflicts have been inversely associated with WBP productivity. Recently, mountain pine beetles (Dendroctonus ponderosae) have killed many cone-producing WBP trees. We used fall (15 August-30 September) Global Positioning System locations from 89 bear years to investigate temporal changes in habitat use and movements during 2000-2011. We calculated Manly-Chesson (MC) indices for selectivity of WBP habitat and secure habitat (≥500 m from roads and human developments), determined dates of WBP use, and documented net daily movement distances and activity radii. To evaluate temporal trends, we used regression, model selection, and candidate model sets consisting of annual WBP production, sex, and year. One-third of sampled grizzly bears had fall ranges with little or no mapped WBP habitat. Most other bears (72%) had a MC index above 0.5, indicating selection for WBP habitats. From 2000 to 2011, mean MC index decreased and median date of WBP use shifted about 1 week later. We detected no trends in movement indices over time. Outside of national parks, there was no correlation between the MC indices for WBP habitat and secure habitat, and most bears (78%) selected for secure habitat. Nonetheless, mean MC index for secure habitat decreased over the study period during years of good WBP productivity. The wide diet breadth and foraging plasticity of grizzly bears likely allowed them to adjust to declining WBP. Bears reduced use of WBP stands without increasing movement rates, suggesting they obtained alternative fall foods within their local surroundings. However, the reduction in mortality risk historically associated with use of secure, high-elevation WBP habitat may be diminishing for bears residing in multiple-use areas.

  20. Influence of whitebark pine decline on fall habitat use and movements of grizzly bears in the Greater Yellowstone Ecosystem

    Science.gov (United States)

    Costello, Cecily M.; van Manen, Frank T.; Haroldson, Mark A.; Ebinger, Michael R.; Cain, Steven L.; Gunther, Kerry A.; Bjornlie, Daniel D.

    2014-01-01

    When abundant, seeds of the high-elevation whitebark pine (WBP; Pinus albicaulis) are an important fall food for grizzly bears (Ursus arctos) in the Greater Yellowstone Ecosystem. Rates of bear mortality and bear/human conflicts have been inversely associated with WBP productivity. Recently, mountain pine beetles (Dendroctonus ponderosae) have killed many cone-producing WBP trees. We used fall (15 August–30 September) Global Positioning System locations from 89 bear years to investigate temporal changes in habitat use and movements during 2000–2011. We calculated Manly–Chesson (MC) indices for selectivity of WBP habitat and secure habitat (≥500 m from roads and human developments), determined dates of WBP use, and documented net daily movement distances and activity radii. To evaluate temporal trends, we used regression, model selection, and candidate model sets consisting of annual WBP production, sex, and year. One-third of sampled grizzly bears had fall ranges with little or no mapped WBP habitat. Most other bears (72%) had a MC index above 0.5, indicating selection for WBP habitats. From 2000 to 2011, mean MC index decreased and median date of WBP use shifted about 1 week later. We detected no trends in movement indices over time. Outside of national parks, there was no correlation between the MC indices for WBP habitat and secure habitat, and most bears (78%) selected for secure habitat. Nonetheless, mean MC index for secure habitat decreased over the study period during years of good WBP productivity. The wide diet breadth and foraging plasticity of grizzly bears likely allowed them to adjust to declining WBP. Bears reduced use of WBP stands without increasing movement rates, suggesting they obtained alternative fall foods within their local surroundings. However, the reduction in mortality risk historically associated with use of secure, high-elevation WBP habitat may be diminishing for bears residing in multiple-use areas.

  1. Controls on Magmatic and Hydrothermal Processes at Yellowstone Supervolcano: The Wideband Magnetotelluric Component of an Integrated MT/Seismic Investigation

    Science.gov (United States)

    Schultz, A.; Bennington, N. L.; Bowles-martinez, E.; Imamura, N.; Cronin, R. A.; Miller, D. J.; Hart, L.; Gurrola, R. M.; Neal, B. A.; Scholz, K.; Fry, B.; Carbonari, R.

    2017-12-01

    Previous seismic and magnetotelluric (MT) studies beneath Yellowstone (YS) have provided insight into the origin and migration of magmatic fluids within the volcanic system. However, important questions remain concerning the generation of magmatism at YS, the migration and storage of these magmatic fluids, as well as their relationships to hydrothermal expressions. Analysis of regional-scale EarthScope MT data collected previously suggests a relative absence of continuity in crustal partial melt accumulations directly beneath YS. This is in contrast to some seismic interpretations, although such long-period MT data have limited resolving power in the upper-to-mid crustal section. A wideband MT experiment was designed as a component of an integrated MT/seismic project to examine: the origin and location of magmatic fluids at upper mantle/lower crustal depths, the preferred path of migration for these magmatic fluids into the mid- to upper-crust, the resulting distribution of the magma reservoir, the composition of the magma reservoir, and implications for future volcanism at YS. A high-resolution wideband MT survey was carried out in the YS region in the summer of 2017, with more than forty-five wideband stations installed within and immediately surrounding the YS National Park boundary. These data provided nearly six decades of bandwidth ( 10-3 Hz -to- 103 Hz). Extraordinary permitting restrictions prevented us from using conventional installation methods at many of our sites, and an innovative "no-dig" subaerial method of wideband MT was developed and used successfully. Using these new data along with existing MT datasets, we are inverting for the 3D resistivity structure at upper crustal through upper mantle scales at YS. Complementary to this MT work, a joint inversion for the 3D crustal velocity structure is being carried out using both ambient noise and earthquake travel time data. Taken together, these data should better constrain the crustal velocity

  2. Predation risk, elk, and aspen: tests of a behaviorally mediated trophic cascade in the Greater Yellowstone Ecosystem.

    Science.gov (United States)

    Winnie, John A

    2012-12-01

    Aspen in the Greater Yellowstone Ecosystem are hypothesized to be recovering from decades of heavy browsing by elk due to a behaviorally mediated trophic cascade (BMTC). Several authors have suggested that wolves interact with certain terrain features, creating places of high predation risk at fine spatial scales, and that elk avoid these places, which creates refugia for plants. This hypothesized BMTC could release aspen from elk browsing pressure, leading to a patchy recovery in places of high risk. I tested whether four specific, hypothesized fine-scale risk factors are correlated with changes in current elk browsing pressure on aspen, or with aspen recruitment since wolf reintroduction, in the Daly Creek drainage in Yellowstone National Park, and near two aspen enclosures outside of the park boundary. Aspen were not responding to hypothesized fine-scale risk factors in ways consistent with the current BMTC hypothesis.

  3. Aerial radiological survey of the Fermi National accelerator Laboratory and surrounding area, Batavia, Illinois. Data of survey: 12-14 May 1977

    International Nuclear Information System (INIS)

    Jobst, J.E.

    1981-01-01

    An aerial radiological survey was conducted over the facilities of the Fermi National Accelerator Laboratory (Fermilab) near Batavia, Illinois on 12-14 May 1977. The survey was flown at an altitude of 91 m by a helicopter containing 20 sodium iodide detectors. The line spacing was also 91 m. Enhanced gamma exposure rate levels, which could be attributed to Fermilab operations, were observed at seven locations. One additional anomaly, not related to the Laboratory, was also discovered

  4. Radiological survey of the area surrounding the National Reactor Testing Station, Idaho Falls, Idaho. Date of survey: 1 and 2 February 1972

    International Nuclear Information System (INIS)

    1974-01-01

    The Aerial Radiological Measuring System (ARMS) was used to survey the National Reactor Testing Station (NRTS) during February 1972. The purpose of the survey was primarily to identify the presence of Ru-106 and Rh-106 in a release from the Chemical Processing Plant at NRTS. Additionally, the gamma-ray terrestrial exposure rate levels were mapped and the distribution of any man-made isotopes was located and defined

  5. Uplift, thermal unrest and magma intrusion at Yellowstone caldera.

    Science.gov (United States)

    Wicks, Charles W; Thatcher, Wayne; Dzurisin, Daniel; Svarc, Jerry

    2006-03-02

    The Yellowstone caldera, in the western United States, formed approximately 640,000 years ago when an explosive eruption ejected approximately 1,000 km3 of material. It is the youngest of a series of large calderas that formed during sequential cataclysmic eruptions that began approximately 16 million years ago in eastern Oregon and northern Nevada. The Yellowstone caldera was largely buried by rhyolite lava flows during eruptions that occurred from approximately 150,000 to approximately 70,000 years ago. Since the last eruption, Yellowstone has remained restless, with high seismicity, continuing uplift/subsidence episodes with movements of approximately 70 cm historically to several metres since the Pleistocene epoch, and intense hydrothermal activity. Here we present observations of a new mode of surface deformation in Yellowstone, based on radar interferometry observations from the European Space Agency ERS-2 satellite. We infer that the observed pattern of uplift and subsidence results from variations in the movement of molten basalt into and out of the Yellowstone volcanic system.

  6. Density-dependent intraspecific aggression regulates survival in northern Yellowstone wolves (Canis lupus).

    Science.gov (United States)

    Cubaynes, Sarah; MacNulty, Daniel R; Stahler, Daniel R; Quimby, Kira A; Smith, Douglas W; Coulson, Tim

    2014-11-01

    Understanding the population dynamics of top-predators is essential to assess their impact on ecosystems and to guide their management. Key to this understanding is identifying the mechanisms regulating vital rates. Determining the influence of density on survival is necessary to understand the extent to which human-caused mortality is compensatory or additive. In wolves (Canis lupus), empirical evidence for density-dependent survival is lacking. Dispersal is considered the principal way in which wolves adjust their numbers to prey supply or compensate for human exploitation. However, studies to date have primarily focused on exploited wolf populations, in which density-dependent mechanisms are likely weak due to artificially low wolf densities. Using 13 years of data on 280 collared wolves in Yellowstone National Park, we assessed the effect of wolf density, prey abundance and population structure, as well as winter severity, on age-specific survival in two areas (prey-rich vs. prey-poor) of the national park. We further analysed cause-specific mortality and explored the factors driving intraspecific aggression in the prey-rich northern area of the park. Overall, survival rates decreased during the study. In northern Yellowstone, density dependence regulated adult survival through an increase in intraspecific aggression, independent of prey availability. In the interior of the park, adult survival was less variable and density-independent, despite reduced prey availability. There was no effect of prey population structure in northern Yellowstone, or of winter severity in either area. Survival was similar among yearlings and adults, but lower for adults older than 6 years. Our results indicate that density-dependent intraspecific aggression is a major driver of adult wolf survival in northern Yellowstone, suggesting intrinsic density-dependent mechanisms have the potential to regulate wolf populations at high ungulate densities. When low prey availability or high

  7. Animal migration amid shifting patterns of phenology and predation: lessons from a Yellowstone elk herd.

    Science.gov (United States)

    Middleton, Arthur D; Kauffman, Matthew J; McWhirter, Douglas E; Cook, John G; Cook, Rachel C; Nelson, Abigail A; Jimenez, Michael D; Klaver, Robert W

    2013-06-01

    Migration is a striking behavioral strategy by which many animals enhance resource acquisition while reducing predation risk. Historically, the demographic benefits of such movements made migration common, but in many taxa the phenomenon is considered globally threatened. Here we describe a long-term decline in the productivity of elk (Cervus elaphus) that migrate through intact wilderness areas to protected summer ranges inside Yellowstone National Park, USA. We attribute this decline to a long-term reduction in the demographic benefits that ungulates typically gain from migration. Among migratory elk, we observed a 21-year, 70% reduction in recruitment and a 4-year, 19% depression in their pregnancy rate largely caused by infrequent reproduction of females that were young or lactating. In contrast, among resident elk, we have recently observed increasing recruitment and a high rate of pregnancy. Landscape-level changes in habitat quality and predation appear to be responsible for the declining productivity of Yellowstone migrants. From 1989 to 2009, migratory elk experienced an increasing rate and shorter duration of green-up coincident with warmer spring-summer temperatures and reduced spring precipitation, also consistent with observations of an unusually severe drought in the region. Migrants are also now exposed to four times as many grizzly bears (Ursus arctos) and wolves (Canis lupus) as resident elk. Both of these restored predators consume migratory elk calves at high rates in the Yellowstone wilderness but are maintained at low densities via lethal management and human disturbance in the year-round habitats of resident elk. Our findings suggest that large-carnivore recovery and drought, operating simultaneously along an elevation gradient, have disproportionately influenced the demography of migratory elk. Many migratory animals travel large geographic distances between their seasonal ranges. Changes in land use and climate that disparately influence

  8. Animal migration amid shifting patterns of phenology and predation: Lessons from a Yellowstone elk herd

    Science.gov (United States)

    Middleton, Arthur D.; Kauffman, Matthew J.; McWhirter, Douglas E.; Cook, John G.; Cook, Rachel C.; Nelson, Abigail A.; Jimenez, Michael D.; Klaver, Robert W.

    2013-01-01

    Migration is a striking behavioral strategy by which many animals enhance resource acquisition while reducing predation risk. Historically, the demographic benefits of such movements made migration common, but in many taxa the phenomenon is considered globally threatened. Here we describe a long-term decline in the productivity of elk (Cervus elaphus) that migrate through intact wilderness areas to protected summer ranges inside Yellowstone National Park, USA. We attribute this decline to a long-term reduction in the demographic benefits that ungulates typically gain from migration. Among migratory elk, we observed a 21-year, 70% reduction in recruitment and a 4-year, 19% depression in their pregnancy rate largely caused by infrequent reproduction of females that were young or lactating. In contrast, among resident elk, we have recently observed increasing recruitment and a high rate of pregnancy. Landscape-level changes in habitat quality and predation appear to be responsible for the declining productivity of Yellowstone migrants. From 1989 to 2009, migratory elk experienced an increasing rate and shorter duration of green-up coincident with warmer spring–summer temperatures and reduced spring precipitation, also consistent with observations of an unusually severe drought in the region. Migrants are also now exposed to four times as many grizzly bears (Ursus arctos) and wolves (Canis lupus) as resident elk. Both of these restored predators consume migratory elk calves at high rates in the Yellowstone wilderness but are maintained at low densities via lethal management and human disturbance in the year-round habitats of resident elk. Our findings suggest that large-carnivore recovery and drought, operating simultaneously along an elevation gradient, have disproportionately influenced the demography of migratory elk. Many migratory animals travel large geographic distances between their seasonal ranges. Changes in land use and climate that disparately influence

  9. What is “natural”? : Yellowstone elk population - A case study

    Science.gov (United States)

    Keigley, R.B.; Wagner, Frederic H.

    2000-01-01

    Ecology analyzes the structure and function of ecosystems at all points along the continuum of human disturbance, from so-called pristine forests to urban backyards. Undisturbed systems provide reference points at one end of the spectrum, and nature reserves and parks are highly valued because they can provide unique examples of such ecosystems. Unfortunately the concept of “natural” or pristine is not that easy to define. Indeed, although ecologists have considered pre-Columbian, western-hemisphere ecosystems to have been largely unaltered by human action, and have termed their state “natural” or “pristine,” evidence from archaeology challenges this view. U.S. and Canadian national parks are charged with preserving the “natural,” and thus need to be able to understand and manage for the “natural.” A pivotal “natural” question in Yellowstone National Park management is the size of the northern-range, wintering elk population at Park establishment in 1872, argued both to have been small and large. Integrating and quantifying several sources of evidence provides a consistent picture of a low population (ca. 5,000–6,000), largely migrating out of the northern range in winter, with little vegetation impact. If we accept this conclusion about what is natural for the Yellowstone ecosystem, then it dramatically alters how we view management alternatives for the Park, which currently supports a northern wintering herd of up to ˜ 25,000 elk.

  10. Protocols for geologic hazards response by the Yellowstone Volcano Observatory

    Science.gov (United States)

    ,

    2010-01-01

    The Yellowstone Plateau hosts an active volcanic system, with subterranean magma (molten rock), boiling, pressurized waters, and a variety of active faults with significant earthquake hazards. Within the next few decades, light-to-moderate earthquakes and steam explosions are certain to occur. Volcanic eruptions are less likely, but are ultimately inevitable in this active volcanic region. This document summarizes protocols, policies, and tools to be used by the Yellowstone Volcano Observatory (YVO) during earthquakes, hydrothermal explosions, or any geologic activity that could lead to a volcanic eruption.

  11. Noninvasive genetic population survey of snow leopards (Panthera uncia in Kangchenjunga conservation area, Shey Phoksundo National Park and surrounding buffer zones of Nepal

    Directory of Open Access Journals (Sweden)

    Karmacharya Dibesh B

    2011-11-01

    Full Text Available Abstract Background The endangered snow leopard is found throughout major mountain ranges of Central Asia, including the remote Himalayas. However, because of their elusive behavior, sparse distribution, and poor access to their habitat, there is a lack of reliable information on their population status and demography, particularly in Nepal. Therefore, we utilized noninvasive genetic techniques to conduct a preliminary snow leopard survey in two protected areas of Nepal. Results A total of 71 putative snow leopard scats were collected and analyzed from two different areas; Shey Phoksundo National Park (SPNP in the west and Kangchanjunga Conservation Area (KCA in the east. Nineteen (27% scats were genetically identified as snow leopards, and 10 (53% of these were successfully genotyped at 6 microsatellite loci. Two samples showed identical genotype profiles indicating a total of 9 individual snow leopards. Four individual snow leopards were identified in SPNP (1 male and 3 females and five (2 males and 3 females in KCA. Conclusions We were able to confirm the occurrence of snow leopards in both study areas and determine the minimum number present. This information can be used to design more in-depth population surveys that will enable estimation of snow leopard population abundance at these sites.

  12. Grizzly bear predation links the loss of native trout to the demography of migratory elk in Yellowstone.

    Science.gov (United States)

    Middleton, Arthur D; Morrison, Thomas A; Fortin, Jennifer K; Robbins, Charles T; Proffitt, Kelly M; White, P J; McWhirter, Douglas E; Koel, Todd M; Brimeyer, Douglas G; Fairbanks, W Sue; Kauffman, Matthew J

    2013-07-07

    The loss of aquatic subsidies such as spawning salmonids is known to threaten a number of terrestrial predators, but the effects on alternative prey species are poorly understood. At the heart of the Greater Yellowstone ecosystem, an invasion of lake trout has driven a dramatic decline of native cutthroat trout that migrate up the shallow tributaries of Yellowstone Lake to spawn each spring. We explore whether this decline has amplified the effect of a generalist consumer, the grizzly bear, on populations of migratory elk that summer inside Yellowstone National Park (YNP). Recent studies of bear diets and elk populations indicate that the decline in cutthroat trout has contributed to increased predation by grizzly bears on the calves of migratory elk. Additionally, a demographic model that incorporates the increase in predation suggests that the magnitude of this diet shift has been sufficient to reduce elk calf recruitment (4-16%) and population growth (2-11%). The disruption of this aquatic-terrestrial linkage could permanently alter native species interactions in YNP. Although many recent ecological changes in YNP have been attributed to the recovery of large carnivores--particularly wolves--our work highlights a growing role of human impacts on the foraging behaviour of grizzly bears.

  13. Grizzly bear predation links the loss of native trout to the demography of migratory elk in Yellowstone

    Science.gov (United States)

    Middleton, Arthur D.; Morrison, Thomas A.; Fortin, Jennifer K.; Robbins, Charles T.; Proffitt, Kelly M.; White, P.J.; McWhirter, Douglas E.; Koel, Todd M.; Brimeyer, Douglas G.; Fairbanks, W. Sue; Kauffman, Matthew J.

    2013-01-01

    The loss of aquatic subsidies such as spawning salmonids is known to threaten a number of terrestrial predators, but the effects on alternative prey species are poorly understood. At the heart of the Greater Yellowstone ecosystem, an invasion of lake trout has driven a dramatic decline of native cutthroat trout that migrate up the shallow tributaries of Yellowstone Lake to spawn each spring. We explore whether this decline has amplified the effect of a generalist consumer, the grizzly bear, on populations of migratory elk that summer inside Yellowstone National Park (YNP). Recent studies of bear diets and elk populations indicate that the decline in cutthroat trout has contributed to increased predation by grizzly bears on the calves of migratory elk. Additionally, a demographic model that incorporates the increase in predation suggests that the magnitude of this diet shift has been sufficient to reduce elk calf recruitment (4–16%) and population growth (2–11%). The disruption of this aquatic–terrestrial linkage could permanently alter native species interactions in YNP. Although many recent ecological changes in YNP have been attributed to the recovery of large carnivores—particularly wolves—our work highlights a growing role of human impacts on the foraging behaviour of grizzly bears.

  14. Ghosts of yellowstone: multi-decadal histories of wildlife populations captured by bones on a modern landscape.

    Directory of Open Access Journals (Sweden)

    Joshua H Miller

    Full Text Available Natural accumulations of skeletal material (death assemblages have the potential to provide historical data on species diversity and population structure for regions lacking decades of wildlife monitoring, thereby contributing valuable baseline data for conservation and management strategies. Previous studies of the ecological and temporal resolutions of death assemblages from terrestrial large-mammal communities, however, have largely focused on broad patterns of community composition in tropical settings. Here, I expand the environmental sampling of large-mammal death assemblages into a temperate biome and explore more demanding assessments of ecological fidelity by testing their capacity to record past population fluctuations of individual species in the well-studied ungulate community of Yellowstone National Park (Yellowstone. Despite dramatic ecological changes following the 1988 wildfires and 1995 wolf re-introduction, the Yellowstone death assemblage is highly faithful to the living community in species richness and community structure. These results agree with studies of tropical death assemblages and establish the broad capability of vertebrate remains to provide high-quality ecological data from disparate ecosystems and biomes. Importantly, the Yellowstone death assemblage also correctly identifies species that changed significantly in abundance over the last 20 to ∼80 years and the directions of those shifts (including local invasions and extinctions. The relative frequency of fresh versus weathered bones for individual species is also consistent with documented trends in living population sizes. Radiocarbon dating verifies the historical source of bones from Equus caballus (horse: a functionally extinct species. Bone surveys are a broadly valuable tool for obtaining population trends and baseline shifts over decadal-to-centennial timescales.

  15. Trophic cascades from wolves to grizzly bears in Yellowstone.

    Science.gov (United States)

    Ripple, William J; Beschta, Robert L; Fortin, Jennifer K; Robbins, Charles T

    2014-01-01

    We explored multiple linkages among grey wolves (Canis lupus), elk (Cervus elaphus), berry-producing shrubs and grizzly bears (Ursus arctos) in Yellowstone National Park. We hypothesized competition between elk and grizzly bears whereby, in the absence of wolves, increases in elk numbers would increase browsing on berry-producing shrubs and decrease fruit availability to grizzly bears. After wolves were reintroduced and with a reduced elk population, we hypothesized there would be an increase in the establishment of berry-producing shrubs, such as serviceberry (Amelanchier alnifolia), which is a major berry-producing plant. We also hypothesized that the percentage fruit in the grizzly bear diet would be greater after than before wolf reintroduction. We compared the frequency of fruit in grizzly bear scats to elk densities prior to wolf reintroduction during a time of increasing elk densities (1968-1987). For a period after wolf reintroduction, we calculated the percentage fruit in grizzly bear scat by month based on scats collected in 2007-2009 (n = 778 scats) and compared these results to scat data collected before wolf reintroduction. Additionally, we developed an age structure for serviceberry showing the origination year of stems in a northern range study area. We found that over a 19-year period, the percentage frequency of fruit in the grizzly diet (6231 scats) was inversely correlated (P wolves and other large carnivores on elk, a reduced and redistributed elk population, decreased herbivory and increased production of plant-based foods that may aid threatened grizzly bears. © 2013 The Authors. Journal of Animal Ecology © 2013 British Ecological Society.

  16. Yellowstone as an Analog for Thermal-Hydrological-Chemical Processes at Yucca Mountain

    International Nuclear Information System (INIS)

    Dobson, P. F.; Kneafsey, T. J.; Simmons, A.; Hulen, J.

    2001-01-01

    Enhanced water-rock interaction resulting from the emplacement of heat-generating nuclear waste in the potential geologic repository at Yucca Mountain, Nevada, may result in changes to fluid flow (resulting from mineral dissolution and precipitation in condensation and boiling zones, respectively). Studies of water-rock interaction in active and fossil geothermal systems (natural analogs) provide evidence for changes in permeability and porosity resulting from thermal-hydrological-chemical (THC) processes. The objective of this research is to document the effects of coupled THC processes at Yellowstone and then examine how differences in scale could influence the impact that these processes may have on the Yucca Mountain system. Subsurface samples from Yellowstone National Park, one of the largest active geothermal systems in the world, contain some the best examples of hydrothermal self-sealing found in geothermal systems. We selected core samples from two USGS research drill holes from the transition zone between conductive and convective portions of the geothermal system (where sealing was reported to occur). We analyzed the core, measuring the permeability, porosity, and grain density of selected samples to evaluate how lithology, texture, and degree of hydrothermal alteration influence matrix and fracture permeability

  17. Coefficients of productivity for Yellowstone's grizzly bear habitat

    Science.gov (United States)

    Mattson, David John; Barber, Kim; Maw, Ralene; Renkin, Roy

    2004-01-01

    This report describes methods for calculating coefficients used to depict habitat productivity for grizzly bears in the Yellowstone ecosystem. Calculations based on these coefficients are used in the Yellowstone Grizzly Bear Cumulative Effects Model to map the distribution of habitat productivity and account for the impacts of human facilities. The coefficients of habitat productivity incorporate detailed information that was collected over a 20-year period (1977-96) on the foraging behavior of Yellowstone's bears and include records of what bears were feeding on, when and where they fed, the extent of that feeding activity, and relative measures of the quantity consumed. The coefficients also incorporate information, collected primarily from 1986 to 1992, on the nutrient content of foods that were consumed, their digestibility, characteristic bite sizes, and the energy required to extract and handle each food. Coefficients were calculated for different time periods and different habitat types, specific to different parts of the Yellowstone ecosystem. Stratifications included four seasons of bear activity (spring, estrus, early hyperphagia, late hyperphagia), years when ungulate carrion and whitebark pine seed crops were abundant versus not, areas adjacent to (bear activity in each region, habitat type, and time period were incorporated into calculations, controlling for the effects of proximity to human facilities. The coefficients described in this report and associated estimates of grizzly bear habitat productivity are unique among many efforts to model the conditions of bear habitat because calculations include information on energetics derived from the observed behavior of radio-marked bears.

  18. Sagebrush-ungulate relationships on the Northern Yellowstone Winter Range

    Science.gov (United States)

    Carl L. Wambolt

    2005-01-01

    Sagebrush (Artemisia) taxa have historically been the landscape dominants over much of the Northern Yellowstone Winter Range (NYWR). Their importance to the unnaturally large ungulate populations on the NYWR throughout the twentieth century has been recognized since the 1920s. Sagebrush-herbivore ecology has been the focus of research on the NYWR for...

  19. Changing numbers of spawning cutthroat trout in tributary streams of Yellowstone Lake and estimates of grizzly bears visiting streams from DNA

    Science.gov (United States)

    Haroldson, M.A.; Gunther, K.A.; Reinhart, Daniel P.; Podruzny, S.R.; Cegelski, C.; Waits, L.; Wyman, T.C.; Smith, J.

    2005-01-01

    to produce annual estimates of grizzly bears visiting streams. Approximately 68 grizzly bears visited the vicinity of cutthroat trout spawning streams annually. Thus, approximately 14–21% of grizzly bears in the Greater Yellowstone Ecosystem (GYE) may have used this threatened food resource annually. Yellowstone National Park (YNP) is attempting to control the lake trout population in Yellowstone Lake; our results underscore the importance of that effort to grizzly bears.

  20. 36 CFR 7.13 - Yellowstone National Park.

    Science.gov (United States)

    2010-07-01

    ... characteristics. (iv) Modifying or disabling a snowcoach's original pollution control equipment is prohibited... Loop Road from its junction with the East Entrance Road to Canyon Junction. (J) The South Canyon Rim... noon and 9 p.m. each day. (N) North Canyon Rim Drive, between noon and 9 p.m. each day. (O) Riverside...

  1. Iron Homeostasis in Yellowstone National Park Hot Spring Microbial Communities

    Science.gov (United States)

    Brown, I.; Tringe, S. G.; Franklin, H.; Bryant, D. A.; Klatt, C. G.; Sarkisova, S. A.; Guevara, M.

    2010-01-01

    It has been postulated that life may have originated on Earth, and possibly on Mars, in association with hydrothermal activity and high concentrations of ferrous iron. However, it is not clear how an iron-rich thermal hydrosphere could be hospitable to microbes, since reduced iron appears to stimulate oxidative stress in all domains of life and particularly in oxygenic phototrophs. Therefore, the study of microbial diversity in iron-depositing hot springs (IDHS) and the mechanisms of iron homeostasis and suppression of oxidative stress may help elucidate how Precambrian organisms could withstand the extremely high concentrations of reactive oxygen species (ROS) produced by interaction between environmental Fe(2+) and O2. Proteins and clusters of orthologous groups (COGs) involved in the maintenance of Fe homeostasis found in cyanobacteria (CB) inhabiting environments with high and low [Fe] were main target of this analysis. Preliminary results of the analysis suggest that the Chocolate Pots (CP) microbial community is heavily dominated by phototrophs from the cyanobacteria (CB), Chloroflexi and Chlorobi phyla, while the Mushroom Spring (MS) effluent channel harbors a more diverse community in which Chloroflexi are the dominant phototrophs. It is speculated that CB inhabiting IDHS have an increased tolerance to both high concentrations of Fe(2+) and ROS produced in the Fenton reaction. This hypothesis was explored via a comparative analysis of the diversity of proteins and COGs involved in Fe and redox homeostasis in the CP and MS microbiomes.

  2. Denning of grizzly bears in the Yellowstone National Park area

    Science.gov (United States)

    Judd, Steven L.; Knight, Richard R.; Blanchard, Bonnie M.

    1986-01-01

    Radiotelemetry was used to locate 101 grizzly bear (Ursus arctos) dens from 1975 to 1980; 35 dens were examined on the ground. Pregnant females denned in late October, and most other bears denned by mid-November. Duration of denning average 113, 132, and 170 days for males, females, and females with new cubs, respectively. Males emerged from mid-February to late March, followed by single females and females with yearlings and 2-year-olds. Females with new cubs emerged from early mid-April. Den sites were associated with moderate tree cover (26%-75% canopy cover) on 30°-60° slopes. Dens occurred on all aspects, although northerly exposures were most common. Grizzly bears usually dug new dens but occasionally used natural cavities or a den from a previous year. Males usually dug larger dens than females with young. Eight excavated and 2 natural dens of the 35 examined dens were used for more than 1 year.

  3. Alternative Fuels Data Center: Yellowstone National Park Commits to

    Science.gov (United States)

    Alternative Fuels Sept. 16, 2017 Photo of a truck Phoenix Utility Fleet Drives Smarter with Biodiesel Aug. 26 Cooking Oil Powers Biodiesel Vehicles in Rhode Island July 14, 2017 Photo of a truck Idaho Transports Mail Home Runs on Biodiesel in North Carolina June 9, 2017 Photo of a bus New Hampshire Cleans up with

  4. Faunal isotope records reveal trophic and nutrient dynamics in twentieth century Yellowstone grasslands.

    Science.gov (United States)

    Fox-Dobbs, Kena; Nelson, Abigail A; Koch, Paul L; Leonard, Jennifer A

    2012-10-23

    Population sizes and movement patterns of ungulate grazers and their predators have fluctuated dramatically over the past few centuries, largely owing to overharvesting, land-use change and historic management. We used δ(13)C and δ(15)N values measured from bone collagen of historic and recent gray wolves and their potential primary prey from Yellowstone National Park to gain insight into the trophic dynamics and nutrient conditions of historic and modern grasslands. The diet of reintroduced wolves closely parallels that of the historic population. We suggest that a significant shift in faunal δ(15)N values over the past century reflects impacts of anthropogenic environmental changes on grassland ecosystems, including grazer-mediated shifts in grassland nitrogen cycle processes.

  5. Yellowstone wolf (Canis lupus) denisty predicted by elk (Cervus elaphus) biomass

    Science.gov (United States)

    Mech, L. David; Barber-Meyer, Shannon

    2015-01-01

    The Northern Range (NR) of Yellowstone National Park (YNP) hosts a higher prey biomass density in the form of elk (Cervus elaphus L., 1758) than any other system of gray wolves (Canis lupus L., 1758) and prey reported. Therefore, it is important to determine whether that wolf–prey system fits a long-standing model relating wolf density to prey biomass. Using data from 2005 to 2012 after elk population fluctuations dampened 10 years subsequent to wolf reintroduction, we found that NR prey biomass predicted wolf density. This finding and the trajectory of the regression extend the validity of the model to prey densities 19% higher than previous data and suggest that the model would apply to wolf–prey systems of even higher prey biomass.

  6. Population size estimation in Yellowstone wolves with error-prone noninvasive microsatellite genotypes.

    Science.gov (United States)

    Creel, Scott; Spong, Goran; Sands, Jennifer L; Rotella, Jay; Zeigle, Janet; Joe, Lawrence; Murphy, Kerry M; Smith, Douglas

    2003-07-01

    Determining population sizes can be difficult, but is essential for conservation. By counting distinct microsatellite genotypes, DNA from noninvasive samples (hair, faeces) allows estimation of population size. Problems arise because genotypes from noninvasive samples are error-prone, but genotyping errors can be reduced by multiple polymerase chain reaction (PCR). For faecal genotypes from wolves in Yellowstone National Park, error rates varied substantially among samples, often above the 'worst-case threshold' suggested by simulation. Consequently, a substantial proportion of multilocus genotypes held one or more errors, despite multiple PCR. These genotyping errors created several genotypes per individual and caused overestimation (up to 5.5-fold) of population size. We propose a 'matching approach' to eliminate this overestimation bias.

  7. Yellowstone wolves and the forces that structure natural systems.

    Science.gov (United States)

    Dobson, Andy P

    2014-12-01

    Since their introduction in 1995 and 1996, wolves have had effects on Yellowstone that ripple across the entire structure of the food web that defines biodiversity in the Northern Rockies ecosystem. Ecological interpretations of the wolves have generated a significant amount of debate about the relative strength of top-down versus bottom-up forces in determining herbivore and vegetation abundance in Yellowstone. Debates such as this are central to the resolution of broader debates about the role of natural enemies and climate as forces that structure food webs and modify ecosystem function. Ecologists need to significantly raise the profile of these discussions; understanding the forces that structure food webs and determine species abundance and the supply of ecosystem services is one of the central scientific questions for this century; its complexity will require new minds, new mathematics, and significant, consistent funding.

  8. Yellowstone wolves and the forces that structure natural systems.

    Directory of Open Access Journals (Sweden)

    Andy P Dobson

    2014-12-01

    Full Text Available Since their introduction in 1995 and 1996, wolves have had effects on Yellowstone that ripple across the entire structure of the food web that defines biodiversity in the Northern Rockies ecosystem. Ecological interpretations of the wolves have generated a significant amount of debate about the relative strength of top-down versus bottom-up forces in determining herbivore and vegetation abundance in Yellowstone. Debates such as this are central to the resolution of broader debates about the role of natural enemies and climate as forces that structure food webs and modify ecosystem function. Ecologists need to significantly raise the profile of these discussions; understanding the forces that structure food webs and determine species abundance and the supply of ecosystem services is one of the central scientific questions for this century; its complexity will require new minds, new mathematics, and significant, consistent funding.

  9. Seismic Evidence for Lower Mantle Plume Under the Yellowstone Hotspot

    Science.gov (United States)

    Nelson, P.; Grand, S.

    2017-12-01

    The mantle plume hypothesis for the origin of intraplate volcanism has been controversial since its inception in the 1970s. The hypothesis proposes hot narrow upwelling of rock rooted at the core mantle boundary (CMB) rise through the mantle and interact with the base of the lithosphere forming linear volcanic systems such as Hawaii and Yellowstone. Recently, broad lower mantle (>500 km in diameter) slow velocity conduits, most likely thermochemical in origin, have been associated with some intraplate volcanic provinces (French and Romanowicz, 2015). However, the direct detection of a classical thin thermal plume in the lower mantle using travel time tomography has remained elusive (Anderson and Natland, 2014). Here we present a new shear wave tomography model for the mantle beneath the western United States that is optimized to find short wavelength, sub-vertical structures in the lower mantle. Our approach uses carefully measured SKS and SKKS travel times recorded by dense North American seismic networks in conjunction with finite frequency kernels to build on existing tomography models. We find the presence of a narrow ( 300 km diameter) well isolated cylindrically shaped slow anomaly in the lower most mantle which we associate with the Yellowstone Hotspot. The conduit has a 2% reduction in shear velocity and is rooted at the CMB near the California/Arizona/Nevada border. A cross sectional view through the anomaly shows that it is slightly tilted toward the north until about 1300 km depth where it appears to weaken and deflect toward the surficial positon of the hotspot. Given the anomaly's strength, proximity to the Yellowstone Hotspot, and morphology we argue that a thermal plume interpretation is the most reasonable. Our results provide strong support for a lower mantle plume origin of the Yellowstone hotspot and more importantly the existence of deep thermal plumes.

  10. Volcanic calderas delineate biogeographic provinces among Yellowstone thermophiles.

    Science.gov (United States)

    Takacs-Vesbach, Cristina; Mitchell, Kendra; Jackson-Weaver, Olan; Reysenbach, Anna-Louise

    2008-07-01

    It has been suggested that the distribution of microorganisms should be cosmopolitan because of their enormous capacity for dispersal. However, recent studies have revealed that geographically isolated microbial populations do exist. Geographic distance as a barrier to dispersal is most often invoked to explain these distributions. Here we show that unique and diverse sequences of the bacterial genus Sulfurihydrogenibium exist in Yellowstone thermal springs, indicating that these sites are geographically isolated. Although there was no correlation with geographic distance or the associated geochemistry of the springs, there was a strong historical signal. We found that the Yellowstone calderas, remnants of prehistoric volcanic eruptions, delineate biogeographical provinces for the Sulfurihydrogenibium within Yellowstone (chi(2): 9.7, P = 0.002). The pattern of distribution that we have detected suggests that major geological events in the past 2 million years explain more of the variation in sequence diversity in this system than do contemporary factors such as habitat or geographic distance. These findings highlight the importance of historical legacies in determining contemporary microbial distributions and suggest that the same factors that determine the biogeography of macroorganisms are also evident among bacteria.

  11. Use of lodgepole pine cover types by Yellowstone grizzly bears

    Science.gov (United States)

    Mattson, D.J.

    1997-01-01

    Lodgepole pine (Pinus contorta) forests are a large and dynamic part of grizzly bear (Ursus arctos) habitat in the Yellowstone ecosystem. Research in other areas suggests that grizzly bears select for young open forest stands, especially for grazing and feeding on berries. Management guidelines accordingly recommend timber harvest as a technique for improving habitat in areas potentially dominated by lodgepole pine. In this paper I examine grizzly bear use of lodgepole pine forests in the Yellowstone area, and test several hypotheses with relevance to a new generation of management guidelines. Differences in grizzly bear selection of lodgepole pine cover types (defined on the basis of stand age and structure) were not pronounced. Selection furthermore varied among years, areas, and individuals. Positive selection for any lodgepole pine type was uncommon. Estimates of selection took 5-11 years or 4-12 adult females to stabilize, depending upon the cover type. The variances of selection estimates tended to stabilize after 3-5 sample years, and were more-or-less stable to slightly increasing with progressively increased sample area. There was no conclusive evidence that Yellowstone's grizzlies favored young (<40 yr) stands in general or for their infrequent use of berries. On the other hand, these results corroborated previous observations that grizzlies favored open and/or young stands on wet and fertile sites for grazing. These results also supported the proposition that temporally and spatially robust inferences require extensive, long-duration studies, especially for wide-ranging vertebrates like grizzly bears.

  12. Carnivore re-colonisation: Reality, possibility and a non-equilibrium century for grizzly bears in the southern Yellowstone ecosystem

    Science.gov (United States)

    Pyare, Sanjay; Cain, S.; Moody, D.; Schwartz, C.; Berger, J.

    2004-01-01

    Most large native carnivores have experienced range contractions due to conflicts with humans, although neither rates of spatial collapse nor expansion have been well characterised. In North America, the grizzly bear (Ursus arctos) once ranged from Mexico northward to Alaska, however its range in the continental USA has been reduced by 95-98%. Under the U. S. Endangered Species Act, the Yellowstone grizzly bear population has re-colonised habitats outside Yellowstone National Park. We analysed historical and current records, including data on radio-collared bears, (1) to evaluate changes in grizzly bear distribution in the southern Greater Yellowstone Ecosystem (GYE) over a 100-year period, (2) to utilise historical rates of re-colonisation to project future expansion trends and (3) to evaluate the reality of future expansion based on human limitations and land use. Analysis of distribution in 20-year increments reflects range reduction from south to north (1900-1940) and expansion to the south (1940-2000). Expansion was exponential and the area occupied by grizzly bears doubled approximately every 20 years. A complementary analysis of bear occurrence in Grand Teton National Park also suggests an unprecedented period of rapid expansion during the last 20-30 years. The grizzly bear population currently has re-occupied about 50% of the southern GYE. Based on assumptions of continued protection and ecological stasis, our model suggests total occupancy in 25 years. Alternatively, extrapolation of linear expansion rates from the period prior to protection suggests total occupancy could take > 100 years. Analyses of historical trends can be useful as a restoration tool because they enable a framework and timeline to be constructed to pre-emptively address the social challenges affecting future carnivore recovery. ?? 2004 The Zoological Society of London.

  13. Survey of selected pathogens and blood parameters of northern yellowstone elk: Wolf sanitation effect implications

    Science.gov (United States)

    Barber-Meyer, S. M.; White, P.J.; Mech, L.D.

    2007-01-01

    The restoration or conservation of predators could reduce seroprevalences of certain diseases in prey if predation selectively removes animals exhibiting clinical signs. We assessed disease seroprevalences and blood parameters of 115 adult female elk (Cervus elaphus) wintering on the northern range of Yellowstone National Park [YNP] during 2000-2005 and compared them to data collected prior to wolf (Canis lupus) restoration (WR) in 1995 and to two other herds in Montana to assess this prediction. Blood parameters were generally within two standard deviations of the means observed in other Montana herds (Gravelly-Snowcrest [GS] and Garnet Mountain [GM]), but Yellowstone elk had higher seroprevalences of parainfluenza-3 virus (95% CI YNP = 61.1-78.6, GS = 30.3-46.5) and bovine-virus-diarrhea virus type 1 (95% CI YNP = 15.9-31.9, GM = 0). In comparisons between pre-wolf restoration [pre-WR] (i.e., prior to 1995) seroprevalences with those post-wolf restoration [post-WR] in Yellowstone, we found lower seroprevalences for some disease-causing agents post-wolf restoration (e.g., bovine-virus-diarrhea virus type-1 [95% CI pre-WR = 73.1-86.3, post-WR = 15.9-31.9] and bovine-respiratory syncytial virus [95% CI pre-WR = 70.0-83.8, post-WR = 0]), but similar (e.g., Brucella abortus [95% CI pre-WR = 0-4.45, post-WR = 0-4.74] and epizootic hemorrhagic disease virus [95% CI pre-WR = 0, post-WR = 0]) or higher for others (e.g., Anaplasma marginale [95% CI pre-WR = 0, post-WR = 18.5-38.7] and Leptospira spp. [95% CI pre-WR = 0.5-6.5, post-WR = 9.5-23.5]). Though we did not detect an overall strong predation effect through reduced disease seroprevalence using retrospective comparisons with sparse data, our reference values will facilitate future assessments of this issue.

  14. State-space modeling to support management of brucellosis in the Yellowstone bison population

    Science.gov (United States)

    Hobbs, N. Thompson; Geremia, Chris; Treanor, John; Wallen, Rick; White, P.J.; Hooten, Mevin B.; Rhyan, Jack C.

    2015-01-01

    The bison (Bison bison) of the Yellowstone ecosystem, USA, exemplify the difficulty of conserving large mammals that migrate across the boundaries of conservation areas. Bison are infected with brucellosis (Brucella abortus) and their seasonal movements can expose livestock to infection. Yellowstone National Park has embarked on a program of adaptive management of bison, which requires a model that assimilates data to support management decisions. We constructed a Bayesian state-space model to reveal the influence of brucellosis on the Yellowstone bison population. A frequency-dependent model of brucellosis transmission was superior to a density-dependent model in predicting out-of-sample observations of horizontal transmission probability. A mixture model including both transmission mechanisms converged on frequency dependence. Conditional on the frequency-dependent model, brucellosis median transmission rate was 1.87 yr−1. The median of the posterior distribution of the basic reproductive ratio (R0) was 1.75. Seroprevalence of adult females varied around 60% over two decades, but only 9.6 of 100 adult females were infectious. Brucellosis depressed recruitment; estimated population growth rate λ averaged 1.07 for an infected population and 1.11 for a healthy population. We used five-year forecasting to evaluate the ability of different actions to meet management goals relative to no action. Annually removing 200 seropositive female bison increased by 30-fold the probability of reducing seroprevalence below 40% and increased by a factor of 120 the probability of achieving a 50% reduction in transmission probability relative to no action. Annually vaccinating 200 seronegative animals increased the likelihood of a 50% reduction in transmission probability by fivefold over no action. However, including uncertainty in the ability to implement management by representing stochastic variation in the number of accessible bison dramatically reduced the probability of

  15. Contrasting past and current numbers of bears visiting Yellowstone cutthroat trout streams

    Science.gov (United States)

    Haroldson, Mark A.; Schwartz, Charles C.; Teisberg, Justin E.; Gunther, Kerry A.; Fortin, Jennifer K.; Robbins, Charles T.

