WorldWideScience

Sample records for height leaf area

  1. The relationship between tree height and leaf area: sapwood area ratio.

    Science.gov (United States)

    McDowell, N; Barnard, H; Bond, B; Hinckley, T; Hubbard, R; Ishii, H; Köstner, B; Magnani, F; Marshall, J; Meinzer, F; Phillips, N; Ryan, M; Whitehead, D

    2002-06-01

    The leaf area to sapwood area ratio (A l :A s ) of trees has been hypothesized to decrease as trees become older and taller. Theory suggests that A l :A s must decrease to maintain leaf-specific hydraulic sufficiency as path length, gravity, and tortuosity constrain whole-plant hydraulic conductance. We tested the hypothesis that A l :A s declines with tree height. Whole-tree A l :A s was measured on 15 individuals of Douglas-fir (Pseudotsuga menziesii var. menziesii) ranging in height from 13 to 62 m (aged 20-450 years). A l :A s declined substantially as height increased (P=0.02). Our test of the hypothesis that A l :A s declines with tree height was extended using a combination of original and published data on nine species across a range of maximum heights and climates. Meta-analysis of 13 whole-tree studies revealed a consistent and significant reduction in A l :A s with increasing height (P<0.05). However, two species (Picea abies and Abies balsamea) exhibited an increase in A l :A s with height, although the reason for this is not clear. The slope of the relationship between A l :A s and tree height (ΔA l :A s /Δh) was unrelated to mean annual precipitation. Maximum potential height was positively correlated with ΔA l :A s /Δh. The decrease in A l :A s with increasing tree size that we observed in the majority of species may be a homeostatic mechanism that partially compensates for decreased hydraulic conductance as trees grow in height.

  2. Leaf area compounds height-related hydraulic costs of water transport in Oregon White Oak trees.

    Science.gov (United States)

    N. Phillips; B. J. Bond; N. G. McDowell; Michael G. Ryan; A. Schauer

    2003-01-01

    The ratio of leaf to sapwood area generally decreases with tree size, presumably to moderate hydraulic costs of tree height. This study assessed consequences of tree size and leaf area on water flux in Quercus garryana Dougl. ex. Hook (Oregon White Oak), a species in which leaf to sapwood area ratio increases with tree size. We tested hypotheses that...

  3. Allometric relationships predicting foliar biomass and leaf area:sapwood area ratio from tree height in five Costa Rican rain forest species.

    Science.gov (United States)

    Calvo-Alvarado, J C; McDowell, N G; Waring, R H

    2008-11-01

    We developed allometric equations to predict whole-tree leaf area (A(l)), leaf biomass (M(l)) and leaf area to sapwood area ratio (A(l):A(s)) in five rain forest tree species of Costa Rica: Pentaclethra macroloba (Willd.) Kuntze (Fabaceae/Mim), Carapa guianensis Aubl. (Meliaceae), Vochysia ferru-gi-nea Mart. (Vochysiaceae), Virola koshnii Warb. (Myristicaceae) and Tetragastris panamensis (Engl.) Kuntze (Burseraceae). By destructive analyses (n = 11-14 trees per species), we observed strong nonlinear allometric relationships (r(2) > or = 0.9) for predicting A(l) or M(l) from stem diameters or A(s) measured at breast height. Linear relationships were less accurate. In general, A(l):A(s) at breast height increased linearly with tree height except for Penta-clethra, which showed a negative trend. All species, however, showed increased total A(l) with height. The observation that four of the five species increased in A(l):A(s) with height is consistent with hypotheses about trade--offs between morphological and anatomical adaptations that favor efficient water flow through variation in the amount of leaf area supported by sapwood and those imposed by the need to respond quickly to light gaps in the canopy.

  4. Equations for predicting diameter, height, crown width, and leaf area of San Joaquin Valley street trees

    Science.gov (United States)

    P.J. Peper; E.G. McPherson; S.M. Mori

    2001-01-01

    Although the modeling of energy-use reduction, air pollution uptake, rainfall interception, and microclimate modification associated with urban trees depends on data relating diameter at breast height (dbh) , crown height, crown diameter, and leaf area to tree age or dbh, scant information is available for common municipal tree species . I n this study , tree height ,...

  5. LEAF MICROMORPHOMETRY OF Schinus molle L. (ANARCADIACEAE IN DIFFERENT CANOPY HEIGHTS.

    Directory of Open Access Journals (Sweden)

    Marinês Ferreira Pires

    2015-03-01

    Full Text Available Leaf characterization of trees is essential for its identification and use, as well as to understand its relationships with environment. The objective of this work is to study the leaflet anatomy and leaf biometrical characteristics at different canopy heights of Schinus molle plants as a function of its environmental and physiological modifications. Leaves were collected at three different canopy heights: base, middle and upper canopy in a plantation of S. molle. Leaves were used for anatomical and biometrical analysis. For the anatomical analysis, leaves were fixed in FAA and stored in ethanol 70% and further submitted to transversal and paradermical sections. Slides were photomicrographed and image analysis was performed in UTHSCSA-Imagetool. For biometrical analysis leaf area, length, width, dry mass and specific leaf area were evaluated. The leaflets exhibited single layer epidermis, anomocytic and ciclocytic stomata, isobilateral mesophyll, subepidermal parenchyma layer in both adaxial and abaxial faces of epidermis, secretory vessels and lamellar collenchyma in midrib and leaf border. Leaf anatomy modifications occurred in cuticle and mesophyll thickness, vascular system, phloem thickness, and stomatal density in accordance with leaf canopy position. Leaves were smaller and with reduced leaf area at higher canopy positions. S. molle leaf anatomy is different from other species within Schinus genre with modifications under different environmental and physiological modifications promoted by its canopy height.

  6. Height-related changes in leaf photosynthetic traits in diverse Bornean tropical rain forest trees.

    Science.gov (United States)

    Kenzo, Tanaka; Inoue, Yuta; Yoshimura, Mitsunori; Yamashita, Megumi; Tanaka-Oda, Ayumi; Ichie, Tomoaki

    2015-01-01

    Knowledge of variations in morphophysiological leaf traits with forest height is essential for quantifying carbon and water fluxes from forest ecosystems. Here, we examined changes in leaf traits with forest height in diverse tree species and their role in environmental acclimation in a tropical rain forest in Borneo that does not experience dry spells. Height-related changes in leaf physiological and morphological traits [e.g., maximum photosynthetic rate (Amax), stomatal conductance (gs), dark respiration rate (Rd), carbon isotope ratio (δ(13)C), nitrogen (N) content, and leaf mass per area (LMA)] from understory to emergent trees were investigated in 104 species in 29 families. We found that many leaf area-based physiological traits (e.g., A(max-area), Rd, gs), N, δ(13)C, and LMA increased linearly with tree height, while leaf mass-based physiological traits (e.g., A(max-mass)) only increased slightly. These patterns differed from other biomes such as temperate and tropical dry forests, where trees usually show decreased photosynthetic capacity (e.g., A(max-area), A(max-mass)) with height. Increases in photosynthetic capacity, LMA, and δ(13)C are favored under bright and dry upper canopy conditions with higher photosynthetic productivity and drought tolerance, whereas lower R d and LMA may improve shade tolerance in lower canopy trees. Rapid recovery of leaf midday water potential to theoretical gravity potential during the night supports the idea that the majority of trees do not suffer from strong drought stress. Overall, leaf area-based photosynthetic traits were associated with tree height and the degree of leaf drought stress, even in diverse tropical rain forest trees.

  7. How should leaf area, sapwood area and stomatal conductance vary with tree height to maximize growth?

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    Buckley, Thomas N; Roberts, David W

    2006-02-01

    Conventional wisdom holds that the ratio of leaf area to sapwood area (L/S) should decline during height (H) growth to maintain hydraulic homeostasis and prevent stomatal conductance (g(s)) from declining. We contend that L/S should increase with H based on a numerical simulation, a mathematical analysis and a conceptual argument: (1) numerical simulation--a tree growth model, DESPOT (Deducing Emergent Structure and Physiology Of Trees), in which carbon (C) allocation is regulated to maximize C gain, predicts L/S should increase during most of H growth; (2) mathematical analysis--the formal criterion for optimal C allocation, applied to a simplified analytical model of whole tree carbon-water balance, predicts L/S should increase with H if leaf-level gas exchange parameters including g(s) are conserved; and (3) conceptual argument--photosynthesis is limited by several substitutable resources (chiefly nitrogen (N), water and light) and H growth increases the C cost of water transport but not necessarily of N and light capture, so if the goal is to maximize C gain or growth, allocation should shift in favor of increasing photosynthetic capacity and irradiance, rather than sustaining g(s). Although many data are consistent with the prediction that L/S should decline with H, many others are not, and we discuss possible reasons for these discrepancies.

  8. Changes in photosynthesis and leaf characteristics with tree height in five dipterocarp species in a tropical rain forest.

    Science.gov (United States)

    Kenzo, Tanaka; Ichie, Tomoaki; Watanabe, Yoko; Yoneda, Reiji; Ninomiya, Ikuo; Koike, Takayoshi

    2006-07-01

    Variations in leaf photosynthetic, morphological and biochemical properties with increasing plant height from seedlings to emergent trees were investigated in five dipterocarp species in a Malaysian tropical rain forest. Canopy openness increased significantly with tree height. Photosynthetic properties, such as photosynthetic capacity at light saturation, light compensation point, maximum rate of carboxylation and maximum rate of photosynthetic electron transport, all increased significantly with tree height. Leaf morphological and biochemical traits, such as leaf mass per area, palisade layer thickness, nitrogen concentration per unit area, chlorophyll concentration per unit dry mass and chlorophyll to nitrogen ratio, also changed significantly with tree height. Leaf properties had simple and significant relationships with tree height, with few intra- and interspecies differences. Our results therefore suggest that the photosynthetic capacity of dipterocarp trees depends on tree height, and that the trees adapt to the light environment by adjusting their leaf morphological and biochemical properties. These results should aid in developing models that can accurately estimate carbon dioxide flux and biomass production in tropical rain forests.

  9. Vertical leaf mass per area gradient of mature sugar maple reflects both height-driven increases in vascular tissue and light-driven increases in palisade layer thickness.

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    Coble, Adam P; Cavaleri, Molly A

    2017-10-01

    A key trait used in canopy and ecosystem function modeling, leaf mass per area (LMA), is influenced by changes in both leaf thickness and leaf density (LMA = Thickness × Density). In tall trees, LMA is understood to increase with height through two primary mechanisms: (i) increasing palisade layer thickness (and thus leaf thickness) in response to light and/or (ii) reduced cell expansion and intercellular air space in response to hydrostatic constraints, leading to increased leaf density. Our objective was to investigate within-canopy gradients in leaf anatomical traits in order to understand environmental factors that influence leaf morphology in a sugar maple (Acer saccharum Marshall) forest canopy. We teased apart the effects of light and height on anatomical traits by sampling at exposed and closed canopies that had different light conditions at similar heights. As expected, palisade layer thickness responded strongly to cumulative light exposure. Mesophyll porosity, however, was weakly and negatively correlated with light and height (i.e., hydrostatic gradients). Reduced mesophyll porosity was not likely caused by limitations on cell expansion; in fact, epidermal cell width increased with height. Palisade layer thickness was better related to LMA, leaf density and leaf thickness than was mesophyll porosity. Vein diameter and fraction of vascular tissue also increased with height and LMA, density and thickness, revealing that greater investment in vascular and support tissue may be a third mechanism for increased LMA with height. Overall, decreasing mesophyll porosity with height was likely due to palisade cells expanding into the available air space and also greater investments in vascular and support tissue, rather than a reduction of cell expansion due to hydrostatic constraints. Our results provide evidence that light influences both palisade layer thickness and mesophyll porosity and indicate that hydrostatic gradients influence leaf vascular and support

  10. Sapwood area - leaf area relationships for coast redwood

    OpenAIRE

    Stancioiu, P T; O'Hara, K L

    2005-01-01

    Coast redwood (Sequoia sempervirens (D. Don) Endl.) trees in different canopy strata and crown positions were sampled to develop relationships between sapwood cross-sectional area and projected leaf area. Sampling occurred during the summers of 2000 and 2001 and covered tree heights ranging from 7.7 to 45.2 m and diameters at breast height ranging from 9.4 to 92.7 cm. Foliage morphology varied greatly and was stratified into five types based on needle type (sun or shade) and twig color. A str...

  11. Height is more important than light in determining leaf morphology in a tropical forest.

    Science.gov (United States)

    Cavaleri, Molly A; Oberbauer, Steven F; Clark, David B; Clark, Deborah A; Ryan, Michael G

    2010-06-01

    Both within and between species, leaf physiological parameters are strongly related to leaf dry mass per area (LMA, g/m2), which has been found to increase from forest floor to canopy top in every forest where it has been measured. Although vertical LMA gradients in forests have historically been attributed to a direct phenotypic response to light, an increasing number of recent studies have provided evidence that water limitation in the upper canopy can constrain foliar morphological adaptations to higher light levels. We measured height, light, and LMA of all species encountered along 45 vertical canopy transects across a Costa Rican tropical rain forest. LMA was correlated with light levels in the lower canopy until approximately 18 m sample height and 22% diffuse transmittance. Height showed a remarkably linear relationship with LMA throughout the entire vertical canopy profile for all species pooled and for each functional group individually (except epiphytes), possibly through the influence of gravity on leaf water potential and turgor pressure. Models of forest function may be greatly simplified by estimating LMA-correlated leaf physiological parameters solely from foliage height profiles, which in turn can be assessed with satellite- and aircraft-based remote sensing.

  12. Climate influences the leaf area/sapwood area ratio in Scots pine.

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    Mencuccini, M; Grace, J

    1995-01-01

    We tested the hypothesis that the leaf area/sapwood area ratio in Scots pine (Pinus sylvestris L.) is influenced by site differences in water vapor pressure deficit of the air (D). Two stands of the same provenance were selected, one in western Scotland and one in eastern England, so that effects resulting from age, genetic variability, density and fertility were minimized. Compared with the Scots pine trees at the cooler and wetter site in Scotland, the trees at the warmer and drier site in England produced less leaf area per unit of conducting sapwood area both at a stem height of 1.3 m and at the base of the live crown, whereas stem permeability was similar at both sites. Also, trees at the drier site had less leaf area per unit branch cross-sectional area at the branch base than trees at the wetter site. For each site, the average values for leaf area, sapwood area and permeability were used, together with values of transpiration rates at different D, to calculate average stem water potential gradients. Changes in the leaf area/sapwood area ratio acted to maintain a similar water potential gradient in the stems of trees at both sites despite climatic differences between the sites.

  13. An evolutionary attractor model for sapwood cross section in relation to leaf area.

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    Westoby, Mark; Cornwell, William K; Falster, Daniel S

    2012-06-21

    Sapwood cross-sectional area per unit leaf area (SA:LA) is an influential trait that plants coordinate with physical environment and with other traits. We develop theory for SA:LA and also for root surface area per leaf area (RA:LA) on the premise that plants maximizing the surplus of revenue over costs should have competitive advantage. SA:LA is predicted to increase in water-relations environments that reduce photosynthetic revenue, including low soil water potential, high water vapor pressure deficit (VPD), and low atmospheric CO(2). Because sapwood has costs, SA:LA adjustment does not completely offset difficult water relations. Where sapwood costs are large, as in tall plants, optimal SA:LA may actually decline with (say) high VPD. Large soil-to-root resistance caps the benefits that can be obtained from increasing SA:LA. Where a plant can adjust water-absorbing surface area of root per leaf area (RA:LA) as well as SA:LA, optimal RA:SA is not affected by VPD, CO(2) or plant height. If selection favours increased height more so than increased revenue-minus-cost, then height is predicted to rise substantially under improved water-relations environments such as high-CO(2) atmospheres. Evolutionary-attractor theory for SA:LA and RA:LA complements models that take whole-plant conductivity per leaf area as a parameter. Copyright © 2012 Elsevier Ltd. All rights reserved.

  14. Influence of Vegetation Structure on Lidar-derived Canopy Height and Fractional Cover in Forested Riparian Buffers During Leaf-Off and Leaf-On Conditions

    Science.gov (United States)

    Wasser, Leah; Day, Rick; Chasmer, Laura; Taylor, Alan

    2013-01-01

    Estimates of canopy height (H) and fractional canopy cover (FC) derived from lidar data collected during leaf-on and leaf-off conditions are compared with field measurements from 80 forested riparian buffer plots. The purpose is to determine if existing lidar data flown in leaf-off conditions for applications such as terrain mapping can effectively estimate forested riparian buffer H and FC within a range of riparian vegetation types. Results illustrate that: 1) leaf-off and leaf-on lidar percentile estimates are similar to measured heights in all plots except those dominated by deciduous compound-leaved trees where lidar underestimates H during leaf off periods; 2) canopy height models (CHMs) underestimate H by a larger margin compared to percentile methods and are influenced by vegetation type (conifer needle, deciduous simple leaf or deciduous compound leaf) and canopy height variability, 3) lidar estimates of FC are within 10% of plot measurements during leaf-on periods, but are underestimated during leaf-off periods except in mixed and conifer plots; and 4) depth of laser pulse penetration lower in the canopy is more variable compared to top of the canopy penetration which may influence within canopy vegetation structure estimates. This study demonstrates that leaf-off lidar data can be used to estimate forested riparian buffer canopy height within diverse vegetation conditions and fractional canopy cover within mixed and conifer forests when leaf-on lidar data are not available. PMID:23382966

  15. Applicability of non-destructive substitutes for leaf area in different stands of Norway spruce (Picea abies L. Karst.) focusing on traditional forest crown measures

    Science.gov (United States)

    Laubhann, Daniel; Eckmüllner, Otto; Sterba, Hubert

    2010-01-01

    Since individual tree leaf area is an important measure for productivity as well as for site occupancy, it is of high interest in many studies about forest growth. The exact determination of leaf area is nearly impossible. Thus, a common way to get information about leaf area is to use substitutes. These substitutes are often variables which are collected in a destructive way which is not feasible for long term studies. Therefore, this study aimed at testing the applicability of using substitutes for leaf area which could be collected in a non-destructive way, namely crown surface area and crown projection area. In 8 stands of Norway spruce (Picea abies L. Karst.), divided into three age classes and two thinning treatments, a total of 156 trees were felled in order to test the relationship between leaf area and crown surface area and crown projection area, respectively. Individual tree leaf area of the felled sample trees was estimated by 3P-branch sampling with an accuracy of ±10%. Crown projection area and crown surface area were compared with other, more commonly used, but destructive predictors of leaf area, namely sapwood area at different heights on the bole. Our investigations confirmed findings of several studies that sapwood area is the most precise measure for leaf area because of the high correlation between sapwood area and the leaf area. But behind sapwood area at crown base and sapwood area at three tenth of the tree height the predictive ability of crown surface area was ranked third and even better than that of sapwood area at breast height (R2 = 0.656 compared with 0.600). Within the stands leaf area is proportional to crown surface area. Using the pooled data of all stands a mixed model approach showed that additionally to crown surface area dominant height and diameter at breast height (dbh) improved the leaf area estimates. Thus, taking dominant height and dbh into account, crown surface area can be recommended for estimating the leaf area of

  16. Applicability of non-destructive substitutes for leaf area in different stands of Norway spruce (Picea abies L. Karst.) focusing on traditional forest crown measures.

    Science.gov (United States)

    Laubhann, Daniel; Eckmüllner, Otto; Sterba, Hubert

    2010-09-30

    Since individual tree leaf area is an important measure for productivity as well as for site occupancy, it is of high interest in many studies about forest growth. The exact determination of leaf area is nearly impossible. Thus, a common way to get information about leaf area is to use substitutes. These substitutes are often variables which are collected in a destructive way which is not feasible for long term studies. Therefore, this study aimed at testing the applicability of using substitutes for leaf area which could be collected in a non-destructive way, namely crown surface area and crown projection area. In 8 stands of Norway spruce (Picea abies L. Karst.), divided into three age classes and two thinning treatments, a total of 156 trees were felled in order to test the relationship between leaf area and crown surface area and crown projection area, respectively. Individual tree leaf area of the felled sample trees was estimated by 3P-branch sampling with an accuracy of ±10%. Crown projection area and crown surface area were compared with other, more commonly used, but destructive predictors of leaf area, namely sapwood area at different heights on the bole. Our investigations confirmed findings of several studies that sapwood area is the most precise measure for leaf area because of the high correlation between sapwood area and the leaf area. But behind sapwood area at crown base and sapwood area at three tenth of the tree height the predictive ability of crown surface area was ranked third and even better than that of sapwood area at breast height (R(2) = 0.656 compared with 0.600). Within the stands leaf area is proportional to crown surface area. Using the pooled data of all stands a mixed model approach showed that additionally to crown surface area dominant height and diameter at breast height (dbh) improved the leaf area estimates. Thus, taking dominant height and dbh into account, crown surface area can be recommended for estimating the leaf area

  17. Investigating the Alometric Relationships between Leaf Area and Some of Vegetative Characteristics in SC704 Corn Hybrid

    Directory of Open Access Journals (Sweden)

    E Zeinali

    2016-10-01

    Full Text Available Introduction Since the leaves are the main source of production of photosynthetic substances in plants, dry matter production and crop yield potential is largely dependent on the leaf surface, and many environmental changes affect growth and yield through changes in leaf area. Hence, green leaf area per plant and leaf area index is measured in almost all studies of crop physiology to understand the mechanism of yield alteration. However, measurement of leaf area compared with the other traits such as plant height and total plant dry weight is very difficult, need to precision instruments and spend more time and cost. Therefore, according to the allometric relationships in plants, extensive studies were done to find the relationship between leaf area and the other plant traits that their measurement is easier, faster and cheaper, and does not require expensive equipment. Using these relationships will be used to estimate plant leaf area with acceptable accuracy without measuring. Plant traits that have high correlation with leaf area and usually use to estimate the plant leaf area are the number of leaves or nodes per main stem, plant height, leaf dry weight and dry weight of vegetative parts of the plant. Allometric equations was used successfully to calculate leaf area for various crops such as cotton, wheat, chickpea, faba bean, peanuts, soybean and sweet sorghum. This study was conducted to obtain the allometric relationships between green leaf area (cm2 per plant with number of leaves or nodes per main stem, plant height, green leaf dry weight and dry weight of vegetative parts of the plant (gram per plant, and investigating the effect of plant density and planting date on these relationships in SC704 corn (Zea mays L. hybrid. Materials and Methods This study was conducted at Gorgan University of Agricultural Sciences and Natural Resources farm located at latitude 36 o 51’ N, longitude 54 o27’ E and altitude of 13 meters above sea level

  18. Sapwood area as an estimator of leaf area and foliar weight in cherrybark oak and green ash

    Science.gov (United States)

    James S. Meadows; John D. Hodges

    2002-01-01

    The relationships between foliar weight/leaf area and four stem dimensions (d.b.h., total stem cross-sectional area, total sapwood area, and current sapwood area at breast height) were investigated in two important bottomland tree species of the Southern United States, cherrybark oak (Quercus falcata var. pagodifolia ...

  19. Leaf-IT: An Android application for measuring leaf area.

    Science.gov (United States)

    Schrader, Julian; Pillar, Giso; Kreft, Holger

    2017-11-01

    The use of plant functional traits has become increasingly popular in ecological studies because plant functional traits help to understand key ecological processes in plant species and communities. This also includes changes in diversity, inter- and intraspecific interactions, and relationships of species at different spatiotemporal scales. Leaf traits are among the most important traits as they describe key dimensions of a plant's life history strategy. Further, leaf area is a key parameter with relevance for other traits such as specific leaf area, which in turn correlates with leaf chemical composition, photosynthetic rate, leaf longevity, and carbon investment. Measuring leaf area usually involves the use of scanners and commercial software and can be difficult under field conditions. We present Leaf-IT, a new smartphone application for measuring leaf area and other trait-related areas. Leaf-IT is free, designed for scientific purposes, and runs on Android 4 or higher. We tested the precision and accuracy using objects with standardized area and compared the area measurements of real leaves with the well-established, commercial software WinFOLIA using the Altman-Bland method. Area measurements of standardized objects show that Leaf-IT measures area with high accuracy and precision. Area measurements with Leaf-IT of real leaves are comparable to those of WinFOLIA. Leaf-IT is an easy-to-use application running on a wide range of smartphones. That increases the portability and use of Leaf-IT and makes it possible to measure leaf area under field conditions typical for remote locations. Its high accuracy and precision are similar to WinFOLIA. Currently, its main limitation is margin detection of damaged leaves or complex leaf morphologies.

  20. Leaf area and tree increment dynamics of even-aged and multiaged lodgepole pine stands in Montana

    Science.gov (United States)

    Cassandra L. Kollenberg; Kevin L. O' Hara

    1999-01-01

    Age structure and distribution of leaf area index (LAI) of even and multiaged lodgepole pine (Pinus contorta var. latifolia Engelm.) stands were examined on three study areas in western and central Montana. Projected leaf area was determined based on a relationship with sapwood cross-sectional area at breast height. Stand structure and LAI varied considerably between...

  1. Predictive equations for dimensions and leaf area of coastal Southern California street trees

    Science.gov (United States)

    P.J. Peper; E.G. McPherson; S.M. Mori

    2001-01-01

    Tree height, crown height, crown width, diameter at breast height (dbh), and leaf area were measured for 16 species of commonly planted street trees in the coastal southern California city of Santa Monica, USA. The randomly sampled trees were planted from 1 to 44 years ago. Using number of years after planting or dbh as explanatory variables, mean values of dbh, tree...

  2. Easy Leaf Area: Automated digital image analysis for rapid and accurate measurement of leaf area.

    Science.gov (United States)

    Easlon, Hsien Ming; Bloom, Arnold J

    2014-07-01

    Measurement of leaf areas from digital photographs has traditionally required significant user input unless backgrounds are carefully masked. Easy Leaf Area was developed to batch process hundreds of Arabidopsis rosette images in minutes, removing background artifacts and saving results to a spreadsheet-ready CSV file. • Easy Leaf Area uses the color ratios of each pixel to distinguish leaves and calibration areas from their background and compares leaf pixel counts to a red calibration area to eliminate the need for camera distance calculations or manual ruler scale measurement that other software methods typically require. Leaf areas estimated by this software from images taken with a camera phone were more accurate than ImageJ estimates from flatbed scanner images. • Easy Leaf Area provides an easy-to-use method for rapid measurement of leaf area and nondestructive estimation of canopy area from digital images.

  3. Measurement of Leaf Mass and Leaf Area of Oaks In A Mediterranean-climate Region For Biogenic Emission Estimation

    Science.gov (United States)

    Karlik, J.

    Given the key role played by biogenic volatile organic compounds (BVOC) in tro- pospheric chemistry and regional air quality, it is critical to generate accurate BVOC emission inventories. Because several oak species have high BVOC emission rates, and oak trees are often of large stature with corresponding large leaf masses, oaks may be the most important genus of woody plants for BVOC emissions modeling in the natural landscapes of Mediterranean-climate regions. In California, BVOC emis- sions from oaks may mix with anthropogenic emissions from urban areas, leading to elevated levels of ozone. Data for leaf mass and leaf area for a stand of native blue oaks (Quercus douglasii) were obtained through harvest and leaf removal from 14 trees lo- cated in the Sierra Nevada foothills of central California. Trees ranged in height from 4.2 to 9.9 m, with trunk diameters at breast height of 14 to 85 cm. Mean leaf mass density was 730 g m-2 for the trees and had an overall value of 310 g m-2 for the site. Consideration of the surrounding grassland devoid of trees resulted in a value of about 150 g m-2, less than half of reported values for eastern U.S. oak woodlands, but close to a reported value for oaks found in St. Quercio, Italy. The mean value for leaf area index (LAI) for the trees at this site was 4.4 m2 m-2. LAI for the site was 1.8 m2 m-2, but this value was appropriate for the oak grove only; including the surrounding open grassland resulted in an overall LAI value of 0.9 m2 m-2 or less. A volumetric method worked well for estimating the leaf mass of the oak trees. Among allometric relationships investigated, trunk circumference, mean crown radius, and crown projec- tion were well correlated with leaf mass. Estimated emission of isoprene (mg C m-2 h-1) for the site based these leaf mass data and experimentally determined emission rate was similar to that reported for a Mediterranean oak woodland in France.

  4. Easy Leaf Area: Automated Digital Image Analysis for Rapid and Accurate Measurement of Leaf Area

    Directory of Open Access Journals (Sweden)

    Hsien Ming Easlon

    2014-07-01

    Full Text Available Premise of the study: Measurement of leaf areas from digital photographs has traditionally required significant user input unless backgrounds are carefully masked. Easy Leaf Area was developed to batch process hundreds of Arabidopsis rosette images in minutes, removing background artifacts and saving results to a spreadsheet-ready CSV file. Methods and Results: Easy Leaf Area uses the color ratios of each pixel to distinguish leaves and calibration areas from their background and compares leaf pixel counts to a red calibration area to eliminate the need for camera distance calculations or manual ruler scale measurement that other software methods typically require. Leaf areas estimated by this software from images taken with a camera phone were more accurate than ImageJ estimates from flatbed scanner images. Conclusions: Easy Leaf Area provides an easy-to-use method for rapid measurement of leaf area and nondestructive estimation of canopy area from digital images.

  5. Remote Sensing of Leaf Area Index from LiDAR Height Percentile Metrics and Comparison with MODIS Product in a Selectively Logged Tropical Forest Area in Eastern Amazonia

    Directory of Open Access Journals (Sweden)

    Yonghua Qu

    2018-06-01

    Full Text Available Leaf area index (LAI is an important parameter to describe the capacity of forests to intercept light and thus affects the microclimate and photosynthetic capacity of canopies. In general, tropical forests have a higher leaf area index and it is a challenge to estimate LAI in a forest with a very dense canopy. In this study, it is assumed that the traditional Light Detection and Ranging (LiDAR-derived fractional vegetation cover (fCover has weak relationship with leaf area index in a dense forest. We propose a partial least squares (PLS regression model using the height percentile metrics derived from airborne LiDAR data to estimate the LAI of a dense forest. Ground inventory and airborne LiDAR data collected in a selectively logged tropical forest area in Eastern Amazonia are used to map LAI from the plot level to the landscape scale. The results indicate that the fCover, derived from the first return or the last return, has no significant correlations with the ground-based LAI. The PLS model evaluated by the leave-one-out validation shows that the estimated LAI is significantly correlated with the ground-based LAI with an R2 of 0.58 and a root mean square error (RMSE of 1.13. A data comparison indicates that the Moderate Resolution Imaging Spectrometer (MODIS LAI underestimates the landscape-level LAI by about 22%. The MODIS quality control data show that in the selected tile, the cloud state is not the primary factor affecting the MODIS LAI performance; rather, the LAI from the main radiative transfer (RT algorithm contributes much to the underestimation of the LAI in the tropical forest. In addition, the results show that the LiDAR-based LAI has a better response to the logging activities than the MODIS-based LAI, and that the leaf area reduction caused by logging is about 13%. In contrast, the MODIS-based LAI exhibits no apparent spatial correlation with the LiDAR-based LAI. It is suggested that the main algorithm of MODIS should be

  6. Restoration thinning and influence of tree size and leaf area to sapwood area ratio on water relations of Pinus ponderosa.

    Science.gov (United States)

    Simonin, K; Kolb, T E; Montes-Helu, M; Koch, G W

    2006-04-01

    Ponderosa pine (Pinus ponderosa Dougl. ex P. Laws) forest stand density has increased significantly over the last century (Covington et al. 1997). To understand the effect of increased intraspecific competition, tree size (height and diameter at breast height (DBH)) and leaf area to sapwood area ratio (A(L):A(S)) on water relations, we compared hydraulic conductance from soil to leaf (kl) and transpiration per unit leaf area (Q(L)) of ponderosa pine trees in an unthinned plot to trees in a thinned plot in the first and second years after thinning in a dense Arizona forest. We calculated kl and Q(L) based on whole- tree sap flux measured with heat dissipation sensors. Thinning increased tree predawn water potential within two weeks of treatment. Effects of thinning on kl and Q(L) depended on DBH, A(L):A(S) and drought severity. During severe drought in the first growing season after thinning, kl and Q(L) of trees with low A(L):A(S) (160-250 mm DBH; 9-11 m height) were lower in the thinned plot than the unthinned plot, suggesting a reduction in stomatal conductance (g(s)) or reduced sapwood specific conductivity (K(S)), or both, in response to thinning. In contrast kl and Q(L) were similar in the thinned plot and unthinned plot for trees with high A(L):A(S) (260-360 mm DBH; 13-16 m height). During non-drought periods, kl and Q(L) were greater in the thinned plot than in the unthinned plot for all but the largest trees. Contrary to previous studies of ponderosa pine, A(L):A(S) was positively correlated with tree height and DBH. Furthermore, kl and Q(L) showed a weak negative correlation with tree height and a strong negative correlation with A(S) and thus A(L):A(S) in both the thinned and unthinned plots, suggesting that trees with high A(L):A(S) had lower g(s). Our results highlight the important influence of stand competitive environment on tree-size-related variation in A(L):A(S) and the roles of A(L):A(S) and drought on whole-tree water relations in response to

  7. Total belowground carbon flux in subalpine forests is related to leaf area index, soil nitrogen, and tree height

    Science.gov (United States)

    Berryman, Erin Michele; Ryan, Michael G.; Bradford, John B.; Hawbaker, Todd J.; Birdsey, R.

    2016-01-01

    In forests, total belowground carbon (C) flux (TBCF) is a large component of the C budget and represents a critical pathway for delivery of plant C to soil. Reducing uncertainty around regional estimates of forest C cycling may be aided by incorporating knowledge of controls over soil respiration and TBCF. Photosynthesis, and presumably TBCF, declines with advancing tree size and age, and photosynthesis increases yet C partitioning to TBCF decreases in response to high soil fertility. We hypothesized that these causal relationships would result in predictable patterns of TBCF, and partitioning of C to TBCF, with natural variability in leaf area index (LAI), soil nitrogen (N), and tree height in subalpine forests in the Rocky Mountains, USA. Using three consecutive years of soil respiration data collected from 22 0.38-ha locations across three 1-km2 subalpine forested landscapes, we tested three hypotheses: (1) annual soil respiration and TBCF will show a hump-shaped relationship with LAI; (2) variability in TBCF unexplained by LAI will be related to soil nitrogen (N); and (3) partitioning of C to TBCF (relative to woody growth) will decline with increasing soil N and tree height. We found partial support for Hypothesis 1 and full support for Hypotheses 2 and 3. TBCF, but not soil respiration, was explained by LAI and soil N patterns (r2 = 0.49), and the ratio of annual TBCF to TBCF plus aboveground net primary productivity (ANPP) was related to soil N and tree height (r2 = 0.72). Thus, forest C partitioning to TBCF can vary even within the same forest type and region, and approaches that assume a constant fraction of TBCF relative to ANPP may be missing some of this variability. These relationships can aid with estimates of forest soil respiration and TBCF across landscapes, using spatially explicit forest data such as national inventories or remotely sensed data products.

  8. Leaf area index from litter collection: impact of specific leaf area variability within a beech stand

    Energy Technology Data Exchange (ETDEWEB)

    Bouriaud, O. [Inst. National de la Recherche Agronomique, Centre de Recherches Forestieres de Nancy, Champenoux (France); Soudani, K. [Univ. Paris-Sud XI, Dept. d' Ecophysiologie Vegetale, Lab. Ecologie Systematique et Evolution, Orsay Cedex (France); Breda, N. [Inst. National de la Recherche Agronomique, Centre de Recherches Forestieres de Nancy, Champenoux (France)

    2003-06-01

    Litter fall collection is a direct method widely used to estimate leaf area index (LAI) in broad-leaved forest stands. Indirect measurements using radiation transmittance and gap fraction theory are often compared and calibrated against litter fall, which is considered as a reference method, but few studies address the question of litter specific leaf area (SLA) measurement and variability. SLA (leaf area per unit of dry weight, m{sup 2}{center_dot}g{sup -1}) is used to convert dry leaf litter biomass (g .m{sup -}2) into leaf area per ground unit area (m{sup 2}{center_dot}m{sup -2}). We paid special attention to this parameter in two young beech stands (dense and thinned) in northeastern France. The variability of both canopy (closure, LAI) and site conditions (soil properties, vegetation) was investigated as potential contributing factors to beech SLA variability. A systematic description of soil and floristic composition was performed and three types of soil were identified. Ellenberg's indicator values were averaged for each plot to assess nitrogen soil content. SLA of beech litter was measured three times during the fall in 23 plots in the stands (40 ha). Litter was collected bimonthly in square-shaped traps (0.5 m{sup 2}) and dried. Before drying, 30 leaves per plot and for each date were sampled, and leaf length, width, and area were measured with the help of a LI-COR areameter. SLA was calculated as the ratio of cumulated leaf area to total dry weight of the 30 leaves. Leaves characteristics per plot were averaged for the three dates of litter collection. Plant area index (PAI), estimated using the LAI-2000 plant canopy analyser and considering only the upper three rings, ranged from 2.9 to 8.1. Specific leaf area of beech litter was also highly different from one plot to the other, ranging from 150 to 320 cm{sup 2}{center_dot}g{sup -1}. Nevertheless, no relationship was found between SLA and stand canopy closure or PAI On the contrary, a significant

  9. Leaf area index from litter collection: impact of specific leaf area variability within a beech stand

    International Nuclear Information System (INIS)

    Bouriaud, O.; Soudani, K.; Breda, N.

    2003-01-01

    Litter fall collection is a direct method widely used to estimate leaf area index (LAI) in broad-leaved forest stands. Indirect measurements using radiation transmittance and gap fraction theory are often compared and calibrated against litter fall, which is considered as a reference method, but few studies address the question of litter specific leaf area (SLA) measurement and variability. SLA (leaf area per unit of dry weight, m 2 ·g -1 ) is used to convert dry leaf litter biomass (g .m - 2) into leaf area per ground unit area (m 2 ·m -2 ). We paid special attention to this parameter in two young beech stands (dense and thinned) in northeastern France. The variability of both canopy (closure, LAI) and site conditions (soil properties, vegetation) was investigated as potential contributing factors to beech SLA variability. A systematic description of soil and floristic composition was performed and three types of soil were identified. Ellenberg's indicator values were averaged for each plot to assess nitrogen soil content. SLA of beech litter was measured three times during the fall in 23 plots in the stands (40 ha). Litter was collected bimonthly in square-shaped traps (0.5 m 2 ) and dried. Before drying, 30 leaves per plot and for each date were sampled, and leaf length, width, and area were measured with the help of a LI-COR areameter. SLA was calculated as the ratio of cumulated leaf area to total dry weight of the 30 leaves. Leaves characteristics per plot were averaged for the three dates of litter collection. Plant area index (PAI), estimated using the LAI-2000 plant canopy analyser and considering only the upper three rings, ranged from 2.9 to 8.1. Specific leaf area of beech litter was also highly different from one plot to the other, ranging from 150 to 320 cm 2 ·g -1 . Nevertheless, no relationship was found between SLA and stand canopy closure or PAI On the contrary, a significant relationship between SLA and soil properties was observed. Both SLA

  10. Interaction between sapwood and foliage area in alpine ash (Eucalyptus delegatensis) trees of different heights.

    Science.gov (United States)

    Mokany, Karel; McMurtrie, Ross E; Atwell, Brian J; Keith, Heather

    2003-10-01

    In native stands of Eucalyptus delegatensis R. T. Baker, sapwood area (As) to foliage area (Af) ratios (As:Af) decreased as tree height increased, contradicting the common interpretation of the Pipe Model Theory as well as the generally observed trend of increasing As:Af ratios with tree height. To clarify this relationship, we estimated sapwood hydraulic conductivity theoretically based on measurements of sapwood vessel diameters and Poiseuille's law for fluid flow through pipes. Despite the observed decrease in As:Af ratios with tree height, leaf specific conductivity increased with total tree height, largely as a result of an increase in the specific conductivity of sapwood. This observation supports the proposition that the stem's ability to supply foliage with water must increase as trees grow taller, to compensate for the increased hydraulic path length. The results presented here highlight the importance of measuring sapwood hydraulic conductivity in analyses of sapwood-foliage interactions, and suggest that measurements of sapwood hydraulic conductivity may help to resolve conflicting observations of how As:Af ratios change as trees grow taller.

  11. Leaf Area Estimation Models for Ginger ( Zingibere officinale Rosc ...

    African Journals Online (AJOL)

    The study was carried out to develop leaf area estimation models for three cultivars (37/79, 38/79 and 180/73) and four accessions (29/86, 30/86, 47/86 and 52/86) of ginger. Significant variations were observed among the tested genotypes in leaf length (L), leaf width (W) and actual leaf area (ALA). Leaf area was highly ...

  12. Rapid, high-resolution measurement of leaf area and leaf orientation using terrestrial LiDAR scanning data

    International Nuclear Information System (INIS)

    Bailey, Brian N; Mahaffee, Walter F

    2017-01-01

    The rapid evolution of high performance computing technology has allowed for the development of extremely detailed models of the urban and natural environment. Although models can now represent sub-meter-scale variability in environmental geometry, model users are often unable to specify the geometry of real domains at this scale given available measurements. An emerging technology in this field has been the use of terrestrial LiDAR scanning data to rapidly measure the three-dimensional geometry of trees, such as the distribution of leaf area. However, current LiDAR methods suffer from the limitation that they require detailed knowledge of leaf orientation in order to translate projected leaf area into actual leaf area. Common methods for measuring leaf orientation are often tedious or inaccurate, which places constraints on the LiDAR measurement technique. This work presents a new method to simultaneously measure leaf orientation and leaf area within an arbitrarily defined volume using terrestrial LiDAR data. The novelty of the method lies in the direct measurement of the fraction of projected leaf area G from the LiDAR data which is required to relate projected leaf area to total leaf area, and in the new way in which radiation transfer theory is used to calculate leaf area from the LiDAR data. The method was validated by comparing LiDAR-measured leaf area to (1) ‘synthetic’ or computer-generated LiDAR data where the exact area was known, and (2) direct measurements of leaf area in the field using destructive sampling. Overall, agreement between the LiDAR and reference measurements was very good, showing a normalized root-mean-squared-error of about 15% for the synthetic tests, and 13% in the field. (paper)

  13. Effect of Plant Growth Regulators on Leaf Number, Leaf Area and Leaf Dry Matter in Grape

    Directory of Open Access Journals (Sweden)

    Zahoor Ahmad BHAT

    2011-03-01

    Full Text Available Influence of phenylureas (CPPU and brassinosteriod (BR along with GA (gibberellic acid were studied on seedless grape vegetative characteristics like leaf number, leaf area and leaf dry matter. Growth regulators were sprayed on the vines either once (7 days after fruit set or 15 days after fruit set or twice (7+15 days after fruit set. CPPU 2 ppm+BR 0.4 ppm+GA 25 ppm produced maximum number of leaves (18.78 while as untreated vines produced least leaf number (16.22 per shoot. Maximum leaf area (129.70 cm2 and dry matter content (26.51% was obtained with higher CPPU (3 ppm and BR (0.4 ppm combination along with GA 25 ppm. Plant growth regulators whether naturally derived or synthetic are used to improve the productivity and quality of grapes. The relatively high value of grapes justifies more expensive inputs. A relatively small improvement in yield or fruit quality can justify the field application of a very costly product. Application of new generation growth regulators like brassinosteroids and phenylureas like CPPU have been reported to increase the leaf number as well as leaf area and dry matter thereby indirectly influencing the fruit yield and quality in grapes.

  14. Neighbor and Height Effects on Crown Properties Associated with the Uniform-Stress Principle of Stem Formation

    Directory of Open Access Journals (Sweden)

    Thomas J. Dean

    2018-06-01

    Full Text Available According to the uniform-stress principle of stem formation, the amount of leaf area a tree carries and the leverage it exerts on the stem determine the stem dimensions. Within an even-aged monoculture, the leaf area per tree and the leverage placed on the stem are functions of tree density and tree height. The uniform-stress principle presents the means to translate density effects on crown characteristics into stem dimensions and total standing volume. This approach is truly a top-down method of simulating growth tree and stand growth because leaf area and other crown properties must be determined before stem size and taper can be calculated. Each crown property influences either the sail area or the leverage placed on the stem, but the degree to which a specific crown property affects these parameters changes with stand density and height. Leverage is the more complicated of the two variables, being a function of the height to the base of the live crown and the vertical distribution of leaf area. The purpose of this brief review is to summarize the effects of stand density on the height to the base of the live tree and the vertical distribution of leaf area and the various ways these variables have been quantified.

  15. A non-destructive method for estimating onion leaf area

    Directory of Open Access Journals (Sweden)

    Córcoles J.I.

    2015-06-01

    Full Text Available Leaf area is one of the most important parameters for characterizing crop growth and development, and its measurement is useful for examining the effects of agronomic management on crop production. It is related to interception of radiation, photosynthesis, biomass accumulation, transpiration and gas exchange in crop canopies. Several direct and indirect methods have been developed for determining leaf area. The aim of this study is to develop an indirect method, based on the use of a mathematical model, to compute leaf area in an onion crop using non-destructive measurements with the condition that the model must be practical and useful as a Decision Support System tool to improve crop management. A field experiment was conducted in a 4.75 ha commercial onion plot irrigated with a centre pivot system in Aguas Nuevas (Albacete, Spain, during the 2010 irrigation season. To determine onion crop leaf area in the laboratory, the crop was sampled on four occasions between 15 June and 15 September. At each sampling event, eight experimental plots of 1 m2 were used and the leaf area for individual leaves was computed using two indirect methods, one based on the use of an automated infrared imaging system, LI-COR-3100C, and the other using a digital scanner EPSON GT-8000, obtaining several images that were processed using Image J v 1.43 software. A total of 1146 leaves were used. Before measuring the leaf area, 25 parameters related to leaf length and width were determined for each leaf. The combined application of principal components analysis and cluster analysis for grouping leaf parameters was used to reduce the number of variables from 25 to 12. The parameter derived from the product of the total leaf length (L and the leaf diameter at a distance of 25% of the total leaf length (A25 gave the best results for estimating leaf area using a simple linear regression model. The model obtained was useful for computing leaf area using a non

  16. Relationships of leaf dark respiration to leaf nitrogen, specific leaf area and leaf life-span: a test across biomes and functional groups.

    Science.gov (United States)

    Reich, Peter B; Walters, Michael B; Ellsworth, David S; Vose, James M; Volin, John C; Gresham, Charles; Bowman, William D

    1998-05-01

    Based on prior evidence of coordinated multiple leaf trait scaling, we hypothesized that variation among species in leaf dark respiration rate (R d ) should scale with variation in traits such as leaf nitrogen (N), leaf life-span, specific leaf area (SLA), and net photosynthetic capacity (A max ). However, it is not known whether such scaling, if it exists, is similar among disparate biomes and plant functional types. We tested this idea by examining the interspecific relationships between R d measured at a standard temperature and leaf life-span, N, SLA and A max for 69 species from four functional groups (forbs, broad-leafed trees and shrubs, and needle-leafed conifers) in six biomes traversing the Americas: alpine tundra/subalpine forest, Colorado; cold temperate forest/grassland, Wisconsin; cool temperate forest, North Carolina; desert/shrubland, New Mexico; subtropical forest, South Carolina; and tropical rain forest, Amazonas, Venezuela. Area-based R d was positively related to area-based leaf N within functional groups and for all species pooled, but not when comparing among species within any site. At all sites, mass-based R d (R d-mass ) decreased sharply with increasing leaf life-span and was positively related to SLA and mass-based A max and leaf N (leaf N mass ). These intra-biome relationships were similar in shape and slope among sites, where in each case we compared species belonging to different plant functional groups. Significant R d-mass -N mass relationships were observed in all functional groups (pooled across sites), but the relationships differed, with higher R d at any given leaf N in functional groups (such as forbs) with higher SLA and shorter leaf life-span. Regardless of biome or functional group, R d-mass was well predicted by all combinations of leaf life-span, N mass and/or SLA (r 2 ≥ 0.79, P morphological, chemical and metabolic traits.

  17. Estimating leaf area and leaf biomass of open-grown deciduous urban trees

    Science.gov (United States)

    David J. Nowak

    1996-01-01

    Logarithmic regression equations were developed to predict leaf area and leaf biomass for open-grown deciduous urban trees based on stem diameter and crown parameters. Equations based on crown parameters produced more reliable estimates. The equations can be used to help quantify forest structure and functions, particularly in urbanizing and urban/suburban areas.

  18. Leaf area prediction models for Tsuga canadensis in Maine

    Science.gov (United States)

    Laura S. Kenefic; R.S. Seymour

    1999-01-01

    Tsuga canadensis (L.) Carr. (eastern hemlock) is a common species throughout the Acadian forest. Studies of leaf area and growth efficiency in this forest type have been limited by the lack of equations to predict leaf area of this species. We found that sapwood area was an effective leaf area surrogate in T. canadensis, though...

  19. Estimation of leaf area in tropical maize

    NARCIS (Netherlands)

    Elings, A.

    2000-01-01

    Leaf area development of six tropical maize cultivars grown in 1995 and 1996 in several tropical environments in Mexico (both favourable and moisture-and N-limited) was observed and analysed. First, the validity of a bell-shaped curve describing the area of individual leaves as a function of leaf

  20. Tree Diversity Enhances Stand Carbon Storage but Not Leaf Area in a Subtropical Forest.

    Science.gov (United States)

    Castro-Izaguirre, Nadia; Chi, Xiulian; Baruffol, Martin; Tang, Zhiyao; Ma, Keping; Schmid, Bernhard; Niklaus, Pascal A

    2016-01-01

    Research about biodiversity-productivity relationships has focused on herbaceous ecosystems, with results from tree field studies only recently beginning to emerge. Also, the latter are concentrated largely in the temperate zone. Tree species diversity generally is much higher in subtropical and tropical than in temperate or boreal forests, with reasons not fully understood. Niche overlap and thus complementarity in the use of resources that support productivity may be lower in forests than in herbaceous ecosystems, suggesting weaker productivity responses to diversity change in forests. We studied stand basal area, vertical structure, leaf area, and their relationship with tree species richness in a subtropical forest in south-east China. Permanent forest plots of 30 x 30 m were selected to span largely independent gradients in tree species richness and secondary successional age. Plots with higher tree species richness had a higher stand basal area. Also, stand basal area increases over a 4-year census interval were larger at high than at low diversity. These effects translated into increased carbon stocks in aboveground phytomass (estimated using allometric equations). A higher variability in tree height in more diverse plots suggested that these effects were facilitated by denser canopy packing due to architectural complementarity between species. In contrast, leaf area was not or even negatively affected by tree diversity, indicating a decoupling of carbon accumulation from leaf area. Alternatively, the same community leaf area might have assimilated more C per time interval in more than in less diverse plots because of differences in leaf turnover and productivity or because of differences in the display of leaves in vertical and horizontal space. Overall, our study suggests that in species-rich forests niche-based processes support a positive diversity-productivity relationship and that this translates into increased carbon storage in long-lived woody

  1. Influences of radiation and leaf area vertical distribution on the growth of Chinese fir young plantation with different densities

    International Nuclear Information System (INIS)

    Wang Lili

    1990-01-01

    A study on the radiation and leaf area vertical distribution in relation to the growth of 8-year-old Chinese fir plantations of 5 densities was conducted. The leaf area vertical distribution and LAI were closely related to stem density. The crown form varies from conic to cylindric with the increase of stem density. The LAI rises at first and then declines with the increase of density. The extinction of radiation sharpened when the crown density increased. The extinction leveled at the depth of 3/4 forest heights from the tops of forest canopies. Calculating the extinction coefficients by means of accumulated leaf area index separately for each crown layer can minimize the errors caused by the irregularity of leaf distribution. Four indices, i.e., absorption of radiation, LAI,biomass of individual tree and averaged annual increment of biomass were used to have a comprehensive evaluation on the growth of Chinese fir of 5 densities. The results showed that the plantation with a stem density of 2m × 1 m was the best one among the 5 young plantations

  2. Specific leaf area estimation from leaf and canopy reflectance through optimization and validation of vegetation indices

    NARCIS (Netherlands)

    Ali, A.M.; Darvishzadeh, R.; Skidmore, A.K.; van Duren, I.C.

    2017-01-01

    Specific leaf area (SLA), which is defined as the leaf area per unit of dry leaf mass is an important component when assessing functional diversity and plays a key role in ecosystem modeling, linking plant carbon and water cycles as well as quantifying plant physiological processes. However, studies

  3. An evolutionary perspective on leaf economics : Phylogenetics of leaf mass per area in vascular plants

    NARCIS (Netherlands)

    Flores, Olivier; Garnier, Eric; Wright, Ian J.; Reich, Peter B.; Pierce, Simon; Diaz, Sandra; Pakeman, Robin J.; Rusch, Graciela M.; Bernard-Verdier, Maud; Testi, Baptiste; Bakker, Jan P.; Bekker, Renee M.; Cerabolini, Bruno E. L.; Ceriani, Roberta M.; Cornu, Guillaume; Cruz, Pablo; Delcamp, Matthieu; Dolezal, Jiri; Eriksson, Ove; Fayolle, Adeline; Freitas, Helena; Golodets, Carly; Gourlet-Fleury, Sylvie; Hodgson, John G.; Brusa, Guido; Kleyer, Michael; Kunzmann, Dieter; Lavorel, Sandra; Papanastasis, Vasilios P.; Perez-Harguindeguy, Natalia; Vendramini, Fernanda; Weiher, Evan

    In plant leaves, resource use follows a trade-off between rapid resource capture and conservative storage. This "worldwide leaf economics spectrum" consists of a suite of intercorrelated leaf traits, among which leaf mass per area, LMA, is one of the most fundamental as it indicates the cost of leaf

  4. Leaf area estimation of cassava from linear dimensions

    Directory of Open Access Journals (Sweden)

    SAMARA ZANETTI

    2017-08-01

    Full Text Available ABSTRACT The objective of this study was to determine predictor models of leaf area of cassava from linear leaf measurements. The experiment was carried out in greenhouse in the municipality of Botucatu, São Paulo state, Brazil. The stem cuttings with 5-7 nodes of the cultivar IAC 576-70 were planted in boxes filled with about 320 liters of soil, keeping soil moisture at field capacity, monitored by puncturing tensiometers. At 80 days after planting, 140 leaves were randomly collected from the top, middle third and base of cassava plants. We evaluated the length and width of the central lobe of leaves, number of lobes and leaf area. The measurements of leaf areas were correlated with the length and width of the central lobe and the number of lobes of the leaves, and adjusted to polynomial and multiple regression models. The linear function that used the length of the central lobe LA = -69.91114 + 15.06462L and linear multiple functions LA = -69.9188 + 15.5102L + 0.0197726K - 0.0768998J or LA = -69.9346 + 15.0106L + 0.188931K - 0.0264323H are suitable models to estimate leaf area of cassava cultivar IAC 576-70.

  5. Simple models for predicting leaf area of mango (Mangifera indica L.

    Directory of Open Access Journals (Sweden)

    Maryam Ghoreishi

    2012-01-01

    Full Text Available Mango (Mangifera indica L., one of the most popular tropical fruits, is cultivated in a considerable part of southern Iran. Leaf area is a valuable parameter in mango research, especially plant physiological and nutrition field. Most of available methods for estimating plant leaf area are difficult to apply, expensive and destructive which could in turn destroy the canopy and consequently make it difficult to perform further tests on the same plant. Therefore, a non-destructive method which is simple, inexpensive, and could yield an accurate estimation of leaf area will be a great benefit to researchers. A regression analysis was performed in order to determine the relationship between the leaf area and leaf width, leaf length, dry and fresh weight. For this purpose 50 mango seedlings of local selections were randomly took from a nursery in the Hormozgan province, and different parts of plants were separated in laboratory. Leaf area was measured by different method included leaf area meter, planimeter, ruler (length and width and the fresh and dry weight of leaves were also measured. The best regression models were statistically selected using Determination Coefficient, Maximum Error, Model Efficiency, Root Mean Square Error and Coefficient of Residual Mass. Overall, based on regression equation, a satisfactory estimation of leaf area was obtained by measuring the non-destructive parameters, i.e. number of leaf per seedling, length of the longest and width of widest leaf (R2 = 0.88 and also destructive parameters, i.e. dry weight (R2 = 0.94 and fresh weight (R2= 0.94 of leaves.

  6. [Seasonal differences in the leaf hydraulic conductance of mature Acacia mangium in response to its leaf water use and photosynthesis].

    Science.gov (United States)

    Zhao, Ping; Sun, Gu-Chou; Ni, Guang-Yan; Zeng, Xiao-Ping

    2013-01-01

    In this study, measurements were made on the leaf water potential (psi1), stomatal conductance (g(s)), transpiration rate, leaf area index, and sapwood area of mature Acacia mangium, aimed to understand the relationships of the leaf hydraulic conductance (K1) with the leaf water use and photosynthetic characteristics of the A. mangium in wet season (May) and dry season (November). The ratio of sapwood area to leaf area (A(sp)/A(cl)) of the larger trees with an average height of 20 m and a diameter at breast height (DBH) of 0.26 m was 8.5% higher than that of the smaller trees with an average height of 14.5 m and a DBH of 0.19 m, suggesting that the larger trees had a higher water flux in their leaf xylem, which facilitated the water use of canopy leaf. The analysis on the vulnerability curve of the xylem showed that when the K1 decreased by 50%, the psi1 in wet season and dry season was -1.41 and -1.55 MPa, respectively, and the vulnerability of the xylem cavitation was higher in dry season than in wet season. The K1 peak value in wet season and dry season was 5.5 and 4.5 mmol x m(-2) x s(-1) x MPa(-1), and the maximum transpiration rate (T(r max)) was 3.6 and 1.8 mmol x m(-2) x s(-1), respectively. Both the K1 and T(r max), were obviously higher in wet season than in dry season. Within a day, the K1 and T(r), fluctuated many times, reflecting the reciprocated cycle of the xylem cavitation and refilling. The leaf stomatal closure occurred when the K1 declined over 50% or the psi1 reached -1.6 MPa. The g(s) would be maintained at a high level till the K1 declined over 50%. The correlation between the hydraulic conductance and photosynthetic rate was more significant in dry season than in wet season. The loss of leaf hydraulic conductance induced by seasonal change could be the causes of the decrease of T(r) and CO2 gas exchange.

  7. Community-weighted mean of leaf traits and divergence of wood traits predict aboveground biomass in secondary subtropical forests.

    Science.gov (United States)

    Ali, Arshad; Yan, En-Rong; Chang, Scott X; Cheng, Jun-Yang; Liu, Xiang-Yu

    2017-01-01

    Subtropical forests are globally important in providing ecological goods and services, but it is not clear whether functional diversity and composition can predict aboveground biomass in such forests. We hypothesized that high aboveground biomass is associated with high functional divergence (FDvar, i.e., niche complementarity) and community-weighted mean (CWM, i.e., mass ratio; communities dominated by a single plant strategy) of trait values. Structural equation modeling was employed to determine the direct and indirect effects of stand age and the residual effects of CWM and FDvar on aboveground biomass across 31 plots in secondary forests in subtropical China. The CWM model accounted for 78, 20, 6 and 2% of the variation in aboveground biomass, nitrogen concentration in young leaf, plant height and specific leaf area of young leaf, respectively. The FDvar model explained 74, 13, 7 and 0% of the variation in aboveground biomass, plant height, twig wood density and nitrogen concentration in young leaf, respectively. The variation in aboveground biomass, CWM of leaf nitrogen concentration and specific leaf area, and FDvar of plant height, twig wood density and nitrogen concentration in young leaf explained by the joint model was 86, 20, 13, 7, 2 and 0%, respectively. Stand age had a strong positive direct effect but low indirect positive effects on aboveground biomass. Aboveground biomass was negatively related to CWM of nitrogen concentration in young leaf, but positively related to CWM of specific leaf area of young leaf and plant height, and FDvar of plant height, twig wood density and nitrogen concentration in young leaf. Leaf and wood economics spectra are decoupled in regulating the functionality of forests, communities with diverse species but high nitrogen conservative and light acquisitive strategies result in high aboveground biomass, and hence, supporting both the mass ratio and niche complementarity hypotheses in secondary subtropical forests

  8. CO2 and temperature effects on leaf area production in two annual plant species

    International Nuclear Information System (INIS)

    Ackerly, D.D.; Coleman, J.S.; Morse, S.R.; Bazzaz, F.A.

    1992-01-01

    The authors studied leaf area production in two annual plant species, Abutilon theophrasti and Amaranthus retroflexus, under three day/night temperature regimes and two concentrations of carbon dioxide. The production of whole-plant leaf area during the first 30 d of growth was analyzed in terms of the leaf initiation rate, leaf expansion, individual leaf area, and, in Amaranthus, production of branch leaves. Temperature and CO 2 influenced leaf area production through effects on the rate of development, determined by the production of nodes on the main stem, and through shifts in the relationship between whole-plant leaf area and the number of main stem nodes. In Abutilon, leaf initiation rate was highest at 38 degree, but area of individual leaves was greatest at 28 degree. Total leaf area was greatly reduced at 18 degree due to slow leaf initiation rates. Elevated CO 2 concentration increased leaf initiation rate at 28 degree, resulting in an increase in whole-part leaf area. In Amaranthus, leaf initiation rate increased with temperature, and was increased by elevated CO 2 at 28 degree. Individual leaf area was greatest at 28 degree, and was increased by elevated CO 2 at 28 degree but decreased at 38 degree. Branch leaf area displayed a similar response to CO 2 , butt was greater at 38 degree. Overall, wholeplant leaf area was slightly increased at 38 degree relative to 28 degree, and elevated CO 2 levels resulted in increased leaf area at 28 degree but decreased leaf area at 38 degree

  9. Leaf density explains variation in leaf mass per area in rice between cultivars and nitrogen treatments.

    Science.gov (United States)

    Xiong, Dongliang; Wang, Dan; Liu, Xi; Peng, Shaobing; Huang, Jianliang; Li, Yong

    2016-05-01

    Leaf mass per area (LMA) is an important leaf trait; however, correlations between LMA and leaf anatomical features and photosynthesis have not been fully investigated, especially in cereal crops. The objectives of this study were (a) to investigate the correlations between LMA and leaf anatomical traits; and (b) to clarify the response of LMA to nitrogen supply and its effect on photosynthetic nitrogen use efficiency (PNUE). In the present study, 11 rice varieties were pot grown under sufficient nitrogen (SN) conditions, and four selected rice cultivars were grown under low nitrogen (LN) conditions. Leaf anatomical traits, gas exchange and leaf N content were measured. There was large variation in LMA across selected rice varieties. Regression analysis showed that the variation in LMA was more closely related to leaf density (LD) than to leaf thickness (LT). LMA was positively related to the percentage of mesophyll tissue area (%mesophyll), negatively related to the percentage of epidermis tissue area (%epidermis) and unrelated to the percentage of vascular tissue area (%vascular). The response of LMA to N supplementation was dependent on the variety and was also mainly determined by the response of LD to N. Compared with SN, photosynthesis was significantly decreased under LN, while PNUE was increased. The increase in PNUE was more critical in rice cultivars with a higher LMA under SN supply. Leaf density is the major cause of the variation in LMA across rice varieties and N treatments, and an increase in LMA under high N conditions would aggravate the decrease in PNUE. © The Author 2016. Published by Oxford University Press on behalf of the Annals of Botany Company. All rights reserved. For Permissions, please email: journals.permissions@oup.com.

  10. Non-destructive linear model for leaf area estimation in Vernonia ferruginea Less

    Directory of Open Access Journals (Sweden)

    MC. Souza

    Full Text Available Leaf area estimation is an important biometrical trait for evaluating leaf development and plant growth in field and pot experiments. We developed a non-destructive model to estimate the leaf area (LA of Vernonia ferruginea using the length (L and width (W leaf dimensions. Different combinations of linear equations were obtained from L, L2, W, W2, LW and L2W2. The linear regressions using the product of LW dimensions were more efficient to estimate the LA of V. ferruginea than models based on a single dimension (L, W, L2 or W2. Therefore, the linear regression “LA=0.463+0.676WL” provided the most accurate estimate of V. ferruginea leaf area. Validation of the selected model showed that the correlation between real measured leaf area and estimated leaf area was very high.

  11. Converging patterns of vertical variability in leaf morphology and nitrogen across seven Eucalyptus plantations in Brazil and Hawaii, USA

    Science.gov (United States)

    Adam P. Coble; Alisha Autio; Molly A. Cavaleri; Dan Binkley; Michael G. Ryan

    2014-01-01

    Across sites in Brazil and Hawaii, LMA and Nmass were strongly correlated with height and shade index, respectively, which may help simplify canopy function modeling of Eucalyptus plantations. Abstract Within tree canopies, leaf mass per area (LMA) and leaf nitrogen per unit area (Narea) commonly increase with height. Previous research has suggested that these patterns...

  12. Comparison of dwarf bamboos (Indocalamus sp.) leaf parameters to determine relationship between spatial density of plants and total leaf area per plant.

    Science.gov (United States)

    Shi, Pei-Jian; Xu, Qiang; Sandhu, Hardev S; Gielis, Johan; Ding, Yu-Long; Li, Hua-Rong; Dong, Xiao-Bo

    2015-10-01

    The relationship between spatial density and size of plants is an important topic in plant ecology. The self-thinning rule suggests a -3/2 power between average biomass and density or a -1/2 power between stand yield and density. However, the self-thinning rule based on total leaf area per plant and density of plants has been neglected presumably because of the lack of a method that can accurately estimate the total leaf area per plant. We aimed to find the relationship between spatial density of plants and total leaf area per plant. We also attempted to provide a novel model for accurately describing the leaf shape of bamboos. We proposed a simplified Gielis equation with only two parameters to describe the leaf shape of bamboos one model parameter represented the overall ratio of leaf width to leaf length. Using this method, we compared some leaf parameters (leaf shape, number of leaves per plant, ratio of total leaf weight to aboveground weight per plant, and total leaf area per plant) of four bamboo species of genus Indocalamus Nakai (I. pedalis (Keng) P.C. Keng, I. pumilus Q.H. Dai and C.F. Keng, I. barbatus McClure, and I. victorialis P.C. Keng). We also explored the possible correlation between spatial density and total leaf area per plant using log-linear regression. We found that the simplified Gielis equation fit the leaf shape of four bamboo species very well. Although all these four species belonged to the same genus, there were still significant differences in leaf shape. Significant differences also existed in leaf area per plant, ratio of leaf weight to aboveground weight per plant, and leaf length. In addition, we found that the total leaf area per plant decreased with increased spatial density. Therefore, we directly demonstrated the self-thinning rule to improve light interception.

  13. Models for leaf area estimation in dwarf pigeon pea by leaf dimensions

    Directory of Open Access Journals (Sweden)

    Rafael Vieira Pezzini

    2018-03-01

    Full Text Available ABSTRACT This study aims to determine the most suitable model to estimate the leaf area of dwarf pigeon pea in function of the leaf central leaflet dimension. Six samplings of 200 leaves were performed in the first experiment, at 36, 42, 50, 56, 64, and 72 days after emergence (DAE. In the second experiment, seven samplings of 200 leaves were performed at 29, 36, 43, 49, 57, 65, and 70 DAE, totaling 2600 leaves. The length (L and width (W of the central leaflet were measured in all leaves composed by left, central, and right leaflets, the product of length times width (LW was calculated, and the leaf area (Y – sum of left, central, and right leaflet areas was determined by digital images. Linear, power, quadratic, and cubic models of Y as function of L, W, and LW were built using data from the second experiment. Leaves from the first experiment were used to validate the models. In dwarf pigeon pea, the linear (Ŷ = – 0.4088 + 1.6669x, R2 = 0.9790 is preferable, but power (Ŷ = 1.6097x1.0065, R2 = 0.9766, quadratic (Ŷ = – 0.3625 + 1.663x + 0.00007x2, R2 = 0.9790, and cubic (Ŷ = 0.7216 + 1.522x + 0.005x2 – 5E–05x3, R2 = 0.9791 models in function of LW are also suitable to estimate the leaf area obtained by digital images. The power model (Ŷ = 5.2508x1.7868, R2 = 0.95 based on the central leaflet width is less laborious because requires only one variable, but it presents accuracy reduction.

  14. Light drives vertical gradients of leaf morphology in a sugar maple (Acer saccharum) forest.

    Science.gov (United States)

    Coble, Adam P; Cavaleri, Molly A

    2014-02-01

    Leaf mass per area (LMA, g m(-2)) is an essential trait for modeling canopy function due to its strong association with photosynthesis, respiration and leaf nitrogen. Leaf mass per area, which is influenced by both leaf thickness and density (LMA = thickness × density), generally increases from the bottom to the top of tree canopies, yet the mechanisms behind this universal pattern are not yet resolved. For decades, the light environment was assumed to be the most influential driver of within-canopy variation in LMA, yet recent evidence has shown hydrostatic gradients to be more important in upper canopy positions, especially in tall evergreen trees in temperate and tropical forests. The aim of this study was to disentangle the importance of various environmental drivers on vertical LMA gradients in a mature sugar maple (Acer saccharum Marshall) forest. We compared LMA, leaf density and leaf thickness relationships with height, light and predawn leaf water potential (ΨPre) within a closed and an exposed canopy to assess leaf morphological traits at similar heights but different light conditions. Contrary to our expectations and recent findings in the literature, we found strong evidence that light was the primary driver of vertical gradients in leaf morphology. At similar heights (13-23 m), LMA was greater within the exposed canopy than the closed canopy, and light had a stronger influence over LMA compared with ΨPre. Light also had a stronger influence over both leaf thickness and density compared with ΨPre; however, the increase in LMA within both canopy types was primarily due to increasing leaf thickness with increasing light availability. This study provides strong evidence that canopy structure and crown exposure, in addition to height, should be considered as a parameter for determining vertical patterns in LMA and modeling canopy function.

  15. Effect of Cover Crops on Vertical Distribution of Leaf Area and Dry Matter of Soybean (Glycine max L. in Competition with Weeds

    Directory of Open Access Journals (Sweden)

    seyyedeh samaneh hashemi

    2017-08-01

    Full Text Available Introduction Amount and vertical distribution of leaf area are essential for estimating interception and utilization of solar radiation of crop canopies and, consequently dry matter accumulation (Valentinuz & Tollenaar, 2006. Vertical distribution of leaf area is leaf areas per horizontal layers, based on height (Boedhram et al., 2001. Above-ground biomass is one of the central traits in functional plant ecology and growth analysis. It is a key parameter in many allometric relationships (Niklas & Enquist, 2002. The vertical biomass distribution is considered to be the main determinant of competitive strength in plant species. The presence of weeds intensifies competition for light, with the effect being determined by plant height, position of the branches, and location of the maximum leaf area. So, this experiment was conducted to study the vertical distribution of leaf area and dry matter of soybean canopy in competition with weeds and cover crops. Materials and methods This experiment was performed based on complete randomized block design with 3 replications in center of Agriculture of Joybar in 2013. Soybean was considered as main crop and soybean and Persian clover (Trifolium resupinatum L., fenugreek (Trigonella foenum–graecum L., chickling pea (Lathyrus sativus L. and winter vetch (Vicia sativa L. were the cover crops. Treatments were included cover crops (Persian clover, fenugreek, chickling pea and winter vetch and cover crop planting times (simultaneous planting of soybean with cover crops and planting cover crops three weeks after planting of soybeans and also monoculture of soybeans both in weedy and weed free conditions were considered as controls. Soybean planted in 50 cm row spacing with 5 cm between plants in the same row. Each plot was included 5 rows soybeans. Cover crop inter-seeded simultaneously in the main crop. Crops were planted on 19 May 2013 for simultaneous planting of soybean. The dominant weed species were green

  16. Joint Leaf chlorophyll and leaf area index retrieval from Landsat data using a regularized model inversion system

    Science.gov (United States)

    Leaf area index (LAI) and leaf chlorophyll (Chl) content represent key biophysical and biochemical controls on water, energy and carbon exchange processes in the terrestrial biosphere. In combination, LAI and leaf Chl content provide critical information on vegetation density, vitality and photosynt...

  17. DETERMINATION OF LEAF AREA AND PLANT COVER BY USING DIGITAL IMAGE PROCESSING

    OpenAIRE

    LŐKE, ZS.; SOÓS, G.

    2002-01-01

    The development of different crop models, and crop simulation models in particular, pointed out the importance of quantifying the exact value of the leaf area. To measure the leaf size of plants of pinnatifid form, automatic, portable leaf area meters are necessary. In most places these instruments are not available to measure the assimilatory surface size of crops with special leaf shapes. Any cheap and effective method, that could replace the application of expensive portable area meters co...

  18. Structure of forest ecosystems and leaf area index of wood plants -results of monitoring over the years 1991-1994

    International Nuclear Information System (INIS)

    Oszlanyi, J.

    1995-01-01

    Monitored characteristics and their dynamics over the last four vegetation seasons reveal the following conclusions: 1) Changes of monitored parameters (e.g. the structure of tree and shrub layer, the leaf area index) are slow, drab and insignificant at the permanent monitoring representing a major part of forest ecosystems of the area affected by the Hydroelectric power structures Gabcikovo. Despite the absence of floods, the ground water level is at a sufficient height to contact rhisosphere of wood plants and the recorded changes are in accord with growth regularities. 2) An increase of the ground water level in the upper part of the monitored territory and a partial renaturation of hydropedological conditions led to an improvement of production-ecological parameters of the area. Changes of its structure are of positive tendency, the leaf area index is stabilised at high values and somewhere even increased (in 1994 being by 70-80% higher than in 1991). 3) Localities with a permanent decrease of the ground water level (band along the old river-bed of the Danube, a dry triangle among the old river-bed of the Danube, the inlet canal and the river arm supplied by the intake structure at Dobrohost and other places) were afflicted by negative changes, locally indicating destruction of tree and shrub layers, with the leaf area index significantly reduced by 20-30%. (author). 1 tab., 5 refs [sk

  19. Effect of solution and leaf surface polarity on droplet spread area and contact angle.

    Science.gov (United States)

    Nairn, Justin J; Forster, W Alison; van Leeuwen, Rebecca M

    2016-03-01

    How much an agrochemical spray droplet spreads on a leaf surface can significantly influence efficacy. This study investigates the effect solution polarity has on droplet spreading on leaf surfaces and whether the relative leaf surface polarity, as quantified using the wetting tension dielectric (WTD) technique, influences the final spread area. Contact angles and spread areas were measured using four probe solutions on 17 species. Probe solution polarity was found to affect the measured spread area and the contact angle of the droplets on non-hairy leaves. Leaf hairs skewed the spread area measurement, preventing investigation of the influence of surface polarity on hairy leaves. WTD-measured leaf surface polarity of non-hairy leaves was found to correlate strongly with the effect of solution polarity on spread area. For non-polar leaf surfaces the spread area decreases with increasing solution polarity, for neutral surfaces polarity has no effect on spread area and for polar leaf surfaces the spread area increases with increasing solution polarity. These results attest to the use of the WTD technique as a means to quantify leaf surface polarity. © 2015 Society of Chemical Industry. © 2015 Society of Chemical Industry.

  20. Trade-offs between seed and leaf size (seed-phytomer-leaf theory): functional glue linking regenerative with life history strategies … and taxonomy with ecology?

    Science.gov (United States)

    Hodgson, John G; Santini, Bianca A; Montserrat Marti, Gabriel; Royo Pla, Ferran; Jones, Glynis; Bogaard, Amy; Charles, Mike; Font, Xavier; Ater, Mohammed; Taleb, Abdelkader; Poschlod, Peter; Hmimsa, Younes; Palmer, Carol; Wilson, Peter J; Band, Stuart R; Styring, Amy; Diffey, Charlotte; Green, Laura; Nitsch, Erika; Stroud, Elizabeth; Romo-Díez, Angel; de Torres Espuny, Lluis; Warham, Gemma

    2017-11-10

    While the 'worldwide leaf economics spectrum' (Wright IJ, Reich PB, Westoby M, et al. 2004. The worldwide leaf economics spectrum. Nature : 821-827) defines mineral nutrient relationships in plants, no unifying functional consensus links size attributes. Here, the focus is upon leaf size, a much-studied plant trait that scales positively with habitat quality and components of plant size. The objective is to show that this wide range of relationships is explicable in terms of a seed-phytomer-leaf (SPL) theoretical model defining leaf size in terms of trade-offs involving the size, growth rate and number of the building blocks (phytomers) of which the young shoot is constructed. Functional data for 2400+ species and English and Spanish vegetation surveys were used to explore interrelationships between leaf area, leaf width, canopy height, seed mass and leaf dry matter content (LDMC). Leaf area was a consistent function of canopy height, LDMC and seed mass. Additionally, size traits are partially uncoupled. First, broad laminas help confer competitive exclusion while morphologically large leaves can, through dissection, be functionally small. Secondly, leaf size scales positively with plant size but many of the largest-leaved species are of medium height with basally supported leaves. Thirdly, photosynthetic stems may represent a functionally viable alternative to 'small seeds + large leaves' in disturbed, fertile habitats and 'large seeds + small leaves' in infertile ones. Although key elements defining the juvenile growth phase remain unmeasured, our results broadly support SPL theory in that phytometer and leaf size are a product of the size of the initial shoot meristem (≅ seed mass) and the duration and quality of juvenile growth. These allometrically constrained traits combine to confer ecological specialization on individual species. Equally, they appear conservatively expressed within major taxa. Thus, 'evolutionary canalization' sensu Stebbins (Stebbins GL

  1. Leaf Area Prediction Using Three Alternative Sampling Methods for Seven Sierra Nevada Conifer Species

    Directory of Open Access Journals (Sweden)

    Dryw A. Jones

    2015-07-01

    Full Text Available Prediction of projected tree leaf area using allometric relationships with sapwood cross-sectional area is common in tree- and stand-level production studies. Measuring sapwood is difficult and often requires destructive sampling. This study tested multiple leaf area prediction models across seven diverse conifer species in the Sierra Nevada of California. The best-fit whole tree leaf area prediction model for overall simplicity, accuracy, and utility for all seven species was a nonlinear model with basal area as the primary covariate. A new non-destructive procedure was introduced to extend the branch summation approach to leaf area data collection on trees that cannot be destructively sampled. There were no significant differences between fixed effects assigned to sampling procedures, indicating that data from the tested sampling procedures can be combined for whole tree leaf area modeling purposes. These results indicate that, for the species sampled, accurate leaf area estimates can be obtained through partially-destructive sampling and using common forest inventory data.

  2. Canopy gradients in leaf functional traits for species that differ in growth strategies and shade tolerance.

    Science.gov (United States)

    Coble, Adam P; Fogel, Marilyn L; Parker, Geoffrey G

    2017-10-01

    In temperate deciduous forests, vertical gradients in leaf mass per area (LMA) and area-based leaf nitrogen (Narea) are strongly controlled by gradients in light availability. While there is evidence that hydrostatic constraints on leaf development may diminish LMA and Narea responses to light, inherent differences among tree species may also influence leaf developmental and morphological response to light. We investigated vertical gradients in LMA, Narea and leaf carbon isotope composition (δ13C) for three temperate deciduous species (Carpinus caroliniana Walter, Fagus grandifolia Ehrh., Liriodendron tulipifera L.) that differed in growth strategy (e.g., indeterminate and determinate growth), shade tolerance and leaf area to sapwood ratio (Al:As). Leaves were sampled across a broad range of light conditions within three vertical layers of tree crowns to maximize variation in light availability at each height and to minimize collinearity between light and height. All species displayed similar responses to light with respect to Narea and δ13C, but not for LMA. Light was more important for gradients in LMA for the shade-tolerant (C. caroliniana) and -intolerant (L. tulipifera) species with indeterminate growth, and height (e.g., hydrostatic gradients) and light were equally important for the shade-tolerant (F. grandifolia) species with determinate growth. Fagus grandifolia had a higher morphological plasticity in response to light, which may offer a competitive advantage in occupying a broader range of light conditions throughout the canopy. Differences in responses to light and height for the taller tree species, L. tulipifera and F. grandifolia, may be attributed to differences in growth strategy or Al:As, which may alter morphological and functional responses to light availability. While height was important in F. grandifolia, height was no more robust in predicting LMA than light in any of the species, confirming the strong role of light availability in

  3. Mapping of QTLs for leaf area and the association with winter ...

    African Journals Online (AJOL)

    Variations in plant architecture are often associated with the ability of plants to survive cold stress during winter. In studies of winter hardiness in lentil, it appeared that small leaf area was associated with improved winter survival. Based on this observation, the inheritance of leaf area and the relationship with winter ...

