Sample records for subalpine fir engelmann

  1. Similarity of nutrient uptake and root dimensions of Engelmann spruce and subalpine fir at two contrasting sites in Colorado

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    Yanai, R; McFarlane, K; Lucash, M; Kulpa, S; Wood, D


    Nutrient uptake capacity is an important parameter in modeling nutrient uptake by plants. Researchers commonly assume that uptake capacity measured for a species can be used across sites. We tested this assumption by measuring the nutrient uptake capacity of intact roots of Engelmann spruce (Picea engelmanni Parry) and subalpine fir (Abies lasiocarpa (Hook.) Nutt.) at Loch Vale Watershed and Fraser Experimental Forest in the Rocky Mountains of central Colorado. Roots still attached to the tree were exposed to one of three concentrations of nutrient solutions for time periods ranging from 1 to 96 hours, and solutions were analyzed for ammonium, nitrate, calcium, magnesium, and potassium. Surprisingly, the two species were indistinguishable in nutrient uptake within site for all nutrients (P > 0.25), but uptake rates differed by site. In general, nutrient uptake was higher at Fraser (P = 0.01, 0.15, 0.03, 0.18 for NH{sub 4}{sup +}, NO{sub 3}{sup -}, Ca{sup 2+}, and K{sup +}, respectively), which is west of the Continental Divide and has lower atmospheric deposition of N than Loch Vale. Mean uptake rates by site for ambient solution concentrations were 0.12 {micro}mol NH{sub 4}{sup +} g{sub fwt}{sup -1} h{sup -1}, 0.02 {micro}mol NO{sub 3}{sup -} g{sub fwt}{sup -1}, 0.21 {micro}mol Ca{sup 2+} g{sub fwt}{sup -1} h{sup -1}, and 0.01 {micro}mol Mg{sup 2+} g{sub fwt}{sup -1} h{sup -1} at Loch Vale, and 0.21 {micro}mol NH{sub 4}{sup +} f{sub fwt}{sup -1}h{sup -1}, 0.04 {micro}mol NO{sub 3}{sup -} g{sub fwt}{sup -1} h{sup -1}, 0.51 {micro}mol Ca{sup 2+}g{sub fwt}{sup -1}h{sup -1}, and 0.07 {micro}mol Mg{sup 2+} f{sub fwt}{sup -1}h{sup -1} at Fraser. The importance of site conditions in determining uptake capacity should not be overlooked when parameterizing nutrient uptake models. We also characterized the root morphology of these two species and compared them to other tree species we have measured at various sites in the northeastern USA. Engelman spruce and subalpine fir

  2. Provenance variability in nursery growth of subalpine fir (United States)

    Charlie Cartwright; Cheng Ying


    Subalpine fir (Abies lasiocarpa [Hook] Nutt.) is a wide-ranging, high-elevation species in the interior of British Columbia. It is commonly harvested for lumber, but replanting of it is limited. Some reticence is based upon wood quality and rate of growth, but there are also seed and nursery culturing difficulties. This study investigated seedling growth traits of 111...

  3. Multi-scale patterns of subalpine fir mortality are driven by complex interactions among broad-scale climate, local topography, stand structure, and tree characteristics (United States)

    Harvey, B. J.; Andrus, R. A.; Orrego, A.; Battaglia, M.; Negrón, J. F.; Veblen, T. T.


    Recent tree mortality events across much of western North America have been associated with warming temperatures and elevated drought stress, which can interact with forest stand-structure and tree vigor to drive outbreaks of native insect species. Although cross-scale interactions among drivers of tree-mortality events have been described for some beetle species (e.g., mountain pine beetle or spruce beetle) and their hosts (lodgepole pine and Engelmann spruce), less is known about how drivers at different scales interact to kill subalpine fir (Abies lasiocarpa) in a phenomenon simply dubbed "subalpine fir mortality" or "subalpine fir decline." Understanding the fate of subalpine fir is important, however, because this is commonly the tree species expected to establish and exhibit growth releases following outbreaks of mountain pine beetle or spruce beetle. In this study, we use three decades of field and remote data that span spatial scales from individual trees to sub-continental ecoregions to explore factors associated with subalpine fir mortality and how drivers interact across scales. Between 1991 and 2015, >5 million hectares (ha) of subalpine forest in the US Rocky Mountains have been affected by subalpine fir mortality. At the eco-region scale (1,000s of km), mortality was temporally associated with increases in regional drought stress, suggesting that climate is an important broad-scale driver. However, at the eco-region, landscape, and stand scales (several km to sub ha), mortality was greatest in cooler/wetter topographic locations and in areas with greater pre-mortality subalpine fir dominance. Conversely, mortality was lowest for fir trees in warmer/drier topographic locations and in areas of lesser pre-mortality subalpine fir dominance. This suggests that topographically driven differences in stand structure drive mortality dynamics at meso-scales and moderate the broad-scale influence of climate. Finally, at the tree and tree-neighborhood scale

  4. Long-term landscape changes in a subalpine spruce-fir forest in central Utah, USA

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    Jesse L. Morris1


    Full Text Available Background: In Western North America, increasing wildfire and outbreaks of native bark beetles have been mediated by warming climate conditions. Bioclimatic models forecast the loss of key high elevation species throughout the region. This study uses retrospective vegetation and fire history data to reconstruct the drivers of past disturbance and environmental change. Understanding the relationship among climate, antecedent disturbances, and the legacy effects of settlement-era logging can help identify the patterns and processes that create landscapes susceptible to bark beetle epidemics. Methods: Our analysis uses data from lake sediment cores, stand inventories, and historical records. Sediment cores were dated with radiometric techniques (14C and 210Pb/137Cs and subsampled for pollen and charcoal to maximize the temporal resolution during the historical period (1800 CE to present and to provide environmental baseline data (last 10,500 years. Pollen data for spruce were calibrated to carbon biomass (C t/ha using standard allometric equations and a transfer function. Charcoal samples were analyzed with statistical models to facilitate peak detection and determine fire recurrence intervals. Results: The Wasatch Plateau has been dominated by Engelmann spruce forests for the last ~10,500 years, with subalpine fir becoming more prominent since 6000 years ago. This landscape has experienced a dynamic fire regime, where burning events are more frequent and of higher magnitude during the last 3000 years. Two important disturbances have impacted Engelmann spruce in the historical period: 1 high-grade logging during the late 19th century; and (2 a high severity spruce beetle outbreak in the late 20th century that killed >90 % of mature spruce (>10 cm dbh. Conclusions: Our study shows that spruce-dominated forests in this region are resilient to a range of climate and disturbance regimes. Several lines of evidence suggest that 19th century logging

  5. Airflow patterns in a small subalpine basin (United States)

    G. Wooldridge; R. Musselman; B. Connell; D. Fox


    A study of mean wind speeds and directions has been completed in the Snowy Range of Southern Wyoming, U.S.A. It was conducted in a subalpine ecosystem at an altitude of 3 200 m to 3 400 m above sea level during the summers of 1988 and 1989. Indexes of deformation and axes of asymmetry due to wind shaping of Engelmann spruce (Picea engelmannii) and subalpine fir (Abies...

  6. Development of spruce-fir stands following spruce beetle outbreaks (United States)

    J. M. Schmid; T. E. Hinds


    Logged and unlogged stands of Engelmann spruce-subalpine fir were evaluated in spruce beetle outbreak areas infested about 15, 25, 50, and 100 years ago. Seedling regeneration was generally adequate except in heavily logged areas, although seedlings were often damaged, apparently by animals. Species composition was dramatically altered in favor of fir in the unlogged...

  7. Survival, frost susceptibility, growth, and disease resistance of corkbark and subalpine fir grown for landscape and Christmas trees (United States)

    Trees from six corkbark fir (Abies lasiocarpa var. arizonica) and 10 subalpine fir (A. lasiocarpa var. lasiocarpa) seed sources were grown at the University of Idaho Sandpoint Research and Extension Center (SREC) and two commercial nurseries in Idaho and Oregon. Post transplant mortality was highest...

  8. Mean wind patterns and snow depths in an alpine-subalpine ecosystem as measured by damage to coniferous trees (United States)

    G. L. Wooldridge; R. C. Musselman; R. A. Sommerfeld; D. G. Fox; B. H. Connell


    1. Deformations of Engelmann spruce and subalpine fir trees were surveyed for the purpose of determining climatic wind speeds and directions and snow depths in the Glacier Lakes Ecosystem Experiments Site (GLEES) in the Snowy Range of southeastern Wyoming, USA. Tree deformations were recorded at 50- and 100-m grid intervals over areas of c. 30 ha and 300 ha,...

  9. Disturbance and Stand Development of a Colorado Subalpine Forest


    Veblen, Thomas T.; Hadley, Keith S; Reid, Marion S


    Stand development patterns were examined in an Engelmann spruce (Picea engelmannii), subalpine fir (Abies lasiocarpa) and lodgepole pine (Pinus contorta var. latifolia) forest in Rocky Mountain National Park in northern Colorado. Two old-growth stands (with fine-scale windthrows dominating dynamics) and a 260-yr-old post-fire stand were sampled for tree ages, sizes, growth, and replacement patterns in windthrow gaps. Visual assessment of frequency of growth releases in increment cores, and de...

  10. Climate Risk Modelling of Balsam Woolly Adelgid Damage Severity in Subalpine Fir Stands of Western North America. (United States)

    Hrinkevich, Kathryn H; Progar, Robert A; Shaw, David C


    The balsam woolly adelgid (Adelges piceae (Ratzeburg) (Homoptera: Adelgidae)) (BWA) is a nonnative, invasive insect that threatens Abies species throughout North America. It is well established in the Pacific Northwest, but continues to move eastward through Idaho and into Montana and potentially threatens subalpine fir to the south in the central and southern Rocky Mountains. We developed a climatic risk model and map that predicts BWA impacts to subalpine fir using a two-step process. Using 30-year monthly climate normals from sites with quantitatively derived BWA damage severity index values, we built a regression model that significantly explained insect damage. The sites were grouped into two distinct damage categories (high damage and mortality versus little or no mortality and low damage) and the model estimates for each group were used to designate distinct value ranges for four climatic risk categories: minimal, low, moderate, and high. We then calculated model estimates for each cell of a 4-kilometer resolution climate raster and mapped the risk categories over the entire range of subalpine fir in the western United States. The spatial variation of risk classes indicates a gradient of climatic susceptibility generally decreasing from the Olympic Peninsula in Washington and the Cascade Range in Oregon and Washington moving eastward, with the exception of some high risk areas in northern Idaho and western Montana. There is also a pattern of decreasing climatic susceptibility from north to south in the Rocky Mountains. Our study provides an initial step for modeling the relationship between climate and BWA damage severity across the range of subalpine fir. We showed that September minimum temperature and a metric calculated as the maximum May temperature divided by total May precipitation were the best climatic predictors of BWA severity. Although winter cold temperatures and summer heat have been shown to influence BWA impacts in other locations, these

  11. Climate Risk Modelling of Balsam Woolly Adelgid Damage Severity in Subalpine Fir Stands of Western North America.

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    Kathryn H Hrinkevich

    Full Text Available The balsam woolly adelgid (Adelges piceae (Ratzeburg (Homoptera: Adelgidae (BWA is a nonnative, invasive insect that threatens Abies species throughout North America. It is well established in the Pacific Northwest, but continues to move eastward through Idaho and into Montana and potentially threatens subalpine fir to the south in the central and southern Rocky Mountains. We developed a climatic risk model and map that predicts BWA impacts to subalpine fir using a two-step process. Using 30-year monthly climate normals from sites with quantitatively derived BWA damage severity index values, we built a regression model that significantly explained insect damage. The sites were grouped into two distinct damage categories (high damage and mortality versus little or no mortality and low damage and the model estimates for each group were used to designate distinct value ranges for four climatic risk categories: minimal, low, moderate, and high. We then calculated model estimates for each cell of a 4-kilometer resolution climate raster and mapped the risk categories over the entire range of subalpine fir in the western United States. The spatial variation of risk classes indicates a gradient of climatic susceptibility generally decreasing from the Olympic Peninsula in Washington and the Cascade Range in Oregon and Washington moving eastward, with the exception of some high risk areas in northern Idaho and western Montana. There is also a pattern of decreasing climatic susceptibility from north to south in the Rocky Mountains. Our study provides an initial step for modeling the relationship between climate and BWA damage severity across the range of subalpine fir. We showed that September minimum temperature and a metric calculated as the maximum May temperature divided by total May precipitation were the best climatic predictors of BWA severity. Although winter cold temperatures and summer heat have been shown to influence BWA impacts in other

  12. Estimating under-canopy ablation in a subalpine red-fir forest, southern Sierra Nevada, California (United States)

    Kirchner, P. B.; Bales, R. C.; Rice, R.; Musselman, K. N.; Molotch, N. P.


    Snow ablation in forested environments is a result of the multi-component energy balance between the snow surface, radiation, topography, and vegetation. While these processes have been successfully described and modeled over small to moderate spatial extents the required data are available from few locations and existing models are computationally intensive. The problem of applying these principals to determining snow coverage for large spatial extents and frequent time steps, required by satellite observations, has not been solved. We present a simplified approach for determining a melt-out date based on modeled incident radiation, percent canopy cover, and leaf area index. This method was tested using results from instrumental data, field observations, and readily available spatial data sets by calibrating the MODIS Snow Covered Area and Grain size/albedo (MODSCAG) model from a snow-dominated site in the Wolverton basin Sequoia National Park; part of the Southern Sierra Nevada Critical Zone Observatory. The percent snow cover determined by MODSCAG from peak accumulation and melt out during the 2008 and 2009 water years were compared to ground observations of both forest gaps and under canopies. Ground based measurements indicated that under-canopy melt out of snow-covered area began earlier and ended 1 to 4 weeks after that indicated by satellite observations, which can only view snow in forest gaps. In our study ablation rates, snow cover duration, leaf area index, canopy closure, and Incoming short and long wave radiation were measured on north and southeast facing plots in a subalpine red fir forest. Results from regression analysis yield an R2=0.99 between modeled and measured short wave radiation and an R2=0.82 between leaf area index and the difference between open and under canopy thermal infrared radiation. Canopy cover and leaf area index were also found to be good predictors of observed melt rates and the melt off date of snow under tree canopies. This

  13. Spectral evidence of early-stage spruce beetle infestation in Engelmann spruce (United States)

    Adrianna C. Foster; Jonathan A. Walter; Herman H. Shugart; Jason Sibold; Jose Negron


    Spruce beetle (Dendroctonus rufipennis (Kirby)) outbreaks cause widespread mortality of Engelmann spruce (Picea engelmannii (Parry ex Engelm)) within the subalpine forests of the western United States. Early detection of infestations could allow forest managers to mitigate outbreaks or anticipate a response to tree mortality and the potential effects on ecosystem...

  14. Camurati-Engelmann disease

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    Neil Bellew


    Full Text Available Camurati-Engelmann disease (CED, or progressive diaphyseal dysplasia, is a rare sclerosing dysplasia of which 250 cases have been described in the English literature. The disease affects one in a million people and is autosomal dominant with variable penetrance. It was initially described by Cockayne in 1920; Camurati was the first to suggest its hereditary nature in 1922. A single case of muscular wasting and marked bone involvement was reported by Engelmann in 1929. As the name suggests, there is progressive hyperostosis and predominant involvement of the diaphyses. The onset of the disease is usually during childhood; patients usually present by puberty and usually before age 30, with limb pain, muscular weakness, waddling gait and easy fatigue. Other symptoms and signs may include delayed growth, reduced muscle mass, anorexia and enlargement of the arms and legs. Systemic manifestations of hepatosplenomegaly, bone marrow dysfunction (anaemia and leucopaenia and delayed sexual development occasionally occur. In a few patients, abnormal values of bone resorption and formation have been described. Radiologically, the hallmark of the disorder is bilateral, symmetrical cortical thickening of the diaphyses of the long bones on both the periosteal and endosteal sides of the diaphyses. In decreasing order of frequency, the tibia, femur, fibula, humerus, ulna and radius are affected. CED results from disturbance of intramembranous ossification (Fig.1 affecting the long bones, calvaria, mandible and facial bones. There are a few reported cases of involvement of the skull base (a site of endochondral ossification, but these occur in advanced stages.

  15. Comparisons of photosynthesis-related traits of 27 abundant or subordinate bryophyte species in a subalpine old-growth fir forest. (United States)

    Wang, Zhe; Bader, Maaike Y; Liu, Xin; Zhu, Zhangming; Bao, Weikai


    Bryophyte communities can exhibit similar structural and taxonomic diversity as vascular plant communities, just at a smaller scale. Whether the physiological diversity can be similarly diverse, and whether it can explain local abundance patterns is unknown, due to a lack of community-wide studies of physiological traits. This study re-analyzed data on photosynthesis-related traits (including the nitrogen, phosphorus and chlorophyll concentrations, photosynthetic capacities, and photosynthetic nutrient use efficiencies) of 27 bryophyte species in a subalpine old-growth fir forest on the eastern Tibetan Plateau. We explored differences between taxonomic groups and hypothesized that the most abundant bryophyte species had physiological advantages relative to other subdominant species. Principal component analysis (PCA) was used to summarize the differences among species and trait values of the most abundant and other co-occurring subdominant species. Species from the Polytrichaceae were separated out on both PCA axes, indicating their high chlorophyll concentrations and photosynthetic capacities (axis 1) and relatively high-light requirements (axis 2). Mniaceae species also had relatively high photosynthetic capacities, but their light saturation points were low. In contrast, Racomitrium joseph-hookeri and Lepidozia reptans , two species with a high shoot mass per area, had high-light requirements and low nutrient and chlorophyll concentrations and photosynthetic capacities. The nutrient concentrations, photosynthetic capacities, and photosynthetic nutrient use efficiencies of the most abundant bryophyte species did not differ from co-occurring subdominant species. Our research confirms the links between the photosynthesis-related traits and adaptation strategies of bryophytes. However, species relative abundance was not related to these traits.

  16. Identifying Subalpine Fir (Abies lasiocarpa Attacked by the Balsam Woolly Adelgid (Adelges piceae Using Spectral Measurements of the Foliage

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    Stephen P. Cook


    Full Text Available Balsam woolly adelgid is an invasive pest of firs in the United States. Aerial surveys are conducted for detection of adelgid infestations but other remotely sensed data may also be useful. Our objective was to determine if high spectral resolution, branch-level data can be used to distinguish infested from noninfested trees. Stepwise discriminant analysis yielded a three-variable model (the red-green index and two narrow-bands (one at 670 nm and the other at 1912 nm that classified infested versus non-infested trees with 94% accuracy compared with the 83% accuracy obtained with a single-variable model. The response of trees in narrow spectral bands was integrated across wavebands to simulate measurements from the multispectral SPOT5-HRVIR sensor. Stepwise discriminant analysis again yielded a three-variable model (simple ratio, the SPOT5-HRVIR band in the SWIR region and NDVI with similar accuracy (93% at discriminating infested from non-infested trees compared with the 83% accuracy obtained with a single-variable model.

  17. A Bayesian Analysis Shows That a Spruce Beetle Outbreak Reduces Snow Interception and Sublimation in a Subalpine Spruce-Fir Forest (United States)

    Frank, J. M.; Massman, W. J.; Ewers, B. E.; Williams, D. G.


    Sublimation is a key component in the water cycle of cold, snow dominated ecosystems. In many high elevation spruce-fir forests of western North America, recent spruce beetle outbreaks have caused widespread tree mortality, opened the canopy, and potentially altered the processes that control sublimation. This study evaluates three hypotheses: in these ecosystems the dominant source for sublimation originates from canopy intercepted snow, the loss of canopy following a beetle disturbance leads to significantly less sublimation, and major sublimation events are driven by the flow of sensible heat into the canopy. Bayesian analysis is used to evaluate a two source energy and canopy mass model that explains seventeen years (2000-2016) of winter eddy-covariance flux data at the GLEES AmeriFlux sites where a spruce beetle outbreak caused 75-85% basal area mortality. The model estimated that the resistance to snow sublimation from the canopy was an order of magnitude less than from the snowpack and that the maximum snow loading in the canopy was reduced to 25-56% of its pre-outbreak capacity. Comparing model results obtained using the observed decrease in leaf area index versus a "no beetle" condition, there has been a significant decrease in ecosystem sublimation since 2011. In the past few years, a 5-11% increase in snowpack sublimation has been offset by 28-32% less sublimation from canopy intercepted snow, with the net being 17-25% less total sublimation. This is equivalent to 3-6% of the total precipitation. Informing the model with information other than the above-canopy fluxes indicates that a near snowpack eddy covariance system decreases the canopy contribution to sublimation, including observed sensible heat fluxes requires a correction to resolve the surface energy imbalance, and stable isotopes of water vapor extend sublimation events. Because tree growth and ecological succession are slow in spruce-fir forests, these results could persist for decades.

  18. Warming and provenance limit tree recruitment across and beyond the elevation range of subalpine forest. (United States)

    Kueppers, Lara M; Conlisk, Erin; Castanha, Cristina; Moyes, Andrew B; Germino, Matthew J; de Valpine, Perry; Torn, Margaret S; Mitton, Jeffry B


    Climate niche models project that subalpine forest ranges will extend upslope with climate warming. These projections assume that the climate suitable for adult trees will be adequate for forest regeneration, ignoring climate requirements for seedling recruitment, a potential demographic bottleneck. Moreover, local genetic adaptation is expected to facilitate range expansion, with tree populations at the upper forest edge providing the seed best adapted to the alpine. Here, we test these expectations using a novel combination of common gardens, seeded with two widely distributed subalpine conifers, and climate manipulations replicated at three elevations. Infrared heaters raised temperatures in heated plots, but raised temperatures more in the forest than at or above treeline because strong winds at high elevation reduced heating efficiency. Watering increased season-average soil moisture similarly across sites. Contrary to expectations, warming reduced Engelmann spruce recruitment at and above treeline, as well as in the forest. Warming reduced limber pine first-year recruitment in the forest, but had no net effect on fourth-year recruitment at any site. Watering during the snow-free season alleviated some negative effects of warming, indicating that warming exacerbated water limitations. Contrary to expectations of local adaptation, low-elevation seeds of both species initially recruited more strongly than high-elevation seeds across the elevation gradient, although the low-provenance advantage diminished by the fourth year for Engelmann spruce, likely due to small sample sizes. High- and low-elevation provenances responded similarly to warming across sites for Engelmann spruce, but differently for limber pine. In the context of increasing tree mortality, lower recruitment at all elevations with warming, combined with lower quality, high-provenance seed being most available for colonizing the alpine, portends range contraction for Engelmann spruce. The lower

  19. Warming and provenance limit tree recruitment across and beyond the elevation range of subalpine forest (United States)

    Kueppers, Lara M.; Conlisk, Erin; Castanha, Cristina; Moyes, Andrew B.; Germino, Matthew; de Valpine, Perry; Torn, Margaret S.; Mitton, Jeffry B.


    Climate niche models project that subalpine forest ranges will extend upslope with climate warming. These projections assume that the climate suitable for adult trees will be adequate for forest regeneration, ignoring climate requirements for seedling recruitment, a potential demographic bottleneck. Moreover, local genetic adaptation is expected to facilitate range expansion, with tree populations at the upper forest edge providing the seed best adapted to the alpine. Here, we test these expectations using a novel combination of common gardens, seeded with two widely distributed subalpine conifers, and climate manipulations replicated at three elevations. Infrared heaters raised temperatures in heated plots, but raised temperatures more in the forest than at or above treeline because strong winds at high elevation reduced heating efficiency. Watering increased season-average soil moisture similarly across sites. Contrary to expectations, warming reduced Engelmann spruce recruitment at and above treeline, as well as in the forest. Warming reduced limber pine first-year recruitment in the forest, but had no net effect on fourth-year recruitment at any site. Watering during the snow-free season alleviated some negative effects of warming, indicating that warming exacerbated water limitations. Contrary to expectations of local adaptation, low-elevation seeds of both species initially recruited more strongly than high-elevation seeds across the elevation gradient, although the low-provenance advantage diminished by the fourth year for Engelmann spruce, likely due to small sample sizes. High- and low-elevation provenances responded similarly to warming across sites for Engelmann spruce, but differently for limber pine. In the context of increasing tree mortality, lower recruitment at all elevations with warming, combined with lower quality, high-provenance seed being most available for colonizing the alpine, portends range contraction for Engelmann spruce. The lower

  20. Camurati-Engelmann's disease: a case report | Kusaba Byanyima ...

    African Journals Online (AJOL)

    Camurati-Engelmann's disease: a case report. Rosemary Kusaba Byanyima, Jennifer Batuuka Nabawesi. Abstract. Camurati-Engelmann's disease is a rare condition worldwide. No cases have been documented in Uganda. A 26 year old female presented with a history of grinding pain in the limbs for over 20 years. Strong ...

  1. Testing remote sensing estimates of bark beetle induced mortality in lodgepole pine and Engelmann spruce with ground data (United States)

    King, A.; Ewers, B. E.; Sivanpillai, R.; Pendall, E.


    Bark beetles have caused widespread regional mortality in both lodgepole and Engelmann spruce forests across western North America, and while studies have addressed the impact on water partitioning caused by the mountain pine beetle, spruce beetle which often occur at high elevations with larger snowpack might have a disproportional impact. Beetle caused mortality can have significant effects on the hydrology of a watershed and therefore needs to be considered when evaluating increased runoff. The objective of this project was to generate maps showing beetle caused mortality for lodgepole pine and spruce fir forests that capture changes to the landscape to improve hydrologic models. Our study area in southeast Wyoming covered an area of approximately 2 by 4 km from 2700 to 2800m elevation range. High spatial resolution (0.5m) aerial imagery acquired by the Airborne Environmental Research Observational Camera (AEROCam) in fall 2011, provided by the Upper Midwest Aerospace Consortium (UMAC), was manually classified into four conifer thematic classes: live and dead lodgepole pine, and live and dead spruce/fir. The classified high resolution image was then verified by tree surveys conducted July-September, 2012 documenting species, tree diameter at breast height (dbh), and the stage of beetle infestation for each tree. After verification the high resolution aerial images were used to train and evaluate the accuracy of a supervised classification of a Landsat 5 Thematic Mapper image from the same time period and area. The preliminary results of a supervised classification show that map accuracy was 57%, 77%, 44%, and 83% for lodgepole live and dead, and spruce/fir live and dead respectively. The highest commission error, 24%, was for dead lodgepole pine being falsely labeled dead spruce/fir. The second highest commission error, 22%, was for live spruce/fir falsely labeled dead spruce/fir. The results indicate high spectral overlap between dead spruce/fir and dead

  2. Indicators and associated decay of Engelmann spruce in Colorado (United States)

    Thomas E. Hinds; Frank G. Hawksworth


    Average cull deductions for 11 cull indicators were determined from over 2,000 abnormalities on 1,027 merchantable Engelmann spruce in 21 stands throughout Colorado. On a board-foot basis, Fomes pini punk knots or sporophores caused an 81 percent deduction. Deduction for broken tops or dead tops with adjacent dead rust brooms amounted to 24 percent....

  3. Climate, geography, and tree establishment in subalpine meadows of the Olympic Mountains, Washington, USA (United States)

    Woodward, Andrea; Schreiner, Edward G.; Silsbee, D.G.


    Noticeable changes in vegetation distribution have occurred in the Pacific Northwest during the last century as trees have established in some subalpine meadows. To study the relationship of this process to climate, recently established trees were aged in six subalpine meadows in the Olympic Mountains, Washington. The sites represent three points along a steep precipitation gradient. Subalpine fir (Abies lasiocarpa) has been establishing at the dry end of the gradient, mountain hemlock (Tsuga mertensiana) at the wet end, and both species in the center. Establishment patterns were compared with deviations from the century-long average for these weather variables: winter precipitation, Palmer Drought Severity Index, and winter, October, and May temperatures. Results show that establishment occurred in dry areas when weather conditions were wetter than average, and in wet areas under drier than average conditions. Establishment at central sites did not show consistent relationships with climate. If future climatic conditions continue to warm, establishment of subalpine fir in subalpine meadows in dry areas may cease and mountain hemlock may resume in wet areas.

  4. Declines in low-elevation subalpine tree populations outpace growth in high-elevation populations with warming (United States)

    Conlisk, Erin; Castanha, Cristina; Germino, Matthew J.; Veblen, Thomas T; Smith, Jeremy M.; Kueppers, Lara M.


    Species distribution shifts in response to climate change require that recruitment increase beyond current range boundaries. For trees with long life spans, the importance of climate-sensitive seedling establishment to the pace of range shifts has not been demonstrated quantitatively.Using spatially explicit, stochastic population models combined with data from long-term forest surveys, we explored whether the climate-sensitivity of recruitment observed in climate manipulation experiments was sufficient to alter populations and elevation ranges of two widely distributed, high-elevation North American conifers.Empirically observed, warming-driven declines in recruitment led to rapid modelled population declines at the low-elevation, ‘warm edge’ of subalpine forest and slow emergence of populations beyond the high-elevation, ‘cool edge’. Because population declines in the forest occurred much faster than population emergence in the alpine, we observed range contraction for both species. For Engelmann spruce, this contraction was permanent over the modelled time horizon, even in the presence of increased moisture. For limber pine, lower sensitivity to warming may facilitate persistence at low elevations – especially in the presence of increased moisture – and rapid establishment above tree line, and, ultimately, expansion into the alpine.Synthesis. Assuming 21st century warming and no additional moisture, population dynamics in high-elevation forests led to transient range contractions for limber pine and potentially permanent range contractions for Engelmann spruce. Thus, limitations to seedling recruitment with warming can constrain the pace of subalpine tree range shifts.

  5. Population dynamics and bioenergetics of a fossorial herbivore, Thomomys talpoides (Rodentia: Geomyidae), in a spruce-fir sere (United States)

    Andersen, Douglas C.; MacMahon, James A.


    Studies of the bioenergetics of the northern pocket gopher, Thomomys talpoides, are coupled with data on demography, activity budgets, and microclimates to model the energy requirements of individuals and populations in the Wasatch Mountains of Utah during 1976-1979. Metabolic rates during rest increased linearly with decreasing ambient temperature, but burrowing metabolic rates (16.3 mL O2 • h-1 • g-9.75) were independent of both temperature and physical properties of the soil. Radio-telemetry studies indicated that free-ranging gophers are active =50% of each day. Conservative estimates of true energy consumption were calculated using estimates of habitat-specific minimum daily burrowing requirements. Rates of burrowing measured in the laboratory were either ∞ 0.0 or ∞ 2.0 cm/min. The low burrowing rate was observed when the soil was frozen or saturated with water, as would occur in the field in early winter and in spring, respectively. Gophers burrowed through soil at the study site at an average rate of ∞ 1.5 cm/min. Belowground food energy densities at gopher foraging depth declined from 24.6 to 3.2 J/cm3 along a successional gradient (subalpine forb meadow to Engelmann spruce dominated forest). We conclude that individual gophers are food limited within the climax spruce seral stage. Further, daily energy costs associated with reproduction in females may exceed the belowground energy supply available in intermediate seral stages (aspen and subalpine fir). Reduction of burrowing rates for any reason will affect gophers in the late seral stages proportionately more than those resident in the meadow. The peak gopher densities recorded (from 62 individuals/ha in the meadow to 2 individuals/ha in spruce forest) support these inferences. Detailed demographic information was obtained only in the meadow seral stage. Adult survivorship was lower in winter than in summer and varied greatly between years (0.18-0.70 yr-1). Juvenile survivorship from weaning

  6. The Response of Subalpine Vegetation to Climate Change and Bark Beetle Infestations: A Multi-Scale Interaction. (United States)

    Foster, A.; Shuman, J. K.; Shugart, H. H., Jr.; Negrón, J. F.


    Mean annual temperatures in the western United States have increased in the last few decades, and are predicted to continue warming. In the subalpine zone of the Rocky Mountains, this warming is also predicted to increase the frequency and severity of spruce beetle outbreaks. Climate change itself may affect this vegetation, potentially leading to shifts in species compositions. These forests are a crucial part of the US's carbon budget, thus it is important to analyze how climate change and bark beetles in conjunction will affect the biomass and species composition of vegetation in subalpine zone. UVAFME is an individual-based gap model that simulates biomass and species composition of a forest. This model has been quantitatively tested at various Rocky Mountain sites in the Front Range, and has been shown to accurately simulate the vegetation dynamics in the region. UVAFME has been updated with a spruce beetle subroutine that calculates the probability for beetle infestation of each tree on a plot. This probability is based on site, climate, and individual tree characteristics, such as temperature; stand structure; and tree stress level, size, and age. These governing characteristics are based on data from the US Forest Service, and other studies on spruce susceptibility and spruce beetle phenology. UVAFME is then run with multiple climate change and beetle scenarios to determine the net effect of both variables on subalpine vegetation. These results are compared among the different scenarios and to current forest inventory data. We project that increasing temperatures due to climate change will cause an increase in the frequency and severity of spruce beetle outbreaks, leading to a decrease in the biomass and dominance of Engelmann spruce. These results are an important step in understanding the possible futures for the vegetation of subalpine zone in the Rocky Mountains.

  7. Spruce aphid, Elatobium abietinum (Walker): Life history and damage to Engelmann spruce in the Pinaleno Mountains, Arizona (United States)

    Ann M. Lynch


    Spruce aphid is an exotic insect recently introduced to the Pinaleno Mountains. It feeds on dormant Engelmann spruce, and possible effects include tree-growth suppression, tree mortality, and reduction in seed and cone production. Potential longer-term effects include changes in forest structure and species composition - primarily through reduction in Engelmann spruce...

  8. Resistance to galling adelgids varies among families of Engelmann spruce (Picea engelmani P.) (United States)

    William J. Mattson; Alvin Yanchuk; Gyula Kiss; Bruce Birr


    Cooley gall adelgids, Adelges cooleyi, and round gall adelgids, Adelges abietis, differentially infested 110 half-sib families of Engelmann spruce, Picea engelmannii at 9 study sites in British Columbia. There was a negative genetic correlation (-0.53) between the infestations of the two gall-forming species....

  9. Fire ecology and management of the major ecosystems of southern Utah (United States)

    Sharon M. Hood; Melanie Miller


    This document provides managers with a literature synthesis of the historical conditions, current conditions, fire regime condition classes (FRCC), and recommended treatments for the major ecosystems in southern Utah. Sections are by ecosystems and include: 1) coniferous forests (ponderosa pine, mixed conifer, and Engelmann spruce-subalpine fir), 2) aspen, 3) pinyon-...

  10. Species composition influences management outcomes following mountain pine beetle in lodgepole pine-dominated forests (United States)

    Kristen Pelz; C. C. Rhoades; R. M. Hubbard; M. A. Battaglia; F. W. Smith


    Mountain pine beetle outbreaks have killed lodgepole pine on more than one million hectares of Colorado and southern Wyoming forest during the last decade and have prompted harvest operations throughout the region. In northern Colorado, lodgepole pine commonly occurs in mixed stands with subalpine fir, Engelmann spruce, and aspen. Variation in tree species composition...

  11. Seed maturity in white fir and red fir (United States)

    William W. Oliver


    White fir and red fir seed collected over a 2-month period in northern California was tested for germination of fresh and stratified seed. Ratio of embryo length to embryo cavity length was found to be the most useful index of seed maturity for white and red fir. Cone specific gravity also was correlated with nearly all measures of seed germination. Data suggest that...

  12. Hydrological Features on Subalpine Forest Zone in the East of Qinghai-Tibet Plateau (United States)

    Zhong, X.; Cheng, G.; Guo, W.


    The Hengduan mountain chains of China is situated on the east of Qinghai-Tibet Plateau with area of more than 400,000 km2. Mountains and rivers run through in north-south direction, and are collocated side by side on east-west. Elevation difference between ridges and valleys has great disparity, normally of 1000-2500m, so the vertical zones of vegetation are very distinct. Subalpine coniferous forest zone, mainly composed of fir (Abies) and spruce (Picea), is on altitude of 2800-4200m, which is a chief component of the forested area in southwest China, and an important region for water conservation of several international rivers inlcuding Nujiang River and Lancangjiang River, as well as the world-famous Changjiang River. Thus, it has both theoretical and practical significance to study hydrological process and laws of forest in this region. The study area is located at the Gongga Mountain, on the east edge of the Hengduan mountain chains. Elevation of the main peak is 7556m, and elevation difference between ridge and valley on the eastern slope is 6400m. An ecological observation station was built at altitude of 3000m on the eastern slope of Gongga Mountain in 1988, mainly for alpine ecology and forest hydrology research. Based on the analysis of 20- years observation data from this station, it is revealed that hydrological process of forest in this area has several features as follows: (1) Canopy interception of primitive fir (Abies) forest is obviously greater than other tree species, and interception rate is 30-40%. Maximal canopy interception of one-time precipitation of primitive fir forest is commonly 2-5mm. According to observation data of canopy interception, a conceptual model of canopy interception of fir forest is established: R=1.69[(1-exp(-0.41P))+0.19P (P is precipitation in mm); (2) Natural valid moisture holding capacity in layer of moss-decayed wood and leaves beneath trees is up to 5.6mm. Porosity in soil surface layer and non-capillary porosity

  13. Douglas-Fir Tussock Moth (United States)

    Boyd E. Wickman; Richard R. Mason; Galen C. Trostle


    The Douglas-fir tussock moth (Orgyia pseudotsugata McDunnough) is an important defoliator of true firs and Douglas-fir in Western North America. Severe tussock moth outbreaks have occurred in British Columbia, Idaho, Washington, Oregon, Nevada, California, Arizona, and New Mexico, but the area subject to attack is more extensive

  14. Rooting cuttings from douglas-fir, white-fir, and California red fir christmas trees (United States)

    C. M. Blankensop; R. Z. Callaham


    Christmas tree growers in California have asked geneticists to help improve the characteristics of the - wild species they are cultivating. The preferred Christmas trees of California are Shasta red fir (Abies magnifica A. Murr.), white fir (A. concolor (Gord. & Glend.) Lindl.), and Douglas-fir (Pseudotsuga...

  15. Germination Characteristics of Engelmann Oak and Coast Live Oak from the Santa Rosa Plateau, Riverside County, California (United States)

    Gerald E. Snow


    Over 2,000 acorns of Quercus agrifolia (coast live oak) and over 500 acorns of Q. engelmannii (Engelmann oak) were collected in the Jim Knight pasture area of the Santa Rosa Plateau. These were used to test for temperature and moisture conditions on germination of viable acorns in the laboratory under controlled environmental...

  16. Sensitivity of subalpine tree seedlings and alpine plants to natural and manipulated climate variation: Initial results from an Alpine Treeline Warming Experiment (Invited) (United States)

    Kueppers, L. M.


    Niche models and paleoecological studies indicate that future climate change will alter the geographic distributions of plant species. Changes in temperature, snowmelt timing, or moisture conditions at one edge of a species’ range may have different consequences for recruitment, carbon exchange, phenology, and survival than changes at another edge. Similarly, local genetic adaptation may constrain species and community responses to climate change. We have established a new experiment to investigate potential shifts in the distribution of subalpine tree species, and the alpine species they might replace. We are asking how tree species recruitment and alpine species growth and reproduction vary within their current ranges, and in response to temperature and soil moisture manipulations. We are also examining whether genetic provenance and ecosystem processes constrain tree seedling and alpine herb responses. Our Alpine Treeline Warming Experiment is located across three sites at Niwot Ridge, CO, ranging from near the lower limit of subalpine forest to alpine tundra. We use infrared heaters to raise growing season surface soil temperatures by 4-5°C, and to lengthen the growing season. The warming treatment is crossed with a soil moisture manipulation to distinguish effects due to higher temperatures from those due to drier soil. Each plot is a common garden sown with high and low elevation provenances of limber pine (Pinus flexilis) and Engelmann spruce (Picea engelmannii). We established an additional set of experimental plots to examine treatment effects on alpine species phenology, growth and reproduction. Under ambient conditions in 2009, tree seedling germination rate, lifespan, and first season survival was higher within the species’ current range than in the alpine, and for Engelmann spruce, was higher at the low elevation limit than the high elevation limit. Source population (low vs. high elevation) was a significant factor explaining natural variation in

  17. Camurati-Engelmann's Disease on {sup 99m}Tc-MDP Bone Scan

    Energy Technology Data Exchange (ETDEWEB)

    Yoon, Hai Jeon; Oh, So Won; Paeng, Jin Chul; Lee, You Kyung; Choi, In Ho; Lee, Dong Soo [Seoul National University College of Medicine, Seoul (Korea, Republic of)


    A 24 year-old female presented for a {sup 99m}Tc-methylene diphosphonate (MDP) whole body bone scan due to chronic pain in the bilateral lower extremities that has aggravated since 2002. She was diagnosed with Camurati-Engelmann disease (CED) based on the clinical and radiological findings in 2002, and she re-visited our institute to evaluate disease status at this time. CED is a rare autosomal dominant type of bone dysplasia characterized by progressive cortical thickening of long bones, and narrowing of medullary cavity, and thus presents with typical clinical symptoms and signs such as chronic pain in the extremities, muscle weakness, and waddling gait. On the {sup 99m}Tc-MDP bone scan performed to evaluate disease status, intense increased uptake was seen in the skull, facial bones, bilateral scapulae, bilateral long bones, and bilateral pelvic bones, which clearly demonstrated the extent of CED involvement.

  18. Provenance variation in subalpine fir grown as an exotic tree species in Denmark and Iceland

    DEFF Research Database (Denmark)

    Skúlason, Brynjar

    in a period of 15 growing seasons after establishment. Provenances showed significant differences for all measured traits. The northernmost provenances showed earliest bud set, highest autumn frost tolerance and a latitudinal cline was delineated, while the southernmost provenances showed earliest flushing...

  19. Long-term landscape changes in a subalpine spruce-fir forest in central Utah, USA (United States)

    Jesse L. Morris; R. Justin DeRose; Andrea R. Brunelle


    In Western North America, increasing wildfire and outbreaks of native bark beetles have been mediated by warming climate conditions. Bioclimatic models forecast the loss of key high elevation species throughout the region. This study uses retrospective vegetation and fire history data to reconstruct the drivers of past disturbance and environmental change....

  20. Climate driven changes in Engelmann spruce stands at timberline in the La Sal Mountains (United States)

    James F. Fowler; Steven Overby; Barb Smith


    Due to global warming spruce-fir forest and associated vegetation may experience elevational displacement and altered species composition at the timberline-treeline ecotone. These forests and their component species are predicted to migrate upslope and thus landscape features such as timberline and treeline may move upslope as well. Prior to this study, baseline data...

  1. Comparison of terpene composition in Engelmann spruce (Picea engelmannii) using hydrodistillation, SPME and PLE. (United States)

    Mardarowicz, Marek; Wianowska, Dorota; Dawidowicz, Andrzej L; Sawicki, Ryszard


    Terpenes emitted by conifer trees are generally determined by analysing plant extracts or essential oils, prepared from foliage and cones using steam distillation. The application of these procedures limits experiments to cut plant materials. Recently headspace techniques have been adopted to examine terpene emission by living plants. This paper deals with the application of solid-phase micro-extraction (SPME) for the analysis of terpenes emitted by conifers foliage of Engelmann spruce (Picea engelmannii), including its seedlings. The compositions of SPME extracts obtained for destroyed and non-destroyed old and juvenile spruce needles were compared with the compositions of essential oils and pressurised liquid extraction (PLE) extracts corresponding to the same plant materials. No substantial differences have been found in the qualitative terpene composition estimated by analysing essential oil and PLE and SPME extracts from non-destroyed old and juvenile foliage. The disintegration of spruce needles results in the formation of a significant amount of myrcene in the case of the old conifer foliage and non-terpenoic compounds in the case of juvenile conifer foliage. This phenomenon can be attributed to enzymatic reactions occurring in the destroyed plant cells.

  2. Damage from wind and other causes in mixed white fir-red fir stands adjacent to clearcuttings (United States)

    Donald T. Gordon


    Damage to timber surrounding clearcuttings and in one light selection cutting in mixed white fir-red fir stands was monitored for 6 years in northeastern California. In some years, bark beetles apparently killed more trees than did wind damage, but in two of the study years, severe wind storms caused much damage. One storm produced mainly break-age, apparently...

  3. Leader growth in Nordman fir christmas trees

    DEFF Research Database (Denmark)

    Christensen, Claus Jerram; Pedersen, Lars Bo

    Leader Griowth in Nordman fir Christmas trees: Growth visualization and effects of fertilization, irrigation and drought......Leader Griowth in Nordman fir Christmas trees: Growth visualization and effects of fertilization, irrigation and drought...

  4. Silvical characteristics of balsam fir (Abies balsamea) (United States)

    Arthur C. Hart


    Balsam fir takes its name from the Latin word for balm. Some people know the tree as the Balm-of-Gilead fir. It has also been called the blister fir, because of the bark blisters that yield Canada balsam, a resin that is used for, among other things, mounting microscope slides. The needles of balsam fir have a spicy aroma that Donald Culross Peattie has called "...

  5. Factors influencing the spatial and temporal dynamics of engelmann spruce mortality during a spruce beetle outbreak on the Markagunt Plateau, Utah (United States)

    R. Justin DeRose; James N. Long


    Host conditions are known to influence spruce beetle population levels, but whether they influence the spatial and temporal patterns of beetle-caused mortality during an outbreak is unknown. Using dendrochronological techniques, we quantified the spatiotemporal dynamics of a modern (late 1980s through the early 2000s) spruce beetle outbreak in Engelmann spruce on the...

  6. Can spruce beetle (Dendroctonus rufipennis Kirky) pheromone trap catches or stand conditions predict Engelmann spruce (Picea engelmannii Parry ex Engelm.) tree mortality in Colorado? (United States)

    Jose F. Negron; John B. Popp


    1) Bark beetles (Coleoptera: Curculionidae: Scolytinae) can cause extensive tree mortality in forests dominated by their hosts. Among these, the spruce beetle (Dendroctonus rufipennis) is one of the most important beetles in western North America causing Engelmann spruce (Picea engelmannii) tree mortality. 2) Although pheromone traps with attractants are commonly used...

  7. Evaluation of storage and filtration protocols for alpine/subalpine lake water quality samples (United States)

    John L. Korfmacher; Robert C. Musselman


    Many government agencies and other organizations sample natural alpine and subalpine surface waters using varying protocols for sample storage and filtration. Simplification of protocols would be beneficial if it could be shown that sample quality is unaffected. In this study, samples collected from low ionic strength waters in alpine and subalpine lake inlets...

  8. Ozone and modeled stomatal conductance at a high elevation subalpine site in southeastern Wyoming (United States)

    Robert C. Musselman; Karl F. Zeller; Nedialko T. Nikolov


    Ozone concentrations have been monitored at the Glacier Lakes Ecosystem Experiment Site (GLEES) in the Snowy Range of the Medicine Bow Mountains 55 km west of Laramie, Wyoming, USA. The site is located at 3,186 m elevation in a large subalpine meadow of a mature subalpine forest near timberline. Continuous ozone and meteorological monitoring are a part of the GLEES...

  9. Fire, fuel composition and resilience threshold in subalpine ecosystem.

    Directory of Open Access Journals (Sweden)

    Olivier Blarquez

    Full Text Available BACKGROUND: Forecasting the effects of global changes on high altitude ecosystems requires an understanding of the long-term relationships between biota and forcing factors to identify resilience thresholds. Fire is a crucial forcing factor: both fuel build-up from land-abandonment in European mountains, and more droughts linked to global warming are likely to increase fire risks. METHODS: To assess the vegetation response to fire on a millennium time-scale, we analyzed evidence of stand-to-local vegetation dynamics derived from sedimentary plant macroremains from two subalpine lakes. Paleobotanical reconstructions at high temporal resolution, together with a fire frequency reconstruction inferred from sedimentary charcoal, were analyzed by Superposed Epoch Analysis to model plant behavior before, during and after fire events. PRINCIPAL FINDINGS: We show that fuel build-up from arolla pine (Pinus cembra always precedes fires, which is immediately followed by a rapid increase of birch (Betula sp., then by ericaceous species after 25-75 years, and by herbs after 50-100 years. European larch (Larix decidua, which is the natural co-dominant species of subalpine forests with Pinus cembra, is not sensitive to fire, while the abundance of Pinus cembra is altered within a 150-year period after fires. A long-term trend in vegetation dynamics is apparent, wherein species that abound later in succession are the functional drivers, loading the environment with fuel for fires. This system can only be functional if fires are mainly driven by external factors (e.g. climate, with the mean interval between fires being longer than the minimum time required to reach the late successional stage, here 150 years. CONCLUSION: Current global warming conditions which increase drought occurrences, combined with the abandonment of land in European mountain areas, creates ideal ecological conditions for the ignition and the spread of fire. A fire return interval of less

  10. Thinning balsam fir thickets with soil sterilants (United States)

    Arthur C. Hart


    Under certain conditions that we do not yet fully understand, balsam fir has a tendency to form dense thickets that cause stagnation of growth. This condition is common throughout the spruce-fir region, and it presents the landowner with one of his most perplexing management problems. A typical thicket averaging 15 feet tall may contain 5,000 to 10,000 stems per acre (...

  11. Veneer recovery from Douglas-fir logs. (United States)

    E.H. Clarke; A.C. Knauss


    During 1956, the Pacific Northwest Forest and Range Experiment Station made a series of six veneer-recovery studies in the Douglas-fir region of Oregon and Washington. The net volume of logs involved totaled approximately 777 M board-feet. Purpose of these studies was to determine volume recovery, by grade of veneer, from the four principal grades of Douglas-fir logs...

  12. Radial growth of grand fir and Douglas-fir 10 years after defoliation by the Douglas-fir tussock moth in the Blue Mountains outbreak. (United States)

    Boyd E. Wickman


    Radial-growth recovery related to amount of tree defoliation was measured 10 years after a severe outbreak of Douglas-fir tussock moth (Orgyia pseudotsugata (McDunnough). For the period 1978-82, growth of qrand fir surpassed and was significantly greater than in the preoutbreak period, 1968-72. Douglas-fir growth during the postoutbreak period...

  13. Douglas-fir tussock moth- and Douglas-fir beetle-caused mortality in a ponderosa pine/Douglas-fir forest in the Colorado Front Range, USA (United States)

    Jose F. Negron; Ann M. Lynch; Willis C. Schaupp; Vladimir Bocharnikov


    An outbreak of the Douglas-fir tussock moth, Orgyia pseudotsugata McDunnough, occurred in the South Platte River drainage on the Pike-San Isabel National Forest in the Colorado Front Range attacking Douglas-fir, Pseudotsuga menziesii (Mirb.) Franco. Stocking levels, species composition, and tree size in heavily and lightly defoliated stands were similar. Douglas-fir...

  14. Decay in white fir top-killed by Douglas-fir tussock moth. (United States)

    Boyd E. Wickman; Robert F. Scharpf


    Stands heavily defoliated in 1936-37 by the Douglas-fir tussock moth, Hemerocampa pseudotsugata McD., at Mammoth Lakes, California, were studied to determine the incidence and extent of decay in top-damaged trees. This was done by dissecting the tops of trees felled during logging. Comparisons were made with white fir in a nearby logged area that was...

  15. [Contribution of soil fauna to the mass loss of Betula albosinensis leaf litter at early decomposition stage of subalpine forest litter in western Sichuan]. (United States)

    Xia, Lei; Wu, Fu-Zhong; Yang, Wan-Qin; Tan, Bo


    In order to quantify the contribution of soil fauna to the decomposition of birch (Betula albosinensis) leaf litter in subalpine forests in western Sichuan of Southwest China during freeze-thaw season, a field experiment with different mesh sizes (0.02, 0.125, 1 and 3 mm) of litterbags was conducted in a representative birch-fir (Abies faxoniana) forest to investigate the mass loss rate of the birch leaf litter from 26 October, 2010 to 18 April, 2011, and the contributions of micro-, meso- and macro-fauna to the decomposition of the leaf litter. Over the freeze-thaw season, 11.8%, 13.2%, 15.4% and 19.5% of the mass loss were detected in the litterbags with 0.02, 0. 125, 1 and 3 mm mesh sizes, respectively. The total contribution of soil fauna to the litter decomposition accounted for 39.5% of the mass loss, and the taxa and individual relative density of the soil fauna in the litterbags had the similar variation trend with that of the mass loss rate. The contribution rate of soil fauna to the leaf litter mass loss showed the order of micro- fauna, with the highest contribution of micro-fauna (7.9%), meso-fauna (11.9%), and macro-fauna (22.7%) at the onset of freezing stage, deeply frozen stage, and thawing stage, respectively. The results demonstrated that soil fauna played an important role in the litter decomposition in subalpine forests of western Sichuan during freeze-thaw season.

  16. Comparative phytosocioogical investigation of subalpine alder thickets in southwestern Alaska and the North Pacific (United States)

    US Fish and Wildlife Service, Department of the Interior — We present the first vegetation analysis of subalpine alder (Alnus viridis) thickets in southwestern Alaska. The data are primarily from mesic, hilly and mountainous...

  17. Determination of fir strengh charachteristics


    Газизов, А. М.; Юсупов, С. А.


    Пихтовая древесина имеет наименьшую популярность в области столярных и строительных работ в связи с низкими физико-механическими свойствами. Показателем механических свойств древесины является ее прочность. Fir wood has the least popularity in the field of carpentry and construction in connection with low mechanical properties. An index of mechanical properties of wood is its durability....

  18. HOW to Identify and Manage Needlecast Diseases on Balsam Fir (United States)

    Mike Albers; Jana Albers; Jane Cummings Carlson; Linda Haugen; Nancy Wenner


    Needlecast diseases are common in balsam fir stands and Christmas tree plantations in the northeastern and north central United States and in southern Ontario, Quebec, and New Brunswick. Three different needlecast fungi, Lirula nervata, Lirula mirabilis and Isthmiella faullii, cause similar disease symptoms on balsam fir. These diseases may affect other firs planted in...

  19. FIR-MMC in rantsoenen van roze-vleeskalveren

    NARCIS (Netherlands)

    Plomp, M.; Heeres-van der Tol, J.J.; Driegen, J.


    FIR-MMC is een koolstofhoudend kleimineraal. Ervaringen uit de praktijk zouden aangeven dat FIR-MMC de voederconversie kan verbeteren. Toevoegen van FIR-MMC had in dit onderzoek bij geen van de rantsoenen invloed op voeropname, groei, voederconversie en karkaskwaliteit van de kalveren.

  20. System Verilog modelling of FIR filters (United States)

    Pawlus, Łukasz; Wegrzyn, Marek


    In the paper modelling of FIR filters by means of Verilog and SystemVerilog is presented. Hardware/software co-design approach for such systems is applied in the presented design. As a final technology for a FIR filters system implementation, a FPSLIC device is considered. Filters system demonstrates example methods of communication between FPGA and AVR microcontroller in a FPSLIC structure, i.e. the communication through SRAM memory, addressing lines, data bus, interrupts. It also demonstrates how to serve peripheral elements in FPSLIC device by means of DPI interface. FIR filters model contains also interface which implements a FPSLIC cache logic and gives opportunity to a dynamical reconfiguration of FPGA in a FPSLIC structure.

  1. Douglas-Fir Tussock Moth- and Douglas-Fir Beetle-Caused Mortality in a Ponderosa Pine/Douglas-Fir Forest in the Colorado Front Range, USA

    Directory of Open Access Journals (Sweden)

    José F. Negrón


    Full Text Available An outbreak of the Douglas-fir tussock moth, Orgyia pseudotsugata McDunnough, occurred in the South Platte River drainage on the Pike-San Isabel National Forest in the Colorado Front Range attacking Douglas-fir, Pseudotsuga menziesii (Mirb. Franco. Stocking levels, species composition, and tree size in heavily and lightly defoliated stands were similar. Douglas-fir tussock moth defoliation resulted in significant Douglas-fir mortality in the heavily defoliated stands, leading to a change in dominance to ponderosa pine, Pinus ponderosa Lawson. Douglas-fir beetle, Dendroctonus pseudotsuqae Hopkins, populations increased following the defoliation event but caused less mortality, and did not differ between heavily and lightly defoliated stands. Douglas-fir tussock moth-related mortality was greatest in trees less than 15 cm dbh (diameter at 1.4 m above the ground that grew in suppressed and intermediate canopy positions. Douglas-fir beetle-related mortality was greatest in trees larger than 15 cm dbh that grew in the dominant and co-dominant crown positions. Although both insects utilize Douglas-fir as its primary host, stand response to infestation is different. The extensive outbreak of the Douglas-fir tussock moth followed by Douglas-fir beetle activity may be associated with a legacy of increased host type growing in overstocked conditions as a result of fire exclusion.

  2. Some effects of douglas fir terpenes on certain microorganisms. (United States)

    Andrews, R E; Parks, L W; Spence, K D


    The Douglas fir terpene alpha-pinene was shown to inhibit the growth of a variety of bacteria and a yeast. Other terpenes of the Douglas fir, including limonene, camphene, and isobornyl acetate, were also inhibitory to Bacillus thuringiensis. All terpenes were inhibitory at concentrations normally present in the fir needle diet of Douglas fir tussock moth larvae. The presence of such terpenes in the diet of these insects was found to strongly influence the infectivity of B. thuringiensis spores for the Douglas fir tussock moth larvae. The terpene alpha-pinene destroyed the cellular integrity and modified mitochondrial activity in certain microorganisms.

  3. Management of silver fir (Abies alba Mill.) in Slovenia


    Bončina, Andrej; Ficko, Andrej; Klopčič, Matija; Matijašič, Dragan; Poljanec, Aleš


    In the paper, we analysed the structure and developmental characteristics of forest stands with silver fir in Slovenia, the management and cut in four forest site strata, where silver fir occurs. We used databases from the Slovenia Forest Service. In growing stock (GS) of silver fir, large (dbh=30-49cm) and very large (d=50 cm and more) diameter trees account for 84.9 % at the national level. The highest share of very large diameter silver fir trees (45 %) is in Dinaric silver fir forests and...

  4. Tall oil precursors of Douglas fir (United States)

    Daniel O. Foster; Duane F. Zinkel; Anthony H. Conner


    The sapwood and heartwood extractives of Douglas fir [Pseudotsuga menziesii (Mirb.) Franco] and the tall oil in the kraft black liquor were characterized. On pulping, isomerization and conversion of conjugated resin acids to dehydroabietic acid was observed. Recovery of both fatty and resin acids from pulping was lower than predicted from the extractive composition....


    Energy Technology Data Exchange (ETDEWEB)

    Stephen Benson, David Douglas, George Neil, Michelle D. Shinn, Gwyn Williams


    We describe a dual free-electron laser (FEL) configuration on the UV Demo FEL at Jefferson Lab that allows simultaneous lasing at FIR/THz and UV wavelengths. The FIR/THz source would be an FEL oscillator with a short wiggler providing nearly diffraction-limited pulses with pulse energy exceeding 50 microJoules. The FIR source would use the exhaust beam from a UVFEL. The coherent harmonics in the VUV from the UVFEL are out-coupled through a hole. The FIR source uses a shorter resonator with either hole or edge coupling to provide very high power FIR pulses. Simulations indicate excel-lent spectral brightness in the FIR region with over 100 W/cm-1 output.

  6. Phenology of plants in relation to ambient environment in a subalpine forest of Uttarakhand, western Himalaya. (United States)

    Bisht, Vinod K; Kuniyal, Chandra P; Bhandari, Arvind K; Nautiyal, Bhagwati P; Prasad, P


    Observations on phenology of some representative trees, shrubs, under-shrubs and herbs in a subalpine forest of Uttarakhand, western Himalaya were recorded. With the commencement of favorable growth season in April, occurrence of leaf fall was indicatory growth phenomenon in Quercus semecarpifolia, Q. floribunda and Abies spectabilis. However, active vegetative growth in herbaceous species starts onward April and fruit maturation and seed dehiscence are completed from mid of September to October. In general, vegetative growth and reproductive stages in majority of the studied species seems to be dependent on adequate moisture content and also flowering and fruiting in subalpine plants correlate ambient temperature.

  7. Linkages between grazing history and herbivore exclusion on decomposition rates in mineral soils of subalpine grasslands (United States)

    Alan G. Haynes; Martin Schutz; Nina Buchmann; Deborah S. Page-Dumroese; Matt D. Busse; Anita C. Risch


    Herbivore-driven changes to soil properties can influence the decomposition rate of organic material and therefore soil carbon cycling within grassland ecosystems. We investigated how aboveground foraging mammalian and invertebrate herbivores affect mineral soil decomposition rates and associated soil properties in two subalpine vegetation types (shortgrass and tall-...

  8. Human disturbance provides foraging opportunities for birds in primary subalpine forest

    DEFF Research Database (Denmark)

    DuBay, Shane G.; Hart Reeve, Andrew; Wu, Yongjie


    to species that naturally occur in edge, open, or disturbed habitats. With observations and experiments we provide evidence of insectivorous birds exploiting human disturbance in primary subalpine forest in the mountains of southern China, displaying behavioral flexibility to gain novel foraging...

  9. Net primary productivity of subalpine meadows in Yosemite National Park in relation to climate variability (United States)

    Peggy E. Moore; Jan W. van Wagtendonk; Julie L. Yee; Mitchel P. McClaran; David N. Cole; Neil K. McDougald; Matthew L. Brooks


    Subalpine meadows are some of the most ecologically important components of mountain landscapes, and primary productivity is important to the maintenance of meadow functions. Understanding how changes in primary productivity are associated with variability in moisture and temperature will become increasingly important with current and anticipated changes in climate....

