Sample records for hemlock cicuta douglasii

  1. Toxicoses in livestock from the hemlocks (Conium and Cicuta spp.).

    Panter, K E; Keeler, R F; Baker, D C


    The hemlocks, Conium maculatum (poison-hemlock) and Cicuta spp. (waterhemlock), are poisonous plants that cause sizeable losss to the livestock industry. Clinical signs of poisonhemlock toxicosis are similar in all species of livestock and include muscular weakness, incordination, trembling, initial central nervous system stimulation, depression and death from respiratory paralysis. Poison-hemlock also causes skeletal defects in the offspring of cattle, pigs and sheep and cleft palate in pigs when ingested during specific periods of gestation. The primary toxicants in poison-hemlock are coniine and gamma-coniceine. Coniine predominates in mature plants and seed, whereas gamma-coniceine predominates in early growth of the plant. Waterhemlock is the most violently toxic poisonous plant known. The toxicant is cicutoxin, which acts on the central nervous system, causing violent convulsions and death. Clinical signs of poisoning appear within 15 min after ingestion of a lethal dose and include excessive salivation, nervousness, tremors, muscular weakness and convulsive seizures interspersed by intermittent periods of relaxation and a final paralytic seizure resulting in anoxia and death. Elevated activities of lactic dehydrogenase, aspartate aminotransferase and creatine kinase in blood are observed, indicative of muscular damage. Toxicoses from poisonhemlock and waterhemlock generally occur in early spring when both plants emerge before other, more palatable plants begin to grow. All parts of the poison-hemlock plant are toxic. The root or tubers of waterhemlock are toxic; however, experimental evidence concerning the toxicity of other plant parts is inconclusive.

  2. Genetic analysis of a Microseris douglasii (Asteraceae) population polymorphic for an alien chloroplast type

    Roelofs, Dick; Bachmann, Konrad


    Recent evidence suggests chloroplast introgression from Microseris bigelovii into M. douglasii. We have examined 23 plants from a population of M. douglasii polymorphic for M. douglasii and M. bigelovii chloroplast types. All 23 plants were completely homozygous for morphological and RAPD markers,

  3. NCRN Hemlock Inventory Data

    Department of the Interior — ​Data associated with the 2015 hemlock inventory project in NCR. Eastern hemlock (Tsuga canadensis) is a coniferous tree native to the NE and Appalachian regions of...

  4. Hemlock Dwarf Mistletoe (FIDL)

    Paul E. Hennon; Jerome S. Beatty; Diane Hildebrand


    Hemlock dwarf mistletoe, Arceuthobium tsugense (Rosendahl) G.N. Jones, causes a serious disease of western hemlock and several other tree species along the Pacific Coast of North America. This small, seed-bearing plant lives exclusively as a parasite on living trees. Throughout its range, hemlock dwarf mistletoe occurs in patch-like patterns in the forests. Some...

  5. Maintenance of eastern hemlock forests: Factors associated with hemlock vulnerability to hemlock woolly adelgid

    Mary Ann Fajvan; Petra Bohall Wood


    Eastern hemlock (Tsuga canadensis [L.]) is the most shade-tolerant and long-lived tree species in eastern North America. The hemlock woolly adelgid (Adelges tsugae) (HWA), is a nonnative invasive insect that feeds on eastern hemlock and Carolina hemlock (Tsuga caroliniana Engelm.). HWA currently is established in...

  6. Impacts of hemlock decline and ecological considerations for hemlock stand restoration following hemlock woolly adelgid outbreaks

    Sarah Z. Jonas; Weimin Xi; John D. Waldron; Robert N. Coulson


    We present a synthesis of current knowledge and information of hemlock woolly adelgid (HWA, Adelges tsugae Annand) impact on hemlock forests and conceptual framework of restoring damaged hemlock stands by HWA infestation. Native to Asia, HWA has been thriving in the eastern United States since the early 1950s and has become a serious pathological agent of both eastern...

  7. Biology and control of hemlock woolly adelgid

    Nathan P. Havill; Ligia C. Vieira; Scott M. Salom


    This publication is a substantial revision of FHTET 2001-03, Hemlock Woolly Adelgid, which was published in 2001. This publication contains information on the native range of hemlock and range of hemlock woolly adelgid, the importance of hemlocks in eastern forest ecosystems, and on hosts, life cycle, control, and population trends of the hemlock woolly adelgid.

  8. Breeding biology and bee guild of Douglas' dustymaiden, Chaenactis douglasii (Asteraceae, Helenieae)

    James H. Cane; Byron Love; Katharine Swoboda


    Douglas' dustymaiden, Chaenactis douglasii (Hook.) Hook. & Arn., is a widespread, inconspicuous, short-lived perennial wildflower that blooms in early summer and is found in basin sagesteppe to upper montane areas throughout the U.S. Intermountain West. The species is proving practical to grow for seed and is expected to be used for western rangeland...

  9. Dynamics of Connecticut hemlock stands

    Jeffrey S. Ward; David M. Smith


    The stand dynamics and production of two one-acre plots of eastern hemlock (Tsuga canadensis L) in Connecticut have been followed for more than six decades. Data were recorded for all individual trees. One plot (Saltonstall) was established in 1924 after the removal of a hardwood overstory. This stand had a nearly pure, almost fully closed understory...

  10. Using dendrochronology to model hemlock woolly adelgid effects on eastern hemlock growth and vulnerability

    James Rentch; Mary Ann Fajvan; Richard Evans; Brad. Onken


    This study examined the relationship between eastern hemlock (Tsuga canadensis (L.) Carr.) crown condition and changes in radial growth associated with infestation by hemlock woolly adelgid (HWA), Adelges tsugae (Hemiptera: Adelgidae). Tree-ring chronologies of eastern hemlock were used to develop a binomial decline index based on...

  11. Hemlock (Conium Maculatum) Poisoning In A Child.

    Konca, Capan; Kahramaner, Zelal; Bosnak, Mehmet; Kocamaz, Halil


    Poison hemlock (Conium maculatum) is a plant that is poisonous for humans and animals. Accidental ingestion of the plant may result in central nervous system depression, respiratory failure, acute rhabdomyolysis, acute renal failure and even death. The main treatment of hemlock poisoning is supportive care. The case of a 6-year-old girl who was admitted to the emergency department with complaints of burning sensation in mouth, hypersalivation, tremor in hands and ataxia after ingestion of poison hemlock is presented here with clinical and laboratory features. In this case, we aim to report that accidental ingestion of plants resembling vegetables that are consumed daily can lead to serious complications and even death.

  12. Poison hemlock (Conium maculatum L.).

    Vetter, J


    One of the most poisonous species amongst higher plants is Conium maculatum. It is a very common nitrophile weed species, belonging to the Apiaceae (formerly Umbelliferae) family. It contains some piperidine alkaloids (coniine, N-methyl-coniine, conhydrine, pseudoconhydrine, gamma-coniceine), which are formed by the cyclisation of an eight-carbon chain derived from four acetate units. gamma-Coniceine is the precursor of the other hemlock alkaloids. All vegetative organs, flowers and fruits contain alkaloids. The concentrations (both absolute and relative) of the different alkaloids depend on plant varieties, on ecological conditions and on the age of the plant. The characteristic biological effects of the plants are summarised on cattle, sheep, goat, swine, rabbit, elk, birds and insects and the symptoms of the human toxicosis (some cases of poisonings) are discussed according to the literature data. The general symptoms of hemlock poisoning are effects on nervous system (stimulation followed by paralysis of motor nerve endings and CNS stimulation and later depression), vomiting, trembling, problems in movement, slow and weak later rapid pulse, rapid respiration, salivation, urination, nausea, convulsions, coma and death.

  13. Hemlock (Conium Maculatum) Poisoning In A Child

    KONCA, Capan; KAHRAMANER, Zelal; BOSNAK, Mehmet; KOCAMAZ, Halil


    SUMMARY Poison hemlock (Conium maculatum) is a plant that is poisonous for humans and animals. Accidental ingestion of the plant may result in central nervous system depression, respiratory failure, acute rhabdomyolysis, acute renal failure and even death. The main treatment of hemlock poisoning is supportive care. The case of a 6-year-old girl who was admitted to the emergency department with complaints of burning sensation in mouth, hypersalivation, tremor in hands and ataxia after ingestio...

  14. Recovery of hemlock in Vermont from defoliation by the spring hemlock looper, Lambdina athasaria (Walker)

    Barbara S. Burns; Henry, Jr. Trial


    Following an outbreak of spring hemlock looper in 1991, ten hemlocks in each of fifteen study plots were monitored annually through 1999. Although some mortality occurred within two years after defoliation, and additional mortality occurred in plots which were subsequently disturbed by logging, most defoliated trees recovered. Twenty-four percent of the trees with 90%...

  15. Influences of eastern hemlock mortality on nutrient cycling

    Thad E. Yorks; Jennifer C. Jenkins; Donald J. Leopold; Dudley J. Raynal; David A. Orwig


    Mortality of eastern hemlock (Tsuga canadensis (L.) Carriere) may be caused by a variety of agents, but hemlock trees of all sizes over a large geographic area are currently threatened by an outbreak of the hemlock woolly adelgid (HWA: Adelges tsugae Annand) in the eastern United States. In this paper, we review what is currently...

  16. Hemlock (Conium Maculatum Poisoning In A Child

    Capan KONCA


    Full Text Available Poison hemlock (Conium maculatum is a plant that is poisonous for humans and animals. Accidental ingestion of the plant may result in central nervous system depression, respiratory failure, acute rhabdomyolysis, acute renal failure and even death. The main treatment of hemlock poisoning is supportive care. The case of a 6-year-old girl who was admitted to the emergency department with complaints of burning sensation in mouth, hypersalivation, tremor in hands and ataxia after ingestion of poison hemlock is presented here with clinical and laboratory features. In this case, we aim to report that accidental ingestion of plants resembling vegetables that are consumed daily can lead to serious complications and even death.

  17. Western hemlock as a veneer resource.

    Thomas D. Fahey; Jr. Woodfin


    Presents recovery of veneer grade and volume from western hemlock from Oregon and Washington. Veneer grade recovery varied by grade and size of logs. Veneer volume recovered was about 45 percent of the cubic volume of the log and varied somewhat with log diameter.

  18. Prioritizing conservation seed banking locations for imperiled hemlock species using multi-attribute frontier mapping

    John M. Hastings; Kevin M. Potter; Frank H. Koch; Mark Megalos; Robert M. Jetton


    Hemlock woolly adelgid (Adelges tsugae Annand) (HWA) is an invasive forest insect sweeping across the native range of eastern (Tsuga canadensis [L.] Carr.) and Carolina (Tsuga caroliniana Engelm.) hemlocks, threatening to severely reduce eastern hemlock extent and to push Carolina hemlock to extirpation. HWA poses a significant threat to these eastern US natives, now...

  19. Hemlock woolly adelgid in the southern Appalachians: Control strategies, ecological impacts, and potential management responses

    James M. Vose; David N. Wear; Albert E. Mayfield; C. Dana Nelson


    Hemlock woolly adelgid (Adelges tsugae Annad; or HWA) is a non-native invasive pest that attacks and kills eastern hemlock (Tsuga canadensis (L.) Carrière) and Carolina hemlock (Tsuga caroliniana Engelm.). Hemlock is a ‘‘foundation species’’ due to its strong influence on ecosystem structure and function,...

  20. Early effect of two successive thinnings in western hemlock.

    George R. Staebler


    The Hemlock Experimental Forest near Grays Harbor in western Washington was established in 1949 in cooperation with the St. Regis Paper Company. A major effort in this cooperative research program is a study of commercial thinning in a stand of nearly pure, well stocked, even-aged western hemlock that originated in 1903, after logging.

  1. Physiological responses of eastern hemlock (Tsuga Canadensis) to biological control and silvicultural release: implications for hemlock restoration

    Chelcy F. Miniat; David Zeitlow; Steven T. Brantley; Albert (Bud) Mayfield; Rusty Rhea; Robert Jetton; Paul.  Arnold


    The rapid loss of eastern hemlock (Tsuga canadensis) from riparian zones in the southern Appalachian Mountains due to Hemlock Woolly Adelgid (Adelgis tsugae, HWA) infestation has resulted in changes to watershed structure and function. Several restoration strategies have been proposed, including silvicultural treatments that increase incident light in forest...

  2. A landscape-scale remote sensing/GIS tool to assess eastern hemlock vulnerability to hemlock woolly adelgid-induced decline

    Jennifer Pontius; Richard Hallett; Mary Martin; Lucie. Plourde


    The hemlock woolly adelgid (Adelges tsugae Annand) (HWA) is an invasive insect pest that is causing widespread mortality of eastern hemlock (Tsuga canadensis (L.) Carr.). However, some stands remain living more than a decade after infestation. The ability to target management efforts in locations where hemlock is most likely to...

  3. Successful transplantation of donor organs from a hemlock poisoning victim.

    Foster, Preston F; McFadden, Robert; Trevino, Raul; Galliardt, Scott; Kopczewski, Lea Ann; Gugliuzza, Kristene; Gonzalez, Zulma; Wright, Francis


    The poison hemlock plant (Conium maculatum) has been a known poison since early in human history, most notably as the agent used for the execution/suicide of Socrates in ancient Greece. No experience has been reported regarding the suitability of a hemlock victim's organs for transplantation. This report documents successful transplantation of the liver, kidney, and pancreas from a 14-year-old girl who died of anoxic encephalopathy from asphyxia after the accidental ingestion of fresh hemlock while on a nature hike. Predonation laboratory values were not remarkable, and liver and kidney biopsy results were normal. All organs in the three recipients had immediate function, and no recipient had any clinical evidence of transmitted toxin. All recipients are well, with functioning transplants at greater than 6 months after transplantation. Poison hemlock intoxication does not seem to be a contraindication to organ donation.

  4. Water use and carbon exchange of red oak- and eastern hemlock-dominated forests in the northeastern USA : implications for ecosystem-level effects of hemlock woolly adelgid

    Hadley, J.L.; Kuzeja, P.S.; Singh, S.


    This study used the eddy flux method to measure water use and carbon exchange of a red oak forest and an eastern hemlock-dominated forest located in north-central Massachusetts over a period of 2 years. The study demonstrated that water use by the red oak reached approximately 4 mm per day -1 . A maximum rate of 2 mm per day -1 was used by the eastern hemlock forest. Carbon (C) uptake rates were higher in the red oak forest than in the eastern hemlock forest. Measurements of sap flux suggested that transpiration of red oak and black birches in the eastern hemlock forest were approximately twice as high as transpiration rates observed for eastern hemlock. However, annual C storage of eastern hemlock was almost equal to C storage rates of the red oak forest, due to net C uptake by the hemlock during an unusually warm fall and spring. The study showed that C storage by eastern hemlock forests will increase as a result of climatic warming. Although forest water use will decrease as a result of eastern hemlock due to insect disturbances, the replacement of eastern hemlock by deciduous species such as red oak will increase water use during the summer-time in areas where hemlock is a predominant species. 24 refs., 5 tabs., 11 figs

  5. Allozyme Variation in Hemlock Woolly Adelgid (Hemiptera: Adelgidae) from the United States and China

    V. S& #225; nchez; M.A. Keena; M.A. Keena


    The hemlock woolly adelgid, Adelges tsugae Annand (Hemiptera: Adelgidae), is a major introduced pest of eastern hemlock, Tsuga canadensis (L.) Carriere. Hemlock woolly adelgid in the United States is anholocyclic and an obligate parthenogen, because no suitable primary host (on which sexual reproduction occurs in Asia) is...

  6. Distribution and Abundance of Adelges tsugae (Hemiptera: Adelgidae) Within Hemlock Trees

    S.V. Joseph; J.L. Hanula; S.K. Braman


    We studied the distribution of hemlock woolly adelgid, Adelges tsugae Annand (Hemiptera: Adelgidae), within hemlock trees for three summer (progrediens) and two winter (sistens) generations in northern Georgia. Eastern hemlock, Tsuga canadensis (L.) Carrie` re, trees were treated with 0, 10, or 25% of 1.5 g of imidacloprid per 2.5 cm of tree diameter at breast height...

  7. Use of water by eastern hemlock: implications for systemic insecticide application

    Chelcy R. Ford; James M. Vose; Michael Daley; Nathan Phillips


    The hemlock woolly adelgid (HWA, Adelges tsugae Annand) is causing widespread decline and mortality of eastern hemlock (Tsuga canadensis (L.) Carr.) throughout most of the range of eastern hemlock. Stem injection of insecticide is widely used as a chemical control measure, but the effectiveness of this method depends on the...

  8. Classification and spatial analysis of eastern hemlock health using remote sensing and GIS

    Laurent R. Bonneau; Kathleen S. Shields; Daniel L. Civco; David R. Mikus


    Over the past decade hemlock stands in southern Connecticut have undergone significant decline coincident with the arrival in 1985 of an exotic insect pest, the hemlock woolly adelgid (Adelges tsugae Annand). The objective of this study was to evaluate image enhancement techniques for rating the health of hemlocks at the landscape level using...

  9. Growth and stocking of eastern hemlock (Tsuga canadensis) in New England

    Dale S. Solomon; William B. Leak


    Summarization of the limited growth information in mixed-species stands in New England indicates that eastern hemlock (Tsuga canadensis) may be one of the fastest growing species in diameter, second only to white pine. However, on some sites hemlock diameter growth is about equal to that of associated hardwoods. Hemlock grows slowly in height and...

  10. Impact of Imidacloprid for control of Hemlock Woolly Adelgid on nearby aquatic macroinvertebrate asseblages.

    Melissa Churchel; James Hanula; C. Wayne Berisford; James Vose; Mark Dalusky


    Imidacloprid, a systemic insecticide that acts on the nervous system, is currently being used to control hemlock woolly adelgid (Adelges tsugae Annand), which is damaging hemlock trees. The objective of this study was to determine whether soil injection with imidacloprid for hemlock woolly adelgid control near streams adversely affects aquatic invertebrates. Eastern...

  11. Silvicultural and integrated pest management strategies for restoring eastern hemlock to degraded southern Appalachian mountain ecosystems.

    W.A. Whittier; A.E. Mayfield III; R.M. Jetton


    The ecologically foundational species eastern hemlock, Tsuga canadensis, is being functionally eliminated from southern Appalachian forests by the hemlock woolly adelgid (HWA, Adelges tsugae). The management of HWA has focused on chemical and biological control, conservation of hemlock genetic resources, and host resistance...

  12. Hemlock alkaloids from Socrates to poison aloes.

    Reynolds, Tom


    Hemlock (Conium maculatum L. Umbelliferae) has long been known as a poisonous plant. Toxicity is due to a group of piperidine alkaloids of which the representative members are coniine and gamma-coniceine. The latter is the more toxic and is the first formed biosynthetically. Its levels in relation to coniine vary widely according to environmental conditions and to provenance of the plants. Surprisingly, these piperidine alkaloids have turned up in quite unrelated species in the monocotyledons as well as the dicotyledons. Aloes, for instance, important medicinal plants, are not regarded as poisonous although some species are very bitter. Nevertheless a small number of mostly local species contain the alkaloids, especially gamma-coniceine and there have been records of human poisoning. The compounds are recognized by their characteristic mousy smell. Both acute and chronic symptoms have been described. The compounds are neurotoxins and death results from respiratory failure, recalling the effects of curare. Chronic non-lethal ingestion by pregnant livestock leads to foetal malformation. Both acute and chronic toxicity are seen with stock in damp meadows and have been recorded as problems especially in North America. The alkaloids derive biosynthetically from acetate units via the polyketide pathway in contrast to other piperidine alkaloids which derive from lysine.

  13. Grading options for western hemlock "pulpwood" logs from southeastern Alaska.

    David W. Green; Kent A. McDonald; John Dramm; Kenneth Kilborn

    Properties and grade yield are estimated for structural lumber produced from No. 3, No. 4, and low-end No. 2 grade western hemlock logs of the type previously used primarily for the production of pulp chips. Estimates are given for production in the Structural Framing, Machine Stress Rating, and Laminating Stock grading systems. The information shows that significant...

  14. Initial survey of predacious diptera on hemlocks in Japan

    Hisashi Ohishi; Shigehiko Shiyake; Yorio Miyatake; Ashley Lamb; Michael E. Montgomery


    Some species of Coleoptera and Diptera are specialist predators of adelgids. Previously, we reported our survey of predacious Coleoptera on hemlocks in Japan (Shiyake et al. 2008). Two of these beetles, Sasajiscymnus tsugae and Laricobius sp. nov., have been exported to the U.S. for biological control. Here, we provide the first...

  15. True fir-hemlock spacing trials: design and first results.

    Robert O. Curtis; Gary W. Clendenen; Jan A. Henderson


    A series of 18 precommercial thinning trials was established in true fir-hemlock stands in the Olympic Mountains and along the west side of the Cascade Range in Washington and Oregon from 1987 through 1994. This paper documents establishment of these installations and presents some preliminary observations and results. Substantial differences in growth rates in height...

  16. Vascular plant propagule banks of six eastern hemlock stands and potential response to the hemlock woolly adelgid in the Catskill Mountains of New York

    Thad E. Yorks; Donald J. Leopold; Dudley J. Raynal


    We examined propagule banks in six eastern hemlock (Tsuga canadensis (L.) Carriere) stands in the Catskill Mountains of New York. These stands are at risk of mortality due to the hemlock woolly adelgid (Adelges tsugae Annand), but potential effects of mortality on species composition are uncertain.

  17. Intravenous Poison Hemlock Injection Resulting in Prolonged Respiratory Failure and Encephalopathy.

    Brtalik, Douglas; Stopyra, Jason; Hannum, Jennifer


    Poison hemlock (Conium maculatum) is a common plant with a significant toxicity. Data on this toxicity is sparse as there have been few case reports and never a documented poisoning after intravenous injection. We present a case of intravenous poison hemlock injection encountered in the emergency department. We describe a 30-year-old male who presented to the emergency department after a brief cardiac arrest after injecting poison hemlock. The patient had return of spontaneous circulation in the emergency department but had prolonged muscular weakness and encephalopathy later requiring tracheostomy. Intravenous injection of poison hemlock alkaloids can result in significant toxicity, including cardiopulmonary arrest, prolonged weakness, and encephalopathy.

  18. Evaluation of hemlock (Tsuga) species and hybrids for resistance to Adelges tsugae (Hemiptera: Adelgidae) using artificial infestation

    Michael E. Montgomery; S.E. Bentz; Richard T. Olsen


    Hemlock (Tsuga) species and hybrids were evaluated for resistance to the hemlock woolly adelgid, Adelges tsugae Annand (Hemiptera: Adelgidae). The adelgid was accidentally introduced from Asia to the eastern United States, where it is causing widespread mortality of the native hemlocks, Tsuga canadensis (L.)...

  19. Orientation behavior of the predator Laricobius nigrinus (Coleoptera: Derodontidae) to hemlock woolly adelgid and host tree odors in a multi-chambered olfactometer

    Kimberly F. Wallin; Tanya M. Latty; Darrell W. Ross


    We studied the adult ambulatory response of the predator, Laricobius nigrinus Fender (Coleoptera: Derodontidae), to odors from its prey, Adelges tsugae Annand, the hemlock woolly adelgid, and foliage of hemlock woolly adelgid, host hemlocks (Tsuga spp.), and other conifers. Both the predator and hemlock woolly...

  20. The range and response of Neonicotinoids on hemlock woolly adelgid, Adelges tsugae (Hemiptera: Adelgidae)

    Shimat V. Joseph; S. Kristine Braman; Jim Quick; James L. Hanula


    Hemlock woolly adelgid (HWA), Adelges tsugae Annand is a serious pest of eastern and Carolina hemlock in the eastern United States. A series of experiments compared commercially available and experimental insecticides, rates, application methods and timing for HWA control in Georgia and North Carolina. Safari 20 SG (dinotefuran) provided an average of 79 to 87%...

  1. Phenology of the Hemlock Woolly Adelgid (Hemiptera: Adelgidae) in Northern Georgia

    Shimar V. Joseph; Albert E. Mayfield; Mark J. Dalusky; Christopher Asaro; C. Wayne. Berisford


    Understanding the seasonal phenology of an insect pest in a specific geographic region is essential for optimizing the timing of management actions or research activities. We examined the phenology of hemlock woolly adelgid, Adelges tsugae Annand, near the southern limit of the range of eastern hemlock, Tsuga canadensis (L.) Carriere, in the Appalachians of northern...

  2. Imidacloprid movement in soils and impacts on soil microarthropods in southern Appalachian eastern hemlock stands

    Jennifer D. Knoepp; James M. Vose; Jerry L. Michael; Barbara C. Reynolds


    Imidacloprid is a systemic insecticide effective in controlling the exotic pest Adelges tsugae (hemlock woolly adelgid) in eastern hemlock (Tsuga canadensis) trees. Concerns over imidacloprid impacts on nontarget species have limited its application in southern Appalachian ecosystems. We quantified the movement and adsorption of imidacloprid in forest soils after soil...

  3. Chinese Coccinellidae for biological control of the hemlock woolly adelgid: description of native habitat

    Michael E. Montgomery; Defu Yao; Hongbin Wang


    The hemlock woolly adelgid, Adelges tsugae Annand, is generally believed to be native to Asia and is known to occur in India, Japan, and China. In China, there are approximately four species of hemlock that grow in 14 provinces. We explored regions of the Southwestern Plateau in Yunnan, Sichuan, and Shaanxi Provinces for the adelgid and its natural...

  4. Hemlock resources at risk in the Great Smoky Mountains National Park

    Kristine D. Johnson; Fred P. Hain; Katherine S. Johnson; Felton Hastings


    Eastern hemlock (Tsuga canadensis (L.) Carr) is the dominant species in a variety of sites in Great Smoky Mountains National Park. Hemlock covers approximately 3820 acres (1528 hectares) or one percent of the Park, which at 524,856 acres is the largest area managed as wilderness in the eastern United States. Since timber was never harvested in about...

  5. Use of satellite image data to identify changes in hemlock health over space and time

    Laurent R. Bonneau; Kathleen S. Shields; Daniel L. Civco; David R. Mikus


    Eastern hemlock (Tsuga canadensis (L.) Carriere), is an important component of ecosystems in the northeastern United States and is the primary coniferous species in southern Connecticut. Hemlocks play a unique role in the region by providing spatial and structural habitat diversity that supports many wildlife and fish populations. Widespread damage...

  6. Management Strategies for Annosus Root Disease in Pacific Northwest Coastal Western Hemlock

    Kenelm W. Russell


    Actual loss from annosus root disease infections in hemlock stands is difficult to determine. As political trends move toward protecting old-growth timber, greater market demand will be placed on second growth western hemlock. These stands must be kept healthy for maximum productivity. The paper compares the following 70-year rotation timber management scenarios: The...

  7. Vegetation and invertebrate community response to eastern hemlock decline in southern new England

    Laura L. Ingwell; Mailea Miller-Pierce; R. Talbot Trotter; Evan L. Preisser


    The introduction of Adelges tsugae (Hemlock Woolly Adelgid [HWA]) to the eastern United States has had a devastating impact on Tsuga canadensis (Eastern Hemlock). Although much research has been done to assess HWA impacts on ecosystem processes and vegetation structure, few researchers have examined community-level changes in...

  8. Abundance of Armillaria within old-growth eastern hemlock stands in South-Central Pennsylvania

    Matthew S. Fromm; Donald D. Davis


    Abstract—In early summer 2002, 329 soil-sampling pits were dug within an old-growth, eastern hemlock (Tsuga canadensis [L.] Carrière) stand in south-central Pennsylvania recently infested with the hemlock woolly adelgid (Adelges tsugae Annand). For comparison, 199 similar pits were dug in an adjacent hardwood stand. Rhizomorphs of...

  9. Xylem transport models optimize effectiveness of systemic insecticide applications for controlling hemlock woolly adelgid (Adelges tsugae)

    Chelcy R. Ford; Barbara C. Reynolds; James Vose


    The hemlock woolly adelgid (HWA, Adelges tsugae Annand) is causing widespread decline and mortality of eastern hemlock trees (Tsuga Canadensis (L.) Carr.). Stem injection of insecticide is widely used as a control measure, but its effectiveness depends on individual tree hydraulic characteristics. Recent work has shown that eastern...

  10. Stylet bundle morphology and trophically related enzymes of the hemlock woolly adelgid (Hemiptera: Adelgidae)

    Kelly L.F. Oten; Allen C. Cohen; Fred P. Hain


    The hemlock woolly adelgid, Adelges tsugae Annand (Hemiptera: Adelgidae), is a pest of eastern and Carolina hemlocks (Tsuga canadensis (L.) Carriere and Tsuga caroliniana Engelmann, respectively) in the eastern United States and has already caused catastrophic changes to eastern forests. As one of the significant...

  11. Using climate and genetic diversity data to prioritize conservation seed banking for imperiled hemlock species

    J.M. Hastings; K.M. Potter; F.H. Koch; M.A. Megalos; R.M. Jetton


    Hemlock woolly adelgid (HWA, Adelges tsugae) is an invasive forest insect that has caused mortality of eastern (Tsuga canadensis) and Carolina hemlock (T. caroliniana) at an alarming rate. Now infesting 19 states and over 400 counties of the eastern United States, HWA poses a significant threat to native host species. The current biological and chemical methods for...

  12. 77 FR 46373 - Availability of an Environmental Assessment for a Biological Control Agent for Hemlock Woolly...


    ...] Availability of an Environmental Assessment for a Biological Control Agent for Hemlock Woolly Adelgid AGENCY... States for use as a biological control agent to reduce the severity of hemlock woolly adelgid... beetle from the western United States, into the eastern United States for use as a biological control...

  13. 75 FR 28232 - Availability of an Environmental Assessment for a Biological Control Agent for Hemlock Woolly...


    ...] Availability of an Environmental Assessment for a Biological Control Agent for Hemlock Woolly Adelgid AGENCY..., into the continental United States for use as a biological control agent to reduce the severity of... biological control agent to reduce the severity of hemlock woolly adelgid (HWA) infestations. HWA, an...

  14. Hemlock woolly adelgid (Homoptera: Adelgidae): stylet bundle insertion and feeding sites

    Rebecca F. Young; Kathleen S. Shields; Graeme P. Berlyn


    Stylet bundle insertion site, path traveled, and feeding site were examined for the hemlock woolly adelgid, Adelges tsugae Annand, on needles from current and previous years of eastern hemlock, Tsuga canadensis Carriere. The stylet bundle is composed of 4 individual stylets--2 outer mandibular stylets and 2 inner maxillary stylets...

  15. Influence of Hemlock Woolly Adelgid Infestation Levels on Water Stress in Eastern Hemlocks within the Great Smoky Mountains National Park, U.S.A.

    Carla Coots


    Full Text Available Extensive mortality of eastern hemlock, Tsuga canadensis (L. Carrière, resulting from infestation by hemlock woolly adelgid, Adelges tsugae Annand (Hemiptera: Adelgidae, has occurred throughout the eastern United States. Although imidacloprid treatment can reduce tree mortality, its effectiveness can be influenced by several factors including tree water stress. The relationship between water stress and infestation rates is unknown, and an understanding of these could greatly increase the efficiency of management for this invasive insect. The primary objective of this study was to assess water stress at three levels of hemlock woolly adelgid infestations. Water stress was measured monthly for 13 months in eastern hemlocks classified as <25%, 25%–75%, and >75% infested. The highest level of water stress was found in those trees with hemlock woolly adelgid infestation levels greater than 75%. The lowest level of water stress was found in those trees with less than 25% hemlock woolly adelgid infestation levels. Knowledge of these effects can contribute to development of more effective chemical management strategies.

  16. Poison hemlock-induced respiratory failure in a toddler.

    West, Patrick L; Horowitz, B Zane; Montanaro, Marc T; Lindsay, James N


    The ingestion of poison hemlock, or Conium maculatum, is described in a 2-year-old boy. He had the onset of abdominal pain and weakness after being fed C. maculatum picked by his sister from the roadside 2 hours earlier. He had a rapidly progressive muscular weakness and was intubated for respiratory failure. His symptoms completely resolved within 24 hours of the ingestion. Conium maculatum is a common weed that causes toxicity by its primary toxin, coniine, which stimulates nicotinic receptors and causes a syndrome of rapidly progressive muscle weakness and paralysis. We describe the course of a benign-appearing plant ingestion resulting in respiratory failure.

  17. Polyketide synthases from poison hemlock (Conium maculatum L.).

    Hotti, Hannu; Seppänen-Laakso, Tuulikki; Arvas, Mikko; Teeri, Teemu H; Rischer, Heiko


    Coniine is a toxic alkaloid, the biosynthesis of which is not well understood. A possible route, supported by evidence from labelling experiments, involves a polyketide formed by the condensation of one acetyl-CoA and three malonyl-CoAs catalysed by a polyketide synthase (PKS). We isolated PKS genes or their fragments from poison hemlock (Conium maculatum L.) by using random amplification of cDNA ends (RACE) and transcriptome analysis, and characterized three full-length enzymes by feeding different starter-CoAs in vitro. On the basis of our in vitro experiments, two of the three characterized PKS genes in poison hemlock encode chalcone synthases (CPKS1 and CPKS2), and one encodes a novel type of PKS (CPKS5). We show that CPKS5 kinetically favours butyryl-CoA as a starter-CoA in vitro. Our results suggest that CPKS5 is responsible for the initiation of coniine biosynthesis by catalysing the synthesis of the carbon backbone from one butyryl-CoA and two malonyl-CoAs. © 2015 FEBS.

  18. Identification of host fruit volatiles from domestic apple (Malus domestica), native black hawthorn (Crataegus douglasii) and introduced ornamental hawthorn (C. monogyna) attractive to Rhagoletis pomonella flies from the western United States.

    Cha, Dong H; Yee, Wee L; Goughnour, Robert B; Sim, Sheina B; Powell, Thomas H Q; Feder, Jeffrey L; Linn, Charles E


    The apple maggot fly, Rhagoletis pomonella, infests apple (Malus domestica) and hawthorn species (most notably the downy hawthorn, Crataegus mollis) in the eastern USA. Evidence suggests that the fly was introduced into the western USA sometime in the last 60 years. In addition to apple, R. pomonella also infests two species of hawthorns in the western USA as major hosts: the native black hawthorn (C. douglasii) and the introduced ornamental English hawthorn, C. monogyna. Apple and downy hawthorn-origin flies in the eastern USA use volatile blends emitted from the surface of their respective ripening fruit to find and discriminate among host trees. To test whether the same is true for western flies, we used coupled gas chromatography and electroantennographic detection (GC-EAD) and developed a 7-component apple fruit blend for western apple-origin flies, an 8-component black hawthorn fruit blend for flies infesting C. douglasii, and a 9-component ornamental hawthorn blend for flies from C. monogyna. Crataegus douglasii and C. monogyna-origin flies showed similar levels of upwind directed flight to their respective natal synthetic fruit blends in flight tunnel assays compared to whole fruit adsorbent extracts, indicating that the blends contain all the behaviorally relevant fruit volatiles to induce maximal response levels. The black and ornamental hawthorn blends shared four compounds in common including 3-methylbutan-1-ol, which appears to be a key volatile for R. pomonella populations in the eastern, southern, and western USA that show a preference for fruit from different Crataegus species. However, the blends also differed from one another and from domesticated apple in several respects that make it possible that western R. pomonella flies behaviorally discriminate among fruit volatiles and form ecologically differentiated host races, as is the case for eastern apple and hawthorn flies.

  19. Scymnus camptodromus (Coleoptera: Coccinellidae) larval development and predation of hemlock woolly adelgid (Hemiptera: Adelgidae)

    Samita Limbu; Melody A. Keena; David Long; Nancy Ostiguy; Kelli. Hoover


    Development time and prey consumption of Scymnus (Neopullus) camptodromus Yu and Liu (Coleoptera: Coccinellidae) larvae by instar, strain, and temperature were evaluated. S. camptodromus, a specialist predator of hemlock woolly adelgid Adelges tsugae (Annand) (Hemiptera:...

  20. Conium maculatum (poison hemlock) toxicosis in a flock of range turkeys.

    Frank, A A; Reed, W M


    Five 20-week-old tom turkeys from a flock of range turkeys were presented for examination; the flock had a history of salivation, tremors, paralysis, and increased mortality. Necropsy revealed numerous seeds identified as seeds from Conium maculatum (poison hemlock) within the crop, proventriculus, and gizzard. Histopathologic alterations were limited to catarrhal enteritis. Clinical signs of Conium maculatum toxicosis abated after the turkeys were removed from their range, which was infested with poison hemlock.

  1. Influence of eastern hemlock (Tsuga canadensis L.) on fish community structure and function in headwater streams of the Delaware River basin

    Ross, R.M.; Bennett, R.M.; Snyder, C.D.; Young, J.A.; Smith, D.R.; Lemarie, D.P.


    Hemlock (Tsuga canadensis) forest of the eastern U.S. are in decline due to invasion by the exotic insect hemlock woolly adelgid (Adelges tsugae). Aquatic biodiversity in hemlock ecosystems has not been documented; thus the true impact of the infestation cannot be assessed. We compared ichthyofaunal assemblages and trophic structure of streams draining hemlock and hardwood forests by sampling first- and second-order streams draining 14 paired hemlock and hardwood stands during base flows in July 1997 at the Delaware Water Gap National Recreation Area of Pennsylvania and New Jersey. Over 1400 fish of 15 species and 7 families were collected, but hemlock and hardwood streams individually harbored only one to four species. Brook trout (Salvelinus fontinalis) and brown trout (Salmo trutta) were two to three times as prevalent in hemlock than hardwood streams. Insectivorous fishes occurred in significantly higher proportion in streams of hardwood (0.90) than hemlock (0.46) stands, while piscivores occurred more often in hemlock (0.85) than hardwood (0.54) stands. Functional (trophic) diversity of fishes in hemlock and second-order streams was numerically greater than that of hardwood and first-order streams. Species composition also differed by stream order and terrain type. Biodiversity is threatened at several levels within hemlock ecosystems at risk to the hemlock woolly adelgid in eastern U.S. forests.

  2. Multivariate statistical analysis of hemlock (Tsuga) volatiles by SPME/GC/MS: insights into the phytochemistry of the hemlock woolly adelgid (Adelges tsugae Annand)

    Anthony Lagalante; Frank Calvosa; Michael Mirzabeigi; Vikram Iyengar; Michael Montgomery; Kathleen Shields


    A previously developed single-needle, SPME/GC/MS technique was used to measure the terpenoid content of T. canadensis growing in a hemlock forest at Lake Scranton, PA (Lagalante and Montgomery 2003). The volatile terpenoid composition was measured over a 1-year period from June 2003 to May 2004 to follow the annual cycle of foliage development from...

  3. Response of Red-Backed Salamanders (Plethodon Cinereus to Changes in Hemlock Forest Soil Driven by Invasive Hemlock Woolly Adelgid (Adelges Tsugae

    Alison Ochs


    Full Text Available Hemlock forests of the northeastern United States are declining due to the invasive hemlock woolly adelgid (HWA (Adelges tsugae. Hardwood species replace these forests, which affects soil properties that may influence other communities, such as red-backed salamanders (red-backs (Plethodon cinereus. This study examined the effects of HWA invasion on soil properties and how this affects red-backs at the Hemlock Removal Experiment at Harvard Forest, which consists of eight 0.8 ha plots treated with girdling to simulate HWA invasion, logging to simulate common management practices, or hemlock- or hardwood-dominated controls. Coverboard surveys were used to determine the relative abundance of red-backs between plots during June and July 2014 and soil cores were collected from which the bulk density, moisture, pH, temperature, leaf litter, and carbon-nitrogen ratio were measured. Ordination provided a soil quality index based on temperature, pH, and carbon-to-nitrogen ratio, which was significantly different between plot treatments (p < 0.05 and showed a significant negative correlation with the red-back relative abundance (p < 0.05. The findings support the hypothesis that red-backs are affected by soil quality, which is affected by plot treatment and thus HWA invasion. Further studies should explore how salamanders react in the long term towards changing environments and consider the use of red-backs as indicator species.

  4. Population isolation results in low genetic variation and high differentiation in Carolina hemlock (tsuga caroliniana), an imperiled southern Appalachian conifer

    Kevin M. Potter; Lia Campbell; Sedley A. Josserand; C. Dana Nelson; Robert M. Jetton


    Carolina hemlock (Tsuga caroliniana) is a rare conifer species that grows in small, isolated populations in the southern Appalachian Mountains of Virginia, North Carolina, South Carolina, Tennessee, and Georgia. The species is additionally imperiled by the hemlock woolly adelgid (Adelges tsugae), an invasive insect that can...

  5. Effects of fertilization of four hemlock species on Adelges tsugae (Hemiptera: Adelgidae) growth and feeding preference of predators.

    S.V. Joseph; James Hanula


    Understanding how fertilization affects host resistance to hemlock woolly adelgid, Adelges tsugae Annand (Hemiptera:Adelgidae), is important because fertilizers are often used to grow resistant selections to a suitable size for testing. We evaluated four hemlock species (Tsuga) under three different fertilizer regimes to assess whether fertility affected resistance to...

  6. Variation in winter survival of the invasive hemlock woolly adelgid (Hemiptera: Adelgidae) across the eastern United States

    R. Talbot, III Trotter; Kathleen S. Shields


    The hemlock woolly adelgid (Adelges tsugae Annand) is a small, aphid-like insect native to East Asia and western North America. First documented in the eastern United States in Richmond, VA, in 1951, it has spread to at least 17 states, where it causes increased mortality among both eastern and Carolina hemlocks (Tsuga canadensis...