  4. Evaluation of four methods for estimating leaf area of isolated trees

    Science.gov (United States)

    P.J. Peper; E.G. McPherson

    2003-01-01

    The accurate modeling of the physiological and functional processes of urban forests requires information on the leaf area of urban tree species. Several non-destructive, indirect leaf area sampling methods have shown good performance for homogenous canopies. These methods have not been evaluated for use in urban settings where trees are typically isolated and...

  5. The effect of air pollution and other environmental stressors on leaf fluctuating asymmetry and specific leaf area of Salix alba L

    Energy Technology Data Exchange (ETDEWEB)

    Wuytack, Tatiana, E-mail: tatiana.wuytack@ua.ac.be [Department of Bioscience Engineering, University of Antwerp, Groenenborgerlaan 171, B-2020 Antwerp (Belgium); Wuyts, Karen, E-mail: karen.wuyts@ugent.be [Department of Bioscience Engineering, University of Antwerp, Groenenborgerlaan 171, B-2020 Antwerp (Belgium); Laboratory of Forestry, Department of Forest and Water Management, Ghent University, Geraardsbergsesteenweg 267, B-9090 Gontrode (Melle) (Belgium); Van Dongen, Stefan, E-mail: stefan.vandongen@ua.ac.be [Department of Biology, University of Antwerp, Groenenborgerlaan 171, B-2020 Antwerp (Belgium); Baeten, Lander, E-mail: lander.baeten@ugent.be [Laboratory of Forestry, Department of Forest and Water Management, Ghent University, Geraardsbergsesteenweg 267, B-9090 Gontrode (Melle) (Belgium); Kardel, Fatemeh, E-mail: fatemeh.kardel@ua.ac.be [Department of Bioscience Engineering, University of Antwerp, Groenenborgerlaan 171, B-2020 Antwerp (Belgium); Verheyen, Kris, E-mail: kris.verheyen@ugent.be [Laboratory of Forestry, Department of Forest and Water Management, Ghent University, Geraardsbergsesteenweg 267, B-9090 Gontrode, Melle (Belgium); Samson, Roeland, E-mail: roeland.samson@ua.ac.be [Department of Bioscience Engineering, University of Antwerp, Groenenborgerlaan 171, B-2020 Antwerp (Belgium)

    2011-10-15

    We aimed at evaluating the effect of low-level air pollution on leaf area fluctuating asymmetry (FAA) and specific leaf area (SLA) of Salix alba L., taking into account other environmental factors. Cuttings were grown in standardized conditions in the near vicinity of air quality measuring stations in Belgium. Variability of SLA and FAA between measuring stations explained 83% and 7.26%, respectively, of the total variability. FAA was not influenced by air pollution or environmental factors such as shading, herbivory, air temperature and humidity. SLA was increased by an increase in shadow, while NO{sub x} and O{sub 3} concentrations had only a marginal influence. The influence of SO{sub 2} concentration was negligible. Although our data analysis suggests a relationship between SLA and NO{sub x}/O{sub 3} concentration, the absence of a straightforward relationship between FAA and SLA and air pollution still questions the usefulness of these bio-indicators for monitoring air pollution. - Highlights: > Leaf characteristics of white willow as possible bio-indicators for air quality. > Fluctuating asymmetry is not a good bio-indicator for monitoring the air quality. > Shadow increases specific leaf area. > NO{sub x} and O{sub 3} change specific leaf area of white willow. - Specific leaf area of S. alba increased with increasing shade and, in less extent, with increasing NO{sub x} and decreasing O{sub 3} concentration, while leaf asymmetry did not respond to air pollution

  6. The effect of air pollution and other environmental stressors on leaf fluctuating asymmetry and specific leaf area of Salix alba L

    International Nuclear Information System (INIS)

    Wuytack, Tatiana; Wuyts, Karen; Van Dongen, Stefan; Baeten, Lander; Kardel, Fatemeh; Verheyen, Kris; Samson, Roeland

    2011-01-01

    We aimed at evaluating the effect of low-level air pollution on leaf area fluctuating asymmetry (FAA) and specific leaf area (SLA) of Salix alba L., taking into account other environmental factors. Cuttings were grown in standardized conditions in the near vicinity of air quality measuring stations in Belgium. Variability of SLA and FAA between measuring stations explained 83% and 7.26%, respectively, of the total variability. FAA was not influenced by air pollution or environmental factors such as shading, herbivory, air temperature and humidity. SLA was increased by an increase in shadow, while NO x and O 3 concentrations had only a marginal influence. The influence of SO 2 concentration was negligible. Although our data analysis suggests a relationship between SLA and NO x /O 3 concentration, the absence of a straightforward relationship between FAA and SLA and air pollution still questions the usefulness of these bio-indicators for monitoring air pollution. - Highlights: → Leaf characteristics of white willow as possible bio-indicators for air quality. → Fluctuating asymmetry is not a good bio-indicator for monitoring the air quality. → Shadow increases specific leaf area. → NO x and O 3 change specific leaf area of white willow. - Specific leaf area of S. alba increased with increasing shade and, in less extent, with increasing NO x and decreasing O 3 concentration, while leaf asymmetry did not respond to air pollution

  7. Mango leaf gall formation: varietal susceptibility and within tree distribution

    International Nuclear Information System (INIS)

    Khan, H.A.; Akram, W.; Khan, M.A.

    2012-01-01

    The present study was carried out to screen most commonly cultivated mango, Mangifera indica L., cultivars for their susceptibility to gall formation. Sarooli cultivar proved to be the most resistant one by having a minimum number of galls per 100 leaves. The abundance of galls in four quadrants of the tree i.e., east, west, north and south, was also studied which revealed that east quadrant had maximum number of galls while the abundance of galls in the remaining quadrants was variable. Gall formation on mango leaves seemed to increase gradually with increasing height from the ground level, reached a maximum at the height 12 ft to 16 ft and then declined. Leaf area measurements and nutrient analysis of the leaves were also done to see their impact on gall formation. Correlation analysis revealed that gall formation was positively linked with leaf area and the amount of Zn (ppm), P (%), K (%) while N (%) had negative correlation (P<0.05) with gall formation. In conclusion, the findings of the present study could be helpful in the management of mango leaf gall formation. (author)

  8. Community Characteristics and Leaf Stoichiometric Traits of Desert Ecosystems Regulated by Precipitation and Soil in an Arid Area of China

    Science.gov (United States)

    Guan, Tianyu; Zhou, Jihua; Cai, Wentao; Gao, Nannan; Du, Hui; Jiang, Lianhe; Lai, Liming; Zheng, Yuanrun

    2018-01-01

    Precipitation is a key environmental factor determining plant community structure and function. Knowledge of how community characteristics and leaf stoichiometric traits respond to variation in precipitation is crucial for assessing the effects of global changes on terrestrial ecosystems. In this study, we measured community characteristics, leaf stoichiometric traits, and soil properties along a precipitation gradient (35–209 mm) in a desert ecosystem of Northwest China to explore the drivers of these factors. With increasing precipitation, species richness, aboveground biomass, community coverage, foliage projective cover (FPC), and leaf area index (LAI) all significantly increased, while community height decreased. The hyperarid desert plants were characterized by lower leaf carbon (C) and nitrogen/phosphorus (N/P) levels, and stable N and P, and these parameters did not change significantly with precipitation. The growth of desert plants was limited more by N than P. Soil properties, rather than precipitation, were the main drivers of desert plant leaf stoichiometric traits, whereas precipitation made the biggest contribution to vegetation structure and function. These results test the importance of precipitation in regulating plant community structure and composition together with soil properties, and provide further insights into the adaptive strategy of communities at regional scale in response to global climate change. PMID:29320458

  9. Community Characteristics and Leaf Stoichiometric Traits of Desert Ecosystems Regulated by Precipitation and Soil in an Arid Area of China.

    Science.gov (United States)

    Zhang, Xiaolong; Guan, Tianyu; Zhou, Jihua; Cai, Wentao; Gao, Nannan; Du, Hui; Jiang, Lianhe; Lai, Liming; Zheng, Yuanrun

    2018-01-10

    Precipitation is a key environmental factor determining plant community structure and function. Knowledge of how community characteristics and leaf stoichiometric traits respond to variation in precipitation is crucial for assessing the effects of global changes on terrestrial ecosystems. In this study, we measured community characteristics, leaf stoichiometric traits, and soil properties along a precipitation gradient (35-209 mm) in a desert ecosystem of Northwest China to explore the drivers of these factors. With increasing precipitation, species richness, aboveground biomass, community coverage, foliage projective cover (FPC), and leaf area index (LAI) all significantly increased, while community height decreased. The hyperarid desert plants were characterized by lower leaf carbon (C) and nitrogen/phosphorus (N/P) levels, and stable N and P, and these parameters did not change significantly with precipitation. The growth of desert plants was limited more by N than P. Soil properties, rather than precipitation, were the main drivers of desert plant leaf stoichiometric traits, whereas precipitation made the biggest contribution to vegetation structure and function. These results test the importance of precipitation in regulating plant community structure and composition together with soil properties, and provide further insights into the adaptive strategy of communities at regional scale in response to global climate change.

  10. Relationships between stem diameter, sapwood area, leaf area and transpiration in a young mountain ash forest.

    Science.gov (United States)

    Vertessy, R A; Benyon, R G; O'Sullivan, S K; Gribben, P R

    1995-09-01

    We examined relationships between stem diameter, sapwood area, leaf area and transpiration in a 15-year-old mountain ash (Eucalyptus regnans F. Muell.) forest containing silver wattle (Acacia dealbata Link.) as a suppressed overstory species and mountain hickory (Acacia frigescens J.H. Willis) as an understory species. Stem diameter explained 93% of the variation in leaf area, 96% of the variation in sapwood area and 88% of the variation in mean daily spring transpiration in 19 mountain ash trees. In seven silver wattle trees, stem diameter explained 87% of the variation in sapwood area but was a poor predictor of the other variables. When transpiration measurements from individual trees were scaled up to a plot basis, using stem diameter values for 164 mountain ash trees and 124 silver wattle trees, mean daily spring transpiration rates of the two species were 2.3 and 0.6 mm day(-1), respectively. The leaf area index of the plot was estimated directly by destructive sampling, and indirectly with an LAI-2000 plant canopy analyzer and by hemispherical canopy photography. All three methods gave similar results.

  11. Estimating the total leaf area of the green dwarf coconut tree (Cocos nucifera L.

    Directory of Open Access Journals (Sweden)

    Sousa Elias Fernandes de

    2005-01-01

    Full Text Available Leaf area has significant effect on tree transpiration, and its measurement is important to many study areas. This work aimed at developing a non-destructive, practical, and empirical method to estimate the total leaf area of green dwarf coconut palms (Cocos nucifera L. in plantations located at the northern region of Rio de Janeiro state, Brazil. A mathematical model was developed to estimate total leaf area values (TLA as function of the average lengths of the last three leaf raquis (LR3, and of the number of leaves in the canopy (NL. The model has satisfactory degree of accuracy for agricultural engineering purposes.

  12. A new method for estimating the leaf area index of cucumber and tomato plants Um novo método para estimar o índice de área foliar de plantas de pepino e tomate

    Directory of Open Access Journals (Sweden)

    Flávio Favaro Blanco

    2003-12-01

    Full Text Available Non-destructive methods of leaf area measurement are useful for small plant populations, such as experiments with potted plants, and allow the measurement of the same plant several times during the growing period. A methodology was developed to estimate the leaf area index (LAI of cucumber and tomato plants through the evaluation of the leaf area distribution pattern (LADP of the plants and the relative height of the leaves in the plants. Plant and leaf height, as well as the length and width of all leaves were measured and the area of some leaves was determined by a digital area meter. The obtained regression equations were used to estimate the leaf area for all relative heights along the plant. The LADP adjusted to a quadratic model for both crops and LAI were estimated by measuring the length and width of the leaves located at the relative heights representing the mean leaf area of the plants. The LAI estimations presented high precision and accuracy when the proposed methodology was used resulting in time and effort savings and being useful for both crops.Métodos não destrutivos para a medição da área foliar são úteis para pequena população de plantas, como experimentos com plantas conduzidas em vasos, e permitem que a mesma planta seja medida várias vezes durante o período de cultivo. O objetivo desse trabalho foi desenvolver uma metodologia para a estimativa do índice de área foliar (IAF do pepino e do tomate pela determinação do padrão de distribuição de área foliar (PDAF das plantas e da altura relativa da folha que representa a área foliar média da planta. A altura da planta e da folha, assim como o comprimento e a largura de todas as folhas, foram medidos e algumas folhas tiveram sua área determinada por um medidor digital de área foliar. As equações de regressão obtidas foram utilizadas para estimar a área foliar para todas as alturas relativas ao longo da planta. O PDAF ajustou-se a um modelo quadr

  13. The Design and Implementation of the Leaf Area Index Sensor

    Directory of Open Access Journals (Sweden)

    Xiuhong Li

    2015-03-01

    Full Text Available The quick and accurate acquisition of crop growth parameters on a large scale is important for agricultural management and food security. The combination of photographic and wireless sensor network (WSN techniques can be used to collect agricultural information, such as leaf area index (LAI, over long distances and in real time. Such acquisition not only provides farmers with photographs of crops and suggestions for farmland management, but also the collected quantitative parameters, such as LAI, can be used to support large scale research in ecology, hydrology, remote sensing, etc. The present research developed a Leaf Area Index Sensor (LAIS to continuously monitor the growth of crops in several sampling points, and applied 3G/WIFI communication technology to remotely collect (and remotely setup and upgrade crop photos in real-time. Then the crop photos are automatically processed and LAI is estimated based on the improved leaf area index of Lang and Xiang (LAILX algorithm in LAIS. The research also constructed a database of images and other information relating to crop management. The leaf length and width method (LAILLW can accurately measure LAI through direct field harvest. The LAIS has been tested in several exemplary applications, and validation with LAI from LAILLW. The LAI acquired by LAIS had been proved reliable.

  14. Optimal allocation of leaf epidermal area for gas exchange

    OpenAIRE

    de Boer, Hugo J.; Price, Charles A.; Wagner-Cremer, Friederike; Dekker, Stefan C.; Franks, Peter J.; Veneklaas, Erik J.

    2016-01-01

    Summary A long?standing research focus in phytology has been to understand how plants allocate leaf epidermal space to stomata in order to achieve an economic balance between the plant's carbon needs and water use. Here, we present a quantitative theoretical framework to predict allometric relationships between morphological stomatal traits in relation to leaf gas exchange and the required allocation of epidermal area to stomata. Our theoretical framework was derived from first principles of ...

  15. Non-destructive estimation of leaf area for different plant ages and accessions of Capsicum annuum L.

    NARCIS (Netherlands)

    Swart, de E.A.M.; Groenwold, R.; Kanne, H.J.; Stam, P.; Marcelis, L.F.M.; Voorrips, R.E.

    2004-01-01

    Accurate measurements of leaf area are important for agronomic and physiological studies. To be able to perform repeated measurements of leaf area on single (genetically unique) plants, a method was developed to estimate leaf area from non-destructive measurements in Capsicum annuum L. independent

  16. Leaf area index estimation of Eucalyptus grandis W.Hill. in plantations

    Directory of Open Access Journals (Sweden)

    Dubal Papamija-Muñoz

    2012-12-01

    Full Text Available We estimated leaf area index (LAI in Eucalyptus grandis W.Hill. plantations in four farms in the Smurfit Kappa Carton de Colombia (SKCC with three farms located in the city of Popayan (Cauca and one located in the municipality of Restrepo (Valle del Cauca. Each farm had three fertilized and three unfertilized plots with 64 individuals in each. We used three methods, Plant Canopy Analyzer 2000 (PCA 2000, flat photograph PIPEcv software and a destructive method, which was generated using a mathematical model. The first two methods were measured bimonthly for a year and the final method required trees being cut to measure their diameter. Estimation of leaf area index was 2.01 for PCA 2000, 3.12 for PIPEcv and 2.83 for the mathematical model. These values correspond to the average and range of leaf area indices obtained for each method on all farms. Statistically the three methodologies developed in this study were not closely related.

  17. Leaf area and net photosynthesis during development of Prunus serotina seedlings

    Science.gov (United States)

    Stephen B. Horsley; Kurt W. Gottschalk

    1993-01-01

    We used the plastochron index to study the relationship between plant age, leaf age and development, and net photosynthesis of black cherry (Prtmus serotina Ehrh.) seedlings. Leaf area and net photosynthesis were measured on all leaves >=75 mm of plants ranging in age from 7 to 20 plastochrons. Effects of plant developmental stage...

  18. A better way of representing stem area index in two-big-leaf models: the application and impact on canopy integration of leaf nitrogen content

    Science.gov (United States)

    Chen, M.; Butler, E. E.; Wythers, K. R.; Kattge, J.; Ricciuto, D. M.; Thornton, P. E.; Atkin, O. K.; Flores-Moreno, H.; Reich, P. B.

    2017-12-01

    In order to better estimate the carbon budget of the globe, accurately simulating gross primary productivity (GPP) in earth system models is critical. When upscaling leaf level photosynthesis to the canopy, climate models uses different big-leaf schemes. About half of the state-of-the-art earth system models use a "two-big-leaf" scheme that partitions canopies into direct and diffusively illuminated fractions to reduce high bias of GPP simulated by one-big-leaf models. Some two-big-leaf models, such as ACME (identical in this respect to CLM 4.5) add leaf area index (LAI) and stem area index (SAI) together when calculating canopy radiation transfer. This treatment, however, will result in higher fraction of sunlit leaves. It will also lead to an artificial overestimation of canopy nitrogen content. Here we introduce a new algorithm of simulating SAI in a two-big-leaf model. The new algorithm reduced the sunlit leave fraction of the canopy and conserved the nitrogen content from leaf to canopy level. The lower fraction of sunlit leaves reduced global GPP especially in tropical area. Compared to the default model, for the past 100 years (1909-2009), the averaged global annual GPP is lowered by 4.11 PgC year-1 using this new algorithm.

  19. Age-related effects on leaf area/sapwood area relationships, canopy transpiration and carbon gain of Norway spruce stands (Picea abies) in the Fichtelgebirge, Germany.

    Science.gov (United States)

    Köstner, B; Falge, E; Tenhunen, J D

    2002-06-01

    Stand age is an important structural determinant of canopy transpiration (E(c)) and carbon gain. Another more functional parameter of forest structure is the leaf area/sapwood area relationship, A(L)/A(S), which changes with site conditions and has been used to estimate leaf area index of forest canopies. The interpretation of age-related changes in A(L)/A(S) and the question of how A(L)/A(S) is related to forest functions are of current interest because they may help to explain forest canopy fluxes and growth. We conducted studies in mature stands of Picea abies (L.) Karst. varying in age from 40 to 140 years, in tree density from 1680 to 320 trees ha(-1), and in tree height from 15 to 30 m. Structural parameters were measured by biomass harvests of individual trees and stand biometry. We estimated E(c) from scaled-up xylem sap flux of trees, and canopy-level fluxes were predicted by a three-dimensional microclimate and gas exchange model (STANDFLUX). In contrast to pine species, A(L)/A(S) of P. abies increased with stand age from 0.26 to 0.48 m(2) cm(-2). Agreement between E(c) derived from scaled-up sap flux and modeled canopy transpiration was obtained with the same parameterization of needle physiology independent of stand age. Reduced light interception per leaf area and, as a consequence, reductions in net canopy photosynthesis (A(c)), canopy conductance (g(c)) and E(c) were predicted by the model in the older stands. Seasonal water-use efficiency (WUE = A(c)/E(c)), derived from scaled-up sap flux and stem growth as well as from model simulation, declined with increasing A(L)/A(S) and stand age. Based on the different behavior of age-related A(L)/A(S) in Norway spruce stands compared with other tree species, we conclude that WUE rather than A(L)/A(S) could represent a common age-related property of all species. We also conclude that, in addition to hydraulic limitations reducing carbon gain in old stands, a functional change in A(L)/A(S) that is related to

  20. Effects of canopy structural variables on retrieval of leaf dry matter content and specific leaf area from remotely sensed data

    NARCIS (Netherlands)

    Ali, A.M.; Darvishzadeh, R.; Skidmore, A.K.; van Duren, I.C.

    2016-01-01

    Leaf dry matter content (LDMC) and specific leaf area (SLA) are two important traits in measuring biodiversity. To use remote sensing for the estimation of these traits, it is essential to understand the underlying factors that influence their relationships with canopy reflectance. The effect of

  1. Why Does Not the Leaf Weight-Area Allometry of Bamboos Follow the 3/2-Power Law?

    Directory of Open Access Journals (Sweden)

    Shuyan Lin

    2018-05-01

    Full Text Available The principle of similarity (Thompson, 1917 states that the weight of an organism follows the 3/2-power law of its surface area and is proportional to its volume on the condition that the density is constant. However, the allometric relationship between leaf weight and leaf area has been reported to greatly deviate from the 3/2-power law, with the irregularity of leaf density largely ignored for explaining this deviation. Here, we choose 11 bamboo species to explore the allometric relationships among leaf area (A, density (ρ, length (L, thickness (T, and weight (W. Because the edge of a bamboo leaf follows a simplified two-parameter Gielis equation, we could show that A ∝ L2 and that A ∝ T2. This then allowed us to derive the density-thickness allometry ρ ∝ Tb and the weight-area allometry W ∝ A(b+3/2 ≈ A9/8, where b approximates −3/4. Leaf density is strikingly negatively associated with leaf thickness, and it is this inverse relationship that results in the weight-area allometry to deviate from the 3/2-power law. In conclusion, although plants are prone to invest less dry mass and thus produce thinner leaves when the leaf area is sufficient for photosynthesis, such leaf thinning needs to be accompanied with elevated density to ensure structural stability. The findings provide the insights on the evolutionary clue about the biomass investment and output of photosynthetic organs of plants. Because of the importance of leaves, plants could have enhanced the ratio of dry material per unit area of leaf in order to increase the efficiency of photosynthesis, relative the other parts of plants. Although the conclusion is drawn only based on 11 bamboo species, it should also be applicable to the other plants, especially considering previous works on the exponent of the weight-area relationship being less than 3/2 in plants.

  2. Environmental Assessment: Eagle Heights Housing Area Revitalization Dover Air Force Base, Delaware

    Science.gov (United States)

    2004-07-01

    tidal species. Butterflies were the only insects surveyed, and nine were found on base. Approximately 51 species of birds were recorded on base...Jones River adjacent to the northern border of the housing area, the fro-fruit (Phyla lanceolata) and the hyssop-leaf hedge- nettle (Stachys...other sites in Delaware that this species is found. The hyssop-leaf hedge- nettle thrives in moist sandy soil along the coast and shoreline and occurs

  3. Assessment of plant biomass and nitrogen nutrition with plant height in early-to mid-season corn.

    Science.gov (United States)

    Yin, Xinhua; Hayes, Robert M; McClure, M Angela; Savoy, Hubert J

    2012-10-01

    The physiological basis for using non-destructive high-resolution measurements of plant height through plant height sensing to guide variable-rate nitrogen (N) applications on corn (Zea mays L.) during early (six-leaf growth stage, V6) to mid (V12) season is largely unknown. This study was conducted to assess the relationships of plant biomass and leaf N with plant height in early- to mid-season corn under six different N rate treatments. Corn plant biomass was significantly and positively related to plant height under an exponential model when both were measured at V6. This relationship explained 62-78% of the variations in corn biomass production. Leaf N concentration was, in general, significantly and positively related to plant height when both were measured at V6, V8, V10 and V12. This relationship became stronger as the growing season progressed from V6 to V12. The relationship of leaf N with plant height in early- to mid-season corn was affected by initial soil N fertility and abnormal weather conditions. The relationship of leaf N concentration with plant height may provide a physiological basis for using plant height sensing to guide variable-rate N applications on corn. Copyright © 2012 Society of Chemical Industry.

  4. Estimating leaf functional traits by inversion of PROSPECT: Assessing leaf dry matter content and specific leaf area in mixed mountainous forest

    Science.gov (United States)

    Ali, Abebe Mohammed; Darvishzadeh, Roshanak; Skidmore, Andrew K.; Duren, Iris van; Heiden, Uta; Heurich, Marco

    2016-03-01

    Assessments of ecosystem functioning rely heavily on quantification of vegetation properties. The search is on for methods that produce reliable and accurate baseline information on plant functional traits. In this study, the inversion of the PROSPECT radiative transfer model was used to estimate two functional leaf traits: leaf dry matter content (LDMC) and specific leaf area (SLA). Inversion of PROSPECT usually aims at quantifying its direct input parameters. This is the first time the technique has been used to indirectly model LDMC and SLA. Biophysical parameters of 137 leaf samples were measured in July 2013 in the Bavarian Forest National Park, Germany. Spectra of the leaf samples were measured using an ASD FieldSpec3 equipped with an integrating sphere. PROSPECT was inverted using a look-up table (LUT) approach. The LUTs were generated with and without using prior information. The effect of incorporating prior information on the retrieval accuracy was studied before and after stratifying the samples into broadleaf and conifer categories. The estimated values were evaluated using R2 and normalized root mean square error (nRMSE). Among the retrieved variables the lowest nRMSE (0.0899) was observed for LDMC. For both traits higher R2 values (0.83 for LDMC and 0.89 for SLA) were discovered in the pooled samples. The use of prior information improved accuracy of the retrieved traits. The strong correlation between the estimated traits and the NIR/SWIR region of the electromagnetic spectrum suggests that these leaf traits could be assessed at canopy level by using remotely sensed data.

  5. AfSIS MODIS Collection: Leaf Area Index - FPAR, 2012 Release

    Data.gov (United States)

    Center for International Earth Science Information Network, Columbia University — The Africa Soil Information Service (AfSIS) Moderate Resolution Imaging Spectroradiometer (MODIS) Collection Leaf Area Index (LAI) and Photosynthetically Active...

  6. Tree differences in primary and secondary growth drive convergent scaling in leaf area to sapwood area across Europe

    NARCIS (Netherlands)

    Petit, Giai; Arx, von Georg; Kiorapostolou, Natasa; Lechthaler, Silvia; Prendin, Angela Luisa; Anfodillo, Tommaso; Caldeira, Maria C.; Cochard, Hervé; Copini, Paul; Crivellaro, Alan; Delzon, Sylvain; Gebauer, Roman; Gričar, Jožica; Grönholm, Leila; Hölttä, Teemu; Jyske, Tuula; Lavrič, Martina; Lintunen, Anna; Lobo-do-Vale, Raquel; Peltoniemi, Mikko; Peters, Richard L.; Robert, Elisabeth M.R.; Roig Juan, Sílvia; Senfeldr, Martin; Steppe, Kathy; Urban, Josef; Camp, Van Janne; Sterck, Frank

    2018-01-01

    Trees scale leaf (AL) and xylem (AX) areas to couple leaf transpiration and carbon gain with xylem water transport. Some species are known to acclimate in AL: AX balance in response to climate conditions, but whether trees of different species acclimate in AL: AX in similar ways over their entire

  7. 76 FR 77696 - Establishment of the Naches Heights Viticultural Area

    Science.gov (United States)

    2011-12-14

    ... Decision. SUMMARY: This final rule establishes the 13,254-acre ``Naches Heights'' viticultural area in... origin of their wines and to allow consumers to better identify wines they may purchase. DATES: Effective... consumers to identify wines they may purchase. Establishment of a viticultural area is neither an approval...

  8. Estimativa da área foliar de Crambe abyssinica por discos foliares e por fotos digitais Estimate leaf area of Crambe abyssinica for leaf discs and digital photos

    Directory of Open Access Journals (Sweden)

    Marcos Toebe

    2010-02-01

    Full Text Available A área foliar é importante na determinação do crescimento e desenvolvimento das culturas agrícolas. Assim, os objetivos do trabalho foram comparar os métodos de discos foliares e de fotos digitais na estimativa da área foliar de Crambe abyssinica e modelar a área foliar em função do comprimento (C, da largura (L e ou do produto comprimento vezes largura (CxL de diferentes tamanhos de folhas. Para isso, em 308 folhas, foram determinados a área foliar, o comprimento, a largura e o produto comprimento vezes largura por meio dos métodos de discos foliares e de fotos digitais. Em seguida, foram comparados os métodos por meio do coeficiente de correlação linear entre a área foliar. A seguir, em cada método, modelou-se a área foliar (Y em função do C, da L e do CxL, por meio dos modelos: linear, linear simples, quadrático, geométrico e exponencial. Os coeficientes de correlação linear de Pearson e de Spearman entre a área foliar dos métodos de discos foliares e de fotos digitais foram de 0,9917 e 0,9889, respectivamente, o que revela métodos concordantes. Em ambos os métodos, os modelos quadráticos e geométricos apresentaram os melhores coeficientes de determinação da área foliar em função do comprimento e da largura das folhas. A largura da folha é a variável que melhor estima a área foliar. O método de fotos digitais pode ser utilizado para estimar a área foliar de crambe.Leaf area is important in determining the growth and development of agricultural crops. The aim of this study was to compare the methods of leaf discs and digital photos in estimating leaf area of Crambe abyssinica, and model leaf area according to length (C, width (L and/ or the product of length width (CxL for different sizes of leaves. For this, in 308 leaves it was determined the leaf area, length, width and the product of length width using the methods of leaf discs and digital photos. Then the methods were compared using the linear

  9. Optimal allocation of leaf epidermal area for gas exchange.

    Science.gov (United States)

    de Boer, Hugo J; Price, Charles A; Wagner-Cremer, Friederike; Dekker, Stefan C; Franks, Peter J; Veneklaas, Erik J

    2016-06-01

    A long-standing research focus in phytology has been to understand how plants allocate leaf epidermal space to stomata in order to achieve an economic balance between the plant's carbon needs and water use. Here, we present a quantitative theoretical framework to predict allometric relationships between morphological stomatal traits in relation to leaf gas exchange and the required allocation of epidermal area to stomata. Our theoretical framework was derived from first principles of diffusion and geometry based on the hypothesis that selection for higher anatomical maximum stomatal conductance (gsmax ) involves a trade-off to minimize the fraction of the epidermis that is allocated to stomata. Predicted allometric relationships between stomatal traits were tested with a comprehensive compilation of published and unpublished data on 1057 species from all major clades. In support of our theoretical framework, stomatal traits of this phylogenetically diverse sample reflect spatially optimal allometry that minimizes investment in the allocation of epidermal area when plants evolve towards higher gsmax . Our results specifically highlight that the stomatal morphology of angiosperms evolved along spatially optimal allometric relationships. We propose that the resulting wide range of viable stomatal trait combinations equips angiosperms with developmental and evolutionary flexibility in leaf gas exchange unrivalled by gymnosperms and pteridophytes. © 2016 The Authors New Phytologist © 2016 New Phytologist Trust.

  10. Two Inexpensive and Non-destructive Techniques to Correct for Smaller-Than-Gasket Leaf Area in Gas Exchange Measurements

    Directory of Open Access Journals (Sweden)

    Andreas M. Savvides

    2018-04-01

    Full Text Available The development of technology, like the widely-used off-the-shelf portable photosynthesis systems, for the quantification of leaf gas exchange rates and chlorophyll fluorescence offered photosynthesis research a massive boost. Gas exchange parameters in such photosynthesis systems are calculated as gas exchange rates per unit leaf area. In small chambers (<10 cm2, the leaf area used by the system for these calculations is actually the internal gasket area (AG, provided that the leaf covers the entire AG. In this study, we present two inexpensive and non-destructive techniques that can be used to easily quantify the enclosed leaf area (AL of plant species with leaves of surface area much smaller than the AG, such as that of cereal crops. The AL of the cereal crop species studied has been measured using a standard image-based approach (iAL and estimated using a leaf width-based approach (wAL. iAL and wAL did not show any significant differences between them in maize, barley, hard and soft wheat. Similar results were obtained when the wAL was tested in comparison with iAL in different positions along the leaf in all species studied. The quantification of AL and the subsequent correction of leaf gas exchange parameters for AL provided a precise quantification of net photosynthesis and stomatal conductance especially with decreasing AL. This study provides two practical, inexpensive and non-destructive solutions to researchers dealing with photosynthesis measurements on small-leaf plant species. The image-based technique can be widely used for quantifying AL in many plant species despite their leaf shape. The leaf width-based technique can be securely used for quantifying AL in cereal crop species such as maize, wheat and barley along the leaf. Both techniques can be used for a wide range of gasket shapes and sizes with minor technique-specific adjustments.

  11. Growth characteristics of mangrove seedling in silvofishery pond – the allometric relationship of height, diameter and leaf abundance

    Science.gov (United States)

    Dwi Hastuti, Endah; Budi Hastuti, Rini

    2018-03-01

    Dynamic environment condition of the silvofishery pond should provide an effect on the growth of mangrove seedling. This research aimed to observe the morphometric growth rate of mangrove seedling of Avicennia marina and Rhizophora mucronata planted in the silvofishery pond and to analyze the morphometric growth relationship of height, diameter and leaf number development of mangrove seedling. The research was conducted through field experiment involving mangrove species of A. marina and R. mucronata for 18 months during March 2015 to September 2016, both single structured and mixed structure. The observation was conducted every 13 weeks including seedling height, diameter and number of leaves. Data analysis was conducted by regression to provide the statistical relation between the growth of diameter – height, diameter – number of leaves and height – number of leaves. The result showed that the growth rate of A. marina in single structured pond was ranged from 0.38 – 3.00 cm.wk-1, 0.0015 – 0.0969 cm.wk‑1 and 0.1 – 13.7 leaves.wk‑1 respectively for height, diameter and number of leaves, while in mixed structure was 0.23 – 1.69 cm.wk‑1, 0.0169 – 0.0731 cm.wk‑1 and 0.5 – 14.0 leaves.wk-. The growth of R. mucronata respectively in single and mixed structure were 0.08 – 2.00 cm.wk‑1 and 0.15 – 2.62 cm.wk‑1, 0.0031 – 0.1369 cm.wk‑1 and 0.0008 – 0.0831 cm.wk‑1 and 0.0 – 1.9 leaves.wk‑1 and 0.0 – 1.6 leaves.wk-1respectively for height, diameter and number of leaves. Data analysis showed that the growth of seedling height of Avicennia in the mixed structure was significantly affected by its diameter growth and the number of leaves of Avicennia in single structured was significantly affected by its diameter. While the height, diameter and number of leaves of R. mucronata both in mixed and single structured silvofishery ponds were independent to each other. This research concluded that mangrove seedling growth is varied among

  12. Vertical leaf area distribution, light transmittance, and application of the Beer-Lambert Law in four mature hardwood stands in the southern Appalachians

    Science.gov (United States)

    James M. Vose; Neal H. Sullivan; Barton D. Clinton; Paul V. Bolstad

    1995-01-01

    We quantified stand leaf area index and vertical leaf area distribution, and developed canopy extinction coefficients (k), in four mature hardwood stands. Leaf area index, calculated from litter fall and specific leaf area (cm²·g-1), ranged from 4.3 to 5.4 m²·m-2. In three of the four stands, leaf area was distributed in...

  13. Effects of Temporal and Interspecific Variation of Specific Leaf Area on Leaf Area Index Estimation of Temperate Broadleaved Forests in Korea

    Directory of Open Access Journals (Sweden)

    Boram Kwon

    2016-09-01

    Full Text Available This study investigated the effects of interspecific and temporal variation of specific leaf area (SLA, cm2·g−1 on leaf area index (LAI estimation for three deciduous broadleaved forests (Gwangneung (GN, Taehwa (TH, and Gariwang (GRW in Korea with varying ages and composition of tree species. In fall of 2014, fallen leaves were periodically collected using litter traps and classified by species. LAI was estimated by obtaining SLAs using four calculation methods (A: including both interspecific and temporal variation in SLA; B: species specific mean SLA; C: period-specific mean SLA; and D: overall mean, then multiplying the SLAs by the amount of leaves. SLA varied across different species in all plots, and SLAs of upper canopy species were less than those of lower canopy species. The LAIs calculated using method A, the reference method, were GN 6.09, TH 5.42, and GRW 4.33. LAIs calculated using method B showed a difference of up to 3% from the LAI of method A, but LAIs calculated using methods C and D were overestimated. Therefore, species specific SLA must be considered for precise LAI estimation for broadleaved forests that include multiple species.

  14. Effect of weed control treatments on total leaf area of plantation black walnut (Juglans nigra)

    Science.gov (United States)

    Jason Cook; Michael R. Saunders

    2013-01-01

    Determining total tree leaf area is necessary for describing tree carbon balance, growth efficiency, and other measures used in tree-level and stand-level physiological growth models. We examined the effects of vegetation control methods on the total leaf area of sapling-size plantation black walnut trees using allometric approaches. We found significant differences in...

  15. ISLSCP II Leaf Area Index (LAI) from Field Measurements, 1932-2000

    Data.gov (United States)

    National Aeronautics and Space Administration — ABSTRACT: Leaf Area Index (LAI) data from the scientific literature, covering the period from 1932-2000, have been compiled at the Oak Ridge National Laboratory...

  16. Leaf structural characteristics are less important than leaf chemical properties in determining the response of leaf mass per area and photosynthesis of Eucalyptus saligna to industrial-age changes in [CO2] and temperature.

    Science.gov (United States)

    Xu, Cheng-Yuan; Salih, Anya; Ghannoum, Oula; Tissue, David T

    2012-10-01

    The rise in atmospheric [CO(2)] is associated with increasing air temperature. However, studies on plant responses to interactive effects of [CO(2)] and temperature are limited, particularly for leaf structural attributes. In this study, Eucalyptus saligna plants were grown in sun-lit glasshouses differing in [CO(2)] (290, 400, and 650 µmol mol(-1)) and temperature (26 °C and 30 °C). Leaf anatomy and chloroplast parameters were assessed with three-dimensional confocal microscopy, and the interactive effects of [CO(2)] and temperature were quantified. The relative influence of leaf structural attributes and chemical properties on the variation of leaf mass per area (LMA) and photosynthesis within these climate regimes was also determined. Leaf thickness and mesophyll size increased in higher [CO(2)] but decreased at the warmer temperature; no treatment interaction was observed. In pre-industrial [CO(2)], warming reduced chloroplast diameter without altering chloroplast number per cell, but the opposite pattern (reduced chloroplast number per cell and unchanged chloroplast diameter) was observed in both current and projected [CO(2)]. The variation of LMA was primarily explained by total non-structural carbohydrate (TNC) concentration rather than leaf thickness. Leaf photosynthetic capacity (light- and [CO(2)]-saturated rate at 28 °C) and light-saturated photosynthesis (under growth [CO(2)] and temperature) were primarily determined by leaf nitrogen contents, while secondarily affected by chloroplast gas exchange surface area and chloroplast number per cell, respectively. In conclusion, leaf structural attributes are less important than TNC and nitrogen in affecting LMA and photosynthesis responses to the studied climate regimes, indicating that leaf structural attributes have limited capacity to adjust these functional traits in a changing climate.

  17. Indirect Field Measurement of Wine-Grape Vineyard Canopy Leaf Area Index

    Science.gov (United States)

    Johnson, Lee F.; Pierce, Lars L.; Skiles, J. W. (Technical Monitor)

    2002-01-01

    Leaf area index (LAI) indirect measurements were made at 12 study plots in California's Napa Valley commercial wine-grape vineyards with a LI-COR LI-2000 Plant Canopy Analyzer (PCA). The plots encompassed different trellis systems, biological varieties, and planting densities. LAI ranged from 0.5 - 2.25 sq m leaf area/ sq m ground area according to direct (defoliation) measurements. Indirect LAI reported by the PCA was significantly related to direct LAI (r(exp 2) = 0.78, p less than 001). However, the PCA tended to underestimate direct LAI by about a factor of two. Narrowing the instrument's conical field of view from 148 deg to 56 deg served to increase readings by approximately 30%. The PCA offers a convenient way to discern relative differences in vineyard canopy density. Calibration by direct measurement (defoliation) is recommended in cases where absolute LAI is desired. Calibration equations provided herein may be inverted to retrieve actual vineyard LAI from PCA readings.

  18. Measurements methods and variability assesment of the Norway spruce total leaf area. Implications for remote sensing

    Czech Academy of Sciences Publication Activity Database

    Homolová, L.; Lukeš, Petr; Malenovský, Z.; Lhotáková, Z.; Kaplan, Věroslav; Hanuš, Jan

    2013-01-01

    Roč. 27, č. 1 (2013), s. 111-121 ISSN 0931-1890 R&D Projects: GA ČR GA205/09/ 1989 Institutional support: RVO:67179843 Keywords : chlorophyll content * conversion factor * Picea abies * projected leaf area * remote sensing * total leaf area Subject RIV: EH - Ecology, Behaviour Impact factor: 1.869, year: 2013

  19. Branch age and light conditions determine leaf-area-specific conductivity in current shoots of Scots pine.

    Science.gov (United States)

    Grönlund, Leila; Hölttä, Teemu; Mäkelä, Annikki

    2016-08-01

    Shoot size and other shoot properties more or less follow the availability of light, but there is also evidence that the topological position in a tree crown has an influence on shoot development. Whether the hydraulic properties of new shoots are more regulated by the light or the position affects the shoot acclimation to changing light conditions and thereby to changing evaporative demand. We investigated the leaf-area-specific conductivity (and its components sapwood-specific conductivity and Huber value) of the current-year shoots of Scots pine (Pinus sylvestris L.) in relation to light environment and topological position in three different tree classes. The light environment was quantified in terms of simulated transpiration and the topological position was quantified by parent branch age. Sample shoot measurements included length, basal and tip diameter, hydraulic conductivity of the shoot, tracheid area and density, and specific leaf area. In our results, the leaf-area-specific conductivity of new shoots declined with parent branch age and increased with simulated transpiration rate of the shoot. The relation to transpiration demand seemed more decisive, since it gave higher R(2) values than branch age and explained the differences between the tree classes. The trend of leaf-area-specific conductivity with simulated transpiration was closely related to Huber value, whereas the trend of leaf-area-specific conductivity with parent branch age was related to a similar trend in sapwood-specific conductivity. © The Author 2016. Published by Oxford University Press. All rights reserved. For Permissions, please email: journals.permissions@oup.com.

  20. Leaf area and light use efficiency patterns of Norway spruce under different thinning regimes and age classes

    Science.gov (United States)

    Gspaltl, Martin; Bauerle, William; Binkley, Dan; Sterba, Hubert

    2013-01-01

    Silviculture focuses on establishing forest stand conditions that improve the stand increment. Knowledge about the efficiency of an individual tree is essential to be able to establish stand structures that increase tree resource use efficiency and stand level production. Efficiency is often expressed as stem growth per unit leaf area (leaf area efficiency), or per unit of light absorbed (light use efficiency). We tested the hypotheses that: (1) volume increment relates more closely with crown light absorption than leaf area, since one unit of leaf area can receive different amounts of light due to competition with neighboring trees and self-shading, (2) dominant trees use light more efficiently than suppressed trees and (3) thinning increases the efficiency of light use by residual trees, partially accounting for commonly observed increases in post-thinning growth. We investigated eight even-aged Norway spruce (Picea abies (L.) Karst.) stands at Bärnkopf, Austria, spanning three age classes (mature, immature and pole-stage) and two thinning regimes (thinned and unthinned). Individual leaf area was calculated with allometric equations and absorbed photosynthetically active radiation was estimated for each tree using the three-dimensional crown model Maestra. Absorbed photosynthetically active radiation was only a slightly better predictor of volume increment than leaf area. Light use efficiency increased with increasing tree size in all stands, supporting the second hypothesis. At a given tree size, trees from the unthinned plots were more efficient, however, due to generally larger tree sizes in the thinned stands, an average tree from the thinned treatment was superior (not congruent in all plots, thus only partly supporting the third hypothesis). PMID:25540477

  1. Tree differences in primary and secondary growth drive convergent scaling in leaf area to sapwood area across Europe.

    Science.gov (United States)

    Petit, Giai; von Arx, Georg; Kiorapostolou, Natasa; Lechthaler, Silvia; Prendin, Angela Luisa; Anfodillo, Tommaso; Caldeira, Maria C; Cochard, Hervé; Copini, Paul; Crivellaro, Alan; Delzon, Sylvain; Gebauer, Roman; Gričar, Jožica; Grönholm, Leila; Hölttä, Teemu; Jyske, Tuula; Lavrič, Martina; Lintunen, Anna; Lobo-do-Vale, Raquel; Peltoniemi, Mikko; Peters, Richard L; Robert, Elisabeth M R; Roig Juan, Sílvia; Senfeldr, Martin; Steppe, Kathy; Urban, Josef; Van Camp, Janne; Sterck, Frank

    2018-06-01

    Trees scale leaf (A L ) and xylem (A X ) areas to couple leaf transpiration and carbon gain with xylem water transport. Some species are known to acclimate in A L  : A X balance in response to climate conditions, but whether trees of different species acclimate in A L  : A X in similar ways over their entire (continental) distributions is unknown. We analyzed the species and climate effects on the scaling of A L vs A X in branches of conifers (Pinus sylvestris, Picea abies) and broadleaved (Betula pendula, Populus tremula) sampled across a continental wide transect in Europe. Along the branch axis, A L and A X change in equal proportion (isometric scaling: b ˜ 1) as for trees. Branches of similar length converged in the scaling of A L vs A X with an exponent of b = 0.58 across European climates irrespective of species. Branches of slow-growing trees from Northern and Southern regions preferentially allocated into new leaf rather than xylem area, with older xylem rings contributing to maintaining total xylem conductivity. In conclusion, trees in contrasting climates adjust their functional balance between water transport and leaf transpiration by maintaining biomass allocation to leaves, and adjusting their growth rate and xylem production to maintain xylem conductance. © 2018 The Authors. New Phytologist © 2018 New Phytologist Trust.

  2. Estimation of Leaf Area Index and its Sunlit Portion from DSCOVR EPIC data

    Science.gov (United States)

    Knyazikhin, Y.; Yang, B.; Mottus, M.; Rautiainen, M.; Stenberg, P.; Yan, L.; Chen, C.; Yan, K.; Park, T.; Myneni, R. B.; Song, W.

    2016-12-01

    The NASA's Earth Polychromatic Imaging Camera (EPIC) onboard NOAA's Deep Space Climate Observatory (DSCOVR) mission was launched on February 11, 2015 to the Sun-Earth Lagrangian L1 point where it began to collect radiance data of the entire sunlit Earth at 16 km resolution (in equatorial zone) every 65 to 110 min in June 2015. It provides imageries in near backscattering directions with the scattering angle between 168o and 176o at ten UV to Near-IR narrow spectral bands centered at 317.5 (band width 1.0) nm, 325.0 (1.0) nm, 340.0 (3.0) nm, 388.0 (3.0) nm, 433.0 (3.0) nm, 551.0 (3.0) nm, 680.0 (1.7) nm, 687.8 (0.6) nm, 764.0 (1.7) nm and 779.5 (2.0) nm. This poster presents the theoretical basis of the algorithm designed for the generation of leaf area index (LAI) and diurnal course of sunlit leaf area index (SLAI) from EPIC Bidirectional Reflectance Factor of vegetated land. LAI and SLAI are defined as the total hemi-surface and sunlit leaf semi-surface per unit ground area. Whereas LAI is a standard product of many satellite the SLAI is a new satellite-derived parameter. Sunlit and shaded leaves exhibit different radiative response to incident Photosynthetically Active Radiation (400-700 nm), which in turn triggers various physiological and physical processes required for the functioning of plants. Leaf area and its sunlit portion are key state parameters in most ecosystem productivity and carbon/nitrogen cycle. Status of the EPIC LAI/SLAI product and its validation strategy are also discussed in this poster.

  3. Leaf wetness distribution within a potato crop

    Science.gov (United States)

    Heusinkveld, B. G.

    2010-07-01

    The Netherlands has a mild maritime climate and therefore the major interest in leaf wetness is associated with foliar plant diseases. During moist micrometeorological conditions (i.e. dew, fog, rain), foliar fungal diseases may develop quickly and thereby destroy a crop quickly. Potato crop monocultures covering several hectares are especially vulnerable to such diseases. Therefore understanding and predicting leaf wetness in potato crops is crucial in crop disease control strategies. A field experiment was carried out in a large homogeneous potato crop in the Netherlands during the growing season of 2008. Two innovative sensor networks were installed as a 3 by 3 grid at 3 heights covering an area of about 2 hectares within two larger potato crops. One crop was located on a sandy soil and one crop on a sandy peat soil. In most cases leaf wetting starts in the top layer and then progresses downward. Leaf drying takes place in the same order after sunrise. A canopy dew simulation model was applied to simulate spatial leaf wetness distribution. The dew model is based on an energy balance model. The model can be run using information on the above-canopy wind speed, air temperature, humidity, net radiation and within canopy air temperature, humidity and soil moisture content and temperature conditions. Rainfall was accounted for by applying an interception model. The results of the dew model agreed well with the leaf wetness sensors if all local conditions were considered. The measurements show that the spatial correlation of leaf wetness decreases downward.

  4. Effect of nitrogen supply on leaf appearance, leaf growth, leaf nitrogen economy and photosynthetic capacity in maize (Zea mays L.)

    NARCIS (Netherlands)

    Vos, J.; Putten, van der P.E.L.; Birch, C.J.

    2005-01-01

    Leaf area growth and nitrogen concentration per unit leaf area, Na (g m-2 N) are two options plants can use to adapt to nitrogen limitation. Previous work indicated that potato (Solanum tuberosum L.) adapts the size of leaves to maintain Na and photosynthetic capacity per unit leaf area. This paper

  5. Leaf area index uncertainty estimates for model-data fusion applications

    Science.gov (United States)

    Andrew D. Richardson; D. Bryan Dail; D.Y. Hollinger

    2011-01-01

    Estimates of data uncertainties are required to integrate different observational data streams as model constraints using model-data fusion. We describe an approach with which random and systematic uncertainties in optical measurements of leaf area index [LAI] can be quantified. We use data from a measurement campaign at the spruce-dominated Howland Forest AmeriFlux...

  6. Taxonomy and remote sensing of leaf mass per area (LMA) in humid tropical forests

    Science.gov (United States)

    Gregory P. Asner; Roberta E. Martin; Raul Tupayachi; Ruth Emerson; Paola Martinez; Felipe Sinca; George V.N. Powell; S. Joseph Wright; Ariel E. Lugo

    2011-01-01

    Leaf mass per area (LMA) is a trait of central importance to plant physiology and ecosystem function, but LMA patterns in the upper canopies of humid tropical forests have proved elusive due to tall species and high diversity. We collected top-of-canopy leaf samples from 2873 individuals in 57 sites spread across the Neotropics, Australasia, and Caribbean and Pacific...

  7. USING COMBINATION OF PLANAR AND HEIGHT FEATURES FOR DETECTING BUILT-UP AREAS FROM HIGH-RESOLUTION STEREO IMAGERY

    Directory of Open Access Journals (Sweden)

    F. Peng

    2017-09-01

    Full Text Available Within-class spectral variation and between-class spectral confusion in remotely sensed imagery degrades the performance of built-up area detection when using planar texture, shape, and spectral features. Terrain slope and building height are often used to optimize the results, but extracted from auxiliary data (e.g. LIDAR data, DSM. Moreover, the auxiliary data must be acquired around the same time as image acquisition. Otherwise, built-up area detection accuracy is affected. Stereo imagery incorporates both planar and height information unlike single remotely sensed images. Stereo imagery acquired by many satellites (e.g. Worldview-4, Pleiades-HR, ALOS-PRISM, and ZY-3 can be used as data source of identifying built-up areas. A new method of identifying high-accuracy built-up areas from stereo imagery is achieved by using a combination of planar and height features. The digital surface model (DSM and digital orthophoto map (DOM are first generated from stereo images. Then, height values of above-ground objects (e.g. buildings are calculated from the DSM, and used to obtain raw built-up areas. Other raw built-up areas are obtained from the DOM using Pantex and Gabor, respectively. Final high-accuracy built-up area results are achieved from these raw built-up areas using the decision level fusion. Experimental results show that accurate built-up areas can be achieved from stereo imagery. The height information used in the proposed method is derived from stereo imagery itself, with no need to require auxiliary height data (e.g. LIDAR data. The proposed method is suitable for spaceborne and airborne stereo pairs and triplets.

  8. Using Combination of Planar and Height Features for Detecting Built-Up Areas from High-Resolution Stereo Imagery

    Science.gov (United States)

    Peng, F.; Cai, X.; Tan, W.

    2017-09-01

    Within-class spectral variation and between-class spectral confusion in remotely sensed imagery degrades the performance of built-up area detection when using planar texture, shape, and spectral features. Terrain slope and building height are often used to optimize the results, but extracted from auxiliary data (e.g. LIDAR data, DSM). Moreover, the auxiliary data must be acquired around the same time as image acquisition. Otherwise, built-up area detection accuracy is affected. Stereo imagery incorporates both planar and height information unlike single remotely sensed images. Stereo imagery acquired by many satellites (e.g. Worldview-4, Pleiades-HR, ALOS-PRISM, and ZY-3) can be used as data source of identifying built-up areas. A new method of identifying high-accuracy built-up areas from stereo imagery is achieved by using a combination of planar and height features. The digital surface model (DSM) and digital orthophoto map (DOM) are first generated from stereo images. Then, height values of above-ground objects (e.g. buildings) are calculated from the DSM, and used to obtain raw built-up areas. Other raw built-up areas are obtained from the DOM using Pantex and Gabor, respectively. Final high-accuracy built-up area results are achieved from these raw built-up areas using the decision level fusion. Experimental results show that accurate built-up areas can be achieved from stereo imagery. The height information used in the proposed method is derived from stereo imagery itself, with no need to require auxiliary height data (e.g. LIDAR data). The proposed method is suitable for spaceborne and airborne stereo pairs and triplets.

  9. Effect of brushwood transposition on the leaf litter arthropod fauna in a cerrado area

    Directory of Open Access Journals (Sweden)

    Paula Cristina Benetton Vergílio

    2013-10-01

    Full Text Available The results of ecological restoration techniques can be monitored through biological indicators of soil quality such as the leaf litter arthropod fauna. This study aimed to determine the immediate effect of brushwood transposition transferred from an area of native vegetation to a disturbed area, on the leaf litter arthropod fauna in a degraded cerrado area. The arthropod fauna of four areas was compared: a degraded area with signal grass, two experimental brushwood transposition areas, with and without castor oil plants, and an area of native cerrado. In total, 7,660 individuals belonging to 23 taxa were sampled. Acari and Collembola were the most abundant taxa in all studied areas, followed by Coleoptera, Diptera, Hemiptera, Hymenoptera, and Symphyla. The brushwood transposition area without castor oil plants had the lowest abundance and dominance and the highest diversity of all areas, providing evidence of changes in the soil community. Conversely, the results showed that the presence of castor oil plants hampered early succession, negatively affecting ecological restoration in this area.

  10. Importance of the method of leaf area measurement to the interpretation of gas exchange of complex shoots

    Science.gov (United States)

    W. K. Smith; A. W. Schoettle; M. Cui

    1991-01-01

    Net CO(2) uptake in full sunlight, total leaf area (TLA), projected leaf area of detached leaves (PLA), and the silhouette area of attached leaves in their natural orientation to the sun at midday on June 1 (SLA) were measured for sun shoots of six conifer species. Among species, TLA/SLA ranged between 5.2 and 10.0 (x bar = 7.3), TLA/PLA ranged between 2.5 and 2.9 (x...

  11. Biophysical constraints on leaf expansion in a tall conifer.

    Science.gov (United States)

    Fredrick C. Meinzer; Barbara J. Bond; Jennifer A. Karanian

    2008-01-01

    The physiological mechanisms responsible for reduced extension growth as trees increase in height remain elusive. We evaluated biophysical constraints on leaf expansion in old-growth Douglas-fir (Psuedotsuga menziesii (Mirb.) Franco) trees. Needle elongation rates, plastic and elastic extensibility, bulk leaf water, (L...

  12. Solar radiation measurements and Leaf Area Index (LAI) from vegetal covers

    International Nuclear Information System (INIS)

    Wandelli, E.V.; Marques Filho, A. de O.

    1999-01-01

    A method by which a physical model of the solar radiation transfer in a vegetal medium is inverted to estimate the leaf area index (LAI) for different types of vegetation is presented here, as an alternative to the destructive experiments, which are a hard task to implement on the vegetation covers. Radiation data were obtained during the dry season — 1996, at the Embrapa Experimental Station, (BR 174 - km 54, 2° 31' S, 60° 01' W), Manaus, Brazil. The method yielded convergent values for the LAI between different adopted radiation classes with more stable estimates at time when there is a predominant diffuse radiation. The application of the inversion algorithm yields the following values for the leaf area index and respective annual foliage increments: 3.5 (0.35 yr. -1 ) for the intact secondary forest; 2.0 (0.5 yr -1 ) for the palm agroforestry system; and 1.6 (0.4 yr -1 ) for the multi-layer ones [pt

  13. Towards ground-truthing of spaceborne estimates of above-ground biomass and leaf area index in tropical rain forests

    Science.gov (United States)

    Köhler, P.; Huth, A.

    2010-05-01

    The canopy height of forests is a key variable which can be obtained using air- or spaceborne remote sensing techniques such as radar interferometry or lidar. If new allometric relationships between canopy height and the biomass stored in the vegetation can be established this would offer the possibility for a global monitoring of the above-ground carbon content on land. In the absence of adequate field data we use simulation results of a tropical rain forest growth model to propose what degree of information might be generated from canopy height and thus to enable ground-truthing of potential future satellite observations. We here analyse the correlation between canopy height in a tropical rain forest with other structural characteristics, such as above-ground biomass (AGB) (and thus carbon content of vegetation) and leaf area index (LAI). The process-based forest growth model FORMIND2.0 was applied to simulate (a) undisturbed forest growth and (b) a wide range of possible disturbance regimes typically for local tree logging conditions for a tropical rain forest site on Borneo (Sabah, Malaysia) in South-East Asia. It is found that for undisturbed forest and a variety of disturbed forests situations AGB can be expressed as a power-law function of canopy height h (AGB=a·hb) with an r2~60% for a spatial resolution of 20 m×20 m (0.04 ha, also called plot size). The regression is becoming significant better for the hectare wide analysis of the disturbed forest sites (r2=91%). There seems to exist no functional dependency between LAI and canopy height, but there is also a linear correlation (r2~60%) between AGB and the area fraction in which the canopy is highly disturbed. A reasonable agreement of our results with observations is obtained from a comparison of the simulations with permanent sampling plot data from the same region and with the large-scale forest inventory in Lambir. We conclude that the spaceborne remote sensing techniques have the potential to

  14. Leaf-jams - A new and unique leaf deposit in the ephemeral Hoanib River, NW Namibia: Origin and plant taphonomic implications

    Energy Technology Data Exchange (ETDEWEB)

    Hofmann, Christa-Ch. [University of Vienna, Department of Palaeontology, Palaeobotany Studies Group, Althanstrasse 14, 1090, Vienna (Austria); Rice, A. Hugh N. [University of Vienna, Department of Geodynamics and Sedimentology, Althanstrasse 14, 1090, Vienna (Austria)

    2010-08-01

    This paper documents a previously unrecorded type of leaf deposit, comprising essentially monospecific linear accumulations of Colophospermum mopane leaves on a point bar of the ephemeral Hoanib River, NW Namibia. In these 'leaf-jams', leaf laminae stand on edge, orientated more-or-less normal to bedding. Leaf-jams, which formed upstream of cobbles, clumps of grass and sticks wedged against the former two, were orientated subparallel to the adjacent meandering river-bed, such that over the 40 m of their occurrence, their mean azimuth changed by 59 anticlockwise downstream. The longest leaf-jam was 50 cm and contained approximately 500 leaves, as well as grass culms, twigs (C. mopane, Tamarix usneoides and unidentified) and medium- to fine-grained sand and silt. Individual leaf-jams were partially buried in the point bar sediments up to a depth of 3 cm. Leaf-jam formation occurred in the austral summer of 2006, during the waning stage of a major flood caused by anomalous tropical to extra-tropical storms. Their monospecifity is due to the overwhelming preponderance of the zonal taxon C. mopane in the catchment area, although the Khowarib Gorge contains a quite diverse azonal plant association due to the presence of a permanent water-seep. During leaf-jam formation, the water depth was less than the height of the cobbles (0.1 m), with stream flow-rates competent to transport medium-grained sand (velocity estimated at 0.5 m s{sup -} {sup 1}). Leaves must have been partially or fully waterlogged to inhibit buoyancy forces tending to lift them out of the developing leaf-jams, which propagated upstream in a manner comparable to longitudinal bars in a braided river. If fossilised, such deposits would probably lead to a very biased interpretation of the composition of the surrounding flora; the correct interpretation would be the one least favoured by palaeobotanists. (author)

  15. The influence of branch order on optimal leaf vein geometries: Murray's law and area preserving branching.

    Directory of Open Access Journals (Sweden)

    Charles A Price

    Full Text Available Models that predict the form of hierarchical branching networks typically invoke optimization based on biomechanical similitude, the minimization of impedance to fluid flow, or construction costs. Unfortunately, due to the small size and high number of vein segments found in real biological networks, complete descriptions of networks needed to evaluate such models are rare. To help address this we report results from the analysis of the branching geometry of 349 leaf vein networks comprising over 1.5 million individual vein segments. In addition to measuring the diameters of individual veins before and after vein bifurcations, we also assign vein orders using the Horton-Strahler ordering algorithm adopted from the study of river networks. Our results demonstrate that across all leaves, both radius tapering and the ratio of daughter to parent branch areas for leaf veins are in strong agreement with the expectation from Murray's law. However, as veins become larger, area ratios shift systematically toward values expected under area-preserving branching. Our work supports the idea that leaf vein networks differentiate roles of leaf support and hydraulic supply between hierarchical orders.

  16. Determination of coefficient defining leaf area development in different genotypes, plant types and planting densities in peanut (Arachis hypogeae L.).

    Science.gov (United States)

    Halilou, Oumarou; Hissene, Halime Mahamat; Clavijo Michelangeli, José A; Hamidou, Falalou; Sinclair, Thomas R; Soltani, Afshin; Mahamane, Saadou; Vadez, Vincent

    2016-12-01

    Rapid leaf area development may be attractive under a number of cropping conditions to enhance the vigor of crop establishment and allow rapid canopy closure for maximizing light interception and shading of weed competitors. This study was undertaken to determine (1) if parameters describing leaf area development varied among ten peanut ( Arachis hypogeae L.) genotypes grown in field and pot experiments, (2) if these parameters were affected by the planting density, and (3) if these parameters varied between Spanish and Virginia genotypes. Leaf area development was described by two steps: prediction of main stem number of nodes based on phyllochron development and plant leaf area dependent based on main stem node number. There was no genetic variation in the phyllochron measured in the field. However, the phyllochron was much longer for plants grown in pots as compared to the field-grown plants. These results indicated a negative aspect of growing peanut plants in the pots used in this experiment. In contrast to phyllochron, there was no difference in the relationship between plant leaf area and main stem node number between the pot and field experiments. However, there was genetic variation in both the pot and field experiments in the exponential coefficient (PLAPOW) of the power function used to describe leaf area development from node number. This genetic variation was confirmed in another experiment with a larger number of genotypes, although possible G × E interaction for the PLAPOW was found. Sowing density did not affect the power function relating leaf area to main stem node number. There was also no difference in the power function coefficient between Spanish and Virginia genotypes. SSM (Simple Simulation model) reliably predicted leaf canopy development in groundnut. Indeed the leaf area showed a close agreement between predicted and observed values up to 60000 cm 2  m -2 . The slightly higher prediction in India and slightly lower prediction in

  17. SAFARI 2000 Leaf Area Measurements at the Mongu Tower Site, Zambia, 2000-2002

    Data.gov (United States)

    National Aeronautics and Space Administration — ABSTRACT: Data from the LAI-2000 instrument were processed to determine the leaf area index (LAI) at the EOS Validation Core Site in Kataba Local Forest,...

  18. Leaf area index, biomass carbon and growth rate of radiata pine genetic types and relationships with LiDAR

    Science.gov (United States)

    Peter N. Beets; Stephen Reutebuch; Mark O. Kimberley; Graeme R. Oliver; Stephen H. Pearce; Robert J. McGaughey

    2011-01-01

    Relationships between discrete-return light detection and ranging (LiDAR) data and radiata pine leaf area index (LAI), stem volume, above ground carbon, and carbon sequestration were developed using 10 plots with directly measured biomass and leaf area data, and 36 plots with modelled carbon data. The plots included a range of genetic types established on north- and...

  19. Indirect estimations and spatial variation in leaf area index of coniferous, deciduous and mixed forest stands in Forsmark and Laxemar

    International Nuclear Information System (INIS)

    Tagesson, Torbern

    2006-12-01

    Two sites in Sweden are investigated for a potential deep repository of the nuclear waste, the Laxemar investigation area (57 deg 5 min N, 16 deg 7 min E) and the Forsmark investigation area (60 deg 4 min N, 18 deg 2 min E). In the characterisation of these sites, development of site descriptive models is an important part. Leaves are the main surface were an exchange of matter and energy between the atmosphere and the biosphere takes place, and leaf area index (LAI) of the vegetation cover is an important variable correlated to a number of ecophysiological parameters and hereby an important parameter in ecosystem models. In the investigation areas, LAI of boreal and temperate ecosystems were therefore estimated indirectly through optical measurements using the LAI-2000 (LI-COR, Cambridge UK) and TRAC (Tracing Radiation and Architecture of Canopies). On average, measured maximum LAI was 3.40 in Laxemar and 3.43 in Forsmark; minimum LAI was 1.65 in Laxemar and 1.97 in Forsmark. Forest inventory data showed that LAI is positively correlated with basal area, stand height, stand volume and breast height tree diameter. For the coniferous stands, there was also a linearly negative relationship with age. In the Laxemar investigation area, there were no significant relationships for LAI with a satellite derived kNN (kNearest Neighbor) data set with stand height, stand volume and stand age. The kNN data set can therefore not be used to extrapolate measured LAI over the Laxemar investigation area. There were significant relationships between LAI and the normalized difference vegetation index (NDVI) for coniferous, deciduous and mixed forest stands in the Laxemar investigation area. A NDVI image could be used to extrapolate LAI over the entire investigation area. For the Forsmark investigation area, effective LAI for all stands were correlated to NDVI and this relationship could then be used for extrapolation. The effective LAI image was afterwards corrected for average

  20. Indirect estimations and spatial variation in leaf area index of coniferous, deciduous and mixed forest stands in Forsmark and Laxemar

    Energy Technology Data Exchange (ETDEWEB)

    Tagesson, Torbern [Dept. of Physical Geography and Ecosystem Analysis, Lund Univ., Lund (Sweden)

    2006-12-15

    Two sites in Sweden are investigated for a potential deep repository of the nuclear waste, the Laxemar investigation area (57 deg 5 min N, 16 deg 7 min E) and the Forsmark investigation area (60 deg 4 min N, 18 deg 2 min E). In the characterisation of these sites, development of site descriptive models is an important part. Leaves are the main surface were an exchange of matter and energy between the atmosphere and the biosphere takes place, and leaf area index (LAI) of the vegetation cover is an important variable correlated to a number of ecophysiological parameters and hereby an important parameter in ecosystem models. In the investigation areas, LAI of boreal and temperate ecosystems were therefore estimated indirectly through optical measurements using the LAI-2000 (LI-COR, Cambridge UK) and TRAC (Tracing Radiation and Architecture of Canopies). On average, measured maximum LAI was 3.40 in Laxemar and 3.43 in Forsmark; minimum LAI was 1.65 in Laxemar and 1.97 in Forsmark. Forest inventory data showed that LAI is positively correlated with basal area, stand height, stand volume and breast height tree diameter. For the coniferous stands, there was also a linearly negative relationship with age. In the Laxemar investigation area, there were no significant relationships for LAI with a satellite derived kNN (kNearest Neighbor) data set with stand height, stand volume and stand age. The kNN data set can therefore not be used to extrapolate measured LAI over the Laxemar investigation area. There were significant relationships between LAI and the normalized difference vegetation index (NDVI) for coniferous, deciduous and mixed forest stands in the Laxemar investigation area. A NDVI image could be used to extrapolate LAI over the entire investigation area. For the Forsmark investigation area, effective LAI for all stands were correlated to NDVI and this relationship could then be used for extrapolation. The effective LAI image was afterwards corrected for average

  1. A method for reconstructing the development of the sapwood area of balsam fir.

    Science.gov (United States)

    Coyea, M R; Margolis, H A; Gagnon, R R

    1990-09-01

    Leaf area is commonly estimated as a function of sapwood area. However, because sapwood changes to heartwood over time, it has not previously been possible to reconstruct either the sapwood area or the leaf area of older trees into the past. In this study, we report a method for reconstructing the development of the sapwood area of dominant and codominant balsam fir (Abies balsamea (L.) Mill.). The technique is based on establishing a species-specific relationship between the number of annual growth rings in the sapwood area and tree age. Because the number of annual growth rings in the sapwood of balsam fir at a given age was found to be independent of site quality and stand density, the number of rings in sapwood (NRS) can be predicted from the age of a tree thus: NRS = 14.818 (1 - e(-0.031 age)), unweighted R(2) = 0.80, and NRS = 2.490 (1 - e(-0.038 age)), unweighted R(2) = 0.64, for measurements at breast height and at the base of the live crown, respectively. These nonlinear asymptotic regression models based only on age, were not improved by adding other tree variables such as diameter at breast height, diameter at the base of the live crown, total tree height or percent live crown.

  2. Analysis, improvement and application of the MODIS leaf area index products

    Science.gov (United States)

    Yang, Wenze

    Green leaf area governs the exchanges of energy, mass and momentum between the Earth's surface and the atmosphere. Therefore, leaf area index (LAI) and fraction of incident photosynthetically active radiation (0.4-0.7 mum) absorbed by the vegetation canopy (FPAR) are widely used in vegetation monitoring and modeling. The launch of Terra and Aqua satellites with the moderate resolution imaging spectroradiometer (MODIS) instrument onboard provided the first global products of LAI and FPAR, derived mainly from an algorithm based on radiative transfer. The objective of this research is to comprehensively evaluate the Terra and Aqua MODIS LAI/FPAR products. Large volumes of these products have been analyzed with the goal of understanding product quality with respect to version (Collection 3 versus 4), algorithm (main versus back-up), snow (snow-free versus snow on the ground) and cloud (cloud-free versus cloudy) conditions. Field validation efforts identified several key factors that influence the accuracy of algorithm retrievals. The strategy of validation efforts guiding algorithm refinements has led to progressively more accurate LAI/FPAR products. The combination of products derived from the Terra and Aqua MODIS sensors increases the success rate of the main radiative transfer algorithm by 10-20 percent over woody vegetation. The Terra Collection 4 LAI data reveal seasonal swings in green leaf area of about 25 percent in a majority of the Amazon rainforests caused by variability in cloud cover and light. The timing and the influence of this seasonal cycle are critical to understanding tropical plant adaptation patterns and ecological processes. The results presented in this dissertation suggest how the product quality has gradually improved largely through the efforts of validation activities. The Amazon case study highlights the utility of these data sets for monitoring global vegetation dynamics. Thus, these results can be seen as a benchmark for evaluation of

  3. Multiyear Multiseasonal Changes in Leaf and Canopy Traits Measured by AVIRIS over Ecosystems with Different Functional Type Characteristics Through the Progressive California Drought 2013-2015

    Science.gov (United States)

    Ustin, S.; Roth, K. L.; Huesca, M.; Casas, A.; Adeline, K.; Drewry, D.; Koltunov, A.; Ramirez, C.

    2015-12-01

    Given the known heterogeneity in ecological processes within plant communities in California, we questioned whether the concept of conventional plant functional types (cPFTs) was adequate to characterize the functionality of the dominant species in these communities. We examined seasonal (spring, summer, fall) airborne AVIRIS and MASTER imagery collected during three years of progressive drought in California, and airborne LiDAR acquired once, for ecosystems that represent a wide range of plant functional types, from annual agriculture and herbaceous perennial wetlands, to forests and shrublands, including broadleaf deciduous and evergreen species and conifer species. These data were used to determine the extent to which changes in canopy chemistry could be detected, quantified, and related to leaf and canopy traits that are indicators of physiological functioning (water content, Leaf Mass Area, total C, N, and pigments (chlorophyll a, b, and carotenoids). At the canopy scale we measured leaf area index, and for forests — species, height, canopy area, DBH, deciduous or evergreen, broadleaf or needleleaf, and gap size. Strong correlations between leaf and canopy traits were predictable and quantifiable from spectroscopy data. Key structural properties of canopy height, biomass and complexity, a measure of spatial and vertical heterogeneity, were predicted by AVIRIS and validated against LiDAR data. Our data supports the hypothesis that optical sensors provide more detailed information about the distribution and variability in leaf and canopy traits related to plant functionality than cPFTs.

  4. First direct landscape-scale measurement of tropical rain forest Leaf Area Index, a key driver of global primary productivity

    Science.gov (United States)

    David B. Clark; Paulo C. Olivas; Steven F. Oberbauer; Deborah A. Clark; Michael G. Ryan

    2008-01-01

    Leaf Area Index (leaf area per unit ground area, LAI) is a key driver of forest productivity but has never previously been measured directly at the landscape scale in tropical rain forest (TRF). We used a modular tower and stratified random sampling to harvest all foliage from forest floor to canopy top in 55 vertical transects (4.6 m2) across 500 ha of old growth in...

  5. Diagnostic test of predicted height model in Indonesian elderly: a study in an urban area

    Directory of Open Access Journals (Sweden)

    Fatmah Fatmah

    2010-08-01

    Full Text Available Aim In an anthropometric assessment, elderly are frequently unable to measure their height due to mobility and skeletal deformities. An alternative is to use a surrogate value of stature from arm span, knee height, and sitting height. The equations developed for predicting height in Indonesian elderly using these three predictors. The equations put in the nutritional assessment card (NSA of older people. Before the card which is the first new technology in Indonesia will be applied in the community, it should be tested. The study aimed was to conduct diagnostic test of predicted height model in the card compared to actual height.Methods Model validation towards 400 healthy elderly conducted in Jakarta City with cross-sectional design. The study was the second validation test of the model besides Depok City representing semi urban area which was undertaken as the first study.Result Male elderly had higher mean age, height, weight, arm span, knee height, and sitting height as compared to female elderly. The highest correlation between knee height and standing height was similar in women (r = 0.80; P < 0.001 and men (r = 0.78; P < 0.001, and followed by arm span and sitting height. Knee height had the lowest difference with standing height in men (3.13 cm and women (2.79 cm. Knee height had the biggest sensitivity (92.2%, and the highest specificity on sitting height (91.2%.Conclusion Stature prediction equation based on knee-height, arm span, and sitting height are applicable for nutritional status assessment in Indonesian elderly. (Med J Indones 2010;19:199-204Key words: diagnostic test, elderly, predicted height model

  6. Mapping Forest Canopy Height Across Large Areas by Upscaling ALS Estimates with Freely Available Satellite Data

    Directory of Open Access Journals (Sweden)

    Phil Wilkes

    2015-09-01

    Full Text Available Operational assessment of forest structure is an on-going challenge for land managers, particularly over large, remote or inaccessible areas. Here, we present an easily adopted method for generating a continuous map of canopy height at a 30 m resolution, demonstrated over 2.9 million hectares of highly heterogeneous forest (canopy height 0–70 m in Victoria, Australia. A two-stage approach was utilized where Airborne Laser Scanning (ALS derived canopy height, captured over ~18% of the study area, was used to train a regression tree ensemble method; random forest. Predictor variables, which have a global coverage and are freely available, included Landsat Thematic Mapper (Tasselled Cap transformed, Moderate Resolution Imaging Spectroradiometer Normalized Difference Vegetation Index time series, Shuttle Radar Topography Mission elevation data and other ancillary datasets. Reflectance variables were further processed to extract additional spatial and temporal contextual and textural variables. Modeled canopy height was validated following two approaches; (i random sample cross validation; and (ii with 108 inventory plots from outside the ALS capture extent. Both the cross validation and comparison with inventory data indicate canopy height can be estimated with a Root Mean Square Error (RMSE of ≤ 31% (~5.6 m at the 95th percentile confidence interval. Subtraction of the systematic component of model error, estimated from training data error residuals, rescaled canopy height values to more accurately represent the response variable distribution tails e.g., tall and short forest. Two further experiments were carried out to test the applicability and scalability of the presented method. Results suggest that (a no improvement in canopy height estimation is achieved when models were constructed and validated for smaller geographic areas, suggesting there is no upper limit to model scalability; and (b training data can be captured over a small

  7. Estimation of leaf area index in cereal crops using red-green images

    DEFF Research Database (Denmark)

    Kirk, Kristian; Andersen, Hans Jørgen; Thomsen, Anton G

    2009-01-01

    A new method for estimating the leaf area index (LAI) in cereal crops based on red-green images taken from above the crop canopy is introduced. The proposed method labels pixels into vegetation and soil classes using a combination of greenness and intensity derived from the red and green colour b...

  8. Floristic-functional variation of tree component along an altitudinal gradient in araucaria forest areas, in Southern Brazil.

    Science.gov (United States)

    Soboleski, Vanessa F; Higuchi, Pedro; Silva, Ana Carolina DA; Silva, Mariele A F DA; Nunes, Amanda S; Loebens, Rodineli; Souza, Karine DE; Ferrari, Jheniffer; Lima, Carla L; Kilca, Ricardo V

    2017-01-01

    We aimed to investigate the taxonomic and functional variations of tree component of Araucaria Forest (AF) areas located along an altitudinal gradient (700, 900 and 1,600 m asl), in the southern region of Brazil. The functional traits determined were leaf area, specific leaf area, wood density, maximum potential height and dispersal syndromes and deciduousness. The data were analyzed through a functional and taxonomic dissimilarity dendrograms, community-weighted mean trait values, parametric and nonparametric tests, and Principal Component Analysis. The largest floristic-structural similarity was observed between the lower altitude areas (700 and 900 m asl), whose Bray-Curtis distance was 0.63. The area at 700 m asl was characterized by a predominance of deciduous and semi-deciduous species, with a high number of self- and wind-dispersed species, whereas the area at 1,600 m asl exhibited a predominance of animal-dispersed and evergreen species. It was also observed that there were significant variations for leaf traits, basic wood density and maximum potential height. Over all altitudinal gradient, the ordinations indicated that there was no evidence of functional differentiation among dispersal and deciduousness groups. In conclusion, the evaluated Araucaria Forest areas presented high floristic-functional variation of the tree component along the altitudinal gradient.