  10. Drought-driven disturbance history characterizes a southern Rocky Mountain subalpine forest (United States)

    R. Justin DeRose; James N. Long


    The view that subalpine forest vegetation dynamics in western North America are "driven" by a particular disturbance type (i.e., fire) has shaped our understanding of their disturbance regimes. In the wake of a recent (1990s) landscape- extent spruce beetle (Dendroctonus rufipennis Kirby) outbreak in the southern Rocky Mountains, we re-examined the temporal...

  11. Water use patterns of three species in subalpine forest, Southwest China: the deuterium isotope approach (United States)

    Qing Xu; Harbin Li; Jiquan Chen; Jiquan Cheng; Xiaoli Cheng; Shirong Liu; Shuqing An


    Determination of water sources of plant species in a community is critical for understanding the hydrological processes and their importance in ecosystem functions. Such partitioning of plant xylem water into specific sources (i.e. precipitation, groundwater) can be achieved by analyzing deuterium isotopic composition (δD) values for source waters. A subalpine dark...

  12. Comparison of the abundance and composition of litter fauna in tropical and subalpine forests (United States)

    G. Gonzalez; T.R. Seastedt


    In this study, we quantify the abundance and composition of the litter fauna in dry and wet tropical forests and north- and south-facing subalpine forests. We used the same litter species contained in litterbags across study sites to standardize for substrate conditions, and a single method of fauna extraction from the litter (Tullgren method). Fauna densities were...

  13. Shelterwood regeneration of true fir: conclusion after 8 years (United States)

    Robert J. Laacke; Jeanne H. Tomascheski


    Shelterwood cuttings on Swain Mountain Experimental Forest were measured to determine performance six to eight years after the shelterwood cutting and before shelterwood removal. Use of appropriate selecting criteria minimized windthrow of seed trees. Regeneration remained red fir, even where the major seed source was white fir. Density of seed trees may affect...

  14. Balsam Fir Wooly Adlegid Survey on CVNWR 2009 (United States)

    US Fish and Wildlife Service, Department of the Interior — On July 17, 2009 Amber Myers and I conducted a survey to check the status of Balsam Fir Wooly Adelgid on one hundred Balsam Fir Trees off of Freeland Trail. We...

  15. Financial analysis of pruning coast Douglas-fir. (United States)

    Roger D. Fight; James M. Cahlll; Thomas D. Fahey; Thomas A. Snellgrove


    Pruning of coast Douglas-fir was evaluated; recent product recovery information for pruned and unpruned logs for both sawn and peeled products was used. Dimensions of pruned and unpruned trees were simulated with the Douglas-fir stand simulator (DFSIM). Results are presented for a range of sites, ages at time of pruning, ages at time of harvest, product prices, and...

  16. A model to estimate noble fir bough weight. (United States)

    Keith A. Blatner; Roger D. Fight; Nan Vance; Mark Savage; Roger. Chapman


    The harvesting of noble fir (Abies procera) for the production of Christmas wreaths and related products has been a mainstay of the nontimber forest products industry in the Pacific Northwest (PNW) for decades. Although noble fir is the single most important bough product harvested in the PNW, little or no work has been published concerning the...

  17. First report of Phytophthora ramorum infecting grand fir in California (United States)

    K.L. Riley; G.A. Chastagner


    Phytophthora ramorum was detected on grand fir in 2003 and 2005 in a Christmas tree plantation near Los Gatos, CA, in association with infected California bay laurel. Isolates derived from stem lesions were used to inoculate grand fir seedlings in two tests. Isolations from lesions on inoculated plants were positive for P. ramorum...

  18. Mortality and growth reduction of white fir following defoliation by the Douglas-fir tussock moth (United States)

    Boyd E. Wickman


    In 5 years after a 1954-56 outbreak of Hemerocampa pseudotsugata in Calaveras and Tuolumne Counties, California, 20 percent of the merchantable white fir, or 11,071 board feet per acre, died in heavily defoliated stands. Another 1,113 board feet per acre was lost owing to radial growth reductions in partly defoliated trees; 12 percent of these trees...

  19. Human-Related Forest Fires in the Subalpine Belt of the Spanish Pyrenees (United States)

    Sanjuán, Yasmina; María García-Ruiz, José; Beguería, Santiago; Serrano-Muela, María Pilar; González-Sampériz, Penélope; Valero-Garcés, Blas; Arnáez, José


    The subalpine belt of the Central Pyrenees ranges approximately between 1700 and 2200 m a.s.l. This area should be covered with dense forests of Pinus sylvestris and P. uncinata, with increasingly open formations towards the upper forest limit. At present, most of the subalpine belt is occupied with grasslands due to human-induced deforestation for enlarging the area occupied by summer pastures. Two are the most important scientific problems related to deforestation of the subalpine belt: (i) the timing of deforestation, and (ii) the geomorphic consequences of a sudden substitution of forests by grasslands. Up to now, intense deforestation is clearly recorded in regional palaeoenvironmental sequences since the Middle Ages and, traditionally, this practice was usually attributed to large fires with the purpose of balance the winter and summer pasture resources. Nevertheless, the presence of abundant remnants of prehistoric monuments (dolmens, cromlechs, tumulus) in the subalpine belt induced to think in a previous seasonal presence of human populations, most probably practicing some primitive type of transhumance. This would only be possible if part of the subalpine forests would be burnt to allow a limited expansion of grasslands, despite the consequences in the landscape of this kind of practices were not permanent in time. We present here new dates of fire occurrence from charcoal obtained from soils in the hillslopes and from lacustrine sediments. Two periods of human-induced fires have been identified: (i) between 2500 and 2000 cal. yr BP, and (ii) between 1100 and 900 cal yr BP. The consequences of deforestation can be easily observed in the landscape, particularly shallow landslide activity, gelifluction, solifluction and the rapid development of parallel incisions in the steepest slopes.

  20. Manipulation of subalpine and alpine microclimate in the Alpine Treeline Warming Experiment (United States)

    Kueppers, L. M.; Moyes, A. B.; Ferrenberg, S. M.; Christianson, D. S.; Castanha, C.; Germino, M. J.


    To experimentally test model projections of subalpine tree species' uphill migration with climate change, we have established the Alpine Treeline Warming Experiment at Niwot Ridge, CO. Common gardens subject to full factorial warming and watering experiments are replicated across three sites: near the lower limit of subalpine forest, within the alpine-treeline ecotone, and in the alpine tundra, beyond the current elevation ranges of the species. In 2010, differences in ambient climate among the three sites included 5.1° C greater growing-season air temperature and 0.5 kPa greater vapor pressure deficit in the lowest compared to the upper two sites. The lower subalpine site also experienced lower soil moisture compared to the upper two sites. Snowmelt date varied substantially between sites, with the longest snow-free period in the lower subalpine site and the shortest in the treeline site. In all sites, we observed advances in the timing of snowmelt in heated relative to control plots. The warming treatment also raised 5 cm soil temperatures by 3° C at the lower subalpine site, and by 1° C in the upper two sites, averaged over the growing season. More substantial wind in the alpine diminished the heating effect through sensible heat loss. Seasonal average volumetric soil moisture at 5-10 cm did not vary strongly among treatments even though seedling survival and gas exchange data suggest that water additions alleviated drought stress in some plots. These preliminary findings for identical warming and watering treatments across our three high mountain sites suggest that microclimate responses vary with radiation environment, patterns of snow accumulation, and wind speed. Some of these differences are realistic for a future warmer world, while others are artifacts of the experimental approach. Microclimate differences in 2011 reflect modified heating methods and a different spring hydroclimate (later snow accumulation and melt), highlighting the importance of

  1. A Comparison of Infrared Gas Analyzers Above a Subalpine Forest (United States)

    Burns, S. P.; Metzger, S.; Blanken, P.; Burba, G. G.; Swiatek, E.; Li, J.; Conrad, B.


    Infrared gas analyzers (IRGAs) are a key component in theeddy-covariance measurement of water vapor and carbon dioxide exchangebetween the surface and atmosphere. Historically, closed-path IRGAshave been used for the fast (> 10 Hz) measurement of atmospheric H2Oand CO2. In order to use them in the field, these IRGAs were typicallyhoused in temperature-controlled enclosures or buildings that weretens of meters away from the measurement location. This necessitatedthe use of long tubing and pumps to bring the air sample to the IRGAcell. Attenuation of H2O and CO2 fluctuations within the tubing was apersistent problem with such a setup, especially for H2O. As analternative, open-path IRGAs have frequently been utilized, but thekey trade-offs with the open-path design are: (i) precipitation anddew-related data gaps, and (ii) the need to account for WPL densityeffects. Over the past five years a new type of closed-path IRGA hasemerged which is weather-proof, compact, and low-maintenance. Becauseof its small size, short intake tubing can be used, which places thesampling cell close to the sonic anemometer and reduces high frequencysignal loss. Two such IRGAs are the LI-COR LI-7200 and the CampbellScientific EC155, which is part of the CPEC200 eddy covariance system.The Niwot Ridge AmeriFlux tower has used a LI-COR LI-6262 IRGA tomeasure CO2 fluxes above a subalpine forest since November, 1998.Starting in summer 2013, a LI-7200 (along with an open-path LI-7500)were deployed at 21.5 m on the AmeriFlux tower. In Fall 2013, aEC155/CPEC200 was added so that a side-by-side comparison between allfour IRGAs was possible. The preliminary results presented in ourstudy compare the CO2 and H2O mean and variance measured by each IRGA,the vertical wind statistics from three side-by-side sonicanemometers, as well as the corresponding spectra and cospectra fromthese sensors as well as other important aspects of systemperformance.

  2. Growth of White fir after Douglas-fir tussock moth outbreaks: long-term records in the Sierra Nevada. (United States)

    Boyd E. Wickman


    Radial growth of white fir trees, Abies concolor (Gord. and Glend.) Lindl. ex Hildebr., defoliated almost 30 years ago by Douglas-fir tussock moth, Orgyia pseudotsugata (McDunnough), in the central Sierra Nevada was compared with 22 years of growth prior to the outbreak. There was little difference in growth between the two...

  3. Tree growth in thinned and unthinned White fir stands 20 years after a Douglas-fir tussock moth outbreak. (United States)

    Boyd E. Wickman


    Twenty-year postoutbreak growth was compared in thinned and unthinned, severely defoliated stands. Basal area of unthinned white fir has declined 37 percent and pine basal area has increased 32 percent since 1964. The stand thinned in 1960 has the lowest basal area in the study area, but the greatest tree growth before and after the outbreak. All defoliated fir are...

  4. Symptoms associated with inoculation of stems on living Douglas-fir and Grand Fir Trees with Phytophthora ramorum (United States)

    Gary Chastagner; Kathy Riley; Katie Coats; Marianne Elliott; Annie DeBauw; Norm Dart


    To obtain a better understanding of the potential risk of infection and colonization of living Douglas-fir (Pseudotsuga menziesii) and grand fir (Abies grandis) stems, the stems on over 150 trees of each species were inoculated at a Christmas tree farm near Los Gatos, California. This study had the following objectives: 1)...

  5. Silver fir and Douglas fir are more tolerant to extreme droughts than Norway spruce in south-western Germany. (United States)

    Vitali, Valentina; Büntgen, Ulf; Bauhus, Jürgen


    Improving our understanding of the potential of forest adaptation is an urgent task in the light of predicted climate change. Long-term alternatives for susceptible yet economically important tree species such as Norway spruce (Picea abies) are required, if the frequency and intensity of summer droughts will continue to increase. Although Silver fir (Abies alba) and Douglas fir (Pseudotsuga menziesii) have both been described as drought-tolerant species, our understanding of their growth responses to drought extremes is still limited. Here, we use a dendroecological approach to assess the resistance, resilience, and recovery of these important central Europe to conifer species the exceptional droughts in 1976 and 2003. A total of 270 trees per species were sampled in 18 managed mixed-species stands along an altitudinal gradient (400-1200 m a.s.l.) at the western slopes of the southern and central Black Forest in southwest Germany. While radial growth in all species responded similarly to the 1976 drought, Norway spruce was least resistant and resilient to the 2003 summer drought. Silver fir showed the overall highest resistance to drought, similarly to Douglas fir, which exhibited the widest growth rings. Silver fir trees from lower elevations were more drought prone than trees at higher elevations. Douglas fir and Norway spruce, however, revealed lower drought resilience at higher altitudes. Although the 1976 and 2003 drought extremes were quite different, Douglas fir maintained consistently the highest radial growth. Although our study did not examine population-level responses, it clearly indicates that Silver fir and Douglas fir are generally more resistant and resilient to previous drought extremes and are therefore suitable alternatives to Norway spruce; Silver fir more so at higher altitudes. Cultivating these species instead of Norway spruce will contribute to maintaining a high level of productivity across many Central European mountain forests under

  6. Temporal dynamics of abiotic and biotic factors on leaf litter of three plant species in relation to decomposition rate along a subalpine elevation gradient.

    Directory of Open Access Journals (Sweden)

    Jianxiao Zhu

    Full Text Available Relationships between abiotic (soil temperature and number of freeze-thaw cycles or biotic factors (chemical elements, microbial biomass, extracellular enzymes, and decomposer communities in litter and litter decomposition rates were investigated over two years in subalpine forests close to the Qinghai-Tibet Plateau in China. Litterbags with senescent birch, fir, and spruce leaves were placed on the forest floor at 2,704 m, 3,023 m, 3,298 m, and 3,582 m elevation. Results showed that the decomposition rate positively correlated with soil mean temperature during the plant growing season, and with the number of soil freeze-thaw cycles during the winter. Concentrations of soluble nitrogen (N, phosphorus (P and potassium (K had positive effects but C:N and lignin:N ratios had negative effects on the decomposition rate (k, especially during the winter. Meanwhile, microbial biomass carbon (MBC, N (MBN, and P (MBP were positively correlated with k values during the first growing season. These biotic factors accounted for 60.0% and 56.4% of the variation in decomposition rate during the winter and the growing season in the first year, respectively. Specifically, litter chemistry (C, N, P, K, lignin, C:N and lignin:N ratio independently explained 29.6% and 13.3%, and the microbe-related factors (MBC, MBN, MBP, bacterial and fungal biomass, sucrase and ACP activity explained 22.9% and 34.9% during the first winter and the first growing season, respectively. We conclude that frequent freeze-thaw cycles and litter chemical properties determine the winter decomposition while microbe-related factors play more important roles in determining decomposition in the subsequent growing season.

  7. Grand Fir Nutrient Management in the Inland Northwestern USA

    Directory of Open Access Journals (Sweden)

    Dennis R. Parent


    Full Text Available Grand fir (Abies grandis (Douglas ex D. Don Lindley is widely distributed in the moist forests of the Inland Northwest. It has high potential productivity, its growth being nearly equal to western white pine, the most productive species in the region. There are large standing volumes of grand fir in the region. Nutritionally, the species has higher foliage cation concentrations than associated conifers, especially potassium (K and calcium (Ca. In contrast, it has lower nitrogen (N foliage concentrations, which creates favorable nutrient balance on N-limited sites. Despite concentration differences, grand fir stores proportionally more nutrients per tree than associated species because of greater crown biomass. Although few fertilization trials have examined grand fir specifically, its response is inferred from its occurrence in many monitored mixed conifer stands. Fertilization trials including grand fir either as a major or minor component show that it has a strong diameter and height growth response ranging from 15% to 50% depending in part on site moisture availability and soil geology. Grand fir tends to have a longer response duration than other inland conifers. When executed concurrently with thinning, fertilization often increases the total response. Late rotation application of N provides solid investment returns in carefully selected stands. Although there are still challenges with the post-fertilization effects on tree mortality, grand fir will continue to be an important species with good economic values and beneficial responses to fertilization and nutrient management.

  8. Holocene vegetation and fire regimes in subalpine and mixed conifer forests, southern Rocky Mountains, USA (United States)

    Anderson, R. Scott; Allen, Craig D.; Toney, J.L.; Jass, R.B.; Bair, A.N.


    Our understanding of the present forest structure of western North America hinges on our ability to determine antecedent forest conditions. Sedimentary records from lakes and bogs in the southern Rocky Mountains of Colorado and New Mexico provide information on the relationships between climate and vegetation change, and fire history since deglaciation. We present a new pollen record from Hunters Lake (Colorado) as an example of a high-elevation vegetation history from the southern Rockies. We then present a series of six sedimentary records from ???2600 to 3500-m elevation, including sites presently at the alpine?subalpine boundary, within the Picea engelmannii?Abies lasiocarpa forest and within the mixed conifer forest, to determine the history of fire in high-elevation forests there. High Artemisia and low but increasing percentages of Picea and Pinus suggest vegetation prior to 13 500 calendar years before present (cal yr BP) was tundra or steppe, with open spruce woodland to ???11 900 cal yr BP. Subalpine forest (Picea engelmannii, Abies lasiocarpa) existed around the lake for the remainder of the Holocene. At lower elevations, Pinus ponderosa and/or contorta expanded 11 900 to 10 200 cal yr BP; mixed conifer forest expanded ???8600 to 4700 cal yr BP; and Pinus edulis expanded after ???4700 cal yr BP. Sediments from lake sites near the alpine?subalpine transition contained five times less charcoal than those entirely within subalpine forests, and 40 times less than bog sites within mixed conifer forest. Higher fire episode frequencies occurred between ???12 000 and 9000 cal yr BP (associated with the initiation or expansion of south-west monsoon and abundant lightning, and significant biomass during vegetation turnover) and at ???2000?1000 cal yr BP (related to periodic droughts during the long-term trend towards wetter conditions and greater biomass). Fire episode frequencies for subalpine?alpine transition and subalpine sites were on average 5 to 10 fire

  9. Variation in winter snowpack depth and duration influences summer soil respiration in a subalpine meadow (United States)

    Arnold, C. L.; Ghezzehei, T. A.; Berhe, A. A.


    Subalpine meadows in the Sierra Nevada rely on the depth and duration of the winter snowpack to supply ample water to restore the water table in the meadow during the spring snowmelt. This study examines the role that interannual variability in the winter snowpack plays in the overall rate of summer soil respiration along a hydrologic gradient in a subalpine meadow. Carbon dioxide efflux from the meadow was measured from June through September in 2011 and 2012 using soil collars and a LICOR 8100A infrared gas analyzer. Preliminary results show that soil respiration rates are influenced by the hydrologic gradient across the meadow, with drier regions peaking earlier in the summer as compared to wetter regions. We also show that high snowpack years can suppress soil respiration in the meadow until late in the summer season as compared to low snowpack years, where soil respiration peaks early in the summer.

  10. Phenology of plants in relation to ambient environment in a subalpine forest of Uttarakhand, western Himalaya


    Bisht, Vinod K.; Kuniyal, Chandra P.; Bhandari, Arvind K.; Nautiyal, Bhagwati P.; Prasad, P.


    Observations on phenology of some representative trees, shrubs, under-shrubs and herbs in a subalpine forest of Uttarakhand, western Himalaya were recorded. With the commencement of favorable growth season in April, occurrence of leaf fall was indicatory growth phenomenon in Quercus semecarpifolia, Q. floribunda and Abies spectabilis. However, active vegetative growth in herbaceous species starts onward April and fruit maturation and seed dehiscence are completed from mid of September to Octo...

  11. Bleaching of leaf litter and associated microfungi in subboreal and subalpine forests. (United States)

    Hagiwara, Yusuke; Matsuoka, Shunsuke; Hobara, Satoru; Mori, Akira S; Hirose, Dai; Osono, Takashi


    Fungal decomposition of lignin leads to the whitening, or bleaching, of leaf litter, especially in temperate and tropical forests, but less is known about such bleaching in forests of cooler regions, such as boreal and subalpine forests. The purposes of the present study were to examine the extent of bleached area on the surface of leaf litter and its variation with environmental conditions in subboreal and subalpine forests in Japan and to examine the microfungi associated with the bleaching of leaf litter by isolating fungi from the bleached portions of the litter. Bleached area accounted for 21.7%-32.7% and 2.0%-10.0% of total leaf area of Quercus crispula and Betula ermanii, respectively, in subboreal forests, and for 6.3% and 18.6% of total leaf area of B. ermanii and Picea jezoensis var. hondoensis, respectively, in a subalpine forest. In subboreal forests, elevation, C/N ratio and pH of the FH layer, and slope aspect were selected as predictor variables for the bleached leaf area. Leaf mass per area and lignin content were consistently lower in the bleached area than in the nonbleached area of the same leaves, indicating that the selective decomposition of acid unhydrolyzable residue (recalcitrant compounds such as lignin, tannins, and cutins) enhanced the mass loss of leaf tissues in the bleached portions. Isolates of a total of 11 fungal species (6 species of Ascomycota and 5 of Basidiomycota) exhibited leaf-litter-bleaching activity under pure culture conditions. Two fungal species (Coccomyces sp. and Mycena sp.) occurred in both subboreal and subalpine forests, which were separated from each other by approximately 1100 km.

  12. Genecology of Douglas fir in western Oregon and Washington. (United States)

    J. Bradley St Clair; Nancy L. Mandel; Kenneth W. Vance-Borland


    Background and Aims. Genecological knowledge is important for understanding evolutionary processes and for managing genetic resources. Previous studies of coastal Douglas fir (Pseudotsuga menziesii var. menziesii) have been inconclusive with respect to geographical patterns of variation, due in part to...

  13. Growth after thinning in 110-year old Douglas-fir. (United States)

    Vern P. Yerkes


    A 6-year record of growth after thinning indicates that 110-year-old Douglas-fir stands on site 111 land have passed the age when growth capacity can be fully transferred to fewer stems through thinning.

  14. Understanding the physiology of postharvest needle abscission in balsam fir

    Directory of Open Access Journals (Sweden)

    Mason Thane Macdonald


    Full Text Available Balsam fir (Abies balsamea trees are commonly used as a specialty horticultural species for Christmas trees and associated greenery in eastern Canada and United States. Postharvest needle abscission has always been a problem, but is becoming an even bigger challenge in recent years presumably due to increased autumn temperatures and earlier harvesting practices. An increased understanding of postharvest abscission physiology in balsam fir may benefit the Christmas tree industry while simultaneously advancing our knowledge in senescence and abscission of conifers in general. Our paper describes the dynamics of needle abscission in balsam fir while identifying key factors that modify abscission patterns. Concepts such as genotypic abscission resistance, nutrition, environmental factors, and postharvest changes in water conductance and hormone evolution are discussed as they relate to our understanding of the balsam fir abscission physiology. Our paper ultimately proposes a pathway for needle abscission via ethylene and also suggests other potential alternative pathways based on our current understanding.


    Morphological differences between old growth and sapling (Pseudotsuga menziesii, (Mirb.) Franco) Douglas fir trees may extend to differences in needle anatomy. We used microscopy with image analysis to compare and quantify anatomical parameters in cross-sections of previous year...

  16. Simulations of the Newark FIR FEL Operation

    CERN Document Server

    Zheng, Jianjun


    The operation of the NJIT/Rutgers FIR FEL is simulated for the first time sing the slightly modified UH code. The lasering behavior is explored for a wide range of parameters. Particularly, we studied the effects on the operation of the microtron based FEL due to the electron beam pulse phase instability which is possibly caused by the microtron cathode back heating. The study shows that for a small phase slew, systematic or random, there is little effect on the normal operation of the FEL. However, a dramatic phase slew of the e-pulse lasting for 1μs kills the operation of the FEL. We estimated the tolerance of the phase instability and discussed the proper operation condition of the device.

  17. Coaxial visible and FIR camera system with accurate geometric calibration (United States)

    Ogino, Yuka; Shibata, Takashi; Tanaka, Masayuki; Okutomi, Masatoshi


    A far-infrared (FIR) image contains important invisible information for various applications such as night vision and fire detection, while a visible image includes colors and textures in a scene. We present a coaxial visible and FIR camera system accompanied to obtain the complementary information of both images simultaneously. The proposed camera system is composed of three parts: a visible camera, a FIR camera, and a beam-splitter made from silicon. The FIR radiation from the scene is reflected at the beam-splitter, while the visible radiation is transmitted through this beam-splitter. Even if we use this coaxial visible and FIR camera system, the alignment between the visible and FIR images are not perfect. Therefore, we also present the joint calibration method which can simultaneously estimate accurate geometric parameters of both cameras, i.e. the intrinsic parameters of both cameras and the extrinsic parameters between both cameras. In the proposed calibration method, we use a novel calibration target which has a two-layer structure where thermal emission property of each layer is different. By using the proposed calibration target, we can stably and precisely obtain the corresponding points of the checker pattern in the calibration target from both the visible and the FIR images. Widely used calibration tools can accurately estimate both camera parameters. We can obtain aligned visible and FIR images by the coaxial camera system with precise calibration using two-layer calibration target. Experimental results demonstrate that the proposed camera system is useful for various applications such as image fusion, image denoising, and image up-sampling.

  18. Subthreshold FIR Filter Architecture for Ultra Low Power Applications


    Mishra, Biswajit; M. Al-Hashimi, Bashir


    Subthreshold design has been proposed as an effective technique for designing signal processing circuits needed in wireless sensor nodes powered by sources with limited energy. In this paper we propose a subthreshold FIR architecture which brings the benefits of reducedleakage energy, reduced minimum energy point, reduced operating voltage and increased operating frequency when compared with recently reported subthreshold designs. We shall demonstrate this through the design of a 9-tap FIR fi...

  19. FIR filter-based online jerk-constrained trajectory generation


    BESSET, Pierre; BEAREE, Richard


    International audience; In the context of human-robot manipulation interaction for service or industrial robotics, the robot controller must be able to quickly react to unpredictable events in dynamic environments. In this paper, a FIR filter-based trajectory generation methodology is presented, combining the simplicity of the analytic second-order trajectory generation, i.e. acceleration-limited trajectory, with the flexibility and computational efficiency of FIR filtering, to generate on th...

  20. Characterizing water, energy and CO2 exchange for a Sky Island subalpine forest in the southwestern USA (United States)

    Brown-Mitic, C. M.; Shuttleworth, J.; Harlow, C.; Bales, R.


    Among the ecosystems present in the semi-arid environment of the Southwestern U.S., Sky Island forest is unique and it has a unique relationship to the sparse surface-water resources available in the region. This ecosystem exists only at the top of mountains because it is only here that, as a long-term average, precipitation input exceeds evapotranspiration to the extent that forest vegetation can survive. Sky Island Forests, therefore, command potentially significant source areas for the water (some originally falling as snow) that ultimately leaves topographically high ground to recharge aquifers in the plains below by mountain-front recharge. They are also very recently recognized as important carbon sinks where very little or no understanding exist of the exchange/cycling dynamics. The Mount Bigelow project provides an empirically based understanding of the hydro- micrometeorological dynamics of a sky island sub-alpine forest in the southwestern U.S. It is the first study to attempt to document, understand, and model the water, energy, and (related) carbon exchanges of the uniquely interesting and, from the water resource standpoint, uniquely important Sky Island forest ecosystem. The fundamental science issues addressed are: the characteristics of the surface-atmosphere exchanges of water, energy and carbon; the storage of moisture and energy in plants and soil; partitioning of winter snow and rain between evapotranspiration/sublimation, deep drainage and the near-surface environmental water resource that sustains the forest. In order to achieve our objective, a network of four below canopy hydro-micrometeorological stations 10 m tall, and one above canopy 30 m tall high resolution eddy correlation tower, were deployed within a predominantly douglas fir/pine second growth forest. Our observations indicate that the surface flux potential (i.e. sensible heat flux (H) potential), defined as the surface-air temperature gradient varies significantly over space as

  1. A small animal model study of perlite and fir bark dust on guinea pig lungs. (United States)

    McMichael, R F; DiPalma, J R; Blumenstein, R; Amenta, P S; Freedman, A P; Barbieri, E J


    Fir bark (Abies) and perlite (noncrystalline silicate) dusts have been reported to cause pulmonary disease in humans. Guinea pigs were exposed to either fir bark or perlite dust in a special chamber. Severe pathologic changes occurred in the lungs, consisting of lymphoid aggregated and a perivascular inflammatory response. Both dusts caused similar changes although one was vegetable (fir bark) and the other mineral (perlite). Fir bark and perlite dust appeared to be more than just nuisance dusts.

  2. Stability of nuclear DNA content among divergent and isolated populations of Fraser fir (United States)

    L.D. Auckland; J.S. Johnston; H.J. Price; F.E. Bridgwater


    Fraser fir (Abies fraseri (Pursh) Poir.) is an endemic species consisting of six major disjunct populations in the Appalachian Mountains, U.S.A. Nuclear DNA content was measured with laser flow cytometry to determine if genome size differences could be detected among the disjunct populations of Fraser fir and its close relatives, balsam fir

  3. Resurgence of human bothriocephalosis (Diphyllobothrium latum in the subalpine lake region

    Directory of Open Access Journals (Sweden)



    Full Text Available Human bothriocephalosis is once again being found in various catchment basins in the subalpine region, including Lago Maggiore and Lac Léman, which however are not isolated cases. Domestic animals are thought to be responsible for the survival of the parasite during the period when no human cases were reported. The new phenomenon of eating raw or poorly cooked fish is responsible for the resurgence of human bothriocephalosis, which affects various lake districts in Europe. This habit of eating raw fish might lead to the resurgence of a much more dangerous human parasitosis, transmitted in a similar way: infestation by Anisakis.

  4. Gospodarjenje z jelko v Sloveniji: Management of silver fir (Abies alba Mill.) in Slovenia:


    Bončina, Andrej; Ficko, Andrej; Klopčič, Matija; Matijašič, Dragan; Poljanec, Aleš


    In the paper, we analysed the structure and developmental characteristics of forest stands with silver fir in Slovenia, the management and cut in four forest site strata, where silver fir occurs. We used databases from the Slovenia Forest Service. In growing stock (GS) of silver fir, large (dbh=30-49cm) and very large (d=50 cm and more) diameter trees account for 84.9 % at the national level. The highest share of very large diameter silver fir trees (45 %) is in Dinaric silver fir forests and...

  5. Phylogeny and ecophysiology of opportunistic "snow molds" from a subalpine forest ecosystem. (United States)

    Schmidt, S K; Wilson, K L; Meyer, A F; Gebauer, M M; King, A J


    Mats of coenocytic "snow molds" are commonly observed covering the soil and litter of alpine and subalpine areas immediately following snow melt. Here, we describe the phylogenetic placement, growth rates, and metabolic potential of cold-adapted fungi from under-snow mats in the subalpine forests of Colorado. SSU rDNA sequencing revealed that these fungi belong to the zygomycete orders Mucorales and Mortierellales. All of the isolates could grow at temperatures observed under the snow at our sites (0 degrees C and -2 degrees C) but were unable to grow at temperatures above 25 degrees C and were unable to grow anaerobically. Growth rates for these fungi were very high at -2 degrees C, approximately an order of magnitude faster than previously studied cold-tolerant fungi from Antarctic soils. Given the rapid aerobic growth of these fungi at low temperatures, we propose that they are uniquely adapted to take advantage of the flush of nutrient that occurs at the soil-snow interface beneath late winter snow packs. In addition, extracellular enzyme production was relatively high for the Mucorales, but quite low for the Mortierellales, perhaps indicating some niche separation between these fungi beneath the late winter snow pack.

  6. Subalpine Pyrenees received higher nitrogen deposition than predicted by EMEP and CHIMERE chemistry-transport models (United States)

    Boutin, Marion; Lamaze, Thierry; Couvidat, Florian; Pornon, André


    Deposition of reactive nitrogen (N) from the atmosphere is expected to be the third greatest driver of biodiversity loss by the year 2100. Chemistry-transport models are essential tools to estimate spatially explicit N deposition but the reliability of their predictions remained to be validated in mountains. We measured N deposition and air concentration over the subalpine Pyrenees. N deposition was found to range from 797 to 1,463 mg N m-2 year-1. These values were higher than expected from model predictions, especially for nitrate, which exceeded the estimations of EMEP by a factor of 2.6 and CHIMERE by 3.6. Our observations also displayed a reversed reduced-to-oxidized ratio in N deposition compared with model predictions. The results highlight that the subalpine Pyrenees are exposed to higher levels of N deposition than expected according to standard predictions and that these levels exceed currently recognized critical loads for most high-elevation habitats. Our study reveals a need to improve the evaluation of N deposition in mountains which are home to a substantial and original part of the world’s biodiversity.

  7. Biogeochemical impacts of wildfires over four millennia in a Rocky Mountain subalpine watershed. (United States)

    Dunnette, Paul V; Higuera, Philip E; McLauchlan, Kendra K; Derr, Kelly M; Briles, Christy E; Keefe, Margaret H


    Wildfires can significantly alter forest carbon (C) storage and nitrogen (N) availability, but the long-term biogeochemical legacy of wildfires is poorly understood. We obtained a lake-sediment record of fire and biogeochemistry from a subalpine forest in Colorado, USA, to examine the nature, magnitude, and duration of decadal-scale, fire-induced ecosystem change over the past c. 4250 yr. The high-resolution record contained 34 fires, including 13 high-severity events within the watershed. High-severity fires were followed by increased sedimentary N stable isotope ratios (δ15N) and bulk density, and decreased C and N concentrations--reflecting forest floor destruction, terrestrial C and N losses, and erosion. Sustained low sediment C : N c. 20-50 yr post-fire indicates reduced terrestrial organic matter subsidies to the lake. Low sedimentary δ15N c. 50-70 yr post-fire, coincident with C and N recovery, suggests diminishing terrestrial N availability during stand development. The magnitude of post-fire changes generally scaled directly with inferred fire severity. Our results support modern studies of forest successional C and N accumulation and indicate pronounced, long-lasting biogeochemical impacts of wildfires in subalpine forests. However, even repeated high-severity fires over millennia probably did not deplete C or N stocks, because centuries between high-severity fires allowed for sufficient biomass recovery. © 2014 The Authors. New Phytologist © 2014 New Phytologist Trust.

  8. Biological and physical influences on the carbon isotope content of CO2 in a subalpine forest snowpack, Niwot Ridge, Colorado (United States)

    D. R. Bowling; W. J. Massman; S. M. Schaeffer; S. P. Burns; R. K. Monson; M. W. Williams


    Considerable research has recently been devoted to understanding biogeochemical processes under winter snow cover, leading to enhanced appreciation of the importance of many winter ecological processes. In this study, a comprehensive investigation of the stable carbon isotope composition (δ 13C) of CO2 within a high-elevation subalpine...

  9. Mycorrhiza-plant colonization patterns on a subalpine glacier forefront as a model system of primary succession (United States)

    Efren Cazares; James M. Trappe; Ari Jumpponen


    Lyman glacier in the North Cascades Mountains of Washington has a subalpine forefront characterized by a well-developed terminal moraine, inconspicuous successional moraines, fluting, and outwash. These deposits were depleted of symbiotic fungi when first exposed but colonized by them over time after exposure. Four major groups of plant species in this system are (1)...

  10. On mycorrhiza development of spruces and firs in damaged stands

    Energy Technology Data Exchange (ETDEWEB)

    Ritter, T.; Weber, G.; Kottke, I.; Oberwinkler, F.


    The authors studied the very fine roots of sick spruces and firs and established the following: 1. a surprising stability of mycorrhiza development, 2. differences in the dynamism of development and 3. modifications in the composition of the accompanying microfungi. The results suggest connections in the chain of causes of forest disease which have received little attention so far.

  11. Dust in FIR-bright ADF-S galaxies (United States)

    Małek, K.; Pollo, A.; Takeuchi, T. T.; Buat, V.; Burgarella, D.; Malkan, M.


    Multiwavelength Spectral Energy Distributions (SEDs) of far-infrared (FIR) galaxies detected in the AKARI South Ecliptic Poles Survey (ADF-S) allow to trace differences between [Ultra]-Luminous Infrared Galaxies ([U]LIRGS) and other types of star-forming galaxies (SF).

  12. Animal damage to young spruce and fir in Maine (United States)

    Barton M. Blum


    The loss of terminal buds on small balsam fir (Abies balsamea (L.) Mill.) and spruce (Picea spp.) trees because of nipping by mammals or birds has increased on the Penobscot Experimental Forest in recent years. The cut stem is smooth and slightly angled; there is no sign of tearing. Unnipped trees grew about 13 percent more than...

  13. Developmental decline in height growth in Douglas-fir. (United States)

    Barbara J. Bond; Nicole M. Czarnomski; Clifton Cooper; Michael E. Day; Michael S. Greenwood


    The characteristic decline in height growth that occurs over a tree's lifespan is often called "age-related decline." But is the reduction in height growth in aging trees a function of age or of size? We grafted shoot tips across different ages and sizes of Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) trees to determine whether...

  14. Biology of bats in Douglas-fir forests. (United States)

    Robin E. Christy; Stephen D. West


    Twelve species of bats occur in Douglas-fir forests of the Pacific Northwest, of which nine are known to roost in tree cavities, bark crevices, or foliage, and several are closely associated with old-growth forests. Thus bat populations may be detrimentally affected by forest management practices involving the removal of large, old trees and snags. We review the life...

  15. The biology of arboreal rodents in Douglas-fir forests. (United States)

    Andrew B. Carey


    Arboreal rodents in Douglas-fir forests west of the Cascade crest in Oregon and Washington include (listed in decreasing order of dependence on trees) red tree vole (Phenacomys longicaucfus), northern flying squirrel (Glaucomys sabrinus), Douglas' squirrel (Tamiasciurus douglasii), dusky-footed woodrat...

  16. Variation in seedling morphology of Turkish fir ( Abies ...

    African Journals Online (AJOL)

    In this study, the genetic variation of some seedling characteristics of Turkish fir was investigated. A total of 303 trees were selected from 17 plots and 10 seedlings from each tree were used. Fifteen morphological characteristics were determined, including root collar diameter, seedling height, total needle, bud length and ...

  17. Seed production of Douglas-fir increased by thinning. (United States)

    Donald L. Reukema


    In planning thinnings and final harvest cuttings for stands of young-growth Douglas-fir, foresters need reliable information on the capacity of young-growth stands to bear seed, on the periodicity of seed crops, and on the effects of thinning and other forest practices on seed production. One of the first studies designed to help provide this information was begun in...

  18. Effects of bear damage on Douglas-fir lumber recovery (United States)

    Eini C. Lowell; Dennis Dykstra; George McFadden


    Bear activily resulting in injury to Douglas-fir (Pseudotsuga menziesii [Mirb.] Franco) trees has been documented as early as the mid-1850s in the Pacific Northwest. The study reported in this article was designed to help managers decide whether the common practice of removing the damaged but potentially valuable butt section of the bottom log and...

  19. Power and Aging Characterization of Digital FIR Filters Architectures

    DEFF Research Database (Denmark)

    Calimera, Andrea; Liu, Wei; Macii, Enrico


    -variation, temperature and aging induced variations pose new challenges in the fabrication of the next generation of ICs. This work presents a detailed power and aging characterization of digital FIR filters in an industrial 45nm CMOS technology, and a design space exploration of different filter architectures...

  20. Hormonal control of second flushing in Douglas-fir shoots. (United States)

    Morris Cline; Mark Yoders; Dipti Desai; Constance Harrington; William. Carlson


    Spring-flushing, over-wintered buds of Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) produce new buds that may follow various developmental pathways. These include second flushing in early summer or dormancy before flushing during the following spring. Second flushing usually entails an initial release of apical dominance as some of the...

  1. Predicting Douglas-fir's response to a warming climate (United States)

    Andrea Watts; Sheel Bansal; Connie Harrington; Brad. St. Clair


    Douglas-fir is an iconic tree in the Pacific Northwest. Although individual trees may appear to be identical, genetic differences within each tree have resulted from adaptation to the local environment. These genetic differences over time have resulted in differences among populations that are important to the species' survival and growth in changing climates....

  2. Evaluating the Importance of Plant Functional Traits: the Subalpine and Alpine (United States)

    Sanchez, A.; Smith, W. K.


    Over the past several decades, researchers have attempted to characterize plant groups according to traits that are considered functional, i.e. contributing significantly to fitness. Due to the complexity of measuring fitness, the capability for photosynthetic carbon gain is often used as a proxy. Thus, this approach correlates structural differences to photosynthetic performance, especially those differences that are known to be associated with photosynthesis, are easily measured and inexpensive. At the often sharp boundary between the subalpine forest and alpine community (treeline ecotone), plant structural traits change dramatically, i.e. tall evergreen trees give way abruptly to low-stature shrubs, grasses, forbs, and herbs. Yet, the differences in functional traits, so abundant in the literature for a variety of species and communities, have not been compared contiguous communities such as the subalpine forest and alpine. Can differences in functional traits already identified in the literature also be used to characterize species of these two contrasting communities? Or are there other traits that are most functional and/or, possibly, unique to each community and not the most popular traits reported so far in the literature. Also, does the community structure itself help determine functional traits? For example, the top ten most frequently studied traits (145 total papers from approximately 63 different refereed journals) considered functional include the following (% of the 145 publications): specific leaf area or mass (SLA or SLM 39%), plant height (36%), leaf nitrogen content (34%), leaf size (19%), leaf area (16%), leaf photosynthetic performance (15%), leaf dry matter content (LDMC 15%), leaf mass per unit leaf area (LMA 15%), leaf thickness (15%), and seed mass (14%). In addition, another 120 traits were mentioned as functional, although all fell below a 14% citation rate. Particular focus was placed on this group due to the possibility that they might

  3. Evolutionary history and population genetics of fraser fir and intermediate fir, southern Appalachian endemic conifers imperiled by an exotic pest and climate change (United States)

    Kevin M. Potter; John Frampton; Sedley Josserand; C. Dana. Nelson


    Two Abies (true fir) taxa are endemic to high elevations of the Appalachian Mountains, where both are restricted to small populations and are imperiled by the same exotic insect. Fraser fir (Abies fraseri) exists in a handful of island-like populations on mountain ridges in the southern Appalachians of North Carolina, Tennessee and...

  4. A preliminary study of the deterioration of alder and Douglas-fir chips in outdoor piles. (United States)

    Ernest. Wright


    In the fall of 1952, E. E. Matson of the Pacific Northwest Forest and Range Experiment Station learned that the Fir-Tex Insulating Board Company bf St. Helens, Oregon was considering mixing alder with Douglas-fir chips for outside storage. Since alder heartwood i s more susceptible to decay than that of Douglas-fir, the question arose whether mixing the two might...

  5. Degrading Precision Arithmetics for Low-power FIR Implementation

    DEFF Research Database (Denmark)

    Albicocco, Pietro; Cardarilli, Gian Carlo; Nannarelli, Alberto


    In this paper a review of different techniques used to implement highly optimized DSP systems is presented. The case of study is the implementation of parallel FIR filters aimed to applications characterized by high speed and high selectivity in frequency where at the same time low power dissipat......In this paper a review of different techniques used to implement highly optimized DSP systems is presented. The case of study is the implementation of parallel FIR filters aimed to applications characterized by high speed and high selectivity in frequency where at the same time low power...... on selective bit freezing, DPA-II, based on VDD voltage scaling, and DPA-III, based on power gating. Some theoreticaVsimuiative analysis of the introduced arithmetic errors and some implementation results are shown. A discussion on the suitability of these methodologies on standard cell technologies and FPGAs...

  6. Noniterative Design of 2-Channel FIR Orthogonal Filters

    Directory of Open Access Journals (Sweden)

    Jiménez M Elena Domínguez


    Full Text Available This paper addresses the problem of obtaining an explicit expression of all real FIR paraunitary filters. In this work, we present a general parameterization of 2-channel FIR orthogonal filters. Unlike other approaches which make use of a lattice structure, we show that our technique designs any orthogonal filter directly, with no need of iteration procedures. Moreover, in order to design an -tap 2-channel paraunitary filterbank, it suffices to choose independent parameters, and introduce them in a simple expression which provides the filter coefficients directly. Some examples illustrate how this new approach can be used for designing filters with certain desired properties. Further conditions can be eventually imposed on the parameters so as to design filters for specific applications.

  7. A compact CO bipolar outflow from NGC 2024 FIR6

    Energy Technology Data Exchange (ETDEWEB)

    Richer, J.S. (Mullard Radio Astronomy Observatory, Cambridge (UK). Cavendish Lab.)


    The discovery is reported of a bipolar outflow source in the NGC 2024 cloud core, which appears to be associated with both the 1300-{mu}m continuum source FIR6, and a water maser. This flow is compact (size {approx} 0.05 pc), displays bright, wide line-wings (total width {approx} 125 km s{sup -1}) and has a mechanical luminosity L{sub CO} {approx} 15 times the solar luminosity. These properties suggest that FIR6 is a young stellar object of high mass: from the correlation of mechanical luminosity with bolometric luminosity, the source luminosity is > {approx} 10{sup 3.5} times the solar luminosity. The short dynamical timescale of this flow suggests that the system is very young, but the high inferred luminosity implies that the object has evolved beyond the stage of isothermal protostellar contraction. (author).

  8. Cyclotron Resonance Gain for FIR and THz Radiation in Graphene

    CERN Document Server

    Cole, Nightvid


    A cyclotron resonance maser source using low-effective-mass conduction electrons in graphene, if successful, would allow for generation of Far Infrared (FIR) and Terahertz (THz) radiation without requiring magnetic fields running into the tens of Tesla. In order to investigate this possibility, we consider a device in which electrons are effectively injected via pumping from the valence band to the conduction band using an infrared (IR) laser source, subsequently gyrate in a magnetic field applied perpendicular to the plane of the graphene, and give rise to gain for a FIR/THz wave crossing the plane of the graphene. A set of integral expressions is derived by assuming that the non-radiative energy loss processes of the electrons can be adequately represented by a damping force proportional and antiparallel to their momentum. Minimal gain may occur at very short electron damping times of hundreds of femtoseconds.

  9. Dowel-nut connection in Douglas-fir peeler cores (United States)

    Ronald W. Wolfe; John R. King; Agron. Gjinolli

    As part of an effort to encourage more efficient use of small-diameter timber, the Forest Products Laboratory cooperated with Geiger Engineers in a study of the structural properties of Douglas-fir peeler cores and the efficacy of a bdowel-nutc connection detail for application in the design of a space frame roof system. A 44.5-mm- (1.75-in.-) diameter dowel-nut...

  10. Predicting fir trees stem diameters using Artificial Neural Network ...

    African Journals Online (AJOL)

    The aim of this paper is to examine the applicability of Artificial Neural Network models (ANNs), in the prediction of fir trees stem over bark diameters at 5.3, 9.3, 13.3, 17.3, 21.3, 25.3, 29.3 and 33.3 meters above ground. The values of these diameters are necessary for an efficient estimation of a single tree volume using the ...

  11. Douglas-fir displays a range of growth responses to ... (United States)

    Douglas-fir (Pseudotsuga menziesii var. menziesii (Mirb.) Franco) growth in the Pacific Northwest is affected by climatic, edaphic factors and Swiss needle cast (SNC) disease. We examine Douglas-fir growth responses to temperature, dewpoint deficit (DPD), soil moisture, and SNC using time series intervention analysis of intra-annual tree-ring width data collected at nine forest stands in western Oregon, USA. The effects of temperature and SNC were similar in importance on tree growth at all sites. Previous-year DPD during the annual drought period was a key factor limiting growth regionally. Winter temperature was more important at high elevation cool sites, whereas summer temperature was more important at warm and dry sites. Growth rate increased with summer temperature to an optimum (Topt) then decreased at higher temperatures. At drier sites, temperature and water affected growth interactively such that Topt decreased with decreasing summer soil moisture. With climate change, growth rates increased at high elevation sites and declined at mid-elevation inland sites since ~1990. Growth response to climate is masked by SNC regionally. We conclude that as temperature rises and precipitation patterns shift towards wetter winters and drier summers, Douglas-fir will experience greater temperature and water stress and an increase in severity of SNC. By the end of the 21st century, climate models predict hotter, drier summers and warmer, wetter winters in the Pac

  12. Comparison of remote sensing and plant trait-based modelling to predict ecosystem services in subalpine grasslands

    Czech Academy of Sciences Publication Activity Database

    Homolová, Lucie; Schaepman, M. E.; Lamarque, L.; Clevers, J.G.P.W.; de Bello, Francesco; Thuiller, W.; Lavorel, S.


    Roč. 5, č. 8 (2014), č. článku 100. ISSN 2150-8925 Institutional support: RVO:67179843 ; RVO:67985939 Keywords : land-use change * leaf chlorophyll content * imaging spectroscopy * water-content * aviris data * spectral reflectance * hyperspectral data * species richness * area index * vegetation * aisa * biomass * ecosystem properties * ecosystem services * linear regression * remote sensing * spatial heterogeneity * subalpine grasslands Subject RIV: EH - Ecology, Behaviour; EF - Botanics (BU-J) Impact factor: 2.255, year: 2014

  13. The Effect of Re-Planting Trees on Soil Microbial Communities in a Wildfire-Induced Subalpine Grassland


    Ed-Haun Chang; Guanglong Tian; Chih-Yu Chiu


    Wildfire often causes tremendous changes in ecosystems, particularly in subalpine and alpine areas, which are vulnerable due to severe climate conditions such as cold temperature and strong wind. This study aimed to clarify the effect of tree re-planting on ecosystem services such as the soil microbial community after several decades. We compared the re-planted forest and grassland with the mature forest as a reference in terms of soil microbial biomass C and N (Cmic and Nmic), enzyme activit...

  14. Effects of long-term population fluctuations of a top predator on invertebrate communities in subalpine ponds in Colorado


    Wissinger, S.; Whiteman, H.; Denoël, Mathieu; Greig, H.; Butkas, K.


    Experimental and comparative data from subalpine ponds with and without tiger salamanders (Ambystoma tigrinum nebulosum) suggest that this species is a keystone predator on benthic and planktonic prey communities. At our study site in central Colorado, the population size of salamanders has fluctuated cyclically over the past 20 years from fewer than 100 to over 5000 individuals. Here we present long-term benthic data that reveal taxon- and habitat-specific correlations with fluctuations in s...

  15. The Effect of Re-Planting Trees on Soil Microbial Communities in a Wildfire-Induced Subalpine Grassland

    Directory of Open Access Journals (Sweden)

    Ed-Haun Chang


    Full Text Available Wildfire often causes tremendous changes in ecosystems, particularly in subalpine and alpine areas, which are vulnerable due to severe climate conditions such as cold temperature and strong wind. This study aimed to clarify the effect of tree re-planting on ecosystem services such as the soil microbial community after several decades. We compared the re-planted forest and grassland with the mature forest as a reference in terms of soil microbial biomass C and N (Cmic and Nmic, enzyme activities, phospholipid fatty acids (PLFA composition, and denaturing gradient gel electrophoresis (DGGE. The Cmic and Nmic did not differ among the grassland, re-planted forest and mature forest soil; however, ratios of Cmic/Corg and Nmic/Ntot decreased from the grassland to re-planted forest and mature forest soil. The total PLFAs and those attributed to bacteria and Gram-positive and Gram-negative bacteria did not differ between the re-planted forest and grassland soil. Principle component analysis of the PLFA content separated the grassland from re-planted forest and mature forest soil. Similarly, DGGE analysis revealed changes in both bacterial and fungal community structures with changes in vegetation. Our results suggest that the microbial community structure changes with the re-planting of trees after a fire event in this subalpine area. Recovery of the soil microbial community to the original state in a fire-damaged site in a subalpine area may require decades, even under a re-planted forest.

  16. Responses Of Subalpine Conifer Seedling Germination And Survival To Soil Microclimate In The Alpine Treeline Warming Experiment (United States)

    Castanha, C.; Moyes, A. B.; Torn, M. S.; Germino, M. J.; Kueppers, L. M.


    At Niwot Ridge, Colorado, we used common gardens and climate manipulations to investigate potential subalpine tree species range shifts due to climate change. In Fall 2009 we harvested seed from local populations of limber pine and Englemann spruce, which we sowed in 3 experimental sites spanning an elevation gradient from lower subalpine forest (3080m asl), to the upper subalpine treeline ecotone (3400m asl), to the alpine tundra (3550m asl). In October we turned on overhead infrared heaters designed to increase growing season surface soil temperature by 4-5°C, and following snowmelt in 2010 we crossed this heating treatment with manual watering, adding 3mm of water each week. Here we report on the species, site, and treatment effects on seedling emergence and survival as mediated by snowmelt date, soil temperature, and soil moisture. Depending on the site and plot, heating accelerated germination by 1 to 4 weeks. Germination degree days (heat accumulation required for seed germination) were greater for pine than for spruce and greater in drier plots. Seedling survival was explained by date of emergence, with older seedlings more likely to survive the season. Survival was also explained by drought degree days -- the number of days below critical soil moisture thresholds compounded by high temperature -- with lower thresholds for spruce than for pine. Our preliminary results indicate that a warmer environment will stimulate germination for both species, but that, survival - especially for spruce - will be critically modulated by summer soil moisture.

  17. Seasonal fluctuation in concentrations and content of sugar, starch and nutrient in nordmanns fir (Abies nordmannia) Christmas tree

    DEFF Research Database (Denmark)

    Pedersen, Lars Bo; Christensen, Claus Jerram

    Seasonal fluctuation in concentrations and content of sugar, starch and nutrient in nordmanns fir (Abies nordmannia) Christmas tree......Seasonal fluctuation in concentrations and content of sugar, starch and nutrient in nordmanns fir (Abies nordmannia) Christmas tree...

  18. Growth Response of Silver Fir and Bosnian Pine from Kosovo

    Directory of Open Access Journals (Sweden)

    Elvin Toromani


    Full Text Available Background and Purpose: This paper explore the growth-climate relationships in total ring width chronologies of silver fir (Abies alba Mill. and Bosnian pine (Pinus heldreichii Christ. The objective of this study is to quantify the climate influence on radial growth of both species. The relationships between climate and ring widths were analyzed using extreme growing years (called pointer years, simple correlations and response functions analysis (bootstrapped coefficients. The objectives of this study were: (1 to define the pattern of climatic response of each species, (2 to highlight the influence of local ecological conditions on tree's growth, and (3 to compare the response of silver fir and Bosnian pine to climate. Responses of total ring width to climate were estimated by establishing the mean relationship between growth and climate through simple correlations analysis and bootstrapped response functions. The response to climatic variability was also assessed by analyzing pointer years which correspond to abrupt changes in growth pattern and revealing the tree-growth response to extreme climatic events. For the period 1908-2008 the mean sensitivity (MS of total ring width chronology for Bosnian pine (0.209 was higher than silver fir (0.169 suggesting that Bosnian pine is more sensitive to climate (pointer years were more frequent in ring width chronology of Bosnian pine than in silver fir ring width chronology. The high values of first-order autocorrelations for Bosnian pine (0.674 indicated a strong dependence of current growth on the previous year’s growth. Pointer years analysis underlined the high sensitivity to spring temperatures and precipitation for both species. Radial growth for both species depends strongly on spring climate variables (temperatures and precipitation which play a significant role particularly for earlywood production. Material and Methods: We selected 12 silver fir trees and 15 Bosnian pine trees and took two 5

  19. Ten-year variability in fluxes, meteorology, and environmental conditions at a Colorado subalpine forest site (United States)

    Burns, S. P.; Turnipseed, A.; Bowling, D. R.; Hu, J.; Monson, R. K.


    Changing meteorological and environmental conditions affect fluxes; model analysis has shown that environmental variability directly accounts for about half the interannual variability in net ecosystem exchange (NEE) of CO2 whereas the other 50% is due to biotic responses to these changing variables (Richardson et al. 2007). In our study, ten years (1998-2008) of turbulent flux measurements of heat, water vapor, and CO2 at the Niwot Ridge AmeriFlux site (Monson et al. 2002) are examined with respect to meteorological conditions (atmospheric temperature, stability, precipitation, and cloudiness) as well as changes in environmental conditions, such as snow depth and soil moisture. The typical yearly cycle and an estimate of the magnitude of year-to-year variability in the diurnal fluxes and other variables for a high-elevation subalpine forest ecosystem are presented. Wintertime ecosystem respiration has an average 30-min NEE of 0.62 μmol m-2 s-1 with an interannual range between 0.5-1 μmol m-2 s-1. Uptake of CO2 in late summer has an average NEE of -0.71 μmol m-2 s-1 with an interannual range between -0.1 to -1.5 μmol m-2 s-1. Previous studies at the Niwot Ridge AmeriFlux site have described the importance of photosynthetic photon flux density (PPFD) (Monson et al. 2002) and also growing season length (Hu et al. 2009) on NEE. Water isotope ratios analyzed by Hu et al. (2009) have shown that trees at the site primarily rely on water from snowmelt to sustain them throughout the summer; combining this result with the SIPNET model, Hu et al. conclude that there is a limited connection between summer precipitation and the cumulative annual gross primary production (GPP). We have tested this conclusion more explicitly by examining the response of NEE to specific precipitation events and the effect of extended dry periods on the diel cycle of the fluxes, CO2 mole fraction, sap flow, and other meteorological and soil variables. Additionally, we examine the connection

  20. Response of subalpine grasslands communities to clear-cut and prescribed burnt to control shrub encroachment in Pyrenees (United States)

    Alados, Concepción L.; Gartzia, Maite; Nuche, Paloma; Saiz, Hugo; Pueyo, Yolanda


    Anthropogenic activities have modified vegetation in subalpine belts for long time, lowering treeline ecotone and influencing landscape mainly through grazing and fire. During the last decades the abandonment of traditional land use practices and global warming are contributing largely to the colonization of woody species in subalpine grasslands causing irreversible changes in ecosystem functioning. To prevent those changes a variety of management strategies are carried out to stop the expansion of the highly encroaching shrubs, which require the use of fire and/or clear-cutting, particularly for the conservation of grasslands in subhumid high productive ecosystems. However, it is still poorly understood how different management strategies affect the recovery of subalpine grasslands. Using a field experiment we tested the impact of management treatments on soil properties and vegetation characteristics, including species richness, community structure, interspecies interaction, and complexity of network association. Vegetation was monitored during four years in eight stands (two stands per treatment) where the vegetation was removed by prescribed fire (Burnt treatment), or by mechanical removal (Clear-cut treatment). Two undisturbed E. horridum stands were used as a control (C-Erizón) and two grassland communities regularly grazed (C-Grass) were used as a control for subalpine grassland. Soils nutrients declined in Burnt treatment 3 years after fire, but not differences between Clear-cut and C-Erizón were observed. Species richness and diversity were larger in C-Grass and lower in C-Erizón. Burnt and Clear-cut treatments increased species diversity and richness gradually after 4 year treatment. The proportion of legume forbs, grasses and non-legume forbs did not reach the levels of C-Grass after 4 years of Clear-cut or Burnt treatments. Shrubs and sub-shrubs increased faster after 4 years of burning than after 4 years of clearing, although they did not reach the

  1. Assessing the Climate Sensitivity of Cold Content and Snowmelt in Seasonal Alpine and Subalpine Snowpacks (United States)

    Jennings, K. S.; Molotch, N. P.


    In cold, high-elevation sites, snowpack cold content acts as a buffer against climate warming by resisting snowmelt during periods of positive energy fluxes. To test the climate sensitivity of cold content and snowmelt, we employed the physical SNOWPACK snow model, forced with a 23-year, hourly, quality-controlled, gap-filled meteorological dataset from the Niwot Ridge Long Term Ecological Research (LTER) site in the Front Range mountains of Colorado. SNOWPACK was run at two points with seasonal snowpacks within the LTER, one in the alpine (3528 m) and one in the subalpine (3022 m). Model output was validated using snow water equivalent (SWE), snowpack temperature, and cold content data from snow pits dug near the met stations and automated SWE data from nearby SNOTEL snow pillows. Cold content accumulates primarily through additions of new snow, while negative energy fluxes—cooling through longwave emission and sublimation—play a lesser role, particularly in the deeper snowpack of the alpine. On average, the snowpack energy balance becomes positive on April 1 in the alpine and March 8 in the subalpine. Peak SWE occurs after these dates and its timing is primarily determined by the amount of precipitation received after peak cold content, with persistent snowfall delaying the main snowmelt pulse. Years with lower cold content, due to reduced precipitation and/or increased air temperature, experience an earlier positive energy balance with more melt events occurring before the date of peak SWE, which has implications for soil moisture, streamflow volume and timing, water uptake by vegetation, and microbial respiration. Synthetic warming experiments show significant cold content reductions and increased late-winter/early-spring melt as positive energy balances occur earlier in the snow season (a forward shift between 5.1 and 21.0 days per °C of warming). These results indicate cold, high-elevation sites, which are critical for water resources in the western

  2. [Dynamics of species diversity in artificial restoration process of subalpine coniferous forest]. (United States)

    Wu, Yan; Liu, Qing; He, Hai; Lin, Bo


    Through plot investigation and by adopting the concept of space as a substitute for time, the developments of species diversity of trees, shrubs and herbaceous plants in subalpine coniferous plantations at different restoration stages were studied, and the correlation coefficients of species in each layer were discussed. The results indicated that in the restoration process, the species richness, diversity and evenness in subalpine coniferous plantations were gradually increased in a fluctuating way. The restoration process of Picea asperata plantations showed a tendency of development that in favor of resuming species diversity. The indices of species richness (species number and Margalef index) and species diversity (Shannon-Wiener index and Simpson index) of trees increased rapidly from the early stages of plantation establishment to the stage of canopy closing (about 30 yr of stand age) and then presented a tendency of decrease with some slight fluctuations, while the index of species evenness showed a periodical rising trend. For the shrub layer, the indices of species richness (Simpson index and Macintosh index) gradually increased with increasing restoration years, whereas the indices of species diversity (Shannon-Wiener index) decreased in the early stages, sharply increased during the stages of canopy closing, and then slowly decreased, which exhibited a tendency of high-->low-->high. In the layer of herbaceous plants, the indices of species richness (Margalef index and number of species) and species diversity (Simpson index, Macintosh index and Shannon-Wiener index) presented a trend of decrease in the early stages of plantations establishment to canopy closing and increased later on. During this process, herbaceous species and their life forms changed greatly, with shade tolerant species gradually substituting the intolerant species. Among the plantations of different stand ages, the average correlation coefficients of trees, shrubs and herbaceous

  3. Early development of matched planted and naturally regenerated Douglas-fir stands after slash burning in the Cascade Range. (United States)

    R.E. Miller; R.E. Bigley; S. Webster


    We compared matched planted and naturally regenerated plots in 35- to 38- year-old Douglas-fir (Pseudotsuga menziesii var. menziesii) stands at seven locations in western Washington and Oregon. Total number of live stems is similar, but stands planted to Douglas fir average 26 more live stemslac of Douglas-fir and 39 fewer...