    2014-01-01

    Spawning cutthroat trout (Oncorhynchus clarkii bouvieri) were historically abundant within tributary streams of Yellowstone Lake within Yellowstone National Park and were a highly digestible source of energy and protein for Yellowstone’s grizzly bears (Ursus arctos) and black bears (U. americanus). The cutthroat trout population has subsequently declined since the introduction of non-native lake trout (Salvelinus namaycush), and in response to effects of drought and whirling disease (Myxobolus cerebralis). The trout population, duration of spawning runs, and indices of bear use of spawning streams had declined in some regions of the lake by 1997–2000. We initiated a 3-year study in 2007 to assess whether numbers of spawning fish, black bears, and grizzly bears within and alongside stream corridors had changed since 1997– 2000. We estimated numbers of grizzly bears and black bears by first compiling encounter histories of individual bears visiting 48 hair-snag sites along 35 historically fished streams.We analyzed DNA encounter histories with Pradel-recruitment and Jolly-Seber (POPAN) capture-mark-recapture models. When compared to 1997–2000, the current number of spawning cutthroat trout per stream and the number of streams with cutthroat trout has decreased. We estimated that 48 (95% CI¼42–56) male and 23 (95% CI¼21–27) female grizzly bears visited the historically fished tributary streams during our study. In any 1- year, 46 to 59 independent grizzly bears (8–10% of estimated Greater Yellowstone Ecosystem population) visited these streams. When compared with estimates from the 1997 to 2000 study and adjusted for equal effort, the number of grizzly bears using the stream corridors decreased by 63%. Additionally, the number of black bears decreased between 64% and 84%. We also document an increased proportion of bears of both species visiting front-country (i.e., near human development) streams. With the recovery of cutthroat trout, we suggest bears

  16. Demonstration of Decision Support Tools for Sustainable Development - An Application on Alternative Fuels in the Greater Yellowstone-Teton Region

    Energy Technology Data Exchange (ETDEWEB)

    Shropshire, D.E.; Cobb, D.A.; Worhach, P.; Jacobson, J.J.; Berrett, S.

    2000-12-30

    The Demonstration of Decision Support Tools for Sustainable Development project integrated the Bechtel/Nexant Industrial Materials Exchange Planner and the Idaho National Engineering and Environmental Laboratory System Dynamic models, demonstrating their capabilities on alternative fuel applications in the Greater Yellowstone-Teton Park system. The combined model, called the Dynamic Industrial Material Exchange, was used on selected test cases in the Greater Yellow Teton Parks region to evaluate economic, environmental, and social implications of alternative fuel applications, and identifying primary and secondary industries. The test cases included looking at compressed natural gas applications in Teton National Park and Jackson, Wyoming, and studying ethanol use in Yellowstone National Park and gateway cities in Montana. With further development, the system could be used to assist decision-makers (local government, planners, vehicle purchasers, and fuel suppliers) in selecting alternative fuels, vehicles, and developing AF infrastructures. The system could become a regional AF market assessment tool that could help decision-makers understand the behavior of the AF market and conditions in which the market would grow. Based on this high level market assessment, investors and decision-makers would become more knowledgeable of the AF market opportunity before developing detailed plans and preparing financial analysis.

  17. Post-glacial inflation-deflation cycles, tilting, and faulting in the Yellowstone Caldera based on Yellowstone Lake shorelines

    Science.gov (United States)

    Pierce, Kenneth L.; Cannon, Kenneth P.; Meyer, Grant A.; Trebesch, Matthew J.; Watts, Raymond D.

    2002-01-01

    The Yellowstone caldera, like many other later Quaternary calderas of the world, exhibits dramatic unrest. Between 1923 and 1985, the center of the Yellowstone caldera rose nearly one meter along an axis between its two resurgent domes (Pelton and Smith, 1979, Dzurisin and Yamashita, 1987). From 1985 until 1995-6, it subsided at about two cm/yr (Dzurisin and others, 1990). More recent radar interferometry studies show renewed inflation of the northeastern resurgent dome between 1995 and 1996; this inflation migrated to the southwestern resurgent dome from 1996 to 1997 (Wicks and others, 1998). We extend this record back in time using dated geomorphic evidence of postglacial Yellowstone Lake shorelines around the northern shore, and Yellowstone River levels in the outlet area. We date these shorelines using carbon isotopic and archeological methods. Following Meyer and Locke (1986) and Locke and Meyer (1994), we identify the modern shoreline as S1 (1.9 ? 0.3 m above the lake gage datum), map paleoshoreline terraces S2 to S6, and infer that the prominent shorelines were cut during intracaldera uplift episodes that produced rising water levels. Doming along the caldera axis reduces the gradient of the Yellowstone River from Le Hardys Rapids to the Yellowstone Lake outlet and ultimately causes an increase in lake level. The 1923-1985 doming is part of a longer uplift episode that has reduced the Yellowstone River gradient to a ?pool? with a drop of only 0.25 m over most of this 5 km reach. We also present new evidence that doming has caused submergence of some Holocene lake and river levels. Shoreline S5 is about 14 m above datum and estimated to be ~12.6 ka, because it post-dates a large hydrothermal explosion deposit from the Mary Bay area (MB-II) that occurred ~13 ka. S4 formed about 8 m above datum ~10.7 ka as dated by archeology and 14C, and was accompanied by offset on the Fishing Bridge fault. About 9.7 ka, the Yellowstone River eroded the ?S-meander?, followed

  18. The evolution of Yellowstone's magmatic system over the past 630 kyr: Insights from the crystal record

    Science.gov (United States)

    Stelten, M. E.

    2017-12-01

    The Yellowstone Plateau volcanic field in northwestern Wyoming is one of the world's largest, active silicic volcanic centers, and has produced three caldera-forming "super eruptions" over the past 2.1 Myr. As a result, the petrologic evolution of Yellowstone's magmatic system has been the focus of numerous studies over the past 60 years. Early studies at Yellowstone focused on characterizing whole-rock chemical and isotopic variations observed in magmas erupted over Yellowstone's lifetime. While these have provided important insights into the source of Yellowstone magmas and the processes controlling their compositional evolution though time, whole-rock studies are limited in their ability to identify the mechanisms and timescales of rhyolite generation. In contrast, much of the recent work at Yellowstone has focused on applying micro-analytical techniques to characterize the age and composition of phenocrysts hosted in Yellowstone rhyolites. These studies have greatly advanced our understanding of the magmatic system at Yellowstone and have provided crucial new insights into the mechanisms and timescales of rhyolite generation. In particular, recent work has focused on applying micro-analytical techniques to study the age and origin of the [1] three caldera-forming eruptions that produced the Huckleberry Ridge, Mesa Falls, Lava Creek tuffs and [2] post-Lava Creek tuff intracaldera rhyolites that compose the Plateau Rhyolite. As a result, a wealth of crystal-chemical data now exists for rhyolites erupted throughout Yellowstone's 2.1 Myr history. These data provide a unique opportunity to create a detailed reconstruction of Yellowstone's magmatic system through time. In this contribution, I integrate available age, chemical, and isotopic data for phenocrysts hosted in Yellowstone rhyolites to construct a model for the evolution of Yellowstone's magmatic system from the caldera-forming eruption of the Lava Creek tuff at ca. 0.63 Ma to the present day. In particular

  19. Recovering aspen follow changing elk dynamics in Yellowstone: evidence of a trophic cascade?

    Science.gov (United States)

    Painter, Luke E; Beschta, Robert L; Larsen, Eric J; Ripple, William J

    2015-01-01

    To investigate the extent and causes of recent quaking aspen (Populus tremuloides) recruitment in northern Yellowstone National Park, we measured browsing intensity and height of young aspen in 87 randomly selected aspen stands in 2012, and compared our results to similar data collected in 1997-1998. We also examined the relationship between aspen recovery and the distribution of Rocky Mountain elk (Cervus elaphus) and bison (Bison bison) on the Yellowstone northern ungulate winter range, using ungulate fecal pile densities and annual elk count data. In 1998, 90% of young aspen were browsed and none were taller-than 200 cm, the height at which aspen begin to escape from elk browsing. In 2012, only 37% in the east and 63% in the west portions of the winter range were browsed, and 65% of stands in the east had young aspen taller than 200 cm. Heights of young aspen were inversely related to browsing intensity, with the least browsing and greatest heights in the eastern portion of the range, corresponding with recent changes in elk density and distribution. In contrast with historical elk distribution (1930s-1990s), the greatest densities of elk recently (2006-2012) have been north of the park boundary (approximately 5 elk/km2), and in the western part of the range (2-4 elk/km2), with relatively few elk in the eastern portion of the range (wolves (Canis lupius) in 1995-1996 played a role in these changing elk population dynamics, interacting with other influences including increased predation by bears (Ursus spp.), competition with an expanding bison population, and shifting patterns of human land use and hunting outside the park. The resulting new aspen recruitment is evidence of a landscape-scale trophic cascade in which a resurgent large carnivore community, combined with other ecological changes, has benefited aspen through effects on ungulate prey.

  20. Are wolves saving Yellowstone's aspen? A landscape-level test of a behaviorally mediated trophic cascade.

    Science.gov (United States)

    Kauffman, Matthew J; Brodie, Jedediah F; Jules, Erik S

    2010-09-01

    Behaviorally mediated trophic cascades (BMTCs) occur when the fear of predation among herbivores enhances plant productivity. Based primarily on systems involving small-bodied predators, BMTCs have been proposed as both strong and ubiquitous in natural ecosystems. Recently, however, synthetic work has suggested that the existence of BMTCs may be mediated by predator hunting mode, whereby passive (sit-and-wait) predators have much stronger effects than active (coursing) predators. One BMTC that has been proposed for a wide-ranging active predator system involves the reintroduction of wolves (Canis lupus) to Yellowstone National Park, USA, which is thought to be leading to a recovery of trembling aspen (Populus tremuloides) by causing elk (Cervus elaphus) to avoid foraging in risky areas. Although this BMTC has been generally accepted and highly popularized, it has never been adequately tested. We assessed whether wolves influence aspen by obtaining detailed demographic data on aspen Stands using tree rings and by monitoring browsing levels in experimental elk exclosures arrayed across a gradient of predation risk for three years. Our study demonstrates that the historical failure of aspen to regenerate varied widely among stands (last recruitment year ranged from 1892 to 1956), and our data do not indicate an abrupt cessation of recruitment. This pattern of recruitment failure appears more consistent with a gradual increase in elk numbers rather than a rapid behavioral shift in elk foraging following wolf extirpation. In addition, our estimates of relative survivorship of young browsable aspen indicate that aspen are not currently recovering in Yellowstone, even in the presence of a large wolf population. Finally, in an experimental test of the BMTC hypothesis we found that the impacts of elk browsing on aspen demography are not diminished in sites where elk are at higher risk of predation by wolves. These findings suggest the need to further evaluate how trophic

  1. Cougar survival and source-sink structure on Greater Yellowstone's Northern Range

    Science.gov (United States)

    Ruth, T.K.; Haroldson, M.A.; Murphy, K.M.; Buotte, P.C.; Hornocker, M.G.; Quigley, H.B.

    2011-01-01

    We studied survival and causes of mortality of radiocollared cougars (Puma concolor) on the Greater Yellowstone Northern Range (GYNR) prior to (1987–1994) and after wolf (Canis lupus) reintroduction (1998–2005) and evaluated temporal, spatial, and environmental factors that explain variation in adult, subadult, and kitten survival. Using Program MARK and multimodel inference, we modeled cougar survival based on demographic status, season, and landscape attributes. Our best models for adult and independent subadults indicated that females survived better than males and survival increased with age until cougars reached older ages. Lower elevations and increasing density of roads, particularly in areas open to cougar hunting north of Yellowstone National Park (YNP), increased mortality risks for cougars on the GYNR. Indices of ungulate biomass, cougar and wolf population size, winter severity, rainfall, and individual characteristics such as the presence of dependent young, age class, and use of Park or Wilderness were not important predictors of survival. Kitten survival increased with age, was lower during winter, increased with increasing minimum estimates of elk calf biomass, and increased with increasing density of adult male cougars. Using our best model, we mapped adult cougar survival on the GYNR landscape. Results of receiver operating characteristic (ROC) analysis indicated a good model fit for both female (area under the curve [AUC] = 0.81, 95%CI = 0.70–0.92, n = 35 locations) and male cougars (AUC = 0.84, 95%CI = 0.74–0.94, n = 49 locations) relative to hunter harvest locations in our study area. Using minimum estimates of survival necessary to sustain the study population, we developed a source-sink surface and we identify several measures that resource management agencies can take to enhance cougar population management based on a source-sink strategy.

  2. iTRAQ analysis of hepatic proteins in free-living Mus spretus mice to assess the contamination status of areas surrounding Doñana National Park (SW Spain)

    Energy Technology Data Exchange (ETDEWEB)

    Abril, Nieves; Chicano-Gálvez, Eduardo; Michán, Carmen; Pueyo, Carmen; López-Barea, Juan, E-mail: bb1lobaj@uco.es

    2015-08-01

    This work aims to develop and integrate new -omics tools that would be applicable to different ecosystem types for a technological updating of environmental evaluations. We used a 2nd-generation (iTRAQ-8plex) proteomic approach to identify/quantify proteins differentially expressed in the liver of free-living Mus spretus mice from Doñana National Park or its proximities. Mass spectrometry was performed in an LTQ Orbitrap system for iTRAQ reporter ion quantitation and protein identification using a Mus musculus database as reference. A prior IEF step improved the separation of the complex peptide mixture. Over 2000 identified proteins were altered, of which 118 changed by ≥ 2.5-fold in mice from at least two problem sites. Part of the results obtained with the iTRAQ analysis was confirmed by Western blot. Over 75% of the 118 proteins were upregulated in animals captured at polluted sites and only 16 proteins were downregulated. Upregulated proteins were involved in stress response; cell proliferation and apoptosis; signal transduction; metastasis or tumour suppression; xenobiotic export or vesicular trafficking; and metabolism. The downregulated proteins, all potentially harmful, were classified as oncoproteins and proteins favouring genome instability. The iTRAQ results presented here demonstrated that the survival of hepatic cells is compromised in animals living at polluted sites, which showed deep alterations in metabolism and the signalling pathways. The identified proteins may be useful as biomarkers of environmental pollution and provide insight about the metabolic pathways and/or physiological processes affected by pollutants in DNP and its surrounding areas. - Highlights: • iTRAQ quantitation was used for the first time to monitor a wildlife reserve • Over 2,000 proteins with altered expression were identified in problem Doñana sites • Of them, 118 changed over 2.5-fold in, at least, two problem sites • Upregulation of protective proteins

  3. Months between rejuvenation and volcanic eruption at Yellowstone caldera, Wyoming

    Science.gov (United States)

    Till, Christy B.; Vazquez, Jorge A.; Boyce, Jeremy W

    2015-01-01

    Rejuvenation of previously intruded silicic magma is an important process leading to effusive rhyolite, which is the most common product of volcanism at calderas with protracted histories of eruption and unrest such as Yellowstone, Long Valley, and Valles, USA. Although orders of magnitude smaller in volume than rare caldera-forming super-eruptions, these relatively frequent effusions of rhyolite are comparable to the largest eruptions of the 20th century and pose a considerable volcanic hazard. However, the physical pathway from rejuvenation to eruption of silicic magma is unclear particularly because the time between reheating of a subvolcanic intrusion and eruption is poorly quantified. This study uses geospeedometry of trace element profiles with nanometer resolution in sanidine crystals to reveal that Yellowstone’s most recent volcanic cycle began when remobilization of a near- or sub-solidus silicic magma occurred less than 10 months prior to eruption, following a 220,000 year period of volcanic repose. Our results reveal a geologically rapid timescale for rejuvenation and effusion of ~3 km3 of high-silica rhyolite lava even after protracted cooling of the subvolcanic system, which is consistent with recent physical modeling that predict a timescale of several years or less. Future renewal of rhyolitic volcanism at Yellowstone is likely to require an energetic intrusion of mafic or silicic magma into the shallow subvolcanic reservoir and could rapidly generate an eruptible rhyolite on timescales similar to those documented here.

  4. Circumstances surrounding aneurysmal subarachnoid hemorrhage

    NARCIS (Netherlands)

    Schievink, W. I.; Karemaker, J. M.; Hageman, L. M.; van der Werf, D. J.

    1989-01-01

    The circumstances surrounding aneurysmal subarachnoid hemorrhage were investigated in a group of 500 consecutive patients admitted to a neurosurgical center. Subarachnoid hemorrhage occurred during stressful events in 42.8% of the patients, during nonstrenuous activities in 34.4%, and during rest or

  5. Mountain big sagebrush age distribution and relationships on the northern Yellowstone Winter Range

    Science.gov (United States)

    Carl L. Wambolt; Trista L. Hoffman

    2001-01-01

    This study was conducted within the Gardiner Basin, an especially critical wintering area for native ungulates utilizing the Northern Yellowstone Winter Range. Mountain big sagebrush plants on 33 sites were classified as large (≥22 cm canopy cover), small (

  6. The Bear Facts: Implications of Whitebark Pine Loss for Yellowstone Grizzlies

    OpenAIRE

    Willcox, Louisa

    2009-01-01

    Whitebark pine is a foundation species, and barometer of the health of high elevation forests ecosystems in the West. It provides food and cover for numerous wildlife species, including the Clark’s nutcracker, crossbill, grosbeak, red squirrel and chipmunk. Whitebark pine is particularly important in the Greater Yellowstone Ecosystem (GYE), where it provides an essential food source for the imperiled Yellowstone grizzly bear. We will review the current scientific knowledge about the relations...

  7. A field trip guide to the petrology of Quaternary volcanism on the Yellowstone Plateau

    Science.gov (United States)

    Vazquez, Jorge A.; Stelten, Mark; Bindeman, Ilya N.; Cooper, Kari

    2017-12-19

    The Yellowstone Plateau is one of the largest manifestations of silicic volcanism on Earth, and marks the youngest focus of magmatism associated with the Yellowstone Hot Spot. The earliest products of Yellowstone Hot Spot volcanism are from ~17 million years ago, but may be as old as ~32 Ma, and include contemporaneous eruption of voluminous mafic and silicic magmas, which are mostly located in the region of northwestern Nevada and southeastern Oregon. Since 17 Ma, the main locus of Yellowstone Hot Spot volcanism has migrated northeastward producing numerous silicic caldera complexes that generally remain active for ~2–4 million years, with the present-day focus being the Yellowstone Plateau. Northeastward migration of volcanism associated with the Yellowstone Hot Spot resulted in the formation of the Snake River Plain, a low relief physiographic feature extending ~750 kilometers from northern Nevada to eastern Idaho. Most of the silicic volcanic centers along the Snake River Plain have been inundated by younger basalt volcanism, but many of their ignimbrites and lava flows are exposed in the extended regions at the margins of the Snake River Plain. 

  8. Thunder on the Yellowstone revisited: An assessment of management of native ungulates by natural regulation, 1968-1993

    Science.gov (United States)

    Singer, F.J.; Swift, D.M.; Coughenour, M.B.; Varley, J.D.

    1998-01-01

    Natural regulation of native ungulates was initiated in 1968 in Yellowstone National Park (YNP) based on the premise that ungulates would reach an equilibrium with their plant resources. The natural-regulation management model stated: density dependence will regulate ungulates (i.e., a dynamic equilibrium will result between ungulates and their food supply, within some bounds of vegetation and soil effects); and no retrogression of soil and vegetation will occur from elk (Cervus elaphus) grazing during this process. The historical record indicated that elk were abundant in the system and elk were primarily food limited before settlement by European man (i.e., wolves [Canis lupus] and Native Americans were only an adjunct to the density dependent population regulation of ungulates). Density dependence was demonstrated in elk, but not in bison (Bison bison). No widespread evidence of overgrazing was observed through 1993 in study sites within vegetation communities that comprised about 97% of the winter range. No evidence of increased exotics, increased sediment yield, warming or drying of the soil, changes in soil nutrients, or differences in aboveground standing-crop biomass of plants was found between grazed and ungrazed plots. Ungulate herbivory apparently stimulated aboveground production of grasses, enhanced nitrogen and macronutrients in grasses, increased nutrient cycling, and enhanced measures of fitness in 6 common plants. However, exposed soil surface (bare ground and pebbles combined) was 11-18% greater on grazed than ungrazed plots, apparently due to a 71% decline in dead and standing litter on grazed plots. Percent live-plant basal cover, however, did not differ on grazed versus ungrazed plots, and there was no difference in soil microclimate or sediment yield. Differences in the abundance of 12% of the herbaceous species were found in grazed versus ungrazed sites (16 of 128 species); 10 were declines and 6 were increases. Willow (Salix spp.) and aspen

  9. Energy development and water options in the Yellowstone River Basin

    Energy Technology Data Exchange (ETDEWEB)

    Narayanan, R.; MacIntyre, D.D.; Torpy, M.F.

    1980-08-01

    Using a mixed-integer programming model, the impacts of institutional constraints on the marginal capacity for energy development in the Yellowstone River Basin and consequent hydrologic changes were examined. Under average annual flow conditions, energy outputs in the Yellowstone Basin can increase roughly nine times by 1985 and 12 to 18 times by 2000. In contrast, water availability is limiting energy development in the Tongue and Powder River Basins in Wyoming. Variability in hydrologic regime causes model solutions to change drastically. If flows decrease to 80 and 60% of average annual levels, the energy production is decreased by 17 and 95%, respectively. If development strategies in the basin are followed on the basis of 80% average annual flows, the Buffalo Bill enlargement (271,300 acre-ft), Tongue River Modification (58,000 acre-ft), and the two reservoirs at Sweetgrass Creek (each 27,000 acre-ft) will be necessary, in addition to several small storage facilities, to best meet the instream flow needs in Montana and to deliver the waters apportioned by compact between Wyoming and Montana. Furthermore, the results indicate that relaxing the instream flow requirements from recommended levels by 10% could increase regional energy output by 19% in 1985 and 35% in 2000. This model illustrates that modifications in institutional restrictions to achieve greater water mobility between users in a given state, as well as flexible practices for transferring water between states, can assist economic growth. Thus, the probability for restricted energy development at this juncture appears to be affected to a greater degree by institutional constraints than by water availability constraints.

  10. Advancing Site-Based Data Curation for Geobiology: The Yellowstone Exemplar (Invited)

    Science.gov (United States)

    Palmer, C. L.; Fouke, B. W.; Rodman, A.; Choudhury, G. S.

    2013-12-01

    While advances in the management and archiving of scientific digital data are proceeding apace, there is an urgent need for data curation services to collect and provide access to high-value data fit for reuse. The Site-Based Data Curation (SBDC) project is establishing a framework of guidelines and processes for the curation of research data generated at scientifically significant sites. The project is a collaboration among information scientists, geobiologists, data archiving experts, and resource managers at Yellowstone National Park (YNP). Based on our previous work with the Data Conservancy on indicators of value for research data, several factors made YNP an optimal site for developing the SBDC framework, including unique environmental conditions, a permitting process for data collection, and opportunities for geo-located longitudinal data and multiple data sources for triangulation and context. Stakeholder analysis is informing the SBDC requirements, through engagement with geologists, geochemists, and microbiologists conducting research at YNP and personnel from the Yellowstone Center for Resources and other YNP units. To date, results include data value indicators specific to site-based research, minimum and optimal parameters for data description and metadata, and a strategy for organizing data around sampling events. New value indicators identified by the scientists include ease of access to park locations for verification and correction of data, and stable environmental conditions important for controlling variables. Researchers see high potential for data aggregated from the many individual investigators conducting permitted research at YNP, however reuse is clearly contingent on detailed and consistent sampling records. Major applications of SBDC include identifying connections in dynamic systems, spatial temporal synthesis, analyzing variability within and across geological features, tracking site evolution, assessing anomalies, and greater awareness

  11. Land use planning: A potential force for retaining habitat connectivity in the Greater Yellowstone Ecosystem and Beyond

    Directory of Open Access Journals (Sweden)

    Craig L. Shafer

    2015-01-01

    Full Text Available The grizzly bear (Ursus arctos horribilis population in the Greater Yellowstone Ecosystem (GYE is perceived to have been isolated from the population in the Northern Continental Divide Ecosystem for a century. Better land use planning is needed to thwart progressive intra- and inter-ecosystem habitat fragmentation, especially due to private land development. The dilemma of private lands being intermixed in large landscapes is addressed. This review attempts to identify some land use planning levels and tools which might facilitate dispersal by the grizzly bear and other large mammals. The planning levels discussed include national, regional, state, county and municipal, and federal land management agency. Specific potential federal tools mentioned include zoning, Landscape Conservation Cooperatives, the Endangered Species Act, beyond boundary authority, land exchanges, less-than-fee acquisition and other incentives, the Northern Rockies Ecosystem Protection Act, and federal land annexation. Besides summarizing existing recommendations, some derived observations are offered.

  12. Parasite invasion following host reintroduction: a case study of Yellowstone's wolves.

    Science.gov (United States)

    Almberg, Emily S; Cross, Paul C; Dobson, Andrew P; Smith, Douglas W; Hudson, Peter J

    2012-10-19

    Wildlife reintroductions select or treat individuals for good health with the expectation that these individuals will fare better than infected animals. However, these individuals, new to their environment, may also be particularly susceptible to circulating infections and this may result in high morbidity and mortality, potentially jeopardizing the goals of recovery. Here, using the reintroduction of the grey wolf (Canis lupus) into Yellowstone National Park as a case study, we address the question of how parasites invade a reintroduced population and consider the impact of these invasions on population performance. We find that several viral parasites rapidly invaded the population inside the park, likely via spillover from resident canid species, and we contrast these with the slower invasion of sarcoptic mange, caused by the mite Sarcoptes scabiei. The spatio-temporal patterns of mange invasion were largely consistent with patterns of host connectivity and density, and we demonstrate that the area of highest resource quality, supporting the greatest density of wolves, is also the region that appears most susceptible to repeated disease invasion and parasite-induced declines. The success of wolf reintroduction appears not to have been jeopardized by infectious disease, but now shows signs of regulation or limitation modulated by parasites.

  13. Tracking hydrothermal feature changes in response to seismicity and deformation at Mud Volcano thermal area, Yellowstone

    Science.gov (United States)

    Diefenbach, A. K.; Hurwitz, S.; Murphy, F.; Evans, W.

    2013-12-01

    The Mud Volcano thermal area in Yellowstone National Park comprises many hydrothermal features including fumaroles, mudpots, springs, and thermal pools. Observations of hydrothermal changes have been made for decades in the Mud Volcano thermal area, and include reports of significant changes (the appearance of new features, increased water levels in pools, vigor of activity, and tree mortality) following an earthquake swarm in 1978 that took place beneath the area. However, no quantitative method to map and measure surface feature changes through time has been applied. We present an analysis of aerial photographs from 1954 to present to track temporal changes in the boundaries between vegetated and thermally barren areas, as well as location, extent, color, clarity, and runoff patterns of hydrothermal features within the Mud Volcano thermal area. This study attempts to provide a detailed, long-term (>50 year) inventory of hydrothermal features and change detection at Mud Volcano thermal area that can be used to identify changes in hydrothermal activity in response to seismicity, uplift and subsidence episodes of the adjacent Sour Creek resurgent dome, or other potential causes.

  14. Anatomy of Old Faithful from subsurface seismic imaging of the Yellowstone Upper Geyser Basin

    KAUST Repository

    Wu, Sin-Mei

    2017-10-03

    The Upper Geyser Basin in Yellowstone National Park contains one of the highest concentrations of hydrothermal features on Earth including the iconic Old Faithful geyser. Although this system has been the focus of many geological, geochemical, and geophysical studies for decades, the shallow (<200 m) subsurface structure remains poorly characterized. To investigate the detailed subsurface geologic structure including the hydrothermal plumbing of the Upper Geyser Basin, we deployed an array of densely spaced three-component nodal seismographs in November of 2015. In this study, we extract Rayleigh-wave seismic signals between 1-10 Hz utilizing non-diffusive seismic waves excited by nearby active hydrothermal features with the following results. 1) imaging the shallow subsurface structure by utilizing stationary hydrothermal activity as a seismic source, 2) characterizing how local geologic conditions control the formation and location of the Old Faithful hydrothermal system, and 3) resolving a relatively shallow (10-60 m) and large reservoir located ~100 m southwest of Old Faithful geyser.

  15. Opportunity's Surroundings on Sol 1818

    Science.gov (United States)

    2009-01-01

    NASA's Mars Exploration Rover Opportunity used its navigation camera to take the images combined into this full-circle view of the rover's surroundings during the 1,818th Martian day, or sol, of Opportunity's surface mission (March 5, 2009). South is at the center; north at both ends. The rover had driven 80.3 meters (263 feet) southward earlier on that sol. Tracks from the drive recede northward in this view. The terrain in this portion of Mars' Meridiani Planum region includes dark-toned sand ripples and lighter-toned bedrock. This view is presented as a cylindrical projection with geometric seam correction.

  16. Understanding the Yellowstone magmatic system using 3D geodynamic inverse models

    Science.gov (United States)

    Kaus, B. J. P.; Reuber, G. S.; Popov, A.; Baumann, T.

    2017-12-01

    The Yellowstone magmatic system is one of the largest magmatic systems on Earth. Recent seismic tomography suggest that two distinct magma chambers exist: a shallow, presumably felsic chamber and a deeper much larger, partially molten, chamber above the Moho. Why melt stalls at different depth levels above the Yellowstone plume, whereas dikes cross-cut the whole lithosphere in the nearby Snake River Plane is unclear. Partly this is caused by our incomplete understanding of lithospheric scale melt ascent processes from the upper mantle to the shallow crust, which requires better constraints on the mechanics and material properties of the lithosphere.Here, we employ lithospheric-scale 2D and 3D geodynamic models adapted to Yellowstone to better understand magmatic processes in active arcs. The models have a number of (uncertain) input parameters such as the temperature and viscosity structure of the lithosphere, geometry and melt fraction of the magmatic system, while the melt content and rock densities are obtained by consistent thermodynamic modelling of whole rock data of the Yellowstone stratigraphy. As all of these parameters affect the dynamics of the lithosphere, we use the simulations to derive testable model predictions such as gravity anomalies, surface deformation rates and lithospheric stresses and compare them with observations. We incorporated it within an inversion method and perform 3D geodynamic inverse models of the Yellowstone magmatic system. An adjoint based method is used to derive the key model parameters and the factors that affect the stress field around the Yellowstone plume, locations of enhanced diking and melt accumulations. Results suggest that the plume and the magma chambers are connected with each other and that magma chamber overpressure is required to explain the surface displacement in phases of high activity above the Yellowstone magmatic system.

  17. Opportunity's Surroundings on Sol 1687

    Science.gov (United States)

    2009-01-01

    NASA's Mars Exploration Rover Opportunity used its navigation camera to take the images combined into this 360-degree view of the rover's surroundings on the 1,687th Martian day, or sol, of its surface mission (Oct. 22, 2008). Opportunity had driven 133 meters (436 feet) that sol, crossing sand ripples up to about 10 centimeters (4 inches) tall. The tracks visible in the foreground are in the east-northeast direction. Opportunity's position on Sol 1687 was about 300 meters southwest of Victoria Crater. The rover was beginning a long trek toward a much larger crater, Endeavour, about 12 kilometers (7 miles) to the southeast. This view is presented as a cylindrical projection with geometric seam correction.

  18. Opportunity's Surroundings on Sol 1798

    Science.gov (United States)

    2009-01-01

    NASA's Mars Exploration Rover Opportunity used its navigation camera to take the images combined into this 180-degree view of the rover's surroundings during the 1,798th Martian day, or sol, of Opportunity's surface mission (Feb. 13, 2009). North is on top. The rover had driven 111 meters (364 feet) southward on the preceding sol. Tracks from that drive recede northward in this view. For scale, the distance between the parallel wheel tracks is about 1 meter (about 40 inches). The terrain in this portion of Mars' Meridiani Planum region includes dark-toned sand ripples and lighter-toned bedrock. This view is presented as a cylindrical projection with geometric seam correction.

  19. Bathymetry (Alaska and surrounding waters)

    Data.gov (United States)

    National Oceanic and Atmospheric Administration, Department of Commerce — This dataset is consists of point data taken from numerous depth surveys from the last century. These data were processed and imported into a geographic information...

  20. Binaural Rendering in MPEG Surround

    Directory of Open Access Journals (Sweden)

    Kristofer Kjörling

    2008-04-01

    Full Text Available This paper describes novel methods for evoking a multichannel audio experience over stereo headphones. In contrast to the conventional convolution-based approach where, for example, five input channels are filtered using ten head-related transfer functions, the current approach is based on a parametric representation of the multichannel signal, along with either a parametric representation of the head-related transfer functions or a reduced set of head-related transfer functions. An audio scene with multiple virtual sound sources is represented by a mono or a stereo downmix signal of all sound source signals, accompanied by certain statistical (spatial properties. These statistical properties of the sound sources are either combined with statistical properties of head-related transfer functions to estimate “binaural parameters” that represent the perceptually relevant aspects of the auditory scene or used to create a limited set of combined head-related transfer functions that can be applied directly on the downmix signal. Subsequently, a binaural rendering stage reinstates the statistical properties of the sound sources by applying the estimated binaural parameters or the reduced set of combined head-related transfer functions directly on the downmix. If combined with parametric multichannel audio coders such as MPEG Surround, the proposed methods are advantageous over conventional methods in terms of perceived quality and computational complexity.

  1. Radiofrequency fields in our surroundings

    International Nuclear Information System (INIS)

    2011-01-01

    In 2010, the National Radiation Protection Authority (NRPA) with the Post and Telecommunications Authority carried out a project where it is measured radiofrequency fields from various telecom systems in homes, kindergartens, schools, offices, and urban environments. Close to 99 percent of the measurement points were found values of less than one thousandth of the maximum. No values were near the limits. (AG)

  2. Consumption of pondweed rhizomes by Yellowstone grizzly bears

    Science.gov (United States)

    Mattson, D.J.; Podruzny, S.R.; Haroldson, M.A.

    2005-01-01

    Pondweeds (Potamogeton spp.) are common foods of waterfowl throughout the Northern Hemisphere. However, consumption of pondweeds by bears has been noted only once, in Russia. We documented consumption of pondweed rhizomes by grizzly bears (Ursus arctos) in the Yellowstone region, 1977-96, during investigations of telemetry locations obtained from 175 radiomarked bears. We documented pondweed excavations at 25 sites and detected pondweed rhizomes in 18 feces. We observed grizzly bears excavating and consuming pondweed on 2 occasions. All excavations occurred in wetlands that were inundated during and after snowmelt, but dry by late August or early September of most years. These wetlands were typified by the presence of inflated sedge (Carex vesicaria) and occurred almost exclusively on plateaus of Pliocene-Pleistocene detrital sediments or volcanic rhyolite flows. Bears excavated wetlands with pondweeds when they were free of standing water, most commonly during October and occasionally during spring prior to the onset of terminal snowmelt. Most excavations were about 4.5 cm deep, 40 cubic decimeter (dm3) in total volume, and targeted the thickened pondweed rhizomes. Starch content of rhizomes collected near grizzly bear excavations averaged 28% (12% SD; n = 6). These results add to the documented diversity of grizzly bear food habits and, because pondweed is distributed circumboreally, also raise the possibility that consumption of pondweed by grizzly bears has been overlooked in other regions.

  3. 78 FR 22470 - Special Regulations; Areas of the National Park System; Yellowstone National Park; Winter Use

    Science.gov (United States)

    2013-04-16

    ..., natural soundscapes, visitor use and experience, and park operations. Impacts associated with each of the... monitoring, including data regarding air quality, wildlife, soundscapes, and health and safety, were used in... impacts to wildlife, air quality, natural soundscapes, and visitor and employee safety, the NPS is...

  4. 76 FR 39048 - Special Regulations; Areas of the National Park System, Yellowstone National Park

    Science.gov (United States)

    2011-07-05

    ..., natural soundscapes, visitor use and experience, and visitor accessibility. Impacts associated with each... oversnow vehicles on the park's soundscapes. NPS Approved Snowmobiles and Snowcoaches The Superintendent..., air quality, natural soundscapes, and visitor and employee safety, the NPS is proposing to continue...

  5. 78 FR 63069 - Special Regulations; Areas of the National Park System; Yellowstone National Park; Winter Use

    Science.gov (United States)

    2013-10-23

    ... soundscapes, visitor use and experience, and park operations. Impacts associated with each of the alternatives..., soundscapes, and health and safety, were used in formulating the alternatives in the Plan/SEIS. Applies the... To mitigate impacts to wildlife, air quality, natural soundscapes, and visitor and employee safety...

  6. 77 FR 73919 - Special Regulations; Areas of the National Park System, Yellowstone National Park

    Science.gov (United States)

    2012-12-12

    ... effect on the economy of $100 million or more. (b) Will not cause a major increase in costs or prices for... contain any new collection of information that requires approval by the Office of Management and Budget... reserving time off from work, booking airfares and hotel accommodations, making reservations for snowmobile...

  7. 77 FR 53826 - Special Regulations; Areas of the National Park System, Yellowstone National Park

    Science.gov (United States)

    2012-09-04

    ... effect on the economy of $100 million or more. (b) Will not cause a major increase in costs or prices for... approval by the Office of Management and Budget (OMB) under the PRA of 1995 (44 U.S.C. 3501 et seq.). OMB... reserving time off from work, booking airfares and hotel accommodations, making reservations for snowmobile...

  8. Songbird response to increased willow (Salix spp.) growth in Yellowstone's northern range.

    Science.gov (United States)

    Baril, Lisa M; Hansen, Andrew J; Renkin, Roy; Lawrence, Rick

    2011-09-01

    After nearly a century of height suppression, willows (Salix spp.) in the northern range of Yellowstone National Park, U.S.A., are increasing in height growth as a possible consequence of wolf (Canis lupus) restoration, climate change, or other factors. Regardless of the drivers, the recent release of this rare but important habitat type could have significant implications for associated songbirds that are exhibiting declines in the region. Our objective was to evaluate bird response to releasing willows by comparing willow structure and bird community composition across three willow growth conditions: height suppressed, recently released, and previously tall (i.e., tall prior to the height increase of released willows). Released and previously tall willows exhibited high and similar vertical structure, but released willows were significantly lower in horizontal structure. Suppressed willows were significantly shorter and lower in horizontal cover than released or previously tall willows. Bird richness increased along a gradient from lowest in suppressed to highest in previously tall willows, but abundance and diversity were similar between released and previously tall willows, despite lower horizontal cover in the released condition. Common Yellowthroat (Geothlypis trichas) and Lincoln's Sparrow (Melospiza lincolnii) were found in all three growth conditions; however, Yellow Warbler (Dendroica petechia), Warbling Vireo (Vireo gilvus), Willow Flycatcher (Empidonax traillii), and Song Sparrow (Melospiza melodii) were present in released and previously tall willows only. Wilson's Warbler (Wilsonia pusilla) was found in previously tall willows only, appearing to specialize on tall, dense willows. The results of our a priori habitat models indicated that foliage height diversity was the primary driver of bird richness, abundance, and diversity. These results indicate that vertical structure was a more important driver of bird community variables than horizontal

  9. 3 CFR 8362 - Proclamation 8362 of April 17, 2009. National Park Week, 2009

    Science.gov (United States)

    2010-01-01

    ... occurred over the long course of our history. Our system of National Parks is entrusted to each generation... Lincoln Memorial and Ellis Island to the Great Smoky Mountains and Yellowstone, National Parks attract... shared history. From the Prehistoric Trackways National Monument to the Civil War battlefield at...

  10. Heat flow in vapor dominated areas of the Yellowstone Plateau volcanic field: implications for the thermal budget of the Yellowstone Caldera

    Science.gov (United States)

    Hurwitz, Shaul; Harris, Robert; Werner, Cynthia Anne; Murphy, Fred

    2012-01-01

    Characterizing the vigor of magmatic activity in Yellowstone requires knowledge of the mechanisms and rates of heat transport between magma and the ground surface. We present results from a heat flow study in two vapor dominated, acid-sulfate thermal areas in the Yellowstone Caldera, the 0.11 km2 Obsidian Pool Thermal Area (OPTA) and the 0.25 km2 Solfatara Plateau Thermal Area (SPTA). Conductive heat flux through a low permeability layer capping large vapor reservoirs is calculated from soil temperature measurements at >600 locations and from laboratory measurements of soil properties. The conductive heat output is 3.6 ± 0.4 MW and 7.5 ± 0.4 MW from the OPTA and the SPTA, respectively. The advective heat output from soils is 1.3 ± 0.3 MW and 1.2 ± 0.3 MW from the OPTA and the SPTA, respectively and the heat output from thermal pools in the OPTA is 6.8 ± 1.4 MW. These estimates result in a total heat output of 11.8 ± 1.4 MW and 8.8 ± 0.4 MW from OPTA and SPTA, respectively. Focused zones of high heat flux in both thermal areas are roughly aligned with regional faults suggesting that faults in both areas serve as conduits for the rising acid vapor. Extrapolation of the average heat flux from the OPTA (103 ± 2 W·m−2) and SPTA (35 ± 3 W·m−2) to the ~35 km2 of vapor dominated areas in Yellowstone yields 3.6 and 1.2 GW, respectively, which is less than the total heat output transported by steam from the Yellowstone Caldera as estimated by the chloride inventory method (4.0 to 8.0 GW).

  11. Dietary breadth of grizzly bears in the Greater Yellowstone Ecosystem

    Science.gov (United States)

    Gunther, Kerry A.; Shoemaker, Rebecca; Frey, Kevin L.; Haroldson, Mark A.; Cain, Steven L.; van Manen, Frank T.; Fortin, Jennifer K.

    2014-01-01

    Grizzly bears (Ursus arctos) in the Greater Yellowstone Ecosystem (GYE) are opportunistic omnivores that eat a great diversity of plant and animal species. Changes in climate may affect regional vegetation, hydrology, insects, and fire regimes, likely influencing the abundance, range, and elevational distribution of the plants and animals consumed by GYE grizzly bears. Determining the dietary breadth of grizzly bears is important to document future changes in food resources and how those changes may affect the nutritional ecology of grizzlies. However, no synthesis exists of all foods consumed by grizzly bears in the GYE. We conducted a review of available literature and compiled a list of species consumed by grizzly bears in the GYE. We documented >266 species within 200 genera from 4 kingdoms, including 175 plant, 37 invertebrate, 34 mammal, 7 fungi, 7 bird, 4 fish, 1 amphibian, and 1 algae species as well as 1 soil type consumed by grizzly bears. The average energy values of the ungulates (6.8 kcal/g), trout (Oncorhynchus spp., 6.1 kcal/g), and small mammals (4.5 kcal/g) eaten by grizzlies were higher than those of the plants (3.0 kcal/g) and invertebrates (2.7 kcal/g) they consumed. The most frequently detected diet items were graminoids, ants (Formicidae), whitebark pine seeds (Pinus albicaulis), clover (Trifolium spp.), and dandelion (Taraxacum spp.). The most consistently used foods on a temporal basis were graminoids, ants, whitebark pine seeds, clover, elk (Cervus elaphus), thistle (Cirsium spp.), and horsetail (Equisetum spp.). Historically, garbage was a significant diet item for grizzlies until refuse dumps were closed. Use of forbs increased after garbage was no longer readily available. The list of foods we compiled will help managers of grizzly bears and their habitat document future changes in grizzly bear food habits and how bears respond to changing food resources.

  12. Leaders' perspectives in the Yellowstone to Yukon Conservation Initiative

    Science.gov (United States)

    Mattson, D.J.; Clark, S.G.; Byrd, K.L.; Brown, S.R.; Robinson, B.