  9. Estimativa da área foliar de Sida cordifolia e Sida rhombifolia usando dimensões lineares do limbo foliar Estimate of Sida cordifolia and Sida rhombifolia leaf area using leaf blade linear dimensions

    Directory of Open Access Journals (Sweden)

    S. Bianco

    2008-01-01

    Full Text Available A estimativa da área foliar pode auxiliar na compreensão de relações de interferência entre plantas daninhas e cultivadas. Com o objetivo de obter uma equação que, por meio de parâmetros lineares dimensionais das folhas, permita a estimativa da área foliar de Sida cordifolia e Sida rhombifolia, estudaram-se as correlações entre área foliar real (Af e parâmetros dimensionais do limbo foliar, como o comprimento (C ao longo da nervura principal e a largura máxima (L perpendicular à nervura principal. Foram analisados 200 limbos foliares de cada espécie, coletados em diferentes agroecossistemas na Universidade Estadual Paulista, campus de Jaboticabal. Os modelos estatísticos utilizados foram linear: Y = a + bx; linear simples: Y = bx; geométrico: Y = ax b; e exponencial: Y = ab x. Todos os modelos analisados podem ser empregados para estimação da área foliar de S. cordifolia e S. rhombifolia. Sugere-se optar pela equação linear simples, envolvendo o produto C*L, considerando-se o coeficiente linear igual a zero, em função da praticidade desta. Desse modo, a estimativa da área foliar de S. cordifolia pode ser obtida pela fórmula Af = 0,7878*(C*L, com coeficiente de determinação de 0,9307, enquanto para S. rhombifolia a estimativa da área foliar pode ser obtida pela fórmula Af = 0,6423*(C*L, com coeficiente de determinação de 0,9711.Leaf area estimate may contribute to understand the relationship of interference between weeds and crops. The objective of this research was to obtain a mathematical equation to estimate Sida cordifolia and Sida rhombifolia leaf area based on linear measures of leaf blade. Correlation studies were conducted between real leaf area (Af and dimensional leaf blade parameters such as leaf length (C and maximum leaf width (L. Around 200 leaf blades of each species were analyzed, collected from several agro-ecosystems at São Paulo State University, in Jaboticabal, SP, Brazil. The statistical

  10. Leaf litter production of mahogany along street and campus forest of Universitas Negeri Semarang, Indonesia

    Science.gov (United States)

    Martin, F. P.; Abdullah, M.; Solichin; Hadiyanti, L. N.; Widianingrum, K.

    2018-03-01

    The leaf litter of trees along the existing streets on campus UNNES if not managed properly will be scattered and become garbage. Leaf litter Production in UNNES campus is not known for certain. UNNES does not own mapping of leaf litter Production of dominant tree species on campus. This cause leaf waste management is not optimal yet. There is still a lot of leaf litter that is discharged (not processed) because it exceeds the capacity of the fertilizer production equipment in the compost house. Aims of this study were to examine leaf litter production of dominant trees in Universitas Negeri Semarang and evaluate the relationship between leaf litter and average rainfall. Purposive sampling method placed pouches of nylon gauze measuring 1 × 1 mm2 as litter trap container with size 1 x l m2 (10 points mounted along street and campus forest). Litter trap mounted at the height of 50 cm above ground level. Leaf litter will be taken once a week for three months to observe the litter production. The litter was then dried by the oven at 70 ° C for 48 hours to obtain constant dry weight. Based on the results of the research, it was known that Mahogany tree in UNNES campus area has the potential to produce the litter of about 10 ton/ha / 3months in campus forest area and 2.5 ton/ha / 3months along campus street. There is a significant relationship between litter production of Mahogany leaves and precipitation during August - October 2017.

  11. LCE: leaf carbon exchange data set for tropical, temperate, and boreal species of North and Central America.

    Science.gov (United States)

    Smith, Nicholas G; Dukes, Jeffrey S

    2017-11-01

    Leaf canopy carbon exchange processes, such as photosynthesis and respiration, are substantial components of the global carbon cycle. Climate models base their simulations of photosynthesis and respiration on an empirical understanding of the underlying biochemical processes, and the responses of those processes to environmental drivers. As such, data spanning large spatial scales are needed to evaluate and parameterize these models. Here, we present data on four important biochemical parameters defining leaf carbon exchange processes from 626 individuals of 98 species at 12 North and Central American sites spanning ~53° of latitude. The four parameters are the maximum rate of Rubisco carboxylation (V cmax ), the maximum rate of electron transport for the regeneration of Ribulose-1,5,-bisphosphate (J max ), the maximum rate of phosphoenolpyruvate carboxylase carboxylation (V pmax ), and leaf dark respiration (R d ). The raw net photosynthesis by intercellular CO 2 (A/C i ) data used to calculate V cmax , J max , and V pmax rates are also presented. Data were gathered on the same leaf of each individual (one leaf per individual), allowing for the examination of each parameter relative to others. Additionally, the data set contains a number of covariates for the plants measured. Covariate data include (1) leaf-level traits (leaf mass, leaf area, leaf nitrogen and carbon content, predawn leaf water potential), (2) plant-level traits (plant height for herbaceous individuals and diameter at breast height for trees), (3) soil moisture at the time of measurement, (4) air temperature from nearby weather stations for the day of measurement and each of the 90 d prior to measurement, and (5) climate data (growing season mean temperature, precipitation, photosynthetically active radiation, vapor pressure deficit, and aridity index). We hope that the data will be useful for obtaining greater understanding of the abiotic and biotic determinants of these important biochemical

  12. Towards ground-truthing of spaceborne estimates of above-ground life biomass and leaf area index in tropical rain forests

    Directory of Open Access Journals (Sweden)

    P. Köhler

    2010-08-01

    Full Text Available The canopy height h of forests is a key variable which can be obtained using air- or spaceborne remote sensing techniques such as radar interferometry or LIDAR. If new allometric relationships between canopy height and the biomass stored in the vegetation can be established this would offer the possibility for a global monitoring of the above-ground carbon content on land. In the absence of adequate field data we use simulation results of a tropical rain forest growth model to propose what degree of information might be generated from canopy height and thus to enable ground-truthing of potential future satellite observations. We here analyse the correlation between canopy height in a tropical rain forest with other structural characteristics, such as above-ground life biomass (AGB (and thus carbon content of vegetation and leaf area index (LAI and identify how correlation and uncertainty vary for two different spatial scales. The process-based forest growth model FORMIND2.0 was applied to simulate (a undisturbed forest growth and (b a wide range of possible disturbance regimes typically for local tree logging conditions for a tropical rain forest site on Borneo (Sabah, Malaysia in South-East Asia. In both undisturbed and disturbed forests AGB can be expressed as a power-law function of canopy height h (AGB = a · hb with an r2 ~ 60% if data are analysed in a spatial resolution of 20 m × 20 m (0.04 ha, also called plot size. The correlation coefficient of the regression is becoming significant better in the disturbed forest sites (r2 = 91% if data are analysed hectare wide. There seems to exist no functional dependency between LAI and canopy height, but there is also a linear correlation (r2 ~ 60% between AGB and the area fraction of gaps in which the canopy is highly disturbed. A reasonable agreement of our results with observations is obtained from a

  13. Towards ground-truthing of spaceborne estimates of above-ground life biomass and leaf area index in tropical rain forests

    Science.gov (United States)

    Köhler, P.; Huth, A.

    2010-08-01

    The canopy height h of forests is a key variable which can be obtained using air- or spaceborne remote sensing techniques such as radar interferometry or LIDAR. If new allometric relationships between canopy height and the biomass stored in the vegetation can be established this would offer the possibility for a global monitoring of the above-ground carbon content on land. In the absence of adequate field data we use simulation results of a tropical rain forest growth model to propose what degree of information might be generated from canopy height and thus to enable ground-truthing of potential future satellite observations. We here analyse the correlation between canopy height in a tropical rain forest with other structural characteristics, such as above-ground life biomass (AGB) (and thus carbon content of vegetation) and leaf area index (LAI) and identify how correlation and uncertainty vary for two different spatial scales. The process-based forest growth model FORMIND2.0 was applied to simulate (a) undisturbed forest growth and (b) a wide range of possible disturbance regimes typically for local tree logging conditions for a tropical rain forest site on Borneo (Sabah, Malaysia) in South-East Asia. In both undisturbed and disturbed forests AGB can be expressed as a power-law function of canopy height h (AGB = a · hb) with an r2 ~ 60% if data are analysed in a spatial resolution of 20 m × 20 m (0.04 ha, also called plot size). The correlation coefficient of the regression is becoming significant better in the disturbed forest sites (r2 = 91%) if data are analysed hectare wide. There seems to exist no functional dependency between LAI and canopy height, but there is also a linear correlation (r2 ~ 60%) between AGB and the area fraction of gaps in which the canopy is highly disturbed. A reasonable agreement of our results with observations is obtained from a comparison of the simulations with permanent sampling plot (PSP) data from the same region and with the

  14. BigFoot Leaf Area Index Surfaces for North and South American Sites, 2000-2003

    Data.gov (United States)

    National Aeronautics and Space Administration — ABSTRACT: The BigFoot project gathered leaf area index (LAI) data for nine EOS Land Validation Sites located from Alaska to Brazil from 2000 to 2003. Each site is...

  15. Estimation of leaf area in coffee leaves (Coffea arabica L. of the Castillo® variety

    Directory of Open Access Journals (Sweden)

    Carlos Andrés Unigarro-Muñoz

    2015-01-01

    Full Text Available Allometric models based on measurements of single leaf dimensions or a combination there are useful tools for determining individual leaf area (LA because they are non-destructive, precise, simple and economical methods. The present study was carried out at the Central Station Naranjal of Cenicafé, located in the Department of Caldas (Colombia, four models were defined using the variables length (L and/or width (W to estimate LA in coffee leaves of the Castillo® variety (Coffea arabica L.. Estimation of regression coefficients was performed using information recorded from 6,441 leaves (group 1, and their validation was performed using records from another 992 leaves (group 2. Leaves were collected from all strata of the canopy and ranged from 0.76 to 140 cm2 in LA. In addition to exhibiting coefficients of variation differing from zero based on t-tests at 1%, the evaluated models possess coefficients of determination between 0.93 and 0.99. Four expressions have developed and adjusted to estimate leaf area in individual leaves, based on the measurement of simple variables and non-destructive.

  16. Leaf mass per area is independent of vein length per area: avoiding pitfalls when modelling phenotypic integration (reply to Blonder et al. 2014).

    Science.gov (United States)

    Sack, Lawren; Scoffoni, Christine; John, Grace P; Poorter, Hendrik; Mason, Chase M; Mendez-Alonzo, Rodrigo; Donovan, Lisa A

    2014-10-01

    It has been recently proposed that leaf vein length per area (VLA) is the major determinant of leaf mass per area ( MA), and would thereby determine other traits of the leaf economic spectrum (LES), such as photosynthetic rate per mass (A(mass)), nitrogen concentration per mass (N(mass)) and leaf lifespan (LL). In a previous paper we argued that this 'vein origin' hypothesis was supported only by a mathematical model with predestined outcomes, and that we found no support for the 'vein origin' hypothesis in our analyses of compiled data. In contrast to the 'vein origin' hypothesis, empirical evidence indicated that VLA and LMA are independent mechanistically, and VLA (among other vein traits) contributes to a higher photosynthetic rate per area (A(area)), which scales up to driving a higher A(mass), all independently of LMA, N(mass) and LL. In their reply to our paper, Blonder et al. (2014) raised questions about our analysis of their model, but did not address our main point, that the data did not support their hypothesis. In this paper we provide further analysis of an extended data set, which again robustly demonstrates the mechanistic independence of LMA from VLA, and thus does not support the 'vein origin' hypothesis. We also address the four specific points raised by Blonder et al. (2014) regarding our analyses. We additionally show how this debate provides critical guidance for improved modelling of LES traits and other networks of phenotypic traits that determine plant performance under contrasting environments. © The Author 2014. Published by Oxford University Press on behalf of the Society for Experimental Biology.

  17. ESTIMATION OF LEAF AREA INDEX IN OPEN-CANOPY PONDEROSA PINE FORESTS AT DIFFERENT SUCCESSIONAL STAGES AND MANAGEMENT REGIMES IN OREGON. (R828309)

    Science.gov (United States)

    AbstractLeaf area and its spatial distribution are key parameters in describing canopy characteristics. They determine radiation regimes and influence mass and energy exchange with the atmosphere. The evaluation of leaf area in conifer stands is particularly challengi...

  18. Leaf reflectance variation along a vertical crown gradient of two deciduous tree species in a Belgian industrial habitat

    International Nuclear Information System (INIS)

    Khavaninzadeh, Ali Reza; Veroustraete, Frank; Van Wittenberghe, Shari; Verrelst, Jochem; Samson, Roeland

    2015-01-01

    The reflectometry of leaf asymmetry is a novel approach in the bio-monitoring of tree health in urban or industrial habitats. Leaf asymmetry responds to the degree of environmental pollution and reflects structural changes in a leaf due to environmental pollution. This paper describes the boundary conditions to scale up from leaf to canopy level reflectance, by describing the variability of adaxial and abaxial leaf reflectance, hence leaf asymmetry, along the crown height gradients of two tree species. Our findings open a research pathway towards bio-monitoring based on the airborne remote sensing of tree canopies and their leaf asymmetric properties. - Highlights: • Reflectometry of leaf asymmetry is a novel approach in tree health bio-monitoring. • Leaf asymmetry reflects degrees of structural changes by environmental pollution. • Conditions to scale up from leaf to canopy level reflectance are described. • A research pathway is opened towards airborne pollution bio-assessment. - Tree leaf asymmetry responds to the degree of environmental pollution and reflects leaf structural changes differentially according to species and height in the crown

  19. Sapwood Area Related to Tree Size, Tree Age, and Leaf Area Index in Cedrus libani

    OpenAIRE

    Güney, Aylin

    2018-01-01

    Sapwoodincludes the water conducting part of the stem which transports water andminerals from roots to leaves. Studies using sap flow gauges have to determinethe area of the sapwood in order to scale measured sap flow densities to thetree or stand level. The aim of this study was to investigate the relationshipbetween sapwood area at breast height and other tree parameters which are easyto measure of the montane Mediterranean conifer Cedrus libani, including a total number of 92 study trees o...

  20. Morphological analysis of plant density effects on early leaf area growth in maize

    NARCIS (Netherlands)

    Bos, H.J.; Vos, J.; Struik, P.C.

    2000-01-01

    The mechanisms of density-related reduced leaf area per plant in non-tillering maize (Zea mays) were investigated. Maize cv. Luna crops with a wide range of plant densities were grown in the field at Wageningen for two years. Half of the plots were shaded (50% transmittance). Detailed measurements

  1. A phenomics approach to the analysis of the influence of glutathione on leaf area and abiotic stress tolerance in Arabidopsis thaliana

    Directory of Open Access Journals (Sweden)

    Daniel eSchnaubelt

    2013-11-01

    Full Text Available Reduced glutathione (GSH is an abundant low molecular weight plant thiol. It fulfils multiple functions in plant biology, many of which remain poorly characterised. A phenomics approach was therefore used to investigate the effects of glutathione homeostasis on growth and stress tolerance in Arabidopsis thaliana. Rosette leaf area was compared in mutants that are either defective in GSH synthesis (cad2, pad2 and rax1 or the export of γ-glutamyl cysteine and GSH from the chloroplast (clt and in wild type plants under standard growth conditions and following exposure to a range of abiotic stress treatments, including oxidative stress, water stress and high salt. In the absence of stress, the GSH synthesis mutants had a significantly lower leaf area than the wild type. Conversely, the clt mutant has a greater leaf area and a significantly reduced lateral root density than the wild type. These findings demonstrate that cellular glutathione homeostasis exerts an influence on root architecture and on rosette area. An impaired capacity to synthesise GSH or a specific depletion of the cytosolic GSH pool did not adversely affect leaf area in plants exposed to short term abiotic stress. However, the negative effects of long term exposure to oxidative stress and high salt on leaf area were less marked in the GSH synthesis mutants than the wild type. These findings demonstrate the importance of cellular glutathione homeostasis in the regulation of plant growth under optimal and stress conditions.

  2. A phenomics approach to the analysis of the influence of glutathione on leaf area and abiotic stress tolerance in Arabidopsis thaliana.

    Science.gov (United States)

    Schnaubelt, Daniel; Schulz, Philipp; Hannah, Matthew A; Yocgo, Rosita E; Foyer, Christine H

    2013-01-01

    Reduced glutathione (GSH) is an abundant low molecular weight plant thiol. It fulfills multiple functions in plant biology, many of which remain poorly characterized. A phenomics approach was therefore used to investigate the effects of glutathione homeostasis on growth and stress tolerance in Arabidopsis thaliana. Rosette leaf area was compared in mutants that are either defective in GSH synthesis (cad2, pad2, and rax1) or the export of γ-glutamylcysteine and GSH from the chloroplast (clt) and in wild-type plants under standard growth conditions and following exposure to a range of abiotic stress treatments, including oxidative stress, water stress, and high salt. In the absence of stress, the GSH synthesis mutants had a significantly lower leaf area than the wild type. Conversely, the clt mutant has a greater leaf area and a significantly reduced lateral root density than the wild type. These findings demonstrate that cellular glutathione homeostasis exerts an influence on root architecture and on rosette area. An impaired capacity to synthesize GSH or a specific depletion of the cytosolic GSH pool did not adversely affect leaf area in plants exposed to short-term abiotic stress. However, the negative effects of long-term exposure to oxidative stress and high salt on leaf area were less marked in the GSH synthesis mutants than the wild type. These findings demonstrate the importance of cellular glutathione homeostasis in the regulation of plant growth under optimal and stress conditions.

  3. A Global Database of Field-observed Leaf Area Index in Woody Plant Species, 1932-2011

    Data.gov (United States)

    National Aeronautics and Space Administration — ABSTRACT: This data set provides global leaf area index (LAI) values for woody species. The data are a compilation of field-observed data from 1,216 locations...

  4. A Global Data Set of Leaf Photosynthetic Rates, Leaf N and P, and Specific Leaf Area

    Data.gov (United States)

    National Aeronautics and Space Administration — ABSTRACT: This global data set of photosynthetic rates and leaf nutrient traits was compiled from a comprehensive literature review. It includes estimates of Vcmax...

  5. A Global Data Set of Leaf Photosynthetic Rates, Leaf N and P, and Specific Leaf Area

    Data.gov (United States)

    National Aeronautics and Space Administration — This global data set of photosynthetic rates and leaf nutrient traits was compiled from a comprehensive literature review. It includes estimates of Vcmax (maximum...

  6. A Global Database of Field-observed Leaf Area Index in Woody Plant Species, 1932-2011

    Data.gov (United States)

    National Aeronautics and Space Administration — This data set provides global leaf area index (LAI) values for woody species. The data are a compilation of field-observed data from 1,216 locations obtained from...

  7. Combined use of leaf size and economics traits allows direct comparison of hydrophyte and terrestrial herbaceous adaptive strategies.

    Science.gov (United States)

    Pierce, Simon; Brusa, Guido; Sartori, Matteo; Cerabolini, Bruno E L

    2012-04-01

    Hydrophytes generally exhibit highly acquisitive leaf economics. However, a range of growth forms is evident, from small, free-floating and rapidly growing Lemniden to large, broad-leaved Nymphaeiden, denoting variability in adaptive strategies. Traits used to classify adaptive strategies in terrestrial species, such as canopy height, are not applicable to hydrophytes. We hypothesize that hydrophyte leaf size traits and economics exhibit sufficient overlap with terrestrial species to allow a common classification of plant functional types, sensu Grime's CSR theory. Leaf morpho-functional traits were measured for 61 species from 47 water bodies in lowland continental, sub-alpine and alpine bioclimatic zones in southern Europe and compared against the full leaf economics spectrum and leaf size range of terrestrial herbs, and between hydrophyte growth forms. Hydrophytes differed in the ranges and mean values of traits compared with herbs, but principal components analysis (PCA) demonstrated that both groups shared axes of trait variability: PCA1 encompassed size variation (area and mass), and PCA2 ranged from relatively dense, carbon-rich leaves to nitrogen-rich leaves of high specific leaf area (SLA). Most growth forms exhibited trait syndromes directly equivalent to herbs classified as R adapted, although Nymphaeiden ranged between C and SR adaptation. Our findings support the hypothesis that hydrophyte adaptive strategy variation reflects fundamental trade-offs in economics and size that govern all plants, and that hydrophyte adaptive strategies can be directly compared with terrestrial species by combining leaf economics and size traits.

  8. Non-destructive equations to estimate the leaf area of Styrax pohlii and Styrax ferrugineus

    Directory of Open Access Journals (Sweden)

    MC Souza

    Full Text Available We developed linear equations to predict the leaf area (LA of the species Styrax pohlii and Styrax ferrugineus using the width (W and length (L leaf dimensions. For both species the linear regression (Y=α+bX using LA as a dependent variable vs. W × L as an independent variable was more efficient than linear regressions using L, W, L2 and W2 as independent variables. Therefore, the LA of S. pohlii can be estimated with the equation LA=0.582+0.683WL, while the LA of S. ferrugineus follows the equation LA=−0.666+0.704WL.

  9. Measurement and comparison of remotely derived leaf area index predictors

    Science.gov (United States)

    Jensen, Ryan Russell

    Environmental change occurs in response to both natural and anthropogenic causes. As the world's human population continues to increase, anthropogenic change will also increase. These changes affect the health and vigor of forests throughout the world, including those in north central Florida. Leaf Area Index (LAI), the amount of leaf area per unit ground area, is an important biophysical variable that is directly related to rates of atmospheric gas exchange, biomass partitioning, and productivity. While global and local models that map biophysical parameters are prevalent in the literature, landscape to regional scale models are less common. Therefore, the ability to map and monitor LAI over landscape to regional scale areas is essential for understanding medium scale biophysical properties and how these properties affect biogeochemical cycling, biomass accumulation, and primary productivity. This study develops and verifies several new models to estimate LAI using in situ field measurements throughout north central Florida, Landsat Thematic Mapper remotely sensed imagery, remotely derived vegetation indices, simple and multiple regression, and artificial neural networks (ANNs). This study concludes that while multiple band regression and regression with individual vegetation indices (Normalized Difference Vegetation Index, Soil Adjusted Vegetation Index, Simple Ratio, and Greenness Vegetation Index) can estimate LAI, the most accurate way to estimate regional scale LAI is to train an ANN using in situ LAI data and remote sensing brightness values measured from six different portions of the electromagnetic spectrum. The new ANN method of estimating LAI is then applied to two forest ecology studies. The first study analyzes LAI in longleaf pine/turkey oak sandhills as a function of time since last burn. It concludes that in the absence of fire, sandhill LAI increases, and this may be useful for identifying where prescribed burns need to be done. The second study

  10. Investigation the Vertical Distribution of Leaf Area and Dry Matter of Sweet Basil (Ocimum basilicum L., Borage (Borago officinalis L. and Cover Crops in Competition with Weeds

    Directory of Open Access Journals (Sweden)

    zeinab shirzadi margavi

    2017-10-01

    laboratory, leaves and stem were separated and for every sample the area of green leaves was measured with a leaf area meter LICOR- 3000 A (LI-COR, Lincoln, NE, USA. Afterwards all samples were oven-dried at 70 ºC for 48 hours and weighted. Analysis of variance (ANOVA was performed by application SAS (Ver. 9.2. In order to compare the average, least significant differences test (LSD were used. Excel was used for charting. Results and discussion Results presented that vertical distribution of sweet basil leaf area was limited in top layer of canopy (40-60 cm in all treatments except presence of weeds. But dry matter of sweet basil canopy was distributed in all layers in weed free treatment. In borage, omitting the competition with weeds, causes to distribute leaf area and dry matter in all three layers. Results showed that mung bean and Persian clover have same distribution of leaf area and dry matter in all layers in neighboring with sweet basil. Generally, the vertical distribution of dry matter of weeds in the fields of borage and sweet basil had more uniformity in the absence of cover crops. Planting cover crops between rows of main plants causes rapid occupation of empty spaces and lack of weed seed germination and also prevent seedling growth and development of weeds. Conclusion Shading can cause leaf abscission of sweet basil and borage in lower layers when competing with weeds. While, presence of cover crop made dry matter allocation to these lower layers in sweet basil. Moreover, borage allocated its dry matter in presence of Persian clover and also in weed free. No applying cover crops caused weed height increment, so with planting suitable cover crop, weed growing can be decreased.

  11. LBA-ECO CD-04 Leaf Area Index, km 83 Tower Site, Tapajos National Forest, Brazil

    Data.gov (United States)

    National Aeronautics and Space Administration — ABSTRACT: Leaf area index was estimated in an 18 ha plot at the logged forest tower site, km 83, Tapajos National Forest, Para, Brazil. The plot was adjacent to the...

  12. Seasonal variation in specific leaf area, epicuticular wax and pigments in 15 woody species from northeastern mexico during summer and winter

    International Nuclear Information System (INIS)

    Rodriguez, H.G.; Maiti, R.; Kumari, A.

    2017-01-01

    The present study has been undertaken on the variability in specific leaf area, epicuticular wax and pigment content of 15 native woody species in northeastern Mexico. The species showed considerable variability in responses of these leaf traits. Majority of the species showed a decline in specific leaf area and epicuticular wax content. With respect to pigments, only few species showed a decrease, but some species showed an increase in pigments (chlorophyll a, b and total chlorophyll (a+b)) showing mechanism of adaptation to winter season.However, in few species there was a decline in pigment contents showing susceptibility to winter. (author)

  13. DMSA scan nomograms for renal length and area: Related to patient age and to body weight, height or surface area

    International Nuclear Information System (INIS)

    Hassan, I.M.; Que, L.; Rutland, M.D.

    2002-01-01

    Aim: To create nomograms for renal size as measured from DMSA renal studies, and to test the nomograms for their ability to separate normal from abnormal kidneys. Method: Renal length was measured from posterior oblique views and renal area from posterior views. Results from 253 patients with bilateral normal kidneys were used to create nomograms for renal size relative to patient age, body height, weight or body surface area (BSA). The nomograms enclosed 95% of the normal kidneys, thus indicating the range for 95% confidence limits, and hence the specificity. Each nomogram was then tested against 46 hypertrophied kidneys and 46 damaged kidneys. Results: The results from nomograms of renal length and renal area, compared to age, body height, body weight and BSA are presented. For each nomogram, the range is presented as a fraction of the mean value, and the number of abnormal kidneys (hypertrophied or damaged) outside the normal range is presented as a percentage (indicating the sensitivity). Conclusion: Renal Area was no better than renal length for detecting abnormal kidneys. Patient age was the least useful method of normalisation. BSA normalisation produced the best results most frequently (narrower ranges and highest detection of abnormal kidneys)

  14. BIOMONITORING OF URBAN AREA BY ANATOMICAL LEAF CHANGES

    Directory of Open Access Journals (Sweden)

    Elena IRIZA

    2012-01-01

    Full Text Available Plants play a vital role as indicators of pollution. The automobile emissions are high particularly at the traffic intersections. Plants growing under the stress of air pollution show differences in leaf surface characteristics. Light microscopic studies of leaf surface revealed an increase in the number of stomata and trichomes of polluted populations in comparison to control populations of Plantago major and Plantago lanceolata. These changes can be considered as indicators of environmental stress.

  15. LBA-ECO CD-04 Leaf Area Index, km 83 Tower Site, Tapajos National Forest, Brazil

    Data.gov (United States)

    National Aeronautics and Space Administration — Leaf area index was estimated in an 18 ha plot at the logged forest tower site, km 83, Tapajos National Forest, Para, Brazil. The plot was adjacent to the eddy flux...

  16. Seasonal variability of leaf area index and foliar nitrogen in contrasting dry-mesic tundras

    DEFF Research Database (Denmark)

    Campioli, Matteo; Michelsen, Anders; Lemeur, Raoul

    2009-01-01

    Assimilation and exchange of carbon for arctic ecosystems depend strongly on leaf area index (LAI) and total foliar nitrogen (TFN). For dry-mesic tundras, the seasonality of these characteristics is unexplored. We addressed this knowledge gap by measuring variations of LAI and TFN at five contras...

  17. Wave Height Estimation from Shadowing Based on the Acquired X-Band Marine Radar Images in Coastal Area

    Directory of Open Access Journals (Sweden)

    Yanbo Wei

    2017-08-01

    Full Text Available In this paper, the retrieving significant wave height from X-band marine radar images based on shadow statistics is investigated, since the retrieving accuracy can not be seriously affected by environmental factors and the method has the advantage of without any external reference to calibrate. However, the accuracy of the significant wave height estimated from the radar image acquired at the near-shore area is not ideal. To solve this problem, the effect of water depth is considered in the theoretical derivation of estimated wave height based on the sea surface slope. And then, an improved retrieving algorithm which is suitable for both in deep water area and shallow water area is developed. In addition, the radar data are sparsely processed in advance in order to achieve high quality edge image for the requirement of shadow statistic algorithm, since the high resolution radar images will lead to angle-blurred for the image edge detection and time-consuming in the estimation of sea surface slope. The data acquired from Pingtan Test Base in Fujian Province were used to verify the effectiveness of the proposed algorithm. The experimental results demonstrate that the improved method which takes into account the water depth is more efficient and effective and has better performance for retrieving significant wave height in the shallow water area, compared to the in situ buoy data as the ground truth and that of the existing shadow statistic method.

  18. Forage accumulation in brachiaria grass under continuous grazing with single or variable height during the seasons of the year

    Directory of Open Access Journals (Sweden)

    Manoel Eduardo Rozalino Santos

    2013-05-01

    Full Text Available The objective of this study was to evaluate grazing management strategies of Brachiaria decumbens cv. Basilisk managed with different heights under continuous grazing with cattle. Two grazing management strategies were evaluated: maintenance of pasture with an average height of 25 cm throughout the experimental period and maintenance of pasture on the average of 15 cm in height during winter, up to 25 cm from the beginning of spring. The split-plot scheme and the randomized block design with four replications were adopted. The grazing management strategies corresponded to the primary factor, while the seasons (winter, spring and summer corresponded to secondary factor. The reduction of the average sward height to 15 cm in the winter resulted, when compared with pasture maintained at 25 cm, in overall higher growth rates (95 kg/ha.day DM and leaf blade (66.1 kg/ha.day DM, as well as higher rates of total accumulation (81.5 kg/ha.day DM and leaf blade (52.6 kg/ha.day DM. The accumulated forage production (from winter to summer was higher in the pasture lowered to 15 cm in winter (25.6 t/ha DM compared with that managed with an average height of 25 cm (22.2 t/ha DM. Regarding the seasons of the year, in the winter, there were lower rates of overall growth (6.4 kg/ha.day DM, leaf blade (5.6 kg/ha.day DM and pseudostem (0.8 kg/ha.day DM, and also lower total (-6.6 kg/ha.day DM and leaf blade (-7.5 kg/ha.day DM accumulation rates. In the spring there was a higher rate of leaf senescence (22.4 kg/ha.day DM. The accumulation of forage is incremented when the pasture of B. decumbens is lowered to 15 cm during the winter, and in the spring and summer, its average height is increased to 25 cm.

  19. Joint leaf chlorophyll content and leaf area index retrieval from Landsat data using a regularized model inversion system (REGFLEC)

    KAUST Repository

    Houborg, Rasmus

    2015-01-19

    Leaf area index (LAI) and leaf chlorophyll content (Chll) represent key biophysical and biochemical controls on water, energy and carbon exchange processes in the terrestrial biosphere. In combination, LAI and Chll provide critical information on vegetation density, vitality and photosynthetic potentials. However, simultaneous retrieval of LAI and Chll from space observations is extremely challenging. Regularization strategies are required to increase the robustness and accuracy of retrieved properties and enable more reliable separation of soil, leaf and canopy parameters. To address these challenges, the REGularized canopy reFLECtance model (REGFLEC) inversion system was refined to incorporate enhanced techniques for exploiting ancillary LAI and temporal information derived from multiple satellite scenes. In this current analysis, REGFLEC is applied to a time-series of Landsat data.A novel aspect of the REGFLEC approach is the fact that no site-specific data are required to calibrate the model, which may be run in a largely automated fashion using information extracted entirely from image-based and other widely available datasets. Validation results, based upon in-situ LAI and Chll observations collected over maize and soybean fields in central Nebraska for the period 2001-2005, demonstrate Chll retrieval with a relative root-mean-square-deviation (RMSD) on the order of 19% (RMSD=8.42μgcm-2). While Chll retrievals were clearly influenced by the version of the leaf optical properties model used (PROSPECT), the application of spatio-temporal regularization constraints was shown to be critical for estimating Chll with sufficient accuracy. REGFLEC also reproduced the dynamics of in-situ measured LAI well (r2 =0.85), but estimates were biased low, particularly over maize (LAI was underestimated by ~36 %). This disparity may be attributed to differences between effective and true LAI caused by significant foliage clumping not being properly accounted for in the canopy

  20. Validation of Leaf Area Index measurements based on the Wireless Sensor Network platform

    Science.gov (United States)

    Song, Q.; Li, X.; Liu, Q.

    2017-12-01

    The leaf area index (LAI) is one of the important parameters for estimating plant canopy function, which has significance for agricultural analysis such as crop yield estimation and disease evaluation. The quick and accurate access to acquire crop LAI is particularly vital. In the study, LAI measurement of corn crops is mainly through three kinds of methods: the leaf length and width method (LAILLW), the instruments indirect measurement method (LAII) and the leaf area index sensor method(LAIS). Among them, LAI value obtained from LAILLW can be regarded as approximate true value. LAI-2200,the current widespread LAI canopy analyzer,is used in LAII. LAIS based on wireless sensor network can realize the automatic acquisition of crop images,simplifying the data collection work,while the other two methods need person to carry out field measurements.Through the comparison of LAIS and other two methods, the validity and reliability of LAIS observation system is verified. It is found that LAI trend changes are similar in three methods, and the rate of change of LAI has an increase with time in the first two months of corn growth when LAIS costs less manpower, energy and time. LAI derived from LAIS is more accurate than LAII in the early growth stage,due to the small blade especially under the strong light. Besides, LAI processed from a false color image with near infrared information is much closer to the true value than true color picture after the corn growth period up to one and half months.

  1. Características morfogênicas e estruturais de perfilhos de capim-braquiária em locais do pasto com alturas variáveis Morphogenic and structural characteristics of tillers on areas with signalgrass pasture varying on height

    Directory of Open Access Journals (Sweden)

    Manoel Eduardo Rozalino Santos

    2011-03-01

    replications. Leaf appearance rate of signalgrass was linearly and negatively influenced by heights of the plants on the same pasture. The tillers on the same sites with the highest heights showed greater phyllochron (9.3 days. The higher plant heights on the same pasture promoted both higher leaf senescence and pseudoculm elongation rates for the tillers. Leaf lifespan (37.8 days and leaf elongation rate (1.3 cm/tillers.day were not influenced by the plant height. The number of green (4.36 and dead (2.15 leaves was not changed by the plants height either. However, the higher height of plants resulted in a linear increase of 69% on the number of leaves showing defoliation. Pseudoculm and leaf blade lengths responded linearly and positively to the increase in the plants height on the same pasture. The natural height variance of plants found on the same pasture changes the morphogenetic and structural characteristics of signalgrass. There is spatial variability of the vegetation on Brachiaria decumbens cv. Basilisk pastures managed under continuous stocking with bovine.

  2. Use of remotely sensed precipitation and leaf area index in a distributed hydrological model

    DEFF Research Database (Denmark)

    Andersen, J.; Dybkjær, G.; Jensen, Karsten Høgh

    2002-01-01

    Remotely sensed precipitation from METEOSAT data and leaf area index (LAI) from NOAA AVHRR data is used as input data to the distributed hydrological modelling of three sub catchments (82.000 km(2)) in the Senegal River Basin. Further, root depths of annual vegetation are related to the temporal...

  3. Estimation of leaf area index in the sunflower as a function of thermal time1

    Directory of Open Access Journals (Sweden)

    Dioneia Daiane Pitol Lucas

    Full Text Available The aim of this study was to obtain a mathematical model for estimating the leaf area index (LAI of a sunflower crop as a function of accumulated thermal time. Generating the models and testing their coefficients was carried out using data obtained from experiments carried out for different sowing dates in the crop years of 2007/08, 2008/09, 2009/10 and 2010/11 with two sunflower hybrids, Aguará 03 and Hélio 358. Linear leaf dimensions were used for the non-destructive measurement of the leaf area, and thermal time was used to quantify the biological time. With the data for accumulated thermal time (TTa and LAI known for any one day after emergence, mathematical models were generated for estimating the LAI. The following models were obtained, as they presented the best fit (lowest rootmean- square error, RMSE: gaussian peak, cubic polynomial, sigmoidal and an adjusted compound model, the modified sigmoidal. The modified sigmoidal model had the best fit to the generation data and the highest value for the coefficient of determination (R2. In testing the models, the lowest values for root-mean-square error, and the highest R2 between the observed and estimated values were obtained with the modified sigmoidal model.

  4. Relationship between fruit weight and the fruit-to-leaf area ratio, at the spur and whole-tree level, for three sweet cherry varieties

    NARCIS (Netherlands)

    Cittadini, E.D.; Ridder, de N.; Peri, P.L.; Keulen, van H.

    2008-01-01

    Fruit weight is the main quality parameter of sweet cherries and leaf area/fruit is the most important characteristic influencing fruit weight. The objective of this study was to determine the relationship between Mean Fruit Weight (MFW) and the Fruit Number to Leaf Area Ratio (FNLAR) for `Bing¿,

  5. Use of middle infrared radiation to estimate the leaf area index of a boreal forest

    Energy Technology Data Exchange (ETDEWEB)

    Boyd, D.S. [Kingston Univ., Surrey (United Kingdom). Centre for Earth and Environmental Science Research, School of Geography; Wicks, T. E.; Curran, P.J. [Southampton Univ., Southampton, Hampshire (United Kingdom). Dept. of Geography

    2000-06-01

    Reflected radiation recorded by satellite sensors is a common procedure to estimate the leaf area index (LAI) of boreal forest. The normalized difference vegetation index (NDVI), derived from measurements of visible and near infrared radiation were commonly used to estimate LAI. But research in tropical forest has shown that LAI is more closely related to radiation of middle infrared wavelengths than that of visible wavelengths. This research calculated a vegetation index (VI3) using radiation from vegetation recorded at near and middle infrared wavelengths. In the case of boreal forest, VI3 and LAI displayed a closer relationship than NDVI and LAI. Also, the use of VI3 explained approximately 76 per cent of the variation in field estimates of LAI, versus approximately 46 per cent for NDVI. The authors concluded that consideration should be given to information provided by middle infrared radiation to estimate the leaf area index of boreal forest. The research area was located in the Southern Study Area (SSA) of the BOReal Ecosystem-Atmospher Study (BOREAS), situated on the southern edge of the Canadian boreal forest, 40 km north of Prince Albert, Saskatchewan. 1 tab., 4 figs., 46 refs.

  6. Leaf Physiological and Morphological Responses to Shade in Grass-Stage Seedlings and Young Trees of Longleaf Pine

    Directory of Open Access Journals (Sweden)

    Lisa J. Samuelson

    2012-08-01

    Full Text Available Longleaf pine has been classified as very shade intolerant but leaf physiological plasticity to light is not well understood, especially given longleaf pine’s persistent seedling grass stage. We examined leaf morphological and physiological responses to light in one-year-old grass-stage seedlings and young trees ranging in height from 4.6 m to 6.3 m to test the hypothesis that young longleaf pine would demonstrate leaf phenotypic plasticity to light environment. Seedlings were grown in a greenhouse under ambient levels of photosynthetically active radiation (PAR or a 50% reduction in ambient PAR and whole branches of trees were shaded to provide a 50% reduction in ambient PAR. In seedlings, shading reduced leaf mass per unit area (LMA, the light compensation point, and leaf dark respiration (RD, and increased the ratio of light-saturated photosynthesis to RD and chlorophyll b and total chlorophyll expressed per unit leaf dry weight. In trees, shading reduced LMA, increased chlorophyll a, chlorophyll b and total chlorophyll on a leaf dry weight basis, and increased allocation of total foliar nitrogen to chlorophyll nitrogen. Changes in leaf morphological and physiological traits indicate a degree of shade tolerance that may have implications for even and uneven-aged management of longleaf pine.

  7. Effective leaf area index retrieving from terrestrial point cloud data: coupling computational geometry application and Gaussian mixture model clustering

    Science.gov (United States)

    Jin, S.; Tamura, M.; Susaki, J.

    2014-09-01

    Leaf area index (LAI) is one of the most important structural parameters of forestry studies which manifests the ability of the green vegetation interacted with the solar illumination. Classic understanding about LAI is to consider the green canopy as integration of horizontal leaf layers. Since multi-angle remote sensing technique developed, LAI obliged to be deliberated according to the observation geometry. Effective LAI could formulate the leaf-light interaction virtually and precisely. To retrieve the LAI/effective LAI from remotely sensed data therefore becomes a challenge during the past decades. Laser scanning technique can provide accurate surface echoed coordinates with densely scanned intervals. To utilize the density based statistical algorithm for analyzing the voluminous amount of the 3-D points data is one of the subjects of the laser scanning applications. Computational geometry also provides some mature applications for point cloud data (PCD) processing and analysing. In this paper, authors investigated the feasibility of a new application for retrieving the effective LAI of an isolated broad leaf tree. Simplified curvature was calculated for each point in order to remove those non-photosynthetic tissues. Then PCD were discretized into voxel, and clustered by using Gaussian mixture model. Subsequently the area of each cluster was calculated by employing the computational geometry applications. In order to validate our application, we chose an indoor plant to estimate the leaf area, the correlation coefficient between calculation and measurement was 98.28 %. We finally calculated the effective LAI of the tree with 6 × 6 assumed observation directions.

  8. Crescimento do capim-tifton 85 sob doses de nitrogênio e alturas de corte Growth analysis of Tifton 85 bermudagrass under nitrogen fertilization and plant height

    Directory of Open Access Journals (Sweden)

    Odilon Gomes Pereira

    2012-01-01

    subplots. For all variables, the interaction N rate × plant height was not significant. The net assimilatory rate and relative growth rate of Tifton 85 increased with increasing N rates. The leaf area ratio (LAR decreased linearly with plant height at harvesting. In contrast, the LAR increased linearly with N rate. Linearly, the leaf weight ratio decreased with plant height at harvesting, and increased with N rate. The specific leaf area of Tifton 85 bermudagrass was influenced linear and positively only by nitrogen fertilization. The leaf area index (LAI of Tifton 85 bermudagrass increased by about 0.05 unit for each centimeter increase in plant height at harvesting. There was a quadratic response of N in LAI of Tifton 85 bermudagrass. The increase of bermudagrass height at harvesting season decreases the participation of leaf blade in the forage. The nitrogen increases the growth rate of Tifton 85 bermudagrass. To ensure efficient harvesting of forage, the increase of N rate for Tifton 85 Bermudagrass implies a reduction in the harvest interval.

  9. Why do leaf-tying caterpillars abandon their leaf ties?

    Directory of Open Access Journals (Sweden)

    Michelle Sliwinski

    2013-09-01

    Full Text Available Leaf-tying caterpillars act as ecosystem engineers by building shelters between overlapping leaves, which are inhabited by other arthropods. Leaf-tiers have been observed to leave their ties and create new shelters (and thus additional microhabitats, but the ecological factors affecting shelter fidelity are poorly known. For this study, we explored the effects of resource limitation and occupant density on shelter fidelity and assessed the consequences of shelter abandonment. We first quantified the area of leaf material required for a caterpillar to fully develop for two of the most common leaf-tiers that feed on white oak, Quercus alba. On average, Psilocorsis spp. caterpillars consumed 21.65 ± 0.67 cm2 leaf material to complete development. We also measured the area of natural leaf ties found in a Maryland forest, to determine the distribution of resources available to caterpillars in situ. Of 158 natural leaf ties examined, 47% were too small to sustain an average Psilocorsis spp. caterpillar for the entirety of its development. We also manipulated caterpillar densities within experimental ties on potted trees to determine the effects of cohabitants on the likelihood of a caterpillar to leave its tie. We placed 1, 2, or 4 caterpillars in ties of a standard size and monitored the caterpillars twice daily to track their movement. In ties with more than one occupant, caterpillars showed a significantly greater propensity to leave their tie, and left sooner and at a faster rate than those in ties as single occupants. To understand the consequences of leaf tie abandonment, we observed caterpillars searching a tree for a site to build a shelter in the field. This is a risky behavior, as 17% of the caterpillars observed died while searching for a shelter site. Caterpillars that successfully built a shelter traveled 110 ± 20 cm and took 28 ± 7 min to find a suitable site to build a shelter. In conclusion, leaf-tying caterpillars must frequently

  10. Height-related trends in leaf xylem anatomy and shoot hydraulic characteristics in a tall conifer: safety versus efficiency in water transport

    Science.gov (United States)

    D.R. Woodruff; F.C. Meinzer; B. Lachenbruch

    2008-01-01

    Growth and aboveground biomass accumulation follow a common pattern as tree size increases, with productivity peaking when leaf area reaches its maximum and then declining as tree age and size increase. Age- and size-related declines in forest productivity are major considerations in setting the rotational age of commercial forests, and relate to issues of carbon...

  11. Family differences in equations for predicting biomass and leaf area in Douglas-fir (Pseudotsuga menziesii var. menziesii).

    Science.gov (United States)

    J.B. St. Clair

    1993-01-01

    Logarithmic regression equations were developed to predict component biomass and leaf area for an 18-yr-old genetic test of Douglas-fir (Pseudotsuga menziesii [Mirb.] Franco var. menziesii) based on stem diameter or cross-sectional sapwood area. Equations did not differ among open-pollinated families in slope, but intercepts...

  12. Amazon forest carbon dynamics predicted by profiles of canopy leaf area and light environment

    Science.gov (United States)

    S. C. Stark; V. Leitold; J. L. Wu; M. O. Hunter; C. V. de Castilho; F. R. C. Costa; S. M. McMahon; G. G. Parker; M. Takako Shimabukuro; M. A. Lefsky; M. Keller; L. F. Alves; J. Schietti; Y. E. Shimabukuro; D. O. Brandao; T. K. Woodcock; N. Higuchi; P. B de Camargo; R. C. de Oliveira; S. R. Saleska

    2012-01-01

    Tropical forest structural variation across heterogeneous landscapes may control above-ground carbon dynamics. We tested the hypothesis that canopy structure (leaf area and light availability) – remotely estimated from LiDAR – control variation in above-ground coarse wood production (biomass growth). Using a statistical model, these factors predicted biomass growth...

  13. Comparison of different ground techniques to map leaf area index of Norway spruce forest canopy

    NARCIS (Netherlands)

    Homolova, L.; Malenovsky, Z.; Hanus, J.; Tomaskova, I.; Dvoráková, M.; Pokorny, R.

    2007-01-01

    The leaf area index (LAI) of three monocultures of Norway spruce (Picea abies (L.) Karst), different in age and structure, was measured by means of two indirect optical techniques of LAI field mapping: 1/ plant canopy analyser LAI-2000, and 2/ digital hemispherical photographs (DHP). The supportive

  14. An antithetic variate to facilitate upper-stem height measurements for critical height sampling with importance sampling

    Science.gov (United States)

    Thomas B. Lynch; Jeffrey H. Gove

    2013-01-01

    Critical height sampling (CHS) estimates cubic volume per unit area by multiplying the sum of critical heights measured on trees tallied in a horizontal point sample (HPS) by the HPS basal area factor. One of the barriers to practical application of CHS is the fact that trees near the field location of the point-sampling sample point have critical heights that occur...

  15. Non-destructive measurement of soybean leaf thickness via X-ray computed tomography allows the study of diel leaf growth rhythms in the third dimension.

    Science.gov (United States)

    Pfeifer, Johannes; Mielewczik, Michael; Friedli, Michael; Kirchgessner, Norbert; Walter, Achim

    2018-01-01

    Present-day high-resolution leaf growth measurements provide exciting insights into diel (24-h) leaf growth rhythms and their control by the circadian clock, which match photosynthesis with oscillating environmental conditions. However, these methods are based on measurements of leaf area or elongation and neglect diel changes of leaf thickness. In contrast, the influence of various environmental stress factors to which leaves are exposed to during growth on the final leaf thickness has been studied extensively. Yet, these studies cannot elucidate how variation in leaf area and thickness are simultaneously regulated and influenced on smaller time scales. Only few methods are available to measure the thickness of young, growing leaves non-destructively. Therefore, we evaluated X-ray computed tomography to simultaneously and non-invasively record diel changes and growth of leaf thickness and area. Using conventional imaging and X-ray computed tomography leaf area, thickness and volume growth of young soybean leaves were simultaneously and non-destructively monitored at three cardinal time points during night and day for a period of 80 h under non-stressful growth conditions. Reference thickness measurements on paperboards were in good agreement to CT measurements. Comparison of CT with leaf mass data further proved the consistency of our method. Exploratory analysis showed that measurements were accurate enough for recording and analyzing relative diel changes of leaf thickness, which were considerably different to those of leaf area. Relative growth rates of leaf area were consistently positive and highest during 'nights', while diel changes in thickness fluctuated more and were temporarily negative, particularly during 'evenings'. The method is suitable for non-invasive, accurate monitoring of diel variation in leaf volume. Moreover, our results indicate that diel rhythms of leaf area and thickness show some similarity but are not tightly coupled. These

  16. Stomatal conductance, canopy temperature, and leaf area index estimation using remote sensing and OBIA techniques

    Science.gov (United States)

    S. Panda; D.M. Amatya; G. Hoogenboom

    2014-01-01

    Remotely sensed images including LANDSAT, SPOT, NAIP orthoimagery, and LiDAR and relevant processing tools can be used to predict plant stomatal conductance (gs), leaf area index (LAI), and canopy temperature, vegetation density, albedo, and soil moisture using vegetation indices like normalized difference vegetation index (NDVI) or soil adjusted...

  17. Mapping Vineyard Leaf Area Using Mobile Terrestrial Laser Scanners: Should Rows be Scanned On-the-Go or Discontinuously Sampled?

    Directory of Open Access Journals (Sweden)

    Ignacio del-Moral-Martínez

    2016-01-01

    Full Text Available The leaf area index (LAI is defined as the one-side leaf area per unit ground area, and is probably the most widely used index to characterize grapevine vigor. However, LAI varies spatially within vineyard plots. Mapping and quantifying this variability is very important for improving management decisions and agricultural practices. In this study, a mobile terrestrial laser scanner (MTLS was used to map the LAI of a vineyard, and then to examine how different scanning methods (on-the-go or discontinuous systematic sampling may affect the reliability of the resulting raster maps. The use of the MTLS allows calculating the enveloping vegetative area of the canopy, which is the sum of the leaf wall areas for both sides of the row (excluding gaps and the projected upper area. Obtaining the enveloping areas requires scanning from both sides one meter length section along the row at each systematic sampling point. By converting the enveloping areas into LAI values, a raster map of the latter can be obtained by spatial interpolation (kriging. However, the user can opt for scanning on-the-go in a continuous way and compute 1-m LAI values along the rows, or instead, perform the scanning at discontinuous systematic sampling within the plot. An analysis of correlation between maps indicated that MTLS can be used discontinuously in specific sampling sections separated by up to 15 m along the rows. This capability significantly reduces the amount of data to be acquired at field level, the data storage capacity and the processing power of computers.

  18. Detection of Chlorophyll and Leaf Area Index Dynamics from Sub-weekly Hyperspectral Imagery

    Science.gov (United States)

    Houborg, Rasmus; McCabe, Matthew F.; Angel, Yoseline; Middleton, Elizabeth M.

    2016-01-01

    Temporally rich hyperspectral time-series can provide unique time critical information on within-field variations in vegetation health and distribution needed by farmers to effectively optimize crop production. In this study, a dense time series of images were acquired from the Earth Observing-1 (EO-1) Hyperion sensor over an intensive farming area in the center of Saudi Arabia. After correction for atmospheric effects, optimal links between carefully selected explanatory hyperspectral vegetation indices and target vegetation characteristics were established using a machine learning approach. A dataset of in-situ measured leaf chlorophyll (Chll) and leaf area index (LAI), collected during five intensive field campaigns over a variety of crop types, were used to train the rule-based predictive models. The ability of the narrow-band hyperspectral reflectance information to robustly assess and discriminate dynamics in foliar biochemistry and biomass through empirical relationships were investigated. This also involved evaluations of the generalization and reproducibility of the predictions beyond the conditions of the training dataset. The very high temporal resolution of the satellite retrievals constituted a specifically intriguing feature that facilitated detection of total canopy Chl and LAI dynamics down to sub-weekly intervals. The study advocates the benefits associated with the availability of optimum spectral and temporal resolution spaceborne observations for agricultural management purposes.

  19. Detection of chlorophyll and leaf area index dynamics from sub-weekly hyperspectral imagery

    KAUST Repository

    Houborg, Rasmus; McCabe, Matthew; Angel, Yoseline; Middleton, Elizabeth M.

    2016-01-01

    Temporally rich hyperspectral time-series can provide unique time critical information on within-field variations in vegetation health and distribution needed by farmers to effectively optimize crop production. In this study, a dense timeseries of images were acquired from the Earth Observing-1 (EO-1) Hyperion sensor over an intensive farming area in the center of Saudi Arabia. After correction for atmospheric effects, optimal links between carefully selected explanatory hyperspectral vegetation indices and target vegetation characteristics were established using a machine learning approach. A dataset of in-situ measured leaf chlorophyll (Chll) and leaf area index (LAI), collected during five intensive field campaigns over a variety of crop types, were used to train the rule-based predictive models. The ability of the narrow-band hyperspectral reflectance information to robustly assess and discriminate dynamics in foliar biochemistry and biomass through empirical relationships were investigated. This also involved evaluations of the generalization and reproducibility of the predictions beyond the conditions of the training dataset. The very high temporal resolution of the satellite retrievals constituted a specifically intriguing feature that facilitated detection of total canopy Chl and LAI dynamics down to sub-weekly intervals. The study advocates the benefits associated with the availability of optimum spectral and temporal resolution spaceborne observations for agricultural management purposes.

  20. Detection of chlorophyll and leaf area index dynamics from sub-weekly hyperspectral imagery

    KAUST Repository

    Houborg, Rasmus

    2016-10-25

    Temporally rich hyperspectral time-series can provide unique time critical information on within-field variations in vegetation health and distribution needed by farmers to effectively optimize crop production. In this study, a dense timeseries of images were acquired from the Earth Observing-1 (EO-1) Hyperion sensor over an intensive farming area in the center of Saudi Arabia. After correction for atmospheric effects, optimal links between carefully selected explanatory hyperspectral vegetation indices and target vegetation characteristics were established using a machine learning approach. A dataset of in-situ measured leaf chlorophyll (Chll) and leaf area index (LAI), collected during five intensive field campaigns over a variety of crop types, were used to train the rule-based predictive models. The ability of the narrow-band hyperspectral reflectance information to robustly assess and discriminate dynamics in foliar biochemistry and biomass through empirical relationships were investigated. This also involved evaluations of the generalization and reproducibility of the predictions beyond the conditions of the training dataset. The very high temporal resolution of the satellite retrievals constituted a specifically intriguing feature that facilitated detection of total canopy Chl and LAI dynamics down to sub-weekly intervals. The study advocates the benefits associated with the availability of optimum spectral and temporal resolution spaceborne observations for agricultural management purposes.

  1. Ozone Flux Measurement and Modelling on Leaf/Shoot and Canopy Scale

    Directory of Open Access Journals (Sweden)

    Ludger Grünhage

    calculated by SVAT models often based on the energy balance of the soil-vegetation-atmosphere system and on the big-leaf concept. This latter assumes the canopy as equivalent to a single leaf having a leaf area equal to the total area of all the plant’s leaves and lying at a certain height above the ground. The complexity of SVAT models ranges from one-dimensional to three-dimensional models. The most used are one-dimensional models in single-layer, dual-source or multi-layer version. The main uncertainties in flux modelling are currently associated to the estimation of the non-stomatal flux component and to the up-scaling process from leaf to canopy and stand level. For the latter a separate representation of sunlit and shaded leaves is recommended.

  2. Retrieving Leaf Area Index (LAI) Using Remote Sensing: Theories, Methods and Sensors.

    Science.gov (United States)

    Zheng, Guang; Moskal, L Monika

    2009-01-01

    The ability to accurately and rapidly acquire leaf area index (LAI) is an indispensable component of process-based ecological research facilitating the understanding of gas-vegetation exchange phenomenon at an array of spatial scales from the leaf to the landscape. However, LAI is difficult to directly acquire for large spatial extents due to its time consuming and work intensive nature. Such efforts have been significantly improved by the emergence of optical and active remote sensing techniques. This paper reviews the definitions and theories of LAI measurement with respect to direct and indirect methods. Then, the methodologies for LAI retrieval with regard to the characteristics of a range of remotely sensed datasets are discussed. Remote sensing indirect methods are subdivided into two categories of passive and active remote sensing, which are further categorized as terrestrial, aerial and satellite-born platforms. Due to a wide variety in spatial resolution of remotely sensed data and the requirements of ecological modeling, the scaling issue of LAI is discussed and special consideration is given to extrapolation of measurement to landscape and regional levels.

  3. How do leaf veins influence the worldwide leaf economic spectrum? Review and synthesis.

    Science.gov (United States)

    Sack, Lawren; Scoffoni, Christine; John, Grace P; Poorter, Hendrik; Mason, Chase M; Mendez-Alonzo, Rodrigo; Donovan, Lisa A

    2013-10-01

    Leaf vein traits are implicated in the determination of gas exchange rates and plant performance. These traits are increasingly considered as causal factors affecting the 'leaf economic spectrum' (LES), which includes the light-saturated rate of photosynthesis, dark respiration, foliar nitrogen concentration, leaf dry mass per area (LMA) and leaf longevity. This article reviews the support for two contrasting hypotheses regarding a key vein trait, vein length per unit leaf area (VLA). Recently, Blonder et al. (2011, 2013) proposed that vein traits, including VLA, can be described as the 'origin' of the LES by structurally determining LMA and leaf thickness, and thereby vein traits would predict LES traits according to specific equations. Careful re-examination of leaf anatomy, published datasets, and a newly compiled global database for diverse species did not support the 'vein origin' hypothesis, and moreover showed that the apparent power of those equations to predict LES traits arose from circularity. This review provides a 'flux trait network' hypothesis for the effects of vein traits on the LES and on plant performance, based on a synthesis of the previous literature. According to this hypothesis, VLA, while virtually independent of LMA, strongly influences hydraulic conductance, and thus stomatal conductance and photosynthetic rate. We also review (i) the specific physiological roles of VLA; (ii) the role of leaf major veins in influencing LES traits; and (iii) the role of VLA in determining photosynthetic rate per leaf dry mass and plant relative growth rate. A clear understanding of leaf vein traits provides a new perspective on plant function independently of the LES and can enhance the ability to explain and predict whole plant performance under dynamic conditions, with applications towards breeding improved crop varieties.

  4. Microscopic diagnosis of the leaf and stem of Piper solmsianum C.DC.

    Science.gov (United States)

    Bertocco, A R P; Migacz, I P; Santos, V L P; Franco, C R C; Silva, R Z; Yunes, R A; Cechinel-Filho, V; Budel, J M

    2017-08-01

    Piper solmsianum C.DC., which is popularly known as pariparoba, is a shrub that measures 1-3 m in height and it inhabits areas with wet tropical soils. The objective of this study was to analyze the leaf and stem anatomy using light microscopy, scanning electron micrographs, and energy-dispersive X-ray spectroscopy in order to provide information for species identification. The anatomical profile showed the following main microscopic markers: hypostomatic leaf; hypodermis layer on both sides; pearl glands; biconvex midrib shape; five collateral vascular bundles in open arc with the central bundle larger than the others; circular stem shape; collateral vascular bundles arranged in two rings; sinuous sclerenchymatic sheath in the pith; secretory idioblasts; and starch grains in the mesophyll, in the ground parenchyma of the midrib, petiole, and in the stem; and six morphotypes of calcium oxalate crystals (styloids, cuneiform, tabular crystal rosettes, cuneiform crystal rosettes, elongated square dipyramids, as well as very elongated square dipyramids). © 2017 Wiley Periodicals, Inc.

  5. Litterfall and Leaf Area Index in the CONECOFOR Permanent Monitoring Plots

    Directory of Open Access Journals (Sweden)

    Andrea CUTINI

    2002-09-01

    Full Text Available Forest canopies are more sensitive and react more promptly to abiotic and biotic disturbances than other stand structural components. Monitoring crown and canopy characteristics is therefore a crucial issue for intensive and continuous monitoring programs of forest ecosystem status. These observations formed the basis for the measurement of annual litter production and leaf area index (LAI in the Italian permanent monitoring plots (CONECOFOR program established within the EC-UN/ECE program "Intensive Monitoring (Level II of Forest Ecosystems". Preliminary results after three years of observation are presented. The low value of within plot mean relative standard deviation (20.8 ± 1.9% of litter production, which in any case never exceeded 30%, accounted for the good sampling error and accuracy of the chosen method, which seems to be accurate enough to detect changes in litter production through the years. The higher inconsistency of the amount of woody and fruits fractions over the years demonstrated the greater reliability of leaf fraction or, on the other hand, of LAI compared to total litter. Mean values of annual leaf-litter and total litter production and LAI were rather high in comparison with data reported in literature for similar stands, and reflected both a medium-high productivity and a juvenile phase in the development of the selected stands on average. Focusing on changes in litter production through the years, statistical analysis on a sub-sample of plots showed the existence of significant differences both in leaf litter and total litter production. These findings seem to attribute to the "year" factor a driving role in determining changes in litter production and LAI. Temporal intermittence in data collection, together with the shortness of the monitoring period, make it difficult to speculate or arrive at definitive conclusions on changes in litter production due to time-dependent factors. The importance of having a complete

  6. Estimation of leaf area index in cereal crops using red–green images

    DEFF Research Database (Denmark)

    Nielsen, Kristian Kirk; Andersen, Hans Jørgen; Thomsen, Anton

    2009-01-01

    A new method for estimating the leaf area index (LAI) in cereal crops based on red–green images taken from above the crop canopy is introduced. The proposed method labels pixels into vegetation and soil classes using a combination of greenness and intensity derived from the red and green colour b....... Conclusions Acknowledgements Appendix. Modelling the correlation between greenness and brightness References   Fig. 1. Simulated image of a vegetation canopy (left), with distribution of pixel greenness and brightness (right). View Within Article...

  7. Defoliation patterns in signal grass tillers with varying heights in the same pasture - doi: 10.4025/actascianimsci.v35i3.14376

    Directory of Open Access Journals (Sweden)

    Simone Pedro da Silva

    2013-07-01

    Full Text Available The experiment was carried out to evaluate the patterns of defoliation in plants of various heights in the same pasture of Brachiaria decumbens cv. Basilisk under continuous stocking with cattle. Four plant heights were evaluated (10, 20, 30 and 40 cm in the same managed sward, with mean height equal to 25 cm. A randomized blocks design was used, with two replications. Defoliation frequency increased linearly along with plant height in the same pasture. Defoliation intensity, number of defoliations in leaf blade and percentage of leaf blade grazed per tiller also increased linearly along with plant height. The defoliation interval decreased linearly according to plant height. The quadratic model was adequately fitted into grazing efficiency data, reaching the highest value (98% at the sward site with 35 cm plants. The percentage variation in signal grass height showed a negative linear response with initial plant height. The structure of the pasture, characterized by the diversity in plant heights, is simultaneously the cause and consequence of the variability in defoliation patterns in individual tillers, which occur on the horizontal plane of the pasture.  

  8. Improved estimation of leaf area index and leaf chlorophyll content of a potato crop using multi-angle spectral data – potential of unmanned aerial vehicle imagery

    NARCIS (Netherlands)

    Roosjen, Peter P.J.; Brede, Benjamin; Suomalainen, Juha M.; Bartholomeus, Harm M.; Kooistra, Lammert; Clevers, Jan G.P.W.

    2018-01-01

    In addition to single-angle reflectance data, multi-angular observations can be used as an additional information source for the retrieval of properties of an observed target surface. In this paper, we studied the potential of multi-angular reflectance data for the improvement of leaf area index

  9. Multi-Spectral Imaging from an Unmanned Aerial Vehicle Enables the Assessment of Seasonal Leaf Area Dynamics of Sorghum Breeding Lines

    Directory of Open Access Journals (Sweden)

    Andries B. Potgieter

    2017-09-01

    Full Text Available Genetic improvement in sorghum breeding programs requires the assessment of adaptation traits in small-plot breeding trials across multiple environments. Many of these phenotypic assessments are made by manual measurement or visual scoring, both of which are time consuming and expensive. This limits trial size and the potential for genetic gain. In addition, these methods are typically restricted to point estimates of particular traits, such as leaf senescence or flowering and do not capture the dynamic nature of crop growth. In water-limited environments in particular, information on leaf area development over time would provide valuable insight into water use and adaptation to water scarcity during specific phenological stages of crop development. Current methods to estimate plant leaf area index (LAI involve destructive sampling and are not practical in breeding. Unmanned aerial vehicles (UAV and proximal-sensing technologies open new opportunities to assess these traits multiple times in large small-plot trials. We analyzed vegetation-specific crop indices obtained from a narrowband multi-spectral camera on board a UAV platform flown over a small pilot trial with 30 plots (10 genotypes randomized within 3 blocks. Due to variable emergence we were able to assess the utility of these vegetation indices to estimate canopy cover and LAI over a large range of plant densities. We found good correlations between the Normalized Difference Vegetation Index (NDVI and the Enhanced Vegetation Index (EVI with plant number per plot, canopy cover and LAI both during the vegetative growth phase (pre-anthesis and at maximum canopy cover shortly after anthesis. We also analyzed the utility of time-sequence data to assess the senescence pattern of sorghum genotypes known as fast (senescent or slow senescing (stay-green types. The Normalized Difference Red Edge (NDRE index which estimates leaf chlorophyll content was most useful in characterizing the leaf area

  10. Can Leaf Spectroscopy Predict Leaf and Forest Traits Along a Peruvian Tropical Forest Elevation Gradient?

    Science.gov (United States)

    Doughty, Christopher E.; Santos-Andrade, P. E.; Goldsmith, G. R.; Blonder, B.; Shenkin, A.; Bentley, L. P.; Chavana-Bryant, C.; Huaraca-Huasco, W.; Díaz, S.; Salinas, N.; Enquist, B. J.; Martin, R.; Asner, G. P.; Malhi, Y.

    2017-11-01

    High-resolution spectroscopy can be used to measure leaf chemical and structural traits. Such leaf traits are often highly correlated to other traits, such as photosynthesis, through the leaf economics spectrum. We measured VNIR (visible-near infrared) leaf reflectance (400-1,075 nm) of sunlit and shaded leaves in 150 dominant species across ten, 1 ha plots along a 3,300 m elevation gradient in Peru (on 4,284 individual leaves). We used partial least squares (PLS) regression to compare leaf reflectance to chemical traits, such as nitrogen and phosphorus, structural traits, including leaf mass per area (LMA), branch wood density and leaf venation, and "higher-level" traits such as leaf photosynthetic capacity, leaf water repellency, and woody growth rates. Empirical models using leaf reflectance predicted leaf N and LMA (r2 > 30% and %RMSE < 30%), weakly predicted leaf venation, photosynthesis, and branch density (r2 between 10 and 35% and %RMSE between 10% and 65%), and did not predict leaf water repellency or woody growth rates (r2<5%). Prediction of higher-level traits such as photosynthesis and branch density is likely due to these traits correlations with LMA, a trait readily predicted with leaf spectroscopy.

  11. Bayesian estimation of seasonal course of canopy leaf area index from hyperspectral satellite data

    Science.gov (United States)

    Varvia, Petri; Rautiainen, Miina; Seppänen, Aku

    2018-03-01

    In this paper, Bayesian inversion of a physically-based forest reflectance model is investigated to estimate of boreal forest canopy leaf area index (LAI) from EO-1 Hyperion hyperspectral data. The data consist of multiple forest stands with different species compositions and structures, imaged in three phases of the growing season. The Bayesian estimates of canopy LAI are compared to reference estimates based on a spectral vegetation index. The forest reflectance model contains also other unknown variables in addition to LAI, for example leaf single scattering albedo and understory reflectance. In the Bayesian approach, these variables are estimated simultaneously with LAI. The feasibility and seasonal variation of these estimates is also examined. Credible intervals for the estimates are also calculated and evaluated. The results show that the Bayesian inversion approach is significantly better than using a comparable spectral vegetation index regression.

  12. [Latitude variation mechanism of leaf traits of Metasequoia glyptostroboides in eastern coastal China].

    Science.gov (United States)

    Guo, Wei Hong; Wang, Hua; Yu, Mu Kui; Wu, Tong Gui; Han, You Zhi

    2017-03-18

    We analyzed the rules of Metasequoia glyptostroboides along with latitude, including leaf length, leaf width, leaf perimeter, leaf area, ratio of leaf length to width, specific leaf area (SLA), and leaf dry mass based on eight stands growing at different latitudes in the coastal area of eastern China, as well as their relationships with climatic and soil factors. The results showed that the leaf length, leaf width and leaf perimeter increased with increasing latitude, while the leaf area and SLA firstly increased and then decreased. The mean annual temperature and annual precipitation were the major environmental factors affecting the leaf traits along latitude gradient. With the increase of soil N content, the SLA decreased firstly and then increased, while the leaf mass decreased significantly. With the increase of soil P content, the SLA increased, and the leaf mass decreased significantly.

  13. Measurement methods and variability assessment of the Norway spruce total leaf area: Implications for remote sensing

    NARCIS (Netherlands)

    Homolova, L.; Lukes, P.; Malenovsky, Z.; Lhotakova, Z.; Kaplan, V.; Hanus, J.

    2013-01-01

    Estimation of total leaf area (LAT) is important to express biochemical properties in plant ecology and remote sensing studies. A measurement of LAT is easy in broadleaf species, but it remains challenging in coniferous canopies. We proposed a new geometrical model to estimate Norway spruce LAT and

  14. Effects of different potting growing media for Petunia grandiflora and Nicotiana alata Link & Otto on photosynthetic capacity, leaf area, and flowering potential

    Directory of Open Access Journals (Sweden)

    Gheorghe Cristian Popescu

    2015-03-01

    Full Text Available Petunia grandiflora Juss. and Nicotiana alata Link & Otto are two of the most widely spread plants on the market for annual potted ornamental plants. In order to identify the most adequate substrate formula we analyzed the effects of different potting growing media used for P. hybrida grandiflora 'Bravo' and N. alata 'Dinamo' on their photosynthetic capacity, leaf area, and flowering potential. Optimization of growing media formula for petunia and ornamental tobacco was performed by preparing four growing media mixing fallow soil (FS, Biolan peat (BP, acid peat (AP, leaf compost (C, and perlite (P in different proportions. The physiological potential of petunia and ornamental tobacco was investigated by photosynthesis and respiration rate and chlorophyll pigments in leaves, while the vegetative and flowering phenological stages were evaluated by number of leaves per plant, leaf area, number of flowers per plant and leaf area/flowers ratio. These measurements were significantly influenced by the different potting growing media used in this study. In the flowering stage, the highest photosynthesis rates (8.612 μmol CO2 m-2 s-1 as well as leaf area (1.766 dm² of petunias were obtained on growing media with 60% biolan peat, 30% acid peat and 10% perlite (BP60-AP30-P10. Flowering responses to growing conditions vary greatly among plants and the biggest number of ornamental tobacco flowers (22 flowers plant-1 was registered as an effect of BP60-AP30-P10 media. Growing media with the BP60-AP30-P10 formula seem to be the most adequate growth substrate to develop profitable crops for petunias and ornamental tobacco with high decorative value.

  15. Localising QTLs for leaf rust resistance and agronomic traits in barley (¤Hordeum vulgare¤ L.)

    DEFF Research Database (Denmark)

    Kicherer, S.; Backes, G.; Walther, U.

    2000-01-01

    to leaf rust by means of artificial infection, heading date, plant height and Kernel weight were assessed. For leaf rust resistance, 4 QTLs were localised, that explained 96.1% of the genetic variation. One QTL on chromosome 4H confirmed a position found in another genetic background and one mapped...

  16. Testing high spatial resolution WorldView-2 imagery for retrieving the leaf area index

    Science.gov (United States)

    Tarantino, Eufemia; Novelli, Antonio; Laterza, Maurizio; Gioia, Andrea

    2015-06-01

    This work analyzes the potentiality of WorldView-2 satellite data for retrieving the Leaf Area Index (LAI) area located in Apulia, the most Eastern region of Italy, overlooking the Adriatic and Ionian seas. Lacking contemporary in-situ measurements, the semi-empiric method of Clevers (1989) (CLAIR model) was chosen as a feasible image-based LAI retrieval method, which is based on an inverse exponential relationship between the LAI and the WDVI (Weighted Difference Vegetation Index) with relation to different land covers. Results were examined in homogeneous land cover classes and compared with values obtained in recent literature.

  17. Monitoring crop leaf area index time variation from higher resolution remotely sensed data

    International Nuclear Information System (INIS)

    Jiao, Sihong

    2014-01-01

    The leaf area index (LAI) is significant for research on global climate change and ecological environment. China HJ-1 satellite has a revisit cycle of four days, providing CCD data (HJ-1 CCD) with a resolution of 30 m. However, the HJ-1 CCD is incapable of obtaining observations at multiple angles. This is problematic because single angle observations provide insufficient data for determining the LAI. This article proposes a new method for determining LAI using HJ-1 CCD data. The proposed method uses background knowledge of dynamic land surface processes that are extracted from MODerate resolution Imaging Spectroradiometer (MODIS) LAI 1-km resolution data. To process the uncertainties that arise from using two data sources with different spatial resolutions, the proposed method is implemented in a dynamitic Bayesian network scheme by integrating a LAI dynamic process model and a canopy reflectance model with remotely sensed data. Validation results showed that the determination coefficient between estimated and measured LAI was 0.791, and the RMSE was 0.61. This method can enhance the accuracy of the retrieval results while retaining the time series variation characteristics of the vegetation LAI. The results suggest that this algorithm can be widely applied to determining high-resolution leaf area indices using data from China HJ-1 satellite even if information from single angle observations are insufficient for quantitative application

  18. Large-scale Estimates of Leaf Area Index from Active Remote Sensing Laser Altimetry

    Science.gov (United States)

    Hopkinson, C.; Mahoney, C.

    2016-12-01

    Leaf area index (LAI) is a key parameter that describes the spatial distribution of foliage within forest canopies which in turn control numerous relationships between the ground, canopy, and atmosphere. The retrieval of LAI has demonstrated success by in-situ (digital) hemispherical photography (DHP) and airborne laser scanning (ALS) data; however, field and ALS acquisitions are often spatially limited (100's km2) and costly. Large-scale (>1000's km2) retrievals have been demonstrated by optical sensors, however, accuracies remain uncertain due to the sensor's inability to penetrate the canopy. The spaceborne Geoscience Laser Altimeter System (GLAS) provides a possible solution in retrieving large-scale derivations whilst simultaneously penetrating the canopy. LAI retrieved by multiple DHP from 6 Australian sites, representing a cross-section of Australian ecosystems, were employed to model ALS LAI, which in turn were used to infer LAI from GLAS data at 5 other sites. An optimally filtered GLAS dataset was then employed in conjunction with a host of supplementary data to build a Random Forest (RF) model to infer predictions (and uncertainties) of LAI at a 250 m resolution across the forested regions of Australia. Predictions were validated against ALS-based LAI from 20 sites (R2=0.64, RMSE=1.1 m2m-2); MODIS-based LAI were also assessed against these sites (R2=0.30, RMSE=1.78 m2m-2) to demonstrate the strength of GLAS-based predictions. The large-scale nature of current predictions was also leveraged to demonstrate large-scale relationships of LAI with other environmental characteristics, such as: canopy height, elevation, and slope. The need for such wide-scale quantification of LAI is key in the assessment and modification of forest management strategies across Australia. Such work also assists Australia's Terrestrial Ecosystem Research Network, in fulfilling their government issued mandates.

  19. Growth, leaf traits and litter decomposition of roadside hybrid aspen (Populus tremula L. x P. tremuloides Michx.) clones

    International Nuclear Information System (INIS)

    Nikula, Suvi; Manninen, Sirkku; Vapaavuori, Elina; Pulkkinen, Pertti

    2011-01-01

    Road traffic contributes considerably to ground-level air pollution and is therefore likely to affect roadside ecosystems. Differences in growth and leaf traits among 13 hybrid aspen (Populus tremula x P. tremuloides) clones were studied in relation to distance from a motorway. The trees sampled were growing 15 and 30 m from a motorway and at a background rural site in southern Finland. Litter decomposition was also measured at both the roadside and rural sites. Height and diameter growth rate and specific leaf area were lowest, and epicuticular wax amount highest in trees growing 15 m from the motorway. Although no significant distance x clone interactions were detected, clone-based analyses indicated differences in genotypic responses to motorway proximity. Leaf N concentration did not differ with distance from the motorway for any of the clones. Leaf litter decomposition was only temporarily retarded in the roadside environment, suggesting minor effects on nutrient cycling. - Highlights: → Roadside hybrid aspen displayed xeromorphic leaf traits and reduction in growth rate. → These responses were limited to trees close to the motorway and only to some clones. → Leaf litter decomposition was only temporarily retarded in the roadside environment. - Hybrid aspen had more xeromorphic leaves, displayed reduced growth, and showed retarded litter decomposition at an early stage in the roadside environment.

  20. Correlations between some physiographic factors and diameter, height and cone characteristics in stone pine (Pinus pinea L. afforestation areas in Balıkesir-Burhaniye region

    Directory of Open Access Journals (Sweden)

    Ali Ömer Üçler

    2017-11-01

    Full Text Available In this study, effects of some physiographic factors (aspect, slope and elevation on breast diameter, height and cone properties (cone weight, number of cones, cone width and cone length in stone pine (Pinus pinea L. afforestation areas in Balıkesir-Burhaniye region were investigated. To determine to effects of physiographic conditions, study area is separated according to aspect, slope and elevation groups. Aspect is separated two groups which are shady aspect (B1 and sunny aspect (B2. Slope is separated two groups which are small from 36 percent (E1 and big from 36 percent (E2. Elevation also is separated two groups which are under 400 meters (Y1 and over 400 meters (Y2. Sample areas were scattering to land classifications by using Stratified Sampling Method. Thus, each aspect, slope and elevation classifications were obtained from sample area where are provided from 32 different areas. Tree heights, the diameter at breast height and cone number were measured on saplings. In addition, in each sample area five Pinus pinea trees were selected. It was measured cone diameter, cone height and air-dries weight values of cones which are picked from selecting trees. According to statistical analysis results, aspect effects to tree heights, cone number and cone weight; slope effects the diameter at breast height, cone number and cone diameter; elevation effects the diameter at breast height, tree height, cone number and cone weight. In terms of tree height and diameter values; the highest average breast height diameter was found in the B1E1Y1 with 13.6 cm, and the highest average height value was found in the B2E2Y2 by interaction. In terms of cone characteristics; with the highest average cone number of 8.5 in B1E1Y1, the highest mean cone weight in B1E1Y1 with 232 gr and the highest cone diameter in B2E2Y1 with 5.4 cm. According to the results of correlation analysis; it was determined that slope and elevation factors have positive effect on breast

  1. Retrieval of Specific Leaf Area From Landsat-8 Surface Reflectance Data Using Statistical and Physical Models

    NARCIS (Netherlands)

    Ali, Abebe Mohammed; Darvishzadeh, R.; Skidmore, Andrew K.