  4. A forest health inventory assessment of red fir (Abies magnifica) in upper montane California (United States)

    Leif Mortenson; Andrew N. Gray; David C. Shaw


    We investigated the forest health of red fir (Abies magnifica) and how it compared with commonly-associated species Jeffrey pine (Pinus jeffreyi), lodgepole pine (Pinus contorta) and white fir (Abies concolor) in the upper montane forests of California. We evaluated tree mortality rates...

  5. A SNP resource for douglas-fir: de novo transcriptome assembly and SNP detection and validation (United States)

    Glenn R. Howe; Jianbin Yu; Brian Knaus; Richard Cronn; Scott Kolpak; Peter Dolan; W. Walter Lorenz; Jeffrey F.D. Dean


    Douglas-fir (Pseudotsuga menziesii), one of the most economically and ecologically important tree species in the world, also has one of the largest tree breeding programs. Although the coastal and interior varieties of Douglas-fir (vars. menziesii and glauco) are native to North America, the coastal variety is...

  6. Mammoth lakes revisited—50 years after a Douglas-fir tussock moth outbreak. (United States)

    Boyd E. Wickman; G. Lynn Starr


    For five decades after an outbreak of Douglas-fir tussock moth (Orgyia pseudotsugata (McDunnough)), radial growth of defoliated white fir trees (Abies concolor (Gord. & Glend.) Lindl.), was significantly greater than that of nondefoliated host trees nearby. The increased growth probably was due to the thinning effect of...

  7. Belowground competition from overstory trees influences Douglas-fir sapling morphology in thinned stands (United States)

    Warren D. Devine; Timothy B. Harrington


    We evaluated effects of belowground competition on morphology of naturally established coast Douglas-fir (Pseudotsuga menziesii var. menziesii (Mirb.) Franco) saplings in 60- to 80-year-old thinned Douglas-fir stands in southwestern Washington. We separately quantified belowground competition from overstory and understory sources...

  8. Population dynamics of dwarf mistletoe on young true firs in the central Sierra Nevada, California (United States)

    Robert E Scharpf; J. R. Jr. Parmeter


    Young red firs (Abies magnifica A. Murr.) and white firs (A. concolor [Gord. & Glend.] Lindl. ex Hildebr.) on the Stanislaus National Forest, California, were inoculated with seeds of dwarf mistletoe (Arceuthobium abietinum) for 5 successive years. Only 3 to 4 percent of about 7000 seeds placed on branches...

  9. White fir stands killed by tussock moth...70-mm. color photography aids detection (United States)

    Steven L. Wert; Boyd E. Wickman


    The use of large-scale 70 mm. aerial photography proved to be an effective technique for detecting trees in white fir stands killed by Douglas-fir tussock moth in northeastern California. Correlations between ground and photo estimates of dead trees were high. But correlations between such estimates of lesser degrees of tree damage--thin tops and topkill--were much...

  10. Projected future suitable habitat and productivity of Douglas-fir in western North America (United States)

    Aaron R. Weiskittel; Nicholas L. Crookston; Gerald E. Rehfeldt


    Douglas-fir (Pseudotsuga menziesii [Mirb.] Franco) is one of the most common and commercially important species in western North America. The species can occupy a range of habitats, is long-lived (up to 500 years), and highly productive. However, the future of Douglas-fir in western North America is highly uncertain due to the expected changes in climate conditions....

  11. An audio FIR-DAC in a BCD process for high power Class-D amplifiers

    NARCIS (Netherlands)

    Doorn, T.S.; van Tuijl, Adrianus Johannes Maria; Schinkel, Daniel; Annema, Anne J.; Berkhout, M.; Berkhout, M.; Nauta, Bram

    A 322 coefficient semi-digital FIR-DAC using a 1-bit PWM input signal was designed and implemented in a high voltage, audio power bipolar CMOS DMOS (BCD) process. This facilitates digital input signals for an analog class-D amplifier in BCD. The FIR-DAC performance depends on the ISI-resistant

  12. Dwarf Mistletoe on Red Fir . . . infection and control in understory stands (United States)

    Robert F. Scharpf


    Height and age of understory red fir (Abies magnifica A. Murr.) were related to dwarf mistletoe (Arceuthobiilm campylopodum f. abietinum) infection from the surrounding overstory red fir on four National Forests in California. Percentage of trees infected and intensity of infection increased significantly as height of understory...

  13. Conversion of SPORL pretreated Douglas fir forest residues into microbial lipids with oleaginous yeasts (United States)

    Bruce S. Dien; Junyong Zhu; Patricia J. Slininger; Cletus P. Kurtzman; Bryan R. Moser; Patricia J. O' Bryan; Roland Gleisner; Michael A. Cotta


    Douglas fir is the dominant commercial tree grown in the United States. In this study Douglas fir residue was converted to single cell oils (SCO) using oleaginous yeasts. Monosaccharides were extracted from the woody biomass by pretreating with sulfite and dilute sulfuric acid (SPORL process) and hydrolyzing using commercial cellulases. A new SPORL process that uses pH...

  14. Documentation of the Douglas-fir tussock moth outbreak-population model. (United States)

    J.J. Colbert; W. Scott Overton; Curtis. White


    Documentation of three model versions: the Douglas-fir tussock moth population-branch model on (1) daily temporal resolution, (2) instart temporal resolution, and (3) the Douglas-fir tussock moth stand-outbreak model; the hierarchical framework and the conceptual paradigm used are described. The coupling of the model with a normal-stand model is discussed. The modeling...

  15. Multi-decadal establishment for single-cohort Douglas-fir forests (United States)

    James A. Freund; Jerry F. Franklin; Andrew J. Larson; James A. Lutz


    The rate at which trees regenerate following stand-replacing wildfire is an important but poorly understood process in the multi-century development of Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) and western hemlock (Tsuga heterophylla (Raf.) Sarg.) forests. Temporal patterns of Douglas-fir establishment reconstructed from old-growth forests (>450 year) have...

  16. Predictions of fire behavior and resistance to control: for use with photo series for the Douglas fir-hemlock type and the coastal Douglas-fir-hardwood type. (United States)

    David V. SANDBERG; Franklin R. Ward


    This publication presents tables on the behavior of fire and the resistance of fuels to control. The information is to be used with the photos in the publication, "Photo Series for Quantifying Forest Residues in the Coastal Douglas-fir—Hemlock Type, Coastal Douglas-fir—Hardwood Type" (Maxwell, Wayne G.; Ward, Franklin R. 1976. Gen. Tech. Rep. PNW-GTR-051....

  17. Analysis of the growth characteristics of a 450-year-old silver fir tree

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    Pantić Damjan


    Full Text Available The growth characteristics of silver fir are of high importance for selection forest management, and for the current aims laid out in Serbia’s forest management focused on increasing the share of silver firs in Serbia’s growing stock. With the objective of increasing the understanding of the growth characteristics of silver fir, the growth of two silver fir trees felled during forest site production research on Mt. Goč, located in Central Serbia, have been analyzed. Both trees showed significant differences in their growth dynamics over long periods as results of micro-site and micro-stand effects (primarily ambient light regime. The common growth characteristic of the two trees, a 450-year-old tree as the main study object (labeled Tree A and a 270-year-old Tree B is a long stagnation stage. For Tree A the latent phase, with small interruptions, lasted 410 years; one phase lasted 330 years in continuity, which is the longest period of silver fir stagnation recorded in Europe. Tree B showed a long-lasting stagnation stage that lasted 170 years. The long stagnation stage of Tree A, characterized by an average diameter increment of 1.4 mm/year (average growth ring width of 0.7 mm and an average height increment of 0.08 m/year, shows the extraordinary silver fir capacity for physiological survival in complete shade. This study adds to the existing knowledge of the shade tolerance of the silver fir. Therefore, the silver fir belongs to the group of extremely shade-tolerant tree species. This characteristic makes silver fir an irreplaceable tree species in the selection forest structure. It offers a wide range of silvicultural flexibility in the management of these forests, and is applicable to silver fir selection Serbia’s forests. [Projekat Ministarstva nauke Republike Srbije, br. EVBR 37008: Sustainable management of total forest potentials in the Republic of Serbia

  18. Imago feeding preference of the black fir sawyer Monochamus urussovi Fisch. (Coleoptera: Cerambycidae

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    Grodnitsky Dmitry


    Full Text Available Mass outbreaks of the black fir sawyer M. urussovi regularly occur in forest stands dominated by Siberian fir Abies sibirica; outbreaks have been observed only on fir and exclusively in Siberia, although the beetle is also found in the vast territory including Finland, Poland, Ukraine, Kazakhstan, China, Russian Far East and Japan. A closely related common species M. sutor has never produced outbreaks; its larvae have been registered only in Scots pine. Feeding preferences of the two species were compared under experimental conditions; total numbers of tested specimens were 132 (M. urussovi and 40 (M. sutor. M. sutor beetles were caught in nature, M. urussovi pupas and young adults were collected from fir trunks; pupas were reared to beetles. Each beetle was placed into a covered plastic enclosure for 24 hours and offered cut shoots of fir, spruce, Siberian pine, Scots pine, larch, birch and willow; all cuts were of the same length and diameter, needles and leaves had been previously removed. Beetles of the two species showed different feeding preference. M. sutor imagoes completely ignored fir shoots, definitely preferring Scots pine. M. urussovi feeding behavior was different: beetles did consume bark, bast and cambium of fir. However, they were mainly fed on Scots pine shoots. Feeding preference given to Scots pine by beetles grown in fir trunks can be explained by the hypothesis that this species originated on pine. Mass outbreaks of M. urussovi in Siberian fir-dominated forest ecosystems occur, because these plant communities are historically young (evidently much less than 10 000 years and yet unbalanced, transitional sort of ecosystems, so fir did not have enough evolutionary time to get adapted to injure by the sawyer and Ophiostoma fungi carried by beetles.

  19. Protocol for fir tree sampling for provenance studies (United States)

    Meisel, Thomas; Bandoniene, Donata; Zettl, Daniela


    Isotopic (stable and radiogenic) as well as trace element fingerprinting methods used for tracing the geographical origin, rely on databases, that need to contain data sets representative of the measurands of the individual samples for a specific geographic entity. Through this work, we want to assess different sampling strategies for obtaining representative sample of fir trees (Abies sp.). Motivation for this work is the protection of the local Austrian Christmas tree market from wrongly tagged trees of non-Austrian origin. In particular, we studied three typical Christmas trees the most common species sold as Christmas tree, namely Abies nordmanniana (Nordmann Fir), from the same locality in lower Austria. For the initial tests we applied the elemental fingerprinting method, to study the suitability of the different parts of the tree applying ICP-MS analysis after complete acid digestion in a high pressure asher system (HPA-S).Needle samples from each year of life of the tree and stem wood from three different heights were analyzed for their trace element content to prove the repeatability and to find the best sampling protocol. For the analysis of the needles, the natural wax coating had to be removed in order to get reproducible results. For the analysis of stem wood only the bark was removed. As expected the data of all three trees allowed the differentiation of the individual needle ages, but interestingly enough also between the three sampling heights of the needs. Both needles and wood proved to be suitable for successful fingerprinting, but importantly, provided that sample of the same type and ages are compared. The same samples for the three trees will also be used for isotopic analysis studies to better understand the influence of age and sampling height on the representativeness of fir tree samples. Based on elemental fingerprinting alone, a successful discrimination between local (Austrian) and foreign (Danish, Irish) Christmas trees was possible.

  20. Limnological research in the deep southern subalpine lakes: synthesis, directions and perspectives

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    Nico Salmaso


    Full Text Available Based on a selection of 230 papers published during the last 15 years in international journals, the present work aims at evaluating the state of the art of limnological research in the deep southern subalpine lakes (DSL: Garda, Iseo, Como, Lugano and Maggiore. Historically, most of the limnological research was fostered by the need to find solutions to the problems connected with eutrophication and pollution. Many data are available on the thermal structure, algal nutrient concentrations and phytoplankton of the DSL, while other topics still remain more or less constrained to single lakes. Apart from this geographical bias, a number of aspects emerged from this synoptic view. Limnological research is still linked to the concept of scientific monitoring, while experimental studies and modelling are confined to specific niches; the integration of different disciplines is held back by the division of studies on different compartments; integration of studies and synoptic analyses at a macro regional scale have been carried out only for specific research areas. The DSL are increasingly threatened by new pressures (climatic change, excessive proliferation of toxic cyanobacteria, introduction of new species and new micropollutants and by the interactions among these new and old stressors. In this rapidly changing situation, the paper emphasises the need to define criteria to be used to distinguish research able to produce relevant results and predictive models, which are essential elements for an efficient management of water resources.

  1. Patch-Scale Effects of Equine Disturbance on Arthropod Assemblages and Vegetation Structure in Subalpine Wetlands (United States)

    Holmquist, Jeffrey G.; Schmidt-Gengenbach, Jutta; Ballenger, Elizabeth A.


    Assessments of vertebrate disturbance to plant and animal assemblages often contrast grazed versus ungrazed meadows or other larger areas of usage, and this approach can be powerful. Random sampling of such habitats carries the potential, however, for smaller, more intensely affected patches to be missed and for other responses that are only revealed at smaller scales to also escape detection. We instead sampled arthropod assemblages and vegetation structure at the patch scale (400-900 m2 patches) within subalpine wet meadows of Yosemite National Park (USA), with the goal of determining if there were fine-scale differences in magnitude and directionality of response at three levels of grazing intensity. Effects were both stronger and more nuanced than effects evidenced by previous random sampling of paired grazed and ungrazed meadows: (a) greater negative effects on vegetation structure and fauna in heavily used patches, but (b) some positive effects on fauna in lightly grazed patches, suggested by trends for mean richness and total and population abundances. Although assessment of disturbance at either patch or landscape scales should be appropriate, depending on the management question at hand, our patch-scale work demonstrated that there can be strong local effects on the ecology of these wetlands that may not be detected by comparing larger scale habitats.

  2. A molecular investigation of soil organic carbon composition across a subalpine catchment (United States)

    Hsu, Hsiao-Tieh; Lawrence, Corey R.; Winnick, Matthew J.; Bargar, John R.; Maher, Katharine


    The dynamics of soil organic carbon (SOC) storage and turnover are a critical component of the global carbon cycle. Mechanistic models seeking to represent these complex dynamics require detailed SOC compositions, which are currently difficult to characterize quantitatively. Here, we address this challenge by using a novel approach that combines Fourier transform infrared spectroscopy (FT-IR) and bulk carbon X-ray absorption spectroscopy (XAS) to determine the abundance of SOC functional groups, using elemental analysis (EA) to constrain the total amount of SOC. We used this SOC functional group abundance (SOC-fga) method to compare variability in SOC compositions as a function of depth across a subalpine watershed (East River, Colorado, USA) and found a large degree of variability in SOC functional group abundances between sites at different elevations. Soils at a lower elevation are predominantly composed of polysaccharides, while soils at a higher elevation have more substantial portions of carbonyl, phenolic, or aromatic carbon. We discuss the potential drivers of differences in SOC composition between these sites, including vegetation inputs, internal processing and losses, and elevation-driven environmental factors. Although numerical models would facilitate the understanding and evaluation of the observed SOC distributions, quantitative and meaningful measurements of SOC molecular compositions are required to guide such models. Comparison among commonly used characterization techniques on shared reference materials is a critical next step for advancing our understanding of the complex processes controlling SOC compositions.

  3. Phytoplankton association patterns in the deep southern subalpine lakes (Part 2

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    Nico Salmaso


    Full Text Available The three following papers constitute the second, and final, part of a series of papers dedicated to the phytoplankton of the deep subalpine Italian lakes (DSL. The first part, comprising an introduction and three papers on lakes Garda, Como and Maggiore respectively, was published in the volume 61 (1 of this journal (J. Limnol., 61. The research, carried out for three years (two years in L.Como in the period 1997-2000, was a concerted effort by investigators of five Insitutes in Italy and Switzerland. It was generated by the awareness that, despite the large number of papers existing on the phytoplankton of the single DSL, those of a comprehensive nature are very few, and by the perceived interest of a comparative investigation on the phytoplankton of the large lakes in the Insubrian district in the light of the recent progress on the ecology of the freshwater algae. So, in the final paper following those on lakes Iseo and Lugano an effort is made to compare and characterize the different species assemblages in the background of geographical and morphological characteristics, and of recent changes in trophic gradients. Despite often important differences in these respects, and observed various community responses, a common pool of species seems to be identifiable in the district.

  4. Development of partial rock veneers by root throw in a subalpine setting (United States)

    Osterkamp, W.R.; Toy, T.J.; Lenart, M.T.


    Rock veneers stabilize hillslope surfaces, occur especially in areas of immature soil, and form through a variety of process sets that includes root throw. Near Westcliffe, Colorado, USA, data were collected from a 20 ?? 500 m transect on the east slope of the Sangre de Cristo Mountains. Ages of pit/mound complexes with rock fragments exposed at the surface by root throw ranged from recent (freshly toppled tree) to unknown (complete tree decay). Calculations based on dimensions of the pit/mound complexes, estimated time of free topppling, sizes of exposed rock fragments, and percentage rock covers at pit/mound complexes, as well as within the transect area, indicate that recent rates of root throw have resulted in only partial rock veneering since late Pleistocene deglaciation. Weathering of rock fragments prevent development of an extensive rock veneer and causes a balance, achieved within an estimated 700 years, between the rates of rock-fragment exposure by root throw and clast disintegration by chemical reduction. The estimated rate of rock-fragment reduction accounts for part of the fluvial sediment yields observed for forested subalpine areas of western North America. Copyright ?? 2005 John Wiley & Sons, Ltd.

  5. Invasion of subalpine meadows by lodgepole pine in Yellowstone National Park, Wyoming, USA

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    Jakubos, B.; Romme, W.H. (Fort Lewis College, Durango, CO (United States))


    Many of the dry and mesic subalpine meadows in Yellowstone National Park are bordered by bands of small lodgepole pine trees. The authors asked whether these stands of small trees represent a directional process of meadow invasion, or alternatively, (1) small patches of postfire succession or; (2) more-or-less stable populations of trees having small stature because of adverse site conditions. Transect studies revealed that the bands of small trees were consistently younger than adjacent forest stands of obvious fire origin, that they lacked any evidence of fire, and that the trees were progressively younger as they approached the meadow. Soils under the young trees generally were more similar to meadow soils than to coniferous forest soils. The authors concluded, therefore, that meadow invasion has been occurring as a directional process since at least the mid- to late 1800s. Frequency of tree establishment in two dry meadows was positively correlated with mean June temperature and total summer precipitation (R[sup 2] = 0.49, P<0.0001, multiple stepwise regression). Thus, the major cause of tree invasion into dry meadows appears to be a regional climatic trend towards warmer and wetter growing seasons since the end of the Little Ice Age. Tree establishment in two mesic meadows was more weakly and inconsistently correlated with weather variables. Thus, the mechanism of invasion of mesic meadows may involve interactions of episodic seed crops and microhabitat changes at the forest border, as well as regional climatic variability. 30 refs., 7 figs., 1 tab.

  6. Targeted grazing for the restoration of sub-alpine shrub-encroached grasslands

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    Massimiliano Probo


    Full Text Available The decline of agro-pastoral activities has led to a widespread tree and shrub-encroachment of former semi-natural meso-eutrophic grasslands in many European mountain regions. Temporary night camp areas (TNCA and mineral mix supplements for targeted cattle were arranged over shrub-encroached areas to restore grassland vegetation within the Val Troncea Natural Park (Italy. From 2011 to 2015, their effects on vegetation structure and pastoral value of forage were assessed along permanent transects. Four years after treatments, both practices were effective in reducing the shrub cover and increasing the cover and average height of the herbaceous layer, but changes were more remarkable within TNCA. Moreover, the arrangement of TNCA decreased the cover of nanophanerophytes and increased the cover of graminoids and high quality species, as well as the overall forage pastoral value. In conclusion, TNCA were the most effective pastoral practice to contrast shrub-encroachment and increase herbage mass and forage quality of sub-alpine grasslands.

  7. Extensive wildfires, climate change, and an abrupt state change in subalpine ribbon forests, Colorado. (United States)

    Calder, W John; Shuman, Bryan


    Ecosystems may shift abruptly when the effects of climate change and disturbance interact, and landscapes with regularly patterned vegetation may be especially vulnerable to abrupt shifts. Here we use a fossil pollen record from a regularly patterned ribbon forest (alternating bands of forests and meadows) in Colorado to examine whether past changes in wildfire and climate produced abrupt vegetation shifts. Comparing the percentages of conifer pollen with sedimentary δ18 O data (interpreted as an indicator of temperature or snow accumulation) indicates a first-order linear relationship between vegetation composition and climate change with no detectable lags over the past 2,500 yr (r = 0.55, P changed abruptly within a century of extensive wildfires, which were recognized in a previous study to have burned approximately 80% of the surrounding 1,000 km2 landscape 1,000 yr ago when temperatures rose ~0.5°C. The vegetation change was larger than expected from the effects of climate change alone. Pollen assemblages changed from a composition associated with closed subalpine forests to one similar to modern ribbon forests. Fossil pollen assemblages then remained like those from modern ribbon forests for the following ~1,000 yr, providing a clear example of how extensive disturbances can trigger persistent new vegetation states and alter how vegetation responds to climate. © 2017 by the Ecological Society of America.

  8. Litter Production, Decomposition, and Nutrient Release in Subalpine Forest Communities of the Northwest Himalaya

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    Vinod K. Bisht


    Full Text Available Production, decomposition, and release of nutrients from leaf and nonleaf litter were investigated in four subalpine forests of North-West Himalaya, India. Total annual litter fall in four communities varied from 2950.00 to 4040.00 kg ha−1 and was found significant (CD0.05 = 118.2. Decomposition of leaf litter varied from 1.82–3.5% during autumn-winter to 36.14–45.51 during summer rainy season in all stands and percent of mass loss was significantly varied in stands (CD6.00. Similarly, decomposition in nonleaf litter was varied from 0.3–1.1% during autumn-winter to 19.59–30.05% during summer rainy season and was significantly varied irrespective of seasons. However, percent decomposition of leaf litter and the values of decay constant (k were at par in all stands. Total standing state of nutrients in fresh litter as well as release of total nitrogen (N, phosphorus (P, and potassium (K in due course of decomposition (12 months was also varying significantly. The rate of nonleaf litter decomposition was significantly positively correlated with air temperature (r=0.63–0.74 in all communities. The significant correlation (r=0.85 was observed only in Rhododendron-Sorbus forest community (PRS. Study indicates that the air temperature is a major determinant for nonleaf litter decomposition in this region.

  9. Does the aboveground herbivore assemblage influence soil bacterial community composition and richness in subalpine grasslands? (United States)

    Hodel, Melanie; Schütz, Martin; Vandegehuchte, Martijn L; Frey, Beat; Albrecht, Matthias; Busse, Matt D; Risch, Anita C


    Grassland ecosystems support large communities of aboveground herbivores that are known to directly and indirectly affect belowground properties such as the microbial community composition, richness, or biomass. Even though multiple species of functionally different herbivores coexist in grassland ecosystems, most studies have only considered the impact of a single group, i.e., large ungulates (mostly domestic livestock) on microbial communities. Thus, we investigated how the exclusion of four groups of functionally different herbivores affects bacterial community composition, richness, and biomass in two vegetation types with different grazing histories. We progressively excluded large, medium, and small mammals as well as invertebrate herbivores using exclosures at 18 subalpine grassland sites (9 per vegetation type). We assessed the bacterial community composition using terminal restriction fragment length polymorphism (T-RFLP) at each site and exclosure type during three consecutive growing seasons (2009-2011) for rhizosphere and mineral soil separately. In addition, we determined microbial biomass carbon (MBC), root biomass, plant carbon:nitrogen ratio, soil temperature, and soil moisture. Even though several of these variables were affected by herbivore exclusion and vegetation type, against our expectations, bacterial community composition, richness, or MBC were not. Yet, bacterial communities strongly differed between the three growing seasons as well as to some extent between our study sites. Thus, our study indicates that the spatiotemporal variability in soil microclimate has much stronger effects on the soil bacterial communities than the grazing regime or the composition of the vegetation in this high-elevation ecosystem.

  10. Do insects lose flight before they lose their wings? Population genetic structure in subalpine stoneflies. (United States)

    McCulloch, Graham A; Wallis, Graham P; Waters, Jonathan M


    Wing reduction and flightlessness are common features of alpine and subalpine insects, and are typically interpreted as evolutionary adaptations to increase fecundity and promote local recruitment. Here, we assess the impact of wing reduction on dispersal in stoneflies (Plecoptera: Gripopterygidae: Zelandoperla) in southern New Zealand. Specifically, we present comparative phylogeographic analyses (COI; H3) of strong-flying Zelandoperla decorata (144 individuals, 63 localities) vs. the co-distributed but weak-flying Zelandoperla fenestrata species group (186 individuals, 81 localities). The latter group exhibits a variety of morphotypes, ranging from fully winged to completely wingless. Consistent with its capacity for strong flight-mediated dispersal, Z. decorata exhibited no substantial phylogeographic differentiation across its broad South Island range. Conversely the weak-flying fenestrata species group exhibited substantial genetic structure across both fine and broad geographic scales. Intriguingly, the variable degrees of wing development observed within the fenestrata species group had no apparent impact on levels of phylogeographic structure, which were high regardless of morphotype, suggesting that even fully winged specimens of this group do not fly. This finding implies that Zelandoperla flight loss occurs independently of wing loss, and might reflect underlying flight muscle reduction.

  11. Optimum FIR filter for sampled signals in presence of jitter (United States)

    Cattaneo, Paolo Walter


    The requirements of the integrated readout electronics for calorimetry at high luminosity hadron colliders pose new challenges both to hardware design and to the performance of signal processing algorithms. Both aspects have been treated in detail by the FERMI(RD16) collaboration [C. Alippi et al., Nucl. Instr. and Meth. A 344 (1994) 180], from which this work has been motivated. The estimation of the amplitude of sampled signals is usually performed with a digital FIR filter, or with a more sophisticated non linear digital filter using FIR filters as building blocks [S.J. Inkinen and J. Niittylahti, Trainable FIR-order statistic hybrid filters, to be published in IEEE Trans. Circuits and Systems; H. Alexanian et al., FERMI Collaboration, Optimized digital feature extraction in the FERMI microsystem Nucl. Instr. and Meth. A 357 (1995)]. In presence of significant signal phase jitter with respect to the clock, the phase dependence of the filter output can be a major source of error. This is especially true for measurements of large amplitudes for which the effect of electronic noise becomes negligible. This paper reports on the determination of digital FIR filters that optimize the signal over noise ratio due to known jitter distributions for different filter lengths. As the presence of electronic noise is neglected, the results are mainly relevant for measurements of large signals. FERMI is a collaboration with the aim of designing integrated electronics for the read out of calorimeter detectors in particle physics experiments at hadron colliders. It includes: CERN, Geneva, Switzerland; Department of Physics and Measurement Technology, University of Linköping, Sweden; Center for Industrial Microelectronics and Materials Technology, University of Linköping, Sweden; LPNHE Universities Paris VI-VII, Paris, France; Dipartimento di Elettronica, Politecnico di Milano, Italy, Sezine INFN, Pavia, Italy; Dipartimento di Fisica Nucleare e Teorica dell'Universitá e Sezione

  12. Space-time modelling of lightning-caused ignitions in the Blue Mountains, Oregon (United States)

    Diaz-Avalos, Carlos; Peterson, D.L.; Alvarado, Ernesto; Ferguson, Sue A.; Besag, Julian E.


    Generalized linear mixed models (GLMM) were used to study the effect of vegetation cover, elevation, slope, and precipitation on the probability of ignition in the Blue Mountains, Oregon, and to estimate the probability of ignition occurrence at different locations in space and in time. Data on starting location of lightning-caused ignitions in the Blue Mountains between April 1986 and September 1993 constituted the base for the analysis. The study area was divided into a pixela??time array. For each pixela??time location we associated a value of 1 if at least one ignition occurred and 0 otherwise. Covariate information for each pixel was obtained using a geographic information system. The GLMMs were fitted in a Bayesian framework. Higher ignition probabilities were associated with the following cover types: subalpine herbaceous, alpine tundra, lodgepole pine (Pinus contorta Dougl. ex Loud.), whitebark pine (Pinus albicaulis Engelm.), Engelmann spruce (Picea engelmannii Parry ex Engelm.), subalpine fir (Abies lasiocarpa (Hook.) Nutt.), and grand fir (Abies grandis (Dougl.) Lindl.). Within each vegetation type, higher ignition probabilities occurred at lower elevations. Additionally, ignition probabilities are lower in the northern and southern extremes of the Blue Mountains. The GLMM procedure used here is suitable for analysing ignition occurrence in other forested regions where probabilities of ignition are highly variable because of a spatially complex biophysical environment.

  13. Structural properties of laminated Douglas fir/epoxy composite material

    Energy Technology Data Exchange (ETDEWEB)

    Spera, D.A. (National Aeronautics and Space Administration, Cleveland, OH (USA). Lewis Research Center); Esgar, J.B. (Sverdrup Technology, Inc., Cleveland, OH (USA)); Gougeon, M.; Zuteck, M.D. (Gougeon Bros., Bay City, MI (USA))


    This publication contains a compilation of static and fatigue and strength data for laminated-wood material made from Douglas fir and epoxy. Results of tests conducted by several organizations are correlated to provide insight into the effects of variables such as moisture, size, lamina-to-lamina joint design, wood veneer grade, and the ratio of cyclic stress to steady stress during fatigue testing. These test data were originally obtained during development of wood rotor blades for large-scale wind turbines of the horizontal-axis (propeller) configuration. Most of the strength property data in this compilation are not found in the published literature. Test sections ranged from round cylinders 2.25 in. in diameter to rectangular slabs 6 in. by 24 in. in cross section and approximately 30 ft long. All specimens were made from Douglas fir veneers 0.10 in. thick, bonded together with the WEST epoxy system developed for fabrication and repair of wood boats. Loading was usually parallel to the grain. Size effects (reduction in strength with increase in test volume) are observed in some of the test data, and a simple mathematical model is presented that includes the probability of failure. General characteristics of the wood/epoxy laminate are discussed, including features that make it useful for a wide variety of applications. 9 refs.

  14. Long-term reactive nitrogen loading alters soil carbon and microbial community properties in a subalpine forest ecosystem (United States)

    Boot, Claudia M; Hall, Ed K.; Denef, Karolien; Baron, Jill S.


    Elevated nitrogen (N) deposition due to increased fossil fuel combustion and agricultural practices has altered global carbon (C) cycling. Additions of reactive N to N-limited environments are typically accompanied by increases in plant biomass. Soil C dynamics, however, have shown a range of different responses to the addition of reactive N that seem to be ecosystem dependent. We evaluated the effect of N amendments on biogeochemical characteristics and microbial responses of subalpine forest organic soils in order to develop a mechanistic understanding of how soils are affected by N amendments in subalpine ecosystems. We measured a suite of responses across three years (2011–2013) during two seasons (spring and fall). Following 17 years of N amendments, fertilized soils were more acidic (control mean 5.09, fertilized mean 4.68), and had lower %C (control mean 33.7% C, fertilized mean 29.8% C) and microbial biomass C by 22% relative to control plots. Shifts in biogeochemical properties in fertilized plots were associated with an altered microbial community driven by reduced arbuscular mycorrhizal (control mean 3.2 mol%, fertilized mean 2.5 mol%) and saprotrophic fungal groups (control mean 17.0 mol%, fertilized mean 15.2 mol%), as well as a decrease in N degrading microbial enzyme activity. Our results suggest that decreases in soil C in subalpine forests were in part driven by increased microbial degradation of soil organic matter and reduced inputs to soil organic matter in the form of microbial biomass.

  15. Trace elements in sub-alpine forest soils on the eastern edge of the Tibetan Plateau, China (United States)

    Wang, Xiaodan; Cheng, Genwei; Zhong, Xianghao; Li, Mai-He


    Industrial development has increased fast in China during the last decades. This has led to a range of environmental problems. Deposition of trace elements to forest ecosystems via the atmosphere is one potential problem. In this paper, we report the results from a pilot study where the trace element levels of the sub-alpine forest soils on the eastern edge of the Tibetan Plateau have been measured. Possible relationships between soil properties and trace element concentrations have also been investigated. The obtained concentrations (mg kg-1) were boron (B) 48.06-53.70, molybdenum (Mo) 1.53-2.26, zinc (Zn) 68.18-79.53, copper (Cu) 36.81-42.44, selenium (Se) 0.33-0.49, cadmium (Cd) 0.16-0.29, lead (Pb) 25.80-30.71, chromium (Cr) 96.10-110.08, nickel (Ni) 30.16-45.60, mercury (Hg) 0.05-0.11, and arsenic (As) 3.09-4.17. With a few exceptions, the element concentration can be characterized as low in the investigated sub-alpine forest soils. No clear differences in trace element levels were found between topsoil and subsoil samples, indicating that the atmospheric deposition of trace element has been low. The soil parent material plays a key role to determine trace element levels. Soil properties, including pHw, organic carbon (OC), clay fraction, cation-exchange capacity (CEC), total iron (Fe), and total aluminum (Al) concentrations were related to trace element concentration using correlation analysis. Total Fe and Al showed the strongest relationships with concentrations of most trace elements in the sub-alpine forest soils. PCA analyses indicated that a significant increase in the number of cars with the fast development of local tourism may result in higher Pb concentration in the future.

  16. Quantitative characterization of the interactions among c-myc transcriptional regulators FUSE, FBP, and FIR. (United States)

    Hsiao, Hsin-Hao; Nath, Abhinav; Lin, Chi-Yen; Folta-Stogniew, Ewa J; Rhoades, Elizabeth; Braddock, Demetrios T


    Human c-myc is critical for cell homeostasis and growth but is a potent oncogenic factor if improperly regulated. The c-myc far-upstream element (FUSE) melts into single-stranded DNA upon active transcription, and the noncoding strand FUSE recruits an activator [the FUSE-binding protein (FBP)] and a repressor [the FBP-interacting repressor (FIR)] to fine-tune c-myc transcription in a real-time manner. Despite detailed biological experiments describing this unique mode of transcriptional regulation, quantitative measurements of the physical constants regulating the protein-DNA interactions remain lacking. Here, we first demonstrate that the two FUSE strands adopt different conformations upon melting, with the noncoding strand DNA in an extended, linear form. FBP binds to the linear noncoding FUSE with a dissociation constant in the nanomolar range. FIR binds to FUSE more weakly, having its modest dissociation constants in the low micromolar range. FIR is monomeric under near-physiological conditions but upon binding of FUSE dimerizes into a 2:1 FIR(2)-FUSE complex mediated by the RRMs. In the tripartite interaction, our analysis suggests a stepwise addition of FIR onto an activating FBP-FUSE complex to form a quaternary FIR(2)-FBP-FUSE inhibitory complex. Our quantitative characterization enhances understanding of DNA strand preference and the mechanism of the stepwise complex formation in the FUSE-FBP-FIR regulatory system.

  17. Tree Biomass Estimation of Chinese fir (Cunninghamia lanceolata) Based on Bayesian Method (United States)

    Zhang, Jianguo


    Chinese fir (Cunninghamia lanceolata (Lamb.) Hook.) is the most important conifer species for timber production with huge distribution area in southern China. Accurate estimation of biomass is required for accounting and monitoring Chinese forest carbon stocking. In the study, allometric equation was used to analyze tree biomass of Chinese fir. The common methods for estimating allometric model have taken the classical approach based on the frequency interpretation of probability. However, many different biotic and abiotic factors introduce variability in Chinese fir biomass model, suggesting that parameters of biomass model are better represented by probability distributions rather than fixed values as classical method. To deal with the problem, Bayesian method was used for estimating Chinese fir biomass model. In the Bayesian framework, two priors were introduced: non-informative priors and informative priors. For informative priors, 32 biomass equations of Chinese fir were collected from published literature in the paper. The parameter distributions from published literature were regarded as prior distributions in Bayesian model for estimating Chinese fir biomass. Therefore, the Bayesian method with informative priors was better than non-informative priors and classical method, which provides a reasonable method for estimating Chinese fir biomass. PMID:24278198

  18. Tree biomass estimation of Chinese fir (Cunninghamia lanceolata based on Bayesian method.

    Directory of Open Access Journals (Sweden)

    Xiongqing Zhang

    Full Text Available Chinese fir (Cunninghamia lanceolata (Lamb. Hook. is the most important conifer species for timber production with huge distribution area in southern China. Accurate estimation of biomass is required for accounting and monitoring Chinese forest carbon stocking. In the study, allometric equation W = a(D2Hb was used to analyze tree biomass of Chinese fir. The common methods for estimating allometric model have taken the classical approach based on the frequency interpretation of probability. However, many different biotic and abiotic factors introduce variability in Chinese fir biomass model, suggesting that parameters of biomass model are better represented by probability distributions rather than fixed values as classical method. To deal with the problem, Bayesian method was used for estimating Chinese fir biomass model. In the Bayesian framework, two priors were introduced: non-informative priors and informative priors. For informative priors, 32 biomass equations of Chinese fir were collected from published literature in the paper. The parameter distributions from published literature were regarded as prior distributions in Bayesian model for estimating Chinese fir biomass. Therefore, the Bayesian method with informative priors was better than non-informative priors and classical method, which provides a reasonable method for estimating Chinese fir biomass.

  19. Assessing heat fluxes and water quality trends in subalpine lakes from EO (United States)

    Cazzaniga, Ilaria; Giardino, Claudia; Bresciani, Mariano; Elli, Chiara; Valerio, Giulia; Pilotti, Marco


    Lakes play a fundamental role in providing ecosystem services such as water supplying, hydrological regulation, climate change mitigation, touristic recreation (Schallenberg et al., 2013). Preserving and improving of quality of lakes waters, which is a function of either both natural and human influences, is therefore an important action to be considered. Remote Sensing techniques are spreading as useful instrument for lakes, by integrating classical in situ limnological measurements to frequent and synoptic monitoring capabilities. Within this study, Earth Observation data are exploited for understanding the temporal changes of water quality parameters over a decade, as well as for measuring the surface energy fluxes in recent years in deep clear lakes in the European subalpine ecoregion. According to Pareth et al. (2016), subalpine lakes are showing a clear response to climate change with an increase of 0.017 °C /year of lake surface temperature, whilst the human activities contribute to produce a large impact (agriculture, recreation, industry, fishing and drinking) on these lakes. The investigation is focused on Lake Iseo, which has shown a significant deterioration of water quality conditions since the seventies, and on Lake Garda, the largest Italian lake where EO data have been widely used for many purposes and applications (Giardino et al., 2014). Available ENVISAT-MERIS (2002-2012) and Landsat-8-OLI (2013-on going) imagery has been exploited to produce chlorophyll-a (chl-a) concentration maps, while Landsat-8-TIRS imagery has been used for estimating lake surface temperatures. MERIS images were processed through a neural network (namely the C2R processor, Doerffer et al., 2007), to correct the atmospheric effects and to retrieve water constituents concentration in optically complex deep waters. With regard to L8's images, some atmospheric correctors (e.g. ACOLITE and 6SV) were tested and validated to indentify, for each of the two lakes, the more accurate

  20. Attempt to induce lightwood in balsam fir and tamarack by treating with paraquat

    Energy Technology Data Exchange (ETDEWEB)

    Kiatgrajai, P. (Univ. of Wisconsin, Madison); Conner, A.H.; Rowe, J.W.; Peters, W.; Roberts, D.R.


    Treatment of balsam fir (Abies balsamea (L.) Mill.) and tamarack (Larix laricina (Du Roi) K. Koch) with the herbicide paraquat did not induce formation of lightwood. However, a dead zone of phloem extended above the paraquat-treatment site. In balsam fir, the wood directly behind the dead zone was particularly heavily attacked by fungi. In both species, traumatic resin ducts were observed in the last several growth rings of wood adjacent to the treatment site. The ether extract of the tamarack, but not of the balsam fir, was essentially oleoresin.

  1. Hydrologic flow path development varies by aspect during spring snowmelt in complex subalpine terrain (United States)

    Webb, Ryan W.; Fassnacht, Steven R.; Gooseff, Michael N.


    In many mountainous regions around the world, snow and soil moisture are key components of the hydrologic cycle. Preferential flow paths of snowmelt water through snow have been known to occur for years with few studies observing the effect on soil moisture. In this study, statistical analysis of the topographical and hydrological controls on the spatiotemporal variability of snow water equivalent (SWE) and soil moisture during snowmelt was undertaken at a subalpine forested setting with north, south, and flat aspects as a seasonally persistent snowpack melts. We investigated if evidence of preferential flow paths in snow can be observed and the effect on soil moisture through measurements of snow water equivalent and near-surface soil moisture, observing how SWE and near-surface soil moisture vary on hillslopes relative to the toes of hillslopes and flat areas. We then compared snowmelt infiltration beyond the near-surface soil between flat and sloping terrain during the entire snowmelt season using soil moisture sensor profiles. This study was conducted during varying snowmelt seasons representing above-normal, relatively normal, and below-normal snow seasons in northern Colorado. Evidence is presented of preferential meltwater flow paths at the snow-soil interface on the north-facing slope causing increases in SWE downslope and less infiltration into the soil at 20 cm depth; less association is observed in the near-surface soil moisture (top 7 cm). We present a conceptualization of the meltwater flow paths that develop based on slope aspect and soil properties. The resulting flow paths are shown to divert at least 4 % of snowmelt laterally, accumulating along the length of the slope, to increase the snow water equivalent by as much as 170 % at the base of a north-facing hillslope. Results from this study show that snow acts as an extension of the vadose zone during spring snowmelt and future hydrologic investigations will benefit from studying the snow and soil

  2. Conifer density within lake catchments predicts fish mercury concentrations in remote subalpine lakes (United States)

    Eagles-Smith, Collin A.; Herring, Garth; Johnson, Branden L.; Graw, Rick


    Remote high-elevation lakes represent unique environments for evaluating the bioaccumulation of atmospherically deposited mercury through freshwater food webs, as well as for evaluating the relative importance of mercury loading versus landscape influences on mercury bioaccumulation. The increase in mercury deposition to these systems over the past century, coupled with their limited exposure to direct anthropogenic disturbance make them useful indicators for estimating how changes in mercury emissions may propagate to changes in Hg bioaccumulation and ecological risk. We evaluated mercury concentrations in resident fish from 28 high-elevation, sub-alpine lakes in the Pacific Northwest region of the United States. Fish total mercury (THg) concentrations ranged from 4 to 438 ng/g wet weight, with a geometric mean concentration (±standard error) of 43 ± 2 ng/g ww. Fish THg concentrations were negatively correlated with relative condition factor, indicating that faster growing fish that are in better condition have lower THg concentrations. Across the 28 study lakes, mean THg concentrations of resident salmonid fishes varied as much as 18-fold among lakes. We used a hierarchal statistical approach to evaluate the relative importance of physiological, limnological, and catchment drivers of fish Hg concentrations. Our top statistical model explained 87% of the variability in fish THg concentrations among lakes with four key landscape and limnological variables: catchment conifer density (basal area of conifers within a lake's catchment), lake surface area, aqueous dissolved sulfate, and dissolved organic carbon. Conifer density within a lake's catchment was the most important variable explaining fish THg concentrations across lakes, with THg concentrations differing by more than 400 percent across the forest density spectrum. These results illustrate the importance of landscape characteristics in controlling mercury bioaccumulation in fish.

  3. Calcium induces long-term legacy effects in a subalpine ecosystem.

    Directory of Open Access Journals (Sweden)

    Urs Schaffner

    Full Text Available Human activities have transformed a significant proportion of the world's land surface, with profound effects on ecosystem processes. Soil applications of macronutrients such as nitrate, phosphorus, potassium or calcium are routinely used in the management of croplands, grasslands and forests to improve plant health or increase productivity. However, while the effects of continuous fertilization and liming on terrestrial ecosystems are well documented, remarkably little is known about the legacy effect of historical fertilization and liming events in terrestrial ecosystems and of the mechanisms involved. Here, we show that more than 70 years after the last application of lime on a subalpine grassland, all major soil and plant calcium pools were still significantly larger in limed than in unlimed plots, and that the resulting shift in the soil calcium/aluminium ratio continues to affect ecosystem services such as primary production. The difference in the calcium content of the vegetation and the topmost 10 cm of the soil in limed vs. unlimed plots amounts to approximately 19.5 g m(-2, equivalent to 16.3% of the amount that was added to the plots some 70 years ago. In contrast, plots that were treated with nitrogen-phosphorus-potassium fertilizer in the 1930s did not differ from unfertilized plots in any of the soil and vegetation characteristics measured. Our findings suggest that the long-term legacy effect of historical liming is due to long-term storage of added calcium in stable soil pools, rather than a general increase in nutrient availability. Our results demonstrate that single applications of calcium in its carbonated form can profoundly and persistently alter ecosystem processes and services in mountain ecosystems.

  4. Net primary productivity of subalpine meadows in Yosemite National Park in relation to climate variability (United States)

    Moore, Peggy E.; Van Wagtendonk, Jan W.; Yee, Julie L.; McClaran, Mitchel P.; Cole, David N.; McDougald, Neil K.; Brooks, Matthew L.


    Subalpine meadows are some of the most ecologically important components of mountain landscapes, and primary productivity is important to the maintenance of meadow functions. Understanding how changes in primary productivity are associated with variability in moisture and temperature will become increasingly important with current and anticipated changes in climate. Our objective was to describe patterns and variability in aboveground live vascular plant biomass in relation to climatic factors. We harvested aboveground biomass at peak growth from four 64-m2 plots each in xeric, mesic, and hydric meadows annually from 1994 to 2000. Data from nearby weather stations provided independent variables of spring snow water content, snow-free date, and thawing degree days for a cumulative index of available energy. We assembled these climatic variables into a set of mixed effects analysis of covariance models to evaluate their relationships with annual aboveground net primary productivity (ANPP), and we used an information theoretic approach to compare the quality of fit among candidate models. ANPP in the xeric meadow was negatively related to snow water content and thawing degree days and in the mesic meadow was negatively related to snow water content. Relationships between ANPP and these 2 covariates in the hydric meadow were not significant. Increasing snow water content may limit ANPP in these meadows if anaerobic conditions delay microbial activity and nutrient availability. Increased thawing degree days may limit ANPP in xeric meadows by prematurely depleting soil moisture. Large within-year variation of ANPP in the hydric meadow limited sensitivity to the climatic variables. These relationships suggest that, under projected warmer and drier conditions, ANPP will increase in mesic meadows but remain unchanged in xeric meadows because declines associated with increased temperatures would offset the increases from decreased snow water content.

  5. Intraguild predation and cannibalism among larvae of detritivorous caddisflies in subalpine wetlands (United States)

    Wissinger, S.A.; Sparks, G.B.; Rouse, G.L.; Brown, W.S.; Steltzer, Heidi


    Comparative data from subalpine wetlands in Colorado indicate that larvae of the limnephilid caddisflies, Asynarchus nigriculus and Limnephilus externus, are reciprocally abundant among habitats - Limnephilus larvae dominate in permanent waters, whereas Asynarchus larvae dominate in temporary basins. The purpose of this paper is to report on field and laboratory experiments that link this pattern of abundance to biotic interactions among larvae. In the first field experiment, growth and survival were compared in single and mixed species treatments in littoral enclosures. Larvae, which eat mainly vascular plant detritus, grew at similar rates among treatments in both temporary and permanent habitats suggesting that exploitative competition is not important under natural food levels and caddisfly densities. However, the survival of Limnephilus larvae was reduced in the presence of Asynarchus larvae. Subsequent behavioral studies in laboratory arenas revealed that Asynarchus larvae are extremely aggressive predators on Limnephilus larvae. In a second field experiment we manipulated the relative sizes of larvae and found that Limnephilus larvae were preyed on only when Asynarchus larvae had the same size advantage observed in natural populations. Our data suggest that the dominance of Asynarchus larvae in temporary habitats is due to asymmetric intraguild predation (IGP) facilitated by a phenological head start in development. These data do not explain the dominance of Limnephilus larvae in permanent basins, which we show elsewhere to be an indirect effect of salamander predation. Behavioral observations also revealed that Asynarchus larvae are cannibalistic. In contrast to the IGP on Limnephilus larvae, Asynarchus cannibalism occurs among same-sized larvae and often involves the mobbing of one victim by several conspecifics. In a third field experiment, we found that Asynarchus cannibalism was not density-dependent and occurred even at low larval densities. We

  6. Nitrogen deposition but not ozone affects productivity and community composition of subalpine grassland after 3 yr of treatment. (United States)

    Bassin, Seraina; Volk, Matthias; Suter, Matthias; Buchmann, Nina; Fuhrer, Jürg


    A field experiment was established at 2000 m above sea level (asl) in the central Swiss Alps with the aim of investigating the effects of elevated ozone (O(3)) and nitrogen deposition (N), and of their combination, on above-ground productivity and species composition of subalpine grassland. One hundred and eighty monoliths were extracted from a species-rich Geo-Montani-Nardetum pasture and exposed in a free-air O(3)-fumigation system to one of three concentrations of O(3) (ambient, 1.2 x ambient, 1.6 x ambient) and five concentrations of additional N. Above-ground biomass, proportion of functional groups and normalized difference vegetation index (NDVI) were measured annually. After 3 yr of treatment, the vegetation responded to the N input with an increase in above-ground productivity and altered species composition, but without changes resulting from elevated O(3). N input > 10 kg N ha(-1) yr(-1) was sufficient to affect the composition of functional groups, with sedges benefiting over-proportionally. No interaction of O(3) x N was observed, except for NDVI; positive effects of N addition on canopy greenness were counteracted by accelerated leaf senescence in the highest O(3) treatment. The results suggest that effects of elevated O(3) on the productivity and floristic composition of subalpine grassland may develop slowly, regardless of the sensitive response to increasing N.

  7. Effects of tourism and topography on vegetation diversity in the subalpine meadows of the Dongling Mountains of Beijing, China. (United States)

    Zhang, Jin-Tun; Xiang, ChunLing; Li, Min


    Subalpine meadows in the Dongling Mountains (located at E115º26'-115º40', N40º00'-40º05') of Beijing, China are important for tourism and the provision of ecosystem services. However, because of poor management serious degradation has occurred on these subalpine meadows. The aim of this paper is to present a quantitative analysis of effects of tourism disturbance and topography on the status and diversity of montane meadow communities and to provide direction for improved management. Sixty quadrats of 2 × 2 m(2) along 10 transects were set up to collect data on site characteristics and vegetation status. The relationships between community composition and structure, species diversity, and tourism disturbance and topographic variables were analyzed by multivariate methods (TWINSPAN and CCA). The results showed that eight meadow communities were identified by TWINSPAN. Most of them were seriously degraded. The first CCA axis identified an elevation and tourism disturbance intensity gradient, which illustrated that tourism disturbance and elevation were most important factors influencing meadow types, composition and structure. Some resistant species and response species to tourism disturbance were identified and can be used as indicator species of tourism disturbance. Species richness, heterogeneity and evenness were closely related to tourism disturbance and elevation. It is concluded that tourism disturbance must be controlled to enable grassland rehabilitation to occur in the meadows. Measures of effective management of the meadows were discussed.

  8. Effects of Tourism and Topography on Vegetation Diversity in the Subalpine Meadows of the Dongling Mountains of Beijing, China (United States)

    Zhang, Jin-Tun; Xiang, Chunling; Li, Min


    Subalpine meadows in the Dongling Mountains (located at E115º26'-115º40', N40º00'-40º05') of Beijing, China are important for tourism and the provision of ecosystem services. However, because of poor management serious degradation has occurred on these subalpine meadows. The aim of this paper is to present a quantitative analysis of effects of tourism disturbance and topography on the status and diversity of montane meadow communities and to provide direction for improved management. Sixty quadrats of 2 × 2 m2 along 10 transects were set up to collect data on site characteristics and vegetation status. The relationships between community composition and structure, species diversity, and tourism disturbance and topographic variables were analyzed by multivariate methods (TWINSPAN and CCA). The results showed that eight meadow communities were identified by TWINSPAN. Most of them were seriously degraded. The first CCA axis identified an elevation and tourism disturbance intensity gradient, which illustrated that tourism disturbance and elevation were most important factors influencing meadow types, composition and structure. Some resistant species and response species to tourism disturbance were identified and can be used as indicator species of tourism disturbance. Species richness, heterogeneity and evenness were closely related to tourism disturbance and elevation. It is concluded that tourism disturbance must be controlled to enable grassland rehabilitation to occur in the meadows. Measures of effective management of the meadows were discussed.

  9. FIR Detectors/Cameras Based on GaN and Si Field-Effect Devices Project (United States)

    National Aeronautics and Space Administration — SETI proposes to develop GaN and Si based multicolor FIR/THz cameras with detector elements and readout, signal processing electronics integrated on a single chip....

  10. Ternary and Multi-Bit FIR Filter Area-Performance Tradeoffs in FPGA

    Directory of Open Access Journals (Sweden)

    Khalil-Ur-Rahman Dayo


    Full Text Available In this paper, performance and area of conventional FIR (Finite Impulse Responce filters versus ternary sigma delta modulated FIR filter is compared in FPGA (Field Programmable Gate Arrays using VHDL (Verilog Description Language. Two different approaches were designed and synthesized at same spectral performance by obtaining a TIR (Target Impulse Response. Both filters were synthesized on adaptive LUT (Look Up Table FPGA device in pipelined and non-pipelined modes. It is shown that the Ternary FIR filter occupies approximately the same area as the corresponding multi-bit filter, but for a given specification, the ternary FIR filter has 32% better performance in non-pipelined and 72% in pipelined mode, compared to its equivalent Multi-Bit filter at its optimum 12-bit coefficient quantization. These promising results shows that ternary logic based (i.e. +1,0,-1 filters can be used for huge chip area savings and higher performance.

  11. Active cancellation of acoustical resonances with an FPGA FIR filter (United States)

    Ryou, Albert; Simon, Jonathan


    We present a novel approach to enhancing the bandwidth of a feedback-controlled mechanical system by digitally canceling acoustical resonances (poles) and anti-resonances (zeros) in the open-loop response via an FPGA FIR filter. By performing a real-time convolution of the feedback error signal with an inverse filter, we can suppress arbitrarily many poles and zeros below 100 kHz, each with a linewidth down to 10 Hz. We demonstrate the efficacy of this technique by canceling the ten largest mechanical resonances and anti-resonances of a high-finesse optical resonator, thereby enhancing the unity gain frequency by more than an order of magnitude. This approach is applicable to a broad array of stabilization problems including optical resonators, external cavity diode lasers, and scanning tunneling microscopes and points the way to applying modern optimal control techniques to intricate linear acoustical systems.

  12. Subalpine vegetation pattern three decades after stand-replacing fire: Effects of landscape context and topography on plant community composition, tree regeneration, and diversity (United States)

    Jonathan D. Coop; Robert T. Massatti; Anna W. Schoettle


    These subalpine wildfires generated considerable, persistent increases in plant species richness at local and landscape scales, and a diversity of plant communities. The findings suggest that fire suppression in such systems must lead to reduced diversity. Concerns about post-fire invasion by exotic plants appear unwarranted in high-elevation wilderness settings.

  13. Verification of satellite radar remote sensing based estimates of boreal and subalpine growing seasons using an ecosystem process model and surface biophysical measurement network information (United States)

    McDonald, K. C.; Kimball, J. S.; Zimmerman, R.


    We employ daily surface Radar backscatter data from the SeaWinds Ku-band Scatterometer onboard Quikscat to estimate landscape freeze-thaw state and associated length of the seasonal non-frozen period as a surrogate for determining the annual growing season across boreal and subalpine regions of North America for 2000 and 2001.

  14. Carex sempervirens tussocks induce spatial heterogeneity in litter decomposition, but not in soil properties, in a subalpine grassland in the Central Alps (United States)

    Fei-Hai Yu; Martin Schutz; Deborah S. Page-Dumroese; Bertil O. Krusi; Jakob Schneller; Otto Wildi; Anita C. Risch


    Tussocks of graminoids can induce spatial heterogeneity in soil properties in dry areas with discontinuous vegetation cover, but little is known about the situation in areas with continuous vegetation and no study has tested whether tussocks can induce spatial heterogeneity in litter decomposition. In a subalpine grassland in the Central Alps where vegetation cover is...

  15. Deforestation induces shallow landsliding in the montane and subalpine belts of the Urbión Mountains, Iberian Range, Northern Spain (United States)

    García-Ruiz, José M.; Beguería, Santiago; Arnáez, José; Sanjuán, Yasmina; Lana-Renault, Noemí; Gómez-Villar, Amelia; Álvarez-Martínez, Javier; Coba-Pérez, Paz


    In this study the spatial distribution of shallow landslides in the upper montane and subalpine belts of the Urbión Mountains (Iberian Range, northern Spain) was investigated, particularly in relation to the spatial organization of deforestation and land cover. The upper montane and subalpine belts have been deforested several times since the Neolithic Period, to enlarge the area of summer grasslands for feeding transhumant sheep flocks. Consequently, the timberline was lowered by 400-600 m, and increased the occurrence of severe erosion processes, particularly shallow landslides. This study shows that most of the landslide scars are in the summer grasslands area, and that a remarkable extent of the subalpine belt area has been subjected to mass movements. In addition to land use, the soil characteristics and topography help explain the development of conditions most favorable to landsliding. Shallow landslide susceptibility was highest in the upper parts of the slopes near the divides, in areas having slope gradients of 10-30° and deep soils with an increasing proportion of clay with depth. The landslides were clustered and not randomly distributed, and the causes of this spatial distribution are discussed. The current trend of woody encroachment in the upper montane and subalpine belts, resulting from decreasing livestock pressure, will probably reduce the susceptibility of these areas to shallow landslides in the future.

  16. Lab and Field Warming Similarly Advance Germination Date and Limit Germination Rate for High and Low Elevation Provenances of Two Widespread Subalpine Conifers

    Directory of Open Access Journals (Sweden)

    Lara M. Kueppers


    Full Text Available Accurately predicting upslope shifts in subalpine tree ranges with warming requires understanding how future forest populations will be affected by climate change, as these are the seed sources for new tree line and alpine populations. Early life history stages are particularly sensitive to climate and are also influenced by genetic variation among populations. We tested the climate sensitivity of germination and initial development for two widely distributed subalpine conifers, using controlled-environment growth chambers with one temperature regime from subalpine forest in the Colorado Rocky Mountains and one 5 °C warmer, and two soil moisture levels. We tracked germination rate and timing, rate of seedling development, and seedling morphology for two seed provenances separated by ~300 m elevation. Warming advanced germination timing and initial seedling development by a total of ~2 weeks, advances comparable to mean differences between provenances. Advances were similar for both provenances and species; however, warming reduced the overall germination rate, as did low soil moisture, only for Picea engelmannii. A three-year field warming and watering experiment planted with the same species and provenances yielded responses qualitatively consistent with the lab trials. Together these experiments indicate that in a warmer, drier climate, P. engelmannii germination, and thus regeneration, could decline, which could lead to declining subalpine forest populations, while Pinus flexilis forest populations could remain robust as a seed source for upslope range shifts.

  17. Early survival and growth of planted Douglas-fir with red alder in four mixed regimes. (United States)

    Marshall D. Murray; Richard E. Miller


    To quantify between-species interactions, we measured and compared survival and growth of planted Douglas-fir and associated planted and volunteer red alder at a location on the west side of the Cascade Range in Washington. The planted alder were wildlings dug either from a nearby area or from a distant, coastal site and interplanted into a 3-year-old Douglas-fir...

  18. Identification of varieties and gene flow in Douglas fir exemplified in artificially established stands in Germany

    Directory of Open Access Journals (Sweden)

    Barbara Fussi


    Full Text Available Douglas-fir [Pseudotsuga menziesii (Mirb. Franco] is an economically valuable non-native tree species in Germany and is considered very promising in view of global climate change. Therefore, the genetic characterization of Douglas-fir populations and seed stands in Germany is essential. We studied coastal and interior Douglas-fir varieties, both present in Germany, by using eleven isoenzyme and four microsatellite loci. By analyzing eight reference populations of known origin we were able to identify the two varieties on the population level using Bayesian and distance based methods. Seven populations present in Bavaria were then successfully assigned to one of the two varieties. Within varieties we found stronger grouping within the interior variety than within the coastal one. Despite lower differences within coastal Douglas-fir we have first indications for the origin of two populations. For two Bavarian populations, natural regeneration was included and genetic data revealed no significant genetic difference between adults and offspring. The parentage analysis for one of the studied stands revealed that a large proportion of adults took part in the reproduction, but some trees were more successful than others in transferring their genes to the next generation. Our study was able to improve variety identification of Douglas-fir using isoenzyme markers and nuclear microsatellites and study reproductive patterns, both are important issues for the management of Douglas-fir stands in Bavaria.

  19. Importance of nitrogen cycling hot spots in an alpine-subalpine ecosystem (United States)

    Darrouzet-Nardi, A.; Bowman, W. D.


