WorldWideScience

Sample records for nonlinear ontogenetic shift

  1. Ontogenetic shifts in functional morphology of dragonfly legs (Odonata: Anisoptera).

    Science.gov (United States)

    Leipelt, Klaus Guido; Suhling, Frank; Gorb, Stanislav N

    2010-12-01

    Anisopteran leg functions change dramatically from the final larval stadium to the adult. Larvae use legs mainly for locomotion, walking, climbing, clinging, or burrowing. Adults use them for foraging and grasping mates, for perching, clinging to the vegetation, and for repelling rivals. In order to estimate the ontogenetic shift in the leg construction from the larva to the adult, this study quantitatively compared lengths of fore, mid, and hind legs and the relationships between three leg segments, femur, tibia, and tarsus, in larval and adult Anisoptera of the families Gomphidae, Aeshnidae, Cordulegastridae, Corduliidae, and Libellulidae, represented by two species each. We found that leg segment length ratio as well as ontogenetic shift in length ratios was different between families, but rather similar within the families. While little ontogenetic shift occurred in Aeshnidae, there were some modifications in Corduliidae and Libellulidae. The severest shift occurred in Gomphidae and Cordulegastridae, both having burrowing larvae. These two families form a cluster, which is in contrast to their taxonomic relationship within the Anisoptera. Cluster analysis implies that the function of larval legs is primarily responsible for grouping, whereas adult behavior or the taxonomic relationships do not explain the grouping. This result supports the previous hypothesis about the convergent functional shift of leg characters in the dragonfly ontogenesis.

  2. Detecting spatial ontogenetic niche shifts in complex dendritic ecological networks

    Science.gov (United States)

    Fields, William R.; Grant, Evan; Lowe, Winsor H.

    2017-01-01

    Ontogenetic niche shifts (ONS) are important drivers of population and community dynamics, but they can be difficult to identify for species with prolonged larval or juvenile stages, or for species that inhabit continuous habitats. Most studies of ONS focus on single transitions among discrete habitat patches at local scales. However, for species with long larval or juvenile periods, affinity for particular locations within connected habitat networks may differ among cohorts. The resulting spatial patterns of distribution can result from a combination of landscape-scale habitat structure, position of a habitat patch within a network, and local habitat characteristics—all of which may interact and change as individuals grow. We estimated such spatial ONS for spring salamanders (Gyrinophilus porphyriticus), which have a larval period that can last 4 years or more. Using mixture models to identify larval cohorts from size frequency data, we fit occupancy models for each age class using two measures of the branching structure of stream networks and three measures of stream network position. Larval salamander cohorts showed different preferences for the position of a site within the stream network, and the strength of these responses depended on the basin-wide spatial structure of the stream network. The isolation of a site had a stronger effect on occupancy in watersheds with more isolated headwater streams, while the catchment area, which is associated with gradients in stream habitat, had a stronger effect on occupancy in watersheds with more paired headwater streams. Our results show that considering the spatial structure of habitat networks can provide new insights on ONS in long-lived species.

  3. Hitting the moving target: modelling ontogenetic shifts with stable isotopes reveals the importance of isotopic turnover.

    Science.gov (United States)

    Hertz, Eric; Trudel, Marc; El-Sabaawi, Rana; Tucker, Strahan; Dower, John F; Beacham, Terry D; Edwards, Andrew M; Mazumder, Asit

    2016-05-01

    Ontogenetic niche shifts are widely prevalent in nature and are important in shaping the structure and dynamics of ecosystems. Stable isotope analysis is a powerful tool to assess these shifts, with δ(15) N providing a measure of trophic level and δ(13) C a measure of energy source. Previous applications of stable isotopes to study ontogenetic niche shifts have not considered the appreciable time lag between diet and consumer tissue associated with isotopic turnover. These time lags introduce significant complexity into field studies of ontogenetic niche shifts. Juvenile Chinook salmon (Oncorhynchus tshawytscha) migrate from freshwater to marine ecosystems and shift their diet from feeding primarily on invertebrates to feeding primarily on fish. This dual ontogenetic habitat and diet shift, in addition to the long time lag associated with isotopic turnover, suggests that there is potential for a disconnect between the prey sources that juvenile salmon are consuming, and the inferred prey sources from stable isotopes. We developed a model that considered ontogenetic niche shifts and time lags associated with isotopic turnover, and compared this 'ontogeny' model to one that considered only isotopic turnover. We used a Bayesian framework to explicitly account for parameter uncertainty. Data showed overwhelming support for the ontogeny model relative to the isotopic turnover model. Estimated variables from best model fits indicate that the ontogeny model predicts a much greater reliance on fish prey than does the stomach content data. Overall, we found that this method of quantifying ontogenetic niche shifts effectively accounted for both isotopic turnover and ontogenetic diet shifts; a finding that could be widely applicable to a variety of systems.

  4. Growth dynamics of juvenile loggerhead sea turtles undergoing an ontogenetic habitat shift.

    Science.gov (United States)

    Ramirez, Matthew D; Avens, Larisa; Seminoff, Jeffrey A; Goshe, Lisa R; Heppell, Selina S

    2017-04-01

    Ontogenetic niche theory predicts that individuals may undergo one or more changes in habitat or diet throughout their lifetime to maintain optimal growth rates, or to optimize trade-offs between mortality risk and growth. We combine skeletochronological and stable nitrogen isotope (δ(15)N) analyses of sea turtle humeri (n = 61) to characterize the growth dynamics of juvenile loggerhead sea turtles (Caretta caretta) during an oceanic-to-neritic ontogenetic shift. The primary objective of this study was to determine how ontogenetic niche theory extends to sea turtles, and to individuals with different patterns of resource use (discrete shifters, n = 23; facultative shifters n = 14; non-shifters, n = 24). Mean growth rates peaked at the start of the ontogenetic shift (based on change in δ(15)N values), but returned to pre-shift levels within 2 years. Turtles generally only experienced 1 year of relatively high growth, but the timing of peak growth relative to the start of an ontogenetic shift varied among individuals (before, n = 14; during, n = 12; after, n = 8). Furthermore, no reduction in growth preceded the transition, as is predicted by ontogenetic niche theory. Annual growth rates were similar between non-transitioning turtles resident in oceanic and neritic habitats and turtles displaying alternative patterns of resource use. These results suggest that factors other than maximization of size-specific growth may more strongly influence the timing of ontogenetic shifts in loggerhead sea turtles, and that alternative patterns of resource use may have limited influence on somatic growth and age at maturation in this species.

  5. Coexistence of Predator and Prey in Intraguild Predation systems with Ontogenetic Niche Shifts

    NARCIS (Netherlands)

    Hin, V.; Schellekens, T.; Persson, L.; Roos, A.M.

    2011-01-01

    In basic intraguild predation (IGP) systems, predators and prey also compete for a shared resource. Theory predicts that persistence of these systems is possible when intraguild prey is superior in competition and productivity is not too high. IGP often results from ontogenetic niche shifts, in

  6. Shape shifting predicts ontogenetic changes in metabolic scaling in diverse aquatic invertebrates

    DEFF Research Database (Denmark)

    Glazier, Douglas S.; Hirst, Andrew G.; Atkinson, D.

    2016-01-01

    Metabolism fuels all biological activities, and thus understanding its variation is fundamentally important. Much of this variation is related to body size, which is commonly believed to follow a 3/4-power scaling law. However, during ontogeny, many kinds of animals and plants show marked shifts...... that may be applied to ontogenetic shifts in bR shown by many kinds of animals and plants...

  7. Intersexual allometry differences and ontogenetic shifts of coloration patterns in two aquatic turtles, Graptemys oculifera and Graptemys flavimaculata

    Science.gov (United States)

    Ennen, Joshua R.; Lindeman, Peter V.; Lovich, Jeffrey E.

    2015-01-01

    Coloration can play critical roles in a species' biology. The allometry of color patterns may be useful for elucidating the evolutionary mechanisms responsible for shaping the traits. We measured characteristics relating to eight aspects of color patterns from Graptemys oculifera and G. flavimaculata to investigate the allometric differences among male, female, and unsexed juvenile specimens. Additionally, we investigated ontogenetic shifts by incorporating the unsexed juveniles into the male and female datasets. In general, male color traits were isometric (i.e., color scaled with body size), while females and juvenile color traits were hypoallometric, growing in size more slowly than the increase in body size. When we included unsexed juveniles in our male and female datasets, our linear regression analyses found all relationships to be hypoallometric and our model selection analysis found support for nonlinear models describing the relationship between body size and color patterns, suggestive of an ontogenetic shift in coloration traits for both sexes at maturity. Although color is critical for many species' biology and therefore under strong selective pressure in many other species, our results are likely explained by an epiphenomenon related to the different selection pressures on body size and growth rates between juveniles and adults and less attributable to the evolution of color patterns themselves.

  8. Ontogenetic shifts and spatial associations in organ positions for snakes.

    Science.gov (United States)

    Anderson, Gretchen E; Secor, Stephen M

    2015-12-01

    Snakes possess an elongated body form and serial placement of organs which provides the opportunity to explore historic and adaptive mechanisms of organ position. We examined the influence of body size and sex on the position of, and spatial associations between, the heart, liver, small intestine, and right kidney for ten phylogenetically diverse species of snakes that vary in body shape and habitat. Snake snout-vent length explained much of the variation in the position of these four organs. For all ten species, the position of the heart and liver relative to snout-vent length decreased as a function of size. As body size increased from neonate to adult, these two organs shifted anteriorly an average of 4.7% and 5.7% of snout-vent length, respectively. Similarly, the small intestine and right kidney shifted anteriorly with an increase in snout-vent length for seven and five of the species, respectively. The absolute and relative positioning of these organs did not differ between male and female Burmese pythons (Python molurus). However, for diamondback water snakes (Nerodia rhombifer), the liver and small intestine were more anteriorly positioned in females as compared to males, whereas the right kidney was positioned more anteriorly for males. Correlations of residuals of organ position (deviation from predicted position) demonstrated significant spatial associations between organs for nine of the ten species. For seven species, individuals with hearts more anterior (or posterior) than predicted also tended to possess livers that were similarly anteriorly (or posteriorly) placed. Positive associations between liver and small intestine positions and between small intestine and right kidney positions were observed for six species, while spatial associations between the heart and small intestine, heart and right kidney, and liver and right kidney were observed in three or four species. This study demonstrates that size, sex, and spatial associations may have

  9. Characterizing ontogenetic habitat shifts in marine fishes: advancing nascent methods for marine spatial management.

    Science.gov (United States)

    Galaiduk, Ronen; Radford, Ben T; Saunders, Benjamin J; Newman, Stephen J; Harvey, Euan S

    2017-09-01

    Niche requirements and habitat resource partitioning by conspecific fishes of different sizes are significant knowledge gaps in the species distribution modelling domain. Management actions and operations are typically concentrated on static habitats, or specific areas of interest, without considering movement patterns of species associated with ontogenetic shifts in habitat usage. Generalized additive models were used to model the body-length-habitat relationships of six fish species. These models were used to identify subsets of environmental parameters that drive and explain the continuous length-habitat relationships for each of the study species, which vary in their degree of ecological and/or commercial importance. Continuous predictive maps of the length distributions for each of the six study species across approximately 200 km(2) of the study area were created from these models. The spatial patterns in habitat partitioning by individuals of different body lengths for all six study species provide strong evidence for ontogenetic shifts. This highlights the importance of considering ontogenetic processes for marine spatial management. Importantly, predictive hotspot maps were created that identify potential areas that accumulate individuals of similar life stages of multiple species (e.g., multispecies nursery areas). In circumstances where limited resources are available for monitoring and management of fish resources, predictive modelling is a valuable tool for studying previously overlooked processes such as ontogenetic habitat shifts. Predictive modelling provides crucial information that elucidates spatial patterns in community composition across mosaics of benthic habitats. This novel technique can contribute to the spatial management of coastal fish and fisheries by identifying areas that are important for different life history stages of multiple fish species. © 2017 by the Ecological Society of America.

  10. Alternative stable states generated by ontogenetic niche shift in the presence of multiple resource use.

    Directory of Open Access Journals (Sweden)

    Takefumi Nakazawa

    Full Text Available It has been suggested that when juveniles and adults use different resources or habitats, alternative stable states (ASS may exist in systems coupled by an ontogenetic niche shift. However, mainly the simplest system, i.e., the one-consumer-two-resource system, has been studied previously, and little is known about the development of ASS existing in more complex systems. Here, I theoretically investigated the development of ASS caused by an ontogenetic niche shift in the presence of multiple resource use. I considered three independent scenarios; (i additional resources, (ii multiple habitats, and (iii interstage resource sharing. The model analyses illustrate that relative balance between the total resource availability in the juvenile and adult habitats is crucial for the development of ASS. This balance is determined by factors such as local habitat productivity, subsidy inputs, colonization area, and foraging mobility. Furthermore, it is also shown that interstage resource sharing generally suppresses ASS. These results suggest that the anthropogenic impacts of habitat modifications (e.g., fragmentation and destruction or interaction modifications (e.g., changes in ontogeny and foraging behavior propagate through space and may cause or prevent regime shifts in the regional community structure.

  11. Ontogenetic shift in response to prey-derived chemical cues in prairie rattlesnakes Crotalus viridis viridis

    Institute of Scientific and Technical Information of China (English)

    Anthony J.SAVIOLA; David CHISZAR; Stephen P.MACKESSY

    2012-01-01

    Snakes often have specialized diets that undergo a shift from one prey type to another depending on the life stage of the snake.Crotalus viridis viridis (prairie rattlesnake) takes different prey at different life stages,and neonates typically prey on ectotherms,while adults feed almost entirely on small endotherms.We hypothesized that elevated rates of tongue flicking to chemical stimuli should correlate with particular prey consumed,and that this response shifts from one prey type to another as individuals age.To examine if an ontogenetic shift in response to chemical cues occurred,we recorded the rate of tongue flicking for 25 neonate,20 subadult,and 20 adult (average SVL=280.9,552,789.5 mm,respectively) wild-caught C.v.viridis to chemical stimuli presented on a cotton-tipped applicator; water-soluble cues from two ectotherms (prairie lizard,Sceloporus undulatus,and house gecko,Hemidactylusfrenatus),two endotherms (deer mouse,Peromyscus maniculatus and lab mouse,Mus musculus),and water controls were used.Neonates tongue flicked significantly more to chemical cues of their common prey,S.undulatus,than to all other chemical cues; however,the response to this lizard's chemical cues decreased in adult rattlesnakes.Subadults tongue flicked with a higher rate of tongue flicking to both S.undulatus and P.maniculatus than to all other treatments,and adults tongue flicked significantly more to P.maniculatus than to all other chemical cues.In addition,all three sub-classes demonstrated a greater response for natural prey chemical cues over chemical stimuli of prey not encountered in the wild (M.musculus and H.frenatus).This shift in chemosensory response correlated with the previously described ontogenetic shifts in C.v.viridis diet.Because many vipers show a similar ontogenetic shift in diet and venom composition,we suggest that this shift in prey cue discrimination is likely a general phenomenon among viperid snakes.

  12. Ontogenetic Shifts in Brain Organization in the Bluespotted Stingray Neotrygon kuhlii (Chondrichthyes: Dasyatidae).

    Science.gov (United States)

    Lisney, Thomas J; Yopak, Kara E; Camilieri-Asch, Victoria; Collin, Shaun P

    2017-02-28

    Fishes exhibit lifelong neurogenesis and continual brain growth. One consequence of this continual growth is that the nervous system has the potential to respond with enhanced plasticity to changes in ecological conditions that occur during ontogeny. The life histories of many teleost fishes are composed of a series of distinct stages that are characterized by shifts in diet, habitat, and behavior. In many cases, these shifts correlate with changes in overall brain growth and brain organization, possibly reflecting the relative importance of different senses and locomotor performance imposed by the new ecological niches they encounter throughout life. Chondrichthyan (cartilaginous) fishes also undergo ontogenetic shifts in habitat, movement patterns, diet, and behavior, but very little is known about any corresponding shifts in the size and organization of their brains. Here, we investigated postparturition ontogenetic changes in brain-body size scaling, the allometric scaling of seven major brain areas (olfactory bulbs, telencephalon, diencephalon, optic tectum, tegmentum, cerebellum, and medulla oblongata) relative to the rest of the brain, and cerebellar foliation in a chondrichthyan, i.e., the bluespotted stingray Neotrygon kuhlii. We also investigated the unusual morphological asymmetry of the cerebellum in this and other batoids. As in teleosts, the brain continues to grow throughout life, with a period of rapid initial growth relative to body size, before slowing considerably at the onset of sexual maturity. The olfactory bulbs and the cerebellum scale with positive allometry relative to the rest of the brain, whereas the other five brain areas scale with varying degrees of negative allometry. None of the major brain areas showed the stage-specific differences in rates of growth often found in teleosts. Cerebellar foliation also increases at a faster rate than overall brain growth. We speculate that changes in the olfactory bulbs and cerebellum could reflect

  13. Ontogenetic shifts in morphology and resource use of cisco Coregonus artedi.

    Science.gov (United States)

    Muir, A M; Vecsei, P; Pratt, T C; Krueger, C C; Power, M; Reist, J D

    2013-02-01

    Two previously described lacustrine cisco Coregonus spp. morphs [i.e. a small (cisco Coregonus artedi. Geometric body shape did not differ between the two size classes nor could they be differentiated by 24 size-corrected linear measurements, indicating that the two groups had similar phenotypes. Strong, positive correlations between all linear characters and geometric centroid size (a composite variable of fish body length, mass and age) suggested that body morphology changed with age as fish grew. Total gillraker number (N(GR)) increased with L(F) according to: N(GR) = 36.3 + 0.034L(F). Differences in gillraker number and phenotype with age and size were explained by shifts in habitat and trophic resource use. Relative abundance within 0-30, 30-60, 60-90 and >90 m depth strata differed between size classes suggesting that morphology changed when fish shifted their habitat as they grew older. Large C. artedi had lower δ(13)C and slightly higher δ(15)N, indicating greater reliance on pelagic prey resources (i.e. more or larger zooplankton, such as Mysis spp.), compared to small C. artedi, which relied slightly more on benthic prey. Gillraker shape and number have always been used as key diagnostic characters in coregonine taxonomy; based on the findings presented here, ontogenetic shifts should be accounted for in resulting classifications.

  14. Ontogenetic shifts in plant-plant interactions in a rare cycad within angiosperm communities.

    Science.gov (United States)

    Álvarez-Yépiz, Juan C; Búrquez, Alberto; Dovčiak, Martin

    2014-06-01

    Gymnosperms and angiosperms can co-occur within the same habitats but key plant traits are thought to give angiosperms an evolutionary competitive advantage in many ecological settings. We studied ontogenetic changes in competitive and facilitative interactions between a rare gymnosperm (Dioon sonorense, our target species) and different plant and abiotic neighbours (conspecific-cycads, heterospecific-angiosperms, or abiotic-rocks) from 2007 to 2010 in an arid environment of northwestern Mexico. We monitored survival and growth of seedlings, juveniles, and adults of the cycad Dioon sonorense to evaluate how cycad survival and relative height growth rate (RHGR) responded to intra- and interspecific competition, canopy openness, and nearest neighbour. We tested spatial associations among D. sonorense life stages and angiosperm species and measured ontogenetic shifts in cycad shade tolerance. Canopy openness decreased cycad survival while intraspecific competition decreased survival and RHGR during early ontogeny. Seedling survival was higher in association with rocks and heterospecific neighbours where intraspecific competition was lower. Shade tolerance decreased with cycad ontogeny reflecting the spatial association of advanced stages with more open canopies. Interspecific facilitation during early ontogeny of our target species may promote its persistence in spite of increasing interspecific competition in later stages. We provide empirical support to the long-standing assumption that marginal rocky habitats serve as refugia from angiosperm competition for slow-growing gymnosperms such as cycads. The lack of knowledge of plant-plant interactions in rare or endangered species may hinder developing efficient conservation strategies (e.g. managing for sustained canopy cover), especially under the ongoing land use and climatic changes.

  15. Evolution of ontogenetic dietary shifts and associated gut features in prickleback fishes (Teleostei: Stichaeidae).

    Science.gov (United States)

    German, D P; Gawlicka, A K; Horn, M H

    2014-02-01

    We tested the hypothesis that an ontogenetic dietary shift from carnivory to herbivory or omnivory, and concomitant changes in the gut facilitating digestion of algae, are synapomorphies of the tribes Xiphisterini and Esselenichthyini in the family Stichaeidae (pricklebacks). Previous investigations have revealed that two xiphisterine pricklebacks-Xiphister mucosus and Xiphister atropurpureus-become herbivorous or omnivorous, respectively, as their bodies grow larger, and that their guts show related changes in length and function. In this study we found that, with increase in size, the basal member of the Xiphisterini, Phytichthys chirus, showed an increased proportion of algae in its diet, increased activity of α-amylase and decreased activity of aminopeptidase, all of which support the synapomorphy hypothesis. Cebidichthys violaceus, a herbivore in the Esselenichthyini, shows similar ontogenetic changes in diet and digestive tract length and physiology, but these features were not observed in two derived carnivores, Dictyosoma burgeri and Dictyosoma rubrimaculatum, within the clade. These results suggest that herbivory is isolated to C. violaceus within the Esselenichthyini. Allometric relationships of gut length as a function of body size generally follow diet within the Xiphisterini and Esselenichthyini, with herbivores having the longest guts, which become disproportionately longer than body size as the fishes grow, omnivores intermediate gut lengths, and carnivores the shortest. A carnivore from an adjacent clade, Anoplarchus purpurescens, had the shortest gut, which did not change in length relative to body length as the fish grew. Overall, our results clarify the patterns of dietary evolution within the Stichaeidae and lay the foundation for more detailed studies of dietary and digestive specialization in fishes in the family.

  16. A Parent-Offspring Trade-Off Limits the Evolution of an Ontogenetic Niche Shift.

    Science.gov (United States)

    Ten Brink, Hanna; de Roos, André M

    2017-07-01

    Many free-living animal species, including the majority of fish, insects, and amphibians, change their food and habitat during their life. Even though these ontogenetic changes in niche are common, it is not well understood which ecological conditions have favored the evolution of these shifts. Using an adaptive dynamics approach, we show that it is evolutionarily advantageous to switch to an alternative food source in the course of ontogeny when this results in a higher intake rate for the switching consumers. Individuals are, however, not able to specialize on this new food source when this negatively affects the performance early in life on the original food source. Selection on these early life stages is so strong that in species with a complete diet shift, evolution results in large juveniles and adults that are maladapted to the alternative food source while their offspring are specialized on the original food source when young. These outcomes suggest strong selection to decouple the different life stages, such that they can maximize their performance on different food sources independently from each other. Metamorphosis could be a way to decouple the different life stages and therefore evolve in species that feed on multiple food sources during their life.

  17. Coexistence of predator and prey in intraguild predation systems with ontogenetic niche shifts.

    Science.gov (United States)

    Hin, Vincent; Schellekens, Tim; Persson, Lennart; de Roos, André M

    2011-12-01

    In basic intraguild predation (IGP) systems, predators and prey also compete for a shared resource. Theory predicts that persistence of these systems is possible when intraguild prey is superior in competition and productivity is not too high. IGP often results from ontogenetic niche shifts, in which the diet of intraguild predators changes as a result of growth in body size (life-history omnivory). As a juvenile, a life-history omnivore competes with the species that becomes its prey later in life. Competition can hence limit growth of young predators, while adult predators can suppress consumers and therewith neutralize negative effects of competition. We formulate and analyze a stage-structured model that captures both basic IGP and life-history omnivory. The model predicts increasing coexistence of predators and consumers when resource use of stage-structured predators becomes more stage specific. This coexistence depends on adult predators requiring consumer biomass for reproduction and is less likely when consumers outcompete juvenile predators, in contrast to basic IGP. Therefore, coexistence occurs when predation structures the community and competition is negligible. Consequently, equilibrium patterns over productivity resemble those of three-species food chains. Life-history omnivory thus provides a mechanism that allows intraguild predators and prey to coexist over a wide range of resource productivity.

  18. Retrospective characterization of ontogenetic shifts in killer whale diets via δ13C and δ15N analysis of teeth

    Science.gov (United States)

    Newsome, Seth D.; Etnier, Michael A.; Monson, Daniel H.; Fogel, Marilyn L.

    2009-01-01

    Metabolically inert, accretionary structures such as the dentin growth layers in teeth provide a life history record of individual diet with near-annual resolution. We constructed ontogenetic δ13C and δ15N profiles by analyzing tooth dentin growth layers from 13 individual killer whales Orcinus orca collected in the eastern northeast Pacific Ocean between 1961 and 2003. The individuals sampled were 6 to 52 yr old, representing 2 ecotypes—resident and transient—collected across ~25° of latitude. The average isotopic values of transient individuals (n = 10) are consistent with a reliance on mammalian prey, while the average isotopic values of residents (n = 3) are consistent with piscivory. Regardless of ecotype, most individuals show a decrease in δ15N values of ~2.5‰ through the first 3 yr of life, roughly equivalent to a decrease of one trophic level. We interpret this as evidence of gradual weaning, after which, ontogenetic shifts in isotopic values are highly variable. A few individuals (n = 2) maintained relatively stable δ15N and δ13C values throughout the remainder of their lives, whereas δ15N values of most (n = 11) increased by ~1.5‰, suggestive of an ontogenetic increase in trophic level. Significant differences in mean δ13C and δ15N values among transients collected off California suggest that individuality in prey preferences may be prevalent within this ecotype. Our approach provides retrospective individual life history and dietary information that cannot be obtained through traditional field observations of free-ranging and elusive species such as killer whales, including unique historic ecological information that pre-dates modern studies. By providing insights into individual diet composition, stable isotope analysis of teeth and/or bones may be the only means of evaluating a number of hypothesized historical dietary shifts in killer whales of the northeast Pacific Ocean

  19. Integrating ontogenetic shift, growth and mortality to determine a species' ecological role from isotopic signatures.

    Directory of Open Access Journals (Sweden)

    Nelson F Fontoura

    Full Text Available Understanding species linkages and energy transfer is a basic goal underlying any attempt at ecosystem analysis. Although the first food-web studies were based on gut contents of captured specimens, the assessment of stable isotopes, mainly δ13C and δ15N, has become a standard methodology for wide-range analyses in the last 30 years. Stable isotopes provide information on the trophic level of species, food-web length, and origin of organic matter ingested by consumers. In this study, we analyzed the ontogenetic variability of δ13C and δ15N obtained from samples of three Neotropical fish species: silver sardine (Lycengraulis grossidens, n=46, white lambari (Cyanocharax alburnus, n= 26, and the red-tail lambari (Astyanax fasciatus, n=23 in Pinguela Lagoon, southern Brazil. We developed a new metric, called the Weighted Isotopic Signature (φ 15N or φ 13C, ‰, that incorporates ontogenetic variability, body growth, and natural mortality into a single number.

  20. Light-shift-induced photonic nonlinearities

    Energy Technology Data Exchange (ETDEWEB)

    Brandao, F G S L; Hartmann, M J; Plenio, M B [Institute for Mathematical Sciences, Imperial College London, 53 Exhibition Road, SW7 2PE (United Kingdom)], E-mail: fernando@brandao@imperial.ac.uk

    2008-04-15

    We propose a new method to produce self- and cross-Kerr photonic nonlinearities, using light-induced Stark shifts due to the interaction of a cavity mode with atoms. The proposed experimental set-up is simpler than in previous approaches, while the strength of the nonlinearity obtained with a single atom is the same as in the setting based on electromagnetically induced transparency. Furthermore our scheme can be applied to engineer effective photonic nonlinear interactions whose strength increases with the number of atoms coupled to the cavity mode, leading to photon-photon interactions several orders of magnitude larger than previously considered possible.

  1. The tooth, the whole tooth and nothing but the tooth: tooth shape and ontogenetic shift dynamics in the white shark Carcharodon carcharias.

    Science.gov (United States)

    French, G C A; Stürup, M; Rizzuto, S; van Wyk, J H; Edwards, D; Dolan, R W; Wintner, S P; Towner, A V; Hughes, W O H

    2017-10-01

    Results from this study of the white shark Carcharodon carcharias include measurements obtained using a novel photographic method that reveal significant differences between the sexes in the relationship between tooth cuspidity and shark total length, and a novel ontogenetic change in male tooth shape. Males exhibit broader upper first teeth and increased distal inclination of upper third teeth with increasing length, while females do not present a consistent morphological change. Substantial individual variation, with implications for pace of life syndrome, was present in males and tooth polymorphism was suggested in females. Sexual differences and individual variation may play major roles in ontogenetic changes in tooth morphology in C. carcharias, with potential implications for their foraging biology. Such individual and sexual differences should be included in studies of ontogenetic shift dynamics in other species and systems. © 2017 The Fisheries Society of the British Isles.

  2. Characterization of the juvenile green turtle (Chelonia mydas) microbiome throughout an ontogenetic shift from pelagic to neritic habitats.

    Science.gov (United States)

    Price, James T; Paladino, Frank V; Lamont, Margaret M; Witherington, Blair E; Bates, Scott T; Soule, Tanya

    2017-01-01

    The gut microbiome of herbivorous animals consists of organisms that efficiently digest the structural carbohydrates of ingested plant material. Green turtles (Chelonia mydas) provide an interesting model of change in these microbial communities because they undergo a pronounced shift from a surface-pelagic distribution and omnivorous diet to a neritic distribution and herbivorous diet. As an alternative to direct sampling of the gut, we investigated the cloacal microbiomes of juvenile green turtles before and after recruitment to neritic waters to observe any changes in their microbial community structure. Cloacal swabs were taken from individual turtles for analysis of the 16S rRNA gene sequences using Illumina sequencing. One fecal sample was also obtained, allowing for a preliminary comparison with the bacterial community of the cloaca. We found significant variation in the juvenile green turtle bacterial communities between pelagic and neritic habitats, suggesting that environmental and dietary factors support different bacterial communities in green turtles from these habitats. This is the first study to characterize the cloacal microbiome of green turtles in the context of their ontogenetic shifts, which could provide valuable insight into the origins of their gut bacteria and how the microbial community supports their shift to herbivory.

  3. Are ontogenetic shifts in diet linked to shifts in feeding mechanics? Scaling of the feeding apparatus in the banded watersnake Nerodia fasciata.

    Science.gov (United States)

    Vincent, Shawn E; Moon, Brad R; Herrel, Anthony; Kley, Nathan J

    2007-06-01

    The effects of size on animal behaviour, ecology, and physiology are widespread. Theoretical models have been developed to predict how animal form, function, and performance should change with increasing size. Yet, numerous animals undergo dramatic shifts in ecology (e.g. habitat use, diet) that may directly influence the functioning and presumably the scaling of the musculoskeletal system. For example, previous studies have shown that banded watersnakes (Nerodia fasciata) switch from fish prey as juveniles to frog prey as adults, and that fish and frogs represent functionally distinct prey types to watersnakes. We therefore tested whether this ontogenetic shift in diet was coupled to changes in the scaling patterns of the cranial musculoskeletal system in an ontogenetic size series (70-600 mm snout-vent length) of banded watersnakes. We found that all cranial bones and gape size exhibited significant negative allometry, whereas the muscle physiological cross-sectional area (pCSAs) scaled either isometrically or with positive allometry against snout-vent length. By contrast, we found that gape size, most cranial bones, and muscle pCSAs exhibited highly significant positive allometry against head length. Furthermore, the mechanical advantage of the jaw-closing lever system remained constant over ontogeny. Overall, these cranial allometries should enable watersnakes to meet the functional requirements of switching from fusiform fish to bulky frog prey. However, recent studies have reported highly similar allometries in a wide diversity of vertebrate taxa, suggesting that positive allometry within the cranial musculoskeletal system may actually be a general characteristic of vertebrates.

  4. Ontogenetic shifts in male mating preference and morph-specific polyandry in a female colour polymorphic insect.

    Science.gov (United States)

    Sánchez-Guillén, Rosa Ana; Hammers, Martijn; Hansson, Bengt; Van Gossum, Hans; Cordero-Rivera, Adolfo; Galicia Mendoza, Dalia Ivette; Wellenreuther, Maren

    2013-06-06

    Sexual conflict over mating rates may favour the origin and maintenance of phenotypes with contrasting reproductive strategies. The damselfly Ischnura elegans is characterised by a female colour polymorphism that consists of one androchrome and two gynochrome female morphs. Previous studies have shown that the polymorphism is genetic and to a high extent maintained by negative frequency-dependent mating success that varies temporally and spatially. However, the role of learning in male mating preferences has received little attention. We used molecular markers to investigate differences in polyandry between female morphs. In addition, we experimentally investigated innate male mating preferences and experience-dependent shifts in male mating preferences for female morphs. Field and molecular data show that androchrome females were less polyandrous than gynochrome females. Interestingly, we found that naïve males showed significantly higher sexual preferences to androchrome than to gynochrome females in experimental trials. In contrast, experienced males showed no preference for androchrome females. The ontogenetic change in male mate preferences occurs most likely because of learned mate recognition after experience with females, which in this case does not result in a preference for one of the morphs, but rather in the loss of an innate preference for androchrome females.

  5. Ontogenetic niche shifts in dinosaurs influenced size, diversity and extinction in terrestrial vertebrates.

    Science.gov (United States)

    Codron, Daryl; Carbone, Chris; Müller, Dennis W H; Clauss, Marcus

    2012-08-23

    Given the physiological limits to egg size, large-bodied non-avian dinosaurs experienced some of the most extreme shifts in size during postnatal ontogeny found in terrestrial vertebrate systems. In contrast, mammals--the other dominant vertebrate group since the Mesozoic--have less complex ontogenies. Here, we develop a model that quantifies the impact of size-specific interspecies competition on abundances of differently sized dinosaurs and mammals, taking into account the extended niche breadth realized during ontogeny among large oviparous species. Our model predicts low diversity at intermediate size classes (between approx. 1 and 1000 kg), consistent with observed diversity distributions of dinosaurs, and of Mesozoic land vertebrates in general. It also provides a mechanism--based on an understanding of different ecological and evolutionary constraints across vertebrate groups--that explains how mammals and birds, but not dinosaurs, were able to persist beyond the Cretaceous-Tertiary (K-T) boundary, and how post-K-T mammals were able to diversify into larger size categories.

  6. Parasites as biological tags to track an ontogenetic shift in the feeding behaviour of Gadus morhua off West and East Greenland.

    Science.gov (United States)

    Münster, Julian; Klimpel, Sven; Fock, Heino O; MacKenzie, Ken; Kuhn, Thomas

    2015-07-01

    Parasites, being an integral part of every ecosystem and trophically transmitted along the food webs, can provide detailed insights into the structure of food webs and can close the information gap between short-term stomach content analyses and long-term fish otolith analyses. They are useful for tracking ontogenetic shifts in the host's diet, the occurrence of specific organisms or migratory behaviour of their hosts, even in inaccessible environments. In the present study, stomach content analyses and parasitological examinations were performed on 70 Atlantic cod Gadus morhua, one of the most important high-level predators of small fish in the North Atlantic, caught during one research vessel cruise from West and East Greenlandic waters. Analyses revealed significant differences in fish size with higher values for East Greenland (average total length (TL) of 50.5 cm) compared to West Greenland (average TL of 33.3 cm). Clear differences were also present in prey and parasite composition. Crustacea was the main food source for all fish (IRI = 10082.70), while the importance of teleosts increased with fish size. With a prevalence of 85 % in West Greenland and 100 % in East Greenland, Nematoda were the most abundant parasite group. The results indicate an ontogenetic shift in the diet, which are discussed in the context of the common distribution theory, stock dynamics and migratory behaviour.

  7. Integrated "omics" profiling indicates that miRNAs are modulators of the ontogenetic venom composition shift in the Central American rattlesnake, Crotalus simus simus.

    Science.gov (United States)

    Durban, Jordi; Pérez, Alicia; Sanz, Libia; Gómez, Aarón; Bonilla, Fabián; Rodríguez, Santos; Chacón, Danilo; Sasa, Mahmood; Angulo, Yamileth; Gutiérrez, José M; Calvete, Juan J

    2013-04-10

    Understanding the processes that drive the evolution of snake venom is a topic of great research interest in molecular and evolutionary toxinology. Recent studies suggest that ontogenetic changes in venom composition are genetically controlled rather than environmentally induced. However, the molecular mechanisms underlying these changes remain elusive. Here we have explored the basis and level of regulation of the ontogenetic shift in the venom composition of the Central American rattlesnake, Crotalus s. simus using a combined proteomics and transcriptomics approach. Proteomic analysis showed that the ontogenetic shift in the venom composition of C. s. simus is essentially characterized by a gradual reduction in the expression of serine proteinases and PLA2 molecules, particularly crotoxin, a β-neurotoxic heterodimeric PLA2, concominantly with an increment of PI and PIII metalloproteinases at age 9-18 months. Comparison of the transcriptional activity of the venom glands of neonate and adult C. s. simus specimens indicated that their transcriptomes exhibit indistinguisable toxin family profiles, suggesting that the elusive mechanism by which shared transcriptomes generate divergent venom phenotypes may operate post-transcriptionally. Specifically, miRNAs with frequency count of 1000 or greater exhibited an uneven distribution between the newborn and adult datasets. Of note, 590 copies of a miRNA targeting crotoxin B-subunit was exclusively found in the transcriptome of the adult snake, whereas 1185 copies of a miRNA complementary to a PIII-SVMP mRNA was uniquely present in the newborn dataset. These results support the view that age-dependent changes in the concentration of miRNA modulating the transition from a crotoxin-rich to a SVMP-rich venom from birth through adulthood can potentially explain what is observed in the proteomic analysis of the ontogenetic changes in the venom composition of C. s. simus. Existing snake venom toxins are the result of early

  8. Integrated “omics” profiling indicates that miRNAs are modulators of the ontogenetic venom composition shift in the Central American rattlesnake, Crotalus simus simus

    Science.gov (United States)

    2013-01-01

    Background Understanding the processes that drive the evolution of snake venom is a topic of great research interest in molecular and evolutionary toxinology. Recent studies suggest that ontogenetic changes in venom composition are genetically controlled rather than environmentally induced. However, the molecular mechanisms underlying these changes remain elusive. Here we have explored the basis and level of regulation of the ontogenetic shift in the venom composition of the Central American rattlesnake, Crotalus s. simus using a combined proteomics and transcriptomics approach. Results Proteomic analysis showed that the ontogenetic shift in the venom composition of C. s. simus is essentially characterized by a gradual reduction in the expression of serine proteinases and PLA2 molecules, particularly crotoxin, a β-neurotoxic heterodimeric PLA2, concominantly with an increment of PI and PIII metalloproteinases at age 9–18 months. Comparison of the transcriptional activity of the venom glands of neonate and adult C. s. simus specimens indicated that their transcriptomes exhibit indistinguisable toxin family profiles, suggesting that the elusive mechanism by which shared transcriptomes generate divergent venom phenotypes may operate post-transcriptionally. Specifically, miRNAs with frequency count of 1000 or greater exhibited an uneven distribution between the newborn and adult datasets. Of note, 590 copies of a miRNA targeting crotoxin B-subunit was exclusively found in the transcriptome of the adult snake, whereas 1185 copies of a miRNA complementary to a PIII-SVMP mRNA was uniquely present in the newborn dataset. These results support the view that age-dependent changes in the concentration of miRNA modulating the transition from a crotoxin-rich to a SVMP-rich venom from birth through adulhood can potentially explain what is observed in the proteomic analysis of the ontogenetic changes in the venom composition of C. s. simus. Conclusions Existing snake venom

  9. Ontogenetic shifts in fishes between vegetated and unvegetated tidepools: assessing the effect of physical structure on fish habitat selection.

    Science.gov (United States)

    Oliveira, R R de S; Macieira, R M; Giarrizzo, T

    2016-07-01

    The aim of this study of tidepool fishes was analyse variation in their use of intertidal habitats (rocky shore, mangrove and salt marsh). Specimens were collected during wet and dry periods from 18 tidepools in the three habitats. A total of 7690 specimens, belonging to 19 families and 30 species, was captured. The fish assemblage in rocky shore pools was clearly distinct from that of vegetated habitats (mangrove and salt marshes). The rocky shore fauna was dominated by permanent resident species, whereas pools in mangrove and salt marsh habitats were inhabited primarily by opportunistic and transient species. Habitat segregation by ontogenetic stage (e.g. smaller individuals in mangroves, intermediate size classes in salt marsh and sub-adults/adults on rocky shores) indicates age-related migration in response to the physical structure of these habitats and to the natural history of each fish species. These findings are important for the development of effective conservation and management plans for intertidal fishes.

  10. The reaction of European lobster larvae (Homarus gammarus) to different quality food: effects of ontogenetic shifts and pre-feeding history.

    Science.gov (United States)

    Schoo, Katherina L; Aberle, Nicole; Malzahn, Arne M; Schmalenbach, Isabel; Boersma, Maarten

    2014-02-01

    Young larval stages of many organisms represent bottlenecks in the life-history of many species. The high mortality commonly observed in, for example, decapod larvae has often been linked to poor nutrition, with most studies focussing on food quantity. Here, we focus instead on the effects of quality and have investigated its effects on the nutritional condition of lobster larvae. We established a tri-trophic food chain consisting of the cryptophyte Rhodomonas salina, the calanoid copepod Acartia tonsa and larvae of the European lobster Homarus gammarus. In a set of experiments, we manipulated the C:N:P stoichiometry of the primary producers, and accordingly those of the primary consumer. In a first experiment, R. salina was grown under N- and P-limitation and the nutrient content of the algae was manipulated by addition of the limiting nutrient to create a food quality gradient. In a second experiment, the effect on lobster larvae of long- and short-term exposure to food of varying quality during ontogenetic development was investigated. The condition of the lobster larvae was negatively affected even by subtle N- and P-nutrient limitations of the algae. Furthermore, younger lobster larvae were more vulnerable to nutrient limitation than older ones, suggesting an ontogenetic shift in the capacity of lobster larvae to cope with low quality food. The results presented here might have substantial consequences for the survival of lobster larvae in the field, as, in the light of future climate change and re-oligotrophication of the North Sea, lobster larvae might face marked changes in temperature and nutrient conditions, thus significantly altering their condition and growth.

  11. The Gouy phase shift in nonlinear interactions of waves

    Science.gov (United States)

    Lastzka, Nico; Schnabel, Roman

    2007-06-01

    We theoretically analyze the influence of the Gouy phase shift on the nonlinear interaction between waves of different frequencies. We focus on χ(2)interaction of optical fields, e.g. through birefringent crystals, and show that focussing, stronger than suggested by the Boyd-Kleinman factor, can further improve nonlinear processes. An increased value of 3.32 for the optimal focussing parameter for a single pass process is found. The new value builds on the compensation of the Gouy phase shift by a spatially varying, instead constant, wave vector phase mismatch. We analyze the single-ended, singly resonant standing wave nonlinear cavity and show that in this case the Gouy phase shift leads to an additional phase during backreflection. Our numerical simulations may explain ill-understood experimental observations in such devices.

  12. Time shift of pulses due to dispersion slope and nonlinearity

    Energy Technology Data Exchange (ETDEWEB)

    Marcuse, D.; Menyuk, C.R.; Holzloehner, R.

    1999-12-01

    The authors show that the time delay of optical pulses traveling in long fibers is influenced by the dispersion slope and the fiber nonlinearity. Consequently, one or more new pulses that are inserted by add-drop operations into a pulse train that has already traveled a long distance may shift relative to the old pulses. This time shift delays the initial pulses more than the newly inserted ones, so that the newly inserted pulses can leave their time frames, leading to errors.

  13. Ontogenetic shifts in the diet of plains hog-nosed snakes (Heterodon nasicus) revealed by stable isotope analysis.

    Science.gov (United States)

    Durso, Andrew M; Mullin, Stephen J

    2017-02-01

    Wild snake diets are difficult to study using traditional methods, but stable isotopes offer several advantages, including integrating dietary information over time, providing data from individuals that have not fed recently, and avoiding bias towards slowly-digesting prey items. We used stable isotope signatures of carbon and nitrogen from scale tissue, red blood cells, and blood plasma to assess the diet of wild plains hog-nosed snakes (Heterodon nasicus) in Illinois. We developed Bayesian mixing models which, taken together, predicted that H. nasicus shifted from a juvenile diet predominantly (31-63%) composed of six-lined racerunners (Aspidoscelis sexlineatus) and their eggs to an adult diet predominantly (44-56%) composed of eggs of the aquatic turtles Chrysemys picta and Chelydra serpentina, with a contribution from toads (Anaxyrus sp.; 6-27%) during their adolescent years. These results agreed with sparse data from gut contents. Combining traditional and isotopic techniques for studying the diets of wild snakes can increase the utility of both types of data.

  14. Nonlinear frequency shift of electrostatic waves in general collisionless plasma: unifying theory of fluid and kinetic nonlinearities

    CERN Document Server

    Liu, Chang

    2015-01-01

    The nonlinear frequency shift is derived in a transparent asymptotic form for intense Langmuir waves in general collisionless plasma. The formula describes both fluid and kinetic effects simultaneously. The fluid nonlinearity is expressed, for the ?first time, through the plasma dielectric function, and the kinetic nonlinearity accounts for both smooth distributions and trapped-particle beams. Various known limiting scalings are reproduced as special cases. The calculation avoids differential equations and can be extended straightforwardly to other nonlinear plasma waves.

  15. Goos-Hänchen shifts of Helmholtz solitons at nonlocal nonlinear interfaces

    Science.gov (United States)

    Zhiwei, Shi; Jing, Xue; Jilong, Chen; Yang, Li; Huagang, Li

    2015-02-01

    We address the nonlinear Goos-Hänchen shift of Helmholtz solitons at a nonlocal nonlinear interface between a Kerr medium and a nonlocal nonlinear medium. Based on the framework of the Helmholtz theory, we have demonstrated that the Goos-Hänchen shift depends on the angle of the incidence, the linear and nonlinear refractive index mismatch at the interface, the nonparaxial parameter and the degree of nonlocality. Interestingly, internal and external refraction can be introduced when the nonlinear refractive index mismatch is greater than a threshold value. The total reflection will occur when the degree of nonlocality exceeds a value.

  16. Generation of large fifth-order nonlinear phase shifts by use of lossless χ(2) cascaded nonlinear processes

    Institute of Scientific and Technical Information of China (English)

    XU Guang; QIAN Liejia; WANG Tao; FAN Dianyuan; LI Fuming

    2004-01-01

    It is shown that the cascaded fifth-order nonlinear phase shifts will increase with energy loss in the cascaded processes. Essentially different from the multi-photon absorption accompanied with inherent material nonlinearities, the loss of fundamental wave in a cascaded process is controllable and suppressible. By introducing difference frequencies generated from the reaction between the fundamental and its second harmonic after the cascaded processes, the fundamental wave can be free of energy loss, while the large cascaded fifth-order nonlinear phase shift is maintained.

  17. Stochastic ontogenetic growth model

    Science.gov (United States)

    West, B. J.; West, D.

    2012-02-01

    An ontogenetic growth model (OGM) for a thermodynamically closed system is generalized to satisfy both the first and second law of thermodynamics. The hypothesized stochastic ontogenetic growth model (SOGM) is shown to entail the interspecies allometry relation by explicitly averaging the basal metabolic rate and the total body mass over the steady-state probability density for the total body mass (TBM). This is the first derivation of the interspecies metabolic allometric relation from a dynamical model and the asymptotic steady-state distribution of the TBM is fit to data and shown to be inverse power law.

  18. Nonlinear Pulse Compression and Reshaping Using Cross-Phase Modulation in a Dispersion-Shifted Fiber

    Institute of Scientific and Technical Information of China (English)

    S.; W.; Chan; K.; K.; Chow; C.; Shu

    2003-01-01

    Nonlinear pulse compression has been demonstrated by cross-phase modulation in a dispersion-shifted fiber. The output is obtained from filtering of the broadened optical spectrum and a pulse width reduction from 61 to 28 ps is achieved.

  19. Cylindrical PVF2 film based fiber optic phase modulator - Phase shift nonlinearity and frequency response

    Science.gov (United States)

    Sudarshanam, V. S.; Claus, Richard O.

    1993-03-01

    A new cylindrical coil configuration for polyvinylidene flouride (PVF2) film based fiber optic phase modulator is studied for the frequency response and nonlinearity of phase shift at the resonance frequency. This configuration, hitherto unapproached for PVF2 film modulators, offers resonance at well defined, controllable and higher frequencies than possible for the flat-strip configuration. Two versions of this configuration are presented that differ strongly in both the resonance frequency and the phase shift nonlinearity coefficient.

  20. Intermittent Giant Goos-Hanchen shifts from Airy beams at nonlinear interfaces

    CERN Document Server

    Chamorro-Posada, Pedro; Aceves, Alejandro B; McDonald, Graham S

    2013-01-01

    We study the giant Goos-Hanchen shift obtained from an Airy beam impinging on a nonlinear interface. To avoid any angular restriction associated with the paraxial approximation, the analysis is based on the numerical solution of the nonlinear Helmholtz equation. We report the existence of non-standard intermittent and oscillatory regimes for the nonlinear Goos-Hanchen shifts which can be explained in terms of the competition between the critical coupling to a surface mode of the reflected component of the Airy beam and the soliton emission from the refracted beam component.

  1. Nonlinear pressure shifts of alkali-metal atoms in inert gases.

    Science.gov (United States)

    Gong, F; Jau, Y-Y; Happer, W

    2008-06-13

    Precise measurements show that the microwave resonance frequencies of ground-state Rb or Cs atoms have a nonlinear dependence on the pressure of the buffer gases Ar and Kr. No nonlinearities were observed in the gases He or N(2). These observations strongly suggest that the nonlinearities are due to the van der Waals molecules that form in Ar and Kr, but not in He or N(2). The nonlinear part of the shifts is largest in the pressure range of a few tens of torr, similar to the operating pressures of gas-cell atomic clocks. The observed shifts are very well described by a simple function, parametrized by the effective three-body formation rate of molecules and by the effective product of the collisionally limited lifetime times the shift of the hyperfine coupling coefficient in the molecule.

  2. Wavenumber shift due to nonlinear plasma and wave interaction

    Science.gov (United States)

    Gan, Chunyun; Xiang, Nong; Yu, Zhi; Yang, Youlei; Ou, Jing

    2016-06-01

    Wavenumber shift of the ion Bernstein wave has been observed in the particle-in-cell simulations when the input power of the injected wave is sufficiently large. It is demonstrated that the increase of the total kinetic energy of ions, including both the thermal energy related to the random thermal motion and the oscillation energy due to the coherent motion with the wave, gives rise to such change of the wavenumber. However, the velocity distribution function of the ions can approximately be fitted as a Maxwellian distribution function, and thus, the linear dispersion relation still holds, provided that the initial ion temperature is replaced by the effective temperature measured in the simulation.

  3. Tune shift induced by nonlinear resistive wall wake field of flat collimator

    CERN Document Server

    Zimmermann, Frank; Assmann, R W; Burkhardt, H; Caspers, Friedhelm; Gasior, M; Jones, R; Kroyer, T; Métral, E; Redaelli, S; Robert-Démolaize, G; Roncarolo, F; Rumolo, Giovanni; Steinhagen, Ralph J; Wenninger, J

    2006-01-01

    We present formulae for the coherent and incoherent tune shifts due to the nonlinear resistive wall wake field for a single beam traveling between two parallel plates. In particular, we demonstrate that the nonlinear terms of the resistive-wall wake field become important if the gap between the plates is comparable to the transverse rms beam size. We also compare the theoretically predicted tune shift as a function of gap size with measurements for an LHC prototype graphite collimator in the CERN SPS and with simulations.

  4. Dynamic detection of a single bacterium: nonlinear rotation rate shifts of driven magnetic microsphere stages

    CERN Document Server

    McNaughton, B H; Kopelman, R; Agayan, Rodney R.; Kopelman, Raoul; Naughton, Brandon H. Mc

    2006-01-01

    We report on a new technique which was used to detect single Escherichia coli that is based on the changes in the nonlinear rotation of a magnetic microsphere driven by an external magnetic field. The presence of one Escherichia Coli bacterium on the surface of a 2.0 micron magnetic microsphere caused an easily measurable change in the drag of the system and, therefore, in the nonlinear rotation rate. The straight-forward measurement uses standard microscopy techniques and the observed average shift in the nonlinear rotation rate changed by a factor of ~3.8.

  5. Widely varying giant Goos-Hänchen shifts from Airy beams at nonlinear interfaces.

    Science.gov (United States)

    Chamorro-Posada, Pedro; Sánchez-Curto, Julio; Aceves, Alejandro B; McDonald, Graham S

    2014-03-15

    We present a numerical study of the giant Goos-Hänchen shifts (GHSs) obtained from an Airy beam impinging on a nonlinear interface. To avoid any angular restriction associated with the paraxial approximation, the analysis is based on the nonlinear Helmholtz equation. We report the existence of nonstandard nonlinear GHSs displaying an extreme sensitivity to the input intensity and the existence of multiple critical values. These intermittent and oscillatory regimes can be explained in terms of competition between critical coupling to a surface mode and soliton emission from the refracted beam component and how this interplay varies with localization of the initial Airy beam.

  6. Effective supercontinuum generation by using highly nonlinear dispersion-shifted fiber incorporated with Si nanocrystals.

    Science.gov (United States)

    Jeong, Seongmook; Ju, Seongmin; Kim, Youngwoong; Watekar, Pramod R; Jeong, Hyejeong; Lee, Ho-Jae; Boo, Seongjae; Kim, Dug Young; Han, Won-Taek

    2012-01-01

    The dispersion-shifted fiber (DSF) incorporated with Si nanocrystals (Si-NCs) having highly nonlinear optical property was fabricated to investigate the effective supercontinuum generation characteristics by using the MCVD process and the drawing process. Optical nonlinearity was enhanced by incorporating Si nanocrystals in the core of the fiber and the refractive index profile of a dispersion-shifted fiber was employed to match its zero-dispersion wavelength to that of the commercially available pumping source for generating effective supercontinuum. The non-resonant nonlinear refractive index, n2, of the Si-NCs doped DSF measured by the cw-SPM method was measured to be 7.03 x 10(-20) [m2/W] and the coefficient of non-resonant nonlinearity, gamma, was 7.14 [W(-1) km(-1)]. To examine supercontinuum generation of the Si-NCs doped DSF, the femtosecond fiber laser with the pulse width of 150 fs (at 1560 nm) was launched into the fiber core. The output spectrum of the Si-NCs doped DSF was found to broaden from 1300 nm to wavelength well beyond 1700 nm, which can be attributed to the enhanced optical nonlinearity by Si-NCs embedded in the fiber core. The short wavelength of the supercontinuum spectrum in the Si-NCs doped DSF showed shift from 1352 nm to 1220 nm for the fiber length of 2.5 m and 200 m, respectively.

  7. Fluid nonlinear frequency shift of nonlinear ion acoustic waves in multi-ion species plasmas in small wave number region

    CERN Document Server

    Feng, Q S; Wang, Q; Zheng, C Y; Liu, Z J; Cao, L H; He, X T

    2016-01-01

    The properties of the nonlinear frequency shift (NFS) especially the fluid NFS from the harmonic generation of the ion-acoustic wave (IAW) in multi-ion species plasmas have been researched by Vlasov simulation. The pictures of the nonlinear frequency shift from harmonic generation and particles trapping are shown to explain the mechanism of NFS qualitatively. The theoretical model of the fluid NFS from harmonic generation in multi-ion species plasmas is given and the results of Vlasov simulation are consistent to the theoretical result of multi-ion species plasmas. When the wave number $k\\lambda_{De}$ is small, such as $k\\lambda_{De}=0.1$, the fluid NFS dominates in the total NFS and will reach as large as nearly $15\\%$ when the wave amplitude $|e\\phi/T_e|\\sim0.1$, which indicates that in the condition of small $k\\lambda_{De}$, the fluid NFS dominates in the saturation of stimulated Brillouin scattering especially when the nonlinear IAW amplitude is large.

  8. Which way will the circulation shift in a changing climate? Possible nonlinearity of extratropical cloud feedbacks

    Science.gov (United States)

    Tandon, Neil F.; Cane, Mark A.

    2017-06-01

    In a suite of idealized experiments with the Community Atmospheric Model version 3 coupled to a slab ocean, we show that the atmospheric circulation response to CO2 increase is sensitive to extratropical cloud feedback that is potentially nonlinear. Doubling CO2 produces a poleward shift of the Southern Hemisphere (SH) midlatitude jet that is driven primarily by cloud shortwave feedback and modulated by ice albedo feedback, in agreement with earlier studies. More surprisingly, for CO2 increases smaller than 25 %, the SH jet shifts equatorward. Nonlinearities are also apparent in the Northern Hemisphere, but with less zonal symmetry. Baroclinic instability theory and climate feedback analysis suggest that as the CO2 forcing amplitude is reduced, there is a transition from a regime in which cloud and circulation changes are largely decoupled to a regime in which they are highly coupled. In the dynamically coupled regime, there is an apparent cancellation between cloud feedback due to warming and cloud feedback due to the shifting jet, and this allows the ice albedo feedback to dominate in the high latitudes. The extent to which dynamical coupling effects exceed thermodynamic forcing effects is strongly influenced by cloud microphysics: an alternate model configuration with slightly increased cloud liquid (LIQ) produces poleward jet shifts regardless of the amplitude of CO2 forcing. Altering the cloud microphysics also produces substantial spread in the circulation response to CO2 doubling: the LIQ configuration produces a poleward SH jet shift approximately twice that produced under the default configuration. Analysis of large ensembles of the Canadian Earth System Model version 2 demonstrates that nonlinear, cloud-coupled jet shifts are also possible in comprehensive models. We still expect a poleward trend in SH jet latitude for timescales on which CO2 increases by more than 25 %. But on shorter timescales, our results give good reason to expect significant

  9. Goos-Haenchen shift and time delay in dispersive nonlinear media

    Energy Technology Data Exchange (ETDEWEB)

    Ilic, I. [Vinca Institute of Nuclear Sciences, University of Belgrade, P.O. Box 522, 11001 Belgrade (Serbia); Belicev, P.P., E-mail: petrab@vinca.r [Vinca Institute of Nuclear Sciences, University of Belgrade, P.O. Box 522, 11001 Belgrade (Serbia); Milanovic, V.; Radovanovic, J. [Faculty of Electrical Engineering, University of Belgrade, Bul. kralja Aleksandra 73, 11120 Belgrade (Serbia); Hadzievski, Lj. [Vinca Institute of Nuclear Sciences, University of Belgrade, P.O. Box 522, 11001 Belgrade (Serbia)

    2011-03-07

    We present an analysis of the influence of the Goos-Haenchen effect on tunneling times, group delay and dwell time, of electromagnetic waves propagating through an obstacle made of left-handed metamaterial embedded in a dielectric which exhibits saturable type of nonlinearity. The derived equations show that only the group delay, is affected by the Goos-Haenchen shift without any impact on the dwell time. Besides the reduction of the group delay, the most remarkable result is the possibility for total reduction of the Goos-Haenchen shift for finite incident angles. These phenomena are observable in the frequency region for which metamaterial exhibits negative index of refraction.

  10. Nonlinear modification of the laser noise power spectrum induced by a frequency-shifted optical feedback

    CERN Document Server

    Lacot, Eric; Girardeau, Vadim; Hugon, Olivier; Jacquin, Olivier

    2016-01-01

    In this article, we study the non-linear coupling between the stationary (i.e. the beating modulation signal) and transient (i.e. the laser quantum noise) dynamics of a laser subjected to frequency shifted optical feedback. We show how the noise power spectrum and more specifically the relaxation oscillation frequency of the laser are modified under different optical feedback condition. Specifically we study the influence of (i) the amount of light returning to the laser cavity and (ii) the initial detuning between the frequency shift and intrinsic relaxation frequency. The present work shows how the relaxation frequency is related to the strength of the beating signal and the shape of the noise power spectrum gives an image of the Transfer Modulation Function (i.e. of the amplification gain) of the nonlinear-laser dynamics.The theoretical predictions, confirmed by numerical resolutions, are in good agreements with the experimental data.

  11. Nonlinear frequency shift in Raman backscattering and its implications for plasma diagnostics

    Science.gov (United States)

    Kaganovich, D.; Hafizi, B.; Palastro, J. P.; Ting, A.; Helle, M. H.; Chen, Y.-H.; Jones, T. G.; Gordon, D. F.

    2016-12-01

    Raman backscattered radiation of intense laser pulses in plasmas is investigated for a wide range of intensities relevant to laser wakefield acceleration. The weakly nonlinear dispersion relation for Raman backscattering predicts an intensity and density dependent frequency shift that is opposite to that suggested by a simple relativistic consideration. This observation has been benchmarked against experimental results, providing a novel diagnostic for laser-plasma interactions.

  12. Nonlinear Frequency Shift in Raman Backscattering and its Implications for Plasma Diagnostics

    CERN Document Server

    Kaganovich, D; Palastro, J P; Ting, A; Helle, M H; Chen, Y -H; Jones, T G; Gordon, D F

    2016-01-01

    Raman backscattered radiation of intense laser pulses in plasma is investigated for a wide range of intensities relevant to laser wakefield acceleration. The weakly nonlinear dispersion relation for Raman backscattering predicts an intensity and density dependent frequency shift that is opposite to that suggested by a simple relativistic consideration. This observation has been benchmarked against experimental results, providing a novel diagnostic for laser-plasma interactions.

  13. Dynamic Nonlinear Focal Shift in Amplitude Modulated Moderately Focused Acoustic Beams

    CERN Document Server

    Jiménez, Noé; González-Salido, Nuria

    2016-01-01

    The phenomenon of the displacement of the position of the pressure, intensity and acoustic radiation force maxima along the axis of focused acoustic beams under increasing driving amplitudes (nonlinear focal shift) is studied for the case of a moderately focused beam excited with continuous and 25 kHz amplitude modulated signals, both in water and tissue. We prove that in amplitude modulated beams the linear and nonlinear propagation effects coexist in a semi-period of modulation, giving place to a complex dynamic behaviour, where the singular points of the beam (peak pressure, rarefaction, intensity and acoustic radiation force) locate at different points on axis as a function of time. These entire phenomena are explained in terms of harmonic generation and absorption during the propagation in a lossy nonlinear medium both, for a continuous and an amplitude modulated beam. One of the possible applications of the acoustic radiation force displacement is the generation of shear waves at different locations by ...

  14. Dynamical patterns and regime shifts in the nonlinear model of soil microorganisms growth

    Science.gov (United States)

    Zaitseva, Maria; Vladimirov, Artem; Winter, Anna-Marie; Vasilyeva, Nadezda

    2017-04-01

    Dynamical model of soil microorganisms growth and turnover is formulated as a system of nonlinear partial differential equations of reaction-diffusion type. We consider spatial distributions of concentrations of several substrates and microorganisms. Biochemical reactions are modelled by chemical kinetic equations. Transport is modelled by simple linear diffusion for all chemical substances, while for microorganisms we use different transport functions, e.g. some of them can actively move along gradient of substrate concentration, while others cannot move. We solve our model in two dimensions, starting from uniform state with small initial perturbations for various parameters and find parameter range, where small initial perturbations grow and evolve. We search for bifurcation points and critical regime shifts in our model and analyze time-space profile and phase portraits of these solutions approaching critical regime shifts in the system, exploring possibility to detect such shifts in advance. This work is supported by NordForsk, project #81513.

  15. Nonlinear focal shift beyond the geometrical focus in moderately focused acoustic beams.

    Science.gov (United States)

    Camarena, Francisco; Adrián-Martínez, Silvia; Jiménez, Noé; Sánchez-Morcillo, Víctor

    2013-08-01

    The phenomenon of the displacement of the position along the axis of the pressure, intensity, and radiation force maxima of focused acoustic beams under increasing driving voltages (nonlinear focal shift) is studied for the case of a moderately focused beam. The theoretical and experimental results show the existence of this shift along the axis when the initial pressure in the transducer increases until the acoustic field reaches the fully developed nonlinear regime of propagation. Experimental data show that at high amplitudes and for moderate focusing, the position of the on-axis pressure maximum and radiation force maximum can surpass the geometrical focal length. On the contrary, the on-axis pressure minimum approaches the transducer under increasing driving voltages, increasing the distance between the positive and negative peak pressure in the beam. These results are in agreement with numerical KZK model predictions and the existed data of other authors and can be explained according to the effect of self-refraction characteristic of the nonlinear regime of propagation.