  7. Spatial and population characteristics of dwarf mistletoe infected trees in an old-growth Douglas-fir - western hemlock forest.

    David C. Shaw; Jiquan Chen; Elizabeth A. Freeman; David M. Braun


    We investigated the distribution and severity of trees infected with western hemlock dwarf mistletoe (Arceuthobium tsugense (Rosendahl) G.N. Jones subsp. tsugense) in an old-growth Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) - western hemlock (Tsuga heterophylla (Raf.) Sarg.)...

  8. Impacts of Hemlock Loss on Nitrogen Retention Vary with Soil Nitrogen Availability in the Southern Appalachian Mountains

    Corinne E. Block; Jennifer D. Knoepp; Katherine J. Elliott; Jennifer M. Fraterrigo


    The impacts of exotic insects and pathogens on forest ecosystems are increasingly recognized, yet the factors influencing the magnitude of effects remain poorly understood. Eastern hemlock (Tsuga canadensis) exerts strong control on nitrogen (N) dynamics, and its loss due to infestation by the hemlock woolly adelgid (Adelges tsugae...

  9. Influence of Imidacloprid and Horticultural Oil on Spider Abundance on Eastern Hemlock in the Southern Appalachians.

    Hakeem, A; Grant, J F; Lambdin, P L; Hale, F A; Rhea, J R; Wiggins, G J; Coots, C


    Hemlock woolly adelgid, Adelges tsugae Annand (Hemiptera: Adelgidae), is an exotic pest of eastern hemlock, Tsuga canadensis (L.) Carrière (Pinales: Pinaceae), in the eastern United States. Two commonly used insecticides to manage adelgid are imidacloprid, a systemic neonicotinoid insecticide, and horticultural oil, a refined petroleum oil foliar spray. We have investigated the influence of imidacloprid and horticultural oil on spider abundance at different canopy strata in eastern hemlock. In total, 2,084 spiders representing 11 families were collected from the canopies of eastern hemlock. In beat-sheet and direct observation samples, the families Theridiidae, Araneidae, Salticidae, and Anyphaenidae were the most abundant. Significantly higher numbers of spiders were recorded on untreated control trees compared with trees treated with imidacloprid using soil drench and soil injection applications. Spider abundance in trees injected with imidacloprid and horticultural oil applications did not significantly differ from control trees. Spider abundance was significantly greater in the top and middle strata of the canopy than in the bottom stratum, where imidacloprid concentrations were the highest. Regression analysis showed that spider abundance was inversely associated with imidacloprid concentration. This research demonstrates that imidacloprid, when applied with selected methods, has the potential to result in reductions of spider densities at different strata. However, slight reductions in spider abundance may be an acceptable short-term ecological impact compared with the loss of an untreated hemlock and all the associated ecological benefits that it provides. Future studies should include investigations of long-term impact of imidacloprid on spiders associated with eastern hemlock.

  10. Assessment of Imidacloprid and Its Metabolites in Foliage of Eastern Hemlock Multiple Years Following Treatment for Hemlock Woolly Adelgid, Adelges tsugae (Hemiptera: Adelgidae), in Forested Conditions.

    Benton, E P; Grant, J F; Webster, R J; Nichols, R J; Cowles, R S; Lagalante, A F; Coots, C I


    Widespread decline and mortality of eastern hemlock, Tsuga canadensis (L.) Carrière, have been caused by hemlock woolly adelgid, Adelges tsugae (Annand) (HWA) (Hemiptera: Adelgidae). The current study is a retrospective analysis conducted in collaboration with Great Smoky Mountains National Park (GRSM) to determine longevity of imidacloprid and its insecticidal metabolites (imidacloprid olefin, 5-hydroxy, and dihydroxy) in GRSM's HWA integrated pest management (IPM) program. Foliage samples were collected from three canopy strata of hemlocks that were given imidacloprid basal drench treatments 4-7 yr prior to sampling. Foliage was analyzed to assess concentrations in parts per billion (ppb) of imidacloprid and its metabolites. Imidacloprid and its olefin metabolite were present in most, 95 and 65%, respectively, branchlets 4-7 yr post-treatment, but the 5-hydroxy and dihydroxy metabolites were present in only 1.3 and 11.7%, respectively, of the branchlets. Imidacloprid and olefin concentrations significantly decreased between 4 and 7 yr post-treatment. Concentrations of both imidacloprid and olefin were below the LC50 for HWA 5-7 yr post-treatment. Knowledge of the longevity of imidacloprid treatments and its metabolite olefin can help maximize the use of imidacloprid in HWA IPM programs. © The Authors 2015. Published by Oxford University Press on behalf of Entomological Society of America. All rights reserved. For Permissions, please email:

  11. Sharing rotting wood in the shade: ectomycorrhizal communities of co-occurring birch and hemlock seedlings

    Sarah K. Poznanovic; Erik A. Lilleskov; Christopher R. Webster


    Coarse woody debris (CWD) is an important nursery environment for many tree species. Understanding the communities of ectomycorrhizal fungi (ECMF) and the effect of ECMF species on tree seedling condition in CWD will elucidate the potential for ECMF-mediated effects on seedling dynamics. In hemlock-dominated stands, we characterized ECMF communities associated with...

  12. Comparison of fecundity and survival of hemlock woolly adelgid (Hemiptera: Adelgidae) in northern and southern populations

    Artemis Roehrig; Joseph. Elkinton


    The hemlock woolly adelgid (HWA), Adelges tsugae Annand (Hemiptera: Adelgidae), is an introduced species first reported in the eastern United States in 1951. The infestation has since spread in all directions from its initial sighting in Virginia, to its current range from northern Georgia, to southern Maine, and westward into Tennessee, causing...

  13. Induction of cold hardiness in an invasive herbivore: The case of hemlock woolly adelgid (Hemiptera: Adelgidae)

    Joseph S. Elkinton; Jeffrey A. Lombardo; Artemis D. Roehrig; Thomas J. McAvoy; Albert Mayfield; Mark Whitmore


    As a measure of cold hardiness, we tested the supercooling points or freezing temperatures of individual hemlock woolly adelgids (Adelges tsugae Annand) collected from 15 locations across the north to south range of the adelgid in eastern North America at different times during two winters. Adelgids from the northern interior locations with USDA hardiness zones of 5B–...

  14. Enhancement of foreign collection and quarantine evaluation of hemlock woolly adelgid natural enemies

    Mike Montgomery; Roy Van Driesche; Scott Salom; Wenhua Lu; Guoyue Yu; Jianhua Zhou; Li Li; Shigehiko Shiyake


    The objectives of this newly funded Forest Service activity are to promote the biological control of hemlock woolly adelgid (Adelges tsugae Annand) by (1) hiring an entomologist to work in Asia; (2) revisit known collecting sites and find promising new ones; (3) study the life history and phenology of HWA and its predators; (4) establish protocols...

  15. Hemlock woolly adelgid and its natural enemies in Sichuan Province, China, 2005

    Jianhua Zhou; Yinbo Xiao; Yugui Xiao; Wenhua Lu; Michael Montgomery; Roy Van Driesche; Scott Salom


    A partnership of Chinese and American institutions was formed in 2005 to obtain natural enemies for biological control of Adelges tsugae Annand, the hemlock woolly adelgid (HWA), in the eastern United States. We report here the first 6 months (June-November) of studies done at three sites in Kangding and Baoxing Counties in Sichuan Province.

  16. Developing Biomass Equations for Western Hemlock and Red Alder Trees in Western Oregon Forests

    Krishna Poudel; Hailemariam Temesgen


    Biomass estimates are required for reporting carbon, assessing feedstock availability, and assessing forest fire threat. We developed diameter- and height-based biomass equations for Western hemlock (Tsuga heterophylla (Raf.) Sarg.) and red alder (Alnus rubra Bong.) trees in Western Oregon. A system of component biomass...

  17. Characterization of terpenoid volatiles from cultivars of eastern hemlock (Tsuga canadensis).

    Lagalante, Anthony F; Montgomery, Michael E; Calvosa, Frank C; Mirzabeigi, Michael N


    The volatile terpenoid fraction from needles in 13 cultivars of Tsuga canadensis L. (Carriere) was analyzed by gas chromatography with mass spectrometry (GC-MS). The results of this study are considered along with previously reported results for foliar terpenoid levels of the Asian (T. sieboldii, T. chinensis, T. diversifolia), western North American (T. mertensiana, T. heterophylla), and eastern North American species (T. canadensis, T. caroliniana) of hemlock to draw conclusions about the potential of cultivar host resistance to the hemlock woolly adelgid (Adelges tsugae Annand). It is suggested that hemlocks in eastern North America have adapted their terpenoid chemistry for protection against endemic defoliators and that this has made them vulnerable to non-native, sucking pests such as adelgids and scales. Some cultivars of T. canadensis have a terpenoid profile that resembles that of the resistant noneastern North American species and are candidates for biological screening for resistance. Among the cultivars, the variation in terpenoid chemistry did not absolutely correspond with the considerable differences in morphological characters observed, indicating that the terpenoid chemistry is not definitively coupled with hemlock morphology.

  18. Development of Sirococcus shoot blight following thinning in western hemlock regeneration.

    Charles G. Shaw; Thomas H. Laurent; Spencer. Israelson


    Shoot mortality from Sirococcus strobilinus Preuss. and other causes was recorded by crown position from April 1978 through October 1979 in younggrowth western hemlock (Tsuga heterophylla (Raf.) Sarg.) crop trees released in a 1977 thinning at Thomas Bay, Alaska. All study trees contained some infected shoots, but no terminal...

  19. Ecosystem Function in Appalachian Headwater Streams during an Active Invasion by the Hemlock Woolly Adelgid

    Robert M. Northington; Jackson R. Webster; Ernest F. Benfield; Beth M. Cheever; Barbara R. Niederlehner


    Forested ecosystems in the southeastern United States are currently undergoing an invasion by the hemlock woolly adelgid (HWA). Previous studies in this area have shown changes to forest structure, decreases in canopy cover, increases in organic matter, and changes to nutrient cycling on the forest floor and soil. Here, we were interested in how the effects of canopy...

  20. Prey suitability and phenology of Leucopis spp. (Diptera: Chamaemyiidae) associated with hemlock woolly adelgid (Hemiptera: Adelgidae) in the Pacific Northwest

    Sarah M. Grubin; Darrell W. Ross; Kimberly F. Wallin


    Leucopis spp. (Diptera: Chamaemyiidae) from the Pacific Northwest previously were identified as potential biological control agents for the hemlock woolly adelgid, Adelges tsugae Annand (Hemiptera: Adelgidae), in the eastern United States. We collected Leucopis spp. larvae from A. tsugae...

  1. Aerial application of the insect-killing fungus Lecanicillium muscarium in a microfactory formulation for hemlock woolly adelgid suppression

    Scott Costa; Karen Felton; Bradley Onken; Richard Reardon; Rusty. Rhea


    Forest populations of hemlock woolly adelgid (HWA) were reduced using an operational formulation of the insect-killing fungus Lecanicillium muscarium when it was supported by microfactory formulation technology.

  2. A 4-year record of sitka spruce and western hemlock seed fall on the Cascade Head Experimental Forest.

    Robert H. Ruth; Carl M. Berntsen


    Four years' measurement of seed fall in the spruce-hemlock type on the Cascade Head Experimental Forest indicates that an ample supply of seed is distributed over clear-cut areas under staggered-setting cutting. The largest tract sampled was 81 acres; in spite of a seed crop failure in 1950, it received an average of 243,000 viable spruce and hemlock seeds per...

  3. Variation in plant defense against invasive herbivores: evidence for a hypersensitive response in eastern hemlocks (Tsuga canadensis).

    Radville, Laura; Chaves, Arielle; Preisser, Evan L


    Herbivores can trigger a wide array of morphological and chemical changes in their host plants. Feeding by some insects induces a defensive hypersensitive response, a defense mechanism consisting of elevated H(2)O(2) levels and tissue death at the site of herbivore feeding. The invasive hemlock woolly adelgid Adelges tsugae ('HWA') and elongate hemlock scale Fiorinia externa ('EHS') feed on eastern hemlocks; although both are sessile sap feeders, HWA causes more damage than EHS. The rapid rate of tree death following HWA infestation has led to the suggestion that feeding induces a hypersensitive response in hemlock trees. We assessed the potential for an herbivore-induced hypersensitive response in eastern hemlocks by measuring H(2)O(2) levels in foliage from HWA-infested, EHS-infested, and uninfested trees. Needles with settled HWA or EHS had higher H(2)O(2) levels than control needles, suggesting a localized hypersensitive plant response. Needles with no direct contact to settled HWA also had high H(2)O(2) levels, suggesting that HWA infestation may induce a systemic defense response in eastern hemlocks. There was no similar systemic defensive response in the EHS treatment. Our results showed that two herbivores in the same feeding guild had dramatically different outcomes on the health of their shared host.

  4. Congenital skeletal malformations induced by maternal ingestion of Conium maculatum (poison hemlock) in newborn pigs.

    Panter, K E; Keeler, R F; Buck, W B


    Skeletal malformations were induced in newborn pigs from gilts fed Conium maculatum seed or plant during gestation days 43 through 53 and 51 through 61. The teratogenic effects in groups dosed during gestation days 43 through 53 were more severe than those in groups dosed during the later period, with many newborn pigs showing arthrogryposis and twisted and malaligned bones in the limbs and with 1 pig showing scoliosis and deformity of the thoracic cage. The pigs born to gilts given C maculatum during gestation days 51 through 61 had excessive flexure primarily in the carpal joints, without scoliosis or bone malalignment in the limbs. The teratogenicity of poison hemlock depends on the alkaloid concentration and content. Based on the data presented, we speculate that gamma-coniceine is the teratogenic alkaloid in the poison hemlock fed to the gilts.

  5. Lupines, poison-hemlock and Nicotiana spp: toxicity and teratogenicity in livestock.

    Panter, K E; James, L F; Gardner, D R


    Many species of lupines contain quinolizidine or piperidine alkaloids known to be toxic or teratogenic to livestock. Poison-hemlock (Conium maculatum) and Nicotiana spp. including N. tabacum and N. glauca contain toxic and teratogenic piperidine alkaloids. The toxic and teratogenic effects from these plant species have distinct similarities including maternal muscular weakness and ataxia and fetal contracture-type skeletal defects and cleft palate. It is believed that the mechanism of action of the piperidine and quinolizidine alkaloid-induced teratogenesis is the same; however, there are some differences in incidence, susceptible gestational periods, and severity between livestock species. Wildlife species have also been poisoned after eating poison-hemlock but no terata have been reported. The most widespread problem for livestock producers in recent times has been lupine-induced "crooked calf disease." Crooked calf disease is characterized as skeletal contracture-type malformations and occasional cleft palate in calves after maternal ingestion of lupines containing the quinolizidine alkaloid anagyrine during gestation days 40-100. Similar malformations have been induced in cattle and goats with lupines containing the piperidine alkaloids ammodendrine, N-methyl ammodendrine, and N-acetyl hystrine and in cattle, sheep, goats, and pigs with poison-hemlock containing predominantly coniine or gamma-coniceine and N. glauca containing anabasine. Toxic and teratogenic effects have been linked to structural aspects of these alkaloids, and the mechanism of action is believed to be associated with an alkaloid-induced inhibition of fetal movement during specific gestational periods. This review presents a historical perspective, description and distribution of lupines, poison-hemlock and Nicotiana spp., toxic and teratogenic effects and management information to reduce losses.

  6. Experimental Evidence that Hemlock Mortality Enhances Carbon Stabilization in Southern Appalachian Forest Soils

    Fraterrigo, J.; Ream, K.; Knoepp, J.


    Forest insects and pathogens (FIPs) can cause uncertain changes in forest carbon balance, potentially influencing global atmospheric carbon dioxide (CO2) concentrations. We quantified the effects of hemlock (Tsuga canadensis L. Carr.) mortality on soil carbon fluxes and pools for a decade following either girdling or natural infestation by hemlock woolly adelgid (HWA; Adelges tsugae) to improve mechanistic understanding of soil carbon cycling response to FIPs. Although soil respiration (Rsoil) was similar among reference plots and plots with hemlock mortality, both girdled and HWA-infested plots had greater activities of β-glucosidase, a cellulose-hydrolyzing extracellular enzyme, and decreased O-horizon mass and fine root biomass from 2005 to 2013. During this period, total mineral soil carbon accumulated at a higher rate in disturbed plots than in reference plots in both the surface (0-10 cm) and subsurface (10-30 cm); increases were predominantly in the mineral-associated fraction of the soil organic matter. In contrast, particulate organic matter carbon accrued slowly in surface soils and declined in the subsurface of girdled plots. δ13C values of this fraction demonstrate that particulate organic matter carbon in the surface soil has become more microbially processed over time, suggesting enhanced decomposition of organic matter in this pool. Together, these findings indicate that hemlock mortality and subsequent forest regrowth has led to enhanced soil carbon stabilization in southern Appalachian forests through the translocation of carbon from detritus and particulate soil organic matter pools to the mineral-associated organic matter pool. These findings have implications for ecosystem management and modeling, demonstrating that forests may tolerate moderate disturbance without diminishing soil carbon storage when there is a compensatory growth response by non-host trees.

  7. Effects of rhododendron removal on the water use of hardwood species following eastern hemlock mortality

    Hawthorne, S. N.; Miniat, C.; Elliott, K.


    Forest disturbance that alters vegetation species composition can affect ecosystem productivity and function. The loss of eastern hemlock (Tsuga canadensis) to hemlock woolly adelgid infestations in southern Appalachian Mountains has resulted in more than a two-fold increase in growth of co-occurring rhododendron (Rhododendron maximum) understory, evergreen shrubs. In contrast, the growth of hardwood species increased by 1.2 fold during the same 5 year period following infestation. This study examines the effects of mechanically removing the rhododendron shrub layer on water use and growth of hardwood species. The treatment—hypothesized to speed ecosystem recovery of structure and function—involved cutting, spreading and burning rhododendron stems to remove both rhododendron and soil O-horizon. Sap flow, soil moisture and micro-climate (humidity, temperature) were measured in a pair of reference and treated plots. Preliminary results from the relatively dry summer/fall 2016 have shown that the mean daily transpiration (Et) of the treated plot was 24% greater than the mean daily Et of hardwood trees in the reference plot (t-test, p treatment plots compared to the reference plots. This suggests that the removal of the shrub layer reduced competition for resources for the canopy and seedling trees, which may increase tree growth and recruitment. Thus, in the wake of hemlock loss, recovery of riparian forest structure and function may be aided with shrub layer removal.

  8. Ancient and modern colonization of North America by hemlock woolly adelgid, Adelges tsugae (Hemiptera: Adelgidae), an invasive insect from East Asia

    Nathan P. Havill; Shigehiko Shiyake; Ashley Lamb Galloway; Robert G. Foottit; Guoyue Yu; Annie Paradis; Joseph Elkinton; Michael E. Montgomery; Masakazu Sano; Adalgisa Caccone


    Hemlock woolly adelgid, Adelges tsugae, is an invasive pest of hemlock trees (Tsuga) in eastern North America. We used 14 microsatellites and mitochondrial COI sequences to assess its worldwide genetic structure and reconstruct its colonization history. The resulting information about its life cycle, biogeography and host...

  9. Eastern hemlock response to even- and uneven-age management in the Acadian forest: results from the Penobscot Experimental Forest long-term silviculture study

    John C. Brissette; Laura S. Kenefic


    Eastern hemlock (Tsuga canadensis (L.) Carr.) is an important tree species in the mixed-species conifer forests of northern New England and adjacent Canada. Hemlock is very tolerant of understory conditions; consequently, it responds differently to various silvicultural treatments. In a long-term study at the Penobscot Experimental Forest in east-...

  10. Radio ultrasound observations of the fetotoxic effects in sheep from ingestion of Conium maculatum (poison-hemlock).

    Panter, K E; Bunch, T D; Keeler, R F; Sisson, D V


    Fetal movement in pregnant ewes gavaged with Conium maculatum (poison-hemlock) was reduced significantly, but temporarily. Fetal movement was observed by radio ultrasound at 45, 54 and 60 days of gestation in control ewes and on days 45, 54, and 60 of gestation immediately before and 1 hour following poison-hemlock feeding in treated ewes. Fetal movement was significantly reduced (P less than 0.01) 1 hour after poison-hemlock administration, but returned to normal within 18 hours post treatment. At parturition seven of eleven lambs born to seven treated ewes had varying degrees of front limb abnormalities. Modest to moderate flexure of the carpal joints, some lateral deviation in the front limbs at the pastern joint and kinked tails were observed. These malformations were transient and resolved spontaneously by 8 weeks after lambing.

  11. Responses of sugar maple and hemlock seedlings to elevated carbon dioxide under altered above and below ground nitrogen sources

    Eller, Allyson S.D.; McGuire, Krista L.; Sparks, Jed


    The aim of this study is to determine the influence of CO2, NO2 and nitrate deposition (NO3-) increase at the same time on tree seedlings. Experiments were conducted on sugar maple and eastern hemlock in an open field over a two year period. They were grown under ambient or elevated CO2 and NO2 and with or without wet deposition of NO3-. Results showed that the effects of one treatment can be eliminated by another treatment thus demonstrating these effects are not additive. The growth of both sugar maple and eastern hemlock was found to be similar under the influence of the three treatments and under control conditions. .

  12. Hemlock Alkaloids in Aloes. Occurrence and Distribution of gamma-Coniceine.

    Dring, J V; Nash, R J; Roberts, M F; Reynolds, T


    The hemlock alkaloid gamma-coniceine was identified in a number of ALOE species, namely A. GILLILANDII, Reynolds A. BALLYI Reynolds, A. RUSPOLIANA Baker, A. IBITIENSIS Perrier and A. DELTOIDEODONTA Baker. Coniine was identified in A. VIGUIERI Perrier. The levels of gamma-coniceine are higher than those found in CONIUM MACULATUM L. Some species also contained trace amounts of conhydrinone and pseudoconhydrin. Three of the species are Madagascan endemics, one is restricted to Arabia, while the rest are remote from each other in East Africa. Some of the species are loosely related but there is no overall taxonomic affinity between them.

  13. Maternal and fetal toxicity of poison hemlock (Conium maculatum) in sheep.

    Panter, K E; Bunch, T D; Keeler, R F


    Poison hemlock (Conium maculatum) was toxic to pregnant ewes and their fetuses when fed during gestation days 30 through 60. Maternal effects included trembling, muscular weakness in the neck initially, then progressing to the limbs, ataxia, frequent urination and defecation, and death. Convulsive seizures were not observed. Fetotoxic effects included excessive flexure of the carpal joints with lateral deviation in the front limbs and kinked tails. At term, 7 of 11 lambs had varying degrees of the limb abnormalities, but all lambs appeared clinically normal at 8 weeks after parturition.

  14. Fire-mediated pathways of stand development in Douglas-fir/western hemlock forests of the Pacific Northwest, USA

    A.J. Tepley; F.J. Swanson; T.A. Spies


    Forests dominated by Douglas-fir and western hemlock in the Pacific Northwest of the United States have strongly influenced concepts and policy concerning old-growth forest conservation. Despite the attention to their old-growth characteristics, a tendency remains to view their disturbance ecology in relatively simple terms, emphasizing infrequent, stand-replacing (SR...

  15. Contamination delays the release of Laricobius osakensis for biological control of hemlock woolly adelgid: cryptic diversity in Japanese Laricobius spp

    Melissa J. Fischer; Nathan P. Havill; Carrie S. Jubb; Sean W. Prosser; Brent D. Opell; Scott M. Salom; Loke T. Kok


    Laricobius osakensis (Coleoptera: Derodontidae) was imported from Japan to the United States in 2006 for study in quarantine facilities as a potential biological control of Hemlock Woolly Adelgid. Laricobius osakensis was released from quarantine in 2010, but it was soon discovered that the colony also contained a cryptic species...

  16. Measurement of imidacloprid in xylem fluid from eastern hemlock (Tsuga canadensis) by derivitization/GC/MS and ELISA

    Anthony Lagalante; Peter Greenbacker; Jonathan Jones; Richard Turcotte; Bradley Onken


    Imidacloprid is a nonvolatile insecticide and its direct quantification is not possible by gas chromatography. In order to ascertain imidacloprid levels in soil and trunk injection treated trees, a sensitive and selective method has been developed using GC/MS to measure the imidacloprid levels in xylem fluid exudates. In May 2005, a stand of hemlock trees in West...

  17. Host Range Specificity of Scymnus camptodromus (Coleoptera: Coccinellidae), A Predator of Hemlock Woolly Adelgid (Hemiptera: Adelgidae).

    Limbu, Samita; Cassidy, Katie; Keena, Melody; Tobin, Patrick; Hoover, Kelli


    Scymnus (Neopullus) camptodromus Yu and Liu (Coleoptera: Coccinellidae) was brought to the United States from China as a potential biological control agent for hemlock woolly adelgid (Adelges tsugae Annand) (Hemiptera: Adelgidae). Scymnus camptodromus phenology is closely synchronized with that of A. tsugae and has several characteristics of a promising biological control agent. As a prerequisite to field release, S. camptodromus was evaluated for potential nontarget impacts. In host range studies, the predator was given the choice of sympatric adelgid and nonadelgid prey items. Nontarget testing showed that S. camptodromus will feed to some degree on other adelgid species, but highly prefers A. tsugae. We also evaluated larval development of S. camptodromus on pine bark adelgid (Pineus strobi (Hartig)) (Hemiptera: Adelgidae) and larch adelgid (Adelges laricis Vallot) (Hemiptera: Adelgidae); a small proportion of predator larvae was able to develop to adulthood on P. strobi or A. laricis alone. Scymnus camptodromus showed no interest in feeding on woolly alder aphid (Paraprociphilus tessellatus Fitch) (Hemiptera: Aphididae) or woolly apple aphid (Eriosoma lanigerum (Hausmann)) (Hemiptera: Aphididae), and minimal interest in cotton aphid (Aphis gossypii Glover) (Hemiptera: Aphididae) in choice and no-choice experiments. Scymnus camptodromus females did not oviposit on any host material other than A. tsugae-infested hemlock. Under the circumstances of the study, S. camptodromus appears to be a specific predator of A. tsugae, with minimal risk to nontarget species. Although the predator can develop on P. strobi, the likelihood that S. camptodromus would oviposit on pine hosts of this adelgid is small.

  18. Comparative toxicity of coniine, an alkaloid of Conium maculatum (poison hemlock), in chickens, quails, and turkeys.

    Frank, A A; Reed, W M


    Coniine, an alkaloid of Conium maculatum (poison hemlock), was administered by gavage to immature chickens, quails, and turkeys at 0, 25, 50, and 100 mg/kg body weight. At 25 mg coniine/kg body weight, clinical signs were observed only in quails (2/10) and consisted of excitement, depression, hypermetria, seizures, opisthotonos, and flaccid paralysis. Chickens (9/10) and quails (8/10) dosed at 50 mg/kg body weight were affected, and several birds of each species died (2/10 and 5/10, respectively). Turkeys (7/10) were affected only when dosed at 100 mg/kg body weight, and quails (6/10), turkeys (4/10), and chickens (10/10) died at this dose. There were no gross or microscopic lesions. Coniine was detected in skeletal muscle and liver of birds dying after ingestion and was present in some survivors 7 days post-treatment.

  19. Anti-Pseudomonas aeruginosa activity of hemlock (Conium maculatum, Apiaceae) essential oil.

    Di Napoli, Michela; Varcamonti, Mario; Basile, Adriana; Bruno, Maurizio; Maggi, Filippo; Zanfardino, Anna


    Conium maculatum is a nitrophilous weed belonging to the Apiaceae family and occurring in hedgerows, pastures, waste ground, along rivers and roadsides. Little is known on the chemistry and bioactivity of other secondary metabolites occurring in the plant. In the present work, we have analysed the chemical composition and antimicrobial activity of the essential oils hydrodistilled from leaves and inflorescenes of C. maculatum growing in Sicily, Italy. The composition of essential oils was achieved by gas chromatography-mass spectrometry (GC-MS) analysis, whereas the inhibitory effects on the growth of two Gram negative strains, namely Escherichia coli and Pseudomonas aeruginosa were assessed by two different analysis. The essential oils exhibited different chemical profiles (1-butylpiperidine and myrcene in the inflorescenes), (mostly (E)-caryophyllene in the leaves). The latter oil was particularly active in inhibiting the growth of P. aeruginosa. These results shed light on the possible application of hemlock essential oils as antimicrobial agents.

  20. Windthrow and salvage logging in an old-growth hemlock-northern hardwoods forest

    Lang, K.D.; Schulte, L.A.; Guntenspergen, G.R.


    Although the initial response to salvage (also known as, post-disturbance or sanitary) logging is known to vary among system components, little is known about longer term forest recovery. We examine forest overstory, understory, soil, and microtopographic response 25 years after a 1977 severe wind disturbance on the Flambeau River State Forest in Wisconsin, USA, a portion of which was salvage logged. Within this former old-growth hemlock-northern hardwoods forest, tree dominance has shifted from Eastern hemlock (Tsuga canadensis) to broad-leaf deciduous species (Ulmus americana, Acer saccharum, Tilia americana, Populus tremuloides, and Betula alleghaniensis) in both the salvaged and unsalvaged areas. While the biological legacies of pre-disturbance seedlings, saplings, and mature trees were initially more abundant in the unsalvaged area, regeneration through root suckers and stump sprouts was common in both areas. After 25 years, tree basal area, sapling density, shrub layer density, and seedling cover had converged between unsalvaged and salvaged areas. In contrast, understory herb communities differed between salvaged and unsalvaged forest, with salvaged forest containing significantly higher understory herb richness and cover, and greater dominance of species benefiting from disturbance, especially Solidago species. Soil bulk density, pH, organic carbon content, and organic nitrogen content were also significantly higher in the salvaged area. The structural legacy of tip-up microtopography remains more pronounced in the unsalvaged area, with significantly taller tip-up mounds and deeper pits. Mosses and some forest herbs, including Athyrium filix-femina and Hydrophyllum virginianum, showed strong positive responses to this tip-up microrelief, highlighting the importance of these structural legacies for understory biodiversity. In sum, although the pathways of recovery differed, this forest appeared to be as resilient to the compound disturbances of windthrow

  1. A 10-Year Assessment of Hemlock Decline in the Catskill Mountain Region of New York State Using Hyperspectral Remote Sensing Techniques.

    Hanavan, Ryan P; Pontius, Jennifer; Hallett, Richard


    The hemlock woolly adelgid is a serious pest of Eastern and Carolina hemlock in the eastern United States. Successfully managing the hemlock resource in the region depends on careful monitoring of the spread of this invasive pest and the targeted application of management options such as biological control, chemical, or silvicultural treatments. To inform these management activities and test the applicability of a landscape-scale remote sensing effort to monitor hemlock condition, hyperspectral collections, and concurrent ground-truthing in 2001 and 2012 of hemlock condition were compared with field metrics spanning a 10-yr survey in the Catskills region of New York. Fine twig dieback significantly increased from 9 to 15% and live crown ratio significantly decreased from 67 to 56% in 2001 and 2012, respectively. We found a significant shift from 59% "healthy" hemlock in 2001 to only 16% in 2012. However, this shift from healthy to declining classifications was mostly a shift to decline class 2 "early decline". These results indicate that while there has been significant increase in decline symptoms as measured in both field and remote sensing assessments, a majority of the declining areas identified in the resulting spatial coverages remain in the "early decline" category and widespread mortality has not yet occurred. While this slow decline across the region stands in contrast to many reports of mortality within 10 yr, the results from this work are in line with other long-term monitoring studies and indicate that armed with the spatial information provided here, continued management strategies can be focused on particular areas to help control the further decline of hemlock in the region. Published by Oxford University Press on behalf of Entomological Society of America 2015. This work is written by US Government employees and is in the public domain in the US.

  2. Effect of the habitat on the mass and germinative capacity of poison hemlock (Conium maculatum L. achenes

    Jan Winkler


    Full Text Available Poison hemlock (Conium maculatum L. is a overwintering annual or biennial herb, which propagates only generatively. We estimated the germinative capacity and weight of its achenes harvested on different habitats. The territory where achenes were picked up is situated in the municipality Holice near Olomouc, Czech Republic (altitude of 220–235 m. Material was collected in the years 2002 and 2003 on two different habitats. The first one was on a field after sugar beet and maize as preceding crops, the second one was an adjacent balk. The mass of seeds was estimated in ten replications (100 achenes each. Thereafter the achenes were weighed on an analytical scales. The germinative capacity was assessed under different conditions of germination. Altogether there were 8 variants of germination conditions in 10 replications (20 achenes each. The first and the last evaluation were performed on the 9th and 30th day of the experiment, respectively. The total average mass of 100 achenes originating from all habitats and both experimental years was 0.23 g. The total average germinative capacity of poison hemlock achenes was 53%. The mass of achenes from the balk site was significantly higher than those from the field but the difference in their germinative capacity was statistically insignificant. It was concluded that the hemlock can be considered as a negatively photoblastic plant. Its achenes which become ripe in the proximity of fields can show an important effect on their weed infestation.

  3. Steroids from Poison Hemlock (Conium maculatum L.: A GC-MS analysis

    Radulović Niko S.


    Full Text Available The steroid content of Conium maculatum L. (Poison Hemlock, Apiaceae, a well-known weed plant species, was studied herein for the first time. This was achieved by detailed GC-MS analyses of twenty two samples (dichloromethane extracts of different plant organs of C. maculatum at three or four different stages of phenological development, collected from three locations. In total, twenty four different steroids were identified. Six steroids had an ergostane nucleus while the other ones possessed a stigmastane carbon framework. The identity of these compounds was determined by spectral means (MS fragmentation, GC co-injections with authentic standards and chemical transformation (silylation. Steroid compounds were noted to be the main chemical constituents of root extracts (up to 70 % of this plant species in the last phase of development. The predominant ones were stigmasta-5,22- dien-3β-ol (stigmasterol and stigmasta-5-en-3β-ol (β-sitosterol. In an attempt to classify the samples, principal component analysis (PCA and agglomerative hierarchical clustering (AHC were performed using steroid percentages as variables.

  4. Residual densities affect growth of overstory trees and planted Douglas-fir, western hemlock, and western redcedar: results from the first decade

    Leslie Chandler Brodie; Dean S. DeBell


    In recent years, interest has increased in silvicultural systems and harvest cuts that retain partial overstories, but there are few data available on the growth of the understory trees in such stands. We studied the response of overstory trees and underplanted seedlings, Douglas-fir (Pseudotsuga menziesii), western hemlock (Tsuga...

  5. Assessment of the potential for hybridisation between Laricobius nigrinus (Coleoptera: Derodontidae) and Laricobius osakensis, predators of the hemlock woolly adelgid (Hemiptera: Adelgidae)

    Melissa J. Fischer; Carlyle C. Brewster; Nathan P. Havill; Scott M. Salom; Loke T. Kok


    In 2003, Laricobius nigrinus Fender was introduced into the eastern United States as a biological control agent of the hemlock woolly adelgid (Adelges tsugae Annand). Following its release, it was discovered that L. nigrinus was hybridising and producing viable progeny with Laricobius rubidus...

  6. Identifying core habitat and connectivity for focal species in the interior cedar-hemlock forest of North America to complete a conservation area design

    Lance Craighead; Baden Cross


    To identify the remaining areas of the Interior Cedar- Hemlock Forest of North America and prioritize them for conservation planning, the Craighead Environmental Research Institute has developed a 2-scale method for mapping critical habitat utilizing 1) a broad-scale model to identify important regional locations as the basis for a Conservation Area Design (CAD), and 2...

  7. Growth and yield of all-aged Douglas-fir -- western hemlock forest stands: a matrix model with stand diversity effects.

    Jingjing Liang; Joseph Buonglorno; Robert A. Monserud


    A density-dependent matrix model was developed for Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) -- western hemlock (Tsuga heterophylla (Raf.) Sarg.) forest stands in the Pacific Northwest of the United States. The model predicted the number and volume of trees for 4 species groups and 19 diameter classes. The parameters...

  8. Behavioral responses of Laricobius spp.and hybrids (Coleoptera: Derodontidae) to hemlock woolly adelgid and adelgid host tree odors in an olfactometer

    Arielle L. Arsenault; Nathan P. Havill; Albert E. Mayfield; Kimberly F. Wallin


    The predatory species Laricobius nigrinus (Fender) and Laricobius osakensis (Shiyake and Montgomery) (Coleoptera: Derodontidae) have been released for biological control of hemlock woolly adelgid (Adelges tsugae; Hemiptera: Adelgidae) in eastern North America. L. osakensis is native to Japan, whereas L. nigrinus is endemic to the Pacific Northwest of the United States...

  9. Macro-scale assessment of demographic and environmental variation within genetically derived evolutionary lineages of eastern hemlock (Tsuga canadensis), an imperiled conifer of the eastern United States

    Anantha M. Prasad; Kevin M. Potter


    Eastern hemlock (Tsuga canadensis) occupies a large swath of eastern North America and has historically undergone range expansion and contraction resulting in several genetically separate lineages. This conifer is currently experiencing mortality across most of its range following infestation of a non-native insect. With the goal of better...

  10. Toxicosis in dairy cattle exposed to poison hemlock (Conium maculatum) in hay: isolation of Conium alkaloids in plants, hay, and urine.

    Galey, F D; Holstege, D M; Fisher, E G


    Cattle in two herds developed signs of bloating, increased salivation and lacrimation, depression, respiratory distress, ataxia, and death after ingestion of hay that contained large amounts of poison hemlock (Conium maculatum). Twenty of 30 Angus cows and calves were affected in the first herd (2 died). In the second herd, 5 of 30 Holstein heifers were affected (1 died). The Conium alkaloids, coniine and gamma-coniceine, were quantified in the hay, the plants from the responsible hayfield, and the urine of affected animals.

  11. [Child poisoning after ingestion of a wild apiaceae: a case report].

    Durand, M-F; Pommier, P; Chazalette, A; de Haro, L


    Apiaceae family (formerly Umbelliferae) contains several highly toxic species, including Poison Hemlock (Conium maculatum), Water Hemlock (Cicuta virosa) and Hemlock Water Dropwort (Oenanthe crocata) which are the three main poisonous Apiaceae species growing in France. Thinking he was identifying wild carrots, an 11-year-old boy without previous history ingested the root from a wild Apiaceae. One hour later, he was confused, had drowsiness, headache as well as abdominal pain, vomiting and diarrhoea. Upon hospital admission, myosis, ophtalmoplegia and a moderate rhabdomyolysis were noted. The patient recovered after 24 h of symptomatic treatments. In this case, the description of the ingested plant allowed to identify the Apiaceae family but not the species involved. The geographical location (Southern France in a humid area), the clinical features and the aspect of the ingested root, with an orange secretion led to implicate Oenanthe crocata as the origin of this unusual poisoning.

  12. Cryptic east-west divergence and molecular diagnostics for two species of silver flies (Diptera: Chamaemyiidae: Leucopis ) from North America being evaluated for biological control of hemlock woolly adelgid

    Nathan P. Havill; Stephen D. Gaimari; Adalgisa. Caccone


    Exploring genetic diversity within species of biological control agents can expose previously overlooked beneficial genotypes. This may be the case for two species of silver flies, Leucopis argenticollis and L. piniperda, predators of the hemlock woolly adelgid (Adelges tsugae) in the Pacific Northwest of...

  13. Predators collected from balsam woolly adelgid and Cooley spruce gall adelgid in western Oregon and Washington, U.S.A., with reference to biological control of hemlock woolly adelgid (Hemiptera: Adelgidae)

    Darrell W. Ross; Glenn R. Kohler; Kimberly F. Wallin


    As part of a comprehensive study to survey predators associated with hemlock woolly adelgid, Adelges tsugae Annand, 1928 in the Pacific Northwest (PNW), U.S.A. (Kohler et al. 2008), predators of balsam woolly adelgid, Adelges piceae (Ratzeburg, 1844) and Cooley spruce gall adelgid, Adelges cooleyi (Gillette...

  14. Plant guide: Douglas' dusty-maiden (Chaenactic douglasii)

    Derek Tilley; Dan Ogle; Loren St. John


    Douglas' dustymaiden can be used as part of a native forb component in wildland seedings to increase biodiversity, improve wildlife habitat, and provide food for numerous birds and mammals. Douglas' dustymaiden is readily visited by pollinators and other insect species. It is considered an important species for sage grouse during brood rearing because of its...