    Nitrogen (N) availability in soils is influenced by many microbially catalyzed reactions such as N fixation, denitrification, and N mineralization from soil organic matter (SOM). Reaction rates for these processes are heterogeneous across landscapes, often forming hot spots that have disproportionately high N cycling activity. N cycling hot spots have been documented in many ecosystems at hourly to weekly times scales; however, much less is known about the persistence and importance of N cycling hot spots over longer times scales. Furthermore, few studies have investigated multiple landscape-level measurements of different N cycling processes at the same site. Using a mathematical definition of hot spots, a time series of short-term measurements, ion exchange resins deployed for one year, and nitrogen isotopic signatures in SOM, we investigated the importance of hot spots over longer time scales in a 0.89 km2 alpine-subalpine ecosystem at the Niwot Ridge LTER site. Measurements of KCl-extractable inorganic N taken on multiple measurement dates showed that hot moments of N availabilty occurred in some but not all parts of the study site and at varying times throughout the season. Ion exchange resins deployed for one year showed that N availability hot spots were important though not completely dominant over one year (14% of values accounted for 58% of total resin-extractable inorganic N observed). In contrast, isotopic signatures in 219 SOM samples were well approximated by a normal distribution, suggesting that landscape-level N losses through leaching or gas efflux were more constrained. Denitrification was the possible exception: we saw evidence for several likely hot spots in the wetland areas of our study site. The results of this study suggest that short-term hot spots are important for plant ecological dynamics at our study site and for denitrification, but that long-term N cycling hot spots are less important for other parts of the soil N cycle such as N

  20. Fuel deposition rates of montane and subalpine conifers in the central Sierra Nevada, California, USA (United States)

    van Wagtendonk, J.W.; Moore, P.E.


    Fire managers and researchers need information on fuel deposition rates to estimate future changes in fuel bed characteristics, determine when forests transition to another fire behavior fuel model, estimate future changes in fuel bed characteristics, and parameterize and validate ecosystem process models. This information is lacking for many ecosystems including the Sierra Nevada in California, USA. We investigated fuel deposition rates and stand characteristics of seven montane and four subalpine conifers in the Sierra Nevada. We collected foliage, miscellaneous bark and crown fragments, cones, and woody fuel classes from four replicate plots each in four stem diameter size classes for each species, for a total of 176 sampling sites. We used these data to develop predictive equations for each fuel class and diameter size class of each species based on stem and crown characteristics. There were consistent species and diameter class differences in the annual amount of foliage and fragments deposited. Foliage deposition rates ranged from just over 50 g m-2 year-1 in small diameter mountain hemlock stands to ???300 g m-2 year-1 for the three largest diameter classes of giant sequoia. The deposition rate for most woody fuel classes increased from the smallest diameter class stands to the largest diameter class stands. Woody fuel deposition rates varied among species as well. The rates for the smallest woody fuels ranged from 0.8 g m-2 year-1 for small diameter stands of Jeffrey pine to 126.9 g m-2 year-1 for very large diameter stands of mountain hemlock. Crown height and live crown ratio were the best predictors of fuel deposition rates for most fuel classes and species. Both characteristics reflect the amount of crown biomass including foliage and woody fuels. Relationships established in this study allow predictions of fuel loads to be made on a stand basis for each of these species under current and possible future conditions. These predictions can be used to

  1. Soil, plant, and transport influences on methane in a subalpine forest under high ultraviolet irradiance

    Directory of Open Access Journals (Sweden)

    D. Baer


    Full Text Available Recent studies have demonstrated direct methane emission from plant foliage under aerobic conditions, particularly under high ultraviolet (UV irradiance. We examined the potential importance of this phenomenon in a high-elevation conifer forest using micrometeorological techniques. Vertical profiles of methane and carbon dioxide in forest air were monitored every 2 h for 6 weeks in summer 2007. Day to day variability in above-canopy CH4 was high, with observed values in the range 1790 to 1910 nmol mol−1. High CH4 was correlated with high carbon monoxide and related to wind direction, consistent with pollutant transport from an urban area by a well-studied mountain-plain wind system. Soils were moderately dry during the study. Vertical gradients of CH4 were small but detectable day and night, both near the ground and within the vegetation canopy. Gradients near the ground were consistent with the forest soil being a net CH4 sink. Using scalar similarity with CO2, the magnitude of the summer soil CH4 sink was estimated at ~1.7 mg CH4 m−2 h−1, which is similar to other temperate forest upland soils. The high-elevation forest was naturally exposed to high UV irradiance under clear sky conditions, with observed peak UVB irradiance >2 W m−2. Gradients and means of CO2 within the canopy under daytime conditions showed net uptake of CO2 due to photosynthetic drawdown as expected. No evidence was found for a significant foliar CH4 source in the vegetation canopy, even under high UV conditions. While the possibility of a weak foliar source cannot be excluded given the observed soil sink, overall this subalpine forest was a net sink for atmospheric methane during the growing season.

  2. Ice duration, winter stratification, and mixing behavior of subalpine lakes in western Maine (United States)

    Daly, J.; Adams, S.; Abrams, R.; Engel, B.


    The timing and duration of both winter and summer periods of stratification periods is not well known for subalpine ponds in the northeastern United States. The remote nature of many of these lakes precludes detailed manual monitoring during the winter and the visual identification of major ice phenological events. These lakes are associated with ecological niches at or near their local elevation maximum that may be at risk due to climate change; historic ice-out records for larger regional lakes indicates a significant trend toward earlier ice-out in response to climate warming. We are using low-cost data loggers to develop high-resolution records characterizing water temperature variability at multiple depths in fifteen lakes 600 - 1000 m elevation. These lakes are located along a 175 km transect in western Maine; most are along the Appalachian Trail. The loggers are recording sub-hourly water temperature and light at the surface, two meters depth, and the bottom of each lake. The timing and duration of winter stratification and ice cover on these lakes are determined by distinctive temperature patterns recorded by the data loggers. The onset of winter stratification is marked by the expected temperature inversion and a slowly increasing hypolimnetic temperature; ice-on follows and is represented by the transition from daily heating cycles to a steady temperature over several days as the ice freezes in around the near-surface logger and energy loss is accomplished through a phase change rather than a drop in temperature. Once the ice is established, temperatures recorded by the near-surface logger vary daily, and are frequently below freezing in response to changing air temperature. Evaluation of winter 2009-2011 records shows that the duration of winter stratification exceeds ice duration at nearly every site. The timing of the onset of stratification is nearly uniform across the study area within each year of data, suggesting a nearly simultaneous response to

  3. Performance Evaluation of FIR Filter After Implementation on Different FPGA and SOC and Its Utilization in Communication and Network

    DEFF Research Database (Denmark)

    Pandey, Bishwajeet; Das, Bhagwan; Kaur, Amanpreet


    There are many areas of communication and network, which have open scope to use FIR filter. Therefore, energy efficient FIR filter will increase lifetime of network and FIR filter with less delay and latency will increase performance of network. In this work, we are going to design an FIR filter ...... provide by Xilinx. We transform that C code into HDL using Vivado HLS 2016.2 before power analysis on Vivado 2016.2. Ultrascale FPGA is generally used for packet processing in 100G networking and heterogeneous wireless infrastructure....

  4. [Effects of snow pack on soil nitrogen transformation enzyme activities in a subalpine Abies faxioniana forest of western Sichuan, China]. (United States)

    Xiong, Li; Xu, Zhen-Feng; Wu, Fu-Zhong; Yang, Wan-Qin; Yin, Rui; Li, Zhi-Ping; Gou, Xiao-Lin; Tang, Shi-Shan


    This study characterized the dynamics of the activities of urease, nitrate reductase and nitrite reductase in both soil organic layer and mineral soil layer under three depths of snow pack (deep snowpack, moderate snowpack and shallow snowpack) over the three critical periods (snow formed period, snow stable period, and snow melt period) in the subalpine Abies faxoniana forest of western Sichuan in the winter of 2012 and 2013. Throughout the winter, soil temperature under deep snowpack increased by 46.2% and 26.2%, respectively in comparison with moderate snowpack and shallow snowpack. In general, the three nitrogen-related soil enzyme activities under shallow snowpack were 0.8 to 3.9 times of those under deep snowpack during the winter. In the beginning and thawing periods of seasonal snow pack, shallow snowpack significantly increased the activities of urease, nitrate and nitrite reductase enzyme in both soil organic layer and mineral soil layer. Although the activities of the studied enzymes in soil organic layer and mineral soil layer were observed to be higher than those under deep- and moderate snowpacks in deep winter, no significant difference was found under the three snow packs. Meanwhile, the effects of snowpack on the activities of the measured enzymes were related with season, soil layer and enzyme type. Significant variations of the activities of nitrogen-related enzymes were found in three critical periods over the winter, and the three measured soil enzymes were significantly higher in organic layer than in mineral layer. In addition, the activities of the three measured soil enzymes were closely related with temperature and moisture in soils. In conclusion, the decrease of snow pack induced by winter warming might increase the activities of soil enzymes related with nitrogen transformation and further stimulate the process of wintertime nitrogen transformation in soils of the subalpine forest.

  5. Characterization of Atmospheric Nitrate Dynamics in a Sub-Alpine Watershed Using Δ17O and δ15N (United States)

    Bourgeois, I.; Savarino, J. P.; Clement, J. C.


    Remote subalpine ecosystems are usually characterized by nutrient-poor soils (Körner, 2004; Seastedt et al., 2004), making them particularly susceptible to undergo changes due to increased atmospheric N deposition (Vitousek et al., 1997; Preunkert et al., 2003). Using Δ17O, a conserved tracer of atmospheric nitrate (NO3 atm) (Michalski et al., 2004; Tsunogai et al., 2010), and δ15N, indicator of NO3 biological sources (Kendall, 1998; Casciotti et al., 2009), we measured the seasonal variations of NO3 atm stable isotopic composition and concentration in several streams and soils originating from two sub-alpine watersheds in the French Alps. Our objective was to investigate whether or not NO3 atm impacts the soil N biogeochemical cycle by increasing nutrients availability for plants and bacteria. We coupled streams and soils measurements with snow-pits sampling and aerosols collection at the Lautaret Pass, to better emphasize the correlation between atmospheric deposition, soil retention and watersheds effluents response. Our results reveal that different temporal dynamics govern our study site: stream measurements show that in spring, snowmelt results in a NO3 atm impulse, accounting for ca. 31 % of the total stream NO3 budget; on the opposite in autumn, NO3 atm accounts only for ca. 3 % of the total stream NO3 budget, highlighting the presence of a NO3 bacterial pool (nitrification). We also inferred from the observed Δ17O variations two distinct phenomena in the spring/summer season: a fast snow run-off and a slower snow-water percolation. The later is believed to affect most the soil N cycle as it directly increases available NO3. Measured soil leachates and extracts confirm this hypothesis and point out the potential importance of anthropogenic N deposition as on average 7 to 10 % of the soil solutions NO3 derives directly from the atmosphere.

  6. Pheno-anomalies of sub-alpine Vaccinium heaths in response to climatic variations (United States)

    Puppi, Giovanna; Monti, Alessandra; Bonafede, Fausto; Vignodelli, Michele; Zanotti, Anna Letizia


    reduction in seed production and could have therefore be disadvantaged in turn-over. This hypothesis is consistent with the results on vegetation changes: in fact, the comparison of the actual vegetation with the historical observations in the same sites, shows a reduction of herb (Hemicryptophytes) diversity and cover in time. It is noteworthy that many of the declining species flower in the driest and hottest weeks of the year. In a climate-warming scenario, the low extension of these sub-alpine islands of the Apennines leads to a high extinction risk of the most sensible species. So, the monitoring of this vulnerable vegetation type seems necessary in order to detect the current trends and should be continued in the future. Puppi and Speranza 1980, Arch. Bot. Biogeogr. Ital. 56(3/4) Puppi et al. 1994, Fitosociologia 26: 63-79

  7. Impact of land use change on soil organic matter dynamics in subalpine grassland (United States)

    Meyer, Stefanie; Leifeld, Jens; Bahn, Michael; Fuhrer, Jürg


    Information regarding the response of soil organic matter (SOM) in soils to past and expected future land use changes in the European Alps is scarce. Understanding this response requires knowledge of size and residence times of SOM fractions with distinct stabilities. In order to quantify differences between types of land use in the amount, distribution and turnover rates of soil organic carbon (SOC) in subalpine grassland soils, we used soil aggregate and SOM density fractionation in combination with 14C dating. Samples were taken along gradients of different types of land use from meadow (M) to pasture (P) and to abandoned grassland (A) in the Stubai Valley and in the Matsch Valley. Sampling sites in both areas were located at equal altitude (1880 m and 1820 m, respectively) with the same parent material and soil type, but the Matsch Valley receives 400-500 mm less annual rainfall. SOC stocks in the top 10 cm were 2.47 ± 0.32 (M), 2.75 ± 0.32 (P), and 2.50 ± 0.31 kg C/m2 (A) in the Stubai Valley and 2.25 ± 0.14 (M), 3.45 ± 0.22 (P), 3.16 ± 0.27 kg C/m2(A) in the Matsch Valley. Three aggregate size classes were separated by wet sieving: 2 mm. The light floating fraction (wPOM, ρ >1 g/cm3) was included in the analysis. Free (f-) and occluded particulate organic matter (oPOM) were isolated from each aggregate size class (ρ >1.6 g/cm3). At both locations, more than 80% of SOC was stored in small (0.25-2 mm) and large (>2 mm) macroaggregates, but no trend in relation to the different types of land use could be detected. The fraction of C in fPOM and in oPOM in all aggregate size classes was highest for soil from abandoned grasslands. The bulk soil of the abandoned site in the Stubai Valley showed a significantly higher share of fPOM-C and oPOM-C and a higher amount of wPOM-C as compared to the soil from managed grassland, whereas in the Matsch Valley pasture soil had a significantly higher wPOM-C content. At both sites, 13C natural abundance analyses revealed

  8. Autotrophic and Heterotrophic Controls over Winter Soil Carbon Cycling in a Subalpine Forest Ecosystem (United States)

    Monson, R. K.; Scott-Denton, L. E.; Lipson, D. A.; Weintrub, M. N.; Rosenstiel, T. N.; Schmidt, S. K.; Williams, M. W.; Burns, S. P.; Delany, A. E.; Turnipseed, A. A.


    that interannual variation in winter ecosystem respiration is positively correlated to interannual variation in the spring snow depth. Years with a with a deeper spring snow pack exhibited higher soil temperatures, and concomitantly higher soil respiration rates. Given the recently reported decadal-scale trend in decreasing snow pack in the Western U.S., which is coupled to warm climate anomalies, our observations indicate the potential for higher wintertime soil carbon sequestration due to lower winter ecosystem respiration rates in subalpine forests. Our studies of processes beneath the winter snow pack demonstrate that contrary to previous assumptions, winter biogeochemical processing of soil organic matter is an important component of ecosystem carbon budgets. Despite low temperatures and an inactive plant rhizosphere, winter microbial communities and exoenzymes appear to be active, carbon substrates appear to be in relatively high abundance and soil respiration rates appear to be sensitive to seasonal and interannual winter climate variability.

  9. Use of passive UAS imaging to measure biophysical parameters in a southern Rocky Mountain subalpine forest (United States)

    Caldwell, M. K.; Sloan, J.; Mladinich, C. S.; Wessman, C. A.


    Unmanned Aerial Systems (UAS) can provide detailed, fine spatial resolution imagery for ecological uses not otherwise obtainable through standard methods. The use of UAS imagery for ecology is a rapidly -evolving field, where the study of forest landscape ecology can be augmented using UAS imagery to scale and validate biophysical data from field measurements to spaceborne observations. High resolution imagery provided by UAS (30 cm2 pixels) offers detailed canopy cover and forest structure data in a time efficient and inexpensive manner. Using a GoPro Hero2 (2 mm focal length) camera mounted in the nose cone of a Raven unmanned system, we collected aerial and thermal data monthly during the summer 2013, over two subalpine forests in the Southern Rocky Mountains in Colorado. These forests are dominated by lodgepole pine (Pinus ponderosae) and have experienced insect-driven (primarily mountain pine beetle; MPB, Dendroctonus ponderosae) mortality. Objectives of this study include observations of forest health variables such as canopy water content (CWC) from thermal imagery and leaf area index (LAI), biomass and forest productivity from the Normalized Difference Vegetation Index (NDVI) from UAS imagery. Observations were, validated with ground measurements. Images were processed using a combination of AgiSoft Photoscan professional software and ENVI remote imaging software. We utilized the software Leaf Area Index Calculator (LAIC) developed by Córcoles et al. (2013) for calculating LAI from digital images and modified to conform to leaf area of needle-leaf trees as in Chen and Cihlar (1996) . LAIC uses a K-means cluster analysis to decipher the RGB levels for each pixel and distinguish between green aboveground vegetation and other materials, and project leaf area per unit of ground surface area (i.e. half total needle surface area per unit area). Preliminary LAIC UAS data shows summer average LAI was 3.8 in the most dense forest stands and 2.95 in less dense

  10. Fast Adaptive Blind MMSE Equalizer for Multichannel FIR Systems

    Directory of Open Access Journals (Sweden)

    Abed-Meraim Karim


    Full Text Available We propose a new blind minimum mean square error (MMSE equalization algorithm of noisy multichannel finite impulse response (FIR systems, that relies only on second-order statistics. The proposed algorithm offers two important advantages: a low computational complexity and a relative robustness against channel order overestimation errors. Exploiting the fact that the columns of the equalizer matrix filter belong both to the signal subspace and to the kernel of truncated data covariance matrix, the proposed algorithm achieves blindly a direct estimation of the zero-delay MMSE equalizer parameters. We develop a two-step procedure to further improve the performance gain and control the equalization delay. An efficient fast adaptive implementation of our equalizer, based on the projection approximation and the shift invariance property of temporal data covariance matrix, is proposed for reducing the computational complexity from to , where is the number of emitted signals, the data vector length, and the dimension of the signal subspace. We then derive a statistical performance analysis to compare the equalization performance with that of the optimal MMSE equalizer. Finally, simulation results are provided to illustrate the effectiveness of the proposed blind equalization algorithm.

  11. FIR Filter Sharpening by Frequency Masking and Pipelining-Interleaving Technique

    Directory of Open Access Journals (Sweden)

    CIRIC, M. P.


    Full Text Available This paper focuses on the improvements of digital filters with a highly sharp transition zone on the Xilinx FPGA chips by combining a sharpening method based on the amplitude change function and frequency masking and PI (Pipelining-Interleaving techniques. A linear phase requires digital filter realizations with Finite Impulse Response (FIR filters. On the other hand, a drawback of FIR filters applications is a low computational efficiency, especially in applications such as filter sharpening techniques, because this technique uses processing the data by repeated passes through the same filter. Computational efficiency of FIR filters can be significantly improved by using some of the multirate techniques, and such a degree of computation savings cannot be achieved in multirate implementations of IIR (Infinite Impulse Response filters. This paper shows the realization of a filter sharpening method with FIR filters combined with frequency masking and PI (Pipelining-Interleaving technique in order to effectively realize the filter with improved characteristic. This realization at the same time keeps the good features of FIR filters such as the linear phase characteristic.

  12. Repeated manual release in a young plantation: effect on Douglas-fir seedlings, hardwoods, shrubs, forbs, and grasses. (United States)

    Philip M. McDonald; Gary O. Fiddler; Henry Harrison


    Douglas-fir seedlings on the Arcata Resource Area, Bureau of Land Management, U.S. Department of the Interior, in central coastal California, were released by chain sawing and grubbing competing vegetation around them at different frequencies (0, 2, and 3 grubbings) over a 5-year period. After 5 years, average Douglas-fir stem diameter (measured at 12 inches above mean...

  13. Genetic diversity and seed production in Santa Lucia fir (Abies bracteata),a relict of the Miocene broadleaved evergreen forest (United States)

    F. Thomas Ledig; Paul D. Hodgskiss; David R. Johnson


    Santa Lucia fir (Abies bracteata), is a unique fir, the sole member of the subgenus Pseudotorreya. It is a relict of the Miocene broadleaved evergreen sclerophyll forest, and is now restricted to a highly fragmented range in the Santa Lucia Mountains of central coastal California. Expected heterozygosity for 30 isozyme loci in 18 enzyme systems...

  14. Douglas-fir ectomycorrhizae in 40- and 400-year-old stands: mycobiont availability to late successional western hemlock. (United States)

    T. R. Horton; R. Molina; K. Hood


    We investigated ectomycorrhizal (EM) fungi in forest stands containing both early successional Douglas-fir and late successional western hemlock at two points in the typical stand development by identifying EM fungi from roots of Douglas-fir and western hemlock in mixed stands. Tn an early seral stage forest, EM roots of western hemlock seedlings and intermingling 40-...

  15. Flight periodicity of the Douglas-fir beetle, Dendroctonus pseudotsugae Hopkins (Coleoptera: Curculionidae: Scolytinae) in Colorado, U.S.A (United States)

    Jose F. Negron; Willis C. Schaupp; Lee Pederson


    There are about 500 species of bark beetles (Coleoptera: Curculionidae: Scolytinae) in the United States (Wood 1982). A number of them are important disturbance agents in forested ecosystems, occasionally creating large tracts of dead trees. One eruptive species is the Douglas-fir beetle, Dendroctonus pseudotsugae Hopkins, which utilizes Douglas-fir, Pseudotsuga...

  16. An examination of the genetic control of Douglas-fir vascular tissue phytochemicals: implications for black bear foraging. (United States)

    Bruce A. Kimball; G.R. Johnson; Dale L. Nolte; Doreen L. Griffin


    Silvicultural practices can influence black bear (Ursus americanus) foraging preferences for Douglas-fir (Pseudotsuga menziesii) cambial-zone vascular tissues, but little is known about the role of genetics. To study the impact of genetic selection, vascular tissue samples were collected from Douglas-fir trees in six half-sib families from five...

  17. Changes in wood product proportions in the Douglas-fir region with respect to size, age, and time. (United States)

    R.A. Monserud; X. Zhou


    We examine both the variation and the changing proportions of different wood products obtained from trees and logs in the Douglas-fir region of the Northwestern United States. Analyses are based on a large product recovery database covering over 40 years of recovery studies; 13 studies are available for Douglas-fir (Pseudotsuga menziesii (Mirb.)...

  18. Morphological defects in native Japanese fir trees around the Fukushima Daiichi Nuclear Power Plant. (United States)

    Watanabe, Yoshito; Ichikawa, San'ei; Kubota, Masahide; Hoshino, Junko; Kubota, Yoshihisa; Maruyama, Kouichi; Fuma, Shoichi; Kawaguchi, Isao; Yoschenko, Vasyl I; Yoshida, Satoshi


    After the accident at the Fukushima Daiichi Nuclear Power Plant (F1NPP) in March 2011, much attention has been paid to the biological consequences of the released radionuclides into the surrounding area. We investigated the morphological changes in Japanese fir, a Japanese endemic native conifer, at locations near the F1NPP. Japanese fir populations near the F1NPP showed a significantly increased number of morphological defects, involving deletions of leader shoots of the main axis, compared to a control population far from the F1NPP. The frequency of the defects corresponded to the radioactive contamination levels of the observation sites. A significant increase in deletions of the leader shoots became apparent in those that elongated after the spring of 2012, a year after the accident. These results suggest possibility that the contamination by radionuclides contributed to the morphological defects in Japanese fir trees in the area near the F1NPP.

  19. Effect of stand edge on the natural regeneration of spruce, beech and Douglas-fir

    Directory of Open Access Journals (Sweden)

    Lumír Dobrovolný


    Full Text Available Our work aimed at studying the strategy of woody plants regeneration during the regeneration of a spruce stand with the admixture of beech and Douglas-fir by border cutting (NW-SE aspect on acidic sites of higher elevations in the Bohemian-Moravian Upland. Spruce is better adapted to bear shade than Douglas-fir. Nevertheless, in optimal light conditions up to a distance of ca. 35 m (about 16% DIFFSF from the stand edge, the Douglas-fir can put the spruce into danger as to height growth. By contrast to beech, the density of spruce is significantly higher within the distance of 45 m (about 15% DIFFSF from the stand edge but further on the situation would change to the benefit of beech. The density of Douglas-fir significantly dominates over beech within a distance of 35 m from the stand edge; from 55 m (less than 15% DIFFSF, the situation changes in favour of beech. Beech can survive in full shade deep in the stand core waiting for its opportunity to come. As compared to spruce and Douglas-fir, the height growth of beech was at all times significantly greater at a distance of 25 m from the stand edge. Converted to practical conditions, spruce and Douglas-fir with individually admixed beech seedlings showed good prosperity approximately up to a distance of one stand height from the edge. A mixture of spruce and beech did well at a greater distance but good prosperity at a distance of 2–3 stand heights was shown only by beech. Thus, border regeneration eliminates disadvantages of the climatic extremes of clear-cutting and specifics of shelterwood felling during which one – usually shade-tolerant tree species dominates in the natural regeneration (e.g. beech.

  20. Climate Change, Wildfires and Fir Forests in Greece: Perceptions of Forest Managers

    Directory of Open Access Journals (Sweden)

    Ioannis Mitsopoulos


    Full Text Available Background and Purpose: The potential impact that climate change may have on fire regime in ecosystems that are not fire-dependent emerges from fires that are nowadays spreading over higher altitudes and northern latitudes. The effects of fire occurrence in high elevation forests of Greece became apparent during the last few years when a number of large forest fires burned a significant number of high altitude fir ecosystems. This paper describes a study that investigated perceptions of wildfire risk to fir ecosystems of Greece in the context of climate change among a sample of Greek forestry experts by conducting personal interviews. Materials and Methods: A total of 63 forest experts answered the developed survey from 43 different forest management units covering all forestry agencies which manage fir forests in the country. The perceived fire risk and management to fir forest ecosystems was assessed by means of a subset of scales previously identified as relevant to climate change and to all fire management aspects, such as fire prevention, fire suppression and post fire treatments. Results: Increased fire risk was rated as a quite important issue in fir forests during the last few years, while the adapting options to climate change are not implemented mainly due to budget and personnel constrains. According to forest experts, fire prevention in fir forests under climate change should be focused on public awareness and fuel management. Nevertheless, the survey results indicate that there is also a need for specific fire prevention measures such as the type of logging activities and other technical measures. The result of this survey highlighted the need for the establishment of a new firefighting organization and for the better coordination of all involved parties during fire suppression. Conclusions: The findings of the current study can be critically important in determining how forest fire management considerations are incorporated into

  1. Damage by Neonectria neomacrospora and Adelges piceae in provenance trials of subalpine fir (Abies lasiocarpa) in Denmark

    DEFF Research Database (Denmark)

    Skúlason, Brynjar; Hansen, Ole Kim; Thomsen, Iben Margrete


    Field trials of Abies lasiocarpa were undertaken with the aim of assessing the potential for Christmas tree production in Denmark. Twenty-six provenances originating from Alaska to New Mexico were tested. Damage by the insect Adelges piceae and the fungus Neonectria neomacrospora was recorded for...

  2. Modified FIR thermometry for surface temperature sensing by using high power laser. (United States)

    Wang, Ran; Zhang, Xinlu; Zhang, Zhilin; Zhong, Hujiang; Chen, Yujin; Zhao, Enming; Vasilescu, Steven; Liu, Lu


    The FIR (fluorescence intensity ratio) technique for optical thermometry has attracted considerable attention over recent years due to its high sensitivity and high spatial resolution. However, it is thought that a heating effect induced by incident light may lead to temperature overestimations, which in turn impedes the reliability of this technique for applications which require high levels of accuracy. To further improve the FIR technique, this paper presents a modified calibration expression, which is suitable for surface temperature sensing, based on the temperature distribution (calculated through COMSOL software). In addition, this modified method is verified by the experimental data.

  3. Identification and characterization of small non-coding RNAs from Chinese fir by high throughput sequencing

    Directory of Open Access Journals (Sweden)

    Wan Li-Chuan


    Full Text Available Abstract Background Small non-coding RNAs (sRNAs play key roles in plant development, growth and responses to biotic and abiotic stresses. At least four classes of sRNAs have been well characterized in plants, including repeat-associated siRNAs (rasiRNAs, microRNAs (miRNAs, trans-acting siRNAs (tasiRNAs and natural antisense transcript-derived siRNAs. Chinese fir (Cunninghamia lanceolata is one of the most important coniferous evergreen tree species in China. No sRNA from Chinese fir has been described to date. Results To obtain sRNAs in Chinese fir, we sequenced a sRNA library generated from seeds, seedlings, leaves, stems and calli, using Illumina high throughput sequencing technology. A comprehensive set of sRNAs were acquired, including conserved and novel miRNAs, rasiRNAs and tasiRNAs. With BLASTN and MIREAP we identified a total of 115 conserved miRNAs comprising 40 miRNA families and one novel miRNA with precursor sequence. The expressions of 16 conserved and one novel miRNAs and one tasiRNA were detected by RT-PCR. Utilizing real time RT-PCR, we revealed that four conserved and one novel miRNAs displayed developmental stage-specific expression patterns in Chinese fir. In addition, 209 unigenes were predicted to be targets of 30 Chinese fir miRNA families, of which five target genes were experimentally verified by 5' RACE, including a squamosa promoter-binding protein gene, a pentatricopeptide (PPR repeat-containing protein gene, a BolA-like family protein gene, AGO1 and a gene of unknown function. We also demonstrated that the DCL3-dependent rasiRNA biogenesis pathway, which had been considered absent in conifers, existed in Chinese fir. Furthermore, the miR390-TAS3-ARF regulatory pathway was elucidated. Conclusions We unveiled a complex population of sRNAs in Chinese fir through high throughput sequencing. This provides an insight into the composition and function of sRNAs in Chinese fir and sheds new light on land plant sRNA evolution.

  4. [Dynamic changes of topsoil organic carbon in subalpine spruce plantation at different succession stages in western Sichuan Province]. (United States)

    Jiang, Fa-Yan; Sun, Hui; Lin, Bo; Liu, Qing


    Soil samples at 0-30 cm depth were collected from the primary spruce (Picea asperata) forest and its plantations at different succession stages (22-, 47-, and 65-year-old) in subalpine zone of western Sichuan Province to study the dynamic changes of soil organic carbon stock and labile organic carbon content. The soil total organic carbon (TOC) stocks in 0-10, 10-20, and 20-30 cm soil layers decreased from 95.87, 79.08, and 71.55 t x hm(-2) in 22-year-old plantation to 56.12, 34.75, and 31.06 t x hm(-2) in 65-year-old plantation, respectively, and the TOC stocks in these soil layers in 47 and 65-year-old plantations were less than those (88.08, 71.16 and 64.81 t x hm(-2), respectively) in primary forest. The easily oxidizable organic carbon (EOC) contents in 0-10, 10-20, and 20-30 cm soil layers decreased from 35.89, 26.91, and 26.00 g x kg(-1) in primary forest to 20.25, 14.50, and 12.36 g x kg L(-1) in 65-year-old plantation, the microbial biomass carbon (MBC) contents decreased from 524.44, 273.26, and 257.97 mg x kg(-1) in primary forest to 312.41, 186.95, and 152.18 mg x kg(-1) in 65-year-old plantation, and the particulate organic carbon (POC) contents decreased from 40.23, 27.10, and 19.55 g x kg(-1) in primary forest to 12.33, 7.31, and 5.32 g x kg(-1) in 65-year-old plantation, respectively. The results suggested that within the long succession period of primary P. asperata forest to its plantations in the subalpine zone of western Sichuan Province, soil TOC and labile organic carbon were in the state of net consumption.

  5. Subalpine Conifer Seedling Demographics: Species Responses to Climate Manipulations Across an Elevational Gradient at Niwot Ridge, Colorado (United States)

    Castanha, C.; Germino, M. J.; Torn, M. S.; Ferrenberg, S.; Harte, J.; Kueppers, L. M.


    The effect of climate change on future ranges of treeline species is poorly understood. For example, it is not known whether trees will recruit into the alpine, above the current treeline, and whether population-level differences in trees will mediate range shifts. At Niwot Ridge, Colorado, we used common gardens and climate manipulations to test predictions that warming will lead to greater recruitment at and beyond the cold edge of these species ranges, and will reduce recruitment at the warm edge. Seed from local populations of limber pine and Englemann spruce was harvested and reciprocally planted in 3 experimental sites spanning an elevation gradient from lower subalpine forest (10,000’), to the upper subalpine treeline ecotone (11,000’), to the alpine tundra (11,300’). In Fall 2009 seeds were sown into 20 plots at each site. Overhead infrared heaters targeted increases in growing season surface soil temperature of 4-5°C. The heating treatment, which began in October 2009, was crossed with manual watering, which was initiated following snowmelt in 2010. Over the 2010 growing season, we surveyed seedling germination and mortality weekly. Germination began in early May at the forest site, in early June at the krummholz site, and in early July at the alpine site. Depending on the site and plot, heating accelerated germination by 1 to 4 weeks. Seed source elevation, species, and site all affected germination, with effects for the two species also depending on site. At all sites, lower elevation, warm-edge populations had higher germination rates than high-elevation, cool-edge populations, indicating a potential bottleneck for germination of the high elevation seed sources in the adjacent alpine tundra. At all sites, survival was generally higher for pine than for spruce. Watering tended to enhance pine germinant survival while heating tended to depress spruce germinant survival. Our results indicate that the alpine tundra, generally considered an

  6. Genetics Home Reference: Camurati-Engelmann disease (United States)

    ... 68. Review. Citation on PubMed Janssens K, Vanhoenacker F, Bonduelle M, Verbruggen L, Van Maldergem L, Ralston S, Guañabens N, Migone N, Wientroub S, Divizia MT, Bergmann C, Bennett C, Simsek S, Melançon S, Cundy T, ...

  7. Early warning system for Douglas-fir tussock moth outbreaks in the Western United States. (United States)

    Gary E. Daterman; John M. Wenz; Katharine A. Sheehan


    The Early Warning System is a pheromone-based trapping system used to detect outbreaks of Douglas-fir tussock moth (DFTM, Orgyia pseudotsugata) in the western United States. Millions of acres are susceptible to DFTM defoliation, but Early Warning System monitoring focuses attention only on the relatively limited areas where outbreaks may be...

  8. Using silviculture to influence carbon sequestration in southern Appalachian spruce-fir forests (United States)

    Patrick T. Moore; R. Justin DeRose; James N. Long; Helga. van Miegroet


    Enhancement of forest growth through silvicultural modification of stand density is one strategy for increasing carbon (C) sequestration. Using the Fire and Fuels Extension of the Forest Vegetation Simulator, the effects of even-aged, uneven-aged and no-action management scenarios on C sequestration in a southern Appalachian red spruce-Fraser fir forest were modeled....

  9. Is long primary growth associated with stem sinuosity in Douglas-fir? (United States)

    Barbara L. Gartner; G.R. Johnson


    Stem sinuosity is a highly visible stem-form trait in the leaders of fast-growing Douglas-fir (Pseudotsuga menziesii var. menziesii (Mirb.) Franco) trees, yet its cause is unknown. We tested the hypotheses that sinuous stems have longer expanses of primary growth than nonsinuous stems (putting the leader at higher risk for...

  10. Lacinipolia Patalis grote (Lepidoptera: Noctuidae) infesting Douglas-fir cones: A new host record (United States)

    Nancy G. Rappaport


    Larvai of Lacinipolia patalis (Grote) (Lepidoptera: Noctuidae) were discovered in Douglas-fir (Pseudotsuga menziessi [Mirb.} Franco) cones collected from the Louisiana-Pacific Corporation's Little River Seed Orchard near Trinidad Head in Humboldt County, CA (elevation 91 m) during the fall of 1985. Previous surveys have not...

  11. EuroFIR quality approach for managing food composition data; where are we in 2014? (United States)

    Westenbrink, Susanne; Roe, Mark; Oseredczuk, Marine; Castanheira, Isabel; Finglas, Paul


    A EuroFIR quality management framework was developed to assure data quality of food composition data, incorporating several recommendations developed or improved during the EuroFIR projects. A flow chart of the compilation process with standard operating procedures to assure critical steps was the starting point. Recommendations for food description, component identification, value documentation, recipe calculation, quality evaluation of values, guidelines to assess analytical methods, document and data repositories and training opportunities were harmonized as elements of the quality framework. European food composition database organizations reached consensus on the EuroFIR quality framework and started implementation. Peer reviews of the European compiler organizations were organized to evaluate the quality framework, focusing on what was achieved and on improvements needed. The reviews demonstrated that European food database compilers have made good use of standards and guidelines produced by EuroFIR, as well as a common understanding that a quality framework is essential to assure food composition data quality. Copyright © 2015 Elsevier Ltd. All rights reserved.

  12. Estimating biomass of shrubs and forbs in central Washington Douglas-fir stands. (United States)

    Craig M. Olson; Robert E. Martin


    Understory plants in closed 70-year-old even-aged Douglas-fir stands in north central Washington were destructively sampled to determine the relationship of ground cover and height to dry weight. Weight of plant material can be estimated from the product of plant height and percentage of ground cover on 50- x 100-centimeter (cm) quadrats. Correlation coefficients for...

  13. Assessing the specific energy consumption and physical properties of comminuted Douglas-fir chips for bioconversion (United States)

    Yalan Liu; Jinwu Wang; Michael P. Wolcott


    Size reduction homogenizes the bulk biomass and facilitates downstream feedstock handling, transportation, and storage. Effects of feeding rate, mill-type (hammer and knife mill), screen size, and moisture content on comminution energy consumption of commercial Douglas-fir (Pseudotsuga menziesii) pulp chips were quantified. The resulting particles...

  14. Stratification period and germination of Douglas-fir seed from Oregon seed orchards: two case studies. (United States)

    Frank C. Sorenson


    Effect of stratification period (S) and incubation temperature (T) on germination be- havior were tested by using two groups of Douglas-fir orchard seedlots that had given low germination percentages in standard tests. One group of seedlots that had experienced different cone-drying regimes, but otherwise were treated comparably, were germinated at T = 15 and 25 °C...

  15. Effect of thinning on form of young-growth Douglas-fir trees. (United States)

    Vern P. Yerkes


    With the advent of increased thinning activity in managed stands of young-growth Douglas-fir (Pseudotsuga menziesii), knowledge of growth and form development of released trees becomes necessary for calculations of total volume and growth. Any appreciable change in the rate of radial increment at various points along the stem of a released tree...

  16. The New Zealand douglas-fir breeding program: proposed adjustments for a changing climate (United States)

    Heidi Dungey; Charlie Low; Mark Miller; Kane Fleet; Alvin D. Yanchuk


    Genetic improvement of Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) in New Zealand was initiated in 1955 with large provenance trials established in the late 1950s. These trials showed that material of Oregon and Californian origin was growing faster than other provenances. Additional collections were made to further evaluate provenance...

  17. The economic significance of mortality in old-growth Douglas-fir management. (United States)

    R.O. McMahon


    Current mortality in the Douglas-fir subregion, exclusive of catastrophic mortality, approximates a billion feet a year. The Forest Service report "Timber Resources for America's Future" recommended "...utilizing a substantial portion of the unsalvaged mortality loss..." as one means of permanently increasing the Nation's timber supply and...

  18. Douglas-fir growth in mountain ecosystems: water limits tree growth from stand to region (United States)

    Jeremy S. Littell; David L. Peterson; Michael Tjoelker


    The purpose of this work is to understand the nature of growth-climate relationships for Douglas-fir (Pseudotsuga menziesii) across the climatic dimensions of its niche. We used a combination of biophysically informed sampling (to identify sample sites) and dendroclimatology (to identify growth-climate relationships) along a climate gradient in...

  19. CO2 uptake by a stand of Douglas fir: flux measurements compared with model calculations

    NARCIS (Netherlands)

    Vermetten, A.W.M.; Ganzeveld, L.; Jeuken, A.; Hofschreuder, P.; Mohren, G.M.J.


    Fluxes of CO2 were calculated by the gradient method from concentration differences, measured in the surface roughness layer above a Douglas fir stand in the Netherlands during a full year (1989). The annual course of the CO2 flux density clearly showed the influence of temperature and incoming

  20. Revised Site Index Curves for Balsam Fir and White Spruce in the Lake States (United States)

    Willard H. Carmean; Jerold T. Hahn


    The original site index curves for balsam fir and white spruce are revised from a breast height age to a total age basis. Site index values from these revised curves are thus comparable to index values for other species that are based upon total tree age. This note also includes formulations for estimating site index by using computers or programmable, hand-...

  1. Growth phenology of coast Douglas-fir seed sources planted in diverse environments (United States)

    Peter J. Gould; Constance A. Harrington; J. Bradley. St. Clair


    The timing of periodic life cycle events in plants (phenology) is an important factor determining how species and populations will react to climate change. We evaluated annual patterns of basal-area and height growth of coast Douglas-fir (Pseudotusuga menziesii var. menziesii (Mirb.) Franco) seedlings from four seed sources...

  2. Fungal endophytes in woody roots of Douglas-fir (Pseudotsuga menziesii) and ponderosa pine (Pinus ponderosa) (United States)

    J. A. Hoff; Ned B. Klopfenstein; Geral I. McDonald; Jonalea R. Tonn; Mee-Sook Kim; Paul J. Zambino; Paul F. Hessburg; J. D. Rodgers; T. L. Peever; L. M. Carris


    The fungal community inhabiting large woody roots of healthy conifers has not been well documented. To provide more information about such communities, a survey was conducted using increment cores from the woody roots of symptomless Douglas-fir (Pseudotsuga menziesii) and ponderosa pine (Pinus ponderosa) growing in dry forests...

  3. Screening Douglas-fir for rapid early growth in common-garden tests in Spain (United States)

    Gabriel Toval Hernandez; Guillermo Vega Alonso; Gonzalo Puerto Arribas; James L. Jenkinson


    Douglas-firs from 91 seed sources in North America were evaluated after 5 and 6 years in 15 common-garden tests in the mountainous regions of northwest and north central Spain. Analyses of tallest trees showed that most of the sources of highest potential for reforestation in Spain are found in regions where the Pacific Ocean air mass dominates climate. Fast growers...

  4. Structural lumber from dense stands of small-diameter Douglas-fir trees. (United States)

    David W. Green; Eini C. Lowell; Roland. Hernandez


    Small-diameter trees growing in overstocked dense stands are often targeted for thinning to reduce fire hazard and improve forest health and ecosystem diversity. In the Pacific Northwest and Intermountain regions, Douglas-fir can be a predominant species in such stands. In this study, mechanical properties and grade yield of structural products were estimated for 2 by...

  5. The Douglas-Fir Genome Sequence Reveals Specialization of the Photosynthetic Apparatus in Pinaceae

    Directory of Open Access Journals (Sweden)

    David B. Neale


    Full Text Available A reference genome sequence for Pseudotsuga menziesii var. menziesii (Mirb. Franco (Coastal Douglas-fir is reported, thus providing a reference sequence for a third genus of the family Pinaceae. The contiguity and quality of the genome assembly far exceeds that of other conifer reference genome sequences (contig N50 = 44,136 bp and scaffold N50 = 340,704 bp. Incremental improvements in sequencing and assembly technologies are in part responsible for the higher quality reference genome, but it may also be due to a slightly lower exact repeat content in Douglas-fir vs. pine and spruce. Comparative genome annotation with angiosperm species reveals gene-family expansion and contraction in Douglas-fir and other conifers which may account for some of the major morphological and physiological differences between the two major plant groups. Notable differences in the size of the NDH-complex gene family and genes underlying the functional basis of shade tolerance/intolerance were observed. This reference genome sequence not only provides an important resource for Douglas-fir breeders and geneticists but also sheds additional light on the evolutionary processes that have led to the divergence of modern angiosperms from the more ancient gymnosperms.

  6. Effect of natural inbreeding on variance structure in tests of wind pollination Douglas-fir progenies. (United States)

    Frank C. Sorensen; T.L. White


    Studies of the mating habits of Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) have shown that wind-pollination families contain a small proportion of very slow-growing natural inbreds.The effect of these very small trees on means, variances, and variance ratios was evaluated for height and diameter in a 16-year-old plantation by...

  7. Genetic variation in response to shade in coastal Douglas-fir. (United States)

    J. Bradley St. Clair; Richard A. Sniezko


    Tree improvement programs have generally relied on testing families in open light environments. With increased interest in multiaged silvicultural systems, some people have questioned whether families selected in the open are appropriate for planting in the shade. We grew Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco var. menziesii...

  8. Growth and nutrition of Douglas fir, Scots pine and pedunculate oak in relation to soil acidification

    NARCIS (Netherlands)

    Visser, de P.H.B.


    In a Douglas fir ( Pseudotsuga menziesii (Mirb.) Franco) and in a Scots pine ( Pinus sylvestris L.) stand on sandy soil in the Netherlands, inputs of water, nutrients and acid loads were changed for four years. Effects of soil changes on growth and

  9. FIBER handbook: a growth model for spruce-fir and northern hardwood types (United States)

    Dale S. Solomon; Richard A. Hosmer; Homer T., Jr. Hayslett; Homer T. Hayslett


    A matrix model, FIBER, has been developed to provide the forest manager with a means of simulating the management and growth of forest stands in the Northeast. Instructional material is presented for the management of even-aged and multi-aged spruce-fir, mixedwood, and northern hardwood stands. FIBER allows the user to simulate a range of silvicultural treatments for a...

  10. Herbs and brush on California red fir regeneration sites: a species and frequency sampling (United States)

    Donald T. Gordon; Eugene E. Bowen


    In a sampling of 32 logged areas in red fir forests of northern California, 58 plant species were found. Species corresponded with less than one-fifth of the 93 species reported in an earlier study of the virgin forest. Sampling of uncut areas adjacent to cuttings does not seem useful for predicting the earlier successional vegetation that will appear when the areas...

  11. Estimating effect of Megastigmus spermotrophus (Hymenoptera: Torymidae) on Douglas-fir seed production: the new paradigm (United States)

    Nancy Rappaport; Alain Roques; Sylvia Mori


    In a pollen exclusion experiment performed on the cones of five Douglas-fir (Pseudotsuga menziesii [Mirbel] Franco) trees, the number of seeds infested by a seed chalcid, Megastigmus spermotrophus Wachtl, did not differ significantly between pollinated and unpollinated cones from the same tree. This finding led to a revision of the...

  12. Genetic variation and population structure in fraser fir (Abies fraseri): a microsatellite assessment of young trees (United States)

    Kevin M. Potter; John Framton; Sedley A. Josserand; C. Dana Nelson


    The island-like populations of Fraser fir (Abies fraseri (Pursh) Poir.) have been isolated since the end of the late-Wisconsinian glaciation on the highest peaks of the Southern Appalachian Mountains and therefore offer an opportunity to investigate the genetic dynamics of a long-fragmented forest tree species. An analysis of eight microsatellite...

  13. Logging damage in thinned, young-growth true fir stands in California and recommendations for prevention. (United States)

    Paul E. Aho; Gary Fiddler; Mike. Srago


    Logging-damage surveys and tree-dissection studies were made in commercially thinned, naturally established young-growth true fir stands in the Lassen National Forest in northern California. Significant damage occurred to residual trees in stands logged by conventional methods. Logging damage was substantially lower in stands thinned using techniques designed to reduce...

  14. Predicting regeneration in the grand fir-cedar-hemlock ecosystem of the northern Rocky Mountains (United States)

    Dennis E. Ferguson; Albert R. Stage; Raymond J. Boyd


    Conifer establishment following regeneration treatments can be predicted in the grand fir-cedar-hemlock ecosystem of the northern Rocky Mountains. Alternative treatments can be evaluated by a model that represents regeneration establishment and early development. This model is designed to be used with the Intermountain Forest and Range Experiment Station's...

  15. Impact of douglas-fir tussock moth... color aerial phtography evaluates mortality (United States)

    Steven L. Wert; Boyd E. Wickman


    Thorough evaluation of insect impact on forest stands is difficult and expensive on the ground. In a study of tree damage following Douglas-fir tussock moth defoliation in Modoc County, California, large-scale (1:1,584)70-mm. color aerial photography was an effective sampling tool and took lesstime and expense than ground methods. Comparison of the photo...

  16. Partial DNA sequencing of Douglas-fir cDNAs used in RFLP mapping (United States)

    K.D. Jermstad; D.L. Bassoni; C.S. Kinlaw; D.B. Neale


    DNA sequences from 87 Douglas-fir (Pseudotsuga menziesii [Mirb.] Franco) cDNA RFLP probes were determined. Sequences were submitted to the GenBank dbEST database and searched for similarity against nucleotide and protein databases using the BLASTn and BLASTx programs. Twenty-one sequences (24%) were assigned putative functions; 18 of which...

  17. Simulation of carbon and water budgets of a Douglas-fir forest

    NARCIS (Netherlands)

    Wijk, van M.T.; Dekker, S.C.; Bouten, W.; Kohsiek, W.; Mohren, G.M.J.


    The forest growth/hydrology model FORGRO–SWIF, consisting of a forest growth and a soil water model, was applied to quantify the inter-annual variability of the carbon and water budgets of a Douglas-fir forest (Pseudotsuga menziessii (Mirb.) Franco) in The Netherlands. With these budgets, the water

  18. Dynamic phenotypic plasticity in photosynthesis and biomass patterns in Douglas-fir seedlings (United States)

    A. C. Koehn; G. I. McDonald; D. L. Turner; D. L. Adams


    As climate changes, understanding the mechanisms long-lived conifers use to adapt becomes more important. Light gradients within a forest stand vary constantly with the changes in climate, and the minimum light required for survival plays a major role in plant community dynamics. This study focuses on the dynamic plasticity of Douglas-fir (Pseudotsuga menziesii var....

  19. Genetic maladaptation of coastal Douglas-fir seedlings to future climates (United States)

    Brad St. Clair;  Glenn T. Howe


    Climates are expected to warm considerably over the next century, resulting in expectations that plant populations will not be adapted to future climates.We estimated the risk of maladaptation of current populations of coastal Douglas-fir (Pseudotsuga menziesii var. menziesii) to future climates as the proportion of nonoverlap between two normal...

  20. Predicting the effects of tropospheric ozone on regional productivity of ponderosa pine and white fir. (United States)

    D.A. Weinstein; J.A. Laurence; W.A. Retzlaff; J.S. Kern; E.H. Lee; W.E. Hogsett; J. Weber


    We simulated forest dynamics of the regional ponderosa pine-white fir conifer forest of the San Bernadino and Sierra Nevada mountains of California to determine the effects of high ozone concentrations over the next century and to compare the responses to our similar study for loblolly pine forests of the southeast. As in the earlier study, we linked two models, TREGRO...

  1. A tool for automatic generation of RTL-level VHDL description of RNS FIR filters

    DEFF Research Database (Denmark)

    Re, Andrea Del; Nannarelli, Alberto; Re, Marco


    Although digital filters based on the Residue Number System (RNS) show high performance and low power dissipation, RNS filters are not widely used in DSP systems, because of the complexity of the algorithms involved. We present a tool to design RNS FIR filters which hides the RNS algorithms...

  2. Estimating the weight of Douglas-fir tree boles and logs with an iterative computer model. (United States)

    Dale R. Waddell; Dale L Weyermann; Michael B. Lambert


    A computer model that estimates the green weights of standing trees was developed and validated for old-growth Douglas-fir. The model calculates the green weight for the entire bole, for the bole to any merchantable top, and for any log length within the bole. The model was validated by estimating the bias and accuracy of an independent subsample selected from the...

  3. Analysis and comparison of nonlinear tree height prediction strategies for Douglas-fir forest. (United States)

    H. Temesgen; V.J. Monleon; D.W. Hann


    Using an extensive Douglas-fir data set from southwest Oregon, we examined the (I) performance and suitability of selected prediction strategies, (2) contribution of relative position and stand-density measures in improving tree height (h) prediction values, and (3) effect of different subsampling designs to fill in missing h values in a new stand using a regional...

  4. Pest management in Douglas-fir seed orchards: a microcomputer decision method (United States)

    James B. Hoy; Michael I. Haverty


    The computer program described provides a Douglas-fir seed orchard manager (user) with a quantitative method for making insect pest management decisions on a desk-top computer. The decision system uses site-specific information such as estimates of seed crop size, insect attack rates, insecticide efficacy and application costs, weather, and crop value. At sites where...

  5. Establishment and growth of native hardwood and conifer seedlings underplanted in thinned Douglas-fir stands. (United States)

    Kathleen G. Maas-Hebner; William H. Emmingham; David L. Larson; Samuel S. Chan


    Five conifers and two hardwoods native to the Pacific Northwest were planted under four overstory densities of 30-year-old plantations of Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) in the Oregon Coast Range, USA. Stand treatments were unthinned (547 trees ha-1), narrow thin (252 trees ha-1),...

  6. Carbon Sequestration in Soils Affected by Douglas Fir Reforestation in Apennines (Northern Italy

    Directory of Open Access Journals (Sweden)

    Giampaolo Di Biase


    Full Text Available Douglas fir reforestation plays an important role in Italian forest because no indigenous conifer has similar characteristics of productivity and timber quality. Few studies on physicochemical properties of soils under Douglas fire are noticeable. The aim of this work is to evaluate the organic C stock into soils under Douglas fir plantation in different selected areas. The areas of study are located in the North Apennine (Italy; Corno alle Scale (COR, Vallombrosa (VAL, Mulino Mengoni (MEN, respectively are chosen for the presence of Douglas fir reforestation of 60 years old. Two soil profiles for each area have been open and described. The pH value decreased along the profile depth. The organic C amount in organic layers was higher in Val and Men pedons than that determined in COR one. Higher amount of organic C were detected in organo-mineral horizons of Co pedons, highlighting a rapid turnover of soil organic matter. The C stock calculated in the first 30 cm of soil showed that the higher C amount is stored in highest altitudes profiles (COR6 and VAL6 than the other. The soil are classified as Lithic Dystrudepts in the highest altitudes (COR 6, 7 and VAL 6, 7, respectively while as Humic Dystrudepts in MEN (4 and 5 pedons. We conclude that no dangerous effects on soil quality of Douglas fir were investigated and they seem to be similar to those of native tree species, even if other different aspects should be investigated.

  7. Effects of water suspension and wet-dry cycling on fertility of Douglas-fir pollen. (United States)

    Donald L. Copes; Nan C. Vance


    Studies were made to determine how long Douglas-fir pollen remains viable after suspension in cool water form 0 to 34 days. Linear regression analysis of in vivo and in vitro tests indicated that filled seed efficiency and pollen viability, respectively, decreased about 3 percent per day. The relation may have been nonlinear the first 6 days, as little decrease...

  8. Characterization of wood strands from young, small-diameter Douglas-fir and western hemlock trees (United States)

    Vikram Yadama; Eini C. Lowell; Christopher E. Langum


    Tensile properties of strands processed from small-diameter Douglas-fir and western hemlock trees grown on the Washington coast were analyzed and effects of location within the tree on properties was examined. Reduction factors for strand properties relative to small, clear solid wood specimen properties were determined by correlating strand properties to previously...

  9. DFSIM with economics: A financial analysis option for the DFSIM Douglas-fir simulator. (United States)

    Roger O. Fight; Judith M. Chittester; Gary W. Clendenen


    A modified version of the DFSIM Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco var. menziesii) growth and yield simulator, DFSIM WITH ECONOMICS, now has an economics option that allows the user to estimate present net worth at the same time a silvicultural regime is simulated. If desired, the economics option will apply a...

  10. Northern flying squirrel mycophagy and truffle production in fir forests in northeastern California (United States)

    J.R. Waters; K.S. McKelvey; C.J. Zabel; D.L. Luoma


    In this paper we summarize the results of four studies in which we either examined the feeding habits of the northern flying squirrel (Glaucomys sabrinus), a mycophagous (consuming fungi) small mammal, or compared the abundance of truffles (sporocarps of hypogeous mycorrhizal fungi) among different types of fir (Abies) forest....

  11. Three different criteria for the design of two-dimensional zero phase FIR digital filters

    DEFF Research Database (Denmark)

    Gislason, E.; Johansen, M.; Conradsen, Knut


    An error criterion for the design of FIR filters is proposed. Filters with relatively many free filter coefficients are designed using the Chebyshev, the weighted-least-squares (WLS), and a new partitioned minimax error criterion, and the performance of the filters is compared. A general and fast...

  12. Silvicultural research and the evolution of forest practices in the Douglas-fir region. (United States)

    Robert O. Curtis; Dean S. DeBell; Richard E. Miller; Michael Newton; J. Bradley St. Clair; William I. Stein


    Silvicultural practices in the Douglas-fir region evolved through a combination of formal research, observation, and practical experience of forest managers and silviculturists, and changing economic and social factors. This process began more than a century ago and still continues. It has had a great influence on the economic well-being of the region and on the...

  13. Ground-based forest harvesting effects on soil physical properties and Douglas-fir growth. (United States)

    Adrian Ares; Thomas A. Terry; Richard E. Miller; Harry W. Anderson; Barry L. Flaming


    Soil properties and forest productivity can be affected by heavy equipment used for harvest and site preparation but these impacts vary greatly with site conditions and operational practices. We assessed the effects of ground-based logging on soil physical properties and subsequent Douglas-fir [Pseudotsuga menziesii (Mirb) Franco] growth on a highly...

  14. Growth of California red fir advance regeneration after overstory removal and thinning (United States)

    William W. Oliver


    Advance regeneration is common under decadent, old-growth stands of California red fir (Abies magnifica A. Murr.). Intense competition for the site's resources can create sapling stands of poor vigor and advanced age. When competition is reduced by overstory removal and thinning, suppressed advance regeneration has been shown to respond with...

  15. Preparation and morphology of porous SiO2 ceramics derived from fir flour templates

    Directory of Open Access Journals (Sweden)



    Full Text Available The preparation of SiO2 ceramics with controllable porous structure from fir flour templates via sol–gel processing was investigated. The specific size the fir flour, which was treated with 20 % NaOH solution, was infiltrated with a low viscous silica sol and subsequently calcined in air, which resulted in the formation of highly porous SiO2 ceramics. X-Ray diffraction (XRD, Fourier transform infrared spectroscopy (FTIR and field emission scanning electron microscopy (FESEM were employed to investigate the microstructure and phase formation during processing as well as of the SiO2 ceramics. N2 adsorption measurements were used to analyze the pore size distributions (PSD of the final ceramics. The results indicated that the surface topography was changed and the proportion of the amorphous material was increased in NaOH-treated fir flour. The final oxide products retained ordered structures of the pores and showed unique pore sizes and distributions with hierarchy on the nanoscale derived from the fir flour.

  16. Overstory response to alternative thinning treatments in young Douglas-fir forests of Western Oregon. (United States)

    Liane R. Davis; Klaus J. Puettmann; Gabriel F. Tucker


    An increase in land dominated by young second-growth Douglas-fir forests in the Pacific Northwest has coincided with heightened concerns over loss of old-growth habitat. In search of options for managing young forests to provide late-successional forest structures, the Young Stand Thinning and Diversity Study was designed to test the effectiveness of modified thinning...

  17. Can a fake fir tell the truth about Swiss needle cast? (paper) ... (United States)

    A key question in dendrochronology to reconstruct forest disturbance history is how to distinguish between the effects of Swiss needle cast (SNC) and other forest disturbance agents (e.g., Arceuthobium spp., Armillaria, Phaseolus schweinitzii, Dendroctonus ponderosae, Dendroctonus pseudotsugae, Choristoneura occidentalis Freeman, Orgyia pseudotsugata McDunnough) on radial stem growth of Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco). SNC impacts physiological processes of carbon and water relations by stomatal occlusion and early needle abscission resulting in a reduction of tree growth with a distinct periodicity, whereas phytophagous pests reduce tree growth by defoliation with epidemics following less regular pseudo-periodicities. Outbreaks of the various forest disturbance agents differ in their magnitude, frequency, and duration. In particular, SNC impacts on Douglas-fir growth display a primary periodicity of 6-30 years and a secondary periodicity of 3-5 years which is unique to the causal fungus Phaeocryptopus gaeumannii (Rhode) Petrak. We use frequency domain analysis of tree-ring chronologies of Douglas-fir to identify the SNC disease cycle and separate the confounding effects of climate and SNC. We demonstrate the dendroecological reconstruction of SNC impacts on ancient Douglas-fir trees dated ~65K radioactive years B.P. from Eddyville, OR that were unearthed by the Oregon Department of Transportation. By the end of the 21st century, climate

  18. Realized gains from block-plot coastal Douglas-fir trials in the northern Oregon Cascades (United States)

    Terrence Z. Ye; Keith J.S. Jayawickrama; J. Bradley. St. Clair


    Realized gains for coastal Douglas-fir (Pseudotsuga menziesii var. menziesii) were evaluated using data collected from 15-year-old trees from five field trials planted in large block plots in the northern Oregon Cascades. Three populations with different genetic levels (elite--high predicted gain; intermediate--moderate predicted gain; and an...

  19. Douglas-fir seedlings exhibit metabolic responses to increased temperature and atmospheric drought.

    Directory of Open Access Journals (Sweden)

    Kirstin Jansen

    Full Text Available In the future, periods of strongly increased temperature in concert with drought (heat waves will have potentially detrimental effects on trees and forests in Central Europe. Norway spruce might be at risk in the future climate of Central Europe. However, Douglas-fir is often discussed as an alternative for the drought and heat sensitive Norway spruce, because some provenances are considered to be well adapted to drier and warmer conditions. In this study, we identified the physiological and growth responses of seedlings from two different Douglas-fir provenances to increased temperature and atmospheric drought during a period of 92 days. We analysed (i plant biomass, (ii carbon stable isotope composition as an indicator for time integrated intrinsic water use efficiency, (iii apparent respiratory carbon isotope fractionation as well as (iv the profile of polar low molecular metabolites. Plant biomass was only slightly affected by increased temperatures and atmospheric drought but the more negative apparent respiratory fractionation indicated a temperature-dependent decrease in the commitment of substrate to the tricarboxylic acid cycle. The metabolite profile revealed that the simulated heat wave induced a switch in stress protecting compounds from proline to polyols. We conclude that metabolic acclimation successfully contributes to maintain functioning and physiological activity in seedlings of both Douglas-fir provenances under conditions that are expected during heat waves (i.e. elevated temperatures and atmospheric drought. Douglas-fir might be a potentially important tree species for forestry in Central Europe under changing climatic conditions.