  16. Cochannel and Adjacent-Channel Interference in Nonlinear Minimum-Shift-Keyed Satellite System

    Science.gov (United States)

    Yu, John

    1995-01-01

    The interference susceptibility of a serial-minimum-shift-keyed (SMSK) modulation system to an interfering signal transmitted through a satellite link with cascaded nonlinear elements was investigated through computer simulation. The satellite link evaluated in this study represented NASA's Advanced Communications Technology Satellite (ACTS) system. Specifically, nonlinear characteristics were used that had specified amplitude-modulation to amplitude-modulation and amplitude-modulation to phase-modulation transfer characteristics obtained from the actual ACTS hardware. Two measurement scenarios were analyzed: degradation of an MSK satellite link from cochannel interference and from adjacent-channel interference. Interference was evaluated in terms of the probability of bit error rate (BER) versus energy per bit over noise power density Eb/No.

  17. Non Stationary And Non Gaussian Character Of ENSO: The Role Of Climate Shifts And Nonlinearities

    Science.gov (United States)

    Boucharel, J.; Dewitte, B.; Garel, B.; Du Penhoat, Y.

    2008-12-01

    ENSO is the dominant climate mode of variability in the Pacific, having socio-economical impacts on surrounding regions. ENSO exhibits a significant modulation at decadal to interdecadal timescales which is associated to changes of its characteristics (onset, amplitude, frequency, propagation, and predictability). Among these characteristics, some of them are generally ignored in ENSO studies, such as its asymmetry and the deviation of its statistics from those of the Gaussian distribution. These properties could be related to the ability of the current generation of coupled models to predict ENSO and its modulation. Here, the non-Gaussian nature and asymmetry of ENSO is diagnosed from in situ data and the outputs of a variety of models (from intermediate complexity models to full physics coupled general circulation models) using robust statistical tools. In particular alpha-stable laws are used as theoretical background material to quantify the non-Gaussian character of ENSO time series. It is shown that the Alpha-stable character of ENSO may result from the presence of climate shifts inducing non stationnarity in the time series. Also, cool (warm) periods are associated with ENSO statistics having a larger (weaker) tendency towards Gaussianity and a weaker (larger) asymmetry. This supports the hypothesis of ENSO being rectified by changes in mean state through non-linear processes. The relationship between change in mean state and non-linearity is investigated both in the Zebiak and Cane model and the IPCC models, which indicate that the propensity of a model to reproduce extreme events is related to its tendency to emphasize the non-linear interaction between mean state and ENSO variability. More particularly, high statistical moments i.e. high order nonlinearities seem to be involved in the feedback between extreme events occurrence and mean state shift triggering through inverse energy cascade, emphasizing the ENSO multifractal nature.

  18. Nonlinear Goos-Haenchen shifts due to surface polariton resonance in Kretschmann configuration with a Kerr-type substrate

    Energy Technology Data Exchange (ETDEWEB)

    Xu Guoding, E-mail: guodingxu@163.co [Department of Physics, Suzhou University of Science and Technology, Suzhou 215009 (China); Zang Taocheng; Mao Hongmin; Pan Tao [Department of Physics, Suzhou University of Science and Technology, Suzhou 215009 (China)

    2010-07-26

    As the surface polaritons are excited in Kretschmann configuration with a Kerr-type substrate, the nonlinear Goos-Haenchen (GH) shifts exhibit the optically hysteretic response to the intensity of incident light. For thicker metal films, the GH shifts become very sensitive to the intensity of incident light and the angle of incidence.

  19. Dynamic nonlinear focal shift in amplitude modulated moderately focused acoustic beams.

    Science.gov (United States)

    Jiménez, Noé; Camarena, Francisco; González-Salido, Nuria

    2017-03-01

    The phenomenon of the displacement of the position of the pressure, intensity and acoustic radiation force maxima along the axis of focused acoustic beams under increasing driving amplitudes (nonlinear focal shift) is studied for the case of a moderately focused beam excited with continuous and 25kHz amplitude modulated signals, both in water and tissue. We prove that in amplitude modulated beams the linear and nonlinear propagation effects coexist in a semi-period of modulation, giving place to a complex dynamic behavior, where the singular points of the beam (peak pressure, rarefaction, intensity and acoustic radiation force) locate at different points on axis as a function of time. These entire phenomena are explained in terms of harmonic generation and absorption during the propagation in a lossy nonlinear medium both for a continuous and an amplitude modulated beam. One of the possible applications of the acoustic radiation force displacement is the generation of shear waves at different locations by using a focused mono-element transducer excited by an amplitude modulated signal.

  20. Non Gaussian and Non stationary characters of ENSO: the role of climate shifts and nonlinearities

    Science.gov (United States)

    Boucharel, J.; Dewitte, B.; Garel, B.; Du Penhoat, Y.

    2009-04-01

    ENSO is the dominant climate mode of variability in the Pacific, having socio-economical impacts on surrounding regions. ENSO exhibits a significant modulation at decadal to multidecadal timescales which is associated to changes of its characteristics (onset, amplitude, frequency, propagation, and predictability). Among these characteristics, some of them are generally ignored in ENSO studies, such as its asymmetry and the deviation of its statistics from those of the Gaussian distribution. These properties could be related to the ability of the current generation of coupled models to predict ENSO and its modulation. Here, the non-Gaussian nature and asymmetry of ENSO is diagnosed from in situ data and a variety of models (from intermediate complexity models to full physics coupled general circulation models) using robust statistical tools. In particular α-stable laws are used as theoretical background material to quantify the non-Gaussian character of ENSO time series. It is shown that the α-stable character of ENSO may result from the presence of climate shifts inducing non stationnarity in the time series. Also, cool (warm) periods are associated with ENSO statistics having a larger (weaker) tendency towards Gaussianity and a weaker (larger) asymmetry. This supports the hypothesis of ENSO being rectified by changes in mean state through non-linear processes. The relationship between change in mean state and non-linearity is investigated both in the Zebiak and Cane model and the IPCC models, which indicate that the propensity of a model to reproduce extreme events is related to its tendency to emphasize the non-linear interaction between mean state and ENSO variability. More particularly, high statistical moments i.e high order nonlinearities seem to be involved in the feedback between extreme events occurrence and mean state shit triggering through energy cascade, emphasizing the ENSO multifractal nature.

  1. Large enhancement of nonlinear Goos-Hänchen shifts and optical bistability due to surface plasmon excitations

    Science.gov (United States)

    Kim, Kihong

    2015-12-01

    The Goos-Hänchen shift of p wave beams incident on a metal-nonlinear dielectric bilayer in the Kretschmann configuration is studied theoretically. The reflectance, the phase of the reflection coefficient and the Goos-Hänchen shift are calculated in a numerically precise manner by using the invariant imbedding method. The Goos-Hänchen shift has been found to be able to take both extremely large positive and negative values due to surface plasmon excitations and very strong bistability and unique hysteresis phenomena appear. In addition, several previous results on the intensity dependence of the Goos-Hänchen shift are pointed out to be erroneous.

  2. Fiber optical parametric oscillator based on highly nonlinear dispersion-shifted fiber

    Institute of Scientific and Technical Information of China (English)

    Sigang YANG; Kenneth K. Y. WONG; Minghua CHEN; Shizhong XIE

    2013-01-01

    The development of fiber optical parametric oscillators (FOPO) based on highly nonlinear dispersion- shifted fiber is reviewed in this paper. Firstly, the background and motivation are introduced, and it is pointed out that the FOPO is promising to act as optical source in non-conventional wavelength bands. Subsequently, the context focuses principally on the problem of inherent multiple-longitudinal-mode characteristic of FOPO and the corresponding solutions to it. The primary technique is by locking the phase of multiple longitudinal modes. The first reported actively mode locked FOPO is also presented in this article. However, it is not probable to realize passively mode locked FOPO because of the random phase dithering of the pump required for suppressing stimulated Brillouin scattering. Furthermore, a regeneratively mode locked FOPO is demonstrated, which can generate wide band tunable radiation in non- conventional wavelengths. Besides mode locked FOPO, the single-longitudinal-mode FOPO is also introduced. Finally, potential future directions are discussed.

  3. Phase sensitivity in deformed-state superposition considering nonlinear phase shifts

    Science.gov (United States)

    Berrada, K.

    2016-07-01

    We study the problem of the phase estimation for the deformation-state superposition (DSS) under perfect and lossy (due to a dissipative interaction of DSS with their environment) regimes. The study is also devoted to the phase enhancement of the quantum states resulting from a generalized non-linearity of the phase shifts, both without and with losses. We find that such a kind of superposition can give the smallest variance in the phase parameter in comparison with usual Schrödinger cat states in different order of non-linearity even if for a larger average number of photons. Due to the significance of how a system is quantum correlated with its environment in the construction of a scalable quantum computer, the entanglement between the DSS and its environment is investigated during the dissipation. We show that partial entanglement trapping occurs during the dynamics depending on the kind of deformation and mean photon number. These features make the DSS with a larger average number of photons a good candidate for implementation of schemes of quantum optics and information with high precision.

  4. Numerical study of nonlinear singular fractional differential equations arising in biology by operational matrix of shifted Legendre polynomials

    Directory of Open Access Journals (Sweden)

    D. Jabari Sabeg

    2016-10-01

    Full Text Available In this paper, we present a new computational method for solving nonlinear singular boundary value problems of fractional order arising in biology. To this end, we apply the operational matrices of derivatives of shifted Legendre polynomials to reduce such problems to a system of nonlinear algebraic equations. To demonstrate the validity and applicability of the presented method, we present some numerical examples.

  5. Bragg-Scattering Four-Wave Mixing in Nonlinear Fibers with Intracavity Frequency-Shifted Laser Pumps

    Directory of Open Access Journals (Sweden)

    Katarzyna Krupa

    2012-01-01

    Full Text Available We experimentally study four-wave mixing in highly nonlinear fibers using two independent and partially coherent laser pumps and a third coherent signal. We focus our attention on the Bragg-scattering frequency conversion. The two pumps were obtained by amplifying two Intracavity frequency-shifted feedback lasers working in a continuous wave regime.

  6. Study on self-frequency-shift of femtosecond pulse in nonlinear dispersion medium using time-resolved cross-phase modulation method

    Institute of Scientific and Technical Information of China (English)

    赵应桥; 朱鹤元; 刘建华; 孙迭篪; 李富铭

    1997-01-01

    A time-resolved cross-phase modulation method combined with a modified nonlinear Schrodinger equation is used to study the effects of nonlinear response time on the propagation of ultrashort pulses in nonlinear dispersion media. Evolution of cross-phase modulation spectrum with the different time delay between the probe pulse and pump pulse is simulated using split-step Fourier method. It is shown that both normal self-frequency-shift-red-shift and abnormal self-frequency-shift-blue-shift can occur in the frequency domain for the probe pulse, and a satisfactory theoretical interpretation is given.

  7. Beware the two-beam coupling phase shift: Interpreting pump-probe measurements of the nonlinear refractive index

    CERN Document Server

    Wahlstrand, J K; McCole, E T; Cheng, Y -H; Palastro, J P; Levis, R J; Milchberg, H M

    2013-01-01

    Nonlinear optics experiments measuring phase shifts induced in a weak probe pulse by a strong pump pulse must account for coherent effects that only occur when the pump and probe pulses are temporally overlapped. It is well known that a weak probe beam experiences a greater phase shift from a strong pump beam than the pump beam induces on itself. The physical mechanism behind the enhanced phase shift is diffraction of pump light into the probe direction by a nonlinear refractive index grating produced by interference between the two beams. For an instantaneous third-order response, the effect of the grating is to simply double the probe phase shift, but when delayed nonlinearities are considered, the effect is more complex. A comprehensive treatment is given for both degenerate and nondegenerate pump-probe experiments in noble and diatomic gases. Results of numerical calculations are compared to a recent transient birefringence measurement [Loriot et al., Opt. Express 17, 13429 (2009)] and a recent spectral i...

  8. Two—photon Nonlinear Jaynes—Cummings Model with Stark Shift

    Institute of Scientific and Technical Information of China (English)

    董传华; 卢俊

    2002-01-01

    Two-photon Jaynes-Cummimgs model is generalized to the case of Kerr medium in this paper,The field and atom are prepared initially in two-photon superposition state and ground state respectively.Nonlinear coefficient affects the dynamic behaviors of the field and atom.Evolutions of the squeezing for the operators of field and atom and the quantum inversion are discussed.In particular,the higher-order squeezing for atomic dipole and the effects of nonlinearity on it,which have not been studied by other authors,are investigated,Increasing the nonlinear coefficient will decrease the squeezing depth of atomic dipole.

  9. Nonlinear Goos–Hänchen shifts of reflected light from inhomogeneous Kerr-like slabs

    Energy Technology Data Exchange (ETDEWEB)

    Mao, Hongmin, E-mail: hongminmao@hotmail.com; Zang, Taocheng; Sun, Jian; Pan, Tao; Xu, Guoding

    2013-09-02

    We investigate the Goos–Hänchen (GH) shifts of reflected light from Kerr-like slabs, whose permittivities are inhomogeneous in space as well as light intensity dependent. The GH shifts exhibit bistable, multivalued properties or a more complicated hysteretic response to the input light intensity, and the different spatial dependences of the permittivity have a great effect on the hysteretic response. The bistable or multivalued GH shifts can be modulated by various parameters, such as the angle of incidence and the thickness of slab. - Highlights: • The Goos–Hänchen (GH) shifts of reflected light from inhomogeneous Kerr-like slabs are investigated. • The GH shifts exhibit bistable, multivalued properties or a more complicated hysteretic response to the input light intensity. • The bistable or multivalued GH shifts can be modulated by the angle of incidence and the thickness of slab.

  10. Band gap shift and the optical nonlinear absorption of sputtered ZnO-TiO2 films.

    Science.gov (United States)

    Han, Yi-Bo; Han, Jun-Bo; Hao, Zhong-Hua

    2011-06-01

    ZnO-TiO2 composite films with different Zn/Ti atomic ratios were prepared with radio frequency reactive sputtering method. The Zn percentage composition (f(Zn)) dependent optical band gap and optical nonlinear absorption were investigated using the transmittance spectrum and the Z-scan technique, respectively. The results showed that composite films with f(Zn) in the range of 23.5%-88.3% are poor crystallized and their optical properties are anomalous which exhibit adjustable optical band gap and large optical nonlinear absorption. The optical absorption edge shifted to the blue wavelength direction with the increasing of f(Zn) and reached the minimum value of 285 nm for the sample with f(Zn) = 70.5%, which has the largest direct band gap of 4.30 eV. Further increasing of f(Zn) resulted in the red-shift of the optical absorption edge. The maximum optical nonlinear absorption coefficient of 1.5 x 10(3) cm/GW was also obtained for the same sample with f(Zn) = 70.5%, which is more than 40 times larger than those of pure TiO2 and ZnO films.

  11. Stable isotope analysis of vertebrae reveals ontogenetic changes in habitat in an endothermic pelagic shark.

    Science.gov (United States)

    Carlisle, Aaron B; Goldman, Kenneth J; Litvin, Steven Y; Madigan, Daniel J; Bigman, Jennifer S; Swithenbank, Alan M; Kline, Thomas C; Block, Barbara A

    2015-01-22

    Ontogenetic changes in habitat are driven by shifting life-history requirements and play an important role in population dynamics. However, large portions of the life history of many pelagic species are still poorly understood or unknown. We used a novel combination of stable isotope analysis of vertebral annuli, Bayesian mixing models, isoscapes and electronic tag data to reconstruct ontogenetic patterns of habitat and resource use in a pelagic apex predator, the salmon shark (Lamna ditropis). Results identified the North Pacific Transition Zone as the major nursery area for salmon sharks and revealed an ontogenetic shift around the age of maturity from oceanic to increased use of neritic habitats. The nursery habitat may reflect trade-offs between prey availability, predation pressure and thermal constraints on juvenile endothermic sharks. The ontogenetic shift in habitat coincided with a reduction of isotopic niche, possibly reflecting specialization upon particular prey or habitats. Using tagging data to inform Bayesian isotopic mixing models revealed that adult sharks primarily use neritic habitats of Alaska yet receive a trophic subsidy from oceanic habitats. Integrating the multiple methods used here provides a powerful approach to retrospectively study the ecology and life history of migratory species throughout their ontogeny.

  12. Nonlinear phase shifts of modulated light waves with slow and superluminal group delay in stimulated Brillouin scattering.

    Science.gov (United States)

    Arditi, Tal; Granot, Er'el; Sternklar, Shmuel

    2007-09-15

    Brillouin amplification with counterpropagating modulated pump and Stokes light leads to nonlinear modulation-phase shifts of the interacting intensity waves. This is due to a partial transformation of the nonmodulated light component at the input into modulated light at the output as a result of a mixing process with the counterpropagating modulated component of the pump and results in an advance or delay of the input modulation. This occurs for interactions over less than half of a modulation wavelength. Milliwatts of power in a kilometer of standard single-mode fiber give significant tunability of the modulation phase.

  13. Numerical Solutions of Odd Order Linear and Nonlinear Initial Value Problems Using a Shifted Jacobi Spectral Approximations

    Directory of Open Access Journals (Sweden)

    A. H. Bhrawy

    2012-01-01

    Full Text Available A shifted Jacobi Galerkin method is introduced to get a direct solution technique for solving the third- and fifth-order differential equations with constant coefficients subject to initial conditions. The key to the efficiency of these algorithms is to construct appropriate base functions, which lead to systems with specially structured matrices that can be efficiently inverted. A quadrature Galerkin method is introduced for the numerical solution of these problems with variable coefficients. A new shifted Jacobi collocation method based on basis functions satisfying the initial conditions is presented for solving nonlinear initial value problems. Through several numerical examples, we evaluate the accuracy and performance of the proposed algorithms. The algorithms are easy to implement and yield very accurate results.

  14. Non-linear DSGE Models, The Central Difference Kalman Filter, and The Mean Shifted Particle Filter

    DEFF Research Database (Denmark)

    Andreasen, Martin Møller

    This paper shows how non-linear DSGE models with potential non-normal shocks can be estimated by Quasi-Maximum Likelihood based on the Central Difference Kalman Filter (CDKF). The advantage of this estimator is that evaluating the quasi log-likelihood function only takes a fraction of a second. T...

  15. Analysis of contributions of nonlinear material constants to temperature-induced velocity shifts of quartz surface acoustic wave resonators.

    Science.gov (United States)

    Zhang, Haifeng; Kosinski, John A; Zuo, Lei

    2016-09-01

    In this paper, we examine the significance of the various higher-order effects regarding calculating temperature behavior from a set of material constants and their temperature coefficients. Temperature-induced velocity shifts have been calculated for quartz surface acoustic wave (SAW) resonators and the contributions of different groups of nonlinear material constants (third-order elastic constants (TOE), third-order piezoelectric constants (TOP), third-order dielectric constants (TOD) and electrostrictive constants (EL)) to the temperature-induced velocity shifts have been analyzed. The analytical methodology has been verified through the comparison of experimental and analytical results for quartz resonators. In general, the third-order elastic constants were found to contribute most significantly to the temperature-induced shifts in the SAW velocity. The contributions from the third-order dielectric constants and electrostrictive constants were found to be negligible. For some specific cases, the third-order piezoelectric constants were found to make a significant contribution to the temperature-induced shifts. The significance of each third-order elastic constant as a contributor to the temperature-velocity effect was analyzed by applying a 10% variation to each of the third-order elastic constants separately. Additionally, we have considered the issues arising from the commonly used thermoelastic expansions that provide a good but not exact description of the temperature effects on frequency in piezoelectric resonators as these commonly used expansions do not include the effects of higher-order material constants.

  16. Ontogenetic and interspecific metabolic scaling in insects.

    Science.gov (United States)

    Maino, James L; Kearney, Michael R

    2014-12-01

    Design constraints imposed by increasing size cause metabolic rate in animals to increase more slowly than mass. This ubiquitous biological phenomenon is referred to as metabolic scaling. However, mechanistic explanations for interspecific metabolic scaling do not apply to ontogenetic size changes within a species, implying different mechanisms for scaling phenomena. Here, we show that the dynamic energy budget theory approach of compartmentalizing biomass into reserve and structural components provides a unified framework for understanding ontogenetic and interspecific metabolic scaling. We formulate the theory for insects and show that it can account for ontogenetic metabolic scaling during the embryonic and larval phases, as well as the U-shaped respiration curve during pupation. After correcting for the predicted ontogenetic scaling effects, which we show to follow universal curves, the scaling of respiration between species is approximated by a three-quarters power law, supporting past empirical studies on insect metabolic scaling and our theoretical predictions. The ability to explain ontogenetic and interspecific metabolic scaling effects under one consistent framework suggests that the partitioning of biomass into reserve and structure is a necessary foundation to a general metabolic theory.

  17. Ontogenetic and seasonal shifts in the diet of the pemecou sea catfish Sciades herzbergii (Siluriformes: Ariidae, from a macrotidal mangrove creek in the Curuçá estuary, Northern Brazil

    Directory of Open Access Journals (Sweden)

    Tommaso Giarrizzo

    2008-06-01

    Full Text Available The feeding ecology of pemecou sea catfish (Sciades herzbergii was studied bimonthly from September 2003 to July 2004 in a macrotidal mangrove creek at Curuçá estuary, Brazil. Feeding activity and relative importance of prey groups were assessed in 528 and 226 specimens (6.2 to 36.0 cm total length, respectively, using stomach contents. A total of 1 820 specimens were caught. The fish left the creek with an average of 2.2 % of their body weight in prey. Feeding activity was lower in the dry season (September and November 2003 than in the rainy season (January to May 2004. The diet was dominated by Ocypodidae and Grapsidae. Feeding strategies of the pemecou sea catfish indicated a slight variation between the diets of adults and juveniles; however; the main factor leading to diversification in the diet were the seasons. Opportunistic feeding behavior was observed in the dry season, shifting to more specialized feeding in the wet season. This temporal diet shift may be explained by the environmental seasonal variations (e.g. transparency of the water, high water level in the intertidal mangrove creek habitat. Rev. Biol. Trop. 56 (2: 861-873. Epub 2008 June 30.Estudiamos la ecología trófica del bagre guatero Sciades herzbergii, con periodicidad bimestral, desde septiembre de 2003 a julio de 2004, en un canal de marea del estuario del río Curuçá, norte de Brasil. Del total de 1820 ejemplares capturados, analizamos la actividad alimentaría y la importancia de las diferentes fuentes de alimento por evaluación de los contenidos estomacales de 528 y 226 ejemplares (longitud corporal 6,2 a 36,0 cm, respectivamente. S. herzbergii sale del canal de marea con un valor promedio de 2,2 % de su peso corporal como alimento. El ritmo alimentario fue menor durante el período seco (septiembre a noviembre de 2003 que en el lluvioso (enero a mayo de 2004. Los análisis de los contenidos estomacales de 226 ejemplares indicaron una dieta con predominio de

  18. Nonlinear vegetation phenology shifts over northern China during 1982-2006

    Science.gov (United States)

    An, Youzhi; Liu, Wenbo; Gao, Wei; Gao, Zhiqiang; Liu, Chaoshun; Shi, Runhe

    2016-09-01

    The response of vegetation phenology change to climate change effects in the Northern China has been reported in the past several decades. Phenological change is a critical understanding in terrestrial carbon cycling. This study aims to investigate linear and nonlinear change trends and nonlinear response change trends in climate in vegetation phenology over Northern China in the last three decades. We analyzed the vegetation phenology over the Northern China by the new released Global Inventory Modeling and Mapping Studies (GIMMS) Normalized Difference Vegetation Index (NDVIg) dataset from 1982 to 2006.Results show that based on linear method, we can found that SOS was gradually advanced, EOS gradually delayed, and then LOS gradually lengthened. But on the basis of nonlinear method, phenological trends in the SOS, EOS and LOS are not continuous, we can found extended LOS with advanced SOS and delayed EOS before the turning point (TP) of spring SOS and autumn EOS trends and shortened LOS with delayed SOS and advanced EOS after the turning point (TP) of spring SOS and autumn EOS trends.

  19. Non-linear DSGE Models, The Central Difference Kalman Filter, and The Mean Shifted Particle Filter

    DEFF Research Database (Denmark)

    Andreasen, Martin Møller

    . The second contribution of this paper is to derive a new particle filter which we term the Mean Shifted Particle Filter (MSPFb). We show that the MSPFb outperforms the standard Particle Filter by delivering more precise state estimates, and in general the MSPFb has lower Monte Carlo variation in the reported...

  20. Optical encryption of parallel quadrature phase shift keying signals based on nondegenerate four-wave mixing in highly nonlinear fiber

    Science.gov (United States)

    Cui, Yue; Zhang, Min; Zhan, Yueying; Wang, Danshi; Huang, Shanguo

    2016-08-01

    A scheme for optical parallel encryption/decryption of quadrature phase shift keying (QPSK) signals is proposed, in which three QPSK signals at 10 Gb/s are encrypted and decrypted simultaneously in the optical domain through nondegenerate four-wave mixing in a highly nonlinear fiber. The results of theoretical analysis and simulations show that the scheme can perform high-speed wiretapping against the encryption of parallel signals and receiver sensitivities of encrypted signal and the decrypted signal are -25.9 and -23.8 dBm, respectively, at the forward error correction threshold. The results are useful for designing high-speed encryption/decryption of advanced modulated signals and thus enhancing the physical layer security of optical networks.

  1. A nonlinear training set superposition filter derived by neural network training methods for implementation in a shift-invariant optical correlator

    Science.gov (United States)

    Kypraios, Ioannis; Young, Rupert C. D.; Birch, Philip M.; Chatwin, Christopher R.

    2003-08-01

    The various types of synthetic discriminant function (sdf) filter result in a weighted linear superposition of the training set images. Neural network training procedures result in a non-linear superposition of the training set images or, effectively, a feature extraction process, which leads to better interpolation properties than achievable with the sdf filter. However, generally, shift invariance is lost since a data dependant non-linear weighting function is incorporated in the input data window. As a compromise, we train a non-linear superposition filter via neural network methods with the constraint of a linear input to allow for shift invariance. The filter can then be used in a frequency domain based optical correlator. Simulation results are presented that demonstrate the improved training set interpolation achieved by the non-linear filter as compared to a linear superposition filter.

  2. Mitigation of nonlinear effects through frequency shift free mid-span spectral inversion using counter-propagating dual pumped FWM in fiber

    Science.gov (United States)

    Anchal, Abhishek; Kumar, Pradeep; Landais, Pascal

    2016-10-01

    We propose and numerically verify a scheme of frequency-shift free mid-span spectral inversion (MSSI) for nonlinearity mitigation in an optical transmission system. Spectral inversion is accomplished by optical phase conjugation, realized by counter-propagating dual pumped four-wave mixing in a highly nonlinear fiber. We examine the performance of MSSI due to critical parameters such as nonlinear fiber length, pump and signal power. We demonstrate the near complete nonlinearity mitigation of 40 Gbps DQPSK modulated data transmitted over 1000 km standard single mode fiber using our method of MSSI. We perform simulation of bit-error rate as a function of optical signal to noise ratio to corroborate the effect of frequency-shift free MSSI.

  3. Spectroscopic investigations of dispersion-shifted fiber with thin active Bi-doped ring and high nonlinear refractive index

    Science.gov (United States)

    Zlenko, A. S.; Akhmetshin, U. G.; Bogatyrjov, V. A.; Bulatov, L. I.; Dvoyrin, V. V.; Firstov, S. V.; Dianov, E. M.

    2009-10-01

    A germanium-doped silica-core fiber with an active region in the form of a thin ring of silica doped with bismuth ions was fabricated. Bismuth doping in the ring surrounding the core allows to stabilize bismuth in silica glass, and it does not impose any restrictions on the composition of the core. The bismuth concentration in the ring is less than 0.2 wt.%. The GeO2 concentration in the core is more than 15 mol.%. A high germanium concentration in the core allows to shift the zero dispersion wavelength to 1860 nm and to obtain a high nonlinear refractive index (n2 more than 3,2*10-20 m2/W). Spectroscopic investigations were carried out in the visible and near infrared (800-1700 nm) spectral range. Despite the small concentration of bismuth, we observed the absorption and luminescence characteristic bands, confirming the presence of bismuth active centers in silica glass. Upon pumping at 1350 nm the on/off gain spectrum was measured on a 20-m fiber. The gain was observed throughout investigated range of 1430-1530 nm. The maximal gain of ~9.5 dB was obtained near 1430 nm. The results of the spectroscopic investigations of the fiber with a thin active Bi-doped ring showed prospects of the creation and application of such fiber type for laser and nonlinear optics.

  4. 80-Gb/s wavelength conversion based on cross-phase modulation in high-nonlinearity dispersion-shifted fiber and optical filtering

    DEFF Research Database (Denmark)

    Yu, Jianjun; Jeppesen, Palle

    2001-01-01

    Using cross-phase modulation in a 1-km high-nonlinearity dispersion-shifted fiber with subsequent filtering by a tunable optical filter, 80-Gb/s pulsewidth maintained wavelength conversion is realized. Penalty-free transmission over 80-km conventional single-mode fiber and 12-km dispersion...

  5. Optical Solitary Waves in Fourth-Order Dispersive Nonlinear Schr(o)dinger Equation with Self-steepening and Self-frequency Shift

    Institute of Scientific and Technical Information of China (English)

    2006-01-01

    By making use of the generalized sine-Gordon equation expansion method, we find cnoidal periodic wave solutions and fundamental bright and dark optical solitarywave solutions for the fourth-order dispersive and the quintic nonlinear Schrodinger equation with self-steepening, and self-frequency shift. Moreover, we discuss the formation conditions of the bright and dark solitary waves.

  6. The ontogenetic osteohistology of Tenontosaurus tilletti.

    Directory of Open Access Journals (Sweden)

    Sarah Werning

    Full Text Available Tenontosaurus tilletti is an ornithopod dinosaur known from the Early Cretaceous (Aptian-Albian Cloverly and Antlers formations of the Western United States. It is represented by a large number of specimens spanning a number of ontogenetic stages, and these specimens have been collected across a wide geographic range (from central Montana to southern Oklahoma. Here I describe the long bone histology of T. tilletti and discuss histological variation at the individual, ontogenetic and geographic levels. The ontogenetic pattern of bone histology in T. tilletti is similar to that of other dinosaurs, reflecting extremely rapid growth early in life, and sustained rapid growth through sub-adult ontogeny. But unlike other iguanodontians, this dinosaur shows an extended multi-year period of slow growth as skeletal maturity approached. Evidence of termination of growth (e.g., an external fundamental system is observed in only the largest individuals, although other histological signals in only slightly smaller specimens suggest a substantial slowing of growth later in life. Histological differences in the amount of remodeling and the number of lines of arrested growth varied among elements within individuals, but bone histology was conservative across sampled individuals of the species, despite known paleoenvironmental differences between the Antlers and Cloverly formations. The bone histology of T. tilletti indicates a much slower growth trajectory than observed for other iguanodontians (e.g., hadrosaurids, suggesting that those taxa reached much larger sizes than Tenontosaurus in a shorter time.

  7. Ontogenetic foraging activity and feeding selectivity of the Brazilian endemic parrotfish Scarus zelindae

    Science.gov (United States)

    Santos, Marcus; Lippi, Daniel L.; Silva, Pedro

    2016-01-01

    Parrotfish are fundamental species in controlling algal phase-shifts and ensuring the resilience of coral reefs. Nevertheless, little is known on their ecological role in the south-western Atlantic Ocean. The present study analysed the ontogenetic foraging activity and feeding selectivity of the Brazilian endemic parrotfish Scarus zelindae using behavioural observation and benthic composition analyses. We found a significant negative relationship between fish size and feeding rates for S. zelindae individuals. Thus, terminal phase individuals forage with lower feeding rates compared to juveniles and initial phase individuals. The highest relative foraging frequency of S. zelindae was on epilithic algae matrix (EAM) with similar values for juveniles (86.6%), initial phase (88.1%) and terminal phase (88.6%) individuals. The second preferred benthos for juveniles was sponge (11.6%) compared with initial (4.5%) and terminal life phases (1.3%). Different life phases of S. zelindae foraged on different benthos according to their availability. Based on Ivlev’s electivity index, juveniles selected EAM and sponge, while initial phase and terminal phase individuals only selected EAM. Our findings demonstrate that the foraging frequency of the endemic parrotfish S. zelindae is reduced according to body size and that there is a slight ontogenetic change in feeding selectivity. Therefore, ecological knowledge of ontogenetic variations on resource use is critical for the remaining parrotfish populations which have been dramatically reduced in the Southwestern Atlantic Ocean. PMID:27761330

  8. 北部湾鲔(Euthynnus affinis)的摄食习性及其随生长发育的变化%Feeding Habits and Ontogenetic Diet Shifts of Mackerel Tuna (Euthynnus affinis) in the Beibu Gulf, South China Sea

    Institute of Scientific and Technical Information of China (English)

    宿鑫; 李永东; 何雄波; 卢伙胜; 颜云榕

    2015-01-01

    To study the mackerel tuna (Euthynnus affinis) feeding habits and its ontogenetic diet shifts, 435 mackerel tuna were collected for an annual research from September 2008 to September 2009 in the main fishing ports such as Beihai, Basuo, Sanya and Jianghong of the Beibu Gulf, South China Sea. Results showed that the mackerel tuna of the Beibu Gulf was omnivorous and it predated not only the small pelagic fish but also the benthic fish, crustaceans and cephalopods. Pisces were dominant in the diets with the percentage of Index of Relative Importance (IRI%) of 98.57%. Among these, 21 species were identified. As indicated with IRI%, Bregmaceros rarisquamosus (45.50%), Stolephorus zollingeri (11.81%), Theyssa setirostris (5.89%), and Sardinella aurita (5.87%) were principal in the food cons-ituent. The main species changed monthly and it was Sardinella aurita in January; Bregmaceros rarisquamosus in February, March, November, December; Unidentified Pisces from April to July;Sardinella aurita in August; Leignathus riviulatus in September; Stolephorus zollingeri in October. The prey weight and numbers increased when body size of mackerel tuna was less than 340 mm (SL<340 mm), but there were decreased when body size of mackerel tuna was from 341 to 520 (341-520 mm). The spawning periods of the mackerel tuna were spring and summer, mainly from April to August. There were two spawn peaks, June and August, which were recognized by the female gonad somatic index (GSI) that maintained strong feeding intensity during the reproduction periods. Moreover, the peak periods were consistent with the water temperature of pelagic layers provided by France CATSAT fisheries remote sensing system.%基于2008年9月-2009年9月在北部湾逐月采集的435尾鲔(Euthynnus affinis)生物学数据,对鲔生长、摄食习性及其随生长发育的变化进行了分析。结果显示,北部湾鲔属于广食性鱼类,主要摄食中上层小型鱼类,另外还包括底栖鱼类、

  9. Color Shift Failure Prediction for Phosphor-Converted White LEDs by Modeling Features of Spectral Power Distribution with a Nonlinear Filter Approach.

    Science.gov (United States)

    Fan, Jiajie; Mohamed, Moumouni Guero; Qian, Cheng; Fan, Xuejun; Zhang, Guoqi; Pecht, Michael

    2017-07-18

    With the expanding application of light-emitting diodes (LEDs), the color quality of white LEDs has attracted much attention in several color-sensitive application fields, such as museum lighting, healthcare lighting and displays. Reliability concerns for white LEDs are changing from the luminous efficiency to color quality. However, most of the current available research on the reliability of LEDs is still focused on luminous flux depreciation rather than color shift failure. The spectral power distribution (SPD), defined as the radiant power distribution emitted by a light source at a range of visible wavelength, contains the most fundamental luminescence mechanisms of a light source. SPD is used as the quantitative inference of an LED's optical characteristics, including color coordinates that are widely used to represent the color shift process. Thus, to model the color shift failure of white LEDs during aging, this paper first extracts the features of an SPD, representing the characteristics of blue LED chips and phosphors, by multi-peak curve-fitting and modeling them with statistical functions. Then, because the shift processes of extracted features in aged LEDs are always nonlinear, a nonlinear state-space model is then developed to predict the color shift failure time within a self-adaptive particle filter framework. The results show that: (1) the failure mechanisms of LEDs can be identified by analyzing the extracted features of SPD with statistical curve-fitting and (2) the developed method can dynamically and accurately predict the color coordinates, correlated color temperatures (CCTs), and color rendering indexes (CRIs) of phosphor-converted (pc)-white LEDs, and also can estimate the residual color life.

  10. Global nonlinear optimization for the estimation of static shift and interpretation of 1-D magnetotelluric sounding data

    Directory of Open Access Journals (Sweden)

    Arkoprovo Biswas

    2011-07-01

    Full Text Available In the presence of conducting inhomogeneities in near-surface structures, apparent resistivity data in magnetotelluric sounding can be severely distorted. This is due to electric fields generated from boundary charges on surficial inhomogeneities. Such distortion persists throughout the entire recording range and is known as static shift in magnetotellurics. Frequency-independent static shifts manifest as vertical, parallel shifts that occur in plots of the dual logarithmic scale of apparent resistivity versus time period. The phase of magnetotelluric sounding data remains unaffected by the static shift and can be used to remove the static shift to some extent. However, individual inversion of phase data yields highly nonunique results, and alone it will not work to correctly remove the static shift. Inversions of uncorrected magnetotelluric data yield erroneous and unreliable estimations, while static-shift-corrected magnetotelluric data provide better and reliable estimations of the resistivities and thicknesses of subsurface structures. In the present study, static shift (a frequency-independent real constant is also considered as one of the model parameters and is optimized together with other model parameters (resistivity and thickness using the very fast simulated annealing global inversion technique. This implies that model parameters are determined simultaneously with the estimate of the static shift in the data. Synthetic and noisy data generated for a number of models are interpreted, to demonstrate the efficacy of the approach to yield reliable estimates of subsurface structures when the apparent resistivity data are affected by static shift. Individual inversions of static-shift-affected apparent resistivity data and phase data yield unreliable estimations of the model parameters. Furthermore, the estimated model parameters after individual data inversions do not show any systematic correlations with the amount of static shift in the

  11. Multiwavelength Raman fiber ring laser with the spectrum profile broadened by parametric four wave mixing in highly nonlinear dispersion-shifted fibers

    Institute of Scientific and Technical Information of China (English)

    Yan Wang; Qing Wang; Wei Zhang; Xiaoming Liu; Jiangde Peng

    2005-01-01

    @@ A broadband multiwavelength Raman fiber ring laser (RFRL) covering the whole C-band at room temperature are presented. The effect of the intracavity highly nonlinear dispersion-shifted fiber on broadening and flattening the output spectrum envelope is discussed and experimentally demonstrated. More than 45-dB extinction-ratio multiwavelength output from 1527.76 to 1566.86 nm with 100-GHz channel spacing and 2.1-dB power ripple has been achieved by carefully controlling the individual powers of three pump lasers.

  12. An ontogenetic perspective on individual differences.

    Science.gov (United States)

    Senner, Nathan R; Conklin, Jesse R; Piersma, Theunis

    2015-09-07

    Phenotypic differences among individuals can arise during any stage of life. Although several distinct processes underlying individual differences have been defined and studied (e.g. parental effects, senescence), we lack an explicit, unified perspective for understanding how these processes contribute separately and synergistically to observed variation in functional traits. We propose a conceptual framework based on a developmental view of life-history variation, linking each ontogenetic stage with the types of individual differences originating during that period. In our view, the salient differences among these types are encapsulated by three key criteria: timing of onset, when fitness consequences are realized, and potential for reversibility. To fill a critical gap in this framework, we formulate a new term to refer to individual differences generated during adulthood-reversible state effects. We define these as 'reversible changes in a functional trait resulting from life-history trade-offs during adulthood that affect fitness', highlighting how the adult phenotype can be repeatedly altered in response to environmental variation. Defining individual differences in terms of trade-offs allows explicit predictions regarding when and where fitness consequences should be expected. Moreover, viewing individual differences in a developmental context highlights how different processes can work in concert to shape phenotype and fitness, and lays a foundation for research linking individual differences to ecological and evolutionary theory.

  13. Giant Goos-Hänchen shifts and radiation-induced trapping of Helmholtz solitons at nonlinear interfaces.

    Science.gov (United States)

    Sánchez-Curto, Julio; Chamorro-Posada, Pedro; McDonald, Graham S

    2011-09-15

    Giant Goos-Hänchen shifts and radiation-induced trapping are studied at the planar boundary separating two focusing Kerr media within the framework of the Helmholtz theory. The analysis, valid for all angles of incidence, reveals that interfaces exhibiting linear external refraction can also accommodate both phenomena. Numerical evidence of these effects is provided, based on analytical predictions derived from a generalized Snell's law.

  14. Use of isotopic analysis of vertebrae in reconstructing ontogenetic feeding ecology in white sharks.

    Science.gov (United States)

    Estrada, James A; Rice, Aaron N; Natanson, Lisa J; Skomal, Gregory B

    2006-04-01

    We conducted stable 13C and 15N analysis on white shark vertebrae and demonstrated that incremental analysis of isotopes along the radius of a vertebral centrum produces a chronological record of dietary information, allowing for reconstruction of an individual's trophic history. Isotopic data showed significant enrichments in 15N with increasing sampling distance from the centrum center, indicating a correlation between body size and trophic level. Additionally, isotopic values verified two distinct ontogenetic trophic shifts in the white shark: one following parturition, marking a dietary switch from yolk to fish; and one at a total length of >341 cm, representing a known diet shift from fish to marine mammals. Retrospective trophic-level reconstruction using vertebral tissue will have broad applications in future studies on the ecology of threatened, endangered, or extinct species to determine life-long feeding patterns, which would be impossible through other methods.

  15. Variability in expression of Bothrops insularis snake venom proteases: an ontogenetic approach.

    Science.gov (United States)

    Zelanis, André; de Souza Ventura, Janaina; Chudzinski-Tavassi, Ana Marisa; de Fátima Domingues Furtado, Maria

    2007-05-01

    Bothrops insularis is a threatened snake endemic to Queimada Grande Island, southern coast of São Paulo, Brazil, and the occurrence of sexual abnormalities in males, females and intersexes (females with functional ovaries and rudimentary hemipenis) has been reported in this population. The aim of this study was to identify ontogenetic shifts in protease expression of offspring of captive-bred B. insularis. Three neonates from a single litter were maintained at the facilities of Laboratory of Herpetology, Institute Butantan, for 41 months. The snakes were individually milked and venoms were analyzed both by SDS-PAGE, under reducing conditions, and for biochemical activities. The venoms from the mother and from a pool of adult specimens were used as references. In regard to the electrophoretic patterns, common bands were identified mainly between 14 and 50 kDa among snakes. The occurrence of proteolytic activity was noticed predominantly between 27 and 45 kDa in zymograms. Inhibitory assays with 1,10-phenantroline (10 mM) and PMSF (5 mM) showed that venoms possessed both metalloproteases and serine proteases. Venoms of young specimens showed a higher coagulant activity than those of adults, especially upon factors X and II. All venoms presented fibrino(geno)lytic activity, degrading Aalpha and Bbeta chains of fibrinogen, and lysing fibrin plate. These findings can reflect important individual, ontogenetic and sexual differences on venom composition and are likely correlated with diet habits of this species.

  16. Morphological, ontogenetic and molecular characterization of Scutellospora reticulata (Glomeromycota)

    NARCIS (Netherlands)

    De Souza, F.A.; Declerck, S.; Smit, E.; Kowalchuk, G.A.

    2005-01-01

    The arbuscular mycorrhizal (AM) fungus Scutellospora reticulata (CNPAB11) was characterized using morphological, ontogenetic and molecular approaches. Spore ontogenesis was studied using Ri T-DNA transformed carrot roots and observations were compared with those published for eight other, pot-cultur

  17. Ontogenetic variation in the body stoichiometry of two fish species.

    Science.gov (United States)

    Boros, Gergely; Sály, Péter; Vanni, Michael J

    2015-10-01

    One of the central questions of ecological stoichiometry theory is to what extent animal species maintain constant elemental composition in their bodies. Although several recent studies demonstrate intraspecific variation in animal elemental composition, relatively little is known about ontogenetic changes in vertebrates, especially during early life stages. We studied the intraspecific and interspecific ontogenetic variation in the body stoichiometry of two fish species in two different orders; fathead minnow (Pimephales promelas) and sheepshead minnow (Cyprinodon variegatus), reared under controlled laboratory conditions. During ontogeny, we measured the chemical composition of fish bodies, including carbon (C), nitrogen (N), phosphorus (P), calcium (Ca), and ribonucleic acid (RNA) contents. We found that N and RNA contents were relatively high in early life stages and declined substantially during development. In contrast, body C and C:N ratios were relatively low in embryos, post-embryos and larvae, and increased remarkably thereafter. Concentrations and ratios of some elements (e.g., Ca, P, Ca:P) did not exhibit consistent ontogenetic trends, but fluctuated dynamically between consecutive developmental stages in both species. Specific growth rates correlated significantly with RNA contents in both species. Analyses of the relative importance of different P pools at each developmental stage revealed that RNA was a considerable P pool in post-embryos, while bone-associated P was the dominant body P pool in later stages. Our results suggest that the elemental composition of fish bodies changes considerably during ontogeny. Each ontogenetic stage has its own stoichiometric signature, but the timing, magnitude and direction of ontogenetic changes can vary substantially between taxa.

  18. Ontogenetic, spatial and temporal variation in trophic level and diet of Chukchi Sea fishes

    Science.gov (United States)

    Marsh, Jennifer M.; Mueter, Franz J.; Iken, Katrin; Danielson, Seth

    2017-01-01

    Climate warming and increasing development are expected to alter the ecosystem of the Chukchi Sea, including its fish communities. As a component of the Arctic Ecosystem Integrated Survey, we assessed the ontogenetic, spatial and temporal variability of the trophic level and diet of key fish species in the Chukchi Sea using N and C stable isotopes. During August and September of 2012 and 2013, 16 common fish species and two primary, invertebrate consumers were collected from surface, midwater and bottom trawls within the eastern Chukchi Sea. Linear mixed-effects models were used to detect possible variation in the relationship between body length and either δ13C or δ15N values among water masses and years for 13 fish species with an emphasis on Arctic cod (Boreogadus saida). We also examined the fish community isotopic niche space, trophic redundancy, and trophic separation within each water mass as measures of resiliency of the fish food web. Ontogenetic shifts in trophic level and diet were observed for most species and these changes tended to vary by water mass. As they increased in length, most fish species relied more on benthic prey with the exception of three forage fish species (walleye pollock, Gadus chalcogrammus, capelin, Mallotus villosus, and Pacific sandlance, Ammodytes hexapterus). Species that exhibited interannual differences in diet and trophic level were feeding at lower trophic levels and consumed a more pelagic diet in 2012 when zooplankton densities were higher. Fish communities occupied different isotopic niche spaces depending on water mass association. In more northerly Arctic waters, the fish community occupied the smallest isotopic niche space and relied heavily on a limited range of intermediate δ13C prey, whereas in warmer, nutrient-rich Bering Chukchi Summer Water, pelagic prey was important. In the warmest, Pacific-derived coastal water, fish consumed both benthic and pelagic prey. Examining how spatial gradients in trophic

  19. Allometric growth in the extant coelacanth lung during ontogenetic development.

    Science.gov (United States)

    Cupello, Camila; Brito, Paulo M; Herbin, Marc; Meunier, François J; Janvier, Philippe; Dutel, Hugo; Clément, Gaël

    2015-09-15

    Coelacanths are lobe-finned fishes known from the Devonian to Recent that were long considered extinct, until the discovery of two living species in deep marine waters of the Mozambique Channel and Sulawesi. Despite extensive studies, the pulmonary system of extant coelacanths has not been fully investigated. Here we confirm the presence of a lung and discuss its allometric growth in Latimeria chalumnae, based on a unique ontogenetic series. Our results demonstrate the presence of a potentially functional, well-developed lung in the earliest known coelacanth embryo, and its arrested growth at later ontogenetic stages, when the lung is clearly vestigial. The parallel development of a fatty organ for buoyancy control suggests a unique adaptation to deep-water environments. Furthermore, we provide the first evidence for the presence of small, hard, flexible plates around the lung in L. chalumnae, and consider them homologous to the plates of the 'calcified lung' of fossil coelacanths.

  20. Ontogenetic patterns in the dreams of women across the lifespan.

    Science.gov (United States)

    Dale, Allyson; Lortie-Lussier, Monique; De Koninck, Joseph

    2015-12-01

    The present study supports and extends previous research on the developmental differences in women's dreams across the lifespan. The participants included 75 Canadian women in each of 5 age groups from adolescence to old age including 12-17, 18-24, 25-39, 40-64, and 65-85, totaling 375 women. One dream per participant was scored by two independent judges using the method of content analysis. Trend analysis was used to determine the ontogenetic pattern of the dream content categories. Results demonstrated significant ontogenetic decreases (linear trends) for female and familiar characters, activities, aggression, and friendliness. These patterns of dream imagery reflect the waking developmental patterns as proposed by social theories and recognized features of aging as postulated by the continuity hypothesis. Limitations and suggestions for future research including the examining of developmental patterns in the dreams of males are discussed.

  1. Marine reserve design theory for species with ontogenetic migration.

    Science.gov (United States)

    White, J Wilson

    2015-01-01

    Models for marine reserve design have been developed primarily with 'reef fish' life histories in mind: sedentary adults in patches connected by larval dispersal. However, many fished species undertake ontogenetic migrations, such as from nursery grounds to adult spawning habitats, and current theory does not fully address the range of reserve options posed by that situation. I modelled a generic species with ontogenetic migration to investigate the possible benefits of reserves under three alternative scenarios. First, the fishery targets adult habitat, and reserves can sustain yields under high exploitation, unless habitat patches are well connected. Second, the fishery targets the nursery, and reserves are highly effective, regardless of connectivity patterns. Third, the fishery targets both habitats, and reserves only succeed if paired on adjacent, well-connected nursery and adult patches. In all cases, reserves can buffer populations against overexploitation but would not enhance fishery yield beyond that achievable by management without reserves. These results summarize the general situations in which management using reserves could be useful for ontogenetically migrating species, and the type of connectivity data needed to inform reserve design.

  2. Excitation mechanism of A1g mode and origin of nonlinear temperature dependence of Raman shift of CVD-grown mono- and few-layer MoS2 films.

    Science.gov (United States)

    Yang, Tianqi; Huang, Xiaoting; Zhou, Hong; Wu, Guangheng; Lai, Tianshu

    2016-05-30

    MoS2 films are grown on SiO2/Si substrates by chemical vapor deposition. The vibrational properties of optical phonons of mono-, bi- and multilayer MoS2 are studied by Raman scattering spectroscopy over temperature range from 90 to 540 K with 514.5 nm and 785 nm lasers. The Raman peaks of E2g1 and A1g modes are observed simultaneously for mono-, bi- and multilayer MoS2 with 514.5 nm laser, but only the Raman peak of E2g1 mode is seen for monolayer MoS2 as 785 nm laser is used, revealing electron-phonon exchange excitation mechanism of A1g mode for the first time. The Raman shifts of E2g1 and A1g modes present obvious nonlinear temperature dependence. A semi-quantitative model is used to fit the nonlinear temperature dependence of Raman shifts which matches well to experimental data. Meanwhile, the fitting results reveal that the nonlinear temperature dependence of Raman shifts of E2g1 mode mainly originates from three-phonon anharmonic effect, while one of A1g mode is contributed by stronger three- and weaker four-phonon anharmonic effects cooperatively but two contributions are comparable in intensity.

  3. Ontogenetic Shape Change in the Chicken Brain: Implications for Paleontology.

    Science.gov (United States)

    Kawabe, Soichiro; Matsuda, Seiji; Tsunekawa, Naoki; Endo, Hideki

    2015-01-01

    Paleontologists have investigated brain morphology of extinct birds with little information on post-hatching changes in avian brain morphology. Without the knowledge of ontogenesis, assessing brain morphology in fossil taxa could lead to misinterpretation of the phylogeny or neurosensory development of extinct species. Hence, it is imperative to determine how avian brain morphology changes during post-hatching growth. In this study, chicken brain shape was compared at various developmental stages using three-dimensional (3D) geometric morphometric analysis and the growth rate of brain regions was evaluated to explore post-hatching morphological changes. Microscopic MRI (μMRI) was used to acquire in vivo data from living and post-mortem chicken brains. The telencephalon rotates caudoventrally during growth. This change in shape leads to a relative caudodorsal rotation of the cerebellum and myelencephalon. In addition, all brain regions elongate rostrocaudally and this leads to a more slender brain shape. The growth rates of each brain region were constant and the slopes from the growth formula were parallel. The dominant pattern of ontogenetic shape change corresponded with interspecific shape changes due to increasing brain size. That is, the interspecific and ontogenetic changes in brain shape due to increased size have similar patterns. Although the shape of the brain and each brain region changed considerably, the volume ratio of each brain region did not change. This suggests that the brain can change its shape after completing functional differentiation of the brain regions. Moreover, these results show that consideration of ontogenetic changes in brain shape is necessary for an accurate assessment of brain morphology in paleontological studies.

  4. Ontogenetic Shape Change in the Chicken Brain: Implications for Paleontology.

    Directory of Open Access Journals (Sweden)

    Soichiro Kawabe

    Full Text Available Paleontologists have investigated brain morphology of extinct birds with little information on post-hatching changes in avian brain morphology. Without the knowledge of ontogenesis, assessing brain morphology in fossil taxa could lead to misinterpretation of the phylogeny or neurosensory development of extinct species. Hence, it is imperative to determine how avian brain morphology changes during post-hatching growth. In this study, chicken brain shape was compared at various developmental stages using three-dimensional (3D geometric morphometric analysis and the growth rate of brain regions was evaluated to explore post-hatching morphological changes. Microscopic MRI (μMRI was used to acquire in vivo data from living and post-mortem chicken brains. The telencephalon rotates caudoventrally during growth. This change in shape leads to a relative caudodorsal rotation of the cerebellum and myelencephalon. In addition, all brain regions elongate rostrocaudally and this leads to a more slender brain shape. The growth rates of each brain region were constant and the slopes from the growth formula were parallel. The dominant pattern of ontogenetic shape change corresponded with interspecific shape changes due to increasing brain size. That is, the interspecific and ontogenetic changes in brain shape due to increased size have similar patterns. Although the shape of the brain and each brain region changed considerably, the volume ratio of each brain region did not change. This suggests that the brain can change its shape after completing functional differentiation of the brain regions. Moreover, these results show that consideration of ontogenetic changes in brain shape is necessary for an accurate assessment of brain morphology in paleontological studies.

  5. Ontogenetic differentiation of swimming performance and behaviour in relation to habitat availability in the endangered North Sea houting (Coregonus oxyrinchus)

    DEFF Research Database (Denmark)

    Poulsen, Søren Brandt; Jensen, Lasse Fast; Schulz, Carsten

    2012-01-01

    The survival of the highly endangered, anadromous fish species North Sea houting (Coregonus oxyrinchus) depends on the correct timing of downstream dispersal during its early ontogenetic stages. To date, however, no studies have investigated the ontogenetic differentiation of swimming performance...

  6. Ontogenetic scaling of fish metabolism in the mouse-to-elephant mass magnitude range

    DEFF Research Database (Denmark)

    Moran, Damian; Wells, R.M.G.

    2007-01-01

    Intraspecific or ontogenetic analyses of mass-metabolism relationships do not often conform to the same allometric correlations as those seen in interspecific analyses. A commonly cited reason for this discrepancy is that ontogenetic studies examine smaller mass ranges than interspecific studies,...

  7. Ontogenetic origins of floral bilateral symmetry in Moringaceae (Brassicales).

    Science.gov (United States)

    Olson, Mark E

    2003-01-01

    Floral morphology of the 13 species of Moringa ranges from actinomorphic flowers with little hypanthium to highly zygomorphic flowers with well-developed hypanthia. Scanning electron and light microscopy were used to identify ontogenetic differences among two actinomorphic and eight zygomorphic species. All species show traces of zygomorphy between petal organogenesis and anther differentiation. At late organogenesis, zygomorphy is manifest by one petal being larger than the others, slight unidirectional maturation of the anthers, and in many species, some staminodes may be missing. At organ differentiation and beyond, the actinomorphic species show a trend toward increasing actinomorphy, whereas the zygomorphic features of early ontogeny are progressively accentuated throughout the ontogeny of the zygomorphic species. Because of the early traces of zygomorphy throughout the family, ontogeny in Moringa does not resemble that known from the sister taxon Caricaceae, which has flowers that are actinomorphic throughout ontogeny. Great intraspecific variation was found in floral plan in the actinomorphic-flowered species in contrast to the zygomorphic species. Each of the main clades in the family is distinguished by at least one feature of floral ontogeny. In general, ontogenetic differences that are congruent with deeper phylogenetic splits tend to occur earlier in ontogeny than those congruent with more recent divergences.

  8. Ontogenetic changes of phyllotaxis in Anagallis arvensis L.

    Directory of Open Access Journals (Sweden)

    Dorota Kwiatkowska

    2014-01-01

    Full Text Available During the ontogeny of Anagallis spontaneous changes of phyllotaxis appear in a regular sequence. The initial decussate pattern is followed by spiral Fibonacci phyllotaxis, this in turn, by a trimerous pattern, and finally Lucas spiral phyllotaxis is formed. In the course of the first and most common phyllotactic transition, from the decussate to spiral Fibonacci pattern, changes in primordia arrangement occur only within a limited sector of the apex circumference. In the complementary sector, primordia emerge as if the decussate phyllotaxis continued. It is likely that similar circumferential discontinuity accounts for further transitions. The common ontogenetic sequence of patterns in Anagallis is such that, theoretically, each transition requires minimal changes in shoot apex geometry. Although the meristem in Anagallis is able to produce primordia either in whorls or spirally, the elongated shoots of this plant seem to have leaves exclusively in whorls. It appeared that in shoots with an initially spiral pattern, leaves can be clustered in pseudo-whorls due to the uneven internode elongation. Pseudowhorls are composed usually of three (Fibonacci or four (Lucas leaves of successive nodes. The number of leaves in a pseudo-whorl equals the number of leaves positioned on one revolution of the ontogenetic helix, which is different in these two spiral patterns. In shoot apices with whorled phyllotaxis, the leaf and flower primordia of a whorl are of different size. On elongated shoots, flower buds emerging in the axils of leaves of one whorl also differ in size.

  9. Two-Photon Nonlinear Jaynes-Cummings Model with Stark Shift%具有Stark位移的非线性双光子Jaynes-Cummings模型

    Institute of Scientific and Technical Information of China (English)

    董传华; 卢俊

    2002-01-01

    Two-photon Jaynes-Cummimgs model is generalized to the case of Kerr medium in this paper. The field and atom are prepared initially in two-photon superposition state and ground state respectively. Nonlinear coefficient affects the dynamic behaviors of the field and atom. Evolutions of the squeezing for the operators of field and atom and the quantum inversion are discussed. In particular, the higher-order squeezing for atomic dipole and the effects of nonlinearity on it, which have not been studied by other authors,are investigated. Increasing the nonlinear coefficient will decrease the squeezing depth of atomic dipole.

  10. Beyond body size: muscle biochemistry and body shape explain ontogenetic variation of anti-predatory behaviour in the lizard Salvator merianae.

    Science.gov (United States)

    de Barros, Fábio Cury; de Carvalho, José Eduardo; Abe, Augusto Shinya; Kohlsdorf, Tiana

    2016-06-01

    Anti-predatory behaviour evolves under the strong action of natural selection because the success of individuals avoiding predation essentially defines their fitness. Choice of anti-predatory strategies is defined by prey characteristics as well as environmental temperature. An additional dimension often relegated in this multilevel equation is the ontogenetic component. In the tegu Salvator merianae, adults run away from predators at high temperatures but prefer fighting when it is cold, whereas juveniles exhibit the same flight strategy within a wide thermal range. Here, we integrate physiology and morphology to understand ontogenetic variation in the temperature-dependent shift of anti-predatory behaviour in these lizards. We compiled data for body shape and size, and quantified enzyme activity in hindlimb and head muscles, testing the hypothesis that morphophysiological models explain ontogenetic variation in behavioural associations. Our prediction is that juveniles exhibit body shape and muscle biochemistry that enhance flight strategies. We identified biochemical differences between muscles mainly in the LDH:CS ratio, whereby hindlimb muscles were more glycolytic than the jaw musculature. Juveniles, which often use evasive strategies to avoid predation, have more glycolytic hindlimb muscles and are much smaller when compared with adults 1-2 years old. Ontogenetic differences in body shape were identified but marginally contributed to behavioural variation between juvenile and adult tegus, and variation in anti-predatory behaviour in these lizards resides mainly in associations between body size and muscle biochemistry. Our results are discussed in the ecological context of predator avoidance by individuals differing in body size living at temperature-variable environments, where restrictions imposed by the cold could be compensated by specific phenotypes. © 2016. Published by The Company of Biologists Ltd.

  11. Ontogenetic niche shifts in three Vaccinium species on a sub-alpine mountain side

    DEFF Research Database (Denmark)

    Auffret, Alistair G.; Meineri, Eric; Bruun, Hans Henrik

    2010-01-01

    . uliginosum and V. vitis-idaea on a mountain slope in northern Sweden in relation to current adult occurrence. Methods: We combined a seed-sowing experiment in seven community types with adult occurrence observations and species distribution mapping. Results: Emergence of V. myrtillus and V. vitis...

  12. Ontogenetic habitat shift, population growth, and burrowing behavior of the Indo-Pacific beach star,

    NARCIS (Netherlands)

    Bos, A.R.; Gumanao, G.S.; van Katwijk, M.M.; Mueller, B.; Saceda, M.M.; Tejada, R.L.P.

    2011-01-01

    Archaster typicus, a common sea star in Indo-Pacific regions, has been a target for the ornamental trade, even though little is known about its population biology. Spatial and temporal patterns of abundance and size structure of A. typicus were studied in the Davao Gulf, the Philippines (125°42.7'E,

  13. Ontogenetic shifts in terrestrial reliance of stream-dwelling brown trout

    Directory of Open Access Journals (Sweden)

    Javier Sánchez-Hernández

    2016-07-01

    Full Text Available This study focuses on terrestrial reliance of brown trout (Salmo trutta and compared it to the potential prey available (macrozoobenthos and drifting invertebrates in three temperate rivers (Galicia, NW Spain, with special emphasis on variations in terrestrial energy intake through the ontogeny of brown trout. Additionally, we paid particular attention to individual variation of terrestrial resource use within and between age classes. Prey items were grouped in four categories: i aquatic invertebrates; ii imagoes of aquatic invertebrates; iii terrestrial invertebrates; and iv fish prey. Next, energy composition was measured according to dry weight-energy equations for each individual in line with above-mentioned prey categories. Our findings illustrate that terrestrial invertebrates appeared to be scarce in the environment, whereas aquatic food resources were rather abundant and accessible. The use of terrestrial invertebrates tended to increase with age, but with a high degree of inter-individual variation in resource use. In fact, the individual reliance of brown trout on terrestrial invertebrates may vary considerably (between 0% and 76.9%. Besides, the frequency of terrestrial foragers, i.e., individuals with terrestrial invertebrates in their stomachs, increased with age, except in one population which had the maximum value in the age-2 class. The acquisition of terrestrial invertebrates thus appears to be a process strongly dependent upon the actual food availability in the environment, but with a high degree of individual variance in resource use within the same age class. Finally, we discuss that terrestrial invertebrates may largely contribute to cover the energy intake of the species, highlighting the interface between terrestrial and aquatic ecosystems, and thereby the importance of riparian canopy cover as a key factor for food supply of stream-dwelling salmonids species.

  14. Ontogenetic and phylogenetic shifts in the cognitive representation of the conditioned response.

    Science.gov (United States)

    Lindberg, M A

    1982-03-01

    The retention of the conditioned response was tested in a retroactive interference paradigm using subjects from kindergarten, Grades 3 and 6, and college. The conditioned stimuli (CS) were either a tone or light, and the unconditioned stimuli (US) were vibratory tactual stimulations delivered to either the right or left index finger which produced a withdrawal response. Original AB learning consisted of pairing one of the CSs with one of the USs (e.g., tone-left). Interpolated CD learning consisted of pairing the other CS with the other US (e.g., light-right). When the first stimulus was presented alone to test for retroactive interference, the college students gave the response associated with it during original learning, the B response, but the kindergartners and third graders gave the D response, which was never associated with the CS during training. A second experiment used avoidance conditioning instructions and replicated these results while including rest-control and AD groups. These data suggest that what is learned by children in simple conditioning paradigms is different than what is learned by adults in the same paradigms. The results are discussed in terms of the following two hypotheses: (a) Russian theories of conditioning and extensions of Tighe and Tighe's and Kendler's theories of the development of perception and discrimination learning in children, and (b) theories of development of separability in perceptual development.