  15. Multi-decade biomass dynamics in an old-growth hemlock-northern hardwood forest, Michigan, USA

    Kerry D. Woods


    Full Text Available Trends in living aboveground biomass and inputs to the pool of coarse woody debris (CWD in an undisturbed, old-growth hemlock-northern hardwood forest in northern MI were estimated from multi-decade observations of permanent plots. Growth and demographic data from seven plot censuses over 47 years (1962–2009, combined with one-time measurement of CWD pools, help assess biomass/carbon status of this landscape. Are trends consistent with traditional notions of late-successional forests as equilibrial ecosystems? Specifically, do biomass pools and CWD inputs show consistent long-term trends and relationships, and can living and dead biomass pools and trends be related to forest composition and history? Aboveground living biomass densities, estimated using standard allometric relationships, range from 360–450 Mg/ha among sampled stands and types; these values are among the highest recorded for northeastern North American forests. Biomass densities showed significant decade-scale variation, but no consistent trends over the full study period (one stand, originating following an 1830 fire, showed an aggrading trend during the first 25 years of the study. Even though total above-ground biomass pools are neither increasing nor decreasing, they have been increasingly dominated, over the full study period, by very large (>70 cm dbh stems and by the most shade-tolerant species (Acer saccharum and Tsuga canadensis.CWD pools measured in 2007 averaged 151 m3/ha, with highest values in Acer-dominated stands. Snag densities averaged 27/ha, but varied nearly ten-fold with canopy composition (highest in Tsuga-dominated stands, lowest in Acer-dominated; snags constituted 10–50% of CWD biomass. Annualized CWD inputs from tree mortality over the full study period averaged 1.9–3.2 Mg/ha/yr, depending on stand and species composition. CWD input rates tended to increase over the course of the study. Input rates may be expected to increase over longer

  16. Growth and Its Relationship to Individual Genetic Diversity of Mountain Hemlock (Tsuga mertensiana at Alpine Treeline in Alaska: Combining Dendrochronology and Genomics

    Jeremy S. Johnson


    Full Text Available Globally, alpine treelines are characterized as temperature-limited environments with strong controls on tree growth. However, at local scales spatially heterogeneous environments generally have more variable impacts on individual patterns of tree growth. In addition to the landscape spatial heterogeneity there is local variability in individual tree genetic diversity (level of individual heterozygosity. It has been hypothesized that higher individual heterozygosity will result in more consistent patterns of growth. In this article, we combine genomics and dendrochronology to explore the relationship between individual genetic diversity and tree growth at a mountain hemlock (Tsuga mertensiana Bong. Carr alpine treeline on the Kenai Peninsula, Alaska, USA. We correlated average observed individual heterozygosity with average tree-ring width and variance in tree-ring width within individuals to test the hypothesis that trees with higher individual heterozygosity will also have more consistent growth patterns, suggesting that they may be more resilient to climate and environmental fluctuations at the alpine treeline. Our results showed that there was no significant relationship between tree growth and individual heterozygosity. However, there was a significant positive relationship between average tree-ring width and variance in tree-ring width implying that overall, fast growing trees in stressful environments, such as the alpine treeline, grow unstably regardless of the level of individual heterozygosity.

  17. Thematic Mapper Analysis of Blue Oak (Quercus douglasii) in Central California

    Paul A. Lefebvre Jr.; Frank W. Davis; Mark Borchert


    Digital Thematic Mapper (TM) satellite data from September 1986 and December 1985 were analyzed to determine seasonal reflectance properties of blue oak rangeland in the La Panza mountains of San Luis Obispo County. Linear regression analysis was conducted to examine relationships between TM reflectance and oak canopy cover, basal area, and site topographic variables....

  18. The future of bioethics: three dogmas and a cup of hemlock.

    Dawson, Angus


    In this paper I argue that bioethics is in crisis and that it will not have a future unless it begins to embrace a more Socratic approach to its leading assumptions. The absence of a critical and sceptical spirit has resulted in little more than a dominant ideology. I focus on three key issues. First, that too often bioethics collapses into medical ethics. Second, that medical ethics itself is beset by a lack of self-reflection that I characterize here as a commitment to three dogmas. Third, I offer a more positive perspective by suggesting how bioethics may benefit from looking towards public health ethics as a new source of inspiration and direction.

  19. Induction of cleft palate in newborn pigs by maternal ingestion of poison hemlock (Conium maculatum).

    Panter, K E; Keeler, R F; Buck, W B


    Cleft palates were induced in newborn pigs of gilts fed Conium maculatum seed or plant during gestation days 30 through 45. Twelve of 23 newborn pigs born to 3 gilts given Utah-grown C maculatum seed and 9 of 12 newborn pigs born to a single gilt given the fresh Utah spring-growth C maculatum plant had cleft palates. The cleft palates ranged from a unilateral cleft, involving only 1 side of the palate, to a full bilateral cleft. Brachygnathia was also observed in some of these newborn pigs with cleft palate. Other malformations were not observed. Chemical analysis of seed and plant samples indicated that gamma-coniceine was the responsible teratogenic alkaloid. A daily dose of plant or seed that provided greater than or equal to 1.07 mg of gamma-coniceine/kg of body weight fed to gilts during the 30th through the 45th day of pregnancy resulted in teratogenic effects.

  20. A novel toxic alkaloid from poison hemlock (Conium maculatum L., Apiaceae): identification, synthesis and antinociceptive activity.

    Radulović, Niko; Dorđević, Nevenka; Denić, Marija; Pinheiro, Mariana Martins Gomes; Fernandes, Patricia Dias; Boylan, Fabio


    2-Pentylpiperidine, named conmaculatin, a novel volatile alkaloid related to coniine was identified from the renowned toxic weed Conium maculatum L. (Apiaceae). The structure of conmaculatin was corroborated by synthesis (8 steps starting from cyclohexanol, overall yield 12%). Conmaculatin's strong peripheral and central antinociceptive activity in mice was observed in a narrow dose range (10-20mg/kg). It was found to be lethal in doses higher than 20mg/kg. Copyright © 2011 Elsevier Ltd. All rights reserved.

  1. Some results of chemical debarking on sitka spruce, western hemlock, and red alder.

    Carl M. Berntsen


    Chemical treatment of standing trees to facilitate bark removal has received a great deal of attention in Eastern United States and Canada during the past decade. The potential advantages of removing bark or extending easy peeling throughout we year have stimulated tests of many chemicals and methods of application.

  2. Tsuga canadensis (L.) Carr, mortality will impact hydrologic processes in southern Appalachian forest ecosystems

    Chelcy R. Ford; James M. Vose


    Eastern hemlock (Tsuga canadensis (L.) Carr.) is one of the principal riparian and cove canopy species in the southern Appalachian Mountains. Throughout its range, eastern hemlock is facing potential widespread mortality from the hemlock woolly adelgid (HWA). If HWA-induced eastern hemlock mortality alters hydrologic function, land managers...


    Francisco López-Muñoz


    Full Text Available Los textos literarios del Siglo de Oro constituyen una interesante fuente para el estudio de la sociedad española tardorrenacentista y novobarroca, incluyendo sus figuras más marginales. En el presente trabajo, hemos analizado el mundo mágico de brujas y hechiceras desde la perspectiva del uso extraterapéutico de agentes farmacológicos y psicotrópicos, a través de los principales autores áureos, centrándonos, básicamente, en Miguel de Cervantes y Lope de Vega. Se han estudiado los principales agentes empleados en la elaboración de las pócimas venenosas de bruja, destacando las plantas alucinógenas de la familia de las Solanaceae (beleño, mandrágora, belladona, estramonio, solano, eléboro, además de otras, como el acónito, la cicuta, la adelfa, la verbena o la adormidera. Otras sustancias de procedencia animal o mineral también se emplearon en la confección de estos preparados (sapos, arsénico. Finalmente, se han analizado las posibles fuentes documentales en materia científica que pudieron utilizar estos dos destacados literatos, como el Dioscórides comentado por Andrés Laguna en ambos casos, y la Historia Natural de Plinio, comentada por Francisco Hernández y Gerónimo de Huerta, y el opúsculo Il Sapere Util’e Delettevole de Constantino Castriota, en el caso particular de Lope de Vega. WITCHES’ POTIONS IN THE LITERATURE OF THE SPANISH GOLDEN AGE: THE OTHER SIDE OF THE THERAPEUTIC AND PSYCHOTROPIC AGENTS ABSTRACT The literary texts of the Golden Age are an interesting source for the study of Spanish late Renaissance and early Baroque society, including their most marginal figures. In the present work, we have analyzed the magical world of witches and sorceresses from the perspective of extra-therapeutic use of pharmacological and psychotropic substances, through the main golden authors, focusing basically on Miguel de Cervantes and Lope de Vega. The main agents used in the production of witches poisonous potions


    Although tree- and stand-level estimates of forest water use are increasingly common, relatively little is known about partitioning of soil water resources among co-occurring tree species. We studied seasonal courses of soil water utilization in a 450-year-old Pseudotsuga menzies...

  5. Seasonal changes in chemical composition and nutritive value of native forages in a spruce-hemlock forest, southeastern Alaska.

    Thomas A. Hanley; Jay D. McKendrick


    Twenty-two forages from Admiralty Island, southeastern Alaska, were monitored bimonthly for one year to assess seasonal changes in their chemical composition: neutral detergent fiber, acid detergent fiber, cellulose, lignin/cutin, invitro dry-matter digestibility, total nitrogen, phosphorus, potassium, calcium, magnesium, sodium, copper, manganese, iron, and zinc....

  6. Testing Methods for Challenging the National Wetland Plant List: Using Tsuga canadensis (L.) Carr. (Eastern Hemlock) as a Case Study


    frame were constructed based on GIS data (Table 1) and habitat descriptions of areas where T. canadensis was likely to occur, such as wooded swamps...6 2.1 Small study areas (100 km2) ......................................................................................... 6 2.1.1 Constructing a...Western Ecology Division are thanked for their procedural reviews and recommendations for improving the study design. Ms. Lindsey Lefebvre, Ms

  7. Biochemistry of hemlock (Conium maculatum L.) alkaloids and their acute and chronic toxicity in livestock. A review.

    López, T A; Cid, M S; Bianchini, M L


    The literature on Conium maculatum biochemistry and toxicology, dispersed in a large number of scientific publications, has been put together in this review. C. maculatum is a weed known almost worldwide by its toxicity to many domestic animals and to human beings. It is an Umbelliferae, characterized by long, hollow stems, reaching up to 2 m height at maturity, producing a large amount of lush foliage during its vegetative growth. Its flowers are white, grouped in umbels formed by numerous umbellules. It produces a large number of seeds that allow the plant to form thick stands in modified soils, sometimes encroaching on cultivated fields, to the extent of impeding the growth of any other vegetation inside the C. maculatum area of growth. Eight piperidinic alkaloids have been identified in this species. Two of them, gamma-coniceine and coniine are generally the most abundant and they account for most of the plant acute and chronic toxicity. These alkaloids are synthesized by the plant from eight acetate units from the metabolic pool, forming a polyketoacid which cyclises through an aminotransferase and forms gamma-coniceine as the parent alkaloid via reduction by a NADPH-dependent reductase. The acute toxicity is observed when animals ingest C. maculatum vegetative and flowering plants and seeds. In a short time the alkaloids produce a neuromuscular blockage conducive to death when the respiratory muscles are affected. The chronic toxicity affects only pregnant animals. When they are poisoned by C. maculatum during the fetuses organ formation period, the offspring is born with malformations, mainly palatoschisis and multiple congenital contractures (MCC; frequently described as arthrogryposis). Acute toxicity, if not lethal, may resolve in the spontaneous recovery of the affected animals provided further exposure to C. maculatum is avoided. It has been observed that poisoned animals tend to return to feed on this plant. Chronic toxicity is irreversible and although MCC can be surgically corrected in some cases, most of the malformed animals are lost. Since no specific antidote is available, prevention is the only way to deal with the production loses caused by this weed. Control with herbicides and grazing with less susceptible animals (such as sheep) have been suggested. C. maculatum alkaloids can be transferred to milk and to fowl muscle tissue through which the former can reach the human food chain. The losses produced by C. maculatum chronic toxicity may be largely underestimated, at least in some regions, because of the difficulty in associate malformations in offspring with the much earlier maternal poisoning.

  8. Cooperative advanced-generation breeding and testing of coastal Douglas-fir and western hemlock—strategy and implementation.

    K. J.S. Jayawickrama; G.R. Johnson; T. Ye


    As in many temperate regions of the world, forest tree improvement got underway in the Pacific Northwest of the USA in the 1950s, with a number of companies and agencies starting independent tree improvement programs. Booth-Kelly Lumber Co., Crown Zellerbach Corp., the Industrial Forestry Association, Port Blakely Mill Co., Simpson Timber Co., Timber Service Co., the...

  9. 78 FR 21099 - Grant of Authority for Subzone Status, Hemlock Semiconductor, L.L.C., (Polysilicon), Clarksville, TN


    ... DEPARTMENT OF COMMERCE Foreign-Trade Zones Board [Order No. 1895] Grant of Authority for Subzone... foreign commerce, and for other purposes,'' and authorizes the Foreign-Trade Zones Board to grant to..., and when the activity results in a significant public benefit and is in the public interest; Whereas...

  10. Multi-scale spatial controls of understory vegetation in Douglas-fir–western hemlock forests of western Oregon, USA

    Julia I. Burton; Lisa M. Ganio; Klaus J. Puettmann


    Forest understory vegetation is influenced by broad-scale variation in climate, intermediate scale variation in topography, disturbance and neighborhood interactions. However, little is known about how these multi-scale controls interact to influence observed spatial patterns. We examined relationships between the aggregated cover of understory plant species (%...

  11. Distribution of nitrogen-15 tracers applied to the canopy of a mature spruce-hemlock stand, Howland, Maine, USA

    David Bryan Dail; David Y. Hollinger; Eric A. Davidson; Ivan Fernandez; Herman C. Sievering; Neal A. Scott; Elizabeth Gaige


    In N-limited ecosystems, fertilization by N deposition may enhance plant growth and thus impact C sequestration. In many N deposition-C sequestration experiments, N is added directly to the soil, bypassing canopy processes and potentially favoring N immobilization by the soil. To understand the impact of enhanced N deposition on a low fertility unmanaged forest and...

  12. Early lessons from commercial thinning in a 30-year-old Sitka Spruce-Western Hemlock forest.

    Sarah E. Greene; William H. Emmingham


    A commercial thinning study was undertaken in a 30-year-old stand, pre-commercially thinned at 15 years of age, at Cascade Head Experimental Forest on the Oregon coast. Measurements obtained after three different thinning treatments are presented and include stand volume, basal area, current growth rate, scar damage, crown ratio, and sapwood radius. Method of...

  13. Exotic Annual Grasses in Western Rangelands: Predicting Resistance and Resilience of Native Ecosystems to Invasion (Draft)


    herbivores on Ipomopsis arizonica . In addition, litter and neighboring plants can influence herbivore damage by concealing seeds and small seedlings...1991. Facilitation and interference of Quercus douglasii on understory productivity in central California. - Ecology 72: 1484- 1499. 66 Callaway, R

  14. Intraspecific variation in Tsuga canadensis foliar chemistry

    Laura Ingwell; Joseph Brady; Matthew Fitzpatrick; Brian Maynard; Richard Casagrande; Evan Preisser


    Three groups of eastern hemlock (Tsuga canadensis Carr.) trees were analyzed to compare their chemical composition and the potential for naturally occurring resistance to hemlock woolly adelgid (Adelges tsguae...

  15. Effects of fertilizer and low rates of Imidacloprid on Adelges tsugae (Hemiptera: Adelgidae).

    S. V. Joseph; James Hanula; S. K. Braman; F. J. Byrne


    Healthy hemlock trees, Tsuga canadensis (L.) Carriere, and hemlock woolly adelgid, Adelges tsugae Annand (Hemiptera: Adelgidae), populations should favor retention and population growth of adelgid predators such as Laricobius nigrinus Fender (Coleoptera: Derodontidae) and Sasajiscymnus tsugae (Sasaji&McClure) (Coleoptera: Coccinellidae). Eastern hemlock trees...

  16. Comparison of tree size structure and growth for partially harvested and even-aged hemlock-spruce stands in southeast Alaska

    Robert L. Deal; Troy Heithecker; Eric K. Zenner


    The effects of partial cutting on tree size structure and stand growth were evaluated in 52 plots in 13 stands in southeast Alaska that were partially harvested 53 to 96 years ago and compared with 50-year-old even-aged stands that developed after clearcutting. The net basal-area growth was greater in the partially cut plots than in the uncut plots, and basal-area...

  17. Emergence, seasonality, and hybridization of Laricobius nigrinus (Coleoptera: Derodontidae), an introduced predator of hemlock woolly adelgid (Hemiptera: Adelgidae), in the Tennessee Appalachians

    Gregory J. Wiggins; Jerome F. Grant; Rusty Rhea; Albert (Bud) Mayfield; Abdul Hakeem; Paris L. Lambdin; A. B. Lamb Galloway


    From 2010 through 2013, adult emergence and seasonality of Laricobius nigrinus Fender, an introduced predatoryspecies native to western North America, as well as hybridization with the native species Laricobius rubidus(LeConte), were evaluated using emergence traps and beat-sheet sampling in areas of...

  18. Nondestructive Testing Technique to Quantify Deterioration from Marine Borer Attack in Sitka Spruce and Western Hemlock Logs: Observations from a Pilot Test

    Robert Ross; John W. Forsman; John R. Erickson; Allen M. Brackley


    Stress-wave nondestructive evaluation (NDE) techniques are used widely in the forest products industry—from the grading of wood veneer to inspection of timber structures. Inspection professionals frequently use stress-wave NDE techniques to locate internal voids and decayed or deteriorated areas in large timbers. Although these techniques have proven useful, little...

  19. Development over 25 years of Douglas-fir, western hemlock, and western redcedar planted at various spacings on a very good site in British Columbia.

    Donald L. Reukema; J. Harry G. Smith


    Results of five spacing trials on the University of British Columbia Research Forest, covering a range of plantation spacings from 1 to 5 meters, showed that choice of initial spacing is among the most important factors influencing bole and crown development and stand growth and yield. The trials include Douglas-fir (Pseudotsuga menziesi), western...

  20. Concentrations of Ca and Mg in early stages of sapwood decay in red spruce, eastern hemlock, red maple, and paper birch

    Kevin T. Smith; Walter C. Shortle; Jody Jellison; Jon Connolly; Jonathan Schilling


    The decay of coarse woody debris is a key component in the formation of forest soil and in the biogeochemical cycles of Ca and Mg. We tracked changes in density and concentration of Ca and Mg in sapwood of red maple (Acer rubrum L.), red spruce (Picea rubens Sarg.), paper birch (Betula papyrifera Marsh.), and...

  1. The foliar chemistry dynamic in eastern hemlock and how it relates to the competitive mechanism between two invasive herbivores: Adelges tsugae and Fiorinia externa.

    Mailea R. Miller-Pierce; Evan L. Preisser; Dave A. Orwig


    While invasive species themselves have been examined, little work has addressed the question of competition between two invasive specialists on a shared host. An example of this situation exists in the eastern United States, where...

  2. Seasonal differences in freezing tolerance of yellow-cedar and western hemlock trees at a site affected by yellow-cedar decline

    Paul G. Schaberg; Paul E. Hennon; Amore, David V. D; Gary J. Hawley; Catherine H. Borer; Catherine H. Borer


    To assess whether inadequate cold hardiness could be a contributor to yellow-cedar (Chamaecyparis nootkatensis (D. Don) Spach) decline, we measured the freezing tolerance of foliage from yellow-cedar trees in closed-canopy (nondeclining) and open-canopy (declining at elevations below 130 m) stands at three sites along an elevational gradient in the heart of the decline...

  3. Examining Wetland Frequency Discrepancies Produced by Data Collected at Wetland Boundaries and across the Landscape: Using Tsuga canadensis (L.) Carr. (Eastern Hemlock) as a Case Study


    Continental Canada. The Bryologist 98:218–227. Wiens, J. A. 1989. Spatial Scaling in Ecology . Functional Ecology 3:385–397. REPORT DOCUMENTATION...EPA U.S. Environmental Protection Agency FAC Facultative FACU Facultative Upland FACW Facultative Wetland FWS U.S. Fish and Wildlife Service...National and Regional Panel members from four cooperat- ing federal agencies: the Corps, the U.S. Environmental Protection Agency (EPA), the U.S. Fish and

  4. Pruning high-value Douglas-fir can reduce dwarf mistletoe severity and increase longevity in Central Oregon

    Helen M. Maffei; Gregory M. Filip; Nancy E. Grulke; Brent W. Oblinger; Ellis Q. Margolis; Kristen L. Chadwick


    Mid- to very large-sized Douglas-fir (Pseudotsuga menzieseii var. menziesii) that were lightly- to moderately-infected by dwarf mistletoe (Arceuthobium douglasii) were analyzed over a 14-year period to evaluate whether mechanical pruning could eradicate mistletoe (or at least delay the onset of severe infection) without...

  5. Coarse woody debris metrics in a California oak woodland

    William D. Tietje; Michael A. Hardy; Christopher C. Yim


    Little information is available on the metrics of coarse woody debris (CWD) in California oak woodland, most notably at the scale of the stand and woodland type. In a remote part of the National Guard Post, Camp Roberts, that has not burned in over a half century, we tallied 314 pieces of CWD in a blue oak (Quercus douglasii)-coast live oak (

  6. Modeling the effectiveness of tree planting to mitigate habitat loss in blue oak woodlands

    Richard B. Standiford; Douglas McCreary; William Frost


    Many local conservation policies have attempted to mitigate the loss of oak woodland habitat resulting from conversion to urban or intensive agricultural land uses through tree planting. This paper models the development of blue oak (Quercus douglasii) stand structure attributes over 50 years after planting. The model uses a single tree, distance...

  7. Vegetation Change in Blue Oak Woodlands in California

    Barbara A. Holzman; Barbara H. Allen-Diaz


    A preliminary report of a statewide project investigating vegetation change in blue oak (Quercus douglasii) woodlands in California is presented. Vegetation plots taken in the 1930s, as part of a statewide vegetation mapping project, were relocated and surveyed. Species composition, cover and tree stand structure data from the earlier study were...

  8. Effects of fire and browsing on regeneration of blue oak

    James W. Bartolome; Mitchel P. McClaran; Barbara H. Allen-Diaz; Jim Dunne; Lawrence D. Ford; Richard B. Standiford; Neil K. McDougald; Larry C. Forero


    Blue oaks (Quercus douglasii) are not regenerating well over much of California. The roles of fire and browsing in regeneration are probably significant, but poorly understood. We burned two foothill blue oak woodland sites which contained significant numbers of small trees between 40 and 70 cm tall, then compared height growth over 14 years among 48...

  9. Acorn Yield During 1988 and 1989 on California's Central Coast

    Sergio L. Garcia; Wayne A. Jensen; William H. Weitkamp; William D. Tietje


    In 1988, a study was began to evaluate acorn yield of valley oak (Quercus lobata), coast live oak (Q. agrifolia), and blue oak (Q. douglasii) in three of California's central coast counties: Santa Barbara, San Luis Obispo, and San Benito. The purpose of the study was to examine the degree and variability of...

  10. The Influence of Epiphytic Lichens on the Nutrient Cycling of a Blue Oak Woodland

    Johannes M. Knops; Thomas H. H. Nash III; William H. Schlesinger


    We evaluated the importance of epiphytic lichens in the nutrient cycling of a blue oak (Quercus douglasii) woodland in California. Each oak tree contained an average of 3.8 kg lichen biomass, totaling 590 kg per ha. For comparison, oak leaf biomass was 958 kg per ha. We compared tree growth, volume and composition of throughfall (rainfall falling...

  11. Blue oak plant communities of southern San Luis Obispo and northern Santa Barbara Counties, California

    Mark I. Borchert; Nancy D. Cunha; Patricia C. Krosse; Marcee L. Lawrence


    An ecological classification system has been developed for the Pacific Southwest Region of the Forest Service. As part of that classification effort, blue oak (Quercus douglasii) woodlands and forests of southern San Luis Obispo and northern Santa Barbara Counties in Los Padres National Forest were classified into I3 plant communities using...

  12. Blue Oak Canopy Effect on Seasonal Forage Production and Quality

    William E. Frost; Neil K. McDougald; Montague W. Demment


    Forage production and forage quality were measured seasonally beneath the canopy of blue oak (Quercus douglasii) and in open grassland at the San Joaquin Experimental Range. At the March and peak standing crop sampling dates forage production was significantly greater (p=.05) beneath blue oak compared to open grassland. At most sampling dates, the...

  13. Field-cage evaluation of the survival, feeding and reproduction of Laricobius osakensis (Coleoptera: Derodontidae), a predator of Adelges tsugae (Hemiptera: Adelgidae)

    L.C. Viera; S.M. Salom; M.E. Montgomery; L.T. Kok


    The hemlock woolly adelgid, Adelges tsugae Annand, is a serious, non-native pest of hemlock in eastern North America. Laricobius osakensis Montgomery and Shiyake was identified as a key predator in Japan, where A. tsugae is native. Performance of adult and immature stages of L. osakensis was...

  14. Annosus Root disease of Western Conifers (FIDL)

    Craig L. Schmitt; John R. Parmeter; John T. Kliejunas


    Annosus root disease is found on all western conifer species but is of most concern on true firs, hemlocks, and pines. Incense cedar, coast redwood and sequoia are sometimes infected in California. Western juniper is infected throughout its range. Annosus is common and causes extensive decay in old-growth western and mountain hemlock stands. Many mixed conifer stands...

  15. Cherry Creek Research Natural Area: guidebook supplement 41

    Reid Schuller; Jennie Sperling; Tim Rodenkirk


    This guidebook describes Cherry Creek Research Natural Area, a 239-ha (590-ac) area that supports old-growth Douglas-fir-western hemlock (Pseudotsuga menziesii- Tsuga heterophylla) forest occurring on sedimentary materials in the southern Oregon Coast Range. Major plant associations present within the area include the western hemlock/Oregon oxalis...

  16. Finding of No Significant Impact: SLC-4 to SLC-6 Replacement Waterline Vandenberg Air Force Base, California


    marianum). stinging nettle (Urtica dioica). poison hemlock ( Conium maculatum ). poison oak and introduced grasses dominate the understory. This site is...thistle * Coni cos a pugioniformis Slender leaved iceplant * * C ’onium maculatum Poison hemlock * Coreopsis gigantean Giant coreopsis * * C ’ortaderia

  17. Potential concerns for tree response from stem injection

    Kevin T. Smith; Phillip A. Lewis


    Stem injection of imidacloprid is an available component of management strategies for hemlock woolly adelgid. Preliminary observations of similar treatments of maple and ash show that the injury sustained by injection warrants investigation of the wound response in eastern hemlock. Such investigations need adequate experimental controls to identify the role of...

  18. Responses of southeast Alaska understory species to variation in light and soil environments

    Thomas A. Hanley; Bernard T. Bormann; Jeffrey C. Barnard; S. Mark Nay


    Aboveground growth rates of seedlings of bunchberry (Cornus canadensis L.), oval-leaf blueberry (Vaccinium ovalifolium Sm.), salmonberry (Rubus spectabilis Pursh), devilsclub (Oplopanax horridus (Sm.) Miq.), and western hemlock (Tsuga heterophylla (Raf.) Sarg.) were...

  19. Influences of Forest Tree Species and Early Spring Temperature on Surface-Atmosphere Transfers of Water and Carbon in the Northeastern U.S.

    Hadley, J. L.; Kuzeja, P.; Mulcahy, T.; Singh, S.


    Influences of Forest Tree Species and Early Spring Temperature on Surface-Atmosphere Transfers of Water and Carbon in the Northeastern U.S. Julian Hadley, Paul Kuzeja, Safina Singh and Thomas Mulcahy Transfers of water vapor from terrestrial ecosystems to the atmosphere affect regional hydrology, weather and climate over short time scales, and forest-atmosphere CO2 exchange affects global climate over long timescales. To better understand these effects for forests dominated by two very different tree species, we measured forest-atmosphere water vapor and CO2 transfers by the eddy flux technique to at two sites in central Massachusetts USA for three years. Average annual evapotranspiration (ET) for a young deciduous forest dominated by red oak (Quercus rubra L., the most abundant tree species in the area), was about 430 mm or 25 percent greater than for a coniferous forest dominated by 100 to 230 year old eastern hemlock (Tsuga canadensis L.). The difference in ET was most pronounced in July and August when the deciduous forest lost about 50 percent more water by ET in the average year (192 mm for oak forest versus 130 mm for hemlock). These data indicate that if deciduous trees with similar physiology to red oak replace hemlocks, summertime ET will increase while summer streamflow, soil water content and the extent of year- round wetlands will decrease. Increased summertime ET should also lead to slightly higher regional atmospheric humidity and precipitation. Hemlock-to-deciduous forest conversion has occurred from North Carolina to southern New England and is continuing northward as a lethal insect pest, the hemlock woolly adelgid (Adelges tsugae Annand) continues to kill hemlocks. Average annual carbon storage for the old hemlock forest in our study was about 3.3 Mg C/ha, nearly equal to the average for the deciduous forest, 3.5 Mg C/ha. This calls into question ecological theory that predicts large declines in the rate of carbon uptake for old forests, and

  20. Post-disturbance plant community dynamics following a rare natural-origin fire in a Tsuga canadensis forest.

    Bryan D Murray

    Full Text Available Opportunities to directly study infrequent forest disturbance events often lead to valuable information about vegetation dynamics. In mesic temperate forests of North America, stand-replacing crown fire occurs infrequently, with a return interval of 2000-3000 years. Rare chance events, however, may have profound impacts on the developmental trajectories of forest ecosystems. For example, it has been postulated that stand-replacing fire may have been an important factor in the establishment of eastern hemlock (Tsuga canadensis stands in the northern Great Lakes region. Nevertheless, experimental evidence linking hemlock regeneration to non-anthropogenic fire is limited. To clarify this potential relationship, we monitored vegetation dynamics following a rare lightning-origin crown fire in a Wisconsin hemlock-hardwood forest. We also studied vegetation in bulldozer-created fire breaks and adjacent undisturbed forest. Our results indicate that hemlock establishment was rare in the burned area but moderately common in the scarified bulldozer lines compared to the reference area. Early-successional, non-arboreal species including Rubus spp., Vaccinium angustifolium, sedges (Carex spp., grasses, Epilobium ciliatum, and Pteridium aquilinium were the most abundant post-fire species. Collectively, our results suggest that competing vegetation and moisture stress resulting from drought may reduce the efficacy of scarification treatments as well as the usefulness of fire for preparing a suitable seedbed for hemlock. The increasing prevalence of growing-season drought suggests that silvicultural strategies based on historic disturbance regimes may need to be reevaluated for mesic species.

  1. Functional co-operation between the nuclei of Saccharomyces cerevisiae and mitochondria from other yeast species

    Spirek, M.; Horvath, A.; Piskur, Jure


    We elaborated a simple method that allows the transfer of mitochondria from collection yeasts to Saccharomyces cerevisiae. Protoplasts prepared from different yeasts were fused to the protoplasts of the ade2-1, ura3-52, kar1-1, rho (0) strain of S. cerevisiae and were selected for respiring cybrids....... italicus, S, oviformis, S. capensis and S. chevalieri) exhibited complete compatibility with S. cerevisiae nuclei. The closely related S. douglasii mitochondrial genome could also partially restore respiration-deficiency in rho (0) S. cerevisiae, whereas mitochondrial genomes from phylogenetically less...

  2. A hierarchical approach for simulating northern forest dynamics

    Don C. Bragg; David W. Roberts; Thomas R. Crow


    Complexity in ecological systems has challenged forest simulation modelers for years, resulting in a number of approaches with varying degrees of success. Arguments in favor of hierarchical modeling are made, especially for considering a complex environmental issue like widespread eastern hemlock regeneration failure. We present the philosophy and basic framework for...

  3. Previsual detection of two conifer-infesting adelgid species in North American forest

    Stephen Cook; Karen Humes; Ryan Hruska; Christopher Williams; Grant Fraley


    The balsam woolly adelgid, Adelges piceae, and hemlock woolly adelgid, A. tsugae (Homoptera: Adelgidae), are invasive pests of coniferous forests in both the Eastern and Western United States. Balsam woolly adelgid is capable of attacking and killing native North American firs, with Fraser fir (Abies fraseri...

  4. Physical and biological responses to an alternative removal strategy of a moderate-sized dam in Washington, USA.

    Shannon Claeson; B. Coffin


    Dam removal is an increasingly practised river restoration technique, and ecological responses vary with watershed, dam and reservoir properties, and removal strategies. Moderate-sized dams, like Hemlock Dam (7.9m tall and 56m wide), are large enough that removal effects could be significant, but small enough that mitigation may be possible through a modified dam...

  5. Understanding Federal regulations as guidelines for classical biological control programs

    Michael E. Montgomery


    This chapter reviews the legislation and rules that provide the foundation for federal regulation of the introduction of natural enemies of insects as biological control agents. It also outlines the steps for complying with regulatory requirements, using biological control of Adelges tsugae Annand, the hemlock woolly adelgid (HWA), as an example. The...

  6. Voluntary Euthanasia and the Right to Die: A Dialogue with Derek Humphry.

    Sinnett, E. Robert; And Others


    Presents interview with Derek Humphry, founder of the Hemlock Society (an international right-to-die organization), who shares his personal experiences, as well as his efforts to educate the public and stimulate legal reform. Notes Humphry has dedicated more than a decade to this highly charged universal problem. (Author/ABL)

  7. Multi-decadal establishment for single-cohort Douglas-fir forests

    James A. Freund; Jerry F. Franklin; Andrew J. Larson; James A. Lutz


    The rate at which trees regenerate following stand-replacing wildfire is an important but poorly understood process in the multi-century development of Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) and western hemlock (Tsuga heterophylla (Raf.) Sarg.) forests. Temporal patterns of Douglas-fir establishment reconstructed from old-growth forests (>450 year) have...

  8. Invasions by two non-native insects alter regional forest species composition and successional trajectories

    Randall S. Morin; Andrew M. Liebhold


    While invasions of individual non-native phytophagous insect species are known to affect growth and mortality of host trees, little is known about how multiple invasions combine to alter forest dynamics over large regions. In this study we integrate geographical data describing historical invasion spread of the hemlock woolly adelgid, Adelges tsugae...

  9. Gypsy moth IPM

    Michael L. McManus; Andrew M. Liebhold


    Over the last 50 years, North American forests have been inundated by a multitude of alien pest invasions. Among these, noteworthy invaders include the hemlock woolly adelgid (Adelges tsugae), emerald ash borer (Agrilus planipennis), chestnut blight and Dutch elm disease. These species have greatly altered both the ecological and...

  10. Aluminum solubility and mobility in relation to organic carbon in surface soils affected by six tree species of the northeastern United States

    Dijkstra, F.A.; Fitzhugh, R.D.


    We compared Al solubility and mobility in surface soils among six tree species (sugar maple [Acer saccharum], white ash [Fraxinus americana], red maple [Acer rubrum, L.], American beech [Fagus grandifolia, Ehrh.], red oak [Quercus rubra, L.], and hemlock [Tsuga canadensis, Carr.]) in a mixed

  11. Weyerhaeuser Export Facility at DuPont. Volume I.


    exists on the DuPont site, in which the most common trees are lodgepole pine ( Pinus contorta) and western hemlock (Tsuga heterophtlla). Bog birch (Betula...Weyerhaeuser production facilities and storage at the proposed export facility. Tropolone methyl ethers have also been mentioned as potential contam

  12. Placing our northern hardwood woodlots under management

    Russell J. Hutnik


    Do you own a woodlot? Does it contain mostly northern hardwoods - that is, beech, birch, maple, and ash, with some hemlock and spruce? If the answers to these two questions are "yes," then you may be interested in the work that is carried on at the Bartlett Experimental Forest in New Hampshire. This is one of the field laboratories established by the U. S....

  13. Cost and productivity of new technology for harvesting and in-woods processing small-diameter trees.

    Michael B Lambert; James O. Howard


    A study was conducted on the productivity and cost of an integrated harvesting and processing system operating in small-diameter timber (western hemlock-type) on the Olympic Peninsula of western Washington. The system uses a new steep-slope fellerbuncher, a clam-bunk grapple-skidder (forwarder), a prototype chain-flail debarker delimber, a chipper, a conveyor system,...

  14. Tree improvement in the Pacific Northwest.

    R. Johnson


    Advanced-generation tree breeding programs are underway for Douglas-fir and coastal western hemlock. These programs will continue to improve growth rates and other traits. Regardless of whether seeds is from a seed orchard or natural collection, it must be used in its appropriated breeding zone or seed zone. These zones vary by species. Breeding programs are underway...

  15. Selected yield tables for plantations and natural stands in Inland Northwest Forests

    Albert R. Stage; David L. Renner; Roger C. Chapman


    Yields arrayed by site index and age have been tabulated for plantations of 500 trees per acre, with five thinning regimes, for Douglas-fir, grand fir, and western larch. Yields were also tabulated for naturally regenerated stands of the grand fir-cedar-hemlock ecosystem of the Inland Empire. All yields were estimated with the Prognosis Model for Stand Development,...

  16. Conservation of biodiversity: a useful paradigm for forest ecosystem management.

    A.B. Carey; R.O. Curtis


    The coniferous forests of the Western Hemlock Zone of western Oregon and western Washington are remarkable in the longevity and stature of their trees, long intervals between stand-replacing events, capacity to produce timber, diversity of life forms and species, and controversy over their management. The controversy is hardly new (Overton and Hunt 1974). But the...

  17. Environmental Assessment, Minuteman III and Peacekeeper Silo Elimination, Malmstrom AFB, Montana; F. E. Warren AFB, Wyoming; and Vandenberg AFB, California


    Baccharis pilularis), California sagebrush (Artemisia californica), and poison hemlock ( Conium maculatum ) are common species in the area (Vandenberg AFB...spotted bat (Euderma maculatum ) and Preble’s shrew (Sorex preblei). Habitat for the spotted bat is most often in rough, rocky, semiarid, and arid

  18. Stereoselective potencies and relative toxicities of y-Coniceine and N-Methylconiine enantiomers

    '-Coniceine, coniine, and N-methylconiine are toxic alkaloids present in poison hemlock (Conium maculatum). We previously reported the comparison of the relative potencies of (+)- and (-)-coniine enantiomers. In this study, we synthesized '-coniceine and the enantiomers of N-methylconiine and dete...

  19. Fetal-muscle type nicotinic acetylcholine receptor activation in TE-671 cells, and inhibition of fetal movement in a day 40 pregnant goat model by optical isomers of the piperidine alkaloid coniine

    Coniine is an optically active toxic piperidine alkaloid and nicotinic acetylcholine receptor (nAChR) agonist found in poison hemlock (Conium maculatum L.). Coniine teratogenicity is hypothesized to be due to the binding, activation, and prolonged desensitization of fetal muscle-type nAChR which re...

  20. Crooked Calf Syndrome: Managing Lupines on Rangelands of the Channel Scablands of East-Central Washington State

    “Crooked calf syndrome”, the contracture-type skeletal defects and cleft palate caused by velvet lupine (Lupinus leucophyllus) on the channel Scablands of east-central Washington State are the same as those defects induced by Conium maculatum (poison-hemlock) and Nicotiana spp. (wild tobacco) in rum...

  1. Steroselective Potencies and Relative Toxicities of Coniine Enantiomers

    Coniine, one of the major toxic alkaloids present in poison hemlock (Conium maculatum), occurs in two optically active forms. A comparison of the relative potencies of (+)- and (-)-coniine enantiomers has not been previously reported. In this study, we separated the enantiomers of coniine and dete...

  2. True fir spacing trials: 10-year results.

    Robert O. Curtis


    Eighteen precommercial thinning trials were established in true fir-hemlock stands in the Olympic Mountains and the west side of the Cascade Range during the period 1987 through 1994. This paper updates a previous report, with results for the first 10 years after establishment. Results are given for (1) all trees, (2) the largest 80 per acre of any species, and (3)...

  3. Preliminary test of two stump surface protectants against Fomes annosus.

    E.E. Nelson; C.Y. Li


    Two materials, monolaurin (at two concentrations) and an unidentified species of the genus Streptomyces, were tested along with borax for ability to protect freshly cut stump surfaces of western hemlock (Tsuga heterophylla (Raf.) Sarg.) from colonization by Fomes annosus. Protectants were significantly (P...

  4. Role of construction debris in release of copper, chromium, and arsenic from treated wood structures

    Stan T. Lebow; Steven A. Halverson; Jeffrey J. Morrell; John. Simonsen

    Recent research on the release of wood preservatives from treated wood used in sensitive environments has not considered the potential contribution from construction residues. This study sought to develop leaching rate data for small construction debris and compare those to the release rate from treated wood itself. Western hemlock boards were pressure treated with...

  5. Impact of Market-Based Disturbance on the Composition of West Virginia's Forest Resource

    William G. Luppold; John E. Baumgras; John E. Baumgras


    The eastern hardwood resource has been shaped by a combination of human and natural disturbances. This impact on the forest resources of West Virginia has been especially dramatic. This resource has changed from a virgin forest dominated white oak, chestnut, spruce, white pine, and hemlock in the late 19th century, to one dominated by red oak in the 1950's, to...

  6. Restoring pine barrens for avian conservation

    Greg Corace


    At first glance, many visitors to Michigan's Upper Peninsula (U.P.) see a fairly uniform forested region. Although northern hardwood forests comprised of sugar maple (Acer saccharum), American basswood (Tilia americana), yellow birch (Betula alleghaniensis) and eastern hemlock (Tsuga canadensis) predominate, the U.P. is a fact a mosaic of forest cover types...

  7. Impacts of dwarf mistletoe on the physiology of host Tsuga heterophylla trees as recorded in tree-ring C and O stable isotopes

    D. E. Marias; F. C. Meinzer; D. R. Woodruff; D. C. Shaw; S. L. Voelker; Steven W. Oak; William J. Otrosina; William D. Smith; Kamal J.K. Gandhi


    Dwarf mistletoes, obligate, parasitic plants with diminutive aerial shoots, have long-term effects on host tree water relations, hydraulic architecture and photosynthetic gas exchange and can eventually induce tree death. To investigate the long-term (1886–2010) impacts of dwarf mistletoe on the growth and gas exchange characteristics of host western hemlock, we...