    2017-01-01

    One of the key traits in the assessment of ecosystem functions is a specific leaf area (SLA). The main aim of this study was to examine the potential of new generation satellite images, such as Landsat-8 imagery, for the retrieval of SLA at regional and global scales. Therefore, both statistical and

  2. Leaf gas exchange, fv/fm ratio, ion content and growth conditions of the two moringa species under magnetic water treatment

    International Nuclear Information System (INIS)

    Hasan, M.M.; Alharby, H.F.; Hajar, A.; Hakeem, K.R.

    2017-01-01

    The current greenhouse experiment investigates the role of magnetic water on the two Moringa species (Moringa oleifera and Moringa peregrina). Both species were exposed to the magnetic field (30 mT). The magnetic water increased the plant height, leaf number, leaflet number, and internode distances in both the species, respectively. Relative water content (RWC) and leaf area in both the species showed changes under magnetic water treatment. The results showed in magnetic water treatment, the leaf gas exchange parameters such as assimilation (A), stomatal conductance (gs), transpiration rate (E), and vapor pressure deficit (VPD) were increased. Similarly, Photosynthetic pigments (Chl a, Chl b, Chl (a+b), Carotenoids), photosynthetic water use efficiency (WUE) were also increased significantly. Magnetized water had also significant effects on the maximal efficiency of PSII photochemistry (Fv/Fm). Our study suggested that magnetic water treatment could be used as an environment-friendly technology for improving the growth and physiology of Moringa species. In addition, this technology could be further incorporated into the traditional methods of agriculture for the improvement of crop plants, particularly in the arid and sub-arid areas of the world. (author)

  3. SOIL EXCHANGEABLE ALUMINUM INFLUENCING THE GROWTH AND LEAF TISSUE MACRONUTRIENTS CONTENT OF CASTOR PLANTS

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    ROSIANE DE LOURDES SILVA DE LIMA

    2014-01-01

    Full Text Available Three castor ( Ricinus communis genotypes were studied regarding tolerance to high exchange factorial distribution of five doses of exchangeable aluminum added to the soil (0, 0.15, 0.30, 0.60, and 1.20 cmol c dm - 3 and three castor genotypes (BRS Nordestina, BRS Paraguaçu, and Lyra. The plants were raised in pots in a greenhouse. At 53 days after emergence, data were taken on plant height, leaf area, dry mass of shoot and root, and leaf tissue content of macronutrients. The most sensitive genotype was the cv. BRS Nordestina, in which the shoot and root dry weight in the highest aluminum content were reduced to 12.9% and 16.2% of the control treatment, respectively. The most tolerant genotype was the hybrid Lyra, in which the shoot and root dry weight in the maximum content of aluminum were reduced to 43.5% and 42.7% of the control treatment, respectively.The increased exchangeable aluminum affected the leaf nutrient content, and the intensity of the response was different among cultivars. The aluminum toxicity increased N, Ca, and Mg contents and reduced on P, K, and S contents. The cv. BRS Nordestina had a drastic shoot dry weight reduction associated with an intense increment in the N leaf content. Thus, the N increment was caused by a concentration effect caused by the limited growth.

  4. Digital Cover Photography for Estimating Leaf Area Index (LAI in Apple Trees Using a Variable Light Extinction Coefficient

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    Carlos Poblete-Echeverría

    2015-01-01

    Full Text Available Leaf area index (LAI is one of the key biophysical variables required for crop modeling. Direct LAI measurements are time consuming and difficult to obtain for experimental and commercial fruit orchards. Devices used to estimate LAI have shown considerable errors when compared to ground-truth or destructive measurements, requiring tedious site-specific calibrations. The objective of this study was to test the performance of a modified digital cover photography method to estimate LAI in apple trees using conventional digital photography and instantaneous measurements of incident radiation (Io and transmitted radiation (I through the canopy. Leaf area of 40 single apple trees were measured destructively to obtain real leaf area index (LAID, which was compared with LAI estimated by the proposed digital photography method (LAIM. Results showed that the LAIM was able to estimate LAID with an error of 25% using a constant light extinction coefficient (k = 0.68. However, when k was estimated using an exponential function based on the fraction of foliage cover (ff derived from images, the error was reduced to 18%. Furthermore, when measurements of light intercepted by the canopy (Ic were used as a proxy value for k, the method presented an error of only 9%. These results have shown that by using a proxy k value, estimated by Ic, helped to increase accuracy of LAI estimates using digital cover images for apple trees with different canopy sizes and under field conditions.

  5. Digital cover photography for estimating leaf area index (LAI) in apple trees using a variable light extinction coefficient.

    Science.gov (United States)

    Poblete-Echeverría, Carlos; Fuentes, Sigfredo; Ortega-Farias, Samuel; Gonzalez-Talice, Jaime; Yuri, Jose Antonio

    2015-01-28

    Leaf area index (LAI) is one of the key biophysical variables required for crop modeling. Direct LAI measurements are time consuming and difficult to obtain for experimental and commercial fruit orchards. Devices used to estimate LAI have shown considerable errors when compared to ground-truth or destructive measurements, requiring tedious site-specific calibrations. The objective of this study was to test the performance of a modified digital cover photography method to estimate LAI in apple trees using conventional digital photography and instantaneous measurements of incident radiation (Io) and transmitted radiation (I) through the canopy. Leaf area of 40 single apple trees were measured destructively to obtain real leaf area index (LAI(D)), which was compared with LAI estimated by the proposed digital photography method (LAI(M)). Results showed that the LAI(M) was able to estimate LAI(D) with an error of 25% using a constant light extinction coefficient (k = 0.68). However, when k was estimated using an exponential function based on the fraction of foliage cover (f(f)) derived from images, the error was reduced to 18%. Furthermore, when measurements of light intercepted by the canopy (Ic) were used as a proxy value for k, the method presented an error of only 9%. These results have shown that by using a proxy k value, estimated by Ic, helped to increase accuracy of LAI estimates using digital cover images for apple trees with different canopy sizes and under field conditions.

  6. Digital Cover Photography for Estimating Leaf Area Index (LAI) in Apple Trees Using a Variable Light Extinction Coefficient

    Science.gov (United States)

    Poblete-Echeverría, Carlos; Fuentes, Sigfredo; Ortega-Farias, Samuel; Gonzalez-Talice, Jaime; Yuri, Jose Antonio

    2015-01-01

    Leaf area index (LAI) is one of the key biophysical variables required for crop modeling. Direct LAI measurements are time consuming and difficult to obtain for experimental and commercial fruit orchards. Devices used to estimate LAI have shown considerable errors when compared to ground-truth or destructive measurements, requiring tedious site-specific calibrations. The objective of this study was to test the performance of a modified digital cover photography method to estimate LAI in apple trees using conventional digital photography and instantaneous measurements of incident radiation (Io) and transmitted radiation (I) through the canopy. Leaf area of 40 single apple trees were measured destructively to obtain real leaf area index (LAID), which was compared with LAI estimated by the proposed digital photography method (LAIM). Results showed that the LAIM was able to estimate LAID with an error of 25% using a constant light extinction coefficient (k = 0.68). However, when k was estimated using an exponential function based on the fraction of foliage cover (ff) derived from images, the error was reduced to 18%. Furthermore, when measurements of light intercepted by the canopy (Ic) were used as a proxy value for k, the method presented an error of only 9%. These results have shown that by using a proxy k value, estimated by Ic, helped to increase accuracy of LAI estimates using digital cover images for apple trees with different canopy sizes and under field conditions. PMID:25635411

  7. Seasonal patterns of leaf gas exchange and water relations in dry rain forest trees of contrasting leaf phenology.

    Science.gov (United States)

    Choat, Brendan; Ball, Marilyn C; Luly, Jon G; Donnelly, Christine F; Holtum, Joseph A M

    2006-05-01

    Diurnal and seasonal patterns of leaf gas exchange and water relations were examined in tree species of contrasting leaf phenology growing in a seasonally dry tropical rain forest in north-eastern Australia. Two drought-deciduous species, Brachychiton australis (Schott and Endl.) A. Terracc. and Cochlospermum gillivraei Benth., and two evergreen species, Alphitonia excelsa (Fenzal) Benth. and Austromyrtus bidwillii (Benth.) Burret. were studied. The deciduous species had higher specific leaf areas and maximum photosynthetic rates per leaf dry mass in the wet season than the evergreens. During the transition from wet season to dry season, total canopy area was reduced by 70-90% in the deciduous species and stomatal conductance (g(s)) and assimilation rate (A) were markedly lower in the remaining leaves. Deciduous species maintained daytime leaf water potentials (Psi(L)) at close to or above wet season values by a combination of stomatal regulation and reduction in leaf area. Thus, the timing of leaf drop in deciduous species was not associated with large negative values of daytime Psi(L) (greater than -1.6 MPa) or predawn Psi(L) (greater than -1.0 MPa). The deciduous species appeared sensitive to small perturbations in soil and leaf water status that signalled the onset of drought. The evergreen species were less sensitive to the onset of drought and g(s) values were not significantly lower during the transitional period. In the dry season, the evergreen species maintained their canopies despite increasing water-stress; however, unlike Eucalyptus species from northern Australian savannas, A and g(s) were significantly lower than wet season values.

  8. Drought adaptation strategies of four grapevine cultivars (Vitis vinifera L.: modification of the properties of the leaf area

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    María Gómez-del-Campo

    2003-09-01

    Full Text Available This essay studies the morphological and anatomical properties of the leaves of Garnacha tinta, Tempranillo, Chardonnay and Airén grapevines in order to discover the drought adaptation strategies present in Vitis vinifera L. The grapevines were grown under two water availability conditions: water limitation and non-water limitation. There was a significantly lower development of leaf area under conditions of water limitation compared to non-water limitation due to a reduction in the size of main and lateral shoot leaves, and a smaller number of leaves on lateral shoots. The development of the leaf area under water limitation conditions occurred on earlier dates than under non-water limitation conditions. Significantly lower stomatal density was observed under water limitation conditions rather than non-water limitation conditions exclusively in the Airén cultivar.

  9. Increase the foliage area of Asparagus Officinalis L. Cv. UC 157 F1 “asparagus” by the spraying of Gibberellin (AG3 and 6 – Benzilaminopurine (6 – BAP

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    José Paraguay Mercado

    2010-12-01

    Full Text Available The present work had as objective to study the increment of the leaf area of Asparagus Officinalis L., for they were used it asparagus crown with 5 yolks, to those that were applied by aspersion, different concentrations of the hormones of gibberellic acid and 6-bencilaminopurina which were applied at the 25, 60, 110 and 160 days respectively. It was found that in the different rehearsed treatments one doesn't observe differential significant in the increment of the number of plants, plant height, number of yolks and dry weight of foliage. However it was observed that the combined application of the phytohormonas in the range of 0.110 at 0.230 m of gibberllic acid and 0.037 to 0.075 m of 6-Bencilaminopurin a bigger number of yolk is achieved (13%, sprout (7%, plant height (11.3% and dry weight (7.3% in the treated plants. Concluded to continue making studies of sinergism of these two phytohormonas, gibberellic (AG3 and 6-bencilaminopurina (6-BAP, seeking to achieve adequate levels of leaf area increase of the leaf area in the asparagus.

  10. Linear relations between leaf mass per area (LMA) and seasonal climate discovered through Linear Manifold Clustering (LMC)

    Science.gov (United States)

    Kiang, N. Y.; Haralick, R. M.; Diky, A.; Kattge, J.; Su, X.

    2016-12-01

    Leaf mass per area (LMA) is a critical variable in plant carbon allocation, correlates with leaf activity traits (photosynthetic activity, respiration), and is a controller of litterfall mass and hence carbon substrate for soil biogeochemistry. Recent advances in understanding the leaf economics spectrum (LES) show that LMA has a strong correlation with leaf life span, a trait that reflects ecological strategy, whereas physiological traits that control leaf activity scale with each other when mass-normalized (Osnas et al., 2013). These functional relations help reduce the number of independent variables in quantifying leaf traits. However, LMA is an independent variable that remains a challenge to specify in dynamic global vegetation models (DGVMs), when vegetation types are classified into a limited number of plant functional types (PFTs) without clear mechanistic drivers for LMA. LMA can range orders of magnitude across plant species, as well as vary within a single plant, both vertically and seasonally. As climate relations in combination with alternative ecological strategies have yet to be well identified for LMA, we have assembled 22,000 records of LMA spanning 0.004 - 33 mg/m2 from the numerous contributors to the TRY database (Kattge et al., 2011), with observations distributed over several climate zones and plant functional categories (growth form, leaf type, phenology). We present linear relations between LMA and climate variables, including seasonal temperature, precipitation, and radiation, as derived through Linear Manifold Clustering (LMC). LMC is a stochastic search technique for identifying linear dependencies between variables in high dimensional space. We identify a set of parsimonious classes of LMA-climate groups based on a metric of minimum description to identify structure in the data set, akin to data compression. The relations in each group are compared to Köppen-Geiger climate classes, with some groups revealing continuous linear relations

  11. Measuring device and method for dimples height differences of 17 x 17 grid

    International Nuclear Information System (INIS)

    Xu Yilan; Zheng Zhihui; Yan Liwei; Wang Xihe

    2001-01-01

    There are 264 cell for fastening fuel rods in the grid of 17 x 17 fuel assembly of PWR. The height differences of top and bottom dimples in a grid is an important quality characteristic of the grid. The report deals with measuring machine and method for dimples height differences of the grid. The device has two measuring probes. The Parallel Leaf Spring is used for transmitting the little displacement between two probes. The uncertainty of the device is σ≤4 μm. The measuring method is shown to be practicable

  12. Pasture height at the beginning of deferment as a determinant of signal grass structure and potential selectivity by cattle - doi: 10.4025/actascianimsci.v35i4.20421

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    Manoel Eduardo Rozalino Santos

    2013-10-01

    Full Text Available Current experiment identified the height of signal grass {Urochloa decumbens (Stapf R. D. Webster cv. Basilisk [syn. Brachiaria decumbens Stapf cv. Basilisk]} at the beginning of deferment that provided an appropriate pasture structure and potential selectivity by cattle on deferred pastures. Four pasture heights at the beginning of deferment (10, 20, 30 and 40 cm and two forage samples (available on pasture and simulated grazing were studied. The experimental design was set in completely randomized blocks, with two replications, in a split-plot arrangement. Higher percentage of live leaf blades and lower percentage of live stems and senescent forage were recorded in the forage sample from simulated grazing. The increase in pasture height increased the percentage of senescent forage and reduced the percentage of live leaf blades in forage samples. Pasture height at the beginning of deferment did not affect the potential selectivity index by cattle for the percentage of live leaf blade. The potential selectivity index varied quadratically for the percentage of live stems and increased linearly for the percentage of senescent forage with pasture height. A 10-to-20 cm reduction in pasture height at the beginning of deferment improved the structure of deferred signal grass and optimized selectivity by cattle.  

  13. Effects of structural complexity on within-canopy light environments and leaf traits in a northern mixed deciduous forest.

    Science.gov (United States)

    Fotis, Alexander T; Curtis, Peter S

    2017-10-01

    Canopy structure influences forest productivity through its effects on the distribution of radiation and the light-induced changes in leaf physiological traits. Due to the difficulty of accessing and measuring forest canopies, few field-based studies have quantitatively linked these divergent scales of canopy functioning. The objective of our study was to investigate how canopy structure affects light profiles within a forest canopy and whether leaves of mature trees adjust morphologically and biochemically to the light environments characteristic of canopies with different structural complexity. We used a combination of light detection and ranging (LiDAR) data and hemispherical photographs to quantify canopy structure and light environments, respectively, and a telescoping pole to sample leaves. Leaf mass per area (LMA), nitrogen on an area basis (Narea) and chlorophyll on a mass basis (Chlmass) were measured in red maple (Acer rubrum), american beech (Fagus grandifolia), white pine (Pinus strobus), and northern red oak (Quercus rubra) at different heights in plots with similar leaf area index but contrasting canopy complexity (rugosity). We found that more complex canopies had greater porosity and reduced light variability in the midcanopy while total light interception was unchanged relative to less complex canopies. Leaf phenotypes of F. grandifolia, Q. rubra and P. strobus were more sun-acclimated in the midstory of structurally complex canopies while leaf phenotypes of A. rubrum were more shade-acclimated (lower LMA) in the upper canopy of more complex stands, despite no differences in total light interception. Broadleaf species showed further differences in acclimation with increased Narea and reduced Chlmass in leaves with higher LMA, while P. strobus showed no change in Narea and Chlmass with higher LMA. Our results provide new insight on how light distribution and leaf acclimation in mature trees might be altered when natural and anthropogenic

  14. Estrutura do pasto de capim-braquiária com variação de alturas Structure of signal grass pasture with height variation

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    Manoel Eduardo Rozalino Santos

    2010-10-01

    experimental design with two repetitions. Variation in the initial height of the signal grass pasture plant showed a linear and a negative response. The masses of green leaf blade mass, live stem, dead material, live forage and total forage linearly increased with heights of the plants on the pasture. However, it was observed a linear reduction in green leaf blade/live stem ratio and the height of the plants on the pasture (from 1.16 to 0.58. Plant height on the pasture had a stronger effect on the increase of live stem and the dead material masses than it had on the green leaf blade mass. Volumetric density of green leaf blade decreased (from 97.09 to 39.07 kg/cm ha DM with plant height on the pasture. The same response was found for volumetric densities of live forage (from 181.62 to 107.85 kg/cm.ha dry matter and total forage (from 283.39 to 195.72 kg/cm.ha dry matter. On the other hand, the plant heights on pasture did not affect volumetric density of live stem and dead material, which showed mean values 80.95 and 95.61 kg/cm.ha DM, respectively. The heights of the plants on pasture had a quadratic effect on light interception by the canopy. Grazing areas with heights of 10, 20, 30 and 40 cm showed light interception values of 56.15, 85.16, 93.39 and 96.57%, respectively. It occurs and temporal variabilities on the B. decumbens pasture structure managed under continuous stocking.

  15. Modelos para estimativa da área foliar de Curcuma alismatifolia e Vurcuma zedoaria Leaf area prediction models for Curcuma alismatifolia and Curcuma zedoaria

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    Ana Christina Rossini Pinto

    2008-01-01

    Full Text Available O presente estudo determina modelos para estimativa da área foliar de Curcuma alismatifolia e de Curcuma zedoaria. Para utilização destas espécies como ornamentais, é necessário o estabelecimento de técnicas de produção adequadas. Assim, a determinação da área foliar é importante, pois é usada para avaliar a resposta da planta a fatores ambientais e técnicas culturais. O uso de modelos para estimar a área foliar é um método simples, de boa precisão e não destrutivo. No estádio de floração foram coletadas cem folhas de C.alismatifolia ('Pink' e 'White' e de C.zedoaria. Determinaram-se o comprimento (C e a largura (L máximos e a área foliar real (AFR, com auxílio de integrador de área foliar (LI-3100. Estudaram-se as relações entre a AFR e o C, L e CL (produto do comprimento pela largura da folha, por meio de modelos de regressão linear. Os modelos AFR = 0,59048 CL (C.alismatifolia 'Pink', AFR = 6,08410 + 0,52162 CL (C.alismatifolia 'White' e AFR = 0,70233 CL (C.zedoaria são estatisticamente adequados para estimar a área foliar real.The present work establishes regression models to estimate leaf area of Curcuma alismatifolia and Curcuma zedoaria. To use these of species as ornamental plants is necessary to establish adequate cultivation techniques. Thus, the determination of leaf area is very important, once it is used to evaluate plant response to environmental factors and crop techniques. The use of prediction models to estimate leaf area is a simple, accurate and nondestructive method. At the stage of flowering, a hundred leaves of C.alismatifolia ('Pink' and 'White' and C.zedoaria were collected for each species and cultivar. Maximum length (L, maximum width (W and real leaf area (RLA were measured with a leaf area meter (LI-3100. The relation between RLA and the L, W and the product of length by width (LW, was studied through linear regression models. The models RLA = 0.59048 LW (C.alismatifolia 'Pink', RLA = 6

  16. Seasonal variation of photosynthetic model parameters and leaf area index from global Fluxnet eddy covariance data

    Science.gov (United States)

    Groenendijk, M.; Dolman, A. J.; Ammann, C.; Arneth, A.; Cescatti, A.; Dragoni, D.; Gash, J. H. C.; Gianelle, D.; Gioli, B.; Kiely, G.; Knohl, A.; Law, B. E.; Lund, M.; Marcolla, B.; van der Molen, M. K.; Montagnani, L.; Moors, E.; Richardson, A. D.; Roupsard, O.; Verbeeck, H.; Wohlfahrt, G.

    2011-12-01

    Global vegetation models require the photosynthetic parameters, maximum carboxylation capacity (Vcm), and quantum yield (α) to parameterize their plant functional types (PFTs). The purpose of this work is to determine how much the scaling of the parameters from leaf to ecosystem level through a seasonally varying leaf area index (LAI) explains the parameter variation within and between PFTs. Using Fluxnet data, we simulate a seasonally variable LAIF for a large range of sites, comparable to the LAIM derived from MODIS. There are discrepancies when LAIF reach zero levels and LAIM still provides a small positive value. We find that temperature is the most common constraint for LAIF in 55% of the simulations, while global radiation and vapor pressure deficit are the key constraints for 18% and 27% of the simulations, respectively, while large differences in this forcing still exist when looking at specific PFTs. Despite these differences, the annual photosynthesis simulations are comparable when using LAIF or LAIM (r2 = 0.89). We investigated further the seasonal variation of ecosystem-scale parameters derived with LAIF. Vcm has the largest seasonal variation. This holds for all vegetation types and climates. The parameter α is less variable. By including ecosystem-scale parameter seasonality we can explain a considerable part of the ecosystem-scale parameter variation between PFTs. The remaining unexplained leaf-scale PFT variation still needs further work, including elucidating the precise role of leaf and soil level nitrogen.

  17. Simultaneous growth of self-patterned carbon nanotube forests with dual height scales

    Science.gov (United States)

    Sam, Ebru Devrim; Kucukayan-Dogu, Gokce; Baykal, Beril; Dalkilic, Zeynep; Rana, Kuldeep; Bengu, Erman

    2012-05-01

    In this study, we report on a unique, one-step fabrication technique enabling the simultaneous synthesis of vertically aligned multi-walled carbon nanotubes (VA-MWCNTs) with dual height scales through alcohol catalyzed chemical vapor deposition (ACCVD). Regions of VA-MWCNTs with different heights were well separated from each other leading to a self-patterning on the surface. We devised a unique layer-by-layer process for application of catalyst and inhibitor precursors on oxidized Si (100) surfaces before the ACCVD step to achieve a hierarchical arrangement. Patterning could be controlled by adjusting the molarity and application sequence of precursors. Contact angle measurements on these self-patterned surfaces indicated that manipulation of these hierarchical arrays resulted in a wide range of hydrophobic behavior changing from that of a sticky rose petal to a lotus leaf.In this study, we report on a unique, one-step fabrication technique enabling the simultaneous synthesis of vertically aligned multi-walled carbon nanotubes (VA-MWCNTs) with dual height scales through alcohol catalyzed chemical vapor deposition (ACCVD). Regions of VA-MWCNTs with different heights were well separated from each other leading to a self-patterning on the surface. We devised a unique layer-by-layer process for application of catalyst and inhibitor precursors on oxidized Si (100) surfaces before the ACCVD step to achieve a hierarchical arrangement. Patterning could be controlled by adjusting the molarity and application sequence of precursors. Contact angle measurements on these self-patterned surfaces indicated that manipulation of these hierarchical arrays resulted in a wide range of hydrophobic behavior changing from that of a sticky rose petal to a lotus leaf. Electronic supplementary information (ESI) available: Fig. S1; AFM image of the Co-O layer which was first dried at 40 °C and then oxidized at 200 °C. Fig. S2; graph relative to the area of CNT islands for different

  18. Genetic variance of sunflower yield components - Heliantus annuus L.

    Directory of Open Access Journals (Sweden)

    Hladni Nada

    2003-01-01

    Full Text Available The main goals of sunflower breeding in Yugoslavia and abroad are increased seed yield and oil content per unit area and increased resistance to diseases, insects and stress conditions via an optimization of plant architecture. In order to determine the mode of inheritance, gene effects and correlations of total leaf number per plant, total leaf area and plant height, six genetically divergent inbred lines of sunflower were subjected to half diallel crosses. Significant differences in mean values of all the traits were found in the F1 and F2 generations. Additive gene effects were more important in the inheritance of total leaf number per plant and plant height, while in the case of total leaf area per plant the nonadditive ones were more important looking at all the combinations in the F1 and F2 generations. The average degree of dominance (Hi/D1/2 was lower than one for total leaf number per plant and plant height, so the mode of inheritance was partial dominance, while with total leaf area the value was higher than one, indicating super dominance as the mode of inheritance. Significant positive correlation was found: between total leaf area per plant and total leaf number per plant (0.285* and plant height (0.278*. The results of the study are of importance for further sunflower breeding work.

  19. Scaling up stomatal conductance from leaf to canopy using a dual-leaf model for estimating crop evapotranspiration.

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    Risheng Ding

    Full Text Available The dual-source Shuttleworth-Wallace model has been widely used to estimate and partition crop evapotranspiration (λET. Canopy stomatal conductance (Gsc, an essential parameter of the model, is often calculated by scaling up leaf stomatal conductance, considering the canopy as one single leaf in a so-called "big-leaf" model. However, Gsc can be overestimated or underestimated depending on leaf area index level in the big-leaf model, due to a non-linear stomatal response to light. A dual-leaf model, scaling up Gsc from leaf to canopy, was developed in this study. The non-linear stomata-light relationship was incorporated by dividing the canopy into sunlit and shaded fractions and calculating each fraction separately according to absorbed irradiances. The model includes: (1 the absorbed irradiance, determined by separately integrating the sunlit and shaded leaves with consideration of both beam and diffuse radiation; (2 leaf area for the sunlit and shaded fractions; and (3 a leaf conductance model that accounts for the response of stomata to PAR, vapor pressure deficit and available soil water. In contrast to the significant errors of Gsc in the big-leaf model, the predicted Gsc using the dual-leaf model had a high degree of data-model agreement; the slope of the linear regression between daytime predictions and measurements was 1.01 (R2 = 0.98, with RMSE of 0.6120 mm s-1 for four clear-sky days in different growth stages. The estimates of half-hourly λET using the dual-source dual-leaf model (DSDL agreed well with measurements and the error was within 5% during two growing seasons of maize with differing hydrometeorological and management strategies. Moreover, the estimates of soil evaporation using the DSDL model closely matched actual measurements. Our results indicate that the DSDL model can produce more accurate estimation of Gsc and λET, compared to the big-leaf model, and thus is an effective alternative approach for estimating and

  20. Leaf habit and woodiness regulate different leaf economy traits at a given nutrient supply.

    Science.gov (United States)

    Ordoñez, Jenny C; van Bodegom, Peter M; Witte, Jan-Philip M; Bartholomeus, Ruud P; van Dobben, Han F; Aerts, Rien

    2010-11-01

    The large variation in the relationships between environmental factors and plant traits observed in natural communities exemplifies the alternative solutions that plants have developed in response to the same environmental limitations. Qualitative attributes, such as growth form, woodiness, and leaf habit can be used to approximate these alternative solutions. Here, we quantified the extent to which these attributes affect leaf trait values at a given resource supply level, using measured plant traits from 105 different species (254 observations) distributed across 50 sites in mesic to wet plant communities in The Netherlands. For each site, soil total N, soil total P, and water supply estimates were obtained by field measurements and modeling. Effects of growth forms, woodiness, and leaf habit on relations between leaf traits (SLA, specific leaf area; LNC, leaf nitrogen concentration; and LPC, leaf phosphorus concentration) vs. nutrient and water supply were quantified using maximum-likelihood methods and Bonferroni post hoc tests. The qualitative attributes explained 8-23% of the variance within sites in leaf traits vs. soil fertility relationships, and therefore they can potentially be used to make better predictions of global patterns of leaf traits in relation to nutrient supply. However, at a given soil fertility, the strength of the effect of each qualitative attribute was not the same for all leaf traits. These differences may imply a differential regulation of the leaf economy traits at a given nutrient supply, in which SLA and LPC seem to be regulated in accordance to changes in plant size and architecture while LNC seems to be primarily regulated at the leaf level by factors related to leaf longevity.

  1. Simulation Models of Leaf Area Index and Yield for Cotton Grown with Different Soil Conditioners.

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    Lijun Su

    Full Text Available Simulation models of leaf area index (LAI and yield for cotton can provide a theoretical foundation for predicting future variations in yield. This paper analyses the increase in LAI and the relationships between LAI, dry matter, and yield for cotton under three soil conditioners near Korla, Xinjiang, China. Dynamic changes in cotton LAI were evaluated using modified logistic, Gaussian, modified Gaussian, log normal, and cubic polynomial models. Universal models for simulating the relative leaf area index (RLAI were established in which the application rate of soil conditioner was used to estimate the maximum LAI (LAIm. In addition, the relationships between LAIm and dry matter mass, yield, and the harvest index were investigated, and a simulation model for yield is proposed. A feasibility analysis of the models indicated that the cubic polynomial and Gaussian models were less accurate than the other three models for simulating increases in RLAI. Despite significant differences in LAIs under the type and amount of soil conditioner applied, LAIm could be described by aboveground dry matter using Michaelis-Menten kinetics. Moreover, the simulation model for cotton yield based on LAIm and the harvest index presented in this work provided important theoretical insights for improving water use efficiency in cotton cultivation and for identifying optimal application rates of soil conditioners.

  2. Effects of LiDAR point density, sampling size and height threshold on estimation accuracy of crop biophysical parameters.

    Science.gov (United States)

    Luo, Shezhou; Chen, Jing M; Wang, Cheng; Xi, Xiaohuan; Zeng, Hongcheng; Peng, Dailiang; Li, Dong

    2016-05-30

    Vegetation leaf area index (LAI), height, and aboveground biomass are key biophysical parameters. Corn is an important and globally distributed crop, and reliable estimations of these parameters are essential for corn yield forecasting, health monitoring and ecosystem modeling. Light Detection and Ranging (LiDAR) is considered an effective technology for estimating vegetation biophysical parameters. However, the estimation accuracies of these parameters are affected by multiple factors. In this study, we first estimated corn LAI, height and biomass (R2 = 0.80, 0.874 and 0.838, respectively) using the original LiDAR data (7.32 points/m2), and the results showed that LiDAR data could accurately estimate these biophysical parameters. Second, comprehensive research was conducted on the effects of LiDAR point density, sampling size and height threshold on the estimation accuracy of LAI, height and biomass. Our findings indicated that LiDAR point density had an important effect on the estimation accuracy for vegetation biophysical parameters, however, high point density did not always produce highly accurate estimates, and reduced point density could deliver reasonable estimation results. Furthermore, the results showed that sampling size and height threshold were additional key factors that affect the estimation accuracy of biophysical parameters. Therefore, the optimal sampling size and the height threshold should be determined to improve the estimation accuracy of biophysical parameters. Our results also implied that a higher LiDAR point density, larger sampling size and height threshold were required to obtain accurate corn LAI estimation when compared with height and biomass estimations. In general, our results provide valuable guidance for LiDAR data acquisition and estimation of vegetation biophysical parameters using LiDAR data.

  3. Environmental parameters affecting the structure of leaf-litter frog (Amphibia: Anura communities in tropical forests: a case study from an Atlantic Rainforest area in southeastern Brazil

    Directory of Open Access Journals (Sweden)

    Carla C. Siqueira

    2014-04-01

    Full Text Available Despite a recent increase of information on leaf litter frog communities from Atlantic rainforests, few studies have analyzed the relationship between environmental parameters and community structure of these animals. We analyzed the effects of some environmental factors on a leaf litter frog community at an Atlantic Rainforest area in southeastern Brazil. Data collection lasted ten consecutive days in January 2010, at elevations ranging between 300 and 520 m above sea level. We established 50 quadrats of 5 x 5 m on the forest floor, totaling 1,250 m² of sampled area, and recorded the mean leaf-litter depth and the number of trees within the plot, as well as altitude. We found 307 individuals belonging to ten frog species within the plots. The overall density of leaf-litter frogs estimated from the plots was 24.6 ind/100m², with Euparkerella brasiliensis (Parker, 1926, Ischnocnema guentheri (Steindachner, 1864, Ischnocnema parva (Girard, 1853 and Haddadus binotatus (Spix, 1824 presenting the highest estimated densities. Among the environmental variables analyzed, only altitude influenced the parameters of anuran community. Our results indicate that the study area has a very high density of forest floor leaf litter frogs at altitudes of 300-500 m. Future estimates of litter frog density might benefit from taking the local altitudinal variation into consideration. Neglecting such variation might result in underestimated/overestimated values if they are extrapolated to the whole area.

  4. NOAA Climate Data Record (CDR) of Leaf Area Index (LAI) and Fraction of Absorbed Photosynthetically Active Radiation (FAPAR), Version 4

    Data.gov (United States)

    National Oceanic and Atmospheric Administration, Department of Commerce — This dataset contains gridded daily Leaf Area Index (LAI) and Fraction of Absorbed Photosynthetically Active Radiation (FAPAR) derived from the NOAA Climate Data...

  5. Variation in chlorophyll content per unit leaf area in spring wheat and implications for selection in segregating material.

    Directory of Open Access Journals (Sweden)

    John Hamblin

    Full Text Available Reduced levels of leaf chlorophyll content per unit leaf area in crops may be of advantage in the search for higher yields. Possible reasons include better light distribution in the crop canopy and less photochemical damage to leaves absorbing more light energy than required for maximum photosynthesis. Reduced chlorophyll may also reduce the heat load at the top of canopy, reducing water requirements to cool leaves. Chloroplasts are nutrient rich and reducing their number may increase available nutrients for growth and development. To determine whether this hypothesis has any validity in spring wheat requires an understanding of genotypic differences in leaf chlorophyll content per unit area in diverse germplasm. This was measured with a SPAD 502 as SPAD units. The study was conducted in series of environments involving up to 28 genotypes, mainly spring wheat. In general, substantial and repeatable genotypic variation was observed. Consistent SPAD readings were recorded for different sampling positions on leaves, between different leaves on single plant, between different plants of the same genotype, and between different genotypes grown in the same or different environments. Plant nutrition affected SPAD units in nutrient poor environments. Wheat genotypes DBW 10 and Transfer were identified as having consistent and contrasting high and low average SPAD readings of 52 and 32 units, respectively, and a methodology to allow selection in segregating populations has been developed.

  6. Forest height estimation from mountain forest areas using general model-based decomposition for polarimetric interferometric synthetic aperture radar images

    Science.gov (United States)

    Minh, Nghia Pham; Zou, Bin; Cai, Hongjun; Wang, Chengyi

    2014-01-01

    The estimation of forest parameters over mountain forest areas using polarimetric interferometric synthetic aperture radar (PolInSAR) images is one of the greatest interests in remote sensing applications. For mountain forest areas, scattering mechanisms are strongly affected by the ground topography variations. Most of the previous studies in modeling microwave backscattering signatures of forest area have been carried out over relatively flat areas. Therefore, a new algorithm for the forest height estimation from mountain forest areas using the general model-based decomposition (GMBD) for PolInSAR image is proposed. This algorithm enables the retrieval of not only the forest parameters, but also the magnitude associated with each mechanism. In addition, general double- and single-bounce scattering models are proposed to fit for the cross-polarization and off-diagonal term by separating their independent orientation angle, which remains unachieved in the previous model-based decompositions. The efficiency of the proposed approach is demonstrated with simulated data from PolSARProSim software and ALOS-PALSAR spaceborne PolInSAR datasets over the Kalimantan areas, Indonesia. Experimental results indicate that forest height could be effectively estimated by GMBD.

  7. Final report on the safety assessment of AloeAndongensis Extract, Aloe Andongensis Leaf Juice,aloe Arborescens Leaf Extract, Aloe Arborescens Leaf Juice, Aloe Arborescens Leaf Protoplasts, Aloe Barbadensis Flower Extract, Aloe Barbadensis Leaf, Aloe Barbadensis Leaf Extract, Aloe Barbadensis Leaf Juice,aloe Barbadensis Leaf Polysaccharides, Aloe Barbadensis Leaf Water, Aloe Ferox Leaf Extract, Aloe Ferox Leaf Juice, and Aloe Ferox Leaf Juice Extract.

    Science.gov (United States)

    2007-01-01

    Plant materials derived from the Aloe plant are used as cosmetic ingredients, including Aloe Andongensis Extract, Aloe Andongensis Leaf Juice, Aloe Arborescens Leaf Extract, Aloe Arborescens Leaf Juice, Aloe Arborescens Leaf Protoplasts, Aloe Barbadensis Flower Extract, Aloe Barbadensis Leaf, Aloe Barbadensis Leaf Extract, Aloe Barbadensis Leaf Juice, Aloe Barbadensis Leaf Polysaccharides, Aloe Barbadensis Leaf Water, Aloe Ferox Leaf Extract, Aloe Ferox Leaf Juice, and Aloe Ferox Leaf Juice Extract. These ingredients function primarily as skin-conditioning agents and are included in cosmetics only at low concentrations. The Aloe leaf consists of the pericyclic cells, found just below the plant's skin, and the inner central area of the leaf, i.e., the gel, which is used for cosmetic products. The pericyclic cells produce a bitter, yellow latex containing a number of anthraquinones, phototoxic compounds that are also gastrointestinal irritants responsible for cathartic effects. The gel contains polysaccharides, which can be acetylated, partially acetylated, or not acetylated. An industry established limit for anthraquinones in aloe-derived material for nonmedicinal use is 50 ppm or lower. Aloe-derived ingredients are used in a wide variety of cosmetic product types at concentrations of raw material that are 0.1% or less, although can be as high as 20%. The concentration of Aloe in the raw material also may vary from 100% to a low of 0.0005%. Oral administration of various anthraquinone components results in a rise in their blood concentrations, wide systemic distribution, accumulation in the liver and kidneys, and excretion in urine and feces; polysaccharide components are distributed systemically and metabolized into smaller molecules. aloe-derived material has fungicidal, antimicrobial, and antiviral activities, and has been effective in wound healing and infection treatment in animals. Aloe barbadensis (also known as Aloe vera)-derived ingredients were not toxic

  8. The effect of water deficit stress and nitrogen fertilizer levels on morphology traits, yield and leaf area index in maize

    International Nuclear Information System (INIS)

    Moosavi, S.G.

    2012-01-01

    In order to study the effect of water deficit stress at different growth stages and N fertilizer levels on morphological traits, yield and yield components of maize cv. Single Cross 704, an experiment was conducted as a split-plot based on a Randomized Complete Block Design with three replications. The main plot included irrigation at four levels (irrigation stop at 10-leaf, tasselling and grain-filling stages and optimum irrigation) and the sub-plot was N fertilizer at three levels (75, 150 and 225 kg N/ha). The results of analysis of variance showed that water-deficit stress and N fertilizer level significantly affected leaf area index at silking stage, ear length, grain number per ear, 1000-grain weight and grain yield. Stem diameter, ear diameter and harvest index were only affected by irrigation treatments and the interaction between irrigation and N level did not significantly affect the studied traits. Means comparison indicated that ear diameter under optimum irrigation was higher than that under the treatments of irrigation stop at 8-leaf, tasselling and grain-filling stages by 29.9, 19.1 and 33.5%, respectively; and ear length was higher than them by 38.1, 28.9 and 25.2%, respectively. Moreover, the highest grain number per ear, 1000-grain weight and grain yield were obtained under optimum irrigation treatment, and irrigation stop at 10-leaf, tasselling and grain-filling stages decreased grain yield by 52.8, 66.4 and 44.9%, respectively; and it decreased grain number/ear by 45.9, 59.3 and 30.1%, respectively. In addition, optimum irrigation treatment with mean 1000-grain weight of 289.2 g was significantly superior over other irrigation stop treatments by 27.6-42.8% and produced the highest leaf area index at silking stage (4.1). Means comparison of traits at different N levels indicated that N level of 225 kg/ha produced the highest ear length (17.82 cm), grain number per ear (401.9), 1000-grain weight (258.8 g), leaf area index at silking stage (4

  9. Thematic mapper detection of changes in the leaf area of closed canopy pine plantations in central Massachusetts

    International Nuclear Information System (INIS)

    Herwitz, S.R.; Peterson, D.L.; Eastman, J.R.

    1989-01-01

    Remote sensing studies of conifer forests have previously reported that the Thematic Mapper Band 4/Band 3 ratio is positively correlated with regional differences in leaf area index (LAI). Our study was an attempt to determine whether Landsat Thematic Mapper data can be used to detect differences and changes in the LAI of closed canopy pine plantations on a local scale in central Massachusetts. Field measurements of LAI were obtained using locally-derived allometric relationships between leaf area and trunk diameter (DBH). A thinning treatment, which reduced the LAI of one of the larger plantations by more than 25%, resulted in a significant decrease (P < 0.001) in the 4/3 ratio from the prethinned value. No significant change in the 4/3 ratio was found in a nearby broadleaved hardwood forest which served as a radiometric control. However, a decrease in the 4/3 ratio similar to that observed in the thinned plantation was observed in nearby unthinned pine plantations. This change in the reflectance of the unthinned stands may be attributable to a moderate natural reduction in LAI. Such a reduction in LAI would demonstrate the limitations of allometric equations for evaluating LAI under conditions in which the relationship between leaf area and DBH may be changing from year to year. It also would explain why no significant relationship (P > 0.1) was found between the 4/3 ratio and the LAI of the different unthinned plantations which had LAI values ranging from 3.96 to 7.01. We conclude that the TM sensor may be a better guide to moderate changes and differences in the LAI of closed canopy pine plantations at local scales than field measurements involving allometric equations. (author)

  10. Prediction of the competitive effects of weeds on crop yields based on the relative leaf area of weeds

    DEFF Research Database (Denmark)

    Lotz, L. A. P.; Christensen, Svend; Cloutier, D.

    1996-01-01

    . alba whereas the density model did not. A parameter that allows the maximum yield loss to be smaller than 100% was mostly not needed to describe the effects of weed competition. The parameter that denotes the competitiveness of the weed species with respect to the crop decreased the later the relative......For implementation of simple yield loss models into threshold-based weed management systems, a thorough validation is needed over a great diversity of sites. Yield losses by competition wsth Sinapis alba L. (white mustard) as a model weed, were studied in 12 experiments in sugar beet (Beta vulgaris...... L.) and in 11 experiments in spring wheat (Triticum aestivum L.). Most data sets were heller described by a model based on the relative leaf area of the weed than by a hyperbolic model based on weed density. This leaf area model accounted for (part of) the effect of different emerging times of the S...

  11. Can adaptive modulation of traits to urban environments facilitate Ricinus communis L. invasiveness?

    Science.gov (United States)

    Goyal, Neha; Pardha-Saradhi, P; Sharma, Gyan P

    2014-11-01

    This paper addresses the phenotypic variation among Ricinus communis L. populations in four urban habitat types (road verges, garbage dumps, construction debris, and natural area) in Delhi, India, by evaluating important traits such as plant height, basal circumference, seeds per plant, seed size, seed weight, specific leaf area, and reproductive index. An important biochemical marker, proline, considered as a good plant performance indicator under stress was also quantified in leaves of R. communis to evaluate its response in different habitats. Interestingly, the species showed significant variation in plant height, specific leaf area, seed size, seed weight, and leaf proline content in different habitat types. Leaf proline content was positively related to plant height, specific leaf area, and seed size while negatively related to the total number of seeds/plant. Interestingly, reproductive index, calculated as a ratio of the total number of seeds to the plant height also showed a negative relation with leaf proline content. Results indicated that R. communis exhibits adaptive modulation of growth, reproductive traits, and leaf proline content in various urban habitats which contributes to invasiveness, range expansion, and establishment of the species. The study also gives evidence of how morphological and physiological traits could directly affect invasiveness of R. communis.

  12. Worldwide Historical Estimates of Leaf Area Index, 1932-2000

    Science.gov (United States)

    Scurlock, J. M. O.; Asner, G. P.; Gower, S. T.

    2001-01-01

    Approximately 1000 published estimates of leaf area index (LAI) from nearly 400 unique field sites, covering the period 1932-2000, have been compiled into a single data set. LA1 is a key parameter for global and regional models of biosphere/atmosphere exchange of carbon dioxide, water vapor, and other materials. It also plays an integral role in determining the energy balance of the land surface. This data set provides a benchmark of typical values and ranges of LA1 for a variety of biomes and land cover types, in support of model development and validation of satellite-derived remote sensing estimates of LA1 and other vegetation parameters. The LA1 data are linked to a bibliography of over 300 originalsource references.This report documents the development of this data set, its contents, and its availability on the Internet from the Oak Ridge National Laboratory Distributed Active Archive Center for Biogeochemical Dynamics. Caution is advised in using these data, which were collected using a wide range of methodologies and assumptions that may not allow comparisons among sites.

  13. Leaf Area Index (LAI Estimation of Boreal Forest Using Wide Optics Airborne Winter Photos

    Directory of Open Access Journals (Sweden)

    Pauline Stenberg

    2009-12-01

    Full Text Available A new simple airborne method based on wide optics camera is developed for leaf area index (LAI estimation in coniferous forests. The measurements are carried out in winter, when the forest floor is completely snow covered and thus acts as a light background for the hemispherical analysis of the images. The photos are taken automatically and stored on a laptop during the flights. The R2 value of the linear regression of the airborne and ground based LAI measurements was 0.89.

  14. Effect of Wind on the Relation of Leaf N, P Stoichiometry with Leaf Morphology in Quercus Species

    Directory of Open Access Journals (Sweden)

    Peng Zhang

    2018-02-01

    Full Text Available Leaf nitrogen (N and phosphorus (P stoichiometry correlates closely to leaf morphology, which is strongly impacted by wind at multiple scales. However, it is not clear how leaf N, P stoichiometry and its relationship to leaf morphology changes with wind load. We determined the leaf N and P concentrations and leaf morphology—including specific leaf area (SLA and leaf dissection index (LDI—for eight Quercus species under a simulated wind load for seven months. Leaf N and P concentrations increased significantly under these conditions for Quercus acutissima, Quercus rubra, Quercus texana, and Quercus palustris—which have elliptic leaves—due to their higher N, P requirements and a resultant leaf biomass decrease, which is a tolerance strategy for Quercus species under a wind load. Leaf N:P was relatively stable under wind for all species, which supports stoichiometric homeostasis. Leaf N concentrations showed a positive correlation to SLA, leaf N and P concentrations showed positive correlations to LDI under each wind treatment, and the slope of correlations was not affected by wind, which indicates synchronous variations between leaf stoichiometry and leaf morphology under wind. However, the intercept of correlations was affected by wind, and leaf N and P use efficiency decreased under the wind load, which suggests that the Quercus species changes from “fast investment-return” in the control to “slow investment-return” under windy conditions. These results will be valuable to understanding functional strategies for plants under varying wind loads, especially synchronous variations in leaf traits along a wind gradient.

  15. Leaf area index estimation with MODIS reflectance time series and model inversion during full rotations of Eucalyptus plantations

    NARCIS (Netherlands)

    Maire, Le G.; Marsden, C.; Verhoef, W.; Ponzoni, F.J.; Seen, Lo D.; Bégué, A.; Stape, J.L.; Nouvellon, Y.

    2011-01-01

    The leaf area index (LAI) of fast-growing Eucalyptus plantations is highly dynamic both seasonally and inter-annually, and is spatially variable depending on pedo-climatic conditions. LAI is very important in determining the carbon and water balance of a stand, but is difficult to measure during a

  16. Leaf Potential Productivity at Different Canopy Levels in Densely-planted and Intermediately-thinned Apple Orchards

    Directory of Open Access Journals (Sweden)

    Ying SUN

    2016-07-01

    Full Text Available Most apple orchards in the apple production districts in China were densely planted with vigorous rootstocks during the 1980s. These orchards have suffered micro-environmental deterioration and loss of fruit quality because of the closed canopy. Modification of the densely-planted orchards is a priority in current apple production. Intermediate thinning is a basic technique used to transform densely-planted apple orchards in China. Our goal was to provide theoretical basis for studying the effect of thinning on the efficiency of photosynthetically active radiation (PAR, fruit quality, and yield. We measured leaf area, solar radiation, and leaf air exchange at different tree canopy levels and by fitting relevant photosynthetic models, vertical distribution characteristics of leaf photosynthetic potentials and PAR were analyzed in various levels within canopies in densely-planted and intermediately-thinned orchards. Intermediate thinning significantly improved the radiant environment inside the canopies. PAR distribution within the canopies in the intermediately-thinned orchard was better distributed than in the densely-planted orchards. The invalid space under 30.0% of relative photosynthetically active radiation (PARr was nearly zero in the intermediately-thinned orchard; but minimum PARr was 17.0% and the space under 0.30 of the relative height of the canopy was invalid for photosynthesis in the densely-planted orchard. The leaf photosynthetic efficiency in the intermediately-thinned orchard was improved. Photosynthetic rates (Pn at the middle and bottom levels of the canopy, respectively, were increased by 7.80% and 10.20% in the intermediately-thinned orchard. Leaf development, which influences photosynthetic potential, was closely related to the surrounding micro-environment, especially light. Leaf photosynthetic potentials were correlated with leaf nitrogen content (Nl and specific leaf weight (Ml at various levels of canopies. Compared

  17. Canopy characteristics, animal behavior and forage intake by goats grazing on Tanzania-grass pasture with different heights - doi: 10.4025/actascianimsci.v34i4.14544

    Directory of Open Access Journals (Sweden)

    Maurílio Souza dos Santos

    2012-10-01

    Full Text Available This study evaluated the influence of Tanzania-grass sward height (30, 50, 70 and 90 cm on the morphological characteristics of the canopy, grazing behavior and forage intake by adult Anglo Nubian female goats. A completely randomized experimental design was employed, with two replicates in space and two replicates in time. Six animals were used to assess the grazing behavior, and four, the ingestion process. The rise in sward height increased the forage and leaf mass, the percentages of stem and dead material, and reduced the leaf stem-1 ratio. Above 50 cm there was an increase in grazing time and a decrease in leisure time. A positive linear correlation was detected between sward height and bite depth. The consumed forage mass, ingestion rate and daily intake were higher at 50 cm, indicating that the other heights reduced the intake process. The sward height was negatively correlated to the bite rate and positively to the bite time. The sward height of 50 cm presents the best combination of features, favoring the grazing and ingestive behavior of female adult goats.

  18. Determining the K coefficient to leaf area index estimations in a tropical dry forest

    Science.gov (United States)

    Magalhães, Sarah Freitas; Calvo-Rodriguez, Sofia; do Espírito Santo, Mário Marcos; Sánchez Azofeifa, Gerardo Arturo

    2018-03-01

    Vegetation indices are useful tools to remotely estimate several important parameters related to ecosystem functioning. However, improving and validating estimations for a wide range of vegetation types are necessary. In this study, we provide a methodology for the estimation of the leaf area index (LAI) in a tropical dry forest (TDF) using the light diffusion through the canopy as a function of the successional stage. For this purpose, we estimated the K coefficient, a parameter that relates the normalized difference vegetation index (NDVI) to LAI, based on photosynthetically active radiation (PAR) and solar radiation. The study was conducted in the Mata Seca State Park, in southeastern Brazil, from 2012 to 2013. We defined four successional stages (very early, early, intermediate, and late) and established one optical phenology tower at one plot of 20 × 20 m per stage. Towers measured the incoming and reflected solar radiation and PAR for NDVI calculation. For each plot, we established 24 points for LAI sampling through hemispherical photographs. Because leaf cover is highly seasonal in TDFs, we determined ΔK (leaf growth phase) and K max (leaf maturity phase). We detected a strong correlation between NDVI and LAI, which is necessary for a reliable determination of the K coefficient. Both NDVI and LAI varied significantly between successional stages, indicating sensitivity to structural changes in forest regeneration. Furthermore, the K values differed between successional stages and correlated significantly with other environmental variables such as air temperature and humidity, fraction of absorbed PAR, and soil moisture. Thus, we established a model based on spectral properties of the vegetation coupled with biophysical characteristics in a TDF that makes possible to estimate LAI from NDVI values. The application of the K coefficient can improve remote estimations of forest primary productivity and gases and energy exchanges between vegetation and atmosphere

  19. Leaf anatomical traits determine the 18O enrichment of leaf water in coastal halophytes

    Science.gov (United States)

    Liang, J.; Lin, G., Sr.; Sternberg, L. O.

    2017-12-01

    Foliar anatomical adaptations to high-salinity environment in mangroves may be recorded by leaf water isotopes. Recent studies observed that a few mangrove species have lower 18O enrichment of leaf water (ΔL) relative to source water than the adjacent terrestrial trees, but what factors actually control this phenomenon is still disputable at present. To resolve this issue, we collected 15 species of true mangrove plants, 14 species of adjacent freshwater trees and 4 species of semi-mangrove plants at five study sites on the southeastern coast of China. Leaf stomatal density and pore size, water content, ΔL and other related leaf physiological traits were determined for the selected leaves of these plants. Our results confirmed that ΔL values of mangroves were generally 3 4 ‰ lower than those of the adjacent freshwater or semi-mangrove species. Higher leaf water per area (LWC) and lower leaf stomatal density (LS) of mangroves played co-dominant roles in lowering ΔL through elongating effective leaf mixing length by about 20%. The Péclet model incorporated by LWC and LS performed well in predicting ΔL. The demonstrated general law between leaf anatomy and ΔL in this paper based on a large pool of species bridges the gap between leaf functional traits and metabolic proxies derived ΔL, which will have considerable potential applications in vegetation succession and reconstruction of paleoclimate research.

  20. Interacting Effects of Leaf Water Potential and Biomass on Vegetation Optical Depth

    Science.gov (United States)

    Momen, M.; Wood, J. D.; Novick, K. A.; Pockman, W.; Konings, A. G.

    2017-12-01

    Remotely-sensed microwave observations of vegetation optical depth (VOD) have been widely used to examine vegetation responses to climate. Such studies have alternately found that VOD is sensitive to both biomass and canopy water content. However, the relative impacts of changes in phenology or water stress on VOD have not been disentangled. In particular, understanding whether leaf water potential (LWP) affects VOD may permit the assimilation of satellite observations into new large-scale plant hydraulic models. Despite extensive validation of the relationship between satellite-derived VOD estimates and vegetation density, relatively few studies have explicitly sought to validate the sensitivity of VOD to canopy water status, and none have studied the effect of variations in LWP on VOD. In this work, we test the sensitivity of VOD to variations in LWP, and present a conceptual framework which relates VOD to a combination of leaf water potential and total biomass including leaves, whose dynamics can be measured through leaf area index, and woody biomass. We used in-situ measurements of LWP data to validate the conceptual model in mixed deciduous forests in Indiana and Missouri, as well as a pinion-juniper woodland in New Mexico. Observed X-band VOD from the AMSR-E and AMSR2 satellites showed dynamics similar to those reconstructed VOD signals based on the new conceptual model which employs in-situ LWP data (R2=0.60-0.80). Because LWP data are not available at global scales, we further estimated ecosystem LWP based on remotely sensed surface soil moisture to better understand the sensitivity of VOD across ecosystems. At the global scale, incorporating a combination of biomass and water potential in the reconstructed VOD signal increased correlations with VOD about 15% compared to biomass alone and about 30% compared to water potential alone. In wetter regions with denser and taller canopy heights, VOD has a higher correlation with leaf area index than with water

  1. Leaf, woody, and root biomass of Populus irrigated with landfill leachate

    Science.gov (United States)

    Jill A. Zalesny; Ronald S., Jr. Zalesny; D.R. Coyle; R.B. Hall

    2007-01-01

    Poplar (Populus spp.) trees can be utilized for ecological leachate disposal when applied as an irrigation source for managed tree systems. Our objective was to evaluate differences in tree height, diameter, volume, and biomass of leaf, stem, branch, and root tissues of Populus trees after two seasons of irrigation with municipal...

  2. Yielding of leaf celery Apium graveolens L. var. secalinum Alef. depending on the number of harvests and irrigation

    Directory of Open Access Journals (Sweden)

    Ewa Rożek

    2013-04-01

    Full Text Available Leaf celery (Apium graveolensvar. secalinum is a vegetable with medicinal and spicy properties. Its numerous intensely fragrant leaves can be cut several times during the plant growing period. The aim of this study was to evaluate the effect of irrigation and number of harvests on leaf celery yield of the cultivars ‘Afina’ and‘Gewone Snij’. Plant irrigation significantly increased leaf yield and plant height of leaf celery. Higher total yield was obtained from non-irrigated plants when leaves were harvested three times, whereas for irrigated plants yield was higher in the case of two leaf harvests. Irrespective of the experimental factors, higher yield was obtained from the cultivar ‘Gewone Snij’.

  3. Leaf Wetness within a Lily Canopy

    NARCIS (Netherlands)

    Jacobs, A.F.G.; Heusinkveld, B.G.; Klok, E.J.

    2005-01-01

    A wetness duration experiment was carried out within a lily field situated adjacent to coastal dunes in the Netherlands. A within-canopy model was applied to simulate leaf wetness in three layers, with equal leaf area indices, within the canopy. This simulation model is an extension of an existing

  4. Estimativa da área da folha da batateira utilizando medidas lineares Evaluation of the potato plant leaf area using linear measures

    Directory of Open Access Journals (Sweden)

    Marcelo CC Silva

    2008-03-01

    Full Text Available O objetivo deste experimento foi determinar o modelo mais apropriado para estimar a área da folha da batateira, utilizando-se medidas de comprimento e largura da folha. Foram coletadas 300 folhas de 300 plantas de batata, cultivar Monalisa, de forma aleatória, aos 21 e 56 dias após a emergência (DAE. Em laboratório, foram medidos o comprimento (C, a largura (L e a área de cada folha (AF. Os dados foram submetidos à análise de regressão com o valor da AF sendo considerado a variável dependente e os valores de comprimento e largura de folha as variáveis independentes. Foram testados três modelos estatísticos: linear, exponencial e logarítmico. A AF da batateira foi mais precisamente estimada (R² = 0,88, usando as medidas, L e C (AF = 0,2798**LC + 71,267. Para maior rapidez e praticidade, a AF da batateira, foi também apropriadamente estimada medindo-se apenas L ou C da folha e utilizando-se as equações AF = 0,0479**L + 10,777 (R² = 0,83 ou AF = 0,0659**C + 12,979 (R² = 0,82. A área foliar estimada 21 DAE, utilizando o modelo linear foi de 234,41 cm², sendo que o valor real medido, foi de 185,52 cm². Aos 56 DAE, a área foliar estimada pelo mesmo modelo foi de 175,60 cm², o valor real medido, foi de 176,01 cm². Com um dos modelos propostos, a área da folha pode ser estimada em tempo real, de forma rápida e sem a necessidade de coletar a folha.The objective of this experiment was to determine the most appropriate model to estimate potato leaf area through the leaf length and width. 300 leaves of 300 potato plants, cv. Monalisa were collected in an aleatory way, 21 and 56 days after the plant emergence (DAE. In laboratory, the length (C, width (L and area of each leaf (AF were measured. The data were submitted to the regression analysis with the AF value as a dependent variable and the leaf length and width values as the independent variables. Three statistical models were tested (linear, exponential and logarithmic. Potato

  5. A leaf gas exchange model that accounts for intra-canopy variability by considering leaf nitrogen content and local acclimation to radiation in grapevine (Vitis vinifera L.).

    Science.gov (United States)

    Prieto, Jorge A; Louarn, Gaëtan; Perez Peña, Jorge; Ojeda, Hernán; Simonneau, Thierry; Lebon, Eric

    2012-07-01

    Understanding the distribution of gas exchange within a plant is a prerequisite for scaling up from leaves to canopies. We evaluated whether leaf traits were reliable predictors of the effects of leaf ageing and leaf irradiance on leaf photosynthetic capacity (V(cmax) , J(max) ) in field-grown vines (Vitis vinifera L). Simultaneously, we measured gas exchange, leaf mass per area (LMA) and nitrogen content (N(m) ) of leaves at different positions within the canopy and at different phenological stages. Daily mean leaf irradiance cumulated over 10 d (PPFD(10) ) was obtained by 3D modelling of the canopy structure. N(m) decreased over the season in parallel to leaf ageing while LMA was mainly affected by leaf position. PPFD(10) explained 66, 28 and 73% of the variation of LMA, N(m) and nitrogen content per area (N(a) ), respectively. Nitrogen content per unit area (N(a) = LMA × N(m) ) was the best predictor of the intra-canopy variability of leaf photosynthetic capacity. Finally, we developed a classical photosynthesis-stomatal conductance submodel and by introducing N(a) as an input, the model accurately simulated the daily pattern of gas exchange for leaves at different positions in the canopy and at different phenological stages during the season. © 2012 Blackwell Publishing Ltd.

  6. Serious complications in experiments in which UV doses are effected by using different lamp heights.

    Science.gov (United States)

    Flint, Stephan D; Ryel, Ronald J; Hudelson, Timothy J; Caldwell, Martyn M

    2009-10-06

    Many experiments examining plant responses to enhanced ultraviolet-B radiation (280-315nm) simply compare an enhanced UV-B treatment with ambient UV-B (or no UV-B radiation in most greenhouse and controlled-environment studies). Some more detailed experiments utilize multiple levels of UV-B radiation. A number of different techniques have been used to adjust the UV dose. One common technique is to place racks of fluorescent UV-emitting lamps at different heights above the plant canopy. However, the lamps and associated support structure cast shadows on the plant bed below. We calculated one example of the sequence of shade intervals for two common heights of lamp racks and show the patterns and duration of shade which the plants receive is distributed differently over the course of the day for different heights of the lamp racks. We also conducted a greenhouse experiment with plants (canola, sunflower and maize) grown under unenergized lamp racks suspended at the same two heights above the canopy. Growth characteristics differed in unpredictable ways between plants grown under the two heights of lamp racks. These differences could enhance or obscure potential UV-B effects. Also, differences in leaf mass per unit foliage area, which were observed in this experiment, could contribute to differences in plant UV-B sensitivity. We recommend the use of other techniques for achieving multiple doses of UV-B radiation. These range from simple and inexpensive approaches (e.g., wrapping individual fluorescent tubes in layers of a neutral-density filter such as cheese cloth) to more technical and expensive alternatives (e.g., electronically modulated lamp control systems). These choices should be determined according to the goals of the particular experiment.

  7. Modelos de determinação não-destrutiva da área foliar em girassol Models for estimating leaf area in sunflower

    Directory of Open Access Journals (Sweden)

    Ivan Carlos Maldaner

    2009-08-01

    Full Text Available Os objetivos deste trabalho foram obter e testar modelos matemáticos de estimativa da área do limbo foliar em função das suas dimensões lineares para o girassol. Foram conduzidos dois experimentos na área experimental do departamento de Fitotecnia da Universidade Federal de Santa Maria. As plantas de girassol foram coletadas a partir dos 27 dias após emergência (DAE. A área foliar (AF foi determinada pelo método dos discos. Ajustaram-se modelos lineares, quadráticos, cúbicos e potenciais entre área foliar e comprimento ou largura e seus produtos (comprimento*largura, sendo eliminados os que apresentaram coeficiente de determinação menor do que 0,90. A estatística utilizada para avaliar o desempenho dos modelos foi a raiz do quadrado médio do erro (RQME. Os modelos que melhor se ajustaram aos dados foram: potência, quadrático e cúbico, considerando a largura como variável independente. A área foliar de girassol pode ser estimada com o modelo potência, por ser o mais preciso, e a largura da folha.The objective of this study was to obtain and to numerical models to estimate the leaf area in function leaves linear dimension in sunflower. Two experiments were conducted at the experimental area of the Plant Science Department of the Federal University of Santa Maria, Santa Maria, RS, Brazil. Plants of sunflower were collected starting 27 days after emergency (DAE. The disks method was used to determine the leaf area (LA. Leaves were dried in oven at 65°C until constant weight. Linear, quadratic, cubic and power models between leaf area and length or width, and the product (length * width, were fitted. Models that apresented coefficient of determination lower than 0.90 were not selected. The statistic used to evaluate the performance of the models was the root mean square error (RQME. Models that had the best fit were power, quadratic and cubic using blade width as the independent variable. Leaf area in sunflower can be

  8. Brain structure mediates the association between height and cognitive ability.

    Science.gov (United States)

    Vuoksimaa, Eero; Panizzon, Matthew S; Franz, Carol E; Fennema-Notestine, Christine; Hagler, Donald J; Lyons, Michael J; Dale, Anders M; Kremen, William S

    2018-05-11

    Height and general cognitive ability are positively associated, but the underlying mechanisms of this relationship are not well understood. Both height and general cognitive ability are positively associated with brain size. Still, the neural substrate of the height-cognitive ability association is unclear. We used a sample of 515 middle-aged male twins with structural magnetic resonance imaging data to investigate whether the association between height and cognitive ability is mediated by cortical size. In addition to cortical volume, we used genetically, ontogenetically and phylogenetically distinct cortical metrics of total cortical surface area and mean cortical thickness. Height was positively associated with general cognitive ability and total cortical volume and cortical surface area, but not with mean cortical thickness. Mediation models indicated that the well-replicated height-general cognitive ability association is accounted for by individual differences in total cortical volume and cortical surface area (highly heritable metrics related to global brain size), and that the genetic association between cortical surface area and general cognitive ability underlies the phenotypic height-general cognitive ability relationship.

  9. A finger leaf design for dual layer MLCs

    International Nuclear Information System (INIS)

    Cui Weijie; Dai Jianrong

    2010-01-01

    Objective: To introduce a finger leaf design that is applied to dual layer MLCs. Methods: An optimization model was firstly constructed to describe the problem of determining leaf end shapes,and the corresponding problems were then solved by the simplex search method or the simulated annealing technique. Optimal parameters for arc shapes of leaf end projections were obtained, and a comparison was done between optimized MLCs and conventional MLCs in terms of field conformity. The optimization process was based on 634 target fields selected from the patient data base of a treatment planning system. Areas of these fields ranged from 20.0 to 602.7 cm with a mean and its standard deviation of (125.7 ± 0.0) cm 2 . Results: The optimized leaf end shapes projected to the isocenter plane were semicircles. With the finger leaf design, the total area of discrepancy regions between MLC fields and target fields was reduced by 32.3%. Conclusions: The finger leaf design improves the conformity of the MLC shaped fields to the desired target fields. (authors)

  10. Comparison of different ground techniques to map leaf area index of Norway spruce forest canopy

    OpenAIRE

    Homolova, L.; Malenovsky, Z.; Hanus, J.; Tomaskova, I.; Dvoráková, M.; Pokorny, R.

    2007-01-01

    The leaf area index (LAI) of three monocultures of Norway spruce (Picea abies (L.) Karst), different in age and structure, was measured by means of two indirect optical techniques of LAI field mapping: 1/ plant canopy analyser LAI-2000, and 2/ digital hemispherical photographs (DHP). The supportive measurements with the TRAC instrument were conducted to produce mainly the element clumping index. The aim of the study was to compare the performances of LAI-2000 and DHP and to evaluate effect of...

  11. A Comparison of Leaf Area Index Maps Derived from Multi-Sensor Optical Data Acquired over Agricultural Areas

    Directory of Open Access Journals (Sweden)

    Giuseppe Satalino

    2010-06-01

    Full Text Available The objective of this study is to retrieve and compare Leaf Area Index (LAI maps from temporal series of SPOT, IKONOS and MERIS images acquired, from 2006 to 2008, over an agricultural site in Southern Italy. Results show that the root mean square error (RMSE of LAI derived from MERIS data is approximately 1 m2 m-2, slightly larger than the one obtained by using SPOT and IKONOS data. In addition, LAI retrieved from MERIS data tends to underestimate LAI retrieved from SPOT and IKONOS data, particularly at low LAI values. Nevertheless, the paper gives examples highlighting the strength of MERIS with respect to SPOT and IKONOS data in providing long and dense temporal series of LAI maps suitable to feature the temporal evolution of vegetation growth at regional scale.

  12. What is the relationship between changes in canopy leaf area and changes in photosynthetic CO² flux in artic ecosystems?

    NARCIS (Netherlands)

    Street, L.E.; Shaver, G.R.; Williams, M.; Wijk, van M.T.

    2007-01-01

    1 The arctic environment is highly heterogeneous in terms of plant distribution and productivity. If we are to make regional scale predictions of carbon exchange it is necessary to find robust relationships that can simplify this variability. One such potential relationship is that of leaf area to

  13. The leaf litter ant fauna of an Atlantic Forest area in the Cantareira State Park – São Paulo, Brazil

    Directory of Open Access Journals (Sweden)

    Andre Soliva Ribeiro

    2005-11-01

    Full Text Available The present work surveys the leaf litter ant fauna of an Atlantic Forest area in Cantareira State Park – SP, Brazil as a complement to the project “Richness and diversity of Hymenoptera and Isoptera along a latitudinal gradient in the Atlantic Forest – the eastern Brazilian rain forest” that forms part of the BIOTA-FAPESP program. The general protocol of the project was to collect 50 leaf litter samples of 1 m2 which were then sifted and submitted to Winkler extractors for 48 hours. Sixty-two species of 25 genera in eight ant subfamilies were collected. Myrmicinae was the richest with 39 species, followed by Ponerinae (14, Ectatomminae, Heteroponerinae and Formicinae (two species each, Amblyoponinae, Proceratiinae and Dolichoderinae (one species each. The richest genera were Solenopsis and Hypoponera (12 morph-species each, and Pheidole (eight. Richness estimators indicated that the total number of species in the area should be between 68 and 85, in a confidence interval of 95%. In comparison, other locations of the evergreen Atlantic Forest have shown a significantly higher richness. Our hypothesis is that the proximity of regions of great urban concentration, allied to the factors that act on a local scale, modifies the structure of the local community of leaf litter ants.

  14. Leaf area and foliar biomass relationships in northern hardwood forests located along an 800 km acid deposition gradient

    International Nuclear Information System (INIS)

    Burton, A.J.; Pregitzer, K.S.; Reed, D.D.

    1991-01-01

    The canopies of northern hardwood forests dominated by sugar maple (Acer saccharum Marsh.) were examined at five locations spanning 800 km along an acid deposition and climatic gradient in the Great Lakes region. Leaf area index (LAI) calculated from litterfall ranged from 6.0 to 8.0 in 1988, from 4.9 to 7.9 in 1989, and from 5.3 to 7.8 in 1990. The data suggest that maximum LAI for the sites is between 7 and 8. Insect defoliation and the allocation of assimilates to reproductive parts in large seed years reduced LAI by up to 34%. Allometric equations for leaf area and foliar biomass were not significantly different among sites. They predicted higher LAI values than were estimated from litterfall and could not account for the influences of defoliation and seed production. Canopy transmittance was a viable alternative for estimating LAI. Extinction coefficients (K) of 0.49 to 0.65 were appropriate for solar elevations of 63 degree to 41 degree. Patterns of specific leaf area (SLA) were similar for the sites. Average sugar maple SLA increased from 147 cm 2 g -1 in the upper 5 m of the canopy to 389 cm 2 g -1 in the seeding layer. Litterfall SLA averaged 196 cm 2 g -1 for all species and 192 cm 2 g -1 for sugar maple. Similarity among the sites in allometric relationships, maximum LAI, canopy transmittance, and patterns of SLA suggests these characteristics were controlled primarily by the similar nutrient and moisture availability at the sites. A general increasing trend in litter production along the gradient could not be attributed to N deposition or length of growing season due to year to year variability resulting from insect defoliation and seed production

  15. Simulation of ICESat-2 canopy height retrievals for different ecosystems

    Science.gov (United States)

    Neuenschwander, A. L.

    2016-12-01

    Slated for launch in late 2017 (or early 2018), the ICESat-2 satellite will provide a global distribution of geodetic measurements from a space-based laser altimeter of both the terrain surface and relative canopy heights which will provide a significant benefit to society through a variety of applications ranging from improved global digital terrain models to producing distribution of above ground vegetation structure. The ATLAS instrument designed for ICESat-2, will utilize a different technology than what is found on most laser mapping systems. The photon counting technology of the ATLAS instrument onboard ICESat-2 will record the arrival time associated with a single photon detection. That detection can occur anywhere within the vertical distribution of the reflected signal, that is, anywhere within the vertical distribution of the canopy. This uncertainty of where the photon will be returned from within the vegetation layer is referred to as the vertical sampling error. Preliminary simulation studies to estimate vertical sampling error have been conducted for several ecosystems including woodland savanna, montane conifers, temperate hardwoods, tropical forest, and boreal forest. The results from these simulations indicate that the canopy heights reported on the ATL08 data product will underestimate the top canopy height in the range of 1 - 4 m. Although simulation results indicate the ICESat-2 will underestimate top canopy height, there is, however, a strong correlation between ICESat-2 heights and relative canopy height metrics (e.g. RH75, RH90). In tropical forest, simulation results indicate the ICESat-2 height correlates strongly with RH90. Similarly, in temperate broadleaf forest, the simulated ICESat-2 heights were also strongly correlated with RH90. In boreal forest, the simulated ICESat-2 heights are strongly correlated with RH75 heights. It is hypothesized that the correlations between simulated ICESat-2 heights and canopy height metrics are a

  16. Interacting Effects of Leaf Water Potential and Biomass on Vegetation Optical Depth: Effects of LWP and Biomass on VOD

    Energy Technology Data Exchange (ETDEWEB)

    Momen, Mostafa [Department of Earth System Science, Stanford University, Stanford CA USA; Wood, Jeffrey D. [School of Natural Resources, University of Missouri, Columbia MO USA; Novick, Kimberly A. [School of Public and Environmental Affairs, Indiana University-Bloomington, Bloomington IN USA; Pangle, Robert [Department of Biology, University of New Mexico, Albuquerque NM USA; Pockman, William T. [Department of Biology, University of New Mexico, Albuquerque NM USA; McDowell, Nate G. [Pacific Northwest National Laboratory, Richland WA USA; Konings, Alexandra G. [Department of Earth System Science, Stanford University, Stanford CA USA

    2017-11-01

    Remotely sensed microwave observations of vegetation optical depth (VOD) have been widely used for examining vegetation responses to climate. Nevertheless, the relative impacts of phenological changes in leaf biomass and water stress on VOD have not been explicitly disentangled. In particular, determining whether leaf water potential (ψL) affects VOD may allow these data sets as a constraint for plant hydraulic models. Here we test the sensitivity of VOD to variations in ψL and present a conceptual framework that relates VOD to ψL and total biomass including leaves, whose dynamics are measured through leaf area index, and woody components. We used measurements of ψL from three sites across the US—a mixed deciduous forests in Indiana and Missouri and a piñon-juniper woodland in New Mexico—to validate the conceptual model. The temporal dynamics of X-band VOD were similar to those of the VOD signal estimated from the new conceptual model with observed ψL (R2 = 0.6–0.8). At the global scale, accounting for a combination of biomass and estimated ψL (based on satellite surface soil moisture data) increased correlations with VOD by ~ 15% and 30% compared to biomass and water potential, respectively. In wetter regions with denser and taller canopy heights, VOD has a higher correlation with leaf area index than with water stress and vice versa in drier regions. Our results demonstrate that variations in both phenology and ψL must be considered to accurately interpret the dynamics of VOD observations for ecological applications.

  17. Effects of age-related increases in sapwood area, leaf area, and xylem conductivity on height-related hydraulic costs in two contrasting coniferous species

    Science.gov (United States)

    Jean-Christophe Domec; Barbara Lachenbruch; Michele L. Pruyn; Rachel Spicer

    2012-01-01

    Introduction: Knowledge of vertical variation in hydraulic parameters would improve our understanding of individual trunk functioning and likely have important implications for modeling water movement to the leaves. Specifically, understanding how foliage area (Al), sapwood area (As), and hydraulic specific...

  18. Effects of leaf movement on leaf temperature, transpiration and radiation interception in soybean under water stress conditions

    International Nuclear Information System (INIS)

    Isoda, A.; Wang, P.

    2001-01-01

    Varietal differences in leaf movement were examined in terms of radiation interception, leaf temperature and transpiration under water stressed conditions. Five cultivars (Qindou 7232, Gaofei 16, Dongnong 87 - 138, 8285 - 8 and 8874) were grown in a concrete frame field in Xinjiang, China. Irrigation treatments (irrigation and no irrigation) were made from the flowering to the pod filling stage. A leaflet in the uppermost layer of the canopy was restrained horizontally. Leaf temperatures, transpiration rate (stem sap flow rate of the main stem per unit leaf area) and intercepted radiation of each leaflet were measured. There were greater varietal differences in leaf movement, leaf temperature and transpiration rate. Leaf temperature seemed to be adjusted by leaf movement and transpiration. The extent to which is adjusted by leaf movement and transpiration differed among the cultivars; leaf temperature was influenced mainly by leaf movement for Gaofei 16 and Dongnong 87 - 138, mainly by transpiration for Qindou 7232 and 8874, and by both for 8285 - 8. Intercepted radiation in the upper two layers of the canopy (20 cm from the uppermost) was greater in the irrigated plot, although the mean values of total leaflets of the irrigated plot were not different as compared to the non-irrigated plot. Although paraheliotropic leaf movement decreased radiation interception, it offers some possibilities for the improvement in radiation penetration within a dense canopy. Cumulated amount of transpiration during a day was compared between the restrained-leaf and the non-leaf-restrained plants in 8874. Paraheliotropic leaf movement reduced water loss by 23% in the irrigated and 71% in the non-irrigated plots

  19. Effects of some growth regulating applications on leaf yield, raw ...

    African Journals Online (AJOL)

    This study investigated the effects of repetitive applications of herbagreen (HG), humic acid (HA), combined foliar fertilizer (CFF) and HG+CFF performed in the Müsküle grape variety grafted on 5 BB rootstock on fresh or pickled leaf size and leaf raw cellulose content. HA application increased leaf area and leaf water ...

  20. Temporal dynamics and spatial variability in the enhancement of canopy leaf area under elevated atmospheric CO2

    Science.gov (United States)

    Heather R. McCarthy; Ram Oren; Adrien C. Finzi; David S. Ellsworth; Hyun-Seok Kim; Kurt H. Johnsen; Bonnie Millar

    2007-01-01

    Increased canopy leaf area (L) may lead to higher forest productivity and alter processes such as species dynamics and ecosystem mass and energy fluxes. Few CO2enrichment studies have been conducted in closed canopy forests and none have shown a sustained enhancement of L. We reconstructed 8 years (1996–2003) of L at Duke’s Free Air CO...

  1. Assessing urban habitat quality based on specific leaf area and stomatal characteristics of Plantago lanceolata L

    International Nuclear Information System (INIS)

    Kardel, F.; Wuyts, K.; Babanezhad, M.; Vitharana, U.W.A.; Wuytack, T.; Potters, G.; Samson, R.

    2010-01-01

    This study has evaluated urban habitat quality by studying specific leaf area (SLA) and stomatal characteristics of the common herb Plantago lanceolata L. SLA and stomatal density, pore surface and resistance were measured at 169 locations in the city of Gent (Belgium), distributed over four land use classes, i.e., sub-urban green, urban green, urban and industry. SLA and stomatal density significantly increased from sub-urban green towards more urbanised land use classes, while the reverse was observed for stomatal pore surface. Stomatal resistance increased in the urban and industrial land use class in comparison with the (sub-) urban green, but differences between land use classes were less pronounced. Spatial distribution maps for these leaf characteristics showed a high spatial variation, related to differences in habitat quality within the city. Hence, stomatal density and stomatal pore surface are assumed to be potentially good bio-indicators for urban habitat quality. - Stomatal characteristics of Plantago lanceolata can be used for biomonitoring of urban habitat quality.