  20. Winter injury among white fir seedlings...unusual pattern in seed source study (United States)

    M. Thompson Conkle; W. J. Libby; J. L. Hamrick


    White fir seeds collected from 43 sources were sown at the Institute of Forest Genetics, Placerville, Calif. in 1963. Observations made 3 years later showed that seedlings from northern sources sustained more winter injury than did southern origin seedlings. Seedlings from low elevations were less severely damaged than seedlings from higher elevations in the same...

  1. Physiological responses of planting frozen and thawed Douglas-fir seedlings (United States)

    M. Anisul Islam; Kent G. Apostol; Douglass F. Jacobs; R. Kasten Dumroese


    We studied the short-term (7-day) physiological responses of planting thawed and frozen root plugs of Douglas-fir (Pseudotsuga menziesii) seedlings in 2 separate experiments under cool-moist and warm-dry growing conditions, respectively. Our results showed that shoot water potential, root hydraulic conductance, net photosynthesis (A), and...

  2. Training aspects in the use and production of food composition databases. The EuroFIR experience.

    NARCIS (Netherlands)

    Hollman, P.C.H.; Witthöft, C.; Busstra, M.C.; Elburg, L.; Hulshof, P.J.M.


    An increasing number of national food databases have been published on the internet. However, these internet databases can only be searched individually, and the data have been compiled at a national level, resulting in incompatibilities. To resolve these problems, the Network of Excellence EuroFIR

  3. Prediction of wood Quality in Small-Diameter Douglas-Fir using site and Stand Characteristics (United States)

    C.D. Morrow; T.M. Gorman; J.W. Evans; D.E. Kretschmann; C.A. Hatfield


    Standing stress wave measurements were taken on 274 small-diameter Douglas-fir trees in western Montana. Stand, site, and soil measurements collected in the field and remotely through geographical information system (GIS) data layers were used to model dynamic modulus of elasticity (DMOE) in those trees. The best fit linear model developed resulted in an adjusted

  4. Impact of the foliar pathogen Swiss needle cast on wood quality of Douglas-fir. (United States)

    G.R. Johnson; Amy T. Grotta; Barbara L. Gartner; Geoff. Downes


    Many stands of Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) near coastal areas of Oregon and Washington are heavily infected with the foliar pathogen causing Swiss needle cast (SNC) disease, and yet there is very little research on the resulting wood quality. Modulus of elasticity(MOE), modulus of rupture (MOR), microfibril angle (MFA), wood...

  5. Early genetic evaluation of open-pollinated Douglas-fir families (United States)

    Kurt H. Riitters; David A. Perry


    In a test of early genetic evaluation of the growth potential of 14 families of open-pollinated Douglas-fir (Pseudotsuga menziesii) [Mirb.] Franco), measures of growth and phenology of seedligns grown in a coldframe were correlated with height of saplings in evaluation plantations at 9, 12, and 15 years. fifteen-year height was most strongly...

  6. Far infrared ray (FIR) therapy: An effective and oncological safe treatment modality for breast cancer related lymphedema. (United States)

    Li, Ke; Xia, Liang; Liu, Ning Fei; Nicoli, Fabio; Constantinides, Joannis; D'Ambrosia, Christopher; Lazzeri, Davide; Tremp, Mathias; Zhang, Ju Fang; Zhang, Yi Xin


    The incidence of breast cancer related lymphedema is approximately 5%. Far infrared ray (FIR) treatment can potentially reduce fluid volume and extremity circumference as well as the frequency of dermato-lymphangitis (DLA). However, there is no published data on the oncological safety of FIR and the potential for activation of any residual breast cancer cells. The aim of this study is to investigate the safety of far infrared ray (FIR) treatment of postmastectomy lymphedema, clinically and in vitro. Patients who underwent mastectomy more than 5years ago complicated by upper extremity lymphedema for more than 1year were included. The enrolled patients were divided into an FIR treatment group and a control group (conservative treatment using bandage compression). Outcome measures included tumor markers (CA153, CA125), ultrasonography of relevant structures and monitoring for adverse reactions 1year after treatment. For the in vitro part of the study, the effects of FIR on human breast adenocarcinoma cell lines (MCF7, MDA-MB231) compared to the effects of FIR on human dermal fibroblasts as a control were considered. The viability, proliferation, cell cycle and apoptotic statistics of the adenocarcinoma and human dermal fibroblast cell lines were analyzed and compared. Results demonstrated that after treatment with FIR, tumor marker (CA153, CA125) concentrations in both the FIR and control groups were not elevated. There was no statistically significant difference between FIR and control group marker expression (p>0.05). Furthermore, no patients were diagnosed with lymphadenectasis or newly enlarged lymph nodes in these two groups. Importantly, there were no adverse events in either group. The in vitro experiment indicated that FIR radiation does not affect viability, proliferation, cell cycle and apoptosis of fibroblasts, MCF-7 and MDA-MB-231 cells. FIR should be considered as feasible and safe for the treatment of breast cancer related lymphedema patients 5years

  7. Inter-annual climate variability and zooplankton: applying teleconnection indices to two deep subalpine lakes in Italy

    Directory of Open Access Journals (Sweden)

    Marina Manca


    Full Text Available Investigating relation between meteo-climatic indices and between-year variation in Daphnia population density and phenology is crucial for e.g. predicting impact of climate change on lake ecosystem structure and functioning. We tested whether and how two teleconnection indices calculated for the winter period, namely the East Atlantic pattern (EADJF and the Eastern Mediterranean Pattern (EMPDJF were correlated with Daphnia population growth in two Italian subalpine lakes, Garda and Maggiore. We investigated between-lake temporal coherence in: i water temperature within the water layer in which Daphnia is distributed; ii timing of Daphnia initial and spring maximum population density peak and iii the level of Daphnia spring maximum population density peak over an eleven-year period (1998-2008 of unchanged predation pressure by fish and invertebrates, and of common oligotrophy. Between-lake temporal coherence was high for an earlier start, an earlier, and lower, Daphnia population spring density peak after milder winters. Peak density level was coherently, positively correlated with soluble reactive phosphorus (SRP concentration. We hypothesized that Daphnia peak densities were related to atmospheric modes of variability in winter and to the degree of late winter mixing promoting replenishment of algal nutrients into upper water layers and phytoplankton growth, enhancing food availability and Daphnia fecundity, promoting Daphnia peak. 

  8. Equine Grazing in Managed Subalpine Wetlands: Effects on Arthropods and Plant Structure as a Function of Habitat (United States)

    Holmquist, Jeffrey G.; Schmidt-Gengenbach, Jutta; Haultain, Sylvia A.


    Grazing management necessarily emphasizes the most spatially extensive vegetation assemblages, but landscapes are mosaics, often with more mesic vegetation types embedded within a matrix of drier vegetation. Our primary objective was to contrast effects of equine grazing on both subalpine vegetation structure and associated arthropods in a drier reed grass ( Calamagrostis muiriana) dominated habitat versus a wetter, more productive sedge habitat ( Carex utriculata). A second objective was to compare reed grass and sedge as habitats for fauna, irrespective of grazing. All work was done in Sequoia National Park (CA, USA), where detailed, long-term records of stock management were available. We sampled paired grazed and control wet meadows that contained both habitats. There were moderate negative effects of grazing on vegetation, and effects were greater in sedge than in reed grass. Conversely, negative grazing effects on arthropods, albeit limited, were greater in the drier reed grass, possibly due to microhabitat differences. The differing effects on plants and animals as a function of habitat emphasize the importance of considering both flora and fauna, as well as multiple habitat types, when making management decisions. Sedge supported twice the overall arthropod abundance of reed grass as well as greater diversity; hemipteran and dipteran taxa were particularly abundant in sedge. Given the greater grazing effects on sedge vegetation, greater habitat provision for terrestrial arthropods, and value as aquatic arthropod habitat, the wetter sedge assemblage is worthy of additional consideration by managers when planning for grazing and other aspects of land usage.

  9. The effects of winter recreation on alpine and subalpine fauna: a systematic review and meta-analysis.

    Directory of Open Access Journals (Sweden)

    Chloe F Sato

    Full Text Available The ski industry is often perceived as having a negative impact on sensitive alpine and subalpine communities. However, empirical evidence of such impacts is lacking. We reviewed the available literature from the last 35 years to quantify the reported effects of winter recreation on faunal communities. Overall, using one-sample binomial tests ('sign tests' we found that the effects of all types of winter recreation-related disturbances (i.e. ski runs, resort infrastructure and winter tourism were more likely to be negative or have no effect, than be positive for wildlife. More specifically, in Europe, where the majority of the available research was conducted, the impacts of winter recreation were most often negative for fauna. In terms of specific taxa, birds and to a lesser extent mammals and arthropods, responded negatively to disturbance. Results from our meta-analysis confirmed the results from our binomial tests. Richness, abundance and diversity of fauna were lower in areas affected by winter recreation when compared with undisturbed areas. For most regions and taxa, however, empirical evidence remains too limited to identify clear impacts of winter recreation. We therefore conclude that the majority of ski resorts are operating in the absence of knowledge needed to inform effective strategies for biodiversity conservation and ecologically-sound management. Thus, there is an urgent need for more empirical research to be conducted throughout this increasingly threatened ecological community, especially given the indication from the available literature that fauna often respond negatively to winter recreation.

  10. Nutrient status in soil of Ski runs in the sub-alpine belt of Uludag Mountain, Bursa, Turkey. (United States)

    Guleryuz, Gurcan; Kirmizi, Serap; Arslan, Hulya


    Large areas of land are disturbed in sensitive bio-diverse mountain environments by Skiruns. Restoration of vegetation on such disturbed mountain sites may be hampered by soil degradation but the severity and nature of the constraints is not well understood. This study was designed to compare the water holding and nutritional status of soil in three Ski runs which had different construction dates and disturbance levels, and the adjacent undisturbed site in the Abies bommuelleriana forest community in the sub-alpine belt of Uludag Mountain (Bithynian Olympus). The values of soil parameters were depressed in proportion to the disturbance level. Water holding capacity (WHC), total nitrogen (N), organic carbon (C) and calcium (Ca2+), magnesium (Mg2+) and potassium (K+) contents (mg kg(-1) dry weight) of soils in the Ski run which had the highest disturbance level were lower than that of the undisturbed adjacent sites. However the results indicated that the soil parameters were less degraded when secondary vegetation was growing on the disturbed areas.

  11. The Effects of Winter Recreation on Alpine and Subalpine Fauna: A Systematic Review and Meta-Analysis (United States)

    Sato, Chloe F.; Wood, Jeff T.; Lindenmayer, David B.


    The ski industry is often perceived as having a negative impact on sensitive alpine and subalpine communities. However, empirical evidence of such impacts is lacking. We reviewed the available literature from the last 35 years to quantify the reported effects of winter recreation on faunal communities. Overall, using one-sample binomial tests (‘sign tests’) we found that the effects of all types of winter recreation-related disturbances (i.e. ski runs, resort infrastructure and winter tourism) were more likely to be negative or have no effect, than be positive for wildlife. More specifically, in Europe, where the majority of the available research was conducted, the impacts of winter recreation were most often negative for fauna. In terms of specific taxa, birds and to a lesser extent mammals and arthropods, responded negatively to disturbance. Results from our meta-analysis confirmed the results from our binomial tests. Richness, abundance and diversity of fauna were lower in areas affected by winter recreation when compared with undisturbed areas. For most regions and taxa, however, empirical evidence remains too limited to identify clear impacts of winter recreation. We therefore conclude that the majority of ski resorts are operating in the absence of knowledge needed to inform effective strategies for biodiversity conservation and ecologically-sound management. Thus, there is an urgent need for more empirical research to be conducted throughout this increasingly threatened ecological community, especially given the indication from the available literature that fauna often respond negatively to winter recreation. PMID:23691190

  12. Using time scales to characterize phytoplankton assemblages in a deep subalpine lake during the thermal stratification period: Lake Iseo, Italy (United States)

    Marti, Clelia Luisa; Imberger, Jörg; Garibaldi, Letizia; Leoni, Barbara


    A combination of field observations and 3-D hydrodynamic simulations were used to identify the phytoplankton species and to estimate the various time scales of the dominant physical and biological processes in Lake Iseo, a deep subalpine lake located in northern Italy, during a stratified period (July 2010). By ordering the rate processes time scales, we derive a phytoplankton patch categorization and growth interpretation that provides a general framework for the spatial distribution of phytoplankton concentration in Lake Iseo and illuminates the characteristics of their ecological niches. The results show that the diurnal surface layer was well mixed, received strong diurnal radiation, had low phosphorus concentrations and the phytoplankton biomass was sustained by the green alga Sphaerocystis schroeterii. The vertical mixing time scales were much shorter than horizontal mixing time scales causing a depth-uniform chlorophyll a concentration. The horizontal patch scale was determined by horizontal dispersion balancing the phytoplankton growth time scale, dictating the success of the observed green algae. The strongly stratified nutrient-rich metalimnion had mild light conditions and Diatoma elongatum and Planktothrix rubescens made up the largest proportions of the total phytoplankton biomass at the intermediate and deeper metalimnetic layers. The vertical transport time scales were much shorter than horizontal transport and vertical dispersion leading to growth niche for the observed phytoplankton. The study showed that time-scale hierarchy mandates the essential phytoplankton attributes or traits for success in a particular section of the water column and/or water body.

  13. Using functional traits to assess the resistance of subalpine grassland to trampling by mountain biking and hiking. (United States)

    Pickering, Catherine Marina; Barros, Agustina


    Functional traits reflect plant responses to disturbance, including from visitor impacts. The impacts of mountain biking and hiking on functional composition were compared using a common experimental protocol in a subalpine grassland in the Australian Alps. The overlapping cover of all species was recorded two weeks after different intensities of hiking (200 and 500 passes) and mountain biking (none, 25, 75, 200 and 500 passes). Species' functional trait data were combined with their relative cover to calculate community trait weighted means for plant height, leaf area, percentage leaf dry matter content and Specific Leaf Area (SLA). Species such as Poa fawcettiae with larger leaves and SLA but lower dry weight content of leaves were more resistant to use, with differences between bikers and hikers only apparent at the highest levels of use tested. This differs from some vegetation communities in Europe where plants with smaller leaves were more resistant to hiking. More research using functional traits may account for differences in species responses to trampling. Managers of conservation areas used for hiking and biking need to minimise off trail use by both user groups. Copyright © 2015 Elsevier Ltd. All rights reserved.

  14. Virtual disjunct eddy covariance measurements of organic compound fluxes from a subalpine forest using proton transfer reaction mass spectrometry

    Directory of Open Access Journals (Sweden)

    T. G. Karl


    Full Text Available A `virtual' disjunct eddy covariance (vDEC device was tested with field measurements of biogenic VOC fluxes at a subalpine forest site in the Rocky Mountains of the USA. A PTR-MS instrument was used as the VOC sensor. Daily peak emission fluxes of 2-methyl-3-buten-2-ol (MBO, methanol, acetone and acetaldehyde were around 1.5, 1, 0.8 and 0.4 mg m-2 h-1, respectively. High pass filtering due to long sampling lines was investigated in laboratory experiments, and suggested that VOC losses in PTFA lines are generally governed by diffusion laws. Memory effects and surface reactions did not seem to play a dominant role. Model estimates of MBO fluxes compared well with measured fluxes. The results also suggest that latent heat and sensible heat fluxes are reasonably well correlated with VOC fluxes and could be used to predict variations in VOC emissions. The release of MBO, methanol, acetone and acetaldehyde resulted in significant change of tropospheric oxidant levels and a 10--40% increase in ozone levels, as inferred from a photochemical box model. We conclude that vDEC with a PTR-MS instrument is a versatile tool for simultaneous field analysis of multiple VOC fluxes.

  15. Complex terrain alters temperature and moisture limitations of forest soil respiration across a semiarid to subalpine gradient (United States)

    Berryman, Erin Michele; Barnard, H.R.; Adams, H.R.; Burns, M.A.; Gallo, E.; Brooks, P.D.


    Forest soil respiration is a major carbon (C) flux that is characterized by significant variability in space and time. We quantified growing season soil respiration during both a drought year and a nondrought year across a complex landscape to identify how landscape and climate interact to control soil respiration. We asked the following questions: (1) How does soil respiration vary across the catchments due to terrain-induced variability in moisture availability and temperature? (2) Does the relative importance of moisture versus temperature limitation of respiration vary across space and time? And (3) what terrain elements are important for dictating the pattern of soil respiration and its controls? Moisture superseded temperature in explaining watershed respiration patterns, with wetter yet cooler areas higher up and on north facing slopes yielding greater soil respiration than lower and south facing areas. Wetter subalpine forests had reduced moisture limitation in favor of greater seasonal temperature limitation, and the reverse was true for low-elevation semiarid forests. Coincident climate poorly predicted soil respiration in the montane transition zone; however, antecedent precipitation from the prior 10 days provided additional explanatory power. A seasonal trend in respiration remained after accounting for microclimate effects, suggesting that local climate alone may not adequately predict seasonal variability in soil respiration in montane forests. Soil respiration climate controls were more strongly related to topography during the drought year highlighting the importance of landscape complexity in ecosystem response to drought.

  16. The effects of winter recreation on alpine and subalpine fauna: a systematic review and meta-analysis. (United States)

    Sato, Chloe F; Wood, Jeff T; Lindenmayer, David B


    The ski industry is often perceived as having a negative impact on sensitive alpine and subalpine communities. However, empirical evidence of such impacts is lacking. We reviewed the available literature from the last 35 years to quantify the reported effects of winter recreation on faunal communities. Overall, using one-sample binomial tests ('sign tests') we found that the effects of all types of winter recreation-related disturbances (i.e. ski runs, resort infrastructure and winter tourism) were more likely to be negative or have no effect, than be positive for wildlife. More specifically, in Europe, where the majority of the available research was conducted, the impacts of winter recreation were most often negative for fauna. In terms of specific taxa, birds and to a lesser extent mammals and arthropods, responded negatively to disturbance. Results from our meta-analysis confirmed the results from our binomial tests. Richness, abundance and diversity of fauna were lower in areas affected by winter recreation when compared with undisturbed areas. For most regions and taxa, however, empirical evidence remains too limited to identify clear impacts of winter recreation. We therefore conclude that the majority of ski resorts are operating in the absence of knowledge needed to inform effective strategies for biodiversity conservation and ecologically-sound management. Thus, there is an urgent need for more empirical research to be conducted throughout this increasingly threatened ecological community, especially given the indication from the available literature that fauna often respond negatively to winter recreation.

  17. Transcriptome analysis of immature xylem in the Chinese fir at different developmental phases

    Directory of Open Access Journals (Sweden)

    Yunxing Zhang


    Full Text Available Background.Chinese fir [Cunninghamia lanceolata (Lamb. Hook.] is one of the most important native tree species for timber production in southern China. An understanding of overall fast growing stage, stem growth stage and senescence stage cambium transcriptome variation is lacking. We used transcriptome sequencing to identify the repertoire of genes expressed during development of xylem tissue in Chinese fir, aiming to delineate the molecular mechanisms of wood formation. Results. We carried out transcriptome sequencing at three different cultivation ages (7Y, 15Y and 21Y generating 68.71 million reads (13.88 Gbp. A total of 140,486 unigenes with a mean size of 568.64 base pairs (bp were obtained via de novo assembly. Of these, 27,427 unigenes (19.52% were further annotated by comparison to public protein databases. A total of 5,331 (3.79% unigenes were mapped into 118 pathways by searching against the Kyoto Encyclopedia of Genes and Genomes Pathway database (KEGG. Differentially expressed genes (DEG analysis identified 3, 16 and 5,899 DEGs from the comparison of 7Y vs. 15Y, 7Y vs. 21Y and 15Y vs. 21Y, respectively, in the immature xylem tissues, including 2,638 significantly up-regulated and 3,280 significantly down-regulated genes. Besides, five NAC transcription factors, 190 MYB transcription factors, and 34 WRKY transcription factors were identified respectively from Chinese fir transcriptome. Conclusion. Our results revealed the active transcriptional pathways and identified the DEGs at different cultivation phases of Chinese fir wood formation. This transcriptome dataset will aid in understanding and carrying out future studies on the molecular basis of Chinese fir wood formation and contribute to future artificial production and applications.

  18. A census of radio-selected AGNs on the COSMOS field and of their FIR properties (United States)

    Magliocchetti, M.; Popesso, P.; Brusa, M.; Salvato, M.


    We use the new catalogue by Laigle et al. to provide a full census of VLA-COSMOS radio sources. We identify 90 per cent of such sources and sub-divide them into active galactic nuclei (AGNs) and star-forming galaxies on the basis of their radio luminosity. The AGN sample is complete with respect to radio selection at all z ≲ 3.5. Out of 704 AGNs, 272 have a counterpart in the Herschel maps. By exploiting the better statistics of the new sample, we confirm the results of Magliocchetti et al.: the probability for a radio-selected AGN to be detected at far-infrared (FIR) wavelengths is both a function of radio luminosity and redshift, whereby powerful sources are more likely FIR emitters at earlier epochs. Such an emission is due to star-forming processes within the host galaxy. FIR emitters and non-FIR emitters only differentiate in the z ≲ 1 universe. At higher redshifts, they are indistinguishable from each other, as there is no difference between FIR-emitting AGNs and star-forming galaxies. Lastly, we focus on radio AGNs which show AGN emission at other wavelengths. We find that mid-infrared (MIR) emission is mainly associated with ongoing star formation and with sources which are smaller, younger and more radio luminous than the average parent population. X-ray emitters instead preferentially appear in more massive and older galaxies. We can therefore envisage an evolutionary track whereby the first phase of a radio-active AGN and of its host galaxy is associated with MIR emission, while at later stages the source becomes only active at radio wavelengths and possibly also in the X-ray.

  19. Variation in phenology and monoterpene patterns of defoliated and nondefoliated Douglas-fir (Pseudotsuga menziesii var. glauca). (United States)

    Rose-Marie Muzika; Judith Engle; Catherine Parks; Boyd. Wickman


    Foliage was collected from paired Douglas-fir (Pseudotsuga menziesii var. glauca (Beissn.) Franco) trees characterized as either "resistant" or "susceptible" western spruce budworm (Choristoneura occidentalis Freeman (Lepidoptera: Tortricidae) attack. Resistant trees produced more...

  20. Role of Nurse Logs in Forest Expansion at Timberline (United States)

    Johnson, A. C.; Yeakley, A.


    Nurselogs, known to be key sites of forest regeneration in lower elevation temperate forests, may be important sites for seedling establishment at expanding timberline forests. To determine factors associated with seedling establishment and survival on nurselogs at timberline, fourteen sites, located across a precipitation gradient in the Washington North Cascades Mountains, were examined. Site attributes including seedling type and height, disturbance process introducing downed wood, wood decay type, shading, slope gradient, aspect, and temperature and water content of wood and adjacent soil were determined along 60 m long transects. Nurselogs were found at 13 out of 14 sites; sites typically associated with greater than 80% shade and downed wood having a high level of wood decay. Downed wood serving as nurselogs originated from blowdown, snow avalanches, and forest fires. In total, 46 of 136 downed wood pieces observed served as nurselogs. Seedlings on nurselogs included mountain hemlock (Tsuga mertensiana), Pacific silver fir (Abies amabilis), yellow cedar (Chamaecyparis nootkatensis), subalpine fir (Abies lasiocarpa), Engelmann spruce (Picea engelmannii), and western larch (Larix occidentalis). Nurselogs had significantly higher temperatures (p = 0.015) and higher moisture contents (p = 0.019) than the adjacent soil. Per equal volumes weighed, nurselogs had on average of 23.8 g more water than the adjacent soil. Given predictions of climate warming and associated summer drought conditions in Pacific Northwest forests, the moisture provided by nurselogs may be integral for conifer survival and subsequent timberline expansion in some landscapes.

  1. Effects of manganese and manganese-nitrogen applications on growth and nutrition of Douglas-fir seedlings. (United States)

    M. A. Radwan; John S. Shumway; Dean S. DeBell


    Effects of manganese (Mn) on Douglas-fir grown in soil, with and without urea, and in nutrient solution were investigated. In addition, Mn sorption by forest soils was evaluated. Results show that Douglas-fir does not respond to added Mn and is quite tolerant to high Mn levels. Moreover, Mn sorption by soils is high. It is doubtful that Mn toxicity is of practical...

  2. Mercury in forest mushrooms and topsoil from the Yunnan highlands and the subalpine region of the Minya Konka summit in the Eastern Tibetan Plateau. (United States)

    Falandysz, Jerzy; Saba, Martyna; Liu, Hong-Gao; Li, Tao; Wang, Ji-Peng; Wiejak, Anna; Zhang, Ji; Wang, Yuan-Zhong; Zhang, Dan


    This study aimed to investigate and discuss the occurrence and accumulation of mercury in the fruiting bodies of wild-growing fungi (Macromycetes) collected from montane forests in two regions of southwestern China with differences in soil geochemistry, climate and geographical conditions. Fungal mycelia in soils of the subalpine region of the Minya Konka (Gongga Mountain) in Sichuan and in the highlands of Yunnan efficiently accumulated mercury in fruiting bodies (mushrooms). The examined sites in Yunnan with highly mineralized red and yellow soils showed Hg contents ranging from 0.066 to 0.28 mg kg -1 dry biomass (db) which is roughly similar to the results obtained for samples collected from sites with dark soils relatively rich in organic matter from a remote, the subalpine region of Minya Konka. Due to the remoteness of the subalpine section of Minya Konka, as well as its elevation and climate, airborne mercury from long-range transport could be deposited preferentially on the topsoil and the Hg levels determined in soil samples taken beneath the fruiting bodies were up to 0.48 mg kg -1 dry matter. In Yunnan, with polymetallic soils (Circum-Pacific Mercuriferous Belt), Amanita mushrooms showed mercury in caps of fruiting bodies of up to 7.3 mg kg -1 dry biomass. Geogenic Hg from the mercuriferous belt seems to be the overriding source of mercury accumulated in mushrooms foraged in the regions of Yunnan, while long-range atmospheric transport and subsequent deposition are the mercury sources for specimens foraged in the region of Minya Konka.

  3. Diphyllobothrium latum (Cestoda: Diphyllobothriidea in perch (Perca fluviatilis in three sub-alpine lakes: influence of biotic and abiotic factors on prevalence

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    Orlando PETRINI


    Full Text Available In recent years, human diphyllobothriosis has staged a comeback in Swiss, French and Italian sub-alpine regions. The main putative infective source of the causative agent (the tapeworm Diphyllobothrium latum in these areas is perch (Perca fluviatilis. Therefore, the occurrence of D. latum in this fish species was investigated between 2005 and 2008 in the sub-alpine lakes Maggiore, Lugano and Geneva. Prevalence in fish of Lake Maggiore was 14% (n = 880. In Lake Geneva, 5.1% fillets (n = 532 were infected, whereas perch from Lake Lugano were free from the parasite. These results are discussed in relation to previous studies. Data on fish size and weight indicate that infection of perch by D. latum is independent of age and sex. Abiotic factors considered critical for D. latum life cycle (water temperature and oxygen concentration characterize the three basins and were related to their infestation frequencies. The presence of this parasite was most likely favoured by warmer, well oxygenated waters. Previous studies indicate that the lake’s trophic state (i.e. content of total phosphorus influenced the availability of the first intermediate hosts (copepods of some pseudophyllideans. In our study, no correlation was observed between the amount of phosphorus and the number of copepods in populations of zooplankton. Nevertheless, the trophic states of the three lakes seemed to affect the degree of infection in fish. In conclusion, at least in sub-alpine lakes, abiotic factors such as water temperature, oxygenation and trophic state seem to have an influence on maintaining or preventing perch infection with D. latum.

  4. Wood quality and value production in mixed fir-spruce-beech stands: long-term research in the Western Carpathians

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    Petráš Rudolf


    Full Text Available Stem quality and damage was evaluated in mixed spruce-fir-beech stands. Moreover, an assortments structure was determined with their financial value. Results were compared with pure spruce (Picea abies [L.] Karst., fir (Abies alba Mill. and beech (Fagus sylvatica L. stands. Repeated measurements on 31 long-term research plots, stand assortment models, assortment yield models and value yield models were used. Stem quality of fir and spruce was only slightly lower in mixed stands compared to pure stands but beech stem quality was considerably worse in mixed stands. Fir and spruce had slightly lower proportions of better IIIA quality logs and higher proportions of IIIB quality in mixed stands. Beech had worse assortment structure than spruce and fir, in general. Pure beech stands had higher proportions of better I–IIIA quality assortments than mixed stands by 1–7%. Fir and spruce average value production (€ m−3 culminated at about 56 and 62 cm mean diameters. Almost the same value production was found in pure stands. In these stands it culminated at the mean diameter of 58 and 60 cm. Beech produced substantially less value on the same sites. In mixed stands, its value production culminated at the mean diameter of 40 cm. In pure stands, it culminated at the mean diameter of 36 cm. Although the production was found to be similar in both mixed and pure forests, higher damage intensity and less stem quality in mixed forests suggest that the pure forests can be more profitable.

  5. Volatile and Within-Needle Terpene Changes to Douglas-fir Trees Associated With Douglas-fir Beetle (Coleoptera: Curculionidae) Attack. (United States)

    Giunta, A D; Runyon, J B; Jenkins, M J; Teich, M


    Mass attack by tree-killing bark beetles (Curculionidae: Scolytinae) brings about large chemical changes in host trees that can have important ecological consequences. For example, mountain pine beetle (Dendroctonus ponderosae Hopkins) attack increases emission of terpenes by lodgepole pine (Pinus contorta Dougl. ex Loud.), affecting foliage flammability with consequences for wildfires. In this study, we measured chemical changes to Douglas-fir (Pseudotsuga menziesii var. glauca (Mirb.) Franco) foliage in response to attack by Douglas-fir beetles (Dendroctonus pseudotsugae Hopkins) as trees die and crowns transitioned from green/healthy, to green-infested (year of attack), to yellow (year after attack), and red (2 yr after attack). We found large differences in volatile and within-needle terpene concentrations among crown classes and variation across a growing season. In general, emissions and concentrations of total and individual terpenes were greater for yellow and red needles than green needles. Douglas-fir beetle attack increased emissions and concentrations of terpene compounds linked to increased tree flammability in other conifer species and compounds known to attract beetles (e.g., [Formula: see text]-pinene, camphene, and D-limonene). There was little relationship between air temperature or within-needle concentrations of terpenes and emission of terpenes, suggesting that passive emission of terpenes (e.g., from dead foliage) does not fully explain changes in volatile emissions. The potential physiological causes and ecological consequences of these bark beetle-associated chemical changes are discussed. © The Authors 2016. Published by Oxford University Press on behalf of Entomological Society of America. All rights reserved. For Permissions, please email:

  6. Winter ecology of a subalpine grassland: Effects of snow removal on soil respiration, microbial structure and function. (United States)

    Gavazov, Konstantin; Ingrisch, Johannes; Hasibeder, Roland; Mills, Robert T E; Buttler, Alexandre; Gleixner, Gerd; Pumpanen, Jukka; Bahn, Michael


    Seasonal snow cover provides essential insulation for mountain ecosystems, but expected changes in precipitation patterns and snow cover duration due to global warming can influence the activity of soil microbial communities. In turn, these changes have the potential to create new dynamics of soil organic matter cycling. To assess the effects of experimental snow removal and advanced spring conditions on soil carbon (C) and nitrogen (N) dynamics, and on the biomass and structure of soil microbial communities, we performed an in situ study in a subalpine grassland in the Austrian Alps, in conjunction with soil incubations under controlled conditions. We found substantial winter C-mineralisation and high accumulation of inorganic and organic N in the topsoil, peaking at snowmelt. Soil microbial biomass doubled under the snow, paralleled by a fivefold increase in its C:N ratio, but no apparent change in its bacteria-dominated community structure. Snow removal led to a series of mild freeze-thaw cycles, which had minor effects on in situ soil CO2 production and N mineralisation. Incubated soil under advanced spring conditions, however, revealed an impaired microbial metabolism shortly after snow removal, characterised by a limited capacity for C-mineralisation of both fresh plant-derived substrates and existing soil organic matter (SOM), leading to reduced priming effects. This effect was transient and the observed recovery in microbial respiration and SOM priming towards the end of the winter season indicated microbial resilience to short-lived freeze-thaw disturbance under field conditions. Bacteria showed a higher potential for uptake of plant-derived C substrates during this recovery phase. The observed temporary loss in microbial C-mineralisation capacity and the promotion of bacteria over fungi can likely impede winter SOM cycling in mountain grasslands under recurrent winter climate change events, with plausible implications for soil nutrient availability and

  7. Distribution Pattern, Conservation Status, and Associated Flora of the Genus Juniperus in Subalpine Pastures of the Kashmir Himalayas

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    Hamayun Shaheen


    Full Text Available Juniperus is an evergreen gymnosperm genus with a broad geographical distribution in the Northern Hemisphere. Juniperus constitutes important vegetation associations in the Himalayan highlands that have significant ecological and socioeconomic importance. This research investigated the distribution pattern, community structure, and ecosystem services provided byJuniperus -dominated subalpine vegetation in the upper Neelum Valley, Pakistan. Vegetation attributes and geographical characteristics were systematically recorded at 4 selected sites. Two species of Juniperus, Juniperus communis L. and J. excelsa M. Bieb., were found to have average importance values of 23.4 and 20.02%, respectively. J. excelsa showed an average basal area of 0.30 m2 ha−1 and an average stem density of 46.95 ha−1; J. communis had an average basal area of 0.25 m2 ha−1 and an average stem density of 33.21 ha−1. A total of 56 Juniperus-associated plant species from 29 families were recorded, with Asteraceae as the dominant family, followed by Lamiaceae, Polygonaceae, Rosaceae, Caryophyllaceae, and Apiaceae. Predominant associated species included Thymus linearis, Aster falconeri, Rosa webbiana, Berberis lyceum, Anagallis arvensis, Rumex nepalensis, Poa alpina, Bistorta affinis, and Iris hookeriana. The calculated average values were Shannon's diversity, 3.07; Simpson's diversity, 0.94; species richness, 1.11; species evenness, 0.90; and maturity index, 45.90. Hemicryptophytes were the dominant lifeform in the area (57.14%, and microphylls (46.42% were the dominant leaf type. Overgrazing and fuelwood cutting were identified as serious threats to both Juniperus species. Restoration of the degraded juniper stands through collective efforts by government and local communities and regular monitoring is recommended.

  8. Thirty Years of Change in Subalpine Forest Cover from Landsat Image Analysis in the Sierra Nevada Mountains of California (United States)

    Potter, Christopher


    Landsat imagery was analyzed to understand changes in subalpine forest stands since the mid-1980s in the Sierra-Nevada region of California. At locations where long-term plot measurements have shown that stands are becoming denser in the number of small tree stems (compared to the early 1930s), the 30-year analysis of Landsat greenness index (NDVI) indicated that no consistent increases in canopy leaf cover have occurred at these same locations since the mid-1980s. Interannual variations in stand NDVI closely followed snow accumulation amounts recorded at nearby stations. In contrast, at eastern Sierra whitebark pine stand locations where it has been observed that widespread tree mortality has occurred, decreasing NDVI trends over the past 5-10 years were consistent with rapid loss of forest canopy cover. Landsat imagery was further analyzed to understand patterns of post-wildfire vegetation recovery, focusing on high burn severity (HBS) patches within burned areas dating from the late 1940s. Analysis of landscape metrics showed that the percentage of total HBS area comprised by the largest patch of recovered woody cover was relatively small in all fires that occurred since 1995, but increased rapidly with time since fire. Patch complexity of recovered woody cover decreased notably after more than 50 years of regrowth, but was not readily associated with time for fires that occurred since the mid 1990s. The aggregation level of patches with recovery of woody cover increased steadily with time since fire. The study approach using satellite remote sensing can be expanded to assess the consequences of stand-replacing wildfires in all forests of the region.

  9. Vegetation types and climate conditions reflected by the modern phytolith assemblages in the subalpine Dalaoling Forest Reserve, central China (United States)

    Traoré, Djakanibé Désiré; Gu, Yansheng; Liu, Humei; Shemsanga, Ceven; Ge, Jiwen


    This research describes modern phytolith records and distributions from subalpine surface soils in the Dalaoling Forest Reserve, and reveals its implications for local climate conditions with respect to the altitude gradient. Well-preserved phytolith morpho-types, assemblages, and climatic indices were used to study the relationship between local vegetation and climate conditions. The phytolith classification system is mainly based on the characteristics of detailed morpho-types described for anatomical terms, which are divided into seven groups: long cells, short cells, bulliform cells, hair cells, pteridophyte type, broad-leaved type, and gymnosperm type. Phytoliths originating from the Poaceae are composed of Pooideae (rondel and trapeziform), Panicoideae (bilobate, cross, and polylobate), Chloridoideae (short/square saddle), and Bambusoideae (oblong concave saddle). Based on the altitudinal distribution of the phytolith assemblages and the indices of aridity (Iph), climate (Ic), and tree cover density (D/P), five phytolith assemblage zones have revealed the five types of climatic conditions ranging from 1,169 m to 2,005 m in turn: warm-wet, warm-xeric to warm-mesic, warm-xeric to cool-mesic, cool-xeric, and cool-mesic to cool-xeric. The Bambusoideae, Panicoideae, and Chloridoideae are the dominant vegetation at the lower-middle of the mountains, while Pooideae is mainly distributed in the higher mountains. The close relationship between phytolith assembleages and changes of altitude gradient suggest that vegetation distribution patterns and plant ecology in the Dalaoling mountains are controlled by temperature and humidity conditions. Our results highlight the importance of phytolith records as reliable ecoclimatic indicators for vegetation ecology in subtropical regions.

  10. Mycorrhiza-plant colonization patterns on a subalpine glacier forefront as a model system of primary succession. (United States)

    Cázares, Efrén; Trappe, James M; Jumpponen, Ari


    Lyman glacier in the North Cascades Mountains of Washington has a subalpine forefront characterized by a well-developed terminal moraine, inconspicuous successional moraines, fluting, and outwash. These deposits were depleted of symbiotic fungi when first exposed but colonized by them over time after exposure. Four major groups of plant species in this system are (1) mycorrhiza-independent or facultative mycotrophic, (2) dependent on arbuscular mycorrhizae (AM) (3) dependent on ericoid mycorrhiza (ERM) or ectomycorrhizae (EM), and (4) colonized by dark-septate (DS) endophytes. We hypothesized that availability of mycorrhizal propagules was related to the success of mycorrhiza-dependent plants in colonizing new substrates in naturally evolved ecosystems. To test this hypothesis roots samples of 66 plant species were examined for mycorrhizal colonization. The plants were sampled from communities at increasing distances from the glacier terminus to compare the newest communities with successively older ones. Long established, secondary successional dry meadow communities adjacent to the glacier forefront, and nearby high alpine communities were sampled for comparison. DS were common on most plant species on the forefront. Nonmycorrhizal plants predominated in the earlier successional sites, whereas the proportion of mycorrhizal plants generally increased with age of community. AM were present, mostly at low levels, and nearly absent in two sites of the forefront. ERM were present in all species of Ericaceae sampled, and EM in all species of Pinaceae and Salicaceae. Roots of plants in the long established meadow and heath communities adjacent to the forefront and the high alpine community all had one or another of the colonization types, with DS and AM predominating.

  11. High-intensity coherent FIR radiation from sub-picosecond electron bunches

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    Kung, P.H.; Lihn, Hung-chi; Wiedemann, H.; Bocek, D.


    A facility to generate high-intensity, ultra-short pulses of broad-band far-infrared radiation has been assembled and tested at Stanford. The device uses sub-picosecond relativistic electron bunches to generate coherent radiation through transition or synchrotron radiation in the far-infrared (FIR) regime between millimeter waves and wavelengths of about 100 {mu}m and less. Experimental results show a peak radiation power of greater than 0.33 MW within a micro-bunch and an average FIR radiation power of 4 mW. The average bunch length of 2856 micro-bunches within a 1 {mu}sec macro-pulse is estimated to be about 480 sec. Simulations experimental setup and results will be discussed.

  12. Optimization of high speed pipelining in FPGA-based FIR filter design using genetic algorithm (United States)

    Meyer-Baese, Uwe; Botella, Guillermo; Romero, David E. T.; Kumm, Martin


    This paper compares FPGA-based full pipelined multiplierless FIR filter design options. Comparison of Distributed Arithmetic (DA), Common Sub-Expression (CSE) sharing and n-dimensional Reduced Adder Graph (RAG-n) multiplierless filter design methods in term of size, speed, and A*T product are provided. Since DA designs are table-based and CSE/RAG-n designs are adder-based, FPGA synthesis design data are used for a realistic comparison. Superior results of a genetic algorithm based optimization of pipeline registers and non-output fundamental coefficients are shown. FIR filters (posted as open source by Kastner et al.) for filters in the length from 6 to 151 coefficients are used.

  13. Increased Biomass of Nursery-Grown Douglas-Fir Seedlings upon Inoculation with Diazotrophic Endophytic Consortia

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    Zareen Khan


    Full Text Available Douglas-fir (Pseudotsuga menziesii seedlings are periodically challenged by biotic and abiotic stresses. The ability of endophytes to colonize the interior of plants could confer benefits to host plants that may play an important role in plant adaptation to environmental changes. In this greenhouse study, nursery-grown Douglas-fir seedlings were inoculated with diazotrophic endophytes previously isolated from poplar and willow trees and grown for fifteen months in nutrient-poor conditions. Inoculated seedlings had significant increases in biomass (48%, root length (13% and shoot height (16% compared to the control seedlings. Characterization of these endophytes for symbiotic traits in addition to nitrogen fixation revealed that they can also solubilize phosphate and produce siderophores. Colonization was observed through fluorescent microscopy in seedlings inoculated with gfp- and mkate-tagged strains. Inoculation with beneficial endophytes could prove to be valuable for increasing the production of planting stocks in forest nurseries.

  14. The effects of Douglas fir monoculture on stand characteristics in a zone of Montane beech forest

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    Kostić Olga


    Full Text Available The right choice of tree species to form forest cultures is of paramount importance to the preservation of the diversity, fertility and ecological stability of forest ecosystems. To that end, we examined the effect of a 40-year-long cultivation of Douglas fir (Pseudotsuga menziesii (Mirb Franco on the floristic composition, characteristics of the forest floor, physical and chemical properties of the soil and the intensity of organic matter decomposition in a beech forest in western Serbia (Mt. Maljen. It was found that the cultivation of Douglas fir caused a reduction in biodiversity, changes in the chemical properties of the soil, that were most pronounced in the surface layers (0-10 cm, and a slowing down in the metabolism of the beech stand. The absence of many plant species characteristic to natural beech forests was observed in the Douglas fir plantation, these were reflected in the detected changes in the chemical properties of the soil, such as lower substitutional acidity (p<0.05, depletion of the adsorption of basis in the cation complex (p<0.001 and lower amounts of C, N, P (p<0.001 and K (p<0.01 in relation to the beech stand (control. No differences were found in soil moisture and active acidity levels. The higher value of the C/N ratio of the Douglas fir litter (p<0.001 provided proof for its lower decomposition rate compared to beech litter (p<0.05. Over time, all these changes could lead to further acidification and degradation of the soil and a reduction in this ecosystem’s productivity.

  15. Influence of ultrasounds on some mechanical properties of fir wood (Abies alba Mill). Microscopic sections (United States)

    Parpala, V.; Pastirnac, A.; Paraschiv, N.


    It was found that as a result of ultrasonic treatment fir wood (Abies alba Mill) shows the effects of homogenization, and the modulus of elasticity for static flexure drops on an average by 50% for 45 min of treatment. The drop is more pronounced for test pieces with 8 to 12 annual rings. Study of microscopic sections disclosed that early wood with one annual ring undergoes the most powerful effect.

  16. Inheritance of germinative energy and germinative capacity in Douglas-fir (United States)

    Thomas E. Greathouse


    In the West foresters have had considerable difficulty in reforesting south-facing slopes. We considered this problem when we selected plus-trees for the first Douglas-fir seed orchard in Region 6. We were, however, faced with the need to answer such questions as these: (1) Should we try to produce seed inherently suited for south slopes? (2) If so, should we strive to...

  17. Manipulating stand structure of Douglas-fir plantations for wildlife habitat and wood production (United States)

    Timothy B. Harrington


    In southwestern Oregon, a study was initiated by Oregon State University (OSU) in 1983 to determine effects of hardwood competition on Douglas-fir plantation development. The use of herbicides to uniformly suppress competing vegetation—especially fast-growing hardwoods—proved to be an effective approach to ensure dominance and a high level of productivity for planted...

  18. Changes after partial cutting of a spruce-fir stand in Maine (United States)

    Arthur C. Hart


    In 1945 a partial-cutting experiment in spruce-fir silviculture was begun by the Penobscot Research Center of the Northeastern Forest Experiment Station. The Eastern Pulp Wood Company of Calais, Maine, made available for the study a tract of forest land, in Dyer Township, Washington County, Maine. This 20.6-acre tract is part of a large area that had been burned over...

  19. Structural DMR: A Technique for Implementation of Soft Error Tolerant FIR Filters


    Reviriego, P.; Bleakley, Chris J.; Maestro, J.A.


    In this brief, an efficient technique for implementation of soft-error-tolerant finite impulse response (FIR) filters is presented. The proposed technique uses two implementations of the basic filter with different structures operating in parallel. A soft error occurring in either filter causes the outputs of the filters to differ, or mismatch, for at least one sample. The filters are specifically designed so that, when a soft error occurs, they produce distinct error patterns at the filter o...

  20. Seasonal dynamics of structure and functional activity of ectomycorrhizal roots of the Siberian fir

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    T. A. Sizonenko


    Full Text Available The aim of our work was to study seasonal dynamics of the Siberian fir Abies sibirica Ledeb. ectomycorrhizal morpho-anatomical structure, respiration rate and fluorescence. The study was carried out in the bilberry-sphagnum spruce forest in the middle taiga of the Komi Republic, Russia. The morpho-anatomical structure and fluorescence parameters were studied by light and luminescence microscopy. Thin root respiration was studied in intact fine roots in the field using an infrared gas analyzer. 12 subtypes of fungal mantels were revealed in ectomycorrhizal fir roots; their amount and composition demonstrated seasonal dynamic changes. At the beginning vegetation stage, the diversity and proportion of pseudoparenchymatous and double covers were maximal. Plant component of ectomycorrhizae that includes cortical parenchyma and stele had high activity of fluorescence during the entire vegetation period. The dynamics of staining of fungal component (fungal mantel and Hartig net was more contrasting. The highest fluorescence intensity of cortical parenchyma was found in ectomycorrhizae with maximal fungal mantel thickness. High proportion of tannin cells in cortical parenchyma was related with low intensity of fungal mantel and Hartig net fluorescence. During vegetation season, maximal amount of intensively strained ectomycorrhizal elements occurred in July and unstrained – in June and August. Relation between fine roots respiration and an increase of brightly strained ectomycorrhizal structural elements in fir roots was not statistically significant. Root CO2-emission was lower in May and September in comparison with summer months. For respiration rate of fir fine roots we found its strong positive correlation with the litter temperature.

  1. Genetic variation in basic density and modulus of elasticity of coastal Douglas-fir. (United States)

    G.R. Johnson; B.L. Gartner


    Douglas-fir trees from 39 open-pollinated families at four test locations were assessed to estimate heritability of modulus of elasticity (MOE) and basic density. Heritability estimates of MOE (across-site h = 0.55) were larger than those for total height (0.15) and diameter at breast height (DBH; 0.29), and similar to those for density (0.59)....

  2. Dynamics and management of silver fir stand in Tuscany: a review of long-term research

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    Bianchi L


    Full Text Available The paper aims to outline the guidelines for the management of silver fir stands in Tuscan Apennines, according to a review of research activities carried out by the Silviculture Institute of Florence, nowadays Department of Forest Environmental Technologies (DISTAF, from 1960 up to today. The study cases are placed in the Vallombrosa Forest Reserve (province of Florence, in the National Park of Foreste Casentinesi Monte Falterona e Campigna (province of Arezzo and in Amiata Mountain (province of Siena. The main research activities concern different issues linked to the dynamics of silver fir stands: 1 ecology of natural regeneration, 2 stands dynamics in time, 3 sapling growth rates in relation with different canopy cover, 4 impact of wildlife on regeneration storey, 5 forest typology according to vegetation dynamics. As regards management aspects, the decrease of the importance of silver fir pure stands for timber production, as well as the increase of their role from naturalistic, recreation and landscape point of view, encourage close-to-nature sylviculture interventions aimed to promote stand dynamics towards mixed forest. Selective cuttings are proposed in order to progressively reduce the tree canopy cover and to favour the growth of regeneration layer. Nevertheless, wherever silver fir artificial stands play an important role from an historical point of view, the conservation of pure stand by means of clear cutting and artificial regeneration is recommended. Finally, the paper outlines the necessity to face the interactions between forest regeneration and wild ungulate populations, which currently represents one of the most dramatic problems of sustainable management.

  3. Transient physiological responses of planting frozen root plugs of Douglas-fir seedlings (United States)

    M. Anisul Islam; Douglass F. JAcobs; Kent G. Apostol; R. Kasten Dumroese


    Short-term physiological responses of planting frozen (FR) and rapidly thawed (TR) root plugs of Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) seedlings were examined through time series (0 h, 6 h, 12 h, 1 day, 3 days, and 7 days) measurements in two separate experiments: 10 C day: 6 C night, RH 75% and 30 C day: 20 C night, RH 50%, respectively...

  4. On the Nature of the Deeply Embedded Protostar OMC-2 FIR 4 (United States)

    Furlan, E.; Megeath, S. T.; Osorio, M.; Stutz, A. M.; Fischer, W. J.; Ali, B.; Stanke, T.; Manoj, P.; Adams, J. D.; Tobin, J. J.


    We use mid-infrared to submillimeter data from the Spitzer, Herschel, and Atacama Pathfinder Experiment telescopes to study the bright submillimeter source OMC-2 FIR 4. We find a point source at 8, 24, and 70 μm, and a compact, but extended source at 160, 350, and 870 μm. The peak of the emission from 8 to 70 μm, attributed to the protostar associated with FIR 4, is displaced relative to the peak of the extended emission; the latter represents the large molecular core the protostar is embedded within. We determine that the protostar has a bolometric luminosity of 37 L ⊙, although including more extended emission surrounding the point source raises this value to 86 L ⊙. Radiative transfer models of the protostellar system fit the observed spectral energy distribution well and yield a total luminosity of most likely less than 100 L ⊙. Our models suggest that the bolometric luminosity of the protostar could be as low as 12-14 L ⊙, while the luminosity of the colder (~20 K) extended core could be around 100 L ⊙, with a mass of about 27 M ⊙. Our derived luminosities for the protostar OMC-2 FIR 4 are in direct contradiction with previous claims of a total luminosity of 1000 L ⊙. Furthermore, we find evidence from far-infrared molecular spectra and 3.6 cm emission that FIR 4 drives an outflow. The final stellar mass the protostar will ultimately achieve is uncertain due to its association with the large reservoir of mass found in the cold core.

  5. Automated knot detection with visual post-processing of Douglas-fir veneer images (United States)

    C.L. Todoroki; Eini C. Lowell; Dennis Dykstra


    Knots on digital images of 51 full veneer sheets, obtained from nine peeler blocks crosscut from two 35-foot (10.7 m) long logs and one 18-foot (5.5 m) log from a single Douglas-fir tree, were detected using a two-phase algorithm. The algorithm was developed using one image, the Development Sheet, refined on five other images, the Training Sheets, and then applied to...

  6. Lumber and veneer recovery from intensively managed young-growth Douglas-fir. (United States)

    Thomas D. Fahey; James M. Cahill; Thomas A. Snellgrove; Linda S. Heath


    The objective of the study was to develop models that predict lumber and veneer recovery as a function of young-growth log characteristics. Empirical lumber and veneer recovery data for logs cut from more than 300 young-growth Douglas-fir trees were used in model development. Trees were sampled from 15 stands in western Oregon and Washington and represent a wide range...

  7. FIR: An Effective Scheme for Extracting Useful Metadata from Social Media. (United States)

    Chen, Long-Sheng; Lin, Zue-Cheng; Chang, Jing-Rong


    Recently, the use of social media for health information exchange is expanding among patients, physicians, and other health care professionals. In medical areas, social media allows non-experts to access, interpret, and generate medical information for their own care and the care of others. Researchers paid much attention on social media in medical educations, patient-pharmacist communications, adverse drug reactions detection, impacts of social media on medicine and healthcare, and so on. However, relatively few papers discuss how to extract useful knowledge from a huge amount of textual comments in social media effectively. Therefore, this study aims to propose a Fuzzy adaptive resonance theory network based Information Retrieval (FIR) scheme by combining Fuzzy adaptive resonance theory (ART) network, Latent Semantic Indexing (LSI), and association rules (AR) discovery to extract knowledge from social media. In our FIR scheme, Fuzzy ART network firstly has been employed to segment comments. Next, for each customer segment, we use LSI technique to retrieve important keywords. Then, in order to make the extracted keywords understandable, association rules mining is presented to organize these extracted keywords to build metadata. These extracted useful voices of customers will be transformed into design needs by using Quality Function Deployment (QFD) for further decision making. Unlike conventional information retrieval techniques which acquire too many keywords to get key points, our FIR scheme can extract understandable metadata from social media.

  8. Swiss Needle Cast in Western Oregon Douglas-Fir Plantations: 20‐Year Monitoring Results

    Directory of Open Access Journals (Sweden)

    Gabriela Ritóková


    Full Text Available Swiss needle cast (SNC, a foliar disease specific to Douglas-fir (Pseudotsuga menziesii, is caused by an endemic Ascomycete fungus (Phaeocryptopus gaeumannii. In the late 1980s and early 1990s significant symptoms began to appear in coastal Oregon, and these have been associated with the planting of Douglas-fir in the Sitka spruce zone, leaf wetness during potential spore dispersal in May–August, and mild winter temperature. The first annual aerial survey was initiated in 1996 and has continued through 2015, which indicates a significant increase in area of visible symptoms from the air, increasing from 53,050 ha in 1996 to 238,705 ha in 2015. Monitoring plots in the NW Oregon Coast Range verified impacts of SNC on tree growth and productivity, with growth reductions averaging about 23% in the epidemic area linked to needle retention. A series of monitoring plots was set up in the western Cascade Mountains of Oregon and 590 10–23-year old Douglas-fir trees in 59 stands were tracked for 10 years, measured in 2001, 2006, and 2011. No measureable growth impacts were noted in this region of Oregon. A new plot network is being installed throughout the Oregon and southwest Washington coastal ranges as a means of monitoring future disease impact and providing framework for additional studies.

  9. An advanced strategy for wind speed forecasting using expert 2-D FIR filters

    Directory of Open Access Journals (Sweden)



    Full Text Available Renewable energies such as wind and solar have become the most attractive means of electricity generation nowadays. Social and environmental benefits as well as economical issues result in further utilization of such these energy resources. In this regard, wind energy plays an important roll in operation of small-scale power systems like Micro Grid. On the other hand, wind stochastic nature in different time and place horizons, makes accurate forecasting of its behavior an inevitable task for market planners and energy management systems. In this paper an advanced strategy for wind speed estimation has been purposed and its superior performance is compared to that of conventional methods. The model is based on linear predictive filtering and image processing principles using 2-D FIR filters. To show the efficiency of purposed predictive model different FIR filters are designed and tested through similar data. Wind speed data have been collected during the period January 1, 2009 to December 31, 2009 from Casella automatic weather station at Plymouth. It is observed that 2-D FIR filters act more accurately in comparison with 1-D conventional representations; however, their prediction ability varies considerably through different filter sizing.

  10. A CS Recovery Algorithm for Model and Time Delay Identification of MISO-FIR Systems

    Directory of Open Access Journals (Sweden)

    Yanjun Liu


    Full Text Available This paper considers identifying the multiple input single output finite impulse response (MISO-FIR systems with unknown time delays and orders. Generally, parameters, orders and time delays of an MISO system are separately identified from different algorithms. In this paper, we aim to perform the model identification and time delay estimation simultaneously from a limited number of observations. For an MISO-FIR system with many inputs and unknown input time delays, the corresponding identification model contains a large number of parameters, requiring a great number of observations for identification and leading to a heavy computational burden. Inspired by the compressed sensing (CS recovery theory, a threshold orthogonal matching pursuit algorithm (TH-OMP is presented to simultaneously identify the parameters, the orders and the time delays of the MISO-FIR systems. The proposed algorithm requires only a small number of sampled data compared to the conventional identification methods, such as the least squares method. The effectiveness of the proposed algorithm is verified by simulation results.

  11. Design optimisation of powers-of-two FIR filter using self-organising random immigrants GA (United States)

    Chandra, Abhijit; Chattopadhyay, Sudipta


    In this communication, we propose a novel design strategy of multiplier-less low-pass finite impulse response (FIR) filter with the aid of a recent evolutionary optimisation technique, known as the self-organising random immigrants genetic algorithm. Individual impulse response coefficients of the proposed filter have been encoded as sum of signed powers-of-two. During the formulation of the cost function for the optimisation algorithm, both the frequency response characteristic and the hardware cost of the discrete coefficient FIR filter have been considered. The role of crossover probability of the optimisation technique has been evaluated on the overall performance of the proposed strategy. For this purpose, the convergence characteristic of the optimisation technique has been included in the simulation results. In our analysis, two design examples of different specifications have been taken into account. In order to substantiate the efficiency of our proposed structure, a number of state-of-the-art design strategies of multiplier-less FIR filter have also been included in this article for the purpose of comparison. Critical analysis of the result unambiguously establishes the usefulness of our proposed approach for the hardware efficient design of digital filter.

  12. How Do Trees Know When to Flower? Predicting Reproductive Phenology of Douglas-fir with Changing Winter and Spring Temperatures (United States)

    Prevey, J.; St Clair, B.; Harrington, C.


    Flowering at the right time is one of the primary ways that plants are adapted to their environment. Trees that flower too early risk cold damage to vulnerable new tissues and those that flower too late miss peak resources or may mistime flowering to coincide with other trees, altering outcrossing rates and gene flow. Past observations indicate that temperature cues over winter and spring influence the timing of flowering in many tree species. Understanding these cues is important for predicting how flowering phenology of trees will change with a changing climate.We developed predictive models of flowering for Douglas-fir, an abundant and commercially important tree in the Pacific Northwest. We assembled over 10,000 flowering observations of trees from 11 sites across western Oregon and Washington. We modeled the dates of flowering using hourly temperature data; our models of flowering were adapted from previous models of vegetative budburst and height growth initiation developed for Douglas-fir. Preliminary results show that both chilling (cold) and forcing (warm) temperatures over winter and spring are important determinants of flowering time for Douglas-fir. This suggests that as spring temperatures warm in the future, Douglas-fir across the Pacific Northwest will flower earlier, unless plants experience insufficient chilling over winter, in which case it is possible that Douglas-fir may flower later than in the past, or not flower at all. At one site, Douglas-fir genotypes from different geographic regions flowered in the same order from year to year, indicating that both temperature and heredity influence flowering. Knowledge of the environmental and genetic cues that drive the timing of flowering can help predict how changes in temperature under various climate models could change flowering time across sites. These models may also indicate the geographic areas where future climate could enhance or reduce flowering of Douglas-fir in the future.