  8. Dwarf mistletoe and host tree interactions in managed forests of the Pacific Northwest.

    Donald M. Knutson; Robert. Tinnin


    Dwarf mistletoes in the Pacific Northwest infect true firs, larch, pine, Douglas-fir, and hemlock. Forty-one percent of all stands east of the crest of the Cascade Range and 10 percent of west-side stands are infected. General characteristics of dwarf mistletoe are discussed including mortality and growth losses rate of spread within a tree and within stands. Relation...

  9. Pennsylvania boreal conifer forests and their bird communities: past, present, and potential

    Douglas A. Gross


    Pennsylvania spruce (Picea spp.)- and eastern hemlock (Tsuga canadensis)-dominated forests, found primarily on glaciated parts of the Allegheny Plateau, are relicts of boreal forest that covered the region following glacial retreat. The timber era of the late 1800s and early 1900s (as late as 1942) destroyed most of the boreal...

  10. Tree species effects on calcium cycling: The role of calcium uptake in deep soils

    Dijkstra, F.A.; Smits, M.M.


    Soil acidity and calcium (Ca) availability in the surface soil differ substantially beneath sugar maple (Acer saccharum) and eastern hemlock (Tsuga canadensis) trees in a mixed forest in northwestern Connecticut. We determined the effect of pumping of Ca from deep soil (rooting zone below 20-cm

  11. Vegetation Sampling for Wetland Delineation: A Review and Synthesis of Methods and Sampling Issues


    which trees are sampled via use of an angle gauge or basal area prism (Husch et al. 2003; Packard and Radtke 2007). Basal area data can be used to...refuges from fungal pathogens for seeds of eastern hemlock (Tsuga canadensis). Ecology 85(1): 284– 289. Packard, K. C., and P. J. Radtke . 2007. Forest

  12. Seasonal food selection and digestibility by tame white-tailed deer in central Maine

    Hewlette S. Crawford


    Seasonal food selection and digestibility by tame white-tailed deer (Odocoileus virginianus) were studied in the white pine (Pinus strobus)–Canada hemlock (Tsuga canadensis) and lowland conifer types, areas representative of important deer habitat in the northeastern United States. Deer selected highly...

  13. Production rates and costs of cable yarding wood residue from clearcut units

    Chris B. LeDoux


    Wood residue is a little used source of fiber, chips, and fuel because harvest costs are largely unknown. This study calculates incremental production rates and costs for yarding and loading logging residue in clearcut old-growth Douglas-fir/western hemlock forests. Harvest operations were observed for two timber sales in western Oregon. Three different cable yarding...

  14. Mapping host-species abundance of three major exotic forest pests

    Randall S. Morin; Andrew M. Liebhold; Eugene R. Luzader; Andrew J. Lister; Kurt W. Gottschalk; Daniel B. Twardus


    Periodically over the last century, forests of the Eastern United States devastated by invasive pests. We used existing data to predict the geographical extent of future damage from beech bark disease (BBD), hemlock woolly adelgid (HWA), and gypsy moth. The distributions of host species of these alien pests were mapped in 1-km2 cells by interpolating host basal area/ha...

  15. Estimating tree bole and log weights from green densities measured with the Bergstrom Xylodensimeter.

    Dale R. Waddell; Michael B. Lambert; W.Y. Pong


    The performance of the Bergstrom xylodensimeter, designed to measure the green density of wood, was investigated and compared with a technique that derived green densities from wood disk samples. In addition, log and bole weights of old-growth Douglas-fir and western hemlock were calculated by various formulas and compared with lifted weights measured with a load cell...

  16. Development of epicormic sprouts in Sitka spruce following thinning and pruning in south-east Alaska.

    Robert L. Deal; R. James Barbour; Michael H. McClellan; Dean L. Parry


    The frequency and size of epicormic sprouts in Sitka spruce (Picea sitchensis (Bong.) Carr.) were assessed in five 23-29 year-old mixed Sitka spruce-western hemlock (Tsuga heterophylla (Raf.) Sarg.) stands that were uniformly thinned and pruned to 2.4, 3.7 and 5.2 m lift heights. Six to nine years after treatment sprouts were...

  17. Landscape- vs gap-level controls on the abundance of a fire-sensitive, late-successional tree species.

    Michael C. Wimberly; Thomas A. Spies


    Tsuga heterophylla (western hemlock), a fire-sensitive, late-successional tree species, is an important component of old-growth forests in the Pacific Northwest, USA. In the Oregon Coast Range, however, T. heterophylla occurs at low densities in or is completely absent from many conifer stands. We used a cellular automata-based...

  18. The care and handling of the forest gene pool

    Roy R. Silen; Ivan Doig


    What must be the world's most magnificent pool of forest genes has timbered our Pacific slopes.Why else do the tallest firs, pines, spruces, hemlocks, redwoods, and larches all rise along the Pacific Coast of North America? Does their hugeness simply thrust up from our deep soils and mild, rainy climate? From a vantage point of three...

  19. Rhododendron maximum impacts seed bank composition and richness following Tsuga canadensis loss in riparian forests

    Tristan M. Cofer; Katherine J. Elliott; Janis K. Bush; Chelcy F. Miniat


    Southern Appalachian riparian forests have undergone changes in composition and function from invasive pathogens and pests. Castanea dentata mortality in the 1930s from chestnut blight (Cryphonectria parasitica) and Tsuga canadensis mortality in the 2000s from the hemlock woolly adelgid (Adelges tsugae) have led to the expansion and...

  20. Detection and monitoring of invasive exotic plants: a comparison of four sampling methods

    Cynthia D. Huebner


    The ability to detect and monitor exotic invasive plants is likely to vary depending on the sampling method employed. Methods with strong qualitative thoroughness for species detection often lack the intensity necessary to monitor vegetation change. Four sampling methods (systematic plot, stratified-random plot, modified Whittaker, and timed meander) in hemlock and red...

  1. DNA barcodes and molecular diagnostics to distinguish an introduced and native Laricobius (Coleoptera: Derodontidae) species in eastern North America

    G.A. Davis; N.P. Havill; Z.N. Adelman; A. Caccone; L.T. Kok; S.M. Salom


    Molecular diagnostics based on DNA barcodes can be powerful identification tools in the absence of distinctive morphological characters for distinguishing between closely related species. A specific example is distinguishing the endemic species Laricobius rubidus from Laricobius nigrinus, a biological control agent of hemlock...

  2. Species of Concern (SOC) on Department of Defense Installations



  3. 78 FR 14509 - Availability of an Environmental Assessment and Finding of No Significant Impact for a Biological...


    ... 2013. Kevin Shea, Acting Administrator, Animal and Plant Health Inspection Service. [FR Doc. 2013-05141... DEPARTMENT OF AGRICULTURE Animal and Plant Health Inspection Service [Docket No. APHIS-2012-0060... for Hemlock Woolly Adelgid AGENCY: Animal and Plant Health Inspection Service, USDA. ACTION: Notice...

  4. 76 FR 42675 - Availability of an Environmental Assessment and Finding of No Significant Impact for a Biological...


    ...). Done in Washington, DC, this 13th day of July 2011. Kevin Shea, Acting Administrator, Animal and Plant... DEPARTMENT OF AGRICULTURE Animal and Plant Health Inspection Service [Docket No. APHIS-2010-0029... for Hemlock Woolly Adelgid AGENCY: Animal and Plant Health Inspection Service, USDA. ACTION: Notice...

  5. Changes to southern Appalachian water yield and stormflow after loss of a foundation species

    Steven T. Brantley; Chelcy Ford Miniat; Katherine J. Elliott; Stephanie H. Laseter; James M. Vose


    Few studies have examined how insect outbreaks affect landscape-level hydrologic processes. We report the hydrologic effects of the invasive, exotic hemlock woolly adelgid (HWA) in a headwater catchment in the southern Appalachian Mountains. The study watershed experienced complete mortality of an evergreen tree species, Tsuga canadensis (L.) Carr. (...

  6. Leaf area prediction models for Tsuga canadensis in Maine

    Laura S. Kenefic; R.S. Seymour


    Tsuga canadensis (L.) Carr. (eastern hemlock) is a common species throughout the Acadian forest. Studies of leaf area and growth efficiency in this forest type have been limited by the lack of equations to predict leaf area of this species. We found that sapwood area was an effective leaf area surrogate in T. canadensis, though...

  7. Diagnosis of Annosus Root Disease in Mixed Conifer Forests in the Northwestern United States

    Craig L. Schmitt


    Recognizing annosus root disease affecting conifers in northwestern United States forests is discussed. Field diagnosis can bemade by observing characteristic stand patterns, wood stain and decay, ectotrophic mycelium, and sporophores. Most seriously affected trees include hemlocks, grand fir, white fir and Pacific silver fir. Ponderosa pine and other true firs may...

  8. A new species of Laricobius (Coleoptera: Derodontidae) from Japan with phylogeny and a key for native and introduced congeners in North America

    Montgomery Michael E.; S. Shiyake; Nathan P. Havill


    Laricobius osakensis Montgomery and Shiyake sp. nov., collected from Adelges tsugae Annand on hemlock [Tsuga sieboldii Carr. and Tsuga diversifolia (Maxim.) Mast.] in Japan, is described and illustrated. The new species was collected from several localities on Honshu, Shikokou, and Kyushu...

  9. Bedrock type significantly affects individual tree mortality for various conifers in the inland Northwest, U.S.A

    James A. Moore; David A Hamilton; Yu Xiao; John Byrne


    Individual tree mortality models for western white pine (Pinus monticola Dougl. ex D. Don), Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco), grand fir (Abies grandis (Dougl. ex D. Don) Lindl.), western redcedar (Thuja plicata Donn ex. D. Don), western hemlock (Tsuga heterophylla (Raf.) Sarg.), and western larch (Larix occidentalis Nutt.) were developed using data...

  10. Linking Science and Management in an Interactive Geospatial, Mutli-Criterion, Structured Decision Support Framework: Use Case Studies of the "Future Forests Geo-visualization and Decision Support Tool

    Pontius, J.; Duncan, J.


    Land managers are often faced with balancing management activities to accomplish a diversity of management objectives, in systems faced with many stress agents. Advances in ecosystem modeling provide a rich source of information to inform management. Coupled with advances in decision support techniques and computing capabilities, interactive tools are now accessible for a broad audience of stakeholders. Here we present one such tool designed to capture information on how climate change may impact forested ecosystems, and how that impact varies spatially across the landscape. This tool integrates empirical models of current and future forest structure and function in a structured decision framework that allows users to customize weights for multiple management objectives and visualize suitability outcomes across the landscape. Combined with climate projections, the resulting products allow stakeholders to compare the relative success of various management objectives on a pixel by pixel basis and identify locations where management outcomes are most likely to be met. Here we demonstrate this approach with the integration of several of the preliminary models developed to map species distributions, sugar maple health, forest fragmentation risk and hemlock vulnerability to hemlock woolly adelgid under current and future climate scenarios. We compare three use case studies with objective weightings designed to: 1) Identify key parcels for sugarbush conservation and management, 2) Target state lands that may serve as hemlock refugia from hemlock woolly adelgid induced mortality, and 3) Examine how climate change may alter the success of managing for both sugarbush and hemlock across privately owned lands. This tool highlights the value of flexible models that can be easily run with customized weightings in a dynamic, integrated assessment that allows users to hone in on their potentially complex management objectives, and to visualize and prioritize locations across the

  11. Shrinking windows of opportunity for oak seedling establishment in southern California mountains

    Davis, Frank W.; Sweet, Lynn C.; Serra-Diaz, Josep M.; Franklin, Janet; McCullough, Ian M.; Flint, Alan L.; Flint, Lorraine E.; Dingman, John; Regan, Helen M.; Syphard, Alexandra D.; Hannah, Lee; Redmond, Kelly; Moritz, Max A.


    Seedling establishment is a critical step that may ultimately govern tree species’ distribution shifts under environmental change. Annual variation in the location of seed rain and microclimates results in transient “windows of opportunity” for tree seedling establishment across the landscape. These establishment windows vary at fine spatiotemporal scales that are not considered in most assessments of climate change impacts on tree species range dynamics and habitat displacement. We integrate field seedling establishment trials conducted in the southern Sierra Nevada and western Tehachapi Mountains of southern California with spatially downscaled grids of modeled water-year climatic water deficit (CWDwy) and mean August maximum daily temperature (Tmax) to map historical and projected future microclimates suitable for establishment windows of opportunity for Quercus douglasii, a dominant tree species of warm, dry foothill woodlands, and Q. kelloggii, a dominant of cooler, more mesic montane woodlands and forests. Based on quasi-binomial regression models, Q. douglasii seedling establishment is significantly associated with modeled CWDwy and to a lesser degree with modeled Tmax. Q. kelloggii seedling establishment is most strongly associated with Tmax and best predicted by a two-factor model including CWDwy and Tmax. Establishment niche models are applied to explore recruitment window dynamics in the western Tehachapi Mountains, where these species are currently widespread canopy dominants. Establishment windows are projected to decrease by 50–95%, shrinking locally to higher elevations and north-facing slopes by the end of this century depending on the species and climate scenario. These decreases in establishment windows suggest the potential for longer-term regional population declines of the species. While many additional processes regulate seedling establishment and growth, this study highlights the need to account for topoclimatic controls and

  12. Diagenesis of conifer needles in a coastal marine environment

    Hedges, John I.; Weliky, K.


    Physically intact fir, hemlock and cedar needles were isolated from different horizons of a sediment core from a coastal marine bay (Dabob Bay, Washington State, U.S.A.) and from nearby trees and forest litter. Green fir, hemlock and cedar needles were all characterized by glucose-rich aldose mixtures (~30% of tissue carbon), the production of vanillyl and cinnamyl CuO-derived phenols (~8% of tissue carbon) and the presence of both pinitol and myo-inositol (1-2% of tissue carbon). Needles from forest litter were enriched in lignin phenols and non-glucose aldoses and depleted in glucose and cyclitols. The sediment core contained an average of 10 mg/1 of physically intact fir, hemlock and cedar needles, which occurred in similar relative abundances and accounted for less than 1% of the total nonwoody gymnosperm tissue. Compared to the green and litter counterparts, all sedimentary needles were greatly depleted in cyclitols, glucose and p-coumaric acid and enriched in vanillyl phenol precursors. The degree of elevation of vanillyl phenol yield from the degraded needles was used to estimate minimal carbon losses from the samples, which ranged from near 40% for needle litter to almost 70% for the deepest (~100 years old) sedimentary fir/hemlock samples. Although downcore increases in carbon loss and refractory organic components indicated in situ diagenesis, the bulk of overall degradation occurred either on land or during the first 10-20 years after deposition. Atomic C/N ratios of degraded needles were lower than for green counterparts, but nitrogen was lost overall. These relative changes indicate the following stability series: vanillyl phenols > N > ferulic acid, p-hydroxy phenols, most aldoses and bulk tissue > glucose and p-coumaric acid > cyclitols (near 100% loss). Vanillic acid to vanillin ratios, (Ad/Al)v, of the green fir and hemlock needles were unusually high (0.36-0.38) and decreased downcore. Diagenesis also decreased the cinnamyl/vanillyl phenol ratio

  13. Chronic impacts of invasive herbivores on a foundational forest species: a whole-tree perspective.

    Wilson, Claire M; Schaeffer, Robert N; Hickin, Mauri L; Rigsby, Chad M; Sommi, Amanda F; Thornber, Carol S; Orians, Colin M; Preisser, Evan L


    Forests make up a large portion of terrestrial plant biomass, and the long-lived woody plants that dominate them possess an array of traits that deter consumption by forest pests. Although often extremely effective against native consumers, invasive species that avoid or overcome these defenses can wreak havoc on trees and surrounding ecosystems. This is especially true when multiple invasive species co-occur, since interactions between invasive herbivores may yield non-additive effects on the host. While the threat posed by invasive forest pests is well known, long-term field experiments are necessary to explore these consumer-host interactions at appropriate spatial and temporal scales. Moreover, it is important to measure multiple variables to get a 'whole-plant' picture of their combined impact. We report the results of a four-year field experiment addressing the individual and combined impacts of two invasive herbivores, the hemlock woolly adelgid (Adelges tsugae) and elongate hemlock scale (Fiorinia externa), on native eastern hemlock (Tsuga canadensis) in southern New England. In 2011, we planted 200 hemlock saplings into a temperate forest understory and experimentally manipulated the presence/absence of both herbivore species; in 2015, we harvested the 88 remaining saplings and assessed plant physiology, growth, and resource allocation. Adelgids strongly affected hemlock growth: infested saplings had lower above/belowground biomass ratios, more needle loss, and produced fewer new needles than control saplings. Hemlock scale did not alter plant biomass allocation or growth, and its co-occurrence did not alter the impact of adelgid. While both adelgid and scale impacted the concentrations of primary metabolites, adelgid effects were more pronounced. Adelgid feeding simultaneously increased free amino acids local to feeding sites and a ~30% reduction in starch. The cumulative impact of adelgid-induced needle loss, manipulation of nitrogen pools, and the loss

  14. Changes in canopy structure and ant assemblages affect soil ecosystem variables as a foundation species declines

    Kendrick, Joseph A.; Ribbons, Relena Rose; Classen, Aimee Taylor


    in ant species composition would interact to alter soil ecosystem variables. In the Harvard Forest Hemlock Removal Experiment (HF-HeRE), established in 2003, T. canadensis in large plots were killed in place or logged and removed to mimic adelgid infestation or salvage harvesting, respectively. In 2006...... (richness and abundance) of ants increases rapidly as T. canadensis is lost from the stands. Because ants live and forage at the litter-soil interface, we hypothesized that environmental changes caused by hemlock loss (e.g., increased light and warmth at the forest floor, increased soil pH) and shifts......, we built ant exclosure subplots within all of the canopy manipulation plots to examine direct and interactive effects of canopy change and ant assemblage composition on soil and litter variables. Throughout HF-HeRE, T. canadensis was colonized by the adelgid in 2009, and the infested trees are now...

  15. Tree rings as monitors of heavy metal air pollution histories

    Kennedy, G.; Bergeron, S.


    The potential of five species of trees as historical monitors of heavy metal air pollution has been investigated. The study was carried out at a site 2 km from an industrial complex including several metal refineries. Using neutron activation, heavy metal concentrations were measured in the xylem as a function of the year of wood formation. The manganese concentrations were by far the highest. In maple trees the high natural level of this essential trace element masked any increases due to pollution. In ash and cedar increased Mn concentrations were found, relative to control trees, but there is evidence for radial translocation. In hemlock the time variations of the average Mn concentrations followed the production rates of the refineries but large variations among individual trees were observed. Hemlock was estimated to accumulate up to 0.3% of the atmospheric Mn input. (author) 13 refs.; 3 figs

  16. Interim Regional Supplement to the Corps of Engineers Wetland Delineation Manual: Western Mountains, Valleys, and Coast Region


    elevations in the local area dominated by conifers (e.g., spruce (Picea), fir (Abies), hemlock (Tsuga), Douglas-fir (Pseudotsuga), coast redwood...annual temperature is 33 to 62 ºF (1 to 17 ºC) (USDA Natural Resources Conservation Service 2006a). Low-elevation mixed conifer forests are...redox concentrations and those that do not, for hue 10YR, to meet the definition of a depleted matrix. Due to inaccurate color reproduction , do not use

  17. Green Jobs in Tennessee: Economic Impact of Green Investments

    Murat Arik


    The term green jobs has been widely used to describe jobs in businesses that are particularly related to renewable energy, energy efficiency, or environmental sustainability. The Business and Economic Research Center has partnered with the Tennessee Department of Labor and Workforce Development to estimate the economic impact of six ground-breaking green investments in Tennessee: Hemlock Semiconductor, Wacker Chemie AG, Volkswagen, Nissan Leaf and Storage Battery Manufacturing, Tennessee Sola...

  18. Effects of vegetation type on microbial biomass carbon and nitrogen in subalpine mountain forest soils.

    Ravindran, Anita; Yang, Shang-Shyng


    Microbial biomass plays an important role in nutrient transformation and conservation of forest and grassland ecosystems. The objective of this study was to determine the microbial biomass among three vegetation types in subalpine mountain forest soils of Taiwan. Tatachia is a typical high-altitude subalpine temperate forest ecosystem in Taiwan with an elevation of 1800-3952 m and consists of three vegetation types: spruce, hemlock, and grassland. Three plots were selected in each vegetation type. Soil samples were collected from the organic layer, topsoil, and subsoil. Microbial biomass carbon (Cmic) was determined by the chloroform fumigation-extraction method, and microbial biomass nitrogen (Nmic) was determined from the total nitrogen (Ntot) released during fumigation-extraction. Bacteria, actinomycetes, fungi, cellulolytic microbes, phosphate-solubilizing microbes, and nitrogen-fixing microbes were also counted. The Cmic and Nmic were highest in the surface soil and declined with the soil depth. These were also highest in spruce soils, followed by in hemlock soils, and were lowest in grassland soils. Cmic and Nmic had the highest values in the spring season and the lowest values in the winter season. Cmic and Nmic had significantly positive correlations with total organic carbon (Corg) and Ntot. Contributions of Cmic and Nmic, respectively, to Corg and Ntot indicated that the microbial biomass was immobilized more in spruce and hemlock soils than in grassland soils. Microbial populations of the tested vegetation types decreased with increasing soil depth. Cmic and Nmic were high in the organic layer and decreased with the depth of layers. These values were higher for spruce and hemlock soils than for grassland soils. Positive correlations were observed between Cmic and Nmic and between Corg and Ntot. Copyright © 2014. Published by Elsevier B.V.

  19. Field performance of timber bridges. 9, Big Erick`s stress-laminated deck bridge

    J. A. Kainz; J. P. Wacker; M. Nelson

    The Big Erickas bridge was constructed during September 1992 in Baraga County, Michigan. The bridge is 72 ft long, 16 ft wide, and consists of three simple spans: two stress-laminated deck approach spans and a stress-laminated box center span. The bridge is unique in that it is one of the first known stress-laminated timber bridge applications to use Eastern Hemlock...

  20. Environmental Assessment: Invasive Pest Plant Management


    Silvilagus floridanus Amphibians Eastern newt Notophthalmus viridescens Spotted salamander Ambystoma maculatum Two-lined salamander Eurycea...Annual EnQeron annuus c A Geranium. Carolina Geranium carotinia.num - c WA!B Geranium, Cranesbill Geranium maculatum c c WA!B Ground 01erry...Physalis heterophyffa c p Hemlock, Poison Conium macula/urn c 6 B Hen bit Lamium amplexicau/e c 3 WA/B Hoary Cress Cardarfa spp. c p Houndstongue

  1. Gods behaving badly.

    Retsas, Spyros


    This paper addresses the myths surrounding the birth and death of Asclepios, the popular healing God of the Greeks and his place among other deities of the Greek Pantheon. The enigmatic invocation of Asclepios by Socrates, the Athenian philosopher condemned to take the hemlock, in his final moments is also discussed. © The Author(s) 2013 Reprints and permissions:


    Elustondo, Diego; Avramidis, Stavros


    This paper presents an experimental evaluation of the first commercial scale dry-sort-redry (DSRD) strategy for drying of 2x4 Pacific coast hemlock (PCH) lumber. The DSRD strategy is a methodology designed to reduce final moisture content variability in kiln dried lumber by complementing conventional drying with radio frequency vacuum (RFV) drying technology. The strategy′s objective is to avoid producing over-dried lumber in conventional drying by setting the target moisture content to...

  3. Ecological setting of the Wind River old-growth forest.

    David C. Shaw; Jerry F. Franklin; Ken Bible; Jeffrey Klopatek; Elizabeth Freeman; Sarah Greene; Geoffrey G. Parker


    The Wind River old-growth forest, in the southern Cascade Range of Washington State, is a cool (average annual temperature, 8.7°C), moist (average annual precipitation, 2223 mm), 500-year-old Douglas-fir-western hemlock forest of moderate to low productivity at 371-m elevation on a less than 10% slope. There is a seasonal snowpack (November-March), and rain-on-snow and...

  4. Forest insects and diseases in Fundy National Park in 1994. Technical note No. 310

    Meikle, O.A.


    Personnel of the Forest Insect and Disease Survey regularly survey national parks for forest insect and disease conditions. This document discusses briefly some of the conditions encountered in Fundy National Park during the year, including insects and diseases found throughout the Park that are likely to recur: Gypsy moth, winter drying, sirococcus shoot blight, forest tent caterpillar, balsam fir needle cast and yellow witches` broom, birch decline, and hemlock looper.

  5. Forest insects and diseases in Fundy National Park in 1993. Technical note No. 296

    Meikle, O.A.


    Personnel of the Forest Insect and Disease Survey regularly survey national parks for forest insect and disease conditions. This document discusses briefly some of the conditions encountered in Fundy National Park during the year, including insects and diseases found throughout the Park that are likely to recur: Gypsy moth, winter drying, sirococcus shoot blight, forest tent caterpillar, balsam fir needle cast and yellow witches' broom, birch decline, and hemlock looper.

  6. Jackson Mills and Mine Falls Dams, Nashua, New Hampshire. Reconnaissance Report, Hydroelectric Feasibility. Volume 1, Jackson Mills Dam.


    Catalpa Solidago sp. Goldenrod Aster novae - angliae New England Aster Acer saccharum Sugar Maple Ulmus rubra Slippery elm Solanum hi rum pine, and hemlock are the common softwood species, and the common hardwood species include red maple, silver maple, white oak, willow, slippery ... elm and birch. In 1972, between 70 and 7S percent of the total area of the watershed consisted of forests and primarily wooded land. (Reference 3

  7. Cultural Resources in the Southern Lake Erie Basin: A Predictive Study.


    American elm , black ash, and silver maple; also associated were slippery elm , burr oak, swamp white oak, and bitternut hickory. Another type of swamp...American elm -red maple-black ash association dominated the wetter sites. Associated were slippery elm , burr oak, swamp white oak, and bitternut...americana Red elm ( slippery elm ) U. rubra Balsam fir Abies balsanea Hackberry Celtis occidentalis American hazel Corylus americana Eastern hemlock

  8. Flambeau Mining Corporation, Ladysmith, Rusk County, Wisconsin. Proposed Open Pit Copper Mine and Waste Containment Area, Draft Environmental Impact Statement.


    American elm Lonicera tatarica - tartarian honeysuckle Ulmus rubra - slippery elm Siiibucus canadenis - common elder Ulmus thoiii~sii - cork elm borders the marshes and swamps. 2.060 The predominant species are the trembling aspen (Populus tremuloides), red maple (Acer rubrum), the elms ...succession. The most numerous trees (in descending order) are: white birch, red maple, aspen, sugar maple, black ash, basswood, elm (Ulmus sp.), hemlock

  9. Postglacial vegetation history of Mitkof Island, Alexander Archipelago, southeastern Alaska

    Ager, Thomas A.; Carrara, Paul E.; Smith, Jane L.; Anne, Victoria; Johnson, Joni


    An AMS radiocarbon-dated pollen record from a peat deposit on Mitkof Island, southeastern Alaska provides a vegetation history spanning ˜12,900 cal yr BP to the present. Late Wisconsin glaciers covered the entire island; deglaciation occurred > 15,400 cal yr BP. The earliest known vegetation to develop on the island (˜12,900 cal yr BP) was pine woodland ( Pinus contorta) with alder ( Alnus), sedges (Cyperaceae) and ferns (Polypodiaceae type). By ˜12,240 cal yr BP, Sitka spruce ( Picea sitchensis) began to colonize the island while pine woodland declined. By ˜11,200 cal yr BP, mountain hemlock ( Tsuga mertensiana) began to spread across the island. Sitka spruce-mountain hemlock forests dominated the lowland landscapes of the island until ˜10,180 cal yr BP, when western hemlock ( Tsuga heterophylla) began to colonize, and soon became the dominant tree species. Rising percentages of pine, sedge, and sphagnum after ˜7100 cal yr BP may reflect an expansion of peat bog habitats as regional climate began to shift to cooler, wetter conditions. A decline in alders at that time suggests that coastal forests had spread into the island's uplands, replacing large areas of alder thickets. Cedars ( Chamaecyparis nootkatensis, Thuja plicata) appeared on Mitkof Island during the late Holocene.

  10. Tree species and soil nutrient profiles in old-growth forests of the Oregon Coast Range

    Cross, Alison; Perakis, Steven S.


    Old-growth forests of the Pacific Northwest provide a unique opportunity to examine tree species – soil relationships in ecosystems that have developed without significant human disturbance. We characterized foliage, forest floor, and mineral soil nutrients associated with four canopy tree species (Douglas-fir (Pseudotsuga menziesii (Mirbel) Franco), western hemlock (Tsuga heterophylla (Raf.) Sarg.), western redcedar (Thuja plicata Donn ex D. Don), and bigleaf maple (Acer macrophyllum Pursh)) in eight old-growth forests of the Oregon Coast Range. The greatest forest floor accumulations of C, N, P, Ca, Mg, and K occurred under Douglas-fir, primarily due to greater forest floor mass. In mineral soil, western hemlock exhibited significantly lower Ca concentration and sum of cations (Ca + Mg + K) than bigleaf maple, with intermediate values for Douglas-fir and western redcedar. Bigleaf maple explained most species-based differences in foliar nutrients, displaying high concentrations of N, P, Ca, Mg, and K. Foliar P and N:P variations largely reflected soil P variation across sites. The four tree species that we examined exhibited a number of individualistic effects on soil nutrient levels that contribute to biogeochemical heterogeneity in these ecosystems. Where fire suppression and long-term succession favor dominance by highly shade-tolerant western hemlock, our results suggest a potential for declines in both soil Ca availability and soil biogeochemical heterogeneity in old-growth forests.

  11. Red Alder-Conifer Stands in Alaska: An Example of Mixed Species Management to Enhance Structural and Biological Complexity

    Robert L. Deal


    Full Text Available There is worldwide interest in managing forests to improve biodiversity, enhance ecosystem services and assure long-term sustainability of forest resources. An increasingly important goal of forest management is to increase stand diversity and improve wildlife and aquatic habitat. Well-planned silvicultural systems containing a mixture of broadleaf-conifer species have potential to enhance stand diversity and provide other ecosystem services earlier than typical even-aged conifer plantations. Here, we use the example of mixed Sitka spruce/western hemlock and red alder in young, managed stands in southeast Alaska to achieve these goals. We briefly describe the silvics of Sitka spruce, western hemlock and red alder plantations as pure conifer stands or pure broadleaf stands. Then, we synthesize studies of mixed red alder-Sitka spruce/western hemlock stands in southeast Alaska and present their potential for improving stand structural complexity, biodiversity and other ecosystem services over pure conifer forests. Finally, we discuss some of the opportunities and potential tradeoffs for managing mixed broadleaf-conifer stands for providing a number of natural resources and the influence of these broadleaf-conifer forests on ecosystem linkages and processes.

  12. The killer of Socrates: Coniine and Related Alkaloids in the Plant Kingdom.

    Hotti, Hannu; Rischer, Heiko


    Coniine, a polyketide-derived alkaloid, is poisonous to humans and animals. It is a nicotinic acetylcholine receptor antagonist, which leads to inhibition of the nervous system, eventually causing death by suffocation in mammals. Coniine's most famous victim is Socrates who was sentenced to death by poison chalice containing poison hemlock in 399 BC. In chemistry, coniine holds two historical records: It is the first alkaloid the chemical structure of which was established (in 1881), and that was chemically synthesized (in 1886). In plants, coniine and twelve closely related alkaloids are known from poison hemlock ( Conium maculatum L.), and several Sarracenia and Aloe species. Recent work confirmed its biosynthetic polyketide origin. Biosynthesis commences by carbon backbone formation from butyryl-CoA and two malonyl-CoA building blocks catalyzed by polyketide synthase. A transamination reaction incorporates nitrogen from l-alanine and non-enzymatic cyclization leads to γ-coniceine, the first hemlock alkaloid in the pathway. Ultimately, reduction of γ-coniceine to coniine is facilitated by NADPH-dependent γ-coniceine reductase. Although coniine is notorious for its toxicity, there is no consensus on its ecological roles, especially in the carnivorous pitcher plants where it occurs. Lately there has been renewed interest in coniine's medical uses particularly for pain relief without an addictive side effect.

  13. The killer of Socrates: Coniine and Related Alkaloids in the Plant Kingdom

    Hannu Hotti


    Full Text Available Coniine, a polyketide-derived alkaloid, is poisonous to humans and animals. It is a nicotinic acetylcholine receptor antagonist, which leads to inhibition of the nervous system, eventually causing death by suffocation in mammals. Coniine’s most famous victim is Socrates who was sentenced to death by poison chalice containing poison hemlock in 399 BC. In chemistry, coniine holds two historical records: It is the first alkaloid the chemical structure of which was established (in 1881, and that was chemically synthesized (in 1886. In plants, coniine and twelve closely related alkaloids are known from poison hemlock (Conium maculatum L., and several Sarracenia and Aloe species. Recent work confirmed its biosynthetic polyketide origin. Biosynthesis commences by carbon backbone formation from butyryl-CoA and two malonyl-CoA building blocks catalyzed by polyketide synthase. A transamination reaction incorporates nitrogen from l-alanine and non-enzymatic cyclization leads to γ-coniceine, the first hemlock alkaloid in the pathway. Ultimately, reduction of γ-coniceine to coniine is facilitated by NADPH-dependent γ-coniceine reductase. Although coniine is notorious for its toxicity, there is no consensus on its ecological roles, especially in the carnivorous pitcher plants where it occurs. Lately there has been renewed interest in coniine’s medical uses particularly for pain relief without an addictive side effect.

  14. Risk assessment of imidacloprid use in forest settings on the aquatic macroinvertebrate community.

    Benton, Elizabeth P; Grant, Jerome F; Nichols, Rebecca J; Webster, R Jesse; Schwartz, John S; Bailey, Joseph K


    The isolated effects of a single insecticide can be difficult to assess in natural settings because of the presence of numerous pollutants in many watersheds. Imidacloprid use for suppressing hemlock woolly adelgid, Adelges tsugae (Annand) (Hemiptera: Adelgidae), in forests offers a rare opportunity to assess potential impacts on aquatic macroinvertebrates in relatively pristine landscapes. Aquatic macroinvertebrate communities were assessed in 9 streams in Great Smoky Mountains National Park (southern Appalachian Mountains, USA). The streams flow through hemlock conservation areas where imidacloprid soil drench treatments were applied for hemlock woolly adelgid suppression. Sites were located upstream and downstream of the imidacloprid treatments. Baseline species presence data (pre-imidacloprid treatment) were available from previous sample collections at downstream sites. Downstream and upstream sites did not vary in numerous community measures. Although comparisons of paired upstream and downstream sites showed differences in diversity in 7 streams, higher diversity was found more often in downstream sites. Macroinvertebrate functional feeding groups and life habits were similar between downstream and upstream sites. Downstream and baseline stream samples were similar. While some functional feeding group and life habit species richness categories varied, variations did not indicate poorer quality downstream communities. Imidacloprid treatments applied according to US Environmental Protection Agency federal restrictions did not result in negative effects to aquatic macroinvertebrate communities, which indicates that risks of imidacloprid use in forest settings are low. Environ Toxicol Chem 2017;36:3108-3119. © 2017 SETAC. © 2017 SETAC.

  15. A comparative toxicity assessment of materials used in aquatic construction.

    Lalonde, Benoit A; Ernst, William; Julien, Gary; Jackman, Paula; Doe, Ken; Schaefer, Rebecca


    Comparative toxicity testing was performed on selected materials that may be used in aquatic construction projects. The tests were conducted on the following materials: (1) untreated wood species (hemlock [Tsuga ssp], Western red cedar (Thuja plicata), red oak [Quercus rubra], Douglas fir [Pseudotsuga menziesii], red pine [Pinus resinosa], and tamarack [Larix ssp]); (2) plastic wood; (3) Ecothermo wood hemlock stakes treated with preservatives (e.g., chromated copper arsenate [CCA], creosote, alkaline copper quaternary [ACQ], zinc naphthenate, copper naphthenate, and Lifetime Wood Treatment); (4) epoxy-coated steel; (5) hot-rolled steel; (6) zinc-coated steel; and (7) concrete. Those materials were used in acute lethality tests with rainbow trout, Daphnia magna, Vibrio fischeri and threespine stickleback. The results indicated the following general ranking of the materials (from the lowest to highest LC(50) values); ACQ > creosote > zinc naphthenate > copper naphthenate > CCA (treated at 22.4 kg/m(3)) > concrete > red pine > western red cedar > red oak > zinc-coated steel > epoxy-coated steel > CCA (6.4 kg/m(3)). Furthermore, the toxicity results indicated that plastic wood, certain untreated wood species (hemlock, tamarack, Douglas fir, and red oak), hot-rolled steel, Ecothermo wood, and wood treated with Lifetime Wood Treatment were generally nontoxic to the test species. © Springer Science+Business Media, LLC 2011

  16. Invasive insect effects on nitrogen cycling and host physiology are not tightly linked.

    Rubino, Lucy; Charles, Sherley; Sirulnik, Abby G; Tuininga, Amy R; Lewis, James D


    Invasive insects may dramatically alter resource cycling and productivity in forest ecosystems. Yet, although responses of individual trees should both reflect and affect ecosystem-scale responses, relationships between physiological- and ecosystem-scale responses to invasive insects have not been extensively studied. To address this issue, we examined changes in soil nitrogen (N) cycling, N uptake and allocation, and needle biochemistry and physiology in eastern hemlock (Tsuga canadensis (L) Carr) saplings, associated with infestation by the hemlock woolly adelgid (HWA) (Adelges tsugae Annand), an invasive insect causing widespread decline of eastern hemlock in the eastern USA. Compared with uninfested saplings, infested saplings had soils that exhibited faster nitrification rates, and more needle (15)N uptake, N and total protein concentrations. However, these variables did not clearly covary. Further, within infested saplings, needle N concentration did not vary with HWA density. Light-saturated net photosynthetic rates (Asat) declined by 42% as HWA density increased from 0 to 3 adelgids per needle, but did not vary with needle N concentration. Rather, Asat varied with stomatal conductance, which was highest at the lowest HWA density and accounted for 79% of the variation in Asat. Photosynthetic light response did not differ among HWA densities. Our results suggest that the effects of HWA infestation on soil N pools and fluxes, (15)N uptake, needle N and protein concentrations, and needle physiology may not be tightly coupled under at least some conditions. This pattern may reflect direct effects of the HWA on N uptake by host trees, as well as effects of other scale-dependent factors, such as tree hydrology, affected by HWA activity. © The Author 2015. Published by Oxford University Press. All rights reserved. For Permissions, please email:

  17. Acceleration of exotic plant invasion in a forested ecosystem by a generalist herbivore.

    Eschtruth, Anne K; Battles, John J


    The successful invasion of exotic plants is often attributed to the absence of coevolved enemies in the introduced range (i.e., the enemy release hypothesis). Nevertheless, several components of this hypothesis, including the role of generalist herbivores, remain relatively unexplored. We used repeated censuses of exclosures and paired controls to investigate the role of a generalist herbivore, white-tailed deer (Odocoileus virginianus), in the invasion of 3 exotic plant species (Microstegium vimineum, Alliaria petiolata, and Berberis thunbergii) in eastern hemlock (Tsuga canadensis) forests in New Jersey and Pennsylvania (U.S.A.). This work was conducted in 10 eastern hemlock (T. canadensis) forests that spanned gradients in deer density and in the severity of canopy disturbance caused by an introduced insect pest, the hemlock woolly adelgid (Adelges tsugae). We used maximum likelihood estimation and information theoretics to quantify the strength of evidence for alternative models of the influence of deer density and its interaction with the severity of canopy disturbance on exotic plant abundance. Our results were consistent with the enemy release hypothesis in that exotic plants gained a competitive advantage in the presence of generalist herbivores in the introduced range. The abundance of all 3 exotic plants increased significantly more in the control plots than in the paired exclosures. For all species, the inclusion of canopy disturbance parameters resulted in models with substantially greater support than the deer density only models. Our results suggest that white-tailed deer herbivory can accelerate the invasion of exotic plants and that canopy disturbance can interact with herbivory to magnify the impact. In addition, our results provide compelling evidence of nonlinear relationships between deer density and the impact of herbivory on exotic species abundance. These findings highlight the important role of herbivore density in determining impacts on

  18. Professor Stewart's casebook of mathematical mysteries

    Stewart, Ian


    Like its wildly popular predecessors Cabinet of Mathematical Curiosities and Hoard of Mathematical Treasures, Professor Stewart's brand-new book is a miscellany of over 150 mathematical curios and conundrums, packed with trademark humour and numerous illustrations. In addition to the fascinating formulae and thrilling theorems familiar to Professor Stewart's fans, the Casebook follows the adventures of the not-so-great detective Hemlock Soames and his sidekick Dr John Watsup (immortalised in the phrase 'Watsup, Doc?'). By a remarkable coincidence they live at 222B Baker Street, just a

  19. Forest pest conditions in the maritimes in 1991. Information report No. M-X-181E. Annual publication

    Magasi, L.P.


    This report reviews the status of forest insects and diseases in the Maritimes region in 1991 and forecasts conditions for 1992, when appropriate. Pests and problems of conifers, hardwoods, and high-value areas, such as nurseries, seed orchards, plantations, and Christmas tree areas, are described as observed in 1991. Control operations against spruce budworm, hemlock looper, and Sirococcus shoot blight are summarized. A section on forest health monitoring brings together the various aspects of work dealing with changes in forest conditions, some of which are still unexplained. Forest insect monitoring systems, pheromones, and light traps are briefly described. A list of reports and publications relating to forest pest conditions is included.

