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Sample records for fern cave lava

  1. Rock paintings in Fern Cave, Lava Beds National Monument, California: Not the 1054 A.D. (Crab Nebula) Supernova

    International Nuclear Information System (INIS)

    Armitage, R.A.; Hyman, M.; Rowe, M. W.

    1997-01-01

    On July 4, 1054 A.D. a supernova brighter than Venus appeared in the sky, remaining visible for approximately 23 days and 650 nights. It was chronicled in five independent historic accounts, four in China and one in Japan. For at least 40 years investigators have attributed certain distinctive rock paintings and carvings in the western United States as recordings of the 1054 A.D. supernova. More than twenty such depictions (circle or star-like symbols and a crescent) have been located. Two panels of rock paintings in Lava Beds National Monument, California, one at Fern Cave and one at Symbol Bridge, were listed as recording the 1054 A.D. supernova. The only direct means of assessing the likelihood that a 'supernova' representation records the 1054 A.D. event is to date the rock painting or carving. At Texas A and M University, was developed a plasma-chemical extraction technique that permits to analyze the 14 C in rock paintings, whether the pigments used were charcoal or inorganic Fe- and Mn- oxides and hydroxides with organic binder/vehicles. This paper presents direct 14 C age estimates on a rock painting suggested to represent the 1054 A.D. supernova. Charcoal pigment samples were collected from three figures in proximity at Fern Cave: a crescent pointing downward and two nearby circles, one above and one below the crescent. The AMS 14 C analysis on each sample using this technique show that these images do not represent the 1054 A.D. supernova

  2. The genesis of a lava cave in the Deccan Volcanic Province (Maharashtra, India

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    Nikhil R. Pawar

    2016-01-01

    Full Text Available Lava tubes and channels forming lava distributaries have been recognized from different parts of western Deccan Volcanic Province (DVP. Openings of smaller dimension have been documented from the pāhoehoe flows around Pune, in the western DVP. A small lava cave is exposed in Ghoradeshwar hill, near Pune. Detailed field studies of the physical characteristics, structure and morphology of the flows hosting the lava tube has been carried out. This is the first detailed documentation of a lava cave from the DVP. The lava cave occurs in a compound pāhoehoe flow of Karla Formation, characterized by the presence of lobes, toes and small scale features like squeeze-ups. Field observations and measurements reveal that the dimensions of the cave are small, with low roof and a maximum width of 108 cm. The cave morphology along the 20 m passage varies from circular to semi-circular, with a twilight zone to the north. The gentle micro-topography at Ghoradeshwar controlled the advancement of pāhoehoe lobes and toes within the sheet lobe. The pre-flow gradients towards the north led to the progression of flow from the east, where the cave opening is presently seen. Dimensions and related morphology of the lava cave suggest that it can be best described as a small sub-crustal cave formed by draining of an inflated of pāhoehoe lava lobe. At Ghoradeshwar, besides the natural lava cave, Buddhist caves carved in pāhoehoe lava flows are also observed, indicating that early man took advantage of the existing openings in pāhoehoe flows and sculpted the caves to suit their requirements.

  3. Erosion by flowing lava: Geochemical evidence in the Cave Basalt, Mount St. Helens, Washington

    Science.gov (United States)

    Williams, D.A.; Kadel, S.D.; Greeley, R.; Lesher, C.M.; Clynne, M.A.

    2004-01-01

    We sampled basaltic lava flows and underlying dacitic tuff deposits in or near lava tubes of the Cave Basalt, Mount St. Helens, Washington to determine whether the Cave Basalt lavas contain geochemical evidence of substrate contamination by lava erosion. The samples were analyzed using a combination of wavelength-dispersive X-ray fluorescence spectrometry and inductively-coupled plasma mass spectrometry. The results indicate that the oldest, outer lava tube linings in direct contact with the dacitic substrate are contaminated, whereas the younger, inner lava tube linings are uncontaminated and apparently either more evolved or enriched in residual liquid. The most heavily contaminated lavas occur closer to the vent and in steeper parts of the tube system, and the amount of contamination decreases with increasing distance downstream. These results suggest that erosion by lava and contamination were limited to only the initially emplaced flows and that erosion was localized and enhanced by vigorous laminar flow over steeper slopes. After cooling, the initial Cave Basalt lava flows formed an insulating lining within the tubes that prevented further erosion by later flows. This interpretation is consistent with models of lava erosion that predict higher erosion rates closer to sources and over steeper slopes. A greater abundance of xenoliths and xenocrysts relative to xenomelts in hand samples indicates that mechanical erosion rather than thermal erosion was the dominant erosional process in the Cave Basalt, but further sampling and petrographic analyses must be performed to verify this hypothesis. ?? Springer-Verlag 2003.

  4. Pisgah Lava Cave Communication Test: Science Case Study for the Networked Constellations Initiative

    Science.gov (United States)

    Belov, K.; Ellison, D.; Fraeman, A.

    2017-01-01

    As part of the science case study for the Networked Constellations initiative, a team of JPL scientists explore the possibility of a mission to study the lava caves on Mars. Natural caves on Mars and the Moon present a unique opportunity to learn about the planetary geology and to provide a shelter for human explorers. Due to power and communication challenges, a network of assets has significant advantages over a single asset sent inside a cave. However, communication between the assets and the data downlink present significant difficulties due to the presence of rough walls, boulders, and other obstacles with unknown dielectric constant inside a typical cave, disturbing the propagation of the radio waves. A detailed study is needed to establish the limitations of the current communication technologies and to develop requirements for the new communication technology applicable to the cave environment. On May 4 of 2017, Konstantin Belov, Doug Ellison, and Abby Fraeman visited a lava cave in Pisgah, CA. The purpose of the visit was to build a 3D map of the cave, which could be used to create a model of radio wave propagation, and to conduct a series of communication tests using off-the-shelf equipment to verify the in-cave communication challenges. This experiment should be considered as a simple 'proof of concept' and is the subject of this report.

  5. Wintering bats of the upper Snake River Plain: occurrence in lava-tube caves

    Energy Technology Data Exchange (ETDEWEB)

    Genter, D.L.

    1986-04-30

    Distribution and habitat selection of hibernating bats at the Idaho National Engineering Laboratory (INEL) and adjacent area are reported. Exploration of over 30 lava-tube caves revealed that two species, Myotis leibii and Plecotus townsendii, hibernate in the upper Snake River Plain. Five species, M. lucifugus, M. evotis, Eptesicus fuscus, Lasionycteris noctivagans, and Lasiurus cinereus are considered migratory. Myotis leibii and P. townsendii hibernate throughout much of the area, occasionally in mixed-species groups. Myotis leibii uses the dark and protected regions of the cave, usually wedged into tiny pockets and crevices near or at the highest portion of the ceiling. Individuals of P. townsendii may be found at any height or depth in the cave. Temperature appears to be primary limiting factor in habitat selection. Myotis leibii was found in significantly cooler air temperatures than P. townsendii. Neither species tolerated continuous temperatures below 1.5 C. Relative humidity does not seem to be a significant factor in the distribution or habitat selection of the two species in lava-tube caves. 18 references, 1 figure, 1 table.

  6. Selected caves and lava-tube systems in and near Lava Beds National Monument, California

    Science.gov (United States)

    Waters, Aaron Clement; Donnelly-Nolan, Julie M.; Rogers, Bruce W.

    1990-01-01

    Lava Beds National Monument (fig. 1) lies on the north slope of the huge Medicine Lake shield (fig. 2), a complex volcanic edifice of greater volume than the steep-sided Mount Shasta volcanic cone, which towers as a snowclad land mark 40 mi southwest of the monument (fig. 3).

  7. Assessing the origin of unusual organic formations in lava caves from Canary Islands (Spain)

    Science.gov (United States)

    Miller, Ana Z.; de la Rosa, Jose M.; Garcia-Sanchez, Angela M.; Pereira, Manuel F. C.; Jurado, Valme; Fernández, Octavio; Knicker, Heike; Saiz-Jimenez, Cesareo

    2016-04-01

    Lava tubes, like other caves, contain a variety of speleothems formed in the initial stage of a lava tube formation or due to leaching and subsequent precipitation of secondary minerals. Primary and secondary mineral formations in lava caves are mainly composed of silicate minerals, although secondary minerals common in limestone caves have been also reported in this type of caves. In addition, unusual colored deposits have been found on the walls and ceilings of lava tubes, some of them of unknown origin and composition. A brown to black-colored mud-like deposits was observed in "Llano de los Caños" Cave, La Palma Island, Canary Islands, Spain. These black deposits coat the wall and ceiling of the lava tube where sub-horizontal fractures occur. FESEM-EDS, X-ray micro-computed tomography and mineralogical analyses were conducted for morphological, 3D microstructural and compositional characterization of these unusual speleothem samples. These techniques revealed that they are mainly composed of amorphous materials, suggesting an organic carbon composition. Hence, analytical pyrolysis (Py-GC/MS), solid-state 13C Nuclear Magnetic Resonance (NMR) and stable isotope analysis were applied to assess the nature and origin of the black deposits. The combination of these analytical tools permits the identification of specific biomarkers (di- and triterpenoids) for tracing the potential sources of the organic compounds in the speleothems. For comparison purposes, samples from the topsoil and overlaying vegetation were also analyzed. Chromatograms resulting from the Py-GC/MS showed an abundance of polysaccharides, lipids and terpenoids typically derived from the vegetation of the area (Erica arborea). In addition, levoglucosan, polycyclic aromatic hydrocarbons and N-containing heterocyclic compounds were detected. They probably derived from the leaching of charred vegetation resulting from a wildfire occurred in the area in 2012. The lack of the typical pattern of odd

  8. Lava cave microbial communities within mats and secondary mineral deposits: implications for life detection on other planets.

    Science.gov (United States)

    Northup, D E; Melim, L A; Spilde, M N; Hathaway, J J M; Garcia, M G; Moya, M; Stone, F D; Boston, P J; Dapkevicius, M L N E; Riquelme, C

    2011-09-01

    Lava caves contain a wealth of yellow, white, pink, tan, and gold-colored microbial mats; but in addition to these clearly biological mats, there are many secondary mineral deposits that are nonbiological in appearance. Secondary mineral deposits examined include an amorphous copper-silicate deposit (Hawai'i) that is blue-green in color and contains reticulated and fuzzy filament morphologies. In the Azores, lava tubes contain iron-oxide formations, a soft ooze-like coating, and pink hexagons on basaltic glass, while gold-colored deposits are found in lava caves in New Mexico and Hawai'i. A combination of scanning electron microscopy (SEM) and molecular techniques was used to analyze these communities. Molecular analyses of the microbial mats and secondary mineral deposits revealed a community that contains 14 phyla of bacteria across three locations: the Azores, New Mexico, and Hawai'i. Similarities exist between bacterial phyla found in microbial mats and secondary minerals, but marked differences also occur, such as the lack of Actinobacteria in two-thirds of the secondary mineral deposits. The discovery that such deposits contain abundant life can help guide our detection of life on extraterrestrial bodies.

  9. Cultivation and complete genome sequencing of Gloeobacter kilaueensis sp. nov., from a lava cave in Kīlauea Caldera, Hawai'i.

    Directory of Open Access Journals (Sweden)

    Jimmy H W Saw

    Full Text Available The ancestor of Gloeobacter violaceus PCC 7421(T is believed to have diverged from that of all known cyanobacteria before the evolution of thylakoid membranes and plant plastids. The long and largely independent evolutionary history of G. violaceus presents an organism retaining ancestral features of early oxygenic photoautotrophs, and in whom cyanobacteria evolution can be investigated. No other Gloeobacter species has been described since the genus was established in 1974 (Rippka et al., Arch Microbiol 100:435. Gloeobacter affiliated ribosomal gene sequences have been reported in environmental DNA libraries, but only the type strain's genome has been sequenced. However, we report here the cultivation of a new Gloeobacter species, G. kilaueensis JS1(T, from an epilithic biofilm in a lava cave in Kīlauea Caldera, Hawai'i. The strain's genome was sequenced from an enriched culture resembling a low-complexity metagenomic sample, using 9 kb paired-end 454 pyrosequences and 400 bp paired-end Illumina reads. The JS1(T and G. violaceus PCC 7421(T genomes have little gene synteny despite sharing 2842 orthologous genes; comparing the genomes shows they do not belong to the same species. Our results support establishing a new species to accommodate JS1(T, for which we propose the name Gloeobacter kilaueensis sp. nov. Strain JS1(T has been deposited in the American Type Culture Collection (BAA-2537, the Scottish Marine Institute's Culture Collection of Algae and Protozoa (CCAP 1431/1, and the Belgian Coordinated Collections of Microorganisms (ULC0316. The G. kilaueensis holotype has been deposited in the Algal Collection of the US National Herbarium (US# 217948. The JS1(T genome sequence has been deposited in GenBank under accession number CP003587. The G+C content of the genome is 60.54 mol%. The complete genome sequence of G. kilaueensis JS1(T may further understanding of cyanobacteria evolution, and the shift from anoxygenic to oxygenic photosynthesis.

  10. Comparing flow-through and static ice cave models for Shoshone Ice Cave

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    Kaj E. Williams

    2015-05-01

    Full Text Available In this paper we suggest a new ice cave type: the “flow-through” ice cave. In a flow-through ice cave external winds blow into the cave and wet cave walls chill the incoming air to the wet-bulb temperature, thereby achieving extra cooling of the cave air. We have investigated an ice cave in Idaho, located in a lava tube that is reported to have airflow through porous wet end-walls and could therefore be a flow-through cave. We have instrumented the site and collected data for one year. In order to determine the actual ice cave type present at Shoshone, we have constructed numerical models for static and flow-through caves (dynamic is not relevant here. The models are driven with exterior measurements of air temperature, relative humidity and wind speed. The model output is interior air temperature and relative humidity. We then compare the output of both models to the measured interior air temperatures and relative humidity. While both the flow-through and static cave models are capable of preserving ice year-round (a net zero or positive ice mass balance, both models show very different cave air temperature and relative humidity output. We find the empirical data support a hybrid model of the static and flow-through models: permitting a static ice cave to have incoming air chilled to the wet-bulb temperature fits the data best for the Shoshone Ice Cave.

  11. Between Two Fern Genomes

    Science.gov (United States)

    2014-01-01

    Ferns are the only major lineage of vascular plants not represented by a sequenced nuclear genome. This lack of genome sequence information significantly impedes our ability to understand and reconstruct genome evolution not only in ferns, but across all land plants. Azolla and Ceratopteris are ideal and complementary candidates to be the first ferns to have their nuclear genomes sequenced. They differ dramatically in genome size, life history, and habit, and thus represent the immense diversity of extant ferns. Together, this pair of genomes will facilitate myriad large-scale comparative analyses across ferns and all land plants. Here we review the unique biological characteristics of ferns and describe a number of outstanding questions in plant biology that will benefit from the addition of ferns to the set of taxa with sequenced nuclear genomes. We explain why the fern clade is pivotal for understanding genome evolution across land plants, and we provide a rationale for how knowledge of fern genomes will enable progress in research beyond the ferns themselves. PMID:25324969

  12. Astrobiology Training in Lava Tubes (ATiLT): Characterizing coralloid speleothems in basaltic lava tubes as a Mars analogue

    Science.gov (United States)

    Ni, J.; Leveille, R. J.; Douglas, P.

    2017-12-01

    Coralloid speleothems or cave corals are small mineralised nodes that can take a variety of forms, and which develop through groundwater seepage and water-rock interaction in caves. They are found commonly on Earth in a plethora of caves, including lava tubes. Since lava tubes have been identified on the surface of Mars from remotely sensed images, there has been interest in studying Earth's lava tube systems as an analogue for understanding Martian lava environments. If cave minerals were found on Mars, they could indicate past or present water-rock interaction in the Martian subsurface. Martian lava tubes could also provide insights into habitable subsurface environments as well as conditions favourable for the synthesis and preservation of biosignatures. One of the aims of the Astrobiology Training in Lava Tubes (ATiLT) project is to analyze biosignatures and paleoenvironmental indicators in secondary cave minerals, which will be looked at in-situ and compared to collected field samples. In this study, secondary mineralization in lava cave systems from Lava Beds National Monument, CA is examined. In the field, coralloid speleothems have been observed growing on all surfaces of the caves, including cave ceilings, floors, walls and overhangs. They are also observed growing adjacent to biofilms, which sometimes fill in the cracks of the coralloid nodes. Preliminary results show the presence of opal, calcite, quartz and other minor minerals in the speleothems. This study seeks to understand the formation mechanism and source of these secondary minerals, as well as determine their possible relation to the biofilms. This will be done through the analysis of the water chemistry, isotope geochemistry and microscale mineralogy.

  13. Gypsum speleothems in lava tubes from Lanzarote, Canary Islands. Did you say gypsum?

    OpenAIRE

    Huerta, Pedro; Martín-García, Rebeca; Rodríguez-Berriguete, Álvaro; Iglesia, A. la; Martín-Pérez, Andrea; Alonso-Zarza, Ana María

    2015-01-01

    Lanzarote is the easternmost island of the volcanic Canary archipielago considered together with Fuerteventura the low relief islands of the archipielago. These island receive less rain than 300 mm/year. Basaltic lava flows preserves lava tubes formed during cooling and solidification of external parts of lava, while internal parts were still hot and flowing. When lava flow stopped the lava abandoned the tubes, and the tubes preserved empty. These tubes actuate as caves and som...

  14. {sup 14}C AMS dating Yongcheon cave

    Energy Technology Data Exchange (ETDEWEB)

    Lee, J.H., E-mail: jefflee@snu.ac.kr [AMS Lab., NCIRF, Seoul National University, Seoul 151-742 (Korea, Republic of); Choe, K. [AMS Lab., NCIRF, Seoul National University, Seoul 151-742 (Korea, Republic of); Kim, J.C. [Dept. of Physics and Astronomy, Seoul National University, Seoul 151-742 (Korea, Republic of); Choi, S.H.; Kang, J.; Song, S.; Song, Y.M. [AMS Lab., NCIRF, Seoul National University, Seoul 151-742 (Korea, Republic of); Jang, J.G. [Jeju National Museum, Jeju 690-782 (Korea, Republic of)

    2013-01-15

    The biggest island in South Korea is Jeju Island, which lies 80 km south of the mainland and has one shield volcano, Mt. Halla. The volcanic island and its lava tubes were added to the world heritage list by UNESCO in 2007. Among the many lava tubes on the island, a unique cave had been accidentally found in 2005 while some workers were replacing a telephone pole. Until the discovery, it had been completely isolated from the outside by naturally-built sand blocks. Yongcheon cave is a lime-decorated lava tube showing both the properties of a volcanic lava tube and a limestone cave. This cave, about 3 km in length, is acknowledged to be the best of this type in the world and includes a large clean-water lake, lava falls, and richly developed speleothems inside it. Even though there is archaeological evidence from well preserved pottery that ancient people entered this place, the preservation of artifacts was ensured by a geological change that made later entrance difficult. We have collected charcoal samples scattered around the cave and dated them using AMS. Ages were in the range of ca. 1570-1260 BP (A.D. 340-880) and this corresponds to the Ancient Three Kingdoms and the Unified Silla era in Korean history. The {sup 14}C AMS measurement results presented in this paper on wood charcoal provide precise dates which will be very useful not only to clarify the nature of human activities in this cave but also to provide reference dates when comparing other dating methods.

  15. 14C AMS dating Yongcheon cave

    International Nuclear Information System (INIS)

    Lee, J.H.; Choe, K.; Kim, J.C.; Choi, S.H.; Kang, J.; Song, S.; Song, Y.M.; Jang, J.G.

    2013-01-01

    The biggest island in South Korea is Jeju Island, which lies 80 km south of the mainland and has one shield volcano, Mt. Halla. The volcanic island and its lava tubes were added to the world heritage list by UNESCO in 2007. Among the many lava tubes on the island, a unique cave had been accidentally found in 2005 while some workers were replacing a telephone pole. Until the discovery, it had been completely isolated from the outside by naturally-built sand blocks. Yongcheon cave is a lime-decorated lava tube showing both the properties of a volcanic lava tube and a limestone cave. This cave, about 3 km in length, is acknowledged to be the best of this type in the world and includes a large clean-water lake, lava falls, and richly developed speleothems inside it. Even though there is archaeological evidence from well preserved pottery that ancient people entered this place, the preservation of artifacts was ensured by a geological change that made later entrance difficult. We have collected charcoal samples scattered around the cave and dated them using AMS. Ages were in the range of ca. 1570-1260 BP (A.D. 340–880) and this corresponds to the Ancient Three Kingdoms and the Unified Silla era in Korean history. The 14 C AMS measurement results presented in this paper on wood charcoal provide precise dates which will be very useful not only to clarify the nature of human activities in this cave but also to provide reference dates when comparing other dating methods.

  16. Chemical deposits in volcanic caves of Argentina

    Directory of Open Access Journals (Sweden)

    Carlos Benedetto

    1998-01-01

    Full Text Available During the last Conference of the FEALC (Speleological Federation of Latin America and Caribbean Islands which was held in the town of Malargue, Mendoza, in February 1997, two volcanic caves not far from that town were visited and sampled for cave mineral studies. The first cave (Cueva del Tigre opens close to the Llancanelo lake, some 40 kms far from Malargue and it is a classical lava tube. Part of the walls and of the fallen lava blocks are covered by white translucent fibres and grains. The second visited cave is a small tectonic cavity opened on a lava bed some 100 km southward of Malargue. The cave “El Abrigo de el Manzano” is long no more than 10-12 meters with an average width of 3 meters and it hosts several bird nests, the larger of which is characterized by the presence of a relatively thick pale yellow, pale pink flowstone. Small broken or fallen samples of the secondary chemical deposits of both these caves have been collected in order to detect their mineralogical composition. In the present paper the results of the detailed mineralogical analyses carried out on the sampled material are shortly reported. In the Cueva del Tigre lava tube the main detected minerals are Sylvite, Thenardite, Bloedite and Kieserite, all related to the peculiar dry climate of that area. The flowstone of “El Abrigo de el Manzano” consists of a rather complex admixture of several minerals, the large majority of which are phosphates but also sulfates and silicates, not all yet identified. The origin of all these minerals is related to the interaction between bird guano and volcanic rock.

  17. Candidate cave entrances on Mars

    Science.gov (United States)

    Cushing, Glen E.

    2012-01-01

    This paper presents newly discovered candidate cave entrances into Martian near-surface lava tubes, volcano-tectonic fracture systems, and pit craters and describes their characteristics and exploration possibilities. These candidates are all collapse features that occur either intermittently along laterally continuous trench-like depressions or in the floors of sheer-walled atypical pit craters. As viewed from orbit, locations of most candidates are visibly consistent with known terrestrial features such as tube-fed lava flows, volcano-tectonic fractures, and pit craters, each of which forms by mechanisms that can produce caves. Although we cannot determine subsurface extents of the Martian features discussed here, some may continue unimpeded for many kilometers if terrestrial examples are indeed analogous. The features presented here were identified in images acquired by the Mars Odyssey's Thermal Emission Imaging System visible-wavelength camera, and by the Mars Reconnaissance Orbiter's Context Camera. Select candidates have since been targeted by the High-Resolution Imaging Science Experiment. Martian caves are promising potential sites for future human habitation and astrobiology investigations; understanding their characteristics is critical for long-term mission planning and for developing the necessary exploration technologies.

  18. The omnipresent water fern

    NARCIS (Netherlands)

    Nierop, K.G.J.; Speelman, E.N.; de Leeuw, J.W.; Reichart, G.J.

    2011-01-01

    Several studies have reported the presence of large amounts of lignin in ubiquitously occurring species of the freshwater fern Azolla. Molecular analysis using flash pyrolysis and thermally assisted hydrolysis and methylation shows, however, that neither the leaves nor the roots of Azolla contain

  19. Fern of the future?

    NARCIS (Netherlands)

    Brinkhuis, H.; Bijl, P.K.

    Held in the palm of a human hand, a single specimen of Azolla filiculoides looks downright inconsequential. Even with the scaly leaves of this miniature aquatic fern spread flat, it barely spans the distance between the creases that cleave the flesh. Its fibrous root tendrils dangle like a lock of

  20. The morphology of ferns

    NARCIS (Netherlands)

    Holttum, R.E.

    1959-01-01

    A fern plant consists of a stem, bearing leaves and roots. The leaves (or some of them) bear dehiscent sporangia, each sporangium containing unicellular spores, which are in most cases Wind-dispersed. A spore germinates to produce a small green plant called a prothallus. The Prothallus bears sexual

  1. Drained Lava Tubes and Lobes From Eocretaceous Paraná-Etendeka Province, Brazil

    Science.gov (United States)

    Waichel, B. L.; Lima, E. F. D.; Mouro, L. D.; Briske, D. R.; Tratz, E. B.

    2017-12-01

    The identification of lava tubes in continental flood basalt provinces (CFBP) is difficult and reports of preserved drained tubes and lobes are rare. The large extension of CFBP must be related to an efficient transport of lava and tubes are the most efficient mechanism to transport lava in insulated pathways, like observed in modern volcanic fields. Looking for caves in the central portion of Paraná-Etendeka Province, we discovered drained lava tubes (4) and lobes (6) in a volcanic sequence constituted by pahoehoe flows. Lava tubes are: Casa de Pedra, Perau Branco, Dal Pae and Pinhão. The Casa de Pedra tube system is composed of two principal chambers with similar dimensions, reaching up to 10 m long and 4.0 m high connected by a narrow passage. The general form of the chamber is hemispherical, with re-entrances of ellipsoidal shape probably formed by small lava lobes and collapse structures in the roof. The second chamber is connected with three secondary lava tubes. Columns in the cave are formed when the flowing lava separates in two lava channels that join again further down the system, forming and anastomosing tube network. Lateral lava benches and lava drainings at the walls are observed in secondary tubes. The general lava flow is to SW. The Perau Branco system is composed of five tubes with ellipsoidal openings. The main features are the long tubes that emerge from the small flattened chambers. One tube is more than 20 m long, with alternating circular and flattened ellipsoidal sections. The general lava flow is to NE. Pinhão tube is spherical with 3 meters diameter and 15 m long, with lava flow orientation to NW. This tube has a bottleneck shape with linings (up to 3 cm thick), which are observed in the roof and walls. Dal Pae Tube is 10 m long with an ellipsoidal opening, bottleneck shape and orientation to NE. The lava flow directions measured in the tubes is to SW (Casa de Pedra, Pinhão) and NE (Perau Branco, Dal Pae) and this pattern is related to

  2. Cave age determination

    International Nuclear Information System (INIS)

    Anon.

    1998-01-01

    To determine the age of a cave we have to determine the age of elements which were created at the same moment when the cave was. Usually a cave is dug out by infiltration of acid waters. During this alteration process rocks free some aluminium and potassium ions. In Carlsbad and Lechuguilla caves, scientists have found alunite, this aluminium and potassium sulfate can be dated by using the carbon method of age determination. (A.C.)

  3. New species of Malaysian ferns

    NARCIS (Netherlands)

    Holttum, R.E.

    1962-01-01

    The present paper includes descriptions of several new species of ferns found among recent collections from various parts of Malaysia; also two new combinations of names of species which are of interest on account of their taxonomic history.

  4. Lunar Lava Tube Sensing

    Science.gov (United States)

    York, Cheryl Lynn; Walden, Bryce; Billings, Thomas L.; Reeder, P. Douglas

    1992-01-01

    Large (greater than 300 m diameter) lava tube caverns appear to exist on the Moon and could provide substantial safety and cost benefits for lunar bases. Over 40 m of basalt and regolith constitute the lava tube roof and would protect both construction and operations. Constant temperatures of -20 C reduce thermal stress on structures and machines. Base designs need not incorporate heavy shielding, so lightweight materials can be used and construction can be expedited. Identification and characterization of lava tube caverns can be incorporated into current precursor lunar mission plans. Some searches can even be done from Earth. Specific recommendations for lunar lava tube search and exploration are (1) an Earth-based radar interferometer, (2) an Earth-penetrating radar (EPR) orbiter, (3) kinetic penetrators for lunar lava tube confirmation, (4) a 'Moon Bat' hovering rocket vehicle, and (5) the use of other proposed landers and orbiters to help find lunar lava tubes.

  5. Introducing Kansas Lava

    Science.gov (United States)

    Gill, Andy; Bull, Tristan; Kimmell, Garrin; Perrins, Erik; Komp, Ed; Werling, Brett

    Kansas Lava is a domain specific language for hardware description. Though there have been a number of previous implementations of Lava, we have found the design space rich, with unexplored choices. We use a direct (Chalmers style) specification of circuits, and make significant use of Haskell overloading of standard classes, leading to concise circuit descriptions. Kansas Lava supports both simulation (inside GHCi), and execution via VHDL, by having a dual shallow and deep embedding inside our Signal type. We also have a lightweight sized-type mechanism, allowing for MATLAB style matrix based specifications to be directly expressed in Kansas Lava.

  6. How early ferns became trees.

    Science.gov (United States)

    Galtier, J; Hueber, F M

    2001-09-22

    A new anatomically preserved fern, discovered from the basalmost Carboniferous of Australia, shows a unique combination of very primitive anatomical characters (solid centrarch cauline protostele) with the elaboration of an original model of the arborescent habit. This plant possessed a false trunk composed of a repetitive branching system of very small stems, which established it as the oldest tree-fern known to date. The potential of this primitive zygopterid fern to produce such an unusual growth form-without real equivalent among living plants-is related to the possession of two kinds of roots that have complementary functional roles: (i) large roots produced by stems with immediate positive geotropism, strongly adapted to mechanical support and water uptake from the soil; and (ii) small roots borne either on large roots or on petiole bases for absorbing humidity inside the false trunk.

  7. Screening for ptaquiloside in ferns

    DEFF Research Database (Denmark)

    Rasmussen, Lars Holm; Pedersen, Henrik Ærenlund

    2017-01-01

    . Objective: To set-up a methodology for large-scale qualitative studies on the distribution of PTA in ferns using already available herbarium specimens as source. Methodology: PTA and the main degradation product pterosin B (PtB) were quantified in aqueous frond extracts by HPLC-DAD. PTA was quantified after...... of the analyte was obtained. Preservation of fronds in a plant press increase formation of PtB. Hence, the method is only suitable for qualitative studies. Presence of PTA and PtB were found in samples up to 50 years old. Among 21 ferns tested, the compounds were only found in Pteridium aquilinum. Conclusion...

  8. Hollow volcanic tumulus caves of Kilauea Caldera, Hawaii County, Hawaii

    Directory of Open Access Journals (Sweden)

    William R. Halliday

    1998-01-01

    Full Text Available In addition to lava tube caves with commonly noted features, sizable subcrustal spaces of several types exist on the floor of Kilauea Caldera. Most of these are formed by drainage of partially stabilized volcanic structures enlarged or formed by injection of very fluid lava beneath a plastic crust. Most conspicuous are hollow tumuli, possibly first described by Walker in 1991. Walker mapped and described the outer chamber of Tumulus E-I Cave. Further exploration has revealed that it has a hyperthermic inner room beneath an adjoining tumulus with no connection evident on the surface. Two lengthy, sinuous hollow tumuli also are present in this part of the caldera. These findings support Walkers conclusions that hollow tumuli provide valuable insights into tumulus-forming mechanisms, and provide information about the processes of emplacement of pahoehoe sheet flows.

  9. Hawaii Volcanism: Lava Forms

    Data.gov (United States)

    National Oceanic and Atmospheric Administration, Department of Commerce — Over the last several million years the Hawaiian Islands have been built of successive lava flows. They are the most recent additions in a long line of volcanoes...

  10. Clastic cave deposits in Botovskaya cave (Eastern Siberia, Russian Federation)

    Czech Academy of Sciences Publication Activity Database

    Kadlec, Jaroslav; Chadima, Martin; Lisá, Lenka; Hercman, H.; Osintsev, A.; Oberhänsli, H.

    2008-01-01

    Roč. 70, č. 3 (2008), s. 142-155 ISSN 1090-6924 Institutional research plan: CEZ:AV0Z30130516 Keywords : cave sediments * mineral magnetism * Botovskaya Cave Subject RIV: DE - Earth Magnetism, Geodesy, Geography Impact factor: 0.532, year: 2008 http://www.caves.org/pub/journal/Journal_of_Cave_and_Karst_Studies_volume_70.htm

  11. Salt ingestion caves.

    Directory of Open Access Journals (Sweden)

    Lundquist Charles A.

    2006-01-01

    Full Text Available Large vertebrate herbivores, when they find a salt-bearing layer of rock, say in a cliff face, can produce sizable voids where, overgenerations, they have removed and consumed salty rock. The cavities formed by this natural animal process constitute a uniqueclass of caves that can be called salt ingestion caves. Several examples of such caves are described in various publications. Anexample in Mississippi U.S.A., Rock House Cave, was visited by the authors in 2000. It seems to have been formed by deer orbison. Perhaps the most spectacular example is Kitum Cave in Kenya. This cave has been excavated to a length over 100 metersby elephants. An ancient example is La Cueva del Milodon in Chile, which is reported to have been excavated by the now extinctmilodon, a giant ground sloth. Still other possible examples can be cited. This class of caves deserves a careful definition. First, thecavity in rock should meet the size and other conventions of the locally accepted definition of a cave. Of course this requirement differsin detail from country to country, particularly in the matter of size. The intent is to respect the local conventions. The characteristicthat human entry is possible is judged to be a crucial property of any recognized cave definition. Second, the cavity should besignificantly the result of vertebrate animal consumption of salt-bearing rock. The defining process is that rock removed to form thecave is carried away in the digestive track of an animal. While sodium salts are expected to be the norm, other salts for which thereis animal hunger are acceptable. Also some other speleogenesis process, such as solution, should not be excluded as long as it issecondary in formation of a cave in question.

  12. Nerillidae (Annelida) from the Corona lava tube, Lanzarote, with description of Meganerilla cesari, n. sp

    DEFF Research Database (Denmark)

    Worsaae, Katrine; Martínez, A; Núñez, J

    2009-01-01

    Five species of Nerillidae are previously known from Atlantic cave systems. Another four species of Nerillidae are reported here from the Corona lava tube (Lanzarote, Canary Islands) presenting the first records of Mesonerilla and Meganerilla from anchialine environments. We here describe...

  13. Anchialine fauna of the Corona lava tunnel (Lanzarote,Canary Islands): diversity, endemism and distribution

    DEFF Research Database (Denmark)

    Martínez, Alexandro; Palmero, A M; Brito, M C

    2009-01-01

    A checklist of 77 taxa recorded from the anchialine sections of the Corona lava tube is provided, including information on habitats, faunal distribution within the cave, and main references. Of the nine major groups recorded, Crustacea shows the highest diversity with 31 species and the highest d...

  14. Ecological anatomy of ferns fronds

    Directory of Open Access Journals (Sweden)

    Nina M. Derzhavina

    2014-04-01

    Full Text Available Structural types of frond anatomy are distinguished on the basis of investigation of 30 species of homosporous ferns and with regard for literature: hydromorphic, hygromorphic, mesomorphic, subxeromorphic, and subsucculent (cryptic succulent. Following frond traits are of highest adaptive value: their area and thickness, type of mesophyll, dry weight of an area unit – specific superficial density, cellular volume, and number of cells per unit of frond area.

  15. In situ NIR reflectance and LIBS measurements in lava tubes in preparation for future Mars missions

    Science.gov (United States)

    Leveille, R.; Sobron, P.

    2017-12-01

    The ATiLT (Astrobiology Training in Lava Tubes) program addresses Mars astrobiology exploration objectives by performing field work and instrumental analyses in lava tubes as high fidelity analog environments to putative lava tubes on Mars. The main field location for ATiLT is the Lava Beds National Monument (LABE) in Northern California. LABE is situated on the lower north flank of the Medicine Lake Volcano of the Cascade arc. This location features hundreds of caves, most of which are relatively shallow, typically well above the water table, reaching 20-45m below land surface at their maximum depth. Some LABE caves feature `cold sinks' where cold air sinks and becomes trapped in deeper cave passages, thus allowing perennial ice to accumulate despite above freezing surface temperatures. Several lava tube caves in LABE also contain seasonal or perennial ice accumulations, which makes them excellent analogs to Mars lava tubes where the presence of ice has been predicted. While lava tubes are very attractive systems to test hypotheses related to habitability and the possibility for life on Mars, at present there are no comprehensive in-situ instrument-driven characterizations of the mineralogy and geochemistry of lava tubes. ATiLT fills this gap by providing detailed, in-situ investigations with scientific instruments relevant to Mars exploration. Our aim is to help constrain future exploration targets on Mars and define future mission operations and requirements. For this purpose, in May 2017 we carried out a field campaign in several lava tubes at LABE. We deployed two miniature spectroscopic sensors suitable for dark, humid, cave conditions: NIR reflectance (1-5 μm) and LIBS (300-900 nm). The advantages of combining NIR reflectance and LIBS are evident: LIBS can reveal the relative concentration of major (and often trace) elements present in a bulk sample, whereas NIR reflectance yields information on the individual mineral species and their chemical and

  16. Ferns diversified in the shadow of angiosperms.

    Science.gov (United States)

    Schneider, Harald; Schuettpelz, Eric; Pryer, Kathleen M; Cranfill, Raymond; Magallón, Susana; Lupia, Richard

    2004-04-01

    The rise of angiosperms during the Cretaceous period is often portrayed as coincident with a dramatic drop in the diversity and abundance of many seed-free vascular plant lineages, including ferns. This has led to the widespread belief that ferns, once a principal component of terrestrial ecosystems, succumbed to the ecological predominance of angiosperms and are mostly evolutionary holdovers from the late Palaeozoic/early Mesozoic era. The first appearance of many modern fern genera in the early Tertiary fossil record implies another evolutionary scenario; that is, that the majority of living ferns resulted from a more recent diversification. But a full understanding of trends in fern diversification and evolution using only palaeobotanical evidence is hindered by the poor taxonomic resolution of the fern fossil record in the Cretaceous. Here we report divergence time estimates for ferns and angiosperms based on molecular data, with constraints from a reassessment of the fossil record. We show that polypod ferns (> 80% of living fern species) diversified in the Cretaceous, after angiosperms, suggesting perhaps an ecological opportunistic response to the diversification of angiosperms, as angiosperms came to dominate terrestrial ecosystems.

  17. Speleothem (Cave Deposit) Data

    Data.gov (United States)

    National Oceanic and Atmospheric Administration, Department of Commerce — Records of past temperature, precipitation, and other aspects of climate derived from mineral deposits found in caves. Parameter keywords describe what was measured...

  18. Energy expenditure in caving.

    Directory of Open Access Journals (Sweden)

    Giorgia Antoni

    Full Text Available The aim of this study was to determine the energy expenditure of a group of cavers of both genders and different ages and experience during a 10 hour subterranean exploration, using portable metabolimeters. The impact of caving activity on body composition and hydration were also assessed through bioelectrical impedance, and nutritional habits of cavers surveyed. During cave activity, measured total energy expenditure (TEE was in the range 225-287 kcal/h for women-men (MET = 4.1, respectively; subjects had an energy intake from food in the range 1000-1200 kcal, thus inadequate to restore lost calories. Bayesian statistical analysis estimated the effect of predictive variables on TEE, revealing that experienced subjects had a 5% lower TEE than the less skilled ones and that women required a comparatively larger energy expenditure than men to perform the same task. BIVA (bioelectrical impedance vector analysis showed that subjects were within the range of normal hydration before and after cave activity, but bioelectrical changes indicated a reduction of extracellular water in men, which might result in hypo-osmolal dehydration in the case of prolonged underground exercise. All these facts should be considered when planning cave explorations, preparing training programs for subjects practising caving, and optimizing a diet for cavers. Further, information gathered through this study could be of value to reduce accidents in caves related to increase in fatigue.

  19. Cryogenic Cave Pearls In the Periglacial Zones of Ice Caves

    Czech Academy of Sciences Publication Activity Database

    Žák, Karel; Orvošová, M.; Filippi, Michal; Vlček, M.; Onac, B. P.; Persoiu, A.; Rohovec, Jan; Světlík, Ivo

    2013-01-01

    Roč. 83, č. 2 (2013), s. 207-220 ISSN 1527-1404 R&D Projects: GA ČR GAP210/10/1760 Institutional research plan: CEZ:AV0Z30130516 Institutional support: RVO:67985831 ; RVO:61389005 Keywords : caves * cryogenic caves * ice caves * periglacial zones Subject RIV: DB - Geology ; Mineralogy Impact factor: 1.943, year: 2013

  20. Winter distribution and use of high elevation caves as foraging sites by the endangered Hawaiian hoary bat, Lasiurus cinereus semotus

    Science.gov (United States)

    Bonaccorso, Frank; Montoya-Aiona, Kristina; Pinzari, Corinna A.; Todd, Christopher M.

    2016-01-01

    We examine altitudinal movements involving unusual use of caves by Hawaiian hoary bats, Lasiurus cinereus semotus, during winter and spring in the Mauna Loa Forest Reserve (MLFR), Hawai‘i Island. Acoustic detection of hoary bat vocalizations, were recorded with regularity outside 13 lava tube cave entrances situated between 2,200 to 3,600 m asl from November 2012 to April 2013. Vocalizations were most numerous in November and December with the number of call events and echolocation pulses decreasing through the following months. Bat activity was positively correlated with air temperature and negatively correlated with wind speed. Visual searches found no evidence of hibernacula nor do Hawaiian hoary bats appear to shelter by day in these caves. Nevertheless, bats fly deep into caves as evidenced by numerous carcasses found in cave interiors. The occurrence of feeding buzzes around cave entrances and visual observations of bats flying in acrobatic fashion in cave interiors point to the use of these spaces as foraging sites. Peridroma moth species (Noctuidae), the only abundant nocturnal, flying insect sheltering in large numbers in rock rubble and on cave walls in the MLFR, apparently serve as the principal prey attracting hoary bats during winter to lava tube caves in the upper MLFR. Caves above 3,000 m on Mauna Loa harbor temperatures suitable for Pseudogymnoascus destructansfungi, the causative agent of White-nose Syndrome that is highly lethal to some species of North American cave-dwelling bats. We discuss the potential for White-nose Syndrome to establish and affect Hawaiian hoary bats.

  1. Lava tubes and aquifer vulnerability in the upper Actopan River basin, Veracruz, México

    Science.gov (United States)

    Espinasa-Pereña, R.; Delgado Granados, H.

