Sample records for parkfield cholame monarch

  1. Delayed dynamic triggering of deep tremor along the Parkfield-Cholame section of the San Andreas Fault following the 2014 M6.0 South Napa earthquake (United States)

    Peng, Zhigang; Shelly, David R.; Ellsworth, William L.


    Large, distant earthquakes are known to trigger deep tectonic tremor along the San Andreas Fault and in subduction zones. However, there are relatively few observations of triggering from regional distance earthquakes. Here we show that a small tremor episode about 12–18 km NW of Parkfield was triggered during and immediately following the passage of surface waves from the 2014 Mw 6.0 South Napa main shock. More notably, a major tremor episode followed, beginning about 12 h later, and centered SE of Parkfield near Cholame. This major episode is one of the largest seen over the past several years, containing intense activity for ~3 days and taking more than 3 weeks to return to background levels. This episode showed systematic along-strike migration at ~5 km/d, suggesting that it was driven by a slow-slip event. Our results suggest that moderate-size earthquakes are capable of triggering major tremor and deep slow slip at regional distances.

  2. Along-strike variations in fault frictional properties along the San Andreas Fault near Cholame, California from joint earthquake and low-frequency earthquake relocations (United States)

    Harrington, Rebecca M.; Cochran, Elizabeth S.; Griffiths, Emily M.; Zeng, Xiangfang; Thurber, Clifford H.


    Recent observations of low‐frequency earthquakes (LFEs) and tectonic tremor along the Parkfield–Cholame segment of the San Andreas fault suggest slow‐slip earthquakes occur in a transition zone between the shallow fault, which accommodates slip by a combination of aseismic creep and earthquakes (fault, which accommodates slip by stable sliding (>35  km depth). However, the spatial relationship between shallow earthquakes and LFEs remains unclear. Here, we present precise relocations of 34 earthquakes and 34 LFEs recorded during a temporary deployment of 13 broadband seismic stations from May 2010 to July 2011. We use the temporary array waveform data, along with data from permanent seismic stations and a new high‐resolution 3D velocity model, to illuminate the fine‐scale details of the seismicity distribution near Cholame and the relation to the distribution of LFEs. The depth of the boundary between earthquakes and LFE hypocenters changes along strike and roughly follows the 350°C isotherm, suggesting frictional behavior may be, in part, thermally controlled. We observe no overlap in the depth of earthquakes and LFEs, with an ∼5  km separation between the deepest earthquakes and shallowest LFEs. In addition, clustering in the relocated seismicity near the 2004 Mw 6.0 Parkfield earthquake hypocenter and near the northern boundary of the 1857 Mw 7.8 Fort Tejon rupture may highlight areas of frictional heterogeneities on the fault where earthquakes tend to nucleate.

  3. Shallow deformation of the San Andreas fault 5 years following the 2004 Parkfield earthquake (Mw6) combining ERS2 and Envisat InSAR. (United States)

    Bacques, Guillaume; de Michele, Marcello; Raucoules, Daniel; Aochi, Hideo; Rolandone, Frédérique


    This study focuses on the shallow deformation that occurred during the 5 years following the Parkfield earthquake (28/09/2004, Mw 6, San Andreas Fault, California). We use Synthetic Aperture Radar interferometry (InSAR) to provide precise measurements of transient deformations after the Parkfield earthquake between 2005 and 2010. We propose a method to combine both ERS2 and ENVISAT interferograms to increase the temporal data sampling. Firstly, we combine 5 years of available Synthetic Aperture Radar (SAR) acquisitions including both ERS-2 and Envisat. Secondly, we stack selected interferograms (both from ERS2 and Envisat) for measuring the temporal evolution of the ground velocities at given time intervals. Thanks to its high spatial resolution, InSAR could provide new insights on the surface fault motion behavior over the 5 years following the Parkfield earthquake. As a complement to previous studies in this area, our results suggest that shallow transient deformations affected the Creeping-Parkfield-Cholame sections of the San Andreas Fault after the 2004 Mw6 Parkfield earthquake.

  4. Analysis of nonvolcanic tremor on the San Andreas fault near Parkfield, CA using U. S. Geological Survey Parkfield Seismic Array (United States)

    Fletcher, Jon B.; Baker, Lawrence M.


    Reports by Nadeau and Dolenc (2005) that tremor had been detected near Cholame Valley spawned an effort to use UPSAR (U. S. Geological Survey Parkfield Seismic Array) to study characteristics of tremor. UPSAR was modified to record three channels of velocity at 40-50 sps continuously in January 2005 and ran for about 1 month, during which time we recorded numerous episodes of tremor. One tremor, on 21 January at 0728, was recorded with particularly high signal levels as well as another episode 3 days later. Both events were very emergent, had a frequency content between 2 and 8 Hz, and had numerous high-amplitude, short-duration arrivals within the tremor signal. Here using the first episode as an example, we discuss an analysis procedure, which yields azimuth and apparent velocity of the tremor at UPSAR. We then provide locations for both tremor episodes. The emphasis here is how the tremor episode evolves. Twelve stations were operating at the time of recording. Slowness of arrivals was determined using cross correlation of pairs of stations; the same method used in analyzing the main shock data from 28 September 2004. A feature of this analysis is that 20 s of the time series were used at a time to calculate correlation; the longer windows resulted in more consistent estimates of slowness, but lower peak correlations. These values of correlation (peaks of about 0.25), however, are similar to that obtained for the S wave of a microearthquake. Observed peaks in slowness were traced back to source locations assumed to lie on the San Andreas fault. Our inferred locations for the two tremor events cluster near the locations of previously observed tremor, south of the Cholame Valley. Tremor source depths are in the 14-24 km range, which is below the seismogenic brittle zone, but above the Moho. Estimates of error do not preclude locations below the Moho, however. The tremor signal is very emergent but contains packets that are several times larger than the background

  5. Strong-Motion Data From the Parkfield Earthquake of September 28, 2004 (United States)

    Shakal, A. F.; Borcherdt, R. D.; Graizer, V.; Haddadi, H.; Huang, M.; Lin, K.; Stephens, C.


    Very complex ground motion with high spatial variability was recorded in the near field of the M6 Parkfield earthquake of 9/28/04 by a strong motion array. The array provided the highest density of recording stations in the near field of any earthquake recorded to date. A total of 56 stations were located within 20 km of the fault; 48 were within 10 km of the fault, more than for many other earthquakes combined. Most (45) of the stations were part of a specialized array of classic analog instruments installed by CGS in the early 1980s, and 11 were digital high resolution instruments installed by the USGS. The set of recordings obtained provide a wealth of information on near field ground motion. Processing and analysis of the strong-motion data, available at, is underway. The spatial variation of the ground motion, even over relatively short distances, is great. For example, a peak acceleration of 0.30 g was recorded in the town of Parkfield, but several stations, within about 2 km, that surround this station recorded acceleration levels well over 1 g. The strong shaking at these stations, near the termination end of the rupture, is consistent with directivity focusing, as the rupture propagated from the epicenter near Gold Hill to the northwest. However, some of the strongest shaking occurs well south of the rupture, at stations near Hwy 46 at the south end of the Cholame Valley, incompatible with directivity focusing from a simple rupture. An additional aspect is that several near-fault stations have very low shaking, despite being directly over the rupturing fault. This may provide a quantitative basis to understand observed cases of low-strength buildings immediately near a fault being only slightly damaged.

  6. The October 1992 Parkfield, California, earthquake prediction (United States)

    Langbein, J.


    A magnitude 4.7 earthquake occurred near Parkfield, California, on October 20, 992, at 05:28 UTC (October 19 at 10:28 p.m. local or Pacific Daylight Time).This moderate shock, interpreted as the potential foreshock of a damaging earthquake on the San Andreas fault, triggered long-standing federal, state and local government plans to issue a public warning of an imminent magnitude 6 earthquake near Parkfield. Although the predicted earthquake did not take place, sophisticated suites of instruments deployed as part of the Parkfield Earthquake Prediction Experiment recorded valuable data associated with an unusual series of events. this article describes the geological aspects of these events, which occurred near Parkfield in October 1992. The accompnaying article, an edited version of a press conference b Richard Andrews, the Director of the California Office of Emergency Service (OES), describes governmental response to the prediction.   

  7. Navigational Strategies of Migrating Monarch Butterflies (United States)


    AFRL-OSR-VA-TR-2014-0339 NAVIGATIONAL STRATEGIES OF MIGRATING MONARCH BUTTERFLIES Steven Reppert UNIVERSITY OF MASSACHUSETTS Final Report 11/10/2014...Final Progress Statement to (Dr. Patrick Bradshaw) Contract/Grant Title: Navigational Strategies of Migrating Monarch Butterflies Contract...Grant #: FA9550-10-1-0480 Reporting Period: 01-Sept-10 to 31-Aug-14 Overview of accomplishments: Migrating monarch butterflies (Danaus

  8. Analysis of nonvolcanic tremor on the San Andreas Fault near Parkfield, CA using U.S. Geological Survey Parkfield Seismic Array (United States)

    Fletcher, Jon B.; Baker, Lawrence M.


    Reports by Nadeau and Dolenc (2005) that tremor had been detected near Cholame Valley spawned an effort to use UPSAR (U. S. Geological Survey Parkfield Seismic Array) to study characteristics of tremor. UPSAR was modified to record three channels of velocity at 40–50 sps continuously in January 2005 and ran for about 1 month, during which time we recorded numerous episodes of tremor. One tremor, on 21 January at 0728, was recorded with particularly high signal levels as well as another episode 3 days later. Both events were very emergent, had a frequency content between 2 and 8 Hz, and had numerous high-amplitude, short-duration arrivals within the tremor signal. Here using the first episode as an example, we discuss an analysis procedure, which yields azimuth and apparent velocity of the tremor at UPSAR. We then provide locations for both tremor episodes. The emphasis here is how the tremor episode evolves. Twelve stations were operating at the time of recording. Slowness of arrivals was determined using cross correlation of pairs of stations; the same method used in analyzing the main shock data from 28 September 2004. A feature of this analysis is that 20 s of the time series were used at a time to calculate correlation; the longer windows resulted in more consistent estimates of slowness, but lower peak correlations. These values of correlation (peaks of about 0.25), however, are similar to that obtained for the S wave of a microearthquake. Observed peaks in slowness were traced back to source locations assumed to lie on the San Andreas fault. Our inferred locations for the two tremor events cluster near the locations of previously observed tremor, south of the Cholame Valley. Tremor source depths are in the 14–24 km range, which is below the seismogenic brittle zone, but above the Moho. Estimates of error do not preclude locations below the Moho, however. The tremor signal is very emergent but contains packets that are several times larger than the

  9. Monarch Butterflies: Spirits of Loved Ones (United States)

    Crumpecker, Cheryl


    The study of the beautiful monarch butterfly lends itself to a vast array of subject matter, and offers the opportunity to meet a large and varied number of standards and objectives for many grade levels. Art projects featuring monarchs may include many cross-curricular units such as math (symmetry and number graphing), science (adaptation and…

  10. Teleseismically-induced tremor near Parkfield, CA - a cacophony or a symphony? (United States)

    Vidale, J. E.; Peng, Z.; Creager, K. C.; Bodin, P.


    The tremor triggered near Parkfield, CA by the 2002 Denali and 2004 Sumatra earthquakes was strong and well recorded by the dense regional CISN and the borehole HRSN networks. Peng et al. (this meeting) survey tremors triggered by a larger set of 12 regional and teleseismic events, providing a broader context. In the case of both the 2002 M7.9 Denali and 2004 M9.1 Sumatra earthquakes, the tremor emanates from at least two source regions deep within the SAF. The first source region is 40 km NW of the SAFOD in the creeping section of the SAF, and the second region is 40 km SE of the SAFOD near Cholame, close to the location where most of the non-triggered tremor has been found previously (Nadeau and Dolenc, Science, 2005). The Denali earthquake triggered tremor is in phase with the surface waves for about 400s. The northern region started tremoring first by about 100s, and both regions quieted before the end of the surface waves. The wavetrain for the 2004 M9.1 Sumatra earthquake was long enough that tremors were also excited by the weak diffracted P waves, and tremor turned up the volume for an hour upon the arrival of the surface waves, underwent a sudden and curious hiatus for 500s before the end of the surface waves, then re-started and continued for at least an hour after the passage of the surface waves. It is easy to suggest that the tremor was accompanied by deep slip on the SAF, but creep and strain data indicate any slip was too small to generate a detectable surface deformation. These observations suggest a component of driven, instantaneous, perhaps Coulomb-friction response with an added dose of self-sustaining, dribbling activity more suggestive of the oozing of fluids.

  11. Long term electromagnetic monitoring at Parkfield, CA (United States)

    Kappler, Karl Neil

    Electric and magnetic fields in the (10-4-1.0) Hz band were monitored at two sites adjacent to the San Andreas Fault near Parkfield and Hollister, California. Observed fields typically comprise natural magnetotelluric fields, with cultural and instrument noise. A data window [2002-2005], enclosing the September 28, 2004 M6 Parkfield earthquake, was analyzed to determine if anomalous electric or magnetic fields, or changes in ground conductivity, occurred before the earthquake. The data were edited, removing intervals of instrument malfunction, leaving 875 days left in the four-year period. Frequent, local spike-like disturbances were removed. The distribution of these spikes was not biased around the time of the earthquake. Signal to noise ratios, estimated via magnetotelluric processing techniques, provided an index of data quality. Plots of signal and noise amplitude spectra, showed the behavior of the ULF fields to be remarkably constant over the period of analysis. From these first-order plots, it is clear that most of the recorded energy is coherent over the spatial extent of the array. Three main statistical techniques were employed to separate local anomalous electrical or magnetic fields from the dominant coherent natural fields: transfer function estimates between components at each site were employed to subtract the dominant field, and look deeper at the 'residual' fields; the data were decomposed into principal components to identify linear combinations of array channels, which are maximally uncorrelated; the technique of canonical coherences was employed to distinguish anomalous fields which are spatially broad from anomalies which occur at a single site only, and furthermore to distinguish anomalies which are present in both the electric and magnetic fields form those which are present in only one field type. Standard remote reference apparent resistivity estimates were generated daily at Parkfield. Most of the variation was observed to be seasonal

  12. Do Healthy Monarchs Migrate Farther? Tracking Natal Origins of Parasitized vs. Uninfected Monarch Butterflies Overwintering in Mexico. (United States)

    Altizer, Sonia; Hobson, Keith A; Davis, Andrew K; De Roode, Jacobus C; Wassenaar, Leonard I


    Long-distance migration can lower parasite prevalence if strenuous journeys remove infected animals from wild populations. We examined wild monarch butterflies (Danaus plexippus) to investigate the potential costs of the protozoan Ophryocystis elektroscirrha on migratory success. We collected monarchs from two wintering sites in central Mexico to compare infection status with hydrogen isotope (δ2H) measurements as an indicator of latitude of origin at the start of fall migration. On average, uninfected monarchs had lower δ2H values than parasitized butterflies, indicating that uninfected butterflies originated from more northerly latitudes and travelled farther distances to reach Mexico. Within the infected class, monarchs with higher quantitative spore loads originated from more southerly latitudes, indicating that heavily infected monarchs originating from farther north are less likely to reach Mexico. We ruled out the alternative explanation that lower latitudes give rise to more infected monarchs prior to the onset of migration using citizen science data to examine regional differences in parasite prevalence during the summer breeding season. We also found a positive association between monarch wing area and estimated distance flown. Collectively, these results emphasize that seasonal migrations can help lower infection levels in wild animal populations. Our findings, combined with recent declines in the numbers of migratory monarchs wintering in Mexico and observations of sedentary (winter breeding) monarch populations in the southern U.S., suggest that shifts from migratory to sedentary behavior will likely lead to greater infection prevalence for North American monarchs.

  13. Do Healthy Monarchs Migrate Farther? Tracking Natal Origins of Parasitized vs. Uninfected Monarch Butterflies Overwintering in Mexico.

    Directory of Open Access Journals (Sweden)

    Sonia Altizer

    Full Text Available Long-distance migration can lower parasite prevalence if strenuous journeys remove infected animals from wild populations. We examined wild monarch butterflies (Danaus plexippus to investigate the potential costs of the protozoan Ophryocystis elektroscirrha on migratory success. We collected monarchs from two wintering sites in central Mexico to compare infection status with hydrogen isotope (δ2H measurements as an indicator of latitude of origin at the start of fall migration. On average, uninfected monarchs had lower δ2H values than parasitized butterflies, indicating that uninfected butterflies originated from more northerly latitudes and travelled farther distances to reach Mexico. Within the infected class, monarchs with higher quantitative spore loads originated from more southerly latitudes, indicating that heavily infected monarchs originating from farther north are less likely to reach Mexico. We ruled out the alternative explanation that lower latitudes give rise to more infected monarchs prior to the onset of migration using citizen science data to examine regional differences in parasite prevalence during the summer breeding season. We also found a positive association between monarch wing area and estimated distance flown. Collectively, these results emphasize that seasonal migrations can help lower infection levels in wild animal populations. Our findings, combined with recent declines in the numbers of migratory monarchs wintering in Mexico and observations of sedentary (winter breeding monarch populations in the southern U.S., suggest that shifts from migratory to sedentary behavior will likely lead to greater infection prevalence for North American monarchs.

  14. Fire creates host plant patches for monarch butterflies (United States)

    Baum, Kristen A.; Sharber, Wyatt V.


    Monarch butterflies (Danaus plexippus) depend on the presence of host plants (Asclepias spp.) within their breeding range for reproduction. In the southern Great Plains, Asclepias viridis is a perennial that flowers in May and June, and starts to senesce by August. It is locally abundant and readily used by monarchs as a host plant. We evaluated the effects of summer prescribed fire on A. viridis and the use of A. viridis by monarch butterflies. Summer prescribed fire generated a newly emergent population of A. viridis that was absent in other areas. Pre-migrant monarch butterflies laid eggs on A. viridis in summer burned plots in late August and September, allowing adequate time for a new generation of adult monarchs to emerge and migrate south to their overwintering grounds. Thus, summer prescribed fire may provide host plant patches and/or corridors for pre-migrant monarchs during a time when host plant availability may be limited in other areas. PMID:22859559

  15. Public Knowledge of Monarchs and Support for Butterfly Conservation

    Directory of Open Access Journals (Sweden)

    Jerrod Penn


    Full Text Available Pollinator populations in North America are in decline, including the iconic monarch butterfly. In order to determine if public knowledge of monarchs informs opinions on butterfly conservation, we surveyed the public to assess their knowledge of monarchs. We also asked participants about their attitudes towards general butterfly conservation and if they believe that butterfly gardens contribute to conservation. Respondents generally had some knowledge of monarchs but were unaware of monarch population declines and the necessity of milkweed to their life cycle. Respondent knowledge was correlated with more positive attitudes about butterfly conservation. Furthermore, membership in an environmental organization increased the likelihood that the participant had prior knowledge of monarchs and cared about monarch conservation. Respondent socioeconomic factors of age and sex were also significantly correlated with conservation attitudes—older and female participants had more positive attitudes towards general butterfly conservation. Interestingly, females were also less likely than males to admit having prior knowledge of monarchs, indicating that gender may also play an important role in conservation outreach efforts. Our study indicates that educational efforts need to be directed more toward individuals not already associated with an environmental organization as these individuals are predisposed to regard conservation positively.

  16. Propagating native milkweeds for restoring monarch butterfly habitat (United States)

    Thomas D. Landis; R. Kasten. Dumroese


    The number of monarch butterflies, charismatic nomads of North America, is rapidly declining. Milkweeds (Asclepias spp.), which are the sole food source for monarch caterpillars, have also experienced a decline throughout the breeding range of this butterfly. Milkweeds can be grown from seeds or vegetatively from root cuttings or rhizomes. Seed germination is often...

  17. Stress diffusion along the san andreas fault at parkfield, california. (United States)

    Malin, P E; Alvarez, M G


    Beginning in January 1990, the epicenters of microearthquakes associated with a 12-month increase in seismicity near Parkfield, California, moved northwest to southeast along the San Andreas fault. During this sequence of events, the locally variable rate of cumulative seismic moment increased. This increase implies a local increase in fault slip. These data suggest that a southeastwardly diffusing stress front propagated along the San Andreas fault at a speed of 30 to 50 kilometers per year. Evidently, this front did not load the Parkfield asperities fast enough to produce a moderate earthquake; however, a future front might do so.

  18. Fueling the fall migration of the monarch butterfly. (United States)

    Brower, Lincoln P; Fink, Linda S; Walford, Peter


    Monarch butterflies in eastern North America accumulate lipids during their fall migration to central Mexico, and use them as their energy source during a 5 month overwintering period. When and where along their migratory journey the butterflies accumulate these lipids has implications for the importance of fall nectar sources in North America. We analyzed the lipid content of 765 summer breeding and fall migrant monarch butterflies collected at 1 nectaring site in central Virginia over 4 years (1998-2001), and compared them with 16 additional published and unpublished datasets from other sites, dating back to 1941. Virginia migrants store significantly more lipid than summer butterflies, and show significant intraseason and between-year variation. None of the Virginia samples, and none of the historical samples, with one exception, had lipid levels comparable with those found in migrants that had reached Texas and northern Mexico. This evidence suggests that upon reaching Texas, the butterflies undergo a behavioral shift and spend more time nectaring. The one exceptional sample led us to the discovery that monarchs that form roosts along their migratory routes have higher lipid contents than monarchs collected while nectaring at flowers. We propose that for much of their journey monarchs are opportunistic migrants, and the variation within and between samples reflects butterflies' individual experiences. The stored lipids appear to be of less importance as fuel for the butterflies' migration than for their survival during their overwintering period, in part because soaring on favorable winds reduces the energetic cost of flying. The conservation of nectar plants in Texas and northern Mexico is crucial to sustaining the monarch's migratory spectacle, and nectar abundance throughout eastern North America is also important. As generalists in their selection of nectar sources and nectaring habitats, monarchs are unlikely to be affected by small changes in plant

  19. Conductivity Structure of the San Andreas Fault, Parkfield, Revisited (United States)

    Park, S. K.; Roberts, J. J.


    Laboratory measurements of samples of sedimentary rocks from the Parkfield syncline reveal resistivities as low as 1 ohm m when saturated with fluids comparable to those found in nearby wells. The syncline lies on the North American side of the San Andreas fault at Parkfield and plunges northwestward into the fault zone. A previous interpretation of a high resolution magnetotelluric profile across the San Andreas fault at Parkfield identified an anomalously conductive (1-3 ohm m) region just west of the fault and extending to depths of 3 km. These low resistivity rocks were inferred to be crushed rock in the fault zone that was saturated with brines. As an alternative to this interpretation, we suggest that this anomalous region is actually the Parkfield syncline and that the current trace of the San Andreas fault at Middle Mountain does not form the boundary between the Salinian block and the North American plate. Instead, that boundary is approximately 1 km west and collocated with current seismicity. This work was performed under the auspices of the U.S. Department of Energy by the University of California Lawrence Livermore National Laboratory under contract W-7405-ENG-48 and supported specifically by the Office of Basic Energy Science. Additional support was provided by the U.S. Geological Survey (USGS), Department of the Interior, under USGS Award number 03HQGR0041. The views and conclusions contained in this document are those of the authors and should not be interpreted as necessarily representing the official policies, either expressed or implied, of the U.S. Government.

  20. Coherency analysis of accelerograms recorded by the UPSAR array during the 2004 Parkfield earthquake

    DEFF Research Database (Denmark)

    Konakli, Katerina; Kiureghian, Armen Der; Dreger, Douglas


    Spatial variability of near-fault strong motions recorded by the US Geological Survey Parkfield Seismograph Array (UPSAR) during the 2004 Parkfield (California) earthquake is investigated. Behavior of the lagged coherency for two horizontal and the vertical components is analyzed by separately...

  1. Monarchs (Danaus plexippus) and milkweeds (Asclepias species): The current situation and methods for propagating milkweeds (United States)

    Tara Luna; R. Kasten Dumroese


    An international effort is under way to conserve populations of the monarch butterfly (Danaus plexippus L. [Lepidoptera: Nymphalidae]). Monarchs complete an impressive migration each year, flying from winter roosts on the California coast and the central mountains of Mexico to breeding areas throughout North America. Monarchs depend on habitats along their migratory...

  2. The USGS plan for short-term prediction of the anticipated Parkfield earthquake (United States)

    Bakun, W.H.


    Aside from the goal of better understanding the Parkfield earthquake cycle, it is the intention of the U.S Geological Survey to attempt to issue a warning shortly before the anticipated earthquake. Although short-term earthquake warnings are not yet generally feasible, the wealth of information available for the previous significant Parkfield earthquakes suggests that if the next earthquake follows the pattern of "characteristic" Parkfield shocks, such a warning might be possible. Focusing on earthquake precursors reported for the previous  "characteristic" shocks, particulary the 1934 and 1966 events, the USGS developed a plan* in late 1985 on which to base earthquake warnings for Parkfield and has assisted State, county, and local officials in the Parkfield area to prepare a coordinated, reasonable response to a warning, should one be issued. 

  3. Density estimates of monarch butterflies overwintering in central Mexico

    Directory of Open Access Journals (Sweden)

    Wayne E. Thogmartin


    Full Text Available Given the rapid population decline and recent petition for listing of the monarch butterfly (Danaus plexippus L. under the Endangered Species Act, an accurate estimate of the Eastern, migratory population size is needed. Because of difficulty in counting individual monarchs, the number of hectares occupied by monarchs in the overwintering area is commonly used as a proxy for population size, which is then multiplied by the density of individuals per hectare to estimate population size. There is, however, considerable variation in published estimates of overwintering density, ranging from 6.9–60.9 million ha−1. We develop a probability distribution for overwinter density of monarch butterflies from six published density estimates. The mean density among the mixture of the six published estimates was ∼27.9 million butterflies ha−1 (95% CI [2.4–80.7] million ha−1; the mixture distribution is approximately log-normal, and as such is better represented by the median (21.1 million butterflies ha−1. Based upon assumptions regarding the number of milkweed needed to support monarchs, the amount of milkweed (Asclepias spp. lost (0.86 billion stems in the northern US plus the amount of milkweed remaining (1.34 billion stems, we estimate >1.8 billion stems is needed to return monarchs to an average population size of 6 ha. Considerable uncertainty exists in this required amount of milkweed because of the considerable uncertainty occurring in overwinter density estimates. Nevertheless, the estimate is on the same order as other published estimates. The studies included in our synthesis differ substantially by year, location, method, and measures of precision. A better understanding of the factors influencing overwintering density across space and time would be valuable for increasing the precision of conservation recommendations.

  4. Density estimates of monarch butterflies overwintering in central Mexico (United States)

    Thogmartin, Wayne E.; Diffendorfer, James E.; Lopez-Hoffman, Laura; Oberhauser, Karen; Pleasants, John M.; Semmens, Brice X.; Semmens, Darius J.; Taylor, Orley R.; Wiederholt, Ruscena


    Given the rapid population decline and recent petition for listing of the monarch butterfly (Danaus plexippus L.) under the Endangered Species Act, an accurate estimate of the Eastern, migratory population size is needed. Because of difficulty in counting individual monarchs, the number of hectares occupied by monarchs in the overwintering area is commonly used as a proxy for population size, which is then multiplied by the density of individuals per hectare to estimate population size. There is, however, considerable variation in published estimates of overwintering density, ranging from 6.9–60.9 million ha−1. We develop a probability distribution for overwinter density of monarch butterflies from six published density estimates. The mean density among the mixture of the six published estimates was ∼27.9 million butterflies ha−1 (95% CI [2.4–80.7] million ha−1); the mixture distribution is approximately log-normal, and as such is better represented by the median (21.1 million butterflies ha−1). Based upon assumptions regarding the number of milkweed needed to support monarchs, the amount of milkweed (Asclepias spp.) lost (0.86 billion stems) in the northern US plus the amount of milkweed remaining (1.34 billion stems), we estimate >1.8 billion stems is needed to return monarchs to an average population size of 6 ha. Considerable uncertainty exists in this required amount of milkweed because of the considerable uncertainty occurring in overwinter density estimates. Nevertheless, the estimate is on the same order as other published estimates. The studies included in our synthesis differ substantially by year, location, method, and measures of precision. A better understanding of the factors influencing overwintering density across space and time would be valuable for increasing the precision of conservation recommendations.

  5. Host Diet Affects the Morphology of Monarch Butterfly Parasites. (United States)

    Hoang, Kevin; Tao, Leiling; Hunter, Mark D; de Roode, Jacobus C


    Understanding host-parasite interactions is essential for ecological research, wildlife conservation, and health management. While most studies focus on numerical traits of parasite groups, such as changes in parasite load, less focus is placed on the traits of individual parasites such as parasite size and shape (parasite morphology). Parasite morphology has significant effects on parasite fitness such as initial colonization of hosts, avoidance of host immune defenses, and the availability of resources for parasite replication. As such, understanding factors that affect parasite morphology is important in predicting the consequences of host-parasite interactions. Here, we studied how host diet affected the spore morphology of a protozoan parasite ( Ophryocystis elektroscirrha ), a specialist parasite of the monarch butterfly ( Danaus plexippus ). We found that different host plant species (milkweeds; Asclepias spp.) significantly affected parasite spore size. Previous studies have found that cardenolides, secondary chemicals in host plants of monarchs, can reduce parasite loads and increase the lifespan of infected butterflies. Adding to this benefit of high cardenolide milkweeds, we found that infected monarchs reared on milkweeds of higher cardenolide concentrations yielded smaller parasites, a potentially hidden characteristic of cardenolides that may have important implications for monarch-parasite interactions.

  6. San andreas fault zone head waves near parkfield, california. (United States)

    Ben-Zion, Y; Malin, P


    Microearthquake seismograms from the borehole seismic network on the San Andreas fault near Parkfield, California, provide three lines of evidence that first P arrivals are "head" waves refracted along the cross-fault material contrast. First, the travel time difference between these arrivals and secondary phases identified as direct P waves scales linearly with the source-receiver distance. Second, these arrivals have the emergent wave character associated in theory and practice with refracted head waves instead of the sharp first breaks associated with direct P arrivals. Third, the first motion polarities of the emergent arrivals are reversed from those of the direct P waves as predicted by the theory of fault zone head waves for slip on the San Andreas fault. The presence of fault zone head waves in local seismic network data may help account for scatter in earthquake locations and source mechanisms. The fault zone head waves indicate that the velocity contrast across the San Andreas fault near Parkfield is approximately 4 percent. Further studies of these waves may provide a way of assessing changes in the physical state of the fault system.

  7. Acceleration and volumetric strain generated by the Parkfield 2004 earthquake on the GEOS strong-motion array near Parkfield, California (United States)

    Borcherdt, Rodger D.; Johnston, Malcolm J.S.; Dietel, Christopher; Glassmoyer, Gary; Myren, Doug; Stephens, Christopher


    An integrated array of 11 General Earthquake Observation System (GEOS) stations installed near Parkfield, CA provided on scale broad-band, wide-dynamic measurements of acceleration and volumetric strain of the Parkfield earthquake (M 6.0) of September 28, 2004. Three component measurements of acceleration were obtained at each of the stations. Measurements of collocated acceleration and volumetric strain were obtained at four of the stations. Measurements of velocity at most sites were on scale only for the initial P-wave arrival. When considered in the context of the extensive set of strong-motion recordings obtained on more than 40 analog stations by the California Strong-Motion Instrumentation Program (Shakal, et al., 2004 and those on the dense array of Spudich, et al, (1988), these recordings provide an unprecedented document of the nature of the near source strong motion generated by a M 6.0 earthquake. The data set reported herein provides the most extensive set of near field broad band wide dynamic range measurements of acceleration and volumetric strain for an earthquake as large as M 6 of which the authors are aware. As a result considerable interest has been expressed in these data. This report is intended to describe the data and facilitate its use to resolve a number of scientific and engineering questions concerning earthquake rupture processes and resultant near field motions and strains. This report provides a description of the array, its scientific objectives and the strong-motion recordings obtained of the main shock. The report provides copies of the uncorrected and corrected data. Copies of the inferred velocities, displacements, and Psuedo velocity response spectra are provided. Digital versions of these recordings are accessible with information available through the internet at several locations: the National Strong-Motion Program web site (, the COSMOS Virtual Data Center Web site

  8. Sensory basis of lepidopteran migration: Focus on the monarch butterfly (United States)

    Guerra, Patrick A.; Reppert, Steven M.


    In response to seasonal habitats, migratory lepidopterans, exemplified by the monarch butterfly, have evolved migration to deal with dynamic conditions. During migration, monarchs use orientation mechanisms, exploiting a time-compensated sun compasses and a light-sensitive inclination magnetic compass to facilitate fall migration south. The sun compass is bidirectional with overwintering coldness triggering the change in orientation direction for remigration northward in the spring. The timing of the remigration and milkweed emergence in the southern US have co-evolved for propagation of the migration. Current research is uncovering the anatomical and molecular substrates that underlie migratory-relevant sensory mechanisms with the antennae being critical components. Orientation mechanisms may be detrimentally affected by environmental factors such as climate change and sensory interference from human-generated sources. PMID:25625216

  9. Non-target effects of clothianidin on monarch butterflies (United States)

    Pecenka, Jacob R.; Lundgren, Jonathan G.


    Monarch butterflies ( Danaus plexippus) frequently consume milkweed in and near agroecosystems and consequently may be exposed to pesticides like neonicotinoids. We conducted a dose response study to determine lethal and sublethal doses of clothianidin using a 36-h exposure scenario. We then quantified clothianidin levels found in milkweed leaves adjacent to maize fields. Toxicity assays revealed LC10, LC50, and LC90 values of 7.72, 15.63, and 30.70 ppb, respectively. Sublethal effects (larval size) were observed at 1 ppb. Contaminated milkweed plants had an average of 1.14 ± 0.10 ppb clothianidin, with a maximum of 4 ppb in a single plant. This research suggests that clothianidin could function as a stressor to monarch populations.

  10. Monarch (Danaus plexippus L. Nymphalidae) migration, nectar resources and fire regimes in the Ouachita Mountains of Arkansas (United States)

    D. Craig Rudolph; Charles A. Ely; Richard R. Schaefer; J. Howard Williamson; Ronald E. Thill


    Monarchs (Danaus plexippus) pass through the Ouachita Mountains in large numbers in September and October on their annual migration to overwintering sites in the Transvolcanic Belt of central Mexico. Monarchs are dependent on nectar resources to fuel their migratory movements. In the Ouachita Mountains of west-central Arkansas migrating monarchs...

  11. Moments, magnitudes, and radiated energies of non-volcanic tremor near Cholame, CA, from ground motion spectra at UPSAR (United States)

    Fletcher, J. B.; McGarr, A.


    By averaging the spectra of events within two episodes of tremor (on Jan. 21 and 24, 2005) across the 12 stations of UPSAR, we improved the S/N sufficiently to define source spectra. Analysis of eleven impulsive events revealed attenuation-corrected spectra of displacement similar to those of earthquakes, with a low-frequency plateau, a corner frequency, and a high frequency decay proportional to f-2. Seismic moments, M0, estimated from these spectra range from about 3 to 10 × 1011 N-m or moment magnitudes in the range 1.6 to 1.9. The corner frequencies range from 2.6 to 7.2 Hz and, if interpreted in the same way as for earthquakes, indicate low stress drops that vary from 0.001 to 0.04 MPa. Seismic energies, estimated from the ground motion spectra, vary from 0.2 × 105 to 4.4 × 105 J, or apparent stresses in the range 0.002 to 0.02 MPa. The low stress parameters are consistent with a weak fault zone in the lower crust at the depth of tremor. In contrast, the same analysis on a micro-earthquake, located near Cholame (depth = 10.3 km), revealed a stress drop of 0.5 MPa and an apparent stress of 0.02 MPa. Residual spectra from ω-2 model fits to the displacement spectra of the non-volcanic tremor events show peaks near 4 Hz that are not apparent in the spectra for the microearthquake nor for the spectrum of earth noise. These spectral peaks may indicate that tremor entails more than shear failure reminiscent of mechanisms, possibly entailing fluid flow, associated with volcanic tremor or deep volcanic earthquakes.

  12. Abies religiosa habitat prediction in climatic change scenarios and implications for monarch butterfly conservation in Mexico (United States)

    Cuauhtemoc Saenz-Romero; Gerald E. Rehfeldt; Pierre Duval; Roberto A. Lindig-Cisneros


    Abies religiosa (HBK) Schl. & Cham. (oyamel fir) is distributed in conifer-dominated mountain forests at high altitudes along the Trans-Mexican Volcanic Belt. This fir is the preferred host for overwintering monarch butterfly (Danaus plexippus) migratory populations which habitually congregate within a few stands now located inside a Monarch Butterfly Biosphere...

  13. Monarch butterfly oviposition preference supports a diversity of milkweed species on the landscape (United States)

    Over the past two decades, the population of monarch butterflies east of the Rocky Mountains has experienced a significant decline in overwintering numbers. Habitat restoration that includes planting milkweeds is essential to boost monarch numbers within the breeding range. Milkweeds are the only ho...

  14. National valuation of monarch butterflies indicates an untapped potential for incentive-based conservation (United States)

    Diffendorfer, Jay E.; Loomis, John B.; Ries, Leslie; Oberhauser, Karen; Semmens, Darius; Semmens, Brice; Butterfield, Bruce; Bagstad, Ken; Goldstein, Josh; Wiederholt, Ruscena; Mattsson, Brady; Thogmartin, Wayne E.


    The annual migration of monarch butterflies (Danaus plexippus) has high cultural value and recent surveys indicate monarch populations are declining. Protecting migratory species is complex because they cross international borders and depend on multiple regions. Understanding how much, and where, humans place value on migratory species can facilitate market-based conservation approaches. We performed a contingent valuation study of monarchs to understand the potential for such approaches to fund monarch conservation. The survey asked U.S. respondents about the money they would spend, or have spent, growing monarch-friendly plants, and the amount they would donate to monarch conservation organizations. Combining planting payments and donations, the survey indicated U.S. households valued monarchs as a total one-time payment of $4.78–$6.64 billion, levels similar to many endangered vertebrate species. The financial contribution of even a small percentage of households through purchases or donations could generate new funding for monarch conservation through market-based approaches.

  15. A trans-national monarch butterfly population model and implications for regional conservation priorities (United States)

    Oberhauser, Karen; Wiederholt, Ruscena; Diffendorfer, James E.; Semmens, Darius J.; Ries, Leslie; Thogmartin, Wayne E.; Lopez-Hoffman, Laura; Semmens, Brice


    1. The monarch has undergone considerable population declines over the past decade, and the governments of Mexico, Canada, and the United States have agreed to work together to conserve the species.2. Given limited resources, understanding where to focus conservation action is key for widespread species like monarchs. To support planning for continental-scale monarch habitat restoration, we address the question of where restoration efforts are likely to have the largest impacts on monarch butterfly (Danaus plexippus Linn.) population growth rates.3. We present a spatially explicit demographic model simulating the multi-generational annual cycle of the eastern monarch population, and use the model to examine management scenarios, some of which focus on particular regions of North America.4. Improving the monarch habitat in the north central or southern parts of the monarch range yields a slightly greater increase in the population growth rate than restoration in other regions. However, combining restoration efforts across multiple regions yields population growth rates above 1 with smaller simulated improvements in habitat per region than single-region strategies.5. Synthesis and applications: These findings suggest that conservation investment in projects across the full monarch range will be more effective than focusing on one or a few regions, and will require international cooperation across many land use categories.

  16. Migratory monarchs wintering in California experience low infection risk compared to monarchs breeding year-round on non-native milkweed. (United States)

    Satterfield, Dara A; Villablanca, Francis X; Maerz, John C; Altizer, Sonia


    Long-distance migration can lower infection risk for animal populations by removing infected individuals during strenuous journeys, spatially separating susceptible age classes, or allowing migrants to periodically escape from contaminated habitats. Many seasonal migrations are changing due to human activities including climate change and habitat alteration. Moreover, for some migratory populations, sedentary behaviors are becoming more common as migrants abandon or shorten their journeys in response to supplemental feeding or warming temperatures. Exploring the consequences of reduced movement for host-parasite interactions is needed to predict future responses of animal pathogens to anthropogenic change. Monarch butterflies (Danaus plexippus) and their specialist protozoan parasite Ophryocystis elektroscirrha (OE) provide a model system for examining how long-distance migration affects infectious disease processes in a rapidly changing world. Annual monarch migration from eastern North America to Mexico is known to reduce protozoan infection prevalence, and more recent work suggests that monarchs that forego migration to breed year-round on non-native milkweeds in the southeastern and south central Unites States face extremely high risk of infection. Here, we examined the prevalence of OE infection from 2013 to 2016 in western North America, and compared monarchs exhibiting migratory behavior (overwintering annually along the California coast) with those that exhibit year-round breeding. Data from field collections and a joint citizen science program of Monarch Health and Monarch Alert showed that infection frequency was over nine times higher for monarchs sampled in gardens with year-round milkweed as compared to migratory monarchs sampled at overwintering sites. Results here underscore the importance of animal migrations for lowering infection risk and motivate future studies of pathogen transmission in migratory species affected by environmental change. © The

  17. Postseismic relaxation along the San Andreas fault at Parkfield from continuous seismological observations. (United States)

    Brenguier, F; Campillo, M; Hadziioannou, C; Shapiro, N M; Nadeau, R M; Larose, E


    Seismic velocity changes and nonvolcanic tremor activity in the Parkfield area in California reveal that large earthquakes induce long-term perturbations of crustal properties in the San Andreas fault zone. The 2003 San Simeon and 2004 Parkfield earthquakes both reduced seismic velocities that were measured from correlations of the ambient seismic noise and induced an increased nonvolcanic tremor activity along the San Andreas fault. After the Parkfield earthquake, velocity reduction and nonvolcanic tremor activity remained elevated for more than 3 years and decayed over time, similarly to afterslip derived from GPS (Global Positioning System) measurements. These observations suggest that the seismic velocity changes are related to co-seismic damage in the shallow layers and to deep co-seismic stress change and postseismic stress relaxation within the San Andreas fault zone.

  18. Tracking climate impacts on the migratory monarch butterfly (United States)

    Zipkin, Elise F.; Ries, Leslie; Reeves, Rick; Regetz, James; Oberhauser, Karen S.


    Understanding the impacts of climate on migratory species is complicated by the fact that these species travel through several climates that may be changing in diverse ways throughout their complete migratory cycle. Most studies are not designed to tease out the direct and indirect effects of climate at various stages along the migration route. We assess the impacts of spring and summer climate conditions on breeding monarch butterflies, a species that completes its annual migration cycle over several generations. No single, broad-scale climate metric can explain summer breeding phenology or the substantial year-to-year fluctuations observed in population abundances. As such, we built a Poisson regression model to help explain annual arrival times and abundances in the Midwestern United States. We incorporated the climate conditions experienced both during a spring migration/breeding phase in Texas as well as during subsequent arrival and breeding during the main recruitment period in Ohio. Using data from a state-wide butterfly monitoring network in Ohio, our results suggest that climate acts in conflicting ways during the spring and summer seasons. High spring precipitation in Texas is associated with the largest annual population growth in Ohio and the earliest arrival to the summer breeding ground, as are intermediate spring temperatures in Texas. On the other hand, the timing of monarch arrivals to the summer breeding grounds is not affected by climate conditions within Ohio. Once in Ohio for summer breeding, precipitation has minimal impacts on overall abundances, whereas warmer summer temperatures are generally associated with the highest expected abundances, yet this effect is mitigated by the average seasonal temperature of each location in that the warmest sites receive no benefit of above average summer temperatures. Our results highlight the complex relationship between climate and performance for a migrating species and suggest that attempts to

  19. Dynamic mechanical oscillations during metamorphosis of the monarch butterfly (United States)

    Pelling, Andrew E; Wilkinson, Paul R; Stringer, Richard; Gimzewski, James K


    The mechanical oscillation of the heart is fundamental during insect metamorphosis, but it is unclear how morphological changes affect its mechanical dynamics. Here, the micromechanical heartbeat with the monarch chrysalis (Danaus plexippus) during metamorphosis is compared with the structural changes observed through in vivo magnetic resonance imaging (MRI). We employ a novel ultra-sensitive detection approach, optical beam deflection, in order to measure the microscale motions of the pupae during the course of metamorphosis. We observed very distinct mechanical contractions occurring at regular intervals, which we ascribe to the mechanical function of the heart organ. Motion was observed to occur in approximately 15 min bursts of activity with frequencies in the 0.4–1.0 Hz range separated by periods of quiescence during the first 83 per cent of development. In the final stages, the beating was found to be uninterrupted until the adult monarch butterfly emerged. Distinct stages of development were characterized by changes in frequency, amplitude, mechanical quality factor and de/repolarization times of the mechanical pulsing. The MRI revealed that the heart organ remains functionally intact throughout metamorphosis but undergoes morphological changes that are reflected in the mechanical oscillation. PMID:18682363

  20. Incorporating fault zone head wave and direct wave secondary arrival times into seismic tomography: Application at Parkfield, California (United States)

    Bennington, Ninfa L.; Thurber, Clifford; Peng, Zhigang; Zhang, Haijiang; Zhao, Peng


    We present a three-dimensional (3D) P wave velocity (Vp) model of the Parkfield region that utilizes existing P wave arrival time data, including fault zone head waves (FZHWs), and data from direct wave secondary arrivals (DWSAs). The first-arrival and DWSA travel times are obtained as the global- and local-minimum travel time paths, respectively. The inclusion of FZHWs and DWSAs results in as much as a 5% and a 10% increase in the across-fault velocity contrast, respectively, for the Vp model at Parkfield relative to that of Thurber et al. [2006]. Viewed along strike, three pronounced velocity contrast regions are observed: a pair of strong positive velocity contrasts (SW fast), one NW of the 1966 Parkfield earthquake hypocenter and the other SE of the 2004 Parkfield earthquake hypocenter, and a strong negative velocity contrast (NE fast) between the two hypocenters. The negative velocity contrast partially to entirely encompasses peak coseismic slip estimated in several slip models for the 2004 earthquake, suggesting that the negative velocity contrast played a part in defining the rupture patch of the 2004 Parkfield earthquake. Following Ampuero and Ben-Zion (2008), the pattern of velocity contrasts is consistent with the observed bilateral rupture propagation for the 2004 Parkfield earthquake. Although the velocity contrasts also suggest bilateral rupture propagation for the 1966 Parkfield earthquake, the fault is creeping to the NW here, i.e., exhibiting velocity-strengthening behavior. Thus, it is not surprising that rupture propagated only SE during this event.

  1. Three-component ambient noise beamforming in the Parkfield area (United States)

    Löer, Katrin; Riahi, Nima; Saenger, Erik H.


    We apply a three-component beamforming algorithm to an ambient noise data set recorded at a seismic array to extract information about both isotropic and anisotropic surface wave velocities. In particular, we test the sensitivity of the method with respect to the array geometry as well as to seasonal variations in the distribution of noise sources. In the earth's crust, anisotropy is typically caused by oriented faults or fractures and can be altered when earthquakes or human activities cause these structures to change. Monitoring anisotropy changes thus provides time-dependent information on subsurface processes, provided they can be distinguished from other effects. We analyse ambient noise data at frequencies between 0.08 and 0.52 Hz recorded at a three-component array in the Parkfield area, California (US), between 2001 November and 2002 April. During this time, no major earthquakes were identified in the area and structural changes are thus not expected. We compute dispersion curves of Love and Rayleigh waves and estimate anisotropy parameters for Love waves. For Rayleigh waves, the azimuthal source coverage is too limited to perform anisotropy analysis. For Love waves, ambient noise sources are more widely distributed and we observe significant and stable surface wave anisotropy for frequencies between 0.2 and 0.4 Hz. Synthetic data experiments indicate that the array geometry introduces apparent anisotropy, especially when waves from multiple sources arrive simultaneously at the array. Both the magnitude and the pattern of apparent anisotropy, however, differ significantly from the anisotropy observed in Love wave data. Temporal variations of anisotropy parameters observed at frequencies below 0.2 Hz and above 0.4 Hz correlate with changes in the source distribution. Frequencies between 0.2 and 0.4 Hz, however, are less affected by these variations and provide relatively stable results over the period of study.

  2. Anthropogenic Impacts on Mortality and Population Viability of the Monarch Butterfly. (United States)

    Malcolm, Stephen B


    Monarch butterflies (Danaus plexippus) are familiar herbivores of milkweeds of the genus Asclepias, and most monarchs migrate each year to locate these host plants across North American ecosystems now dominated by agriculture. Eastern migrants overwinter in high-elevation forests in Mexico, and western monarchs overwinter in trees on the coast of California. Both populations face three primary threats to their viability: (a) loss of milkweed resources for larvae due to genetically modified crops, pesticides, and fertilizers; (b) loss of nectar resources from flowering plants; and (c) degraded overwintering forest habitats due to commercially motivated deforestation and other economic activities. Secondary threats to population viability include (d) climate change effects on milkweed host plants and the dynamics of breeding, overwintering, and migration; (e) the influence of invasive plants and natural enemies; (f) habitat fragmentation and coalescence that promote homogeneous, species-depleted landscapes; and (g) deliberate culture and release of monarchs and invasive milkweeds.

  3. Interpreting surveys to estimate the size of the monarch butterfly population: Pitfalls and prospects.

    Directory of Open Access Journals (Sweden)

    John M Pleasants

    Full Text Available To assess the change in the size of the eastern North American monarch butterfly summer population, studies have used long-term data sets of counts of adult butterflies or eggs per milkweed stem. Despite the observed decline in the monarch population as measured at overwintering sites in Mexico, these studies found no decline in summer counts in the Midwest, the core of the summer breeding range, leading to a suggestion that the cause of the monarch population decline is not the loss of Midwest agricultural milkweeds but increased mortality during the fall migration. Using these counts to estimate population size, however, does not account for the shift of monarch activity from agricultural fields to non-agricultural sites over the past 20 years, as a result of the loss of agricultural milkweeds due to the near-ubiquitous use of glyphosate herbicides. We present the counter-hypotheses that the proportion of the monarch population present in non-agricultural habitats, where counts are made, has increased and that counts reflect both population size and the proportion of the population observed. We use data on the historical change in the proportion of milkweeds, and thus monarch activity, in agricultural fields and non-agricultural habitats to show why using counts can produce misleading conclusions about population size. We then separate out the shifting proportion effect from the counts to estimate the population size and show that these corrected summer monarch counts show a decline over time and are correlated with the size of the overwintering population. In addition, we present evidence against the hypothesis of increased mortality during migration. The milkweed limitation hypothesis for monarch decline remains supported and conservation efforts focusing on adding milkweeds to the landscape in the summer breeding region have a sound scientific basis.

  4. Lipid reserves and immune defense in healthy and diseased migrating monarchs Danaus plexippus

    Directory of Open Access Journals (Sweden)



    Full Text Available Recent studies suggest that the energetic demands of long-distance migration might lower the pool of resources available for costly immune defenses. Moreover, migration could amplify the costs of parasitism if animals suffering from parasite-induced damage or depleted energy reserves are less able to migrate long distances. We investigated relationships between long-distance migration, infection, and immunity in wild fall-migrating monarch butterflies Danaus plexippus. Monarchs migrate annually from eastern North America to central Mexico, accumulating lipids essential for migration and winter survival as they travel southward. Monarchs are commonly infected by the debilitating protozoan parasite Ophryocystis elektroscirrha (OE. We collected data on lipid reserves, parasite loads, and two immune measures (hemocyte concentration and phenoloxidase activity from wild monarchs migrating through north GA (USA to ask whether (1 parasite infection negatively affects lipid reserves, and (2 greater investment in lipid reserves is associated with lower immune measures. Results showed that monarchs sampled later in the fall migration had lower but not significantly different immune measures and significantly higher lipid reserves than those sampled earlier. Lipid measures correlated negatively but only nearly significantly with one measure of immune defense (phenoloxidase activity in both healthy and infected monarchs, but did not depend on monarch infection status or parasite load. These results provide weak support for a trade-off between energy reserves and immune defense in migrants, and suggest that previously-demonstrated costs of OE infection for monarch migration are not caused by depleted lipid reserves [Current Zoology 59 (3: 393–402, 2013].

  5. A Monarch Butterfly Optimization for the Dynamic Vehicle Routing Problem

    Directory of Open Access Journals (Sweden)

    Shifeng Chen


    Full Text Available The dynamic vehicle routing problem (DVRP is a variant of the Vehicle Routing Problem (VRP in which customers appear dynamically. The objective is to determine a set of routes that minimizes the total travel distance. In this paper, we propose a monarch butterfly optimization (MBO algorithm to solve DVRPs, utilizing a greedy strategy. Both migration operation and the butterfly adjusting operator only accept the offspring of butterfly individuals that have better fitness than their parents. To improve performance, a later perturbation procedure is implemented, to maintain a balance between global diversification and local intensification. The computational results indicate that the proposed technique outperforms the existing approaches in the literature for average performance by at least 9.38%. In addition, 12 new best solutions were found. This shows that this proposed technique consistently produces high-quality solutions and outperforms other published heuristics for the DVRP.

  6. The Effects of Scales on Autorotation of Monarch Butterfly Forewings (United States)

    Dechello, Nicole; Lang, Amy


    The wings of Monarch butterflies (Danus plexippus) have scales of approximately 100 micrometers that cover their wings in a roof-shingle pattern, and these scales are hypothesized to help improve flight efficiency for their long migration. The aerodynamic effects of the scales, particularly involving the leading edge vortex formation and resulting lift, were investigated by observing the natural autorotation of forewing specimen when dropped in quiescent air. A high-speed camera recorded drop tests of 32 forewings both with scales and after removal of the scales. It was found that the scales, on average, comprised 17% of the forewing mass. Tracking software was used to analyze the videos for several parameters, including descent speed and radius of rotation. NSF ECE Grant #1358991 supported the first author as an research experience for undergraduate (REU) student.

  7. Middle Miocene carnivorans from the Monarch Mill Formation, Nevada

    Directory of Open Access Journals (Sweden)

    Kent Smith


    Full Text Available he lowest part of the Monarch Mill Formation in the Middlegate basin, west-central Nevada, has yielded a middle Miocene (Barstovian Land Mammal Age vertebrate assemblage, the Eastgate local fauna. Paleobotanical evidence from nearby, nearly contemporaneous fossil leaf assemblages indicates that the Middle Miocene vegetation in the area was mixed coniferous and hardwood forest and chaparral-sclerophyllous shrubland, and suggests that the area had been uplifted to 2700–2800 m paleoaltitude before dropping later to near its present elevation of 1600 m. Thus, the local fauna provides a rare glimpse at a medium- to high-altitude vertebrate community in the intermountain western interior of North America. The local fauna includes the remains of fish, amphibians, reptiles, birds, and 25 families of mammals. Carnivorans, the focus of this study, include six taxa (three of which are new belonging to four families. Canidae are represented by the borophagine Tomarctus brevirostris and the canine Leptocyon sp. indet. The earliest record and second North American occurrence of the simocyonine ailurid Actiocyon is represented by A. parverratis sp. nov. Two new mustelids, Brevimalictis chikasha gen. et sp. nov. and Negodiaetictis rugatrulleum gen. et sp. nov., may represent Galictinae but are of uncertain subfamilial and tribal affinity. The fourth family is represented by the felid Pseudaelurus sp. indet. Tomarctus brevirostris is limited biochronologically to the Barstovian land mammal age and thus is consistent with the age indicated by other members of the Eastgate local fauna as well as by indirect tephrochronological dates previously associated with the Monarch Mill Formation. Actiocyon parverratis sp. nov. extends the temporal range of the genus Actiocyon from late Clarendonian back to the Barstovian. The Eastgate local fauna improves our understanding of mammalian successions and evolution, during and subsequent to the Mid-Miocene Climatic Optimum

  8. An Evaluation of Butterfly Gardens for Restoring Habitat for the Monarch Butterfly (Lepidoptera: Danaidae). (United States)

    Cutting, Brian T; Tallamy, Douglas W


    The eastern migratory monarch butterfly (Danaus plexippus L.) population in North America hit record low numbers during the 2013-2014 overwintering season, prompting pleas by scientists and conservation groups to plant the butterfly's milkweed host plants (Asclepias spp.) in residential areas. While planting butterfly gardens with host plants seems like an intuitive action, no previous study has directly compared larval survival in gardens and natural areas to demonstrate that gardens are suitable habitats for Lepidoptera. In this study, milkweed was planted in residential gardens and natural areas. In 2009 and 2010, plants were monitored for oviposition by monarch butterflies and survival of monarch eggs and caterpillars. Monarchs oviposited significantly more frequently in gardens than in natural sites, with 2.0 and 6.2 times more eggs per plant per observation in 2009 and 2010, respectively. There were no significant differences in overall subadult survival between gardens and natural areas. Significant differences in survival were measured for egg and larval cohorts when analyzed separately, but these were not consistent between years. These results suggest that planting gardens with suitable larval host plants can be an effective tool for restoring habitat for monarch butterflies. If planted over a large area, garden plantings may be useful as a partial mitigation for dramatic loss of monarch habitat in agricultural settings. © The Authors 2015. Published by Oxford University Press on behalf of Entomological Society of America. All rights reserved. For Permissions, please email:

  9. Climate Change May Alter Breeding Ground Distributions of Eastern Migratory Monarchs (Danaus plexippus) via Range Expansion of Asclepias Host Plants (United States)

    Lemoine, Nathan P.


    Climate change can profoundly alter species’ distributions due to changes in temperature, precipitation, or seasonality. Migratory monarch butterflies (Danaus plexippus) may be particularly susceptible to climate-driven changes in host plant abundance or reduced overwintering habitat. For example, climate change may significantly reduce the availability of overwintering habitat by restricting the amount of area with suitable microclimate conditions. However, potential effects of climate change on monarch northward migrations remain largely unknown, particularly with respect to their milkweed (Asclepias spp.) host plants. Given that monarchs largely depend on the genus Asclepias as larval host plants, the effects of climate change on monarch northward migrations will most likely be mediated by climate change effects on Asclepias. Here, I used MaxEnt species distribution modeling to assess potential changes in Asclepias and monarch distributions under moderate and severe climate change scenarios. First, Asclepias distributions were projected to extend northward throughout much of Canada despite considerable variability in the environmental drivers of each individual species. Second, Asclepias distributions were an important predictor of current monarch distributions, indicating that monarchs may be constrained as much by the availability of Asclepias host plants as environmental variables per se. Accordingly, modeling future distributions of monarchs, and indeed any tightly coupled plant-insect system, should incorporate the effects of climate change on host plant distributions. Finally, MaxEnt predictions of Asclepias and monarch distributions were remarkably consistent among general circulation models. Nearly all models predicted that the current monarch summer breeding range will become slightly less suitable for Asclepias and monarchs in the future. Asclepias, and consequently monarchs, should therefore undergo expanded northern range limits in summer months

  10. Climate change may alter breeding ground distributions of eastern migratory monarchs (Danaus plexippus via range expansion of Asclepias host plants.

    Directory of Open Access Journals (Sweden)

    Nathan P Lemoine

    Full Text Available Climate change can profoundly alter species' distributions due to changes in temperature, precipitation, or seasonality. Migratory monarch butterflies (Danaus plexippus may be particularly susceptible to climate-driven changes in host plant abundance or reduced overwintering habitat. For example, climate change may significantly reduce the availability of overwintering habitat by restricting the amount of area with suitable microclimate conditions. However, potential effects of climate change on monarch northward migrations remain largely unknown, particularly with respect to their milkweed (Asclepias spp. host plants. Given that monarchs largely depend on the genus Asclepias as larval host plants, the effects of climate change on monarch northward migrations will most likely be mediated by climate change effects on Asclepias. Here, I used MaxEnt species distribution modeling to assess potential changes in Asclepias and monarch distributions under moderate and severe climate change scenarios. First, Asclepias distributions were projected to extend northward throughout much of Canada despite considerable variability in the environmental drivers of each individual species. Second, Asclepias distributions were an important predictor of current monarch distributions, indicating that monarchs may be constrained as much by the availability of Asclepias host plants as environmental variables per se. Accordingly, modeling future distributions of monarchs, and indeed any tightly coupled plant-insect system, should incorporate the effects of climate change on host plant distributions. Finally, MaxEnt predictions of Asclepias and monarch distributions were remarkably consistent among general circulation models. Nearly all models predicted that the current monarch summer breeding range will become slightly less suitable for Asclepias and monarchs in the future. Asclepias, and consequently monarchs, should therefore undergo expanded northern range limits in

  11. Climate change may alter breeding ground distributions of eastern migratory monarchs (Danaus plexippus) via range expansion of Asclepias host plants. (United States)

    Lemoine, Nathan P


    Climate change can profoundly alter species' distributions due to changes in temperature, precipitation, or seasonality. Migratory monarch butterflies (Danaus plexippus) may be particularly susceptible to climate-driven changes in host plant abundance or reduced overwintering habitat. For example, climate change may significantly reduce the availability of overwintering habitat by restricting the amount of area with suitable microclimate conditions. However, potential effects of climate change on monarch northward migrations remain largely unknown, particularly with respect to their milkweed (Asclepias spp.) host plants. Given that monarchs largely depend on the genus Asclepias as larval host plants, the effects of climate change on monarch northward migrations will most likely be mediated by climate change effects on Asclepias. Here, I used MaxEnt species distribution modeling to assess potential changes in Asclepias and monarch distributions under moderate and severe climate change scenarios. First, Asclepias distributions were projected to extend northward throughout much of Canada despite considerable variability in the environmental drivers of each individual species. Second, Asclepias distributions were an important predictor of current monarch distributions, indicating that monarchs may be constrained as much by the availability of Asclepias host plants as environmental variables per se. Accordingly, modeling future distributions of monarchs, and indeed any tightly coupled plant-insect system, should incorporate the effects of climate change on host plant distributions. Finally, MaxEnt predictions of Asclepias and monarch distributions were remarkably consistent among general circulation models. Nearly all models predicted that the current monarch summer breeding range will become slightly less suitable for Asclepias and monarchs in the future. Asclepias, and consequently monarchs, should therefore undergo expanded northern range limits in summer months

  12. Possible deep fault slip preceding the 2004 Parkfield earthquake, inferred from detailed observations of tectonic tremor (United States)

    Shelly, David R.


    Earthquake predictability depends, in part, on the degree to which sudden slip is preceded by slow aseismic slip. Recently, observations of deep tremor have enabled inferences of deep slow slip even when detection by other means is not possible, but these data are limited to certain areas and mostly the last decade. The region near Parkfield, California, provides a unique convergence of several years of high-quality tremor data bracketing a moderate earthquake, the 2004 magnitude 6.0 event. Here, I present detailed observations of tectonic tremor from mid-2001 through 2008 that indicate deep fault slip both before and after the Parkfield earthquake that cannot be detected with surface geodetic instruments. While there is no obvious short-term precursor, I find unidirectional tremor migration accompanied by elevated tremor rates in the 3 months prior to the earthquake, which suggests accelerated creep on the fault ∼16 km beneath the eventual earthquake hypocenter.

  13. Stability and uncertainty of finite-fault slip inversions: Application to the 2004 Parkfield, California, earthquake (United States)

    Hartzell, S.; Liu, P.; Mendoza, C.; Ji, C.; Larson, K.M.


    The 2004 Parkfield, California, earthquake is used to investigate stability and uncertainty aspects of the finite-fault slip inversion problem with different a priori model assumptions. We utilize records from 54 strong ground motion stations and 13 continuous, 1-Hz sampled, geodetic instruments. Two inversion procedures are compared: a linear least-squares subfault-based methodology and a nonlinear global search algorithm. These two methods encompass a wide range of the different approaches that have been used to solve the finite-fault slip inversion problem. For the Parkfield earthquake and the inversion of velocity or displacement waveforms, near-surface related site response (top 100 m, frequencies above 1 Hz) is shown to not significantly affect the solution. Results are also insensitive to selection of slip rate functions with similar duration and to subfault size if proper stabilizing constraints are used. The linear and nonlinear formulations yield consistent results when the same limitations in model parameters are in place and the same inversion norm is used. However, the solution is sensitive to the choice of inversion norm, the bounds on model parameters, such as rake and rupture velocity, and the size of the model fault plane. The geodetic data set for Parkfield gives a slip distribution different from that of the strong-motion data, which may be due to the spatial limitation of the geodetic stations and the bandlimited nature of the strong-motion data. Cross validation and the bootstrap method are used to set limits on the upper bound for rupture velocity and to derive mean slip models and standard deviations in model parameters. This analysis shows that slip on the northwestern half of the Parkfield rupture plane from the inversion of strong-motion data is model dependent and has a greater uncertainty than slip near the hypocenter.

  14. Spatial-temporal variation of low-frequency earthquake bursts near Parkfield, California (United States)

    Wu, Chunquan; Guyer, Robert; Shelly, David R.; Trugman, D.; Frank, William; Gomberg, Joan S.; Johnson, P.


    Tectonic tremor (TT) and low-frequency earthquakes (LFEs) have been found in the deeper crust of various tectonic environments globally in the last decade. The spatial-temporal behaviour of LFEs provides insight into deep fault zone processes. In this study, we examine recurrence times from a 12-yr catalogue of 88 LFE families with ∼730 000 LFEs in the vicinity of the Parkfield section of the San Andreas Fault (SAF) in central California. We apply an automatic burst detection algorithm to the LFE recurrence times to identify the clustering behaviour of LFEs (LFE bursts) in each family. We find that the burst behaviours in the northern and southern LFE groups differ. Generally, the northern group has longer burst duration but fewer LFEs per burst, while the southern group has shorter burst duration but more LFEs per burst. The southern group LFE bursts are generally more correlated than the northern group, suggesting more coherent deep fault slip and relatively simpler deep fault structure beneath the locked section of SAF. We also found that the 2004 Parkfield earthquake clearly increased the number of LFEs per burst and average burst duration for both the northern and the southern groups, with a relatively larger effect on the northern group. This could be due to the weakness of northern part of the fault, or the northwesterly rupture direction of the Parkfield earthquake.

  15. Searching for geodetic transient slip signals along the Parkfield segment of the San Andreas Fault (United States)

    Rousset, B.; Burgmann, R.


    The Parkfield section of the San Andreas fault is at the transition between a segment locked since the 1857 Mw 7.9 Fort Tejon earthquake to its south and a creeping segment to the north. It is particularly well instrumented since it is the many previous studies have focused on studying the coseismic and postseismic phases of the two most recent earthquake cycles, the interseismic phase is exhibiting interesting dynamics at the down-dip edge of the seismogenic zone, characterized by a very large number of low frequency earthquakes (LFE) with different behaviors depending on location. Interseismic fault creep rates appear to vary over a wide range of spatial and temporal scales, from the Earth's surface to the base of crust. In this study, we take advantage of the dense Global Positioning System (GPS) network, with 77 continuous stations located within a circle of radius 80 km centered on Parkfield. We correct these time series for the co- and postseismic signals of the 2003 Mw 6.3 San Simeon and 2004 Mw 6.0 Parkfield earthquakes. We then cross-correlate the residual time series with synthetic slow-slip templates following the approach of Rousset et al. (2017). Synthetic tests with transient events contained in GPS time series with realistic noise show the limit of detection of the method. In the application with real GPS time series, the highest correlation amplitudes are compared with micro-seismicity rates, as well as tremor and LFE observations.

  16. Searching for evidence of a preferred rupture direction in small earthquakes at Parkfield (United States)

    Kane, D. L.; Shearer, P. M.; Allmann, B.; Vernon, F. L.


    Theoretical modeling of strike-slip ruptures along a bimaterial interface suggests that the interface will have a preferred rupture direction and will produce asymmetric ground motion (Shi and Ben-Zion, 2006). This could have widespread implications for earthquake source physics and for hazard analysis on mature faults because larger ground motions would be expected in the direction of rupture propagation. Studies have shown that many large global earthquakes exhibit unilateral rupture, but a consistently preferred rupture direction along faults has not been observed. Some researchers have argued that the bimaterial interface model does not apply to natural faults, noting that the rupture of the M 6 2004 Parkfield earthquake propagated in the opposite direction from previous M 6 earthquakes along that section of the San Andreas Fault (Harris and Day, 2005). We analyze earthquake spectra from the Parkfield area to look for evidence of consistent rupture directivity along the San Andreas Fault. We separate the earthquakes into spatially defined clusters and quantify the differences in high-frequency energy among earthquakes recorded at each station. Propagation path effects are minimized in this analysis because we compare earthquakes located within a small volume and recorded by the same stations. By considering a number of potential end-member models, we seek to determine if a preferred rupture direction is present among small earthquakes at Parkfield.

  17. The redder the better: wing color predicts flight performance in monarch butterflies.

    Directory of Open Access Journals (Sweden)

    Andrew K Davis

    Full Text Available The distinctive orange and black wings of monarchs (Danaus plexippus have long been known to advertise their bitter taste and toxicity to potential predators. Recent work also showed that both the orange and black coloration of this species can vary in response to individual-level and environmental factors. Here we examine the relationship between wing color and flight performance in captive-reared monarchs using a tethered flight mill apparatus to quantify butterfly flight speed, duration and distance. In three different experiments (totaling 121 individuals we used image analysis to measure body size and four wing traits among newly-emerged butterflies prior to flight trials: wing area, aspect ratio (length/width, melanism, and orange hue. Results showed that monarchs with darker orange (approaching red wings flew longer distances than those with lighter orange wings in analyses that controlled for sex and other morphometric traits. This finding is consistent with past work showing that among wild monarchs, those sampled during the fall migration are darker in hue (redder than non-migratory monarchs. Together, these results suggest that pigment deposition onto wing scales during metamorphosis could be linked with traits that influence flight, such as thorax muscle size, energy storage or metabolism. Our results reinforce an association between wing color and flight performance in insects that is suggested by past studies of wing melansim and seasonal polyphenism, and provide an important starting point for work focused on mechanistic links between insect movement and color.

  18. The Redder the Better: Wing Color Predicts Flight Performance in Monarch Butterflies (United States)

    Davis, Andrew K.; Chi, Jean; Bradley, Catherine; Altizer, Sonia


    The distinctive orange and black wings of monarchs (Danaus plexippus) have long been known to advertise their bitter taste and toxicity to potential predators. Recent work also showed that both the orange and black coloration of this species can vary in response to individual-level and environmental factors. Here we examine the relationship between wing color and flight performance in captive-reared monarchs using a tethered flight mill apparatus to quantify butterfly flight speed, duration and distance. In three different experiments (totaling 121 individuals) we used image analysis to measure body size and four wing traits among newly-emerged butterflies prior to flight trials: wing area, aspect ratio (length/width), melanism, and orange hue. Results showed that monarchs with darker orange (approaching red) wings flew longer distances than those with lighter orange wings in analyses that controlled for sex and other morphometric traits. This finding is consistent with past work showing that among wild monarchs, those sampled during the fall migration are darker in hue (redder) than non-migratory monarchs. Together, these results suggest that pigment deposition onto wing scales during metamorphosis could be linked with traits that influence flight, such as thorax muscle size, energy storage or metabolism. Our results reinforce an association between wing color and flight performance in insects that is suggested by past studies of wing melansim and seasonal polyphenism, and provide an important starting point for work focused on mechanistic links between insect movement and color. PMID:22848463

  19. Tracking multi-generational colonization of the breeding grounds by monarch butterflies in eastern North America (United States)

    Flockhart, D. T. Tyler; Wassenaar, Leonard I.; Martin, Tara G.; Hobson, Keith A.; Wunder, Michael B.; Norris, D. Ryan


    Insect migration may involve movements over multiple breeding generations at continental scales, resulting in formidable challenges to their conservation and management. Using distribution models generated from citizen scientist occurrence data and stable-carbon and -hydrogen isotope measurements, we tracked multi-generational colonization of the breeding grounds of monarch butterflies (Danaus plexippus) in eastern North America. We found that monarch breeding occurrence was best modelled with geographical and climatic variables resulting in an annual breeding distribution of greater than 12 million km2 that encompassed 99% occurrence probability. Combining occurrence models with stable isotope measurements to estimate natal origin, we show that butterflies which overwintered in Mexico came from a wide breeding distribution, including southern portions of the range. There was a clear northward progression of monarchs over successive generations from May until August when reproductive butterflies began to change direction and moved south. Fifth-generation individuals breeding in Texas in the late summer/autumn tended to originate from northern breeding areas rather than regions further south. Although the Midwest was the most productive area during the breeding season, monarchs that re-colonized the Midwest were produced largely in Texas, suggesting that conserving breeding habitat in the Midwest alone is insufficient to ensure long-term persistence of the monarch butterfly population in eastern North America. PMID:23926146

  20. Climate change and an invasive, tropical milkweed: an ecological trap for monarch butterflies. (United States)

    Faldyn, Matthew J; Hunter, Mark D; Elderd, Bret D


    While it is well established that climate change affects species distributions and abundances, the impacts of climate change on species interactions has not been extensively studied. This is particularly important for specialists whose interactions are tightly linked, such as between the monarch butterfly (Danaus plexippus) and the plant genus Asclepias, on which it depends. We used open-top chambers (OTCs) to increase temperatures in experimental plots and placed either nonnative Asclepias curassavica or native A. incarnata in each plot along with monarch larvae. We found, under current climatic conditions, adult monarchs had higher survival and mass when feeding on A. curassavica. However, under future conditions, monarchs fared much worse on A. curassavica. The decrease in adult survival and mass was associated with increasing cardenolide concentrations under warmer temperatures. Increased temperatures alone reduced monarch forewing length. Cardenolide concentrations in A. curassavica may have transitioned from beneficial to detrimental as temperature increased. Thus, the increasing cardenolide concentrations may have pushed the larvae over a tipping point into an ecological trap; whereby past environmental cues associated with increased fitness give misleading information. Given the ubiquity of specialist plant-herbivore interactions, the potential for such ecological traps to emerge as temperatures increase may have far-reaching consequences. © 2018 by the Ecological Society of America.