  20. Operational Draft Regional Guidebook for the Functional Assessment of High-gradient Ephemeral and Intermittent Headwater Streams in Western West Virginia and Eastern Kentucky


    Liriodendron tulipifera tuliptree Ulmus rubra slippery elm Magnolia acuminata cucumber-tree Ulmus thomasii rock elm ERDC/EL TR-10-11 103 Watershed...americana American elm ERDC/EL TR-10-11 41 0 0.5 1 1.5 2 2.5 SRV per 100 feet of stream reach 0 0.1 0.2 0.3 0.4 0.5 0.6 0.7 0.8 0.9 1 Va ria bl e S...canadensis eastern hemlock Juglans nigra black walnut Ulmus americana American elm Liquidambar styraciflua sweetgum Ulmus parvifolia Chinese elm

  1. Ozone exposure and stomatal sluggishness in different plant physiognomic classes

    Paoletti, Elena, E-mail: e.paoletti@ipp.cnr.i [IPP-CNR, Via Madonna del Piano 10, I-50019 Sesto Fiorentino, Florence (Italy); Grulke, Nancy E. [US Forest Service, 4955 Canyon Crest Drive, Riverside, CA 92507 (United States)


    Gas exchange responses to static and variable light were tested in three species: snap bean (Phaseolus vulgaris, two cultivars), California black oak (Quercus kelloggii), and blue oak (Q. douglasii). The effects of 1-month (snap beans) and 2-month (oaks) O{sub 3} (ozone) exposure (70 ppb over 8 h per day in open-top chambers) were investigated. A delay in stomatal responses (i.e., 'sluggish' responses) to variable light was found to be both an effect of O{sub 3} exposure and a reason for increased O{sub 3} sensitivity in snap bean cultivars, as it implied higher O{sub 3} uptake during times of disequilibrium. Sluggishness increased the time to open (thus limiting CO{sub 2} uptake) and close stomata (thus increasing transpirational water loss) after abrupt changes in light level. Similar responses were shown by snap beans and oaks, suggesting that O{sub 3}-induced stomatal sluggishness is a common trait among different plant physiognomic classes. - Sluggish stomatal responses are suggested to be both an effect of O{sub 3} exposure and a reason of increased O{sub 3} sensitivity in plants.

  2. Rising synchrony controls western North American ecosystems

    Black, Bryan A.; van der Sleen, Peter; Di Lorenzo, Emanuele; Griffin, Daniel; Sydeman, William J.; Dunham, Jason B.; Rykaczewski, Ryan R.; Garcia-Reyes, Marisol; Safeeq, Mohammad; Arismendi, Ivan; Bograd, Steven J.


    Along the western margin of North America, the winter expression of the North Pacific High (NPH) strongly influences interannual variability in coastal upwelling, storm track position, precipitation, and river discharge. Coherence among these factors induces covariance among physical and biological processes across adjacent marine and terrestrial ecosystems. Here, we show that over the past century the degree and spatial extent of this covariance (synchrony) has substantially increased, and is coincident with rising variance in the winter NPH. Furthermore, centuries‐long blue oak (Quercus douglasii) growth chronologies sensitive to the winter NPH provide robust evidence that modern levels of synchrony are among the highest observed in the context of the last 250 years. These trends may ultimately be linked to changing impacts of the El Niño Southern Oscillation on mid‐latitude ecosystems of North America. Such a rise in synchrony may destabilize ecosystems, expose populations to higher risks of extinction, and is thus a concern given the broad biological relevance of winter climate to biological systems.

  3. Data from camera surveys identifying co-occurrence and occupancy linkages between fishers (Pekania pennanti, rodent prey, mesocarnivores, and larger predators in mixed-conifer forests

    Rick A. Sweitzer


    Full Text Available These data provide additional information relevant to the frequency of fisher detections by camera traps, and single-season occupancy and local persistence of fishers in small patches of forest habitats detailed elsewhere, “Landscape Fuel Reduction, Forest Fire, and Biophysical Linkages to Local Habitat Use and Local Persistence of Fishers (Pekania pennanti in Sierra Nevada Mixed-conifer Forests” [10]. The data provides insight on camera trap detections of 3 fisher predators (bobcat [Lynx rufus]. Coyote [Canis latrans], mountain lion [Puma concolor], 5 mesocarnivores in the same foraging guild as fishers (gray fox [Urocyon cinereoargenteus] ringtail [Bassariscus astutus], marten [Martes americana], striped skunk [Mephitis mephitis] spotted skunk [Spilogale gracilis], and 5 Sciuridae rodents that fishers consume as prey (Douglas squirrel [Tamiasciurus douglasii], gray squirrel [Sciurus griseus], northern flying squirrel [Glaucomys sabrinus], long-eared chipmunk [Neotamias quadrimaculatus], California ground squirrel [Spermophilus beecheyi]. We used these data to identify basic patterns of co-occurrence with fishers, and to evaluate the relative importance of presence of competing mesocarnivores, rodent prey, and predators for fisher occupancy of small, 1 km2 grid cells of forest habitat. Keywords: Carnivores, Competition, Distribution, Foraging guild, Predation, Tree squirrels

  4. Averaged 30 year climate change projections mask opportunities for species establishment

    Serra-Diaz, Josep M.; Franklin, Janet; Sweet, Lynn C.; McCullough, Ian M.; Syphard, Alexandra D.; Regan, Helen M.; Flint, Lorraine E.; Flint, Alan L.; Dingman, John; Moritz, Max A.; Redmond, Kelly T.; Hannah, Lee; Davis, Frank W.


    Survival of early life stages is key for population expansion into new locations and for persistence of current populations (Grubb 1977, Harper 1977). Relative to adults, these early life stages are very sensitive to climate fl uctuations (Ropert-Coudert et al. 2015), which often drive episodic or ‘event-limited’ regeneration (e.g. pulses) in long-lived plant species (Jackson et al. 2009). Th us, it is diffi cult to mechanistically associate 30-yr climate norms to dynamic processes involved in species range shifts (e.g. seedling survival). What are the consequences of temporal aggregation for estimating areas of potential establishment? We modeled seedling survival for three widespread tree species in California, USA ( Quercus douglasii, Q. kelloggii , Pinus sabiniana ) by coupling a large-scale, multi-year common garden experiment to high-resolution downscaled grids of climatic water defi cit and air temperature (Flint and Flint 2012, Supplementary material Appendix 1). We projected seedling survival for nine climate change projections in two mountain landscapes spanning wide elevation and moisture gradients. We compared areas with windows of opportunity for seedling survival – defi ned as three consecutive years of seedling survival in our species, a period selected based on studies of tree niche ontogeny (Supplementary material Appendix 1) – to areas of 30-yr averaged estimates of seedling survival. We found that temporal aggregation greatly underestimated the potential for species establishment (e.g. seedling survival) under climate change scenarios.

  5. Anthropogenic Disturbances Create a New Vegetation Toposequence in the Gatineau River Valley, Quebec

    Jason Laflamme


    Full Text Available This study measured changes in forest composition that have occurred since the preindustrial era along the toposequence of the Gatineau River Valley, Quebec, Canada (5650 km2, based on survey records prior to colonization (1804–1864 and recent forest inventories (1982–2006. Changes in forest cover composition over time were found to be specific to toposequence position. Maple and red oak are now more frequent on upper toposequence positions (+26%, +21%, respectively, whereas yellow birch, eastern hemlock, and American beech declined markedly (−34% to −17%. Poplar is more frequent throughout the landscape, but particularly on mid-toposequence positions (+40%. In contrast, white pine, frequent on all toposequence positions in the preindustrial forest, is now confined to shallow and coarse-textured soils (−20%. The preindustrial forest types of the study area were mostly dominated by maple, yellow birch, and beech, with strong components of white pine, hemlock, and eastern white cedar, either as dominant or codominant species. In a context of ongoing anthropogenic disturbances and environmental changes, it is probably not possible to restore many of these types, except where targeted silvicultural interventions could increase the presence of certain species. The new forest types observed should be managed to ensure continuity of vital ecosystem services and functions as disturbance regimes evolve.

  6. Effects of climate change, land-use change, and invasive species on the ecology of the Cumberland forests

    Dale, V.H.; Fogel, J.


    The mixed mesophytic forests of the Cumberland Plateau and Mountains in Tennessee and Kentucky are among the most diverse forests in North America. However, land use changes and climatic warming will have a significant impact on the forest biomass and composition in the region, which currently experiences mild winters and hot, humid summers. In this study, 3 general circulation models projected climatic warming throughout 2030 to 2080 as well as changes in precipitation patterns. Predicted changes from 1980 to 2100 were used in a forest ecosystem model to estimate transient changes in forest biomass and species composition over time. Results of the study demonstrated that climatic warning will cause an initial decline in forest stand biomass before a recovery period caused by forest species composition shifts. A landscape model showed that forest composition will change as a result of the spread of hemlock adelgid. Loss of the hemlocks will cause changes in soil nutrients and moisture conditions in mesic forests of the region. Land cover changes will be large and cause declines in forested lands as well as in a number of large, contiguous forest patches that provide a necessary habitat for species particular to the Cumberland area. 53 refs., 2 tabs., 8 figs

  7. Tree-ring stable isotopes record the impact of a foliar fungal pathogen on CO(2) assimilation and growth in Douglas-fir.

    Saffell, Brandy J; Meinzer, Frederick C; Voelker, Steven L; Shaw, David C; Brooks, J Renée; Lachenbruch, Barbara; McKay, Jennifer


    Swiss needle cast (SNC) is a fungal disease of Douglas-fir (Pseudotsuga menziesii) that has recently become prevalent in coastal areas of the Pacific Northwest. We used growth measurements and stable isotopes of carbon and oxygen in tree-rings of Douglas-fir and a non-susceptible reference species (western hemlock, Tsuga heterophylla) to evaluate their use as proxies for variation in past SNC infection, particularly in relation to potential explanatory climate factors. We sampled trees from an Oregon site where a fungicide trial took place from 1996 to 2000, which enabled the comparison of stable isotope values between trees with and without disease. Carbon stable isotope discrimination (Δ(13)C) of treated Douglas-fir tree-rings was greater than that of untreated Douglas-fir tree-rings during the fungicide treatment period. Both annual growth and tree-ring Δ(13)C increased with treatment such that treated Douglas-fir had values similar to co-occurring western hemlock during the treatment period. There was no difference in the tree-ring oxygen stable isotope ratio between treated and untreated Douglas-fir. Tree-ring Δ(13)C of diseased Douglas-fir was negatively correlated with relative humidity during the two previous summers, consistent with increased leaf colonization by SNC under high humidity conditions that leads to greater disease severity in following years. © 2013 John Wiley & Sons Ltd.

  8. Asymmetric biotic interactions and abiotic niche differences revealed by a dynamic joint species distribution model.

    Lany, Nina K; Zarnetske, Phoebe L; Schliep, Erin M; Schaeffer, Robert N; Orians, Colin M; Orwig, David A; Preisser, Evan L


    A species' distribution and abundance are determined by abiotic conditions and biotic interactions with other species in the community. Most species distribution models correlate the occurrence of a single species with environmental variables only, and leave out biotic interactions. To test the importance of biotic interactions on occurrence and abundance, we compared a multivariate spatiotemporal model of the joint abundance of two invasive insects that share a host plant, hemlock woolly adelgid (HWA; Adelges tsugae) and elongate hemlock scale (EHS; Fiorina externa), to independent models that do not account for dependence among co-occurring species. The joint model revealed that HWA responded more strongly to abiotic conditions than EHS. Additionally, HWA appeared to predispose stands to subsequent increase of EHS, but HWA abundance was not strongly dependent on EHS abundance. This study demonstrates how incorporating spatial and temporal dependence into a species distribution model can reveal the dependence of a species' abundance on other species in the community. Accounting for dependence among co-occurring species with a joint distribution model can also improve estimation of the abiotic niche for species affected by interspecific interactions. © 2018 by the Ecological Society of America.

  9. Entomopathogenic Fungi Associated with Exotic Invasive Insect Pests in Northeastern Forests of the USA

    Gouli, Vladimir; Gouli, Svetlana; Marcelino, José A. P.; Skinner, Margaret; Parker, Bruce L.


    Mycopathogens of economically important exotic invasive insects in forests of northeastern USA have been the subject of research at the Entomology Research Laboratory, University of Vermont, for the last 20 years. Elongate hemlock scale, European fruit lecanium, hemlock woolly adelgid and pear thrips were analyzed for the presence of mycopathogens, in order to consider the potential for managing these pests with biological control. Fungal cultures isolated from insects with signs of fungal infection were identified based on morphological characters and DNA profiling. Mycopathogens recovered from infected insects were subdivided into three groups, i.e., specialized entomopathogenic; facultative entomopathogens; ubiquitous opportunistic contaminants. Epizootics were caused by fungi in the specialized group with the exception of M. microspora, P. marquandii and I. farinosa. Inoculation of insects in laboratory and field conditions with B. bassiana, L. muscarium and Myriangium sp. caused insect mortality of 45 to 95%. Although pest populations in the field seemed severely compromised after treatment, the remnant populations re-established themselves after the winter. Although capable of inducing high mortality, a single localized aerial application of a soil-dwelling fungus does not maintain long-time suppression of pests. However, it can halt their range expansion and maintain populations below the economic threshold level without the use of expensive insecticides which have a negative impact on the environment. PMID:26462527

  10. Effects of climate change, land-use change, and invasive species on the ecology of the Cumberland forests

    Dale, V.H. [Oak Ridge National Laboratory, Oak Ridge, TN (United States). Environmental Sciences Div.; Lannom, K.O.; Hodges, D.G. [Tennessee Univ., Knoxville, TN (United States). Natural Resource Policy Center; Tharp, M.L. [CompSci Consulting LLC, McRae, GA (United States); Fogel, J. [Virginia Tech Univ., Richmond, VA (United States). Virginia Cooperative Extension, Northeast District Office


    The mixed mesophytic forests of the Cumberland Plateau and Mountains in Tennessee and Kentucky are among the most diverse forests in North America. However, land use changes and climatic warming will have a significant impact on the forest biomass and composition in the region, which currently experiences mild winters and hot, humid summers. In this study, 3 general circulation models projected climatic warming throughout 2030 to 2080 as well as changes in precipitation patterns. Predicted changes from 1980 to 2100 were used in a forest ecosystem model to estimate transient changes in forest biomass and species composition over time. Results of the study demonstrated that climatic warning will cause an initial decline in forest stand biomass before a recovery period caused by forest species composition shifts. A landscape model showed that forest composition will change as a result of the spread of hemlock adelgid. Loss of the hemlocks will cause changes in soil nutrients and moisture conditions in mesic forests of the region. Land cover changes will be large and cause declines in forested lands as well as in a number of large, contiguous forest patches that provide a necessary habitat for species particular to the Cumberland area. 53 refs., 2 tabs., 8 figs.

  11. Detecting influences on California drought intervals using isotopes in tree-ring cellulose

    Kanner, L. C.; Buenning, N. H.; Stott, L. D.; Stahle, D. W.


    Multi-decadal drought events have characterized climate variability in California over the last century. However, the causes of interannual precipitation variability and the origins of multi-decadal drought in California remain unclear. We utilize the oxygen isotopic composition (δ18O) of tree-ring cellulose in combination with previously developed ring-width measurements to trace the delivery of moisture to the region and investigate ocean-atmosphere patterns that might generate prolonged drought. Of the 36 Quercus douglasii (blue oak) sites in the California central valley, we have focused our work at two locations - one north of Los Angeles (34.74°N, 120°W, 1036 masl) and the other east of San Francisco (37.88°N 121.97°W, 182 masl). Using cores from at least five different trees at each location, tree-ring cellulose δ18O was measured for each year of growth from 1954 to 2004. The δ18O values of tree-ring cellulose range from 29‰ to 34‰ (VSMOW) at both sites and exhibit shared interannual variance (r = 0.43, p shifts in the moisture source region are of primary importance because moisture from high latitude sources has a lower isotopic composition compared to subtropical regions. Using NCAR reanalysis data, wind field anomalies suggest that moisture is derived from the north during dry years (low δ18O) and from the subtropics during wet years (high δ18O). Additional processes such as condensation height and post-condensation effects may also be important in controlling isotopic variability.

  12. Seasonal distributions of the western cherry fruit fly (Diptera: Tephritidae) among host and nonhost fruit trees.

    Yee, Wee L


    Seasonal distributions of the western cherry fruit fly, Rhagoletis indifferens Curran (Diptera: Tephritidae), in sweet cherry (Prunus avium (L.) L.) (major host), black hawthorn (occasional developmental host) (Crataegus douglasii Lindley), and other trees were determined in a ponderosa pine ecosystem in Washington state, USA. The hypothesis that most fly dispersal from cherry trees occurs after fruit senesce or drop was tested, with emphasis on movement to black hawthorn trees. Sweet cherry fruit developed earlier than black hawthorn, bitter cherry (common host), choke cherry, and apple fruit. Flies were usually captured first in sweet cherry trees but were caught in bitter cherry and other trees throughout the season. Peak fly capture periods in sweet cherry began around the same time or slightly earlier than in other trees. However, peak fly capture periods in black hawthorn and other nonsweet cherry trees continued after peak periods in sweet cherry ended, or relative fly numbers within sweet cherry declined more quickly than those within other trees. Larvae were reared from sweet and bitter cherry but not black hawthorn fruit. Results provide partial support for the hypothesis in that although R. indifferens commonly disperses from sweet cherry trees with fruit, it could disperse more, or more flies are retained in nonsweet cherry trees after than before sweet cherries drop. This could allow opportunities for the flies to use other fruit for larval development. Although R. indifferens infestation in black hawthorn was not detected, early season fly dispersal to this and other trees and fly presence in bitter cherry could make fly management in sweet cherry difficult. Published by Oxford University Press on behalf of the Entomological Society of America 2014. This work is written by a US Government employee and is in the public domain in the US.

  13. Seasonality and Interannual Variability of Carbon Uptake and Respiration in a California Oak Savanna

    Ma, S.; Baldocchi, D.; Xu, L.


    Estimating terrestrial carbon sink with large-scale modeling research requires understanding the physiological and ecological processes associated with the carbon uptake and respiration of ecosystems and their variability in seasons and years. This study was conducted in an oak/grass savanna ecosystem in California, USA. The savanna ecosystem consists of blue oak trees ( Quercus douglasii) in the overstory and annual C3 grasses in the understory. Fluxes of CO2 were measured above the canopy (overstory) and the grasses (understory) from 2001 to 2005 with two eddy covariance systems. Under typical Mediterranean Climate, net ecosystem exchange of CO2 (NEE), ecosystem respiration (Reco), and gross primary production (GPP) in this savanna ecosystem had a distinctive dry-wet seasonal pattern. Leaf area index, leaf nitrogen concentration, and leaf carbon stable isotope discrimination reflected the responses of leaf to the seasonality and interannual variability. Light- use efficiency, the ratio of GPP to absorbed photosynthetically active radiation (aPAR), was not consistent within a year or from year to year, indicating that photosynthesis process was constrained with low temperature during the beginning of the wet season and limited by precipitation during the summer drought. Annual NEE, Reco, and GPP above the canopy varied significantly between years, varying from -108 - 133 gC m-2, 780 - 988 gC m-2, and 646 - 963 gC m-2, respectively. The difference of interannual Reco was 1.2 times of that of interannual GPP. There was a tight relationship between annual NEE and the precipitation during the period with daily mean temperature varying between 10 - 20°C, equivalent to precipitation during March and April. The longer the period lasted, the higher carbon uptake occurred. Estimated annual NEE from 1949 - 2005 in the savanna ecosystem varied between ~-400 - 200 gC m-2.

  14. Late Glacial-Holocene Pollen-Based Vegetation History from Pass Lake, Prince of Wales Island, Southeastern Alaska

    Ager, Thomas A.; Rosenbaum, Joseph G.


    A radiocarbon-dated history of vegetation development since late Wisconsin deglaciation has been reconstructed from pollen evidence preserved in a sediment core from Pass Lake on Prince of Wales Island, southeastern Alaska. The shallow lake is in the south-central part of the island and occupies a low pass that was filled by glacial ice of local origin during the late Wisconsin glaciation. The oldest pollen assemblages indicate that pine woodland (Pinus contorta) had developed in the area by ~13,715 cal yr B.P. An abrupt decline in the pine population, coinciding with expansion of alder (Alnus) and ferns (mostly Polypodiaceae) began ~12,875 yr B.P., and may have been a response to colder, drier climates during the Younger Dryas climatic interval. Mountain hemlock (Tsuga mertensiana) began to colonize central Prince of Wales Island by ~11,920 yr B.P. and was soon followed by Sitka spruce (Picea sitchensis). Pollen of western hemlock (Tsuga heterophylla) began to appear in Pass Lake sediments soon after 11,200 yr B.P. The abundance of western hemlock pollen in the Pass Lake core during most of the Holocene appears to be the result of wind transport from trees growing at lower altitudes on the island. The late Holocene pollen record from Pass Lake is incomplete because of one or more unconformities, but the available record suggests that a vegetation change occurred during the late Holocene. Increases in pollen percentages of pine, cedar (probably yellow cedar, Chamaecyparis nootkatensis), and heaths (Ericales) suggest an expansion of muskeg vegetation occurred in the area during the late Holocene. This vegetation change may be related to the onset of cooler, wetter climates that began as early as ~3,774 yr B.P. in the region. This vegetation history provides the first radiocarbon-dated Late Glacial-Holocene terrestrial paleoecological framework for Prince of Wales Island. An analysis of magnetic properties of core sediments from Pass Lake suggests that unconformities

  15. Transformation of Values, Identities & Life Styles at Iranian Society in Cinema Narration

    Jamal Mohammadi


    Full Text Available This article is to do a sociologically analysis on transformation of values, Identities and life styles at Iranian post Islamic revolution society in cinema narrations. The method of study is qualitative; first the selected movies have been analyzed by applying semiotic narration analysis based on elements such as images, shots, actions, mise en secene, dialogues and scenes. Then their main social categories have been extracted and sociologically analyzed through thematic analysis strategy based on theoretical foundation of research. The population of research includes all post revolution Iranian cinema movies which 12 movies have been selected by purposive sampling method. These movies include; Narges, Fair People Wedding, Glass Agency, Under City Cover, Protest, Blue Scarf, Hemlock, Boutique, Wednesday Celebration, Trial in Street, about Eli & Separation of Nader & Simin. Findings show that selected movies narrate gaps, distinctness of social life domains, of instrumental rationality expansion & split of value systems

  16. Alkaloid-Containing Plants Poisonous to Cattle and Horses in Europe

    Cristina Cortinovis


    Full Text Available Alkaloids, nitrogen-containing secondary plant metabolites, are of major interest to veterinary toxicology because of their occurrence in plant species commonly involved in animal poisoning. Based on epidemiological data, the poisoning of cattle and horses by alkaloid-containing plants is a relatively common occurrence in Europe. Poisoning may occur when the plants contaminate hay or silage or when forage alternatives are unavailable. Cattle and horses are particularly at risk of poisoning by Colchicum autumnale (meadow saffron, Conium maculatum (poison hemlock, Datura stramonium (jimson weed, Equisetum palustre (marsh horsetail, Senecio spp. (ragwort and groundsel and Taxus baccata (European yew. This review of poisonous alkaloid-containing plants describes the distribution of these plants, conditions under which poisoning occurs, active toxic principles involved and subsequent clinical signs observed.

  17. Stereoselective potencies and relative toxicities of coniine enantiomers.

    Lee, Stephen T; Green, Benedict T; Welch, Kevin D; Pfister, James A; Panter, Kip E


    Coniine, one of the major toxic alkaloids present in poison hemlock ( Conium maculatum), occurs in two optically active forms. A comparison of the relative potencies of (+)- and (-)-coniine enantiomers has not been previously reported. In this study, we separated the enantiomers of coniine and determined the biological activity of each enantiomer in vitro and in vivo. The relative potencies of these enantiomers on TE-671 cells expressing human fetal nicotinic neuromuscular receptors had the rank order of (-)-coniine > (+/-)-coniine > (+)-coniine. A mouse bioassay was used to determine the relative lethalities of (-)-, (+/-)-, and (+)-coniine in vivo. The LD 50 values of the coniine enantiomers were 7.0, 7.7, and 12.1 mg/kg for the (-)-, (+/-)-, and (+)- forms of coniine, respectively. The results from this study demonstrate that there is a stereoselective difference in the in vitro potencies of the enantiomers of coniine that directly correlates with the relative toxicities of the enantiomers in vivo.

  18. [The attempts at drug therapy of cancer by Anton Störck (1731-1803). History of experimental pharmacology in the old Vienna Medical School].

    Schweppe, K W; Probst, C


    The essay deals with the development of medical research in Vienna - especially the development of therapeutic drugs. This progress is related to the philosophical, historical, and political background of the enlightened absolutism and the reformatory efforts of van Swieten during the regency of Maria Theresia in Austria. Anton Störck's research on hemlock (Conium maculatum) is used as an example. The method of Störck's research-work is described. Furthermore it is demonstrated to what extent Störck's data, deduced from empirical examinations, are integrated in the official medical system, i.e. Boerhaave's iatromechanic system. Finally the attempt is made to correlate these processes of medical history with the scientific-historical model of Thomas Kuhn.

  19. Antifeedant compounds from three species of Apiaceae active against the field slug, Deroceras reticulatum (Muller).

    Birkett, Michael A; Dodds, Catherine J; Henderson, Ian F; Leake, Lucy D; Pickett, John A; Selby, Martin J; Watson, Peter


    Extracts of volatiles from foliage of three plants in the Apiaceae, Conium maculatum L. (hemlock), Coriandrum sativum L. (coriander), and Petroselinum crispum Mill. (Nym.) (parsley), previously shown to exhibit antifeedant activity in assays with the field slug, Deroceras reticulatum (Muller) (Limacidae: Pulmonata), were studied further to identify the active components. Coupled gas chromatography-mass spectrometry (GC-MS) and neurophysiological assays using tentacle nerve preparations resulted in the identification of 11 active compounds from the three extracts. Wheat flour feeding bioassays were used to determine which of these compounds had the highest antifeedant activity. One of the most active compounds was the alkaloid gamma-coniceine, from C. maculatum. The role of potentially toxic alkaloids as semiochemicals and the potential for using such compounds as crop protection agents to prevent slug feeding damage is discussed.

  20. Alkaloid-Containing Plants Poisonous to Cattle and Horses in Europe.

    Cortinovis, Cristina; Caloni, Francesca


    Alkaloids, nitrogen-containing secondary plant metabolites, are of major interest to veterinary toxicology because of their occurrence in plant species commonly involved in animal poisoning. Based on epidemiological data, the poisoning of cattle and horses by alkaloid-containing plants is a relatively common occurrence in Europe. Poisoning may occur when the plants contaminate hay or silage or when forage alternatives are unavailable. Cattle and horses are particularly at risk of poisoning by Colchicum autumnale (meadow saffron), Conium maculatum (poison hemlock), Datura stramonium (jimson weed), Equisetum palustre (marsh horsetail), Senecio spp. (ragwort and groundsel) and Taxus baccata (European yew). This review of poisonous alkaloid-containing plants describes the distribution of these plants, conditions under which poisoning occurs, active toxic principles involved and subsequent clinical signs observed.

  1. Piperidine alkaloids: human and food animal teratogens.

    Green, Benedict T; Lee, Stephen T; Panter, Kip E; Brown, David R


    Piperidine alkaloids are acutely toxic to adult livestock species and produce musculoskeletal deformities in neonatal animals. These teratogenic effects include multiple congenital contracture (MCC) deformities and cleft palate in cattle, pigs, sheep, and goats. Poisonous plants containing teratogenic piperidine alkaloids include poison hemlock (Conium maculatum), lupine (Lupinus spp.), and tobacco (Nicotiana tabacum) [including wild tree tobacco (Nicotiana glauca)]. There is abundant epidemiological evidence in humans that link maternal tobacco use with a high incidence of oral clefting in newborns; this association may be partly attributable to the presence of piperidine alkaloids in tobacco products. In this review, we summarize the evidence for piperidine alkaloids that act as teratogens in livestock, piperidine alkaloid structure-activity relationships and their potential implications for human health. Copyright © 2012 Elsevier Ltd. All rights reserved.

  2. Congenital skeletal malformations and cleft palate induced in goats by ingestion of Lupinus, Conium and Nicotiana species.

    Panter, K E; Keeler, R F; Bunch, T D; Callan, R J


    Three piperidine alkaloid containing plants, Conium maculatum (poison-hemlock), Nicotiana glauca (tree tobacco) and Lupinus formosus (lunara lupine), induced multiple congenital contractures (MCC) and palatoschisis in goat kids when their dams were gavaged with the plant during gestation days 30-60. The skeletal abnormalities included fixed extension or flexure of the carpal, tarsal, and fetlock joints, scoliosis, lordosis, torticollis and rib cage abnormalities. Clinical signs of toxicity included those reported in sheep, cattle and pigs--ataxia, incoordination, muscular weakness, prostration and death. One quinolizidine alkaloid containing plant, Lupinus caudatus (tailcup lupine), on the other hand, which is also known to cause MCC in cows, caused only slight signs of toxicity in pregnant goats and no teratogenic effects in their offspring.

  3. Assessing Structure and Condition of Temperate And Tropical Forests: Fusion of Terrestrial Lidar and Airborne Multi-Angle and Lidar Remote Sensing

    Saenz, Edward J.

    Forests provide vital ecosystem functions and services that maintain the integrity of our natural and human environment. Understanding the structural components of forests (extent, tree density, heights of multi-story canopies, biomass, etc.) provides necessary information to preserve ecosystem services. Increasingly, remote sensing resources have been used to map and monitor forests globally. However, traditional satellite and airborne multi-angle imagery only provide information about the top of the canopy and little about the forest structure and understory. In this research, we investigative the use of rapidly evolving lidar technology, and how the fusion of aerial and terrestrial lidar data can be utilized to better characterize forest stand information. We further apply a novel terrestrial lidar methodology to characterize a Hemlock Woolly Adelgid infestation in Harvard Forest, Massachusetts, and adapt a dynamic terrestrial lidar sampling scheme to identify key structural vegetation profiles of tropical rainforests in La Selva, Costa Rica.

  4. Exploratory radio-frequency/vacuum drying of three B.C. coastal softwoods

    Avramidis, S.; Zwick, R.L.


    Five exploratory drying runs were carried out in a commercial size (23 m3) radio-frequency/vacuum (RF/V) kiln. The species investigated were Pacific Coast hemlock, Douglas-fir, and western redcedar of different sizes and grades. Evaluation of the dried lumber showed that the three species can be dried in very short times with a low amount of degrade with the exception of clear cedar, which exhibited severe internal honeycombing; indicating the need to develop proper kiln schedules for this drying process. The thermodynamic and sensitivity analyses performed showed that above certain efficiency levels, RF/V drying is economical and fast. This finding was particularly true with lumber thicknesses over 8 cm

  5. Radiation curing of mixture of diallylphthalate prepolymer and vinyl monomer, 11

    Gotoda, Masao; Yagi, Toshiaki; Yoshizawa, Iwao; Okugawa, Hideo.


    Preparation of Wood-Polymer Composite (WPC) piled plywood for flooring, was attempted by electron beam irradiation of American Hemlock veneer, impregnated with low molecular weight diallylphthalate prepolymer (DAPsub(p).L)/vinyl monomer mixtures, in comparison with that by thermal curing reported previously. For the improvement of the mechanical properties of WPC piled plywood, 5-15 parts of low molecular weight polyvinylacetate (PVAc) was added to DAPsub(p).L/methyl acrylate/hydroxyethyl acrylate/benzoyl peroxide (50/40/10/2) mixture. The addition of PVAc did not give much influence to the degree of impregnation in the veneer and to the curability by electron beam irradiation. As to the resistance to hot and cold cycle test, WPC piled plywood, prepared by the resin containing 5 parts of PVAc gave the best results. As a general conclusion, due to the superior mechanical properties, the WPC piled plywood processed by electron beam irradiation is suitable for flooring use. (author)

  6. Evaluating the Contribution of Climate Forcing and Forest Dynamics to Accelerating Carbon Sequestration by Forest Ecosystems in the Northeastern U.S.: Final Report

    Munger, J. William [Harvard University, SEAS; Foster, David R. [Harvard University, Harvard Forest; Richardson, Andrew D. [Harvard University, OEB


    This report summarizes work to improve quantitative understanding of the terrestrial ecosystem processes that control carbon sequestration in unmanaged forests It builds upon the comprehensive long-term observations of CO2 fluxes, climate and forest structure and function at the Harvard Forest in Petersham, MA. This record includes the longest CO2 flux time series in the world. The site is a keystone for the AmeriFlux network. Project Description The project synthesizes observations made at the Harvard Forest HFEMS and Hemlock towers, which represent the dominant mixed deciduous and coniferous forest types in the northeastern United States. The 20+ year record of carbon uptake at Harvard Forest and the associated comprehensive meteorological and biometric data, comprise one of the best data sets to challenge ecosystem models on time scales spanning hourly, daily, monthly, interannual and multi-decadal intervals, as needed to understand ecosystem change and climate feedbacks.

  7. Draft Environmental Impact Report/Environmental Impact Statement. Cullinan Ranch Specific Plan. Appendices.


    0XandA±liA Poison hemlock Cgn~i lfAQUIAtW Hottentot-fig 14espmbrvunthe~mmgjj jaumea 1LU31A carnos * Sea lavender Lim~nium califgrnicum miner’s lettuce ...MeagubryAnthomuw odu~lef Jaumea. JAI.mA ZAXn2M Sea lavender Liumnim £&JJ.±oLnicim Miner’s lettuce Montia perflnaa Li nanthus LTfAnDnhILE granAilau...PROPUCT IN P 100 W 4tM POLLINATION Il 12.10 190 1320 TOTAL 1901 93,00 II I ECONOMIC FEASIBILITY OF THE ". CULLINAN RANCH FOR AGR I CULTURAL PRODUCT I ON 1

  8. Ecosystem, location, and climate effects on foliar secondary metabolites of lodgepole pine populations from central British Columbia.

    Wallis, Christopher M; Huber, Dezene P W; Lewis, Kathy J


    Lodgepole pines, Pinus contorta Douglas ex Louden var. latifolia Engelm. ex S. Watson, are encountering increased abiotic stress and pest activity due to recent increases in temperature and changes in precipitation throughout their range. This tree species counters these threats by producing secondary metabolites, including phenolics and terpenoids. We examined foliar levels of lignin, soluble phenolics, monoterpenoids, sesquiterpenoids, and diterpenoids in 12 stands in British Columbia, Canada. We used these data to assess associations among foliar secondary metabolite levels and ecosystem, geographic, and climatic variables. Regressions were also performed to observe which combinations of variables best explained secondary metabolite variance. Stands of P. c. latifolia in the Coastal Western Hemlock and Interior Cedar/Hemlock biogeoclimatic zones had consistently greater foliar levels of almost all measured secondary metabolites than did other stands. Lignin was present in greater amounts in Boreal White/Black Spruce ecosystem (i.e., northern) stands than in southern stands, suggesting a role for this metabolite in pine survival in the boreal forest. Attempts to develop regression models with geographic and climatic variables to explain foliar secondary metabolite levels resulted in multiple models with similar predictive capability. Since foliar secondary metabolite levels appeared to vary most between stand ecosystem types and not as much due to geographic and climatic variables, metabolic profiles appeared best matched to the stress levels within local environments. It is unknown if differences in secondary metabolite levels are the result of genetic adaptation or phenotypic plasticity, but results from this and other studies suggest that both are important. These results are interpreted in light of ongoing efforts to assist in the migration of certain populations of P. c. latifolia northward in an effort to counter predicted effects of climate change.

  9. The potential to characterize ecological data with terrestrial laser scanning in Harvard Forest, MA

    Boucher, P.; Saenz, E.; Li, Z.


    Contemporary terrestrial laser scanning (TLS) is being used widely in forest ecology applications to examine ecosystem properties at increasing spatial and temporal scales. Harvard Forest (HF) in Petersham, MA, USA, is a long-term ecological research (LTER) site, a National Ecological Observatory Network (NEON) location and contains a 35 ha plot which is part of Smithsonian Institution's Forest Global Earth Observatory (ForestGEO). The combination of long-term field plots, eddy flux towers and the detailed past historical records has made HF very appealing for a variety of remote sensing studies. Terrestrial laser scanners, including three pioneering research instruments: the Echidna Validation Instrument, the Dual-Wavelength Echidna Lidar and the Compact Biomass Lidar, have already been used both independently and in conjunction with airborne laser scanning data and forest census data to characterize forest dynamics. TLS approaches include three-dimensional reconstructions of a plot over time, establishing the impact of ice storm damage on forest canopy structure, and characterizing eastern hemlock (Tsuga canadensis) canopy health affected by an invasive insect, the hemlock woolly adelgid (Adelges tsugae). Efforts such as those deployed at HF are demonstrating the power of TLS as a tool for monitoring ecological dynamics, identifying emerging forest health issues, measuring forest biomass and capturing ecological data relevant to other disciplines. This paper highlights various aspects of the ForestGEO plot that are important to current TLS work, the potential for exchange between forest ecology and TLS, and emphasizes the strength of combining TLS data with long-term ecological field data to create emerging opportunities for scientific study. PMID:29503723

  10. The potential to characterize ecological data with terrestrial laser scanning in Harvard Forest, MA.

    Orwig, D A; Boucher, P; Paynter, I; Saenz, E; Li, Z; Schaaf, C


    Contemporary terrestrial laser scanning (TLS) is being used widely in forest ecology applications to examine ecosystem properties at increasing spatial and temporal scales. Harvard Forest (HF) in Petersham, MA, USA, is a long-term ecological research (LTER) site, a National Ecological Observatory Network (NEON) location and contains a 35 ha plot which is part of Smithsonian Institution's Forest Global Earth Observatory (ForestGEO). The combination of long-term field plots, eddy flux towers and the detailed past historical records has made HF very appealing for a variety of remote sensing studies. Terrestrial laser scanners, including three pioneering research instruments: the Echidna Validation Instrument, the Dual-Wavelength Echidna Lidar and the Compact Biomass Lidar, have already been used both independently and in conjunction with airborne laser scanning data and forest census data to characterize forest dynamics. TLS approaches include three-dimensional reconstructions of a plot over time, establishing the impact of ice storm damage on forest canopy structure, and characterizing eastern hemlock ( Tsuga canadensis ) canopy health affected by an invasive insect, the hemlock woolly adelgid ( Adelges tsugae ). Efforts such as those deployed at HF are demonstrating the power of TLS as a tool for monitoring ecological dynamics, identifying emerging forest health issues, measuring forest biomass and capturing ecological data relevant to other disciplines. This paper highlights various aspects of the ForestGEO plot that are important to current TLS work, the potential for exchange between forest ecology and TLS, and emphasizes the strength of combining TLS data with long-term ecological field data to create emerging opportunities for scientific study.

  11. Recognizing Non-Stationary Climate Response in Tree Growth for Southern Coastal Alaska, USA

    Wiles, G. C.; Jarvis, S. K.; D'Arrigo, R.; Vargo, L. J.; Appleton, S. N.


    Stationarity in growth response of trees to climate over time is assumed in dendroclimatic studies. Recent studies of Alaskan yellow-cedar (Chamaecyparis nootkatensis (D. Don) Spach) have identified warming-induced early loss of insulating snowpack and frost damage as a mechanism that can lead to decline in tree growth, which for this species is documented over the last century. A similar stress may be put on temperature-sensitive mountain hemlock (Tsuga mertensiana (Bong.) Carrière) trees at low elevations, which in some cases show a decline in tree growth with warming temperatures. One of the challenges of using tree-ring based SAT, SST, PDO and PNA-related reconstructions for southern coastal Alaska has been understanding the response of tree-ring chronologies to the warming temperatures over the past 50 years. Comparisons of tree growth with long meteorological records from Sitka Alaska that extend back to 1830 suggest many mountain hemlock sites at low elevations are showing decreasing ring-widths, at mid elevations most sites show a steady increasing growth tracking warming, and at treeline a release is documented. The recognition of this recent divergence or decoupling of tree-ring and temperature trends allows for divergence-free temperature reconstructions using trees from moderate elevations. These reconstructions now provide a better perspective for comparing recent warming to Medieval warming and a better understanding of forest dynamics as biomes shift in response to the transition from the Little Ice Age to contemporary warming. Reconstructed temperatures are consistent with well-established, entirely independent tree-ring dated ice advances of land-terminating glaciers along the Gulf of Alaska providing an additional check for stationarity in the reconstructed interval.