    2011-12-01

    Rapid infiltration leads to very dry conditions on the surface of some volcanic terrains, with large allogenic streams sometimes sinking underground upon reaching a lava flow. Aquifers in lava flows tend to be heterogeneous and discontinuous, generally unconfined and fissured, and have high transmissivity. Springs associated with basalts may be very large but are typically restricted to lava-flow margins. Concern has been expressed regarding the potential for lava-tube caves to facilitate groundwater contamination similar to that afflicting some karst aquifers (Kempe et al., 2003; Kiernan et al., 2002; Halliday 2003). The upper Actopan River basin is a series of narrow valleys excavated in Tertiary volcanic brechias. Several extensive Holocene basaltic tube-fed lava flows have partially filled these valleys. The youngest and longest flow originates at El Volcancillo, a 780 ybP monogenetic volcano. It is over 50 km long, and was fed through a major master tube, the remains of which form several lava-tube caves (Gassos and Espinasa-Pereña, 2008). Another tube-fed flow initiates at a vent at the bottom of Barranca Huichila and can be followed for 7 km to where it is covered by the Volcancillo flow. The Huichila River is captured by this system of lava tubes and can be followed through several underground sections. In dry weather the stream disappears at a sump in one of these caves, although during hurricanes it overflows the tube, floods the Tengonapa plain, and finally sinks through a series of skylights into the master tube of the Volcancillo flow. Near villages, the cave entrances are used as trash dumps, which are mobilized during floods. These include household garbage, organic materials associated with agriculture and even medical supplies. This is a relatively recent phenomenon, caused by population growth and the building of houses above the lava flows. The water resurges at El Descabezadero, gushing from fractures in the lava above the underlying brechias

  2. rFerns: An Implementation of the Random Ferns Method for General-Purpose Machine Learning

    Directory of Open Access Journals (Sweden)

    Miron B. Kursa

    2014-11-01

    Full Text Available Random ferns is a very simple yet powerful classification method originally introduced for specific computer vision tasks. In this paper, I show that this algorithm may be considered as a constrained decision tree ensemble and use this interpretation to introduce a series of modifications which enable the use of random ferns in general machine learning problems. Moreover, I extend the method with an internal error approximation and an attribute importance measure based on corresponding features of the random forest algorithm. I also present the R package rFerns containing an efficient implementation of this modified version of random ferns.

  3. Studies of Young Hawai'ian Lava Tubes: Implications for Planetary Habitability and Human Exploration

    Science.gov (United States)

    McAdam, Amy; Bleacher, Jacob; Young, Kelsey; Johnson, Sarah Stewart; Needham, Debra; Schmerr, Nicholas; Shiro, Brian; Garry, Brent; Whelley, Patrick; Knudson, Christine; hide

    2017-01-01

    Habitability: Subsurface environments may preserve records of habitability or biosignatures, with more stable environmental conditions compared to surface (e.g., smaller variations in temperature and humidity) and reduced exposure to radiation; Lava tubes are expected on Mars, and candidates are observed from orbit; Few detailed studies of microbial populations in terrestrial lava caves; Also contain a variety of secondary minerals; Microbial activity may play a role in mineral formation or be preserved in these minerals; Minerals can provide insight into fluids (e.g., pH, temperature).

  4. SURVEY OF ORNAMENTAL FERNS, THEIR MORPHOLOGY AND ...

    African Journals Online (AJOL)

    ornamental ferns for environmental management and economic values. ... little variations in the structures of rhizome except in Phymatodes .... forms such as erect, tall, open, bushy and .... linear leaflet with acute apex and flat, glabrous lamina.

  5. Random clustering ferns for multimodal object recognition

    OpenAIRE

    Villamizar Vergel, Michael Alejandro; Garrell Zulueta, Anais; Sanfeliu Cortés, Alberto; Moreno-Noguer, Francesc

    2017-01-01

    The final publication is available at link.springer.com We propose an efficient and robust method for the recognition of objects exhibiting multiple intra-class modes, where each one is associated with a particular object appearance. The proposed method, called random clustering ferns, combines synergically a single and real-time classifier, based on the boosted assembling of extremely randomized trees (ferns), with an unsupervised and probabilistic approach in order to recognize efficient...

  6. Trends and concepts in fern classification

    Science.gov (United States)

    Christenhusz, Maarten J. M.; Chase, Mark W.

    2014-01-01

    Background and Aims Throughout the history of fern classification, familial and generic concepts have been highly labile. Many classifications and evolutionary schemes have been proposed during the last two centuries, reflecting different interpretations of the available evidence. Knowledge of fern structure and life histories has increased through time, providing more evidence on which to base ideas of possible relationships, and classification has changed accordingly. This paper reviews previous classifications of ferns and presents ideas on how to achieve a more stable consensus. Scope An historical overview is provided from the first to the most recent fern classifications, from which conclusions are drawn on past changes and future trends. The problematic concept of family in ferns is discussed, with a particular focus on how this has changed over time. The history of molecular studies and the most recent findings are also presented. Key Results Fern classification generally shows a trend from highly artificial, based on an interpretation of a few extrinsic characters, via natural classifications derived from a multitude of intrinsic characters, towards more evolutionary circumscriptions of groups that do not in general align well with the distribution of these previously used characters. It also shows a progression from a few broad family concepts to systems that recognized many more narrowly and highly controversially circumscribed families; currently, the number of families recognized is stabilizing somewhere between these extremes. Placement of many genera was uncertain until the arrival of molecular phylogenetics, which has rapidly been improving our understanding of fern relationships. As a collective category, the so-called ‘fern allies’ (e.g. Lycopodiales, Psilotaceae, Equisetaceae) were unsurprisingly found to be polyphyletic, and the term should be abandoned. Lycopodiaceae, Selaginellaceae and Isoëtaceae form a clade (the lycopods) that is

  7. Trends and concepts in fern classification.

    Science.gov (United States)

    Christenhusz, Maarten J M; Chase, Mark W

    2014-03-01

    Throughout the history of fern classification, familial and generic concepts have been highly labile. Many classifications and evolutionary schemes have been proposed during the last two centuries, reflecting different interpretations of the available evidence. Knowledge of fern structure and life histories has increased through time, providing more evidence on which to base ideas of possible relationships, and classification has changed accordingly. This paper reviews previous classifications of ferns and presents ideas on how to achieve a more stable consensus. An historical overview is provided from the first to the most recent fern classifications, from which conclusions are drawn on past changes and future trends. The problematic concept of family in ferns is discussed, with a particular focus on how this has changed over time. The history of molecular studies and the most recent findings are also presented. Fern classification generally shows a trend from highly artificial, based on an interpretation of a few extrinsic characters, via natural classifications derived from a multitude of intrinsic characters, towards more evolutionary circumscriptions of groups that do not in general align well with the distribution of these previously used characters. It also shows a progression from a few broad family concepts to systems that recognized many more narrowly and highly controversially circumscribed families; currently, the number of families recognized is stabilizing somewhere between these extremes. Placement of many genera was uncertain until the arrival of molecular phylogenetics, which has rapidly been improving our understanding of fern relationships. As a collective category, the so-called 'fern allies' (e.g. Lycopodiales, Psilotaceae, Equisetaceae) were unsurprisingly found to be polyphyletic, and the term should be abandoned. Lycopodiaceae, Selaginellaceae and Isoëtaceae form a clade (the lycopods) that is sister to all other vascular plants, whereas

  8. Hawaii Lava Flows

    Science.gov (United States)

    2001-01-01

    This sequence of ASTER nighttime thermal images shows the Pu'u O'o lava flows entering the sea at Kamokuna on the southeast side of the Island of Hawaii. Each image covers an area of 9 x 12 km. The acquisition dates are April 4 2000, May 13 2000, May 22 2000 (upper row) and June 30 2000, August 1 2000 and January 1 2001 (lower row). Thermal band 14 has been color coded from black (coldest) through blue, red, yellow and white (hottest). The first 5 images show a time sequence of a single eruptive phase; the last image shows flows from a later eruptive phase. The images are located at 19.3 degrees north latitude, 155 degrees west longitude. The U.S. science team is located at NASA's Jet Propulsion Laboratory, Pasadena, Calif. The Terra mission is part of NASA's Science Mission Directorate.

  9. Degassing driving crystallization of plagioclase phenocrysts in lava tube stalactites on Mount Etna (Sicily, Italy)

    Science.gov (United States)

    Lanzafame, Gabriele; Ferlito, Carmelo

    2014-10-01

    Basaltic lava flows can form tubes in response to the cooling of the outer surface. We collected lava stalactites (frozen lava tears) and sampled lava from the ceilings of three lava tubes on Mount Etna. Comparison of the petrographic characters between ceiling lavas and relative stalactites reveals surprising differences in the groundmass textures and crystal compositions. Major and trace element contents in stalactites show only a slight increase in alkali and SiO2 compared to ceiling lava, whereas significant differences exist in composition and textures between plagioclases within the ceiling lava and those within the stalactites, being in the last case definitively more An-rich. We advance the hypothesis that the high temperature reached in the cave caused the exsolution of the volatiles still trapped in the dripping melt. The volatiles, mainly H2O, formed bubbles and escaped from the melt; such a water-loss might have promoted the silicate polymerization in the stalactites resulting in the growth of An-rich plagioclase phenocrysts. Our results have important implications: in fact plagioclase phenocrysts are usually associated with intratelluric growth and are often considered as the main petrologic evidence for the existence of a magma chamber. The textural and chemical features of plagioclases in stalactites prove that phenocryst growth in syn to post-eruptive conditions is plausible and clearly explains the relatively low viscosity of many phenocryst-rich lava flows on Mount Etna, as well as on many other volcanoes around the world. Therefore, we can conclude that plagioclase phenocrysts cannot exclusively be considered as having originated within a magma chamber.

  10. Chloroplast Genome Evolution in Early Diverged Leptosporangiate Ferns

    OpenAIRE

    Kim, Hyoung Tae; Chung, Myong Gi; Kim, Ki-Joong

    2014-01-01

    In this study, the chloroplast (cp) genome sequences from three early diverged leptosporangiate ferns were completed and analyzed in order to understand the evolution of the genome of the fern lineages. The complete cp genome sequence of Osmunda cinnamomea (Osmundales) was 142,812 base pairs (bp). The cp genome structure was similar to that of eusporangiate ferns. The gene/intron losses that frequently occurred in the cp genome of leptosporangiate ferns were not found in the cp genome of O. c...

  11. Stoichiometry of ferns in Hawaii: implications for nutrient cycling.

    Science.gov (United States)

    Amatangelo, Kathryn L; Vitousek, Peter M

    2008-10-01

    We asked if element concentrations in ferns differ systematically from those in woody dicots in ways that could influence ecosystem properties and processes. Phylogenetically, ferns are deeply separated from angiosperms; for our analyses we additionally separated leptosporangiate ferns into polypod ferns, a monophyletic clade of ferns which radiated after the rise of angiosperms, and all other leptosporangiate (non-polypod) ferns. We sampled both non-polypod and polypod ferns on a natural fertility gradient and within fertilized and unfertilized plots in Hawaii, and compared our data with shrub and tree samples collected previously in the same plots. Non-polypod ferns in particular had low Ca concentrations under all conditions and less plasticity in their N and P stoichiometry than did polypod ferns or dicots. Polypod ferns were particularly rich in N and P, with low N:P ratios, and their stoichiometry varied substantially in response to differences in nutrient availability. Distinguishing between these two groups has the potential to be useful both in and out of Hawaii, as they have distinct properties which can affect ecosystem function. These differences could contribute to the widespread abundance of polypod ferns in an angiosperm-dominated world, and to patterns of nutrient cycling and limitation in sites where ferns are abundant.

  12. Survey and hydrogeology of Carroll Cave

    Science.gov (United States)

    Carroll Cave, located in Camden County, Missouri, is the largest known cave formed in the Gasconade Dolomite of the Salem Plateau. Despite extensive visitation over the last 50 years and multiple survey efforts, a comprehensive map of the cave has never been produced. In 2002, the Carroll Cave Conse...

  13. Spiders in caves.

    Science.gov (United States)

    Mammola, Stefano; Isaia, Marco

    2017-04-26

    World experts of different disciplines, from molecular biology to macro-ecology, recognize the value of cave ecosystems as ideal ecological and evolutionary laboratories. Among other subterranean taxa, spiders stand out as intriguing model organisms for their ecological role of top predators, their unique adaptations to the hypogean medium and their sensitivity to anthropogenic disturbance. As the description of the first eyeless spider ( Stalita taenaria ), an array of papers on subterranean spider biology, ecology and evolution has been published, but a comprehensive review on these topics is still lacking. We provide a general overview of the spider families recorded in hypogean habitats worldwide, we review the different adaptations of hypogean spiders to subterranean life, and we summarize the information gathered so far about their origin, population structure, ecology and conservation status. Finally, we point out the limits of the knowledge we currently have regarding hypogean spiders, aiming to stimulate future research. © 2017 The Author(s).

  14. Cave breakdown by vadose weathering.

    Directory of Open Access Journals (Sweden)

    Osborne R. Armstrong L.

    2002-01-01

    Full Text Available Vadose weathering is a significant mechanism for initiating breakdown in caves. Vadose weathering of ore bodies, mineral veins, palaeokarst deposits, non-carbonate keystones and impure, altered or fractured bedrock, which is intersected by caves, will frequently result in breakdown. Breakdown is an active, ongoing process. Breakdown occurs throughout the vadose zone, and is not restricted to large diameter passages, or to cave ceilings. The surfaces of disarticulated blocks are commonly coated, rather than having fresh broken faces, and blocks continue to disintegrate after separating from the bedrock. Not only gypsum, but also hydromagnesite and aragonite are responsible for crystal wedging. It is impossible to study or identify potential breakdown foci by surface surveys alone, in-cave observation and mapping are essential.

  15. Complete chloroplast genome sequence of a tree fern Alsophila spinulosa: insights into evolutionary changes in fern chloroplast genomes.

    Science.gov (United States)

    Gao, Lei; Yi, Xuan; Yang, Yong-Xia; Su, Ying-Juan; Wang, Ting

    2009-06-11

    Ferns have generally been neglected in studies of chloroplast genomics. Before this study, only one polypod and two basal ferns had their complete chloroplast (cp) genome reported. Tree ferns represent an ancient fern lineage that first occurred in the Late Triassic. In recent phylogenetic analyses, tree ferns were shown to be the sister group of polypods, the most diverse group of living ferns. Availability of cp genome sequence from a tree fern will facilitate interpretation of the evolutionary changes of fern cp genomes. Here we have sequenced the complete cp genome of a scaly tree fern Alsophila spinulosa (Cyatheaceae). The Alsophila cp genome is 156,661 base pairs (bp) in size, and has a typical quadripartite structure with the large (LSC, 86,308 bp) and small single copy (SSC, 21,623 bp) regions separated by two copies of an inverted repeat (IRs, 24,365 bp each). This genome contains 117 different genes encoding 85 proteins, 4 rRNAs and 28 tRNAs. Pseudogenes of ycf66 and trnT-UGU are also detected in this genome. A unique trnR-UCG gene (derived from trnR-CCG) is found between rbcL and accD. The Alsophila cp genome shares some unusual characteristics with the previously sequenced cp genome of the polypod fern Adiantum capillus-veneris, including the absence of 5 tRNA genes that exist in most other cp genomes. The genome shows a high degree of synteny with that of Adiantum, but differs considerably from two basal ferns (Angiopteris evecta and Psilotum nudum). At one endpoint of an ancient inversion we detected a highly repeated 565-bp-region that is absent from the Adiantum cp genome. An additional minor inversion of the trnD-GUC, which is possibly shared by all ferns, was identified by comparison between the fern and other land plant cp genomes. By comparing four fern cp genome sequences it was confirmed that two major rearrangements distinguish higher leptosporangiate ferns from basal fern lineages. The Alsophila cp genome is very similar to that of the

  16. Complete chloroplast genome sequence of a tree fern Alsophila spinulosa: insights into evolutionary changes in fern chloroplast genomes

    Directory of Open Access Journals (Sweden)

    Yang Yong-Xia

    2009-06-01

    Full Text Available Abstract Background Ferns have generally been neglected in studies of chloroplast genomics. Before this study, only one polypod and two basal ferns had their complete chloroplast (cp genome reported. Tree ferns represent an ancient fern lineage that first occurred in the Late Triassic. In recent phylogenetic analyses, tree ferns were shown to be the sister group of polypods, the most diverse group of living ferns. Availability of cp genome sequence from a tree fern will facilitate interpretation of the evolutionary changes of fern cp genomes. Here we have sequenced the complete cp genome of a scaly tree fern Alsophila spinulosa (Cyatheaceae. Results The Alsophila cp genome is 156,661 base pairs (bp in size, and has a typical quadripartite structure with the large (LSC, 86,308 bp and small single copy (SSC, 21,623 bp regions separated by two copies of an inverted repeat (IRs, 24,365 bp each. This genome contains 117 different genes encoding 85 proteins, 4 rRNAs and 28 tRNAs. Pseudogenes of ycf66 and trnT-UGU are also detected in this genome. A unique trnR-UCG gene (derived from trnR-CCG is found between rbcL and accD. The Alsophila cp genome shares some unusual characteristics with the previously sequenced cp genome of the polypod fern Adiantum capillus-veneris, including the absence of 5 tRNA genes that exist in most other cp genomes. The genome shows a high degree of synteny with that of Adiantum, but differs considerably from two basal ferns (Angiopteris evecta and Psilotum nudum. At one endpoint of an ancient inversion we detected a highly repeated 565-bp-region that is absent from the Adiantum cp genome. An additional minor inversion of the trnD-GUC, which is possibly shared by all ferns, was identified by comparison between the fern and other land plant cp genomes. Conclusion By comparing four fern cp genome sequences it was confirmed that two major rearrangements distinguish higher leptosporangiate ferns from basal fern lineages. The

  17. LAVA Pressure Transducer Trade Study

    Science.gov (United States)

    Oltman, Samuel B.

    2016-01-01

    The Regolith and Environment Science and Oxygen and Lunar Volatile Extraction (RESOLVE) payload will transport the (LAVA) subsystem to hydrogen-rich locations on the moon supporting NASA's in-situ resource utilization (ISRU) programs. There, the LAVA subsystem will analyze volatiles that evolve from heated regolith samples in order to quantify how much water is present. To do this, the system needs resilient pressure transducers (PTs) to calculate the moles in the gas samples. The PT trade study includes a comparison of newly-procured models to a baseline unit with prior flight history in order to determine the PT model with the best survivability in flight-forward conditions.

  18. The fern sporangium: an ultrafast natural catapult

    Science.gov (United States)

    Noblin, Xavier; Argentina, Mederic; Westbrook, Jared; Llorens, Coraline; Rojas, Nicolas; Dumais, Jacques

    2012-02-01

    Plants have developed fascinating mechanisms to create ultra fast movements that often reach the upper limit allowed by physical laws. Inspiration for new technologies is one of the reasons for the strong interest for these mechanisms, along with the deep interest of understanding complex, natural systems. The fern sporangium is a capsule that contains the spores, it is surrounded by a row of cells called the annulus which acts as a beam. Due to the water evaporation from its cells, the annulus bends strongly and induces elastic energy storage during an opening phase. The tension in the cells breaks when cavitation bubbles appear in the cells, leading to a fast release of the elastic energy. The fern sporangium then acts as a catapult which ejects rapidly its spores by closing back to the initial closed shape. We have analyzed the slow opening motion and the fast catapulting mechanism. We found that the catapult motion involves two time scales, showing a very original behavior. In man-made catapults, the recoil motion needs to be arrested by a cross bar so that the projectile is released from the arm. We show here that the fern sporangium replaces the essential cross bar by an elegant poroelastic damping, leading to a completely autonomous, efficient device.

  19. Tear ferning test in healthy dogs.

    Science.gov (United States)

    Oriá, Arianne P; Raposo, Ana Claudia S; Araújo, Nayone L L C; Lima, Felipe B; Masmali, Ali M

    2017-11-07

    To evaluate and compare three tear sampling methods using two grading scales for administering the tear ferning test (TFT) to healthy dogs. In total, 90 dogs (180 eyes) were subjected to tear sampling using millimetered strips, reused after the Schirmer tear test (STT) (Schirmer group, SG). Then, the dogs were subdivided into three groups according to sampling approach: micropipette (MPG), microcapillary (MCG), and Schirmer sample 2 (S2G). The collected tears were dried on a clean microscope glass slide at room temperature and humidity. The ferning patterns were observed under a polarized light microscope and classified according to the Rolando and Masmali grading scales. Although all three methods were feasible, the STT was easier to perform in clinical settings. Type I and Grade 1 were the most commonly observed (64.17% and 61.7%, respectively) regardless of collection method. There was no significant difference between the STT median values and the TFT classifications. The TFT is appropriate for dogs and can be performed using the three suggested sampling methods, with a higher frequency of Type I and Grade 1. Thus, it is possible to use both grading scales in the classification of tear ferning in dogs. © 2017 American College of Veterinary Ophthalmologists.

  20. Exposure to radon in Cuban tourist caves

    International Nuclear Information System (INIS)

    Carrazana Gonzalez, Jorge; Dominguez Ley, Orlando; Gil Castillo, Reinaldo; Molerio Leon, Leslie

    2008-01-01

    With the objective of estimating the dose, due to Radon-222, received by tour guides and other people who work inside some of the most important tourist caves in the Republic of Cuba, measurements of radon concentrations were carried out in four of these caves: Santo Tomas Cave, Tapiada Cave, Del Indio Cave and Jose Miguel Cave. All these caves are located in the Vinales Valley (Pinar del Rio City), a very important tourist area in the country. The relationship among radon concentration, the concentration of natural radionuclides inside the caves and the geologic characteristics of the specific locations was analyzed. In order to select the most appropriates measurement places, different criteria were taken into account: representatives points of the geology of the caves, level of gamma radiation, exchange of air, places of work inside the caves (cafeterias, restaurants and shops) and places more frequently visited by tour guides. The maximum radon concentration was found in Jose Miguel Cave with a value of 220 Bq/m 3 . The measurement of gamma dose rate inside the caves was carried out with a scintillation detector type Scintrex BGS-3 previously calibrated at SSI, Sweden. The radon concentrations were measured with the equipment Alpha Guard PQ2000/MC50 and SARAD RM2000, both also calibrated at SSI, Sweden. The obtained results indicated that, in the studied caves, the annual dose is not greater than 1 mSv considering a work year of 2000 hours. (author)

  1. Lava Flow at Kilauea, Hawaii

    Science.gov (United States)

    2007-01-01

    On July 21, 2007, the world's most active volcano, Kilauea on Hawaii's Big Island, produced a new fissure eruption from the Pu'u O'o vent, which fed an open lava channel and lava flows toward the east. Access to the Kahauale'a Natural Area Reserve was closed due to fire and gas hazards. The two Advanced Spaceborne Thermal Emission and Reflection Radiometer (ASTER) nighttime thermal infrared images were acquired on August 21 and August 30, 2007. The brightest areas are the hottest lava flows from the recent fissure eruption. The large lava field extending down to the ocean is part of the Kupaianaha field. The most recent activity there ceased on June 20, but the lava is still hot and appears bright on the images. Magenta areas are cold lava flows from eruptions that occurred between 1969 and 2006. Clouds are cold (black) and the ocean is a uniform warm temperature, and light gray in color. These images are being used by volcanologists at the U.S. Geological Survey Hawaii Volcano Observatory to help monitor the progress of the lava flows. With its 14 spectral bands from the visible to the thermal infrared wavelength region, and its high spatial resolution of 15 to 90 meters (about 50 to 300 feet), ASTER images Earth to map and monitor the changing surface of our planet. ASTER is one of five Earth-observing instruments launched December 18, 1999, on NASA's Terra spacecraft. The instrument was built by Japan's Ministry of Economy, Trade and Industry. A joint U.S./Japan science team is responsible for validation and calibration of the instrument and the data products. The broad spectral coverage and high spectral resolution of ASTER provides scientists in numerous disciplines with critical information for surface mapping, and monitoring of dynamic conditions and temporal change. Example applications are: monitoring glacial advances and retreats; monitoring potentially active volcanoes; identifying crop stress; determining cloud morphology and physical properties

  2. GEOMORPHIC ANALYSIS OF MAWSMAI CAVE, MEGHALAYA, INDIA

    Directory of Open Access Journals (Sweden)

    Vibhash Chandra Jha

    2011-12-01

    Full Text Available ABSTRACT Caving is an art which can be best experienced in the mystic Mawsmai Cave of Meghalaya. This Cave is fast becoming a hot tourist’s destination offering great adventurous opportunity to the tourists. The cave is located in the East Khasi Hill region near Cherrapunjee and is made of limestone. The emotion of thrill heightens after entering the cave. The entrance has a narrow vertical opening and is well lit. Due to its location in the world’s largest rainfall region, dripping of water from the cave roofs occurs almost throughout the year. The formation of stalactites and stalagmites create wonderful phenomena specially found in this caves. The conspicuous pillars formed due to the joining of the roof and the floors are an awe-inspiring creativity of the creator of this world. Keywords: Cave, Limestone, Stalactite, Stalagmite, Pillar, Solution.

  3. The future of the CAVE

    KAUST Repository

    DeFanti, Thomas; Acevedo-Feliz, Daniel; Ainsworth, Richard; Brown, Maxine; Cutchin, Steven; Dawe, Gregory; Doerr, Kai-Uwe; Johnson, Andrew; Knox, Christopher; Kooima, Robert; Kuester, Falko; Leigh, Jason; Long, Lance; Otto, Peter; Petrovic, Vid; Ponto, Kevin; Prudhomme, Andrew; Rao, Ramesh; Renambot, Luc; Sandin, Daniel; Schulze, Jurgen; Smarr, Larry; Srinivasan, Madhusudhanan; Weber, Philip; Wickham, Gregory

    2011-01-01

    The CAVE, a walk-in virtual reality environment typically consisting of 4–6 3 m-by-3 m sides of a room made of rear-projected screens, was first conceived and built in 1991. In the nearly two decades since its conception, the supporting technology has improved so that current CAVEs are much brighter, at much higher resolution, and have dramatically improved graphics performance. However, rear-projection-based CAVEs typically must be housed in a 10 m-by-10 m-by-10 m room (allowing space behind the screen walls for the projectors), which limits their deployment to large spaces. The CAVE of the future will be made of tessellated panel displays, eliminating the projection distance, but the implementation of such displays is challenging. Early multi-tile, panel-based, virtual-reality displays have been designed, prototyped, and built for the King Abdullah University of Science and Technology (KAUST) in Saudi Arabia by researchers at the University of California, San Diego, and the University of Illinois at Chicago. New means of image generation and control are considered key contributions to the future viability of the CAVE as a virtual-reality device.

  4. Basaltic caves at Craters of the Moon National Monument and Preserve as analogs for Mars

    Science.gov (United States)

    Hinman, N. W.; Richardson, C. D.; McHenry, L.; Scott, J. R.

    2010-12-01

    Basaltic caves and lava tubes offer stable physicochemical conditions for formation of secondary minerals. Such features, putatively observed on Mars, intercept groundwater to weather country rock, leading to formation of secondary minerals. Further, caves are stable environments to search for evidence of past life, as they could offer protection from the oxidizing martian atmosphere. Searching for signs of life in a cave that could protect bio/organic compounds would preclude the need for risky drilling on Mars. Craters of the Moon National Monument (COM) offers an opportunity to study caves in Holocene iron-rich basalt flows to characterize secondary mineral deposits and search for organic compounds associated with secondary minerals; COM basalts are a good analog for martian basalts because of their high iron but other elements are higher at COM than on Mars. The Blue Dragon flow (~2.1 ka) contains the majority of the accessible caves and lava tubes. Two types of secondary mineral deposits were observed in these caves: ceiling coatings and crack or floor precipitates. Hematite, silica, and calcite comprise ceiling coatings. The crack and floor precipitates are white, efflorescent deposits in cavities along cave walls and ceilings or in localized mounds on cave floors. The secondary minerals in crack and floor precipitates are mainly thenardite and mirabilite with some minor concentrations of trona and/or burkeite. Organic compounds were found associated with the efflorescent deposits. Formation of the deposits is likely due to chemical leaching of basalt by meteoritic water. To test this, fluids collected from the ceiling and walls of the caves were analyzed. Solutions were modeled with the geochemical code, PHREEQC. The model tracked composition as water evaporated. Selected minerals were allowed to precipitate as they became oversaturated. Among the first minerals to become oversaturated were quartz and calcite, which are observed in ceiling deposits. Iron

  5. Chronology of guitarrero cave, peru.

    Science.gov (United States)

    Lynch, T F; Gillespie, R; Gowlett, J A; Hedges, R E

    1985-08-30

    Dating by accelerator mass spectrometry of wooden artifacts, cord, and charcoal samples from Guitarrero Cave, Peru, supports the antiquity of South America's earliest textiles and other perishable remains. The new dates are consistent with those obtained from disintegration counters and leave little doubt about the integrity of the lower Preceramic layers and their early cultivars. Re-evaluation of the mode of deposition suggests that most of the remains resulted from short-term use of the cave in the eighth millennium B.C., with a possible brief human visit as early as 12,560 years ago.

  6. Two new fern chloroplasts and decelerated evolution linked to the long generation time in tree ferns.

    Science.gov (United States)

    Zhong, Bojian; Fong, Richard; Collins, Lesley J; McLenachan, Patricia A; Penny, David

    2014-04-30

    We report the chloroplast genomes of a tree fern (Dicksonia squarrosa) and a "fern ally" (Tmesipteris elongata), and show that the phylogeny of early land plants is basically as expected, and the estimates of divergence time are largely unaffected after removing the fastest evolving sites. The tree fern shows the major reduction in the rate of evolution, and there has been a major slowdown in the rate of mutation in both families of tree ferns. We suggest that this is related to a generation time effect; if there is a long time period between generations, then this is probably incompatible with a high mutation rate because otherwise nearly every propagule would probably have several lethal mutations. This effect will be especially strong in organisms that have large numbers of cell divisions between generations. This shows the necessity of going beyond phylogeny and integrating its study with other properties of organisms. © The Author(s) 2014. Published by Oxford University Press on behalf of the Society for Molecular Biology and Evolution.

  7. LavaSIM: the effect of heat transfer in 3D on lava flow characteristics (Invited)

    Science.gov (United States)

    Fujita, E.

    2013-12-01

    Characteristics of lava flow are governed by many parameters like lava viscosity, effusion rate, ground topography, etc. The accuracy and applicability of lava flow simulation code is evaluated whether the numerical simulation can reproduce these features quantitatively, which is important from both strategic and scientific points of views. Many lava flow simulation codes are so far proposed, and they are classified into two categories, i.e., the deterministic and the probabilistic models. LavaSIM is one of the former category models, and has a disadvantage of time consuming. But LavaSIM can solves the equations of continuity, motion, energy by step and has an advantage in the calculation of three-dimensional analysis with solid-liquid two phase flow, including the heat transfer between lava, solidified crust, air, water and ground, and three-dimensional convection in liquid lava. In other word, we can check the detailed structure of lava flow by LavaSIM. Therefore, this code can produce both channeled and fan-dispersive flows. The margin of the flow is solidified by cooling and these solidified crusts control the behavior of successive lava flow. In case of a channel flow, the solidified margin supports the stable central main flow and elongates the lava flow distance. The cross section of lava flow shows that the liquid lava flows between solidified crusts. As for the lava extrusion flow rate, LavaSIM can include the time function as well as the location of the vents. In some cases, some parts of the solidified wall may be broken by the pressure of successive flow and/or re-melting. These mechanisms could characterize complex features of the observed lava flows at many volcanoes in the world. To apply LavaSIM to the benchmark tests organized by V-hub is important to improve the lava flow evaluation technique.

  8. 3N Cave: Longest salt cave in the world

    Czech Academy of Sciences Publication Activity Database

    Bruthans, J.; Filippi, Michal; Zare, M.; Asadi, N.; Vilhelm, Z.

    2006-01-01

    Roč. 64, č. 9 (2006), s. 10-18 ISSN 0027-7010 R&D Projects: GA AV ČR(CZ) KJB301110501 Institutional research plan: CEZ:AV0Z30130516 Keywords : salt cave * salt karst * Iran * expedition Namak Subject RIV: DB - Geology ; Mineralogy

  9. Distribution and dynamics of hayscented fern following stand harvest

    Science.gov (United States)

    Songlin Fei; Peter J. Gould; Melanie J. Kaeser; Kim C. Steiner

    2008-01-01

    The distribution and dynamics of hayscented fern were examined as part of a large-scale study of oak regeneration in Pennsylvania. The study included 69 stands covering 3,333 acres in two physiographic provinces. Hayscented fern was more widely distributed and occurred at higher densities in the Allegheny Plateau physiographic provinces versus the Ridge and Valley...

  10. Resolution of lava tubes with ground penetrating radar: preliminary results from the TubeX project

    Science.gov (United States)

    Esmaeili, S.; Kruse, S.; Garry, W. B.; Whelley, P.; Young, K.; Jazayeri, S.; Bell, E.; Paylor, R.

    2017-12-01

    As early as the mid 1970's it was postulated that planetary tubes or caves on other planetary bodies (i.e., the Moon or Mars) could provide safe havens for human crews, protect life and shield equipment from harmful radiation, rapidly fluctuating surface temperatures, and even meteorite impacts. What is not clear, however, are the exploration methods necessary to evaluate a potential tube-rich environment to locate suitable tubes suitable for human habitation. We seek to address this knowledge gap using a suite of instruments to detect and document tubes in a terrestrial analog study at Lava Beds National Monument, California, USA. Here we describe the results of ground penetrating radar (GPR) profiles and light detection and ranging (LiDAR) scans. Surveys were conducted from the surface and within four lava tubes (Hercules Leg, Skull, Valentine and, Indian Well Caves) with varying flow composition, shape, and complexity. Results are shown across segments of these tubes where the tubes are 10 m in height and the ceilings are 1 - 10 m below the surface. The GPR profiles over the tubes are, as expected, complex, due to scattering from fractures in roof material and three-dimensional heterogeneities. Point clouds derived from the LiDAR scans of both the interior and exterior of the lava tubes provide precise positioning of the tube geometry and depth of the ceiling and floor with respect to the surface topography. GPR profiles over LiDAR-mapped tube cross-sections are presented and compared against synthetic models of radar response to the measured geometry. This comparison will help to better understand the origins of characteristic features in the radar profiles. We seek to identify the optimal data processing and migration approaches to aid lava tube exploration of planetary surfaces.

  11. Towards resolving the complete fern tree of life.

    Science.gov (United States)

    Lehtonen, Samuli

    2011-01-01

    In the past two decades, molecular systematic studies have revolutionized our understanding of the evolutionary history of ferns. The availability of large molecular data sets together with efficient computer algorithms, now enables us to reconstruct evolutionary histories with previously unseen completeness. Here, the most comprehensive fern phylogeny to date, representing over one-fifth of the extant global fern diversity, is inferred based on four plastid genes. Parsimony and maximum-likelihood analyses provided a mostly congruent results and in general supported the prevailing view on the higher-level fern systematics. At a deep phylogenetic level, the position of horsetails depended on the optimality criteria chosen, with horsetails positioned as the sister group either of Marattiopsida-Polypodiopsida clade or of the Polypodiopsida. The analyses demonstrate the power of using a 'supermatrix' approach to resolve large-scale phylogenies and reveal questionable taxonomies. These results provide a valuable background for future research on fern systematics, ecology, biogeography and other evolutionary studies.

  12. The microbiology of Lascaux Cave

    Czech Academy of Sciences Publication Activity Database

    Bastian, F.; Jurado, V.; Nováková, Alena; Alabouvette, C.; Saiz-Jimenez, C.

    2010-01-01

    Roč. 156, č. 3 (2010), s. 644-652 ISSN 1350-0872 Institutional research plan: CEZ:AV0Z60660521 Keywords : Lascaux Cave * microbiology * Paleolithic paintings Subject RIV: EH - Ecology, Behaviour Impact factor: 2.957, year: 2010

  13. Chloroplast genome evolution in early diverged leptosporangiate ferns.

    Science.gov (United States)

    Kim, Hyoung Tae; Chung, Myong Gi; Kim, Ki-Joong

    2014-05-01

    In this study, the chloroplast (cp) genome sequences from three early diverged leptosporangiate ferns were completed and analyzed in order to understand the evolution of the genome of the fern lineages. The complete cp genome sequence of Osmunda cinnamomea (Osmundales) was 142,812 base pairs (bp). The cp genome structure was similar to that of eusporangiate ferns. The gene/intron losses that frequently occurred in the cp genome of leptosporangiate ferns were not found in the cp genome of O. cinnamomea. In addition, putative RNA editing sites in the cp genome were rare in O. cinnamomea, even though the sites were frequently predicted to be present in leptosporangiate ferns. The complete cp genome sequence of Diplopterygium glaucum (Gleicheniales) was 151,007 bp and has a 9.7 kb inversion between the trnL-CAA and trnVGCA genes when compared to O. cinnamomea. Several repeated sequences were detected around the inversion break points. The complete cp genome sequence of Lygodium japonicum (Schizaeales) was 157,142 bp and a deletion of the rpoC1 intron was detected. This intron loss was shared by all of the studied species of the genus Lygodium. The GC contents and the effective numbers of codons (ENCs) in ferns varied significantly when compared to seed plants. The ENC values of the early diverged leptosporangiate ferns showed intermediate levels between eusporangiate and core leptosporangiate ferns. However, our phylogenetic tree based on all of the cp gene sequences clearly indicated that the cp genome similarity between O. cinnamomea (Osmundales) and eusporangiate ferns are symplesiomorphies, rather than synapomorphies. Therefore, our data is in agreement with the view that Osmundales is a distinct early diverged lineage in the leptosporangiate ferns.

  14. Bioaccumulation of eight heavy metals in cave animals from Dashui ...

    African Journals Online (AJOL)

    ajl2

    karst caves and water systems in the caves are well developed. So, heavy metals can contaminate cave envi- ronment and affect cave animals. Karst topography is widely distributed in Guizhou province, China, accounting for 73.8% of the total land area. So, the examination of heavy metal pollution in cave soil and water ...

  15. Environmental conditions of Borra Cave, Visakhapattanam, India

    Directory of Open Access Journals (Sweden)

    Haraprasad Bairagya

    2014-05-01

    Full Text Available Caving is an art which can be best experienced in the mystic Borra of Eastern Ghats and ranked as the second largest cave of India just after Belum Caves situated in the same state Andhrapradesh, India. This Cave is fast becoming a hot tourist?s destination offering great adventurous opportunity to the tourists in the Eastern Ghats. The cave is located in the Ananthagiri hills of the Eastern Ghats region near Visakhapattanam and is made of limestone. The emotion of thrill heightens after entering the cave. The entrance has a narrow vertical opening and is well lit. Due to its location in the sub-equatorial region, dripping of water from the cave roofs occurs almost throughout the year. The formation of stalactites and stalagmites create wonderful phenomena specially found in this cave. The conspicuous pillars formed due to the joining of the roof and the floors are an awe-inspiring creativity of the creator of this world. Various viruses and bacteria are in the cave interior along with different other creatures. The Borra cave helps the Govt. of Andhra Pradesh, India, to earn huge economic benefits for the sake of tourism industry. DOI: http://dx.doi.org/10.3126/ije.v3i2.10526 International Journal of the Environment Vol.3(2 2014: 150-166

  16. Probabilistically modeling lava flows with MOLASSES

    Science.gov (United States)

    Richardson, J. A.; Connor, L.; Connor, C.; Gallant, E.

    2017-12-01

    Modeling lava flows through Cellular Automata methods enables a computationally inexpensive means to quickly forecast lava flow paths and ultimate areal extents. We have developed a lava flow simulator, MOLASSES, that forecasts lava flow inundation over an elevation model from a point source eruption. This modular code can be implemented in a deterministic fashion with given user inputs that will produce a single lava flow simulation. MOLASSES can also be implemented in a probabilistic fashion where given user inputs define parameter distributions that are randomly sampled to create many lava flow simulations. This probabilistic approach enables uncertainty in input data to be expressed in the model results and MOLASSES outputs a probability map of inundation instead of a determined lava flow extent. Since the code is comparatively fast, we use it probabilistically to investigate where potential vents are located that may impact specific sites and areas, as well as the unconditional probability of lava flow inundation of sites or areas from any vent. We have validated the MOLASSES code to community-defined benchmark tests and to the real world lava flows at Tolbachik (2012-2013) and Pico do Fogo (2014-2015). To determine the efficacy of the MOLASSES simulator at accurately and precisely mimicking the inundation area of real flows, we report goodness of fit using both model sensitivity and the Positive Predictive Value, the latter of which is a Bayesian posterior statistic. Model sensitivity is often used in evaluating lava flow simulators, as it describes how much of the lava flow was successfully modeled by the simulation. We argue that the positive predictive value is equally important in determining how good a simulator is, as it describes the percentage of the simulation space that was actually inundated by lava.

  17. A molecular phylogeny of scaly tree ferns (Cyatheaceae).

    Science.gov (United States)

    Korall, Petra; Conant, David S; Metzgar, Jordan S; Schneider, Harald; Pryer, Kathleen M

    2007-05-01

    Tree ferns recently were identified as the closest sister group to the hyperdiverse clade of ferns, the polypods. Although most of the 600 species of tree ferns are arborescent, the group encompasses a wide range of morphological variability, from diminutive members to the giant scaly tree ferns, Cyatheaceae. This well-known family comprises most of the tree fern diversity (∼500 species) and is widespread in tropical, subtropical, and south temperate regions of the world. Here we investigate the phylogenetic relationships of scaly tree ferns based on DNA sequence data from five plastid regions (rbcL, rbcL-accD IGS, rbcL-atpB IGS, trnG-trnR, and trnL-trnF). A basal dichotomy resolves Sphaeropteris as sister to all other taxa and scale features support these two clades: Sphaeropteris has conform scales, whereas all other taxa have marginate scales. The marginate-scaled clade consists of a basal trichotomy, with the three groups here termed (1) Cyathea (including Cnemidaria, Hymenophyllopsis, Trichipteris), (2) Alsophila sensu stricto, and (3) Gymnosphaera (previously recognized as a section within Alsophila) + A. capensis. Scaly tree ferns display a wide range of indusial structures, and although indusium shape is homoplastic it does contain useful phylogenetic information that supports some of the larger clades recognised.

  18. The evolution, morphology and development of fern leaves

    Directory of Open Access Journals (Sweden)

    Alejandra eVasco

    2013-09-01

    Full Text Available Leaves are lateral determinate structures formed in a predictable sequence (phyllotaxy on the flanks of an indeterminate shoot apical meristem. The origin and evolution of leaves in vascular plants has been widely debated. Being the main conspicuous organ of nearl all vascular plants and often easy to recognize as such, it seems surprising that leaves have had multiple origins. For decades, morphologists, anatomists, paleobotanists, and systematists have contributed data to this debate. More recently, molecular genetic studies have provided insight into leaf evolution and development mainly within angiosperms and, to a lesser extent, lycophytes. There has been recent interest in extending leaf evolutionary developmental studies to other species and lineages, particularly in lycophytes and ferns. Therefore, a review of fern leaf morphology, evolution and development is timely. Here we discuss the theories of leaf evolution in ferns, morphology and diversity of fern leaves, and experimental results of fern leaf development. We summarize what is known about the molecular genetics of fern leaf development and what future studies might tell us about the evolution of fern leaf development.

  19. Modeling Submarine Lava Flow with ASPECT

    Science.gov (United States)

    Storvick, E. R.; Lu, H.; Choi, E.