    2011-01-01

    The Yellowstone to Yukon Conservation Initiative (Y2Y) was created in 1993 to advance conservation in a 1.2 million km2 portion of the North American Rocky Mountains. We assembled 21 people with influence over Y2Y in a workshop to elucidate perspectives on challenges and solutions for this organization at a key point in its evolution, and used Q method to define four perspectives on challenges and three on solutions. Participants were differentiated by four models for effecting change-vision-based advocacy, practice-based learning, political engagement, and scientific management-with emphasis on the first three. Those with authority in Y2Y aligned with vision-based advocacy and expressed ambivalence about practice-based adaptive learning and rigorous appraisals of existing strategies. Workshop results were consistent with an apparent trend toward organizational maturation focused on stabilizing revenues, developing formal organizational arrangements, and focusing strategies. Consolidation of power in Y2Y around a long-standing formula does not bode well for the effectiveness of Y2Y. We recommend that leaders in Y2Y and similar organizations focused on large-scale conservation to create and maintain an open system-philosophically and operationally-that capitalizes on the diverse perspectives and skills of individuals who are attracted to such efforts. We also recommend that the Y2Y initiative be followed closely to harvest additional lessons for potential application to large-scale conservation efforts elsewhere. ?? Springer Science+Business Media, LLC(outside the USA).2011.

  13. Low-δD hydration rinds in Yellowstone perlites record rapid syneruptive hydration during glacial and interglacial conditions

    Science.gov (United States)

    Bindeman, Ilya N.; Lowenstern, Jacob B.

    2016-01-01

    Hydration of silicic volcanic glass forms perlite, a dusky, porous form of altered glass characterized by abundant “onion-skin” fractures. The timing and temperature of perlite formation are enigmatic and could plausibly occur during eruption, during post-eruptive cooling, or much later at ambient temperatures. To learn more about the origin of natural perlite, and to fingerprint the hydration waters, we investigated perlitic glass from several synglacial and interglacial rhyolitic lavas and tuffs from the Yellowstone volcanic system. Perlitic cores are surrounded by a series of conchoidal cracks that separate 30- to 100-µm-thick slivers, likely formed in response to hydration-induced stress. H2O and D/H profiles confirm that most D/H exchange happens together with rapid H2O addition but some smoother D/H variations may suggest separate minor exchange by deuterium atom interdiffusion following hydration. The hydrated rinds (2–3 wt% H2O) transition rapidly (within 30 µm, or by 1 wt% H2O per 10 µm) to unhydrated glass cores. This is consistent with quenched “hydration fronts” where H2O diffusion coefficients are strongly dependent on H2O concentrations. The chemical, δ18O, and δD systematics of bulk glass records last equilibrium between ~110 and 60 °C without chemical exchange but with some δ18O exchange. Similarly, the δ18O of water extracted from glass by rapid heating suggests that water was added to the glass during cooling at higher rates of diffusion at 60–110 °C temperatures, compared with values expected from extrapolation of high-temperature (>400 °C) experimental data. The thick hydration rinds in perlites, measuring hundreds of microns, preserve the original D/H values of hydrating water as a recorder of paleoclimate conditions. Measured δD values in perlitic lavas are −150 to −191 or 20–40 ‰ lower than glass hydrated by modern Yellowstone waters. This suggests that Yellowstone perlites record the low-δD signature

  14. The climate adaptation programs and activities of the Yellowstone to Yukon Conservation Initiative

    Science.gov (United States)

    Wendy L. Francis

    2011-01-01

    The Yellowstone to Yukon Conservation Initiative (Y2Y) is an innovative transboundary effort to protect biodiversity and facilitate climate adaptation by linking large protected core areas through compatible land uses on matrix lands. The Y2Y organization acts as the keeper of the Y2Y vision and implements two interconnected programs - Science and Action, and Vision...

  15. Climate influences on whitebark pine mortality from mountain pine beetle in the Greater Yellowstone Ecosystem

    Science.gov (United States)

    Polly C. Buotte; Jeffrey A. Hicke; Haiganoush K. Preisler; John T. Abatzoglou; Kenneth F. Raffa; Jesse A. Logan

    2016-01-01

    Extensive mortality of whitebark pine, beginning in the early to mid-2000s, occurred in the Greater Yellowstone Ecosystem (GYE) of the western USA, primarily from mountain pine beetle but also from other threats such as white pine blister rust. The climatic drivers of this recent mortality and the potential for future whitebark pine mortality from mountain pine beetle...

  16. Monitoring white pine blister rust infection and mortality in whitebark pine in the Greater Yellowstone ecosystem

    Science.gov (United States)

    Cathie Jean; Erin Shanahan; Rob Daley; Gregg DeNitto; Dan Reinhart; Chuck Schwartz

    2011-01-01

    There is a critical need for information on the status and trend of whitebark pine (Pinus albicaulis) in the Greater Yellowstone Ecosystem (GYE). Concerns over the combined effects of white pine blister rust (WPBR, Cronartium ribicola), mountain pine beetle (MPB, Dendroctonus ponderosae), and climate change prompted an interagency working group to design and implement...

  17. Performance of Yellowstone and Snake River Cutthroat Trout Fry Fed Seven Different Diets.

    Science.gov (United States)

    Five commercial diets and two formulated feeds were fed to initial-feeding Yellowstone cutthroat trout Oncorhynchus clarkii bouvieri fry and Snake River cutthroat trout O. clarkii spp. (currently being petitioned for classification as O. clarkii behnkei) fry for 18 weeks to evaluate fish performance...

  18. Lower-mantle plume beneath the Yellowstone hotspot revealed by core waves

    Science.gov (United States)

    Nelson, Peter L.; Grand, Stephen P.

    2018-04-01

    The Yellowstone hotspot, located in North America, is an intraplate source of magmatism the cause of which is hotly debated. Some argue that a deep mantle plume sourced at the base of the mantle supplies the heat beneath Yellowstone, whereas others claim shallower subduction or lithospheric-related processes can explain the anomalous magmatism. Here we present a shear wave tomography model for the deep mantle beneath the western United States that was made using the travel times of core waves recorded by the dense USArray seismic network. The model reveals a single narrow, cylindrically shaped slow anomaly, approximately 350 km in diameter that we interpret as a whole-mantle plume. The anomaly is tilted to the northeast and extends from the core-mantle boundary to the surficial position of the Yellowstone hotspot. The structure gradually decreases in strength from the deepest mantle towards the surface and if it is purely a thermal anomaly this implies an initial excess temperature of 650 to 850 °C. Our results strongly support a deep origin for the Yellowstone hotspot, and also provide evidence for the existence of thin thermal mantle plumes that are currently beyond the resolution of global tomography models.

  19. Continued warming could transform Greater Yellowstone fire regimes by mid-21st century

    Science.gov (United States)

    Anthony L. Westerling; Monica G. Turner; Erica A. H. Smithwick; William H. Romme; Michael G. Ryan

    2011-01-01

    Climate change is likely to alter wildfire regimes, but the magnitude and timing of potential climate-driven changes in regional fire regimes are not well understood. We considered how the occurrence, size, and spatial location of large fires might respond to climate projections in the Greater Yellowstone ecosystem (GYE) (Wyoming), a large wildland ecosystem dominated...

  20. Attributes of Yellowstone cutthroat trout redds in a tributary of the Snake River, Idaho

    Science.gov (United States)

    Russell F. Thurow; John G. King

    1994-01-01

    We characterized spawning sites of Yellowstone cutthroat trout Oncorhynchus clarki bouvieri, described the microhabitat of completed redds, and tested the influence of habitat conditions on the morphology of completed redds in Pine Creek, Idaho. Cutthroat trout spawned in June as flows subsided after peak stream discharge. During spawning, minimum and maximum water...

  1. Twenty Years After the 1988 Yellowstone Fires: Lessons About Disturbance and Ecosystems

    Science.gov (United States)

    Romme, W.H.; Boyce, M.S.; Gresswell, R.; Merrill, E.H.; Minshall, G.W.; Whitlock, C.; Turner, M.G.

    2011-01-01

    The 1988 Yellowstone fires were among the first in what has proven to be an upsurge in large severe fires in the western USA during the past 20 years. At the time of the fires, little was known about the impacts of such a large severe disturbance because scientists had had few previous opportunities to study such an event. Ecologists predicted short- and long-term effects of the 1988 fires on vegetation, biogeochemistry, primary productivity, wildlife, and aquatic ecosystems based on scientific understanding of the time. Twenty-plus years of subsequent study allow these early predictions to be evaluated. Most of the original predictions were at least partially supported, but some predictions were refuted, others nuanced, and a few postfire phenomena were entirely unexpected. Post-1988 Yellowstone studies catalyzed advances in ecology focused on the importance of spatial and temporal heterogeneity, contingent influences, and multiple interacting drivers. Post-1988 research in Yellowstone also has changed public perceptions of fire as an ecological process and attitudes towards fire management. Looking ahead to projected climate change and more frequent large fires, the well-documented ecological responses to the 1988 Yellowstone fires provide a foundation for detecting and evaluating potential changes in fire regimes of temperate mountainous regions. ?? 2011 Springer Science+Business Media, LLC.

  2. Low rates of antimicrobial-resistant Enterobacteriaceae in wildlife in Taï National Park, Côte d'Ivoire, surrounded by villages with high prevalence of multiresistant ESBL-producing Escherichia coli in people and domestic animals.

    Science.gov (United States)

    Albrechtova, Katerina; Papousek, Ivo; De Nys, Helene; Pauly, Maude; Anoh, Etile; Mossoun, Arsene; Dolejska, Monika; Masarikova, Martina; Metzger, Sonya; Couacy-Hymann, Emmanuel; Akoua-Koffi, Chantal; Wittig, Roman M; Klimes, Jiri; Cizek, Alois; Leendertz, Fabian H; Literak, Ivan

    2014-01-01

    Antimicrobial resistance genes can be found in all ecosystems, including those where antibiotic selective pressure has never been exerted. We investigated resistance genes in a collection of faecal samples of wildlife (non-human primates, mice), people and domestic animals (dogs, cats) in Côte d'Ivoire; in the chimpanzee research area of Taï National Park (TNP) and adjacent villages. Single bacteria isolates were collected from antibiotic-containing agar plates and subjected to molecular analysis to detect Enterobacteriaceae isolates with plasmid-mediated genes of extended-spectrum beta-lactamases (ESBLs) and plasmid-mediated quinolone resistance (PMQR). While the prevalence of ESBL-producing E. coli in the villages was 27% in people (n = 77) and 32% in dogs (n = 38), no ESBL-producer was found in wildlife of TNP (n = 75). PMQR genes, mainly represented by qnrS1, were also present in human- and dog-originating isolates from the villages (36% and 42% in people and dogs, respectively), but no qnrS has been found in the park. In TNP, different variants of qnrB were detected in Citrobacter freundii isolates originating non-human primates and mice. In conclusion, ESBL and PMQR genes frequently found in humans and domestic animals in the villages were rather exceptional in wildlife living in the protected area. Although people enter the park, the strict biosecurity levels they are obliged to follow probably impede transmission of bacteria between them and wildlife.

  3. Survival of adult female elk in yellowstone following wolf restoration

    Science.gov (United States)

    Evans, S.B.; Mech, L.D.; White, P.J.; Sargeant, G.A.

    2006-01-01

    Counts of northern Yellowstone elk (Cervus elaphus) in northwestern Wyoming and adjacent Montana, USA, have decreased at an average rate of 6-8% per year since wolves (Canis lupus) were reintroduced in 1995. Population growth rates of elk are typically sensitive to variations in adult female survival; populations that are stable or increasing exhibit high adult female survival. We used survival records for 85 radiocollared adult female elk 1-19 years old to estimate annual survival from March 2000 to February 2004. Weighted average annual survival rates were approximately 0.83 (95% CI = 0.77-0.89) for females 1-15 years old and 0.80 (95% CI = 0.73-0.86) for all females. Our estimates were much lower than the rate of 0.99 observed during 1969-1975 when fewer elk were harvested by hunters, wolves were not present, and other predators were less numerous. Of 33 documented deaths included in our analysis, we attributed 11 to hunter harvest, 14 to predation (10 wolf, 2 unknown, 1 cougar [Puma concolor], and 1 bear [Ursus sp.]), 6 to unknown causes, and 2 to winter-kill. Most deaths occurred from December through March. Estimates of cause-specific annual mortality rates were 0.09 (0.05-0.14) for all predators, 0.08 (0.04-0.13) for hunting, and 0.07 (0.03-0.11) for wolves specifically. Wolf-killed elk were typically older (median = 12 yr) than hunter-killed elk (median = 9 yr, P = 0.03). However, elk that winter outside the park where they were exposed to hunting were also younger (median = 7 yr) than elk that we did not observe outside the park (median = 9 yr, P wolves and hunters may reflect characteristics of elk exposed to various causes of mortality, as well as differences in susceptibility. Unless survival rates of adult females increase, elk numbers are likely to continue declining. Hunter harvest is the only cause of mortality that is amenable to management at the present time.

  4. Comunidade de aves de sub-bosque em uma área de entorno do Parque Nacional do Itatiaia, Rio de Janeiro, Brasil Understory bird community in a surrounded area of Itatiaia National Park, Rio de Janeiro, Brazil

    Directory of Open Access Journals (Sweden)

    Eliana R. Maia-Gouvêa

    2005-12-01

    Full Text Available Este trabalho foi realizado em mata secundária no Município de Itatiaia, Estado do Rio de Janeiro (22º30'S e 44º30'W próximo ao Parque Nacional do Itatiaia, com o objeitvo de descrever a comunidade de aves ali presente. Foram amostrados três tipos vegetacionais distintos: reflorestamento, bosque e pomar, tendo sido realizadas 19 excursões entre 1984 e 1999. As aves foram amostradas através de capturas com redes de neblina e anilhamento, tendo sido utilizadas de 15 a 31 redes de 12 x 2 m e malha 36 mm. Foram também obtidas medidas morfométricas (comprimento total, asa, bico, cauda e tarso e dados biológicos (sexo, idade reprodução e muda. Depois de 5.621,79 horas-rede, foi registrado um total de 553 capturas, com 71 recapturas (12,84%; 417 indivíduos foram anilhados, e 65 beija-flores deixaram de ser marcados por falta de anilhas específicas. A comunidade estudada esteve representada por 77 espécies e 18 famílias, apresentando índice de diversidade H' = -1,594 e a curva do coletor com tendência à estabilização. As famílias com maior número de espécies foram Emberizidae (n = 21; 27,27% e Tyrannidae (n = 15; 19,48%. As espécies com maior abundância relativa foram Turdus leucomelas (n = 40; 9,59% e Turdus rufiventris (n = 36; 8,63%. Seis das espécies amostradas (7,8% são endêmicas do bioma Mata Atlântica. Na estação chuvosa foram amostradas 68 espécies, e na estação seca, 42; e as capturas estiveram relacionadas com as chuvas (rs = -0,6778; p = 0,05. O período reprodutivo ocorreu de outubro a março estando correlacionado com o início da estação chuvosa (rs = -0,702; p = 0,052.This study was conduced in a second growth woodland close to Itatiaia National Park (22º30'S e 44º30'W, Rio de Janeiro state, Brazil, and aimed to describe the understory bird community living in this area. We sampled three different vegetation types, reforestation, wood and orchard, through 19 field trips between 1984 and 1999. Birds

  5. Projecting the spatiotemporal carbon dynamics of the Greater Yellowstone Ecosystem from 2006 to 2050.

    Science.gov (United States)

    Huang, Shengli; Liu, Shuguang; Liu, Jinxun; Dahal, Devendra; Young, Claudia; Davis, Brian; Sohl, Terry L; Hawbaker, Todd J; Sleeter, Ben; Zhu, Zhiliang

    2015-12-01

    Climate change and the concurrent change in wildfire events and land use comprehensively affect carbon dynamics in both spatial and temporal dimensions. The purpose of this study was to project the spatial and temporal aspects of carbon storage in the Greater Yellowstone Ecosystem (GYE) under these changes from 2006 to 2050. We selected three emission scenarios and produced simulations with the CENTURY model using three General Circulation Models (GCMs) for each scenario. We also incorporated projected land use change and fire occurrence into the carbon accounting. The three GCMs showed increases in maximum and minimum temperature, but precipitation projections varied among GCMs. Total ecosystem carbon increased steadily from 7,942 gC/m 2 in 2006 to 10,234 gC/m 2 in 2050 with an annual rate increase of 53 gC/m 2 /year. About 56.6% and 27% of the increasing rate was attributed to total live carbon and total soil carbon, respectively. Net Primary Production (NPP) increased slightly from 260 gC/m 2 /year in 2006 to 310 gC/m 2 /year in 2050 with an annual rate increase of 1.22 gC/m 2 /year. Forest clear-cutting and fires resulted in direct carbon removal; however, the rate was low at 2.44 gC/m 2 /year during 2006-2050. The area of clear-cutting and wildfires in the GYE would account for 10.87% of total forested area during 2006-2050, but the predictive simulations demonstrated different spatial distributions in national forests and national parks. The GYE is a carbon sink during 2006-2050. The capability of vegetation is almost double that of soil in terms of sequestering extra carbon. Clear-cutting and wildfires in GYE will affect 10.87% of total forested area, but direct carbon removal from clear-cutting and fires is 109.6 gC/m 2 , which accounts for only 1.2% of the mean ecosystem carbon level of 9,056 gC/m 2 , and thus is not significant.

  6. Seismic and GPS constraints on the dynamics and kinematics of the Yellowstone volcanic field

    Science.gov (United States)

    Smith, R. B.; Farrell, J.; Jordan, M.; Puskas, C.; Waite, G. P.

    2007-12-01

    The seismically and volcanically Yellowstone hotspot resulted from interaction of a mantle plume with the overriding North America plate. This feature and related processes have modified continental lithosphere producing the Yellowstone-Snake River Plain-Newberry silicic volcanic field (YSRPN) system, with its NE volcanically active Yellowstone volcanic field. The size and accessibility of the Yellowstone area has allowed a range of geophysical experiments including earthquake monitoring and seismic and GPS imaging of this system. Seismicity is dominated by small-magnitude normal- to oblique-slip faulting earthquake swarms with shallow focal depths, maximum of ~5 km, restricted by high temperatures and a weak elastic layer. There is developing evidence of non-double couple events. Outside the caldera, earthquakes are deeper, ~20 km, and capable of M 7+ earthquakes. We integrate the results from a multi-institution experiment that recorded data from 110 seismic stations and 180 GPS stations for 1999-2004. The tomographic images confirm the existence of a low Vp-body beneath the Yellowstone caldera at depths greater than 8 km, possibly representing hot, crystallizing magma. A key result of our study is a volume of anomalously low Vp and Vp/Vs in the northwestern part of the volcanic field at shallow depths of stress field inverted from seismic and GPS data is dominated by regional SW extension with superimposed volumetric expansion and uplift from local volcanic sources. Mantle tomography derived from integrated inversion of teleseismic and local earthquake data constrained by geoid, crustal structure, discontinuity structure reveals an upper-mantle low P and S velocity body extends from 80 km to ~250 km directly beneath Yellowstone and then continues to 650 km with unexpected westward tilt to the west at ~60° with a 1% to 2% melt. This geometry is consistent with the ascent of the buoyant magma entrained in eastward return-flow of the upper mantle. Some remaining

  7. 78 FR 77709 - Notice of Extension of Concession Contracts

    Science.gov (United States)

    2013-12-24

    ... Jackson Hole Llamas.... Yellowstone National Park. YELL107-04 Wyoming Backcountry Yellowstone National... Creek Yellowstone National Outfitters, Inc. Park. YELL121-04 Yellowstone Llamas..... Yellowstone...... Yellowstone National Park. YELL168-04 Llama Trips in Yellowstone National Yellowstone. Park. YELL170-04 Rockin...

  8. Dual stable isotopes of CH 4 from Yellowstone hot-springs suggest hydrothermal processes involving magmatic CO 2

    Energy Technology Data Exchange (ETDEWEB)

    Moran, James J.; Whitmore, Laura M.; Jay, Zackary J.; Jennings, Ryan deM.; Beam, Jacob P.; Kreuzer, Helen W.; Inskeep, William P.

    2017-07-01

    Volcanism and post-magmatism contribute both significant annual CH4 fluxes to the atmosphere (on par with other natural sources such as forest fire and wild animal emissions) and have been implicated in past climate-change events. The Yellowstone hot spot is one of the largest volcanic systems on Earth and is known to emit methane in addition to other greenhouse gases (e.g. carbon dioxide) but the ultimate source of this methane flux has not been elucidated. Here we use dual stable isotope analysis (δ2H and δ13C) of CH4(g) sampled from ten high-temperature geothermal pools in Yellowstone National Park to show that the predominant flux of CH4(g) is abiotic. The average δ13C and δ2H values of CH4(g) emitted from hot springs (-26.7 (±2.4) and -236.9 (±12.0) ‰, respectively) are not consistent with biotic (microbial or thermogenic) methane sources, but are within previously reported ranges for abiotic methane production. Correlation between δ13CCH4 and δ13C-dissolved inorganic C (DIC) also suggests that CO2 is a parent C source for the observed CH4(g). Moreover, CH4-CO2 isotopic geothermometry was used to estimate CH4(g) formation temperatures ranging from ~ 250 - 350°C, which is just below the temperature estimated for the hydrothermal reservoir and consistent with the hypothesis that subsurface, rock-water interactions are responsible for large methane fluxes from this volcanic system. An understanding of conditions leading to the abiotic production of methane and associated isotopic signatures are central to understanding the evolutionary history of deep carbon sources on Earth.

  9. History of surface displacements at the Yellowstone Caldera, Wyoming, from leveling surveys and InSAR observations, 1923-2008

    Science.gov (United States)

    Dzurisin, Daniel; Wicks, Charles W.; Poland, Michael P.

    2012-01-01

    Modern geodetic studies of the Yellowstone caldera, Wyoming, and its extraordinary tectonic, magmatic, and hydrothermal systems date from an initial leveling survey done throughout Yellowstone National Park in 1923 by the U.S. Coast and Geodetic Survey. A repeat park-wide survey by the U.S. Geological Survey (USGS) and the University of Utah during 1975-77 revealed that the central part of the caldera floor had risen more than 700 mm since 1923, at an average rate of 14±1 mm/yr. From 1983 to 2007, the USGS conducted 15 smaller surveys of a single level line that crosses the northeast part of the caldera, including the area where the greatest uplift had occurred from 1923 to 1975-77. The 1983 and 1984 surveys showed that uplift had continued at an average rate of 22±1 mm/yr since 1975-77, but no additional uplift occurred during 1984-85 (-2±5 mm/yr), and during 1985-95 the area subsided at an average rate of 19±1 mm/yr. The change from uplift to subsidence was accompanied by an earthquake swarm, the largest ever recorded in the Yellowstone area (as of March 2012), starting in October 1985 and located near the northwest rim of the caldera. Interferometric synthetic aperture radar (InSAR) images showed that the area of greatest subsidence migrated from the northeast part of the caldera (including the Sour Creek resurgent dome) during 1992-93 to the southwest part (including the Mallard Lake resurgent dome) during 1993-95. Thereafter, uplift resumed in the northeast part of the caldera during 1995-96, while subsidence continued in the southwest part. The onset of uplift migrated southwestward, and by mid-1997, uplift was occurring throughout the entire caldera (essentially rim to rim, including both domes). Consistent with these InSAR observations, leveling surveys indicated 24±3 mm of uplift in the northeast part of the caldera during 1995-98. The beginning of uplift was coincident with or followed shortly after an earthquake swarm near the north caldera rim

  10. Religion's relationship with social boundaries surrounding gender ...

    African Journals Online (AJOL)

    Religion's relationship with social boundaries surrounding gender. ... is associated with segregation, marginalization and differentiation between men and women. ... are necessary in the society it should not be mistaken for gender inequality.

  11. Magnetotelluric Investigations of the Yellowstone Caldera: Understanding the Emplacement of Crustal Magma Bodies

    Science.gov (United States)

    Gurrola, R. M.; Neal, B. A.; Bennington, N. L.; Cronin, R.; Fry, B.; Hart, L.; Imamura, N.; Kelbert, A.; Bowles-martinez, E.; Miller, D. J.; Scholz, K. J.; Schultz, A.

    2017-12-01

    Wideband magnetotellurics (MT) presents an ideal method for imaging conductive shallow magma bodies associated with contemporary Yellowstone-Snake River Plain (YSRP) magmatism. Particularly, how do these magma bodies accumulate in the mid to upper crust underlying the Yellowstone Caldera, and furthermore, what role do hydrothermal fluids play in their ascent? During the summer 2017 field season, two field teams from Oregon State University and the University of Wisconsin-Madison installed forty-four wideband MT stations within and around the caldera, and using data slated for joint 3-D inversion with existing seismic data, two 2-D vertical conductivity sections of the crust and upper mantle were constructed. These models, in turn, provide preliminary insight into the emplacement of crustal magma bodies and hydrothermal processes in the YSRP region.

  12. Grizzly bear use of army cutworm moths in the Yellowstone Ecosystem

    Science.gov (United States)

    French, Steven P.; French, Marilynn G.; Knight, Richard R.

    1994-01-01

    The ecology of alpine aggregations of army cutworm moths (Euxoa auxiliaris) and the feeding behavior of grizzly bears (Ursus arctos horribilis) at these areas were studied in the Yellowstone ecosystem from 1988 to 1991. Army cutworm moths migrate to mountain regions each summer to feed at night on the nectar of alpine and subalpine flowers, and during the day they seek shelter under various rock formations. Grizzly bears were observed feeding almost exclusively on moths up to 3 months each summer at the 10 moth-aggregation areas we identified. Fifty-one different grizzly bears were observed feeding at 4 of these areas during a single day in August 1991. Army cutworm moths are a preferred source of nutrition for many grizzly bears in the Yellowstone ecosystem and represent a high quality food that is available during hyperphagia.

  13. Tree-ring 14C links seismic swarm to CO2 spike at Yellowstone, USA

    Science.gov (United States)

    Evans, William C.; Bergfeld, D.; McGeehin, J.P.; King, J.C.; Heasler, H.

    2010-01-01

    Mechanisms to explain swarms of shallow seismicity and inflation-deflation cycles at Yellowstone caldera (western United States) commonly invoke episodic escape of magma-derived brines or gases from the ductile zone, but no correlative changes in the surface efflux of magmatic constituents have ever been documented. Our analysis of individual growth rings in a tree core from the Mud Volcano thermal area within the caldera links a sharp ~25% drop in 14C to a local seismic swarm in 1978. The implied fivefold increase in CO2 emissions clearly associates swarm seismicity with upflow of magma-derived fluid and shows that pulses of magmatic CO2 can rapidly traverse the 5-kmthick brittle zone, even through Yellowstone's enormous hydrothermal reservoir. The 1978 event predates annual deformation surveys, but recognized connections between subsequent seismic swarms and changes in deformation suggest that CO2 might drive both processes. ?? 2010 Geological Society of America.

  14. Monitoring changes in Greater Yellowstone Lake water quality following the 1988 wildfires

    Science.gov (United States)

    Lathrop, Richard G., Jr.; Vande Castle, John D.; Brass, James A.

    1994-01-01

    The fires that burned the Greater Yellowstone Area (GYA) during the summer of 1988 were the largest ever recorded for the region. Wildfire can have profound indirect effects on associated aquatic ecosystems by increased nutrient loading, sediment, erosion, and runoff. Satellite remote sensing and water quality sampling were used to compare pre- versus post-fire conditions in the GYA's large oliotrophic (high transparency, low productivity) lakes. Inputs of suspended sediment to Jackson Lake appear to have increased. Yellowstone Lake has not shown any discernable shift in water quality. The insights gained separately from the Landsat Thematic and NOAA Advanced Very High Resolution Radiometer (AVHRR) remote sensing systems, along with conventional in-situ sampling, can be combined into a useful water quality monitoring tool.

  15. Water tables constrain height recovery of willow on Yellowstone's northern range.

    Science.gov (United States)

    Bilyeu, Danielle M; Cooper, David J; Hobbs, N Thompson

    2008-01-01

    Excessive levels of herbivory may disturb ecosystems in ways that persist even when herbivory is moderated. These persistent changes may complicate efforts to restore ecosystems affected by herbivores. Willow (Salix spp.) communities within the northern range in Yellowstone National Park have been eliminated or degraded in many riparian areas by excessive elk (Cervus elaphus L.) browsing. Elk browsing of riparian willows appears to have diminished following the reintroduction of wolves (Canis lupis L.), but it remains uncertain whether reduced herbivory will restore willow communities. The direct effects of elk browsing on willows have been accompanied by indirect effects from the loss of beaver (Castor canadensis Kuhl) activity, including incision of stream channels, erosion of fine sediments, and lower water tables near streams historically dammed by beaver. In areas where these changes have occurred, lowered water tables may suppress willow height even in the absence of elk browsing. We conducted a factorial field experiment to understand willow responses to browsing and to height of water tables. After four years of protection from elk browsing, willows with ambient water tables averaged only 106 cm in height, with negligible height gain in two of three study species during the last year of the experiment. Willows that were protected from browsing and had artificially elevated water tables averaged 147 cm in height and gained 19 cm in the last year of the experiment. In browsed plots, elevated water tables doubled height gain during a period of slightly reduced browsing pressure. We conclude that water availability mediates the rate of willow height gain and may determine whether willows grow tall enough to escape the browse zone of elk and gain resistance to future elk browsing. Consequently, in areas where long-term beaver absence has resulted in incised stream channels and low water tables, a reduction in elk browsing alone may not be sufficient for recovery

  16. Quantifying Carbon Consequences of Recent Land Management, Disturbances and Subsequent Forest Recovery in the Greater Yellowstone Ecosystems (GYE)

    Science.gov (United States)

    Zhao, F. R.; Healey, S. P.; McCarter, J. B.; Garrard, C.; Zhu, Z.; Huang, C.

    2016-12-01

    Natural disturbances and land management directly alter C stored in biomass and soil pools, and forest recovery following these events are critical for long-term regional C balance. The Greater Yellowstone Ecosystem (GYE), located in Central Rocky Mountains of United States, is of different land ownerships within similar environmental settings, making it an ideal site to examine the impacts of land management, disturbance and forest recovery on regional C dynamics. Recent advances in the remote sensing of vegetation condition and change, along with new techniques linking remote sensing with inventory records, have allowed investigations that are much more tightly constrained to actual landscape environments instead of hypothetical or generalized conditions. These new capabilities are built into the Forest Carbon Management Framework (ForCaMF), which is being used by the National Forest System to not only model, but to monitor across very specific management units, the impact of different kinds of disturbance on carbon storage. In this study, we used the ForCaMF approach to examine three C related management questions in GYE National Parks and National Forests: 1) what was the carbon storage impact of fire disturbance and management activities from 1985 to 2010 in the GYE National Parks and National Forests? 2) Using an historic fire that occurred in 1988 as a basis for comparison, what difference would active post-fire forest restoration make in subsequent C storage? 3) In light of the fact that GYE National Forests significantly reduced harvest rates in the 1990s, how would maintaining high harvest rates of the 1980s impacted C storage? Simulation results show that recent forest fires in the GYE National Parks induced an accumulative C storage loss of about 12 Mg/ha, compared with C storage loss up to 2 Mg/ha in the GYE National Forests by harvests. If the high harvest rates as of the 1980s had been maintained, C emissions from the National Forests ( 11 Mg

  17. Multiscale Genetic Structure of Yellowstone Cutthroat Trout in the Upper Snake River Basin.

    Energy Technology Data Exchange (ETDEWEB)

    Cegelski, Christine C.; Campbell, Matthew R.

    2006-05-30

    Populations of Yellowstone cutthroat trout Oncorhynchus clarkii bouvierii have declined throughout their native range as a result of habitat fragmentation, overharvest, and introductions of nonnative trout that have hybridized with or displaced native populations. The degree to which these factors have impacted the current genetic population structure of Yellowstone cutthroat trout populations is of primary interest for their conservation. In this study, we examined the genetic diversity and genetic population structure of Yellowstone cutthroat trout in Idaho and Nevada with data from six polymorphic microsatellite loci. A total of 1,392 samples were analyzed from 45 sample locations throughout 11 major river drainages. We found that levels of genetic diversity and genetic differentiation varied extensively. The Salt River drainage, which is representative of the least impacted migration corridors in Idaho, had the highest levels of genetic diversity and low levels of genetic differentiation. High levels of genetic differentiation were observed at similar or smaller geographic scales in the Portneuf River, Raft River, and Teton River drainages, which are more altered by anthropogenic disturbances. Results suggested that Yellowstone cutthroat trout are naturally structured at the major river drainage level but that habitat fragmentation has altered this structuring. Connectivity should be restored via habitat restoration whenever possible to minimize losses in genetic diversity and to preserve historical processes of gene flow, life history variation, and metapopulation dynamics. However, alternative strategies for management and conservation should also be considered in areas where there is a strong likelihood of nonnative invasions or extensive habitat fragmentation that cannot be easily ameliorated.

  18. A Long-Term Comparison of Yellowstone Cutthroat Trout Abundance and Size Structure in Their Historical Range in Idaho.

    Energy Technology Data Exchange (ETDEWEB)

    Meyer, Kevin A.; Schill, Daniel J.; Elle, F. Steven

    2002-05-23

    We compared estimates of population abundance and size structure for Yellowstone cutthroat trout Oncorhynchus clarki bouvieri obtained by electrofishing 77 stream segments across southeastern Idaho in the 1980s and again in 1999-2000 to test whether populations of Yellowstone cutthroat trout had changed. Sites sampled in the 1980s were relocated in 1999-2000 by using maps and photographs or by finding original site-boundary stakes, so that the same reach of stream was sampled during both periods. Abundance of Yellowstone cutthroat trout longer than 10 cm did not change, averaging 41 fish/100 m of stream during both the 1980s and 1999-2000. The proportion of the total catch of trout composed of Yellowstone cutthroat trout also did not change, averaging 82% in the 1980s and 78% in 1999-2000. At the 48 sites where size structure could be estimated for both periods, the proportion of Yellowstone cutthroat trout that were 10-20 cm long declined slightly (74% versus 66%), but the change was due entirely to the shift in size structure at the Teton River sites. The number of sites that contained rainbow trout O. mykiss or cutthroat trout 3 rainbow trout hybrids rose from 23 to 37, but the average proportion of the catch composed of rainbow trout and hybrids did not increase (7% in both the 1980s and 1999-2000). Although the distribution and abundance of Yellowstone cutthroat trout have been substantially reduced in Idaho over the last century, our results indicate that Yellowstone cutthroat trout abundance and size structure in Idaho have remained relatively stable at a large number of locations for the last 10-20 years. The expanding distribution of rainbow trout and hybrids in portions of the upper Snake River basin, however, calls for additional monitoring and active management actions.

  19. Use of naturally occurring mercury to determine the importance of cutthroat trout to Yellowstone grizzly bears

    Science.gov (United States)

    Felicetti, L.A.; Schwartz, C.C.; Rye, R.O.; Gunther, K.A.; Crock, J.G.; Haroldson, M.A.; Waits, L.; Robbins, C.T.

    2004-01-01

    Spawning cutthroat trout (Oncorhynchus clarki (Richardson, 1836)) are a potentially important food resource for grizzly bears (Ursus arctos horribilis Ord, 1815) in the Greater Yellowstone Ecosystem. We developed a method to estimate the amount of cutthroat trout ingested by grizzly bears living in the Yellowstone Lake area. The method utilized (i) the relatively high, naturally occurring concentration of mercury in Yellowstone Lake cutthroat trout (508 ± 93 ppb) and its virtual absence in all other bear foods (6 ppb), (ii) hair snares to remotely collect hair from bears visiting spawning cutthroat trout streams between 1997 and 2000, (iii) DNA analyses to identify the individual and sex of grizzly bears leaving a hair sample, (iv) feeding trials with captive bears to develop relationships between fish and mercury intake and hair mercury concentrations, and (v) mercury analyses of hair collected from wild bears to estimate the amount of trout consumed by each bear. Male grizzly bears consumed an average of 5 times more trout/kg bear than did female grizzly bears. Estimated cutthroat trout intake per year by the grizzly bear population was only a small fraction of that estimated by previous investigators, and males consumed 92% of all trout ingested by grizzly bears.

  20. Simulation of water-rock interaction in the yellowstone geothermal system using TOUGHREACT

    International Nuclear Information System (INIS)

    Dobson, P.F.; Salah, S.; Spycher, N.; Sonnenthal, E.

    2003-01-01

    The Yellowstone geothermal system provides an ideal opportunity to test the ability of reactive transport models to accurately simulate water-rock interaction. Previous studies of the Yellowstone geothermal system have characterized water-rock interaction through analysis of rocks and fluids obtained from both surface and downhole samples. Fluid chemistry, rock mineralogy, permeability, porosity, and thermal data obtained from the Y-8 borehole in Upper Geyser Basin were used to constrain a series of reactive transport simulations of the Yellowstone geothermal system using TOUGHREACT. Three distinct stratigraphic units were encountered in the 153.4 m deep Y-8 drill core: volcaniclastic sandstone, perlitic rhyolitic lava, and nonwelded pumiceous tuff. The main alteration phases identified in the Y-8 core samples include clay minerals, zeolites, silica polymorphs, adularia, and calcite. Temperatures observed in the Y-8 borehole increase with depth from sub-boiling conditions at the surface to a maximum of 169.8 C at a depth of 104.1 m, with near-isothermal conditions persisting down to the well bottom. 1-D models of the Y-8 core hole were constructed to determine if TOUGHREACT could accurately predict the observed alteration mineral assemblage given the initial rock mineralogy and observed fluid chemistry and temperatures. Preliminary simulations involving the perlitic rhyolitic lava unit are consistent with the observed alteration of rhyolitic glass to form celadonite

  1. Growth, morphology, and developmental instability of rainbow trout, Yellowstone cutthroat trout, and four hybrid generations

    Science.gov (United States)

    Ostberg, C.O.; Duda, J.J.; Graham, J.H.; Zhang, S.; Haywood, K. P.; Miller, B.; Lerud, T.L.

    2011-01-01

    Hybridization of cutthroat trout Oncorhynchus clarkii with nonindigenous rainbow trout O. mykiss contributes to the decline of cutthroat trout subspecies throughout their native range. Introgression by rainbow trout can swamp the gene pools of cutthroat trout populations, especially if there is little selection against hybrids. We used rainbow trout, Yellowstone cutthroat trout O. clarkii bouvieri, and rainbow trout × Yellowstone cutthroat trout F1 hybrids as parents to construct seven different line crosses: F1 hybrids (both reciprocal crosses), F2 hybrids, first-generation backcrosses (both rainbow trout and Yellowstone cutthroat trout), and both parental taxa. We compared growth, morphology, and developmental instability among these seven crosses reared at two different temperatures. Growth was related to the proportion of rainbow trout genome present within the crosses. Meristic traits were influenced by maternal, additive, dominant, overdominant, and (probably) epistatic genetic effects. Developmental stability, however, was not disturbed in F1 hybrids, F2 hybrids, or backcrosses. Backcrosses were morphologically similar to their recurrent parent. The lack of developmental instability in hybrids suggests that there are few genetic incompatibilities preventing introgression. Our findings suggest that hybrids are not equal: that is, growth, development, character traits, and morphology differ depending on the genomic contribution from each parental species as well as the hybrid generation.

  2. Nation

    DEFF Research Database (Denmark)

    Østergaard, Uffe

    2014-01-01

    Nation er et gammelt begreb, som kommer af det latinske ord for fødsel, natio. Nationalisme bygger på forestillingen om, at mennesker har én og kun én national identitet og har ret til deres egen nationalstat. Ordet og forestillingen er kun godt 200 år gammel, og i 1900-tallet har ideologien bredt...

  3. Response of Yellowstone grizzly bears to changes in food resources: A synthesis. Final report to the Interagency Grizzly Bear Committee and Yellowstone Ecosystem Subcommittee

    Science.gov (United States)

    ,; van Manen, Frank T.; Costello, Cecily M.; Haroldson, Mark A.; Bjornlie, Daniel D.; Ebinger, Michael R.; Gunther, Kerry A.; Mahalovich, Mary Frances; Thompson, Daniel J.; Higgs, Megan D.; Irvine, Kathryn M.; Legg, Kristin; Tyers, Daniel B.; Landenburger, Lisa; Cain, Steven L.; Frey, Kevin L.; Aber, Bryan C.; Schwartz, Charles C.

    2013-01-01

    The Yellowstone grizzly bear (Ursus arctos) was listed as a threatened species in 1975 (Federal Register 40 FR:31734-31736). Since listing, recovery efforts have focused on increasing population size, improving habitat security, managing bear mortalities, and reducing bear-human conflicts. The Interagency Grizzly Bear Committee (IGBC; partnership of federal and state agencies responsible for grizzly bear recovery in the lower 48 states) and its Yellowstone Ecosystem Subcommitte (YES; federal, state, county, and tribal partners charged with recovery of grizzly bears in the Greater Yelowston Ecosystem [GYE]) tasked the Interagency Grizzly Bear Study Team to provide information and further research relevant to three concerns arising from the 9th Circuit Court of Appeals November 2011 decision: 1) the ability of grizzly bears as omnivores to find alternative foods to whitebark pine seeds; 2) literature to support their conclusions; and 3) the non-intuitive biological reality that impacts can occur to individuals without causing the overall population to decline. Specifically, the IGBC and YES requested a comprehensive synthesis of the current state of knowledge regarding whitebark pinbe decline and individual and population-level responses of grizzly bears to changing food resources in the GYE. This research was particularly relevant to grizzly bear conservation given changes in the population trajectory observed during the last decade.

  4. Hurricane Gustav Aerial Photography: Rapid ResponseImagery of the Surrounding Regions After Landfall

    Data.gov (United States)

    National Oceanic and Atmospheric Administration, Department of Commerce — The imagery posted on this site is of the surrounding regionsafter Hurricane Gustav made landfall. The aerial photography missions wereconducted by the NOAA Remote...

  5. Hurricane Dennis Aerial Photography: Draft Image Mosaics of the Florida Panhandle and Surrounding Regions After Landfall

    Data.gov (United States)

    National Oceanic and Atmospheric Administration, Department of Commerce — The imagery posted on this site is of the Florida panhandle and surrounding regions after Hurricane Dennis made landfall. The regions photographed range from...

  6. Hurricane Humberto Aerial Photography: Rapid Response Imagery of the Surrounding Regions After Landfall

    Data.gov (United States)

    National Oceanic and Atmospheric Administration, Department of Commerce — The imagery posted on this site is of the surrounding regions after Hurricane Humberto made landfall. The aerial photography missions were conducted by the NOAA...

  7. Hurricane Ike Aerial Photography: Rapid ResponseImagery of the Surrounding Regions After Landfall

    Data.gov (United States)

    National Oceanic and Atmospheric Administration, Department of Commerce — The imagery posted on this site is of the surrounding regionsafter Hurricane Ike made landfall. The aerial photography missions wereconducted by the NOAA Remote...

  8. Tropical Storm Ernesto Aerial Photography: Rapid Response Imagery of the Surrounding Regions After Landfall

    Data.gov (United States)

    National Oceanic and Atmospheric Administration, Department of Commerce — The imagery posted on this site is of the surrounding regions after Tropical Storm Ernesto made landfall. The aerial photography missions were conducted by the NOAA...

  9. Enhancement of Afterimage Colors by Surrounding Contours

    Directory of Open Access Journals (Sweden)

    Takao Sato

    2011-05-01

    Full Text Available Presenting luminance contours surrounding the adapted areas in test phase enhances color afterimages in both duration and color appearance. The presence of surrounding contour is crucial to some color phenomenon such as van Lier's afterimage, but the contour-effect itself has not been seriously examined. In this paper, we compared the contour-effect to color afterimages and to actually colored patches to examine the nature of color information subserving color-aftereffect. In the experiment, observers were adapted for 1 sec to a small colored square (red, green, yellow, or blue presented on a gray background. Then, a test field either with or without surrounding contour was presented. Observers matched the color of a test-patch located near the afterimage to the color of afterimage. It was found that the saturation of negative afterimage was almost doubled by the presence of surrounding contours. There was no effect of luminance contrast or polarity of contours. In contrast, no enhancement of saturation by surrounding contours was observed for actually colored patches even though the colors of patches were equalized to that of afterimage without contours. This dissociation in the contour-effect demonstrates the crucial difference between the color information for aftereffects and for ordinary bottom-up color perception.

  10. Chronology of awareness about US National Park external threats.

    Science.gov (United States)

    Shafer, Craig L

    2012-12-01

    The objective of this paper is to raise understanding of the history of protected area external threat awareness in the United States and at World Protected Area Congresses. The earliest concerns about external threats to US national parks began in the late nineteenth century: a potential railroad transgression of Yellowstone National Park in the 1880s. During the early and mid 1930s, George Wright and colleagues focused on outside boundary concerns like of hunting and trapping of furbearers, grazing, logging, disease and hybridization between species. In the 1960s, a worldwide recognition began about the role of outside habitat fragmentation/isolation on nature reserves and human generated stressors crossing their boundaries. The State of the Park Report 1980 added a plethora of threats: oil/gas and geothermal exploration and development, hydropower and reclamation projects, urban encroachment, roads, resorts, and recreational facilities. The early 1980s ushered in political interference with NPS threats abatement efforts as well as Congressional legislative initiatives to support the abatement challenges of the agency. By 1987, the Government Accounting Office issued its first report on National Park Service (NPS) progress in dealing with external threats. Climate change impacts on parks, especially in terms of animals adjusting their temperature and moisture requirements by latitude and altitude, surfaced in the technical literature by the mid-1980s. By 1992, the world parks community stressed the need to integrate protected areas into the surrounding landscape and human community. The importance of the matrix has gradually gained appreciation in the scientific community. This chronology represents one example of national park and protected areas' institutional history contributing to the breath of modern conservation science.