  12. Pruning high-value Douglas-fir can reduce dwarf mistletoe severity and increase longevity in central Oregon

    Maffei, Helen M; Filip, Gregory M; Gruelke, Nancy E; Oblinger, Brent W; Margolis, Ellis; Chadwick, Kristen L


    Mid- to very large-sized Douglas-fir (Pseudotsuga menzieseii var. menziesii) that were lightly- to moderately-infected by dwarf mistletoe (Arceuthobium douglasii) were analyzed over a 14-year period to evaluate whether mechanical pruning could eradicate mistletoe (or at least delay the onset of severe infection) without significantly affecting tree vitality and by inference, longevity. Immediate and longterm pruning effects on mistletoe infection severity were assessed by comparing pruned trees (n = 173) to unpruned trees (n = 55) with respect to: (1) percentage of trees with no visible infections 14 years post-pruning, (2) Broom Volume Rating (BVR), and (3) rate of BVR increase 14 years postpruning. Vitality/longevity (compared with unpruned trees) was assessed using six indicators: (1) tree survival, (2) the development of severe infections, (3) the development of dead tops, (4) tree-ring width indices, (5) Normalized Difference Vegetation Index (NDVI) from high-resolution multi-spectral imagery, and (6) live-crown ratio (LCR) and increment. Twenty-four percent of the pruned trees remained free of mistletoe 14 years post-pruning. Pruning is most likely to successfully eradicate mistletoe in lightly infected trees (BVR 1 or 2) without infected neighbors. Pruning significantly decreased mean BVR in the pruned versus the unpruned trees. However, the subsequent average rate of intensification (1.3–1.5 BVR per decade) was not affected, implying that a single pruning provides ~14 years respite in the progression of infection levels. Post-pruning infection intensification was slower on dominant and co-dominants than on intermediate or suppressed trees. The success of mistletoe eradication via pruning and need for follow-up pruning should be evaluated no sooner than 14 years after pruning to allow for the development of detectable brooms. Based on six indicators, foliage from witches brooms contribute little to long-term tree vitality since removal appears to have

  13. Measurement of Leaf Mass and Leaf Area of Oaks In A Mediterranean-climate Region For Biogenic Emission Estimation

    Karlik, J.

    Given the key role played by biogenic volatile organic compounds (BVOC) in tro- pospheric chemistry and regional air quality, it is critical to generate accurate BVOC emission inventories. Because several oak species have high BVOC emission rates, and oak trees are often of large stature with corresponding large leaf masses, oaks may be the most important genus of woody plants for BVOC emissions modeling in the natural landscapes of Mediterranean-climate regions. In California, BVOC emis- sions from oaks may mix with anthropogenic emissions from urban areas, leading to elevated levels of ozone. Data for leaf mass and leaf area for a stand of native blue oaks (Quercus douglasii) were obtained through harvest and leaf removal from 14 trees lo- cated in the Sierra Nevada foothills of central California. Trees ranged in height from 4.2 to 9.9 m, with trunk diameters at breast height of 14 to 85 cm. Mean leaf mass density was 730 g m-2 for the trees and had an overall value of 310 g m-2 for the site. Consideration of the surrounding grassland devoid of trees resulted in a value of about 150 g m-2, less than half of reported values for eastern U.S. oak woodlands, but close to a reported value for oaks found in St. Quercio, Italy. The mean value for leaf area index (LAI) for the trees at this site was 4.4 m2 m-2. LAI for the site was 1.8 m2 m-2, but this value was appropriate for the oak grove only; including the surrounding open grassland resulted in an overall LAI value of 0.9 m2 m-2 or less. A volumetric method worked well for estimating the leaf mass of the oak trees. Among allometric relationships investigated, trunk circumference, mean crown radius, and crown projec- tion were well correlated with leaf mass. Estimated emission of isoprene (mg C m-2 h-1) for the site based these leaf mass data and experimentally determined emission rate was similar to that reported for a Mediterranean oak woodland in France.

  14. PREFACE: The Eighth Liquid Matter Conference The Eighth Liquid Matter Conference

    Dellago, Christoph; Kahl, Gerhard; Likos, Christos N.


    The Eighth Liquid Matter Conference (LMC8) was held at the Universität Wien from 6-10 September 2011. Initiated in 1990, the conferences of this series cover a broad range of highly interdisciplinary topics, ranging from simple liquids to soft matter and biophysical systems. The vast spectrum of scientific subjects presented and discussed at the LMC8 is reflected in the themes of the ten symposia: Ionic and quantum liquids, liquid metals Water, solutions and reaction dynamics Liquid crystals Polymers, polyelectrolytes, biopolymers Colloids Films, foams, surfactants, emulsions, aerosols Confined fluids, interfacial phenomena Supercooled liquids, glasses, gels Non-equilibrium systems, rheology, nanofluids Biofluids, active matter This special issue contains scientific papers, authored by participants of the LMC8, which provide a cross-section of the scientific activities in current liquid matter science, as discussed at the conference, and demonstrate the scientific as well as methodological progress made in this field over the past couple of years. The Eighth Liquid Matter Conference contents The Eighth Liquid Matter ConferenceChristoph Dellago, Gerhard Kahl and Christos N Likos Comparing light-induced colloidal quasicrystals with different rotational symmetriesMichael Schmiedeberg and Holger Stark Hydrogen bond network relaxation in aqueous polyelectrolyte solutions: the effect of temperatureS Sarti, D Truzzolillo and F Bordi Equilibrium concentration profiles and sedimentation kinetics of colloidal gels under gravitational stressS Buzzaccaro, E Secchi, G Brambilla, R Piazza and L Cipelletti The capillary interaction between two vertical cylindersHimantha Cooray, Pietro Cicuta and Dominic Vella Hydrodynamic and viscoelastic effects in polymer diffusionJ Farago, H Meyer, J Baschnagel and A N Semenov A density-functional theory study of microphase formation in binary Gaussian mixturesM Carta, D Pini, A Parola and L Reatto Microcanonical determination of the

  15. Detection of long-term trends in carbon accumulation by forests in Northeastern U. S. and determination of causal factors: Final report

    J. William Munger; Steven C. Wofsy; David R. Foster


    The overall project goal was to quantify the trends and variability for Net ecosystem exchange of CO{sub 2}, H{sub 2}O, and energy by northeastern forests, with particular attention to the role of succession, differences in species composition, legacies of past land use, and disturbances. Measurements included flux measurements and observations of biomass accumulation using ecosystem modeling as a framework for data interpretation. Continuation of the long-term record at the Environmental Measurement Site (EMS) Tower was a priority. The final quality-assured CO{sub 2}-flux data now extend through 2010. Data through 2011 are collected but not yet finalized. Biomass observations on the plot array centered on the tower are extended to 2011. Two additional towers in a hemlock stand (HEM) and a younger deciduous stand (LPH) complement the EMS tower by focusing on stands with different species composition or age distribution and disturbance history, but comparable climate and soil type. Over the period since 1993 the forest has added 24.4 Mg-C ha{sup -1} in the living trees. Annual net carbon uptake had been increasing from about 2 Mg-C ha{sup -1}y{sup -1} in the early 1990s to nearly 6 Mg-C ha{sup -1}y{sup -1} by 2008, but declined in 2009-2010. We attribute the increasing carbon uptake to a combination of warmer temperatures, increased photosynthetic efficiency, and increased influence by subcanopy hemlocks that are active in the early spring and late autumn when temperatures are above freezing but the deciduous canopy is bare. Not all of the increased carbon accumulation was found in woody biomass. Results from a study using data to optimize parameters in an ecosystem process model indicate that significant changes in model parameters for photosynthetic capacity and shifts in allocation to slow cycling soil organic matter are necessary for the model to match the observed trends. The emerging working hypothesis is that the pattern of increasing carbon uptake over the

  16. Kootenai River White Sturgeon Investigations; White Sturgeon Spawning and Recruitment Evaluation, 2003-2004 Annual Report.

    Rust, Pete; Wakkinen, Virginia (Idaho Department of Fish and Game, Boise, ID)


    The objective of this research was to determine the environmental requirements for successful spawning and recruitment of the Kootenai River white sturgeon Acipenser transmontanus population. Annual tasks include monitoring and evaluating the various life stages of Kootenai River white sturgeon. Sampling for adult Kootenai River white sturgeon in 2003 began in March and continued through April. Eighty-one adult white sturgeon were captured with 3,576 hours of angling and set-lining effort in the Kootenai River. Discharge from Libby Dam and river stage at Bonners Ferry in 2003 peaked in May and early June. Flows remained above 500 m{sup 3}/s throughout June, decreased rapidly through mid July, and increased back to near 500 m{sup 3}/s after mid July and through mid August. By late August, flows had decreased to below 400 m{sup 3}/s. We monitored the movements of 24 adult sturgeon in Kootenay Lake, British Columbia (BC) and the Kootenai River from March 15, 2003 to August 31, 2003. Some of the fish were radio or sonic tagged in previous years. Twelve adult white sturgeon were moved upstream to the Hemlock Bar reach (rkm 260.0) and released as part of the Set and Jet Program. Transmitters were attached to seven of these fish, and their movements were monitored from the time of release until they moved downstream of Bonners Ferry. Eight additional radio-tagged white sturgeon adults were located in the traditional spawning reach (rkm 228-240) during May and June. Sampling with artificial substrate mats began May 21, 2003 and ended June 30, 2003. We sampled 717 mat d (a mat d is one 24 h set) during white sturgeon spawning. Three white sturgeon eggs were collected near Shortys Island on June 3, 2003, and five eggs were collected from the Hemlock Bar reach on June 5, 2003. Prejuvenile sampling began June 17, 2003 and continued until July 31, 2003. Sampling occurred primarily at Ambush Rock (rkm 244.0) in an attempt to document any recruitment that might have occurred from

  17. Holocene Paleohydrological Changes in Northern Michigan: Interpretations of Biomarker Distributions and Compound Specific Stable Isotope Analysis from Peatlands

    Nichols, J. E.; Booth, R. K.; Jackson, S. T.; Pendall, E. G.; Huang, Y.


    Sediments of ombrotrophic peatlands are excellent archives for reconstructing past changes in precipitation/evaporation (P/E) balance. Multiproxy analysis of these sediments is critical for better understanding of climatic events experienced by these highly sensitive systems, as each proxy may respond to different climate parameters. In this study, we use distributions of n-alkanes and δD of Sphagnum biomarkers to interpret paleohydrology from sediments of Irwin Smith Bog, northern Michigan. We then integrate these data with pollen data and testate amoebae-inferred water table depth. Sphagnum moss is the dominant peat former in ombrotrophic bogs, but vascular plants become abundant when water tables are drawn down. Thus, the abundance of Sphagnum relative to vascular plants is indicative of peatland hydrology. The n-alkanes produced by Sphagnum differ from vascular plants in the relative abundance of the different homologues, with the former having excess amounts of shorter chain C23 n-alkane. We use several measures (compound ratios, PCA) to show changes in then-alkane distributions in the sediments, and thus changes in the peatland plant community. Our data provide high- resolution, quantitative paleohydrological records for the study region that are consistent with other records. We also show that the relative abundance of a newly identified Sphagnum biomarker, 2-heptacosanone, can be used to reconstruct changing plant communities. Because ombrotrophic systems lose water by evaporation, drier/warmer conditions cause hydrogen isotopic enrichment of bog water and Sphagnum biomarkers. We measured the δD of C23 n-alkane and 2-heptacosanone to provide additional paleoclimate information. Our multiproxy approach allows us to better understand the climate changes during key intervals of the Holocene. For example, a sharp decrease in the abundance of Tsuga canadensis (hemlock) pollen has been previously identified in records from many places throughout eastern North

  18. Experiments based on blue intensity for reconstructing North Pacific temperatures along the Gulf of Alaska

    Wilson, Rob; D'Arrigo, Rosanne; Andreu-Hayles, Laia; Oelkers, Rose; Wiles, Greg; Anchukaitis, Kevin; Davi, Nicole


    Ring-width (RW) records from the Gulf of Alaska (GOA) have yielded a valuable long-term perspective for North Pacific changes on decadal to longer timescales in prior studies but contain a broad winter to late summer seasonal climate response. Similar to the highly climate-sensitive maximum latewood density (MXD) proxy, the blue intensity (BI) parameter has recently been shown to correlate well with year-to-year warm-season temperatures for a number of sites at northern latitudes. Since BI records are much less labour intensive and expensive to generate than MXD, such data hold great potential value for future tree-ring studies in the GOA and other regions in mid- to high latitudes. Here we explore the potential for improving tree-ring-based reconstructions using combinations of RW- and BI-related parameters (latewood BI and delta BI) from an experimental subset of samples at eight mountain hemlock (Tsuga mertensiana) sites along the GOA. This is the first study for the hemlock genus using BI data. We find that using either inverted latewood BI (LWBinv) or delta BI (DB) can improve the amount of explained temperature variance by > 10 % compared to RW alone, although the optimal target season shrinks to June-September, which may have implications for studying ocean-atmosphere variability in the region. One challenge in building these BI records is that resin extraction did not remove colour differences between the heartwood and sapwood; thus, long term trend biases, expressed as relatively warm temperatures in the 18th century, were noted when using the LWBinv data. Using DB appeared to overcome these trend biases, resulting in a reconstruction expressing 18th-19th century temperatures ca. 0.5 °C cooler than the 20th-21st centuries. This cool period agrees well with previous dendroclimatic studies and the glacial advance record in the region. Continuing BI measurement in the GOA region must focus on sampling and measuring more trees per site (> 20) and compiling

  19. Experiments based on blue intensity for reconstructing North Pacific temperatures along the Gulf of Alaska

    R. Wilson


    Full Text Available Ring-width (RW records from the Gulf of Alaska (GOA have yielded a valuable long-term perspective for North Pacific changes on decadal to longer timescales in prior studies but contain a broad winter to late summer seasonal climate response. Similar to the highly climate-sensitive maximum latewood density (MXD proxy, the blue intensity (BI parameter has recently been shown to correlate well with year-to-year warm-season temperatures for a number of sites at northern latitudes. Since BI records are much less labour intensive and expensive to generate than MXD, such data hold great potential value for future tree-ring studies in the GOA and other regions in mid- to high latitudes. Here we explore the potential for improving tree-ring-based reconstructions using combinations of RW- and BI-related parameters (latewood BI and delta BI from an experimental subset of samples at eight mountain hemlock (Tsuga mertensiana sites along the GOA. This is the first study for the hemlock genus using BI data. We find that using either inverted latewood BI (LWBinv or delta BI (DB can improve the amount of explained temperature variance by > 10 % compared to RW alone, although the optimal target season shrinks to June–September, which may have implications for studying ocean–atmosphere variability in the region. One challenge in building these BI records is that resin extraction did not remove colour differences between the heartwood and sapwood; thus, long term trend biases, expressed as relatively warm temperatures in the 18th century, were noted when using the LWBinv data. Using DB appeared to overcome these trend biases, resulting in a reconstruction expressing 18th–19th century temperatures ca. 0.5 °C cooler than the 20th–21st centuries. This cool period agrees well with previous dendroclimatic studies and the glacial advance record in the region. Continuing BI measurement in the GOA region must focus on sampling and measuring more trees per

  20. Development and testing of a snow interceptometer to quantify canopy water storage and interception processes in the rain/snow transition zone of the North Cascades, Washington, USA

    Martin, Kael A.; Van Stan, John T.; Dickerson-Lange, Susan E.; Lutz, James A.; Berman, Jeffrey W.; Gersonde, Rolf; Lundquist, Jessica D.


    Tree canopy snow interception is a significant hydrological process, capable of removing up to 60% of snow from the ground snowpack. Our understanding of canopy interception has been limited by our ability to measure whole canopy water storage in an undisturbed forest setting. This study presents a relatively inexpensive technique for directly measuring snow canopy water storage using an interceptometer, adapted from Friesen et al. (2008). The interceptometer is composed of four linear motion position sensors distributed evenly around the tree trunk. We incorporate a trunk laser-mapping installation method for precise sensor placement to reduce signal error due to sensor misalignment. Through calibration techniques, the amount of canopy snow required to produce the measured displacements can be calculated. We demonstrate instrument performance on a western hemlock (Tsuga heterophylla) for a snow interception event in November 2011. We find a snow capture efficiency of 83 ± 15% of accumulated ground snowfall with a maximum storage capacity of 50 ± 8 mm snow water equivalent (SWE). The observed interception event is compared to simulated interception, represented by the variable infiltration capacity (VIC) hydrologic model. The model generally underreported interception magnitude by 33% using a leaf area index (LAI) of 5 and 16% using an LAI of 10. The interceptometer captured intrastorm accumulation and melt rates up to 3 and 0.75 mm SWE h-1, respectively, which the model failed to represent. While further implementation and validation is necessary, our preliminary results indicate that forest interception magnitude may be underestimated in maritime areas.

  1. Actual state of plant decline phenomena due to acid rain and others

    Koshiji, Masashi


    Kanagawa Prefecture presents diversified aspects in its topography and weather from Sagami Bay to the highest point, Hirugatake (1673 m) in Tanzawa mountains, with plain, terrace, hill and mountain. When the state of forest areas is observed, evergreen broadleaf tree forests are in coastal areas and to the inland of 700 - 800 m elevation. Deciduous broadleaf tree forests are in the mountains exceeding 700 - 800 m. Further, fir and Japanese hemlock trees distribute in the area where both forests adjoin. At present, from terraces and hills to mountains, the plantations of Japanese cedar, Hinoki cypress and pine trees and the secondary forests of oak trees are seen. The decline of Japanese cedar trees in this Prefecture was first seen around 1960, and it was investigated in 1972 by dividing the tree form into five steps corresponding to the degree of decline. The progress of the decline, and the outline of fir tree forest in Oyama and its decline are reported. The secular change of growth ring width is utilized as the recorder of environmental change in the past such as weather, natural calamity and atmospheric pollution. The analysis of the course of decline using growth rings was carried out. The relation of the decline of forests to atmospheric pollution, and the decline of Japanese beech and fir trees in Tanzawa mountains are reported. (K.I.)

  2. Transport and concentration of abscisic acid (ABA) and auxin (IAA) in deciduous and coniferous trees. Transport und Gehalt von Abscisinsaeure (ABA) und Auxin (IAA) in Laub- und Nadelblaettern

    Hartung, W.


    Abscisic acid and indoleacetic acid were chosen to examine whether intact deciduous and coniferous tissues from spruce, hemlock fir, spinage, barley and sorrel or isolated mesophyll protoplasts from barley and closing cell preparations from Valerianella locusta are affected by sulphur dioxide in terms of changes in the concentration, transportation and distribution of such plant hormones. The distribution of phytohormones like ABA and IAA over the individual cell compartments is determined by the different pH gradients of the latter. Owing to their acidity these hormones are accumulated in alkaline cell inclusion bodies like chloroplasts and cytosol. Potentially acid air pollutants like SO{sub 2} and NO{sub x} lead to acidification of previously alkaline cell compartments, due to which fact the cellular pH gradients are reduced. This, in turn, gives rise to a redistribution of phytohormones to the effect that certain target cells such as closing cells of leaves or meristem cells come under the influence of altered hormone concentrations and compositions. This is bound to affect the processes controlling the development, growth and stress behaviour of plants. (orig.) With 55 refs., 2 tabs., 16 figs.

  3. Diversity of Mat-Forming Fungi in Relation to Soil Properties, Disturbance, and Forest Ecotype at Crater Lake National Park, Oregon, USA

    James M. Trappe


    Full Text Available In forest ecosystems, fungal mats are functionally important in nutrient and water uptake in litter and wood decomposition processes, in carbon resource allocation, soil weathering and in cycling of soil resources. Fungal mats can occur abundantly in forests and are widely distributed globally. We sampled ponderosa pine/white fir and mountain hemlock/noble fir communities at Crater Lake National Park for mat-forming soil fungi. Fungus collections were identified by DNA sequencing. Thirty-eight mat-forming genotypes were identified; members of the five most common genera (Gautieria, Lepiota, Piloderma, Ramaria, and Rhizopogon comprised 67% of all collections. The mycorrhizal genera Alpova and Lactarius are newly identified as ectomycorrhizal mat-forming taxa, as are the saprotrophic genera Flavoscypha, Gastropila, Lepiota and Xenasmatella. Twelve typical mat forms are illustrated, representing both ectomycorrhizal and saprotrophic fungi that were found. Abundance of fungal mats was correlated with higher soil carbon to nitrogen ratios, fine woody debris and needle litter mass in both forest ecotypes. Definitions of fungal mats are discussed, along with some of the challenges in defining what comprises a fungal “mat”.

  4. The Year Without a Ski Season: An Analysis of the Winter of 2015 for Three Ski Resorts in Western Canada Using Historical and Simulation Model Forecasted Climate Data

    Pidwirny, M. J.; Goode, J. D.; Pedersen, S.


    The winter of 2015 will go down as "the year without a ski season" for many ski resorts located close to the west coast of Canada and the USA. During this winter season, a large area of the eastern North Pacific Ocean had extremely high sea surface temperatures. These high sea surface temperatures influenced weather patterns on the west coast of North America producing very mild temperatures inland. Further, in alpine environments precipitation that normally arrives in the form of snow instead fell as rain. This research examines the climate characteristics of the winter of 2015 in greater detail for three ski resorts in British Columbia, Canada: Mount Washington, Cypress Mountain and Hemlock Valley. For these resorts, historical (1901 to 2013) and IPCC AR5 climate model forecasted climate data (RCP8.5 for 2025, 2055, and 2085) was generated for the variable winter degree days climate database ClimateBC. A value for winter degree days climate data at nearby meteorological stations for comparative analysis. For all three resorts, the winter of 2015 proved to be warmer than any individual year in the period 1901 to 2013. Interpolations involving the multi-model ensemble forecast means suggest that the climate associated with winter of 2015 will become the average normal for these resorts in only 35 to 45 years under the RCP8.5 emission scenario.

  5. Cytochemical localization of cellulases in decayed and nondecayed wood

    Murmanis, L.; Highley, T.L.; Palmer, J.G.


    Sawdust from undecayed western hemlock wood and from wood previously decayed by the brown-rot fungus Poria placenta or by the white-rot fungus Ganoderma applanatum was incubated with commercial cellulase from Trichoderma viride. Samples were treated cytochemically to locate cellulase activity and examined by TEM. Results showed that cellulase degraded undecayed wood extensively, with the attack starting on the outer border of a cell wall and progressing inside. Wood decayed by P. placenta, with or without cellulase incubation, and treated by the cytochemical test showed uniform distribution of electron dense particles throughout the cell walls. In wood decayed by G. applanatum, cellulase degradation was similar to that in undecayed wood. From measurements of particle diameter it is suggested that electron dense particles are cellulase. It is concluded that brown-rot and white-rot fungi have different effects on the microstructure of wood. The brown-rot fungus appears to open the wood microstructure so that cellulase can diffuse throughout the degraded tracheid wall.

  6. Kevorkian v. Thompson.


    Court Decision: 947 Federal Supplement 1152; 1997 Jan 6 (date of decision). The United States District Court of the Eastern District of Michigan held that a mentally competent patient who is terminally ill or intractably suffering does not have a liberty interest in assisted suicide under the due process clause of the Fourth Amendment and is not denied equal protection under the Fourteenth Amendment. Plaintiff Jack Kevorkian is a physician who advocates the right to die and assisted patients to commit suicide. Plaintiff Janet Good, the former president of the Michigan Hemlock Society, suffers from terminal pancreatic cancer. The plaintiffs claimed that Michigan's statute prohibiting physician-assisted suicide is unconstitutional. The District Court held that there is no cognizable constitutional right to assisted suicide because the right to suicide or assisted suicide is not deeply rooted in the nation's history and traditions, and because the statute does not infringe on any fundamental right or liberty. The court noted a difference between the withdrawal of life support and acts to hasten death by assisted suicide. The court also held that the law against physician-assisted suicide furthered legitimate state interests in denying to physicians "the role of killers of their patients," in regulating circumstances under which life may be ended, and in protecting the vulnerable but viable from "self-interested importuning of third parties."

  7. Spring and Autumn Phenological Variability across Environmental Gradients of Great Smoky Mountains National Park, USA

    Steven P. Norman


    Full Text Available Mountainous regions experience complex phenological behavior along climatic, vegetational and topographic gradients. In this paper, we use a MODIS time series of the Normalized Difference Vegetation Index (NDVI to understand the causes of variations in spring and autumn timing from 2000 to 2015, for a landscape renowned for its biological diversity. By filtering for cover type, topography and disturbance history, we achieved an improved understanding of the effects of seasonal weather variation on land surface phenology (LSP. Elevational effects were greatest in spring and were more important than site moisture effects. The spring and autumn NDVI of deciduous forests were found to increase in response to antecedent warm temperatures, with evidence of possible cross-seasonal lag effects, including possible accelerated green-up after cold Januarys and early brown-down following warm springs. Areas that were disturbed by the hemlock woolly adelgid and a severe tornado showed a weaker sensitivity to cross-year temperature and precipitation variation, while low severity wildland fire had no discernable effect. Use of ancillary datasets to filter for disturbance and vegetation type improves our understanding of vegetation’s phenological responsiveness to climate dynamics across complex environmental gradients.

  8. Potyviruses, novel and known, in cultivated and wild species of the family Apiaceae in Australia.

    Moran, J; van Rijswijk, B; Traicevski, V; Kitajima, E W; Mackenzie, A M; Gibbs, A J


    Three potyviruses were identified by gene sequencing and found to be widespread in species of Apiaceae in Australia. Only celery mosaic virus was found in celery crops and in one of 180 specimens of feral carrot ( Daucus carota). Another related but distinct novel potyvirus, carrot virus Y, was the only virus found in carrot crops and all except one feral carrot. A more distantly related novel potyvirus, apium virus Y, was found in plants of sea celery ( Apium prostratum), cultivated parsley ( Petroselinum crispum) and the immigrant weed species poison hemlock ( Conium maculatum). These three potyviruses, together with celery yellow mosaic virus of South America and the closely related carrot thin leaf virus and carrot virus B of North America, form a distinct subgenus of the Potyviridae most closely related to turnip mosaic virus and two potyviruses of yam; yam mosaic virus from the Ivory Coast and Japanese yam mosaic virus. Celery mosaic and carrot virus Y are probably recent migrants to Australia, but apium virus Y may have been endemic longer. In ELISA tests using polyclonal antibodies against virions of celery mosaic virus, some isolates of carrot virus Y were indistinguishable from celery mosaic virus, whereas others gave smaller absorbancy values, and those of apium virus Y did not react. This study shows the value of virus identification based on gene sequencing for planning control measures.

  9. [Neurological syndromes linked with the intake of plants and fungi containing a toxic component (I). Neurotoxic syndromes caused by the ingestion of plants, seeds and fruits].

    Carod-Artal, F J

    A wide range of plants, seeds and fruits used for nutritional and medicinal purposes can give rise to neurotoxic symptoms. We review the neurological pathology associated with the acute or chronic consumption of plants, seeds and fruits in human beings and in animals. Of the plants that can trigger acute neurotoxic syndromes in humans, some of the most notable include Mandragora officinalis, Datura stramonium, Conium maculatum (hemlock), Coriaria myrtifolia (redoul), Ricinus communis, Gloriosa superba, Catharanthus roseus, Karwinskia humboldtiana and Podophyllum pelatum. We also survey different neurological syndromes linked with the ingestion of vegetable foodstuffs that are rich in cyanogenic glycosides, Jamaican vomiting sickness caused by Blighia sapida, Parkinson dementia ALS of Guam island and exposition to Cycas circinalis, Guadeloupean parkinsonism and exposition to Annonaceae, konzo caused by ingestion of wild manioc and neurolathyrism from ingestion of Lathyrus sativus, the last two being models of motor neurone disease. Locoism is a chronic disease that develops in livestock feeding on plants belonging to Astragalus and Oxytropis sp., Sida carpinifolia and Ipomea carnea, which are rich in swainsonine, a toxin that inhibits the enzyme alpha mannosidase and induces a cerebellar syndrome. The ingestion of neurotoxic seeds, fruits and plants included in the diet and acute poisoning by certain plants can give rise to different neurological syndromes, some of which are irreversible.

  10. Ultrasound studies of the effects of certain poisonous plants on uterine function and fetal development in livestock.

    Bunch, T D; Panter, K E; James, L F


    Ingestion of locoweed (Astragalus spp. and Oxytropis spp.) by pregnant livestock may result in fetal malformations, delayed placentation, reduced placental and uterine vascular development, hydrops amnii, hydrops allantois, abnormal cotyledonary development, interruption of fetal fluid balance, and abortion. Ultrasonography of pregnant sheep fed locoweed demonstrated that abortion was first preceded by changes in fetal heart rate and strength of contraction and structural changes of the cotyledons, followed by increased accumulation of fetal fluid within the placental membranes and death of the fetus. During pregnancy the toxic agent in locoweed (swainsonine) apparently passes through the placental barrier to the fetus and during lactation through the milk to the neonate. Poison-hemlock (Conium maculatum), wild tree tobacco (Nicotiana glauca), and lunara lupine (Lupinus formosus) all contain piperidine alkaloids and induce fetal malformations, including multiple congenital contractures and cleft palate in livestock. Ultrasonography studies of pregnant sheep and goats gavaged with these plants during 30 to 60 d of gestation suggests that the primary cause of multiple congenital contractures and cleft palate is the degree and the duration of the alkaloid-induced fetal immobilization.

  11. Fetal muscle-type nicotinic acetylcholine receptor activation in TE-671 cells and inhibition of fetal movement in a day 40 pregnant goat model by optical isomers of the piperidine alkaloid coniine.

    Green, Benedict T; Lee, Stephen T; Welch, Kevin D; Pfister, James A; Panter, Kip E


    Coniine is an optically active toxic piperidine alkaloid and nicotinic acetylcholine receptor (nAChR) agonist found in poison hemlock (Conium maculatum L.). Coniine teratogenicity is hypothesized to be attributable to the binding, activation, and prolonged desensitization of fetal muscle-type nAChR, which results in the complete inhibition of fetal movement. However, pharmacological evidence of coniine actions at fetal muscle-type nAChR is lacking. The present study compared (-)-coniine, (+)-coniine, and nicotine for the ability to inhibit fetal movement in a day 40 pregnant goat model and in TE-671 cells that express fetal muscle-type nAChR. Furthermore, α-conotoxins (CTx) EI and GI were used to antagonize the actions of (+)- and (-)-coniine in TE-671 cells. (-)-Coniine was more effective at eliciting electrical changes in TE-671 cells and inhibiting fetal movement than was (+)-coniine, suggesting stereoselectivity by the receptor. The pyridine alkaloid nicotine did not inhibit fetal movement in a day 40 pregnant goat model, suggesting agonist specificity for the inhibition of fetal movement. Low concentrations of both CTxs potentiated the TE-671 cell response and higher concentrations of CTx EI, and GI antagonized the actions of both coniine enantiomers demonstrating concentration-dependent coagonism and selective antagonism. These results provide pharmacological evidence that the piperidine alkaloid coniine is acting at fetal muscle-type nAChR in a concentration-dependent manner.

  12. Stereoselective potencies and relative toxicities of γ-coniceine and N-methylconiine enantiomers.

    Lee, Stephen T; Green, Benedict T; Welch, Kevin D; Jordan, Glenn T; Zhang, Qian; Panter, Kip E; Hughes, David; Chang, Cheng-Wei Tom; Pfister, James A; Gardner, Dale R


    γ-Coniceine, coniine, and N-methylconiine are toxic alkaloids present in poison hemlock (Conium maculatum). We previously reported the comparison of the relative potencies of (+)- and (-)-coniine enantiomers. In this study, we synthesized γ-coniceine and the enantiomers of N-methylconiine and determined the biological activity of γ-coniceine and each of the N-methylconiine enantiomers in vitro and in vivo. The relative potencies of these piperidine alkaloids on cells expressing human fetal muscle-type nicotinic acetylcholine receptors had the rank order of γ-coniceine > (-)-N-methylconiine > (±)-N-methylconiine > (+)-N-methylconiine. The relative lethalities of γ-coniceine and (-)-, (±)-, and (+)-N-methylconiine in vivo using a mouse bioassay were 4.4, 16.1, 17.8, and 19.2 mg/kg, respectively. The results from this study suggest γ-coniceine is a more potent agonist than the enantiomers of N-methylconiine and that there is a stereoselective difference in the in vitro potencies of the enantiomers of N-methylconiine that correlates with the relative toxicities of the enantiomers in vivo.

  13. The effect of coniine on presynaptic nicotinic receptors.

    Erkent, Ulkem; Iskit, Alper B; Onur, Rustu; Ilhan, Mustafa


    Toxicity of coniine, an alkaloid of Conium maculatum (poison hemlock), is manifested by characteristic nicotinic clinical signs including excitement, depression, hypermetria, seizures, opisthotonos via postsynaptic nicotinic receptors. There is limited knowledge about the role of presynaptic nicotinic receptors on the pharmacological and toxicological effects of coniine in the literature. The present study was undertaken to evaluate the possible role of presynaptic nicotinic receptors on the pharmacological and toxicological effects of coniine. For this purpose, the rat anococcygeus muscle and guinea-pig atria were used in vitro. Nicotine (100 μM) elicited a biphasic response composed of a relaxation followed by contraction through the activation of nitrergic and noradrenergic nerve terminals in the phenylephrine-contracted rat anococcygeus muscle. Coniine inhibited both the nitrergic and noradrenergic response in the muscle (-logIC(50) = 3.79 ± 0.11 and -logIC(50) = 4.57 ± 0.12 M, respectively). The effect of coniine on nicotinic receptor-mediated noradrenergic transmission was also evaluated in the guinea-pig atrium (-logIC(50) = 4.47 ± 0.12 M) and did not differ from the -logIC(50) value obtained in the rat anococcygeus muscle. This study demonstrated that coniine exerts inhibitory effects on nicotinic receptor-mediated nitrergic and noradrenergic transmitter response.

  14. Alternative Carrier Solvents for Pigments Extracted from Spalting Fungi

    Lauren Pittis


    Full Text Available The use of both naturally occurring and synthetic pigmented wood has been prevalent in woodcraft for centuries. Modern manifestations generally involve either woodworkers’ aniline dyes, or pigments derived from a special class of fungi known as spalting fungi. While fungal pigments are more renewable than anilines and pose less of an environmental risk, the carrier required for these pigments—dichloromethane (DCM—is both problematic for humans and tends to only deposit the pigments on the surface of wood instead of evenly within the material. Internal coloration of wood is key to adoption of a pigmenting system by woodworkers. To address this issue, five solvents that had moderate solubility with the pigments extracted from Chlorociboria aeruginosa and Scytalidium cuboideum were identified, in the hopes that a reduction in solubility would result in a greater amount of the pigment deposited inside the wood. Of the tested solvents, acetonitrile was found to produce the highest internal color in ash, Douglas-fir, madrone, mountain hemlock, Port-Orford cedar, Pacific silver fir, red alder and sugar maple. While these carrier solvents are not ideal for extracting the pigments from the fungi, acetonitrile in particular does appear to allow for more pigment to be deposited within wood. The use of acetonitrile over DCM offers new opportunities for possible industrial spalting applications, in which larger pieces of wood could be uniformly pigmented and sold to the end user in larger quantities than are currently available with spalted wood.

  15. Aerospace Toxicology and Microbiology

    James, John T.; Parmet, A. J.; Pierson, Duane L.


    Toxicology dates to the very earliest history of humanity with various poisons and venom being recognized as a method of hunting or waging war with the earliest documentation in the Evers papyrus (circa 1500 BCE). The Greeks identified specific poisons such as hemlock, a method of state execution, and the Greek word toxos (arrow) became the root of our modern science. The first scientific approach to the understanding of poisons and toxicology was the work during the late middle ages of Paracelsus. He formulated what were then revolutionary views that a specific toxic agent or "toxicon" caused specific dose-related effects. His principles have established the basis of modern pharmacology and toxicology. In 1700, Bernardo Ramazzini published the book De Morbis Artificum Diatriba (The Diseases of Workers) describing specific illnesses associated with certain labor, particularly metal workers exposed to mercury, lead, arsenic, and rock dust. Modern toxicology dates from development of the modern industrial chemical processes, the earliest involving an analytical method for arsenic by Marsh in 1836. Industrial organic chemicals were synthesized in the late 1800 s along with anesthetics and disinfectants. In 1908, Hamilton began the long study of occupational toxicology issues, and by WW I the scientific use of toxicants saw Haber creating war gases and defining time-dosage relationships that are used even today.

  16. An appraisal of the classic forest succession paradigm with the shade tolerance index.

    Lienard, Jean; Florescu, Ionut; Strigul, Nikolay


    In this paper we revisit the classic theory of forest succession that relates shade tolerance and species replacement and assess its validity to understand patch-mosaic patterns of forested ecosystems of the USA. We introduce a macroscopic parameter called the "shade tolerance index" and compare it to the classic continuum index in southern Wisconsin forests. We exemplify shade tolerance driven succession in White Pine-Eastern Hemlock forests using computer simulations and analyzing approximated chronosequence data from the USDA FIA forest inventory. We describe this parameter across the last 50 years in the ecoregions of mainland USA, and demonstrate that it does not correlate with the usual macroscopic characteristics of stand age, biomass, basal area, and biodiversity measures. We characterize the dynamics of shade tolerance index using transition matrices and delimit geographical areas based on the relevance of shade tolerance to explain forest succession. We conclude that shade tolerance driven succession is linked to climatic variables and can be considered as a primary driving factor of forest dynamics mostly in central-north and northeastern areas in the USA. Overall, the shade tolerance index constitutes a new quantitative approach that can be used to understand and predict succession of forested ecosystems and biogeographic patterns.

  17. An appraisal of the classic forest succession paradigm with the shade tolerance index.

    Jean Lienard

    Full Text Available In this paper we revisit the classic theory of forest succession that relates shade tolerance and species replacement and assess its validity to understand patch-mosaic patterns of forested ecosystems of the USA. We introduce a macroscopic parameter called the "shade tolerance index" and compare it to the classic continuum index in southern Wisconsin forests. We exemplify shade tolerance driven succession in White Pine-Eastern Hemlock forests using computer simulations and analyzing approximated chronosequence data from the USDA FIA forest inventory. We describe this parameter across the last 50 years in the ecoregions of mainland USA, and demonstrate that it does not correlate with the usual macroscopic characteristics of stand age, biomass, basal area, and biodiversity measures. We characterize the dynamics of shade tolerance index using transition matrices and delimit geographical areas based on the relevance of shade tolerance to explain forest succession. We conclude that shade tolerance driven succession is linked to climatic variables and can be considered as a primary driving factor of forest dynamics mostly in central-north and northeastern areas in the USA. Overall, the shade tolerance index constitutes a new quantitative approach that can be used to understand and predict succession of forested ecosystems and biogeographic patterns.

  18. Ozone for removal of acute toxicity from wood yard run-off

    Zenaitis, G.; Duff, S.J.B. [British Columbia Univ., Vancouver, BC (Canada)


    Environmental regulators are increasingly concerned about stormwater run-off from wood handling facilities. The authors discussed the use of ozone in the development of treatment methods to deal with toxic run-off from wood yard and dryland sorts. Two sawmills were selected, one located on the north coast of British Columbia (mill A), while mill B was located on Vancouver Island. Balsam fir, western hemlock, Sitka spruce,and western red cedar were processed at mill A, while mill B processed a wide range of species, where it is not unusual to undergo a complete change in a 24-hour period. Water samples from the run-off were obtained at the two mills, which were then kept in 20-L plastic containers stored in the dark at 4 Celsius upon receipt. Ozone was used to treat the centrifuged samples in a lab-scale reactor. Results indicated an 80-90 per cent reduction in toxicity through ozonation, a 90-95 per cent reduction for tannin and lignin (TL), and a 95-100 per cent reduction in dehydroabietic acid (DHA). Neutral solutions tended to improve slightly the toxicity and DHA removal when compared to acidic solutions. The influence of mass transfer effects, the influence of non-toxic dissolved organics, and the potential gains through the use of catalysts are still being investigated. 25 refs., 2 tabs., 6 figs.

  19. Building Resilience into Sitka Spruce (Picea sitchensis (Bong. Carr. Forests in Scotland in Response to the Threat of Climate Change

    Andrew D. Cameron


    Full Text Available It is expected that a warming climate will have an impact on the future productivity of European spruce forests. In Scotland, Sitka spruce (Picea sitchensis (Bong. Carr. dominates the commercial forestry sector and there is growing pressure to develop alternative management strategies to limit potential economic losses through climate change. This review considers management options to increase the resilience of Sitka spruce dominated forests in Scotland. Given the considerable uncertainty over the potential long-term impacts of climate change, it is recommended that Sitka spruce should continue to be planted where it already grows well. However, new planting and restocking should be established in mixtures where silviculturally practicable, even if no-thin regimes are adopted, to spread future risks of damage. Three potentially compatible species with Sitka spruce are western hemlock (Tsuga heterophylla (Raf. Sarg., grand fir (Abies grandis (Lamb. Lindl. and Douglas fir (Pseudotsuga menziesii (Mirb. Franco and all form natural mixtures in its native range in North America. The predicted windier climate will require a range of management inputs, such as early cutting of extraction racks and early selective thinning, to improve stability. The potential to improve resilience to particularly abiotic damage through transforming even-aged stands into irregular structures and limiting the overall size of the growing stock is discussed.

  20. Dreams, Nightmares and Haunted Houses: Televisual Horror as Domestic Imaginary

    Ruth Griffin


    Full Text Available It has been widely proclaimed that U.S.Television is currently experiencing a Golden Age with horror at its vanguard, in part enabled by technological innovations that have seen audiences engage with TV in ever more diverse ways, enabled by the advent of Smart T.V. Meanwhile, television has historically positioned itself as a humble and domesticated medium and yet its increasingly sophisticated channels penetrate into the very heart of the contemporary home. With this in mind, I view Suburban Gothic TV series such as American Horror Story (2011 and Hemlock Grove (2013 through the lens of psychoanalytic concepts such as The Uncanny, considering the extent to which such dramas invoke the dark side of the domestic imaginary which haunts that most cherished of spaces, the home. Why does Gothic Horror continue to engage the imaginations of the contemporary home’s technologically orientated inhabitants? And how has technology helped to drive the resurgence of a genre so firmly rooted in a historical-literary form? These are just some of the questions that this article explores.