    2017-12-01

    Submarine lava flow is not easily observed and experimented on due to limited accessibility and challenges posed by the fast solidification of lava and the associated drastic changes in rheology. However, recent advances in numerical modeling techniques might address some of these challenges and provide unprecedented insight into the mechanics of submarine lava flow and conditions determining its wide-ranging morphologies. In this study, we explore the applicability ASPECT, Advanced Solver for Problems in Earth's ConvecTion, to submarine lava flow. ASPECT is a parallel finite element code that solves problems of thermal convection in the Earth's mantle. We will assess ASPECT's capability to model submarine lava flow by observing models of lava flow morphology simulated with GALE, a long-term tectonics finite element analysis code, with models created using comparable settings and parameters in ASPECT. From these observations we will contrast the differing models in order to identify the benefits of each code. While doing so, we anticipate we will learn about the conditions required for end-members of lava flow morphology, for example, pillows and sheet flows. With ASPECT specifically we focus on 1) whether the lava rheology can be implemented; 2) how effective the AMR is in resolving morphologies of the solidified crust; 3) whether and under what conditions the end-members of the lava flow morphologies, pillows and sheets, can be reproduced.

  20. Lava tubes - Potential shelters for habitats

    Science.gov (United States)

    Horz, F.

    Natural caverns occur on the moon in the form of 'lava tubes', which are the drained conduits of underground lava rivers. The inside dimensions of these tubes measure tens to hundreds of meters, and their roofs are expected to be thicker than 10 meters. Consequently, lava tube interiors offer an environment that is naturally protected from the hazards of radiation and meteorite impact. Further, constant, relatively benign temperatures of -20 C prevail. These are extremely favorable environmental conditions for human activities and industrial operations. Significant operational, technological, and economical benefits might result if a lunar base were constructed inside a lava tube.

  1. What factors control superficial lava dome explosivity?

    Science.gov (United States)

    Boudon, Georges; Balcone-Boissard, Hélène; Villemant, Benoît; Morgan, Daniel J

    2015-09-30

    Dome-forming eruption is a frequent eruptive style and a major hazard on numerous volcanoes worldwide. Lava domes are built by slow extrusion of degassed, viscous magma and may be destroyed by gravitational collapse or explosion. The triggering of lava dome explosions is poorly understood: here we propose a new model of superficial lava-dome explosivity based upon a textural and geochemical study (vesicularity, microcrystallinity, cristobalite distribution, residual water contents, crystal transit times) of clasts produced by key eruptions. Superficial explosion of a growing lava dome may be promoted through porosity reduction caused by both vesicle flattening due to gas escape and syn-eruptive cristobalite precipitation. Both processes generate an impermeable and rigid carapace allowing overpressurisation of the inner parts of the lava dome by the rapid input of vesiculated magma batches. The relative thickness of the cristobalite-rich carapace is an inverse function of the external lava dome surface area. Explosive activity is thus more likely to occur at the onset of lava dome extrusion, in agreement with observations, as the likelihood of superficial lava dome explosions depends inversely on lava dome volume. This new result is of interest for the whole volcanological community and for risk management.

  2. Pathogenic and opportunistic microorganisms in caves

    Directory of Open Access Journals (Sweden)

    Sanchez-Moral Sergio

    2010-01-01

    Full Text Available With today’s leisure tourism, the frequency of visits to many caves makes it necessary to know about possible potentially pathogenic microorganisms in caves, determine their reservoirs, and inform the public about the consequences of such visits. Our data reveal that caves could be a potential danger to visitors because of the presence of opportunistic microorganisms, whose existence and possible development in humans is currently unknown.

  3. Measurements of radon levels inside Mexican caves

    International Nuclear Information System (INIS)

    Borau, J.; Gonzalez, A.; Espinosa, G.; Golzarri, J.I.

    1993-01-01

    Living animal species on earth have been exposed to environmental radon from the very beginning of time. The effects of radiation, combined with other natural parameters such as temperature, humidity, salt contents, etc., have most likely influenced the evaluation of different species. Thus, it is important to know and to evaluate the radon levels, among other radioactive elements present in enclosed environments such as caves, especially since those caves were also the dwellings and refuge of the predecessor of man. In this work we present radon level measurements inside some caves with vestiges of ancient inhabitats and some recently discovered natural caves, using Nuclear Track Detectors. (author)

  4. Geologic field-trip guide to Medicine Lake Volcano, northern California, including Lava Beds National Monument

    Science.gov (United States)

    Donnelly-Nolan, Julie M.; Grove, Timothy L.

    2017-08-17

    Medicine Lake volcano is among the very best places in the United States to see and walk on a variety of well-exposed young lava flows that range in composition from basalt to rhyolite. This field-trip guide to the volcano and to Lava Beds National Monument, which occupies part of the north flank, directs visitors to a wide range of lava flow compositions and volcanic phenomena, many of them well exposed and Holocene in age. The writing of the guide was prompted by a field trip to the California Cascades Arc organized in conjunction with the International Association of Volcanology and Chemistry of the Earth’s Interior (IAVCEI) quadrennial meeting in Portland, Oregon, in August of 2017. This report is one of a group of three guides describing the three major volcanic centers of the southern Cascades Volcanic Arc. The guides describing the Mount Shasta and Lassen Volcanic Center parts of the trip share an introduction, written as an overview to the IAVCEI field trip. However, this guide to Medicine Lake volcano has descriptions of many more stops than are included in the 2017 field trip. The 23 stops described here feature a range of compositions and volcanic phenomena. Many other stops are possible and some have been previously described, but these 23 have been selected to highlight the variety of volcanic phenomena at this rear-arc center, the range of compositions, and for the practical reason that they are readily accessible. Open ground cracks, various vent features, tuffs, lava-tube caves, evidence for glaciation, and lava flows that contain inclusions and show visible evidence of compositional zonation are described and visited along the route.

  5. Spectroscopic Analysis of Arsenic Uptake in Pteris Ferns

    Directory of Open Access Journals (Sweden)

    Terrence Slonecker

    2009-09-01

    Full Text Available Two arsenic-accumulating Pteris ferns (Pteris cretica mayii and Pteris multifida, along with a non-accumulating control fern (Nephrolepis exaltata were grown in greenhouse conditions in clean sand spiked with 0, 20, 50, 100 and 200 ppm sodium arsenate. Spectral data were collected for each of five replicates prior to harvest at 4-week intervals. Fern samples were analyzed for total metals content and Partial Least Squares and Stepwise Linear Regression techniques were used to develop models from the spectral data. Results showed that Pteris cretica mayii and Pteris multifida are confirmed hyperaccumulators of inorganic arsenic and that reasonably accurate predictive models of arsenic concentration can be developed from the first derivative of spectral reflectance of the hyperaccumulating Pteris ferns. Both the arsenic uptake and spectral results indicate that there is some species-specific variability but the results compare favorably with previously published data and additional research is recommended.

  6. THE ANTIMICROBIAL ACTIVITY OF SOME EXTRACTS OF FERN GAMETOPHYTES

    Directory of Open Access Journals (Sweden)

    Ionica Deliu

    2013-12-01

    Full Text Available The nature freely offers us many resources for health and beauty. The ferns and their therapeutic properties are less exploit in Romania, except Lycopodium clavatum and Equisetum arvense. Some of the fern properties were demonstrated, like antioxidant, antimicrobial, antiviral, antihelmintic properties. Plants are reasonable alternative to synthetic drugs, avoid the side effect and high cost of synthetic drugs production. Also, the drug resistance bacteria can be controlled using plant derived remedies. In this study the antimicrobial effect of methanolic and ethanolic extracts from three fern species were tested. The extracts were gained from gametophytic stage of ferns obtained in vitro. The most obvious effect was observed for Asplenium trichomanes-ramosum extract. The total polyphenols and flavonoids content were established, too.

  7. Organelle Genome Inheritance in Deparia Ferns (Athyriaceae, Aspleniineae, Polypodiales

    Directory of Open Access Journals (Sweden)

    Li-Yaung Kuo

    2018-04-01

    Full Text Available Organelle genomes of land plants are predominately inherited maternally but in some cases can also be transmitted paternally or biparentally. Compared to seed plants (>83% genera of angiosperms and >12% genera of gymnosperms, plastid genome (plastome inheritance has only been investigated in fewer than 2% of fern genera, and mitochondrial genome (mitogenome from only one fern genus. We developed a new and efficient method to examine plastome and mitogenome inheritance in a fern species—Deparia lancea (Athyriaceae, Aspleniineae, Polypodiales, and found that plastid and mitochondrial DNAs were transmitted from only the maternal parentage to a next generation. To further examine whether both organelle genomes have the same manner of inheritance in other Deparia ferns, we sequenced both plastid and mitochondrial DNA regions of inter-species hybrids, and performed phylogenetic analyses to identify the origins of organellar DNA. Evidence from our experiments and phylogenetic analyses support that both organelle genomes in Deparia are uniparentally and maternally inherited. Most importantly, our study provides the first report of mitogenome inheritance in eupolypod ferns, and the second one among all ferns.

  8. Evolutionary and ecological perspectives of Late Paleozoic ferns. Part III. Anachoropterid ferns (including Anachoropteris, Tubicaulis, the Sermayaceae, Kaplanopteridaceae and Psalixochlaenaceae)

    OpenAIRE

    Galtier, Jean; Phillips, Tom L.

    2014-01-01

    The anachoropterid ferns, previously assigned to the family Anachoropteridaceae, are a group of anatomically preserved late Paleozoic filicalean ferns characterized by a C-shaped foliar xylem with abaxially recurved arms (inversicatenalean anatomy) and two main protoxylem strands. The variously curved to strongly inrolled foliar xylem certainly reflects different evolutionary trends within the morphogenus Anachoropteris. The occurrence of two groups of Tubicaulis is supported by differences i...

  9. Millipedes (Diplopoda) from caves of Portugal

    DEFF Research Database (Denmark)

    P. S. Reboleira, Ana Sofia; Enghoff, Henrik

    2014-01-01

    in caves of the mainland and the island of Madeira has provided new data about the distribution and diversity of millipedes. A review of millipedes from caves of Portugal is presented, listing fourteen species belonging to eight families, among which six species are considered troglobionts...

  10. Microbiological and environmental issues in show caves.

    Science.gov (United States)

    Saiz-Jimenez, Cesareo

    2012-07-01

    Cultural tourism expanded in the last half of the twentieth century, and the interest of visitors has come to include caves containing archaeological remains. Some show caves attracted mass tourism, and economical interests prevailed over conservation, which led to a deterioration of the subterranean environment and the rock art. The presence and the role of microorganisms in caves is a topic that is often ignored in cave management. Knowledge of the colonisation patterns, the dispersion mechanisms, and the effect on human health and, when present, over rock art paintings of these microorganisms is of the utmost importance. In this review the most recent advances in the study of microorganisms in caves are presented, together with the environmental implications of the findings.

  11. The Mammoth Cave system, Kentucky, USA

    International Nuclear Information System (INIS)

    Palmer, A. N.

    2016-01-01

    Mammoth Cave is the main attraction of Mammoth Cave National Park. For several decades it has been the longest known cave in the world and currently contains 652 km in 2016 of surveyed passages. It is located in the heart of an extensive karst plateau, in which the stratal dip averages only one degree. The cave is part of a drainage basin of more than 200 km 2 . The cave has been known to local inhabitants for several millennia and contains a rich trove of archaeological and historical artifacts. It contains many speleo biota including several rare and endangered species and has been designated a World Heritage Site and an International Biosphere Reserve (UNESCO). Its many passage levels and sediments contain a record of the fluvial history of most of south-eastern North America. (Author)

  12. Reconstructed tree fern Alethopteris zeilleri (Carboniferous, Medullosales)

    Energy Technology Data Exchange (ETDEWEB)

    Zodrow, Erwin L. [Department of Earth Sciences, University College of Cape Breton, Sydney, Nova, Scotia (Canada B1P 6L2)

    2007-01-02

    From a smaller open-pit area in the roof shale of the basal Cantabrian coal seam in Sydney Coalfield, Cape Breton Island, Nova Scotia, Canada, large amounts of the pteridosperm foliage Alethopteris zeilleri (Ragot) were found. This foliage is associated with abundant, naked medullosalean axes and dichotomies of varying sizes, up to 0.80-m long, cauline structures 0.90 m and 1.3 m long, detached ovules assigned to Pachytesta incrassata Brongniart, rare male-pollen organs of the type Dolerotheca Halle, rooted tree ferns in life position, and one specimen each of a juvenile medullosalean frond and root mantle. The fossils are compression/impression-preserved, and the foliage yielded thickly cutinized cuticles with unoriented cells (57-103 by 27-57 {mu}m) in intercostal fields. Ultimate rachises, and abaxial surfaces (excluding costal fields) show a mixture of simple and complexly-branched trichomes, and two different structural bases. These, together with fractal dimensionality of curvatures of anticlinal walls in intercostal fields, have taxonomic potential for alethopterids. The finds suggest reconstructing A. zeilleri (Ragot) as a tree, 5-7 m high, that bore both P. incrassata Brongniart and Dolerotheca-type fructifications. Its habitat was low-land coastal plains in the Pennsylvanian coal swamps of ancestral Sydney Coalfield. (author)

  13. Geomagnetic polarity zones for icelandic lavas

    Science.gov (United States)

    Dagley, P.; Wilson, R.L.; Ade-Hall, J. M.; Walker, G.P.L.; Haggerty, S.E.; Sigurgeirsson, T.; Watkins, N.D.; Smith, P.J.; Edwards, J.; Grasty, R.L.

    1967-01-01

    Analysis of cores collected from a sequence of lavas in Eastern Iceland has made possible an accurate calculation of the average rate of reversal of the Earth's magnetic field. ?? 1967 Nature Publishing Group.

  14. Taylor instability in rhyolite lava flows

    Science.gov (United States)

    Baum, B. A.; Krantz, W. B.; Fink, J. H.; Dickinson, R. E.

    1989-01-01

    A refined Taylor instability model is developed to describe the surface morphology of rhyolite lava flows. The effect of the downslope flow of the lava on the structures resulting from the Taylor instability mechanism is considered. Squire's (1933) transformation is developed for this flow in order to extend the results to three-dimensional modes. This permits assessing why ridges thought to arise from the Taylor instability mechanism are preferentially oriented transverse to the direction of lava flow. Measured diapir and ridge spacings for the Little and Big Glass Mountain rhyolite flows in northern California are used in conjunction with the model in order to explore the implications of the Taylor instability for flow emplacement. The model suggests additional lava flow features that can be measured in order to test whether the Taylor instability mechanism has influenced the flows surface morphology.

  15. Towards resolving the complete fern tree of life.

    Directory of Open Access Journals (Sweden)

    Samuli Lehtonen

    Full Text Available In the past two decades, molecular systematic studies have revolutionized our understanding of the evolutionary history of ferns. The availability of large molecular data sets together with efficient computer algorithms, now enables us to reconstruct evolutionary histories with previously unseen completeness. Here, the most comprehensive fern phylogeny to date, representing over one-fifth of the extant global fern diversity, is inferred based on four plastid genes. Parsimony and maximum-likelihood analyses provided a mostly congruent results and in general supported the prevailing view on the higher-level fern systematics. At a deep phylogenetic level, the position of horsetails depended on the optimality criteria chosen, with horsetails positioned as the sister group either of Marattiopsida-Polypodiopsida clade or of the Polypodiopsida. The analyses demonstrate the power of using a 'supermatrix' approach to resolve large-scale phylogenies and reveal questionable taxonomies. These results provide a valuable background for future research on fern systematics, ecology, biogeography and other evolutionary studies.

  16. Internal fabric development in complex lava domes

    Czech Academy of Sciences Publication Activity Database

    Závada, Prokop; Kratinová, Zuzana; Kusbach, V.; Schulmann, K.

    2009-01-01

    Roč. 466, č. 1-2 (2009), s. 101-113 ISSN 0040-1951 R&D Projects: GA AV ČR KJB301110703; GA AV ČR KJB300120702 Grant - others:GA ČR(CZ) GA205/03/0204 Institutional research plan: CEZ:AV0Z30120515 Keywords : analogue modeling * lava extrusion * exogenous growth * crystal-rich lava * AMS Subject RIV: DB - Geology ; Mineralogy Impact factor: 1.935, year: 2009

  17. PIXE analysis of cave sediments, prehispanic paintings and obsidian cutting tools from Baja California Sur caves

    International Nuclear Information System (INIS)

    Miranda, J.; Oliver, A.; Dacal, A.; Ruvalcaba, J.L.; Cruz, F.; Ortiz, M.E.; Vinas, R.

    1993-01-01

    Elemental PIXE analysis of cave sediments, minerals, pigments of the prehispanic paintings and obsidian cutting tools from caves in Baja California Sur has been carried out with a 0.7 MeV proton beam. The elements analysed in this sample set (Al to Co) provide an idea of the environment of the caves. The obsidian data analysis suggests that the human communities who made these painting used more than one obsidian source to manufacture cutting tools. (orig.)

  18. Bacterial community survey of sediments at Naracoorte Caves, Australia

    Directory of Open Access Journals (Sweden)

    Ball Andrew S.

    2012-07-01

    Full Text Available Bacterial diversity in sediments at UNESCO World Heritage listed Naracoorte Caves was surveyed as part of an investigation carried out in a larger study on assessing microbial communities in caves. Cave selection was based on tourist accessibility; Stick Tomato and Alexandra Cave (> 15000 annual visits and Strawhaven Cave was used as control (no tourist access. Microbial analysis showed that Bacillus was the most commonly detected microbial genus by culture dependent and independent survey of tourist accessible and inaccessible areas of show (tourist accessible and control caves. Other detected sediment bacterial groups were assigned to the Firmicutes, Actinobacteria and Proteobacteria. The survey also showed differences in bacterial diversity in caves with human access compared to the control cave with the control cave having unique microbial sequences (Acinetobacter, Agromyces, Micrococcus and Streptomyces. The show caves had higher bacterial counts, different 16S rDNA based DGGE cluster patterns and principal component groupings compared to Strawhaven. Different factors such as human access, cave use and configurations could have been responsible for the differences observed in the bacterial community cluster patterns (tourist accessible and inaccessible areas of these caves. Cave sediments can therefore act as reservoirs of microorganisms. This might have some implications on cave conservation activities especially if these sediments harbor rock art degrading microorganisms in caves with rock art.

  19. Analyis on the vascular epiphytes of tree ferns in a montane rain forest in Costa Rica

    OpenAIRE

    Jens Bittner; Javier Trejos Zelaya

    2009-01-01

    The relationships between epiphytes and host specifity are known. We investigated the relation between specific epiphytes and tree fern trunks. Only some epiphytes are frequent or very frequent on tree ferns. Most of the epiphyte species are unspecific on these trunks. Blechnum fragile are the only found exclusively on tree ferns. We also observed different epiphyte communities dependent on the tree fern species and the morphology of the tree trunk.

  20. Analyis on the vascular epiphytes of tree ferns in a montane rain forest in Costa Rica

    Directory of Open Access Journals (Sweden)

    Jens Bittner

    2009-02-01

    Full Text Available The relationships between epiphytes and host specifity are known. We investigated the relation between specific epiphytes and tree fern trunks. Only some epiphytes are frequent or very frequent on tree ferns. Most of the epiphyte species are unspecific on these trunks. Blechnum fragile are the only found exclusively on tree ferns. We also observed different epiphyte communities dependent on the tree fern species and the morphology of the tree trunk.

  1. NAK WP-Cave project

    International Nuclear Information System (INIS)

    Svemar, C.

    1985-11-01

    WP-Cave is designed as an egg-shaped underground structure for intermediate storing and final disposal of high-level nuclear waste. Its height, when storing 1600 tonnes of spent fuel, is about 250 m and its diameter 110 m at mid-height. The waste storage has a compact layout and is surrounded by two engineered barriers. The innermost one is a 5 m-wide shield consisting of a mixture of bentonite clay and same which has a low hydraulic conductivity. This shield is surrounded by a so-called hydraulic cage, which initially drains the storage rock mass and, in the long-term diverts, the ground water flow past the storage. In this way an initial dry supervision period can be maintained. After sealing-off the storage and after water-filling, a stagnant chemical environment is established inside the bentonite-sand barrier preventing the disposed waste from being dissolved and from migrating to the geosphere. The programme, as outlined by the Project Board, has considered R and D questions with specific relation to the WP-Cave such as: properties of low-graded bentonite mixtures, function of the hydraulic cage, full-face boring of the storage, geomechanics of the storage and the bentonite-sand barrier, dry ventilation of the storage, temperature distributions and thermal stresses. An initial safety analysis has also been conducted. The hydraulic conductivity of low-grade bentonite mixtures has been measured in laboratory tests and found to be higher than expected. Tests of gas conductivity, for instance, confirm that only low gas pressures would build up inside the bentonite-sand barrier. The initial safety analysis indicates that a compact storage, such as that presented, would allow for the safe isolation of the spent nuclear fuel and would fulfull the radiation protection criterion of 0.1 mSv/year. With 27 refs. (Author)

  2. El historiador Fernández de Piedrahita

    Directory of Open Access Journals (Sweden)

    Carlos López Narváez

    1964-10-01

    Full Text Available Varón de Dios, las letras y la patria, Lucas Fernández de Piedrahita nació en Santafé de Bogotá, el 6 de marzo de 1624, de hidalgo linaje por su padre español, y de principesca sangre materna, como que su progenitora, doña Catalina Collantes, esposa de don Domingo Fernández de Soto y Piedrahita, era nieta de doña Francisca Coya de la realeza inca del Perú.

  3. Uptake of rare earth elements by dryopteris erythrosora (autumn fern)

    International Nuclear Information System (INIS)

    Ozaki, Takuo; Enomoto, Shuichi

    2001-01-01

    Mechanisms of uptake of rare earth elements (REEs) were investigated, particularly those by REE accumulator species (autumn fern). Rare earth elements are practically insoluble under natural conditions, suggesting some unknown mechanisms in REE accumulator species. In the present investigation, two notable phenomena were observed. (1) Concerning the ionic-radius dependence of REE uptake by leaves, nonaccumulator species showed an extremely high uptake for Y compared with the adjacent-ionic-radius REEs in the multitracer, while accumulator species showed no anomaly. (2) REE uptake by autumn fern was influenced by the addition of chelating chemical reagents in the uptake solution, while no effect was observed for nonaccumulator species. (author)

  4. Visualization framework for CAVE virtual reality systems

    OpenAIRE

    Kageyama, Akira; Tomiyama, Asako

    2016-01-01

    We have developed a software framework for scientific visualization in immersive-type, room-sized virtual reality (VR) systems, or Cave automatic virtual environment (CAVEs). This program, called Multiverse, allows users to select and invoke visualization programs without leaving CAVE’s VR space. Multiverse is a kind of immersive “desktop environment” for users, with a three-dimensional graphical user interface. For application developers, Multiverse is a software framework with useful class ...

  5. Stress analysis of longwall top coal caving

    Energy Technology Data Exchange (ETDEWEB)

    Alehossein, H.; Poulsen, B.A. [CSIRO Exploration & Mining, Brisbane, Qld. (Australia)

    2010-01-15

    Longwall top coal caving (LTCC) is a relatively new method of mining thick coal seams that is currently achieving high productivity and efficiency. The technique is similar to traditional longwall mining in that a cutting head slices coal from the lower section of the coal seam onto a conveyor belt installed in front of the hydraulic support near the cutting face. In modern LTCC an additional rear conveyor belt is located behind the support, to which the flow of the caved coal from the upper part of the seam can be controlled by a moveable flipper attached to the canopy of the support. The mining method relies on the fracturing of the top coal by the front abutment pressure to achieve satisfactory caving into the rear conveyor. This paper develops a yield and caveability criterion based on in situ conditions in the top coal in advance of the mining face (yield) and behind the supports (caveability). Yielding and caving effects are combined into one single number called caving number (CN), which is the multiplication result of caving factor (CF) and yield factor (YF). Analytical derivations are based on in situ stress conditions, Mohr-Coulomb and/or Hoek-Brown rock failure criteria and an on-associated elastoplastic strain softening material behaviour. The yield and caveability criteria are in agreement with results from both numerical studies and mine data. The caving number is normalised to mining conditions of a reference Chinese mine (LMX mine) and is used to assess LTCC performance at fourteen other Chinese working longwalls that have had varying success with the LTCC technology. As a predictive model, results of this analytical/numerical study are useful to assess the potential success of caving in new LTCC operations and in different mining conditions.

  6. Distinction between epigenic and hypogenic maze caves

    Science.gov (United States)

    Palmer, Arthur N.

    2011-11-01

    Certain caves formed by dissolution of bedrock have maze patterns composed of closed loops in which many intersecting fractures or pores have enlarged simultaneously. Their origin can be epigenic (by shallow circulation of meteoric groundwater) or hypogenic (by rising groundwater or production of deep-seated solutional aggressiveness). Epigenic mazes form by diffuse infiltration through a permeable insoluble caprock or by floodwater supplied by sinking streams. Most hypogenic caves involve deep sources of aggressiveness. Transverse hypogenic cave origin is a recently proposed concept in which groundwater of mainly meteoric origin rises across strata in the distal portions of large flow systems, to form mazes in soluble rock sandwiched between permeable but insoluble strata. The distinction between maze types is debated and is usually based on examination of diagnostic cave features and relation of caves to their regional setting. In this paper, the principles of mass transfer are applied to clarify the limits of each model, to show how cave origin is related to groundwater discharge, dissolution rate, and time. The results show that diffuse infiltration and floodwater can each form maze caves at geologically feasible rates (typically within 500 ka). Transverse hypogenic mazes in limestone, to enlarge significantly within 1 Ma, require an unusually high permeability of the non-carbonate beds (generally ≥ 10-4 cm/s), large discharge, and calcite saturation no greater than 90%, which is rare in deep diffuse flow in sedimentary rocks. Deep sources of aggressiveness are usually required. The origin of caves by transverse hypogenic flow is much more favorable in evaporite rocks than in carbonate rocks.

  7. Hypogene caves of the central Appalachian Shenandoah Valley in Virginia

    Science.gov (United States)

    Doctor, Daniel H.; Orndorff, Wil

    2017-01-01

    Several caves in the Shenandoah Valley in Virginia show evidence for early hypogenic conduit development with later-enhanced solution under partly confined phreatic conditions guided by geologic structures. Many (but not all) of these caves have been subsequently invaded by surface waters as a result of erosion and exhumation. Those not so affected are relict phreatic caves, bearing no relation to modern drainage patterns. Field and petrographic evidence shows that carbonate rocks hosting certain relict phreatic caves were dolomitized and/or silicified by early hydrothermal fluid migration in zones that served to locally enhance rock porosity, thus providing preferential pathways for later solution by groundwater flow, and making the surrounding bedrock more resistant to surficial weathering to result in caves that reside within isolated hills on the land surface. Features suggesting that deep phreatic processes dominated the development of these relict caves include (1) cave passage morphologies indicative of ascending fluids, (2) cave plans of irregular pattern, reflecting early maze or anastomosing development, (3) a general lack of cave breakdown and cave streams or cave stream deposits, and (4) calcite wall and pool coatings within isolated caves intersecting the local water table, and within unroofed caves at topographic locations elevated well above the local base level. Episodes of deep karstification were likely separated by long periods of geologic time, encompassing multiple phases of sedimentary fill and excavation within caves, and reflect a complex history of deep fluid migration that set the stage for later shallow speleogenesis that continues today.

  8. Exposure to radon in Australian tourist caves

    International Nuclear Information System (INIS)

    Solomon, S.B.; Langroo, R.; Lyons, R.G.

    1994-01-01

    In 1991 the International Commission on Radiological Protection(ICRP) produced guidelines and recommendations dealing with workplace exposure to elevated background radiation, in particular, the risk associated with the inhalation of radon and radon progeny. An intervention level of 1000 Bq m -3 has been proposed. Australia has over 40 tourist caves, under the management of the various State Departments or private groups. The limited data available on radon levels in Australian caves would suggest that some of these caves may be in excess of the proposed intervention level, thus presenting a potential health risk for the cave guides. This paper summarises the current information on radon in Australian caves and describes the proposed methodologies to be used for a Worksafe Australia-funded survey of radon levels in Tourist caves within Australia. This survey is to be carried out jointly by researchers at the Australian Radiation Laboratory, University of Auckland and the University of Sydney, during 1994 and 1995. 7 refs., 3 tabs

  9. Genomic sequencing of Pleistocene cave bears

    Energy Technology Data Exchange (ETDEWEB)

    Noonan, James P.; Hofreiter, Michael; Smith, Doug; Priest, JamesR.; Rohland, Nadin; Rabeder, Gernot; Krause, Johannes; Detter, J. Chris; Paabo, Svante; Rubin, Edward M.

    2005-04-01

    Despite the information content of genomic DNA, ancient DNA studies to date have largely been limited to amplification of mitochondrial DNA due to technical hurdles such as contamination and degradation of ancient DNAs. In this study, we describe two metagenomic libraries constructed using unamplified DNA extracted from the bones of two 40,000-year-old extinct cave bears. Analysis of {approx}1 Mb of sequence from each library showed that, despite significant microbial contamination, 5.8 percent and 1.1 percent of clones in the libraries contain cave bear inserts, yielding 26,861 bp of cave bear genome sequence. Alignment of this sequence to the dog genome, the closest sequenced genome to cave bear in terms of evolutionary distance, revealed roughly the expected ratio of cave bear exons, repeats and conserved noncoding sequences. Only 0.04 percent of all clones sequenced were derived from contamination with modern human DNA. Comparison of cave bear with orthologous sequences from several modern bear species revealed the evolutionary relationship of these lineages. Using the metagenomic approach described here, we have recovered substantial quantities of mammalian genomic sequence more than twice as old as any previously reported, establishing the feasibility of ancient DNA genomic sequencing programs.

  10. Nornahraun lava morphology and mode of emplacement

    Science.gov (United States)

    Pedersen, Gro B. M.; Höskuldsson, Armann; Riishuus, Morten S.; Jónsdóttir, Ingibjörg; Gudmundsson, Magnús T.; Sigmundsson, Freysteinn; Óskarsson, Birgir V.; Drouin, Vincent; Gallagher, Catherine; Askew, Rob; Moreland, William M.; Dürig, Tobias; Dumont, Stephanie; Þórdarson, Þór

    2015-04-01

    The ongoing Nornahraun eruption is the largest effusive eruption in Iceland since the Laki eruption in 1783-84, with an estimated lava volume of ~1.15 km3 covering an area of ~83.4 km2 (as of 5 JAN 2015). The eruption provides an unprecedented opportunity to study i) lava morphologies and their emplacement styles, ii) the transition from from open to closed lava pathways and iii) lava pond formation. Tracking of the lava advancement and morphology has been performed by GPS and GoPro cameras installed in 4×4 vehicles as well as video footage. Complimentary observations have been provided from aircraft platforms and by satellite data. Of particular importance for lava morphology observations are 1-12 m/pixel airborne SAR images (x-band). The Nornahraun flow field comprises a continuum of morphologies from pāhoehoe to 'a'ā, which have varied tem-porally and spatially. At the onset of the eruption 31 AUG, lava flows advanced rapidly (400-800 m/hr) from the 1.5 km long fissure as large slabby pāhoehoe [1-3] sheet lobes, 100-500 m wide and 0.3-1 m thick at the flow fronts. By 1 SEPT, the flows began channeling towards the NE constrained by the older Holuhraun I lava field and the to-pography of flood plain itself. A central open channel developed, feeding a 1-2 km wide active 'a'ā frontal lobe that advanced 1-2 km/day. In addition to its own caterpillar motion, the frontal lobe advanced in a series of 30-50 m long breakouts, predominantly slabby and rubbly pāhoehoe [4,5]. These breakouts had initial velocities of 10-30 m/hr and reached their full length within tens of minutes and subsequently inflated over hours. With the continuous advancement of the 'a'ā flow front, the breakouts were incorporated into the 'a'ā flow fronts and seldom preserved. At the margins of the frontal lava lobe, the breakouts were more sporadic, but predominantly rubbly pāhoehoe and slabby pāhoehoe, as at the flow front. The lava flow advanced ENE into Jökulsá á Fjöllum on 7 SEPT

  11. Eleven bones: More fossil remains of Cave Lions and Cave Hyaenas from the North Sea area

    NARCIS (Netherlands)

    Bosscha Erdbrink, D.P.

    1983-01-01

    Six fossil Cave Lion bones and five fossil Cave Hyaena bones are described. One lion bone and one hyaena bone were dredged from the Westerschelde ( = Western Scheldt, southwestern part of the Netherlands). The other specimens were recovered from the bottom of the North Sea, in the area West and

  12. Millipedes (Diplopoda of twelve caves in Western Mecsek, Southwest Hungary

    Directory of Open Access Journals (Sweden)

    Angyal, D.

    2013-11-01

    Full Text Available Twelve caves of Western Mecsek, Southwest Hungary were examined between September 2010 and April 2013from the millipede (Diplopoda faunistical point of view. Ten species were found in eight caves, which consistedeutroglophile and troglobiont elements as well. The cave with the most diverse fauna was the Törökpince Sinkhole, while thetwo previously also investigated caves, the Abaligeti Cave and the Mánfai-kőlyuk Cave provided less species, which couldbe related to their advanced touristic and industrial utilization.

  13. New observations and synthesis of paleogene heterosporous water ferns

    NARCIS (Netherlands)

    Collinson, M.E.; Smith, S.Y.; van Konijnenburg-van Cittert, J.H.A.; Batten, D.J.; Refidaff, C.; Barke, J.; Marone, F.

    Premise of research. Reproductive structures of modern genera of heterosporous water ferns (Marsileaceae and Salviniaceae) are widespread and abundant in plant mesofossil assemblages from the Paleogene. For Salviniaceae, whole fertile fossil plants give a good understanding of morphology. These

  14. In vitro mutagenesis of commercial fern, Asplenium nidus from spores

    International Nuclear Information System (INIS)

    Norazlina Noordin

    2004-01-01

    Asplenium is a largest, most diverse fern genera. One of the common species is Asplenium nidus, well known as Bird's-nest fern, a medium to large fern with erect, stout, unbranched rhizomes. In creating variability of ferns for the benefit of the ornamental plant industry, in vitro mutagenesis is used. In this study, spores of Asplenium nidus were collected from frond bearing mature sporangia. Spores were cultured in modified 1/2 MS basal medium supplemented with various combinations of 6-Benzylaminopurine (BAP) and Naphtalene Acetic Acid (NAA). Spore cultures were incubated in incubation room at 24 degree C with 16 hours photoperiod (3500 lux). It was found that, the most effective combinations were 1 mg/1 BAP + 0. 1 mg/1 NAA and 2mg/1 BAP + 0. 1 mg/1 NAA. Prothallus was formed after 10 days of cultures and gametophytes were formed 1 month later. These gametophytes were irradiated with Gamma ray at doses of 0, 20, 90, 120, 150 and 180 Gy. From the preliminary result obtained from this study, for generating variations and desired phenotypic expression for Asplenium nidus, recommended doses for in vitro mutagenesis using spores are between 90 Gy to 150 Gy. Gametophytes were subcultured at monthly interval to ensure further development and propagation. Frequent monitoring for any changes in the morphology of the irradiated Asplenium nidus plants were carried out. (Author)

  15. The omnipresent water fern Azolla caroliniana does not contain lignin

    NARCIS (Netherlands)

    Nierop, K.G.J.; Speelman, E.N.; Leeuw, J.W. de; Reichart, G.-J.

    2011-01-01

    Several studies have reported the presence of large amounts of lignin in ubiquitously occurring species of the freshwater fern Azolla. Molecular analysis using flash pyrolysis and thermally assisted hydrolysis and methylation shows, however, that neither the leaves nor the roots of Azolla contain

  16. Tear ferning in normal dogs and dogs with keratoconjunctivitis sicca ...

    African Journals Online (AJOL)

    This study evaluates tear ferning as an ancillary technique for the evaluation of the canine tear film in normal eyes and eyes affected by keratoconjunctivitis sicca (KCS). Thirty dogs with KCS and 50 control dogs with normal tear film were evaluated with a full ophthalmoscopic examination and a Schirmer tear test type 1 ...

  17. First megafossil evidence of Cyatheaceous tree fern from the Indian

    Indian Academy of Sciences (India)

    A part of the compressed tree fern axis with leaf and adventitious root scars in unusual arrangement from Plio–Pleistocene sediments of Arunachal Pradesh, India is described as Cyathea siwalika sp. nov. This record suggests that Cyathea was an important component of tropical evergreen forest in the area during the ...

  18. Instructions for collecting tree ferns of the family Cyatheaceae

    NARCIS (Netherlands)

    Holttum, R.E.

    1957-01-01

    The next monographic study which will be undertaken for the series Pteridophyta of the Flora Malesiana will be devoted to the tree ferns of the Cyatheaceae. In connection with the large size of these plants and the desirability of having more and complete material at our disposal, the following

  19. The colonisation of woodland gaps by ferns and horsetails

    NARCIS (Netherlands)

    Bremer, P.

    2010-01-01

    In the Voorsterbos, a planted woodland on a former sea-floor (the Netherlands), artificial gaps within stands of Fagus sylvatica on boulder clay were monitored for five or six years after cutting. Ten fern species and three species of horsetail established in these gaps, with Dryopteris cristata,

  20. New fern records for Kilimanjaro | Hemp | Journal of East African ...

    African Journals Online (AJOL)

    Working on the flora and vegetation of Mt Kilimanjaro, Tanzania, 17 ferns in 10 families were found that had not yet been recorded for the floral region T2. The altitudinal range, localities and habitat description are given for Adiantum raddianum, Asplenium bugoiense, Blechnum ivohibense, Blotiella stipitata, Dryopteris ...

  1. Three new fern records for Kilimanjaro | Hemp | Journal of East ...

    African Journals Online (AJOL)

    Working on the flora and vegetation of Mt. Kilimanjaro, Tanzania, three ferns in three families were found that are not yet recorded for the floral region T2. The altitudinal range, localities and habitat description are given for Adiantum reniforme, Azolla africana and Trichomanes radicans. Journal of East African Natural History ...

  2. Stasis and convergence characterize morphological evolution in eupolypod II ferns.

    Science.gov (United States)

    Sundue, Michael A; Rothfels, Carl J

    2014-01-01

    Patterns of morphological evolution at levels above family rank remain underexplored in the ferns. The present study seeks to address this gap through analysis of 79 morphological characters for 81 taxa, including representatives of all ten families of eupolypod II ferns. Recent molecular phylogenetic studies demonstrate that the evolution of the large eupolypod II clade (which includes nearly one-third of extant fern species) features unexpected patterns. The traditional 'athyrioid' ferns are scattered across the phylogeny despite their apparent morphological cohesiveness, and mixed among these seemingly conservative taxa are morphologically dissimilar groups that lack any obvious features uniting them with their relatives. Maximum-likelihood and maximum-parsimony character optimizations are used to determine characters that unite the seemingly disparate groups, and to test whether the polyphyly of the traditional athyrioid ferns is due to evolutionary stasis (symplesiomorphy) or convergent evolution. The major events in eupolypod II character evolution are reviewed, and character and character state concepts are reappraised, as a basis for further inquiries into fern morphology. Characters were scored from the literature, live plants and herbarium specimens, and optimized using maximum-parsimony and maximum-likelihood, onto a highly supported topology derived from maximum-likelihood and Bayesian analysis of molecular data. Phylogenetic signal of characters were tested for using randomization methods and fitdiscrete. The majority of character state changes within the eupolypod II phylogeny occur at the family level or above. Relative branch lengths for the morphological data resemble those from molecular data and fit an ancient rapid radiation model (long branches subtended by very short backbone internodes), with few characters uniting the morphologically disparate clades. The traditional athyrioid ferns were circumscribed based upon a combination of

  3. The Mammoth Cave system, Kentucky, USA; El sistema de la Mammoth Cave, Kentucky, EE.UU

    Energy Technology Data Exchange (ETDEWEB)

    Palmer, A. N.

    2016-07-01

    Mammoth Cave is the main attraction of Mammoth Cave National Park. For several decades it has been the longest known cave in the world and currently contains 652 km in 2016 of surveyed passages. It is located in the heart of an extensive karst plateau, in which the stratal dip averages only one degree. The cave is part of a drainage basin of more than 200 km{sup 2}. The cave has been known to local inhabitants for several millennia and contains a rich trove of archaeological and historical artifacts. It contains many speleo biota including several rare and endangered species and has been designated a World Heritage Site and an International Biosphere Reserve (UNESCO). Its many passage levels and sediments contain a record of the fluvial history of most of south-eastern North America. (Author)

  4. The Payun-Matru lava field: a source of analogues for Martian long lava flows

    Science.gov (United States)

    Giacomini, L.; Pasquarè, G.; Massironi, M.; Frigeri, A.; Bistacchi, A.; Frederico, C.

    2007-08-01

    The Payun Matru Volcanic complex is a Quaternary fissural structure belonging to the back-arc extensional area of the Andes in the Mendoza Province (Argentina). The eastern portion of the volcanic structure is covered by a basaltic field of pahoehoe lava flows advanced over more than 180 km from the fissural feeding vents that are aligned with a E-W fault system (Carbonilla fault). Thanks to their widespread extension, these flows represent some of the largest lava flows in the world and the Pampas Onduladas flow can be considered the longest sub-aerial individual lava flow on the Earth surface [1,2]. These gigantic flows propagated over the nearly flat surface of the Pampean foreland, moving on a 0.3 degree slope. The very low viscosity of the olivine basalt lavas, coupled with the inflation process and an extensive system of lava tubes are the most probable explanation for their considerable length. The inflation process likely develop under a steady flow rate sustained for a long time [3]. A thin viscoelastic crust, built up at an early stage, is later inflated by the underlying fluid core, which remains hot and fluid thanks to the thermal-shield effect of the crust. The crust is progressively thickened by accretion from below and spreading is due to the continuous creation of new inflated lobes, which develop at the front of the flow. Certain morphological features are considered to be "fingerprints" of inflation [4, 5, 6]; these include tumuli, lava rises, lava lobes and ridges. All these morphologies are present in the more widespread Payun Matru lava flows that, where they form extensive sheetflows, can reach a maximum thickness of more than 20 meters. After the emplacement of the major flows, a second eruptive cycle involved the Payun Matru volcanic structure. During this stage thick and channelized flows of andesitic and dacitic lavas, accompanied the formation of two trachitic and trachiandesitic strato-volcanoes (Payun Matru and Payun Liso) culminated

  5. The show cave of Diros vs. wild caves of Peloponnese, Greece - distribution patterns of Cyanobacteria

    Directory of Open Access Journals (Sweden)

    Vasiliki Lamprinou

    2014-09-01

    Full Text Available The karst cave ‘Vlychada’of Diros, one of the oldest show caves in Peloponnese, sustains extended phototrophic biofilms on various substrata – on rocks inside the cave including speleothems, and especially near the artificial lighting installation (‘Lampenflora’. After a survey of the main abiotic parameters (Photosynthetically Active Radiation -PAR, Temperature -T, Relative Humidity -RH, Carbon Dioxide -CO2 three clusters of sampling sites were revealed according to Principal Component Analysis (PCA: i the water gallery section predominately influenced by CO2, ii the dry passages influenced by RH and PAR, and iii the area by the cave exit at the dry section influenced by temperature. The collected samples from the water gallery section and the dry passages of the cave revealed a total of 43 taxa of Cyanobacteria, with the unicellular/colonial forms being the most abundant. The applied non-metric Multi-dimensional Scaling Ordination (nMDS of the cumulative species composition showed a clear distinction between the water gallery section and the dry passages of the cave. Further comparison with previous data from other wild caves of Peloponnese (‘Kastria’, ‘Francthi’, and ‘Selinitsa’ was conducted revealing a distinction between the show cave and the wild ones. Apart from the human impact on cave ecosystems – through aesthetic alteration (‘greening’ of cave decorations by the ‘Lampenflora’, and by the cleaning treatments and restoration projects on the speleothems – identification of the organisms constituting the ‘Lampenflora’ might provide taxonomically and ecologically significant taxa.