  1. Neo-sex Chromosomes in the Monarch Butterfly, Danaus plexippus

    Directory of Open Access Journals (Sweden)

    Andrew J. Mongue


    Full Text Available We report the discovery of a neo-sex chromosome in the monarch butterfly, Danaus plexippus, and several of its close relatives. Z-linked scaffolds in the D. plexippus genome assembly were identified via sex-specific differences in Illumina sequencing coverage. Additionally, a majority of the D. plexippus genome assembly was assigned to chromosomes based on counts of one-to-one orthologs relative to the butterfly Melitaea cinxia (with replication using two other lepidopteran species, in which genome scaffolds have been mapped to linkage groups. Sequencing coverage-based assessments of Z linkage combined with homology-based chromosomal assignments provided strong evidence for a Z-autosome fusion in the Danaus lineage, involving the autosome homologous to chromosome 21 in M. cinxia. Coverage analysis also identified three notable assembly errors resulting in chimeric Z-autosome scaffolds. Cytogenetic analysis further revealed a large W chromosome that is partially euchromatic, consistent with being a neo-W chromosome. The discovery of a neo-Z and the provisional assignment of chromosome linkage for >90% of D. plexippus genes lays the foundation for novel insights concerning sex chromosome evolution in this female-heterogametic model species for functional and evolutionary genomics.

  2. Which native milkweeds are acceptable host plants for larval monarch butterflies (Danaus plexippus) within the Midwestern U.S. (United States)

    Over the past two decades, the population of monarch butterflies east of the Rocky Mountains has experienced a significant decline. Habitat restoration within the summer breeding range is crucial to boost population numbers. Monarch butterfly larvae use milkweeds as their only host plant. However, l...

  3. Finite-Source Inversion for the 2004 Parkfield Earthquake using 3D Velocity Model Green's Functions (United States)

    Kim, A.; Dreger, D.; Larsen, S.


    We determine finite fault models of the 2004 Parkfield earthquake using 3D Green's functions. Because of the dense station coverage and detailed 3D velocity structure model in this region, this earthquake provides an excellent opportunity to examine how the 3D velocity structure affects the finite fault inverse solutions. Various studies (e.g. Michaels and Eberhart-Phillips, 1991; Thurber et al., 2006) indicate that there is a pronounced velocity contrast across the San Andreas Fault along the Parkfield segment. Also the fault zone at Parkfield is wide as evidenced by mapped surface faults and where surface slip and creep occurred in the 1966 and the 2004 Parkfield earthquakes. For high resolution images of the rupture process"Ait is necessary to include the accurate 3D velocity structure for the finite source inversion. Liu and Aurchuleta (2004) performed finite fault inversions using both 1D and 3D Green's functions for 1989 Loma Prieta earthquake using the same source paramerization and data but different Green's functions and found that the models were quite different. This indicates that the choice of the velocity model significantly affects the waveform modeling at near-fault stations. In this study, we used the P-wave velocity model developed by Thurber et al (2006) to construct the 3D Green's functions. P-wave speeds are converted to S-wave speeds and density using by the empirical relationships of Brocher (2005). Using a finite difference method, E3D (Larsen and Schultz, 1995), we computed the 3D Green's functions numerically by inserting body forces at each station. Using reciprocity, these Green's functions are recombined to represent the ground motion at each station due to the slip on the fault plane. First we modeled the waveforms of small earthquakes to validate the 3D velocity model and the reciprocity of the Green"fs function. In the numerical tests we found that the 3D velocity model predicted the individual phases well at frequencies lower than 0

  4. Decline of Monarch Butterflies Overwintering in Mexico- Is the Migratory Phenomenon at Risk? (United States)

    Brower, Lincoln; Taylor, Orley R.; Williams, Ernest H.; Slayback, Daniel; Zubieta, Raul R.; Ramirez, M. Isabel


    1.During the 2009-2010 overwintering season and following a 15-year downward trend, the total area in Mexico occupied by the eastern North American population of overwintering monarch butterflies reached an all-time low. Despite an increase, it remained low in 2010-2011. 2. Although the data set is small, the decline in abundance is statistically significant using both linear and exponential regression models. 3. Three factors appear to have contributed to reduce monarch abundance: degradation of the forest in the overwintering areas; the loss of breeding habitat in the United States due to the expansion ofGM herbicide-resistant crops, with consequent loss of milkweed host plants, as well as continued land development; and severe weather. 4. This decline calls into question the long-term survival of the monarchs' migratory phenomenon

  5. Long-term trends in midwestern milkweed abundances and their relevance to monarch butterfly declines (United States)

    Zaya, David N.; Pearse, Ian; Spyreas, Gregory


    Declines in monarch butterfly populations have prompted investigation into the sensitivity of their milkweed host plants to land-use change. Documented declines in milkweed abundance in croplands have spurred efforts to promote milkweeds in other habitats. Nevertheless, our current understanding of milkweed populations is poor. We used a long-term plant survey from Illinois to evaluate whether trends in milkweed abundance have caused monarch decline and to highlight the habitat-management practices that promote milkweeds. Milkweed abundance in natural areas has not declined precipitously, although when croplands are considered, changes in agricultural weed management have led to a 68% loss of milkweed available for monarchs across the region. Midsuccessional plant communities with few invasive species provide optimal milkweed habitat. The augmentation of natural areas and the management of existing grasslands, such as less frequent mowing and woody- and exotic-species control, may replace some of the milkweed that has been lost from croplands.

  6. Restoring monarch butterfly habitat in the Midwestern US: 'All hands on deck' (United States)

    Thogmartin, Wayne E.; Lopez-Hoffman, Laura; Rohweder, Jason; Diffendorfer, James E.; Drum, Ryan G.; Semmens, Darius J.; Black, Scott; Caldwell, Iris; Cotter, Donita; Drobney, Pauline; Jackson, Laura L.; Gale, Michael; Helmers, Doug; Hilburger, Steven B.; Howard, Elizabeth; Oberhauser, Karen S.; Pleasants, John M.; Semmens, Brice X.; Taylor, Orley R.; Ward, Patrick; Weltzin, Jake F.; Wiederholt, Ruscena


    The eastern migratory population of monarch butterflies (Danaus plexippus plexippus) has declined by >80% within the last two decades. One possible cause of this decline is the loss of ≥1.3 billion stems of milkweed (Asclepias spp.), which monarchs require for reproduction. In an effort to restore monarchs to a population goal established by the US Fish and Wildlife Service and adopted by Mexico, Canada, and the US, we developed scenarios for amending the Midwestern US landscape with milkweed. Scenarios for milkweed restoration were developed for protected area grasslands, Conservation Reserve Program land, powerline, rail and roadside rights of way, urban/suburban lands, and land in agricultural production. Agricultural land was further divided into productive and marginal cropland. We elicited expert opinion as to the biological potential (in stems per acre) for lands in these individual sectors to support milkweed restoration and the likely adoption (probability) of management practices necessary for affecting restoration. Sixteen of 218 scenarios we developed for restoring milkweed to the Midwestern US were at levels (>1.3 billion new stems) necessary to reach the monarch population goal. One of these scenarios would convert all marginal agriculture to conserved status. The other 15 scenarios converted half of marginal agriculture (730 million stems), with remaining stems contributed by other societal sectors. Scenarios without substantive agricultural participation were insufficient for attaining the population goal. Agricultural lands are essential to reaching restoration targets because they occupy 77% of all potential monarch habitat. Barring fundamental changes to policy, innovative application of economic tools such as habitat exchanges may provide sufficient resources to tip the balance of the agro-ecological landscape toward a setting conducive to both robust agricultural production and reduced imperilment of the migratory monarch butterfly.

  7. Milkweed Matters: Monarch Butterfly (Lepidoptera: Nymphalidae) Survival and Development on Nine Midwestern Milkweed Species. (United States)

    Pocius, V M; Debinski, D M; Pleasants, J M; Bidne, K G; Hellmich, R L; Brower, L P


    The population of monarch butterflies east of the Rocky Mountains has experienced a significant decline over the past 20 yr. In order to increase monarch numbers in the breeding range, habitat restoration that includes planting milkweed plants is essential. Milkweeds in the genus Asclepias and Cynanchum are the only host plants for larval monarch butterflies in North America, but larval performance and survival across nine milkweeds native to the Midwest is not well documented. We examined development and survival of monarchs from first-instar larval stages to adulthood on nine milkweed species native to Iowa. The milkweeds included Asclepias exaltata (poke milkweed) (Gentianales: Apocynaceae), Asclepias hirtella (tall green milkweed) (Gentianales: Apocynaceae), Asclepias incarnata (swamp milkweed) (Gentianales: Apocynaceae), Asclepias speciosa (showy milkweed) (Gentianales: Apocynaceae), Asclepias sullivantii (prairie milkweed) (Gentianales: Apocynaceae), Asclepias syriaca (common milkweed) (Gentianales: Apocynaceae), Asclepias tuberosa (butterfly milkweed) (Gentianales: Apocynaceae), Asclepias verticillata (whorled milkweed) (Gentianales: Apocynaceae), and Cynanchum laeve (honey vine milkweed) (Gentianales: Apocynaceae). In greenhouse experiments, fewer larvae that fed on Asclepias hirtella and Asclepias sullivantii reached adulthood compared with larvae that fed on the other milkweed species. Monarch pupal width and adult dry mass differed among milkweeds, but larval duration (days), pupal duration (days), pupal mass, pupal length, and adult wet mass were not significantly different. Both the absolute and relative adult lipids were different among milkweed treatments; these differences are not fully explained by differences in adult dry mass. Monarch butterflies can survive on all nine milkweed species, but the expected survival probability varied from 30 to 75% among the nine milkweed species. © The Author 2017. Published by Oxford University Press on behalf

  8. Restoring monarch butterfly habitat in the Midwestern US: ‘all hands on deck’ (United States)

    Thogmartin, Wayne E.; López-Hoffman, Laura; Rohweder, Jason; Diffendorfer, Jay; Drum, Ryan; Semmens, Darius; Black, Scott; Caldwell, Iris; Cotter, Donita; Drobney, Pauline; Jackson, Laura L.; Gale, Michael; Helmers, Doug; Hilburger, Steve; Howard, Elizabeth; Oberhauser, Karen; Pleasants, John; Semmens, Brice; Taylor, Orley; Ward, Patrick; Weltzin, Jake F.; Wiederholt, Ruscena


    The eastern migratory population of monarch butterflies (Danaus plexippus plexippus) has declined by >80% within the last two decades. One possible cause of this decline is the loss of ≥1.3 billion stems of milkweed (Asclepias spp.), which monarchs require for reproduction. In an effort to restore monarchs to a population goal established by the US Fish and Wildlife Service and adopted by Mexico, Canada, and the US, we developed scenarios for amending the Midwestern US landscape with milkweed. Scenarios for milkweed restoration were developed for protected area grasslands, Conservation Reserve Program land, powerline, rail and roadside rights of way, urban/suburban lands, and land in agricultural production. Agricultural land was further divided into productive and marginal cropland. We elicited expert opinion as to the biological potential (in stems per acre) for lands in these individual sectors to support milkweed restoration and the likely adoption (probability) of management practices necessary for affecting restoration. Sixteen of 218 scenarios we developed for restoring milkweed to the Midwestern US were at levels (>1.3 billion new stems) necessary to reach the monarch population goal. One of these scenarios would convert all marginal agriculture to conserved status. The other 15 scenarios converted half of marginal agriculture (730 million stems), with remaining stems contributed by other societal sectors. Scenarios without substantive agricultural participation were insufficient for attaining the population goal. Agricultural lands are essential to reaching restoration targets because they occupy 77% of all potential monarch habitat. Barring fundamental changes to policy, innovative application of economic tools such as habitat exchanges may provide sufficient resources to tip the balance of the agro-ecological landscape toward a setting conducive to both robust agricultural production and reduced imperilment of the migratory monarch butterfly.

  9. Consequences of Food Restriction for Immune Defense, Parasite Infection, and Fitness in Monarch Butterflies. (United States)

    McKay, Alexa Fritzsche; Ezenwa, Vanessa O; Altizer, Sonia


    Organisms have a finite pool of resources to allocate toward multiple competing needs, such as development, reproduction, and enemy defense. Abundant resources can support investment in multiple traits simultaneously, but limited resources might promote trade-offs between fitness-related traits and immune defenses. We asked how food restriction at both larval and adult life stages of the monarch butterfly (Danaus plexippus) affected measures of immunity, fitness, and immune-fitness interactions. We experimentally infected a subset of monarchs with a specialist protozoan parasite to determine whether parasitism further affected these relationships and whether food restriction influenced the outcome of infection. Larval food restriction reduced monarch fitness measures both within the same life stage (e.g., pupal mass) as well as later in life (e.g., adult lifespan); adult food restriction further reduced adult lifespan. Larval food restriction lowered both hemocyte concentration and phenoloxidase activity at the larval stage, and the effects of larval food restriction on phenoloxidase activity persisted when immunity was sampled at the adult stage. Adult food restriction reduced only adult phenoloxidase activity but not hemocyte concentration. Parasite spore load decreased with one measure of larval immunity, but food restriction did not increase the probability of parasite infection. Across monarchs, we found a negative relationship between larval hemocyte concentration and pupal mass, and a trade-off between adult hemocyte concentration and adult life span was evident in parasitized female monarchs. Adult life span increased with phenoloxidase activity in some subsets of monarchs. Our results emphasize that food restriction can alter fitness and immunity across multiple life stages. Understanding the consequences of resource limitation for immune defense is therefore important for predicting how increasing constraints on wildlife resources will affect fitness and

  10. Slip deficit on the san andreas fault at parkfield, california, as revealed by inversion of geodetic data. (United States)

    Segall, P; Harris, R


    A network of geodetic lines spanning the San Andreas fault near the rupture zone of the 1966 Parkfield, California, earthquake (magnitude M = 6) has been repeatedly surveyed since 1959. In the study reported here the average rates of line-length change since 1966 were inverted to determine the distribution of interseismic slip rate on the fault. These results indicate that the Parkfield rupture surface has not slipped significantly since 1966. Comparison of the geodetically determined seismic moment of the 1966 earthquake with the interseismic slip-deficit rate suggests that the strain released by the latest shock will most likely be restored between 1984 and 1989, although this may not occur until 1995. These results lend independent support to the earlier forecast of an M = 6 earthquake near Parkfield within 5 years of 1988.

  11. Remote triggering of fault-strength changes on the San Andreas fault at Parkfield. (United States)

    Taira, Taka'aki; Silver, Paul G; Niu, Fenglin; Nadeau, Robert M


    Fault strength is a fundamental property of seismogenic zones, and its temporal changes can increase or decrease the likelihood of failure and the ultimate triggering of seismic events. Although changes in fault strength have been suggested to explain various phenomena, such as the remote triggering of seismicity, there has been no means of actually monitoring this important property in situ. Here we argue that approximately 20 years of observation (1987-2008) of the Parkfield area at the San Andreas fault have revealed a means of monitoring fault strength. We have identified two occasions where long-term changes in fault strength have been most probably induced remotely by large seismic events, namely the 2004 magnitude (M) 9.1 Sumatra-Andaman earthquake and the earlier 1992 M = 7.3 Landers earthquake. In both cases, the change possessed two manifestations: temporal variations in the properties of seismic scatterers-probably reflecting the stress-induced migration of fluids-and systematic temporal variations in the characteristics of repeating-earthquake sequences that are most consistent with changes in fault strength. In the case of the 1992 Landers earthquake, a period of reduced strength probably triggered the 1993 Parkfield aseismic transient as well as the accompanying cluster of four M > 4 earthquakes at Parkfield. The fault-strength changes produced by the distant 2004 Sumatra-Andaman earthquake are especially important, as they suggest that the very largest earthquakes may have a global influence on the strength of the Earth's fault systems. As such a perturbation would bring many fault zones closer to failure, it should lead to temporal clustering of global seismicity. This hypothesis seems to be supported by the unusually high number of M >or= 8 earthquakes occurring in the few years following the 2004 Sumatra-Andaman earthquake.

  12. Variation in wing characteristics of monarch butterflies during migration: Earlier migrants have redder and more elongated wings

    Directory of Open Access Journals (Sweden)

    Satterfield Dara A.


    Full Text Available The migration of monarch butterflies (Danaus plexippus in North America has a number of parallels with long-distance bird migration, including the fact that migratory populations of monarchs have larger and more elongated forewings than residents. These characteristics likely serve to optimize flight performance in monarchs, as they also do with birds. A question that has rarely been addressed thus far in birds or monarchs is if and how wing characteristics vary within a migration season. Individuals with superior flight performance should migrate quickly, and/or with minimal stopovers, and these individuals should be at the forefront of the migratory cohort. Conversely, individuals with poor flight performance and/or low endurance would be more likely to fall behind, and these would comprise the latest migrants. Here we examined how the wing morphology of migrating monarchs varies to determine if wing characteristics of early migrants differ from late migrants. We measured forewing area, elongation (length/width, and redness, which has been shown to predict flight endurance in monarchs. Based on a collection of 75 monarchs made one entire season (fall 2010, results showed that the earliest migrants (n = 20 in this cohort had significantly redder and more elongated forewings than the latest migrants (n = 17. There was also a non-significant tendency for early migrants to have larger forewing areas. These results suggest that the pace of migration in monarchs is at least partly dependent on the properties of their wings. Moreover, these data also raise a number of questions about the ultimate fate of monarchs that fall behind

  13. A look inside the San Andreas Fault at Parkfield through vertical seismic profiling. (United States)

    Chavarria, J Andres; Malin, Peter; Catchings, Rufus D; Shalev, Eylon


    The San Andreas Fault Observatory at Depth pilot hole is located on the southwestern side of the Parkfield San Andreas fault. This observatory includes a vertical seismic profiling (VSP) array. VSP seismograms from nearby microearthquakes contain signals between the P and S waves. These signals may be P and S waves scattered by the local geologic structure. The collected scattering points form planar surfaces that we interpret as the San Andreas fault and four other secondary faults. The scattering process includes conversions between P and S waves, the strengths of which suggest large contrasts in material properties, possibly indicating the presence of cracks or fluids.

  14. The image of the blessed monarch, the Holy King of Georgia David the Builder

    Directory of Open Access Journals (Sweden)

    Efimov Vladimir Fedorovich


    Full Text Available The article considers the biography of the saint Georgian monarch, David the Builder, analyzes his actions, church, external and internal policy. Finally it draws a conclusion that all his life was dedicated to the service of God and neighbor. Thus, his life was a model of Christian Ministry, he occupied a responsible position in society.

  15. Effects of in situ climate warming on monarch caterpillar (Danaus plexippus development

    Directory of Open Access Journals (Sweden)

    Nathan P. Lemoine


    Full Text Available Climate warming will fundamentally alter basic life history strategies of many ectothermic insects. In the lab, rising temperatures increase growth rates of lepidopteran larvae but also reduce final pupal mass and increase mortality. Using in situ field warming experiments on their natural host plants, we assessed the impact of climate warming on development of monarch (Danaus plexippus larvae. Monarchs were reared on Asclepias tuberosa grown under ‘Ambient’ and ‘Warmed’ conditions. We quantified time to pupation, final pupal mass, and survivorship. Warming significantly decreased time to pupation, such that an increase of 1 °C corresponded to a 0.5 day decrease in pupation time. In contrast, survivorship and pupal mass were not affected by warming. Our results indicate that climate warming will speed the developmental rate of monarchs, influencing their ecological and evolutionary dynamics. However, the effects of climate warming on larval development in other monarch populations and at different times of year should be investigated.

  16. Milkweed: A resource for increasing stink bug parasitism and aiding insect pollinator and monarch butterfly conservation (United States)

    The flowers of milkweed species can produce a rich supply of nectar, and therefore, planting an insecticide-free milkweed habitat in agricultural farmscapes could possibly conserve monarch butterflies, bees and other insect pollinators, as well as enhance parasitism of insect pests. In peanut-cotton...

  17. Dietary risk assessment of v-ATPase A dsRNAs on monarch butterfly larvae (United States)

    The goal of this study is to assess the risks of RNA interference (RNAi)-based genetically engineered crops on a non-target arthropod, monarch butterfly, Danaus plexippus. We hypothesize that an insecticidal double-stranded (ds) RNA targeting western corn rootworm, Diabrotica virgifera virgifera, ha...

  18. Secondary Defense Chemicals in Milkweed Reduce Parasite Infection in Monarch Butterflies, Danaus plexippus. (United States)

    Gowler, Camden D; Leon, Kristoffer E; Hunter, Mark D; de Roode, Jacobus C


    In tri-trophic systems, herbivores may benefit from their host plants in fighting parasitic infections. Plants can provide parasite resistance in two contrasting ways: either directly, by interfering with the parasite, or indirectly, by increasing herbivore immunity or health. In monarch butterflies, the larval diet of milkweed strongly influences the fitness of a common protozoan parasite. Toxic secondary plant chemicals known as cardenolides correlate strongly with parasite resistance of the host, with greater cardenolide concentrations in the larval diet leading to lower parasite growth. However, milkweed cardenolides may covary with other indices of plant quality including nutrients, and a direct experimental link between cardenolides and parasite performance has not been established. To determine if the anti-parasitic activity of milkweeds is indeed due to secondary chemicals, as opposed to nutrition, we supplemented the diet of infected and uninfected monarch larvae with milkweed latex, which contains cardenolides but no nutrients. Across three experiments, increased dietary cardenolide concentrations reduced parasite growth in infected monarchs, which consequently had longer lifespans. However, uninfected monarchs showed no differences in lifespan across treatments, confirming that cardenolide-containing latex does not increase general health. Our results suggest that cardenolides are a driving force behind plant-derived resistance in this system.

  19. Monarch butterfly population decline in North America: identifying the threatening processes (United States)

    Thogmartin, Wayne E.; Wiederholt, Ruscena; Oberhauser, Karen; Drum, Ryan G.; Diffendorfer, Jay E.; Altizer, Sonia; Taylor, Orley R.; Pleasants, John M.; Semmens, Darius J.; Semmens, Brice X.; Erickson, Richard A.; Libby, Kaitlin; Lopez-Hoffman, Laura


    The monarch butterfly (Danaus plexippus) population in North America has sharply declined over the last two decades. Despite rising concern over the monarch butterfly's status, no comprehensive study of the factors driving this decline has been conducted. Using partial least-squares regressions and time-series analysis, we investigated climatic and habitat-related factors influencing monarch population size from 1993 to 2014. Potential threats included climatic factors, habitat loss (milkweed and overwinter forest), disease and agricultural insecticide use (neonicotinoids). While climatic factors, principally breeding season temperature, were important determinants of annual variation in abundance, our results indicated strong negative relationships between population size and habitat loss variables, principally glyphosate use, but also weaker negative effects from the loss of overwinter forest and breeding season use of neonicotinoids. Further declines in population size because of glyphosate application are not expected. Thus, if remaining threats to habitat are mitigated we expect climate-induced stochastic variation of the eastern migratory population of monarch butterfly around a relatively stationary population size.

  20. Using a modified time-reverse imaging technique to locate low-frequency earthquakes on the San Andreas Fault near Cholame, California (United States)

    Horstmann, Tobias; Harrington, Rebecca M.; Cochran, Elizabeth S.


    We present a new method to locate low-frequency earthquakes (LFEs) within tectonic tremor episodes based on time-reverse imaging techniques. The modified time-reverse imaging technique presented here is the first method that locates individual LFEs within tremor episodes within 5 km uncertainty without relying on high-amplitude P-wave arrivals and that produces similar hypocentral locations to methods that locate events by stacking hundreds of LFEs without having to assume event co-location. In contrast to classic time-reverse imaging algorithms, we implement a modification to the method that searches for phase coherence over a short time period rather than identifying the maximum amplitude of a superpositioned wavefield. The method is independent of amplitude and can help constrain event origin time. The method uses individual LFE origin times, but does not rely on a priori information on LFE templates and families.We apply the method to locate 34 individual LFEs within tremor episodes that occur between 2010 and 2011 on the San Andreas Fault, near Cholame, California. Individual LFE location accuracies range from 2.6 to 5 km horizontally and 4.8 km vertically. Other methods that have been able to locate individual LFEs with accuracy of less than 5 km have mainly used large-amplitude events where a P-phase arrival can be identified. The method described here has the potential to locate a larger number of individual low-amplitude events with only the S-phase arrival. Location accuracy is controlled by the velocity model resolution and the wavelength of the dominant energy of the signal. Location results are also dependent on the number of stations used and are negligibly correlated with other factors such as the maximum gap in azimuthal coverage, source–station distance and signal-to-noise ratio.

  1. Frequency-Dependent Tidal Triggering of Low Frequency Earthquakes Near Parkfield, California (United States)

    Xue, L.; Burgmann, R.; Shelly, D. R.


    The effect of small periodic stress perturbations on earthquake generation is not clear, however, the rate of low-frequency earthquakes (LFEs) near Parkfield, California has been found to be strongly correlated with solid earth tides. Laboratory experiments and theoretical analyses show that the period of imposed forcing and source properties affect the sensitivity to triggering and the phase relation of the peak seismicity rate and the periodic stress, but frequency-dependent triggering has not been quantitatively explored in the field. Tidal forcing acts over a wide range of frequencies, therefore the sensitivity to tidal triggering of LFEs provides a good probe to the physical mechanisms affecting earthquake generation. In this study, we consider the tidal triggering of LFEs near Parkfield, California since 2001. We find the LFEs rate is correlated with tidal shear stress, normal stress rate and shear stress rate. The occurrence of LFEs can also be independently modulated by groups of tidal constituents at semi-diurnal, diurnal and fortnightly frequencies. The strength of the response of LFEs to the different tidal constituents varies between LFE families. Each LFE family has an optimal triggering frequency, which does not appear to be depth dependent or systematically related to other known properties. This suggests the period of the applied forcing plays an important role in the triggering process, and the interaction of periods of loading history and source region properties, such as friction, effective normal stress and pore fluid pressure, produces the observed frequency-dependent tidal triggering of LFEs.

  2. Migratory connectivity of the monarch butterfly (Danaus plexippus: patterns of spring re-colonization in eastern North America.

    Directory of Open Access Journals (Sweden)

    Nathan G Miller

    Full Text Available Each year, millions of monarch butterflies (Danaus plexippus migrate up to 3000 km from their overwintering grounds in central Mexico to breed in eastern North America. Malcolm et al. (1993 articulated two non-mutually exclusive hypotheses to explain how Monarchs re-colonize North America each spring. The 'successive brood' hypothesis proposes that monarchs migrate from Mexico to the Gulf Coast, lay eggs and die, leaving northern re-colonization of the breeding range to subsequent generations. The 'single sweep' hypothesis proposes that overwintering monarchs continue to migrate northward after arriving on the Gulf coast and may reach the northern portion of the breeding range, laying eggs along the way. To examine these hypotheses, we sampled monarchs throughout the northern breeding range and combined stable-hydrogen isotopes (δD to estimate natal origin with wing wear scores to differentiate between individuals born in the current vs. previous year. Similar to Malcolm et al. (1993, we found that the majority of the northern breeding range was re-colonized by the first generation of monarchs (90%. We also estimated that a small number of individuals (10% originated directly from Mexico and, therefore adopted a sweep strategy. Contrary to Malcolm et al. (1993, we found that 62% of monarchs sampled in the Great Lakes originated from the Central U.S., suggesting that this region is important for sustaining production in the northern breeding areas. Our results provide new evidence of re-colonization patterns in monarchs and contribute important information towards identifying productive breeding regions of this unique migratory insect.

  3. Local and cross-seasonal associations of climate and land use with abundance of monarch butterflies Danaus plexippus (United States)

    Saunders, Sarah P.; Ries, Leslie; Oberhasuer, Karen S.; Thogmartin, Wayne E.; Zipkin, Elise F.


    Quantifying how climate and land use factors drive population dynamics at regional scales is complex because it depends on the extent of spatial and temporal synchrony among local populations, and the integration of population processes throughout a species’ annual cycle. We modeled weekly, site-specific summer abundance (1994–2013) of monarch butterflies Danaus plexippus at sites across Illinois, USA to assess relative associations of monarch abundance with climate and land use variables during the winter, spring, and summer stages of their annual cycle. We developed negative binomial regression models to estimate monarch abundance during recruitment in Illinois as a function of local climate, site-specific crop cover, and county-level herbicide (glyphosate) application. We also incorporated cross-seasonal covariates, including annual abundance of wintering monarchs in Mexico and climate conditions during spring migration and breeding in Texas, USA. We provide the first empirical evidence of a negative association between county-level glyphosate application and local abundance of adult monarchs, particularly in areas of concentrated agriculture. However, this association was only evident during the initial years of the adoption of herbicide-resistant crops (1994–2003). We also found that wetter and, to a lesser degree, cooler springs in Texas were associated with higher summer abundances in Illinois, as were relatively cool local summer temperatures in Illinois. Site-specific abundance of monarchs averaged approximately one fewer per site from 2004–2013 than during the previous decade, suggesting a recent decline in local abundance of monarch butterflies on their summer breeding grounds in Illinois. Our results demonstrate that seasonal climate and land use are associated with trends in adult monarch abundance, and our approach highlights the value of considering fine-resolution temporal fluctuations in population-level responses to environmental

  4. An experimental displacement and over 50 years of tag-recoveries show that monarch butterflies are not true navigators. (United States)

    Mouritsen, Henrik; Derbyshire, Rachael; Stalleicken, Julia; Mouritsen, Ole Ø; Frost, Barrie J; Norris, D Ryan


    Monarch butterflies (Danaus plexippus) breeding in eastern North America are famous for their annual fall migration to their overwintering grounds in Mexico. However, the mechanisms they use to successfully reach these sites remain poorly understood. Here, we test whether monarchs are true navigators who can determine their location relative to their final destination using both a "compass" and a "map". Using flight simulators, we recorded the orientation of wild-caught monarchs in southwestern Ontario and found that individuals generally flew in a southwest direction toward the wintering grounds. When displaced 2,500 km to the west, the same individuals continued to fly in a general southwest direction, suggesting that monarchs use a simple vector-navigation strategy (i.e., use a specific compass bearing without compensating for displacement). Using over 5 decades of field data, we also show that the directional concentration and the angular SD of recoveries from tagged monarchs largely conformed to two mathematical models describing the directional distribution of migrants expected under a vector-navigation strategy. A third analysis of tagged recoveries shows that the increasing directionality of migration from north to south is largely because of the presence of geographic barriers that guide individuals toward overwintering sites. Our work suggests that monarchs breeding in eastern North America likely combine simple orientation mechanisms with geographic features that funnel them toward Mexican overwintering sites, a remarkable achievement considering that these butterflies weigh less than a gram and travel thousands of kilometers to a site they have never seen.

  5. Cryptochromes define a novel circadian clock mechanism in monarch butterflies that may underlie sun compass navigation.

    Directory of Open Access Journals (Sweden)

    Haisun Zhu


    Full Text Available The circadian clock plays a vital role in monarch butterfly (Danaus plexippus migration by providing the timing component of time-compensated sun compass orientation, a process that is important for successful navigation. We therefore evaluated the monarch clockwork by focusing on the functions of a Drosophila-like cryptochrome (cry, designated cry1, and a vertebrate-like cry, designated cry2, that are both expressed in the butterfly and by placing these genes in the context of other relevant clock genes in vivo. We found that similar temporal patterns of clock gene expression and protein levels occur in the heads, as occur in DpN1 cells, of a monarch cell line that contains a light-driven clock. CRY1 mediates TIMELESS degradation by light in DpN1 cells, and a light-induced TIMELESS decrease occurs in putative clock cells in the pars lateralis (PL in the brain. Moreover, monarch cry1 transgenes partially rescue both biochemical and behavioral light-input defects in cry(b mutant Drosophila. CRY2 is the major transcriptional repressor of CLOCK:CYCLE-mediated transcription in DpN1 cells, and endogenous CRY2 potently inhibits transcription without involvement of PERIOD. CRY2 is co-localized with clock proteins in the PL, and there it translocates to the nucleus at the appropriate time for transcriptional repression. We also discovered CRY2-positive neural projections that oscillate in the central complex. The results define a novel, CRY-centric clock mechanism in the monarch in which CRY1 likely functions as a blue-light photoreceptor for entrainment, whereas CRY2 functions within the clockwork as the transcriptional repressor of a negative transcriptional feedback loop. Our data further suggest that CRY2 may have a dual role in the monarch butterfly's brain-as a core clock element and as an output that regulates circadian activity in the central complex, the likely site of the sun compass.

  6. Seismomagnetic effects from the long-awaited 28 September 2004 M 6.0 parkfield earthquake (United States)

    Johnston, M.J.S.; Sasai, Y.; Egbert, G.D.; Mueller, R.J.


    Precise measurements of local magnetic fields have been obtained with a differentially connected array of seven synchronized proton magnetometers located along 60 km of the locked-to-creeping transition region of the San Andreas fault at Parkfield, California, since 1976. The M 6.0 Parkfield earthquake on 28 September 2004, occurred within this array and generated coseismic magnetic field changes of between 0.2 and 0.5 nT at five sites in the network. No preseismic magnetic field changes exceeding background noise levels are apparent in the magnetic data during the month, week, and days before the earthquake (or expected in light of the absence of measurable precursive deformation, seismicity, or pore pressure changes). Observations of electric and magnetic fields from 0.01 to 20 Hz are also made at one site near the end of the earthquake rupture and corrected for common-mode signals from the ionosphere/magnetosphere using a second site some 115 km to the northwest along the fault. These magnetic data show no indications of unusual noise before the earthquake in the ULF band (0.01-20 Hz) as suggested may have preceded the 1989 ML 7.1 Loma Prieta earthquake. Nor do we see electric field changes similar to those suggested to occur before earthquakes of this magnitude from data in Greece. Uniform and variable slip piezomagnetic models of the earthquake, derived from strain, displacement, and seismic data, generate magnetic field perturbations that are consistent with those observed by the magnetometer array. A higher rate of longer-term magnetic field change, consistent with increased loading in the region, is apparent since 1993. This accompanied an increased rate of secular shear strain observed on a two-color EDM network and a small network of borehole tensor strainmeters and increased seismicity dominated by three M 4.5-5 earthquakes roughly a year apart in 1992, 1993, and 1994. Models incorporating all of these data indicate increased slip at depth in the region

  7. Fitness costs of animal medication: antiparasitic plant chemicals reduce fitness of monarch butterfly hosts. (United States)

    Tao, Leiling; Hoang, Kevin M; Hunter, Mark D; de Roode, Jacobus C


    The emerging field of ecological immunology demonstrates that allocation by hosts to immune defence against parasites is constrained by the costs of those defences. However, the costs of non-immunological defences, which are important alternatives to canonical immune systems, are less well characterized. Estimating such costs is essential for our understanding of the ecology and evolution of alternative host defence strategies. Many animals have evolved medication behaviours, whereby they use antiparasitic compounds from their environment to protect themselves or their kin from parasitism. Documenting the costs of medication behaviours is complicated by natural variation in the medicinal components of diets and their covariance with other dietary components, such as macronutrients. In the current study, we explore the costs of the usage of antiparasitic compounds in monarch butterflies (Danaus plexippus), using natural variation in concentrations of antiparasitic compounds among plants. Upon infection by their specialist protozoan parasite Ophryocystis elektroscirrha, monarch butterflies can selectively oviposit on milkweed with high foliar concentrations of cardenolides, secondary chemicals that reduce parasite growth. Here, we show that these antiparasitic cardenolides can also impose significant costs on both uninfected and infected butterflies. Among eight milkweed species that vary substantially in their foliar cardenolide concentration and composition, we observed the opposing effects of cardenolides on monarch fitness traits. While high foliar cardenolide concentrations increased the tolerance of monarch butterflies to infection, they reduced the survival rate of caterpillars to adulthood. Additionally, although non-polar cardenolide compounds decreased the spore load of infected butterflies, they also reduced the life span of uninfected butterflies, resulting in a hump-shaped curve between cardenolide non-polarity and the life span of infected butterflies

  8. Clustering and periodic recurrence of microearthquakes on the san andreas fault at parkfield, california. (United States)

    Nadeau, R M; Foxall, W; McEvilly, T V


    The San Andreas fault at Parkfield, California, apparently late in an interval between repeating magnitude 6 earthquakes, is yielding to tectonic loading partly by seismic slip concentrated in a relatively sparse distribution of small clusters (<20-meter radius) of microearthquakes. Within these clusters, which account for 63% of the earthquakes in a 1987-92 study interval, virtually identical small earthquakes occurred with a regularity that can be described by the statistical model used previously in forecasting large characteristic earthquakes. Sympathetic occurrence of microearthquakes in nearby clusters was observed within a range of about 200 meters at communication speeds of 10 to 100 centimeters per second. The rate of earthquake occurrence, particularly at depth, increased significantly during the study period, but the fraction of earthquakes that were cluster members decreased.

  9. Detection of aseismic creep along the San Andreas fault near Parkfield, California with ERS-1 radar interferometry (United States)

    Werner, Charles L.; Rosen, Paul; Hensley, Scott; Fielding, Eric; Buckley, Sean


    The differential interferometric analysis of ERS data from Parkfield (CA) observations revealed the wide area distribution of creep along the moving fault segment of the San Andreas fault over a 15 month interval. The removal of the interferometric phase related to the surface topography was carried out. The fault was clearly visible in the differential interferogram. The magnitude of the tropospheric water vapor phase distortions is greater than the signal and hinders quantitative analysis beyond order of magnitude calculations.

  10. Monarchical Activities of the Yoruba Kings of South Western Nigeria: A Cultural Heritage in Printmaking Visual Documentary.

    Directory of Open Access Journals (Sweden)

    Emmanuel Bankole Oladumiye


    Full Text Available Printmaking is a visual documentary media of art which was used as a medium of expression in analyzing myth and mythology monarchical activities of the Yorubas in South Western Nigeria in this study. The  monarchical activities of the Yoruba Kings, is  the cultural heritage and legacy that people do guide jealously and considered to be of high cultural value. The Yoruba Kings of South Western Nigeria are traditional entity which passed through the rites of installing kings for the throne fore fathers as a leader with symbol of authority between the people and the spirit world. The kings in Yoruba kingdom is so much respected that they are seen as divine and representative of God on earth and they are exalted into the position of deity because of his monarchical duties to his subjects at large. The funfairs that accompany the monarch roles  are worth documenting using printmaking as vehicle of visual and historical expression of myths and mythologies demonstrating African culture which stands out as sacred. The discourse also relies on oral testimonies written and archival documents. The materials used for the execution of the prints are rubber, wood, plate, offset printing inks and glass which records the events as an alternative to the use of photographic documentation. The research examine the philosophy behind the monarchical roles of the Yoruba Kings in print visuals based on the cultural heritage of the Yoruba people it employs an exploratory qualitative methods rely on literature review.

  11. Characterization of the San Andreas Fault near Parkfield, California by fault-zone trapped waves (United States)

    Li, Y.; Vidale, J.; Cochran, E.


    In October, 2002, coordinated by the Pre-EarthScope/SAFOD, we conducted an extensive seismic experiment at the San Andreas fault (SAF), Parkfield to record fault-zone trapped waves generated by explosions and microearthquakes using dense linear seismic arrays of 52 PASSCAL 3-channel REFTEKs deployed across and along the fault zone. We detonated 3 explosions within and out of the fault zone during the experiment, and also recorded other 13 shots of PASO experiment of UWM/RPI (Thurber and Roecker) detonated around the SAFOD drilling site at the same time. We observed prominent fault-zone trapped waves with large amplitudes and long duration following S waves at stations close to the main fault trace for sources located within and close to the fault zone. Dominant frequencies of trapped waves are 2-3 Hz for near-surface explosions and 4-5 Hz for microearthquakes. Fault-zone trapped waves are relatively weak on the north strand of SAF for same sources. In contrast, seismograms registered for both the stations and shots far away from the fault zone show a brief S wave and lack of trapped waves. These observations are consistent with previous findings of fault-zone trapped waves at the SAF [Li et al., 1990; 1997], indicating the existence of a well-developed low-velocity waveguide along the main fault strand (principal slip plan) of the SAF. The data from denser arrays and 3-D finite-difference simulations of fault-zone trapped waves allowed us to delineate the internal structure, segmentation and physical properties of the SAF with higher resolution. The trapped-wave inferred waveguide on the SAF Parkfield segment is ~150 m wide at surface and tapers to ~100 m at seismogenic depth, in which Q is 20-50 and S velocities are reduced by 30-40% from wall-rock velocities, with the greater velocity reduction at the shallow depth and to southeast of the 1966 M6 epicenter. We interpret this low-velocity waveguide on the SAF main strand as being the remnant of damage zone caused

  12. Defining behavioral and molecular differences between summer and migratory monarch butterflies (United States)

    Zhu, Haisun; Gegear, Robert J; Casselman, Amy; Kanginakudru, Sriramana; Reppert, Steven M


    Background In the fall, Eastern North American monarch butterflies (Danaus plexippus) undergo a magnificent long-range migration. In contrast to spring and summer butterflies, fall migrants are juvenile hormone deficient, which leads to reproductive arrest and increased longevity. Migrants also use a time-compensated sun compass to help them navigate in the south/southwesterly direction en route for Mexico. Central issues in this area are defining the relationship between juvenile hormone status and oriented flight, critical features that differentiate summer monarchs from fall migrants, and identifying molecular correlates of behavioral state. Results Here we show that increasing juvenile hormone activity to induce summer-like reproductive development in fall migrants does not alter directional flight behavior or its time-compensated orientation, as monitored in a flight simulator. Reproductive summer butterflies, in contrast, uniformly fail to exhibit directional, oriented flight. To define molecular correlates of behavioral state, we used microarray analysis of 9417 unique cDNA sequences. Gene expression profiles reveal a suite of 40 genes whose differential expression in brain correlates with oriented flight behavior in individual migrants, independent of juvenile hormone activity, thereby molecularly separating fall migrants from summer butterflies. Intriguing genes that are differentially regulated include the clock gene vrille and the locomotion-relevant tyramine beta hydroxylase gene. In addition, several differentially regulated genes (37.5% of total) are not annotated. We also identified 23 juvenile hormone-dependent genes in brain, which separate reproductive from non-reproductive monarchs; genes involved in longevity, fatty acid metabolism, and innate immunity are upregulated in non-reproductive (juvenile-hormone deficient) migrants. Conclusion The results link key behavioral traits with gene expression profiles in brain that differentiate migratory

  13. 3D P and S Wave Velocity Structure and Tremor Locations in the Parkfield Region (United States)

    Zeng, X.; Thurber, C. H.; Shelly, D. R.; Bennington, N. L.; Cochran, E. S.; Harrington, R. M.


    We have assembled a new dataset to refine the 3D seismic velocity model in the Parkfield region. The S arrivals from 184 earthquakes recorded by the Parkfield Experiment to Record MIcroseismicity and Tremor array (PERMIT) during 2010-2011 were picked by a new S wave picker, which is based on machine learning. 74 blasts have been assigned to four quarries, whose locations were identified with Google Earth. About 1000 P and S wave arrivals from these blasts at permanent seismic network were also incorporated. Low frequency earthquakes (LFEs) occurring within non-volcanic tremor (NVT) are valuable for improving the precision of NVT location and the seismic velocity model at greater depths. Based on previous work (Shelley and Hardebeck, 2010), waveforms of hundreds of LFEs in same family were stacked to improve signal qualify. In a previous study (McClement et al., 2013), stacked traces of more than 30 LFE families at the Parkfileld Array Seismic Observatory (PASO) have been picked. We expanded our work to include LFEs recorded by the PERMIT array. The time-frequency Phase Weight Stacking (tf-PWS) method was introduced to improve the stack quality, as direct stacking does not produce clear S-wave arrivals on the PERMIT stations. This technique uses the coherence of the instantaneous phase among the stacked signals to enhance the signal-to-noise ratio (SNR) of the stack. We found that it is extremely effective for picking LFE arrivals (Thurber et al., 2014). More than 500 P and about 1000 S arrivals from 58 LFE families were picked at the PERMIT and PASO arrays. Since the depths of LFEs are much deeper than earthquakes, we are able to extend model resolution to lower crustal depths. Both P and S wave velocity structure have been obtained with the tomoDD method. The result suggests that there is a low velocity zone (LVZ) in the lower crust and the location of the LVZ is consistent with the high conductivity zone beneath the southern segment of the Rinconada fault that

  14. An experimental displacement and over 50 years of tag-recoveries show that monarch butterflies are not true navigators

    DEFF Research Database (Denmark)

    Mourtisen, Henrik; Derbyshirec, Rachael; Stalleickena, Julia


    grounds. When displaced 2,500 km to the west, the same individuals continued to fly in a general southwest direction, suggesting that monarchs use a simple vector-navigation strategy (i.e., use a specific compass bearing without compensating for displacement). Using over 5 decades of field data, we also...... directionality of migration from north to south is largely because of the presence of geographic barriers that guide individuals toward overwintering sites. Our work suggests that monarchs breeding in eastern North America likely combine simple orientation mechanisms with geographic features that funnel them...

  15. Discordant timing between antennae disrupts sun compass orientation in migratory monarch butterflies (United States)

    Guerra, Patrick A; Merlin, Christine; Gegear, Robert J; Reppert, Steven M


    To navigate during their long-distance migration, monarch butterflies (Danaus plexippus) use a time-compensated sun compass. The sun compass timing elements reside in light-entrained circadian clocks in the antennae. Here we show that either antenna is sufficient for proper time compensation. However, migrants with either antenna painted black (to block light entrainment) and the other painted clear (to permit light entrainment) display disoriented group flight. Remarkably, when the black-painted antenna is removed, re-flown migrants with a single, clear-painted antenna exhibit proper orientation behaviour. Molecular correlates of clock function reveal that period and timeless expression is highly rhythmic in brains and clear-painted antennae, while rhythmic clock gene expression is disrupted in black-painted antennae. Our work shows that clock outputs from each antenna are processed and integrated together in the monarch time-compensated sun compass circuit. This dual timing system is a novel example of the regulation of a brain-driven behaviour by paired organs. PMID:22805565

  16. Forbs: Foundation for restoration of monarch butterflies, other pollinators, and greater sage-grouse in the western United States (United States)

    Kas Dumroese; Tara Luna; Jeremy Pinto; Thomas D. Landis


    Monarch butterflies (Danaus plexippus), other pollinators, and Greater Sage-Grouse (Centrocercus urophasianus) are currently the focus of increased conservation efforts. Federal attention on these fauna is encouraging land managers to develop conservation strategies, often without corresponding financial resources. This could foster a myopic approach when...

  17. Establishment of a Monarch Butterfly (Danaus plexippus, Lepidoptera: Danaidae) Cell Line and its Susceptibility to Insect Viruses (United States)

    A cell line from the monarch butterfly Danaus plexippus designated BCIRL-DP-AM/JG was established from adult ovaries. The cell line consisted mainly of round cells and took a prolonged period of time in the growth medium ExCell 401 containing 10% fetal bovine serum and antibiotics before it could be...

  18. Quasi-extinction risk and population targets for the Eastern, migratory population of monarch butterflies (Danaus plexippus) (United States)

    Semmens, Brice X.; Semmens, Darius J.; Thogmartin, Wayne E.; Wiederholt, Ruscena; Lopez-Hoffman, Laura; Diffendorfer, James E.; Pleasants, John M.; Oberhauser, Karen S.; Taylor, Orley R.


    The Eastern, migratory population of monarch butterflies (Danaus plexippus), an iconic North American insect, has declined by ~80% over the last decade. The monarch’s multi-generational migration between overwintering grounds in central Mexico and the summer breeding grounds in the northern U.S. and southern Canada is celebrated in all three countries and creates shared management responsibilities across North America. Here we present a novel Bayesian multivariate auto-regressive state-space model to assess quasi-extinction risk and aid in the establishment of a target population size for monarch conservation planning. We find that, given a range of plausible quasi-extinction thresholds, the population has a substantial probability of quasi-extinction, from 11–57% over 20 years, although uncertainty in these estimates is large. Exceptionally high population stochasticity, declining numbers, and a small current population size act in concert to drive this risk. An approximately 5-fold increase of the monarch population size (relative to the winter of 2014–15) is necessary to halve the current risk of quasi-extinction across all thresholds considered. Conserving the monarch migration thus requires active management to reverse population declines, and the establishment of an ambitious target population size goal to buffer against future environmentally driven variability.

  19. Chasing migration genes: a brain expressed sequence tag resource for summer and migratory monarch butterflies (Danaus plexippus.

    Directory of Open Access Journals (Sweden)

    Haisun Zhu


    Full Text Available North American monarch butterflies (Danaus plexippus undergo a spectacular fall migration. In contrast to summer butterflies, migrants are juvenile hormone (JH deficient, which leads to reproductive diapause and increased longevity. Migrants also utilize time-compensated sun compass orientation to help them navigate to their overwintering grounds. Here, we describe a brain expressed sequence tag (EST resource to identify genes involved in migratory behaviors. A brain EST library was constructed from summer and migrating butterflies. Of 9,484 unique sequences, 6068 had positive hits with the non-redundant protein database; the EST database likely represents approximately 52% of the gene-encoding potential of the monarch genome. The brain transcriptome was cataloged using Gene Ontology and compared to Drosophila. Monarch genes were well represented, including those implicated in behavior. Three genes involved in increased JH activity (allatotropin, juvenile hormone acid methyltransfersase, and takeout were upregulated in summer butterflies, compared to migrants. The locomotion-relevant turtle gene was marginally upregulated in migrants, while the foraging and single-minded genes were not differentially regulated. Many of the genes important for the monarch circadian clock mechanism (involved in sun compass orientation were in the EST resource, including the newly identified cryptochrome 2. The EST database also revealed a novel Na+/K+ ATPase allele predicted to be more resistant to the toxic effects of milkweed than that reported previously. Potential genetic markers were identified from 3,486 EST contigs and included 1599 double-hit single nucleotide polymorphisms (SNPs and 98 microsatellite polymorphisms. These data provide a template of the brain transcriptome for the monarch butterfly. Our "snap-shot" analysis of the differential regulation of candidate genes between summer and migratory butterflies suggests that unbiased, comprehensive

  20. Population Genetics of Overwintering Monarch Butterflies, Danaus plexippus (Linnaeus), from Central Mexico Inferred from Mitochondrial DNA and Microsatellite Markers (United States)

    Pfeiler, Edward; Nazario-Yepiz, Nestor O.; Pérez-Gálvez, Fernan; Chávez-Mora, Cristina Alejandra; Laclette, Mariana Ramírez Loustalot; Rendón-Salinas, Eduardo


    Abstract Population genetic variation and demographic history in Danaus plexippus (L.), from Mexico were assessed based on analyses of mitochondrial cytochrome c oxidase subunit I (COI; 658 bp) and subunit II (COII; 503 bp) gene segments and 7 microsatellite loci. The sample of 133 individuals included both migratory monarchs, mainly from 4 overwintering sites within the Monarch Butterfly Biosphere Reserve (MBBR) in central Mexico (states of Michoacán and México), and a nonmigratory population from Irapuato, Guanajuato. Haplotype (h) and nucleotide (π) diversities were relatively low, averaging 0.466 and 0.00073, respectively, for COI, and 0.629 and 0.00245 for COII. Analysis of molecular variance of the COI data set, which included additional GenBank sequences from a nonmigratory Costa Rican population, showed significant population structure between Mexican migratory monarchs and nonmigratory monarchs from both Mexico and Costa Rica, suggesting limited gene flow between the 2 behaviorally distinct groups. Interestingly, while the COI haplotype frequencies of the nonmigratory populations differed from the migratory, they were similar to each other, despite the great physical distance between them. Microsatellite analyses, however, suggested a lack of structure between the 2 groups, possibly owing to the number of significant deviations from Hardy–Weinberg equilibrium resulting from heterzoygote deficiencies found for most of the loci. Estimates of demographic history of the combined migratory MBBR monarch population, based on the mismatch distribution and Bayesian skyline analyses of the concatenated COI and COII data set (n = 89) suggested a population expansion dating to the late Pleistocene (~35000–40000 years before present) followed by a stable effective female population size (Nef) of about 6 million over the last 10000 years. PMID:28003372

  1. Chasing Migration Genes: A Brain Expressed Sequence Tag Resource for Summer and Migratory Monarch Butterflies (Danaus plexippus) (United States)

    Zhu, Haisun; Casselman, Amy; Reppert, Steven M.


    North American monarch butterflies (Danaus plexippus) undergo a spectacular fall migration. In contrast to summer butterflies, migrants are juvenile hormone (JH) deficient, which leads to reproductive diapause and increased longevity. Migrants also utilize time-compensated sun compass orientation to help them navigate to their overwintering grounds. Here, we describe a brain expressed sequence tag (EST) resource to identify genes involved in migratory behaviors. A brain EST library was constructed from summer and migrating butterflies. Of 9,484 unique sequences, 6068 had positive hits with the non-redundant protein database; the EST database likely represents ∼52% of the gene-encoding potential of the monarch genome. The brain transcriptome was cataloged using Gene Ontology and compared to Drosophila. Monarch genes were well represented, including those implicated in behavior. Three genes involved in increased JH activity (allatotropin, juvenile hormone acid methyltransfersase, and takeout) were upregulated in summer butterflies, compared to migrants. The locomotion-relevant turtle gene was marginally upregulated in migrants, while the foraging and single-minded genes were not differentially regulated. Many of the genes important for the monarch circadian clock mechanism (involved in sun compass orientation) were in the EST resource, including the newly identified cryptochrome 2. The EST database also revealed a novel Na+/K+ ATPase allele predicted to be more resistant to the toxic effects of milkweed than that reported previously. Potential genetic markers were identified from 3,486 EST contigs and included 1599 double-hit single nucleotide polymorphisms (SNPs) and 98 microsatellite polymorphisms. These data provide a template of the brain transcriptome for the monarch butterfly. Our “snap-shot” analysis of the differential regulation of candidate genes between summer and migratory butterflies suggests that unbiased, comprehensive transcriptional profiling

  2. Constraints on Dynamic Triggering from very Short term Microearthquake Aftershocks at Parkfield (United States)

    Ampuero, J.; Rubin, A.


    The study of microearthquakes helps bridge the gap between laboratory experiments and data from large earthquakes, the two disparate scales that have contributed so far to our understanding of earthquake physics. Although they are frequent, microearthquakes are difficult to analyse. Applying high precision relocation techniques, Rubin and Gillard (2000) observed a pronounced asymmetry in the spatial distribution of the earliest and nearest aftershocks of microearthquakes along the San Andreas fault (they occur more often to the NW of the mainshock). It was suggested that this could be related to the velocity contrast across the fault. Preferred directivity of dynamic rupture pulses running along a bimaterial interface (to the SE in the case of the SAF) is expected on theoretical grounds. Our numerical simulations of crack-like rupture on such interfaces show a pronounced asymmetry of the stress histories beyond the rupture ends, and suggest two possible mechanisms for the observed asymmetry: First, that it results from an asymmmetry in the static stress field following arrest of the mainshock (closer to failure to the NW), or second, that it is due to a short-duration tensile pulse that propagates to the SE, which could reduce the number of aftershocks to the SE by dynamic triggering of any nucleation site close enough to failure to have otherwise produced an aftershock. To distinguish betwen these mechanisms we need observations of dynamic triggering in microseismicity. For small events triggered at a distance of some mainshock radii, triggering time scales are so short that seismograms of both events overlap. To detect the occurrence of compound events and very short term aftershocks in the HRSN Parkfield archived waveforms we have developed an automated search algorithm based on empirical Green's function (EGF) deconvolution. Optimal EGFs are first selected by the coherency of the cross-component convolution with respect to the target event. Then Landweber

  3. Joint inversion for Vp, Vs, and Vp/Vs at SAFOD, Parkfield, California (United States)

    Zhang, H.; Thurber, C.; Bedrosian, P.


    We refined the three-dimensional (3-D) Vp, Vs and Vp/Vs models around the San Andreas Fault Observatory at Depth (SAFOD) site using a new double-difference (DD) seismic tomography code (tomoDDPS) that simultaneously solves for earthquake locations and all three velocity models using both absolute and differential P, S, and S-P times. This new method is able to provide a more robust Vp/Vs model than that from the original DD tomography code (tomoDD), obtained simply by dividing Vp by Vs. For the new inversion, waveform cross-correlation times for earthquakes from 2001 to 2002 were also used, in addition to arrival times from earthquakes and explosions in the region. The Vp values extracted from the model along the SAFOD trajectory match well with the borehole log data, providing in situ confirmation of our results. Similar to previous tomographic studies, the 3-D structure around Parkfield is dominated by the velocity contrast across the San Andreas Fault (SAF). In both the Vp and Vs models, there is a clear low-velocity zone as deep as 7 km along the SAF trace, compatible with the findings from fault zone guided waves. There is a high Vp/Vs anomaly zone on the southwest side of the SAF trace that is about 1-2 km wide and extends as deep as 4 km, which is interpreted to be due to fluids and fractures in the package of sedimentary rocks abutting the Salinian basement rock to the southwest. The relocated earthquakes align beneath the northeast edge of this high Vp/Vs zone. We carried out a 2-D correlation analysis for an existing resistivity model and the corresponding profiles through our model, yielding a classification that distinguishes several major lithologies. ?? 2009 by the American Geophysical Union.

  4. Postearthquake relaxation after the 2004 M6 Parkfield, California, earthquake and rate-and-state friction (United States)

    Savage, J.C.; Langbein, J.


    An unusually complete set of measurements (including rapid rate GPS over the first 10 days) of postseismic deformation is available at 12 continuous GPS stations located close to the epicenter of the 2004 M6.0 Parkfield earthquake. The principal component modes for the relaxation of the ensemble of those 12 GPS stations were determined. The first mode alone furnishes an adequate approximation to the data. Thus, the relaxation at all stations can be represented by the product of a common temporal function and distinct amplitudes for each component (north or east) of relaxation at each station. The distribution in space of the amplitudes indicates that the relaxation is dominantly strike slip. The temporal function, which spans times from about 5 min to 900 days postearthquake, can be fit by a superposition of three creep terms, each of the form ??l loge(1 + t/??l), with characteristic times ??, = 4.06, 0.11, and 0.0001 days. It seems likely that what is actually involved is a broad spectrum of characteristic times, the individual components of which arise from afterslip on different fault patches. Perfettini and Avouac (2004) have shown that an individual creep term can be explained by the spring-slider model with rate-dependent (no state variable) friction. The observed temporal function can also be explained using a single spring-slider model (i.e., single fault patch) that includes rate-and-state-dependent friction, a single-state variable, and either of the two commonly used (aging and slip) state evolution laws. In the latter fits, the rate-and-state friction parameter b is negative.

  5. Preliminary analysis of strong-motion recordings from the 28 September 2004 Parkfield, California earthquake (United States)

    Shakal, A.; Graizer, V.; Huang, M.; Borcherdt, R.; Haddadi, H.; Lin, K.-W.; Stephens, C.; Roffers, P.


    The Parkfield 2004 earthquake yielded the most extensive set of strong-motion data in the near-source region of a magnitude 6 earthquake yet obtained. The recordings of acceleration and volumetric strain provide an unprecedented document of the near-source seismic radiation for a moderate earthquake. The spatial density of the measurements alon g the fault zone and in the linear arrays perpendicular to the fault is expected to provide an exceptional opportunity to develop improved models of the rupture process. The closely spaced measurements should help infer the temporal and spatial distribution of the rupture process at much higher resolution than previously possible. Preliminary analyses of the peak a cceleration data presented herein shows that the motions vary significantly along the rupture zone, from 0.13 g to more than 2.5 g, with a map of the values showing that the larger values are concentrated in three areas. Particle motions at the near-fault stations are consistent with bilateral rupture. Fault-normal pulses similar to those observed in recent strike-slip earthquakes are apparent at several of the stations. The attenuation of peak ground acceleration with distance is more rapid than that indicated by some standard relationships but adequately fits others. Evidence for directivity in the peak acceleration data is not strong. Several stations very near, or over, the rupturing fault recorded relatively low accelerations. These recordings may provide a quantitative basis to understand observations of low near-fault shaking damage that has been reported in other large strike-slip earthquak.

  6. Tremor reveals stress shadowing, deep postseismic creep, and depth-dependent slip recurrence on the lower-crustal San Andreas fault near Parkfield (United States)

    Shelly, David R.; Johnson, Kaj M.


    The 2003 magnitude 6.5 San Simeon and the 2004 magnitude 6.0 Parkfield earthquakes induced small, but significant, static stress changes in the lower crust on the central San Andreas fault, where recently detected tectonic tremor sources provide new constraints on deep fault creep processes. We find that these earthquakes affect tremor rates very differently, consistent with their differing transferred static shear stresses. The San Simeon event appears to have cast a "stress shadow" north of Parkfield, where tremor activity was stifled for 3-6 weeks. In contrast, the 2004 Parkfield earthquake dramatically increased tremor activity rates both north and south of Parkfield, allowing us to track deep postseismic slip. Following this event, rates initially increased by up to two orders of magnitude for the relatively shallow tremor sources closest to the rupture, with activity in some sources persisting above background rates for more than a year. We also observe strong depth dependence in tremor recurrence patterns, with shallower sources generally exhibiting larger, less-frequent bursts, possibly signaling a transition toward steady creep with increasing temperature and depth. Copyright 2011 by the American Geophysical Union.

  7. Seismicity Precursors of the M6.0 2004 Parkfield and M7.0 1989Loma Prieta Earthquakes

    Energy Technology Data Exchange (ETDEWEB)

    Korneev, Valeri A.


    The M6.0 2004 Parkfield and M7.0 1989 Loma Prietastrike-slip earthquakes on the San Andreas Fault (SAF) were preceded byseismicity peaks occurring several months prior to the main events.Earthquakes directly within the SAF zone were intentionally excluded fromthe analysis because they manifest stress-release processes rather thanstress accumulation. The observed increase in seismicity is interpretedas a signature of the increasing stress level in the surrounding crust,whereas the peaks and the subsequent decrease in seismicity areattributed to damage-induced softening processes. Furthermore, in bothcases there is a distinctive zone of low seismic activity that surroundsthe epicentral region in the pre-event period. The increase of seismicityin the crust surrounding a potential future event and the development ofa low-seismicity epicentral zone can be regarded as promising precursoryinformation that could help signal the arrival of large earthquakes. TheGutenberg-Richter relationship (GRR) should allow extrapolation ofseismicity changes down to seismic noise level magnitudes. Thishypothesis is verified by comparison of seismic noise at 80 Hz with theParkfield M4 1993-1994 series, where noise peaks 5 months before theseries to about twice the background level.

  8. 3-D P- and S-wave velocity structure and low-frequency earthquake locations in the Parkfield, California region (United States)

    Zeng, Xiangfang; Thurber, Clifford H.; Shelly, David R.; Harrington, Rebecca M.; Cochran, Elizabeth S.; Bennington, Ninfa L.; Peterson, Dana; Guo, Bin; McClement, Kara


    To refine the 3-D seismic velocity model in the greater Parkfield, California region, a new data set including regular earthquakes, shots, quarry blasts and low-frequency earthquakes (LFEs) was assembled. Hundreds of traces of each LFE family at two temporary arrays were stacked with time–frequency domain phase weighted stacking method to improve signal-to-noise ratio. We extend our model resolution to lower crustal depth with LFE data. Our result images not only previously identified features but also low velocity zones (LVZs) in the area around the LFEs and the lower crust beneath the southern Rinconada Fault. The former LVZ is consistent with high fluid pressure that can account for several aspects of LFE behaviour. The latter LVZ is consistent with a high conductivity zone in magnetotelluric studies. A new Vs model was developed with S picks that were obtained with a new autopicker. At shallow depth, the low Vs areas underlie the strongest shaking areas in the 2004 Parkfield earthquake. We relocate LFE families and analyse the location uncertainties with the NonLinLoc and tomoDD codes. The two methods yield similar results.

  9. Aftershock distribution as a constraint on the geodetic model of coseismic slip for the 2004 Parkfield earthquake (United States)

    Bennington, Ninfa; Thurber, Clifford; Feigl, Kurt; ,


    Several studies of the 2004 Parkfield earthquake have linked the spatial distribution of the event’s aftershocks to the mainshock slip distribution on the fault. Using geodetic data, we find a model of coseismic slip for the 2004 Parkfield earthquake with the constraint that the edges of coseismic slip patches align with aftershocks. The constraint is applied by encouraging the curvature of coseismic slip in each model cell to be equal to the negative of the curvature of seismicity density. The large patch of peak slip about 15 km northwest of the 2004 hypocenter found in the curvature-constrained model is in good agreement in location and amplitude with previous geodetic studies and the majority of strong motion studies. The curvature-constrained solution shows slip primarily between aftershock “streaks” with the continuation of moderate levels of slip to the southeast. These observations are in good agreement with strong motion studies, but inconsistent with the majority of published geodetic slip models. Southeast of the 2004 hypocenter, a patch of peak slip observed in strong motion studies is absent from our curvature-constrained model, but the available GPS data do not resolve slip in this region. We conclude that the geodetic slip model constrained by the aftershock distribution fits the geodetic data quite well and that inconsistencies between models derived from seismic and geodetic data can be attributed largely to resolution issues.

  10. Dispersal of the monarch butterfly (Danaus plexippus over southern Spain from its breeding grounds

    Directory of Open Access Journals (Sweden)

    Obregón, R.


    Full Text Available From 2000–2016, monarch butterflies were detected at 127 locations away from their usual coastal breeding areas in the south of the Iberian peninsula. These findings were recorded in the summer–autumn period, coinciding with the highest abundance of individuals and the highest proportion of patches occupied in their reproduction areas near the Strait of Gibraltar. These dispersing individuals have no chance of successfully establishing new colonies at these sites because the food plants for egg laying do not grow in the localities where they were detected. However, these dispersive movements could be the source of their successful colonisation on food plants growing in other areas of the Iberian peninsula and in other Mediterranean countries.

  11. The incorporation of fault zone head wave and direct wave secondary arrival times and arrival polarizations into seismic tomography: Application to the Parkfield, California area (United States)

    Bennington, N. L.; Thurber, C. H.; Peng, Z.; Zhao, P.


    We present a 3D P-wave velocity (Vp) model of the Parkfield region that utilizes existing P-wave arrival time data, including fault zone head waves (FZHW), plus new data from direct wave secondary arrivals (DWSA). The first-arrival and DWSA travel times are obtained as the global and local minimum travel time paths, respectively. The inclusion of DWSA results in as much as a 10% increase in the across-fault velocity contrast for the Vp model at Parkfield relative to Thurber et al. (2006). Viewed along strike, three pronounced velocity contrast regions are observed: a pair of strong positive velocity contrasts (SW fast), one NW of the 1966 Parkfield hypocenter and the other SE of the 2004 Parkfield hypocenter, and a strong negative velocity contrast (NE fast) between the two hypocenters. The negative velocity contrast partially to entirely encompasses peak coseismic slip estimated in several slip models for the 2004 earthquake, suggesting that the negative velocity contrast played a part in defining the rupture patch of the 2004 Parkfield earthquake. We expand on this work by modifying our seismic tomography algorithm to incorporate arrival polarizations (azimuths). Synthetic tests will be presented to demonstrate the improvements in velocity structure when arrival polarizations are incorporated. These tests will compare the synthetic model recovered when FZHW/DWSA arrivals as well as existing P-wave arrival time data are inverted to that recovered with the same dataset with the inclusion of arrival polarizations. We plan to extend this work to carry out a full scale seismic tomography/relocation inversion at Parkfield, CA utilizing arrival polarizations from all first-P arrivals, and FZHW/DWSA arrivals as well as existing P-wave arrival time data. This effort requires the determination of polarization data for all P-waves and FZHW's at Parkfield. To this end, we use changes in the arrival azimuth from fault normal to source-receiver direction to identify FZHW and

  12. Experimental examination of intraspecific density-dependent competition during the breeding period in monarch butterflies (Danaus plexippus.

    Directory of Open Access Journals (Sweden)

    D T Tyler Flockhart

    Full Text Available A central goal of population ecology is to identify the factors that regulate population growth. Monarch butterflies (Danaus plexippus in eastern North America re-colonize the breeding range over several generations that result in population densities that vary across space and time during the breeding season. We used laboratory experiments to measure the strength of density-dependent intraspecific competition on egg laying rate and larval survival and then applied our results to density estimates of wild monarch populations to model the strength of density dependence during the breeding season. Egg laying rates did not change with density but larvae at high densities were smaller, had lower survival, and weighed less as adults compared to lower densities. Using mean larval densities from field surveys resulted in conservative estimates of density-dependent population reduction that varied between breeding regions and different phases of the breeding season. Our results suggest the highest levels of population reduction due to density-dependent intraspecific competition occur early in the breeding season in the southern portion of the breeding range. However, we also found that the strength of density dependence could be almost five times higher depending on how many life-stages were used as part of field estimates. Our study is the first to link experimental results of a density-dependent reduction in vital rates to observed monarch densities in the wild and show that the effects of density dependent competition in monarchs varies across space and time, providing valuable information for developing robust, year-round population models in this migratory organism.

  13. Acoustic Emission Precursors of M6.0 2004 Parkfield and M7.0 1989Loma Prieta Earthquakes

    Energy Technology Data Exchange (ETDEWEB)

    Korneev, Valeri


    Two recent strike-slip earthquakes on the San Andreas Fault(SAF) in California, the M6.0 2004 Parkfield and M7.0 1989 Loma Prietaevents, revealed peaks in the acoustic emission (AE) activity in thesurrounding crust several months prior to the main events. Earthquakesdirectly within the SAF zone were intentionally excluded from theanalysis. The observed increase in AE is assumed to be a signature of theincreasing stress level in the surrounding crust, while the peak andsubsequent decrease in AE starting several months prior to the mainevents is attributed to damage-induced softening processes as discussedherein. Further, distinctive zones of low seismic activity surroundingthe epicentral regions in the pre-event time period are present for thetwo studied events. Both AE increases in the crust surrounding apotential future event and the development of a low-seismicity epicentralzone can be regarded as promising precursory information that could helpsignal the arrival of large earthquakes.

  14. The Monarch Initiative: an integrative data and analytic platform connecting phenotypes to genotypes across species

    International Nuclear Information System (INIS)

    Mungall, Christopher J.; McMurry, Julie A.; Köhler, Sebastian; Balhoff, James P.; Borromeo, Charles


    The correlation of phenotypic outcomes with genetic variation and environmental factors is a core pursuit in biology and biomedicine. Numerous challenges impede our progress: patient phenotypes may not match known diseases, candidate variants may be in genes that have not been characterized, model organisms may not recapitulate human or veterinary diseases, filling evolutionary gaps is difficult, and many resources must be queried to find potentially significant genotype-phenotype associations. Nonhuman organisms have proven instrumental in revealing biological mechanisms. Advanced informatics tools can identify phenotypically relevant disease models in research and diagnostic contexts. Large-scale integration of model organism and clinical research data can provide a breadth of knowledge not available from individual sources and can provide contextualization of data back to these sources. The Monarch Initiative ( is a collaborative, open science effort that aims to semantically integrate genotype-phenotype data from many species and sources in order to support precision medicine, disease modeling, and mechanistic exploration. Our integrated knowledge graph, analytic tools, and web services enable diverse users to explore relationships between phenotypes and genotypes across species.

  15. Classification of osteoporosis by artificial neural network based on monarch butterfly optimisation algorithm. (United States)

    Devikanniga, D; Joshua Samuel Raj, R


    Osteoporosis is a life threatening disease which commonly affects women mostly after their menopause. It primarily causes mild bone fractures, which on advanced stage leads to the death of an individual. The diagnosis of osteoporosis is done based on bone mineral density (BMD) values obtained through various clinical methods experimented from various skeletal regions. The main objective of the authors' work is to develop a hybrid classifier model that discriminates the osteoporotic patient from healthy person, based on BMD values. In this Letter, the authors propose the monarch butterfly optimisation-based artificial neural network classifier which helps in earlier diagnosis and prevention of osteoporosis. The experiments were conducted using 10-fold cross-validation method for two datasets lumbar spine and femoral neck. The results were compared with other similar hybrid approaches. The proposed method resulted with the accuracy, specificity and sensitivity of 97.9% ± 0.14, 98.33% ± 0.03 and 95.24% ± 0.08, respectively, for lumbar spine dataset and 99.3% ± 0.16%, 99.2% ± 0.13 and 100, respectively, for femoral neck dataset. Further, its performance is compared using receiver operating characteristics analysis and Wilcoxon signed-rank test. The results proved that the proposed classifier is efficient and it outperformed the other approaches in all the cases.