  13. Structure and productivity of mixed spruce and fir forests on Mt. Kopaonik

    Directory of Open Access Journals (Sweden)

    Šljukić Biljana


    Full Text Available The subject of this research are mixed forests of spruce and fir in the area of NP Kopaonik, which belong to the community of spruce and fir - Abieti-Piceetum abietis Mišić et Popović, 1978. The basis for the study of the structural development and production potential of these forests are data from 12 sample plots, with the average size of 0.18 ha. In terms of coenoecological affiliation all the sample plots belong to the group of ecological units - forests of spruce and fir (Abieti-Piceetum abietis, Mišić et Popović, 1978 on acid brown and brown podzolic soils, which are differentiated into 5 ecological units: Abieti-Piceetum abietis oxalidetosum on brown podzolic soil, Abieti-Piceetum abietis oxalidetosum on acid brown soil, Abieti-Piceetum abietis vaccinietosum on brown podzolic soil, Abieti-Piceetum abietis typicum on brown podzolic soil and Abieti-Piceetum abietis Dr.ymetosum on brown pozolic soil. In structural terms, these forests are characterized by very diverse structural forms, ranging from the structure of even-aged stands to typical multi-storey, unevenaged-aged stands. The form of cumulative curves of tree distribution is in most cases determined by spruce as the dominant species. At the same time, thin and medium-thick trees dominate, while the presence of stems with large dimensions is minimal. The average volume of these forestse is 777 m3•ha-1, with a mixture ratio of 0.7: 0.3 in favor of spruce. The average value of the current volume increment is 14 m3•ha-1, with a 68% share of spruce and 32% of fir. The percentage of increment ranges from 1.6% to 2.5% in all sample plots and is somewhat higher for fir. The site potential, stand characteristics and relations among the tree species have resulted in structural complexity, high productivity and ecological stability of these forests. Therefore, future forest management should avoid radical measures and procedures that would violate the established relationships and

  14. Distinct genecological patterns in seedlings of Norway spruce and silver fir from a mountainous landscape. (United States)

    Frank, Aline; Sperisen, Christoph; Howe, Glenn Thomas; Brang, Peter; Walthert, Lorenz; St Clair, John Bradley; Heiri, Caroline


    Understanding the genecology of forest trees is critical for gene conservation, for predicting the effects of climate change and climate change adaptation, and for successful reforestation. Although common genecological patterns have emerged, species-specific details are also important. Which species are most vulnerable to climate change? Which are the most important adaptive traits and environmental drivers of natural selection? Even though species have been classified as adaptive specialists vs. adaptive generalists, large-scale studies comparing different species in the same experiment are rare. We studied the genecology of Norway spruce (Picea abies) and silver fir (Abies alba), two co-occurring but ecologically distinct European conifers in Central Europe. For each species, we collected seed from more than 90 populations across Switzerland, established a seedling common-garden test, and developed genecological models that associate population variation in seedling growth and phenology to climate, soil properties, and site water balance. Population differentiation and associations between seedling traits and environmental variables were much stronger for Norway spruce than for silver fir, and stronger for seedling height growth than for bud phenology. In Norway spruce, height growth and second flushing were strongly associated with temperature and elevation, with seedlings from the lowlands being taller and more prone to second flush than seedlings from the Alps. In silver fir, height growth was more weakly associated with temperature and elevation, but also associated with water availability. Soil characteristics explained little population variation in both species. We conclude that Norway spruce has become an adaptive specialist because trade-offs between rapid juvenile growth and frost avoidance have subjected it to strong diversifying natural selection based on temperature. In contrast, because silver fir has a more conservative growth habit, it has

  15. Ash, Asterionella, and Anglers: A Paleolimnological Approach to Understanding Anthropogenic and Volcanogenic Disturbances in a Small Sub-Alpine Watershed (United States)

    Howard, K. L.; Noble, P. J.


    This poster summarizes geochemical, biological, hydrological, and watershed data that characterize Manzanita Lake, a small sub-alpine catchment in Lassen Volcanic National Park, CA. The future objective is to use characteristics of this system to interpret variations in diatom and sediment composition dating back to the 1914 Mt. Lassen eruption from a recently recovered lake core. Manzanita Lake is a small (0.18 km2) lake with a ~30 km2 watershed area situated on the northwest flank of Mt. Lassen, one of the most active Cascade volcanoes, and is a valuable recreational spot for anglers and visitors. Hydraulic residence time is short; roughly 119 days, and is derived from lake volume (1.0 X 106m3) and estimates of stream inflow (~6 ft3/sec) and outflow (~3 ft3/sec) that were made from May 2014 sampling data. Limnological sampling in 2012-2014 suggests that Manzanita Lake exhibits stable thermal stratification in the summer months, which is unusual given the shallow depth (~10m), but possibly supported by the morphometry of the lake basin and inputs of cold snowmelt from the flank of Lassen Peak. The lake is a moderately conductive (100-114 μS), mesotrophic system with secchi depths ranging from 8m to the bottom (~10m). Total phosphorus (TP) ranges from 15-25 ppb and dissolved inorganic nitrogen (DIN) from 2-15 ppb, with ammonium in the epilimnion being the largest contributor. A high concentration of silica (Si) in surface water inputs (34 mg/L) to Manzanita Lake likely reflects the rhyodacitic bedrock geology and large drainage ratio (164) of the watershed. Variations in Si concentration in the lake seem to be coupled with diatom production. During the sampling period Manzanita Lake is has been dominated by diatom blooms throughout the summer and fall months. There is a seasonal succession in the diatom species present, with abundant Asterionella formosa in the spring, transitioning to abundant Fragilaria crotonensis in the summer months, to a mixed dominance of

  16. Spatio-seasonal variability in dissolved organic matter optical properties and its bioavailability in a subalpine lake (United States)

    Stadler, Masumi; Ejarque, Elisabet; Kainz, Martin J.


    Allochthonous and autochothonous dissolved organic matter (DOM) in lakes mainly originate from terrestrial and aquatic primary production, respectively. Due to their differing biochemical composition the degradability of DOM by microorganisms is expected to vary. The carbon use efficiency of bacteria and DOM biodegradability determine whether the consumed DOM is incorporated into microbial biomass or respired to CO2 and ultimately emitted into the atmosphere. Thus, understanding the interaction of biodegradable DOM and its consumers is crucial to increase our knowledge on the role of lakes in the global carbon cycling. However, interactions of specific aquatic DOM signatures and the microbial population still remain widely debated. The aim of this study was to explore how DOM biodegradability changes along a stream-lake continuum at different seasons of the year. We monitored DOM quantity and its optical properties, inorganic nutrients, CO2 and bacterial growth over 20 days in dark bioassays with water from the inflow, outflow and at three layers of an oligotrophic subalpine lake. Preliminary results reveal highest microbial abundance in the metalimnion in winter and summer (0.7 106 and 2.5 106 cells mL-1, respectively) and the inflow in spring and autumn (1 106 and 1.4 106 cells mL-1, respectively) after 20 days. Surprisingly, with the exception of winter samples final inflow bacterial abundance results high, despite its lowest initial natural cell concentration, providing evidence for effective utilisation of terrestrial DOM, even with its high humic signature as indicated by the humification index (HIX). Nonetheless, after a microbial biomass peak with the inflow yielding mostly highest after three days, at the final experimental stage microbial biomass does only marginally differ between all sites with the exception of autumn samples where outflow and metalimnion turn out most productive. Even though the DOM of all lake sites and the lake outflow were

  17. Drought induced changes of plant belowground carbon allocation affect soil microbial community function in a subalpine meadow (United States)

    Fuchslueger, L.; Bahn, M.; Fritz, K.; Hasibeder, R.; Richter, A.


    There is growing evidence that climate extremes may affect ecosystem carbon dynamics more strongly than gradual changes in temperatures or precipitation. Climate projections suggest more frequent heat waves accompanied by extreme drought periods in many parts of Europe, including the Alps. Drought is considered to decrease plant C uptake and turnover, which may in turn decrease belowground C allocation and potentially has significant consequences for microbial community composition and functioning. However, information on effects of drought on C dynamics at the plant-soil interface in real ecosystems is still scarce. Our study aimed at understanding how summer drought affects soil microbial community composition and the uptake of recently assimilated plant C by different microbial groups in grassland. We hypothesized that under drought 1) the microbial community shifts, fungi being less affected than bacteria, 2) plants decrease belowground C allocation, which further reduces C transfer to soil microbes and 3) the combined effects of belowground C allocation, reduced soil C transport due to reduced soil moisture and shift in microbial communities cause an accumulation of extractable organic C in the soil. Our study was conducted as part of a rain-exclusion experiment in a subalpine meadow in the Austrian Central Alps. After eight weeks of rain exclusion we pulse labelled drought and control plots with 13CO2 and traced C in plant biomass, extractable organic C (EOC) and soil microbial communities using phospholipid fatty acids (PLFA). Drought induced a shift of the microbial community composition: gram-positive bacteria became more dominant, whereas gram-negative bacteria were not affected by drought. Also the relative abundance of fungal biomass was not affected by drought. While total microbial biomass (as estimated by total microbial PLFA content) increased during drought, less 13C was taken up. This reduction was pronounced for bacterial biomarkers. It reflects

  18. Soil attributes and microclimate are important drivers of initial deadwood decay in sub-alpine Norway spruce forests. (United States)

    Fravolini, Giulia; Egli, Markus; Derungs, Curdin; Cherubini, Paolo; Ascher-Jenull, Judith; Gómez-Brandón, María; Bardelli, Tommaso; Tognetti, Roberto; Lombardi, Fabio; Marchetti, Marco


    Deadwood is known to significantly contribute to global terrestrial carbon stocks and carbon cycling, but its decay dynamics are still not thoroughly understood. Although the chemistry of deadwood has been studied as a function of decay stage in temperate to subalpine environments, it has generally not been related to time. We therefore studied the decay (mass of deadwood, cellulose and lignin) of equal-sized blocks of Picea abies wood in soil-mesocosms over two years in the Italian Alps. The 8 sites selected were along an altitudinal sequence, reflecting different climate zones. In addition, the effect of exposure (north- and south-facing slopes) was taken into account. The decay dynamics of the mass of deadwood, cellulose and lignin were related to soil parameters (pH, soil texture, moisture, temperature) and climatic data. The decay rate constants of Picea abies deadwood were low (on average between 0.039 and 0.040y(-1)) and of lignin close to zero (or not detectable), while cellulose reacted much faster with average decay rate constants between 0.110 and 0.117y(-1). Our field experiments showed that local scale factors, such as soil parameters and topographic properties, influenced the decay process: higher soil moisture and clay content along with a lower pH seemed to accelerate wood decay. Interestingly, air temperature negatively correlated with decay rates or positively with the amount of wood components on south-facing sites. It exerted its influence rather on moisture availability, i.e. the lower the temperature the higher the moisture availability. Topographic features were also relevant with generally slower decay processes on south-facing sites than on north-facing sites owing to the drier conditions, the higher pH and the lower weathering state of the soils (less clay minerals). This study highlights the importance of a multifactorial consideration of edaphic parameters to unravel the complex dynamics of initial wood decay. Copyright © 2016 Elsevier

  19. In vivo evaluation of the effect of stimulus distribution on FIR statistical efficiency in event-related fMRI. (United States)

    Jansma, J Martijn; de Zwart, Jacco A; van Gelderen, Peter; Duyn, Jeff H; Drevets, Wayne C; Furey, Maura L


    Technical developments in MRI have improved signal to noise, allowing use of analysis methods such as Finite impulse response (FIR) of rapid event related functional MRI (er-fMRI). FIR is one of the most informative analysis methods as it determines onset and full shape of the hemodynamic response function (HRF) without any a priori assumptions. FIR is however vulnerable to multicollinearity, which is directly related to the distribution of stimuli over time. Efficiency can be optimized by simplifying a design, and restricting stimuli distribution to specific sequences, while more design flexibility necessarily reduces efficiency. However, the actual effect of efficiency on fMRI results has never been tested in vivo. Thus, it is currently difficult to make an informed choice between protocol flexibility and statistical efficiency. The main goal of this study was to assign concrete fMRI signal to noise values to the abstract scale of FIR statistical efficiency. Ten subjects repeated a perception task with five random and m-sequence based protocol, with varying but, according to literature, acceptable levels of multicollinearity. Results indicated substantial differences in signal standard deviation, while the level was a function of multicollinearity. Experiment protocols varied up to 55.4% in standard deviation. Results confirm that quality of fMRI in an FIR analysis can significantly and substantially vary with statistical efficiency. Our in vivo measurements can be used to aid in making an informed decision between freedom in protocol design and statistical efficiency. Published by Elsevier B.V.

  20. An Innovations-Based Noise Cancelling Technique on Inverse Kepstrum Whitening Filter and Adaptive FIR Filter in Beamforming Structure (United States)

    Jeong, Jinsoo


    This paper presents an acoustic noise cancelling technique using an inverse kepstrum system as an innovations-based whitening application for an adaptive finite impulse response (FIR) filter in beamforming structure. The inverse kepstrum method uses an innovations-whitened form from one acoustic path transfer function between a reference microphone sensor and a noise source so that the rear-end reference signal will then be a whitened sequence to a cascaded adaptive FIR filter in the beamforming structure. By using an inverse kepstrum filter as a whitening filter with the use of a delay filter, the cascaded adaptive FIR filter estimates only the numerator of the polynomial part from the ratio of overall combined transfer functions. The test results have shown that the adaptive FIR filter is more effective in beamforming structure than an adaptive noise cancelling (ANC) structure in terms of signal distortion in the desired signal and noise reduction in noise with nonminimum phase components. In addition, the inverse kepstrum method shows almost the same convergence level in estimate of noise statistics with the use of a smaller amount of adaptive FIR filter weights than the kepstrum method, hence it could provide better computational simplicity in processing. Furthermore, the rear-end inverse kepstrum method in beamforming structure has shown less signal distortion in the desired signal than the front-end kepstrum method and the front-end inverse kepstrum method in beamforming structure. PMID:22163987

  1. Diameter Growth, Biological Rotation Age and Biomass of Chinese Fir in Burning and Clearing Site Preparations in Subtropical China

    Directory of Open Access Journals (Sweden)

    Hua Zhou


    Full Text Available Sustained forest management of Cunninghamia lanceolata (Chinese fir plantations in subtropical China is restricted by the limited availability of quantitative data. This study combines inventory data and tree-ring analysis of Chinese fir from natural and plantation forests that were subjected to controlled burning or brush clearing site preparations. Inter-annual variation of Chinese fir tree-ring widths were measured for the controlled burning, brush clearing and natural forest sites. The mean annual diametric growth of Chinese fir was 0.56 cm·year−1 for the natural forest, 0.80 cm·year−1 for the brush clearing site and 1.10 cm·year−1 for the controlled burning site. The time needed to reach the minimum cutting/logging diameter of 15 cm was 14 years in the controlled burning site, 19 years in the brush clearing site and >40 years in the natural forest. The biological rotation ages for the burning, cutting and natural forest sites were 15, 26 and >100 years, respectively. The total aboveground biomasses for the burning and clearing sites were 269.8 t·ha−1 and 252 t·ha−1, respectively. These results suggest that the current 25-year cutting cycle greatly underestimates the growth rate of Chinese fir plantations.

  2. Pityophthorus orarius Bright (Coleoptera: Scolytidae) in a northern California Douglas-fir seed orchard: effect of clone, tree vigor, and cone crop on rate of attack (United States)

    Nancy G. Rappaport; David L. Wood


    The geographic range of the Douglas-fir twig beetle, Pityophthorus orarius Bright, was extended beyond the original provenance of southern British Columbia to northern California. A survey of 457 Douglas-fir [Pseudotsuga menziesii (Mirb.) Franco] trees in 1985 revealed that those with heavy cone crops were more likely to be...

  3. Resource use and clonal differences in attack rate by the Douglas-fir seed chalcid, Megastigmus spermotrophus Wachtl (Hymenoptera: Torymidae), in France (United States)

    Nancy Rappaport; Alain Roques


    The within-cone distribution of Megastigmus spermotrophus Wachtl (Hymenoptera: Torymidae), the Douglas-fir seed chalcid, infesting Douglas-fir [Pseudotsuga menziesii (Mirb.) Franco] cones from north-central France was compared with that in samples from California. Results indicate that the mid-region of cones was more intensively...

  4. Five-year field trials using preservative-treated, second-growth Douglas-fir exposed in ground contact in Australia (United States)

    Rodney C. De Groot; Douglas M. Crawford; Jack Norton; John Keith


    The durability of preservative-treated stakes of second-growth Coastal Douglas-fir was evaluated in a field plot in northern Queensland, Australia. Results from this field trial indicate that second-growth Douglas-fir can be treated with preservatives to meet Australian standards and will provide long-term durability in adverse environments. Data presented indicate...

  5. Ecological impacts of using chloropicrin to control laminated root rot in Northwest conifer forests: growth and mycorrhiza formation of planted Douglas-fir seedlings after two growing seasons. (United States)

    Michael A. Castellano; Donaraye McKay; Walter G. Thles


    Bareroot Douglas-fir seedlings inoculated with Rhizopogon sp. and processed by standard nursery and reforestation procedures performed equally well whether planted near Douglas-fir stumps previously fumigated with two dosages of chloropicrin to control Phellinus weirii infection or near stumps not fumigated. Before stump-...

  6. Demonstrating a New Census of Infrared Galaxies with ALMA (DANCING-ALMA). I. FIR Size and Luminosity Relation at z = 0-6 Revealed with 1034 ALMA Sources (United States)

    Fujimoto, Seiji; Ouchi, Masami; Shibuya, Takatoshi; Nagai, Hiroshi


    We present the large statistics of the galaxy effective radius R e in the rest-frame far-infrared (FIR) wavelength {R}{{e}({FIR})} obtained from 1627 Atacama Large Millimeter/submillimeter Array (ALMA) 1 mm band maps that become public by 2017 July. Our ALMA sample consists of 1034 sources with the star formation rate ˜ 100{--}1000 {M}⊙ {{yr}}-1 and the stellar mass ˜ {10}10{--}{10}11.5 {M}⊙ at z = 0-6. We homogeneously derive {R}{{e}({FIR})} and FIR luminosity L FIR of our ALMA sources via the uv-visibility method with the exponential disk model, carefully evaluating selection and measurement incompletenesses by realistic Monte-Carlo simulations. We find that there is a positive correlation between {R}{{e}({FIR})} and L FIR at the >99% significance level. The best-fit power-law function, {R}{{e}({FIR})}\\propto {L}{FIR}α , provides α =0.28+/- 0.07, and shows that {R}{{e}({FIR})} at a fixed L FIR decreases toward high redshifts. The best-fit α and the redshift evolution of {R}{{e}({FIR})} are similar to those of R e in the rest-frame UV (optical) wavelength {R}{{e}({UV})} ({R}{{e}({Opt}.)}) revealed by Hubble Space Telescope (HST) studies. We identify that our ALMA sources have significant trends of {R}{{e}({FIR})}≲ {R}{{e}({UV})} and {R}{{e}({Opt}.)}, which suggests that the dusty starbursts take place in compact regions. Moreover, {R}{{e}({FIR})} of our ALMA sources is comparable to {R}{{e}({Opt}.)} of quiescent galaxies at z ˜ 1-3 as a function of stellar mass, supporting the evolutionary connection between these two galaxy populations. We also investigate rest-frame UV and optical morphologies of our ALMA sources with deep HST images, and find that ˜30%-40% of our ALMA sources are classified as major mergers. This indicates that dusty starbursts are triggered by not only the major mergers but also the other mechanism(s).

  7. The influence of changes in soil moisture in association with geomorphic change on the formation of a subalpine coniferous forest on Mt. Akita-Komagatake, northern Japan (United States)

    Konno, A.


    The coniferous forest (largely composed of Abies mariesii) is presently the typical vegetation of the subalpine zone in Japan. Pollen analysis revealed that few A. mariesii were present during the last glacial period, and the species began to expand to the subalpine zone during the Holocene (Morita, 1992). However, on Mt. Akita-Komagatake in northern Japan, the expected predominance of A. mariesii is not extensively observed, and the predominant vegetation is instead the dwarf bamboo (Sasa kurilensis). It is unknown why the area under coniferous forest is small in this region. Therefore, I examined this issue from the perspectives of (1) distribution of vegetation, (2) geomorphology, (3) soil moisture, and (4) vegetation history. (1) Precise digital elevation model data and photographic interpretation showed that this coniferous forest was densely distributed in a flat segment considered to be formed by a landslide; (2) this landslide is thought to have occurred up to 3,699 ± 26 yr BP because a boring-core specimen from the landslide included the AK-3 tephra layer (2,300-2,800 yr BP: Wachi et al, 1997) and the radiocarbon date of the lowermost humic soil layer was 3,699 ± 26 yr BP; (3) the soil in the forest area had higher volumetric water content than that in the non-forest area; and (4) phytolith analysis revealed that the main species in the study site was initially dwarf bamboo, but coniferous forest replaced it after the Towada-a tephra (1035 cal. BP, Machida and Arai, 1992) layer fell. These results suggest that soil water conditions changed because of the formation of the flat segment by the landslide, and the coniferous forest was consequently established. However, the landslide only indirectly affected the formation of the coniferous forest, because the forest developed over several thousand years after the landslide occurred. In other words, more direct reasons for the establishment of the coniferous forest may involve changes in soil moisture. This

  8. Calculation methods of reactivity using derivatives of nuclear power and Filter fir; Metodos para o calculo da reatividade usando derivadas da potencia nuclear e o filtro FIR

    Energy Technology Data Exchange (ETDEWEB)

    Diaz, Daniel Suescun


    This work presents two new methods for the solution of the inverse point kinetics equation. The first method is based on the integration by parts of the integral of the inverse point kinetics equation, which results in a power series in terms of the nuclear power in time dependence. Applying some conditions to the nuclear power, the reactivity is represented as first and second derivatives of this nuclear power. This new calculation method for reactivity has special characteristics, amongst which the possibility of using different sampling periods, and the possibility of restarting the calculation, after its interruption associated it with a possible equipment malfunction, allowing the calculation of reactivity in a non-continuous way. Apart from this reactivity can be obtained with or without dependency on the nuclear power memory. The second method is based on the Laplace transform of the point kinetics equations, resulting in an expression equivalent to the inverse kinetics equation as a function of the power history. The reactivity can be written in terms of the summation of convolution with response to impulse, characteristic of a linear system. For its digital form the Z-transform is used, which is the discrete version of the Laplace transform. In this method it can be pointed out that the linear part is equivalent to a filter named Finite Impulse Response (Fir). The Fir filter will always be, stable and non-varying in time, and, apart from this, it can be implemented in the non-recursive way. This type of implementation does not require feedback, allowing the calculation of reactivity in a continuous way. The proposed methods were validated using signals with random noise and showing the relationship between the reactivity difference and the degree of the random noise. (author)

  9. Star Formation Under the Outflow: The Discovery of a Non-thermal Jet from OMC-2 FIR 3 and Its Relationship to the Deeply Embedded FIR 4 Protostar (United States)

    Osorio, Mayra; Díaz-Rodríguez, Ana K.; Anglada, Guillem; Megeath, S. Thomas; Rodríguez, Luis F.; Tobin, John J.; Stutz, Amelia M.; Furlan, Elise; Fischer, William J.; Manoj, P.; Gómez, José F.; González-García, Beatriz; Stanke, Thomas; Watson, Dan M.; Loinard, Laurent; Vavrek, Roland; Carrasco-González, Carlos


    We carried out multiwavelength (0.7-5 cm), multi-epoch (1994-2015) Very Large Array (VLA) observations toward the region enclosing the bright far-IR sources FIR 3 (HOPS 370) and FIR 4 (HOPS 108) in OMC-2. We report the detection of 10 radio sources, 7 of them identified as young stellar objects. We image a well-collimated radio jet with a thermal free-free core (VLA 11) associated with the Class I intermediate-mass protostar HOPS 370. The jet features several knots (VLA 12N, 12C, 12S) of non-thermal radio emission (likely synchrotron from shock-accelerated relativistic electrons) at distances of ˜7500-12,500 au from the protostar, in a region where other shock tracers have been previously identified. These knots are moving away from the HOPS 370 protostar at ˜100 km s-1. The Class 0 protostar HOPS 108, which itself is detected as an independent, kinematically decoupled radio source, falls in the path of these non-thermal radio knots. These results favor the previously proposed scenario in which the formation of HOPS 108 is triggered by the impact of the HOPS 370 outflow with a dense clump. However, HOPS 108 has a large proper motion velocity of ˜30 km s-1, similar to that of other runaway stars in Orion, whose origin would be puzzling within this scenario. Alternatively, an apparent proper motion could result because of changes in the position of the centroid of the source due to blending with nearby extended emission, variations in the source shape, and/or opacity effects.

  10. Star Formation Under the Outflow: The Discovery of a Non-thermal Jet from OMC-2 FIR 3 and Its Relationship to the Deeply Embedded FIR 4 Protostar

    Energy Technology Data Exchange (ETDEWEB)

    Osorio, Mayra; Díaz-Rodríguez, Ana K.; Anglada, Guillem; Gómez, José F. [Instituto de Astrofísica de Andalucía (CSIC) Glorieta de la Astronomía s/n E-18008 Granada (Spain); Megeath, S. Thomas [Ritter Astrophysical Research Center, Department of Physics and Astronomy University of Toledo 2801 West Bancroft Street Toledo, OH 43606 (United States); Rodríguez, Luis F.; Loinard, Laurent; Carrasco-González, Carlos [Instituto de Radioastronomía y Astrofísica, UNAM Apartado Postal 3-72 (Xangari), 58089 Morelia, Michoacán (Mexico); Tobin, John J. [Homer L. Dodge Department of Physics and Astronomy University of Oklahoma, Norman, OK 73019 (United States); Stutz, Amelia M. [Department of Astronomy, University of Concepción Concepción (Chile); Furlan, Elise [IPAC, Mail Code 314-6, Caltech 1200 E. California Boulevard, Pasadena, CA 91125 (United States); Fischer, William J. [Space Telescope Science Institute 3700 San Martin Drive, Baltimore, MD 21218 (United States); Manoj, P. [Tata Institute of Fundamental Research Homi Bhabha Road, Mumbai 400 005 (India); González-García, Beatriz; Vavrek, Roland [European Space Astronomy Center, ESA P.O. Box 78, E-28691 Villanueva de la Cañada, Madrid (Spain); Stanke, Thomas [European Southern Observatory Garching bei München (Germany); Watson, Dan M., E-mail: [Department of Physics and Astronomy, University of Rochester Rochester, NY 14627 (United States)


    We carried out multiwavelength (0.7–5 cm), multi-epoch (1994–2015) Very Large Array (VLA) observations toward the region enclosing the bright far-IR sources FIR 3 (HOPS 370) and FIR 4 (HOPS 108) in OMC-2. We report the detection of 10 radio sources, 7 of them identified as young stellar objects. We image a well-collimated radio jet with a thermal free–free core (VLA 11) associated with the Class I intermediate-mass protostar HOPS 370. The jet features several knots (VLA 12N, 12C, 12S) of non-thermal radio emission (likely synchrotron from shock-accelerated relativistic electrons) at distances of ∼7500–12,500 au from the protostar, in a region where other shock tracers have been previously identified. These knots are moving away from the HOPS 370 protostar at ∼100 km s{sup −1}. The Class 0 protostar HOPS 108, which itself is detected as an independent, kinematically decoupled radio source, falls in the path of these non-thermal radio knots. These results favor the previously proposed scenario in which the formation of HOPS 108 is triggered by the impact of the HOPS 370 outflow with a dense clump. However, HOPS 108 has a large proper motion velocity of ∼30 km s{sup −1}, similar to that of other runaway stars in Orion, whose origin would be puzzling within this scenario. Alternatively, an apparent proper motion could result because of changes in the position of the centroid of the source due to blending with nearby extended emission, variations in the source shape, and/or opacity effects.

  11. Comparative study of long-term water uptake of Norway spruce and Douglas-fir in Moravian upland

    Directory of Open Access Journals (Sweden)

    Nadezhdina Nadezhda


    Full Text Available Long-term water uptake of Douglas-fir and Norway spruce trees, growing in condition of Moravian upland, was studied with aim of comparing sap flow in small roots with flow in stems. Sap flow was measured by the heat field deformation method using multi-point sensors for stems and single-point sensors for roots. Differences between species were found in relationships between sap flow in tree stems and water uptake by roots, suggesting that Douglas-fir is able to take water from deeper soil more efficiently than spruce. This allows Douglas-fir to transpire more water especially during drought and grow faster than spruce. These biological features should be taken into account for future forest species compositions because they may have impact on both, forestry and hydrology.

  12. Filtered Iterative Reconstruction (FIR) via Proximal Forward-Backward Splitting: A Synergy of Analytical and Iterative Reconstruction Method for CT

    CERN Document Server

    Gao, Hao


    This work is to develop a general framework, namely filtered iterative reconstruction (FIR) method, to incorporate analytical reconstruction (AR) method into iterative reconstruction (IR) method, for enhanced CT image quality. Specifically, FIR is formulated as a combination of filtered data fidelity and sparsity regularization, and then solved by proximal forward-backward splitting (PFBS) algorithm. As a result, the image reconstruction decouples data fidelity and image regularization with a two-step iterative scheme, during which an AR-projection step updates the filtered data fidelity term, while a denoising solver updates the sparsity regularization term. During the AR-projection step, the image is projected to the data domain to form the data residual, and then reconstructed by certain AR to a residual image which is in turn weighted together with previous image iterate to form next image iterate. Since the eigenvalues of AR-projection operator are close to the unity, PFBS based FIR has a fast convergenc...

  13. Design of Multiplier-less FIR filters with Simultaneously Variable Bandwidth and Fractional Delay

    Directory of Open Access Journals (Sweden)

    Aravind Illa


    Full Text Available Low complexity and reconfigurability are the key features in emerging communication applications in order to support multi-standards and operation modes. To obtain these features, an efficient implementation of finite impulse response (FIR filter, with Variable Bandwidth and Fractional Delay, is proposed in this paper. The reduction in the implementation complexity is achieved by converting the continuous filter coefficients to signed power of two (SPT space. This may lead to performance degradation. Hence, in this paper, Artificial Bee Colony (ABC optimization is deployed for finding the near optimal solution in the discrete space.

  14. Properties of predictor based on relative neighborhood graph localized FIR filters

    DEFF Research Database (Denmark)

    Sørensen, John Aasted


    of the training vectors belonging to the lunes formed by the nodes. The predictor training is carried out by iteration through 3 stages: initialization of the RNG of the training signal by vector quantization, LS estimation of the FIR filters localized in the input space by RNG nodes and adaptation of the RNG...... nodes by equalizing the LS approximation error among the lunes formed by the nodes of the RNG. The training properties of the predictor is exemplified on a burst signal and characterized by the normalized mean square error (NMSE) and the mean valence of the RNG nodes through the adaptation...

  15. Design of broadly tuned FIR FEL based on a variable-period microwiggler

    Energy Technology Data Exchange (ETDEWEB)

    Qing-Xiang Liu [Institute of Applied Electronics, Sichuan (China)]|[Southwest Jiaotong Univ., Sichuan (China); Yong Xu [Southwest Jiatong Univ., Sichuan (China)


    A varible-period microwiggler is proposed and investigated. The fundamental period of the microwiggler is designed as {lambda}o=2mm, and the period of the microwiggler can be turned from {lambda}o to n{lambda}o (n=1,2,3,{hor_ellipsis}) The wiggler fields with the period 3{lambda}o, 4{lambda}o, and 5{lambda}o are measured and compared with the theoretical results. Finally, a broadly tuned FIR FEL is designed based on the performance of the variable-period microwiggler.

  16. Modified Particle Swarm Optimization for Blind Deconvolution and Identification of Multichannel FIR Filters

    Directory of Open Access Journals (Sweden)

    Khanagha Ali


    Full Text Available Blind identification of MIMO FIR systems has widely received attentions in various fields of wireless data communications. Here, we use Particle Swarm Optimization (PSO as the update mechanism of the well-known inverse filtering approach and we show its good performance compared to original method. Specially, the proposed method is shown to be more robust against lower SNR scenarios or in cases with smaller lengths of available data records. Also, a modified version of PSO is presented which further improves the robustness and preciseness of PSO algorithm. However the most important promise of the modified version is its drastically faster convergence compared to standard implementation of PSO.

  17. Energy-Aware Scheduling of FIR Filter Structures using a Timed Automata Model

    DEFF Research Database (Denmark)

    Wognsen, Erik Ramsgaard; Hansen, Rene Rydhof; Larsen, Kim Guldstrand


    structures where we initially derive data flow graphs and precedence graphs using the Synchronous Data Flow (SDF) notation. Based on actual measurements on the Altera Cyclone IV FPGA, we derive power and timing estimates for addition and multiplication, including idling power consumption. We next model...... the FIR structures in UPPAAL CORA and employ model checking to find energyoptimal solutions in linearly priced timed automata. In conclusion we state that there are significant energy-versus-time differences between the four structures when we experiment with varying numbers of adders and multipliers...

  18. An Experimental Assessment of Transverse Adaptive Fir Filters as Applied to Vibrating Structures Identification

    Directory of Open Access Journals (Sweden)

    Daniel A. Castello


    Full Text Available The present work is aimed at assessing the performance of adaptive Finite Impulse Response (FIR filters on the identification of vibrating structures. Four adaptive algorithms were used: Least Mean Squares (LMS, Normalized Least Mean Squares (NLMS, Transform-Domain Least Mean Squares (TD – LMS and Set-Membership Binormalized Data-Reusing LMS Algorithm (SM – BNDRLMS. The capability of these filters to perform the identification of vibrating structures is shown on real experiments. The first experiment consists of an aluminum cantilever beam containing piezoelectric sensors and actuators and the second one is a steel pinned-pinned beam instrumented with accelerometers and an electromechanical shaker.

  19. Floristic and community diversity of sub-alpine and alpine grasslands and grazed dwarf-shrub heaths in the Romanian Carpathians

    Directory of Open Access Journals (Sweden)

    Coldea, Gheorghe


    Full Text Available Floristic diversity at community level and community diversity at landscape level are presented for the main grassland and dwarf-shrub communities in the sub-alpine and alpine zones in five mountain ranges of the Romanian Carpathians. The 30 plant communities studied had their floristic composition and distribution determined by geological substratum and pedo-climatic factors. The most diverse grasslands (Oxytropido-Elynetum, Seslerio haynaldianae-Caricetun were on calcareous substratum and on the mountain slopes (Festucetum pictae, whilst the lowest diversity was in the oligo-mesotrophic sub-alpine grasslands (Scorzonero- Festucetum nigricantis, Alchemillo-Poetum alpinae, Violo-Nardetum. The Shannon-Weaver diversity index for the communities increased almost linearly with the number of species per community, whilst community diversity in the five mountain ranges was a function of the number of communities per mountain range.

    [fr] Nous présentons dans ce travail la diversité floristique au niveau de la communauté et la diversité de communautés au niveau du paysage pour les principaux types de pâturages et de communautés à petits arbustes des étages subalpin et alpin de cinq chaînes montagneuses des Carpathes en Roumanie. Aussi bien la composition floristique que la distribution des 30 communautés étudiées dépendent de la nature géologique du terrain et des facteurs édapho-climatiques. Les pâturages les plus riches en diversité (Oxytropido-Elynetum, Seslerio haynaldianae- Caricetun se développent sur substrat calcaire et sur les versants des montagnes (Testucetum pictae; par contre la plus faible diversité a été rencontrée dans les pâturages oligo-mesotrophes subalpins (Scorzonero-Festucetum nigricantis, Alchemillo-Poetum alpinae, Violo-Nardetum. L'index de diversité de Shannon- Weaver augmente quasi linéairement avec le nombre d'espèces de chaque communauté, tandis que

  20. Responses of assemblages of Orthoptera to management and use of ski slopes on upper sub-alpine meadows in the Austrian Alps. (United States)

    Illich, Ingeborg P; Haslett, John R


    The Orthoptera assemblages occurring on sub-alpine ski slopes were compared with those found on neighbouring unskied meadows by making frequent transect counts at two pairs of sites in the Gastein valley in the Austrian Central Alps. On one of the ski slopes no Orthoptera were present, although two species were abundant on the control meadow a few meters away. On the second ski slope, the Orthoptera assemblage exhibited reduced species richness, lower densities of individuals and a generally accelerated rate of nymphal development compared to the control meadow populations. These results may be explained in terms of the changed habitat conditions on the ski slopes and the known biologies of the species concerned. The implications of the findings for winter tourism management in high altitude ecosystems are briefly discussed.

  1. Effects of elevated CO₂ and temperature on photosynthesis and leaf traits of an understory dwarf bamboo in subalpine forest zone, China. (United States)

    Li, Yongping; Zhang, Yuanbin; Zhang, Xiaolu; Korpelainen, Helena; Berninger, Frank; Li, Chunyang


    The dwarf bamboo (Fargesia rufa Yi), growing understory in subalpine dark coniferous forest, is one of the main foods for giant panda, and it influences the regeneration of subalpine coniferous forests in southwestern China. To investigate the effects of elevated CO₂, temperature and their combination, the dwarf bamboo plantlets were exposed to two CO₂ regimes (ambient and double ambient CO₂ concentration) and two temperatures (ambient and +2.2°C) in growth chambers. Gas exchange, leaf traits and carbohydrates concentration were measured after the 150-day experiment. Elevated CO₂ significantly increased the net photosynthetic rate (Anet ), intrinsic water-use efficiency (WUEi ) and carbon isotope composition (δ¹³C) and decreased stomatal conductance (g(s)) and total chlorophyll concentration based on mass (Chl(m)) and area (Chl(a)). On the other hand, elevated CO₂ decreased specific leaf area (SLA), which was increased by elevated temperature. Elevated CO₂ also increased foliar carbon concentration based on mass (C(m)) and area (C(a)), nitrogen concentration based on area (N(a)), carbohydrates concentration (i.e. sucrose, sugar, starch and non-structural carbohydrates) and the slope of the A(net)-N(a) relationship. However, elevated temperature decreased C(m), C(a) and N(a). The combination of elevated CO₂ and temperature hardly affected SLA, C(m), C(a), N(m), N(a), Chl(m) and Chl(a). Variables Anet and Na had positive linear relationships in all treatments. Our results showed that photosynthetic acclimation did not occur in dwarf bamboo at elevated CO₂ and it could adjust physiology and morphology to enable the capture of more light, to increase WUE and improve nutritional conditions. Copyright © Physiologia Plantarum 2012.

  2. Leap frog in slow motion: Divergent responses of tree species and life stages to climatic warming in Great Basin subalpine forests. (United States)

    Smithers, Brian V; North, Malcolm P; Millar, Constance I; Latimer, Andrew M


    In response to climate warming, subalpine treelines are expected to move up in elevation since treelines are generally controlled by growing season temperature. Where treeline is advancing, dispersal differences and early life stage environmental tolerances are likely to affect how species expand their ranges. Species with an establishment advantage will colonize newly available habitat first, potentially excluding species that have slower establishment rates. Using a network of plots across five mountain ranges, we described patterns of upslope elevational range shift for the two dominant Great Basin subalpine species, limber pine and Great Basin bristlecone pine. We found that the Great Basin treeline for these species is expanding upslope with a mean vertical elevation shift of 19.1 m since 1950, which is lower than what we might expect based on temperature increases alone. The largest advances were on limber pine-dominated granitic soils, on west aspects, and at lower latitudes. Bristlecone pine juveniles establishing above treeline share some environmental associations with bristlecone adults. Limber pine above-treeline juveniles, in contrast, are prevalent across environmental conditions and share few environmental associations with limber pine adults. Strikingly, limber pine is establishing above treeline throughout the region without regard to site characteristic such as soil type, slope, aspect, or soil texture. Although limber pine is often rare at treeline where it coexists with bristlecone pine, limber pine juveniles dominate above treeline even on calcareous soils that are core bristlecone pine habitat. Limber pine is successfully "leap-frogging" over bristlecone pine, probably because of its strong dispersal advantage and broader tolerances for establishment. This early-stage dominance indicates the potential for the species composition of treeline to change in response to climate change. More broadly, it shows how species differences in dispersal

  3. Combustion influences on natural abundance nitrogen isotope ratio in soil and plants following a wildfire in a sub-alpine ecosystem. (United States)

    Huber, Edith; Bell, Tina L; Adams, Mark A


    This before-and-after-impact study uses the natural abundance N isotope ratio (δ(15)N) to investigate the effects of a wildfire on sub-alpine ecosystem properties and processes. We measured the (15)N signatures of soil, charred organic material, ash and foliage in three sub-alpine plant communities (grassland, heathland and woodland) in south-eastern Australia. Surface bulk soil was temporarily enriched in (15)N immediately after wildfire compared to charred organic material and ash in all plant communities. We associated the enrichment of bulk soil with fractionation of N during combustion and volatilization of N, a process that also explains the sequential enrichment of (15)N of unburnt leaves > ash > charred organic material in relation to duration and intensity of heating. The rapid decline in (15)N of bulk soil to pre-fire values indicates that depleted ash, containing considerable amounts of total N, was readily incorporated into the soil. Foliar δ(15)N also increased with values peaking 1 year post-fire. Foliar enrichment was foremost coupled with the release of enriched NH4(+) into the soil owing to isotopic discrimination during volatilization of soluble N and combustion of organic material. The mode of post-fire regeneration influenced foliar (15)N enrichment in two species indicating use of different sources of N following fire. The use of natural abundance of (15)N in soil, ash and foliage as a means of tracing transformation of N during wildfire has established the importance of combustion products as an important, albeit temporary source of inorganic N for plants regenerating after wildfire.

  4. Using stable water isotopes and borehole NMR to inform an ecohydrological model in a subalpine and upper montane catchment in the Rocky Mountains (United States)

    Millar, D.; Parsekian, A.; Mercer, J.; Speckman, H. N.; Beverly, D.; Ewers, B. E.; Mackay, D. S.


    Recent work using stable water isotopes has revealed that vegetation across a range of different biomes preferentially take up tightly bound soil water over more mobile pools. This so called two water worlds hypothesis (TWWH) has important implications for hydrological modeling efforts in ecosystems where it holds true, since few if any ecohydrological models incorporate this phenomenon. Further, in ecosystems where the TWWH is supported, information regarding the proportion of soil water in the bound and mobile pools is necessary to inform plant-soil water dynamics in models. In this study, we investigate which soil water pools are used by dominant vegetation in an upper montane and subalpine catchment in the Rocky Mountains of southern Wyoming, and use this information to inform the Terrestrial Regional Ecosystem Exchange Simulator (TREES). Within each catchment, we test the TWWH using stable water isotope analyses in an upland coniferous forest and an adjacent, downgradient groundwater-supported wetland. The proportion of soil water in each pool within each ecosystem was inferred from borehole nuclear magnetic resonance (NMR). These field data are being incorporated into TREES, by partitioning plant water uptake between bound and mobile pools. NMR analyses were conducted in all four ecosystems down to a depth of approximately 75 cm and revealed that while mid growing season soil water content was approximately two-fold higher in the subalpine forest versus that of the upper montane forest, the vast majority of soil water, 86% on average, existed in the bound pool in both ecosystems. Alternatively, soils in both wetlands were saturated throughout their profiles, with a majority of the soil water existing in the mobile pool, 63% on average. These initial findings highlight the importance of bound soil water pools in both upland forests, as opposed to the wetlands, which had an abundance of water in both pools.

  5. Seasonal dynamics of the plant community and soil seed bank along a successional gradient in a subalpine meadow on the Tibetan Plateau.

    Directory of Open Access Journals (Sweden)

    Miaojun Ma

    Full Text Available BACKGROUND: Knowledge about how change the importance of soil seed bank and relationship between seed mass and abundance during vegetation succession is crucial for understanding vegetation dynamics. Many studies have been conducted, but their ecological mechanisms of community assembly are not fully understood. METHODOLOGY: We examined the seasonal dynamics of the vegetation and soil seed bank as well as seed size distribution along a successional gradient. We also explored the potential role of the soil seed bank in plant community regeneration, the relationship between seed mass and species abundance, and the relative importance of deterministic and stochastic processes along a successional gradient. PRINCIPAL FINDINGS: Species richness of seed bank increased (shallow layer and the total and seed density decreased (each layer and the total significantly with succession. Species richness and seed density differed significantly between different seasons and among soil depths. Seed mass showed a significant negative relationship with relative abundance in the earliest successional stage, but the relationships were not significant in later stages. Seed mass showed no relationship with relative abundance in the whole successional series in seed bank. Results were similar for both July 2005 and April 2006. CONCLUSIONS: The seed mass and abundance relationship was determined by a complex interaction between small and larger seeded species and environmental factors. Both stochastic processes and deterministic processes were important determinants of the structure of the earliest stage. The importance of seed bank decreased with succession. The restoration of abandoned farmed and grazed meadows to the species-rich subalpine meadow in Tibetan Plateau can be successfully achieved from the soil seed bank. However, at least 20 years are required to fully restore an abandoned agricultural meadow to a natural mature subalpine meadow.

  6. A multi-proxy record of hydroclimate, vegetation, fire, and post-settlement impacts for a subalpine plateau, Central Rocky Mountains U.S.A (United States)

    Anderson, Lesleigh; Brunelle, Andrea; Thompson, Robert S.


    Apparent changes in vegetation distribution, fire, and other disturbance regimes throughout western North America have prompted investigations of the relative importance of human activities and climate change as potential causal mechanisms. Assessing the effects of Euro-American settlement is difficult because climate changes occur on multi-decadal to centennial time scales and require longer time perspectives than historic observations can provide. Here, we report vegetation and environmental changes over the past ~13,000 years as recorded in a sediment record from Bison Lake, a subalpine lake on a high plateau in northwestern Colorado. Results are based on multiple independent proxies, which include pollen, charcoal, and elemental geochemistry, and are compared with previously reported interpretations of hydroclimatic changes from oxygen isotope ratios. The pollen data indicate a slowly changing vegetation sequence from sagebrush steppe during the late glacial to coniferous forest through the late Holocene. The most dramatic vegetation changes of the Holocene occurred during the ‘Medieval Climate Anomaly’ (MCA) and ‘Little Ice Age’ (LIA) with rapid replacement of conifer forest by grassland followed by an equally rapid return to conifer forest. Late Holocene vegetation responses are mirrored by changes in fire, lake biological productivity, and watershed erosion. These combined records indicate that subsequent disturbance related to Euro-American settlement, although perhaps significant, had acted upon a landscape that was already responding to MCA-LIA hydroclimatic change. Results document both rapid and long-term subalpine grassland ecosystem dynamics driven by agents of change that can be anticipated in the future and simulated by ecosystem models.

  7. Mercury bioaccumulation in fishes from subalpine lakes of the Wallowa-Whitman National Forest, northeastern Oregon and western Idaho (United States)

    Eagles-Smith, Collin A.; Herring, Garth; Johnson, Branden L.; Graw, Rick


    Mercury (Hg) is a globally distributed pollutant that poses considerable risks to human and wildlife health. Over the past 150 years since the advent of the industrial revolution, approximately 80 percent of global emissions have come from anthropogenic sources, largely fossil fuel combustion. As a result, atmospheric deposition of Hg has increased by up to 4-fold above pre-industrial times. Because of their isolation, remote high-elevation lakes represent unique environments for evaluating the bioaccumulation of atmospherically deposited Hg through freshwater food webs, as well as for evaluating the relative importance of Hg loading versus landscape influences on Hg bioaccumulation. The increase in Hg deposition to these systems over the past century, coupled with their limited exposure to direct anthropogenic disturbance make them useful indicators for estimating how changes in Hg emissions may propagate to changes in Hg bioaccumulation and ecological risk. In this study, we evaluated Hg concentrations in fishes of high-elevation, sub-alpine lakes in the Wallowa-Whitman National Forest in northeastern Oregon and western Idaho. Our goals were to (1) assess the magnitude of Hg contamination in small-catchment lakes to evaluate the risk of atmospheric Hg to human and wildlife health, (2) quantify the spatial variability in fish Hg concentrations, and (3) determine the ecological, limnological, and landscape factors that are best correlated with fish total mercury (THg) concentrations in these systems. Across the 28 study lakes, mean THg concentrations of resident salmonid fishes varied as much as 18-fold among lakes. Importantly, our top statistical model explained 87 percent of the variability in fish THg concentrations among lakes with four key landscape and limnological variables— catchment conifer density (basal area of conifers within a lake’s catchment), lake surface area, aqueous dissolved sulfate, and dissolved organic carbon. The basal area of conifers

  8. Progress on FIR interferometry and Thomson Scattering measurements on HIT-SI3 (United States)

    Everson, Christopher; Jarboe, Thomas; Morgan, Kyle


    Spatially resolved measurements of the electron temperature (Te) and density (ne) will be fundamental in assessing the degree to which HIT-SI3 demonstrates closed magnetic flux and energy confinement. Further, electron temperature measurements have not yet been made on an inductively-driven spheromak. Far infrared (FIR) interferometer and Thomson Scattering (TS) systems have been installed on the HIT-SI3 spheromak. The TS system currently implemented on HIT-SI3 was originally designed for other magnetic confinement experiments, and progress continues toward modifying and optimizing for HIT-SI3 plasmas. Initial results suggest that the electron temperature is of order 10 eV. Plans to modify the TS system to provide more sensitivity and accuracy at low temperatures are presented. The line-integrated ne is measured on one chord by the FIR interferometer, with densities near 5x1019 m-3. Four cylindrical volumes have been added to the HIT-SI3 apparatus to enhance passive pumping. It is hoped that this will allow for more control of the density during the 2 ms discharges. Density measurements from before and after the installation of the passive pumping volumes are presented for comparison.

  9. Tolerance to multiple climate stressors: A case study of Douglas-fir drought and cold hardiness (United States)

    Bansal, Sheel; Harrington, Constance A; St. Clair, John Bradley


    Summary: 1. Drought and freeze events are two of the most common forms of climate extremes which result in tree damage or death, and the frequency and intensity of both stressors may increase with climate change. Few studies have examined natural covariation in stress tolerance traits to cope with multiple stressors among wild plant populations. 2. We assessed the capacity of coastal Douglas-fir (Pseudotsuga menziesii var. menziesii), an ecologically and economically important species in the northwestern USA, to tolerate both drought and cold stress on 35 populations grown in common gardens. We used principal components analysis to combine drought and cold hardiness trait data into generalized stress hardiness traits to model geographic variation in hardiness as a function of climate across the Douglas-fir range. 3. Drought and cold hardiness converged among populations along winter temperature gradients and diverged along summer precipitation gradients. Populations originating in regions with cold winters had relatively high tolerance to both drought and cold stress, which is likely due to overlapping adaptations for coping with winter desiccation. Populations from regions with dry summers had increased drought hardiness but reduced cold hardiness, suggesting a trade-off in tolerance mechanisms. 4. Our findings highlight the necessity to look beyond bivariate trait–climate relationships and instead consider multiple traits and climate variables to effectively model and manage for the impacts of climate change on widespread species.

  10. Evaluation of the FIR Example using Xilinx Vivado High-Level Synthesis Compiler

    Energy Technology Data Exchange (ETDEWEB)

    Jin, Zheming [Argonne National Lab. (ANL), Argonne, IL (United States); Finkel, Hal [Argonne National Lab. (ANL), Argonne, IL (United States); Yoshii, Kazutomo [Argonne National Lab. (ANL), Argonne, IL (United States); Cappello, Franck [Argonne National Lab. (ANL), Argonne, IL (United States)


    Compared to central processing units (CPUs) and graphics processing units (GPUs), field programmable gate arrays (FPGAs) have major advantages in reconfigurability and performance achieved per watt. This development flow has been augmented with high-level synthesis (HLS) flow that can convert programs written in a high-level programming language to Hardware Description Language (HDL). Using high-level programming languages such as C, C++, and OpenCL for FPGA-based development could allow software developers, who have little FPGA knowledge, to take advantage of the FPGA-based application acceleration. This improves developer productivity and makes the FPGA-based acceleration accessible to hardware and software developers. Xilinx Vivado HLS compiler is a high-level synthesis tool that enables C, C++ and System C specification to be directly targeted into Xilinx FPGAs without the need to create RTL manually. The white paper [1] published recently by Xilinx uses a finite impulse response (FIR) example to demonstrate the variable-precision features in the Vivado HLS compiler and the resource and power benefits of converting floating point to fixed point for a design. To get a better understanding of variable-precision features in terms of resource usage and performance, this report presents the experimental results of evaluating the FIR example using Vivado HLS 2017.1 and a Kintex Ultrascale FPGA. In addition, we evaluated the half-precision floating-point data type against the double-precision and single-precision data type and present the detailed results.

  11. EuroFIR Guideline on calculation of nutrient content of foods for food business operators. (United States)

    Machackova, Marie; Giertlova, Anna; Porubska, Janka; Roe, Mark; Ramos, Carlos; Finglas, Paul


    This paper presents a Guideline for calculating nutrient content of foods by calculation methods for food business operators and presents data on compliance between calculated values and analytically determined values. In the EU, calculation methods are legally valid to determine the nutrient values of foods for nutrition labelling (Regulation (EU) No 1169/2011). However, neither a specific calculation method nor rules for use of retention factors are defined. EuroFIR AISBL (European Food Information Resource) has introduced a Recipe Calculation Guideline based on the EuroFIR harmonized procedure for recipe calculation. The aim is to provide food businesses with a step-by-step tool for calculating nutrient content of foods for the purpose of nutrition declaration. The development of this Guideline and use in the Czech Republic is described and future application to other Member States is discussed. Limitations of calculation methods and the importance of high quality food composition data are discussed. Copyright © 2017. Published by Elsevier Ltd.

  12. A Method for Designing FIR Filters with Arbitrary Magnitude Characteristic Used for Modeling Human Audiogram

    Directory of Open Access Journals (Sweden)

    SZOPOS, E.


    Full Text Available This paper presents an iterative method for designing FIR filters that implement arbitrary magnitude characteristics, defined by the user through a set of frequency-magnitude points (frequency samples. The proposed method is based on the non-uniform frequency sampling algorithm. For each iteration a new set of frequency samples is generated, by processing the set used in the previous run; this implies changing the samples location around the previous frequency values and adjusting their magnitude through interpolation. If necessary, additional samples can be introduced, as well. After each iteration the magnitude characteristic of the resulting filter is determined by using the non-uniform DFT and compared with the required one; if the errors are larger than the acceptable levels (set by the user a new iteration is run; the length of the resulting filter and the values of its coefficients are also taken into consideration when deciding a re-run. To demonstrate the efficiency of the proposed method a tool for designing FIR filters that match human audiograms was implemented in LabVIEW. It was shown that the resulting filters have smaller coefficients than the standard one, and can also have lower order, while the errors remain relatively small.

  13. [Identification and analysis of differentially expressed genes during wood formation in Chinese fir by SSH]. (United States)

    Wang, Gui-Feng; Gao, Yan; Yang, Li-Wei; Shi, Ji-Sen


    Wood is an important raw material for the global industry with rapidly increasing demand. To isolate the differentially expressed genes in xylogenesis of Chinese fir [Cunninghamia lanceolata (Lamb.) Hook], a forward subtractive cDNA library was constructed using suppression subtractive hybridization (SSH) method, which was performed using the cDNA from the mutant Dugansha clone as the tester and the cDNA from the normal Jurong 0 clone as the driver. Six hundred and eighteen clones were obtained. Recombinants were identified using PCR with universal T7 and SP6 primers and using EcoR digestion. To further eliminate false positive, dot hybridization was used with four DIG-labeled probes (FSP, RSP, UTP, and UDP). Real-time PCR was performed to confirm the results. A total of 260 unique ESTs were obtained, 60% of the ESTs exhibiting homologies with proteins of known function fell into 4 major classes: metabolism, cell wall biogenesis and remodeling, signal transduction and stress. The systematic analysis of genes involved in wood formation in Chinese fir provides valuable insights into the molecular mechanisms involved in xylem differentiation, is important resources for forest research directed toward understanding the genetic control of wood formation and future endeavors to modify wood and fiber properties for industrial use.

  14. Investigation of process of interception in beech-fir stand on mountain Goch

    Directory of Open Access Journals (Sweden)

    Ristić Ratko


    Full Text Available Part of the precipitation is intercepted by vegetation before reaching the ground. The portion of intercepted water, which is retained in storage of the vegetal cover and evaporates, is called the interception loss. Interception represents important component of water balance, because of reduction of total rainfall. It decreases potential for forming of surface runoff. Depends on complex of vegetative and climate factors. In humid forested regions about 25% of the annual precipitation may become interception loss. Results of investigation in beech-fir stand on mountain Goch are presented in this paper Investigation was carried out on experimental catchment area Vaona IV (A=0.098 km2, in association Abieti-fagetum, on mountain Goch, in Central Serbia. Standard rain gauges were used to determine total precipitation and throughfalls, during vegetation period (1992-1997. Total interception (Ic depends on total precipitation (Pb, kind of trees, shape and density of the crown, position in the stand and age. Average values of interception (Icsr during vegetation period (1992-1997, amount to Icsr=43.7-53.3% of total precipitation (fir, and Icsr=17.7-22.8% (beech.

  15. Survival and growth of northern white-cedar and balsam fir seedlings in riparian management zones in northern Minnesota, USA (United States)

    Brian J. Palik; Brooke K. Haworth; Andrew J. David; Randall K. Kolka


    Northern white-cedar (Thuja occidentalis) and balsam fir (Abies balsamea.) are co-occurring species in riparian forests of the western Great Lakes region. Throughout much of the region, northern white-cedar has been experiencing population declines due to herbivory by white-tailed deer (Odocoileus virginianus)....

  16. EuroFIR-BASIS - a combined composition and biological activity database for bioactive compounds in plant-based foods

    DEFF Research Database (Denmark)

    Gry, Jørn; Black, Lucinda; Eriksen, Folmer Damsted


    Mounting evidence suggests that certain non-nutrient bioactive compounds promote optimal human health and reduce the risk of chronic disease. An Internet-deployed database, EuroFIR-BASIS, which uniquely combines food composition and biological effects data for plant-based bioactive compounds, is ...

  17. Relationships among foliar phenology, radial growth rate, and xylem density in a young Douglas-fir plantation (United States)

    Warren D. Devine; Constance A. Harrington


    We related intraannual patterns in radial growth rate and xylem density to foliar phenology and second growth flushes in a young Douglas-fir plantation in western Washington. Three foliar maturity classes were defined: (1) shoots and needles elongating; (2) elongation complete, needles maturing; and (3) needles mature. Diameter growth rate had two peaks, one about the...

  18. The wind stability of different silvicultural systems for Douglas-fir in The Netherlands: a model-based approach

    NARCIS (Netherlands)

    Schelhaas, M.J.


    The aim of this study was to evaluate different silvicultural systems for Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) in the Netherlands in terms of timber production and wind stability over a full rotation. This was done using the forest genetics, ecology, management and wind model

  19. Modeling regional and climatic variation of wood density and ring width in intensively managed Douglas-fir (United States)

    Cosmin N. Filipescue; Eini C. Lowell; Ross Koppenaal; Al K. Mitchell


    Characteristics of annual rings are reliable indicators of growth and wood quality in trees. The main objective of our study was to model the variation in annual ring attributes due to intensive silviculture and inherent regional differences in climate and site across a wide geographic range of Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco)....

  20. Interim definitions for old growth Douglas-fir and mixed-conifer forests in the Pacific Northwest and California. (United States)

    J.F. Franklin; F. Hall; W. Laudenslayer; C. Maser; J. Nunan; J. Poppino; C.J. Ralph; T. Spies


    Interim definitions of old-growth forests are provided to guide efforts in land-management planning until comprehensive definitions based on research that is currently underway can be formulated. The basic criteria for identifying old-growth Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) and mixed-conifer forests in western Washington and...

  1. Hot Water and Copper Coatings in Reused Containers Decrease Inoculum of Fusarium and Cylindrocarpon and Increase Douglas Fir Seedling Growth (United States)

    R. Kasten Dumroese; Robert L. James; David L. Wenny


    Inoculum of Douglas fir root diseases caused by the fungi Fusarium and Cylindrocarpon is carried from crop to crop in reused containers. Soaking containers for 90 seconds in 80 °C water removed ~99% of Fusarium and 100% of Cylindrocarpon inoculum between growing cycles. Overall seedling growth was also improved:...

  2. Hydraulic architecture and photosynthetic capacity as constraints on release from suppression in Douglas-fir and western hemlock. (United States)

    Renninger H.J.; Meinzer F.C.; B.L. Gartner


    We compared hydraulic architecture, photosynthesis, and growth in Douglas-fir with that of a shade-tolerant western hemlock. The study was conducted in a site that had been thinned to release suppressed trees, and one that remained unthinned. Release seemed to be constrained initially by photosynthetic capacity in both species. After released trees increased their...

  3. Bioassays of TH6038 and difluron applied to western spruce budworm and Douglas-fir Tussock moth (United States)

    Nancy L. Gillette; Jacqueline L. Robertson; Robert L. Lyon


    Two insects molt inhibitors, TH6038 N-[[4-cholorphenyl)amino]carbonyl]-2,6-dichlorobenzamide) and difluron (N-[[(4-chlorophenyl)amino]carbonyl]-2,6-difluorobenzamide), were tested for topical and feeding toxicity to the western spruce budworm, Choristoneura occidentalis Freeman, and the Douglas-fir tussock moth, Orgyia pseudotsugata...

  4. Spatial and population characteristics of dwarf mistletoe infected trees in an old-growth Douglas-fir - western hemlock forest. (United States)

    David C. Shaw; Jiquan Chen; Elizabeth A. Freeman; David M. Braun


    We investigated the distribution and severity of trees infected with western hemlock dwarf mistletoe (Arceuthobium tsugense (Rosendahl) G.N. Jones subsp. tsugense) in an old-growth Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) - western hemlock (Tsuga heterophylla (Raf.) Sarg.)...

  5. Respiration , nitrogen fixation, and mineralizable nitrogen spatial and temporal patterns within two Oregon Douglas-fir stands. (United States)

    Sharon M. Hope; Ching-Yan. Li


    Substrate respiration, mineralizable nitrogen, and nitrogen fixation rates, substrate moisture,content, and temperature were measured in trenched and undisturbed plots within two western Oregon Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) stands. The stands represent two different environments and ages. Woods Creek, the site of the lower...

  6. Development of a hybrid modeling approach for predicting intensively managed Douglas-fir growth at multiple scales. (United States)

    A. Weiskittel; D. Maguire; R. Monserud


    Hybrid models offer the opportunity to improve future growth projections by combining advantages of both empirical and process-based modeling approaches. Hybrid models have been constructed in several regions and their performance relative to a purely empirical approach has varied. A hybrid model was constructed for intensively managed Douglas-fir plantations in the...

  7. Acoustic Evaluation of Thinning and Biosolid Fertilization Effects on Wood Quality of a Douglas-fir stand (United States)

    Xiping Wang; Robert J. Ross; Steve Verrill; Eini Lowell; Jamie Barbour


    In this study, we examined the potential of using a time-of-flight (TOF) acoustic wave method to evaluate thinning and biosolid fertilization effects on acoustic velocity of trees and modulus of elasticity (MOE) of structural lumber in a 76-year-old Douglas-fir (Pseudotsuga menziesii, (Mirb., Franco)) experimental stand. The stand consisted of four...