  6. Dating the Lascaux cave gour formation

    International Nuclear Information System (INIS)

    Genty, D.; Touma, M.; Konik, S.; Konik, S.; Valladas, H.; Hellstrom, J.; Moreau, C.; Dumoulin, J.P.; Nouet, J.; Dauphin, Y.; Weil, R.

    2011-01-01

    Lascaux Cave is renowned for its outstanding prehistoric paintings, strikingly well-preserved over about 18,000 yr. While stalagmites and stalactites are almost absent in the cave, there is an extensive calcite flow-stone that covered a large part of the cave until its opening for tourists during the 1950's. The deposit comprises a succession of calcite rims, or 'gours', which allowed seepage water to pond in large areas in the cave. Their possible role in preservation of the cave paintings has often been evoked, but until now this deposit has not been studied in detail. Here, we present 24 new radiocarbon accelerator mass spectrometry (AMS) and 6 uranium-thorium (U-Th) analyses from the calcite of the gours, 4 AMS 14 C dates from charcoals trapped in the calcite, and 4 AMS 14 C analyses on organic matter extracted from the calcite. Combining the calibrated 14 C ages obtained on charcoals and organic matter and U-Th ages from 14 C analyses made on the carbonate, has allowed the calculation of the dead carbon proportion (dcp) of the carbonate deposits. The latter, used with the initial atmospheric 14 C activities reconstructed with the new IntCal09 calibration data, allows high-resolution age estimation of the gour calcite samples and their growth rates. The carbonate deposit grew between 9530 and 6635 yr cal BP (for dcp = 10.7 ± 1.8%; 2 s) or between 8518 and 5489 yr cal BP (for dcp = 20.5 ± 1.9%; 2 s). This coincides with humid periods that can be related to the Atlantic period in Europe and to Sapropel 1 in the eastern Mediterranean Sea. However, geomorphological changes at the cave entrance might also have played a role in the gour development. In the 1940's, when humans entered the cave for the first time since its prehistoric occupation, the calcite gours had already been inactive for several thousand years. (authors)

  7. Ridge-like lava tube systems in southeast Tharsis, Mars

    Science.gov (United States)

    Zhao, Jiannan; Huang, Jun; Kraft, Michael D.; Xiao, Long; Jiang, Yun

    2017-10-01

    Lava tubes are widely distributed in volcanic fields on a planetary surface and they are important means of lava transportation. We have identified 38 sinuous ridges with a lava-tube origin in southeast Tharsis. The lengths vary between 14 and 740 km, and most of them occur in areas with slopes rate, low lava viscosity, and sustained magma supply during a long period. Besides, lava flow inflation is also important in the formation of the ridge-like lava tubes and some associated features. These lava tubes provide efficient lateral pathways for magma transportation over the relatively low topographic slopes in southeast Tharsis, and they are important for the formation of long lava flows in this region. The findings of this study provide an alternative formation mechanism for sinuous ridges on the martian surface.

  8. Solar activity influence on air temperature regimes in caves

    Science.gov (United States)

    Stoeva, Penka; Mikhalev, Alexander; Stoev, Alexey

    Cave atmospheres are generally included in the processes that happen in the external atmosphere as circulation of the cave air is connected with the most general circulation of the air in the earth’s atmosphere. Such isolated volumes as the air of caves are also influenced by the variations of solar activity. We discuss cave air temperature response to climate and solar and geomagnetic activity for four show caves in Bulgaria studied for a period of 46 years (1968 - 2013). Everyday noon measurements in Ledenika, Saeva dupka, Snezhanka and Uhlovitsa cave have been used. Temperatures of the air in the zone of constant temperatures (ZCT) are compared with surface temperatures recorded at meteorological stations situated near about the caves - in the towns of Vratsa, Lovech, Peshtera and Smolyan, respectively. For comparison, The Hansen cave, Middle cave and Timpanogos cave from the Timpanogos Cave National Monument, Utah, USA situated nearly at the same latitude have also been examined. Our study shows that the correlation between cave air temperature time series and sunspot number is better than that between the cave air temperature and Apmax indices; that t°ZCT is rather connected with the first peak in geomagnetic activity, which is associated with transient solar activity (CMEs) than with the second one, which is higher and connected with the recurrent high speed streams from coronal holes. Air temperatures of all examined show caves, except the Ledenika cave, which is ice cave show decreasing trends. On the contrary, measurements at the meteorological stations show increasing trends in the surface air temperatures. The trend is decreasing for the Timpanogos cave system, USA. The conclusion is that surface temperature trends depend on the climatic zone, in which the cave is situated, and there is no apparent relation between temperatures inside and outside the caves. We consider possible mechanism of solar cosmic rays influence on the air temperatures in caves

  9. Diverting lava flows in the lab

    Science.gov (United States)

    Dietterich, Hannah; Cashman, Katharine V.; Rust, Alison C.; Lev, Einat

    2015-01-01

    Recent volcanic eruptions in Hawai'i, Iceland and Cape Verde highlight the challenges of mitigating hazards when lava flows threaten infrastructure. Diversion barriers are the most common form of intervention, but historical attempts to divert lava flows have met with mixed success and there has been little systematic analysis of optimal barrier design. We examine the interaction of viscous flows of syrup and molten basalt with barriers in the laboratory. We find that flows thicken immediately upslope of an obstacle, forming a localized bow wave that can overtop barriers. Larger bow waves are generated by faster flows and by obstacles oriented at a high angle to the flow direction. The geometry of barriers also influences flow behaviour. Barriers designed to split or dam flows will slow flow advance, but cause the flow to widen, whereas oblique barriers can effectively divert flows, but may also accelerate flow advance. We argue that to be successful, mitigation of lava-flow hazards must incorporate the dynamics of lava flow–obstacle interactions into barrier design. The same generalizations apply to the effect of natural topographic features on flow geometry and advance rates.

  10. Possible lava tube system in a hummocky lava flow at Daund ...

    Indian Academy of Sciences (India)

    R. Narasimhan (Krishtel eMaging) 1461 1996 Oct 15 13:05:22

    The presence of a branching and meandering lava tube system in the Daund flow, which represents the ..... is entirely related to the process of differential ero- sion and exhumation. Thus ... illuminating and thought provoking. References.

  11. Cave Tourism: The Potential of Asar Cave as a Natural Tourism Asset at Lenggong Valley, Perak

    Directory of Open Access Journals (Sweden)

    Rindam Main

    2014-01-01

    Full Text Available The Lenggong Valley, from a standpoint of natural tourism research, presents strengths, weaknesses, opportunities and challenges that can be utilized to help increase the opportunities for the local community to increase their standard of living. Asar Cave comprises one of the caves that are found in Lenggong. A series of external studies have been done on Asar Cave in order to measure its potential for natural tourism in Lenggong. The objective of this study is to discuss caves as a natural resource that has great potential in the growth of the economy of the residents of the Lenggong Valley. Marketing caves as a source of nature tourism helps the government’s achievements in National Key Result Areas, apart from being a form of environmental control as well as helping to increase awareness about environmental education, specifically those associated with caves. The research results find that SWOT analysis presents huge potential for caves to become a source of nature tourism development in Lenggong. Great potential can also be seen from a standpoint of increasing the standard of living of its residents through their involvement in the tourism sector based on local natural assets.

  12. Črna Jama as a cold air trap cave within Postojna Cave, Slovenia

    Science.gov (United States)

    Šebela, Stanka; Turk, Janez

    2017-10-01

    Črna Jama is the coldest section of cave within the Postojna Cave System. Mean annual air temperatures at the Črna Jama 2 site are 5.6 °C (2015) and 5.7 °C (2016), and at the Črna Jama 3 site 7.1 °C (2015) and 7.2 (2016), whereas the mean external air temperature was 10.3 °C (2015) and 10.0 °C (2016). In Lepe Jame, the passage most heavily visited by tourists, the mean cave-air temperature is 10.7 °C (2014-2017). Črna Jama exhibits winter and summer temperature regimes. During warm periods (Tcave Tout), ventilation takes place and dense, cold, outside air sinks into Črna Jama because of the favourable cave entrance morphology. Recent Črna Jama air temperature data (2014-2017) indicate a < 0.5 °C higher temperature than that recorded in historical data since 1933. Črna Jama is the most appropriate place within the Postojna Cave System to study long-term climatic changes. There are hardly any tourist visits to the cave, and human impacts on the cave climate are essentially reduced.

  13. Large-scale phylogenomic analysis resolves a backbone phylogeny in ferns

    Science.gov (United States)

    Shen, Hui; Jin, Dongmei; Shu, Jiang-Ping; Zhou, Xi-Le; Lei, Ming; Wei, Ran; Shang, Hui; Wei, Hong-Jin; Zhang, Rui; Liu, Li; Gu, Yu-Feng; Zhang, Xian-Chun; Yan, Yue-Hong

    2018-01-01

    Abstract Background Ferns, originated about 360 million years ago, are the sister group of seed plants. Despite the remarkable progress in our understanding of fern phylogeny, with conflicting molecular evidence and different morphological interpretations, relationships among major fern lineages remain controversial. Results With the aim to obtain a robust fern phylogeny, we carried out a large-scale phylogenomic analysis using high-quality transcriptome sequencing data, which covered 69 fern species from 38 families and 11 orders. Both coalescent-based and concatenation-based methods were applied to both nucleotide and amino acid sequences in species tree estimation. The resulting topologies are largely congruent with each other, except for the placement of Angiopteris fokiensis, Cheiropleuria bicuspis, Diplaziopsis brunoniana, Matteuccia struthiopteris, Elaphoglossum mcclurei, and Tectaria subpedata. Conclusions Our result confirmed that Equisetales is sister to the rest of ferns, and Dennstaedtiaceae is sister to eupolypods. Moreover, our result strongly supported some relationships different from the current view of fern phylogeny, including that Marattiaceae may be sister to the monophyletic clade of Psilotaceae and Ophioglossaceae; that Gleicheniaceae and Hymenophyllaceae form a monophyletic clade sister to Dipteridaceae; and that Aspleniaceae is sister to the rest of the groups in eupolypods II. These results were interpreted with morphological traits, especially sporangia characters, and a new evolutionary route of sporangial annulus in ferns was suggested. This backbone phylogeny in ferns sets a foundation for further studies in biology and evolution in ferns, and therefore in plants. PMID:29186447

  14. Large-scale phylogenomic analysis resolves a backbone phylogeny in ferns.

    Science.gov (United States)

    Shen, Hui; Jin, Dongmei; Shu, Jiang-Ping; Zhou, Xi-Le; Lei, Ming; Wei, Ran; Shang, Hui; Wei, Hong-Jin; Zhang, Rui; Liu, Li; Gu, Yu-Feng; Zhang, Xian-Chun; Yan, Yue-Hong

    2018-02-01

    Ferns, originated about 360 million years ago, are the sister group of seed plants. Despite the remarkable progress in our understanding of fern phylogeny, with conflicting molecular evidence and different morphological interpretations, relationships among major fern lineages remain controversial. With the aim to obtain a robust fern phylogeny, we carried out a large-scale phylogenomic analysis using high-quality transcriptome sequencing data, which covered 69 fern species from 38 families and 11 orders. Both coalescent-based and concatenation-based methods were applied to both nucleotide and amino acid sequences in species tree estimation. The resulting topologies are largely congruent with each other, except for the placement of Angiopteris fokiensis, Cheiropleuria bicuspis, Diplaziopsis brunoniana, Matteuccia struthiopteris, Elaphoglossum mcclurei, and Tectaria subpedata. Our result confirmed that Equisetales is sister to the rest of ferns, and Dennstaedtiaceae is sister to eupolypods. Moreover, our result strongly supported some relationships different from the current view of fern phylogeny, including that Marattiaceae may be sister to the monophyletic clade of Psilotaceae and Ophioglossaceae; that Gleicheniaceae and Hymenophyllaceae form a monophyletic clade sister to Dipteridaceae; and that Aspleniaceae is sister to the rest of the groups in eupolypods II. These results were interpreted with morphological traits, especially sporangia characters, and a new evolutionary route of sporangial annulus in ferns was suggested. This backbone phylogeny in ferns sets a foundation for further studies in biology and evolution in ferns, and therefore in plants. © The Authors 2017. Published by Oxford University Press.

  15. Cave dwellings in the Mediterranean basin

    Directory of Open Access Journals (Sweden)

    Viedma Urdiales Eugenia María

    2011-01-01

    Full Text Available The construction and use of subterranean caves for different functions has been relatively commonplace throughout history in different regions around the world, especially in the Mediterranean basin. Some of them are still standing at the beginning of the 21st century, and are a good example of adaptation to the geographic environment, and a part of the historical heritage. Following a short overview of the different Mediterranean countries, this work pays special attention to the present use of caves as dwelling spaces in Italy, and particularly in Spain where the caves are currently in an interesting process of renovation to meet the needs of the present population. This process is helping to boost the local economy, and it is funded by both private and public sources in several towns in Andalusia (Spain.

  16. Radon in New Zealand tourist caves

    International Nuclear Information System (INIS)

    Lyons, R.G.; Solomon, S.B.; Langroo, R.

    1998-01-01

    Seasonal average radon concentrations in 112 sites in 22 NZ tourist caves have been measured using track etch detectors over an annual cycle. Values ranged from -3 to nearly 10,000 Bq m -3 . Seasonal variation was also very marked with factors of over 50 for the same site in different seasons being recorded. Thirty six percent of the sites exceeded the ICRP guideline of 1000 Bq m -3 at which action to reduce exposure is recommended. Caves are fragile ecological and chemical systems, therefore, may be seriously adversely affected by standard techniques to reduce radon concentrations. They are also complex physical systems, and an understanding of the principles and parameters governing cave aerodynamics is essential when considering options to reduce exposure. This paper discusses possible causes for variations in radon concentrations observed in this study and the implications for viable actions

  17. Diurnal Variation of Radon Concentration in the Postojna Cave

    International Nuclear Information System (INIS)

    Gregoric, A.; Vaupotic, J.

    2011-01-01

    Postojna Cave, with 20 km of galleries, is the longest known cave system and also the largest of about 20 show caves in Slovenia and one of the most visited show caves in the world. It is well known that high concentrations of radon are common in karstic caves, although quantities of uranium (238U) in limestone are rather low. The reason for this is low natural ventilation of the underground cavities. Tectonic faults constitute an additional source of radon. Variations of radon concentration in cave air arise from a balance of the emission from cave surfaces and drip waters, decay in cave air, and exchange with the outside atmosphere. Because of its elevated radon concentrations, Postojna Cave has been under permanent radon survey since 1995. The influence of meteorological conditions on the radon levels and their temporal variations depends mostly on the shape of the cave, and the number and directions of cracks, corridors and fissures connecting the cave rooms with the outside atmosphere. The driving force for air movement in horizontal caves, and thus the inflow of fresh air and release of the cave air to the atmosphere, is the temperature difference between the cave air and outdoors, which causes seasonal pattern of radon concentration in the cave with high levels in summer and low in winter. However, on a daily scale different behaviour of radon can be observed at different locations in the cave. In this paper diurnal variation of radon concentration at two locations is presented and discussed. Postojna Cave is a horizontal cave with a stable yearly temperature around 10 degrees of @C. Continuous measurements of radon concentration were carried out from 2005 to 2010 at two locations along the guided tourist trail. Radon concentration was measured with Radim 5 WP monitors (SMM Company, Prague, Czech Republic) with sampling frequency once an hour. The evaluation of five-year radon monitoring at two sites in the Postojna Cave reveals significant diurnal and

  18. Dating Petroglyphs from Fugoppe Cave, Japan

    Directory of Open Access Journals (Sweden)

    Masaru Ogawa

    2014-01-01

    Full Text Available For over 20 years, I have tried to establish a relative date for petroglyphs in Fugoppe Cave, Japan. Unsuspected amidst debris accumulating from about 1300 years ago, the petroglyphs were rediscovered accidentally in 1950. From an analysis of petroglyphs on fallen rocks scattered randomly on the site floor, I argue that the artworks date from ca.1900 years ago. The cave itself, formed by wave action, saw its main occupation by pottery-making people from 1700–1500 years ago; although the petroglyphs on the rock walls predated their occupation, it seems unlikely that the occupants attached any meaning to them.

  19. Alvaro Fernández Pérez

    Directory of Open Access Journals (Sweden)

    Díaz Piedrahita Santiago

    1995-11-01

    Full Text Available Fue el Dr. Fernández miembro destacado de la comunidad botánica latinoamericana formada hacia la mitad del siglo, y en Colombia desempeñó un papel importante como miembro de planta del Herbario Nacional Colombiano y del Instituto de Ciencias Naturales de la Facultad de Ciencias de la Universidad Nacional de Colombia. Nació el Dr. Fernández el 11 de febrero de 1920 en la procera ciudad de Popayán, donde tras adelantar los estudios primarios y secundarios obtuvo en 1940 el título de bachiller en el Liceo de la Universidad del Cauca; cumplida esta etapa se desplazó a Bogotá, ingresando a la Universidad Nacional donde completó el plan de estudios de la carrera de Farmacia e inició el curso del doctorado que se ofrecía para la época.

  20. PIXE study on arsenic accumulation by a fern. Pteris vittata

    International Nuclear Information System (INIS)

    Yamazaki, H.; Ishii, K.; Matsuyama, S.

    2010-01-01

    Pteris vittata is a fern reported to be an arsenic hyper-accumulator. To develop the practical application of the fern to a phytoremediation technique, it is necessary to explicate the effective accumulation mechanism. In this study, the arsenic distribution and the elemental correlation in the cellular level were examined in the fronds supplied with arsenate and arsenite separately via xylem vessel using an in-air micro-PIXE system at Tohoku University. The difference in transportation rate between arsenate and arsenite as well as the translocation of elements necessary for plant metabolism was revealed in different tissues of the fronds accumulating arsenic in high concentration. Hence, the in-air micro-PIXE analysis is an effective measure for undertaking phytoremediation research of hyper-accumulator plants. (author)

  1. Environmental Fate and Analysis of Ptaquiloside from the Bracken Fern

    DEFF Research Database (Denmark)

    Clauson-Kaas, Frederik

    The naturally occurring phytotoxin ptaquiloside (PTA) has long been known to be both acute toxic and carcinogenic. Contents of more than 1% ptaquiloside on dry weight has been detected in bracken (Pteridium spp.), a fern distributed across the globe in often dense populations. This work focused...... on the fate of PTA in the soil-water system, from where it may leach to drinking water sources. PTA was detected in concentrations up to 2.2 µg/L in natural waters receiving drainage from bracken populations, and was found in both surface and groundwater. It was shown that ptaquiloside leached off bracken...... fronds (the leaves of ferns) in concentrations up to 169 µ/L during rainfall events. Rainfall further determined the concentration in a stream that drained a bracken-covered catchment, suggesting that this is a potent driver of ptaquiloside exposure in the environment. In both pure and natural waters, p...

  2. Investigation of clay sediments and bedrock morphology in caves with seismic traveltime tomography: an application at Alepotrypa Cave (Diros, Greece

    Directory of Open Access Journals (Sweden)

    Lazaros Polymenakos

    2017-01-01

    Full Text Available The deposition of unconsolidated clay sediments in caves, in relation to the buried morphology of the karstic conduit, are important parameters for the study of cave evolution. We introduce the application of an active seismic imaging technique to investigate the clay deposits and bedrock morphology in caves. Seismic traveltime tomography, applied for the first time in cave studies, can assist with the interpretation of cave geomorphology. Utilizing the P-wave velocity contrast between the clay sediments and the surrounding rock mass, we map the buried rock surface and significant sediment interfaces and provide an estimate of the sediment thickness and volume. Our study focuses on the Alepotrypa Cave located in Diros (Peloponnese, Greece, revealing important information for the evolution of the cave. The proposed technique could be applied in caves with significant clay deposits, in order to constrain the clay volume and reconstruct the buried floor shape of the cave. The technique exploits fully the ground morphology and access points in a cave, so it is suitable for a detailed three-dimensional exploration of cave deposits and the underlying cave morphology.

  3. The importance of ants in cave ecology, with new records and behavioral observations of ants in Arizona caves

    Directory of Open Access Journals (Sweden)

    Robert B. Pape

    2016-09-01

    Full Text Available The importance of ants as elements in cave ecology has been mostly unrecognized. A global list of ant species recorded from caves, compiled from a review of existing literature, is presented. This paper also reviews what is currently known about ants occurring in Arizona (USA caves. The diversity and distribution represented in these records suggests ants are relatively common cave visitors (trogloxenes. A general utilization of caves by ants within both temperate and tropical latitudes may be inferred from this combined evidence. Observations of ant behavior in Arizona caves demonstrate a low level and sporadic, but persistent, use of these habitats and their contained resources by individual ant colonies. Documentation of Neivamyrmex sp. preying on cave-inhabiting arthropods is reported here for the first time. Observations of hypogeic army ants in caves suggests they may not penetrate to great vertical depth in search of prey, but can be persistent occupants in relatively shallow, horizontal sections of caves where they may prey on endemic cave animals. First cave records for ten ant species are reported from Arizona caves. These include two species of Neivamyrmex (N. nigrescens Cresson and Neivamyrmex sp.; Formicidae: Dorylinae, four myrmicines (Pheidole portalensis Wilson, Pheidole cf. porcula Wheeler, Solenopsis aurea Wheeler and Stenamma sp. Westwood, one dolichoderine (Forelius keiferi Wheeler and three formicines (Lasius arizonicus Wheeler, L. sitiens Wilson, and Camponotus sp. Mayr.

  4. Somatic embryogenesis in ferns: a new experimental system.

    Science.gov (United States)

    Mikuła, Anna; Pożoga, Mariusz; Tomiczak, Karolina; Rybczyński, Jan J

    2015-05-01

    Somatic embryogenesis has never been reported in ferns. The study showed that it is much easier to evoke the acquisition and expression of embryogenic competence in ferns than in spermatophytes. We discovered that the tree fern Cyathea delgadii offers an effective model for the reproducible and rapid formation of somatic embryos on hormone-free medium. Our study provides cyto-morphological evidence for the single cell origin and development of somatic embryos. Somatic embryogenesis (SE) in both primary and secondary explants was induced on half-strength micro- and macro-nutrients Murashige and Skoog medium without the application of exogenous plant growth regulators, in darkness. The early stage of SE was characterized by sequential perpendicular cell divisions of an individual epidermal cell of etiolated stipe explant. These resulted in the formation of a linear pro-embryo. Later their development resembled that of the zygotic embryo. We defined three morphogenetic stages of fern somatic embryo development: linear, early and late embryonic leaf stage. The transition from somatic embryo to juvenile sporophyte was quick and proceeded without interruption caused by dormancy. Following 9 weeks of culture the efficiency of somatic embryogenesis reached 12-13 embryos per responding explant. Spontaneous formation of somatic embryos and callus production, which improved the effectiveness of the process sevenfold in 10-month-long culture, occurred without subculturing. The tendency for C. delgadii to propagate by SE in vitro makes this species an excellent model for studies relating to asexual embryogenesis and the endogenous hormonal regulation of that process and opens new avenues of experimentation.

  5. The scaly tree ferns (Cyatheaceae-Polypodiopsida) of Brazil

    OpenAIRE

    Weigand, Anna; Lehnert, Marcus

    2016-01-01

    ABSTRACT A synopsis of all scaly tree fern species (Cyatheaceae) occurring in Brazil is presented. We recognize 51 species in three genera [Sphaeropteris one species, Alsophila four species (one subspecies, two varieties), and Cyathea 45 species (one variety) ] with 17 taxa being endemic to Brazil. One hybrid endemic to Brazil is recognized. Further included are five species that have not yet been recorded in Brazil, but are expected here because they are found in adjacent countries and occur...

  6. Sorption equilibrium of mercury onto ground-up tree fern.

    Science.gov (United States)

    Ho, Yuh-Shan; Wang, Chung-Chi

    2008-08-15

    The sorption behavior of mercury at different temperatures onto ground-up tree fern was investigated. The experimental results were fitted to two two-parameter isotherms, the Freundlich and Langmuir isotherms, as well as to two three-parameter isotherms, the Redlich-Peterson and Sips isotherms to obtain the characteristic parameters of each model. A comparison of best-fitting was performed using the coefficient of determination and Chi-square test. Both the Langmuir and Redlich-Peterson isotherms were found to well represent the measured sorption data. According to the evaluation using the Langmuir equation, the saturated monolayer sorption capacity of mercury ions onto ground-up tree fern was 26.5 mg/g at 298 K. It was noted that an increase in temperature resulted in a higher mercury ion loading per unit weight of the tree fern. In addition, various thermodynamic parameters, such as DeltaG degrees, DeltaH degrees, and DeltaS degrees, were calculated and compared with the sorption of mercury by other sorbents.

  7. Ferns: the missing link in shoot evolution and development

    Directory of Open Access Journals (Sweden)

    Andrew Robert George Plackett

    2015-11-01

    Full Text Available Shoot development in land plants is a remarkably complex process that gives rise to an extreme diversity of forms. Our current understanding of shoot developmental mechanisms comes almost entirely from studies of angiosperms (flowering plants, the most recently diverged plant lineage. Shoot development in angiosperms is based around a layered multicellular apical meristem that produces lateral organs and/or secondary meristems from populations of founder cells at its periphery. In contrast, non-seed plant shoots develop from either single apical initials or from a small population of morphologically distinct apical cells. Although developmental and molecular information is becoming available for non-flowering plants, such as the model moss Physcomitrella patens, making valid comparisons between highly divergent lineages is extremely challenging. As sister group to the seed plants, the monilophytes (ferns and relatives represent an excellent phylogenetic midpoint of comparison for unlocking the evolution of shoot developmental mechanisms, and recent technical advances have finally made transgenic analysis possible in the emerging model fern Ceratopteris richardii. This review compares and contrasts our current understanding of shoot development in different land plant lineages with the aim of highlighting the potential role that the fern C. richardii could play in shedding light on the evolution of underlying genetic regulatory mechanisms.

  8. Sorption equilibrium of mercury onto ground-up tree fern

    Energy Technology Data Exchange (ETDEWEB)

    Ho, Y.-S. [Department of Chemical Engineering, I-Shou University, No. 1, Section 1, Hsueh-Cheng Road, Ta-Hsu Hsiang, Kaohsiung County 840, Taiwan (China)], E-mail: ysho@isu.edu.tw; Wang, C.-C. [Department of Chemical Engineering, I-Shou University, No. 1, Section 1, Hsueh-Cheng Road, Ta-Hsu Hsiang, Kaohsiung County 840, Taiwan (China)

    2008-08-15

    The sorption behavior of mercury at different temperatures onto ground-up tree fern was investigated. The experimental results were fitted to two two-parameter isotherms, the Freundlich and Langmuir isotherms, as well as to two three-parameter isotherms, the Redlich-Peterson and Sips isotherms to obtain the characteristic parameters of each model. A comparison of best-fitting was performed using the coefficient of determination and Chi-square test. Both the Langmuir and Redlich-Peterson isotherms were found to well represent the measured sorption data. According to the evaluation using the Langmuir equation, the saturated monolayer sorption capacity of mercury ions onto ground-up tree fern was 26.5 mg/g at 298 K. It was noted that an increase in temperature resulted in a higher mercury ion loading per unit weight of the tree fern. In addition, various thermodynamic parameters, such as {delta}G{sup o}, {delta}H{sup o}, and {delta}S{sup o}, were calculated and compared with the sorption of mercury by other sorbents.

  9. Sorption equilibrium of mercury onto ground-up tree fern

    International Nuclear Information System (INIS)

    Ho, Y.-S.; Wang, C.-C.

    2008-01-01

    The sorption behavior of mercury at different temperatures onto ground-up tree fern was investigated. The experimental results were fitted to two two-parameter isotherms, the Freundlich and Langmuir isotherms, as well as to two three-parameter isotherms, the Redlich-Peterson and Sips isotherms to obtain the characteristic parameters of each model. A comparison of best-fitting was performed using the coefficient of determination and Chi-square test. Both the Langmuir and Redlich-Peterson isotherms were found to well represent the measured sorption data. According to the evaluation using the Langmuir equation, the saturated monolayer sorption capacity of mercury ions onto ground-up tree fern was 26.5 mg/g at 298 K. It was noted that an increase in temperature resulted in a higher mercury ion loading per unit weight of the tree fern. In addition, various thermodynamic parameters, such as ΔG o , ΔH o , and ΔS o , were calculated and compared with the sorption of mercury by other sorbents

  10. Morphology, Physiology, and Anatomy of Penny Fern (Drymoglossum phyloselloidesand Its Effect on Cocoa

    Directory of Open Access Journals (Sweden)

    Fitria Yuliasmara

    2013-08-01

    Full Text Available This study aims to determine the anatomy, physiology and morphology of penny fern (Drimoglosum phylloseloides and its effect on cocoa. Morphological observation of penny fern used microscope to observe the roots, stems, leaves and spores. Physiology of penny fern was observed based on number of stomata and stomatal conductance using stomata printing method, while the amount of chlorophyll based on spectrophotometric method and rate of transpiration used cobalt chloride paper. Penny fern anatomy on cross-sectional and longitudinal in roots, stems and leaves. Penny fern growth was observed based the length of tendrils once a week during rainy and dry season. While the effect of penny fern invasion was observed based on variable leaf area with gravimetric method, the cross-section of attacked cacao branch using microtom and microscope and chlorophyll content by chlorophyll meter. Results showed that penny fern is a epiphytic weed which was crassulaceae acid metabolism plants that have the ability to absorb carbon dioxide at night and carry out photosynthesis during the day with closed stomata. Penny ferns reproduce using spores. The growth rate of penny fern 2.18 cm/week during the dry season and while in rainy season 3.89 cm/week. Penny fern leaf contains 0.0212 mg/g chlorophyll. Penny fern stomata density was 18.33/mm 2 with a width of opening stomata at night 26.3 µm which caused a veryslow rate of transpiration of 0.69 mm 2 /seconds. The existence penny fern on cocoa decreased leaf area and chlorophyll content decreased crop productivity which was indicated by decreasing in number of flowers, number of small, medium fruit, and large pods. However it had no effect on the number of leaves on one side flush cocoa. Key words: Drimoglosum phylloseloides, weeds, decrease productivity, Theobroma cacao

  11. Colonisation of epiphytic ferns by skinks and geckos in the high canopy of a Bornean rainforest

    OpenAIRE

    Donald, J.; Clegg, J.; Ellwood, M. D. F.

    2017-01-01

    Nest site availability limits the fitness and survival of skinks and geckos, particularly in the canopy of tall tropical rainforests. We document the systematic colonisation and nest use of epiphytic bird’s nest ferns (Asplenium spp) by the gecko Hemiphyllodactylus typus and the skink Lipinia cf. vittigera. As part of a controlled experiment we placed 32 ferns of similar sizes in the high canopy of a lowland dipterocarp rainforest in Sabah, Malaysian Borneo. Half of these ferns, sampled after...

  12. Cryogenic Minerals in Caves of the Vizhay River (Northern Urals

    Directory of Open Access Journals (Sweden)

    E. P. Bazarova

    2018-03-01

    Full Text Available New information on the cryogenic mineral formations at the two Vizhay River caves (Northern Urals is given.   Calcite with the insignificant gypsum admixture predominates in the cryogenic material composition from both caves. In addition, the metastable phases of calcite, such as monohydrocalcite and ikaite were found. In the Saksofon Cave, calcite forms both spherulites and complanate grains. In Lednikovaya Cave, the major part of cryomaterial is presented by spherulites, which may suggests the significant supersaturation of solution. In Lednikovaya Сave, the distinct concentric structure with the growth zones denotes the cryogenic material formation in a thin water film under the partial thawing of upper part of long-term ice mound in summer. In Saksofon Cave the growth zones in crystals are poorly developed that probably caused by the seasonal glaciation in the cave and cryogenic minerals are younger than those in the Lednikovaya Cave.

  13. Instrumenting caves to collect hydrologic and geochemical data: case study from James Cave, Virginia

    Science.gov (United States)

    Schreiber, Madeline E.; Schwartz, Benjamin F.; Orndorff, William; Doctor, Daniel H.; Eagle, Sarah D.; Gerst, Jonathan D.

    2015-01-01

    Karst aquifers are productive groundwater systems, supplying approximately 25 % of the world’s drinking water. Sustainable use of this critical water supply requires information about rates of recharge to karst aquifers. The overall goal of this project is to collect long-term, high-resolution hydrologic and geochemical datasets at James Cave, Virginia, to evaluate the quantity and quality of recharge to the karst system. To achieve this goal, the cave has been instrumented for continuous (10-min interval) measurement of the (1) temperature and rate of precipitation; (2) temperature, specific conductance, and rate of epikarst dripwater; (3) temperature of the cave air; and (4) temperature, conductivity, and discharge of the cave stream. Instrumentation has also been installed to collect both composite and grab samples of precipitation, soil water, the cave stream, and dripwater for geochemical analysis. This chapter provides detailed information about the instrumentation, data processing, and data management; shows examples of collected datasets; and discusses recommendations for other researchers interested in hydrologic and geochemical monitoring of cave systems. Results from the research, briefly described here and discussed in more detail in other publications, document a strong seasonality of the start of the recharge season, the extent of the recharge season, and the geochemistry of recharge.

  14. Helium Isotopes and Noble Gas Abundances of Cave Dripping Water in Three Caves in East Asia

    Science.gov (United States)

    Chen, A. T.; Shen, C. C.; Tan, M.; Li, T.; Uemura, R.; Asami, R.

    2015-12-01

    Paleo-temperature recorded in nature archives is a critical parameter to understand climate change in the past. With advantages of unique inert chemical characteristics and sensitive solubilities with temperature, dissolved noble gases in speleothem inclusion water were recently proposed to retrieve terrestrial temperature history. In order to accurately apply this newly-developed speleothem noble gas temperature (NGT) as a reliable proxy, a fundamental issue about behaviors of noble gases in the karst should be first clarified. In this study, we measured noble gas contents in air and dripping water to evaluate any ratio deviation between noble gases. Cave dripping water samples was collected from three selected caves, Shihua Cave in northern China, Furong Cave in southwestern, and Gyukusen Cave in an island located in the western Pacific. For these caves are characterized by a thorough mixing and long-term storage of waters in a karst aquifer by the absence of seasonal oxygen isotope shifts. Ratios of dripping water noble gases are statistically insignificant from air data. Helium isotopic ratios in the dripping water samples match air value. The results indicate that elemental and isotopic signatures of noble gases from air can be frankly preserved in the epikarst and support the fidelity of NGT techniques.

  15. Is radon emission in caves causing deletions in satellite DNA sequences of cave-dwelling crickets?

    Directory of Open Access Journals (Sweden)

    Giuliana Allegrucci

    Full Text Available The most stable isotope of radon, 222Rn, represents the major source of natural radioactivity in confined environments such as mines, caves and houses. In this study, we explored the possible radon-related effects on the genome of Dolichopoda cave crickets (Orthoptera, Rhaphidophoridae sampled in caves with different concentrations of radon. We analyzed specimens from ten populations belonging to two genetically closely related species, D. geniculata and D. laetitiae, and explored the possible association between the radioactivity dose and the level of genetic polymorphism in a specific family of satellite DNA (pDo500 satDNA. Radon concentration in the analyzed caves ranged from 221 to 26,000 Bq/m3. Specimens coming from caves with the highest radon concentration showed also the highest variability estimates in both species, and the increased sequence heterogeneity at pDo500 satDNA level can be explained as an effect of the mutation pressure induced by radon in cave. We discovered a specific category of nuclear DNA, the highly repetitive satellite DNA, where the effects of the exposure at high levels of radon-related ionizing radiation are detectable, suggesting that the satDNA sequences might be a valuable tool to disclose harmful effects also in other organisms exposed to high levels of radon concentration.

  16. Food sources of selected terrestrial cave arthropods

    Czech Academy of Sciences Publication Activity Database

    Smrž, J.; Kováč, L.; Mikeš, J.; Šustr, Vladimír; Lukešová, Alena; Tajovský, Karel; Nováková, Alena; Režňáková, P.

    2015-01-01

    Roč. 16, č. 1 (2015), s. 37-46 ISSN 1768-1448 Grant - others:Vega(SK) 1/0139/09 Institutional research plan: CEZ:AV0Z60660521 Institutional support: RVO:60077344 Keywords : Acari * caves * Collembola * Diplopoda * feeding habits * Isopoda Subject RIV: EG - Zoology

  17. Measurements of Aerosol Characteristics in Skocjan Caves

    International Nuclear Information System (INIS)

    Jovanovic, P.

    2013-01-01

    Measurements of radon concentration and radon progeny concentration (attached and unattached) have been performed in Skocjan caves. In the same time also aerosol concentration (PM 10 ), aerosol size distribution with ten stage Hauke impactor and Scanning Mobility Particle Sizer - SMPS have been performed. The idea was to find impact of outer air and visitors to the aerosol characteristics of cave air. Measurements with impactor have been implemented in summer and winter period, with SMPS only in summer period. Radon concentrations ranged in winter period in region from 500 to 1000 Bq/m 3 , equilibrium factor was about 55 %. In summer period radon concentration increased up to 10 kBq/m 3 , equilibrium factor was about 45 %, and unattached fraction went up to 20 %. Measurements of aerosol size distribution show lower aerosol sizes in winter season (around 1 μm) and bigger aerosol sizes in summer season (around 3 - 6 μm). We could not find good correlation between unattached fraction and aerosol size distribution. Also we could not find clear impact of visitors to the air characteristics in cave. Probably our measuring location was too close to the entrance and the impact of outer air was too high. We will repeat measurements deeper in cave to find better results.(author)

  18. LAVA: Large scale Automated Vulnerability Addition

    Science.gov (United States)

    2016-05-23

    LAVA: Large-scale Automated Vulnerability Addition Brendan Dolan -Gavitt∗, Patrick Hulin†, Tim Leek†, Fredrich Ulrich†, Ryan Whelan† (Authors listed...released, and thus rapidly become stale. We can expect tools to have been trained to detect bugs that have been released. Given the commercial price tag...low TCN) and dead (low liveness) program data is a powerful one for vulnera- bility injection. The DUAs it identifies are internal program quantities

  19. Modeling steam pressure under martian lava flows

    Science.gov (United States)

    Dundas, Colin M.; Keszthelyi, Laszlo P.

    2013-01-01

    Rootless cones on Mars are a valuable indicator of past interactions between lava and water. However, the details of the lava–water interactions are not fully understood, limiting the ability to use these features to infer new information about past water on Mars. We have developed a model for the pressurization of a dry layer of porous regolith by melting and boiling ground ice in the shallow subsurface. This model builds on previous models of lava cooling and melting of subsurface ice. We find that for reasonable regolith properties and ice depths of decimeters, explosive pressures can be reached. However, the energy stored within such lags is insufficient to excavate thick flows unless they draw steam from a broader region than the local eruption site. These results indicate that lag pressurization can drive rootless cone formation under favorable circumstances, but in other instances molten fuel–coolant interactions are probably required. We use the model results to consider a range of scenarios for rootless cone formation in Athabasca Valles. Pressure buildup by melting and boiling ice under a desiccated lag is possible in some locations, consistent with the expected distribution of ice implanted from atmospheric water vapor. However, it is uncertain whether such ice has existed in the vicinity of Athabasca Valles in recent history. Plausible alternative sources include surface snow or an aqueous flood shortly before the emplacement of the lava flow.

  20. Simulated oxygen isotopes in cave drip water and speleothem calcite in European caves

    Directory of Open Access Journals (Sweden)

    A. Wackerbarth

    2012-11-01

    Full Text Available Interpreting stable oxygen isotope (δ18O records from stalagmites is still one of the complex tasks in speleothem research. Here, we present a novel model-based approach, where we force a model describing the processes and modifications of δ18O from rain water to speleothem calcite (Oxygen isotope Drip water and Stalagmite Model – ODSM with the results of a state-of-the-art atmospheric general circulation model enhanced by explicit isotope diagnostics (ECHAM5-wiso. The approach is neither climate nor cave-specific and allows an integrated assessment of the influence of different varying climate variables, e.g. temperature and precipitation amount, on the isotopic composition of drip water and speleothem calcite.

    First, we apply and evaluate this new approach under present-day climate conditions using observational data from seven caves from different geographical regions in Europe. Each of these caves provides measured δ18O values of drip water and speleothem calcite to which we compare our simulated isotope values. For six of the seven caves modeled δ18O values of drip water and speleothem calcite are in good agreement with observed values. The mismatch of the remaining caves might be caused by the complexity of the cave system, beyond the parameterizations included in our cave model.

    We then examine the response of the cave system to mid-Holocene (6000 yr before present, 6 ka climate conditions by forcing the ODSM with ECHAM5-wiso results from 6 ka simulations. For a set of twelve European caves, we compare the modeled mid-Holocene-to-modern difference in speleothem calcite δ18O to available measurements. We show that the general European changes are simulated well. However, local discrepancies are found, and might be explained either by a too low model resolution, complex local soil-atmosphere interactions affecting evapotranspiration or by cave specific factors

  1. Radon survey in caves from Mallorca Island, Spain

    Energy Technology Data Exchange (ETDEWEB)

    Dumitru, Oana A. [Department of Geology, Babeș-Bolyai University, Kogălniceanu 1, 400084 Cluj-Napoca (Romania); School of Geosciences, University of South Florida, 4202 E. Fowler Ave., NES 107 Tampa (United States); Onac, Bogdan P. [School of Geosciences, University of South Florida, 4202 E. Fowler Ave., NES 107 Tampa (United States); Fornós, Joan J. [Departament de Ciències de la Terra, Universitat de les Illes Balears, Crta. Valldemossa km 7.5, 07122 Palma (Mallorca) (Spain); Cosma, Constantin [Environmental Radioactivity and Nuclear Dating Center, Babeș-Bolyai University, Fântânele 30, 400294 Cluj-Napoca (Romania); Ginés, Angel; Ginés, Joaquín [Departament de Ciències de la Terra, Universitat de les Illes Balears, Crta. Valldemossa km 7.5, 07122 Palma (Mallorca) (Spain); Merino, Antoni [Grup Espeleològic de Llubí, Federació Balear d' Espeleologia, c/Uruguai s/n, Palma Arena, 07010 Palma, Illes Balears (Spain)

    2015-09-01

    This study reports radon concentration in the most representative caves of Mallorca, identifying those in which the recommended action level is exceeded, thus posing health risks. Two show caves (Campanet and Artà) and three non-touristic caves (Font, Drac, Vallgornera) were investigated. Data were collected at several locations within each cave for three different periods, from March 2013 to March 2014. Except for Vallgornera, where only one monitoring period was possible, and Artà in which low values were recorded throughout the year, a clear seasonal variability, with higher values during the warm seasons and lower during winter time is prominent. Radon concentrations differed markedly from one cave to another, as well as within the same cave, ranging from below detection limit up to 3060 Bq·m{sup −3}. The results of this study have significant practical implications, making it possible to provide some recommendation to cave administrators and other agencies involved in granting access to the investigated caves. - Highlights: • A survey of radon was carried out in caves from Mallorca, Spain using CR 39 detectors. • Three different seasons are covered: spring, summer, and winter. • Radon level ranges from below detection limit up to 3060 Bq·m{sup −3}. • Seasonal variation is evident (higher values in summer and lower during winter). • Particular recommendations were made to each cave administration.