  16. The sonority of the daily life of the Castilian cities in times of the Catholic Monarchs

    Directory of Open Access Journals (Sweden)

    Gisela Beatriz Coronado Schwindt


    Full Text Available Throughout history, societies have experienced their everyday lives through sensory models built by them, determining a field of possibilities of the visible and the invisible, the tactile and non-tactile, olfactory and odorless, the taste and the insipid thing. The senses, in addition to be a means of perception of physical experiences, can be conceptualised as social phenomena and historical formations since their meanings are modified over time. Actively involved in the social construction of a culture due to sensory perceptions include, while at the same time define, the areas in which the economic and political activities, and social practices are developed. Different sounds of human beings, issued by themselves or caused by words, deeds, gestures, etc., tell us about their attitudes, practices and conflicts within the framework of their social reality. Gathered in a time and space they form a specific soundscape plausible to analyze in their social and historical significance. Through these pages, we propose to understand the intervention that exercises various sounds in the social configuration of the Castilian cities during the reign of the Catholic Monarchs. This analysis is carried out through the narration of daily life in late medieval and early modern times, based on different written sources of the period. The exercise we set ourselves is to reread the documentation available to the historian from a cultural and sensory perspective.

  17. Regional climate on the breeding grounds predicts variation in the natal origin of monarch butterflies overwintering in Mexico over 38 years. (United States)

    Flockhart, D T Tyler; Brower, Lincoln P; Ramirez, M Isabel; Hobson, Keith A; Wassenaar, Leonard I; Altizer, Sonia; Norris, D Ryan


    Addressing population declines of migratory insects requires linking populations across different portions of the annual cycle and understanding the effects of variation in weather and climate on productivity, recruitment, and patterns of long-distance movement. We used stable H and C isotopes and geospatial modeling to estimate the natal origin of monarch butterflies (Danaus plexippus) in eastern North America using over 1000 monarchs collected over almost four decades at Mexican overwintering colonies. Multinomial regression was used to ascertain which climate-related factors best-predicted temporal variation in natal origin across six breeding regions. The region producing the largest proportion of overwintering monarchs was the US Midwest (mean annual proportion = 0.38; 95% CI: 0.36-0.41) followed by the north-central (0.17; 0.14-0.18), northeast (0.15; 0.11-0.16), northwest (0.12; 0.12-0.16), southwest (0.11; 0.08-0.12), and southeast (0.08; 0.07-0.11) regions. There was no evidence of directional shifts in the relative contributions of different natal regions over time, which suggests these regions are comprising the same relative proportion of the overwintering population in recent years as in the mid-1970s. Instead, interannual variation in the proportion of monarchs from each region covaried with climate, as measured by the Southern Oscillation Index and regional-specific daily maximum temperature and precipitation, which together likely dictate larval development rates and food plant condition. Our results provide the first robust long-term analysis of predictors of the natal origins of monarchs overwintering in Mexico. Conservation efforts on the breeding grounds focused on the Midwest region will likely have the greatest benefit to eastern North American migratory monarchs, but the population will likely remain sensitive to regional and stochastic weather patterns. © 2017 John Wiley & Sons Ltd.

  18. Trends in deforestation and forest degradation after a decade of monitoring in the Monarch Butterfly Biosphere Reserve in Mexico. (United States)

    Vidal, Omar; López-García, José; Rendón-Salinas, Eduardo


    We used aerial photographs, satellite images, and field surveys to monitor forest cover in the core zones of the Monarch Butterfly Biosphere Reserve in Mexico from 2001 to 2012. We used our data to assess the effectiveness of conservation actions that involved local, state, and federal authorities and community members (e.g., local landowners and private and civil organizations) in one of the world's most iconic protected areas. From 2001 through 2012, 1254 ha were deforested (i.e., cleared areas had <10% canopy cover), 925 ha were degraded (i.e., areas for which canopy forest decreased), and 122 ha were affected by climatic conditions. Of the total 2179 ha of affected area, 2057 ha were affected by illegal logging: 1503 ha by large-scale logging and 554 ha by small-scale logging. Mexican authorities effectively enforced efforts to protect the monarch reserve, particularly from 2007 to 2012. Those efforts, together with the decade-long financial support from Mexican and international philanthropists and businesses to create local alternative-income generation and employment, resulted in the decrease of large-scale illegal logging from 731 ha affected in 2005-2007 to none affected in 2012, although small-scale logging is of growing concern. However, dire regional social and economic problems remain, and they must be addressed to ensure the reserve's long-term conservation. The monarch butterfly (Danaus plexippus) overwintering colonies in Mexico-which engage in one of the longest known insect migrations-are threatened by deforestation, and a multistakeholder, regional, sustainable-development strategy is needed to protect the reserve. © 2013 Society for Conservation Biology.

  19. Volcanic ash modeling with the NMMB-MONARCH-ASH model: quantification of offline modeling errors (United States)

    Marti, Alejandro; Folch, Arnau


    Volcanic ash modeling systems are used to simulate the atmospheric dispersion of volcanic ash and to generate forecasts that quantify the impacts from volcanic eruptions on infrastructures, air quality, aviation, and climate. The efficiency of response and mitigation actions is directly associated with the accuracy of the volcanic ash cloud detection and modeling systems. Operational forecasts build on offline coupled modeling systems in which meteorological variables are updated at the specified coupling intervals. Despite the concerns from other communities regarding the accuracy of this strategy, the quantification of the systematic errors and shortcomings associated with the offline modeling systems has received no attention. This paper employs the NMMB-MONARCH-ASH model to quantify these errors by employing different quantitative and categorical evaluation scores. The skills of the offline coupling strategy are compared against those from an online forecast considered to be the best estimate of the true outcome. Case studies are considered for a synthetic eruption with constant eruption source parameters and for two historical events, which suitably illustrate the severe aviation disruptive effects of European (2010 Eyjafjallajökull) and South American (2011 Cordón Caulle) volcanic eruptions. Evaluation scores indicate that systematic errors due to the offline modeling are of the same order of magnitude as those associated with the source term uncertainties. In particular, traditional offline forecasts employed in operational model setups can result in significant uncertainties, failing to reproduce, in the worst cases, up to 45-70 % of the ash cloud of an online forecast. These inconsistencies are anticipated to be even more relevant in scenarios in which the meteorological conditions change rapidly in time. The outcome of this paper encourages operational groups responsible for real-time advisories for aviation to consider employing computationally

  20. Rural aquaculture as a sustainable alternative for forest conservation in the Monarch Butterfly Biosphere Reserve, Mexico. (United States)

    López-García, José; Manzo-Delgado, Lilia L; Alcántara-Ayala, Irasema


    Forest conservation plays a significant role in environmental sustainability. In Mexico only 8.48 million ha of forest are used for conservation of biodiversity. Payment for Environmental Services in the Monarch Butterfly Biosphere Reserve, one of the most important national protected areas, contributes to the conservation of these forests. In the Reserve, production of rainbow trout has been important for the rural communities who need to conserve the forest cover in order to maintain the hibernation cycle of the butterfly. Aquaculture is a highly productive activity for these protected areas, since it harnesses the existing water resources. In this study, changes from 1999 to 2012 in vegetation and land-use cover in the El Lindero basin within the Reserve were evaluated in order to determine the conservation status and to consider the feasibility of aquaculture as a means of sustainable development at community level. Evaluation involved stereoscopic interpretation of digital aerial photographs from 1999 to 2012 at 1:10,000 scale, comparative analysis by orthocorrected mosaics and restitution on the mosaics. Between 1999 and 2012, forested land recovered by 28.57 ha (2.70%) at the expense of non-forested areas, although forest degradation was 3.59%. Forest density increased by 16.87%. In the 46 ha outside the Reserve, deforestation spread by 0.26%, and land use change was 0.11%. The trend towards change in forest cover is closely related to conservation programmes, particularly payment for not extracting timber, reforestation campaigns and surveillance, whose effects have been exploited for the development of rural aquaculture; this is a new way to improve the socio-economic status of the population, to avoid logging and to achieve environmental sustainability in the Reserve. Copyright © 2014 Elsevier Ltd. All rights reserved.

  1. Tidal triggering of low frequency earthquakes near Parkfield, California: Implications for fault mechanics within the brittle-ductile transition (United States)

    Thomas, A.M.; Burgmann, R.; Shelly, David R.; Beeler, Nicholas M.; Rudolph, M.L.


    Studies of nonvolcanic tremor (NVT) have established the significant impact of small stress perturbations on NVT generation. Here we analyze the influence of the solid earth and ocean tides on a catalog of ∼550,000 low frequency earthquakes (LFEs) distributed along a 150 km section of the San Andreas Fault centered at Parkfield. LFE families are identified in the NVT data on the basis of waveform similarity and are thought to represent small, effectively co-located earthquakes occurring on brittle asperities on an otherwise aseismic fault at depths of 16 to 30 km. We calculate the sensitivity of each of these 88 LFE families to the tidally induced right-lateral shear stress (RLSS), fault-normal stress (FNS), and their time derivatives and use the hypocentral locations of each family to map the spatial variability of this sensitivity. LFE occurrence is most strongly modulated by fluctuations in shear stress, with the majority of families demonstrating a correlation with RLSS at the 99% confidence level or above. Producing the observed LFE rate modulation in response to shear stress perturbations requires low effective stress in the LFE source region. There are substantial lateral and vertical variations in tidal shear stress sensitivity, which we interpret to reflect spatial variation in source region properties, such as friction and pore fluid pressure. Additionally, we find that highly episodic, shallow LFE families are generally less correlated with tidal stresses than their deeper, continuously active counterparts. The majority of families have weaker or insignificant correlation with positive (tensile) FNS. Two groups of families demonstrate a stronger correlation with fault-normal tension to the north and with compression to the south of Parkfield. The families that correlate with fault-normal clamping coincide with a releasing right bend in the surface fault trace and the LFE locations, suggesting that the San Andreas remains localized and contiguous down

  2. Improving attenuation tomography by novel inversions for t* and Q: application to Parkfield, California and Okmok volcano, Alaska (United States)

    Pesicek, J. D.; Bennington, N. L.; Thurber, C. H.; Zhang, H.


    Standard methods for mapping variations in seismic attenuation (Q) structure using local earthquake data involve a two-step process. First, values of the whole path attenuation operator t* are determined from earthquake data recorded on a seismic array by inverting observed spectra for source and attenuation parameters. Then, these t* data are used to invert tomographically for frequency-independent Q models. The observed earthquake amplitude spectra depend on both source parameters and site effects. However, quantification of site effects is often neglected. Bennington et al. [2008] determined site response jointly with source parameters for small groups of events and then computed each station's site response as the average over all groups. Building on this work, we have adapted the method to model all events simultaneously to more accurately determine site response from the earthquake spectra. This in turn allows us to more accurately determine t*. However, resolution analysis of the results shows that some parameters are not well resolved in the joint inversion. Thus, an alternating inversion scheme is tested and adapted to alleviate poor resolution and parameter trade-offs. The new scheme produces better fits to the earthquake spectra than previous methods, and the resulting t* data should allow for more accurate determination of the Q structure. Because the equation relating t* to Q is nonlinear, the typical approach to determining Q is to solve for changes to a starting model iteratively, similar to methods commonly used in travel time tomography. However, if we instead solve for the inverse of Q, the equation becomes linear and the solution no longer depends on the starting model. This simple modification may allow us to more accurately determine Q. We test these new t* and Q methods using earthquake data from Parkfield, California and Okmok volcano, Alaska. We present the results for real and synthetic data and compare and contrast these results to more

  3. Attenuation of the jasmonate burst, plant defensive traits, and resistance to specialist monarch caterpillars on shaded common milkweed (Asclepias syriaca). (United States)

    Agrawal, Anurag A; Kearney, Emily E; Hastings, Amy P; Ramsey, Trey E


    Plant responses to herbivory and light competition are often in opposing directions, posing a potential conflict for plants experiencing both stresses. For sun-adapted species, growing in shade typically makes plants more constitutively susceptible to herbivores via reduced structural and chemical resistance traits. Nonetheless, the impact of light environment on induced resistance has been less well-studied, especially in field experiments that link physiological mechanisms to ecological outcomes. Accordingly, we studied induced resistance of common milkweed (Asclepias syriaca, a sun-adapted plant), and linked hormonal responses, resistance traits, and performance of specialist monarch caterpillars (Danaus plexippus) in varying light environments. In natural populations, plants growing under forest-edge shade showed reduced levels of resistance traits (lower leaf toughness, cardenolides, and trichomes) and enhanced light-capture traits (higher specific leaf area, larger leaves, and lower carbon-to-nitrogen ratio) compared to paired plants in full sun. In a field experiment repeated over two years, only milkweeds growing in full sun exhibited induced resistance to monarchs, whereas plants growing in shade were constitutively more susceptible and did not induce resistance. In a more controlled field experiment, plant hormones were higher in the sun (jasmonic acid, salicylic acid, abscisic acid, indole acidic acid) and were induced by herbivory (jasmonic acid and abscisic acid). In particular, the jasmonate burst following herbivory was halved in plants raised in shaded habitats, and this correspondingly reduced latex induction (but not cardenolide induction). Thus, we provide a mechanistic basis for the attenuation of induced plant resistance in low resource environments. Additionally, there appears to be specificity in these interactions, with light-mediated impacts on jasmonate-induction being stronger for latex exudation than cardenolides.

  4. Vertebrate-like CRYPTOCHROME 2 from monarch regulates circadian transcription via independent repression of CLOCK and BMAL1 activity. (United States)

    Zhang, Ying; Markert, Matthew J; Groves, Shayna C; Hardin, Paul E; Merlin, Christine


    Circadian repression of CLOCK-BMAL1 by PERIOD and CRYPTOCHROME (CRY) in mammals lies at the core of the circadian timekeeping mechanism. CRY repression of CLOCK-BMAL1 and regulation of circadian period are proposed to rely primarily on competition for binding with coactivators on an α-helix located within the transactivation domain (TAD) of the BMAL1 C terminus. This model has, however, not been tested in vivo. Here, we applied CRISPR/Cas9-mediated mutagenesis in the monarch butterfly ( Danaus plexippus ), which possesses a vertebrate-like CRY (dpCRY2) and an ortholog of BMAL1, to show that insect CRY2 regulates circadian repression through TAD α-helix-dependent and -independent mechanisms. Monarch mutants lacking the BMAL1 C terminus including the TAD exhibited arrhythmic eclosion behavior. In contrast, mutants lacking the TAD α-helix but retaining the most distal C-terminal residues exhibited robust rhythms during the first day of constant darkness (DD1), albeit with a delayed peak of eclosion. Phase delay in this mutant on DD1 was exacerbated in the presence of a single functional allele of dpCry2 , and rhythmicity was abolished in the absence of dpCRY2. Reporter assays in Drosophila S2 cells further revealed that dpCRY2 represses through two distinct mechanisms: a TAD-dependent mechanism that involves the dpBMAL1 TAD α-helix and dpCLK W328 and a TAD-independent mechanism involving dpCLK E333. Together, our results provide evidence for independent mechanisms of vertebrate-like CRY circadian regulation on the BMAL1 C terminus and the CLK PAS-B domain and demonstrate the importance of a BMAL1 TAD-independent mechanism for generating circadian rhythms in vivo.

  5. Automatic identification of fault zone head waves and direct P waves and its application in the Parkfield section of the San Andreas Fault, California (United States)

    Li, Zefeng; Peng, Zhigang


    Fault zone head waves (FZHWs) are observed along major strike-slip faults and can provide high-resolution imaging of fault interface properties at seismogenic depth. In this paper, we present a new method to automatically detect FZHWs and pick direct P waves secondary arrivals (DWSAs). The algorithm identifies FZHWs by computing the amplitude ratios between the potential FZHWs and DSWAs. The polarities, polarizations and characteristic periods of FZHWs and DSWAs are then used to refine the picks or evaluate the pick quality. We apply the method to the Parkfield section of the San Andreas Fault where FZHWs have been identified before by manual picks. We compare results from automatically and manually picked arrivals and find general agreement between them. The obtained velocity contrast at Parkfield is generally 5-10 per cent near Middle Mountain while it decreases below 5 per cent near Gold Hill. We also find many FZHWs recorded by the stations within 1 km of the background seismicity (i.e. the Southwest Fracture Zone) that have not been reported before. These FZHWs could be generated within a relatively wide low velocity zone sandwiched between the fast Salinian block on the southwest side and the slow Franciscan Mélange on the northeast side. Station FROB on the southwest (fast) side also recorded a small portion of weak precursory signals before sharp P waves. However, the polarities of weak signals are consistent with the right-lateral strike-slip mechanisms, suggesting that they are unlikely genuine FZHW signals.

  6. Does skipping a meal matter to a butterfly's appearance? Effects of larval food stress on wing morphology and color in monarch butterflies.

    Directory of Open Access Journals (Sweden)

    Haley Johnson

    Full Text Available In animals with complex life cycles, all resources needed to form adult tissues are procured at the larval stage. For butterflies, the proper development of wings involves synthesizing tissue during metamorphosis based on the raw materials obtained by larvae. Similarly, manufacture of pigment for wing scales also requires resources acquired by larvae. We conducted an experiment to test the effects of food deprivation in the larval stage on multiple measures of adult wing morphology and coloration of monarch butterflies (Danaus plexippus, a species in which long-distance migration makes flight efficiency critical. In a captive setting, we restricted food (milkweed from late-stage larvae for either 24 hrs or 48 hrs, then after metamorphosis we used image analysis methods to measure forewing surface area and elongation (length/width, which are both important for migration. We also measured the brightness of orange pigment and the intensity of black on the wing. There were correlations between several wing features, including an unexpected association between wing elongation and melanism, which will require further study to fully understand. The clearest effect of food restriction was a reduction in adult wing size in the high stress group (by approximately 2%. Patterns observed for other wing traits were ambiguous: monarchs in the low stress group (but not the high had less elongated and paler orange pigmentation. There was no effect on wing melanism. Although some patterns obtained in this study were unclear, our results concerning wing size have direct bearing on the monarch migration. We show that if milkweed is limited for monarch larvae, their wings become stunted, which could ultimately result in lower migration success.

  7. Milkweed (Gentianales: Apocynaceae): a farmscape resource for increasing parasitism of stink bugs (Hemiptera: Pentatomidae) and providing nectar to insect pollinators and monarch butterflies. (United States)

    Tillman, P G; Carpenter, J E


    In peanut-cotton farmscapes in Georgia, the stink bugs Nezara viridula (L.) and Chinavia hilaris (Say) (Hemiptera: Pentatomidae) and the leaffooted bug, Leptoglossus phyllopus (L.) (Hemiptera: Coreidae), disperse at crop-to-crop interfaces to feed on bolls in cotton. The main objective of this study was to determine whether insecticide-free tropical milkweed (Asclepias curassavica L.), a nectar-producing plant, can increase parasitism of these bugs by Trichopoda pennipes (F.) (Diptera: Tachinidae) and provide nectar to monarch butterflies and insect pollinators in these farmscapes. Peanut-cotton plots with and without flowering milkweed plants were established in 2009 and 2010. Adult T. pennipes, monarch butterflies, honey bees, and native insect pollinators readily fed on floral nectar of milkweed. Monarch larvae feeding on milkweed vegetation successfully developed into pupae. In 2009, N. viridula was the primary host of T. pennipes in cotton, and parasitism of this pest by the parasitoid was significantly higher in milkweed cotton (61.6%) than in control cotton (13.3%). In 2010, parasitism of N. viridula, C. hilaris, and L. phyllopus by T. pennipes was significantly higher in milkweed cotton (24.0%) than in control cotton (1.1%). For both years of the study, these treatment differences were not owing to a response by the parasitoid to differences in host density, because density of hosts was not significantly different between treatments. In conclusion, incorporation of milkweed in peanut-cotton plots increased stink bug parasitism in cotton and provided nectar to insect pollinators and monarch butterflies.

  8. Does Skipping a Meal Matter to a Butterfly's Appearance? Effects of Larval Food Stress on Wing Morphology and Color in Monarch Butterflies (United States)

    Johnson, Haley; Solensky, Michelle J.; Satterfield, Dara A.; Davis, Andrew K.


    In animals with complex life cycles, all resources needed to form adult tissues are procured at the larval stage. For butterflies, the proper development of wings involves synthesizing tissue during metamorphosis based on the raw materials obtained by larvae. Similarly, manufacture of pigment for wing scales also requires resources acquired by larvae. We conducted an experiment to test the effects of food deprivation in the larval stage on multiple measures of adult wing morphology and coloration of monarch butterflies (Danaus plexippus), a species in which long-distance migration makes flight efficiency critical. In a captive setting, we restricted food (milkweed) from late-stage larvae for either 24 hrs or 48 hrs, then after metamorphosis we used image analysis methods to measure forewing surface area and elongation (length/width), which are both important for migration. We also measured the brightness of orange pigment and the intensity of black on the wing. There were correlations between several wing features, including an unexpected association between wing elongation and melanism, which will require further study to fully understand. The clearest effect of food restriction was a reduction in adult wing size in the high stress group (by approximately 2%). Patterns observed for other wing traits were ambiguous: monarchs in the low stress group (but not the high) had less elongated and paler orange pigmentation. There was no effect on wing melanism. Although some patterns obtained in this study were unclear, our results concerning wing size have direct bearing on the monarch migration. We show that if milkweed is limited for monarch larvae, their wings become stunted, which could ultimately result in lower migration success. PMID:24695643

  9. Does skipping a meal matter to a butterfly's appearance? Effects of larval food stress on wing morphology and color in monarch butterflies. (United States)

    Johnson, Haley; Solensky, Michelle J; Satterfield, Dara A; Davis, Andrew K


    In animals with complex life cycles, all resources needed to form adult tissues are procured at the larval stage. For butterflies, the proper development of wings involves synthesizing tissue during metamorphosis based on the raw materials obtained by larvae. Similarly, manufacture of pigment for wing scales also requires resources acquired by larvae. We conducted an experiment to test the effects of food deprivation in the larval stage on multiple measures of adult wing morphology and coloration of monarch butterflies (Danaus plexippus), a species in which long-distance migration makes flight efficiency critical. In a captive setting, we restricted food (milkweed) from late-stage larvae for either 24 hrs or 48 hrs, then after metamorphosis we used image analysis methods to measure forewing surface area and elongation (length/width), which are both important for migration. We also measured the brightness of orange pigment and the intensity of black on the wing. There were correlations between several wing features, including an unexpected association between wing elongation and melanism, which will require further study to fully understand. The clearest effect of food restriction was a reduction in adult wing size in the high stress group (by approximately 2%). Patterns observed for other wing traits were ambiguous: monarchs in the low stress group (but not the high) had less elongated and paler orange pigmentation. There was no effect on wing melanism. Although some patterns obtained in this study were unclear, our results concerning wing size have direct bearing on the monarch migration. We show that if milkweed is limited for monarch larvae, their wings become stunted, which could ultimately result in lower migration success.

  10. Evaluation of hypotheses for right-lateral displacement of Neogene strata along the San Andreas Fault between Parkfield and Maricopa, California (United States)

    Stanley, Richard G.; Barron, John A.; Powell, Charles L.


    We used geological field studies and diatom biostratigraphy to test a published hypothesis that Neogene marine siliceous strata in the Maricopa and Parkfield areas, located on opposite sides of the San Andreas Fault, were formerly contiguous and then were displaced by about 80–130 kilometers (km) of right-lateral slip along the fault. In the Maricopa area on the northeast side of the San Andreas Fault, the upper Miocene Bitterwater Creek Shale consists of hard, siliceous shale with dolomitic concretions and turbidite sandstone interbeds. Diatom assemblages indicate that the Bitterwater Creek Shale was deposited about 8.0–6.7 million years before present (Ma) at the same time as the uppermost part of the Monterey Formation in parts of coastal California. In the Parkfield area on the southwest side of the San Andreas Fault, the upper Miocene Pancho Rico Formation consists of soft to indurated mudstone and siltstone and fossiliferous, bioturbated sandstone. Diatom assemblages from the Pancho Rico indicate deposition about 6.7–5.7 Ma (latest Miocene), younger than the Bitterwater Creek Shale and at about the same time as parts of the Sisquoc Formation and Purisima Formation in coastal California. Our results show that the Bitterwater Creek Shale and Pancho Rico Formation are lithologically unlike and of different ages and therefore do not constitute a cross-fault tie that can be used to estimate rightlateral displacement along the San Andreas Fault.In the Maricopa area northeast of the San Andreas Fault, the Bitterwater Creek Shale overlies conglomeratic fan-delta deposits of the upper Miocene Santa Margarita Formation, which in turn overlie siliceous shale of the Miocene Monterey Formation from which we obtained a diatom assemblage dated at about 10.0–9.3 Ma. Previous investigations noted that the Santa Margarita Formation in the Maricopa area contains granitic and metamorphic clasts derived from sources in the northern Gabilan Range, on the opposite side of

  11. S-wave triggering of tremor beneath the Parkfield, California, section of the San Andreas fault by the 2011 Tohoku, Japan earthquake: observations and theory (United States)

    Hill, David P.; Peng, Zhigang; Shelly, David R.; Aiken, Chastity


    The dynamic stresses that are associated with the energetic seismic waves generated by the Mw 9.0 Tohoku earthquake off the northeast coast of Japan triggered bursts of tectonic tremor beneath the Parkfield section of the San Andreas fault (SAF) at an epicentral distance of ∼8200  km. The onset of tremor begins midway through the ∼100‐s‐period S‐wave arrival, with a minor burst coinciding with the SHSH arrival, as recorded on the nearby broadband seismic station PKD. A more pronounced burst coincides with the Love arrival, followed by a series of impulsive tremor bursts apparently modulated by the 20‐ to 30‐s‐period Rayleigh wave. The triggered tremor was located at depths between 20 and 30 km beneath the surface trace of the fault, with the burst coincident with the S wave centered beneath the fault 30 km northwest of Parkfield. Most of the subsequent activity, including the tremor coincident with the SHSH arrival, was concentrated beneath a stretch of the fault extending from 10 to 40 km southeast of Parkfield. The seismic waves from the Tohoku epicenter form a horizontal incidence angle of ∼14°, with respect to the local strike of the SAF. Computed peak dynamic Coulomb stresses on the fault at tremor depths are in the 0.7–10 kPa range. The apparent modulation of tremor bursts by the small, strike‐parallel Rayleigh‐wave stresses (∼0.7  kPa) is likely enabled by pore pressure variations driven by the Rayleigh‐wave dilatational stress. These results are consistent with the strike‐parallel dynamic stresses (δτs) associated with the S, SHSH, and surface‐wave phases triggering small increments of dextral slip on the fault with a low friction (μ∼0.2). The vertical dynamic stresses δτd do not trigger tremor with vertical or oblique slip under this simple Coulomb failure model.

  12. Within-wing isotopic (δ2H, δ13C, δ15N variation of monarch butterflies: implications for studies of migratory origins and diet

    Directory of Open Access Journals (Sweden)

    Hobson Keith A.


    Full Text Available Increasingly, stable isotope measurements are being used to assign individuals to broad geographic origins based on established relationships between animal tissues and tissue-specific isoscapes. In particular, the eastern North American population of the monarch butterfly (Danaus plexippus has been the subject of several studies using established δ2H and δ13C wingtissue isoscapes to infer natal origins of migrating and overwintering individuals. However, there has been no study investigating potential variance that can derive from subsampling different regions of the wings, especially those regions differing in pigmentation (orange versus black. Within-wing isotopic (δ2H, δ13C, δ15N variance of 40 monarch butterflies collected from natural overwinter mortality on Mexican roost sites were split evenly into two groups: unwashed samples and those washed in a 2:1 chloroform:methanol solvent. Isotopic variance in δ2H and δ13C was related to pigment (within-wing range 5‰ and 0.5‰, respectively, but not region of subsampling. This variance was reduced 3 to 4 fold through solvent washing that removed pigmented surface scales and any adhered oils. Wing δ15N was similarly influenced by pigment (range 0.3‰, but this effect was not reduced through washing. We recommend future isotopic studies of monarchs and other butterflies for migration research to use the same region for subsampling consistently and to wash samples with solvent to reduce isotopic variance related to uncontrolled variance in discrimination (δ2H, δ13C, δ15N and/or adsorbed water vapor (δ2H. These data also need to be included in description of methods.

  13. Role of forest conservation in lessening land degradation in a temperate region: the Monarch Butterfly Biosphere Reserve, Mexico. (United States)

    Manzo-Delgado, Lilia; López-García, José; Alcántara-Ayala, Irasema


    With international concern about the rates of deforestation worldwide, particular attention has been paid to Latin America. Forest conservation programmes in Mexico include Payment for Environmental Services (PES), a scheme that has been successfully introduced in the Monarch Butterfly Biosphere Reserve. To seek further evidence of the role of PES in lessening land degradation processes in a temperate region, the conservation state of the Cerro Prieto ejido within the Reserve was assessed by an analysis of changes in vegetation cover and land-use between 1971 and 2013. There were no changes in the total forest surface area, but the relative proportions of the different classes of cover density had changed. In 1971, closed and semi-closed forest occupied 247.81 ha and 5.38 ha, 82.33% and 1.79% of the total area of the ejido, respectively. By 2013, closed forest had decreased to 230.38 ha (76.54% of the ejido), and semi-closed cover was 17.23 ha (5.72% of the ejido), suggesting that some semi-closed forest had achieved closed status. The final balance between forest losses and recovery was: 29.63 ha were lost, whereas 13.72 ha were recovered. Losses were mainly linked to a sanitation harvest programme to control the bark beetle Scolytus mundus. Ecotourism associated with forest conservation in the Cerro Prieto ejido has been considered by inhabitants as a focal alternative for economic development. Consequently, it is essential to develop a well-planned and solidly structured approach based on social cohesion to foster a community-led sustainable development at local level. Copyright © 2013 Elsevier Ltd. All rights reserved.

  14. Assimilation of MODIS Dark Target and Deep Blue Observations in the Dust Aerosol Component of NMMB-MONARCH version 1.0 (United States)

    Di Tomaso, Enza; Schutgens, Nick A. J.; Jorba, Oriol; Perez Garcia-Pando, Carlos


    A data assimilation capability has been built for the NMMB-MONARCH chemical weather prediction system, with a focus on mineral dust, a prominent type of aerosol. An ensemble-based Kalman filter technique (namely the local ensemble transform Kalman filter - LETKF) has been utilized to optimally combine model background and satellite retrievals. Our implementation of the ensemble is based on known uncertainties in the physical parametrizations of the dust emission scheme. Experiments showed that MODIS AOD retrievals using the Dark Target algorithm can help NMMB-MONARCH to better characterize atmospheric dust. This is particularly true for the analysis of the dust outflow in the Sahel region and over the African Atlantic coast. The assimilation of MODIS AOD retrievals based on the Deep Blue algorithm has a further positive impact in the analysis downwind from the strongest dust sources of the Sahara and in the Arabian Peninsula. An analysis-initialized forecast performs better (lower forecast error and higher correlation with observations) than a standard forecast, with the exception of underestimating dust in the long-range Atlantic transport and degradation of the temporal evolution of dust in some regions after day 1. Particularly relevant is the improved forecast over the Sahara throughout the forecast range thanks to the assimilation of Deep Blue retrievals over areas not easily covered by other observational datasets.The present study on mineral dust is a first step towards data assimilation with a complete aerosol prediction system that includes multiple aerosol species.

  15. Climate-change and mass mortality events in overwintering monarch butterflies Eventos de mortandad masiva y cambio climático en poblaciones invernales de la mariposa monarca

    Directory of Open Access Journals (Sweden)

    Narayani Barve


    Full Text Available Monarch butterflies (Danaus plexippus have a unique yearly life cycle, in which successive generations breed and move northward from the southern USA in spring to the northern US and southern Canada by late summer; they overwinter in extremely restricted areas in central Mexico and along the California coast. Mexican overwintering populations have experienced significant mortality events recently, which have been hypothesized as increasing in frequency owing to climate change. Here, we test the hypothesis of climate-change causation of these mortality events, at least in part, finding significant local weather trends toward conditions lethal for monarch survival. We use ecological niche estimates and future climate projections to estimate future overwintering distributions; results anticipate dramatic reductions in suitability of present overwintering areas, and serious implications for local human economies.La mariposa monarca (Danaus plexippus tiene un ciclo de vida singular, en el cual generaciones sucesivas se reproducen y migran hacia el norte, empezando en el sur de los Estados Unidos en la primavera y terminando en el norte de los Estados Unidos y sur del Canadá en verano. Pasan el invierno en unas pocas zonas muy restringidas del centro de México y la costa del estado de California. En tiempos recientes, las poblaciones en México han experimentado mortalidades significativas y se ha hipotetizado que la causa puede ser el cambio climático. En este artículo probamos, al menos en parte, la hipótesis del cambio climático como causa de estos eventos de mortalidad y encontramos un desplazamiento significativo del clima local hacia condiciones que son letales para la mariposa. Utilizamos estimados de nicho ecológico y proyecciones de climas futuros para definir futuras áreas de invernación. Nuestros resultados anticipan una reducción dramática en la calidad de estas áreas actuales e implicaciones serias para las economías locales.

  16. Detection of trees damaged by pests in Abies religiosa forests in the Monarch Butterfly Biosphere Reserve using infrared aerial photography

    Directory of Open Access Journals (Sweden)

    Pablo Leautaud Valenzuela


    photographic mosaic of the sampling area. The unassisted and assisted spectral classification technique was carried out in the ERDAS Imagine image-processing software package. For the unassisted classification, tests were carried out considering various numbers of categories: 5, 10 and 15; the assisted classification included the spectral properties of each category used for the partition to group images into five categories: healthy forest, diseased forest, Juniperus scrubland, bare soil and shaded areas. The accuracy of the technique for the detection of damaged trees was verified through field work, visiting different checkpoints where the health status of the tree was corroborated by direct observation and infrared photography at ground level. A representative sampling area of the A. religiosa forest was established in the Monarch Butterfly Biosphere Reserve (RBMM, sufficient to encompass the largest number of damaged trees, but not so large as to excessively prolong the information-processing phases and make field sampling unattainable.  The analysis comprised an area of 1907 ha in Sierra Chincua, where the greatest affectation was observed in a core zone including 97 points (62% with more than twice the density of individuals (11 trees/km2, relative to the buffer zone (4 trees/km2. This greater damage is the result of forest management policies, which have set no management (including sanitation in the core zone. At the end of this research work, we concluded that digital aerial photographs proved useful for the detection of damaged trees in Abies religiosa forests of RBMM. It is possible to obtain multispectral images using a low-cost photographic technology that is relatively simple and widely available. Our study showed that the best method to detect damage in A. religiosa forests in RBMM is the visual interpretation of aerial photographs, yielding a detection efficiency of over 98%. The method used has a greater costeffectiveness compared to helicopter overflight

  17. Transient stresses al Parkfield, California, produced by the M 7.4 Landers earthquake of June 28, 1992: implications for the time-dependence of fault friction

    Directory of Open Access Journals (Sweden)

    J. B. Fletcher


    Full Text Available he M 7.4 Landers earthquake triggered widespread seismicity in the Western U.S. Because the transient dynamic stresses induced at regional distances by the Landers surface waves are much larger than the expected static stresses, the magnitude and the characteristics of the dynamic stresses may bear upon the earthquake triggering mechanism. The Landers earthquake was recorded on the UPSAR array, a group of 14 triaxial accelerometers located within a 1-square-km region 10 km southwest of the town of Parkfield, California, 412 km northwest of the Landers epicenter. We used a standard geodetic inversion procedure to determine the surface strain and stress tensors as functions of time from the observed dynamic displacements. Peak dynamic strains and stresses at the Earth's surface are about 7 microstrain and 0.035 MPa, respectively, and they have a flat amplitude spectrum between 2 s and 15 s period. These stresses agree well with stresses predicted from a simple rule of thumb based upon the ground velocity spectrum observed at a single station. Peak stresses ranged from about 0.035 MPa at the surface to about 0.12 MPa between 2 and 14 km depth, with the sharp increase of stress away from the surface resulting from the rapid increase of rigidity with depth and from the influence of surface wave mode shapes. Comparison of Landers-induced static and dynamic stresses at the hypocenter of the Big Bear aftershock provides a clear example that faults are stronger on time scales of tens of seconds than on time scales of hours or longer.

  18. Correlation between deep fluids, tremor and creep along the central San Andreas fault. (United States)

    Becken, Michael; Ritter, Oliver; Bedrosian, Paul A; Weckmann, Ute


    The seismicity pattern along the San Andreas fault near Parkfield and Cholame, California, varies distinctly over a length of only fifty kilometres. Within the brittle crust, the presence of frictionally weak minerals, fault-weakening high fluid pressures and chemical weakening are considered possible causes of an anomalously weak fault northwest of Parkfield. Non-volcanic tremor from lower-crustal and upper-mantle depths is most pronounced about thirty kilometres southeast of Parkfield and is thought to be associated with high pore-fluid pressures at depth. Here we present geophysical evidence of fluids migrating into the creeping section of the San Andreas fault that seem to originate in the region of the uppermost mantle that also stimulates tremor, and evidence that along-strike variations in tremor activity and amplitude are related to strength variations in the lower crust and upper mantle. Interconnected fluids can explain a deep zone of anomalously low electrical resistivity that has been imaged by magnetotelluric data southwest of the Parkfield-Cholame segment. Near Cholame, where fluids seem to be trapped below a high-resistivity cap, tremor concentrates adjacent to the inferred fluids within a mechanically strong zone of high resistivity. By contrast, subvertical zones of low resistivity breach the entire crust near the drill hole of the San Andreas Fault Observatory at Depth, northwest of Parkfield, and imply pathways for deep fluids into the eastern fault block, coincident with a mechanically weak crust and the lower tremor amplitudes in the lower crust. Fluid influx to the fault system is consistent with hypotheses of fault-weakening high fluid pressures in the brittle crust.

  19. Alberdi’s monarchic temptation

    Directory of Open Access Journals (Sweden)

    Horacio Crespo


    Full Text Available This article studies Alberdi’s book Del Gobierno De Sud-América Según Las Miras De Su Revolución Fundamental, which was written between 1863 and 1867 – when a monarchy was imposed in Mexico and it was assumed that many South American countries would follow the same path – and published posthumously in 1896. In this controversial text, the author discussed the convenience and adaptation of monarchy to the necessary conditions for progress in Latin America and showed, in the book’s most provocative reflection, an ambiguous preference for constitutional monarchy over republican government. In the end, Alberdi comes out in favor of a centralist republic with a civilizing mission and a secure grip on power.

  20. Continuous borehole strain and pore pressure in the near field of the 28 September 2004 M 6.0 parkfield, California, earthquake: Implications for nucleation, fault response, earthquake prediction and tremor (United States)

    Johnston, M.J.S.; Borcherdt, R.D.; Linde, A.T.; Gladwin, M.T.


    Near-field observations of high-precision borehole strain and pore pressure, show no indication of coherent accelerating strain or pore pressure during the weeks to seconds before the 28 September 2004 M 6.0 Parkfield earthquake. Minor changes in strain rate did occur at a few sites during the last 24 hr before the earthquake but these changes are neither significant nor have the form expected for strain during slip coalescence initiating fault failure. Seconds before the event, strain is stable at the 10-11 level. Final prerupture nucleation slip in the hypocentral region is constrained to have a moment less than 2 ?? 1012 N m (M 2.2) and a source size less than 30 m. Ground displacement data indicate similar constraints. Localized rupture nucleation and runaway precludes useful prediction of damaging earthquakes. Coseismic dynamic strains of about 10 microstrain peak-to-peak were superimposed on volumetric strain offsets of about 0.5 microstrain to the northwest of the epicenter and about 0.2 microstrain to the southeast of the epicenter, consistent with right lateral slip. Observed strain and Global Positioning System (GPS) offsets can be simply fit with 20 cm of slip between 4 and 10 km on a 20-km segment of the fault north of Gold Hill (M0 = 7 ?? 1017 N m). Variable slip inversion models using GPS data and seismic data indicate similar moments. Observed postseismic strain is 60% to 300% of the coseismic strain, indicating incomplete release of accumulated strain. No measurable change in fault zone compliance preceding or following the earthquake is indicated by stable earth tidal response. No indications of strain change accompany nonvolcanic tremor events reported prior to and following the earthquake.

  1. Dendroclimatic analysis of Pinus pseudostrobus and Pinus devoniana in the municipalities of Áporo and Zitácuaro (Michoacán, Monarch Butterfly Biosphere Reserve

    Directory of Open Access Journals (Sweden)

    Jaume Marlès Magre


    Full Text Available This article presents the first study on dendroclimatology of Pinus pseudostrobus and Pinus devoniana in the state of Michoacán (Mexico, specifically in the municipalities of Áporo and Zitácuaro, both municipalities within the Monarch Butterfly Biosphere Reserve (MBBR. The sampling in Áporo, northwest of the MBBR, was held in Los Ejidos del Rincón del Soto and Arroyo Seco, in Sierra Chincúa (May 2011. In Zitácuaro, southwest of the reserve, a sampling was performed in the Ejido de San Juan de Zitácuaro, in the area of Ocotal and Palma, and Meso Sedano (June 2011. There were a total of 38 Pinus pseudostrobus and 12 Pinus devoniana sampled in both areas of the study and distributed in 28 trees in the municipality of Áporo and 22 in Zitácuaro. Two samples per tree were taken at 1.3 m height, resulting in a total of 100 tree cores. The dendrochronological series in Áporo for the species Pinus pseudostrobus were extended to 62 years (1949-2010 and for Pinus devoniana 86 years (1925-2010; and the series in Zitácuaro for Pinus pseudostrobus and Pinus devoniana were extended to 47 years (1964-2010 and 44 years (1967-2010, respectively. The ring chronologies were validated using the program COFECHA, which calculates the cross correlations between individual series of the tree-growth, five series were eliminated due to very low or negative correlations. The climate data from Zitácuaro were obtained from two weather stations located in the same municipality. And, in the case of Áporo, the data was obtained from stations located in Senguio. The growth rates related to the climate were obtained by removing the growth trend of each tree due to the age, size and other factors such as the competition, using the program ARSTAN. The following statistics were used to evaluate the quality of the residual chronologies and to determine the potential dendrochronology of species for the different populations: the average correlation between series (Rbar

  2. Electromagnetic Imaging of Fluids in the San Andreas Fault; FINAL

    International Nuclear Information System (INIS)

    Martyn Unsworth


    OAK 270 - Magnetotelluric data were collected on six profiles across the san Andreas Fault at Cholame,Parkfield, and Hollister in Central California. On each profile, high electrical resistivities were imaged west of the fault, and are due to granitic rocks of the Salinian block. East of the fault, lower electrical resistivities are associated with rocks of the Fanciscan formation. On the seismically active Parkfield and Hollister segments, a region of low resistivity was found in the fault zone that extends to a depth of several kilometers. This is due to a zone of fracturing (the damaged zone) that has been infiltrated by saline ground water. The shallowest micro-earthquakers occur at a depth that is coincident with the base of the low resistivity wedge. This strongly suggests that above this depth, the fault rocks are too weak to accumulate sufficient stress for earthquake rupture to occur and fault motion is accommodated through aseismic creep

  3. monarchical monstrosity in postcolonial literature: a reading of femi ...

    African Journals Online (AJOL)

    Dean SPGS NAU

    This paper is a thematic study of Femi Osofisan's The Chattering and the .... democracy and from a lack of consensus on the rules of ...... *Clement Olujide Ajidahun, PhD, Department of English Studies, ... Developments in Chile and Argentina.

  4. Paul Downes. Democracy, Revolution and Monarchism in Early American Literature.

    Directory of Open Access Journals (Sweden)

    Jean-Pierre Martin


    Full Text Available L’auteur, Associate Professor, Department of English, University of Toronto, part implicitement d’une évidence : pour l’ensemble hétérogène des colonies américaines rebelles, le lien à la Couronne est le seul commun dénominateur, et le trajet le plus court de Charleston à Boston passe par Londres…D’où, pour un tiers environ de la population — les Loyalistes— une fidèlité à la monarchie poussée parfois jusqu’à l’exil volontaire ; mais pour les Indépendantistes, nolentes volentes, des référence...

  5. Monarchical monstrosity in postcolonial literature: a reading of Femi ...

    African Journals Online (AJOL)

    This paper is a thematic study of Femi Osofisan's The Chattering and the Song and Yungba-Yungba and the Dance Contest. Femi Osofisan, in the two plays, exposes the political leadership in Africa as characterized by dictatorship, despotism, tyranny and corruption. The paper provides a theoretical framework where ...

  6. Neo-sex chromosomes in the monarch butterfly, Danaus plexippus

    Czech Academy of Sciences Publication Activity Database

    Mongue, A. J.; Nguyen, Petr; Voleníková, Anna; Walters, J. R.


    Roč. 7, č. 10 (2017), s. 3281-3294 ISSN 2160-1836 R&D Projects: GA ČR(CZ) GA14-22765S; GA ČR(CZ) GP14-35819P Grant - others:GA JU(CZ) 159/2016/P Institutional support: RVO:60077344 Keywords : sex chromosomes * evolution * Lepidoptera Subject RIV: EB - Genetics ; Molecular Biology OBOR OECD: Genetics and heredity (medical genetics to be 3) Impact factor: 2.861, year: 2016

  7. Pre-monarchical political leadership among the Gas, with special ...

    African Journals Online (AJOL)

    ... Gas to the coast. The transition from priestly theocracy to chiefly rule became necessary as a response to the pressure on the Gas as the middlemen between the merchants on the coast and the other inland groups. Institute of African Studies: Research Review Volume Research Review Supplement 17 2006: pp. 137-147 ...

  8. Locating Very-Low-Frequency Earthquakes in the San Andreas Fault. (United States)

    Peña-Castro, A. F.; Harrington, R. M.; Cochran, E. S.


    The portion of tectonic fault where rheological properties transtition from brittle to ductile hosts a variety of seismic signals suggesting a range of slip velocities. In subduction zones, the two dominantly observed seismic signals include very-low frequency earthquakes ( VLFEs), and low-frequency earthquakes (LFEs) or tectonic tremor. Tremor and LFE are also commonly observed in transform faults, however, VLFEs have been reported dominantly in subduction zone environments. Here we show some of the first known observations of VLFEs occurring on a plate boundary transform fault, the San Andreas Fault (SAF) between the Cholame-Parkfield segment in California. We detect VLFEs using both permanent and temporary stations in 2010-2011 within approximately 70 km of Cholame, California. We search continous waveforms filtered from 0.02-0.05 Hz, and remove time windows containing teleseismic events and local earthquakes, as identified in the global Centroid Moment Tensor (CMT) and the Northern California Seismic Network (NCSN) catalog. We estimate the VLFE locations by converting the signal into envelopes, and cross-correlating them for phase-picking, similar to procedures used for locating tectonic tremor. We first perform epicentral location using a grid search method and estimate a hypocenter location using Hypoinverse and a shear-wave velocity model when the epicenter is located close to the SAF trace. We account for the velocity contrast across the fault using separate 1D velocity models for stations on each side. Estimated hypocentral VLFE depths are similar to tremor catalog depths ( 15-30 km). Only a few VLFEs produced robust hypocentral locations, presumably due to the difficulty in picking accurate phase arrivals with such a low-frequency signal. However, for events for which no location could be obtained, the moveout of phase arrivals across the stations were similar in character, suggesting that other observed VLFEs occurred in close proximity.

  9. Precise Relative Location of San Andreas Fault Tremors Near Cholame, CA, Using Seismometer Clusters: Slip on the Deep Extension of the Fault? (United States)

    Shelly, D. R.; Ellsworth, W. L.; Ryberg, T.; Haberland, C.; Fuis, G.; Murphy, J.; Nadeau, R.; Bürgmann, R.


    Non-volcanic tremor, similar in character to that generated at some subduction zones, was recently identified beneath the strike-slip San Andreas Fault (SAF) in central California (Nadeau and Dolenc, 2005). Using a matched filter method, we closely examine a 24-hour period of active SAF tremor and show that, like tremor in the Nankai Trough subduction zone, this tremor is composed of repeated similar events. We take advantage of this similarity to locate detected similar events relative to several chosen events. While low signal-to-noise makes location challenging, we compensate for this by estimating event-pair differential times at 'clusters' of nearby temporary and permanent stations rather than at single stations. We find that the relative locations consistently form a near-linear structure in map view, striking parallel to the surface trace of the SAF. Therefore, we suggest that at least a portion of the tremor occurs on the deep extension of the fault, similar to the situation for subduction zone tremor. Also notable is the small depth range (a few hundred meters or less) of many of the located tremors, a feature possibly analogous to earthquake streaks observed on the shallower portion of the fault. The close alignment of the tremor with the SAF slip orientation suggests a shear slip mechanism, as has been argued for subduction tremor. At times, we observe a clear migration of the tremor source along the fault, at rates of 15-40 km/hr.

  10. A deep crustal fluid channel into the San Andreas Fault system near Parkfield, California (United States)

    Becken, M.; Ritter, O.; Park, S.K.; Bedrosian, P.A.; Weckmann, U.; Weber, M.


    Magnetotelluric (MT) data from 66 sites along a 45-km-long profile across the San Andreas Fault (SAF) were inverted to obtain the 2-D electrical resistivity structure of the crust near the San Andreas Fault Observatory at Depth (SAFOD). The most intriguing feature of the resistivity model is a steeply dipping upper crustal high-conductivity zone flanking the seismically defined SAF to the NE, that widens into the lower crust and appears to be connected to a broad conductivity anomaly in the upper mantle. Hypothesis tests of the inversion model suggest that upper and lower crustal and upper-mantle anomalies may be interconnected. We speculate that the high conductivities are caused by fluids and may represent a deep-rooted channel for crustal and/or mantle fluid ascent. Based on the chemical analysis of well waters, it was previously suggested that fluids can enter the brittle regime of the SAF system from the lower crust and mantle. At high pressures, these fluids can contribute to fault-weakening at seismogenic depths. These geochemical studies predicted the existence of a deep fluid source and a permeable pathway through the crust. Our resistivity model images a conductive pathway, which penetrates the entire crust, in agreement with the geochemical interpretation. However, the resistivity model also shows that the upper crustal branch of the high-conductivity zone is located NE of the seismically defined SAF, suggesting that the SAF does not itself act as a major fluid pathway. This interpretation is supported by both, the location of the upper crustal high-conductivity zone and recent studies within the SAFOD main hole, which indicate that pore pressures within the core of the SAF zone are not anomalously high, that mantle-derived fluids are minor constituents to the fault-zone fluid composition and that both the volume of mantle fluids and the fluid pressure increase to the NE of the SAF. We further infer from the MT model that the resistive Salinian block basement to the SW of the SAFOD represents an isolated body, being 5-8km wide and reaching to depths >7km, in agreement with aeromagnetic data. This body is separated from a massive block of Salinian crust farther to the SW. The NE terminus of resistive Salinian crust has a spatial relationship with a near-vertical zone of increased seismic reflectivity ???15km SW of the SAF and likely represents a deep-reaching fault zone. ?? 2008 The Authors Journal compilation ?? 2008 RAS.

  11. Steep-dip seismic imaging of the shallow San Andreas fault near Parkfield. (United States)

    Hole, J A; Catchings, R D; St Clair, K C; Rymer, M J; Okaya, D A; Carney, B J


    Seismic reflection and refraction images illuminate the San Andreas Fault to a depth of 1 kilometer. The prestack depth-migrated reflection image contains near-vertical reflections aligned with the active fault trace. The fault is vertical in the upper 0.5 kilometer, then dips about 70 degrees to the southwest to at least 1 kilometer subsurface. This dip reconciles the difference between the computed locations of earthquakes and the surface fault trace. The seismic velocity cross section shows strong lateral variations. Relatively low velocity (10 to 30%), high electrical conductivity, and low density indicate a 1-kilometer-wide vertical wedge of porous sediment or fractured rock immediately southwest of the active fault trace.

  12. Relating seismicity to the velocity structure of the San Andreas Fault near Parkfield, CA (United States)

    Lippoldt, Rachel; Porritt, Robert W.; Sammis, Charles G.


    The central section of the San Andreas Fault (SAF) displays a range of seismic phenomena including normal earthquakes, low-frequency earthquakes (LFE), repeating microearthquakes (REQ) and aseismic creep. Although many lines of evidence suggest that LFEs are tied to the presence of fluids, their geological setting is still poorly understood. Here, we map the seismic velocity structures associated with LFEs beneath the central SAF using surface wave tomography from ambient seismic noise to provide constraints on the physical conditions that control LFE occurrence. Fault perpendicular sections show that the SAF, as revealed by lateral contrasts in relative velocities, is contiguous to depths of 50 km and appears to be relatively localized at depths between about 15 and 30 km. This is consistent with the hypothesis that LFEs are shear-slip events on a deep extension of the SAF. We find that along strike variations in seismic behaviour correspond to changes in the seismic structure, which support proposed connections between fluids and seismicity. LFEs and REQs occur within low-velocity structures, suggesting that the presence of fluids, weaker minerals, or hydrous phase minerals may play an important role in the generation of slow-slip phenomena.

  13. Modular Open Network ARCHitecture (MONARCH): Transitioning plug-and-play to aerospace (United States)

    Martin, M.; Lyke, J.

    The Air Force Research Laboratory (AFRL) developed an initial plug-and-play (PnP) capability for spacecraft, similar to USB on personal computers, which better defines hardware and software interfaces and incorporates self-discovery and auto-configuration in order to simplify spacecraft development and reduce cost and schedule. PnP technology was matured through a suborbital PnP flight experiment in September 2007 and a secondary Spacecraft Avionics Experiment (SAE) payload on the TacSat-3 satellite, which launched in May 2009. AFRL developed and submitted a complete set of PnP standards through the American Institute of Aeronautics and Astronautics (AIAA) in 2011. Space electronics to adapt existing satellite components and implement full PnP on satellites in accordance with these AFRL standards was independently developed in alternate hardware implementations by Goodrich Corp under AFRL and by Northrop Grumman under Operationally Responsive Space (ORS). In 2011, AFRL conducted a cost-benefit analysis of PnP and assembled a collaborative review board (CRB) in Sept 2011 to evaluate PnP. This CRB was comprised of representatives from Space and Missiles Center (SMC), National Reconnaissance Organization (NRO), Naval Research Laboratory (NRL), John Hopkins University (JHU) Applied Physics Laboratory (APL), The Aerospace Corporation, and several large commercial and DOD satellite developers. This CRB laid out a transition path to develop and implement PnP standards for implementation in large (> 1000 kg) DOD and commercial satellites. Transition of PnP technology into operational systems continues in PnP architecture studies for SMC, PnP products from multiple space industry vendors, commercial implementations of PnP, and the Northrop Grumman ORS-2 spacecraft currently project to fly in 2014-2015. This paper provides details related to development of PnP technology, AFRL's cost-benefit analysis of PnP, recommendations of the PnP CRB, and on-going efforts to mature - nd fly PnP technology.

  14. Antimasonian activity of Russian monarchical emigration of «first wave» (1917–1940

    Directory of Open Access Journals (Sweden)

    Ermakov V.A.


    Full Text Available summary: The work shows that in the modern domestic liberal historiography of Russian emigration, there is a tendency to conceal and hide political problems raised by Russian monarchists. And, on the contrary, in the works of representatives of the patriotic direction, a wide range of views of Russian monarchists, centered around the Masonic problem and the struggle against Bolshevism, are being explored. The system of estimations by Russian right-wing representatives of the «philosophical steamer» as «apologists of Freemasonry» and the bearers of the ideas of «Masonic democracy» and also as «reformers of Orthodoxy» was reconstructed. The specifics of the struggle of Russian patriotic circles of white emigration with the Russian Masonic lodges restored abroad are shown. The anti-Masonic activities of the Russian Church abroad are considered. As the highest achievement of anti-Masonic publicism is considered the religious and political philosophy of the history of Russian monarchists. The author believe that the main historiographic criterion for distinguishing the political activity of Russian emigration should be recognized as its pro-Masonic or anti-Masonic orientation. As a result of the research, the author come to the conclusion that the anti-Masonic activities of the Russian right largely objectively reflected the main contradiction of the Russian history of «post-October abroad» as the struggle of Russian patriotic monarchists and the Orthodox Church abroad with the Russophobic forces of the West, whose concentrated expression was Freemasonry.

  15. Create a pollinator garden at your nursery: An emphasis on monarch butterflies (United States)

    Thomas D. Landis; R. Kasten Dumroese; Matthew E. Horning


    We realize that this type of article is a departure for FNN readers but feel that it is important for forest, conservation, and native plant nurseries to be good environmental stewards. In addition, establishing a pollinator garden at your nursery can be good for business, too. Demonstrating the role and beauty of native plants and their pollinators, particulary in a...

  16. Famous people and genetic disorders: from monarchs to geniuses--a portrait of their genetic illnesses. (United States)

    Ho, Nicola C; Park, Susan S; Maragh, Kevin D; Gutter, Emily M


    Famous people with genetic disorders have always been a subject of interest because such news feeds the curiosity the public has for celebrities. It gives further insight into their lives and provides a medical basis for any unexplained or idiosyncratic feature or behavior they exhibit. It draws admiration from society of those who excel in their specialized fields despite the impositions of their genetic illnesses and also elicits sympathy even in the most casual observer. Such news certainly catapults a rare genetic disorder into the realm of public awareness. We hereby present six famous figures: King George III, Toulouse-Lautrec, Queen Victoria, Nicolo Paganini, Abraham Lincoln, and Vincent van Gogh, all of whom made a huge indelible mark in either the history of politics or that of the arts. Copyright 2003 Wiley-Liss, Inc.

  17. Using an Earthquake Simulator to Model Tremor Along a Strike Slip Fault (United States)

    Cochran, E. S.; Richards-Dinger, K. B.; Kroll, K.; Harrington, R. M.; Dieterich, J. H.


    We employ the earthquake simulator, RSQSim, to investigate the conditions under which tremor occurs in the transition zone of the San Andreas fault. RSQSim is a computationally efficient method that uses rate- and state- dependent friction to simulate a wide range of event sizes for long time histories of slip [Dieterich and Richards-Dinger, 2010; Richards-Dinger and Dieterich, 2012]. RSQSim has been previously used to investigate slow slip events in Cascadia [Colella et al., 2011; 2012]. Earthquakes, tremor, slow slip, and creep occurrence are primarily controlled by the rate and state constants a and b and slip speed. We will report the preliminary results of using RSQSim to vary fault frictional properties in order to better understand rupture dynamics in the transition zone using observed characteristics of tremor along the San Andreas fault. Recent studies of tremor along the San Andreas fault provide information on tremor characteristics including precise locations, peak amplitudes, duration of tremor episodes, and tremor migration. We use these observations to constrain numerical simulations that examine the slip conditions in the transition zone of the San Andreas Fault. Here, we use the earthquake simulator, RSQSim, to conduct multi-event simulations of tremor for a strike slip fault modeled on Cholame section of the San Andreas fault. Tremor was first observed on the San Andreas fault near Cholame, California near the southern edge of the 2004 Parkfield rupture [Nadeau and Dolenc, 2005]. Since then, tremor has been observed across a 150 km section of the San Andreas with depths between 16-28 km and peak amplitudes that vary by a factor of 7 [Shelly and Hardebeck, 2010]. Tremor episodes, comprised of multiple low frequency earthquakes (LFEs), tend to be relatively short, lasting tens of seconds to as long as 1-2 hours [Horstmann et al., in review, 2013]; tremor occurs regularly with some tremor observed almost daily [Shelly and Hardebeck, 2010; Horstmann

  18. Joint innversion of seismic and magnetotelluric data in the Parkfield Region of California using the normalized cross-gradient constraint (United States)

    Bennington, Ninfa L.; Zhang, Haijiang; Thurber, Cliff; Bedrosian, Paul A.


    We present jointly inverted models of P-wave velocity (Vp) and electrical resistivity for a two-dimensional profile centered on the San Andreas Fault Observatory at Depth (SAFOD). Significant structural similarity between main features of the separately inverted Vp and resistivity models is exploited by carrying out a joint inversion of the two datasets using the normalized cross-gradient constraint. This constraint favors structurally similar Vp and resistivity images that adequately fit the seismic and magnetotelluric (MT) datasets. The new inversion code, tomoDDMT, merges the seismic inversion code tomoDD and the forward modeling and sensitivity kernel subroutines of the MT inversion code OCCAM2DMT. TomoDDMT is tested on a synthetic dataset and demonstrates the code’s ability to more accurately resolve features of the input synthetic structure relative to the separately inverted resistivity and velocity models. Using tomoDDMT, we are able to resolve a number of key issues raised during drilling at SAFOD. We are able to infer the distribution of several geologic units including the Salinian granitoids, the Great Valley sequence, and the Franciscan Formation. The distribution and transport of fluids at both shallow and great depths is also examined. Low values of velocity/resistivity attributed to a feature known as the Eastern Conductor (EC) can be explained in two ways: the EC is a brine-filled, high porosity region, or this region is composed largely of clay-rich shales of the Franciscan. The Eastern Wall, which lies immediately adjacent to the EC, is unlikely to be a fluid pathway into the San Andreas Fault’s seismogenic zone due to its observed higher resistivity and velocity values.

  19. Characterization and application of microearthquake clusters to problems of scaling, fault zone dynamics, and seismic monitoring at Parkfield, California

    Energy Technology Data Exchange (ETDEWEB)

    Nadeau, Robert Michael [Univ. of California, Berkeley, CA (United States)


    This document contains information about the characterization and application of microearthquake clusters and fault zone dynamics. Topics discussed include: Seismological studies; fault-zone dynamics; periodic recurrence; scaling of microearthquakes to large earthquakes; implications of fault mechanics and seismic hazards; and wave propagation and temporal changes.

  20. Using Online Active-Learning Techniques to Convey Time Compensated Sun Compass Orientation in the Eastern North American Monarch


    Noah Hammond Green; Douglas G. McMahon; Cynthia Brame


    A common tool that animals use to navigate in a constant direction is known as ?time compensated sun compass orientation.? This is a process by which animals use the position of the sun along with information from their internal circadian clocks to determine and maintain a directional heading. Many circadian scientists and educators use this process as an example of how the internal circadian clock can directly influence animal behavior. However, many students have difficulty grasping this bi...

  1. Coal and Oil: The Dark Monarchs of Global Energy: Understanding Supply and Extraction Patterns and their Importance for Future Production

    International Nuclear Information System (INIS)

    Hoeoek, Mikael


    The formation of modern society has been dominated by coal and oil, and together these two fossil fuels account for nearly two thirds of all primary energy used by mankind. This makes future production a key question for future social development and this thesis attempts to answer whether it is possible to rely on an assumption of ever increasing production of coal and oil. Both coal and oil are finite resources, created over long time scales by geological processes. It is thus impossible to extract more fossil fuels than geologically available. In other words, there are limits to growth imposed by nature. The concept of depletion and exhaustion of recoverable resources is a fundamental question for the future extraction of coal and oil. Historical experience shows that peaking is a well established phenomenon in production of various natural resources. Coal and oil are no exceptions, and historical data shows that easily exploitable resources are exhausted while more challenging deposits are left for the future. For oil, depletion can also be tied directly to the physical laws governing fluid flows in reservoirs. Understanding and predicting behaviour of individual fields, in particularly giant fields, are essential for understanding future production. Based on comprehensive databases with reserve and production data for hundreds of oilfields, typical patterns were found. Alternatively, depletion can manifest itself indirectly through various mechanisms. This has been studied for coal. Over 60% of the global crude oil production is derived from only around 330 giant oilfields, where many of them are becoming increasingly mature. The annual decline in existing oil production has been determined to be around 6% and it is unrealistic that this will be offset by new field developments, additional discoveries or unconventional oil. This implies that the peak of the oil age is here. For coal a similar picture emerges, where 90% of the global coal production originates from only 6 countries. Some of them, such as the USA show signs of increasing maturity and exhaustion of the recoverable amounts. However, there is a greater uncertainty about the recoverable reserves and coal production may yield a global maximum somewhere between 2030 and 2060. This analysis shows that the global production peaks of both oil and coal can be expected comparatively soon. This has significant consequences for the global energy supply and society, economy and environment. The results of this thesis indicate that these challenges should not be taken lightly

  2. On the origin of mixed-layered clay minerals from the San Andreas Fault at 2.5-3 km vertical depth (SAFOD drillhole at Parkfield, California) (United States)

    Schleicher, A. M.; Warr, L. N.; van der Pluijm, B. A.


    A detailed mineralogical study is presented of the matrix of mudrocks sampled from spot coring at three key locations along the San Andreas Fault Observatory at depth (SAFOD) drill hole. The characteristics of authigenic illite-smectite (I-S) and chlorite-smectite (C-S) mixed-layer mineral clays indicate a deep diagenetic origin. A randomly ordered I-S mineral with ca. 20-25% smectite layers is one of the dominant authigenic clay species across the San Andreas Fault zone (sampled at 3,066 and 3,436 m measured depths/MD), whereas an authigenic illite with ca. 2-5% smectite layers is the dominant phase beneath the fault (sampled at 3,992 m MD). The most smectite-rich mixed-layered assemblage with the highest water content occurs in the actively deforming creep zone at ca. 3,300-3,353 m (true vertical depth of ca. 2.7 km), with I-S (70:30) and C-S (50:50). The matrix of all mudrock samples show extensive quartz and feldspar (both plagioclase and K-feldspar) dissolution associated with the crystallization of pore-filling clay minerals. However, the effect of rock deformation in the matrix appears only minor, with weak flattening fabrics defined largely by kinked and fractured mica grains. Adopting available kinetic models for the crystallization of I-S in burial sedimentary environments and the current borehole depths and thermal structure, the conditions and timing of I-S growth can be evaluated. Assuming a typical K+ concentration of 100-200 ppm for sedimentary brines, a present-day geothermal gradient of 35°C/km and a borehole temperature of ca. 112°C for the sampled depths, most of the I-S minerals can be predicted to have formed over the last 4-11 Ma and are probably still in equilibrium with circulating fluids. The exception to this simple burial pattern is the occurrence of the mixed layered phases with higher smectite content than predicted by the burial model. These minerals, which characterize the actively creeping section of the fault and local thin film clay coating on polished brittle slip surfaces, can be explained by the influence of either cooler fluids circulating along this segment of the fault or the flow of K+-depleted brines.

  3. Global search of triggered non-volcanic tremor (United States)

    Chao, Tzu-Kai Kevin

    chapter focuses on a systematic comparison of triggered tremor around the Calaveras Fault (CF) in northern California (NC), the Parkfield-Cholame section of the San Andreas Fault (SAF) in central California (CC), and the San Jacinto Fault (SJF) in southern California (SC). Out of 42 large (Mw ≥7.5) earthquakes between 2001 and 2010, only the 2002 Mw 7.9 Denali fault earthquake triggered clear tremor in NC and SC. In comparison, abundant triggered and ambient tremor has been observed in CC. Further analysis reveal that the lack of triggered tremor observations in SC and NC is not simply a consequence of their different background noise levels as compared to CC, but rather reflects different background tremor rates in these regions. In the final chapter, I systematically search for triggered tremor following the 2011 Mw9.0 Tohoku-Oki earthquake in the regions where ambient or triggered tremor has been found before. The main purpose is to check whether triggered tremor is observed in regions when certain conditions (e.g., surface wave amplitudes) are met. Triggered tremor is observed in southwest Japan, Taiwan, the Aleutian Arc, south-central Alaska, northern Vancouver Island, the Parkfield-Cholame section of the SAF in CC and the SJF in SC, and the North Island of New Zealand. Such a widespread triggering of tremor is not too surprising because of the large amplitude surface waves (minimum peak value of ˜0.1 cm/s) and the associated dynamic stresses (at least ˜7-8 kPa), which is one of the most important factors in controlling the triggering threshold. The triggered tremor in different region is located close to or nearby the ambient tremor active area. In addition, the amplitudes of triggered tremor have positive correlations with the amplitudes of teleseismic surface waves among many regions. Moreover, both Love and Rayleigh waves participate in triggering tremor in different regions, and their triggering potential is somewhat controlled by the incident angles. In

  4. Prime Contract Awards Alphabetically by Contractor, by State or Country, and Place, FY 88. Part 13. (Marshall Electronics. Inc.-Monarch Crown Corp.) (United States)


    000000a 1-0 1-00000000000 -00-4 39 w0 moon dewca00 acca QCUQ0 w9 100000000 300-4 310 if- 24ɜ IN"" IN 14 4 < 4300 w-4 34 -C- C2 44 * 4 C4 X 1000-4 130...MMC) @) N4 Ř 0 (0 ZCN NNN ON >. 1 W00O4C~ U) 0wa ccd c cO ON 444- - 44 U. I o a006. I0 6 U.w mop01. F6 m00 @0w-4 r- ,CV) N- 0 w W 90N C14 M 0) It0) N...6r- 6I.- r - f6 -6 000 0 00 -.- W004 6 -4 CV0 m mm 000000 40wo P.V-.- 0v Owlm M000 I mn 6n( )40040 P 00 000 r-0- 4 600 0 a1 .N MCC00 1 .4 ON04 N NO4

  5. Listening to their voices: The essence of the experience of special and regular education students as they learn monarch, Danaus plexippus, biology and ecology (United States)

    Koomen, Michele Jean Hollingsworth

    This dissertation reports on a phenomenological study of nine regular and special education students as they studied insect biology and ecology in their inclusive seventh grade life science class. Three fundamental data collection methods of interpretive research (student observations, interviews and artifact analysis) framed the data collection of this study. Hermeneutic phenomenological analysis and a seven-step framework, beginning with establishment of the unit of analysis and ending in theory generation, were used to systematically analyze the data resulting in the emergence of four main themes. The essence of the lived experience of the study participants reveal a variety of ways working with others in groups supported their learning. Students reported that it was easier to share ideas, ask questions and complete their work when they worked together with other classmates. A second finding of this study, It's kind of hard in learning science, exposes some of the anxiety and the challenges that are part of the experiences of both regular and special education students in learning science. A third finding reveals that for the students in this study the practice of inquiry learning in science is fragile. Despite daily opportunities in inquiry activities, many students are fixated in finding the "right" answers and just getting their "work done." The practice of inquiry is also fragile because of the perceptions of how we go about doing and learning science. The perception of practicing science for the special education students was moderated and limited by their viewpoint that science is coupled with language arts. The last major theme describes the manner in which both student groups navigate through science learning using various strategies that contribute to their learning or engaging in behaviors that seem to conceal their learning differences. The results of this research have implication for inclusive classroom teachers, special educators, teacher educators and administrators. Listening to their voices serves to "prime" us to consider and value their perspectives as we make decisions that affect their learning and their lives.

  6. Prime Contract Awards Alphabetically by Contractor, by State or Country, and Place, FY 88. Part 14. (Monarch Engineering-Ogie Trinidad, Inc.) (United States)


    Q 0e - 0 0 at l’- 000000 0 l ON IZOCo 00 0 D 0 ) 0 -a 000000 lMON 10-000 w-3 w <- C- ac 0l’~- < 000000 ajI I 00 0- 3-4 0 coo l’ L2 x- 3I- C-0C CV (a 00 d -Iw’ aJI 00 am -4 -a 00 ml ini a OqW I tow Ci 09 0 1%1 P z IOOW I 00 0 w ILL 4 aj 0o : 0 4 -4 4 .4 ((wawOCI) N .4 W in V WCV) al a a N...C) -4 - - - -A Cl) ll 00 00 W 0 0 a 0 0 0 0 0 0 rojo ) w ro-O rll_ ro-O to-O 0 r-- M 0 " I _SOO :3 CD :Jtc, OO.DOOOOOOoOOOQOOCoOCo (-) c rl) (1) C

  7. Description and Evaluation of the Multiscale Online Nonhydrostatic AtmospheRe CHemistry Model (NMMB-MONARCH) Version 1.0: Gas-Phase Chemistry at Global Scale (United States)

    Badia, Alba; Jorba, Oriol; Voulgarakis, Apostolos; Dabdub, Donald; Garcia-Pando, Carlos Perez; Hilboll, Andreas; Goncalves, Maria; Janjic, Zavisa


    This paper presents a comprehensive description and benchmark evaluation of the tropospheric gas-phase chemistry component of the Multiscale Online Nonhydrostatic AtmospheRe CHemistry model (NMMBMONARCH), formerly known as NMMB/BSC-CTM, that can be run on both regional and global domains. Here, we provide an extensive evaluation of a global annual cycle simulation using a variety of background surface stations (EMEP, WDCGG and CASTNET), ozonesondes (WOUDC, CMD and SHADOZ), aircraft data (MOZAIC and several campaigns), and satellite observations (SCIAMACHY and MOPITT).We also include an extensive discussion of our results in comparison to other state-of-the-art models. We note that in this study, we omitted aerosol processes and some natural emissions (lightning and volcano emissions). The model shows a realistic oxidative capacity across the globe. The seasonal cycle for CO is fairly well represented at different locations (correlations around 0.3-0.7 in surface concentrations), although concentrations are underestimated in spring and winter in the Northern Hemisphere, and are overestimated throughout the year at 800 and 500 hPa in the Southern Hemisphere. Nitrogen species are well represented in almost all locations, particularly NO2 in Europe (root mean square error - RMSE - below 5 ppb). The modeled vertical distributions of NOx and HNO3 are in excellent agreement with the observed values and the spatial and seasonal trends of tropospheric NO2 columns correspond well to observations from SCIAMACHY, capturing the highly polluted areas and the biomass burning cycle throughout the year. Over Asia, the model underestimates NOx from March to August, probably due to an underestimation of NOx emissions in the region. Overall, the comparison of the modeled CO and NO2 with MOPITT and SCIAMACHY observations emphasizes the need for more accurate emission rates from anthropogenic and biomass burning sources (i.e., specification of temporal variability).