  15. Phenological mismatch and ontogenetic diet shifts interactively affect offspring condition in a passerine

    NARCIS (Netherlands)

    Samplonius, Jelmer M.; Kappers, Elena F.; Brands, Stef; Both, Christiaan

    1. Climate change may cause phenological asynchrony between trophic levels, which can lead to mismatched reproduction in animals. Although indirect effects of mismatch on fitness are well described, direct effects on parental prey choice are not. Moreover, direct effects of prey variation on

  16. Ontogenetic comparisons of standard metabolism in three species of crocodilians

    Science.gov (United States)

    Brien, Matthew L.; Tracy, Christopher R.; Manolis, S. Charlie; Webb, Grahame J. W.; Seymour, Roger S.; Christian, Keith A.

    2017-01-01

    Due in part to their large size, aggressive temperament, and difficulty in handling, there are few physiological studies of adult crocodilians in the literature. As a result, studies comparing individuals across an ontogenetic series and comparisons among species are also lacking. We addressed this gap in knowledge by measuring standard metabolic rates (SMR) of three species of crocodilians (Crocodylus porosus, C. johnsoni, and Alligator mississippiensis), and included individuals that ranged from 0.22 to 114 kg. Allometric scaling of SMR with body mass was similar among the species, but C. porosus had significantly higher SMR than did C. johnsoni or A. mississippiensis. Differences in SMR among species are potentially related to behavioural differences in levels of aggression; C. porosus are the most aggressive of the crocodilians measured, and have rates of standard metabolism that are approximately 36% higher at the grand mean body size than those measured for C. johnsoni or A. mississippiensis, which are among the least aggressive crocodilians. PMID:28182696

  17. Comparative vs ontogenetic paradigms for tests of the intrinsic mutagenesis hypothesis of aging

    Energy Technology Data Exchange (ETDEWEB)

    Sacher, G.A.

    1979-01-01

    This paper examines certain aspects of the biology of longevity and aging that bear on the role of DNA in the ontogenetic aging process, and on the genetic basis for the differences of longevity among mammal species. (PCS)

  18. Exploring the ontogenetic scaling hypothesis during the diversification of pollination syndromes in Caiophora (Loasaceae, subfam. Loasoideae)

    National Research Council Canada - National Science Library

    Strelin, Marina M; Benitez-Vieyra, Santiago; Fornoni, Juan; Klingenberg, Christian Peter; Cocucci, Andrea A

    2016-01-01

    ... (ontogenetic scaling hypothesis), the rarity of reversion to ancestral states following adaptive radiations of pollination syndromes suggests that changes in the ancestral developmental programme of the flower are common during...

  19. Ontogenetic variations in the venom proteome of the Amazonian snake Bothrops atrox

    Directory of Open Access Journals (Sweden)

    Sousa Marcelo V

    2006-05-01

    Full Text Available Abstract Background Bothrops atrox is responsible for the majority of snakebite accidents in the Brazilian Amazon region. Previous studies have demonstrated that the biological and pharmacological activities of B. atrox venom alter with the age of the animal. Here, we present a comparative proteome analysis of B. atrox venom collected from specimens of three different stages of maturation: juveniles, sub-adults and adults. Results Optimized conditions for two-dimensional gel electrophoresis (2-DE of pooled venom samples were achieved using immobilized pH gradient (IPG gels of non-linear 3–10 pH range during the isoelectric focusing step and 10–20% gradient polyacrylamide gels in the second dimension. Software-assisted analysis of the 2-DE gels images demonstrated differences in the number and intensity of spots in juvenile, sub-adult and adult venoms. Although peptide mass fingerprinting (PMF failed to identify even a minor fraction of spots, it allowed us to group spots that displayed similar peptide maps. The spots were subjected to a combination of tandem mass spectrometry and Mascot and MS BLAST database searches that identified several classes of proteins, including metalloproteinases, serine proteinases, lectins, phospholipases A2, L-amino oxidases, nerve growth factors, vascular endothelial growth factors and cysteine-rich secretory proteins. Conclusion The analysis of B. atrox samples from specimens of different ages by 2-DE and mass spectrometry suggested that venom proteome alters upon ontogenetic development. We identified stage specific and differentially expressed polypeptides that may be responsible for the activities of the venom in each developmental stage. The results provide insight into the molecular basis of the relation between symptomatology of snakebite accidents in humans and the venom composition. Our findings underscore the importance of the use of venoms from individual specimen at various stages of maturation for

  20. Ontogenetic taurine biosynthesis ability in rainbow trout (Oncorhynchus mykiss).

    Science.gov (United States)

    Wang, Xuan; He, Gen; Mai, Kangsen; Xu, Wei; Zhou, Huihui

    2015-07-01

    Taurine (2-aminoethane sulfonic acid) plays important roles in multiple physiological processes including osmoregulation, bile salt conjugation and membrane protection. It is known that taurine biosynthesis varies in different fish species. However, its ontogenetic regulation has not been clear. In the present study, we found that the hepatic concentrations of taurine increased marginally with rainbow trout growth. The mRNA expression, protein levels and enzyme activities of key enzymes involved in taurine biosynthesis, cysteine dioxygenase (CDO) and cysteine sulfinate decarboxylase (CSD), were analyzed. Our results showed that the mRNA levels and protein abundances of CSD increased dramatically with the development of rainbow trout stages while the enzyme activities showed a slight improvement. However, the expression and activities of CDO decreased with rainbow trout growth. These results provide valuable information on defining the exact supplementation of taurine in diets for different stages of rainbow trout and give new insights into elucidating the regulation of taurine metabolism in rainbow trout. Copyright © 2015 Elsevier Inc. All rights reserved.

  1. Ontogenetic data and the evolutionary origin of the mammalian scapula

    Science.gov (United States)

    Sánchez-Villagra, Marcelo; Maier, Wolfgang

    2002-09-01

    One of the most useful diagnostic therian features is the division of the lateral surface of the scapula into two fossae separated by a scapular spine. A new model to describe this evolutionary innovation based on ontogenetic data is provided, consistent with information provided by recent fossil discoveries. The development of the scapula in three didelphid and one dasyurid marsupials using histological sections was studied. Only the ventral, acromial portion of the scapular spine, which originates from the anterior margin of the scapular blade, is preformed in cartilage. The major dorsal portion is formed at a later stage by appositional bone, which expands from the perichondral ossification of the scapula into an intermuscular aponeurosis between the supra-and infraspinatus muscles. This intermuscular aponeurosis inserts more or less in the middle of the lateral surface of the developing scapula. Thus, the floor of the supraspinous fossa is already present at the beginning of scapular development, simultaneous with the infraspinous fossa. The portion of the scapular spine that is situated dorsal to the acromial process is hypothesized to be a neomorphic structure of therians. The dorsal portion of the scapular spine evolved as additional attachment for powerful supraspinatus and infraspinatus muscles meeting near the middle of the lateral scapula.

  2. Ontogenetic and among-individual variation in foraging strategies of northeast Pacific white sharks based on stable isotope analysis

    Science.gov (United States)

    Kim, S.L.; Tinker, M. Tim; Estes, J.A.; Koch, P.L.

    2012-01-01

    There is growing evidence for individuality in dietary preferences and foraging behaviors within populations of various species. This is especially important for apex predators, since they can potentially have wide dietary niches and a large impact on trophic dynamics within ecosystems. We evaluate the diet of an apex predator, the white shark (Carcharodon carcharias), by measuring the stable carbon and nitrogen isotope composition of vertebral growth bands to create lifetime records for 15 individuals from California. Isotopic variations in white shark diets can reflect within-region differences among prey (most importantly related to trophic level), as well as differences in baseline values among the regions in which sharks forage, and both prey and habitat preferences may shift with age. The magnitude of isotopic variation among sharks in our study (>5‰ for both elements) is too great to be explained solely by geographic differences, and so must reflect differences in prey choice that may vary with sex, size, age and location. Ontogenetic patterns in δ15N values vary considerably among individuals, and one third of the population fit each of these descriptions: 1) δ15N values increased throughout life, 2) δ15N values increased to a plateau at ~5 years of age, and 3) δ15N values remained roughly constant values throughout life. Isotopic data for the population span more than one trophic level, and we offer a qualitative evaluation of diet using shark-specific collagen discrimination factors estimated from a 3+ year captive feeding experiment (Δ13Cshark-diet and Δ15Nshark-diet equal 4.2‰ and 2.5‰, respectively). We assess the degree of individuality with a proportional similarity index that distinguishes specialists and generalists. The isotopic variance is partitioned among differences between-individual (48%), within-individuals (40%), and by calendar year of sub-adulthood (12%). Our data reveal substantial ontogenetic and individual dietary

  3. The Making of a Monster: Postnatal Ontogenetic Changes in Craniomandibular Shape in the Great Sabercat Smilodon

    Science.gov (United States)

    Christiansen, Per

    2012-01-01

    Derived sabercats had craniomandibular morphologies that in many respects were highly different from those of extant felids, and this has often been interpreted functionally as adaptations for predation at extreme gape angles with hypertrophied upper canines. It is unknown how much of this was a result of intraspecific postnatal ontogeny, since juveniles of sabercats are rare and no quantitative study has been made of craniomandibular ontogeny. Postnatal ontogenetic craniomandibular shape changes in two morphologically derived sabercats, Smilodon fatalis and S. populator, were analysed using geometric morphometrics and compared to three species of extant pantherines, the jaguar, tiger, and Sunda clouded leopard. Ontogenetic shape changes in Smilodon usually involved the same areas of the cranium and mandible as in extant pantherines, and large-scale modularization was similar, suggesting that such may have been the case for all felids, since it followed the same trends previously observed in other mammals. However, in other respects Smilodon differed from extant pantherines. Their crania underwent much greater and more localised ontogenetic shape changes than did the mandibles, whereas crania and mandibles of extant pantherines underwent smaller, fewer and less localised shape changes. Ontogenetic shape changes in the two species of Smilodon are largely similar, but differences are also present, notably those which may be tied to the presence of larger upper canines in S. populator. Several of the specialized cranial characters differentiating adult Smilodon from extant felids in a functional context, which are usually regarded as evolutionary adaptations for achieving high gape angles, are ontogenetic, and in several instances ontogeny appears to recapitulate phylogeny to some extent. No such ontogenetic evolutionary adaptive changes were found in the extant pantherines. Evolution in morphologically derived sabercats involved greater cranial ontogenetic changes

  4. The making of a monster: postnatal ontogenetic changes in craniomandibular shape in the great sabercat Smilodon.

    Directory of Open Access Journals (Sweden)

    Per Christiansen

    Full Text Available Derived sabercats had craniomandibular morphologies that in many respects were highly different from those of extant felids, and this has often been interpreted functionally as adaptations for predation at extreme gape angles with hypertrophied upper canines. It is unknown how much of this was a result of intraspecific postnatal ontogeny, since juveniles of sabercats are rare and no quantitative study has been made of craniomandibular ontogeny. Postnatal ontogenetic craniomandibular shape changes in two morphologically derived sabercats, Smilodon fatalis and S. populator, were analysed using geometric morphometrics and compared to three species of extant pantherines, the jaguar, tiger, and Sunda clouded leopard. Ontogenetic shape changes in Smilodon usually involved the same areas of the cranium and mandible as in extant pantherines, and large-scale modularization was similar, suggesting that such may have been the case for all felids, since it followed the same trends previously observed in other mammals. However, in other respects Smilodon differed from extant pantherines. Their crania underwent much greater and more localised ontogenetic shape changes than did the mandibles, whereas crania and mandibles of extant pantherines underwent smaller, fewer and less localised shape changes. Ontogenetic shape changes in the two species of Smilodon are largely similar, but differences are also present, notably those which may be tied to the presence of larger upper canines in S. populator. Several of the specialized cranial characters differentiating adult Smilodon from extant felids in a functional context, which are usually regarded as evolutionary adaptations for achieving high gape angles, are ontogenetic, and in several instances ontogeny appears to recapitulate phylogeny to some extent. No such ontogenetic evolutionary adaptive changes were found in the extant pantherines. Evolution in morphologically derived sabercats involved greater cranial

  5. Exploring the ontogenetic scaling hypothesis during the diversification of pollination syndromes in Caiophora (Loasaceae, subfam. Loasoideae).

    Science.gov (United States)

    Strelin, Marina M; Benitez-Vieyra, Santiago; Fornoni, Juan; Klingenberg, Christian Peter; Cocucci, Andrea A

    2016-04-01

    Phenotypic diversification of flowers is frequently attributed to selection by different functional groups of pollinators. During optimization of floral phenotype, developmental robustness to genetic and non-genetic perturbations is expected to limit the phenotypic space available for future evolutionary changes. Although adaptive divergence can occur without altering the basic developmental programme of the flower (ontogenetic scaling hypothesis), the rarity of reversion to ancestral states following adaptive radiations of pollination syndromes suggests that changes in the ancestral developmental programme of the flower are common during such evolutionary transitions. Evidence suggests that flower diversification into different pollination syndromes in the Loasoideae genus Caiophora took place during a recent adaptive radiation in the central Andes. This involved transitions from bee to hummingbird and small rodent pollination. The aim of this work was to examine if the adaptive radiation of pollination syndromes in Caiophora occurred through ontogenetic scaling or involved a departure from the ontogenetic pattern basal to this genus. We used geometric morphometric variables to describe the shape and size of floral structures taking part in the pollination mechanism of Loasoideae. This approach was used to characterize the developmental trajectories of three species basal to the genus Caiophora through shape-size relationships (ontogenetic allometry). We then tested if the shape-size combinations of these structures in mature flowers of derived Caiophora species fall within the phenotypic space predicted by the development of basal species. Variation in the size and shape of Caiophora flowers does not overlap with the pattern of ontogenetic allometry of basal species. Derived bee-, hummingbird- and rodent-pollinated species had divergent ontogenetic patterns of floral development from that observed for basal bee-pollinated species. The adaptive radiation of

  6. Seasonal and ontogenetic variation of skin microbial communities and relationships to natural disease dynamics in declining amphibians

    Science.gov (United States)

    Longo, Ana V.; Savage, Anna E.; Hewson, Ian; Zamudio, Kelly R.

    2015-01-01

    Recently, microbiologists have focused on characterizing the probiotic role of skin bacteria for amphibians threatened by the fungal disease chytridiomycosis. However, the specific characteristics of microbial diversity required to maintain health or trigger disease are still not well understood in natural populations. We hypothesized that seasonal and developmental transitions affecting susceptibility to chytridiomycosis could also alter the stability of microbial assemblages. To test our hypothesis, we examined patterns of skin bacterial diversity in two species of declining amphibians (Lithobates yavapaiensis and Eleutherodactylus coqui) affected by the pathogenic fungus Batrachochytrium dendrobatidis (Bd). We focused on two important transitions that affect Bd susceptibility: ontogenetic (from juvenile to adult) shifts in E. coqui and seasonal (from summer to winter) shifts in L. yavapaiensis. We used a combination of community-fingerprinting analyses and 16S rRNA amplicon sequencing to quantify changes in bacterial diversity and assemblage composition between seasons and developmental stages, and to investigate the relationship between bacterial diversity and pathogen load. We found that winter-sampled frogs and juveniles, two states associated with increased Bd susceptibility, exhibited higher diversity compared with summer-sampled frogs and adult individuals. Our findings also revealed that hosts harbouring higher bacterial diversity carried lower Bd infections, providing support for the protective role of bacterial communities. Ongoing work to understand skin microbiome resilience after pathogen disturbance has the potential to identify key taxa involved in disease resistance. PMID:26587253

  7. The ontogenetic switch between odonate life history stages: effects on fitness when time and food are limited.

    Science.gov (United States)

    Plaistow; Siva-jothy

    1999-09-01

    During the course of ontogeny, odonates switch from being aquatic larvae to being terrestrial adults. Ontogenetic niche shift theory proposes that such shifts are adaptive and have evolved to maximize a growth rate (size) to mortality rate ratio. Individuals should therefore switch from one niche to the other at an optimal size or state. Since the majority of odonates are seasonal breeders, the extent to which the switch is optimal will depend upon the time and the resources available during postembryonic development. We collected a cohort of larvae that varied in how close they were to eclosion and reared them on either a high-nutrition or a low-nutrition diet. We then determined the relative influence of both time and nutritional constraints on survival and development rate, as well as the body size, size-corrected flight muscle mass and fat reserves of individuals at eclosion. Damselflies in both high- and low-nutrition treatments responded to a short development period by developing faster and reducing their body size, but did not change their proportional investment in fat reserves and flight muscle. Reduced larval nutrition resulted in decreased body size, flight muscle mass and fat reserves at eclosion. However, it had no effect on survival to eclosion, or development rate. We discuss these results in terms of the influence that time and nutritional constraints have on odonate development patterns and fitness. Copyright 1999 The Association for the Study of Animal Behaviour.

  8. Nonlinear magnetic metamaterials.

    Science.gov (United States)

    Shadrivov, Ilya V; Kozyrev, Alexander B; van der Weide, Daniel W; Kivshar, Yuri S

    2008-12-08

    We study experimentally nonlinear tunable magnetic metamaterials operating at microwave frequencies. We fabricate the nonlinear metamaterial composed of double split-ring resonators where a varactor diode is introduced into each resonator so that the magnetic resonance can be tuned dynamically by varying the input power. We demonstrate that at higher powers the transmission of the metamaterial becomes power-dependent and, as a result, such metamaterial can demonstrate various nonlinear properties. In particular, we study experimentally the power-dependent shift of the transmission band and demonstrate nonlinearity-induced enhancement (or suppression) of wave transmission. (c) 2008 Optical Society of America

  9. Diet-morphology relationship in the stream-dwelling characid Deuterodon stigmaturus (Gomes, 1947 (Characiformes: Characidae is partially conditioned by ontogenetic development

    Directory of Open Access Journals (Sweden)

    Renato Bolson Dala-Corte

    Full Text Available ABSTRACT We tested whether interindividual variations in diet composition within a population of Deuterodon stigmaturus can be explained by morphological differences between individuals, and whether diet-morphology relationships are dependent on the ontogenetic development. We analyzed diet of 75 specimens sampled in a coastal stream of Southern Brazil. Variation in stomach content was summarized with a Principal Coordinate Analysis (PCoA. The retained PCoA axes were tested as response to standard length (SL, and to values of intestine length (IL and mouth length (ML independent of body size, using linear mixed-effects models (LMM. The most consumed food items by D. stigmaturus were filamentous algae (41%, terrestrial plants (20.3%, detritus (12%, and aquatic invertebrates (8.8%. The LMMs showed that SL was positively related to consumption of terrestrial plants, whereas IL independent of SL was negatively related to aquatic invertebrates and positively related to filamentous algae. When body sized was held constant, ML was not related to diet variation. Interindividual diet differences conditioned to body size suggest that individuals shift their trophic niche and function in the ecosystem along the ontogenetic development. Relationships between intestine length and diet composition suggest interindividual differences in foraging ability and digestibility of distinct food items.

  10. Fast Programming Algorithm to Find Non-Linear Feedback Shift Register%快速寻找非线性反馈移位寄存器的编程算法

    Institute of Scientific and Technical Information of China (English)

    叶炜晨; 陈克非

    2014-01-01

    Non-linear feedback shift register (NLFSR) is a common device to generate pseudo-random sequences in stream cipher. However, there is still no effective mathematical algorithm to find NLFSRs for a given period or output sequence. This paper provides a quick method to find NLFSRs. This method is a programming algorithm based on compute unified device architecture (CUDA) and parallel computing, and can quickly find NLFSRs for the given period or output sequence. This method has very good performance on both simple and complex NLFSRs. With this new method, people can easily get a large amount of experimental data about NLFSRs. It will be a great help for the future research on the mathematical algorithm to find NLFSRs.%在流密码中,非线性反馈移位寄存器(non-linear feedback shift register,NLFSR)是一种常用的安全性较高的伪随机序列生成器。目前仍然没有一种普遍有效的数学算法,能够根据给定的序列或者序列周期,直接推导出NLFSR。提出了一种快速寻找NLFSR的编程算法。该算法基于统一计算架构(compute unified device architecture,CUDA)和并行计算来实现,计算速度快,尤其适用于处理高次数的复杂NLFSR。并且该算法可以快速大规模地计算出NLFSR,为未来研究寻找NLFSR的数学算法提供了大量的实验数据。

  11. Interindividual variability and ontogenetic effects on Mg and Sr incorporation in the planktonic foraminifer Globigerinoides sacculifer

    NARCIS (Netherlands)

    Duenas-Bohorquez, A.; Da Rocha, R.E.; Kuroyanagi, A.; Nooijer, L.J. de; Bijma, J.; Reichart, G.-J.

    2011-01-01

    In order to investigate the interindividual and ontogenetic effects on Mg and Sr incorporation, magnesium/calcium (Mg/Ca) and strontium/calcium (Sr/Ca) ratios of cultured planktonic foraminifera have been determined. Specimens of Globigerinoides sacculifer were grown under controlled physical and

  12. Functional specialization and ontogenetic scaling of limb anatomy in Alligator mississippiensis

    Science.gov (United States)

    Allen, Vivian; Elsey, Ruth M; Jones, Nicola; Wright, Jordon; Hutchinson, John R

    2010-01-01

    Crocodylians exhibit a fascinating diversity of terrestrial gaits and limb motions that remain poorly described and are of great importance to understanding their natural history and evolution. Their musculoskeletal anatomy is pivotal to this diversity and yet only qualitative studies of muscle-tendon unit anatomy exist. The relative masses and internal architecture (fascicle lengths and physiological cross-sectional areas) of muscles of the pectoral and pelvic limbs of American alligators (Alligator mississippiensis Daudin 1801) were recorded for an ontogenetic series of wild specimens (n = 15, body masses from 0.5 to 60 kg). The data were analysed by reduced major axis regression to determine scaling relationships with body mass. Physiological cross-sectional areas and therefore muscle force-generating capacity were found to be greater in the extensor (anti-gravity) muscles of the pelvic limb than in the pectoral limb, reflecting how crocodylians differ from mammals in having greater loading of the hindlimbs than the forelimbs. Muscle masses and architecture were generally found to scale isometrically with body mass, suggesting an ontogenetic decrease in terrestrial athleticism. This concurs with the findings of previous studies showing ontogenetic decreases in limb bone length and the general scaling principle of a decline of strength : weight ratios with increasing size in animals. Exceptions to isometric scaling found included positive allometry in fascicle length for extensor musculature of both limbs, suggesting an ontogenetic increase in working range interpreted as increasing postural variability – in particular the major hip extensors – the interpretation of which is complicated by previous described ontogenetic increase of moment arms for these muscles. PMID:20148991

  13. Heritability of hsp70 expression in the beetle Tenebrio molitor: Ontogenetic and environmental effects.

    Science.gov (United States)

    Lardies, Marco A; Arias, María Belén; Poupin, María Josefina; Bacigalupe, Leonardo D

    2014-08-01

    Ectotherms constitute the vast majority of terrestrial biodiversity and are especially likely to be vulnerable to climate warming because their basic physiological functions such as locomotion, growth, and reproduction are strongly influenced by environmental temperature. An integrated view about the effects of global warming will be reached not just establishing how the increase in mean temperature impacts the natural populations but also establishing the effects of the increase in temperature variance. One of the molecular responses that are activated in a cell under a temperature stress is the heat shock protein response (HSP). Some studies that have detected consistent differences among thermal treatments and ontogenetic stages in HSP70 expression have assumed that these differences had a genetic basis and consequently expression would be heritable. We tested for changes in quantitative genetic parameters of HSP70 expression in a half-sib design where individuals of the beetle Tenebrio molitor were maintained in constant and varying thermal environments. We estimated heritability of HSP70 expression using a linear mixed modelling approach in different ontogenetic stages. Expression levels of HSP70 were consistently higher in the variable environment and heritability estimates were low to moderate. The results imply that within each ontogenetic stage additive genetic variance was higher in the variable environment and in adults compared with constant environment and larvae stage, respectively. We found that almost all the genetic correlations across ontogenetic stages and environment were positive. These suggest that directional selection for higher levels of expression in one environment will result in higher expression levels of HSP70 on the other environment for the same ontogenetic stage.

  14. Shifting Attention

    Science.gov (United States)

    Ingram, Jenni

    2014-01-01

    This article examines the shifts in attention and focus as one teacher introduces and explains an image that represents the processes involved in a numeric problem that his students have been working on. This paper takes a micro-analytic approach to examine how the focus of attention shifts through what the teacher and students do and say in the…

  15. Shifting Attention

    Science.gov (United States)

    Ingram, Jenni

    2014-01-01

    This article examines the shifts in attention and focus as one teacher introduces and explains an image that represents the processes involved in a numeric problem that his students have been working on. This paper takes a micro-analytic approach to examine how the focus of attention shifts through what the teacher and students do and say in the…

  16. Tough Shift

    DEFF Research Database (Denmark)

    Brewer, Robert S.; Verdezoto, Nervo; Holst, Thomas;

    2015-01-01

    in a student dormitory and found that players did not shift their electricity use, because they were unwilling to change their schedules and found it easier to focus on reducing electricity use. Based on our findings, we discuss the implications for encouraging shifting, and also the challenges of integrating...

  17. Life stage, not climate change, explains observed tree range shifts.

    Science.gov (United States)

    Máliš, František; Kopecký, Martin; Petřík, Petr; Vladovič, Jozef; Merganič, Ján; Vida, Tomáš

    2016-05-01

    Ongoing climate change is expected to shift tree species distribution and therefore affect forest biodiversity and ecosystem services. To assess and project tree distributional shifts, researchers may compare the distribution of juvenile and adult trees under the assumption that differences between tree life stages reflect distributional shifts triggered by climate change. However, the distribution of tree life stages could differ within the lifespan of trees, therefore, we hypothesize that currently observed distributional differences could represent shifts over ontogeny as opposed to climatically driven changes. Here, we test this hypothesis with data from 1435 plots resurveyed after more than three decades across the Western Carpathians. We compared seedling, sapling and adult distribution of 12 tree species along elevation, temperature and precipitation gradients. We analyzed (i) temporal shifts between the surveys and (ii) distributional differences between tree life stages within both surveys. Despite climate warming, tree species distribution of any life stage did not shift directionally upward along elevation between the surveys. Temporal elevational shifts were species specific and an order of magnitude lower than differences among tree life stages within the surveys. Our results show that the observed range shifts among tree life stages are more consistent with ontogenetic differences in the species' environmental requirements than with responses to recent climate change. The distribution of seedlings substantially differed from saplings and adults, while the distribution of saplings did not differ from adults, indicating a critical transition between seedling and sapling tree life stages. Future research has to take ontogenetic differences among life stages into account as we found that distributional differences recently observed worldwide may not reflect climate change but rather the different environmental requirements of tree life stages.

  18. Analysis of ontogenetic spectra of populations of plants and lichens via ordinal regression

    Science.gov (United States)

    Sofronov, G. Yu.; Glotov, N. V.; Ivanov, S. M.

    2015-03-01

    Ontogenetic spectra of plants and lichens tend to vary across the populations. This means that if several subsamples within a sample (or a population) were collected, then the subsamples would not be homogeneous. Consequently, the statistical analysis of the aggregated data would not be correct, which could potentially lead to false biological conclusions. In order to take into account the heterogeneity of the subsamples, we propose to use ordinal regression, which is a type of generalized linear regression. In this paper, we study the populations of cowberry Vaccinium vitis-idaea L. and epiphytic lichens Hypogymnia physodes (L.) Nyl. and Pseudevernia furfuracea (L.) Zopf. We obtain estimates for the proportions of between-sample variability in the total variability of the ontogenetic spectra of the populations.

  19. Extreme Postnatal Scaling in Bat Feeding Performance: A View of Ecomorphology from Ontogenetic and Macroevolutionary Perspectives.

    Science.gov (United States)

    Santana, Sharlene E; Miller, Kimberly E

    2016-09-01

    Ecomorphology studies focus on understanding how anatomical and behavioral diversity result in differences in performance, ecology, and fitness. In mammals, the determinate growth of the skeleton entails that bite performance should change throughout ontogeny until the feeding apparatus attains its adult size and morphology. Then, interspecific differences in adult phenotypes are expected to drive food resource partitioning and patterns of lineage diversification. However, Formal tests of these predictions are lacking for the majority of mammal groups, and thus our understanding of mammalian ecomorphology remains incomplete. By focusing on a fundamental measure of feeding performance, bite force, and capitalizing on the extraordinary morphological and dietary diversity of bats, we discuss how the intersection of ontogenetic and macroevolutionary changes in feeding performance may impact ecological diversity in these mammals. We integrate data on cranial morphology and bite force gathered through longitudinal studies of captive animals and comparative studies of free-ranging individuals. We demonstrate that ontogenetic trajectories and evolutionary changes in bite force are highly dependent on changes in body and head size, and that bats exhibit dramatic, allometric increases in bite force during ontogeny. Interspecific variation in bite force is highly dependent on differences in cranial morphology and function, highlighting selection for ecological specialization. While more research is needed to determine how ontogenetic changes in size and bite force specifically impact food resource use and fitness in bats, interspecific diversity in cranial morphology and bite performance seem to closely match functional differences in diet. Altogether, these results suggest direct ecomorphological relationships at ontogenetic and macroevolutionary scales in bats.

  20. The ontogenetic scaling of hydrodynamics and swimming performance in jellyfish (Aurelia aurita).

    Science.gov (United States)

    McHenry, Matthew J; Jed, Jason

    2003-11-01

    It is not well understood how ontogenetic changes in the motion and morphology of aquatic animals influence the performance of swimming. The goals of the present study were to understand how changes in size, shape and behavior affect the hydrodynamics of jet propulsion in the jellyfish Aurelia aurita and to explore how such changes affect the ontogenetic scaling of swimming speed and cost of transport. We measured the kinematics of jellyfish swimming from video recordings and simulated the hydrodynamics of swimming with two computational models that calculated thrust generation by paddle and jet mechanisms. Our results suggest that thrust is generated primarily by jetting and that there is negligible thrust generation by paddling. We examined how fluid forces scaled with body mass using the jet model. Despite an ontogenetic increase in the range of motion by the bell diameter and a decrease in the height-to-diameter ratio, we found that thrust and acceleration reaction scaled with body mass as predicted by kinematic similarity. However, jellyfish decreased their pulse frequency with growth, and speed consequently scaled at a lower exponential rate than predicted by kinematic similarity. Model simulations suggest that the allometric growth in Aurelia results in swimming that is slower, but more energetically economical, than isometric growth with a prolate bell shape. The decrease in pulse frequency over ontogeny allows large Aurelia medusae to avoid a high cost of transport but generates slower swimming than if they maintained a high pulse frequency. Our findings suggest that ontogenetic change in the height-to-diameter ratio and pulse frequency of Aurelia results in swimming that is relatively moderate in speed but is energetically economical.

  1. Z-scan theory for nonlocal nonlinear media with simultaneous nonlinear refraction and nonlinear absorption.

    Science.gov (United States)

    Rashidian Vaziri, Mohammad Reza

    2013-07-10

    In this paper, the Z-scan theory for nonlocal nonlinear media has been further developed when nonlinear absorption and nonlinear refraction appear simultaneously. To this end, the nonlinear photoinduced phase shift between the impinging and outgoing Gaussian beams from a nonlocal nonlinear sample has been generalized. It is shown that this kind of phase shift will reduce correctly to its known counterpart for the case of pure refractive nonlinearity. Using this generalized form of phase shift, the basic formulas for closed- and open-aperture beam transmittances in the far field have been provided, and a simple procedure for interpreting the Z-scan results has been proposed. In this procedure, by separately performing open- and closed-aperture Z-scan experiments and using the represented relations for the far-field transmittances, one can measure the nonlinear absorption coefficient and nonlinear index of refraction as well as the order of nonlocality. Theoretically, it is shown that when the absorptive nonlinearity is present in addition to the refractive nonlinearity, the sample nonlocal response can noticeably suppress the peak and enhance the valley of the Z-scan closed-aperture transmittance curves, which is due to the nonlocal action's ability to change the beam transverse dimensions.

  2. Chicken-sized oviraptorid dinosaurs from central China and their ontogenetic implications.

    Science.gov (United States)

    Lü, Junchang; Currie, Philip J; Xu, Li; Zhang, Xingliao; Pu, Hanyong; Jia, Songhai

    2013-02-01

    Oviraptorids are a group of specialized non-avian theropod dinosaurs that were generally one to 8 m in body length. New specimens of baby oviraptorids from the Late Cretaceous of Henan Province are some of the smallest individuals known. They include diagnostic characters such as the relative position of the antorbital fenestra and the external naris, distinct opening in the premaxilla anteroventral to the external naris, antorbital fossa partly bordered by premaxilla posterodorsally, lacrimal process of premaxilla does not contact the anterodorsal process of the lacrimal, parietal almost as long as frontal; in dorsal view, posterior margin forms a straight line between the postzygapophyses in each of the fourth and fifth cervicals; femur longer than ilium. They also elucidate the ontogenetic processes of oviraptorids, including fusion of cranial elements and changes in relative body proportions. Hind limb proportions are constant in oviraptorids, regardless of absolute body size or ontogenetic stage. This suggests a sedentary lifestyle that did not involve the pursuit of similar-sized prey. The functional implications for bite force and therefore dietary preferences are better understood through the study of such small animals. The comparison of the measurements of 115 skeletons indicates that oviraptorids maintain their hind limb proportions regardless of ontogenetic stage or absolute size, which is a pattern seen more commonly in herbivores than in carnivores. This may weakly support the hypothesis that oviraptorids are herbivores rather than active carnivores.

  3. Ontogenetic change in molecular and functional properties of blood of garter snakes, Thamnophis sirtalis.

    Science.gov (United States)

    Pough, F H

    1977-07-01

    At birth the blood of garter snakes (Thamnophis sirtalis) has high oxygen affinity (P50 at 25 degrees C and pH 7.2 = 21 mm Hg), low hematocrit (14%), and a high proportion of inactive hemoglobin (44%). Growth of the snakes is accompanied by a steady decrease in oxygen affinity (P50 = 44 mm Hg in adults), increased hematocrit (to 44%), and decreased inactive hemoglobin (to 3%). These adult values are reached in the second or third year of a snake's life. There is a continuous change in the electrophoretically separable hemoglobin fractions during this period, but there is no ontogenetic change in the oxygen affinity of phosphate-stripped hemoglobin in solution. Organic triphosphates do not fully explain the ontogenetic change in blood oxygen affinity. Similar ontogenetic changes in blood oxygen affinity were found in two other species of snakes. The unusual morphology of snake lungs and consequent changes in pulmonary pO2 and pCO2 during ontogeny may be the basis for the unusual relationship between body size and blood oxygen affinity in snakes.

  4. Power Shift

    Institute of Scientific and Technical Information of China (English)

    2008-01-01

    @@ "We are entering a new era of world history: the end of Western domination and the arrival of the Asian century. The question is: will Washington wake up to this reality?" This is the central premise of Kishore Mahbubani's provocative new book The New Asian Hemisphere: The Irresistible Shift of Global Power to the East.

  5. Analyzing algorithms for nonlinear and spatially nonuniform phase shifts in the liquid crystal point diffraction interferometer. 1998 summer research program for high school juniors at the University of Rochester`s Laboratory for Laser Energetics: Student research reports

    Energy Technology Data Exchange (ETDEWEB)

    Jain, N. [Pittsford Sutherland High School, NY (United States)

    1999-03-01

    Phase-shifting interferometry has many advantages, and the phase shifting nature of the Liquid Crystal Point Diffraction Interferometer (LCPDI) promises to provide significant improvement over other current OMEGA wavefront sensors. However, while phase-shifting capabilities improve its accuracy as an interferometer, phase-shifting itself introduces errors. Phase-shifting algorithms are designed to eliminate certain types of phase-shift errors, and it is important to chose an algorithm that is best suited for use with the LCPDI. Using polarization microscopy, the authors have observed a correlation between LC alignment around the microsphere and fringe behavior. After designing a procedure to compare phase-shifting algorithms, they were able to predict the accuracy of two particular algorithms through computer modeling of device-specific phase shift-errors.

  6. Impacts of ontogenetically migrating copepods on downward carbon flux in the western subarctic Pacific Ocean

    Science.gov (United States)

    Kobari, Toru; Steinberg, Deborah K.; Ueda, Ai; Tsuda, Atsushi; Silver, Mary W.; Kitamura, Minoru

    2008-07-01

    To evaluate the impacts of ontogenetically (seasonally) migrating copepods on carbon transport to the mesopelagic zone, we investigated depth distribution, population structure, and feeding activity of the ontogentic copepod community in the western subarctic Pacific Ocean from day-night pairs of zooplankton samples down to 1000 m during the VERtical Transport In the Global Ocean (VERTIGO) program. Over the 31 July-16 August 2005 study period, the biomass of Neocalanus cristatus and Neocalanus plumchrus predominated in the near surface waters, while Neocalanus flemingeri was already dormant at depth. We observed a strong diel migration for Metridia pacifica, and a seasonal downward migration for Eucalanus bungii. Based on gut pigment analysis, ingestion rate of the copepod community was 214-375 mg C m -2 day -1, which was equal to 26-37% of the concurrent primary production. However, comparison of grazing estimated from gut pigments to calculated carbon demand of the copepod community indicates that phytoplankton comprised 37-59% of the ingested carbon. Thus, the copepod community appears to have also relied on detritus and microzooplankton for their nutrition, likely because primary production during this time was dominated by picophytoplankton too small to be grazed by these large copepods. Fecal pellet flux by the copepod community was estimated to account for 141-223% of the sedimentary particulate organic carbon (POC) flux at 150 m, suggesting considerable fragmentation and consumption of pellets in the upper layers. Fecal pellets alone were adequate to meet copepod carbon demand in the surface 0-150 m layer. Active carbon flux by diel migration of M. pacifica (respiration, egestion, and mortality) was 4-17 mg C m -2 day -1, equal to 6-44% of sedimentary POC flux at 150 m. Active carbon flux by N. flemingeri ontogenetic migration (i.e., respiration and mortality at depth) contributed 246 mg C m -2 year -1, equal to 9% of sedimentary POC flux at 1000 m. The

  7. Snake venomics across genus Lachesis. Ontogenetic changes in the venom composition of Lachesis stenophrys and comparative proteomics of the venoms of adult Lachesis melanocephala and Lachesis acrochorda.

    Science.gov (United States)

    Madrigal, Marvin; Sanz, Libia; Flores-Díaz, Marietta; Sasa, Mahmood; Núñez, Vitelbina; Alape-Girón, Alberto; Calvete, Juan J

    2012-12-21

    We report the proteomic analysis of ontogenetic changes in venom composition of the Central American bushmaster, Lachesis stenophrys, and the characterization of the venom proteomes of two congeneric pitvipers, Lachesis melanocephala (black-headed bushmaster) and Lachesis acrochorda (Chochoan bushmaster). Along with the previous characterization of the venom proteome of Lachesis muta muta (from Bolivia), our present outcome enables a comparative overview of the composition and distribution of the toxic proteins across genus Lachesis. Comparative venomics revealed the close kinship of Central American L. stenophrys and L. melanocephala and support the elevation of L. acrochorda to species status. Major ontogenetic changes in the toxin composition of L. stenophrys venom involves quantitative changes in the concentration of vasoactive peptides and serine proteinases, which steadily decrease from birth to adulthood, and age-dependent de novo biosynthesis of Gal-lectin and snake venom metalloproteinases (SVMPs). The net result is a shift from a bradykinin-potentiating and C-type natriuretic peptide (BPP/C-NP)-rich and serine proteinase-rich venom in newborns and 2-years-old juveniles to a (PI>PIII) SVMP-rich venom in adults. Notwithstanding minor qualitative and quantitative differences, the venom arsenals of L. melanocephala and L. acrochorda are broadly similar between themselves and also closely mirror those of adult L. stenophrys and L. muta venoms. The high conservation of the overall composition of Central and South American bushmaster venoms provides the ground for rationalizing the "Lachesis syndrome", characterized by vagal syntomatology, sensorial disorders, hematologic, and cardiovascular manifestations, documented in envenomings by different species of this wide-ranging genus. This finding let us predict that monospecific Lachesic antivenoms may exhibit paraspecificity against all congeneric species. Copyright © 2012 Elsevier B.V. All rights reserved.

  8. Ontogenetic habitat shift, population growth, and burrowing behavior of the Indo-Pacific beach star, Archaster typicus (Echinodermata; Asteroidea).

    Science.gov (United States)

    Bos, Arthur R; Gumanao, Girley S; van Katwijk, Marieke M; Mueller, Benjamin; Saceda, Marjho M; Tejada, Rosie Lynn P

    2011-01-01

    Archaster typicus, a common sea star in Indo-Pacific regions, has been a target for the ornamental trade, even though little is known about its population biology. Spatial and temporal patterns of abundance and size structure of A. typicus were studied in the Davao Gulf, the Philippines (125°42.7'E, 7°0.6'N), from February 2008 to December 2009. Specimens of A. typicus were associated with intertidal mangrove prop roots, seagrass meadows, sandy beaches, and shoals. Among prop roots, specimens were significantly smaller and had highest densities (131 ind. m(-2)) between November and March. High organic matter in sediment and a relatively low predation rate seemed to support juvenile life among mangroves. Size and density analyses provided evidence that individuals gradually move to seagrass, sandy habitats, and shoals as they age. Specimens were significantly larger at a shoal (maximum radius R = 81 mm). New recruits were found between August and November in both 2008 and 2009. Timing of recruitment and population size frequencies confirmed a seasonal reproductive cycle. Juveniles had relatively high growth rates (2-7 mm month(-1)) and may reach an R of 20-25 mm after 1 year. Growth rates of larger specimens (R > 30 mm) were generally stars and provides new biological information as a basis for management of harvested A. typicus populations.

  9. Ontogenetic shifts in male mating preference and morph-specific polyandry in a female colour polymorphic insect

    NARCIS (Netherlands)

    Sanchez-Guillen, Rosa Ana; Hammers, Martijn; Hansson, Bengt; Van Gossum, Hans; Cordero-Rivera, Adolfo; Galicia Mendoza, Dalia Ivette; Wellenreuther, Maren

    2013-01-01

    Background: Sexual conflict over mating rates may favour the origin and maintenance of phenotypes with contrasting reproductive strategies. The damselfly Ischnura elegans is characterised by a female colour polymorphism that consists of one androchrome and two gynochrome female morphs. Previous

  10. 基于非线性滤波估计的北斗B1 QPSK信号跟踪方法%BeiDou B1 quadrature phase shift keying carrier tracking method based on nonlinear Kalman filter

    Institute of Scientific and Technical Information of China (English)

    李荣冰; 黄隽祎; 刘建业; 谢非

    2014-01-01

    Doppler shift and signal power attenuation in the complex environment both can make damage in the accuracy of carrier tracking. Therefore, the non-linear Kalman filter for carrier tracking is designed, which makes correlated observations in the EKF and UKF model based on the analysis of the structure of BeiDou B1 signal. By using measurements from the estimation of filtering in feedback control of the carrier tracking loop, higher and more stable performance can be given in high dynamic and weak signal environments. Finally, the test results show that the feedback control-based EKF and UKF model can perform precise carrier tracking, and make a good limitation of loop error, both of which lead to realization of high performance of signal tracking.%复杂环境下的多普勒频移变化及信号功率衰减均会对载波准确跟踪造成影响。在研究北斗卫星B1频点信号结构的基础上,建立以环路中相关积分值为观测量的非线性EKF模型和UKF模型,并提出利用滤波估计状态量进行状态反馈控制的方法,从而解决了载波跟踪环路在高动态及弱信号环境中难以高性能工作的问题。实验结果表明,状态反馈控制的EKF模型和UKF滤波模型能准确地跟踪弱信号及高动态下的信号变化,从而有效控制跟踪误差,为实现快速准确的载波跟踪奠定了基础。

  11. Ontogenetic investigation of underwater hearing capabilities in loggerhead sea turtles (Caretta caretta) using a dual testing approach.

    Science.gov (United States)

    Lavender, Ashley L; Bartol, Soraya M; Bartol, Ian K

    2014-07-15

    Sea turtles reside in different acoustic environments with each life history stage and may have different hearing capacity throughout ontogeny. For this study, two independent yet complementary techniques for hearing assessment, i.e. behavioral and electrophysiological audiometry, were employed to (1) measure hearing in post-hatchling and juvenile loggerhead sea turtles Caretta caretta (19-62 cm straight carapace length) to determine whether these migratory turtles exhibit an ontogenetic shift in underwater auditory detection and (2) evaluate whether hearing frequency range and threshold sensitivity are consistent in behavioral and electrophysiological tests. Behavioral trials first required training turtles to respond to known frequencies, a multi-stage, time-intensive process, and then recording their behavior when they were presented with sound stimuli from an underwater speaker using a two-response forced-choice paradigm. Electrophysiological experiments involved submerging restrained, fully conscious turtles just below the air-water interface and recording auditory evoked potentials (AEPs) when sound stimuli were presented using an underwater speaker. No significant differences in behavior-derived auditory thresholds or AEP-derived auditory thresholds were detected between post-hatchling and juvenile sea turtles. While hearing frequency range (50-1000/1100 Hz) and highest sensitivity (100-400 Hz) were consistent in audiograms pooled by size class for both behavior and AEP experiments, both post-hatchlings and juveniles had significantly higher AEP-derived than behavior-derived auditory thresholds, indicating that behavioral assessment is a more sensitive testing approach. The results from this study suggest that post-hatchling and juvenile loggerhead sea turtles are low-frequency specialists, exhibiting little differences in threshold sensitivity and frequency bandwidth despite residence in acoustically distinct environments throughout ontogeny.

  12. Fluid Shifts

    Science.gov (United States)

    Stenger, M. B.; Hargens, A. R.; Dulchavsky, S. A.; Arbeille, P.; Danielson, R. W.; Ebert, D. J.; Garcia, K. M.; Johnston, S. L.; Laurie, S. S.; Lee, S. M. C.; Liu, J.; Macias, B.; Martin, D. S.; Minkoff, L.; Ploutz-Snyder, R.; Ribeiro, L. C.; Sargsyan, A.; Smith, S. M.

    2017-01-01

    Introduction. NASA's Human Research Program is focused on addressing health risks associated with long-duration missions on the International Space Station (ISS) and future exploration-class missions beyond low Earth orbit. Visual acuity changes observed after short-duration missions were largely transient, but now more than 50 percent of ISS astronauts have experienced more profound, chronic changes with objective structural findings such as optic disc edema, globe flattening and choroidal folds. These structural and functional changes are referred to as the visual impairment and intracranial pressure (VIIP) syndrome. Development of VIIP symptoms may be related to elevated intracranial pressure (ICP) secondary to spaceflight-induced cephalad fluid shifts, but this hypothesis has not been tested. The purpose of this study is to characterize fluid distribution and compartmentalization associated with long-duration spaceflight and to determine if a relation exists with vision changes and other elements of the VIIP syndrome. We also seek to determine whether the magnitude of fluid shifts during spaceflight, as well as any VIIP-related effects of those shifts, are predicted by the crewmember's pre-flight status and responses to acute hemodynamic manipulations, specifically posture changes and lower body negative pressure. Methods. We will examine a variety of physiologic variables in 10 long-duration ISS crewmembers using the test conditions and timeline presented in the figure below. Measures include: (1) fluid compartmentalization (total body water by D2O, extracellular fluid by NaBr, intracellular fluid by calculation, plasma volume by CO rebreathe, interstitial fluid by calculation); (2) forehead/eyelids, tibia, and calcaneus tissue thickness (by ultrasound); (3) vascular dimensions by ultrasound (jugular veins, cerebral and carotid arteries, vertebral arteries and veins, portal vein); (4) vascular dynamics by MRI (head/neck blood flow, cerebrospinal fluid

  13. Nonlinear Peltier effect in semiconductors

    Science.gov (United States)

    Zebarjadi, Mona; Esfarjani, Keivan; Shakouri, Ali

    2007-09-01

    Nonlinear Peltier coefficient of a doped InGaAs semiconductor is calculated numerically using the Monte Carlo technique. The Peltier coefficient is also obtained analytically for single parabolic band semiconductors assuming a shifted Fermi-Dirac electronic distribution under an applied bias. Analytical results are in agreement with numerical simulations. Key material parameters affecting the nonlinear behavior are doping concentration, effective mass, and electron-phonon coupling. Current density thresholds at which nonlinear behavior is observable are extracted from numerical data. It is shown that the nonlinear Peltier effect can be used to enhance cooling of thin film microrefrigerator devices especially at low temperatures.

  14. Evaluation of long bone surface textures as ontogenetic indicators in centrosaurine ceratopsids.

    Science.gov (United States)

    Tumarkin-Deratzian, Allison R

    2009-09-01

    The search for criteria for aging non-mammalian fossil vertebrates has preoccupied paleobiologists in recent years. Previous studies of the long bones of pterosaurs and modern and subfossil birds as well as of cranial material of centrosaurine ceratopsid dinosaurs have documented variations in surface textures that seem to be ontogenetically related. In this study, long bones from the centrosaurine ceratopsid genera Centrosaurus, Einiosaurus, and Pachyrhinosaurus are examined to test the hypothesis that changes in bone surface textures and reduction of surface porosity could be correlated with size (and presumably age) classes, as has been previously documented in pterosaurs and birds. The data set includes 141 bones representing all six long bone elements, collected from monodominant centrosaurine bone beds. Bone surface patterns are documented by macroscopic visual examination, and a sequence of five texture classes ordered by decreasing surface porosity is described based on the common distributions of these patterns. Calculations of Spearman's rank correlation coefficients reveal significant correlations between texture class and size. The smallest bones are invariably associated with porous midshaft textures that grade to fibrous and long-grained patterns proximally and distally [Texture Class (TC) 1]. Post-hoc analysis after Kruskal-Wallis ANOVA on ranks confirms that the mean size of TC1 bones is, in most cases, significantly different than the mean size of bones in other texture classes. Results of this study suggest the presence of an ontogenetic surface textural signal in centrosaurine long bones; however, comparison of texture classes with size-independent maturity criteria is needed to clarify further the potential ontogenetic significance of higher texture classes.

  15. Nonlinear evolution of drift instabilities

    Energy Technology Data Exchange (ETDEWEB)

    Lee, W.W.; Krommes, J.A.; Oberman, C.R.; Smith, R.A.

    1984-01-01

    The nonlinear evolution of collisionless drift instabilities in a shear-free magnetic field has been studied by means of gyrokinetic particle simulation as well as numerical integration of model mode-coupling equations. The purpose of the investigation is to identify relevant nonlinear mechanisms responsible for the steady-state drift wave fluctuations. It is found that the saturation of the instability is mainly caused by the nonlinear E x B convection of the resonant electrons and their associated velocity space nonlinearity. The latter also induces energy exchange between the competing modes, which, in turn, gives rise to enhanced diffusion. The nonlinear E x B convection of the ions, which contributes to the nonlinear frequency shift, is also an important ingredient for the saturation.

  16. Sizing the Jurassic theropod dinosaur Allosaurus: assessing growth strategy and evolution of ontogenetic scaling of limbs.

    Science.gov (United States)

    Bybee, Paul J; Lee, Andrew H; Lamm, Ellen-Thérèse

    2006-03-01

    Allosaurus is one of the most common Mesozoic theropod dinosaurs. We present a histological analysis to assess its growth strategy and ontogenetic limb bone scaling. Based on an ontogenetic series of humeral, ulnar, femoral, and tibial sections of fibrolamellar bone, we estimate the ages of the largest individuals in the sample to be between 13-19 years. Growth curve reconstruction suggests that maximum growth occurred at 15 years, when body mass increased 148 kg/year. Based on larger bones of Allosaurus, we estimate an upper age limit of between 22-28 years of age, which is similar to preliminary data for other large theropods. Both Model I and Model II regression analyses suggest that relative to the length of the femur, the lengths of the humerus, ulna, and tibia increase in length more slowly than isometry predicts. That pattern of limb scaling in Allosaurus is similar to those in other large theropods such as the tyrannosaurids. Phylogenetic optimization suggests that large theropods independently evolved reduced humeral, ulnar, and tibial lengths by a phyletic reduction in longitudinal growth relative to the femur.

  17. Genetic and ontogenetic variation in an endangered tree structures dependent arthropod and fungal communities.

    Directory of Open Access Journals (Sweden)

    Benjamin J Gosney

    Full Text Available Plant genetic and ontogenetic variation can significantly impact dependent fungal and arthropod communities. However, little is known of the relative importance of these extended genetic and ontogenetic effects within a species. Using a common garden trial, we compared the dependent arthropod and fungal community on 222 progeny from two highly differentiated populations of the endangered heteroblastic tree species, Eucalyptus morrisbyi. We assessed arthropod and fungal communities on both juvenile and adult foliage. The community variation was related to previous levels of marsupial browsing, as well as the variation in the physicochemical properties of leaves using near-infrared spectroscopy. We found highly significant differences in community composition, abundance and diversity parameters between eucalypt source populations in the common garden, and these were comparable to differences between the distinctive juvenile and adult foliage. The physicochemical properties assessed accounted for a significant percentage of the community variation but did not explain fully the community differences between populations and foliage types. Similarly, while differences in population susceptibility to a major marsupial herbivore may result in diffuse genetic effects on the dependent community, this still did not account for the large genetic-based differences in dependent communities between populations. Our results emphasize the importance of maintaining the populations of this rare species as separate management units, as not only are the populations highly genetically structured, this variation may alter the trajectory of biotic colonization of conservation plantings.

  18. Revisiting a model of ontogenetic growth: estimating model parameters from theory and data.

    Science.gov (United States)

    Moses, Melanie E; Hou, Chen; Woodruff, William H; West, Geoffrey B; Nekola, Jeffery C; Zuo, Wenyun; Brown, James H

    2008-05-01

    The ontogenetic growth model (OGM) of West et al. provides a general description of how metabolic energy is allocated between production of new biomass and maintenance of existing biomass during ontogeny. Here, we reexamine the OGM, make some minor modifications and corrections, and further evaluate its ability to account for empirical variation on rates of metabolism and biomass in vertebrates both during ontogeny and across species of varying adult body size. We show that the updated version of the model is internally consistent and is consistent with other predictions of metabolic scaling theory and empirical data. The OGM predicts not only the near universal sigmoidal form of growth curves but also the M(1/4) scaling of the characteristic times of ontogenetic stages in addition to the curvilinear decline in growth efficiency described by Brody. Additionally, the OGM relates the M(3/4) scaling across adults of different species to the scaling of metabolic rate across ontogeny within species. In providing a simple, quantitative description of how energy is allocated to growth, the OGM calls attention to unexplained variation, unanswered questions, and opportunities for future research.

  19. Ontogenetic Variation in Biological Activities of Venoms from Hybrids between Bothrops erythromelas and Bothrops neuwiedi Snakes

    Science.gov (United States)

    Santoro, Marcelo Larami; do Carmo, Thaís; Cunha, Bruna Heloísa Lopes; Alves, André Fonseca; Zelanis, André; Serrano, Solange Maria de Toledo; Grego, Kathleen Fernandes; Sant’Anna, Savio Stefanini; Barbaro, Katia Cristina; Fernandes, Wilson

    2015-01-01

    Lance-headed snakes are found in Central and South America, and they account for most snakebites in Brazil. The phylogeny of South American pitvipers has been reviewed, and the presence of natural and non-natural hybrids between different species of Bothrops snakes demonstrates that reproductive isolation of several species is still incomplete. The present study aimed to analyze the biological features, particularly the thrombin-like activity, of venoms from hybrids born in captivity, from the mating of a female Bothrops erythromelas and a male Bothrops neuwiedi, two species whose venoms are known to display ontogenetic variation. Proteolytic activity on azocoll and amidolytic activity on N-benzoyl-DL-arginine-p-nitroanilide hydrochloride (BAPNA) were lowest when hybrids were 3 months old, and increased over body growth, reaching values similar to those of the father when hybrids were 12 months old. The clotting activity on plasma diminished as hybrids grew; venoms from 3- and 6-months old hybrids showed low clotting activity on fibrinogen (i.e., thrombin-like activity), like the mother venom, and such activity was detected only when hybrids were older than 1 year of age. Altogether, these results point out that venom features in hybrid snakes are genetically controlled during the ontogenetic development. Despite the presence of the thrombin-like enzyme gene(s) in hybrid snakes, they are silenced during the first six months of life. PMID:26714190

  20. Ontogenetic Variation in Biological Activities of Venoms from Hybrids between Bothrops erythromelas and Bothrops neuwiedi Snakes.

    Directory of Open Access Journals (Sweden)

    Marcelo Larami Santoro

    Full Text Available Lance-headed snakes are found in Central and South America, and they account for most snakebites in Brazil. The phylogeny of South American pitvipers has been reviewed, and the presence of natural and non-natural hybrids between different species of Bothrops snakes demonstrates that reproductive isolation of several species is still incomplete. The present study aimed to analyze the biological features, particularly the thrombin-like activity, of venoms from hybrids born in captivity, from the mating of a female Bothrops erythromelas and a male Bothrops neuwiedi, two species whose venoms are known to display ontogenetic variation. Proteolytic activity on azocoll and amidolytic activity on N-benzoyl-DL-arginine-p-nitroanilide hydrochloride (BAPNA were lowest when hybrids were 3 months old, and increased over body growth, reaching values similar to those of the father when hybrids were 12 months old. The clotting activity on plasma diminished as hybrids grew; venoms from 3- and 6-months old hybrids showed low clotting activity on fibrinogen (i.e., thrombin-like activity, like the mother venom, and such activity was detected only when hybrids were older than 1 year of age. Altogether, these results point out that venom features in hybrid snakes are genetically controlled during the ontogenetic development. Despite the presence of the thrombin-like enzyme gene(s in hybrid snakes, they are silenced during the first six months of life.

  1. Ontogenetic mechanisms underlying sexual size dimorphism in Urodele amphibians: An across-species approach

    Institute of Scientific and Technical Information of China (English)

    Lixia ZHANG; Xin LU

    2013-01-01

    Why do two sexes of the same species differ in body size holds a long-standing question of evolutionary biology.While many across-species comparisons have focused on ultimate causes behind sexual size dimorphism (SSD),only have a few been directed toward elucidating its ontogenetic basis.Urodeles are an amphibian group in which the direction and degree of SSD vary greatly among species.Using demographic data yielded by skeletochronology for 33 urodele species,the current study reveals a positive across-species correlation between SSD and the sex difference in mean age of adult animals,and the latter increases with the corresponding difference in age at maturity; annual growth rate does not differ between the sexes.We conclude that extended longevities in one sex,which is mediated by delayed maturation,would allow it to grow for longer and get larger,with growth rate making a weak contribution to body size.The sex-specific divergence in ontogenetic trajectory might be explained by potentially high growth costs of reproduction to females in association with stronger fecundity selection,and to males that are expected to experience stronger sexual selection [Current Zoology 59 (1):142-150,2013].

  2. Ontogenetic Variation in the Thermal Biology of Yarrow's Spiny Lizard, Sceloporus jarrovii.

    Science.gov (United States)

    Gilbert, Anthony L; Lattanzio, Matthew S

    2016-01-01

    Climate change is rapidly altering the way current species interact with their environment to satisfy life-history demands. In areas anticipated to experience extreme warming, rising temperatures are expected to diminish population growth, due either to environmental degradation, or the inability to tolerate novel temperature regimes. Determining how at risk ectotherms, and lizards in particular, are to changes in climate traditionally emphasizes the thermal ecology and thermal sensitivity of physiology of adult members of a population. In this study, we reveal ontogenetic differences in thermal physiological and ecological traits that have been used to anticipate how ectotherms will respond to climate change. We show that the thermal biological traits of juvenile Yarrow's Spiny Lizards (Sceloporus jarrovii) differ from the published estimates of the same traits for adult lizards. Juvenile S. jarrovii differ in their optimal performance temperature, field field-active body temperature, and critical thermal temperatures compared to adult S. jarrovii. Within juvenile S. jarrovii, males and females exhibit differences in field-active body temperature and desiccation tolerance. Given the observed age- and sex-related variation in thermal physiology, we argue that not including physiological differences in thermal biology throughout ontogeny may lead to misinterpretation of patterns of ecological or evolutionary change due to climate warming. Further characterizing the potential for ontogenetic changes in thermal biology would be useful for a more precise and accurate estimation of the role of thermal physiology in mediating population persistence in warmer environments.

  3. Ontogenetic mechanisms underlying sexual size dimorphism in Urodele amphibians: An across-species approach

    Directory of Open Access Journals (Sweden)

    Lixia ZHANG, Xin LU

    2013-02-01

    Full Text Available Why do two sexes of the same species differ in body size holds a long-standing question of evolutionary biology. While many across-species comparisons have focused on ultimate causes behind sexual size dimorphism (SSD, only have a few been directed toward elucidating its ontogenetic basis. Urodeles are an amphibian group in which the direction and degree of SSD vary greatly among species. Using demographic data yielded by skeletochronology for 33 urodele species, the current study reveals a positive across-species correlation between SSD and the sex difference in mean age of adult animals, and the latter increases with the corresponding difference in age at maturity; annual growth rate does not differ between the sexes. We conclude that extended longevities in one sex, which is mediated by delayed maturation, would allow it to grow for longer and get larger, with growth rate making a weak contribution to body size. The sex-specific divergence in ontogenetic trajectory might be explained by potentially high growth costs of reproduction to females in association with stronger fecundity selection, and to males that are expected to experience stronger sexual selection [Current Zoology 59 (1: 142–150, 2013].

  4. Ontogenetic allometry in the foot size of Oligoryzomys flavescens (Waterhouse, 1837 (Rodentia, Sigmodontinae

    Directory of Open Access Journals (Sweden)

    R. Maestri

    Full Text Available Ontogenetic allometry is the study of how the size or shape of certain structures changes over the course of an animal’s development. In this study, using Huxley's formula of allometric growth (1932, we assessed the changes in the rate of growth of the feet size of the sigmodontine rodent Oligoryzomys flavescens during its ontogeny and compared differences between males and females. We find evidence of a change of polarity during the ontogenetic development of the species, with the presence of positive allometry during pregnancy and negative allometry in adulthood. Moreover, we note the presence of sexual dimorphism in the size of the feet, in which males of the species have a higher rate of growth than females. This growth pattern is positively related to escape from predators in childhood in both sexes and, in adulthood, provides a higher encounter rate of females by males, due to the larger displacement of the latter. We suggest that both the forces of natural selection and sexual selection have acted to shape the evolution of foot size in this species.

  5. Ontogenetic Variation in the Thermal Biology of Yarrow's Spiny Lizard, Sceloporus jarrovii.

    Directory of Open Access Journals (Sweden)

    Anthony L Gilbert

    Full Text Available Climate change is rapidly altering the way current species interact with their environment to satisfy life-history demands. In areas anticipated to experience extreme warming, rising temperatures are expected to diminish population growth, due either to environmental degradation, or the inability to tolerate novel temperature regimes. Determining how at risk ectotherms, and lizards in particular, are to changes in climate traditionally emphasizes the thermal ecology and thermal sensitivity of physiology of adult members of a population. In this study, we reveal ontogenetic differences in thermal physiological and ecological traits that have been used to anticipate how ectotherms will respond to climate change. We show that the thermal biological traits of juvenile Yarrow's Spiny Lizards (Sceloporus jarrovii differ from the published estimates of the same traits for adult lizards. Juvenile S. jarrovii differ in their optimal performance temperature, field field-active body temperature, and critical thermal temperatures compared to adult S. jarrovii. Within juvenile S. jarrovii, males and females exhibit differences in field-active body temperature and desiccation tolerance. Given the observed age- and sex-related variation in thermal physiology, we argue that not including physiological differences in thermal biology throughout ontogeny may lead to misinterpretation of patterns of ecological or evolutionary change due to climate warming. Further characterizing the potential for ontogenetic changes in thermal biology would be useful for a more precise and accurate estimation of the role of thermal physiology in mediating population persistence in warmer environments.

  6. Nonlinear Multiantenna Detection Methods

    Directory of Open Access Journals (Sweden)

    Chen Sheng

    2004-01-01

    Full Text Available A nonlinear detection technique designed for multiple-antenna assisted receivers employed in space-division multiple-access systems is investigated. We derive the optimal solution of the nonlinear spatial-processing assisted receiver for binary phase shift keying signalling, which we refer to as the Bayesian detector. It is shown that this optimal Bayesian receiver significantly outperforms the standard linear beamforming assisted receiver in terms of a reduced bit error rate, at the expense of an increased complexity, while the achievable system capacity is substantially enhanced with the advent of employing nonlinear detection. Specifically, when the spatial separation expressed in terms of the angle of arrival between the desired and interfering signals is below a certain threshold, a linear beamformer would fail to separate them, while a nonlinear detection assisted receiver is still capable of performing adequately. The adaptive implementation of the optimal Bayesian detector can be realized using a radial basis function network. Two techniques are presented for constructing block-data-based adaptive nonlinear multiple-antenna assisted receivers. One of them is based on the relevance vector machine invoked for classification, while the other on the orthogonal forward selection procedure combined with the Fisher ratio class-separability measure. A recursive sample-by-sample adaptation procedure is also proposed for training nonlinear detectors based on an amalgam of enhanced -means clustering techniques and the recursive least squares algorithm.