  11. Explaining preferences for home surroundings and locations

    DEFF Research Database (Denmark)

    Andersen, Hans Skifter

    2011-01-01

    This article is based on a survey carried out in Denmark that asked a random sample of the population about their preferences for home surroundings and locations. It shows that the characteristics of social surroundings are very important and can be divided into three independent dimensions......: avoiding social nuisances, preferring social homogeneity and living close to one’s social network and place of origin. The study shows that most people have many detailed preferences, whereas some have very few. This confirms an earlier theory that some people are very connected to certain places...... with given characteristics and thus do not have priorities regarding home surroundings and locations. For others, mostly young people and singles, home is just a place to sleep and relax, whereas life is lived elsewhere. For this group, there are only preferences for location and there are few specific...

  12. Smart Chips for Smart Surroundings -- 4S

    NARCIS (Netherlands)

    Schuler, Eberhard; König, Ralf; Becker, Jürgen; Rauwerda, G.K.; van de Burgwal, M.D.; Smit, Gerardus Johannes Maria; Cardoso, João M.P.; Hübner, Michael

    2011-01-01

    The overall mission of the 4S project (Smart Chips for Smart Surroundings) was to define and develop efficient flexible, reconfigurable core building blocks, including the supporting tools, for future Ambient System Devices. Reconfigurability offers the needed flexibility and adaptability, it

  13. Childhood Suicide and Myths Surrounding It.

    Science.gov (United States)

    Greene, Dorothea B.

    1994-01-01

    Dispels five misconceptions surrounding the suicide of children: that children under the age of six do not commit suicide; that suicide in latency years is extremely rare; that psychodynamically and developmentally true depression is not possible in childhood; that child cannot understand finality of death; and that children are cognitively and…

  14. Identification of β-SiC surrounded by relatable surrounding diamond ...

    Indian Academy of Sciences (India)

    β-SiC is identified in the presence of a relatable surrounding diamond medium using subtle, but discernible Raman ... Change in the nature of the surrounding material structure and its .... intensity implies very low graphite content in thin film. In.

  15. Volcano crisis response at Yellowstone volcanic complex - after-action report for exercise held at Salt Lake City, Utah, November 15, 2011

    Science.gov (United States)

    Pierson, Thomas C.; Driedger, Carolyn L.; Tilling, Robert I.

    2013-01-01

    A functional tabletop exercise was run on November 14-15, 2011 in Salt Lake City, Utah, to test crisis response capabilities, communication protocols, and decision-making by the staff of the multi-agency Yellowstone Volcano Observatory (YVO) as they reacted to a hypothetical exercise scenario of accelerating volcanic unrest at the Yellowstone caldera. The exercise simulated a rapid build-up of seismic activity, ground deformation, and hot-spring water-chemistry and temperature anomalies that culminated in a small- to moderate-size phreatomagmatic eruption within Yellowstone National Park. The YVO scientific team's responses to the unfolding events in the scenario and to simulated requests for information by stakeholders and the media were assessed by (a) the exercise organizers; (b) several non-YVO scientists, who observed and queried participants, and took notes throughout the exercise; and (c) the participants themselves, who kept logs of their actions during the exercise and later participated in a group debriefing session and filled out detailed questionnaires. These evaluations were tabulated, interpreted, and summarized for this report, and on the basis of this information, recommendations have been made. Overall, the YVO teams performed their jobs very well. The exercise revealed that YVO scientists were able to successfully provide critical hazards information, issue information statements, and appropriately raise alert levels during a fast-moving crisis. Based on the exercise, it is recommended that several measures be taken to increase YVO effectiveness during a crisis: 1. Improve role clarification within and between YVO science teams. 2. Improve communications tools and protocols for data-sharing and consensus-building among YVO scientists, who are geographically and administratively dispersed among various institutions across the United States. 3. Familiarize YVO staff with Incident Command System (ICS) procedures and protocols, and provide more in

  16. A sense of place: Ecoregional design at Mesa Verde National Park

    Science.gov (United States)

    Robert G. Bailey

    2012-01-01

    When the National Park Service was established in 1916, the new agency inherited an architectural legacy developed by private interests, particularly the railroads. This legacy included Northern Pacific's Old Faithful Inn in Yellowstone and Santa Fe's El Tovar at the south rim of the Grand Canyon, both built in the Swiss Chalet­Norway Villa tradition. This...

  17. Travel Times, Streamflow Velocities, and Dispersion Rates in the Yellowstone River, Montana

    Science.gov (United States)

    McCarthy, Peter M.

    2009-01-01

    The Yellowstone River is a vital natural resource to the residents of southeastern Montana and is a primary source of water for irrigation and recreation and the primary source of municipal water for several cities. The Yellowstone River valley is the primary east-west transportation corridor through southern Montana. This complex of infrastructure makes the Yellowstone River especially vulnerable to accidental spills from various sources such as tanker cars and trucks. In 2008, the U.S. Geological Survey (USGS), in cooperation with the Montana Department of Environmental Quality, initiated a dye-tracer study to determine instream travel times, streamflow velocities, and dispersion rates for the Yellowstone River from Lockwood to Glendive, Montana. The purpose of this report is to describe the results of this study and summarize data collected at each of the measurement sites between Lockwood and Glendive. This report also compares the results of this study to estimated travel times from a transport model developed by the USGS for a previous study. For this study, Rhodamine WT dye was injected at four locations in late September and early October 2008 during reasonably steady streamflow conditions. Streamflows ranged from 3,490 to 3,770 cubic feet per second upstream from the confluence of the Bighorn River and ranged from 6,520 to 7,570 cubic feet per second downstream from the confluence of the Bighorn River. Mean velocities were calculated for each subreach between measurement sites for the leading edge, peak concentration, centroid, and trailing edge at 10 percent of the peak concentration. Calculated velocities for the centroid of the dye plume for subreaches that were completely laterally mixed ranged from 1.83 to 3.18 ft/s within the study reach from Lockwood Bridge to Glendive Bridge. The mean of the completely mixed centroid velocity for the entire study reach, excluding the subreach between Forsyth Bridge and Cartersville Dam, was 2.80 ft/s. Longitudinal

  18. Origin and Evolution of the Yellowstone Hotspot from Seismic-GPS Imaging and Geodynamic Modeling

    Science.gov (United States)

    Smith, R. B.; Jordan, M.; Puskas, C. M.; Farrell, J.; Waite, G. P.

    2006-12-01

    The Yellowstone hotspot resulted from interaction of a mantle plume with the overriding North America plate. This feature and related processes have influenced a large part of the western U.S., producing the 16 Ma Yellowstone-Snake River Plain-Newberry silicic-basalt volcanic field (YSRPN). We integrate results from a multi-institution experiment that deployed 80 seismic stations and 160 campaign and 21 permanent GPS stations for 1999-2003. Crust and mantle velocity models were derived from inversion of teleseismic and local earthquake data. Kinematic and dynamic models were derived from inversion of GPS velocities constrained by stresses associated the topography and the +15 m geoid anomaly. Tomography revealed a P- and S-wave low-velocity body at depths of 8-16 km beneath the caldera that is interpreted as partial melt of 8-15% that feeds the youthful Yellowstone volcanic field. Volume changes in the magma chamber are responsible for GPS-measured episodes of uplift and subsidence of the caldera at decadal scales with average rates of ~20 mm/yr but much higher short-term rates of up to 80 mm/yr. An upper-mantle low-velocity body was imaged by inverting teleseismic data constrained by the geoid structure, crustal structure, and the upper mantle discontinuities. This low P and S velocity body extends from 80 km to ~250 km directly beneath Yellowstone and then continues to 650 km with unexpected tilt to the west at ~60°. The tilt is consistent with the ascent of the buoyant magma entrained in eastward return-flow of the upper mantle. We estimate this body has an excess temperature from 85K to 120K, depending on the water content and with up to 1.5% melt. Using the inclined plume-geometry and plate motion history, we extrapolate the Yellowstone mantle source southwestward ~800 km as a plume-head in oceanic lithosphere centered beneath the Columbia Plateau basalt field at 16 Ma. Magma ascent was truncated there by the passage of thicker continental lithosphere over

  19. A fluid-driven earthquake swarm on the margin of the Yellowstone caldera

    Science.gov (United States)

    Shelly, David R.; Hill, David P.; Massin, Frederick; Farrell, Jamie; Smith, Robert B.; Taira, Taka'aki

    2013-01-01

    Over the past several decades, the Yellowstone caldera has experienced frequent earthquake swarms and repeated cycles of uplift and subsidence, reflecting dynamic volcanic and tectonic processes. Here, we examine the detailed spatial-temporal evolution of the 2010 Madison Plateau swarm, which occurred near the northwest boundary of the Yellowstone caldera. To fully explore the evolution of the swarm, we integrated procedures for seismic waveform-based earthquake detection with precise double-difference relative relocation. Using cross-correlation of continuous seismic data and waveform templates constructed from cataloged events, we detected and precisely located 8710 earthquakes during the three-week swarm, nearly four times the number of events included in the standard catalog. This high-resolution analysis reveals distinct migration of earthquake activity over the course of the swarm. The swarm initiated abruptly on January 17, 2010 at about 10 km depth and expanded dramatically outward (both shallower and deeper) over time, primarily along a NNW-striking, ~55º ENE-dipping structure. To explain these characteristics, we hypothesize that the swarm was triggered by the rupture of a zone of confined high-pressure aqueous fluids into a pre-existing crustal fault system, prompting release of accumulated stress. The high-pressure fluid injection may have been accommodated by hybrid shear and dilatational failure, as is commonly observed in exhumed hydrothermally affected fault zones. This process has likely occurred repeatedly in Yellowstone as aqueous fluids exsolved from magma migrate into the brittle crust, and it may be a key element in the observed cycles of caldera uplift and subsidence.

  20. The Yellowstone ‘hot spot’ track results from migrating Basin Range extension

    Science.gov (United States)

    Foulger, Gillian R.; Christiansen, Robert L.; Anderson, Don L.; Foulger, Gillian R.; Lustrino, Michele; King, Scott D.

    2015-01-01

    Whether the volcanism of the Columbia River Plateau, eastern Snake River Plain, and Yellowstone (western U.S.) is related to a mantle plume or to plate tectonic processes is a long-standing controversy. There are many geological mismatches with the basic plume model as well as logical flaws, such as citing data postulated to require a deep-mantle origin in support of an “upper-mantle plume” model. USArray has recently yielded abundant new seismological results, but despite this, seismic analyses have still not resolved the disparity of opinion. This suggests that seismology may be unable to resolve the plume question for Yellowstone, and perhaps elsewhere. USArray data have inspired many new models that relate western U.S. volcanism to shallow mantle convection associated with subduction zone processes. Many of these models assume that the principal requirement for surface volcanism is melt in the mantle and that the lithosphere is essentially passive. In this paper we propose a pure plate model in which melt is commonplace in the mantle, and its inherent buoyancy is not what causes surface eruptions. Instead, it is extension of the lithosphere that permits melt to escape to the surface and eruptions to occur—the mere presence of underlying melt is not a sufficient condition. The time-progressive chain of rhyolitic calderas in the eastern Snake River Plain–Yellowstone zone that has formed since basin-range extension began at ca. 17 Ma results from laterally migrating lithospheric extension and thinning that has permitted basaltic magma to rise from the upper mantle and melt the lower crust. We propose that this migration formed part of the systematic eastward migration of the axis of most intense basin-range extension. The bimodal rhyolite-basalt volcanism followed migration of the locus of most rapid extension, not vice versa. This model does not depend on seismology to test it but instead on surface geological observations.

  1. Thermal controls of Yellowstone cutthroat trout and invasive fishes under climate change

    Science.gov (United States)

    Al-Chokhachy, Robert K.; Alder, Jay R.; Hostetler, Steven W.; Gresswell, Robert E.; Shepard, Bradley

    2013-01-01

    We combine large observed data sets and dynamically downscaled climate data to explore historic and future (2050–2069) stream temperature changes over the topographically diverse Greater Yellowstone Ecosystem (elevation range = 824–4017 m). We link future stream temperatures with fish growth models to investigate how changing thermal regimes could influence the future distribution and persistence of native Yellowstone cutthroat trout (YCT) and competing invasive species. We find that stream temperatures during the recent decade (2000–2009) surpass the anomalously warm period of the 1930s. Climate simulations indicate air temperatures will warm by 1 °C to >3 °C over the Greater Yellowstone by mid-21st century, resulting in concomitant increases in 2050–2069 peak stream temperatures and protracted periods of warming from May to September (MJJAS). Projected changes in thermal regimes during the MJJAS growing season modify the trajectories of daily growth rates at all elevations with pronounced growth during early and late summer. For high-elevation populations, we find considerable increases in fish body mass attributable both to warming of cold-water temperatures and to extended growing seasons. During peak July to August warming, mid-21st century temperatures will cause periods of increased thermal stress, rendering some low-elevation streams less suitable for YCT. The majority (80%) of sites currently inhabited by YCT, however, display minimal loss (changes in total body mass by midcentury; we attribute this response to the fact that many low-elevation populations of YCT have already been extirpated by historical changes in land use and invasions of non-native species. Our results further suggest that benefits to YCT populations due to warmer stream temperatures at currently cold sites could be offset by the interspecific effects of corresponding growth of sympatric, non-native species, underscoring the importance of developing climate adaptation

  2. Agricultural implications of reduced water supplies in the Green and Upper Yellowstone River Basins

    Energy Technology Data Exchange (ETDEWEB)

    Lansford, R. R.; Roach, F.; Gollehon, N. R.; Creel, B. J.

    1982-02-01

    The growth of the energy sector in the energy-rich but water-restricted Western US has presented a potential conflict with the irrigated agricultural sector. This study measures the direct impacts on farm income and employment resulting from the transfer of water from agriculture to energy in two specific geographical areas - the Green and Upper Yellowstone River Basins. We used a linear programming model to evaluate the impacts of reduced water supplies. Through the use of regional multipliers, we expanded our analysis to include regional impacts. Volume I provides the major analysis of these impacts. Volume II provides further technical data.

  3. Beyond the Inventory: An Interagency Collaboration to Reduce Greenhouse Gas Emissions in the Greater Yellowstone Area

    Energy Technology Data Exchange (ETDEWEB)

    Kandt, A.; Hotchkiss, E.; Fiebig, M.

    2010-10-01

    As one of the largest, intact ecosystems in the continental United States, land managers within the Greater Yellowstone Area (GYA) have recognized the importance of compiling and understanding agency greenhouse gas (GHG) emissions. The 10 Federal units within the GYA have taken an active role in compiling GHG inventories on a unit- and ecosystem-wide level, setting goals for GHG mitigation, and identifying mitigation strategies for achieving those goals. This paper details the processes, methodologies, challenges, solutions, and lessons learned by the 10 Federal units within the GYA throughout this ongoing effort.

  4. Temporal, spatial, and environmental influences on the demographics of grizzly bears in the Greater Yellowstone Ecosystem

    Science.gov (United States)

    Schwartz, Charles C.; Haroldson, Mark A.; White, Gary C.; Harris, Richard B.; Cherry, Steve; Keating, Kim A.; Moody, Dave; Servheen, Christopher

    2006-01-01

    During the past 2 decades, the grizzly bear (Ursus arctos) population in the Greater Yellowstone Ecosystem (GYE) has increased in numbers and expanded in range. Understanding temporal, environmental, and spatial variables responsible for this change is useful in evaluating what likely influenced grizzly bear demographics in the GYE and where future management efforts might benefit conservation and management. We used recent data from radio-marked bears to estimate reproduction (1983–2002) and survival (1983–2001); these we combined into models to evaluate demographic vigor (lambda [λ]). We explored the influence of an array of individual, temporal, and spatial covariates on demographic vigor.

  5. Filamentous Morphology as a Means for Thermophilic Bacteria to Survive Steep Physical and Chemical Gradients in Yellowstone Hot Springs

    Science.gov (United States)

    Dong, Y.; Srivastava, V.; Bulone, V.; Keating, K. M.; Khetani, R. S.; Fields, C. J.; Inskeep, W.; Sanford, R. A.; Yau, P. M.; Imai, B. S.; Hernandez, A. G.; Wright, C.; Band, M.; Cann, I. K.; Ahrén, D.; Fouke, K. W.; Sivaguru, M.; Fried, G.; Fouke, B. W.

    2017-12-01

    The filamentous heat-loving bacterium Sulfurihydrogenibium yellowstonense makes up more than 90% of the microbial community that inhabits turbulent, dysoxic hot spring outflow channels (66-71°C, 6.2-6.5 pH, 0.5-0.75 m/s flow rate) at Mammoth Hot Spring in Yellowstone National Park. These environments contain abundantly available inorganic substrates (e.g., CO2, sulfide and thiosulfate) and are associated with extensive CaCO3 (travertine) precipitation driven in part by CO2 off-gassing. Evidence from integrated Meta-Omics analyses of DNA, RNA, and proteins (metagenomics, metatranscriptomics and metaproteomics) extracted from these S. yellowstonense-dominated communities have detected 1499 non-rRNA open reading frames (ORFs), their transcripts and cognate proteins. During chemoautotrophy and CO2 carbon fixation, chaperons facilitate enzymatic stability and functionalities under elevated temperature. High abundance transcripts and proteins for Type IV pili and exopolysaccharides (EPS) are consistent with S. yellowstonense forming strong (up to 0.5 m) intertwined microbial filaments (fettuccini streamers) composed of linked individual cells that withstand hydrodynamic shear forces and extremely rapid travertine mineralization. Their primary energy source is the oxidation of reduced sulfur (e.g., sulphide, sulfur or thiosulfate) and the simultaneous uptake of extremely low concentrations of dissolved O2 facilitated by bd-type cytochromes. Field observations indicate that the fettuccini microbial filaments build up ridged travertine platforms on the bottom of the springs, parallel to the water flow, where living filaments attach almost exclusively to the top of each ridge. This maximizes their access to miniscule amounts of dissolved oxygen, while optimizing their ability to rapidly form down-flow branched filaments and thus survive in these stressful environments that few other microbes can inhabit.

  6. Real-time Remote Data Online For Norris Geyser Basin in Yellowstone National Park

    Science.gov (United States)

    Perry, J. E.; Lowenstern, J. B.; Clor, L.; Cervelli, P. F.; Allen, S. T.; Heasler, H.; Moloney, T.

    2010-12-01

    Perry, John, Lowenstern, Jacob, Cervelli, Peter, Clor, Laura, Heasler, Henry, Allen, Scott, Moloney, Tim During June 2010, ten 900MHz wireless temperature data loggers (nodes) were installed around the Norris Geyser Basin to monitor geothermal features, streams and soil temperatures. The loggers can provide near real-time updates on temperature variations within 0.2 deg C due to hydrothermal discharges or subsurface fluid migration. Each sensor node is programmed to measure the temperature every two minutes and automatically upload data to the base station computer daily. The hardware consists of a waterproof case containing an M5 logger (made by Marathon Products, Inc.®) with internal memory, lithium D-cell batteries and a 900 MHz, 1-W-transceiver and 5 meter long Teflon-coated probe with a thermistor sensor. Tethered stub or panel antennas are oriented to optimize signal strength to the base station near the Norris Museum. A 0.61 meter-long base-station antenna located 10m high provides signal to the furthest node over 850 meters away with most being “line-of-sight”. A 20-meter coaxial cable and lightning grounding wire connects the base-station antenna to an Ethernet-radio connected to the YNP local-area network. A server located 26-km north at Mammoth Hot Springs requests data at regular intervals (normally daily), archives the information, and then sends it to the USGS for further archiving and internet distribution. During periods of unusual hydrothermal behavior, data can be requested as needed, and it is possible to set user-programmable alarm limits for notification. The RF network is designed to monitor changes from three different sub-basins at Norris (Gray Lakes, Steamboat-Echinus and Porcelain Basin), the main Tantalus Creek drainage, and five individual thermal features (Constant, Porkchop, Steamboat and Echinus Geysers, and Opalescent Spring). The logger installed in Nuphar Lake provides ambient temperatures controlled solely by local meteorological conditions. Soil temperature is measured near Vixen Geyser. The system will permit land managers, scientists and the public to keep track of changes to the hydrothermal system, including geyser eruptions, periodic basin-wide disturbances, or fluid-release events that may accompany or follow seismic activity.

  7. 77 FR 30321 - Proposed Concession Contract for Yellowstone National Park-Alternative Formula for Calculating...

    Science.gov (United States)

    2012-05-22

    .... Elective Franchise Fee Reduction/LSI Buy Down Provision The NPS also points out that it intends to include an elective franchise fee reduction/LSI buy down provision in the terms of the prospectus for YELL007... it would be in its best interest to elect to accept the elective franchise fee reduction/ LSI buy...

  8. Mapping forest canopy fuels in Yellowstone National Park using lidar and hyperspectral data

    Science.gov (United States)

    Halligan, Kerry Quinn

    The severity and size of wildland fires in the forested western U.S have increased in recent years despite improvements in fire suppression efficiency. This, along with increased density of homes in the wildland-urban interface, has resulted in high costs for fire management and increased risks to human health, safety and property. Crown fires, in comparison to surface fires, pose an especially high risk due to their intensity and high rate of spread. Crown fire models require a range of quantitative fuel parameters which can be difficult and costly to obtain, but advances in lidar and hyperspectral sensor technologies hold promise for delivering these inputs. Further research is needed, however, to assess the strengths and limitations of these technologies and the most appropriate analysis methodologies for estimating crown fuel parameters from these data. This dissertation focuses on retrieving critical crown fuel parameters, including canopy height, canopy bulk density and proportion of dead canopy fuel, from airborne lidar and hyperspectral data. Remote sensing data were used in conjunction with detailed field data on forest parameters and surface reflectance measurements. A new method was developed for retrieving Digital Surface Model (DSM) and Digital Canopy Models (DCM) from first return lidar data. Validation data on individual tree heights demonstrated the high accuracy (r2 0.95) of the DCMs developed via this new algorithm. Lidar-derived DCMs were used to estimate critical crown fire parameters including available canopy fuel, canopy height and canopy bulk density with linear regression model r2 values ranging from 0.75 to 0.85. Hyperspectral data were used in conjunction with Spectral Mixture Analysis (SMA) to assess fuel quality in the form of live versus dead canopy proportions. Severity and stage of insect-caused forest mortality were estimated using the fractional abundance of green vegetation, non-photosynthetic vegetation and shade obtained from SMA. Proportion of insect attack was estimated with a linear model producing an r2 of 0.6 using SMA and bark endmembers from image and reference libraries. Fraction of red attack, with a possible link to increased crown fire risk, was estimated with an r2 of 0.45.

  9. The battle for Yellowstone: Morality and the  sacred roots of environmental conflict, by Justin Farrell

    Science.gov (United States)

    John Schelhas

    2017-01-01

    A growing number of intractable environmental conflicts involving interest groups with deeply held beliefs are resisting resolution in spite of extensive scientific analysis and legal and bureaucratic attention. Justin Farrell addresses three such conflicts in the Greater Yellowstone Ecosystem (GYE) as moral and spiritual conflicts, each uniquely animated by history,...

  10. Opportunity's Surroundings on Sol 1818 (Vertical)

    Science.gov (United States)

    2009-01-01

    NASA's Mars Exploration Rover Opportunity used its navigation camera to take the images combined into this full-circle view of the rover's surroundings during the 1,818th Martian day, or sol, of Opportunity's surface mission (March 5, 2009). South is at the center; north at both ends. This view is presented as a vertical projection with geometric seam correction. North is at the top. The rover had driven 80.3 meters (263 feet) southward earlier on that sol. Tracks from the drive recede northward in this view. The terrain in this portion of Mars' Meridiani Planum region includes dark-toned sand ripples and lighter-toned bedrock.

  11. Opportunity's Surroundings on Sol 1818 (Polar)

    Science.gov (United States)

    2009-01-01

    NASA's Mars Exploration Rover Opportunity used its navigation camera to take the images combined into this full-circle view of the rover's surroundings during the 1,818th Martian day, or sol, of Opportunity's surface mission (March 5, 2009). South is at the center; north at both ends. This view is presented as a polar projection with geometric seam correction. North is at the top. The rover had driven 80.3 meters (263 feet) southward earlier on that sol. Tracks from the drive recede northward in this view. The terrain in this portion of Mars' Meridiani Planum region includes dark-toned sand ripples and lighter-toned bedrock.

  12. Evaluation of rules to distinguish unique female grizzly bears with cubs in Yellowstone

    Science.gov (United States)

    Schwartz, C.C.; Haroldson, M.A.; Cherry, S.; Keating, K.A.

    2008-01-01

    The United States Fish and Wildlife Service uses counts of unduplicated female grizzly bears (Ursus arctos) with cubs-of-the-year to establish limits of sustainable mortality in the Greater Yellowstone Ecosystem, USA. Sightings are dustered into observations of unique bears based on an empirically derived rule set. The method has never been tested or verified. To evaluate the rule set, we used data from radiocollared females obtained during 1975-2004 to simulate populations under varying densities, distributions, and sighting frequencies. We tested individual rules and rule-set performance, using custom software to apply the rule-set and duster sightings. Results indicated most rules were violated to some degree, and rule-based dustering consistently underestimated the minimum number of females and total population size derived from a nonparametric estimator (Chao2). We conclude that the current rule set returns conservative estimates, but with minor improvements, counts of unduplicated females-with-cubs can serve as a reasonable index of population size useful for establishing annual mortality limits. For the Yellowstone population, the index is more practical and cost-effective than capture-mark-recapture using either DNA hair snagging or aerial surveys with radiomarked bears. The method has useful application in other ecosystems, but we recommend rules used to distinguish unique females be adapted to local conditions and tested.

  13. The YNP Metagenome Project: Environmental Parameters Responsible for Microbial Distribution in the Yellowstone Geothermal Ecosystem

    Directory of Open Access Journals (Sweden)

    William P. Inskeep

    2013-05-01

    Full Text Available The Yellowstone geothermal complex contains over 10,000 diverse geothermal features that host numerous phylogenetically deeply-rooted and poorly understood archaea, bacteria and viruses. Microbial communities in high-temperature environments are generally less diverse than soil, marine, sediment or lake habitats and therefore offer a tremendous opportunity for studying the structure and function of different model microbial communities using environmental metagenomics. One of the broader goals of this study was to establish linkages among microbial distribution, metabolic potential and environmental variables. Twenty geochemically distinct geothermal ecosystems representing a broad spectrum of Yellowstone hot-spring environments were used for metagenomic and geochemical analysis and included approximately equal numbers of: (1 phototrophic mats, (2 ‘filamentous streamer’ communities, and (3 archaeal-dominated sediments. The metagenomes were analyzed using a suite of complementary and integrative bioinformatic tools, including phylogenetic and functional analysis of both individual sequence reads and assemblies of predominant phylotypes. This volume identifies major environmental determinants of a large number of thermophilic microbial lineages, many of which have not been fully described in the literature nor previously cultivated to enable functional and genomic analyses. Moreover, protein family abundance comparisons and in-depth analyses of specific genes and metabolic pathways relevant to these hot-spring environments reveal hallmark signatures of metabolic capabilities that parallel the distribution of phylotypes across specific types of geochemical environments.

  14. Rapid heterogeneous assembly of multiple magma reservoirs prior to Yellowstone supereruptions.

    Science.gov (United States)

    Wotzlaw, Jörn-Frederik; Bindeman, Ilya N; Stern, Richard A; D'Abzac, Francois-Xavier; Schaltegger, Urs

    2015-09-10

    Large-volume caldera-forming eruptions of silicic magmas are an important feature of continental volcanism. The timescales and mechanisms of assembly of the magma reservoirs that feed such eruptions as well as the durations and physical conditions of upper-crustal storage remain highly debated topics in volcanology. Here we explore a comprehensive data set of isotopic (O, Hf) and chemical proxies in precisely U-Pb dated zircon crystals from all caldera-forming eruptions of Yellowstone supervolcano. Analysed zircons record rapid assembly of multiple magma reservoirs by repeated injections of isotopically heterogeneous magma batches and short pre-eruption storage times of 10(3) to 10(4) years. Decoupled oxygen-hafnium isotope systematics suggest a complex source for these magmas involving variable amounts of differentiated mantle-derived melt, Archean crust and hydrothermally altered shallow-crustal rocks. These data demonstrate that complex magma reservoirs with multiple sub-chambers are a common feature of rift- and hotspot related supervolcanoes. The short duration of reservoir assembly documents rapid crustal remelting and two to three orders of magnitude higher magma production rates beneath Yellowstone compared to continental arc volcanoes. The short pre-eruption storage times further suggest that the detection of voluminous reservoirs of eruptible magma beneath active supervolcanoes may only be possible prior to an impending eruption.

  15. Yersinia enterocolitica: an unlikely cause of positive brucellosis tests in greater yellowstone ecosystem bison (Bison bison).

    Science.gov (United States)

    See, Wade; Edwards, William H; Dauwalter, Stacey; Almendra, Claudia; Kardos, Martin D; Lowell, Jennifer L; Wallen, Rick; Cain, Steven L; Holben, William E; Luikart, Gordon

    2012-07-01

    Yersinia enterocolitica serotype O:9 has identical O-antigens to those of Brucella abortus and has apparently caused false-positive reactions in numerous brucellosis serologic tests in elk (Cervus canadensis) from southwest Montana. We investigated whether a similar phenomenon was occurring in brucellosis antibody-positive bison (Bison bison) using Y. enterocolitica culturing techniques and multiplex PCR of four diagnostic loci. Feces from 53 Yellowstone bison culled from the population and 113 free-roaming bison from throughout the Greater Yellowstone Ecosystem (GYE) were tested. Yersinia enterocolitica O:9 was not detected in any of 53 the bison samples collected at slaughter facilities or in any of the 113 fecal samples from free-ranging bison. One other Y. enterocolitica serotype was isolated; however, it is not known to cause cross-reaction on B. abortus serologic assays because it lacks the perosamine synthetase gene and thus the O-antigens. These findings suggest that Y. enterocolitica O:9 cross-reactivity with B. abortus antigens is unlikely to have been a cause of false-positive serology tests in GYE bison and that Y. enterocolitica prevalence was low in bison in the GYE during this study.

  16. Crust Structure Data of Seas Surrounding Turkey

    International Nuclear Information System (INIS)

    Maden, N.; Gelisli, K.

    2007-01-01

    Black Sea, Aegean, Mediterranean and Marmara Sea, which surround the Turkey, have not been examined with respect to the Geological, Geophysical and other natural sciences sufficiently. In fact, it is not attach importance the Turkish seas adequately and abandoned with respect to the scientific researches. The most important reason of this situation is the lack of the education of the Marine Sciences in the Turkish Universities. In this study, it is tried to construct a crustal structure data base of the surrounding seas of the Turkey by collecting crustal structure data sets done by different authors in different times so far. The data acquired in the base are collected from different data base sources by dragging. The Moho depth in the eastern and western basin of the Black sea is 22 km and 19 km, respectively. In the Marmara Sea the Moho depth is 24 km. The moho value in the southern Aegean is 20 km, in the northern Aegean the moho depth is 30 km. on the other hand, the moho depth value in the eastern and western basin of the Mediterranean Sea are 15-20 km and 25-30 km, respectively

  17. Opportunity's Surroundings on Sol 1798 (Polar)

    Science.gov (United States)

    2009-01-01

    NASA's Mars Exploration Rover Opportunity used its navigation camera to take the images combined into this 180-degree view of the rover's surroundings during the 1,798th Martian day, or sol, of Opportunity's surface mission (Feb. 13, 2009). North is on top. This view is presented as a polar projection with geometric seam correction. The rover had driven 111 meters (364 feet) southward on the preceding sol. Tracks from that drive recede northward in this view. For scale, the distance between the parallel wheel tracks is about 1 meter (about 40 inches). The terrain in this portion of Mars' Meridiani Planum region includes dark-toned sand ripples and lighter-toned bedrock.

  18. Opportunity's Surroundings After Sol 1820 Drive (Polar)

    Science.gov (United States)

    2009-01-01

    NASA's Mars Exploration Rover Opportunity used its navigation camera to take the images combined into this full-circle view of the rover's surroundings during the 1,820th to 1,822nd Martian days, or sols, of Opportunity's surface mission (March 7 to 9, 2009). This view is presented as a polar projection with geometric seam correction. North is at the top. The rover had driven 20.6 meters toward the northwest on Sol 1820 before beginning to take the frames in this view. Tracks from that drive recede southwestward. For scale, the distance between the parallel wheel tracks is about 1 meter (about 40 inches). The terrain in this portion of Mars' Meridiani Planum region includes dark-toned sand ripples and small exposures of lighter-toned bedrock.

  19. Opportunity's Surroundings on Sol 1798 (Vertical)

    Science.gov (United States)

    2009-01-01

    NASA's Mars Exploration Rover Opportunity used its navigation camera to take the images combined into this 180-degree view of the rover's surroundings during the 1,798th Martian day, or sol, of Opportunity's surface mission (Feb. 13, 2009). North is on top. This view is presented as a vertical projection with geometric seam correction. The rover had driven 111 meters (364 feet) southward on the preceding sol. Tracks from that drive recede northward in this view. For scale, the distance between the parallel wheel tracks is about 1 meter (about 40 inches). The terrain in this portion of Mars' Meridiani Planum region includes dark-toned sand ripples and lighter-toned bedrock.

  20. Opportunity's Surroundings on Sol 1687 (Vertical)

    Science.gov (United States)

    2009-01-01

    NASA's Mars Exploration Rover Opportunity used its navigation camera to take the images combined into this 360-degree view of the rover's surroundings on the 1,687th Martian day, or sol, of its surface mission (Oct. 22, 2008). Opportunity had driven 133 meters (436 feet) that sol, crossing sand ripples up to about 10 centimeters (4 inches) tall. The tracks visible in the foreground are in the east-northeast direction. Opportunity's position on Sol 1687 was about 300 meters southwest of Victoria Crater. The rover was beginning a long trek toward a much larger crater, Endeavour, about 12 kilometers (7 miles) to the southeast. This view is presented as a vertical projection with geometric seam correction.

  1. Opportunity's Surroundings After Sol 1820 Drive (Vertical)

    Science.gov (United States)

    2009-01-01

    NASA's Mars Exploration Rover Opportunity used its navigation camera to take the images combined into this full-circle view of the rover's surroundings during the 1,820th to 1,822nd Martian days, or sols, of Opportunity's surface mission (March 7 to 9, 2009). This view is presented as a vertical projection with geometric seam correction. North is at the top. The rover had driven 20.6 meters toward the northwest on Sol 1820 before beginning to take the frames in this view. Tracks from that drive recede southwestward. For scale, the distance between the parallel wheel tracks is about 1 meter (about 40 inches). The terrain in this portion of Mars' Meridiani Planum region includes dark-toned sand ripples and small exposures of lighter-toned bedrock.

  2. Opportunity's Surroundings on Sol 1687 (Polar)

    Science.gov (United States)

    2009-01-01

    NASA's Mars Exploration Rover Opportunity used its navigation camera to take the images combined into this 360-degree view of the rover's surroundings on the 1,687th Martian day, or sol, of its surface mission (Oct. 22, 2008). Opportunity had driven 133 meters (436 feet) that sol, crossing sand ripples up to about 10 centimeters (4 inches) tall. The tracks visible in the foreground are in the east-northeast direction. Opportunity's position on Sol 1687 was about 300 meters southwest of Victoria Crater. The rover was beginning a long trek toward a much larger crater, Endeavour, about 12 kilometers (7 miles) to the southeast. This view is presented as a polar projection with geometric seam correction.

  3. Opportunity's Surroundings After Sol 1820 Drive

    Science.gov (United States)

    2009-01-01

    NASA's Mars Exploration Rover Opportunity used its navigation camera to take the images combined into this full-circle view of the rover's surroundings during the 1,820th to 1,822nd Martian days, or sols, of Opportunity's surface mission (March 7 to 9, 2009). South is at the center; north at both ends. The rover had driven 20.6 meters toward the northwest on Sol 1820 before beginning to take the frames in this view. Tracks from that drive recede southwestward. For scale, the distance between the parallel wheel tracks is about 1 meter (about 40 inches). The terrain in this portion of Mars' Meridiani Planum region includes dark-toned sand ripples and small exposures of lighter-toned bedrock. This view is presented as a cylindrical projection with geometric seam correction.

  4. Towards Semantic Understanding of Surrounding Vehicular Maneuvers

    DEFF Research Database (Denmark)

    Kristoffersen, Miklas Strøm; Dueholm, Jacob Velling; Satzoda, Ravi K.

    2016-01-01

    This paper proposes the use of multiple low-cost visual sensors to obtain a surround view of the ego-vehicle for semantic understanding. A multi-perspective view will assist the analysis of naturalistic driving studies (NDS), by automating the task of data reduction of the observed sequences...... into events. A user-centric vision-based framework is presented using a vehicle detector and tracker in each separate perspective. Multi-perspective trajectories are estimated and analyzed to extract 14 different events, including potential dangerous behaviors such as overtakes and cut-ins. The system...... is tested on ten sequences of real-world data collected on U. S. highways. The results show the potential use of multiple low-cost visual sensors for semantic understanding around the ego-vehicle....

  5. Lovelock black holes surrounded by quintessence

    Energy Technology Data Exchange (ETDEWEB)

    Ghosh, Sushant G. [University of KwaZulu-Natal, Astrophysics and Cosmology Research Unit, School of Mathematics, Statistics and Computer Science, Durban (South Africa); Centre for Theoretical Physics, Multidisciplinary Centre for Advanced Research and Studies (MCARS), New Delhi (India); Maharaj, Sunil D.; Baboolal, Dharmanand; Lee, Tae-Hun [University of KwaZulu-Natal, Astrophysics and Cosmology Research Unit, School of Mathematics, Statistics and Computer Science, Durban (South Africa)

    2018-02-15

    Lovelock gravity consisting of the dimensionally continued Euler densities is a natural generalization of general relativity to higher dimensions such that equations of motion are still second order, and the theory is free of ghosts. A scalar field with a positive potential that yields an accelerating universe has been termed quintessence. We present exact black hole solutions in D-dimensional Lovelock gravity surrounded by quintessence matter and also perform a detailed thermodynamical study. Further, we find that the mass, entropy and temperature of the black hole are corrected due to the quintessence background. In particular, we find that a phase transition occurs with a divergence of the heat capacity at the critical horizon radius, and that specific heat becomes positive for r{sub h} < r{sub c} allowing the black hole to become thermodynamically stable. (orig.)

  6. Lovelock black holes surrounded by quintessence

    Science.gov (United States)

    Ghosh, Sushant G.; Maharaj, Sunil D.; Baboolal, Dharmanand; Lee, Tae-Hun

    2018-02-01

    Lovelock gravity consisting of the dimensionally continued Euler densities is a natural generalization of general relativity to higher dimensions such that equations of motion are still second order, and the theory is free of ghosts. A scalar field with a positive potential that yields an accelerating universe has been termed quintessence. We present exact black hole solutions in D-dimensional Lovelock gravity surrounded by quintessence matter and also perform a detailed thermodynamical study. Further, we find that the mass, entropy and temperature of the black hole are corrected due to the quintessence background. In particular, we find that a phase transition occurs with a divergence of the heat capacity at the critical horizon radius, and that specific heat becomes positive for r_h

  7. Linking rapid magma reservoir assembly and eruption trigger mechanisms at evolved Yellowstone-type supervolcanoes

    Science.gov (United States)

    Wotzlaw, J.F.; Bindeman, I.N.; Watts, Kathryn E.; Schmitt, A.K.; Caricchi, L.; Schaltegger, U.

    2014-01-01

    The geological record contains evidence of volcanic eruptions that were as much as two orders of magnitude larger than the most voluminous eruption experienced by modern civilizations, the A.D. 1815 Tambora (Indonesia) eruption. Perhaps nowhere on Earth are deposits of such supereruptions more prominent than in the Snake River Plain–Yellowstone Plateau (SRP-YP) volcanic province (northwest United States). While magmatic activity at Yellowstone is still ongoing, the Heise volcanic field in eastern Idaho represents the youngest complete caldera cycle in the SRP-YP, and thus is particularly instructive for current and future volcanic activity at Yellowstone. The Heise caldera cycle culminated 4.5 Ma ago in the eruption of the ∼1800 km3 Kilgore Tuff. Accessory zircons in the Kilgore Tuff display significant intercrystalline and intracrystalline oxygen isotopic heterogeneity, and the vast majority are 18O depleted. This suggests that zircons crystallized from isotopically distinct magma batches that were generated by remelting of subcaldera silicic rocks previously altered by low-δ18O meteoric-hydrothermal fluids. Prior to eruption these magma batches were assembled and homogenized into a single voluminous reservoir. U-Pb geochronology of isotopically diverse zircons using chemical abrasion–isotope dilution–thermal ionization mass spectrometry yielded indistinguishable crystallization ages with a weighted mean 206Pb/238U date of 4.4876 ± 0.0023 Ma (MSWD = 1.5; n = 24). These zircon crystallization ages are also indistinguishable from the sanidine 40Ar/39Ar dates, and thus zircons crystallized close to eruption. This requires that shallow crustal melting, assembly of isolated batches into a supervolcanic magma reservoir, homogenization, and eruption occurred extremely rapidly, within the resolution of our geochronology (103–104 yr). The crystal-scale image of the reservoir configuration, with several isolated magma batches, is very similar to the

  8. Evaluation of ML-MC as a Depth Discriminant in Yellowstone, USA and Italy

    Science.gov (United States)

    Li, Z.; Koper, K. D.; Burlacu, R.; Sun, D.; D'Amico, S.

    2017-12-01

    Recent work has shown that the difference between two magnitude scales, ML (local Richter magnitude) and MC (coda/duration magnitude), acts as a depth discriminant in Utah. Shallow seismic sources, such as mining induced earthquakes and explosions, have strongly negative ML-MC values, while deeper tectonic earthquakes have ML-MC values near zero. These observations imply that ML-MC might be effective at discriminating small explosions from deeper natural earthquakes at local distances. In this work, we examine seismicity catalogs for the Yellowstone region and Italy to determine if ML-MCacts as a depth discriminant in these regions as well. We identified 4,780 earthquakes that occurred in the Yellowstone region between Sept. 24, 1994 and March 31, 2017 for which both ML and MC were calculated. The ML-MC distribution is well described by a Gaussian function with a mean of 0.102 and a standard deviation of 0.326. We selected a subset of these events with accurate depths and determined mean ML-MC values in various depth bins. An event depth was considered accurate if the formal depth error was less than 2 km and either (1) the nearest station was within one focal depth or (2) the distance to the nearest station was smaller than the bin size. We find that ML-MC decreases as event depths become shallower than about 10 km. Similar to the results for Utah, the decrease is statistically significant and is robust with respect to small changes in bin size and the criteria used to define accurate depths. We used a similar process to evaluate whether ML-MC was a function of source depth for 63,555 earthquakes that occurred between April 16, 2005 and April 30, 2012 in Italy. The ML-MC values in Italy are also well described by a normal distribution, with a mean of -0.477 and standard deviation of 0.315. We again find a statistically significant decrease in ML-MC for shallow earthquakes. In contrast to the Yellowstone results, for Italy ML-MC decreases at a nearly constant rate

  9. Infecciones parasitarias del coyote, Canis latrans (Carnivora: Canidae en un Parque Nacional y una zona agrícola en Costa Rica Parasitic infections of coyote, Canis latrans (Carnivora: Canidae in a Costa Rican National Park and a surrounding agricultural area

    Directory of Open Access Journals (Sweden)

    Carmen Niehaus

    2012-06-01

    Full Text Available Conforme las poblaciones humanas se expanden hacia los hábitats silvestres con sus mascotas y ganado, el potencial de transmisión de enfermedades hacia los animales silvestres -y viceversa- aumenta, y hace necesario identificar interacciones zoonóticas potenciales. Los cánidos domésticos y silvestres pueden funcionar como reservorios o diseminadores de enfermedades infecciosas (se incluyen parásitos, por lo que el coyote (Canis latrans puede también servir como indicador de la salud ecológica. Asimismo, se estudiaron los parásitos de 209 muestras de heces de coyotes en una zona mixta de área silvestre protegida y campo agrícola del Parque Nacional Volcán Irazú (PNVI en Costa Rica. La recolección fue realizada mensualmente durante un año en tres sub-áreas denominadas: Irazú (la más cercana al volcán, papales (por el cultivo de papas, y Prusia (un sector del PNVI. Entonces, se empleó examen directo y concentración mecánica, se obtuvo 36.84% de muestras positivas por al menos un helminto. La presencia de parásitos fue muy similar para ambos sectores boscosos del PNVI (33.3% en Prusia y 37.4% en Irazú, pero contrastó con el 63.63% observado en los papales. También, se identificaron uncinarias (probablemente Ancylostoma caninum, estrongilidios (posiblemente Strongyloides sp., Toxocara canis, Trichuris sp. y Taenia pisiformis, así como Hymenolepis diminuta, probablemente un parásito espurio proveniente de roedores ingeridos por los coyotes. Se comenta la importancia de estos primeros hallazgos y se concluye que las estaciones seca y lluviosa influyen en la presencia de los parásitos.Parasitic infections of coyote, Canis latrans (Carnivora: Canidae in a Costa Rican National Park and a surrounding agricultural area. As human populations expand into wild habitats with their pets and livestock, the potential spread of disease to wildlife or vice versa increases. Because, wild and domestic canids may pose as reservoirs or

  10. A spatially explicit model for an Allee effect: why wolves recolonize so slowly in Greater Yellowstone.

    Science.gov (United States)

    Hurford, Amy; Hebblewhite, Mark; Lewis, Mark A

    2006-11-01

    A reduced probability of finding mates at low densities is a frequently hypothesized mechanism for a component Allee effect. At low densities dispersers are less likely to find mates and establish new breeding units. However, many mathematical models for an Allee effect do not make a distinction between breeding group establishment and subsequent population growth. Our objective is to derive a spatially explicit mathematical model, where dispersers have a reduced probability of finding mates at low densities, and parameterize the model for wolf recolonization in the Greater Yellowstone Ecosystem (GYE). In this model, only the probability of establishing new breeding units is influenced by the reduced probability of finding mates at low densities. We analytically and numerically solve the model to determine the effect of a decreased probability in finding mates at low densities on population spread rate and density. Our results suggest that a reduced probability of finding mates at low densities may slow recolonization rate.