  8. Selection of reference genes for RT-qPCR analysis in the monarch butterfly, Danaus plexippus (L.), a migrating bio-indicator (United States)

    Quantitative real-time PCR (qRT-PCR) is a reliable and reproducible technique for measuring and evaluating changes in gene expression. To facilitate gene expression studies and obtain more accurate qRT-PCR data, normalization relative to stable housekeeping genes is required. In this study, expres...

  9. Milkweed (Gentianales: Apocynaceae): A farmscape resource for increasing parasitism of stink bugs (Hemiptera: Pentatomidae) and providing nectar to insect pollinators and monarch butterflies (United States)

    In peanut-cotton farmscapes in Georgia, stink bugs, i.e., Nezara viridula (L.), Euschistus servus (Say), and Chinavia hilaris (Say), develop in peanut and then disperse at the crop-to-crop interface to feed on fruit in cotton. The main objective of this study was to examine the influence of a habit...

  10. The Dirt on Outdoor Classrooms. (United States)

    Rich, Steve


    Explains the planning procedure for outdoor classrooms and introduces an integrated unit on monarch butterflies called the Monarch Watch program. Makes recommendations to solve financial problems of outdoor classrooms. (YDS)

  11. American Illuminations

    DEFF Research Database (Denmark)

    Nye, David

    Illuminated fêtes and civic celebrations began in Renaissance Italy and spread through the courts of Europe. Their fireworks, torches, lamps, and special effects glorified the monarch, marked the birth of a prince, or celebrated military victory. Nineteenth-century Americans rejected such monarch...

  12. Occurrence and host specificity of a neogregarine protozoan in four milkweed butterfly hosts (Danaus spp.). (United States)

    Barriga, Paola A; Sternberg, Eleanore D; Lefèvre, Thierry; de Roode, Jacobus C; Altizer, Sonia


    Throughout their global range, wild monarch butterflies (Danaus plexippus) are infected with the protozoan Ophryocystis elektroscirrha (OE). In monarchs, OE infection reduces pupal eclosion, adult lifespan, adult body size and flight ability. Infection of other butterfly hosts with OE is rare or unknown, and the only previously published records of OE infection were on monarch and queen butterflies (D. gilippus). Here we explored the occurrence and specificity of OE and OE-like parasites in four Danaus butterfly species. We surveyed wild D. eresimus (soldier), D. gilippus (queen), D. petilia (lesser wanderer), and D. plexippus (monarch) from five countries to determine the presence of infection. We conducted five cross-infection experiments, on monarchs and queen butterflies and their OE and OE-like parasites, to determine infection probability and the impact of infection on their hosts. Our field survey showed that OE-like parasites were present in D. gilippus, D. petilia, and D. plexippus, but were absent in D. eresimus. Infection probability varied geographically such that D. gilippus and D. plexippus populations in Puerto Rico and Trinidad were not infected or had low prevalence of infection, whereas D. plexippus from S. Florida and Australia had high prevalence. Cross-infection experiments showed evidence for host specificity, in that OE strains from monarchs were more effective at infecting monarchs than queens, and monarchs were less likely to be infected by OE-like strains from queens and lesser wanderers relative to their own natal strains. Our study showed that queens are less susceptible to OE and OE-like infection than monarchs, and that the reduction in adult lifespan following infection is more severe in monarchs than in queens. Copyright © 2016 Elsevier Inc. All rights reserved.

  13. Genetics Home Reference: familial cold autoinflammatory syndrome (United States)

    ... inflammatory response. Monarch-1 is involved in the inhibition of the inflammatory response. Mutations in the NLRP12 ... cold autoinflammatory syndrome Orphanet: Familial cold urticaria Patient Support and Advocacy Resources (3 links) Autoinflammatory Alliance National ...

  14. 'Praise beyond Words': Psalm 150 as grand finale of the crescendo ...

    African Journals Online (AJOL)



    Sep 1, 2010 ... of this earthly monarch, Books I-III theologically express the history of ..... considering its theological impact on the final Hallel, on the. 16. ...... Herders Theologischer Kommentar zum Alten Testament, Herder, Freiburg/Basel/.

  15. Razum bez nassilija / Mark Ostanin

    Index Scriptorium Estoniae

    Ostanin, Mark


    Laeval Ocean Monarch keskkonnasümpoosioni "Läänemeri: ühine pärand, jagatud vastutus" plenaaristungist, kus ürituse patroon Konstantinoopoli patriarh Bartholomeos kohtus president Arnold Rüütliga

  16. The impacts of the transgenes on the modified crops, non-target soil ...

    African Journals Online (AJOL)


    reduction of emission of greenhouse gases, caused several changes in the modified crop plants, interacted with soil ...... fitness and behaviour of monarch butterfly larvae exposed to a combination of ..... climate change hypothesis. Am. Antiq.

  17. Old Hickory Lake Appendix M To Park Management Shoreline Management Plan (United States)


    attractors, establishment of nesting/forage habitat such as monarch butterfly way stations, construction of chimney swift towers, etc. 22. Leases...Elaeagnus umbellata Thunb.) Japanese Bush honeysuckles (Lonicera japonica.) Amur Bush honeysuckle (Lonicera maackii.) Marrows Bush honeysuckle (Lonicera

  18. DefenseLink Feature: (United States)

    courtesy of U.S. Department of Agriculture. Photo of Jackie Robinson in Kansas City Monarchs uniform African-Americans Have Legacy Of Military Service, Sacrifice WASHINGTON, Feb. 12, 2008 - Since the birth

  19. Genetic Factors and Host Traits Predict Spore Morphology for a Butterfly Pathogen

    Directory of Open Access Journals (Sweden)

    Jacobus C. de Roode


    Full Text Available Monarch butterflies (Danaus plexippus throughout the world are commonly infected by the specialist pathogen Ophryocystis elektroscirrha (OE. This protozoan is transmitted when larvae ingest infectious stages (spores scattered onto host plant leaves by infected adults. Parasites replicate internally during larval and pupal stages, and adult monarchs emerge covered with millions of dormant spores on the outsides of their bodies. Across multiple monarch populations, OE varies in prevalence and virulence. Here, we examined geographic and genetic variation in OE spore morphology using clonal parasite lineages derived from each of four host populations (eastern and western North America, South Florida and Hawaii. Spores were harvested from experimentally inoculated, captive-reared adult monarchs. Using light microscopy and digital image analysis, we measured the size, shape and color of 30 replicate spores per host. Analyses examined predictors of spore morphology, including parasite source population and clone, parasite load, and the following host traits: family line, sex, wing area, and wing color (orange and black pigmentation. Results showed significant differences in spore size and shape among parasite clones, suggesting genetic determinants of morphological variation. Spore size also increased with monarch wing size, and monarchs with larger and darker orange wings tended to have darker colored spores, consistent with the idea that parasite development depends on variation in host quality and resources. We found no evidence for effects of source population on variation in spore morphology. Collectively, these results provide support for heritable variation in spore morphology and a role for host traits in affecting parasite development.

  20. Size distribution of Parkfield’s microearthquakes reflects changes in surface creep rate (United States)

    Tormann, Theresa; Wiemer, Stefan; Metzger, Sabrina; Michael, Andrew J.; Hardebeck, Jeanne L.


    The nucleation area of the series of M6 events in Parkfield has been shown to be characterized by low b-values throughout the seismic cycle. Since low b-values represent high differential stresses, the asperity structure seems to be always stably stressed and even unaffected by the latest main shock in 2004. However, because fault loading rates and applied shear stress vary with time, some degree of temporal variability of the b-value within stable blocks is to be expected. We discuss in this study adequate techniques and uncertainty treatment for a detailed analysis of the temporal evolution of b-values. We show that the derived signal for the Parkfield asperity correlates with changes in surface creep, suggesting a sensitive time resolution of the b-value stress meter, and confirming near-critical loading conditions within the Parkfield asperity.

  1. Determination of Focal Depths of Earthquakes in the Mid-Oceanic Ridges from Amplitude Spectra of Surface Waves (United States)


    Foreshock , mainshock and aftershock of the Parkfield, California earthquake of June 28, 1966. b. The Denver earthquake of August 9, 1967. Let us look...into the results of these tests in more details. (1) Test on the main shock, foreshock and aftershock of the Parkfield earthquake of June 28, 1966...According to McEvilly et. al. (1967), the origin times and locations of.these events were the following: Foreshock June 28, 1966, 04:08:56.2 GMT; 350 57.6

  2. Nonvolcanic tremors deep beneath the San Andreas Fault. (United States)

    Nadeau, Robert M; Dolenc, David


    We have discovered nonvolcanic tremor activity (i.e., long-duration seismic signals with no clear P or S waves) within a transform plate boundary zone along the San Andreas Fault near Cholame, California, the inferred epicentral region of the 1857 Fort Tejon earthquake (moment magnitude approximately 7.8). The tremors occur between 20 to 40 kilometers' depth, below the seismogenic zone (the upper approximately 15 kilometers of Earth's crust where earthquakes occur), and their activity rates may correlate with variations in local earthquake activity.

  3. Simulations of tremor-related creep reveal a weak crustal root of the San Andreas Fault (United States)

    Shelly, David R.; Bradley, Andrew M.; Johnson, Kaj M.


    Deep aseismic roots of faults play a critical role in transferring tectonic loads to shallower, brittle crustal faults that rupture in large earthquakes. Yet, until the recent discovery of deep tremor and creep, direct inference of the physical properties of lower-crustal fault roots has remained elusive. Observations of tremor near Parkfield, CA provide the first evidence for present-day localized slip on the deep extension of the San Andreas Fault and triggered transient creep events. We develop numerical simulations of fault slip to show that the spatiotemporal evolution of triggered tremor near Parkfield is consistent with triggered fault creep governed by laboratory-derived friction laws between depths of 20–35 km on the fault. Simulated creep and observed tremor northwest of Parkfield nearly ceased for 20–30 days in response to small coseismic stress changes of order 104 Pa from the 2003 M6.5 San Simeon Earthquake. Simulated afterslip and observed tremor following the 2004 M6.0 Parkfield earthquake show a coseismically induced pulse of rapid creep and tremor lasting for 1 day followed by a longer 30 day period of sustained accelerated rates due to propagation of shallow afterslip into the lower crust. These creep responses require very low effective normal stress of ~1 MPa on the deep San Andreas Fault and near-neutral-stability frictional properties expected for gabbroic lower-crustal rock.

  4. Earthquake recurrence models fail when earthquakes fail to reset the stress field (United States)

    Tormann, Thessa; Wiemer, Stefan; Hardebeck, Jeanne L.


    Parkfield's regularly occurring M6 mainshocks, about every 25 years, have over two decades stoked seismologists' hopes to successfully predict an earthquake of significant size. However, with the longest known inter-event time of 38 years, the latest M6 in the series (28 Sep 2004) did not conform to any of the applied forecast models, questioning once more the predictability of earthquakes in general. Our study investigates the spatial pattern of b-values along the Parkfield segment through the seismic cycle and documents a stably stressed structure. The forecasted rate of M6 earthquakes based on Parkfield's microseismicity b-values corresponds well to observed rates. We interpret the observed b-value stability in terms of the evolution of the stress field in that area: the M6 Parkfield earthquakes do not fully unload the stress on the fault, explaining why time recurrent models fail. We present the 1989 M6.9 Loma Prieta earthquake as counter example, which did release a significant portion of the stress along its fault segment and yields a substantial change in b-values.

  5. Fault rocks from the SAFOD core samples : implications for weakening at shallow depths along the San Andreas Fault, California

    NARCIS (Netherlands)

    Holdsworth, R.E.; van Diggelen, E.W.E.; Spiers, C.J.; Bresser, J.H.P. de; Walker, R.J.; Bown, L.


    The drilling of a deep borehole across the actively creeping Parkfield segment of the San Andreas Fault Zone (SAFZ), California, and collection of core materials permit direct geological study of fault zone processes at 2–3 km depth. The three drill cores sample both host and fault rocks and pass

  6. Seismic trapped modes in the oroville and san andreas fault zones. (United States)

    Li, Y G; Leary, P; Aki, K; Malin, P


    Three-component borehole seismic profiling of the recently active Oroville, California, normal fault and microearthquake event recording with a near-fault three-component borehole seismometer on the San Andreas fault at Parkfield, California, have shown numerous instances of pronounced dispersive wave trains following the shear wave arrivals. These wave trains are interpreted as fault zone-trapped seismic modes. Parkfield earthquakes exciting trapped modes have been located as deep as 10 kilometers, as shallow as 4 kilometers, and extend 12 kilometers along the fault on either side of the recording station. Selected Oroville and Parkfield wave forms are modeled as the fundamental and first higher trapped SH modes of a narrow low-velocity layer at the fault. Modeling results suggest that the Oroville fault zone is 18 meters wide at depth and has a shear wave velocity of 1 kilometer per second, whereas at Parkfield, the fault gouge is 100 to 150 meters wide and has a shear wave velocity of 1.1 to 1.8 kilometers per second. These low-velocity layers are probably the rupture planes on which earthquakes occur.

  7. Loss of migratory behaviour increases infection risk for a butterfly host (United States)

    Satterfield, Dara A.; Maerz, John C.; Altizer, Sonia


    Long-distance animal migrations have important consequences for infectious disease dynamics. In some cases, migration lowers pathogen transmission by removing infected individuals during strenuous journeys and allowing animals to periodically escape contaminated habitats. Human activities are now causing some migratory animals to travel shorter distances or form sedentary (non-migratory) populations. We focused on North American monarch butterflies and a specialist protozoan parasite to investigate how the loss of migratory behaviours affects pathogen spread and evolution. Each autumn, monarchs migrate from breeding grounds in the eastern US and Canada to wintering sites in central Mexico. However, some monarchs have become non-migratory and breed year-round on exotic milkweed in the southern US. We used field sampling, citizen science data and experimental inoculations to quantify infection prevalence and parasite virulence among migratory and sedentary populations. Infection prevalence was markedly higher among sedentary monarchs compared with migratory monarchs, indicating that diminished migration increases infection risk. Virulence differed among parasite strains but was similar between migratory and sedentary populations, potentially owing to high gene flow or insufficient time for evolutionary divergence. More broadly, our findings suggest that human activities that alter animal migrations can influence pathogen dynamics, with implications for wildlife conservation and future disease risks. PMID:25589600

  8. Komparace íránské a ruské revoluce v dílech dvou současných exilových íránských autorů

    Directory of Open Access Journals (Sweden)

    Marek Německý


    Full Text Available Article describes the books of two authors, who try to compare Russian and Iranian revolutions and political development generally. Whereas Ghoncheh Tazmini uses Hegelian structuralist approach, that sees these revolutions as an inevitable result of modernization from above, another Iranian political scientist Zhand Shakibi puts stress on the role of human actor – pre-revolutionary monarch and his (missmanagement of the situation in explaining the origins of revolution. Shakibi applies the human agency perspective to the analysis of the character of pre-revolutionary monarch and his modus operandi. According to Shakibi the structural variables did not by themselves cause the revolution, they create only potential for revolution. The revolutions must be explained by describing complex interactions between structural variables and modus operandi of the monarchs.


    Directory of Open Access Journals (Sweden)

    Laura García Durán


    Full Text Available In this paper we shall focus on the analysis of Innocent III’s power over Philip Augustus and Alfonso IX’s marital issues. Innocent challenged these kings because he could not allow Western monarchs to maintain marital unions which could be censurable in the eyes of his subjects since they were bigamous or had consanguinity ties below the fourth degree. Innocent was well acquainted with the canonical doctrine and proved himself willing to apply it to kings regardless of the special rights of the sovereigns of France and León over whom the shadow of excommunication and interdict (seven years for the Leonese and two decades for the French king threateningly loomed; before this unexpected challenge the monarchs dared oppose Rome. This also allows us to compare and contrast the relationship between Innocent and the two monarchs.

  10. Locating non-volcanic tremor along the San Andreas Fault using a multiple array source imaging technique (United States)

    Ryberg, T.; Haberland, C.H.; Fuis, G.S.; Ellsworth, W.L.; Shelly, D.R.


    Non-volcanic tremor (NVT) has been observed at several subduction zones and at the San Andreas Fault (SAF). Tremor locations are commonly derived by cross-correlating envelope-transformed seismic traces in combination with source-scanning techniques. Recently, they have also been located by using relative relocations with master events, that is low-frequency earthquakes that are part of the tremor; locations are derived by conventional traveltime-based methods. Here we present a method to locate the sources of NVT using an imaging approach for multiple array data. The performance of the method is checked with synthetic tests and the relocation of earthquakes. We also applied the method to tremor occurring near Cholame, California. A set of small-aperture arrays (i.e. an array consisting of arrays) installed around Cholame provided the data set for this study. We observed several tremor episodes and located tremor sources in the vicinity of SAF. During individual tremor episodes, we observed a systematic change of source location, indicating rapid migration of the tremor source along SAF. ?? 2010 The Authors Geophysical Journal International ?? 2010 RAS.

  11. «The mission of the russian intelligentsia» in the works of Lev Tikhomirov

    Directory of Open Access Journals (Sweden)

    Pykharev Filipp


    Full Text Available This paper is devoted to the work of political activist and writer Lev Alexandrovich Tikhomirov. Considered and reviewed diaries, memoirs and journalism Tikhomirov, reflecting the two periods of his life: while promoting the ideology of the party «People’s Freedom» («Narodnaya Volya» and service to the monarchical idea. The author shows that the main factor that influenced the development of creativity Lev Tikhomirov as a populist, and the monarchical period, was the conviction of the special mission of the Russian intelligentsia in Russian history.

  12. Blue blood or true blood

    DEFF Research Database (Denmark)

    Møller, Fenja Søndergaard


    directly but assumed to be embedded in the monarchical regime type, and the results show that alternative covariates are unable to fully explain the monarchical peace. Moreover, the study finds that horizontal discrimination increases the risk of intrastate conflict in authoritarian republics...... but that discrimination has no effect in monarchies. Future conflict studies should therefore consider legitimacy connected to authoritarian regime types.......In the aftermath of the Arab uprisings, the difference between monarchies and republics appears more profound than ever. Aside from Bahrain, all the Middle Eastern monarchies avoided major anti-governmental protests, and no armed conflict has occurred in any of them since 1979. Inspired by Middle...

  13. Women in History--Queen Liliuokalani (United States)

    Koeppe, Tina


    This article profiles Queen Liliuokalani, Hawaii's last monarch. Liliuokalani was born in Hawaii in 1838 into the family of a high chief. She attended the Royal School, run by American missionaries, and received a high quality education and learned to love music, writing and politics. Liliuokalani was given the Christian name "Lydia" as…

  14. Valgust pimedusse / Heikki Parviainen, Tormi Soorsk

    Index Scriptorium Estoniae

    Parviainen, Heikki


    TM võrdleb LED-taskulampe: CRX 202, Duracell Daylite, Energizer Lithium Led, Fenix LD20 Premium 4, iTP SA2 Eluma, Leatherman Monarch 500, Led Lenser P6, Maxell MAFL-Q01, Mini Maglite Led, Smith&Wesson Galaxy Elite, Stanley FatMax

  15. Cultural Identity and Regional Security in the Western Balkans (United States)


    collectivist goals of humanitarian intervention.”45 Certainly, one of the most prominent representatives of this school is Barry Buzan. He has...sovereignty of the Hungarian monarch in exchange of holding some degree of local political autonomy inside Croatia. Although, some recent...

  16. Culturas y modelos políticos en la construcción del estado contemporáneo

    Directory of Open Access Journals (Sweden)

    Ángeles Lario


    Full Text Available Se analizan las culturas políticas monárquica y republicana; sus significados, sus diferencias y puntos comunes. Su proyección en América y Europa.Are monarchical and republican political cultures analyzed; his meanings, his differences and common points. His development in America and Europa.

  17. Dismantling the Perceived Barriers to the Implementation of National Higher Education Accreditation Guidelines in the Kingdom of Saudi Arabia (United States)

    Onsman, Andrys


    The Kingdom of Saudi Arabia has invested heavily in its social sector, especially in setting up new schools and universities. The aim of the development is to prepare the Kingdom for a future that is not dependent on its oil resources which are predicted to run out in less than a hundred years. Driven by the country's monarch, King Abdullah, many…

  18. Niche separation in flycatcher-like species in the lowland rainforests of Malaysia. (United States)

    Mansor, Mohammad Saiful; Ramli, Rosli


    Niche theory suggests that sympatric species reduce interspecific competition through segregation of shared resources by adopting different attack manoeuvres. However, the fact that flycatcher-like bird species exclusively use the sally manoeuvre may thus challenge this view. We studied the foraging ecology of three flycatcher-like species (i.e. Paradise-flycatcher Terpsiphone sp., Black-naped Monarch Hypothymis azurea, and Rufous-winged Philentoma Philentoma pyrhoptera) in the Krau Wildlife Reserve in central Peninsular Malaysia. We investigated foraging preferences of each bird species and the potential niche partitioning via spatial or behavioural segregation. Foraging substrate was important parameter that effectively divided paradise-flycatcher from Black-naped Monarch and Rufous-winged Philentoma, where monarch and philentoma foraged mainly on live green leaves, while paradise-flycatcher foraged on the air. They also exhibited different foraging height preferences. Paradise-flycatcher, for instance, preferred the highest studied strata, while Black-naped Monarch foraged mostly in lower strata, and Rufous-winged Philentoma made use of the lowest strata. This study indicates that niche segregation occurs among sympatric species through foraging substrate and attack manoeuvres selection. Copyright © 2017 Elsevier B.V. All rights reserved.

  19. Treating ER+ Breast Cancer with CDK4/6 Inhibitors. (United States)


    Data from the MONARCH2, PALOMA-1, and TREnd trials strongly support using CDK4/6 inhibitors alongside standard endocrine therapy for advanced ER-positive breast cancer. Including these targeted agents not only improves progression-free survival but may reverse acquired resistance to hormone treatment. ©2017 American Association for Cancer Research.

  20. Latin Hagiography in Medieval Iceland

    DEFF Research Database (Denmark)

    Jensson, Gottskálk


    I. INTRODUCTION A. Latin literacy in Iceland and the cult of saints B. Icelandic quasi-hagiography and the Christian monarchs of Norway II. THE ICELANDIC SAINTS AND THEIR LATIN TEXTS A. S. Thorlacus Scalotensis episcopus. – 1. The texts: the Latin hagiography about St Þorlákur. – 2. The historical...

  1. Was Montesquieu a Liberal Republican?

    NARCIS (Netherlands)

    de Dijn, A.


    This paper sets out to criticize Thomas Pangle's and Paul Rahe's reading of The Spirit of the Laws as a contribution to liberal republicanism, arguing instead that Montesquieu's text is better understood as a defense of liberal monarchism. Pangle's and Rahe's interpretation of The Spirit of the Laws

  2. La politique intellectuelle de Mvemba N'zinga (Dom Afonso 1er ...

    African Journals Online (AJOL)

    Le Kongo venait de rater un grand rendez-vous avec l'histoire. Mots-Clés: enseignement; élite intellectuelle; Bakongo; Mvemba N'zinga; Mbanza Kongo; Lisbonne; Mani Kongo Summary Taking advantage of the opportunities that he offered to the Portuguese monarch - consignment of missionaries, study of the young " ...

  3. The Absolutist Reformism: Projects of Political Reforms in Russia (2nd half of 18th century – 1st quarter of 19th century

    Directory of Open Access Journals (Sweden)

    Konstantin D. Bugrov


    Full Text Available The article deals with the phenomenon of absolutist reformism – a form of political culture that chronologically spans from the mid-18th century to 19th century, and is determined by both communicative context (genre, pragmatic purpose, and the social and political status of its participants, the members of court-administrative elite. The author argues that the principal reformers, who belonged to the court and administrative elite of Russian Empire, were competing with each other, and the reform proposals allowed the competitors to simultaneously improve their own positions within the structure of state governance and enact the absolute power of the monarch to bring the reform forth. However, that meant that the monarch was appearing in the reform proposals as an omnipotent arbiter capable of creating the social and political institutions by his will. Consequently, these reform proposals – starting from the early projects of the 1750es – 1760es, and finishing with the intense production of reform plans under Alexander I – were aimed at increasing the power of monarch, assuring its benevolent character, and protecting it from the potential usurpation from the inside of the bureaucratic apparatus. This logic of argumentation, which places the monarch against the bureaucracy, was to flourish later on in Russian 19th century.

  4. Clericalization of the Russian Excursion Tourism

    Directory of Open Access Journals (Sweden)

    Boris Rodoman


    Full Text Available In the modern Russia almost every secular excursion is starting to resemble a religious pilgrimage, while reducing itself to a visit to Orthodox monasteries and temples. A lot of stories are told about saints and their actions, an unconcealed religious propaganda is carried on, and the history of the country is presented in a cleric-monarchical manner.

  5. Fulltext PDF

    Indian Academy of Sciences (India)


    citizens in order to have absolute power. The “Goettinger Sieben” rebelled against this action by sending a “protest letter” to the monarch. The outcome was tragic. On 12th December 1837, Ernst August. I dismissed the seven professors and ordered three of them –. Dahlmann, J Grimm and Gervinus – to leave the country.

  6. Restoring arid western habitats: Native plants maximize wildlife conservation effectiveness (United States)

    Kas Dumroese; Jeremy Pinto; Deborah M. Finch


    Greater sage-grouse (Centrocercus urophasianus) and monarch butterflies (Danaus plexippus) and other pollinating insects have garnered a lot of attention recently from federal and state wildlife officials. These two species and pollinators share dwindling sagebrush habitat in the western United States that is putting their populations at risk. Sagebrush...

  7. Building a New Business Model (United States)

    Berkey, Lisa


    Monarch High School in Boulder, Colorado, is one of 25 schools piloting the High School of Business program, an accelerated business administration program developed by Columbus, Ohio-based MBA"Research" and Curriculum Center. This article describes the program which uses a heavily project-based pedagogy to teach a curriculum modeled…

  8. U.S. Army Staffs-Are They Broken? (United States)


    routine business that could be reduced to a few rules and thus would not require the presence of the decision maker, nmally the monarch.. 4 Information...equip=*ni, billeting, military policy arx moalle G2 - Reponsible far all inteligence fwrctiou. G3 - IRmsP l zuible for cpmratiros to inludezm zmrtu of

  9. Leadership for the church: The shepherd model

    African Journals Online (AJOL)


    May 30, 2014 ... also monarchical leaders (2 Sm 5:2) and God himself (Is 40:11; Ps 23:1). The prophetic ... referred to are Abraham (Gn 12:16); Rachel (Gn 29:9); Jacob ... these functions are contrary to the shepherds of Ezekiel's era, ..... Volf, M., 1998, After our likeness: The church as the image of the trinity, Wm. B.

  10. 77 FR 9657 - Ocean Transportation Intermediary License Applicants (United States)


    ..., Owner, Application Type: Add OFF Service. Cargo Tours International, Inc. dba CTI Global Logistics (NVO.... Deckwell Sky (USA) Inc. dba Monarch Container Line (NVO & OFF), 14343 E. Don Julian Road, City of Industry.... Logistics, Inc. (NVO & OFF), 3905 West Albany Street, McHenry, IL 60050, Officers: Arthur N. Nutig, Chief...

  11. Raising Butterflies from Your Own Garden. (United States)

    Howley-Pfeifer, Patricia


    Describes how raising monarch, black swallowtail, and mourning cloak butterflies in a kindergarten class garden can provide opportunities for observation experiences. Includes detailed steps for instruction and describes stages of growth. Excerpts children's journal dictations to illustrate ways to support the discovery process. Describes related…

  12. Laat de kindertjes tot mij komen... Franse vorsten geportretteerd op een onbekende miniatuur

    NARCIS (Netherlands)

    Manuth, V.; Leeuwen, R. van


    Let the Children come unto Me: French monarchs portrayed on an unknown miniature This article identifies eleven portraits of the French kings and queens from the reigns of Henry II to Henry IV in a miniature depicting Christ Suffering the Little Children To Come unto Him. It was painted by an

  13. Native plant development and deployment [Section VII (United States)

    Jessica Wright; Kas Dumroese; Amy Symstad; Theresa Pitts-Singer; Jim Cane; Gary Krupnick; Peggy Olwell; Byron Love; Elizabeth Sellers; John Englert; Troy Wood


    Native plant materials are needed to create, enhance, or restore pollinator habitat. They provide critical foraging and breeding areas for wild and managed pollinator species, including transnational migratory species such as hummingbirds and monarch butterflies. Although many pollinators and plants are generalists, some have limited, obligate relationships (i.e., one...

  14. Delisting Process for Endangered Species and Relevance to Populations on Army Lands (United States)


    09/21/2004 Tinian monarch Recovered 11/11/1977 11/22/1983 Treefrog, pine barrens (FL pop.) New information discovered 09/13/1996 04/26/2000...Sand Association et al. 10-25-2001 Delist Positive (68 FR 52784) Plymouth redbelly turtle National Wilderness Institute 02-03-1997 Delist NOT

  15. Precise tremor source locations and amplitude variations along the lower-crustal central San Andreas Fault (United States)

    Shelly, David R.; Hardebeck, Jeanne L.


    We precisely locate 88 tremor families along the central San Andreas Fault using a 3D velocity model and numerous P and S wave arrival times estimated from seismogram stacks of up to 400 events per tremor family. Maximum tremor amplitudes vary along the fault by at least a factor of 7, with by far the strongest sources along a 25 km section of the fault southeast of Parkfield. We also identify many weaker tremor families, which have largely escaped prior detection. Together, these sources extend 150 km along the fault, beneath creeping, transitional, and locked sections of the upper crustal fault. Depths are mostly between 18 and 28 km, in the lower crust. Epicenters are concentrated within 3 km of the surface trace, implying a nearly vertical fault. A prominent gap in detectible activity is located directly beneath the region of maximum slip in the 2004 magnitude 6.0 Parkfield earthquake.

  16. Multistation magnetotellurics. Final report, 1 January 1996--30 June 1997

    Energy Technology Data Exchange (ETDEWEB)

    Egbert, G.D.


    The author has developed the foundations of a practical multivariate approach to processing magnetotelluric array data. Compared to current standards for magnetotelluric data processing, the multivariate approach is unique in that all available data channels are used simultaneously. The approach is outlined in this report. Using Multmtrn, a program for multiple station analysis of magnetotelluric data, the author achieved significant improvements in apparent resistivity and phase estimates in initial tests. Examples of the use of this approach are given including: Carrizo Plain and Parkfield electromagnetic profiling data; sea floor magnetotelluric (MT) data from the Gulf of Mexico; MT survey in a culturally noisy area of Bavaria; and Parkfield/Hollister earthquake monitoring array data. Experience with these projects has resulted in an improved program. The new version of the code is available at or by contacting egbert{at} Appendices of this report present documentation for Multmtrn.

  17. A modification scheme for seismic acceleration - time histories

    International Nuclear Information System (INIS)

    Bethell, J.


    A technique is described for the modification of recorded earthquake acceleration-time histories which gives reduced peak accelerations whilst leaving other significant characteristics unchanged. Such modifications are of use in constructing design basis acceleration-time histories such that all important parameters conform to a specified return period. The technique is applied to two recordings from the 1966 Parkfield earthquake, their peak accelerations being reduced in each case from about 40% g to 25% g. (author)

  18. Quantifying slip balance in the earthquake cycle: Coseismic slip model constrained by interseismic coupling

    KAUST Repository

    Wang, Lifeng


    The long-term slip on faults has to follow, on average, the plate motion, while slip deficit is accumulated over shorter time scales (e.g., between the large earthquakes). Accumulated slip deficits eventually have to be released by earthquakes and aseismic processes. In this study, we propose a new inversion approach for coseismic slip, taking interseismic slip deficit as prior information. We assume a linear correlation between coseismic slip and interseismic slip deficit, and invert for the coefficients that link the coseismic displacements to the required strain accumulation time and seismic release level of the earthquake. We apply our approach to the 2011 M9 Tohoku-Oki earthquake and the 2004 M6 Parkfield earthquake. Under the assumption that the largest slip almost fully releases the local strain (as indicated by borehole measurements, Lin et al., 2013), our results suggest that the strain accumulated along the Tohoku-Oki earthquake segment has been almost fully released during the 2011 M9 rupture. The remaining slip deficit can be attributed to the postseismic processes. Similar conclusions can be drawn for the 2004 M6 Parkfield earthquake. We also estimate the required time of strain accumulation for the 2004 M6 Parkfield earthquake to be ~25 years (confidence interval of [17, 43] years), consistent with the observed average recurrence time of ~22 years for M6 earthquakes in Parkfield. For the Tohoku-Oki earthquake, we estimate the recurrence time of~500-700 years. This new inversion approach for evaluating slip balance can be generally applied to any earthquake for which dense geodetic measurements are available.

  19. Quantifying slip balance in the earthquake cycle: Coseismic slip model constrained by interseismic coupling

    KAUST Repository

    Wang, Lifeng; Hainzl, Sebastian; Mai, Paul Martin


    The long-term slip on faults has to follow, on average, the plate motion, while slip deficit is accumulated over shorter time scales (e.g., between the large earthquakes). Accumulated slip deficits eventually have to be released by earthquakes and aseismic processes. In this study, we propose a new inversion approach for coseismic slip, taking interseismic slip deficit as prior information. We assume a linear correlation between coseismic slip and interseismic slip deficit, and invert for the coefficients that link the coseismic displacements to the required strain accumulation time and seismic release level of the earthquake. We apply our approach to the 2011 M9 Tohoku-Oki earthquake and the 2004 M6 Parkfield earthquake. Under the assumption that the largest slip almost fully releases the local strain (as indicated by borehole measurements, Lin et al., 2013), our results suggest that the strain accumulated along the Tohoku-Oki earthquake segment has been almost fully released during the 2011 M9 rupture. The remaining slip deficit can be attributed to the postseismic processes. Similar conclusions can be drawn for the 2004 M6 Parkfield earthquake. We also estimate the required time of strain accumulation for the 2004 M6 Parkfield earthquake to be ~25 years (confidence interval of [17, 43] years), consistent with the observed average recurrence time of ~22 years for M6 earthquakes in Parkfield. For the Tohoku-Oki earthquake, we estimate the recurrence time of~500-700 years. This new inversion approach for evaluating slip balance can be generally applied to any earthquake for which dense geodetic measurements are available.

  20. The morphology of strike-slip faults - Examples from the San Andreas Fault, California (United States)

    Bilham, Roger; King, Geoffrey


    The dilatational strains associated with vertical faults embedded in a horizontal plate are examined in the framework of fault kinematics and simple displacement boundary conditions. Using boundary element methods, a sequence of examples of dilatational strain fields associated with commonly occurring strike-slip fault zone features (bends, offsets, finite rupture lengths, and nonuniform slip distributions) is derived. The combinations of these strain fields are then used to examine the Parkfield region of the San Andreas fault system in central California.


    Directory of Open Access Journals (Sweden)

    Hristina Nikolic


    Full Text Available This article mainly deals with the Plumpungan manuscript discourse, an ancient stone inscription found in the Salatiga area. The manuscript dates back to the 8th century and is one of the main historical sources of the Salatiga municipality. There is a tight connection involving the above mentioned inscription with the legitimacy of the Javanese court at that time, as well as with the monarch, who was seen as half- man, half-deity. The monarch drew on the labor of his subjects in maintaining reli- gious sites to ensure his place on earth, and in heaven. The Plumpungan manuscript was a ‘legal document’ used to reassure inhabitants of the area that the monarch is the legitimate ruler, and to prevent further revolt. The monarch’s subjects in the Salatiga area at the time were farmers disheartened with high taxes and the fear of volcanic eruptions, which later caused great migrations to East Java. Consequently, the mon- arch, using the Plumpungan manuscript as a medium, decreed that the Hampra vil- lage (present-day Salatiga become a tax-free area due to the excellent care that its citizens provided for the religious sites, in addition to the obeisance shown to the monarch. It becomes clear, however, that all the way through the discourse analysis, the king wanted to ensure his legitimacy. Socio-historical context confirms that the monarch, Bhanu, was a successful ruler who held power over four regions, analogi- cally with Indra, the king of all deities. Discourse analysis is detained under three main dimensions, which is grammatical, lexical and contextual, unified from be- ginning to end in a particular style that reveals genuinely hidden meaning.     Key words: manuscript, Plumpungan, kodikologi, dan analisis stilistika.

  2. Heterogeneous slip and rupture models of the San Andreas fault zone based upon three-dimensional earthquake tomography

    Energy Technology Data Exchange (ETDEWEB)

    Foxall, William [Univ. of California, Berkeley, CA (United States)


    Crystal fault zones exhibit spatially heterogeneous slip behavior at all scales, slip being partitioned between stable frictional sliding, or fault creep, and unstable earthquake rupture. An understanding the mechanisms underlying slip segmentation is fundamental to research into fault dynamics and the physics of earthquake generation. This thesis investigates the influence that large-scale along-strike heterogeneity in fault zone lithology has on slip segmentation. Large-scale transitions from the stable block sliding of the Central 4D Creeping Section of the San Andreas, fault to the locked 1906 and 1857 earthquake segments takes place along the Loma Prieta and Parkfield sections of the fault, respectively, the transitions being accomplished in part by the generation of earthquakes in the magnitude range 6 (Parkfield) to 7 (Loma Prieta). Information on sub-surface lithology interpreted from the Loma Prieta and Parkfield three-dimensional crustal velocity models computed by Michelini (1991) is integrated with information on slip behavior provided by the distributions of earthquakes located using, the three-dimensional models and by surface creep data to study the relationships between large-scale lithological heterogeneity and slip segmentation along these two sections of the fault zone.

  3. "Courageous, Zealous, Learned, Wise, and Chaste" - Queen Elizabeth I's Biblical Analogies After Her Death

    Directory of Open Access Journals (Sweden)

    Aidan Norrie


    Full Text Available During her reign, Queen Elizabeth I of England was the subject of various biblical analogies. Much of the current historiography, however, does not continue analysis of these literary devices after the Queen's death in 1603. Primary source documents show that their use did not cease with the Queen's death. These analogies appear to have continued for two primary purposes. Analogies legitimised a questionable event that had occurred during Elizabeth's reign, and portrayed these decisions as an example for the current Protestant monarch to imitate. Also, in the years after the Queen's death, analogies reinforced England's Protestantism (and its divine sanction, and gave the Stuart monarchs an example to emulate in religio-political matters.

  4. Working through a psychotherapy group's political cultures. (United States)

    Ettin, Mark F; Cohen, Bertram D


    Macropolitical evolution, starting with authoritarian monarchism, has moved through anarchistic transitions either to the totalitarianism of fascism and communism or to liberal and social democracy. We posit analogous micropolitical development in process-oriented therapy groups: "dependence" and "counterdependence" corresponding to monarchism and anarchism; and "independence" and "interdependence" to liberal and social democracy, respectively. Transition from counterdependence to independence and interdependence may be: (1) facilitated through group members' cooperative experience of rebellion, or (2) blocked by collective identification, the internalization of dystopian or utopian fantasies that coalesce as "group-self" perceptions. We explore how group therapists work clinically with and through these several "political cultures" in the service of group and self transformation.

  5. The Return of the Loving Father: Masculinity, Legitimacy and the French and Dutch Restoration

    Directory of Open Access Journals (Sweden)

    Matthijs Lok


    Full Text Available Historians of gender often see the construction of hegemonic images of masculinity as the result of long-term cultural processes. In this article we investigate the influence of short-term political events on the shaping of dominant political masculinities by comparing the representations of the early French and Dutch Restoration monarchies. The events of the political transition of 1813-1815 greatly influenced the competition of different models of masculinity existing in the early nineteenth century. In both countries the newly established monarchs aimed to legitimate their insecure rule by presenting themselves as 'loving fathers' returning to their despairing children after the dark years of exile. The Dutch monarchy differed from the French case with regards to the role of women in the monarchical representation and the duality of the representation of William I as father and hero. Unlike Louis XVIII, William could present his fatherly rule as a return to the national tradition of domesticity (huiselijkheid.

  6. The Romanian Social Democratic Party versus the authoritarian monarchy

    Directory of Open Access Journals (Sweden)

    Florin Grecu


    Full Text Available The article approaches the Romanian social-democratic collaboration during 1938-1940 with the authoritarian monarch regime. Even though the party leaders had diverging political views, regards to the acceptance or the non-acceptance of the authoritarian regime, the influential PSDR members held leading positions within the single party and the corporate parliament and within the union structures. The positions were offered by the regime, so that the union leaders would stop instigating workers to go on strike, and to accept the governmental policies. The freedom of speech and the political actions were ceded to the monarch, who governed at the place of the political parties and he controlled the unions, by using the guilds.

  7. The Shifting Frontier: The Achaemenid Empire's Treatment of Western Colonies

    Directory of Open Access Journals (Sweden)

    Jon L. Berquist


    Full Text Available Until recently, most formulations of ancient Israel's history within the biblical time-frame separated the time-line into four broad segments: pre-monarchic (also called patriarchal, monarchic, exilic, and postexilic. This outline allowed the construction of many major interpretations, based upon presumed differences between these periods. Newer presentations of that history, however, have called into question many parts of this reconstruction. Other terms are more descriptive than the appellation "postexilic," which has two chief drawbacks. The first is that it is open-ended; the last 25 centuries have been after the exile, and so will the centuries. The second is that "postexilic" defines the period in terms of its predecessor, and it is not surprising that much scholarship of this period has been reductionistic.

  8. A Well Regulated Militia Political and Military Organisation in Pre-Napoleonic Switzerland (1550-1799

    Directory of Open Access Journals (Sweden)

    Gassmann Jürg


    Full Text Available The period sees the transition of the ordinary fighter from feudal levy, yeoman or city burgher militia, to subject in an absolute polity, to today’s concept of the free citizen in a democratic state. In the period, the Swiss Confederacy was the only major polity that was not monarchical, but republican, and at the same time eschewed a standing army in favour of continued reliance on militia throughout.

  9. Style and elegance in Bessarabian photographs of the 2nd half of the 19th century


    Mihail Dohot


    The author presents the fashion trends of Western Europe, Romania and Bessarabia in the second half of the 19th century through the prism of photography. The influence of European monarchic houses on the fashion of that time is considered, as well as the role of artistic and cultural emancipation, which has left its imprint on society through the visual arts. The article lists fashion designers who contributed to the development of fashion and whose work was reflected in the photographs.

  10. Beneficial Insect Attraction to Milkweeds (Asclepias speciosa, Asclepias fascicularis) in Washington State, USA


    David G. James; Lorraine Seymour; Gerry Lauby; Katie Buckley


    Native plant and beneficial insect associations are relatively unstudied yet are important in native habitat restoration programs for improving and sustaining conservation biological control of arthropod pests in agricultural crops. Milkweeds (Asclepias spp.) are currently the focus of restoration programs in the USA aimed at reversing a decline in populations of the milkweed-dependent monarch butterfly (Danaus plexippus); however, little is known of the benefits of these plants to other bene...

  11. The chemistry of antipredator defense by secondary compounds in neotropical lepidoptera: facts, perspectives and caveats


    Trigo, José R.


    Chemical defense against predation in butterflies and moths has been studied since nineteenth century. A classical example is that of the larvae of the monarch butterfly Danaus plexippus, which feed on leaves of Asclepias curassavica (Asclepiadaceae), sequestering cardenolides. The adults are protected against predation by birds. Several other substances may be involved in chemical defense, such as iridoid glycosides, cyanogenic glycosides, glucosinolates, pyrrolizidine and tropane alkaloids,...

  12. Information Processing and Collective Behavior in a Model Neuronal System (United States)


    were invited to visit the 711th Human Performance Wing at Wright Patterson AFB to give a briefing on our work. We also presented our work at AFRL in...for an AFOSR project headed by Steve Reppert on Monarch Butterfly navigation. We visited the Reppert lab at the UMASS Medical School and have had many...the requested briefing at the 711th Human Performance Wing . It is a natural extension of the original aims of the grant, as the suprachiasmatic

  13. Structural colours of nickel bioreplicas of butterfly wings (United States)

    Tolenis, Tomas; Swiontek, Stephen E.; Lakhtakia, Akhlesh


    The two-angle conformally evaporated-film-by-rotation technique (TA-CEFR) was devised to coat the wings of the monarch butterfly with nickel in order to form a 500-nm thick bioreplica thereof. The bioreplica exhibits structural colours that are completely obscured in actual wings by pigmental colours. Thus, the TA-CEFR technique provides a way to replicate, study and exploit hidden morphologies of biological surfaces.

  14. "Identity Monarchy": Interrogating Heritage for a Divided Malaysia


    Milner, Anthony


    Malaysia, it has been observed, is currently experiencing a "revival" of "Malay kingship" with the growing importance of "proactive and participating constitutional rulers." In fact, modern Malaysia has since independence been characterized by monarchy--by a multiplicity of Rulers and elaborate royal ceremony and hierarchy --as well as by its "plural society." But the modern monarchs--though they have never become quite "constitutional Rulers"--cannot be seen as merely "traditional," because ...

  15. East Europe Report, Political, Sociological and Military Affairs, No. 2199. (United States)


    countries. Had the libel been true, these monarchs obviously would not have tolerated the Jesuits in their countries. In 1814, Pope Pius VII re- stored the...with Pope John Paul II during his first pilgrimage to Poland requires a separate discussion. The growth in this form of manifestation of the other side of the argument. It is true that in 1773 Pope Clement XIV dissolved the Jesuit Order, but he did so under pressure from the

  16. Modélisation des propriétés diélectriques des nanocomposites ...

    African Journals Online (AJOL)

    Dans ce travail, nous nous intéressons aux propriétés diélectriques dans le domaine des micro-ondes (2.45, 9.50 et 35 GHz) des composites constitués des inclusions de noir de carbone dans une matrice résine époxyde. Nous avons utilisé deux types de carbone : le monarch 700 et le sterling, dont les diamètres des ...

  17. Contribución geográfica al programa integral de desarrollo mariposa monarca

    Directory of Open Access Journals (Sweden)

    Carlos Melo Gallegos


    Full Text Available The present study deals with the ecological and socioeconomic problems that constitute the conservation, management and deve­lopment of the Ecological Reservation "Mariposa Monarca" (Monarch Butterfly at regional level, including geographic basis to the ordaining and planification of the environment and its natural resources. An ecogeographic outline is established so as to help and guide norm–operative actions of the Programa Integral de De­sarrollo (Integral Programme of Development set by SEDUE in that reservation.

  18. Princes and Merchants: European City Growth before the Industrial Revolution.


    De Long, J Bradford; Shleifer, Andrei


    As measured by the pace of city growth in western Europe from 1000 to 1800. absolutist monarchs stunted the growth of commerce and industry. A region ruled by an absolutist prince saw its total urban population shrink by one hundred thousand people per century relative to a region without absolutist government. This might be explained by higher rates of taxation under revenue-maximizing absolutist governments than under non-absolutist governments. which care more about general economic prospe...

  19. America First Isolationism in U.S. Foreign Policy from the 19th to the 21st Century (United States)


    his third diplomatic mission to Europe to offer mediation to the belligerent parties, but, as Link points out, the British and French leaders were...represented the German people and “not the military masters and the monarchical autocrats ”; Germany accepted the conditions in their entirety on October...the Republican leader of the Senate, Henry Cabot Lodge, generally opposed any democratic attempts and personally disliked Wilson.145 Duroselle

  20. Proměny dětství a vzdělávání v Českých zemích


    Musil, Jan


    The diploma work is concetrating on childhood development and it's dependency on spreading and improving in educational system. The work would give an interpretation how education had influenced the period of childhood in the past in Czech countries. What effect had education on childhood duration and quality. The work has chronological system from the beginning of education in Czech countries (monarch St. Václav) to the end of rule Joseph II. Individual chapter give interpretations about edu...

  1. Temple Wars: Cambodia’s Dispute Over Preah Vihear Ownership and its Effects on National Power (United States)


    patronage of Buddhism and aspects of kingship that led to territorial expansion. Suryavarman was a unifying monarch that expanded the Khmer Empire Buddhism may be the reason why elements of the religion are found in the temple’s architecture. Likewise, the thirteenth century brought about a...decline of Hindu worship in the Khmer Empire and the Preah Vihear Temple was then dedicated to Buddhism . 23 Cambodia’s Geography and its

  2. Area Handbook Series. Sri Lanka, A Country Study (United States)


    power- ful Indian monarch, Asoka , nurtured the new comprehensive religio-philosophical system in the third century B.C. Asoka’s con- version to...right collarbone and his revered alms bowl from Asoka and to have built the Thuparama Dagoba, or stupa (Buddhist shrine), to honor these highly revered...Association. The state has similarly retained close ties with the sangha. Since the time of Asoka , the first great Indian emperor (third century B.C.), the

  3. Food plant derived disease tolerance and resistance in a natural butterfly-plant-parasite interactions. (United States)

    Sternberg, Eleanore D; Lefèvre, Thierry; Li, James; de Castillejo, Carlos Lopez Fernandez; Li, Hui; Hunter, Mark D; de Roode, Jacobus C


    Organisms can protect themselves against parasite-induced fitness costs through resistance or tolerance. Resistance includes mechanisms that prevent infection or limit parasite growth while tolerance alleviates the fitness costs from parasitism without limiting infection. Although tolerance and resistance affect host-parasite coevolution in fundamentally different ways, tolerance has often been ignored in animal-parasite systems. Where it has been studied, tolerance has been assumed to be a genetic mechanism, unaffected by the host environment. Here we studied the effects of host ecology on tolerance and resistance to infection by rearing monarch butterflies on 12 different species of milkweed food plants and infecting them with a naturally occurring protozoan parasite. Our results show that monarch butterflies experience different levels of tolerance to parasitism depending on the species of milkweed that they feed on, with some species providing over twofold greater tolerance than other milkweed species. Resistance was also affected by milkweed species, but there was no relationship between milkweed-conferred resistance and tolerance. Chemical analysis suggests that infected monarchs obtain highest fitness when reared on milkweeds with an intermediate concentration, diversity, and polarity of toxic secondary plant chemicals known as cardenolides. Our results demonstrate that environmental factors-such as interacting species in ecological food webs-are important drivers of disease tolerance. © 2012 The Author(s). Evolution© 2012 The Society for the Study of Evolution.

  4. To The Historical Portrait of Theophan Prokopovych

    Directory of Open Access Journals (Sweden)

    А. І. Makhinko


    Full Text Available The article highlights the stages of life Theophan Prokopovych, an outstanding theologian, philosopher, teacher and writer. Its artistic heritage became the property of the Russian and Ukrainian cultures. Theophan Prokopovych life is divided into two periods, which differ from each other by the nature of its activities. In the first period (1687–1715 years he studies and then engaged in research and teaching activities in the Kiev Academy. In the second period (1715–1736 years it is at the request of Peter I moved to St. Petersburg, where he became the archbishop and is engaged in administrative and social activities / During this time he wrote a treatise «Spiritual regulations» and «The Truth will monarch», which became the conceptual basics of church reform and the reform of succession to the throne. As a result of the reform of the Church patriarchate was abolished and the church became subordinated to the state. This similarity can be traced with some Protestant countries of Europe, where the head of state led the church. «The truth will monarch» to overturn the traditional order of succession to the throne in the male downlink and gave the right to appoint a monarch testament heir. In these treatises have been realized his philosophical views on the state and its institutions.

  5. Disease ecology across soil boundaries: effects of below-ground fungi on above-ground host-parasite interactions. (United States)

    Tao, Leiling; Gowler, Camden D; Ahmad, Aamina; Hunter, Mark D; de Roode, Jacobus C


    Host-parasite interactions are subject to strong trait-mediated indirect effects from other species. However, it remains unexplored whether such indirect effects may occur across soil boundaries and connect spatially isolated organisms. Here, we demonstrate that, by changing plant (milkweed Asclepias sp.) traits, arbuscular mycorrhizal fungi (AMF) significantly affect interactions between a herbivore (the monarch butterfly Danaus plexippus) and its protozoan parasite (Ophryocystis elektroscirrha), which represents an interaction across four biological kingdoms. In our experiment, AMF affected parasite virulence, host resistance and host tolerance to the parasite. These effects were dependent on both the density of AMF and the identity of milkweed species: AMF indirectly increased disease in monarchs reared on some species, while alleviating disease in monarchs reared on other species. The species-specificity was driven largely by the effects of AMF on both plant primary (phosphorus) and secondary (cardenolides; toxins in milkweeds) traits. Our study demonstrates that trait-mediated indirect effects in disease ecology are extensive, such that below-ground interactions between AMF and plant roots can alter host-parasite interactions above ground. In general, soil biota may play an underappreciated role in the ecology of many terrestrial host-parasite systems. © 2015 The Author(s).

  6. Disease ecology across soil boundaries: effects of below-ground fungi on above-ground host–parasite interactions (United States)

    Tao, Leiling; Gowler, Camden D.; Ahmad, Aamina; Hunter, Mark D.; de Roode, Jacobus C.


    Host–parasite interactions are subject to strong trait-mediated indirect effects from other species. However, it remains unexplored whether such indirect effects may occur across soil boundaries and connect spatially isolated organisms. Here, we demonstrate that, by changing plant (milkweed Asclepias sp.) traits, arbuscular mycorrhizal fungi (AMF) significantly affect interactions between a herbivore (the monarch butterfly Danaus plexippus) and its protozoan parasite (Ophryocystis elektroscirrha), which represents an interaction across four biological kingdoms. In our experiment, AMF affected parasite virulence, host resistance and host tolerance to the parasite. These effects were dependent on both the density of AMF and the identity of milkweed species: AMF indirectly increased disease in monarchs reared on some species, while alleviating disease in monarchs reared on other species. The species-specificity was driven largely by the effects of AMF on both plant primary (phosphorus) and secondary (cardenolides; toxins in milkweeds) traits. Our study demonstrates that trait-mediated indirect effects in disease ecology are extensive, such that below-ground interactions between AMF and plant roots can alter host–parasite interactions above ground. In general, soil biota may play an underappreciated role in the ecology of many terrestrial host–parasite systems. PMID:26468247

  7. Genetic variation in plant volatile emission does not result in differential attraction of natural enemies in the field. (United States)

    Wason, Elizabeth L; Hunter, Mark D


    Volatile organic chemical (VOC) emission by plants may serve as an adaptive plant defense by attracting the natural enemies of herbivores. For plant VOC emission to evolve as an adaptive defense, plants must show genetic variability for the trait. To date, such variability has been investigated primarily in agricultural systems, yet relatively little is known about genetic variation in VOCs emitted by natural populations of native plants. Here, we investigate intraspecific variation in constitutive and herbivore-induced plant VOC emission using the native common milkweed plant (Asclepias syriaca) and its monarch caterpillar herbivore (Danaus plexippus) in complementary field and common garden greenhouse experiments. In addition, we used a common garden field experiment to gauge natural enemy attraction to milkweed VOCs induced by monarch damage. We found evidence of genetic variation in the total constitutive and induced concentrations of VOCs and the composition of VOC blends emitted by milkweed plants. However, all milkweed genotypes responded similarly to induction by monarchs in terms of their relative change in VOC concentration and blend. Natural enemies attacked decoy caterpillars more frequently on damaged than on undamaged milkweed, and natural enemy visitation was associated with higher total VOC concentrations and with VOC blend. Thus, we present evidence that induced VOCs emitted by milkweed may function as a defense against herbivores. However, plant genotypes were equally attractive to natural enemies. Although milkweed genotypes diverge phenotypically in their VOC concentrations and blends, they converge into similar phenotypes with regard to magnitude of induction and enemy attraction.

  8. The frequency dependence of friction in experiment, theory, and observations of low frequency earthquakes (United States)

    Thomas, A.; Beeler, N. M.; Burgmann, R.; Shelly, D. R.


    Low frequency earthquakes (LFEs) are small amplitude, short duration events composing tectonic tremor, probably generated by shear slip on asperities downdip of the seismogenic zone. In Parkfield, Shelly and Hardebeck [2010] have identified 88 LFE families, or hypocentral locations, that contain over half a million LFEs since 2001 on a 160-km-long section of the San Andreas fault between 16 and 30 km depth. A number of studies have demonstrated the extreme sensitivity of low frequency earthquakes (LFEs) near Parkfield to stress changes ranging from contingent upon the amplitude and frequency content of the applied stress. We attempt to test this framework by comparing observations of LFEs triggered in response to stresses spanning several orders of magnitude in both frequency and amplitude (e.g. tides, teleseismic surface waves, static stress changes, etc.) to the predicted response of a single degree of freedom slider block model with rate and state dependent strength. The sensitivity of failure time in the friction model as developed in previous studies does not distinguish between shear and normal stresses; laboratory experiments show a more complicated sensitivity of failure time to normal stress change than in the published model. Because the shear and normal tidal stresses at Parkfield have different amplitudes and are not in phase, we have modified the model to include the expected sensitivity to normal stress. Our prior investigations of the response of both regular and low frequency earthquakes to tidal stresses [Thomas et al., 2009; Shelly and Johnson, 2011] are qualitatively consistent with the predictions of the friction model , as both the timing and degree (probability) of correlation are in agreement.

  9. Toward Expanding Tremor Observations in the Northern San Andreas Fault System in the 1990s (United States)

    Damiao, L. G.; Dreger, D. S.; Nadeau, R. M.; Taira, T.; Guilhem, A.; Luna, B.; Zhang, H.


    The connection between tremor activity and active fault processes continues to expand our understanding of deep fault zone properties and deformation, the tectonic process, and the relationship of tremor to the occurrence of larger earthquakes. Compared to tremors in subduction zones, known tremor signals in California are ~5 to ~10 smaller in amplitude and duration. These characteristics, in addition to scarce geographic coverage, lack of continuous data (e.g., before mid-2001 at Parkfield), and absence of instrumentation sensitive enough to monitor these events have stifled tremor detection. The continuous monitoring of these events over a relatively short time period in limited locations may lead to a parochial view of the tremor phenomena and its relationship to fault, tectonic, and earthquake processes. To help overcome this, we have embarked on a project to expand the geographic and temporal scope of tremor observation along the Northern SAF system using available continuous seismic recordings from a broad array of 100s of surface seismic stations from multiple seismic networks. Available data for most of these stations also extends back into the mid-1990s. Processing and analysis of tremor signal from this large and low signal-to-noise dataset requires a heavily automated, data-science type approach and specialized techniques for identifying and extracting reliable data. We report here on the automated, envelope based methodology we have developed. We finally compare our catalog results with pre-existing tremor catalogs in the Parkfield area.

  10. Scientific drilling into the San Andreas fault and site characterization research: Planning and coordination efforts. Final technical report

    Energy Technology Data Exchange (ETDEWEB)

    Zoback, M.D.


    The fundamental scientific issue addressed in this proposal, obtaining an improved understanding of the physical and chemical processes responsible for earthquakes along major fault zones, is clearly of global scientific interest. By sampling the San Andreas fault zone and making direct measurements of fault zone properties to 4.0 km at Parkfield they will be studying an active plate-boundary fault at a depth where aseismic creep and small earthquakes occur and where a number of the scientific questions associated with deeper fault zone drilling can begin to be addressed. Also, the technological challenges associated with drilling, coring, downhole measurements and borehole instrumentation that may eventually have to be faced in deeper drilling can first be addressed at moderate depth and temperature in the Parkfield hole. Throughout the planning process leading to the development of this proposal they have invited participation by scientists from around the world. As a result, the workshops and meetings they have held for this project have involved about 350 scientists and engineers from about a dozen countries.

  11. Episodic radon changes in subsurface soil gas along active faults and possible relation to earthquakes

    International Nuclear Information System (INIS)

    King, C.


    Subsurface soil gas along active faults in central California has been continuously monitored by the Track Etch method to test whether its radon-isotope content may show any premonitory changes useful for earthquake prediction. The monitoring network was installed in May 1975 and has since been gradually expanded to consist of more than 60 stations along a 380-km section of the San Andreas fault system between Santa Rosa and Cholame. This network has recorded several episodes, each lasting several weeks to several months, during which the radon concentration increased by a factor of approximately 2 above average along some long, but limited, fault segments (approx.100 km). These episodes occurred in different seasons and do not appear to be systematically related to changes in meteorological conditions. However, they coincided reasonably well in time and space with larger local earthquakes above a threshold magnitude of about 4.0. These episodic radon changes may be caused by a changing outgassing rate in the fault zones in response to some episodic strain changes, which incidentally caused the earthquakes

  12. Trans-generational parasite protection associated with paternal diet. (United States)

    Sternberg, Eleanore D; de Roode, Jacobus C; Hunter, Mark D


    Multiple generations of hosts are often exposed to the same pathogens, favouring the evolution of trans-generational defences. Because females have more opportunities to transfer protective molecules to offspring, many studies have focused on maternally derived protection. However, males of many species can transfer compounds along with sperm, including chemicals that could provide protection. Here, we assess maternally and paternally derived protection in a monarch butterfly-protozoan parasite system where parasite resistance is heavily influenced by secondary plant chemicals, known as cardenolides, present in the larval diet of milkweed plants. We reared monarch butterflies on medicinal and non-medicinal milkweed species and then measured resistance of their offspring to infection. We also measured cardenolide content in adult monarchs reared on the two species, and in the eggs that they produced. We found that offspring were more resistant to infection when their fathers were reared on medicinal milkweed, while maternal diet had less of an effect. We also found that eggs contained the highest levels of cardenolides when both parents were reared on the medicinal species. Moreover, females reared on non-medicinal milkweed produced eggs with significantly higher levels of cardenolides if they mated with males reared on the medicinal milkweed species. However, we found an equivocal relationship between the cardenolides present in eggs and parasite resistance in the offspring. Our results demonstrate that males reared on medicinal plants can transfer protection to their offspring, but the exact mechanism remains unresolved. This suggests that paternal protection from parasitism might be important, particularly when there are environmental sources of parasite resistance and when males transfer spermatophores during mating. © 2014 The Authors. Journal of Animal Ecology © 2014 British Ecological Society.

  13. Environmental Persistence Influences Infection Dynamics for a Butterfly Pathogen.

    Directory of Open Access Journals (Sweden)

    Dara A Satterfield

    Full Text Available Many pathogens, including those infecting insects, are transmitted via dormant stages shed into the environment, where they must persist until encountering a susceptible host. Understanding how abiotic conditions influence environmental persistence and how these factors influence pathogen spread are crucial for predicting patterns of infection risk. Here, we explored the consequences of environmental transmission for infection dynamics of a debilitating protozoan parasite (Ophryocystis elektroscirrha that infects monarch butterflies (Danaus plexippus. We first conducted an experiment to observe the persistence of protozoan spores exposed to natural conditions. Experimental results showed that, contrary to our expectations, pathogen doses maintained high infectivity even after 16 days in the environment, although pathogens did yield infections with lower parasite loads after environmental exposure. Because pathogen longevity exceeded the time span of our experiment, we developed a mechanistic model to better explore environmental persistence for this host-pathogen system. Model analysis showed that, in general, longer spore persistence led to higher infection prevalence and slightly smaller monarch population sizes. The model indicated that typical parasite doses shed onto milkweed plants must remain viable for a minimum of 3 weeks for prevalence to increase during the summer-breeding season, and for 11 weeks or longer to match levels of infection commonly reported from the wild, assuming moderate values for parasite shedding rate. Our findings showed that transmission stages of this butterfly pathogen are long-lived and indicated that this is a necessary condition for the protozoan to persist in local monarch populations. This study provides a modeling framework for future work examining the dynamics of an ecologically important pathogen in an iconic insect.

  14. Disc-retained tubes for radiologically inserted gastrostomy (RIG): Not up to the job?

    International Nuclear Information System (INIS)

    Kibriya, N.; Wilbraham, L.; Mullan, D.; Puro, P.; Vasileuskaya, S.; Edwards, D.W.; Laasch, H.-U.


    Aim: To assess the insertion procedure and performance of disc-retained gastrostomy tubes, recording complications and accidental displacements by prospective audit, and to determine whether primary placement of the tube off-licence was feasible. Materials and methods: Disc-retained 12 F single-lumen Monarch gastrostomy tubes (Enteral UK, Selby, UK) were inserted by three gastrointestinal interventional radiologists in a supra-regional cancer centre. The 12 F tubes required a 20 F peel-away sheath with four-point gastropexy fixation and were placed under conscious sedation, using electrocardiogram (EEG) bispectral index monitoring. Follow-up was performed in an in-house gastrostomy drop-in clinic at 1 week and 1 month, supplemented with weekly telephone follow-up. Patients also had open access to the gastrostomy drop-in clinic for immediate advice and complication management. Results: Eighteen patients underwent primary insertion of a Monarch gastrostomy tube over 5 months. A total of 6/18 (33%) tubes displaced; 4/18 (22%) completely, 2/18 (11%) occult into the peritoneum. Four of 18 (22%) patients developed infection at the stoma site. Due to the unexpectedly poor performance of the tube, the study was terminated early. Conclusion: Initial experience with the Monarch disc-retained gastrostomy tube demonstrates it unsuitable for primary placement with current protocols. In view of the potentially serious complications, the Medicines and Healthcare Products Regulatory Agency (MHRA) has been informed. A request has been made to the distributer to reassess the tube design and/or review the procedure promoted for primary placement


    Institute of Scientific and Technical Information of China (English)

    James; J.Stewart


    Cartimandua's role has been much discussed by modern historians.1 Her position as queen of the Brigantes, wife of Venutius and client of Rome has been defined by scholars in the previous fifty years. One area not thoroughly examined is Tacitus' portrayal of her character, which changes dramatically from the lengthy passage in Annals 12.32-40 to the short account of Histories 3.45. While neither section flatters the monarch, Tacitus destroys her character in the Histories, equating her with figures such as G...

  16. Status and trends of the land bird avifauna on Tinian and Aguiguan, Mariana Islands (United States)

    Camp, Richard J.; Pratt, Thane K.; Amidon, Fred; Marshall, Ann P.; Kremer, Shelly; Laut, Megan


    Avian surveys were conducted on the islands of Tinian and Aguiguan, Marianas Islands, in 2008 by the U.S. Fish and Wildlife Service to provide current baseline densities and abundances and assess population trends using data collected from previous surveys. On Tinian, during the three surveys (1982, 1996, and 2008), 18 species were detected, and abundances and trends were assessed for 12 species. Half of the 10 native species—Yellow Bittern (Ixobrychus sinensis), White-throated Ground-Dove (Gallicolumba xanthonura), Collared Kingfisher (Todiramphus chloris), Rufous Fantail (Rhipidura rufifrons), and Micronesian Starling (Aplonis opaca)—and one alien bird—Island Collared-Dove (Streptopelia bitorquata)—have increased since 1982. Three native birds—Mariana Fruit-Dove (Ptilinopus roseicapilla), Micronesian Honeyeater (Myzomela rubratra), and Tinian Monarch (Monarcha takatsukasae)—have decreased since 1982. Trends for the remaining two native birds—White Tern (Gygis alba) and Bridled White-eye (Zosterops saypani)—and one alien bird—Eurasian Tree Sparrow (Passer montanus)—were considered relatively stable. Only five birds—White-throated Ground-Dove, Mariana Fruit-Dove, Tinian Monarch, Rufous Fantail, and Bridled White-eye—showed significant differences among regions of Tinian by year. Tinian Monarch was found in all habitat types, with the greatest monarch densities observed in limestone forest, secondary forest, and tangantangan (Leucaena leucocephala) thicket and the smallest densities found in open fields and urban/residential habitats. On Aguiguan, 19 species were detected on one or both of the surveys (1982 and 2008), and abundance estimates were produced for nine native and one alien species. Densities for seven of the nine native birds—White-throated Ground-Dove, Mariana Fruit-Dove, Collared Kingfisher, Rufous Fantail, Bridled White-eye, Golden White-eye (Cleptornis marchei), and Micronesian Starling—and the alien bird— Island

  17. The Prison Songs of Lili'uokalani


    Morris, Cynthia L.


    This thesis is a study of the music composed by Lili'uokalani, the last monarch of Hawai'i during her 1895 incarceration at the hands of the Republic of Hawai'i. Aspects of text and context are considered for each of the songs. This thesis also engages in a discussion of the alleged politicization of a copy of "Aloha Oe," notated by the Queen during the incarceration. My research consisted of visits to the Hawai'i State Archives and the Bishop Museum Archives in Honolulu. I enlisted the help ...

  18. Style and elegance in Bessarabian photographs of the 2nd half of the 19th century

    Directory of Open Access Journals (Sweden)

    Mihail Dohot


    Full Text Available The author presents the fashion trends of Western Europe, Romania and Bessarabia in the second half of the 19th century through the prism of photography. The influence of European monarchic houses on the fashion of that time is considered, as well as the role of artistic and cultural emancipation, which has left its imprint on society through the visual arts. The article lists fashion designers who contributed to the development of fashion and whose work was reflected in the photographs.

  19. The agreement to exchange (permutatio in roman, Byzantine and Serbian mediaeval law

    Directory of Open Access Journals (Sweden)

    Šarkić Srđan


    Full Text Available The first part of this paper is dedicated to the definition of the agreement to exchange (permutatio and emphasizes the difference, made by Roman lawyers, between exchange (barter and sale (emptio-venditio. The second part analyses Byzantine legal sources that mention this old contract, while the third part is dedicated to Serbian legal documents. In Serbian legal documents the exchange was mentioned as the agreement between a monarch and a monastery or a natural person (individual, concerning donations that were given to the Church.

  20. Man with a Hoe

    Directory of Open Access Journals (Sweden)

    David Middleton


    Full Text Available He leans on the short handle, knotted oak,Its flat blade pressed on brambled clay and stone.A boulder  shoulders thorns up from the soilWhile oxen plow a far-off pastoral farm.Whose stubble-fires smoke white toward skies in haze.He dominates the land as serf and lord,The subject monarch of his stark domain,His thistle-crown root-bound in freehold earth.Not fallen from some paradise whose cropsTurned golden while he plucked a harp’s ripe strings,He’s come down long hard centuries the same,Man’...

  1. Saying the Unsayable

    DEFF Research Database (Denmark)

    analyses important cultural, historical, political, religious, and legal forces shaping the popular image of the monarchy and, in particular, of King Bhumibol Adulyadej. In this manner, the book offers valuable insights into the relationships between monarchy, religion and democracy in Thailand - topics...... that, after the September 2006 coup d'état, gained renewed national and international interest. By addressing such contentious issues as Thai-style democracy, lése majesté legislation, religious symbolism and politics, monarchical traditions, and the royal sufficiency economy, this volume...

  2. La emigración política y la oposición violenta a la monarquía restaurada (1897-1931

    Directory of Open Access Journals (Sweden)

    Eduardo González Calleja


    Full Text Available This work examines the strategies of violent contention adopted by the various factions of the exiled Spanish anti-monarchical opposition during the first third of the twentieth century. In parallel, governmental responses to these threats are also analysed, with attention to their evolution from the first attempts at building international coalitions against anarchist terrorism in late nineteenth century, to the hardening of foreign defence policies during the Primo de Rivera dictatorship. Both the subversive activities of dissident groups and the intransigence of repressive political power contributed to the irreversible damage to the international image of the monarchy from the 1920s onwards.

  3. The Democratic Surplus that Constitutionalised the European Union

    DEFF Research Database (Denmark)

    Harste, Gorm


    This article questions the very foundation of the doctrine of a so-called “democratic deficit” in the EU. Yet in order to argue beyond nationalist myths, clear-cut concepts are necessary. Speaking about democracy in the EU, the article exposes four dimensions that constitute a “democratic surplus...... nation-state, the EU, secondly, is not build by nobles and monarchs, nor by war. Third, a separation of powers is obvious. And fourth, this article demonstrates how the EU rescued the democratic nation-state....

  4. Baking the first bread in space (United States)


    This Getaway Special program is a joint venture between Spar, Monarch flour and Telesat, with Telesat being responsible for the design, manufacture and implementation of the equipment. The purpose of the experiment is to investigate the behavior of bread yeast in the absence of gravity and in the presence of normal atmospheric pressure. The proposed design mixes flour, water and yeast on-orbit, allows the mixture to prove and then bakes it. This paper outlines the development history of the experiment, the various test programs and some of the problems encountered, with their solutions.

  5. Coins and maps: taxation and politics in the making of Brazilian new provinces; early 19th century

    Directory of Open Access Journals (Sweden)

    Vitor Marcos Gregório


    Full Text Available There was a lot of economic and political questions used as arguments during the decision making process which culminated with the emancipation of Amazonas and Paraná provinces in the mid-nineteenth century. Among them, the fiscality has a great importance, having been approached by those who advocated a new territorial organization through the creation of these two new provinces, as well for those who disagreed these proposals. This paper aims to analyse these element as indicators of the importance of the territorial organization questions as a historical object, and as a important instrument for the comprehension of the monarchical Brazilian State making process.