  7. Nonlinear optics

    CERN Document Server

    Bloembergen, Nicolaas

    1996-01-01

    Nicolaas Bloembergen, recipient of the Nobel Prize for Physics (1981), wrote Nonlinear Optics in 1964, when the field of nonlinear optics was only three years old. The available literature has since grown by at least three orders of magnitude.The vitality of Nonlinear Optics is evident from the still-growing number of scientists and engineers engaged in the study of new nonlinear phenomena and in the development of new nonlinear devices in the field of opto-electronics. This monograph should be helpful in providing a historical introduction and a general background of basic ideas both for expe

  8. Femtosecond nonlinear polarization evolution based on cascade quadratic nonlinearities.

    Science.gov (United States)

    Liu, X; Ilday, F O; Beckwitt, K; Wise, F W

    2000-09-15

    We experimentally demonstrate that one can exploit nonlinear phase shifts produced in type I phase-mismatched second-harmonic generation to produce intensity-dependent polarization evolution with 100-fs pulses. An amplitude modulator based on nonlinear polarization rotation provides passive amplitude-modulation depth of up to ~50%. Applications of the amplitude and phase modulations to mode locking of femtosecond bulk and fiber lasers are promising and are discussed.

  9. Nonlinear supratransmission

    Energy Technology Data Exchange (ETDEWEB)

    Geniet, F; Leon, J [Physique Mathematique et Theorique, CNRS-UMR 5825, 34095 Montpellier (France)

    2003-05-07

    A nonlinear system possessing a natural forbidden band gap can transmit energy of a signal with a frequency in the gap, as recently shown for a nonlinear chain of coupled pendulums (Geniet and Leon 2002 Phys. Rev. Lett. 89 134102). This process of nonlinear supratransmission, occurring at a threshold that is exactly predictable in many cases, is shown to have a simple experimental realization with a mechanical chain of pendulums coupled by a coil spring. It is then analysed in more detail. First we go to different (nonintegrable) systems which do sustain nonlinear supratransmission. Then a Josephson transmission line (a one-dimensional array of short Josephson junctions coupled through superconducting wires) is shown to also sustain nonlinear supratransmission, though being related to a different class of boundary conditions, and despite the presence of damping, finiteness, and discreteness. Finally, the mechanism at the origin of nonlinear supratransmission is found to be a nonlinear instability, and this is briefly discussed here.

  10. Effects of competition on root-shoot allocation in Plantago lanceolata L.: adaptive plasticity or ontogenetic drift?

    NARCIS (Netherlands)

    Berendse, F.; Möller, F.

    2009-01-01

    We investigated how shoot and root allocation in plants responds to increasing levels of competitive stress at different levels of soil fertility. In addition, we analyzed whether different responses were due to adaptive plasticity or should be attributed to ontogenetic drift. Plantago lanceolata pl

  11. Entropy squeezing and atomic inversion in the k-photon Jaynes—Cummings model in the presence of the Stark shift and a Kerr medium: A full nonlinear approach

    Science.gov (United States)

    H, R. Baghshahi; M, K. Tavassoly; A, Behjat

    2014-07-01

    The interaction between a two-level atom and a single-mode field in the k-photon Jaynes—Cummings model (JCM) in the presence of the Stark shift and a Kerr medium is studied. All terms in the Hamiltonian, such as the single-mode field, its interaction with the atom, the contribution of the Stark shift and the Kerr medium effects are considered to be f-deformed. In particular, the effect of the initial state of the radiation field on the dynamical evolution of some physical properties such as atomic inversion and entropy squeezing are investigated by considering different initial field states (coherent, squeezed and thermal states).

  12. The ontogenetic origins of mirror neurons: evidence from 'tool-use' and 'audiovisual' mirror neurons.

    Science.gov (United States)

    Cook, Richard

    2012-10-23

    Since their discovery, mirror neurons--units in the macaque brain that discharge both during action observation and execution--have attracted considerable interest. Whether mirror neurons are an innate endowment or acquire their sensorimotor matching properties ontogenetically has been the subject of intense debate. It is widely believed that these units are an innate trait; that we are born with a set of mature mirror neurons because their matching properties conveyed upon our ancestors an evolutionary advantage. However, an alternative view is that mirror neurons acquire their matching properties during ontogeny, through correlated experience of observing and performing actions. The present article re-examines frequently overlooked neurophysiological reports of 'tool-use' and 'audiovisual' mirror neurons within the context of this debate. It is argued that these findings represent compelling evidence that mirror neurons are a product of sensorimotor experience, and not an innate endowment.

  13. Environment-dependent plasticity and ontogenetic changes in the brain of hatchery-reared Atlantic salmon

    DEFF Research Database (Denmark)

    Näslund, J.; Larsen, Martin Hage; Thomassen, S.T.;

    2017-01-01

    enhancement strategies, like environmental enrichment. Here, we investigated the size of the brain in hatcheryreared Atlantic salmon Salmo salar kept at standard (high) and reduced (low) tank densities. In contrast to our predictions, we found that fish reared at high density had larger dry mass of cerebellum......Lowered rearing density has repeatedly been shown to increase the performance of hatchery-reared salmonids stocked into natural environments. One possible mechanism for this pattern could be that lower densities enhance brain development, which has been shown to be the case in other hatchery...... the opposite pattern was observed for telencephalon. Overall, these results reveal substantial brain plasticity depending on the surrounding environment as well as ontogenetic adaptive changes in the brain of the Atlantic salmon...

  14. Practical Nonlinearities

    Science.gov (United States)

    2016-07-01

    Advanced Research Projects Agency (DARPA) Dynamics-Enabled Frequency Sources (DEFYS) program is focused on the convergence of nonlinear dynamics and...Early work in this program has shown that nonlinear dynamics can provide performance advantages. However, the pathway from initial results to...dependent nonlinear stiffness observed in these devices. This work is ongoing, and will continue through the final period of this program . Reference 9

  15. Ontogenetic convergence and evolution of foot morphology in European cave salamanders (Family: Plethodontidae

    Directory of Open Access Journals (Sweden)

    Nistri Annamaria

    2010-07-01

    Full Text Available Abstract Background A major goal in evolutionary biology is to understand the evolution of phenotypic diversity. Both natural and sexual selection play a large role in generating phenotypic adaptations, with biomechanical requirements and developmental mechanisms mediating patterns of phenotypic evolution. For many traits, the relative importance of selective and developmental components remains understudied. Results We investigated ontogenetic trajectories of foot morphology in the eight species of European plethodontid cave salamander to test the hypothesis that adult foot morphology was adapted for climbing. Using geometric morphometrics and other approaches, we found that developmental patterns in five species displayed little morphological change during growth (isometry, where the extensive interdigital webbing in adults was best explained as the retention of the juvenile morphological state. By contrast, three species exhibited significant allometry, with an increase in interdigital webbing during growth. Phylogenetic analyses revealed that multiple evolutionary transitions between isometry and allometry of foot webbing have occurred in this lineage. Allometric parameters of foot growth were most similar to those of a tropical species previously shown to be adapted for climbing. Finally, interspecific variation in adult foot morphology was significantly reduced as compared to variation among juveniles, indicating that ontogenetic convergence had resulted in a common adult foot morphology across species. Conclusions The results presented here provide evidence of a complex history of phenotypic evolution in this clade. The common adult phenotype exhibited among species reveals that selection plays an important part in generating patterns of foot diversity in the group. However, developmental trajectories arriving at this common morphology are distinct; with some species displaying developmental stasis (isometry, while others show an increase

  16. Ontogenetic scaling patterns and functional anatomy of the pelvic limb musculature in emus (Dromaius novaehollandiae

    Directory of Open Access Journals (Sweden)

    Luis P. Lamas

    2014-12-01

    Full Text Available Emus (Dromaius novaehollandiae are exclusively terrestrial, bipedal and cursorial ratites with some similar biomechanical characteristics to humans. Their growth rates are impressive, as their body mass increases eighty-fold from hatching to adulthood whilst maintaining the same mode of locomotion throughout life. These ontogenetic characteristics stimulate biomechanical questions about the strategies that allow emus to cope with their rapid growth and locomotion, which can be partly addressed via scaling (allometric analysis of morphology. In this study we have collected pelvic limb anatomical data (muscle architecture, tendon length, tendon mass and bone lengths and calculated muscle physiological cross sectional area (PCSA and average tendon cross sectional area from emus across three ontogenetic stages (n = 17, body masses from 3.6 to 42 kg. The data were analysed by reduced major axis regression to determine how these biomechanically relevant aspects of morphology scaled with body mass. Muscle mass and PCSA showed a marked trend towards positive allometry (26 and 27 out of 34 muscles respectively and fascicle length showed a more mixed scaling pattern. The long tendons of the main digital flexors scaled with positive allometry for all characteristics whilst other tendons demonstrated a less clear scaling pattern. Finally, the two longer bones of the limb (tibiotarsus and tarsometatarsus also exhibited positive allometry for length, and two others (femur and first phalanx of digit III had trends towards isometry. These results indicate that emus experience a relative increase in their muscle force-generating capacities, as well as potentially increasing the force-sustaining capacities of their tendons, as they grow. Furthermore, we have clarified anatomical descriptions and provided illustrations of the pelvic limb muscle–tendon units in emus.

  17. Ecological interactions in dinosaur communities: influences of small offspring and complex ontogenetic life histories.

    Science.gov (United States)

    Codron, Daryl; Carbone, Chris; Clauss, Marcus

    2013-01-01

    Because egg-laying meant that even the largest dinosaurs gave birth to very small offspring, they had to pass through multiple ontogenetic life stages to adulthood. Dinosaurs' successors as the dominant terrestrial vertebrate life form, the mammals, give birth to live young, and have much larger offspring and less complex ontogenetic histories. The larger number of juveniles in dinosaur as compared to mammal ecosystems represents both a greater diversity of food available to predators, and competitors for similar-sized individuals of sympatric species. Models of population abundances across different-sized species of dinosaurs and mammals, based on simulated ecological life tables, are employed to investigate how differences in predation and competition pressure influenced dinosaur communities. Higher small- to medium-sized prey availability leads to a normal body mass-species richness (M-S) distribution of carnivorous dinosaurs (as found in the theropod fossil record), in contrast to the right-skewed M-S distribution of carnivorous mammals (as found living members of the order Carnivora). Higher levels of interspecific competition leads to a left-skewed M-S distribution in herbivorous dinosaurs (as found in sauropods and ornithopods), in contrast to the normal M-S distribution of large herbivorous mammals. Thus, our models suggest that differences in reproductive strategy, and consequently ontogeny, explain observed differences in community structure between dinosaur and mammal faunas. Models also show that the largest dinosaurian predators could have subsisted on similar-sized prey by including younger life stages of the largest herbivore species, but that large predators likely avoided prey much smaller than themselves because, despite predicted higher abundances of smaller than larger-bodied prey, contributions of small prey to biomass intake would be insufficient to satisfy meat requirements. A lack of large carnivores feeding on small prey exists in mammals

  18. Scaling of feeding biomechanics in the horn shark Heterodontus francisci: ontogenetic constraints on durophagy.

    Science.gov (United States)

    Kolmann, Matthew A; Huber, Daniel R

    2009-01-01

    Organismal performance changes over ontogeny as the musculoskeletal systems underlying animal behavior grow in relative size and shape. As performance is a determinant of feeding ecology, ontogenetic changes in the former can influence the latter. The horn shark Heterodontus francisci consumes hard-shelled benthic invertebrates, which may be problematic for younger animals with lower performance capacities. Scaling of feeding biomechanics was investigated in H. francisci (n=16, 19-59cm standard length (SL)) to determine the biomechanical basis of allometric changes in feeding performance and whether this performance capacity constrains hard-prey consumption over ontogeny. Positive allometry of anterior (8-163N) and posterior (15-382N) theoretical bite force was attributed to positive allometry of cross-sectional area in two jaw adducting muscles and mechanical advantage at the posterior bite point (0.79-1.26). Mechanical advantage for anterior biting scaled isometrically (0.52). Fracture forces for purple sea urchins Strongylocentrotus purpuratus consumed by H. francisci ranged from 24 to 430N. Comparison of these fracture forces to the bite force of H. francisci suggests that H. francisci is unable to consume hard prey early in its life history, but can consume the majority of S. purpuratus by the time it reaches maximum size. Despite this constraint, positive allometry of biting performance appears to facilitate an earlier entry into the durophagous niche than would an isometric ontogenetic trajectory. The posterior gape of H. francisci is significantly smaller than the urchins capable of being crushed by its posterior bite force. Thus, the high posterior bite forces of H. francisci cannot be fully utilized while consuming prey of similar toughness and size to S. purpuratus, and its potential trophic niche is primarily determined by anterior biting capacity.

  19. Prickly poppies can get pricklier: ontogenetic patterns in the induction of physical defense traits.

    Science.gov (United States)

    Hoan, Ryan P; Ormond, Rhys A; Barton, Kasey E

    2014-01-01

    Plant ontogeny is a common source of variation in defense and herbivory. Yet, few studies have investigated how the induction of physical defense traits changes across plant ontogeny. Physical defense traits are costly to produce, and thus, it was predicted that induction as a cost-saving strategy would be particularly favorable for seedlings, leading to ontogenetic declines in the inducibility of these traits. We tested for induction of three different physical defense traits (prickles, latex and leaf toughness) in response to mechanical defoliation and jasmonic acid application using prickly poppies (Argemone glauca and A. mexicana, Papaveraceae) as a model system. Genetic variation in the induction of physical defenses was tested using maternal sib-ships sampled from multiple populations. Both species induced higher densities of laminar prickles, although the magnitude of induction was much higher in the endemic Hawaiian prickly poppy, A. glauca, than in the cosmopolitan A. mexicana. The magnitude of prickle induction was also higher in young compared to older juvenile plant stages in A. glauca, demonstrating a strong role of ontogeny. Neither latex exudation nor leaf toughness was induced in either species. Although significant genetic variation was detected within and among populations for constitutive expression of physical defense traits in Argemone, there was no evidence for genetic variation in the induction of these traits. This study provides the first evidence for the induction of physical defenses in prickly poppies, emphasizing how an ontogenetically explicit framework can reveal new insights into plant defense. Moreover, this study illustrates how sister species comparisons between island vs. continental plants can provide new insights into plant functional and evolutionary ecology, highlighting a fruitful area for future research on more species pairs.

  20. Ecological interactions in dinosaur communities: influences of small offspring and complex ontogenetic life histories.

    Directory of Open Access Journals (Sweden)

    Daryl Codron

    Full Text Available Because egg-laying meant that even the largest dinosaurs gave birth to very small offspring, they had to pass through multiple ontogenetic life stages to adulthood. Dinosaurs' successors as the dominant terrestrial vertebrate life form, the mammals, give birth to live young, and have much larger offspring and less complex ontogenetic histories. The larger number of juveniles in dinosaur as compared to mammal ecosystems represents both a greater diversity of food available to predators, and competitors for similar-sized individuals of sympatric species. Models of population abundances across different-sized species of dinosaurs and mammals, based on simulated ecological life tables, are employed to investigate how differences in predation and competition pressure influenced dinosaur communities. Higher small- to medium-sized prey availability leads to a normal body mass-species richness (M-S distribution of carnivorous dinosaurs (as found in the theropod fossil record, in contrast to the right-skewed M-S distribution of carnivorous mammals (as found living members of the order Carnivora. Higher levels of interspecific competition leads to a left-skewed M-S distribution in herbivorous dinosaurs (as found in sauropods and ornithopods, in contrast to the normal M-S distribution of large herbivorous mammals. Thus, our models suggest that differences in reproductive strategy, and consequently ontogeny, explain observed differences in community structure between dinosaur and mammal faunas. Models also show that the largest dinosaurian predators could have subsisted on similar-sized prey by including younger life stages of the largest herbivore species, but that large predators likely avoided prey much smaller than themselves because, despite predicted higher abundances of smaller than larger-bodied prey, contributions of small prey to biomass intake would be insufficient to satisfy meat requirements. A lack of large carnivores feeding on small prey

  1. Ontogenetic influence on neural spine bifurcation in Diplodocoidea (Dinosauria: Sauropoda): a critical phylogenetic character.

    Science.gov (United States)

    Woodruff, D Cary; Fowler, Denver W

    2012-07-01

    Within Diplodocoidea (Dinosauria: Sauropoda), phylogenetic position of the three subclades Rebbachisauridae, Dicraeosauridae, and Diplodocidae is strongly influenced by a relatively small number of characters. Neural spine bifurcation, especially within the cervical vertebrae, is considered to be a derived character, with taxa that lack this feature regarded as relatively basal. Our analysis of dorsal and cervical vertebrae from small-sized diplodocoids (representing at least 18 individuals) reveals that neural spine bifurcation is less well developed or absent in smaller specimens. New preparation of the roughly 200-cm long diplodocid juvenile Sauriermuseum Aathal 0009 reveals simple nonbifurcated cervical neural spines, strongly reminiscent of more basal sauropods such as Omeisaurus. An identical pattern of ontogenetically linked bifurcation has also been observed in several specimens of the basal macronarian Camarasaurus, suggesting that this is characteristic of several clades of Sauropoda. We suggest that neural spine bifurcation performs a biomechanical function related to horizontal positioning of the neck that may become significant only at the onset of a larger body size, hence, its apparent absence or weaker development in smaller specimens. These results have significant implications for the taxonomy and phylogenetic position of taxa described from specimens of small body size. On the basis of shallow bifurcation of its cervical and dorsal neural spines, the small diplodocid Suuwassea is more parsimoniously interpreted as an immature specimen of an already recognized diplodocid taxon. Our findings emphasize the view that nonmature dinosaurs often exhibit morphologies more similar to their ancestral state and may therefore occupy a more basal position in phylogenetic analyses than would mature specimens of the same species. In light of this, we stress the need for phylogenetic reanalysis of sauropod clades where vital characters may be ontogenetically

  2. Ontogenetic changes in limb bone structural proportions in mountain gorillas (Gorilla beringei beringei).

    Science.gov (United States)

    Ruff, Christopher B; Burgess, M Loring; Bromage, Timothy G; Mudakikwa, Antoine; McFarlin, Shannon C

    2013-12-01

    Behavioral studies indicate that adult mountain gorillas (Gorilla beringei) are the most terrestrial of all nonhuman hominoids, but that infant mountain gorillas are much more arboreal. Here we examine ontogenetic changes in diaphyseal strength and length of the femur, tibia, humerus, radius, and ulna in 30 Virunga mountain gorillas, including 18 immature specimens and 12 adults. Comparisons are also made with 14 adult western lowland gorillas (Gorilla gorilla gorilla), which are known to be more arboreal than adult mountain gorillas. Infant mountain gorillas have significantly stronger forelimbs relative to hind limbs than older juveniles and adults, but are nonsignificantly different from western lowland gorilla adults. The change in inter-limb strength proportions is abrupt at about two years of age, corresponding to the documented transition to committed terrestrial quadrupedalism in mountain gorillas. The one exception is the ulna, which shows a gradual increase in strength relative to the radius and other long bones during development, possibly corresponding to the gradual adoption of stereotypical fully pronated knuckle-walking in older juvenile gorillas. Inter-limb bone length proportions show a contrasting developmental pattern, with hind limb/forelimb length declining rapidly from birth to five months of age, and then showing no consistent change through adulthood. The very early change in length proportions, prior to significant independent locomotion, may be related to the need for relatively long forelimbs for climbing in a large-bodied hominoid. Virunga mountain gorilla older juveniles and adults have equal or longer forelimb relative to hind limb bones than western lowland adults. These findings indicate that both ontogenetically and among closely related species of Gorilla, long bone strength proportions better reflect actual locomotor behavior than bone length proportions.

  3. Prickly poppies can get pricklier: ontogenetic patterns in the induction of physical defense traits.

    Directory of Open Access Journals (Sweden)

    Ryan P Hoan

    Full Text Available Plant ontogeny is a common source of variation in defense and herbivory. Yet, few studies have investigated how the induction of physical defense traits changes across plant ontogeny. Physical defense traits are costly to produce, and thus, it was predicted that induction as a cost-saving strategy would be particularly favorable for seedlings, leading to ontogenetic declines in the inducibility of these traits. We tested for induction of three different physical defense traits (prickles, latex and leaf toughness in response to mechanical defoliation and jasmonic acid application using prickly poppies (Argemone glauca and A. mexicana, Papaveraceae as a model system. Genetic variation in the induction of physical defenses was tested using maternal sib-ships sampled from multiple populations. Both species induced higher densities of laminar prickles, although the magnitude of induction was much higher in the endemic Hawaiian prickly poppy, A. glauca, than in the cosmopolitan A. mexicana. The magnitude of prickle induction was also higher in young compared to older juvenile plant stages in A. glauca, demonstrating a strong role of ontogeny. Neither latex exudation nor leaf toughness was induced in either species. Although significant genetic variation was detected within and among populations for constitutive expression of physical defense traits in Argemone, there was no evidence for genetic variation in the induction of these traits. This study provides the first evidence for the induction of physical defenses in prickly poppies, emphasizing how an ontogenetically explicit framework can reveal new insights into plant defense. Moreover, this study illustrates how sister species comparisons between island vs. continental plants can provide new insights into plant functional and evolutionary ecology, highlighting a fruitful area for future research on more species pairs.

  4. Long bone histology of the subterranean rodent Bathyergus suillus (Bathyergidae): ontogenetic pattern of cortical bone thickening.

    Science.gov (United States)

    Montoya-Sanhueza, Germán; Chinsamy, Anusuya

    2017-02-01

    Patterns of bone development in mammals are best known from terrestrial and cursorial groups, but there is a considerable gap in our understanding of how specializations for life underground affect bone growth and development. Likewise, studies of bone microstructure in wild populations are still scarce, and they often include few individuals and tend to be focused on adults. For these reasons, the processes generating bone microstructural variation at intra- and interspecific levels are not fully understood. This study comprehensively examines the bone microstructure of an extant population of Cape dune molerats, Bathyergus suillus (Bathyergidae), the largest subterranean mammal endemic to the Western Cape of South Africa. The aim of this study is to investigate the postnatal bone growth of B. suillus using undecalcified histological sections (n = 197) of the femur, humerus, tibia-fibula, ulna and radius, including males and females belonging to different ontogenetic and reproductive stages (n = 42). Qualitative histological features demonstrate a wide histodiversity with thickening of the cortex mainly resulting from endosteal and periosteal bone depositions, whilst there is scarce endosteal resorption and remodeling throughout ontogeny. This imbalanced bone modeling allows the tissues deposited during ontogeny to remain relatively intact, thus preserving an excellent record of growth. The distribution of the different bone tissues observed in the cortex depends on ontogenetic status, anatomical features (e.g. muscle attachment structures) and location on the bone (e.g. anterior or lateral). The type of bone microstructure and modeling is discussed in relation to digging behavior, reproduction and physiology of this species. This study is the first histological assessment describing the process of cortical thickening in long bones of a fossorial mammal.

  5. Ontogenetically-regulated male sterility in tissue culture - induced and spontaneous sorghum mutants

    Directory of Open Access Journals (Sweden)

    Elkonin L.A.

    2003-01-01

    Full Text Available Variability of male fertility expression in the AS-1 line, a somaclonal variant obtained from tissue culture of CMS-plant, and in the progeny of revenant '124-1' obtained from fertile tiller, which developed on CMS-plant transferred from the field to the greenhouse, was investigated. Both revertants were characterized by similar expression of male fertility during plant ontogenesis: the panicle on the main tiller was almost completely sterile whereas formation of fertile pollen grains and seed set were observed on the panicles of the shoot tillers. A clear basipetal gradient of male fertility was manifested on all panicles: the base had significantly higher per cent of fertile pollen grains in comparison with the middle part, while in the top the anthers were either absent or had few sterile pollen grains. Such an ontogenetically-regulated restoration of male fertility was controlled by nuclear genes and could be transferred through the pollen in crosses with progenitor CMS-line. Growing of AS-1 plants in the growth chambers simultaneously under a long (16/8 and a short (12/12 daylength conditions demonstrated that differences of fertility level in different tillers was not caused by change of photoperiod during plant ontogenesis and functioning of photoperiod-sensitive fertility restoring gene. Whereas, the ontogenetically-regulated expression of male fertility in both revenants was temperature-dependent and was clearly manifested under relatively cool conditions during 2-week period before the beginning of anthesis of the first panicle (average daily temperature 21°C. The increase of the average daily temperature by 2-3 С resulted in sharp increase of male fertility level. Possibility of using AS-1 line in a new "two-line system" of hybrid seed production, which require only two lines (sterile mutant and fertility restorer, is discussed.

  6. Nonlinear effects in Thomson backscattering

    Science.gov (United States)

    Maroli, C.; Petrillo, V.; Tomassini, P.; Serafini, L.

    2013-03-01

    We analyze the nonlinear classical effects of the X/γ radiation produced by Thomson/Compton sources. We confirm the development of spectral fringes of the radiation on axis, which comports broadening, shift, and deformation of the spectrum. For the nominal parameters of the SPARC-LAB Thomson scattering and of the European Proposal for the gamma source ELI-NP, however, the radiation, when collected in the suitable acceptance angle, does not reveal many differences from that predicted by the linear model and the nonlinear redshift is subdominant with respect to the quantum recoil. An experiment aimed to the study of the nonlinearities is proposed on the SPARC-LAB source.

  7. Nonlinear oscillations

    CERN Document Server

    Nayfeh, Ali Hasan

    1995-01-01

    Nonlinear Oscillations is a self-contained and thorough treatment of the vigorous research that has occurred in nonlinear mechanics since 1970. The book begins with fundamental concepts and techniques of analysis and progresses through recent developments and provides an overview that abstracts and introduces main nonlinear phenomena. It treats systems having a single degree of freedom, introducing basic concepts and analytical methods, and extends concepts and methods to systems having degrees of freedom. Most of this material cannot be found in any other text. Nonlinear Oscillations uses sim

  8. Nonlinear Science

    CERN Document Server

    Yoshida, Zensho

    2010-01-01

    This book gives a general, basic understanding of the mathematical structure "nonlinearity" that lies in the depths of complex systems. Analyzing the heterogeneity that the prefix "non" represents with respect to notions such as the linear space, integrability and scale hierarchy, "nonlinear science" is explained as a challenge of deconstruction of the modern sciences. This book is not a technical guide to teach mathematical tools of nonlinear analysis, nor a zoology of so-called nonlinear phenomena. By critically analyzing the structure of linear theories, and cl

  9. Nonlinear analysis

    CERN Document Server

    Nanda, Sudarsan

    2013-01-01

    "Nonlinear analysis" presents recent developments in calculus in Banach space, convex sets, convex functions, best approximation, fixed point theorems, nonlinear operators, variational inequality, complementary problem and semi-inner-product spaces. Nonlinear Analysis has become important and useful in the present days because many real world problems are nonlinear, nonconvex and nonsmooth in nature. Although basic concepts have been presented here but many results presented have not appeared in any book till now. The book could be used as a text for graduate students and also it will be useful for researchers working in this field.

  10. Hydrogen isotope effect on the Dimits shift

    Science.gov (United States)

    Itoh, S.-I.; Itoh, K.

    2016-10-01

    The hydrogen isotope effect on the Dimits shift in drift wave turbulence (Dimits et al 2000 Phys. Plasmas 7 969) is discussed using the theory of zonal flows, in which the nonlinear damping rate of zonal flows is taken into account. The up-shift of the critical linear growth rate of the drift waves, above which drift wave fluctuations develop, is investigated. The dependence on the mass number of the hydrogen isotope is discussed.

  11. Nonlinear theory of kinetic instabilities near threshold

    Energy Technology Data Exchange (ETDEWEB)

    Berk, H.L.; Pekker, M.S. [Univ. of Texas, Austin, TX (United States). Inst. for Fusion Studies; Breizman, B.N. [Texas Univ., Austin, TX (United States). Inst. for Fusion Studies]|[Budker Inst. of Nuclear Physics, Novosibirsk (Russian Federation)

    1997-05-01

    A new nonlinear equation has been derived and solved for the evolution of an unstable collective mode in a kinetic system close to the threshold of linear instability. The resonant particle response produces the dominant nonlinearity, which can be calculated iteratively in the near-threshold regime as long as the mode doe snot trap resonant particles. With sources and classical relaxation processes included, the theory describes both soft nonlinear regimes, where the mode saturation level is proportional to an increment above threshold, and explosive nonlinear regimes, where the mode grows to a level that is independent of the closeness to threshold. The explosive solutions exhibit mode frequency shifting. For modes that exist in the absence of energetic particles, the frequency shift is both upward and downward. For modes that require energetic particles for their existence, there is a preferred direction of the frequency shift. The frequency shift continues even after the mode traps resonant particles.

  12. Non-linear scaling of oxygen consumption and heart rate in a very large cockroach species (Gromphadorhina portentosa): correlated changes with body size and temperature.

    Science.gov (United States)

    Streicher, Jeffrey W; Cox, Christian L; Birchard, Geoffrey F

    2012-04-01

    Although well documented in vertebrates, correlated changes between metabolic rate and cardiovascular function of insects have rarely been described. Using the very large cockroach species Gromphadorhina portentosa, we examined oxygen consumption and heart rate across a range of body sizes and temperatures. Metabolic rate scaled positively and heart rate negatively with body size, but neither scaled linearly. The response of these two variables to temperature was similar. This correlated response to endogenous (body mass) and exogenous (temperature) variables is likely explained by a mutual dependence on similar metabolic substrate use and/or coupled regulatory pathways. The intraspecific scaling for oxygen consumption rate showed an apparent plateauing at body masses greater than about 3 g. An examination of cuticle mass across all instars revealed isometric scaling with no evidence of an ontogenetic shift towards proportionally larger cuticles. Published oxygen consumption rates of other Blattodea species were also examined and, as in our intraspecific examination of G. portentosa, the scaling relationship was found to be non-linear with a decreasing slope at larger body masses. The decreasing slope at very large body masses in both intraspecific and interspecific comparisons may have important implications for future investigations of the relationship between oxygen transport and maximum body size in insects.

  13. On the nymphs of lantana lace bug Teleonemia scrupulosa Stål (Hemiptera: Heteroptera: Tingidae: Tinginae): ontogenetic features of integumentary structures highlighted.

    Science.gov (United States)

    Guidoti, Marcus; Barcellos, Aline

    2013-02-11

    The five instars of Teleonemia scrupulosa Stål are described. The postembryonic ontogenetic development of integumentary structures is emphasized, with a discussion on its potential use in cladistic studies within Tingidae.

  14. Goos-Hänchen shifts in harmonic generation from metals.

    Science.gov (United States)

    Yallapragada, V J; Gopal, Achanta Venu; Agarwal, G S

    2013-05-06

    We present the first calculation of the Goos-Hänchen shifts in the context of the nonlinear generation of fields. We specifically concentrate on shifts of second harmonic generated at metallic surfaces. At metallic surfaces the second harmonic primarily arises from discontinuities of the field at surfaces which not only result in large harmonic generation but also in significant Goos-Hänchen shifts of the generated second harmonic. Our results can be extended to other shifts like angular shifts and Fedorov-Imbert shifts.

  15. A NONLINEAR FEASIBILITY PROBLEM HEURISTIC

    Directory of Open Access Journals (Sweden)

    Sergio Drumond Ventura

    2015-04-01

    Full Text Available In this work we consider a region S ⊂ given by a finite number of nonlinear smooth convex inequalities and having nonempty interior. We assume a point x 0 is given, which is close in certain norm to the analytic center of S, and that a new nonlinear smooth convex inequality is added to those defining S (perturbed region. It is constructively shown how to obtain a shift of the right-hand side of this inequality such that the point x 0 is still close (in the same norm to the analytic center of this shifted region. Starting from this point and using the theoretical results shown, we develop a heuristic that allows us to obtain the approximate analytic center of the perturbed region. Then, we present a procedure to solve the problem of nonlinear feasibility. The procedure was implemented and we performed some numerical tests for the quadratic (random case.

  16. Nonlinear Reconstruction

    CERN Document Server

    Zhu, Hong-Ming; Pen, Ue-Li; Chen, Xuelei; Yu, Hao-Ran

    2016-01-01

    We present a direct approach to non-parametrically reconstruct the linear density field from an observed non-linear map. We solve for the unique displacement potential consistent with the non-linear density and positive definite coordinate transformation using a multigrid algorithm. We show that we recover the linear initial conditions up to $k\\sim 1\\ h/\\mathrm{Mpc}$ with minimal computational cost. This reconstruction approach generalizes the linear displacement theory to fully non-linear fields, potentially substantially expanding the BAO and RSD information content of dense large scale structure surveys, including for example SDSS main sample and 21cm intensity mapping.

  17. Nonlinear optics

    CERN Document Server

    Boyd, Robert W

    2013-01-01

    Nonlinear Optics is an advanced textbook for courses dealing with nonlinear optics, quantum electronics, laser physics, contemporary and quantum optics, and electrooptics. Its pedagogical emphasis is on fundamentals rather than particular, transitory applications. As a result, this textbook will have lasting appeal to a wide audience of electrical engineering, physics, and optics students, as well as those in related fields such as materials science and chemistry.Key Features* The origin of optical nonlinearities, including dependence on the polarization of light* A detailed treatment of the q

  18. Prolixus (Acari: Trombidiformes: Tenuipalpidae) newly recorded from New Zealand: A new species from Cyperaceae and its ontogenetic patterns in chaetotaxy.

    Science.gov (United States)

    Xu, Yun; Zhang, Zhi-Qiang

    2014-12-19

    The genus Prolixus (Acari: Tenuipalpidae) was represented by two species from Australian sedges prior to this study. A new species, Prolixus meyerae sp. nov., is here described and illustrated from leaves of Gahnia (Cyperaceae) in Auckland, New Zealand. In this paper, we present the ontogenetic additions in idiosomal and leg chaetotaxy from larva to adult. A key to world species of Prolixus is also proposed.

  19. Ontogenetic change of body color patterns in laboratory-raised juveniles of six terrestrial hermit crab species.

    Science.gov (United States)

    Hamasaki, Katsuyuki; Tsuru, Takuma; Sanda, Tetsuya; Fujikawa, Shunsuke; Dan, Shigeki; Kitada, Shuichi

    2017-01-30

    We examined the ontogenetic change of body color patterns in the laboratory-raised juveniles of six terrestrial hermit crab species, including Birgus latro, Coenobita brevimanus, C. cavipes, C. purpureus, C. rugosus, and C. violascens, which commonly occur in the southern islands, Japan. The body color patterns of coenobitid juveniles were species-specific. The diagnostic features of body color patterns enable identification of juveniles of coenobitid crab species in the wild, thereby helping to understand the precise habitats of each coenobitid species.

  20. Precocial hindlimbs and altricial forelimbs: partitioning ontogenetic strategies in mallards (Anas platyrhynchos).

    Science.gov (United States)

    Dial, Terry R; Carrier, David R

    2012-11-01

    Precocial development, in which juveniles are relatively mature at hatching or birth, is more common among vertebrates than altricial development, and is likely to be the basal condition. Altricial development characterizes many birds and mammals and is generally viewed as an alternate strategy, promoting fast growth rates, short developmental periods and relatively poor locomotor performance prior to attaining adult size. Many aquatic birds such as Anseriformes (ducks, geese and swans), Charadriformes (gulls and terns) and Gruiformes (rails) undergo distinctive developmental trajectories, in that hatchlings are able to run and swim the day they hatch, yet they do not begin to fly until fully grown. We hypothesized that there should be tradeoffs in apportioning bone and muscle mass to the hindlimb and forelimb that could account for these patterns in locomotor behavior within the mallard (Anas platyrhynchos). Growth of the musculoskeletal system in the forelimbs and hindlimbs was measured and compared with maximal aquatic and terrestrial sprint speeds and aerial descent rates throughout the 2-month-long mallard ontogenetic period. At 30 days post hatching, when body mass is 50% of adult values, hindlimb muscle mass averages 90% and forelimb muscle mass averages 10% of adult values; similarly, bone growth (length and width) in the hindlimbs and forelimbs averages 90 and 60% of adult values, respectively. The attainment of mallard locomotor performance parallels the morphological maturation of forelimb and hindlimb morphometrics - hindlimb performance initiates just after hatching at a relatively high level (~50% adult values) and gradually improves throughout the first month of development, while forelimb performance is relatively non-existent at hatching (~10% adult values), experiencing delayed and dramatic improvement in function, and maturing at the time of fledging. This divergence in ontogenetic strategy between locomotor modules could allow developing

  1. Brief communication: Endocranial volumes in an ontogenetic sample of chimpanzees from the Taï Forest National Park, Ivory Coast.

    Science.gov (United States)

    Neubauer, Simon; Gunz, Philipp; Schwarz, Uta; Hublin, Jean-Jacques; Boesch, Christophe

    2012-02-01

    Ontogenetic samples of endocranial volumes (EVs) from great apes and humans are critical for understanding the evolution of the brain growth pattern in the hominin lineage. However, high quality ontogenetic data are scarce, especially for nonhuman primates. Here, we provide original data derived from an osteological collection of a wild population of Pan troglodytes verus from the Taï Forest National Park, Ivory Coast. This sample is unique, because age, sex, and pedigree information are available for many specimens from behavioral observations in the wild. We scanned crania of all 30 immature specimens and 13 adult individuals using high-resolution computed tomography. We then created virtual casts of the bony braincase (endocasts) to measure EVs. We also measured cranial length, width, and height and attempted to relate cranial distances to EV via regression analysis. Our data are consistent with previous studies. The only neonate in the sample has an EV of 127 cm(3) or 34% of the adult mean. EV increases rapidly during early ontogeny. The average adult EV in this sample is 378.7 ± 30.1 cm(3) . We found sexual dimorphism in adults; males seem to be already larger than females before adult EV is attained. Regressions on cranial width and multiple regression provide better estimates for EV than regressions on cranial length or height. Increasing the sample size and compiling more high quality ontogenetic data of EV will help to reconcile ongoing discussions about the evolution of hominin brain growth. Copyright © 2011 Wiley Periodicals, Inc.

  2. Nonlinear optimization

    CERN Document Server

    Ruszczynski, Andrzej

    2011-01-01

    Optimization is one of the most important areas of modern applied mathematics, with applications in fields from engineering and economics to finance, statistics, management science, and medicine. While many books have addressed its various aspects, Nonlinear Optimization is the first comprehensive treatment that will allow graduate students and researchers to understand its modern ideas, principles, and methods within a reasonable time, but without sacrificing mathematical precision. Andrzej Ruszczynski, a leading expert in the optimization of nonlinear stochastic systems, integrates t

  3. Analyses and computations of asymmetric Z-scan for large phase shift from diffraction theory

    Institute of Scientific and Technical Information of China (English)

    Liyong Ren(任立勇); Baoli Yao(姚保利); Xun Hou(侯洵); Liren Liu(刘立人); Changhe Zhou(周常河)

    2003-01-01

    Based on Fresnel-Kirchhoff diffraction theory, we set up a diffraction model of nonlinear optical media toGaussian beam, which can interpret the Z-scan phenomenon from a new way. This theory is not onlywell consistent with the conventional Z-scan theory in the case of small nonlinear phase shift, but alsocan fit for the lager nonlinear phase shift. Numeric computations indicate the shape of the Z-scan curveis greatly affected by the value of the nonlinear phase shift. The symmetric dispersion-like Z-scan curveis only valid for small nonlinear phase shift (|△φo| <π), but with increasing the nonlinear phase shift, thevalley of the transmittance is severely suppressed and the peak is greatly enhanced. Further calculationsshow some new interesting results.

  4. Probing hysteretic elasticity in weakly nonlinear materials

    Energy Technology Data Exchange (ETDEWEB)

    Johnson, Paul A [Los Alamos National Laboratory; Haupert, Sylvain [UPMC UNIV PARIS; Renaud, Guillaume [UPMC UNIV PARIS; Riviere, Jacques [UPMC UNIV PARIS; Talmant, Maryline [UPMC UNIV PARIS; Laugier, Pascal [UPMC UNIV PARIS

    2010-12-07

    Our work is aimed at assessing the elastic and dissipative hysteretic nonlinear parameters' repeatability (precision) using several classes of materials with weak, intermediate and high nonlinear properties. In this contribution, we describe an optimized Nonlinear Resonant Ultrasound Spectroscopy (NRUS) measuring and data processing protocol applied to small samples. The protocol is used to eliminate the effects of environmental condition changes that take place during an experiment, and that may mask the intrinsic elastic nonlinearity. As an example, in our experiments, we identified external temperature fluctuation as a primary source of material resonance frequency and elastic modulus variation. A variation of 0.1 C produced a frequency variation of 0.01 %, which is similar to the expected nonlinear frequency shift for weakly nonlinear materials. In order to eliminate environmental effects, the variation in f{sub 0} (the elastically linear resonance frequency proportional to modulus) is fit with the appropriate function, and that function is used to correct the NRUS calculation of nonlinear parameters. With our correction procedure, we measured relative resonant frequency shifts of 10{sup -5} , which are below 10{sup -4}, often considered the limit to NRUS sensitivity under common experimental conditions. Our results show that the procedure is an alternative to the stringent control of temperature often applied. Applying the approach, we report nonlinear parameters for several materials, some with very small nonclassical nonlinearity. The approach has broad application to NRUS and other Nonlinear Elastic Wave Spectroscopy approaches.

  5. Terreneuvian orthothecid (Hyolitha digestive tracts from northern Montagne Noire, France; taphonomic, ontogenetic and phylogenetic implications.

    Directory of Open Access Journals (Sweden)

    Léa Devaere

    Full Text Available More than 285 specimens of Conotheca subcurvata with three-dimensionally preserved digestive tracts were recovered from the Terreneuvian (early Cambrian Heraultia Limestone of the northern Montagne Noire, southern France. They represent one of the oldest occurrences of such preserved guts. The newly discovered operculum of some complete specimens provides additional data allowing emendation of the species diagnosis. Infestation of the U-shaped digestive tracts by smooth uniseriate, branching to anastomosing filaments along with isolated botryoidal coccoids attests to their early, microbially mediated phosphatisation. Apart from taphonomic deformation, C. subcurvata exhibits three different configurations of the digestive tract: (1 anal tube and gut parallel, straight to slightly undulating; (2 anal tube straight and loosely folded gut; and (3 anal tube straight and gut straight with local zigzag folds. The arrangement of the digestive tracts and its correlation with the mean apertural diameter of the specimens are interpreted as ontogenetically dependent. The simple U-shaped gut, usually considered as characteristic of the Hyolithida, developed in earlier stages of C. subcurvata, whereas the more complex orthothecid type-3 only appears in largest specimens. This growth pattern suggests a distinct phylogenetic relationship between these two hyolith orders through heterochronic processes.

  6. Terreneuvian orthothecid (Hyolitha) digestive tracts from northern Montagne Noire, France; taphonomic, ontogenetic and phylogenetic implications.

    Science.gov (United States)

    Devaere, Léa; Clausen, Sébastien; Alvaro, J Javier; Peel, John S; Vachard, Daniel

    2014-01-01

    More than 285 specimens of Conotheca subcurvata with three-dimensionally preserved digestive tracts were recovered from the Terreneuvian (early Cambrian) Heraultia Limestone of the northern Montagne Noire, southern France. They represent one of the oldest occurrences of such preserved guts. The newly discovered operculum of some complete specimens provides additional data allowing emendation of the species diagnosis. Infestation of the U-shaped digestive tracts by smooth uniseriate, branching to anastomosing filaments along with isolated botryoidal coccoids attests to their early, microbially mediated phosphatisation. Apart from taphonomic deformation, C. subcurvata exhibits three different configurations of the digestive tract: (1) anal tube and gut parallel, straight to slightly undulating; (2) anal tube straight and loosely folded gut; and (3) anal tube straight and gut straight with local zigzag folds. The arrangement of the digestive tracts and its correlation with the mean apertural diameter of the specimens are interpreted as ontogenetically dependent. The simple U-shaped gut, usually considered as characteristic of the Hyolithida, developed in earlier stages of C. subcurvata, whereas the more complex orthothecid type-3 only appears in largest specimens. This growth pattern suggests a distinct phylogenetic relationship between these two hyolith orders through heterochronic processes.

  7. Large ontogenetic declines in intra-crown leaf area index in two temperate deciduous tree species.

    Science.gov (United States)

    Nock, C A; Caspersen, J P; Thomas, S C

    2008-03-01

    The widespread occurrence of age-related changes in leaf morphology and allocation suggests that the leaf area index of individual trees (intra-crown LAI) may decline late in ontogeny. We used direct, within-canopy measurements to quantify the LAI of canopy trees with exposed crowns of two temperate deciduous species. Intra-crown LAI declined from approximately 7 to 4 in Acer saccharum, and from approximately 9.5 to 6.5 in Betula alleghaniensis, as tree size increased (from 15 to 72 cm diameter at breast height [dbh]). For A. saccharum, age (which varied from 30 to 160 years) was a significantly better predictor of LAI decline than dbh. We also modeled the effect of ontogenetic declines in LAI on understory light availability and found that light transmission increases significantly as canopy trees grow and mature. Our results thus suggest that gradual declines in LAI with tree age may play an important and overlooked role in contributing to the heterogeneity of sub-canopy light regimes in mature forests.

  8. Ontogenetic differences in the feeding biomechanics of oviparous and viviparous caecilians (Lissamphibia: Gymnophiona).

    Science.gov (United States)

    Kleinteich, Thomas

    2010-10-01

    Caecilians have a unique dual jaw-closing system in that jaw closure is driven by the ancestral jaw-closing muscles (mm. levatores mandibulae) plus a secondarily recruited hyobranchial muscle (m. interhyoideus posterior). There is a variety of feeding habits (suction feeding, skin feeding, intrauterine scraping, and biting) during ontogeny that relate to reproductive modes in different caecilian species. This study examines the cranial biomechanics of caecilians in the suction-feeding larva of Ichthyophis cf. kohtaoensis, in the embryo and juvenile of the skin-feeding Boulengerula taitana, and in a newborn of the intrauterine feeder Typhlonectes natans. A lever arm model was applied to calculate effective mechanical advantages of jaw-closing muscles over gape angles and to predict total bite force in developing caecilians. In I. cf. kohtaoensis, Notable differences were found in the larval jaw-closing system compared to that of the adult. The suction-feeding larva of I. cf. kohtaoensis has comparatively large mm. levatores mandibulae that insert with an acute muscle fiber angle to the lower jaw and a m. interhyoideus posterior that has its optimal leverage at small gape angles. Conversely, the skin-feeding juvenile of B. taitana and the neonate T. natans are very similar in the feeding parameters considered herein compared to adult caecilians. Some ontogenetic variation in the feeding system of B. taitana before the onset of feeding was present. This study contributes to our understanding of the functional demands that feeding habits put on the development of cranial structures.

  9. Manganese-induced oxidative stress in two ontogenetic stages of chamomile and amelioration by nitric oxide.

    Science.gov (United States)

    Kováčik, Jozef; Babula, Petr; Hedbavny, Josef; Švec, Pavel

    2014-02-01

    Impact of manganese (Mn(2+)) excess (100, 500 and 1000 μM over 7 days) on two ontogenetic stages (7-week-old plants and 7-day-old seedlings) of Matricaria chamomilla was compared. Mn excess depressed growth of seedlings (but not germination) and stimulated oxidative stress (ROS and lipid peroxidation) in both plants and seedlings. Growth inhibition could be evoked by higher Mn uptake and higher translocation factor in seedlings than in plants. Total thiols staining revealed elevation in almost all treatments. In 7-week-old plants, activity of peroxidases increased slightly and rather decreased under high Mn doses. Superoxide rather than hydrogen peroxide contributed to visualized ROS presence. Fluorescence of nitric oxide (NO) showed stimulation in plants but decrease in seedlings. Impact of exogenous nitric oxide donor (sodium nitroprusside/SNP) was therefore tested and results showed amelioration of 1000 μM Mn-induced oxidative stress in seedlings (decrease in H2O2 and increase in NO content while antioxidative enzyme activities were variably affected) concomitantly with depleted Mn accumulation. It is concluded that NO participates in tolerance to Mn excess but negative effects of the highest SNP dose were also observed. Extensive fluorescence microscopy is also explanatively discussed.

  10. Vertical movement patterns and ontogenetic niche expansion in the tiger shark, Galeocerdo cuvier.

    Science.gov (United States)

    Afonso, André S; Hazin, Fábio H V

    2015-01-01

    Sharks are top predators in many marine ecosystems and can impact community dynamics, yet many shark populations are undergoing severe declines primarily due to overfishing. Obtaining species-specific knowledge on shark spatial ecology is important to implement adequate management strategies for the effective conservation of these taxa. This is particularly relevant concerning highly-mobile species that use wide home ranges comprising coastal and oceanic habitats, such as tiger sharks, Galeocerdo cuvier. We deployed satellite tags in 20 juvenile tiger sharks off northeastern Brazil to assess the effect of intrinsic and extrinsic factors on depth and temperature usage. Sharks were tracked for a total of 1184 d and used waters up to 1112 m in depth. The minimum temperature recorded equaled 4°C. All sharks had a clear preference for surface (sharks used mostly shallow (sharks spending considerably more time in surface (shark habitat was observed, with generalized linear models estimating a ~4-fold increase in maximum diving depth from 150- to 300-cm size-classes. The time spent in the upper 5 m of the water column did not vary ontogenetically but shark size was the most important factor explaining the utilization of deeper water layers. Young-of-the-year tiger sharks seem to associate with shallow, neritic habitats but they progressively move into deeper oceanic habitats as they grow larger. Such an early plasticity in habitat use could endow tiger sharks with access to previously unavailable prey, thus contributing to a wider ecological niche.

  11. Ontogenetic cell death and phagocytosis in the visual system of vertebrates.

    Science.gov (United States)

    Francisco-Morcillo, Javier; Bejarano-Escobar, Ruth; Rodríguez-León, Joaquín; Navascués, Julio; Martín-Partido, Gervasio

    2014-10-01

    Programmed cell death (PCD), together with cell proliferation, cell migration, and cell differentiation, is an essential process during development of the vertebrate nervous system. The visual system has been an excellent model on which to investigate the mechanisms involved in ontogenetic cell death. Several phases of PCD have been reported to occur during visual system ontogeny. During these phases, comparative analyses demonstrate that dying cells show similar but not identical spatiotemporally restricted patterns in different vertebrates. Additionally, the chronotopographical coincidence of PCD with the entry of specialized phagocytes in some regions of the developing vertebrate visual system suggests that factors released from degenerating cells are involved in the cell migration of macrophages and microglial cells. Contradicting this hypothesis however, in many cases the cell corpses generated during degeneration are rapidly phagocytosed by neighboring cells, such as neuroepithelial cells or Müller cells. In this review, we describe the occurrence and the sites of PCD during the morphogenesis and differentiation of the retina and optic pathways of different vertebrates, and discuss the possible relationship between PCD and phagocytes during ontogeny.

  12. Ontogenetic changes in the chemistry and morphology of oil glands in Hermannia convexa (Acari: Oribatida).

    Science.gov (United States)

    Raspotnig, Günther; Krisper, Günther; Schuster, Reinhart

    2005-01-01

    As a first example for the chemistry of oil gland secretions in the Hermannioidea (one of the three superfamilies of desmonomatan Oribatida), the oil gland secretion of Hermannia convexa was investigated by gas chromatography-mass spectrometry. Hexane extracts of all juvenile stages showed a multicomponent chromatographic pattern, mainly consisting of well-known oil gland secretion components such as neral, geranial, gamma-acaridial and the unsaturated C17-hydrocarbons, 6,9-heptadecadiene and 8-heptadecene. The secretion profiles of juveniles varied slightly between samples of two different collections, namely in the presence of gamma-acaridial and 8-heptadecene. Furthermore, a minor component, identified as 1,8-cineole (= eucalyptol) and hitherto not known from oil gland secretions of other species, was recorded in both juvenile and adult extracts. In adult profiles, 1,8-cineole, in low amounts, represented the only detectable component; thus, their profiles fundamentally differed from those of juveniles. A subsequent histological investigation revealed well developed oil glands in all juvenile stages, but degenerated oil glands in adults, consistent with the chemical data. So far, apart from H. convexa, degeneration of oil glands in the course of ontogenetic development is only known from a brachypylid species; on the other hand, chemical oil gland-polymorphism between juveniles and adults may occur in closely related Nothridae while it does not occur in oil glands of early- and middle-derivative Oribatida (Parhyposomata, Mixonomata, trhypochthoniid Desmonomata), nor in astigmatid mites.

  13. Variations of Salminus hilarii diet (Ostariophysi, Characidae): seasonal and ontogenetic effects.

    Science.gov (United States)

    Villares Junior, G A; Goitein, R

    2015-08-01

    This study described the variations seasonal and ontogenetic of Salminus hilarii diet. Samples were collected in the Sorocaba River, São Paulo, Brazil, one of the few rivers where individuals of the species still occur in a higher frequency. The preys consumed were analyzed by Importance Alimentary Index (AIi). To determine similarities between year seasons, the AIi data were analyzed by the Morisita-Horn index and reduced in cluster analysis, along with a statistical comparison made by one-way ANOSIM test (5%). The feeding activity was analyzed according to the stomach repletion index and compared among the year seasons using non parametric variance analysis Kruskal-Wallis test (5%). Comparison of prey consumed between immature and adult individuals was made by Spearman correlation (5%). A Pearson correlation (5%) was applied between the standard length of the fish and prey consumed, as well as between the mouth and prey heights. The analyzes of stomach contents showed that the diet of this species was exclusively piscivorous, with significant difference of prey consumption during the period, the same happening among adult and immature individuals. It was observed that these fishes use to swallow their prey whole and that significant correlations between size of predator and prey size can be observed. There is also correlation between the mouth height and the maximum prey depth. Salminus hilarii feeds on the available prey, and the species food composition and feeding activity depends on prey`s abundance, their size and morphology, as do the water temperatures.

  14. Chlorophyll b in angiosperms: Functions in photosynthesis, signaling and ontogenetic regulation.

    Science.gov (United States)

    Voitsekhovskaja, O V; Tyutereva, E V

    2015-09-15

    Chlorophyll b (Chlb) is an antenna chlorophyll. The binding of Chlb by antenna proteins is crucial for the correct assembly of the antenna complexes in thylakoid membranes. Since the levels of the proteins of major and minor antenna are affected to different extents by Chlb binding, the availability of Chlb influences the composition and the size of antenna complexes which in turn determine the supramolecular organization of the thylakoid membranes in grana. Therefore, Chlb synthesis levels have a major impact on lateral mobility and diffusion of membrane molecules, and thus affect not only light harvesting and thermal energy dissipation processes, but also linear electron transport and repair processes in grana. Furthermore, in angiosperms Chlb synthesis affects plant functions beyond chloroplasts. First, the stability of pigment-protein complexes in the antennae, which depends on Chlb, is an important factor in the regulation of plant ontogenesis, and Chlb levels were recently shown to influence plant ontogenetic signaling. Second, the amounts of minor antenna proteins in chloroplasts, which depend on the availability of Chlb, were recently shown to affect ABA levels and signaling in plants. These mechanisms can be examined in mutants where Chlb synthesis is reduced or abolished. The dramatic effects caused by the lack of Chlb on plant productivity are interpreted in this review in light of the pleiotropic effects on photosynthesis and signaling, and the potential to manipulate Chlb biosynthesis for the improvement of crop production is discussed. Copyright © 2015 Elsevier GmbH. All rights reserved.

  15. Ontogenetic scaling of the humerus in sea turtles and its implications for locomotion.

    Science.gov (United States)

    Nishizawa, Hideaki; Asahara, Masakazu; Kamezaki, Naoki

    2013-03-01

    In the present study, we analyzed the ontogenetic scaling of humeri in the green turtle (Chelonia mydas) and loggerhead turtle (Caretta caretta). Green turtles have relatively thicker humeri than loggerhead turtles, indicating that the humerus of the green turtle can resist greater loads. Our results are consistent with isometry, or slightly negative allometry, of diameter in relation to length of the humerus in both species. Geometric similarity or isometry of the humerus in relation to body mass is supported by estimates of the cross-sectional properties of green turtles. Sea turtles are adapted for aquatic life, but also perform terrestrial locomotion. Thus, during terrestrial locomotion, which requires support against gravity, the observed scaling relationships indicate that there may be greater stress and fracture risk on the humeri of larger green turtles than on the humeri of smaller turtles. In aquatic habitats, in which limbs are mainly used for propulsion, the stress and fracture risk for green turtle humeri are estimated to increase with greater speed. This scaling pattern may be related to the possibility that smaller turtles swim at a relatively faster speed per body length.

  16. Ontogenetic variation of plastral spotting pattern in Phrynops hilarii (Testudines, Chelidae

    Directory of Open Access Journals (Sweden)

    Cabrera Mario R.

    2001-01-01

    Full Text Available The plastral spotting variation in the chelid turtle Phrynops hilarii (Duméril & Bibron, 1835 in relation to sex, size, and geographic procedence of individuals was analyzed. States for qualitative characters were analyzed using non-parametric tests. Quantitative characters (shell and scute measurements were standardized for body size by linear regression against carapace length, and were subjected to principal components analysis and canonical discriminant function analysis. Results suggest that increased plastral spotting is a polymorphic ontogenetic trait in P. hilarii. Neither hatchlings nor juveniles have plastral pattern moderately or heavily pigmented. The simplest pattern, however, may persist without changes in some adults. There are no differences between sexes. The spatial distribution of the plastral pattern is not ordered latitudinally or longitudinally, showing no relationship with gradients of elevation, temperature, or precipitation. This pattern trait lacks of taxonomic significance. The morphometric analysis failed to reveal any character of diagnostic utility in the plastron to support the possibility that these patterns correspond to different sympatric taxa.

  17. Changes of motor abilities during ontogenetic development in Lurcher mutant mice.

    Science.gov (United States)

    Markvartová, V; Cendelín, J; Vozeh, F

    2010-07-14

    Lurcher mutant mice represent a natural model of olivocerebellar degeneration. This degeneration is caused by a mutation of the gene for the delta2 glutamate receptor. Lurcher mutants suffer from cerebellar ataxia and cognitive functions deficiency as a consequence of excitotoxic apoptosis of Purkinje cells in the cerebellar cortex and a secondary decrease of granule cells and inferior olive neurons. This process finishes by the 90th day of postnatal life, but already by 14 days, the Purkinje cells are damaged and the ataxia is fully developed. Purkinje cells die by apoptosis within the first 3 weeks of life. The aim of our work was to study the development of motor functions in the course of the ontogenetic development in Lurcher mutant mice of the B6CBA strain and to compare it with wild type mice of the same strain. Mice aged 2, 3, 6, 9, and 22 weeks were used in our experiment. Motor skills were examined using four standard tests: the horizontal wire, rotating cylinder, footbridge and slanting ladder. Our findings in Lurcher mutant mice show a significant increase of motor abilities up to the sixth postnatal week and selective decrease early after this period. This improvement of motor skills is caused by the physiological development of musculature and the nervous system, probably with some contribution of plasticity of the maturing brain. The cause of the decline of these abilities immediately after the completion of the development is unknown.

  18. Ontogenetic variations and structural adjustments in mammals evolving prolonged to continuous dental growth

    Science.gov (United States)

    Gomes Rodrigues, Helder; Lefebvre, Rémi; Fernández-Monescillo, Marcos; Mamani Quispe, Bernardino; Billet, Guillaume

    2017-01-01

    Studying dental ontogeny in mammals can provide valuable insight on the evolution of their masticatory apparatus and their related adaptations. The multiple acquisitions of a prolonged to continuous growth of teeth in herbivorous mammals in response to high abrasion represent an intensively investigated issue. However, the ontogenetic and architectural patterns associated with these repeated dental innovations remain poorly known. Here, we focused on two case studies corresponding to distant mammalian clades, the extinct Mesotheriidae (Notoungulata), which shared some striking dental features with the extant Ctenodactylidae (Rodentia). We studied the impact of prolonged to continuous growth of molars on their occlusal complexity, their relative size and their dynamics in the jaw. We found that variations of occlusal complexity patterns are the result of paedomorphic or peramorphic heterochronic processes impacting dental crown. We showed that variations in both upper and lower molar proportions generally follow the inhibitory developmental cascade model. In that context, prolonged dental growth implies transitory adjustments due to wear, and also involves dental migration and loss when combined with molar lengthening. Interestingly, these features may be present in many mammals having prolonged dental growth, and emphasize the crucial need of considering these aspects in future evolutionary and developmental studies. PMID:28791172

  19. DESCRIPTION OF AN EARLY ONTOGENETIC EVOLUTIONARY STEP IN LEPIDORBITOIDES (LEPIDORBITOIDES BISAMBERGENSIS ASYMMETRICA, EARLY MAASTRICHTIAN (CENTRAL TURKEY

    Directory of Open Access Journals (Sweden)

    ERCAN ÖZCAN

    2001-03-01

    Full Text Available Lepidorbitoides bisambergensis is characterised by having a ‘quadriserial’ embryo without any chamberlet directly arising from the deuteroconch and is a very diagnostic and common species in Lower Maastrichtian flysch successions in Anatolia. Some populations of this species present an early ontogenetic morphologic feature which is characterised by distinctly asymmetric early chamber arrangement recognised in the horizontal sections. This asymmetry is mainly caused by the pronounced difference in the size of auxiliary chamberlets which rest on both, protoconch and deuteroconch and also enhanced by the development of unequal number of chamberlets in the series arising from these auxiliary chamberlets on the protoconchal side. These asymmetric specimens are commonly identified in stratigraphic horizons below the symmetric ones after the introduction of a new auxiliary chamberlet and progressively replaced by symmetric ones in the younger populations. Asymmetric "quadriserial" specimens representing the early phylogenetic stage of L. bisambergensis described in the stratigraphic horizons corresponding to G. havanensis and G. aegyptiaca (? zones are thought to deserve a particular taxonomic status and are attributed to Lepidorbitoides bisambergensis asymmetrica Özcan & Özkan-Altiner, 1999a. 

  20. Ontogenetic variations and structural adjustments in mammals evolving prolonged to continuous dental growth

    Science.gov (United States)

    Gomes Rodrigues, Helder; Lefebvre, Rémi; Fernández-Monescillo, Marcos; Mamani Quispe, Bernardino; Billet, Guillaume

    2017-07-01

    Studying dental ontogeny in mammals can provide valuable insight on the evolution of their masticatory apparatus and their related adaptations. The multiple acquisitions of a prolonged to continuous growth of teeth in herbivorous mammals in response to high abrasion represent an intensively investigated issue. However, the ontogenetic and architectural patterns associated with these repeated dental innovations remain poorly known. Here, we focused on two case studies corresponding to distant mammalian clades, the extinct Mesotheriidae (Notoungulata), which shared some striking dental features with the extant Ctenodactylidae (Rodentia). We studied the impact of prolonged to continuous growth of molars on their occlusal complexity, their relative size and their dynamics in the jaw. We found that variations of occlusal complexity patterns are the result of paedomorphic or peramorphic heterochronic processes impacting dental crown. We showed that variations in both upper and lower molar proportions generally follow the inhibitory developmental cascade model. In that context, prolonged dental growth implies transitory adjustments due to wear, and also involves dental migration and loss when combined with molar lengthening. Interestingly, these features may be present in many mammals having prolonged dental growth, and emphasize the crucial need of considering these aspects in future evolutionary and developmental studies.