  2. Fish mitigate trophic depletion in marine cave ecosystems.

    Science.gov (United States)

    Bussotti, Simona; Di Franco, Antonio; Bianchi, Carlo Nike; Chevaldonné, Pierre; Egea, Lea; Fanelli, Emanuela; Lejeusne, Christophe; Musco, Luigi; Navarro-Barranco, Carlos; Pey, Alexis; Planes, Serge; Vieux-Ingrassia, Jean Vincent; Guidetti, Paolo

    2018-06-15

    Dark marine habitats are often characterized by a food-limited condition. Peculiar dark habitats include marine caves, characterized by the absence of light and limited water flow, which lead to reduced fluxes of organic matter for cave-dwelling organisms. We investigated whether the most abundant and common cave-dwelling fish Apogon imberbis has the potential to play the role of trophic vector in Mediterranean marine caves. We first analysed stomach contents to check whether repletion changes according to a nycthemeral cycle. We then identified the prey items, to see whether they belong to species associated with cave habitats or not. Finally, we assessed whether A. imberbis moves outside marine caves at night to feed, by collecting visual census data on A. imberbis density both inside and outside caves, by day and by night. The stomach repletion of individuals sampled early in the morning was significantly higher than later in the day. Most prey were typical of habitats other than caves. A. imberbis was on average more abundant within caves during the day and outside during the night. Our study supports the hypothesis regarding the crucial trophic role of A. imberbis in connecting Mediterranean marine caves with external habitats.

  3. Book Review: Caves and Karst of the Yorkshire Dales

    Science.gov (United States)

    Westaway, Rob

    2015-10-01

    The British Cave Research Association (BCRA) is the research division of the British Caving Association (BCA), itself the principal society in Britain for those interested in caving, with activities including provision of training and safety certification for cavers and managing access to caves. Although some UK cave-related research is carried out by academics, this tends to be restricted to archaeological investigations of caves that have served as human habitations, and to be focused more on the occupants than the caves themselves. In contrast, most cave exploration is undertaken as a leisure activity, under the auspices of clubs affiliated to the BCA/BCRA, this being indeed virtually the only field of Earth science where amateur investigators can continue to make significant discoveries. Many cave explorers are also affiliated with academic researchers, such as managers of dating laboratories; the synergy between these two groups is highly productive, having resulted for instance in the discovery and exploration in recent years of the vast Ogof Draenen cave system in South Wales, which probably dates back to the Early Pleistocene (e.g., Farrant et al., 2014).

  4. Cryogenic cave carbonates from the Cold Wind Cave, Nízke Tatry Mountains, Slovakia: Extending the age range of cryogenic cave carbonate formation to the Saalian

    Directory of Open Access Journals (Sweden)

    Zak K.

    2009-07-01

    Full Text Available Cold Wind Cave, located at elevations ranging between 1,600 and 1,700 m a. s. l. in the main range of the Nízke Tatry Mountains(Slovakia, is linked in origin with the adjacent Dead Bats Cave. Together, these caves form a major cave system located within anarrow tectonic slice of Triassic sediments. Both caves have undergone complex multiphase development. A system of sub-horizontalcave levels characterized by large, tunnel-like corridors was formed during the Tertiary, when elevation differences surroundingthe cave were less pronounced than today. The central part of the Nízke Tatry Mountains, together with the cave systems, wasuplifted during the Neogene and Lower Pleistocene, which changed the drainage pattern of the area completely. The formation ofnumerous steep-sloped vadose channels and widespread cave roof frost shattering characterized cave development throughout theQuaternary.In the Cold Wind Cave, extensive accumulations of loose, morphologically variable crystal aggregates of secondary cave carbonateranging in size between less than 1 mm to about 35 mm was found on the surface of fallen limestone blocks. Based on the C andO stable isotope compositions of the carbonate (δ13C: 0.72 to 6.34 ‰, δ18O: –22.61 to –13.68 ‰ V-PDB and the negative relationbetween δ13C and δ18O, the carbonate crystal aggregates are interpreted as being cryogenic cave carbonate (CCC. Publishedmodels suggest the formation of CCC in slowly freezing water pools, probably on the surface of cave ice, most probably duringtransitions from stadials to interstadials. Though the formation of these carbonates is likely one of the youngest events in thesequence of formation of cave sediments of the studied caves, the 230Th/234U ages of three samples (79.7±2.3, 104.0±2.9, and180.0±6.3 ka are the oldest so far obtained for CCC in Central Europe. This is the first description of CCC formation in one caveduring two glacial periods (Saalian and Weichselian.

  5. Contextualizing Cave Maps as Geospatial Information: Case Study of Indonesia

    Science.gov (United States)

    Reinhart, H.

    2017-12-01

    Caves are the result of solution processes. Because they are happened from geochemical and tectonic activity, they can be considered as geosphere phenomena. As one of the geosphere phenomena, especially at karst landform, caves have spatial dimensions and aspects. Cave’s utilizations and developments are increasing in many sectors such as hydrology, earth science, and tourism industry. However, spatial aspects of caves are poorly concerned dues to the lack of recognition toward cave maps. Many stakeholders have not known significances and importance of cave maps in determining development of a cave. Less information can be considered as the cause. Therefore, it is strongly necessary to put cave maps into the right context in order to make stakeholders realize the significance of it. Also, cave maps will be officially regarded as tools related to policy, development, and conservation act of caves hence they will have regulation in the usages and applications. This paper aims to make the contextualization of cave maps toward legal act. The act which is used is Act Number 4 Year 2011 About Geospatial Information. The contextualization is done by scrutinizing every articles and clauses related to cave maps and seek the contextual elements from both of them. The results are that cave maps can be regarded as geospatial information and classified as thematic geospatial information. The usages of them can be regulated through the Act Number 4 Year 2011. The regulations comprised by data acquisition, database, authorities, surveyor, and the obligation of providing cave maps in planning cave’s development and the environment surrounding.

  6. Minerogenetic mechanisms occurring in the cave environment: an overview

    Directory of Open Access Journals (Sweden)

    Forti Paolo

    2011-07-01

    Full Text Available Perhaps man’s first motivation to explore caves, beyond using them as shelter, was the search for substances that were not availableelsewhere: most of them were minerals. However, for a long time it was believed that the cave environment was not very interestingfrom the mineralogical point of view. This was due to the fact that most cave deposits are normally composed of a singlecompound: calcium carbonate. Therefore, the systematic study of cave mineralogy is of only recent origin. However, although onlya limited number of natural cavities have been investigated in detail, about 350 cave minerals have already been observed, someof which are new to science. The presence of such unexpected richness is a direct consequence of the variety of rocks traversedby water or other fluids before entering a cave and the sediments therein. Different cave environments allow the development ofvarious minerogenetic mechanisms, the most important of which are double exchange reactions, evaporation, oxidation, hydrationdehydration,sublimation, deposition from aerosols and vapors, and segregation. The cave temperature and pH/Eh strictly controlmost of them, although some are driven by microorganisms. The cave environment, due to its long-term stability, can sometimesallow for the development of huge euhedral crystals, such as those found in the Naica caves (Mexico, but the presence of extremelysmall yet complex aggregates of different minerals is far more common. Future development in the field of cave mineralogy will likelybe focused mainly on hydrothermal and sulfuric-acid caves and on the role played by micro-organisms in controlling some of the mostimportant minerogenetic processes in caves.

  7. Crowdfunding the Azolla fern genome project: a grassroots approach.

    Science.gov (United States)

    Li, Fay-Wei; Pryer, Kathleen M

    2014-01-01

    Much of science progresses within the tight boundaries of what is often seen as a "black box". Though familiar to funding agencies, researchers and the academic journals they publish in, it is an entity that outsiders rarely get to peek into. Crowdfunding is a novel means that allows the public to participate in, as well as to support and witness advancements in science. Here we describe our recent crowdfunding efforts to sequence the Azolla genome, a little fern with massive green potential. Crowdfunding is a worthy platform not only for obtaining seed money for exploratory research, but also for engaging directly with the general public as a rewarding form of outreach.

  8. A Gompertz regression model for fern spores germination

    Directory of Open Access Journals (Sweden)

    Gabriel y Galán, Jose María

    2015-06-01

    Full Text Available Germination is one of the most important biological processes for both seed and spore plants, also for fungi. At present, mathematical models of germination have been developed in fungi, bryophytes and several plant species. However, ferns are the only group whose germination has never been modelled. In this work we develop a regression model of the germination of fern spores. We have found that for Blechnum serrulatum, Blechnum yungense, Cheilanthes pilosa, Niphidium macbridei and Polypodium feuillei species the Gompertz growth model describe satisfactorily cumulative germination. An important result is that regression parameters are independent of fern species and the model is not affected by intraspecific variation. Our results show that the Gompertz curve represents a general germination model for all the non-green spore leptosporangiate ferns, including in the paper a discussion about the physiological and ecological meaning of the model.La germinación es uno de los procesos biológicos más relevantes tanto para las plantas con esporas, como para las plantas con semillas y los hongos. Hasta el momento, se han desarrollado modelos de germinación para hongos, briofitos y diversas especies de espermatófitos. Los helechos son el único grupo de plantas cuya germinación nunca ha sido modelizada. En este trabajo se desarrolla un modelo de regresión para explicar la germinación de las esporas de helechos. Observamos que para las especies Blechnum serrulatum, Blechnum yungense, Cheilanthes pilosa, Niphidium macbridei y Polypodium feuillei el modelo de crecimiento de Gompertz describe satisfactoriamente la germinación acumulativa. Un importante resultado es que los parámetros de la regresión son independientes de la especie y que el modelo no está afectado por variación intraespecífica. Por lo tanto, los resultados del trabajo muestran que la curva de Gompertz puede representar un modelo general para todos los helechos leptosporangiados

  9. The scaly tree ferns (Cyatheaceae-Polypodiopsida of Brazil

    Directory of Open Access Journals (Sweden)

    Anna Weigand

    2016-01-01

    Full Text Available ABSTRACT A synopsis of all scaly tree fern species (Cyatheaceae occurring in Brazil is presented. We recognize 51 species in three genera [Sphaeropteris one species, Alsophila four species (one subspecies, two varieties, and Cyathea 45 species (one variety ] with 17 taxa being endemic to Brazil. One hybrid endemic to Brazil is recognized. Further included are five species that have not yet been recorded in Brazil, but are expected here because they are found in adjacent countries and occur literally on the border with Brazil. We present the first key covering the family for the whole territory of Brazil.

  10. Changes in soil solution chemistry of Andisols following invasion by bracken fern

    Science.gov (United States)

    J. L. Johnson-Maynard; P. A. McDaniel; D. E. Ferguson; A. L. Falen

    1998-01-01

    Disturbed areas within the Grand Fir Mosaic (GFM) ecosystem of northern Idaho show little to no natural conifer regeneration. Clear-cut sites are invaded quickly by bracken fern successional communities and seem to be in an arrested state of secondary succession. This study compared the soil solution composition of Andisols supporting bracken fern successional...

  11. Asplenium bird’s nest ferns in rainforest canopies are climate-contingent refuges for frogs

    Directory of Open Access Journals (Sweden)

    Brett R. Scheffers

    2014-12-01

    Full Text Available Epiphytes are important for canopy dwelling organisms because they provide a cool and moist microhabitat in the relatively hot and dry canopy. Here we examine whether epiphytic Asplenium ferns act as important habitats for arboreal frogs. We conducted extensive fern and habitat surveys for frogs in the Philippines, and complimented these surveys with roaming day and night canopy surveys to identify the full extent of habitat use across the vertical strata. We artificially dried ferns of various sizes to identify relationships between water and temperature buffering. Ferns are the preferred diurnal microhabitat and breeding habitat for arboreal frogs. A strong positive relationship exists between fern size and frog usage and abundance. Our drying experiments show that large ferns buffer maximum temperatures and reduce variability in temperatures, and buffering is directly linked to their hydration. Frogs are likely using large ferns for their moist, cool, environments for breeding and daytime retreat, which supports the buffered microhabitat hypothesis—these plants promote species coexistence through habitat creation and amelioration of physical stress. However, drying experiments suggest that this buffering is contingent on regular rainfall. Altered rainfall regimes could lead to the unexpected loss of the functional capacity of these important fern habitats. Keywords: Climate change, Functionality, Microhabitat, Refuge, Ectotherm, Precipitation

  12. Spatial Characteristics of Edible Wild Fern Harvesting in Mountainous Villages in Northeastern Japan Using GPS Tracks

    Directory of Open Access Journals (Sweden)

    Toshiya Matsuura

    2014-02-01

    Full Text Available Wild plants in forests provide valuable living resources for rural communities. The location where local people harvest various species is important to the wise use of forest ecosystem services. Using global positioning system (GPS tracking of harvesters’ activities as well as geographic information system (GIS and a generalized linear model (GLM, this study analyzed the spatial differences among harvesting sites of three popular edible ferns, i.e., ostrich fern (Matteuccia struthiopteris, bracken (Pteridium aquilinum, and royal fern (Osmunda japonica, in mountainous villages of Northeastern Japan. The explanatory variables used were vegetation classes, terrain features, and proximity to roadways. The GLM yielded clear differences in harvesting sites among species that were affected by both the species’ ecological characteristics and human behavior. Ostrich fern was harvested mainly in canopy openings along valley floors, whereas royal fern harvest sites were frequently located in snow avalanche scrublands. Bracken was mainly harvested in deforested areas or young conifer plantations. Whereas ostrich fern and bracken harvest sites were restricted by the accessibility from roadways, this was not the case for royal fern. Potential harvest sites of ferns were estimated with the highest value for bracken. Our results suggest that local harvesters seriously consider various natural and anthropogenic factors to maintain effective and sustainable harvesting.

  13. Cave Entrance dependent Spore Dispersion of Filamentous Fungi Isolated from Various Sediments of Iron Ore Cave in Brazil: a colloquy on human threats while caving

    Directory of Open Access Journals (Sweden)

    Erika Linzi Silva Taylor

    2014-04-01

    Full Text Available Caves are stable environments with characteristics favoring the development of fungi. The fungal community present in a cave also includes pathogenic and opportunistic species out of which some are also served as energy sources in such energy stared ecosystems. Studies on microbial diversity and their role on such energy starved ecosystem are scarce. The present study was aimed to identify the cultivable filamentous fungi present in the various sediment of an iron ore cave and to recognize them as pathogenic and/or opportunistic species. Further the impact of cave entrance on the spore depositions on various distances dependent sediments were analyzed. The results suggest a diverse microbial community inhabiting the cave and an influence of cave entrance over spore deposition on various sediments. We counted a total of 4,549 filamentous fungi that included 34 species of 12 genera: Acremonium, Aspergillus, Cladosporium, Fusarium, Geotrichum, Paecilomyces, Purpureocillium, Penicillium, Torula, Trichoderma, Mucor and Rhizopus. A positive significant relation was observed between spore deposition and distance from cave entrance (p= 0.001. Areas of potential mycoses risks were recognized. This is the first study on microbiological community of an iron ore cave in the country.

  14. Hotspot mitigation in the StarCAVE

    KAUST Repository

    Rhee, Jordan; Schulze, Jurgen; DeFanti, Thomas A.

    2010-01-01

    Rear-projected screens such as those in Digital Light Projection (DLP) televisions suffer from an image quality problem called hot spotting, where the image is brightest at a point dependent on the viewing angle. In rear-projected mulit-screen configurations such as the StarCAVE at Calit2, this causes discontinuities in brightness at the edges where screens meet, and thus in the 3D image perceived by the user. In the StarCAVE we know the viewer's position in 3D space and we have programmable graphics hardware, so we can mitigate this effect by performing post-processing in the inverse of the pattern, yielding a homogenous image at the output. Our implementation improves brightness homogeneity by a factor of 4 while decreasing frame rate by only 1-3 fps.

  15. Hotspot mitigation in the StarCAVE

    KAUST Repository

    Rhee, Jordan

    2010-01-27

    Rear-projected screens such as those in Digital Light Projection (DLP) televisions suffer from an image quality problem called hot spotting, where the image is brightest at a point dependent on the viewing angle. In rear-projected mulit-screen configurations such as the StarCAVE at Calit2, this causes discontinuities in brightness at the edges where screens meet, and thus in the 3D image perceived by the user. In the StarCAVE we know the viewer\\'s position in 3D space and we have programmable graphics hardware, so we can mitigate this effect by performing post-processing in the inverse of the pattern, yielding a homogenous image at the output. Our implementation improves brightness homogeneity by a factor of 4 while decreasing frame rate by only 1-3 fps.

  16. Sensing Structures Inspired by Blind Cave Fish

    Science.gov (United States)

    McConney, Michael E.; Chen, Nannan; Lu, David; Anderson, Kyle D.; Hu, Huan; Liu, Chang; Tsukruk, Vladimir V.

    2009-03-01

    Blind cave fish, with degenerated non-functioning eyes, have evolved to ``see'' their hydrodynamic environment by using the flow receptors of the lateral line system. The hair-cell receptors are encapsulated in a hydrogel-like material, called a cupula, which increases the sensitivity of the hair-cell receptors by coupling their motion to the surrounding flowing media. We characterized the viscoelastic properties and of blind cave fish cupulae by using colloidal-probe spectroscopy in fluid. A photo-patternable hydrogel with similar properties was developed to mimic the fish receptor coupling structure. Flow-based measurements indicated that the hydrogels enhance drag through increased surface area, but also inherent material properties. These bio-inspired structures endowed micro-fabricated flow sensors with sensitivities rivaling that of fish.

  17. Hawaiian lavas: a window into mantle dynamics

    Science.gov (United States)

    Jones, Tim; Davies, Rhodri; Campbell, Ian

    2017-04-01

    The emergence of double track volcanism at Hawaii has traditionally posed two problems: (i) the physical emergence of two parallel chains of volcanoes at around 3 Ma, named the Loa and Kea tracks after the largest volcanoes in their sequence, and (ii) the systematic geochemical differences between the erupted lavas along each track. In this study, we dissolve this distinction by providing a geodynamical explanation for the physical emergence of double track volcanism at 3 Ma and use numerical models of the Hawaiian plume to illustrate how this process naturally leads to each volcanic track sampling distinct mantle compositions, which accounts for much of the geochemical characteristics of the Loa and Kea trends.

  18. The effect of nitrogen additions on bracken fern and its insect herbivores at sites with high and low atmospheric pollution

    Science.gov (United States)

    M.E. Jones; M.E. Fenn; T.D. Paine

    2011-01-01

    The impact of atmospheric pollution, including nitrogen deposition, on bracken fern herbivores has never been studied. Bracken fern is globally distributed and has a high potential to accumulate nitrogen in plant tissue. We examined the response of bracken fern and its herbivores to N fertilization at a high and low pollution site in forests downwind of Los Angeles,...

  19. Variation of functional clonal traits along elevation in two fern species

    International Nuclear Information System (INIS)

    Song, Y.B.; Chen, L.Y.; Xiong, W.

    2015-01-01

    Phenotypical plasticity is generally considered among adaptive strategies by which plants can cope with environmental variation in space and time. Although much is known about plasticity in seed plants in terms of functional clonal traits while little is known about ferns. Variation of functional clonal traits of two ferns Dicranopteris dichotoma and Diplopterygium glaucum among plots differing in elevation in a subtropical evergreen broad-leaved forest in southern China was investigated. Along with elevation increasing the two fern species showed similar variation pattern of functional clonal traits: stable spacer length, increasing specific spacer length and decreasing spacer weight per ramet and specific spacer weight. The two ferns species had similar variation pattern of ramet performance traits but different variation pattern of ramet population properties. These results suggest an evolutionary trade-off between functions of foraging for and storing of resources in the two ferns, with a functional preference for the foraging in response to environmental change. (author)

  20. In Vitro Conservation of Some Threatened and Economically Important Ferns Belonging to the Indian Subcontinent

    Directory of Open Access Journals (Sweden)

    Shastri P. Shukla

    2014-01-01

    Full Text Available This study was undertaken to identify methods of mass multiplication for five ornamental, economically important ferns (Nephrolepis biserrata (Sw. Schott., N. cordifolia cv. ‘‘duffii’’ (L. Presl., N. exaltata cv. bostoniensis (L. Schott., Pteris vittata L., and Cyclosorus dentatus Link., and three threatened ferns, namely, Cyathea spinulosa Wall. ex. Hook, Pityrogramma calomelanos (L. Link., and Microsorum punctatum (L. Schott., through in vitro techniques. Collections were made from different biodiversity zones of India including Northeast Himalayas, Kumaon Himalayas, and Western Ghat and successfully introduced and grown in a fern-house. Aseptic cultures were raised at the morphogenic level of callus, axillary shoot, multiple shoot, and rooted plants. An optimized medium is described for each fern species. Plantlets were also produced from spore culture of Cyathea spinulosa and successfully hardened under fern house conditions.

  1. Calcium carbonate concretions in caves : an overview

    International Nuclear Information System (INIS)

    Gewelt, M.; Ek, C.

    1988-01-01

    The scientific work of the last twenty years on calcium carbonate cave deposits (dripstones and flowstones) is presented. Recent studies on speleothems composition, growth, age and paleoclimatic environment are examined. Main new results are related with the development of isotopic and radiometric dating methods. Increasing numbers of dates allow for statistical speleothem repartition studies which could be related with paleoclimates. Two new frequency curves of U-series ages data of speleothems are given. (M.C.B.)

  2. 76 FR 42654 - Endangered and Threatened Wildlife and Plants; Petition To List Grand Canyon Cave Pseudoscorpion

    Science.gov (United States)

    2011-07-19

    ... from 1977 to 1978. Eight caves were examined including Babylon Cave, Crystal Forest Cave, Land's End...) describes examining the walls, ceilings, and floors for animals and invertebrates. He identified 12...

  3. Calculating flux to predict future cave radon concentrations

    Czech Academy of Sciences Publication Activity Database

    Rowberry, Matthew David; Martí, Xavier; Frontera, C.; Van De Wiel, M.J.; Briestenský, Miloš

    2016-01-01

    Roč. 157, JUN (2016), 16-26 ISSN 0265-931X R&D Projects: GA MŠk LM2010008 Institutional support: RVO:67985891 ; RVO:68378271 Keywords : cave radon concentration * cave radon flux * cave ventilation * radioactive decay * fault slip * numerical modelling Subject RIV: DC - Siesmology, Volcanology, Earth Structure; BG - Nuclear, Atomic and Molecular Physics, Colliders (FZU-D) Impact factor: 2.310, year: 2016

  4. Depth Perception in Cave and Panorama

    DEFF Research Database (Denmark)

    Mullins, Michael; Strojan, Tadeja Zupancic

    2004-01-01

    This study compares aspects of spatial perception in a physical environment and its virtual representations in a CAVE and Panorama, derived from recent research. To measure accuracy of spatial perception, participants in an experiment were asked to look at identical objects in the three environme......, learning and training in virtual environments; in architectural education; and participatory design processes, in which the dialogue between real and imagined space may take place in virtual . reality environments...... environments and then locate them and identify their shape on scaled drawings.  Results are presented together with statistical analysis. In a discussion of the results, the paper addresses the two hypothetical assertions ? that depth perception in physical reality and its virtual representations in CAVE......This study compares aspects of spatial perception in a physical environment and its virtual representations in a CAVE and Panorama, derived from recent research. To measure accuracy of spatial perception, participants in an experiment were asked to look at identical objects in the three...

  5. Radiocarbon intercomparison program for Chauvet Cave

    International Nuclear Information System (INIS)

    Cuzange, M.T.; Delque-Kolic, E.; Oberlin, C.; Goslar, T.; Grootes, P.M.; Nadeau, M.J.; Higham, T.; Ramsey, C.B.; Kaltnecker, E.; Paterne, M.; Valladas, H.; Van der Plicht, J.; Van der Plicht, J.; Clottes, J.; Geneste, J.M.

    2007-01-01

    We present the first results of an accelerator mass spectrometry (AMS) radiocarbon intercomparison program on 3 different charcoal samples collected in one of the hearths of the Megaceros gallery of Chauvet Cave (Ardeche, France). This cave, rich in parietal decoration, is important for the study of the appearance and evolution of prehistoric art because certain drawings have been 14 C dated to the Aurignacian period at the beginning of the Upper Paleolithic. The new dates indicate an age of about 32,000 BP, which is consistent with this attribution and in agreement with the results from the same sector of the cave measured previously at the Laboratoire des Sciences du Climat et de l'Environnement (LSCE). Six laboratories were involved in the intercomparison. Samples were measured in 4 AMS facilities: Center for Isotope Research, Groningen University, the Netherlands; the Oxford Radiocarbon Accelerator Unit, UK; the Centre de datation par le carbone 14, Univ. Claude Bernard Lyon 1, France (measured by AMS facilities of Poznan University, Poland); and the LSCE, UMR CEA-CNRS-UVSQ, France (measured by the Leibniz-Labor of Christian-Albrechts-Universitat Kiel, Germany). (authors)

  6. Eruptive behavior of the Marum/Mbwelesu lava lake, Vanuatu and comparisons with lava lakes on Earth and Io

    Science.gov (United States)

    Radebaugh, Jani; Lopes, Rosaly M.; Howell, Robert R.; Lorenz, Ralph D.; Turtle, Elizabeth P.

    2016-08-01

    Observations from field remote sensing of the morphology, kinematics and temperature of the Marum/Mbwelesu lava lake in the Vanuatu archipelago in 2014 reveal a highly active, vigorously erupting lava lake. Active degassing and fountaining observed at the 50 m lava lake led to large areas of fully exposed lavas and rapid ( 5 m/s) movement of lava from the centers of upwelling outwards to the lake margins. These rapid lava speeds precluded the formation of thick crust; there was never more than 30% non-translucent crust. The lava lake was observed with several portable, handheld, low-cost, near-infrared imagers, all of which measured temperatures near 1000 °C and one as high as 1022 °C, consistent with basaltic temperatures. Fine-scale structure in the lava fountains and cooled crust was visible in the near infrared at 5 cm/pixel from 300 m above the lake surface. The temperature distribution across the lake surface is much broader than at more quiescent lava lakes, peaking 850 °C, and is attributed to the highly exposed nature of the rapidly circulating lake. This lava lake has many characteristics in common with other active lava lakes, such as Erta Ale in Ethiopia, being confined, persistent and high-temperature; however it was much more active than is typical for Erta Ale, which often has > 90% crust. Furthermore, it is a good analogue for the persistent, high-temperature lava lakes contained within volcanic depressions on Jupiter's moon Io, such as Pele, also believed from spacecraft and ground-based observations to exhibit similar behavior of gas emission, rapid overturn and fountaining.

  7. An Investigation of Meromixis in Cave Pools, Lechuguilla Cave, New Mexico.

    Directory of Open Access Journals (Sweden)

    David B. Levy

    2008-07-01

    Full Text Available Chemical characteristics of permanent stratification in cave pools (meromixis may provide insight into the geochemical origin and evolution of cave pool waters. The objective of this study was to test the hypothesis that some pools in Lechuguilla Cave may be subject to ectogenic meromixis, where permanent chemical stratification is induced by input of relatively saline or fresh water from an external source. However, because organic C concentrations in Lechuguilla waters are low (typically 0.9 m, and are probably the result of localized and transient atmospheric CO2(g concentrations. At LOBG, an EC increase of 93 µS cm-1 at the 0.9-m depth suggests meromictic conditions which are ectogenic, possibly due to surface inflow of fresh water as drips or seepage into a pre-existing layer of higher salinity.

  8. Ultrasonic synthesis of fern-like ZnO nanoleaves and their enhanced photocatalytic activity

    International Nuclear Information System (INIS)

    Ma, Qing Lan; Xiong, Rui; Zhai, Bao-gai; Huang, Yuan Ming

    2015-01-01

    Graphical abstract: - Highlights: • Fern-like ZnO nanoleaves were synthesized by ultrasonicating Zn microcrystals in water. • A fern-like ZnO nanoleaf is a self-assembly of ZnO nanoplates along one ZnO nanorod. • Fern-like ZnO nanoleaves exhibit enhanced photocatalytic activity than ZnO nanocrystals. • The branched hierarchical structures are responsible for the enhanced photocatalytic activity. - Abstract: Two-dimensional fern-like ZnO nanoleaves were synthesized by ultrasonicating zinc microcrystals in water. The morphology, crystal structure, optical property and photocatalytic activity of the fern-like ZnO nanoleaves were characterized with scanning electron microscopy, X-ray diffraction, transmission electron microscopy, photoluminescence spectroscopy and ultraviolet–visible spectroscopy, respectively. It is found that one fern-like ZnO nanoleaf is composed of one ZnO nanorod as the central trunk and a number of ZnO nanoplates as the side branches in opposite pairs along the central ZnO nanorod. The central ZnO nanorod in the fern-like nanoleaves is about 1 μm long while the side-branching ZnO nanoplates are about 100 nm long and 20 nm wide. Further analysis has revealed that ZnO nanocrystals are the building blocks of the central ZnO nanorod and the side-branching ZnO nanoplates. Under identical conditions, fern-like ZnO nanoleaves exhibit higher photocatalytic activity in photodegrading methyl orange in aqueous solution than spherical ZnO nanocrystals. The first-order photocatalytic rate constant of the fern-like ZnO nanoleaves is about four times as large as that of the ZnO nanoparticles. The branched architecture of the hierarchical nanoleaves is suggested be responsible for the enhanced photocatalytic activity of the fern-like ZnO nanoleaves

  9. Some deep caves in Biokovo Mountain (Croatia)

    Science.gov (United States)

    Garasic, Mladen; Garasic, Davor

    2014-05-01

    The investigation of 3 caves explored more than 1000 meters in depth in the Dinaric karst area in Croatia, has been in progress for a considerable period of time. These are complex speleological features situated in the longest mountain range of the Dinaric karst, i.e. at the Northern Velebit mountain range. In fact, these caves have been studied for over two decades now. The first one is a cave system of Lukina jama (Luke's Cave) - Trojama cave, which has been investigated until the depth of 1421 meters (Jalžić, 2007; Šmida, 1993). Its total length is 3731 meters and a new expedition will soon continue to investigate this pit through speleodiving in siphons. The second greatest cave by depth is Slovačka jama (Slovak Cave), 1320 meters in depth, with cave chanals measuring 5677 meters in total length. The third greatest cave by depth is the Cave system of Velebita, reaching down to 1026 m in depth, with the chanal length of 3176 meters (Bakšić, 2006a; 2006b). However, another 3 speleological sites, which can rightly be added to those deeper than 1000 m, have recently been discovered. These are three caverns that were discovered during construction of the Sveti Ilija Tunnel that passes through Mt. Biokovo, in the Dinaric karst area. These caverns undoubtedly point to the link with the ground surface, while the rock overburden above the tunnel in the zone where the caverns were discovered ranges from 1250 and 1350 meters. Bats from the ground surface were found in the caverns and, according to measurements, they are situated in the depth from 200 and 300 meters below the tunnel level. This would mean that the depth of these newly found caves ranges from 1450 and 1650 m, when observed from the ground surface. There are several hundreds of known caves in Biokovo, and the deepest ones discovered so far are Jama Mokre noge (Wet Feet Cave) 831 m in depth, and Jama Amfora (Amphora Cave) 788 m in depth (Bockovac, 1999; Bakšić & all, 2002; Lacković & all, 2001

  10. Discovery of a diverse cave flora in China

    Science.gov (United States)

    Wen, Fang

    2018-01-01

    Few studies document plants in caves. Our field observations of a widespread and seemingly angiosperm-rich cave flora in SW China lead us to test the following hypotheses, 1) SW China caves contain a diverse vascular plant flora, 2) that this is a relic of a largely absent forest type lacking endemic species, and 3) that the light environment plants occupy in caves is not distinct from non-cave habitats. To do so we surveyed 61 caves and used species accumulation curves (SAC) to estimate the total diversity of this flora and used a subsample of 14 caves to characterise the light environment. We used regional floras and existing conservation assessments to evaluate the conservation value of this flora. We used observations on human disturbance within caves to evaluate anthropogenic activities. Four-hundred-and-eighteen vascular plant species were documented with SACs predicting a total diversity of 529–846. Ninety-three percent of the species documented are known karst forest species, 7% are endemic to caves and 81% of the species are angiosperms. We demonstrate that the light environment in caves is distinct to that of terrestrial habitats and that a subset of the flora likely grow in the lowest light levels documented for vascularised plants. Our results suggest that the proportion of species threatened with extinction is like that for the terrestrial habitat and that almost half of the entrance caverns sampled showed signs of human disturbance. We believe that this is the first time that such an extensive sample of cave flora has been undertaken and that such a diverse vascular plant flora has been observed in caves which we predict occurs elsewhere in SE Asia. We argue that the cave flora is an extension of the karst forest understory present prior to catastrophic deforestation in the 20thC. We suggest that within SW China caves serve as both refuges and a valuable source of germplasm for the restoration of karst forest. We also propose that caves represent a

  11. Discovery of a diverse cave flora in China.

    Science.gov (United States)

    Monro, Alexandre K; Bystriakova, Nadia; Fu, Longfei; Wen, Fang; Wei, Yigang

    2018-01-01

    Few studies document plants in caves. Our field observations of a widespread and seemingly angiosperm-rich cave flora in SW China lead us to test the following hypotheses, 1) SW China caves contain a diverse vascular plant flora, 2) that this is a relic of a largely absent forest type lacking endemic species, and 3) that the light environment plants occupy in caves is not distinct from non-cave habitats. To do so we surveyed 61 caves and used species accumulation curves (SAC) to estimate the total diversity of this flora and used a subsample of 14 caves to characterise the light environment. We used regional floras and existing conservation assessments to evaluate the conservation value of this flora. We used observations on human disturbance within caves to evaluate anthropogenic activities. Four-hundred-and-eighteen vascular plant species were documented with SACs predicting a total diversity of 529-846. Ninety-three percent of the species documented are known karst forest species, 7% are endemic to caves and 81% of the species are angiosperms. We demonstrate that the light environment in caves is distinct to that of terrestrial habitats and that a subset of the flora likely grow in the lowest light levels documented for vascularised plants. Our results suggest that the proportion of species threatened with extinction is like that for the terrestrial habitat and that almost half of the entrance caverns sampled showed signs of human disturbance. We believe that this is the first time that such an extensive sample of cave flora has been undertaken and that such a diverse vascular plant flora has been observed in caves which we predict occurs elsewhere in SE Asia. We argue that the cave flora is an extension of the karst forest understory present prior to catastrophic deforestation in the 20thC. We suggest that within SW China caves serve as both refuges and a valuable source of germplasm for the restoration of karst forest. We also propose that caves represent a

  12. Radon in an underground cave system in Victoria

    International Nuclear Information System (INIS)

    Hedt, J.C.; Boal, T.J.

    1998-01-01

    Radon levels in a cave system in Victoria have been measured. The variation of radon and radon progeny levels with time, position throughout the cave and season have been determined. The radiation exposure of tour guides were estimated. The data is being used to develop a radiation management plan for the tour guides. Radon concentration within a cave system was proven to be dependent to a large extent upon the rate of air exchange with outside. Cave ventilation is the single most important factor in determining if there is diurnal variation in the radon concentration

  13. Caves as observatories for atmospheric thermal tides: an example from Ascunsă Cave, Romania

    Directory of Open Access Journals (Sweden)

    Virgil Drăgușin

    2018-01-01

    Full Text Available As part of a microclimate study at Ascunsă Cave, Romania, we used Gemini Tinytag Plus 2 data loggers to record cave air temperature variability. At one of the monitoring points we recognized the presence of semidiurnal cycles on the order of a few thousands of a degree Celsius that could be produced under the influence of the semidiurnal tidal components of the Sun (S2 or the Moon (M2. Using a Gemini Tinytag Plus 2 data logger with an external probe we measured core rock temperature and showed that it does not influence the cave air temperature on such short time scales. We thus rejected the possibility that Earth tides, mostly produced by the lunar tidal influence on the Earth’s crust, would have had a semidiurnal influence on cave air temperature. Moreover, time series analysis revealed a 12.00-hour periodicity in temperature data, specific for the S2, allowing us to assign these variations to the influence of the thermo-tidal action of the Sun. Using the Ideal Gas Law and assuming a constant volume and amount of air, we calculated that a theoretical change in atmospheric pressure of around 40 Pa was needed to produce the temperature changes we observed. This agrees with published values of atmospheric pressure changes induced by the semidiurnal solar component of the thermal tides (S2(t. We thus can assign the observed temperature changes to semidiurnal atmospheric pressure changes (S2(p induced by the thermal excitation of the Sun. Our study signals the possibility that readily available data from cave monitoring studies around the world could be used in the study of atmospheric tides. Moreover, it appears that Ascunsă Cave acts as a natural meteorological filter on a short time scale, removing the direct thermal influences of the Sun (especially night and day differences and preserving only the barometric information from the surface.

  14. Lava tube shatter rings and their correlation with lava flux increases at Kīlauea Volcano, Hawai‘i

    Science.gov (United States)

    Orr, T.R.

    2011-01-01

    Shatter rings are circular to elliptical volcanic features, typically tens of meters in diameter, which form over active lava tubes. They are typified by an upraised rim of blocky rubble and a central depression. Prior to this study, shatter rings had not been observed forming, and, thus, were interpreted in many ways. This paper describes the process of formation for shatter rings observed at Kīlauea Volcano during November 2005–July 2006. During this period, tilt data, time-lapse images, and field observations showed that episodic tilt changes at the nearby Pu‘u ‘Ō‘ō cone, the shallow magmatic source reservoir, were directly related to fluctuations in the level of lava in the active lava tube, with periods of deflation at Pu‘u ‘Ō‘ō correlating with increases in the level of the lava stream surface. Increases in lava level are interpreted as increases in lava flux, and were coincident with lava breakouts from shatter rings constructed over the lava tube. The repetitive behavior of the lava flux changes, inferred from the nearly continuous tilt oscillations, suggests that shatter rings form from the repeated rise and fall of a portion of a lava tube roof. The locations of shatter rings along the active lava tube suggest that they form where there is an abrupt decrease in flow velocity through the tube, e.g., large increase in tube width, abrupt decrease in tube slope, and (or) sudden change in tube direction. To conserve volume, this necessitates an abrupt increase in lava stream depth and causes over-pressurization of the tube. More than a hundred shatter rings have been identified on volcanoes on Hawai‘i and Maui, and dozens have been reported from basaltic lava fields in Iceland, Australia, Italy, Samoa, and the mainland United States. A quick study of other basaltic lava fields worldwide, using freely available satellite imagery, suggests that they might be even more common than previously thought. If so, this confirms that episodic

  15. Conservation Priorities for Tree Ferns (Cyatheaceae in Sri Lanka

    Directory of Open Access Journals (Sweden)

    R. H. G. Ranil

    2011-09-01

    Full Text Available Diversity, phytogeography and conservation status of Sri Lankan tree-ferns are discussed in this paper. The family Cyatheaceae is represented by eight taxa (seven species and one doubtful variety in Sri Lanka with a high rate of endemism of 75%. Apart from Cyathea walkerae and C. gigantea, the other species are restricted to geographically isolated areas in the country with limited population sizes. Fortunately, all Sri Lankan species of Cyathea occur within the protected areas of the wet zone. However, ex situ conservation is limited to C. walkerae and C. crinita at Botanic Gardens in Hakgala. Despite the family being listed in Annex II of CITES, its species have not yet been assessed in Sri Lanka for the Red Listing criteria. Identification of the nature and level of threat to Sri Lankan Cyathea species is therefore a major priority, followed by the monitoring of populations in situ in protected areas in the wet zone. Ex situ conservation of rare species and cultivation of Cyathea species from spores have also been identified as priority areas. A strong programme should be developed with the National Herbarium to explore little known forest patches in the wet zone to enhance our knowledge of Cyathea species in Sri Lanka. Such information will provide a strong basis for preparing a conservation and management plan for tree-ferns in the country.

  16. Invariant Hough Random Ferns for Object Detection and Tracking

    Directory of Open Access Journals (Sweden)

    Yimin Lin

    2014-01-01

    Full Text Available This paper introduces an invariant Hough random ferns (IHRF incorporating rotation and scale invariance into the local feature description, random ferns classifier training, and Hough voting stages. It is especially suited for object detection under changes in object appearance and scale, partial occlusions, and pose variations. The efficacy of this approach is validated through experiments on a large set of challenging benchmark datasets, and the results demonstrate that the proposed method outperforms state-of-the-art conventional methods such as bounding-box-based and part-based methods. Additionally, we also propose an efficient clustering scheme based on the local patches’ appearance and their geometric relations that can provide pixel-accurate, top-down segmentations from IHRF back-projections. This refined segmentation can be used to improve the quality of online object tracking because it avoids the drifting problem. Thus, an online tracking framework based on IHRF, which is trained and updated in each frame to distinguish and segment the object from the background, is established. Finally, the experimental results on both object segmentation and long-term object tracking show that this method yields accurate and robust tracking performance in a variety of complex scenarios, especially in cases of severe occlusions and nonrigid deformations.

  17. Hydrological characterization of cave drip waters in a porous limestone: Golgotha Cave, Western Australia

    Science.gov (United States)

    Mahmud, Kashif; Mariethoz, Gregoire; Baker, Andy; Treble, Pauline C.

    2018-02-01

    Cave drip water response to surface meteorological conditions is complex due to the heterogeneity of water movement in the karst unsaturated zone. Previous studies have focused on the monitoring of fractured rock limestones that have little or no primary porosity. In this study, we aim to further understand infiltration water hydrology in the Tamala Limestone of SW Australia, which is Quaternary aeolianite with primary porosity. We build on our previous studies of the Golgotha Cave system and utilize the existing spatial survey of 29 automated cave drip loggers and a lidar-based flow classification scheme, conducted in the two main chambers of this cave. We find that a daily sampling frequency at our cave site optimizes the capture of drip variability with the least possible sampling artifacts. With the optimum sampling frequency, most of the drip sites show persistent autocorrelation for at least a month, typically much longer, indicating ample storage of water feeding all stalactites investigated. Drip discharge histograms are highly variable, showing sometimes multimodal distributions. Histogram skewness is shown to relate to the wetter-than-average 2013 hydrological year and modality is affected by seasonality. The hydrological classification scheme with respect to mean discharge and the flow variation can distinguish between groundwater flow types in limestones with primary porosity, and the technique could be used to characterize different karst flow paths when high-frequency automated drip logger data are available. We observe little difference in the coefficient of variation (COV) between flow classification types, probably reflecting the ample storage due to the dominance of primary porosity at this cave site. Moreover, we do not find any relationship between drip variability and discharge within similar flow type. Finally, a combination of multidimensional scaling (MDS) and clustering by k means is used to classify similar drip types based on time series

  18. Hydrological characterization of cave drip waters in a porous limestone: Golgotha Cave, Western Australia

    Directory of Open Access Journals (Sweden)

    K. Mahmud

    2018-02-01

    Full Text Available Cave drip water response to surface meteorological conditions is complex due to the heterogeneity of water movement in the karst unsaturated zone. Previous studies have focused on the monitoring of fractured rock limestones that have little or no primary porosity. In this study, we aim to further understand infiltration water hydrology in the Tamala Limestone of SW Australia, which is Quaternary aeolianite with primary porosity. We build on our previous studies of the Golgotha Cave system and utilize the existing spatial survey of 29 automated cave drip loggers and a lidar-based flow classification scheme, conducted in the two main chambers of this cave. We find that a daily sampling frequency at our cave site optimizes the capture of drip variability with the least possible sampling artifacts. With the optimum sampling frequency, most of the drip sites show persistent autocorrelation for at least a month, typically much longer, indicating ample storage of water feeding all stalactites investigated. Drip discharge histograms are highly variable, showing sometimes multimodal distributions. Histogram skewness is shown to relate to the wetter-than-average 2013 hydrological year and modality is affected by seasonality. The hydrological classification scheme with respect to mean discharge and the flow variation can distinguish between groundwater flow types in limestones with primary porosity, and the technique could be used to characterize different karst flow paths when high-frequency automated drip logger data are available. We observe little difference in the coefficient of variation (COV between flow classification types, probably reflecting the ample storage due to the dominance of primary porosity at this cave site. Moreover, we do not find any relationship between drip variability and discharge within similar flow type. Finally, a combination of multidimensional scaling (MDS and clustering by k means is used to classify similar drip

  19. What factors control the superficial lava dome explosivity?

    Science.gov (United States)

    Boudon, Georges; Balcone-Boissard, Hélène; Villemant, Benoit; Morgan, Daniel J.