  2. Leaf-cutting ant attack in initial pine plantations and growth of defoliated plants

    Directory of Open Access Journals (Sweden)

    Mariane Aparecida Nickele

    2012-07-01

    Full Text Available The objective of this work was to evaluate the natural attack by Acromyrmex crassispinus in initial Pinus taeda plantations without control measures against ants, as well as the effect of defoliation in seedlings of P. taeda. Evaluations of the attack of leaf-cutting ants on P. taeda plantations were done monthly in the first six months, then 9 and 12 months after planting. The percentages of plants that were naturally attacked by ants were registered. The effect of defoliation was evaluated by artificial defoliation, simulating the natural patterns of attack by A. crassispinus on P. taeda seedlings. The natural attack of A. crassispinus was greater during the first months after planting, being more intense in the first 30 days. Artificial defoliation indicated that there were no significant losses in diameter and height in plants with less than 75% defoliation. However, there were significant losses in diameter and height in plants with 100% defoliation, independently of the cut of the apical meristem, and also plant death. The control of leaf-cutting ants in P. taeda plantings, in which A. crassispinus is the most frequent leaf-cutting ant, should be intense only at the beginning of planting, since the most severe attacks occur during this time.

  3. Tree height-diameter allometry across the United States.

    Science.gov (United States)

    Hulshof, Catherine M; Swenson, Nathan G; Weiser, Michael D

    2015-03-01

    The relationship between tree height and diameter is fundamental in determining community and ecosystem structure as well as estimates of biomass and carbon storage. Yet our understanding of how tree allometry relates to climate and whole organismal function is limited. We used the Forest Inventory and Analysis National Program database to determine height-diameter allometries of 2,976,937 individuals of 293 tree species across the United States. The shape of the allometric relationship was determined by comparing linear and nonlinear functional forms. Mixed-effects models were used to test for allometric differences due to climate and floristic (between angiosperms and gymnosperms) and functional groups (leaf habit and shade tolerance). Tree allometry significantly differed across the United States largely because of climate. Temperature, and to some extent precipitation, in part explained tree allometric variation. The magnitude of allometric variation due to climate, however, had a phylogenetic signal. Specifically, angiosperm allometry was more sensitive to differences in temperature compared to gymnosperms. Most notably, angiosperm height was more negatively influenced by increasing temperature variability, whereas gymnosperm height was negatively influenced by decreasing precipitation and increasing altitude. There was little evidence to suggest that shade tolerance influenced tree allometry except for very shade-intolerant trees which were taller for any given diameter. Tree allometry is plastic rather than fixed and scaling parameters vary around predicted central tendencies. This allometric variation provides insight into life-history strategies, phylogenetic history, and environmental limitations at biogeographical scales.

  4. Active moss biomonitoring applied to an industrial area in Romania: variation of element contents with the height of exposure site

    International Nuclear Information System (INIS)

    Culicov, O.; Yurukova, L.; Mocanu, R.; Frontasyeva, V.; Sarbu, C.

    2007-01-01

    Neutron activation analysis and the moss-bag transplant technique were used to investigate the variation of element contents with the height of the exposure site of Sphagnum girgensohnii samples in the strongly polluted town of Baia Mare, Romania, according to a novel sampling design. Moss collected from the background area in Moscow region, Russia, was hanged in bags at 3 locations spread within an area of one square kilometre, and analyzed after 4 months of exposure. At each location a number of 15 samples were suspended at 6 different levels above the ground: 6, 12, 18, 24 and 30 m. A total of 35 elements were determined by NAA at IBR-2 reactor in Dubna, Russia. Discriminant Analysis allowed depicting the differences between the accumulation patterns of moss at different levels of exposure and at various locations. The concentration of only twenty four and twenty five elements of thirty five provided sufficient information to enable classification rules to be developed for identifying moss samples according with their height of exposure and location, respectively. The most discriminant elements have the F values (indicating the statistical significance in the discrimination between groups) descending in the following order: K, Rb, Sb, Cs, Zn, Cl for the location model and Ca, Cd, As, Sr, Ba, Cl for the height model. A good agreement have been observed between the groups obtained by applying Discriminant Analysis and the origin of the samples. A small number of elements are significant for the separation of moss-bags by height in individual locations. The parameter ?variation in concentration by height? is less significant than the parameter ? variation in concentration by location?

  5. The leaf size-twig size spectrum in evergreen broadleaved forest of ...

    African Journals Online (AJOL)

    Compared to deciduous broad-leaved species, the evergreen broad-leaved species were smaller in total leaf area for a given cross-sectional area or stem mass. This suggests that the species would support less leaf area at a given twig cross-sectional area with increasing environmental stress. And the life form can modify ...

  6. Sugarcane leaf area estimate obtained from the corrected Normalized Difference Vegetation Index (NDVI

    Directory of Open Access Journals (Sweden)

    Rodrigo Moura Pereira

    2016-06-01

    Full Text Available Large farmland areas and the knowledge on the interaction between solar radiation and vegetation canopies have increased the use of data from orbital remote sensors in sugarcane monitoring. However, the constituents of the atmosphere affect the reflectance values obtained by imaging sensors. This study aimed at improving a sugarcane Leaf Area Index (LAI estimation model, concerning the Normalized Difference Vegetation Index (NDVI subjected to atmospheric correction. The model generated by the NDVI with atmospheric correction showed the best results (R2 = 0.84; d = 0.95; MAE = 0.44; RMSE = 0.55, in relation to the other models compared. LAI estimation with this model, during the sugarcane plant cycle, reached a maximum of 4.8 at the vegetative growth phase and 2.3 at the end of the maturation phase. Thus, the use of atmospheric correction to estimate the sugarcane LAI is recommended, since this procedure increases the correlations between the LAI estimated by image and by plant parameters.

  7. BOREAS TE-9 NSA Leaf Chlorophyll Density

    Science.gov (United States)

    Hall, Forrest G. (Editor); Curd, Shelaine (Editor); Margolis, Hank; Sy, Mikailou

    2000-01-01

    The BOREAS TE-9 team collected several data sets related to chemical and photosynthetic properties of leaves in boreal forest tree species. These data were collected to help provide an explanation of potential seasonal and spatial changes of leaf pigment properties in boreal forest species at the NSA. At different dates (FFC-Winter, FFC-Thaw, IFC-1, IFC-2, and IMC-3), foliage samples were collected from the upper third of the canopy for five NSA sites (YJP, OJP, OBS, UBS, and OA) near Thompson, Manitoba. Subsamples of 100 needles for black spruce, 20 needles for jack pine, and single leaf for trembling aspen were cut into pieces and immersed in a 20-mL DMF aliquot in a Nalgene test tube. The extracted foliage materials were then oven-dried at 68 C for 48 hours and weighed. Extracted leaf dry weight was converted to a total leaf area basis to express the chlorophyll content in mg/sq cm of total leaf area. The data are provided in tabular ASCII files. The data files are available on a CD-ROM (see document number 20010000884), or from the Oak Ridge National Laboratory (ORNL) Distributed Active Archive Center (DAAC).

  8. Moisture availability constraints on the leaf area to sapwood area ratio: analysis of measurements on Australian evergreen angiosperm trees

    Science.gov (United States)

    Togashi, Henrique; Prentice, Colin; Evans, Bradley; Forrester, David; Drake, Paul; Feikema, Paul; Brooksbank, Kim; Eamus, Derek; Taylor, Daniel

    2014-05-01

    The leaf area to sapwood area ratio (LA:SA) is a key plant trait that links photosynthesis to transpiration. Pipe model theory states that the sapwood cross-sectional area of a stem or branch at any point should scale isometrically with the area of leaves distal to that point. Optimization theory further suggests that LA:SA should decrease towards drier climates. Although acclimation of LA:SA to climate has been reported within species, much less is known about the scaling of this trait with climate among species. We compiled LA:SA measurements from 184 species of Australian evergreen angiosperm trees. The pipe model was broadly confirmed, based on measurements on branches and trunks of trees from one to 27 years old. We found considerable scatter in LA:SA among species. However quantile regression showed strong (0.2

  9. Morphological and moisture availability controls of the leaf area-to-sapwood area ratio: analysis of measurements on Australian trees.

    Science.gov (United States)

    Togashi, Henrique Furstenau; Prentice, Iain Colin; Evans, Bradley John; Forrester, David Ian; Drake, Paul; Feikema, Paul; Brooksbank, Kim; Eamus, Derek; Taylor, Daniel

    2015-03-01

    The leaf area-to-sapwood area ratio (LA:SA) is a key plant trait that links photosynthesis to transpiration. The pipe model theory states that the sapwood cross-sectional area of a stem or branch at any point should scale isometrically with the area of leaves distal to that point. Optimization theory further suggests that LA:SA should decrease toward drier climates. Although acclimation of LA:SA to climate has been reported within species, much less is known about the scaling of this trait with climate among species. We compiled LA:SA measurements from 184 species of Australian evergreen angiosperm trees. The pipe model was broadly confirmed, based on measurements on branches and trunks of trees from one to 27 years old. Despite considerable scatter in LA:SA among species, quantile regression showed strong (0.2 < R1 < 0.65) positive relationships between two climatic moisture indices and the lowermost (5%) and uppermost (5-15%) quantiles of log LA:SA, suggesting that moisture availability constrains the envelope of minimum and maximum values of LA:SA typical for any given climate. Interspecific differences in plant hydraulic conductivity are probably responsible for the large scatter of values in the mid-quantile range and may be an important determinant of tree morphology.

  10. Assessing the Transferability of Statistical Predictive Models for Leaf Area Index Between Two Airborne Discrete Return LiDAR Sensor Designs Within Multiple Intensely Managed Loblolly Pine Forest Locations in the South-Eastern USA

    Science.gov (United States)

    Sumnall, Matthew; Peduzzi, Alicia; Fox, Thomas R.; Wynne, Randolph H.; Thomas, Valerie A.; Cook, Bruce

    2016-01-01

    Leaf area is an important forest structural variable which serves as the primary means of mass and energy exchange within vegetated ecosystems. The objective of the current study was to determine if leaf area index (LAI) could be estimated accurately and consistently in five intensively managed pine plantation forests using two multiple-return airborne LiDAR datasets. Field measurements of LAI were made using the LiCOR LAI2000 and LAI2200 instruments within 116 plots were established of varying size and within a variety of stand conditions (i.e. stand age, nutrient regime and stem density) in North Carolina and Virginia in 2008 and 2013. A number of common LiDAR return height and intensity distribution metrics were calculated (e.g. average return height), in addition to ten indices, with two additional variants, utilized in the surrounding literature which have been used to estimate LAI and fractional cover, were calculated from return heights and intensity, for each plot extent. Each of the indices was assessed for correlation with each other, and was used as independent variables in linear regression analysis with field LAI as the dependent variable. All LiDAR derived metrics were also entered into a forward stepwise linear regression. The results from each of the indices varied from an R2 of 0.33 (S.E. 0.87) to 0.89 (S.E. 0.36). Those indices calculated using ratios of all returns produced the strongest correlations, such as the Above and Below Ratio Index (ABRI) and Laser Penetration Index 1 (LPI1). The regression model produced from a combination of three metrics did not improve correlations greatly (R2 0.90; S.E. 0.35). The results indicate that LAI can be predicted over a range of intensively managed pine plantation forest environments accurately when using different LiDAR sensor designs. Those indices which incorporated counts of specific return numbers (e.g. first returns) or return intensity correlated poorly with field measurements. There were

  11. Impact of anatomical traits of maize (Zea mays L.) leaf as affected by nitrogen supply and leaf age on bundle sheath conductance.

    Science.gov (United States)

    Retta, Moges; Yin, Xinyou; van der Putten, Peter E L; Cantre, Denis; Berghuijs, Herman N C; Ho, Quang Tri; Verboven, Pieter; Struik, Paul C; Nicolaï, Bart M

    2016-11-01

    The mechanism of photosynthesis in C 4 crops depends on the archetypal Kranz-anatomy. To examine how the leaf anatomy, as altered by nitrogen supply and leaf age, affects the bundle sheath conductance (g bs ), maize (Zea mays L.) plants were grown under three contrasting nitrogen levels. Combined gas exchange and chlorophyll fluorescence measurements were done on fully grown leaves at two leaf ages. The measured data were analysed using a biochemical model of C 4 photosynthesis to estimate g bs . The leaf microstructure and ultrastructure were quantified using images obtained from micro-computed tomography and microscopy. There was a strong positive correlation between g bs and leaf nitrogen content (LNC) while old leaves had lower g bs than young leaves. Leaf thickness, bundle sheath cell wall thickness and surface area of bundle sheath cells per unit leaf area (S b ) correlated well with g bs although they were not significantly affected by LNC. As a result, the increase of g bs with LNC was little explained by the alteration of leaf anatomy. In contrast, the combined effect of LNC and leaf age on S b was responsible for differences in g bs between young leaves and old leaves. Future investigations should consider changes at the level of plasmodesmata and membranes along the CO 2 leakage pathway to unravel LNC and age effects further. Copyright © 2016 Elsevier Ireland Ltd. All rights reserved.

  12. In defense of the classical height system

    Science.gov (United States)

    Foroughi, Ismael; Vaníček, Petr; Sheng, Michael; Kingdon, Robert William; Santos, Marcelo C.

    2017-11-01

    In many European countries, normal heights referred to the quasi-geoid as introduced by Molodenskij in the mid-20th century are preferred to the classical height system that consists of orthometric heights and the geoid as a reference surface for these heights. The rationale for this choice is supposed to be that in the classical height system, neither the geoid, nor the orthometric height can be ever known with centimetre level accuracy because one would need to know the topographical mass density to a level that can never be achieved. The aim of this paper is to question the validity of this rationale. The common way of assessing the congruency of a local geoid model and the orthometric heights is to compare the geoid heights with the difference between orthometric heights provided by leveling and geodetic heights provided by GNSS. On the other hand, testing the congruency of a quasi-geoidal model with normal height a similar procedure is used, except that instead of orthometric heights, normal heights are employed. For the area of Auvergne, France, which is now a more or less standard choice for precise geoid or quasi-geoid testing, only the normal heights are supplied by the Institute Geographic National, the provider of the data. This is clearly the consequence of the European preference for the Molodenskij system. The quality of the height system is to be judged by the congruency of the difference of the geoid/quasi-geoid heights subtracted from the geodetic heights and orthometric/normal heights. To assess the congruency of the classical height system, the Helmert approximation of orthometric heights is typically used as the transformation between normal and Helmert's heights is easily done. However, the evaluation of the differences between Helmert's and the rigorous orthometric heights is somewhat more involved as will be seen from the review in this paper. For the area of interest, the differences between normal and Helmert's heights at the control

  13. Estimativa da área foliar de meloeiro em estádios fenológicos por fotos digitais Estimate of the leaf area of melon plant in growing stages for digital photos

    Directory of Open Access Journals (Sweden)

    Sidinei José Lopes

    2007-08-01

    Full Text Available O objetivo deste trabalho foi avaliar a precisão do método de fotos digitais na estimativa da área foliar de meloeiro e encontrar modelos matemáticos de estimativa da área foliar em função de medidas lineares da folha para diferentes estádios fenológicos. Foram fotografadas todas as folhas ímpares de 8 plantas após transplante definitivo, através de câmera fotográfica digital, resultando, durante todo o ciclo da cultura, em 4.188 fotos, das quais mediu-se a área foliar, o comprimento e a largura da folha, por meio do software Sigma Scan Pro v. 5.0, Jandel Scientific. Para verificar a precisão do método de fotos digitais, retirou-se uma amostra de 40 folhas, de onde foram obtidas a área foliar através do método padrão de discos foliares e pelo método de fotos. Foi encontrada uma correlação de 0,99 entre o método padrão (discos e o de fotos. O método de fotos digitais pode ser utilizado para estimar a área foliar da cultura de meloeiro, e a estimativa da área foliar de meloeiro por estádio fenológico apresenta maior precisão, sendo a maior variabilidade na estimativa da área da folha observada no período reprodutivo. A largura máxima da folha de meloeiro é a medida linear que melhor estima a área foliar.This experiment was aimed at evaluating the precision of digital photos in estimating the leaf area of watermelon plants and to find mathematical models that estimates leaf area as a function of leaf linear measurements at different growth stages. All odd leaves of eight plants were photographed after being established on the field using a digital camera that resulted in 4,188 photos in which length and width were measured using a Sigma Scan Pro v. 5.0 Jandel Scientific software. In order to estimate the precision of the digital photos method, a sample consisting of 40 leaves was taken and leaf area measured using the standard leaf disks and the photo method. A 0.99 correlation coefficient was detected between

  14. Leaf Area Index (LAI) in different type of agroforestry systems based on hemispherical photographs in Cidanau Watershed

    Science.gov (United States)

    Nur Khairiah, Rahmi; Setiawan, Yudi; Budi Prasetyo, Lilik; Ayu Permatasari, Prita

    2017-01-01

    Ecological functions of agroforestry systems have perceived benefit to people around Cidanau Watershed, especially in the protection of water quality. The main causes of the problems encountered in the Cidanau Watershed are associated with the human factors, especially encroachment and conversion of forest into farmland. The encroachment has made most forest in Cidanau Watershed become bare land. To preserve the ecological function of agroforestry systems in Cidanau Watershed, monitoring of the condition of the vegetation canopy in agroforestry systems is really needed. High intensity thinning of crown density due to deforestation can change stand leaf area index dramatically. By knowing LAI, we can assess the condition of the vegetation canopy in agroforestry systems. LAI in this research was obtained from Hemispherical Photographs analysis using the threshold method in HemiView Canopy Analysis Software. Our research results indicate that there are six types of agroforestry in Cidanau Watershed i.e. Sengon Agroforestry, Clove Agroforestry, Melinjo Agroforestry, Chocolate Agroforestry, Coffee Agroforestry, and Complex Agroforestry. Several factors potentially contribute to variations in the value of LAI in different types of agroforestry. The simple assumptions about differences ranges of LAI values on six types of agroforestry is closely related to leaf area and plant population density.

  15. Modelo para determinção da área foliar de Kalanchoe blossfeldiana Poelln Model for leaf area determination in Kalanchoe blossfeldiana Poelln

    Directory of Open Access Journals (Sweden)

    Marcia Xavier Peiter

    2006-12-01

    Full Text Available O presente trabalho teve por objetivo a verificação de um procedimento matemático que permita a descrição do crescimento foliar de Kalanchoe (Kalanchoe blossfeldiana Poelln. e possa prever a sua área foliar fotossinteticamente ativa a partir de medidas não destrutivas de folhas. As mudas de Kalanchoe Cv. "Gold Jewel" foram cultivadas para o procedimento experimental em vasos irrigados com doses recomendadas para a cultura. Semanalmente, foram retirados três vasos da estufa e as plantas tiveram suas folhas cortadas, identificadas e submetidas a tomadas de medidas de sua posição na planta, do máximo comprimento longitudinal e do máximo comprimento transversal. Foram realizadas um total de nove coletas semanalmente, desde 04/04/2003 até o início da floração. Em cada coleta, três plantas eram amostradas e a área foliar calculada com a utilização do método de Gauss (GARCIA & PIEDADE, 1944 implementado em Visual Basic especificamente para este objetivo. Foram amostradas um total de 979 folhas e a verificação da possibilidade de uso de um fator de correção médio (FCM para o cálculo da área de uma folha, independentemente de sua posição na planta ou fase do ciclo de crescimento, foi averiguada por análise de regressão entre os valores obtidos pelo método padrão (Gauss e os valores estimados pelo método do FCM. Os resultados experimentais indicam que o valor FCM=1,1134 pode ser utilizado para estimar a área foliar pela multiplicação pelos valores de comprimento e largura de folha em qualquer fase do cultivo e sem qualquer posição da folha na planta.This research was aimed at versifying a mathematical procedure that allows the description of leaf of Kalanchoe (Kalanchoe blossfeldiana Poelln. and the estimation of its photosynthetically active leaf area starting from a non destructive leaf determination. Seedlings of Kalanchoe cv Gold Jewel were cultivated in irrigated vases with recommended doses for the culture

  16. Benchmarking sensitivity of biophysical processes to leaf area changes in land surface models

    Science.gov (United States)

    Forzieri, Giovanni; Duveiller, Gregory; Georgievski, Goran; Li, Wei; Robestson, Eddy; Kautz, Markus; Lawrence, Peter; Ciais, Philippe; Pongratz, Julia; Sitch, Stephen; Wiltshire, Andy; Arneth, Almut; Cescatti, Alessandro

    2017-04-01

    Land surface models (LSM) are widely applied as supporting tools for policy-relevant assessment of climate change and its impact on terrestrial ecosystems, yet knowledge of their performance skills in representing the sensitivity of biophysical processes to changes in vegetation density is still limited. This is particularly relevant in light of the substantial impacts on regional climate associated with the changes in leaf area index (LAI) following the observed global greening. Benchmarking LSMs on the sensitivity of the simulated processes to vegetation density is essential to reduce their uncertainty and improve the representation of these effects. Here we present a novel benchmark system to assess model capacity in reproducing land surface-atmosphere energy exchanges modulated by vegetation density. Through a collaborative effort of different modeling groups, a consistent set of land surface energy fluxes and LAI dynamics has been generated from multiple LSMs, including JSBACH, JULES, ORCHIDEE, CLM4.5 and LPJ-GUESS. Relationships of interannual variations of modeled surface fluxes to LAI changes have been analyzed at global scale across different climatological gradients and compared with satellite-based products. A set of scoring metrics has been used to assess the overall model performances and a detailed analysis in the climate space has been provided to diagnose possible model errors associated to background conditions. Results have enabled us to identify model-specific strengths and deficiencies. An overall best performing model does not emerge from the analyses. However, the comparison with other models that work better under certain metrics and conditions indicates that improvements are expected to be potentially achievable. A general amplification of the biophysical processes mediated by vegetation is found across the different land surface schemes. Grasslands are characterized by an underestimated year-to-year variability of LAI in cold climates

  17. Substrates with green manure compost and leaf application of biofertilizer on seedlings of yellow passion fruit plants

    Directory of Open Access Journals (Sweden)

    Cristiane Muniz Barbosa Barros

    2013-12-01

    Full Text Available Substrates and fertilization are fundamental for seedling production, which well nourished can produce earlier and are more resistant to stresses. Animal manures are often used in non-industrialized substrates with good results, but their costs are increasing. Other residues may be used for plant nutrition, in substrates or in leaf fertilization. The aim of this work was to evaluate substrates prepared with green manure composts and the leaf application of biofertilizer on the formation of yellow passion fruit seedlings. A greenhouse experiment was conducted between December 2009 and February 2010, with a split-plot random block design. Plots received or not leaf application of supermagro biofertilizer. Subplots consisted of different substrates: soil; soil + cattle manure; soil + cattle manure composted with black oats straw; soil + cattle manure composted with ryegrass straw; soil + cattle manure composted with turnip straw; and soil + cattle manure composted with vetch straw. There were three dates of leaf fertilization: 10, 25 and 40 days after emergence (DAE. At 50 DAE plants were collected for evaluation of growth and accumulation of biomass and nutrients: N, P, K, Ca, Mg, Cu, Mn and Zn. Data were submitted to analysis of variance and means compared by Tukey test. The substrate soil + cattle manure promoted higher stem diameter, plant height, leaf area, root length and volume and nutrient accumulation. Among substrates with green manure composts, those prepared with black oats and turnip straw outranked the others. The use of leaf biofertilizer showed diverse results on seedling formation, being beneficial when combined to substrates with black oats composted straw, and prejudicial when combined to soil + cattle manure and soil + turnip composted straw substrates. The accumulation of nutrients by the seedlings occurred in the following order: K>Ca>N>Mg>P>Zn>Cu=Mn.

  18. The plant economics spectrum is structured by leaf habits and growth forms across subtropical species.

    Science.gov (United States)

    Zhao, Yan-Tao; Ali, Arshad; Yan, En-Rong

    2017-02-01

    The plant economics spectrum that integrates the combination of leaf and wood syndromes provides a useful framework for the examination of species strategies at the whole-plant level. However, it remains unclear how species that differ in leaf habits and growth forms are integrated within the plant economics spectrum in subtropical forests. We measured five leaf and six wood traits across 58 subtropical plant species, which represented two leaf habits (evergreen vs deciduous) and two growth forms (tree vs shrub) in eastern China. Principal component analysis (PCA) was employed separately to construct the leaf (LES), wood (WES) and whole-plant (WPES) economics spectra. Leaf and wood traits are highly intra- and intercorrelated, thus defining not only the LES and WES, but also a WPES. Multi-trait variations in PCAs revealed that the traits which were representative of the acquisitive strategy, i.e., cheap tissue investment and rapid returns on that investment, were clustered at one end, while traits that represented the conservative strategy, i.e., expensive tissue investment and slower returns, were clustered at other end in each of the axes of the leaf and wood syndromes (PC1-axis) and the plant height strategy (PC2-axis). The local WPES, LES and WES were tightly correlated with each other. Evergreens shaped the conservative side, while deciduous species structured the acquisitive side of the WPES and LES. With respect to plant height strategies, trees formulated the acquisitive side and shrub species made up the conservative side of the WPES, LES and WES. In conclusion, our results suggested that the LES and WES were coordinated to a WPES for subtropical species. The finding of this local spectrum of plant form and function would be beneficial for modeling nutrient fluxes and species compositions in the changing climate, but also for understanding species strategies in an evolutionary context. © The Author 2016. Published by Oxford University Press. All rights

  19. Nota científica: métodos para estimativa da area foliar de plantas daninhas: 2: Wissadula subpeltata (Kuntze Fries Methods for estimation of leaf area of weeds: 2: Wissadula subpeltata (Kuntze Fries

    Directory of Open Access Journals (Sweden)

    S. Bianco

    1983-06-01

    Full Text Available Com o objetivo de obter uma equação que, através de parâmetros lineares dimensionais das folhas, permitisse estimar a área foliar de Wissadula subpeltata (Kuntze Fries, estudaram- se correlações entre a área foliar real e o comprimento da folha ao longo da nervura principal (C , largura máxi ma da folha (L , comprimento do espaço entre o ponto de inserção do pecíolo na folha até a primeira ramificação da nervura principal (CE, L + C, L x C e L x CE. Todas as equações, geométricas ou lineares simples, permitiram boas estimativas da área foliar . Do pont o de vista prático, sugere- se optar pela equação linear simples envolvendo o produto C x L, considerando o coeficiente linear igual a zero. Deste modo, a estimativa da área foliar de W. subpeltata pode ser feita pel a fórmula Y = 0, 85 49 (C x L, ou seja 85 ,49% do produto entre o comprimento da nervura principal e a largura máxima da folha.In order to final an equation that make poss ible to estimate the leaf area of Wissadula usbpeltata (Kuntze Fries , were studied correlations between truelea far ea (Y and the lea flenght in the mid rib direct ion (C, maximum leaf width (L , lenght of the segment between the petiole insert ion point in the leaf and the first rami fication of leaf mid rib (CE, L + C, L x C and L x CE. Al l equations, geometric and simple linear, permited good lea fare a estimatives. It is suggested to decid e for simple linear equations envolving the C x L, considering zero the linear coefficient. Thus , the leaf area of W. subpeltata can be estimated by the equation Y = 0.8549 (C x L, or else 85,49% of the multiplication between the leaf lenght in the mid rib direction and the maximum leaf width.

  20. From leaf to whole-plant water use efficiency (WUE in complex canopies: Limitations of leaf WUE as a selection target

    Directory of Open Access Journals (Sweden)

    Hipólito Medrano

    2015-06-01

    Full Text Available Plant water use efficiency (WUE is becoming a key issue in semiarid areas, where crop production relies on the use of large volumes of water. Improving WUE is necessary for securing environmental sustainability of food production in these areas. Given that climate change predictions include increases in temperature and drought in semiarid regions, improving crop WUE is mandatory for global food production. WUE is commonly measured at the leaf level, because portable equipment for measuring leaf gas exchange rates facilitates the simultaneous measurement of photosynthesis and transpiration. However, when those measurements are compared with daily integrals or whole-plant estimates of WUE, the two sometimes do not agree. Scaling up from single-leaf to whole-plant WUE was tested in grapevines in different experiments by comparison of daily integrals of instantaneous water use efficiency [ratio between CO2 assimilation (AN and transpiration (E; AN/E] with midday AN/E measurements, showing a low correlation, being worse with increasing water stress. We sought to evaluate the importance of spatial and temporal variation in carbon and water balances at the leaf and plant levels. The leaf position (governing average light interception in the canopy showed a marked effect on instantaneous and daily integrals of leaf WUE. Night transpiration and respiration rates were also evaluated, as well as respiration contributions to total carbon balance. Two main components were identified as filling the gap between leaf and whole plant WUE: the large effect of leaf position on daily carbon gain and water loss and the large flux of carbon losses by dark respiration. These results show that WUE evaluation among genotypes or treatments needs to be revised.

  1. Leaf size indices and structure of the peat swamp forest

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    L.G. Aribal

    2017-12-01

    Full Text Available Leaf size indices of the tree species in the peatland of Agusan del Sur in Mindanao in Philippines was examined to deduce the variation of forest structure and observed forest zonation.  Using raunkiaer and webb’s leaf size classification, the leaf morphometrics of seven tree species consistently found on the established sampling plots were determined.  The species includes Ternstroemia philippinensis Merr., Polyscias aherniana Merr. Lowry and G.M. Plunkett, Calophyllum sclerophyllum Vesque, Fagraea racemosa Jack, Ilex cymosa Blume, Syzygium tenuirame (Miq. Merr. and Tristaniopsis micrantha Merr. Peter G.Wilson and J.T.Waterh.The LSI were correlated against the variables of the peat physico-chemical properties (such as bulk density, acrotelm thickness, peat depth, total organic carbon, nitrogen, phosphorus, and potassium, pH; water (pH, ammonium, nitrate, phosphate; and leaf tissue elements (nitrogen, phosphorus and potassium.  Result showed a decreasing leaf size indices and a three leaf size category consisting of mesophyllous, mesophyllous-notophyllous and microphyllous were observed which corresponds to the structure of vegetation i.e., from the tall-pole forest having the biggest average leaf area of 6,142.29 mm2 to the pygmy forest with average leaf area of 1,670.10 mm2.  Such decreased leaf size indices were strongly correlated to soil nitrogen, acrotelm thickness, peat depth, phosphate in water, nitrogen and phosphorus in the plant tissue.

  2. Optimal leaf sequencing with elimination of tongue-and-groove underdosage

    Energy Technology Data Exchange (ETDEWEB)

    Kamath, Srijit [Department of Computer and Information Science and Engineering, University of Florida, Gainesville, FL (United States); Sahni, Sartaj [Department of Computer and Information Science and Engineering, University of Florida, Gainesville, FL (United States); Palta, Jatinder [Department of Radiation Oncology, University of Florida, Gainesville, FL (United States); Ranka, Sanjay [Department of Computer and Information Science and Engineering, University of Florida, Gainesville, FL (United States); Li, Jonathan [Department of Radiation Oncology, University of Florida, Gainesville, FL (United States)

    2004-02-07

    The individual leaves of a multileaf collimator (MLC) have a tongue-and-groove or stepped-edge design to minimize leakage radiation between adjacent leaves. This design element has a drawback in that it creates areas of underdosages in intensity-modulated photon beams unless a leaf trajectory is specifically designed such that for any two adjacent leaf pairs, the direct exposure under the tongue-and-groove is equal to the lower of the direct exposures of the leaf pairs. In this work, we present a systematic study of the optimization of a leaf sequencing algorithm for segmental multileaf collimator beam delivery that completely eliminates areas of underdosages due to tongue-and-groove or stepped-edge design of the MLC. Simultaneous elimination of tongue-and-groove effect and leaf interdigitation is also studied. This is an extension of our previous work (Kamath et al 2003a Phys. Med. Biol. 48 307) in which we described a leaf sequencing algorithm that is optimal for monitor unit (MU) efficiency under most common leaf movement constraints that include minimum leaf separation. Compared to our previously published algorithm (without constraints), the new algorithms increase the number of sub-fields by approximately 21% and 25%, respectively, but are optimal in MU efficiency for unidirectional schedules. (note)

  3. Optimal leaf sequencing with elimination of tongue-and-groove underdosage

    International Nuclear Information System (INIS)

    Kamath, Srijit; Sahni, Sartaj; Palta, Jatinder; Ranka, Sanjay; Li, Jonathan

    2004-01-01

    The individual leaves of a multileaf collimator (MLC) have a tongue-and-groove or stepped-edge design to minimize leakage radiation between adjacent leaves. This design element has a drawback in that it creates areas of underdosages in intensity-modulated photon beams unless a leaf trajectory is specifically designed such that for any two adjacent leaf pairs, the direct exposure under the tongue-and-groove is equal to the lower of the direct exposures of the leaf pairs. In this work, we present a systematic study of the optimization of a leaf sequencing algorithm for segmental multileaf collimator beam delivery that completely eliminates areas of underdosages due to tongue-and-groove or stepped-edge design of the MLC. Simultaneous elimination of tongue-and-groove effect and leaf interdigitation is also studied. This is an extension of our previous work (Kamath et al 2003a Phys. Med. Biol. 48 307) in which we described a leaf sequencing algorithm that is optimal for monitor unit (MU) efficiency under most common leaf movement constraints that include minimum leaf separation. Compared to our previously published algorithm (without constraints), the new algorithms increase the number of sub-fields by approximately 21% and 25%, respectively, but are optimal in MU efficiency for unidirectional schedules. (note)

  4. Comparison of leaf-on and leaf-off ALS data for mapping riparian tree species

    Science.gov (United States)

    Laslier, Marianne; Ba, Antoine; Hubert-Moy, Laurence; Dufour, Simon

    2017-10-01

    Forest species composition is a fundamental indicator of forest study and management. However, describing forest species composition at large scales and of highly diverse populations remains an issue for which remote sensing can provide significant contribution, in particular, Airborne Laser Scanning (ALS) data. Riparian corridors are good examples of highly valuable ecosystems, with high species richness and large surface areas that can be time consuming and expensive to monitor with in situ measurements. Remote sensing could be useful to study them, but few studies have focused on monitoring riparian tree species using ALS data. This study aimed to determine which metrics derived from ALS data are best suited to identify and map riparian tree species. We acquired very high density leaf-on and leaf-off ALS data along the Sélune River (France). In addition, we inventoried eight main riparian deciduous tree species along the study site. After manual segmentation of the inventoried trees, we extracted 68 morphological and structural metrics from both leaf-on and leaf-off ALS point clouds. Some of these metrics were then selected using Sequential Forward Selection (SFS) algorithm. Support Vector Machine (SVM) classification results showed good accuracy with 7 metrics (0.77). Both leaf-on and leafoff metrics were kept as important metrics for distinguishing tree species. Results demonstrate the ability of 3D information derived from high density ALS data to identify riparian tree species using external and internal structural metrics. They also highlight the complementarity of leaf-on and leaf-off Lidar data for distinguishing riparian tree species.

  5. Algorithm for retrieving vegetative canopy and leaf parameters from multi- and hyperspectral imagery

    Science.gov (United States)

    Borel, Christoph

    2009-05-01

    In recent years hyper-spectral data has been used to retrieve information about vegetative canopies such as leaf area index and canopy water content. For the environmental scientist these two parameters are valuable, but there is potentially more information to be gained as high spatial resolution data becomes available. We developed an Amoeba (Nelder-Mead or Simplex) based program to invert a vegetative canopy radiosity model coupled with a leaf (PROSPECT5) reflectance model and modeled for the background reflectance (e.g. soil, water, leaf litter) to a measured reflectance spectrum. The PROSPECT5 leaf model has five parameters: leaf structure parameter Nstru, chlorophyll a+b concentration Cab, carotenoids content Car, equivalent water thickness Cw and dry matter content Cm. The canopy model has two parameters: total leaf area index (LAI) and number of layers. The background reflectance model is either a single reflectance spectrum from a spectral library() derived from a bare area pixel on an image or a linear mixture of soil spectra. We summarize the radiosity model of a layered canopy and give references to the leaf/needle models. The method is then tested on simulated and measured data. We investigate the uniqueness, limitations and accuracy of the retrieved parameters on canopy parameters (low, medium and high leaf area index) spectral resolution (32 to 211 band hyperspectral), sensor noise and initial conditions.

  6. Simulation of leaf area index on site scale based on model data fusion

    Science.gov (United States)

    Yang, Y.; Wang, J. B.

    2017-12-01

    The world's grassland area is about 24 × 108hm2, accounting for about one-fifth of the global land area. It is one of the most widely distributed terrestrial ecosystems on Earth. And currently, it is the most affected area of human activity. A considerable portion of the global CO2 emissions are fixed by grassland, and the grassland carbon cycle plays an important role in the global carbon cycle (Li Bo, Yongshen Peng, Li Yao, China's Prairie, 1990). In recent years, the carbon cycle and its influencing factors of grassland ecosystems have become one of the hotspots in ecology, geology, botany and agronomy under the background of global change ( Mu Shaojie, 2014) . And the model is now as a popular and effective method of research. However, there are still some uncertainties in this approach. CEVSA ( Carbon Exchange between Vegetation, Soil and Atmosphere) is a biogeochemical cycle model based on physiological and ecological processes to simulate plant-soil-atmosphere system energy exchange and water-carbon-nitrogen coupling cycles (Cao at al., 1998a; 1998b; Woodward et al., 1995). In this paper, the remote sensing observation data of leaf area index are integrated into the model, and the CEVSA model of site version is optimized by Markov chain-Monte Carlo method to achieve the purpose of increasing the accuracy of model results.

  7. Leaf temperature and stomatal influences on sap velocity diurnal hysteresis in the Amazon rainforest

    Science.gov (United States)

    Jardine, K.; Gimenez, B.; Negron Juarez, R. I.; Koven, C.; Powell, T.; Higuchi, N.; Chambers, J.; Varadharajan, C.

    2016-12-01

    In order to improve our ability to predict terrestrial evapotranspiration fluxes, an understanding of the interactions between plant physiology and environmental conditions is necessary, but remains poorly characterized, especially in tropical ecosystems. In this study we show a tight positive correlation between sap velocity (at 1 m of height) and leaf surface temperature (LST, 20-30 m of height) in canopy dominant trees in two primary rainforest sites in the Amazon basin (Santarém and Manaus, Brazil). As leaf temperatures varied throughout the day, sap velocity responded with little delay (<15 min). Positive sap velocity was often observed at night, but also closely followed night time LSTs. When plotted versus LST, sap velocity showed an exponential increase before reaching a reflection point and a plateau and is characterized as a sigmoidal curve, in all observed trees. Moreover, a clear diurnal hysteresis in sap velocity was evident with morning periods showing higher temperature sensitivities than afternoon and night periods. Diurnal leaf observations showed a morning peak in stomatal conductance ( 10:00-10:30), but a mid-day to afternoon peak in transpiration and leaf temperature (12:00-14:00). Our observations suggest the sap velocity-LST hysteresis pattern arises due to the temporal offset between stomatal conductance and vapor pressure deficits (VPD) and demonstrates the dominating effect of VPD over stomatal conductance in maintaining high transpiration/sap flow rates under elevated temperatures. Our results have important implications for modeling tropical forest transpiration and suggests the possibility of predicting evapotranspiration fluxes at the ecosystem to regional scales based on remote sensed vegetation temperature.

  8. Patterns of leaf morphology and leaf N content in relation to winter temperatures in three evergreen tree species

    Science.gov (United States)

    Mediavilla, Sonia; Gallardo-López, Victoria; González-Zurdo, Patricia; Escudero, Alfonso

    2012-09-01

    The competitive equilibrium between deciduous and perennial species in a new scenario of climate change may depend closely on the productivity of leaves along the different seasons of the year and on the morphological and chemical adaptations required for leaf survival during the different seasons. The aim of the present work was to analyze such adaptations in the leaves of three evergreen species ( Quercus ilex, Q. suber and Pinus pinaster) and their responses to between-site differences in the intensity of winter harshness. We explore the hypothesis that the harshness of winter would contribute to enhancing the leaf traits that allow them to persist under conditions of stress. The results revealed that as winter harshness increases a decrease in leaf size occurs in all three species, together with an increase in the content of nitrogen per unit leaf area and a greater leaf mass per unit area, which seems to be achieved only through increased thickness, with no associated changes in density. P. pinaster was the species with the most intense response to the harshening of winter conditions, undergoing a more marked thickening of its needles than the two Quercus species. Our findings thus suggest that lower winter temperatures involve an increase in the cost of leaf production of evergreen species, which must be taken into account in the estimation of the final cost and benefit balance of evergreens. Such cost increases would be more pronounced for those species that, like P. pinaster, show a stronger response to the winter cold.

  9. Sexual Orientation, Objective Height, and Self-Reported Height.

    Science.gov (United States)

    Skorska, Malvina N; Bogaert, Anthony F

    2017-01-01

    Studies that have used mostly self-reported height have found that androphilic men and women are shorter than gynephilic men and women, respectively. This study examined whether an objective height difference exists or whether a psychosocial account (e.g., distortion of self-reports) may explain these putative height differences. A total of 863 participants, recruited at a Canadian university, the surrounding region, and through lesbian, gay, bisexual, and transgender (LGBT) events across Canada, self-reported their height and had their height measured. Androphilic men were shorter, on average, than gynephilic men. There was no objective height difference between gynephilic, ambiphilic, and androphilic women. Self-reported height, statistically controlling for objective height, was not related to sexual orientation. These findings are the first to show an objective height difference between androphilic and gynephilic men. Also, the findings suggest that previous studies using self-reported height found part of a true objective height difference between androphilic and gynephilic men. These findings have implications for existing biological theories of men's sexual orientation development.

  10. Leaf cuticle variations in amaranthus spinousus as indicators of environmental pollution

    International Nuclear Information System (INIS)

    Otoide, J.E.; Kayode, J.

    2007-01-01

    Investigation of the leaf epidermal characteristics of Amaranthus spinosus from polluted and non-polluted populations revealed that the stomatal pores of the leaves of the plants of the polluted areas were closed whereas those of the non-polluted areas were open. Mean length x mean width of stomatal pores on the upper leaf surface were 0.86 micro x 0.43 micro and 1.23 micro x 0.45 micro on the lower leaf surface of the non polluted microhabitats. Also, the leaves of the polluted population were smaller than those of the non-polluted population. The average leaf area of the plants of the Polluted population was 7.64 cm/sub -2/ against 12.13 cm/sub 2/ of the plants of the non-polluted areas. The results were attributed to the combined effects of air pollutant that predominated roadsides from where the samples were taken. Thus it is inferred that this plant could serve as bio-indicator of air pollution. (author)

  11. Examining variation in the leaf mass per area of dominant species across two contrasting tropical gradients in light of community assembly

    NARCIS (Netherlands)

    Neyret, Margot; Bentley, Lisa Patrick; Oliveras Menor, Imma; Marimon, Beatriz S.; Marimon-Junior, Ben Hur; Almeida de Oliveira, Edmar; Barbosa Passos, Fábio; Castro Ccoscco, Rosa; Santos, dos Josias; Matias Reis, Simone; Morandi, Paulo S.; Rayme Paucar, Gloria; Robles Cáceres, Arturo; Valdez Tejeira, Yolvi; Yllanes Choque, Yovana; Salinas, Norma; Shenkin, Alexander; Asner, Gregory P.; Díaz, Sandra; Enquist, Brian J.; Malhi, Yadvinder

    2016-01-01

    Understanding variation in key functional traits across gradients in high diversity systems and the ecology of community changes along gradients in these systems is crucial in light of conservation and climate change. We examined inter- and intraspecific variation in leaf mass per area (LMA) of

  12. Modeling canopy-level productivity: is the "big-leaf" simplification acceptable?

    Science.gov (United States)

    Sprintsin, M.; Chen, J. M.

    2009-05-01

    The "big-leaf" approach to calculating the carbon balance of plant canopies assumes that canopy carbon fluxes have the same relative responses to the environment as any single unshaded leaf in the upper canopy. Widely used light use efficiency models are essentially simplified versions of the big-leaf model. Despite its wide acceptance, subsequent developments in the modeling of leaf photosynthesis and measurements of canopy physiology have brought into question the assumptions behind this approach showing that big leaf approximation is inadequate for simulating canopy photosynthesis because of the additional leaf internal control on carbon assimilation and because of the non-linear response of photosynthesis on leaf nitrogen and absorbed light, and changes in leaf microenvironment with canopy depth. To avoid this problem a sunlit/shaded leaf separation approach, within which the vegetation is treated as two big leaves under different illumination conditions, is gradually replacing the "big-leaf" strategy, for applications at local and regional scales. Such separation is now widely accepted as a more accurate and physiologically based approach for modeling canopy photosynthesis. Here we compare both strategies for Gross Primary Production (GPP) modeling using the Boreal Ecosystem Productivity Simulator (BEPS) at local (tower footprint) scale for different land cover types spread over North America: two broadleaf forests (Harvard, Massachusetts and Missouri Ozark, Missouri); two coniferous forests (Howland, Maine and Old Black Spruce, Saskatchewan); Lost Creek shrubland site (Wisconsin) and Mer Bleue petland (Ontario). BEPS calculates carbon fixation by scaling Farquhar's leaf biochemical model up to canopy level with stomatal conductance estimated by a modified version of the Ball-Woodrow-Berry model. The "big-leaf" approach was parameterized using derived leaf level parameters scaled up to canopy level by means of Leaf Area Index. The influence of sunlit

  13. Área foliar de duas trepadeiras infestantes de cana-de-açúcar utilizando dimensões lineares de folhas Foliar area estimate of two sugarcane-infesting weeds using leaf blade linear dimensions

    Directory of Open Access Journals (Sweden)

    N.P. Cardozo

    2009-01-01

    Full Text Available Esta pesquisa teve como objetivo obter uma equação, por meio de medidas lineares dimensionais das folhas, que permitisse a estimativa da área foliar de Momordica charantia e Pyrostegia venusta. Entre maio e dezembro de 2007, foram estudadas as correlações entre a área folia real (Sf e as medidas dimensionais do limbo foliar, como o comprimento ao longo da nervura principal (C e a largura máxima (L perpendicular à nervura principal. Todas as equações, exponenciais geométricas ou lineares simples, permitiram boas estimativas da área foliar. Do ponto de vista prático, sugere-se optar pela equação linear simples envolvendo o produto C x L, considerando-se o coeficiente linear igual a zero. Desse modo, a estimativa da área foliar de Momordica charantia pode ser feita pela fórmula Sf = 0,4963 x (C x L, e a de Pyrostegia venusta, por Sf = 0,6649 x (C x L.The aim of this study was to obtain a mathematical equation to estimate the leaf area of Momordica charantia and Pyrostegia venusta using linear leaf blade measurements. Correlation studies were conducted involving real leaf area (Sf and leaf length (C, maximum leaf width (L and C x L. The linear and geometric equations involving parameter C provided good leaf area estimates. From a practical viewpoint, the simple linear equation of the regression model is suggested using the C x L parameter, i.e., considering the linear coefficient equal to zero. Thus, leaf area estimate of Momordica charantia can be obtained by using the equation Sf = 0.4963 x (C x L, and that of Pyrostegia venusta by using equation Sf = 0.6649 x (C x L.

  14. Modelo matemático para estimativa da área foliar total de bananeira 'Prata-anã' Esteem method of total leaf area of 'Prata anã' banana tree

    Directory of Open Access Journals (Sweden)

    Moises Zucoloto

    2008-12-01

    Full Text Available O objetivo deste trabalho foi desenvolver um modelo para estimar a área foliar total de bananeira, cultivar Prata-Anã, utilizando dimensões lineares da terceira folha, como o comprimento, a largura e o número total de folhas na emissão da inflorescência. As regressões lineares foram determinadas considerando-se a área foliar total de cada planta (AFT como variável dependente e o comprimento (C e a largura (L da terceira folha, o produto de CxL, o número total de folhas por planta (N e o produto de CxLxN como variáveis independentes. O modelo linear que melhor estimou a área foliar total (AFTe da bananeira 'Prata-Anã', ao nível de 5% de significância com R² de 0,89, foi a equação AFTe = 0,5187(CxLxN + 9603,5.The objective of this work was to estimate the total leaf area of banana, cultivar Prata Anã, according to the linear dimensions of the third leaf, such as the length and the width and the total number of leves in the inflorescence emission. The linear regressions were determined considering total leaf area of each plant (AFT such as dependent variable and the length (C and the width (L of the third leaf, the product of CxL, the total number of leaf per plant (N and the product of CxLxN as independent variables. The best linear model that estimated the total leaf area (AFTe of banana 'Prata Anã' at the level of 5% of significance with R² of 0,89 was the equation AFTe = 0.5187 (CxLxN + 9603.5.

  15. [Relationships among leaf traits and their expression in different vegetation zones in Yanhe River basin, Northwest China].

    Science.gov (United States)

    Guo, Ru; Wen, Zhong-ming; Wang, Hong-xia; Qi, De-hui

    2015-12-01

    This article selected zonal plant communities as the research objects in different vegetation zones in Yanhe River basin. We measured six leaf traits of the dominant species and main accompanying species in each community, and then analyzed the relationships and their changes along with environmental gradients between these traits in order to understand the plant adaptation strategies to the environment changes. The results showed that the specific leaf area was significantly negatively correlated to leaf tissue density, area-based leaf nitrogen and phosphorus concentrations, and significantly positively correlated to mass-based leaf phosphorus concentration. Both the scaling relationships among these traits and plant life strategies were different among the three vegetation zones, the scaling-dependent relationship between leaf tissue density and specific leaf area was stronger in steppe and forest-steppe zones than in forest zone, but the correlations among area-based leaf nitrogen/phosphorus concentrations and specific leaf area and leaf tissue density were more significant in forest zone than in steppe zone. In the arid grassland and forest-steppe zone, plants give priority to defensive and stress resistance strategies, and in relatively moist nutrient-rich forest zone, plants give priority to fast growth and resource optimization allocation strategies.

  16. Peach leaf responses to soil and cement dust pollution.

    Science.gov (United States)

    Maletsika, Persefoni A; Nanos, George D; Stavroulakis, George G

    2015-10-01

    Dust pollution can negatively affect plant productivity in hot, dry and with high irradiance areas during summer. Soil or cement dust were applied on peach trees growing in a Mediterranean area with the above climatic characteristics. Soil and cement dust accumulation onto the leaves decreased the photosynthetically active radiation (PAR) available to the leaves without causing any shade effect. Soil and mainly cement dust deposition onto the leaves decreased stomatal conductance, photosynthetic and transpiration rates, and water use efficiency due possibly to stomatal blockage and other leaf cellular effects. In early autumn, rain events removed soil dust and leaf functions partly recovered, while cement dust created a crust partially remaining onto the leaves and causing more permanent stress. Leaf characteristics were differentially affected by the two dusts studied due to their different hydraulic properties. Leaf total chlorophyll decreased and total phenol content increased with dust accumulation late in the summer compared to control leaves due to intense oxidative stress. The two dusts did not cause serious metal imbalances to the leaves, except of lower leaf K content.

  17. An application of plot-scale NDVI in predicting carbon dioxide exchange and leaf area index in heterogeneous subarctic tundra

    Energy Technology Data Exchange (ETDEWEB)

    Dagg, J.; Lafleur, P.

    2010-07-01

    This paper reported on a study that examined the flow of carbon into and out of tundra ecosystems. It is necessary to accurately predict carbon dioxide (CO{sub 2}) exchange in the Tundra because of the impacts of climate change on carbon stored in permafrost. Understanding the relationships between the normalized difference vegetation index (NDVI) and vegetation and CO{sub 2} exchange may explain how small-scale variation in vegetation community extends to remotely sensed estimates of landscape characteristics. In this study, CO{sub 2} fluxes were measured with a portable chamber in a range of Tundra vegetation communities. Biomass and leaf area were measured with destructive harvest, and NDVI was obtained using a hand-held infrared camera. There was a weak correlation between NDVI and leaf area index in some vegetation communities, but a significant correlation between NDVI and biomass, including mosses. NDVI was found to be strongly related to photosynthetic activity and net CO{sub 2} uptake in all vegetation groups. However, NDVI related to ecosystem respiration only in wet sedge. It was concluded that at plot scale, the ability of NDVI to predict ecosystem properties and CO{sub 2} exchange in heterogeneous Tundra vegetation is variable.

  18. An application of plot-scale NDVI in predicting carbon dioxide exchange and leaf area index in heterogeneous subarctic tundra

    International Nuclear Information System (INIS)

    Dagg, J.; Lafleur, P.

    2010-01-01

    This paper reported on a study that examined the flow of carbon into and out of tundra ecosystems. It is necessary to accurately predict carbon dioxide (CO 2 ) exchange in the Tundra because of the impacts of climate change on carbon stored in permafrost. Understanding the relationships between the normalized difference vegetation index (NDVI) and vegetation and CO 2 exchange may explain how small-scale variation in vegetation community extends to remotely sensed estimates of landscape characteristics. In this study, CO 2 fluxes were measured with a portable chamber in a range of Tundra vegetation communities. Biomass and leaf area were measured with destructive harvest, and NDVI was obtained using a hand-held infrared camera. There was a weak correlation between NDVI and leaf area index in some vegetation communities, but a significant correlation between NDVI and biomass, including mosses. NDVI was found to be strongly related to photosynthetic activity and net CO 2 uptake in all vegetation groups. However, NDVI related to ecosystem respiration only in wet sedge. It was concluded that at plot scale, the ability of NDVI to predict ecosystem properties and CO 2 exchange in heterogeneous Tundra vegetation is variable.

  19. Generality of leaf trait relationships: A test across six biomes

    Energy Technology Data Exchange (ETDEWEB)

    Reich, P.B. [Univ. of Minnesota, Saint Paul, MN (United States). Dept. of Forest Resources; Ellsworth, D.S. [Brookhaven National Lab., Upton, NY (United States). Dept. of Applied Science; Walters, M.B. [Michigan State Univ., East Lansing, MI (United States). Dept. of Forestry; Vose, J.M. [Forest Service, Otto, NC (United States). Coweeta Hydrological Lab.; Gresham, C. [Clemson Univ., Georgetown, SC (United States). Baruch Forest Inst.; Volin, J.C. [Florida Atlantic Univ., Davie, FL (United States). Div. of Science; Bowman, W.D. [Inst. of Arctic and Alpine Research, Boulder, CO (United States). Mountain Research Station]|[Univ. of Colorado, Boulder, CO (United States). Dept. of Evolutionary, Population, and Organismic Biology

    1999-09-01

    Convergence in interspecific leaf trait relationships across diverse taxonomic groups and biomes would have important evolutionary and ecological implications. Such convergence has been hypothesized to result from trade-offs that limit the combination of plant traits for any species. Here the authors address this issue by testing for biome differences in the slope and intercept of interspecific relationships among leaf traits: longevity, net photosynthetic capacity (A{sub max}), leaf diffusive conductance (G{sub S}), specific leaf area (SLA), and nitrogen (N) status, for more than 100 species in six distinct biomes of the Americas. The six biomes were: alpine tundra-subalpine forest ecotone, cold temperate forest-prairie ecotone, montane cool temperate forest, desert shrubland, subtropical forest, and tropical rain forest. Despite large differences in climate and evolutionary history, in all biomes mass-based leaf N (N{sub mass}), SLA, G{sub S}, and A{sub max} were positively related to one another and decreased with increasing leaf life span. The relationships between pairs of leaf traits exhibited similar slopes among biomes, suggesting a predictable set of scaling relationships among key leaf morphological, chemical, and metabolic traits that are replicated globally among terrestrial ecosystems regardless of biome or vegetation type. However, the intercept (i.e., the overall elevation of regression lines) of relationships between pairs of leaf traits usually differed among biomes. With increasing aridity across sites, species had greater A{sub max} for a given level of G{sub S} and lower SLA for any given leaf life span. Using principal components analysis, most variation among species was explained by an axis related to mass-based leaf traits (A{sub max}, N, and SLA) while a second axis reflected climate, G{sub S}, and other area-based leaf traits.

  20. Influence of sewage sludge, as a substrate, in the plasticity of functional characteristics of plants.

    Science.gov (United States)

    da Silva, Vicente Elício Porfiro Sales Gonçalves; Buarque, Patrícia Marques Carneiro; Ferreira, Wanessa Nepomuceno; Buarque, Hugo Leonardo de Brito; Silva, Maria Amanda Menezes

    2018-04-24

    This work aimed to evaluate the effect of sewage sludge application as fertilizer on the plasticity of functional characteristics of species commonly found in the Caatinga. The research was developed in the nursery of the Federal Institute of Education, Science and Technology of Ceará (IFCE), Quixadá campus, located in northeastern Brazil. Three treatments were applied: raw sludge, sanitized sludge, and no manipulation. In each treatment, five species were planted, each with five individuals, totaling 75 individuals, which were tagged, and 4 months after germination, they were destroyed to obtain dry matter content (TMSF) from leaf, stem (TMSC), fine root (TMSRF), and thick root (TMSRG); leaf area; height and diameter of the seedling; and length above and below the ground. The sanitized sludge was responsible for giving higher values for leaf area, height of the seedlings, and diameter and length of stem and root. However, the dry matter content of the fine roots was higher in the treatment without manipulation. At the community level, as TMSRG increased, TMSC also increased, the same occurred between TMSRG and TMSRF, TMSC and TMSRF, and stem length and leaf area. In the treatment without manipulation, there was a positive correlation between leaf area, height and plant diameter, and negative correlation between root length and plant diameter. Thus, it can be concluded that the use of sanitized sludge is a good tool to increase the availability of soil resources, conferring to individuals' greater dry matter content, greater leaf area, and higher height and diameter above the ground.

  1. Accumulation of three different sizes of particulate matter on plant leaf surfaces: Effect on leaf traits

    Directory of Open Access Journals (Sweden)

    Chen Xiaoping

    2015-01-01

    Full Text Available Plants not only improve air quality by adsorbing particulate matter (PM on leaf surfaces but can also be affected by their accumulation. In this study, a field investigation was performed in Wuhan, China, into the relationship between seven leaf traits and the accumulation of three different sizes of PM (PM11, PM2.5 and PM0.2 on leaves. The retention abilities of plant leaves with respect to the three sizes of PM differed significantly at different sites and species. The average PM retention capabilities of plant leaves and specific leaf area (SLA were significantly greater in a seriously polluted area, whereas the average values of chlorophyll a (Chl a, chlorophyll b (Chl b, total chlorophyll, carotenoid, pH and relative water content (RWC were greater at the control site. SLA significantly positively correlated with the size of PM, but Chl a, Chl b, total chlorophyll, RWC significantly negatively correlated with the size of PM, whereas the pH did not correlate significantly with the the PM fractions. Additionally, SLA was found to be affected by large particles (PM11, p<0.01; PM2.5 had a more obvious effect on plant leaf traits than the other PM (p<0.05. Overall, the findings from this study provide useful information regarding the selection of plants to reduce atmospheric pollution.

  2. Comparable Analysis of the Distribution Functions of Runup Heights of the 1896, 1933 and 2011 Japanese Tsunamis in the Sanriku Area

    Science.gov (United States)

    Choi, B. H.; Min, B. I.; Yoshinobu, T.; Kim, K. O.; Pelinovsky, E.

    2012-04-01

    Data from a field survey of the 2011 tsunami in the Sanriku area of Japan is presented and used to plot the distribution function of runup heights along the coast. It is shown that the distribution function can be approximated using a theoretical log-normal curve [Choi et al, 2002]. The characteristics of the distribution functions derived from the runup-heights data obtained during the 2011 event are compared with data from two previous gigantic tsunamis (1896 and 1933) that occurred in almost the same region. The number of observations during the last tsunami is very large (more than 5,247), which provides an opportunity to revise the conception of the distribution of tsunami wave heights and the relationship between statistical characteristics and number of observations suggested by Kajiura [1983]. The distribution function of the 2011 event demonstrates the sensitivity to the number of observation points (many of them cannot be considered independent measurements) and can be used to determine the characteristic scale of the coast, which corresponds to the statistical independence of observed wave heights.

  3. Comparable analysis of the distribution functions of runup heights of the 1896, 1933 and 2011 Japanese Tsunamis in the Sanriku area

    Directory of Open Access Journals (Sweden)

    B. H. Choi

    2012-05-01

    Full Text Available Data from a field survey of the 2011 Tohoku-oki tsunami in the Sanriku area of Japan is used to plot the distribution function of runup heights along the coast. It is shown that the distribution function can be approximated by a theoretical log-normal curve. The characteristics of the distribution functions of the 2011 event are compared with data from two previous catastrophic tsunamis (1896 and 1933 that occurred in almost the same region. The number of observations during the last tsunami is very large, which provides an opportunity to revise the conception of the distribution of tsunami wave heights and the relationship between statistical characteristics and the number of observed runup heights suggested by Kajiura (1983 based on a small amount of data on previous tsunamis. The distribution function of the 2011 event demonstrates the sensitivity to the number of measurements (many of them cannot be considered independent measurements and can be used to determine the characteristic scale of the coast, which corresponds to the statistical independence of observed wave heights.

  4. Determination of normal heights in the area of Polish Economic Zone

    Directory of Open Access Journals (Sweden)

    Jerzy B. Rogowski

    2017-12-01

    Full Text Available The article presents a method of determining the level of the seabed in the Polish reference system. The authors show how to determine the ellipsoidal height of the seabed using GNSS measurements and single-beam echo sounders. The authors propose the transition to the system of normal heights referred to the average level of the North Sea as defined by the tide-gauge in Amsterdam to be made using the EGM 2008 model and data from the official Polish quasi-geoid model as well as data from another model distributed by GUGiK (Head Office of Geodesy and Cartography. The article presents also potential errors of the presented method.

  5. Soil Characteristics, Microbial Compostion of Plot, Leaf Count and ...

    African Journals Online (AJOL)

    Soil Characteristics, Microbial Compostion of Plot, Leaf Count and Sprout Studies of Cocoyam ( Colocasia [Schott] and Xanthosoma [Schott], Araceae) Collected in Edo State, ... Science, Technology and Arts Research Journal ... Government Areas (LGA) in Edo state and describe them based on leaf count and sprout

  6. Clinical measurement of the height of the interproximal contact area in maxillary anterior teeth.

    Science.gov (United States)

    Sghaireen, Mohd G; Al-Zarea, Bader K; Al-Shorman, Hisham M; Al-Omiri, Mahmoud K

    2013-11-01

    To clinically quantify the apicoincisal height of interproximal areas directly in patients' mouths. Thirty participants (11 females and 9 males, mean age=26±1.5 years) were recruited into this study. Measurement of interproximal contact areas was carried out directly in patients' mouths using digital caliper (TERENSA, USA) with measuring accuracy of 0.01 mm. The interproximal contact areas that were measured are: central incisor to central incisor, central incisor to lateral incisor, lateral incisor to canine, and canine to first premolar on both sides of the jaw. Statistical significance was based on probability values less than 0.05 (pcontact point was the one present between central incisors and it ranged from 2.9 to 6.5 mm. On the other hand, the contact point between canine and first premolar was the smallest on both sides of the arch and ranged from 0.6 to 2.5 mm. The dimensions of the contact points declined as we move from anterior area backwards. Statistical analysis using t-test showed that there were significant differences between the measurements of interproximal points of each tooth (Pcontact point decreased as we moved from anterior to posterior teeth. The contact area between the central incisors was largest and the one between canine and premolar was the smallest. This study is the first to report direct intra-oral clinical measurement of contact points. Clinical evaluation of contact point dimensions using digital caliber is a viable, quick and accurate method to use.

  7. Estimação da área foliar do algodoeiro por meio de dimensões e massa das folhas Cotton leaf area estimates based on leaf dimensions and dry mass methods

    Directory of Open Access Journals (Sweden)

    José Eduardo B. A. Monteiro

    2005-01-01

    Full Text Available O objetivo deste trabalho foi avaliar dois métodos de estimação da área foliar do algodoeiro, por meio de suas dimensões e massa seca das folhas. Foram utilizadas as cultivares IAC 23 e Coodetec 401. No método que utilizou dimensões, as folhas do algodoeiro foram agrupadas em novas, cordiformes e maduras. Para cada tipo de folha, de cada cultivar, foi determinado um fator de forma (FF por meio de análise de regressão entre o produto do comprimento (C pela largura (L e a área das folhas. Avaliou-se a correlação entre a área foliar estimada pelo fator FF e sua medida direta, utilizando-se dados independentes. Testou-se, ainda, um fator único para cada cultivar, independente do estádio da cultura e, também, um fator geral para as duas cultivares. No método que utilizou a massa seca, as folhas foram agrupadas em novas e maduras. Determinou-se o fator de massa seca (FM por meio da análise de regressão entre a massa seca de folhas e respectivas áreas foliares. Em seguida, avaliou-se a correlação entre dados estimados por FM e dados medidos de forma direta, em nova amostra. O método das dimensões é viável para a estimação de área foliar do algodoeiro, por apresentar boa precisão e exatidão, com r² entre 0,71 e 0,98 e com coeficiente angular da regressão entre 0,87 e 0,95. No entanto, pelo método da massa seca, observaram-se precisão e exatidão maiores, com r² entre 0,94 e 0,98, e coeficiente angular da regressão entre 0,97 e 1,00, com a vantagem de ser menos trabalhoso.The objective of this study was to evaluate two different methods to estimate cotton leaf area (LA, based on leaf dimensions (length - L and width - W and leaf dry mass (DM. Two cultivars, IAC 23 and Coodetec 401, were used. For leaf dimensions method, leaves were classified by age: young, heart-shape, and mature. For each age class, a leaf shape factor (LSF was obtained by simple linear regression between L*W and LA. For leaf dry mass method, leaves

  8. Fear of heights and visual height intolerance.

    Science.gov (United States)

    Brandt, Thomas; Huppert, Doreen

    2014-02-01

    The aim of this review is, first, to cover the different aspects of visual height intolerance such as historical descriptions, definition of terms, phenomenology of the condition, neurophysiological control of gaze, stance and locomotion, and therapy, and, second, to identify warranted epidemiological and experimental studies. Vivid descriptions of fear of heights can be found in ancient texts from the Greek, Roman, and Chinese classics. The life-time prevalence of visual height intolerance is as high as 28% in the general population, and about 50% of those who are susceptible report an impact on quality of life. When exposed to heights, visual exploration by eye and head movements is restricted, and the velocity of locomotion is reduced. Therapy for fear of heights is dominated by the behavioral techniques applied during real or virtual reality exposure. Their efficacy might be facilitated by the administration of D-cycloserine or glucocorticoids. Visual height intolerance has a considerable impact on daily life and interpersonal interactions. It is much more frequent than fear of heights, which is defined as an environmental subtype of a specific phobia. There is certainly a continuum stretching from acrophobia to a less-pronounced visual height intolerance, to which the categorical distinction of a specific phobia does not apply.

  9. Weighting of field heights for sharpness and noisiness

    Science.gov (United States)

    Keelan, Brian W.; Jin, Elaine W.

    2009-01-01

    Weighting of field heights is important in cases when a single numerical value needs to be calculated that characterizes an attribute's overall impact on perceived image quality. In this paper we report an observer study to derive the weighting of field heights for sharpness and noisiness. One-hundred-forty images were selected to represent a typical consumer photo space distribution. Fifty-three sample points were sampled per image, representing field heights of 0, 14, 32, 42, 51, 58, 71, 76, 86% and 100%. Six observers participated in this study. The field weights derived in this report include both: the effect of area versus field height (which is a purely objective, geometric factor); and the effect of the spatial distribution of image content that draws attention to or masks each of these image structure attributes. The results show that relative to the geometrical area weights, sharpness weights were skewed to lower field heights, because sharpness-critical subject matter was often positioned relatively near the center of an image. Conversely, because noise can be masked by signal, noisiness-critical content (such as blue skies, skin tones, walls, etc.) tended to occur farther from the center of an image, causing the weights to be skewed to higher field heights.

  10. [Models for biomass estimation of four shrub species planted in urban area of Xi'an city, Northwest China].

    Science.gov (United States)

    Yao, Zheng-Yang; Liu, Jian-Jun

    2014-01-01

    Four common greening shrub species (i. e. Ligustrum quihoui, Buxus bodinieri, Berberis xinganensis and Buxus megistophylla) in Xi'an City were selected to develop the highest correlation and best-fit estimation models for the organ (branch, leaf and root) and total biomass against different independent variables. The results indicated that the organ and total biomass optimal models of the four shrubs were power functional model (CAR model) except for the leaf biomass model of B. megistophylla which was logarithmic functional model (VAR model). The independent variables included basal diameter, crown diameter, crown diameter multiplied by height, canopy area and canopy volume. B. megistophylla significantly differed from the other three shrub species in the independent variable selection, which were basal diameter and crown-related factors, respectively.

  11. Characterization of leaf area of Agave fourcroydes Lem. plants obtained from asexual propagation

    Directory of Open Access Journals (Sweden)

    Maryla Sosa del Castillo

    2014-01-01

    Full Text Available The henequen (Agave fourcroydes Lem. is a crop of great economic importance. This study was aimed to characterize the leaf surface of henequen plants variety `Sac Ki' obtained by asexual propagation methods. In vitro plants, shoots of bulbils of in vitro plants, shoots of rhizomes of in vitro plants, shoots of bulbils of field plants and shoots of field rhizomes were used. At 7 and 15 months after planting in the nursery, the epidermis was characterized through the stomatal index and stomatal density. Moreover, the conductor vessels and fiber bundles in leaf mesophyll, were characterized. It was found that the leaf surface of henequen plants variety `Sac Ki' obtained by different methods of asexual propagation showed similar anatomical structures. However, it was observed that in vitro plants were different from the rest in terms of stomatal index and stomatal density in both time points. It was suggesting a accommodate response to environmental conditions. Key words: stomatic density, stomatic index, in vitro plants

  12. Dynamics of production and forage utilization on elephant grass pastures managed with different post-grazing heights

    Directory of Open Access Journals (Sweden)

    Braulio Maia de Lana Sousa

    2012-08-01

    Full Text Available This study was conducted to evaluate the daily production of forage and its morphological components, as well as the potential of forage utilization in pastures of Pennisetum purpureum cv. Napier managed with three post-grazing heights (30, 50 and 70 cm. Two experiments were carried out: one from February to May 2009 and another from December 2009 to May 2010, characterizing months of summer and fall. The experimental design was of completely randomized blocks with three replicates. The grazing was performed by crossbred heifers of approximately 270 kg body weight, when the sward intercepted 95% of the incoming light. In both experiments, the pastures managed with post-grazing height of 30 cm, in the summer months (December to March, presented lower daily production of leaves and stems, as well as less daily leaf senescence, which resulted in lower daily forage production and accumulation in comparison with those managed at 70 and 50 cm. In the period from February to March 2009 (experiment 1 and December 2009 to March 2010 (experiment 2, pastures presented greater daily production of leaves and forage, greater daily forage accumulation and more daily leaf senescence in relation to the months of April and May 2009 and 2010. On the other hand, the daily production of stems was higher in the fall, in comparison with the summer. Therefore, elevation in the post-grazing height, especially in the summer, increases the regrowth vigor of elephant grass cv. Napier.

  13. Biomass, Leaf Area, and Resource Availability of Kudzu Dominated Plant Communities Following Herbicide Treatment

    Energy Technology Data Exchange (ETDEWEB)

    L.T. Rader

    2001-10-01

    Kudzu is an exotic vine that threatens the forests of the southern U.S. Five herbicides were tested with regard to their efficacy in controlling kudzu, community recover was monitored, and interactions with planted pines were studied. The sites selected were old farm sites dominated by kudzu.These were burned following herbicide treatment. The herbicides included triclopyr, clopyralid, metsulfuron, tebuthiuron, and picloram plus 2,4-D. Pine seedlings were planted the following year. Regression equations were developed for predicting biomass and leaf area. Four distinct plant communities resulted from the treatments. The untreated check continued to be kudzu dominated. Blackberry dominated the clopyradid treatment. Metsulfron, trychlopyr and picloram treated sites resulted in herbaceous dominated communities. The tebuthiuron treatment maintained all vegetation low.

  14. Using the Normalized Differential Wetness Index to Scale Leaf Area Index, Create Three-Dimensional Classification Maps, and Scale Seasonal Evapotranspiration Depletions in Canopies Along the Middle Rio Grande Riparian CorridorCorridor

    Science.gov (United States)

    McDonnell, D. E.; Cleverly, J. R.; Dahm, C. N.; Coonrod, J. A.

    2005-12-01

    This research creates temporally and spatially explicit data layers of vegetation, leaf area index (LAI), three dimensional (3D) vegetation classification maps, and seasonal evapotranspiration (ET) depletions along the middle Rio Grande riparian corridor. The first part of this work produces two dimensional (2D) classification maps of native and non-native canopy vegetation using temporal patterns and the decision tree classifier in ENVI 4.0 (Research Systems Inc. Boulder, Colorado). The second part of this work correlates the normalized differential wetness index (NDWI) with field measurements of plant area index (PAI), stem area index (SAI), and leaf area index (LAI) using the LAI-2000 Plant Canopy Analyzer (PCA) (LICOR Inc., Lincoln, Nebraska). SAI is measured in winter to capture only branches and stems. PAI is measured during the growing season. Field measurements taken within 10 days of image capture dates provide adequate correlations though the closer the dates the better the correlation. LAI represents the surface area of active green leafy vegetation. NDWI correlates with both PAI and estimated LAI in both Tamarisk chinensis and Populus deltoides ssp. Wislizeni sites better than the more traditional normalized differential vegetation index (NDVI). This study also suggests that winter PCA measurements approximate SAI which should be subtracted from PAI in woody vegetation like T. chinensis and Salix exigua stands. The results show that correcting for leaf geometry by multiplying T. chinensis areas with cylindrical cladophylls by pi and the remaining flat leaf vegetation by two yields the best relationship between NDWI and total LAI. The 2Dclassification maps can be placed on top of relief maps of LAI to produce 3D classification maps. The final part of this research scales ET from four 3D eddy covariance towers located in two T. chinensis and two P. deltoides study sites. ET is regressed with LAI, percent daylight (PD), and average hourly incoming net

  15. Adaptation of European beech (Fagus silvatica L.) to different ecological conditions: leaf size variation

    International Nuclear Information System (INIS)

    Barna, M.

    2004-01-01

    In beech trees, both leaf morphology and leaf area show considerable adaptation capabilities to the local radiation climate. The plants adapting to shade conditions create large leaf area with high chlorophyll concentration and high water content in the living tissues. On the other hand, the leaves of plants exposed to radiation of higher intensity have smaller area, several layers of mesophyll, thick epidermis and cuticle, higher dry weight, higher energy potential of the dry mass and several other characteristic properties

  16. The importance of hydraulic architecture to the distribution patterns of trees in a central Amazonian forest.

    Science.gov (United States)

    Cosme, Luiza H M; Schietti, Juliana; Costa, Flávia R C; Oliveira, Rafael S

    2017-07-01

    Species distributions and assemblage composition may be the result of trait selection through environmental filters. Here, we ask whether filtering of species at the local scale could be attributed to their hydraulic architectural traits, revealing the basis of hydrological microhabitat partitioning in a Central Amazonian forest. We analyzed the hydraulic characteristics at tissue (anatomical traits, wood specific gravity (WSG)), organ (leaf area, specific leaf area (SLA), leaf area : sapwood area ratio) and whole-plant (height) levels for 28 pairs of congeneric species from 14 genera restricted to either valleys or plateaus of a terra-firme forest in Central Amazonia. On plateaus, species had higher WSG, but lower mean vessel area, mean vessel hydraulic diameter, sapwood area and SLA than in valleys; traits commonly associated with hydraulic safety. Mean vessel hydraulic diameter and mean vessel area increased with height for both habitats, but leaf area and leaf area : sapwood area ratio investments with tree height declined in valley vs plateau species. [Correction added after online publication 29 March 2017: the preceding sentence has been reworded.] Two strategies for either efficiency or safety were detected, based on vessel size or allocation to sapwood. In conclusion, contrasting hydrological conditions act as environmental filters, generating differences in species composition at the local scale. This has important implications for the prediction of species distributions under future climate change scenarios. © 2017 The Authors. New Phytologist © 2017 New Phytologist Trust.

  17. Differences between height- and light-dependent changes in shoot traits in five deciduous tree species.

    Science.gov (United States)

    Osada, Noriyuki; Okabe, Yoshihiko; Hayashi, Daisuke; Katsuyama, Tomonori; Tokuchi, Naoko

    2014-01-01

    The effects of tree height on shoot traits may in some cases differ in magnitude and direction from the effects of light. Nevertheless, general patterns of change in shoot traits in relation to variations in height and light have not so far been revealed. A comprehensive analysis of the differences between the effects of height and light on a range of leaf and shoot traits is important for the scaling of these traits to individual trees. We investigated the biomass allocation and structure of current-year shoots at the top of the crowns of five deciduous tree species in Japan. Height effect was investigated by comparing shoot traits among trees of different heights growing under a high light environment. The effects of light were examined by comparing saplings growing in high- and low-light environments. The effects of light were significant for most traits, while those of height were not significant for some traits. The magnitudes of the effects of light were larger than those of height for most traits related to biomass allocation. There was an extreme difference between the effects of height and light in the direction of change in the length of current-year shoots and in the number of standing leaves. The measures of both parameters increased with the increase in light, but decreased with the increase in tree height. Thus, the effects of height and light on diverse traits at the level of current-year shoots were not always similar. These results suggest that great care must be taken when scaling shoot traits from small trees to tall trees because the effects of height and light can be complex.

  18. Chemical and mechanical changes during leaf expansion of four woody species of dry Restinga woodland.

    Science.gov (United States)

    Schlindwein, C C D; Fett-Neto, A G; Dillenburg, L R

    2006-07-01

    Young leaves are preferential targets for herbivores, and plants have developed different strategies to protect them. This study aimed to evaluate different leaf attributes of presumed relevance in protection against herbivory in four woody species (Erythroxylum argentinum, Lithrea brasiliensis, Myrciaria cuspidata, and Myrsine umbellata), growing in a dry restinga woodland in southern Brazil. Evaluation of leaf parameters was made through single-point sampling of leaves (leaf mass per area and leaf contents of nitrogen, carbon, and pigments) at three developmental stages and through time-course sampling of expanding leaves (area and strength). Leaves of M. umbellata showed the highest leaf mass per area (LMA), the largest area, and the longest expansion period. On the other extreme, Myrc. cuspidata had the smallest LMA and leaf size, and the shortest expansion period. Similarly to L. brasiliensis, it displayed red young leaves. None of the species showed delayed-greening, which might be related to the high-irradiance growth conditions. Nitrogen contents reduced with leaf maturity and reached the highest values in the young leaves of E. argentinum and Myrc. cuspidata and the lowest in M. umbellata. Each species seems to present a different set of protective attributes during leaf expansion. Myrciaria cuspidata appears to rely mostly on chemical defences to protect its soft leaves, and anthocyanins might play this role at leaf youth, while M. umbellata seems to invest more on mechanical defences, even at early stages of leaf growth, as well as on a low allocation of nitrogen to the leaves. The other species display intermediate characteristics.

  19. Endophytic fungi reduce leaf-cutting ant damage to seedlings

    Science.gov (United States)

    Bittleston, L. S.; Brockmann, F.; Wcislo, W.; Van Bael, S. A.