  11. Disparate stakeholder management: the case of elk and bison feeding in southern Greater Yellowstone

    Science.gov (United States)

    Koontz, Lynne; Hoag, Dana; DeLong, Don

    2012-01-01

    For resource decisions to make the most possible progress toward achieving agency mandates, managers must work with stakeholders and may need to at least partially accommodate some of their key underlying interests. To accommodate stakeholder interests, while also substantively working toward fulfilling legal mandates, managers must understand the sociopolitical factors that influence the decision-making process. We coin the phrase disparate stakeholder management (DSM) to describe situations with disparate stakeholders and disparate management solutions. A DSM approach (DSMA) requires decision makers to combine concepts from many sciences, thus releasing them from disciplinary bonds that often constrain innovation and effectiveness. We combined three distinct approaches to develop a DSMA that assisted in developing a comprehensive range of elk and bison management alternatives in the Southern Greater Yellowstone Area. The DSMA illustrated the extent of compromise between meeting legal agency mandates and accommodating the preferences of certain stakeholder groups.

  12. Opportunity's Surroundings on Sol 1798 (Stereo)

    Science.gov (United States)

    2009-01-01

    [figure removed for brevity, see original site] Left-eye view of a color stereo pair for PIA11850 [figure removed for brevity, see original site] Right-eye view of a color stereo pair for PIA11850 NASA's Mars Exploration Rover Opportunity used its navigation camera to take the images combined into this stereo 180-degree view of the rover's surroundings during the 1,798th Martian day, or sol, of Opportunity's surface mission (Feb. 13, 2009). North is on top. This view combines images from the left-eye and right-eye sides of the navigation camera. It appears three-dimensional when viewed through red-blue glasses with the red lens on the left. The rover had driven 111 meters (364 feet) southward on the preceding sol. Tracks from that drive recede northward in this view. For scale, the distance between the parallel wheel tracks is about 1 meter (about 40 inches). The terrain in this portion of Mars' Meridiani Planum region includes dark-toned sand ripples and lighter-toned bedrock. This view is presented as a cylindrical-perspective projection with geometric seam correction.

  13. Opportunity's Surroundings on Sol 1818 (Stereo)

    Science.gov (United States)

    2009-01-01

    [figure removed for brevity, see original site] Left-eye view of a color stereo pair for PIA11846 [figure removed for brevity, see original site] Right-eye view of a color stereo pair for PIA11846 NASA's Mars Exploration Rover Opportunity used its navigation camera to take the images combined into this full-circle view of the rover's surroundings during the 1,818th Martian day, or sol, of Opportunity's surface mission (March 5, 2009). South is at the center; north at both ends. This view combines images from the left-eye and right-eye sides of the navigation camera. It appears three-dimensional when viewed through red-blue glasses with the red lens on the left. The rover had driven 80.3 meters (263 feet) southward earlier on that sol. Tracks from the drive recede northward in this view. The terrain in this portion of Mars' Meridiani Planum region includes dark-toned sand ripples and lighter-toned bedrock. This view is presented as a cylindrical-perspective projection with geometric seam correction.

  14. Opportunity's Surroundings on Sol 1687 (Stereo)

    Science.gov (United States)

    2009-01-01

    [figure removed for brevity, see original site] Left-eye view of a color stereo pair for PIA11739 [figure removed for brevity, see original site] Right-eye view of a color stereo pair for PIA11739 NASA's Mars Exploration Rover Opportunity used its navigation camera to take the images combined into this stereo, 360-degree view of the rover's surroundings on the 1,687th Martian day, or sol, of its surface mission (Oct. 22, 2008). The view appears three-dimensional when viewed through red-blue glasses. Opportunity had driven 133 meters (436 feet) that sol, crossing sand ripples up to about 10 centimeters (4 inches) tall. The tracks visible in the foreground are in the east-northeast direction. Opportunity's position on Sol 1687 was about 300 meters southwest of Victoria Crater. The rover was beginning a long trek toward a much larger crater, Endeavour, about 12 kilometers (7 miles) to the southeast. This panorama combines right-eye and left-eye views presented as cylindrical-perspective projections with geometric seam correction.

  15. Opportunity's Surroundings After Sol 1820 Drive (Stereo)

    Science.gov (United States)

    2009-01-01

    [figure removed for brevity, see original site] Left-eye view of a color stereo pair for PIA11841 [figure removed for brevity, see original site] Right-eye view of a color stereo pair for PIA11841 NASA's Mars Exploration Rover Opportunity used its navigation camera to take the images combined into this full-circle view of the rover's surroundings during the 1,820th to 1,822nd Martian days, or sols, of Opportunity's surface mission (March 7 to 9, 2009). This view combines images from the left-eye and right-eye sides of the navigation camera. It appears three-dimensional when viewed through red-blue glasses with the red lens on the left. The rover had driven 20.6 meters toward the northwest on Sol 1820 before beginning to take the frames in this view. Tracks from that drive recede southwestward. For scale, the distance between the parallel wheel tracks is about 1 meter (about 40 inches). The terrain in this portion of Mars' Meridiani Planum region includes dark-toned sand ripples and small exposures of lighter-toned bedrock. This view is presented as a cylindrical-perspective projection with geometric seam correction.

  16. The lithosphere-asthenosphere: Italy and surroundings

    International Nuclear Information System (INIS)

    Panza, G.F.; Aoudia, A.; Pontevivo, A.; Chimera, G.; Raykova, R.

    2003-02-01

    The velocity-depth distribution of the lithosphere-asthenosphere in the Italian region and surroundings is imaged, with a lateral resolution of about 100 km, by surface wave velocity tomography and non-linear inversion. Maps of the Moho depth, of the thickness of the lithosphere and of the shear-wave velocities, down to depths of 200 km and more, are constructed. A mantle wedge, identified in the uppermost mantle along the Apennines and the Calabrian Arc, underlies the principal recent volcanoes, and partial melting can be relevant in this part of the uppermost mantle. In Calabria a lithospheric doubling is seen, in connection with the subduction of the Ionian lithosphere. The asthenosphere is shallow in the Southern Tyrrhenian Sea. High velocity bodies, cutting the asthenosphere, outline the Adria-lonian subduction in the Tyrrhenian Sea and the deep-reaching lithospheric root in the Western Alps. Less deep lithospheric roots are seen in the Central Apennines. The lithosphere-asthenosphere properties delineate a differentiation between the northern and the southern sectors of the Adriatic Sea, likely attesting the fragmentation of Adria. (author)

  17. The lithosphere-asthenosphere Italy and surroundings

    CERN Document Server

    Panza, G F; Chimera, G; Pontevivo, A; Raykova, R

    2003-01-01

    The velocity-depth distribution of the lithosphere-asthenosphere in the Italian region and surroundings is imaged, with a lateral resolution of about 100 km, by surface wave velocity tomography and non-linear inversion. Maps of the Moho depth, of the thickness of the lithosphere and of the shear-wave velocities, down to depths of 200 km and more, are constructed. A mantle wedge, identified in the uppermost mantle along the Apennines and the Calabrian Arc, underlies the principal recent volcanoes, and partial melting can be relevant in this part of the uppermost mantle. In Calabria a lithospheric doubling is seen, in connection with the subduction of the Ionian lithosphere. The asthenosphere is shallow in the Southern Tyrrhenian Sea. High velocity bodies, cutting the asthenosphere, outline the Adria-lonian subduction in the Tyrrhenian Sea and the deep-reaching lithospheric root in the Western Alps. Less deep lithospheric roots are seen in the Central Apennines. The lithosphere-asthenosphere properties delineat...

  18. INTERSTELLAR MAGNETIC FIELD SURROUNDING THE HELIOPAUSE

    International Nuclear Information System (INIS)

    Whang, Y. C.

    2010-01-01

    This paper presents a three-dimensional analytical solution, in the limit of very low plasma β-ratio, for the distortion of the interstellar magnetic field surrounding the heliopause. The solution is obtained using a line dipole method that is the integration of point dipole along a semi-infinite line; it represents the magnetic field caused by the presence of the heliopause. The solution allows the variation of the undisturbed magnetic field at any inclination angle. The heliosphere is considered as having blunt-nosed geometry on the upwind side and it asymptotically approaches a cylindrical geometry having an open exit for the continuous outflow of the solar wind on the downwind side. The heliopause is treated as a magnetohydrodynamic tangential discontinuity; the interstellar magnetic field lines at the boundary are tangential to the heliopause. The interstellar magnetic field is substantially distorted due to the presence of the heliopause. The solution shows the draping of the field lines around the heliopause. The magnetic field strength varies substantially near the surface of the heliopause. The effect on the magnetic field due to the presence of the heliopause penetrates very deep into the interstellar space; the depth of penetration is of the same order of magnitude as the scale length of the heliosphere.

  19. Body condition and pregnancy in northern Yellowstone elk: evidence for predation risk effects?

    Science.gov (United States)

    White, P J; Garrott, Robert A; Hamlin, Kenneth L; Cook, Rachel C; Cook, John G; Cunningham, Julie A

    2011-01-01

    S. Creel et al. reported a negative correlation between fecal progesterone concentrations and elk:wolf ratios in greater Yellowstone elk (Cervus elaphus) herds and interpreted this correlation as evidence that pregnancy rates of elk decreased substantially in the presence of wolves (Canis lupus). Apparently, the hypothesized mechanism is that decreased forage intake reduces body condition and either results in elk failing to conceive during the autumn rut or elk losing the fetus during winter. We tested this hypothesis by comparing age-specific body condition (percentage ingesta-free body fat) and pregnancy rates for northern Yellowstone elk, one of the herds sampled by Creel et al., before (1962-1968) and after (2000-2006) wolf restoration using indices developed and calibrated for Rocky Mountain elk. Mean age-adjusted percentage body fat of female elk was similarly high in both periods (9.0%-0.9% pre-wolf; 8.9%-0.8% post-wolf). Estimated pregnancy rates (proportion of females that were pregnant) were 0.91 pre-wolf and 0.87 post-wolf for 4-9 year-old elk (95% CI on difference = -0.15 to 0.03, P = 0.46) and 0.64 pre-wolf and 0.78 post-wolf for elk > 9 years old (95% CI on difference = -0.01 to 0.27, P = 0.06). Thus, there was little evidence in these data to support strong effects of wolf presence on elk pregnancy. We caution that multiple lines of evidence and/or strong validation should be brought to bear before relying on indirect measures of how predators affect pregnancy rates.

  20. Plumbing the depths of Yellowstone's hydrothermal system from helicopter magnetic and electromagnetic data

    Science.gov (United States)

    Finn, C.; Bedrosian, P.; Holbrook, W. S.; Auken, E.; Lowenstern, J. B.; Hurwitz, S.; Sims, K. W. W.; Carr, B.; Dickey, K.

    2017-12-01

    Although Yellowstone's iconic hydrothermal systems and lava flows are well mapped at the surface, their groundwater flow systems and thickness are almost completely unknown. In order to track the geophysical signatures of geysers, hot springs, mud pots, steam vents, hydrothermal explosion craters and lava flows at depths to hundreds of meters, we collected helicopter electromagnetic and magnetic (HEM) data. The data cover significant portions of the caldera including a majority of the known thermal areas. HEM data constrain electrical resistivity which is sensitive to groundwater salinity and temperature, phase distribution (liquid-vapor), and clay formed during chemical alteration of rocks. The magnetic data are sensitive to variations in the magnetization of lava flows, faults and hydrothermal alteration. The combination of electromagnetic and magnetic data is ideal for mapping zones of cold fresh water, hot saline water, steam, clay, and altered and unaltered rock. Preliminary inversion of the HEM data indicates very low resistivity directly beneath the northern part of Yellowstone Lake, intersecting with the lake bottom in close correspondence with mapped vents, fractures and hydrothermal explosion craters and are also associated with magnetic lows. Coincident resistivity and magnetic lows unassociated with mapped alteration occur, for example, along the southeast edge of the Mallard Lake dome and along the northeastern edge of Sour Creek Dome, suggesting the presence of buried alteration. Low resistivities unassociated with magnetic lows may relate to hot and/or saline groundwater or thin (<50 m) layers of early lake sediments to which the magnetic data are insensitive. Resistivity and magnetic lows follow interpreted caldera boundaries in places, yet deviate in others. In the Norris-Mammoth Corridor, NNE-SSW trending linear resistivity and magnetic lows align with mapped faults. This pattern of coincident resistivity and magnetic lows may reflect fractures

  1. Grizzly bear-human conflicts in the Yellowstone Ecosystem, 1992-2000

    Science.gov (United States)

    Gunther, K.A.; Haroldson, M.A.; Cain, S.L.; Copeland, J.; Frey, K.; Schwartz, C.C.

    2004-01-01

    For many years, the primary strategy for managing grizzly bears (Ursus arctos) that came into conflict with humans in the Greater Yellowstone Ecosystem (GYE) was to capture and translocate the offending bears away from conflict sites. Translocation usually only temporarily alleviated the problems and most often did not result in long-term solutions. Wildlife managers needed to be able to predict the causes, types, locations, and trends of conflicts to more efficiently allocate resources for pro-active rather than reactive management actions. To address this need, we recorded all grizzly bear-human conflicts reported in the GYE during 1992-2000. We analyzed trends in conflicts over time (increasing or decreasing), geographic location on macro- (inside or outside of the designated Yellowstone Grizzly Bear Recovery Zone [YGBRZ]) and micro- (geographic location) scales, land ownership (public or private), and relationship to the seasonal availability of bear foods. We recorded 995 grizzly bear-human conflicts in the GYE. Fifty-three percent of the conflicts occurred outside and 47% inside the YGBRZ boundary. Fifty-nine percent of the conflicts occurred on public and 41% on private land. Incidents of bears damaging property and obtaining anthropogenic foods were inversely correlated to the abundance of naturally occurring bear foods. Livestock depredations occurred independent of the availability of bear foods. To further aid in prioritizing management strategies to reduce conflicts, we also analyzed conflicts in relation to subsequent human-caused grizzly bear mortality. There were 74 human-caused grizzly bear mortalities during the study, primarily from killing bears in defense of life and property (43%) and management removal of bears involved in bear-human conflicts (28%). Other sources of human-caused mortality included illegal kills, electrocution by downed power-lines, mistaken identification by American black bear (Ursus americanus) hunters, and vehicle strikes

  2. Anomalous shear wave delays and surface wave velocities at Yellowstone Caldera, Wyoming

    International Nuclear Information System (INIS)

    Daniel, R.G.; Boore, D.M.

    1982-01-01

    To investigate the effects of a geothermal area on the propagation of intermediate-period (1--30 s) teleseismic body waves and surface waves, a specially designed portable seismograph system was operated in Yellowstone Caldera, Wyoming. Travel time residuals, relative to a station outside the caldera, of up to 2 s for compressional phases are in agreement with short-period residuals for P phases measured by other investigators. Travel time delays for shear arrivals in the intermediate-period band range from 2 to 9 s and decrease with increasing dT/dΔ. Measured Rayleigh wave phase velocities are extremely low, ranging from 3.2 km/s at 27-s period to 2.0 km/s at 7-s period; the estimated uncertainty associated with these values is 15%. We propose a model for compressional and shear velocities and Poisson's ratio beneath the Yellowstone caldera which fits the teleseismic body and surface wave data: it consists of a highly anomalous crust with an average shear velocity of 3.0 km/s overlying an upper mantle with average velocity of 4.1 km/s. The high average value of Poisson's ratio in the crust (0.34) suggests the presence of fluids there; Poisson's ratio in the mantle between 40 and approximately 200 km is more nearly normal (0.29) than in the crust. A discrepancy between normal values of Poisson's ratio in the crust calculated from short-period data and high values calculated from teleseismic data can be resolved by postulating a viscoelastic crustal model with frequency-dependent shear velocity and attenuation

  3. Body and diet composition of sympatric black and grizzly bears in the Greater Yellowstone Ecosystem

    Science.gov (United States)

    Schwartz, Charles C.; Fortin, Jennifer K.; Teisberg, Justin E.; Haroldson, Mark A.; Servheen, Christopher; Robbins, Charles T.; van Manen, Frank T.

    2013-01-01

    The Greater Yellowstone Ecosystem (GYE) has experienced changes in the distribution and availability of grizzly bear (Ursus arctos) food resources in recent decades. The decline of ungulates, fish, and whitebark pine seeds (Pinus albicaulis) has prompted questions regarding their ability to adapt. We examined body composition and diet of grizzly bears using bioelectrical impedance and stable isotopes to determine if 1) we can detect a change in diet quality associated with the decline in either ungulates or whitebark pine, and 2) the combined decline in ungulates, fish, and pine seeds resulted in a change in grizzly bear carrying capacity in the GYE. We contrasted body fat and mass in grizzly bears with a potential competitor, the American black bear (Ursus americanus), to address these questions. Grizzly bears assimilated more meat into their diet and were in better body condition than black bears throughout the study period, indicating the decline in ungulate resources did not affect grizzly bears more than black bears. We also found no difference in autumn fat levels in grizzly bears in years of good or poor pine seed production, and stable isotope analyses revealed this was primarily a function of switching to meat resources during poor seed-producing years. This dietary plasticity was consistent over the course of our study. We did not detect an overall downward trend in either body mass or the fraction of meat assimilated into the diet by grizzly bears over the past decade, but we did detect a downward trend in percent body fat in adult female grizzly bears after 2006. Whether this decline is an artifact of small sample size or due to the population reaching the ecological carrying capacity of the Yellowstone ecosystem warrants further investigation.

  4. A perfect storm: multiple stressors interact to drive postfire regeneration failure of lodgepole pine and Douglas-fir forests in Yellowstone

    Science.gov (United States)

    Hansen, W. D.; Braziunas, K. H.; Rammer, W.; Seidl, R.; Turner, M. G.

    2017-12-01

    Twenty-first century forests will experience increased stress as environmental conditions and disturbance regimes change. Whether forests retain their structure or transitions to alternate states, particularly when affected by multiple stressors, remains unresolved. Subalpine forests in Yellowstone National Park, WY experience large severe wildfires, and postfire-tree regeneration is necessary to assure resilience. Drying is projected, causing frequent larger wildfires that could reduce seed supply and drought that could constrain postfire-seedling establishment. We asked what combinations of warming-drying conditions, increased fire frequency, and increased burned-patch size cause postfire tree-regeneration failure in Yellowstone? We conducted a simulation experiment to identify combinations of fire frequency, fire size, postfire climate, substrate type, and elevation where lodgepole-pine and Douglas-fir regeneration failed. We expected postfire densities to be reduced if burned-patch sizes exceeded effective dispersal distance, sequential fires burned before trees reached reproductive maturity, or drought occurred after fire. We also expected regeneration failure only where multiple stressors occurred simultaneously at low elevation or on poor substrates.Douglas-fir stands were most vulnerable to regeneration failure. 98% of simulated Douglas-fir stands located in the middle of large burned patches failed to regenerate 30 years post fire. Lodgepole-pine stands in the middle of large burned patches failed to regenerate if they were also located at low elevations (93%) or at higher elevations on soils with poor water retention (73%). Stands of serotinous lodgepole (i.e., trees with closed cones that open when heated) also failed to regenerate if fire recurred before trees were reproductively mature (82%). Drought constrained postfire regeneration, yet, enhanced establishment due to release from cold-temperatures during mid-to-late 21st century often outweighed

  5. Hurricane Ivan Aerial Photography: High-Resolution Imagery of the Florida Panhandle and Surrounding Regions After Landfall

    Data.gov (United States)

    National Oceanic and Atmospheric Administration, Department of Commerce — The imagery posted on this site is of the Florida panhandle and surrounding regions after Hurricane Ivan made landfall. The regions photographed range from Gulf...

  6. Marine Resource Survey in waters surrounding Guam and the Commonwealth of the Northern Mariana Islands (SE1002, EK60)

    Data.gov (United States)

    National Oceanic and Atmospheric Administration, Department of Commerce — The goals of the cruise were to collect a variety of data to assess the status of marine resources in waters surrounding Guam and CNMI. Marine resource habitats were...

  7. Understanding community norms surrounding tobacco sales.

    Directory of Open Access Journals (Sweden)

    Patricia A McDaniel

    Full Text Available In the US, denormalizing tobacco use is key to tobacco control; less attention has been paid to denormalizing tobacco sales. However, some localities have placed limits on the number and type of retailers who may sell tobacco, and some retailers have abandoned tobacco sales voluntarily. Understanding community norms surrounding tobacco sales may help accelerate tobacco denormalization.We conducted 15 focus groups with customers of California, New York, and Ohio retailers who had voluntarily discontinued tobacco sales to examine normative assumptions about where cigarettes should or should not be sold, voluntary decisions to discontinue tobacco sales, and government limits on such sales.Groups in all three states generally agreed that grocery stores that sold healthy products should not sell tobacco; California groups saw pharmacies similarly, while this was a minority opinion in the other two states. Convenience stores were regarded as a natural place to sell tobacco. In each state, it was regarded as normal and commendable for some stores to want to stop selling tobacco, although few participants could imagine convenience stores doing so. Views on government's role in setting limits on tobacco sales varied, with California and New York participants generally expressing support for restrictions, and Ohio participants expressing opposition. However, even those who expressed opposition did not approve of tobacco sales in all possible venues. Banning tobacco sales entirely was not yet normative.Limiting the ubiquitous availability of tobacco sales is key to ending the tobacco epidemic. Some limits on tobacco sales appear to be normative from the perspective of community members; it may be possible to shift norms further by problematizing the ubiquitous presence of cigarettes and drawing connections to other products already subject to restrictions.

  8. Understanding community norms surrounding tobacco sales.

    Science.gov (United States)

    McDaniel, Patricia A; Malone, Ruth E

    2014-01-01

    In the US, denormalizing tobacco use is key to tobacco control; less attention has been paid to denormalizing tobacco sales. However, some localities have placed limits on the number and type of retailers who may sell tobacco, and some retailers have abandoned tobacco sales voluntarily. Understanding community norms surrounding tobacco sales may help accelerate tobacco denormalization. We conducted 15 focus groups with customers of California, New York, and Ohio retailers who had voluntarily discontinued tobacco sales to examine normative assumptions about where cigarettes should or should not be sold, voluntary decisions to discontinue tobacco sales, and government limits on such sales. Groups in all three states generally agreed that grocery stores that sold healthy products should not sell tobacco; California groups saw pharmacies similarly, while this was a minority opinion in the other two states. Convenience stores were regarded as a natural place to sell tobacco. In each state, it was regarded as normal and commendable for some stores to want to stop selling tobacco, although few participants could imagine convenience stores doing so. Views on government's role in setting limits on tobacco sales varied, with California and New York participants generally expressing support for restrictions, and Ohio participants expressing opposition. However, even those who expressed opposition did not approve of tobacco sales in all possible venues. Banning tobacco sales entirely was not yet normative. Limiting the ubiquitous availability of tobacco sales is key to ending the tobacco epidemic. Some limits on tobacco sales appear to be normative from the perspective of community members; it may be possible to shift norms further by problematizing the ubiquitous presence of cigarettes and drawing connections to other products already subject to restrictions.

  9. Events surrounding the construction of Nimrod

    International Nuclear Information System (INIS)

    Wilkinson, D.

    1979-01-01

    Using quotations from many letters, memoranda and notes of meetings the author gives a personal rundown of the attitudes prevailing among nuclear physicists both in universities and in the UKAEA during the 1950's concerning the vexed question of the provision of expensive facilities for high energy particle research in the UK. The outcome of these discussions was the foundation in 1957 of the National Institute for Research in Nuclear Science. (U.K.)

  10. Influence of whitebark pine decline on fall habitat use and movements of grizzly bears in the Greater Yellowstone Ecosystem

    OpenAIRE

    Costello, Cecily M; van Manen, Frank T; Haroldson, Mark A; Ebinger, Michael R; Cain, Steven L; Gunther, Kerry A; Bjornlie, Daniel D

    2014-01-01

    When abundant, seeds of the high-elevation whitebark pine (WBP; Pinus albicaulis) are an important fall food for grizzly bears (Ursus arctos) in the Greater Yellowstone Ecosystem. Rates of bear mortality and bear/human conflicts have been inversely associated with WBP productivity. Recently, mountain pine beetles (Dendroctonus ponderosae) have killed many cone-producing WBP trees. We used fall (15 August–30 September) Global Positioning System locations from 89 bear years to investigate tempo...

  11. The duration of a Yellowstone super-eruption cycle and implications for the age of the Olduvai subchron

    Science.gov (United States)

    Rivera, Tiffany A.; Darata, Rachel; Lippert, Peter C.; Jicha, Brian R.; Schmitz, Mark D.

    2017-12-01

    Small-volume rhyolitic eruptions preceding and following a caldera-forming eruption can provide insights into the tempo of eruption cycles and timing of magmatic recharge. In this contribution, high-precision 40Ar/39Ar eruption ages were obtained on the three effusive eruptions bracketing the Huckleberry Ridge Tuff, which comprise Yellowstone's first volcanic cycle. These dates are supplemented with detailed paleomagnetic and rock magnetic analyses to resolve discrepancies with previous reported stratigraphy. The Huckleberry Ridge Tuff (2.08 Ma) was preceded by an eruption at 2.14 Ma, and followed by eruptions at 1.98 and 1.95 Ma, all of which occurred during four distinct periods of geomagnetic instability within the Matuyama chron. The first volcanic cycle of Yellowstone has now been constrained to within a 200 kyr timespan, or half of the previously proposed duration, and similar to the duration of volcanic activity for caldera-forming systems in the Jemez Volcanic Field. The maximum duration for magmatic recharge for the first Yellowstone volcanic cycle is no greater than 100 kyr, and likely closer to 40 kyr. Furthermore, the combined 40Ar/39Ar eruption ages and paleomagnetic results provide polarity anchors for the Pre-Olduvai excursion and Olduvai subchron, which are often used as tie-points in studies of early Pleistocene hominin evolution.

  12. Impacts of Artificial Reefs on Surrounding Ecosystems

    Science.gov (United States)

    Manoukian, Sarine

    Artificial reefs are becoming a popular biological and management component in shallow water environments characterized by soft seabed, representing both important marine habitats and tools to manage coastal fisheries and resources. An artificial reef in the marine environment acts as an open system with exchange of material and energy, altering the physical and biological characteristics of the surrounding area. Reef stability will depend on the balance of scour, settlement, and burial resulting from ocean conditions over time. Because of the unstable nature of sediments, they require a detailed and systematic investigation. Acoustic systems like high-frequency multibeam sonar are efficient tools in monitoring the environmental evolution around artificial reefs, whereas water turbidity can limit visual dive and ROV inspections. A high-frequency multibeam echo sounder offers the potential of detecting fine-scale distribution of reef units, providing an unprecedented level of resolution, coverage, and spatial definition. How do artificial reefs change over time in relation to the coastal processes? How accurately does multibeam technology map different typologies of artificial modules of known size and shape? How do artificial reefs affect fish school behavior? What are the limitations of multibeam technology for investigating fish school distribution as well as spatial and temporal changes? This study addresses the above questions and presents results of a new approach for artificial reef seafloor mapping over time, based upon an integrated analysis of multibeam swath bathymetry data and geoscientific information (backscatter data analysis, SCUBA observations, physical oceanographic data, and previous findings on the geology and sedimentation processes, integrated with unpublished data) from Senigallia artificial reef, northwestern Adriatic Sea (Italy) and St. Petersburg Beach Reef, west-central Florida continental shelf. A new approach for observation of fish

  13. Yellowstone-Snake River Plain seismic profilling experiment: Crustal structure of the eastern Snake River Plain

    International Nuclear Information System (INIS)

    Braile, L.W.; Smith, R.B.; Ansorge, J.; Baker, M.R.; Sparlin, M.A.; Prodehl, C.; Schilly, M.M.; Healy, J.H.; Mueller, S.; Olsen, K.H.

    1982-01-01

    Seismic refraction profiles recorded along the eastern Snake River Plain (ESRP) in southeastern Idaho during the 1978 Yellowstone-Snake River Plain cooperative seismic profiling experiment are interpreted to infer the crustal velocity and attenuation (Q-1) structure of the ESRP. Travel-time and synthetic seismogram modeling of a 250 km reversed refraction profile as well as a 100 km detailed profile indicate that the crust of the ESRP is highly anomalous. Approximately 3 to 6 km of volcanic rocks (with some interbedded sediments) overlie an upper-crustal layer (compressional velocity approx. =6.1 km/s) which thins southwestward along the ESRP from a thickness of 10 km near Island Park Caldera to 2 to 3 km beneath the central and southwestern portions of the ESRP. An intermediate-velocity (approx. =6.5 km/s) layer extends from approx. =10 to approx. =20 km depth. a thick (approx. =22 km) lower crust of compressional velocity 6.8 km/s, a total crustall thickness of approx. =42 km, and a P/sub n/ velocity of approx. =7.9 km/s is observed in the ESRP, similar to the western Snake River Plain and the Rocky Mountains Provinces. High attenuation is evident on the amplitude corrected seismic data due to low-Q values in the volcanic rocks (Q/sub p/ = 20 to 200) and throughout the crust (Q/sub p/ = 160 to 300). Based on these characteristics of the crustal structure and volcanic-age progression data, it is suggested that the ESRP has resulted from an intensitive period of intrusion of mantle-derived basaltic magma into the upper crust generating explosive silicic volcanism and associated regional uplift and caldera collapse. This activity began about 15 m.y. ago in southwestern Idaho and has migrated northeast to its present position at Yellowstone. Subsequent cooling of the intruded upper crust results in the 6.5 km/s velocity intermediate layer. Crustal subsidence and periodic basaltic volcanism as represented by the ESRP complete the sequence of crustal evolution

  14. Volcanism at 1.45 Ma within the Yellowstone Volcanic Field, United States

    Science.gov (United States)

    Rivera, Tiffany A.; Furlong, Ryan; Vincent, Jaime; Gardiner, Stephanie; Jicha, Brian R.; Schmitz, Mark D.; Lippert, Peter C.

    2018-05-01

    Rhyolitic volcanism in the Yellowstone Volcanic Field has spanned over two million years and consisted of both explosive caldera-forming eruptions and smaller effusive flows and domes. Effusive eruptions have been documented preceding and following caldera-forming eruptions, however the temporal and petrogenetic relationships of these magmas to the caldera-forming eruptions are relatively unknown. Here we present new 40Ar/39Ar dates for four small-volume eruptions located on the western rim of the second-cycle caldera, the source of the 1.300 ± 0.001 Ma Mesa Falls Tuff. We supplement our new eruption ages with whole rock major and trace element chemistry, Pb isotopic ratios of feldspar, and paleomagnetic and rock magnetic analyses. Eruption ages for the effusive Green Canyon Flow (1.299 ± 0.002 Ma) and Moonshine Mountain Dome (1.302 ± 0.003 Ma) are in close temporal proximity to the eruption age of the Mesa Falls Tuff. In contrast, our results indicate a period of volcanism at ca 1.45 Ma within the Yellowstone Volcanic Field, including the eruption of the Bishop Mountain Flow (1.458 ± 0.002 Ma) and Tuff of Lyle Spring (1.450 ± 0.003 Ma). These high-silica rhyolites are chemically and isotopically distinct from the Mesa Falls Tuff and related 1.3 Ma effusive eruptions. The 40Ar/39Ar data from the Tuff of Lyle Spring demonstrate significant antecrystic inheritance, prevalent within the upper welded ash-flow tuff matrix, and minimal within individual pumice. Antecrysts are up to 20 kyr older than the eruption, with subpopulations of grains occurring every few thousand years. We interpret these results as an indicator for the timing of magmatic pulses into a growing magmatic system that would ultimately erupt the Tuff of Lyle Spring, and which we more broadly interpret as the tempo of crustal accumulation associated with bimodal magmatism. We propose a system whereby chemically, isotopically, and temporally distinct, isolated small-volume magma batches are

  15. Monitoring super-volcanoes: geophysical and geochemical signals at Yellowstone and other large caldera systems.

    Science.gov (United States)

    Lowenstern, Jacob B; Smith, Robert B; Hill, David P

    2006-08-15

    Earth's largest calderas form as the ground collapses during immense volcanic eruptions, when hundreds to thousands of cubic kilometres of magma are explosively withdrawn from the Earth's crust over a period of days to weeks. Continuing long after such great eruptions, the resulting calderas often exhibit pronounced unrest, with frequent earthquakes, alternating uplift and subsidence of the ground, and considerable heat and mass flux. Because many active and extinct calderas show evidence for repetition of large eruptions, such systems demand detailed scientific study and monitoring. Two calderas in North America, Yellowstone (Wyoming) and Long Valley (California), are in areas of youthful tectonic complexity. Scientists strive to understand the signals generated when tectonic, volcanic and hydrothermal (hot ground water) processes intersect. One obstacle to accurate forecasting of large volcanic events is humanity's lack of familiarity with the signals leading up to the largest class of volcanic eruptions. Accordingly, it may be difficult to recognize the difference between smaller and larger eruptions. To prepare ourselves and society, scientists must scrutinize a spectrum of volcanic signals and assess the many factors contributing to unrest and toward diverse modes of eruption.

  16. Whitebark pine vulnerability to climate-driven mountain pine beetle disturbance in the Greater Yellowstone Ecosystem.

    Science.gov (United States)

    Logan, Jesse A; MacFarlane, William W; Willcox, Louisa

    2010-06-01

    Widespread outbreaks of mountain pine beetles (MPB) are occurring throughout the range of this native insect. Episodic outbreaks are a common occurrence in the beetles' primary host, lodgepole pine. Current outbreaks, however, are occurring in habitats where outbreaks either did not previously occur or were limited in scale. Herein, we address widespread, ongoing outbreaks in high-elevation, whitebark pine forests of the Greater Yellowstone Ecosystem, where, due to an inhospitable climate, past outbreaks were infrequent and short lived. We address the basic question: are these outbreaks truly unprecedented and a threat to ecosystem continuity? In order to evaluate this question we (1) present evidence that the current outbreak is outside the historic range of variability; (2) examine system resiliency to MPB disturbance based on adaptation to disturbance and host defenses to MPB attack; and (3) investigate the potential domain of attraction to large-scale MPB disturbance based on thermal developmental thresholds, spatial structure of forest types, and the confounding influence of an introduced pathogen. We conclude that the loss of dominant whitebark pine forests, and the ecological services they provide, is likely under continuing climate warming and that new research and strategies are needed to respond to the crisis facing whitebark pine.

  17. Microbial communities and chemosynthesis in Yellowstone Lake sublacustrine hydrothermal vent waters

    Directory of Open Access Journals (Sweden)

    Tingting eYang

    2011-06-01

    Full Text Available Abstract. Five sublacustrine hydrothermal vent locations from 1-109 m water depth in Yellowstone Lake were surveyed by ribosomal RNA sequencing in relation to their chemical composition and dark CO2 fixation rates. They harbor distinct chemosynthetic bacterial communities, depending on temperature (16 - 110ºC and electron donor supply (H2S <1 - >100µM; NH3 <0.5 - >10µM. Members of the Aquificales, most closely affiliated with the genus Sulfurihydrogenibium, are the most frequently recovered bacterial 16S rRNA gene phylotypes in the hottest samples; the detection of these thermophilic sulfur-oxidizing autotrophs coincided with maximal dark CO2 fixation rates reaching near 9 µM C h-1 at temperatures of 50 to 60°C. Vents at lower temperatures yielded mostly phylotypes related to the mesophilic gammaproteobacterial sulfur oxidizer Thiovirga. In contrast, cool vent water with low chemosynthetic activity yielded predominantly phylotypes related to freshwater Actinobacterial clusters with a cosmopolitan distribution.

  18. Genetic analysis of individual origins supports isolation of grizzly bears in the Greater Yellowstone Ecosystem

    Science.gov (United States)

    Haroldson, Mark A.; Schwartz, Charles; Kendall, Katherine C.; Gunther, Kerry A.; Moody, David S.; Frey, Kevin L.; Paetkau, David

    2010-01-01

    The Greater Yellowstone Ecosystem (GYE) supports the southernmost of the 2 largest remaining grizzly bear (Ursus arctos) populations in the contiguous United States. Since the mid-1980s, this population has increased in numbers and expanded in range. However, concerns for its long-term genetic health remain because of its presumed continued isolation. To test the power of genetic methods for detecting immigrants, we generated 16-locus microsatellite genotypes for 424 individual grizzly bears sampled in the GYE during 1983–2007. Genotyping success was high (90%) and varied by sample type, with poorest success (40%) for hair collected from mortalities found ≥1 day after death. Years of storage did not affect genotyping success. Observed heterozygosity was 0.60, with a mean of 5.2 alleles/marker. We used factorial correspondence analysis (Program GENETIX) and Bayesian clustering (Program STRUCTURE) to compare 424 GYE genotypes with 601 existing genotypes from grizzly bears sampled in the Northern Continental Divide Ecosystem (NCDE) (FST  =  0.096 between GYE and NCDE). These methods correctly classified all sampled individuals to their population of origin, providing no evidence of natural movement between the GYE and NCDE. Analysis of 500 simulated first-generation crosses suggested that over 95% of such bears would also be detectable using our 16-locus data set. Our approach provides a practical method for detecting immigration in the GYE grizzly population. We discuss estimates for the proportion of the GYE population sampled and prospects for natural immigration into the GYE.

  19. Effects of management and climate on elk brucellosis in the Greater Yellowstone Ecosystem

    Science.gov (United States)

    Cross, P.C.; Edwards, W.H.; Scurlock, B.M.; Maichak, E.J.; Rogerson, J.D.

    2007-01-01

    Every winter, government agencies feed ???6000 metric tons (6 ?? 106 kg) of hay to elk in the southern Greater Yellowstone Ecosystem (GYE) to limit transmission of Brucella abortus, the causative agent of brucellosis, from elk to cattle. Supplemental feeding, however, is likely to increase the transmission of brucellosis in elk, and may be affected by climatic factors, such as snowpack. We assessed these possibilities using snowpack and feeding data from 1952 to 2006 and disease testing data from 1993 to 2006. Brucellosis seroprevalence was strongly correlated with the timing of the feeding season. Longer feeding seasons were associated with higher seroprevalence, but elk population size and density had only minor effects. In other words, the duration of host aggregation and whether it coincided with peak transmission periods was more important than just the host population size. Accurate modeling of disease transmission depends upon incorporating information on how host contact rates fluctuate over time relative to peak transmission periods. We also found that supplemental feeding seasons lasted longer during years with deeper snowpack. Therefore, milder winters and/or management strategies that reduce the length of the feeding season may reduce the seroprevalence of brucellosis in the elk populations of the southern GYE. ?? 2007 by the Ecological Society of America.

  20. Brucellosis Transmission between Wildlife and Livestock in the Greater Yellowstone Ecosystem: Inferences from DNA Genotyping.

    Science.gov (United States)

    O'Brien, Michael P; Beja-Pereira, Albano; Anderson, Neil; Ceballos, Ruben M; Edwards, William H; Harris, Beth; Wallen, Rick L; Costa, Vânia

    2017-04-01

    The wildlife of the Greater Yellowstone Ecosystem carries brucellosis, which was first introduced to the area by cattle in the 19th century. Brucellosis transmission between wildlife and livestock has been difficult to study due to challenges in culturing the causative agent, Brucella abortus . We examined B. abortus transmission between American bison ( Bison bison ), Rocky Mountain elk ( Cervus elaphus nelsoni), and cattle ( Bos taurus ) using variable number tandem repeat (VNTR) markers on DNA from 98 B. abortus isolates recovered from populations in Idaho, Montana, and Wyoming, US. Our analyses reveal interspecies transmission. Two outbreaks (2007, 2008) in Montana cattle had B. abortus genotypes similar to isolates from both bison and elk. Nevertheless, similarity in elk and cattle isolates from the 2008 outbreak suggest that elk are the likely source of brucellosis transmission to cattle in Montana and Wyoming. Brucella abortus isolates from sampling in Montana appear to be divided in two clusters: one found in local Montana elk, cattle, and bison; and another found mainly in elk and a bison from Wyoming, which is consistent with brucellosis having entered Montana via migration of infected elk from Wyoming. Our findings illustrate complex patterns of brucellosis transmission among elk, bison, and cattle as well as the utility of VNTRs to infer the wildlife species of origin for disease outbreaks in livestock.

  1. HIV behavioural surveillance among refugees and surrounding host ...

    African Journals Online (AJOL)

    We used a standardised behavioural surveillance survey (BSS), modified to be directly relevant to populations in conflict and post-conflict settings as well as to their surrounding host populations, to survey the populations of a refugee settlement in south-western Uganda and its surrounding area. Two-stage probability ...

  2. Investigation of the readout electronics of DELPHI surround muon chamber

    International Nuclear Information System (INIS)

    Khovanskij, N.; Krumshtejn, Z.; Ol'shevskij, A.; Sadovskij, A.; Sedykh, Yu.; Molnar, J.; Sicho, P.; Tomsa, Z.

    1995-01-01

    The characteristics of the readout electronics of the DELPHI surround muon chambers with various AMPLEX chips (AMPLEX 16 and AMPLEX-SICAL) are presented. This electronics is studied in a cosmic rays test of the real surround muon chamber model. 4 refs., 6 figs., 1 tab

  3. Stimulus size dependence of hue changes induced by chromatic surrounds.

    Science.gov (United States)

    Kellner, Christian Johannes; Wachtler, Thomas

    2016-03-01

    A chromatic surround induces a change in the perceived hue of a stimulus. This shift in hue depends on the chromatic difference between the stimulus and the surround. We investigated how chromatic induction varies with stimulus size and whether the size dependence depends on the surround hue. Subjects performed asymmetric matching of color stimuli with different sizes in surrounds of different chromaticities. Generally, induced hue shifts decreased with increasing stimulus size. This decrease was quantitatively different for different surround hues. However, when size effects were normalized to an overall induction strength, the chromatic specificity was largely reduced. The separability of inducer chromaticity and stimulus size suggests that these effects are mediated by different neural mechanisms.

  4. Enhancing fire science exchange: The Joint Fire Science Program's National Network of Knowledge Exchange Consortia

    Science.gov (United States)

    Vita Wright; Crystal Kolden; Todd Kipfer; Kristine Lee; Adrian Leighton; Jim Riddering; Leana Schelvan

    2011-01-01

    The Northern Rocky Mountain region is one of the most fire-prone regions in the United States. With a history of large fires that have shaped national policy, including the fires of 1910 and 2000 in Idaho and Montana and the Yellowstone fires of 1988, this region is projected to have many large severe fires in the future. Communication about fire science needs and...

  5. Multiple estimates of effective population size for monitoring a long-lived vertebrate: an application to Yellowstone grizzly bears.