    2015-04-01

    Dome-forming eruption is a frequent eruptive style; lava domes result from intermittent, slow extrusion of viscous lava. Most dome-forming eruptions produce highly microcrystallized and highly- to almost totally-degassed magmas which have a low explosive potential. During lava dome growth, recurrent collapses of unstable parts are the main destructive process of the lava dome, generating concentrated pyroclastic density currents (C-PDC) channelized in valleys. These C-PDC have a high, but localized, damage potential that largely depends on the collapsed volume. Sometimes, a dilute ash cloud surge develops at the top of the concentrated flow with an increased destructive effect because it may overflow ridges and affect larger areas. In some cases, large lava dome collapses can induce a depressurization of the magma within the conduit, leading to vulcanian explosions. By contrast, violent, laterally directed, explosions may occur at the base of a growing lava dome: this activity generates dilute and turbulent, highly-destructive, pyroclastic density currents (D-PDC), with a high velocity and propagation poorly dependent on the topography. Numerous studies on lava dome behaviors exist, but the triggering of lava dome explosions is poorly understood. Here, seven dome-forming eruptions are investigated: in the Lesser Antilles arc: Montagne Pelée, Martinique (1902-1905, 1929-1932 and 650 y. BP eruptions), Soufrière Hills, Montserrat; in Guatemala, Santiaguito (1929 eruption); in La Chaîne des Puys, France (Puy de Dome and Puy Chopine eruptions). We propose a new model of superficial lava-dome explosivity based upon a textural and geochemical study (vesicularity, microcrystallinity, cristobalite distribution, residual water contents, crystal transit times) of clasts produced by these key eruptions. Superficial explosion of a growing lava dome may be promoted through porosity reduction caused by both vesicle flattening due to gas escape and syn-eruptive cristobalite

  20. A flexible open-source toolkit for lava flow simulations

    Science.gov (United States)

    Mossoux, Sophie; Feltz, Adelin; Poppe, Sam; Canters, Frank; Kervyn, Matthieu

    2014-05-01

    Lava flow hazard modeling is a useful tool for scientists and stakeholders confronted with imminent or long term hazard from basaltic volcanoes. It can improve their understanding of the spatial distribution of volcanic hazard, influence their land use decisions and improve the city evacuation during a volcanic crisis. Although a range of empirical, stochastic and physically-based lava flow models exists, these models are rarely available or require a large amount of physical constraints. We present a GIS toolkit which models lava flow propagation from one or multiple eruptive vents, defined interactively on a Digital Elevation Model (DEM). It combines existing probabilistic (VORIS) and deterministic (FLOWGO) models in order to improve the simulation of lava flow spatial spread and terminal length. Not only is this toolkit open-source, running in Python, which allows users to adapt the code to their needs, but it also allows users to combine the models included in different ways. The lava flow paths are determined based on the probabilistic steepest slope (VORIS model - Felpeto et al., 2001) which can be constrained in order to favour concentrated or dispersed flow fields. Moreover, the toolkit allows including a corrective factor in order for the lava to overcome small topographical obstacles or pits. The lava flow terminal length can be constrained using a fixed length value, a Gaussian probability density function or can be calculated based on the thermo-rheological properties of the open-channel lava flow (FLOWGO model - Harris and Rowland, 2001). These slope-constrained properties allow estimating the velocity of the flow and its heat losses. The lava flow stops when its velocity is zero or the lava temperature reaches the solidus. Recent lava flows of Karthala volcano (Comoros islands) are here used to demonstrate the quality of lava flow simulations with the toolkit, using a quantitative assessment of the match of the simulation with the real lava flows. The

  1. Biomineralization and biosignatures of coralloid-type speleothems from lava tubes of Galapagos Islands: evidences on the fossil record of prokaryotes

    Science.gov (United States)

    Miller, Ana Z.; Garcia-Sanchez, Angela M.; Pereira, Manuel F. C.; Gazquez, Fernando; Calaforra, José M.; Forti, Paolo; Toulkeridis, Theofilos; Martínez-Frías, Jesús; Saiz-Jimenez, Cesareo

    2016-04-01

    Lava tubes have traditionally been considered of little interest from a mineralogical point of view. Recently, this type of volcanic caves has received particular attention because lava tubes have been described on Mars. Speleothems, or secondary mineral deposits in lava tubes are mainly composed of siliceous minerals. Coralloid-type speleothems are found either on basaltic cave walls or on the surface of other speleothems. Several authors attribute a microbially mediated origin to their formation. This type of speleothems was recorded within Royal Palm Cave of Santa Cruz Island in Galapagos Archipelago (Ecuador), a lava tube 600 m long, 5 to 15 m height and 2 to 10 m width. The Galapagos Islands are an archipelago of 19 volcanic islands located some 1500 km west of Ecuador, in the Pacific Ocean. These islands host one of the most biodiverse settings on Earth, studied by Charles Darwin. Beige and greyish small coralloids were collected in Royal Palm Cave and analysed by field emission scanning electron microscopy with energy dispersive X-ray spectroscopy (FESEM-EDS), X-ray micro-computed tomography (micro-CT) and mineralogical analyses for morphological, 3D microstructural and compositional characterization, as well as for assessing microbe-mineral interactions and biogenicity. In addition, 16S rRNA gene analyses were performed to identify microbial communities associated with the coralloid-type speleothems. The coralloids showed internal compositional zonation along the growth direction of the speleothems, according to micro-CT data. Internal layering was clearly discernable by the differences in opacity of the distinct mineralogical phases to X-rays, being dominated by alteration products of siliceous composition, whereas more opaque phases, usually Ca-rich minerals, were dominant in the outermost part of the speleothems. X-ray diffraction and infrared spectroscopy reinforced that the first stage of deposition is mainly composed of opal A and clay minerals

  2. Lava inundation zone maps for Mauna Loa, Island of Hawaiʻi, Hawaii

    Science.gov (United States)

    Trusdell, Frank A.; Zoeller, Michael H.

    2017-10-12

    Lava flows from Mauna Loa volcano, on the Island of Hawaiʻi, constitute a significant hazard to people and property. This report addresses those lava flow hazards, mapping 18 potential lava inundation zones on the island.

  3. Characteristics and genesis of porphyroclastic lava rock in Xiangshan

    International Nuclear Information System (INIS)

    Zhou Xiaohua; Wang Zhuning

    2012-01-01

    Due to the transitional characteristics of porphyroclastic lava rock in Xiangshan of Jiangxi province, there are a variety of views on its genesis, petrographic attribution. This is because the marginal facies of the porphyroclastic lava is with ignimbrite and tuff characteristics, its transition phase has the characteristics of lava, and its intermediate phase has the feature of sub-volcanic rocks, further more, different texture of the rocks bears transition relationship. By the study of mineral composition, REE pattern, trace elements, isotopes, we put forward that the porphyroclastic lava is formed by the remelting of basement metamorphic rocks. The rocks was believed to be formed in the environment similar to volcanics and subvolcanics, and quite different to plutonic rocks due to the features of low-structure of potassium feldspar phenocrysts and solution mechanism, because the porphyroclastic lava phenocrysts occurs as fragments and maybe related to cryptoexplosion. Therefore the rocks was believed to belong to the volcano extrusive facies. (authors)

  4. A geochronological approach for cave evolution in the Cantabrian Coast (Pindal Cave, NW Spain)

    Science.gov (United States)

    Jimenez-Sanchez, M.; Bischoff, J.L.; Stoll, H.; Aranburu, A.

    2006-01-01

    Some of the oldest speleothems in the North Cantabrian Coast (Spain) are reported for the first time in this work. Pindal Cave is developed at 24 m above sea level, in a karstic massif reaching its highest surface in a marine terrace (rasa) located at 50-64 m above the present sea level. Several phases of evolution were previously recognized into the cave, including block collapse of the roof, episodic flooding and detrital sedimentation, and chemical precipitation of at least four speleothem generations over both alluvial and collapse deposits. Three of these speleothem generations have been dated by U/Th. The first generation yielded ages from 124,2 ?? 1, 5 ka BP to 73,1 ?? 0,9 ka BP, giving a minimum age for the main detritic sediments in the cave. The second one is not dated. The third generation gives an age of 3,71 ?? 0,4 ka BP (mathematically corrected to 2.7 ?? 0.5 ka BP), while for the youngest generation, with actively growing stalagmites in the cave, basal ages of 200 years BP are estimated by counting annual laminae. The data suggest a tentative maximum elevation rate close to 0, 2 mm/yr for the Cantabrian Margin in this area, although further chronological studies will be needed to check this hypothesis. ?? 2006 Gebru??der Borntraeger.

  5. Environmental record in detrital cave sediments in the Botovskaya and Dolganskaya Jama caves (Russian Federation)

    Czech Academy of Sciences Publication Activity Database

    Lisá, Lenka; Kadlec, Jaroslav; Chadima, Martin; Hercman, H.; Oberhansli, H.; Osincev, A.

    2006-01-01

    Roč. 8, - (2006), A10273-A10273 ISSN 1029-7006. [European Geosciences Union General Assembly. 02.04.2006-07.04.2006, Vienna] Institutional research plan: CEZ:AV0Z30130516 Keywords : cave * sediments * micromorphology * magnetic properties * provenance Subject RIV: DB - Geology ; Mineralogy http://www.cosis.net/abstracts/EGU06/10273/EGU06-J-10273-2.pdf

  6. The evolutionary history of ferns inferred from 25 low-copy nuclear genes.

    Science.gov (United States)

    Rothfels, Carl J; Li, Fay-Wei; Sigel, Erin M; Huiet, Layne; Larsson, Anders; Burge, Dylan O; Ruhsam, Markus; Deyholos, Michael; Soltis, Douglas E; Stewart, C Neal; Shaw, Shane W; Pokorny, Lisa; Chen, Tao; dePamphilis, Claude; DeGironimo, Lisa; Chen, Li; Wei, Xiaofeng; Sun, Xiao; Korall, Petra; Stevenson, Dennis W; Graham, Sean W; Wong, Gane K-S; Pryer, Kathleen M

    2015-07-01

    • Understanding fern (monilophyte) phylogeny and its evolutionary timescale is critical for broad investigations of the evolution of land plants, and for providing the point of comparison necessary for studying the evolution of the fern sister group, seed plants. Molecular phylogenetic investigations have revolutionized our understanding of fern phylogeny, however, to date, these studies have relied almost exclusively on plastid data.• Here we take a curated phylogenomics approach to infer the first broad fern phylogeny from multiple nuclear loci, by combining broad taxon sampling (73 ferns and 12 outgroup species) with focused character sampling (25 loci comprising 35877 bp), along with rigorous alignment, orthology inference and model selection.• Our phylogeny corroborates some earlier inferences and provides novel insights; in particular, we find strong support for Equisetales as sister to the rest of ferns, Marattiales as sister to leptosporangiate ferns, and Dennstaedtiaceae as sister to the eupolypods. Our divergence-time analyses reveal that divergences among the extant fern orders all occurred prior to ∼200 MYA. Finally, our species-tree inferences are congruent with analyses of concatenated data, but generally with lower support. Those cases where species-tree support values are higher than expected involve relationships that have been supported by smaller plastid datasets, suggesting that deep coalescence may be reducing support from the concatenated nuclear data.• Our study demonstrates the utility of a curated phylogenomics approach to inferring fern phylogeny, and highlights the need to consider underlying data characteristics, along with data quantity, in phylogenetic studies. © 2015 Botanical Society of America, Inc.

  7. Tree fern trunks facilitate seedling regeneration in a productive lowland temperate rain forest.

    Science.gov (United States)

    Gaxiola, Aurora; Burrows, Larry E; Coomes, David A

    2008-03-01

    Seedling regeneration on forest floors is often impaired by competition with established plants. In some lowland temperate rain forests, tree fern trunks provide safe sites on which tree species establish, and grow large enough to take root in the ground and persist. Here we explore the competitive and facilitative effects of two tree fern species, Cyathea smithii and Dicksonia squarrosa, on the epiphytic regeneration of tree species in nutrient-rich alluvial forests in New Zealand. The difficulties that seedlings have in establishing on vertical tree fern trunks were indicated by the following observations. First, seedling abundance was greatest on the oldest sections of tree fern trunks, near the base, suggesting that trunks gradually recruited more and more seedlings over time, but many sections of trunk were devoid of seedlings, indicating the difficulty of establishment on a vertical surface. Second, most seedlings were from small-seeded species, presumably because smaller seeds can easily lodge on tree fern trunks. Deer browsing damage was observed on 73% of epiphytic seedlings growing within 2 m of the ground, whereas few seedlings above that height were browsed. This suggests that tree ferns provide refugia from introduced deer, and may slow the decline in population size of deer-preferred species. We reasoned that tree ferns would compete with epiphytic seedlings for light, because below the tree fern canopy photosynthetically active radiation (PAR) was about 1% of above-canopy PAR. Frond removal almost tripled %PAR on the forest floor, leading to a significant increase in the height growth rate (HGR) of seedlings planted on the forest floor, but having no effects on the HGRs of epiphytic seedlings. Our study shows evidence of direct facilitative interactions by tree ferns during seedling establishment in plant communities associated with nutrient-rich soils.

  8. World Register of marine Cave Species (WoRCS: a new Thematic Species Database for marine and anchialine cave biodiversity

    Directory of Open Access Journals (Sweden)

    Vasilis Gerovasileiou

    2016-09-01

    Full Text Available Scientific exploration of marine cave environments and anchialine ecosystems over recent decades has led to outstanding discoveries of novel taxa, increasing our knowledge of biodiversity. However, biological research on underwater caves has taken place only in a few areas of the world and relevant information remains fragmented in isolated publications and databases. This fragmentation makes assessing the conservation status of marine cave species especially problematic, and this issue should be addressed urgently given the stresses resulting from planned and rampant development in the coastal zone worldwide. The goal of the World Register of marine Cave Species (WoRCS initiative is to create a comprehensive taxonomic and ecological database of known species from marine caves and anchialine systems worldwide and to present this as a Thematic Species Database (TSD of the World Register of marine Species (WoRMS. WoRCS will incorporate ecological data (e.g., type of environment, salinity regimes, and cave zone as well as geographical information on the distribution of species in cave and anchialine environments. Biodiversity data will be progressively assembled from individual database sources at regional, national or local levels, as well as from literature sources (estimate: >20,000 existing records of cave-dwelling species scattered in several databases. Information will be organized in the WoRCS database following a standard glossary based on existing terminology. Cave-related information will be managed by the WoRCS thematic editors with all data dynamically linked to WoRMS and its team of taxonomic editors. In order to mobilize data into global biogeographic databases, a Gazetteer of the Marine and Anchialine Caves of the World will be established. The presence records of species could be eventually georeferenced for submission to the Ocean Biogeographic Information System (OBIS and constitute an important dataset for biogeographical and

  9. Remote sensing evidence of lava-ground ice interactions associated with the Lost Jim Lava Flow, Seward Peninsula, Alaska

    Science.gov (United States)

    Marcucci, Emma C.; Hamilton, Christopher W.; Herrick, Robert R.

    2017-12-01

    Thermokarst terrains develop when ice-bearing permafrost melts and causes the overlying surface to subside or collapse. This process occurs widely throughout Arctic regions due to environmental and climatological factors, but can also be induced by localized melting of ground ice by active lava flows. The Lost Jim Lava Flow (LJLF) on the Seward Peninsula of Alaska provides evidence of former lava-ground ice interactions. Associated geomorphic features, on the scale of meters to tens of meters, were identified using satellite orthoimages and stereo-derived digital terrain models. The flow exhibits positive- and mixed-relief features, including tumuli ( N = 26) and shatter rings ( N = 4), as well as negative-relief features, such as lava tube skylights ( N = 100) and irregularly shaped topographic depressions ( N = 1188) that are interpreted to include lava-rise pits and lava-induced thermokarst terrain. Along the margins of the flow, there are also clusters of small peripheral pits that may be the products of meltwater or steam escape. On Mars, we observed morphologically similar pits near lava flow margins in northeastern Elysium Planitia, which suggests a common formation mechanism. Investigating the LJLF may therefore help to elucidate processes of lava-ground ice interaction on both Earth and Mars.

  10. Mesolithic burial place in La Martina Cave (Dinant, Belgium)

    International Nuclear Information System (INIS)

    Dewez, M.; Gilot, E.; Groessens-Van-Dyck, M.C.; Cordy, J.M.

    1995-01-01

    The ''La Martina'' cave is located near Dinant (Belgium). Although the sediments had been shoveled out in the mid XIXth century, a calcic breccia has provided prehistoric bones. We can distinguish a Pleistocene fauna with cave bear, one Mesolithic burial place with two cromagnoid skeletons, from the 6th millennium BC, and some Holocene faunal remains. (authors). 7 refs

  11. Radon Exposures in the Caves of Tenerife (Canary Islands)

    International Nuclear Information System (INIS)

    Pinza-Molina, C.; Alcaide, J.M.; Rodriques-Bethencourt, R.; Hernandez-Armas, J.

    1999-01-01

    There are more than a hundred volcanic caves and pits of various lengths on the island of Tenerife (Canary Islands, Spain). The results are presented of atmospheric radon measurements performed in three of these caves and in three pits. One of the caves selected, El Viento Cave, is nearly 20 km in length and is the longest volcanic tube in the Canarian archipelago and the second longest in the world. The measurements were performed over two distinct periods during the year using passive polycarbonate detectors. The mean radon concentrations range between 0.3 and 8 kBq.m -3 , the maximum value corresponds to a site located at 1850 m from the mouth of El Viento Cave. Possible touristic development of these caves has been taken into account in estimating the effective doses for visitors and guides (considered separately). The values obtained range from 0.3 to 100 μSv per visit for visitors. The largest effective dose would correspond to that for guides at 41 mSv.y -1 in the Viento Cave. This result would make protection against radiological hazards obligatory if the cave were to be developed as a site for tourism. (author)

  12. Measurements of radon concentrations at caves in Jeju

    Energy Technology Data Exchange (ETDEWEB)

    Go, S. H.; Kang, D. H.; Jung, B. J. [Cheju National University, Cheju (Korea, Republic of)

    2004-07-01

    Radon is a radioactive gas emitting {alpha} particles. It is chemically stable due to its inert characteristic. While its daughter products, {sup 218}Po, {sup 214}Bi, {sup 214}Pb and {sup 214}Po, attached with aerosol particles, is known to cause lung cancer. As radon is produced from uranium and thorium, it accumulates in poorly ventilative underground voids such as caves and mine. Radon concentrations at caves in Jeju were measured in this study. The measurements were made by setting three CR-39 detectors for 70 days at 2 {approx} 4 positions in Manjang, Hyupjae and Ssangyong caves. The radon levels of the caves spread 403.1 . 606.7 Bq/m{sup 3}. With these results, it is concluded that the Jeju caves have 6 times higher radon concentrations than ordinary house of 65.3 Bq/m{sup 3} and that they are higher than Seoul subway stations due to poor ventilation. While, the caves in Jeju have lower radon concentrations than limestone caves of Robin Hood. The radon concentration in the middle of Manjang cave is slightly higher than the action level in the work place of 500 Bq/m{sup 3} suggested by the ICRP. The measurement errors are estimated to be less than 5 % from its calibration factor.

  13. Measurements of radon concentrations at caves in Jeju

    International Nuclear Information System (INIS)

    Go, S. H.; Kang, D. H.; Jung, B. J.

    2004-01-01

    Radon is a radioactive gas emitting α particles. It is chemically stable due to its inert characteristic. While its daughter products, 218 Po, 214 Bi, 214 Pb and 214 Po, attached with aerosol particles, is known to cause lung cancer. As radon is produced from uranium and thorium, it accumulates in poorly ventilative underground voids such as caves and mine. Radon concentrations at caves in Jeju were measured in this study. The measurements were made by setting three CR-39 detectors for 70 days at 2 ∼ 4 positions in Manjang, Hyupjae and Ssangyong caves. The radon levels of the caves spread 403.1 . 606.7 Bq/m 3 . With these results, it is concluded that the Jeju caves have 6 times higher radon concentrations than ordinary house of 65.3 Bq/m 3 and that they are higher than Seoul subway stations due to poor ventilation. While, the caves in Jeju have lower radon concentrations than limestone caves of Robin Hood. The radon concentration in the middle of Manjang cave is slightly higher than the action level in the work place of 500 Bq/m 3 suggested by the ICRP. The measurement errors are estimated to be less than 5 % from its calibration factor

  14. Pollination implications of the diverse diet of tropical nectar-feeding bats roosting in an urban cave.

    Science.gov (United States)

    Lim, Voon-Ching; Ramli, Rosli; Bhassu, Subha; Wilson, John-James

    2018-01-01

    Intense landscaping often alters the plant composition in urban areas. Knowing which plant species that pollinators are visiting in urban areas is necessary for understanding how landscaping impacts biodiversity and associated ecosystem services. The cave nectar bat, Eonycteris spelaea , is an important pollinator for many plants and is often recorded in human-dominated habitats. Previous studies of the diet of E. spelaea relied on morphological identification of pollen grains found in faeces and on the body of bats and by necessity disregarded other forms of digested plant material present in the faeces (i.e., plant juice and remnants). The main objective of this study was to examine the diet of the nectarivorous bat, E. spelaea, roosting in an urban cave at Batu Caves, Peninsular Malaysia by identifying the plant material present in the faeces of bats using DNA metabarcoding. Faeces were collected under the roost of E. spelaea once a week from December 2015 to March 2016. Plant DNA was extracted from the faeces, Polymerase chain reaction (PCR) amplified at ITS2 and rbcL regions and mass sequenced. The resultant plant operational taxonomic units were searched against NCBI GenBank for identification. A total of 55 species of plants were detected from faeces of E. spelaea including Artocarpus heterophyllus, Duabanga grandiflora and Musa spp. which are likely to be important food resources for the cave nectar bat. Many native plant species that had not been reported in previous dietary studies of E. spelaea were detected in this study including Bauhinia strychnoidea and Urophyllum leucophlaeum , suggesting that E. spelaea remains a crucial pollinator for these plants even in highly disturbed habitats. The detection of many introduced plant species in the bat faeces indicates that E. spelaea are exploiting them, particularly Xanthostemon chrysanthus, as food resources in urban area. Commercial food crops were detected from all of the faecal samples, suggesting that E

  15. Pollination implications of the diverse diet of tropical nectar-feeding bats roosting in an urban cave

    Directory of Open Access Journals (Sweden)

    Voon-Ching Lim

    2018-03-01

    Full Text Available Background Intense landscaping often alters the plant composition in urban areas. Knowing which plant species that pollinators are visiting in urban areas is necessary for understanding how landscaping impacts biodiversity and associated ecosystem services. The cave nectar bat, Eonycteris spelaea, is an important pollinator for many plants and is often recorded in human-dominated habitats. Previous studies of the diet of E. spelaea relied on morphological identification of pollen grains found in faeces and on the body of bats and by necessity disregarded other forms of digested plant material present in the faeces (i.e., plant juice and remnants. The main objective of this study was to examine the diet of the nectarivorous bat, E. spelaea, roosting in an urban cave at Batu Caves, Peninsular Malaysia by identifying the plant material present in the faeces of bats using DNA metabarcoding. Methods Faeces were collected under the roost of E. spelaea once a week from December 2015 to March 2016. Plant DNA was extracted from the faeces, Polymerase chain reaction (PCR amplified at ITS2 and rbcL regions and mass sequenced. The resultant plant operational taxonomic units were searched against NCBI GenBank for identification. Results A total of 55 species of plants were detected from faeces of E. spelaea including Artocarpus heterophyllus, Duabanga grandiflora and Musa spp. which are likely to be important food resources for the cave nectar bat. Discussion Many native plant species that had not been reported in previous dietary studies of E. spelaea were detected in this study including Bauhinia strychnoidea and Urophyllum leucophlaeum, suggesting that E. spelaea remains a crucial pollinator for these plants even in highly disturbed habitats. The detection of many introduced plant species in the bat faeces indicates that E. spelaea are exploiting them, particularly Xanthostemon chrysanthus, as food resources in urban area. Commercial food crops were

  16. Moonshot Laboratories' Lava Relief Google Mapping Project

    Science.gov (United States)

    Brennan, B.; Tomita, M.

    2016-12-01

    The Moonshot Laboratories were conceived at the University Laboratory School (ULS) on Oahu, Hawaii as way to develop creative problem solvers able to resourcefully apply 21st century technologies to respond to the problems and needs of their communities. One example of this was involved students from ULS using modern mapping and imaging technologies to assist peers who had been displaced from their own school in Pahoe on the Big Island of Hawaii. During 2015, lava flows from the eruption of Kilauea Volcano were slowly encroaching into the district of Puna in 2015. The lava flow was cutting the main town of Pahoa in half, leaving no safe routes of passage into or out of the town. One elementary school in the path of the flow was closed entirely and a new one was erected north of the flow for students living on that side. Pahoa High School students and teachers living to the north were been forced to leave their school and transfer to Kea'au High School. These students were separated from friends, family and the community they grew up in and were being thrust into a foreign environment that until then had been their local rival. Using Google Mapping technologies, Moonshot Laboratories students created a dynamic map to introduce the incoming Pahoa students to their new school in Kea'au. Elements included a stylized My Maps basemap, YouTube video descriptions of the building, videos recorded by Google Glass showing first person experiences, and immersive images of classrooms were created using 360 cameras. During the first day of orientation at Kea'au for the 200 Pahoa students, each of them were given a tablet to view the map as they toured and got to know their new campus. The methods and technologies, and more importantly innovative thinking, used to create this map have enormous potential for how to educate all students about the world around us, and the issues facing it. http://www.moonshotincubator.com/

  17. THE DIVERSITY OF EPIPHYTIC FERN ON THE OIL PALM TREE (Elaeis guineensis Jacq. IN PEKANBARU, RIAU

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    Nery Sofiyanti

    2015-02-01

    Full Text Available Oil palm (Elaeis guineensis is one  main commodity in Riau Province. Morphologically, the trunk of oil palm  has suitable environment for the growth of epiphytic fern, due to its broaden base of petiole that may accumulate organic and anorganic debrish. The objective of this study was to investigate the diversity of epiphytic fern on the oil palm tree. A total of 125 oil palm trees from seven  study sites in Pekanbaru, Riau were observed. The number of epiphytic ferns identified in this study was 16 species belongs to six families.

  18. Correlation of the Deccan and Rajahmundry Trap lavas: Are these the longest and largest lava flows on Earth?

    Science.gov (United States)

    Self, S.; Jay, A. E.; Widdowson, M.; Keszthelyi, L. P.

    2008-05-01

    We propose that the Rajahmundry Trap lavas, found near the east coast of peninsular India , are remnants of the longest lava flows yet recognized on Earth (˜ 1000 km long). These outlying Deccan-like lavas are shown to belong to the main Deccan Traps. Several previous studies have already suggested this correlation, but have not demonstrated it categorically. The exposed Rajahmundry lavas are interpreted to be the distal parts of two very-large-volume pāhoehoe flow fields, one each from the Ambenali and Mahabaleshwar Formations of the Wai Sub-group in the Deccan Basalt Group. Eruptive conditions required to emplace such long flows are met by plausible values for cooling and eruption rates, and this is shown by applying a model for the formation of inflated pāhoehoe sheet flow lobes. The model predicts flow lobe thicknesses similar to those observed in the Rajahmundry lavas. For the last 400 km of flow, the lava flows were confined to the pre-existing Krishna valley drainage system that existed in the basement beyond the edge of the gradually expanding Deccan lava field, allowing the flows to extend across the subcontinent to the eastern margin where they were emplaced into a littoral and/or shallow marine environment. These lavas and other individual flow fields in the Wai Sub-group may exceed eruptive volumes of 5000 km 3, which would place them amongst the largest magnitude effusive eruptive units yet known. We suggest that the length of flood basalt lava flows on Earth is restricted mainly by the size of land masses and topography. In the case of the Rajahmundry lavas, the flows reached estuaries and the sea, where their advance was perhaps effectively terminated by cooling and/or disruption. However, it is only during large igneous province basaltic volcanism that such huge volumes of lava are erupted in single events, and when the magma supply rate is sufficiently high and maintained to allow the formation of very long lava flows. The Rajahmundry lava

  19. Petrographic and geochemical study on cave pearls from Kanaan Cave (Lebanon

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    Nader Fadi H.

    2007-01-01

    Full Text Available The Kanaan cave is situated at the coastal zone, north of Beirut City (capital of Lebanon. The cave is located within the upper part of the Jurassic Kesrouane Formation (Liassic to Oxfordian which consists mainly of micritic limestone. Twenty seven cave pearls were subjected to petrographic (conventional and scanning electron microscopy and geochemical analyses (major/trace elements and stable isotopes. The cave pearls were found in an agitated splash-pool with low mud content. They are believed to have formed through chemical precipitation of calcite in water over-saturated with calcium. The nucleus and micritic laminae show δ18OV-PDB values of about -5.0‰ and δ13C V-PDB values of -11.8‰, while the surrounding calcite spar laminae resulted in δ18OV-PDB ranging between -5.3 and -5.2‰, and δ13C V-PDB between -12.3 and -12.1‰. A genesis/diagenesis model for these speleothems is proposed involving recrystallization which has selectively affected the inner layers of the cave pearls. This is chiefly invoked by sparry calcite crystals ‘invading’ the inner micrite cortical laminae and the nuclei (cross-cutting the pre-existing mud-envelopes, and the slight depletion in δ18O values from inner to outer cortical layers. The calculated δ18OV-SMOW of the water (-4.2‰ matches with data on meteoric water signature for the central eastern Mediterranean region.

  20. Urang Cave Karst Environmental Development, as Tourism Object

    Directory of Open Access Journals (Sweden)

    Srijono Srijono

    2013-12-01

    Full Text Available Karst environment become an alternative tourist destination as well as to boost local revenues. In karst environments in Grobogan District, Central Java Province, formed Urang Cave, with an interesting endokarst phenomenon. This study aims to do zoning district Urang Cave as tourist sites. The research method is using contour maps as a base map of Urang Cave karst environment geomorphological mapping. Geomorphological data processing is using ArcView GIS 3.3 program. Land use map refers to RBI, scale 1:25.000. Geomorphological analysis refers to the Ministry of Energy and Mineral Resources No. 1456.K/20/MEM/2000, and petrography. Each development zone is analyzed its geophysical environmental element, then set scoring and value summation. For comprehensive environmental element analysis, chemical analysis of rocks, and water-soil chemistry. In reference to Minister of Energy Mineral Resource decrees No. 1456/K/20/MEM/2000, Urang Cave zoning defined into 3 (three zone, as follow: the Protected Zone, Cultivation Zone 1, and Cultivation Zone 2. Protected Zone, consists of Urang Cave tunnel/hallway with a unique spheleothem in it. This zone as a cave tracking site tourism, potential to produce karst water as a decent drinking water while maintaining hardness. Cultivation Zone 1 is spreading about 200 m in distance from outer appearance of spring around the cave hallway. In this zone mining of cave sediments may be done in the inactive form caves, without changing the state of the existing major exokarst morphology. Cultivation Zone 2, an outer zone, located farthest from the tunnel/hallway Urang Cave. Utilization of this zone as a limestone mining quarry, although only on a small scale.

  1. A cave response to environmental changes in the Late Pleistocene: a study of Budimirica Cave sediments, Macedonia

    Czech Academy of Sciences Publication Activity Database

    Temovski, M.; Pruner, Petr; Hercman, H.; Bosák, Pavel

    2016-01-01

    Roč. 69, č. 3 (2016), s. 307-316 ISSN 1330-030X Institutional support: RVO:67985831 Keywords : cave sediments * palaeoenvironmental changes * Late Pleistocene * Budimirica Cave * Macedonia Subject RIV: DB - Geology ; Mineralogy Impact factor: 0.595, year: 2016 http://www. geologia -croatica.hr/ojs/index.php/GC

  2. World Register of marine Cave Species (WoRCS): a new Thematic Species Database for marine and anchialine cave biodiversity

    NARCIS (Netherlands)

    Gerovasileiou, Vasilis; Martínez, Alejandro; Álvarez, Fernando; Boxshall, Geoff; Humphreys, William F.; Jaume, Damià; Becking, L.E.; Muricy, Guilherme; Hengstum, van Peter J.; Dekeyzer, Stefanie; Decock, Wim; Vanhoorne, Bart; Vandepitte, Leen; Bailly, Nicolas; Iliffe, Thomas M.

    2016-01-01

    Scientific exploration of marine cave environments and anchialine ecosystems over recent decades has led to outstanding discoveries of novel taxa, increasing our knowledge of biodiversity. However, biological research on underwater caves has taken place only in a few areas of the world and relevant

  3. Hazard Monitoring of Growing Lava Flow Fields Using Seismic Tremor

    Science.gov (United States)

    Eibl, E. P. S.; Bean, C. J.; Jónsdottir, I.; Hoskuldsson, A.; Thordarson, T.; Coppola, D.; Witt, T.; Walter, T. R.

    2017-12-01

    An effusive eruption in 2014/15 created a 85 km2 large lava flow field in a remote location in the Icelandic highlands. The lava flows did not threaten any settlements or paved roads but they were nevertheless interdisciplinarily monitored in detail. Images from satellites and aircraft, ground based video monitoring, GPS and seismic recordings allowed the monitoring and reconstruction of a detailed time series of the growing lava flow field. While the use of satellite images and probabilistic modelling of lava flows are quite common tools to monitor the current and forecast the future growth direction, here we show that seismic recordings can be of use too. We installed a cluster of seismometers at 15 km from the vents and recorded the ground vibrations associated with the eruption. This seismic tremor was not only generated below the vents, but also at the edges of the growing lava flow field and indicated the parts of the lava flow field that were most actively growing. Whilst the time resolution is in the range of days for satellites, seismic stations easily sample continuously at 100 Hz and could therefore provide a much better resolution and estimate of the lava flow hazard in real-time.

  4. Seven Possible Cave Skylights on Mars

    Science.gov (United States)

    2007-01-01

    [figure removed for brevity, see original site] Figure 1 Seven very dark holes on the north slope of a Martian volcano have been proposed as possible cave skylights, based on day-night temperature patterns suggesting they are openings to subsurface spaces. These six excerpts of images taken in visible-wavelength light by the Thermal Emission Imaging System camera on NASA's Mars Odyssey orbiter show the seven openings. Solar illumination comes from the left in each frame. The volcano is Arsia Mons, at 9 degrees south latitude, 239 degrees east longitude. The features have been given informal names to aid comparative discussion (see figure 1). They range in diameter from about 100 meters (328 feet) to about 225 meters (738 feet). The candidate cave skylights are (A) 'Dena,' (B) 'Chloe,' (C) 'Wendy,' (D) 'Annie,' (E) 'Abby' (left) and 'Nikki,' and (F) 'Jeanne.' Arrows signify north and the direction of illumination. Mars Odyssey is managed by NASA's Jet Propulsion Laboratory, a division of the California Institute of Technology, Pasadena, for NASA's Science Mission Directorate, Washington. Lockheed Martin Space Systems, Denver, is the prime contractor for the project and built the spacecraft. The orbiter's Thermal Emission Imaging System was developed by Arizona State University, Tempe, in collaboration with Raytheon Santa Barbara Remote Sensing, Santa Barbara, Calif., and is operated by Arizona State University.

  5. Tree ferns: monophyletic groups and their relationships as revealed by four protein-coding plastid loci.

    Science.gov (United States)

    Korall, Petra; Pryer, Kathleen M; Metzgar, Jordan S; Schneider, Harald; Conant, David S

    2006-06-01

    Tree ferns are a well-established clade within leptosporangiate ferns. Most of the 600 species (in seven families and 13 genera) are arborescent, but considerable morphological variability exists, spanning the giant scaly tree ferns (Cyatheaceae), the low, erect plants (Plagiogyriaceae), and the diminutive endemics of the Guayana Highlands (Hymenophyllopsidaceae). In this study, we investigate phylogenetic relationships within tree ferns based on analyses of four protein-coding, plastid loci (atpA, atpB, rbcL, and rps4). Our results reveal four well-supported clades, with genera of Dicksoniaceae (sensu ) interspersed among them: (A) (Loxomataceae, (Culcita, Plagiogyriaceae)), (B) (Calochlaena, (Dicksonia, Lophosoriaceae)), (C) Cibotium, and (D) Cyatheaceae, with Hymenophyllopsidaceae nested within. How these four groups are related to one other, to Thyrsopteris, or to Metaxyaceae is weakly supported. Our results show that Dicksoniaceae and Cyatheaceae, as currently recognised, are not monophyletic and new circumscriptions for these families are needed.

  6. Is there foul play in the leaf pocket? The metagenome of floating fern

    NARCIS (Netherlands)

    Dijkhuizen, L.W.; Brouwer, P.; Bolhuis, H.; Reichart, G.-J.; Koppers, N.; Huettel, B.; Bolger, A.M.; Li, F.-W.; Cheng, S.; Liu, X.; Wong, G.K.-S.; Pryer, K.; Weber, A.; Bräutigam, A.; Schluepmann, H.

    2018-01-01

    Dinitrogen fixation by Nostoc azollae residing in specialized leaf pockets supports prolific growth of the floating fern Azolla filiculoides. To evaluate contributions by further microorganisms, the A. filiculoides microbiome and nitrogen metabolism in bacteria persistently associated with Azolla

  7. Using a Microscale Approach to Rapidly Separate and Characterize Three Photosynthetic Pigment Species from Fern

    Science.gov (United States)

    Ayudhya, Theppawut Israsena Na; Posey, Frederick T.; Tyus, Jessica C.; Dingra, Nin N.

    2015-01-01

    A rapid separation of three photosynthetic pigments (chlorophyll "a" and "b" and xanthophyll) from fern ("Polystichum acrostichoides") is described using microscale solvent extraction and traditional thin layer chromatography that minimizes use of harmful chemicals and lengthy procedures. The experiment introduces…

  8. Elevated uptake of Th and U by netted chain fern (Woodwardia areolata)

    International Nuclear Information System (INIS)

    Knox, A.S.; Kaplan, D.I.; Hinton, T.G.

    2008-01-01

    We assessed the ability of netted chain fern (Woodwardia areolata) to uptake U and Th from wetland soils on the U.S. Department of Energy's Savannah River Site in South Carolina. Netted chain fern had the highest Th and U concentrations of all plants collected from the wetland. Ferns grown in contaminated soil (329 mg x kg -1 Th, 44 mg x kg -1 U) in a greenhouse contained 6.4 mg x kg -1 Th and 5.3 mg x kg -1 U compared with 0.13 mg x kg -1 Th and 0.035 mg x kg -1 U in Bermuda grass (Cynodon dactylon). Netted chain fern has potential for the phytoremediation of soils contaminated with Th and U. (author)

  9. Heat transfer measurements of the 1983 kilauea lava flow.

    Science.gov (United States)

    Hardee, H C

    1983-10-07

    Convective heat flow measurements of a basaltic lava flow were made during the 1983 eruption of Kilauea volcano in Hawaii. Eight field measurements of induced natural convection were made, giving heat flux values that ranged from 1.78 to 8.09 kilowatts per square meter at lava temperatures of 1088 and 1128 degrees Celsius, respectively. These field measurements of convective heat flux at subliquidus temperatures agree with previous laboratory measurements in furnace-melted samples of molten lava, and are useful for predicting heat transfer in magma bodies and for estimating heat extraction rates for magma energy.

  10. Under the volcano: phylogeography and evolution of the cave-dwelling Palmorchestia hypogaea (Amphipoda, Crustacea at La Palma (Canary Islands

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    Oromí Pedro

    2008-01-01

    Full Text Available Abstract Background The amphipod crustacean Palmorchestia hypogaea occurs only in La Palma (Canary Islands and is one of the few terrestrial amphipods in the world that have adapted to a strictly troglobitic life in volcanic cave habitats. A surface-dwelling closely related species (Palmorchestia epigaea lives in the humid laurel forest on the same island. Previous studies have suggested that an ancestral littoral Orchestia species colonized the humid forests of La Palma and that subsequent drought episodes in the Canaries reduced the distribution of P. epigaea favouring the colonization of lava tubes through an adaptive shift. This was followed by dispersal via the hypogean crevicular system. Results P. hypogaea and P. epigaea did not form reciprocally monophyletic mitochondrial DNA clades. They showed geographically highly structured and genetically divergent populations with current gene flow limited to geographically close surface locations. Coalescence times using Bayesian estimations assuming a non-correlated relaxed clock with a normal prior distribution of the age of La Palma, together with the lack of association of habitat type with ancestral and recent haplotypes, suggest that their adaptation to cave life is relatively ancient. Conclusion The data gathered here provide evidence for multiple invasions of the volcanic cave systems that have acted as refuges. A re-evaluation of the taxonomic status of the extant species of Palmorchestia is needed, as the division of the two species by habitat and ecology is unnatural. The information obtained here, and that from previous studies on hypogean fauna, shows the importance of factors such as the uncoupling of morphological and genetic evolution, the role of climatic change and regressive evolution as key processes in leading to subterranean biodiversity.

  11. PIXE study on absorption of arsenate and arsenite by arsenic hyperaccumulating fern (Pteris vittata)

    International Nuclear Information System (INIS)

    Yamazaki, H.; Ishii, K.; Matsuyama, S.