    2011-01-01

    Our study examines how the mutualism between Atta colombica leaf-cutting ants and their cultivated fungus is influenced by the presence of diverse foliar endophytic fungi (endophytes) at high densities in tropical leaf tissues. We conducted laboratory choice trials in which ant colonies chose between Cordia alliodora seedlings with high (Ehigh) or low (Elow) densities of endophytes. The Ehigh seedlings contained 5.5 times higher endophyte content and a greater diversity of fungal morphospecies than the Elow treatment, and endophyte content was not correlated with leaf toughness or thickness. Leaf-cutting ants cut over 2.5 times the leaf area from Elow relative to Ehigh seedlings and had a tendency to recruit more ants to Elow plants. Our findings suggest that leaf-cutting ants may incur costs from cutting and processing leaves with high endophyte loads, which could impact Neotropical forests by causing variable damage rates within plant communities. PMID:20610420

  20. Remote sensing based mapping of leaf nitrogen and leaf area index in European landscapes using the REGularized canopy reFLECtance (REGFLEC) model

    DEFF Research Database (Denmark)

    Boegh, E.; Houborg, R.; Bienkowski, J.

    2011-01-01

    index (LAI) are important determinants of the maximum CO2 Methods/Approach uptake by plants and trees. In the EU project NitroEurope, high spatial resolution (10-20 m) remote sensing data from the HRG and HRVIR sensors onboard the SPOT satellites were acquired to derive maps of leaf N and LAI for 5...... European landscapes. The estimations of leaf N, Cab and LAI soil reflectance parameters and canopy parameters are discussed in relation to the prevailing soil types and vegetation characteristics of land cover classes across the 5 European landscapes....

  1. Model estimates of leaf area and reference canopy stomatal conductance suggest correlation between phenology and physiology in both trembling aspen and red pine

    Science.gov (United States)

    Mackay, D. S.; Ewers, B. E.; Kruger, E. L.

    2006-12-01

    Phenological variations impact water and carbon fluxes, as evidenced by the large interannual variability of net ecosystem exchange of carbon dioxide and evapotranspiration (ET). In northern Wisconsin we observed daily variations of canopy transpiration from hardwoods from 1.0 to 1.7 mm/day during the leaf unfolding period and 1.7 to 2.6 mm/day with leaves fully out. Correlations between such flux rates and phenology have not been extensively tested and mechanistic connections are in their infancy. Some data suggest that stomatal conductance and photosynthesis increases up to full expansion. Moreover, in conifers, the interaction of phenology and physiology is more complicated than in deciduous trees because needles are retained for several years. Using inverse modeling with a coupled photosynthesis-transpiration model we estimated reference canopy stomatal conductance, Gsref, for red pine (Pinus resinosa), and Gsref and leaf area index, L, for trembling aspen (Populus tremuloides), using 30-min continuous sap flux data spanning a period from just prior to the start of leaf expansion to just after leaf senescence. The red pine showed Gsref ramp up from 105 to 179 mmol m-2 leaf s-1, which represented a 37 to 50 percent increase in Gsref after accounting for maximum possible changes in L. After full leaf out, the trembling aspen were almost immediately defoliated, and then reflushed after three weeks. Model estimates of L reflected this pattern and were consistent with measurements. However, Gsref never exceeded 45 mmol m-2 s-1 prior to defoliation, but peaked at 112 mmol m-2 s-1 after reflushing. These results support the need for further work that aims to separate phenology and physiology.

  2. Tradeoff between stem hydraulic efficiency and mechanical strength affects leaf-stem allometry in 28 Ficus tree species

    NARCIS (Netherlands)

    Fan, Ze Xin; Sterck, Frank; Zhang, Shi Bao; Fu, Pei Li; Hao, Guang You

    2017-01-01

    Leaf-stem allometry is an important spectrum that linked to biomass allocation and life history strategy in plants, although the determinants and evolutionary significance of leaf-stem allometry remain poorly understood. Leaf and stem architectures - including stem area/mass, petiole area/mass,

  3. Leaf area development, dry weight accumulation and solar energy conversion efficiencies of Phaseolus vulgaris L. under different soil moisture levels near Nairobi, Kenya

    NARCIS (Netherlands)

    Muniafu, M.M.; Macharia, J.N.M.; Stigter, C.J.; Coulson, G.L.

    1999-01-01

    Leaf area development, dry weight accumulation and solar energy conversion efficiencies of Phaseolus vulgaris L. cv GLP-2 under two soil moisture levels in two contrasting seasons near Nairobi, Kenya were investigated. The experiment confirms that dry weights and yields of Phaseolus vulgaris are

  4. IN SITU AND MODIS MOD15A2 LEAF AREA INDEX MEASUREMENTS OF A MID-ATLANTIC DECIDOUS FOREST SITE: PERSPECTIVES FROM FOUR-YEARS OF FIELD STUDIES

    Science.gov (United States)

    The U.S. Environmental Protection Agency is interested in leaf area index as it pertains to biogenic emissions, atmospheric pollutant deposition, ecological indicators, vegetation phenology, and land cover mapping.

  5. Leaf phenotypic variation and developmental instability in relation to different light regimes

    Directory of Open Access Journals (Sweden)

    Henrique Venâncio

    2016-06-01

    Full Text Available ABSTRACT For pioneer plants, shaded habitats represent a stressful condition, where sunlight exposure is below the optimum level and so leaves expand in order to intercept a greater amount of light. We investigated changes in both phenotypic variation and stress of Bauhinia brevipes in sunny and shaded microhabitats. Leaf area was used as a measure of phenotypic variation, whereas leaf asymmetry (difference between right and left sides of leaves, was used as a measure of stress. We hypothesized an increase in leaf area and stress in shaded locations, which might indicate that B. brevipes was compensating for low light absorption, and elevated levels of stress, respectively. Plants in the sun fitted a fluctuating asymmetry pattern (normal distribution of right minus left sides, while shaded plants were clearly antisymmetric (bimodal distribution of leaf side differences. Leaf asymmetry and area were 5% and 26.8% higher in plants in the shade compared to plants in the sun, respectively. These results were expected since B. brevipes is found predominantly in open areas; so sunlight exposure is important for its development. The presence of antisymmetry is rare in studies of developmental instability, and here it might indicate higher stress compared to plants with fluctuating asymmetry.

  6. QTL mapping of flag leaf-related traits in wheat (Triticum aestivum L.).

    Science.gov (United States)

    Liu, Kaiye; Xu, Hao; Liu, Gang; Guan, Panfeng; Zhou, Xueyao; Peng, Huiru; Yao, Yingyin; Ni, Zhongfu; Sun, Qixin; Du, Jinkun

    2018-04-01

    QTL controlling flag leaf length, flag leaf width, flag leaf area and flag leaf angle were mapped in wheat. This study aimed to advance our understanding of the genetic mechanisms underlying morphological traits of the flag leaves of wheat (Triticum aestivum L.). A recombinant inbred line (RIL) population derived from ND3331 and the Tibetan semi-wild wheat Zang1817 was used to identify quantitative trait loci (QTLs) controlling flag leaf length (FLL), flag leaf width (FLW), flag leaf area (FLA), and flag leaf angle (FLANG). Using an available simple sequence repeat genetic linkage map, 23 putative QTLs for FLL, FLW, FLA, and FLANG were detected on chromosomes 1B, 2B, 3A, 3D, 4B, 5A, 6B, 7B, and 7D. Individual QTL explained 4.3-68.52% of the phenotypic variance in different environments. Four QTLs for FLL, two for FLW, four for FLA, and five for FLANG were detected in at least two environments. Positive alleles of 17 QTLs for flag leaf-related traits originated from ND3331 and 6 originated from Zang1817. QTLs with pleiotropic effects or multiple linked QTL were also identified on chromosomes 1B, 4B, and 5A; these are potential target regions for fine-mapping and marker-assisted selection in wheat breeding programs.

  7. Global meta-analysis of leaf area index in wetlands indicates uncertainties in understanding of their ecosystem function

    Science.gov (United States)

    Dronova, I.; Taddeo, S.; Foster, K.

    2017-12-01

    Projecting ecosystem responses to global change relies on the accurate understanding of properties governing their functions in different environments. An important variable in models of ecosystem function is canopy leaf area index (LAI; leaf area per unit ground area) declared as one of the Essential Climate Variables in the Global Climate Observing System and extensively measured in terrestrial landscapes. However, wetlands have been largely under-represented in these efforts, which globally limits understanding of their contribution to carbon sequestration, climate regulation and resilience to natural and anthropogenic disturbances. This study provides a global synthesis of >350 wetland-specific LAI observations from 182 studies and compares LAI among wetland ecosystem and vegetation types, biomes and measurement approaches. Results indicate that most wetland types and even individual locations show a substantial local dispersion of LAI values (average coefficient of variation 65%) due to heterogeneity of environmental properties and vegetation composition. Such variation indicates that mean LAI values may not sufficiently represent complex wetland environments, and the use of this index in ecosystem function models needs to incorporate within-site variation in canopy properties. Mean LAI did not significantly differ between direct and indirect measurement methods on a pooled global sample; however, within some of the specific biomes and wetland types significant contrasts between these approaches were detected. These contrasts highlight unique aspects of wetland vegetation physiology and canopy structure affecting measurement principles that need to be considered in generalizing canopy properties in ecosystem models. Finally, efforts to assess wetland LAI using remote sensing strongly indicate the promise of this technology for cost-effective regional-scale modeling of canopy properties similar to terrestrial systems. However, such efforts urgently require more

  8. Maize YABBY genes drooping leaf1 and drooping leaf2 affect agronomic traits by regulating leaf architecture

    Science.gov (United States)

    Leaf architectural traits, such as length, width and angle, directly influence canopy structure and light penetration, photosynthate production and overall yield. We discovered and characterized a maize (Zea mays) mutant with aberrant leaf architecture we named drooping leaf1 (drl1), as leaf blades ...

  9. Leaf life span plasticity in tropical seedlings grown under contrasting light regimes.

    Science.gov (United States)

    Vincent, Gregoire

    2006-02-01

    The phenotypic plasticity of leaf life span in response to low resource conditions has a potentially large impact on the plant carbon budget, notably in evergreen species not subject to seasonal leaf shedding, but has rarely been well documented. This study evaluates the plasticity of leaf longevity, in terms of its quantitative importance to the plant carbon balance under limiting light. Seedlings of four tropical tree species with contrasting light requirements (Alstonia scholaris, Hevea brasiliensis, Durio zibethinus and Lansium domesticum) were grown under three light regimes (full sunlight, 45 % sunlight and 12 % sunlight). Their leaf dynamics were monitored over 18 months. All species showed a considerable level of plasticity with regard to leaf life span: over the range of light levels explored, the ratio of the range to the mean value of life span varied from 29 %, for the least plastic species, to 84 %, for the most. The common trend was for leaf life span to increase with decreasing light intensity. The plasticity apparent in leaf life span was similar in magnitude to the plasticity observed in specific leaf area and photosynthetic rate, implying that it has a significant impact on carbon gain efficiency when plants acclimate to different light regimes. In all species, median survival time was negatively correlated with leaf photosynthetic capacity (or its proxy, the nitrogen content per unit area) and leaf emergence rate. Longer leaf life spans under low light are likely to be a consequence of slower ageing as a result of a slower photosynthetic metabolism.

  10. Relating Stomatal Conductance to Leaf Functional Traits.

    Science.gov (United States)

    Kröber, Wenzel; Plath, Isa; Heklau, Heike; Bruelheide, Helge

    2015-10-12

    Leaf functional traits are important because they reflect physiological functions, such as transpiration and carbon assimilation. In particular, morphological leaf traits have the potential to summarize plants strategies in terms of water use efficiency, growth pattern and nutrient use. The leaf economics spectrum (LES) is a recognized framework in functional plant ecology and reflects a gradient of increasing specific leaf area (SLA), leaf nitrogen, phosphorus and cation content, and decreasing leaf dry matter content (LDMC) and carbon nitrogen ratio (CN). The LES describes different strategies ranging from that of short-lived leaves with high photosynthetic capacity per leaf mass to long-lived leaves with low mass-based carbon assimilation rates. However, traits that are not included in the LES might provide additional information on the species' physiology, such as those related to stomatal control. Protocols are presented for a wide range of leaf functional traits, including traits of the LES, but also traits that are independent of the LES. In particular, a new method is introduced that relates the plants' regulatory behavior in stomatal conductance to vapor pressure deficit. The resulting parameters of stomatal regulation can then be compared to the LES and other plant functional traits. The results show that functional leaf traits of the LES were also valid predictors for the parameters of stomatal regulation. For example, leaf carbon concentration was positively related to the vapor pressure deficit (vpd) at the point of inflection and the maximum of the conductance-vpd curve. However, traits that are not included in the LES added information in explaining parameters of stomatal control: the vpd at the point of inflection of the conductance-vpd curve was lower for species with higher stomatal density and higher stomatal index. Overall, stomata and vein traits were more powerful predictors for explaining stomatal regulation than traits used in the LES.

  11. Leaf Dynamics of Panicum maximum under Future Climatic Changes.

    Science.gov (United States)

    Britto de Assis Prado, Carlos Henrique; Haik Guedes de Camargo-Bortolin, Lívia; Castro, Érique; Martinez, Carlos Alberto

    2016-01-01

    Panicum maximum Jacq. 'Mombaça' (C4) was grown in field conditions with sufficient water and nutrients to examine the effects of warming and elevated CO2 concentrations during the winter. Plants were exposed to either the ambient temperature and regular atmospheric CO2 (Control); elevated CO2 (600 ppm, eC); canopy warming (+2°C above regular canopy temperature, eT); or elevated CO2 and canopy warming (eC+eT). The temperatures and CO2 in the field were controlled by temperature free-air controlled enhancement (T-FACE) and mini free-air CO2 enrichment (miniFACE) facilities. The most green, expanding, and expanded leaves and the highest leaf appearance rate (LAR, leaves day(-1)) and leaf elongation rate (LER, cm day(-1)) were observed under eT. Leaf area and leaf biomass were higher in the eT and eC+eT treatments. The higher LER and LAR without significant differences in the number of senescent leaves could explain why tillers had higher foliage area and leaf biomass in the eT treatment. The eC treatment had the lowest LER and the fewest expanded and green leaves, similar to Control. The inhibitory effect of eC on foliage development in winter was indicated by the fewer green, expanded, and expanding leaves under eC+eT than eT. The stimulatory and inhibitory effects of the eT and eC treatments, respectively, on foliage raised and lowered, respectively, the foliar nitrogen concentration. The inhibition of foliage by eC was confirmed by the eC treatment having the lowest leaf/stem biomass ratio and by the change in leaf biomass-area relationships from linear or exponential growth to rectangular hyperbolic growth under eC. Besides, eC+eT had a synergist effect, speeding up leaf maturation. Therefore, with sufficient water and nutrients in winter, the inhibitory effect of elevated CO2 on foliage could be partially offset by elevated temperatures and relatively high P. maximum foliage production could be achieved under future climatic change.

  12. Climate controls photosynthetic capacity more than leaf nitrogen contents

    Science.gov (United States)

    Ali, A. A.; Xu, C.; McDowell, N. G.

    2013-12-01

    Global vegetation models continue to lack the ability to make reliable predictions because the photosynthetic capacity varies a lot with growth conditions, season and among species. It is likely that vegetation models link photosynthetic capacity to concurrent changes in leaf nitrogen content only. To improve the predictions of the vegetation models, there is an urgent need to review species growth conditions and their seasonal response to changing climate. We sampled the global distribution of the Vcmax (maximum carboxylation rates) data of various species across different environmental gradients from the literature and standardized its value to 25 degree Celcius. We found that species explained the largest variation in (1) the photosynthetic capacity and (2) the proportion of nitrogen allocated for rubisco (PNcb). Surprisingly, climate variables explained more variations in photosynthetic capacity as well as PNcb than leaf nitrogen content and/or specific leaf area. The chief climate variables that explain variation in photosynthesis and PNcb were radiation, temperature and daylength. Our analysis suggests that species have the greatest control over photosynthesis and PNcb. Further, compared to leaf nitrogen content and/or specific leaf area, climate variables have more control over photosynthesis and PNcb. Therefore, climate variables should be incorporated in the global vegetation models when making predictions about the photosynthetic capacity.

  13. Memory for target height is scaled to observer height.

    Science.gov (United States)

    Twedt, Elyssa; Crawford, L Elizabeth; Proffitt, Dennis R

    2012-04-01

    According to the embodied approach to visual perception, individuals scale the environment to their bodies. This approach highlights the central role of the body for immediate, situated action. The present experiments addressed whether body scaling--specifically, eye-height scaling--occurs in memory when action is not immediate. Participants viewed standard targets that were either the same height as, taller than, or shorter than themselves. Participants then viewed a comparison target and judged whether the comparison was taller or shorter than the standard target. Participants were most accurate when the standard target height matched their own heights, taking into account postural changes. Participants were biased to underestimate standard target height, in general, and to push standard target height away from their own heights. These results are consistent with the literature on eye-height scaling in visual perception and suggest that body scaling is not only a useful metric for perception and action, but is also preserved in memory.

  14. Características morfogênicas e estruturais do capim-tifton 85 sob doses de nitrogênio e alturas de corte Morphogenic and structural characteristics of tifton 85 bermudagrass under different nitrogen doses and harvesting heights

    Directory of Open Access Journals (Sweden)

    Odilon Gomes Pereira

    2011-09-01

    Full Text Available Objetivou-se avaliar características morfogênicas e estruturais do capim-tifton 85 adubado com nitrogênio nas doses de 0, 33, 66, 100 ou 133 kg/ha.ano em três alturas à data de corte (30, 40 ou 50 cm. O delineamento foi inteiramente casualizado em esquema de parcelas subdivididas com três repetições. As doses de nitrogênio (N corresponderam às parcelas e as alturas ao tempo de corte, às subparcelas. A taxa de aparecimento foliar do capim-tifton 85 foi influenciada de forma negativa pela altura da planta na época do corte e de maneira positiva pela dose de nitrogênio. A taxa de senescência foliar e o número total de folhas por perfilho diminuíram, enquanto a taxa de alongamento foliar aumentou com a adubação nitrogenada. Por outro lado, a taxa de senescência foliar e o número total de folhas por perfilho aumentaram com a altura das plantas na época da colheita. Já o número de folhas vivas por perfilho não foi influenciado pela dose de nitrogênio nem pela altura da planta na época do corte. Para uma mesma altura do capim-tifton 85 na época da colheita, a adubação nitrogenada aumenta o fluxo de tecidos e a frequência de desfolhação. Na condição de desfolhação intermitente, o capim-tifton 85 deve ser colhido com altura de até 30 cm.The objective of this study was to evaluate the morphogenic and structural characteristics of tifton 85 bermudagrass under different nitrogen doses (0, 33, 66, 100 or 133 kg/ha.year of N and three different plant heights (30, 40 or 50 cm at harvesting time. A split plot scheme in a completely randomized design with three replications was used. The nitrogen (N doses stood for the portions and the haversting heights, for the subportions. The leaf appearance rate of tifton 85 bermudagrass was negatively influenced by plant height at harvesting time, but positively influenced by N dose. Leaf senescence rate and total number of leaves per tiller decreased, whereas leaf elongation rate

  15. Phytoextractor Potential of Cultivated Species in Industrial Area Contaminated by Lead

    Directory of Open Access Journals (Sweden)

    Silvânia Maria de Souza Gomes Nascimento

    Full Text Available ABSTRACT: High growth rate is one of the criteria used for the selection of species to be used in metal phytoextraction programs. This study was carried out to characterize the growth characteristics of sunflower (Helianthus annuus L., castor bean (Ricinus communis L., corn (Zea mays L, and vetiver [Vetiveria zizanioides (L. Nash] grown on a soil contaminated with lead (Pb, with and without pH correction, to improve agronomic practices regarding phytoremediation programs. The experiment was designed as a randomized block with four replications; treatments were arranged in a split-plot arrangement, with the main plot representing the species (sunflower, castor bean, corn, and vetiver, with or without pH correction and soil fertilization, and the split-plot representing harvest periods (60, 90, and 120 days after planting. After variance analysis and mean comparison analysis of the data by the Tukey test (p≤0.05, a significant effect was observed from soil pH correction for vetiver in all of the growth variables evaluated, except for the leaf area index at 120 days after planting (DAP. Castor bean and sunflower plants in soil with high acidity conditions, without pH correction (pH˂4.0, were affected by soil Pb levels. Corn plants benefited from soil pH correction and had improved results for the plant height, diameter, and leaf area variables at 60 and 90 DAP, as well as leaf area index at 60 DAP. There was no increase in these variables between the harvest periods evaluated. Regarding phytoextraction potential, corn and vetiver had the highest Pb translocation to the plant shoots at 90 DAP and were therefore considered the most suitable species for phytoremediation of the area under study. Overall, liming was essential for improving species biomass production for all the species studied in soils with high Pb availability in solution.

  16. Experimental manipulation of leaf litter colonization by aquatic invertebrates in a third order tropical stream.

    Science.gov (United States)

    Uieda, V S; Carvalho, E M

    2015-05-01

    Through a manipulative experiment, the colonization of leaf litter by invertebrates was investigated in two sections of a tropical stream (spatial scale) that differed in function of the canopy cover, one with the presence (closed area) and another without riparian vegetation (open area), during one month of the dry and one of the wet season (temporal scale). The work aimed to verify differences related to four variables: season, canopy cover, leaf type and leaf condition. Litter bags containing arboreal and herbaceous leaves (leaf type variable), non-conditioned and preconditioned (leaf condition variable) were placed at the bottom of the stream in each area (canopy cover variable) and season (dry and wet), and removed after 13-day colonization. The analysis of the remaining litter dry mass per leaf bag emphasizes differences related mainly to seasonality, canopy cover and leaf type, although leaf condition was also important when combined with those three factors. Comparing the abundance of invertebrates per treatment, there was a tendency of high predominance of Chironomidae during the dry season and greater taxa diversity and evenness during the wet season, when the water flow increase could alter the availability of microhabitats for local fauna. Even though canopy cover alone was not a significant source of variation in the abundance of invertebrates, the results showed a tendency of a combined effect of canopy cover with seasonality and leaf condition.

  17. Inheritance of okra leaf type in different genetic backgrounds and its ...

    African Journals Online (AJOL)

    use

    2011-11-21

    Nov 21, 2011 ... trait was studied in three cross combinations involving a common okra leaf parent ... pests with reduced leaf area allowing better air flow and maximum sunlight .... Irrigation both by canal and turbine water was applied to the.

  18. Can biomass responses to warming at plant to ecosystem levels be predicted by leaf-level responses?

    Science.gov (United States)

    Xia, J.; Shao, J.; Zhou, X.; Yan, W.; Lu, M.

    2015-12-01

    Global warming has the profound impacts on terrestrial C processes from leaf to ecosystem scales, potentially feeding back to climate dynamics. Although numerous studies had investigated the effects of warming on C processes from leaf to plant and ecosystem levels, how leaf-level responses to warming scale up to biomass responses at plant, population, and community levels are largely unknown. In this study, we compiled a dataset from 468 papers at 300 experimental sites and synthesized the warming effects on leaf-level parameters, and plant, population and ecosystem biomass. Our results showed that responses of plant biomass to warming mainly resulted from the changed leaf area rather than the altered photosynthetic capacity. The response of ecosystem biomass to warming was weaker than those of leaf area and plant biomass. However, the scaling functions from responses of leaf area to plant biomass to warming were different in diverse forest types, but functions were similar in non-forested biomes. In addition, it is challenging to scale the biomass responses from plant up to ecosystem. These results indicated that leaf area might be the appropriate index for plant biomass response to warming, and the interspecific competition might hamper the scaling of the warming effects on plant and ecosystem levels, suggesting that the acclimation capacity of plant community should be incorporated into land surface models to improve the prediction of climate-C cycle feedback.

  19. Childhood height, adult height, and the risk of prostate cancer

    DEFF Research Database (Denmark)

    Bjerregaard, Lise Geisler; Aarestrup, Julie; Gamborg, Michael

    2016-01-01

    PURPOSE: We previously showed that childhood height is positively associated with prostate cancer risk. It is, however, unknown whether childhood height exerts its effects independently of or through adult height. We investigated whether and to what extent childhood height has a direct effect...... on the risk of prostate cancer apart from adult height. METHODS: We included 5,871 men with height measured at ages 7 and 13 years in the Copenhagen School Health Records Register who also had adult (50-65 years) height measured in the Danish Diet, Cancer and Health study. Prostate cancer status was obtained...... through linkage to the Danish Cancer Registry. Direct and total effects of childhood height on prostate cancer risk were estimated from Cox regressions. RESULTS: From 1996 to 2012, 429 prostate cancers occurred. Child and adult heights were positively and significantly associated with prostate cancer risk...

  20. Use of GLM approach to assess the responses of tropical trees to urban air pollution in relation to leaf functional traits and tree characteristics.

    Science.gov (United States)

    Mukherjee, Arideep; Agrawal, Madhoolika

    2018-05-15

    Responses of urban vegetation to air pollution stress in relation to their tolerance and sensitivity have been extensively studied, however, studies related to air pollution responses based on different leaf functional traits and tree characteristics are limited. In this paper, we have tried to assess combined and individual effects of major air pollutants PM 10 (particulate matter ≤ 10 µm), TSP (total suspended particulate matter), SO 2 (sulphur dioxide), NO 2 (nitrogen dioxide) and O 3 (ozone) on thirteen tropical tree species in relation to fifteen leaf functional traits and different tree characteristics. Stepwise linear regression a general linear modelling approach was used to quantify the pollution response of trees against air pollutants. The study was performed for six successive seasons for two years in three distinct urban areas (traffic, industrial and residential) of Varanasi city in India. At all the study sites, concentrations of air pollutants, specifically PM (particulate matter) and NO 2 were above the specified standards. Distinct variations were recorded in all the fifteen leaf functional traits with pollution load. Caesalpinia sappan was identified as most tolerant species followed by Psidium guajava, Dalbergia sissoo and Albizia lebbeck. Stepwise regression analysis identified maximum response of Eucalyptus citriodora and P. guajava to air pollutants explaining overall 59% and 58% variability's in leaf functional traits, respectively. Among leaf functional traits, maximum effect of air pollutants was observed on non-enzymatic antioxidants followed by photosynthetic pigments and leaf water status. Among the pollutants, PM was identified as the major stress factor followed by O 3 explaining 47% and 33% variability's in leaf functional traits. Tolerance and pollution response were regulated by different tree characteristics such as height, canopy size, leaf from, texture and nature of tree. Outcomes of this study will help in urban forest

  1. Creating high-resolution bare-earth digital elevation models (DEMs) from stereo imagery in an area of densely vegetated deciduous forest using combinations of procedures designed for lidar point cloud filtering

    Science.gov (United States)

    DeWitt, Jessica D.; Warner, Timothy A.; Chirico, Peter G.; Bergstresser, Sarah E.

    2017-01-01

    For areas of the world that do not have access to lidar, fine-scale digital elevation models (DEMs) can be photogrammetrically created using globally available high-spatial resolution stereo satellite imagery. The resultant DEM is best termed a digital surface model (DSM) because it includes heights of surface features. In densely vegetated conditions, this inclusion can limit its usefulness in applications requiring a bare-earth DEM. This study explores the use of techniques designed for filtering lidar point clouds to mitigate the elevation artifacts caused by above ground features, within the context of a case study of Prince William Forest Park, Virginia, USA. The influences of land cover and leaf-on vs. leaf-off conditions are investigated, and the accuracy of the raw photogrammetric DSM extracted from leaf-on imagery was between that of a lidar bare-earth DEM and the Shuttle Radar Topography Mission DEM. Although the filtered leaf-on photogrammetric DEM retains some artifacts of the vegetation canopy and may not be useful for some applications, filtering procedures significantly improved the accuracy of the modeled terrain. The accuracy of the DSM extracted in leaf-off conditions was comparable in most areas to the lidar bare-earth DEM and filtering procedures resulted in accuracy comparable of that to the lidar DEM.

  2. Distribution of leaf characteristics in relation to orientation within the canopy of woody species

    Science.gov (United States)

    Escudero, Alfonso; Fernández, José; Cordero, Angel; Mediavilla, Sonia

    2013-04-01

    Over the last few decades considerable effort has been devoted to research of leaf adaptations to environmental conditions. Many studies have reported strong differences in leaf mass per unit area (LMA) within a single tree depending on the photosynthetic photon flux density (PPFD) incident on different locations in the crown. There are fewer studies, however, of the effects of differences in the timing of light incidence during the day on different crown orientations. Leaves from isolated trees of Quercus suber and Quercus ilex in a cold Mediterranean climate were sampled to analyze differences in LMA and other leaf traits among different crown orientations. Gas-exchange rates, leaf water potentials, leaf temperatures and PPFD incident on leaf surfaces in different crown orientations were also measured throughout one entire summer day for each species. Mean daily PPFD values were similar for the leaves from the eastern and western sides of the canopy. On the western side, PPFD reached maximum values during the afternoon. Maximum leaf temperatures were approximately 10-20% higher on the west side, whereas minimum leaf water potentials were between 10 and 24% higher on the east side. Maximum transpiration rates were approximately 22% greater on the west, because of the greater leaf-to-air vapor pressure deficits (LAVPD). Mean individual leaf area was around 10% larger on the east than on the west side of the trees. In contrast, there were no significant differences in LMA between east and west sides of the crown. Contrary to our expectations, more severe water stress on the west side did not result in increases in LMA, although it was associated with lower individual leaf area. We conclude that increases in LMA measured by other authors along gradients of water stress would be due to differences in light intensity between dry and humid sites.

  3. Leaf water 18 O and 2 H enrichment along vertical canopy profiles in a broadleaved and a conifer forest tree.

    Science.gov (United States)

    Bögelein, Rebekka; Thomas, Frank M; Kahmen, Ansgar

    2017-07-01

    Distinguishing meteorological and plant-mediated drivers of leaf water isotopic enrichment is prerequisite for ecological interpretations of stable hydrogen and oxygen isotopes in plant tissue. We measured input and leaf water δ 2 H and δ 18 O as well as micrometeorological and leaf morpho-physiological variables along a vertical gradient in a mature angiosperm (European beech) and gymnosperm (Douglas fir) tree. We used these variables and different enrichment models to quantify the influence of Péclet and non-steady state effects and of the biophysical drivers on leaf water enrichment. The two-pool model accurately described the diurnal variation of leaf water enrichment. The estimated unenriched water fraction was linked to leaf dry matter content across the canopy heights. Non-steady state effects and reduced stomatal conductance caused a higher enrichment of Douglas fir compared to beech leaf water. A dynamic effect analyses revealed that the light-induced vertical gradients of stomatal conductance and leaf temperature outbalanced each other in their effects on evaporative enrichment. We conclude that neither vertical canopy gradients nor the Péclet effect is important for estimates and interpretation of isotopic leaf water enrichment in hypostomatous trees. Contrarily, species-specific non-steady state effects and leaf temperatures as well as the water vapour isotope composition need careful consideration. © 2017 John Wiley & Sons Ltd.

  4. Respostas do capim-Tifton 85 à aplicação de nitrogênio: cobertura do solo, índice de área foliar e interceptação da radiação solar Responses of the grass Tifton 85 to the application of nitrogen: soil cover, leaf area index, and solar radiation interception

    Directory of Open Access Journals (Sweden)

    A.P.P. Oliveira

    2010-04-01

    Full Text Available Estudaram-se os efeitos da adubação nitrogenada sobre o crescimento do capim Tifton 85 (Cynodon spp. cv. Tifton 85, em termos de ocorrência de plantas e altura do dossel, índice de área foliar e interceptação de radiação fotossinteticamente ativa. O estudo foi desenvolvido sobre um planossolo de ocorrência em campo experimental. Os tratamentos consistiram de níveis de N (0; 150; 300; 450 e 600kg/ha de N-ureia, aplicados em duas vezes, arranjados em um desenho experimental de blocos inteiramente ao acaso, com quatro repetições. As avaliações foram realizadas semanalmente, entre 10 e 40 dias após o corte. Tanto a ocorrência de plantas como a altura do dossel responderam à adubação nitrogenada, conforme um padrão de resposta que variou, sazonalmente, em função das doses de N em interação com as condições ambientais. A expansão da área foliar e a interceptação da radiação fotossinteticamente ativa associada, também foram controladas direta e proporcionalmente pelas doses de N aplicadas, sendo maximizadas a valores de índice de área foliar em torno de 4,0.The effects of nitrogen (N fertilization levels on the growth patterns of the grass Tifton 85 (Cynodon spp. cv.Tifton 85 were studied in terms of occurrence of plants, sward height, leaf area index, and photosynthetically active radiation interception (PARint. The field trial was made in an experimental area. Treatments consisted of N levels (0; 150; 300; 450; and 600kg/ha N-urea, split in two applications arranged in an experimental design of entirely randomized blocks with four repetitions. Plots were weekly sampled, from the day 10th to 40th after cutting. Both occurrence of plants as well as sward height were responsive to nitrogen fertilization according to a response pattern that seasonally varied, depending on N rates and environmental conditions; thus, demonstrating their sensitivity as indicators of growth conditions and canopy structure. Leaf expansion

  5. Light distribution in leaf chambers and its consequences for photosynthesis measurements

    NARCIS (Netherlands)

    Hogewoning, S.W.; Trouwborst, G.; Harbinson, J.; Ieperen, van W.

    2010-01-01

    The impact of a heterogeneous distribution of actinic light within a leaf chamber for photosynthetic measurements by gas exchange on the photosynthesis-irradiance relationship was investigated. High-resolution light distributions were measured over the area of a commercially available clamp-on leaf

  6. Fall from heights: does height really matter?

    Science.gov (United States)

    Alizo, G; Sciarretta, J D; Gibson, S; Muertos, K; Romano, A; Davis, J; Pepe, A

    2018-06-01

    Fall from heights is high energy injuries and constitutes a fraction of all fall-related trauma evaluations while bearing an increase in morbidity and mortality. We hypothesize that despite advancements in trauma care, the overall survivability has not improved in this subset of trauma patients. All adult trauma patients treated after sustaining a fall from heights during a 40-month period were retrospectively reviewed. Admission demographics, clinical data, fall height (ft), injury patterns, ISS, GCS, length of stay, and mortality were reviewed. 116 patients sustained a fall from heights, 90.4% accidental. A mean age of 37± 14.7 years, 86% male, and a fall height of 19 ± 10 ft were encountered. Admission GCS was 13 ± 2 with ISS 10 ± 11. Overall LOS was 6.6 ± 14.9 days and an ICU LOS of 2.8 ± 8.9 days. Falls ≥ 25 ft.(16%) had lower GCS 10.4 ± 5.8, increased ISS 22.6 ± 13.8, a fall height 37.9 ± 13.1 ft and associated increased mortality (p < 0.001). Mortality was 5.2%, a mean distance fallen of 39 ± 22 ft. and an ISS of 31.5 ±16.5. Brain injury was the leading cause of death, 50% with open skull fractures. Level of height fallen is a good predictor of overall outcome and survival. Despite advances in trauma care, death rates remain unchanged. Safety awareness and injury prevention programs are needed to reduce the risk of high-level falls.

  7. Measuring Leaf Area in Soy Plants by HSI Color Model Filtering and Mathematical Morphology

    International Nuclear Information System (INIS)

    Benalcázar, M; Padín, J; Brun, M; Pastore, J; Ballarin, V; Peirone, L; Pereyra, G

    2011-01-01

    There has been lately a significant progress in automating tasks for the agricultural sector. One of the advances is the development of robots, based on computer vision, applied to care and management of soy crops. In this task, digital image processing plays an important role, but must solve some important problems, like the ones associated to the variations in lighting conditions during image acquisition. Such variations influence directly on the brightness level of the images to be processed. In this paper we propose an algorithm to segment and measure automatically the leaf area of soy plants. This information is used by the specialists to evaluate and compare the growth of different soy genotypes. This algorithm, based on color filtering using the HSI model, detects green objects from the image background. The segmentation of leaves (foliage) was made applying Mathematical Morphology. The foliage area was estimated counting the pixels that belong to the segmented leaves. From several experiments, consisting in applying the algorithm to measure the foliage of about fifty plants of various genotypes of soy, at different growth stages, we obtained successful results, despite the high brightness variations and shadows in the processed images.

  8. Measuring Leaf Area in Soy Plants by HSI Color Model Filtering and Mathematical Morphology

    Science.gov (United States)

    Benalcázar, M.; Padín, J.; Brun, M.; Pastore, J.; Ballarin, V.; Peirone, L.; Pereyra, G.

    2011-12-01

    There has been lately a significant progress in automating tasks for the agricultural sector. One of the advances is the development of robots, based on computer vision, applied to care and management of soy crops. In this task, digital image processing plays an important role, but must solve some important problems, like the ones associated to the variations in lighting conditions during image acquisition. Such variations influence directly on the brightness level of the images to be processed. In this paper we propose an algorithm to segment and measure automatically the leaf area of soy plants. This information is used by the specialists to evaluate and compare the growth of different soy genotypes. This algorithm, based on color filtering using the HSI model, detects green objects from the image background. The segmentation of leaves (foliage) was made applying Mathematical Morphology. The foliage area was estimated counting the pixels that belong to the segmented leaves. From several experiments, consisting in applying the algorithm to measure the foliage of about fifty plants of various genotypes of soy, at different growth stages, we obtained successful results, despite the high brightness variations and shadows in the processed images.

  9. Plant growth and leaf-spot severity on eucalypt at different CO2 concentrations in the air

    Directory of Open Access Journals (Sweden)

    Carlos Eduardo Oliveira da Silva

    2014-03-01

    Full Text Available The objective of this work was to evaluate the effects of increased air-CO2 concentration on plant growth and on leaf-spot caused by Cylindrocladium candelabrum in Eucalyptus urophylla. Seedlings were cultivated for 30 days at 451, 645, 904, and 1,147 µmol mol-1 CO2 ; then, they were inoculated with the pathogen and kept under the same conditions for seven days. Increased CO2 concentration increased plant height and shoot dry matter mass, and decreased disease incidence and severity. Stem diameter was not affected by the treatments. Increased concentrations of atmospheric CO2 favorably affect eucalypt growth and reduce leaf-spot severity.

  10. Satellite remote sensing for estimating leaf area index, FPAR and primary production. A literature review

    International Nuclear Information System (INIS)

    Boresjoe Bronge, Laine

    2004-03-01

    Land vegetation is a critical component of several biogeochemical cycles that have become the focus of concerted international research effort. Most ecosystem productivity models, carbon budget models, and global models of climate, hydrology and biogeochemistry require vegetation parameters to calculate land surface photosynthesis, evapotranspiration and net primary production. Therefore, accurate estimates of vegetation parameters are increasingly important in the carbon cycle, the energy balance and in environmental impact assessment studies. The possibility of quantitatively estimating vegetation parameters of importance in this context using satellite data has been explored by numerous papers dealing with the subject. This report gives a summary of the present status and applicability of satellite remote sensing for estimating vegetation productivity by using vegetation index for calculating leaf area index (LAI) and fraction of absorbed photosynthetically active radiation (FPAR). Some possible approaches for use of satellite data for estimating LAI, FPAR and net primary production (NPP) on a local scale are suggested. Recommendations for continued work in the Forsmark and Oskarshamn investigation areas, where vegetation data and NDVI-images based on satellite data have been produced, are also given

  11. Satellite remote sensing for estimating leaf area index, FPAR and primary production. A literature review

    Energy Technology Data Exchange (ETDEWEB)

    Boresjoe Bronge, Laine [SwedPower AB, Stockholm (Sweden)

    2004-03-01

    Land vegetation is a critical component of several biogeochemical cycles that have become the focus of concerted international research effort. Most ecosystem productivity models, carbon budget models, and global models of climate, hydrology and biogeochemistry require vegetation parameters to calculate land surface photosynthesis, evapotranspiration and net primary production. Therefore, accurate estimates of vegetation parameters are increasingly important in the carbon cycle, the energy balance and in environmental impact assessment studies. The possibility of quantitatively estimating vegetation parameters of importance in this context using satellite data has been explored by numerous papers dealing with the subject. This report gives a summary of the present status and applicability of satellite remote sensing for estimating vegetation productivity by using vegetation index for calculating leaf area index (LAI) and fraction of absorbed photosynthetically active radiation (FPAR). Some possible approaches for use of satellite data for estimating LAI, FPAR and net primary production (NPP) on a local scale are suggested. Recommendations for continued work in the Forsmark and Oskarshamn investigation areas, where vegetation data and NDVI-images based on satellite data have been produced, are also given.

  12. The effects of planting media and leaf fertilizers on the growth of jamrud orchid (Dendrobium macrophyllum A. Rich.

    Directory of Open Access Journals (Sweden)

    I GEDE TIRTA

    2006-01-01

    Full Text Available Jamrud orchid (Dendrobium macrophyllum A. Rich. have attractive flowers which make the orchid become one of high economic ornamental plants. The orchid is one of endangered species. Its growth is slow, however appropriate planting media and leaf fertilizers can improve the growth of the orchid. The experiment was conducted from February to June 2003, at “Eka Karya” Bali Botanic Garden. The design used in the experiment was completely randomized block with two treatments and four replicates. The first factor were six kinds of planting medias (roots of C. contaminans, roots of Asplenium nidus, charcoal, roots of C. contaminans+roots of A. nidus, roots of C. contaminans+charcoal and roots of A. nidus+charcoal. The second factor were four kinds of fertilizers (plant catalyst, super bionik, inabio and subur inti persada and one treatment without fertilizer. The results of experiment showed that the interaction between planting medias and leaf fertilizers significantly affected increment of plant height at 12, 14, 16 and 18 weeks after planting, of leaf number at 14, 16 and 18 weeks after planting, of root length, of plant fresh weight and oven dry weight. Treatment of C. contaminans roots and of A. nidus roots combined with inabio fertilizer produced the highest vegetative growth. This treatment increased the total oven dry weight of plant (54.81%, increased the weight of plant (67.48%, of root length (41.63%, of total leaf number (70.73%, of plant height (59.01% and bud number (72.22% compared with treatment without fertilizer in the same media.

  13. Effects of leaf movement on radiation interception in field grown leguminous crops, 1: Peanut (Arachis hypogaea L.)

    International Nuclear Information System (INIS)

    Isoda, A.; Yoshimura, T.; Ishikawa, T.; Nojima, H.; Takasaki, Y.

    1993-01-01

    The effects of leaf movement of peanut on radiation interception were examined. A peanut cultivar (c.v. Nakateyutaka) was planted at three planting densities (20, 30 and 40 cm equidistant spacings). In the treatment plots, the upper layer of the canopy was covered horizontally with a nylon net to restrain the movement of the leaflets. Intercepted radiation of each leaflet was measured by integrated solarimeter films for two consecutive days. It was observed that the leaflets of the upper layer oriented paraheliotropically to the sun rays in midday. Intercepted radiation per unit leaf area and unit ground area of the control were larger in the 20 cm pacing, almost similar in the 30 cm spacing and smaller in the 40 cm spacing as compared with the treatment. The leaf movement of the upper layer of the canopy played a significant role in radiation interception in the 20 cm plot, no discernible effect in the 30 cm plot and a rather adverse role in the 40 cm plot. Leaf area of the 20 cm spacing was concentrated densely at the upper layer. Leaf area of the 30 and 40 cm spacing was larger at the middle layers. It was assumed that effectiveness of the leaf movement of the upper layer would depend mainly on spatial leaf area distribution and density

  14. 76 FR 30060 - Proposed Establishment of the Naches Heights Viticultural Area (2009R-107P)

    Science.gov (United States)

    2011-05-24

    ... acres of wine grape producing vines. Name Evidence The ``Naches Heights'' name applies to an elevated... carved much of the basin geography within the Columbia Valley AVA. The proposed Naches Heights.... Distinguishing Features The petition states that geology, geography, and soils distinguish the proposed...

  15. Specific leaf areas of the tank bromeliad Guzmania monostachia perform distinct functions in response to water shortage.

    Science.gov (United States)

    Freschi, Luciano; Takahashi, Cassia Ayumi; Cambui, Camila Aguetoni; Semprebom, Thais Ribeiro; Cruz, Aline Bertinatto; Mioto, Paulo Tamoso; de Melo Versieux, Leonardo; Calvente, Alice; Latansio-Aidar, Sabrina Ribeiro; Aidar, Marcos Pereira Marinho; Mercier, Helenice

    2010-05-01

    Leaves comprise most of the vegetative body of tank bromeliads and are usually subjected to strong longitudinal gradients. For instance, while the leaf base is in contact with the water accumulated in the tank, the more light-exposed middle and upper leaf sections have no direct access to this water reservoir. Therefore, the present study attempted to investigate whether different leaf portions of Guzmania monostachia, a tank-forming C(3)-CAM bromeliad, play distinct physiological roles in response to water shortage, which is a major abiotic constraint in the epiphytic habitat. Internal and external morphological features, relative water content, pigment composition and the degree of CAM expression were evaluated in basal, middle and apical leaf portions in order to allow the establishment of correlations between the structure and the functional importance of each leaf region. Results indicated that besides marked structural differences, a high level of functional specialization is also present along the leaves of this bromeliad. When the tank water was depleted, the abundant hydrenchyma of basal leaf portions was the main reservoir for maintaining a stable water status in the photosynthetic tissues of the apical region. In contrast, the CAM pathway was intensified specifically in the upper leaf section, which is in agreement with the presence of features more suitable for the occurrence of photosynthesis at this portion. Gas exchange data indicated that internal recycling of respiratory CO(2) accounted for virtually all nighttime acid accumulation, characterizing a typical CAM-idling pathway in the drought-exposed plants. Altogether, these data reveal a remarkable physiological complexity along the leaves of G. monostachia, which might be a key adaptation to the intermittent water supply of the epiphytic niche. Copyright 2009 Elsevier GmbH. All rights reserved.

  16. Evaluation of Indirect Measurement Method of Seasonal Patterns of Leaf Area Index in a High-Density Short Rotation Coppice Culture of Poplar

    Czech Academy of Sciences Publication Activity Database

    Tripathi, Abishek; Fischer, Milan; Orság, Matěj; Marek, Michal V.; Žalud, Zdeněk; Trnka, Miroslav

    2016-01-01

    Roč. 64, č. 2 (2016), s. 549-556 ISSN 1211-8516 R&D Projects: GA MŠk(CZ) LD13030; GA MŠk(CZ) ED1.1.00/02.0073; GA MŠk(CZ) EE2.3.20.0248 Institutional support: RVO:67179843 Keywords : SunScan Plant Canopy Analyzer * litterfall * specific leaf area * poplar clone J-105 Subject RIV: GC - Agronomy

  17. "Breath figures" on leaf surfaces-formation and effects of microscopic leaf wetness.

    Science.gov (United States)

    Burkhardt, Juergen; Hunsche, Mauricio

    2013-01-01

    "Microscopic leaf wetness" means minute amounts of persistent liquid water on leaf surfaces which are invisible to the naked eye. The water is mainly maintained by transpired water vapor condensing onto the leaf surface and to attached leaf surface particles. With an estimated average thickness of less than 1 μm, microscopic leaf wetness is about two orders of magnitude thinner than morning dewfall. The most important physical processes which reduce the saturation vapor pressure and promote condensation are cuticular absorption and the deliquescence of hygroscopic leaf surface particles. Deliquescent salts form highly concentrated solutions. Depending on the type and concentration of the dissolved ions, the physicochemical properties of microscopic leaf wetness can be considerably different from those of pure water. Microscopic leaf wetness can form continuous thin layers on hydrophobic leaf surfaces and in specific cases can act similar to surfactants, enabling a strong potential influence on the foliar exchange of ions. Microscopic leaf wetness can also enhance the dissolution, the emission, and the reaction of specific atmospheric trace gases e.g., ammonia, SO2, or ozone, leading to a strong potential role for microscopic leaf wetness in plant/atmosphere interaction. Due to its difficult detection, there is little knowledge about the occurrence and the properties of microscopic leaf wetness. However, based on the existing evidence and on physicochemical reasoning it can be hypothesized that microscopic leaf wetness occurs on almost any plant worldwide and often permanently, and that it significantly influences the exchange processes of the leaf surface with its neighboring compartments, i.e., the plant interior and the atmosphere. The omission of microscopic water in general leaf wetness concepts has caused far-reaching, misleading conclusions in the past.

  18. The impact of long-term water stress on relative growth rate and morphology of needles and shoots of Metasequoia glyptostroboides seedlings: research toward identifying mechanistic models.

    Science.gov (United States)

    Zhang, Yanxiang; Equiza, Maria Alejandra; Zheng, Quanshui; Tyree, Melvin T

    2011-09-01

    Leaf morphology in the upper canopy of trees tends to be different from that lower down. The effect of long-term water stress on leaf growth and morphology was studied in seedlings of Metasequoia glyptostroboides to understand how tree height might affect leaf morphology in larger trees. Tree height increases water stress on growing leaves through increased hydraulic resistance to water flow and increased gravitational potential, hence we assume that water stress imposed by soil dehydration will have an effect equivalent to stress induced by height. Seedlings were subjected to well-watered and two constant levels of long-term water stress treatments. Drought treatment significantly reduced final needle count, area and mass per area (leaf mass area, LMA) and increased needle density. Needles from water-stressed plants had lower maximum volumetric elastic modulus (ε(max)), osmotic potential at full turgor (Ψ¹⁰⁰(π)) (and at zero turgor (Ψ⁰(π)) (than those from well-watered plants. Palisade and spongy mesophyll cell size and upper epidermal cell size decreased significantly in drought treatments. Needle relative growth rate, needle length and cell sizes were linear functions of the daily average water potential at the time of leaf growth (r² 0.88-0.999). We conclude that water stress alone does mimic the direction and magnitude of changes in leaf morphology observed in tall trees. The results are discussed in terms of various models for leaf growth rate. Copyright © Physiologia Plantarum 2011.

  19. Height and Biomass of Mangroves in Africa from ICEsat/GLAS and SRTM

    Science.gov (United States)

    Fatoyinbo, Temilola E.; Simard, Marc

    2012-01-01

    The accurate quantification of forest 3-D structure is of great importance for studies of the global carbon cycle and biodiversity. These studies are especially relevant in Africa, where deforestation rates are high and the lack of background data is great. Mangrove forests are ecologically significant and it is important to measure mangrove canopy heights and biomass. The objectives of this study are to estimate: 1. The total area, 2. Canopy height distributions and 3. Aboveground biomass of mangrove forests in Africa. To derive mangrove 3-D structure and biomass maps, we used a combination of mangrove maps derived from Landsat ETM+, LiDAR canopy height estimates from ICEsat/GLAS (Ice, Cloud, and land Elevation Satellite/Geoscience Laser Altimeter System) and elevation data from SRTM (Shuttle Radar Topography Mission) for the African continent. More specifically, we extracted mangrove forest areas on the SRTM DEM using Landsat based landcover maps. The LiDAR (Light Detection and Ranging) measurements from the large footprint GLAS sensor were used to derive local estimates of canopy height and calibrate the Interferometric Synthetic Aperture Radar (InSAR) data from SRTM. We then applied allometric equations relating canopy height to biomass in order to estimate above ground biomass (AGB) from the canopy height product. The total mangrove area of Africa was estimated to be 25 960 square kilometers with 83% accuracy. The largest mangrove areas and greatest total biomass was 29 found in Nigeria covering 8 573 km2 with 132 x10(exp 6) Mg AGB. Canopy height across Africa was estimated with an overall root mean square error of 3.55 m. This error also includes the impact of using sensors with different resolutions and geolocation error which make comparison between measurements sensitive to canopy heterogeneities. This study provides the first systematic estimates of mangrove area, height and biomass in Africa. Our results showed that the combination of ICEsat/GLAS and

  20. Predicting tropical plant physiology from leaf and canopy spectroscopy.

    Science.gov (United States)

    Doughty, Christopher E; Asner, Gregory P; Martin, Roberta E

    2011-02-01

    A broad regional understanding of tropical forest leaf photosynthesis has long been a goal for tropical forest ecologists, but it has remained elusive due to difficult canopy access and high species diversity. Here we develop an empirical model to predict sunlit, light-saturated, tropical leaf photosynthesis using leaf and simulated canopy spectra. To develop this model, we used partial least squares (PLS) analysis on three tropical forest datasets (159 species), two in Hawaii and one at the biosphere 2 laboratory (B2L). For each species, we measured light-saturated photosynthesis (A), light and CO(2) saturated photosynthesis (A(max)), respiration (R), leaf transmittance and reflectance spectra (400-2,500 nm), leaf nitrogen, chlorophyll a and b, carotenoids, and leaf mass per area (LMA). The model best predicted A [r(2) = 0.74, root mean square error (RMSE) = 2.9 μmol m(-2) s(-1))] followed by R (r(2) = 0.48), and A(max) (r(2) = 0.47). We combined leaf reflectance and transmittance with a canopy radiative transfer model to simulate top-of-canopy reflectance and found that canopy spectra are a better predictor of A (RMSE = 2.5 ± 0.07 μmol m(-2) s(-1)) than are leaf spectra. The results indicate the potential for this technique to be used with high-fidelity imaging spectrometers to remotely sense tropical forest canopy photosynthesis.

  1. Coordination of Leaf Photosynthesis, Transpiration, and Structural Traits in Rice and Wild Relatives (Genus Oryza).

    Science.gov (United States)

    Giuliani, Rita; Koteyeva, Nuria; Voznesenskaya, Elena; Evans, Marc A; Cousins, Asaph B; Edwards, Gerald E

    2013-07-01

    The genus Oryza, which includes rice (Oryza sativa and Oryza glaberrima) and wild relatives, is a useful genus to study leaf properties in order to identify structural features that control CO(2) access to chloroplasts, photosynthesis, water use efficiency, and drought tolerance. Traits, 26 structural and 17 functional, associated with photosynthesis and transpiration were quantified on 24 accessions (representatives of 17 species and eight genomes). Hypotheses of associations within, and between, structure, photosynthesis, and transpiration were tested. Two main clusters of positively interrelated leaf traits were identified: in the first cluster were structural features, leaf thickness (Thick(leaf)), mesophyll (M) cell surface area exposed to intercellular air space per unit of leaf surface area (S(mes)), and M cell size; a second group included functional traits, net photosynthetic rate, transpiration rate, M conductance to CO(2) diffusion (g(m)), stomatal conductance to gas diffusion (g(s)), and the g(m)/g(s) ratio.While net photosynthetic rate was positively correlated with gm, neither was significantly linked with any individual structural traits. The results suggest that changes in gm depend on covariations of multiple leaf (S(mes)) and M cell (including cell wall thickness) structural traits. There was an inverse relationship between Thick(leaf) and transpiration rate and a significant positive association between Thick(leaf) and leaf transpiration efficiency. Interestingly, high g(m) together with high g(m)/g(s) and a low S(mes)/g(m) ratio (M resistance to CO(2) diffusion per unit of cell surface area exposed to intercellular air space) appear to be ideal for supporting leaf photosynthesis while preserving water; in addition, thick M cell walls may be beneficial for plant drought tolerance.

  2. Spatial-temporal variability of leaf chlorophyll and its relationship with cocoa yield

    Directory of Open Access Journals (Sweden)

    Caique C. Medauar

    Full Text Available ABSTRACT The objective of this study was to evaluate the spatial-temporal variability of leaf chlorophyll index and its relationship with cocoa yield. The experiment was carried out in an experimental area of cocoa production located in Ilhéus, Bahia State, Brazil. Leaf chlorophyll content was measured in September, October, January, February, March and April in the 2014/2015 season, at each sampling point of a regular grid by using a portable chlorophyll meter. Under the same conditions, yield was evaluated and the data were submitted to descriptive statistics and a linear correlation study. Geostatistical analysis was used to determine and quantify the spatial and temporal variability of leaf chlorophyll index and yield. Leaf chlorophyll index varied over the period evaluated, but the months of February, March and April showed no spatial dependence in the study area, indicating absence of temporal stability. Cocoa monthly yield, except in January, presented high spatial variability. Under the conditions of this study, it was not possible to establish a relationship between leaf chlorophyll index and cocoa yield.

  3. Monocot leaves are eaten less than dicot leaves in tropical lowland rain forests: correlations with toughness and leaf presentation

    DEFF Research Database (Denmark)

    Grubb, P.J.; Jackson, R.V.; Barberis, I.M.

    2008-01-01

    . It was hypothesized that (a) losses of leaf area to herbivorous invertebrates are generally greatest during leaf expansion and smaller for monocots than for dicots, and (b) where losses after expansion are appreciable any difference between monocots and dicots then is smaller than that found during expansion. Methods......: At six sites on four continents, estimates were made of lamina area loss from the four most recently mature leaves of focal monocots and of the nearest dicot shoot. Measurements of leaf mass per unit area, and the concentrations of water and nitrogen were made for many of the species. In Panama...... of leaf mass per unit area, or concentrations of water or nitrogen. At only one site was the increase in loss from first to fourth mature leaf significant (also large and the same in monocots and dicots), but the losses sustained during expansion were much smaller in the monocots. In the leaf-cutter ant...

  4. A global boundary-layer height climatology

    Energy Technology Data Exchange (ETDEWEB)

    Dop, H. van; Krol, M.; Holtslag, B. [Inst. for Marine and Atmospheric Research Utrecht, IMAU, Utrecht (Netherlands)

    1997-10-01

    In principle the ABL (atmospheric boundary layer) height can be retrieved from atmospheric global circulation models since they contain algorithms which determine the intensity of the turbulence as a function of height. However, these data are not routinely available, or on a (vertical) resolution which is too crude in view of the application. This justifies the development of a separate algorithm in order to define the ABL. The algorithm should include the generation of turbulence by both shear and buoyancy and should be based on readily available atmospheric parameters. There is obviously a wide application for boundary heights in off-line global and regional chemistry and transport modelling. It is also a much used parameter in air pollution meteorology. In this article we shall present a theory which is based on current insights in ABL dynamics. The theory is applicable over land and sea surfaces in all seasons. The theory is (for various reasons) not valid in mountainous areas. In areas where boundary-layer clouds or deep cumulus convection are present the theory does not apply. However, the same global atmospheric circulation models contain parameterizations for shallow and deep convection from which separate estimates can be obtained for the extent of vertical mixing. (au)

  5. Undecomposed Twigs in the Leaf Litter as Nest-Building Resources for Ants (Hymenoptera: Formicidae in Areas of the Atlantic Forest in the Southeastern Region of Brazil

    Directory of Open Access Journals (Sweden)

    Tae Tanaami Fernandes

    2012-01-01

    Full Text Available In tropical forests, the leaf-litter stratum exhibits one of the greatest abundances of ant species. This diversity is associated with the variety of available locations for nest building. Ant nests can be found in various microhabitats, including tree trunks and fallen twigs in different stages of decomposition. In this study, we aimed to investigate undecomposed twigs as nest-building resources in the leaf litter of dense ombrophilous forest areas in the southeastern region of Brazil. Demographic data concerning the ant colonies, the physical characteristics of the nests, and the population and structural of the forest were observed. Collections were performed manually over four months in closed canopy locations that did not have trails or flooded areas. A total of 294 nests were collected, and 34 ant species were recorded. Pheidole, Camponotus, and Hypoponera were the richest genera observed; these genera were also among the most populous and exhibited the greatest abundance of nests. We found no association between population size and nest diameter. Only tree cover influenced the nest abundance and species richness. Our data indicate that undecomposed twigs may be part of the life cycle of many species and are important for maintaining ant diversity in the leaf litter.

  6. [Effects of simulated warming on the growth, leaf phenology, and leaf traits of Salix eriostachya in sub-alpine timberline ecotone of western Sichuan, China].

    Science.gov (United States)

    Xu, Zhen-feng; Hu, Ting-xing; Zhang, Li; Zhang, Yuan-bin; Xian, Jun-ren; Wang, Kai-yun

    2009-01-01

    By using open-top chamber (OTC), the effects of simulated warming on the growth, leaf phenology, and leaf traits of Salix eriostachya in sub-alpine timberline ecotone of Western Sichuan were studied. The results showed that comparing with the control, the mean air temperature at 1.2 m above the ground throughout S. eriostachya growth season in OTC increased by 2.9 degrees C, while the soil temperature at the depth of 5 cm only increased by 0.4 degrees C. The temperature increase in OTC made S. eriostachya budding advanced and defoliation postponed obviously, and the leaf life-span longer. The leaf and branch growth rates as well as the specific leaf area in OTC increased obviously, whereas the leaf nitrogen concentration decreased significantly. In OTC, the stomata conductance, net photosynthetic rate, photorespiration, and dark respiration rate of S. eriostachya all exhibited an increasing trend. It was suggested that S. eriostachya had stronger capability to adapt to warming, and, under the background of future global climate change, the elevation of S. eriostachya distribution in the timberline ecotone would be likely to ascend.

  7. Edge, height and visibility effects on nest predation by birds and mammals in the Brazilian cerrado

    Science.gov (United States)

    Dodonov, Pavel; Paneczko, Ingrid Toledo; Telles, Marina

    2017-08-01

    Edge influence is one of the main impacts in fragmented landscapes; yet, most of studies on edge influence have focused on high-contrast edges, and the impacts of low-contrast edges and narrow linear openings are less understood. Edge influence often affects bird nest predation, but these effects are not ubiquitous and may depend on characteristics such as nest height and visibility. We performed an experiment on nest predation in a migratory passerine, Elaenia chiriquensis (Lesser Elaenia; Passeriformes: Tyrannidae), in a savanna vegetation of the Brazilian Cerrado biome in South-Eastern Brazil. We used 89 real E. chiriquensis nests, collected during previous reproductive seasons, with two plasticine eggs in each, and randomly distributed them at two locations (edge - up to 20 m from a firebreak edge and interior - approx. 150-350 m from the edge) and two heights (low - 60-175 cm and high - 190-315 cm above ground). We also measured leaf and branch density around each nest. We performed this study on two 15-days campaigns, checking the nests every 2-3 days and removing those with predation marks. We sorted the predation marks into those made by birds, mammals, or unidentified predators, and used generalized linear models to assess the effects of location, height and leaf density on survival time and predator type. Only four nests had not been predated during the experiment; 55 nests were predated by birds, 7 by mammals, and 23 by unidentified predators. Low nests in the interior tended to have larger survival times whereas high nests at the edge tended to be more predated by birds and less predated by mammals. Thus, even a low-contrast (firebreak) edge may significantly increase nest predation, which is also affected by the nest's height, mainly due to predation by birds. These effects may be due to predator movement along the edge as well as to edge-related changes in vegetation structure. We suggest that higher-contrast edges which may also be used as movement

  8. Occurrence of barley leaf disease and control strategies in Denmark

    DEFF Research Database (Denmark)

    Jørgensen, Lise Nistrup; Ørum, Jens Erik; Heick, Thies Marten

    Barley (Hordeum vulgare) is one of the major crops in Denmark and of special importance for malting and for pig feed. In 2016, the crop was grown covering a total area of 700,000 ha; approximately 25% of arable area in Denmark. To ensure high yield of around 60 dt ha-1, disease-tolerant cultivars...... have proven to be quite effective against all leaf diseases, aside from brown rust and mildew. Denmark has a national record system for pesticide usages. All farmers upload their fungicide use by crop, creating a good basis for assessing the differences in use pattern across different regions...... and fungicide treatments are required. Each year, barley cultivars are assessed for susceptibility towards leaf diseases in national observation plots. The most predominant fungal leaf diseases in Denmark are barley scald (Rhynchosporium secalis), net blotch (Pyrenophora teres), brown rust (Puccinia hordei...

  9. Estimation of leaf area for greenhouse cucumber by linear measurements under salinity and grafting Estimativa da área foliar do pepino em ambiente protegido por medidas lineares sob salinidade e enxertia

    Directory of Open Access Journals (Sweden)

    Flávio Favaro Blanco

    2005-08-01

    Full Text Available The measurement of leaf area by linear parameters is a useful tool when plants cannot be destroyed for direct measurement. The objectives of this study were to establish equations to estimate the leaf area of greenhouse-cucumber and to evaluate the effects of salinity and grafting on this estimative. Non-grafted cucumber seedlings, cv. 'Hokushin', were transplanted in a greenhouse and were irrigated with water of different salinities (1.0, 3.2 and 5.0 dS m-1. In the second growing period, the same cultivar was grafted on Cucurbita spp. and the plants were irrigated with water of 1.4, 3.0 and 5.3 dS m-1. Leaves of different sizes were collected from both experiments and leaf area was determined by an integrating area meter. Leaf length (L and width (W were also recorded. An equation for estimating the leaf area from L and W was developed for a given salinity level or grafting condition and estimated well the area of leaves collected in the other treatments. The leaf area (LA of cucumber 'Hokushin' could be estimated using the equation LA = 0.88LW - 4.27, for any grafting and salinity conditions.A determinação da área foliar por medidas lineares é uma ferramenta útil quando as plantas não podem ser destruídas para que a medição direta seja realizada. Os objetivos desse trabalho foram definir equações para a estimativa da área foliar do pepino em ambiente protegido e avaliar os efeitos da salinidade e da a enxertia nessa estimativa. Mudas de pepino, cv. 'Hokushin', não enxertadas, foram transplantadas em um ambiente protegido e irrigadas com água de diferentes salinidades (1,0, 3,2 e 5,0 dS m-1. No segundo período de cultivo, a mesma cultivar foi enxertada sobre Cucurbita spp., sendo as plantas irrigadas com água de 1,4, 3,0 e 5,3 dS m-1. Foram coletadas folhas de diferentes tamanhos dos dois cultivos e dos três tratamentos e a área foliar foi determinada por um medidor de área foliar. O comprimento (C e a largura (L da folha

  10. Crop quality control system: a tool to control the visual quality of pot plants

    NARCIS (Netherlands)

    Dijkshoorn-Dekker, M.W.C.

    2002-01-01

    Key words: quality, growth, model, leaf unfolding rate, internode, plant height, plant width, leaf area, temperature, plant spacing, season, light, development, image processing, grading, neural network, pot plant, Ficus benjamina

  11. Acclimation of photosynthetic capacity to irradiance in tree canopies in relation to leaf nitrogen concentration and leaf mass per unit area

    NARCIS (Netherlands)

    Meir, P.; Kruijt, B.; Broadmeadow, M.; Barbosa, E.; Kull, O.; Carswell, F.; Nobre, A.; Jarvis, P.G.

    2002-01-01

    The observation of acclimation in leaf photosynthetic capacity to differences in growth irradiance has been widely used as support for a hypothesis that enables a simplification of some soil-vegetation-atmosphere transfer (SVAT) photosynthesis models. The acclimation hypothesis requires that

  12. The determination of the mixing height. Current progress and problems

    Energy Technology Data Exchange (ETDEWEB)

    Gryning, S.E.; Beyrich, F.; Batchvarova, E. [eds.

    1997-10-01

    This report contains extended abstracts of presentations given at a EURASAP Workshop on The Determination of the Mixing Height - Current Progress and Problems. The Workshop, initiated from discussions with Peter Builtjes, was held at Risoe National Laboratory 1-3 October 1997 within the framework of EURASAP (European Association for the Sciences of Air Pollution). The specific topics and chairpersons of the Workshop were: Theoretical Considerations (Sven-Erik Gryning), Mixing Height Estimation from Turbulence Measurements and In-Situ Soundings (Douw Steyn), Mixing Height Determination from NWP-Models (Han van Dop), Climatology and Global Aspects (Werner Klug), Mixing Height Determination from Remote Systems (Werner Klug), Verification of Mixing Height Parameterizations and Models (Frank Beyrich), Mixing Height over Complex Terrain (Ekaterina Batchvarova), Internal Boundary Layers: Mixing Height in Coastal Areas and Over Cities (Allen White). The discussion at the end of the Workshop was chaired by Robert Bornstein. (au)

  13. Remote sensing of LAI, chlorophyll and leaf nitrogen pools of crop- and grasslands in five European landscapes

    DEFF Research Database (Denmark)

    Bøgh, Eva; Houborg, R; Bienkowski, J

    2013-01-01

    Leaf nitrogen and leaf surface area influence the exchange of gases between terrestrial ecosystems and the atmosphere, and they play a significant role in the global cycles of carbon, nitrogen and water. Remote sensing data from satellites can be used to estimate leaf area index (LAI), leaf......). Predictabilities of SVIs and REGFLEC simulations generally improved when constrained to single land use categories (wheat, maize, barley, grass) across the European landscapes, reflecting sensitivity to canopy structures. Predictability further improved when constrained to local (10 × 10 km2) landscapes, thereby...

  14. Impact of Vertical Canopy Position on Leaf Spectral Properties and Traits across Multiple Species

    Directory of Open Access Journals (Sweden)

    Tawanda W. Gara

    2018-02-01

    Full Text Available Understanding the vertical pattern of leaf traits across plant canopies provide critical information on plant physiology, ecosystem functioning and structure and vegetation response to climate change. However, the impact of vertical canopy position on leaf spectral properties and subsequently leaf traits across the entire spectrum for multiple species is poorly understood. In this study, we examined the ability of leaf optical properties to track variability in leaf traits across the vertical canopy profile using Partial Least Square Discriminatory Analysis (PLS-DA. Leaf spectral measurements together with leaf traits (nitrogen, carbon, chlorophyll, equivalent water thickness and specific leaf area were studied at three vertical canopy positions along the plant stem: lower, middle and upper. We observed that foliar nitrogen (N, chlorophyll (Cab, carbon (C, and equivalent water thickness (EWT were higher in the upper canopy leaves compared with lower shaded leaves, while specific leaf area (SLA increased from upper to lower canopy leaves. We found that leaf spectral reflectance significantly (P ≤ 0.05 shifted to longer wavelengths in the ‘red edge’ spectrum (685–701 nm in the order of lower > middle > upper for the pooled dataset. We report that spectral bands that are influential in the discrimination of leaf samples into the three groups of canopy position, based on the PLS-DA variable importance projection (VIP score, match with wavelength regions of foliar traits observed to vary across the canopy vertical profile. This observation demonstrated that both leaf traits and leaf reflectance co-vary across the vertical canopy profile in multiple species. We conclude that canopy vertical position has a significant impact on leaf spectral properties of an individual plant’s traits, and this finding holds for multiple species. These findings have important implications on field sampling protocols, upscaling leaf traits to canopy level

  15. Calcium oxalate druses affect leaf optical properties in selenium-treated Fagopyrum tataricum.

    Science.gov (United States)

    Golob, Aleksandra; Stibilj, Vekoslava; Nečemer, Marijan; Kump, Peter; Kreft, Ivan; Hočevar, Anja; Gaberščik, Alenka; Germ, Mateja

    2018-03-01

    Plants of the genus Fagopyrum contain high levels of crystalline calcium oxalate (CaOx) deposits, or druses, that can affect the leaf optical properties. As selenium has been shown to modify the uptake and accumulation of metabolically important elements such as calcium, we hypothesised that the numbers of druses can be altered by selenium treatment, and this would affect the leaf optical properties. Tartary buckwheat (Fagopyrum tataricum Gaertn.) was grown outdoors in an experimental field. At the beginning of flowering, plants were foliarly sprayed with sodium selenate solution at 10 mg selenium L -1 or only with water. Plant morphological, biochemical, physiological and optical properties were examined, along with leaf elemental composition and content. Se spraying did not affect leaf biochemical and functional properties. However, it increased leaf thickness and the contents of Se in the leaves, and decreased the density of calcium oxalate druses in the leaves. Except Se content, Se spraying did not affect contents of other elements in leaves, including total calcium per dry mass of leaf tissue. Redundancy analysis showed that of all parameters tested, only the calcium oxalate druses parameters were significant in explaining the variability of the leaf reflectance and transmittance spectra. The density of CaOx druses positively correlated with the reflectance in the blue, green, yellow and UV-B regions of the spectrum, while the area of CaOx druses per mm 2 of leaf transection area positively correlated with the transmittance in the green and yellow regions of the spectrum. Copyright © 2018 Elsevier B.V. All rights reserved.

  16. Method of determination of the leaf area of the potato cultivar Ágata from linear dimensionsMétodo de determinação da área foliar da cultivar de batata Ágata a partir de dimensões lineares

    Directory of Open Access Journals (Sweden)

    Sidnei Osmar Jadoski

    2012-12-01

    Full Text Available The cultivar Ágata is currently the most widely cultivated in areas of potato production in Brazil, however, are necessaries basic information about its characteristics, especially on the development of the canopy. The work was developed in experimental area of the Unit of Research in Potato Culture and Microclimate for Agriculture, in the department of Agronomy of the UniversidadeEstadual do Centro Oeste (Unicentro, in Guarapuava (PR. The aim was to evaluate the efficiency of the adjustment of different mathematic models to determine the leaf area of potato plants of the cultivar Ágata. The evaluations were performed considering the dimensions of a 100 seeds of several sizes which were randomly collected in the phase of maximum vegetative development of plants. The data obtained were submitted to analysis of regression, being tested six different mathematic models: linear, quadratic, cubic functions and the exponential models from growth, Gauss and Lorentz, using the length and the height of leaves as independent variable to the adjustment in function of the real leaf area measured in laboratory. The variable length and the product of length x width of the leaf were those who presented the best results of adjustments to the models tested. With more emphasis to the length, for which all the models testes present statistical significance. It was concluded that the leaf length is the most appropriated parameter to be used in evaluation of the leaf area of potato plants cultivar Ágata, being the exponential function of growth and Lorentz the less recommended since its lowest adjustment coefficient. A cultivar Ágata é atualmente a mais cultivada nas lavouras de produção de batata no Brasil, contudo, ainda são necessárias informações básicas sobre suas características, especialmente sobre o desenvolvimento do dossel vegetativo.O trabalho foi desenvolvido em área experimental da Unidade de Pesquisa em Bataticultura e Microclima para a

  17. Innovative LIDAR 3D Dynamic Measurement System to estimate fruit-tree leaf area.

    Science.gov (United States)

    Sanz-Cortiella, Ricardo; Llorens-Calveras, Jordi; Escolà, Alexandre; Arnó-Satorra, Jaume; Ribes-Dasi, Manel; Masip-Vilalta, Joan; Camp, Ferran; Gràcia-Aguilá, Felip; Solanelles-Batlle, Francesc; Planas-DeMartí, Santiago; Pallejà-Cabré, Tomàs; Palacin-Roca, Jordi; Gregorio-Lopez, Eduard; Del-Moral-Martínez, Ignacio; Rosell-Polo, Joan R

    2011-01-01

    In this work, a LIDAR-based 3D Dynamic Measurement System is presented and evaluated for the geometric characterization of tree crops. Using this measurement system, trees were scanned from two opposing sides to obtain two three-dimensional point clouds. After registration of the point clouds, a simple and easily obtainable parameter is the number of impacts received by the scanned vegetation. The work in this study is based on the hypothesis of the existence of a linear relationship between the number of impacts of the LIDAR sensor laser beam on the vegetation and the tree leaf area. Tests performed under laboratory conditions using an ornamental tree and, subsequently, in a pear tree orchard demonstrate the correct operation of the measurement system presented in this paper. The results from both the laboratory and field tests confirm the initial hypothesis and the 3D Dynamic Measurement System is validated in field operation. This opens the door to new lines of research centred on the geometric characterization of tree crops in the field of agriculture and, more specifically, in precision fruit growing.

  18. SU-E-T-430: Modeling MLC Leaf End in 2D for Sliding Window IMRT and Arc Therapy

    International Nuclear Information System (INIS)

    Liang, X; Zhu, T

    2014-01-01

    Purpose: To develop a 2D geometric model for MLC accounting for leaf end dose leakage for dynamic IMRT and Rapidarc therapy. Methods: Leaf-end dose leakage is one of the problems for MLC dose calculation and modeling. Dosimetric leaf gap used to model the MLC and to count for leakage in dose calculation, but may not be accurate for smaller leaf gaps. We propose another geometric modeling method to compensate for the MLC round-shape leaf ends dose leakage, and improve the accuracy of dose calculation and dose verification. A triangular function is used to geometrically model the MLC leaf end leakage in the leaf motion direction, and a step function is used in the perpendicular direction. Dose measurements with different leaf gap, different window width, and different window height were conducted, and the results were used to fit the analytical model to get the model parameters. Results: Analytical models have been obtained for stop-and-shoot and dynamic modes for MLC motion. Parameters a=0.4, lw'=5.0 mm for 6X and a=0.54, lw'=4.1 mm for 15x were obtained from the fitting process. The proposed MLC leaf end model improves the dose profile at the two ends of the sliding window opening. This improvement is especially significant for smaller sliding window openings, which are commonly used for highly modulated IMRT plans and arc therapy plans. Conclusion: This work models the MLC round leaf end shape and movement pattern for IMRT dose calculation. The theory, as well as the results in this work provides a useful tool for photon beam IMRT dose calculation and verification

  19. Leaf photosynthetic traits scale with hydraulic conductivity and wood density in Panamanian forest canopy trees.

    Science.gov (United States)

    L.S. Santiago; G. Goldstein; F.C. Meinzer; J.B. Fisher; K. Maehado; D. Woodruff; T. Jones

    2004-01-01

    We investigated how water transport capacity, wood density and wood anatomy were related to leaf photosynthetic traits in two lowland forests in Panama. Leaf-specific hydraulic conductivity (kL) of upper branches was positively correlated with maximum rates of net CO2, assimilation per unit leaf area (Aarea...

  20. Schottky Barrier Height of Pd/MoS2 Contact by Large Area Photoemission Spectroscopy.

    Science.gov (United States)

    Dong, Hong; Gong, Cheng; Addou, Rafik; McDonnell, Stephen; Azcatl, Angelica; Qin, Xiaoye; Wang, Weichao; Wang, Weihua; Hinkle, Christopher L; Wallace, Robert M

    2017-11-08

    MoS 2 , as a model transition metal dichalcogenide, is viewed as a potential channel material in future nanoelectronic and optoelectronic devices. Minimizing the contact resistance of the metal/MoS 2 junction is critical to realizing the potential of MoS 2 -based devices. In this work, the Schottky barrier height (SBH) and the band structure of high work function Pd metal on MoS 2 have been studied by in situ X-ray photoelectron spectroscopy (XPS). The analytical spot diameter of the XPS spectrometer is about 400 μm, and the XPS signal is proportional to the detection area, so the influence of defect-mediated parallel conduction paths on the SBH does not affect the measurement. The charge redistribution by Pd on MoS 2 is detected by XPS characterization, which gives insight into metal contact physics to MoS 2 and suggests that interface engineering is necessary to lower the contact resistance for the future generation electronic applications.

  1. Estimating the Fractional Vegetation Cover from GLASS Leaf Area Index Product

    Directory of Open Access Journals (Sweden)

    Zhiqiang Xiao

    2016-04-01

    Full Text Available The fractional vegetation cover (FCover is an essential biophysical variable and plays a critical role in the carbon cycle studies. Existing FCover products from satellite observations are spatially incomplete and temporally discontinuous, and also inaccurate for some vegetation types to meet the requirements of various applications. In this study, an operational method is proposed to calculate high-quality, accurate FCover from the Global LAnd Surface Satellite (GLASS leaf area index (LAI product to ensure physical consistency between LAI and FCover retrievals. As a result, a global FCover product (denoted by TRAGL were generated from the GLASS LAI product from 2000 to present. With no missing values, the TRAGL FCover product is spatially complete. A comparison of the TRAGL FCover product with the Geoland2/BioPar version 1 (GEOV1 FCover product indicates that these FCover products exhibit similar spatial distribution pattern. However, there were relatively large discrepancies between these FCover products over equatorial rainforests, broadleaf crops in East-central United States, and needleleaf forests in Europe and Siberia. Temporal consistency analysis indicates that TRAGL FCover product has continuous trajectories. Direct validation with ground-based FCover estimates demonstrated that TRAGL FCover values were more accurate (RMSE = 0.0865, and R2 = 0.8848 than GEOV1 (RMSE = 0.1541, and R2 = 0.7621.