  1. Short communication: Dairy bedding type affects survival of Prototheca in vitro.

    Adhikari, N; Bonaiuto, H E; Lichtenwalner, A B


    Protothecae are algal pathogens, capable of causing bovine mastitis, that are unresponsive to treatment; they are believed to have an environmental reservoir. The role of bedding management in control of protothecal mastitis has not been studied. The purpose of this study was to evaluate the growth of either environmental or mastitis-associated Prototheca genotypes in dairy bedding materials that are commonly used in Maine. Prototheca zopfii genotypes 1 and 2 (gt1 and gt2) were inoculated into sterile broth only (control ), kiln-dried spruce shavings, "green" hemlock sawdust, sand, or processed manure-pack beddings with broth, and incubated for 2 d. Fifty microliters of each isolate was then cultured onto plates and the resulting colonies counted at 24 and 48 h postinoculation. Shavings were associated with significantly less total Prototheca growth than other bedding types. Growth of P. zopfii gt1 was significantly higher than that of gt2 in the manure-pack bedding material. Spruce shavings, compared with manure, sand, or sawdust, may be a good bedding type to prevent growth of Prototheca. Based on these in vitro findings, bedding type may affect Prototheca infection of cattle in vivo. Copyright © 2013 American Dairy Science Association. Published by Elsevier Inc. All rights reserved.

  2. Regional and Large-Scale Climate Influences on Tree-Ring Reconstructed Null Zone Position in San Francisco Bay

    Stahle, D.; Griffin, D.; Cleaveland, M.; Fye, F.; Meko, D.; Cayan, D.; Dettinger, M.; Redmond, K.


    A new network of 36 moisture sensitive tree-ring chronologies has been developed in and near the drainage basins of the Sacramento and San Joaquin Rivers. The network is based entirely on blue oak (Quercus douglasii), which is a California endemic found from the lower forest border up into the mixed conifer zone in the Coast Ranges, Sierra Nevada, and Cascades. These blue oak tree-ring chronologies are highly correlated with winter-spring precipitation totals, Sacramento and San Joaquin streamflow, and with seasonal variations in salinity and null zone position in San Francisco Bay. Null zone is the non-tidal bottom water location where density-driven salinity and river-driven freshwater currents balance (zero flow). It is the area of highest turbidity, water residence time, sediment accumulation, and net primary productivity in the estuary. Null zone position is measured by the distance from the Golden Gate of the 2 per mil bottom water isohaline and is primarily controlled by discharge from the Sacramento and San Joaquin Rivers (and ultimately by winter-spring precipitation). The location of the null zone is an estuarine habitat indicator, a policy variable used for ecosystem management, and can have a major impact on biological resources in the San Francisco estuary. Precipitation-sensitive blue oak chronologies can be used to estimate null zone position based on the strong biogeophysical interaction among terrestrial, aquatic, and estuarine ecosystems, orchestrated by precipitation. The null zone reconstruction is 626-years long and provides a unique long term perspective on the interannual to decadal variability of this important estuarine habitat indicator. Consecutive two-year droughts (or longer) allow the null zone to shrink into the confined upper reaches of Suisun Bay, causing a dramatic reduction in phytoplankton production and favoring colonization of the estuary by marine biota. The reconstruction indicates an approximate 10 year recurrence interval

  3. Plants promote mating and dispersal of the human pathogenic fungus Cryptococcus.

    Deborah J Springer

    Full Text Available Infections due to Cryptococcus are a leading cause of fungal infections worldwide and are acquired as a result of environmental exposure to desiccated yeast or spores. The ability of Cryptococcus to grow, mate, and produce infectious propagules in association with plants is important for the maintenance of the genetic diversity and virulence factors important for infection of animals and humans. In the Western United States and Canada, Cryptococcus has been associated with conifers and tree species other than Eucalyptus; however, to date Cryptococcus has only been studied on live Arabidopsis thaliana, Eucalyptus sp., and Terminalia catappa (almond seedlings. Previous research has demonstrated the ability of Cryptococcus to colonize live plants, leaves, and vasculature. We investigated the ability of Cryptococcus to grow on live seedlings of the angiosperms, A. thaliana, Eucalyptus camaldulensis, Colophospermum mopane, and the gymnosperms, Pseudotsuga menziesii (Douglas fir, and Tsuga heterophylla (Western hemlock. We observed a broad-range ability of Cryptococcus to colonize both traditional infection models as well as newly tested conifer species. Furthermore, C. neoformans, C. deneoformans, C. gattii (VGI, C. deuterogattii (VGII and C. bacillisporus (VGIII were able to colonize live plant leaves and needles but also undergo filamentation and mating on agar seeded with plant materials or in saprobic association with dead plant materials. The ability of Cryptococcus to grow and undergo filamentation and reproduction in saprobic association with both angiosperms and gymnosperms highlights an important role of plant debris in the sexual cycle and exposure to infectious propagules. This study highlights the broad importance of plants (and plant debris as the ecological niche and reservoirs of infectious propagules of Cryptococcus in the environment.

  4. Elevated Annual Runoff Ratios in Pacific Northwest Catchments Impacted by Epidemic Foliage Disease of Douglas-fir

    Bladon, K. D.; Bywater-Reyes, S.; LeBoldus, J. M.; Segura, C.; Ritokova, G.; Shaw, D. C.


    Catchments in the Western United States are undergoing unprecedented levels of tree die-off and/or reduced vigor due to increased severity of wildfire, drought, insect outbreaks, and disease. In the U.S. Pacific Northwest, Swiss needle cast (SNC) is the most damaging foliar disease of Douglas-fir (Pseudotsuga menziesii), physically obstructing stomata and preventing CO2 uptake and transpiration. A recent analysis in coastal Oregon indicated a substantial increase in area affected by the disease, from 530.5 km2 in 1996 to 2,387.1 km2 in 2015. Deforestation or reduced tree vigor can have profound impacts on catchment hydrology, in theory, producing increased streamflow due to reduced interception and transpiration. However, these increases have not always been detectable as impacts also depend on factors such as climate and vegetation composition. Moreover, press disturbances, such as insect outbreaks or disease, often do not result in complete removal of understorey or canopy vegetation. We analyzed trends in annual runoff ratios (quotient of discharge divided by precipitation) from 1990-2015 in 12 catchments (183-1,744 km2) in western Oregon. In general, runoff ratios increased by 10-27% in catchments with a total area of SNC >10%, with the most substantial runoff increases in catchments with SNC impacting >25% of the area. Interestingly, the most severely impacted catchment ( 90.5% SNC) showed a decrease in runoff. This is consistent with a potential compensatory response from understory western hemlock (Tsuga heterophylla) trees, a phenomenon observed in the most severely impacted sites. Findings from this study are important for assessing the impacts of biotic forest disturbances on water supply and aquatic ecosystem health.

  5. Anesthesia and Pain Relief in the History of Islamic Medicine.

    Alembizar, Faranak; Hosseinkhani, Ayda; Salehi, Alireza


    Since diseases and surgeries could be very painful, the annihilation of pain has been the most important goal of physicians. The history of Iranian-Islamic medicine includes distinguished physicians that attempted to find different methods of anesthesia. This research aims at reviewing approaches for anesthesia throughout the history of the Iranian-Islamic medicine, in order to identify a variety of drugs used during that period. In this research, the information was mainly collected from medical history, traditional literature and various search engines (e.g. Google Scholar, PubMed, Medline, Scopus, SIDS and NoorMags). The search keywords were Anesthetic, Tbnj (sedation), Tnvym (sedative), and Hypnotic. Finally, a detailed analytical study was performed on all notes and the results were presented. Mohammad Ibn-Zakaria Al-Razi (known to the Western world as Razes) in the 10th century was the first physician who used general inhalation for anesthesia in surgeries. Drugs used to relieve pain and anesthesia can be divided into two categories: (i) single drug and (ii) compound drugs. Usually, these are consumed by eating, drinking, inhalation, or as topical. Drugs such as Hemlock, Mandrake, Henbane, Hyocyamus, Mandragora, Loiseuria, Opium Poppy, and Black Nightshade were used. Beyond these herbs, Aghili (18 th century) in his book "Makhzan al-adviyah" also explained the topical application of ice for pain management. The choice for the type of medication and its form of consumption is commensurate to pain and the speed by which the drug has an effect. Anesthesia was usually done in two ways: (i) using a substance called "Mokhader" which was consumed via the mouth or nose, and (ii) "Tnvym" which means putting a patient to sleep to block the sensation of pain. Typically, anesthesia methods and drug recipes were kept as secret to prevent misuse and abuse by unauthorized people. Based on our study, Islamic physicians proposed inspiring methods in using drugs for anesthesia

  6. Wind River Watershed Restoration Project; Underwood Conservation District, Annual Report 2002-2003.

    White, Jim


    The goal of the Wind River project is to preserve, protect and restore Wind River steelhead. In March, 1998, the National Marine Fisheries Service listed the steelhead of the lower Columbia as 'threatened' under the Endangered Species Act. In 1997, the Washington Department of Fish and Wildlife rated the status of the Wind River summer run steelhead as critical. Due to the status of this stock, the Wind River summer steelhead have the highest priority for recovery and restoration in the state of Washington's Lower Columbia Steelhead Conservation Initiative. The Wind River Project includes four cooperating agencies. Those are the Underwood Conservation District (UCD), United States Geological Service (USGS), US Forest Service (USFS), and Washington State Department of Fish & Wildlife (WDFW). Tasks include monitoring steelhead populations (USGS and WDFW), Coordinating a Watershed Committee and Technical Advisory Group (UCD), evaluating physical habitat conditions (USFS and UCD), assessing watershed health (all), reducing road sediments sources (USFS), rehabilitating riparian corridors, floodplains, and channel geometry (UCD, USFS), evaluate removal of Hemlock Dam (USFS), and promote local watershed stewardship (UCD, USFS). UCD's major efforts have included coordination of the Wind River Watershed Committee and Technical Advisory Committee (TAC), water temperature and water chemistry monitoring, riparian habitat improvement projects, and educational activities. Our coordination work enables the local Watershed Committee and TAC to function and provide essential input to Agencies, and our habitat improvement work focuses on riparian revegetation. Water chemistry and temperature data collection provide information for monitoring watershed conditions and fish habitat, and are comparable with data gathered in previous years. Water chemistry information collected on Trout Creek should, with 2 years data, determine whether pH levels make conditions

  7. Taking the pulse of mountains: Ecosystem responses to climatic variability

    Fagre, Daniel B.; Peterson, David L.; Hessl, Amy E.


    An integrated program of ecosystem modeling and field studies in the mountains of the Pacific Northwest (U.S.A.) has quantified many of the ecological processes affected by climatic variability. Paleoecological and contemporary ecological data in forest ecosystems provided model parameterization and validation at broad spatial and temporal scales for tree growth, tree regeneration and treeline movement. For subalpine tree species, winter precipitation has a strong negative correlation with growth; this relationship is stronger at higher elevations and west-side sites (which have more precipitation). Temperature affects tree growth at some locations with respect to length of growing season (spring) and severity of drought at drier sites (summer). Furthermore, variable but predictable climate-growth relationships across elevation gradients suggest that tree species respond differently to climate at different locations, making a uniform response of these species to future climatic change unlikely. Multi-decadal variability in climate also affects ecosystem processes. Mountain hemlock growth at high-elevation sites is negatively correlated with winter snow depth and positively correlated with the winter Pacific Decadal Oscillation (PDO) index. At low elevations, the reverse is true. Glacier mass balance and fire severity are also linked to PDO. Rapid establishment of trees in subalpine ecosystems during this century is increasing forest cover and reducing meadow cover at many subalpine locations in the western U.S.A. and precipitation (snow depth) is a critical variable regulating conifer expansion. Lastly, modeling potential future ecosystem conditions suggests that increased climatic variability will result in increasing forest fire size and frequency, and reduced net primary productivity in drier, east-side forest ecosystems. As additional empirical data and modeling output become available, we will improve our ability to predict the effects of climatic change

  8. Evaluation of developmental toxicity of coniine to rats and rabbits.

    Forsyth, C S; Frank, A A


    Conium maculatum (poison hemlock, CM) is teratogenic in several domestic species, presumably due to its piperidine alkaloids, including coniine, which has been verified to be teratogenic in cattle. Coniine/CM teratogenicity culminates in production of arthrogryposis. The purpose of this study was to evaluate coniine-induced teratogenicity in two laboratory animal species, Sprague-Dawley rats and New Zealand white rabbits. Pregnant rats were given coniine (25 mg/kg body weight) by oral gavage at 8-hour intervals on gestation days 16-18. Pregnant rabbits were given coniine (40 mg/kg body weight) by oral gavage at 8-hour intervals on gestation days 20-24. Rats were killed on day 19 and rabbits on day 29. Fetuses were immediately removed, weighed, and examined for external abnormalities. Alternate fetuses were either stained for skeletal examinations with alizarin red-S or fixed in Bouin's solution for visceral examination. Symptoms of maternal intoxication due to coniine administration were observed in both the rat and the rabbit, and higher doses were uniformly lethal. Rabbits treated with coniine appeared to lose more weight and eat less than controls, but there was no statistically significant difference between groups. Fetal weights were significantly lower in coniine-exposed rat and rabbit fetuses indicating fetotoxicity. The only statistically significant treatment-related visceral or skeletal malformation was a reduction of cranial ossification of rabbit fetuses, probably related to maternal toxicity. Coniine-exposed rabbit litters tended to be affected by arthrogryposis (no bony deformities noted on skeletal exam) more than controls (2/6 vs. 0/9).

  9. Rapid in situ detection of alkaloids in plant tissue under ambient conditions using desorption electrospray ionization.

    Talaty, Nari; Takáts, Zoltán; Cooks, R Graham


    Desorption electrospray ionization (DESI) mass spectrometry is applied to the in situ detection of alkaloids in the tissue of poison hemlock (Conium maculatum), jimsonweed (Datura stramonium) and deadly nightshade (Atropa belladonna). The experiment is carried out by electrospraying micro-droplets of solvent onto native or freshly-cut plant tissue surfaces. No sample preparation is required and the mass spectra are recorded under ambient conditions, in times of a few seconds. The impact of the sprayed droplets on the surface produces gaseous ions from organic compounds originally present in the plant tissue. The effects of operating parameters, including the electrospray high voltage, heated capillary temperature, the solvent infusion rate and the carrier gas pressure on analytical performance are evaluated and optimized. Different types of plant material are analyzed including seeds, stems, leaves, roots and flowers. All the previously reported alkaloids have been detected in C. maculatum, while fifteen out of nineteen known alkaloids for D. stramonium and the principal alkaloids of A. belladonna were also identified. All identifications were confirmed by tandem mass spectrometry. Results obtained show similar mass spectra, number of alkaloids, and signal intensities to those obtained when extraction and separation processes are performed prior to mass spectrometric analysis. Evidence is provided that DESI ionization occurs by both a gas-phase ionization process and by a droplet pick-up mechanism. Quantitative precision of DESI is compared with conventional electrospray ionization mass spectrometry (after sample workup) and the RSD values for the same set of 25 dicotyledonous C. maculatum seeds (one half of each seed analyzed by ESI and the other by DESI) are 9.8% and 5.2%, respectively.

  10. Favorable fragmentation: river reservoirs can impede downstream expansion of riparian weeds.

    Rood, Stewart B; Braatne, Jeffrey H; Goater, Lori A


    River valleys represent biologically rich corridors characterized by natural disturbances that create moist and barren sites suitable for colonization by native riparian plants, and also by weeds. Dams and reservoirs interrupt the longitudinal corridors and we hypothesized that this could restrict downstream weed expansion. To consider this "reservoir impediment" hypothesis we assessed the occurrences and abundances of weeds along a 315-km river valley corridor that commenced with an unimpounded reach of the Snake River and extended through Brownlee, Oxbow, and Hells Canyon reservoirs and dams, and downstream along the Snake River. Sampling along 206 belt transects with 3610 quadrats revealed 16 noxious and four invasive weed species. Ten weeds were upland plants, with Canada thistle (Cirsium arvense) restricted to the upstream reaches, where field morning glory (Convolvulus arvensis) was also more common. In contrast, St. John's wort (Hypericum perforatum) was more abundant below the dams, and medusahead wildrye (Taeniatherum caput-medusae) occurred primarily along the reservoirs. All seven riparian species were abundant in the upstream zones but sparse or absent below the dams. This pattern was observed for the facultative riparian species, poison hemlock (Conium maculatum) and perennial pepperweed (Lepidium latifolium), the obligate riparian, yellow nut sedge (Cyperus esculentus), the invasive perennial, reed canary grass (Phalaris arundinacea), and three invasive riparian trees, Russian olive (Elaeagnus angustifolia), false indigo (Amorpha fruticosa), and tamarisk (Tamarix spp.). The hydrophyte purple loosestrife (Lythrum salicaria) was also restricted to the upstream zone. These longitudinal patterns indicate that the reservoirs have impeded the downstream expansion of riparian weeds, and this may especially result from the repetitive draw-down and refilling of Brownlee Reservoir that imposes a lethal combination of drought and flood stress. The dams and

  11. Comparison of nicotinic receptor binding and biotransformation of coniine in the rat and chick.

    Forsyth, C S; Speth, R C; Wecker, L; Galey, F D; Frank, A A


    Coniine, an alkaloid from Conium maculatum (poison hemlock), is a known teratogen in many domestic species with maternal ingestion resulting in arthrogryposis of the offspring. We have previously shown that rats are not susceptible and rabbits only weakly susceptible to coniine-induced arthrogryposis. However, the chick embryo does provide a reproducible laboratory animal model of coniine-induced teratogenesis. The reason for this cross-species variation is unknown. The purpose of this study was to evaluate coniine binding to nicotinic receptors and to measure coniine metabolism in vitro between susceptible and non-susceptible species. Using the chick model, neither the peripheral nicotinic receptor antagonist d-tubocurarine chloride nor the central nicotinic receptor antagonist trimethaphan camsylate blocked the teratogenesis or lethality of 1.5% coniine (50 microliters/egg). Trimethaphan camsylate enhanced coniine-induced lethality in a dose-dependent manner. Neither nicotinic receptor blocker prevented nicotine sulfate-induced malformations but d-tubocurarine chloride did block lethality in a dose-dependent manner. Competition by coniine for [125I]-alpha-bungarotoxin to nicotinic receptors isolated from adult rat diaphragm and chick thigh muscle and competition by coniine for [3H]-cytisine to receptors from rat and chick brain were used to assess coniine binding to nicotinic receptors. The IC50 for coniine in rat diaphragm was 314 microM while that for chick leg muscle was 70 microM. For neuronal nicotinic receptors, the IC50s of coniine for maternal rat brain, fetal rat brain, and chick brain were 1100 microM, 820 microM, and 270 microM, respectively. There were no differences in coniine biotransformation in vitro by microsomes from rat or chick livers. Differences in apparent affinity of coniine for nicotinic receptors or differences in the quantity of the nicotinic receptor between the rat and chick may explain, in part, the differences in susceptibility of

  12. Effect of coniine on the developing chick embryo.

    Forsyth, C S; Frank, A A; Watrous, B J; Bohn, A A


    Coniine, an alkaloid from Conium maculatum (poison hemlock), has been shown to be teratogenic in livestock. The major teratogenic outcome is arthrogryposis, presumably due to nicotinic receptor blockade. However, coniine has failed to produce arthrogryposis in rats or mice and is only weakly teratogenic in rabbits. The purpose of this study was to evaluate and compare the effects of coniine and nicotine in the developing chick. Concentrations of coniine and nicotine sulfate were 0.015%, 0.03%, 0.075%, 0.15%, 0.75%, 1.5%, 3%, and 6% and 1%, 5%, and 10%, respectively. Both compounds caused deformations and lethality in a dose-dependent manner. All concentrations of nicotine sulfate caused some lethality but a no effect level for coniine lethality was 0.75%. The deformations caused by both coniine and nicotine sulfate were excessive flexion or extension of one or more toes. No histopathological alterations or differences in bone formation were seen in the limbs or toes of any chicks from any group; however, extensive cranial hemorrhage occurred in all nicotine sulfate-treated chicks. There was a statistically significant (P < or = 0.01) decrease in movement in coniine and nicotine sulfate treated chicks as determined by ultrasound. Control chicks were in motion an average of 33.67% of the time, while coniine-treated chicks were only moving 8.95% of a 5-min interval, and no movement was observed for nicotine sulfate treated chicks. In summary, the chick embryo provides a reliable and simple experimental animal model of coniine-induced arthrogryposis. Data from this model support a mechanism involving nicotinic receptor blockade with subsequent decreased fetal movement.

  13. Pharmacologically active plant metabolites as survival strategy products.

    Attardo, C; Sartori, F


    The fact that plant organisms produce chemical substances that are able to positively or negatively interfere with the processes which regulate human life has been common knowledge since ancient times. One of the numerous possible examples in the infusion of Conium maculatum, better known as Hemlock, a plant belonging to the family umbelliferae, used by the ancient Egyptians to cure skin diseases. The current official pharmacopoeia includes various chemical substances produced by secondary plant metabolisms. For example, the immunosuppressive drugs used to prevent organ transplant rejection and the majority of antibiotics are metabolites produced by fungal organisms, pilocarpin, digitalis, strophantus, salicylic acid and curare are examples of plant organism metabolites. For this reason, there has been an increase in research into plants, based on information on their medicinal use in the areas where they grow. The study of plants in relation to local culture and traditions is known as "ethnobotany". Careful study of the behaviour of sick animals has also led to the discovery of medicinal plants. The study of this subject is known as "zoopharmacognosy". The aim of this article is to discuss the fact that "ad hoc" production of such chemical substances, defined as "secondary metabolites", is one of the modes in which plant organisms respond to unfavourable environmental stimuli, such as an attack by predatory phytophagous animals or an excessive number of plant individuals, even of the same species, in a terrain. In the latter case, the plant organisms produce toxic substances, called "allelopathic" which limit the growth of other individuals. "Secondary metabolites" are produced by metabolic systems that are shunts of the primary systems which, when required, may be activated from the beginning, or increased to the detriment of others. The study of the manner in which such substances are produced is the subject of a new branch of learning called "ecological

  14. The influence of variable snowpacks on habitat use by mountain caribou

    Trevor A. Kinley


    Full Text Available Mountain caribou (Rangifer tarandus caribou in southeastern British Columbia subsist for most of the winter on arboreal hair lichen, mostly Bryoria spp. Foraging occurs mainly in old subalpine fir (Abies lasiocarpa forests near treeline. Here, the lower limit of Bryoria in the canopy is dictated by snowpack depth because hair lichens die when buried in snow. Bryoria is often beyond the reach of caribou in early winter, prompting caribou to move downslope to where lichen occurs lower in the canopy and other foraging modes are possible. Snowpacks are normally deep enough by late winter that caribou can reach Bryoria where it is most abundant, at high elevations. Extending this to inter-annual comparisons, Bryoria should be less accessible during late winter of low-snow years following normal winters, or of normal to low-snow years after deep-snow winters. We hypothesized that when maximum snowpack in late winter is low relative to the deepest of the previous 5 years, mountain caribou will use lower elevations to facilitate foraging (“lichen-snow-caribou” or LSC hypothesis. We tested this with late-winter data from 13 subpopulations. In the dry climatic region generally and for minor snowfall differences in wet and very wet regions, caribou did not shift downslope or in fact were at higher elevations during relatively low-snow years, possibly reflecting the ease of locomotion. The LSC hypothesis was supported within wet and very wet regions when snowpacks were about 1 m or more lower than in recent years. Elevation declined by 300 m (median to 600 m (25th percentile for snowpack differences of at least 1.5 m. Greater use of lodgepole pine and western hemlock stands sometimes also occurred. Management strategies emphasizing subalpine fir stands near treeline should be re-examined to ensure protection of a broader range of winter habitats used by caribou under variable snowpack conditions.

  15. Broadening the ecological context of ungulate-ecosystem interactions: the importance of space, seasonality, and nitrogen.

    Murray, Bryan D; Webster, Christopher R; Bump, Joseph K


    Spatial heterogeneity of soil resources, particularly nitrogen availability, affects herbaceous-layer cover and diversity in temperate forest ecosystems. Current hypotheses predict that ungulate herbivores influence nitrogen availability at the stand scale, but how ungulates affect nitrogen availability at finer spatial scales that are relevant to the herb layer is less understood. We tested the hypothesis that ungulate exclusion reduces the spatial complexity of nitrogen availability at neighborhood scales (1-26 m) apart from mean stand scale effects. This outcome was expected due to a lack of ungulate nitrogenous waste deposition within exclosures and seasonally variable ungulate habitat use. To test this hypothesis we examined spatial patterning of ammonium and nitrate availability, herb-layer cover and diversity, and under-canopy solar radiation using geostatistical models. Our study sites included six stands of eastern hemlock (Tsuga canadensis) forest: three where white-tailed deer (Odocoileus virginianus) were excluded and three that were accessible to deer. Where deer were present, patch sizes of ammonium availability, cover, and diversity were smaller compared to deer exclosures, whereas mean site-level effects were not significant. Within deer exclosures cover and solar radiation were more similar in patch size than were cover and nitrogen availability. Our results suggest that browsing ungulates affect spatial patterns of herb-layer cover and diversity through the excretion of nitrogenous wastes in small, discrete patches. Ungulate-excreted nitrogen deposition and herbivory were concentrated in the dormant season, allowing herb-layer plants a greater opportunity to benefit from nitrogen additions. Therefore, the impact of ungulates on nitrogen cycling in forest ecosystems varies with spatial scale and the seasonal timing of ungulate impacts. In this way, ungulates may function as a seasonally dependent link between fine-scale and landscape

  16. Tradeoffs between chilling and forcing in satisfying dormancy requirements for Pacific Northwest tree species.

    Harrington, Constance A; Gould, Peter J


    Many temperate and boreal tree species have a chilling requirement, that is, they need to experience cold temperatures during fall and winter to burst bud normally in the spring. Results from trials with 11 Pacific Northwest tree species are consistent with the concept that plants can accumulate both chilling and forcing units simultaneously during the dormant season and they exhibit a tradeoff between amount of forcing and chilling. That is, the parallel model of chilling and forcing was effective in predicting budburst and well chilled plants require less forcing for bud burst than plants which have received less chilling. Genotypes differed in the shape of the possibility line which describes the quantitative tradeoff between chilling and forcing units. Plants which have an obligate chilling requirement (Douglas-fir, western hemlock, western larch, pines, and true firs) and received no or very low levels of chilling did not burst bud normally even with long photoperiods. Pacific madrone and western redcedar benefited from chilling in terms of requiring less forcing to promote bud burst but many plants burst bud normally without chilling. Equations predicting budburst were developed for each species in our trials for a portion of western North America under current climatic conditions and for 2080. Mean winter temperature was predicted to increase 3.2-5.5°C and this change resulted in earlier predicted budburst for Douglas-fir throughout much of our study area (up to 74 days earlier) but later budburst in some southern portions of its current range (up to 48 days later) as insufficient chilling is predicted to occur. Other species all had earlier predicted dates of budburst by 2080 than currently. Recent warming trends have resulted in earlier budburst for some woody plant species; however, the substantial winter warming predicted by some climate models will reduce future chilling in some locations such that budburst will not consistently occur earlier.

  17. Plant recolonization in the Himalaya from the southeastern Qinghai-Tibetan Plateau: Geographical isolation contributed to high population differentiation.

    Cun, Yu-Zhi; Wang, Xiao-Quan


    The Himalaya-Hengduan Mountains region (HHM) in the southern and southeastern Qinghai-Tibetan Plateau (QTP) is considered an important reservoir and a differentiation center for temperate and alpine plants in the Cenozoic. To reveal how plants responded to the Quaternary climatic oscillations in the QTP, the phylogeographical histories of a few subalpine and alpine plants have been investigated, but nearly all studies used only uniparentally inherited cytoplasmic DNA markers, and only a couple of them included sampling from the Himalaya. In this study, range-wide genetic variation of the Himalayan hemlock (Tsuga dumosa), an important forest species in the HHM, was surveyed using DNA markers from three genomes. All markers revealed genetic depauperation in the Himalaya and richness in the Hengduan Mountains populations. Surprisingly, population differentiation of this wind-pollinated conifer is very high in all three genomes, with few common and many private nuclear gene alleles. These results, together with fossil evidence, clearly indicate that T. dumosa recolonized the Himalaya from the Hengduan Mountains before the Last Glacial Maximum (LGM), accompanied with strong founder effects, and the influence of the earlier glaciations on demographic histories of the QTP plants could be much stronger than that of the LGM. The strong population differentiation in T. dumosa could be attributed to restricted gene flow caused by the complicated topography in the HHM that formed during the uplift of the QTP, and thus sheds lights on the importance of geographical isolation in the development of high plant species diversity in this biodiversity hotspot. Copyright 2010 Elsevier Inc. All rights reserved.

  18. Kootenai River fisheries investigations: rainbow and bull trout recruitment: annual progress report 1999; ANNUAL

    Walters, Jody P.; Downs, Christopher Charles


    Our 1999 objectives were to determine sources of rainbow trout Oncorhynchus mykiss and bull trout Salvelinus confluentus spawning and recruitment in the Idaho reach of the Kootenai River. We used a rotary-screw trap to capture juvenile trout to determine age at out-migration and to estimate total out-migration from the Boundary Creek drainage to the Kootenai River. The out-migrant estimate for March through August 1999 was 1,574 (95% C. I.= 825-3,283) juvenile rainbow trout. Most juveniles out-migrated at age-2 and age-3. No out-migrating bull trout were caught. Five of 17 rainbow trout radio-tagged in Idaho migrated upstream into Montana waters during the spawning season. Five bull trout originally radio-tagged in O'Brien Creek, Montana in early October moved downstream into Idaho and British Columbia by mid-October. Annual angler exploitation for the rainbow trout population upstream of Bonners Ferry, Idaho was estimated to be 58%. Multi-pass depletion estimates for index reaches of Caboose, Curley, and Debt creeks showed 0.20, 0.01, and 0.13 rainbow trout juveniles/m(sup 2), respectively. We estimated rainbow trout (180-415 mm TL) standing stock of 1.6 kg/ha for the Hemlock Bar reach (29.4 ha) of the Kootenai River, similar to the 1998 estimate. Recruitment of juvenile rainbow and bull trout from Idaho tributaries is not sufficient to be the sole source of subsequent older fish in the mainstem Kootenai River. These populations are at least partly dependent on recruitment from Montana waters. The low recruitment and high exploitation rate may be indicators of a rainbow trout population in danger of further decline

  19. Extrinsic and Intrinsic Responses to Environmental Change: Insights from Terrestrial Paleoecological Archives

    Seddon, A. W. R.; Mackay, A. W.


    Current understanding of ecological behaviour indicates that systems can experience sudden and abrupt changes in state, driven either by a large external change in environmental conditions (extrinsically forced), or the result of a set local feedbacks and site-specific interactions (intrinsically mediated responses). Responses mediated by intrinsic processes are notoriously diffi- cult to predict, they can occur as slow environmental variables gradually erode the resilience of the system eventually resulting in threshold transitions between alternative stable states. Finding ways to identify, model and predict such complex ecosystem behavior has been identified as a priority research challenge for both ecology and paleoecology. The paleoecological record can play a role in understanding the processes behind abrupt ecological change because it enables the reconstruction of processes occurring over decadal-centennial timescales or longer. Therefore, paleoecological data can be used to identify the existence of ecological thresholds and to investigate the environmental processes that can lead to loss of resilience and abrupt transitions between alternate states. In addition, incidences of abrupt vegetation changes in the past can serve as palaeoecological model systems; analogues of abrupt dynamics which can be used to test theories surrounding ecological responses to climate change. Here, I present examples from a range of terrestrial ecosystems (Holocene environmental changes from a coastal lagoon in the Galapagos Islands; Northern European vegetation changes since the last deglaciation; the North American hemlock decline) demonstrating evidence of abrupt ecosystem change. For each system I present a set of statistical techniques tailored to distin- guish between extrinsic versus intrinsically mediated ecological responses. Examples are provided from both single sites (i.e. landscape scale) and multiple sites (regional-continental scale). These techniques provide a

  20. Warm summer nights and the growth decline of shore pine in Southeast Alaska

    Sullivan, Patrick F.; Mulvey, Robin L.; Brownlee, Annalis H.; Barrett, Tara M.; Pattison, Robert R.


    Shore pine, which is a subspecies of lodgepole pine, was a widespread and dominant tree species in Southeast Alaska during the early Holocene. At present, the distribution of shore pine in Alaska is restricted to coastal bogs and fens, likely by competition with Sitka spruce and Western hemlock. Monitoring of permanent plots as part of the United States Forest Service Forest Inventory and Analysis program identified a recent loss of shore pine biomass in Southeast Alaska. The apparent loss of shore pine is concerning, because its presence adds a vertical dimension to coastal wetlands, which are the richest plant communities of the coastal temperate rainforest in Alaska. In this study, we examined the shore pine tree-ring record from a newly established plot network throughout Southeast Alaska and explored climate-growth relationships. We found a steep decline in shore pine growth from the early 1960s to the present. Random Forest regression revealed a strong correlation between the decline in shore pine growth and the rise in growing season diurnal minimum air temperature. Warm summer nights, cool daytime temperatures and a reduced diurnal temperature range are associated with greater cloud cover in Southeast Alaska. This suite of conditions could lead to unfavorable tree carbon budgets (reduced daytime photosynthesis and greater nighttime respiration) and/or favor infection by foliar pathogens, such as Dothistroma needle blight, which has recently caused widespread tree mortality on lodgepole pine plantations in British Columbia. Further field study that includes experimental manipulation (e.g., fungicide application) will be necessary to identify the proximal cause(s) of the growth decline. In the meantime, we anticipate continuation of the shore pine growth decline in Southeast Alaska.

  1. Investigating substrate use efficiency across different microbial physiologies in soil-extracted, solubilized organic matter (SESOM)

    Cyle, K. T.; Martinez, C. E.


    Recent experimental work has elevated the importance of microbial processing for the stabilization of fresh carbon inputs within the soil mineral matrix. Enhancing our understanding of soil carbon and nitrogen dynamics therefore requires a better understanding of how efficiently microbial metabolism can process low molecular weight carbon substrates (carbon use efficiency, CUE) under environmentally relevant conditions. One approach to better understanding microbial uptake rates and CUE is the ecophysiological study of soil isolates in liquid media culture consisting of soil-extracted solubilized organic matter (SESOM). We are using SESOM from an Oa horizon under hemlock hardwood vegetation in upstate New York as liquid media for the growth of 12 isolates from the Oa and B horizon of the same site. Here we seek to test the uptake rates as well as CUE of 5 different low molecular weight substrates spanning compound class and nominal oxidation state (glucose, acetate, formate, glycine, valine) by isolates differing in phylogeny and physiology. The use of a spike of each of the 13C-labeled substrates into SESOM, along with a 0.2 μm filtration step, allows accurate partitioning of labeled carbon between biomass, gaseous CO2 as well as the exometabolome. Coupled UHPLC-MS measurements are being used to identify and determine uptake rates of over 80 potential C substrates present in the extract as well as our labeled substrate of interest along the course of the isolate growth curve. This work seeks to utilize a gradient in substrate class as well as microbial physiologies to inform our understanding of C and N cycling under relevant soil solution conditions. Future experiments may also use labeled biomass from stationary phase to investigate the stabilization potential of anabolic products formed from each substrate with a clay fraction isolated from the same site.

  2. Forest structure and light regimes following moderate wind storms: implications for multi-cohort management.

    Hanson, Jacob J; Lorimer, Craig G


    Moderate-severity disturbances appear to be common throughout much of North America, but they have received relatively little detailed study compared to catastrophic disturbances and small gap dynamics. In this study, we examined the immediate impact of moderate-intensity wind storms on stand structure, opening sizes, and light regimes in three hemlock-hardwood forests of northeastern Wisconsin. These were compared to three stands managed by single-tree and group selection, the predominant forest management system for northern hardwoods in the region. Wind storms removed an average of 41% of the stand basal area, compared to 27% removed by uneven-aged harvests, but both disturbances removed trees from a wide range of size classes. The removal of nearly half of the large trees by wind in two old-growth stands caused partial retrogression to mature forest structure, which has been hypothesized to be a major disturbance pathway in the region. Wind storms resulted in residual stand conditions that were much more heterogeneous than in managed stands. Gap sizes ranged from less than 10 m2 up to 5000 m2 in wind-disturbed stands, whereas the largest opening observed in managed stands was only 200 m2. Wind-disturbed stands had, on average, double the available solar radiation at the forest floor compared to managed stands. Solar radiation levels were also more heterogeneous in wind-disturbed stands, with six times more variability at small scales (0.1225 ha) and 15 times more variability at the whole-stand level. Modification of uneven-aged management regimes to include occasional harvests of variable intensity and spatial pattern may help avoid the decline in species diversity that tends to occur after many decades of conventional uneven-aged management. At the same time, a multi-cohort system with these properties would retain a high degree of average crown cover, promote structural heterogeneity typical of old-growth forests, and maintain dominance by late

  3. Five-Year Impacts of Swiss Needle Cast on Douglas-fir in Interior Forests of Oregon, USA

    FILIP, Gregory


    Full Text Available In 2001 and 2006, we examined 590 Douglas-firs in 59 stands age 10-23 years in the northern Cascade Mountain foothills in Oregon, USA. Mean 5-year-dbh growth was 6.1 cm and total-height growth was 3.6 m. Mean needle-retention index increased by 3.4 over 5 years, and mid-crown retention increased by 1.2 years. Mean percentages of stomata occluded by pseudothecia of Phaeocryptopus gaeumannii were 13.6% for 2000-(2-year-old needles and 1.7% for 2001-(1-year-old needles sampled in 2002, and 13.3% for 2004 (2-year-old needles sampled in 2006. Mean crown-length to sapwood-area ratio was 5.2 cm/cm2 in 2006. There were poor correlations (R2 <0.3 among all variables except for a moderate correlation between stand elevation and either 2000-stomata occluded (R2 = 0.43 or 2004-stomata occluded (R2 = 0.50, where there were fewer pseudothecia at the higher elevations. Either 5 years is not enough time to evaluate the affects of Swiss needle cast on Douglas-fir growth in the Oregon Cascades or there was no significant effect of Swiss needle cast during the latest outbreak on Douglas-fir growth. Based on our results and their interpretation, forest managers may need not alter their current practices in the northern Oregon Cascades, and managing a mix of Douglas-fir and western hemlock at lower elevations and noble fir at higher elevations will help offset any future stand-growth declines due to Swiss needle cast.

  4. Feeding by Leucopis argenticollis and Leucopis piniperda (Diptera: Chamaemyiidae) from the western USA on Adelges tsugae (Hemiptera: Adelgidae) in the eastern USA.

    Motley, K; Havill, N P; Arsenault-Benoit, A L; Mayfield, A E; Ott, D S; Ross, D; Whitmore, M C; Wallin, K F


    Leucopis argenticollis (Zetterstedt) and Leucopis piniperda (Malloch) are known to feed on the lineage of Adelges tsugae Annand that is native to western North America, but it is not known if they will survive on the lineage that was introduced from Japan to the eastern USA. In 2014, western Leucopis spp. larvae were brought to the laboratory and placed on A. tsugae collected in either Washington (North American A. tsugae lineage) or Connecticut (Japanese lineage). There were no significant differences in survival or developmental times between flies reared on the two different adelgid lineages. In 2015 and 2016, western Leucopis spp. adults were released at two different densities onto enclosed branches of A. tsugae infested eastern hemlock (Tsuga canadensis (L.) Carr.) in Tennessee and New York. Cages were recovered and their contents examined 4 weeks after release at each location. Leucopis spp. larvae and puparia of the F1 generation were recovered at both release locations and adults of the F1 generation were collected at the Tennessee location. The number of Leucopis spp. offspring collected increased with increasing adelgid density, but did not differ by the number of adult flies released. Flies recovered from cages and flies collected from the source colony were identified as L.argenticollis and L. piniperda using DNA barcoding. These results demonstrate that Leucopis spp. from the Pacific Northwest are capable of feeding and developing to the adult stage on A. tsugae in the eastern USA and they are able to tolerate environmental conditions during late spring and early summer at the southern and northern extent of the area invaded by A. tsugae in the eastern USA.

  5. Education and distance learning: changing the trends.

    Merrell, Ronald C


    Training and instruction are activities deeply ingrained in human relations and derive from the critical need for the young to learn survival skills. The responsibility in primitive society for such training almost certainly fell to parents who continued their pedagogical role after childhood issues to include hunting, gathering, fine motor activities and other life skills needed for personal or family survival. Such instruction only ended when the young were ready for independent life and- contribution to tribal well-being. Delegation of teaching to others was inevitable. Teaching has become a specialty and has at least one interesting story in ancient literature. Ulysses was certain to be away at the Trojan War and subsequent adventures for many years. He would not be able to provide his son, Telemachus, with the guidance and training to prepare him for adulthood. Therefore, he asked Mentor to act In Loco Parentis and instruct the young man toward competence and adult success. Teaching as a profession and discipline has been through many stages and many controversies. Socrates was a great teacher with a distinct technique for learning by questioning. His influence on his students was profound. Plato was such a good student he recorded all the master's works. Socrates has never been credited with even the briefest lecture note. As great as he was Socrates was forced to drink the bitter hemlock because his teaching was considered a corruption of youth rather than a proper preparation for effective adulthood. Dissonance between the expectations of learners, parents and teachers has a rich history. Certainly even now education is not something the professoriate may invent for the naïve learner and then expect grateful acquiesce with faithful learning. Learning has -dimensions in human psychology and communication. The learners' autonomy, privacy and motivation cannot be denied. Learning is collaboration with teacher and the endpoint is the acquisition of new

  6. Dendrogeomorphic Assessment of the Rattlesnake Gulf Landslide in the Tully Valley, Onondaga County, New York

    Tamulonis, Kathryn L.; Kappel, William M.