    2008-01-01

    Pytoremediation using an arsenic hyperaccumulator, Petris vittata L., has generated an increasing interest worldwide due to both environmentally sound and cost effectiveness. However the mechanism of arsenic accumulation by this fern is not clear at this time. This study examined the uptake of arsenate (As(V)) and arsenite (As(III)) by a hydroponic culture of Pteris vittata using both in-air submilli-PIXE for different parts of the fern and in-air micro-PIXE for the tissue cells. These PIXE analysis systems used 3 MeV proton beams from a 4.5-MV single-ended Dynamitron accelerator at Tohoku University, Japan. The fern took up both arsenate and arsenite from hydroponic solutions which were spiked with 50 mg of arsenic per litter. Final amount of arsenic accumulation in the fern is 1,500 mg per kg (wet weight) of the plant biomass in arsenite treatment and 1,100 mg per kg in arsenate treatment. Arsenic accumulation was not observed at the root parts of the ferns. The in-vivo mapping of elements by submilli-PIXE analyses on the fern laminas showed the arsenic accumulation in the edges of a pinna. The micro-PIXE analyses revealed arsenic maps homogeneously distributed in cells of the lamina, stem and rhizome of the fern. These results indicate that arsenic, both arsenate and arsenite in a contaminated medium are translocated quickly from roots to fronds of Pteris vittata, and distributes homogeneously into tissue cells of the fern laminas. (author)

  12. Global biogeography of scaly tree ferns (Cyatheaceae): evidence for Gondwanan vicariance and limited transoceanic dispersal

    OpenAIRE

    Korall, Petra; Pryer, Kathleen

    2014-01-01

    Aim Scaly tree ferns, Cyatheaceae, are a well-supported group of mostly tree-forming ferns found throughout the tropics, the subtropics and the south-temperate zone. Fossil evidence shows that the lineage originated in the Late Jurassic period. We reconstructed large-scale historical biogeographical patterns of Cyatheaceae and tested the hypothesis that some of the observed distribution patterns are in fact compatible, in time and space, with a vicariance scenario related to the break-up of G...

  13. Fern Stomatal Responses to ABA and CO2 Depend on Species and Growth Conditions.

    Science.gov (United States)

    Hõrak, Hanna; Kollist, Hannes; Merilo, Ebe

    2017-06-01

    Changing atmospheric CO 2 levels, climate, and air humidity affect plant gas exchange that is controlled by stomata, small pores on plant leaves and stems formed by guard cells. Evolution has shaped the morphology and regulatory mechanisms governing stomatal movements to correspond to the needs of various land plant groups over the past 400 million years. Stomata close in response to the plant hormone abscisic acid (ABA), elevated CO 2 concentration, and reduced air humidity. Whether the active regulatory mechanisms that control stomatal closure in response to these stimuli are present already in mosses, the oldest plant group with stomata, or were acquired more recently in angiosperms remains controversial. It has been suggested that the stomata of the basal vascular plants, such as ferns and lycophytes, close solely hydropassively. On the other hand, active stomatal closure in response to ABA and CO 2 was found in several moss, lycophyte, and fern species. Here, we show that the stomata of two temperate fern species respond to ABA and CO 2 and that an active mechanism of stomatal regulation in response to reduced air humidity is present in some ferns. Importantly, fern stomatal responses depend on growth conditions. The data indicate that the stomatal behavior of ferns is more complex than anticipated before, and active stomatal regulation is present in some ferns and has possibly been lost in others. Further analysis that takes into account fern species, life history, evolutionary age, and growth conditions is required to gain insight into the evolution of land plant stomatal responses. © 2017 American Society of Plant Biologists. All Rights Reserved.

  14. Distribution and dynamics of the invasive native hay-scented fern

    Science.gov (United States)

    Songlin Fei; Peter Gould; Melanie Kaeser; Kim. Steiner

    2010-01-01

    The spread and dominance of the invasive native hay-scented fern in the understory is one of the most significant changes to affect the forest ecosystems in the northeastern United States in the last century. We studied changes in the distribution and dynamics of hay-scented fern at a large scale over a 10-yr period in Pennsylvania. The study included 56 stands...

  15. Eogenetic caves in conglomerate: an example from Udin Boršt, Slovenia

    Directory of Open Access Journals (Sweden)

    Lipar Matej

    2011-01-01

    Full Text Available Udin Boršt is a karstified terrace of carbonate rock, which is of fluvioglacial origin, and is situated in the north-western part of Slovenia. There are 15 registered caves, which have been interpreted as caves in conglomerate, while karst of Udin Boršt itself was interpreted as conglomerate karst, shallow karst or isolated karst. In this article, caves in Udin Boršt have been interpreted as eogenetic caves. Based on porosity and bedding material, different types of caves and cave passages have developed. Four general types of eogenetic caves found in Udin Boršt are; linear stream caves, shelter caves, breakdown caves and vadose shafts.

  16. Hardened Lava Meets Wind on Mars

    Science.gov (United States)

    2006-01-01

    NASA's Mars Exploration Rover Spirit used its microscopic imager to capture this spectacular, jagged mini-landscape on a rock called 'GongGong.' Measuring only 3 centimeters (1.2 inches) across, this surface records two of the most important and violent forces in the history of Mars -- volcanoes and wind. GongGong formed billions of years ago in a seething, stirring mass of molten rock. It captured bubbles of gases that were trapped at great depth but had separated from the main body of lava as it rose to the surface. Like taffy being stretched and tumbled, the molten rock was deformed as it moved across an ancient Martian landscape. The tiny bubbles of gas were deformed as well, becoming elongated. When the molten lava solidified, the rock looked like a frozen sponge. Far from finished with its life, the rock then withstood billions of years of pelting by small sand grains carried by Martian dust storms that sometimes blanketed the planet. The sand wore away the surface until, little by little, the delicate strands that enclosed the bubbles of gas were breached and the spiny texture we see today emerged. Even now, wind continues to deposit sand and dust in the holes and crevices of the rock. Similar rocks can be found on Earth where the same complex interplay of volcanoes and weathering occur, whether it be the pelting of rocks by sand grains in the Mojave desert or by ice crystals in the frigid Antarctic. GongGong is one of a group of rocks studied by Spirit and informally named by the Athena Science Team to honor the Chinese New Year (the Year of the Dog). In Chinese mythology, GongGong was the god-king of water in the North Land. When he sacrificed his life to knock down Mount BuZhou, he defeated the bad Emperor in Heaven, freed the sun, moon and stars to go from east to west, and caused all the rivers in China to flow from west to east. Spirit's microscopic imager took this image during on the rover's 736th day, or sol, of exploring Mars (Jan. 28, 2006). The

  17. Management in a neotropical show cave: planning for invertebrates conservation

    Directory of Open Access Journals (Sweden)

    Thais Giovannini Pellegrini

    2012-07-01

    Full Text Available Lapa Nova is a dolomitic cave about 4.5 km long located in northwestern Minas Gerais state, Brazil. The cave experiences intense tourism, concentrated over a single period of the year, during the Feast of Our Lady of Lapa. In order to evaluate the impacts felt by the invertebrate community from this tourism, a new methodology was proposed. Four types of areas (intense visitation area, outlying visitation areas, moderate visitation areas and no-visitation areas were sampled for invertebrates. There was one sampling prior and another on the last day of the 128th feast, to evaluate the effects of visitation on cave-dwelling invertebrates. Results show that invertebrate populations residing in more intensely visited areas of the cave undergo changes in distribution following the event. As a consequence of tourism, invertebrates shift to outlying locations from the visited area, which serve as refuges to the communities. Apparently, the fact that there are places inside Lapa Nova inaccessible to tourists reduces the impact suffered by the invertebrate community, as those sites serve as refuges for cave-dwelling organisms during the pilgrimage. A proper management plan was devised for the tourism/religious use of the cave. It consists basically of delimiting marked pathways for tourists, allowing invertebrates to seek shelter at locations outside visited areas and keeping no-visitation areas off-limits to tourism based on the results of the visitation effects on cave-dwelling invertebrates.

  18. Radon in the Creswell Crags Permian limestone caves

    International Nuclear Information System (INIS)

    Gillmore, G.K.; Phillips, P.S.; Denman, A.R.; Gilbertson, D.D.

    2002-01-01

    An investigation of radon levels in the caves of Creswell Crags, Derbyshire, an important Site of Special Scientific Interest (SSSI) shows that the Lower Magnesian Limestone (Permian) caves have moderate to raised radon gas levels (27-7800 Bq m -3 ) which generally increase with increasing distance into the caves from the entrance regions. This feature is partly explained in terms of cave ventilation and topography. While these levels are generally below the Action Level in the workplace (400 Bq m -3 in the UK), they are above the Action Level for domestic properties (200 Bq m -3 ). Creswell Crags has approximately 40,000 visitors per year and therefore a quantification of effective dose is important for both visitors and guides to the Robin Hood show cave. Due to short exposure times the dose received by visitors is low (0.0016 mSv/visit) and regulations concerning exposure are not contravened. Similarly, the dose received by guides is fairly low (0.4 mSv/annum) due in part to current working practice. However, the risk to researchers entering the more inaccessible areas of the cave system is higher (0.06 mSv/visit). This survey also investigated the effect of seasonal variations on recorded radon concentration. From this work summer to winter ratios of between 1.1 and 9.51 were determined for different locations within the largest cave system

  19. The fungal colonisation of rock-art caves: experimental evidence.

    Science.gov (United States)

    Jurado, Valme; Fernandez-Cortes, Angel; Cuezva, Soledad; Laiz, Leonila; Cañaveras, Juan Carlos; Sanchez-Moral, Sergio; Saiz-Jimenez, Cesareo

    2009-09-01

    The conservation of rock-art paintings in European caves is a matter of increasing interest. This derives from the bacterial colonisation of Altamira Cave, Spain and the recent fungal outbreak of Lascaux Cave, France-both included in the UNESCO World Heritage List. Here, we show direct evidence of a fungal colonisation of rock tablets in a testing system exposed in Altamira Cave. After 2 months, the tablets, previously sterilised, were heavily colonised by fungi and bacteria. Most fungi isolated were labelled as entomopathogens, while the bacteria were those regularly identified in the cave. Rock colonisation was probably promoted by the dissolved organic carbon supplied with the dripping and condensation waters and favoured by the displacement of aerosols towards the interior of the cave, which contributed to the dissemination of microorganisms. The role of arthropods in the dispersal of spores may also help in understanding fungal colonisation. This study evidences the fragility of rock-art caves and demonstrates that microorganisms can easily colonise bare rocks and materials introduced into the cavity.

  20. Radon concentration measurements in the desert caves of Saudi Arabia

    International Nuclear Information System (INIS)

    Al-Mustafa, Hanan; Al-Jarallah, M.I.; Fazal-ur-Rehman; Abu-Jarad, F.

    2005-01-01

    Beneath the harsh deserts of Saudi Arabia lie dark chambers and complex mazes filled with strange shapes and wondrous beauty. Radon concentration measurements have been carried out in the desert caves of Al-Somman Plateau in the Eastern Province of Saudi Arabia. Passive radon dosimeters, based on alpha particle etch track detectors with an inlet filter, were used in this study. A total of 59 dosimeters were placed in five caves for a period of six months. Out of 59 dosimeters, 37 could be collected for analysis. Measurements showed significant variations in radon concentrations in caves depending upon their natural ventilation. The results of the study show that the average radon concentration in the different caves ranges from 74 up to 451Bqm -3 . The average radon concentration in four of the caves was low in the range 74-114Bqm -3 . However, one cave showed an average radon concentration of 451Bqm -3 . Radon is not a problem for tourists in the majority of caves. However, sometimes it may imply some limitation to the working time of guides

  1. Radon concentration measurements in the desert caves of Saudi Arabia

    Energy Technology Data Exchange (ETDEWEB)

    Al-Mustafa, Hanan [Women College, P. O. Box 838, Dammam 31113 (Saudi Arabia); Al-Jarallah, M.I. [Department of Physics, King Fahd University of Petroleum and Minerals, Dhahran 31261 (Saudi Arabia)]. E-mail: mibrahim@kfupm.edu.sa; Fazal-ur-Rehman [Department of Physics, King Fahd University of Petroleum and Minerals, Dhahran 31261 (Saudi Arabia); Abu-Jarad, F. [Radiation Protection Unit, Environmental Protection Department, Saudi Aramco P.O. Box 13027, Dhahran 31311 (Saudi Arabia)

    2005-11-15

    Beneath the harsh deserts of Saudi Arabia lie dark chambers and complex mazes filled with strange shapes and wondrous beauty. Radon concentration measurements have been carried out in the desert caves of Al-Somman Plateau in the Eastern Province of Saudi Arabia. Passive radon dosimeters, based on alpha particle etch track detectors with an inlet filter, were used in this study. A total of 59 dosimeters were placed in five caves for a period of six months. Out of 59 dosimeters, 37 could be collected for analysis. Measurements showed significant variations in radon concentrations in caves depending upon their natural ventilation. The results of the study show that the average radon concentration in the different caves ranges from 74 up to 451Bqm{sup -3}. The average radon concentration in four of the caves was low in the range 74-114Bqm{sup -3}. However, one cave showed an average radon concentration of 451Bqm{sup -3}. Radon is not a problem for tourists in the majority of caves. However, sometimes it may imply some limitation to the working time of guides.

  2. Arsenic accumulation by two brake ferns growing on an arsenic mine and their potential in phytoremediation.

    Science.gov (United States)

    Wei, Chao-Yang; Chen, Tong-Bin

    2006-05-01

    In an area near an arsenic mine in Hunan Province of south China, soils were often found with elevated arsenic levels. A field survey was conducted to determine arsenic accumulation in 8 Cretan brake ferns (Pteris cretica) and 16 Chinese brake ferns (Pteris vittata) growing on these soils. Three factors were evaluated: arsenic concentration in above ground parts (fronds), arsenic bioaccumulation factor (BF; ratio of arsenic in fronds to soil) and arsenic translocation factor (TF; ratio of arsenic in fronds to roots). Arsenic concentrations in the fronds of Chinese brake fern were 3-704 mg kg-1, the BFs were 0.06-7.43 and the TFs were 0.17-3.98, while those in Cretan brake fern were 149-694 mg kg-1, 1.34-6.62 and 1.00-2.61, respectively. Our survey showed that both ferns were capable of arsenic accumulation under field conditions. With most of the arsenic being accumulated in the fronds, these ferns have potential for use in phytoremediation of arsenic contaminated soils.

  3. Taking the Temperature of a Lava Planet

    Science.gov (United States)

    Kreidberg, Laura; Lopez, Eric; Cowan, Nick; Lupu, Roxana; Stevenson, Kevin; Louden, Tom; Malavolta, Luca

    2018-05-01

    Ultra-short period rocky planets (USPs) are an exotic class of planet found around less than 1% of stars. With orbital periods shorter than 24 hours, these worlds are blasted with stellar radiation that is expected to obliterate any traces of a primordial atmosphere and melt the dayside surface into a magma ocean. Observations of USPs have yielded several surprising results, including the measurement of an offset hotspot in the thermal phase curve of 55 Cancri e (which may indicate a thick atmosphere has survived), and a high Bond albedo for Kepler-10b, which suggests the presence of unusually reflective lava on its surface. To further explore the properties of USPs and put these results in context, we propose to observe a thermal phase curve of the newly discovered USP K2- 141b. This planet is a rocky world in a 6.7 hour orbit around a bright, nearby star. When combined with optical phase curve measured by K2, our observations will uniquely determine the planet's Bond albedo, precisely measure the offset of the thermal curve, and determine the temperature of the dayside surface. These results will cement Spitzer's role as a pioneer in the study of terrestrial planets beyond the Solar System, and provide a critical foundation for pursuing the optimal follow-up strategy for K2-141b with JWST.

  4. Uranium-series dating of fossil bones from alpine caves

    International Nuclear Information System (INIS)

    Leitner-Wild, E.; Steffan, I.

    1993-01-01

    During the course of an investigation of fossil cave bear populations the uranium-series method for absolute age determination has been applied to bone material. The applicability of the method to bone samples from alpine caves is demonstrated by the concordance of U/Th and U/Pa ages and cross-checks with the radiocarbon method. Stratigraphic agreement between bone ages and carbonate speleothem ages also indicates the potential of the uranium-series method as a suitable tool for the age determination of fossil bones from alpine cave environments. (Author)

  5. Terraced margins of inflated lava flows on Earth and Mars

    Science.gov (United States)

    Zimbelman, J. R.; Garry, W. B.; Bleacher, J. E.; Crumpler, L. S.

    2011-12-01

    When fluid basaltic lava flows are emplaced over a shallow regional slope (typically much less than one degree), the lava flows often display impressive characteristics of inflation. Here we describe a distinctive marginal characteristic that is often developed along the margins of endogenously inflated basaltic lava flows; discreet topographic levels of the emplaced lava that are here termed 'terraced margins'. Terraced margins were first noted at the distal end of the Carrizozo lava flow in central New Mexico, where they are particularly well expressed, but terraces have also been observed along some margins of the McCartys lava flow (NM), the distal end of the 1859 Mauna Loa lava flow (HI), and lava flows at Craters of the Moon (ID). Differential Global Positioning System surveys across several terraced margins reveal consistent topographic characteristics: the upper surface of each terrace level is at roughly one half the height of the sheet lobe from which it emerges; when a terrace becomes the source of an additional outbreak, the upper surface of the second terrace is at roughly one half the height of the source terrace; often a subtle topographic depression is present along the contact between a terrace and its source sheet lobe, suggesting that the terrace outflow starts at a level roughly one-third the height of the source lobe; the upper surfaces of both the source sheet lobe and associated terraces are level to within tens of centimeters across length scales of many tens to hundreds of meters, indicative of inflation of all components. The field observations will be used as the constraints for modeling of the inflation and terracing mechanisms, an effort that has only recently started. The multiple imaging data sets now available for Mars have revealed the presence of terraced margins on some lava flows on Mars. Although detailed topographic data are not currently available for the Martian examples identified so far, the presence of terraced margins for

  6. The Disruption of Tephra Fall Deposits by Basaltic Lava Flows

    Science.gov (United States)

    Brown, R. J.; Thordarson, T.; Self, S.; Blake, S.

    2010-12-01

    Complex physical and stratigraphic relationships between lava and proximal tephra fall deposits around vents of the Roza Member in the Columbia River Basalt Province, (CRBP), USA, illustrate how basaltic lavas can disrupt, dissect (spatially and temporally) and alter tephra fall deposits. Thin pahoehoe lobes and sheet lobes occur intercalated with tephra deposits and provide evidence for synchronous effusive and explosive activity. Tephra that accumulated on the tops of inflating pahoehoe flows became disrupted by tumuli, which dissected the overlying sheet into a series of mounds. During inflation of subjacent tumuli tephra percolated down into the clefts and rubble at the top of the lava, and in some cases came into contact with lava hot enough to thermally alter it. Lava breakouts from the tumuli intruded up through the overlying tephra deposit and fed pahoehoe flows that spread across the surface of the aggrading tephra fall deposit. Non-welded scoria fall deposits were compacted and welded to a depth of ~50 cm underneath thick sheet lobes. These processes, deduced from the field relationships, have resulted in considerable stratigraphic complexity in proximal regions. We also demonstrate that, when the advance of lava and the fallout of tephra are synchronous, the contacts of some tephra sheets can be diachronous across their extent. The net effect is to reduce the usefulness of pyroclastic deposits in reconstructing eruption dynamics.

  7. ComputerApplications and Virtual Environments (CAVE)

    Science.gov (United States)

    1993-01-01

    Virtual Reality (VR) can provide cost effective methods to design and evaluate components and systems for maintenance and refurbishment operations. The Marshall Space Flight Centerr (MSFC) in Huntsville, Alabama began to utilize VR for design analysis in the X-34 experimental reusable space vehicle. Analysts at MSFC's Computer Applications and Virtual Environments (CAVE) used Head Mounted Displays (HMD) (pictured), spatial trackers and gesture inputs as a means to animate or inhabit a properly sized virtual human model. These models were used in a VR scenario as a way to determine functionality of space and maintenance requirements for the virtual X-34. The primary functions of the virtual X-34 mockup was to support operations development and design analysis for engine removal, the engine compartment and the aft fuselage. This capability provided general visualization support to engineers and designers at MSFC and to the System Design Freeze Review at Orbital Sciences Corporation (OSC). The X-34 program was cancelled in 2001.

  8. Lava delta deformation as a proxy for submarine slope instability

    Science.gov (United States)

    Di Traglia, Federico; Nolesini, Teresa; Solari, Lorenzo; Ciampalini, Andrea; Frodella, William; Steri, Damiano; Allotta, Benedetto; Rindi, Andrea; Marini, Lorenzo; Monni, Niccolò; Galardi, Emanuele; Casagli, Nicola

    2018-04-01

    The instability of lava deltas is a recurrent phenomenon affecting volcanic islands, which can potentially cause secondary events such as littoral explosions (due to interactions between hot lava and seawater) and tsunamis. It has been shown that Interferometric Synthetic Aperture Radar (InSAR) is a powerful technique to forecast the collapse of newly emplaced lava deltas. This work goes further, demonstrating that the monitoring of lava deltas is a successful strategy by which to observe the long-term deformation of subaerial-submarine landslide systems on unstable volcanic flanks. In this paper, displacement measurements derived from Synthetic Aperture Radar (SAR) imagery were used to detect lava delta instability at Stromboli volcano (Italy). Recent flank eruptions (2002-2003, 2007 and 2014) affected the Sciara del Fuoco (SdF) depression, created a "stacked" lava delta, which overlies a pre-existing scar produced by a submarine-subaerial tsunamigenic landslide that occurred on 30 December 2002. Space-borne X-band COSMO-SkyMED (CSK) and C-band SENTINEL-1A (SNT) SAR data collected between February 2010 and October 2016 were processed using the SqueeSAR algorithm. The obtained ground displacement maps revealed the differential ground motion of the lava delta in both CSK and SNT datasets, identifying a stable area (characterized by less than 2 mm/y in both datasets) within the northern sector of the SdF and an unstable area (characterized by velocity fields on the order of 30 mm/y and 160 mm/y in the CSK and SNT datasets, respectively) in the central sector of the SdF. The slope stability of the offshore part of the SdF, as reconstructed based on a recently performed multibeam bathymetric survey, was evaluated using a 3D Limit Equilibrium Method (LEM). In all the simulations, Factor of Safety (F) values between 0.9 and 1.1 always characterized the submarine slope between the coastline and -250 m a.s.l. The critical surfaces for all the search volumes corresponded to

  9. First assessment on the air CO2 dynamic in the show caves of tropical karst, Vietnam

    Directory of Open Access Journals (Sweden)

    Duc A. Trinh

    2018-01-01

    Full Text Available In this study, air, water, and host rock in show caves in a Vietnam’s karst region was monitored and analyzed to identify the ventilation regime and track the cave air CO2 sources. In general, the studied caves are well ventilated. In dynamic – multiple entrance caves, air ventilation is described with the use of U shape model. In static – single entrance cave, air circulation is explained by cold air trap model. Both ventilation models suggest that air is more circulated in winter than in summer. Seasonally, the cave air CO2 increases from early spring to summer. Value in the deepest part of the single-entrance cave is approximately 1,000 ppmv and 8,000 ppmv in early spring and summer, respectively. In multiple-entrance and wet caves, CO2 level is fairly constant all over the show section, increasing from 500 ppmv in early spring to 2,000 ppmv in summer. Data of microclimate, CO2 content, and particularly δ13C show that cave air, particularly in single entrance cave, has higher CO2 concentration during summer due to a stagnation of cave air circulation and an elevated CO2 input from soil and epikarst. The cave air CO2 increase is also observed after intense rainfalls. A factor that increase cave air CO2 in show caves during the festive days could probably be huma n exhaling but the extent of human factor in these studied cave systems should be further investigated. Cave waters including cave pools and streams mediate CO2 level in wet caves. Above all, the atmospheric fraction of CO2 is always dominant (>60% in all cave sections.

  10. Glycoside Hydrolase (GH) 45 and 5 Candidate Cellulases in Aphelenchoides besseyi Isolated from Bird?s-Nest Fern

    OpenAIRE

    Wu, Guan-Long; Kuo, Tzu-Hao; Tsay, Tung-Tsuan; Tsai, Isheng J.; Chen, Peichen J.

    2016-01-01

    Five Aphelenchoides besseyi isolates collected from bird's-nest ferns or rice possess different parasitic capacities in bird's-nest fern. Two different glycoside hydrolase (GH) 45 genes were identified in the fern isolates, and only one was found in the rice isolates. A Abe GH5-1 gene containing an SCP-like family domain was found only in the fern isolates. Abe GH5-1 gene has five introns suggesting a eukaryotic origin. A maximum likelihood phylogeny revealed that Abe GH5-1 is part of the nem...

  11. Survival and growth of epiphytic ferns depend on resource sharing

    Directory of Open Access Journals (Sweden)

    Hua-Zheng eLu

    2016-03-01

    Full Text Available Locally available resources can be shared within clonal plant systems through physiological integration, thus enhancing their survival and growth. Most epiphytes exhibit clonal growth habit, but few studies have tested effects of physiological integration (resource sharing on survival and growth of epiphytes and whether such effects vary with species. We conducted two experiments, one on individuals (single ramets and another on groups (several ramets within a plot, with severed and intact rhizome treatments (without and with physiological integration on two dominant epiphytic ferns (Polypodiodes subamoena and Lepisorus scolopendrium in a subtropical montane moist forest in Southwest China. Rhizome severing (preventing integration significantly reduced ramet survival in the individual experiment and number of surviving ramets in the group experiment, and it also decreased biomass of both species in both experiments. However, the magnitude of such integration effects did not vary significantly between the two species. We conclude that resource sharing may be a general strategy for clonal epiphytes to adapt to forest canopies where resources are limited and heterogeneously distributed in space and time.

  12. Arsenic in the rhizosphere soil solution of ferns.

    Science.gov (United States)

    Wei, Chaoyang; Zheng, Huan; Yu, Jiangping

    2012-12-01

    The aim of this study was to explore the evidence of arsenic hyperaccumulation in plant rhizosphere solutions. Six common fern plants were selected and grown in three types of substrate: arsenic (As) -tailings, As-spiked soil, and soil-As-tailing composites. A rhizobox was designed with an in-situ collection of soil solutions to analyze changes in the As concentration and valence as well as the pH, dissolved organic carbon (DOC) and total nitrogen (TN). Arsenite composed less than 20% of the total As, and As depletion was consistent with N depletion in the rhizosphere solutions of the various treatments. The As concentrations in the rhizosphere and non-rhizosphere solutions in the presence of plants were lower than in the respective controls without plants, except for in the As-spiked soils. The DOC concentrations were invariably higher in the rhizosphere versus non-rhizosphere solutions from the various plants; however, no significant increase in the DOC content was observed in Pteris vittata, in which only a slight decrease in pH appeared in the rhizosphere compared to non-rhizosphere solutions. The results showed that As reduction by plant roots was limited, acidification-induced solubilization was not the mechanism for As hyperaccumulation.

  13. Organochlorine residues in bat guano from nine Mexican caves, 1991

    Science.gov (United States)

    Clark, D.R.; Moreno-Valdez, A.; Mora, M.A.

    1995-01-01

    Samples of bat guano, primarily from Mexican free-tailed bats (Tadarida brasiliensis), were collected at nine bat roosts in caves in northern and eastern Mexico and analysed for organochlorine residues. DDE, the most abundant residue found in each cave, was highest (0.99 p.p.m. dry weight) at Ojuela Cave, Durango. Other studies of DDE in bat guano indicate that this concentration is too low to reflect harmful concentrations in the bats themselves. The DDE at Ojuela may represent either lingering residues from use of DDT years ago in the Ojuela area of perhaps depuration loss from migrant bats with summer maternity roost(s) in a DDE-contaminated area such as Carlsbad Cavern, New Mexico. Presence of o,p-DDT at Tio Bartolo Cave, Nuevo Leon, indicates recent use of DDT, but the concentration of this contaminant was low. Possible impacts on bat colonies of the organophosphorus and carbamate insecticides now in extensive use are unknown.

  14. Urban landscape and architectural workshop Škocjan Caves 2007

    Directory of Open Access Journals (Sweden)

    Andrej Mahovič

    2012-01-01

    Full Text Available The fundamental aim and purpose of the Škocjan Caves 2007 student workshop was for students participating in the seminar of the Faculty of Architecture in Ljubljana to recognise and address some specific spatial and design issues for the further sustainable development of the Škocjan Caves. The working hypothesis of the workshop was that, in spite of significant results in the protection and development of the Caves which have been achieved in the past ten years, it is still necessary to address some spatial issues through an academic process, and to offer different architectural and spatial solutions. The aforesaid is even more important as the Caves are recognised as the only natural and cultural heritage of the Republic of Slovenia to be listed as a UNESCO World Heritage site since 1986.

  15. Center for Advanced Energy Studies: Computer Assisted Virtual Environment (CAVE)

    Data.gov (United States)

    Federal Laboratory Consortium — The laboratory contains a four-walled 3D computer assisted virtual environment - or CAVE TM — that allows scientists and engineers to literally walk into their data...

  16. Luminescence dating at Rose cottage cave: a progress report

    CSIR Research Space (South Africa)

    Woodborne, S

    1997-10-01

    Full Text Available Deal with infrared-stimulated luminescence and thermoluminescence dates from Rose Cottage Cave in South Africa. Discrepancy between luminescence and radiocarbon dates; Concentration of radioactive elements in sediments before and after leaching...

  17. Ancient photosynthetic eukaryote biofilms in an Atacama Desert coastal cave

    Science.gov (United States)

    Azua-Bustos, A.; Gonzalez-Silva, C.; Mancilla, R.A.; Salas, L.; Palma, R.E.; Wynne, J.J.; McKay, C.P.; Vicuna, R.

    2009-01-01

    Caves offer a stable and protected environment from harsh and changing outside prevailing conditions. Hence, they represent an interesting habitat for studying life in extreme environments. Here, we report the presence of a member of the ancient eukaryote red algae Cyanidium group in a coastal cave of the hyperarid Atacama Desert. This microorganism was found to form a seemingly monospecific biofilm growing under extremely low photon flux levels. Our work suggests that this species, Cyanidium sp. Atacama, is a new member of a recently proposed novel monophyletic lineage of mesophilic "cave" Cyanidium sp., distinct from the remaining three other lineages which are all thermo-acidophilic. The cave described in this work may represent an evolutionary island for life in the midst of the Atacama Desert. ?? Springer Science + Business Media, LLC 2009.

  18. Simultaneous caving and surface restoration system for oil shale mining

    Energy Technology Data Exchange (ETDEWEB)

    Allsman, P.T.

    1968-10-01

    A modified caving method is introduced for mining oil shale and simultaneous restoration of the land surface by return of spent shale onto the subsided area. Other methods have been designed to mine the relatively thin richer beds occurring near outcrops in the Piceance Creek Basin of NW. Colorado. Since the discovery of the much thicker beds in the N.-central part of the basin, some attention has focused on in situ and open-pit methods of recovery. Although caving has been recognized as a possible means of mining shale, most people have been skeptical of its success. This stems from the unknown and salient factors of cavability and size of broken rock with caving. Wisdom would seem to dictate that serious evaluation of the caving method be made along with the other methods.

  19. Minerals cave, Volcan Irazu, Costa Rica: description, mineralogy and origin

    International Nuclear Information System (INIS)

    Ulloa, Andres; Campos-Fernandez, Cristian S.; Rojas, Leonardo

    2013-01-01

    Two caves are found in the NW sector of the main crater of the Volcan Irazu at the base of the crown of a glide. The caves are located in an area of structural weakness of the massif developed in sequences of pyroclasts with hydrothermal alteration. Several explorations are organized for the recognition of the caves. The purpose has been of collecting samples and photographs for mineralogical and topographic analyses, through X-ray diffractometry. The minerals present in the samples are compared and identified by means of PDF-2 power x-ray diffraction database of the 2007 ICDD, International Center for Diffraction Data. The origin of the caves has been studied, and it is suggested to carry out other of complement studies [es

  20. The biogeochemistry of anchialine caves: Progress and possibilities

    Science.gov (United States)

    Pohlman, John W.

    2011-01-01

    Recent investigations of anchialine caves and sinkholes have identified complex food webs dependent on detrital and, in some cases, chemosynthetically produced organic matter. Chemosynthetic microbes in anchialine systems obtain energy from reduced compounds produced during organic matter degradation (e.g., sulfide, ammonium, and methane), similar to what occurs in deep ocean cold seeps and mud volcanoes, but distinct from dominant processes operating at hydrothermal vents and sulfurous mineral caves where the primary energy source is mantle derived. This review includes case studies from both anchialine and non-anchialine habitats, where evidence for in situ chemosynthetic production of organic matter and its subsequent transfer to higher trophic level metazoans is documented. The energy sources and pathways identified are synthesized to develop conceptual models for elemental cycles and energy cascades that occur within oligotrophic and eutrophic anchialine caves. Strategies and techniques for testing the hypothesis of chemosynthesis as an active process in anchialine caves are also suggested.

  1. Phylogenetic Relationships of the Fern Cyrtomium falcatum (Dryopteridaceae from Dokdo Island Based on Chloroplast Genome Sequencing

    Directory of Open Access Journals (Sweden)

    Gurusamy Raman

    2016-12-01

    Full Text Available Cyrtomium falcatum is a popular ornamental fern cultivated worldwide. Native to the Korean Peninsula, Japan, and Dokdo Island in the Sea of Japan, it is the only fern present on Dokdo Island. We isolated and characterized the chloroplast (cp genome of C. falcatum, and compared it with those of closely related species. The genes trnV-GAC and trnV-GAU were found to be present within the cp genome of C. falcatum, whereas trnP-GGG and rpl21 were lacking. Moreover, cp genomes of Cyrtomium devexiscapulae and Adiantum capillus-veneris lack trnP-GGG and rpl21, suggesting these are not conserved among angiosperm cp genomes. The deletion of trnR-UCG, trnR-CCG, and trnSeC in the cp genomes of C. falcatum and other eupolypod ferns indicates these genes are restricted to tree ferns, non-core leptosporangiates, and basal ferns. The C. falcatum cp genome also encoded ndhF and rps7, with GUG start codons that were only conserved in polypod ferns, and it shares two significant inversions with other ferns, including a minor inversion of the trnD-GUC region and an approximate 3 kb inversion of the trnG-trnT region. Phylogenetic analyses showed that Equisetum was found to be a sister clade to Psilotales-Ophioglossales with a 100% bootstrap (BS value. The sister relationship between Pteridaceae and eupolypods was also strongly supported by a 100% BS, but Bayesian molecular clock analyses suggested that C. falcatum diversified in the mid-Paleogene period (45.15 ± 4.93 million years ago and might have moved from Eurasia to Dokdo Island.

  2. Exposure of tour guides to Radon at the Cango caves

    International Nuclear Information System (INIS)

    Pule, O.J.; Lindsay, R.

    2008-01-01

    Full text: A study was commissioned by the National Nuclear Regulator of South Africa to investigate the radon levels in the Cango caves and the associated radiological exposure to tour guides. The Cango caves are about 1.3 km in length and are visited by about 300000 tourists per annum due to their natural beauty. This study followed an earlier investigation by the Department of Health which indicated levels in excess of 1000 Bqm -3 which could lead to excess exposure of the cave tour guides. Radon in the various cave chambers and radon exposure of tour guides was measured using RAD-7 continuous radon monitor, electrets ion chambers and personnel monitoring electrets respectively. The measurements in the cave were done during summer and winter seasons to determine any variations between the seasons. The occupancy time for individual guides and equilibrium factor were also investigated. The radon concentration in the cave range from 1000 Bqm -3 to more than 2000 Bqm -3 with equilibrium factor of approximately 0.4, and the variation between winter and summer measurements are insignificant. The radon exposure levels to tour guides differ due to various time periods they spent in the caves. The average dose to tour guides due to radon is 7 mSv -a and the highest exposure is about 10 mSv -a . The exposure to tourists is found to be insignificant due to time they spent in the cave. The regulatory authority is currently investigating what action is necessary to protect the tour guides. (author)

  3. Hurricane Impact on Seepage Water in Larga Cave, Puerto Rico

    Science.gov (United States)

    Vieten, Rolf; Warken, Sophie; Winter, Amos; Schröder-Ritzrau, Andrea; Scholz, Denis; Spötl, Christoph

    2018-03-01

    Hurricane-induced rainfall over Puerto Rico has characteristic δ18O values which are more negative than local rainfall events. Thus, hurricanes may be recorded in speleothems from Larga cave, Puerto Rico, as characteristic oxygen isotope excursions. Samples of 84 local rainfall events between 2012 and 2013 ranged from -6.2 to +0.3‰, whereas nine rainfall samples belonging to a rainband of hurricane Isaac (23-24 August 2012) ranged from -11.8 to -7.1‰. Cave monitoring covered the hurricane season of 2014 and investigated the impact of hurricane rainfall on drip water chemistry. δ18O values were measured in cumulative monthly rainwater samples above the cave. Inside the cave, δ18O values of instantaneous drip water samples were analyzed and drip rates were recorded at six drip sites. Most effective recharge appears to occur during the wet months (April-May and August-November). δ18O values of instantaneous drip water samples ranged from -3.5 to -2.4‰. In April 2014 and April 2015 some drip sites showed more negative δ18O values than the effective rainfall (-2.9‰), implying an influence of hurricane rainfall reaching the cave via stratified seepage flow months to years after the event. Speleothems from these drip sites in Larga cave have a high potential for paleotempestology studies.

  4. Cave-dwelling pholcid spiders (Araneae, Pholcidae: a review

    Directory of Open Access Journals (Sweden)

    Bernhard A. Huber

    2018-06-01

    Full Text Available Pholcidae are ubiquitous spiders in tropical and subtropical caves around the globe, yet very little is known about cave-dwelling pholcids beyond what is provided in taxonomic descriptions and faunistic papers. This paper provides a review based on a literature survey and unpublished information, while pointing out potential biases and promising future projects. A total of 473 native (i.e. non-introduced species of Pholcidae have been collected in about 1000 caves. The large majority of cave-dwelling pholcids are not troglomorphic; a list of 86 troglomorphic species is provided, including 21 eyeless species and 21 species with strongly reduced eyes. Most troglomorphic pholcids are representatives of only two genera: Anopsicus Chamberlin & Ivie, 1938 and Metagonia Simon, 1893. Mexico is by far the richest country in terms of troglomorphic pholcids, followed by several islands and mainland SE Asia. The apparent dominance of Mexico may partly be due to collectors’ and taxonomists’ biases. Most caves harbor only one pholcid species, but 91 caves harbor two and more species (up to five species. Most troglomorphic pholcids belong to two subfamilies (Modisiminae, Pholcinae, very few belong to Smeringopinae and Arteminae, none to Ninetinae. This is in agreement with the recent finding that within Pholcidae, microhabitat changes in general are concentrated in Modisiminae and Pholcinae.

  5. The Influence of Topographic Obstacles on Basaltic Lava Flow Morphologies

    Science.gov (United States)

    von Meerscheidt, H. C.; Brand, B. D.; deWet, A. P.; Bleacher, J. E.; Hamilton, C. W.; Samuels, R.

    2014-12-01

    Smooth pāhoehoe and jagged ´áā represent two end-members of a textural spectrum that reflects the emplacement characteristics of basaltic lava flows. However, many additional textures (e.g., rubbly and slabby pāhoehoe) reflect a range of different process due to lava flow dynamics or interaction with topography. Unfortunately the influence of topography on the distribution of textures in basaltic lava flows is not well-understood. The 18 ± 1.0 ka Twin Craters lava flow in the Zuni-Bandera field (New Mexico, USA) provides an excellent site to study the morphological changes of a lava flow that encountered topographic obstacles. The flow field is 0.2-3.8 km wide with a prominent central tube system that intersects and wraps around a 1000 m long ridge, oriented perpendicular to flow. Upstream of the ridge, the flow has low-relief inflation features extending out and around the ridge. This area includes mildly to heavily disrupted pāhoehoe with interdispersed agglutinated masses, irregularly shaped rubble and lava balls. Breakouts of ´áā and collapse features are also common. These observations suggest crustal disruption due to flow-thickening upstream from the ridge and the movement of lava out and around the obstacle. While the ridge influenced the path of the tube, which wraps around the southern end of the ridge, the series of collapse features and breakouts of ´áā along the tube system are more likely a result of changes in flux throughout the tube system because these features are found both upstream and downstream of the obstacle. This work demonstrates that topography can significantly influence the formation history and surface disruption of a flow field, and in some cases the influence of topography can be separated from the influences of changes in flux along a tube system.

  6. Cave monitoring in the Béke and Baradla caves (Northeastern Hungary: implications for the conditions for the formation cave carbonates

    Directory of Open Access Journals (Sweden)

    György Czuppon

    2018-01-01

    Full Text Available In order to use speleothems in the reconstruction of past climate and environmental changes it is necessary to understand the environmental and hydrological processes that determine the physico-chemical conditions of carbonate precipitation and hence speleothem formation. Therefore, in this study an extended monitoring program was conducted in the Béke and Baradla caves located in the Aggtelek region (Northeastern Hungary. The studied caves are rich in speleothem and flowstone occurrences with great potential for paleoclimatology studies. The monitoring activity included measurements of atmospheric and cave temperatures, CO2 concentration in cave air, as well as chemical and isotopic compositions of water samples (drip water, precipitation and in situ carbonate precipitates. The hydrogen and oxygen isotope compositions of drip waters showed no seasonal variation at any of the collection sites, indicating a well-mixed karstic aquifer. This implies that the isotopic compositions of local speleothems were able to record multiannual isotopic changes inherited from stable isotopes in the drip water. CO2 concentration showed seasonality (high values in summer and low values in winter in both caves, likely affecting carbonate precipitation or corrosion and consequently stalagmite growth. Systematic variations among Mg/Ca and Sr/Ca, Na/Ca, and Si/Ca element ratios were detected in the drip water suggesting Prior Calcite Precipitation (PCP. As PCP is characteristic of periods of reduced infiltration during drier weather conditions, the variations in drip water chemistry and drip rates indicate that the hydrological conditions also varied significantly during the studied period. This hydrological variability appears to affect not only trace element composition but also the isotopic composition of modern carbonate precipitates. In summary, these findings imply that the speleothems from the studied caves were able to record the hydrological changes

  7. rbcL gene sequences provide evidence for the evolutionary lineages of leptosporangiate ferns.

    Science.gov (United States)

    Hasebe, M; Omori, T; Nakazawa, M; Sano, T; Kato, M; Iwatsuki, K

    1994-06-07

    Pteriodophytes have a longer evolutionary history than any other vascular land plant and, therefore, have endured greater loss of phylogenetically informative information. This factor has resulted in substantial disagreements in evaluating characters and, thus, controversy in establishing a stable classification. To compare competing classifications, we obtained DNA sequences of a chloroplast gene. The sequence of 1206 nt of the large subunit of the ribulose-bisphosphate carboxylase gene (rbcL) was determined from 58 species, representing almost all families of leptosporangiate ferns. Phlogenetic trees were inferred by the neighbor-joining and the parsimony methods. The two methods produced almost identical phylogenetic trees that provided insights concerning major general evolutionary trends in the leptosporangiate ferns. Interesting findings were as follows: (i) two morphologically distinct heterosporous water ferns, Marsilea and Salvinia, are sister genera; (ii) the tree ferns (Cyatheaceae, Dicksoniaceae, and Metaxyaceae) are monophyletic; and (iii) polypodioids are distantly related to the gleichenioids in spite of the similarity of their exindusiate soral morphology and are close to the higher indusiate ferns. In addition, the affinities of several "problematic genera" were assessed.