  6. Hanford Tank Farms Waste Feed Flow Loop Phase VI: PulseEcho System Performance Evaluation

    Energy Technology Data Exchange (ETDEWEB)

    Denslow, Kayte M.; Bontha, Jagannadha R.; Adkins, Harold E.; Jenks, Jeromy WJ; Hopkins, Derek F.


    This document presents the visual and ultrasonic PulseEcho critical velocity test results obtained from the System Performance test campaign that was completed in September 2012 with the Remote Sampler Demonstration (RSD)/Waste Feed Flow Loop cold-test platform located at the Monarch test facility in Pasco, Washington. This report is intended to complement and accompany the report that will be developed by WRPS on the design of the System Performance simulant matrix, the analysis of the slurry test sample concentration and particle size distribution (PSD) data, and the design and construction of the RSD/Waste Feed Flow Loop cold-test platform.

  7. Lucrezia d’Alagno o la celebración literaria y pública de la consorte de facto de Alfonso el Magnánimo


    Rodríguez Mesa, Francisco José


    Alfonso el Magnánimo, primer monarca napolitano de la Casa de Aragón, tuvo una relación en sus últimos años con la dama napolitana Lucrezia d’Alagno. Su entrada en la escena pública -y política- de Nápoles la situará habitualmente en celebraciones de la ciudad y la corte. Con testimonios literarios e históricos proponemos un análisis de su papel en los años finales del rey. The first Neapolitan monarch of the House of Aragon, Alfonso the Magnanimous, had an affair with the Neapolitan lady ...

  8. Specificity of herbivore-induced hormonal signaling and defensive traits in five closely related milkweeds (Asclepias spp.). (United States)

    Agrawal, Anurag A; Hastings, Amy P; Patrick, Eamonn T; Knight, Anna C


    Despite the recognition that phytohormonal signaling mediates induced responses to herbivory, we still have little understanding of how such signaling varies among closely related species and may generate herbivore-specific induced responses. We studied closely related milkweeds (Asclepias) to link: 1) plant damage by two specialist chewing herbivores (milkweed leaf beetles Labidomera clivicolis and monarch caterpillars Danaus plexippus); 2) production of the phytohormones jasmonic acid (JA), salicylic acid (SA), and abscisic acid (ABA); 3) induction of defensive cardenolides and latex; and 4) impacts on Danaus caterpillars. We first show that A. syriaca exhibits induced resistance following monarch herbivory (i.e., reduced monarch growth on previously damaged plants), while the defensively dissimilar A. tuberosa does not. We next worked with a broader group of five Asclepias, including these two species, that are highly divergent in defensive traits yet from the same clade. Three of the five species showed herbivore-induced changes in cardenolides, while induced latex was found in four species. Among the phytohormones, JA and ABA showed specific responses (although they generally increased) to insect species and among the plant species. In contrast, SA responses were consistent among plant and herbivore species, showing a decline following herbivore attack. Jasmonic acid showed a positive quantitative relationship only with latex, and this was strongest in plants damaged by D. plexippus. Although phytohormones showed qualitative tradeoffs (i.e., treatments that enhanced JA reduced SA), the few significant individual plant-level correlations among hormones were positive, and these were strongest between JA and ABA in monarch damaged plants. We conclude that: 1) latex exudation is positively associated with endogenous JA levels, even among low-latex species; 2) correlations among milkweed hormones are generally positive, although herbivore damage induces a

  9. Reimagining Society in 18th Century French Literature

    DEFF Research Database (Denmark)

    Kjærgård, Jonas Ross

    of interpersonal interaction flourished in the rich generic landscape of late eighteenth century French literature. These works of literature, the forgotten as well as the canonized ones, continuously intervened in that burgeoning social imaginary within which the struggle to define the happiness of all took place.......The French revolutionary shift from monarchical to popular sovereignty came clothed in a new political language, a significant part of which was a strange coupling of happiness and rights. In Old Regime ideology, Frenchmen were considered subjects who had no need of understanding why what...

  10. The fall of Edward II: Failures in kingship and masculinity, the letter of Manuel Fieschi and the cult of resurrected celebrities


    McAdam, Katie


    Edward II is a monarch whose name is almost synonymous with scandal and failure, and the infamous tale of his murder by red-hot poker at Berkeley Castle is one that looms over the bloody history of English royalty. The theories and historiography surrounding his political errors, his sexuality, the nature of his death and even the date of his death have changed continually since the end of his reign in 1327, both to further changing political agendas and to explore new historical narratives. ...

  11. Odpovědnost členů představenstva a dozorčí rady akciové společnosti ve srovnávacím pohledu


    Dubanská, Barbora


    "… being a director is a privilege be earned every day…" This quote was taken from a decision rendered by the Australian court in the case of ASIC v Rich, where a substantial pecuniary fine and other sanctions were imposed upon a board member, John David Rich, for breaches of his duties towards the company. The present-day directors of companies hold the positions of modern monarchs; their powers would remain unlimited if not for the current regulations in place. Members of the board of direc...

  12. Talking Division Up

    DEFF Research Database (Denmark)

    Villadsen, Lisa Storm

    In 1864, Denmark lost a border-war against Prussia and Austria and 2/5 of its territory: Holstein, Lauenborg and Slesvig. These areas had for centuries been both subjects to the Danish monarch and associated with the German Realm as quasi-independent duchies. In this paper, I turn to the beginning...... of the political program of inflated self-conception that led Denmark to war. While its initial spark emerged in an effort to gain democratic influence via a constitution (Denmark was an absolute monarchy until 1849), a key event in the nationalist surge was a movement dedicated to the celebration and idealization...

  13. Memória e missão: O Paiz e Gazeta Nacional. Imprensa do Rio De Janeiro (1884-1888

    Directory of Open Access Journals (Sweden)

    Andréa Santos da Silva Pessanha


    Full Text Available Newspapers constitute, featuring as object or font of study, a special field for the XIX century Brazilian society analysis. This study intends to show which representations about journalists, the press and its role were spread through the papers O Paiz and Gazeta Nacional, linked to the Court republican groups. Chronologically speaking, the paper covers the effervescence of the abolitionist movement in Rio de Janeiro, I mean, exactly the period when press plays the central role on the making of the campaign, bringing the political discussing about the end of slave labor and the crisis of the monarchical system to the public circle.

  14. [The historian's competence tested by authority: on an academic debate of the 18th century]. (United States)

    Schandeler, Jean-Pierre


    The debate which took place in the 1720s at the Royal Academy of Inscriptions and Letters on the possibility or impossibility of understanding the history of the first centuries of Rome is generally interpreted to be less of a debate than an important epistemological clarification. A contextualization which takes into account the political stakes of the debate allows one to understand that the debate was the beginning of a larger process of the autonomisation of the field of historical studies, not only from the perspective of disciplinary divides, but also in relation to monarchal power.

  15. Laat de kindertjes tot mij komen... Franse vorsten geportretteerd op een onbekende miniatuur


    Manuth, V.; Leeuwen, R. van


    Let the Children come unto Me: French monarchs portrayed on an unknown miniature This article identifies eleven portraits of the French kings and queens from the reigns of Henry II to Henry IV in a miniature depicting Christ Suffering the Little Children To Come unto Him. It was painted by an unknown French artist in the first decade of the 17th century. Remarkable is the exotic garb of the mothers, which is identified as the typical dress worn by gypsies and was thought at the time to be of ...

  16. La monarquía visigoda y su política matrimonial: el Reino visigodo de Toledo

    Directory of Open Access Journals (Sweden)

    María del Rosario VALVERDE CASTRO


    Full Text Available RESUMEN: Inscribiendo los matrimonios regios visigodos en su contexto histórico, pretendemos desvelar la relación existente entre las transformaciones de la institución monárquica visigoda y los cambios que experimentan las estrategias matrimoniales que dicha institución de poder pone en práctica durante su etapa toledana.ABSTRACT: By placing the royal Visigothic marriages within their historical context, we intend to show the relation between the transformation of the Visigothic monarchical institution and the changes undergone by the marriage strategies which this power institution put into practice during its Toletan stage.

  17. Women's royal holiness as the church-historical phenomenon

    Directory of Open Access Journals (Sweden)

    Efimov Vladimir Fedorovich


    Full Text Available The article under the topic of the Royal Holiness examines its specific subtype, the righteousness of the Royal women. Among the saint queens, empresses and princesses the authors distinguish the independent rulers, the widows, and wives of monarchs. The Holy Byzantine Empress and canonized European Queen, the Queen of the Georgian and the Russian Princess defended the Christian statehood brightened by feats of patience, mercy, by building the temples. In the conclusion of the article briefly tells about the Royal martyrs and Confessors of the 20th century.

  18. Centralizare versus „descentralizare”. „Reforma administrativă” de la 1938 (Centralization versus „decentralization”. „Administrative reform” from 1938

    Directory of Open Access Journals (Sweden)

    Florin GRECU


    Full Text Available The „decentralization” of the Romanian public administration during the authoritarian monarchy was meant to be achieved through the administrative reform from August 14, 1938, but the effect of the law consisted in an excessive centralization and militarization. Thus, the decentralization strengthened the state power, by nominating the officer corps in the administration and in the single party, the National Renaissance Front, focused decisions at the government level, at the expense of the 10 counties manufactured by the monarchical authoritarian regime.

  19. A monarquia segundo Jean-Jacques Rousseau

    Directory of Open Access Journals (Sweden)

    Renato Moscateli


    Full Text Available The Swiss philosopher Jean-Jacques Rousseau didn't limit himself to consider how a democratic State could be constructed in substitution to the absolutist monarchies existing at his time. He also dedicated his efforts to understand the monarchic regime itself, being conscientious of the obstacles to be overstepped so that this political form was surpassed in direction to other ones more next to his ideals, in which all the citizens could participate actively of the administration of the government. Thus, the article will handle some aspects of the Rousseau's reflection about the monarchy, from his criticism to the regime until his concrete proposals for possible reforms on it.

  20. The heresy of tzarebozhie as the "reincarnation" of the cult of the deified rulers of the Ancient World

    Directory of Open Access Journals (Sweden)

    Nikolsky Evgeny Vladimirovich


    Full Text Available From theological perspectives, this paper studies the current movement of Russian Rastafarianes. The author analyzes the origins of this movement rooted in pre-revolutionary sectarianism; also, he conducts typological parallels with the cult of the deified rulers of the Ancient World and shows the similarity of views of Russian Rastafarianes with the idea of the monarch in ancient paganism. In conclusion, the article shows this new religious movement with its external Christian attributes promotes pure pagan world where people worship not the God but creation. Final dechristianization of Russian Rastafarianes is predictable.

  1. The British Monarchy On Screen


    Merck, Mandy


    Moving images of the British monarchy, in fact and fiction, are almost as old as the moving image itself, dating back to an 1895 American drama, The Execution of Mary Queen of Scots. British monarchs even appeared in the new ‘animated photography’ from 1896, led by Queen Victoria. Half a century later, the 1953 coronation of Elizabeth II was a milestone in the adoption of television, watched by 20 million Britons and 100 million North Americans. At the century’s end, Princess Diana’s funeral ...

  2. Satellite Infrared Radiation Measurements Prior to the Major Earthquakes (United States)

    Ouzounov, Dimitar; Pulintes, S.; Bryant, N.; Taylor, Patrick; Freund, F.


    This work describes our search for a relationship between tectonic stresses and increases in mid-infrared (IR) flux as part of a possible ensemble of electromagnetic (EM) phenomena that may be related to earthquake activity. We present and &scuss observed variations in thermal transients and radiation fields prior to the earthquakes of Jan 22, 2003 Colima (M6.7) Mexico, Sept. 28 .2004 near Parkfield (M6.0) in California and Northern Sumatra (M8.5) Dec. 26,2004. Previous analysis of earthquake events has indicated the presence of an IR anomaly, where temperatures increased or did not return to its usual nighttime value. Our procedures analyze nighttime satellite data that records the general condtion of the ground after sunset. We have found from the MODIS instrument data that five days before the Colima earthquake the IR land surface nighttime temperature rose up to +4 degrees C in a 100 km radius around the epicenter. The IR transient field recorded by MODIS in the vicinity of Parkfield, also with a cloud free environment, was around +1 degree C and is significantly smaller than the IR anomaly around the Colima epicenter. Ground surface temperatures near the Parkfield epicenter four days prior to the earthquake show steady increase. However, on the night preceding the quake, a significant drop in relative humidity was indicated, process similar to those register prior to the Colima event. Recent analyses of continuous ongoing long- wavelength Earth radiation (OLR) indicate significant and anomalous variability prior to some earthquakes. The cause of these anomalies is not well understood but could be the result of a triggering by an interaction between the lithosphere-hydrosphere and atmospheric related to changes in the near surface electrical field and/or gas composition prior to the earthquake. The OLR anomaly usually covers large areas surrounding the main epicenter. We have found strong anomalies signal (two sigma) along the epicentral area signals on Dec 21

  3. Time-lapse imaging of fault properties at seismogenic depth using repeating earthquakes, active sources and seismic ambient noise (United States)

    Cheng, Xin


    The time-varying stress field of fault systems at seismogenic depths plays the mort important role in controlling the sequencing and nucleation of seismic events. Using seismic observations from repeating earthquakes, controlled active sources and seismic ambient noise, five studies at four different fault systems across North America, Central Japan, North and mid-West China are presented to describe our efforts to measure such time dependent structural properties. Repeating and similar earthquakes are hunted and analyzed to study the post-seismic fault relaxation at the aftershock zone of the 1984 M 6.8 western Nagano and the 1976 M 7.8 Tangshan earthquakes. The lack of observed repeating earthquakes at western Nagano is attributed to the absence of a well developed weak fault zone, suggesting that the fault damage zone has been almost completely healed. In contrast, the high percentage of similar and repeating events found at Tangshan suggest the existence of mature fault zones characterized by stable creep under steady tectonic loading. At the Parkfield region of the San Andreas Fault, repeating earthquake clusters and chemical explosions are used to construct a scatterer migration image based on the observation of systematic temporal variations in the seismic waveforms across the occurrence time of the 2004 M 6 Parkfield earthquake. Coseismic fluid charge or discharge in fractures caused by the Parkfield earthquake is used to explain the observed seismic scattering properties change at depth. In the same region, a controlled source cross-well experiment conducted at SAFOD pilot and main holes documents two large excursions in the travel time required for a shear wave to travel through the rock along a fixed pathway shortly before two rupture events, suggesting that they may be related to pre-rupture stress induced changes in crack properties. At central China, a tomographic inversion based on the theory of seismic ambient noise and coda wave interferometry

  4. Alfonso el Regenerador. Performing Monarchy and Spanish Nationalist Imaginary, from a comparative perspective (1902-1913

    Directory of Open Access Journals (Sweden)

    Javier Moreno Luzón


    Full Text Available This article studies the links between monarchy and Spanish nationalism in the first decade of Alfonso XIII’s reign. It uses a comparative perspective to analyse the Spanish case in the context of the performing monarchies than emerged in Europe in the period 1870-1914. It focuses on three different aspects: the great monarchical ceremonies, specially the royal oath to the Constitution —or coronation—; the royal trips, extraordinarily developed; and the military ceremonies containing a strong nationalist meaning as the annual swearing of loyalty to the national flag by new soldiers. Through those performances, the crown was integrated in a national imaginary dominated by discourses and practices of regeneration of the fatherland in the aftermath of the colonial disaster of 1898: in such a regeneration, the king was thought as a necessary force. In fact, Alfonso was known as el Regenerador (the regenerator. At a lower level of pomp and splendor than other european monarchies, the Spanish king was perceived as a national symbol by different groups. Among them, local elites, various associations and the heterogeneus public of the royal spectacles, shown by the mass media. Those performances reinforced the political role of a king that enjoyed constitutional executive powers, like most of the European monarchs, and whose decisions were highly controversial.

  5. Praising the Ruler: Panegyrical Poetry and Russian Absolutism

    Directory of Open Access Journals (Sweden)

    Joachim Klein


    Full Text Available It is difficult to overrate the importance of the panegyric tradition for early modern Russian literature. Between the middle of the 17th to the end of the 18th century, it was practiced in many different genres—almost all Russian poets praised the ruler. This poetry deserves our interest as a specific form of political literature. As such it is not only relevant for the cult of the Russian monarchs, but it also sheds some light on the political mentality of their loyal—and literate—subjects in the age of Russian absolutism. Panegyrical poetry is per definitionem a thoroughly affirmative, noncritical form of political literature. But this did not prevent it from offering a certain scope for the expression of diverse and even contradictory political ideals. This can be exemplified by the panegyrical poems written in the early 1760s in the context of the coup d’état staged by Catherine II and against the backdrop of the Russo-Prussian peace treaty initiated by her predecessor, Peter III. In this situation, a fundamental difference of opinion about the tasks of the monarch and the mission of the Russian state emerged.

  6. Induced responses to herbivory and jasmonate in three milkweed species. (United States)

    Rasmann, Sergio; Johnson, M Daisy; Agrawal, Anurag A


    We studied constitutive and induced defensive traits (latex exudation, cardenolides, proteases, and C/N ratio) and resistance to monarch caterpillars (Danaus plexippus) in three closely related milkweed species (Asclepias angustifolia, A. barjoniifolia and A. fascicularis). All traits showed significant induction in at least one of the species. Jasmonate application only partially mimicked the effect of monarch feeding. We found some correspondence between latex and cardenolide content and reduced larval growth. Larvae fed cut leaves of A. angustifolia grew better than larvae fed intact plants. Addition of the cardenolide digitoxin to cut leaves reduced larval growth but ouabain (at the same concentration) had no effect. We, thus, confirm that latex and cardenolides are major defenses in milkweeds, effective against a specialist herbivore. Other traits such as proteases and C/N ratio additionally may be integrated in the defense scheme of those plants. Induction seems to play an important role in plants that have an intermediate level of defense, and we advocate incorporating induction as an additional axis of the plant defense syndrome hypothesis.

  7. Feeding on toxic prey. The praying mantis (Mantodea) as predator of poisonous butterfly and moth (Lepidoptera) caterpillars. (United States)

    Mebs, Dietrich; Wunder, Cora; Pogoda, Werner; Toennes, Stefan W


    Caterpillars of the monarch butterfly, Danaus plexippus, feed on milkweed plants, Asclepias spp. (Apocynaceae), and sequester their toxic cardenolides aimed at deterring predators. Nevertheless, Chinese praying mantids, Tenodera sinensis, consume these caterpillars after removing the midgut ("gutting") including its plant content. In the present study, monarch caterpillars raised on A. curassavica, and those of the death's-head hawkmoth, Acherontia atropos, raised on Atropa belladonna containing atropine, were fed to mantids, Hierodula membranacea, which removed the gut of both species discarding about 59% of cardenolides and more than 90% of atropine, respectively. The ingestion of these compounds produced no apparent ill effects in the mantids and both were excreted with faeces. On the other hand, when mantids were fed with larvae of two moth species, Amata mogadorensis and Brahmaea certia, raised on non-poisonous host plants, the mantids showed the same gutting behaviour, thereby discarding indigestible plant material. As polar compounds, e.g. cardenolides and atropine, are not absorbed from the mantids midgut and do not pass the gut membrane, this enables the mantids to feed on toxic prey. Copyright © 2017 Elsevier Ltd. All rights reserved.

  8. Ecosystem-Based Incorporation of Nectar-Producing Plants for Stink Bug Parasitoids

    Directory of Open Access Journals (Sweden)

    Glynn Tillman


    Full Text Available Adult parasitoids of pest insects rely on floral resources for survival and reproduction, but can be food-deprived in intensively managed agricultural systems lacking these resources. Stink bugs are serious pests for crops in southwest Georgia. Provisioning nectar-producing plants for parasitoids of stink bugs potentially can enhance biocontrol of these pests. Knowledge of spatial and temporal availability and distribution of stink bugs in host plants is necessary for appropriate timing and placement of flowering plants in agroecosystems. Stink bugs move between closely associated host plants throughout the growing season in response to deteriorating suitability of their host plants. In peanut-cotton farmscapes, stink bugs develop in peanut, and subsequently the adults disperse into adjacent cotton. Parasitism of Nezara viridula (L. adults by Trichopoda pennipes (F. at the peanut-cotton interface was significantly higher in cotton with a strip of milkweed or buckwheat between the two crops than in cotton alone. Milkweed and buckwheat also provided nectar to a wide range of insect pollinators. Monarch butterflies fed on milkweed. When placed between peanut and cotton, a strip of soybean was an effective trap crop for cotton, reducing economic damage. Incorporation of buckwheat near soybean enhanced parasitism of Euschistus servus (Say eggs by Telenomus podisi Ashmead in cotton. In conclusion, nectar provision enhances biocontrol of stink bugs, acts together with other management tactics for stink bug control, and aids in conservation of natural enemies, insect pollinators, and the monarch butterfly.

  9. Stepwise evolution of resistance to toxic cardenolides via genetic substitutions in the Na+/K+ -ATPase of milkweed butterflies (lepidoptera: Danaini). (United States)

    Petschenka, Georg; Fandrich, Steffi; Sander, Nils; Wagschal, Vera; Boppré, Michael; Dobler, Susanne


    Despite the monarch butterfly (Danaus plexippus) being famous for its adaptations to the defensive traits of its milkweed host plants, little is known about the macroevolution of these traits. Unlike most other animal species, monarchs are largely insensitive to cardenolides, because their target site, the sodium pump (Na(+)/K(+) -ATPase), has evolved amino acid substitutions that reduce cardenolide binding (so-called target site insensitivity, TSI). Because many, but not all, species of milkweed butterflies (Danaini) are associated with cardenolide-containing host plants, we analyzed 16 species, representing all phylogenetic lineages of milkweed butterflies, for the occurrence of TSI by sequence analyses of the Na(+)/K(+) -ATPase gene and by enzymatic assays with extracted Na(+)/K(+) -ATPase. Here we report that sensitivity to cardenolides was reduced in a stepwise manner during the macroevolution of milkweed butterflies. Strikingly, not all Danaini typically consuming cardenolides showed TSI, but rather TSI was more strongly associated with sequestration of toxic cardenolides. Thus, the interplay between bottom-up selection by plant compounds and top-down selection by natural enemies can explain the evolutionary sequence of adaptations to these toxins. © 2013 The Author(s). Evolution © 2013 The Society for the Study of Evolution.

  10. Honor and war. Tension between the reality of war and ideological discourse in castilian chronicles of the first half of the fourteenth century

    Directory of Open Access Journals (Sweden)

    Fernando Arias Guillén


    Full Text Available Chronicles represented one of the principal instruments used by different medieval powers to project ideas that legitimized and eulogized their authority. Warfare therefore frequently constitutes a key element of these texts. During the reign of Alfonso XI, the Crown of Castile promoted works that not only justified and glorified the king’s political activity, but also invoked an idealised image of warfare. This image was based on the idea of warfare as an honourable activity and the exclusive domain of a social elite, and therefore did not reflect real military practice. The authors of these works also considered the defence of the king’s honour a valid justification for war, even when this conflicted with more pragmatic attitudes. The Royal Chronicler Fernán Sánchez de Valladolid managed to reconcile the two in order to maintain the monarch beyond criticism. onicler, achieved to combine both in order to maintain the monarch beyond criticism. This approach was not exclusive to Castilian Chronicles, as it also constituted a characteristic element of other contemporary texts, like the works of Froissart or the French Royal Chronicles. This paper concludes that, rather than providing a reliable account of an event, chronicles often based their narratives on a series of common images and ideas that reflected a predetermined ideological discourse.

  11. Bird populations on the Island of Tinian: persistence despite wholesale loss of native forests (United States)

    Camp, Richard J.; Amidon, Frederick A.; Marshall, Ann P.; Pratt, Thane K.


    Bird habitat on the island of Tinian, Mariana Islands, has been substantially altered, and only around 5% of the island has native forest today. The modern bird fauna is likely to be a subset of the original avifauna where only species tolerant to native forest loss and human disturbance have survived. Avian surveys were conducted on the island in 2008 by the U.S. Fish and Wildlife Service to provide current densities and abundances of the remaining species, and assess population trends using data collected from previous surveys. During the three surveys (1982, 1996, and 2008), 18 species were detected, and abundances and trends were assessed for 11 species. Five of the nine native species and one alien bird have increased since 1982. Three native birds—Mariana Fruit-Dove (Ptilinopusroseicapilla), Micronesian Honeyeater (Myzomela rubratra), and Tinian Monarch (Monarcha takatsukasae)—have decreased since 1982. Trends for the remaining two birds (one native and one alien) were considered relatively stable. Only five birds, including the Tinian Monarch, showed significant differences among regions of Tinian by year. Increased development on Tinian may result in increases in habitat clearing and expansion of human-dominated habitats, and declines in some bird populations would likely continue or be exacerbated with these actions. Expanded development activities on Tinian would also mean increased cargo movement between Guam and Tinian, elevating the probability of transporting the Brown Tree Snake (Boiga irregularis) to Tinian, which would lead to precipitous decreases and extinctions.

  12. Storage hexamer utilization in two lepidopterans: differences correlated with the timing of egg formation

    Directory of Open Access Journals (Sweden)

    M.L. Pan


    Full Text Available Most insects produce two or more storage hexamers whose constituents and developmental profiles are sufficiently different to suggest specialization in the ways that they support metamorphosis and reproduction. Hexamerin specializations are compared here in the Cecropia moth (Hyalophora cecropia, which produces eggs during the pupal-adult molt, and the Monarch butterfly (Danaus plexippus, which produces eggs under long-day conditions after adult eclosion. In both sexes of both species, reserves of arylphorin (ArH were exhausted by the end of metamorphosis. In Cecropia, the same was true for the high-methionine hexamerins, V-MtH and M-MtH. But in short day Monarch females 20-30% of the pupal reserves of V-MtH and M-MtH survived metamorphosis, persisting until long-day conditions were imposed to stimulate egg formation. Differences in storage sites have been documented in other lepidopterans, with MtH reserves being found primarily in fat body protein granules and the ArH reserve being found primarily in the hemolymph. Similar differences could explain how a fraction of the MtH's, but not of ArH, escapes utilization during metamorphosis in a species with post-eclosion egg formation. No differences in utilization schedules were detected between V- and M-MtH, despite divergent compositions and antigenic reactivity.

  13. Experimental evidence shows no fractionation of strontium isotopes ((87)Sr/(86)Sr) among soil, plants, and herbivores: implications for tracking wildlife and forensic science. (United States)

    Flockhart, D T Tyler; Kyser, T Kurt; Chipley, Don; Miller, Nathan G; Norris, D Ryan


    Strontium isotopes ((87)Sr/(86)Sr) can be useful biological markers for a wide range of forensic science applications, including wildlife tracking. However, one of the main advantages of using (87)Sr/(86)Sr values, that there is no fractionation from geological bedrock sources through the food web, also happens to be a critical assumption that has never been tested experimentally. We test this assumption by measuring (87)Sr/(86)Sr values across three trophic levels in a controlled greenhouse experiment. Adult monarch butterflies were raised on obligate larval host milkweed plants that were, in turn, grown on seven different soil types collected across Canada. We found no significant differences between (87)Sr/(86)Sr values in leachable Sr from soil minerals, organic soil, milkweed leaves, and monarch butterfly wings. Our results suggest that strontium isoscapes developed from (87)Sr/(86)Sr values in bedrock or soil may serve as a reliable biological marker in forensic science for a range of taxa and across large geographic areas.

  14. Evaluation of Three Dimensional Underground Structure at SAFOD Project

    International Nuclear Information System (INIS)

    Malin, Peter


    In the SAFOD project, the imaging of the fault zone was implemented using data acquired by a pilot hole array of a vertical depth of 2 km and then a main hole was drilled using these data. The trajectory of the main hole below vertical depth of 1.5 km was angled toward/through the fault zone up to a vertical depth of 3 km. An sensor array was located in the hole. As a result, the hypocenter locations of small earthquakes within the fault zone were determined with high accuracy (location error within 10 meters) and the location of the fault zone was able to be identified with high accuracy. Using this data, high resolution underground structure around the San Andreas fault zone was obtained. It was reported that this underground structure revealed the deep structure of the San Andreas Fault at the Parkfield site as well as the branch fault. (author)

  15. Outline and Prospects of SAFOD Project

    International Nuclear Information System (INIS)

    Malin, Peter


    SAFOD is a project to investigate and understand the rupture process of the San Andreas Fault through an investigation of material composition of the fault zone; observation to reveal underground structure; and measurement of physical properties, such as stresses on and around the fault, permeability, and pore pressure, etc. at Parkfield. In addition to observation by a seismometer and tilt-meter which were installed at a depth of about three kilometers in a borehole in the fault zone, core sampling and geophysical logging were carried out as part of this project. High quality data, such as that on guided waves (special seismic waves that propagate in the fault zone) and repeated earthquakes (earthquakes with very similar waveforms) were acquired. Such data can be very useful in understanding the fault rupture process. (authors)

  16. Periodic, chaotic, and doubled earthquake recurrence intervals on the deep San Andreas fault. (United States)

    Shelly, David R


    Earthquake recurrence histories may provide clues to the timing of future events, but long intervals between large events obscure full recurrence variability. In contrast, small earthquakes occur frequently, and recurrence intervals are quantifiable on a much shorter time scale. In this work, I examine an 8.5-year sequence of more than 900 recurring low-frequency earthquake bursts composing tremor beneath the San Andreas fault near Parkfield, California. These events exhibit tightly clustered recurrence intervals that, at times, oscillate between approximately 3 and approximately 6 days, but the patterns sometimes change abruptly. Although the environments of large and low-frequency earthquakes are different, these observations suggest that similar complexity might underlie sequences of large earthquakes.

  17. Periodic, chaotic, and doubled earthquake recurrence intervals on the deep San Andreas Fault (United States)

    Shelly, David R.


    Earthquake recurrence histories may provide clues to the timing of future events, but long intervals between large events obscure full recurrence variability. In contrast, small earthquakes occur frequently, and recurrence intervals are quantifiable on a much shorter time scale. In this work, I examine an 8.5-year sequence of more than 900 recurring low-frequency earthquake bursts composing tremor beneath the San Andreas fault near Parkfield, California. These events exhibit tightly clustered recurrence intervals that, at times, oscillate between ~3 and ~6 days, but the patterns sometimes change abruptly. Although the environments of large and low-frequency earthquakes are different, these observations suggest that similar complexity might underlie sequences of large earthquakes.

  18. Effects of methods of attenuation correction on source parameter determination (United States)

    Sonley, Eleanor; Abercrombie, Rachel E.

    We quantify the effects of using different approaches to model individual earthquake spectra. Applying different approaches can introduce significant variability in the calculated source parameters, even when applied to the same data. To compare large and small earthquake source parameters, the results of multiple studies need to be combined to extend the magnitude range, but the variability introduced by the different approaches hampers the outcome. When studies are combined, there is large uncertainty and large scatter and some systematic differences have been neglected. We model individual earthquake spectra from repeating earthquakes (M˜2) at Parkfield, CA, recorded by a borehole network. We focus on the effects of trade-offs between attenuation (Q) and corner frequency in spectral fitting and the effect of the model shape at the corner frequency on radiated energy. The trade-off between attenuation and corner frequency can increase radiated energy by up to 400% and seismic moment by up to 100%.

  19. Foreshocks and aftershocks of the Great 1857 California earthquake (United States)

    Meltzner, A.J.; Wald, D.J.


    The San Andreas fault is the longest fault in California and one of the longest strike-slip faults anywhere in the world, yet we know little about many aspects of its behavior before, during, and after large earthquakes. We conducted a study to locate and to estimate magnitudes for the largest foreshocks and aftershocks of the 1857 M 7.9 Fort Tejon earthquake on the central and southern segments of the fault. We began by searching archived first-hand accounts from 1857 through 1862, by grouping felt reports temporally, and by assigning modified Mercalli intensities to each site. We then used a modified form of the grid-search algorithm of Bakum and Wentworth, derived from empirical analysis of modern earthquakes, to find the location and magnitude most consistent with the assigned intensities for each of the largest events. The result confirms a conclusion of Sieh that at least two foreshocks ('dawn' and 'sunrise') located on or near the Parkfield segment of the San Andreas fault preceded the mainshock. We estimate their magnitudes to be M ~ 6.1 and M ~ 5.6, respectively. The aftershock rate was below average but within one standard deviation of the number of aftershocks expected based on statistics of modern southern California mainshock-aftershock sequences. The aftershocks included two significant events during the first eight days of the sequence, with magnitudes M ~ 6.25 and M ~ 6.7, near the southern half of the rupture; later aftershocks included a M ~ 6 event near San Bernardino in December 1858 and a M ~ 6.3 event near the Parkfield segment in April 1860. From earthquake logs at Fort Tejon, we conclude that the aftershock sequence lasted a minimum of 3.75 years.

  20. Joint Inversion of Vp, Vs, and Resistivity at SAFOD (United States)

    Bennington, N. L.; Zhang, H.; Thurber, C. H.; Bedrosian, P. A.


    Seismic and resistivity models at SAFOD have been derived from separate inversions that show significant spatial similarity between the main model features. Previous work [Zhang et al., 2009] used cluster analysis to make lithologic inferences from trends in the seismic and resistivity models. We have taken this one step further by developing a joint inversion scheme that uses the cross-gradient penalty function to achieve structurally similar Vp, Vs, and resistivity images that adequately fit the seismic and magnetotelluric MT data without forcing model similarity where none exists. The new inversion code, tomoDDMT, merges the seismic inversion code tomoDD [Zhang and Thurber, 2003] and the MT inversion code Occam2DMT [Constable et al., 1987; deGroot-Hedlin and Constable, 1990]. We are exploring the utility of the cross-gradients penalty function in improving models of fault-zone structure at SAFOD on the San Andreas Fault in the Parkfield, California area. Two different sets of end-member starting models are being tested. One set is the separately inverted Vp, Vs, and resistivity models. The other set consists of simple, geologically based block models developed from borehole information at the SAFOD drill site and a simplified version of features seen in geophysical models at Parkfield. For both starting models, our preliminary results indicate that the inversion produces a converging solution with resistivity, seismic, and cross-gradient misfits decreasing over successive iterations. We also compare the jointly inverted Vp, Vs, and resistivity models to borehole information from SAFOD to provide a "ground truth" comparison.

  1. Imaging Stress Transients and Fault Zone Processes with Crosswell Continuous Active-Source Seismic Monitoring at the San Andreas Fault Observatory at Depth (United States)

    Niu, F.; Taira, T.; Daley, T. M.; Marchesini, P.; Robertson, M.; Wood, T.


    Recent field and laboratory experiments identify seismic velocity changes preceding microearthquakes and rock failure (Niu et al., 2008, Nature; Scuderi et al., 2016, NatureGeo), which indicates that a continuous monitoring of seismic velocity might provide a mean of understanding of the earthquake nucleation process. Crosswell Continuous Active-Source Seismic Monitoring (CASSM) using borehole sources and sensors has proven to be an effective tool for measurements of seismic velocity and its temporal variation at seismogenic depth (Silver, et al, 2007, BSSA; Daley, et al, 2007, Geophysics). To expand current efforts on the CASSM development, in June 2017 we have begun to conduct a year-long CASSM field experiment at the San Andreas Fault Observatory at Depth (SAFOD) in which the preceding field experiment detected the two sudden velocity reductions approximately 10 and 2 hours before microearthquakes (Niu et al., 2008, Nature). We installed a piezoelectric source and a three-component accelerometer at the SAFOD pilot and main holes ( 1 km depth) respectively. A seismic pulse was fired from the piezoelectric source four times per second. Each waveform was recorded 150-ms-long data with a sampling rate of 48 kHz. During this one-year experiment, we expect to have 10-15 microearthquakes (magnitude 1-3) occurring near the SAFOD site, and the data collected from the new experiment would allow us to further explore a relation between velocity changes and the Parkfield seismicity. Additionally, the year-long data provide a unique opportunity to study long-term velocity changes that might be related to seasonal stress variations at Parkfield (Johnson et al., 2017, Science). We will report on initial results of the SAFOD CASSM experiment and operational experiences of the CASSM development.

  2. Constraining earthquake source inversions with GPS data: 1. Resolution-based removal of artifacts (United States)

    Page, M.T.; Custodio, S.; Archuleta, R.J.; Carlson, J.M.


    We present a resolution analysis of an inversion of GPS data from the 2004 Mw 6.0 Parkfield earthquake. This earthquake was recorded at thirteen 1-Hz GPS receivers, which provides for a truly coseismic data set that can be used to infer the static slip field. We find that the resolution of our inverted slip model is poor at depth and near the edges of the modeled fault plane that are far from GPS receivers. The spatial heterogeneity of the model resolution in the static field inversion leads to artifacts in poorly resolved areas of the fault plane. These artifacts look qualitatively similar to asperities commonly seen in the final slip models of earthquake source inversions, but in this inversion they are caused by a surplus of free parameters. The location of the artifacts depends on the station geometry and the assumed velocity structure. We demonstrate that a nonuniform gridding of model parameters on the fault can remove these artifacts from the inversion. We generate a nonuniform grid with a grid spacing that matches the local resolution length on the fault and show that it outperforms uniform grids, which either generate spurious structure in poorly resolved regions or lose recoverable information in well-resolved areas of the fault. In a synthetic test, the nonuniform grid correctly averages slip in poorly resolved areas of the fault while recovering small-scale structure near the surface. Finally, we present an inversion of the Parkfield GPS data set on the nonuniform grid and analyze the errors in the final model. Copyright 2009 by the American Geophysical Union.

  3. Migrating tremors illuminate complex deformation beneath the seismogenic San Andreas fault. (United States)

    Shelly, David R


    The San Andreas fault is one of the most extensively studied faults in the world, yet its physical character and deformation mode beneath the relatively shallow earthquake-generating portion remain largely unconstrained. Tectonic 'non-volcanic' tremor, a recently discovered seismic signal probably generated by shear slip on the deep extension of some major faults, can provide new insight into the deep fate of such faults, including that of the San Andreas fault near Parkfield, California. Here I examine continuous seismic data from mid-2001 to 2008, identifying tremor and decomposing the signal into different families of activity based on the shape and timing of the waveforms at multiple stations. This approach allows differentiation between activities from nearby patches of the deep fault and begins to unveil rich and complex patterns of tremor occurrence. I find that tremor exhibits nearly continuous migration, with the most extensive episodes propagating more than 20 kilometres along fault strike at rates of 15-80 kilometres per hour. This suggests that the San Andreas fault remains a localized through-going structure, at least to the base of the crust, in this area. Tremor rates and recurrence behaviour changed markedly in the wake of the 2004 magnitude-6.0 Parkfield earthquake, but these changes were far from uniform within the tremor zone, probably reflecting heterogeneous fault properties and static and dynamic stresses decaying away from the rupture. The systematic recurrence of tremor demonstrated here suggests the potential to monitor detailed time-varying deformation on this portion of the deep San Andreas fault, deformation which unsteadily loads the shallower zone that last ruptured in the 1857 magnitude-7.9 Fort Tejon earthquake.

  4. Three-Dimensional Investigation of a 5 m Deflected Swale along the San Andreas Fault in the Carrizo Plain

    KAUST Repository

    Akciz, S. O.; Ludwig, L. G.; Zielke, Olaf; Arrowsmith, J. R.


    Topographic maps produced from Light Detection and Ranging (LiDAR) data are useful for paleoseismic and neotectonic research because they provide submeter representation of faulting-related surface features. Offset measurements of geomorphic features, made in the field or on a remotely sensed imagery, commonly assume a straight or smooth (i.e., undeflected) pre-earthquake geometry. Here, we present results from investigation of an ∼20 cm deep and >5 m wide swale with a sharp bend along the San Andreas fault (SAF) at the Bidart fan site in the Carrizo Plain, California. From analysis of LiDAR topography images and field measurements, the swale was initially interpreted as a channel tectonically offset ∼4:7 m. Our observations from exposures in four backhoe excavations and 25 hand-dug trenchettes show that even though a sharp bend in the swale coincides with the trace of the A.D. 1857 fault rupture, the swale formed after the 1857 earthquake and was not tectonically offset. Subtle fractures observed within a surficial gravel unit overlying the 1857 rupture trace are similar to fractures previously documented at the Phelan fan and LY4 paleoseismic sites 3 and 35 km northwest of Bidart fan, respectively. Collectively, the fractures suggest that a post-1857 moderate-magnitude earthquake caused ground cracking in the Carrizo and Cholame stretches of the SAF. Our observations emphasize the importance of excavation at key locations to validate remote and ground-based measurements, and we advocate more geomorphic characterization for each site if excavation is not possible.

  5. Semiautomated tremor detection using a combined cross-correlation and neural network approach (United States)

    Horstmann, Tobias; Harrington, Rebecca M.; Cochran, Elizabeth S.


    Despite observations of tectonic tremor in many locations around the globe, the emergent phase arrivals, low‒amplitude waveforms, and variable event durations make automatic detection a nontrivial task. In this study, we employ a new method to identify tremor in large data sets using a semiautomated technique. The method first reduces the data volume with an envelope cross‒correlation technique, followed by a Self‒Organizing Map (SOM) algorithm to identify and classify event types. The method detects tremor in an automated fashion after calibrating for a specific data set, hence we refer to it as being “semiautomated”. We apply the semiautomated detection algorithm to a newly acquired data set of waveforms from a temporary deployment of 13 seismometers near Cholame, California, from May 2010 to July 2011. We manually identify tremor events in a 3 week long test data set and compare to the SOM output and find a detection accuracy of 79.5%. Detection accuracy improves with increasing signal‒to‒noise ratios and number of available stations. We find detection completeness of 96% for tremor events with signal‒to‒noise ratios above 3 and optimal results when data from at least 10 stations are available. We compare the SOM algorithm to the envelope correlation method of Wech and Creager and find the SOM performs significantly better, at least for the data set examined here. Using the SOM algorithm, we detect 2606 tremor events with a cumulative signal duration of nearly 55 h during the 13 month deployment. Overall, the SOM algorithm is shown to be a flexible new method that utilizes characteristics of the waveforms to identify tremor from noise or other seismic signals.

  6. Talc-bearing serpentinite and the creeping section of the San Andreas fault. (United States)

    Moore, Diane E; Rymer, Michael J


    The section of the San Andreas fault located between Cholame Valley and San Juan Bautista in central California creeps at a rate as high as 28 mm yr(-1) (ref. 1), and it is also the segment that yields the best evidence for being a weak fault embedded in a strong crust. Serpentinized ultramafic rocks have been associated with creeping faults in central and northern California, and serpentinite is commonly invoked as the cause of the creep and the low strength of this section of the San Andreas fault. However, the frictional strengths of serpentine minerals are too high to satisfy the limitations on fault strength, and these minerals also have the potential for unstable slip under some conditions. Here we report the discovery of talc in cuttings of serpentinite collected from the probable active trace of the San Andreas fault that was intersected during drilling of the San Andreas Fault Observatory at Depth (SAFOD) main hole in 2005. We infer that the talc is forming as a result of the reaction of serpentine minerals with silica-saturated hydrothermal fluids that migrate up the fault zone, and the talc commonly occurs in sheared serpentinite. This discovery is significant, as the frictional strength of talc at elevated temperatures is sufficiently low to meet the constraints on the shear strength of the fault, and its inherently stable sliding behaviour is consistent with fault creep. Talc may therefore provide the connection between serpentinite and creep in the San Andreas fault, if shear at depth can become localized along a talc-rich principal-slip surface within serpentinite entrained in the fault zone.

  7. Three-Dimensional Investigation of a 5 m Deflected Swale along the San Andreas Fault in the Carrizo Plain

    KAUST Repository

    Akciz, S. O.


    Topographic maps produced from Light Detection and Ranging (LiDAR) data are useful for paleoseismic and neotectonic research because they provide submeter representation of faulting-related surface features. Offset measurements of geomorphic features, made in the field or on a remotely sensed imagery, commonly assume a straight or smooth (i.e., undeflected) pre-earthquake geometry. Here, we present results from investigation of an ∼20 cm deep and >5 m wide swale with a sharp bend along the San Andreas fault (SAF) at the Bidart fan site in the Carrizo Plain, California. From analysis of LiDAR topography images and field measurements, the swale was initially interpreted as a channel tectonically offset ∼4:7 m. Our observations from exposures in four backhoe excavations and 25 hand-dug trenchettes show that even though a sharp bend in the swale coincides with the trace of the A.D. 1857 fault rupture, the swale formed after the 1857 earthquake and was not tectonically offset. Subtle fractures observed within a surficial gravel unit overlying the 1857 rupture trace are similar to fractures previously documented at the Phelan fan and LY4 paleoseismic sites 3 and 35 km northwest of Bidart fan, respectively. Collectively, the fractures suggest that a post-1857 moderate-magnitude earthquake caused ground cracking in the Carrizo and Cholame stretches of the SAF. Our observations emphasize the importance of excavation at key locations to validate remote and ground-based measurements, and we advocate more geomorphic characterization for each site if excavation is not possible.

  8. Political Repressions in the Mongol Empire, Golden Horde and Other Turkic-Mongol States, and their Justifications (13th–16th cc.

    Directory of Open Access Journals (Sweden)

    R.Yu. Pochekaev


    Full Text Available In this article the author analyses the cases of political repressions in the Mongol Empire, Golden Horde and other Turkic-Mongol states of the 13th–16th centuries. Author investigates different types of repressions: against rivals during the struggle for the throne, officials who incurred the anger of monarchs, rebellious cities and their citizens. So, the political rivals often justified their right to the throne referring to the Great Yasa of Chinggis Khan, and hence, the punishment of the vanquished rivals usually was based as well on the Chinggis Khan’s principles of the “Law and Order”: ambiguity of these principles (since the Great Yasa, as it seems, was not a written code of laws but only a system of rules and principles proclaimed by Chinggis Khan or his successors, who attributed them to him allowed the winners to avenge their rivals following the formal legal norms. Thus, the charge of violation of the Great Yasa was a universal one allowing to solve the problem of of getting rid of a dangerous rival. The punishment of disgraced officials was justified by other arguments that differed from charges of rebellion of the Chinggisid family members. But Chinggisid rulers also used some “standard” accusations such as treason, support of usurpers, bribery. Since in most cases such acts also contradicted to the principles of the Great Yasa (as they were interpreted by the Chinggisids, the formula “put to yasa” was frequently used in verdicts on such cases. At last, we can also include the destruction of the resisting and insurgent cities in terms of political repressions. Reprisals against foreign cities that resisted the Mongol conquerors, was an integral part of the military strategy of terror facilitating the voluntary surrender of the following cities. In this case, the Chinggisids did not need any legal basis for the slaughter and destruction. However, in case of the rebellion of their own cities against the legitimate

  9. Para una historia fiscal de la Mallorca cristiana (siglos XIII-XIV

    Directory of Open Access Journals (Sweden)

    López Bonet, Juan Francisco


    Full Text Available Constituted like an independent kingdom, with the later aggregate of Ibiza -1235-, Majorca happens to integrate - 1276- in a set with own monarch divided between the Balearic islands (will also include built-in Menorca in 1294 and possessions in the French south, but it continues being fiscally independent, in spite of to have seen forced its first monarch to swear fidelity to the one of Aragon. It is when the Aragonese crown absorbs the kingdom of Majorca in 1349 that its fiscality happens to be employee of the necessities of the peninsular territories and is subordinated to outer companies that exceed their tax collecting capacity. The pressure of the monarch, surrounded in multiple outer warlike conflicts, the climatic and epidemics crises, the interests of their social groups and a high level of corruption, constitutes a conglomerate of causes that at the beginning of the five hundred ruins the Majorcan public finances, that are in situation of dependency of the creditors until entered century XIX.

    Constituida como reino independiente, con el agregado posterior de Ibiza –1235-, Mallorca pasa a integrarse –1276- en un conjunto con monarca propio dividido entre las islas Baleares (incluirá también Menorca incorporada en 1294 y posesiones en el sur francés, pero sigue siendo fiscalmente autónoma, a pesar de haberse visto obligado su primer monarca a jurar fidelidad al de Aragón. Es cuando el reino de Mallorca es absorbido por la corona aragonesa en 1349 que su fiscalidad pasa a ser dependiente de las necesidades de los territorios peninsulares y queda subordinada a empresas exteriores que sobrepasan su capacidad recaudatoria. La presión del monarca, envuelto en múltiples conflictos bélicos exteriores, las crisis climáticas y epidémicas, los intereses de sus grupos sociales y un elevado nivel de corrupción, constituyen un conglomerado de causas que a principios del quinientos arruina las finanzas públicas mallorquinas que

  10. King Chulalongkorn: biography and his activities in medicine and public health. (United States)

    Charulukananan, Somrat; Sueblinvong, Tada


    King Rama V, or Chulalongkorn, was the fifth monarch of the Chakri Dynasty. He was one of the most beloved of the Thai kings due to his many activities including abolishing slavery without bloodshed and especially his skillful diplomacy which succeeded in steering Siam out of the grips of the colonial powers. His activities also included reform of the administration of the kingdom according to the European model and in bringing Siam into the modern era with such exquisite skills that he is still vividly remembered today. His reign also saw many developments in medicine and public health. The King's role in these areas, however, were clouded by his more visible activities in politics and diplomacy. The result is that the Thai public learned rather little about his role in these areas. This article aims at collecting this and to show the King's very important role in modernizing medicine and public health in Siam.

  11. The Chivalric Order of the ‘Nodo’, or of the Holy Spirit, and the Patronage of Niccolò Orsini in the church of Santa Maria Jacobi in Nola

    Directory of Open Access Journals (Sweden)

    Cristiana Di Cerbo


    Full Text Available In the Angevin Kingdom of Naples, in the early fourteenth century the Orsini family initiated an urban enrichment in the Campanian city of Nola, that lasted for the entire century. This renovatio urbis focused on the representation of Orsini identity and authority in a variety of settings: during the 1354-1359, Count Niccolò, in fact, promoted the decoration of the conventual female church of Santa Maria Jacobi ordinis sanctae Clarae, making use of both secular and sacred themes, inspired by the medieval bestiaries, the courtly literature (the Lay of Aristotle, the Ovid's Metamorphoses and the st. Augustine's De Musica. The elaborate iconographic program, with its ties to the monarchical order of the Knot - or the Holy Spirit -, shows the role attributed to the nobility as champion of virtue and defensor of the Divine Order upon earth.

  12. Authority, Power and Reason of State in the Theater of Antonio Enríquez Gómez

    Directory of Open Access Journals (Sweden)

    Manuel Calderón Calderón


    Full Text Available Comedies of political content written by Antonio Enríquez Gómez reflect, through the analysis of its four recurring themes —the dichotomy between law and power, the reason of state, the law enforcement from a political point of view and forgiveness—, the outlines of the nascent Leviathan or modern state. Although the patriarchal relationship between the Monarch and his people is based on a iusnaturalist conception, love should be subject to the effectiveness in the exercise of power. The characters in this theatre sometimes hesitate between Aristotelian-Thomistic rationalism and Machiavellian-Hobbesian reason of state and, on other occasions, they oppose the people’s strength, risk and will to the divine reason of Law. In the context of the Hispanic Monarchy crisis, Enríquez Gómez’s critics are not limited to Government representatives and civil servants, but also include the Church and common people (Third State.

  13. El discurso directo en la Crónica Real Ccstellana del siglo XV

    Directory of Open Access Journals (Sweden)

    Luis Fernández Gallardo


    Full Text Available In the fifteenth Century, the Castilian royal chronicle reached a literary maturity that is evident in the use of direct speech. There is no linear evolution, but swings from the rich plurality of voices in García de Santa María´s chronicle, to the extreme selection of Barrientos, conditioned by various propaganda strategies. Enriquez del Castillo expresses the aspiration to eloquence of humanistic historiography. Nevertheless, it is only thanks to Fernando de Pulgar that full awareness of humanistic writing of history is taken, by means of the imitatio of its model, Livy. Humanistic eloquence became a means of exalting the monarchy of the Catholic Monarchs.

  14. History of Smallpox and Its Spread in Human Populations. (United States)

    Thèves, Catherine; Crubézy, Eric; Biagini, Philippe


    Smallpox is considered among the most devastating of human diseases. Its spread in populations, initiated for thousands of years following a probable transmission from an animal host, was concomitant with movements of people across regions and continents, trade and wars. Literature permitted to retrace the occurrence of epidemics from ancient times to recent human history, smallpox having affected all levels of past society including famous monarchs. The disease was officially declared eradicated in 1979 following intensive vaccination campaigns.Paleomicrobiology dedicated to variola virus is restricted to few studies, most unsuccessful, involving ancient material. Only one recent approach allowed the identification of viral DNA fragments from lung tissue of a 300-year-old body excavated from permafrost in Eastern Siberia; phylogenetic analysis revealed that this ancient strain was distinct from those described during the 20th century.

  15. [Study on the liver-protective and choleretic effect of zhizi baipi soup and its disassembled prescription]. (United States)

    Xiao, Xu; Zhu, Ji-Xiao; Luo, Guang-Ming; Li, Lei; Zhu, Yu-Ye; Zeng, Jin-Xiang; Wang, Xiao-Yun; Wu, Bo


    To investigate the effect of Zhizi Baipi soup and its disassembled prescription on protecting liver and improving choleresis and explore the regularity of Zhizi Baipi soup composition. The model of mouse liver injury induced by carbon tetraehlofide (CCl4) was used to observe the effects of Zhizi Baipi soup and its disassembled prescription by oral adminstration, the bile volume was determinied by common bile duct drainage. Zhizi Baipi soup and each treatment group with gardenia could significantly inhibit the increased serum ATL and AST activities, reduce liver MDA level, and significantly promote the bile flow and bilirubin in bile in normal rats. Zhizi Baipi soup has effects on protecting liver and increasing bile secretion, its monarch drug, gardenia plays an important role in the decoction, the effect of eliminating dampness and heat are mainly ascribed to the synergic effect of gardenia and phellodendron.

  16. The impact of Genetically Modified (GM) crops in modern agriculture: A review. (United States)

    Raman, Ruchir


    Genetic modification in plants was first recorded 10,000 years ago in Southwest Asia where humans first bred plants through artificial selection and selective breeding. Since then, advancements in agriculture science and technology have brought about the current GM crop revolution. GM crops are promising to mitigate current and future problems in commercial agriculture, with proven case studies in Indian cotton and Australian canola. However, controversial studies such as the Monarch Butterfly study (1999) and the Séralini affair (2012) along with current problems linked to insect resistance and potential health risks have jeopardised its standing with the public and policymakers, even leading to full and partial bans in certain countries. Nevertheless, the current growth rate of the GM seed market at 9.83-10% CAGR along with promising research avenues in biofortification, precise DNA integration and stress tolerance have forecast it to bring productivity and prosperity to commercial agriculture.

  17. Johann G. Herder: intellectual profile of an illuminated radical

    Directory of Open Access Journals (Sweden)

    Luis Gonzalo Díez Álvarez


    Full Text Available The intellectual origins of Nationalism and its idea of Culture found an unavoidable reference in Johann G. Herder’s work (1744-1803. Herder put forward an approach to Enlightenment contrary to the official one, critical of the policies of bureaucratic Reformism of Absolutism and Kantian Philosophical Rationalism. Herder’s take on Enlightenment holds a sense of History open to cultural diversity. Language and Culture emerge from Herder’s approach as the codes of a people’s world conceived of in a utopian way, beyond the logic of monarchic and aristocratic power. The nationalist idea of culture developed by the German thinker makes us reconsider, in a critical way, the Romantic origins of Nationalism.

  18. The specificity of folklore and mythological motifs in the novel “Tsar Maiden” by Vsevolod Solovyov

    Directory of Open Access Journals (Sweden)

    Lyapina Svetlana Mitrofanovna


    Full Text Available The article deals with folklore motifs in the novel by Vsevolod Solovyov “Tsar-maiden”, and reveals the link between this work and a magic tale. The author comes to the conclusion that the appeal to the image of the Tsar-maiden due to the desire of the writer to show the irrational spirit of pre-Petrine Russia, judgment of the people of the rulers of Imperial power. In the popular view of the nation the fact that the woman has become a monarch it was beyond their comprehension and considered a miracle akin to a fairy tale. Therefore, from Vsevolod Solovyov’s viewpoint, a fabulous image of the Tsar-maiden in the minds of the people coincided with the image of Princess Sophia.

  19. The body of the nation: government positions on physical education during the Brazilian monarchy. (United States)

    de Melo, Victor Andrade; Peres, Fabio de Faria


    In association with its nation building projects, the imperial government in Brazil under monarchic rule took some concrete actions based on proposals for physical education. The aim of this article is to investigate the meanings and significations attributed to this subject in the legislation and the annual reports issued by the Ministry of Business of the Empire (1831-1889), giving special attention to Rio de Janeiro. The approach to the subject in the sources researched demonstrates that the views of physical education took shape through a web of ideas that associated moral, health and civilization conceptions, in a bid to deal with the concrete circumstances of a newly independent peripheral nation with a bureaucratic structure in the process of formation.

  20. Los inventarios documentales del cabildo colonial de Medellín. Un proceso de racionalización hacia el ''buen gobierno''

    Directory of Open Access Journals (Sweden)

    Alfonso Rubio


    Full Text Available We focus since an academic field not much explored: the ''Archivistic History''. After a thorough survey and analysis of the documentation containing in the Historical Archive of Medellín, precisely in its fund called Cabildo, we show the concerns and needs of the colonial cabildo of Medellín by the organization and description of the documents in his archive through documentary inventories directed to ''good governance'', as claimed by the own documentation of the institution. The archive and the inventory construction that describe his documentary units, as part of administration and information Monarchic system, who should control the economics, socials and juridical issues, show us that the information is a strong factor of cohesion and permanence of all system, understanding that the information system is , in essence, a regulation and homogenization structure.

  1. Los indígenas de la Guajira en la independencia de las provincias caribeñas de la Nueva Granada: una aproximación

    Directory of Open Access Journals (Sweden)

    José Polo Acuña


    Full Text Available The involvement of the native Guajiros during the independence was the result of motley factors, determined by the specific type ofrelationship held by each indigenous group with the Creole society, the authorities and the sectors which favored the independence or those which supported the reinstatement of a monarchic order, as well as the degree of subjection or autonomy they maintained facing colonial rule within the context of its collapse and the existing needs of the communities. We focused on the indigenous groups ofthe southern peninsula of the Guajira, known also as Guajira Abajo, for two reasons: first, because this area presents the most markedmestization between Indigenous people and Creoles, and second, because the sources insufficiently depict the activities of the natives inhabiting the extreme North of the peninsula, the Guajira Arriba

  2. The Armenian enigma

    Directory of Open Access Journals (Sweden)

    Aptin Khanbaghi


    Full Text Available The Armenians’ financial elevation under the Safavids has been depicted as sudden by a number of historians. This article questions this unforeseen success and investigates the development of the Armenians’ commercial network before their being pursued and deported by the Safavid monarchs in the 16th and 17th centuries. It surveys the scattering of Armenians in West Asia from the beginning of the mediaeval period. Although the majority of old sources do not refer to the Armenians’ involvement in trade before the 16th century, they refer to their geographical presence and discuss their political association with various rulers and communities in West Asia. By puzzling out the Armenians ties with various people and regions and following their trek from the Byzantine times to the Ottoman and Safavid era, this article extracts the keystone to the Armenians’ ascendancy in the history of world trade.

  3. Aristocracia y monarquía en los reinos de Castilla y León: el caso de la familia Lara

    Directory of Open Access Journals (Sweden)

    Simon Doubleday


    Full Text Available Between the twelfth century and the fourteenth, there was a fundamental change in the basis of the Lara family s power. At the outset, their influence derived primarily from personal access to, and control over, the monarch; throughout the twelfth century, the family s wealth rested heavily on possession of the royal offices known as tenencias, which were the principal fruit of courtly influence. But from the middle of the thirteenth century, the aftershocks of the reconquests of Seville and Córdoba, and the expansion of royal administration, meant that symbiosis was increasingly supplanted by a confrontational political climate marked by an acceleration of patrimonial accumulation by aristocratic lineages such as the Laras. The new structure of aristocratic power would long outlast the Lara family itself, surviving well into the modem age and finding expression in a culture in which, more than in any other western European society, the authority and values of nobility were dominant.


    Directory of Open Access Journals (Sweden)

    S. Mehralizadeh


    Full Text Available The historical garden complex of Mofakham was built in the late nineteenth century as a formal pavilion garden in the city of Bojnourd in north-east Iran for the Persian monarch's local governor (hakem of the time. At least five main buildings and other secondary buildings of the garden complex have disappeared over the years. Although the surrounding context has changed totally, the axial route of the garden and two buildings of the complex remain. The documentation includes: 1. A precise two dimensional map of the complex (Figures 1–3, 2. A number of three dimensional images based on total 3D making (Images 4–5, 3. A 1/150 model

  5. Die Jerusalemse tempelkuItus se huweliksmaatreel versus Christelike waardes

    Directory of Open Access Journals (Sweden)

    A.G. van Aarde


    Full Text Available The marriage arrangements in the Jerusalem Temple cult in opposition to Christian values. Equal access for everyone to God's grace in an unmediated way is a central aspect of Jesus' presentation of the Kingdom of God. Inclusivity and egalitarity should be regarded as essential aspects of Christian selfunderstanding. This article aims to show how these values stood in opposition to the marriage arrangements in the Jerusalem Temple cult. Marriage strategies during the patriarchal and monarchical periods prior to the first-century Jerusalem Temple cult are also briefly touched upon. In a following up article the author will argue that Paul's use of the notion "adopted as children of God" should be seen as an expression of the Christian values advocated within an inclusive and egalitarian community.

  6. Conflict between the Monarchy and the Aristocracy for Control of the almojarifazgos and Other Customs Duties on the Atlantic Coast of Andalusia in the Reign of the Catholic Kings

    Directory of Open Access Journals (Sweden)

    José Damián González Arce


    Full Text Available Between the conquest of the Muslims, in the middle of the thirteenth century, and the first half of the fifteenth century, virtually the entire the Atlantic coast of Andalusia was handed over to powerful lords by the monarchy. They were for the most part great aristocrats who hoped to collect the tax on foreign trade by sea known as the almojarifazgo in their domains, along with other revenues. This tax theoretically belonged to the Crown. Its collection was centralized in the nearby city of Seville and its customs house. After several centuries of monarchical weakness, in the late fifteenth century the Catholic Kings managed, through numerous lengthy judicial processes, to recover these levies for the royal treasury and to almost completely restore control of taxation to the Crown.

  7. Regele Mihai între „est” și „vest”. Percepții, poziționări, simboluri (King Michael between east and west. Perceptions, positionings, symbols

    Directory of Open Access Journals (Sweden)

    Alexandru MURARU


    Full Text Available The article investigates three strategic directions within the political perceptions of Romanian King Michael involving the East/West paradigm. The article will try to highlight the main positioning, both in Exile and the complicated recent history, immediately after the December 1989 events. King Michael’s relations with Western powers and attitude throughout Romania after 1989, the fate of Bessarabia as a direct „confrontation” between East -West and the significances of the „August 23” Act in reshaping the post-revolutionary memorial debate, represent three fundamental perspectives in understanding the „East” and „West” thinking of the Romanian monarch. These three themes, rich in meanings, have three major connotations: moral, geographic and memorials. Each of them are explained and decrypted with many examples, quotes, and historical and geographical references.

  8. Kings, Nobles, and Frontier: Violence and Kinship around the Galician-Portuguese border (12th-13th centuries

    Directory of Open Access Journals (Sweden)

    Inés Calderón Medina


    Full Text Available The birth of the frontier between Galicia and Portugal was an arduous process. The control over this area relied, to a great extent, on the success of the kings of Leon and Portugal in assuring the local nobles’ loyalty and service. The role played by the local nobility in the creation and military defense of this border can be analysed throughout chronicles and diplomas. These sources cast light on the nobility’s key military role in southern Galicia and the political consequences derived from these nobles’ loyalty or infidelity. The kings of Portugal and Leon actively seek the local nobility’s support; however they also relied on royal marriages in order to solve border conflicts. In that regard, disputed landholdings and castles were given to the Portuguese monarchs as part of the Leonese queens’ dowry. Some nobles were chosen as executors of these marriages agreements, so they acted as intermediaries between both kings.

  9. Lope de Vega y el poder monárquico: una puesta al día

    Directory of Open Access Journals (Sweden)

    Marcella Trambaioli


    Full Text Available The essay tries to outline the ‘state-of-the-art’ about the relationship between Lope deVega and the contemporary monarchic power. After some early studies merely concerning the politicaltheory present inside the texts, the scholars have started to analyze his historical theater in relation withhis construction of the Spanish Empire mythology, even if only recently some of them have realized thatthis glorification of both the Austrias and the Spanish nobles gets on with Lope’s self-promotionstrategies, in order to obtain an official post at court. On the contrary, some scholars, mainlyAnglosaxon, have tried to interpret his works in a critical and satyrical perspective, without consideringthe necessary contextualization of his theater. The reception theory may help studying this controversialmatter, only if it tries to match in the right way both the historical and cultural context of the authorand the cultural background of the contemporary audience.

  10. constitucional. A adesão das câmaras no processo de autonomização do Brasil

    Directory of Open Access Journals (Sweden)

    Iara Lis Schiavinatto


    Full Text Available The article is about the work of municipal governments as autonomous political bodies in the Brazilian constitutional process in 1820-26. Emphasis is placed on the historiographical debate of which this research is part, and the specific meaning of the term “adesão” developed in the links between municipal government, localities, and the Vintista Cortes in Lisbon, and the court in Rio de Janeiro. This research thus recovers social memory and memories of the empire, of local social, ethnic and political tensions, and ways of celebrating the “adesão” political pact that simultaneously linked the foundation of independent Brazil and constitutional monarchic rule.

  11. [Substance basis research on Chinese materia medica is one of key scientific problems of inheriting, development and innovation of Chinese materia medica]. (United States)

    Yang, Xiu-wei


    The compound Chinese materia medica is the medication pattern of the traditional Chinese medicine for the disease prevention and treatment. The single Chinese materia medica (mostly in decoction pieces) is the prescription composition of the compound Chinese materia medica. The study of the effective substance basis of Chinese materia medica should be based on the chemical compositions of the compound Chinese materia medica as an entry point considering the different status of "Monarch, Minister, Assistant, and Guide" for a certain single Chinese materia medica in the different compound Chinese materia medica while substance basis research of a certain single Chinese materia medica should be a full component analysis as well as both stable and controllable quality. Substance basis research on Chinese materia medica is one of key scientific problems of inheriting, development and innovation of Chinese materia medica.

  12. Первые военные действия Великой Северной войны или что предшествовало битве под Нарвой (1700, в свете польской и шведской историографий

    Directory of Open Access Journals (Sweden)

    Katarzyna Wagner


    Full Text Available After the death of Charles XI the throne of Sweden took his 15-year-old son, Charles XII, and it was the beginning of Sweden’s international problems with other countries. Monarchs of Denmark, Russia and Saxony started negotiations, because they wanted to sign the anti-Swedish treaty. Every of them had plans to divide Sweden’s territory and took a part for itself. Nobody expected that Charles XII with almost no military experience will enforce the Danes to submit to the Peace of Travendal in August 1700, Saxony overcame after the siege of Riga and Russia during the battle of Narva. After the battle of Narva, where the Russians suffered a crushing defeat, Charles XII turned his attention upon the other powerful neighbor – Frederic I of Saxony (who was also the king of Poland as August II.

  13. "Letters to a lady": questions of gender and the dissemination of Darwinism in Brazil

    Directory of Open Access Journals (Sweden)

    Moema de Rezende Vergara

    Full Text Available Recent works on the history of scientific dissemination in Brazil have practically ignored the category of gender in their analysis. Thus, this article is intended to contribute to the content of a study of a specific practice of scientific disclosure of the 1800s, as exemplified by"Letters to a lady" written by Rangel S. Paio and published in the "O Vulgarizador". In this case, the concept of gender makes it possible to understand the tensions between the masculine and feminine appearing in a series of letters meant for purposes of scientific dissemination, in which the author himself anticipated their gender conten, insofar as he directed his attention to a public of 'ladies' in the Second Monarchical reign in Brazil.

  14. SNS vil høre vores kommentarer til spørgsmålene. Spørgsmål vedr. Cumulative long-term effects of GMO. Modtaget 16-01-2006, deadline 23-01-2006, svar 23-01-2006

    DEFF Research Database (Denmark)

    Strandberg, Morten Tune; Kjellsson, Gøsta; Damgaard, Christian


    where environmental risks of non-GM-crops are assessed in parallel fashion to GM-crops have a point regarding that the environmental risks of non-GM crops should also be assessed. 11) Are you aware of any reports of unexpected effects of the release of a GMO? Re 11): This is a matter of definition. We...... with larvae which feed on plants in field surroundings. Furthermore, contamination of non-GM crops by seed spillage or accidental mixing of seed parties, which are perhaps not unexpected, has been reported from USA, Canada and of cause the Teosinte maize from Mexico. "...... are not aware of any major unexpected environmental effects of released crops. The potential adverse effects of Bt-pollen on monarch butterfly larvae, which turned out to be of minor importance in actual field situations, was an important reminder of a general problem to small populations of butterflies...

  15. Napoléon mémorialiste, entre gloire et insignifiance ou « le plus grand homme dévoré d’inquiétude » // Napoleon memorialist, between grandeur and insignificance

    Directory of Open Access Journals (Sweden)

    Aleš Pohorský


    Full Text Available During his exile at island of Saint Helena, Napoleon is condemned to look back upon his life. Le Mémorial, dictated by Napoleon and edited by Las Cases, is the very foundation of Napoleonian legend. Napoleon mentions the ingratitude of those monarchs which he installed to the throne himself and who later denied him. The work is the bases of the legend that oscillates between monarchy and citizenship, the king and the nation and the dictatorship and liberty. As a literary work Le Mémorial inspired Musset, Chateaubriand, Mme de Staël, Bourrienne, la duchesse d’Abrantès, Balzac, Mme de Rémusat and many others. It has also influenced all European cultures, including Czech literature. The one, who “changed the world” died in solitude and obscurity, in deep meditation about his own failure.

  16. Human Rights in the West

    Directory of Open Access Journals (Sweden)

    Jorgen S. Nielsen


    Full Text Available One of the areas of conflict between Islam and the West in today’s world is the concern for human rights. This has sometimes been criticized in the Muslim world as a form of neo-imperialism. It is therefore necessary to understand the various dimensions of human rights, and the various phases through which this concern has grown. In the earliest form, it was an assertion of the rights of the landed aristocracy against those of the monarch. The French revolution, with its emphasis on "liberty, equality and fraternity," for all individuals, provided another dimension. There were many occasions on which individual and organized religion came into conflict during the Middle Ages. The experience of World War II, particularly the atrocities of the Nazis, led to the internationalization of individual rights.

  17. Optimization Method of Fusing Model Tree into Partial Least Squares

    Directory of Open Access Journals (Sweden)

    Yu Fang


    Full Text Available Partial Least Square (PLS can’t adapt to the characteristics of the data of many fields due to its own features multiple independent variables, multi-dependent variables and non-linear. However, Model Tree (MT has a good adaptability to nonlinear function, which is made up of many multiple linear segments. Based on this, a new method combining PLS and MT to analysis and predict the data is proposed, which build MT through the main ingredient and the explanatory variables(the dependent variable extracted from PLS, and extract residual information constantly to build Model Tree until well-pleased accuracy condition is satisfied. Using the data of the maxingshigan decoction of the monarch drug to treat the asthma or cough and two sample sets in the UCI Machine Learning Repository, the experimental results show that, the ability of explanation and predicting get improved in the new method.

  18. Reflection in Russian public opinion accession to the throne of Emperor Alexander II

    Directory of Open Access Journals (Sweden)

    Maxim N. Krot


    Full Text Available The article deals with the perception of Russian society coming to the throne of Emperor Alexander II in 1855. The author analyzes the internal and external situation in Russia in this period, identifying the most significant factors that influenced the public's attitude to the new monarch, and the expectations that were associated with his coming to power. The article draws parallels between the initial period of the reign of the two liberal Russian monarchs of the XIX century – Alexander I and Alexander II, most clearly expressed not so much in the circumstances of their accession, but in the public mood that prevailed in the country. It presents a broad picture of the "awakening" of public consciousness and activity after the stagnation and oppression of the preceding reign, which was reflected primarily in an effort to think freely and to discuss pressing issues facing the country, as well as contribute to the supreme power in the modernization of social and political relations, awareness where necessary, at this time it becomes clearly. The author identifies semantic levels of social consciousness of the period, indicating the particular circumstances that have shaped them. Much attention is paid to the influence of Russian society of the Crimean War, which in many ways was the starting point of social upheaval transformed in 1855 from a national-patriotic in the socio-political. The author points out that at this time has unique conditions for overcoming the contradictions between state and society and their successful cooperation in the reform of the country. This suggests a significant, perhaps - the crucial role of the public in the selection of the vector conversion of Russia, which has become a feature of the reign of Alexander II.