    Science.gov (United States)

    Kamath, Pauline L; Haroldson, Mark A; Luikart, Gordon; Paetkau, David; Whitman, Craig; van Manen, Frank T

    2015-11-01

    Effective population size (N(e)) is a key parameter for monitoring the genetic health of threatened populations because it reflects a population's evolutionary potential and risk of extinction due to genetic stochasticity. However, its application to wildlife monitoring has been limited because it is difficult to measure in natural populations. The isolated and well-studied population of grizzly bears (Ursus arctos) in the Greater Yellowstone Ecosystem provides a rare opportunity to examine the usefulness of different N(e) estimators for monitoring. We genotyped 729 Yellowstone grizzly bears using 20 microsatellites and applied three single-sample estimators to examine contemporary trends in generation interval (GI), effective number of breeders (N(b)) and N(e) during 1982-2007. We also used multisample methods to estimate variance (N(eV)) and inbreeding N(e) (N(eI)). Single-sample estimates revealed positive trajectories, with over a fourfold increase in N(e) (≈100 to 450) and near doubling of the GI (≈8 to 14) from the 1980s to 2000s. N(eV) (240-319) and N(eI) (256) were comparable with the harmonic mean single-sample N(e) (213) over the time period. Reanalysing historical data, we found N(eV) increased from ≈80 in the 1910s-1960s to ≈280 in the contemporary population. The estimated ratio of effective to total census size (N(e) /N(c)) was stable and high (0.42-0.66) compared to previous brown bear studies. These results support independent demographic evidence for Yellowstone grizzly bear population growth since the 1980s. They further demonstrate how genetic monitoring of N(e) can complement demographic-based monitoring of N(c) and vital rates, providing a valuable tool for wildlife managers. © 2015 John Wiley & Sons Ltd.

  6. Migrations and swimming capabilities of endangered pallid sturgeon (Scaphirhynchus albus) to guide passage designs in the fragmented Yellowstone River

    Science.gov (United States)

    Braaten, P. J.; Elliott, Caroline M.; Rhoten, Jason C.; Fuller, D. B.; McElroy, Brandon J.

    2015-01-01

    Fragmentation of the Yellowstone River is hypothesized to preclude recruitment of endangered Scaphirhynchus albus (pallid sturgeon) by impeding upstream spawning migrations and access to upstream spawning areas, thereby limiting the length of free-flowing river required for survival of early life stages. Building on this hypothesis, the reach of the Yellowstone River affected by Intake Diversion Dam (IDD) is targeted for modification. Structures including a rock ramp and by-pass channel have been proposed as restoration alternatives to facilitate passage. Limited information on migrations and swimming capabilities of pallid sturgeon is available to guide engineering design specifications for the proposed structures. Migration behavior, pathways (channel routes used during migrations), and swimming capabilities of free-ranging wild adult pallid sturgeon were examined using radiotelemetry, and complemented with hydraulic data obtained along the migration pathways. Migrations of 12–26% of the telemetered pallid sturgeon population persisted to IDD, but upstream passage over the dam was not detected. Observed migration pathways occurred primarily through main channel habitats; however, migrations through side channels up to 3.9 km in length were documented. The majority of pallid sturgeon used depths of 2.2–3.4 m and mean water velocities of 0.89–1.83 m/s while migrating. Results provide inferences on depths, velocities, and habitat heterogeneity of reaches successfully negotiated by pallid sturgeon that may be used to guide designs for structures facilitating passage at IDD. Passage will provide connectivity to potential upstream spawning areas on the Yellowstone River, thereby increasing the likelihood of recruitment for this endangered species.

  7. Multiple estimates of effective population size for monitoring a long-lived vertebrate: An application to Yellowstone grizzly bears

    Science.gov (United States)

    Kamath, Pauline L.; Haroldson, Mark A.; Luikart, Gordon; Paetkau, David; Whitman, Craig L.; van Manen, Frank T.

    2015-01-01

    Effective population size (Ne) is a key parameter for monitoring the genetic health of threatened populations because it reflects a population's evolutionary potential and risk of extinction due to genetic stochasticity. However, its application to wildlife monitoring has been limited because it is difficult to measure in natural populations. The isolated and well-studied population of grizzly bears (Ursus arctos) in the Greater Yellowstone Ecosystem provides a rare opportunity to examine the usefulness of different Ne estimators for monitoring. We genotyped 729 Yellowstone grizzly bears using 20 microsatellites and applied three single-sample estimators to examine contemporary trends in generation interval (GI), effective number of breeders (Nb) and Ne during 1982–2007. We also used multisample methods to estimate variance (NeV) and inbreeding Ne (NeI). Single-sample estimates revealed positive trajectories, with over a fourfold increase in Ne (≈100 to 450) and near doubling of the GI (≈8 to 14) from the 1980s to 2000s. NeV (240–319) and NeI (256) were comparable with the harmonic mean single-sample Ne (213) over the time period. Reanalysing historical data, we found NeV increased from ≈80 in the 1910s–1960s to ≈280 in the contemporary population. The estimated ratio of effective to total census size (Ne/Nc) was stable and high (0.42–0.66) compared to previous brown bear studies. These results support independent demographic evidence for Yellowstone grizzly bear population growth since the 1980s. They further demonstrate how genetic monitoring of Ne can complement demographic-based monitoring of Nc and vital rates, providing a valuable tool for wildlife managers.

  8. Use of sulfur and nitrogen stable isotopes to determine the importance of whitebark pine nuts to Yellowstone grizzly bears

    Science.gov (United States)

    Felicetti, L.A.; Schwartz, C.C.; Rye, R.O.; Haroldson, M.A.; Gunther, K.A.; Phillips, D.L.; Robbins, C.T.

    2003-01-01

    Whitebark pine (Pinus albicaulis) is a masting species that produces relatively large, fat- and protein-rich nuts that are consumed by grizzly bears (Ursus arctos horribilis). Trees produce abundant nut crops in some years and poor crops in other years. Grizzly bear survival in the Greater Yellowstone Ecosystem is strongly linked to variation in pine-nut availability. Because whitebark pine trees are infected with blister rust (Cronartium ribicola), an exotic fungus that has killed the species throughout much of its range in the northern Rocky Mountains, we used stable isotopes to quantify the importance of this food resource to Yellowstone grizzly bears while healthy populations of the trees still exist. Whitebark pine nuts have a sulfur-isotope signature (9.2 ?? 1.3??? (mean ?? 1 SD)) that is distinctly different from those of all other grizzly bear foods (ranging from 1.9 ?? 1.7??? for all other plants to 3.1 ?? 2.6??? for ungulates). Feeding trials with captive grizzly bears were used to develop relationships between dietary sulfur-, carbon-, and nitrogen-isotope signatures and those of bear plasma. The sulfur and nitrogen relationships were used to estimate the importance of pine nuts to free-ranging grizzly bears from blood and hair samples collected between 1994 and 2001. During years of poor pine-nut availability, 72% of the bears made minimal use of pine nuts. During years of abundant cone availability, 8 ?? 10% of the bears made minimal use of pine nuts, while 67 ?? 19% derived over 51% of their assimilated sulfur and nitrogen (i.e., protein) from pine nuts. Pine nuts and meat are two critically important food resources for Yellowstone grizzly bears.

  9. Quantifying Carbon Consequences of Recent Land Management and Disturbances in the Greater Yellowstone Ecosystems (GYE) by linking inventory data, remote sensing and carbon modeling

    Science.gov (United States)

    Zhao, F.; Huang, C.; Healey, S. P.; McCarter, J. B.; Garrard, C.; Zhu, Z.

    2015-12-01

    Natural disturbances and land management directly change C stored in biomass and soil pools, and can have indirect impacts on long-term C balance. The Greater Yellowstone Ecosystem (GYE), located in Central Rocky Mountains of United States, is of different land ownerships within similar environmental settings, making it an ideal site to examine the impacts of management and disturbances on regional carbon dynamics. Recent advances in the remote sensing of vegetation condition and change, along with new techniques linking remote sensing with inventory records, have allowed investigations that are much more tightly constrained to actual landscape environment, instead of hypothetical or generalized conditions. These new capabilities are built into the Forest Carbon Management Framework (ForCaMF), which is being used by the National Forest System to not only model, but to monitor across very specific management units, the impact of different kinds of disturbance on carbon storage. In this study, we used the ForCaMF approach to evaluate carbon effects of natural disturbances (e.g. wildfire) and land management (e.g. harvests) in GYE National Parks, Wilderness Area and National Forests. As might be expected, wildfire has been the dominant disturbance factor in the carbon cycle of GYE's administratively protected areas since the mid-1980s, while harvests have dominated storage trends on the managed land in the region's National Forests. Moving beyond this monitoring result but maintaining the same fidelity to historical vegetation patterns, we are also able to simulate alternative disturbance scenarios to provide landscape-specific insights to forest managers. We can estimate likely carbon storage impacts in GYE protected areas, for example, if more active fire suppression had been pursued since the mid-1980s. Likewise, we can identify differences in current carbon storage on managed lands if high harvest rates during the same period had been moderated. We discuss

  10. Drainage and Landscape Evolution in the Bighorn Basin Accompanying Advection of the Yellowstone Hotspot Swell Through North America

    Science.gov (United States)

    Guerrero, E. F.; Meigs, A.

    2012-12-01

    Mantle plumes have been recognized to express themselves on the surface as long wavelength and low amplitude topographic swells. These swells are measured as positive geoid anomalies and include shorter wavelength topographic features such as volcanic edifices and pre-exisitng topography. Advection of the topographic swell is expected as the lithosphere passes over the plume uplift source. The hot spot swell occurs in the landscape as transient signal that is expressed with waxing and waning topography. Waxing topography occurs at the leading edge of the swell and is expressed as an increase in rock uplift that is preserved by rivers and landscapes. Advection of topography predicts a shift in a basin from deposition to incision, an increase in convexity of a transverse river's long profile and a lateral river migration in the direction of advection. The Yellowstone region has a strong positive geoid anomaly and the volcanic signal, which have been interpreted as the longer and shorter wavelength topographic expressions of the hot spot. These expressions of the hot spot developed in a part of North America with a compounded deformation and topographic history. Previous studies of the Yellowstone topographic swell have concentrated on the waning or trailing signal preserved in the Snake River Plain. Our project revisits the classic geomorphology study area in the Bighorn Basin of Wyoming and Montana, which is in leading edge of the swell. Present models identify the swell as having a 400 km in diameter and that it is centered on the Yellowstone caldera. If we assume advection to occur in concert with the caldera eruptive track, the Yellowstone swell has migrated to the northeast at a rate of 3 cm yr-1 and began acting on the Bighorn Basin's landscape between 3 and 2 Ma. The Bighorn Basin has an established history of a basin-wide switch from deposition to incision during the late Pliocene, yet the age control on the erosional evolution of the region is relative. This

  11. Changes in unique hues induced by chromatic surrounds.

    Science.gov (United States)

    Klauke, Susanne; Wachtler, Thomas

    2016-03-01

    A chromatic surround can have a strong influence on the perceived hue of a stimulus. We investigated whether chromatic induction has similar effects on the perception of colors that appear pure and unmixed (unique red, green, blue, and yellow) as on other colors. Subjects performed unique hue settings of stimuli in isoluminant surrounds of different chromaticities. Compared with the settings in a neutral gray surround, unique hue settings altered systematically with chromatic surrounds. The amount of induced hue shift depended on the difference between stimulus and surround hues, and was similar for unique hue settings as for settings of nonunique hues. Intraindividual variability in unique hue settings was roughly twice as high as for settings obtained in asymmetric matching experiments, which may reflect the presence of a reference stimulus in the matching task. Variabilities were also larger with chromatic surrounds than with neutral gray surrounds, for both unique hue settings and matching of nonunique hues. The results suggest that the neural representations underlying unique hue percepts are influenced by the same neural processing mechanisms as the percepts of other colors.

  12. Beaver Activity, Holocene Climate and Riparian Landscape Change Across Stream Scales in the Greater Yellowstone Ecosystem

    Science.gov (United States)

    Levine, R.; Meyer, G. A.

    2013-12-01

    Beaver (Castor canadensis) have been part of the fluvial and riparian landscape across much of North America since the Pleistocene, increasing channel habitat complexity and expanding riparian landscapes. The fur trade, however, decimated beaver populations by the 1840s, and other human activities have limited beaver in many areas, including parts of the Greater Yellowstone Ecosystem (GYE). Understanding fluctuations in beaver occupation through the Holocene will aid in understanding the natural range of variability in beaver activity as well as climatic and anthropogenic impacts to fluvial systems. We are developing a detailed chronology of beaver-assisted sedimentation and overall fluvial activity for Odell and Red Rock Creeks (basin areas 83 and 99 km2) in Centennial Valley (CV), Montana, to augment related studies on the long-term effects of beaver on smaller GYE fluvial systems (basin areas 0.1-50 km2). In developing the CV chronology, we use the presence of concentrations of beaver-chewed sticks as a proxy for beaver occupancy. Beaver-stick deposits are found in paleochannel and fluvial terrace exposures. The relative ages of exposures were determined by elevation data from airborne LiDAR and ground surveys. Numerical ages were obtained from 36 14C ages (~30 more are pending) of beaver-stick wood collected during investigation of the stratigraphy. Most beaver-stick deposits are associated with ~ 1 meter of fine-grained sediment, interpreted as overbank deposits, commonly overlying gravelly sand or pebble gravel channel deposits which is consistent with enhanced overbank sedimentation associated with active beaver dams in CV streams. The CV deposits differ from those on smaller GYE streams where beaver-stick deposits are associated with abandoned dams (berms), infilled ponds and laminated sediments. The lack of pond-related deposition associated with CV beaver-stick deposits is consistent with frequent dam breaching (≤ 5 years) in the modern channel of Odell

  13. Grizzly bear denning chronology and movements in the Greater Yellowstone Ecosystem

    Science.gov (United States)

    Haroldson, Mark A.; Ternent, Mark A.; Gunther, Kerry A.; Schwartz, Charles C.

    2002-01-01

    Den entrance and emergence dates of grizzly bears (Ursus arctos) in the Greater Yellowstone Ecosystem are important to management agencies that wish to minimize impacts of human activities on bears. Current estimates for grizzly bear denning events use data that were collected from 1975–80. We update these estimates by including data obtained from 1981–99. We used aerial telemetry data to estimate week of den entry and emergence by determining the midpoint between the last known active date and the first known date denned, as well as the last known date denned and the first known active date. We also investigated post emergence movement patterns relative to den locations. Mean earliest and latest week of den entry and emergence were also determined. Den entry for females began during the fourth week in September, with 90% denned by the fourth week of November. Earliest den entry for males occurred during the second week of October, with 90% denned by the second week of December. Mean week of den entry for known pregnant females was earlier than males. Earliest week of den entry for known pregnant females was earlier than other females and males. Earliest den emergence for males occurred during the first week of February, with 90% of males out of dens by the fourth week of April. Earliest den emergence for females occurred during the third week of March; by the first week of May, 90% of females had emerged. Male bears emerged from dens earlier than females. Denning period differed among classes and averaged 171 days for females that emerged from dens with cubs, 151 days for other females, and 131 days for males. Known pregnant females tended to den at higher elevations and, following emergence, remained at higher elevation until late May. Females with cubs remained relatively close (grizzly bear populations in the southern Rocky Mountains. 

  14. Methods to estimate distribution and range extent of grizzly bears in the Greater Yellowstone Ecosystem

    Science.gov (United States)

    Haroldson, Mark A.; Schwartz, Charles C.; Thompson, Daniel J.; Bjornlie, Daniel D.; Gunther, Kerry A.; Cain, Steven L.; Tyers, Daniel B.; Frey, Kevin L.; Aber, Bryan C.

    2014-01-01

    The distribution of the Greater Yellowstone Ecosystem grizzly bear (Ursus arctos) population has expanded into areas unoccupied since the early 20th century. Up-to-date information on the area and extent of this distribution is crucial for federal, state, and tribal wildlife and land managers to make informed decisions regarding grizzly bear management. The most recent estimate of grizzly bear distribution (2004) utilized fixed-kernel density estimators to describe distribution. This method was complex and computationally time consuming and excluded observations of unmarked bears. Our objective was to develop a technique to estimate grizzly bear distribution that would allow for the use of all verified grizzly bear location data, as well as provide the simplicity to be updated more frequently. We placed all verified grizzly bear locations from all sources from 1990 to 2004 and 1990 to 2010 onto a 3-km × 3-km grid and used zonal analysis and ordinary kriging to develop a predicted surface of grizzly bear distribution. We compared the area and extent of the 2004 kriging surface with the previous 2004 effort and evaluated changes in grizzly bear distribution from 2004 to 2010. The 2004 kriging surface was 2.4% smaller than the previous fixed-kernel estimate, but more closely represented the data. Grizzly bear distribution increased 38.3% from 2004 to 2010, with most expansion in the northern and southern regions of the range. This technique can be used to provide a current estimate of grizzly bear distribution for management and conservation applications.

  15. Use of isotopic sulfur to determine whitebark pine consumption by Yellowstone bears: a reassessment

    Science.gov (United States)

    Schwartz, Charles C.; Teisberg, Justin E.; Fortin, Jennifer K.; Haroldson, Mark A.; Servheen, Christopher; Robbins, Charles T.; van Manen, Frank T.

    2014-01-01

    Use of naturally occurring stable isotopes to estimate assimilated diet of bears is one of the single greatest breakthroughs in nutritional ecology during the past 20 years. Previous research in the Greater Yellowstone Ecosystem (GYE), USA, established a positive relationship between the stable isotope of sulfur (δ34S) and consumption of whitebark pine (Pinus albicaulis) seeds. That work combined a limited sample of hair, blood clots, and serum. Here we use a much larger sample to reassess those findings. We contrasted δ34S values in spring hair and serum with abundance of seeds of whitebark pine in samples collected from grizzly (Ursus arctos) and American black bears (U. americanus) in the GYE during 2000–2010. Although we found a positive relationship between δ34S values in spring hair and pine seed abundance for grizzly bears, the coefficients of determination were small (R2 ≤ 0.097); we failed to find a similar relationship with black bears. Values of δ34S in spring hair were larger in black bears and δ34S values in serum of grizzly bears were lowest in September and October, a time when we expect δ34S to peak if whitebark pine seeds were the sole source of high δ34S. The relationship between δ34S in bear tissue and the consumption of whitebark pine seeds, as originally reported, may not be as clean a method as proposed. Data we present here suggest other foods have high values of δ34S, and there is spatial heterogeneity affecting the δ34S values in whitebark pine, which must be addressed.

  16. Sex-biased gene flow among elk in the greater Yellowstone ecosystem

    Science.gov (United States)

    Hand, Brian K.; Chen, Shanyuan; Anderson, Neil; Beja-Pereira, Albano; Cross, Paul C.; Ebinger, Michael R.; Edwards, Hank; Garrott, Robert A.; Kardos, Marty D.; Kauffman, Matthew J.; Landguth, Erin L.; Middleton, Arthur; Scurlock, Brandon M.; White, P.J.; Zager, Pete; Schwartz, Michael K.; Luikart, Gordon

    2014-01-01

    We quantified patterns of population genetic structure to help understand gene flow among elk populations across the Greater Yellowstone Ecosystem. We sequenced 596 base pairs of the mitochondrial control region of 380 elk from eight populations. Analysis revealed high mitochondrial DNA variation within populations, averaging 13.0 haplotypes with high mean gene diversity (0.85). The genetic differentiation among populations for mitochondrial DNA was relatively high (FST  =  0.161; P  =  0.001) compared to genetic differentiation for nuclear microsatellite data (FST  =  0.002; P  =  0.332), which suggested relatively low female gene flow among populations. The estimated ratio of male to female gene flow (mm/mf  =  46) was among the highest we have seen reported for large mammals. Genetic distance (for mitochondrial DNA pairwise FST) was not significantly correlated with geographic (Euclidean) distance between populations (Mantel's r  =  0.274, P  =  0.168). Large mitochondrial DNA genetic distances (e.g., FST > 0.2) between some of the geographically closest populations (<65 km) suggested behavioral factors and/or landscape features might shape female gene flow patterns. Given the strong sex-biased gene flow, future research and conservation efforts should consider the sexes separately when modeling corridors of gene flow or predicting spread of maternally transmitted diseases. The growing availability of genetic data to compare male vs. female gene flow provides many exciting opportunities to explore the magnitude, causes, and implications of sex-biased gene flow likely to occur in many species.

  17. A synchronous surround increases the motion strength gain of motion.

    Science.gov (United States)

    Linares, Daniel; Nishida, Shin'ya

    2013-11-12

    Coherent motion detection is greatly enhanced by the synchronous presentation of a static surround (Linares, Motoyoshi, & Nishida, 2012). To further understand this contextual enhancement, here we measured the sensitivity to discriminate motion strength for several pedestal strengths with and without a surround. We found that the surround improved discrimination of low and medium motion strengths, but did not improve or even impaired discrimination of high motion strengths. We used motion strength discriminability to estimate the perceptual response function assuming additive noise and found that the surround increased the motion strength gain, rather than the response gain. Given that eye and body movements continuously introduce transients in the retinal image, it is possible that this strength gain occurs in natural vision.

  18. Surrounding Moving Obstacle Detection for Autonomous Driving Using Stereo Vision

    Directory of Open Access Journals (Sweden)

    Hao Sun

    2013-06-01

    Full Text Available Detection and tracking surrounding moving obstacles such as vehicles and pedestrians are crucial for the safety of mobile robotics and autonomous vehicles. This is especially the case in urban driving scenarios. This paper presents a novel framework for surrounding moving obstacles detection using binocular stereo vision. The contributions of our work are threefold. Firstly, a multiview feature matching scheme is presented for simultaneous stereo correspondence and motion correspondence searching. Secondly, the multiview geometry constraint derived from the relative camera positions in pairs of consecutive stereo views is exploited for surrounding moving obstacles detection. Thirdly, an adaptive particle filter is proposed for tracking of multiple moving obstacles in surrounding areas. Experimental results from real-world driving sequences demonstrate the effectiveness and robustness of the proposed framework.

  19. Contamination of nebulisers and surrounding air at the bedside of ...

    African Journals Online (AJOL)

    An air sampler was used to collect air samples from the surrounding bedside environment. .... individualised resealable plastic bags and stored upside down in a cooler .... conventional and mesh technology nebulisers used at home by adults.

  20. Glow phenomenon surrounding the vertical stabilizer and OMS pods

    Science.gov (United States)

    1994-01-01

    This 35mm frame, photographed as the Space Shuttle Columbia was orbiting Earth during a 'night' pass, documents the glow phenomenon surrounding the vertical stabilizer and the Orbital Maneuvering System (OMS) pods of the spacecraft.

  1. Complex challenges of maintaining whitebark pine in Greater Yellowstone under climate change: A call for innovative research, management, and policy approaches

    Science.gov (United States)

    Andrew Hansen; Kathryn Ireland; Kristin Legg; Robert Keane; Edward Barge; Martha Jenkins; Michiel Pillet

    2016-01-01

    Climate suitability is projected to decline for many subalpine species, raising questions about managing species under a deteriorating climate. Whitebark pine (WBP) (Pinus albicaulis) in the Greater Yellowstone Ecosystem (GYE) crystalizes the challenges that natural resource managers of many high mountain ecosystems will likely face in the coming decades. We...

  2. Monitoring program of surrounding of the NPP SE-EBO

    International Nuclear Information System (INIS)

    Dobis, L.; Kostial, J.

    1997-01-01

    The paper dealt with monitoring program of radiation control of surrounding of the NPP Bohunice, which has the aim: (1) to ensure the control of influence of work of the NPP Bohunice on the environment in their surrounding; (2) to ensure the back-ground for regular brief of control and supervisory organs about condition of the environment in surrounding of the NPP Bohunice; (3) to maintain the expected technical level of control of the NPP Bohunice and to exploit optimally the technical means; (4) to solicit permanently the data about the radioactivity of environment in surrounding of the NPP Bohunice for forming of files of the data; (5) to exploit purposefully the technical equipment, technical workers and to maintain their in permanent emergency and technical eligibility for the case of the breakdown; (6) to obtain permanently the files of the values for qualification of the reference levels. This program of monitoring includes the radiation control of surrounding of the NPP Bohunice, in the time of normal work of power-station's blocks, inclusively of all types of trouble-shooting and repairer works in surrounding of the NPP Bohunice, up to distance 20 km from power-station. The monitoring includes: outlets from the NPP Bohunice, monitoring of radiation characteristics in surrounding of the NPP Bohunice, (aerosols, fall-outs, soil), the links of food chains: (grass and fodder, milk, agriculture products), hydrosphere in surrounding (surface waters, drink water, bores of radiation control in complex of the NPP Bohunice, components of the hydrosphere), measurement of radiation from external sources (measurement of the dose rates, measurement of the doses [sk

  3. Chromatic induction from surrounding stimuli under perceptual suppression.

    Science.gov (United States)

    Horiuchi, Koji; Kuriki, Ichiro; Tokunaga, Rumi; Matsumiya, Kazumichi; Shioiri, Satoshi

    2014-11-01

    The appearance of colors can be affected by their spatiotemporal context. The shift in color appearance according to the surrounding colors is called color induction or chromatic induction; in particular, the shift in opponent color of the surround is called chromatic contrast. To investigate whether chromatic induction occurs even when the chromatic surround is imperceptible, we measured chromatic induction during interocular suppression. A multicolor or uniform color field was presented as the surround stimulus, and a colored continuous flash suppression (CFS) stimulus was presented to the dominant eye of each subject. The subjects were asked to report the appearance of the test field only when the stationary surround stimulus is invisible by interocular suppression with CFS. The resulting shifts in color appearance due to chromatic induction were significant even under the conditions of interocular suppression for all surround stimuli. The magnitude of chromatic induction differed with the surround conditions, and this difference was preserved regardless of the viewing conditions. The chromatic induction effect was reduced by CFS, in proportion to the magnitude of chromatic induction under natural (i.e., no-CFS) viewing conditions. According to an analysis with linear model fitting, we revealed the presence of at least two kinds of subprocesses for chromatic induction that reside at higher and lower levels than the site of interocular suppression. One mechanism yields different degrees of chromatic induction based on the complexity of the surround, which is unaffected by interocular suppression, while the other mechanism changes its output with interocular suppression acting as a gain control. Our results imply that the total chromatic induction effect is achieved via a linear summation of outputs from mechanisms that reside at different levels of visual processing.

  4. Forest Fragments Surrounded by Sugar Cane Are More Inhospitable to Terrestrial Amphibian Abundance Than Fragments Surrounded by Pasture

    Directory of Open Access Journals (Sweden)

    Paula Eveline Ribeiro D’Anunciação

    2013-01-01

    Full Text Available In recent years, there has been increasing interest in matrix-type influence on forest fragments. Terrestrial amphibians are good bioindicators for this kind of research because of low vagility and high philopatry. This study compared richness, abundance, and species composition of terrestrial amphibians through pitfall traps in two sets of semideciduous seasonal forest fragments in southeastern Brazil, according to the predominant surrounding matrix (sugar cane and pasture. There were no differences in richness, but fragments surrounded by sugar cane had the lowest abundance of amphibians, whereas fragments surrounded by pastures had greater abundance. The most abundant species, Rhinella ornata, showed no biometric differences between fragment groups but like many other amphibians sampled showed very low numbers of individuals in fragments dominated by sugar cane fields. Our data indicate that the sugar cane matrix negatively influences the community of amphibians present in fragments surrounded by this type of land use.

  5. Persistence of canine distemper virus in the Greater Yellowstone ecosystem's carnivore community.

    Science.gov (United States)

    Almberg, Emily S; Cross, Paul C; Smith, Douglas W

    2010-10-01

    Canine distemper virus (CDV) is an acute, highly immunizing pathogen that should require high densities and large populations of hosts for long-term persistence, yet CDV persists among terrestrial carnivores with small, patchily distributed groups. We used CDV in the Greater Yellowstone ecosystem's (GYE) wolves (Canis lupus) and coyotes (Canis latrans) as a case study for exploring how metapopulation structure, host demographics, and multi-host transmission affect the critical community size and spatial scale required for CDV persistence. We illustrate how host spatial connectivity and demographic turnover interact to affect both local epidemic dynamics, such as the length and variation in inter-epidemic periods, and pathogen persistence using stochastic, spatially explicit susceptible-exposed-infectious-recovered simulation models. Given the apparent absence of other known persistence mechanisms (e.g., a carrier or environmental state, densely populated host, chronic infection, or a vector), we suggest that CDV requires either large spatial scales or multi-host transmission for persistence. Current GYE wolf populations are probably too small to support endemic CDV. Coyotes are a plausible reservoir host, but CDV would still require 50000-100000 individuals for moderate persistence (> 50% over 10 years), which would equate to an area of 1-3 times the size of the GYE (60000-200000 km2). Coyotes, and carnivores in general, are not uniformly distributed; therefore, this is probably a gross underestimate of the spatial scale of CDV persistence. However, the presence of a second competent host species can greatly increase the probability of long-term CDV persistence at much smaller spatial scales. Although no management of CDV is currently recommended for the GYE, wolf managers in the region should expect periodic but unpredictable CDV-related population declines as often as every 2-5 years. Awareness and monitoring of such outbreaks will allow corresponding adjustments

  6. Elk migration patterns and human activity influence wolf habitat use in the Greater Yellowstone Ecosystem.

    Science.gov (United States)

    Nelson, Abigail A; Kauffman, Matthew J; Middleton, Arthur D; Jimenez, Michael D; McWhirter, Douglas E; Barber, Jarrett; Gerow, Kenneth

    2012-12-01

    Identifying the ecological dynamics underlying human-wildlife conflicts is important for the management and conservation of wildlife populations. In landscapes still occupied by large carnivores, many ungulate prey species migrate seasonally, yet little empirical research has explored the relationship between carnivore distribution and ungulate migration strategy. In this study, we evaluate the influence of elk (Cervus elaphus) distribution and other landscape features on wolf (Canis lupus) habitat use in an area of chronic wolf-livestock conflict in the Greater Yellowstone Ecosystem, USA. Using three years of fine-scale wolf (n = 14) and elk (n = 81) movement data, we compared the seasonal habitat use of wolves in an area dominated by migratory elk with that of wolves in an adjacent area dominated by resident elk. Most migratory elk vacate the associated winter wolf territories each summer via a 40-60 km migration, whereas resident elk remain accessible to wolves year-round. We used a generalized linear model to compare the relative probability of wolf use as a function of GIS-based habitat covariates in the migratory and resident elk areas. Although wolves in both areas used elk-rich habitat all year, elk density in summer had a weaker influence on the habitat use of wolves in the migratory elk area than the resident elk area. Wolves employed a number of alternative strategies to cope with the departure of migratory elk. Wolves in the two areas also differed in their disposition toward roads. In winter, wolves in the migratory elk area used habitat close to roads, while wolves in the resident elk area avoided roads. In summer, wolves in the migratory elk area were indifferent to roads, while wolves in resident elk areas strongly avoided roads, presumably due to the location of dens and summering elk combined with different traffic levels. Study results can help wildlife managers to anticipate the movements and establishment of wolf packs as they expand into areas

  7. Wolf presence and increased willow consumption by Yellowstone elk: implications for trophic cascades.

    Science.gov (United States)

    Creel, Scott; Christianson, David

    2009-09-01

    Recent increases in the height and growth ring width of willow (Salix spp.) and other woody plants in the Greater Yellowstone Ecosystem (GYE) have been attributed to a behaviorally mediated trophic cascade from wolves (Canis lupus) to elk (Cervus elaphus) to willows. This hypothesis predicts that individual elk consume less willow in response to the presence of wolves, but this prediction has not been directly tested with data from elk. We collected 727 fecal samples from elk in the Gallatin Canyon portion of the GYE over three winters and used microhistological methods to quantify the proportion of willow in each sample. We then tested the effect of wolf presence on willow consumption by elk, controlling for the effects of snow conditions, sex, and habitat type. During the period of study, 8-17 wolves occupied the study area, and wolves were locally present on 49% of 260 sampling days, stratified at two-week intervals across three drainages. Over the three years combined, willow consumption was related to snow conditions, wolf presence, and a wolf X sex interaction. As expected, willow consumption increased with deeper and less penetrable snow, and this effect was strong. Contrary to expectation, willow consumption increased in the presence of wolves. As with other aspects of antipredator behavior, wolves had different effects on willow consumption by males and females. Finally, we aggregated the data to estimate winter-long mean willow consumption within each drainage; at this broader scale, willow consumption again increased as predation risk increased. In summary, willow consumption was more strongly affected by snow conditions than by the presence of wolves. Interactions between elk and willow were affected by wolves, but not as predicted by the hypothesis that wolf presence favors willow release through a reduction in the selection of willow by individual elk. If a trophic cascade is operating, our results suggest that a decline in the size of the elk

  8. Persistence of canine distemper virus in the Greater Yellowstone Ecosystem's carnivore community

    Science.gov (United States)

    Almberg, E.S.; Cross, P.C.; Smith, D.W.

    2010-01-01

    Canine distemper virus (CDV) is an acute, highly immunizing pathogen that should require high densities and large populations of hosts for long-term persistence, yet CDV persists among terrestrial carnivores with small, patchily distributed groups. We used CDV in the Greater Yellowstone ecosystem's (GYE) wolves (Canis lupus) and coyotes (Canis latrans) as a case study for exploring how metapopulation structure, host demographics, and multi-host transmission affect the critical community size and spatial scale required for CDV persistence. We illustrate how host spatial connectivity and demographic turnover interact to affect both local epidemic dynamics, such as the length and variation in inter-epidemic periods, and pathogen persistence using stochastic, spatially explicit susceptible-exposed-infectious-recovered simulation models. Given the apparent absence of other known persistence mechanisms (e.g., a carrier or environmental state, densely populated host, chronic infection, or a vector), we suggest that CDV requires either large spatial scales or multi-host transmission for persistence. Current GYE wolf populations are probably too small to support endemic CDV. Coyotes are a plausible reservoir host, but CDV would still require 50 000-100 000 individuals for moderate persistence (>50% over 10 years), which would equate to an area of 1-3 times the size of the GYE (60000-200000 km2). Coyotes, and carnivores in general, are not uniformly distributed; therefore, this is probably a gross underestimate of the spatial scale of CDV persistence. However, the presence of a second competent host species can greatly increase the probability of long-term CDV persistence at much smaller spatial scales. Although no management of CDV is currently recommended for the GYE, wolf managers in the region should expect periodic but unpredictable CDV-related population declines as often as every 2-5 years. Awareness and monitoring of such outbreaks will allow corresponding

  9. Probable causes of increasing brucellosis in free-ranging elk of the Greater Yellowstone Ecosystem

    Science.gov (United States)

    Cross, P.C.; Cole, E.K.; Dobson, A.P.; Edwards, W.H.; Hamlin, K.L.; Luikart, G.; Middleton, A.D.; Scurlock, B.M.; White, P.J.

    2010-01-01

    While many wildlife species are threatened, some populations have recovered from previous overexploitation, and data linking these population increases with disease dynamics are limited. We present data suggesting that free-ranging elk (Cervus elaphus) are a maintenance host for Brucella abortus in new areas of the Greater Yellowstone Ecosystem (GYE). Brucellosis seroprevalence in free-ranging elk increased from 0-7% in 1991-1992 to 8-20% in 2006-2007 in four of six herd units around the GYE. These levels of brucellosis are comparable to some herd units where elk are artificially aggregated on supplemental feeding grounds. There are several possible mechanisms for this increase that we evaluated using statistical and population modeling approaches. Simulations of an age-structured population model suggest that the observed levels of seroprevalence are unlikely to be sustained by dispersal from supplemental feeding areas with relatively high seroprevalence or an older age structure. Increases in brucellosis seroprevalence and the total elk population size in areas with feeding grounds have not been statistically detectable. Meanwhile, the rate of seroprevalence increase outside the feeding grounds was related to the population size and density of each herd unit. Therefore, the data suggest that enhanced elk-to-elk transmission in free-ranging populations may be occurring due to larger winter elk aggregations. Elk populations inside and outside of the GYE that traditionally did not maintain brucellosis may now be at risk due to recent population increases. In particular, some neighboring populations of Montana elk were 5-9 times larger in 2007 than in the 1970s, with some aggregations comparable to the Wyoming feeding-ground populations. Addressing the unintended consequences of these increasing populations is complicated by limited hunter access to private lands, which places many ungulate populations out of administrative control. Agency-landowner hunting access

  10. Elk migration patterns and human activity influence wolf habitat use in the Greater Yellowstone Ecosystem

    Science.gov (United States)

    Nelson, Abigail; Kauffman, Matthew J.; Middleton, Arthur D.; Jimenez, Mike; McWhirter, Douglas; Barber, Jarrett; Gerow, Ken

    2012-01-01

    Identifying the ecological dynamics underlying human–wildlife conflicts is important for the management and conservation of wildlife populations. In landscapes still occupied by large carnivores, many ungulate prey species migrate seasonally, yet little empirical research has explored the relationship between carnivore distribution and ungulate migration strategy. In this study, we evaluate the influence of elk (Cervus elaphus) distribution and other landscape features on wolf (Canis lupus) habitat use in an area of chronic wolf–livestock conflict in the Greater Yellowstone Ecosystem, USA. Using three years of fine-scale wolf (n = 14) and elk (n = 81) movement data, we compared the seasonal habitat use of wolves in an area dominated by migratory elk with that of wolves in an adjacent area dominated by resident elk. Most migratory elk vacate the associated winter wolf territories each summer via a 40–60 km migration, whereas resident elk remain accessible to wolves year-round. We used a generalized linear model to compare the relative probability of wolf use as a function of GIS-based habitat covariates in the migratory and resident elk areas. Although wolves in both areas used elk-rich habitat all year, elk density in summer had a weaker influence on the habitat use of wolves in the migratory elk area than the resident elk area. Wolves employed a number of alternative strategies to cope with the departure of migratory elk. Wolves in the two areas also differed in their disposition toward roads. In winter, wolves in the migratory elk area used habitat close to roads, while wolves in the resident elk area avoided roads. In summer, wolves in the migratory elk area were indifferent to roads, while wolves in resident elk areas strongly avoided roads, presumably due to the location of dens and summering elk combined with different traffic levels. Study results can help wildlife managers to anticipate the movements and establishment of wolf packs as they expand into

  11. Placental vascular responses are dependent on surrounding tissue

    DEFF Research Database (Denmark)

    Brøgger, Torbjørn Halle

    -depth understanding of the mechanism regulating blood flow and perfusion is necessary if we are to come up with new ideas for intervention and treatment. Method: From fresh born placentas stem villi arteries were carefully dissected. The artery branches were divided. The surrounding tissue was removed from one end...... and was left untouched in the other end. Then using wire myography they were investigated in terms of contractility and sensitivity to physiological relevant human-like agonists. Results: Sensitivity to PGF2α, Tx-analog, 5-HT and endothelin-1 was significantly lower in arteries with intact surrounding tissue...... compared to arteries stripped of the tissue. The maximal force development was also significantly lower in arteries with surrounding tissue, when they were depolarized high extracellular [K+] or stimulated with PGF2α or endotheline-1. Conclusion: The perivascular tissue significantly alters stem villi...

  12. Placental vascular responses are dependent on surrounding tissue

    DEFF Research Database (Denmark)

    Brøgger, Torbjørn Halle

    . Materials and methods. From fresh born placentas, stem villi arteries were carefully dissected. The artery branches were divided. The surrounding tissue was removed from one end and was left untouched in the other end.Then, using wire myography, they were investigated in terms of contractility...... and sensitivity to physiological relevant human-like agonists. Results. Sensitivity to PGF2α, Tx-analog, 5-HT and endothelin-1 was significantly lower in arteries with intact surrounding tissue compared to arteries stripped of the tissue. The maximal force development was also significantly lower in arteries...... with surrounding tissue when they were depolarized high extracellular [K+] or stimulated with PGF2α or endotheline-1. Conclusion. The perivascular tissue significantly alters stem villi arteries' sensitivity and force development in a suppressive way. This implicates a new aspect of blood flow regulation...

  13. Surrounding rock stress analysis of underground high level waste repository

    International Nuclear Information System (INIS)

    Liu Wengang; Wang Ju; Wang Guangdi

    2006-01-01

    During decay of nuclear waste, enormous energy was released, which results in temperature change of surrounding rock of depository. Thermal stress was produced because thermal expansion of rock was controlled. Internal structure of surrounding rock was damaged and strength of rock was weakened. So, variation of stress was a dynamic process with the variation of temperature. BeiShan region of Gansu province was determined to be the depository field in the future, it is essential to make research on granite in this region. In the process of experiment, basic physical parameters of granite were analyzed preliminary with MTS. Long range temperature and stress filed was simulated considering the damage effect of surrounding rock, and rules of temperature and stress was achieved. (authors)

  14. Radiological impact of phosphogypsum in the Surrounding Ecosystem

    International Nuclear Information System (INIS)

    Al-Attar, L.; Al-Oudat, M.; Budier, Y.; Khalili, H.; Hamwi, A.; Kanakri, S.

    2011-01-01

    This study was carried out to assess the radiological impact of Syrian PG piles in the compartments of the surrounding ecosystem. Therefore, estimating the distribution of naturally occurring radionuclides (i.e. 2 26 Ra, 2 38 U, 2 32 Th, 2 10 Po and 2 10 Pb) in the raw materials, product and by-product of the Syrian phosphate fertilizer industry was essential. The obtained data revealed that 2 26 Ra retained in PG with a mean activity of 318 Bq kg-1. Uranium content in PG was low since it remained in the produced phosphoric acid. However, over 80% of 2 32 Th, 2 10 Po and 2 10 Pb partitioned in PG. The presence of PG piles did not increase the concentration of 2 22 Rn nor gamma rays exposure dose in the studied area. The annual effective dose was only 0.082 mSv y-1. The geometric mean of total suspended solids was ca. 85 g m-3. The concentration of the radionuclides in filtration and runoff waters were below the detection limits; and were much lower than the permissible limits set for drinking water by the WHO in ground and Qattina Lake waters. Eastern sites soil samples of PG piles were of the highest activity concentrations, due to the characterised western and north-western wind in the area, but remained within the natural levels reported in Syrian soil. The impact of PG piles on plants varied upon the plant species. Significantly, higher concentrations of the radionuclides were recorded for grass in comparison to broad-leaved plants. Among the species that naturally grown on PG piles, Inula, Ecballium and Polygonium may be radionuclides accumulators. Nevertheless, a determined effort is needed on national level to achieve a common and coherent approach to regulate PG piles or to consider it a resource material rather than waste or residue. The presence of PG piles did not increase the concentration of 2 22 Rn nor gamma rays exposure dose in the studied area. The annual effective dose was only 0.082 mSv y -1 . The geometric mean of total suspended solids was ca

  15. Ground penetrating radar documents short-term near-surface hydrological changes around Old Faithful Geyser, Yellowstone National Park, USA

    Science.gov (United States)

    Lynne, Bridget Y.; Heasler, Henry; Jaworowski, Cheryl; Smith, Gary J.; Smith, Isaac J.; Foley, Duncan

    2018-04-01

    In April 2015, Ground Penetrating Radar (GPR) was used to characterize the shallow subsurface (images were collected between two eruptions of Old Faithful Geyser. Each set of time-sequence GPR recordings consisted of four transects aligned to provide coverage near the potential location of the inferred 15 m deep geyser chamber. However, the deepest penetration we could achieve with a 200 MHz GPR antennae was 5 m. Seven time-sequence events were collected over a 48-minute interval to image changes in the near-surface, during pre- and post-eruptive cycles. Time-sequence GPR images revealed a series of possible micro-fractures in a highly porous siliceous sinter in the near-surface that fill and drain repetitively, immediately after an eruption and during the recharge period prior to the next main eruptive event.

  16. Behavior of nuclear waste elements during hydrothermal alteration of glassy rhyolite in an active geothermal system: Yellowstone National Park, Wyoming

    International Nuclear Information System (INIS)

    Sturchio, N.C.; Seitz, M.G.