  8. Occurrence of shrubs and herbaceous vegetation after clear cutting old-growth Douglas-fir in the Oregon Cascades. (United States)

    Vern P. Yerkes


    Land managers often express a need for more complete information about the vegetative cover that develops on cutover areas between harvest of old-growth Douglas-fir (Pseudotsuga menziesii) and establishment of a young-growth forest. The composition and density of this cover frequently determines the management techniques that must be used to...

  9. Direct-Digital Modulation (DIDIMO) Transmitter with -156dBc/Hz Rx-Band Noise using FIR structure

    NARCIS (Netherlands)

    Fukuda, Shuichi; Miya, Shinji; Io, Miho; Hamashita, Koichi; Nauta, Bram


    A direct-digital modulation (DIDIMO) transmitter employing 3-taps 10-bit Finite Impulse Response configuration is proposed and evaluated. FIR-DIDIMO provides simultaneous combination of D/A conversion, up-conversion, power control and also quantization noise filtering at arbitrary offset frequency

  10. Influence of thinning on acoustic velocity of Douglas-fir trees in western Washington and western Oregon (United States)

    David G. Briggs; Gonzalo Thienel; Eric C. Turnblom; Eini Lowell; Dennis Dykstra; Robert J. Ross; Xiping Wang; Peter. Carter


    Acoustic velocity was measured with a time-of-flight method on approximately 50 trees in each of five plots from four test sites of a Douglas-fir (Pseudostuga menziesii (Mirb.) Franco) thinning trial. The test sites reflect two age classes, 33 to 35 and 48 to 50 years, with 50-year site index ranging from 35 to 50 m. The acoustic velocity...

  11. Slow and fast pyrolysis of Douglas-fir lignin: Importance of liquid-intermediate formation on the distribution of products

    NARCIS (Netherlands)

    Zhou, Shuai; Pecha, Brennan; van Kuppevelt, Michiel; McDonald, Armando G.; Garcia-Perez, Manuel


    The formation of liquid intermediates and the distribution of products were studied under slow and fast pyrolysis conditions. Results indicate that monomers are formed from lignin oligomeric products during secondary reactions, rather than directly from the native lignin. Lignin from Douglas-fir

  12. Wolfiporia cocos : a potential agent for composting of bioprocessing Douglas-fir wood treated with copper-based preservatives (United States)

    Rodney. De Groot; Bessie. Woodward


    In laboratory experiments, Douglas-fir wood blocks that were treated with copper- based wood preservatives were challenged with two wood decay fungi known to be tolerant of copper. Factors influencing the amount of decay, as determined by loss of weight in the test blocks, were preservative, then fungus. Within those combinations, the relative importance of...


    Global climatic change as expressed by increased CO2 and temperature has the potential for dramatic effects on trees. To determine what its effects may be on Pacific Northwest forests, Douglas-fir (Pseudotsuga menziesii) seedlings were grown in sun-lit controlled environment cham...

  14. Initial thinning effects in 70- to 150-year-old Douglas-fir--western Oregon and Washington. (United States)

    Richard L. Williamson; Frank E. Price


    Vigorous, mature (post-rotation age) Douglas-fir stands will probably exist for another 50 years or more on some properties in western Oregon and Washington. Intermediate harvests in the form of thinnings were analyzed on nine study areas ranging from 70 to 150 years old when thinned. Recoverable cubic-volume growth, averaging 81 percent of normal...

  15. Effects of leader topping and branch pruning on efficiency of Douglas-fir cone harvesting with a tree shaker. (United States)

    D.L. Copes


    In 1983, a study was conducted to evaluate the effects of leader topping and branch pruning on the efficiency to tree shaking to remove Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) cones. Removal efficiency for three topping and pruning treatments averaged 69 percent, whereas for the uncut control treatment it was 62 percent. The treatment...

  16. Age-class differences in shoot photosynthesis and water relations of Fraser fir (Abies fraseri), southern Appalachian Mountains, USA (United States)

    Keith Reinhardt; Daniel M. Johnson; William K. Smith


    Fraser fir (Abies fraseri (Pursh) Poir.) is an endemic tree species found only in refugial mountain-top forests in the southern Appalachian Mountains, USA. Very few studies have investigated the ecophysiology of this species in its natural environment. We measured and compared photosynthetic gas exchange and water relations of understory germinant...

  17. Effect of organic amendments on Douglas-fir transplants grown in fumigated versus non-fumigated soil (United States)

    Nabil Khudduri


    We transplanted one-year old Douglas-fir (Pseudotsuga menziesii var. menziesii) seedlings into compost-amended soil that had either been spring-fumigated with a methyl bromide/chloropicrin combination or left unfumigated. Seedling nutrient, pathology, morphology, and packout measurements were significantly better for those transplanted into fumigated rather than non-...

  18. Height-growth response to climatic changes differs among populations of Douglas-fir: A novel analysis of historic data (United States)

    Laura P. Leites; Andrew P. Robinson; Gerald E. Rehfeldt; John D. Marshall; Nicholas L. Crookston


    Projected climate change will affect existing forests, as substantial changes are predicted to occur during their life spans. Species that have ample intraspecific genetic differentiation, such as Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco), are expected to display population-specific growth responses to climate change. Using a mixed-effects modeling approach,...

  19. Seasonal carbohydrate dynamics and growth in Douglas-fir trees experiencing chronic, fungal-mediated reduction in functional leaf area (United States)

    B. J. Saffell; F. C. Meinzer; D. R. Woodruff; D. C. Shaw; S. L. Voelker; B. Lachenbruch; K. Falk


    Stored non-structural carbohydrates (NSCs) could play an important role in tree survival in the face of a changing climate and associated stress-related mortality. We explored the effects of the stomata-blocking and defoliating fungal disease called Swiss needle cast on Douglas-fir carbohydrate reserves and growth to evaluate the extent to which NSCs can be mobilized...

  20. Calibration of volume and component biomass equations for Douglas-fir and lodgepole pine in Western Oregon forests (United States)

    Krishna P. Poudel; Temesgen. Hailemariam


    Using data from destructively sampled Douglas-fir and lodgepole pine trees, we evaluated the performance of regional volume and component biomass equations in terms of bias and RMSE. The volume and component biomass equations were calibrated using three different adjustment methods that used: (a) a correction factor based on ordinary least square regression through...

  1. MCOL, frontalin, and ethanol: A potential operational trap lure for Douglas-fir beetle in British Columbia (United States)

    B. Staffan Lingren; Daniel R. Miller; J.P. LaFontaine


    The Douglas-fir beetle, Dedroctonus pseudotsugae (Coleoptera: Curculionidae) is a major pest of Douglas-fire, Pseudotsuga menziesii (Mirb.) in British Columbia (Humphreys 1995). An operational trap lure for D. pseudotsugae could be useful in an integrated pest management program to minimize mortality of Douglas-...

  2. Assessing the Ability of Ground-Penetrating Radar to Detect Fungal Decay in Douglas-Fir Beams (United States)

    Christopher Adam Senalik; James Wacker; Xiping Wang; F. Jalinoos


    This paper describes the testing plan and current progress for assessing the efficacy of using ground-penetrating radar (GPR) to detect fungal decay within Douglas-fir beams. Initially, the beams were assessed using a variety of physical, mechanical, and nondestructive evaluation (NDE) test methods including micro-resistance drilling, Janka hardness, ultrasonic...

  3. Simulating historical disturbance regimes and stand structures in old-forest ponderosa pine/Douglas-fir forests (United States)

    Mike Hillis; Vick Applegate; Steve Slaughter; Michael G. Harrington; Helen Smith


    Forest Service land managers, with the collaborative assistance from research, applied a disturbance based restoration strategy to rehabilitate a greatly-altered, high risk Northern Rocky Mountain old-forest ponderosa pine-Douglas-fir stand. Age-class structure and fire history for the site have been documented in two research papers (Arno and others 1995, 1997)....

  4. Impacts of cloud immersion on microclimate, photosynthesis and water relations of fraser fir in a temperate mountain cloud forest (United States)

    Keith Reinhardt; William K. Smith


    The red spruce-Fraser fir ecosystem (Picea rubens Sarg.-Abies fraseri [Pursh] Poir.) of the southern Appalachian mountains is a temperate zone cloud forest immersed in clouds for 30 to 40 percent of a typical summer day, and experiencing immersion on about 65 percent of all days annually. We compared the microclimate,...

  5. Relative family performance and variance structure of open-pollinated Douglas-fir seedlings grown in three competitive environments. (United States)

    J.B. St. Clair; W.T. Adams


    Open-pollinated Douglas-fir (Pseudotsuga menziesii var menziesii (Mirb.) Franco) families were tested in three contrasting competitive environments to test the hypothesis that relative performance as measured by total seedling dry weight is dependent upon distance or genotype of neighbors. The three competitive environments...

  6. Performance and Complexity Analysis of Blind FIR Channel Identification Algorithms Based on Deterministic Maximum Likelihood in SIMO Systems

    DEFF Research Database (Denmark)

    De Carvalho, Elisabeth; Omar, Samir; Slock, Dirk


    We analyze two algorithms that have been introduced previously for Deterministic Maximum Likelihood (DML) blind estimation of multiple FIR channels. The first one is a modification of the Iterative Quadratic ML (IQML) algorithm. IQML gives biased estimates of the channel and performs poorly at lo...

  7. Ecological adaptations in Douglas-fir (Pseudotsuga menziesii var. glauca) populations: I. North Idaho and North-East Washington (United States)

    Gerald E. Rehfeldt


    Growth, phenology and frost tolerance of seedlings from 50 populations of Douglas-fir (Pseudotsuga menziesii var. glauca) were compared in 12 environments. Statistical analyses of six variables (bud burst, bud set, 3-year height, spring and fall frost injuries, and deviation from regression of 3-year height on 2-year height) showed that populations not only differed in...

  8. Fingerprints of a forest fungus: Swiss needle cast, carbon isotopes, carbohydrates, and growth in Douglas-fir (United States)

    Andrea Watts; Frederick Meinzer; Brandy J. Saffell


    Swiss needle cast is caused by a fungus native to the Pacific Northwest. Its host is Douglas-fir, an iconic evergreen tree in the region. The fungus does not kill its host, but it adversely affects the tree's growth. The fungal fruiting bodies block the stomata, small openings on the underside of the needle where carbon dioxide, water vapor, and other gases are...

  9. Removal of non benzidine direct red dye from aqueous solution by using natural sorbents: Beech and silver fir

    Czech Academy of Sciences Publication Activity Database

    Muntean, S.G.; Todea, A.; Bakardjieva, Snejana; Bologa, C.


    Roč. 66, MAR (2017), s. 235-250 ISSN 1944-3994 Institutional support: RVO:61388980 Keywords : Adsorption * Beech * Direct red * Kinetics * Silver fir Subject RIV: CA - Inorganic Chemistry OBOR OECD: Inorganic and nuclear chemistry Impact factor: 1.631, year: 2016

  10. Structure and dynamics in a virgin northern hardwood-spruce-fir forest--the Bowl, New Hampshire (United States)

    Stanley R. Gemborys


    A phytosociological study was conducted in a virgin northern hardwood- spruce-fir forest in the Bowl in the White Mountains of New Hampshire. There is no evidence of fire or human disturbance but hurricane winds were significant in the past. Bray and Curtis ordination was used to develop an XY vegetational mosaic. Differentiating species were Picea rubens and Acer...

  11. Growth of Douglas-fir near equipment trails used for commercial thinning in the Oregon Coast Range. (United States)

    Richard E. Miller; Jim Smith; Paul W. Adams; Harry W. Anderson


    Soil disturbance is a visually apparent result of using heavy equipment to harvest trees. Subsequent consequences for growth of remaining trees, however, are variable and seldom quantified. We measured tree growth 7 and 11 years after thinning of trees in four stands of coast Douglas-fir (Pseudotsuga menziesii var. menziesii(...

  12. Detecting response of Douglas-fir plantations to urea fertilizer at three locations in the Oregon Coast Range. (United States)

    Richard E. Miller; Jim Smith; Harry. Anderson


    Fertilizer trials in coast Douglas-fir (Pseudotsuga menziesii var. menziesii (Mirb.) Franco) in the Oregon Coast Range usually indicate small and statistically nonsignificant response to nitrogen (N) fertilizers. Inherently weak experimental designs of past trials could make them too insensitive to detect growth differences...

  13. Tree growth and soil relations at the 1925 Wind River spacing test in coast Douglas-fir. (United States)

    Richard E. Miller; Donald L. Reukema; Harry W. Anderson


    The 1925 Wind River spacing test is the earliest field trial seeking to determine the most appropriate spacing for planting Douglas-fir. Spacing treatments were not replicated, although individual spacings were subsampled by two to four tree-measurement plots. Previously, greater growth occurred at the wider spacings (10 and 12 ft) than at the closer spacings (4, 5, 6...

  14. Biomass removal, soil compaction, and vegetation control effects on five-year growth of Douglas-fir in coastal Washington. (United States)

    A. Ares; T. Terry; C. Harrington; W. Devine; D. Peter; J. Bailey


    Sustainable forest production requires an understanding of the effects of site disturbance on tree growth and the consequences of soil amelioration and vegetation control practices. We assessed the impacts of biomass removals at harvest, soil compaction and tillage, and vegetation control on early growth of Douglas-fir in coastal Washington. Harvest treatments included...

  15. Soil compaction after yarding of small-diameter Douglas-fir with a small tractor in southwest Oregon. (United States)

    Michael P. Amaranthus; David E. Steinfeld


    This study evaluated the effect on soil bulk density of yarding small-diameter Douglas-fir (Pseudosuga menziesii var. glauca (Beissn.) Franco) with a small tractor. Levels of compaction were measured before yarding and after one trip, three trips, and six trips by the tractor. Bulk densities in the surface (10 cm) and...

  16. Origin matters! Difference in drought tolerance and productivity of coastal Douglas-fir (Pseudotsuga menziesii (Mirb.)) provenances

    NARCIS (Netherlands)

    Eilmann, B.; Vries, de S.M.G.; Ouden, den J.; Mohren, G.M.J.; Sauren, P.; Sass, U.G.W.


    Forests of the future should be resistant to exacerbating climatic conditions, especially to increasing drought, but at the same time provide a sufficient amount and quality of timber. In this context coastal Douglas-fir (Pseudotsuga menziesii (Mirb.)) is a promising species since it remains

  17. A stump-to-truck cost estimating program for cable logging young-growth Douglas-fir (United States)

    Chris B. LeDoux


    WCOST is a computer program designed to estimate the stump-to-truck logging cost of cable logging young-growth Douglas-fir. The program uses data from stand inventory, cruise data, and the logging plan for the tract in question to produce detailed stump-to-truck cost estimates for specific proposed timber sales. These estimates are then used, in combination with...

  18. Influence of a sand soil plough base on the growth of Douglas fir (Pseudotsuga menziesii (MIRB.) FRANCO).

    NARCIS (Netherlands)

    Gruber, F; Nick, L

    The root/shoot growth of eight year old Douglas fir trees (Pseudotsuga menziesii (MIRB.) FRANCO) planted on ploughed agricultured land in the first generation was investigated. One half of the field was 60 cm deeply ploughed before afforestation. The second half was not deeply ploughed and was

  19. Effects of seed source origin on bark thickness of Douglas-fir (Pseudotsuga menziesii) growing in southwestern Germany (United States)

    Ulrich Kohnle; Sebastian Hein; Frank C. Sorensen; Aaron R. Weiskittel


    Provenance-specific variation in bark thickness in Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) is important for accurate volume calculations and might carry ecological implications as well. To investigate variation, diameter at breast height (dbh) and double bark thickness (dbt) were measured in 10 experiments in southwestern Germany (16...

  20. Height-Diameter Models for Mixed-Species Forests Consisting of Spruce, Fir, and Beech

    Directory of Open Access Journals (Sweden)

    Petráš Rudolf


    Full Text Available Height-diameter models define the general relationship between the tree height and diameter at each growth stage of the forest stand. This paper presents generalized height-diameter models for mixed-species forest stands consisting of Norway spruce (Picea abies Karst., Silver fir (Abies alba L., and European beech (Fagus sylvatica L. from Slovakia. The models were derived using two growth functions from the exponential family: the two-parameter Michailoff and three-parameter Korf functions. Generalized height-diameter functions must normally be constrained to pass through the mean stand diameter and height, and then the final growth model has only one or two parameters to be estimated. These “free” parameters are then expressed over the quadratic mean diameter, height and stand age and the final mathematical form of the model is obtained. The study material included 50 long-term experimental plots located in the Western Carpathians. The plots were established 40-50 years ago and have been repeatedly measured at 5 to 10-year intervals. The dataset includes 7,950 height measurements of spruce, 21,661 of fir and 5,794 of beech. As many as 9 regression models were derived for each species. Although the “goodness of fit” of all models showed that they were generally well suited for the data, the best results were obtained for silver fir. The coefficient of determination ranged from 0.946 to 0.948, RMSE (m was in the interval 1.94-1.97 and the bias (m was -0.031 to 0.063. Although slightly imprecise parameter estimation was established for spruce, the estimations of the regression parameters obtained for beech were quite less precise. The coefficient of determination for beech was 0.854-0.860, RMSE (m 2.67-2.72, and the bias (m ranged from -0.144 to -0.056. The majority of models using Korf’s formula produced slightly better estimations than Michailoff’s, and it proved immaterial which estimated parameter was fixed and which parameters

  1. Selecting Populations for Non-Analogous Climate Conditions Using Universal Response Functions: The Case of Douglas-Fir in Central Europe

    National Research Council Canada - National Science Library

    Chakraborty, Debojyoti; Wang, Tongli; Andre, Konrad; Konnert, Monika; Lexer, Manfred J; Matulla, Christoph; Schueler, Silvio


    ... of seed sources and potential growing areas. In the present study, we analyzed the intraspecific variation in climate growth response of Douglas-fir planted within the non-analogous climate conditions of Central and continental Europe...

  2. Basal area growth, carbon isotope discrimination, and intrinsic water use efficiency after fertilization of Douglas-fir in the Oregon Coast Range (United States)

    Many hectares of intensively managed Douglas-fir (Pseudotsuga menziesii Mirb. Franco) stands in western North America are fertilized with nitrogen to increase growth rates. Understanding the mechanisms of response facilitates prioritization of stands for treatment. The objective ...

  3. Fermentative high-titer ethanol production from Douglas-fir forest residue without detoxification using SPORL: high SO2 loading at low temperature (United States)

    Feng Gu; William Gilles; Roland Gleisner; J.Y. Zhu


    This study evaluated high sulfur dioxide (SO2) loading in applying Sulfite Pretreatment to Overcome the Recalcitrance of Lignocelluloses (SPORL) to Douglas-fir forest residue (FS-10) for ethanol production through yeast fermentation. Three pretreatments were conducted at 140

  4. Ground-Based Submillimeter Spectroscopic Cosmological Surveys and Synergies with Space FIR Surveys (United States)

    Spinoglio, Luigi


    To study the dust obscured processes of both star formation and black hole accretion during galaxy evolution and establish their role, as well as their mutual feedback processes, rest frame IR to submillimeter spectroscopy is needed. At these frequencies dust extinction is at its minimum and a variety of atomic and molecular transitions, tracing most astrophysical domains, occur. A large ground based submillimeter telescope with a large field of view and high sensitivity in the TeraHertz domain will pave the way of future FIR space telescope missions, such as SPICA in the late 2020's and the Origins Telescope later. I will present predictions demonstrating the synergies of such instruments to fully understand galaxy evolution, during its obscured phase, which has built most of the stellar populations in galaxies.

  5. FIR Induced Intrinsic Exciton Transitions in GaAs/AlGaAs Superlattices

    DEFF Research Database (Denmark)

    Dremin, A. A.; Timofeev, V. B.; Birkedal, Dan


    Intrinsic transitions of confined excitons in GaAs/AlGaAs superlattices with different barrier widths have been studied with the use of resonant far-infrared absorption under variation of magnetic field perpendicular and tilted with respect to the growth directions. Few resonances have been obser...... observed in spectra induced by FIR radiation by monitoring the changes of the ground state excitonic photoluminescence. The dominant resonances are the electron cyclotron resonance and the intrinsic 1s --> 2p. intrawell transitions of the heavy-hole excitons.......Intrinsic transitions of confined excitons in GaAs/AlGaAs superlattices with different barrier widths have been studied with the use of resonant far-infrared absorption under variation of magnetic field perpendicular and tilted with respect to the growth directions. Few resonances have been...

  6. Transverse Emittance Measurements In High-power Fir Fel Energy-recovery Linac

    CERN Document Server

    Bolotin, V P; Matveenko, A N; Shevchenko, O A; Vinokurov, N A


    50 MeV accelerator-recirculator of Siberian Center for Photochemical Research has been designed to drive the FIR FEL with an average power of up to 10 kW in the wavelengths region from 5 to 200 μm. The first stage with the beam energy of 14 MeV was put into operation and laser power of about 200 W on 150 μm was achieved recently [1]. Transverse emittance measurements are carried out online in a number of locations along the beam-line. OTR screens and video-cameras are used to capture the beam shape images, video signal is digitized by a frame grabber and the pictures are processed further. Online measurements allow the accelerator parameters to be optimized to minimize the emittance growth, which is essential to the FEL operation. The transverse emittance measurements system and data processing techniques are presented in this paper.

  7. EuroFIR eBASIS: application for health claims submissions and evaluations

    DEFF Research Database (Denmark)

    Kiely, M.; Black, L.J.; Plumb, J.


    Background: The European Food Information Resource (EuroFIR) network has established the eBASIS (Bioactive Substances in Food Information System) online food composition and biological effects database for plant-derived bioactive compounds (phytochemicals). On the basis of submitted evidence...... of phytochemicals. Currently included are 4750 records from 445 studies providing data on 56 validated biomarkers, mainly relating to cardio-metabolic and bone health outcomes. The data cover 144 bioactive compounds from 17 compound classes. Using the EFSA Register of Questions and the database of general function...... health claims, we identified claims relating to phytochemicals made under articles 13.1, 13.5 and 14 and compared them with the eBASIS database to identify overlap between them. Results: The EFSA online health claims database contains 4240 submissions under article 13.1, of which 2157 pertain to plants...

  8. A low-power asynchronous data-path for a FIR filter bank

    DEFF Research Database (Denmark)

    Nielsen, Lars Skovby; Sparsø, Jens


    ways: by reducing the switching activity in the circuit, and by applying adaptive scaling of the supply voltage, in order to exploit the fact that the average case latency as 2-3 times better than the worst case. The paper reports on a study of properties of real life data, and discusses......This paper describes a number of design issues relating to the implementation of low-power asynchronous signal processing circuits. Specifically, the paper addresses the design of a dedicated processor structure that implements an audio FIR filter bank which is part of an industrial application....... The algorithm requires a fixed number of steps and the moderate speed requirement allows a sequential implementation. The latter, in combination with a huge predominance of numerically small data values in the input data stream, is the key to a low-power asynchronous implementation. Power is minimized in two...

  9. Round-Off Noise of Multiplicative FIR Filters Implemented on an FPGA Platform

    Directory of Open Access Journals (Sweden)

    Jean-Jacques Vandenbussche


    Full Text Available The paper analyzes the effects of round-off noise on Multiplicative Finite Impulse Response (MFIR filters used to approximate the behavior of pole filters. General expressions to calculate the signal to round-off noise ratio of a cascade structure of Finite Impulse Response (FIR filters are obtained and applied on the special case of MFIR filters. The analysis is based on fixed-point implementations, which are most common in digital signal processing algorithms implemented in Field-Programmable Gate-Array (FPGA technology. Three well known scaling methods, i.e., L2 bound; infinity bound and absolute bound scaling are considered and compared. The paper shows that the ordering of the MFIR stages, in combination with the scaling methods, have an important impact on the round-off noise. An optimal ordering of the stages for a chosen scaling method can improve the round-off noise performance by 20 dB.

  10. Modeling the hemodynamic response in fMRI using smooth FIR filters

    DEFF Research Database (Denmark)

    Goutte, Cyril; Nielsen, Finn Årup; Hansen, Lars Kai


    -parametric approach based on finite impulse response (FIR) filters. In order to cope with the increase in the number of degrees of freedom, the authors introduce a Gaussian process prior on the filter parameters. They show how to carry on the analysis by incorporating prior knowledge on the filters, optimizing hyper......Modeling the hemodynamic response in functional magnetic resonance (fMRI) experiments is an important aspect of the analysis of functional neuroimages. This has been done in the past using parametric response function, from a limited family. In this contribution, the authors adopt a semi......-parameters using the evidence framework, or sampling using a Markov Chain Monte Carlo (MCMC) approach. The authors present a comparison of their model with standard hemodynamic response kernels on simulated data, and perform a full analysis of data acquired during an experiment involving visual stimulation....

  11. Caribou in British Columbia: A 1996 status report

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    Douglas C. Heard


    Full Text Available Caribou (Rangifer tarandus in British Columbia are classified into mountain, northern and boreal ecotypes based on behavioural and ecological characteristics. We recognized 12 mountain caribou herds, 27 northern caribou herds, and an area occupied by low density boreal caribou dispersed in the boreal forests of the northeast portion of the province. Abundance estimates were usually based on attempts at total counts made from the air. Trends were based on repeated population estimates or the difference between recruitment and mortality rates for each herd. In 1996 there were approximately 18 000 caribou in British Columbia; 2300 mountain and 15 600 northern and boreal. These estimates suggest a slight increase in the numbers of both ecotypes over the last 18 years. Fifteen percent of the herds were reportedly increasing, 10% were decreasing, 31% were stable, but for 44% of the herds the trend was unknown. Historically caribou were found throughout 8 of the 14 biogeoclimatic zones in B.C. Caribou are now rarely found in the Sub-Boreal Spruce zone, likely due to increased predation from wolves that increased in response to increasing moose numbers. Ranges of several herds in the Engelmann Spruce — Subalpine Fir and Alpine Tundra zones of south-eastern British Columbia are also reduced relative to historic conditions, probably because of habitat loss, habitat fragmentation, predation and hunting. Forest harvesting represents the greatest threat to caribou habitat and current research focuses on the mitigation of forest harvesting impacts.

  12. Thinning effect on photosynthesis depends on needle ages in a Chinese fir (Cunninghamia lanceolata) plantation. (United States)

    Li, Ren-Shan; Yang, Qing-Peng; Zhang, Wei-Dong; Zheng, Wen-Hui; Chi, Yong-Gang; Xu, Ming; Fang, Yun-Ting; Gessler, Arthur; Li, Mai-He; Wang, Si-Long


    Canopies in evergreen coniferous plantations often consist of various-aged needles. However, the effect of needle age on the photosynthetic responses to thinning remains ambiguous. Photosynthetic responses of different-aged needles to thinning were investigated in a Chinese fir (Cunninghamia lanceolata) plantation. A dual isotope approach [simultaneous measurements of stable carbon (δ(13)C) and oxygen (δ(18)O) isotopes] was employed to distinguish between biochemical and stomatal limitations to photosynthesis. Our results showed that increases in net photosynthesis rates upon thinning only occurred in the current-year and one-year-old needles, and not in the two- to four-year-old needles. The increased δ(13)C and declined δ(18)O in current year needles of trees from thinned stands indicated that both the photosynthetic capacity and stomatal conductance resulted in increasing photosynthesis. In one-year-old needles of trees from thinned stands, an increased needle δ(13)C and a constant needle δ(18)O were observed, indicating the photosynthetic capacity rather than stomatal conductance contributed to the increasing photosynthesis. The higher water-soluble nitrogen content in current-year and one-year-old needles in thinned trees also supported that the photosynthetic capacity plays an important role in the enhancement of photosynthesis. In contrast, the δ(13)C, δ(18)O and water-soluble nitrogen in the two- to four-year-old needles were not significantly different between the control and thinned trees. Thus, the thinning effect on photosynthesis depends on needle age in a Chinese fir plantation. Our results highlight that the different responses of different-aged needles to thinning have to be taken into account for understanding and modelling ecosystem responses to management, especially under the expected environmental changes in future. Copyright © 2016 Elsevier B.V. All rights reserved.

  13. Changes in the silver fir forest vegetation 50 years after cessation of active management

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    Beata Woziwoda


    Full Text Available Knowledge of the vegetation and the monitoring of its changes in preserved areas is an essential part of effective conservation policy and management. The aim of this study was to assess the effectiveness of traditional methods of conservation of silver fir forests. The study analyses the changes in the structure and species composition of a temperate forest excluded from the commercial silvicultural management for 50 years, and since then protected as a nature reserve. The study is based on a comparative analysis of phytosociological reléves made on permanent plots in 1961, 1982, 1994 and 2011. PCA and ecological indicator values were analyzed, as well as characteristic species based on an indicator value (IndVal index. Results revealed significant and dynamic changes in the forest structure and composition. The mixed coniferous-broadleaved forest with Abies alba and diverse ground flora, considered in the 1960s as valuable and worthy of conservation, was found to have been anthropogenically transformed and unstable. Significant reduction in the human impact was followed by spontaneous regeneration of oak–hornbeam forest. However, the directional process of changes in vegetation was modified by such silvicultural treatments as selective cutting of trees and gap creation, all intended for silver fir maintenance. The results show that Carpinus betulus effectively outcompeted Pinus sylvestris, Picea abies, Quercus robur and A. alba. Changes in the forest overstory and understory caused temporal changes in the habitat conditions reflected in changes in the ground vegetation composition. The proportion of light-demanding and oligotrophic species significantly decreased, while the contribution of species with a wide ecological amplitude, i.e. more shade-tolerant and nutrient-demanding – increased. The share of A. alba was reduced. Species defined in this study as most valuable, should be actively protected, or selection of conservation targets

  14. Transmission of a Gammabaculovirus within Cohorts of Balsam Fir Sawfly (Neodiprion abietis Larvae

    Directory of Open Access Journals (Sweden)

    Christopher J. Lucarotti


    Full Text Available Nucleopolyhedroviruses (NPV: Gammabaculovirus: Baculoviridae of diprionid sawflies (Diprionidae: Hymenoptera are highly host specific and only infect the midgut epithelium. While still alive, infected sawfly larvae excrete NPV-laden diarrhea that contaminates food sources. The diarrhea can then be consumed by conspecific larvae, resulting in rapid horizontal transmission of the virus. To better understand the efficacy of Gammabaculovirus-based biological control products, the horizontal spread of such a virus (NeabNPV within cohorts of balsam fir sawfly (Neodiprion abietis larvae was studied by introducing NeabNPV-treated larvae into single-cohort groups at densities similar to those observed during the increasing (field study and peak (laboratory study phases of an outbreak. In field studies (~200 N. abietis larvae/m2 of balsam fir (Abies balsamea foliage, NeabNPV-induced mortality increased positively in a density-dependent manner, from 23% (in control groups to 51% with the addition of one first-instar NeabNPV-treated larva, to 84% with 10 first–instar-treated larvae. Mortality was 60% and 63% when one or 10 NeabNPV-treated third-instar larva(e, respectively, were introduced into groups. Slightly higher levels of NeabNPV-induced mortality occurring when NeabNPV-treated larvae were introduced into first- rather than third-instar cohorts suggests that early instars are more susceptible to the virus. In the laboratory (~1330 N. abietis larvae/ m2 of foliage, NeabNPV-caused mortality increased from 20% in control groups to over 80% with the introduction of one, five or 10 NeabNPV-treated larvae into treatment groups of first-instar larvae.

  15. Implementasi Filter Finite Impulse Response (FIR Window Hamming dan Blackman menggunakan DSK TMS320C6713

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    Full Text Available ABSTRAK Filter didefinisikan sebagai proses atau rangkaian yang melewatkan pita frekuensi tertentu yang diinginkan dan meredam pita frekuensi lainnya. Salah satu metode perancangan filter digital Finite Impulse Response (FIR adalah metode windowing. Dalam penelitian ini digunakan jenis window Hamming dan Blackman. Simulasi dilakukan dengan menggunakan software Matlab dengan memasukan frekuensi passband, frekuensi stopband, ripple passband, dan stopband attenuation. Dengan frekuensi sampling sebesar 15000 Hz, frekuensi passband sebesar 3000 Hz, frekuensi stopband sebesar 5000 Hz. Setelah simulasi dilakukan implementasi filter dengan parameter yang sama menggunakan DSK TMS 320C6713 dengan bantuan software CCS. Simulasi dan implementasi dilakukan pada semua band frekuensi. Hasil pengujian terhadap implementasi filter adalah respon magnitude, frekuensi cut-off, bandwidth, dan faktor kualitas dengan hasil simulasi tidak menunjukkan perbedaan yang signifikan. Kata kunci: filter digital, windowing, Hamming, Blackman, frekuensi cut-off . ABSTRACT Filter is defined as a process or series that skip certain desired frequency band and other frequency bands drown. One method of designing a digital filter Finite Impulse Response (FIR is a windowing method. This study used the type of window Hamming and Blackman. Simulations performed using Matlab software by inserting a frequency passband, stopband frequency, passband ripple, and stopband attenuation. With a sampling frequency of 15,000 Hz, a frequency of 3000 Hz passband, stopband frequency of 5000 Hz. After the simulation is completed, implementation of the filter with the same parameters using TMS 320C6713 DSK with the help of software CCS. Simulation and implmentasi performed on all frequency bands. The test results of the implementation of the filter is the Magnitude response, the cut-off frequency, bandwidth, and quality factor with simulation results showed no significant difference. Keywords: digital

  16. Les Gorges de Trévans dans le front subalpin – Un site exceptionnel des Alpes de Haute-Provence

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    Jean Nicod


    Full Text Available Les canyons de Trévans sont incisés dans des unités calcaires jurassiques, dans la zone frontale subalpine, proche du piémont de Valensole. Ces reliefs sont en rapport avec les phases néotectoniques. Ce secteur fait principalement partie de la forêt domaniale du Montdenier, avec des écosystèmes montagnards et supra-méditerranéens, bois de hêtres et de chênes pubescents, large extension des brousses et d'importants reboisements en pins noirs. Les hautes surfaces conservent des traces d'un ancien aplanissement et des sols résiduels paléokarstiques. Sur les pentes, sous des escarpements démembrés, les éboulis et convois de blocs témoignent des processus périglaciaires hérités et, localement, des évènements séismiques. De nombreux processus dynamiques s'observent dans les canyons : éboulements provenant des parois affectées des effets de détente, coups de gouge et marmites de géant dans leur fond excavé par les écoulements turbulents dans les cascades… et le Pont de Tuf.The canyon system of Trevans has cut the blocks of the jurassic limestones, in subalpine over thrust front, near the Valensole piedmont. These landforms are in relationship with the neotectonic movements. This area belongs to the State Forest of Montdenier, woodland of various mountain and supra-mediterranean ecosystems, with beeches and white oaks Quercus pubescens, large extent of bush and important reforestation in black pines (Pinus nigra. The high surfaces preserve some relics of the old planation and paleokarstic forms and residual soils. On the slopes, under the break-up escarpments, the screes and landslides give the part of the periglacial processes and, locally, of the seismic events. Numerous dynamic processes occur in the canyons: rock-slides, in relationship with open fractures, scallops and pot-holes in the bottom excavated by the turbulent flows in the waterfalls of the creeks and, only case, a travertine bridge.

  17. Multiphase clock generators with controlled clock impulse width for programmable high order rotator SC FIR filters realized in 0.35 μm CMOS technology (United States)

    Dlugosz, Rafal; Pawlowski, Pawel; Dabrowski, Adam


    Complexity of clock generator is one of the most important parameters in the design and optimization of switched-capacitor (SC) finite impulse response (FIR) filters. There are different SC FIR filter architectures. Some of them need a simple clock generator but the others require a quite complicated multiphase clock system. In the latter case an external clock system (i.e., outside the integrated circuit) is unrealistic because of a great number of the required external pins. We have implemented various SC FIR filter architectures together with complex internal clock generators in the CMOS 0.8 μm and 0.35 μm technologies. One of the most important problems in the design process was the optimization of waveforms and widths of the clock impulses. SC FIR filters are very sensitive to parameters of clock systems. Thus the clock generators must be designed very precisely. We demonstrate results of the design of the 64-phase clock generator for a programmable rotator SC FIR filter. In our approach the width of the clock impulses is controlled by two external signals. This is a very convenient solution, because optimization of the clock impulses, which was difficult in other approaches, is currently much easier. The internal clock generator area is ca. 0.15 mm2 in the CMOS 0.35 μm technology, i.e., only 7 % of the entire SC FIR filter chip area.

  18. Comparison of Wood Quality of Douglas Fir and Spruce from Afforested Agricultural Land and Permanent Forest Land in the Czech Republic

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    Aleš Zeidler


    Full Text Available This study compares the quality of wood from two distinct sites in the Czech Republic—from former afforested agricultural land and forest land. We compared the properties of Norway spruce wood (Picea abies Karst. and Scots pine wood (Pinus sylvestris L., the most important domestic tree species, to Douglas fir (Pseudotsuga menziesii (Mirbel Franco, a North American tree species and a potential substitute for the domestic spruce. Wood density, modulus of elasticity (MOE, modulus of rupture (MOR and impact bending strength were the properties tested that were used for comparing tree species. Without taking into consideration the site, the highest density values from the tested tree species were obtained for Douglas fir (0.568 g·cm−3, followed by the pine (0.508 g·cm−3 and the spruce (0.463 g·cm−3. The Douglas fir also dominated in the remaining assessed properties, whilst the influence of site was not confirmed, with the exception of MOE and MOR, and only for the Douglas fir wood, wherein higher values were obtained for forest land. In terms assessed Douglas fir properties, it exceeds the domestic softwoods and represents a possible suitable replacement for them. The site only plays a role in terms of the Douglas fir, and only for certain properties.

  19. Nevada Monitoring System to Assess Climate Variability and Change (United States)

    Devitt, D. A.; Arnone, J.; Biondi, F.; Fenstermaker, L. F.; Saito, L.; Young, M.; Riddle, B.; Strachan, S. D.; Bird, B.; McCurdy, G.; Lyles, B. F.


    The Nevada System of Higher Education (University of Nevada Las Vegas, University of Nevada Reno and the Desert Research Institute) was awarded a multiyear NSF EPSCoR grant to support infrastructure associated with regional climate change research. The overall project is comprised of 5 components: education, cyberinfrastructure, policy, climate modeling and water/ecology. The water and ecology components are using their infrastructure funding for the assessment of climate variability and change on ecosystem function and hydrologic services. A series of 10 m tall towers are under construction and are being equipped with a wide array of sensors to monitor atmospheric, soil and plant parameters over time. The towers are located within the Mojave and Great Basin Deserts in two transects; the Mojave Desert transect is located in the southern Nevada Sheep Mountain Range and the Great Basin transect is located in the east central Nevada Snake Mountain Range. The towers are centrally positioned in well-defined vegetation zones. In southern Nevada these zones are represented by the following plant species: Creosote/Bursage (Creosotebush scrub zone); Blackbrush/Joshua Tree (Blackbrush zone); Pinyon/ Juniper (pygmy conifer zone), Ponderosa Pine (montane zone) and Bristlecone Pine (subalpine zone). The Snake Mountain transect incorporates the eastern and western valleys on both sides of the mountain range. The vegetation zones are represented by: Greasewood and mixed shrub (salt desert zone); Big Sage (sagebrush zone); Pinyon/Juniper (pygmy conifer zone); White/Douglas Fir, Ponderosa Pine and Aspen (montane zone); and Bristlecone/Limber Pine and Engelmann Spruce (subalpine zone). We are currently in the third year of funding with a goal of having the majority of towers fully operational by winter 2010. In close collaboration with our cyberinfrastructure component team, all data acquired from the transect monitoring stations will be made available to other researchers and the

  20. Net ecosystem exchange of carbon dioxide and evapotranspiration response of a high elevation Rocky Mountain (Wyoming, USA) forest to a bark beetle epidemic (United States)

    Frank, J. M.; Massman, W. J.; Ewers, B. E.


    Bark beetle epidemics have caused major disturbance in the forests of western North America where significant tree mortality alters the balance of ecosystem photosynthesis, carbon balance, and water exchange. In this study we investigate the change in the growing-season light-response of net ecosystem exchange of carbon dioxide (NEE) and evapotranspiration (ET) in a high elevation Rocky Mountain forest over the three years preceding and three years following a bark beetle outbreak. The GLEES AmeriFlux site (southeastern Wyoming, USA) is located in a high elevation subalpine forest dominated by Engelmann spruce (Picea engelmannii) and subalpine fir (Abies lasiocarpa) and recently experienced an epidemic of spruce beetle (Dendroctonus rufipennis). The peak beetle outbreak occurred in 2008, and has impacted 35% of the stems and 90% of the basal area of Engelmann spruce, which accounts for 30% of the trees and 70% of the basal area of the forest. Two semi-empirical light response curves for eddy-covariance carbon flux were compared, with a logistic sigmoid performing better because of residual bias than a rectangular hyperbola (Michaelis-Menten) at estimating the quantum yield of photosynthesis. In the first two years after the peak beetle outbreak the original quantum yield of 0.015 mol mol-1 was reduced by 25%. By the third year it was reduced by a half, which was composed of declines of 45% in the ecosystem's responses to diffuse radiation and 60% to direct radiation. The light-saturated rate of photosynthesis decreased by 10% in the first two years post outbreak, and fell by 40% in the third year. After the peak outbreak, the cumulative NEE over the growing season was reduced by over a half from a sink of 185 gC m-2 to 80 gC m-2, and by the third year it was reduced to near zero, or carbon neutral. The change in the ET response to light was similar in all years after the peak outbreak where the slope of the response curve was decreased by 25%. This led to a

  1. Hydrology of the North Klondike River: carbon export, water balance and inter-annual climate influences within a sub-alpine permafrost catchment. (United States)

    Lapp, Anthony; Clark, Ian; Macumber, Andrew; Patterson, Tim


    Arctic and sub-arctic watersheds are undergoing significant changes due to recent climate warming and degrading permafrost, engendering enhanced monitoring of arctic rivers. Smaller catchments provide understanding of discharge, solute flux and groundwater recharge at the process level that contributes to an understanding of how larger arctic watersheds are responding to climate change. The North Klondike River, located in west central Yukon, is a sub-alpine permafrost catchment, which maintains an active hydrological monitoring station with a record of >40 years. In addition to being able to monitor intra-annual variability, this data set allows for more complex analysis of streamflow records. Streamflow data, geochemistry and stable isotope data for 2014 show a groundwater-dominated system, predominantly recharged during periods of snowmelt. Radiocarbon is shown to be a valuable tracer of soil zone recharge processes and carbon sources. Winter groundwater baseflow contributes 20 % of total annual discharge, and accounts for up to 50 % of total river discharge during the spring and summer months. Although total stream discharge remains unchanged, mean annual groundwater baseflow has increased over the 40-year monitoring period. Wavelet analysis reveals a catchment that responds to El Niño and longer solar cycles, as well as climatic shifts such as the Pacific Decadal Oscillation. Dedicated to Professor Peter Fritz on the occasion of his 80th birthday.

  2. Dynamics of development and variability of surface degradation in the subalpine and alpine zones (an example from the Velká Fatra Mts., Slovakia

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    Lepeška Tomáš


    Full Text Available In the last five centuries, the inappropriate management of the Vel’ká Fatra Mts. sub-alpine and alpine areas has led to the development of different forms of surface destruction. For evaluation of the dynamics and variability of surface degradation the territory of the Hornojelenská valley was chosen. It is a significant avalanche area. It has clearly been destroyed by avalanches, water erosion and cryogenic erosion as well as anthropo-zoogenic processes. The forms of destruction were mapped on a scale of 1:200 based on the aerial photographs and satellite images taken in 1961, 2003, 2009 and 2012. The total area of degradative morphogenetic forms (DMF in 1961 was 5.5780 ha, 4.0650 ha in 2003, 4.5752 ha in 2009 and 4.9431 ha in 2012. The DMF reached its peak in 1961. In the mid-1960s, there were ambitions to reforest the highest areas of the study area that led to the decrease of DMF and the development of vegetation. The present exogenous geomorphologic processes are causing a gradual increase of the total destructed area.

  3. Using the Resistograph®to distinguish different types of wood rot on living silver fir in Molise (Italy

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    Lasserre B


    Full Text Available he study was performed in two silver-fir forests (Abies alba Mill. located in Alto Molise, Province of Isernia: Collemeluccio, near Pescolanciano and Abeti Soprani near Capracotta. The aim of this work was to distinguish different types of wood rot on living silver fir individuals by using the Resistograph® (IML-RESI E400, a device that allows to estimate the variation of wood density by measuring the resistance to micro-perforation. The occurrence of different types of wood rot (white rot and brown rot in living trees was pointed out and discriminated by the device. In the detected deteriorated zones, fungal pathogens and decomposers were isolated and identified, causing either white (Phellinus hartigii, Ganoderma adspersum, Heterobasidion abietinum and Armillaria ostoyae or brown rot (Fomitopsis pinicola.

  4. Optimal design of FIR high pass filter based on L1 error approximation using real coded genetic algorithm

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    Apoorva Aggarwal


    Full Text Available In this paper, an optimal design of linear phase digital finite impulse response (FIR highpass (HP filter using the L1-norm based real-coded genetic algorithm (RCGA is investigated. A novel fitness function based on L1 norm is adopted to enhance the design accuracy. Optimized filter coefficients are obtained by defining the filter objective function in L1 sense using RCGA. Simulation analysis unveils that the performance of the RCGA adopting this fitness function is better in terms of signal attenuation ability of the filter, flatter passband and the convergence rate. Observations are made on the percentage improvement of this algorithm over the gradient-based L1 optimization approach on various factors by a large amount. It is concluded that RCGA leads to the best solution under specified parameters for the FIR filter design on account of slight unnoticeable higher transition width.

  5. Some observations of wood density and anatomical properties in a Douglas-fir sample with suppressed growth (United States)

    J.Y. Zhu; David W. Vahey; C. Tim Scott


    This study used ring width correlations to examine the effects of tree-growth suppression on within-tree local wood density and tracheid anatomical properties. A wood core sample was taken from a 70-yr-old Douglas-fir that grew under various degrees of suppression in a natural forest setting. SilviScan and an imaging technique were used to obtain wood density and...

  6. Do mycorrhizal network benefits to survival and growth of interior Douglas-fir seedlings increase with soil moisture stress? (United States)

    Bingham, Marcus A; Simard, Suzanne W


    Facilitation of tree establishment by ectomycorrhizal (EM) networks (MNs) may become increasingly important as drought stress increases with climate change in some forested regions of North America. The objective of this study was to determine (1) whether temperature, CO(2) concentration ([CO(2)]), soil moisture, and MNs interact to affect plant establishment success, such that MNs facilitate establishment when plants are the most water stressed, and (2) whether transfer of C and water between plants through MNs plays a role in this. We established interior Douglas-fir (Pseudotsuga menziesiivar.glauca) seedlings in root boxes with and without the potential to form MNs with nearby conspecific seedlings that had consistent access to water via their taproots. We varied temperature, [CO(2)], and soil moisture in growth chambers. Douglas-fir seedling survival increased when the potential existed to form an MN. Growth increased with MN potential under the driest soil conditions, but decreased with temperature at 800 ppm [CO(2)]. Transfer of (13)C to receiver seedlings was unaffected by potential to form an MN with donor seedlings, but deuterated water (D(2)O) transfer increased with MN potential under ambient [CO(2)]. Chlorophyll fluorescence was reduced when seedlings had the potential to form an MN under high [CO(2)] and cool temperatures. We conclude that Douglas-fir seedling establishment in laboratory conditions is facilitated by MN potential where Douglas-fir seedlings have consistent access to water. Moreover, this facilitation appears to increase as water stress potential increases and water transfer via networks may play a role in this. These results suggest that conservation of MN potential may be important to forest regeneration where drought stress increases with climate change.

  7. Collective excitations in liquid DMSO : FIR spectrum, Low frequency vibrational density of states and ultrafast dipolar solvation dynamics


    Hazra, Milan; Bagchi, Biman


    Valuable dynamical and structural information about neat liquid DMSO at ambient conditions can be obtained through study of low frequency vibrations in the far infrared (FIR), that is, terahertz regime. For DMSO, collective excitations as well as single molecule stretches and bends have been measured by different kinds of experiments such as OHD-RIKES and terahertz spectroscopy. In the present work we investigate the intermolecular vibrational spectrum of DMSO through three different computat...

  8. Monitoring the Epidemic of Bark Beetles Determined in Uludag Fir Stands in Ilgaz (Derbent and Doruk) via Landsat Satellite Images


    KONDUR, Yalçın; BİLGİLİ, Bayram Cemil; Şimşek, Ziya; ÖNER, Nuri


    Results of the studies carried out in Ilgaz (Derbent and Doruk) in 1999-2000 showed that Cryphalus piceae (Ratz.), Ips acuminatus (Gyll.), Orthotomicus erosus (Woll.), Pityogenes quadridens (Hartig), Ips (Pityokteines) curvidens (Germ.), Tryphodendron lineatus (Oliver) (Scolytidae), Pissodes piceae (Illig.) (Curculionidae) were important insect species where the Uludag fir (Abies nordmanniana subsp. bornmulleriana Mattf.) is the dominant tree species at altitudes between 1,700 and 2,000 m, mo...

  9. Plant-pest interactions in time and space: A Douglas-fir bark beetle outbreak as a case study


    Powers, J. S.; Sollins, P.; Harmon, M. E.; Jones, J. A.


    A conceptual model of Douglas-fir bark beetle (Dendroctonus pseudotsugae) dynamics and associated host tree mortality across multiple spatial and temporal scales was developed, then used to guide a study of the association between the occurrence of beetle-killed trees and factors that might render trees more susceptible to attack. Longterm records of beetle kill showed that beetle epidemics were associated with windstorms and drought at statewide and local spatial scales. At the landscape sca...

  10. [Modified Mechanism of Cell Walls from Chinese Fir Treated with Low-Molecular-Weight Phenol Formaldehyde Resin]. (United States)

    Huang, Yan-hui; Fei, Ben-hua; Zhao, Rong-jun


    Study on the modified mechanism of wood cell walls, it is very important for improving treatment reagents, optimizing treatment technology, and enhancing wood density, mechanical properties, dimensional stability, and so on. Samples of plantation Chinese fir were treated gradually with synthesized water-soluble low-molecular-weight phenol formaldehyde (PF) resins under vacuum and pressure. The correlated physical and chemical properties of the treated and untreated reference samples were determined by X-ray diffractometer (XRD), Fourier transform infrared spectrometer (FTIR), and nuclear magnetic resonance spectrometer(NMR) (Using method of Cross Polarization/Magic Angle Spinning for continuous testing) with high precision and resolution. The results showed that, after treated with water-soluble low-molecular-weight PF resin, the average values of crystallinity from the treated samples were decreased obviously, and the average reduction rate was 12.67%, 11.91% and 6.26%, respectively. Comparing water-soluble, low-molecular-weight PF resin modified Chinese fir with untreated reference samples, no new chemical shifts and characteristic peaks of functional groups from esters, ethers, etc. were present by using FTIR and ¹³C NMR spectrum. It was considered that there was no distinct chemical reaction between the water-soluble low-molecular-weight PF resin and Chinese Fir cell walls. But water-soluble low-molecular-weight PF resin could enter into the structure relatively loose, large size spaces, relatively area large amorphous regions in cell walls of Chinese fir tracheids, and form physical filling, which resulting in the decreasing of relative crystallinity. This study has important reference value for the development of new wood modification reagents and the optimization of wood modification process. The findings also provide important theoretical foundation for further proving the modification mechanisms of wood cell walls and enriching the modified theories of

  11. Transcriptome characteristics and six alternative expressed genes positively correlated with the phase transition of annual cambial activities in Chinese Fir (Cunninghamia lanceolata (Lamb. Hook.

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    Zhanjun Wang

    Full Text Available BACKGROUND: The molecular mechanisms that govern cambial activity in angiosperms are well established, but little is known about these molecular mechanisms in gymnosperms. Chinese fir (Cunninghamia lanceolata (Lamb. Hook, a diploid (2n  = 2x  = 22 gymnosperm, is one of the most important industrial and commercial timber species in China. Here, we performed transcriptome sequencing to identify the repertoire of genes expressed in cambium tissue of Chinese fir. METHODOLOGY/PRINCIPAL FINDINGS: Based on previous studies, the four stage-specific cambial tissues of Chinese fir were defined using transmission electron microscopy (TEM. In total, 20 million sequencing reads (3.6 Gb were obtained using Illumina sequencing from Chinese fir cambium tissue collected at active growth stage, with a mean length of 131 bp and a N50 of 90 bp. SOAPdenovo software was used to assemble 62,895 unigenes. These unigenes were further functionally annotated by comparing their sequences to public protein databases. Expression analysis revealed that the altered expression of six homologous genes (ClWOX1, ClWOX4, ClCLV1-like, ClCLV-like, ClCLE12, and ClPIN1-like correlated positively with changes in cambial activities; moreover, these six genes might be directly involved in cambial function in Chinese fir. Further, the full-length cDNAs and DNAs for ClWOX1 and ClWOX4 were cloned and analyzed. CONCLUSIONS: In this study, a large number of tissue/stage-specific unigene sequences were generated from the active growth stage of Chinese fir cambium. Transcriptome sequencing of Chinese fir not only provides extensive genetic resources for understanding the molecular mechanisms underlying cambial activities in Chinese fir, but also is expected to be an important foundation for future genetic studies of Chinese fir. This study indicates that ClWOX1 and ClWOX4 could be possible reverse genetic target genes for revealing the molecular mechanisms of cambial activities in

  12. Douglas fir (pseudotsuga menziesii) plantlets responses to as, PB, and sb-contaminated soils from former mines. (United States)

    Bonet, Amandine; Pascaud, Grégoire; Faugeron, Céline; Soubrand, Marilyne; Joussein, Emmanuel; Gloaguen, Vincent; Saladin, Gaëlle


    Phytoremediation of metalloids by conifers is not widely studied although they may be relevant for several contaminated sites, especially those located in cold areas and sometimes under dry climates. Here, seeds of Douglas fir were sown in greenhouse on three soils collected in two French former mines: a gold mine (soils L1 and L2) and a lead and silver mine (soil P). These soils are highly contaminated by Pb, As, and Sb at different concentrations. Plants were harvested after ten weeks. Growth parameters, primary metabolite content, and shoot and root ionomes were determined. Douglas firs grown on the soils L1 and P had a lower biomass than controls and a higher oxidation status whereas those grown on the soil L2 exhibited a more developed root system and only slight modifications of carbon and nitrogen nutrition. Based on trace element (TE) concentrations in shoots and roots and their translocation factor (TF), Douglas fir could be a relevant candidate for As phytoextraction (0.8 g. kg(-1) dry weight in shoots and a TF of 1.1) and may be used to phytostabilize Pb and Sb (8.8 g and 127 mg. kg(-1) in roots for Pb and Sb, respectively, and TF lower than 0.1).

  13. Mineral status of plants and soil in silver fir stands of the Silesian and Żywiec Beskidy (Western Carpathians

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    Borgulat Jacek


    Full Text Available For last decades the Silesian and Żywiec Beskidy have been affected by different types of anthropopression. Up to the nineties of the 20th century, forests of the area were affected by a high level of air pollutants transported mainly from the Upper Silesian and Ostrava industrial regions. From 2004 intensive felling and skidding works in declining forest stands combined with their reconstruction have been carried out. Among different species silver fir has been introduced on a large scale. In the paper the results of investigation of plants and soil carried out 7 sites located in Beskidy Mountains (49°41’6”N, 19°1’48”E are presented. Contents of K, Mg, Ca and N in needles of silver fir (Abies alba Mill. and leaves of bilberry (Vaccinium myrtillus L. were determined which was accompanied by a chemical characteristics of forest topsoil from the same sites. Moreover, in both environment compartments, the concentration of S, Pb, Cd, and Zn was also determined to assess the level of anthropopression in individual sites. In general, the obtained results show relatively good mineral status of plants. Some disturbances in the ratios of elements in plants were found in sites located near the Polish and Czech border. In plants and soil in these sites an elevated level of heavy metals was found which testifies the transboundary transport of air pollutants. The bilberry seems to be a better bioindicator of heavy metal pollution than silver fir.

  14. Variability of silver fir (Abies alba Mill. progeny from the Tisovik Reserve expressed in needle traits and chloroplast microsatellite DNA

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    Pawlaczyk Ewa M.


    Full Text Available Progeny from nineteen family lines of silver fir (Abies alba Mill. from the Tisovik Reserve growing in an experimental plot were analyzed based on 4 chloroplast microsatellite DNA loci and 12 morphological and anatomical needle traits. The Tisovik Reserve is located in Białowieża Primeval Forest, 120 km north of the natural range limit of this species, and embraces a small and isolated natural population of silver fir. The aim of this study was to determine genetic variation within and between progeny lines. Analysis of phenotypic variation showed that the traits which differed most among individuals were the needle width and the distance from resin canals to vascular bundle. Those traits, which differed most between the progeny lines, were the number of endodermic cells around the vascular bund and the weight of hypodermic cells. In Tisovik progeny, we detected 107 different haplotypes. In progeny lines, we detected more haplotypes than in maternal trees, and most haplotypes did not exist in maternal trees. This may be the result of pollen influx from other silver fir stands. Progeny from Tisovik showed a higher level of variability in comparison with maternal trees.

  15. ClRTL1 Encodes a Chinese Fir RNase III–Like Protein Involved in Regulating Shoot Branching

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    Xia Li


    Full Text Available Identification of genes controlling shoot branching is crucial for improving plant architecture and increasing crop yield or biomass. A branching mutant of Chinese fir named “Dugansha” (Cunninghamia lanceolata var. dugan. has been isolated in our laboratory. We chose the cDNA-AFLP technique and an effective strategy to screen genes that potentially regulate shoot branching in Chinese fir using this mutant. An RNase III-like1 cDNA fragment named ClRTL1 was identified as a potential positive regulator. To investigate the function of ClRTL1 in regulating shoot branching, we cloned the full-length cDNA sequence from C. lanceolata (Lamb. Hook, deduced its secondary structure and function, and overexpressed the coding sequence in Arabidopsis. The ClRTL1 cDNA is 1045 bp and comprises an open reading frame of 705 bp. It encodes a protein of 235 amino acids. The deduced secondary structure of the ClRTL1 indicates that it is a mini-RNase III-like protein. The expression analysis and phenotypes of 35S: ClRTL1 in A. thaliana implies that ClRTL1 plays a role in promoting shoot branching in Chinese fir.

  16. Southward Pleistocene migration of Douglas-fir into Mexico: phylogeography, ecological niche modeling, and conservation of 'rear edge' populations. (United States)

    Gugger, Paul F; González-Rodríguez, Antonio; Rodríguez-Correa, Hernando; Sugita, Shinya; Cavender-Bares, Jeannine


    • Poleward Pleistocene plant migration has been an important process structuring modern temperate and boreal plant communities, but the contribution of equatorward migration remains poorly understood. Paleobotanical evidence suggests Miocene or Pleistocene origin for temperate 'sky island' plant taxa in Mexico. These 'rear edge' populations situated in a biodiversity hotspot may be an important reserve of genetic diversity in changing climates. • We used mtDNA sequences, cpDNA sequences and chloroplast microsatellites to test hypotheses of Miocene vs Pleistocene colonization of temperate Douglas-fir in Mexico, explore geographic patterns of molecular variation in relation to Pleistocene climate history using ecological niche models, and assess the taxonomic and conservation implications. • We found strong evidence for Pleistocene divergence of Douglas-fir in Mexico (958 thousand yr before present (ka) with the 90% highest posterior density interval ranging from 1.6 million yr before present (Ma) to 491 ka), consistent with the southward Pleistocene migration hypothesis. Genetic diversity was high and strongly partitioned among populations. Spatial patterns of molecular variation and ecological niche models suggest a complex late Pleistocene history involving periods of isolation and expansion along mountain corridors. • These results highlight the importance of southward Pleistocene migration in establishing modern high-diversity plant communities and provide critical insights into proposals to conserve the unique biodiversity of Mexican Douglas-fir and associated taxa. © 2010 The Authors. New Phytologist © 2010 New Phytologist Trust.

  17. Optimized FPGA Implementation of Multi-Rate FIR Filters Through Thread Decomposition (United States)

    Kobayashi, Kayla N.; He, Yutao; Zheng, Jason X.