  2. Variation in light absorption properties of mentha aquatica L. as a function of leaf form: Implications for plant growth

    DEFF Research Database (Denmark)

    Enriquez, Susana; Jensen, Kaj Sand

    2008-01-01

    To understand the association between leaf form and leaf optical properties, we examined light absorption variations in the leaves of Mentha aquatica L., an amphibious freshwater macrophyte. Specific absorption of leaves of M. aquatica showed a 7.5-fold variation, decreasing as pigment per unit...... area increased. This relationship indicates that dispersive samples, such as leaves, although efficient light traps, can also be affected by the "package effect." Mentha aquatica leaves, by expanding their biomass (increased specific leaf area [SLA]), improve their light absorption efficiency per unit...... of both pigment and leaf biomass. Changes in leaf biomass expansion were mainly a result of changes in leaf density, and as a consequence, leaf density appears to be a better descriptor of light absorption efficiency in M. aquatica leaves than does leaf thickness. Light absorption efficiency per unit...

  3. Crescimento inicial de mudas de erva-mate (Ilex paraguariensis St. Hil. em três classes de solos, na região central do Rio Grande do Sul

    Directory of Open Access Journals (Sweden)

    Fabio Luiz Saidelles Fleig

    2003-01-01

    Full Text Available The experiments were installed in three sites with different soil classes but with similar climatic conditions using different fertilization treatment to study initial plant growth of "Erva-mate". The sites were located on an Entisol and two Alfisols and plantation was perfomed using organic compost, no fertilizer, one half and full dosage of N-P-K recommended. The initial growth was evaluated measuring absolute increment in plant height, total plant leaf area and plant leaf number at four times during the first year. The experiments were installed in September of 1998 and in February 1, 1999 plant survival was measured. Chemical fertilization reduced the final survival, whereas, the use of 1 kg of organic compost and nofertilization at planting induced higher plant stand and initial growth as measured by absolute increment in total plant leaf area and height. There was a significant effect of soil type, where, the higher increment in total leaf area and leaf number was observed at the site with shallow and naturally most fertile soil, the Entisol.

  4. Water tables constrain height recovery of willow on Yellowstone's northern range.

    Science.gov (United States)

    Bilyeu, Danielle M; Cooper, David J; Hobbs, N Thompson

    2008-01-01

    Excessive levels of herbivory may disturb ecosystems in ways that persist even when herbivory is moderated. These persistent changes may complicate efforts to restore ecosystems affected by herbivores. Willow (Salix spp.) communities within the northern range in Yellowstone National Park have been eliminated or degraded in many riparian areas by excessive elk (Cervus elaphus L.) browsing. Elk browsing of riparian willows appears to have diminished following the reintroduction of wolves (Canis lupis L.), but it remains uncertain whether reduced herbivory will restore willow communities. The direct effects of elk browsing on willows have been accompanied by indirect effects from the loss of beaver (Castor canadensis Kuhl) activity, including incision of stream channels, erosion of fine sediments, and lower water tables near streams historically dammed by beaver. In areas where these changes have occurred, lowered water tables may suppress willow height even in the absence of elk browsing. We conducted a factorial field experiment to understand willow responses to browsing and to height of water tables. After four years of protection from elk browsing, willows with ambient water tables averaged only 106 cm in height, with negligible height gain in two of three study species during the last year of the experiment. Willows that were protected from browsing and had artificially elevated water tables averaged 147 cm in height and gained 19 cm in the last year of the experiment. In browsed plots, elevated water tables doubled height gain during a period of slightly reduced browsing pressure. We conclude that water availability mediates the rate of willow height gain and may determine whether willows grow tall enough to escape the browse zone of elk and gain resistance to future elk browsing. Consequently, in areas where long-term beaver absence has resulted in incised stream channels and low water tables, a reduction in elk browsing alone may not be sufficient for recovery

  5. Assimilating Remote Sensing Observations of Leaf Area Index and Soil Moisture for Wheat Yield Estimates: An Observing System Simulation Experiment

    Science.gov (United States)

    Nearing, Grey S.; Crow, Wade T.; Thorp, Kelly R.; Moran, Mary S.; Reichle, Rolf H.; Gupta, Hoshin V.

    2012-01-01

    Observing system simulation experiments were used to investigate ensemble Bayesian state updating data assimilation of observations of leaf area index (LAI) and soil moisture (theta) for the purpose of improving single-season wheat yield estimates with the Decision Support System for Agrotechnology Transfer (DSSAT) CropSim-Ceres model. Assimilation was conducted in an energy-limited environment and a water-limited environment. Modeling uncertainty was prescribed to weather inputs, soil parameters and initial conditions, and cultivar parameters and through perturbations to model state transition equations. The ensemble Kalman filter and the sequential importance resampling filter were tested for the ability to attenuate effects of these types of uncertainty on yield estimates. LAI and theta observations were synthesized according to characteristics of existing remote sensing data, and effects of observation error were tested. Results indicate that the potential for assimilation to improve end-of-season yield estimates is low. Limitations are due to a lack of root zone soil moisture information, error in LAI observations, and a lack of correlation between leaf and grain growth.

  6. Ecophysiological function of leaf 'windows' in Lithops species - 'Living Stones' that grow underground.

    Science.gov (United States)

    Martin, C E; Brandmeyer, E A; Ross, R D

    2013-01-01

    Leaf temperatures were lower when light entry at the leaf tip window was prevented through covering the window with reflective tape, relative to leaf temperatures of plants with leaf tip windows covered with transparent tape. This was true when leaf temperatures were measured with an infrared thermometer, but not with a fine-wire thermocouple. Leaf tip windows of Lithops growing in high-rainfall regions of southern Africa were larger than the windows of plants (numerous individuals of 17 species) growing in areas with less rainfall and, thus, more annual insolation. The results of this study indicate that leaf tip windows of desert plants with an underground growth habit can allow entry of supra-optimal levels of radiant energy, thus most likely inhibiting photosynthetic activity. Consequently, the size of the leaf tip windows correlates inversely with habitat solar irradiance, minimising the probability of photoinhibition, while maximising the absorption of irradiance in cloudy, high-rainfall regions. © 2012 German Botanical Society and The Royal Botanical Society of the Netherlands.

  7. Leaf micro-environment influence the altered foliar phenotype of columnar apple (Malus x domestica Borkh.) trees

    DEFF Research Database (Denmark)

    Talwara, Susheela; Grout, Brian William Wilson; Toldam-Andersen, Torben Bo

    2015-01-01

    in the phenotype of the leaves in the leaf clusters that subtend the fruits of CATs, compared to their standard counterparts. This initial investigation considers standard and columnar trees at different levels of genetic relatedness and records significant increases in leaf area, leaf mass per unit area......Columnar apple trees (CATs) have radically-altered architecture (significantly shorter internodes and lateral branches) when compared to standard apple trees, attributed to a mutation of the Co gene involved in apical dominance. These changes in architecture have been associated with changes......, chlorophyll content and competitive shading in the fruiting leaf clusters of columnar cultivars. Additionally, significant increases in intercepted light have been shown to be associated with the columnar structure, and carbon fixation is also increased. We propose that leaf micro-environment of columnar...

  8. Plant functional traits predict green roof ecosystem services.

    Science.gov (United States)

    Lundholm, Jeremy; Tran, Stephanie; Gebert, Luke

    2015-02-17

    Plants make important contributions to the services provided by engineered ecosystems such as green roofs. Ecologists use plant species traits as generic predictors of geographical distribution, interactions with other species, and ecosystem functioning, but this approach has been little used to optimize engineered ecosystems. Four plant species traits (height, individual leaf area, specific leaf area, and leaf dry matter content) were evaluated as predictors of ecosystem properties and services in a modular green roof system planted with 21 species. Six indicators of ecosystem services, incorporating thermal, hydrological, water quality, and carbon sequestration functions, were predicted by the four plant traits directly or indirectly via their effects on aggregate ecosystem properties, including canopy density and albedo. Species average height and specific leaf area were the most useful traits, predicting several services via effects on canopy density or growth rate. This study demonstrates that easily measured plant traits can be used to select species to optimize green roof performance across multiple key services.

  9. Small-scale variations in leaf shape under anthropogenic disturbance in dioecious forest forb mercurialis perennis: A geometric morphometric examination

    Directory of Open Access Journals (Sweden)

    Vujić Vukica

    2016-01-01

    Full Text Available Plants are exposed to increasing levels of diverse human activities that have profound effects on their overall morphology and, specifically, on leaf morphology. Anthropogenic disturbances in urban and suburban forest recreational sites are attracting growing research interest. To explore the persisting recreational impact on leaf shape and size, we conducted a field study on the dioecious forb Mercurialis perennis L. (Euphorbiaceae, typical for undisturbed understory communities. We selected adjacent sites in a suburban forest, which experience contrasting regimes of disturbance by human trampling under otherwise concordant natural conditions. Patterns of leaf shape and size variation and putative sex-specific response to disturbance were analyzed using a geometric morphometric approach. In addition to leaf-level data, plant height, internode and leaf number were analyzed to explore the same response at the whole-plant level. The results show significant variations associated with disturbance at both levels: plants growing under a heavy disturbance regime had shorter stems with a greater number of wider and shorter leaves. Significant differences between sites were also found for leaf size, with larger leaves observed in an undisturbed site. The effects of sex and sex x site interaction on leaf size and shape were nonsignificant, pointing to the absence of sexual dimorphism and sex-specific response to disturbance. Contrary to leaf shape and size, all three analyzed shoot traits showed highly significant sexual dimorphism, with male plants being higher and having higher leaf and internode count. [Projekat Ministarstva nauke Republike Srbije, br. 173025

  10. Concordant preferences for actual height and facial cues to height

    OpenAIRE

    Re, Daniel Edward; Perrett, David I.

    2012-01-01

    Physical height has a well-documented effect on human mate preferences. In general, both sexes prefer opposite-sex romantic relationships in which the man is taller than the woman, while individual preferences for height are affected by a person’s own height. Research in human mate choice has demonstrated that attraction to facial characteristics, such as facial adiposity, may reflect references for body characteristics. Here, we tested preferences for facial cues to height. In general, incre...

  11. Variation of boundary-layer wind spectra with height

    DEFF Research Database (Denmark)

    Larsén, Xiaoli Guo; Petersen, Erik L.; Larsen, Søren Ejling

    2018-01-01

    This study revisits the height dependence of the wind speed power spectrum from the synoptic scale to the spectral gap. Measurements from cup anemometers and sonics at heights of 15 m to 244 m are used. The measurements are from one land site, one coastal land‐based site and three offshore sites...... the atmospheric tide. The second finding regards the height dependence of the general spectrum. We describe the dependence through a so‐called effective roughness, which is calculated from wind spectra and represents the energy removal at different frequencies, and thus surface conditions in the footprint areas....... The generalizable spectral properties of winds presented herein may prove useful for validating numerical models....

  12. The Effect of Foliar Application of Ferrous sulfate on Production of Multi-capsule per Leaf node in Sesame Plant (Sesamum indicum L. under Field Condition

    Directory of Open Access Journals (Sweden)

    S. F Fazeli Kakhki

    2017-06-01

    Full Text Available Introduction Sesame (Sesamum indicum L. belongs to pedaliaceae family and is one of the most important oilseed crops that will grow in relatively dry region. Each leaf node will have one capsule but in some sesame ecotypes the number of capsule per leaf node can increase (Langham and Wiemers, 2002. Factors such as agricultural operations, growing substrates and variety affect yield and harvest index. Iron is one of the main micronutrient that can affect growth and development (Miller et al., 1982. This research was conducted to study the effects of leaf nutrition on the number of capsules per leaf node of sesame. Materials and Methods In order to study the effects of leaf nutrition on the number of capsules per leaf node in sesame plant (sesamum indicum L., an experimental was conducted at Khorasan Razavi Agricultural Education Center, Iran, 1n 2014. This study was done as a factorial arrangement with two factors, kind of seed and the concentrations of ferrous sulphate on the basis completely randomized block design with three replications. The first factor was two seed types (one type has been taken from leaf node with three capsules and the other seeds were taken from leaf node with one capsule in sesame plant. The second factor was three concentration of ferrous sulphate (0, 50 and 100 ppm that applied in %50 vegetation, flowering and capsule phase. Morphological and yield components traits were measured. Data were analyzed with MSTAT-C software and means comparison were done with Duncan multiple range test in 5 percent probability. Results and Discussion The first factor (type of seed showed significant difference on height, length of branch, fresh and dry shoot weight, dry weight of capsule, number capsules per plant, 1000 seed weight and seed weight per plant. Plants growing from the seeds with three capsules in leaf node (Cap2 had the maximum height with 88.2 cm that was 20% more than plants that growth from seeds with one capsule in leaf

  13. Application of Hyperspectral Vegetation Indices to Detect Variations in High Leaf Area Index Temperate Shrub Thicket Canopies

    Science.gov (United States)

    2011-01-01

    litter was collected after leaf fall in November 2008. Litter was dried at 70 °C for 4–5 days, separated into leaf, woody and reproductive (i.e. fruits ...115 (2011) 514–523 Davis, C., Bowles, J., Leathers , R., Korwan, D., Downes, T. V., Snyder, W., et al. (2002). Ocean PHILLS hyperspectral imager

  14. Simulation of three-phase flow and lance height effect on the cavity shape

    Science.gov (United States)

    Dong, Kai; Zhu, Rong; Gao, Wei; Liu, Fu-hai

    2014-06-01

    A three-dimensional computational fluid dynamics (CFD) model was developed to simulate a 150-t top-blown converter. The effect of different lance heights on the cavity shape was investigated using the volume of fluid (VOF) method. Numerical simulation results can reflect the actual molten bath surface waves impinged by the supersonic oxygen jets. With increasing lance height, the cavity depth decreases, and the cavity area, varying like a parabola, increases and then decreases. The cavity area maximizes at the lance height of 1.3 m. Under the three different lance heights simulated in this study, all of the largest impact velocities at the molten bath surface are between 50 m/s and 100 m/s.

  15. Confocal Microscopy for Process Monitoring and Wide-Area Height Determination of Vertically-Aligned Carbon Nanotube Forests

    Directory of Open Access Journals (Sweden)

    Markus Piwko

    2015-08-01

    Full Text Available Confocal microscopy is introduced as a new and generally applicable method for the characterization of the vertically-aligned carbon nanotubes (VACNT forest height. With this technique process control is significantly intensified. The topography of the substrate and VACNT can be mapped with a height resolution down to 15 nm. The advantages of confocal microscopy, compared to scanning electron microscopy (SEM, are demonstrated by investigating the growth kinetics of VACNT using Al2O3 buffer layers with varying thicknesses. A process optimization using confocal microscopy for fast VACNT forest height evaluation is presented.

  16. Zinc deficiency in field-grown pecan trees: changes in leaf nutrient concentrations and structure.

    Science.gov (United States)

    Ojeda-Barrios, Dámaris; Abadía, Javier; Lombardini, Leonardo; Abadía, Anunciación; Vázquez, Saúl

    2012-06-01

    Zinc (Zn) deficiency is a typical nutritional disorder in pecan trees [Carya illinoinensis (Wangenh.) C. Koch] grown under field conditions in calcareous soils in North America, including northern Mexico and south-western United States. The aim of this study was to assess the morphological and nutritional changes in pecan leaves affected by Zn deficiency as well as the Zn distribution within leaves. Zinc deficiency led to decreases in leaf chlorophyll concentrations, leaf area and trunk cross-sectional area. Zinc deficiency increased significantly the leaf concentrations of K and Ca, and decreased the leaf concentrations of Zn, Fe, Mn and Cu. All nutrient values found in Zn-deficient leaves were within the sufficiency ranges, with the only exception of Zn, which was approximately 44, 11 and 9 µg g(-1) dry weight in Zn-sufficient, moderately and markedly Zn-deficient leaves, respectively. Zinc deficiency led to decreases in leaf thickness, mainly due to a reduction in the thickness of the palisade parenchyma, as well as to increases in stomatal density and size. The localisation of Zn was determined using the fluorophore Zinpyr-1 and ratio-imaging technique. Zinc was mainly localised in the palisade mesophyll area in Zn-sufficient leaves, whereas no signal could be obtained in Zn-deficient leaves. The effects of Zn deficiency on the leaf characteristics of pecan trees include not only decreases in leaf chlorophyll and Zn concentrations, but also a reduction in the thickness of the palisade parenchyma, an increase in stomatal density and pore size and the practical disappearance of Zn leaf pools. These characteristics must be taken into account to design strategies to correct Zn deficiency in pecan tree in the field. Copyright © 2012 Society of Chemical Industry.

  17. Leaf size and leaf display of thirty-eight tropical tree species

    NARCIS (Netherlands)

    Poorter, L.; Rozendaal, D.M.A.

    2008-01-01

    Trees forage for light through optimal leaf display. Effective leaf display is determined by metamer traits (i.e., the internode, petiole, and corresponding leaf), and thus these traits strongly co-determine carbon gain and as a result competitive advantage in a light-limited environment. We

  18. Evaluation of Methane from Sisal Leaf Residue and Palash Leaf Litter

    Science.gov (United States)

    Arisutha, S.; Baredar, P.; Deshpande, D. M.; Suresh, S.

    2014-12-01

    The aim of this study is to evaluate methane production from sisal leaf residue and palash leaf litter mixed with different bulky materials such as vegetable market waste, hostel kitchen waste and digested biogas slurry in a laboratory scale anaerobic reactor. The mixture was prepared with 1:1 proportion. Maximum methane content of 320 ml/day was observed in the case of sisal leaf residue mixed with vegetable market waste as the feed. Methane content was minimum (47 ml/day), when palash leaf litter was used as feed. This was due to the increased content of lignin and polyphenol in the feedstock which were of complex structure and did not get degraded directly by microorganisms. Sisal leaf residue mixtures also showed highest content of volatile fatty acids (VFAs) as compared to palash leaf litter mixtures. It was observed that VFA concentration in the digester first increased, reached maximum (when pH was minimum) and then decreased.

  19. The relationship between reference canopy conductance and simplified hydraulic architecture

    Science.gov (United States)

    Novick, Kimberly; Oren, Ram; Stoy, Paul; Juang, Jehn-Yih; Siqueira, Mario; Katul, Gabriel

    2009-06-01

    Terrestrial ecosystems are dominated by vascular plants that form a mosaic of hydraulic conduits to water movement from the soil to the atmosphere. Together with canopy leaf area, canopy stomatal conductance regulates plant water use and thereby photosynthesis and growth. Although stomatal conductance is coordinated with plant hydraulic conductance, governing relationships across species has not yet been formulated at a practical level that can be employed in large-scale models. Here, combinations of published conductance measurements obtained with several methodologies across boreal to tropical climates were used to explore relationships between canopy conductance rates and hydraulic constraints. A parsimonious hydraulic model requiring sapwood-to-leaf area ratio and canopy height generated acceptable agreement with measurements across a range of biomes (r2=0.75). The results suggest that, at long time scales, the functional convergence among ecosystems in the relationship between water-use and hydraulic architecture eclipses inter-specific variation in physiology and anatomy of the transport system. Prognostic applicability of this model requires independent knowledge of sapwood-to-leaf area. In this study, we did not find a strong relationship between sapwood-to-leaf area and physical or climatic variables that are readily determinable at coarse scales, though the results suggest that climate may have a mediating influence on the relationship between sapwood-to-leaf area and height. Within temperate forests, canopy height alone explained a large amount of the variance in reference canopy conductance (r2=0.68) and this relationship may be more immediately applicable in the terrestrial ecosystem models.

  20. Love and fear of heights: the pathophysiology and psychology of height imbalance.

    Science.gov (United States)

    Salassa, John R; Zapala, David A

    2009-01-01

    Individual psychological responses to heights vary on a continuum from acrophobia to height intolerance, height tolerance, and height enjoyment. This paper reviews the English literature and summarizes the physiologic and psychological factors that generate different responses to heights while standing still in a static or motionless environment. Perceptual cues to height arise from vision. Normal postural sway of 2 cm for peripheral objects within 3 m increases as eye-object distance increases. Postural sway >10 cm can result in a fall. A minimum of 20 minutes of peripheral retinal arc is required to detect motion. Trigonometry dictates that a 20-minute peripheral retinal arch can no longer be achieved in a standing position at an eye-object distance of >20 m. At this distance, visual cues conflict with somatosensory and vestibular inputs, resulting in variable degrees of imbalance. Co-occurring deficits in the visual, vestibular, and somatosensory systems can significantly increase height imbalance. An individual's psychological makeup, influenced by learned and genetic factors, can influence reactions to height imbalance. Enhancing peripheral vision and vestibular, proprioceptive, and haptic functions may improve height imbalance. Psychotherapy may improve the troubling subjective sensations to heights.

  1. ‘Breath figures’ on leaf surfaces – formation and effects of microscopic leaf wetness

    Directory of Open Access Journals (Sweden)

    Jürgen eBurkhardt

    2013-10-01

    Full Text Available ‘Microscopic leaf wetness’ means minute amounts of persistent liquid water on leaf surfaces which are invisible to the naked eye. The water is mainly maintained by transpired water vapor condensing onto the leaf surface and to attached leaf surface particles. With an estimated average thickness of less than 1 µm, microscopic leaf wetness it is about 2 orders of magnitude thinner than morning dewfall. The most important physical processes which reduce the saturation vapor pressure and promote condensation are cuticular absorption and the deliquescence of hygroscopic leaf surface particles. Deliquescent salts form highly concentrated solutions. Depending on the amount and concentration of the dissolved ions, the physicochemical properties of microscopic leaf wetness can be considerably different from those of pure water. Microscopic leaf wetness can form continuous thin layers on hydrophobic leaf surfaces and in specific cases can act similar to surfactants, enabling a strong potential influence on the foliar exchange of ions. Microscopic leaf wetness can also enhance the dissolution, the emission, and the reaction of specific atmospheric trace gases e.g. ammonia, SO2, or ozone, leading to a strong potential role for microscopic leaf wetness in plant/atmosphere interaction. Due to its difficult detection, there is little knowledge about the occurrence and the properties of microscopic leaf wetness. However, based on the existing evidence and on physicochemical reasoning it can be hypothesized that microscopic leaf wetness occurs on almost any plant worldwide and often permanently, and that it significantly influences the exchange processes of the leaf surface with its neighboring compartments, i.e., the plant interior and the atmosphere. The omission of microscopic water in general leaf wetness concepts has caused far-reaching, misleading conclusions in the past.

  2. [Impacts of drought stress on the growth and development and grain yield of spring maize in Northeast China].

    Science.gov (United States)

    Ji, Rui-Peng; Che, Yu-Sheng; Zhu, Yong-Ning; Liang, Tao; Feng, Rui; Yu, Wen-Ying; Zhang, Yu-Shu

    2012-11-01

    Taking spring maize variety Danyu-39 as test object, an experiment was conducted in a large-scale agricultural water controlling experimental field to study the impacts of drought stress at three key growth stages, i. e. , 3-leaf-jointing, jointing-silking, and silking-milk ripe, on the growth and development and grain yield of spring maize in Northeast China. Two treatments were installed, including moderate drought stress (MS) and re-watering to suitable water (CK). Compared with CK, the MS at 3-leaf-jointing stage postponed the whole growth period of Danyu-39 by 13 d, and the plant height and leaf area at jointing stage were decreased by 29.8% and 41.2%, respectively. After re-watering, the plant height and grain yield recovered obviously, and the differences in ear characteristics and final yield were insignificant. The MS at jointing-silking stage shortened the whole growth period by 7 d, the plant height and leaf area at silking stage were decreased by 18.6% and 14.1%, respectively, the ear length, grain number per ear, ear dry mass, and grain mass per ear decreased by 6.9%, 19.1%, 28.1%, and 29.4%, respectively, and the blank stem rate increased by 13.3%. When the maize suffered from moderate drought stress at silking-milk ripe stage, the whole growth period was shortened by 15 d, the plant height and leaf area at milk ripe stage were decreased by 2.3% and 37.3%, respectively, the ear length, grain number per ear, ear dry mass, and grain mass per ear decreased by 9.2%, 24.1%, 30.8%, and 27.9%, respectively, and the blank stem rate increased by 24.5%. After re-watering at the latter two stages, the recovery of plant height was little, and the grain yield decreased significantly.

  3. Leaf Area Index Estimation Using Chinese GF-1 Wide Field View Data in an Agriculture Region.

    Science.gov (United States)

    Wei, Xiangqin; Gu, Xingfa; Meng, Qingyan; Yu, Tao; Zhou, Xiang; Wei, Zheng; Jia, Kun; Wang, Chunmei

    2017-07-08

    Leaf area index (LAI) is an important vegetation parameter that characterizes leaf density and canopy structure, and plays an important role in global change study, land surface process simulation and agriculture monitoring. The wide field view (WFV) sensor on board the Chinese GF-1 satellite can acquire multi-spectral data with decametric spatial resolution, high temporal resolution and wide coverage, which are valuable data sources for dynamic monitoring of LAI. Therefore, an automatic LAI estimation algorithm for GF-1 WFV data was developed based on the radiative transfer model and LAI estimation accuracy of the developed algorithm was assessed in an agriculture region with maize as the dominated crop type. The radiative transfer model was firstly used to simulate the physical relationship between canopy reflectance and LAI under different soil and vegetation conditions, and then the training sample dataset was formed. Then, neural networks (NNs) were used to develop the LAI estimation algorithm using the training sample dataset. Green, red and near-infrared band reflectances of GF-1 WFV data were used as the input variables of the NNs, as well as the corresponding LAI was the output variable. The validation results using field LAI measurements in the agriculture region indicated that the LAI estimation algorithm could achieve satisfactory results (such as R² = 0.818, RMSE = 0.50). In addition, the developed LAI estimation algorithm had potential to operationally generate LAI datasets using GF-1 WFV land surface reflectance data, which could provide high spatial and temporal resolution LAI data for agriculture, ecosystem and environmental management researches.

  4. Testing the Suitability of a Terrestrial 2D LiDAR Scanner for Canopy Characterization of Greenhouse Tomato Crops

    Directory of Open Access Journals (Sweden)

    Jordi Llop

    2016-09-01

    Full Text Available Canopy characterization is essential for pesticide dosage adjustment according to vegetation volume and density. It is especially important for fresh exportable vegetables like greenhouse tomatoes. These plants are thin and tall and are planted in pairs, which makes their characterization with electronic methods difficult. Therefore, the accuracy of the terrestrial 2D LiDAR sensor is evaluated for determining canopy parameters related to volume and density and established useful correlations between manual and electronic parameters for leaf area estimation. Experiments were performed in three commercial tomato greenhouses with a paired plantation system. In the electronic characterization, a LiDAR sensor scanned the plant pairs from both sides. The canopy height, canopy width, canopy volume, and leaf area were obtained. From these, other important parameters were calculated, like the tree row volume, leaf wall area, leaf area index, and leaf area density. Manual measurements were found to overestimate the parameters compared with the LiDAR sensor. The canopy volume estimated with the scanner was found to be reliable for estimating the canopy height, volume, and density. Moreover, the LiDAR scanner could assess the high variability in canopy density along rows and hence is an important tool for generating canopy maps.

  5. Variability of the Mixed-Layer Height Over Mexico City

    Science.gov (United States)

    García-Franco, J. L.; Stremme, W.; Bezanilla, A.; Ruiz-Angulo, A.; Grutter, M.

    2018-02-01

    The diurnal and seasonal variability of the mixed-layer height in urban areas has implications for ground-level air pollution and the meteorological conditions. Measurements of the backscatter of light pulses with a commercial lidar system were performed for a continuous period of almost six years between 2011 and 2016 in the southern part of Mexico City. The profiles were temporally and vertically smoothed, clouds were filtered out, and the mixed-layer height was determined with an ad hoc treatment of both the filtered and unfiltered profiles. The results are in agreement when compared with values of mixed-layer height reconstructed from, (i) radiosonde data, and (ii) surface and vertical column densities of a trace gas. The daily maxima of the mean mixed-layer height reach values > 3 km above ground level in the months of March-April, and are clearly lower (pollution episodes and the height of the mixed layer. The growth rate of the convective mixed-layer height has a seasonal behaviour, which is characterized together with the mixed-layer-height anomalies. A clear residual layer is evident from the backscattered signals recorded in days with specific atmospheric conditions, but also from the cloud-filtered mean diurnal profiles. The occasional presence of a residual layer results in an overestimation of the reported mixed-layer height during the night and early morning hours.

  6. Automated estimation of leaf distribution for individual trees based on TLS point clouds

    Science.gov (United States)

    Koma, Zsófia; Rutzinger, Martin; Bremer, Magnus

    2017-04-01

    Light Detection and Ranging (LiDAR) especially the ground based LiDAR (Terrestrial Laser Scanning - TLS) is an operational used and widely available measurement tool supporting forest inventory updating and research in forest ecology. High resolution point clouds from TLS already represent single leaves which can be used for a more precise estimation of Leaf Area Index (LAI) and for higher accurate biomass estimation. However, currently the methodology for extracting single leafs from the unclassified point clouds for individual trees is still missing. The aim of this study is to present a novel segmentation approach in order to extract single leaves and derive features related to leaf morphology (such as area, slope, length and width) of each single leaf from TLS point cloud data. For the study two exemplary single trees were scanned in leaf-on condition on the university campus of Innsbruck during calm wind conditions. A northern red oak (Quercus rubra) was scanned by a discrete return recording Optech ILRIS-3D TLS scanner and a tulip tree (Liliodendron tulpifera) with Riegl VZ-6000 scanner. During the scanning campaign a reference dataset was measured parallel to scanning. In this case 230 leaves were randomly collected around the lower branches of the tree and photos were taken. The developed workflow steps were the following: in the first step normal vectors and eigenvalues were calculated based on the user specified neighborhood. Then using the direction of the largest eigenvalue outliers i.e. ghost points were removed. After that region growing segmentation based on the curvature and angles between normal vectors was applied on the filtered point cloud. On each segment a RANSAC plane fitting algorithm was applied in order to extract the segment based normal vectors. Using the related features of the calculated segments the stem and branches were labeled as non-leaf and other segments were classified as leaf. The validation of the different segmentation

  7. Validation of an assay for quantification of free normetanephrine, metanephrine and methoxytyramine in plasma by high performance liquid chromatography with coulometric detection: Comparison of peak-area vs. peak-height measurements.

    Science.gov (United States)

    Nieć, Dawid; Kunicki, Paweł K

    2015-10-01

    Measurements of plasma concentrations of free normetanephrine (NMN), metanephrine (MN) and methoxytyramine (MTY) constitute the most diagnostically accurate screening test for pheochromocytomas and paragangliomas. The aim of this article is to present the results from a validation of an analytical method utilizing high performance liquid chromatography with coulometric detection (HPLC-CD) for quantifying plasma free NMN, MN and MTY. Additionally, peak integration by height and area and the use of one calibration curve for all batches or individual calibration curve for each batch of samples was explored as to determine the optimal approach with regard to accuracy and precision. The method was validated using charcoal stripped plasma spiked with solutions of NMN, MN, MTY and internal standard (4-hydroxy-3-methoxybenzylamine) with the exception of selectivity which was evaluated by analysis of real plasma samples. Calibration curve performance, accuracy, precision and recovery were determined following both peak-area and peak-height measurements and the obtained results were compared. The most accurate and precise method of calibration was evaluated by analyzing quality control samples at three concentration levels in 30 analytical runs. The detector response was linear over the entire tested concentration range from 10 to 2000pg/mL with R(2)≥0.9988. The LLOQ was 10pg/mL for each analyte of interest. To improve accuracy for measurements at low concentrations, a weighted (1/amount) linear regression model was employed, which resulted in inaccuracies of -2.48 to 9.78% and 0.22 to 7.81% following peak-area and peak-height integration, respectively. The imprecisions ranged from 1.07 to 15.45% and from 0.70 to 11.65% for peak-area and peak-height measurements, respectively. The optimal approach to calibration was the one utilizing an individual calibration curve for each batch of samples and peak-height measurements. It was characterized by inaccuracies ranging from -3

  8. Modelling plant responses to elevated CO2: how important is leaf area index?

    NARCIS (Netherlands)

    Ewert, F.

    2004-01-01

    Background and Aims The problem of increasing CO2 concentration [CO2] and associated climate change has [CO2] on plants. While variation in growth and productivity is generated much interest in modelling effects of closely related to the amount of intercepted radiation, largely determined by leaf

  9. Spring leaf flush in aspen (Populus tremuloides) clones is altered by long-term growth at elevated carbon dioxide and elevated ozone concentration

    International Nuclear Information System (INIS)

    McGrath, Justin M.; Karnosky, David F.; Ainsworth, Elizabeth A.

    2010-01-01

    Early spring leaf out is important to the success of deciduous trees competing for light and space in dense forest plantation canopies. In this study, we investigated spring leaf flush and how long-term growth at elevated carbon dioxide concentration ([CO 2 ]) and elevated ozone concentration ([O 3 ]) altered leaf area index development in a closed Populus tremuloides (aspen) canopy. This work was done at the Aspen FACE experiment where aspen clones have been grown since 1997 in conditions simulating the [CO 2 ] and [O 3 ] predicted for ∼2050. The responses of two clones were compared during the first month of spring leaf out when CO 2 fumigation had begun, but O 3 fumigation had not. Trees in elevated [CO 2 ] plots showed a stimulation of leaf area index (36%), while trees in elevated [O 3 ] plots had lower leaf area index (-20%). While individual leaf area was not significantly affected by elevated [CO 2 ], the photosynthetic operating efficiency of aspen leaves was significantly improved (51%). There were no significant differences in the way that the two aspen clones responded to elevated [CO 2 ]; however, the two clones responded differently to long-term growth at elevated [O 3 ]. The O 3 -sensitive clone, 42E, had reduced individual leaf area when grown at elevated [O 3 ] (-32%), while the tolerant clone, 216, had larger mature leaf area at elevated [O 3 ] (46%). These results indicate a clear difference between the two clones in their long-term response to elevated [O 3 ], which could affect competition between the clones, and result in altered genotypic composition in future atmospheric conditions. - Spring leaf flush is stimulated by elevated [CO 2 ] and suppressed by elevated [O 3 ] in aspen (Populus tremuloides).

  10. Ingestive behavior of supplemented Nellore heifers grazing palisadegrass pastures managed with different sward heights.

    Science.gov (United States)

    Vieira, Bruno Ramalho; Azenha, Mariana Vieira; Casagrande, Daniel Rume; Costa, Diogo Fleury Azevedo; Ruggieri, Ana Cláudia; Berchielli, Telma Teresinha; Reis, Ricardo Andrade

    2017-04-01

    Three sward heights (15, 25 and 35 cm) and three supplement types (energy, energy-protein, and a mineral mix supplement) were evaluated in a 3 × 3 factorial arrangement distributed in a completely randomized design to study changes in forage search patterns in Nellore heifers in a continuous grazing system. Pasture data were collected using two replicates (paddocks) per treatment over four periods during the rainy season. The behavior assessments were made in the first and fourth grazing seasons. It was hypothesized that supplements and pasture management would modify ingestive behavior, considering that animals would require less time grazing if they had energy requirements met through higher digestibility of better managed paddocks, or use of supplements high in energy. Total and green forage masses along with green : dead material ratio were greater in treatments managed with higher sward heights. Sward managed with 35 cm height resulted in lower leaf : stem ratio compared with 15 cm sward height treatments. The animals on the 15 cm pastures spent more time grazing overall and during each meal, but there were no differences observed in meal numbers in comparison to 35 cm treatments. Heifers fed protein and/or energy supplements spent less time grazing in the early afternoon, but overall grazing time was the same for all animals. © 2016 Japanese Society of Animal Science.

  11. Lucas Heights buffer zone: plan of management

    International Nuclear Information System (INIS)

    1986-01-01

    This plan is being used by the Commission as a guide for its management of the Lucas Heights buffer zone, which is essentially a circular area having a 1-6 km radius around the HIFAR reactor. Aspects covered by this plan include past uses, current use, objectives for buffer zone land management, emergency evacuation, resource conservation, archaeology, fire, access, rehabilitation of disturbed areas, resource management and plan implementation

  12. Plant traits and environment: floating leaf blade production and turnover of waterlilies.

    Science.gov (United States)

    Klok, Peter F; van der Velde, Gerard

    2017-01-01

    Floating leaf blades of waterlilies fulfill several functions in wetland ecosystems by production, decomposition and turnover as well as exchange processes. Production and turnover rates of floating leaf blades of three waterlily species, Nuphar lutea (L.) Sm., Nymphaea alba L. and Nymphaea candida Presl, were studied in three freshwater bodies, differing in trophic status, pH and alkalinity. Length and percentages of leaf loss of marked leaf blades were measured weekly during the growing season. Area and biomass were calculated based on leaf length and were used to calculate the turnover rate of floating leaf blades. Seasonal changes in floating leaf production showed that values decreased in the order: Nymphaea alba , Nuphar lutea , Nymphaea candida . The highest production was reached for Nuphar lutea and Nymphaea alba in alkaline, eutrophic water bodies. The production per leaf was relatively high for both species in the acid water body. Nymphaea candida showed a very short vegetation period and low turnover rates. The ratio Total potential leaf biomass/Maximum potential leaf biomass (P/B max ) of the three species ranged from 1.35-2.25. The ratio Vegetation period (Period with floating leaves)/Mean leaf life span ranged from 2.94-4.63, the ratio Growth period (Period with appearance of new floating leaves)/Vegetation period from 0.53-0.73. The clear differences between Nymphaea candida versus Nuphar lutea and Nymphaea alba , may be due to adaptations of Nymphaea candida to an Euro-Siberic climate with short-lasting summer conditions.

  13. Effect of Addition of Moringa Leaf By-Product (Leaf-Waste) on ...

    African Journals Online (AJOL)

    The effects of incorporation of Moringa leaf fibre (a by-product of leaf processing which contains 24% Crude Fibre by dry weight at 0, 5 and 10 % substitution of wheat flour in cookies was investigated. Three products containing wheat flour: Moringa leaf fibre ratios of 100:0, 95:5, and 90:10 respectively were prepared, and a ...

  14. Effects of wind and simulated acid mist on leaf cuticles

    International Nuclear Information System (INIS)

    Hoad, S.P.; Jeffree, C.E.; Grace, J.

    1994-01-01

    The combined effect of wind and simulated acid mist on leaf cuticles was investigated in beech (Fagus sylvatica L.) and birch (Betula pubescens Ehr.). Macroscopic and microscopic features of wind damage are described. Visibly damaged leaf area and the numbers of microscopic cuticular lesions were measured. The cuticular conductance to water vapour (g c ) of the astomatous adaxial surfaces of the leaves was measured by a gravimetric method. Field experimenntal sites were selected to provide either: 1. Direct wind action on widely-spaced plants caused by high speed and impaction of wind-blown particles, but with minimal mutual leaf abrasion 2. Indirect wind action via a high degree of mutual abrasion between closely-spaced plants. Direct wind action increased water loss via the leaf adaxial cuticle two- to three-fold in each species, by increasing the numbers of microscopic cuticular lesions. Indirect wind action caused more visible damage to leaves than direct wind action, increased g c by about threefold compared with complete shelter, and induced the most cuticular lesions. Acid mists at pH 3 or pH 5 were applied to the plants in situ at weekly intervals over a 100-day period. In sheltered plants, no effect of acid mist was detected on visibly damaged leaf area, the numbers of microscopic cuticular lesions, or on g c . However, acid mists in combination with wind exposure caused significant effects on cuticular integrity that were dependent on the type of wind action. Direct wind action combined with pH 3 acid mist resulted in the largest numbers of microscopic cuticular lesions, and the highest g c . By contrast, indirect wind action combined with pH 3 acid mist caused most visible damage to leaf tissue, but fewer microscopic lesions, and lower g c , than in plants treated with water mist. In severely-abraded leaves exposed to indirect wind action and low-pH acid rain, g c may be reduced by wound-isolation of blocks of non-functional leaf tissue. (orig.)

  15. Estimating Tree Height and Diameter at Breast Height (DBH from Digital Surface Models and Orthophotos Obtained with an Unmanned Aerial System for a Japanese Cypress (Chamaecyparis obtusa Forest

    Directory of Open Access Journals (Sweden)

    Kotaro Iizuka

    2017-12-01

    Full Text Available Methods for accurately measuring biophysical parameters are a key component for quantitative evaluation regarding to various forest applications. Conventional in situ measurements of these parameters take time and expense, encountering difficultness at locations with heterogeneous microtopography. To obtain precise biophysical data in such situations, we deployed an unmanned aerial system (UAS multirotor drone in a cypress forest in a mountainous area of Japan. The structure from motion (SfM method was used to construct a three-dimensional (3D model of the forest (tree structures from aerial photos. Tree height was estimated from the 3D model and compared to in situ ground data. We also analyzed the relationships between a biophysical parameter, diameter at breast height (DBH, of individual trees with canopy width and area measured from orthorectified images. Despite the constraints of ground exposure in a highly dense forest area, tree height was estimated at an accuracy of root mean square error = 1.712 m for observed tree heights ranging from 16 to 24 m. DBH was highly correlated with canopy width (R2 = 0.7786 and canopy area (R2 = 0.7923, where DBH ranged from 11 to 58 cm. The results of estimating forest parameters indicate that drone-based remote-sensing methods can be utilized to accurately analyze the spatial extent of forest structures.

  16. Produção de forragem do capim-Tanzânia (Panicum maximum Jacq. cv. Tanzânia-1 pastejado em diferentes alturas Forage production of Tanzaniagrass (Panicum maximum Jacq. cv. Tanzania-1 grazed at different heights

    Directory of Open Access Journals (Sweden)

    Clovenilson Cláudio Perissato Cano

    2004-12-01

    Full Text Available Objetivou-se, com este experimento, avaliar a massa de forragem (MF, massa de lâmina verde (MLV, massa de colmo + bainha verde (MCV, massa de material morto (MMM, massa de forragem verde (MFV, relação folha/colmo (F/C, taxa de acúmulo de massa seca (TAMS, acúmulo de massa de forragem (AMF, índice de área foliar (IAF, porcentagem de solo descoberto (SD e porcentagem de solo coberto com liteira (SCL em pastagem de capim-Tanzânia (Panicum maximum Jacq. cv. Tanzânia-1 manejada em quatro alturas do dossel forrageiro (20, 40, 60 e 80 cm. O método de pastejo utilizado foi o de lotação contínua e taxa de lotação variável, com novilhos da raça Nelore com peso médio de 340 kg. Utilizou-se o delineamento experimental inteiramente casualizado com duas repetições e realizaram-se cinco avaliações. MLV, MCV, MMM, MFV, MF, IAF, TAMS e AMF aumentaram com o avanço da altura do dossel, sendo que a porcentagem de SD, SCL e material morto diminui em pastos mais altos. O manejo do capim-Tanzânia nas alturas de 40 e 60 cm, apresentou as melhores respostas de composição morfológica, garantindo boa oferta de folhas, de cobertura do solo e taxa de acúmulo de massa seca. As alturas de 20 e 80 cm não devem ser recomendadas para o manejo do capim-Tanzânia quando o objetivo for produção com qualidade e quantidade.This experiment was conducted out to evaluate the forage mass (FM, green leaf lamina mass (GLLM, green stem + leaf sheath mass (GSSM, mass of dead material (MDM, green forage mass (GMF, total forage mass (TFM, leaf/stem ratio (L/S, dry matter accumulation rate (DMAR, leaf area index (LAI, % of bare soil (BS and litter cover percentage (LCP in Tanzaniagrass pasture (Panicum maximum Jacq. cv. Tanzania-1 managed at four different sward heights (20, 40, 60 and 80 cm. The grazing method was the continuous stocking with variable stocking rate, and the grazing animals were Nellore steers with average weight of 340 kg. The completely

  17. Effects of an ultraviolet-visible rays translation film on growth of leaf or root vegetables

    International Nuclear Information System (INIS)

    Hamamoto, H.; Ueno, K.; Yamazaki, K.

    2008-01-01

    A new film that absorbs ultraviolet radiation (UV) and fluoresces red light was tested as a rain shelter for the cultivation of turnip (Brassica rapa L.), spinach (Spinacia oleracea L.), and Welsh onion (Allium fistulosum L.). The effect of this UV-visible ray translation film on various growth parameters (height, fresh and dry weight, leaf area and leaf sheath diameter) was compared with those under normal clear film, new UV-cut film, and used UV-cut film respectively. The transmissivity of UV was about 70% for the normal clear film, about 20% for the UV-visible ray translation film and used UV-cut film, and about 10% for the new UV-cut film. The transmissivity of photosynthetically active radiation (PAR) was about 90% for the normal clear film and the new UV-cut film, and about 80% for the used UV-cut film, while the mean transmissivity of PAR was about 80% for the UV-visible ray translation film, with about 60% transmissivity of blue radiation and over 90% of red radiation. The UV-visible ray translation film did not promote the growth of turnip roots but did significantly promote the growth of spinaches and Welsh onions compared with the normal clear film. The UV-visible ray translation film cover promoted the growth of spinaches and Welsh onions to a similar or greater extent compared to the new UV-cut film and also to a greater extent compared to the used UV-cut film

  18. Global variability in leaf respiration in relation to climate and leaf traits

    Science.gov (United States)

    Atkin, Owen K.

    2015-04-01

    Leaf respiration plays a vital role in regulating ecosystem functioning and the Earth's climate. Because of this, it is imperative that that Earth-system, climate and ecosystem-level models be able to accurately predict variations in rates of leaf respiration. In the field of photosynthesis research, the F/vC/B model has enabled modellers to accurately predict variations in photosynthesis through time and space. By contrast, we lack an equivalent biochemical model to predict variations in leaf respiration. Consequently, we need to rely on phenomenological approaches to model variations in respiration across the Earth's surface. Such approaches require that we develop a thorough understanding of how rates of respiration vary among species and whether global environmental gradients play a role in determining variations in leaf respiration. Dealing with these issues requires that data sets be assembled on rates of leaf respiration in biomes across the Earth's surface. In this talk, I will use a newly-assembled global database on leaf respiration and associated traits (including photosynthesis) to highlight variation in leaf respiration (and the balance between respiration and photosynthesis) across global gradients in growth temperature and aridity.

  19. The construction of ventilation turrets in Atta vollenweideri leaf-cutting ants: Carbon dioxide levels in the nest tunnels, but not airflow or air humidity, influence turret structure.

    Directory of Open Access Journals (Sweden)

    Florian Halboth

    Full Text Available Nest ventilation in the leaf-cutting ant Atta vollenweideri is driven via a wind-induced mechanism. On their nests, workers construct small turrets that are expected to facilitate nest ventilation. We hypothesized that the construction and structural features of the turrets would depend on the colony's current demands for ventilation and thus might be influenced by the prevailing environmental conditions inside the nest. Therefore, we tested whether climate-related parameters, namely airflow, air humidity and CO2 levels in the outflowing nest air influenced turret construction in Atta vollenweideri. In the laboratory, we simulated a semi-natural nest arrangement with fungus chambers, a central ventilation tunnel providing outflow of air and an aboveground building arena for turret construction. In independent series, different climatic conditions inside the ventilation tunnel were experimentally generated, and after 24 hours, several features of the built turret were quantified, i.e., mass, height, number and surface area (aperture of turret openings. Turret mass and height were similar in all experiments even when no airflow was provided in the ventilation tunnel. However, elevated CO2 levels led to the construction of a turret with several minor openings and a larger total aperture. This effect was statistically significant at higher CO2 levels of 5% and 10% but not at 1% CO2. The construction of a turret with several minor openings did not depend on the strong differences in CO2 levels between the outflowing and the outside air, since workers also built permeated turrets even when the CO2 levels inside and outside were both similarly high. We propose that the construction of turrets with several openings and larger opening surface area might facilitate the removal of CO2 from the underground nest structure and could therefore be involved in the control of nest climate in leaf-cutting ants.

  20. Far-red enrichment and photosynthetically active radiation level influence leaf senescence in field-grown sunflower

    International Nuclear Information System (INIS)

    Rousseaux, M.C.; Hall, A.J.; Sánchez, R.A.

    1996-01-01

    Basal leaves frequently senesce before anthesis in high population density crops. This paper evaluates the hypothesis that quantitative and qualitative changes in the light environment associated with a high leaf area index (LAI) trigger leaf senescence in sunflower (Helianthus annuus L.) canopies. Mean leaf duration (LD, time from achievement of maximum leaf area) of leaf 8 was significantly (P < 0.05) reduced from 51 to 19 days as crop population density was increased from 0.47 to 4.76 plants m−2. High compared to low plant population density was associated with earlier reduction in the photosynthetically active radiation (PAR) and red/far-red ratio (R/FR) reaching the target leaf. However the changes in R/FR preceded those in PAR. When the light environment of individual leaves of isolated plants growing under field conditions was manipulated using filters and FR-reflecting mirrors, LD was positively and linearly related with the mean daily PAR (MDR) received in the FR- (no FR enrichment) treatments. FR enrichment of light reaching the abaxial surface of the leaf significantly (P < 0.05) reduced LD by 9 days at intermediate PAR levels with respect to FR-controls, but did not affect LD at the maximum PAR used in these experiments. However, when light reaching both leaf surfaces was enriched with FR, LD (for leaves receiving maximum PAR) was 13 days shorter than that of the FR- control. These results show that basal leaf senescence in sunflower is enhanced both by a decrease in PAR and by a decrease in R/FR. (author)

  1. Estimativa da área foliar de plantas de lima ácida 'Tahiti' usando métodos não-destrutivos Leaf area estimative of young 'Tahiti' lime using non-destructive methods

    Directory of Open Access Journals (Sweden)

    Maurício Antonio Coelho Filho

    2005-04-01

    Full Text Available O objetivo desse estudo foi avaliar métodos não-destrutivos para a determinação da área foliar de plantas jovens de lima ácida 'Tahiti'(Citrus latifolia Tan., em campo. Foram utilizadas informações de variáveis biométricas de 28 plantas jovens (0,07 a 1,44 m² e imagens digitais da área frontal de cada planta (silhueta de copa. Essas variáveis foram correlacionadas com medidas diretas (contagem total de folhas x área foliar média. Como resultado, foi possível estimar a área foliar total das plantas (AFT com base na equação: AFT = 88,936 x DI - 1,4017 (R²=0,75, em que DI representa o diâmetro do caule 5 cm abaixo do ponto em que a copa foi enxertada, e a silhueta da planta em m² (IM: AFT = 2,4951 x IM (R²=0,72. A área foliar dos ramos secundários das plantas (AFR pode ser estimada mediante uma equação exponencial envolvendo o diâmetro do ramo (DR: AFR = 0,0144e277,02 x DR (R²=0,71. Estas metodologias podem ser utilizadas quando o interesse for por um valor médio de área foliar no pomar, não sendo indicadas quando é necessária elevada precisão, pois os erros são elevados.The objective of this study was to evaluate non-destructive methods of estimating total leaf area of young 'Tahiti' lime (Citrus latifolia Tan. plants grown in the field. Information of biometrical variables of 28 young plants (0.07 to 1.44 m² and digitized image of front area of each plant (plant silhouette were used. These variables were correlated to the direct measurements (leaves and average leaf area counting. As a result, it was possible to estimate total plant leaf area (AFT based upon the equation: AFT = 88.936 x DI - 1.4017 (R²=0.75, where DI stands for the trunk diameter taken 5 cm below the graft and the silhouette area in m² (IM: AFT = 2.4951 x IM (R²=0.72. The leaf area of secondary branches (AFR can be estimated by an exponential equation with the branch diameter (DR: AFR = 0.0144e277.02 x DR (R²=0.71. These methodologies can

  2. Assessment of alveolar bone height and width using 64-MDCT examination for dental implants

    International Nuclear Information System (INIS)

    Sekiya, Kotaro; Kaneda, Takashi; Sekiya, Keiko; Mori, Shintaro; Sakayanagi, Masashi

    2011-01-01

    There have been many reports showing the usefulness of CT examinations for preoperative dental implant treatment, and some reports on clinical statistics using CT examinations. However, there have been few reports on alveolar bone height and width of over 1,000 Japanese cases. The purpose of this study was to evaluate alveolar bone height and width of 4,123 sites in 1,056 Japanese cases using preoperative CT examinations. The subjects consisted of 4,123 regions in 1,056 cases (370 males and 686 females, mean age 56.1 years old, range 15-87) of preoperative CT examinations conducted from January 2008 to March 2009. The CT examinations were performed using the Aquilion TM 64 (Toshiba Medical Systems Corporation) as the multi detector row CT (MDCT) unit, and ZIOSTATION (ZIOSOFT) as the workstation. The CT images were displayed on the workstation, and the alveolar bone height and width were measured to one decimal place (rounded off to two decimal places). The average alveolar bone height was 14.8 mm (SD±3.8) in the upper anterior area, 11.2 mm (SD±5.5) in the upper premolar area, 6.8 mm (SD±5.4) in the upper molar area, 19.5 mm (SD±5.4) in the lower anterior area, 14.2 mm (SD±3.9) in the lower premolar area, and 13.4 mm (SD±3.4) in the lower molar area. The average alveolar bone width was 4.3 mm (SD±1.9) in the anterior area, 5.7 mm (SD±2.3) in the upper premolar area, 7.9 mm (SD±3.1) in the upper molar area, 4.8 mm (SD±2.1) in the lower anterior area, 5.9 mm (SD±2.2) in the lower premolar area, and 6.9 mm (SD±2.5) in the lower molar area. Our results using preoperative CT examinations indicated that many of the Japanese cases had insufficient alveolar bone height and width for dental implants. (author)

  3. Observations of leaf stomatal conductance at the canopy scale: an atmospheric modeling perspective

    International Nuclear Information System (INIS)

    Avissar, R.

    1993-01-01

    Plant stomata play a key role in the redistribution of energy received on vegetated land into sensible and latent heat. As a result, they have a considerable impact on the atmospheric planetary boundary layer, the hydrologic cycle, the climate, and the weather. Current parameterizations of the stomatal mechanism in state-of-the-art atmospheric models are based on empirical relations that are established at the leaf scale between stomatal conductance and environmental conditions. In order to evaluate these parameterizations, an experiment was carried out on a potato field in New Jersey during the summer of 1989. Stomatal conductances were measured within a small homogeneous area in the middle of the potato field and under a relatively broad range of atmospheric conditions. A large variability of stomatal conductances was observed. This variability, which was associated with the variability of micro-environmental and physiological conditions that is found even in a homogeneous canopy, cannot be simulated explicitly on the scale of a single agricultural field and,a fortiori, on the scale of atmospheric models. Furthermore, this variability could not be related to the environmental conditions measured at a height of 2 m above the plant canopy simultaneously with the conductances, reinforcing the concept of scale decoupling suggested by Jarvis and McNaughton (1986) and McNaughton and Jarvis (1991). Thus, for atmospheric modeling purposes, a parameterization of stomatal conductance at the canopy scale using external environmental forcing conditions seems more appropriate than a parameterization based on leaf-scale stomatal conductance, as currently adopted in state-of-the-art atmospheric models. The measured variability was characterized by a lognormal probability density function (pdf) that remained relatively stable during the entire measuring period. These observations support conclusions by McNaughton and Jarvis (1991) that, unlike current parameterizations, a

  4. Observations of leaf stomatal conductance at the canopy scale: an atmospheric modeling perspective

    International Nuclear Information System (INIS)

    Avissar, R.

    1993-01-01

    Plant stomata play a key role in the redistribution of energy received on vegetated land into sensible and latent heat. As a result, they have a considerable impact on the atmospheric planetary boundary layer, the hydrologic cycle, the climate, and the weather. Current parameterizations of the stomatal mechanism in state-of-the-art atmospheric models are based on empirical relations that are established at the leaf scale between stomatal conductance and environmental conditions. In order to evaluate these parameterizations, an experiment was carried out on a potato field in New Jersey during the summer of 1989. Stomatal conductances were measured within a small homogeneous area in the middle of the potato field and under a relatively broad range of atmospheric conditions. A large variability of stomatal conductances was observed. This variability, which was associated with the variability of micro-environmental and physiological conditions that is found even in a homogeneous canopy, cannot be simulated explicitly on the scale of a single agricultural field and, a fortiori, on the scale of atmospheric models. Furthermore, this variability could not be related to the environmental conditions measured at a height of 2 m above the plant canopy simultaneously with the conductances, reinforcing the concept of scale decoupling suggested by Jarvis and McNaughton (1986) and McNaughton and Jarvis (1991). Thus, for atmospheric modeling purposes, a parameterization of stomatal conductance at the canopy scale using external environmental forcing conditions seems more appropriate than a parameterization based on leaf-scale stomatal conductance, as currently adopted in state-of-the-art atmospheric models. The measured variability was characterized by a lognormal probability density function (pdf) that remained relatively stable during the entire measuring period. These observations support conclusions by McNaughton and Jarvis (1991) that, unlike current parameterizations, a

  5. Towards a unified model for leaf trait and trait-environment relationships

    Science.gov (United States)

    Wang, H.; Harrison, S. P.; Prentice, I. C.; Peng, C.; Yang, Y.

    2016-12-01

    A widely accepted core set of leaf traits describes key aspects of plant function including the coupling among carbon, nitrogen and water cycles at the leaf, plant and ecosystem scales. Our current research focuses on two questions: (1) what dimensions of correlated variation among traits apply across all vascular plants irrespective of environment; (2) how, and to what extent, can variations in community mean values of leaf traits be predicted along environmental gradients? Based on a large quantitative trait data set covering the major environmental gradients across China, we are tackling these questions via two complementary approaches: multivariate analysis of trait-trait, trait-site, and trait-environment relationships, and the development of conceptual models and testable hypotheses for the dependencies of each trait on other traits and/or specific environmental predictors. Preliminary multivariate analyses suggest the existence of at least two independent axes of variation in leaf traits, and show robust relationships between trait syndromes and growing-season climate variables. A minimal conceptual model then considers nitrogen per unit leaf area (Narea) as a function of leaf mass per unit area (LMA) and carboxylation capacity (Vcmax); LMA as a function of irradiance, temperature and water and/or nutrient stress; Vcmax as a function of irradiance, temperature and the long-term ci:ca ratio (indexed by δ13C); and the ci:ca ratio as a function of vapour pressure deficit, temperature and atmospheric pressure. Each of these dependencies has support from observations, pointing the way towards a comprehensive set of equations to predict community-mean values of core traits in next-generation terrestrial ecosystem models.

  6. Agreement between measured height, and height predicted from ...

    African Journals Online (AJOL)

    lower limb measurements, such as knee height, as well as upper limb measures ... had with bone injuries/fractures affecting height or ulna length; and n = 1 had a ... and heels, buttocks and upper back in contact with the vertical surface of the .... found striking similarity in linear growth of infants to five-year- olds among all ...

  7. Accuracy of recumbent height measurement.

    Science.gov (United States)

    Gray, D S; Crider, J B; Kelley, C; Dickinson, L C

    1985-01-01

    Since many patients requiring specialized nutritional support are bedridden, measurement of height for purposes of nutritional assessment or prescription must often be done with the patient in bed. This study examined the accuracy of measuring body height in bed in the supine position. Two measurements were performed on 108 ambulatory inpatients: (1) standing height using a standard height-weight scale, and (2) bed height using a flexible tape. Patients were divided into four groups based on which of two researchers performed each of the two measurements. Each patient was also weighed and self-reported height, weight, sex, and age were recorded. Bed height was significantly longer than standing height by 3.68 cm, but the two measurements were equally precise. It was believed, however, that this 2% difference was probably not clinically significant in most circumstances. Bed height correlated highly with standing height (r = 0.95), and the regression equation was standing height = 13.82 +/- 0.09 bed height. Patients overestimated their heights. Heights recorded by nurses were more accurate when patients were measured than when asked about their heights, but the patients were more often asked than measured.

  8. SU-F-T-350: Continuous Leaf Optimization (CLO) for IMRT Leaf Sequencing

    Energy Technology Data Exchange (ETDEWEB)

    Long, T; Chen, M; Jiang, S; Lu, W [UT Southwestern Medical Center, Dallas, TX (United States)

    2016-06-15

    Purpose: To study a new step-and-shoot IMRT leaf sequencing model that avoids the two main pitfalls of conventional leaf sequencing: (1) target fluence being stratified into a fixed number of discrete levels and/or (2) aperture leaf positions being restricted to a discrete set of locations. These assumptions induce error into the sequence or reduce the feasible region of potential plans, respectively. Methods: We develop a one-dimensional (single leaf pair) methodology that does not make assumptions (1) or (2) that can be easily extended to a multi-row model. The proposed continuous leaf optimization (CLO) methodology takes in an existing set of apertures and associated intensities, or solution “seed,” and improves the plan without the restrictiveness of 1or (2). It then uses a first-order descent algorithm to converge onto a locally optimal solution. A seed solution can come from models that assume (1) and (2), thus allowing the CLO model to improve upon existing leaf sequencing methodologies. Results: The CLO model was applied to 208 generated target fluence maps in one dimension. In all cases for all tested sequencing strategies, the CLO model made improvements on the starting seed objective function. The CLO model also was able to keep MUs low. Conclusion: The CLO model can improve upon existing leaf sequencing methods by avoiding the restrictions of (1) and (2). By allowing for more flexible leaf positioning, error can be reduced when matching some target fluence. This study lays the foundation for future models and solution methodologies that can incorporate continuous leaf positions explicitly into the IMRT treatment planning model. Supported by Cancer Prevention & Research Institute of Texas (CPRIT) - ID RP150485.

  9. A simulation model of distributions of radiational flux at leaf surfaces in crowns of fruit trees

    International Nuclear Information System (INIS)

    Yamamoto, T.

    1988-01-01

    A computer-model was constructed for estimating distributions with time of radiational fluxes at leaf surfaces throughout fruit tree canopies in which leaves did not distribute uniformely in three dimensional space. Several assumptions were set up to construct the model for approximation of using solid geometry. For irregular distribution of leaf area in three dimensional space data were used in the simulation as number of leaves per internal cubic bloc of a cubic grid (n-divided per side). Several main parameters used were peculiar to fruit species which contain parameters (λ, ν) of Beta function to calculate both probability density function of leaf area distribution with respect to inclination angle and leaf extinction coefficient for parallel beam by leaves parameters (A, R i ) to calculate stem extinction coefficient for parallel beam, and parameters (D i ) to calculate leaf extinction coefficient of downward transmission and downward reflection. With these data and parameters solid geometry and Lambert-Beer's law constituted this model

  10. Leaf appearance rate and final main stem leaf number as affected by temperature and photoperiod in cereals grown in Mediterranean environment

    Directory of Open Access Journals (Sweden)

    Ezio Riggi

    2017-09-01

    Full Text Available In the present study, a two-year field trial was carried out with the aim to evaluate daylength and air temperature effects on leaf appearance and related rates in two durum wheat (Triticum durum Desf., two bread wheat (Triticum aestivum L. and two barley (Hordeum vulgare L. cultivars, using six different sowing dates (SD. Significant effects of SD on final main stem leaf number (FLN, thermal leaf appearance rate (TLAR, daily leaf appearance rate (DLAR and phyllochron (PhL were found. Cultivars resulted inversely correlated to mean air temperature in the interval emergence - fifth leaf full expansion (E-V. Linear response of leaf number over days after sowing was shown for all SD and cultivars, with R2 higher than 0.95. FLN linearly decreased from the first to the last SD for durum wheat, while more variable behaviour was observed in bread wheat. TLAR and DLAR showed a linear increment of the rate from the first to the last SD in durum wheat, while did not for bread wheat and barley. PhL in durum wheat decreased from the first to the last SD. Barley and bread wheat showed the highest values on those SDs which did not reach flowering. The increase of TLAR was affected by photoperiod and photothermal units in durum wheat, while by temperatures only in barley and bread wheat. Present results might find practical application in the improvement of phenology simulation models for durum wheat, bread wheat and barley grown in Mediterranean area in absence of water and nutrient stress.

  11. Plant traits and environment: floating leaf blade production and turnover of waterlilies

    Directory of Open Access Journals (Sweden)

    Peter F. Klok

    2017-04-01

    Full Text Available Floating leaf blades of waterlilies fulfill several functions in wetland ecosystems by production, decomposition and turnover as well as exchange processes. Production and turnover rates of floating leaf blades of three waterlily species, Nuphar lutea (L. Sm., Nymphaea alba L. and Nymphaea candida Presl, were studied in three freshwater bodies, differing in trophic status, pH and alkalinity. Length and percentages of leaf loss of marked leaf blades were measured weekly during the growing season. Area and biomass were calculated based on leaf length and were used to calculate the turnover rate of floating leaf blades. Seasonal changes in floating leaf production showed that values decreased in the order: Nymphaea alba, Nuphar lutea, Nymphaea candida. The highest production was reached for Nuphar lutea and Nymphaea alba in alkaline, eutrophic water bodies. The production per leaf was relatively high for both species in the acid water body. Nymphaea candida showed a very short vegetation period and low turnover rates. The ratio Total potential leaf biomass/Maximum potential leaf biomass (P/Bmax of the three species ranged from 1.35–2.25. The ratio Vegetation period (Period with floating leaves/Mean leaf life span ranged from 2.94–4.63, the ratio Growth period (Period with appearance of new floating leaves/Vegetation period from 0.53–0.73. The clear differences between Nymphaea candida versus Nuphar lutea and Nymphaea alba, may be due to adaptations of Nymphaea candida to an Euro-Siberic climate with short-lasting summer conditions.

  12. Nondestructive, stereological estimation of canopy surface area

    DEFF Research Database (Denmark)

    Wulfsohn, Dvora-Laio; Sciortino, Marco; Aaslyng, Jesper M.

    2010-01-01

    We describe a stereological procedure to estimate the total leaf surface area of a plant canopy in vivo, and address the problem of how to predict the variance of the corresponding estimator. The procedure involves three nested systematic uniform random sampling stages: (i) selection of plants from...... a canopy using the smooth fractionator, (ii) sampling of leaves from the selected plants using the fractionator, and (iii) area estimation of the sampled leaves using point counting. We apply this procedure to estimate the total area of a chrysanthemum (Chrysanthemum morifolium L.) canopy and evaluate both...... the time required and the precision of the estimator. Furthermore, we compare the precision of point counting for three different grid intensities with that of several standard leaf area measurement techniques. Results showed that the precision of the plant leaf area estimator based on point counting...

  13. Predicting human height by Victorian and genomic methods.

    Science.gov (United States)

    Aulchenko, Yurii S; Struchalin, Maksim V; Belonogova, Nadezhda M; Axenovich, Tatiana I; Weedon, Michael N; Hofman, Albert; Uitterlinden, Andre G; Kayser, Manfred; Oostra, Ben A; van Duijn, Cornelia M; Janssens, A Cecile J W; Borodin, Pavel M

    2009-08-01

    In the Victorian era, Sir Francis Galton showed that 'when dealing with the transmission of stature from parents to children, the average height of the two parents, ... is all we need care to know about them' (1886). One hundred and twenty-two years after Galton's work was published, 54 loci showing strong statistical evidence for association to human height were described, providing us with potential genomic means of human height prediction. In a population-based study of 5748 people, we find that a 54-loci genomic profile explained 4-6% of the sex- and age-adjusted height variance, and had limited ability to discriminate tall/short people, as characterized by the area under the receiver-operating characteristic curve (AUC). In a family-based study of 550 people, with both parents having height measurements, we find that the Galtonian mid-parental prediction method explained 40% of the sex- and age-adjusted height variance, and showed high discriminative accuracy. We have also explored how much variance a genomic profile should explain to reach certain AUC values. For highly heritable traits such as height, we conclude that in applications in which parental phenotypic information is available (eg, medicine), the Victorian Galton's method will long stay unsurpassed, in terms of both discriminative accuracy and costs. For less heritable traits, and in situations in which parental information is not available (eg, forensics), genomic methods may provide an alternative, given that the variants determining an essential proportion of the trait's variation can be identified.

  14. Height, Relationship Satisfaction, Jealousy, and Mate Retention

    Directory of Open Access Journals (Sweden)

    Gayle Brewer

    2009-07-01

    Full Text Available Male height is associated with high mate value. In particular, tall men are perceived as more attractive, dominant and of a higher status than shorter rivals, resulting in a greater lifetime reproductive success. Female infidelity and relationship dissolution may therefore present a greater risk to short men. It was predicted that tall men would report greater relationship satisfaction and lower jealousy and mate retention behavior than short men. Ninety eight heterosexual men in a current romantic relationship completed a questionnaire. Both linear and quadratic relationships were found between male height and relationship satisfaction, cognitive and behavioral jealousy. Tall men reported greater relationship satisfaction and lower levels of cognitive or behavioral jealousy than short men. In addition, linear and quadratic relationships were found between male height and a number of mate retention behaviors. Tall and short men engaged in different mate retention behaviors. These findings are consistent with previous research conducted in this area detailing the greater attractiveness of tall men.

  15. Characterization of leaf waste based biochar for cost effective hydrogen sulphide removal from biogas.

    Science.gov (United States)

    Sahota, Shivali; Vijay, Virendra Kumar; Subbarao, P M V; Chandra, Ram; Ghosh, Pooja; Shah, Goldy; Kapoor, Rimika; Vijay, Vandit; Koutu, Vaibhav; Thakur, Indu Shekhar

    2018-02-01

    Installation of decentralized units for biogas production along with indigenous upgradation systems can be an effective approach to meet growing energy demands of the rural population. Therefore, readily available leaf waste was used to prepare biochar at different temperatures and employed for H 2 S removal from biogas produced via anaerobic digestion plant. It is found that biochar prepared via carbonization of leaf waste at 400 °C effectively removes 84.2% H 2 S (from 1254 ppm to 201 ppm) from raw biogas for 25 min in a continuous adsorption tower. Subsequently, leaf waste biochar compositional, textural and morphological properties before and after H 2 S adsorption have been analyzed using proximate analysis, CHNS, BET surface area, FTIR, XRD, and SEM-EDX. It is found that BET surface area, pore size, and textural properties of leaf waste biochar plays a crucial role in H 2 S removal from the biogas. Copyright © 2017 Elsevier Ltd. All rights reserved.

  16. Relationship between LiDAR-derived forest canopy height and Landsat images

    Science.gov (United States)

    Cristina Pascual; Antonio Garcia-Abril; Warren B. Cohen; Susana. Martin-Fernandez

    2010-01-01

    The mean and standard deviation (SD) of light detection and ranging (LiDAR)-derived canopy height are related to forest structure. However, LiDAR data typically cover a limited area and have a high economic cost compared with satellite optical imagery. Optical images may be required to extrapolate LiDAR height measurements across a broad landscape. Different spectral...

  17. Effects of water stress on irradiance acclimation of leaf traits in almond trees.

    Science.gov (United States)

    Egea, Gregorio; González-Real, María M; Baille, Alain; Nortes, Pedro A; Conesa, María R; Ruiz-Salleres, Isabel

    2012-04-01

    Photosynthetic acclimation to highly variable local irradiance within the tree crown plays a primary role in determining tree carbon uptake. This study explores the plasticity of leaf structural and physiological traits in response to the interactive effects of ontogeny, water stress and irradiance in adult almond trees that have been subjected to three water regimes (full irrigation, deficit irrigation and rain-fed) for a 3-year period (2006-08) in a semiarid climate. Leaf structural (dry mass per unit area, N and chlorophyll content) and photosynthetic (maximum net CO(2) assimilation, A(max), maximum stomatal conductance, g(s,max), and mesophyll conductance, g(m)) traits and stem-to-leaf hydraulic conductance (K(s-l)) were determined throughout the 2008 growing season in leaves of outer south-facing (S-leaves) and inner northwest-facing (NW-leaves) shoots. Leaf plasticity was quantified by means of an exposure adjustment coefficient (ε=1-X(NW)/X(S)) for each trait (X) of S- and NW-leaves. Photosynthetic traits and K(s-l) exhibited higher irradiance-elicited plasticity (higher ε) than structural traits in all treatments, with the highest and lowest plasticity being observed in the fully irrigated and rain-fed trees, respectively. Our results suggest that water stress modulates the irradiance-elicited plasticity of almond leaves through changes in crown architecture. Such changes lead to a more even distribution of within-crown irradiance, and hence of the photosynthetic capacity, as water stress intensifies. Ontogeny drove seasonal changes only in the ε of area- and mass-based N content and mass-based chlorophyll content, while no leaf age-dependent effect was observed on ε as regards the physiological traits. Our results also indicate that the irradiance-elicited plasticity of A(max) is mainly driven by changes in leaf dry mass per unit area, in g(m) and, most likely, in the partitioning of the leaf N content.

  18. On the temporal variation of leaf magnetic parameters: seasonal accumulation of leaf-deposited and leaf-encapsulated particles of a roadside tree crown.

    Science.gov (United States)

    Hofman, Jelle; Wuyts, Karen; Van Wittenberghe, Shari; Samson, Roeland

    2014-09-15

    Understanding the accumulation behaviour of atmospheric particles inside tree leaves is of great importance for the interpretation of biomagnetic monitoring results. In this study, we evaluated the temporal variation of the saturation isothermal remanent magnetisation (SIRM) of leaves of a roadside urban Platanus × acerifolia Willd. tree in Antwerp, Belgium. We hereby examined the seasonal development of the total leaf SIRM signal as well as the leaf-encapsulated fraction of the deposited dust, by washing the leaves before biomagnetic analysis. On average 38% of the leaf SIRM signal was exhibited by the leaf-encapsulated particles. Significant correlations were found between the SIRM and the cumulative daily average atmospheric PM10 and PM2.5 measurements. Moreover, a steady increase of the SIRM throughout the in-leaf season was observed endorsing the applicability of biomagnetic monitoring as a proxy for the time-integrated PM exposure of urban tree leaves. Strongest correlations were obtained for the SIRM of the leaf-encapsulated particles which confirms the dynamic nature of the leaf surface-accumulated particles. Copyright © 2014 Elsevier B.V. All rights reserved.

  19. Preliminary application of tapered glass capillary microbeam in MeV-PIXE mapping of longan leaf for elemental concentration distribution analysis

    Science.gov (United States)

    Natyanun, S.; Unai, S.; Yu, L. D.; Tippawan, U.; Pussadee, N.

    2017-09-01

    This study was aimed at understanding elemental concentration distribution in local longan leaf for how the plant was affected by the environment or agricultural operation. The analysis applied the MeV-microbeam particle induced X-ray emission (PIXE) mapping technique using a home-developed tapered glass capillary microbeam system at Chiang Mai University. The microbeam was 2-MeV proton beam in 130 µm in diameter. The studying interest was in the difference in the elemental concentrations distributed between the leaf midrib and lamina areas. The micro proton beam analyzed the leaf sample across the leaf midrib edge to the leaf lamina area for total 9 data requisition spots. The resulting data were colored to form a 1D-map of the elemental concentration distribution. Seven dominant elements, Al, S, Cl, K, Ca, Sc and Fe, were identified, the first six of which were found having higher concentrations in the midrib area than in the lamina area, while the Fe concentration was in an opposite trend to that of the others.

  20. Potential of trees leaf/ bark to control atmospheric metals in a gas and petrochemical zone.

    Science.gov (United States)

    Safari, Mojgan; Ramavandi, Bahman; Sanati, Ali Mohammad; Sorial, George A; Hashemi, Seyedenayat; Tahmasebi, Saeid

    2018-05-22

    Leaf and bark of trees are tools for assessing the effects of the heavy metals pollution and monitoring the environmental air quality. The aim of this study was to evaluate the presence of Ni, Pb, V, and Co metals in four tree/shrub species (Conocarpus erectus, Nerium oleander, Bougainvillea spectabilis willd, and Hibiscus rosa-sinensis) in the heavily industrial zone of Asaloyeh, Iran. Two industrial zones (sites 1 and 2), two urban areas (sites 3 and 4), and two rural areas (sites 5 and 6) in the Asaloyeh industrial zone and an uncontaminated area as a control were selected. Sampling from leaf and bark of trees was carried out in spring 2016. The metals content in the washed and unwashed leaf and bark was investigated. The results showed that four studied metals in N. oleander, C. erectus, and B. spectabilis willd in all case sites were significantly higher than that of in the control site (p < 0.05). The highest concentration of metals was found in sites 3, 4, and 6; this was due to dispersion of the pollutants from industrial environments by dominant winds. The highest comprehensive bio-concentration index (CBCI) was found in leaf (0.37) and bark (0.12) of N. oleander. The maximum metal accumulation index (MAI) in the samples was found in leaf of N. oleander (1.58) and in bark of H. rosa-sinensis (1.95). The maximum bio-concentration factor (BCF) was seen for cobalt metal in the N. oleander leaf (0.89). The nickel concentration in washed-leaf samples of C. erectus was measured to be 49.64% of unwashed one. In general, the N. oleander and C. erectus species were found to have the highest absorption rate from the atmosphere and soil than other studied species, and are very suitable tools for managing air pollution in highly industrialized areas. Copyright © 2018 Elsevier Ltd. All rights reserved.

  1. Are leaf physiological traits related to leaf water isotopic enrichment in restinga woody species?

    Directory of Open Access Journals (Sweden)

    BRUNO H.P. ROSADO

    2013-09-01

    Full Text Available During plant-transpiration, water molecules having the lighter stable isotopes of oxygen and hydrogen evaporate and diffuse at a faster rate through the stomata than molecules having the heavier isotopes, which cause isotopic enrichment of leaf water. Although previous models have assumed that leaf water is well-mixed and isotopically uniform, non-uniform stomatal closure, promoting different enrichments between cells, and different pools of water within leaves, due to morpho-physiological traits, might lead to inaccuracies in isotopic models predicting leaf water enrichment. We evaluate the role of leaf morpho-physiological traits on leaf water isotopic enrichment in woody species occurring in a coastal vegetation of Brazil known as restinga. Hydrogen and oxygen stable isotope values of soil, plant stem and leaf water and leaf traits were measured in six species from restinga vegetation during a drought and a wet period. Leaf water isotopic enrichment relative to stem water was more homogeneous among species during the drought in contrast to the wet period suggesting convergent responses to deal to temporal heterogeneity in water availability. Average leaf water isotopic enrichment relative to stem water during the drought period was highly correlated with relative apoplastic water content. We discuss this observation in the context of current models of leaf water isotopic enrichment as a function of the Péclet effect. We suggest that future studies should include relative apoplastic water content in isotopic models.

  2. Are leaf physiological traits related to leaf water isotopic enrichment in restinga woody species?

    Science.gov (United States)

    Rosado, Bruno H P; De Mattos, Eduardo A; Sternberg, Leonel Da S L

    2013-09-01

    During plant-transpiration, water molecules having the lighter stable isotopes of oxygen and hydrogen evaporate and diffuse at a faster rate through the stomata than molecules having the heavier isotopes, which cause isotopic enrichment of leaf water. Although previous models have assumed that leaf water is well-mixed and isotopically uniform, non-uniform stomatal closure, promoting different enrichments between cells, and different pools of water within leaves, due to morpho-physiological traits, might lead to inaccuracies in isotopic models predicting leaf water enrichment. We evaluate the role of leaf morpho-physiological traits on leaf water isotopic enrichment in woody species occurring in a coastal vegetation of Brazil known as restinga. Hydrogen and oxygen stable isotope values of soil, plant stem and leaf water and leaf traits were measured in six species from restinga vegetation during a drought and a wet period. Leaf water isotopic enrichment relative to stem water was more homogeneous among species during the drought in contrast to the wet period suggesting convergent responses to deal to temporal heterogeneity in water availability. Average leaf water isotopic enrichment relative to stem water during the drought period was highly correlated with relative apoplastic water content. We discuss this observation in the context of current models of leaf water isotopic enrichment as a function of the Péclet effect. We suggest that future studies should include relative apoplastic water content in isotopic models.