  1. Cranial ontogenetic variation in early saurischians and the role of heterochrony in the diversification of predatory dinosaurs

    Directory of Open Access Journals (Sweden)

    Christian Foth

    2016-01-01

    Full Text Available Non-avian saurischian skulls underwent at least 165 million years of evolution and shapes varied from elongated skulls, such as in the theropod Coelophysis, to short and box-shaped skulls, such as in the sauropod Camarasaurus. A number of factors have long been considered to drive skull shape, including phylogeny, dietary preferences and functional constraints. However, heterochrony is increasingly being recognized as an important factor in dinosaur evolution. In order to quantitatively analyse the impact of heterochrony on saurischian skull shape, we analysed five ontogenetic trajectories using two-dimensional geometric morphometrics in a phylogenetic framework. This allowed for the comparative investigation of main ontogenetic shape changes and the evaluation of how heterochrony affected skull shape through both ontogenetic and phylogenetic trajectories. Using principal component analyses and multivariate regressions, it was possible to quantify different ontogenetic trajectories and evaluate them for evidence of heterochronic events allowing testing of previous hypotheses on cranial heterochrony in saurischians. We found that the skull shape of the hypothetical ancestor of Saurischia likely led to basal Sauropodomorpha through paedomorphosis, and to basal Theropoda mainly through peramorphosis. Paedomorphosis then led from Orionides to Avetheropoda, indicating that the paedomorphic trend found by previous authors in advanced coelurosaurs may extend back into the early evolution of Avetheropoda. Not only are changes in saurischian skull shape complex due to the large number of factors that affected it, but heterochrony itself is complex, with a number of possible reversals throughout non-avian saurischian evolution. In general, the sampling of complete ontogenetic trajectories including early juveniles is considerably lower than the sampling of single adult or subadult individuals, which is a major impediment to the study of heterochrony on

  2. Hopf Bifurcation in a Nonlinear Wave System

    Institute of Scientific and Technical Information of China (English)

    HE Kai-Fen

    2004-01-01

    @@ Bifurcation behaviour of a nonlinear wave system is studied by utilizing the data in solving the nonlinear wave equation. By shifting to the steady wave frame and taking into account the Doppler effect, the nonlinear wave can be transformed into a set of coupled oscillators with its (stable or unstable) steady wave as the fixed point.It is found that in the chosen parameter regime, both mode amplitudes and phases of the wave can bifurcate to limit cycles attributed to the Hopf instability. It is emphasized that the investigation is carried out in a pure nonlinear wave framework, and the method can be used for the further exploring routes to turbulence.

  3. Nonlinear channelizer

    Science.gov (United States)

    In, Visarath; Longhini, Patrick; Kho, Andy; Neff, Joseph D.; Leung, Daniel; Liu, Norman; Meadows, Brian K.; Gordon, Frank; Bulsara, Adi R.; Palacios, Antonio

    2012-12-01

    The nonlinear channelizer is an integrated circuit made up of large parallel arrays of analog nonlinear oscillators, which, collectively, serve as a broad-spectrum analyzer with the ability to receive complex signals containing multiple frequencies and instantaneously lock-on or respond to a received signal in a few oscillation cycles. The concept is based on the generation of internal oscillations in coupled nonlinear systems that do not normally oscillate in the absence of coupling. In particular, the system consists of unidirectionally coupled bistable nonlinear elements, where the frequency and other dynamical characteristics of the emergent oscillations depend on the system's internal parameters and the received signal. These properties and characteristics are being employed to develop a system capable of locking onto any arbitrary input radio frequency signal. The system is efficient by eliminating the need for high-speed, high-accuracy analog-to-digital converters, and compact by making use of nonlinear coupled systems to act as a channelizer (frequency binning and channeling), a low noise amplifier, and a frequency down-converter in a single step which, in turn, will reduce the size, weight, power, and cost of the entire communication system. This paper covers the theory, numerical simulations, and some engineering details that validate the concept at the frequency band of 1-4 GHz.

  4. Are tree ontogenetic structure and allometric relationship independent of vegetation formation type? A case study with Cordia oncocalyx in the Brazilian caatinga

    Science.gov (United States)

    Silveira, Andréa P.; Martins, Fernando R.; Araújo, Francisca S.

    2012-08-01

    In temperate and tropical rainforests, ontogenetic structure and allometry during tree ontogeny are often associated with light gradients. Light is not considered a limiting resource in deciduous thorny woodland (DTW), but establishment and growth occur during a short rainy period, when the canopy is fully leaved and light in the understory may be modified. Our aim was to investigate whether the light gradient in DTW and the biomechanical limitations of tree growth would be enough to produce an ontogenetic structure and allometric growth similar to rainforest canopy trees. We investigated the ontogenetic stages and diameter-height relationship of Cordia oncocalyx (Boraginaceae), a dominant canopy tree of the DTW of semiarid northeastern Brazil. We tagged, measured and classified the ontogenetic stages of 2.895 individuals in a 1 ha area (5°6'58.1″S and 40°52'19.4″W). In the rainy season only 4.7% of the light falling on the canopy reached the ground. Initial ontogenetic stages, mainly infant (50.9%) and seedling (42.1%), were predominant in the population, with the remaining 7% distributed among juvenile, immature, virginile and reproductive. The ontogenetic structure was similar to that of rainforest tree species, but the population formed both permanent seed and infant banks in response to long dry periods and erratic rainy spells. Like many other Boraginaceae tree species in tropical rainforests, C. oncocalyx has a Prévost architectural model, but allometric growth was quite different from rainforest trees. C. oncocalyx invested slightly more in diameter at first, then in height and finally invested greatly in diameter and attained an asymptotic height. The continued high investment in diameter growth at late stages and the asymptotic height point to low tree density and more frequent xylem embolism as the main drivers of tree allometric shape in DTW. This indicates that tree ontogenetic structure and allometric relationships depend on vegetation

  5. Investigation of High-Nonlinearity Glass Fibers for Potential Applications in Ultrafast Nonlinear Fiber Devices

    OpenAIRE

    Kim, Jong-Kook

    2005-01-01

    Nonlinear fiber devices have been attracting considerable attention in recent years, due to their inherent ultrafast response time and potential applications in optical communication systems. They usually require long fibers to generate sufficient nonlinear phase shifts, since nonlinearities of conventional silica-core silica-clad fibers are too low. These long devices, however, cause the serious problems of pulse walk-off, pulse broadening, and polarization fluctuation which are major limiti...

  6. Nonlinear dynamics

    Energy Technology Data Exchange (ETDEWEB)

    Turchetti, G. (Bologna Univ. (Italy). Dipt. di Fisica)

    1989-01-01

    Research in nonlinear dynamics is rapidly expanding and its range of applications is extending beyond the traditional areas of science where it was first developed. Indeed while linear analysis and modelling, which has been very successful in mathematical physics and engineering, has become a mature science, many elementary phenomena of intrinsic nonlinear nature were recently experimentally detected and investigated, suggesting new theoretical work. Complex systems, as turbulent fluids, were known to be governed by intrinsically nonlinear laws since a long time ago, but received purely phenomenological descriptions. The pioneering works of Boltzmann and Poincare, probably because of their intrinsic difficulty, did not have a revolutionary impact at their time; it is only very recently that their message is reaching a significant number of mathematicians and physicists. Certainly the development of computers and computer graphics played an important role in developing geometric intuition of complex phenomena through simple numerical experiments, while a new mathematical framework to understand them was being developed.

  7. Compressive Shift Retrieval

    Science.gov (United States)

    Ohlsson, Henrik; Eldar, Yonina C.; Yang, Allen Y.; Sastry, S. Shankar

    2014-08-01

    The classical shift retrieval problem considers two signals in vector form that are related by a shift. The problem is of great importance in many applications and is typically solved by maximizing the cross-correlation between the two signals. Inspired by compressive sensing, in this paper, we seek to estimate the shift directly from compressed signals. We show that under certain conditions, the shift can be recovered using fewer samples and less computation compared to the classical setup. Of particular interest is shift estimation from Fourier coefficients. We show that under rather mild conditions only one Fourier coefficient suffices to recover the true shift.

  8. Ontogenetic differences in mesophyll structure and chlorophyll distribution in Eucalyptus globulus ssp. globulus.

    Science.gov (United States)

    James, S A; Smith, W K; Vogelmann, T C

    1999-02-01

    Mesophyll structure has been associated with the photosynthetic performance of leaves via the regulation of internal light and CO(2) profiles. Differences in mesophyll structure and chlorophyll distribution within three ontogenetically different leaf types of Eucalyptus globulus ssp. globulus were investigated. Juvenile leaves are blue-grey in color, dorsiventral (adaxial palisade layer only), hypostomatous, and approximately horizontal in orientation. In contrast, adult leaves are dark green in color, isobilateral (adaxial and abaxial palisade), amphistomatous, and nearly vertical in orientation. The transitional leaf type has structural features that appear intermediate between the juvenile and adult leaves. The ratio of mesophyll cell surface area per unit leaf surface area (A(mes)/A) of juvenile leaves was maximum at the base of a single, adaxial palisade layer and declined through the spongy mesophyll. Chlorophyll a + b content showed a coincident pattern, while the chlorophyll a:b ratio declined linearly from the adaxial to abaxial epidermis. In comparison, the mesophyll of adult leaves had a bimodal distribution of A(mes)/A, with maxima occurring beneath both the adaxial and abaxial surfaces within the first layer of multiple palisade layers. The distribution of chlorophyll a + b content had a similar pattern, although the maximum ratio of chlorophyll a:b occurred immediately beneath the adaxial and abaxial epidermis. The matching distributions of A(mes)/A and chlorophyll provide further evidence that mesophyll structure may act to influence photosynthetic performance. These changes in internal leaf structure at different life stages of E. globulus may be an adaptation for increased xeromorphy under increasing light exposure experienced from the seedling to adult tree, similar to the characteristics reported for different species according to sunlight exposure and water availability within their native habitats.

  9. Astaxanthin, canthaxanthin and astaxanthin esters in the copepod Acartia bifilosa (Copepoda, Calanoida during ontogenetic development

    Directory of Open Access Journals (Sweden)

    Maria £otocka

    2001-12-01

    Full Text Available The contents of astaxanthin, canthaxanthin and astaxanthin esters were studied in natural populations of the copepod Acartia bifilosa from the Pomeranian Bay and Gulf of Gdansk in the southern Baltic Sea. Samples dominated by any one of three developmental groups: (1 nauplii, (2 copepodids I-III and (3 copepodids IV-V and adults of Acartia bifilosa were analysed by means of high performance liquid chromatography (HPLC. As ontogenetic development progressed, significant changes occurred in the proportion of particular pigments in the total pigment pool of the various developmental groups. Astaxanthin and canthaxanthin occurred in all the groups, the former being clearly dominant. However, an increasing percentage of astaxanthin esters was recorded in the copepodids I-III, and even more in the copepodids IV-V and adults group. Most probably, astaxanthin is the main pigment active in copepod lipid metabolism. Carotenoid pigments in copepods very likely act as efficient free-electron quenchers and may be involved as antioxidants in rapid lipid metabolism. The exogenously feeding stages (late nauplii and copepodids transform plant carotenoids taken from food and are evidently capable of metabolising astaxanthin by esterification and further degradation. It is emphasised that, according to literature data, astaxanthin esters may have an even higher quenching ability. It is suggested that crustacean carotenoid pigments, with their electron donor-acceptor abilities, may replace oxygen in peroxidation processes connected with lipid metabolism. The consequences of such a physiological role of astaxanthin for present-day estimations of energy balances in zooplankton communities are mentioned.

  10. Ontogenetic changes in digestive enzyme activities and the amino acid profile of starry flounder Platichthys stellatus

    Science.gov (United States)

    Song, Zhidong; Wang, Jiying; Qiao, Hongjin; Li, Peiyu; Zhang, Limin; Xia, Bin

    2016-09-01

    Ontogenetic changes in digestive enzyme activities and the amino acid (AA) profile of starry flounder, Platichthys stellatus, were investigated and limiting amino acids were estimated compared with the essential AA profile between larvae and live food to clarify starry flounder larval nutritional requirements. Larvae were collected at the egg stage and 0, 2, 4, 7, 12, 17, 24 days after hatching (DAH) for analysis. Larvae grew from 1.91 mm at hatching to 12.13 mm at 24 DAH. Trypsin and chymotrypsin activities changed slightly by 4 DAH and then increased significantly 4 DAH. Pepsin activity increased sharply beginning 17 DAH. Lipase activity increased significantly 4 DAH and increased progressively with larval growth. Amylase activity was also detected in newly hatched larvae and increased 7 DAH followed by a gradual decrease. High free amino acid (FAA) content was detected in starry flounder eggs (110.72 mg/g dry weight). Total FAA content dropped to 43.29 mg/g in 4-DAH larvae and then decreased gradually to 13.74 mg/g in 24-DAH larvae. Most FAAs (except lysine and methionine) decreased >50% in 4-DAH larvae compared with those in eggs and then decreased to the lowest values in 24-DAH larvae. Changes in the protein amino acid (PAA) profile were much milder than those observed for FAAs. Most PAAs increased gradually during larval development, except lysine and phenylalanine. The percentages of free threonine, valine, isoleucine, and leucine decreased until the end of the trial, whereas the protein forms of these four AAs followed the opposite trend. A comparison of the essential AA composition of live food (rotifers, Artemia nauplii, and Artemia metanauplii) and larvae suggested that methionine was potentially the first limiting AA. These results may help develop starry flounder larviculture methods by solving the AA imbalance in live food. Moreover, the increased digestive enzyme activities indicate the possibility of introducing artificial compound feed.

  11. Spatial and ontogenetic variability in the chemical composition of juvenile common sole ( Solea solea) otoliths

    Science.gov (United States)

    Tanner, S. E.; Vasconcelos, R. P.; Reis-Santos, P.; Cabral, H. N.; Thorrold, S. R.

    2011-01-01

    A description of variations in the chemical composition of fish otoliths at different spatial scales and life history stages is a prerequisite for their use as natural tags in fish population connectivity and migration studies. Otolith geochemistry of juvenile common sole ( Solea solea), a marine migrant species collected in six Portuguese estuaries was examined. Elemental ratios (Mg:Ca, Mn:Ca, Cu:Ca, Sr:Ca, Ba:Ca, Pb:Ca) were analysed in two zones of the right otolith (corresponding to late larval and juvenile stages) using laser ablation inductively coupled plasma mass spectrometry (ICP-MS). Stable carbon and oxygen isotopes (δ 13C and δ 18O) were determined in left otoliths using isotopic ratio monitoring mass spectrometry (irm-MS). Significant differences in otolith geochemical signatures were found among estuaries, among sites within estuaries and between otolith zones. Several elemental ratios (Mg:Ca, Mn:Ca, Cu:Ca and Sr:Ca) showed consistent patterns between otolith zones and were likely influenced by environmental factors and ontogenetic effects associated with physiological changes during metamorphosis. Assignment of individuals to their collection estuary based on the otolith geochemical signatures was more accurate at the site level (81%) than among estuaries (69%). Site temperature was not correlated with any of the elemental or isotope ratios, but salinity was significantly correlated with Ba:Ca, δ 13C and δ 18O. Observed spatial variations among estuaries and sites within estuaries indicate that geochemical signatures in otoliths are accurate natural tags of estuarine habitat in common sole. Nevertheless, the significant variations observed between otolith zones should be taken into account in the design of population connectivity studies.

  12. Ontogenetic changes in the shoot primary vasculature of Anagallis arvensis L.

    Directory of Open Access Journals (Sweden)

    Dorota Kwiatkowska

    2014-01-01

    Full Text Available Ontogenetic changes in the primary vasculature of Anagallis shoots are strictly related to phyllotaxis. During the ontogeny of Anagallis, whorled and spiral phyllotactic patterns appear alternately in a regular sequence. The initial decussate phyllotaxis is transformed into the spiral Fibonacci, and then further into trimerous pattern. This in turn may change into the spiral Lucas phyllotaxis. Sporadically the immediate transition from the decussate to trimerous phyllotaxis takes place. The vascular system in Anagallis is always closed, despite that both whorled and spiral phyllotaxes are present. Also the number of vascular traces diverging to the leaf is constant. In the course of a single phyllotactic transition, there is an increase in the number of vascular sympodia and in the number of leaf traces present in the vascular cylinder. Usually only one single sympodium and one or two traces are added to the system. The immediate addition of two sympodia occurs only during infrequent transition from the decussate to trimerous pattern. The increase in the number of sympodia is most often simultaneous with the phyllotactic transition, however, when the trimerous pattern is transformed into the spiral Lucas, the increase is delayed, sometimes for as much as ten plastochrons. In shoots with changing phyllotaxis, a sector within the vascular cylinder can be distinguished, in which the leaf traces are arranged as if the previous phyllotactic pattern continued, whereas rearrangement of traces takes place at the same level but in the complementary sector. This is in agreement with the concept of discontinuous circumferential changes in the shoot apex being responsible for qualitative transformations of phyllotaxis.

  13. Nonlinear Systems.

    Science.gov (United States)

    Seider, Warren D.; Ungar, Lyle H.

    1987-01-01

    Describes a course in nonlinear mathematics courses offered at the University of Pennsylvania which provides an opportunity for students to examine the complex solution spaces that chemical engineers encounter. Topics include modeling many chemical processes, especially those involving reaction and diffusion, auto catalytic reactions, phase…

  14. Evaluating hair as a predictor of blood mercury: the influence of ontogenetic phase and life history in pinnipeds

    Science.gov (United States)

    Peterson, Sarah H.; McHuron, Elizabeth A.; Kennedy, Stephanie N.; Ackerman, Josh; Rea, Lorrie D.; Castellini, J. Margaret; O'Hara, Todd M.; Costa, Daniel P.

    2016-01-01

    Mercury (Hg) biomonitoring of pinnipeds increasingly utilizes nonlethally collected tissues such as hair and blood. The relationship between total Hg concentrations ([THg]) in these tissues is not well understood for marine mammals, but it can be important for interpretation of tissue concentrations with respect to ecotoxicology and biomonitoring. We examined [THg] in blood and hair in multiple age classes of four pinniped species. For each species, we used paired blood and hair samples to quantify the ability of [THg] in hair to predict [THg] in blood at the time of sampling and examined the influence of varying ontogenetic phases and life history of the sampled animals. Overall, we found that the relationship between [THg] in hair and blood was affected by factors including age class, weaning status, growth, and the time difference between hair growth and sample collection. Hair [THg] was moderately to strongly predictive of current blood [THg] for adult female Steller sea lions (Eumetopias jubatus), adult female California sea lions (Zalophus californianus), and adult harbor seals (Phoca vitulina), whereas hair [THg] was poorly predictive or not predictive (different times of year) of blood [THg] for adult northern elephant seals (Mirounga angustirostris). Within species, except for very young pups, hair [THg] was a weaker predictor of blood [THg] for prereproductive animals than for adults likely due to growth, variability in foraging behavior, and transitions between ontogenetic phases. Our results indicate that the relationship between hair [THg] and blood [THg] in pinnipeds is variable and that ontogenetic phase and life history should be considered when interpreting [THg] in these tissues.

  15. A new species of slender coralsnake from Colombia, and its clinal an ontogenetic variation (Serpentes, Elapidae: Leptomicrurus).

    Science.gov (United States)

    Lamar, William W

    2003-01-01

    Leptomicrurus renjifoi is described from tropical semi-deciduous forest of the eastern Colombian Ilanos. It is one of the smallest species in the genus, is most similar to L. scutiventris, and it may be distinguished from known congeners by a combination of color, pattern, and scale characters. Evidence for the recognition of Leptomicrurus is convincing, although its members were recently thought to comprise a closely related assemblage within Micrurus. A supposedly aberrant specimen of L. scutiventris may indicate clinal or ontogenetic variation in pattern.

  16. Nonlinear PDEs

    OpenAIRE

    2015-01-01

    From the Back Cover: The emphasis throughout the present volume is on the practical application of theoretical mathematical models helping to unravel the underlying mechanisms involved in processes from mathematical physics and biosciences. It has been conceived as a unique collection of abstract methods dealing especially with nonlinear partial differential equations (either stationary or evolutionary) that are applied to understand concrete processes involving some important applications re...

  17. Ontogenetic Differences in Dietary Fat Influence Microbiota Assembly in the Zebrafish Gut.

    Science.gov (United States)

    Wong, Sandi; Stephens, W Zac; Burns, Adam R; Stagaman, Keaton; David, Lawrence A; Bohannan, Brendan J M; Guillemin, Karen; Rawls, John F

    2015-09-29

    Gut microbiota influence the development and physiology of their animal hosts, and these effects are determined in part by the composition of these microbial communities. Gut microbiota composition can be affected by introduction of microbes from the environment, changes in the gut habitat during development, and acute dietary alterations. However, little is known about the relationship between gut and environmental microbiotas or about how host development and dietary differences during development impact the assembly of gut microbiota. We sought to explore these relationships using zebrafish, an ideal model because they are constantly immersed in a defined environment and can be fed the same diet for their entire lives. We conducted a cross-sectional study in zebrafish raised on a high-fat, control, or low-fat diet and used bacterial 16S rRNA gene sequencing to survey microbial communities in the gut and external environment at different developmental ages. Gut and environmental microbiota compositions rapidly diverged following the initiation of feeding and became increasingly different as zebrafish grew under conditions of a constant diet. Different dietary fat levels were associated with distinct gut microbiota compositions at different ages. In addition to alterations in individual bacterial taxa, we identified putative assemblages of bacterial lineages that covaried in abundance as a function of age, diet, and location. These results reveal dynamic relationships between dietary fat levels and the microbial communities residing in the intestine and the surrounding environment during ontogenesis. The ability of gut microbiota to influence host health is determined in part by their composition. However, little is known about the relationship between gut and environmental microbiotas or about how ontogenetic differences in dietary fat impact gut microbiota composition. We addressed these gaps in knowledge using zebrafish, an ideal model organism because their

  18. The conceptual framework of ontogenetic trajectories: parallel transport allows the recognition and visualization of pure deformation patterns.

    Science.gov (United States)

    Piras, P; Teresi, L; Traversetti, L; Varano, V; Gabriele, S; Kotsakis, T; Raia, P; Puddu, P E; Scalici, M

    2016-05-01

    Ontogeny is usually studied by analyzing a deformation series spanning over juvenile to adult shapes. In geometric morphometrics, this approach implies applying generalized Procrustes analysis coupled with principal component analysis on multiple individuals or multiple species datasets. The trouble with such a procedure is that it mixes intra- and inter-group variation. While MANCOVA models are relevant statistical/mathematical tools to draw inferences about the similarities of trajectories, if one wants to observe and interpret the morphological deformation alone by filtering inter-group variability, a particular tool, namely parallel transport, is necessary. In the context of ontogenetic trajectories, one should firstly perform separate multivariate regressions between shape and size, using regression predictions to estimate within-group deformations relative to the smallest individuals. These deformations are then applied to a common reference (the mean of per-group smallest individuals). The estimation of deformations can be performed on the Riemannian manifold by using sophisticated connection metrics. Nevertheless, parallel transport can be effectively achieved by estimating deformations in the Euclidean space via ordinary Procrustes analysis. This approach proved very useful in comparing ontogenetic trajectories of species presenting large morphological differences at early developmental stages.

  19. Ontogenetic modulation of branch size, shape, and biomechanics produces diversity across habitats in the Bursera simaruba clade of tropical trees.

    Science.gov (United States)

    Rosell, Julieta A; Olson, Mark E; Aguirre-Hernández, Rebeca; Sánchez-Sesma, Francisco J

    2012-01-01

    Organismal size and shape inseparably interact with tissue biomechanical properties. It is therefore essential to understand how size, shape, and biomechanics interact in ontogeny to produce morphological diversity. We estimated within species branch length-diameter allometries and reconstructed the rates of ontogenetic change along the stem in mechanical properties across the simaruba clade in the tropical tree genus Bursera, measuring 376 segments from 97 branches in nine species in neotropical dry to rain forest. In general, species with stiffer materials had longer, thinner branches, which became stiffer more quickly in ontogeny than their counterparts with more flexible materials. We found a trend from short stature and flexible tissues to tall statures and stiff tissues across an environmental gradient of increasing water availability, likely reflecting a water storage-mechanical support tradeoff. Ontogenetic variation in size, shape, and mechanics results in diversity of habits, for example, rapid length extension, sluggish diameter expansion, and flexible tissues results in a liana, as in Bursera instabilis. Even species of similar habit exhibited notable changes in tissue mechanical properties with increasing size, illustrating the inseparable relationship between organismal proportions and their tissue mechanics in the ontogeny and evolution of morphological diversity.

  20. Biometrie sexual and ontogenetic dimorphism on the marine catfish Genidens genidens (Siluriformes, Ariidae in a tropical estuary

    Directory of Open Access Journals (Sweden)

    Larissa G Paiva

    2015-11-01

    Full Text Available This paper aims to study the ontogenetic sexual dimorphism of Genidens genidens in Guanabara Bay, southeastern coast of Brazil. Altogether 378 specimens were anayzed (233 females and 145 males with total length ranging from 13.3 to 43.5 cm. Specimens were measured for 12 body measurements, sex was identified and maturity stages were recorded and classified. Pearson's linear correlation reveled a significant positive correlation between total length and all other body measures, except for base adipose fin, mouth depth and eye depth for immature females. Analyses nested PERMANOVA desing showed significant differences between maturity stages for each sex, between sexes considering or not maturity stages, indicating a variation in morphometric characteristics driven by sexual dimorphism. Differences among all maturity stages were also found, indicating an ontogenetic morphological difference. But immature individuals didn't differ between sexes indicating that differentiation patterns starts with sexual development. The most important measures differing males and females were related to head characteristics, which appears to be key parameters to evaluate sexual differences. Due to male incubation of fertilized eggs and juvenile individuals <59 mm in their oral cavity, head measures are proposed to be sex dimorphism not related to reproduction, but with post reproductive fase due to ecological and biological needs.

  1. The cooperative Lamb shift in an atomic nanolayer

    CERN Document Server

    Keaveney, James; Krohn, Ulrich; Hughes, Ifan G; Sarkisyan, David; Adams, Charles S

    2012-01-01

    We present an experimental measurement of the cooperative Lamb shift and the Lorentz shift using an atomic nanolayer with tunable thickness and atomic density. The cooperative Lamb shift arises due to the exchange of virtual photons between identical atoms. The interference between the forward and backward propagating virtual fields is confirmed by the thickness dependence of the shift which has a spatial frequency equal to $2k$, i.e. twice that of the optical field. The demonstration of cooperative interactions in an easily scalable system opens the door to a new domain for non-linear optics.

  2. Implementing OpenShift

    CERN Document Server

    Miller, Adam

    2013-01-01

    A standard tutorial-based approach to using OpenShift and deploying custom or pre-built web applications to the OpenShift Online cloud.This book is for software developers and DevOps alike who are interested in learning how to use the OpenShift Platform-as-a-Service for developing and deploying applications, how the environment works on the back end, and how to deploy their very own open source Platform-as-a-Service based on the upstream OpenShift Origin project.

  3. Quantized beam shifts

    CERN Document Server

    Kort-Kamp, W J M; Dalvit, D A R

    2015-01-01

    We predict quantized Imbert-Fedorov, Goos-H\\"anchen, and photonic spin Hall shifts for light beams impinging on a graphene-on-substrate system in an external magnetic field. In the quantum Hall regime the Imbert-Fedorov and photonic spin Hall shifts are quantized in integer multiples of the fine structure constant $\\alpha$, while the Goos- H\\"anchen ones in multiples of $\\alpha^2$. We investigate the influence on these shifts of magnetic field, temperature, and material dispersion and dissipation. An experimental demonstration of quantized beam shifts could be achieved at terahertz frequencies for moderate values of the magnetic field.

  4. Integrated trend assessment of ecosystem changes in the Limfjord (Denmark): evidence of a recent regime shift?

    DEFF Research Database (Denmark)

    Tomczak, Maciej Tomasz; Dinesen, Grete E.; Hoffmann, Erik

    2012-01-01

    ), jellyfish, common shore crab, starfish and blue mussels. We interpret this change as a regime shift that showed a similar temporal pattern to regime shifts identified in adjacent seas. The observed changes in trophic interactions and food web reorganisation suggested a non-linear regime shift. The analyses...

  5. Daily otolith growth and ontogenetic geochemical signatures of age-0 anchovy (Engraulis encrasicolus in the Gulf of Cádiz (SW Spain.

    Directory of Open Access Journals (Sweden)

    I. A. CATALÁN

    2015-01-01

    Full Text Available The European anchovy fishery in the Gulf of Cádiz (ICES Division IXa South is largely influenced by age-0 individuals. Knowledge of young of the year growth dynamics is crucial for management, yet data on daily growth are lacking in the area. Linking growth patterns to the environment requires information on habitat occupancy through ontogeny of the fish that reach the fishery, as anchovy use different areas of the Gulf and the Guadalquivir Estuary through development. We describe the growth dynamics of age-0 anchovy through otolith microstructure analysis, and couple these data with data on microchemical signals in the otoliths to shed light into habitat use and growth dynamics in the area. Age-0 anchovy captured in September, 2011 in the Gulf ranged from 3 to 6 months old for similar sizes, with average growth rates varying twofold. Individual non-linear growth curves showed that maximum otolith growth was positively correlated with the date of spawning, which in turn was negatively correlated with the time to reach maximum growth. There was no correlation between growth parameters and body length or condition (Fulton K at capture. The strontium:calcium (Sr/Ca and magnesium:calcium (Mg/Ca ratios were significantly lower at the edge of the otolith (approximately the age of capture than at ages corresponding to larval and early juveniles (<60 days old, but values fell within typical estuarine-dwelling species. The barium:calcium ratio (Ba/Ca increased significantly in the edge of the otolith, which possibly resulted from residency in highly productive coastal waters or from ontogenetic effects. The variance in otolith elemental ratios was larger at otolith back-calculated ages around 50 days old, age which coincides with the presumed closer dependence of estuarine waters. Our data are a first step towards understanding the contribution of the estuarine system to the fishery of anchovy in the Gulf. The limitations of the approach and future

  6. DSP Approach to the Design of Nonlinear Optical Devices

    Directory of Open Access Journals (Sweden)

    Steve Blair

    2005-06-01

    Full Text Available Discrete-time signal processing (DSP tools have been used to analyze numerous optical filter configurations in order to optimize their linear response. In this paper, we propose a DSP approach to design nonlinear optical devices by treating the desired nonlinear response in the weak perturbation limit as a discrete-time filter. Optimized discrete-time filters can be designed and then mapped onto a specific optical architecture to obtain the desired nonlinear response. This approach is systematic and intuitive for the design of nonlinear optical devices. We demonstrate this approach by designing autoregressive (AR and autoregressive moving average (ARMA lattice filters to obtain a nonlinear phase shift response.

  7. OpenShift cookbook

    CERN Document Server

    Gulati, Shekhar

    2014-01-01

    If you are a web application developer who wants to use the OpenShift platform to host your next big idea but are looking for guidance on how to achieve this, then this book is the first step you need to take. This is a very accessible cookbook where no previous knowledge of OpenShift is needed.

  8. Shifting employment revisited

    NARCIS (Netherlands)

    Cremers, Jan; Gramuglia, Alessia

    2014-01-01

    The CLR-network examined in 2006 the phenomenon of undeclared labour, with specific regard to the construction sector. The resulting study, Shifting Employment: undeclared labour in construction (Shifting-study hereafter), gave evidence that this is an area particularly affected by undeclared activi

  9. Making Shifts toward Proficiency

    Science.gov (United States)

    McGatha, Maggie B.; Bay-Williams, Jennifer M.

    2013-01-01

    The Leading for Mathematical Proficiency (LMP) Framework (Bay-Williams et al.) has three components: (1) The Standards for Mathematical Practice; (2) Shifts in classroom practice; and (3) Teaching skills. This article briefly describes each component of the LMP framework and then focuses more in depth on the second component, the shifts in…

  10. Shifted Independent Component Analysis

    DEFF Research Database (Denmark)

    Mørup, Morten; Madsen, Kristoffer Hougaard; Hansen, Lars Kai

    2007-01-01

    Delayed mixing is a problem of theoretical interest and practical importance, e.g., in speech processing, bio-medical signal analysis and financial data modelling. Most previous analyses have been based on models with integer shifts, i.e., shifts by a number of samples, and have often been carrie...

  11. A novel length back-calculation approach accounting for ontogenetic changes in the fish length - otolith size relationship during the early life of sprat (Sprattus sprattus)

    DEFF Research Database (Denmark)

    Guenther, Claudia C.; Temming, Axel; Baumann, Hannes

    2012-01-01

    An individual-based length back-calculation method was developed for juvenile Baltic sprat (Sprattus sprattus), accounting for ontogenetic changes in the relationship between fish length and otolith length. In sprat, metamorphosis from larvae to juveniles is characterized by the coincidence of low...

  12. Visual sensitivities tuned by heterochronic shifts in opsin gene expression

    Directory of Open Access Journals (Sweden)

    McFarland William N

    2008-05-01

    Full Text Available Abstract Background Cichlid fishes have radiated into hundreds of species in the Great Lakes of Africa. Brightly colored males display on leks and vie to be chosen by females as mates. Strong discrimination by females causes differential male mating success, rapid evolution of male color patterns and, possibly, speciation. In addition to differences in color pattern, Lake Malawi cichlids also show some of the largest known shifts in visual sensitivity among closely related species. These shifts result from modulated expression of seven cone opsin genes. However, the mechanisms for this modulated expression are unknown. Results In this work, we ask whether these differences might result from changes in developmental patterning of cone opsin genes. To test this, we compared the developmental pattern of cone opsin gene expression of the Nile tilapia, Oreochromis niloticus, with that of several cichlid species from Lake Malawi. In tilapia, quantitative polymerase chain reaction showed that opsin gene expression changes dynamically from a larval gene set through a juvenile set to a final adult set. In contrast, Lake Malawi species showed one of two developmental patterns. In some species, the expressed gene set changes slowly, either retaining the larval pattern or progressing only from larval to juvenile gene sets (neoteny. In the other species, the same genes are expressed in both larvae and adults but correspond to the tilapia adult genes (direct development. Conclusion Differences in visual sensitivities among species of Lake Malawi cichlids arise through heterochronic shifts relative to the ontogenetic pattern of the tilapia outgroup. Heterochrony has previously been shown to be a powerful mechanism for change in morphological evolution. We found that altering developmental expression patterns is also an important mechanism for altering sensory systems. These resulting sensory shifts will have major impacts on visual communication and could help

  13. Visual sensitivities tuned by heterochronic shifts in opsin gene expression

    Science.gov (United States)

    Carleton, Karen L; Spady, Tyrone C; Streelman, J Todd; Kidd, Michael R; McFarland, William N; Loew, Ellis R

    2008-01-01

    Background Cichlid fishes have radiated into hundreds of species in the Great Lakes of Africa. Brightly colored males display on leks and vie to be chosen by females as mates. Strong discrimination by females causes differential male mating success, rapid evolution of male color patterns and, possibly, speciation. In addition to differences in color pattern, Lake Malawi cichlids also show some of the largest known shifts in visual sensitivity among closely related species. These shifts result from modulated expression of seven cone opsin genes. However, the mechanisms for this modulated expression are unknown. Results In this work, we ask whether these differences might result from changes in developmental patterning of cone opsin genes. To test this, we compared the developmental pattern of cone opsin gene expression of the Nile tilapia, Oreochromis niloticus, with that of several cichlid species from Lake Malawi. In tilapia, quantitative polymerase chain reaction showed that opsin gene expression changes dynamically from a larval gene set through a juvenile set to a final adult set. In contrast, Lake Malawi species showed one of two developmental patterns. In some species, the expressed gene set changes slowly, either retaining the larval pattern or progressing only from larval to juvenile gene sets (neoteny). In the other species, the same genes are expressed in both larvae and adults but correspond to the tilapia adult genes (direct development). Conclusion Differences in visual sensitivities among species of Lake Malawi cichlids arise through heterochronic shifts relative to the ontogenetic pattern of the tilapia outgroup. Heterochrony has previously been shown to be a powerful mechanism for change in morphological evolution. We found that altering developmental expression patterns is also an important mechanism for altering sensory systems. These resulting sensory shifts will have major impacts on visual communication and could help drive cichlid speciation

  14. Estimating the Ontogenetic Status of an Enantiornithine Bird from the Lower Barremian of El Montsec, Central Pyrenees, Spain

    Directory of Open Access Journals (Sweden)

    Buscalioni, A. D.

    2006-12-01

    Full Text Available An Enantiornithes specimen from El Montsec was initially described as an immature individual based upon qualitative traits such as its relatively large orbit and overall proportions of the skull and the postcranium. In this study we re-evaluate the precise determination of the ontogenetic stage of this individual, establishing a cross-talk among taphonomic, anatomic, and morphometric data. The exceptional preservation of the specimen has allowed pondering ontogenetic influence versus preservational bias in features like the external patterns of bone surfaces, instead of being aprioristically considered due to taphonomic alterations only. The rough texture of the periosteal bone associated with pores in the distal, proximal and mid-shaft areas of the humeral shaft, indicates a subadult stage when compared with long bones of modern birds. Forelimb proportions of embryo and juvenile Enanthiornithes are equivalent to those of adult individuals of other taxa within this clade, though this is not a reliable criterion for establishing a precise ontogenetic stage. The El Montsec specimen may be attributed a close adulthood, yet only if growth regimes in Enantiornithes are considered equivalent to those in Neornithes birds.Un ejemplar de Enantiornithes del Montsec fue inicialmente descrito como un individuo inmaduro sobre la base de caracteres cualitativos tales como su órbita relativamente grande y sus proporciones generales en cuerpo y cráneo. En este estudio se realiza una reevaluación del estado ontogenético preciso de este individuo, estableciendo una argumentación cruzada con datos tafonómicos, anatómicos y morfométricos. La preservación excepcional de este ejemplar ha permitido ponderar la influencia ontogenética versus el sesgo tafonómico en caracteres como los patrones externos de las superficies óseas, en lugar de considerarlos apriorísticamente como debidos únicamente a alteraciones tafonómicas. La textura rugosa del periostio

  15. Ontogenetic characterization of the intestinal microbiota of channel catfish through 16S rRNA gene sequencing reveals insights on temporal shifts and the influence of environmental microbes

    Science.gov (United States)

    Aquaculture recently overtook capture fisheries as the largest producer of food fish; however, to continue increasing fish production the industry is looking for ways of improving fish health and growth. Pre- and probiotic supplementation has gained attention as a means of solving these issues, howe...

  16. The vascular pattern in the flower of some Mesembryanthemaceae: Aptenia cordifolia and Dorotheanthus bellidiformis. The effect of an ontogenetical shifting on the vascular pattern and vascular conservatism

    NARCIS (Netherlands)

    Meulen-Bruijns, van der C.

    1976-01-01

    1. The vascular pattern in the flower at various stages of maturity of Aptenia cordifolia and Dorotheanthus bellidiformis is examined. 2. The vascular pattern of Dorotheanthus has been compared with that of Aptenia: typologically, Dorotheanthus is derived from Aptenia. 3. The vascular pattern of

  17. Nonlinear resonances

    CERN Document Server

    Rajasekar, Shanmuganathan

    2016-01-01

    This introductory text presents the basic aspects and most important features of various types of resonances and anti-resonances in dynamical systems. In particular, for each resonance, it covers the theoretical concepts, illustrates them with case studies, and reviews the available information on mechanisms, characterization, numerical simulations, experimental realizations, possible quantum analogues, applications and significant advances made over the years. Resonances are one of the most fundamental phenomena exhibited by nonlinear systems and refer to specific realizations of maximum response of a system due to the ability of that system to store and transfer energy received from an external forcing source. Resonances are of particular importance in physical, engineering and biological systems - they can prove to be advantageous in many applications, while leading to instability and even disasters in others. The book is self-contained, providing the details of mathematical derivations and techniques invo...

  18. Nonlinear Dynamics

    Institute of Scientific and Technical Information of China (English)

    1996-01-01

    3.1 A Unified Nonlinear Feedback Functional Method for Study Both Control and Synchronization of Spatiotemporal Chaos Fang Jinqing Ali M. K. (Department of Physics, The University of Lethbridge,Lethbridge, Alberta T1K 3M4,Canada) Two fundamental questions dominate future chaos control theories.The first is the problem of controlling hyperchaos in higher dimensional systems.The second question has yet to be addressed:the problem of controlling spatiotemporal chaos in a spatiotemporal system.In recent years, control and synchronization of spatiotemporal chaos and hyperchaos have became a much more important and challenging subject. The reason for this is the control and synchronism of such behaviours have extensive and great potential of interdisciplinary applications, such as security communication, information processing, medicine and so on. However, this subject is not much known and remains an outstanding open.

  19. Non-linear Frequency Scaling Algorithm for FMCW SAR Data

    NARCIS (Netherlands)

    Meta, A.; Hoogeboom, P.; Ligthart, L.P.

    2006-01-01

    This paper presents a novel approach for processing data acquired with Frequency Modulated Continuous Wave (FMCW) dechirp-on-receive systems by using a non-linear frequency scaling algorithm. The range frequency non-linearity correction, the Doppler shift induced by the continuous motion and the ran

  20. The ontogenetic origins of mirror neurons: evidence from ‘tool-use’ and ‘audiovisual’ mirror neurons

    Science.gov (United States)

    Cook, Richard

    2012-01-01

    Since their discovery, mirror neurons—units in the macaque brain that discharge both during action observation and execution—have attracted considerable interest. Whether mirror neurons are an innate endowment or acquire their sensorimotor matching properties ontogenetically has been the subject of intense debate. It is widely believed that these units are an innate trait; that we are born with a set of mature mirror neurons because their matching properties conveyed upon our ancestors an evolutionary advantage. However, an alternative view is that mirror neurons acquire their matching properties during ontogeny, through correlated experience of observing and performing actions. The present article re-examines frequently overlooked neurophysiological reports of ‘tool-use’ and ‘audiovisual’ mirror neurons within the context of this debate. It is argued that these findings represent compelling evidence that mirror neurons are a product of sensorimotor experience, and not an innate endowment. PMID:22573832

  1. Ontogenetic body-mass scaling of nitrogen excretion relates to body surface area in diverse pelagic invertebrates

    DEFF Research Database (Denmark)

    Hirst, Andrew G.; Lilley, M.K.S.; Glazier, D.S.;

    2016-01-01

    Many physiological and ecological processes depend on body size and the supply of limiting nutrients. Hence, it is important to derive quantitative predictions based on a mechanistic understanding of the influence of body size on metabolic rate and on the ratios of consumed to excreted elements. ....... Diverse pelagic invertebrates that dominate vast open water ecosystems falsify the predictions of general metabolic scaling theories built upon resource-transport networks, but support predictions of surface-area dependent theory........ Among diverse pelagic invertebrates that change shape during ontogeny, recent analysis has demonstrated a significant positive correlation between the body-mass allometry of respiration rates (measured as the ontogenetic body mass-scaling exponent bR) and the allometry of body surface area (b...

  2. Setting the reference for the use of Chironomus sancticaroli (Diptera: Chironomidae as bioindicator: Ontogenetic pattern of larval head structures

    Directory of Open Access Journals (Sweden)

    Débora Rebechi

    2012-04-01

    Full Text Available Species of Chironomidae are widely used as bioindicators of water quality, since their larvae undergo morphological deformities when in contact with sediment contaminated with chemicals. In this work we endeavored to study the morphology of head structures (antennae, mandible, mentum, pecten epipharyngis, ventromental plate and premandible throughout the development of the four larval instars of Chironomus sancticaroli Strixino & Strixino, 1981, which can be used in environmental impact analyses. Our results show that it is possible to differentiate among larval instars by doing a quantitative analysis on the number of striae on the ventromental plates. The six structures analyzed changed during larval ontogeny. These changes are part of the ontogeny of the immature stages not exposed to xenobiotics. We believe that the morphological pattern defined in this work can be used for comparisons with ontogenetic changes observed in field studies conducted in polluted environments.

  3. Spatial, seasonal and ontogenetic changes in food resource use by a piscivore fish in two Pantanal lagoons, Brazil

    Directory of Open Access Journals (Sweden)

    Michele de Faveri Gimenes

    Full Text Available We evaluated the factors that affect the food resource use by Plagioscion ternetzi using three approaches: spatial, seasonal, and ontogenetic changes. Fish were sampled between March 2000 and February 2001 and March 2003 and February 2004 in the Sinhá Mariana and Chacororé lagoons, wetlands of the Pantanal Matogrossense. Fish was the predominant food in the diet, but shrimps and insects have been also consumed. The diet of P. ternetzi was significantly different between lagoons, but no difference was detected between dry and flood periods. In Sinhá Mariana lagoon, the diet mainly consisted of Psectrogaster curviventris, in both periods. In the flood period in Chacororé lagoon, fish was the dominant food (especially Astyanax spp., Leporinus spp. and Schizodon borellii, and in dry, shrimp and Eigenmannia spp. The Spearman coefficient indicated no correlation between the abundance of caught fish species and their abundances in the diet of P. ternetzi in Sinhá Mariana lagoon, and a strong negative correlation at Chacororé lagoon, although some very consumed preys were numerically abundant in the environment. Despite the intake of insect and shrimp, P. ternetzi was piscivorous since immature stages. A correlation between each item and the size classes of P. ternetziwas tested by means of Pearson correlation that showed that the changes in the use of resources over ontogenetic development have been represented by increased intake of P. curviventris in Mariana lagoon, and S. borellii at Chacororé lagoon, accompanying the increase in the size of the predator. In this way, the results suggest that several factors can be involved in the use of food resources by P. ternezi, among them the abundance, the size and morphological characteristics of the prey.

  4. Nonlinear Materials Characterization Facility

    Data.gov (United States)

    Federal Laboratory Consortium — The Nonlinear Materials Characterization Facility conducts photophysical research and development of nonlinear materials operating in the visible spectrum to protect...

  5. Choice Shift in Opinion Network Dynamics

    Science.gov (United States)

    Gabbay, Michael

    Choice shift is a phenomenon associated with small group dynamics whereby group discussion causes group members to shift their opinions in a more extreme direction so that the mean post-discussion opinion exceeds the mean pre-discussion opinion. Also known as group polarization, choice shift is a robust experimental phenomenon and has been well-studied within social psychology. In opinion network models, shifts toward extremism are typically produced by the presence of stubborn agents at the extremes of the opinion axis, whose opinions are much more resistant to change than moderate agents. However, we present a model in which choice shift can arise without the assumption of stubborn agents; the model evolves member opinions and uncertainties using coupled nonlinear differential equations. In addition, we briefly describe the results of a recent experiment conducted involving online group discussion concerning the outcome of National Football League games are described. The model predictions concerning the effects of network structure, disagreement level, and team choice (favorite or underdog) are in accord with the experimental results. This research was funded by the Office of Naval Research and the Defense Threat Reduction Agency.

  6. Shift Verification and Validation

    Energy Technology Data Exchange (ETDEWEB)

    Pandya, Tara M. [Oak Ridge National Lab. (ORNL), Oak Ridge, TN (United States); Evans, Thomas M. [Oak Ridge National Lab. (ORNL), Oak Ridge, TN (United States); Davidson, Gregory G [Oak Ridge National Lab. (ORNL), Oak Ridge, TN (United States); Johnson, Seth R. [Oak Ridge National Lab. (ORNL), Oak Ridge, TN (United States); Godfrey, Andrew T. [Oak Ridge National Lab. (ORNL), Oak Ridge, TN (United States)

    2016-09-07

    This documentation outlines the verification and validation of Shift for the Consortium for Advanced Simulation of Light Water Reactors (CASL). Five main types of problems were used for validation: small criticality benchmark problems; full-core reactor benchmarks for light water reactors; fixed-source coupled neutron-photon dosimetry benchmarks; depletion/burnup benchmarks; and full-core reactor performance benchmarks. We compared Shift results to measured data and other simulated Monte Carlo radiation transport code results, and found very good agreement in a variety of comparison measures. These include prediction of critical eigenvalue, radial and axial pin power distributions, rod worth, leakage spectra, and nuclide inventories over a burn cycle. Based on this validation of Shift, we are confident in Shift to provide reference results for CASL benchmarking.

  7. Shift Verification and Validation

    Energy Technology Data Exchange (ETDEWEB)

    Pandya, Tara M. [Oak Ridge National Lab. (ORNL), Oak Ridge, TN (United States); Evans, Thomas M. [Oak Ridge National Lab. (ORNL), Oak Ridge, TN (United States); Davidson, Gregory G [Oak Ridge National Lab. (ORNL), Oak Ridge, TN (United States); Johnson, Seth R. [Oak Ridge National Lab. (ORNL), Oak Ridge, TN (United States); Godfrey, Andrew T. [Oak Ridge National Lab. (ORNL), Oak Ridge, TN (United States)

    2016-09-07

    This documentation outlines the verification and validation of Shift for the Consortium for Advanced Simulation of LightWater Reactors (CASL). Fivemain types of problems were used for validation: small criticality benchmark problems; full-core reactor benchmarks for light water reactors; fixed source coupled neutron-photon dosimetry benchmarks; depletion/burnup benchmarks; and full-core reactor performance benchmarks. We compared Shift results to measured data and other simulated Monte Carlo radiation transport code results and found very good agreement in a variety of comparison measures. These include prediction of critical eigenvalue, radial and axial pin power distributions, rod worth, leakage spectra, and nuclide inventories over a burn cycle. Based on this validation of Shift, we are confident in Shift to provide reference results for CASL benchmarking.

  8. Z-scan measurement of the nonlinear refractive index of graphene.

    Science.gov (United States)

    Zhang, Han; Virally, Stéphane; Bao, Qiaoliang; Ping, Loh Kian; Massar, Serge; Godbout, Nicolas; Kockaert, Pascal

    2012-06-01

    Under strong laser illumination, few-layer graphene exhibits both a transmittance increase due to saturable absorption and a nonlinear phase shift. Here, we unambiguously distinguish these two nonlinear optical effects and identify both real and imaginary parts of the complex nonlinear refractive index of graphene. We show that graphene possesses a giant nonlinear refractive index n(2)≃10(-7) cm(2) W(-1), almost 9 orders of magnitude larger than bulk dielectrics. We find that the nonlinear refractive index decreases with increasing excitation flux but slower than the absorption. This suggests that graphene may be a very promising nonlinear medium, paving the way for graphene-based nonlinear photonics.

  9. Photonic surfaces for designable nonlinear power shaping

    Energy Technology Data Exchange (ETDEWEB)

    Biswas, Roshni, E-mail: rbiswas@usc.edu; Povinelli, Michelle L. [Ming Hsieh Department of Electrical Engineering, University of Southern California, Los Angeles, California 90089 (United States)

    2015-02-09

    We propose a method for designing nonlinear input-output power response based on absorptive resonances of nanostructured surfaces. We show that various power transmission trends can be obtained by placing a photonic resonance mode at the appropriate detuning from the laser wavelength. We demonstrate our results in a silicon photonic crystal slab at a laser wavelength of 808 nm. We quantify the overall spectral red shift as a function of laser power. The shift results from absorptive heating and the thermo-optic effect. We then demonstrate devices with increasing, decreasing, and non-monotonic transmission as a function of laser power. The transmission changes are up to 7.5 times larger than in unpatterned silicon. The strong nonlinear transmission is due to a combination of resonantly enhanced absorption, reduced thermal conductivity, and the resonant transmission lineshape. Our results illustrate the possibility of designing different nonlinear power trends within a single materials platform at a given wavelength of interest.

  10. Allometric shape change of the lower pharyngeal jaw correlates with a dietary shift to piscivory in a cichlid fish

    Science.gov (United States)

    Hellig, Christoph J.; Kerschbaumer, Michaela; Sefc, Kristina M.; Koblmüller, Stephan

    2010-07-01

    The morphological versatility of the pharyngeal jaw of cichlid fishes is assumed to represent a key factor facilitating their unparalleled trophic diversification and explosive radiation. It is generally believed that the functional design of an organism relates to its ecology, and thus, specializations to different diets are typically associated with distinct morphological designs, especially manifested in the cichlids’ pharyngeal jaw apparatus. Thereby, the lower pharyngeal jaw (LPJ) incorporates some of the most predictive features for distinct diet-related morphotypes. Thus, considering that piscivorous cichlids experience an ontogenetic dietary shift from typically various kinds of invertebrates to fish, concomitant morphological changes in the LPJ are expected. Using Lepidiolamprologus elongatus, a top predator in the shallow rocky habitat of Lake Tanganyika, as model, and applying geometric and traditional morphometric techniques, we demonstrate an allometric change in ontogenetic LPJ shape development coinciding with the completion of the dietary shift toward piscivory. The piscivorous LPJ morphotype is initiated in juvenile fish by increasing elongation and narrowing of the LPJ and—when the fish reach a size of 80-90 mm standard length—further refined by the elongation of the posterior muscular processes, which serve as insertion for the fourth musculus levator externus. The enlarged muscular processes of the fully mature piscivorous morphotype provide for the construction of a powerful lever system, which allows the large individuals to process large prey fish and rely on exclusive piscivory.

  11. Evidence of compositional and ultrastructural shifts during the development of calcareous tubes in the biofouling tubeworm, Hydroides elegans.

    Science.gov (United States)

    Chan, Vera Bin San; Vinn, Olev; Li, Chaoyi; Lu, Xingwen; Kudryavtsev, Anatoliy B; Schopf, J William; Shih, Kaimin; Zhang, Tong; Thiyagarajan, Vengatesen

    2015-03-01

    The serpulid tubeworm, Hydroides elegans, is an ecologically and economically important species whose biology has been fairly well studied, especially in the context of larval development and settlement on man-made objects (biofouling). Nevertheless, ontogenetic changes associated with calcareous tube composition and structures have not yet been studied. Here, the ultrastructure and composition of the calcareous tubes built by H. elegans was examined in the three early calcifying juvenile stages and in the adult using XRD, FTIR, ICP-OES, SEM and Raman spectroscopy. Ontogenetic shifts in carbonate mineralogy were observed, for example, juvenile tubes contained more amorphous calcium carbonate and were predominantly aragonitic whereas adult tubes were bimineralic with considerably more calcite. The mineral composition gradually shifted during the tube development as shown by a decrease in Sr/Ca and an increase of Mg/Ca ratios with the tubeworm's age. The inner tube layer contained calcite, whereas the outer layer contained aragonite. Similarly, the tube complexity in terms of ultrastructure was associated with development. The sequential appearance of unoriented ultrastructures followed by oriented ultrastructures may reflect the evolutionary history of serpulid tube biominerals. As aragonitic structures are more susceptible to dissolution under ocean acidification (OA) conditions but are more difficult to be removed by anti-fouling treatments, the early developmental stages of the tubeworms may be vulnerable to OA but act as the important target for biofouling control.

  12. Nonlinear singular vectors and nonlinear singular values

    Institute of Scientific and Technical Information of China (English)

    2000-01-01

    A novel concept of nonlinear singular vector and nonlinear singular value is introduced, which is a natural generalization of the classical linear singular vector and linear singular value to the nonlinear category. The optimization problem related to the determination of nonlinear singular vectors and singular values is formulated. The general idea of this approach is demonstrated by a simple two-dimensional quasigeostrophic model in the atmospheric and oceanic sciences. The advantage and its applications of the new method to the predictability, ensemble forecast and finite-time nonlinear instability are discussed. This paper makes a necessary preparation for further theoretical and numerical investigations.

  13. Enhanced Kerr electro-optic nonlinearity through cascaded Pockels effects

    CERN Document Server

    Li, Guang-Zhen; Jiang, Hao-Wei; Chen, Xian-Feng

    2015-01-01

    We demonstrated a large enhancement of Kerr electro-optic nonlinearity through cascaded Pockels effects in a domain inversion ferroelectric crystal. We designed a structure that can implement the cascaded Pockels effects and second-harmonic generation simultaneously. The energy coupling between the fundamental lights of different polarizations led to a large nonlinear phase shift, and thus an effective electro-optic nonlinear refractive index. The effective nonlinearity can be either positive or negative, causing the second-harmonic spectra to move towards the coupling center, which in turn, offered us a way to measure the effective electro-optic nonlinear refractive index. The corresponding enhanced Kerr electro-optic nonlinearity is more than three orders of magnitude higher than the intrinsic value. These results open a door to manipulate the nonlinear phase by applying external electric field instead of light intensity in noncentrosymmetric crystals.

  14. Giant Goos-Hänchen shift via spontaneous generated coherence

    Science.gov (United States)

    Ziauddin

    2015-11-01

    The influence of spontaneous generated coherence (SGC) on the Goos-Hänchen (GH) shift in the reflected light is presented. A weak probe light is incident on a cavity containing three-level gaseous atomic medium consist of 85Rb atoms. The atom-field interaction follows electromagnetically induced transparency configuration, and the SGC modifies the dispersion and absorption properties of a system [Y. Niu and S. Gong, Phys. Rev. A 73, 053811 (2006)]. The SGC enhances the Kerr nonlinearity which leads to giant negative and positive GH shifts in the reflected light. Further, the control of negative and positive GH shifts is achieved via manipulation of probe field detuning.

  15. Large microwave phase shift and small distortion in an integrated waveguide device

    DEFF Research Database (Denmark)

    Öhman, Filip; Sales, Salvador; Chen, Yaohui;

    2007-01-01

    We have obtained a tunable phase shift of 150 degrees in an integrated semiconductor waveguide by optimizing the interplay of fast and slow light effects. Furthermore, the distortions imposed by device nonlinearities have been quantified....

  16. Protein Chemical Shift Prediction

    CERN Document Server

    Larsen, Anders S

    2014-01-01

    The protein chemical shifts holds a large amount of information about the 3-dimensional structure of the protein. A number of chemical shift predictors based on the relationship between structures resolved with X-ray crystallography and the corresponding experimental chemical shifts have been developed. These empirical predictors are very accurate on X-ray structures but tends to be insensitive to small structural changes. To overcome this limitation it has been suggested to make chemical shift predictors based on quantum mechanical(QM) calculations. In this thesis the development of the QM derived chemical shift predictor Procs14 is presented. Procs14 is based on 2.35 million density functional theory(DFT) calculations on tripeptides and contains corrections for hydrogen bonding, ring current and the effect of the previous and following residue. Procs14 is capable at performing predictions for the 13CA, 13CB, 13CO, 15NH, 1HN and 1HA backbone atoms. In order to benchmark Procs14, a number of QM NMR calculatio...

  17. Ontogenetic resource-use strategies in a rare long-lived cycad along environmental gradients.

    Science.gov (United States)

    Álvarez-Yépiz, Juan C; Cueva, Alejandro; Dovčiak, Martin; Teece, Mark; Yepez, Enrico A

    2014-01-01

    Functional traits can drive plant responses to short- and long-term stressful conditions, with potential effects on species persistence in local habitats, changes in population size and structure, and potential species range shifts in changing environments. We investigated whether ecophysiological traits in a rare cycad vary along environmental gradients and with ontogeny to understand intra-specific resource-use variation (e.g. symbiotic nitrogen fixation, nitrogen- and water-use efficiency) and possible species adaptations for different environments. Environmental gradients were characterized with 14 soil and topographic variables. Nitrogen- and water-use efficiency improved with ontogeny (from seedling to juvenile and adult stages) but declined as soil fertility decreased with increasing elevation. Conversely, reliance on symbiotic nitrogen fixation increased with elevation and varied slightly with ontogeny. Improved water-use efficiency at lower elevation and nitrogen fixation at higher elevation may represent key functional strategies for maintaining the lower and upper altitudinal species range limits, especially in arid environments where stressful conditions are intensifying due to climatic and land-use changes. In addition to facilitation linked to the regeneration niche, improved resource-use efficiency linked to the adult niche may strongly influence cycad distribution and persistence in contemporary environments. A functional approach to conservation of rare or endangered plant species may be needed in order to target the most sensitive stages to changing environmental conditions and to better understand potential range shifts and adaptive responses to global land-use and climate changes.

  18. Experimental demonstration of optical switching and routing via four-wave mixing spatial shift.

    Science.gov (United States)

    Nie, Zhiqiang; Zheng, Huaibin; Zhang, Yanpeng; Zhao, Yan; Zuo, Cuicui; Li, Changbiao; Chang, Hong; Xiao, Min

    2010-01-18

    We demonstrate the shift characteristics of four-wave mixing (FWM) beam spots which are controlled by the strong laser fields via the large cross-Kerr nonlinearity. The shift distances and directions are determined by the nonlinear dispersions. Based on such spatial displacements of the FWM beams, as well as the probe beam, we experimentally demonstrate spatial optical switching for one beam or multiple optical beams, which can be used for all-optical switching, switching arrays and routers.

  19. Ontogenetic Tooth Reduction in Stenopterygius quadriscissus (Reptilia: Ichthyosauria: Negative Allometry, Changes in Growth Rate, and Early Senescence of the Dental Lamina.

    Directory of Open Access Journals (Sweden)

    Daniel G Dick

    Full Text Available We explore the functional, developmental, and evolutionary processes which are argued to produce tooth reduction in the extinct marine reptile Stenopterygius quadriscissus (Reptilia: Ichthyosauria. We analyze the relationship between mandible growth and tooth size, shape, and count, to establish an ontogenetic trend. The pattern in S. quadriscissus is consistent with hypotheses of tooth size reduction by neutral selection, and this unusual morphology (a functionally edentulous rostrum was produced by a series of different evolutionary developmental changes that are known for other taxa showing tooth reduction and loss. Specifically, this species evolved functional edentulism by evolutionary changes in the growth allometry of the dentition and by altering growth rates through ontogeny. This observation supports previous hypotheses that S. quadriscissus underwent ontogenetic tooth reduction. Tooth reduction in S. quadriscissus may be caused by unique selective pressures resulting from prey choice and feeding behavior, expanding our current understanding of the mechanisms producing tooth reduction.

  20. NONLINEAR EXPECTATIONS AND NONLINEAR MARKOV CHAINS

    Institute of Scientific and Technical Information of China (English)

    PENG SHIGE

    2005-01-01

    This paper deals with nonlinear expectations. The author obtains a nonlinear generalization of the well-known Kolmogorov's consistent theorem and then use it to construct filtration-consistent nonlinear expectations via nonlinear Markov chains. Compared to the author's previous results, i.e., the theory of g-expectations introduced via BSDE on a probability space, the present framework is not based on a given probability measure. Many fully nonlinear and singular situations are covered. The induced topology is a natural generalization of Lp-norms and L∞-norm in linear situations.The author also obtains the existence and uniqueness result of BSDE under this new framework and develops a nonlinear type of von Neumann-Morgenstern representation theorem to utilities and present dynamic risk measures.

  1. Switching behaviour of a nonlinear Mach–Zehnder interferometer

    Indian Academy of Sciences (India)

    Arpita Srivastava; Punya Prasanna Paltani; S Medhekar

    2010-04-01

    In the present paper, a detailed investigation on the switching behaviour of a nonlinear Mach–Zehnder interferometer (NMZI) has been carried out using beam propagation method (BPM). A thorough investigation on input vs. output characteristic has been carried out by varying different parameters like length of the arms, refractive index of the linear/nonlinear arm, wavelength of the input beams and nonlinear coefficient of the material of the nonlinear arm. The input vs. output characteristic has also been investigated by shifting the balance point of the NMZI. The present paper provides a physically intuitive understanding of the effect of change in different parameters of the NMZI on its switching behaviour.

  2. Rotational Doppler effect in nonlinear optics

    Science.gov (United States)

    Li, Guixin; Zentgraf, Thomas; Zhang, Shuang

    2016-08-01

    The translational Doppler effect of electromagnetic and sound waves has been successfully applied in measurements of the speed and direction of vehicles, astronomical objects and blood flow in human bodies, and for the Global Positioning System. The Doppler effect plays a key role for some important quantum phenomena such as the broadened emission spectra of atoms and has benefited cooling and trapping of atoms with laser light. Despite numerous successful applications of the translational Doppler effect, it fails to measure the rotation frequency of a spinning object when the probing wave propagates along its rotation axis. This constraint was circumvented by deploying the angular momentum of electromagnetic waves--the so-called rotational Doppler effect. Here, we report on the demonstration of rotational Doppler shift in nonlinear optics. The Doppler frequency shift is determined for the second harmonic generation of a circularly polarized beam passing through a spinning nonlinear optical crystal with three-fold rotational symmetry. We find that the second harmonic generation signal with circular polarization opposite to that of the fundamental beam experiences a Doppler shift of three times the rotation frequency of the optical crystal. This demonstration is of fundamental significance in nonlinear optics, as it provides us with insight into the interaction of light with moving media in the nonlinear optical regime.

  3. A Shift of Power

    Institute of Scientific and Technical Information of China (English)

    2008-01-01

    Administrative reforms are shifting prefecture government powers to the county level in an effort to boost local economies on July 8, the government of China’s southernmost Hainan Province announced that it was to hand over 177 of its administrative powers to county-level governments. The move practically dismantled the powers of the

  4. Middle Jurassic spinicaudatan Shizhuestheria from the Sichuan Basin and its ontogenetic implication

    Institute of Scientific and Technical Information of China (English)

    HIRANO; Hiromichi; KOZAI; Takeshi; SAKAI; Takashi

    2009-01-01

    The diagnosis of the Middle Jurassic spinicaudatan genus Shizhuestheria is revised following the re-examination under a scanning electron microscope of a type specimen from Yanxi of Shizhu, and the newly collected specimens from Xintiangou of Chengjiang Town, Beibei, Chongqing. Small-sized reticulations occur not only in the middle-sized reticulations on growth bands on the dorsal side of the carapace, but also on growth bands on the ventral side of the carapace, which have been overlooked in the original description. The shift from wide dorsal growth bands to narrow ventral ones might indicate an onset of sexual maturity, and the transition in biological priorities from rapid juvenile growth to reproductive activities.