  19. Magna Carta And Its Significant Role For Rule Of Law In The Republic Of Macedonia

    Directory of Open Access Journals (Sweden)

    Shumanovska-Spasovska Ivana


    Full Text Available One of the most important and famous historical documents from the English legal and constitutional legacy is the Magna Carta Libertatum. Signed and sealed in the year 1215 the Magna Carta is further on viewed as the sole inception of the idea of limiting the power of the ruler trough legal rules. That limitation is to be made with legal rules that are binding for everyone, even the monarch. Therefore, the Great Charter is viewed as the first document signed by a monarch with which, the principle of supremacy of the law is set out. That supremacy of the law has been further on developed by eminent scholars and practitioners, eventually leading to the development of the concept of rule of law. Rule of law, as a concept, means that the royal authority (or the executive branch of power is going to be inferior to the law. However, this concept means a lot more than simply that. Unlike the principle of legal state, the rule of law is closely linked to justice, separation of powers and legal certainty. All of these concepts are actually prerequisites for its existence. That is why each of them is separately examined and elaborated. Furthermore, as one of the most important principles the rule of law had a great influence on the constitutional (and legal systems around the world. Since the Republic of Macedonia strives to become a democratic state where the rule of law is established and developed it is important to elaborate the influence of this principle in it. Therefore, the research gravitates over the principle of rule of law in the Republic of Macedonia.

  20. Como en un espejo, la recepción europeade la Constitución de Cádiz

    Directory of Open Access Journals (Sweden)

    Miguel Herrero De Miñón


    Full Text Available La Constitución de Cádiz de 1812 tuvo una amplia difusión en el Viejo Continente por la escasez de modelos constitucionales —Constitución americana 1787, francesa 1791 y británica— por el halo heroico de la resistencia antifrancesa y por su pseudohistoricismo — constitución histórica española—. Sin embargo, el balance político de su recepción presenta una oscura coloración pudiendo calificarse de fracaso, debido a la rígida separación de poderes que persiguió la capitidisminución del ejecutivo monárquico, el monocameralismo que desconocía el marcado carácter estamental de la sociedad española de la época en la que tres cuartas partes de la tierra estaban en manos del monarca, la nobleza y el clero, y el exacerbado centralismo con los funestos perjuicios que irrogó sobre la emancipación de la América española. The Constitution of Cadiz of 1812 had an ample diffusion in the Old Continent by the shortage of constitutional models —American Constitution 1787, French 1791 and British by the pull ahead heroic of the French resistance and by its pseudohistoricismo— Spanish historical constitution. Nevertheless, the political balance of its reception presents/displays a dark coloration being able to be described as failure, due to the rigid separation of powers that persecuted diminution of the executive authority of the monarch, the monocameralismo that did not know the noticeable estamental character of the Spanish society of the time in whom three fourth parts of the Earth were into the hands of the monarch, the nobility and the clergy, and the strong centralism with the unfortunate damages that it caused on the emancipation of Spanish America.

  1. Investigation of the mineral potential of the Clipper Gap, Lone Mountain-Weepah, and Pipe Spring plutons, Nevada

    International Nuclear Information System (INIS)

    Tingley, J.V.; Maldonado, F.


    The Clipper Gap pluton, composed mostly of quartz monzonite with minor granite, granodiorite, and crosscutting alaskite dikes, intrudes Paleozoic western facies strata. A narrow zone of contact metamorphism is present at the intrusive-sediment contact. No mineral production has been recorded from Clipper Gap, but quartz veins containing gold-silver-copper mineral occurrences have been prospected there from the late 1800's to the present. Areas of the Lone Mountain-Weepah plutons that were studied are located in Esmeralda County about 14 km west of Tonopah, Nevada. At Lone Mountain, a Cretaceous intrusive cuts folded Precambrian and Cambrian sediments. Lead-zinc ores have been mined from small replacement ore bodies in the Alpine district, west of Lone Mountain. Copper and molybdenum occurrences have been found along the east flank of Lone Mountain, and altered areas were noted in intrusive outcrops around the south end of Lone Mountain. Mineral occurrences are widespread and varied with mining activity dating back to the 1860's. The Pipe Spring pluton study area is flanked by two important mining districts, Manhattan to the north and Belmont to the northeast. Mining activity at Belmont dates from 1865. Activity at Manhattan was mainly between 1907 and 1947, but the district is active at the present time (1979). Four smaller mining areas, Monarch, Spanish Springs, Baxter Spring, and Willow Springs, are within the general boundary of the area. The Pipe Spring pluton study area contains numerous prospects along the northern contact zone of the pluton. Tungsten-bearing veins occur within the pluton near Spanish Springs, with potential for gold-tungsten placer in the Ralston Valley. Nickel and associated metals occur at Willow Spring and Monarch Ranch, where prospects may be associated with the margin of the Big Ten Peak Caldera

  2. La Orden Militar de Alcántara y la monarquía castellana durante los primeros Trastámaras (1369-1390

    Directory of Open Access Journals (Sweden)

    Novoa Portela, Feliciano


    Full Text Available This work tries to analyze the relations of Alcantara's order with the first two kings of the of dynasty Trastámara, concretly with Enrique II and Juan I. During this period, the Order will suffer a substantial change that is translated, on the one hand, in the purpose of his equivocal politics with the Castilian rown and, for other one, in the decrease of his institutional autonomy that has in the control of the appointment of the masters for his element more significant. All this process places inside a politics led to end for both monarches, whose aim was going to strengthen the power royal and to a progressive increase of means and of political attributions in hands of the monarchic.

    Este artículo trata de analizar las relaciones de la Orden de Alcántara con los dos primeros reyes de la dinastía Trastámara, concretamente, con Enrique II y Juan I. Durante este período, la Orden sufre un cambio sustancial que se traduce, por una parte, en el fin de su política equívoca con la Corona castellana y, por otra, en la disminución de su autonomía institucional, que tiene su elemento más significativo en el control de la designación de sus Maestres. Todo este proceso se sitúa en una política desarrollada por los dos soberanos cuyo objetivo era reforzar el poder real y el aumento

  3. Seed dormancy and germination vary within and among species of milkweeds (United States)

    Kaye, Thomas N; Sandlin, Isaac J; Bahm, Matt A


    Abstract Pollinators in general and monarch butterflies in particular are in decline due to habitat loss. Efforts to restore habitats for insects that rely on specific plant groups as larvae or adults depend on the ability of practitioners to grow and produce these plants. Monarch larvae feed exclusively on milkweed species, primarily in the genus Asclepias, making propagation and restoration of these plants crucial for habitat restoration. Seed germination protocols for milkweeds are not well established, in part due to the large number of milkweed species and conflicting reports of seed dormancy in the genus. We tested for seed dormancy and the optimum period of cold stratification in 15 populations of A. speciosa and 1–2 populations of five additional species, including A. asperula, A. fascicularis, A. subulata, A. subverticillata and A. syriaca. We exposed seeds to cold (5 °C) moist conditions for 0, 2, 4, 6 and 8 weeks and then moved them to 15 °C/25 °C alternating temperatures. In A. speciosa, dormancy was detected in eight populations, and this dormancy was broken by 2–4 weeks of cold stratification. The remaining seven populations showed no dormancy. Seed dormancy was also detected in two populations of A. fascicularis (broken by 4–6 weeks of cold stratification) and a single population of A. syriaca (broken by 2 weeks of cold stratification). No dormancy was detected in A. asperula, A. subulata or A. subverticillata. Seed dormancy appears to be widespread in the genus (confirmed in 15 species) but can vary between populations even within the same species. Variation in seed dormancy and cold stratification requirements within and among Asclepias species suggests local adaptation and maternal environments may drive seedling ecology, and that growers should watch for low germination and use cold stratification as needed to maximize seed germination and retain genetic variability in restored populations. PMID:29593856

  4. A genetically-based latitudinal cline in the emission of herbivore-induced plant volatile organic compounds. (United States)

    Wason, Elizabeth L; Agrawal, Anurag A; Hunter, Mark D


    The existence of predictable latitudinal variation in plant defense against herbivores remains controversial. A prevailing view holds that higher levels of plant defense evolve at low latitudes compared to high latitudes as an adaptive plant response to higher herbivore pressure on low-latitude plants. To date, this prediction has not been examined with respect to volatile organic compounds (VOCs) that many plants emit, often thus attracting the natural enemies of herbivores. Here, we compared genetically-based constitutive and herbivore-induced aboveground vegetative VOC emissions from plants originating across a gradient of more than 10° of latitude (>1,500 km). We collected headspace VOCs from Asclepias syriaca (common milkweed) originating from 20 populations across its natural range and grown in a common garden near the range center. Feeding by specialist Danaus plexippus (monarch) larvae induced VOCs, and field environmental conditions (temperature, light, and humidity) also influenced emissions. Monarch damage increased plant VOC concentrations and altered VOC blends. We found that genetically-based induced VOC emissions varied with the latitude of plant population origin, although the pattern followed the reverse of that predicted-induced VOC concentration increased with increasing latitude. This pattern appeared to be driven by a greater induction of sesquiterpenoids at higher latitudes. In contrast, constitutive VOC emission did not vary systematically with latitude, and the induction of green leafy volatiles declined with latitude. Our results do not support the prevailing view that plant defense is greater at lower than at higher latitudes. That the pattern holds only for herbivore-induced VOC emission, and not constitutive emission, suggests that latitudinal variation in VOCs is not a simple adaptive response to climatic factors.

  5. The evolution of the Muslim monarchy in El Príncipe constante

    Directory of Open Access Journals (Sweden)

    Ricardo Enrique Castells


    Full Text Available Abstract:Don Fernando, the main character of El príncipe constante, has received most of the attention of the specialists in the works of Calderón, but many critics had also analyzed the extraordinary transformation of the King of Fez, but with significant differences of points of views and conclusions. The Muslim monarch treats the captive prince with exquisite courtesy as long as he thinks that the prince can be exchanged for the city of Ceuta. But at the end he behaves like a tyrant as soon as he realized that don Fernando refuses the exchange. This article uses the Prospect Theory of Daniel Kahneman and Amos Tversky to analyze the transformation of the Muslim monarch with the purpose of finding a critical approach that could explain a rather paradoxical or at least incoherent behavior. Resumen:Don Fernando, el protagonista de El príncipe constante, ha recibido la mayor parte de la atención por parte de los estudiosos de la obra de Calderón, pero muchos críticos también han analizado la extraordinaria transformación del rey de Fez, aunque con una marcada diferencia de opiniones y conclusiones. El monarca musulmán atiende al príncipe cautivo con suma cortesía mientras que cree que lo puede cambiar por la ciudad de Ceuta, pero se convierte en un tirano absoluto una vez que don Fernando rechaza el truque que ha arreglado su hermano Enrique. El presente trabajo emplea la teoría de la perspectiva de Daniel Kahneman y Amos Tversky para analizar la transformación del monarca musulmán, con el propósito de encontrar un acercamiento crítico que explique un comportamiento aparentemente paradójico o hasta incoherente.

  6. Interspecific variation in resistance of Asian, European, and North American birches (Betula spp.) to bronze birch borer (Coleoptera: Buprestidae). (United States)

    Nielsen, David G; Muilenburg, Vanessa L; Herms, Daniel A


    Bronze birch borer (Agrilus anxius Gory) is the key pest of birches (Betula spp.) in North America, several of which have been recommended for ornamental landscapes based on anecdotal reports of borer resistance that had not been confirmed experimentally. In a 20-yr common garden experiment initiated in 1979 in Ohio, North American birch species, including paper birch (Betula papyrifera Marshall), 'Whitespire' gray birch (Betula populifolia Marshall), and river birch (Betula nigra L.), were much more resistant to bronze birch borer than species indigenous to Europe and Asia, including European white birch (Betula pendula Roth), downy birch (Betula pubescens Ehrh.), monarch birch (Betula maximowicziana Regel), and Szechuan white birch (Betula szechuanica Jansson). Within 8 yr of planting, every European white, downy, and Szechuan birch had been colonized and killed, although 100% of monarch birch had been colonized and 88% of these plants were killed after nine years. Conversely, 97% of river birch, 76% of paper birch, and 73% Whitespire gray birch were alive 20 yr after planting, and river birch showed no evidence of colonization. This pattern is consistent with biogeographic theory of plant defense: North American birch species that share a coevolutionary history with bronze birch borer were much more resistant than naïve hosts endemic to Europe and Asia, possibly by virtue of evolution of targeted defenses. This information suggests that if bronze birch borer were introduced to Europe or Asia, it could threaten its hosts there on a continental scale. This study also exposed limitations of anecdotal observation as evidence of host plant resistance.

  7. De Atanarico a Valia: Aproximación a los orígenes de la monarquía visigoda

    Directory of Open Access Journals (Sweden)

    María del Rosario VALVERDE CASTRO


    Full Text Available RESUMEN: Durante el periodo en que los visigodos permanecen fuera de las fronteras del Imperio romano, paralelamente al proceso de destribalización social y operando al mismo tiempo como causa y efecto, surge una forma de gobierno, la monarquía, caracterizada por ser electiva, no permanente y por desempeñar funciones esencialmente militares. A partir del asentamiento en territorios imperiales, los monarcas visigodos se convierten en reyes federados y se inicia un proceso de ampliación de las atribuciones regias que acaba modificando la misma naturaleza de la institución monárquica. Al final del periodo migratorio, la monarquía visigoda se ha transformado en el órgano central y permanente de gobierno y se tiende a imponer la herencia como sistema sucesorio.ABSTRACT: During the time the visigoths stayed away from the frontiers of the Roman Empire, a new form of government appeared, running parallel to the process of social destribalization, and operating as causa and effect at the same time: the monarchy, which will be elective and no permanent and will carry out military functions. Once the visigoths are settled on imperial territories, their monarchs turn into federated kings, and a proccess of extension or royal atri- butions is to be started; this process will finally modifícate the own nature of the monarchical institution. By the end of the migratory period, the visigothic monarchy has changed into the central and permanent government body and tends to impose the heritage as the successory system.

  8. El papel de los concilios visigodos en la defensa de los intereses nobiliarios frente al rey

    Directory of Open Access Journals (Sweden)

    Pino Abad, Miguel


    Full Text Available Since 633, a number of mechanisms to safeguard aimed at members of noble estate of continuous samples of royal arbitrariness coming suffering were adopted. In that year, the Fourth Council of Toledo, which was set aside, the prohibition of various monarchs take ownership of the goods, by way of thanks for services rendered, had been donated previously met by his predecessors on the throne. Similarly, it was ordered that henceforth, no monarch could know as sole judge and sentence those cases against the nobles and had rigged the death penalty and/or forfeiture of property. Measures found repeated in successive councils, demonstrating that were consistently unfulfilled, which makes sense if you take into consideration the permanent seizure that was immersed in the Visigothic kingdom.Desde el año 633, se adoptaron una serie de mecanismos tendentes a poner a salvo a los miembros del estamento nobiliario de las continuas muestras de arbitrariedad regia que venían padeciendo. En el citado año, se reunió el Concilio IV de Toledo, en el que se estableció, de un lado, la prohibición de que los distintos monarcas se apropiaran de los bienes que, a modo de agradecimiento por los servicios prestados, habían sido donados previamente por sus predecesores en el trono. Igualmente, quedó ordenado que, en adelante, ningún monarca pudiese conocer y sentenciar como juez único aquellas causas seguidas contra los nobles y que llevaban aparejadas la pena de muerte y/o confiscación de bienes. Medidas que encontramos repetidas en sucesivos concilios, lo que demuestra que fueron sistemáticamente incumplidas, algo lógico si se tiene en consideración la permanente convulsión en que estuvo inmerso el reino visigodo.

  9. Autobiografie v českém středověku

    Directory of Open Access Journals (Sweden)

    Marie Bláhová


    Full Text Available This is an overview of the first instances of the realization of personality in Bohemia, which appear in historical literature from the 12th century and lead to the essay on medieval autobiographies. Although in the Middle Ages autobiographies were rare both in Bohemia as well as in other countries within the Latin cultural circle we are able to identify three significant autobiographies from different social environments, of different literary type and with different functions within two centuries of the late medieval period: First, the autobiography of Charles IV. This was the autobiography of a monarch written as a reflection of a Prince aimed at the author’s heirs to provide an image of the upbringing and behaviour of an exemplary monarch. The second text of this type is the autobiographical letter of former Archbishop of Prague, Jan of Jenštejn, in which the author used his own destiny to explain his political failure. At the very end of the medieval period, Christoph of Tyn, a minor nobleman who gained social success in the Emperor’s army and in diplomatic services, wrote an autobiography, where he wanted to show his descendants and future heirs how to increase the family estates legally, through honest endeavour so that they would not be ashamed of their heritage and doubt its respectable provenace. All of the authors mentioned had their own particular reasons for writing an autobiography. Naturally, all of these autobiographies are subjective, the narration is tailored to its purpose – political goals, justification of one’s failure, “substantiation” for and expression of pride in legally gained property.

  10. Le contrôle de l’activité législative de la nation en 1789 : l’opinion de Dupont de Nemours

    Directory of Open Access Journals (Sweden)

    Anthony Mergey


    Full Text Available On the eve of the French Revolution, Dupont de Nemours became infatuated with constitutional issues that had hitherto been the exclusive domain of some of his fellow physiocrats, in particular Le Mercier de La Rivière. Endorsing the ideas of the time, he was convinced of the necessity of creating a new political regime that would take the features of a moderate monarchy. The prince would thus be submitted to constitutional rules, sovereignty transferred to the nation and the legislative power exercised by its representatives. However, even if sharing authority between the nation and the king was then claimed, the representatives of the nation should not exercise their prerogatives outside a clearly established, defined constitutional framework. Partly consistent with the original physiocratic theories, Dupont de Nemours identified constitutional norms that should be promoted and protected from abuses of the legislature. With this aim in mind, he didn’t plan to resort to a jurisdictional authority, because abuses of Ancien Régime parliaments were still live memories. He decided, in an original way, to entrust the heavy burden of controlling legislative activity to political instances, namely, successively, the people and the monarch. Quite naturally, these suggestions, expressed a few months apart, exposed various forms of control. The examination carried out by the nation appeared as a prerequisite and tended to verify the constitutionality of the law, while the one carried out by the monarch through his veto, as holder of the executive power, was more of a control of opportunity devoid of binding. Although the omnipotence of the revolutionary legicentrism seemed incompatible with such a system of verification of the norm, it remains that Dupont de Nemours was part of a current which considerably developed during the XVIIIth century and which found a certain resonance in the decade following the Revolution.

  11. Vibroseis Monitoring of San Andreas Fault in California

    Energy Technology Data Exchange (ETDEWEB)

    Korneev, Valeri; Nadeau, Robert


    A unique data set of seismograms for 720 source-receiver paths has been collected as part of a controlled source Vibroseis experiment San Andreas Fault (SAF) at Parkfield. In the experiment, seismic waves repeatedly illuminated the epicentral region of the expected M6 event at Parkfield from June 1987 until November 1996. For this effort, a large shear-wave vibrator was interfaced with the 3-component (3-C) borehole High-Resolution Seismic Network (HRSN), providing precisely timed collection of data for detailed studies of changes in wave propagation associated with stress and strain accumulation in the fault zone (FZ). Data collected by the borehole network were examined for evidence of changes associated with the nucleation process of the anticipated M6 earthquake at Parkfield. These investigations reported significant traveltime changes in the S coda for paths crossing the fault zone southeast of the epicenter and above the rupture zone of the 1966 M6 earthquake. Analysis and modeling of these data and comparison with observed changes in creep, water level, microseismicity, slip-at-depth and propagation from characteristic repeating microearthquakes showed temporal variations in a variety of wave propagation attributes that were synchronous with changes in deformation and local seismicity patterns. Numerical modeling suggests 200 meters as an effective thickness of SAF. The observed variations can be explained by velocity 6 percent velocity variation within SAF core. Numerical modeling studies and a growing number of observations have argued for the propagation of fault-zone guided waves (FZGW) within a SAF zone that is 100 to 200 m wide at seismogenic depths and with 20 to 40 percent lower shear-wave velocity than the adjacent unfaulted rock. Guided wave amplitude tomographic inversion for SAF using microearthquakes, shows clearly that FZGW are significantly less attenuated in a well-defined region of the FZ. This region plunges to the northwest along the

  12. Fast Computation of Ground Motion Shaking Map base on the Modified Stochastic Finite Fault Modeling (United States)

    Shen, W.; Zhong, Q.; Shi, B.


    ground shaking intensity, and the results of the comparisons between the simulated and observed MMI for the 2004 Mw 6.0 Parkfield earthquake, the 2008 Mw 7.9Wenchuan earthquake and the 1976 Mw 7.6Tangshan earthquake is fairly well. Take Parkfield earthquake as example, the simulative result reflect the directivity effect and the influence of the shallow velocity structure well. On the other hand, the simulative data is in good agreement with the network data and NGA (Next Generation Attenuation). The consumed time depends on the number of the subfaults and the number of the grid point. For the 2004 Mw 6.0 Parkfield earthquake, the grid size we calculated is 2.5° × 2.5°, the grid space is 0.025°, and the total time consumed is about 1.3hours. For the 2008 Mw 7.9 Wenchuan earthquake, the grid size calculated is 10° × 10°, the grid space is 0.05°, the total number of grid point is more than 40,000, and the total time consumed is about 7.5 hours. For t the 1976 Mw 7.6 Tangshan earthquake, the grid size we calculated is 4° × 6°, the grid space is 0.05°, and the total time consumed is about 2.1 hours. The CPU we used is 3.40GHz, and such computational time could further reduce by using GPU computing technique and other parallel computing technique. This is also our next focus.

  13. Transient Aseismic Slip in the Cascadia Subduction Zone: From Monitoring to Useful Real-time Hazards Information (United States)

    Roeloffs, E. A.; Beeler, N. M.


    The Cascadia subduction zone, extending from northern California to Vancouver Island, has a 10,000 year record of earthquakes > M8.5 at intervals of several hundred years, with the last major event (~M9) in 1700. Agencies in CA, OR, WA, and BC are raising public awareness of the hazards posed by a repeat Cascadia earthquake and its ensuing tsunami. Because most of the subduction interface is now seismically quiet, an interface event M6 or larger would generate intense public concern that it could be a potential foreshock of a great earthquake. Cascadia residents are also interested in the episodic tremor and slip (ETS) events that recur months to years apart: strong evidence implies these aseismic events represent accelerated interface slip downdip of the seismogenic zone. Simple mechanics implies ETS events temporarily increase the stressing rate on the locked zone. ETS events in northern Cascadia recur at fairly regular intervals and produced roughly similar patterns of deformation. However, an unusually large ETS event or increased interface seismicity would certainly prompt public officials and local residents to expect scientists to quickly determine the implications for a major Cascadia earthquake. Earthquake scientists generally agree that such “situations of concern” warrant close monitoring, but attempts to quantify potential probability changes are in very early stages. With >30 borehole strainmeters and >100 GPS stations of the NSF-funded Plate Boundary Observatory (PBO) in Cascadia, geodesists must develop a well-organized real-time monitoring scheme for interpreting aseismic deformation, with an accompanying public communication strategy. Two previously-exercised monitoring and communication protocols could be adapted for Cascadia. During the Parkfield, California, Earthquake Experiment, geodetic signals were assigned alert levels based on their rareness in the past record, on confirmation by more than one instrument, and on consistency with

  14. Stress coupling in the seismic cycle indicated from geodetic measurements (United States)

    Wang, L.; Hainzl, S.; Zoeller, G.; Holschneider, M.


    The seismic cycle includes several phases, the interseismic, coseismic and postseismic phase. In the interseismic phase, strain gradually builds up around the overall locked fault in tens to thousands of years, while it is coseismically released in seconds. In the postseismic interval, stress relaxation lasts months to years, indicated by evident aseismic deformations which have been indicated to release comparable or even higher strain energy than the main shocks themselves. Benefiting from the development of geodetic observatory, e.g., Global Positioning System (GPS) and Interferometric Synthetic Aperture Radar (InSAR) in the last two decades, the measurements of surface deformation have been significantly improved and become valuable information for understanding the stress evolution on the large fault plane. In this study, we utilize the GPS/InSAR data to investigate the slip deficit during the interseismic phase, the coseismic slip and the early postseismic creep on the fault plane. However, it is already well-known that slip inversions based only on the surface measurements are typically non-unique and subject to large uncertainties. To reduce the ambiguity, we utilize the assumption of stress coupling between interseismic and coseismic phases, and between coseismic and postseismic phases. We use a stress constrained joint inversion in Bayesian approach (Wang et al., 2012) to invert simultaneously for (1) interseismic slip deficit and coseismic slip, and (2) coseismic slip and postseismic creep. As case studies, we analyze earthquakes occurred in well-instrumented regions such as the 2004 M6.0 Parkfield earthquake, the 2010 M8.7 earthquake and the 2011 M9.1 Tohoku-Oki earthquake. We show that the inversion with the stress-coupling constraint leads to better constrained slip distributions. Meanwhile, the results also indicate that the assumed stress coupling is reasonable and can be well reflected from the available geodetic measurements. Reference: Lifeng

  15. Generalized Free-Surface Effect and Random Vibration Theory: a new tool for computing moment magnitudes of small earthquakes using borehole data (United States)

    Malagnini, Luca; Dreger, Douglas S.


    Although optimal, computing the moment tensor solution is not always a viable option for the calculation of the size of an earthquake, especially for small events (say, below Mw 2.0). Here we show an alternative approach to the calculation of the moment-rate spectra of small earthquakes, and thus of their scalar moments, that uses a network-based calibration of crustal wave propagation. The method works best when applied to a relatively small crustal volume containing both the seismic sources and the recording sites. In this study we present the calibration of the crustal volume monitored by the High-Resolution Seismic Network (HRSN), along the San Andreas Fault (SAF) at Parkfield. After the quantification of the attenuation parameters within the crustal volume under investigation, we proceed to the spectral correction of the observed Fourier amplitude spectra for the 100 largest events in our data set. Multiple estimates of seismic moment for the all events (1811 events total) are obtained by calculating the ratio of rms-averaged spectral quantities based on the peak values of the ground velocity in the time domain, as they are observed in narrowband-filtered time-series. The mathematical operations allowing the described spectral ratios are obtained from Random Vibration Theory (RVT). Due to the optimal conditions of the HRSN, in terms of signal-to-noise ratios, our network-based calibration allows the accurate calculation of seismic moments down to Mw < 0. However, because the HRSN is equipped only with borehole instruments, we define a frequency-dependent Generalized Free-Surface Effect (GFSE), to be used instead of the usual free-surface constant F = 2. Our spectral corrections at Parkfield need a different GFSE for each side of the SAF, which can be quantified by means of the analysis of synthetic seismograms. The importance of the GFSE of borehole instruments increases for decreasing earthquake's size because for smaller earthquakes the bandwidth available

  16. Impact of a Large San Andreas Fault Earthquake on Tall Buildings in Southern California (United States)

    Krishnan, S.; Ji, C.; Komatitsch, D.; Tromp, J.


    In 1857, an earthquake of magnitude 7.9 occurred on the San Andreas fault, starting at Parkfield and rupturing in a southeasterly direction for more than 300~km. Such a unilateral rupture produces significant directivity toward the San Fernando and Los Angeles basins. The strong shaking in the basins due to this earthquake would have had a significant long-period content (2--8~s). If such motions were to happen today, they could have a serious impact on tall buildings in Southern California. In order to study the effects of large San Andreas fault earthquakes on tall buildings in Southern California, we use the finite source of the magnitude 7.9 2001 Denali fault earthquake in Alaska and map it onto the San Andreas fault with the rupture originating at Parkfield and proceeding southward over a distance of 290~km. Using the SPECFEM3D spectral element seismic wave propagation code, we simulate a Denali-like earthquake on the San Andreas fault and compute ground motions at sites located on a grid with a 2.5--5.0~km spacing in the greater Southern California region. We subsequently analyze 3D structural models of an existing tall steel building designed in 1984 as well as one designed according to the current building code (Uniform Building Code, 1997) subjected to the computed ground motion. We use a sophisticated nonlinear building analysis program, FRAME3D, that has the ability to simulate damage in buildings due to three-component ground motion. We summarize the performance of these structural models on contour maps of carefully selected structural performance indices. This study could benefit the city in laying out emergency response strategies in the event of an earthquake on the San Andreas fault, in undertaking appropriate retrofit measures for tall buildings, and in formulating zoning regulations for new construction. In addition, the study would provide risk data associated with existing and new construction to insurance companies, real estate developers, and

  17. Development of double-pair double difference location algorithm and its application to the regular earthquakes and non-volcanic tremors (United States)

    Guo, H.; Zhang, H.


    can be directly constructed from the station-pair data means that double-pair DD method can be used for improving NVT locations. We have applied the new method to the NVTs beneath the SAF near Cholame, California. Compared to the previous results, the new double-pair DD tremor locations are more concentrated and show more detailed structures.

  18. Automatic picking of direct P, S seismic phases and fault zone head waves (United States)

    Ross, Z. E.; Ben-Zion, Y.


    We develop a set of algorithms for automatic detection and picking of direct P and S waves, as well as fault zone head waves (FZHW), generated by earthquakes on faults that separate different lithologies and recorded by local seismic networks. The S-wave picks are performed using polarization analysis and related filters to remove P-wave energy from the seismograms, and utilize STA/LTA and kurtosis detectors in tandem to lock on the phase arrival. The early portions of P waveforms are processed with STA/LTA, kurtosis and skewness detectors for possible first-arriving FZHW. Identification and picking of direct P and FZHW is performed by a multistage algorithm that accounts for basic characteristics (motion polarities, time difference, sharpness and amplitudes) of the two phases. The algorithm is shown to perform well on synthetic seismograms produced by a model with a velocity contrast across the fault, and observed data generated by earthquakes along the Parkfield section of the San Andreas fault and the Hayward fault. The developed techniques can be used for systematic processing of large seismic waveform data sets recorded near major faults.

  19. Tidal Sensitivity of Declustered Low Frequency Earthquake Families and Inferred Creep Episodes on the San Andreas Fault (United States)

    Babb, A.; Thomas, A.; Bletery, Q.


    Low frequency earthquakes (LFEs) are detected at depths of 16-30 km on a 150 km section of the San Andreas Fault centered at Parkfield, CA. The LFEs are divided into 88 families based on waveform similarity. Each family is thought to represent a brittle asperity on the fault surface that repeatedly slips during aseismic slip of the surrounding fault. LFE occurrence is irregular which allows families to be divided into continuous and episodic. In continuous families a burst of a few LFE events recurs every few days while episodic families experience essentially quiescent periods often lasting months followed by bursts of hundreds of events over a few days. The occurrence of LFEs has also been shown to be sensitive to extremely small ( 1kPa) tidal stress perturbations. However, the clustered nature of LFE occurrence could potentially bias estimates of tidal sensitivity. Here we re-evaluate the tidal sensitivity of LFE families on the deep San Andreas using a declustered catalog. In this catalog LFE bursts are isolated based on the recurrence intervals between individual LFE events for each family. Preliminary analysis suggests that declustered LFE families are still highly sensitive to tidal stress perturbations, primarily right-lateral shear stress (RLSS) and to a lesser extent fault normal stress (FNS). We also find inferred creep episodes initiate preferentially during times of positive RLSS.

  20. Ultra-high resolution four dimensional geodetic imaging of engineered structures for stability assessment (United States)

    Bawden, Gerald W.; Bond, Sandra; Podoski, J. H.; Kreylos, O.; Kellogg, L. H.


    We used ground-based Tripod LiDAR (T-LiDAR) to assess the stability of two engineered structures: a bridge spanning the San Andreas fault following the M6.0 Parkfield earthquake in Central California and a newly built coastal breakwater located at the Kaumālapa`u Harbor Lana'i, Hawaii. In the 10 weeks following the earthquake, we found that the surface under the bridge shifted 7.1 cm with an additional 2.6 cm of motion in the subsequent 13 weeks, which deflected the bridge's northern I-beam support 4.3 cm and 2.1 respectively; the bridge integrity remained intact. T-LiDAR imagery was collected after the completion of armored breakwater with 817 35-ton interlocking concrete armor units, Core-Locs®, in the summers of 2007, 2008 and 2010. We found a wide range of motion of individual Core-Locs, from a few centimeters to >110 cm along the ocean side of the breakwater, with lesser movement along the harbor side.

  1. Wavelet-based blind identification of the UCLA Factor building using ambient and earthquake responses

    International Nuclear Information System (INIS)

    Hazra, B; Narasimhan, S


    Blind source separation using second-order blind identification (SOBI) has been successfully applied to the problem of output-only identification, popularly known as ambient system identification. In this paper, the basic principles of SOBI for the static mixtures case is extended using the stationary wavelet transform (SWT) in order to improve the separability of sources, thereby improving the quality of identification. Whereas SOBI operates on the covariance matrices constructed directly from measurements, the method presented in this paper, known as the wavelet-based modified cross-correlation method, operates on multiple covariance matrices constructed from the correlation of the responses. The SWT is selected because of its time-invariance property, which means that the transform of a time-shifted signal can be obtained as a shifted version of the transform of the original signal. This important property is exploited in the construction of several time-lagged covariance matrices. The issue of non-stationary sources is addressed through the formation of several time-shifted, windowed covariance matrices. Modal identification results are presented for the UCLA Factor building using ambient vibration data and for recorded responses from the Parkfield earthquake, and compared with published results for this building. Additionally, the effect of sensor density on the identification results is also investigated

  2. Preliminary isostatic residual gravity anomaly map of Paso Robles 30 x 60 minute quadrangle, California (United States)

    McPhee, D.K.; Langenheim, V.E.; Watt, J.T.


    This isostatic residual gravity map is part of an effort to map the three-dimensional distribution of rocks in the central California Coast Ranges and will serve as a basis for modeling the shape of basins and for determining the location and geometry of faults within the Paso Robles quadrangle. Local spatial variations in the Earth\\'s gravity field, after accounting for variations caused by elevation, terrain, and deep crustal structure reflect the distribution of densities in the mid- to upper crust. Densities often can be related to rock type, and abrupt spatial changes in density commonly mark lithological or structural boundaries. High-density rocks exposed within the central Coast Ranges include Mesozoic granitic rocks (exposed northwest of Paso Robles), Jurassic to Cretaceous marine strata of the Great Valley Sequence (exposed primarily northeast of the San Andreas fault), and Mesozoic sedimentary and volcanic rocks of the Franciscan Complex [exposed in the Santa Lucia Range and northeast of the San Andreas fault (SAF) near Parkfield, California]. Alluvial sediments and Tertiary sedimentary rocks are characterized by low densities; however, with increasing depth of burial and age, the densities of these rocks may become indistinguishable from those of older basement rocks.

  3. U.S.-Japan Quake Prediction Research (United States)

    Kisslinger, Carl; Mikumo, Takeshi; Kanamori, Hiroo

    For the seventh time since 1964, a seminar on earthquake prediction has been convened under the U.S.-Japan Cooperation in Science Program. The purpose of the seminar was to provide an opportunity for researchers from the two countries to share recent progress and future plans in the continuing effort to develop the scientific basis for predicting earthquakes and practical means for implementing prediction technology as it emerges. Thirty-six contributors, 15 from Japan and 21 from the U.S., met in Morro Bay, Calif.September 12-14. The following day they traveled to nearby sections of the San Andreas fault, including the site of the Parkfield prediction experiment. The conveners of the seminar were Hiroo Kanamori, Seismological Laboratory, California Institute of Technology (Caltech), for the U.S., and Takeshi Mikumo, Disaster Prevention Research Institute, Kyoto University, for Japan . Funding for the participants came from the U.S. National Science Foundation and the Japan Society forthe Promotion of Science, supplemented by other agencies in both countries.

  4. Median Filtering Methods for Non-volcanic Tremor Detection (United States)

    Damiao, L. G.; Nadeau, R. M.; Dreger, D. S.; Luna, B.; Zhang, H.


    Various properties of median filtering over time and space are used to address challenges posed by the Non-volcanic tremor detection problem. As part of a "Big-Data" effort to characterize the spatial and temporal distribution of ambient tremor throughout the Northern San Andreas Fault system, continuous seismic data from multiple seismic networks with contrasting operational characteristics and distributed over a variety of regions are being used. Automated median filtering methods that are flexible enough to work consistently with these data are required. Tremor is characterized by a low-amplitude, long-duration signal-train whose shape is coherent at multiple stations distributed over a large area. There are no consistent phase arrivals or mechanisms in a given tremor's signal and even the durations and shapes among different tremors vary considerably. A myriad of masquerading noise, anthropogenic and natural-event signals must also be discriminated in order to obtain accurate tremor detections. We present here results of the median methods applied to data from four regions of the San Andreas Fault system in northern California (Geysers Geothermal Field, Napa, Bitterwater and Parkfield) to illustrate the ability of the methods to detect tremor under diverse conditions.

  5. Prediction of spectral acceleration response ordinates based on PGA attenuation (United States)

    Graizer, V.; Kalkan, E.


    Developed herein is a new peak ground acceleration (PGA)-based predictive model for 5% damped pseudospectral acceleration (SA) ordinates of free-field horizontal component of ground motion from shallow-crustal earthquakes. The predictive model of ground motion spectral shape (i.e., normalized spectrum) is generated as a continuous function of few parameters. The proposed model eliminates the classical exhausted matrix of estimator coefficients, and provides significant ease in its implementation. It is structured on the Next Generation Attenuation (NGA) database with a number of additions from recent Californian events including 2003 San Simeon and 2004 Parkfield earthquakes. A unique feature of the model is its new functional form explicitly integrating PGA as a scaling factor. The spectral shape model is parameterized within an approximation function using moment magnitude, closest distance to the fault (fault distance) and VS30 (average shear-wave velocity in the upper 30 m) as independent variables. Mean values of its estimator coefficients were computed by fitting an approximation function to spectral shape of each record using robust nonlinear optimization. Proposed spectral shape model is independent of the PGA attenuation, allowing utilization of various PGA attenuation relations to estimate the response spectrum of earthquake recordings.

  6. Post-seismic relaxation from geodetic and seismic data

    Directory of Open Access Journals (Sweden)

    Mikhail V. Rodkin


    Full Text Available We have examined the aftershock sequence and the post-seismic deformation process of the Parkfield earthquake (2004, M = 6, California, USA source area using GPS data. This event was chosen because of the possibility of joint analysis of data from the rather dense local GPS network (from SOPAC Internet archive and of the availability of the rather detailed aftershock sequence data ( The relaxation process of post-seismic deformation prolongs about the same 400 days as the seismic aftershock process does. Thus, the aftershock process and the relaxation process in deformation could be the different sides of the same process. It should be noted that the ratio of the released seismic energy and of the GPS obtained deformation is quite different for the main shock and for the aftershock stage. The ratio of the released seismic energy to the deformation value decreases essentially for the post-shock process. The similar change in the seismic energy/deformation value ratio is valid in a few other strong earthquakes. Thus, this decrease seems typical of aftershock sequences testifying for decrease of ratio of elastic to inelastic deformation in the process of post-shock relaxation when the source area appears to be mostly fractured after the main shock occurs, but the healing process had no yet sufficient time to develop.

  7. Tremor-tide correlations and near-lithostatic pore pressure on the deep San Andreas fault. (United States)

    Thomas, Amanda M; Nadeau, Robert M; Bürgmann, Roland


    Since its initial discovery nearly a decade ago, non-volcanic tremor has provided information about a region of the Earth that was previously thought incapable of generating seismic radiation. A thorough explanation of the geologic process responsible for tremor generation has, however, yet to be determined. Owing to their location at the plate interface, temporal correlation with geodetically measured slow-slip events and dominant shear wave energy, tremor observations in southwest Japan have been interpreted as a superposition of many low-frequency earthquakes that represent slip on a fault surface. Fluids may also be fundamental to the failure process in subduction zone environments, as teleseismic and tidal modulation of tremor in Cascadia and Japan and high Poisson ratios in both source regions are indicative of pressurized pore fluids. Here we identify a robust correlation between extremely small, tidally induced shear stress parallel to the San Andreas fault and non-volcanic tremor activity near Parkfield, California. We suggest that this tremor represents shear failure on a critically stressed fault in the presence of near-lithostatic pore pressure. There are a number of similarities between tremor in subduction zone environments, such as Cascadia and Japan, and tremor on the deep San Andreas transform, suggesting that the results presented here may also be applicable in other tectonic settings.


    Directory of Open Access Journals (Sweden)

    Yury V. Gabsatarov


    Full Text Available Analysis of data from permanent GPS observation stations located in tectonically active regions provides for direct observation of deformation processes of the earth's surface which result from elastic interaction of the lithospheric plates and also occur when accumulated stresses are released by seismic events and postseismic processes.This article describes the methodology of applying the regression analysis of time series of data from GPS-stations for identification of individual components of the stations’ displacements caused by the influence of various deformation processes. Modelling of the stations’ displacements caused only by deformations of the marginal zone, wherein the lithospheric plates interact, allows us to study variations of the steady-state deformation in the marginal zone.he proposed methodology is applied to studies of variations of fields of cumulative surface displacements, surface displacement velocity and maximum shear strain velocity which are determined from the GPS data recorded prior to the Parkfield earthquake of 28 September 2004 (Mw=6.0.Combined analysis of the variations of the above-mentioned fields shows that measurable anomalies of the elastic deformation of the transform fault’s edge took place prior to the seismic event of 28 September 2004, and such anomalies were coincident in space and time with the focal area of the future seismic event.

  9. Seismic response analysis with liquid-structure interaction

    International Nuclear Information System (INIS)

    Thomas, R.G.; Harrop, L.P.


    A linear transient finite element stress analysis of a water filled tank has been carried out using the proprietary computer code ANSYS. The containment structure was represented as rigidly fixed to ground. The flexibility of the tank wall was modelled together with the hydrostatic and hydrodynamic effects of the water contents and attached concentrated masses. The foundations were considered to be laid in solid rock, and no soil-structure interaction effects were included. The seismic input was a ground response spectrum conservatively representing both the Temblor and Parkfield modified time history records. It was found that the response of the structure was greatest at the front end (furthest from the point at which the tank is connected to a rigid internal structure), and that this was dominated by the fundamental mode. Higher modes are important at the back end. Buckling at the front end of the tank has been identified as a potential failure mechanism, and attention has also been called to the tensile capacity of the wall to base junction in this region. The requirement for a proper criterion against which to assess the margin against plastic collapse in a safe shutdown analysis has been noted. In certain regions the structure does not shake-down under the repeated reversed cyclic loading, and the need for an assessment of the implications of this for fatigue resistance has been indicated. (author)

  10. Deep rock damage in the San Andreas Fault revealed by P- and S-type fault-zone-guided waves (United States)

    Ellsworth, William L.; Malin, Peter E.


    Damage to fault-zone rocks during fault slip results in the formation of a channel of low seismic-wave velocities. Within such channels guided seismic waves, denoted by Fg, can propagate. Here we show with core samples, well logs and Fg-waves that such a channel is crossed by the SAFOD (San Andreas Fault Observatory at Depth) borehole at a depth of 2.7 km near Parkfield, California, USA. This laterally extensive channel extends downwards to at least half way through the seismogenic crust, more than about 7 km. The channel supports not only the previously recognized Love-type- (FL) and Rayleigh-type- (FR) guided waves, but also a new fault-guided wave, which we name FF. As recorded 2.7 km underground, FF is normally dispersed, ends in an Airy phase, and arrives between the P- and S-waves. Modelling shows that FF travels as a leaky mode within the core of the fault zone. Combined with the drill core samples, well logs and the two other types of guided waves, FF at SAFOD reveals a zone of profound, deep, rock damage. Originating from damage accumulated over the recent history of fault movement, we suggest it is maintained either by fracturing near the slip surface of earthquakes, such as the 1857 Fort Tejon M 7.9, or is an unexplained part of the fault-creep process known to be active at this site.

  11. La forma de gobierno en la Constitución de Cádiz : reflexiones sobre la configuración de la jefatura del estado monárquica

    Directory of Open Access Journals (Sweden)

    Remedio Sánchez Ferriz


    ítica y el ejercicio efectivo de la potestad ejecutiva (aunque conserve su titularidad formal, llegándose a tal conclusión sobre la base la ausencia de tres presupuestos esenciales: un Gobierno colegiado separado del monarca con funciones ejecutivas y de dirección política, una relación de confianza parlamentaria entre el Ejecutivo y el Parlamento y, finalmente, un núcleo de funciones representativas propias del Jefe del Estado e independientes de su titularidad nominal del poder ejecutivo. Starting from the historical circumstances of the Spanish Parliament («Cortes» of Cádiz and the precedents of the Spanish monarchy, the article analyses the position of the monarchic Head of State in the Constitution of 1812 characterized as a «moderate monarchy» based on the structural principles of national sovereignty and separation of powers that determine the configuration of the King as a constitutionally limited institution in the new form of government. The assertion of national sovereignty pervades the whole of the Constitution without detriment to coexist with the old rhetorical monarchical legitimacy. Moreover, the separation of powers involves the distinction between ownership of sovereignty and its exercise by the constitutionally limited institutions, considering the King the «receiver » of the executive branch against a Parliament that, as it represents the Nation owner of the sovereignty, takes a decisive role in the exercise of the King’s functions. The scope of the broad powers granted to the King is delimited with the numerous restrictions imposed on a system of strict separation of powers. The «moderate monarchy» proclaimed in Cádiz is not a simple variant of the limited European monarchies based on the monarchic principle but a system of government founded on national sovereignty and the distribution of state functions that limits the authority —constitutionally limited rather than sovereign— of the monarchic Executive. Although the

  12. Evaluation of some software measuring displacements using GPS in real-time (United States)

    Langbein, John


    For the past decade, the USGS has been monitoring deformation at various locations in the western United States using continuous GPS. The main focus of these measurements are estimates of displacement averaged over one day. Essentially, these consist of recording at 30 seconds intervals the carrier-frequency phase-data (equivalent to travel-time) between a GPS receiver and the GPS satellite network. In turn, these observations, which are converted to pseudo—ranges, are processed using one of the “research grade” programs (GIPSY, Zumberge et al., or GAMIT, to estimate the position of the GPS receiver averaged over 24 hours. However, it is possible and desirable to estimate the position of the receiver (actually the antenna) more frequently and to do this within a few seconds of the time actual measurement (known as real-time). A recent example, the 2004 Magnitude 6, Parkfield, California earthquake, demonstrated that having GPS estimates of position more frequently than simply a daily average is required if one requires discrimination between co-seismic and post-seismic deformation (Langbein et al., 2006). The high-rate estimates of position obtained at Parkfield show that post-seismic deformation started less than one-hour after the mainshock and that this deformation was roughly the same magnitude as the co-seismic deformation. The high-rate solutions for Parkfield were done by others including Yehuda Bock at UCSD and Kristine Larson at U. of Colorado, but not the USGS. The Parkfield experience points out the need for an in-house capability by the USGS to be able to accurately measure co-seismic displacements and other rapid, deformation signals using GPS. This applies to both the Earthquake and Volcano Hazard programs. Although at many locations where we monitor deformation, we have strainmeters and tiltmeters in addition to GPS which, in principle, are far more sensitive to rapid deformation over periods of less than a day

  13. Monaco. (United States)


    The Principality of Monaco's population characteristics, history, government, political situation, economy, and foreign affairs were briefly described. Monaco, a constitutional monarchy, covers an area of only .6 of a square mile. It is situated on the Mediterranean coast and is surrounded by France on 3 sides. The country was founded as a colony of Genoa, Italy, in 1215, and since 1419, it has been ruled by the House of Grimaldi, except for a brief period of time when the country was under the rule of France. Monaco and France have close ties which have been confirmed by a series of treaties beginning in 1918. In accordance with these treaties, France recognizes the sovereignity of Monaco, and Monaco agrees to exercise its rights only in so far as they accord with French political, military, naval, and economic interests. The monarch was an absolute ruler until 1911 when the constitutional monarchy was established. In 1962, a new constitution was adopted. Under the new constitution, the monarch, or prince of Monaco, shares power with an 18-member elected ledgislative body, the National Council. The Council votes on the budget and endorses laws put forth by the monarch. The executive branch consists of 1) the prince as chief of state; 2) a minister of state, who is a French citizen, appointed by the prince from a list of 3 candidates put forth by the French Government; and 3) a 3-member cabinet. Currently the chief of state is Prince Rainier III, and the minister of state is Jean Herly. The country has a population of 27,000 and an annual population growth rate of 0.9%. The population is 47% French, 16% Italian, and 16% Monegasque. 95% of the inhabitants are Roman Catholic, and French is the official language. School is compulsory between the ages of 6-16 years, and the literacy rate is 100%. The country is an elite tourist center. Its popularity is due, in part, to the casino, which was established in Monaco in 1856. The casino which once was the country's major

  14. Música y fiesta barroca: celebraciones en Tortosa en honor de Felipe V (1701

    Directory of Open Access Journals (Sweden)

    Rosa Montagut, Marian


    Full Text Available This essay describes a specific case of the presence of music during the popular festivities that were held in Tortosa at the beginning of 1701 celebrating and exalting the coronation of the then new and recently crowned Monarch, the Bourbon Prince Felipe de Anjou. The source for analyzing the varied and abundant musical participation in the said festivities is a document printed in Barcelona in 1701 entitled "Festive merrymaking, courteous and loyal demonstrations with which the most faithful and exemplary city of Tortosa celebrated the joyous arrival in his Royal Court of our great Monarch, Felipe de Anjou (Bourbon, King of Spain (may God save him". The importance of this anonymous document lies, on the one hand, in the detailed description it offers us of the essential role played by music - along with other artistic manifestations - in the majority of the activities, and, on the other hand, as a paradigmatic testimony of the relationship between music and "fiesta" during the epoch of the Hispanic Baroque.

    El presente trabajo se basa en un caso concreto de presencia musical en el contexto de unas celebraciones ciudadanas que tuvieron lugar en Tortosa, como festejo y propaganda política a favor de un nuevo y recién coronado monarca, a principios de 1701: el borbón Felipe de Anjou. El vehículo para analizar la variada y abundante participación musical de dichas festividades es un documento impreso en Barcelona en 1701, titulado "Alborozos festivos, leales obsequiosas demostraciones con que la fidelissima y exemplar ciudad de Tortosa celebró el feliz arribo a su real corte de nuestro gran monarca, y señor D. Felipe de Borbón rey de las Españas (que Dios guarde". La importancia de este impreso anónimo radica, por un lado, en la detallada descripción que nos ofrece del papel primordial que desempeñó la música —junto con otras manifestaciones artísticas— en la mayor parte de los actos realizados, y por otro lado, en

  15. «Una columna fortísima del altar y del trono»: Pedro Gutiérrez de Cos, obispo de Huamanga y de Puerto Rico (1750-1833

    Directory of Open Access Journals (Sweden)

    Hernández García, Elizabeth


    Full Text Available Pedro Gutiérrez de Cos has not been sufficiently studies in Peruvian historiography and is scarcely mentioned once he left Perú, due to this it has not been possible to conduct an in-depth study about his option for independence, position which prevailed among his thinking and acting until his death in Puerto Rico. This articles aims to lim three aspects in Gutiérrez de Cos: his academic and professional performance, from his beginnings in Peruvian higher education centers, until the obtention of tthe most important titles in the Hispanic-American ecclesiastic sphere; the political conservative ideas, pillars of the monarchic government and his performance in their respective ecclesiastic destinations; and, among all of these, the independentist political unrest in Perú. The monarchical spirit of Gutiérrez de Cos enabled him to acquire another bishopric beyond this ocean.Pedro Gutiérrez de Cos no ha sido suficientemente trabajado en la historiografía peruana y es poco mencionado una vez que ha salido del Perú, razón por la cual no se ha conseguido profundizar en su opción por la independencia, postura que rigió su pensamiento y actuación hasta su muerte en Puerto Rico. Este artículo se propone entretejer tres aspectos en Gutiérrez de Cos: su trayectoria académica y profesional, desde sus inicios en los centros de enseñanza superior peruanos, hasta la consecución de los nombramientos más importantes en la esfera eclesiástica hispanoamericana; las ideas políticas conservadoras, pilares del gobierno monárquico y de su desempeño en sus respectivos destinos eclesiásticos; y, en medio de todo ello, la convulsión política independentista en el Perú, que consiguió reafirmar su monarquismo y le posibilitó otra mitra en ultramar.

  16. Monarquías en América

    Directory of Open Access Journals (Sweden)

    Manuel Teruel Gregorio de Tejada


    Full Text Available El tema del monarquismo americano no dispone de una obra seria de conjunto. Este artículo trata de ofrecer un ensayo de síntesis global, estructurado en cuatro apartados. El primero se ocupa de los proyectos generales emanados desde la propia Monarquía hispánica: la Memoria del conde de Aranda, con la que se relacionan el informe de Campomanes, del conde de Floridablanca y la representación de José de Ábalos, durante el reinado de Carlos III, así como el proyecto de Manuel de Godoy durante el de Carlos IV. El segundo apartado describe en primer lugar las iniciativas, proyectos o intentos de establecer regímenes monárquicos dentro y después del proceso emancipador, en los que estuvieron implicados Argentina, Colombia, Ecuador, Perú y Bolivia, y a continuación las experiencias del Brasil, México y Haití, donde llegaron a crearse tales regímenes, aunque tuvieron una vida corta y agitada, con la excepción del brasileño, que subsistió durante medio siglo y con relativo sosiego. El tercer apartado esboza las iniciativas de carácter insólito o aventurero, entre los que destacan los casos de Araucania y Redonda.No reliable all-embracing work has dealt with American monarchism. This article tries to offer a syntesis in four parts. The first deals with the general projets originated in the Spanish Monarchy itself: Count of Aranda’s Memoria, which is related to Counts of Campomanes’ and Floridablanca’s Informe and José de Abalos’ Representación, all of them under the reign of Charles III, and the project of Manuel de Godoy under Charles IV’s. The second part depicts the initiatives, projects and attempts to establish monarchic systems during and after the independence process, in which Argentine, Colombia, Ecuador, Peru and Bolivia were involved, and the experiences developed in Brazil, Mexico and Haiti, where such systems were held, although they had a short and uneasy life, excepted for Brazil, whose system survived

  17. Contrasting patterns of evolutionary constraint and novelty revealed by comparative sperm proteomic analysis in Lepidoptera. (United States)

    Whittington, Emma; Forsythe, Desiree; Borziak, Kirill; Karr, Timothy L; Walters, James R; Dorus, Steve


    Rapid evolution is a hallmark of reproductive genetic systems and arises through the combined processes of sequence divergence, gene gain and loss, and changes in gene and protein expression. While studies aiming to disentangle the molecular ramifications of these processes are progressing, we still know little about the genetic basis of evolutionary transitions in reproductive systems. Here we conduct the first comparative analysis of sperm proteomes in Lepidoptera, a group that exhibits dichotomous spermatogenesis, in which males produce a functional fertilization-competent sperm (eupyrene) and an incompetent sperm morph lacking nuclear DNA (apyrene). Through the integrated application of evolutionary proteomics and genomics, we characterize the genomic patterns potentially associated with the origination and evolution of this unique spermatogenic process and assess the importance of genetic novelty in Lepidopteran sperm biology. Comparison of the newly characterized Monarch butterfly (Danaus plexippus) sperm proteome to those of the Carolina sphinx moth (Manduca sexta) and the fruit fly (Drosophila melanogaster) demonstrated conservation at the level of protein abundance and post-translational modification within Lepidoptera. In contrast, comparative genomic analyses across insects reveals significant divergence at two levels that differentiate the genetic architecture of sperm in Lepidoptera from other insects. First, a significant reduction in orthology among Monarch sperm genes relative to the remainder of the genome in non-Lepidopteran insect species was observed. Second, a substantial number of sperm proteins were found to be specific to Lepidoptera, in that they lack detectable homology to the genomes of more distantly related insects. Lastly, the functional importance of Lepidoptera specific sperm proteins is broadly supported by their increased abundance relative to proteins conserved across insects. Our results identify a burst of genetic novelty

  18. The selection of royal figures in the image of power during the Palaiologan epoch: Byzantium - Serbia - Bulgaria

    Directory of Open Access Journals (Sweden)

    Vojvodić Dragan


    Full Text Available The preserved presentations of the Byzantine basileis of the XIII, XIV and XV centuries show that the creators of the late Byzantine monarchical portraits adhered to certain traditional rules when selecting the personages from the ruling house, which they were to portray. Defining which figures were to be depicted in the portrayal of power depended to a large extent on the changing circumstances and events in the imperial house. However, at the same time this was also based on a significantly more profound conception that rested on principles that had evolved in the course of a long history. The understanding of who could personify power was refracted through the prism of ideology and reflected in carefully shaped iconographic matrices. The omission of the images of certain members of the ruler's house, just as much as their inclusion, carried a certain meaning, as did the hierarchical arrangement of those who were portrayed. Generally speaking, this depended on the degree of their kinship with the sovereign, their sex, titles or dignities, and the connection of the members of the dynasty with the emperor's particular marriage. Therefore, one can rather clearly distinguish certain constants, if not rules, according to which some figures were omitted and others included, and, the specific changes that occurred from the end of the Middle Byzantine period till the fall of the Empire. The development of a unique kind of feudalism played a particular role in the specific characteristics in determining who was to appear in the monarchical portraits of the Palaiologan epoch in Byzantium and the states in its neighbourhood. As the preserved portrait ensembles and known written testimonies indicate, we find the images of the rulers' daughters did not feature in presentations of the 'emperors of the Romans' from the Late Byzantine period. In the Palaiologan epoch, they did not participate in the governing of the state nor were they taken into

  19. Literatura y sociedad. Algunos ejemplos de la presencia de Jaime I en la Región de Murcia, a través de diversos contextos escritos. Siglos XIX y XX

    Directory of Open Access Journals (Sweden)

    Ramón Doménech Villa


    Full Text Available Cada lugar se caracteriza por tener su héroe particular. Sería muy difícil encontrar dos héroes compartidos en diferentes espacios físicos, mucho más si estos se encuentran limitando entre sí. En la actualidad podría ser comparable a las rivalidades existentes en ciertas competiciones deportivas, incluso podríamos plantearnos la existencia de ciertos paralelismos entre estos y aquellas situaciones bélicas producidas en siglos anteriores. La Región de Murcia, Andalucía y el País Valencià, compitieron en un particular duelo de protagonismo monárquico, con claros parentescos familiares, durante el siglo XIII. Ante el predominio de uno u otro monarca, y sin contar con su protagonismo destacado podríamos hacernos multitud de preguntas: ¿Desaparecieron todas las referencias al rey Jaime I, dentro del marco de la sociedad murciana, una vez pasados los siglos con otro rey como protagonista? ¿Sería posible encontrar referencias a él, en contextos relacionados sociales de la Región de Murcia? ¿Jugó la literatura de la época algún papel en este sentido? ¿El ciudadano de la Región podía crear un contexto no formal de aprendizaje histórico? Each site is characterized by its particular hero. It would be very difficult to find two different heroes in shared physical spaces, even if they are border regions. In this days and age, this fact could be comparable to existing rivalries in certain sport competitions; besides, we may even look on the existence of certain parallelisms between these latter ones and those warfare situations produced in earlier centuries. The Region of Murcia, Andalusia and the Valencian Country competed each other in a particular feud for a monarchic prominence with clear family relationships during the XIII century. Given the predominance of one or another monarch, and regardless of their leading role we might ask ourselves many questions: Did all references to King James I disappeared, in the context of

  20. On the different meanings of the term law (zakon in Saint Sabba's Life of Saint Simon

    Directory of Open Access Journals (Sweden)

    Šarkić Srđan N.


    Full Text Available In mediaeval Serbian law the central legal term zakon (law indicated a generally obligatory rule (regula iuris which was usually not a result of the activity of a monarch as ultimate holder of state power. Even where a law was made by state authority such a legal rule had primarily the appearence of a customary legal provision, regulating the conditions within one particular manor (vlastelinstvo rather than within the whole national territory. Otherwise such laws prescibed the legal position of different categories of inhabitants and identified particular rules of status. Sometimes a law would be introduced to regulate one paricular problem. The concept of law in this period also includes a legal rule derived from custom or from a private contract. Each of these uses can be illustrated from many hundreds of cases from several sources. The use of the term zakon (law was present in the literary sources as well, such is The Life of Saint Simon (biography of Stefan Nemanja, founder of Serbian mediaeval dynasty Nemanjić, written by his youngest son Rastko, bether known under his monastic name Sabba (Sava. In Sabba's hagiography of his father we found the term zakon six times. Discribing the State Council (Državni sabor that had to decide who will be Nemanja's successor on the throne, Sabba writes that his father pronounced, among other, the following words: 'My sons, do not forget the orthodox law that I established.' The term orthodox law means here orthodox faith, that was established in Serbia after persecutions of Bogomilian heresy. For the second time, term zakon was used in the meaning of monarch's order. Nemanja says to his sons not to forget his laws. Further, giving the instructions to his sons, Nemanja use the citation from the Bible (The Book of Proverbs or Proverbs of Solomon 3, 1-18, where the term zakon corresponds to the latin ius, not lex. Hereinafter the word zakon means Torah or Pentateuch, the first five books of Bible (citation

  1. Ireland and Spain 1931-1933. Divergent Republics

    Directory of Open Access Journals (Sweden)

    Alvaro Jaspe


    Full Text Available This paper will examine the development of the fledgling diplomatic ties between the new Irish state and the recently established Spanish legation based in Dublin.  It will analyse the formal establishment of political, cultural and social links developed in contemporary times between the two old historical allies that had previously been limited to polite lip-service; conditioned, in part, as they were by  monarchical Madrid’s caution in regard to London and previous Spanish reluctance to engage with a rebellious state.  1931 signified a volte face in the relationship developing between Dublin and Madrid since the establishment of a Spanish consulate in Dublin in March 1924, facilitated by their commitment to the League of Nations, to which both were strongly committed in the 1920s.  This paper will illustrate how the declaration of the II Republic in Spain was a crossroads in the relationship between these two nations.  The roles of ‘rebel’ and ‘traditionalist’ state had been instantly switched.  The Church and much of the new political elite in Ireland viewed republican reforms in Spain with ever growing and public distaste creating conflict among Irish republicanism, post-independence.  The main Spanish republican representative in Ireland in this 1931-33 period, Emilio Sanz y Tovar, became very sensitive to these schisms as he tried to cement political and socio-economic ties with his Irish hosts.

  2. Miguel Ciera: um demarcador de limites no interior sul-americano (1750-1760

    Directory of Open Access Journals (Sweden)

    Maria de F\\u00E1tima Costa


    Full Text Available In this paper, we examine the works produced by Miguel Ciera against the backdrop of the Third Delimitation Expedition, which was sent by Portugal to explore the interior of South America between 1752 and 1756. As the astronomer and cosmographer of the demarcation team, this Paduan engineer sailed up Paraguay River all the way to the mouth of Jauru River and there, with his companions, he set the border line, thus substantiating at site the Treaty of Boundaries. It was during this expedition that Ciera gathered information to design his Mappa geographicum quo flumen Argentum, Paranà et Paraguay [¿], a beautiful atlas that he later bestowed upon King Joseph I in 1758. In addition to accurate geographic charts, the atlas included depictions of fauna, popular types and landscapes made in pencil and watercolor, forming the first set of images available of that region, which was named Pantanal as from the 18th century. Here, we analyze this specific piece and seek to demonstrate that it was not merely a beautiful atlas; in fact, it was a refined visual document with which Miguel Ciera offered the Portuguese monarch valuable information about the hinterland here the border line was laid.

  3. Staat als Begriff und Vorstellung

    Directory of Open Access Journals (Sweden)

    Günter Frankenberg


    Full Text Available The article reconstructs the visual representations of the state beginning with the frontispiece of Thomas Hobbes' Leviathan and its variations in »The Elements of Law«. While Hobbes' visual strategy is informed by a distrust of language, Michel Foucault departs from both this distrust as well as the fixation on sovereignty. His deconstruction of the representation paves the way for a democratic aesthetics. The need for symbols of political domination in the post-monarchic era is illustrated by three examples: the symbolizations of national unity, the parliamentary seating arrangements and the domed Berlin Reichstag as the architectural embodiment of the republican promise of transparency. After this visit to the museum of modernity’s pictures the text reintroduces the concept of the modern state. Hobbes' composite structure of the Leviathan reappears under the guise of a farewell to pre-constitutional composite concepts, such as a state under the rule of law, federal state or constitutional state.

  4. Política, economía y fiscalización de un espacio urbano : el establecimiento de la aljama de los judíos en la Mallorca cristiana

    Directory of Open Access Journals (Sweden)

    Jorge Maíz Chacón


    Full Text Available En el texto analizamos la llegada de los judíos a Mallorca después de 1229. Se abordan diversas problemáticas como su instalación inicial cerca de la Almudayna, actividades que desarrollan y las distintas políticas de los monarcas (Cartas de Franqueza, Orden de los Templarios hasta la creación de la aljama balear Analizamos nuevas fuentes sobre las operaciones de compra y venta entre judíos y cristianos momentos antes de la creación del nuevo barrio judío.In the text we analyzed the arrival from the Jews to Majorca after 1229. They are approached diverse problematic like its initial installation near the Almudayna, activities that they develop and the different policies from the monarchs (Cartas de Franqueza, Orden de los Templarios until the creation of the Balearic aljama. We analyzed new sources on the operations of purchase and sale between Jews and Christians moments before the creation of the new Jew district.

  5. On the Cutting Edge of Research to Conserve At-Risk Species: Maximizing Impact through Partnerships. (United States)

    Marquardt, Shauna R; Annis, Mandy; Drum, Ryan G; Hummel, Stephanie Longstaff; Mosby, David E; Smith, Tamara


    Today's conservation challenges are complex. Solving these challenges often requires scientific collaborations that extend beyond the scope, expertise, and capacity of any single agency, organization, or institution. Conservation efforts can benefit from interdisciplinary collaboration, scientific and technological innovations, and the leveraging of capacity and resources among partners. Here we explore a series of case studies demonstrating how collaborative scientific partnerships are furthering the mission of the U.S. Fish and Wildlife Service, including: 1) contaminants of emerging concern in the Great Lakes Basin, 2) Poweshiek skipperling conservation, 3) using technology to improve population survey methods for bats and monarch butterfly, and 4) Big River restoration in the Southeast Missouri lead mining district. These case studies illustrate how strategic and effective scientific collaboration is a multi-stage process that requires investment of time and resources by all participants. Early coordination and communication is crucial to aligning planned work with scientific and decision-making needs. Collaborations between USFWS and external scientists can be mutually beneficial by supporting the agency mission while also providing an avenue for innovative research to be directly applied in conservation decisions and management actions.

  6. Complex Outcomes from Insect and Weed Control with Transgenic Plants: Ecological Surprises?

    Directory of Open Access Journals (Sweden)

    Thomas Bøhn


    Full Text Available Agriculture is fundamental for human survival through food production and is performed in ecosystems that, while simplified, still operate along ecological principles and retain complexity. Agricultural plants are thus part of ecological systems, and interact in complex ways with the surrounding terrestrial, soil, and aquatic habitats. We discuss three case studies that demonstrate how agricultural solutions to pest and weed control, if they overlook important ecological and evolutionary factors, cause “surprises”: (i the fast emergence of resistance against the crop-inserted Bt-toxin in South Africa, (ii the ecological changes generated by Bt-cotton landscapes in China, and (iii the decline of the monarch butterfly, Danaus plexippus, in North America. The recognition that we work with complex systems is in itself important, as it should limit the belief in reductionist solutions. Agricultural practices lacking eco-evolutionary understanding result in “surprises” like resistance evolution both in weeds and pest insects, risking the reappearance of the “pesticide treadmill”—with increased use of toxic pesticides as the follow-up. We recommend prioritization of research that counteracts the tendencies of reductionist approaches. These may be beneficial on a short term, but with trade-off costs on a medium- to long-term. Such costs include loss of biodiversity, ecosystem services, long-term soil productivity, pollution, and reduced food quality.

  7. Integration of polarization and chromatic cues in the insect sky compass. (United States)

    el Jundi, Basil; Pfeiffer, Keram; Heinze, Stanley; Homberg, Uwe


    Animals relying on a celestial compass for spatial orientation may use the position of the sun, the chromatic or intensity gradient of the sky, the polarization pattern of the sky, or a combination of these cues as compass signals. Behavioral experiments in bees and ants, indeed, showed that direct sunlight and sky polarization play a role in sky compass orientation, but the relative importance of these cues are species-specific. Intracellular recordings from polarization-sensitive interneurons in the desert locust and monarch butterfly suggest that inputs from different eye regions, including polarized-light input through the dorsal rim area of the eye and chromatic/intensity gradient input from the main eye, are combined at the level of the medulla to create a robust compass signal. Conflicting input from the polarization and chromatic/intensity channel, resulting from eccentric receptive fields, is eliminated at the level of the anterior optic tubercle and central complex through internal compensation for changing solar elevations, which requires input from a circadian clock. Across several species, the central complex likely serves as an internal sky compass, combining E-vector information with other celestial cues. Descending neurons, likewise, respond both to zenithal polarization and to unpolarized cues in an azimuth-dependent way.

  8. «Since the interest of the community is superior to that of the individuals»: fiscal discourse and political identity in 15th century Cervera

    Directory of Open Access Journals (Sweden)

    Pere Verdés Pijuan


    Full Text Available Just as taxation contributes decisively to the institutional development of the local government, so the discourse used to justify or criticize the fiscal and financial policies of its rulers constitutes a fundamental element in the formation of a communal identity in Catalonia during the later Middle Ages. This is what can be gathered from an analysis of the language and discursive strategies used in Cervera, both by the local authorities and by their critics, throughout the fifteenth century. In light of the references contained in the council minute books and other municipal sources, the conflicts caused by royal exactions and the payments of public debt, by the establishment of one form of taxation or another, and by the administration of communal finances, allow us to appreciate very clearly how far the idea of the common good made its way into the collective urban imagination. As the title of this article suggests, in any given situation we can find references to the idea that the general interest of the universitas and the benefit of its republic always should be given preference to that of an individual (including the monarch, and that this should be done following a specific series of ethical and moral principles, such as charity, trust, credibility and so on -all of them according to the communal rhetoric developed by the Franciscan theorists of the period.

  9. Discursive Practices of Mediatic Violence in the Case of Gezi Park Protests

    Directory of Open Access Journals (Sweden)

    Deniz ÇABA


    Full Text Available Although many ideologies today construct themselves with a liberal discourse, instead of universal understanding of human beings, a reality is drawn over the ideological rhetoric of the state and made dominant over the social. The legitimate violent monarch and the “obedient citizen” argument of the modern state is supposedly constituted by libertarian practices this time. The statist language in the mainstream media is also inevitably questioned on this basis. In this context as a social movement, Gezi Park Protests stand out as an example case in which many concepts can be questioned. The purpose of this study is to determine which ideological codes were used to turn the events of Gezi Park Protests into news in Hürriyet Newspaper, and to question how the concepts of “state” and “citizen” are defined through the “representation of violence”. For this purpose, 16 news items about Gezi Park Protests in Hürriyet Newspaper between 31 May-3 June 2013 were analyzed by using Van Dijk’s method of critical discourse analysis. In the end of the study, it became clear that Hürriyet Newspaper legitimized the monopoly of violence by the state and fixed the meaning in the “obedient citizen” retribution by dividing those who participated in the acts into the acceptable and marginal activist duality.

  10. “…We do not know, who are they, where are they from and what are their aims”. The Mongol Expansion in the Light of Published Latin Sources

    Directory of Open Access Journals (Sweden)

    F.N. Veselov


    Full Text Available This article is primarily dedicated to the monograph of R. Hautala “From ‘David, King of the Indies’ to ‘Detestable Plebs of Satan’: An Anthology of Early Latin Information about the Tatar-Mongols”. One can find that this anthology is the most complete collection of Latin sources, which were created before the Mongol invasion of the European states and concerning the Mongols, their genesis and aims of the Western campaign. The authors state that the mentioned study includes not only the high-level presentation of handwritten artifacts, but extensive research, providing a number of original and decisive conclusions as well. On the other hand, the authors of the paper mention complexes of sources, which could provide more detailed image of diplomatic relations between European monarchs and heads of the Mongol armies, Christian interpretations of threating and unknown enemy who unexpectedly appeared on the boarders of Catholic world. Particularly, the authors point on the fragment from Alberic of Trois-Fontaines, which could be described not only as a source on the Russian-Mongol battle, as well as on the sources on diplomatic relations between the Holy Roman Empire and Mongol rulers, Russian sources that had own impact on the European scholars’ eschatological interpretations of the Tatar invasion.

  11. On the tithes/subsidies dilemma in Castile and the Crown of Aragón in the Late Middle Ages

    Directory of Open Access Journals (Sweden)

    Jordi Morelló Baget


    Full Text Available This work is a diachronic and comparative analysis of the lengthy process in the Crown of Aragon and Castile, involving two forms of extraordinary taxation (tithes and subsidies, which can be considered representative of a major transfer of ecclesiastical income in favour of the emerging tax state. We follow successive chronological phases from the thirteenth century to the reign of the Catholic Monarchs, to ascertain how often the monarchy made use of each type of levy —imposed with or without papal authorization— the different reasons given in each case, and other issues concerning the consent of the clergy and the role of church assemblies in the management of these payments. Despite the divergences and different dynamics that developed in each kingdom, it is clear that both forms of clerical contribution led to the imposition of more regular income taxes, while establishing a fiscality based on the taxation of ecclesiastical gains. The work has been prepared from a selective review of bibliographical output and the consultation of primary sources relating to the Crown of Aragon.