    Dendrogeomorphic techniques were used to assess soil movement within the Rattlesnake Gulf landslide in the Tully Valley of central New York during the last century. This landslide is a postglacial, slow-moving earth slide that covers 23 acres and consists primarily of rotated, laminated, glaciolacustrine silt and clay. Sixty-two increment cores were obtained from 30 hemlock (Tsuga canadensis) trees across the active part of the landslide and from 3 control sites to interpret the soil-displacement history. Annual growth rings were measured and reaction wood was identified to indicate years in which ring growth changed from concentric to eccentric, on the premise that soil movement triggered compensatory growth in displaced trees. These data provided a basis for an 'event index' to identify years of landslide activity over the 108 years of record represented by the oldest trees. Event-index values and total annual precipitation increased during this time, but years with sudden event-index increases did not necessarily correspond to years with above-average precipitation. Multiple-regression and residual-values analyses indicated a possible correlation between precipitation and movement within the landslide and a possible cyclic (decades-long) tree-ring response to displacement within the landslide area from the toe upward to, and possibly beyond, previously formed landslide features. The soil movement is triggered by a sequence of factors that include (1) periods of several months with below-average precipitation followed by persistent above-average precipitation, (2) the attendant increase in streamflow, which erodes the landslide toe and results in an upslope propagation of slumping, and (3) the harvesting of mature trees within this landslide during the last century and continuing to the present.

  7. Seasonal variation in the temperature sensitivity of proteolytic enzyme activity in temperate forest soils

    Brzostek, Edward R.; Finzi, Adrien C.


    Increasing soil temperature has the potential to alter the activity of the extracellular enzymes that mobilize nitrogen (N) from soil organic matter (SOM) and ultimately the availability of N for primary production. Proteolytic enzymes depolymerize N from proteinaceous components of SOM into amino acids, and their activity is a principal driver of the within-system cycle of soil N. The objectives of this study were to investigate whether the soils of temperate forest tree species differ in the temperature sensitivity of proteolytic enzyme activity over the growing season and the role of substrate limitation in regulating temperature sensitivity. Across species and sampling dates, proteolytic enzyme activity had relatively low sensitivity to temperature with a mean activation energy (Ea) of 33.5 kJ mol-1. Ea declined in white ash, American beech, and eastern hemlock soils across the growing season as soils warmed. By contrast, Eain sugar maple soil increased across the growing season. We used these data to develop a species-specific empirical model of proteolytic enzyme activity for the 2009 calendar year and studied the interactive effects of soil temperature (ambient or +5°C) and substrate limitation (ambient or elevated protein) on enzyme activity. Declines in substrate limitation had a larger single-factor effect on proteolytic enzyme activity than temperature, particularly in the spring. There was, however, a large synergistic effect of increasing temperature and substrate supply on proteolytic enzyme activity. Our results suggest limited increases in N availability with climate warming unless there is a parallel increase in the availability of protein substrates.

  8. Incorporating interspecific competition into species-distribution mapping by upward scaling of small-scale model projections to the landscape.

    Mark Baah-Acheamfour

    Full Text Available There are a number of overarching questions and debate in the scientific community concerning the importance of biotic interactions in species distribution models at large spatial scales. In this paper, we present a framework for revising the potential distribution of tree species native to the Western Ecoregion of Nova Scotia, Canada, by integrating the long-term effects of interspecific competition into an existing abiotic-factor-based definition of potential species distribution (PSD. The PSD model is developed by combining spatially explicit data of individualistic species' response to normalized incident photosynthetically active radiation, soil water content, and growing degree days. A revised PSD model adds biomass output simulated over a 100-year timeframe with a robust forest gap model and scaled up to the landscape using a forestland classification technique. To demonstrate the method, we applied the calculation to the natural range of 16 target tree species as found in 1,240 provincial forest-inventory plots. The revised PSD model, with the long-term effects of interspecific competition accounted for, predicted that eastern hemlock (Tsuga canadensis, American beech (Fagus grandifolia, white birch (Betula papyrifera, red oak (Quercus rubra, sugar maple (Acer saccharum, and trembling aspen (Populus tremuloides would experience a significant decline in their original distribution compared with balsam fir (Abies balsamea, black spruce (Picea mariana, red spruce (Picea rubens, red maple (Acer rubrum L., and yellow birch (Betula alleghaniensis. True model accuracy improved from 64.2% with original PSD evaluations to 81.7% with revised PSD. Kappa statistics slightly increased from 0.26 (fair to 0.41 (moderate for original and revised PSDs, respectively.

  9. Mitigating Uncertainty from Vegetation Spatial Complexity with Highly Portable Lidar

    Paynter, I.; Schaaf, C.; Peri, F.; Saenz, E. J.; Genest, D.; Strahler, A. H.; Li, Z.


    To fully utilize the excellent spatial coverage and temporal resolution offered by satellite resources for estimating ecological variables, fine-scale observations are required for comparison, calibration and validation. Lidar instruments have proved effective in estimating the properties of vegetation components of ecosystems, but they are often challenged by occlusion, especially in structurally complex and spatially fragmented ecosystems such as tropical forests. Increasing the range of view angles, both horizontally and vertically, by increasing the number of scans, can mitigate occlusion. However these scans must occur within the window of temporal stability for the ecosystem and vegetation property being measured. The Compact Biomass Lidar (CBL) is a TLS optimized for portability and scanning speed, developed and operated by University of Massachusetts Boston. This 905nm wavelength scanner achieves an angular resolution of 0.25 degrees at a rate of 33 seconds per scan. The ability to acquire many scans within narrow windows of temporal stability for ecological variables has facilitated the more complete investigation of ecosystem structural characteristics, and their expression as a function of view angle. The lightweight CBL has facilitated the use of alternative deployment platforms including towers, trams and masts, allowing analysis of the vertical structure of ecosystems, even in highly enclosed environments such as the sub-canopy of tropical forests where aerial vehicles cannot currently operate. We will present results from view angle analyses of lidar surveys of tropical rainforest in La Selva, Costa Rica where the CBL was deployed at heights up to 10m in Carbono long-term research plots utilizing a portable mast, and on a 25m stationary tower; and temperate forest at Harvard Forest, Massachusetts, USA, where the CBL has been deployed biannually at long-term research plots of hardwood and hemlock, as well as at heights of up to 25m utilizing a

  10. Legal recourse for damages suffered from low-level radiation exposure

    Pesto-Edwards, M.M.


    In the past few years several events involving toxic substances have received widespread coverage by the media, thereby altering an already aware population to the hazards of exposure to toxic agents. Incidents such as Three Mile Islane, Love Canal, and Hemlock, Michigan, the exposure plight of veterans to radiation at the Nevada Test Site and to Agent Orange in Vietnam, and to the exposure of factory workers to asbestos, have been highly publicized. In part because of this publicity, the emphasis of the 1970's on controlling water and air pollution has been shifting slowly during the 1980s to the of control of hazardous waste pollution. Despite this shifting emphasis, legislative and judicial systems have been slow to respond. Few remedies are available to real and imagined victims of toxic substances. From a legal point of view, there is little difference between exposure to low levels of radiation and low levels of toxic chemicals. Both instances fall under the broader domain of environmental law. Depending on the circumstances, one instance might provide legal precedent for the other. This chapter presents examples drawn from both areas in order to illustrate current issues. The discussion is divided into four parts: (1) the common law tort theories that may be asserted when a plaintiff has suffered injury resulting from exposure to low-level radiation or other toxic substances; (2) the difficulties posed by the relief mechanisms rooted in traditional common law; (3) current federal legislation, along with its merits and shortcomings; and (4) solutions to the obstacles now faced by plaintiffs in attempting to recover their damages. Also discussed are suggested judicial and legislative solutions designed to remedy the damages caused to persons exposed to toxic wastes

  11. Formation and maintenance of a forced pool-riffle couplet following loading of large wood

    Thompson, D. M.; Fixler, S. A.


    Pool-riffle maintenance has been documented in numerous studies, but it has been almost impossible to characterize detailed natural pool-riffle formation mechanisms because of the lack of baseline data prior to pool establishment. In 2013, a study was conducted on the Blackledge River in Connecticut to document the formation of a new pool-riffle couplet on a section of river that had previously been studied from 1999 to 2001. In 2001, the study reach contained a scour hole with a residual depth of 0.08 ± 0.09 m downstream of a 1930s paired deflector with no identifiable riffle immediately downstream. At this time, a large, severely undercut, hemlock tree was noted along the left bank. Sometime between fall 2001 and 2004, the tree fell perpendicular to flow across the channel and formed a large wood (LW) jam and new pool-riffle couplet several meters downstream of the old scour hole. Pool spacing along the reach decreased from 4.47 bankfull widths (BFW) in 1999 to 3.83 BFW after the new pool-riffle couplet formed. The new pool has a residual depth, the water depth of the streambed depression below the elevation of the immediate downstream hydraulic control, of 1.36 ± 0.075 to 1.59 ± 0.075 m, which resulted from a combination of 1.32 ± 0.09 m or less of incision below the old scour hole (95.6% or less of the depth increase) and up to 0.18 ± 0.09 m of downstream deposition and associated backwater formation (13.2% or less of the depth increase). To assess dynamic stability of the pool-riffle couplet over several flood cycles, surficial fine-sediment and organic material along the reach were quantified. The 23-m-long pool stores 25.7% of the surficial fine grained sediments and 15.4% of organic material along a 214-m-long reach that includes one additional artificially created pool. An adjacent 50-m-long secondary channel impacted by the LW jam stores 65.3% of the surficial fine-grained sediments and 54.8% of organic material along the full reach.

  12. Novel and Lost Forests in the Upper Midwestern United States, from New Estimates of Settlement-Era Composition, Stem Density, and Biomass.

    Goring, Simon J; Mladenoff, David J; Cogbill, Charles V; Record, Sydne; Paciorek, Christopher J; Jackson, Stephen T; Dietze, Michael C; Dawson, Andria; Matthes, Jaclyn Hatala; McLachlan, Jason S; Williams, John W


    EuroAmerican land-use and its legacies have transformed forest structure and composition across the United States (US). More accurate reconstructions of historical states are critical to understanding the processes governing past, current, and future forest dynamics. Here we present new gridded (8x8km) reconstructions of pre-settlement (1800s) forest composition and structure from the upper Midwestern US (Minnesota, Wisconsin, and most of Michigan), using 19th Century Public Land Survey System (PLSS), with estimates of relative composition, above-ground biomass, stem density, and basal area for 28 tree types. This mapping is more robust than past efforts, using spatially varying correction factors to accommodate sampling design, azimuthal censoring, and biases in tree selection. We compare pre-settlement to modern forests using US Forest Service Forest Inventory and Analysis (FIA) data to show the prevalence of lost forests (pre-settlement forests with no current analog), and novel forests (modern forests with no past analogs). Differences between pre-settlement and modern forests are spatially structured owing to differences in land-use impacts and accompanying ecological responses. Modern forests are more homogeneous, and ecotonal gradients are more diffuse today than in the past. Novel forest assemblages represent 28% of all FIA cells, and 28% of pre-settlement forests no longer exist in a modern context. Lost forests include tamarack forests in northeastern Minnesota, hemlock and cedar dominated forests in north-central Wisconsin and along the Upper Peninsula of Michigan, and elm, oak, basswood and ironwood forests along the forest-prairie boundary in south central Minnesota and eastern Wisconsin. Novel FIA forest assemblages are distributed evenly across the region, but novelty shows a strong relationship to spatial distance from remnant forests in the upper Midwest, with novelty predicted at between 20 to 60km from remnants, depending on historical forest

  13. Biogeomorphic and pedogenic impact of trees in three soil regions

    Pawlik, Łukasz; Šamonil, Pavel


    Vegetation is an important factor of soil formation which together with topography, geology, climate and time modulates chemical and physical soil characteristics. Tree/soils/regolith interaction was recognized in recently uprooted trees and relict treethrow mounds and pits. In our present study we focus on effects of individual standing trees in pedogenesis and biogeomorphic processes. Constant pressure of tree root systems, changing hydric and temperature regime, together with rhizospheric microbes and root mycorrhizal associations may cause multiscale alterations to regolith and soils. We hypothesize different soil chemical properties under old tree stumps compared to unaffected control pedon resulted from affected pedogenetical pathways at the analyzed microsites. The present project highlights changes in soil properties under tree stumps in three different soil regions: Haplic Cambisols (Turbacz Reserve, Gorce Mts., Poland, hereafter HC), Entic Podzols (Zofin Reserve, Novohradske Mts., the Czech Republic, hereafter EP), Albic Podzols (Upper Peninsula, Michigan, USA, hereafter AP). These three regions represent different degrees of soil weathering and leaching. Pedons under fir, beech and hemlock stumps, as well as unaffected control pedons were sampled and laboratory analyzed for several chemical properties; active and exchangeable soil reaction, oxidized carbon, total nitrogen, and various forms of Fe, Al, Mn and Si. At the same time we studied age of the sampled tree stumps, as well as age of their death using radiocarbon technique and dendrochronology. While no effects of the soil-trees interactions can be visible on hillslope surface, we found important evidence of biomechanical activities of tree roots (e.g. root channels) and biochemical changes which add to the discussion about biogeomorphic and pedogenic significance of trees and tree roots as drivers of biomechanical weathering and soil processes in the decadal and centennial time scales. Preliminary

  14. Calibrating and testing a gap model for simulating forest management in the Oregon Coast Range

    Pabst, R.J.; Goslin, M.N.; Garman, S.L.; Spies, T.A.


    The complex mix of economic and ecological objectives facing today's forest managers necessitates the development of growth models with a capacity for simulating a wide range of forest conditions while producing outputs useful for economic analyses. We calibrated the gap model ZELIG to simulate stand-level forest development in the Oregon Coast Range as part of a landscape-scale assessment of different forest management strategies. Our goal was to incorporate the predictive ability of an empirical model with the flexibility of a forest succession model. We emphasized the development of commercial-aged stands of Douglas-fir, the dominant tree species in the study area and primary source of timber. In addition, we judged that the ecological approach of ZELIG would be robust to the variety of other forest conditions and practices encountered in the Coast Range, including mixed-species stands, small-scale gap formation, innovative silvicultural methods, and reserve areas where forests grow unmanaged for long periods of time. We parameterized the model to distinguish forest development among two ecoregions, three forest types and two site productivity classes using three data sources: chronosequences of forest inventory data, long-term research data, and simulations from an empirical growth-and-yield model. The calibrated model was tested with independent, long-term measurements from 11 Douglas-fir plots (6 unthinned, 5 thinned), 3 spruce-hemlock plots, and 1 red alder plot. ZELIG closely approximated developmental trajectories of basal area and large trees in the Douglas-fir plots. Differences between simulated and observed conifer basal area for these plots ranged from -2.6 to 2.4 m2/ha; differences in the number of trees/ha ???50 cm dbh ranged from -8.8 to 7.3 tph. Achieving these results required the use of a diameter-growth multiplier, suggesting some underlying constraints on tree growth such as the temperature response function. ZELIG also tended to overestimate

  15. Effects of different types of moderate severity disturbance on forest structural complexity and ecosystem functioning: A story of ice and fire

    Fahey, R. T.; Atkins, J.; Gough, C. M.; Hardiman, B. S.; Haber, L.; Stuart-Haentjens, E.; David, O.; Campbell, J. L.; Rustad, L.; Duffy, M.


    Disturbances that alter the structure and function of forest ecosystems occur along a continuum of severity. In contrast to the extremes of the disturbance gradient (i.e., stand-replacing disturbance and small gap formation), moderate severity disturbances are poorly understood, even though they make up the majority of the gradient and their spatial extent (and likely overall importance to regional disturbance regimes) often exceeds that of more severe disturbances. Moderate severity disturbances originate from a variety of causes, such as fires, ice storms, or pest and pathogen outbreaks, and each of these could reshape structure and function in different ways. Observational data from a limited number of sites shows that moderate disturbance can increase ecosystem complexity, but the generality of this effect has not been tested across a broad range of disturbance types and severities. Here, we utilize data from a set of five case studies of experimental or natural moderate disturbance to assess the effects of different types and severities of disturbance on forest canopy structural complexity (CSC) and the relationship of canopy structure with ecosystem functioning. Using pre- and post-disturbance measures of CSC derived from aerial and terrestrial LiDAR, UAV imagery, and Landsat data we quantified changes in CSC following an experimental ice storm, a low-severity surface fire, Beech Bark Disease and Hemlock Wooly Adelgid outbreaks, and experimental accelerated succession. Our initial findings indicate that different disturbance types have highly variable effects on CSC, and also that progressive increases in disturbance severity alter CSC differently among disturbance types. Differential effects of variable disturbance types on CSC has implications for the carbon cycle, as forest structure is strongly linked with both growth-limiting resource (e.g., nutrients and light) acquisition and net primary productivity. Understanding how different types and severities of

  16. Poisonous plants in New Zealand: a review of those that are most commonly enquired about to the National Poisons Centre.

    Slaughter, Robin J; Beasley, D Michael G; Lambie, Bruce S; Wilkins, Gerard T; Schep, Leo J


    New Zealand has a number of plants, both native and introduced, contact with which can lead to poisoning. The New Zealand National Poisons Centre (NZNPC) frequently receives enquiries regarding exposures to poisonous plants. Poisonous plants can cause harm following inadvertent ingestion, via skin contact, eye exposures or inhalation of sawdust or smoked plant matter. The purpose of this article is to determine the 15 most common poisonous plant enquiries to the NZNPC and provide a review of current literature, discussing the symptoms that might arise upon exposure to these poisonous plants and the recommended medical management of such poisonings. Call data from the NZNPC telephone collection databases regarding human plant exposures between 2003 and 2010 were analysed retrospectively. The most common plants causing human poisoning were selected as the basis for this review. An extensive literature review was also performed by systematically searching OVID MEDLINE, ISI Web of Science, Scopus and Google Scholar. Further information was obtained from book chapters, relevant news reports and web material. For the years 2003-2010 inclusive, a total of 256,969 enquiries were received by the NZNPC. Of these enquiries, 11,049 involved exposures to plants and fungi. The most common poisonous plant enquiries, in decreasing order of frequency, were: black nightshade (Solanum nigrum), arum lily (Zantedeschia aethiopica), kowhai (Sophora spp.), euphorbia (Euphorbia spp.), peace lily (Spathiphyllum spp.), agapanthus (Agapanthus spp.), stinking iris (Iris foetidissima), rhubarb (Rheum rhabarbarum), taro (Colocasia esculentum), oleander (Nerium oleander), daffodil (Narcissus spp.), hemlock (Conium maculatum), karaka (Corynocarpus laevigatus), foxglove (Digitalis purpurea) and ongaonga/New Zealand tree nettle (Urtica ferox). The combined total of enquiries for these 15 species was 2754 calls (representing approximately 25% of all enquiries regarding plant exposures). The signs

  17. Using Tree-Ring Data to Develop Critical Scientific and Mathematical Thinking Skills in Undergraduate Students

    Fiondella, F.; Davi, N. K.; Wattenberg, F.; Pringle, P. T.; Greidanus, I.; Oelkers, R.


    Tree-ring science provides an engaging, intuitive, and relevant entryway into understanding both climate change and environmental research. It also sheds light on the process of science--from inspiration, to fieldwork, to analysis, to publishing and communication. The basic premise of dendrochronology is that annual rings reflect year-to-year environmental conditions and that by studying long-lived trees we can learn about environmental and climatic conditions going back hundreds to thousands of years. Conceptually, this makes tree-ring studies accessible to students and faculty for a number of reasons. First, in order to collect their data, dendrochronologists often launch expeditions to stunningly picturesque and remote places in search of long-lived, climate sensitive trees. The exciting stories and images that scientists bring back from the field can help connect students to the studies, their motivation, and the data collected. Second, tree rings can be more easily explained as a proxy for climate than ice cores, speleothems and others. Most people have prior knowledge about trees and annual growth rings. It is even possible, for example, for non-expert audiences to see climate variability through time with the naked eye by looking at climate-sensitive tree cores. Third, tree rings are interdisciplinary and illustrate the interplay between the mathematical sciences, the biological sciences, and the geosciences—that is, they show that the biosphere is a fundamental component of the Earth system. Here, we present online, multi-media learning modules for undergraduates that introduce students to several foundational studies in tree-ring science. These include evaluating tree-ring cores from ancient hemlock trees growing on a talus slope in New Paltz, NY to learn about drought in the Northeastern US, evaluating long-term streamflow and drought of the Colorado River based on tree-ring records, and using tree-ring dating techniques to develop construction

  18. The Heidelberg Basin Drilling Project - Sedimentology and Stratigraphy of the Quaternary succession

    Ellwanger, Dietrich; Gabriel, Gerald; Hahne, Jürgen; Hoselmann, Christian; Menzies, John; Simon, Theo; Weidenfeller, Michael; Wielandt-Schuster, Ulrike


    Within the context of the Heidelberg Basin Drilling Project (Gabriel et al. 2008), a detailed sediment succession is presented here based upon deep drillings taken at Heidelberg UniNord and Mannheim Käfertal. Sediment structures, and micromorphological and pollen analyses were conducted and used to reconsider some of the climate transitions within the lower Pleistocene. A new and novel scenario is postulated regarding the preservation of Quaternary sediment packages within the Cenozoic Graben environment of the Heidelberg basin. The palynological evidence comprises the periods of warm climate of the Holsteinian (mainly Abies (fir), some Fagus (beech), Pterocarya & Azolla); the Cromerian (Pinus-Picea-QM (pine-spruce-QM)); the Bavelian (Abies, Tsuga (hemlock fir), QM & phases of increased NAP including Pinus); the Waalian (Abies, Tsuga, QM); and the Tiglian (Fagus & early Pleistocene taxa especially Sciadopytis, downward increasing Tertiary taxa). The sediment package was studied both macroscopically and microscopically. Both techniques provide evidence of fluvial, lacustrine and mass movement sedimentary processes. Some include evidence of periglacial processes (silt droplets within fine grained sands indicative of frozen ground conditions). The periglacial structures are often, not always, accompanied by pollen spectra dominated by pine and NAP. E.g. the Tiglian part of the succession shows periglacial sediment structures at its base and top but not in its middle sections. I.e. it appears not as a series of warm and cold phases but rather as a constant warm period with warm-cold-alternations at its bottom and top. All results illustrate sediment preservation in the Heidelberg basin almost throughout the Quaternary. This may be due to tectonic subsidence, but also to compaction by sediment loading of underlying fine sediments (Oligocene to Quaternary) leading to incomplete but virtually continuous sediment preservation (Tanner et al. 2009). References Gabriel, G

  19. Novel and Lost Forests in the Upper Midwestern United States, from New Estimates of Settlement-Era Composition, Stem Density, and Biomass.

    Simon J Goring

    Full Text Available EuroAmerican land-use and its legacies have transformed forest structure and composition across the United States (US. More accurate reconstructions of historical states are critical to understanding the processes governing past, current, and future forest dynamics. Here we present new gridded (8x8km reconstructions of pre-settlement (1800s forest composition and structure from the upper Midwestern US (Minnesota, Wisconsin, and most of Michigan, using 19th Century Public Land Survey System (PLSS, with estimates of relative composition, above-ground biomass, stem density, and basal area for 28 tree types. This mapping is more robust than past efforts, using spatially varying correction factors to accommodate sampling design, azimuthal censoring, and biases in tree selection.We compare pre-settlement to modern forests using US Forest Service Forest Inventory and Analysis (FIA data to show the prevalence of lost forests (pre-settlement forests with no current analog, and novel forests (modern forests with no past analogs. Differences between pre-settlement and modern forests are spatially structured owing to differences in land-use impacts and accompanying ecological responses. Modern forests are more homogeneous, and ecotonal gradients are more diffuse today than in the past. Novel forest assemblages represent 28% of all FIA cells, and 28% of pre-settlement forests no longer exist in a modern context. Lost forests include tamarack forests in northeastern Minnesota, hemlock and cedar dominated forests in north-central Wisconsin and along the Upper Peninsula of Michigan, and elm, oak, basswood and ironwood forests along the forest-prairie boundary in south central Minnesota and eastern Wisconsin. Novel FIA forest assemblages are distributed evenly across the region, but novelty shows a strong relationship to spatial distance from remnant forests in the upper Midwest, with novelty predicted at between 20 to 60km from remnants, depending on historical

  20. Recent Changes in Tree Species Abundance: Patterns, Trends, and Drivers Across Northeastern US Forests

    Gudex-Cross, D.; Pontius, J.; Adams, A.


    Monitoring trends in the abundance and distribution of tree species is essential to understanding potential impacts of climate change on forested ecosystems. Related studies to date have largely focused on modeling distributional shifts according to future climate scenarios or used field inventory data to examine compositional changes across broader landscapes. Here, we leverage a novel remote sensing technique that utilizes field data, multitemporal Landsat imagery, and spectral unmixing to model regional changes in the abundance (percent basal area) of key northeastern US species over a 30-year period (1985-2015). We examine patterns in how species abundance has changed, as well as their relationship with climate, landscape, and soil characteristics using spatial regression models. Results show significant declines in overall abundance for sugar maple ( 8.6% 30-yr loss), eastern hemlock ( 7.8% 30-yr loss), balsam fir ( 5.0% 30-yr loss), and red spruce ( 3.8% total 30-yr loss). Species that saw significant increasing abundance include American beech ( 7.0% 30-yr gain) and red maple ( 2.5% 30-yr gain). However, these changes were not consistent across the landscape. For example, red spruce is increasing at upper elevations with concurrent losses in balsam fir and birch species. Similarly, sugar maple decreases are concentrated at lower elevations, likely due to increases in American beech. Various abiotic factors were significantly associated with changes in species composition including landscape position (e.g. longitude, elevation, and heat load index) and ecologically-relevant climate variables (e.g. growing season precipitation and annual temperature range). Interestingly, there was a stronger relationship in abundance changes across longitudes, rather than latitudes or elevations as predicted in modeled species migration scenarios.These results indicate that the dominant composition of northeastern forests is changing in ways that run counter to accepted

  1. Alkaloids in the pharmaceutical industry: Structure, isolation and application

    Nikolić Milan


    Full Text Available By the end of the 18th and the beginning of the 19th century a new era began in medicine, pharmaceutics and chemistry that was strongly connected with alkaloids and alkaloid drugs. Even before that it was known that certain drugs administered in limited doses were medicines, and toxic if taken in larger doses (opium, coke leaves, belladonna roots, monkshood tubers crocus or hemlock seeds. However, the identification, isolation and structural characterization of the active ingredients of the alkaloid drugs was only possible in the mid 20th century by the use of modern extraction equipment and instrumental methods (NMR, X-ray diffraction and others.In spite of continuing use over a long time, there is still great interest in investigating new drugs, potential raw materials for the pharmaceutical industry, as well as the more detailed investigation and definition of bio-active components and the indication of their activity range, and the partial synthesis of new alkaloid molecules based on natural alkaloids. The scope of these investigations, especially in the field of semi-synthesis is to make better use of the bio-active ingredients of alkaloid drugs, i.e. to improve the pharmacological effect (stronger and prolonged effect of the medicine, decreased toxicity and side effects, or to extend or change the applications. A combined classification of alkaloids was used, based on the chemical structure and origin, i.e. the source of their isolation to study alkaloid structure. For practical reasons, the following classification of alkaloids was used: ergot alkaloids, poppy alkaloids, tropanic alkaloids purine derivative alkaloids, carbon-cyclic alkaloids, and other alkaloids. The second part of this report presents a table of general procedures for alkaloid isolation from plant drugs (extraction by water non-miscible solvents, extraction by water-miscible solvents and extraction by diluted acid solutions. Also, methods for obtaining chelidonine and

  2. Low Tree-Growth Elasticity of Forest Biomass Indicated by an Individual-Based Model

    Robbie A. Hember


    Full Text Available Environmental conditions and silviculture fundamentally alter the metabolism of individual trees and, therefore, need to be studied at that scale. However, changes in forest biomass density (Mg C ha−1 may be decoupled from changes in growth (kg C year−1 when the latter also accelerates the life cycle of trees and strains access to light, nutrients, and water. In this study, we refer to an individual-based model of forest biomass dynamics to constrain the magnitude of system feedbacks associated with ontogeny and competition and estimate the scaling relationship between changes in tree growth and forest biomass density. The model was driven by fitted equations of annual aboveground biomass growth (Gag, probability of recruitment (Pr, and probability of mortality (Pm parameterized against field observations of black spruce (Picea mariana (Mill. BSP, interior Douglas-fir (Pseudotsuga menziesii var. glauca (Beissn. Franco, and western hemlock (Tsuga heterophylla (Raf. Sarg.. A hypothetical positive step-change in mean tree growth was imposed half way through the simulations and landscape-scale responses were then evaluated by comparing pre- and post-stimulus periods. Imposing a 100% increase in tree growth above calibrated predictions (i.e., contemporary rates only translated into 36% to 41% increases in forest biomass density. This corresponded with a tree-growth elasticity of forest biomass (εG,SB ranging from 0.33 to 0.55. The inelastic nature of stand biomass density was attributed to the dependence of mortality on intensity of competition and tree size, which decreased stand density by 353 to 495 trees ha−1, and decreased biomass residence time by 10 to 23 years. Values of εG,SB depended on the magnitude of the stimulus. For example, a retrospective scenario in which tree growth increased from 50% below contemporary rates up to contemporary rates indicated values of εG,SB ranging from 0.66 to 0.75. We conclude that: (1 effects of

  3. Bioavailability of heavy metals, germanium and rare earth elements at Davidschacht dump-field in mine affected area of Freiberg (Saxony)

    Midula, Pavol; Wiche, Oliver


    with the other elements from this group. High amounts of As, Cd, Pb in mould horizons were proved. The surprisingly highest concentrations were determined for As (in average 3328 mg kg-1). The results of the pH measurement indicates acid conditions (in average 4.86, min. 3.89) for whole mine heap. Due to the mobility of Cd and Pb in acid environment, a high mobility of Cd in mobile soil fractions (in average 0.58 mg kg-1) was found, that seems to be responsible for the Cd pollution of Freiberger Mulde river, situated near the dump-field in the East direction from the studied area. The Pb content was in the average 1513 mg kg-1. SE analyses shows, that only the minor amounts of these metals were accounted in fractions I - IV (As: 7.75 %, Pb: 5.48 %, Cd: 26.77 %). The total Ge content in soil samples was 2.7 average. The concentrations of Nd and Ce were 17.7 mg kg-1and 38.5 mg kg-1, which is even lower than the average Nd and Ce contents in the Earth crust. However, the concentration of Ge was roughly a factor of two higher, than this average showing a large pool of Ge that could be accessed by phytoextraction. The SE analyses shows, that the average in fractions I - IV is even much lower, than in the case of the above mentioned heavy metals in comparison with Ge (1.75 %), Nd (3.28 %) and Ce (3.12 %). The BCF calculated for plants shows, that the only element, which could be possibly used as the object of phytoaccumulation is Cd (the BCF > 1) in species Populus tremula (3.0, 1.7), Spirea douglasii (1.4, 2.2) and Tanacetum vulgare (3.2, 1.3) at the most sampling places. Since these species represent the natural occurring vegetation of the dump, the use of these species together with soil amendments enhancing the plant availability of elements in soil fractions hold promise for phytoextraction of economically valuable metalloids and consequently an in situ bioremediation of the dump field. This work was realised with the support of Christin Jahns on behalf of the

  4. Root phenology at Harvard Forest and beyond

    Abramoff, R. Z.; Finzi, A.


    Roots are hidden from view and heterogeneously distributed making them difficult to study in situ. As a result, the causes and timing of root production are not well understood. Researchers have long assumed that above and belowground phenology is synchronous; for example, most parameterizations of belowground carbon allocation in terrestrial biosphere models are based on allometry and represent a fixed fraction of net C uptake. However, using results from metaanalysis as well as empirical data from oak and hemlock stands at Harvard Forest, we show that synchronous root and shoot growth is the exception rather than the rule. We collected root and shoot phenology measurements from studies across four biomes (boreal, temperate, Mediterranean, and subtropical). General patterns of root phenology varied widely with 1-5 production peaks in a growing season. Surprisingly, in 9 out of the 15 studies, the first root production peak was not the largest peak. In the majority of cases maximum shoot production occurred before root production (Offset>0 in 32 out of 47 plant sample means). The number of days offset between maximum root and shoot growth was negatively correlated with median annual temperature and therefore differs significantly across biomes (ANOVA, F3,43=9.47, pGrowth form (woody or herbaceous) also influenced the relative timing of root and shoot growth. Woody plants had a larger range of days between root and shoot growth peaks as well as a greater number of growth peaks. To explore the range of phenological relationships within woody plants in the temperate biome, we focused on above and belowground phenology in two common northeastern tree species, Quercus rubra and Tsuga canadensis. Greenness index, rate of stem growth, root production and nonstructural carbohydrate content were measured beginning in April 2012 through August 2013 at the Harvard Forest in Petersham, MA, USA. Greenness and stem growth were highest in late May and early June with one clear

  5. Palynostratigraphical correlation of the excavated Miocene lignite seams of the Yataǧan basin (Muǧla Province, south-western Turkey)

    Bouchal, Johannes Martin; Grímsson, Friðgeir; Denk, Thomas


    The excavated main lignite seams and overlying lacustrine sediments of the opencast mines Eskihisar, Salihpaşalar, and Tı naz, Muǧla Province, south-western Turkey were investigated using a high taxonomic resolution palynological approach. The Eskihisar section comprises 47m and 56 samples of which 30 were usable for palynological analysis. The Tı naz section comprises 75 m and 29 samples of which 15 were usable for palynological analysis. Finally, the Salihpaşalar section comprises 25 m and 26 samples of which 16 were usable for palynological analysis. The age of the palynological sections is middle to late Miocene based on radiometric dating and vertebrate fossils. In order to investigate dispersed pollen and spores and their botanical affinities a combined light microscopy and scanning electron microscopy approach was used. The rich palynoflora comprises: seven types of algal cysts (Botryococcus, Zygnemataceae), seventeen spore types belonging to Lycopsida (club mosses), Marsileaceae (water-clover), Osmundaceae, Pteridaceae (brake), and Polypodiaceae; 14 types of gymnosperm pollen belonging to Ephedraceae (Mormon tea), Cupressaceae, Pinaceae (Cathaya, cedar, hemlock, pine, spruce); five types of monocotyledone pollen belonging to Poaceae (grasses, common reed), and Typhaceae (bulrush, bur-reed); ca 90 dicotyledone pollen types belonging to Altingiaceae (sweet gum), Amaranthaceae (goosefoot), Anacardiaceae (sumac family), Apiaceae (parsley family), Aquifoliaceae (holly), Asteraceae (sunflower family), Betulaceae (alder, birch, hazel, hophornbeam, hornbeam), Campanulaceae (bellflower family), Cannabaceae (hackberries), Caprifoliaceae (honeysuckle, teasel family), Caryophyllaceae (pink family), Ericaceae (heather family), Eucommiaceae, Euphorbiaceae (spurge family), Fabaceae (bean family), Fagaceae (beech, oak), Geraniaceae (storkbills), Juglandaceae (hickory, walnut, wingnut), Lamiaceae (bagflower), Linaceae (flax), Lythraceae (waterwillow), Malvaceae

  6. Structural and compositional controls on transpiration in 40- and 450-year-old riparian forests in western Oregon, USA.

    Moore, Georgianne W; Bond, Barbara J; Jones, Julia A; Phillips, Nathan; Meinzer, Federick C


    Large areas of forests in the Pacific Northwest are being transformed to younger forests, yet little is known about the impact this may have on hydrological cycles. Previous work suggests that old trees use less water per unit leaf area or sapwood area than young mature trees of the same species in similar environments. Do old forests, therefore, use less water than young mature forests in similar environments, or are there other structural or compositional components in the forests that compensate for tree-level differences? We investigated the impacts of tree age, species composition and sapwood basal area on stand-level transpiration in adjacent watersheds at the H.J. Andrews Forest in the western Cascades of Oregon, one containing a young, mature (about 40 years since disturbance) conifer forest and the other an old growth (about 450 years since disturbance) forest. Sap flow measurements were used to evaluate the degree to which differences in age and species composition affect water use. Stand sapwood basal area was evaluated based on a vegetation survey for species, basal area and sapwood basal area in the riparian area of two watersheds. A simple scaling exercise derived from estimated differences in water use as a result of differences in age, species composition and stand sapwood area was used to estimate transpiration from late June through October within the entire riparian area of these watersheds. Transpiration was higher in the young stand because of greater sap flux density (sap flow per unit sapwood area) by age class and species, and greater total stand sapwood area. During the measurement period, mean daily sap flux density was 2.30 times higher in young compared with old Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) trees. Sap flux density was 1.41 times higher in young red alder (Alnus rubra Bong.) compared with young P. menziesii trees, and was 1.45 times higher in old P. menziesii compared with old western hemlock (Tsuga heterophylla (Raf

  7. Partitioning CO2 fluxes with isotopologue measurements and modeling to understand mechanisms of forest carbon sequestration

    Saleska, Scott [Univ. of Arizona, Tucson, AZ (United States); Davidson, Eric [Univ. of Arizona, Tucson, AZ (United States); Finzi, Adrien [Univ. of Arizona, Tucson, AZ (United States); Wehr, Richard [Univ. of Arizona, Tucson, AZ (United States); Moorcroft, Paul [Univ. of Arizona, Tucson, AZ (United States)


    photosynthesis (by ~25%) and daytime respiration (by ~100%) in the first half of the growing season at our site, and portrays ecosystem photosynthetic light-use efficiency as declining when in fact it is stable until autumnal senescence. B. Vegetation Phenology and belowground allocation: Findings: 1. Autotrophic respiration (Ra) showed a seasonal pattern, peaking in mid-summer when trees were most active. 2. The effective age of the substrate for belowground respiration is less than 2 weeks. 3. Above and belowground phenology are more synchronous in deciduous hardwood stands than evergreen hemlock stands. 4. The decline in root respiration rates in the fall is related to temperature rather than acclimation of root respiration or substrate limitations. Methodological Issues: 5. The isotopic signatures of autotrophic and heterotrophic respiration are too similar for isotopic partitioning of belowground respiration into these two components at our site—in keeping with the recent findings of Bowling et al. (2015) in a subalpine conifer forest. 6. Artifacts of the trenching method, such as changes in soil moisture and increased carbon substrate from the newly severed roots, are significant and need to be quantified when determining daily to annual estimates of autotrophic and heterotrophic respiration. C. Effects of simulated exudates on priming of microbial decomposition: The stoichiometry of root exudates influences both the amount and the mechanism by which priming occurs. At low C:N, SOC loss is caused by an increase in microbial efficiency. At high C:N, SOC loss is caused by an increase in microbial biomass. D. Modeling with the Ecosystem Demography Model (ED2): 1. Incorporation of 13C tracking to create an isotopically-enabled Ecosystem Demography v2 model (ED2) 2. State-of-the-art parameter optimization methodology developed for improving ED2 model predictions and parameters. 3. Significantly improved model predictions of growth- and maintenance-related carbon fluxes and 13

  8. Partitioning CO2 fluxes with isotopologue measurements and modeling to understand mechanisms of forest carbon sequestration

    Saleska, Scott [Univ. of Arizona, Tucson, AZ (United States); Davidson, Eric [Univ. of Arizona, Tucson, AZ (United States); Finzi, Adrien [Boston Univ., MA (United States); Wehr, Richdard [Harvard Univ., Cambridge, MA (United States); Moorcroft, Paul [Harvard Univ., Cambridge, MA (United States)


    daytime respiration (by ~100%) in the first half of the growing season at our site, and portrays ecosystem photosynthetic light-use efficiency as declining when in fact it is stable until autumnal senescence. B. Vegetation Phenology and belowground allocation: Findings: 1. Autotrophic respiration (Ra) showed a seasonal pattern, peaking in mid-summer when trees were most active. 2. The effective age of the substrate for belowground respiration is less than 2 weeks. 3. Above and belowground phenology are more synchronous in deciduous hardwood stands than evergreen hemlock stands. 4. The decline in root respiration rates in the fall is related to temperature rather than acclimation of root respiration or substrate limitations. Methodological Issues: 5. The isotopic signatures of autotrophic and heterotrophic respiration are too similar for isotopic partitioning of belowground respiration into these two components at our site—in keeping with the recent findings of Bowling et al. (2015) in a subalpine conifer forest. 6. Artifacts of the trenching method, such as changes in soil moisture and increased carbon substrate from the newly severed roots, are significant and need to be quantified when determining daily to annual estimates of autotrophic and heterotrophic respiration. C. Effects of simulated exudates on priming of microbial decomposition: The stoichiometry of root exudates influences both the amount and the mechanism by which priming occurs. At low C:N, SOC loss is caused by an increase in microbial efficiency. At high C:N, SOC loss is caused by an increase in microbial biomass. D. Modeling with the Ecosystem Demography Model (ED2): 1. Incorporation of 13C tracking to create an isotopically-enabled Ecosystem Demography v2 model (ED2) 2. State-of-the-art parameter optimization methodology developed for improving ED2 model predictions and parameters. 3. Significantly improved model predictions of growth- and maintenance-related carbon fluxes and 13C fluxes