  5. Ontogenetic changes in feeding and food preferences of the dog conch Laevistrombus canarium Linnaeus 1758 (Mollusca: Gastropoda) from Merambong shoal, Malaysia

    Science.gov (United States)

    Husna, Wan Nurul Wan Hassan; Mazlan, Abd Ghaffar; Cob, Zaidi Che

    2017-09-01

    Laevistrombus canarium is one of the marine gastropod mollusks that have high commercial value, particularly in the aquaculture sector in Malaysia. This study was conducted to determine the feeding and food items of L. canarium at different ontogenetic stages (juveniles, sub-adults and adults) from Merambong shoals, Malaysia. Field observations on feeding activity were conducted, followed by detailed laboratory analysis on the stomach content. Five-minutes observations on randomly selected individuals were conducted at the field sampling site and their feeding activities were recorded with reference to age stage. Various shell sizes from each ontogenetic stage were randomly collected and quickly anaesthetized with ice and preserved in 10% formalin before being transported to the laboratory for stomach content analyses. Field observations showed that L. canarium mainly grazed on epiphytes occurring on seagrass (46.67%), followed by sediment surface (40%) and epiphytes occurring on macroalgae (13.33%). Stomach content analyses showed a significant difference ( P Food items found in the conch stomach include diatoms, detritus, foraminifera, seagrass and macroalgae fragments, sand particles and shell fragments. The Index of Relative Importance (%IRI) indicates three main types of food dominated the three ontogenetic stages namely diatoms, sand particles and detritus. However, no significant difference ( P >0.05) was detected between the three main food items (diatoms, sand particles and detritus) among the ontogenetic stages. Therefore, feeding activity revealed the role of the dog conch in the marine food network. While, classification of the types of food consumed by L. canarium through stomach content analysis determines the particular position of the gastropod in the food chain. Further studies are needed to provide a better insight between trophic relationships of L. canarium with marine ecosystem.

  6. Ontogenetic changes in feeding and food preferences of the dog conch Laevistrombus canarium Linnaeus 1758 (Mollusca: Gastropoda) from Merambong shoal, Malaysia

    Science.gov (United States)

    Husna, Wan Nurul Wan Hassan; Mazlan, Abd Ghaffar; Cob, Zaidi Che

    2016-10-01

    Laevistrombus canarium is one of the marine gastropod mollusks that have high commercial value, particularly in the aquaculture sector in Malaysia. This study was conducted to determine the feeding and food items of L. canarium at different ontogenetic stages (juveniles, sub-adults and adults) from Merambong shoals, Malaysia. Field observations on feeding activity were conducted, followed by detailed laboratory analysis on the stomach content. Five-minutes observations on randomly selected individuals were conducted at the field sampling site and their feeding activities were recorded with reference to age stage. Various shell sizes from each ontogenetic stage were randomly collected and quickly anaesthetized with ice and preserved in 10% formalin before being transported to the laboratory for stomach content analyses. Field observations showed that L. canarium mainly grazed on epiphytes occurring on seagrass (46.67%), followed by sediment surface (40%) and epiphytes occurring on macroalgae (13.33%). Stomach content analyses showed a significant difference (P Food items found in the conch stomach include diatoms, detritus, foraminifera, seagrass and macroalgae fragments, sand particles and shell fragments. The Index of Relative Importance (%IRI) indicates three main types of food dominated the three ontogenetic stages namely diatoms, sand particles and detritus. However, no significant difference (P >0.05) was detected between the three main food items (diatoms, sand particles and detritus) among the ontogenetic stages. Therefore, feeding activity revealed the role of the dog conch in the marine food network. While, classification of the types of food consumed by L. canarium through stomach content analysis determines the particular position of the gastropod in the food chain. Further studies are needed to provide a better insight between trophic relationships of L. canarium with marine ecosystem.

  7. The development of collective personality: the ontogenetic drivers of behavioral variation across groups

    Directory of Open Access Journals (Sweden)

    Sarah E Bengston

    2014-12-01

    Full Text Available For the past decade, the study of personality has become a topic on the frontier of behavioral ecology. However, most studies have focused on exploring inter-individual behavioral variation in solitary animals, and few account for the role that social interactions may have on the development of an individual’s personality. Moreover, a social group may exhibit collective personality: an emergent behavioral phenotype displayed at the group-level, which is not necessarily the sum or average of individual personalities within that group. The social environment, in many cases, can determine group success, which then influences the relative success of all the individuals in that group. In addition, group-level personality may itself evolve, subject to the same selection pressures as individual-level behavioral variation, when the group is a unit under selection. Therefore, we reason that understanding how collective personalities emerge and change over time will be imperative to understanding individual- and group-level behavioral evolution.Personality is considered to be fixed over an individual’s lifetime. However, behavior may shift throughout development, particularly during adolescence. Therefore, juvenile behavior should not be compared with adult behavior when assessing personality. Similarly, as conditions within a group and/or the local environment can shift, group behavior may similarly fluctuate as it matures. We discuss potential within-group factors, such as group initiation, group maturation, genetic make-up of the group, and the internal social environment, and external factors, such as well as how local environment may play a role in generating group-level personalities. There are a variety of studies that explore group development or quantify group personality, but few that integrate both processes. Therefore, we conclude by discussing potential ways to evaluate development of collective personality, and propose several focal

  8. Ontogenetic changes in the bacterial symbiont community of the tropical demosponge Amphimedon queenslandica: metamorphosis is a new beginning

    Directory of Open Access Journals (Sweden)

    Rebecca A Fieth

    2016-11-01

    Full Text Available Vertical transmission of bacterial symbionts, which is known in many species of sponge (Porifera, is expected to promote strong fidelity between the partners. Combining 16S rRNA gene amplicon sequencing and electron microscopy, we have assayed the relative abundance of vertically-inherited bacterial symbionts in several stages of the life cycle of Amphimedon queenslandica, a tropical coral reef sponge. We reveal that adult A. queenslandica house a low diversity microbiome dominated by just three proteobacterial OTUs, with a single gammaprotebacterium clearly dominant through much of the life cycle. This ontogenetic perspective has revealed that, although vertical transmission occurs very early in development, the inherited symbionts do not maintain proportional dominance of the bacterial community at every developmental stage. A reproductive bottleneck in the A. queenslandica life cycle is larval settlement, when a free-swimming pelagic larva settles out of the water column onto the benthos and completes metamorphoses into the sessile body plan within just 3 to 4 days. During this dramatic life cycle transition, an influx of environmentally-derived bacteria leads to a major reorganization of the microbiome, potentially challenging the fidelity and persistence of the vertically-inherited symbiotic relationships. However, dominance of the primary, vertically-inherited symbionts is restored in adult sponges. The mechanisms underlying ontogenetic changes in the bacterial community are unknown, including how the dominance of the primary symbionts is restored in the adult sponge – does the host or symbiont regulate this process? Using high-resolution transcriptional profiling in multiple stages of the A. queenslandica life cycle combined with this natural perturbation of the microbiome immediately following larval settlement, we are beginning to identify candidate host genes associated with animal-bacterial crosstalk. Among the sponge host genes

  9. Modulation by K+ Plus NH4+ of microsomal (Na+, K+-ATPase activity in selected ontogenetic stages of the diadromous river shrimp Macrobrachium amazonicum (Decapoda, Palaemonidae.

    Directory of Open Access Journals (Sweden)

    Francisco A Leone

    Full Text Available We investigate the synergistic stimulation by K(+ plus NH4 (+ of (Na(+, K(+-ATPase activity in microsomal preparations of whole zoea I and decapodid III, and in juvenile and adult river shrimp gills. Modulation of (Na(+, K(+-ATPase activity is ontogenetic stage-specific, and particularly distinct between juveniles and adults. Although both gill enzymes exhibit two different sites for K(+ and NH4 (+ binding, in the juvenile enzyme, these two sites are equivalent: binding by both ions results in slightly stimulated activity compared to that of a single ionic species. In the adult enzyme, the sites are not equivalent: when one ion occupies its specific binding site, (Na(+, K(+-ATPase activity is stimulated synergistically by ≈ 50% on binding of the complementary ion. Immunolocalization reveals the enzyme to be distributed predominantly throughout the intralamellar septum in the gill lamellae of juveniles and adults. Western blot analyses demonstrate a single immunoreactive band, suggesting a single (Na(+, K(+-ATPase α-subunit isoform that is distributed into different density membrane fractions, independently of ontogenetic stage. We propose a model for the modulation by K(+ and NH4 (+ of gill (Na(+, K(+-ATPase activity. These findings suggest that the gill enzyme may be regulated by NH4 (+ during ontogenetic development in M. amazonicum.

  10. Nonlinear Quantum Optics in Optomechanical Nanoscale Waveguides

    CERN Document Server

    Zoubi, Hashem

    2016-01-01

    We explore the possibility of achieving a significant nonlinear phase shift among photons propagating in nanoscale waveguides exploiting interactions among photons that are mediated by vibrational modes and induced through Stimulated Brillouin Scattering (SBS). We introduce a configuration that allows slowing down the photons by several orders of magnitude via SBS involving sound waves and two pump fields. We extract the conditions for maintaining vanishing amplitude gain or loss for slowly propagating photons while keeping the influence of thermal phonons to the minimum. The nonlinear phase among two counter-propagating photons can be used to realize a deterministic phase gate.

  11. Statistical parametric mapping of the regional distribution and ontogenetic scaling of foot pressures during walking in Asian elephants (Elephas maximus).

    Science.gov (United States)

    Panagiotopoulou, Olga; Pataky, Todd C; Hill, Zoe; Hutchinson, John R

    2012-05-01

    Foot pressure distributions during locomotion have causal links with the anatomical and structural configurations of the foot tissues and the mechanics of locomotion. Elephant feet have five toes bound in a flexible pad of fibrous tissue (digital cushion). Does this specialized foot design control peak foot pressures in such giant animals? And how does body size, such as during ontogenetic growth, influence foot pressures? We addressed these questions by studying foot pressure distributions in elephant feet and their correlation with body mass and centre of pressure trajectories, using statistical parametric mapping (SPM), a neuro-imaging technology. Our results show a positive correlation between body mass and peak pressures, with the highest pressures dominated by the distal ends of the lateral toes (digits 3, 4 and 5). We also demonstrate that pressure reduction in the elephant digital cushion is a complex interaction of its viscoelastic tissue structure and its centre of pressure trajectories, because there is a tendency to avoid rear 'heel' contact as an elephant grows. Using SPM, we present a complete map of pressure distributions in elephant feet during ontogeny by performing statistical analysis at the pixel level across the entire plantar/palmar surface. We hope that our study will build confidence in the potential clinical and scaling applications of mammalian foot pressures, given our findings in support of a link between regional peak pressures and pathogenesis in elephant feet.

  12. The effects of temperature and substrate on ontogenetic behavior of bastard halibut, Paralichthys olivaceus (T. et S.)

    Institute of Scientific and Technical Information of China (English)

    LIU Dong; XIAN Weiwei; ZHU Xinhua

    2005-01-01

    The effects of temperature and substrate on ontogenetic patterns and settlement behavior of bastard halibut, Paralichthys olivaceus (T. et S.) were studied in a series of laboratory experiments. Analysis on stage-based data revealed that except for settlement, and compared to the substrate, the ambient temperature dominantly controlled the hatch success and subsequent development of the species. The oosperm optimum survival rate of 60% during hatching occurred at 20℃, and survival rate of 29%, at 18℃ for larval rearing. The survival rate of larvae reared under variable temperature of 3.5-4.5℃ was higher than that of those reared under stable temperature. The maximum survival rate of 29%, corresponded to variation at temperature of 4℃. With increase in age newly settled juveniles preferred to lie on the fine substrate and gradually moved on the coarse substrate. GLM (general linear model) analysis showed that the combination of temperature and substrate had no significant impact on the survival of settling larvae, but definitely affected the duration of settlement.

  13. An ontogenetic analysis of locustatachykinin-like expression in the central complex of the grasshopper Schistocerca gregaria.

    Science.gov (United States)

    Boyan, G S; Williams, J L D; Herbert, Z

    2008-11-01

    We have investigated the ontogenetic basis of locustatachykinin-like expression in a group of cells located in the pars intercerebralis of the grasshopper midbrain. These cells project fibers to the protocerebral bridge and the central body via a characteristic set of fiber bundles called the w, x, y, z tracts. Lineage analyses associate the immunoreactive cells with one of four neuroblasts (termed W, X, Y, Z) in each protocerebral hemisphere of the early embryo. Locustatachykinin is a ubiquitous myotropic peptide among the insects and its expression in the pars intercerebralis begins at approximately 60-65% of embryogenesis. This coincides with the appearance of the columnar neuroarchitecture characteristic of the central body. The number of immunoreactive cells in a given lineage is initially small, increases significantly in later embryogenesis, and attains the adult situation (about 7% of a lineage) in the first larval instar after hatching. Although each neuroblast generates progeny displaying a spectrum of cell body sizes, there is a clear morphological gradient, which reflects birth order within the lineage. Locustatachykinin expressing cells are located stereotypically at or near the tip of their lineage, which an age profile reveals places them amongst the first born progeny of their respective neuroblasts. Although these neuroblasts begin to generate progeny at approximately 25-27% of embryogenesis, their daughter cells remain quiescent with respect to locustatachykinin expression for over 30% of embryogenesis.

  14. Ontogenetic changes in tracheal structure facilitate deep dives and cold water foraging in adult leatherback sea turtles.

    Science.gov (United States)

    Davenport, John; Fraher, John; Fitzgerald, Ed; McLaughlin, Patrick; Doyle, Tom; Harman, Luke; Cuffe, Tracy; Dockery, Peter

    2009-11-01

    Adult leatherbacks are large animals (300-500 kg), overlapping in size with marine pinniped and cetacean species. Unlike marine mammals, they start their aquatic life as 40-50 g hatchlings, so undergo a 10,000-fold increase in body mass during independent existence. Hatchlings are limited to the tropics and near-surface water. Adults, obligate predators on gelatinous plankton, encounter cold water at depth (<1280 m) or high latitude and are gigantotherms that maintain elevated core body temperatures in cold water. This study shows that there are great ontogenetic changes in tracheal structure related to diving and exposure to cold. Hatchling leatherbacks have a conventional reptilian tracheal structure with circular cartilaginous rings interspersed with extensive connective tissue. The adult trachea is an almost continuous ellipsoidal cartilaginous tube composed of interlocking plates, and will collapse easily in the upper part of the water column during dives, thus avoiding pressure-related structural and physiological problems. It is lined with an extensive, dense erectile vascular plexus that will warm and humidify cold inspired air and possibly retain heat on expiration. A sub-luminal lymphatic plexus is also present. Mammals and birds have independently evolved nasal turbinates to fulfil such a respiratory thermocontrol function; for them, turbinates are regarded as diagnostic of endothermy. This is the first demonstration of a turbinate equivalent in a living reptile.

  15. Venom Profiling of a Population of the Theraphosid Spider Phlogius crassipes Reveals Continuous Ontogenetic Changes from Juveniles through Adulthood.

    Science.gov (United States)

    Santana, Renan C; Perez, David; Dobson, James; Panagides, Nadya; Raven, Robert J; Nouwens, Amanda; Jones, Alun; King, Glenn F; Fry, Bryan G

    2017-03-25

    Theraphosid spiders (tarantulas) are venomous arthropods found in most tropical and subtropical regions of the world. Tarantula venoms are a complex cocktail of toxins with potential use as pharmacological tools, drugs and bioinsecticides. Although numerous toxins have been isolated from tarantula venoms, little research has been carried out on the venom of Australian tarantulas. We therefore investigated the venom profile of the Australian theraphosid spider Phlogius crassipes and examined whether there are ontogenetic changes in venom composition. Spiders were divided into four ontogenic groups according to cephalothorax length, then the venom composition of each group was examined using gel electrophoresis and mass spectrometry. We found that the venom of P. crassipes changes continuously during development and throughout adulthood. Our data highlight the need to investigate the venom of organisms over the course of their lives to uncover and understand the changing functions of venom and the full range of toxins expressed. This in turn should lead to a deeper understanding of the organism's ecology and enhance the potential for biodiscovery.

  16. Ontogenetic histological changes in the wood of mango (Mangifera indica L. cv Deshi) exposed to coal-smoke pollution

    Energy Technology Data Exchange (ETDEWEB)

    Gupta, M.C.; Iqbal, M. [Dept. of Botany, New Delhi (India)

    2005-11-01

    Trees of Mangifera indica L. cv Deshi growing at two sites, one heavily polluted due to coal-smoke emanating from a thermal power plant and the other free from such pollution, were examined across their boles (from pith to cambium) to trace ontogenetic changes induced by coal-smoke pollutants in the wood structure with special reference to vessels and fibres. Wood formation was hampered in the polluted environment, as evident from the width of annual rings, indicating an adverse influence of coal-smoke pollutants on the cambial activity. The pollutants inhibited dimensional growth of tracheal elements and promoted frequency and grouping of vessels since early growth stages. The increasing vessel number per square millimeter of wood and the decreasing dimensions of vessel elements and fibres resulted in low values for vulnerability and mesomorphic ratios in the polluted trees. The ratio of the length of fibres to that of vessel elements also decreased. With the growing age, the decline in vulnerability ratio and mesomorphic ratio was enhanced whereas that in the fibre/vessel-element length ratio was minimized.

  17. Characterization and ontogenetic expression analysis of the myosin light chains from the fast white muscle of mandarin fish Siniperca chuatsi.

    Science.gov (United States)

    Chu, W Y; Chen, J; Zhou, R X; Zhao, F L; Meng, T; Chen, D X; Nong, X X; Liu, Z; Lu, S Q; Zhang, J S

    2011-04-01

    Three full-length complementary DNA (cDNA) clones were isolated encoding the skeletal myosin light chain 1 (MLC1; 1237 bp), myosin light chain 2 (MLC2; 1206 bp) and myosin light chain 3 (MLC3; 1079 bp) from the fast white muscle cDNA library of mandarin fish Siniperca chuatsi. The sequence analysis indicated that MLC1 and MLC3 were not produced from differentially spliced messenger RNAs (mRNA) as reported in birds and rodents but were encoded by different genes. The MLC2 encodes 170 amino acids, which include four EF-hand (helix-loop-helix) structures. The primary structures of the Ca(2+)-binding domain were well conserved among the MLC2s of seven other fish species. The ontogenetic expression analysis by real-time PCR showed that the three light-chain mRNAs were first detected in the gastrula stage, and their expression increased from the tail bud stage to the larval stage. All three MLC mRNAs showed longitudinal expression variation in the fast white muscle of S. chuatsi, especially MLC1 which was highly expressed at the posterior area. Taken together, the study provides a better understanding about the MLC gene structure and their expression pattern in muscle development of S. chuatsi.

  18. Ontogenetic changes in the craniomandibular skeleton of the abelisaurid dinosaur Majungasaurus crenatissimus from the Late Cretaceous of Madagascar

    Directory of Open Access Journals (Sweden)

    Nirina O. Ratsimbaholison

    2016-06-01

    Full Text Available Abelisaurid theropods were one of the most diverse groups of predatory dinosaurs in Gondwana during the Cretaceous. The group is characterized by a tall, wide skull and robust cervical region. This morphology is thought to have facilitated specialized feeding behaviors such as prolonged contact with prey. The Late Cretaceous abelisaurid Majungasaurus crenatissimus typifies this abelisaurid cranial morphotype. Recent fossil discoveries of this species include a partial growth series that allows for the first time an investigation of ontogenetic variation in cranial morphology in a representative abelisaurid. Herein we examine growth trajectories in the shape of individual cranial bones and articulated skulls of Majungasaurus using geometric morphometrics. Several major changes in skull shape were observed through ontogeny, including an increase in the height of the jugal, postorbital, and quadratojugal, an increase in the extent of the contacts between bones, and a decrease in the circumference of the orbit. The skull transitions from relatively short in the smallest individual to tall and robust in large adults, as is seen in other theropods. Such morphological change during ontogeny would likely have resulted in different biomechanical properties and feeding behaviors between small and large individuals. These findings provide a post-hatching developmental framework for understanding the evolution of the distinctive tall skull morphology seen in abelisaurids and other large-sized theropod dinosaurs.

  19. On the Lamb shift

    Energy Technology Data Exchange (ETDEWEB)

    Villarroel, D. [Av. Tobalaba 3696, Puente Alto, Santiago, Metropolitana (Chile)

    2008-02-15

    The Lamb shift is calculated, in an approximate way, considering the hydrogen atom as an isolated physical system; the quantized radiation field does not play any role in the present approach. Our formalism is based on the generalization of the Dirac wave equation that incorporates the effects of the electron self-fields directly into it. Both the physical picture as well as the mathematical formalism have their roots in the classical theory of the electron. (author)

  20. Advances in nonlinear optics

    CERN Document Server

    Chen, Xianfeng; Zeng, Heping; Guo, Qi; She, Weilong

    2015-01-01

    This book presents an overview of the state of the art of nonlinear optics from weak light nonlinear optics, ultrafast nonlinear optics to electro-optical theory and applications. Topics range from the fundamental studies of the interaction between matter and radiation to the development of devices, components, and systems of tremendous commercial interest for widespread applications in optical telecommunications, medicine, and biotechnology.

  1. Distributed nonlinear optical response

    DEFF Research Database (Denmark)

    Nikolov, Nikola Ivanov

    2005-01-01

    The purpose of the research presented here is to investigate basic physical properties in nonlinear optical materials with delayed or nonlocal nonlinearity. Soliton propagation, spectral broadening and the influence of the nonlocality or delay of the nonlinearity are the main focusses in the work...

  2. Noncommutative Nonlinear Supersymmetry

    CERN Document Server

    Nishino, H; Nishino, Hitoshi; Rajpoot, Subhash

    2002-01-01

    We present noncommutative nonlinear supersymmetric theories. The first example is a non-polynomial Akulov-Volkov-type lagrangian with noncommutative nonlinear global supersymmetry in arbitrary space-time dimensions. The second example is the generalization of this lagrangian to Dirac-Born-Infeld lagrangian with nonlinear supersymmetry realized in dimensions D=2,3,4 and 6 (mod 8).

  3. Fiber Nonlinearities: A Tutorial

    Institute of Scientific and Technical Information of China (English)

    Govind P. Agrawal

    2003-01-01

    Fiber nonlinearities have long been regarded as being mostly harmful for fiber-optic communication systems. Over the last few years, however, the nonlinear effects are increasingly being used for practical telecommunications applications,the Raman amplification being only one of the recent examples. In this tutorial I review the vario us nonlinear effects occurring in optical fibers from both standpoints..

  4. Fiber Nonlinearities: A Tutorial

    Institute of Scientific and Technical Information of China (English)

    Govind; P.; Agrawal

    2003-01-01

    Fiber nonlinearities have long been regarded as being mostly harmful for fiber-optic communication systems. Over the last few years, however, the nonlinear effects are increasingly being used for practical telecommunications applications, the Raman amplification being only one of the recent examples. In this tutorial I review the various nonlinear effects occurring in optical fibers from both standpoints..

  5. PBH tests for nonlinear systems

    NARCIS (Netherlands)

    Kawano, Yu; Ohtsuka, Toshiyuki

    2017-01-01

    Recently, concepts of nonlinear eigenvalues and eigenvectors are introduced. In this paper, we establish connections between the nonlinear eigenvalues and nonlinear accessibility/observability. In particular, we provide a generalization of Popov- Belevitch-Hautus (PBH) test to nonlinear accessibilit

  6. The shifting beverage landscape.

    Science.gov (United States)

    Storey, Maureen

    2010-04-26

    STOREY, M.L. The shifting beverage landscape. PHYSIOL BEHAV, 2010. - Simultaneous lifestyle changes have occurred in the last few decades, creating an imbalance in energy intake and energy expenditure that has led to overweight and obesity. Trends in the food supply show that total daily calories available per capita increased 28% since 1970. Total energy intake among men and women has also increased dramatically since that time. Some have suggested that intake of beverages has had a disproportional impact on obesity. Data collected by the Beverage Marketing Corporation between 1988-2008 demonstrate that, in reality, fewer calories per ounce are being produced by the beverage industry. Moreover, data from the National Cancer Institute show that soft drink intake represents 5.5% of daily calories. Data from NHANES 1999-2003 vs. 2003-06 may demonstrate a shift in beverage consumption for age/gender groups, ages 6 to>60years. The beverages provided in schools have significantly changed since 2006 when the beverage industry implemented School Beverage Guidelines. This voluntary action has removed full-calorie soft drinks from participating schools across the country. This shift to lower-calorie and smaller-portion beverages in school has led to a significant decrease in total beverage calories in schools. These data support the concept that to prevent and treat obesity, public health efforts should focus on energy balance and that a narrow focus on sweetened beverages is unlikely to have any meaningful impact on this complex problem. Copyright 2010 Elsevier Inc. All rights reserved.

  7. Catastrophic shifts in ecosystems

    Science.gov (United States)

    Scheffer, Marten; Carpenter, Steve; Foley, Jonathan A.; Folke, Carl; Walker, Brian

    2001-10-01

    All ecosystems are exposed to gradual changes in climate, nutrient loading, habitat fragmentation or biotic exploitation. Nature is usually assumed to respond to gradual change in a smooth way. However, studies on lakes, coral reefs, oceans, forests and arid lands have shown that smooth change can be interrupted by sudden drastic switches to a contrasting state. Although diverse events can trigger such shifts, recent studies show that a loss of resilience usually paves the way for a switch to an alternative state. This suggests that strategies for sustainable management of such ecosystems should focus on maintaining resilience.

  8. Nonlinear dynamics and complexity

    CERN Document Server

    Luo, Albert; Fu, Xilin

    2014-01-01

    This important collection presents recent advances in nonlinear dynamics including analytical solutions, chaos in Hamiltonian systems, time-delay, uncertainty, and bio-network dynamics. Nonlinear Dynamics and Complexity equips readers to appreciate this increasingly main-stream approach to understanding complex phenomena in nonlinear systems as they are examined in a broad array of disciplines. The book facilitates a better understanding of the mechanisms and phenomena in nonlinear dynamics and develops the corresponding mathematical theory to apply nonlinear design to practical engineering.

  9. Construction of a single/multiple wavelength RZ optical pulse source at 40 GHz by use of wavelength conversion in a high-nonlinearity DSF-NOLM

    DEFF Research Database (Denmark)

    Yu, Jianjun; Yujun, Qian; Jeppesen, Palle;

    2001-01-01

    A single or multiple wavelength RZ optical pulse source at 40 GHz is successfully obtained by using wavelength conversion in a nonlinear optical loop mirror consisting of high nonlinearity-dispersion shifted fiber.......A single or multiple wavelength RZ optical pulse source at 40 GHz is successfully obtained by using wavelength conversion in a nonlinear optical loop mirror consisting of high nonlinearity-dispersion shifted fiber....

  10. Resolution enhancement in nonlinear photoacoustic imaging

    Energy Technology Data Exchange (ETDEWEB)

    Goy, Alexandre S.; Fleischer, Jason W. [Department of Electrical Engineering, Princeton University, Olden St., Princeton, New Jersey 08544 (United States)

    2015-11-23

    Nonlinear processes can be exploited to gain access to more information than is possible in the linear regime. Nonlinearity modifies the spectra of the excitation signals through harmonic generation, frequency mixing, and spectral shifting, so that features originally outside the detector range can be detected. Here, we present an experimental study of resolution enhancement for photoacoustic imaging of thin metal layers immersed in water. In this case, there is a threshold in the excitation below which no acoustic signal is detected. Above threshold, the nonlinearity reduces the width of the active area of the excitation beam, resulting in a narrower absorption region and thus improved spatial resolution. This gain is limited only by noise, as the active area of the excitation can be arbitrarily reduced when the fluence becomes closer to the threshold. Here, we demonstrate a two-fold improvement in resolution and quantify the image quality as the excitation fluence goes through threshold.

  11. Nonlinear estimation and control of automotive drivetrains

    CERN Document Server

    Chen, Hong

    2014-01-01

    Nonlinear Estimation and Control of Automotive Drivetrains discusses the control problems involved in automotive drivetrains, particularly in hydraulic Automatic Transmission (AT), Dual Clutch Transmission (DCT) and Automated Manual Transmission (AMT). Challenging estimation and control problems, such as driveline torque estimation and gear shift control, are addressed by applying the latest nonlinear control theories, including constructive nonlinear control (Backstepping, Input-to-State Stable) and Model Predictive Control (MPC). The estimation and control performance is improved while the calibration effort is reduced significantly. The book presents many detailed examples of design processes and thus enables the readers to understand how to successfully combine purely theoretical methodologies with actual applications in vehicles. The book is intended for researchers, PhD students, control engineers and automotive engineers. Hong Chen is a professor at the State Key Laboratory of Automotive Simulation and...

  12. Strongly nonlinear steepening of long interfacial waves

    Directory of Open Access Journals (Sweden)

    N. Zahibo

    2007-06-01

    Full Text Available The transformation of nonlinear long internal waves in a two-layer fluid is studied in the Boussinesq and rigid-lid approximation. Explicit analytic formulation of the evolution equation in terms of the Riemann invariants allows us to obtain analytical results characterizing strongly nonlinear wave steepening, including the spectral evolution. Effects manifesting the action of high nonlinear corrections of the model are highlighted. It is shown, in particular, that the breaking points on the wave profile may shift from the zero-crossing level. The wave steepening happens in a different way if the density jump is placed near the middle of the water bulk: then the wave deformation is almost symmetrical and two phases appear where the wave breaks.

  13. Repetition and Translation Shifts

    Directory of Open Access Journals (Sweden)

    Simon Zupan

    2006-06-01

    Full Text Available Repetition manifests itself in different ways and at different levels of the text. The first basic type of repetition involves complete recurrences; in which a particular textual feature repeats in its entirety. The second type involves partial recurrences; in which the second repetition of the same textual feature includes certain modifications to the first occurrence. In the article; repetitive patterns in Edgar Allan Poe’s short story “The Fall of the House of Usher” and its Slovene translation; “Konec Usherjeve hiše”; are compared. The author examines different kinds of repetitive patterns. Repetitions are compared at both the micro- and macrostructural levels. As detailed analyses have shown; considerable microstructural translation shifts occur in certain types of repetitive patterns. Since these are not only occasional; sporadic phenomena; but are of a relatively high frequency; they reduce the translated text’s potential for achieving some of the gothic effects. The macrostructural textual property particularly affected by these shifts is the narrator’s experience as described by the narrative; which suffers a reduction in intensity.

  14. Ontogenetic behavior and dispersal of Sacramento River white sturgeon, Acipenser transmontanus, with a note on body color

    Science.gov (United States)

    Kynard, B.; Parker, E.

    2005-01-01

    We studied Sacramento River white sturgeon, Acipenser transmontanus, in the laboratory to develop a conceptual model of ontogenetic behavior and provide insight into probable behavior of wild sturgeon. After hatching, free embryos initiated a low intensity, brief downstream dispersal during which fish swam near the bottom and were photonegative. The weak, short dispersal style and behavior of white sturgeon free embryos contrasts greatly with the intense, long dispersal style and behavior (photopositive and swimming far above the bottom) of dispersing free embryos of other sturgeon species. If spawned eggs are concentrated within a few kilometers downstream of a spawning site, the adaptive significance of the free embryo dispersal is likely to move fish away from the egg deposition site to avoid predation and reduce fish density prior to feeding. Larvae foraged on the open bottom, swam white sturgeon populations may be a mis-match between the innate fish dispersal and post-dispersal rearing habitat, which is now highly altered by damming and reservoirs. Sacramento River white sturgeon has a two-step downstream dispersal by the free embryo and juvenile life intervals. Diel activity of all life intervals peaked at night, whether fish were dispersing or foraging. Nocturnal behavior is likely a response to predation, which occurs during both activities. An intense black-tail body color was present on foraging larvae, but was weak or absent on the two life intervals that disperse. Black-tail color may be an adaptation for avoiding predation, signaling among aggregated larvae, or both, but not for dispersal. ?? Springer 2005.

  15. An in situ, individual-based approach to quantify connectivity of marine fish: ontogenetic movements and residency of lingcod.

    Science.gov (United States)

    Bishop, Mary Anne; Reynolds, Brad F; Powers, Sean P

    2010-12-13

    As modern fishery assessments change in an effort to be more accurate and encompass the range of potential ecosystem interactions, critical information on the ecology of species including life history, intra and inter-specific competitive interactions and habitat requirements must be added to the standard fishery-dependent and independent data sets. One species whose movements and habitat associations greatly affects exploitation patterns is lingcod, Ophiodon elongatus, which support an economically important fishery along the coastal waters of the Pacific Coast of North America. High site fidelity and limited movements within nearshore areas are hypothesized to have resulted in high catchability, a major factor that has contributed to overfished stocks. Thus, assessing the level of movement and connectivity among lingcod subpopulations inhabiting nearshore habitats is a prerequisite to determining the condition of lingcod stocks. We used the Pacific Ocean Shelf Tracking (POST) Project acoustic receiver array in Alaska's Prince William Sound to monitor movements and residency of 21 acoustic-tagged lingcod for up to 16 months. Eight of sixteen lingcod (50%) initially aged at 2.5- to 3.5- years-old dispersed from their tag site. Dispersal was highly seasonal, occurring in two, five-week periods from mid-December through January and from mid-April through May. Dispersal in winter may be related to sexually immature lingcod or newly-mature male lingcod being displaced by territorial males. Spring dispersal may be indicative of the onset of migratory behavior where lingcod move out into Prince William Sound and possibly the offshore waters of the Gulf of Alaska. Our results reveal a pattern of ontogenetic dispersal as lingcod approach 4-years-old and exceed 50 cm total length. The large proportion of tagged fish migrating out of Port Gravina, their tagging site, reflects a high level of connectivity among Prince William Sound subpopulations. Our results also support the

  16. Spatial, temporal and ontogenetic variation in diet of anchovy ( Engraulis encrasicolus) on the Algerian coast (SW Mediterranean)

    Science.gov (United States)

    Bacha, M.; Amara, R.

    2009-11-01

    The diet of anchovy Engraulis encrasicolus was studied in three regions (Béjaia, Bénisaf and Ghazaouet) along the Algerian coast. Ontogenetic, spatial and seasonal variations in anchovy diet were investigated using multivariate analyses and analysed in relation with sea surface temperature and chlorophyll- a. 46 prey taxa of varying size between 0.57 mm ( Euterpina acutifrons) and 6.8 mm (fish larvae) were recorded. Whatever the season, the region or the fish size, anchovy is exclusively zooplanktivorous and copepods were the most present prey, constituting 87% by number of the prey taken and found in 98% of the anchovy stomachs examined. However, their occurrence and number varied according to the different areas, seasons and fish size. During its first year of life, anchovy feeds almost exclusively on copepods (mainly small and medium size prey). As anchovy grows, copepods are gradually substituted by large crustaceans such as decapods and amphipods. Hierarchical cluster analysis, analysis of similarities (ANOSIM) and similarities percentage (SIMPER) indicated a distinct diet of anchovy of the bay of Bejaia from those of the bays of Bénisaf and Ghazaouet probably due to differences in hydrologic conditions. Diet differences also occurred between seasons. Summer and spring have distinct prey assemblages each and showed low diet similarities with the two other seasons. More prey species were found in the diet during winter (36) and autumn (30) and the vacuity index was lower in winter. Temporal variability in satellite-derived chlorophyll- a matched the seasonal variability in the diversity of the anchovy prey and feeding intensity as reflected by the vacuity index, suggesting further investigation of the potential use of satellite-derived chlorophyll- a data as a proxy for anchovy feeding intensity.

  17. Nonlinear Elliptic Differential Equations with Multivalued Nonlinearities

    Indian Academy of Sciences (India)

    Antonella Fiacca; Nikolaos Matzakos; Nikolaos S Papageorgiou; Raffaella Servadei

    2001-11-01

    In this paper we study nonlinear elliptic boundary value problems with monotone and nonmonotone multivalued nonlinearities. First we consider the case of monotone nonlinearities. In the first result we assume that the multivalued nonlinearity is defined on all $\\mathbb{R}$. Assuming the existence of an upper and of a lower solution, we prove the existence of a solution between them. Also for a special version of the problem, we prove the existence of extremal solutions in the order interval formed by the upper and lower solutions. Then we drop the requirement that the monotone nonlinearity is defined on all of $\\mathbb{R}$. This case is important because it covers variational inequalities. Using the theory of operators of monotone type we show that the problem has a solution. Finally in the last part we consider an eigenvalue problem with a nonmonotone multivalued nonlinearity. Using the critical point theory for nonsmooth locally Lipschitz functionals we prove the existence of at least two nontrivial solutions (multiplicity theorem).

  18. VARIANCE OF NONLINEAR PHASE NOISE IN FIBER-OPTIC SYSTEM

    OpenAIRE

    RANJU KANWAR; SAMEKSHA BHASKAR

    2013-01-01

    In communication system, the noise process must be known, in order to compute the system performance. The nonlinear effects act as strong perturbation in long- haul system. This perturbation effects the signal, when interact with amplitude noise, and results in random motion of the phase of the signal. Based on the perturbation theory, the variance of nonlinear phase noise contaminated by both self- and cross-phase modulation, is derived analytically for phase-shift- keying system. Through th...

  19. First order tune shift calculations for transverse betatron dynamics

    Energy Technology Data Exchange (ETDEWEB)

    Garavaglia, T.

    1991-09-01

    An effective Hamiltonian, with non-linear magnetic multipole terms and momentum dispersion contributions, is used to obtain the first order tune-shift results for transverse betatron motion for protons in the Superconducting Super Collider (SSC). This Hamiltonian is represented in terms of action angle variables, and analytical results are obtained using symbolic algebra methods. Mathematical derivations of the transverse multipole expansion and of the transverse betatron equations, using an invariant action and curvilinear coordinates, are given in the appendices. Numerical and graphical tune-space results are given that illustrate the dependence of tune-shifts on injection amplitude and momentum spread. 10 refs., 7 figs.

  20. Linear approximation for measurement errors in phase shifting interferometry

    Science.gov (United States)

    van Wingerden, Johannes; Frankena, Hans J.; Smorenburg, Cornelis

    1991-07-01

    This paper shows how measurement errors in phase shifting interferometry (PSI) can be described to a high degree of accuracy in a linear approximation. System error sources considered here are light source instability, imperfect reference phase shifting, mechanical vibrations, nonlinearity of the detector, and quantization of the detector signal. The measurement inaccuracies resulting from these errors are calculated in linear approximation for several formulas commonly used for PSI. The results are presented in tables for easy calculation of the measurement error magnitudes for known system errors. In addition, this paper discusses the measurement error reduction which can be achieved by choosing an appropriate phase calculation formula.

  1. ALL OPTICAL 3-BIT SERIAL INPUT SHIFT REGISTER DESIGN

    Directory of Open Access Journals (Sweden)

    VIKRANT K SRIVASTAVA,

    2010-08-01

    Full Text Available In this Paper, we present all-optical shift Register logic with complete Boolean functionality as a representative circuit for modeling and optimization of monolithically integrated components. Proposed optical logic unit is based on nonlinear effects in semiconductor optical amplifiers (SOA. We show a strategy of optical pulse propagation in SOA based on coupled nonlinear equations describing XGM and FWM effects. These equations are first solved togenerate the pump, probe and conjugate pulses in a SOA. The pulse behavior are analyzed and applied to realize behavior of all-optical NAND gate. Next, the logic is used to implement All-Optical Flip-Flop logic, and its function is verified with the help of truth table. Finally with the help of three Flip Flop a 3-bit shift register is proposed. The full design is simple, compact, economical, thermally stable and integration capable.

  2. [Correlation between the output and composition of essential oil and the level of salicylic acid in mint plants at different ontogenetic stages].

    Science.gov (United States)

    Shelepova, O V; Kondrat'eva, V V; Voronkova, T V; Olekhnovich, L S

    2013-01-01

    Dynamic changes in the content of acetylsalicylic acid and the output and qualitative composition of essential oil have been studied in mint plants (Mentha spicata L. and cultivar Medichka) during their ontogenesis with allowance for changes in weather conditions. Ontogenetic changes in the level of acetylsalicylic acid in leaf tissues are found to be similar in both cv. Medichka and M. spicata. In the case of cv. Medichka, this change is connected with the dynamics of the production and the qualitative composition of essential oil; in the case of M. spicata, this connection is less expressed. The role of acetylsalicylic acid and essential oil in plant adaptation to the environment is discussed.

  3. A new species of Hyla (Anura: Hylidae) from the Sierra Mixes, Oaxaca, Mexico, with comments on ontogenetic variation in the tadpoles

    Science.gov (United States)

    Ustach, P.C.; Mendelson, J.R.; McDiarmid, R.W.; Campbell, J.A.

    2000-01-01

    We describe a new species of Hyla that differs from the similar-loohng species H. pentheter by reaching a larger size, ha\\iing a smaller tympanum, more webbing on the feet, more extensive nuptial excrescences, and a different color pattern on the flanks. \\Ve tentatively place this new species in the phenetic assemblage commonly referred to as the H. bistincta group. \\Ve describe and illlistrate the tadpole and discuss ontogenetic variation among tadpoles, with reference to existing information on tadpoles of other species from the H. bistincta group.

  4. Nonlinear Dirac Equations

    Directory of Open Access Journals (Sweden)

    Wei Khim Ng

    2009-02-01

    Full Text Available We construct nonlinear extensions of Dirac's relativistic electron equation that preserve its other desirable properties such as locality, separability, conservation of probability and Poincaré invariance. We determine the constraints that the nonlinear term must obey and classify the resultant non-polynomial nonlinearities in a double expansion in the degree of nonlinearity and number of derivatives. We give explicit examples of such nonlinear equations, studying their discrete symmetries and other properties. Motivated by some previously suggested applications we then consider nonlinear terms that simultaneously violate Lorentz covariance and again study various explicit examples. We contrast our equations and construction procedure with others in the literature and also show that our equations are not gauge equivalent to the linear Dirac equation. Finally we outline various physical applications for these equations.

  5. Nonlinear, tunable and active metamaterials

    CERN Document Server

    Lapine, Mikhail; Kivshar, Yuri

    2015-01-01

    Metamaterials, artificial electromagnetic media achieved by structuring on the subwave-length-scale were initially suggested for the negative index and superlensing. They became a paradigm for engineering electromagnetic space and controlling propagation of waves. The research agenda is now shifting on achieving tuneable, switchable, nonlinear and sensing functionalities. The time has come to talk about the emerging research field of metadevices employing active and tunable metamaterials with unique functionalities achieved by structuring of functional matter on the subwave-length scale. This book presents the first systematic and comprehensive summary of the reviews written by the pioneers and top-class experts in the field of metamaterials. It addresses many grand challenges of the cutting edge research for creating smaller and more efficient photonic structures and devices.

  6. Nonlinear Cross Gramians

    Science.gov (United States)

    Ionescu, Tudor C.; Scherpen, Jacquelien M. A.

    We study the notion of cross Gramians for nonlinear gradient systems, using the characterization in terms of prolongation and gradient extension associated to the system. The cross Gramian is given for the variational system associated to the original nonlinear gradient system. We obtain linearization results that correspond to the notion of a cross Gramian for symmetric linear systems. Furthermore, first steps towards relations with the singular value functions of the nonlinear Hankel operator are studied and yield promising results.

  7. Nonlinear functional analysis

    Directory of Open Access Journals (Sweden)

    W. L. Fouché

    1983-03-01

    Full Text Available In this article we discuss some aspects of nonlinear functional analysis. It included reviews of Banach’s contraction theorem, Schauder’s fixed point theorem, globalising techniques and applications of homotopy theory to nonlinear functional analysis. The author emphasises that fundamentally new ideas are required in order to achieve a better understanding of phenomena which contain both nonlinear and definite infinite dimensional features.

  8. Nonlinear Electrodynamics and QED

    OpenAIRE

    2003-01-01

    The limits of linear electrodynamics are reviewed, and possible directions of nonlinear extension are explored. The central theme is that the qualitative character of the empirical successes of quantum electrodynamics must be used as a guide for understanding the nature of the nonlinearity of electrodynamics at the subatomic level. Some established theories of nonlinear electrodynamics, namely, those of Mie, Born, and Infeld are presented in the language of the modern geometrical and topologi...

  9. Generation and extraction of second pitch-shift from cochlear biophysics

    CERN Document Server

    Gomez, Florian

    2013-01-01

    It has been long known that the perceived pitch of a complex harmonic sound changes if the partials of the sound are shifted in frequency by a fixed amount. Rules that this pitch-shift can be expected to follow can be derived from simple nonlinear modeling (first pitch-shift), but clear deviations were observed in corresponding psychoacoustic experiments (second pitch-shift effects). This raises the question of whether these deviations are due to the biophysics of the nonlinear hearing sensor, the cochlea, or are an artifact generated higher up in the auditory pathway. In this article, we explicitly confirm that the second pitch-shift originates in the cochlea, and that the key factors responsible for the phenomenon are combination-tone generation, low-pass filtering and feed-forward coupling in the cochlea. We find, in particular, that the scaling laws of Hopf cochlea combination tones fully explain the psychoacoustical pitch data of G.F. Smoorenburg (1970).

  10. Identifying nonlinear biomechanical models by multicriteria analysis

    Science.gov (United States)

    Srdjevic, Zorica; Cveticanin, Livija

    2012-02-01

    In this study, the methodology developed by Srdjevic and Cveticanin (International Journal of Industrial Ergonomics 34 (2004) 307-318) for the nonbiased (objective) parameter identification of the linear biomechanical model exposed to vertical vibrations is extended to the identification of n-degree of freedom (DOF) nonlinear biomechanical models. The dynamic performance of the n-DOF nonlinear model is described in terms of response functions in the frequency domain, such as the driving-point mechanical impedance and seat-to-head transmissibility function. For randomly generated parameters of the model, nonlinear equations of motion are solved using the Runge-Kutta method. The appropriate data transformation from the time-to-frequency domain is performed by a discrete Fourier transformation. Squared deviations of the response functions from the target values are used as the model performance evaluation criteria, thus shifting the problem into the multicriteria framework. The objective weights of criteria are obtained by applying the Shannon entropy concept. The suggested methodology is programmed in Pascal and tested on a 4-DOF nonlinear lumped parameter biomechanical model. The identification process over the 2000 generated sets of parameters lasts less than 20 s. The model response obtained with the imbedded identified parameters correlates well with the target values, therefore, justifying the use of the underlying concept and the mathematical instruments and numerical tools applied. It should be noted that the identified nonlinear model has an improved accuracy of the biomechanical response compared to the accuracy of a linear model.

  11. Geometrically nonlinear creeping mathematic models of shells with variable thickness

    Directory of Open Access Journals (Sweden)

    V.M. Zhgoutov

    2012-08-01

    Full Text Available Calculations of strength, stability and vibration of shell structures play an important role in the design of modern devices machines and structures. However, the behavior of thin-walled structures of variable thickness during which geometric nonlinearity, lateral shifts, viscoelasticity (creep of the material, the variability of the profile take place and thermal deformation starts up is not studied enough.In this paper the mathematical deformation models of variable thickness shells (smoothly variable and ribbed shells, experiencing either mechanical load or permanent temperature field and taking into account the geometrical nonlinearity, creeping and transverse shear, were developed. The refined geometrical proportions for geometrically nonlinear and steadiness problems are given.

  12. [Relationship between shift work and overweight/obesity in male steel workers].

    Science.gov (United States)

    Xiao, M Y; Wang, Z Y; Fan, H M; Che, C L; Lu, Y; Cong, L X; Gao, X J; Liu, Y J; Yuan, J X; Li, X M; Hu, B; Chen, Y P

    2016-11-10

    Objective: To investigate the relationship between shift work and overweight/obesity in male steel workers. Methods: A questionnaire survey was conducted among the male steel workers selected during health examination in Tangshan Steel Company from March 2015 to March 2016. The relationship between shift work and overweight/obesity in the male steel workers were analyzed by using logistic regression model and restricted cubic splinemodel. Results: A total of 7 262 male steel workers were surveyed, the overall prevalence of overweight/obesitywas 64.5% (4 686/7 262), the overweight rate was 34.3% and the obesity rate was 30.2%, respectively. After adjusting for age, educational level and average family income level per month by multivariable logistic regression analysis, shift work was associated with overweight/obesity and obesity in the male steel workers. The OR was 1.19(95% CI: 1.05-1.35) and 1.15(95% CI: 1.00-1.32). Restricted cubic spline model analysis showed that the relationship between shift work years and overweight/obesity in the male steel workers was a nonlinear dose response one (nonlinear test χ(2)=7.43, Pobesity in the male steel workers was a nonlinear dose response one (nonlinear test χ(2)=10.48, Poverweight and obesity in the male steel workers, and shift work years and overweight/obesity had a nonlinear relationship.

  13. Nonlinear Physics of Plasmas

    CERN Document Server

    Kono, Mitsuo

    2010-01-01

    A nonlinearity is one of the most important notions in modern physics. A plasma is rich in nonlinearities and provides a variety of behaviors inherent to instabilities, coherent wave structures and turbulence. The book covers the basic concepts and mathematical methods, necessary to comprehend nonlinear problems widely encountered in contemporary plasmas, but also in other fields of physics and current research on self-organized structures and magnetized plasma turbulence. The analyses make use of strongly nonlinear models solved by analytical techniques backed by extensive simulations and available experiments. The text is written for senior undergraduates, graduate students, lecturers and researchers in laboratory, space and fusion plasmas.

  14. Organic nonlinear optical materials

    Science.gov (United States)

    Umegaki, S.

    1987-01-01

    Recently, it became clear that organic compounds with delocalized pi electrons show a great nonlinear optical response. Especially, secondary nonlinear optical constants of more than 2 digits were often seen in the molecular level compared to the existing inorganic crystals such as LiNbO3. The crystallization was continuously tried. Organic nonlinear optical crystals have a new future as materials for use in the applied physics such as photomodulation, optical frequency transformation, opto-bistabilization, and phase conjugation optics. Organic nonlinear optical materials, e.g., urea, O2NC6H4NH2, I, II, are reviewed with 50 references.

  15. Nonlinearity-reduced interferometer

    Science.gov (United States)

    Wu, Chien-ming

    2007-12-01

    Periodic nonlinearity is a systematic error limiting the accuracy of displacement measurements at the nanometer level. It results from many causes such as the frequency mixing, polarization mixing, polarization-frequency mixing, and the ghost reflections. An interferometer having accuracy in displacement measurement of less than one-nanometer is necessary in nanometrology. To meet the requirement, the periodic nonlinearity should be less than deep sub-nanometer. In this paper, a nonlinearity-reduced interferometry has been proposed. Both the linear- and straightness-interferometer were tested. The developed interferometer demonstrated of a residual nonlinearity less than 25 pm.

  16. Optical limiter based on two-dimensional nonlinear photonic crystals

    Science.gov (United States)

    Belabbas, Amirouche; Lazoul, Mohamed

    2016-04-01

    The aim behind this work is to investigate the capabilities of nonlinear photonic crystals to achieve ultra-fast optical limiters based on third order nonlinear effects. The purpose is to combine the actions of nonlinear effects with the properties of photonic crystals in order to activate the photonic band according to the magnitude of the nonlinear effects, themselves a function of incident laser power. We are interested in designing an optical limiter based nonlinear photonic crystal operating around 1064 nm and its second harmonic at 532 nm. Indeed, a very powerful solid-state laser that can blind or destroy optical sensors and is widely available and easy to handle. In this work, we perform design and optimization by numerical simulations to determine the better structure for the nonlinear photonic crystal to achieve compact and efficient integrated optical limiter. The approach consists to analyze the band structures in Kerr-nonlinear two-dimensional photonic crystals as a function of the optical intensity. We confirm that these bands are dynamically red-shifted with regard to the bands observed in linear photonic crystals or in the case of weak nonlinear effects. The implemented approach will help to understand such phenomena as intensitydriven optical limiting with Kerr-nonlinear photonic crystals.

  17. Comparative venomics of the Prairie Rattlesnake (Crotalus viridis viridis) from Colorado: Identification of a novel pattern of ontogenetic changes in venom composition and assessment of the immunoreactivity of the commercial antivenom CroFab®.

    Science.gov (United States)

    Saviola, Anthony J; Pla, Davinia; Sanz, Libia; Castoe, Todd A; Calvete, Juan J; Mackessy, Stephen P

    2015-05-21

    Here we describe and compare the venomic and antivenomic characteristics of both neonate and adult Prairie Rattlesnake (Crotalus viridis viridis) venoms. Although both neonate and adult venoms contain unique components, similarities among protein family content were seen. Both neonate and adult venoms consisted of myotoxin, bradykinin-potentiating peptide (BPP), phospholipase A2 (PLA2), Zn(2+)-dependent metalloproteinase (SVMP), serine proteinase, L-amino acid oxidase (LAAO), cysteine-rich secretory protein (CRISP) and disintegrin families. Quantitative differences, however, were observed, with venoms of adults containing significantly higher concentrations of the non-enzymatic toxic compounds and venoms of neonates containing higher concentrations of pre-digestive enzymatic proteins such as SVMPs. To assess the relevance of this venom variation in the context of snakebite and snakebite treatment, we tested the efficacy of the common antivenom CroFab® for recognition of both adult and neonate venoms in vitro. This comparison revealed that many of the major protein families (SVMPs, CRISP, PLA2, serine proteases, and LAAO) in both neonate and adult venoms were immunodepleted by the antivenom, whereas myotoxins, one of the major toxic components of C. v. viridis venom, in addition to many of the small peptides, were not efficiently depleted by CroFab®. These results therefore provide a comprehensive catalog of the venom compounds present in C. v. viridis venom and new molecular insight into the potential efficacy of CroFab® against human envenomations by one of the most widely distributed rattlesnake species in North America. Comparative proteomic analysis of venoms of neonate and adult Prairie Rattlesnake (Crotalus viridis viridis) from a discrete population in Colorado revealed a novel pattern of ontogenetic shifts in toxin composition for viperid snakes. The observed stage-dependent decrease of the relative content of disintegrins, catalytically active D49-PLA2s

  18. Measurement-Induced Strong Kerr Nonlinearity for Weak Quantum States of Light

    Science.gov (United States)

    Costanzo, Luca S.; Coelho, Antonio S.; Biagi, Nicola; Fiurášek, Jaromír; Bellini, Marco; Zavatta, Alessandro

    2017-07-01

    Strong nonlinearity at the single photon level represents a crucial enabling tool for optical quantum technologies. Here we report on experimental implementation of a strong Kerr nonlinearity by measurement-induced quantum operations on weak quantum states of light. Our scheme coherently combines two sequences of single photon addition and subtraction to induce a nonlinear phase shift at the single photon level. We probe the induced nonlinearity with weak coherent states and characterize the output non-Gaussian states with quantum state tomography. The strong nonlinearity is clearly witnessed as a change of sign of specific off-diagonal density matrix elements in the Fock basis.

  19. Simple nonlinear interferometer-based all-optical thresholder and its applications for optical CDMA.

    Science.gov (United States)

    Kravtsov, Konstantin; Prucnal, Paul R; Bubnov, Mikhail M

    2007-10-01

    We present an experimental demonstration of an ultrafast all-optical thresholder based on a nonlinear Sagnac interferometer. The proposed design is intended for operation at very small nonlinear phase shifts. Therefore, it requires an in-loop nonlinearity lower than for the classical nonlinear loop mirror scheme. Only 15 meters of conventional (non-holey) silica-based fiber is used as a nonlinear element. The proposed thresholder is polarization insensitive and is good for multi-wavelength operation, meeting all the requirements for autocorrelation detection in various optical CDMA communication systems. The observed cubic transfer function is superior to the quadratic transfer function of second harmonic generation-based thresholders.

  20. The giant mottled eel, Anguilla marmorata, uses blue-shifted rod photoreceptors during upstream migration.

    Directory of Open Access Journals (Sweden)

    Feng-Yu Wang

    Full Text Available Catadromous fishes migrate between ocean and freshwater during particular phases of their life cycle. The dramatic environmental changes shape their physiological features, e.g. visual sensitivity, olfactory ability, and salinity tolerance. Anguilla marmorata, a catadromous eel, migrates upstream on dark nights, following the lunar cycle. Such behavior may be correlated with ontogenetic changes in sensory systems. Therefore, this study was designed to identify changes in spectral sensitivity and opsin gene expression of A. marmorata during upstream migration. Microspectrophotometry analysis revealed that the tropical eel possesses a duplex retina with rod and cone photoreceptors. The λmax of rod cells are 493, 489, and 489 nm in glass, yellow, and wild eels, while those of cone cells are 508, and 517 nm in yellow, and wild eels, respectively. Unlike European and American eels, Asian eels exhibited a blue-shifted pattern of rod photoreceptors during upstream migration. Quantitative gene expression analyses of four cloned opsin genes (Rh1f, Rh1d, Rh2, and SWS2 revealed that Rh1f expression is dominant at all three stages, while Rh1d is expressed only in older yellow eel. Furthermore, sequence comparison and protein modeling studies implied that a blue shift in Rh1d opsin may be induced by two known (N83, S292 and four putative (S124, V189, V286, I290 tuning sites adjacent to the retinal binding sites. Finally, expression of blue-shifted Rh1d opsin resulted in a spectral shift in rod photoreceptors. Our observations indicate that the giant mottled eel is color-blind, and its blue-shifted scotopic vision may influence its upstream migration behavior and habitat choice.

  1. The Nonlinear Field Space Theory

    Science.gov (United States)

    Mielczarek, Jakub; Trześniewski, Tomasz

    2016-08-01

    In recent years the idea that not only the configuration space of particles, i.e. spacetime, but also the corresponding momentum space may have nontrivial geometry has attracted significant attention, especially in the context of quantum gravity. The aim of this letter is to extend this concept to the domain of field theories, by introducing field spaces (i.e. phase spaces of field values) that are not affine spaces. After discussing the motivation and general aspects of our approach we present a detailed analysis of the prototype (quantum) Nonlinear Field Space Theory of a scalar field on the Minkowski background. We show that the nonlinear structure of a field space leads to numerous interesting predictions, including: non-locality, generalization of the uncertainty relations, algebra deformations, constraining of the maximal occupation number, shifting of the vacuum energy and renormalization of the charge and speed of propagation of field excitations. Furthermore, a compact field space is a natural way to implement the "Principle of finiteness" of physical theories, which once motivated the Born-Infeld theory. Thus the presented framework has a variety of potential applications in the theories of fundamental interactions (e.g. quantum gravity), as well as in condensed matter physics (e.g. continuous spin chains), and can shed new light on the issue of divergences in quantum field theories.

  2. The Nonlinear Field Space Theory

    Energy Technology Data Exchange (ETDEWEB)

    Mielczarek, Jakub, E-mail: jakub.mielczarek@uj.edu.pl [Institute of Physics, Jagiellonian University, ul. Łojasiewicza 11, 30-348 Kraków (Poland); Trześniewski, Tomasz, E-mail: tbwbt@ift.uni.wroc.pl [Institute of Physics, Jagiellonian University, ul. Łojasiewicza 11, 30-348 Kraków (Poland); Institute for Theoretical Physics, University of Wrocław, pl. Borna 9, 50-204 Wrocław (Poland)

    2016-08-10

    In recent years the idea that not only the configuration space of particles, i.e. spacetime, but also the corresponding momentum space may have nontrivial geometry has attracted significant attention, especially in the context of quantum gravity. The aim of this letter is to extend this concept to the domain of field theories, by introducing field spaces (i.e. phase spaces of field values) that are not affine spaces. After discussing the motivation and general aspects of our approach we present a detailed analysis of the prototype (quantum) Nonlinear Field Space Theory of a scalar field on the Minkowski background. We show that the nonlinear structure of a field space leads to numerous interesting predictions, including: non-locality, generalization of the uncertainty relations, algebra deformations, constraining of the maximal occupation number, shifting of the vacuum energy and renormalization of the charge and speed of propagation of field excitations. Furthermore, a compact field space is a natural way to implement the “Principle of finiteness” of physical theories, which once motivated the Born–Infeld theory. Thus the presented framework has a variety of potential applications in the theories of fundamental interactions (e.g. quantum gravity), as well as in condensed matter physics (e.g. continuous spin chains), and can shed new light on the issue of divergences in quantum field theories.

  3. The Nonlinear Field Space Theory

    Directory of Open Access Journals (Sweden)

    Jakub Mielczarek

    2016-08-01

    Full Text Available In recent years the idea that not only the configuration space of particles, i.e. spacetime, but also the corresponding momentum space may have nontrivial geometry has attracted significant attention, especially in the context of quantum gravity. The aim of this letter is to extend this concept to the domain of field theories, by introducing field spaces (i.e. phase spaces of field values that are not affine spaces. After discussing the motivation and general aspects of our approach we present a detailed analysis of the prototype (quantum Nonlinear Field Space Theory of a scalar field on the Minkowski background. We show that the nonlinear structure of a field space leads to numerous interesting predictions, including: non-locality, generalization of the uncertainty relations, algebra deformations, constraining of the maximal occupation number, shifting of the vacuum energy and renormalization of the charge and speed of propagation of field excitations. Furthermore, a compact field space is a natural way to implement the “Principle of finiteness” of physical theories, which once motivated the Born–Infeld theory. Thus the presented framework has a variety of potential applications in the theories of fundamental interactions (e.g. quantum gravity, as well as in condensed matter physics (e.g. continuous spin chains, and can shed new light on the issue of divergences in quantum field theories.

  4. Differential ontogenetic exposure to obesogenic environment induces hyperproliferative status and nuclear receptors imbalance in the rat prostate at adulthood.

    Science.gov (United States)

    Pytlowanciv, Eloísa Zanin; Pinto-Fochi, Maria Etelvina; Reame, Vanessa; Gobbo, Marina Guimarães; Ribeiro, Daniele Lisboa; Taboga, Sebastião Roberto; Góes, Rejane Maira

    2016-05-01

    Experimental data indicate that high-fat diet (HFD) may alter proliferative activity and prostate health. However, the consequences of HFD exposure during different periods of ontogenetic development on prostate histophysiology remain to be elucidated. Herein, we compare the influence of obesogenic environment (OE) due to maternal obesity and HFD at different periods of life on proliferative activity and nuclear receptors frequency in the rat ventral prostate and a possible relationship with metabolic and hormonal alterations. Male Wistar rats (19 weeks old), treated with balanced chow (Control group-C; 3% high-fat, 3.5 Kcal/g), were compared with those exposed to HFD (20% high-fat, 4.9 kcal/g) during gestation (G-maternal obesity), gestation and lactation (GL), from post-weaning to adulthood (WA), from lactation to adulthood (LA) and from gestation to adulthood (GA). After the experimental period, the ventral prostate lobes were removed and analyzed with different methods. Metabolic data indicated that G and GL rats became insulin resistant and WA, LA, and GA became insulin resistant and obese. There was a strong inverse correlation between serum testosterone (∼133% lower) and leptin levels (∼467% higher) in WA, LA, and GA groups. Estrogen serum levels increased in GA, and insulin levels increased in all groups, especially in WA (64.8×). OE-groups exhibited prostatic hypertrophy, since prostate weight increased ∼40% in G, GL, LA, and GA and 31% in WA. As indicated by immunohistochemistry, all HFD-groups except G exhibited an increase in epithelial cell proliferation (PCNA-positive) and a decrease in frequency of AR- and ERβ-positive epithelial cells; there was also an increment of ERα-positive stromal cells in comparison with control. Cells containing PPARγ increased in both epithelium and stroma of all OE groups and those expressing LXRα decreased, particularly in groups OE-exposed during gestation (G, GL and GA). OE leads to prostate hypertrophy

  5. The ontogenetic transformation of the mesosaurid tarsus: a contribution to the origin of the primitive amniotic astragalus

    Directory of Open Access Journals (Sweden)

    Graciela Piñeiro

    2016-05-01

    Full Text Available The hypotheses about the origin of the primitive amniotic tarsus are very speculative. Early studies argued that the origin of the astragalus, one of the largest proximal bones in the tarsus of basal amniotes, was produced by either the fusion of two, three, or even four of the original tarsal bones, the intermedium, the tibiale and the proximal centralia (c4 and c3, or that the intermedium alone transforms into the primitive astragalus. More recent studies have shown that the structure of the tarsus in Captorhinus supports the former hypothesis about a fusion of the intermedium, the tibiale, the proximal centrale (c4 and eventually c3, producing a purportedly multipartite structure of the amniotic astragalus, but the issue remained contentious. Very well preserved tarsi of the Early Permian aquatic amniote Mesosaurus tenuidens Gervais, 1864–1865, which represent the most complete ontogenetic succession known for a basal amniote (the other exceptional one is provided by the Late Permian diapsid Hovasaurus boulei Piveteau, 1926, suggest that there is more than one ossification center for the astragalus and that these fuse during late embryonic stages or maybe early after birth. A non-hatched Mesosaurus in an advanced stage of development shows that the tarsus is represented by a single bone, most probably the astragalus, which seems to be formed by the suturing of three bones, here interpreted as being the intermedium, the tibiale, probably already integrated to the c4 in an earlier stage of the development, and the c3. An amniote-like tarsal structure is observed in very basal Carboniferous and Permian tetrapods such as Proterogyrinus, Gephyrostegus, the diadectids Diadectes and Orobates, some microsaurs like Tuditanus and Pantylus and possibly Westlothiana, taxa that were all considered as true amniotes in their original descriptions. Therefore, the structure of the amniotic tarsus, including the configuration of the proximal series formed by

  6. Variability of root traits in common bean genotypes at different levels of phosphorus supply and ontogenetic stages

    Directory of Open Access Journals (Sweden)

    Roberto dos Santos Trindade

    2014-08-01

    Full Text Available Selection of common bean (Phaseolus vulgaris L. cultivars with enhanced root growth would be a strategy for increasing P uptake and grain yield in tropical soils, but the strong plasticity of root traits may compromise their inclusion in breeding programs. The aim of this study was to evaluate the magnitude of the genotypic variability of root traits in common bean plants at two ontogenetic stages and two soil P levels. Twenty-four common bean genotypes, comprising the four growth habits that exist in the species and two wild genotypes, were grown in 4 kg pots at two levels of applied P (20 and 80 mg kg-1 and harvested at the stages of pod setting and early pod filling. Root area and root length were measured by digital image analysis. Significant genotype × P level and genotype × harvest interactions in analysis of variance indicate that the genotypic variation of root traits depended on soil nutrient availability and the stage at which evaluation was made. Genotypes differed for taproot mass, basal and lateral root mass, root area and root length at both P levels and growth stages; differences in specific root area and length were small. Genotypes with growth habits II (upright indeterminate and III (prostrate indeterminate showed better adaptation to limited P supply than genotypes of groups I (determinate and IV (indeterminate climbing. Between the two harvests, genotypes of groups II and III increased the mass of basal and lateral roots by 40 and 50 %, respectively, whereas genotypes of groups I and IV by only 7 and 19 %. Values of the genotypic coefficient of determination, which estimates the proportion of phenotypic variance resulting from genetic effects, were higher at early pod filling than at pod setting. Correlations between shoot mass and root mass, which could indicate indirect selection of root systems via aboveground biomass, were higher at early pod filling than at pod setting. The results indicate that selection for root

  7. The ontogenetic transformation of the mesosaurid tarsus: a contribution to the origin of the primitive amniotic astragalus.