    Multi-rate finite impulse response (MRFIR) filters are among the essential signal-processing components in spaceborne instruments where finite impulse response filters are often used to minimize nonlinear group delay and finite precision effects. Cascaded (multistage) designs of MRFIR filters are further used for large rate change ratio in order to lower the required throughput, while simultaneously achieving comparable or better performance than single-stage designs. Traditional representation and implementation of MRFIR employ polyphase decomposition of the original filter structure, whose main purpose is to compute only the needed output at the lowest possible sampling rate. In this innovation, an alternative representation and implementation technique called TD-MRFIR (Thread Decomposition MRFIR) is presented. The basic idea is to decompose MRFIR into output computational threads, in contrast to a structural decomposition of the original filter as done in the polyphase decomposition. A naive implementation of a decimation filter consisting of a full FIR followed by a downsampling stage is very inefficient, as most of the computations performed by the FIR state are discarded through downsampling. In fact, only 1/M of the total computations are useful (M being the decimation factor). Polyphase decomposition provides an alternative view of decimation filters, where the downsampling occurs before the FIR stage, and the outputs are viewed as the sum of M sub-filters with length of N/M taps. Although this approach leads to more efficient filter designs, in general the implementation is not straightforward if the numbers of multipliers need to be minimized. In TD-MRFIR, each thread represents an instance of the finite convolution required to produce a single output of the MRFIR. The filter is thus viewed as a finite collection of concurrent threads. Each of the threads completes when a convolution result (filter output value) is computed, and activated when the first


    Energy Technology Data Exchange (ETDEWEB)

    Takahashi, Satoko [Joint ALMA Observatory, Alonso de Cordova 3107, Vitacura, Santiago (Chile); Ohashi, Nagayoshi [Academia Sinica Institute of Astronomy and Astrophysics, P.O. Box 23-141, Taipei 10617, Taiwan (China); Bourke, Tyler L., E-mail: [Harvard-Smithsonian Center for Astrophysics, 60 Garden Street, Cambridge, MA 02138 (United States)


    Studying the physical conditions of very low luminosity objects (VeLLOs; L{sub bol} < 0.1 L{sub Sun }) is important for understanding the earliest evolutionary stage of protostars and brown dwarfs. We report interferometric observations of the VeLLO L1521F-IRS, in {sup 12}CO (2-1) line emission and the 1.3 mm continuum emission, using the Submillimeter Array. With the {sup 12}CO (2-1) high-resolution observations, we have spatially resolved a compact but poorly collimated molecular outflow associated with L1521F-IRS for the first time. The blueshifted and redshifted lobes are aligned along the east and west side of L1521F-IRS with a lobe size of Almost-Equal-To 1000 AU. The estimated outflow mass, maximum outflow velocity, and outflow force are (9.0-80) Multiplication-Sign 10{sup -4} M{sub Sun }, 7.2 km s{sup -1}, and (7.4-66) Multiplication-Sign 10{sup -7} M{sub Sun} km s{sup -1} yr{sup -1}, respectively. The estimated outflow parameters such as size, mass, and momentum rate are similar to values derived for other VeLLOs, and are located at the lower end of values compared to previously studied outflows associated with low- to high-mass star-forming regions. Low-velocity less collimated (1.5 km s{sup -1}/1200 AU) and higher-velocity compact (4.0 km s{sup -1}/920 AU) outflow components are suggested by the data. These velocity structures are not consistent with those expected in the jet-driven or wind-driven outflow models, perhaps suggesting a remnant outflow from the first hydrostatic core as well as an undeveloped outflow from the protostar. Detection of an infrared source and compact millimeter continuum emission suggests the presence of the protostar, while its low bolometric luminosity (0.034-0.07 L{sub Sun }) and small outflow suggests that L1521F is in the earliest protostellar stage (<10{sup 4} yr) and contains a substellar mass object. The bolometric (or internal) luminosity of L1521F-IRS suggests that the current mass accretion rate is an order of

  19. Process coupling and control over the response of net ecosystem CO2 exchange to climate variability and insect disturbance in subalpine forests of the Western US (United States)

    Monson, R. K.; Moore, D. J.; Trahan, N. A.; Scott-Denton, L.; Burns, S. P.; Hu, J.; Bowling, D. R.


    Following ten years of studies in subalpine forest ecosystems of the Western US, we have concluded that the tight coupling between gross primary productivity (GPP) and the autotrophic component of soil respiration (Ra) drives responses of net ecosystem CO2 exchange (NEE) to climate variability and insect disturbance. This insight has been gained through long-term eddy flux observations, manipulative plot experiments, analyses of dynamics in the stable isotope compositions of CO2 and H2O, and chamber gas-exchange measurements. Using past observations from these studies, we deployed model-data assimilation techniques and forecast weather/climate modeling to estimate how the coupling between GPP and Ra is likely to affect future (Year 2100) dynamics in NEE. The amount of winter snow and its melting dynamics in the spring represents the dominant control over interannual variation in GPP. Using the SIPNET ecosystem process model, combined with knowledge about the stable isotope content of different water sources, we estimated that approximately 75% of growing season GPP is coupled to the use of snowmelt water, whereas approximately 25% is coupled to summer rain. The tight coupling between GPP and winter snow pack drives a similar tight coupling between soil respiration (Rs) and winter snow pack. Manipulation of snow pack on forest plots has shown that Rs increases with increased snow pack, and this effect disappears when trees are girdled, which stops the transfer of GPP to roots and the soil rhizosphere. Higher-than-normal winter snowpacks cause the carbon isotope ratios of soil-respired CO2 to be depleted in 13C, reflecting a signal of lower photosynthetic water-use efficiency in the GPP that is transferred to the soil rhizosphere. Large-scale forest disturbance due to catastrophic tree mortality from mountain pine beetle attack causes an initial (2-3 year) reduction in Rs, which is attributable to the loss of GPP and its effect on Ra. This near-term reduction in Rs

  20. From static to dynamic mapping of chromospheric magnetism - FIRS and SPIES (United States)

    Schad, Thomas A.; Lin, Haosheng


    Advancements in theoretical forward modeling and observational techniques now allow the mapping of the chromospheric magnetic field vector in some regions. We report on full maps of the chromospheric magnetic field vector across a sunspot and its superpenumbra within NOAA AR 11408. These maps are derived from full Stokes observations of the He I triplet at 1083 nm, which show both Zeeman and atomic-level polarization signatures. Yet, due to the long time to acquire these observations with the slit-based Facility Infrared Spectropolarimeter (FIRS), our measurements primarily probe long-lived chromospheric structures, albeit at very high polarization sensitivity. The fast temporal scales remain difficult to probe with conventional slit-based spectropolarimeters. Alternatively, SPIES is an instrument based on a birefringent fiber optic IFU, designed to multiplex a two-dimensional spatial field with high spectral resolution spectropolarimetry, and is an ideal tool for probing small-scale, dynamic magnetic features. We will present movies of the dynamic chromosphere acquired from SPIES across a sunspot and its fine-scaled superpenumbra.

  1. A Hierarchical Bayesian Model to Predict Self-Thinning Line for Chinese Fir in Southern China.

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    Xiongqing Zhang

    Full Text Available Self-thinning is a dynamic equilibrium between forest growth and mortality at full site occupancy. Parameters of the self-thinning lines are often confounded by differences across various stand and site conditions. For overcoming the problem of hierarchical and repeated measures, we used hierarchical Bayesian method to estimate the self-thinning line. The results showed that the self-thinning line for Chinese fir (Cunninghamia lanceolata (Lamb.Hook. plantations was not sensitive to the initial planting density. The uncertainty of model predictions was mostly due to within-subject variability. The simulation precision of hierarchical Bayesian method was better than that of stochastic frontier function (SFF. Hierarchical Bayesian method provided a reasonable explanation of the impact of other variables (site quality, soil type, aspect, etc. on self-thinning line, which gave us the posterior distribution of parameters of self-thinning line. The research of self-thinning relationship could be benefit from the use of hierarchical Bayesian method.

  2. The vulnerability of silver fir populations to damage from late frosts

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    Klisz Marcin


    Full Text Available The aim of the study was to determine the vulnerability of selected silver fir populations to damage from late frost in the climatic conditions of south-eastern Poland. To determine the vulnerability of apical and lateral shoots to damage caused by late frosts, we observed four test plots in 2009 and 2014, each containing progenies of selected seed stands. Our statistical analyses were based on a model incorporating the following variables: site, year, type of frost damage, population as well as the possible interaction between these variables. Significant differences between the populations were found in terms of their sensitivity to damage from low temperature occurring during the growth period. Furthermore, we indirectly demonstrated differences in the severity of late frost on the experimental plots, as well as the intensity and variability of late frost shoot damage. Based on these results, we divided the studied populations into two groups of low (EF, KRA1 and NAR and high (LES2 and BAL2 sensitivity to late frost damage.

  3. Induced compression wood formation in Douglas fir (Pseudotsuga menziesii) in microgravity (United States)

    Kwon, M.; Bedgar, D. L.; Piastuch, W.; Davin, L. B.; Lewis, N. G.


    In the microgravity environment of the Space Shuttle Columbia (Life and Microgravity Mission STS-78), were grown 1-year-old Douglas fir and loblolly pine plants in a NASA plant growth facility. Several plants were harnessed (at 45 degrees ) to establish if compression wood biosynthesis, involving altered cellulose and lignin deposition and cell wall structure would occur under those conditions of induced mechanical stress. Selected plants were harnessed at day 2 in orbit, with stem sections of specific plants harvested and fixed for subsequent microscopic analyses on days 8, 10 and 15. At the end of the total space mission period (17 days), the remaining healthy harnessed plants and their vertical (upright) controls were harvested and fixed on earth. All harnessed (at 45 degrees ) plant specimens, whether grown at 1 g or in microgravity, formed compression wood. Moreover, not only the cambial cells but also the developing tracheid cells underwent significant morphological changes. This indicated that the developing tracheids from the primary cell wall expansion stage to the fully lignified maturation stage are involved in the perception and transduction of the stimuli stipulating the need for alteration of cell wall architecture. It is thus apparent that, even in a microgravity environment, woody plants can make appropriate corrections to compensate for stress gradients introduced by mechanical bending, thereby enabling compression wood to be formed. The evolutionary implications of these findings are discussed in terms of "variability" in cell wall biosynthesis.

  4. Low-Cost Design of an FIR Filter by Using a Coefficient Mapping Method

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    Ming-Chih Chen


    Full Text Available This work presents a novel coefficient mapping method to reduce the area cost of the finite impulse response (FIR filter design, especially for optimizing its coefficients. Being capable of reducing the area cost and improving the filter performance, the proposed mapping method consists of four steps: quantization of coefficients, import of parameters, constitution of prime coefficients with parameters, and constitution of residual coefficients with prime coefficients. Effectiveness of the proposed coefficient mapping method is verified by selecting the 48-tap filter of IS-95 code division multiple access (CDMA standard as the benchmark. Experimental results indicate that the proposed design with canonical signed digit (CSD coefficients can operate at 86 MHz with an area of 241,813 um2, leading to a throughput rate of 1,382 Mbps. Its ratio of throughput/area is 5,715 Kbps/um2, yielding a higher performance than that of previous designs. In summary, the proposed design reduces 5.7% of the total filter area, shortens 25.7% of the critical path delay, and improves 14.8% of the throughput/area by a value over that of the best design reported before.

  5. Longitudinal Mechano-Sorptive Creep Behavior of Chinese Fir in Tension during Moisture Adsorption Processes. (United States)

    Peng, Hui; Lu, Jianxiong; Jiang, Jiali; Cao, Jinzhen


    To provide comprehensive data on creep behaviors at relative humidity (RH) isohume conditions and find the basic characteristics of mechano-sorptive (MS) creep (MSC), the tensile creep behaviors, "viscoelastic creep (VEC)" at equilibrium moisture content and MSC during adsorption process, were performed on Chinese fir in the longitudinal direction under 20%, 40%, 60% and 80% RH (25 °C) and at 1, 1.3, and 1.6 MPa, respectively. The free swelling behavior was also measured, where the climate conditions corresponded with MSC tests. Based on the databases of free swelling, VEC, and MSC, the existence of MS effect was examined, and the application of the rheological model under the assumption of partitioned strain was investigated. The results revealed that both VEC and MSC increased with magnitude of applied stress, and the increasing RH level. Under all RH isohume conditions, the total strain of MSC was greater than that of VEC. The influence of RH level on VEC was attributed to the water plasticization effect, whereas that on MSC was presumed to be the effect of water plasticization and unstable state in the wood cell wall. In addition, the RH level promoted the relaxation behavior in MSC, while it slightly affected the relaxation behavior in VEC. In the future, the rheological model could consider the link between load configuration and the anatomic structural feature of wood.

  6. Implementación de filtro Fir en sistemas programables en un Chip

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    Idileisy Torres Rodríguez


    Full Text Available Normal 0 21 false false false MicrosoftInternetExplorer4 /* Style Definitions */ table.MsoNormalTable {mso-style-name:"Tabla normal"; mso-tstyle-rowband-size:0; mso-tstyle-colband-size:0; mso-style-noshow:yes; mso-style-parent:""; mso-padding-alt:0cm 5.4pt 0cm 5.4pt; mso-para-margin:0cm; mso-para-margin-bottom:.0001pt; mso-pagination:widow-orphan; font-size:10.0pt; font-family:"Times New Roman"; mso-ansi-language:#0400; mso-fareast-language:#0400; mso-bidi-language:#0400;} Los filtros (analógicos y digitales se usan para la separación de señales combinadas y la restauración de señales que presentan alguna distorsión. Los filtros digitales alcanzan mejores resultados, con precisión limitada por errores de redondeo en la aritmética empleada. El diseño de un filtro digital con respuesta finita al impulso (FIR y la cuantización de sus coeficientes para facilitar su implementación en PSoC es el tema de este trabajo.

  7. Assessing Douglas-Fir Seedling Establishment Using Two Modified Forestry Reclamation Approaches in the Pacific Northwest

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    Colton Miller


    Full Text Available The Forestry Reclamation Approach uses uncompacted, mounded spoils to reforest mined-land and has been successful in hardwood forests in the Appalachian region. A surface coalmine reclamation site in the Pacific Northwest was used to compare the site’s standard reclamation approach (Reference with a modified version of the Forestry Reclamation Approach (FRA along with a modified FRA treatment that also incorporated an amendment of bottom ash from the coal burning power plant on-site (FRA + Ash. Survival and growth were followed for three growing seasons in bareroot and container Douglas-fir seedlings. Soil characteristics and understory cover were also assessed. Considerable variation in microsite characteristics was observed in the study area. Container seedlings did not improve survival compared to bareroot seedlings. In the soil reclamation treatments, seedling survival was significantly higher in FRA + Ash treatments compared to FRA and Reference treatments at the end of the second growing season. Survival declined in each year of the study, but the order of treatment effectiveness did not change. Relativized growth increment was significantly higher in the FRA treatment compared to both the Reference and FRA + Ash treatments during the third growing season. Understory cover was established after three years, but varied substantially across the study area.

  8. Climate-Induced Mortality of Siberian Pine and Fir in the Lake Baikal Watershed, Siberia (United States)

    Kharuk, Viacheslav I.; Im, Sergei T.; Petrova, IIya A.; Golyukov, Alexei S.; Ranson, Kenneth J.; Yagunov, Mikhail N.


    Siberian pine (Pinus sibirica) and fir (Abies sibirica) (so called "dark needle conifers", DNC) showed decreased radial growth increment within the Lake Baikal watershed since the 1980s with increasing mortality recorded since the year 2000. Tree ring width was strongly correlated with vapor pressure deficit, aridity and root zone moisture. Water stress from droughts made trees more susceptible to insect attacks causing mortality in about 10% of DNC stands within the Lake Baikal watershed. Within Siberia DNC mortality increased in the southern part of the DNC range. Biogeographically, tree mortality was located within the DNC - forest-steppes transition. Tree mortality was significantly correlated with drought and soil moisture anomalies. Within the interior of the DNC range mortality occurred within relief features with high water stress risk (i.e., steep convex south facing slopes with shallow well-drained soils). In general, DNC mortality in Siberia was induced by increased aridity and severe drought (inciting factors) in synergy with biotic attacks (contributing factor). In future climate scenarios with predicted increase in aridity DNC could be eliminated from the southern part of its current range and will be replaced by drought-resistant conifers and broadleaf species (e.g., Larix sibirica, Pinus silvestris, and Betula pubescence).

  9. Microwave and FIR Spectroscopy of Dimethylsulfide in the Ground, First and Second Excited Torsional States (United States)

    Ilyushin, V.; Armieieva, Iuliia; Dorovskaya, Olga; Pogrebnyak, Mykola; Krapivin, Igor; Alekseev, E. A.; Margulès, L.; Motiyenko, R. A.; Kwabia Tchana, F.; Jabri, Atef; Manceron, Laurent; Bauerecker, Sigurd; Maul, Christof


    A new study of the dimethylsulfide ((CH_{3})_{2}S) spectrum is reported. The new measurements have been carried out using the Kharkiv spectrometer in the Institute of Radio Astronomy of NASU (Ukraine) and using the Lille spectrometer in the PhLAM laboratory (France). The new millimeter and submillimeter wave measurements cover the frequency range from 49 GHz to 660 GHz. The rotational transitions belonging to the three lowest torsional states of the molecule as well as the new assignments in the FIR torsional band (AILES beamline of the synchrotron SOLEIL) and the microwave data available in the literature have been analyzed using recently developed model for the molecules with two equivalent methyl rotors and C_{2v} symmetry at equilibrium (PAM_C2v_2tops program). In the talk the details of this new study will be discussed. This work was done under support of the Volkswagen foundation. The assistance of Science and Technology Center in Ukraine is acknowledged (STCU partner project P686). A. Jabri, V. Van, H. V. L. Nguyen, H. Mouhib, F. Kwabia Tchana , L. Manceron , W. Stahl, I. Kleiner, A&A 589, A127 (2016). Ilyushin V. V., Hougen J. T. J. Mol. Spectrosc. 289 (2013) 41-49.

  10. Modification of fast-growing Chinese Fir wood with unsaturated polyester resin: Impregnation technology and efficiency (United States)

    Ma, Qing; Zhao, Zijian; Yi, Songlin; Wang, Tianlong

    In this study, Chinese Fir was impregnated with unsaturated polyester resin to enhance its properties. Samples 20 mm × 20 mm × 20 mm in size were split into different sections with epoxy resin and tinfoil and subjected to an impregnation experiment under various parameters. Vacuum degree was -0.04 MPa, -0.06 MPa or -0.08 MPa and vacuum duration was 15 min, 30 min, or 45 min. The results indicated that impregnation weight percent gain is linearly dependent on curing weight percent gain. Vacuum duration appears to have less influence on the curing weight percent gain than vacuum degree, and impregnation was most successful at the transverse section compared to other sections. The optimal impregnation parameters were 30 min modification under -0.08 MPa vacuum followed by 120 min at atmospheric pressure for samples 200 mm × 100 mm × 20 mm in size. Uneven distribution of weight percent gain and cracking during the curing process suggested that 30 min post-processing at -0.09 MPa vacuum was the most effective way to complete the impregnation process. The sample's bending strength and modulus of elasticity increased after impregnation treatment. Bending strength after impregnation without post-processing reached 112.85%, but reached 71.65% with vacuum-processing; modulus of elasticity improved 67.13% and 58.28% without and with post-processing, respectively.

  11. Epicuticular wax on stomata of damaged silver fir trees (Abies alba Mili.

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    Tomislav Bačić


    Full Text Available Condition of epistomatal wax on the abaxial surface of the current and previous-year needles of damaged silver fir trees (Abies alba Mill., both from the polluted Risnjak and "clean" Donja Dobra sites in Gorski Kotar region, both influenced by pollutants coming from Europe, during two years, three times a year, were examined with Scanning Electron Microscope. In the course of time the wax tubules on the epistomatal rims of stomata in polluted, but also in "clean" needles surface, become fused and agglomerated rapidly to various extents of morphologically different types of amorphous wax crusts, primarily compact and particulate ones. This process begins very early, especially in polluted Risnjak site, and may be interpreted as a possible result of air pollution. However, the recrystalization, or production of new tubules, also appears relatively quickly in mostly cases. Quantitative estimations indicate a very large total amount of amorphous wax crusts in the current-year needles, and a very high percentage of the same wax in previous-year needles. Amorphous wax crusts cover stomatal pores, as well as the rims, disturbing the normal gas exchange. Statistically there is a signicant tendency of increase in wax degradation in the needles of the polluted site in comparison with those of the unpolluted one, but there is an insignificant wax degradation among the needles of damaged trees within each site. These results confirmed most of the research done in our preliminary report.

  12. Design of hardware efficient FIR filter: A review of the state-of-the-art approaches

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    Abhijit Chandra


    Full Text Available Digital signal processing (DSP is one of the most powerful technologies which will shape the science, engineering and technology of the twenty-first century. Since 1970, revolutionary changes took place in the broad area of DSP which has made it an essential tool in many engineering applications. Digital filter is considered to be one of the most important components of almost every DSP sub-systems and therefore a number of extensive works had been carried out by researchers on the design of such filter. In order to meet the stringent requirements of filter specification, order of the designed filter is generally assumed to be very large and this leads to high power and area consumption during their implementation. As a matter of fact, design of hardware efficient digital filter has drawn enormous attention which needs to be addressed by various useful means. One popular approach has been to encode the tap coefficients of such filter in the form of sum of signed powers-of-two and thus the operation of multiplication is substituted by simple addition and shifting. This paper presents a detailed review of the basic design approaches applicable for the synthesis of hardware efficient finite duration impulse response (FIR filter. Both the traditional and heuristic search algorithms have been incorporated and properly arranged in this review.

  13. Regulation of Water Use in the Southernmost European Fir (Abies pinsapo Boiss.: Drought Avoidance Matters

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    Raúl Sánchez-Salguero


    Full Text Available The current scenario of global warming has resulted in considerable uncertainty regarding the capacity of forest trees to adapt to increasing drought. Detailed ecophysiological knowledge would provide a basis to forecast expected species dynamics in response to climate change. Here, we compare the water balance (stomatal conductance, xylem water potential, needle osmotic adjustment of Abies pinsapo, a relict drought-sensitive Mediterranean fir, along an altitudinal gradient. We related these variables to soil water and nutrient availability, air temperature, atmospheric water potential, and vapour pressure deficit during two consecutive years. Our results indicate that A. pinsapo closed stomata rapidly over a very narrow range of soil water availability and atmospheric dryness. This isohydric response during water stress suggests that this relict conifer relied on the plant hormone abscisic acid to maintain closed stomata during sustained drought, instead of needle desiccation to passively drive stomatal closure, needle osmotic adjustment or a plastic response of the xylem to different levels of water availability. Both the soil and foliar nutrient contents suggest that the studied populations are not limited by nutrient deficiencies, and drought was stronger in the warmer low-elevation areas.

  14. Transplantation of subalpine wood-pasture turfs along a natural climatic gradient reveals lower resistance of unwooded pastures to climate change compared to wooded ones. (United States)

    Gavazov, Konstantin; Spiegelberger, Thomas; Buttler, Alexandre


    Climate change could impact strongly on cold-adapted mountain ecosystems, but little is known about its interaction with traditional land-use practices. We used an altitudinal gradient to simulate a year-round warmer and drier climate for semi-natural subalpine grasslands across a landscape of contrasting land-use management. Turf mesocosms from three pasture-woodland land-use types-unwooded pasture, sparsely wooded pasture, and densely wooded pasture-spanning a gradient from high to low management intensity were transplanted downslope to test their resistance to two intensities of climate change. We found strong overall effects of intensive (+4 K) experimental climate change (i.e., warming and reduced precipitation) on plant community structure and function, while moderate (+2 K) climate change did not substantially affect the studied land-use types, thus indicating an ecosystem response threshold to moderate climate perturbation. The individual land-use types were affected differently under the +4 K scenario, with a 60% decrease in aboveground biomass (AGB) in unwooded pasture turfs, a 40% decrease in sparsely wooded pasture turfs, and none in densely wooded ones. Similarly, unwooded pasture turfs experienced a 30% loss of species, advanced (by 30 days) phenological development, and a mid-season senescence due to drought stress, while no such effects were recorded for the other land-use types. The observed contrasting effects of climate change across the pasture-woodland landscape have important implications for future decades. The reduced impact of climate change on wooded pastures as compared to unwooded ones should promote the sustainable land use of wooded pastures by maintaining low management intensity and a sparse forest canopy, which buffer the immediate impacts of climate change on herbaceous vegetation.

  15. Exploring the landscape evolution of the subalpine meadow-forest system driven by the geomorphic work performed by the Northern Pocket Gopher (United States)

    Winchell, E. W.; Anderson, R. S.; Lombardi, E. M.; Doak, D. F.


    In the subalpine zone of the Colorado Front Range, field observations suggest that the Northern Pocket Gopher acts as a significant geomorphic agent within meadows, but not within forests. Field surveys during 2014 and 2015 demonstrate that the temporal and spatial digging patterns of gopher-excavated mounds and infilled tunnels are neither steady nor uniform. These include 1) gophers spend the winter near the forest-meadow (FM) edge and the remainder of the year within the meadow, and 2) surface mound generation greatly accelerates in late summer. Hourly subsurface temperatures across the FM pair, and daily digital snow depths at the FM boundary suggest that gophers spend the winter beneath thick snow cover where ground temperatures are warmest. LiDAR-based topography demonstrates that slopes are uniform across the FM pairs, diverging from that expected by extrapolation of the observed pattern of non-uniform geomorphic activity. The topography therefore suggests that the FM boundaries are not stationary. We hypothesize that the landscape is more uniformly impacted by gopher activity in the long term, which requires that 1) FM boundaries migrate significantly and/or 2) meadows are born in different places following forest death via fire followed by rapid gopher habitation. The vertical geomorphic signature of gopher activity is more distinct. Preliminary probing of meadows reveals a 20 cm thick biomantle with a high concentration of stones at ~20 cm depth. The annual surface areas of mounds and infilled tunnels suggest that the entire meadow can be exposed to excavated tillings on century timescales. Further, annual mound volumes suggest that the biomantle is turned over also on century timescales. We will report results of stone line surveys and 137Cs concentration profiles within the forest and meadow that will test the long-term stability of meadows and the timescale over which vertical churning mixes the near-surface material within this landscape.

  16. Use of cosmogenic 35S for comparing ages of water from three alpine-subalpine basins in the Colorado Front Range (United States)

    Sueker, J.K.; Turk, J.T.; Michel, R.L.


    High-elevation basins in Colorado are a major source of water for the central and western United States; however, acidic deposition may affect the quality of this water. Water that is retained in a basin for a longer period of time may be less impacted by acidic deposition. Sulfur-35 (35S), a short-lived isotope of sulfur (t( 1/2 ) = 87 days), is useful for studying short-time scale hydrologic processes in basins where biological influences and water/rock interactions are minimal. When sulfate response in a basin is conservative, the age of water may be assumed to be that of the dissolved sulfate in it. Three alpine-subalpine basins on granitic terrain in Colorado were investigated to determine the influence of basin morphology on the residence time of water in the basins. Fern and Spruce Creek basins are glaciated and accumulate deep snowpacks during the winter. These basins have hydrologic and chemical characteristics typical of systems with rapid hydrologic response times. The age of sulfate leaving these basins, determined from the activity of 35S, averages around 200 days. In contrast, Boulder Brook basin has broad, gentle slopes and an extensive cover of surficial debris. Its area above treeline, about one-half of the basin, is blown free of snow during the winter. Variations in flow and solute concentrations in Boulder Brook are quite small compared to Fern and Spruce Creeks. After peak snowmelt, sulfate in Boulder Brook is about 200 days older than sulfate in Fern and Spruce Creeks. This indicates a substantial source of older sulfate (lacking 35S) that is probably provided from water stored in pore spaces of surficial debris in Boulder Brook basin.

  17. Effects of a long-term disturbance on arthropods and vegetation in subalpine wetlands: manifestations of pack stock grazing in early versus mid-season.

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    Jeffrey G Holmquist

    Full Text Available Conclusions regarding disturbance effects in high elevation or high latitude ecosystems based solely on infrequent, long-term sampling may be misleading, because the long winters may erase severe, short-term impacts at the height of the abbreviated growing season. We separated a long-term effects of pack stock grazing, manifested in early season prior to stock arrival, from b additional pack stock grazing effects that might become apparent during annual stock grazing, by use of paired grazed and control wet meadows that we sampled at the beginning and end of subalpine growing seasons. Control meadows had been closed to grazing for at least two decades, and meadow pairs were distributed across Sequoia National Park, California, USA. The study was thus effectively a landscape-scale, long-term manipulation of wetland grazing. We sampled arthropods at these remote sites and collected data on associated vegetation structure. Litter cover and depth, percent bare ground, and soil strength had negative responses to grazing. In contrast, fauna showed little response to grazing, and there were overall negative effects for only three arthropod families. Mid-season and long-term results were generally congruent, and the only indications of lower faunal diversity on mid-season grazed wetlands were trends of lower abundance across morphospecies and lower diversity for canopy fauna across assemblage metrics. Treatment x Season interactions almost absent. Thus impacts on vegetation structure only minimally cascaded into the arthropod assemblage and were not greatly intensified during the annual growing season. Differences between years, which were likely a response to divergent snowfall patterns, were more important than differences between early and mid-season. Reliance on either vegetation or faunal metrics exclusively would have yielded different conclusions; using both flora and fauna served to provide a more integrative view of ecosystem response.

  18. Effects of a long-term disturbance on arthropods and vegetation in subalpine wetlands: manifestations of pack stock grazing in early versus mid-season. (United States)

    Holmquist, Jeffrey G; Schmidt-Gengenbach, Jutta; Haultain, Sylvia A


    Conclusions regarding disturbance effects in high elevation or high latitude ecosystems based solely on infrequent, long-term sampling may be misleading, because the long winters may erase severe, short-term impacts at the height of the abbreviated growing season. We separated a) long-term effects of pack stock grazing, manifested in early season prior to stock arrival, from b) additional pack stock grazing effects that might become apparent during annual stock grazing, by use of paired grazed and control wet meadows that we sampled at the beginning and end of subalpine growing seasons. Control meadows had been closed to grazing for at least two decades, and meadow pairs were distributed across Sequoia National Park, California, USA. The study was thus effectively a landscape-scale, long-term manipulation of wetland grazing. We sampled arthropods at these remote sites and collected data on associated vegetation structure. Litter cover and depth, percent bare ground, and soil strength had negative responses to grazing. In contrast, fauna showed little response to grazing, and there were overall negative effects for only three arthropod families. Mid-season and long-term results were generally congruent, and the only indications of lower faunal diversity on mid-season grazed wetlands were trends of lower abundance across morphospecies and lower diversity for canopy fauna across assemblage metrics. Treatment x Season interactions almost absent. Thus impacts on vegetation structure only minimally cascaded into the arthropod assemblage and were not greatly intensified during the annual growing season. Differences between years, which were likely a response to divergent snowfall patterns, were more important than differences between early and mid-season. Reliance on either vegetation or faunal metrics exclusively would have yielded different conclusions; using both flora and fauna served to provide a more integrative view of ecosystem response.

  19. Lake biota response to human impact and local climate during the last 200 years: A multi-proxy study of a subalpine lake (Tatra Mountains, W Carpathians). (United States)

    Hamerlík, Ladislav; Dobríková, Daniela; Szarlowicz, Katarzyna; Reczynski, Witold; Kubica, Barbara; Šporka, Ferdinand; Bitušík, Peter


    Element content, loss-on-ignition, chironomid analysis and (210)Pb dating were applied on a sediment core from a subalpine Tatra lake (Popradské pleso) to reveal the response of aquatic biota to eutrophication induced by human activities in the lake catchment. The lead dating indicates that the 0-8 cm section of the core represents the past ca 200 years, ending at ~1814 AD. Comparing the key changes of the proxies with human activities that are historically well documented, four phases of the recent lake development were distinguished: (1) a pre-tourism phase, (2) a phase of increasing touristic activity and early cottage development, (3) a phase of eutrophication, and (4) a phase of post-eutrophication. Neither touristic activity, nor early cottage development around the lake (1st and 2nd phases) had considerable influence on the chironomid assemblage structure or organic content of the lake. The most significant change both in chironomid assemblage structure and loss-on-ignition occurred during the 3rd phase, when a big tourist hotel was built close by the lake and started contaminating it via direct wastewater input. However, the structure of the chironomid assemblage has not changed significantly over time and the dominating taxa remained the same during the whole period. Parallel with the nutrient signal of the paleo assemblage, a secondary signal has been identified as the ratio of rheophilic taxa on total abundance that did not correlate with the sediment's organic content, and is most likely driven by local climatic oscillations. Changes of most of metal elements concentrations reflected rather bigger scale changes of industrial activities than local scale human disturbances. Our results indicate that hydromorphological properties can moderate the impact of organic pollution on the lake biota. Copyright © 2015 Elsevier B.V. All rights reserved.

  20. [A comparative study on soil fauna in native secondary evergreen broad-leaved forest and Chinese fir plantation forests in subtropics]. (United States)

    Yan, Shaokui; Wang, Silong; Hu, Yalin; Gao, Hong; Zhang, Xiuyong


    In this study, we investigated the response of soil animal communities to the replacement of native secondary forest by Chinese fir plantation forest and successive rotation of Chinese fir in subtropics. Three adjacent forest stands, i.e., native secondary evergreen broad-leaved forest stand (control) and Chinese fir plantation stands of first (20 yr) and second (20 yr) rotations were selected for the comparison of soil fauna. All animals were extracted from the floor litter and 0-15 cm soil layer of the stands in Summer, 2003 by using Tullgren method, wet funnel method and hand-sorting method. Compared to two Chinese fir plantation forests, the native secondary evergreen broad-leaved forest had a higher abundance and a higher taxonomic diversity of animals in soil and litter, but there were no significant differences in the biomass and productivity of soil fauna between all study stands. The abundance or diversity did not differ significantly between the first rotation and second rotation stands, too. The results supported that vegetation cover might be one of the main forces driving the development of soil animal communities, and the effect of successive rotation of Chinese fir on the development of soil fauna was a slow-running process.


    Directory of Open Access Journals (Sweden)

    Ali Ömer Üçler


    Full Text Available In this study, naturally regenerated succesfull oriental spruce (Picea orientalis (L. Link and fir (Abies nordmanniana Spach. subsp. nordmanniana mixed stand which encompass 5.8 ha area was examined. For that, seedlings in the area were studied for their relationship between growth and age. In addition, the amount of seedlings reached to area after the cuttings with large shelterwood system were determined. The age and height of fir and oriental spruce seedlings in 24 sampling areas each covering 25 square meters are measured according to square method and sampling areas systematically distributed so that samples could represent the regeneration area. Evaluating gathered data, seedlings reached to the area orderly after 1975 preperation cutting. Measurements and counts showed that orieantal spruce hold rich seeds every 4 years, however, this differentiation is not evident for fir. Evaluating height growth depending on age, correlation coefficients are found 0.9993 for oriental spruce and 0.9973 for fir, therefore the seedlings of fir and oriental spruce realized similar growth during their regeneration stage

  2. Drought effects on root and needle terpenoid content of a coastal and an interior Douglas fir provenance. (United States)

    Kleiber, Anita; Duan, Qiuxiao; Jansen, Kirstin; Verena Junker, Laura; Kammerer, Bernd; Rennenberg, Heinz; Ensminger, Ingo; Gessler, Arthur; Kreuzwieser, Jürgen


    Douglas fir (Pseudotsuga menziesii) is a conifer species that stores large amounts of terpenoids, mainly monoterpenoids in resin ducts of various tissues. The effects of drought on stored leaf terpenoid concentrations in trees are scarcely studied and published data are partially controversial, since reduced, unaffected or elevated terpenoid contents due to drought have been reported. Even less is known on the effect of drought on root terpenoids. In the present work, we investigated the effect of reduced water availability on the terpenoid content in roots and needles of Douglas fir seedlings. Two contrasting Douglas fir provenances were studied: an interior provenance (var. glauca) with assumed higher drought resistance, and a coastal provenance (var. menziesii) with assumed lower drought resistance. We tested the hypothesis that both provenances show specific patterns of stored terpenoids and that the patterns will change in response to drought in both, needles and roots. We further expected stronger changes in the less drought tolerant coastal provenance. For this purpose, we performed an experiment under controlled conditions, in which the trees were exposed to moderate and severe drought stress. According to our expectations, the study revealed clear provenance-specific terpenoid patterns in needles. However, such patterns were not detected in the roots. Drought slightly increased the needle terpenoid contents of the coastal but not of the interior provenance. We also observed increased terpenoid abundance mainly in roots of the moderately stressed coastal provenance. Overall, from the observed provenance-specific reactions with increased terpenoid levels in trees of the coastal origin in response to drought, we conclude on functions of terpenoids for abiotic stress tolerance that might be fulfilled by other, constitutively expressed mechanisms in drought-adapted interior provenances. © The Author 2017. Published by Oxford University Press. All rights

  3. Species distribution models may misdirect assisted migration: insights from the introduction of Douglas-fir to Europe. (United States)

    Boiffin, Juliette; Badeau, Vincent; Bréda, Nathalie


    Species distribution models (SDMs), which statistically relate species occurrence to climatic variables, are widely used to identify areas suitable for species growth under future climates and to plan for assisted migration. When SDMs are projected across times or spaces, it is assumed that species climatic requirements remain constant. However, empirical evidence supporting this assumption is rare, and SDM predictions could be biased. Historical human-aided movements of tree species can shed light on the reliability of SDM predictions in planning for assisted migration. We used Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco), a North American conifer introduced into Europe during the mid-19th century, as a case-study to test niche conservatism. We combined transcontinental data sets of Douglas-fir occurrence and climatic predictors to compare the realized niches between native and introduced ranges. We calibrated a SDM in the native range and compared areas predicted to be climatically suitable with observed presences. The realized niches in the native and introduced ranges showed very limited overlap. The SDM calibrated in North America had very high predictive power in the native range, but failed to predict climatic suitability in Europe where Douglas-fir grows in climates that have no analogue in the native range. We review the ecological mechanisms and silvicultural practices that can trigger such shifts in realized niches. Retrospective analysis of tree species introduction revealed that the assumption of niche conservatism is erroneous. As a result, distributions predicted by SDM are importantly biased. There is a high risk that assisted migration programs may be misdirected and target inadequate species or introduction zones. © 2016 by the Ecological Society of America.

  4. Douglas-fir forests in the Oregon and Washington Cascades: relation of the herpetofauna to stand age and moisture (United States)

    Bury, R. Bruce; Corn, P.S.


    Pitfall traps effectively sampled amphibians but not reptiles in Douglas-fir (Pseudotsuga menziesii) forests. The abundance of only one amphibian species varied across an age gradient or a moisture gradient. Salamanders and frogs that breed in ponds or streams were captured in large numbers in some stands, likely due to the presence of nearby breeding habitat rather than forest conditions. Lizards occurred mostly in dry stands and clearcuts. Time-constrained searches showed different use of downed woody debris among terrestrial salamanders. The occurrence and abundance of species in naturally regenerated forests markedly differed from clearcut stands.

  5. Major characteristics of mixed fir and beech virgin forests in the National park Biogradska Gora in Montenegro

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    Čurović Milić


    Full Text Available In order to manage forest ecosystems at a sufficiently high biodiversity level it is necessary to study the ecological, structural and production characteristics of virgin forests. The research was directed towards identifying the characteristics of mixed fir and beech forests (Abieti-Fagetum s. lat. in the area of the strict reserve of the National Park Biogradska Gora in Montenegro. Basic characteristics of these forests were researched in the process of definition of forest types. In this manner, it is for the first time that a realistic base for typological management of forests and forest ecosystems with similar ecological and structural characteristics was provided for the specific sites.

  6. Landscape-scale effects of fire severity on mixed-conifer and red fir forest structure in Yosemite National Park (United States)

    Kane, Van R.; Lutz, James A.; Roberts, Susan L.; Smith, Douglas F.; McGaughey, Robert J.; Povak, Nicholas A.; Brooks, Matthew L.


    While fire shapes the structure of forests and acts as a keystone process, the details of how fire modifies forest structure have been difficult to evaluate because of the complexity of interactions between fires and forests. We studied this relationship across 69.2 km2 of Yosemite National Park, USA, that was subject to 32 fires ⩾40 ha between 1984 and 2010. Forests types included ponderosa pine (Pinus ponderosa), white fir-sugar pine (Abies concolor/Pinus lambertiana), and red fir (Abies magnifica). We estimated and stratified burned area by fire severity using the Landsat-derived Relativized differenced Normalized Burn Ratio (RdNBR). Airborne LiDAR data, acquired in July 2010, measured the vertical and horizontal structure of canopy material and landscape patterning of canopy patches and gaps. Increasing fire severity changed structure at the scale of fire severity patches, the arrangement of canopy patches and gaps within fire severity patches, and vertically within tree clumps. Each forest type showed an individual trajectory of structural change with increasing fire severity. As a result, the relationship between estimates of fire severity such as RdNBR and actual changes appears to vary among forest types. We found three arrangements of canopy patches and gaps associated with different fire severities: canopy-gap arrangements in which gaps were enclosed in otherwise continuous canopy (typically unburned and low fire severities); patch-gap arrangements in which tree clumps and gaps alternated and neither dominated (typically moderate fire severity); and open-patch arrangements in which trees were scattered across open areas (typically high fire severity). Compared to stands outside fire perimeters, increasing fire severity generally resulted first in loss of canopy cover in lower height strata and increased number and size of gaps, then in loss of canopy cover in higher height strata, and eventually the transition to open areas with few or no trees. However

  7. [Impact of heavy snow storm and freezing rain disasters on soil fauna in Chinese fir plantation in southern China]. (United States)

    Yan, Shao-kui; Zhang, Wei-dong; Liu, Yan-xin; Fu, Sheng-lei; Li, Yuan-liang; Wang, Si-long


    In January 2008, southern China suffered an unusual heavy snowstorm and freezing rain over a large area for almost a month long. This catastrophic event was the worst one in past 50 years, which brought the area a serious impact on the infrastructure, ecology, and environment. To understand the long-term impact of this catastrophic event on the forest ecosystems in this area, a field investigation was conducted on the soil fauna in a pure Chinese fir plantation and a mixed Chinese fir plantation-alder plantation in Huitong County of Hunan Province on March 23, 2008, the date 40 days after the heavy snowstorm and freezing rain. With the abundance and community composition as the main parameters and the monitoring data from the two plantations on March 23, 2007 as the reference, the flexibility and resistance of soil fauna to the disturbances of the catastrophic event was preliminarily evaluated. The results showed that there was a significant deviation of soil fauna communities in the two plantations from the reference. An outbreak increase in microfauna nematode abundance was found from 12216.9 ind x m(-2) to 118343.9 ind x m(-2) in pure Chinese fir plantation and from 25435.9 ind x m(-2) to 84573.0 ind x m(-2) in mixed Chinese fir plantation-alder plantation, while a 27.0% and 85.6% decrease of macrofauna abundance was found in the two plantations, respectively, compared with the reference. Mesofauna abundance also had a significant decrease in litter layer but not in soil. The abundance recovery displayed a trend from quick rate for microfauna to slow rate for macrofauna, which indicated that the soil fauna functional groups, in terms of body size, could be used as a vulnerable indicator in evaluating disturbance event and post-disturbance recovery. By using community ordinations, no shift in soil fauna community composition was detected 40 days after the catastrophic event, suggesting that the community composition of soil invertebrate had a high resistance to

  8. The Impact of Ecological and Phytocenotic Conditions on Fraction Composition of Proteins and Sowing Qualities of the Siberian Fir Seeds

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    S. G. Prokushkin


    Full Text Available The content and fractional composition of nitrogenous compounds and their role in sowing qualities of the Siberian fir has been identified. In particular: there were significant differences in the content of total protein nitrogen and individual fractions of proteins in seeds of site depending on the fir, the rate of growth of the trees, the location of the seeds in the generative layers of the crown and the vitality of the tree. It was found that the viable seeds of the Siberian fir in middle taiga contains the maximum amount of protein nitrogen, the main fraction of which are sparingly soluble and albumin. Whereas, in the seeds from the southern taiga protein nitrogen content is much less prevalent and sparingly soluble fraction and globulins. A positive correlation between protein content in seeds and their nitrogen laboratory germination component with r = 0.88 and the approximation validity with R2 = 0.77, which is typical for the trees of the Ist, and IVth class of growth by Kraft. Positive relationship has been established between laboratory germination and water-soluble fractions which, according to the researchers, serves as a catalyst. Also a relationship between seed germination and the content of salt-soluble fractions and sparingly soluble fractions, which amounted to 0.93 and 0.76, respectively, has been found. The high content of sparingly soluble protein fraction in the seeds of fir and a very high correlation with its germination is quite natural, as insoluble proteins play a role in improving the energy efficiency of respiratory cells, as well as they are a part of the cellular structures. A correlation between the amount of salt-soluble fraction and germination of seeds has been shown. This correlation is natural, since they are mainly seed storage proteins, for a number of different specific features. The relationship between the content of the alcohol and alkali-soluble protein fractions in the seeds and their germination

  9. High resolution FTIR investigation of 12C 2H 2 in the FIR spectral range using synchrotron radiation (United States)

    Amyay, B.; Herman, M.; Fayt, A.; Fusina, L.; Predoi-Cross, A.


    FIR spectra of C 2H 2 have been recorded at 0.00096 cm -1 spectral resolution using the Canadian Light Source synchrotron facility. The analysis allowed us to assign 731 new vibration-rotation lines from 48 bands in 12C 2H 2, 38 of which are reported for the first time. Two additional bands are assigned to 13CH 12CH. The measured line positions and calculated spectra can be made available to help in the remote sensing of acetylene in the terahertz spectral range.

  10. Soil-solution chemistry in a low-elevation spruce-fir ecosystem, Howland, Maine (United States)

    Fernandez, Ivan J.; Lawrence, Gregory B.; Son, Yowhan


    Soil solutions were collected monthly by tension and zero-tension lysimeters in a low-elevation red spruce stand in east-central Maine from May 1987 through December 1992. Soil solutions collected by Oa tension lysimeters had higher concentrations of most constituents than the Oa zero-tension lysimeters. In Oa horizon soil solutions growing season concentrations for SO4, Ca, and Mg averaged 57, 43, and 30 μmol L−1 in tension lysimeters, and 43, 28, and 19 μmol L−1 in zero-tension lysimeters, respectively. Because tension lysimeters remove water held by the soil at tensions up to 10 kPa, solutions are assumed to have more time to react with the soil compared to freely draining solutions collected by zero-tension lysimeters. Solutions collected in the Bs horizon by both types of collectors were similar which was attributed to the frequency of time periods when the water table was above the Bs lysimeters. Concentrations of SO4 and NO3 at this site were lower than concentrations reported for most other eastern U.S. spruce-fir sites, but base cation concentrations fell in the same range. Aluminum concentrations in this study were also lower than reported for other sites in the eastern U.S. and Ca/Al ratios did not suggest inhibition of Ca uptake by roots. Concentrations of SO4, Ca, K, and Cl decreased significantly in both the Oa and Bs horizons over the 56-month sampling period, which could reflect decreasing deposition rates for sulfur and base cations, climatic influences, or natural variation. A longer record of measured fluxes will be needed to adequately define temporal trends in solution chemistry and their causes.

  11. MIR and FIR Analysis of Inorganic Species in a Single Data Acquisition (United States)

    Wang, Peng; Shilov, Sergey


    The extension of the mid IR towards the far IR spectral range below 400 \\wn is of great interest for molecular vibrational analysis for inorganic and organometallic chemistry, for geological, pharmaceutical, and physical applications, polymorph screening and crystallinity analysis as well as for matrix isolation spectroscopy. In these cases, the additional far infrared region offers insight to low energy vibrations which are observable only there. This includes inorganic species, lattice vibrations or intermolecular vibrations in the ordered solid state. The spectral range of a FTIR spectrometer is defined by the major optical components such as the source, beamsplitter, and detector. The globar source covers a broad spectral range from 8000 to 20 \\wn. However a bottle neck exists with respect to the beamsplitter and detector. To extend the spectral range further into the far IR and THz spectral ranges, one or more additional far IR beam splitters and detectors have been previously required. Two new optic components have been incorporated in a spectrometer to achieve coverage of both the mid and far infrared in a single scan: a wide range MIR-FIR beam splitter and the wide range DLaTGS detector that utilizes a diamond window. The use of a standard SiC IR source with these components yields a spectral range of 6000 down to 50 \\wn in one step for all types of transmittance, reflectance and ATR measurements. Utilizing the external water cooled mercury arc high power lamp the spectral range can be ultimately extended down to 10 \\wn. Examples of application will include emission in MIR-THz range, identification of pigments, additives in polymers, and polymorphism studies.

  12. Foliar and cortex oleoresin variability of silver fir (Abies alba Mill.) in Albania. (United States)

    Zeneli, G; Tsitsimpikou, C; Petrakis, P V; Naxakis, G; Habili, D; Roussis, V


    Terpene composition of needle and cortical oleoresin from lateral shoots were analyzed by GC/MS for four Silver fir (Abies alba Mill.) populations scattered in natural species range in Albania. More than sixty compounds were detected in the needle oleoresin, which was characterized by a high content of alpha-pinene, camphene, beta-pinene, limonene and bornyl acetate. Three monoterpenes, alpha-pinene, beta-pinene and limonene, and two sesquiterpenes, beta-caryophyllene and germacrene D, comprised the majority of cortical oleoresin. The terpene composition differences among the populations that led to the recognition of two chemotypes. The needle oleoresin from the provinces of Puka, Bulqiza and Llogara were characterised by high amounts of beta-pinene, camphene and alpha-pinene and low amounts of limonene, while that from Drenova had high amounts of beta-pinene and limonene. A similar pattern was found in the cortical oleoresin with the exception of camphene that was a minor contributor. Geographical and seasonal variation between the populations was, also, investigated. Multivariate analysis of both needle and cortical oleoresin separated Drenova (southeastern population) from the other sites. When both major monoterpenes and sesquiterpenes were considered four chemical profiles could be attributed. Based on their chemical profiles, the populations can be divided into two groups: Populations with high content of beta-pinene and alpha-pinene but a low content of limonene (Puka, Bulqiza and Llogara), typical of most of A. alba populations in all its distribution range. Population with a high content of limonene and a moderate content of beta-pinene and alpha-pinene (Drenova).

  13. Using pixel and object based IKONOS image analysis for studying decay in silver fir stands

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    Ionuț Barnoaiea


    Full Text Available The problem of old silver fir stands decay has appeared in the last decades of the last century with symptoms such as defoliation, wood decay and parasites attacks. The problems in monitoring this phenomenon is that the defoliation and mistletoe attack appear on the crown top, triggering the coronation process - tree develop branches on their lower stem part in order to resist to the defoliation. This reaction makes the attack difficult to notice on aerial or satellite images, due to the reflection of light on the lower branches. The objective of the article is to find a research methodology for identifying and even mapping the mistletoe attack phenomenon at a tree or stand level. For a tree-level analysis, a comparison between the data obtained in a 1 ha sample plot and the data extracted from an IKONOS satellite image has been used. The simple spectral response is less correlated at tree level with the defoliation and mistletoe attacks. We found a very significant correlation in the infrared channel but with low correlation coefficients. Better results have been obtained in a stand-level analysis. In order to improve the separability, the methodology for extracting the remote sensing data did not resume only to the mean spectral response, but we also performed a variability and texture analysis. The obtained correlation coefficients were around 0.7, very significant for the data used in the research. The results obtained with the texture analysis are also related to the biometric characteristics of the stands, mistletoe attacks occurring in stands with a low canopy closure index and is usually accompanied by distorted crowns. The model of forest health analysis should also be tested in similar conditions in order to validate and apply on a large scale inventory.

  14. Using pixel and object based IKONOS image analysis for studying decay in silver fir stands

    Directory of Open Access Journals (Sweden)

    Ionut Barnoaiea


    Full Text Available The problem of old silver fir stands decay has appeared in the lastdecades of the last century with symptoms such as defoliation, wood decay and parasites attacks. The problems in monitoring this phenomenon is that the defoliation and mistletoe attack appear on the crown top, triggering the coronation process - tree develop branches on their lower stem part in order to resist to the defoliation. Thisreaction makes the attack difficult to notice on aerial or satellite images, due to the reflection of light on the lower branches. The objective of the article is to find a research methodology for identifying and even mapping the mistletoe attack phenomenonat a tree or stand level. For a tree-level analysis, a comparison between the data obtained in a 1 ha sample plot and the data extracted from an IKONOS satellite image has been used. The simple spectral response is less correlated at tree level with the defoliation and mistletoe attacks. We found a very significant correlation in the infrared channel but with low correlation coefficients. Better results have been obtained in a stand-level analysis. In order to improve the separability, the methodologyfor extracting the remote sensing data did not resume only to the mean spectral response, but we also performed a variability and texture analysis. The obtained correlation coefficients were around 0.7, very significant for the data used in the research. The results obtained with the texture analysis are also related to the biometriccharacteristics of the stands, mistletoe attacks occurring in stands with a low canopy closure index and is usually accompanied by distorted crowns. The model of forest health analysis should also be tested in similar conditions in order to validate and apply on a large scale inventory.

  15. Deriving Fuel Mass by Size Class in Douglas-fir (Pseudotsuga menziesii Using Terrestrial Laser Scanning

    Directory of Open Access Journals (Sweden)

    Lloyd Queen


    Full Text Available Requirements for describing coniferous forests are changing in response to wildfire concerns, bio-energy needs, and climate change interests. At the same time, technology advancements are transforming how forest properties can be measured. Terrestrial Laser Scanning (TLS is yielding promising results for measuring tree biomass parameters that, historically, have required costly destructive sampling and resulted in small sample sizes. Here we investigate whether TLS intensity data can be used to distinguish foliage and small branches (≤0.635 cm diameter; coincident with the one-hour timelag fuel size class from larger branchwood (>0.635 cm in Douglas-fir (Pseudotsuga menziesii branch specimens. We also consider the use of laser density for predicting biomass by size class. Measurements are addressed across multiple ranges and scan angles. Results show TLS capable of distinguishing fine fuels from branches at a threshold of one standard deviation above mean intensity. Additionally, the relationship between return density and biomass is linear by fuel type for fine fuels (r2 = 0.898; SE 22.7% and branchwood (r2 = 0.937; SE 28.9%, as well as for total mass (r2 = 0.940; SE 25.5%. Intensity decays predictably as scan distances increase; however, the range-intensity relationship is best described by an exponential model rather than 1/d2. Scan angle appears to have no systematic effect on fine fuel discrimination, while some differences are observed in density-mass relationships with changing angles due to shadowing.

  16. The influence of warm-season precipitation on the diel cycle of the surface energy balance and carbon dioxide at a Colorado subalpine forest site (United States)

    Burns, S. P.; Blanken, P. D.; Turnipseed, A. A.; Hu, J.; Monson, R. K.


    Precipitation changes the physical and biological characteristics of an ecosystem. Using a precipitation-based conditional sampling technique and a 14 year data set from a 25 m micrometeorological tower in a high-elevation subalpine forest, we examined how warm-season precipitation affected the above-canopy diel cycle of wind and turbulence, net radiation Rnet, ecosystem eddy covariance fluxes (sensible heat H, latent heat LE, and CO2 net ecosystem exchange NEE) and vertical profiles of scalars (air temperature Ta, specific humidity q, and CO2 dry mole fraction χc). This analysis allowed us to examine how precipitation modified these variables from hourly (i.e., the diel cycle) to multi-day time-scales (i.e., typical of a weather-system frontal passage). During mid-day we found the following: (i) even though precipitation caused mean changes on the order of 50-70 % to Rnet, H, and LE, the surface energy balance (SEB) was relatively insensitive to precipitation with mid-day closure values ranging between 90 and 110 %, and (ii) compared to a typical dry day, a day following a rainy day was characterized by increased ecosystem uptake of CO2 (NEE increased by ≈ 10 %), enhanced evaporative cooling (mid-day LE increased by ≈ 30 W m-2), and a smaller amount of sensible heat transfer (mid-day H decreased by ≈ 70 W m-2). Based on the mean diel cycle, the evaporative contribution to total evapotranspiration was, on average, around 6 % in dry conditions and between 15 and 25 % in partially wet conditions. Furthermore, increased LE lasted at least 18 h following a rain event. At night, even though precipitation (and accompanying clouds) reduced the magnitude of Rnet, LE increased from ≈ 10 to over 20 W m-2 due to increased evaporation. Any effect of precipitation on the nocturnal SEB closure and NEE was overshadowed by atmospheric phenomena such as horizontal advection and decoupling that create measurement difficulties. Above-canopy mean χc during wet conditions was

  17. The effect of warm-season precipitation on the diel cycle of the surface energy balance and carbon dioxide at a Colorado subalpine forest site (United States)

    Burns, S. P.; Blanken, P. D.; Turnipseed, A. A.; Monson, R. K.


    Precipitation changes the physical and biological characteristics of an ecosystem. Using a precipitation-based conditional sampling technique and a 14 year dataset from a 25 m micrometeorological tower in a high-elevation subalpine forest, we examined how warm-season precipitation affected the above-canopy diel cycle of wind and turbulence, net radiation Rnet, ecosystem eddy covariance fluxes (sensible heat H, latent heat LE, and CO2 net ecosystem exchange NEE) and vertical profiles of scalars (air temperature Ta, specific humidity q, and CO2 dry mole fraction χc). This analysis allowed us to examine how precipitation modified these variables from hourly (i.e., the diel cycle) to multi-day time-scales (i.e., typical of a weather-system frontal passage). During mid-day we found: (i) even though precipitation caused mean changes on the order of 50-70% to Rnet, H, and LE, the surface energy balance (SEB) was relatively insensitive to precipitation with mid-day closure values ranging between 70-80%, and (ii) compared to a typical dry day, a day following a rainy day was characterized by increased ecosystem uptake of CO2 (NEE increased by ≈ 10%), enhanced evaporative cooling (mid-day LE increased by ≈ 30 W m-2), and a smaller amount of sensible heat transfer (mid-day H decreased by ≈ 70 W m-2). Based on the mean diel cycle, the evaporative contribution to total evapotranspiration was, on average, around 6% in dry conditions and 20% in wet conditions. Furthermore, increased LE lasted at least 18 h following a rain event. At night, precipitation (and accompanying clouds) reduced Rnet and increased LE. Any effect of precipitation on the nocturnal SEB closure and NEE was overshadowed by atmospheric phenomena such as horizontal advection and decoupling that create measurement difficulties. Above-canopy mean χc during wet conditions was found to be about 2-3 μmol mol-1 larger than χc on dry days. This difference was fairly constant over the full diel cycle

  18. Volatile Profile of Croatian Lime Tree (Tilia sp., Fir Honeydew (Abies alba and Sage (Salvia officinalis Honey

    Directory of Open Access Journals (Sweden)

    Olivera Koprivnjak


    Full Text Available Volatile profiles of lime tree (Tilia sp., fir honeydew (Abies alba and sage (Salvia officinalis honey produced in Croatia have been studied by using headspace solid phase microextraction (HS-SPME followed by gas chromatography-mass spectrometry (GC-MS analysis. Melissopalynological and sensory characterization have been performed in order to check the reliability of botanical origin of the samples. In case of sage honey, sensory characteristics are reported for the first time and are rather uniform including: colour characterized as beige to jade, depending on the consistency; odour characterized as between light and medium intensity, slightly pungent and wooden; taste characterized as low sweetness, expressive acidity and apple caramel, with persistent fruity aftertaste. Characteristic volatile profiles of the analyzed honeys are described. Taking into consideration similarities with lime and fir honey volatile profiles reported in literature, characteristic volatile compounds resulting from qualitative data evaluation are proposed. Sage honey volatile profile has been reported for the first time and it was found quite different compared to the other studied honeys showing the lowest number of peaks among the studied honeys, 34. Several compounds belonging to the sage honey volatile profile have been identified for the first time in honeys. They include tetrahydro-2,2,5,5-tetramethylfuran, 3-hexenyl ester of butanoic acid, 2-methylbenzene, maltol, methyl ester of 3-furanocarboxylic acid and benzeneacetic acid. Based on the obtained results and with the lack of comparative literature data, they are proposed as characteristic volatiles for the volatile pattern of sage honey.

  19. VizieR Online Data Catalog: FIR data of IR-bright dust-obscured galaxies (DOGs) (Toba+, 2017) (United States)

    Toba, Y.; Nagao, T.; Wang, W.-H.; Matsuhara, H.; Akiyama, M.; Goto, T.; Koyama, Y.; Ohyama, Y.; Yamamura, I.


    We investigate the star-forming activity of a sample of infrared (IR)-bright dust-obscured galaxies (DOGs) that show an extreme red color in the optical and IR regime, (i-[22])AB>7.0. Combining an IR-bright DOG sample with the flux at 22μm>3.8mJy discovered by Toba & Nagao (2016ApJ...820...46T) with the IRAS faint source catalog version 2 and AKARI far-IR (FIR) all-sky survey bright source catalog version 2, we selected 109 DOGs with FIR data. For a subsample of seven IR-bright DOGs with spectroscopic redshifts (0.07DOGs and (2) the contribution of the active galactic nucleus to IR luminosity increases with IR luminosity. By comparing the stellar mass and SFR relation for our DOG sample and the literature, we found that most of the IR-bright DOGs lie significantly above the main sequence of star-forming galaxies at similar redshift, indicating that the majority of IRAS- or AKARI-detected IR-bright DOGs are starburst galaxies. (1 data file).

  20. A comparative survey of proteins from recalcitrant tissues of a non-model gymnosperm, Douglas-fir. (United States)

    Dziedzic, Jowita Anna; McDonald, Armando Gabriel


    Most research in plants and other organisms has, for the sake of convenience, focused on the use of model species to identify mechanisms that are conserved throughout the whole kingdom. Nevertheless, unique features and processes such as those related to plant cell wall and fiber formation, and to wood quality, sometimes need to be studied directly in the non-model organism of interest. Such organisms, like the economically and ecologically important gymnosperm Douglas-fir (Pseudotsuga menziesii), which is one of the crucial softwood timber species in Northern America, are often difficult to investigate. High phenolic, resin, and tannin contents in the woody tissues, as well as an incompletely sequenced genome, have contributed greatly to the species' recalcitrance for molecular biology investigations. In this study, we present a complete procedure detailing protein sample preparation, separation, and proteomic analysis based on cross-species identification of Douglas-fir. Proteins from the cambial zone, mature needles, and in vitro callus were extracted, purified, and separated via 1D and 2D SDS-PAGE. One-dimensional electrophoresis coupled with ESI-MS/MS was used for cross-species protein identification in order to evaluate the potential of this approach and reveal major differences in protein profiles among tested tissues. Identified proteins were functionally and developmentally compared. The likely contribution of these proteins to the properties of the cell wall and wood is indicated and discussed. © 2012 WILEY-VCH Verlag GmbH & Co. KGaA, Weinheim.