    1984-01-01

    The behavior of a group of nuclear waste elements (U, Th, Sr, Zr, Sb, Cs, Ba, and Sm) during hydrothermal alteration of glassy rhyolite is investigated through detailed geochemical analyses of whole rocks, glass and mineral separates, and thermal waters. Significant mobility of U, Sr, Sb, Cs, and Ba is found, and the role of sorption processes in their observed behavior is identified. Th, Zr, and Sm are relatively immobile, except on a microscopic scale. 9 references, 2 figures, 2 tables

  17. Chromosome rearrangements, recombination suppression, and limited segregation distortion in hybrids between Yellowstone cutthroat trout (Oncorhynchus clarkii bouvieri) and rainbow trout (O. mykiss)

    Science.gov (United States)

    2013-01-01

    Background Introgressive hybridization is an important evolutionary process that can lead to the creation of novel genome structures and thus potentially new genetic variation for selection to act upon. On the other hand, hybridization with introduced species can threaten native species, such as cutthroat trout (Oncorhynchus clarkii) following the introduction of rainbow trout (O. mykiss). Neither the evolutionary consequences nor conservation implications of rainbow trout introgression in cutthroat trout is well understood. Therefore, we generated a genetic linkage map for rainbow-Yellowstone cutthroat trout (O. clarkii bouvieri) hybrids to evaluate genome processes that may help explain how introgression affects hybrid genome evolution. Results The hybrid map closely aligned with the rainbow trout map (a cutthroat trout map does not exist), sharing all but one linkage group. This linkage group (RYHyb20) represented a fusion between an acrocentric (Omy28) and a metacentric chromosome (Omy20) in rainbow trout. Additional mapping in Yellowstone cutthroat trout indicated the two rainbow trout homologues were fused in the Yellowstone genome. Variation in the number of hybrid linkage groups (28 or 29) likely depended on a Robertsonian rearrangement polymorphism within the rainbow trout stock. Comparison between the female-merged F1 map and a female consensus rainbow trout map revealed that introgression suppressed recombination across large genomic regions in 5 hybrid linkage groups. Two of these linkage groups (RYHyb20 and RYHyb25_29) contained confirmed chromosome rearrangements between rainbow and Yellowstone cutthroat trout indicating that rearrangements may suppress recombination. The frequency of allelic and genotypic segregation distortion varied among parents and families, suggesting few incompatibilities exist between rainbow and Yellowstone cutthroat trout genomes. Conclusions Chromosome rearrangements suppressed recombination in the hybrids. This result

  18. Ornitocenosis of the Sursky pond and its close the surroundings

    International Nuclear Information System (INIS)

    Lacko, J.; Ambrus, B.; Fupso, A.

    2013-01-01

    The paper focuses on the qualitative-quantitative research of fishpond bird community of the Sursky and its surroundings as well as on analysis of seasonal population dynamics of the avifauna as well as on placing the determined species into environmental groups and guilds. Another object is the comparison of our results with recent work focused on research of bird communities on this site.

  19. Review of Ordered Anarchy: Jasay and his Surroundings

    Directory of Open Access Journals (Sweden)

    Aschwin de Wolf

    2009-02-01

    Full Text Available Anthony de Jasay is among the most important social thinkers of our time. His oeuvre offers a sustained critique of government and its defenders. In the book Ordered Anarchy: Jasay and His Surroundings, colleagues and friends pay tribute to the man in the form of an inspiring collection of essays.

  20. Neutron spectrum in small iron pile surrounded by lead reflector

    International Nuclear Information System (INIS)

    Kimura, Itsuro; Hayashi, S.A.; Kobayashi, Katsuhei; Matsumura, Tetsuo; Nishihara, Hiroshi.

    1978-01-01

    In order to save the quantity of sample material, a possibility to assess group constants of a reactor material through measurement and analysis of neutron spectrum in a small sample pile surrounded by a reflector of heavy moderator, was investigated. As the sample and the reflector, we chose iron and lead, respectively. Although the time dispersion in moderation of neutrons was considerably prolonged by the lead reflector, this hardly interferes with the assessment of group constants. Theoretical calculation revealed that both the neutron flux spectrum and the sensitivity coefficient of group constants in an iron sphere, 35 cm in diameter surrounded by the lead reflector, 25 cm thick, were close to those of the bare iron sphere, 108 cm in diameter. The neutron spectra in a small iron pile surrounded by a lead reflector were experimentally obtained by the time-of-flight method with an electron linear accelerator and the result was compared with the predicted values. It could be confirmed that a small sample pile surrounded by a reflector, such as lead, was as useful as a much larger bulk pile for the assessment of group constants of a reactor material. (auth.)

  1. Linking disadvantaged housing areas to the surrounding city

    DEFF Research Database (Denmark)

    Stender, Marie

    Several disadvantaged social housing areas in Denmark are currently undergo-ing thorough physical refurbishments, aiming to integrate them better with the surrounding city. The ambition is to attract new users and residents by opening up the borders of the area and establish attractive, new...

  2. Ecological mechanisms linking protected areas to surrounding lands.

    Science.gov (United States)

    Hansen, Andrew J; DeFries, Ruth

    2007-06-01

    Land use is expanding and intensifying in the unprotected lands surrounding many of the world's protected areas. The influence of this land use change on ecological processes is poorly understood. The goal of this paper is to draw on ecological theory to provide a synthetic framework for understanding how land use change around protected areas may alter ecological processes and biodiversity within protected areas and to provide a basis for identifying scientifically based management alternatives. We first present a conceptual model of protected areas embedded within larger ecosystems that often include surrounding human land use. Drawing on case studies in this Invited Feature, we then explore a comprehensive set of ecological mechanisms by which land use on surrounding lands may influence ecological processes and biodiversity within reserves. These mechanisms involve changes in ecosystem size, with implications for minimum dynamic area, species-area effect, and trophic structure; altered flows of materials and disturbances into and out of reserves; effects on crucial habitats for seasonal and migration movements and population source/sink dynamics; and exposure to humans through hunting, poaching, exotics species, and disease. These ecological mechanisms provide a basis for assessing the vulnerability of protected areas to land use. They also suggest criteria for designing regional management to sustain protected areas in the context of surrounding human land use. These design criteria include maximizing the area of functional habitats, identifying and maintaining ecological process zones, maintaining key migration and source habitats, and managing human proximity and edge effects.

  3. The bird species of pandam wildlife park and the surrounding ...

    African Journals Online (AJOL)

    The effect of time of day as well as vegetation variables on bird species diversity in the park and surrounding farmlands was also conducted. 10 transects in each study site were surveyed twice between during the dry season and vegetation variables (trees, fingers, finger-rings two- hand, grazing, farming, canopy cover, ...

  4. Experiences during the decontamination process of areas surrounding to Fukushima

    International Nuclear Information System (INIS)

    Molina, G.

    2014-10-01

    In this work the experience gained during the decontamination of areas surrounding to Fukushima NPP, rugged during the earthquake and tsunami in 2011 and caused the contamination with fission products in these areas is described. Actions taken by the Japanese government are reported and some of the techniques used, the intervention levels and the progress made and disposal techniques considered are presented. (Author)

  5. Traditional Indian customs surrounding birth A review | Chalmers ...

    African Journals Online (AJOL)

    Since 1960, only a few studies have been made of traditional custOIns surrounding birth in Indian culture. Very few of these have described customs followed by Indians in South Africa. A review of these publications is presented here. Customs described include religious, social and psychological aspects of behaviour in ...

  6. Mechanisms and timescales of generating eruptible rhyolitic magmas at Yellowstone caldera from zircon and sanidine geochronology and geochemistry

    Science.gov (United States)

    Stelten, Mark; Cooper, Kari M.; Vazquez, Jorge A.; Calvert, Andrew T.; Glessner, Justin G

    2015-01-01

    We constrain the physical nature of the magma reservoir and the mechanisms of rhyolite generation at Yellowstone caldera via detailed characterization of zircon and sanidine crystals hosted in three rhyolites erupted during the (ca. 170 – 70 ka) Central Plateau Member eruptive episode – the most recent post-caldera magmatism at Yellowstone. We present 238U-230Th crystallization ages and trace-element compositions of the interiors and surfaces (i.e., unpolished rims) of individual zircon crystals from each rhyolite. We compare these zircon data to 238U- 230Th crystallization ages of bulk sanidine separates coupled with chemical and isotopic data from single sanidine crystals. Zircon age and trace-element data demonstrate that the magma reservoir that sourced the Central Plateau Member rhyolites was long-lived (150 – 250 kyr) and genetically related to the preceding episode of magmatism, which occurred ca. 256 ka. The interiors of most zircons in each rhyolite were inherited from unerupted material related to older stages of Central Plateau Member magmatism or the preceding late Upper Basin Member magmatism (i.e., are antecrysts). Conversely, most zircon surfaces crystallized near the time of eruption from their host liquids (i.e., are autocrystic). The repeated recycling of zircon interiors from older stages of magmatism demonstrates that sequentially erupted Central Plateau Member rhyolites are genetically related. Sanidine separates from each rhyolite yield 238U-230Th crystallization ages at or near the eruption age of their host magmas, coeval with the coexisting zircon surfaces, but are younger than the coexisting zircon interiors. Chemical and isotopic data from single sanidine crystals demonstrate that the sanidines in each rhyolite are in equilibrium with their host melts, which considered along with their near-eruption crystallization ages suggests that nearly all CPM sanidines are autocrystic. The paucity of antecrystic sanidine crystals relative to

  7. Re-evaluation of Yellowstone grizzly bear population dynamics not supported by empirical data: response to Doak & Cutler

    Science.gov (United States)

    van Manen, Frank T.; Ebinger, Michael R.; Haroldson, Mark A.; Harris, Richard B.; Higgs, Megan D.; Cherry, Steve; White, Gary C.; Schwartz, Charles C.

    2014-01-01

    Doak and Cutler critiqued methods used by the Interagency Grizzly Bear Study Team (IGBST) to estimate grizzly bear population size and trend in the Greater Yellowstone Ecosystem. Here, we focus on the premise, implementation, and interpretation of simulations they used to support their arguments. They argued that population increases documented by IGBST based on females with cubs-of-the-year were an artifact of increased search effort. However, we demonstrate their simulations were neither reflective of the true observation process nor did their results provide statistical support for their conclusion. They further argued that survival and reproductive senescence should be incorporated into population projections, but we demonstrate their choice of extreme mortality risk beyond age 20 and incompatible baseline fecundity led to erroneous conclusions. The conclusions of Doak and Cutler are unsubstantiated when placed within the context of a thorough understanding of the data, study system, and previous research findings and publications.

  8. Vasculature surrounding a nodule: A novel lung cancer biomarker.

    Science.gov (United States)

    Wang, Xiaohua; Leader, Joseph K; Wang, Renwei; Wilson, David; Herman, James; Yuan, Jian-Min; Pu, Jiantao

    2017-12-01

    To investigate whether the vessels surrounding a nodule depicted on non-contrast, low-dose computed tomography (LDCT) can discriminate benign and malignant screen detected nodules. We collected a dataset consisting of LDCT scans acquired on 100 subjects from the Pittsburgh Lung Screening study (PLuSS). Fifty subjects were diagnosed with lung cancer and 50 subjects had suspicious nodules later proven benign. For the lung cancer cases, the location of the malignant nodule in the LDCT scans was known; while for the benign cases, the largest nodule in the LDCT scan was used in the analysis. A computer algorithm was developed to identify surrounding vessels and quantify the number and volume of vessels that were connected or near the nodule. A nonparametric receiver operating characteristic (ROC) analysis was performed based on a single nodule per subject to assess the discriminability of the surrounding vessels to provide a lung cancer diagnosis. Odds ratio (OR) were computed to determine the probability of a nodule being lung cancer based on the vessel features. The areas under the ROC curves (AUCs) for vessel count and vessel volume were 0.722 (95% CI=0.616-0.811, plung cancer group 9.7 (±9.6) compared to the non-lung cancer group 4.0 (±4.3) CONCLUSION: Our preliminary results showed that malignant nodules are often surrounded by more vessels compared to benign nodules, suggesting that the surrounding vessel characteristics could serve as lung cancer biomarker for indeterminate nodules detected during LDCT lung cancer screening using only the information collected during the initial visit. Copyright © 2017 Elsevier B.V. All rights reserved.

  9. Neogene fallout tuffs from the Yellowstone hotspot in the Columbia Plateau region, Oregon, Washington and Idaho, USA.

    Directory of Open Access Journals (Sweden)

    Barbara P Nash

    Full Text Available Sedimentary sequences in the Columbia Plateau region of the Pacific Northwest ranging in age from 16-4 Ma contain fallout tuffs whose origins lie in volcanic centers of the Yellowstone hotspot in northwestern Nevada, eastern Oregon and the Snake River Plain in Idaho. Silicic volcanism began in the region contemporaneously with early eruptions of the Columbia River Basalt Group (CRBG, and the abundance of widespread fallout tuffs provides the opportunity to establish a tephrostratigrahic framework for the region. Sedimentary basins with volcaniclastic deposits also contain diverse assemblages of fauna and flora that were preserved during the Mid-Miocene Climatic Optimum, including Sucker Creek, Mascall, Latah, Virgin Valley and Trout Creek. Correlation of ashfall units establish that the lower Bully Creek Formation in eastern Oregon is contemporaneous with the Virgin Valley Formation, the Sucker Creek Formation, Oregon and Idaho, Trout Creek Formation, Oregon, and the Latah Formation in the Clearwater Embayment in Washington and Idaho. In addition, it can be established that the Trout Creek flora are younger than the Mascall and Latah flora. A tentative correlation of a fallout tuff from the Clarkia fossil beds, Idaho, with a pumice bed in the Bully Creek Formation places the remarkably well preserved Clarkia flora assemblage between the Mascall and Trout Creek flora. Large-volume supereruptions that originated between 11.8 and 10.1 Ma from the Bruneau-Jarbidge and Twin Falls volcanic centers of the Yellowstone hotspot in the central Snake River Plain deposited voluminous fallout tuffs in the Ellensberg Formation which forms sedimentary interbeds in the CRBG. These occurrences extend the known distribution of these fallout tuffs 500 km to the northwest of their source in the Snake River Plain. Heretofore, the distal products of these large eruptions had only been recognized to the east of their sources in the High Plains of Nebraska and Kansas.

  10. 238U-230Th dating of chevkinite in high-silica rhyolites from La Primavera and Yellowstone calderas

    Science.gov (United States)

    Vazquez, Jorge A.; Velasco, Noel O.; Schmitt, Axel K.; Bleick, Heather A.; Stelten, Mark E.

    2014-01-01

    Application of 238U-230Th disequilibrium dating of accessory minerals with contrasting stabilities and compositions can provide a unique perspective on magmatic evolution by placing the thermochemical evolution of magma within the framework of absolute time. Chevkinite, a Th-rich accessory mineral that occurs in peralkaline and metaluminous rhyolites, may be particularly useful as a chronometer of crystallization and differentiation because its composition may reflect the chemical changes of its host melt. Ion microprobe 128U-230Th dating of single chevkinite microphenocrysts from pre- and post-caldera La Primavera, Mexico, rhyolites yields model crystallization ages that are within 10's of k.y. of their corresponding K-Ar ages of ca. 125 ka to 85 ka, while chevkinite microphenocrysts from a post-caldera Yellowstone, USA, rhyolite yield a range of ages from ca. 110 ka to 250 ka, which is indistinguishable from the age distribution of coexisting zircon. Internal chevkinite-zircon isochrons from La Primavera yield Pleistocene ages with ~5% precision due to the nearly two order difference in Th/U between both minerals. Coupling chevkinite 238U-230Th ages and compositional analyses reveals a secular trend of Th/U and rare earth elements recorded in Yellowstone rhyolite, likely reflecting progressive compositional evolution of host magma. The relatively short timescale between chevkinite-zircon crystallization and eruption suggests that crystal-poor rhyolites at La Primavera were erupted shortly after differentiation and/or reheating. These results indicate that 238U-230Th dating of chevkinite via ion microprobe analysis may be used to date crystallization and chemical evolution of silicic magmas.

  11. Evaluation of the evolving stress field of the Yellowstone volcanic plateau, 1988 to 2010, from earthquake first-motion inversions

    Science.gov (United States)

    Russo, E.; Waite, G. P.; Tibaldi, A.

    2017-03-01

    Although the last rhyolite eruption occurred around 70 ka ago, the silicic Yellowstone volcanic field is still considered active due to high hydrothermal and seismic activity and possible recent magma intrusions. Geodetic measurements document complex deformation patterns in crustal strain and seismic activity likewise reveal spatial and temporal variations in the stress field. We use earthquake data recorded between 1988 and 2010 to investigate these variations and their possible causes in more detail. Earthquake relocations and a set of 369 well-constrained, double-couple, focal mechanism solutions were computed. Events were grouped according to location and time to investigate trends in faulting. The majority of the events have normal-faulting solutions, subordinate strike-slip kinematics, and very rarely, reverse motions. The dominant direction of extension throughout the 0.64 Ma Yellowstone caldera is nearly ENE, consistent with the perpendicular direction of alignments of volcanic vents within the caldera, but our study also reveals spatial and temporal variations. Stress-field solutions for different areas and time periods were calculated from earthquake focal mechanism inversion. A well-resolved rotation of σ3 was found, from NNE-SSW near the Hebgen Lake fault zone, to ENE-WSW near Norris Junction. In particular, the σ3 direction changed throughout the years around Norris Geyser Basin, from being ENE-WSW, as calculated in the study by Waite and Smith (2004), to NNE-SSW, while the other σ3 directions are mostly unchanged over time. The presence of ;chocolate tablet; structures, with two sets of nearly perpendicular normal faults, was identified in many stages of the deformation history both in the Norris Geyser Basin area and inside the caldera.

  12. Investigation of 1-D crustal velocity structure beneath Izmir Gulf and surroundings by using local earthquakes

    International Nuclear Information System (INIS)

    Polat, Orhan; Özer, Çaglar

    2016-01-01

    In this study; we examined one dimensional crustal velocity structure of Izmir gulf and surroundings. We used nearly one thousand high quality (A and B class) earthquake data which recorded by Disaster and Emergency Management Presidency (AFAD) [1], Bogazici University (BU-KOERI) [2] and National Observatory of Athens (NOA) [3,4]. We tried several synthetic tests to understand power of new velocity structure, and examined phase residuals, RMS values and shifting tests. After evaluating these tests; we decided one dimensional velocity structure and minimum 1-D P wave velocities, hypocentral parameter and earthquake locations from VELEST algorithm. Distribution of earthquakes was visibly improved by using new minimum velocity structure.

  13. Vegetation cover and land use of a protected coastal area and its surroundings, southeast Brazil

    OpenAIRE

    Caris,Elisa Araujo Penna; Kurtz,Bruno Coutinho; Cruz,Carla Bernadete Madureira; Scarano,Fabio Rubio

    2013-01-01

    We applied remote sensing techniques on a TM Landsat 5 image (1:50,000) to map land use and vegetation cover of the Restinga de Jurubatiba National Park and surroundings. The thematic map generated from the digital classification of the image allowed us to spatially characterize and quantify the different land uses and soil covers of the area. Thirteen classes were identified. The most representative classes in the park were the Clusia (31.99%) and Ericaceae formations (29.14%). More than 90%...

  14. Investigation of 1-D crustal velocity structure beneath Izmir Gulf and surroundings by using local earthquakes

    Energy Technology Data Exchange (ETDEWEB)

    Polat, Orhan, E-mail: orhan.polat@deu.edu.tr [Dokuz Eylul University, Faculty of Engineering, Geophysical Engineering Department, Izmir (Turkey); Özer, Çaglar, E-mail: caglar.ozer@deu.edu.tr [Dokuz Eylul University, Faculty of Engineering, Geophysical Engineering Department, Izmir (Turkey); Dokuz Eylul University, The Graduate School of Natural and Applied Sciences, Department of Geophysical Engineering, Izmir-Turkey (Turkey)

    2016-04-18

    In this study; we examined one dimensional crustal velocity structure of Izmir gulf and surroundings. We used nearly one thousand high quality (A and B class) earthquake data which recorded by Disaster and Emergency Management Presidency (AFAD) [1], Bogazici University (BU-KOERI) [2] and National Observatory of Athens (NOA) [3,4]. We tried several synthetic tests to understand power of new velocity structure, and examined phase residuals, RMS values and shifting tests. After evaluating these tests; we decided one dimensional velocity structure and minimum 1-D P wave velocities, hypocentral parameter and earthquake locations from VELEST algorithm. Distribution of earthquakes was visibly improved by using new minimum velocity structure.

  15. Nutritional and Microbial Parameters of Earthworm Cast, Termite Mound and Surrounding Bulk Soil

    OpenAIRE

    Kawaguchi, Sadao; Nishi, Shingo

    2007-01-01

    A comparative analysis of nutritional and microbial parameters was conducted on two types of biogenetic structures of earthworm cast (8.7 cm in height, 7 casts/1m×1m) formed by litter eating Pheretima sp., and mound (64 cm in height, 1.0 mounds/10m×50m) built by fungus growing termite, Macrotermes gilvus, and compared to the surrounding bulk soil as control in the tropical monsoon forest in Cu Chi National Park of Viet Nam. The proportion of the sand in the earthworm cast was higher than in t...

  16. Identificação de conflito de uso da terra em áreas de preservação permanente no entorno do parque nacional do Caparaó, Estado de Minas Gerais Identification of land use conflicts in permanent preservation áreas in the surroundings of the Caparaó national park, Minas Gerais State, Brazil

    Directory of Open Access Journals (Sweden)

    Fernando Soares de Oliveira

    2008-10-01

    Full Text Available Este estudo teve como objetivos elaborar um mapa de uso da terra com base nas imagens do satélite IKONOS II, delimitar de maneira automática as áreas de preservação permanente e identificar a ocorrência de conflitos de usos, tendo como referência legal o Código florestal e a Resolução n.º 303 do CONAMA. A pesquisa foi desenvolvida na entorno do Parque Nacional do Caparaó, pertencente aos municípios de Alto Jequitibá, Alto Caparaó, Caparaó e Espera Feliz, todos situados no estado de Minas Gerais. Utilizando os recursos disponíveis no geoprocessamento, foi possível mapear 8 classes de uso da terra e delimitar as áreas de preservação permanente situadas em áreas com altitudes superior a 1.800 metros (8,42 ha, no terço superior dos morros (18,67 ha; encostas com declividade superior a 45 graus (92,96 ha; nascentes e suas respectivas áreas de contribuição (1.989,44 ha; margens dos cursos d´água com largura inferior a 10 metros (3.957,19 ha; e no terço superior das sub-bacias (6.031,54 ha, perfazendo um total de 12.098,22 ha (48,06% da área total da bacia. A área de uso indevido correspondeu a 8.922,91 ha (73,75%, sendo as classes cafezal (5.183,43 ha e pastagem (3.650,74 ha as principais ocorrências nessas áreas. Apenas 2.160,69 ha (18,40% das áreas de preservação permanente estão protegidas por vegetação nativa.The objective of this work was to prepare a map of land use based on the IKONOS II sattelite images, to automatically determine the permanent preservation areas and to identify the occurrence of use conflicts, using as legal reference the Forest Code and the nº 303 Resolution of the CONAMA. The research was developed around the Caparaó National Park, in lands of the municipalities of Alto Jequitibá, Alto Caparaó, Caparaó and Espera Feliz, all of them located in the State of Minas Gerais. Using the resources available in the geographic processing, it was possible to map 8 classes of land use and to

  17. Ultrastructural relationship of the phagophore with surrounding organelles.

    Science.gov (United States)

    Biazik, Joanna; Ylä-Anttila, Päivi; Vihinen, Helena; Jokitalo, Eija; Eskelinen, Eeva-Liisa

    2015-01-01

    Phagophore nucleates from a subdomain of the endoplasmic reticulum (ER) termed the omegasome and also makes contact with other organelles such as mitochondria, Golgi complex, plasma membrane and recycling endosomes during its formation. We have used serial block face scanning electron microscopy (SB-EM) and electron tomography (ET) to image phagophore biogenesis in 3 dimensions and to determine the relationship between the phagophore and surrounding organelles at high resolution. ET was performed to confirm whether membrane contact sites (MCSs) are evident between the phagophore and those surrounding organelles. In addition to the known contacts with the ER, we identified MCSs between the phagophore and membranes from putative ER exit sites, late endosomes or lysosomes, the Golgi complex and mitochondria. We also show that one phagophore can have simultaneous MCSs with more than one organelle. Future membrane flux experiments are needed to determine whether membrane contacts also signify lipid translocation.

  18. Trajectories and Maneuvers of Surrounding Vehicles with Panoramic Camera Arrays

    DEFF Research Database (Denmark)

    Dueholm, Jacob Velling; Kristoffersen, Miklas Strøm; Satzoda, Ravi K.

    2016-01-01

    Vision-based research for intelligent vehicles have traditionally focused on specific regions around a vehicle, such as a front looking camera for, e.g., lane estimation. Traffic scenes are complex and vital information could be lost in unobserved regions. This paper proposes a framework that uses...... four visual sensors for a full surround view of a vehicle in order to achieve an understanding of surrounding vehicle behaviors. The framework will assist the analysis of naturalistic driving studies by automating the task of data reduction of the observed trajectories. To this end, trajectories...... are estimated using a vehicle detector together with a multiperspective optimized tracker in each view. The trajectories are transformed to a common ground plane, where they are associated between perspectives and analyzed to reveal tendencies around the ego-vehicle. The system is tested on sequences from 2.5 h...

  19. Mechanical Characteristics Analysis of Surrounding Rock on Anchor Bar Reinforcement

    Science.gov (United States)

    Gu, Shuan-cheng; Zhou, Pan; Huang, Rong-bin

    2018-03-01

    Through the homogenization method, the composite of rock and anchor bar is considered as the equivalent material of continuous, homogeneous, isotropic and strength parameter enhancement, which is defined as reinforcement body. On the basis of elasticity, the composite and the reinforcement are analyzed, Based on strengthening theory of surrounding rock and displacement equivalent conditions, the expression of reinforcement body strength parameters and mechanical parameters is deduced. The example calculation shows that the theoretical results are close to the results of the Jia-mei Gao[9], however, closer to the results of FLAC3D numerical simulation, it is proved that the model and surrounding rock reinforcement body theory are reasonable. the model is easy to analyze and calculate, provides a new way for determining reasonable bolt support parameters, can also provides reference for the stability analysis of underground cavern bolting support.

  20. New geologic evidence for additional 16.5-15.5 Ma silicic calderas in northwest Nevada related to initial impingement of the Yellowstone hot spot

    International Nuclear Information System (INIS)

    Coble, Matthew A; Mahood, Gail A

    2008-01-01

    Three silicic calderas have been newly identified in northwest Nevada west of McDermitt caldera. This volcanism is interpreted to have formed during a short interval at 16.5-15.5 Ma, during the waning stage of Steens flood basalt volcanism after the initial impingement of the Yellowstone hot spot. New mapping demonstrates that the area affected by this mid-Miocene silicic volcanism is significantly larger than previously appreciated in the western U.S.

  1. Diagnostic single nucleotide polymorphisms for identifying westslope cutthroat trout (Oncorhynchus clarki lewisi), Yellowstone cutthroat trout (Oncorhynchus clarkii bouvieri) and rainbow trout (Oncorhynchus mykiss).

    Science.gov (United States)

    Kalinowski, S T; Novak, B J; Drinan, D P; Jennings, R deM; Vu, N V

    2011-03-01

    We describe 12 diagnostic single nucleotide polymorphism (SNP) assays for use in species identification among rainbow and cutthroat trout: five of these loci have alleles unique to rainbow trout (Oncorhynchus mykiss), three unique to westslope cutthroat trout (O. clarkii lewisi) and four unique to Yellowstone cutthroat trout (O. clarkii bouvieri). These diagnostic assays were identified using a total of 489 individuals from 26 populations and five fish hatchery strains. © 2010 Blackwell Publishing Ltd.

  2. New geologic evidence for additional 16.5-15.5 Ma silicic calderas in northwest Nevada related to initial impingement of the Yellowstone hot spot

    Energy Technology Data Exchange (ETDEWEB)

    Coble, Matthew A; Mahood, Gail A [Department Geological and Environmental Sciences, 450 Serra Mall, Bldg 320, Stanford University, Stanford, CA 94305-3115 (United States)

    2008-10-01

    Three silicic calderas have been newly identified in northwest Nevada west of McDermitt caldera. This volcanism is interpreted to have formed during a short interval at 16.5-15.5 Ma, during the waning stage of Steens flood basalt volcanism after the initial impingement of the Yellowstone hot spot. New mapping demonstrates that the area affected by this mid-Miocene silicic volcanism is significantly larger than previously appreciated in the western U.S.

  3. Diversity of vascular plants of Piestany and surroundings (presentation)

    International Nuclear Information System (INIS)

    Penzesova, A.; Galusova, T.

    2013-01-01

    In this presentation is a summary of the results of floristic research aimed at determining diversity of vascular plants of Piestany and its surroundings. Plant taxa we determined using the designation keys. We have compiled a list of plant species occurring in the monitored area, we evaluated the selected botanical-phytogeographical characteristics of flora, we've put together a list of local protected, endangered and rare species and a list of local invasive and expansive species according to sources. (Authors)

  4. Methods of Assessing Noise Nuisance of Real Estate Surroundings

    Directory of Open Access Journals (Sweden)

    Szopińska Kinga

    2016-03-01

    Full Text Available Testing what factors create the market value of real estate is key information when preparing property valuations as well as other opinions and professional evaluations on the basis of which court verdicts or administrative decisions are made. One of the factors influencing the value of some real estate is the level of noise present in the surroundings, which can lead to the occurrence of noise nuisance negatively affecting social relations.

  5. Numerical Simulation on Zonal Disintegration in Deep Surrounding Rock Mass

    Directory of Open Access Journals (Sweden)

    Xuguang Chen

    2014-01-01

    Full Text Available Zonal disintegration have been discovered in many underground tunnels with the increasing of embedded depth. The formation mechanism of such phenomenon is difficult to explain under the framework of traditional rock mechanics, and the fractured shape and forming conditions are unclear. The numerical simulation was carried out to research the generating condition and forming process of zonal disintegration. Via comparing the results with the geomechanical model test, the zonal disintegration phenomenon was confirmed and its mechanism is revealed. It is found to be the result of circular fracture which develops within surrounding rock mass under the high geostress. The fractured shape of zonal disintegration was determined, and the radii of the fractured zones were found to fulfill the relationship of geometric progression. The numerical results were in accordance with the model test findings. The mechanism of the zonal disintegration was revealed by theoretical analysis based on fracture mechanics. The fractured zones are reportedly circular and concentric to the cavern. Each fracture zone ruptured at the elastic-plastic boundary of the surrounding rocks and then coalesced into the circular form. The geometric progression ratio was found to be related to the mechanical parameters and the ground stress of the surrounding rocks.

  6. Numerical simulation on zonal disintegration in deep surrounding rock mass.

    Science.gov (United States)

    Chen, Xuguang; Wang, Yuan; Mei, Yu; Zhang, Xin

    2014-01-01

    Zonal disintegration have been discovered in many underground tunnels with the increasing of embedded depth. The formation mechanism of such phenomenon is difficult to explain under the framework of traditional rock mechanics, and the fractured shape and forming conditions are unclear. The numerical simulation was carried out to research the generating condition and forming process of zonal disintegration. Via comparing the results with the geomechanical model test, the zonal disintegration phenomenon was confirmed and its mechanism is revealed. It is found to be the result of circular fracture which develops within surrounding rock mass under the high geostress. The fractured shape of zonal disintegration was determined, and the radii of the fractured zones were found to fulfill the relationship of geometric progression. The numerical results were in accordance with the model test findings. The mechanism of the zonal disintegration was revealed by theoretical analysis based on fracture mechanics. The fractured zones are reportedly circular and concentric to the cavern. Each fracture zone ruptured at the elastic-plastic boundary of the surrounding rocks and then coalesced into the circular form. The geometric progression ratio was found to be related to the mechanical parameters and the ground stress of the surrounding rocks.

  7. Enhanced sources of acoustic power surrounding AR 11429

    International Nuclear Information System (INIS)

    Donea, Alina; Hanson, Christopher

    2013-01-01

    Multi-frequency power maps of the local acoustic oscillations show acoustic enhancements (''acoustic-power halos'') at high frequencies surrounding large active region. Computational seismic holography reveals a high-frequency ''acoustic-emission halo'', or ''seismic glory'' surrounding large active regions. In this study, we have applied computational seismic holography to map the seismic seismic source density surrounding AR 11429. Studies of HMI/SDO Doppler data, shows that the ''acoustic halos'' and the ''seismic glories'' are prominent at high frequencies 5–8 mHz. We investigate morphological properties of acoustic-power and acoustic emission halos around an active region to see if they are spatially correlated. Details about the local magnetic field from vectormagnetograms of AR 11429 are included. We identify a 15'' region of seismic deficit power (dark moat) shielding the white-light boundary of the active region. The size of the seismic moat is related to region of intermediate magnetic field strength. The acoustic moat is circled by the halo of enhanced seismic amplitude as well as enhanced seismic emission. Overall, the results suggest that features are related. However, if we narrow the frequency band to 5.5 – 6.5 mHz, we find that the seismic source density dominates over the local acoustic power, suggesting the existence of sources that emit more energy downward into the solar interior than upward toward the solar surface.

  8. Tissue reaction surrounding miniscrews for orthodontic anchorage: An animal experiment

    Directory of Open Access Journals (Sweden)

    Stephanie Shih-Hsuan Chen

    2012-03-01

    Results and conclusions: (1 Tissue surrounding roots damaged by a miniscrew showed a significant inflammatory response. (2 Root resorption was occasionally observed after 3 weeks following insertion of a miniscrew even if the miniscrew was not in direct contact with the root. (3 Root repair was noted with a cementoblast lining along the resorption surface at as early as 3 weeks after miniscrew insertion. Alveolar bone filled in the lesion when the root damage was large so that the contour of the alveolar bone followed that of the damaged root, with the width of the periodontal ligament space being maintained. (4 Stable miniscrews were mainly those which did not contact adjacent roots, and for which the surrounding tissue showed only a small inflammatory response with some extent of direct bone contact around the miniscrew. On the contrary, most of the failed miniscrews were those which had direct contact with adjacent roots, and which exhibited severe tissue inflammation and were covered by thick layers of soft tissue. Failure was detected 3 weeks after insertion. Surprisingly, the epithelial lining surrounding the miniscrews might not have spontaneously resolved 6 weeks after screw removal. Persistent infection in the sinus tract was noted, and this would require attention.

  9. A permeability barrier surrounds taste buds in lingual epithelia

    Science.gov (United States)

    Dando, Robin; Pereira, Elizabeth; Kurian, Mani; Barro-Soria, Rene; Chaudhari, Nirupa

    2014-01-01

    Epithelial tissues are characterized by specialized cell-cell junctions, typically localized to the apical regions of cells. These junctions are formed by interacting membrane proteins and by cytoskeletal and extracellular matrix components. Within the lingual epithelium, tight junctions join the apical tips of the gustatory sensory cells in taste buds. These junctions constitute a selective barrier that limits penetration of chemosensory stimuli into taste buds (Michlig et al. J Comp Neurol 502: 1003–1011, 2007). We tested the ability of chemical compounds to permeate into sensory end organs in the lingual epithelium. Our findings reveal a robust barrier that surrounds the entire body of taste buds, not limited to the apical tight junctions. This barrier prevents penetration of many, but not all, compounds, whether they are applied topically, injected into the parenchyma of the tongue, or circulating in the blood supply, into taste buds. Enzymatic treatments indicate that this barrier likely includes glycosaminoglycans, as it was disrupted by chondroitinase but, less effectively, by proteases. The barrier surrounding taste buds could also be disrupted by brief treatment of lingual tissue samples with DMSO. Brief exposure of lingual slices to DMSO did not affect the ability of taste buds within the slice to respond to chemical stimulation. The existence of a highly impermeable barrier surrounding taste buds and methods to break through this barrier may be relevant to basic research and to clinical treatments of taste. PMID:25209263

  10. A permeability barrier surrounds taste buds in lingual epithelia.

    Science.gov (United States)

    Dando, Robin; Pereira, Elizabeth; Kurian, Mani; Barro-Soria, Rene; Chaudhari, Nirupa; Roper, Stephen D

    2015-01-01

    Epithelial tissues are characterized by specialized cell-cell junctions, typically localized to the apical regions of cells. These junctions are formed by interacting membrane proteins and by cytoskeletal and extracellular matrix components. Within the lingual epithelium, tight junctions join the apical tips of the gustatory sensory cells in taste buds. These junctions constitute a selective barrier that limits penetration of chemosensory stimuli into taste buds (Michlig et al. J Comp Neurol 502: 1003-1011, 2007). We tested the ability of chemical compounds to permeate into sensory end organs in the lingual epithelium. Our findings reveal a robust barrier that surrounds the entire body of taste buds, not limited to the apical tight junctions. This barrier prevents penetration of many, but not all, compounds, whether they are applied topically, injected into the parenchyma of the tongue, or circulating in the blood supply, into taste buds. Enzymatic treatments indicate that this barrier likely includes glycosaminoglycans, as it was disrupted by chondroitinase but, less effectively, by proteases. The barrier surrounding taste buds could also be disrupted by brief treatment of lingual tissue samples with DMSO. Brief exposure of lingual slices to DMSO did not affect the ability of taste buds within the slice to respond to chemical stimulation. The existence of a highly impermeable barrier surrounding taste buds and methods to break through this barrier may be relevant to basic research and to clinical treatments of taste. Copyright © 2015 the American Physiological Society.

  11. Towards understanding the puzzling lack of acid geothermal springs in Tibet (China): Insight from a comparison with Yellowstone (USA) and some active volcanic hydrothermal systems

    Science.gov (United States)

    Nordstrom, D. Kirk; Guo, Qinghai; McCleskey, R. Blaine

    2014-01-01

    Explanations for the lack of acid geothermal springs in Tibet are inferred from a comprehensive hydrochemical comparison of Tibetan geothermal waters with those discharged from Yellowstone (USA) and two active volcanic areas, Nevado del Ruiz (Colombia) and Miravalles (Costa Rica) where acid springs are widely distributed and diversified in terms of geochemical characteristic and origin. For the hydrothermal areas investigated in this study, there appears to be a relationship between the depths of magma chambers and the occurrence of acid, chloride-rich springs formed via direct magmatic fluid absorption. Nevado del Ruiz and Miravalles with magma at or very close to the surface (less than 1–2 km) exhibit very acidic waters containing HCl and H2SO4. In contrast, the Tibetan hydrothermal systems, represented by Yangbajain, usually have fairly deep-seated magma chambers so that the released acid fluids are much more likely to be fully neutralized during transport to the surface. The absence of steam-heated acid waters in Tibet, however, may be primarily due to the lack of a confining layer (like young impermeable lavas at Yellowstone) to separate geothermal steam from underlying neutral chloride waters and the possible scenario that the deep geothermal fluids below Tibet carry less H2S than those below Yellowstone.

  12. Towards understanding the puzzling lack of acid geothermal springs in Tibet (China): Insight from a comparison with Yellowstone (USA) and some active volcanic hydrothermal systems

    Science.gov (United States)

    Guo, Qinghai; Kirk Nordstrom, D.; Blaine McCleskey, R.

    2014-11-01

    Explanations for the lack of acid geothermal springs in Tibet are inferred from a comprehensive hydrochemical comparison of Tibetan geothermal waters with those discharged from Yellowstone (USA) and two active volcanic areas, Nevado del Ruiz (Colombia) and Miravalles (Costa Rica) where acid springs are widely distributed and diversified in terms of geochemical characteristic and origin. For the hydrothermal areas investigated in this study, there appears to be a relationship between the depths of magma chambers and the occurrence of acid, chloride-rich springs formed via direct magmatic fluid absorption. Nevado del Ruiz and Miravalles with magma at or very close to the surface (less than 1-2 km) exhibit very acidic waters containing HCl and H2SO4. In contrast, the Tibetan hydrothermal systems, represented by Yangbajain, usually have fairly deep-seated magma chambers so that the released acid fluids are much more likely to be fully neutralized during transport to the surface. The absence of steam-heated acid waters in Tibet, however, may be primarily due to the lack of a confining layer (like young impermeable lavas at Yellowstone) to separate geothermal steam from underlying neutral chloride waters and the possible scenario that the deep geothermal fluids below Tibet carry less H2S than those below Yellowstone.

  13. INTERACTIONS OF THE INFRARED BUBBLE N4 WITH ITS SURROUNDINGS

    Energy Technology Data Exchange (ETDEWEB)

    Liu, Hong-Li; Li, Jin-Zeng; Yuan, Jing-Hua; Huang, Maohai; Huang, Ya-Fang; Zhang, Si-Ju [National Astronomical Observatories, Chinese Academy of Sciences, 20A Datun Road, Chaoyang District, Beijing 100012 (China); Wu, Yuefang [Department of Astronomy, Peking University, 100871 Beijing (China); Liu, Tie [Korea Astronomy and Space Science Institute 776, Daedeokdae-ro, Yuseong-gu, Daejeon, 305-348 (Korea, Republic of); Dubner, G.; Paron, S.; Ortega, M. E. [1Instituto de Astronomía y Física del Espacio (IAFE, CONICET-UBA), CC 67, Suc. 28, 1428 Buenos Aires (Argentina); Molinari, Sergio [Istituto di Astrofisica e Planetologia Spaziali—IAPS, Istituto Nazionale di Astrofisica—INAF, via Fosso del Cavaliere 100, I-00133 Roma (Italy); Zavagno, Annie; Samal, Manash R., E-mail: hlliu@nao.cas.cn [Aix Marseille Universit, CNRS, LAM (Laboratoire d’Astrophysique de Marseille) UMR 7326, F-13388, Marseille (France)

    2016-02-10

    The physical mechanisms that induce the transformation of a certain mass of gas in new stars are far from being well understood. Infrared bubbles associated with H ii regions have been considered to be good samples for investigating triggered star formation. In this paper we report on the investigation of the dust properties of the infrared bubble N4 around the H ii region G11.898+0.747, analyzing its interaction with its surroundings and star formation histories therein, with the aim of determining the possibility of star formation triggered by the expansion of the bubble. Using Herschel PACS and SPIRE images with a wide wavelength coverage, we reveal the dust properties over the entire bubble. Meanwhile, we are able to identify six dust clumps surrounding the bubble, with a mean size of 0.50 pc, temperature of about 22 K, mean column density of 1.7 × 10{sup 22} cm{sup −2}, mean volume density of about 4.4 × 10{sup 4} cm{sup −3}, and a mean mass of 320 M{sub ⊙}. In addition, from PAH emission seen at 8 μm, free–free emission detected at 20 cm, and a probability density function in special regions, we could identify clear signatures of the influence of the H ii region on the surroundings. There are hints of star formation, though further investigation is required to demonstrate that N4 is the triggering source.

  14. Blooming Trees: Substructures and Surrounding Groups of Galaxy Clusters

    Science.gov (United States)

    Yu, Heng; Diaferio, Antonaldo; Serra, Ana Laura; Baldi, Marco

    2018-06-01

    We develop the Blooming Tree Algorithm, a new technique that uses spectroscopic redshift data alone to identify the substructures and the surrounding groups of galaxy clusters, along with their member galaxies. Based on the estimated binding energy of galaxy pairs, the algorithm builds a binary tree that hierarchically arranges all of the galaxies in the field of view. The algorithm searches for buds, corresponding to gravitational potential minima on the binary tree branches; for each bud, the algorithm combines the number of galaxies, their velocity dispersion, and their average pairwise distance into a parameter that discriminates between the buds that do not correspond to any substructure or group, and thus eventually die, and the buds that correspond to substructures and groups, and thus bloom into the identified structures. We test our new algorithm with a sample of 300 mock redshift surveys of clusters in different dynamical states; the clusters are extracted from a large cosmological N-body simulation of a ΛCDM model. We limit our analysis to substructures and surrounding groups identified in the simulation with mass larger than 1013 h ‑1 M ⊙. With mock redshift surveys with 200 galaxies within 6 h ‑1 Mpc from the cluster center, the technique recovers 80% of the real substructures and 60% of the surrounding groups; in 57% of the identified structures, at least 60% of the member galaxies of the substructures and groups belong to the same real structure. These results improve by roughly a factor of two the performance of the best substructure identification algorithm currently available, the σ plateau algorithm, and suggest that our Blooming Tree Algorithm can be an invaluable tool for detecting substructures of galaxy clusters and investigating their complex dynamics.

  15. Earthquakes in Switzerland and surrounding regions during 2007

    Energy Technology Data Exchange (ETDEWEB)

    Baer, M.; Deichmann, N.; Clinton, J.; Husen, S.; Faeh, D.; Giardini, D.; Kaestli, P.; Kradolfer, U.; Wiemer, S

    2008-12-15

    This report of the Swiss Seismological Service summarizes the seismic activity in Switzerland and surrounding regions during 2007. During this period, 531 earthquakes and 92 quarry blasts were detected and located in the region under consideration. Of these earthquakes, 30 are aftershocks of the stimulation of a proposed geothermal reservoir beneath the city of Basel in December of 2006. With 20 events with {mu}{sub {iota}} {>=} 2.5, four of which were artificially induced, the seismic activity in the year 2007 was far below the average over the previous 32 years. (author)

  16. Isoperimetric inequalities in surround system and space science

    OpenAIRE

    JiaJin Wen; Jun Yuan; ShanHe Wu

    2016-01-01

    Abstract By means of the algebraic, analysis, convex geometry, computer, and inequality theories we establish the following isoperimetric inequality in the centered 2-surround system S ( 2 ) { P , Γ , l } $S^{(2)} \\{P,\\varGamma ,l \\}$ : ( 1 | Γ | ∮ Γ r ¯ P p ) 1 / p ⩽ | Γ | 4 π sin l π | Γ | [ csc l π | Γ | + cot 2 l π | Γ | ln ( tan l π | Γ | + sec l π | Γ | ) ] , ∀ p ⩽ − 2 . $$\\begin{aligned}& \\biggl(\\frac{1}{|\\varGamma |} \\oint_{\\varGamma }\\bar{r}_{P}^{p} \\biggr)^{1/p}\\leqslant\\frac{|\\varG...