    Science.gov (United States)

    Piñeiro, Graciela; Núñez Demarco, Pablo; Meneghel, Melitta D

    2016-01-01

    The hypotheses about the origin of the primitive amniotic tarsus are very speculative. Early studies argued that the origin of the astragalus, one of the largest proximal bones in the tarsus of basal amniotes, was produced by either the fusion of two, three, or even four of the original tarsal bones, the intermedium, the tibiale and the proximal centralia (c4 and c3), or that the intermedium alone transforms into the primitive astragalus. More recent studies have shown that the structure of the tarsus in Captorhinus supports the former hypothesis about a fusion of the intermedium, the tibiale, the proximal centrale (c4) and eventually c3, producing a purportedly multipartite structure of the amniotic astragalus, but the issue remained contentious. Very well preserved tarsi of the Early Permian aquatic amniote Mesosaurus tenuidens Gervais, 1864-1865, which represent the most complete ontogenetic succession known for a basal amniote (the other exceptional one is provided by the Late Permian diapsid Hovasaurus boulei Piveteau, 1926), suggest that there is more than one ossification center for the astragalus and that these fuse during late embryonic stages or maybe early after birth. A non-hatched Mesosaurus in an advanced stage of development shows that the tarsus is represented by a single bone, most probably the astragalus, which seems to be formed by the suturing of three bones, here interpreted as being the intermedium, the tibiale, probably already integrated to the c4 in an earlier stage of the development, and the c3. An amniote-like tarsal structure is observed in very basal Carboniferous and Permian tetrapods such as Proterogyrinus, Gephyrostegus, the diadectids Diadectes and Orobates, some microsaurs like Tuditanus and Pantylus and possibly Westlothiana, taxa that were all considered as true amniotes in their original descriptions. Therefore, the structure of the amniotic tarsus, including the configuration of the proximal series formed by the astragalus and

  8. An in situ, individual-based approach to quantify connectivity of marine fish: ontogenetic movements and residency of lingcod.

    Directory of Open Access Journals (Sweden)

    Mary Anne Bishop

    Full Text Available As modern fishery assessments change in an effort to be more accurate and encompass the range of potential ecosystem interactions, critical information on the ecology of species including life history, intra and inter-specific competitive interactions and habitat requirements must be added to the standard fishery-dependent and independent data sets. One species whose movements and habitat associations greatly affects exploitation patterns is lingcod, Ophiodon elongatus, which support an economically important fishery along the coastal waters of the Pacific Coast of North America. High site fidelity and limited movements within nearshore areas are hypothesized to have resulted in high catchability, a major factor that has contributed to overfished stocks. Thus, assessing the level of movement and connectivity among lingcod subpopulations inhabiting nearshore habitats is a prerequisite to determining the condition of lingcod stocks. We used the Pacific Ocean Shelf Tracking (POST Project acoustic receiver array in Alaska's Prince William Sound to monitor movements and residency of 21 acoustic-tagged lingcod for up to 16 months. Eight of sixteen lingcod (50% initially aged at 2.5- to 3.5- years-old dispersed from their tag site. Dispersal was highly seasonal, occurring in two, five-week periods from mid-December through January and from mid-April through May. Dispersal in winter may be related to sexually immature lingcod or newly-mature male lingcod being displaced by territorial males. Spring dispersal may be indicative of the onset of migratory behavior where lingcod move out into Prince William Sound and possibly the offshore waters of the Gulf of Alaska. Our results reveal a pattern of ontogenetic dispersal as lingcod approach 4-years-old and exceed 50 cm total length. The large proportion of tagged fish migrating out of Port Gravina, their tagging site, reflects a high level of connectivity among Prince William Sound subpopulations. Our

  9. Ontogenetic exposure of rats to pre- and post-natal manganese enhances behavioral impairments produced by perinatal 6-hydroxydopamine.

    Science.gov (United States)

    Nowak, Przemysław; Bojanek, Kamila; Szkilnik, Ryszard; Jośko, Jadwiga; Boroń, Dariusz; Adwent, Marta; Gorczyca, Piotr; Kostrzewa, Richard M; Brus, Ryszard

    2011-05-01

    Rats lesioned shortly after birth with 6-hydroxydopamine (6-OHDA; 134 μg icv) represent a near-ideal model of severe Parkinson's disease because of the near-total destruction of nigrostriatal dopaminergic fibers. The element manganese, an essential cofactor for many enzymatic reactions, itself in toxic amount, replicates some clinical features similar to those of Parkinson's disease. The aim of this study was to examine the effect of neonatal manganese exposure on 6-OHDA modeling of Parkinson's disease in rats. Manganese (MnCl(2)·4H(2)O) 10,000 ppm was included in the drinking water of pregnant Wistar rats from the time of conception until the 21st day after delivery, the age when neonatal rats were weaned. Control rats consumed tap water. Other groups of neonatal rat pups, on the 3rd day after birth, were pretreated with desipramine (20 mg/kg ip 1 h) prior to bilateral icv administration of 6-OHDA (30, 60, or 137 μg) or its vehicle saline-ascorbic (0.1%) (control). At 2 months after birth, in rats lesioned with 30, 60, or 134 μg 6-OHDA, endogenous striatal dopamine (DA) content was reduced, respectively, by 66, 92, and 98% (HPLC/ED), while co-exposure of these groups to perinatal manganese did not magnify the DA depletion. However, there was prominent enhancement of DA D(1) agonist (i.e., SKF 38393)-induced oral activity in the group of rats exposed perinatally to manganese and also treated neonatally with the 30 mg/kg dose of 6-OHDA. The 30 mg/kg 6-OHDA group, demonstrating cataleptogenic responses to SCH 23390 (0.5 mg/kg) and haloperidol (0.5 mg/kg ip), developed resistance if co-exposed to perinatal manganese. In the group exposed to manganese and lesioned with the 60 mg/kg dose of 6-OHDA, there was a reduction in D(2) agonist (i.e., quinpirole, 0.1 mg/kg)-induced yawning. The series of findings demonstrate that ontogenetic exposure to manganese results in an enhancement of behavioral toxicity to a moderate dose of 6-OHDA, despite the fact that

  10. Nonlinear Magneto-optical Rotation via Alignment-to-Orientation Conversion

    Energy Technology Data Exchange (ETDEWEB)

    Budker, D.; Kimball, D.F.; Rochester, S.M.; Yashchuk, V.V.

    2000-03-10

    Nonlinear magneto-optical rotation (NMOR) is investigated at highlight powers where the rotation is significantly modified by AC Stark shifts. These shifts are shown to change the overall sign of rotation for closed F-->F+1 transitions as light power is increased. The effect is demonstrated by measurements in rubidium and density matrix calculations. The results are important for applications of nonlinear optical rotation such as sensitive magnetometry.

  11. Quadrature phase-shift error analysis using a homodyne laser interferometer.

    Science.gov (United States)

    Gregorcic, Peter; Pozar, Tomaz; Mozina, Janez

    2009-08-31

    The influence of quadrature phase shift on the measured displacement error was experimentally investigated using a two-detector polarizing homodyne laser interferometer with a quadrature detection system. Common nonlinearities, including the phase-shift error, were determined and effectively corrected by a robust data-processing algorithm. The measured phase-shift error perfectly agrees with the theoretically determined phase-shift error region. This error is systematic, periodic and severely asymmetrical around the nominal displacement value. The main results presented in this paper can also be used to assess and correct the detector errors of other interferometric and non-interferometric displacement-measuring devices based on phase-quadrature detection.

  12. Immunohistochemical study on the ontogenetic development of the regional distribution of peptide YY, pancreatic polypeptide, and glucagon-like peptide 1 endocrine cells in bovine gastrointestinal tract.

    Science.gov (United States)

    Pyarokhil, Asadullah Hamid; Ishihara, Miyuki; Sasaki, Motoki; Kitamura, Nobuo

    2012-04-10

    The regional distribution and relative frequency of peptide YY (PYY)-, pancreatic polypeptide (PP)-, and glucagon-like peptide 1 (GLP-1)-immunoreactive (IR) cells were determined immunohistochemically in the gastrointestinal tract at seven ontogenetic stages in pre- and postnatal cattle. Different frequencies of PYY-, PP-, and GLP-1-IR cells were found in the intestines at all stages; they were not found in the esophagus and stomach. The frequencies varied depending on the intestinal segment and the developmental stage. The frequencies of PYY- and PP-IR cells were lower in the small intestine and increased from ileum to rectum, whereas GLP-1-IR cells were more numerous in duodenum and jejunum, decreased in ileum and cecum, and increased again in colon and rectum. The frequencies also varied according to pre- and postnatal stages. All three cell types were most numerous in fetus, and decreased in calf and adult groups, indicating that the frequencies of these three types of endocrine cells decrease with postnatal development. The results suggest that these changes vary depending on feeding habits and adaptation of growth, secretion, and motility of intestine at different ontogenetic stages of cattle.

  13. Lasers for nonlinear microscopy.

    Science.gov (United States)

    Wise, Frank

    2013-03-01

    Various versions of nonlinear microscopy are revolutionizing the life sciences, almost all of which are made possible because of the development of ultrafast lasers. In this article, the main properties and technical features of short-pulse lasers used in nonlinear microscopy are summarized. Recent research results on fiber lasers that will impact future instruments are also discussed.

  14. Nonlinear optical materials.

    Science.gov (United States)

    Eaton, D F

    1991-07-19

    The current state of materials development in nonlinear optics is summarized, and the promise of these materials is critically evaluated. Properties and important materials constants of current commercial materials and of new, promising, inorganic and organic molecular and polymeric materials with potential in second- and third-order nonlinear optical applications are presented.

  15. Estimating nonlinear models

    Science.gov (United States)

    Billings, S. A.

    1988-03-01

    Time and frequency domain identification methods for nonlinear systems are reviewed. Parametric methods, prediction error methods, structure detection, model validation, and experiment design are discussed. Identification of a liquid level system, a heat exchanger, and a turbocharge automotive diesel engine are illustrated. Rational models are introduced. Spectral analysis for nonlinear systems is treated. Recursive estimation is mentioned.

  16. Nonlinear Cross Gramians

    NARCIS (Netherlands)

    Ionescu, T. C.; Scherpen, J. M. A.; Korytowski, A; Malanowski, K; Mitkowski, W; Szymkat, M

    2009-01-01

    We study the notion of cross Gramians for nonlinear gradient systems, using the characterization in terms of prolongation and gradient extension associated to the system. The cross Gramian is given for the variational system associated to the original nonlinear gradient system. We obtain

  17. Engineered nonlinear lattices

    DEFF Research Database (Denmark)

    Clausen, Carl A. Balslev; Christiansen, Peter Leth; Torner, L.

    1999-01-01

    We show that with the quasi-phase-matching technique it is possible to fabricate stripes of nonlinearity that trap and guide light like waveguides. We investigate an array of such stripes and find that when the stripes are sufficiently narrow, the beam dynamics is governed by a quadratic nonlinear...

  18. Controllability in nonlinear systems

    Science.gov (United States)

    Hirschorn, R. M.

    1975-01-01

    An explicit expression for the reachable set is obtained for a class of nonlinear systems. This class is described by a chain condition on the Lie algebra of vector fields associated with each nonlinear system. These ideas are used to obtain a generalization of a controllability result for linear systems in the case where multiplicative controls are present.

  19. Nonlinear Maneuver Autopilot

    Science.gov (United States)

    Menon, P. K. A.; Badgett, M. E.; Walker, R. A.

    1992-01-01

    Trajectory-control laws based on singular-perturbation theory and nonlinear dynamical modeling. Nonlinear maneuver autopilot commands flight-test trajectories of F-15 airplane. Underlying theory of controller enables separation of variables processed in fast and slow control loops, reducing amount of computation required.

  20. Quantized beam shifts in graphene

    Energy Technology Data Exchange (ETDEWEB)

    de Melo Kort-Kamp, Wilton Junior [Los Alamos National Lab. (LANL), Los Alamos, NM (United States); Sinitsyn, Nikolai [Los Alamos National Lab. (LANL), Los Alamos, NM (United States); Dalvit, Diego Alejandro Roberto [Los Alamos National Lab. (LANL), Los Alamos, NM (United States)

    2015-10-08

    We predict the existence of quantized Imbert-Fedorov, Goos-Hanchen, and photonic spin Hall shifts for light beams impinging on a graphene-on-substrate system in an external magnetic field. In the quantum Hall regime the Imbert-Fedorov and photonic spin Hall shifts are quantized in integer multiples of the fine structure constant α, while the Goos-Hanchen ones in multiples of α2. We investigate the influence on these shifts of magnetic field, temperature, and material dispersion and dissipation. An experimental demonstration of quantized beam shifts could be achieved at terahertz frequencies for moderate values of the magnetic field.

  1. Beam shifts and distribution functions

    CERN Document Server

    Aiello, Andrea

    2011-01-01

    When a beam of light is reflected by a smooth surface its behavior deviates from geometrical optics predictions. Such deviations are quantified by the so-called spatial and angular Goos-Haenchen (GH) and Imbert-Fedorov (IF) shifts of the reflected beam. These shifts depend upon the shape of the incident beam, its polarization and on the material composition of the reflecting surface. In this article we suggest a novel approach that allows one to unambiguously isolate the beam-shape dependent aspects of GH and IF shifts. We show that this separation is possible as a result of some universal features of shifted distribution functions which are presented and discussed.

  2. Nonlinear optics and photonics

    CERN Document Server

    He, Guang S

    2015-01-01

    This book provides a comprehensive presentation on most of the major topics in nonlinear optics and photonics, with equal emphasis on principles, experiments, techniques, and applications. It covers many major new topics including optical solitons, multi-photon effects, nonlinear photoelectric effects, fast and slow light , and Terahertz photonics. Chapters 1-10 present the fundamentals of modern nonlinear optics, and could be used as a textbook with problems provided at the end of each chapter. Chapters 11-17 cover the more advanced topics of techniques and applications of nonlinear optics and photonics, serving as a highly informative reference for researchers and experts working in related areas. There are also 16 pages of color photographs to illustrate the visual appearances of some typical nonlinear optical effects and phenomena. The book could be adopted as a textbook for both undergraduates and graduate students, and serve as a useful reference work for researchers and experts in the fields of physics...

  3. Nonlinear optical systems

    CERN Document Server

    Lugiato, Luigi; Brambilla, Massimo

    2015-01-01

    Guiding graduate students and researchers through the complex world of laser physics and nonlinear optics, this book provides an in-depth exploration of the dynamics of lasers and other relevant optical systems, under the umbrella of a unitary spatio-temporal vision. Adopting a balanced approach, the book covers traditional as well as special topics in laser physics, quantum electronics and nonlinear optics, treating them from the viewpoint of nonlinear dynamical systems. These include laser emission, frequency generation, solitons, optically bistable systems, pulsations and chaos and optical pattern formation. It also provides a coherent and up-to-date treatment of the hierarchy of nonlinear optical models and of the rich variety of phenomena they describe, helping readers to understand the limits of validity of each model and the connections among the phenomena. It is ideal for graduate students and researchers in nonlinear optics, quantum electronics, laser physics and photonics.

  4. Nonlinear fiber-optic strain sensor based on four-wave mixing in microstructured optical fiber

    DEFF Research Database (Denmark)

    Gu, Bobo; Yuan, Scott Wu; Frosz, Michael H.

    2012-01-01

    We demonstrate a nonlinear fiber-optic strain sensor, which uses the shifts of four-wave mixing Stokes and anti-Stokes peaks caused by the strain-induced changes in the structure and refractive index of a microstructured optical fiber. The sensor thus uses the inherent nonlinearity of the fiber...

  5. Designing quadratic nonlinear photonic crystal fibers for soliton compression to few-cycle pulses

    DEFF Research Database (Denmark)

    Bache, Morten; Moses, Jeffrey; Lægsgaard, Jesper

    2007-01-01

    Second-harmonic generation (SHG) in the limit of large phase mismatch, given by Deltabeta=beta2-2beta1 effectively induces a Kerr-like nonlinear phase shift on the fundamental wave (FW). The phase mismatch determines the sign and magnitude of the effective Kerr nonlinearity, making large negative...

  6. Regime shifts in resource management

    NARCIS (Netherlands)

    de Zeeuw, A.J.

    2014-01-01

    Resource management has to take account of the possibility of tipping points and regime shifts in ecological systems that provide the resources. This article focuses on the typical model of regime shifts in the ecological literature and analyzes optimal management and common-property issues when tra

  7. Work shifts in Emergency Medicine

    Directory of Open Access Journals (Sweden)

    Roberto Recupero

    2007-06-01

    Full Text Available Emergency Medicine is known as a high stress specialty. The adverse effect of constantly rotating shifts is the single most important reason given for premature attrition from the field. In this work problems tied with night shift work will be taken into account and some solutions to reduce the impact of night work on the emergency physicians will be proposed.

  8. Nonlinear cochlear mechanics.

    Science.gov (United States)

    Zweig, George

    2016-05-01

    An earlier paper characterizing the linear mechanical response of the organ of Corti [J. Acoust. Soc. Am. 138, 1102-1121 (2015)] is extended to the nonlinear domain. Assuming the existence of nonlinear oscillators nonlocally coupled through the pressure they help create, the oscillator equations are derived and examined when the stimuli are modulated tones and clicks. The nonlinearities are constrained by the requirements of oscillator stability and the invariance of zero crossings in the click response to changes in click amplitude. The nonlinear oscillator equations for tones are solved in terms of the fluid pressure that drives them, and its time derivative, presumably a proxy for forces created by outer hair cells. The pressure equation is reduced to quadrature, the integrand depending on the oscillators' responses. The resulting nonlocally coupled nonlinear equations for the pressure, and oscillator amplitudes and phases, are solved numerically in terms of the fluid pressure at the stapes. Methods for determining the nonlinear damping directly from measurements are described. Once the oscillators have been characterized from their tone and click responses, the mechanical response of the cochlea to natural sounds may be computed numerically. Signal processing inspired by cochlear mechanics opens up a new area of nonlocal nonlinear time-frequency analysis.

  9. Fibre-optic nonlinear optical microscopy and endoscopy.

    Science.gov (United States)

    Fu, L; Gu, M

    2007-06-01

    Nonlinear optical microscopy has been an indispensable laboratory tool of high-resolution imaging in thick tissue and live animals. Rapid developments of fibre-optic components in terms of growing functionality and decreasing size provide enormous opportunities for innovations in nonlinear optical microscopy. Fibre-based nonlinear optical endoscopy is the sole instrumentation to permit the cellular imaging within hollow tissue tracts or solid organs that are inaccessible to a conventional optical microscope. This article reviews the current development of fibre-optic nonlinear optical microscopy and endoscopy, which includes crucial technologies for miniaturized nonlinear optical microscopy and their embodiments of endoscopic systems. A particular attention is given to several classes of photonic crystal fibres that have been applied to nonlinear optical microscopy due to their unique properties for ultrashort pulse delivery and signal collection. Furthermore, fibre-optic nonlinear optical imaging systems can be classified into portable microscopes suitable for imaging behaving animals, rigid endoscopes that allow for deep tissue imaging with minimally invasive manners, and flexible endoscopes enabling imaging of internal organs. Fibre-optic nonlinear optical endoscopy is coming of age and a paradigm shift leading to optical microscope tools for early cancer detection and minimally invasive surgery.

  10. Topological nature of nonlinear optical effects in solids.

    Science.gov (United States)

    Morimoto, Takahiro; Nagaosa, Naoto

    2016-05-01

    There are a variety of nonlinear optical effects including higher harmonic generations, photovoltaic effects, and nonlinear Kerr rotations. They are realized by strong light irradiation to materials that results in nonlinear polarizations in the electric field. These are of great importance in studying the physics of excited states of the system as well as for applications to optical devices and solar cells. Nonlinear properties of materials are usually described by nonlinear susceptibilities, which have complex expressions including many matrix elements and energy denominators. On the other hand, a nonequilibrium steady state under an electric field periodic in time has a concise description in terms of the Floquet bands of electrons dressed by photons. We show theoretically, using the Floquet formalism, that various nonlinear optical effects, such as the shift current in noncentrosymmetric materials, photovoltaic Hall response, and photo-induced change of order parameters under the continuous irradiation of monochromatic light, can be described in a unified fashion by topological quantities involving the Berry connection and Berry curvature. We found that vector fields defined with the Berry connections in the space of momentum and/or parameters govern the nonlinear responses. This topological view offers a route to designing nonlinear optical materials.

  11. Si-rich Silicon Nitride for Nonlinear Signal Processing Applications.

    Science.gov (United States)

    Lacava, Cosimo; Stankovic, Stevan; Khokhar, Ali Z; Bucio, T Dominguez; Gardes, F Y; Reed, Graham T; Richardson, David J; Petropoulos, Periklis

    2017-02-02

    Nonlinear silicon photonic devices have attracted considerable attention thanks to their ability to show large third-order nonlinear effects at moderate power levels allowing for all-optical signal processing functionalities in miniaturized components. Although significant efforts have been made and many nonlinear optical functions have already been demonstrated in this platform, the performance of nonlinear silicon photonic devices remains fundamentally limited at the telecom wavelength region due to the two photon absorption (TPA) and related effects. In this work, we propose an alternative CMOS-compatible platform, based on silicon-rich silicon nitride that can overcome this limitation. By carefully selecting the material deposition parameters, we show that both of the device linear and nonlinear properties can be tuned in order to exhibit the desired behaviour at the selected wavelength region. A rigorous and systematic fabrication and characterization campaign of different material compositions is presented, enabling us to demonstrate TPA-free CMOS-compatible waveguides with low linear loss (~1.5 dB/cm) and enhanced Kerr nonlinear response (Re{γ} = 16 Wm(-1)). Thanks to these properties, our nonlinear waveguides are able to produce a π nonlinear phase shift, paving the way for the development of practical devices for future optical communication applications.

  12. Metabolic impact of shift work.

    Science.gov (United States)

    Zimberg, Ioná Zalcman; Fernandes Junior, Silvio A; Crispim, Cibele Aparecida; Tufik, Sergio; de Mello, Marco Tulio

    2012-01-01

    In developing countries, shift work represents a considerable contingent workforce. Recently, studies have shown that overweight and obesity are more prevalent in shift workers than day workers. In addition, shift work has been associated with a higher propensity for the development of many metabolic disorders, such as insulin resistance, diabetes, dislipidemias and metabolic syndrome. Recent data have pointed that decrease of the sleep time, desynchronization of circadian rhythm and alteration of environmental aspects are the main factors related to such problems. Shortened or disturbed sleep is among the most common health-related effects of shift work. The plausible physiological and biological mechanisms are related to the activation of the autonomic nervous system, inflammation, changes in lipid and glucose metabolism, and related changes in the risk for atherosclerosis, metabolic syndrome, and type II diabetes. The present review will discuss the impact of shift work on obesity and metabolic disorders and how disruption of sleep and circadian misalignment may contribute to these metabolic dysfunctions.

  13. Nonlinear fiber optics

    CERN Document Server

    Agrawal, Govind P

    2001-01-01

    The Optical Society of America (OSA) and SPIE - The International Society for Optical Engineering have awarded Govind Agrawal with an honorable mention for the Joseph W. Goodman Book Writing Award for his work on Nonlinear Fiber Optics, 3rd edition.Nonlinear Fiber Optics, 3rd Edition, provides a comprehensive and up-to-date account of the nonlinear phenomena occurring inside optical fibers. It retains most of the material that appeared in the first edition, with the exception of Chapter 6, which is now devoted to the polarization effects relevant for light propagation in optical

  14. Will Nonlinear Backcalculation Help?

    DEFF Research Database (Denmark)

    Ullidtz, Per

    2000-01-01

    demonstrates, that treating the subgrade as a nonlinear elastic material, can result in more realistic moduli and a much better agreement between measured and calculated stresses and strains.The response of nonlinear elastic materials can be calculated using the Finite Element Method (FEM). A much simpler...... approach is to use the Method of Equivalent Thicknesses (MET), modified for a nonlinear subgrade. The paper includes an example where moduli backcalculated using FEM, linear elastic theory and MET are compared. Stresses and strains predicted by the three methods are also compared to measured values...

  15. Nonlinear graphene metamaterial

    CERN Document Server

    Nikolaenko, Andrey E; Atmatzakis, Evangelos; Luo, Zhiqiang; Shen, Ze Xiang; De Angelis, Francesco; Boden, Stuart A; Di Fabrizio, Enzo; Zheludev, Nikolay I

    2012-01-01

    We demonstrate that the broadband nonlinear optical response of graphene can be resonantly enhanced by more than an order of magnitude through hybridization with a plasmonic metamaterial,while retaining an ultrafast nonlinear response time of ~1 ps. Transmission modulation close to ~1% is seen at a pump uence of ~0.03 mJ/cm^2 at the wavelength of ~1600 nm. This approach allows to engineer and enhance graphene's nonlinearity within a broad wavelength range enabling applications in optical switching, mode-locking and pulse shaping.

  16. Nonlinear characterization of GeSbS chalcogenide glass waveguides

    Science.gov (United States)

    Choi, Ju Won; Han, Zhaohong; Sohn, Byoung-Uk; Chen, George F. R.; Smith, Charmayne; Kimerling, Lionel C.; Richardson, Kathleen A.; Agarwal, Anuradha M.; Tan, Dawn T. H.

    2016-12-01

    GeSbS ridge waveguides have recently been demonstrated as a promising mid – infrared platform for integrated waveguide – based chemical sensing and photodetection. To date, their nonlinear optical properties remain relatively unexplored. In this paper, we characterize the nonlinear optical properties of GeSbS glasses, and show negligible nonlinear losses at 1.55 μm. Using self – phase modulation experiments, we characterize a waveguide nonlinear parameter of 7 W‑1/m and nonlinear refractive index of 3.71 × 10‑18 m2/W. GeSbS waveguides are used to generate supercontinuum from 1280 nm to 2120 nm at the ‑30 dB level. The spectrum expands along the red shifted side of the spectrum faster than on the blue shifted side, facilitated by cascaded stimulated Raman scattering arising from the large Raman gain of chalcogenides. Fourier transform infrared spectroscopic measurements show that these glasses are optically transparent up to 25 μm, making them useful for short – wave to long – wave infrared applications in both linear and nonlinear optics.

  17. The Nonlinear Talbot Effect of Rogue Waves

    CERN Document Server

    Zhang, Yiqi; Zheng, Huaibin; Chen, Haixia; Li, Changbiao; Song, Jianping; Zhang, Yanpeng

    2014-01-01

    Akhmediev and Kuznetsov-Ma breathers are rogue wave solutions of the nonlinear Schr\\"odinger equation (NLSE). Talbot effect (TE) is an image recurrence phenomenon in the diffraction of light waves. We report the nonlinear TE of rogue waves in a cubic medium. It is different from the linear TE, in that the wave propagates in a NL medium and is an eigenmode of NLSE. Periodic rogue waves impinging on a NL medium exhibit recurrent behavior, but only at the TE length and at the half-TE length with a \\pi-phase shift; the fractional TE is absent. The NL TE is the result of the NL interference of the lobes of rogue wave breathers. This interaction is related to the transverse period and intensity of breathers, in that the bigger the period and the higher the intensity, the shorter the TE length.

  18. Simplified Model of Nonlinear Landau Damping

    Energy Technology Data Exchange (ETDEWEB)

    N. A. Yampolsky and N. J. Fisch

    2009-07-16

    The nonlinear interaction of a plasma wave with resonant electrons results in a plateau in the electron distribution function close to the phase velocity of the plasma wave. As a result, Landau damping of the plasma wave vanishes and the resonant frequency of the plasma wave downshifts. However, this simple picture is invalid when the external driving force changes the plasma wave fast enough so that the plateau cannot be fully developed. A new model to describe amplification of the plasma wave including the saturation of Landau damping and the nonlinear frequency shift is proposed. The proposed model takes into account the change of the plasma wave amplitude and describes saturation of the Landau damping rate in terms of a single fluid equation, which simplifies the description of the inherently kinetic nature of Landau damping. A proposed fluid model, incorporating these simplifications, is verified numerically using a kinetic Vlasov code.

  19. On the Possibility of Using Nonlinear Elements for Landau Damping in High-Intensity Beams

    Energy Technology Data Exchange (ETDEWEB)

    Alexahin, Y. [Fermilab; Gianfelice-Wendt, E. [Fermilab; Lebedev, V. [Fermilab; Valishev, A. [Fermilab

    2016-09-30

    Direct space-charge force shifts incoherent tunes downwards from the coherent ones breaking the Landau mechanism of coherent oscillations damping at high beam intensity. To restore it nonlinear elements can be employed which move back tunes of large amplitude particles. In the present report we consider the possibility of creating a “nonlinear integrable optics” insertion in the Fermilab Recycler to host either octupoles or hollow electron lens for this purpose. For comparison we also consider the classic scheme with distributed octupole families. It is shown that for the Proton Improvement Plan II (PIP II) parameters the required nonlinear tune shift can be created without destroying the dynamic aperture.

  20. Multipolar nonlinear nanophotonics

    CERN Document Server

    Smirnova, Daria

    2016-01-01

    Nonlinear nanophotonics is a rapidly developing field with many useful applications for a design of nonlinear nanoantennas, light sources, nanolasers, sensors, and ultrafast miniature metadevices. A tight confinement of the local electromagnetic fields in resonant photonic nanostructures can boost nonlinear optical effects, thus offering versatile opportunities for subwavelength control of light. To achieve the desired functionalities, it is essential to gain flexible control over the near- and far-field properties of nanostructures. Thus, both modal and multipolar analyses are widely exploited for engineering nonlinear scattering from resonant nanoscale elements, in particular for enhancing the near-field interaction, tailoring the far-field multipolar interference, and optimization of the radiation directionality. Here, we review the recent advances in this recently emerged research field ranging from metallic structures exhibiting localized plasmonic resonances to hybrid metal-dielectric and all-dielectric...

  1. Solitons in nonlinear lattices

    CERN Document Server

    Kartashov, Yaroslav V; Torner, Lluis

    2010-01-01

    This article offers a comprehensive survey of results obtained for solitons and complex nonlinear wave patterns supported by purely nonlinear lattices (NLs), which represent a spatially periodic modulation of the local strength and sign of the nonlinearity, and their combinations with linear lattices. A majority of the results obtained, thus far, in this field and reviewed in this article are theoretical. Nevertheless, relevant experimental settings are surveyed too, with emphasis on perspectives for implementation of the theoretical predictions in the experiment. Physical systems discussed in the review belong to the realms of nonlinear optics (including artificial optical media, such as photonic crystals, and plasmonics) and Bose-Einstein condensation (BEC). The solitons are considered in one, two, and three dimensions (1D, 2D, and 3D). Basic properties of the solitons presented in the review are their existence, stability, and mobility. Although the field is still far from completion, general conclusions c...

  2. Nonlinear plasmonic antennas

    Directory of Open Access Journals (Sweden)

    Shakeeb Bin Hasan

    2014-12-01

    Full Text Available Contrary to traditional optical elements, plasmonic antennas made from nanostructured metals permit the localization of electromagnetic fields on length scales much smaller than the wavelength of light. This results in huge amplitudes for the electromagnetic field close to the antenna being conducive for the observation of nonlinear effects already at moderate pump powers. Thus, these antennas exhibit a promising potential to achieve optical frequency conversion and all-optical control of light at the nano-scale. This opens unprecedented opportunities for ultrafast nonlinear spectroscopy, sensing devices, on-chip optical frequency conversion, nonlinear optical metamaterials, and novel photon sources. Here, we review some of the recent advances in exploiting the potential of plasmonic antennas to realize robust nonlinear applications.

  3. Ultrafast nonlinear optics

    CERN Document Server

    Leburn, Christopher; Reid, Derryck

    2013-01-01

    The field of ultrafast nonlinear optics is broad and multidisciplinary, and encompasses areas concerned with both the generation and measurement of ultrashort pulses of light, as well as those concerned with the applications of such pulses. Ultrashort pulses are extreme events – both in terms of their durations, and also the high peak powers which their short durations can facilitate. These extreme properties make them powerful experiment tools. On one hand, their ultrashort durations facilitate the probing and manipulation of matter on incredibly short timescales. On the other, their ultrashort durations can facilitate high peak powers which can drive highly nonlinear light-matter interaction processes. Ultrafast Nonlinear Optics covers a complete range of topics, both applied and fundamental in nature, within the area of ultrafast nonlinear optics. Chapters 1 to 4 are concerned with the generation and measurement of ultrashort pulses. Chapters 5 to 7 are concerned with fundamental applications of ultrasho...

  4. Nonlinear Source Emulator

    DEFF Research Database (Denmark)

    Nguyen-Duy, Khiem

    and remains the prime source of energy in non-terrestrial applications such as those in sky-explorers. However, a renewable energy source is expensive, bulky, and its performance is weather dependent, which make testing of downstream converters very difficult. As a result, a nonlinear source emulator (NSE......) is a good solution to solve the problems associated with the use of real nonlinear sources in testing phases. However, a recent technical survey conducted during this work shows that most existing NSEs have only been concerned with simulating nonlinear systems in terrestrial applications. Furthermore......, their dynamic performance were not fast enough in order to imitate how a real nonlinear energy source would react under extreme conditions and operation modes. Particularly, a system in the sky can experience a step change of sunlight irradiation. Moreover, operation modes may include load step between nominal...

  5. Introduction to nonlinear science

    CERN Document Server

    Nicolis, G

    1995-01-01

    One of the most unexpected results in science in recent years is that quite ordinary systems obeying simple laws can give rise to complex, nonlinear or chaotic, behavior. In this book, the author presents a unified treatment of the concepts and tools needed to analyze nonlinear phenomena and to outline some representative applications drawn from the physical, engineering, and biological sciences. Some of the interesting topics covered include: dynamical systems with a finite number of degrees of freedom, linear stability analysis of fixed points, nonlinear behavior of fixed points, bifurcation analysis, spatially distributed systems, broken symmetries, pattern formation, and chaotic dynamics. The author makes a special effort to provide a logical connection between ordinary dynamical systems and spatially extended systems, and to balance the emphasis on chaotic behavior and more classical nonlinear behavior. He also develops a statistical approach to complex systems and compares it to traditional deterministi...

  6. Nonlinear magnetoinductive transmission lines

    CERN Document Server

    Lazarides, Nikos; Tsironis, G P

    2011-01-01

    Power transmission in one-dimensional nonlinear magnetic metamaterials driven at one end is investigated numerically and analytically in a wide frequency range. The nonlinear magnetic metamaterials are composed of varactor-loaded split-ring resonators which are coupled magnetically through their mutual inductances, forming thus a magnetoiductive transmission line. In the linear limit, significant power transmission along the array only appears for frequencies inside the linear magnetoinductive wave band. We present analytical, closed form solutions for the magnetoinductive waves transmitting the power in this regime, and their discrete frequency dispersion. When nonlinearity is important, more frequency bands with significant power transmission along the array may appear. In the equivalent circuit picture, the nonlinear magnetoiductive transmission line driven at one end by a relatively weak electromotive force, can be modeled by coupled resistive-inductive-capacitive (RLC) circuits with voltage-dependent cap...

  7. Optimization under Nonlinear Constraints

    OpenAIRE

    1982-01-01

    In this paper a timesaving method is proposed for maximizing likelihood functions when the parameter space is subject to nonlinear constraints, expressible as second order polynomials. The suggested approach is especially attractive when dealing with systems with many parameters.

  8. Nonlinearity in nanomechanical cantilevers

    DEFF Research Database (Denmark)

    Villanueva Torrijo, Luis Guillermo; Karabalin, R. B.; Matheny, M. H.

    2013-01-01

    Euler-Bernoulli beam theory is widely used to successfully predict the linear dynamics of micro-and nanocantilever beams. However, its capacity to characterize the nonlinear dynamics of these devices has not yet been rigorously assessed, despite its use in nanoelectromechanical systems development....... These findings underscore the delicate balance between inertial and geometric nonlinear effects in the fundamental mode, and strongly motivate further work to develop theories beyond the Euler-Bernoulli approximation. DOI: 10.1103/PhysRevB.87.024304....... In this article, we report the first highly controlled measurements of the nonlinear response of nanomechanical cantilevers using an ultralinear detection system. This is performed for an extensive range of devices to probe the validity of Euler-Bernoulli theory in the nonlinear regime. We find that its...

  9. Nonlinear Stokes Mueller Polarimetry

    CERN Document Server

    Samim, Masood; Barzda, Virginijus

    2015-01-01

    The Stokes Mueller polarimetry is generalized to include nonlinear optical processes such as second- and third-harmonic generation, sum- and difference-frequency generations. The overall algebraic form of the polarimetry is preserved, where the incoming and outgoing radiations are represented by column vectors and the intervening medium is represented by a matrix. Expressions for the generalized nonlinear Stokes vector and the Mueller matrix are provided in terms of coherency and correlation matrices, expanded by higher-dimensional analogues of Pauli matrices. In all cases, the outgoing radiation is represented by the conventional $4\\times 1$ Stokes vector, while dimensions of the incoming radiation Stokes vector and Mueller matrix depend on the order of the process being examined. In addition, relation between nonlinear susceptibilities and the measured Mueller matrices are explicitly provided. Finally, the approach of combining linear and nonlinear optical elements is discussed within the context of polarim...

  10. Adaptive and Nonlinear Control

    Science.gov (United States)

    1992-02-29

    in [22], we also applied the concept of zero dynamics to the problem of exact linearization of a nonlinear control system by dynamic feedback. Exact ...nonlinear systems, although it was well-known that the conditions for exact linearization are very stringent and consequently do not apply to a broad...29th IEEE Conference n Decision and Control, Invited Paper delivered by Dr. Gilliam. Exact Linearization of Zero Dynamics, 29th IEEE Conference on

  11. Nonlinear Optics and Turbulence

    Science.gov (United States)

    1992-10-01

    currently at Queen Mary College, London Patrick Dunne, (Ph.D., 1987, M.I.T., Hydrodynamic Stability, Nonlinear Waves), 1987-1988. Alecsander Dyachenko...U I I I U I I 3 9 3 V. BIOGRAPHIES A. FACULTY BRUCE BAYLY, 31, Ph.D. 1986, Princeton University. Postdoctoral visiting member 1986-88 at Courant...Caputo, A. C. Newell, and M. Shelley , "Nonlinear Wave Propagation Through a Random Medium and Soliton Tunneling", Integrable Systems and

  12. Explaining (Missing) Regulator Paradigm Shifts

    DEFF Research Database (Denmark)

    Wigger, Angela; Buch-Hansen, Hubert

    2014-01-01

    of competition regulation is heaving into sight. It sets out to explain this from the vantage point of a critical political economy perspective, which identifies the circumstances under which a crisis can result in a regulatory paradigm shift. Contrasting the current situation with the shift in EC/EU competition...... capitalism; the social power configuration underpinning the neoliberal order remains unaltered; no clear counter-project has surfaced; the European Commission has been (and remains) in a position to oppose radical changes; and finally, there are no signs of a wider paradigm shift in the EU's regulatory...

  13. Robust Nonlinear Neural Codes

    Science.gov (United States)

    Yang, Qianli; Pitkow, Xaq

    2015-03-01

    Most interesting natural sensory stimuli are encoded in the brain in a form that can only be decoded nonlinearly. But despite being a core function of the brain, nonlinear population codes are rarely studied and poorly understood. Interestingly, the few existing models of nonlinear codes are inconsistent with known architectural features of the brain. In particular, these codes have information content that scales with the size of the cortical population, even if that violates the data processing inequality by exceeding the amount of information entering the sensory system. Here we provide a valid theory of nonlinear population codes by generalizing recent work on information-limiting correlations in linear population codes. Although these generalized, nonlinear information-limiting correlations bound the performance of any decoder, they also make decoding more robust to suboptimal computation, allowing many suboptimal decoders to achieve nearly the same efficiency as an optimal decoder. Although these correlations are extremely difficult to measure directly, particularly for nonlinear codes, we provide a simple, practical test by which one can use choice-related activity in small populations of neurons to determine whether decoding is suboptimal or optimal and limited by correlated noise. We conclude by describing an example computation in the vestibular system where this theory applies. QY and XP was supported by a grant from the McNair foundation.

  14. Nonlinear systems in medicine.

    Science.gov (United States)

    Higgins, John P

    2002-01-01

    Many achievements in medicine have come from applying linear theory to problems. Most current methods of data analysis use linear models, which are based on proportionality between two variables and/or relationships described by linear differential equations. However, nonlinear behavior commonly occurs within human systems due to their complex dynamic nature; this cannot be described adequately by linear models. Nonlinear thinking has grown among physiologists and physicians over the past century, and non-linear system theories are beginning to be applied to assist in interpreting, explaining, and predicting biological phenomena. Chaos theory describes elements manifesting behavior that is extremely sensitive to initial conditions, does not repeat itself and yet is deterministic. Complexity theory goes one step beyond chaos and is attempting to explain complex behavior that emerges within dynamic nonlinear systems. Nonlinear modeling still has not been able to explain all of the complexity present in human systems, and further models still need to be refined and developed. However, nonlinear modeling is helping to explain some system behaviors that linear systems cannot and thus will augment our understanding of the nature of complex dynamic systems within the human body in health and in disease states.

  15. Goos-Hänchen shift in a standing-wave-coupled electromagnetically-induced-transparency medium

    Science.gov (United States)

    Zhang, Xiao-Jun; Wang, Hai-Hua; Liang, Zhi-Peng; Xu, Yan; Fan, Cun-Bo; Liu, Cheng-Zhi; Gao, Jin-Yue

    2015-03-01

    The Goos-Hänchen shift of the system composed by two cavity walls and an intracavity atomic sample is presented. The atomic sample is treated as a four-level double-Λ system, driven by the two counterpropagating coupling fields. The probe field experiences the discontinuous refractive index variation and is reflected. Moreover, under the phase-matching condition, the four-wave mixing effect based on electromagnetically induced transparency can cause effective reflection. The Goos-Hänchen shifts appear in both situations and are carefully investigated in this article. We refer to the first one with the incident and reflected light having identical wavelength as the linear Goos-Hänchen shift, and the second one with the reflection wavelength determined by the phase-matching condition as the nonlinear Goos-Hänchen shift. The differences between the two kinds of shifts, such as the incident angle range, conditions for the shift peaks, and controllability, are discussed.

  16. NONLINEAR DYNAMIC INSTABILITY OF DOUBLE-WALLED CARBON NANOTUBES UNDER PERIODIC EXCITATION

    Institute of Scientific and Technical Information of China (English)

    Yiming Fu; Rengui Bi; Pu Zhang

    2009-01-01

    A multiple-elastic beam model based on Euler-Bernoulli-beam theory is presented to investigate the nonlinear dynamic instability of double-walled nanotubes. Taking the geometric nonlinearity of structure deformation, the effects of van der Waais forces as well as the non-coaxial curvature of each nested tube into account, the nonlinear parametric vibration governing equations are derived. Numerical results indicate that the double-walled nanotube (DWNT) can be considered as a single column when the van der Waals forces are sufficiently strong. The stiffness of medium could substantially reduce the area of the nonlinear dynamic instability region, in particular, the geometric nonlinearity can be out of account when the stiffness is large enough. The area of the principal nonlinear instability region and its shifting distance aroused by the nonlinearity both decrease with the increment of the aspect ratio of the nanotubes.

  17. Modifying the optical path in a nonlinear double-slit experiment

    CERN Document Server

    Paltoglou, Vassilis

    2015-01-01

    In this letter, we study a nonlinear interferometric setup based on diffraction rather than beam combining. It consists of a nonlinear analogue of Young's double-slit experiment where a nonlinear material is placed exactly after one of the slits. The presence of nonlinearity breaks the transverse spatial symmetry of the system and thus modifies the optical path. For moderate nonlinearities this leads to a self-induced shift of the intensity pattern in the transverse plane. A simple theoretical model is developed which is surprisingly accurate in predicting the intensity profile of the main lobes for a wide range of parameters. We discuss about possible applications of our model in nonlinear interferometry, for example in measuring the nonlinearities of optical materials.

  18. Handbook of nonlinear optical crystals

    CERN Document Server

    Dmitriev, Valentin G; Nikogosyan, David N

    1991-01-01

    This Handbook of Nonlinear Optical Crystals provides a complete description of the properties and applications of nonlinear crystals In addition, it presents the most important equations for calculating the main parameters of nonlinear frequency converters This comprehensive reference work will be of great value to all scientists and engineers working in nonlinear optics, quantum electronics and laser physics

  19. Optimal beamforming in MIMO systems with HPA nonlinearity

    KAUST Repository

    Qi, Jian

    2010-09-01

    In this paper, multiple-input multiple-output (MIMO) transmit beamforming (TB) systems under the consideration of nonlinear high-power amplifiers (HPAs) are investigated. The optimal beamforming scheme, with the optimal beamforming weight vector and combining vector, is proposed for MIMO systems with HPA nonlinearity. The performance of the proposed MIMO beamforming scheme in the presence of HPA nonlinearity is evaluated in terms of average symbol error probability (SEP), outage probability and system capacity, considering transmission over uncorrelated quasi-static frequency-flat Rayleigh fading channels. Numerical results are provided and show the effects of several system parameters, namely, parameters of nonlinear HPA, numbers of transmit and receive antennas, and modulation order of phase-shift keying (PSK), on performance. ©2010 IEEE.

  20. Seasonal and ontogenetic variations in the diet of Cichla kelberi Kullander and Ferreira, 2006 introduced in an artificial lake in southeastern Brazil

    Directory of Open Access Journals (Sweden)

    LM Gomiero

    Full Text Available The diet of Cichla kelberi introduced in an artificial lake in Leme-SP was predominantly composed of common fish species (Oreochromis niloticus and C. kelberi. In the spring and summer, the most consumed item was O. niloticus. However, cannibalism was very common for this species. The high frequencies of O. niloticus and C. kelberi reveal that this species is adapted to a seasonal cycle, feeding on the most common prey in each period of the year, with a reduction of foraging activity during the winter. The diets were different among the immature and mature individuals suggesting that there are ontogenetic differences, mainly related to prey type, such as: Ephemeroptera consumed by the immature peacock bass and fish by the mature ones, besides the size of the prey.

  1. Nonlinear Approaches in Engineering Applications

    CERN Document Server

    Jazar, Reza

    2012-01-01

    Nonlinear Approaches in Engineering Applications focuses on nonlinear phenomena that are common in the engineering field. The nonlinear approaches described in this book provide a sound theoretical base and practical tools to design and analyze engineering systems with high efficiency and accuracy and with less energy and downtime. Presented here are nonlinear approaches in areas such as dynamic systems, optimal control and approaches in nonlinear dynamics and acoustics. Coverage encompasses a wide range of applications and fields including mathematical modeling and nonlinear behavior as applied to microresonators, nanotechnologies, nonlinear behavior in soil erosion,nonlinear population dynamics, and optimization in reducing vibration and noise as well as vibration in triple-walled carbon nanotubes. This book also: Provides a complete introduction to nonlinear behavior of systems and the advantages of nonlinearity as a tool for solving engineering problems Includes applications and examples drawn from the el...

  2. VP Anaphors and Object Shift

    DEFF Research Database (Denmark)

    Ørsnes, Bjarne

    2013-01-01

    The article discusses the placement of the VP anaphor det ‘it’ as a complement of verbs selecting VP complements in Danish. With verbs that only allow a VP complement, the VP anaphor must be in SpecCP regardless of its information structure properties. If SpecCP is occupied by an operator, the an...... be in situ. The article argues that a shifted pronominal in Danish must be categorially licensed by the verb and extends this analysis to shifting locatives. An Optimality Theory analysis is proposed that accounts for the observed facts......., the anaphor can be in situ, but it cannot shift. With verbs that allow its VP complement to alternate with an NP complement, the VP anaphor can be in SpecCP, shifted or in situ according to the information structural properties of the anaphor. Only if SpecCP is occupied by an operator, must a topical anaphor...

  3. Shift Work: Improving Daytime Sleep

    Science.gov (United States)

    ... sleeping during the day. Do you have any sleep tips for shift workers? Answers from Timothy Morgenthaler, ... to be awake during the day and to sleep at night. Good daytime sleep is possible, though, ...

  4. Goos-Hänchen shift.

    Science.gov (United States)

    Snyder, A W; Love, J D

    1976-01-01

    An extremely simple derivation of the Goos-Hänchen shift is presented for total internal reflection at a plane interface between two semiinfinite dielectric media, as well as for optical waveguides of plane arid circular cross section. The derivation is based on energy considerations, requires knowledge of Fresnel's equation only, and shows explicitly that the shift is due to the flow of energy across the dielectric boundary.

  5. Ontogenetic improvement of visual function in the medaka Oryzias latipes based on an optomotor testing system for larval and adult fish

    Science.gov (United States)

    Carvalho, Paulo S. M.; Noltie, Douglas B.; Tillitt, D.E.

    2002-01-01

    We developed a system for evaluation of visual function in larval and adult fish. Both optomotor (swimming) and optokinetic (eye movement) responses were monitored and recorded using a system of rotating stripes. The system allowed manipulation of factors such as width of the stripes used, rotation speed of the striped drum, and light illuminance levels within both the scotopic and photopic ranges. Precise control of these factors allowed quantitative measurements of visual acuity and motion detection. Using this apparatus, we tested the hypothesis that significant posthatch ontogenetic improvements in visual function occur in the medaka Oryzias latipes, and also that this species shows significant in ovo neuronal development. Significant improvements in the acuity angle alpha (ability to discriminate detail) were observed from approximately 5 degrees at hatch to 1 degree in the oldest adult stages. In addition, we measured a significant improvement in flicker fusion thresholds (motion detection skills) between larval and adult life stages within both the scotopic and photopic ranges of light illuminance. Ranges of flicker fusion thresholds (X?? ?? SD) at log I=1.96 (photopic) varied from 37.2 ?? 1.6 cycles/s in young adults to 18.6 ?? 1.6 cycles/s in young larvae 10 days posthatch. At log I= - 2.54 (scotopic), flicker fusion thresholds varied from 5.8 ?? 0.7 cycles/s in young adults to 1.7 ?? 0.4 cycles/s in young larvae 10 days posthatch. Light sensitivity increased approximately 2.9 log units from early hatched larval stages to adults. The demonstrated ontogenetic improvements in visual function probably enable the fish to explore new resources, thereby enlarging their fundamental niche. ?? 2002 The Association for the Study of Animal Behaviour. Published by Elsevier Science Ltd. All rights reserved.

  6. Numerical investigation of amplitude-dependent dynamic response in acoustic metamaterials with nonlinear oscillators.

    Science.gov (United States)

    Manimala, James M; Sun, C T

    2016-06-01

    The amplitude-dependent dynamic response in acoustic metamaterials having nonlinear local oscillator microstructures is studied using numerical simulations on representative discrete mass-spring models. Both cubically nonlinear hardening and softening local oscillator cases are considered. Single frequency, bi-frequency, and wave packet excitations at low and high amplitude levels were used to interrogate the models. The propagation and attenuation characteristics of harmonic waves in a tunable frequency range is found to correspond to the amplitude and nonlinearity-dependent shifts in the local resonance bandgap for such nonlinear acoustic metamaterials. A predominant shift in the propagated wave spectrum towards lower frequencies is observed. Moreover, the feasibility of amplitude and frequency-dependent selective filtering of composite signals consisting of individual frequency components which fall within propagating or attenuating regimes is demonstrated. Further enrichment of these wave manipulation mechanisms in acoustic metamaterials using different combinations of nonlinear microstructures presents device implications for acoustic filters and waveguides.

  7. Uncertainty propagation for nonlinear vibrations: A non-intrusive approach

    Science.gov (United States)

    Panunzio, A. M.; Salles, Loic; Schwingshackl, C. W.

    2017-02-01

    The propagation of uncertain input parameters in a linear dynamic analysis is reasonably well established today, but with the focus of the dynamic analysis shifting towards nonlinear systems, new approaches is required to compute the uncertain nonlinear responses. A combination of stochastic methods (Polynomial Chaos Expansion, PCE) with an Asymptotic Numerical Method (ANM) for the solution of the nonlinear dynamic systems is presented to predict the propagation of random input uncertainties and assess their influence on the nonlinear vibrational behaviour of a system. The proposed method allows the computation of stochastic resonance frequencies and peak amplitudes based on multiple input uncertainties, leading to a series of uncertain nonlinear dynamic responses. One of the main challenges when using the PCE is thereby the Gibbs phenomenon, which can heavily impact the resulting stochastic nonlinear response by introducing spurious oscillations. A novel technique to avoid the Gibbs phenomenon is be presented in this paper, leading to high quality frequency response predictions. A comparison of the proposed stochastic nonlinear analysis technique to traditional Monte Carlo simulations, demonstrates comparable accuracy at a significantly reduced computational cost, thereby validating the proposed approach.

  8. Dynamic two state stochastic models for ecological regime shifts

    DEFF Research Database (Denmark)

    Møller, Jan Kloppenborg; Carstensen, Niels Jacob; Madsen, Henrik

    2009-01-01

    A simple non-linear stochastic two state, discrete-time model is presented. The interaction between benthic and pelagic vegetation in aquatic ecosystems subject to changing external nutrient loading is described by the nonlinear functions. The dynamical behavior of the deterministic part...... of regimes, depending on how the noise propagates through the system. The dynamical properties of a system should therefore be described through propagation of the state distributions rather than the state means and consequently, stochastic models should be compared in a probabilistic framework....... of the model illustrates that hysteresis effect and regime shifts can be obtained for a limited range of parameter values only. The effect of multiplicative noise components entering at different levels of the model is presented and discussed. Including noise leads to very different results on the stability...

  9. Nonlinear pulsation masses

    Energy Technology Data Exchange (ETDEWEB)

    Davis, C.G.

    1990-01-01

    The advent of nonlinear pulsation theory really coincides with the development of the large computers after the second world war. Christy and Stobbie were the first to make use of finite difference techniques on computers to model the bumps'' observed in the classical Cepheid light and velocity curves, the so-called Hertzsprung'' sequence. Following this work a more sophisticated analysis of the light and velocity curves from the models was made by Simon and Davis using Fourier techniques. Recently a simpler amplitude equation formalism has been developed that helps explain this resonance mechanism. The determination of Population I Cepheid masses by nonlinear methods will be discussed. For the lower mass objects, such as RR Lyrae and BL Her. stars, we find general agreement using evolutionary masses and nonlinear pulsation theory. An apparent difficulty of nonlinear pulsation theory occurs in the understanding of double'' mode pulsation, which will also be discussed. Recent studies in nonlinear pulsation theory have dealt with the question of mode selection, period doubling and the trends towards chaotic behavior such as is observed in the transition from W Virginis to RV Tauri-like stars. 10 refs., 1 fig., 2 tabs.

  10. Nonlinear elastic waves in materials

    CERN Document Server

    Rushchitsky, Jeremiah J

    2014-01-01

    The main goal of the book is a coherent treatment of the theory of propagation in materials of nonlinearly elastic waves of displacements, which corresponds to one modern line of development of the nonlinear theory of elastic waves. The book is divided on five basic parts: the necessary information on waves and materials; the necessary information on nonlinear theory of elasticity and elastic materials; analysis of one-dimensional nonlinear elastic waves of displacement – longitudinal, vertically and horizontally polarized transverse plane nonlinear elastic waves of displacement; analysis of one-dimensional nonlinear elastic waves of displacement – cylindrical and torsional nonlinear elastic waves of displacement; analysis of two-dimensional nonlinear elastic waves of displacement – Rayleigh and Love nonlinear elastic surface waves. The book is addressed first of all to people working in solid mechanics – from the students at an advanced undergraduate and graduate level to the scientists, professional...

  11. Nonlinear Dynamic Force Spectroscopy

    CERN Document Server

    Björnham, Oscar

    2016-01-01

    Dynamic force spectroscopy (DFS) is an experimental technique that is commonly used to assess information of the strength, energy landscape, and lifetime of noncovalent bio-molecular interactions. DFS traditionally requires an applied force that increases linearly with time so that the bio-complex under investigation is exposed to a constant loading rate. However, tethers or polymers can modulate the applied force in a nonlinear regime. For example, bacterial adhesion pili and polymers with worm-like chain properties are examples of structures that show nonlinear force responses. In these situations, the theory for traditional DFS cannot be readily applied. In this work we expand the theory for DFS to also include nonlinear external forces while still maintaining compatibility with the linear DFS theory. To validate the theory we modeled a bio-complex expressed on a stiff, an elastic and a worm-like chain polymer, using Monte Carlo methods, and assessed the corresponding rupture force spectra. It was found th...

  12. Nonlinear optomechanical paddle nanocavities

    CERN Document Server

    Kaviani, Hamidreza; Wu, Marcelo; Ghobadi, Roohollah; Barclay, Paul E

    2014-01-01

    A photonic crystal optomechanical system combining strong nonlinear optomechanical coupling, low effective mass and large optical mode spacing is introduced. This nanoscale "paddle nanocavity" device supports mechanical resonances with effective mass of 300--600 fg which couple nonlinearly to co-localized optical modes with a quadratic optomechanical coupling coefficient $g^{(2)} > 2\\pi\\times400$ MHz/nm$^2$, and a two phonon to single photon optomechanical coupling rate $\\Delta \\omega_0 > 2\\pi\\times 16$ Hz. This coupling relies on strong phonon-photon interactions in a structure whose optical mode spectrum is highly non--degenerate. Simulations indicate that nonlinear optomechanical readout of thermally driven motion in these devices should be observable for T $> 50 $ mK, and that measurement of phonon shot noise is achievable.

  13. Nonlinear dynamics of structures

    CERN Document Server

    Oller, Sergio

    2014-01-01

    This book lays the foundation of knowledge that will allow a better understanding of nonlinear phenomena that occur in structural dynamics.   This work is intended for graduate engineering students who want to expand their knowledge on the dynamic behavior of structures, specifically in the nonlinear field, by presenting the basis of dynamic balance in non‐linear behavior structures due to the material and kinematics mechanical effects.   Particularly, this publication shows the solution of the equation of dynamic equilibrium for structure with nonlinear time‐independent materials (plasticity, damage and frequencies evolution), as well as those time dependent non‐linear behavior materials (viscoelasticity and viscoplasticity). The convergence conditions for the non‐linear dynamic structure solution  are studied, and the theoretical concepts and its programming algorithms are presented.  

  14. Nonlinear Photonic Crystal Fibers

    DEFF Research Database (Denmark)

    Hansen, Kim Per

    2004-01-01

    , leading to reduced mode confinement and dispersion flexibility. In this thesis, we treat the nonlinear photonic crystal fiber – a special sub-class of photonic crystal fibers, the core of which has a diameter comparable to the wavelength of the light guided in the fiber. The small core results in a large...... nonlinear coefficient and in various applications, it is therefore possible to reduce the required fiber lengths quite dramatically, leading to increased stability and efficiency. Furthermore, it is possible to design these fibers with zero-dispersion at previously unreachable wavelengths, paving the way...... for completely new applications, especially in and near the visible wavelength region. One such application is supercontinuum generation. Supercontinuum generation is extreme broadening of pulses in a nonlinear medium (in this case a small-core fiber), and depending on the dispersion of the fiber, it is possible...

  15. Linearizing nonlinear optics

    CERN Document Server

    Schmidt, Bruno E; Ernotte, Guilmot; Clerici, Matteo; Morandotti, Roberto; Ibrahim, Heide; Legare, Francois

    2016-01-01

    In the framework of linear optics, light fields do not interact with each other in a medium. Yet, when their field amplitude becomes comparable to the electron binding energies of matter, the nonlinear motion of these electrons emits new dipole radiation whose amplitude, frequency and phase differ from the incoming fields. Such high fields are typically achieved with ultra-short, femtosecond (1fs = 10-15 sec.) laser pulses containing very broad frequency spectra. Here, the matter not only couples incoming and outgoing fields but also causes different spectral components to interact and mix through a convolution process. In this contribution, we describe how frequency domain nonlinear optics overcomes the shortcomings arising from this convolution in conventional time domain nonlinear optics1. We generate light fields with previously inaccessible properties because the uncontrolled coupling of amplitudes and phases is turned off. For example, arbitrary phase functions are transferred linearly to the second har...

  16. Nonlinear optomechanics with graphene

    Science.gov (United States)

    Shaffer, Airlia; Patil, Yogesh Sharad; Cheung, Hil F. H.; Wang, Ke; Vengalattore, Mukund

    2016-05-01

    To date, studies of cavity optomechanics have been limited to exploiting the linear interactions between the light and mechanics. However, investigations of quantum signal transduction, quantum enhanced metrology and manybody physics with optomechanics each require strong, nonlinear interactions. Graphene nanomembranes are an exciting prospect for realizing such studies due to their inherently nonlinear nature and low mass. We fabricate large graphene nanomembranes and study their mechanical and optical properties. By using dark ground imaging techniques, we correlate their eigenmode shapes with the measured dissipation. We study their hysteretic response present even at low driving amplitudes, and their nonlinear dissipation. Finally, we discuss ongoing efforts to use these resonators for studies of quantum optomechanics and force sensing. This work is supported by the DARPA QuASAR program through a Grant from the ARO.

  17. Nonlinear Analysis of Buckling

    Directory of Open Access Journals (Sweden)

    Psotný Martin

    2014-06-01

    Full Text Available The stability analysis of slender web loaded in compression was presented. To solve this problem, a specialized computer program based on FEM was created. The nonlinear finite element method equations were derived from the variational principle of minimum of potential energy. To obtain the nonlinear equilibrium paths, the Newton-Raphson iteration algorithm was used. Corresponding levels of the total potential energy were defined. The peculiarities of the effects of the initial imperfections were investigated. Special attention was focused on the influence of imperfections on the post-critical buckling mode. The stable and unstable paths of the nonlinear solution were separated. Obtained results were compared with those gained using ANSYS system.