  12. El juicio de residencia como ritual político en la colonia (Gobernación de Tucumán, siglo XVIII: A political ritual. Tucumán 18th Century Impeachment proceeding in Colonial times

    Directory of Open Access Journals (Sweden)

    Silvina Smietniansky


    Full Text Available Este artículo se propone analizar un juicio de residencia, llevado a cabo en los años 1775 y 1776 en la gobernación de Tucumán, como un ritual político. Entendemos que, al teatralizar una forma ideal de funcionamiento de las instituciones y comportamiento de los funcionarios, el juicio de residencia como ritual político actuaba fortaleciendo el poder monárquico en las colonias, transmitiendo un conjunto de valores y congregando y cohesionando a la comunidad local. Partiendo de esta perspectiva ritual, abordamos el problema de la tensión existente entre las normas y objetivos que ordenaban la institución de la residencia y las implicancias y consecuencias que, en la práctica, acarreaba su celebración.This article analyzes an impeachment proceeding that took place in 1775-1776 in colonial Tucumán, in terms of a political ritual. We understand that by performing the ideal way of government institution's operation and officials' behaviour, the impeachment proceeding acted as a political ritual, reinforcing monarchical power in the colonies, transmitting a set of social values and congregating the local community. From this ritual perspective, we approach the problem of the tension between the norms and aims of the impeachment procedure, and the consequences and implications of its practice.

  13. La singularidad de la monarquía sueca en el contexto europeo: el Rey como símbolo estático del Estado // The singularities of the Swedish monarchy in the European framework: The King as a static symbol of the State

    Directory of Open Access Journals (Sweden)

    Göran Rollnert Liern


    Then, it goes on with the discussion about the King’s powers in the travaux préparatoires of the new Constitution of 1975 that culminated with the so-called Compromise in Turekov in 1971 in which the Swedish political parties finally agreed on a constitutional design restricting the King’s powers, paying attention to the circumstances and reasons that explain why the Swedish monarch lost the formal powers that had coexisted since 1918 with the functioning of the parliamentary system, emphasizing the motivations of the parties represented in the Commission that drafted the new Instrument of Government. The paper concludes by referring to the striking exclusion of the Swedish Head of State from the Cabinet formation process in the Government Instrument of 1975 and the unusual variant of negative parliamentarism adopted in which the vote of the Prime Minister previous to his appointment is not really a vote of confidence but of «no no-confidence». With the constitutional reform of 2011, the novelty introduced was that, following the elections, the Prime Minister must undergo a mandatory vote of confidence, although the principle of negative parliamentarism is maintained essentially as long as he can continue in office if more than half of the members of Parliament do not vote against him, irrespective of the favorable votes he receives.

  14. Plant genotype shapes ant-aphid interactions: implications for community structure and indirect plant defense. (United States)

    Mooney, Kailen A; Agrawal, Anurag A


    Little is known about the mechanisms by which plant genotype shapes arthropod community structure. In a field experiment, we measured the effects of milkweed (Asclepias syriaca) genotype and ants on milkweed arthropods. Populations of the ant-tended aphid Aphis asclepiadis and the untended aphid Myzocallis asclepiadis varied eight- to 18-fold among milkweed genotypes, depending on aphid species and whether ants were present. There was no milkweed effect on predatory arthropods. Ants increased Aphis abundance 59%, decreased Myzocallis abundance 52%, and decreased predator abundance 56%. Milkweed genotype indirectly influenced ants via direct effects on Aphis and Myzocallis abundance. Milkweed genotype also modified ant-aphid interactions, influencing the number of ants attracted per Aphis and Myzocallis. While ant effects on Myzocallis were consistently negative, effects on Aphis ranged from antagonistic to mutualistic among milkweed genotypes. As a consequence of milkweed effects on ant-aphid interactions, ant abundance varied 13-fold among milkweed genotypes, and monarch caterpillar survival was negatively correlated with genetic variation in ant abundance. We speculate that heritable variation in milkweed phloem sap drives these effects on aphids, ants, and caterpillars. In summary, milkweed exerts genetic control over the interactions between aphids and an ant that provides defense against foliage-feeding caterpillars.

  15. Anthropogenic changes in sodium affect neural and muscle development in butterflies (United States)

    Snell-Rood, Emilie C.; Espeset, Anne; Boser, Christopher J.; White, William A.; Smykalski, Rhea


    The development of organisms is changing drastically because of anthropogenic changes in once-limited nutrients. Although the importance of changing macronutrients, such as nitrogen and phosphorus, is well-established, it is less clear how anthropogenic changes in micronutrients will affect organismal development, potentially changing dynamics of selection. We use butterflies as a study system to test whether changes in sodium availability due to road salt runoff have significant effects on the development of sodium-limited traits, such as neural and muscle tissue. We first document how road salt runoff can elevate sodium concentrations in the tissue of some plant groups by 1.5–30 times. Using monarch butterflies reared on roadside- and prairie-collected milkweed, we then show that road salt runoff can result in increased muscle mass (in males) and neural investment (in females). Finally, we use an artificial diet manipulation in cabbage white butterflies to show that variation in sodium chloride per se positively affects male flight muscle and female brain size. Variation in sodium not only has different effects depending on sex, but also can have opposing effects on the same tissue: across both species, males increase investment in flight muscle with increasing sodium, whereas females show the opposite pattern. Taken together, our results show that anthropogenic changes in sodium availability can affect the development of traits in roadside-feeding herbivores. This research suggests that changing micronutrient availability could alter selection on foraging behavior for some roadside-developing invertebrates. PMID:24927579

  16. The organization of executive power in Spain (1808 -1810. Reflections on an unedited text by Jovellanos

    Directory of Open Access Journals (Sweden)

    Ignacio Fernández Sarasola


    Full Text Available The discovery of an unpublished document written by Jovellanos about the organization of the Presidency of the Junta Central allows for a reassessment of his relevance in the configuration of the executive power during the early years after the War of Independence. After explaining where the document was found, as well as confirming its authorship and when it was written, I will analyze the document’s context. The conclusion of this analysis is that many of the measures relevant to the structure of the Junta Central came from Jovellanos, who wanted said organization to be able to exercise genuine political power, yet to be organized in the most convenient way in order to be able to reflect upon the political and administrative reforms once the Cortes met. Throughout the study, I will show how Jovellanos tried to reflect his political theories on sovereignty and the separation of powers in the structure of the Junta Central, but also how he altered his initial beliefs due to the convulsive political situation in Spain during the War of Independence. Additionally, it will be shown that the blueprint of internal executive power obligated Jovellanos to also define the competencies of the future Regency, of the Juntas Provinciales and even of the monarch himself. This work employs a methodology typical to constitutional history, integrating an analysis of political thought, the contents of law and institutional development. In order to create this document, direct sources have been primarily used, many of which are relatively unknown.

  17. Representations of the Spanish Civil War in early Francoist history textbooks

    Directory of Open Access Journals (Sweden)

    Christian ROITH


    Full Text Available Textbooks are effective tools to understand the idiosyncrasies of certain historical time periods as they allow us to examine the ideologies, concepts and values of the social groups in power. Those textbooks which were in use in the areas occupied by the Fascist National troops under the insurgent General Francisco Franco’s command and in all of Spain after his victory permit us to draw conclusions about the characteristics of the Francoist ideology. The analysis of original material collected and classified by the Spanish research project of textbooks (manes, as well as the review of theoretical studies demonstrates that Francoist schoolbooks to a large extent reproduced contents proceeding from the monarchic period before the Second Republic. The representations of the Civil War in these textbooks condense the most important doctrinal principles of the Franco dictatorship and reveal the existing ideological differences between different tendencies represented in the regime. Furthermore, the analysis of these texts shows that the ideological indoctrination through schoolbooks was an essential element of the totalitarian efforts of the Franco regime.

  18. Enfranchised Minors: Women as People in the Middle East after the 2011 Arab Uprisings

    Directory of Open Access Journals (Sweden)

    Rania Maktabi


    Full Text Available The civic status of female citizens in the Middle East and North Africa (MENA region is conceptualized as “enfranchised minorhood” which reflects the confined position of adult women as legal minors under the trusteeship of male kin in family law, criminal law, and nationality law. During and in the aftermath of the Uprisings that erupted throughout MENA in 2011, female lawyers in Morocco, Lebanon, and Kuwait allied with women’s groups and pressured for reforms in patriarchal state laws. By 2015, reforms were manifest in criminal law; incremental in family law; and absent in nationality law. Theoretical conclusions based on comparative analysis of societal pressures in three states indicate that long historical trajectories are imperative for substantiating the expansion of female citizenship following the 2011 Uprisings. Additionally, the civic status of women in the MENA region is being strengthened under authoritarian monarchical rule in Kuwait and Morocco. A third finding is that pressures for reform have more visible reverberations in legal spheres with a clerical imprint such as family law and criminal law, while strengthened pressures in a secular legal sphere such as nationality law have been opposed more forcefully five years after the Uprisings.

  19. Flying Drosophila orient to sky polarization. (United States)

    Weir, Peter T; Dickinson, Michael H


    Insects maintain a constant bearing across a wide range of spatial scales. Monarch butterflies and locusts traverse continents [1, 2], and foraging bees and ants travel hundreds of meters to return to their nests [1, 3, 4], whereas many other insects fly straight for only a few centimeters before changing direction. Despite this variation in spatial scale, the brain region thought to underlie long-distance navigation is remarkably conserved [5, 6], suggesting that the use of a celestial compass is a general and perhaps ancient capability of insects. Laboratory studies of Drosophila have identified a local search mode in which short, straight segments are interspersed with rapid turns [7, 8]. However, this flight mode is inconsistent with measured gene flow between geographically separated populations [9-11], and individual Drosophila can travel 10 km across desert terrain in a single night [9, 12, 13]-a feat that would be impossible without prolonged periods of straight flight. To directly examine orientation behavior under outdoor conditions, we built a portable flight arena in which a fly viewed the natural sky through a liquid crystal device that could experimentally rotate the polarization angle. Our findings indicate that Drosophila actively orient using the sky's natural polarization pattern. Copyright © 2012 Elsevier Ltd. All rights reserved.

  20. Design and Calibration of a Novel Bio-Inspired Pixelated Polarized Light Compass

    Directory of Open Access Journals (Sweden)

    Guoliang Han


    Full Text Available Animals, such as Savannah sparrows and North American monarch butterflies, are able to obtain compass information from skylight polarization patterns to help them navigate effectively and robustly. Inspired by excellent navigation ability of animals, this paper proposes a novel image-based polarized light compass, which has the advantages of having a small size and being light weight. Firstly, the polarized light compass, which is composed of a Charge Coupled Device (CCD camera, a pixelated polarizer array and a wide-angle lens, is introduced. Secondly, the measurement method of a skylight polarization pattern and the orientation method based on a single scattering Rayleigh model are presented. Thirdly, the error model of the sensor, mainly including the response error of CCD pixels and the installation error of the pixelated polarizer, is established. A calibration method based on iterative least squares estimation is proposed. In the outdoor environment, the skylight polarization pattern can be measured in real time by our sensor. The orientation accuracy of the sensor increases with the decrease of the solar elevation angle, and the standard deviation of orientation error is 0 . 15 ∘ at sunset. Results of outdoor experiments show that the proposed polarization navigation sensor can be used for outdoor autonomous navigation.

  1. Defenders and Conquerors: The Rhetoric of Royal Power in Korean Inscriptions from the Fifth to Seventh Centuries

    Directory of Open Access Journals (Sweden)

    Hung-gyu Kim


    Full Text Available This article compares the rhetoric of three inscriptions from the Three Kingdoms period of Korea: the Gwanggaeto inscription, which was carved in 414 on the tomb stele of King Gwanggaeto of Goguryeo; the inscriptions on the monument stones raised between 550 and 568 to record the tours of King Jinheung of Silla; and the King Munmu inscription on the funerary stele of King Munmu of Silla, which was completed in 682. Notably, although all three monarchs were successful warriors, only the Gwanggaeto inscription is characterized by the martial rhetoric of conquest, while the two Silla examples employ the cautious rhetoric of peacemaking. The author analyzes this difference by understanding the inscriptions as situated speech acts, and he suggests that these inscriptions should be understood within the particular political circumstances in which they were situated. Whereas the Gwanggaeto inscription was produced by a powerful Goguryeo acting within a Northeast Asia in which no one state could claim dominance, the Silla inscriptions were produced by a Silla Kingdom that had to struggle first against established Korean rivals and then against an enormously powerful Tang empire.

  2. AHP 35: Review: TIBET WILD

    Directory of Open Access Journals (Sweden)

    William V Bleisch


    Full Text Available Es sieht ein Mondenshcatten Als mein Gefrährte mit, Und aug den wei en Matten Such ich des Wildes Tritt….. Wilhelm Müller, Gute Nacht George Schaller's remarkable career spans nearly six decades of work resulting in field studies of wildlife in the most remote regions, including pioneering investigations on four continents. More than half of that time was spent involved with studies of the wildlife of the Tibetan Plateau and neighboring regions. Following each new phase of his career, from his work on mountain gorillas in Rwanda, tigers in India, lions on the Serengeti, wild sheep in the Himalayas, and Tibetan antelope and other wildlife on the Tibetan steppes, he has made the time to publish a book on each of his expeditions – or more exactly, two (see full list in Appendix. One is always a scholarly monograph full of data, tables, and maps, the other a popular account for the general public. These paired volumes are usually published within one year of each other, and there have been six such pairings so far. For example, Schaller's classic the Mountain Monarchs: Wild Sheep and Goats of the Himalaya was published in 1978; in 1980, he published Stones of Silence: Journeys in the Himalaya; in 1997 he published the popular Tibet's Hidden Wilderness: Wildlife and Nomads of the Chang Tang Reserve; and the next year, 1998, saw the appearance of his scholarly monograph Wildlife of the Tibetan Steppe. ...

  3. Nursing leadership and management effects work environments. (United States)

    Tomey, Ann Marriner


    The aim of this literature search was to identify recent research related to nursing leadership and management effects on work environment using the 14 forces of magnetism. This article gives some historical perspective from the original 1983 American Academy of Nursing study through to the 2002 McClure and Hinshaw update to 2009 publications. Research publications were given a priority for references. The 14 forces of magnetism as identified by Unden and Monarch were: '1. Quality of leadership..., 2. Organizational structure..., 3. Management style..., 4. Personnel policies and programs..., 5. Professional models of care..., 6. Quality of care..., 7 Quality improvement..., 8. Consultation and resources..., 9. Autonomy..., 10. Community and the hospital..., 11. Nurse as teacher..., 12. Image of nursing..., 13. Interdisciplinary relationships... and 14. Professional development....'. Correlations have been found among positive workplace management initiatives, style of transformational leadership and participative management; patient-to-nurse ratios; education levels of nurses; quality of patient care, patient satisfaction, employee health and well-being programmes; nurse satisfaction and retention of nurses; healthy workplace environments and healthy patients and personnel. This article identifies some of the research that provides evidence for evidence-based nursing management and leadership practice.

  4. Physics in Edinburgh: From Napier's Bones to Higgs's Boson (United States)

    Henry, John

    Edinburgh has been the capital of Scotland since 1437 and by any standards is a beautiful city. It makes an immediate impact upon the visitor because major aspects of its history are there for all to see in the general arrangement of the city, and in its buildings. The Castle, sitting on the prominent summit of one of the volcanic plugs that dominate the topography of the city, is a constant focus for the eye of the wandering tourist. From the Castle Esplanade a downward-sloping road running eastward forms the spine of the Old Town. This road is known to tourists as the Royal Mile, but it actually consists of three parts with separate names: the Lawnmarket, the High Street, and the Canongate. The eastern end of the Canongate culminates at the ruined Holyrood Abbey (granted a Royal Charter in 1124) and the adjacent Palace of Holyrood House, the principal residence of the Scottish monarch since the fifteenth century (although rumored to be the least favorite of the present queen’s residences), and now the new Scottish Parliament building (officially opened in 2004). It is a Royal Mile, then, because it stretches from Castle to Palace.

  5. Discussion of FAN Zhong-yan’s Buddhist Thought and Personality Spirit%论范仲淹的佛禅思想及其人格精神

    Institute of Scientific and Technical Information of China (English)

    邢爽; 胡遂


    FAN Zhong-yan’s life ambition is to make the monarch as wise as Yao and Shun.He begged the ancient heart of Habitat for Humanity in his “YueYang Tower Notes”,“not pleased,not to have com-passion,”the implication was obvious Buddhist ideas with.The fusion of Confucianism and Buddhism thought of Fan Zhong-yan for the later generations to establish the “independent personality model un-moved either by gain or loss”.His life is spent in the pursuit and practice of Mahayana Bodhisattva “the highest state of compassion for the world of vision”,“being and not being”in perfect unity.%范仲淹的一生志在致君尧舜。《岳阳楼记》中他“尝求古仁人之心”,“不以物喜,不以己悲”的思想俨然带有佛家的意味。融合儒释思想的范仲淹为后来的士人树立了“宠辱不惊”的独立人格典范。他的一生都在追求和践行大乘菩萨道“悲心宏愿”的最高境界,为世人诠释了“有无之境”的完美统一。

  6. A formação da diplomacia econômica do Brasil The formation of Brazilian economic diplomacy

    Directory of Open Access Journals (Sweden)

    Paulo Roberto de Almeida


    Full Text Available Ensaio de caráter histórico sobre as grandes linhas da diplomacia econômica no Brasil, com ênfase no período monárquico. Depois de uma identificação das questões metodológicas próprias ao estudo da diplomacia econômica no Brasil, em suas etapas formadoras, são apresentados os problemas que mobilizaram a atenção do establishment diplomático imperial, em sua vertente propriamente econômica, bem como ressaltados os elementos de ruptura e de continuidade em relação à diplomacia econômica do século XX.Historical essay focussing the main trends of Brazil's economic diplomacy, in special at its earlier stages, during the monarchic period. Following a brief discussion of methodological issues linked to the study of economic diplomacy in Brazil, the analysis centers on the relevant questions that mobilized the attention of the Brazilian diplomatic establishment. Appropriate consideration is given to elements of innovation or continuity between the economic diplomacy of the XIX century and that of the XX century.

  7. Beneficial Insect Attraction to Milkweeds (Asclepias speciosa, Asclepias fascicularis in Washington State, USA

    Directory of Open Access Journals (Sweden)

    David G. James


    Full Text Available Native plant and beneficial insect associations are relatively unstudied yet are important in native habitat restoration programs for improving and sustaining conservation biological control of arthropod pests in agricultural crops. Milkweeds (Asclepias spp. are currently the focus of restoration programs in the USA aimed at reversing a decline in populations of the milkweed-dependent monarch butterfly (Danaus plexippus; however, little is known of the benefits of these plants to other beneficial insects. Beneficial insects (predators, parasitoids, pollinators attracted to two milkweed species (Asclepias speciosa, Asclepias fascicularis in central Washington State, WA, USA were identified and counted on transparent sticky traps attached to blooms over five seasons. Combining all categories of beneficial insects, means of 128 and 126 insects per trap were recorded for A. speciosa and A. fascicularis, respectively. Predatory and parasitic flies dominated trap catches for A. speciosa while parasitic wasps were the most commonly trapped beneficial insects on A. fascicularis. Bees were trapped commonly on both species, especially A. speciosa with native bees trapped in significantly greater numbers than honey bees. Beneficial insect attraction to A. speciosa and A. fascicularis was substantial. Therefore, these plants are ideal candidates for habitat restoration, intended to enhance conservation biological control, and for pollinator conservation. In central Washington, milkweed restoration programs for enhancement of D. plexippus populations should also provide benefits for pest suppression and pollinator conservation.

  8. Beneficial Insect Attraction to Milkweeds (Asclepias speciosa, Asclepias fascicularis) in Washington State, USA. (United States)

    James, David G; Seymour, Lorraine; Lauby, Gerry; Buckley, Katie


    Native plant and beneficial insect associations are relatively unstudied yet are important in native habitat restoration programs for improving and sustaining conservation biological control of arthropod pests in agricultural crops. Milkweeds (Asclepias spp.) are currently the focus of restoration programs in the USA aimed at reversing a decline in populations of the milkweed-dependent monarch butterfly (Danaus plexippus); however, little is known of the benefits of these plants to other beneficial insects. Beneficial insects (predators, parasitoids, pollinators) attracted to two milkweed species (Asclepias speciosa, Asclepias fascicularis) in central Washington State, WA, USA were identified and counted on transparent sticky traps attached to blooms over five seasons. Combining all categories of beneficial insects, means of 128 and 126 insects per trap were recorded for A. speciosa and A. fascicularis, respectively. Predatory and parasitic flies dominated trap catches for A. speciosa while parasitic wasps were the most commonly trapped beneficial insects on A. fascicularis. Bees were trapped commonly on both species, especially A. speciosa with native bees trapped in significantly greater numbers than honey bees. Beneficial insect attraction to A. speciosa and A. fascicularis was substantial. Therefore, these plants are ideal candidates for habitat restoration, intended to enhance conservation biological control, and for pollinator conservation. In central Washington, milkweed restoration programs for enhancement of D. plexippus populations should also provide benefits for pest suppression and pollinator conservation.

  9. The chemistry of antipredator defense by secondary compounds in neotropical lepidoptera: facts, perspectives and caveats

    Directory of Open Access Journals (Sweden)

    Trigo José R.


    Full Text Available Chemical defense against predation in butterflies and moths has been studied since nineteenth century. A classical example is that of the larvae of the monarch butterfly Danaus plexippus, which feed on leaves of Asclepias curassavica (Asclepiadaceae, sequestering cardenolides. The adults are protected against predation by birds. Several other substances may be involved in chemical defense, such as iridoid glycosides, cyanogenic glycosides, glucosinolates, pyrrolizidine and tropane alkaloids, aristolochic acids, glycosidase inhibitors and pyrazines. The acquisition of these substances by lepidopterans can be due to sequestration from larval or adult host plants or de novo biosynthesis. Many Lepidoptera are known to be unpalatable, including the butterflies Troidini (Papilionidae, Pierinae (Pieridae, Eurytelinae, Melitaeinae, Danainae, Ithomiinae, Heliconiinae and Acraeinae (Nymphalidae, and Arctiidae moths, but knowledge of the chemical substances responsible for property is often scarce. This review discusses mainly three topics: field and laboratory observations on rejection of butterflies and moths by predators, correlation between unpalatability and chemicals found in these insects, and bioassays that test the activity of these chemicals against predators. Perspectives and future directions are suggested for this subject.

  10. Phylogenetic ecology of leaf surface traits in the milkweeds (Asclepias spp.): chemistry, ecophysiology, and insect behavior. (United States)

    Agrawal, Anurag A; Fishbein, Mark; Jetter, Reinhard; Salminen, Juha-Pekka; Goldstein, Jessica B; Freitag, Amy E; Sparks, Jed P


    The leaf surface is the contact point between plants and the environment and plays a crucial role in mediating biotic and abiotic interactions. Here, we took a phylogenetic approach to investigate the function, trade-offs, and evolution of leaf surface traits in the milkweeds (Asclepias). Across 47 species, we found trichome densities of up to 3000 trichomes cm(-2) and epicuticular wax crystals (glaucousness) on 10 species. Glaucous species had a characteristic wax composition dominated by very-long-chain aldehydes. The ancestor of the milkweeds was probably a glaucous species, from which there have been several independent origins of glabrous and pubescent types. Trichomes and wax crystals showed negatively correlated evolution, with both surface types showing an affinity for arid habitats. Pubescent and glaucous milkweeds had a higher maximum photosynthetic rate and lower stomatal density than glabrous species. Pubescent and glaucous leaf surfaces impeded settling behavior of monarch caterpillars and aphids compared with glabrous species, although surface types did not show consistent differentiation in secondary chemistry. We hypothesize that pubescence and glaucousness have evolved as alternative mechanisms with similar functions. The glaucous type, however, appears to be ancestral, lost repeatedly, and never regained; we propose that trichomes are a more evolutionarily titratable strategy.

  11. Evolution of plant growth and defense in a continental introduction. (United States)

    Agrawal, Anurag A; Hastings, Amy P; Bradburd, Gideon S; Woods, Ellen C; Züst, Tobias; Harvey, Jeffrey A; Bukovinszky, Tibor


    Substantial research has addressed adaptation of nonnative biota to novel environments, yet surprisingly little work has integrated population genetic structure and the mechanisms underlying phenotypic differentiation in ecologically important traits. We report on studies of the common milkweed Asclepias syriaca, which was introduced from North America to Europe over the past 400 years and which lacks most of its specialized herbivores in the introduced range. Using 10 populations from each continent grown in a common environment, we identified several growth and defense traits that have diverged, despite low neutral genetic differentiation between continents. We next developed a Bayesian modeling approach to account for relationships between molecular and phenotypic differences, confirming that continental trait differentiation was greater than expected from neutral genetic differentiation. We found evidence that growth-related traits adaptively diverged within and between continents. Inducible defenses triggered by monarch butterfly herbivory were substantially reduced in European populations, and this reduction in inducibility was concordant with altered phytohormonal dynamics, reduced plant growth, and a trade-off with constitutive investment. Freedom from the community of native and specialized herbivores may have favored constitutive over induced defense. Our replicated analysis of plant growth and defense, including phenotypically plastic traits, suggests adaptive evolution following a continental introduction.

  12. The ecology and evolution of animal medication: genetically fixed response versus phenotypic plasticity. (United States)

    Choisy, Marc; de Roode, Jacobus C


    Animal medication against parasites can occur either as a genetically fixed (constitutive) or phenotypically plastic (induced) behavior. Taking the tritrophic interaction between the monarch butterfly Danaus plexippus, its protozoan parasite Ophryocystis elektroscirrha, and its food plant Asclepias spp. as a test case, we develop a game-theory model to identify the epidemiological (parasite prevalence and virulence) and environmental (plant toxicity and abundance) conditions that predict the evolution of genetically fixed versus phenotypically plastic forms of medication. Our model shows that the relative benefits (the antiparasitic properties of medicinal food) and costs (side effects of medicine, the costs of searching for medicine, and the costs of plasticity itself) crucially determine whether medication is genetically fixed or phenotypically plastic. Our model suggests that animals evolve phenotypic plasticity when parasite risk (a combination of virulence and prevalence and thus a measure of the strength of parasite-mediated selection) is relatively low to moderately high and genetically fixed medication when parasite risk becomes very high. The latter occurs because at high parasite risk, the costs of plasticity are outweighed by the benefits of medication. Our model provides a simple and general framework to study the conditions that drive the evolution of alternative forms of animal medication.

  13. Remarkable Women in a Remarkable Age. On the Genesis of the English Public Sphere, 1642-1752

    Directory of Open Access Journals (Sweden)

    Eleonora Cappuccilli


    Full Text Available During the era of the English Revolutions and shortly after that, some spaces, albeit limited, of female visibility open up. Thanks to the window of opportunity caused by the collapse of censorship, the participation in the radical sects and in the Civil war, some remarkable women succeed in introducing themselves in the public sphere, shaping it since its very genesis. Moreover, analysing law institutions as jointure and feoffment, the attempt is to reconstruct some fragments of juridical female autonomy, which belie the total pervasiveness of coverture in the XVII century. As in private law, in public law women, in the role of queens, gain centrality: the principle of female authority, while safeguards the holding of the monarchical regime, destabilizes its patriarchal structure. Going through the works of Katherine Chidley, Margaret Cavendish, Damaris Masham and Mary Astell, the essay aims at reconstructing women's public voice, a voice which upsets the consolidated frames and subverts the established positions, questioning the same social hierarchies.

  14. The queen, the prince, and the ideologue: Alonso Ortiz’s notions of queenship at the court of the Catholic Kings

    Directory of Open Access Journals (Sweden)

    Silleras-Fernández, Núria


    Full Text Available In the court of the Catholic Monarchs (r. 1474-1516, Isabel I of Castile and Fernando II of Aragon, that of a ruling queen, humanists constructed theories of what it meant to be a ruler. Alonso Ortiz, one of the humanists attached to their court, composed the Latin text, Liber de educatione Johannis Serenissimi Principis et primogeniti regum potentissimorum Castelle Aragonum et Siciliae Ferdinandi et Helisabet inclyta prosapia coniugum clarissimorum, a book that reflects an idealized and disempowered humanist vision of elite women that contrasts with the agency they wielded in historical fact.En la corte de los Reyes Católicos (1474-1516, Isabel I de Castilla y Fernando II de Aragón, una corte en la que una reina gobernaba, los humanistas construyeron teorías sobre lo que significaba ser un gobernante. Alonso Ortiz, un humanista de su corte, compuso el texto latino: Liber de educatione Johannis Serenissimi Principis et primogeniti regum potentissimorum Castelle Aragonum et Siciliae Ferdinandi et Helisabet inclyta prosapia coniugum clarissimorum, libro que refleja una visión idealizada y despojada de poder de las mujeres de la élite, hecho este que contrasta con la capacidad de actuación que éstas tuvieron según la evidencia histórica.

  15. Orientalism and the geoculture of the World System: Discursive othering, political economy and the cameralist division of labor in Habsburg Central Europe (1713-1815

    Directory of Open Access Journals (Sweden)

    Klemens Kaps


    Full Text Available This article addresses the question of to what degree the concept of geoculture can be brought in line with research on Orientalist stereotypes and imaginary. Following Said’s original definition of orientalism discourses of the 18th-century political economy are reassessed by focusing on their perception of spatial hierarchies in Eastern Europe. This article reconsiders these discourses as an active factor in the struggle for power and a tool in the hands of the geopolitical interests of absolutist monarchs in Prussia, the Habsburg Monarchy, and Russia in the age of mercantilism, as demonstrated by the Partitions of Poland-Lithuania. By focusing on the Habsburg Monarchy between the Spanish War of Succession and the Congress of Vienna, it is demonstrated here that, territorial landlocked empires within Europe used a similar language as colonial maritime empires in order to justify their geopolitical expansion and territorial domination of Eastern Europe. In a second step, it is shown that this discourse was part of the geopolitical culture of the World System and was instrumental in setting ideological conditions for cameralist-driven institutional transformations in favor of the core regions within the Habsburg dominions in Central Europe.

  16. Kaiser Franz Joseph I und die Serben 1848–1908

    Directory of Open Access Journals (Sweden)

    Wolfgang Rohrbach


    Full Text Available A century from death of Emperor Franz Joseph I (+21.11.1916 triggered a great number of publications about life and almost seven decades long reign of a popular monarch. As the political struggle between Russia, Prussia, Italy and Ottoman Empire are depicted with moderation, Serbia is described unidimensional, with an emphasis on the last decade of Emperors rule. In order to remedy this shortcoming, this contribution is focusing on a period between 1848 and 1908. Today we know much more about relations between Franz Joseph and King Milan Obrenović, as well as with dignitaries of Serbian Orthodox Church, and military, academic and artistic elite. It is, therefore, possible to refute entrenched notions about constant enmity and hostility between Austria and Serbia. In this work, an emphasis is put on cooperation between Austria and Serbia from 1848, when Serbs at Emperors request held a number of important political and military positions, whereas Serbian artists took part in the reconstruction of Imperial Vienna. The university of Vienna hosted the oldest institute for slavistic (1849, becoming, therefore, a crossroad of the cultural and social development of the Balkans.

  17. The alignment of customs duties to the interests of settlers, commerce and politics. Andalusia, 1241-1550

    Directory of Open Access Journals (Sweden)

    José Damián González Arce


    Full Text Available This paper aims to analyze the royal policy of exemptions and reductions of customs duties in Andalusia during the Late Middle Ages. In particular, the taxes studied are the almojarifazgo, tithes, the Moorish half-tithe imposed on imports and exports with Granada, and other marginal taxes, such as rodas, castillerías and tolls. The granting of these franchises reveals the economic and political interests of monarchs in particular periods and circumstances. Their reduction or elimination, on the other hand, shows the changes over time that removed or reduced the causes of depopulation, border insecurity and economic backwardness that motivated the exemptions. The documentation used is from the Archivo General de Simancas and the municipal archives of Seville, Cordoba, Carmona and Jerez. Finally, we conclude that tax advantages served to attract population, and that only the most exposed or strategic populations were able to remain more or less intact over time, especially when they were also key elements for foreign trade.

  18. Roughness Effects on the Formation of a Leading Edge Vortex (United States)

    Elliott, Cassidy; Lang, Amy; Wahidi, Redha; Wilroy, Jacob


    Microscopic scales cover the wings of Monarch butterflies, creating a patterned surface. This patterning is an important natural flow control mechanism that is thought to delay the growth of the leading edge vortex (LEV) produced by the flapping motion of a wing. The increased skin friction caused by the scales leads to a weaker LEV being shed into the butterfly's wake, lessening drag and increasing flight efficiency. To test this theory, a plate of random roughness was designed in SolidWorks and printed on the Objet 30 Pro 3D printer. A 2x3x5 cubic foot tow tank was used to test the rough plate at Reynold's numbers of 1500, 3000, and 6000 (velocities of 8, 16, and 32 mm/s) at an angle of attack of 45 degrees. Particle Image Velocimetry (PIV) captured images of the LEV generated by the plate when towed upwards through the particle-seeded flow. Codes written in MatLab were used to automatically track and determine the strength of the LEV. Circulation values for the randomly-rough plate were then compared to the same values generated in a previous experiment that used a smooth plate and a grooved plate to determine the effect of the patterning on vortex development. Funding provided by NSF REU site Grant EEC 1358991 and CBET 1628600.

  19. Biopolitical mills. Topographies of Power in Early Peripheral Capitalism

    Directory of Open Access Journals (Sweden)

    Wiktor Marzec


    Full Text Available City of Lodz is an exceptional case of industrialsettlement, a focal point of di}erent kinds and techniquesof power typical for nineteenth century industrial capi-talism. Disciplinary power parceling bodies, biopowerproviding with population stability, paternalistic gaze ofthe factory owner and monarchic sovereignty of the tsa-rist rule once met in the Scheibler and Grohman’s indu-strial establishment and nearby workers’ housing estate.{is peripheral capitalism and relations accompanyingit let us verify Foucaultian analysis of power and have anew look at it. Di}erent and less stable pattern of corre-lation of power techniques emerges here; power is nolonger strictly related to the temporal matrix or a func-tional demands of capitalist production, and temporarilyossi/es in relocated and contingent con/gurations. It isan implicit illustration of a new paradigm of power ana-lysis, which Foucault presented when his interest was notan industrial city any more, namely the „topologicalanalysis”, as S. J. Collier aptly called it.

  20. Integrar y excluir (comunión y excomunión en el Medievo

    Directory of Open Access Journals (Sweden)

    Mitre Fernández, Emilio


    Full Text Available The medieval excommunications, in its various manifestations, have biblical, patristic and canonical roots. During the Medieval Ages it is applicable to several situations and to all the people but the most known is that one applied to monarchs. The later Middle Ages increased the critics to this expedient made by relevant authors (Marsilius of Padua or William of de Ockham and politic organisms: vg. the Cortes of Castile.La excomunión medieval, en sus diversas manifestaciones tiene unas raíces bíblicas, patrísticas y canónicas. A lo largo del Medievo se aplicará en diversas situaciones y a toda clase de personas aunque las aplicadas por papas a monarcas sean las mas llamativas. La Baja Edad Media aumentará las críticas a este expediente por parte de destacados autores (Marsilio de Padua y Guillermo de Ockham a la cabeza y por parte de diversos organismos políticos: vg. las Cortes castellanas.

  1. Die »Verrechtlichung« heimlicher Ermittlungsmaßnahmen im Übergang vom Inquisitionsprozess des gemeinen Rechts zum reformierten Strafprozess des 19. Jahrhunderts

    Directory of Open Access Journals (Sweden)

    Pierre Hauck


    Full Text Available The »juridification« of covert investigations in the transition from the ius commune inquisition process to the reformed criminal justice process of the 19th century, understood as a material (i. e. law creating normative distinction between true or wrongful circumstances, is a complex phenomenon. This paper traces the introduction of covert investigations into the law of criminal procedure. It takes its starting point in the uncodified, but nevertheless legally sophisticated search and seizure of letters or the use of informers in the ius commune process. Influences of the French Revolution on the development of law, the 19th century’s early individual legislation, but mainly the tumultuous times before the Revolution of 1848 and the great surge of single procedural codes in the mid 19th century are the subject-matter of this analysis. This research has unearthed some surprising facts: Juridification affects various measures in quite different ways. At the same time, it is shaped by subjects which are hardly comparable with one another (monarchs, parliaments etc., and it is associated with the production of the criminal process as a subject that manifests itself particularly in the redefining of the pre-trial phase.

  2. Reflections on Peter Slezak and the 'Sociology of Scientific Knowledge` (United States)

    Suchting, W. A.

    The paper examines central parts of the first of two papers in this journal by Peter Slezak criticising sociology of scientific knowledge and also considers, independently, some of the main philosophical issues raised by the sociologists of science, in particular David Bloor. The general conclusion is that each account alludes to different and crucial aspects of the nature of knowledge without, severally or jointly, being able to theorise them adequately. The appendix contains epistemological theses central to a more adequate theory of scientific knowledge.... our Histories of six Thousand Moons make no Mention of any other, than the two great Empires of Lilliput and Blefuscu. Which mighty Powers have ... been engaged in a most obstinate War for six and thirty Moons past. It began upon the following Occasion. It is allowed on all Hands, that the primitive Way of breaking Eggs before we eat them, was upon the larger End: But ... the Emperor [of Lilliput] ... published an Edict, commanding all his Subjects, upon great Penalties, to break the smaller End of their Eggs. The People so resented this Law, that ... there have been six Rebellions raised on that Account ... These civil Commotions were constantly fomented by the Monarchs of Blefuscu ... It is computed, that eleven Thousand have, at several Times, suffered Death, rather than break Eggs at the smaller End. Many hundred large Volumes have published upon this Controversy ...

  3. Richard's back: death, scoliosis and myth making. (United States)

    Lund, Mary Ann


    The body of a mediaeval monarch was always under scrutiny, and Richard III's was no exception. In death, however, his body became subject to new forms of examination and interpretation: stripped naked after the battle of Bosworth, his corpse was carried to Leicester and exhibited before being buried. In 2012, it was rediscovered. The revelation that Richard suffered from scoliosis prompts this article to re-evaluate the historical sources about Richard's physique and his posthumous reputation. This article argues that Richard's death and his myth as 'crookback' are inextricably linked and traces attitudes to spinal curvature in the early modern period. It also considers how Shakespeare represented Richard as deformed, and aspects of performance history which suggest physical vulnerability. It then considers Richard's scoliosis from the perspective of medical history, reviewing classical accounts of scoliosis and arguing that Richard was probably treated with a mixture of axial traction and pressure. It demonstrates from the evidence of Richard's medical household that he was well placed to receive hands-on therapies and considers in particular the role of his physician and surgeon, William Hobbes. Finally, it shows how the case of Richard III demonstrates the close relationship between politics and medicine in the period and the contorted process of historical myth making. Published by the BMJ Publishing Group Limited. For permission to use (where not already granted under a licence) please go to

  4. El primer virreinato americano

    Directory of Open Access Journals (Sweden)

    Cassá, Roberto


    Full Text Available This article explores the government of viceroy Christopher Columbus in the American territories. We return to the first Spanish settlement in Santo Domingo and the contradictions inherent to this expansionist proyect. The contradictions were part of the logic of the absolutist state and Columbus’ reaction against the controls imposed by the monarchs. Secondly, we look into the dificulties that the Admiral encountered to develop a mercantilist model. In this context, we examine the rationale behind the first government of the Indies and the features that defined the new West Indian society.

    El artículo trata sobre el gobierno de Cristóbal Colón en tierras americanas. Retomamos el tema del primer emplazamiento español en Santo Domingo y las contradicciones que tuvo aquel proyecto debido a la lógica del estado absolutista, a la ambición desmedida del descubridor y a su reacción ante los controles que desde un principio impusieron los monarcas. En un segundo momento analizamos las dificultades que encontró el Almirante para desarrollar un modelo mercantilista acorde a sus ideas y a los acuerdos a que llegó con la Corona. En ese contexto analizamos la lógica del primer gobierno colombinista en las Indias y los rasgos que definieron la nueva sociedad antillana.

  5. La revolución en Quito: el camino hacia el gobierno mixto

    Directory of Open Access Journals (Sweden)

    Morelli, Federica


    Full Text Available This article studies the language, ideas and political models of the Quito elite in the formative period of the two earlier «juntas». In particular, the ideal of a «mixed government» inspired by various schools of thought such as scholasticism, natural law, and the republican tradition of the Renaissance and the modern period. This is, in our opinion, what explains the special nature of the 1812 constitutions and its two dimensions: monarchical and republican.

    Se estudian aquí el lenguaje, las ideas y los modelos políticos de la elite quiteña en la época de la formación de las dos primeras juntas. Se muestra su ideal de un gobierno mixto con raíces muy variadas que van de la escolástica y del iusnaturalismo a la tradición republicana renacentista y moderna. Se explica así la particularidad de la Constitución de 1812, a la vez monárquica y republicana.

  6. Las Constituciones catalanas de 1706 : la cumbre del sistema pactista catalán

    Directory of Open Access Journals (Sweden)

    Germán Segura García


    Full Text Available En este artículo se estudia el ordenamiento jurídico que supo darse Cataluña en vísperas de la destrucción del sistema pactista que le había vinculado con sus monarcas desde los tiempos medievales. Dejando de lado el análisis de las peculiaridades de dicho sistema, se procederá al examen de su mejor puesta en escena, con el espectro de la guerra civil en ciernes, durante las Cortes de Barcelona de 1705-1706. De esta última reunión de los estamentos catalanes con su monarca surgirían las constituciones de 1706, la ley pactada que quedó malograda por la derrota del bando austracista en la Guerra de Sucesión española.The article deals with the legal corpus that Catalonia was able to achieve for itself before the destruction of the pact system, which had tied the land with its monarchs from the Middle Ages. Leaving aside the analysis of the peculiarities of this system, we will proceed to the examination of its best staging -the spectre of civil war hanging over Spain- during the Courts of Barcelona 1705-1706. From this last assembly that put together the Catalan states and their king, sprang up the constitutions of 1706, the law ruined due to the defeat of the Austrian faction in the Spanish Succession war.

  7. Mark Twain in the Russian Pre-Revolutionary Periodical. Part 2

    Directory of Open Access Journals (Sweden)

    Ekaterina A. Stetsenko


    Full Text Available This article deals with the analysis of interpretation of the works by Mark Twain, famous American author, in the Russian pre-revolutionary periodical press (1872–1916. The objects of research are critical articles, essays, reviews, correspondences, introductions to publications of Twain’s short stories and novels, obituaries, and other materials printed in central and provincial magazines and newspapers. Perception of Twain in Russia was contingent on many factors including political and cultural situation in the country, state of social thought and literary criticism, newspaper and magazine conjuncture etc., always remaining polysemantic and conflicting. In different times, in the years of democratic rising or reaction critics looked for something in Twain’s works that corresponded to the spirit of their time and helped solve ideological and aesthetic problems. Twain had reputation of either a “pure humorist” or a great writer, philosopher, and moralist. Democrats, liberals, conservatives, feminists, adepts of realistic or naturalistic trends in art discussed Twain’s works that became a source of knowledge about the United States and inspired polemics about Russia’s further development. Twain was highly esteemed as the author of books for children and young people. Yet his works that criticized monarchism and imperialism were often ignored or abridged. The history of Twain’s interpretation in the Russian press serves as evidence of the fact that perception of foreign literature is a dynamic and bumpy process, repeating itself and moving backwards but also getting to deeper levels of meanings.

  8. Paradox of the drinking-straw model of the butterfly proboscis. (United States)

    Tsai, Chen-Chih; Monaenkova, Daria; Beard, Charles E; Adler, Peter H; Kornev, Konstantin G


    Fluid-feeding Lepidoptera use an elongated proboscis, conventionally modeled as a drinking straw, to feed from pools and films of liquid. Using the monarch butterfly, Danaus plexippus (Linnaeus), we show that the inherent structural features of the lepidopteran proboscis contradict the basic assumptions of the drinking-straw model. By experimentally characterizing permeability and flow in the proboscis, we show that tapering of the food canal in the drinking region increases resistance, significantly hindering the flow of fluid. The calculated pressure differential required for a suction pump to support flow along the entire proboscis is greater than 1 atm (~101 kPa) when the butterfly feeds from a pool of liquid. We suggest that behavioral strategies employed by butterflies and moths can resolve this paradoxical pressure anomaly. Butterflies can alter the taper, the interlegular spacing and the terminal opening of the food canal, thereby controlling fluid entry and flow, by splaying the galeal tips apart, sliding the galeae along one another, pulsing hemolymph into each galeal lumen, and pressing the proboscis against a substrate. Thus, although physical construction of the proboscis limits its mechanical capabilities, its functionality can be modified and enhanced by behavioral strategies. © 2014. Published by The Company of Biologists Ltd.

  9. Acerca de la contribución militar de la Junta General de la provincia de Guipúzcoa a la Guerra de Granada en 1484

    Directory of Open Access Journals (Sweden)

    García Fernández, Ernesto


    Full Text Available The Guipuzcoans participated with their militias and their armed men in the war against Granada waged by the Catholic Monarchs Isabella and Ferdinand, from their ascension to the throne of the Crown of Castile to the conquest of the Muslim kingdom of Granada in 1492. In this paper we look into how several requests were made from the Province of Guipuzcoa for several ships with their respective armed crews in order to control the different points of access to the Strait of Gibraltar, on the one hand, and, on the other hand, we examine the replies given by the representatives of the Province councils after the Board Meeting at Usarraga.

    Los guipuzcoanos participaron con sus milicias y hombres armados en la Guerra de Granada auspiciada por los Reyes Católicos, Isabel y Fernando, desde su ascenso al trono de la Corona de Castilla hasta la conquista del reino nazarí en 1492. En este artículo se estudian por una parte las demandas a la Provincia Guipúzcoa de varias embarcaciones con sus respectivas tripulaciones armadas para controlar los accesos a Granada a través del Estrecho de Gibraltar y por otra las diversas respuestas dadas por los procuradores generales de los concejos de la Provincia, reunidos en la Junta General de Usarraga.

  10. Excavaciones en el Tíber durante los siglos XVIII y XIX

    Directory of Open Access Journals (Sweden)

    García Sánchez, Jorge


    Full Text Available Since the Renaissance, the Tiber River has been the object of the most unusual archaeological quests, driven mainly by fantastic medieval lore. In the 18th and 19th centuries, the excavations on the river intensified and were epitomized by the scientific atmosphere of the Enlightenment, even tough they resulted in little archaeological worth. In 1819, even the Spanish monarch Fernando VII almost became a Shareholder in one of these archaeological companies. After the Italian unification, many finds were discovered in the works done along the Tiber's urban region, of wich many are now in the Museo Nazionale Romano.Desde el Renacimiento el Tiber ha sido objeto de las más peregrinas búsquedas arqueológicas, emprendidas habitualmente sobre la base de fantásticos relatos medievales. En los siglos XVIII y XIX las excavaciones en el río se intensificaron y se dotaron del cientificismo propio de la Ilustración, siendo no obstante el fruto de estos trabajos muy escaso; en 1819, incluso el propio monarca español Fernando VII estuvo a punto de convertirse en accionista de una de estas empresas arqueológicas. Tras la Unificación italiana, las obras de estructuración del curso urbano del Tiber propiciaron finalmente numerosos hallazgos, muchos de los cuales se conservan en el Museo Nazionale Romano.

  11. O direito de resistência na França renascentista

    Directory of Open Access Journals (Sweden)

    Alberto Ribeiro Gonçalves de Barros


    Full Text Available Com o agravamento dos conflitos religiosos no reino francês, principalmente a partir da segunda metade do século XVI, a questão do direito de resistência ao poder político voltou a ocupar um lugar de destaque no debate público. De um lado, autores que defendiam o direito dos súditos de resistir às ordens do monarca quando seus comandos fossem tirânicos, justificando, inclusive, o tiranicídio; de outro, autores que negavam esse direito e afirmavam o dever irrestrito de obediência à autoridade política legitimamente constituída. O objetivo deste texto é apresentar essa discussão e enfatizar alguns de seus aspectos que antecipam o debate moderno sobre o direito de resistência à autoridade política.In the half of the sixteenth century, with the increase of the religious conflicts in the French kingdom, the question of the right of political resistance retook an important place in public debate. Some authors defend the right of subjects to resist the commands of the monarch when they were tyrannical, justifying even the tyrannicide; others denied this right and affirmed the unrestricted duty of obedience to the political authority. The aim of this paper is to present this quarrel and to emphasize some of its aspects that anticipate the modern debate on the political resistance.

  12. Marshall funds the Camaldolese monastery

    Directory of Open Access Journals (Sweden)

    Jan Kracik


    Full Text Available Mikołaj Wolski (1553-1630, founder of the Camaldolese hermitage at Bielany near Cracow, was an exceptionally colourful and interesting personage. He travelled widely in his youth touring the most important European countries and getting familiar with the Renaissance culture. In spite of the fact that he supported a candidate of the House of Hapsburg to the Polish throne, he managed to get appreciation of king Sigismund III, who entrusted him with a position of the Royal Marshal to the Crown and made him his close advisor. Many a time he was sent as an envoy to the pope and monarchs, playing, at the same time, the role of a patron of artists, whom he won not only for the royal palace, but to beautify his own residence in Krzepica. The last 27 years of his life he devoted to the idea of bringing the Camaldolese Order to Cracow and building a hermitage for them on the Bielany Hill. The founder persoally supervised construction works and resided in one of the first monastery houses. As an expression of humbleness he wished to be buried after his death under the threshold of he church in a monk’s robe.

  13. French Presence in Brazil in the Nineteenth Century: analysis of the accounting archives of Casa Boris’s in the period from 1872 to 1887

    Directory of Open Access Journals (Sweden)

    José Paulo Cosenza


    Full Text Available Founded in Brazilian imperial period, Casa Boris had an important commercial role in the Northeast region of Brazil. Using a competitive business structure, it linked the Ceará Province (branch with France (headquarters, by importing manufactured goods and exporting raw materials. In this paper, we studied the Journal and Ledger of Casa Boris, particularly from 1872 to 1887. The study aimed to analyze the contents of the accounting records contained in these books so as to report facts and points of historical interest to the Brazilian Accounting. The main contribution of the research, in general, is associated with an attempt to understand the influence of political, economic, social and institutional settings on accounting practices and vice-versa. Our findings led to the conclusion that the bookkeeping of Casa Boris was transparent to reveal the quality of accounting control of commercial operations undertaken by the firm and also shows the coexistence of a incipient provincial market that concomitantly linked Brazil to Europe in the second period of Brazilian Monarchical Empire.

  14. From Eyewitness Narratives to Retellings and Literary Adaptations: The Russian Time of Troubles in Early Modern Europe

    Directory of Open Access Journals (Sweden)

    George Prokhorov


    Full Text Available The article focuses on the adaptation strategies used by Lope de Vega in his play El Gran Duque de Moscovia y emperador perseguido (1617. This tragedy, built on material acquired from travelogues, represents the first depiction of the Russian Time of Troubles in fiction. In it, one can follow Lope de Vega’s shift from preserving the factual details collected from different travel sources to creating his own Baroque story placed within a purely Catholic world, as opposed to reality. In doing this, Lope de Vega creates a fictional space filled with mystery and miracles, where Heavens can intervene and punish the guilty party, whereby restoring the original status quo. Key situations turn from illustrations of an alien world into much more general depictions, namely, that of a tyrant versus a legal monarch, and the will of a ruler versus the law. The shift into tyranny provides the story with a new narrative centre and, by following Lope de Vega’s emphasis on the “Muscovian story,” discloses its universal spirit.

  15. Long-range seasonal migration in insects: mechanisms, evolutionary drivers and ecological consequences. (United States)

    Chapman, Jason W; Reynolds, Don R; Wilson, Kenneth


    Myriad tiny insect species take to the air to engage in windborne migration, but entomology also has its 'charismatic megafauna' of butterflies, large moths, dragonflies and locusts. The spectacular migrations of large day-flying insects have long fascinated humankind, and since the advent of radar entomology much has been revealed about high-altitude night-time insect migrations. Over the last decade, there have been significant advances in insect migration research, which we review here. In particular, we highlight: (1) notable improvements in our understanding of lepidopteran navigation strategies, including the hitherto unsuspected capabilities of high-altitude migrants to select favourable winds and orientate adaptively, (2) progress in unravelling the neuronal mechanisms underlying sun compass orientation and in identifying the genetic complex underpinning key traits associated with migration behaviour and performance in the monarch butterfly, and (3) improvements in our knowledge of the multifaceted interactions between disease agents and insect migrants, in terms of direct effects on migration success and pathogen spread, and indirect effects on the evolution of migratory systems. We conclude by highlighting the progress that can be made through inter-phyla comparisons, and identify future research areas that will enhance our understanding of insect migration strategies within an eco-evolutionary perspective. © 2015 John Wiley & Sons Ltd/CNRS.

  16. Shah Shuja’s ‘Hidden History’ and its Implications for the Historiography of Afghanistan

    Directory of Open Access Journals (Sweden)

    Shah Mahmoud Hanifi


    Full Text Available The essay uses colonial archival materials from the Archives of the Punjab Province in Lahore to address the thirty-year period between the two reigns of the Durrani Afghan Monarch Shah Shuja (r. 1803-1809 and 1839-1842. Focusing on the 1809-1839 period, the first part of the essay deals with Mountstuart Elphinstone’s 1809 diplomatic mission and Shuja’s flight from Peshawar. The second part of the article considers the communication between Shah Shuja’s primary wife and colonial officials that culminated in Shuja’s receipt of housing and a monthly British pension in Ludhiana in 1816. The third part of the essay treats Shuja’s aborted attempt to recapture Kabul without British support in 1832-1833 and its consequences for him in Ludhiana. Shuja’s lack of Pashto credentials, his dependency on British capital, and his circular migration pattern are viewed as normative rather than exceptional for Afghan rulers, and as such this essay contributes to a revision of the traditional historiography of Afghanistan that views the country through the incompatible lenses of Pashtun ethnic domination of the Afghan state structure and Pashtun tribal resistance to Afghan state formation.1

  17. Matchmaker Exchange. (United States)

    Sobreira, Nara L M; Arachchi, Harindra; Buske, Orion J; Chong, Jessica X; Hutton, Ben; Foreman, Julia; Schiettecatte, François; Groza, Tudor; Jacobsen, Julius O B; Haendel, Melissa A; Boycott, Kym M; Hamosh, Ada; Rehm, Heidi L


    In well over half of the individuals with rare disease who undergo clinical or research next-generation sequencing, the responsible gene cannot be determined. Some reasons for this relatively low yield include unappreciated phenotypic heterogeneity; locus heterogeneity; somatic and germline mosaicism; variants of uncertain functional significance; technically inaccessible areas of the genome; incorrect mode of inheritance investigated; and inadequate communication between clinicians and basic scientists with knowledge of particular genes, proteins, or biological systems. To facilitate such communication and improve the search for patients or model organisms with similar phenotypes and variants in specific candidate genes, we have developed the Matchmaker Exchange (MME). MME was created to establish a federated network connecting databases of genomic and phenotypic data using a common application programming interface (API). To date, seven databases can exchange data using the API (GeneMatcher, PhenomeCentral, DECIPHER, MyGene2, matchbox, Australian Genomics Health Alliance Patient Archive, and Monarch Initiative; the latter included for model organism matching). This article guides usage of the MME for rare disease gene discovery. © 2017 by John Wiley & Sons, Inc. Copyright © 2017 John Wiley and Sons, Inc.

  18. Anthropogenic changes in sodium affect neural and muscle development in butterflies. (United States)

    Snell-Rood, Emilie C; Espeset, Anne; Boser, Christopher J; White, William A; Smykalski, Rhea


    The development of organisms is changing drastically because of anthropogenic changes in once-limited nutrients. Although the importance of changing macronutrients, such as nitrogen and phosphorus, is well-established, it is less clear how anthropogenic changes in micronutrients will affect organismal development, potentially changing dynamics of selection. We use butterflies as a study system to test whether changes in sodium availability due to road salt runoff have significant effects on the development of sodium-limited traits, such as neural and muscle tissue. We first document how road salt runoff can elevate sodium concentrations in the tissue of some plant groups by 1.5-30 times. Using monarch butterflies reared on roadside- and prairie-collected milkweed, we then show that road salt runoff can result in increased muscle mass (in males) and neural investment (in females). Finally, we use an artificial diet manipulation in cabbage white butterflies to show that variation in sodium chloride per se positively affects male flight muscle and female brain size. Variation in sodium not only has different effects depending on sex, but also can have opposing effects on the same tissue: across both species, males increase investment in flight muscle with increasing sodium, whereas females show the opposite pattern. Taken together, our results show that anthropogenic changes in sodium availability can affect the development of traits in roadside-feeding herbivores. This research suggests that changing micronutrient availability could alter selection on foraging behavior for some roadside-developing invertebrates.

  19. Karyotypes versus Genomes: The Nymphalid Butterflies Melitaea cinxia, Danaus plexippus, and D. chrysippus. (United States)

    Traut, Walther; Ahola, Virpi; Smith, David A S; Gordon, Ian J; Ffrench-Constant, Richard H


    The number of sequenced lepidopteran genomes is increasing rapidly. However, the corresponding assemblies rarely represent whole chromosomes and generally also lack the highly repetitive W sex chromosome. Knowledge of the karyotypes can facilitate genome assembly and further our understanding of sex chromosome evolution in Lepidoptera. Here, we describe the karyotypes of the Glanville fritillary Melitaea cinxia (n = 31), the monarch Danaus plexippus (n = 30), and the African queen D. chrysippus (2n = 60 or 59, depending on the source population). We show by FISH that the telomeres are of the (TTAGG)n type, as found in most insects. M. cinxia and D. plexippus have "conventional" W chromosomes which are heterochromatic in meiotic and somatic cells. In D. chrysippus, the W is inconspicuous. Neither telomeres nor W chromosomes are represented in the published genomes of M. cinxia and D. plexippus. Representation analysis in sequenced female and male D. chrysippus genomes detected an evolutionarily old autosome-Z chromosome fusion in Danaus. Conserved synteny of whole chromosomes, so called "macro synteny", in Lepidoptera permitted us to identify the chromosomes involved in this fusion. An additional and more recent sex chromosome fusion was found in D. chrysippus by karyotype analysis and classical genetics. In a hybrid population between 2 subspecies, D. c. chrysippus and D. c. dorippus, the W chromosome was fused to an autosome that carries a wing colour locus. Thus, cytogenetics and the present state of genome data complement one another to reveal the evolutionary history of the species. © 2017 S. Karger AG, Basel.

  20. El terremoto de Lisboa de 1755: su influencia en la extracción ganadera aPortugal desde la antigua provincia de Tuy (Galicia

    Directory of Open Access Journals (Sweden)

    María Pilar AMARÉ TAFALLA


    Full Text Available RESUMEN: Debido a los daños causados por el terremoto de Lisboa, el rey de España autorizó mediante una real orden la supresión de los derechos aduaneros con Portugal de diversos productos, entre ellos el ganado vacuno. Esta medida ocasionó en la antigua provincia de Tuy escasez de carne y de reses por lo que las autoridades provinciales se quejaron al rey solicitando que se derogara, lo que consiguieron como consecuencia de la presión ejercida por una densa trama de influencias, volviéndose a la situación anterior al terremoto.ABSTRACT: Because of the damage caused by the Lisbon earthquake of November 1st, 1755, the King of Spain approved a Royal Order to abolish the customs duties with Portugal on certain products, including cattle. This measure produced a shortage of beef in ancient Tuy province, on the southern Galician border with Portugal, causing local authorities to make a formal complaint to the monarch in order to restore the customs duties to their former situation before earthquake. They achieved this when the King repealed his Royal Order.

  1. Memories of Serbian victims under Bulgarian occupation in Leskovački glasnik (1915-1918

    Directory of Open Access Journals (Sweden)

    Antić Dejan D.


    Full Text Available The Serbian army was forced to retreat towards Metohija and Albania eventually due to the armed aggression and attacks from the north and east in October 1915 by the united forces of the German and Austro-Hungarian units against the Kingdom of Serbia. Together with the army, the monarch, executive and legislative branch, intellectuals and civilians also retreated, not wanting to fall in the hands of the incoming army formations of the Central Powers. Following the retreat of the government and army bodies, the territory of the Kingdom of Serbia found itself under the control of the enemy forces, which started the 3-year period of occupation. The subject of this paper are the texts written by the newspaper called Leskovački glasnik (Leskovac Messenger in the period between World War I and World War II, which dealt with the sufferings of the Serbian civilians in the Bulgarian occupied zone from 1915 to 1918. Special attention is paid to the Leskovački glasnik's reports about marking the anniversary of the Bulgarian terror and killings of Serbs, about opening and consecration of memorials, about the heroic deeds and the tragic destiny of the people in south Serbia.

  2. Swaziland: country profile. (United States)

    Carrington, L


    Although Swaziland had been independent from colonialism for 20 years, a powerful monarch, King Mswati II, continues to control the country's political, religious, and social system. Swaziland has a population of 676,000, half of whom are under 15 years of age. The infant mortality rate is 105/1000 live births and 25% of children die before they reach their 5th birthday. Life expectancy is 54 years. Tribal chiefs, representing the king, hold and distribute about half of the national land. Most of the fertile land remains in the hands of white settler farmers. The concentration of income in foreign companies and urban centers has exacerbated poverty in rural areas. Depreciation of rand-linked local currency has boosted export earnings, but it has also raised the price of food and medical imports. Swaziland's main exports are sugar, wood pulp, chemicals, and fruit, most of which go to the UK and South Africa. The major food crops are maize, beans, groundnuts, and sorghum. About half of the working population is engaged in small-scale subsistence farming, but food yields are declining. The major producers are foreign companies attracted by Swaziland's low taxes and cheap labor supply.

  3. Intuition beyond the law of the state

    Directory of Open Access Journals (Sweden)

    Stephen Connelly


    Full Text Available This article examines one aspect of the possible influence of Aristotle on Spinoza's thinking of state laws and their limitations.  In the Nicomachean Ethics, the Stagirite sets out a theory of the just city based on appropriate geometrical proportioning of justice, but then proposes the hypothesis of the most excellent man: someone so virtuous that they cannot be bound by the city's laws and so must be banished or elevated to monarch.  The article investigates how Spinoza's own conceptions of geometry and metaphysics inform his view of justice and laws in the city.  It indicates how, in continuing to posit the virtuous as someone both with a higher form of cognition of law, but who must nevertheless live in the city, Spinoza is likely to have been confronted with Aristotle's 'problem of excellence'.  The article examines Spinoza's initial and strikingly modern solution to the problem, but also indicates how Spinoza's own thinking on metaphysics and genetic geometry pushes him beyond this 'answer' in his later political work.

  4. Potential ash impact from Antarctic volcanoes: Insights from Deception Island's most recent eruption. (United States)

    Geyer, A; Marti, A; Giralt, S; Folch, A


    Ash emitted during explosive volcanic eruptions may disperse over vast areas of the globe posing a threat to human health and infrastructures and causing significant disruption to air traffic. In Antarctica, at least five volcanoes have reported historic activity. However, no attention has been paid to the potential socio-economic and environmental consequences of an ash-forming eruption occurring at high southern latitudes. This work shows how ash from Antarctic volcanoes may pose a higher threat than previously believed. As a case study, we evaluate the potential impacts of ash for a given eruption scenario from Deception Island, one of the most active volcanoes in Antarctica. Numerical simulations using the novel MMB-MONARCH-ASH model demonstrate that volcanic ash emitted from Antarctic volcanoes could potentially encircle the globe, leading to significant consequences for global aviation safety. Results obtained recall the need for performing proper hazard assessment on Antarctic volcanoes, and are crucial for understanding the patterns of ash distribution at high southern latitudes with strong implications for tephrostratigraphy, which is pivotal to synchronize palaeoclimatic records.

  5. Dereplication-guided isolation of a new indole alkaloid triglycoside from the hooks of Uncaria rhynchophylla by LC with ion trap time-of-flight MS. (United States)

    Zhang, Jian-Gang; Huang, Xiao-Yan; Ma, Yun-Bao; Zhang, Xue-Mei; Chen, Ji-Jun; Geng, Chang-An


    Uncaria rhynchophylla (Gou-Teng) as the monarch herb of many formulae (Fufang), e.g. "Tian-Ma-Gou-Teng-Yin," "Ling-Jiao-Gou-Teng-Yin," and "Yi-Gan-San", is a famous traditional Chinese medicine documented in the Chinese pharmacopoeia for mental and cardiovascular diseases. In the traditional Chinese medicine system, only the hook-bearing stems are used as the crude materials for Gou-Teng, and the hooks are always considered more effective than the stems. Focusing on the mono-herb and its active constituents from combinatorial formulae is the core idea of reductionism of traditional Chinese medicine theory. Detailed liquid chromatography with mass spectrometry analysis on the hooks of U. rhynchophylla was performed to profile the chemical constituents based on tandem mass spectrometry fragmentation and UV absorption. Under the guidance of liquid chromatography with ion trap/time-of-flight mass spectrometry, one new indole alkaloid triglycoside (1), together with five known compounds 2-6 as the main constituents, were isolated from the hooks of U. rhynchophylla by various column chromatography methods. Compound 1 showed moderate activity on MT 1 and MT 2 melatonin receptors with agonistic rates of 79.6 and 46.3% at the concentration of 1 mM. This dereplication strategy can be equally applicable to rapidly disclose the active constituents of other Chinese herbs through targeted purification. © 2017 WILEY-VCH Verlag GmbH & Co. KGaA, Weinheim.

  6. What Drives The Changes In The Gulf Oil Monarchies?

    Directory of Open Access Journals (Sweden)

    N. Kjurchiski


    Full Text Available The article identifies four main events driving the current changes in thePersian Gulfoil monarchies. Located in a troubled and unstable region of the Middle East with the oil prices declining after the events of the “Arab spring” and election of Donald Trump as president of theUnited States, Gulf oil monarchies are under pressure from inside and outside to undergo significant changes. As a consequence, validity of the theory of «monarchical exclusiveness» is called into question. The political elites in the monarchies recognize the need to depart from the status quo, although the phenomenon of the «king’s dilemma» is still present there. The author argues that the monarchy should undergo conservative liberalization. To avoid political destabilization they should apply a gradual, evolutionary approach to liberalization providing their societies with enough time to adapt to new developments. Along with the gradual liberalisation process there is also need in institutional development to make institutions stronger and more independent.

  7. Empress Elisabeth (‘Sisi’ of Austria and Patriotic Fashionism

    Directory of Open Access Journals (Sweden)

    Christopher M. VanDemark


    Full Text Available In this article, Christopher VanDemark explores the intersections between nationalism, fashion, and the royal figure in Hungary between 1857 and the Compromise of 1867. Focusing on aesthetics as a vehicle for feminine power at a critical junction in Hungarian history, VanDemark contextualizes Empress Elisabeth’s role in engendering a revised political schema in the Habsburg sphere. Foreseeing the power of emblematic politics, the young Empress adeptly situated herself between the Hungarians and the Austrians to recast the Hungarian martyrology narrative promulgated after the failed revolution of 1848. Eminent Hungarian newspapers such as the Pesti Napló, Pester Lloyd, and the Vasárnapi Újság form the backbone of this article, as publications such as these facilitated the dissemination of patriotic sentiment while simultaneously exulting the efficacy of symbolic fashions. The topic of study engages with contemporary works on nationalism, which emphasize gender and aesthetics, and contributes to the emerging body of scholarship on important women in Hungarian history. Seminal texts by Catherine Brice, Sara Maza, Abby Zanger, and Lynn Hunt compliment the wider objective of this brief analysis, namely, the notion that the Queen’s body can both enhance and reform monarchical power within a nineteenth-century milieu.

  8. Gambling in revolutionary Paris - The Palais Royal: 1789-1838. (United States)

    Barnhart, R T


    By the revolution of 1789, the four-story, quadrangular Palais Royal in Paris had become the most glittering tourist center of Europe, with 180 shops and cafes in its ground floor arcades. By 1791, its basement and secondary story contained over 100 separate, illicit gambling operations featuring the most popular dice and card games. The mania for gambling had been transferred from defunct, monarchical Versailles to the thriving, bourgeois Palais Royal, where the five main gaming clubs throbbed from noon till midnight. During the Revolution, Prince Talleyrand won 30,000 francs at one club, and after Waterloo in 1815, Marshal Blucher lost 1,500,000 francs in one night at another. To bring the situation under control and raise taxes for the state, in 1806 Napoleon legalized the main clubs, which from 1819 to 1837 grossed an enormous 137 million francs. When the anti-gambling forces triumphed in 1837 and the clubs were closed down, the National Guard had to be called out to evict the mobs of gamblers who refused to leave the tables. Dramatic reports from Revolutionary police raids, and quotations from the memoirs of humorous French gamblers and shocked foreign visitors, provide anecdotal illustrations of the 49 years during which the Palais Royal was the most intriguing and picturesque gambling mecca of Europe-and probably of the world.

  9. Nuremberg Counting Jetons of XVI–XIX centuries: from Western Europe to Western Siberia

    Directory of Open Access Journals (Sweden)

    Andrey A. Pushkarev


    Full Text Available The article is devoted to the history of Western counting jetons, which have spread over a vast area from Western Europe to Siberia. The history of jetons is very dynamic. For more than three centuries in Western Europe tokens were used as a tool for calculation in trading shops, government offices, etc. In the second half of the XVI century because the spread of the written account the functions of jetons fundamentally changed. They are being used as chips in card games, they were presented as souvenirs, scattered among the crowd at weddings and festivals, etc. At the same time, jetons became the translation tool for public information about the government, in the form of images of the reigning monarch, as well as key political events. Through trade exchange jetons penetrate into the territory of Western Siberia, where their function changed again. All counting tokens from burial graves in Western Siberia, have holes for hanging or sewing, which indicate their use as ornaments. However, in the culture of the indigenous population decorations had not only aesthetic but also a sacred, religious meaning.

  10. The calm before the storm: carlist and neocatholics in the Valencian country between 1849 and 1868 | La calma antes de la tempestad: carlistas y neocatólicos en el País Valenciano entre 1849 y 1868

    Directory of Open Access Journals (Sweden)

    Antonio Caridad Salvador


    Full Text Available Between 1849 and 1868 the Valencian carlism lived a dark age, in which its activity was very small, due to the failures of its previous revolts and the support to moderantism of many former carlists. At the same time, it emerged an absolutist group which accepted Elizabeth II as queen and that took part in the parliamentary activity. They were the pure monarchics (or neocatholics, that defended a return to a more traditional régime, without demanding a change in the monarchy. Starting on 1865, with the recognition of the Italian State, it began the convergence between both political tendencies. | Entre 1849 y 1868 el carlismo valenciano vivió una época oscura, en la que su actividad fue muy reducida, dados los fracasos de sus anteriores revueltas y el apoyo al moderantismo de muchos antiguos carlistas. Al mismo tiempo tanto surgía un grupo absolutista que aceptaba como reina a Isabel II y que llegó a entrar en el juego parlamentario. Se trataba de los monárquicos puros (o neocatólicos, que defendían una vuelta a un régimen más tradicional pero sin buscar un cambio de monarca. A partir de 1865, con el reconocimiento del Estado italiano, empezará la convergencia entre estas dos corrientes.

  11. Sir Astley Paston Cooper (1768-1841): The man and his personality. (United States)

    Doganay, Emre


    The most acclaimed surgeon of his time, Astley Cooper, a man from Norfolk and a student of the eminent John Hunter, was an outstandingly successful surgeon. With his accomplishments in surgery and experiments in dissection he soon became a prominent figure and received recognition worldwide. At the young age of 21 he was appointed Demonstrator in Anatomy at St Thomas's Hospital in London and two years later was made Joint Lecturer in Anatomy and Surgery. With his passion for anatomy, his love for surgery and his expanding knowledge he became Surgeon to Guy's Hospital in 1800 and in the same year was elected a Fellow to the Royal Society. His attainments led him to become surgeon to three successive British monarchs as well as receiving a bestowal of Baronetcy. Through his edifying lectures, fastidious experiments and publications on anatomy and pathology he has inspired and enlightened many budding anatomists and surgeons and the principles of his teachings still prevail in practice today. © The Author(s) 2014.