  17. Induced radioactivity in a 4 MW target and its surroundings

    CERN Document Server

    Agosteo, Stefano; Otto, Thomas; Silari, Marco

    2003-01-01

    An important aspect of a future CERN Neutrino Factory is the material activation arising from a 2.2 GeV, 4 MW proton beam striking a mercury target. An estimation of the hadronic inelastic interactions and the production of residual nuclei in the target, the magnetic horn, the decay tunnel, the surrounding rock and a downstream dump was performed by the Monte Carlo hadronic cascade code FLUKA. The aim was both to assess the dose equivalent rate to be expected during maintenance work and to evaluate the amount of residual radioactivity, which will have to be disposed of after the facility has ceased operation.

  18. Mutual seismic interaction between tunnels and the surrounding granular soil

    Directory of Open Access Journals (Sweden)

    Mohamed Ahmed Abdel-Motaal

    2014-12-01

    Study results show that the maximum exerted straining actions in tunnel lining are directly proportional to the relative stiffness between tunnel and surrounding soil (lining thickness and soil shear modulus. Moreover, it is highly affected by the peak ground acceleration and the tunnel location (embedment depth. A comprehensive study is performed to show the effect of tunnel thickness and tunnel diameter on both the induced bending moment and lining deformation. In general, it is concluded that seismic analysis should be considered in regions subjected to peak ground acceleration greater than 0.15g.

  19. Earthquakes in Switzerland and surrounding regions during 2007

    International Nuclear Information System (INIS)

    Baer, M.; Deichmann, N.; Clinton, J.; Husen, S.; Faeh, D.; Giardini, D.; Kaestli, P.; Kradolfer, U.; Wiemer, S.

    2008-01-01

    This report of the Swiss Seismological Service summarizes the seismic activity in Switzerland and surrounding regions during 2007. During this period, 531 earthquakes and 92 quarry blasts were detected and located in the region under consideration. Of these earthquakes, 30 are aftershocks of the stimulation of a proposed geothermal reservoir beneath the city of Basel in December of 2006. With 20 events with Μ ι ≥ 2.5, four of which were artificially induced, the seismic activity in the year 2007 was far below the average over the previous 32 years. (author)

  20. Diversity of vascular plants of Piestany and surroundings

    International Nuclear Information System (INIS)

    Penzesova, A.; Galusova, T.

    2013-01-01

    In the present work is a summary of the results of floristic research aimed at determining diversity of vascular plants of Piestany and its surroundings. Plant taxa we determined using the designation keys. We have compiled a list of plant species occurring in the monitored area, we evaluated the selected botanical-phytogeographical characteristics of flora, we've put together a list of local protected, endangered and rare species and a list of local invasive and expansive species according to sources. (Authors)

  1. One Japanese case on taxation surrounding foreign trust

    OpenAIRE

    SUZUKI, Yuya

    2015-01-01

    Taxation surrounding trust at cross-border situation is paid attention to byworldwide basis. Japan is not exception. According to recent Japanesejurisprudence, where a trust had been established in accordance with State law ofNew Jersey, the U.S., it was disputed whether or not the act settling that trust fellwithin “shintaku koui (an act of trust)” and one of the related members, who had beena minor child at that time, fell within “jyueki sha (beneficiary)” under JapaneseInheritance Tax Act....

  2. Cortical Surround Interactions and Perceptual Salience via Natural Scene Statistics.

    Directory of Open Access Journals (Sweden)

    Ruben Coen-Cagli

    Full Text Available Spatial context in images induces perceptual phenomena associated with salience and modulates the responses of neurons in primary visual cortex (V1. However, the computational and ecological principles underlying contextual effects are incompletely understood. We introduce a model of natural images that includes grouping and segmentation of neighboring features based on their joint statistics, and we interpret the firing rates of V1 neurons as performing optimal recognition in this model. We show that this leads to a substantial generalization of divisive normalization, a computation that is ubiquitous in many neural areas and systems. A main novelty in our model is that the influence of the context on a target stimulus is determined by their degree of statistical dependence. We optimized the parameters of the model on natural image patches, and then simulated neural and perceptual responses on stimuli used in classical experiments. The model reproduces some rich and complex response patterns observed in V1, such as the contrast dependence, orientation tuning and spatial asymmetry of surround suppression, while also allowing for surround facilitation under conditions of weak stimulation. It also mimics the perceptual salience produced by simple displays, and leads to readily testable predictions. Our results provide a principled account of orientation-based contextual modulation in early vision and its sensitivity to the homogeneity and spatial arrangement of inputs, and lends statistical support to the theory that V1 computes visual salience.

  3. Effectively Communicating the Uncertainties Surrounding Ebola Virus Transmission.

    Directory of Open Access Journals (Sweden)

    Andy Kilianski

    2015-10-01

    Full Text Available The current Ebola virus outbreak has highlighted the uncertainties surrounding many aspects of Ebola virus virology, including routes of transmission. The scientific community played a leading role during the outbreak-potentially, the largest of its kind-as many of the questions surrounding ebolaviruses have only been interrogated in the laboratory. Scientists provided an invaluable resource for clinicians, public health officials, policy makers, and the lay public in understanding the progress of Ebola virus disease and the continuing outbreak. Not all of the scientific communication, however, was accurate or effective. There were multiple instances of published articles during the height of the outbreak containing potentially misleading scientific language that spurred media overreaction and potentially jeopardized preparedness and policy decisions at critical points. Here, we use articles declaring the potential for airborne transmission of Ebola virus as a case study in the inaccurate reporting of basic science, and we provide recommendations for improving the communication about unknown aspects of disease during public health crises.

  4. Effectively Communicating the Uncertainties Surrounding Ebola Virus Transmission.

    Science.gov (United States)

    Kilianski, Andy; Evans, Nicholas G

    2015-10-01

    The current Ebola virus outbreak has highlighted the uncertainties surrounding many aspects of Ebola virus virology, including routes of transmission. The scientific community played a leading role during the outbreak-potentially, the largest of its kind-as many of the questions surrounding ebolaviruses have only been interrogated in the laboratory. Scientists provided an invaluable resource for clinicians, public health officials, policy makers, and the lay public in understanding the progress of Ebola virus disease and the continuing outbreak. Not all of the scientific communication, however, was accurate or effective. There were multiple instances of published articles during the height of the outbreak containing potentially misleading scientific language that spurred media overreaction and potentially jeopardized preparedness and policy decisions at critical points. Here, we use articles declaring the potential for airborne transmission of Ebola virus as a case study in the inaccurate reporting of basic science, and we provide recommendations for improving the communication about unknown aspects of disease during public health crises.

  5. REMOTE SENSING EFFICIENCY FOR URBAN ANALYSIS OF MECCA AND SURROUNDS

    Directory of Open Access Journals (Sweden)

    A. Imam

    2016-06-01

    Full Text Available Situated in the southwest of Saudi Arabia, Mecca is considered the spiritual capital of one and a half billion worldwide Muslims. The city is visited by millions of pilgrims every year. It has undergone significant changes in land cover (LC since the government first embarked on a series of ambitious development projects 20 years ago to accommodate the growing number of pilgrims and citizens. The main objective of our study is to detect, identify, analyze and measure the evolving land cover and urban morphology composition from multi-temporal satellite images. To characterize the morphological change during a period of twenty years, four satellite images, acquired in 1998 by Landsat TM and in 2003, 2008 and 2013 by Landsat ETM+, were classified into five main categories: Urban, Street, Soil and Vegetation. In addition, DEM has been extracted and included as Mountain. Change detection (CD analysis is applied using post-classification comparison and GIS. As part of the study, morphological index, such as, Entropy is included for better understanding of urban structures behaviour. Mecca and its surroundings show a noticeable increase in urban and vegetation cover. Urban cover (UC changes were divided into five radial directions: Northeast, Southeast, Southwest, East, and Northwest. These changes are influenced by mountain ranges surrounding the city and the highways. These revelations can play a significant role towards future planning and development activities, which may further promote urban growth.

  6. Remote Sensing Efficiency for Urban Analysis of Mecca and Surrounds

    Science.gov (United States)

    Imam, Ayman; Alhaddad, Bahaa; Roca, Josep

    2016-06-01

    Situated in the southwest of Saudi Arabia, Mecca is considered the spiritual capital of one and a half billion worldwide Muslims. The city is visited by millions of pilgrims every year. It has undergone significant changes in land cover (LC) since the government first embarked on a series of ambitious development projects 20 years ago to accommodate the growing number of pilgrims and citizens. The main objective of our study is to detect, identify, analyze and measure the evolving land cover and urban morphology composition from multi-temporal satellite images. To characterize the morphological change during a period of twenty years, four satellite images, acquired in 1998 by Landsat TM and in 2003, 2008 and 2013 by Landsat ETM+, were classified into five main categories: Urban, Street, Soil and Vegetation. In addition, DEM has been extracted and included as Mountain. Change detection (CD) analysis is applied using post-classification comparison and GIS. As part of the study, morphological index, such as, Entropy is included for better understanding of urban structures behaviour. Mecca and its surroundings show a noticeable increase in urban and vegetation cover. Urban cover (UC) changes were divided into five radial directions: Northeast, Southeast, Southwest, East, and Northwest. These changes are influenced by mountain ranges surrounding the city and the highways. These revelations can play a significant role towards future planning and development activities, which may further promote urban growth.

  7. MRI of normal pituitary glands and their surrounding structures

    International Nuclear Information System (INIS)

    Sato, Yoshiyuki

    1991-01-01

    Normal MRI appearances of the pituitary glands and their surrounding structures were evaluated in 332 patients without sellar and parasellar diseases. The height of the pituitary gland was maximum at 10-19 years of age reflecting hormonal activity. The width of the pituitary gland decreased, while that of the cavernous sinus increased with aging. This is probably due to atherosclerotic change of the internal carotid artery. Females younger than 30 years of age tended to show a convex upper surface of the pituitary gland and the displacement of the pituitary stalk was common after 50 years of age. Almost all of the anterior lobe of the pituitary gland showed isointensity relative to the pons or cerebral cortex and the majority (85.1%) of the posterior lobe showed hyperintensity. However, the anterior lobe in 2 newborns showed hyperintensity similar to the normal posterior lobe in adults. The posterior lobe was located off the midline in 19.1% of the subjects. One case of pars intermedia cyst was discovered among 14 subjects who were administered Gd-DTPA. The dural membrane between the pituitary gland and cavernous sinus was recognizable only in 8.6% on the right side and 7.5% on the left side. Primary empty sella was identified in 4.5%. Knowledge of the above normal ranges and variations of the pituitary gland and its surrounding structures is important in diagnosing sellar and parasellar lesions. (author) 52 refs

  8. TRIGGERED STAR FORMATION SURROUNDING WOLF-RAYET STAR HD 211853

    Energy Technology Data Exchange (ETDEWEB)

    Liu Tie; Wu Yuefang; Zhang Huawei [Department of Astronomy, Peking University, 100871 Beijing (China); Qin Shengli, E-mail: liutiepku@gmail.com [I. Physikalisches Institut, Universitaet zu Koeln, Zuelpicher Str. 77, 50937 Koeln (Germany)

    2012-05-20

    The environment surrounding Wolf-Rayet (W-R) star HD 211853 is studied in molecular, infrared, as well as radio, and H I emission. The molecular ring consists of well-separated cores, which have a volume density of 10{sup 3} cm{sup -3} and kinematic temperature {approx}20 K. Most of the cores are under gravitational collapse due to external pressure from the surrounding ionized gas. From the spectral energy distribution modeling toward the young stellar objects, the sequential star formation is revealed on a large scale in space spreading from the W-R star to the molecular ring. A small-scale sequential star formation is revealed toward core 'A', which harbors a very young star cluster. Triggered star formations are thus suggested. The presence of the photodissociation region, the fragmentation of the molecular ring, the collapse of the cores, and the large-scale sequential star formation indicate that the 'collect and collapse' process functions in this region. The star-forming activities in core 'A' seem to be affected by the 'radiation-driven implosion' process.

  9. TRIGGERED STAR FORMATION SURROUNDING WOLF-RAYET STAR HD 211853

    International Nuclear Information System (INIS)

    Liu Tie; Wu Yuefang; Zhang Huawei; Qin Shengli

    2012-01-01

    The environment surrounding Wolf-Rayet (W-R) star HD 211853 is studied in molecular, infrared, as well as radio, and H I emission. The molecular ring consists of well-separated cores, which have a volume density of 10 3 cm –3 and kinematic temperature ∼20 K. Most of the cores are under gravitational collapse due to external pressure from the surrounding ionized gas. From the spectral energy distribution modeling toward the young stellar objects, the sequential star formation is revealed on a large scale in space spreading from the W-R star to the molecular ring. A small-scale sequential star formation is revealed toward core 'A', which harbors a very young star cluster. Triggered star formations are thus suggested. The presence of the photodissociation region, the fragmentation of the molecular ring, the collapse of the cores, and the large-scale sequential star formation indicate that the 'collect and collapse' process functions in this region. The star-forming activities in core 'A' seem to be affected by the 'radiation-driven implosion' process.

  10. Cytoplasmic movement profiles of mouse surrounding nucleolus and not-surrounding nucleolus antral oocytes during meiotic resumption.

    Science.gov (United States)

    Bui, Thi Thu Hien; Belli, Martina; Fassina, Lorenzo; Vigone, Giulia; Merico, Valeria; Garagna, Silvia; Zuccotti, Maurizio

    2017-05-01

    Full-grown mouse antral oocytes are classified as surrounding nucleolus (SN) or not-surrounding nucleolus (NSN), depending on the respective presence or absence of a ring of Hoechst-positive chromatin surrounding the nucleolus. In culture, both types of oocytes resume meiosis and reach the metaphase II (MII) stage, but following insemination, NSN oocytes arrest at the two-cell stage whereas SN oocytes may develop to term. By coupling time-lapse bright-field microscopy with image analysis based on particle image velocimetry, we provide the first systematic measure of the changes to the cytoplasmic movement velocity (CMV) occurring during the germinal vesicle-to-MII (GV-to-MII) transition of these two types of oocytes. Compared to SN oocytes, NSN oocytes display a delayed GV-to-MII transition, which can be mostly explained by retarded germinal vesicle break down and first polar body extrusion. SN and NSN oocytes also exhibit significantly different CMV profiles at four main time-lapse intervals, although this difference was not predictive of SN or NSN oocyte origin because of the high variability in CMV. When CMV profile was analyzed through a trained artificial neural network, however, each single SN or NSN oocyte was blindly identified with a probability of 92.2% and 88.7%, respectively. Thus, the CMV profile recorded during meiotic resumption may be exploited as a cytological signature for the non-invasive assessment of the oocyte developmental potential, and could be informative for the analysis of the GV-to-MII transition of oocytes of other species. © 2017 Wiley Periodicals, Inc.

  11. The effects of low pH and elevated aluminum on yellowstone cutthroat trout (Oncorhynchus clarki bouvieri)

    International Nuclear Information System (INIS)

    Farag, A.M.; Woodward, D.F.; Little, E.E.; Steadman, B.; Vertucci, F.A.

    1993-01-01

    Although acid deposition is not considered a problem in the western US, surface waters in high elevations and fish inhabiting these waters may be vulnerable to acidification. This study examined the sensitivity of a wester salmonid to acid and aluminum stress. Yellowstone cutthroat trout (Oncorhynchus clarki bouvieri; YSC) were exposed for 7 d during each of four early life stages, or continuously from fertilization to 40 d post-hatch, to decreased pH and elevated Al. The authors monitored survival, growth, whole-body ion content, and behavior of the exposed fish. Sensitivity of early life stages of YSC may be expressed by survival or by survival and sublethal effects. In their study, eggs were the most sensitive life stage of YSC to low pH if survival alone was considered. However, the sublethal effects on growth, tissue ion content, and behavior revealed the alevins and swim-up larvae were more sensitive to reduced pH and increased Al than eggs or eyed embryos. They also observed that survival was significantly decreased if YSC were exposed to pH 6.0 and 50 μg Al per liter continuously from fertilization to 40 d post-hatch

  12. Whitebark pine, population density, and home-range size of grizzly bears in the greater yellowstone ecosystem.

    Science.gov (United States)

    Bjornlie, Daniel D; Van Manen, Frank T; Ebinger, Michael R; Haroldson, Mark A; Thompson, Daniel J; Costello, Cecily M

    2014-01-01

    Changes in life history traits of species can be an important indicator of potential factors influencing populations. For grizzly bears (Ursus arctos) in the Greater Yellowstone Ecosystem (GYE), recent decline of whitebark pine (WBP; Pinus albicaulis), an important fall food resource, has been paired with a slowing of population growth following two decades of robust population increase. These observations have raised questions whether resource decline or density-dependent processes may be associated with changes in population growth. Distinguishing these effects based on changes in demographic rates can be difficult. However, unlike the parallel demographic responses expected from both decreasing food availability and increasing population density, we hypothesized opposing behavioral responses of grizzly bears with regard to changes in home-range size. We used the dynamic changes in food resources and population density of grizzly bears as a natural experiment to examine hypotheses regarding these potentially competing influences on grizzly bear home-range size. We found that home-range size did not increase during the period of whitebark pine decline and was not related to proportion of whitebark pine in home ranges. However, female home-range size was negatively associated with an index of population density. Our data indicate that home-range size of grizzly bears in the GYE is not associated with availability of WBP, and, for female grizzly bears, increasing population density may constrain home-range size.

  13. Whitebark pine, population density, and home-range size of grizzly bears in the greater yellowstone ecosystem.

    Directory of Open Access Journals (Sweden)

    Daniel D Bjornlie

    Full Text Available Changes in life history traits of species can be an important indicator of potential factors influencing populations. For grizzly bears (Ursus arctos in the Greater Yellowstone Ecosystem (GYE, recent decline of whitebark pine (WBP; Pinus albicaulis, an important fall food resource, has been paired with a slowing of population growth following two decades of robust population increase. These observations have raised questions whether resource decline or density-dependent processes may be associated with changes in population growth. Distinguishing these effects based on changes in demographic rates can be difficult. However, unlike the parallel demographic responses expected from both decreasing food availability and increasing population density, we hypothesized opposing behavioral responses of grizzly bears with regard to changes in home-range size. We used the dynamic changes in food resources and population density of grizzly bears as a natural experiment to examine hypotheses regarding these potentially competing influences on grizzly bear home-range size. We found that home-range size did not increase during the period of whitebark pine decline and was not related to proportion of whitebark pine in home ranges. However, female home-range size was negatively associated with an index of population density. Our data indicate that home-range size of grizzly bears in the GYE is not associated with availability of WBP, and, for female grizzly bears, increasing population density may constrain home-range size.

  14. Whitebark pine, population density, and home-range size of grizzly bears in the greater Yellowstone ecosystem

    Science.gov (United States)

    Bjornlie, Daniel D.; van Manen, Frank T.; Ebinger, Michael R.; Haroldson, Mark A.; Thompson, Daniel J.; Costello, Cecily M.

    2014-01-01

    Changes in life history traits of species can be an important indicator of potential factors influencing populations. For grizzly bears (Ursus arctos) in the Greater Yellowstone Ecosystem (GYE), recent decline of whitebark pine (WBP; Pinus albicaulis), an important fall food resource, has been paired with a slowing of population growth following two decades of robust population increase. These observations have raised questions whether resource decline or density-dependent processes may be associated with changes in population growth. Distinguishing these effects based on changes in demographic rates can be difficult. However, unlike the parallel demographic responses expected from both decreasing food availability and increasing population density, we hypothesized opposing behavioral responses of grizzly bears with regard to changes in home-range size. We used the dynamic changes in food resources and population density of grizzly bears as a natural experiment to examine hypotheses regarding these potentially competing influences on grizzly bear home-range size. We found that home-range size did not increase during the period of whitebark pine decline and was not related to proportion of whitebark pine in home ranges. However, female home-range size was negatively associated with an index of population density. Our data indicate that home-range size of grizzly bears in the GYE is not associated with availability of WBP, and, for female grizzly bears, increasing population density may constrain home-range size.

  15. Speaking to Nations

    DEFF Research Database (Denmark)

    Jensen, Lars

    The ebbs and flows, which typically influences not just rhetoric around, but also how we think through nationalism, suggest nationalism is somehow at the core the same ‘thing’. But is it really? What connects the rise of nationalism in Australia in the 1880s, to the nationalism of the First...... surrounding refugees and migrancy in Australia and Europe (since this is what we are invited to do by the organisers), and how the deconstruction of such discourses might lead to more constructive ways of speaking through nation, might offer a way. I think Stan Grant’s book, Talking to My Country, through its...... combination of incisive criticism and insistence on constructive nation-(re)building offers an interesting launching pad. I am hoping to use Grant’s nation-rebuilding project to suggest ways that could open up similar spaces in equally exclusivist, the-nation-is-white-places in Europe. I am aware Grant has...

  16. MAPPING THE SURROUNDINGS AS A REQUIREMENT FOR AUTONOMOUS DRIVING

    Directory of Open Access Journals (Sweden)

    M. Steininger

    2016-11-01

    Full Text Available Motivated by the hype around driverless cars and the challenges of the sensor integration and data processing, this paper presents a model for using a XBox One Microsoft Kinect stereo camera as sensor for mapping the surroundings. Today, the recognition of the environment of the car is mostly done by a mix of sensors like LiDAR, RADAR and cameras. In the case of the outdoor delivery challenge Robotour 2016 with model cars in scale 1:5, it is our goal to solve the task with one camera only. To this end, a three-stage approach was developed. The test results show that our approach can detect and locate objects at a range of up to eight meters in order to incorporate them as barriers in the navigation process.

  17. Natural occurring radioactivity in Palmyra and its surrounding

    International Nuclear Information System (INIS)

    Al-Masri, M. S.; Shwekani, R.; Raja, G.; Hushari, M.; Al-Hent, R.; Issa, M.

    2006-06-01

    In this work, the natural radiation background has been carried out for Palmyra city and its surroundings area. The study has covered gamma radiation measurements, indoor radon gas concentration and radionuclides levels in environmental samples (soil, water, plat). The results showed that indoor radon gas concentrations and radiation exposure rates are within the background levels. Also, the results showed that there is no artificial radiation in the area and there is no correlation between the natural radiation levels and the reported cancer cases. Therefore, the reported cancer cases in this area may be due to non-radiation cases, which must be investigated, or they are within the natural levels in Syria unless accurate statistics proves the opposite. (author)

  18. Physical geography of the Nete basin and surroundings

    International Nuclear Information System (INIS)

    Beerten, K.

    2011-01-01

    The report briefly describes the main features of the physical geography of the Nete basin (Campine region, Belgium) and its immediate surroundings. First, an integrated overview of the topography, morphology and hydrography is given. This overview serves as the basis for the assessment of the morphological stability of the region and also explains the relationship between the topography and the hydrology. Furthermore, special attention is paid to soil science including a quantitative survey of some soil characteristics data. Another part of this report deals with erosion processes caused by water and wind action, and the (potential) impact on the morphology. Finally, the palaeogeographical evolution during the Quaternary is discussed. This evolution shows that the environment is stable over 10 000 years or more in the current and similar climatic conditions. Altering climatic conditions, notably glacial-interglacial periods, have impacted erosion with periods of strong erosion.

  19. Precision Security: Integrating Video Surveillance with Surrounding Environment Changes

    Directory of Open Access Journals (Sweden)

    Wenfeng Wang

    2018-01-01

    Full Text Available Video surveillance plays a vital role in maintaining the social security although, until now, large uncertainty still exists in danger understanding and recognition, which can be partly attributed to intractable environment changes in the backgrounds. This article presents a brain-inspired computing of attention value of surrounding environment changes (EC with a processes-based cognition model by introducing a ratio value λ of EC-implications within considered periods. Theoretical models for computation of warning level of EC-implications to the universal video recognition efficiency (quantified as time cost of implication-ratio variations from λk to λk+1, k=1,2,… are further established. Imbedding proposed models into the online algorithms is suggested as a future research priority towards precision security for critical applications and, furthermore, schemes for a practical implementation of such integration are also preliminarily discussed.

  20. A Study of the Flow Field Surrounding Interacting Line Fires

    Directory of Open Access Journals (Sweden)

    Trevor Maynard

    2016-01-01

    Full Text Available The interaction of converging fires often leads to significant changes in fire behavior, including increased flame length, angle, and intensity. In this paper, the fluid mechanics of two adjacent line fires are studied both theoretically and experimentally. A simple potential flow model is used to explain the tilting of interacting flames towards each other, which results from a momentum imbalance triggered by fire geometry. The model was validated by measuring the velocity field surrounding stationary alcohol pool fires. The flow field was seeded with high-contrast colored smoke, and the motion of smoke structures was analyzed using a cross-correlation optical flow technique. The measured velocities and flame angles are found to compare reasonably with the predicted values, and an analogy between merging fires and wind-blown flames is proposed.

  1. Teacher Leadership: Everyday Practices Surrounding Work- Related Stress

    Directory of Open Access Journals (Sweden)

    Chiweshe Nigel

    2015-06-01

    Full Text Available This interpretivist study contributes to our scholarly understanding of how everyday practices surrounding work-related stress in education affect teacher leadership and successful learning outcomes. Insights are drawn from our long-standing engagement in the field where we observed how teaching staff, students, and management interacted. These observations were supplemented by in-depth interviews with 20 teaching staff. Our findings reveal competing demands and practices across the individual intrapersonal environment and the work related environment. There were three key themes that emerged in answer to the core research question: 1 the role of relational practices in managing teacher burnout, 2 the role of surveillance practices in education and 3 the role of assimilating practices in education. Drawing insights from these practices, we develop a conceptual framework that will help us to see relations at work anew, and develop a deeper understanding of ‘sickies’, motivation, learning outcomes and teacher leadership opportunities in education

  2. Biogeography of azooxanthellate corals in the Caribbean and surrounding areas

    Science.gov (United States)

    Dawson, J.

    2002-04-01

    Biogeographic patterns for azooxanthellate corals are not as well known as those of zooxanthellate (primarily reef-building) corals. I analyzed occurrences of 129 species of azooxanthellate corals in 19 geopolitical regions in the Caribbean and surrounding areas. I performed an unweighted pair-group method with arithmetic averages (UPGMA) cluster analysis using Bray-Curtis' similarity measure on the complete data set and shallow- and deep-water subsets of the data. The results indicate two provinces, each with a widespread (tropical and subtropical distributions) component to its fauna. One province has a tropical and primarily insular component to it, while the other has a subtropical and primarily continental component. By contrast, zooxanthellate corals have a uniform faunal composition throughout the Caribbean. Moreover, zooxanthellate corals have half as many species in the Caribbean as the azooxanthellate corals even though their global diversities are equal. These differences in diversity and geographic distribution patterns should be considered when developing conservation strategies.

  3. The surrounding tissue modifies the placental stem villous vascular responses

    DEFF Research Database (Denmark)

    Brøgger, Torbjørn; Forman, Axel; Aalkjær, Christian

    2014-01-01

    is available. In-depth understanding of the mechanisms involved in control of placental vascular tone are needed to develop new tissue targets for therapeutic intervention. Method: From fresh born placentas segments of stem villous arteries were carefully dissected. The artery branches were divided....... The surrounding trophoblast was removed from one end and left intact in the other, and the segment was divided to give two ring preparations, with or without trophoblast. The preparations were mounted in wire myographs and responses to vasoactive agents were compared. Results: pD2values for PGF2α, Tx-analog U...... or endotheline-1. These differences partly disappeared in the presence of L-NAME. Conclusion: The perivascular tissue significantly reduces sensitivity and force development of stem villous arteries, partly due to release of NO This represents a new mechanism for control of human stem villous artery tone....

  4. Hurricane Sandy: Rapid Response Imagery of the Surrounding Regions

    Data.gov (United States)

    National Oceanic and Atmospheric Administration, Department of Commerce — The imagery posted on this site is of Hurricane Sandy. The aerial photography missions were conducted by the NOAA Remote Sensing Division. The images were acquired...

  5. Influence of surrounding environment on subcritical crack growth in marble

    Science.gov (United States)

    Nara, Yoshitaka; Kashiwaya, Koki; Nishida, Yuki; , Toshinori, Ii

    2017-06-01

    Understanding subcritical crack growth in rock is essential for determining appropriate measures to ensure the long-term integrity of rock masses surrounding structures and for construction from rock material. In this study, subcritical crack growth in marble was investigated experimentally, focusing on the influence of the surrounding environment on the relationship between the crack velocity and stress intensity factor. The crack velocity increased with increasing temperature and/or relative humidity. In all cases, the crack velocity increased with increasing stress intensity factor. However, for Carrara marble (CM) in air, we observed a region in which the crack velocity still increased with temperature, but the increase in the crack velocity with increasing stress intensity factor was not significant. This is similar to Region II of subcritical crack growth observed in glass in air. Region II in glass is controlled by mass transport to the crack tip. In the case of rock, the transport of water to the crack tip is important. In general, Region II is not observed for subcritical crack growth in rock materials, because rocks contain water. Because the porosity of CM is very low, the amount of water contained in the marble is also very small. Therefore, our results imply that we observed Region II in CM. Because the crack velocity increased in both water and air with increasing temperature and humidity, we concluded that dry conditions at low temperature are desirable for the long-term integrity of a carbonate rock mass. Additionally, mass transport to the crack tip is an important process for subcritical crack growth in rock with low porosity.

  6. Quasars Probing Quasars: the Circumgalactic Medium Surrounding z ~ 2 Quasars

    Science.gov (United States)

    Lau, Marie; Quasars Probing Quasars survey

    2018-01-01

    Understanding the circumgalactic medium--the gaseous halo surrounding a galaxy, is an integral part to understanding galaxy evolution. The z ~ 2-3 universe is interesting as this is when the star formation rate and AGN activity peak. My thesis concludes the decade-long Quasars Probing Quasars survey designed for studying massive galaxy formation and quasar feedback. I use background quasar sightlines that pass close to foreground quasars to study the circumgalactic medium of quasar-host galaxies in absorption. My sample of 149 quasar pairs involve spectra taken with 17 different optical and near IR instruments. I present results on the statistical and physical properties of the circumgalactic medium. The circumgalactic medium is enriched even beyond the virial radius. The alpha/Fe abundance ratio is enhanced, suggesting enrichment from core-collapse supernovae. The cool gas mass within the virial radius is enough to fuel star formation for another Gyr, and may account for 1/3 of the baryonic budget of the galaxy halo. The ionization state increases with projected distance from the quasar, which implies the quasar does not dominate the ionizing radiation flux. However, detection of fluorescent Lyman-alpha emission and NV absorption imply these transverse absorbers are partially illuminated by the quasar. In one peculiar case, the absorbing clump has density >100 cm^-3 and sub-parsec size. The average absorption in the circumgalactic medium exhibits large velocity widths, and is asymmetric about the systemic redshift of the galaxies. The widths are consistent with gravitational motions and Hubble flow, and outflows are not required to explain them. The asymmetry can be explained if the ionizing radiation from the quasar is anisotropic or intermittent and the gas is not in inflow. My results pose challenges for cosmological hydrodynamic simulations to produce a substantial cool gas reservoir surrounding quasars, that is also enriched and shows extreme kinematics.

  7. Ingenuity and National Security

    Science.gov (United States)

    2013-06-01

    is in an unfamiliar situation as the lone super power without a significant existential threat. China is a rising power and Russia aspires to...independence to the most powerful nation on the planet . The time period surrounding World War II demonstrated that governmental processes and

  8. How A Black Hole Lights Up Its Surroundings

    Science.gov (United States)

    Kohler, Susanna

    2017-10-01

    How do the supermassive black holes that live at the centers of galaxies influence their environments? New observations of a distant active galaxy offer clues about this interaction.Signs of CoevolutionPlot demonstrating the m-sigma relation, the empirical correlation between the stellar velocity dispersion of a galactic bulge and the mass of the supermassive black hole at its center. [Msigma]We know that the centers of active galaxies host supermassive black holes with masses of millions to billions of suns. One mystery surrounding these beasts is that they are observed to evolve simultaneously with their host galaxies for instance, an empirical relationship is seen between the growth of a black hole and the growth of its host galaxys bulge. This suggests that there must be a feedback mechanism through which the evolution of a black hole is linked to that of its host galaxy.One proposed source of this coupling is the powerful jets emitted from the poles of these supermassive black holes. These jets are thought to be produced as some of the material accreting onto the black hole is flung out, confined by surrounding gas and magnetic fields. Because the jets of hot gas and radiation extend outward through the host galaxy, they provide a means for the black hole to influence the gas and dust of its surroundings.In our current model of a radio-loud active galactic nuclei,a region of hot, ionized gas the narrow-line region lies beyond the sphere of influence of the supermassive black hole. [C.M. Urry and P. Padovani]Clues in the Narrow-Line RegionThe region of gas thought to sit just outside of the black holes sphere of influence (at a distance of perhaps a thousand to a few thousand light-years) is known as the narrow line region so named because we observe narrow emission lines from this gas. Given its hot, ionized state, this gas must somehow be being pummeled with energy. In the canonical picture, radiation from the black hole heats the gas directly in a process

  9. Logistics system as an effort of integration in the Mandalika Special Economic Zone and its surroundings

    Science.gov (United States)

    Suheri, T.; Melinda, W. G.

    2018-05-01

    Indonesia as one of the maritime countries with its territory in the form of an archipelago. A SEZ with an integrated logistics system is very influential on the value of tourism, especially marine tourism. The creation of logistics activities in the Mandalika SEZ as a tourism area formed from the export and import of products as a form of tourism promotion will attract many domestic and foreign tourists. The importance of a logistics system for both people and goods can support the investment system in the area of Mandalika SEZ. The logistics system can provide agglomeration of facilities and infrastructure to support tourist activities in terms of cargo distribution both locally and globally. Mandalika SEZ requires an integrated logistics system to improve national competitiveness in the form of physical system support such as commodity availability and city/regional infrastructure as well as non-physical infrastructure such as service providers and regulation in order to be sustainable. The logistics system also plays a strategic role in synchronizing and harmonizing progress between economies and between regions in order to avoid an imbalance with the surrounding area. This study aims to find out the performance of the existing logistics systems in the Mandalika area and its surrounding areas in order to formulate policies to integrate the logistics systems within and outside the region. This research used content analysis methodology.

  10. Natural Radioactivity in Abu-Tartor Phosphate Deposits and the Surrounding Region, New Valley, Egypt

    International Nuclear Information System (INIS)

    Khater, A.E.; Higgy, R.H.; Pimpl, M.

    1999-01-01

    Abu-Tartor phosphate mine. New Valley district, is one of the biggest phosphate mines in Egypt which will start full production soon. The planned ore rocks (24.8%P 2 O 5 ) annual production is 4 million tons. The aim of this study is to estimate the natural radioactivity levels in Abu-Tartor phosphate deposits and the surrounding region. The environmental radioactivity levels in the surrounding region will be considered as pre-operational levels which are essential to determine the radiological impacts of phosphate mining later on. Phosphate samples (ore rocks, wet rocks and beneficiation wastes) and environmental samples (soil, water and plant)were collected. The specific activities of Ra-226 (U-238) series, Th-232 series and K-40 were measured using gamma-ray spectrometry based on Hyper pure Germanium detectors. The specific activities of uranium isotopes (U-238, U-235 and U-234) were measured using alpha spectrometry based on surface barrier detectors after radiochemical separation. The specific activity of Pb-210 was measured using low background proportional gas counting system after radiochemical separation . The results were discussed and compared with national and international values

  11. Status of whitebarkpine in the Greater Yellowstone Ecosystem: A step-trend analysis comparing 2004-2007 to 2008-2011

    Science.gov (United States)

    Shanahan, Erin; Irvine, Kathryn M.; Roberts, Dave; Litt, Andrea R.; Legg, Kristin; Daley, Rob; Chambers, Nina

    2014-01-01

    Whitebark pine (Pinus albicaulis) is a foundation and keystone species in upper subalpine environments of the northern Rocky Mountains that strongly influences the biodiversity and productivity of high-elevation ecosystems (Tomback et al. 2001, Ellison et al. 2005). Throughout its historic range, whitebark pine has decreased significantly as a major component of high-elevation forests. As a result, it is critical to understand the challenges to whitebark pine—not only at the tree and stand level, but also as these factors influence the distribution of whitebark pine across the Greater Yellowstone Ecosystem (GYE). In 2003, the National Park Service (NPS) Greater Yellowstone Inventory & Monitoring Network identified whitebark pine as one of twelve significant natural resource indicators or vital signs to monitor (Jean et al. 2005, Fancy et al. 2009) and initiated a long-term, collaborative monitoring program. Partners in this effort include the U.S. Geological Survey, U.S. Forest Service, and Montana State University with representatives from each comprising the Greater Yellowstone Whitebark Pine Monitoring Working Group. The objectives of the monitoring program are to assess trends in (1) the proportion of live, whitebark pine trees (>1.4-m tall) infected with white pine blister rust (blister rust); (2) to document blister rust infection severity by the occurrence and location of persisting and new infections; (3) to determine mortality of whitebark pine trees and describe potential factors contributing to the death of trees; and (4) to assess the multiple components of the recruitment of understory whitebark pine into the reproductive population. In this report we summarize the past eight years (2004-2011) of whitebark pine status and trend monitoring in the GYE. Our study area encompasses six national forests (NF), two national parks (NP), as well as state and private lands in portions of Wyoming, Montana, and Idaho; this area is collectively described as the

  12. Tomography images of the Alpine roots and surrounding upper mantle

    Science.gov (United States)

    Plomerova, Jaroslava; Babuska, Vladislav

    2017-04-01

    Teleseismic body-wave tomography represents powerful tool to study regional velocity structure of the upper mantle and to image velocity anomalies, such as subducted lithosphere plates in collisional zones. In this contribution, we recapitulate 3D models of the upper mantle beneath the Alps, which developed at a collision zone of the Eurasian and African plates. Seismic tomography studies indicate a leading role of the rigid mantle lithosphere that functioned as a major stress guide during the plate collisions. Interactions of the European lithosphere with several micro-plates in the south resulted in an arcuate shape of this mountain range on the surface and in a complicated geometry of the Alpine subductions in the mantle. Early models with one bended lithosphere root have been replaced with more advanced models showing two separate lithosphere roots beneath the Western and Eastern Alps (Babuska et al., Tectonophysics 1990; Lippitsch et al., JGR 2003). The standard isotropic velocity tomography, based on pre-AlpArray data (the currently performed passive seismic experiment in the Alps and surroundings) images the south-eastward dipping curved slab of the Eurasian lithosphere in the Western Alps. On the contrary, beneath the Eastern Alps the results indicate a very steep northward dipping root that resulted from the collision of the European plate with the Adriatic microplate. Dando et al. (2011) interpret high-velocity heterogeneities at the bottom of their regional tomographic model as a graveyard of old subducted lithospheres. High density of stations, large amount of rays and dense ray-coverage of the volume studied are not the only essential pre-requisites for reliable tomography results. A compromise between the amount of pre-processed data and the high-quality of the tomography input (travel-time residuals) is of the high importance as well. For the first time, the existence of two separate roots beneath the Alps has been revealed from carefully pre

  13. Practice of Adult Education--Older Adults, Tourism, and Learning in Yellowstone

    Science.gov (United States)

    Roberson, Donald N., Jr.

    2005-01-01

    The purpose of this article is to present a program of learning for older adults in a national park. Because of the growing trend of tourism among retirees this learning during leisure is gaining prominence. The paper brings together the concepts of aging, self-directed learning, and tourism and leisure. In addition this paper presents a…

  14. The Roles of the Yellowstone Hotspot and Crustal Assimilation in Generating Pleistocene-Holocene Basalts on the Eastern Snake River Plain

    Science.gov (United States)

    Mintz, H.; Chadwick, J.

    2017-12-01

    The southwest motion of the North American plate across the Yellowstone hotspot created a chain of age-progressive rhyolitic calderas over the past 16 myr. in southern Idaho, U.S. The focus of Yellowstone activity now resides in northwest Wyoming, but basaltic volcanism has continued in its wake in southern Idaho on the eastern Snake River Plain (ESRP). These younger basaltic lavas are not age progressive and have buried the Yellowstone rhyolites on the ESRP. The ultimate source of the basalts is commonly ascribed to the passage or presence of the hotspot. However, the mechanisms involved, and the relative roles of the hotspot, other mantle sources, and the North American crust in generating the ESRP basalts remain unclear and have been the subject of recent geochemical and isotopic studies. In this study, the role of crustal assimilation is addressed by analyzing the chemical and isotopic characteristics of some of the youngest Pleistocene-Holocene tholeiitic volcanic fields on the ESRP, which were erupted through varying thicknesses of continental crust. Samples were analyzed from the Hell's Half Acre flow (5,200 years old; all dates Kuntz et al., 1986, 1994), Cerro Grande flow (13,380 years), and Black Butte Crater (a.k.a. Shoshone) flow (10,130 years), which were erupted at distances from between about 200 to 300 km from the current location of the hotspot. The crust of the ESRP thins from northeast to southwest, from about 47 km at the Hells Half Acre flow to 40 km at the Black Butte Crater flow, a thickness difference of about 15%. The apparently similar tectonic and magmatic environments of the three sampled flows suggest the crustal thickness variation may be a primary influence on the magnitude of assimilation and therefore the isotopic characteristics of the lavas. The goal of this work is to constrain the relative role of assimilation and to understand the source(s) of the magmas and the Yellowstone hotspot contribution. Major elements, trace elements

  15. A nebula of gases from Io surrounding Jupiter.

    Science.gov (United States)

    Krimigis, Stamatios M; Mitchell, Donald G; Hamilton, Douglas C; Dandouras, Jannis; Armstrong, Thomas P; Bolton, Scott J; Cheng, Andrew F; Gloeckler, George; Hsieh, K C; Keath, Edwin P; Krupp, Norbert; Lagg, Andreas; Lanzerotti, Louis J; Livi, Stefano; Mauk, Barry H; McEntire, Richard W; Roelof, Edmond C; Wilken, Berend; Williams, Donald J

    2002-02-28

    Several planetary missions have reported the presence of substantial numbers of energetic ions and electrons surrounding Jupiter; relativistic electrons are observable up to several astronomical units (au) from the planet. A population of energetic (>30[?]keV) neutral particles also has been reported, but the instrumentation was not able to determine the mass or charge state of the particles, which were subsequently labelled energetic neutral atoms. Although images showing the presence of the trace element sodium were obtained, the source and identity of the neutral atoms---and their overall significance relative to the loss of charged particles from Jupiter's magnetosphere---were unknown. Here we report the discovery by the Cassini spacecraft of a fast (>103[?]km[?]s-1) and hot magnetospheric neutral wind extending more than 0.5[?]au from Jupiter, and the presence of energetic neutral atoms (both hot and cold) that have been accelerated by the electric field in the solar wind. We suggest that these atoms originate in volcanic gases from Io, undergo significant evolution through various electromagnetic interactions, escape Jupiter's magnetosphere and then populate the environment around the planet. Thus a 'nebula' is created that extends outwards over hundreds of jovian radii.

  16. Earthquakes in Switzerland and surrounding regions during 2006

    Energy Technology Data Exchange (ETDEWEB)

    Baer, M.; Deichmann, N.; Braunmiller, J.; Clinton, J.; Husen, S.; Faeh, D.; Giardini, D.; Kaestli, P.; Kradolfer, U.; Wiemer, S

    2007-12-15

    This report of the Swiss Seismological Service summarizes the seismic activity in Switzerland and surrounding regions during 2006. During this period, 572 earthquakes and 91 quarry blasts were detected and located in the region under consideration. Of these earthquakes, two occurred in conjunction with the construction of the new Gotthard railway tunnel and 165 were induced artificially by the stimulation of a proposed geothermal reservoir beneath the city of Basel. With 20 events with {mu}{sub {iota}} {>=} 2.5, five of which were artificially induced, the seismic activity in the year 2006 was far below the average over the previous 31 years. Nevertheless, six events were felt by the public, most prominently the strongest of the induced Basel events ({mu}{sub {iota}} 3.4), which caused some non-structural building damage. Noteworthy are also the two earthquakes near Cortaillod ({mu}{sub {iota}} 3.2), on the shore of Lake Neuchatel, and in Val Mora ({mu}{sub {iota}} 3.5), between the Engadin and Val Muestair, as well as the 42 aftershocks of the {mu}{sub {iota}} 4.9 Vallorcine earthquake, between Martigny and Chamonix, of September 2005. (author)

  17. Mercury's interior, surface, and surrounding environment latest discoveries

    CERN Document Server

    Clark, Pamela Elizabeth

    2015-01-01<