  3. A model for the height of the internal boundary layer over an area with a irregular coastline

    DEFF Research Database (Denmark)

    Gryning, Sven-Erik; Batchvarova, E.

    1996-01-01

    of subsidence is relatively important in the afternoon under low wind speed high pressure conditions, lowering the height of the internal boundary layer by up to 10%, and it is negligible in the morning hours. The effect of the mixing height over the sea is found to be negligible....

  4. Variability of the Mixed-Layer Height Over Mexico City

    Science.gov (United States)

    García-Franco, J. L.; Stremme, W.; Bezanilla, A.; Ruiz-Angulo, A.; Grutter, M.

    2018-06-01

    The diurnal and seasonal variability of the mixed-layer height in urban areas has implications for ground-level air pollution and the meteorological conditions. Measurements of the backscatter of light pulses with a commercial lidar system were performed for a continuous period of almost six years between 2011 and 2016 in the southern part of Mexico City. The profiles were temporally and vertically smoothed, clouds were filtered out, and the mixed-layer height was determined with an ad hoc treatment of both the filtered and unfiltered profiles. The results are in agreement when compared with values of mixed-layer height reconstructed from, (i) radiosonde data, and (ii) surface and vertical column densities of a trace gas. The daily maxima of the mean mixed-layer height reach values > 3 km above ground level in the months of March-April, and are clearly lower (behaviour, which is characterized together with the mixed-layer-height anomalies. A clear residual layer is evident from the backscattered signals recorded in days with specific atmospheric conditions, but also from the cloud-filtered mean diurnal profiles. The occasional presence of a residual layer results in an overestimation of the reported mixed-layer height during the night and early morning hours.

  5. Long-term Low Radiation Decreases Leaf Photosynthesis, Photochemical Efficiency and Grain Yield in Winter Wheat

    DEFF Research Database (Denmark)

    Mu, H; Jiang, D; Wollenweber, Bernd

    2010-01-01

    the impact of low radiation on crop growth, photosynthesis and yield. Grain yield losses and leaf area index (LAI) reduction were less than the reduction in solar radiation under both shading treatment in both cultivars. Compared with the control (S0), grain yield only reduced 6.4 % and 9.9 % under 22.......0-22.9 % (S1) and 29.5-49.6 % (S2), which was consistent with the reduction in radiation. The reduction in LAI was partially compensated by increases in the fraction of the top and bottom leaf area to the total leaf area, which facilitated to intercept more solar radiation by the canopy. The decrease......Low radiation reduces wheat grain yield in tree-crop intercropping systems in the major wheat planting area of China. Here, two winter wheat (Triticum aestivum L) cultivars, Yangmai 158 (shading tolerant) and Yangmai 11 (shading sensitive), were shaded from jointing to maturity to evaluate...

  6. Final height in survivors of childhood cancer compared with Height Standard Deviation Scores at diagnosis.

    Science.gov (United States)

    Knijnenburg, S L; Raemaekers, S; van den Berg, H; van Dijk, I W E M; Lieverst, J A; van der Pal, H J; Jaspers, M W M; Caron, H N; Kremer, L C; van Santen, H M

    2013-04-01

    Our study aimed to evaluate final height in a cohort of Dutch childhood cancer survivors (CCS) and assess possible determinants of final height, including height at diagnosis. We calculated standard deviation scores (SDS) for height at initial cancer diagnosis and height in adulthood in a cohort of 573 CCS. Multivariable regression analyses were carried out to estimate the influence of different determinants on height SDS at follow-up. Overall, survivors had a normal height SDS at cancer diagnosis. However, at follow-up in adulthood, 8.9% had a height ≤-2 SDS. Height SDS at diagnosis was an important determinant for adult height SDS. Children treated with (higher doses of) radiotherapy showed significantly reduced final height SDS. Survivors treated with total body irradiation (TBI) and craniospinal radiation had the greatest loss in height (-1.56 and -1.37 SDS, respectively). Younger age at diagnosis contributed negatively to final height. Height at diagnosis was an important determinant for height SDS at follow-up. Survivors treated with TBI, cranial and craniospinal irradiation should be monitored periodically for adequate linear growth, to enable treatment on time if necessary. For correct interpretation of treatment-related late effects studies in CCS, pre-treatment data should always be included.

  7. Genetic control and combining ability of flag leaf area and relative water content traits of bread wheat cultivars under drought stress condition

    Directory of Open Access Journals (Sweden)

    Golparvar Ahmad Reza

    2013-01-01

    Full Text Available In order to compare mode of inheritance, combining ability, heterosis and gene action in genetic control of traits flag leaf area, relative water content and grain filling rate of bread wheat under drought stress, a study was conducted on 8 cultivars using of Griffing’s method2 in fixed model. Mean square of general combining ability was significant also for all traits and mean square of specific combining ability was significant also for all traits except relative water content of leaf which show importance of both additive and dominant effects of genes in heredity of these traits under stress. GCA to SCA mean square ratio was significant for none of traits. Results of this study showed that non additive effects of genes were more important than additive effect for all traits. According to results we can understand that genetic improvement of mentioned traits will have low genetic efficiency by selection from the best crosses of early generations. Then it is better to delay selection until advanced generations and increase in heritability of these traits.

  8. Leaf processing behaviour in Atta leafcutter ants: 90% of leaf cutting takes place inside the nest, and ants select pieces that require less cutting.

    Science.gov (United States)

    Garrett, Ryan W; Carlson, Katherine A; Goggans, Matthew Scott; Nesson, Michael H; Shepard, Christopher A; Schofield, Robert M S

    2016-01-01

    Leafcutter ants cut trimmings from plants, carry them to their underground nests and cut them into smaller pieces before inoculating them with a fungus that serves as a primary food source for the colony. Cutting is energetically costly, so the amount of cutting is important in understanding foraging energetics. Estimates of the cutting density, metres of cutting per square metre of leaf, were made from samples of transported leaf cuttings and of fungal substrate from field colonies of Atta cephalotes and Atta colombica. To investigate cutting inside the nest, we made leaf-processing observations of our laboratory colony, A. cephalotes. We did not observe the commonly reported reduction of the leaf fragments into a pulp, which would greatly increase the energy cost of processing. Video clips of processing behaviours, including behaviours that have not previously been described, are linked. An estimated 2.9 (±0.3) km of cutting with mandibles was required to reduce a square metre of leaf to fungal substrate. Only about 12% (±1%) of this cutting took place outside of the nest. The cutting density and energy cost is lower for leaf material with higher ratios of perimeter to area, so we tested for, and found that the laboratory ants had a preference for leaves that were pre-cut into smaller pieces. Estimates suggest that the energy required to transport and cut up the leaf material is comparable to the metabolic energy available from the fungus grown on the leaves, and so conservation of energy is likely to be a particularly strong selective pressure for leafcutter ants.

  9. Caracterización morfofisiológica de Jatropha curcas L. variedad Brasil cultivada en dos zonas de Colombia Morphophysiological characterization of Jatropha curcas L. variety Brazil in two areas of Colombia

    Directory of Open Access Journals (Sweden)

    Erik Alexander Pedraza Sánchez

    2010-01-01

    Full Text Available La Jatrofa o piñón (Jatropha curcas L. es una planta oleaginosa que ha adquirido importancia en Colombia por su alta producción y calidad de aceite para biocombustibles, despertando el interés en la investigación de su fisiología y desempeño productivo. El estudio se realizó en dos zonas de producción (Vichada y Santander, Colombia y en él se midieron la altura de la planta, el número de tallos, el área foliar, el peso seco de la hoja, el área foliar específica, el peso foliar específico, el porcentaje y perfil de ácidos grasos del aceite. Se utilizó un diseño experimental completamente al azar en el cual se consideraron dos zonas de producción, tres repeticiones y seis plantas por repetición. Las condiciones de las zonas de producción influyeron en el desarrollo vegetativo del cultivo; en ambas zonas, las hojas de los estratos inferiores de la planta presentaron mayor área. El área foliar específica y el peso foliar específico también variaron en las zonas de estudio. El contenido de aceite en la semilla no presentó variaciones y solamente el porcentaje de los ácidos grasos linoleico y oleico mostraron diferencias significativas entre las zonas.Piñón or Jatrofa (Jatropha curcas L. is an oleaginous plant that has gained importance in Colombia for their high production and quality of oil for biofuels, it has attracted interest by the investigation of its physiology and production performance. The study was conducted in two production areas (Vichada and Santander, determining plant height, stem number, leaf area, leaf dry weight, specific leaf area, specific leaf weight, percentage of oil extraction and solvent the fatty acid profile of oil by gas chromatography. A complete randomized blocks design, two production areas, three replications and six plants for replication was utilized. The conditions of the production areas influenced the development of the crop growing season, with the lower leaves of the plant showed

  10. BOREAS AFM-6 Boundary Layer Height Data

    Science.gov (United States)

    Wilczak, James; Hall, Forrest G. (Editor); Newcomer, Jeffrey A. (Editor); Smith, David E. (Technical Monitor)

    2000-01-01

    The Boreal Ecosystem-Atmosphere Study (BOREAS) Airborne Fluxes and Meteorology (AFM)-6 team from National Oceanic and Atmospheric Adminsitration/Environment Technology Laboratory (NOAA/ETL) operated a 915-MHz wind/Radio Acoustic Sounding System (RASS) profiler system in the Southern Study Area (SSA) near the Old Jack Pine (OJP) site. This data set provides boundary layer height information over the site. The data were collected from 21 May 1994 to 20 Sep 1994 and are stored in tabular ASCII files. The boundary layer height data are available from the Earth Observing System Data and Information System (EOSDIS) Oak Ridge National Laboratory (ORNL) Distributed Active Archive Center (DAAC). The data files are available on a CD-ROM (see document number 20010000884).

  11. Social inequalities in height: persisting differences today depend upon height of the parents.

    Directory of Open Access Journals (Sweden)

    Bruna Galobardes

    Full Text Available Substantial increases in height have occurred concurrently with economic development in most populations during the last century. In high-income countries, environmental exposures that can limit genetic growth potential appear to have lessened, and variation in height by socioeconomic position may have diminished. The objective of this study is to investigate inequalities in height in a cohort of children born in the early 1990s in England, and to evaluate which factors might explain any identified inequalities.12,830 children from The Avon Longitudinal Study of Parents and Children (ALSPAC, a population based cohort from birth to about 11.5 years of age, were used in this analysis. Gender- and age-specific z-scores of height at different ages were used as outcome variables. Multilevel models were used to take into account the repeated measures of height and to analyze gender- and age-specific relative changes in height from birth to 11.5 years. Maternal education was the main exposure variable used to examine socioeconomic inequalities. The roles of parental and family characteristics in explaining any observed differences between maternal education and child height were investigated. Children whose mothers had the highest education compared to those with none or a basic level of education, were 0.39 cm longer at birth (95% CI: 0.30 to 0.48. These differences persisted and at 11.5 years the height difference was 1.4 cm (95% CI: 1.07 to 1.74. Several other factors were related to offspring height, but few changed the relationship with maternal education. The one exception was mid-parental height, which fully accounted for the maternal educational differences in offspring height.In a cohort of children born in the 1990s, mothers with higher education gave birth to taller boys and girls. Although height differences were small they persisted throughout childhood. Maternal and paternal height fully explained these differences.

  12. Correlation of growth with solar radiation and air temperature on potted miniature rose

    International Nuclear Information System (INIS)

    Yu, W.; Arai, K.; Kato, K.; Imaida, K.; Nishimura, N.; Li, L.; Fukui, H.

    2006-01-01

    To establish systematic year-round production of potted miniature rose, rose growth and environmental factors such as solar radiation and air temperature were investigated for one year and the relationships of growth to these factors were analyzed. The period from the start to end of cultivation was longer in order of summer, spring and autumn cultivation. Leaf area, fresh weight of leaf and plant, leaf number and plant height as response variables were analyzed to explain the relation to environmental factors as explanatory variables using multiple linear regression analysis. The cumulative daily mean solar radiation, cumulative daytime and nighttime temperature within explanatory variables were significant main explanatory variables. Rose growth factors; leaf area, fresh weight of leaf and plant, leaf number and plant height showed close correlation with three environmental factors, respectively. Rose growth factors demonstrated significant multiple linear regressions using three environmental factors, and the parameters in multiple linear regression equations were also significant. Therefore, we demonstrated that the rose growth could be predicted using cumulative daily mean solar radiation, cumulative daytime and nighttime temperature and could be controlled by changing solar radiation and temperature

  13. Climatic and physiological effects on leaf and tree-ring stable isotopes in California redwoods

    Science.gov (United States)

    Ambrose, A. R.; Baxter, W.; Wong, C.; Dawson, T. E.; Carroll, A.; Voelker, S.

    2016-12-01

    Variation in the stable isotope composition of organic matter can provide important information about environmental change and biological responses to it. We analyzed the stable carbon (d13C) and oxygen (d18O) isotope ratios of leaves and of the cellulose from individual tree-rings of California's two redwood species to examine how these trees have responded to environmental variation and change in both time and space. Analyses of leaf d13C for both coast redwood (Sequoia sempervirens) and giant sequoia (Sequoiadendron giganteum) from throughout their geographical ranges show a marked gradient with tree height for trees of all sizes and ages but no clear difference among species or populations. The gradient is best explained by tree response to changes in both microenvironment and physiology that are known to change with height. In contrast, leaf d18O for both species showed no clear relationship with height but very clear differences between species and populations with giant sequoia displaying a much stronger inferred leaf-level response to the higher evaporative conditions present in the Sierra Nevada mountains as compared to the coast. Both species showed population-level differences with the driest and warmest sites most distinct from all of the others. Intra-annual analyses of d13C and d18O in tree-rings over a 21-year period (1974-1994) were also used to explore how climate and tree response to climate was recorded for both species. These analyses revealed unique (local) climatic effects and response to the climate for each species and population of both redwood species. Most pronounced was a significant increase in intrinsic Water Use Efficiency (iWUE) derived from d13C data over the study period in both species, and a distinct d18O response in relation to drought (e.g. 1976/1977) and to warmer days and nights and above-average precipitation (e.g., 1982-1985). Patterns of co-variation in d13C and d18O in both species suggest that during dry and also warm

  14. Monte Carlo simulation of a multi-leaf collimator design for telecobalt machine using BEAMnrc code

    International Nuclear Information System (INIS)

    Ayyangar, Komanduri M.; Narayan, Pradush; Jesuraj, Fenedit; Raju, M.R.; Dinesh Kumar, M.

    2010-01-01

    This investigation aims to design a practical multi-leaf collimator (MLC) system for the cobalt teletherapy machine and check its radiation properties using the Monte Carlo (MC) method. The cobalt machine was modeled using the BEAMnrc Omega-Beam MC system, which could be freely downloaded from the website of the National Research Council (NRC), Canada. Comparison with standard depth dose data tables and the theoretically modeled beam showed good agreement within 2%. An MLC design with low melting point alloy (LMPA) was tested for leakage properties of leaves. The LMPA leaves with a width of 7 mm and height of 6 cm, with tongue and groove of size 2 mm wide by 4 cm height, produced only 4% extra leakage compared to 10 cm height tungsten leaves. With finite 60 Co source size, the interleaf leakage was insignificant. This analysis helped to design a prototype MLC as an accessory mount on a cobalt machine. The complete details of the simulation process and analysis of results are discussed. (author)

  15. Monte Carlo simulation of a multi-leaf collimator design for telecobalt machine using BEAMnrc code

    Directory of Open Access Journals (Sweden)

    Ayyangar Komanduri

    2010-01-01

    Full Text Available This investigation aims to design a practical multi-leaf collimator (MLC system for the cobalt teletherapy machine and check its radiation properties using the Monte Carlo (MC method. The cobalt machine was modeled using the BEAMnrc Omega-Beam MC system, which could be freely downloaded from the website of the National Research Council (NRC, Canada. Comparison with standard depth dose data tables and the theoretically modeled beam showed good agreement within 2%. An MLC design with low melting point alloy (LMPA was tested for leakage properties of leaves. The LMPA leaves with a width of 7 mm and height of 6 cm, with tongue and groove of size 2 mm wide by 4 cm height, produced only 4% extra leakage compared to 10 cm height tungsten leaves. With finite 60 Co source size, the interleaf leakage was insignificant. This analysis helped to design a prototype MLC as an accessory mount on a cobalt machine. The complete details of the simulation process and analysis of results are discussed.

  16. Quantitative study of Xanthosoma violaceum leaf surfaces using RIMAPS and variogram techniques.

    Science.gov (United States)

    Favret, Eduardo A; Fuentes, Néstor O; Molina, Ana M

    2006-08-01

    Two new imaging techniques (rotated image with maximum averaged power spectrum (RIMAPS) and variogram) are presented for the study and description of leaf surfaces. Xanthosoma violaceum was analyzed to illustrate the characteristics of both techniques. Both techniques produce a quantitative description of leaf surface topography. RIMAPS combines digitized images rotation with Fourier transform, and it is used to detect patterns orientation and characteristics of surface topography. Variogram relates the mathematical variance of a surface with the area of the sample window observed. It gives the typical scale lengths of the surface patterns. RIMAPS detects the morphological variations of the surface topography pattern between fresh and dried (herbarium) samples of the leaf. The variogram method finds the characteristic dimensions of the leaf microstructure, i.e., cell length, papillae diameter, etc., showing that there are not significant differences between dry and fresh samples. The results obtained show the robustness of RIMAPS and variogram analyses to detect, distinguish, and characterize leaf surfaces, as well as give scale lengths. Both techniques are tools for the biologist to study variations of the leaf surface when different patterns are present. The use of RIMAPS and variogram opens a wide spectrum of possibilities by providing a systematic, quantitative description of the leaf surface topography.

  17. The effect of fire on habitat selection of mammalian herbivores: the role of body size and vegetation characteristics.

    Science.gov (United States)

    Eby, Stephanie L; Anderson, T Michael; Mayemba, Emilian P; Ritchie, Mark E

    2014-09-01

    Given the role of fire in shaping ecosystems, especially grasslands and savannas, it is important to understand its broader impact on these systems. Post-fire stimulation of plant nutrients is thought to benefit grazing mammals and explain their preference for burned areas. However, fire also reduces vegetation height and increases visibility, thereby potentially reducing predation risk. Consequently, fire may be more beneficial to smaller herbivores, with higher nutritional needs and greater risks of predation. We tested the impacts of burning on different sized herbivores' habitat preference in Serengeti National Park, as mediated by burning's effects on vegetation height, live : dead biomass ratio and leaf nutrients. Burning caused a less than 4 month increase in leaf nitrogen (N), and leaf non-N nutrients [copper (Cu), potassium (K), and magnesium (Mg)] and a decrease in vegetation height and live : dead biomass. During this period, total herbivore counts were higher on burned areas. Generally, smaller herbivores preferred burned areas more strongly than larger herbivores. Unfortunately, it was not possible to determine the vegetation characteristics that explained burned area preference for each of the herbivore species observed. However, total herbivore abundance and impala (Aepyceros melampus) preference for burned areas was due to the increases in non-N nutrients caused by burning. These findings suggest that burned area attractiveness to herbivores is mainly driven by changes to forage quality and not potential decreases in predation risk caused by reductions in vegetation height. © 2014 The Authors. Journal of Animal Ecology © 2014 British Ecological Society.

  18. Plasticity in the Huber value contributes to homeostasis in leaf water relations of a mallee Eucalypt with variation to groundwater depth.

    Science.gov (United States)

    Carter, Jennifer L; White, Donald A

    2009-11-01

    Information on how vegetation adapts to differences in water supply is critical for predicting vegetation survival, growth and water use, which, in turn, has important impacts on site hydrology. Many field studies assess adaptation to water stress by comparing between disparate sites, which makes it difficult to distinguish between physiological or morphological changes and long-term genetic adaptation. When planting trees into new environments, the phenotypic adaptations of a species to water stress will be of primary interest. This study examined the response to water availability of Eucalyptus kochii ssp. borealis (C. Gardner) D. Nicolle, commonly integrated with agriculture in south-western Australia for environmental and economic benefits. By choosing a site where the groundwater depth varied but where climate and soil type were the same, we were able to isolate tree response to water supply. Tree growth, leaf area and stand water use were much larger for trees over shallow groundwater than for trees over a deep water table below a silcrete hardpan. However, water use on a leaf area basis was similar in trees over deep and shallow groundwater, as were the minimum leaf water potential observed over different seasons and the turgor loss point. We conclude that homeostasis in leaf water use and water relations was maintained through a combination of stomatal control and adjustment of sapwood-to-leaf area ratios (Huber value). Differences in the Huber value with groundwater depth were associated with different sapwood-specific conductivity and water use on a sapwood area basis. Knowledge of the coordination between water supply, leaf area, sapwood area and leaf transpiration rate for different species will be important when predicting stand water use.

  19. Characteristic, inheritance and breeding application of rice mutants with greenable albino leaf

    International Nuclear Information System (INIS)

    Fang Xiantao; Ma Hongli; Zhao Fuyuan; Zhang Qingqi; Zhang Shubiao

    2009-01-01

    Inheritance and main agronomic traits of photo-thermo-sensitive genic male sterile line with green-revertible albino leaf were investigated. The results indicated that the mutants might be divided into three types: albino regreening type (W2, W3, W4 and W10), albino to kelly type (W9) and abino-regreening-albino-regreening type (W1 and W7). Genetic study indicated that green-revertible albino leaf color trait of the mutants as controlled by a single recessive gene. These mutants had similar agronomic traits and fertility characteristics to the corresponding male sterile line 'Peiai 64S'. The hybrids of these mutants had similar characteristics with original-hybrids in plant type, developing of tillers and plant height. The yield components of the mutant hybrids were different depending on different mutants. The yield potential of hybrids of W1, W2 and W3 were similar to the original-hybrid. The results also indicated that W1, W2 and W3 had breeding application value. (authors)

  20. Gravity and Height Variations at Medicina, Italy

    Science.gov (United States)

    Bruni, Sara; Zerbini, Susanna; Errico, Maddalena; Santi, Efisio; Wziontek, Hartmut

    2017-04-01

    Since 1996, at the Medicina station, height and gravity variations are monitored continuously by means of GPS, VLBI and superconducting gravimeter (SG) data. Additionally, absolute gravity observations are performed twice a year and environmental parameters, among others water table levels, are regularly acquired. Levelling between the different monuments at the site area is also carried out repeatedly to constrain local ties in the vertical position. Two GPS systems are located very close to each other, and both are in close proximity to the VLBI antenna. Twenty years of data are now available, which allow investigating both long- and short-period height and gravity signals together with their relevant correlations. Natural land subsidence, which is well known to occur in the area, is a major component of the observed long-term behavior; however, non-linear long-period signatures are also present in the time series. On a shorter time scale, fingerprints of the water table seasonal oscillations can be recognized in the data. The Medicina site is characterized by clayey soil subjected to consolidation effects when the water table lowers during summer periods. The pillar on which the SG is installed is especially affected because of its shallow foundation, causing height decreases in the order of 2.5-3 cm for water table lowering of 2 m. This study presents a comparative analysis of the different data sets with the aim of separating mass and deformation contributions in the SG gravity record.

  1. Springtime Leaf Development of Mature Sessile Oak Trees as Based on Multi-Seasonal Monitoring Data

    Directory of Open Access Journals (Sweden)

    NYITRAI, Balázs

    2012-01-01

    Full Text Available Based on a four year leaf growth data-set we modelled the thermal time-dependent leafontogeny in upper and lower canopy layers of mature sessile oak trees, in a Quercetum petraeae-cerrisforest stand (NE Hungary. Our regression models revealed no considerable differences between thetiming of leaf unfolding and leaf expansion of different canopy layers. On the other hand seasonalcourse in leaf mass-to-area ratio (LMA indicated that sun leaves needed considerably longer thermaltime to fully develop their anatomical structures compared to shade leaves. LMA of sun leaves washigher during the whole leaf maturation process suggesting that ‘sun’ and ‘shade’ characteristicsdevelop in very early stage of leaf ontogeny. Functioning of photosynthetic apparatus (Fv/Fo in shadeleaves have built up faster and performed better in all developmental stages which could be attributedto two main factors: 1 very early determination of leaf traits as a function of light environment and 2evolving shading effect of upper canopy layer eliminates photoinhibition in lower leaves.

  2. Contributing factors in foliar uptake of dissolved inorganic nitrogen at leaf level

    Energy Technology Data Exchange (ETDEWEB)

    Wuyts, Karen, E-mail: karen.wuyts@uantwerpen.be [Laboratory of Environmental and Urban Ecology, Research Group ENdEMIC, Dept. Bioscience Engineering, University of Antwerp, Groenenborgerlaan 171, B-2020 Antwerp (Belgium); Forest and Nature Lab (ForNaLab), Dept. Forest and Water Management, Ghent University, Geraardsbergsesteenweg 267, B-9090 Gontrode-Melle (Belgium); Adriaenssens, Sandy, E-mail: adriaenssens@irceline.be [Belgian Interregional Environment Agency (IRCEL-CELINE), Kunstlaan 10–11, B-1210 Brussels (Belgium); Staelens, Jeroen, E-mail: jeroen_staelens@yahoo.com [Flemish Environment Agency (VMM), Kronenburgstraat 45, B-2000 Antwerp (Belgium); Wuytack, Tatiana, E-mail: tatiana.wuytack@uantwerpen.be [Laboratory of Environmental and Urban Ecology, Research Group ENdEMIC, Dept. Bioscience Engineering, University of Antwerp, Groenenborgerlaan 171, B-2020 Antwerp (Belgium); Van Wittenberghe, Shari, E-mail: shari.vanwittenberghe@uantwerpen.be [Laboratory of Environmental and Urban Ecology, Research Group ENdEMIC, Dept. Bioscience Engineering, University of Antwerp, Groenenborgerlaan 171, B-2020 Antwerp (Belgium); Boeckx, Pascal, E-mail: pascal.boeckx@ugent.be [Isotope Bioscience Laboratory (ISOFYS), Dept. Applied Analytical and Physical Chemistry, Ghent University, Coupure Links 653, B-9000 Ghent (Belgium); Samson, Roeland, E-mail: roeland.samson@uantwerpen.be [Laboratory of Environmental and Urban Ecology, Research Group ENdEMIC, Dept. Bioscience Engineering, University of Antwerp, Groenenborgerlaan 171, B-2020 Antwerp (Belgium); Verheyen, Kris, E-mail: kris.verheyen@ugent.be [Forest and Nature Lab (ForNaLab), Dept. Forest and Water Management, Ghent University, Geraardsbergsesteenweg 267, B-9090 Gontrode-Melle (Belgium)

    2015-02-01

    We investigated the influence of leaf traits, rainwater chemistry, and pedospheric nitrogen (N) fertilisation on the aqueous uptake of inorganic N by physiologically active tree leaves. Leaves of juvenile silver birch and European beech trees, supplied with NH{sub 4}NO{sub 3} to the soil at rates from 0 to 200 kg N ha{sup −1} y{sup −1}, were individually exposed to 100 μl of artificial rainwater containing {sup 15}NH{sub 4}{sup +} or {sup 15}NO{sub 3}{sup −} at two concentration levels for one hour. In the next vegetative period, the experiment was repeated with NH{sub 4}{sup +} at the highest concentration only. The N form and the N concentration in the applied rainwater and, to a lesser extent, the pedospheric N treatment and the leaf traits affected the aqueous foliar N uptake. The foliar uptake of NH{sub 4}{sup +} by birch increased when leaves were more wettable. High leaf N concentration and leaf mass per area enhanced the foliar N uptake, and NO{sub 3}{sup −} uptake in particular, by birch. Variation in the foliar N uptake by the beech trees could not be explained by the leaf traits considered. In the first experiment, N fertilisation stimulated the foliar N uptake in both species, which was on average 1.42–1.78 times higher at the highest soil N dose than at the zero dose. However, data variability was high and the effect was not appreciable in the second experiment. Our data suggest that next to rainwater chemistry (N form and concentration) also forest N status could play a role in the partitioning of N entering the ecosystem through the soil and the canopy. Models of canopy uptake of aqueous N at the leaf level should take account of leaf traits such as wettability and N concentration. - Highlights: • Foliar uptake of dissolved inorganic nitrogen (N) by potted trees was studied. • Leaves were individually exposed to rainwater drops containing {sup 15}NH{sub 4}{sup +} or {sup 15}NO{sub 3}{sup −}. • Foliar N uptake efficiency depended on

  3. Morpho-anatomy of the leaf of Myrciaria glomerata

    Directory of Open Access Journals (Sweden)

    Nemes Veiga Pacheco-Silva

    Full Text Available Abstract Myrciaria glomerata O. Berg., Myrtaceae, popularly known as "cabeludinha", has high content of ascorbic acid and anti-inflammatory property and is used in folk medicine. The objectives of this study were the morphological, anatomical and histochemical characterization of the leaves. Leaf studies were made with optical, scanning electron and confocal microscopy. The collection of botanical material was held at the Tijuca Forest, Rio de Janeiro, RJ. Histochemical tests aimed the identification of lipids, starch grains, phenolic compounds and crystals. The leaves are simple, opposite, lanceolate, pinnate, hairy, with involute margins, hypostomatic and dorsiventral. The stomata are anomocytic. The epidermis presents simple trichomes. Epidermal cells show uneven thickening of their periclinal outer walls, mainly on the adaxial side of the leaf. Secretory cavities of essential oils are subepidermal and exceed, in height, the palisade parenchyma, formed by one cell layer. Four to five cellular layers, rich in phenolic compounds and lipids form the spongy parenchyma. The bundles are collateral and there are many crystals of calcium oxalate spread throughout the mesophyll. In the midrib and petiole the bundles are bicollateral. Analysis by scanning electron revealed epicuticular wax rod-shaped and as grains. In confocal microscopy, the adaxial epidermis, the fibers and the secretory epithelium of the cavities show autofluorescence. The data obtained are important in quality control exams of samples of this species.

  4. Leaf area index retrieval using Hyperion EO-1 data-based vegetation indices in Himalayan forest system

    Science.gov (United States)

    Singh, Dharmendra; Singh, Sarnam

    2016-04-01

    Present Study is being taken to retrieve Leaf Area Indexn(LAI) in Himalayan forest system using vegetation indices developed from Hyperion EO-1 hyperspectral data. Hemispherical photograph were captured in the month of March and April, 2012 at 40 locations, covering moist tropical Sal forest, subtropical Bauhinia and pine forest and temperate Oak forest and analysed using an open source GLA software. LAI in the study region was ranging in between 0.076 m2/m2 to 6.00 m2/m2. These LAI values were used to develop spectral models with the FLAASH corrected Hyperion measurements.Normalized difference vegetation index (NDVI) was used taking spectral reflectance values of all the possible combinations of 170 atmospherically corrected channels. The R2 was ranging from lowest 0.0 to highest 0.837 for the band combinations of spectral region 640 nm and 670 nm. The spectral model obtained was, spectral reflectance (y) = 0.02x LAI(x) - 0.0407.

  5. Four novel Talaromyces species isolated from leaf litter from Colombian Amazon rain forests

    DEFF Research Database (Denmark)

    Yilmaz, Neriman; López-Quintero, Carlos A.; Vasco-Palacios, Aída Marcela

    2016-01-01

    Various Talaromyces strains were isolated during a survey of fungi involved in leaf litter decomposition in tropical lowland forests in the Caquetá and Amacayacu areas of the Colombian Amazon. Four new Talaromyces species are described using a polyphasic approach, which includes phenotypic......). In addition to the new species, T. aculeatus and T. macrosporus were isolated during this study on leaf litter decomposition....

  6. Green leaf phenology at Landsat resolution: scaling from the plot to satellite

    Science.gov (United States)

    Fisher, J. I.; Mustard, J. F.; Vadeboncour, M.

    2005-12-01

    Despite the large number of in situ, plot-level phenological measurements and satellite-derived phenological studies, there has been little success to date in merging these records temporally or spatially. In particular, while most phenological patterns and trends derived from satellites appear realistic and coherent, they may not reflect spatial and temporal patterns at the plot level. An obvious explanation is the drastic scale difference from plot-level to most satellite observations. In this research, we bridge this scale gap through higher resolution satellite records (Landsat) and quantify the accuracy of satellite-derived metrics with direct field measurements. We compiled fifty-seven Landsat scenes from southern New England (P12 R51) from 1984 to 2002. Green vegetation areal abundance for each scene was derived from spectral mixture analysis and a single set of endmembers. The leaf area signal was fit with a logistic-growth simulating sigmoid curve to derive phenological markers (half-maximum leaf-onset and offset). Spring leaf-onset dates in homogenous stands of deciduous forests displayed significant and persistent local variability. The local variability was validated with multiple springtime ground observations (r2 = 0.91). The highest degree of verified small-scale variation occurred where contiguous forests displayed leaf-onset gradients of 10-14 days over short distances (example, our results indicate that deciduous forests in the Providence, RI metropolitan area leaf out 5-7 days earlier than comparable rural areas. In preliminary work, we validated the Landsat-derived metrics with similar analyses of MODIS and AVHRR, and demonstrate that aggregating diverse local phenologies into coarse grids may convolute interpretations. Despite these complications, the platform-independent curve-fit methodology may be extended across platforms and field data. The methodologically consistent approach, in tandem with Landsat data, allows us to effectively scale

  7. Beyond leaf color: Comparing camera-based phenological metrics with leaf biochemical, biophysical, and spectral properties throughout the growing season of a temperate deciduous forest

    Science.gov (United States)

    Yang, Xi; Tang, Jianwu; Mustard, John F.

    2014-03-01

    Plant phenology, a sensitive indicator of climate change, influences vegetation-atmosphere interactions by changing the carbon and water cycles from local to global scales. Camera-based phenological observations of the color changes of the vegetation canopy throughout the growing season have become popular in recent years. However, the linkages between camera phenological metrics and leaf biochemical, biophysical, and spectral properties are elusive. We measured key leaf properties including chlorophyll concentration and leaf reflectance on a weekly basis from June to November 2011 in a white oak forest on the island of Martha's Vineyard, Massachusetts, USA. Concurrently, we used a digital camera to automatically acquire daily pictures of the tree canopies. We found that there was a mismatch between the camera-based phenological metric for the canopy greenness (green chromatic coordinate, gcc) and the total chlorophyll and carotenoids concentration and leaf mass per area during late spring/early summer. The seasonal peak of gcc is approximately 20 days earlier than the peak of the total chlorophyll concentration. During the fall, both canopy and leaf redness were significantly correlated with the vegetation index for anthocyanin concentration, opening a new window to quantify vegetation senescence remotely. Satellite- and camera-based vegetation indices agreed well, suggesting that camera-based observations can be used as the ground validation for satellites. Using the high-temporal resolution dataset of leaf biochemical, biophysical, and spectral properties, our results show the strengths and potential uncertainties to use canopy color as the proxy of ecosystem functioning.

  8. Mapping canopy gap fraction and leaf area index at continent-scale from satellite lidar

    Science.gov (United States)

    Mahoney, C.; Hopkinson, C.; Held, A. A.

    2015-12-01

    Information on canopy cover is essential for understanding spatial and temporal variability in vegetation biomass, local meteorological processes and hydrological transfers within vegetated environments. Gap fraction (GF), an index of canopy cover, is often derived over large areas (100's km2) via airborne laser scanning (ALS), estimates of which are reasonably well understood. However, obtaining country-wide estimates is challenging due to the lack of spatially distributed point cloud data. The Geoscience Laser Altimeter System (GLAS) removes spatial limitations, however, its large footprint nature and continuous waveform data measurements make derivations of GF challenging. ALS data from 3 Australian sites are used as a basis to scale-up GF estimates to GLAS footprint data by the use of a physically-based Weibull function. Spaceborne estimates of GF are employed in conjunction with supplementary predictor variables in the predictive Random Forest algorithm to yield country-wide estimates at a 250 m spatial resolution; country-wide estimates are accompanied with uncertainties at the pixel level. Preliminary estimates of effective Leaf Area Index (eLAI) are also presented by converting GF via the Beer-Lambert law, where an extinction coefficient of 0.5 is employed; deemed acceptable at such spatial scales. The need for such wide-scale quantification of GF and eLAI are key in the assessment and modification of current forest management strategies across Australia. Such work also assists Australia's Terrestrial Ecosystem Research Network (TERN), a key asset to policy makers with regards to the management of the national ecosystem, in fulfilling their government issued mandates.

  9. Spatio-temporal evaluation of plant height in corn via unmanned aerial systems

    Science.gov (United States)

    Varela, Sebastian; Assefa, Yared; Vara Prasad, P. V.; Peralta, Nahuel R.; Griffin, Terry W.; Sharda, Ajay; Ferguson, Allison; Ciampitti, Ignacio A.

    2017-07-01

    Detailed spatial and temporal data on plant growth are critical to guide crop management. Conventional methods to determine field plant traits are intensive, time-consuming, expensive, and limited to small areas. The objective of this study was to examine the integration of data collected via unmanned aerial systems (UAS) at critical corn (Zea mays L.) developmental stages for plant height and its relation to plant biomass. The main steps followed in this research were (1) workflow development for an ultrahigh resolution crop surface model (CSM) with the goal of determining plant height (CSM-estimated plant height) using data gathered from the UAS missions; (2) validation of CSM-estimated plant height with ground-truthing plant height (measured plant height); and (3) final estimation of plant biomass via integration of CSM-estimated plant height with ground-truthing stem diameter data. Results indicated a correlation between CSM-estimated plant height and ground-truthing plant height data at two weeks prior to flowering and at flowering stage, but high predictability at the later growth stage. Log-log analysis on the temporal data confirmed that these relationships are stable, presenting equal slopes for both crop stages evaluated. Concluding, data collected from low-altitude and with a low-cost sensor could be useful in estimating plant height.

  10. Constructing seasonal LAI trajectory by data-model fusion for global evergreen needle-leaf forests

    Science.gov (United States)

    Wang, R.; Chen, J.; Mo, G.

    2010-12-01

    For decades, advancements in optical remote sensors made it possible to produce maps of a biophysical parameter--the Leaf Area Index (LAI), which is critically necessary in regional and global modeling of exchanges of carbon, water, energy and other substances, across large areas in a fast way. Quite a few global LAI products have been generated since 2000, e.g. GLOBCARBON (Deng et al., 2006), MODIS Collection 5 (Shabanov et al., 2007), CYCLOPES (Baret et al., 2007), etc. Albeit these progresses, the basic physics behind the technology restrains it from accurate estimation of LAI in winter, especially for northern high-latitude evergreen needle-leaf forests. Underestimation of winter LAI in these regions has been reported in literature (Yang et al., 2000; Cohen et al., 2003; Tian et al., 2004; Weiss et al., 2007; Pisek et al., 2007), and the distortion is usually attributed to the variations of canopy reflectance caused by understory change (Weiss et al., 2007) as well as by the presence of ice and snow on leaves and ground (Cohen, 2003; Tian et al., 2004). Seasonal changes in leaf pigments can also be another reason for low LAI retrieved in winter. Low conifer LAI values in winter retrieved from remote sensing make them unusable for surface energy budget calculations. To avoid these drawbacks of remote sensing approaches, we attempt to reconstruct the seasonal LAI trajectory through model-data fusion. A 1-degree LAI map of global evergreen needle-leaf forests at 10-day interval is produced based on the carbon allocation principle in trees. With net primary productivity (NPP) calculated by the Boreal Ecosystems Productivity Simulator (BEPS) (Chen et al., 1999), carbon allocated to needles is quantitatively evaluated and then can be further transformed into LAI using the specific leaf area (SLA). A leaf-fall scheme is developed to mimic the carbon loss caused by falling needles throughout the year. The seasonally maximum LAI from remote sensing data for each pixel

  11. Final height in survivors of childhood cancer compared with Height Standard Deviation Scores at diagnosis

    NARCIS (Netherlands)

    Knijnenburg, S. L.; Raemaekers, S.; van den Berg, H.; van Dijk, I. W. E. M.; Lieverst, J. A.; van der Pal, H. J.; Jaspers, M. W. M.; Caron, H. N.; Kremer, L. C.; van Santen, H. M.

    2013-01-01

    Our study aimed to evaluate final height in a cohort of Dutch childhood cancer survivors (CCS) and assess possible determinants of final height, including height at diagnosis. We calculated standard deviation scores (SDS) for height at initial cancer diagnosis and height in adulthood in a cohort of

  12. The narrow-leaf syndrome: a functional and evolutionary approach to the form of fog-harvesting rosette plants.

    Science.gov (United States)

    Martorell, Carlos; Ezcurra, Exequiel

    2007-04-01

    Plants that use fog as an important water-source frequently have a rosette growth habit. The performance of this morphology in relation to fog interception has not been studied. Some first-principles from physics predict that narrow leaves, together with other ancillary traits (large number and high flexibility of leaves, caudices, and/or epiphytism) which constitute the "narrow-leaf syndrome" should increase fog-interception efficiency. This was tested using aluminum models of rosettes that differed in leaf length, width and number and were exposed to artificial fog. The results were validated using seven species of Tillandsia and four species of xerophytic rosettes. The total amount of fog intercepted in rosette plants increased with total leaf area, while narrow leaves maximized interception efficiency (measured as interception per unit area). The number of leaves in the rosettes is physically constrained because wide-leafed plants can only have a few blades. At the limits of this constraint, net fog interception was independent of leaf form, but interception efficiency was maximized by large numbers of narrow leaves. Atmospheric Tillandsia species show the narrow-leaf syndrome. Their fog interception efficiencies were correlated to the ones predicted from aluminum-model data. In the larger xerophytic rosette species, the interception efficiency was greatest in plants showing the narrow-leaf syndrome. The adaptation to fog-harvesting in several narrow-leaved rosettes was tested for evolutionary convergence in 30 xerophytic rosette species using a comparative method. There was a significant evolutionary tendency towards the development of the narrow-leaf syndrome the closer the species grew to areas where fog is frequently available. This study establishes convergence in a very wide group of plants encompassing genera as contrasting as Tillandsia and Agave as a result of their dependence on fog.

  13. Generating Global Leaf Area Index from Landsat: Algorithm Formulation and Demonstration

    Science.gov (United States)

    Ganguly, Sangram; Nemani, Ramakrishna R.; Zhang, Gong; Hashimoto, Hirofumi; Milesi, Cristina; Michaelis, Andrew; Wang, Weile; Votava, Petr; Samanta, Arindam; Melton, Forrest; hide

    2012-01-01

    This paper summarizes the implementation of a physically based algorithm for the retrieval of vegetation green Leaf Area Index (LAI) from Landsat surface reflectance data. The algorithm is based on the canopy spectral invariants theory and provides a computationally efficient way of parameterizing the Bidirectional Reflectance Factor (BRF) as a function of spatial resolution and wavelength. LAI retrievals from the application of this algorithm to aggregated Landsat surface reflectances are consistent with those of MODIS for homogeneous sites represented by different herbaceous and forest cover types. Example results illustrating the physics and performance of the algorithm suggest three key factors that influence the LAI retrieval process: 1) the atmospheric correction procedures to estimate surface reflectances; 2) the proximity of Landsatobserved surface reflectance and corresponding reflectances as characterized by the model simulation; and 3) the quality of the input land cover type in accurately delineating pure vegetated components as opposed to mixed pixels. Accounting for these factors, a pilot implementation of the LAI retrieval algorithm was demonstrated for the state of California utilizing the Global Land Survey (GLS) 2005 Landsat data archive. In a separate exercise, the performance of the LAI algorithm over California was evaluated by using the short-wave infrared band in addition to the red and near-infrared bands. Results show that the algorithm, while ingesting the short-wave infrared band, has the ability to delineate open canopies with understory effects and may provide useful information compared to a more traditional two-band retrieval. Future research will involve implementation of this algorithm at continental scales and a validation exercise will be performed in evaluating the accuracy of the 30-m LAI products at several field sites. ©

  14. Bringing satellite winds to hub-height

    DEFF Research Database (Denmark)

    Badger, Merete; Pena Diaz, Alfredo; Bredesen, Rolv Erlend

    2012-01-01

    Satellite observations of the ocean surface can provide detailed information about the spatial wind variability over large areas. This is very valuable for the mapping of wind resources offshore where other measurements are costly and sparse. Satellite sensors operating at microwave frequencies...... measure the amount of radar backscatter from the sea surface, which is a function of the instant wind speed, wind direction, and satellite viewing geometry. A major limitation related to wind retrievals from satellite observations is that existing empirical model functions relate the radar backscatter...... to wind speed at the height 10 m only. The extrapolation of satellite wind fields to higher heights, which are more relevant for wind energy, remains a challenge which cannot be addressed by means of satellite data alone. As part of the EU-NORSEWInD project (2008-12), a hybrid method has been developed...

  15. Determinación del Área Foliar en Fotografías Tomadas con una Cámara Web, un Teléfono Celular o una Cámara Semiprofesional Leaf Area Measurement in Photographs Taken with a Webcam, a Cell Phone or a Semi Professional Camera

    Directory of Open Access Journals (Sweden)

    Natalia Rincón Guerrero

    2012-06-01

    Full Text Available Resumen. Las determinaciones de área foliar son muy usadas en investigación agrícola. Para su cuantificación existen equipos automáticos, pero costosos y de escasa disponibilidad. Para superar estos limitantes, en esta investigación se acoplaron el uso de cámaras digitales (de un teléfono celular, una cámara web o una cámara digital semiprofesional y un software libre para análisis de imágenes (ImageJ, con el fin de determinar el área foliar en forma simple, precisa y a bajo costo. Para estimar la sensibilidad de este sistema, en la primera prueba se compararon las áreas generadas por un medidor de área foliar y por las fotografías tomadas con tres tipos de cámara digital, sobre cuadrados de papel verde de áreas conocidas. En la segunda prueba se correlacionaron los valores de área foliar obtenidos con el medidor de área foliar y los obtenidos a partir del análisis de fotografías de hojas tomadas con las tres cámaras digitales, en cinco especies vegetales, que variaban en morfología y color de hojas. Los resultados indicaron coeficientes de correlación mayores a 0,99 entre los valores de área real de las piezas de papel y los valores de área estimados por el medidor de área foliar o de las fotografías. En la segunda prueba, las mediciones de area foliar obtenidas con el medidor de area foliar y con las cámaras digitales presentaron coeficientes de correlación muy altos (r=1. Los coeficientes de líneas de regresión entre las áreas del medidor de área foliar y de las fotografías del teléfono celular estuvieron entre 0,98 y 1,02, indicando una desviación máxima de 2% comparado con las mediciones del medidor de área foliar. Estos resultados sugieren una gama amplia de posibilidades para este sistema como herramienta para la investigación agrícola y biológica, debido a la disponibilidad de estos equipos que son de bajo costo, fáciles de usar y portátiles.Abstract. Leaf area is a plant trait widely assessed

  16. Urbanization-related changes in European aspen (Populus tremula L.): Leaf traits and litter decomposition

    International Nuclear Information System (INIS)

    Nikula, Suvi; Vapaavuori, Elina; Manninen, Sirkku

    2010-01-01

    We investigated foliar and litter responses of European aspen (Populus tremula L.) to urbanization, including factors such as increased temperature, moisture stress and nitrogen (N) deposition. Leaf samples were collected in 2006-2008 from three urban and three rural forest stands in the Helsinki Metropolitan Area, southern Finland, and reciprocal litter transplantations were established between urban and rural sites. Urban leaves exhibited a higher amount of epicuticular waxes and N concentration, and a lower C:N ratio than rural ones, but there was no difference in specific leaf area. Urban litter had a slightly higher N concentration, lower concentrations of lignin and total phenolics, and was more palatable to a macrofaunal decomposer. Moreover, litter decay was faster at the urban site and for urban litter. Urbanization thus resulted in foliar acclimatization in terms of increased amount of epicuticular waxes, as well as in accelerated decomposition of the N-richer leaf litter. - Urbanization can modify leaf traits of aspen and accelerate litter decomposition through changes in litter traits as well as in environmental conditions at the decomposition site.

  17. Height premium for job performance.

    Science.gov (United States)

    Kim, Tae Hyun; Han, Euna

    2017-08-01

    This study assessed the relationship of height with wages, using the 1998 and 2012 Korean Labor and Income Panel Study data. The key independent variable was height measured in centimeters, which was included as a series of dummy indicators of height per 5cm span (wages to assess the heterogeneity in the height-wage relationship, across the conditional distribution of monthly wages. We found a non-linear relationship of height with monthly wages. For men, the magnitude of the height wage premium was overall larger at the upper quantile of the conditional distribution of log monthly wages than at the median to low quantile, particularly in professional and semi-professional occupations. The height-wage premium was also larger at the 90th quantile for self-employed women and salaried men. Our findings add a global dimension to the existing evidence on height-wage premium, demonstrating non-linearity in the association between height and wages and heterogeneous changes in the dispersion and direction of the association between height and wages, by wage level. Copyright © 2017 Elsevier B.V. All rights reserved.

  18. Red Guava Leaf Harvesting Impact on Flavonoid Optimation in Different Growth Phases

    Directory of Open Access Journals (Sweden)

    MUNIF GHULAMAHDI

    2011-06-01

    Full Text Available Harvesting process is a critical time to identify the quality of raw material for traditional medicine. The time and harvesting techniques, drying process after harvesting, and processing to make the simplicia, are the crucial role to make the good quality of the natural product. On the other hand, there is a lack of general understanding and appreciation about the processes involved in governing shoot and tree growth and development, i.e. red guava. The research objective was to evaluate the influence of leaf harvesting and growth phases on red guava for flavonoid production as antioxidant. Randomized factorial block design in time were laid out with two factors and followed by Duncan’s multiple range test. The treatments were the amount of leaf harvested on tertiary branches (0, 25, 50, and 100% and growth phases of the plant (vegetative and generative. Leaf harvesting 25% on tertiary branches significantly increased the leaf number (766.3 tree-1 and the number of new quarternary branches, decreasing leaf area index (LAI and leaf dry weight at the end of the experiment (22 weeks of observation/WO. The highest leaf dry weight (156.94 g tree-1 and LAI (0.47 was found in harvesting 25% tertiary branches. Harvesting 100% leaf on tertiary branches in vegetative phase significantly produced the lowest flavonoid production (7.82 g tree-1. The result suggested that flavonoid production from red guava leaves should be done by harvesting 50% leaf on tertiary branches in generative phase can be used to produce the highest flavonoid (89.90 g tree-1.

  19. Leaf endophyte load influences fungal garden development in leaf-cutting ants

    Directory of Open Access Journals (Sweden)

    Van Bael Sunshine A

    2012-11-01

    Full Text Available Abstract Background Previous work has shown that leaf-cutting ants prefer to cut leaf material with relatively low fungal endophyte content. This preference suggests that fungal endophytes exact a cost on the ants or on the development of their colonies. We hypothesized that endophytes may play a role in their host plants’ defense against leaf-cutting ants. To measure the long-term cost to the ant colony of fungal endophytes in their forage material, we conducted a 20-week laboratory experiment to measure fungal garden development for colonies that foraged on leaves with low or high endophyte content. Results Colony mass and the fungal garden dry mass did not differ significantly between the low and high endophyte feeding treatments. There was, however, a marginally significant trend toward greater mass of fungal garden per ant worker in the low relative to the high endophyte treatment. This trend was driven by differences in the fungal garden mass per worker from the earliest samples, when leaf-cutting ants had been foraging on low or high endophyte leaf material for only 2 weeks. At two weeks of foraging, the mean fungal garden mass per worker was 77% greater for colonies foraging on leaves with low relative to high endophyte loads. Conclusions Our data suggest that the cost of endophyte presence in ant forage material may be greatest to fungal colony development in its earliest stages, when there are few workers available to forage and to clean leaf material. This coincides with a period of high mortality for incipient colonies in the field. We discuss how the endophyte-leaf-cutter ant interaction may parallel constitutive defenses in plants, whereby endophytes reduce the rate of colony development when its risk of mortality is greatest.

  20. High-Resolution Forest Canopy Height Estimation in an African Blue Carbon Ecosystem

    Science.gov (United States)

    Lagomasino, David; Fatoyinbo, Temilola; Lee, Seung-Kuk; Simard, Marc

    2015-01-01

    Mangrove forests are one of the most productive and carbon dense ecosystems that are only found at tidally inundated coastal areas. Forest canopy height is an important measure for modeling carbon and biomass dynamics, as well as land cover change. By taking advantage of the flat terrain and dense canopy cover, the present study derived digital surface models (DSMs) using stereophotogrammetric techniques on high-resolution spaceborne imagery (HRSI) for southern Mozambique. A mean-weighted ground surface elevation factor was subtracted from the HRSI DSM to accurately estimate the canopy height in mangrove forests in southern Mozambique. The mean and H100 tree height measured in both the field and with the digital canopy model provided the most accurate results with a vertical error of 1.18-1.84 m, respectively. Distinct patterns were identified in the HRSI canopy height map that could not be discerned from coarse shuttle radar topography mission canopy maps even though the mode and distribution of canopy heights were similar over the same area. Through further investigation, HRSI DSMs have the potential of providing a new type of three-dimensional dataset that could serve as calibration/validation data for other DSMs generated from spaceborne datasets with much larger global coverage. HSRI DSMs could be used in lieu of Lidar acquisitions for canopy height and forest biomass estimation, and be combined with passive optical data to improve land cover classifications.

  1. Physilogical and Biochemical Responses of Avena species to ...

    African Journals Online (AJOL)

    mk

    2013-10-23

    Oct 23, 2013 ... Seven species of oat (Avena) were evaluated for their relative drought tolerance under soil moisture stress. The plant height, leaf area production and biomass yield reduced under soil moisture stress. Among the species tested, minimum reduction in height was recorded in Avena vaviloviana, Avena.

  2. Modification of leaf morphology and anatomy as a consequence of columnar architecture in domestic apple (Malus x domestica Borkh.) trees

    DEFF Research Database (Denmark)

    Talwara, Susheela; Grout, Brian William Wilson; Toldam-Andersen, Torben Bo

    2013-01-01

    A quantitative study has been made of the modifications to leaf morphology and anatomy evident in columnar apples trees when compared to standard ones, using the original cultivar and the first columnar mutant derived from it, as well as other closely and more distantly related cultivars. Signifi......A quantitative study has been made of the modifications to leaf morphology and anatomy evident in columnar apples trees when compared to standard ones, using the original cultivar and the first columnar mutant derived from it, as well as other closely and more distantly related cultivars....... Significant increases in leaf number, area, weight per unit area, thickness and midrib angle, together with altered shape, have been recorded consistently for the leaves subtending the developing fruits of the columnar cultivars. Additionally, significant increases in leaf rolling, epicuticular wax, stomatal...... of leaf characteristics is considered in terms of the very open architectural phenotype of columnar trees and the impact this may have on the canopy microclimate that influences leaf development....

  3. Automated Leaf Tracking using Multi-view Image Sequences of Maize Plants for Leaf-growth Monitoring

    Science.gov (United States)

    Das Choudhury, S.; Awada, T.; Samal, A.; Stoerger, V.; Bashyam, S.

    2017-12-01

    Extraction of phenotypes with botanical importance by analyzing plant image sequences has the desirable advantages of non-destructive temporal phenotypic measurements of a large number of plants with little or no manual intervention in a relatively short period of time. The health of a plant is best interpreted by the emergence timing and temporal growth of individual leaves. For automated leaf growth monitoring, it is essential to track each leaf throughout the life cycle of the plant. Plants are constantly changing organisms with increasing complexity in architecture due to variations in self-occlusions and phyllotaxy, i.e., arrangements of leaves around the stem. The leaf cross-overs pose challenges to accurately track each leaf using single view image sequence. Thus, we introduce a novel automated leaf tracking algorithm using a graph theoretic approach by multi-view image sequence analysis based on the determination of leaf-tips and leaf-junctions in the 3D space. The basis of the leaf tracking algorithm is: the leaves emerge using bottom-up approach in the case of a maize plant, and the direction of leaf emergence strictly alternates in terms of direction. The algorithm involves labeling of the individual parts of a plant, i.e., leaves and stem, following graphical representation of the plant skeleton, i.e., one-pixel wide connected line obtained from the binary image. The length of the leaf is measured by the number of pixels in the leaf skeleton. To evaluate the performance of the algorithm, a benchmark dataset is indispensable. Thus, we publicly release University of Nebraska-Lincoln Component Plant Phenotyping dataset-2 (UNL-CPPD-2) consisting of images of the 20 maize plants captured by visible light camera of the Lemnatec Scanalyzer 3D high throughout plant phenotyping facility once daily for 60 days from 10 different views. The dataset is aimed to facilitate the development and evaluation of leaf tracking algorithms and their uniform comparisons.

  4. Using a Statistical Approach to Anticipate Leaf Wetness Duration Under Climate Change in France

    Science.gov (United States)

    Huard, F.; Imig, A. F.; Perrin, P.

    2014-12-01

    Leaf wetness plays a major role in the development of fungal plant diseases. Leaf wetness duration (LWD) above a threshold value is determinant for infection and can be seen as a good indicator of impact of climate on infection occurrence and risk. As LWD is not widely measured, several methods, based on physics and empirical approach, have been developed to estimate it from weather data. Many LWD statistical models do exist, but the lack of standard for measurements require reassessments. A new empirical LWD model, called MEDHI (Modèle d'Estimation de la Durée d'Humectation à l'Inra) was developed for french configuration for wetness sensors (angle : 90°, height : 50 cm). This deployment is different from what is usually recommended from constructors or authors in other countries (angle from 10 to 60°, height from 10 to 150 cm…). MEDHI is a decision support system based on hourly climatic conditions at time steps n and n-1 taking account relative humidity, rainfall and previously simulated LWD. Air temperature, relative humidity, wind speed, rain and LWD data from several sensors with 2 configurations were measured during 6 months in Toulouse and Avignon (South West and South East of France) to calibrate MEDHI. A comparison of empirical models : NHRH (RH threshold), DPD (dew point depression), CART (classification and regression tree analysis dependant on RH, wind speed and dew point depression) and MEDHI, using meteorological and LWD measurements obtained during 5 months in Toulouse, showed that the development of this new model MEHDI was definitely better adapted to French conditions. In the context of climate change, MEDHI was used for mapping the evolution of leaf wetness duration in France from 1950 to 2100 with the French regional climate model ALADIN under different Representative Concentration Pathways (RCPs) and using a QM (Quantile-Mapping) statistical downscaling method. Results give information on the spatial distribution of infection risks

  5. Leaf trait response to nutrients and herbivore exclusion across a globally replicated grassland experiment

    Science.gov (United States)

    Firn, Jennifer

    2017-04-01

    Leaf trait response to nutrients and herbivore exclusion across a globally replicated grassland experiment Jennifer Firn1, James McGree2, Eric Lind3, Elizabeth Borer3, Eric Seabloom3, Lauren Sullivan3, Kimberly Lapierre4 and the Nutrient Network 1Queensland University of Technology (QUT), School of Earth, Environmental and Biological Sciences, Science and Engineering Faculty, Brisbane, QLD, 4001 Australia 2Queensland University of Technology (QUT), School of Mathematical Sciences, Science and Engineering Faculty, Brisbane, QLD, 4001 Australia 3Universtiy of Minnesota, Department of Ecology, Evolution, and Behavior, 1479 Gortner Avenue, 140 Gortner Laboratory, St. Paul, MN 55108 USA 4Department of integrative Biology, University of California, Berkeley, CA 94720 USA Functional trait research has developed with the aim of finding general patterns in how the function of plant assemblages changes with respect to different land-uses. Most studies have compared sites within and across regions with variations in land-use history, but not necessarily with standardized treatments in an experimental framework. The trends that have emerged from this research is that characteristics of leaf traits such as specific leaf area (SLA) correlate with carbon acquisition strategies known to influence ecosystem functioning. SLA has been found to represent a plant's investment in growing light-capturing area per dry mass content. Species with a relatively high SLA tend to have a higher rate of return on the resources invested into making tissue (cheaper leaves in terms of energy and resources needed to produce them) when compared to species with a lower SLA (more expensive leaves to produce). Few studies have examined quantitatively measured traits in an experimental framework. The Nutrient Network experiment, globally distributed experiment, presents a unique opportunity to examine the response of functional traits across grassland ecosystems characterised by a diverse range of

  6. Height Growth of Mahogany Seedlings

    Science.gov (United States)

    C. B. Briscoe; R. W. Nobles

    1962-01-01

    Since the recognition of natural hybridization of small-leaf (West Indies) mahogany (Swietenia mahagoni Jacq.) with bigleaf (Honduras) mahogany (S. macrophylla King) there has been conjecture about their relative growth rates. One would expect small-leaf to be the fastest growing on dry sites, the hybrids to be fastest on intermediate sites, and bigleaf to excel on wet...

  7. Influence of micro-topography and crown characteristics on tree height estimations in tropical forests based on LiDAR canopy height models

    Science.gov (United States)

    Alexander, Cici; Korstjens, Amanda H.; Hill, Ross A.

    2018-03-01

    Tree or canopy height is an important attribute for carbon stock estimation, forest management and habitat quality assessment. Airborne Laser Scanning (ALS) based on Light Detection and Ranging (LiDAR) has advantages over other remote sensing techniques for describing the structure of forests. However, sloped terrain can be challenging for accurate estimation of tree locations and heights based on a Canopy Height Model (CHM) generated from ALS data; a CHM is a height-normalised Digital Surface Model (DSM) obtained by subtracting a Digital Terrain Model (DTM) from a DSM. On sloped terrain, points at the same elevation on a tree crown appear to increase in height in the downhill direction, based on the ground elevations at these points. A point will be incorrectly identified as the treetop by individual tree crown (ITC) recognition algorithms if its height is greater than that of the actual treetop in the CHM, which will be recorded as the tree height. In this study, the influence of terrain slope and crown characteristics on the detection of treetops and estimation of tree heights is assessed using ALS data in a tropical forest with complex terrain (i.e. micro-topography) and tree crown characteristics. Locations and heights of 11,442 trees based on a DSM are compared with those based on a CHM. The horizontal (DH) and vertical displacements (DV) increase with terrain slope (r = 0.47 and r = 0.54 respectively, p tree height are up to 16.6 m on slopes greater than 50° in our study area in Sumatra. The errors in locations (DH) and tree heights (DV) are modelled for trees with conical and spherical tree crowns. For a spherical tree crown, DH can be modelled as R sin θ, and DV as R (sec θ - 1). In this study, a model is developed for an idealised conical tree crown, DV = R (tan θ - tan ψ), where R is the crown radius, and θ and ψ are terrain and crown angles respectively. It is shown that errors occur only when terrain angle exceeds the crown angle, with the

  8. The importance of leaf BRDF in forest canopy bidirectional reflectance : a case study using simulated canopy architecture and PBRT ray tracing

    NARCIS (Netherlands)

    Biliourios, D.; Van der Zande, D.; Verstraeten, W.W.; Stuckens, J.; Muys, B.; Dutre, Ph.; Coppin, P.

    2013-01-01

    Two Fagus sylvatica L. stands with different Leaf Area Index and similar planophile Leaf Angle Distribution were created with L-systems based plant growth software and rendered using both Lambertian and Bousquet’s model leaf Bidirectional Reflectance Distribution Function (BRDF) assumption. During

  9. Estimates of leaf area index from spectral reflectance of wheat under different cultural practices and solar angle

    Science.gov (United States)

    Asrar, G.; Kanemasu, E. T.; Yoshida, M.

    1985-01-01

    The influence of management practices and solar illumination angle on the leaf area index (LAI) was estimated from measurements of wheat canopy reflectance evaluated by two methods, a regression formula and an indirect technique. The date of planting and the time of irrigation in relation to the stage of plant growth were found to have significant effects on the development of leaves in spring wheat. A reduction in soil moisture adversely affected both the duration and magnitude of the maximum LAI for late planting dates. In general, water stress during vegetative stages resulted in a reduction in maximum LAI, while water stress during the reproductive period shortened the duration of green LAI in spring wheat. Canopy geometry and solar angle also affected the spectral properties of the canopies, and hence the estimated LAI. Increase in solar zenith angles resulted in a general increase in estimated LAI obtained from both methods.

  10. Limited acclimation in leaf anatomy to experimental drought in tropical rainforest trees.

    Science.gov (United States)

    Binks, Oliver; Meir, Patrick; Rowland, Lucy; da Costa, Antonio Carlos Lola; Vasconcelos, Steel Silva; de Oliveira, Alex Antonio Ribeiro; Ferreira, Leandro; Mencuccini, Maurizio

    2016-12-01

    Dry periods are predicted to become more frequent and severe in the future in some parts of the tropics, including Amazonia, potentially causing reduced productivity, higher tree mortality and increased emissions of stored carbon. Using a long-term (12 year) through-fall exclusion (TFE) experiment in the tropics, we test the hypothesis that trees produce leaves adapted to cope with higher levels of water stress, by examining the following leaf characteristics: area, thickness, leaf mass per area, vein density, stomatal density, the thickness of palisade mesophyll, spongy mesophyll and both of the epidermal layers, internal cavity volume and the average cell sizes of the palisade and spongy mesophyll. We also test whether differences in leaf anatomy are consistent with observed differential drought-induced mortality responses among taxa, and look for relationships between leaf anatomy, and leaf water relations and gas exchange parameters. Our data show that trees do not produce leaves that are more xeromorphic in response to 12 years of soil moisture deficit. However, the drought treatment did result in increases in the thickness of the adaxial epidermis (TFE: 20.5 ± 1.5 µm, control: 16.7 ± 1.0 µm) and the internal cavity volume (TFE: 2.43 ± 0.50 mm 3 cm -2 , control: 1.77 ± 0.30 mm 3 cm -2 ). No consistent differences were detected between drought-resistant and drought-sensitive taxa, although interactions occurred between drought-sensitivity status and drought treatment for the palisade mesophyll thickness (P = 0.034) and the cavity volume of the leaves (P = 0.025). The limited response to water deficit probably reflects a tight co-ordination between leaf morphology, water relations and photosynthetic properties. This suggests that there is little plasticity in these aspects of plant anatomy in these taxa, and that phenotypic plasticity in leaf traits may not facilitate the acclimation of Amazonian trees to the predicted future reductions in dry

  11. Growth of the C4 dicot Flaveria bidentis: photosynthetic acclimation to low light through shifts in leaf anatomy and biochemistry.

    Science.gov (United States)

    Pengelly, Jasper J L; Sirault, Xavier R R; Tazoe, Youshi; Evans, John R; Furbank, Robert T; von Caemmerer, Susanne

    2010-09-01

    In C(4) plants, acclimation to growth at low irradiance by means of anatomical and biochemical changes to leaf tissue is considered to be limited by the need for a close interaction and coordination between bundle sheath and mesophyll cells. Here differences in relative growth rate (RGR), gas exchange, carbon isotope discrimination, photosynthetic enzyme activity, and leaf anatomy in the C(4) dicot Flaveria bidentis grown at a low (LI; 150 micromol quanta m(2) s(-1)) and medium (MI; 500 micromol quanta m(2) s(-1)) irradiance and with a 12 h photoperiod over 36 d were examined. RGRs measured using a 3D non-destructive imaging technique were consistently higher in MI plants. Rates of CO(2) assimilation per leaf area measured at 1500 micromol quanta m(2) s(-1) were higher for MI than LI plants but did not differ on a mass basis. LI plants had lower Rubisco and phosphoenolpyruvate carboxylase activities and chlorophyll content on a leaf area basis. Bundle sheath leakiness of CO(2) (phi) calculated from real-time carbon isotope discrimination was similar for MI and LI plants at high irradiance. phi increased at lower irradiances, but more so in MI plants, reflecting acclimation to low growth irradiance. Leaf thickness and vein density were greater in MI plants, and mesophyll surface area exposed to intercellular airspace (S(m)) and bundle sheath surface area per unit leaf area (S(b)) measured from leaf cross-sections were also both significantly greater in MI compared with LI leaves. Both mesophyll and bundle sheath conductance to CO(2) diffusion were greater in MI compared with LI plants. Despite being a C(4) species, F. bidentis is very plastic with respect to growth irradiance.

  12. A perspective on the importance of within-tree variation in mortality risk for a leaf-mining insect

    Directory of Open Access Journals (Sweden)

    C. Low

    2012-05-01

    Full Text Available Within-tree variation in abiotic conditions can create a mosaic of fitness gradients for herbivorous insects. To explore these effects, we quantified the patterns of mortality of the solitary oak leafminer, Cameraria hamadryadella (Lepidoptera: Gracillariidae, which lives within leaves of white oak, Quercus alba. We found differential patterns of survival and larval feeding rate within the tree and in association with several abiotic factors: light levels, leaf nitrogen content, and canopy height. We suggest that the leaf scale microhabitat conditions are fundamental to plant-herbivore-enemy interactions because of the differential fitness effects on herbivores. Such effects would be missed by studies that average effects by whole plants. Our study population of C. hamadryadella is located within the Orland E. White State Arboretum of Virginia in Boyce, Virginia, USA.

  13. [Growth and developmental rhythm of Scrophularia ningpoensisin southwest middle mountain area of China].

    Science.gov (United States)

    Zhang, Xue; Chen, Da-Xia; Li, Long-Yun; Yang, Xian; Song, Xu-Hong

    2014-10-01

    Plant samples were collected and investigated periodically. According to the growth of different parts and the characteristics of dry substance accumulation of Scrophularia ningpoensis, the development of S. ningpoensis could be divided into four stages: seeding stage, stem and leaf growth stage, expanding period of root tubers, and dry substance accumulation stage of root tuber. Leaf numbers of S. ningpoensis grew gradually from one at first to 370 at the final stage, main stem leaf were 50 pieces. Leaf size increasesed with the fastest growth at the stem and leaf growth stage, average daily increase amount was 225 cm2. By the middle of August, leaf size reached to 16,270 cm2. Leaf area indexrose sharply in the seeding stage, and remained above 8 among stem and leaf growth stage and expanding period of root tubers, and rapidly reduced to zero in the stage of dry substance accumulation of root tuber. Leaf area ratio has a tendency of obvious dropping. The net assimilation rate had a small change ranges, two small peak were seeding stage and dry substance accumulation of root tuber. The value of specific leaf area was higher in seeding stage, and in the earlier stage of dry substance accumulation of root tuber. Relative growth rate changed with large ranges, higher in seeding stage, rapid decrease in stem and leaf growth stage, rose in expanding period of root tubers, and declined again in the stage of dry substance accumulation of root tuber. Crop growth rate was higher in the first and last stages, and smaller in interim stage. The growth parameters of S. ningpoensis such as relative growth rate, net assimilation rate, leaf area index, leaf area ratio, specific leaf area, crop growth rate changed along with the growth. The rule of dry matter accumulation was as follows: the dry matter increased slowly during the seeding stage and speeded up in the middle and late stages, and in dry substance accumulation of root tuber increased slower, the growth of dry matter all

  14. Height-Deterministic Pushdown Automata

    DEFF Research Database (Denmark)

    Nowotka, Dirk; Srba, Jiri

    2007-01-01

    We define the notion of height-deterministic pushdown automata, a model where for any given input string the stack heights during any (nondeterministic) computation on the input are a priori fixed. Different subclasses of height-deterministic pushdown automata, strictly containing the class...... of regular languages and still closed under boolean language operations, are considered. Several of such language classes have been described in the literature. Here, we suggest a natural and intuitive model that subsumes all the formalisms proposed so far by employing height-deterministic pushdown automata...

  15. Características do pasto e acúmulo de forragem em capim-tanzânia submetido a alturas de manejo do pasto Sward characteristics and herbage accumulation of Tanzania grass submitted to sward heights

    Directory of Open Access Journals (Sweden)

    Marcos Weber do Canto

    2008-03-01

    Full Text Available O objetivo deste experimento foi avaliar alturas de manejo do pasto (20, 40, 60 e 80 cm em capim-tanzânia (Panicum maximum Jacq., em regime de lotação contínua, nas características do dossel e acúmulo de matéria seca. Os animais utilizados foram novilhos Nelore (Bos indicus, e a taxa de lotação foi variável. Foram avaliados: a massa de forragem, a massa de lâmina de folha verde, a razão folha:colmo, a composição morfológica e a taxa de acúmulo de matéria seca. O delineamento experimental foi inteiramente casualizado, com duas repetições. A massa de forragem aumentou linearmente com o aumento da altura do pasto. As médias de massa de forragem foram 2.767, 3.105, 3.657 e 4.436 kg ha-1, respectivamente, para as alturas de 20, 40, 60 e 80 cm. As taxas de acúmulo de matéria seca, a 20, 40, 60 e 80 cm, foram, respectivamente, 104, 108, 90 e 81 kg ha-1 por dia, o que indica que houve redução dessas taxas com a elevação da altura do pasto. A razão folha:colmo decresceu linearmente com o aumento da altura do pasto. Pastagens de capim-tanzânia, sob lotação contínua ao final da primavera e durante o verão, devem ser utilizadas entre 40 e 60 cm de altura.The objective of this experiment was to evaluate different sward height (20, 40, 60 e 80 cm in Tanzania grass (Panicum maximum Jacq. pastures managed under continuous stocking. The animals used were Nellore steers, and the control of sward height was done with put-and-take techniques. Evaluations were made for: forage mass, green leaf mass, leaf:stem ratio, morphological composition and dry matter accumulation rate. The experimental design was completely randomized with two replications. Forage mass increased linearly with sward height with overall mean of 2,767, 3,105, 3,657 and 4,436 kg ha-1 at sward heights 20, 40, 60 and 80 cm, respectively. Rates of dry matter accumulation decreased with increasing sward heights and were 104, 108, 90 and 81 kg ha-1 per day for sward

  16. Assimilation of Soil Wetness Index and Leaf Area Index into the ISBA-A-gs land surface model: grassland case study

    Directory of Open Access Journals (Sweden)

    A. L. Barbu

    2011-07-01

    Full Text Available The performance of the joint assimilation in a land surface model of a Soil Wetness Index (SWI product provided by an exponential filter together with Leaf Area Index (LAI is investigated. The data assimilation is evaluated with different setups using the SURFEX modeling platform, for a period of seven years (2001–2007, at the SMOSREX grassland site in southwestern France. The results obtained with a Simplified Extended Kalman Filter demonstrate the effectiveness of a joint data assimilation scheme when both SWI and Leaf Area Index are merged into the ISBA-A-gs land surface model. The assimilation of a retrieved Soil Wetness Index product presents several challenges that are investigated in this study. A significant improvement of around 13 % of the root-zone soil water content is obtained by assimilating dimensionless root-zone SWI data. For comparison, the assimilation of in situ surface soil moisture is considered as well. A lower impact on the root zone is noticed. Under specific conditions, the transfer of the information from the surface to the root zone was found not accurate. Also, our results indicate that the assimilation of in situ LAI data may correct a number of deficiencies in the model, such as low LAI values in the senescence phase by using a seasonal-dependent error definition for background and observations. In order to verify the specification of the errors for SWI and LAI products, a posteriori diagnostics are employed. This approach highlights the importance of the assimilation design on the quality of the analysis. The impact of data assimilation scheme on CO2 fluxes is also quantified by using measurements of net CO2 fluxes gathered at the SMOSREX site from 2005 to 2007. An improvement of about 5 % in terms of rms error is obtained.

  17. Child Height and Maternal Health Care Knowledge in Mozambique

    DEFF Research Database (Denmark)

    Van Den Broeck, Katleen

    Stunting prevalence rates in Mozambique are very high (41 percent), especially in rural areas (46 percent). Recent research shows that consumption growth alone will not be sufficient to solve the problem of malnutrition. To investigate the role of additional determinants I use a two-stage quantil...... of education and community health care facilities in rural areas and positively affect the height of the most severely stunted children...

  18. Short-term effects of fertilization on photosynthesis and leaf morphology of field-grown loblolly pine following long-term exposure to elevated CO2 concentration

    International Nuclear Information System (INIS)

    Maier, C.A.; Palmroth, S.; Ward, E.

    2008-01-01

    This study examined the effects of an initial nitrogen (N) fertilizer application on the upper-canopy needle morphology and gas exchange of a loblolly pine tree exposed to elevated carbon dioxide (CO 2 ) concentrations over a period of 9 years. Plots in the study were split, and one half of each plot was fertilized with 112 kg ha -1 of elemental N. Measurements included needle length, mass per unit area, N concentrations on a mass and area basis, light-saturated net photosynthesis per unit leaf area, and per unit mass and leaf conductance. Results of the study showed that fertilization had little impact on needle length, mass per unit area, or leaf conductance. Results suggested that although both needle age classes accumulated N following fertilization, current-year foliage incorporated N into its photosynthetic machinery, while 1-year old foliage stored N. No significant interactions were observed between elevated CO 2 and light-saturated net photosynthesis per unit leaf area. The study found few fertilization and CO 2 interaction effects on leaf physiology and morphology. 54 refs., 3 tabs., 3 figs

  19. Global Distribution of Planetary Boundary Layer Height Derived from CALIPSO

    Science.gov (United States)

    Huang, J.

    2015-12-01

    The global distribution of planetary boundary layer (PBL) height, which was estimated from the attenuated back-scatter observations of Cloud-Aerosol Lidar and Infrared Pathfinder Satellite Observation (CALIPSO), is presented. In general, the PBL is capped by a temperature inversion that tends to trap moisture and aerosols. The gradient of back-scatter observed by lidar is almost always associated with this temperature inversion and the simultaneous decrease of moisture content. Thus, the PBL top is defined as the location of the maximum aerosol scattering gradient, which is analogous to the more conventional thermodynamic definition. The maximum standard deviation method, developed by Jordan et al. (2010), is modified and used to derive the global PBL heights. The derived PBL heights are not only consistent with the results of McGrath-Spangler and Denning (2012) but also agree well with the ground-based lidar measurements. It is found that the correlation between CALIPSO and the ground-based lidar was 0.73. The seasonal mean patterns from 4-year mid-day PBL heights over global are demonstrated. Also it is found that the largest PBL heights occur over the Tibetan Plateau and the coastal areas. The smallest PBL heights appear in the Tarim Basin and the northeast of China during the local winter. The comparison of PBL heights from CALIPSO and ECMWF under different land-cover conditions showed that, over ocean and forest surface, the PBL height estimated from the CALIPSO back-scatter climatology is larger than the ones estimated from ECMWF data. However, the PBL heights from ECMWF, over grass land and bare land surface in spring and summer are larger than the ones from CALIPSO.

  20. Measurement for the MLC leaf velocity profile by considering the leaf leakage using a radiographic film

    International Nuclear Information System (INIS)

    Chow, James C L; Grigorov, Grigor N

    2006-01-01

    A method to measure the velocity profile of a multi-leaf collimator (MLC) leaf along its travel range using a radiographic film is reported by considering the intra-leaf leakage. A specific dynamic MLC field with leaves travelling from the field edge to the isocentre line was designed. The field was used to expose a radiographic film, which was then scanned, and the dose profile along the horizontal leaf axis was measured. The velocity at a sampling point on the film can be calculated by considering the horizontal distance between the sampling point and the isocentre line, dose at the sampling point, dose rate of the linear accelerator, the total leaf travel time from the field edge to isocentre line and the pre-measured dose rate of leaf leakage. With the leaf velocities and velocity profiles for all MLC leaves measured routinely, a comprehensive and simple QA for the MLC can be set up to test the consistency of the leaf velocity performance which is essential to the IMRT delivery using a sliding window technique. (note)