  12. Brasileiro tradutor e/ou traidor: Frei José Mariano da Conceição Veloso.

    Directory of Open Access Journals (Sweden)

    Alessandra Ramos de Oliveira Harden


    Full Text Available This article is concerned with a specific phenomenon of the history of translation into Portuguese, the boom of translation of scientific and didactic texts in the turn of the 18th century in Portugal, in which a considerable number of Brazilians were involved. We focus the work of Friar José Mariano da Conceição Veloso, who has lately been referred to as a key figure in the history of sciences and books publishing in Portugal and Brazil. Friar Veloso was translator, editor, and translation coordinator in charge of the work of translators concerned with pragmatic texts in Lisbon. Veloso’s involvement with the still incipient editorial industry in Portugal was linked to Portuguese Enlightenment, especially the dissemination of scientific knowledge that could be applied to the progress of the Portuguese kingdom. Clues of Friar Veloso’s close connection with the Portuguese official policies are found in the prefaces he wrote for his translations, and they are enough to include him withim a patronage system that defined the direction his editorial activities would take. Curiously, despite these ties to the authoritarian Portuguese monarchical regime, Friar Veloso’s work led to unexpected historical developments, such as the growth of the reading public and the improvement of printing techniques.

  13. How the Warriors of Prester John Transformed into Demons from Tartarus. Review of the book: Hautala R. From “David, King of the Indies” to “Detestable Plebs of Satan”: An Anthology of Early Latin Information about the Tatar-Mongols. (Kazan: Sh.Marjani Institute of History of AS RT, 2015. 496 p.

    Directory of Open Access Journals (Sweden)

    R.Yu. Pochekaev


    Full Text Available This paper is a review of recently published book of Finnish researcher Roman Hautala devoted to publication and study of Latin sources on the Mongols of the 13th century. The book is of great importance as it is a compilations of the very valuable sources which demonstrate evolution of the attitude of European authorities and contemporaries towards the Mongols – from potential allies to the most dangerous enemy. Analysis of these sources helps to understand the further policy of Western European monarchs in the Eastern Europe, Ancient Rus’, Mongol Empire and its successors – Golden Horde, Ilkhanate, etc. At the same time, R. Hautala pays substantial attention to a fate of another nomads of Eurasia connected with the Mongol invasion – especially Kumans (Kipchaks in Hungary. The book contains author’s preface, Latin texts and Russian translations of sources as well as very valuable comments of author. Each part could be considered as independent part of this complex research. The main importance of book is put into use of Russian-speaking scientific society of a huge number of sources, most part of which was not accessible for home historians because of lack of knowledge of Latin. The structure of the book completely corresponds to its goals and objectives. Nevertheless, the author of review points that the book has several insignificant weaknesses, which could be kept in mind and removed in the future works of R. Hautala.

  14. Identification and Expression Profiling of the BTB Domain-Containing Protein Gene Family in the Silkworm, Bombyx mori

    Directory of Open Access Journals (Sweden)

    Daojun Cheng


    Full Text Available The BTB domain is a conserved protein-protein interaction motif. In this study, we identified 56 BTB domain-containing protein genes in the silkworm, in addition to 46 in the honey bee, 55 in the red flour beetle, and 53 in the monarch butterfly. Silkworm BTB protein genes were classified into nine subfamilies according to their domain architecture, and most of them could be mapped on the different chromosomes. Phylogenetic analysis suggests that silkworm BTB protein genes may have undergone a duplication event in three subfamilies: BTB-BACK-Kelch, BTB-BACK-PHR, and BTB-FLYWCH. Comparative analysis demonstrated that the orthologs of each of 13 BTB protein genes present a rigorous orthologous relationship in the silkworm and other surveyed insects, indicating conserved functions of these genes during insect evolution. Furthermore, several silkworm BTB protein genes exhibited sex-specific expression in larval tissues or at different stages during metamorphosis. These findings not only contribute to a better understanding of the evolution of insect BTB protein gene families but also provide a basis for further investigation of the functions of BTB protein genes in the silkworm.

  15. Military organization and army command of ancient armies of Northern Black Sea Region

    Directory of Open Access Journals (Sweden)

    K. M. Kolesnykov


    Full Text Available This article deal with the description, analysis and generalization of features organization of command structure of the army ancient states of the Northern Black Sea region: Olbia, Chersonese, Bosporus. Author sure that the foundation of the armed forces of these states was a civil militia – free, economically independent landowners, endowed with the broadest volume of political and social rights men – full citizens. Accordingly, the assemblies of citizens were electing strategists, who led militia forces. In case of need the people also claimed commanders’ mercenary troops. Bosporus kings have attracted a significant number of hired troops that held by the royal treasury income and special direct tax. Bosporus military command came from magistrates of polis only at the lower levels. Middle and higher level of «officer corps» of the Bosporus were appointed by the monarch from among the landowners, officialdom and military aristocracy. For example, in the Roman period on the Bosporus fixed formation units of cathafractarian cavalry. The weapons and tactics of the Sarmatian nomadic nobility (Aspurhianian, Sirak, Aorsy and more were served as a model for Bosporus cataphractarian horsemen.

  16. De conjunto de rentas a impuesto aduanero. La transformación del almojarifazgo durante el siglo XIV en el reino de Murcia

    Directory of Open Access Journals (Sweden)

    González Arce, José Damián


    Full Text Available Before becoming a customs duty, the almojarifazgo consisted of a set of heterogeneous income received by the Castilian monarchs. Originally, most of these rents were demanded by the Muslim rulers of the cities then conquered by the Christian kings. However, during the thirteenth and fourteenth centuries the kings were giving them to the lords and local councils and kept for themselves the most interesting of all, the customs duty. The kingdom of Murcia is almost the only example to study how this transformation because it is the only territory that has retained suffi cient documentation to analyse it.

    Antes de convertirse en un impuesto aduanero, el almojarifazgo consistió en un conjunto de rentas heterogéneas percibidas por los monarcas castellanos. En origen, la mayor parte de esas rentas fueron demandadas por los gobernantes musulmanes de las ciudades luego conquistadas por los reyes cristianos. Sin embargo, durante los siglos XIII y XIV dichos reyes las fueron cediendo a los señores y concejos locales y se quedaron con la más interesante de todas, el arancel aduanero. El reino de Murcia constituye casi el único ejemplo para estudiar cómo se operó esta transformación, porque es el único territorio que ha conservado la sufi ciente documentación para poder analizar tal transformación.

  17. Ecosystem services from transborder migratory species: Implications for conservation governance (United States)

    Lopez-Hoffman, Laura; Chester, Charles C.; Semmens, Darius J.; Thogmartin, Wayne E.; Rodriguez-McGoffin, M. Sofia; Merideth, Robert; Diffendorfer, Jay E.


    This article discusses the conservation challenges of volant migratory transborder species and conservation governance primarily in North America. Many migratory species provide ecosystem service benefits to society. For example, insectivorous bats prey on crop pests and reduce the need for pesticides; birds and insects pollinate food plants; and birds afford recreational opportunities to hunters and birdwatchers. Migration is driven by the seasonal availability of resources; as resources in one area become seasonally scarce, individuals move to locations where resources have become seasonally abundant. The separation of the annual lifecycle means that species management and governance is often fractured across international borders. Because migratory species depend on habitat in different locations, their ability to provide ecosystem services in one area depends on the spatial subsidies, or support, provided by habitat and ecological processes in other areas. This creates telecouplings, or interconnections across geographic space, of areas such that impacts to the habitat of a migratory species in one location will affect the benefits enjoyed by people in other locations. Information about telecoupling and spatial subsidies can be used to craft new governance arrangements such as Payment for Ecosystem Services programs that target specific stakeholder groups and locations. We illustrate these challenges and opportunities with three North American case studies: the Duck Stamp Program, Mexican free-tailed bats (Tadarida brasiliensis mexicana), and monarch butterflies (Danaus plexippus).

  18. Effective stress, friction and deep crustal faulting (United States)

    Beeler, N.M.; Hirth, Greg; Thomas, Amanda M.; Burgmann, Roland


    Studies of crustal faulting and rock friction invariably assume the effective normal stress that determines fault shear resistance during frictional sliding is the applied normal stress minus the pore pressure. Here we propose an expression for the effective stress coefficient αf at temperatures and stresses near the brittle-ductile transition (BDT) that depends on the percentage of solid-solid contact area across the fault. αf varies with depth and is only near 1 when the yield strength of asperity contacts greatly exceeds the applied normal stress. For a vertical strike-slip quartz fault zone at hydrostatic pore pressure and assuming 1 mm and 1 km shear zone widths for friction and ductile shear, respectively, the BDT is at ~13 km. αf near 1 is restricted to depths where the shear zone is narrow. Below the BDT αf = 0 is due to a dramatically decreased strain rate. Under these circumstances friction cannot be reactivated below the BDT by increasing the pore pressure alone and requires localization. If pore pressure increases and the fault localizes back to 1 mm, then brittle behavior can occur to a depth of around 35 km. The interdependencies among effective stress, contact-scale strain rate, and pore pressure allow estimates of the conditions necessary for deep low-frequency seismicity seen on the San Andreas near Parkfield and in some subduction zones. Among the implications are that shear in the region separating shallow earthquakes and deep low-frequency seismicity is distributed and that the deeper zone involves both elevated pore fluid pressure and localization.

  19. A 30-year history of earthquake crisis communication in California and lessons for the future (United States)

    Jones, L.


    The first statement from the US Geological Survey to the California Office of Emergency Services quantifying the probability of a possible future earthquake was made in October 1985 about the probability (approximately 5%) that a M4.7 earthquake located directly beneath the Coronado Bay Bridge in San Diego would be a foreshock to a larger earthquake. In the next 30 years, publication of aftershock advisories have become routine and formal statements about the probability of a larger event have been developed in collaboration with the California Earthquake Prediction Evaluation Council (CEPEC) and sent to CalOES more than a dozen times. Most of these were subsequently released to the public. These communications have spanned a variety of approaches, with and without quantification of the probabilities, and using different ways to express the spatial extent and the magnitude distribution of possible future events. The USGS is re-examining its approach to aftershock probability statements and to operational earthquake forecasting with the goal of creating pre-vetted automated statements that can be released quickly after significant earthquakes. All of the previous formal advisories were written during the earthquake crisis. The time to create and release a statement became shorter with experience from the first public advisory (to the 1988 Lake Elsman earthquake) that was released 18 hours after the triggering event, but was never completed in less than 2 hours. As was done for the Parkfield experiment, the process will be reviewed by CEPEC and NEPEC (National Earthquake Prediction Evaluation Council) so the statements can be sent to the public automatically. This talk will review the advisories, the variations in wording and the public response and compare this with social science research about successful crisis communication, to create recommendations for future advisories

  20. Low strength of deep San Andreas fault gouge from SAFOD core. (United States)

    Lockner, David A; Morrow, Carolyn; Moore, Diane; Hickman, Stephen


    The San Andreas fault accommodates 28-34 mm yr(-1) of right lateral motion of the Pacific crustal plate northwestward past the North American plate. In California, the fault is composed of two distinct locked segments that have produced great earthquakes in historical times, separated by a 150-km-long creeping zone. The San Andreas Fault Observatory at Depth (SAFOD) is a scientific borehole located northwest of Parkfield, California, near the southern end of the creeping zone. Core was recovered from across the actively deforming San Andreas fault at a vertical depth of 2.7 km (ref. 1). Here we report laboratory strength measurements of these fault core materials at in situ conditions, demonstrating that at this locality and this depth the San Andreas fault is profoundly weak (coefficient of friction, 0.15) owing to the presence of the smectite clay mineral saponite, which is one of the weakest phyllosilicates known. This Mg-rich clay is the low-temperature product of metasomatic reactions between the quartzofeldspathic wall rocks and serpentinite blocks in the fault. These findings provide strong evidence that deformation of the mechanically unusual creeping portions of the San Andreas fault system is controlled by the presence of weak minerals rather than by high fluid pressure or other proposed mechanisms. The combination of these measurements of fault core strength with borehole observations yields a self-consistent picture of the stress state of the San Andreas fault at the SAFOD site, in which the fault is intrinsically weak in an otherwise strong crust. ©2011 Macmillan Publishers Limited. All rights reserved

  1. Scientific drilling into the San Andreas Fault Zone - an overview of SAFOD's first five years (United States)

    Zoback, Mark; Hickman, Stephen; Ellsworth, William; ,


    The San Andreas Fault Observatory at Depth (SAFOD) was drilled to study the physical and chemical processes controlling faulting and earthquake generation along an active, plate-bounding fault at depth. SAFOD is located near Parkfield, California and penetrates a section of the fault that is moving due to a combination of repeating microearthquakes and fault creep. Geophysical logs define the San Andreas Fault Zone to be relatively broad (~200 m), containing several discrete zones only 2–3 m wide that exhibit very low P- and S-wave velocities and low resistivity. Two of these zones have progressively deformed the cemented casing at measured depths of 3192 m and 3302 m. Cores from both deforming zones contain a pervasively sheared, cohesionless, foliated fault gouge that coincides with casing deformation and explains the observed extremely low seismic velocities and resistivity. These cores are being now extensively tested in laboratories around the world, and their composition, deformation mechanisms, physical properties, and rheological behavior are studied. Downhole measurements show that within 200 m (maximum) of the active fault trace, the direction of maximum horizontal stress remains at a high angle to the San Andreas Fault, consistent with other measurements. The results from the SAFOD Main Hole, together with the stress state determined in the Pilot Hole, are consistent with a strong crust/weak fault model of the San Andreas. Seismic instrumentation has been deployed to study physics of faulting—earthquake nucleation, propagation, and arrest—in order to test how laboratory-derived concepts scale up to earthquakes occurring in nature.

  2. Relaxation of the south flank after the 7.2-magnitude Kalapana earthquake, Kilauea Volcano, Hawaii (United States)

    Dvorak, John J.; Klein, Fred W.; Swanson, Donald A.


    An M = 7.2 earthquake on 29 November 1975 caused the south flank of Kilauea Volcano, Hawaii, to move seaward several meters: a catastrophic release of compression of the south flank caused by earlier injections of magma into the adjacent segment of a rift zone. The focal mechanisms of the mainshock, the largest foreshock, and the largest aftershock suggest seaward movement of the upper block. The rate of aftershocks decreased in a familiar hyperbolic decay, reaching the pre-1975 rate of seismicity by the mid-1980s. Repeated rift-zone intrusions and eruptions after 1975, which occurred within 25 km of the summit area, compressed the adjacent portion of the south flank, apparently masking continued seaward displacement of the south flank. This is evident along a trilateration line that continued to extend, suggesting seaward displacement, immediately after the M = 7.2 earthquake, but then was compressed during a series of intrusions and eruptions that began in September 1977. Farther to the east, trilateration measurements show that the portion of the south flank above the aftershock zone, but beyond the area of compression caused by the rift-zone intrusions and eruptions, continued to move seaward at a decreasing rate until the mid-1980s, mimicking the decay in aftershock rate. Along the same portion of the south flank, the pattern of vertical surface displacements can be explained by continued seaward movement of the south flank and development of two eruptive fissures along the east rift zone, each of which extended from a depth of ∼3 km to the surface. The aftershock rate and continued seaward movement of the south flank are reminiscent of crustal response to other large earthquakes, such as the 1966 M = 6 Parkfield earthquake and the 1983 M = 6.5 Coalinga earthquake.

  3. Imbricated slip rate processes during slow slip transients imaged by low-frequency earthquakes (United States)

    Lengliné, O.; Frank, W.; Marsan, D.; Ampuero, J. P.


    Low Frequency Earthquakes (LFEs) often occur in conjunction with transient strain episodes, or Slow Slip Events (SSEs), in subduction zones. Their focal mechanism and location consistent with shear failure on the plate interface argue for a model where LFEs are discrete dynamic ruptures in an otherwise slowly slipping interface. SSEs are mostly observed by surface geodetic instruments with limited resolution and it is likely that only the largest ones are detected. The time synchronization of LFEs and SSEs suggests that we could use the recorded LFEs to constrain the evolution of SSEs, and notably of the geodetically-undetected small ones. However, inferring slow slip rate from the temporal evolution of LFE activity is complicated by the strong temporal clustering of LFEs. Here we apply dedicated statistical tools to retrieve the temporal evolution of SSE slip rates from the time history of LFE occurrences in two subduction zones, Mexico and Cascadia, and in the deep portion of the San Andreas fault at Parkfield. We find temporal characteristics of LFEs that are similar across these three different regions. The longer term episodic slip transients present in these datasets show a slip rate decay with time after the passage of the SSE front possibly as t-1/4. They are composed of multiple short term transients with steeper slip rate decay as t-α with α between 1.4 and 2. We also find that the maximum slip rate of SSEs has a continuous distribution. Our results indicate that creeping faults host intermittent deformation at various scales resulting from the imbricated occurrence of numerous slow slip events of various amplitudes.

  4. The effect of lateral variations of friction on crustal faulting

    Directory of Open Access Journals (Sweden)

    M. Cocco


    Full Text Available We propose that lateral variations in fault friction control the heterogeneity of slip observed in large earthquakes, We model these variations using a rate and state-dependent friction law, where we differentiate velocity-weakening into strong and weak-seismic fields, and velocity-strengthening into compliant and viscous fields. The strong-seismic field comprises the seismic slip concentrations, or asperities. The two «intermediate» frictional fields, weak-seismic and compliant, modulate both the tectonic loading and the dynamic rupture process. During the interseismic period, the compliant and viscous regions slip aseismically while the strong-seismic regions remain locked, evolving into stress concentrations that fail only in main shocks. The weak-seismic regions contain most of the interseismic activity and aftershocks, but also «creep seismically», that is, most of the weak-seismic area slips aseismically, actuating the seismicity on the remaining area. This «mixed» frictional behavior can be obtained from a sufficiently heterogenous distribution for the critical slip distance. The interseismic slip provides an inherent rupture resistance: dynamic rupture fronts decelerate as they penetrate into these unloaded compliant or creeping weak-seismic areas, diffusing into broad areas of accelerated afterslip. Aftershocks occur in both the weak-seismic and compliant areas around the fault, but most of the stress is diffused through aseismic slip. Rapid afterslip on these peripheral areas can also produce aftershocks within the main shock rupture area, by reloading weak fault areas that slipped in the main shock and then healed. We test this frictional model by comparing the interevent seismicity and aftershocks to the coseismic slip distribution for the 1966 Parkfield, 1979 Coyote Lake, and 1984 Morgan Hill earthquakes.

  5. Modular Approaches to Earth Science Scientific Computing: 3D Electromagnetic Induction Modeling as an Example (United States)

    Tandon, K.; Egbert, G.; Siripunvaraporn, W.


    We are developing a modular system for three-dimensional inversion of electromagnetic (EM) induction data, using an object oriented programming approach. This approach allows us to modify the individual components of the inversion scheme proposed, and also reuse the components for variety of problems in earth science computing howsoever diverse they might be. In particular, the modularity allows us to (a) change modeling codes independently of inversion algorithm details; (b) experiment with new inversion algorithms; and (c) modify the way prior information is imposed in the inversion to test competing hypothesis and techniques required to solve an earth science problem. Our initial code development is for EM induction equations on a staggered grid, using iterative solution techniques in 3D. An example illustrated here is an experiment with the sensitivity of 3D magnetotelluric inversion to uncertainties in the boundary conditions required for regional induction problems. These boundary conditions should reflect the large-scale geoelectric structure of the study area, which is usually poorly constrained. In general for inversion of MT data, one fixes boundary conditions at the edge of the model domain, and adjusts the earth?s conductivity structure within the modeling domain. Allowing for errors in specification of the open boundary values is simple in principle, but no existing inversion codes that we are aware of have this feature. Adding a feature such as this is straightforward within the context of the modular approach. More generally, a modular approach provides an efficient methodology for setting up earth science computing problems to test various ideas. As a concrete illustration relevant to EM induction problems, we investigate the sensitivity of MT data near San Andreas Fault at Parkfield (California) to uncertainties in the regional geoelectric structure.

  6. How informative are slip models for aftershock forecasting? (United States)

    Bach, Christoph; Hainzl, Sebastian


    Coulomb stress changes (ΔCFS) have been recognized as a major trigger mechanism for earthquakes, in particular aftershock distributions and the spatial patterns of ΔCFS are often found to be correlated. However, the Coulomb stress calculations are based on slip inversions and the receiver fault mechanisms which both contain large uncertainties. In particular, slip inversions are usually non-unique and often differ strongly for the same earthquakes. Here we want to address the information content of those inversions with respect to aftershock forecasting. Therefore we compare the slip models to randomized fractal slip models which are only constrained by fault information and moment magnitude. The uncertainty of the aftershock mechanisms is considered by using many receiver fault orientations, and by calculating ΔCFS at several depth layers. The stress change is then converted into an aftershock probability map utilizing a clock advance model. To estimate the information content of the slip models, we use an Epidemic Type Aftershock Sequence (ETAS) model approach introduced by Bach and Hainzl (2012), where the spatial probability density of direct aftershocks is related to the ΔCFS calculations. Besides the directly triggered aftershocks, this approach also takes secondary aftershock triggering into account. We quantify our results by calculating the information gain of the randomized slip models relative to the corresponding published slip model. As case studies, we investigate the aftershock sequences of several well-known main shocks such as 1992 Landers, 1999 Hector Mine, 2004 Parkfield, 2002 Denali. First results show a huge difference in the information content of slip models. For some of the cases up to 90% of the random slip models are found to perform better than the originally published model, for some other cases only few random models are found performing better than the published slip model.

  7. Shallow soil CO2 flow along the San Andreas and Calaveras Faults, California (United States)

    Lewicki, J.L.; Evans, William C.; Hilley, G.E.; Sorey, M.L.; Rogie, J.D.; Brantley, S.L.


    We evaluate a comprehensive soil CO2 survey along the San Andreas fault (SAF) in Parkfield, and the Calaveras fault (CF) in Hollister, California, in the context of spatial and temporal variability, origin, and transport of CO2 in fractured terrain. CO2 efflux was measured within grids with portable instrumentation and continously with meteorological parameters at a fixed station, in both faulted and unfaulted areas. Spatial and temporal variability of surface CO2 effluxes was observed to be higher at faulted SAF and CF sites, relative to comparable background areas. However, ??13C (-23.3 to - 16.4???) and ??14C (75.5 to 94.4???) values of soil CO2 in both faulted and unfaulted areas are indicative of biogenic CO2, even though CO2 effluxes in faulted areas reached values as high as 428 g m-2 d-1. Profiles of soil CO2 concentration as a function of depth were measured at multiple sites within SAF and CF grids and repeatedly at two locations at the SAF grid. Many of these profiles suggest a surprisingly high component of advective CO2 flow. Spectral and correlation analysis of SAF CO2 efflux and meteorological parameter time series indicates that effects of wind speed variations on atmospheric air flow though fractures modulate surface efflux of biogenic CO2. The resulting areal patterns in CO2 effluxes could be erroneously attributed to a deep gas source in the absence of isotopic data, a problem that must be addressed in fault zone soil gas studies.

  8. Low resistivity and permeability in actively deforming shear zones on the San Andreas Fault at SAFOD (United States)

    Morrow, Carolyn A.; Lockner, David A.; Hickman, Stephen H.


    The San Andreas Fault Observatory at Depth (SAFOD) scientific drillhole near Parkfield, California crosses the San Andreas Fault at a depth of 2.7 km. Downhole measurements and analysis of core retrieved from Phase 3 drilling reveal two narrow, actively deforming zones of smectite-clay gouge within a roughly 200 m-wide fault damage zone of sandstones, siltstones and mudstones. Here we report electrical resistivity and permeability measurements on core samples from all of these structural units at effective confining pressures up to 120 MPa. Electrical resistivity (~10 ohm-m) and permeability (10-21 to 10-22 m2) in the actively deforming zones were one to two orders of magnitude lower than the surrounding damage zone material, consistent with broader-scale observations from the downhole resistivity and seismic velocity logs. The higher porosity of the clay gouge, 2 to 8 times greater than that in the damage zone rocks, along with surface conduction were the principal factors contributing to the observed low resistivities. The high percentage of fine-grained clay in the deforming zones also greatly reduced permeability to values low enough to create a barrier to fluid flow across the fault. Together, resistivity and permeability data can be used to assess the hydrogeologic characteristics of the fault, key to understanding fault structure and strength. The low resistivities and strength measurements of the SAFOD core are consistent with observations of low resistivity clays that are often found in the principal slip zones of other active faults making resistivity logs a valuable tool for identifying these zones.

  9. Deep permeability of the San Andreas Fault from San Andreas Fault Observatory at Depth (SAFOD) core samples (United States)

    Morrow, Carolyn A.; Lockner, David A.; Moore, Diane E.; Hickman, Stephen H.


    The San Andreas Fault Observatory at Depth (SAFOD) scientific borehole near Parkfield, California crosses two actively creeping shear zones at a depth of 2.7 km. Core samples retrieved from these active strands consist of a foliated, Mg-clay-rich gouge containing porphyroclasts of serpentinite and sedimentary rock. The adjacent damage zone and country rocks are comprised of variably deformed, fine-grained sandstones, siltstones, and mudstones. We conducted laboratory tests to measure the permeability of representative samples from each structural unit at effective confining pressures, Pe up to the maximum estimated in situ Pe of 120 MPa. Permeability values of intact samples adjacent to the creeping strands ranged from 10−18 to 10−21 m2 at Pe = 10 MPa and decreased with applied confining pressure to 10−20–10−22 m2 at 120 MPa. Values for intact foliated gouge samples (10−21–6 × 10−23 m2 over the same pressure range) were distinctly lower than those for the surrounding rocks due to their fine-grained, clay-rich character. Permeability of both intact and crushed-and-sieved foliated gouge measured during shearing at Pe ≥ 70 MPa ranged from 2 to 4 × 10−22 m2 in the direction perpendicular to shearing and was largely insensitive to shear displacement out to a maximum displacement of 10 mm. The weak, actively-deforming foliated gouge zones have ultra-low permeability, making the active strands of the San Andreas Fault effective barriers to cross-fault fluid flow. The low matrix permeability of the San Andreas Fault creeping zones and adjacent rock combined with observations of abundant fractures in the core over a range of scales suggests that fluid flow outside of the actively-deforming gouge zones is probably fracture dominated.

  10. Scientific Drilling Into the San Andreas Fault Zone —An Overview of SAFOD’s First Five Years

    Directory of Open Access Journals (Sweden)

    Stephen Hickman


    Full Text Available The San Andreas Fault Observatory at Depth (SAFODwas drilled to study the physical and chemical processes controlling faulting and earthquake generation along an active, plate-bounding fault at depth. SAFOD is located near Parkfield, California and penetrates a section of the fault that is moving due to a combination of repeating microearthquakes and fault creep. Geophysical logs define the SanAndreas Fault Zone to be relatively broad (~200 m, containing several discrete zones only 2–3 m wide that exhibit very low P- and S-wave velocities and low resistivity. Two of these zones have progressively deformed the cemented casing at measured depths of 3192 m and 3302 m. Cores from both deforming zones contain a pervasively sheared, cohesionless, foliated fault gouge that coincides with casing deformation and explains the observed extremely low seismic velocities and resistivity. These cores are being now extensivelytested in laboratories around the world, and their composition, deformation mechanisms, physical properties, and rheological behavior are studied. Downhole measurements show that within 200 m (maximum of the active fault trace, the direction of maximum horizontal stress remains at a high angle to the San Andreas Fault, consistent with other measurements. The results from the SAFOD Main Hole, together with the stress state determined in the Pilot Hole, are consistent with a strong crust/weak fault model of the San Andreas. Seismic instrumentation has been deployed to study physics of faulting—earthquake nucleation, propagation, and arrest—in order to test how laboratory-derived concepts scale up to earthquakes occurring in nature.

  11. Fixed recurrence and slip models better predict earthquake behavior than the time- and slip-predictable models 1: repeating earthquakes (United States)

    Rubinstein, Justin L.; Ellsworth, William L.; Chen, Kate Huihsuan; Uchida, Naoki


    The behavior of individual events in repeating earthquake sequences in California, Taiwan and Japan is better predicted by a model with fixed inter-event time or fixed slip than it is by the time- and slip-predictable models for earthquake occurrence. Given that repeating earthquakes are highly regular in both inter-event time and seismic moment, the time- and slip-predictable models seem ideally suited to explain their behavior. Taken together with evidence from the companion manuscript that shows similar results for laboratory experiments we conclude that the short-term predictions of the time- and slip-predictable models should be rejected in favor of earthquake models that assume either fixed slip or fixed recurrence interval. This implies that the elastic rebound model underlying the time- and slip-predictable models offers no additional value in describing earthquake behavior in an event-to-event sense, but its value in a long-term sense cannot be determined. These models likely fail because they rely on assumptions that oversimplify the earthquake cycle. We note that the time and slip of these events is predicted quite well by fixed slip and fixed recurrence models, so in some sense they are time- and slip-predictable. While fixed recurrence and slip models better predict repeating earthquake behavior than the time- and slip-predictable models, we observe a correlation between slip and the preceding recurrence time for many repeating earthquake sequences in Parkfield, California. This correlation is not found in other regions, and the sequences with the correlative slip-predictable behavior are not distinguishable from nearby earthquake sequences that do not exhibit this behavior.

  12. Velocity Gradient Across the San Andreas Fault and Changes in Slip Behavior as Outlined by Full non Linear Tomography (United States)

    Chiarabba, C.; Giacomuzzi, G.; Piana Agostinetti, N.


    The San Andreas Fault (SAF) near Parkfield is the best known fault section which exhibit a clear transition in slip behavior from stable to unstable. Intensive monitoring and decades of studies permit to identify details of these processes with a good definition of fault structure and subsurface models. Tomographic models computed so far revealed the existence of large velocity contrasts, yielding physical insight on fault rheology. In this study, we applied a recently developed full non-linear tomography method to compute Vp and Vs models which focus on the section of the fault that exhibit fault slip transition. The new tomographic code allows not to impose a vertical seismic discontinuity at the fault position, as routinely done in linearized codes. Any lateral velocity contrast found is directly dictated by the data themselves and not imposed by subjective choices. The use of the same dataset of previous tomographic studies allows a proper comparison of results. We use a total of 861 earthquakes, 72 blasts and 82 shots and the overall arrival time dataset consists of 43948 P- and 29158 S-wave arrival times, accurately selected to take care of seismic anisotropy. Computed Vp and Vp/Vs models, which by-pass the main problems related to linarized LET algorithms, excellently match independent available constraints and show crustal heterogeneities with a high resolution. The high resolution obtained in the fault surroundings permits to infer lateral changes of Vp and Vp/Vs across the fault (velocity gradient). We observe that stable and unstable sliding sections of the SAF have different velocity gradients, small and negligible in the stable slip segment, but larger than 15 % in the unstable slip segment. Our results suggest that Vp and Vp/Vs gradients across the fault control fault rheology and the attitude of fault slip behavior.

  13. Pore Pressure Evolution in Shallow Subduction Earthquake Sequences and Effects on Aseismic Slip Transients -- Numerical Modeling With Rate and State Friction (United States)

    Liu, Y.; Rice, J. R.


    -equilibrate with that of its surroundings). This is consistent with our previous simulations, which show that the aseismic transients migrate along the strike at a higher speed under a lower, constant in time, effective normal stress. As a combination of the two factors, we show the pore pressure evolution with drops (due to dilatancy during slip) and then rises (due to shear heating) on the fault over multiple time scales. We next plan to formulate, and merge with the slip-rupture analysis, fuller fluid release models based on phase equilibria and models of transport in which the average fault-parallel permeability is a decreasing function of the effective normal stress. The thrust fault zone, at seismogenic depths and slightly downdip, is represented in a conceptually similar manner to the well-studied major continental faults, assuming the fault core materials have a lower permeability than the neighboring damaged zone. Heat diffusion in the fault core and damaged zone will also be considered in the modeling. The simulation results may help to improve our understanding of the processes of the aseismic transients observed within a transform plate boundary along the SAF near Cholame, California [Nadeau and Dolenc, 2005].

  14. Building a Global Catalog of Nonvolcanic Tremor Events Using an Automatic Detection Algorithm (United States)

    Bagley, B. C.; Revenaugh, J.


    Nonvolcanic tremor is characterized by a long-period seismic event containing a series of low-frequency earthquakes (LFEs). Tremor has been detected in regions of subduction (e.g. Kao et. al. 2007, 2008; Shelly 2006) and beneath the San Andreas fault near Cholame, California (e.g. Nadeau and Dolenc, 2005). In some cases tremor events seem to have periodicity, and these are often referred to as episodic tremor and slip (ETS). The origin of nonvolcanic tremor has been ascribed to shear slip along plate boundaries and/or high pore-fluid pressure. The apparent periodicity and tectonic setting associated with ETS has led to the suggestion that there may be a link between ETS and megathrust earthquakes. Until recently tremor detection has been a manual process requiring visual inspection of seismic data. In areas that have dense seismic arrays (e.g. Japan) waveform cross correlation techniques have been successfully employed (e.g. Obara, 2002). Kao et al. (2007) developed an algorithm for automatic detection of seismic tremor that can be used in regions without dense arrays. This method has been used to create the Tremor Activity Monitoring System (TAMS), which is used by the Geologic Survey of Canada to monitor northern Cascadia. So far the study of nonvolcanic tremor has been limited to regions of subduction or along major transform faults. It is unknown if tremor events occur in other tectonic settings, or if the current detection schemes will be useful for finding them. We propose to look for tremor events in non-subduction regions. It is possible that if tremor exists in other regions it will have different characteristics and may not trigger the TAMS system or be amenable to other existing detection schemes. We are developing algorithms for searching sparse array data sets for quasi-harmonic energy bursts in hopes of recognizing and cataloging nonvolcanic tremor in an expanded tectonic setting. Statistical comparisons against the TAMS algorithm will be made if

  15. The 'Byzantinisms' of king Stefan Radoslav

    Directory of Open Access Journals (Sweden)

    Maksimović Ljubomir


    Full Text Available The life-style and politics of Stefan Radoslav bear the mark of activities that indicated his special attachment to the Byzantine world. These activities were prompted by a combination of ideological ambitions and political reality, but they were not in keeping with the modest achievements of Radoslav's reign. Moreover, most of these activities belong to the time when Radoslav was heir to the throne. There is no doubt that Stefan Nemanjić the Grand Zhupan and subsequently the first crowned king, had exclusive connections with the Byzantine dynasty of the Angeloi, especially with the emperor Alexios III (1195-1203. In that context, the donor's inscription in the basic ring of the dome in the Church of the Mother of God in Studenica (1208, in which his father Stefan Nemanja, is mentioned as (former 'veleslavni gospodin vse srbske zemlje veli(ki župan i svat cara grčkog kir Alesija', is quite indicative. This ideological construction would acquire a contour in reality by means of a political marriage with one of the female offspring of Angeloi lineage, which would represent an alternative solution to Stefan's failed marriage with Eudocia, daughter of the emperor Alexios. Instead, several years elapsed in waging war with the Latins, the Bulgarians and the State of Epiros. However, efforts to create firmer, more tangible ties with the Angeloi dynasty from Epiros were not forgotten. Therefore, the Serbian monarch brought his eldest son Radoslav into play, intending to have him act as a link with the Angeloi bloodline. As a result of all this, the final attempt to have Radoslav become the husband of a princess from the Angelos dynasty is not surprising. At the end of 1219 or the beginning of 1220, he married Anna Doukaina, the daughter of the epirotic ruler Theodore I Angelos Doukas Komnenos, which at that point represented a marriage connection of the highest possible level between two ruling houses. Stefan's insistence on Serbia acquiring a stake

  16. Honey bees are the dominant diurnal pollinator of native milkweed in a large urban park. (United States)

    MacIvor, James Scott; Roberto, Adriano N; Sodhi, Darwin S; Onuferko, Thomas M; Cadotte, Marc W


    In eastern North America, the field milkweed, Asclepias syriaca L. (Asclepiadaceae), is used in planting schemes to promote biodiversity conservation for numerous insects including the endangered monarch butterfly, Danaus plexippus (Linnaeus) (Nymphalidae). Less is known about its pollinators, and especially in urban habitats where it is planted often despite being under increasing pressure from invasive plant species, such as the related milkweed, the dog-strangling vine (DSV), Vincetoxicum rossicum (Kleopow) Barbar. (Asclepiadaceae). During the A. syriaca flowering period in July 2016, we surveyed bees in open habitats along a DSV invasion gradient and inspected 433 individuals of 25 bee species in 12 genera for pollinia: these were affixed to bees that visited A. syriaca for nectar and contain pollen packets that are vectored (e.g., transferred) between flowers. Of all bees sampled, pollinia were found only on the nonindigenous honeybee, Apis mellifera (43% of all bees identified), as well as one individual bumblebee, Bombus impatiens Cresson. Pollinia were recorded from 45.2% of all honeybees collected. We found no relationship between biomass of DSV and biomass of A. syriaca per site. There was a significant positive correlation between A. syriaca biomass and the number of pollinia, and the proportion vectored. No relationship with DSV biomass was detected for the number of pollinia collected by bees but the proportion of vectored pollinia declined with increasing DSV biomass. Although we find no evidence of DSV flowers attracting potential pollinators away from A. syriaca and other flowering plants, the impacts on native plant-pollinator mutualisms relate to its ability to outcompete native plants. As wild bees do not appear to visit DSV flowers, it could be altering the landscape to one which honeybees are more tolerant than native wild bees.

  17. Fiji. (United States)


    Fiji is a group of volcanic islands located in the South Pacific. Because of the rough terrain in its center, that area is sparsely populated; most of Fiji's population live on the island coasts. Almost all indigenous Fijians are Christians and English is the official language. In 1970 Fiji became a fully sovereign and independent nation within the British Commonwealth. The British monarch appoints the governor general who in turn appoints as prime minister the leader of the majority party in House of Representatives. The transition to independence for Fijians was achieved in a peaceful fashion. While there are some racial tensions between the Indo-Fijians and the indigenous Fijians, the 2 major political parties and the various leaders have succeeded in maintaining order. The government of Fiji, since attaining independence, has worked hard toward economic and social progress and there have been great strides made in education, health, agriculture, and nutrition. The thrust of Fiji's economy is sugar and the 2nd component is tourism. Fiji does import a wide variety of goods but industrial development is proceeding well. Fiji encourages local and foreign investment in the hopes of promoting development and providing industrial jobs. Regional cooperation is the main element in Fiji foreign policy they joined the UN in 1970. Full diplomatic relations exist between the US and Fiji and US and Fijian officials have exchanged visits. In 1985 the US provided $1.5 million in disaster relief funds to Fiji; there is expedcted to be a bilateral aid agreement between the 2 countries in 1986. Travel notes, government and US officials, and further information are included.

  18. o Rio de Janeiro e São Paulo

    Directory of Open Access Journals (Sweden)

    Roseli Maria Martins D'Elboux


    Full Text Available This paper proposes to discuss the transformation of urban landscapes in the Paraíba River Valley as members of the coffee elite emerged in this area and a specific landscape configuration was projected there based on the use of imperial palms ( Roystonea oleracea. Chronologically speaking, the paper covers a period from 1808 to 1911; with regard to space, it focuses on the stretch between Rio de Janeiro and São Paulo, with a closer look at the case of the city of Lorena (SP, so as to encompass all the changes that took place in this region from the introduction to the decline of coffee growing as an economic activity. The urban changes during this period were accompanied by the advent and consolidation of landscapes typical of the society of coffee growers: streets lined with palm trees, a token of their close connections with the royal court, a display of their adherence to “Frenchified customs”. Such configurations were used to characterize public areas and raise them to the same status as the new buildings that gradually replaced those built in colonial style. The paper is structured around three key moments, namely: the introduction of imperial palms in Rio de Janeiro and their association with the idea of nobility and rank, and consequently with neoclassical architecture, which was brought to the colony by the 1816 French Mission; the dissemination of the use of imperial palms as a landscaping resource typical of public spaces from the royal court to the capital of São Paulo, particularly by the coffee barons during the second period of monarchic rule; and, finally, the hypothesis that the use of imperial palms to embellish public areas in São Paulo may have been introduced by a Lorena citizen associated with the coffee elite, albeit later, when Brazil was already a republic

  19. Teologi Proses Mengenai Allah dan Problem Kejahatan: Suatu Tinjauan atas Kasus Al-Nakba

    Directory of Open Access Journals (Sweden)

    Anodya Ariawan Soesilo


    Full Text Available This article presents process theology which is seen as an alternative modus offered by particular group of Christianity in dealing with radical evil. Process theology is a philosophical theology. It passes through the principles of concrescence and prehension offering the idea to identify that God's works are more likely imanent persuasion, namely, the action of love and suavity. Concrescence is acknowledgement that the actual entities not denying if its existence was formed by past objective data (experience, but deliberately work onto prehension that grasp goodness for transforming and rehabilitating the sensitivity of human history. This article will expose the space for acting faith in the mode of human responsibility, keeping an open dialogue as well restraining the false arogancy of identity fanatism of quasi-religious, persisting, and defending humanity. Such concept is based on the hope of the God who is perceptive and saving as a contribution to solve the problem of the complexity of Al-Nakba. In the mechanism of thinking process, the shadow of the failure does not disappear so that humans are asked to be seriously responsible for this. The idea of process has described God as the God who involves in relationships with humans, refuting the monarch description of the God who demands and punishes. God in theology process is a figure who does not exercise power excessively. The God who guides, accompanies, and participates patiently in the world's events still appreciates the independency of humans. That does not mean a justifi cation of irresponsible freedom. The true freedom is aligned with the God's design, namely, the righteousness, sublimity, and kindness.

  20. El asta de la lanza: los mecanismos de financiación de la guerra durante el reinado de Alfonso XI (1312-1350

    Directory of Open Access Journals (Sweden)

    Agrait, Nicolás


    Full Text Available To understand how it was possible for Alfonso XI to triumph militarily over his enemies, it is not enough to look at his courage, leadership and military strategies and tactics. One also has to study in depth the mechanisms he used to cover the high costs of his multiple campaigns and ordinary defense costs. In addition to the Crown’s regular fiscal resources and those taxes specifically related to warfare, Alfonso came to depend more and more on extraordinary levies, like monedas foreras, servicios, alcabalas, the temporary suspension of fiscal exemptions at the local level, the wealth of the Church, loans, and the sale of parts of the royal patrimony. These financial mechanisms opened up new sources of revenue to feed his military ambitions and truly constituted one of the most fundamental reasons for Alfonso’s success as a monarch and commander.

    Para entender cómo fue posible el triunfo militar de Alfonso XI sobre sus enemigos hay que mirar más allá de su valor, liderazgo y estrategias y tácticas bélicas, y estudiar a fondo los mecanismos que utilizó para sufragar los elevados costes de sus múltiples campañas y los gastos de defensa ordinarios. Además de utilizar los recursos fiscales de la Corona y los tributos tradicionales relacionados a la guerra, Alfonso dependió, cada vez más, de impuestos extraordinarios, como monedas foreras, servicios, alcabalas, la suspensión temporal de exenciones fiscales a nivel local, recursos monetarios eclesiásticos, empréstitos, y ventas de patrimonio real. Estos mecanismos financieros abrieron nuevas fuentes de ingreso para financiar sus ambiciones militares y realmente constituyeron una de las razones fundamentales del éxito de Alfonso como monarca y comandante militar.

  1. A current and comprehensive review of cyclin-dependent kinase inhibitors for the treatment of metastatic breast cancer. (United States)

    Bilgin, Burak; Sendur, Mehmet A N; Şener Dede, Didem; Akıncı, Muhammed Bülent; Yalçın, Bülent


    Resistance to endocrine treatment generally occurs over time, especially in the metastatic stage. In this paper, we aimed to review the mechanisms of cyclin-dependent kinase (CDK) 4/6 inhibition and clinical usage of new agents in the light of recent literature updates. A literature search was carried out using PubMed, Medline and ASCO and ESMO annual-meeting abstracts by using the following search keywords; "palbociclib", "abemaciclib", "ribociclib", "cyclin-dependent kinase inhibitors" and "CDK 4/6" in metastatic breast cancer (MBC). The last search was on 10 June 2017. CDKs and cyclins are two molecules that have a key role in cell cycle progression. Today, there are three highly selective CDK4/6 inhibitors in clinical development - palbociclib, ribociclib and abemaciclib. Palbociclib and ribociclib were recently approved by the US FDA in combination with letrozole for the treatment of MBC in a first-line setting, as well as palbociclib in combination with fulvestrant for hormone-receptor (HR)-positive MBC that had progressed while on previous endocrine therapy according to the PALOMA-1, MONALEESA-2 and PALOMA-3 trials, respectively. In the recently published randomized phase III MONARCH 2 trial, abemaciclib plus letrozole had longer progression-free survival and higher objective response rates with less serious adverse events in advanced HR-positive breast cancer previously treated with hormonal treatment. CDK4/6 inhibition is a new and promising target for patients with hormone-receptor-positive MBC. Both palbociclib and ribociclib showed significant additive benefit for patients receiving first-line treatment for HR-positive, epidermal growth factor receptor-2-negative advanced breast cancer. Palbociclib and abemaciclib also had significant activity in combination with fulvestrant for patients with MBC that progressed on previous endocrine therapy.

  2. Cicero and the Mixed Constitution (res publica mixta

    Directory of Open Access Journals (Sweden)

    Mitja Sadek


    Full Text Available The story of the mixed constitution is the story of the most stable and just constitution. In theory, this is a combination of at least two of the three elementary forms of government (monarchy, aristocracy and democracy, with some advantages that elementary forms may lack. It originated with the deliberation of Greek philosophers, who wanted to draw up a constitution safeguarding against the permanent variation of elementary constitutional forms and against coups d’état. For both Plato and Aristotle, the mixed constitution was, above all, the reflection of a search for balance between the two extreme forms of government, direct (Athenian democracy on the one hand and the exclusion of the people from governing on the other. The Greek theory was applied by the historian Polybius to the traditional tripartite constitution of the Roman republic. In his view, the consuls were monarchic elements, the senate an aristocratic element, and the comitia a democratic one. Cicero’s introduction of the idea of the mixed constitution in De re publica can only be understood in the light of the author’s personal situation and contemporary political circumstances. His political engagement at a time when the republic was gradually transforming into a monarchy aimed at restoring the important role of the nobility, represented by the senate. For Cicero, the mixed constitution was mainly an instrument for restoring the lost balance between the consuls, the senate, and the comitia, a last chance to save the decaying republic. The concluding part of the article addresses Alois Riklin’s recent discussion of the modern reception of the mixed constitution idea, which advances the controversial thesis that the paradigm of power division, the foundation of modern representative democracy, originates directly from the mixed constitution.

  3. Deciphering Egyptian Hieroglyphs: Towards a New Strategy for Navigation in Museums

    Directory of Open Access Journals (Sweden)

    Jaime Duque-Domingo


    Full Text Available This work presents a novel strategy to decipher fragments of Egyptian cartouches identifying the hieroglyphs of which they are composed. A cartouche is a drawing, usually inside an oval, that encloses a group of hieroglyphs representing the name of a monarch. Aiming to identify these drawings, the proposed method is based on several techniques frequently used in computer vision and consists of three main stages: first, a picture of the cartouche is taken as input and its contour is localized. In the second stage, each hieroglyph is individually extracted and identified. Finally, the cartouche is interpreted: the sequence of the hieroglyphs is established according to a previously generated benchmark. This sequence corresponds to the name of the king. Although this method was initially conceived to deal with both high and low relief writing in stone, it can be also applied to painted hieroglyphs. This approach is not affected by variable lighting conditions, or the intensity and the completeness of the objects. This proposal has been tested on images obtained from the Abydos King List and other Egyptian monuments and archaeological excavations. The promising results give new possibilities to recognize hieroglyphs, opening a new way to decipher longer texts and inscriptions, being particularly useful in museums and Egyptian environments. Additionally, devices used for acquiring visual information from cartouches (i.e., smartphones, can be part of a navigation system for museums where users are located in indoor environments by means of the combination of WiFi Positioning Systems (WPS and depth cameras, as unveiled at the end of the document.

  4. Church History and the Predicament of the Orthodox Hierarchy in the Russian Empire of the Early 1800s

    Directory of Open Access Journals (Sweden)

    Eugene I. Lyutko


    Full Text Available In this article, the author tries to reflect the emergence of the intellectual concept of “Church History” through a number of theoretical frameworks, setting this discursive turn on the map of the epoch using several narratives. The first is the problem of the cultural gap arising during the 18th century between the intellectual elites of the nobility and clergy. Second, we examine the bureaucratization of the empire leading both to the convergence of parallel “ecclesiastical” and “civil” administrative structures and to the emergence of the bureaucratic layer between episcopate and the monarch, who was considered as the formal “head” of the earthly ecclesiastical structure. Third, we consider the establishment of the administrative bonds between governmental authorities and individuals, which were understood as being in competition for the “pastoral” power of the church hierarchy. We next examine the change in the mode of knowledge distribution, which took place within the emergence of the “public sphere” in the early 19th-century Russian Empire. Finally, we look at the problem of the national identity emerging in the late 18th and early 19th centuries, which was centered around the concept of the ethnic community and political body (and its history rather than on the community of believers actualized in the discourse of the epoch as the concept of Church (and its history. All those narratives on social change strive to explain the global change in Orthodox theology, which became centered on ecclesiology. This change might be effectively problematized as a transition between first and second “orders of theology” within the framework proposed by G. Kaufman. This method of explanation may be especially productive when it comes to drawing an analogy between Russian and Western theology in the modern period.

  5. Transcripción revisada del informe de Pedro de Ayala de 1498 sobre las expediciones inglesas de descubrimiento

    Directory of Open Access Journals (Sweden)

    Robles Macías, Luis A.


    Full Text Available The report sent by Pedro de Ayala to the Catholic Monarchs from London in 1498 is one of the main sources on the earliest English expeditions to North America. The history of its deciphering, interpretation and translation is presented and, as none of the transcriptions carried out up to now is found to be entirely satisfactory, a new one is proposed based on a literal deciphering of the document, along with translations into French and English. Furthermore, some particularly difficult parts of the text are discussed in detail that, having been misunderstood in earlier editions of the report, led to probably erroneous conclusions about the biography of John Cabot, the itinerary of his voyage of 1497, and the name of the monk who went with him on his expedition of 1498.El informe enviado por Pedro de Ayala a los Reyes Católicos desde Londres en 1498 es una de las fuentes fundamentales sobre las primeras expediciones inglesas a Norteamérica. En este estudio se presenta la historia de su descifrado, interpretación y traducción y, constatando que ninguna de las transcripciones realizadas hasta ahora es completamente exacta, se propone una nueva basada en un descifrado literal del documento, acompañada de traducciones al francés y al inglés. Se discuten asimismo en detalle algunas frases particularmente difíciles del texto que, debido a su malinterpretación en ediciones anteriores, condujeron a conclusiones probablemente erróneas sobre la biografía de Juan Caboto, el itinerario de su viaje de 1497 y el nombre del religioso que fue con él en su expedición de 1498.

  6. Una donación entre judíos segovianos, originalmente en hebreo, del año 1487

    Directory of Open Access Journals (Sweden)

    Bonifacio Bartolomé Herrero


    Full Text Available En este trabajo se transcribe y comenta un documento de donación redactado originalmente en hebreo en 1487 y cuyo contenido se ha conservado a través de su traducción al castellano en 1496. La importancia de este texto radica en que es el único testimonio conocido hasta el momento de un documento en hebreo redactado en el seno de la aljama judía de Segovia, una de las más importantes de la península ibérica en el último cuarto del siglo XV. Para situar el documento en su adecuado contexto se ofrece una visión general de la comunidad judía de Segovia durante el reinado de los Reyes Católicos y una breve biografía del receptor de la donación, el herrero judío Yuçe Biton, convertido más tarde al cristianismo con el nombre de Alonso de Palencia.The present work transcribes and comments a document of donation originally written in Hebrew in 1487. The contents of the document have been kept thanks to its translation to the Spanish language in 1496. Its relevance lies on the fact that it is, so far, the only testimony in Hebrew language we have evidence on, written in the Jewish quarter in the town of Segovia, which was one of the most important Hebrew communities in the Iberian Peninsula in the last quarter of the 15th century. In order to set the document in its context, it is offered a general perspective of the Hebrew community in Segovia during the Catholic Monarchs kingdom and a brief biography of the receiver of the donation, the Jewish blacksmith Yuçe Biton, who became a Christian afterwards under the name of Alonso de Palencia.

  7. Evidence for adaptive radiation from a phylogenetic study of plant defenses (United States)

    Agrawal, Anurag A.; Fishbein, Mark; Halitschke, Rayko; Hastings, Amy P.; Rabosky, Daniel L.; Rasmann, Sergio


    One signature of adaptive radiation is a high level of trait change early during the diversification process and a plateau toward the end of the radiation. Although the study of the tempo of evolution has historically been the domain of paleontologists, recently developed phylogenetic tools allow for the rigorous examination of trait evolution in a tremendous diversity of organisms. Enemy-driven adaptive radiation was a key prediction of Ehrlich and Raven's coevolutionary hypothesis [Ehrlich PR, Raven PH (1964) Evolution 18:586–608], yet has remained largely untested. Here we examine patterns of trait evolution in 51 North American milkweed species (Asclepias), using maximum likelihood methods. We study 7 traits of the milkweeds, ranging from seed size and foliar physiological traits to defense traits (cardenolides, latex, and trichomes) previously shown to impact herbivores, including the monarch butterfly. We compare the fit of simple random-walk models of trait evolution to models that incorporate stabilizing selection (Ornstein-Ulenbeck process), as well as time-varying rates of trait evolution. Early bursts of trait evolution were implicated for 2 traits, while stabilizing selection was implicated for several others. We further modeled the relationship between trait change and species diversification while allowing rates of trait evolution to vary during the radiation. Species-rich lineages underwent a proportionately greater decline in latex and cardenolides relative to species-poor lineages, and the rate of trait change was most rapid early in the radiation. An interpretation of this result is that reduced investment in defensive traits accelerated diversification, and disproportionately so, early in the adaptive radiation of milkweeds. PMID:19805160

  8. Spreken vanuit het graf. De stoffelijke resten van Willem van Oranje in hun politiek-culturele betekenis

    Directory of Open Access Journals (Sweden)

    C. Santing


    Full Text Available Voice from the Grave. The politico-cultural relevance of the mortal remains of William of OrangeBalthazar Gerards’ murder of William of Orange on July 10th, 1584 is a canonical fact in Dutch history, but thus far this significant event has not yet been analysed on its own merits, i.e. in terms of the material disposition of his assassinated body. Hence, this article examines the literal and figurative - or cultural and material - handling of the physical remains of William the Silent and their subsequent iconification and deification. The historiographical background and intellectual inspiration for this article are provided by the work of Ernst Kantorowic’s pupil Ralph Giesey, Agostino Paravicini Bagliani and Joanna Woodward. All authors point to the increasingly lavish pomp and circumstance of monarchical funeral culture, which was imitated by Europe’s high aristocracy. The autopsy on William the Silent’s corpse was completed with a meticulous embalming. The preserved body and its subsequent lying in state amidst an extensive funerary apparatus and the full pompa of the burial ceremony contained numerous reminiscences of Charles V Brussels’ funeral, in which the Prince had played a prominent role. Also, after having examined the sculptural iconography of Hendrik de Keyser’s mausoleum in Delft’s Nieuwe Kerk, it can be concluded that the significance of William the Silent’s murdered body was much more royal and catholic than traditional historiographical views on the Dutch Republic would lead us to suspect.

  9. Clavulina-Membranomyces is the most important lineage within the highly diverse ectomycorrhizal fungal community of Abies religiosa. (United States)

    Argüelles-Moyao, Andrés; Garibay-Orijel, Roberto; Márquez-Valdelamar, Laura Margarita; Arellano-Torres, Elsa


    Abies religiosa is an endemic conifer of Mexico, where its monodominant forests are the winter refuge of the monarch butterfly. Due to climate change, it has been estimated that by 2090, A. religiosa populations will decline by 96.5 %. To achieve success, reforestation programs should consider its ectomycorrhizal (ECM) fungi. We used ITS nrDNA sequences to identify the ECM fungi associated with A. religiosa and, based on its abundance and frequency, determined the diversity and community structure in a pure A. religiosa forest near Mexico City. Using sequence metadata, we inferred the species geographic distribution and host preferences. We conducted phylogenetic analyses of the Clavulinaceae (the most important family). The ECM community held 83 species, among which the richest genera were Inocybe (21 species), Tomentella (10 species), and Russula (8 species). Besides its low species richness, the Clavulina-Membranomyces lineage was the most dominant family. Clavulina cf. cinerea and Membranomyces sp. exhibited the highest relative abundance and relative frequency values. Phylogenetic analyses placed the Clavulinaceae genotypes in three different clades: one within Membranomyces and two within Clavulina. A meta-analysis showed that the majority of the ECM fungi (45.78 %) associated with A. religiosa in Mexico have also been sequenced from North America and are shared by Pinaceae and Fagaceae. In contrast, because they have not been sequenced previously, 32.2 % of the species have a restricted distribution. Here, we highlight the emerging pattern that the Clavulina-Membranomyces lineage is dominant in several ECM communities in the Neotropics, including Aldinia and Dicymbe legume tropical forests in the Guyana Shield, the Alnus acuminata subtropical communities, and the A. religiosa temperate forests in Mexico.

  10. Abemaciclib: a CDK4/6 inhibitor for the treatment of HR+/HER2– advanced breast cancer

    Directory of Open Access Journals (Sweden)

    Corona SP


    Full Text Available Silvia Paola Corona,1 Daniele Generali2 1Radiation Oncology Department, Peter MacCallum Cancer Centre, Bentleigh East, VIC, Australia; 2Department of Medical, Surgery and Health Sciences, University of Trieste, Trieste, Italy Abstract: Although early breast cancer (BC is highly curable, advanced or metastatic disease poses numerous challenges in terms of medical management and treatment decisions and is associated with significantly worse prognosis. Among the new targeted agents, anticancer drugs exploiting the cell-cycle machinery have shown great potential in preclinical studies. CDK4/6 inhibitors target the cyclin D/CDK/retinoblastoma signaling pathway, inducing cell-cycle arrest, reduced cell viability and tumor shrinking. As the cyclin D/CDK complex is activated downstream of estrogen signaling, the combination of CDK4/6 inhibitors with standard endocrine therapies represents a rational approach to elicit synergic antitumor activity in hormone receptor-positive BC. The results of clinical trials have indeed confirmed the superiority of the combination of CDK4/6 inhibitors plus endocrine therapies over endocrine therapy alone. Currently approved are three compounds that exhibit similar structural characteristics as well as biological and clinical activities. Abemaciclib is the latest CDK4/6 inhibitor approved by the US Food and Drug Administration (FDA in view of the results of the MONARCH 1 and 2 trials. Further trials are ongoing as other important questions await response. In this review, we focus on abemaciclib to examine preclinical and clinical results, describing current therapeutic indications, open questions and ongoing clinical trials. Keywords: CDK4/6 inhibitor, abemaciclib, breast cancer, hormone receptor-positive BC, metastatic BC, mBC

  11. Diagnosing the Kaiser: Psychiatry, Wilhelm II and the Question of German War Guilt The William Bynum Prize Essay 2016. (United States)

    Freis, David


    After his abdication in November 1918, the German emperor Wilhelm II continued to haunt the minds of his people. With the abolition of the lese-majesty laws in the new republic, many topics that were only discussed privately or obliquely before could now be broached openly. One of these topics was the mental state of the exiled Kaiser. Numerous psychiatrists, physicians and laypeople published their diagnoses of Wilhelm in high-circulation newspaper articles, pamphlets, and books shortly after the end of the war. Whether these diagnoses were accurate and whether the Kaiser really was mentally ill became the issue of a heated debate.This article situates these diagnoses of Wilhelm II in their political context. The authors of these diagnoses - none of whom had met or examined Wilhelm II in person - came from all political camps and they wrote with very different motives in mind. Diagnosing the exiled Kaiser as mentally ill was a kind of exorcism of the Hohenzollern rule, opening the way for either a socialist republic or the hoped-for rule of a new leader. But more importantly, it was a way to discuss and allocate political responsibility and culpability. Psychiatric diagnoses were used to exonerate both the Emperor (for whom the treaty of Versailles provided a tribunal as war criminal) and the German nation. They were also used to blame the Kaiser's entourage and groups that had allegedly manipulated the weak-willed monarch. Medical concepts became a vehicle for a debate on the key political questions in interwar Germany.

  12. Tapada das Necessidades em Lisboa : a historia de um jardim esquecido

    Directory of Open Access Journals (Sweden)

    João Alburquerque Carreiras


    Full Text Available O trabalho apresentado consiste numa análise histórica da Tapada das Necessidades, jardim situado na cidade de Lisboa, formando um dos seus maiores espaços verdes. A Tapada foi construída por D.João V, o monarca que em Portugal mais simbolizou o Absolutismo, reflectindo-o nas suas criações Barrocas. Com o Liberalismo surge D.Fernando, que vai ser o intérprete de uma enorme transformação da tapada, protagonizando o período mais importante na sua evolução. A história da Tapada entranha-se na própria história de Portugal, acompanhando o absolutismo, a transição para o liberalismo, os últimos movimentos da monarquia e os primeiros passos da República.The presented work consists in a historical analysis of the Tapada das Necessidades, a garden situated in the City of Lisbon, forming one of its bigger green spaces. The Tapada was built by D.João V, the monarch that in Portugal more symbolized the Absolutism, reflecting it in its Baroque creations. With the Liberalism appears D.Fernando, who will be the interpreter of an enourmous hashing of the Tapada, carrying out the more important period of its evolution. The history of the Tapada penetrates in Portugal own history, following the absolutism, the transition for the liberalism, the last moves of the monarchy and the first steps of the republic.

  13. Британские истоки архитектурных форм дворца Бржозовского в Одессе

    Directory of Open Access Journals (Sweden)

    Yuri Pismak


    Full Text Available An outstanding British architect Edward Blore (1787–1879 is the main author of project of Vorontsov’s Palace in Alupka (the Crimea, 1830–1848. In the opining of many researchers the Vorontsov’s Palace in Alupka is the best of Edward Blore’s works. Seemingly under impression of Vorontsov’s Alupka Palace, Brzhozovsky’s Palace (Shakh’s Palace. 1851–1852 was built to the project of a renowned Odessa architect Felix Gonsiorovski (1815–1891 in 2 Nadezhdinskaya (Gogol street in Odessa. Apart from Tudor elements and Castel style, these two Palaces are related yet by the fact that they both are situated on the Black Sea coast. Facades of Brzhozovsky’s Palace remind the same of a distinguished monument of architecture, historically connected with the names of British monarchs – the Hampton Court Palace. Hampton Court Palace was recognized as the acme of the English palace architecture. Splendid ensemble sprawling on the bank of the Thames River near London was initiated in 1514. The great vogue for all Gothic sprang up in St. Petersburg under the spell of Walter Scott’s (1771–1832 historic novels. The article tells also about prototypes of Tudor arch – an element which characterizes English architecture of 16th century. This form of the arch was spread to many countries from the United Kingdom. However, long before England, this form of arch was used by architects of Iran and other countries of Central Asia. Formation of Odessa’s and all Southern Region of the Ukraine architectural appearance in the XIXth century was influenced by different architectural European schools. Especially in this article is noted British influence. Consequently it seems proper to consider this process in the retrospect of the European development as a whole.

  14. El ordenamiento de Medina del Campo de 1328 = The Medina del Campo Legal Code of 1328

    Directory of Open Access Journals (Sweden)

    Ana Arranz Guzmán


    Full Text Available Durante los dos últimos siglos la controversia historiográfica ha envuelto en una cierta nebulosa la reunión mantenida por Alfonso XI en Medina del Campo en el año 1328, siendo calificada por algunos historiadores de auténticas Cortes o negando, otros, su propia existencia. Con la transcripción en estas páginas del Ordenamiento elaborado en ella, así como con el análisis realizado del texto y su cotejo con las actas editadas de las Cortes madrileñas de 1329, se espera cerrar, quizá definitivamente, la larga polémica suscitada por esta asamblea, a la vez que aportar algunos datos más sobre la sobresaliente labor legislativa llevada a cabo por el monarca castellano.Scholarship over the last two centuries has debated once and again without reaching a consensus over the meeting held by Alfonso XI in Medina del Campo in 1328, considered by some historians as an authentic parliament (Cortes while others deny it. With the transcription of this legal code (Ordenamiento and its comparison with the edited account of the proceedings of the Cortes held in Madrid in 1329, our purpose is to lay to rest the debate provoked by this assembly. Furthermore, we intend to present additional information relating to the salient legislative work undertaken by this Castilian monarch.

  15. La Real Audiencia de Lima, el sello real y la garantía de la justicia (The Real Audiencia of Lima, the royal seal and ensuring justice

    Directory of Open Access Journals (Sweden)

    José de la Puente Brunke


    Full Text Available Resumen: En este trabajo, referido a la Real Audiencia de Lima, nos proponemos poner de relieve la crucial importancia de la institución audiencial en el contexto de la administración indiana. En primer lugar nos referiremos a los libros y documentos de la Audiencia limeña, y a la trascendencia de su conservación en el contexto político indiano. Luego abordaremos el estudio de la Audiencia como depositaria del Sello Real, lo cual hacía que representara al propio monarca, y que tuviera la autoridad para emitir disposiciones en nombre del rey, y como si el rey las hubiera firmado. Por último, reflexionaremos en torno a la labor jurisdiccional de la Audiencia, y a los criterios en virtud de los cuales se consideraba que sus magistrados podían garantizar que la justicia prevaleciera.Abstract: In this paper, based on the Real Audiencia de Lima, we intend to highlight the crucial importance of audiencial institution in the context of the Indian administration. First we refer to the books and records of the Lima High Court, and the importance of conservation in the Indian political context. Then board the studio audience as depository Privy Seal, which was to represent the monarch himself, and he had the authority to issue rules on behalf of the king, and the king as if it had been signed. Finally, we will think about the judicial work of the Court, and the criteria under which it was considered that its judges could ensure that justice prevails.

  16. As filhas de Eva: religião e relações de gênero na justiça medieval portuguesa Eve's daughters: religion and gender relations in the portuguese medieval justice

    Directory of Open Access Journals (Sweden)

    Edlene Oliveira Silva


    Full Text Available Este artigo analisa as representações de mulheres presentes nas Ordenações Afonsinas, código jurídico português elaborado no século XV que definiu e classificou detalhadamente vários crimes considerados tipicamente femininos e estipulou punições rigorosas. Dentre esses delitos, trataremos de alguns aspectos do adultério, do concubinato e da alcovitagem. Informado pelas representações de gênero, o discurso jurídico do Estado monárquico luso legitimou a perseguição empreendida pela Igreja às mulheres "desviantes". O olhar da justiça era influenciado pelo imaginário religioso cristão e medieval, repleto de ideias patriarcais e misóginas que associavam o feminino ao arquétipo da Eva pecadora, a primeira mulher que se deixou seduzir pelos ardis malignos do demônio.This article analyzes the representations of women in the Ordenações Afonsinas, the Portuguese juridical code elaborated in the 15th century that defined and classified in detail several crimes considered typically feminine, and that stipulated rigorous punishments. Among those crimes, we will discuss some aspects of adultery, concubinage, and panderism. Informed by gender representations, the juridical discourse of the Lusitanian monarchical State legitimated the persecution of "deviating" women undertaken by the Church. The view of justice was influenced by Christian/medieval religious imaginary, full of patriarchal and misogynistic ideas that associated the feminine to Eve's archetype of the sinner, the first woman to be seduced by the Devil's evil artifices.

  17. Hemofilie onder die nasate van koningin Victoria

    Directory of Open Access Journals (Sweden)

    Francois P. Retief


    Full Text Available Hierdie studie bespreek die uitwerking van die sogenaamde Victoriaanse hemofilie op Europese koningshuise gedurende die negentiende en twintigste eeu. Hemofilie as molekulêre defek word verduidelik en die kliniese beeld van die siekte word aangedui. ’n Bespreking van toepaslike terapeutiese ingrepe volg. Dan word ’n historiese oorsig verskaf van die verspreiding van Victoriaanse hemofilie vanaf koningin Victoria (Britse monarg, 1837–1901 via sommige van haar dogters na ander lede van die Britse koningshuis en ook na die Duitse, Russiese en Spaanse koningshuise. Elf bevestigde gevalle van hemofilie onder lede van koningin Victoria se nageslag word vermeld, asook drie ander moontlike gevalle van die siekte. Die effek van hemofilie op die verloop van die geskiedenis word ook ondersoek.Haemophilia amongst the descendants of Queen Victoria. This study discusses the impact of ‘Victorian haemophilia’ on the royal houses of Europe during the 19th and 20th centuries. Haemophilia as a molecular defect is explained and the clinical picture of the condition is indicated. Applicable therapeutic interventions also receive attention. Next, an historical review is provided of how ‘Victorian haemophilia’ spread from Queen Victoria (British monarch, 1837–1901 via some of her daughters to other members of the British royal family and also to the royal houses of Germany, Russia and Spain. Eleven confirmed cases of haemophilia amongst the descendants of Queen Victoria are mentioned, as well as three other possible cases. The effect of haemophilia on the course of history is also investigated.

  18. Algunas gestiones de mediación del Marqués de Villalobar durante la I Guerra Mundial

    Directory of Open Access Journals (Sweden)

    Álvaro Lozano Cutanda


    Full Text Available En este artículo deseo recordar a una figura de primer orden de la diplomacia española: Don Rodrigo Saavedra y Vinent, Marqués de Villalobar. Se trató de un personaje clave dentro la política exterior del Rey Alfonso XIII. Su amistad y vínculos con la Corona, hicieron de él un defensor de la misma en sus puestos diplomáticos y encarnó como nadie la política exterior que quiso llevar a cabo el Rey Alfonso XIII. El gran mérito del Embajador Villalobar fue el de superar una grave incapacidad física y con una gran fuerza de voluntad haber llegado a representar a España. Esa sobrehumana superación personal, unida a su fuerte sentimiento monárquico, aristocrático y humanitario, fueron las constantes en su vida y su profesión.In this article I would like to shed light on an outstanding Spanihs diplomat: Don Rodrigo Saavedra y Vinent, Marquis of Villalobar. He was a key figure in the foreign policy of King Alfonso XIII. His friendship and links with the Spanish royalty made him a strong supporter of the Crown in his diplomatic posts and he embodied like no one else the foreign policy of his King. The greatest merit of Ambassador Villalobar, was to overeóme a big physical handicap, and through sheer will power, to become an ambassador. Without doubts, overcoming that handicap profoundly marked his personality during his entire Ufe. That immense will power and his strong monarchic, aristocratic and humanitarian feelings, were the most outstanding features of his Ufe and work.

  19. Roughness Effects on the Formation of a Leading Edge Vortex (United States)

    Elliott, Cassidy; Lang, Amy; Wahidi, Redha; Wilroy, Jacob


    Microscopic scales cover the wings of Monarch butterflies, creating a patterned surface that acts as a natural energy capture mechanism. This patterning is thought to delay the growth of the leading edge vortex (LEV) produced by the flapping motion of a wing. Increased skin friction caused by the scales leads to a weaker LEV being shed into the butterfly's wake, lessening drag and increasing flight efficiency. To test how this roughness effects LEV formation, a plate of random roughness was designed in SolidWorks and printed on the Objet 30 Pro 3D printer. A 2x3x5 cubic foot tow tank was used to test the rough plate at Reynold's numbers of 1500, 3000, and 6000 (velocities of 8, 16, and 32 mm/s) at an angle of attack of 45 degrees. Images were captured of the LEV generated when the plate was towed upwards through the particle-seeded flow. These images were used to determine the XY velocity of the particles using a technique called Digital Particle Image Velocimetry (DPIV). Codes written in MATLAB were used to track and measure the strength of the LEV. Circulation values for the randomly-rough plate were then compared to the same values generated in a previous experiment that used a smooth plate and a grooved plate to determine the effect of the patterning on vortex development. Funding for this research project was provided by the National Science Foundation under the Research Experience for Undergraduates (REU) program (REU Supplement CBET 1628600 under CBET 1335848).

  20. The effect of butterfly-scale inspired patterning on leading-edge vortex growth (United States)

    Wilroy, Jacob Aaron

    Leading edge vortices (LEVs) are important for generating thrust and lift in flapping flight, and the surface patterning (scales) on butterfly wings is hypothesized to play a role in the vortex formation of the LEV. To simplify this complex flow problem, an experiment was designed to focus on the alteration of 2-D vortex development with a variation in surface patterning. Specifically, the secondary vorticity generated by the LEV interacting at the patterned surface was studied, as well as the subsequent effect on the LEV's growth rate and peak circulation. For this experiment, rapid-prototyped grooves based on the scale geometry of the Monarch butterfly (Danaus plexippus) were created using additive manufacturing and were attached to a flat plate with a chordwise orientation, thus increasing plate surface area. The vortex generated by the grooved plate was then compared to a smooth plate case in an experiment where the plate translated vertically through a 2 x 3 x 5 cubic foot tow tank. The plate was impulsively started in quiescent water and flow fields at Rec = 1416, 2833, and 5667 are examined using Digital Particle Image Velocimetry (DPIV). The maximum vortex formation number is 2.8 and is based on the flat plate travel length and chord length. Flow fields from each case show the generation of a secondary vortex whose interaction with the shear layer and LEV caused different behaviors depending upon the surface type. The vortex development process varied for each Reynolds number and it was found that for the lowest Reynolds number case a significant difference does not exist between surface types, however, for the other two cases the grooves affected the secondary vortex's behavior and the LEV's ability to grow at a rate similar to the smooth plate case.