  18. Nonlinear Metamaterials for Holography

    CERN Document Server

    Almeida, Euclides; Prior, Yehiam

    2015-01-01

    A hologram is an optical element storing phase and possibly amplitude information enabling the reconstruction of a three dimensional image of an object by illumination and scattering of a coherent beam of light, and the image is generated at the same wavelength as the input laser beam. In recent years it was shown that information can be stored in nanometric antennas giving rise to ultrathin components. Here we demonstrate nonlinear multi-layer metamaterial holograms where by the nonlinear process of Third Harmonic Generation, a background free image is formed at a new frequency which is the third harmonic of the illuminating beam. Using e-beam lithography of multilayer plasmonic nanoantennas, we fabricate polarization-sensitive nonlinear elements such as blazed gratings, lenses and other computer-generated holograms. These holograms are analyzed and prospects for future device applications are discussed.

  19. Multidimensional nonlinear descriptive analysis

    CERN Document Server

    Nishisato, Shizuhiko

    2006-01-01

    Quantification of categorical, or non-numerical, data is a problem that scientists face across a wide range of disciplines. Exploring data analysis in various areas of research, such as the social sciences and biology, Multidimensional Nonlinear Descriptive Analysis presents methods for analyzing categorical data that are not necessarily sampled randomly from a normal population and often involve nonlinear relations. This reference not only provides an overview of multidimensional nonlinear descriptive analysis (MUNDA) of discrete data, it also offers new results in a variety of fields. The first part of the book covers conceptual and technical preliminaries needed to understand the data analysis in subsequent chapters. The next two parts contain applications of MUNDA to diverse data types, with each chapter devoted to one type of categorical data, a brief historical comment, and basic skills peculiar to the data types. The final part examines several problems and then concludes with suggestions for futu...

  20. Nonlinear dispersion effects in elastic plates: numerical modelling and validation

    Science.gov (United States)

    Kijanka, Piotr; Radecki, Rafal; Packo, Pawel; Staszewski, Wieslaw J.; Uhl, Tadeusz; Leamy, Michael J.

    2017-04-01

    Nonlinear features of elastic wave propagation have attracted significant attention recently. The particular interest herein relates to complex wave-structure interactions, which provide potential new opportunities for feature discovery and identification in a variety of applications. Due to significant complexity associated with wave propagation in nonlinear media, numerical modeling and simulations are employed to facilitate design and development of new measurement, monitoring and characterization systems. However, since very high spatio- temporal accuracy of numerical models is required, it is critical to evaluate their spectral properties and tune discretization parameters for compromise between accuracy and calculation time. Moreover, nonlinearities in structures give rise to various effects that are not present in linear systems, e.g. wave-wave interactions, higher harmonics generation, synchronism and | recently reported | shifts to dispersion characteristics. This paper discusses local computational model based on a new HYBRID approach for wave propagation in nonlinear media. The proposed approach combines advantages of the Local Interaction Simulation Approach (LISA) and Cellular Automata for Elastodynamics (CAFE). The methods are investigated in the context of their accuracy for predicting nonlinear wavefields, in particular shifts to dispersion characteristics for finite amplitude waves and secondary wavefields. The results are validated against Finite Element (FE) calculations for guided waves in copper plate. Critical modes i.e., modes determining accuracy of a model at given excitation frequency - are identified and guidelines for numerical model parameters are proposed.

  1. Early-warning signals (potentially) reduce uncertainty in forecasted timing of critical shifts in vegetation systems

    NARCIS (Netherlands)

    Karssenberg, D.J.; Bierkens, M.F.P.

    2011-01-01

    Non-linear systems may undergo abrupt transitions as a result of a gradual change in system drivers. Such regime shifts, or critical transitions, have been described for a large range of spatio-temporal systems, including those studied in the social sciences, ecology, geology, and climatology. Regim

  2. Nonlinear airship aeroelasticity

    Science.gov (United States)

    Bessert, N.; Frederich, O.

    2005-12-01

    The aeroelastic derivatives for today's aircraft are calculated in the concept phase using a standard procedure. This scheme has to be extended for large airships, due to various nonlinearities in structural and aerodynamic behaviour. In general, the structural model of an airship is physically as well as geometrically nonlinear. The main sources of nonlinearity are large deformations and the nonlinear material behaviour of membranes. The aerodynamic solution is also included in the nonlinear problem, because the deformed airship influences the surrounding flow. Due to these nonlinearities, the aeroelastic problem for airships can only be solved by an iterative procedure. As one possibility, the coupled aerodynamic and structural dynamic problem was handled using linked standard solvers. On the structural side, the Finite-Element program package ABAQUS was extended with an interface to the aerodynamic solver VSAERO. VSAERO is based on the aerodynamic panel method using potential flow theory. The equilibrium of the internal structural and the external aerodynamic forces leads to the structural response and a trimmed flight state for the specified flight conditions (e.g. speed, altitude). The application of small perturbations around a trimmed state produces reaction forces and moments. These constraint forces are then transferred into translational and rotational acceleration fields by performing an inertia relief analysis of the disturbed structural model. The change between the trimmed flight state and the disturbed one yields the respective aeroelastic derivatives. By including the calculated derivatives in the linearised equation of motion system, it is possible to judge the stability and controllability of the investigated airship.

  3. Knight shifts around vacancies in the 2D Heisenberg model.

    Science.gov (United States)

    Anfuso, Fabrizio; Eggert, Sebastian

    2006-01-13

    The local response to a uniform field around vacancies in the two-dimensional spin-1/2 Heisenberg antiferromagnet is determined by numerical quantum Monte Carlo simulations as a function of temperature. It is possible to separate the Knight shifts into uniform and staggered contributions on the lattice which are analyzed and understood in detail. The contributions show interesting long- and short-range behavior that may be of relevance in NMR and susceptibility measurements. For more than one impurity, remarkable nonlinear enhancement and cancellation effects take place. We predict that the Curie impurity susceptibility will be observable for a random impurity concentration even in the thermodynamic limit.

  4. Contrast Enhancement of Radiographs Using Shift Invariant Wavelet Transform

    Institute of Scientific and Technical Information of China (English)

    2000-01-01

    A novel approach using shift invariant wavelet transform is presented for the contrast enhancement of radiographs. By exploiting cross-scale correlation among wavelet coefficients, edge information of radiographic images is extracted and protected, while noise is smoothed out in the wavelet domain. Radiographs are then reconstructed from the transform coefficients modified at multi-scales by nonlinear enhancement operator. The method can achieve effectively contrast enhancement and edge-preserved denoising simultaneously, yet it is capable of giving visually distinct images and offering considerable benefits in medical diagnosis.

  5. Damping and Frequency Shift of Large Amplitude Electron Plasma Waves

    DEFF Research Database (Denmark)

    Thomsen, Kenneth; Juul Rasmussen, Jens

    1983-01-01

    The initial evolution of large-amplitude one-dimensional electron waves is investigated by applying a numerical simulation. The initial wave damping is found to be strongly enhanced relative to the linear damping and it increases with increasing amplitude. The temporal evolution of the nonlinear...... damping rate γ(t) shows that it increases with time within the initial phase of propagation, t≲π/ωB (ωB is the bounce frequency), whereafter it decreases and changes sign implying a regrowth of the wave. The shift in the wave frequency δω is observed to be positive for t≲π/ωB; then δω changes sign...

  6. Nonlinear fiber optics

    CERN Document Server

    Agrawal, Govind

    2012-01-01

    Since the 4e appeared, a fast evolution of the field has occurred. The 5e of this classic work provides an up-to-date account of the nonlinear phenomena occurring inside optical fibers, the basis of all our telecommunications infastructure as well as being used in the medical field. Reflecting the big developments in research, this new edition includes major new content: slow light effects, which offers a reduction in noise and power consumption and more ordered network traffic-stimulated Brillouin scattering; vectorial treatment of highly nonlinear fibers; and a brand new chapter o

  7. Limits to Nonlinear Inversion

    DEFF Research Database (Denmark)

    Mosegaard, Klaus

    2012-01-01

    For non-linear inverse problems, the mathematical structure of the mapping from model parameters to data is usually unknown or partly unknown. Absence of information about the mathematical structure of this function prevents us from presenting an analytical solution, so our solution depends on our......-heuristics are inefficient for large-scale, non-linear inverse problems, and that the 'no-free-lunch' theorem holds. We discuss typical objections to the relevance of this theorem. A consequence of the no-free-lunch theorem is that algorithms adapted to the mathematical structure of the problem perform more efficiently than...

  8. Fundamentals of nonlinear optics

    CERN Document Server

    Powers, Peter E

    2011-01-01

    Peter Powers's rigorous but simple description of a difficult field keeps the reader's attention throughout. … All chapters contain a list of references and large numbers of practice examples to be worked through. … By carefully working through the proposed problems, students will develop a sound understanding of the fundamental principles and applications. … the book serves perfectly for an introductory-level course for second- and third-order nonlinear optical phenomena. The author's writing style is refreshing and original. I expect that Fundamentals of Nonlinear Optics will fast become pop

  9. Tunable nonlinear graphene metasurfaces

    CERN Document Server

    Smirnova, Daria A; Kivshar, Yuri S; Khanikaev, Alexander B

    2015-01-01

    We introduce the concept of nonlinear graphene metasurfaces employing the controllable interaction between a graphene layer and a planar metamaterial. Such hybrid metasurfaces support two types of subradiant resonant modes, asymmetric modes of structured metamaterial elements ("metamolecules") and graphene plasmons exhibiting strong mutual coupling and avoided dispersion crossing. High tunability of graphene plasmons facilitates strong interaction between the subradiant modes, modifying the spectral position and lifetime of the associated Fano resonances. We demonstrate that strong resonant interaction, combined with the subwavelength localization of plasmons, leads to the enhanced nonlinear response and high efficiency of the second-harmonic generation.

  10. La dieta y la fauna de parásitos metazoos del torito Bovichthys chilensis Regan 1914 (Pisces: Bovichthydae en la costa de Chile centro-sur: variaciones geográficas y ontogenéticas Diet and metazoan parasite fauna of the thornfish Bovichthys chilensis Regan 1914 (Pisces: Bovichthydae on the coast of central-south Chile: geographical and ontogenetic variations

    Directory of Open Access Journals (Sweden)

    GABRIELA MUÑOZ

    2002-12-01

    variación geográfica del parasitismo de los toritos adultos, para dilucidar si en esta especie de hospedador el eje ontogenético es o no más importante que el geográfico para explicar las variaciones de la fauna parasitaria.Knowledge on what, how much, when and where hosts eat and live should help to understand the changes that occur in parasite communities because transmission of endoparasites is tightly linked to diet composition as well as ectoparasites are to the use that hosts make of the habitat. Thus, geographical and ontogenetic variations in diet and parasite fauna of the thornfish Bovichthys chilensis are described, and discussed taking into account the ontogenetic shifts in this host's niche. A total 122 individuals: 108 juveniles collected from the intertidal zone at four localities along the south-central coast of Chile (between 33º and 40º S, and 14 adults collected from the shallow subtidal at a fifth locality (36º S, were examined for food contents and parasite infracommunities. Food items were encountered in almost 70 % of fish examined with 25 prey items being recognized. Only one of the prey items was found in both juvenile and adult fish. The diet of juvenile fish consisted mainly of amphipods and while that of adults was mainly composed of crustacean decapods. Almost 40 % of fish examined harbored parasites with a total of 624 individuals and belonging to 16 taxa being identified, four of which were shared between juvenile and adult fish. Prevalence, total intensity, richness, and diversity of parasites was similarly low among juvenile fish sampled at four different localities, while significant differences in total prevalence, dietary and parasite infracommunity composition among localities were found. Absence of a clear relationship among juvenile fish between the composition of the diet and of infracommunities may be due to the fact that parasitoses are necessarily recent. Adult hosts had infracommunities with greater total prevalence

  11. High-accuracy acoustic detection of nonclassical component of material nonlinearity.

    Science.gov (United States)

    Haupert, Sylvain; Renaud, Guillaume; Rivière, Jacques; Talmant, Maryline; Johnson, Paul A; Laugier, Pascal

    2011-11-01

    The aim is to assess the nonclassical component of material nonlinearity in several classes of materials with weak, intermediate, and high nonlinear properties. In this contribution, an optimized nonlinear resonant ultrasound spectroscopy (NRUS) measuring and data processing protocol applied to small samples is described. The protocol is used to overcome the effects of environmental condition changes that take place during an experiment, and that may mask the intrinsic nonlinearity. External temperature fluctuation is identified as a primary source of measurement contamination. For instance, a variation of 0.1 °C produced a frequency variation of 0.01%, which is similar to the expected nonlinear frequency shift for weakly nonlinear materials. In order to overcome environmental effects, the reference frequency measurements are repeated before each excitation level and then used to compute nonlinear parameters. Using this approach, relative resonant frequency shifts of 10(-5) can be measured, which is below the limit of 10(-4) often considered as the limit of NRUS sensitivity under common experimental conditions. Due to enhanced sensitivity resulting from the correction procedure applied in this work, nonclassical nonlinearity in materials that before have been assumed to only be classically nonlinear in past work (steel, brass, and aluminum) is reported.

  12. Nonlinear effects in optical fibers

    CERN Document Server

    Ferreira, Mario F

    2011-01-01

    Cutting-edge coverage of nonlinear phenomena occurring inside optical fibers Nonlinear fiber optics is a specialized part of fiber optics dealing with optical nonlinearities and their applications. As fiber-optic communication systems have become more advanced and complex, the nonlinear effects in optical fibers have increased in importance, as they adversely affect system performance. Paradoxically, the same nonlinear phenomena also offer the promise of addressing the bandwidth bottleneck for signal processing for future ultra-high speed optical networks. Nonlinear Effects in Optical Fiber

  13. Does the ARFIMA really shift?

    DEFF Research Database (Denmark)

    Monache, Davide Delle; Grassi, Stefano; Santucci de Magistris, Paolo

    Short memory models contaminated by level shifts have long-memory features similar to those associated to processes generated under fractional integration. In this paper, we propose a robust testing procedure, based on an encompassing parametric specification, that allows to disentangle the level...... the highest power compared to other existing tests for spurious long-memory. Finally, we illustrate the usefulness of the proposed approach on the daily series of bipower variation and share turnover and on the monthly inflation series of G7 countries....... shift term from the ARFIMA component. The estimation is carried out via a state-space methodology and it leads to a robust estimate of the fractional integration parameter also in presence of level shifts.The Monte Carlo simulations show that this approach produces unbiased estimates of the fractional...

  14. Nonlinear MHD waves in a Prominence Foot

    CERN Document Server

    Ofman, Leon; Kucera, Therese; Schmieder, Brigitte

    2015-01-01

    We study nonlinear waves in a prominence foot using 2.5D MHD model motivated by recent high-resolution observations with Hinode/SOT in Ca~II emission of a prominence on October 10, 2012 showing highly dynamic small-scale motions in the prominence material. Observations of H$\\alpha$ intensities and of Doppler shifts show similar propagating fluctuations. However the optically thick nature of the emission lines inhibits unique quantitative interpretation in terms of density. Nevertheless, we find evidence of nonlinear wave activity in the prominence foot by examining the relative magnitude of the fluctuation intensity ($\\delta I/I\\sim \\delta n/n$). The waves are evident as significant density fluctuations that vary with height, and apparently travel upward from the chromosphere into the prominence material with quasi-periodic fluctuations with typical period in the range of 5-11 minutes, and wavelengths $\\sim <$2000 km. Recent Doppler shift observations show the transverse displacement of the propagating wav...

  15. Nonlinear spin wave coupling in adjacent magnonic crystals

    Energy Technology Data Exchange (ETDEWEB)

    Sadovnikov, A. V., E-mail: sadovnikovav@gmail.com; Nikitov, S. A. [Laboratory “Metamaterials,” Saratov State University, Saratov 410012 (Russian Federation); Kotel' nikov Institute of Radioengineering and Electronics, Russian Academy of Sciences, Moscow 125009 (Russian Federation); Beginin, E. N.; Morozova, M. A.; Sharaevskii, Yu. P.; Grishin, S. V.; Sheshukova, S. E. [Laboratory “Metamaterials,” Saratov State University, Saratov 410012 (Russian Federation)

    2016-07-25

    We have experimentally studied the coupling of spin waves in the adjacent magnonic crystals. Space- and time-resolved Brillouin light-scattering spectroscopy is used to demonstrate the frequency and intensity dependent spin-wave energy exchange between the side-coupled magnonic crystals. The experiments and the numerical simulation of spin wave propagation in the coupled periodic structures show that the nonlinear phase shift of spin wave in the adjacent magnonic crystals leads to the nonlinear switching regime at the frequencies near the forbidden magnonic gap. The proposed side-coupled magnonic crystals represent a significant advance towards the all-magnonic signal processing in the integrated magnonic circuits.

  16. Hybrid quantum systems for enhanced nonlinear optical susceptibilities

    CERN Document Server

    Sullivan, Dennis; Kuzyk, Mark G

    2016-01-01

    Significant effort has been expended in the search for materials with ultra-fast nonlinear-optical susceptibilities, but most fall far below the fundamental limits. This work applies a theoretical materials development program that has identified a promising new hybrid made of a nanorod and a molecule. This system uses the electrostatic dipole moment of the molecule to break the symmetry of the metallic nanostructure that shifts the energy spectrum to make it optimal for a nonlinear-optical response near the fundamental limit. The structural parameters are varied to determine the ideal configuration, providing guidelines for making the best structures.

  17. Cavity optomechanics with a nonlinear photonic-crystal nanomembrane

    Energy Technology Data Exchange (ETDEWEB)

    Makles, Kevin; Kuhn, Aurélien; Briant, Tristan; Cohadon, Pierre-François; Heidmann, Antoine [Laboratoire Kastler Brossel, UPMC-ENS-CNRS, Case 74, 4 place Jussieu, F75252 Paris Cedex 05 (France); Antoni, Thomas [Laboratoire de Photonique et Nanostructures LPN-CNRS, UPR-20, Route de Nozay, 91460 Marcoussis, France and Laboratoire Kastler Brossel, UPMC-ENS-CNRS, Case 74, 4 place Jussieu, F75252 Paris Cedex 05 (France); Braive, Rémy [Laboratoire de Photonique et Nanostructures LPN-CNRS, UPR-20, Route de Nozay, 91460 Marcoussis, France and Université Paris Diderot, 10, rue Alice Domon et Léonie Duquet, 75205 Paris, Cedex 13 (France); Sagnes, Isabelle; Robert-Philip, Isabelle [Laboratoire de Photonique et Nanostructures LPN-CNRS, UPR-20, Route de Nozay, 91460 Marcoussis (France)

    2014-12-04

    We have designed, fabricated and characterized a nanomembrane which could be used as a moving end mirror of a Fabry-Perot cavity. The high reflectivity and optimized mechanical properties of the membrane should allow us to demonstrate the mechanical ground state of the membrane. As any sub-micron mechanical resonator, our system demonstrates nonlinear dynamical effects. We characterize the mechanical response to a strong pump drive and observe a shift in the oscillation frequency and phase conjugation of the mechanical mode. Such nonlinear effects are expected to play a role in the quantum dynamics of the membrane as well.

  18. Properties of some nonlinear Schroedinger equations motivated through information theory

    Energy Technology Data Exchange (ETDEWEB)

    Yuan, Liew Ding; Parwani, Rajesh R, E-mail: parwani@nus.edu.s [Department of Physics, National University of Singapore, Kent Ridge (Singapore)

    2009-06-01

    We update our understanding of nonlinear Schroedinger equations motivated through information theory. In particular we show that a q-deformation of the basic nonlinear equation leads to a perturbative increase in the energy of a system, thus favouring the simplest q = 1 case. Furthermore the energy minimisation criterion is shown to be equivalent, at leading order, to an uncertainty maximisation argument. The special value eta = 1/4 for the interpolation parameter, where leading order energy shifts vanish, implies the preservation of existing supersymmetry in nonlinearised supersymmetric quantum mechanics. Physically, eta might be encoding relativistic effects.

  19. Modeling and equalization of nonlinear bandlimited satellite channels

    Science.gov (United States)

    Konstantinides, K.; Yao, K.

    1986-01-01

    The problem of modeling and equalization of a nonlinear satellite channel is considered. The channel is assumed to be bandlimited and exhibits both amplitude and phase nonlinearities. A discrete time satellite link is modeled under both uplink and downlink white Gaussian noise. Under conditions of practical interest, a simple and computationally efficient design technique for the minimum mean square error linear equalizer is presented. The bit error probability and some numerical results for a binary phase shift keyed (BPSK) system demonstrate that the proposed equalization technique outperforms standard linear receiver structures.

  20. Nonlinear elliptic systems with exponential nonlinearities

    Directory of Open Access Journals (Sweden)

    Said El Manouni

    2002-12-01

    Full Text Available In this paper we investigate the existence of solutions for {gather*} -mathop{m div}( a(| abla u | ^N| abla u |^{N-2}u = f(x,u,v quad mbox{in } Omega -mathop{m div}(a(| abla v| ^N| abla v |^{N-2}v = g(x,u,v quad mbox{in } Omega u(x = v(x = 0 quad mbox{on }partial Omega. end{gather*} Where $Omega$ is a bounded domain in ${mathbb{R}}^N$, $Ngeq 2$, $f$ and $g$ are nonlinearities having an exponential growth on $Omega$ and $a$ is a continuous function satisfying some conditions which ensure the existence of solutions.

  1. Nonlinearity and disorder: Classification and stability of nonlinear impurity modes

    DEFF Research Database (Denmark)

    Sukhorukov, Andrey A.; Kivshar, Yuri S.; Bang, Ole

    2001-01-01

    We study the effects produced by competition of two physical mechanisms of energy localization in inhomogeneous nonlinear systems. As an example, we analyze spatially localized modes supported by a nonlinear impurity in the generalized nonlinear Schrödinger equation and describe three types of no...

  2. Generalized Nonlinear Yule Models

    Science.gov (United States)

    Lansky, Petr; Polito, Federico; Sacerdote, Laura

    2016-10-01

    With the aim of considering models related to random graphs growth exhibiting persistent memory, we propose a fractional nonlinear modification of the classical Yule model often studied in the context of macroevolution. Here the model is analyzed and interpreted in the framework of the development of networks such as the World Wide Web. Nonlinearity is introduced by replacing the linear birth process governing the growth of the in-links of each specific webpage with a fractional nonlinear birth process with completely general birth rates. Among the main results we derive the explicit distribution of the number of in-links of a webpage chosen uniformly at random recognizing the contribution to the asymptotics and the finite time correction. The mean value of the latter distribution is also calculated explicitly in the most general case. Furthermore, in order to show the usefulness of our results, we particularize them in the case of specific birth rates giving rise to a saturating behaviour, a property that is often observed in nature. The further specialization to the non-fractional case allows us to extend the Yule model accounting for a nonlinear growth.

  3. Nonlinear Regression with R

    CERN Document Server

    Ritz, Christian; Parmigiani, Giovanni

    2009-01-01

    R is a rapidly evolving lingua franca of graphical display and statistical analysis of experiments from the applied sciences. This book provides a coherent treatment of nonlinear regression with R by means of examples from a diversity of applied sciences such as biology, chemistry, engineering, medicine and toxicology.

  4. Methods of nonlinear kinetics

    OpenAIRE

    Gorban, A. N.; Karlin, I.V.

    2003-01-01

    Nonlinear kinetic equations are reviewed for a wide audience of specialists and postgraduate students in physics, mathematical physics, material science, chemical engineering and interdisciplinary research. Contents: The Boltzmann equation, Phenomenology and Quasi-chemical representation of the Boltzmann equation, Kinetic models, Discrete velocity models, Direct simulation, Lattice Gas and Lattice Boltzmann models, Minimal Boltzmann models for flows at low Knudsen number, Other kinetic equati...

  5. Intramolecular and nonlinear dynamics

    Energy Technology Data Exchange (ETDEWEB)

    Davis, M.J. [Argonne National Laboratory, IL (United States)

    1993-12-01

    Research in this program focuses on three interconnected areas. The first involves the study of intramolecular dynamics, particularly of highly excited systems. The second area involves the use of nonlinear dynamics as a tool for the study of molecular dynamics and complex kinetics. The third area is the study of the classical/quantum correspondence for highly excited systems, particularly systems exhibiting classical chaos.

  6. Nonlinear Hamiltonian systems

    DEFF Research Database (Denmark)

    Jørgensen, Michael Finn

    1995-01-01

    It is generally very difficult to solve nonlinear systems, and such systems often possess chaotic solutions. In the rare event that a system is completely solvable, it is said to integrable. Such systems never have chaotic solutions. Using the Inverse Scattering Transform Method (ISTM) two...

  7. Nonlinear phased array imaging

    Science.gov (United States)

    Croxford, Anthony J.; Cheng, Jingwei; Potter, Jack N.

    2016-04-01

    A technique is presented for imaging acoustic nonlinearity within a specimen using ultrasonic phased arrays. Acoustic nonlinearity is measured by evaluating the difference in energy of the transmission bandwidth within the diffuse field produced through different focusing modes. The two different modes being classical beam forming, where delays are applied to different element of a phased array to physically focus the energy at a single location (parallel firing) and focusing in post processing, whereby one element at a time is fired and a focused image produced in post processing (sequential firing). Although these two approaches are linearly equivalent the difference in physical displacement within the specimen leads to differences in nonlinear effects. These differences are localized to the areas where the amplitude is different, essentially confining the differences to the focal point. Direct measurement at the focal point are however difficult to make. In order to measure this the diffuse field is used. It is a statistical property of the diffuse field that it represents the total energy in the system. If the energy in the diffuse field for both the sequential and parallel firing case is measured then the difference between these, within the input signal bandwidth, is largely due to differences at the focal spot. This difference therefore gives a localized measurement of where energy is moving out of the transmission bandwidth due to nonlinear effects. This technique is used to image fatigue cracks and other damage types undetectable with conventional linear ultrasonic measurements.

  8. Generalized Nonlinear Yule Models

    Science.gov (United States)

    Lansky, Petr; Polito, Federico; Sacerdote, Laura

    2016-11-01

    With the aim of considering models related to random graphs growth exhibiting persistent memory, we propose a fractional nonlinear modification of the classical Yule model often studied in the context of macroevolution. Here the model is analyzed and interpreted in the framework of the development of networks such as the World Wide Web. Nonlinearity is introduced by replacing the linear birth process governing the growth of the in-links of each specific webpage with a fractional nonlinear birth process with completely general birth rates. Among the main results we derive the explicit distribution of the number of in-links of a webpage chosen uniformly at random recognizing the contribution to the asymptotics and the finite time correction. The mean value of the latter distribution is also calculated explicitly in the most general case. Furthermore, in order to show the usefulness of our results, we particularize them in the case of specific birth rates giving rise to a saturating behaviour, a property that is often observed in nature. The further specialization to the non-fractional case allows us to extend the Yule model accounting for a nonlinear growth.

  9. Trirefringence in nonlinear metamaterials

    CERN Document Server

    De Lorenci, Vitorio A

    2012-01-01

    We study the propagation of electromagnetic waves in the limit of geometrical optics for a class of nearly transparent nonlinear uniaxial metamaterials for which their permittivity tensors present a negative principal component. Their permeability are assumed positive and dependent on the electric field. We show that light waves experience triple refraction -- trirefringence. Additionally to the ordinary wave, two extraordinary waves propagate in such media.

  10. Nonlinear silicon photonics

    Science.gov (United States)

    Borghi, M.; Castellan, C.; Signorini, S.; Trenti, A.; Pavesi, L.

    2017-09-01

    Silicon photonics is a technology based on fabricating integrated optical circuits by using the same paradigms as the dominant electronics industry. After twenty years of fervid development, silicon photonics is entering the market with low cost, high performance and mass-manufacturable optical devices. Until now, most silicon photonic devices have been based on linear optical effects, despite the many phenomenologies associated with nonlinear optics in both bulk materials and integrated waveguides. Silicon and silicon-based materials have strong optical nonlinearities which are enhanced in integrated devices by the small cross-section of the high-index contrast silicon waveguides or photonic crystals. Here the photons are made to strongly interact with the medium where they propagate. This is the central argument of nonlinear silicon photonics. It is the aim of this review to describe the state-of-the-art in the field. Starting from the basic nonlinearities in a silicon waveguide or in optical resonator geometries, many phenomena and applications are described—including frequency generation, frequency conversion, frequency-comb generation, supercontinuum generation, soliton formation, temporal imaging and time lensing, Raman lasing, and comb spectroscopy. Emerging quantum photonics applications, such as entangled photon sources, heralded single-photon sources and integrated quantum photonic circuits are also addressed at the end of this review.

  11. Nonlinear fibre optics overview

    DEFF Research Database (Denmark)

    Travers, J. C.; Frosz, Michael Henoch; Dudley, J. M.

    2010-01-01

    , provides a background to the associated nonlinear optical processes, treats the generation mechanisms from continuous wave to femtosecond pulse pump regimes and highlights the diverse applications. A full discussion of numerical methods and comprehensive computer code are also provided, enabling readers...

  12. Nonlinear silicon photonics

    Science.gov (United States)

    Tsia, Kevin K.; Jalali, Bahram

    2010-05-01

    An intriguing optical property of silicon is that it exhibits a large third-order optical nonlinearity, with orders-ofmagnitude larger than that of silica glass in the telecommunication band. This allows efficient nonlinear optical interaction at relatively low power levels in a small footprint. Indeed, we have witnessed a stunning progress in harnessing the Raman and Kerr effects in silicon as the mechanisms for enabling chip-scale optical amplification, lasing, and wavelength conversion - functions that until recently were perceived to be beyond the reach of silicon. With all the continuous efforts developing novel techniques, nonlinear silicon photonics is expected to be able to reach even beyond the prior achievements. Instead of providing a comprehensive overview of this field, this manuscript highlights a number of new branches of nonlinear silicon photonics, which have not been fully recognized in the past. In particular, they are two-photon photovoltaic effect, mid-wave infrared (MWIR) silicon photonics, broadband Raman effects, inverse Raman scattering, and periodically-poled silicon (PePSi). These novel effects and techniques could create a new paradigm for silicon photonics and extend its utility beyond the traditionally anticipated applications.

  13. Is this scaling nonlinear?

    CERN Document Server

    Leitao, J C; Gerlach, M; Altmann, E G

    2016-01-01

    One of the most celebrated findings in complex systems in the last decade is that different indexes y (e.g., patents) scale nonlinearly with the population~x of the cities in which they appear, i.e., $y\\sim x^\\beta, \\beta \

  14. Nonlinear Gravitational Lagrangians revisited

    CERN Document Server

    Magnano, Guido

    2016-01-01

    The Legendre transformation method, applied in 1987 to deal with purely metric gravitational Lagrangians with nonlinear dependence on the Ricci tensor, is extended to metric-affine models and is shown to provide a concise and insightful comparison of the dynamical content of the two variational frameworks.

  15. Nonlinearities in Microwave Superconductivity

    OpenAIRE

    Ledenyov, Dimitri O.; Ledenyov, Viktor O.

    2012-01-01

    The research is focused on the modeling of nonlinear properties of High Temperature Superconducting (HTS) thin films, using Bardeen, Cooper, Schrieffer and Lumped Element Circuit theories, with purpose to enhance microwave power handling capabilities of microwave filters and optimize design of microwave circuits in micro- and nano- electronics.

  16. Nonlinear tsunami generation mechanism

    Directory of Open Access Journals (Sweden)

    M. A. Nosov

    2001-01-01

    Full Text Available The nonlinear mechanism of long gravitational surface water wave generation by high-frequency bottom oscillations in a water layer of constant depth is investigated analytically. The connection between the surface wave amplitude and the parameters of bottom oscillations and source length is investigated.

  17. Topics in Nonlinear Dynamics

    DEFF Research Database (Denmark)

    Mosekilde, Erik

    Through a significant number of detailed and realistic examples this book illustrates how the insights gained over the past couple of decades in the fields of nonlinear dynamics and chaos theory can be applied in practice. Aomng the topics considered are microbiological reaction systems, ecological...

  18. Terahertz semiconductor nonlinear optics

    DEFF Research Database (Denmark)

    Turchinovich, Dmitry; Hvam, Jørn Märcher; Hoffmann, Matthias

    2013-01-01

    nonlinearity in doped semiconductors originates from the near-instantaneous heating of free electrons in the ponderomotive potential created by electric field of the THz pulse, leading to ultrafast increase of electron effective mass by intervalley scattering. Modification of effective mass in turn leads...

  19. NONLINEAR MHD WAVES IN A PROMINENCE FOOT

    Energy Technology Data Exchange (ETDEWEB)

    Ofman, L. [Catholic University of America, Washington, DC 20064 (United States); Knizhnik, K.; Kucera, T. [NASA Goddard Space Flight Center, Code 671, Greenbelt, MD 20771 (United States); Schmieder, B. [LESIA, Observatoire de Paris, PSL Research University, CNRS, Sorbonne Universités, UPMC Univ. Paris 06, Univ. Paris-Diderot, Sorbonne Paris Cit, 5 place Jules Janssen, F-92195 Meudon (France)

    2015-11-10

    We study nonlinear waves in a prominence foot using a 2.5D MHD model motivated by recent high-resolution observations with Hinode/Solar Optical Telescope in Ca ii emission of a prominence on 2012 October 10 showing highly dynamic small-scale motions in the prominence material. Observations of Hα intensities and of Doppler shifts show similar propagating fluctuations. However, the optically thick nature of the emission lines inhibits a unique quantitative interpretation in terms of density. Nevertheless, we find evidence of nonlinear wave activity in the prominence foot by examining the relative magnitude of the fluctuation intensity (δI/I ∼ δn/n). The waves are evident as significant density fluctuations that vary with height and apparently travel upward from the chromosphere into the prominence material with quasi-periodic fluctuations with a typical period in the range of 5–11 minutes and wavelengths <2000 km. Recent Doppler shift observations show the transverse displacement of the propagating waves. The magnetic field was measured with the THEMIS instrument and was found to be 5–14 G. For the typical prominence density the corresponding fast magnetosonic speed is ∼20 km s{sup −1}, in qualitative agreement with the propagation speed of the detected waves. The 2.5D MHD numerical model is constrained with the typical parameters of the prominence waves seen in observations. Our numerical results reproduce the nonlinear fast magnetosonic waves and provide strong support for the presence of these waves in the prominence foot. We also explore gravitational MHD oscillations of the heavy prominence foot material supported by dipped magnetic field structure.

  20. Nonlinear Optical Terahertz Technology Project

    Data.gov (United States)

    National Aeronautics and Space Administration — Our approach is based on high-Q optical WGM resonators made with a nonlinear crystal. Such resonators have been demonstrated to dramatically enhance nonlinear...

  1. Phase retrieval using nonlinear diversity.

    Science.gov (United States)

    Lu, Chien-Hung; Barsi, Christopher; Williams, Matthew O; Kutz, J Nathan; Fleischer, Jason W

    2013-04-01

    We extend the Gerchberg-Saxton algorithm to phase retrieval in a nonlinear system. Using a tunable photorefractive crystal, we experimentally demonstrate the noninterferometric technique by reconstructing an unknown phase object from optical intensity measurements taken at different nonlinear strengths.

  2. Strong nonlinear oscillators analytical solutions

    CERN Document Server

    Cveticanin, Livija

    2017-01-01

    This book outlines an analytical solution procedure of the pure nonlinear oscillator system, offering a solution for free and forced vibrations of the one-degree-of-freedom strong nonlinear system with constant and time variable parameter. Includes exercises.

  3. Predicting phase shift of elastic waves in pipes due to fluid flow and imperfections

    DEFF Research Database (Denmark)

    Thomsen, Jon Juel; Dahl, Jonas; Fuglede, Niels

    2009-01-01

    Flexural vibrations of a fluid-conveying pipe is investigated, with special consideration to the spatial shift in phase caused by fluid flow and various imperfections, e.g., non-ideal supports, non-uniform stiffness or mass, non-proportional damping, weak nonlinearity, and flow pulsation. This is......Flexural vibrations of a fluid-conveying pipe is investigated, with special consideration to the spatial shift in phase caused by fluid flow and various imperfections, e.g., non-ideal supports, non-uniform stiffness or mass, non-proportional damping, weak nonlinearity, and flow pulsation....... This is relevant for understanding wave propagation in elastic media in general, and for the design and trouble-shooting of phase-shift measuring devices such as Coriolis mass flowmeters in particular. A multiple time scaling perturbation analysis is employed for a simple model of a fluid-conveying pipe...

  4. Anthropometric changes and fluid shifts

    Science.gov (United States)

    Thornton, W. E.; Hoffler, G. W.; Rummel, J. A.

    1974-01-01

    Several observations of body size, shape, posture, and configuration were made to document changes resulting from direct effects of weightlessness during the Skylab 4 mission. After the crewmen were placed in orbit, a number of anatomical and anthropometric changes occurred including a straightening of the thoracolumbar spine, a general decrease in truncal girth, and an increase in height. By the time of the earliest in-flight measurement on mission day 3, all crewmen had lost more than two liters of extravascular fluid from the calf and thigh. The puffy facies, the bird legs effect, the engorgement of upper body veins, and the reduced volume of lower body veins were all documented with photographs. Center-of-mass measurements confirmed a fluid shift cephalad. This shift remained throughout the mission until recovery, when a sharp reversal occurred; a major portion of the reversal was completed in a few hours. The anatomical changes are of considerable scientific interest and of import to the human factors design engineer, but the shifts of blood and extravascular fluid are of more consequence. It is hypothesized that the driving force for the fluid shift is the intrinsic and unopposed lower limb elasticity that forces venous blood and then other fluid cephalad.

  5. Environmental Protection: a shifting focus

    NARCIS (Netherlands)

    Dr. ir. Jan Venselaar

    2004-01-01

    The last two decades have seen a fundamental change in the way chemistry handles environmental issues. A shift in focus has occurred from 'end-of-pipe' to prevention and process integration. Presently an even more fundamental change is brought about by the need for sustainable development. It is

  6. Wavelength-shifted Cherenkov radiators

    Science.gov (United States)

    Krider, E. P.; Jacobson, V. L.; Pifer, A. E.; Polakos, P. A.; Kurz, R. J.

    1976-01-01

    The scintillation and Cherenkov responses of plastic Cherenkov radiators containing different wavelength-shifting fluors in varying concentrations have been studied in beams of low energy protons and pions. For cosmic ray applications, where large Cherenkov to scintillation ratios are desired, the optimum fluor concentrations are 0.000025 by weight or less.

  7. The Shift Needed for Sustainability

    Science.gov (United States)

    Smith, Peter A. C.; Sharicz, Carol

    2011-01-01

    Purpose: The purpose of this action research is to begin to assess to what extent organizations have in practice begun to make the shift towards triple bottom line (TBL) sustainability. Design/methodology/approach: A definition of TBL sustainability is provided, and key elements of TBL sustainability considered necessary to success are identified…

  8. Crichton's phase-shift ambiguity

    NARCIS (Netherlands)

    Atkinson, D.; Johnson, P.W.; Mehta, N.; Roo, M. de

    1973-01-01

    A re-examination of the SPD phase-shift ambiguity is made with a view to understanding certain singular features of the elastic unitarity constraint. An explicit solution of Crichton's equations is presented, and certain features of this solution are displayed graphically. In particular, it is shown

  9. Application of Nonlinear Elastic Resonance Spectroscopy For Damage Detection In Concrete: An Interesting Story

    Energy Technology Data Exchange (ETDEWEB)

    Byers, Loren W. [Los Alamos National Laboratory; Ten Cate, James A. [Los Alamos National Laboratory; Johnson, Paul A. [Los Alamos National Laboratory

    2012-06-28

    Nonlinear resonance ultrasound spectroscopy experiments conducted on concrete cores, one chemically and mechanically damaged by alkali-silica reactivity, and one undamaged, show that this material displays highly nonlinear wave behavior, similar to many other damaged materials. They find that the damaged sample responds more nonlinearly, manifested by a larger resonant peak and modulus shift as a function of strain amplitude. The nonlinear response indicates that there is a hysteretic influence in the stress-strain equation of state. Further, as in some other materials, slow dynamics are present. The nonlinear response they observe in concrete is an extremely sensitive indicator of damage. Ultimately, nonlinear wave methods applied to concrete may be used to guide mixing, curing, or other production techniques, in order to develop materials with particular desired qualities such as enhanced strength or chemical resistance, and to be used for damage inspection.

  10. Size-Dependent Raman Shifts for nanocrystals.

    Science.gov (United States)

    Gao, Yukun; Zhao, Xinmei; Yin, Penggang; Gao, Faming

    2016-04-22

    Raman spectroscopy is a very sensitive tool for probing semiconductor nanocrystals. The underlying mechanism behind the size-dependent Raman shifts is still quite controversial. Here we offer a new theoretical method for the quantum confinement effects on the Raman spectra of semiconductor nanocrystals. We propose that the shift of Raman spectra in nanocrystals can result from two overlapping effects: the quantum effect shift and surface effect shift. The quantum effect shift is extracted from an extended Kubo formula, the surface effect shift is determined via the first principles calculations. Fairly good prediction of Raman shifts can be obtained without the use of any adjustable parameter. Closer analysis shows that the size-dependent Raman shifts in Si nanocrystals mainly result from the quantum effect shifts. For nanodiamond, the proportion of surface effect shift in Raman shift is up to about 40%. Such model can also provide a good baseline for using Raman spectroscopy as a tool to measure size.

  11. Cubication of Conservative Nonlinear Oscillators

    Science.gov (United States)

    Belendez, Augusto; Alvarez, Mariela L.; Fernandez, Elena; Pascual, Immaculada

    2009-01-01

    A cubication procedure of the nonlinear differential equation for conservative nonlinear oscillators is analysed and discussed. This scheme is based on the Chebyshev series expansion of the restoring force, and this allows us to approximate the original nonlinear differential equation by a Duffing equation in which the coefficients for the linear…

  12. Terahertz Nonlinear Optics in Semiconductors

    DEFF Research Database (Denmark)

    Turchinovich, Dmitry; Hvam, Jørn Märcher; Hoffmann, Matthias C.

    2013-01-01

    We demonstrate the nonlinear optical effects – selfphase modulation and saturable absorption of a single-cycle THz pulse in a semiconductor. Resulting from THz-induced modulation of Drude plasma, these nonlinear optical effects, in particular, lead to self-shortening and nonlinear spectral...

  13. Fault Detection for Nonlinear Systems

    DEFF Research Database (Denmark)

    Stoustrup, Jakob; Niemann, H.H.

    1998-01-01

    The paper describes a general method for designing fault detection and isolation (FDI) systems for nonlinear processes. For a rich class of nonlinear systems, a nonlinear FDI system can be designed using convex optimization procedures. The proposed method is a natural extension of methods based...

  14. The optical nonlinearity of gold nanoparticles prepared by bioreduction method

    Science.gov (United States)

    Balbuena Ortega, A.; Arroyo Carrasco, M. L.; Gayou, V. L.; Orduña Díaz, A.; Delgado Macuil, R.; Rojas López, Marlon

    2013-11-01

    Nonlinear optical and electronic properties of nanosized metal particles have drawn considerable attention because of their strong and size-dependent plasmon resonance absorption. In a metal nanoparticle system such as gold dispersed in a transparent matrix, an absorption peak due to surface plasmon resonance is usually observed in the visible spectral region. Metal nanoparticles are of special interest as nonlinear materials for optical switching and computing because of their relatively large third-order nonlinearity (χ3) and ultrafast response time. The purpose of this study was to analyze the nonlinear optical properties of biosynthesized gold nanoparticles. The samples were prepared by biosynthesis method using yeast extract as reducing agent and the nonlinear optical properties of the nanoparticles were investigated using a single beam Z-scan technique with a beam power of 20 mW and operated at wavelength of 514 nm. The reaction between metal ions and yeast extracts were monitored by UV-visible spectra of Au nanoparticles in aqueous solution with different pH (3-6). The surface plasmon peak position was shifted from 528 nm to 573 nm, according to of pH variation 4 to 6. The average particle size was calculated by the absorption peak position using the Fernig method, from 42 to 103 nm. The z-scan curves showed a negative nonlocal nonlinear refractive index with a magnitude dependent on the nanoparticle size.

  15. Two widespread green Neottia species (Orchidaceae) show mycorrhizal preference for Sebacinales in various habitats and ontogenetic stages.

    Science.gov (United States)

    Těšitelová, Tamara; Kotilínek, Milan; Jersáková, Jana; Joly, François-Xavier; Košnar, Jiří; Tatarenko, Irina; Selosse, Marc-André

    2015-03-01

    Plant dependence on fungal carbon (mycoheterotrophy) evolved repeatedly. In orchids, it is connected with a mycorrhizal shift from rhizoctonia to ectomycorrhizal fungi and a high natural (13)C and (15)N abundance. Some green relatives of mycoheterotrophic species show identical trends, but most of these remain unstudied, blurring our understanding of evolution to mycoheterotrophy. We analysed mycorrhizal associations and (13)C and (15)N biomass content in two green species, Neottia ovata and N. cordata (tribe Neottieae), from a genus comprising green and nongreen (mycoheterotrophic) species. Our study covered 41 European sites, including different meadow and forest habitats and orchid developmental stages. Fungal ITS barcoding and electron microscopy showed that both Neottia species associated mainly with nonectomycorrhizal Sebacinales Clade B, a group of rhizoctonia symbionts of green orchids, regardless of the habitat or growth stage. Few additional rhizoctonias from Ceratobasidiaceae and Tulasnellaceae, and ectomycorrhizal fungi were detected. Isotope abundances did not detect carbon gain from the ectomycorrhizal fungi, suggesting a usual nutrition of rhizoctonia-associated green orchids. Considering associations of related partially or fully mycoheterotrophic species such as Neottia camtschatea or N. nidus-avis with ectomycorrhizal Sebacinales Clade A, we propose that the genus Neottia displays a mycorrhizal preference for Sebacinales and that the association with nonectomycorrhizal Sebacinales Clade B is likely ancestral. Such a change in preference for mycorrhizal associates differing in ecology within the same fungal taxon is rare among orchids. Moreover, the existence of rhizoctonia-associated Neottia spp. challenges the shift to ectomycorrhizal fungi as an ancestral pre-adaptation to mycoheterotrophy in the whole Neottieae.

  16. Nonlinear electrostatic drift Kelvin-Helmholtz instability

    Science.gov (United States)

    Sharma, Avadhesh C.; Srivastava, Krishna M.

    1993-01-01

    Nonlinear analysis of electrostatic drift Kelvin-Helmholtz instability is performed. It is shown that the analysis leads to the propagation of the weakly nonlinear dispersive waves, and the nonlinear behavior is governed by the nonlinear Burger's equation.

  17. Non-linear effects for cylindrical gravitational two-soliton

    CERN Document Server

    Tomizawa, Shinya

    2015-01-01

    Using a cylindrical soliton solution to the four-dimensional vacuum Einstein equation, we study non-linear effects of gravitational waves such as Faraday rotation and time shift phenomenon. In the previous work, we analyzed the single-soliton solution constructed by the Pomeransky's improved inverse scattering method. In this work, we construct a new two-soliton solution with complex conjugate poles, by which we can avoid light-cone singularities unavoidable in a single soliton case. In particular, we compute amplitudes of such non-linear gravitational waves and time-dependence of the polarizations. Furthermore, we consider the time shift phenomenon for soliton waves, which means that a wave packet can propagate at slower velocity than light.

  18. Collisional Effects on Nonlinear Ion Drag Force for Small Grains

    CERN Document Server

    Hutchinson, I H

    2013-01-01

    The ion drag force arising from plasma flow past an embedded spherical grain is calculated self-consistently and non-linearly using particle in cell codes, accounting for ion-neutral collisions. Using ion velocity distribution appropriate for ion drift driven by a force field gives wake potential and force greatly different from a shifted Maxwellian distribution, regardless of collisionality. The low-collisionality forces are shown to be consistent with estimates based upon cross-sections for scattering in a Yukawa (shielded) grain field, but only if non-linear shielding length is used. Finite collisionality initially enhances the drag force, but only by up to a factor of 2. Larger collisionality eventually reduces the drag force. In the collisional regime, the drift distribution gives larger drag than the shift distribution even at velocities where their collisionless drags are equal. Comprehensive practical analytic formulas for force that fit the calculations are provided.

  19. Nonlinear Silicon Photonic Signal Processing Devices for Future Optical Networks

    Directory of Open Access Journals (Sweden)

    Cosimo Lacava

    2017-01-01

    Full Text Available In this paper, we present a review on silicon-based nonlinear devices for all optical nonlinear processing of complex telecommunication signals. We discuss some recent developments achieved by our research group, through extensive collaborations with academic partners across Europe, on optical signal processing using silicon-germanium and amorphous silicon based waveguides as well as novel materials such as silicon rich silicon nitride and tantalum pentoxide. We review the performance of four wave mixing wavelength conversion applied on complex signals such as Differential Phase Shift Keying (DPSK, Quadrature Phase Shift Keying (QPSK, 16-Quadrature Amplitude Modulation (QAM and 64-QAM that dramatically enhance the telecom signal spectral efficiency, paving the way to next generation terabit all-optical networks.

  20. Geographical variation and sexual differences of body length and age composition in Rana temporaria: the ontogenetic development and phenotypic trends

    Directory of Open Access Journals (Sweden)

    Lyapkov Sergey

    2012-06-01

    Full Text Available The analysis of literature data on the mean values of age and body length of adult individuals of widespread species Rana temporaria from about 70 spatially separated populations, including our published data, was conducted. The evident trend in population mean age increase with the decrease of the of activity season length was revealed as well as the absence of that trend in the mean body length, with the maximal mean value in body length being near central part of the range. Our explanation of non-linear trend in the mean values of body length does not contradict other models of geographic variability explaining the correspondence and discrepance with the Bergman rule. In addition our explanation corresponds to the revealed features of interpopulation variation in growth rate. The revealed trend of variation in the mean body length is resulted from both growth rate decrease and mean age increase with the decrease in the length of activity season. The relatively low mean values of body length in populations from south and southern-west borders of the range are explained not only by low mean age but by lower growth rate despite high length of activity season. The interpopulation variation in body length is determined not only by body length but by age composition differences both between and within population. Therefore, the direction and intensity of sexual differences have not distinct trends, and the correspondence to Rensch rule (in contrast to Bergman rule is rarely observed.

  1. Optothermal nonlinearity of silica aerogel

    CERN Document Server

    Braidotti, Maria Chiara; Fleming, Adam; Samuels, Michiel C; Di Falco, Andrea; Conti, Claudio

    2016-01-01

    We report on the characterization of silica aerogel thermal optical nonlinearity, obtained by z-scan technique. The results show that typical silica aerogels have nonlinear optical coefficient similar to that of glass $(\\simeq 10^{-12} $m$^2/$W), with negligible optical nonlinear absorption. The non\\-li\\-near coefficient can be increased to values in the range of $10^{-10} $m$^2/$W by embedding an absorbing dye in the aerogel. This value is one order of magnitude higher than that observed in the pure dye and in typical highly nonlinear materials like liquid crystals.

  2. Essentials of nonlinear optics

    CERN Document Server

    Murti, Y V G S

    2014-01-01

    Current literature on Nonlinear Optics varies widely in terms of content, style, and coverage of specific topics, relative emphasis of areas and the depth of treatment. While most of these books are excellent resources for the researchers, there is a strong need for books appropriate for presenting the subject at the undergraduate or postgraduate levels in Universities. The need for such a book to serve as a textbook at the level of the bachelors and masters courses was felt by the authors while teaching courses on nonlinear optics to students of both science and engineering during the past two decades. This book has emerged from an attempt to address the requirement of presenting the subject at college level. A one-semester course covering the essentials can effectively be designed based on this.

  3. Nonlinear metamaterials for holography

    Science.gov (United States)

    Almeida, Euclides; Bitton, Ora; Prior, Yehiam

    2016-08-01

    A hologram is an optical element storing phase and possibly amplitude information enabling the reconstruction of a three-dimensional image of an object by illumination and scattering of a coherent beam of light, and the image is generated at the same wavelength as the input laser beam. In recent years, it was shown that information can be stored in nanometric antennas giving rise to ultrathin components. Here we demonstrate nonlinear multilayer metamaterial holograms. A background free image is formed at a new frequency--the third harmonic of the illuminating beam. Using e-beam lithography of multilayer plasmonic nanoantennas, we fabricate polarization-sensitive nonlinear elements such as blazed gratings, lenses and other computer-generated holograms. These holograms are analysed and prospects for future device applications are discussed.

  4. Nonlinear metamaterials for holography

    Science.gov (United States)

    Almeida, Euclides; Bitton, Ora

    2016-01-01

    A hologram is an optical element storing phase and possibly amplitude information enabling the reconstruction of a three-dimensional image of an object by illumination and scattering of a coherent beam of light, and the image is generated at the same wavelength as the input laser beam. In recent years, it was shown that information can be stored in nanometric antennas giving rise to ultrathin components. Here we demonstrate nonlinear multilayer metamaterial holograms. A background free image is formed at a new frequency—the third harmonic of the illuminating beam. Using e-beam lithography of multilayer plasmonic nanoantennas, we fabricate polarization-sensitive nonlinear elements such as blazed gratings, lenses and other computer-generated holograms. These holograms are analysed and prospects for future device applications are discussed. PMID:27545581

  5. Nonlinear data assimilation

    CERN Document Server

    Van Leeuwen, Peter Jan; Reich, Sebastian

    2015-01-01

    This book contains two review articles on nonlinear data assimilation that deal with closely related topics but were written and can be read independently. Both contributions focus on so-called particle filters. The first contribution by Jan van Leeuwen focuses on the potential of proposal densities. It discusses the issues with present-day particle filters and explorers new ideas for proposal densities to solve them, converging to particle filters that work well in systems of any dimension, closing the contribution with a high-dimensional example. The second contribution by Cheng and Reich discusses a unified framework for ensemble-transform particle filters. This allows one to bridge successful ensemble Kalman filters with fully nonlinear particle filters, and allows a proper introduction of localization in particle filters, which has been lacking up to now.

  6. Nonlinearity without Superluminality

    CERN Document Server

    Kent, A

    2002-01-01

    Quantum theory is compatible with special relativity. In particular, though measurements on entangled systems are correlated in a way that cannot be reproduced by local hidden variables, they cannot be used for superluminal signalling. As Gisin and Polchinski first pointed out, this is not true for general nonlinear modifications of the Schroedinger equation. Excluding superluminal signalling has thus been taken to rule out most nonlinear versions of quantum theory. The no superluminal signalling constraint has also been used for alternative derivations of the optimal fidelities attainable for imperfect quantum cloning and other operations. These results apply to theories satisfying the rule that their predictions for widely separated and slowly moving entangled systems can be approximated by non-relativistic equations of motion with respect to a preferred time coordinate. This paper describes a natural way in which this rule might fail to hold. In particular, it is shown that quantum readout devices which di...

  7. Monte Carlo and nonlinearities

    CERN Document Server

    Dauchet, Jérémi; Blanco, Stéphane; Caliot, Cyril; Charon, Julien; Coustet, Christophe; Hafi, Mouna El; Eymet, Vincent; Farges, Olivier; Forest, Vincent; Fournier, Richard; Galtier, Mathieu; Gautrais, Jacques; Khuong, Anaïs; Pelissier, Lionel; Piaud, Benjamin; Roger, Maxime; Terrée, Guillaume; Weitz, Sebastian

    2016-01-01

    The Monte Carlo method is widely used to numerically predict systems behaviour. However, its powerful incremental design assumes a strong premise which has severely limited application so far: the estimation process must combine linearly over dimensions. Here we show that this premise can be alleviated by projecting nonlinearities on a polynomial basis and increasing the configuration-space dimension. Considering phytoplankton growth in light-limited environments, radiative transfer in planetary atmospheres, electromagnetic scattering by particles and concentrated-solar-power-plant productions, we prove the real world usability of this advance on four test-cases that were so far regarded as impracticable by Monte Carlo approaches. We also illustrate an outstanding feature of our method when applied to sharp problems with interacting particles: handling rare events is now straightforward. Overall, our extension preserves the features that made the method popular: addressing nonlinearities does not compromise o...

  8. Nonlinear Photonics 2014: introduction.

    Science.gov (United States)

    Akhmediev, N; Kartashov, Yaroslav

    2015-01-12

    International Conference "Nonlinear Photonics-2014" took place in Barcelona, Spain on July 27-31, 2014. It was a part of the "Advanced Photonics Congress" which is becoming a traditional notable event in the world of photonics. The current focus issue of Optics Express contains contributions from the participants of the Conference and the Congress. The articles in this focus issue by no means represent the total number of the congress contributions (around 400). However, it demonstrates wide range of topics covered at the event. The next conference of this series is to be held in 2016 in Australia, which is the home of many researchers working in the field of photonics in general and nonlinear photonics in particular.

  9. Looping through the Lamb Shift

    Energy Technology Data Exchange (ETDEWEB)

    Hazi, A U

    2007-02-06

    Sometimes in science, a small measurement can have big ramifications. For a team of Livermore scientists, such was the case when they measured a small shift in the spectrum of extremely ionized atoms of uranium. The measurement involves the Lamb shift, a subtle change in the energy of an electron orbiting an atom's nucleus. The precision of the Livermore result was 10 times greater than that of existing measurements, making it the best measurement to date of a complicated correction to the simplest quantum description of how atoms behave. The measurement introduces a new realm in the search for deviations between the theory of quantum electrodynamics (QED), which is an extension of quantum mechanics, and the real world. Such deviations, if discovered, would have far-reaching consequences, indicating that QED is not a fundamental theory of nature.

  10. Lamb shift in muonic deuterium

    Energy Technology Data Exchange (ETDEWEB)

    Gorchtein, Mikhail; Vanderhaeghen, Marc [Institut für Kernphysik, Universität Mainz, 55128 Mainz (Germany); Carlson, Carl E. [Department of Physics, College of William and Mary, Williamsburg, Virginia 23187 (United States)

    2013-11-07

    We consider the two-photon exchange contribution to the 2P-2S Lamb shift in muonic deuterium in the framework of forward dispersion relations. The dispersion integrals are evaluated with minimal model dependence using experimental data on elastic deuteron form factors and inelastic electron-deuteron scattering, both in the quasielastic and hadronic range. The subtraction constant that is required to ensure convergence of the dispersion relation for the forward Compton amplitude T{sub 1} (ν,Q{sup 2}) is related to the deuteron magnetic polarizability β(Q{sup 2}) and represents the main source of uncertainty in our analysis. We obtain for the Lamb shift ΔE{sub 2P-2S} = 1.620±0.190 meV and discuss ways to further reduce this uncertainty.

  11. Anthropometric changes and fluid shifts

    Science.gov (United States)

    Thornton, W. E.; Hoffler, G. W.; Rummel, J. A.

    1977-01-01

    In an effort to obtain the most comprehensive and coherent picture of changes under weightlessness, a set of measurements on Skylab 2 was initiated and at every opportunity, additional studies were added. All pertinent information from ancillary sources were gleaned and collated. On Skylab 2, the initial anthropometric studies were scheduled in conjunction with muscle study. A single set of facial photographs was made in-flight. Additional measurements were made on Skylab 3, with photographs and truncal and limb girth measurements in-flight. Prior to Skylab 4, it was felt there was considerable evidence for large and rapid fluid shifts, so a series of in-flight volume and center of mass measurements and infrared photographs were scheduled to be conducted in the Skylab 4 mission. A number of changes were properly documented for the first time, most important of which were the fluid shifts. The following description of Skylab anthropometrics address work done on Skylab 4 primarily.

  12. Nonlinear fractional relaxation

    Indian Academy of Sciences (India)

    A Tofighi

    2012-04-01

    We define a nonlinear model for fractional relaxation phenomena. We use -expansion method to analyse this model. By studying the fundamental solutions of this model we find that when → 0 the model exhibits a fast decay rate and when → ∞ the model exhibits a power-law decay. By analysing the frequency response we find a logarithmic enhancement for the relative ratio of susceptibility.

  13. -Deformed nonlinear maps

    Indian Academy of Sciences (India)

    Ramaswamy Jaganathan; Sudeshna Sinha

    2005-03-01

    Motivated by studies on -deformed physical systems related to quantum group structures, and by the elements of Tsallis statistical mechanics, the concept of -deformed nonlinear maps is introduced. As a specific example, a -deformation procedure is applied to the logistic map. Compared to the canonical logistic map, the resulting family of -logistic maps is shown to have a wider spectrum of interesting behaviours, including the co-existence of attractors – a phenomenon rare in one-dimensional maps.

  14. Controllability of nonlinear systems.

    Science.gov (United States)

    Sussmann, H. J.; Jurdjevic, V.

    1972-01-01

    Discussion of the controllability of nonlinear systems described by the equation dx/dt - F(x,u). Concepts formulated by Chow (1939) and Lobry (1970) are applied to establish criteria for F and its derivatives to obtain qualitative information on sets which can be obtained from x which denotes a variable of state in an arbitrary, real, analytical manifold. It is shown that controllability implies strong accessibility for a large class of manifolds including Euclidean spaces.-

  15. Stochastic Nonlinear Aeroelasticity

    Science.gov (United States)

    2009-01-01

    STOCHASTIC NONLINEAR AEROELASTICITY 5a. CONTRACT NUMBER In- house 5b. GRANT NUMBER 5c. PROGRAM ELEMENT NUMBER 0601102 6. AUTHOR(S) Philip S...ABSTRACT This report documents the culmination of in- house work in the area of uncertainty quantification and probabilistic techniques for... coff U∞ cs ea lw cw Figure 6: Wing and store geometry (left), wing box structural model (middle), flutter distribution (right

  16. Nonlinear Control Systems

    Science.gov (United States)

    2007-03-01

    IEEE Transactions on Automatic Control , AC- 48, pp. 1712-1723, (2003). [14] C.I. Byrnes, A. Isidori...Nonlinear internal models for output regulation,” IEEE Transactions on Automatic Control , AC-49, pp. 2244-2247, (2004). [15] C.I. Byrnes, F. Celani, A...approach,” IEEE Transactions on Automatic Control , 48 (Dec. 2003), 2172–2190. 2. C. I. Byrnes, “Differential Forms and Dynamical Systems,” to appear

  17. INFLUENCE ANALYSIS ON EXPONENTIAL NONLINEAR MODELS WITH RANDOM EFFECTS

    Institute of Scientific and Technical Information of China (English)

    宗序平; 赵俊; 王海斌; 韦博成

    2003-01-01

    This paper presents a unified diagnostic method for exponential nonlinear models with random effects based upon the joint likelihood given by Robinson in 1991.The authors show that the case deletion model is equivalent to mean shift outlier model.From this point of view,several diagnostic measures,such as Cook distance,score statistics are derived.The local influence measure of Cook is also presented.Numerical example illustrates that our method is available.

  18. INFLUENCE ANALYSIS IN NONLINEAR MODELS WITH RANDOM EFFECTS

    Institute of Scientific and Technical Information of China (English)

    WeiBocheng; ZhongXuping

    2001-01-01

    Abstract. In this paper,a unified diagnostic method for the nonlinear models with random ef-fects based upon the joint likelihood given by Robinson in 1991 is presented. It is shown that thecase deletion model is equivalent to the mean shift outlier model. From this point of view ,sever-al diagnostic measures, such as Cook distance, score statistics are derived. The local influencemeasure of Cook is also presented. A numerical example illustrates that the method is avail-able

  19. Filamentation with nonlinear Bessel vortices.

    Science.gov (United States)

    Jukna, V; Milián, C; Xie, C; Itina, T; Dudley, J; Courvoisier, F; Couairon, A

    2014-10-20

    We present a new type of ring-shaped filaments featured by stationary nonlinear high-order Bessel solutions to the laser beam propagation equation. Two different regimes are identified by direct numerical simulations of the nonlinear propagation of axicon focused Gaussian beams carrying helicity in a Kerr medium with multiphoton absorption: the stable nonlinear propagation regime corresponds to a slow beam reshaping into one of the stationary nonlinear high-order Bessel solutions, called nonlinear Bessel vortices. The region of existence of nonlinear Bessel vortices is found semi-analytically. The influence of the Kerr nonlinearity and nonlinear losses on the beam shape is presented. Direct numerical simulations highlight the role of attractors played by nonlinear Bessel vortices in the stable propagation regime. Large input powers or small cone angles lead to the unstable propagation regime where nonlinear Bessel vortices break up into an helical multiple filament pattern or a more irregular structure. Nonlinear Bessel vortices are shown to be sufficiently intense to generate a ring-shaped filamentary ionized channel in the medium which is foreseen as opening the way to novel applications in laser material processing of transparent dielectrics.

  20. Debt Shifting and Ownership Structure

    OpenAIRE

    Dirk Schindler; Guttorm Schjelderup

    2011-01-01

    Previous theoretical studies on the debt shifting behavior of multinationals have assumed affiliates of multinationals to be wholly owned. We develop a model that allows a multinational firm to determine both the leverage and ownership structure in affiliates endogenously. A main finding is that affiliates with minority owners have less debt than wholly owned affiliates and therefore a less tax efficient financing structure. This is due to an externality that arises endogenously in our model,...