  8. Unique expression of a sporophytic character on the gametophytes of notholaenid ferns (Pteridaceae).

    Science.gov (United States)

    Johnson, Anne K; Rothfels, Carl J; Windham, Michael D; Pryer, Kathleen M

    2012-06-01

    Not all ferns grow in moist, shaded habitats; some lineages thrive in exposed, seasonally dry environments. Notholaenids are a clade of xeric-adapted ferns commonly characterized by the presence of a waxy exudate, called farina, on the undersides of their leaves. Although some other lineages of cheilanthoid ferns also have farinose sporophytes, previous studies suggested that notholaenids are unique in also producing farina on their gametophytes. For this reason, consistent farina expression across life cycle phases has been proposed as a potential synapomorphy for the genus Notholaena. Recent phylogenetic studies have shown two species with nonfarinose sporophytes to be nested within Notholaena, with a third nonfarinose species well supported as sister to all other notholaenids. This finding raises the question: are the gametophytes of these three species farinose like those of their close relatives, or are they glabrous, consistent with their sporophytes? We sowed spores of a diversity of cheilanthoid ferns onto culture media to observe and document whether their gametophytes produced farina. To place these species within a phylogenetic context, we extracted genomic DNA, then amplified and sequenced three plastid loci. The aligned data were analyzed using maximum likelihood to generate a phylogenetic tree. Here we show that notholaenids lacking sporophytic farina also lack farina in the gametophytic phase, and notholaenids with sporophytic farina always display gametophytic farina (with a single exception). Outgroup taxa never displayed gametophytic farina, regardless of whether they displayed farina on their sporophytes. Notholaenids are unique among ferns in consistently expressing farina across both phases of the life cycle.

  9. Restoring Native Forest Understory: The Influence of Ferns and Light in a Hawaiian Experiment

    Directory of Open Access Journals (Sweden)

    Robert Shallenberger

    2013-03-01

    Full Text Available Ecological restoration is an increasingly important component of sustainable land management. We explore potential facilitative relationships for enhancing the cost-effectiveness of restoring native forest understory, focusing on two factors: (1 overstory shade and (2 possible facilitation by a fern (Dryopteris wallichiana, one of few native colonists of pasture in our montane Hawaiˈi study system. We planted 720 understory tree seedlings and over 4000 seeds of six species under six planting treatments: a full factorial combination of low, medium and high light, situating plantings in either the presence or absence of a mature fern. After three years, 75% of outplanted seedlings survived. Seedling survivorship was significantly higher in the presence of a fern (79% vs. 71% without a fern and in medium and low light conditions (81% vs. 64% in high light. Relative height was highest at low to medium light levels. After 2.2 years, 2.8% of the planted seeds germinated. We observed no significant differences in seed germination relative to light level or fern presence. Analyzing several approaches, we found nursery germination of seeds followed by outplanting ca. 20% less costly than direct seeding in the field. This study opens new questions about facilitation mechanisms that have the potential to increase the extent and effectiveness of restoration efforts.

  10. Knowledge-Based Estimation of Edible Fern Harvesting Sites in Mountainous Communities of Northeastern Japan

    Directory of Open Access Journals (Sweden)

    Toshiya Matsuura

    2013-12-01

    Full Text Available Once local expert knowledge regarding the harvesting of various non-timber forest products (NTFPs is lost, it is difficult to recover. We investigated whether the knowledge of expert forest harvesters can be used to determine the habitat distribution and harvesting sites of three popular edible wild ferns, i.e., ostrich fern (Matteuccia struthiopteris, bracken (Pteridium aquilinum, and royal fern (Osmunda japonica, in mountainous communities of western Fukushima, Japan. Using multi-criteria evaluation (MCE based on the analytic hierarchy process (AHP and geographic information system (GIS, we found that harvesters were easily able to recognize differences in the spatial characteristics of the habitat distribution of fern species due to both natural and anthropogenic factors. These factors were described by various GIS layers, such as vegetation and terrain features (e.g., gradient, aspect, and slope position derived from a 20-m digital elevation model (DEM. Harvesting sites were limited by their distance from a roadway, which differed among species. By comparison with the GPS records of actual harvesting sites, we estimated the potential harvesting sites of each fern species with reasonable accuracy, particularly for bracken. Our results show that the knowledge of expert forest harvesters can be quantified using MCE and GIS, which is useful for determining the spatial characteristics of NTFP harvesting and ensuring sustainable management practices.

  11. Morphometry and distribution of isolated caves as a guide for phreatic and confined paleohydrological conditions

    Science.gov (United States)

    Frumkin, Amos; Fischhendler, Itay

    2005-04-01

    Isolated caves are a special cave type common in most karst terrains, formed by prolonged slow water flow where aggressivity is locally boosted. The morphometry and distribution of isolated caves are used here to reconstruct the paleohydrology of a karstic mountain range. Within a homogenous karstic rock sequence, two main types of isolated caves are distinguished, and each is associated with a special hydrogeologic setting: maze caves form by rising water in the confined zone of the aquifer, under the Mt. Scopus Group (Israel) confinement, while chamber caves are formed in phreatic conditions, apparently by lateral flow mixing with a vadose input from above.

  12. The WP-CAVE concept for an underground high-level nuclear waste repository

    International Nuclear Information System (INIS)

    Pettersson, G.

    1984-02-01

    A central, nearly spherical cave, of diameter 40 m is excavated in rock and the waste fuel is placed in it. The fuel canisters are placed in cylindrical holes in large concrete balls which are stored at the bottom of a central stack in the cave. Other empty balls fill the rest of the cave. By natural convection, the heat is evenly distributed in the cave and the surrounding central rock body. A clay barrier, which completely surrounds the rock body, prevents ground water circulation for a very long time and protects the cave against tectonic movements. The cave can store approx. 350 tons of fuel after the 10 years of intermediate storage

  13. A Mechanism for Stratifying Lava Flows

    Science.gov (United States)

    Rice, A.

    2005-12-01

    Relict lava flows (e.g., komatiites) are often reported to be zoned in the vertical, each zone separated by a sharp contact. Such stratifications in igneous flows, both intrusive and extrusive, can be treated as analogues of suspended loads of sediments in rivers and streams, and hence amenable to quantitative treatment derived for the hydraulic environment as long as dynamic similitude is assured. Situations typically encountered in the hydraulic environment are streams carrying a bed load at the bottom of the stream, the bed load separated by a sharp horizon from a sediment load carried above it. This sediment load may be topped by others of decreasing density as one moves to the surface of the flow, with perhaps the uppermost layer clear of any suspended matter. Rules exist for estimating the thickness D of these loads: one of them is given by D ~ 4.4V3/rgcvs where V is the shear velocity or average velocity of the flow, r = (ρs - ρl)/ρl where ρs is the density of the suspended solid matter, ρl the density of the fluid, g the acceleration of gravity, c the concentration of the particulate content and vs the settling velocity. The settling velocity is secured through Stoke's Law and the velocity of the flow is given by V = R2/3S1/2/n where R is the hydraulic radius, S the gradient along which the fluid flows and n is the Manning Coefficient. In the igneous case, the bed load would be composed of primocrysts, i.e., of the first crystals to come out of solution as the flow cools along its run. This would leave the upper portions of the flow more evolved except perhaps for a quenched crust riding atop the flow. As the viscosity of the flow is dependent not only on temperature but on composition and crystal content, the mean velocity of each layer will be different from the layer above and below it. This requires shear at the interface of adjoining stratifications, which brings into play another mechanism: dispersive pressure (the Bagnold effect). Dispersive

  14. Petrogenesis of basalt-trachyte lavas from Olmoti Crater, Tanzania

    Science.gov (United States)

    Mollel, Godwin F.; Swisher, Carl C., III; McHenry, Lindsay J.; Feigenson, Mark D.; Carr, Michael J.

    2009-08-01

    Olmoti Crater is part of the Plio-Pleistocene Ngorongoro Volcanic Highland (NVH) in northern Tanzania to the south of Gregory Rift. The Gregory Rift is part of the eastern branch of the East African Rift System (EARS) that stretches some 4000 km from the Read Sea and Gulf of Aden in the north to the Zambezi River in Mozambique. Here, we (1) characterize the chemistry and mineral compositions of lavas from Olmoti Crater, (2) determine the age and duration of Olmoti volcanic activity through 40Ar/ 39Ar dating of Olmoti Crater wall lavas and (3) determine the genesis of Olmoti lavas and the relationship to other NVH and EARS volcanics and (4) their correlation with volcanics in the Olduvai and Laetoli stratigraphic sequences. Olmoti lavas collected from the lower part of the exposed crater wall section (OLS) range from basalt to trachyandesite whereas the upper part of the section (OUS) is trachytic. Petrography and major and trace element data reflect a very low degree partial melt origin for the Olmoti lavas, presumably of peridotite, followed by extensive fractionation. The 87Sr/ 86Sr data overlap whereas Nd and Pb isotope data are distinct between OLS and OUS samples. Interpretation of the isotope data suggests mixing of enriched mantle (EM I) with high-μ-like reservoirs, consistent with the model of Bell and Blenkinsop [Bell, K., Blenkinsop, J., 1987. Nd and Sr isotopic compositions of East African carbonatites: implications for mantle heterogeneity. Geology 5, 99-102] for East African carbonatite lavas. The isotope ratios are within the range of values defined by Oceanic Island Basalt (OIB) globally and moderate normalized Tb/Yb ratios (2.3-1.6) in these lavas suggest melting in the lithospheric mantle consistent with other studies in the region. 40Ar/ 39Ar incremental-heating analyses of matrix and anorthoclase separates from Olmoti OLS and OUS lavas indicate that volcanic activity was short in duration, lasting ˜200 kyr from 2.01 ± 0.03 Ma to 1.80 ± 0

  15. Analogue experiments as benchmarks for models of lava flow emplacement

    Science.gov (United States)

    Garel, F.; Kaminski, E. C.; Tait, S.; Limare, A.

    2013-12-01

    During an effusive volcanic eruption, the crisis management is mainly based on the prediction of lava flow advance and its velocity. The spreading of a lava flow, seen as a gravity current, depends on its "effective rheology" and on the effusion rate. Fast-computing models have arisen in the past decade in order to predict in near real time lava flow path and rate of advance. This type of model, crucial to mitigate volcanic hazards and organize potential evacuation, has been mainly compared a posteriori to real cases of emplaced lava flows. The input parameters of such simulations applied to natural eruptions, especially effusion rate and topography, are often not known precisely, and are difficult to evaluate after the eruption. It is therefore not straightforward to identify the causes of discrepancies between model outputs and observed lava emplacement, whereas the comparison of models with controlled laboratory experiments appears easier. The challenge for numerical simulations of lava flow emplacement is to model the simultaneous advance and thermal structure of viscous lava flows. To provide original constraints later to be used in benchmark numerical simulations, we have performed lab-scale experiments investigating the cooling of isoviscous gravity currents. The simplest experimental set-up is as follows: silicone oil, whose viscosity, around 5 Pa.s, varies less than a factor of 2 in the temperature range studied, is injected from a point source onto a horizontal plate and spreads axisymmetrically. The oil is injected hot, and progressively cools down to ambient temperature away from the source. Once the flow is developed, it presents a stationary radial thermal structure whose characteristics depend on the input flow rate. In addition to the experimental observations, we have developed in Garel et al., JGR, 2012 a theoretical model confirming the relationship between supply rate, flow advance and stationary surface thermal structure. We also provide

  16. The Influence of Slope Breaks on Lava Flow Surface Disruption

    Science.gov (United States)

    Glaze, Lori S.; Baloga, Stephen M.; Fagents, Sarah A.; Wright, Robert

    2014-01-01

    Changes in the underlying slope of a lava flow impart a significant fraction of rotational energy beyond the slope break. The eddies, circulation and vortices caused by this rotational energy can disrupt the flow surface, having a significant impact on heat loss and thus the distance the flow can travel. A basic mechanics model is used to compute the rotational energy caused by a slope change. The gain in rotational energy is deposited into an eddy of radius R whose energy is dissipated as it travels downstream. A model of eddy friction with the ambient lava is used to compute the time-rate of energy dissipation. The key parameter of the dissipation rate is shown to be rho R(sup 2/)mu, where ? is the lava density and mu is the viscosity, which can vary by orders of magnitude for different flows. The potential spatial disruption of the lava flow surface is investigated by introducing steady-state models for the main flow beyond the steepening slope break. One model applies to slow-moving flows with both gravity and pressure as the driving forces. The other model applies to fast-moving, low-viscosity, turbulent flows. These models provide the flow velocity that establishes the downstream transport distance of disrupting eddies before they dissipate. The potential influence of slope breaks is discussed in connection with field studies of lava flows from the 1801 Hualalai and 1823 Keaiwa Kilauea, Hawaii, and 2004 Etna eruptions.

  17. Lead accumulation in the aquatic fern Azolla filiculoides.

    Science.gov (United States)

    Oren Benaroya, Rony; Tzin, Vered; Tel-Or, Elisha; Zamski, Eli

    2004-01-01

    In this study, we characterized lead (Pb2+) accumulation and storage by the aquatic fern Azolla filiculoides. Lead precipitates were detected in the vacuoles of mesophyll cells of Azolla plants cultured for 6 d in rich growth medium containing 20 mg l(-1) Pb2+. Energy dispersive spectroscopy (EDS) analysis of the relative element content of leaves collected from these plants revealed a 100% increase in the levels of P, S, Na and Ca and a 40% decrease in Mg and Cl compared to the untreated plants. Both Azolla whole plants and isolated apoplasts were incubated for 6 d in 20 mg l(-1) Pb2+. Lead content in the whole plant composed 0.37%, 2.3% and 1.8% of the dry weight after 2, 4 and 6 d of growth, respectively, while the isolated Azolla apoplast contained 0.125%, 1.22% and 1.4% Pb2+, respectively. Lead content in Azolla whole plant increase by 200%, 100% and 22% after 2, 4 and 6 d of growth, respectively, when compared to Azolla apoplast. Dark, electron dense deposits of lead were observed in light and transmission electron microscope in leaf cells treated with lead. All the observed lead deposits were localized in vacuoles while larger lead deposits were found in mature leaves than in young leaves. No lead deposits were found in cells of the cyanobiont Anabaena when the plants were exposed to similar conditions. Activity and content of V-H+-ATPase were studied in Azolla plants grown in the presence of 20, 40 and 80 mg l(-1) of lead for a period of 4 d. Activity of V-H+-ATPase was increased by 190%, 210% and 220%, respectively, but the content of V-H+-ATPase was reduced by all lead concentrations. Copyright 2004 Elsevier SAS

  18. Gametophyte and sporophyte of tree ferns in vitro culture

    Directory of Open Access Journals (Sweden)

    Katarzyna Goller

    2011-01-01

    Full Text Available Experiments had been carried out on gametophytes and very young fronds of sporophytes with application of Murashige and Skoog (1962 medium. The paper described the results of 15 years in vitro experiments on 16 species of tree ferns belonging to various genera: Blechnum, Cibotium, Cyathea and Dicksonia. Genus Cyathea was represented by: C. australis (R.Br. Domin., C. capensis (L.f. Sm., C. cooperi (F.Muell. Domin, C. brownii Domin, C. dealbata (G.Forest Sw., C. dregei Kunze, C. leichhardtiana (F.Muell. Copel., C. robertsiana (F.Muell. Domin., C. schanschin Mart., C. smithii Hook.f. and Cyathea sp. In case of genus Dicksonia only two species were introduced into our experiments: D. fibrosa Colenso and D. sellowiana Hook.. Taxa Blechnum was presented by B. brasiliense Desv. and Cibotium by C. glaucum (Sm. Hook. and Arn. and C. schiedei Schltdl. and Cham.. The studied species presented various responses on culture conditions depending on the level of stage of development. Time required for spores germination differed between species and took from only a few to 16 weeks. Prothalium formations showed various types of growth presented by marginal meristems. For all investigated species long term gametophyte in vitro cultures was established. Mature gametophyte possessed functional antheridia and archegonia. Spontaneous fertilization helped to establish the culture of young sporophytes. For all species the ex vitro culture in greenhouse collection was established. Manipulation of sucrose content in the medium stimulated the multiplication of gametophytes, but its lack induced formation of gemmae. Apospory was observed when culture of very young fronds was extended for 6 months and new generation of gametophytes was developed. Finally, sporophytes of 12 species were obtained and they have been growing in our greenhouse.

  19. Cure from the cave: volcanic cave actinomycetes and their potential in drug discovery

    Directory of Open Access Journals (Sweden)

    Cheeptham N.

    2013-01-01

    Full Text Available Volcanic caves have been little studied for their potential as sources of novel microbial species and bioactive compounds with new scaffolds. We present the f irst study of volcanic cave microbiology from Canada and suggest that this habitat has great potential for the isolation of novel bioactive substances. Sample locat ions were plot ted on a contour map that was compiled in ArcView 3.2. Over 400 bacterial isolates were obtained from the Helmcken Falls cave in Wells Gray Provincial Park, British Columbia. From our preliminary screen, of 400 isolates tested, 1% showed activity against extended spectrum ß-lactamase E. coli, 1.75% against Escherichia coli, 2.25% against Acinetobacter baumannii, and 26.50% against Klebsiella pneumoniae. In addition, 10.25% showed activity against Micrococcus luteus, 2% against methicillin resistant Staphylococcus aureus, 9.25% against Mycobacterium smegmatis, 6.25% Pseudomonas aeruginosa and 7.5% against Candida albicans. Chemical and physical characteristics of three rock wall samples were studied using scanning electron microscopy and f lame atomic absorption spectrometry. Calcium (Ca, iron (Fe, and aluminum (Al were the most abundant components while magnesium (Mg, sodium (Na, arsenic (As, lead (Pb, chromium (Cr, and barium (Ba were second most abundant with cadmium (Cd and potassium (K were the least abundant in our samples. Scanning electron microscopy (SEM showed the presence of microscopic life forms in all three rock wall samples. 16S rRNA gene sequencing of 82 isolates revealed that 65 (79.3% of the strains belong to the Streptomyces genus and 5 (6.1% were members of Bacillus, Pseudomonas, Nocardia and Erwinia genera. Interestingly, twelve (14.6% of the 16S rRNA sequences showed similarity to unidentif ied ribosomal RNA sequences in the library databases, the sequences of these isolates need to be further investigated using the EzTaxon-e database (http://eztaxon-e. ezbiocloud.net/ to determine whether

  20. Morphology and evolution of sulphuric acid caves in South Italy

    Science.gov (United States)

    D'Angeli, Ilenia M.; De Waele, Jo; Galdenzi, Sandro; Madonia, Giuliana; Parise, Mario; Vattano, Marco

    2016-04-01

    Sulphuric acid speleogenesis (SAS) related to the upwelling of acid water enriched in H2S and CO2 represents an unusual way of cave development. Since meteoric infiltration waters are not necessarily involved in speleogenesis, caves can form without the typical associated karst expressions (i.e. dolines) at the surface. The main mechanism of sulphuric acid dissolution is the oxidation of H2S (Jones et al., 2015) which can be amplified by bacterial mediation (Engel et al., 2004). In these conditions, carbonate dissolution associated with gypsum replacement, is generally believed to be faster than the normal epigenic one (De Waele et al., 2016). In Italy several SAS caves have been identified, but only few systems have been studied in detail: Frasassi and Acquasanta Terme (Marche)(Galdenzi et al., 2010), Monte Cucco (Umbria) (Galdenzi & Menichetti, 1995), and Montecchio (Tuscany) (Piccini et al., 2015). Other preliminary studies have been carried out in Calabria (Galdenzi, 2007) and Sicily (De Waele et al., 2016). Several less studied SAS cave systems located in South Italy, and in particular in Apulia (Santa Cesarea Terme), Sicily (Acqua Fitusa, Acqua Mintina) and Calabria (Mt. Sellaro and Cassano allo Ionio) have been selected in the framework of a PhD thesis on SAS caves and their speleogenesis. Using both limestone tablet weight loss (Galdenzi et al., 2012) and micro erosion meter (MEM) (Furlani et al., 2010) methods the dissolution rate above and under water in the caves will be quantified. Geomorphological observations, landscape analysis using GIS tools, and the analysis of gypsum and other secondary minerals (alunite and jarosite) (stable isotopes and dating) will help to reconstruct the speleogenetic stages of cave formation. Preliminary microbiological analysis will determine the microbial diversity and ecology in the biofilms. References Engel S.A., Stern L.A., Bennett P.C., 2004 - Microbial contributions to cave formation: New insight into sulfuric acid

  1. Spotted hyena and steppe lion predation behaviours on cave bears of Europe - ?Late Quaternary cave bear extinction as result of predator stress

    Science.gov (United States)

    Diedrich, Cajus G.

    2010-05-01

    Cave bears hibernated in caves all over Eurasia (e.g. Rabeder et al., 2000) including alpine regions using mainly larger caves for this purpose. Late Quaternary spotted hyenas Crocuta crocuta spelaea instead occupied mainly areas close to the cave entrances as their dens (Diedrich and Žák 2006, Diedrich 2010). The largest predator, the steppe lion Panthera leo spelaea was only a sporadic cave dweller (Diedrich 2007b, 2009b). His presence and its remains from caves all over Europe can be recently explained best as result of imported carcasses after killing by their largest antagonists, the Late Quaternary spotted hyenas. In some cases the kill might have happened in the hyena den cave itself during the theft of prey remains by lions (Diedrich 2009a). Another reason of their remains in caves of Europe is the hunting onto the herbivorous cave bears, especially during hibernation times, when megafauna prey was less available in the open environments (Diedrich 2009c). These lion remains from caves of Europe, nearly all of which were from adult animals, provide evidence of active predation by lions onto cave bears even in medium high alpine regions (Diedrich 2009b, in review). Lion skeletons in European cave bear dens were therefore often found amongst originally articulated cave bear skeletons or scattered cave bear remains and even close to their hibernation nests (Diedrich et al. 2009c, in review). Not only lions fed on cave bears documented mainly by the large quantities of chewed, punctured and crushed cave bear long-bones; even damaged skulls reveal that hyenas scavenged primarily on cave bear carcasses which were mainly responsible for the destruction of their carcasses and bones (Diedrich 2005, 2009d). Predation and scavenging on cave bears by the two largest Late Quaternary predators C. c. spelaea and P. l. spelaea explains well the large quantity of fragmented cave bear bones over all European caves in low to medium high mountainous elevations, whereas in

  2. Radon measurements using track detector in Wadi Sannur cave

    International Nuclear Information System (INIS)

    Shahin, F.; Eissa, M.F.; Mostafa, R.M.

    2008-01-01

    The most important contributors to the committed effective dose received by population due to natural sources are the short-lived decay products of radon ( 222 Rn). In natural voids, such as caves, most radon will enter the system from diffusion across the rock-air interface. It is well known that factors such as air pressure and temperature control the distribution of radon. The radon concentration measurements in the Wadi Sannur cave in Beni-Suef governorate in the period from 14 th of July 2005 to 17 th of October 2005. The average radon concentrations in the right, left and whole first cave are 916.12 ± 179.09, 819.63 ± 54.72 and 873.90± 147.11 Bq m -3 respectively. The measurements were performed using track etch detector of type Cr-39. After exposure, all detectors were etched chemically in 6.25 M NaOH solution at 70 C degrees for 6 h. The tracks were counted with an optical microscope magnifying 400 times. The average temperature inside the first cave during the period of measurements is 25-26 C degrees. The annual effective doses for the workers and visitors in the cave have been calculated. The average radon concentration in the Wadi Sannur cave, was 873.90 ± 147.11 Bq m -3 . The annual effective doses for worker and visitor in the cave were 1.33 ± 0.24 and 0.041 ± 0.007 mSv respectively. The doses are within the international recommended dose of 1.15 mSv. (author)

  3. WP-Cave - assessment of feasibility, safety and development potential

    International Nuclear Information System (INIS)

    1989-09-01

    According to SKB R and D-programme 1986, alternative disposal methods will be investigated to provide a basis for selecting a site and a repository system for the Swedish spent nuclear fuel. The present report is a comparison between the WP-Cave and the reference concept KBS-3. The comparison has resulted in the following conclusions: - Both concepts are judged to be able to provide adequate safety. - A utilization of the potential of the WP-Cave requires, however, extensive development in areas where the current state of knowledge and available data are incomplete. - The higher temperatures in the WP-Cave lead to greater uncertainty as to long-term performance. Reducing this uncertainty would require many yaers of research and substantial resources. - Both repositories, including the barriers they incorporate, could be built with a normal adaption of available technology. -It is not possible to say today whether it would be simpler to find suitable sites for one design or the other. - The WP-Cave is considerably more expensive. A future research direction based on a concentrated emplacement of spent fuel along the lines of the WP-Cave is therefore judged to entail greater uncertainty as regards the possibilities of achieving acceptable safety and to require greater resources for research and development, at the same time as the costs of building the repository would be higher. The studies of the WP-Cave as an integral system should therfore be discontinued. Certain barrier designs in the WP-Cave could also be utulized in repository designs with lower temperature, for example the reduction potential of the steel canisters and the hydraulic cage's diversion of groundwater. Studies within these areas are being conducted within SKB and should continue

  4. Analysis of the Condition and Development Opportunities of Cave Tourism in Primorsko-Goranska County

    Directory of Open Access Journals (Sweden)

    Rade Knežević

    2011-04-01

    Full Text Available This paper examines caves as morphological factors of tourism development in Primorsko-Goranska County. The primary aim is to promote cave heritage parallel with the development of cave tourism. The methodological framework is based on analyzing material in the archives of tourist boards, taking an inventory of cave resources (case study, conducting interviews, and making a SWOT analysis of cave tourism development. Research results show that caves represent a complex resource in the tourist trade of Primorsko-Goranska County, to which little importance has been attached up to date (priority being given to mass tourism. In the context of tourism development, caves fall into four groups. The first group comprises caves that have a long tradition, but show signs of aging as a tourism product. The second group includes caves that were once open to the public, but were later closed for some reason. The third group consists of caves on the seaside or seabed visited by speleologists-divers. The fourth group is made up of potential cave sites that have been evaluated as being attractive, but is located in hard-to-access areas, making their valorisation in tourism purpose more difficult. Only Lokvarka Cave has a substantial influence on the development of special interest tourism in Lokve, while in all other cases, the role of caves in the tourism offering needs to be revalorised.

  5. Numerical simulation of formation and preservation of Ningwu ice cave, Shanxi, China

    Science.gov (United States)

    Yang, S.; Shi, Y.

    2015-10-01

    Ice caves exist in locations where annual average air temperature is higher than 0 °C. An example is Ningwu ice cave, Shanxi Province, the largest ice cave in China. In order to quantitatively investigate the mechanism of formation and preservation of the ice cave, we use the finite-element method to simulate the heat transfer process at this ice cave. There are two major control factors. First, there is the seasonal asymmetric heat transfer. Heat is transferred into the ice cave from outside very inefficiently by conduction in spring, summer and fall. In winter, thermal convection occurs that transfers heat very efficiently out of the ice cave, thus cooling it down. Secondly, ice-water phase change provides a heat barrier for heat transfer into the cave in summer. The calculation also helps to evaluate effects of global warming, tourists, colored lights, climatic conditions, etc. for sustainable development of the ice cave as a tourism resource. In some other ice caves in China, managers have installed airtight doors at these ice caves' entrances with the intention of "protecting" these caves, but this in fact prevents cooling in winter and these cave ices will entirely melt within tens of years.

  6. Abiotic factors drives floristic variations of fern's metacommunity in an Atlantic Forest remnant.

    Science.gov (United States)

    Costa, L E N; Farias, R P; Santiago, A C P; Silva, I A A; Barros, I C L

    2018-02-15

    We analyzed floristic variations in fern's metacommunity at the local scale and their relationship with abiotic factors in an Atlantic Forest remnant of northeastern Brazil. Floristic and environmental variations were accessed on ten plots of 10 × 20 m. We performed cluster analyses, based on Bray-Curtis dissimilarity index to establish the floristic relationship. The influence of abiotic factors: luminosity, temperature, relative air humidity and relative soil moisture was evaluated from a redundancy analysis. We found 24 species belonging to 20 genera and 12 families. The fern's flora showed high floristic heterogeneity (>75% for most of the plot's associations). The fern's metacommunity was structured along an abiotic gradient modulated by temperature, luminosity, and relative soil moisture.

  7. Paleo-watertable definition using cave ferromanganese stromatolites and associated cave-wall notches (Sierra de Arnero, Spain)

    Science.gov (United States)

    Rossi, Carlos; Villalaín, Juan J.; Lozano, Rafael P.; Hellstrom, John

    2016-05-01

    The steeply-dipping-dolostone-hosted caves of the Sierra de Arnero (N Spain) contain low-gradient relict canyons with up to ten mapped levels of ferromanganese stromatolites and associated wall notches over a vertical range of 85 m, the highest occurring 460 m above base level. Despite a plausible speleogenetic contribution by pyrite oxidation, and the irregular cave-wall mesomorphologies suggestive of hypogenic speleogenesis, the Arnero relict caves are dominantly epigenic, as indicated by the conduit pattern and the abundant allogenic sediments. Allogenic input declined over time due to a piracy-related decrease in the drainage area of allogenic streams, explaining the large size of the relict Arnero caves relative to the limited present-day outcrop area of the karstified carbonates. Allogenic-sediment input also explains the observed change from watertable canyons to phreatic conduits in the paleo-downstream direction. Stromatolites and notches arguably formed in cave-stream passages at the watertable. The best-defined paleo-watertables show an overall slope of 1.7°, consistent with the present-day relief of the watertable, with higher-slope segments caused by barriers related to sulfide mineralization. The formation of watertable stromatolites favored wall notching by the combined effect of enhanced acidity by Mn-Fe oxidation and shielding of cave floors against erosion. Abrasive bedload further contributed to notch formation by promoting lateral mechanical erosion and protecting passage floors. The irregular wallrock erosional forms of Arnero caves are related partly to paragenesis and partly to the porous nature of the host dolostones, which favored irregular dissolution near passage walls, generating friable halos. Subsequent mechanical erosion contributed to generate spongework patterns. The dolostone porosity also contributes to explain the paradox that virtually all Arnero caves are developed in dolostone despite being less soluble than adjacent

  8. The first fossil of a bolbitidoid fern belongs to the early-divergent lineages of Elaphoglossum (Dryopteridaceae).

    Science.gov (United States)

    Lóriga, Josmaily; Schmidt, Alexander R; Moran, Robbin C; Feldberg, Kathrin; Schneider, Harald; Heinrichs, Jochen

    2014-09-01

    • Closing gaps in the fossil record and elucidating phylogenetic relationships of mostly incomplete fossils are major challenges in the reconstruction of the diversification of fern lineages through time. The cosmopolitan family Dryopteridaceae represents one of the most species-rich families of leptosporangiate ferns, yet its fossil record is sparse and poorly understood. Here, we describe a fern inclusion in Miocene Dominican amber and investigate its relationships to extant Dryopteridaceae.• The morphology of the fossil was compared with descriptions of extant ferns, resulting in it being tentatively assigned to the bolbitidoid fern genus Elaphoglossum. This assignment was confirmed by reconstructing the evolution of the morphological characters preserved in the inclusion on a molecular phylogeny of 158 extant bolbitidoid ferns. To assess the morphology-based assignment of the fossil to Elaphoglossum, we examined DNA-calibrated divergence time estimates against the age of the amber deposits from which it came.• The fossil belongs to Elaphoglossum and is the first of a bolbitidoid fern. Its assignment to a particular section of Elaphoglossum could not be determined; however, sects. Lepidoglossa, Polytrichia, and Setosa can be discounted because the fossil lacks subulate scales or scales with acicular marginal hairs. Thus, the fossil might belong to either sects. Amygdalifolia, Wrightiana, Elaphoglossum, or Squamipedia or to an extinct lineage.• The discovery of a Miocene Elaphoglossum fossil provides remarkable support to current molecular clock-based estimates of the diversification of these ferns. © 2014 Botanical Society of America, Inc.

  9. Relative ages of lava flows at Alba Patera, Mars

    International Nuclear Information System (INIS)

    Schneeberger, D.M.; Pieri, D.C.

    1987-01-01

    Many large lava flows on the flanks of Alba Patera are astonishing in their volume and length. As a suite, these flows suggest tremendously voluminous and sustained eruptions, and provide dimensional boundary conditions typically a factor of 100 larger than terrestrial flows. One of the most striking features associated with Alba Patera is the large, radially oriented lava flows that exhibit a variety of flow morphologies. These include sheet flows, tube fed and tube channel flows, and undifferentiated flows. Three groups of flows were studied; flows on the northwest flank, southeast flank, and the intracaldera region. The lava flows discussed probably were erupted as a group during the same major volcanic episode as suggested by the data presented. Absolute ages are poorly constrained for both the individual flows and shield, due in part to disagreement as to which absolute age curve is representative for Mars. A relative age sequence is implied but lacks precision due to the closeness of the size frequency curves

  10. Building 887: An Aladdin's Cave for Physicists

    CERN Multimedia

    2001-01-01

    Building 887 on the Prévessin site is home to numerous experiments bringing together physicists and engineers from around the world. Its diversity makes the huge building a replica of CERN in miniature. The Installation unit of the SL EA group in front of the support structure for the ATLAS muon chambers. From left to right, seated: Pierre Gimenez, Yves Bonnet, Yves Naveau, Alain Pinget, Christian Becquet, Camille Adenot; standing: Philippe Guillot, Thierry Reynes, Monserrat Zurita-Perez, Claude Ferrari et Denis Gacon. The big wheel to be used for the ATLAS muon chambers (see below) is much the most spectacular installation currently occupying Building 887. But it is far from being the only attraction. Push open the heavy doors of this immense hall and it is a bit like entering a physicists' Aladdin's cave. The building, 55 metres wide and 300 metres long, is a treasure trove of engineering and technology, a CERN in miniature, housing dozens of collaborations from all over the world. With its 150...

  11. Pleistocene cave art from Sulawesi, Indonesia.

    Science.gov (United States)

    Aubert, M; Brumm, A; Ramli, M; Sutikna, T; Saptomo, E W; Hakim, B; Morwood, M J; van den Bergh, G D; Kinsley, L; Dosseto, A

    2014-10-09

    Archaeologists have long been puzzled by the appearance in Europe ∼40-35 thousand years (kyr) ago of a rich corpus of sophisticated artworks, including parietal art (that is, paintings, drawings and engravings on immobile rock surfaces) and portable art (for example, carved figurines), and the absence or scarcity of equivalent, well-dated evidence elsewhere, especially along early human migration routes in South Asia and the Far East, including Wallacea and Australia, where modern humans (Homo sapiens) were established by 50 kyr ago. Here, using uranium-series dating of coralloid speleothems directly associated with 12 human hand stencils and two figurative animal depictions from seven cave sites in the Maros karsts of Sulawesi, we show that rock art traditions on this Indonesian island are at least compatible in age with the oldest European art. The earliest dated image from Maros, with a minimum age of 39.9 kyr, is now the oldest known hand stencil in the world. In addition, a painting of a babirusa ('pig-deer') made at least 35.4 kyr ago is among the earliest dated figurative depictions worldwide, if not the earliest one. Among the implications, it can now be demonstrated that humans were producing rock art by ∼40 kyr ago at opposite ends of the Pleistocene Eurasian world.

  12. Are rates of species diversification and body size evolution coupled in the ferns?

    Science.gov (United States)

    Testo, Weston L; Sundue, Michael A

    2018-03-01

    Understanding the relationship between phenotypic evolution and lineage diversification is a central goal of evolutionary biology. To extend our understanding of the role morphological evolution plays in the diversification of plants, we examined the relationship between leaf size evolution and lineage diversification across ferns. We tested for an association between body size evolution and lineage diversification using a comparative phylogenetic approach that combined a time-calibrated phylogeny and leaf size data set for 2654 fern species. Rates of leaf size change and lineage diversification were estimated using BAMM, and rate correlations were performed for rates obtained for all families and individual species. Rates and patterns of rate-rate correlation were also analyzed separately for terrestrial and epiphytic taxa. We find no significant correlation between rates of leaf area change and lineage diversification, nor was there a difference in this pattern when growth habit is considered. Our results are consistent with the findings of an earlier study that reported decoupled rates of body size evolution and diversification in the Polypodiaceae, but conflict with a recent study that reported a positive correlation between body size evolution and lineage diversification rates in the tree fern family Cyatheaceae. Our findings indicate that lineage diversification in ferns is largely decoupled from shifts in body size, in contrast to several other groups of organisms. Speciation in ferns appears to be primarily driven by hybridization and isolation along elevational gradients, rather than adaptive radiations featuring prominent morphological restructuring. The exceptional diversity of leaf morphologies in ferns appears to reflect a combination of ecophysiological constraints and adaptations that are not key innovations. © 2018 Botanical Society of America.

  13. FERN - a Java framework for stochastic simulation and evaluation of reaction networks.

    Science.gov (United States)

    Erhard, Florian; Friedel, Caroline C; Zimmer, Ralf

    2008-08-29

    Stochastic simulation can be used to illustrate the development of biological systems over time and the stochastic nature of these processes. Currently available programs for stochastic simulation, however, are limited in that they either a) do not provide the most efficient simulation algorithms and are difficult to extend, b) cannot be easily integrated into other applications or c) do not allow to monitor and intervene during the simulation process in an easy and intuitive way. Thus, in order to use stochastic simulation in innovative high-level modeling and analysis approaches more flexible tools are necessary. In this article, we present FERN (Framework for Evaluation of Reaction Networks), a Java framework for the efficient simulation of chemical reaction networks. FERN is subdivided into three layers for network representation, simulation and visualization of the simulation results each of which can be easily extended. It provides efficient and accurate state-of-the-art stochastic simulation algorithms for well-mixed chemical systems and a powerful observer system, which makes it possible to track and control the simulation progress on every level. To illustrate how FERN can be easily integrated into other systems biology applications, plugins to Cytoscape and CellDesigner are included. These plugins make it possible to run simulations and to observe the simulation progress in a reaction network in real-time from within the Cytoscape or CellDesigner environment. FERN addresses shortcomings of currently available stochastic simulation programs in several ways. First, it provides a broad range of efficient and accurate algorithms both for exact and approximate stochastic simulation and a simple interface for extending to new algorithms. FERN's implementations are considerably faster than the C implementations of gillespie2 or the Java implementations of ISBJava. Second, it can be used in a straightforward way both as a stand-alone program and within new

  14. Use of rbcL and trnL-F as a two-locus DNA barcode for identification of NW-European ferns: an ecological perspective

    NARCIS (Netherlands)

    Groot, de G.A.; During, H.J.; Maas, J.W.; Schneider, H.; Erkens, R.H.J.

    2011-01-01

    Although consensus has now been reached on a general two-locus DNA barcode for land plants, the selected combination of markers (rbcL + matK) is not applicable for ferns at the moment. Yet especially for ferns, DNA barcoding is potentially of great value since fern gametophytes—while playing an

  15. Morphometric study of pillow-size spectrum among pillow lavas

    Science.gov (United States)

    Walker, George P. L.

    1992-08-01

    Measurements of H and V (dimensions in the horizontal and vertical directions of pillows exposed in vertical cross-section) were made on 19 pillow lavas from the Azores, Cyprus, Iceland, New Zealand, Tasmania, the western USA and Wales. The median values of H and V plot on a straight line that defines a spectrum of pillow sizes, having linear dimensions five times greater at one end than at the other, basaltic toward the small-size end and andesitic toward the large-size end. The pillow median size is interpreted to reflect a control exercised by lava viscosity. Pillows erupted on a steep flow-foot slope in lava deltas can, however, have a significantly smaller size than pillows in tabular pillowed flows (inferred to have been erupted on a small depositonal slope), indicating that the slope angle also exercised a control. Pipe vesicles, generally abundant in the tabular pillowed flows and absent from the flow-foot pillows, have potential as a paleoslope indicator. Pillows toward the small-size end of the spectrum are smooth-surfaced and grew mainly by stretching of their skin, whereas disruption of the skin and spreading were important toward the large-size end. Disruption involved increasing skin thicknesses with increasing pillow size, and pillows toward the large-size end are more analogous with toothpaste lava than with pahoehoe and are inferred from their thick multiple selvages to have taken hours to grow. Pseudo-pillow structure is also locally developed. An example of endogenous pillow-lava growth, that formed intrusive pillows between ‘normal’ pillows, is described from Sicily. Isolated pillow-like bodies in certain andesitic breccias described from Iceland were previously interpreted to be pillows but have anomalously small sizes for their compositions; it is now proposed that they may lack an essential attribute of pillows, namely, the development of bulbous forms by the inflation of a chilled skin, and are hence not true pillows. Para-pillow lava is

  16. Recent flood lavas in the Elysium region of Mars

    International Nuclear Information System (INIS)

    Plescia, J.B.

    1990-01-01

    A volcanic origin is presently suggested for the Cerberus Formation region of smooth plains in the southeastern Elysium region of Mars, on the basis of its surface morphology, lobate edges, vents, and an embayment relation of the unit with adjacent, older units. The low viscosity lavas that filled a topographic depression in southeastern Elysium subsequently flowed into western Amazonic Planitia via channels formed by an earlier fluvial episode. A young, upper Amazonian dating is indicated by crater frequencies and stratigraphic relations, implying that large-scale eruptions of low-viscosity lava were still possible late in Martian history. 34 refs

  17. Deformation at Lava Lake Volcanoes: Lessons from Karthala

    Science.gov (United States)

    Biggs, J.; Rust, A.; Owens, C.

    2014-12-01

    To remain hot, permanent lava lakes require a continuous connection to a magma reservoir. Depending on the state of the conduit, changes in magma pressure could result in changes in the lake level (hydraulic head) or be accommodated elastically leading to surface deformation. Observing deformation is therefore key to understanding the plumbing system associated with lava lakes. However, the majority of the world's lava lakes lie in difficult socio-economic or remote locations meaning that there are few ground-based observations, and it is often necessary to rely on satellite imagery. Karthala volcano experienced a sequence of eruptions in April 2005, Nov 2005, May 2006 and Jan 2007. The first 3 took place at the Choungou Chahale crater, which typically contains either a water or lava lake; the last formed a new pit crater to the north. Satellite thermal imagery (Hirn et al, 2008) does not show an anomaly during the first eruption, which had a phreatomagmatic component, but large thermal anomalies, associated with an ephemeral lava lake were detected during the Nov 2005 and May 2006 eruptions. The final eruption produced a smaller anomaly attributed to a minor lava flow. Here we present InSAR observations from 2004-2010. We find no significant deformation associated with the first three eruptions, but the January 2007 eruption was associated with ~25 cm of deformation near the volcano's summit, characteristic of a dyke intrusion aligned with the northern rift zone. We also observe an unusual pattern deformation along the coast which may be attributed to rapid settling of soft sediment or recent volcanic deposits triggered by seismic activity. We propose that the first eruption cleared the reservoir-summit connection and interacted with the water in Choungou Chahale. The following eruptions formed a lava lake, but without causing deformation. By the final eruption, the conduit had become blocked and magma intruded along the rift zone causing deformation but no

  18. Determination of heavy metals in soil and different parts of Diplazium esculentum (medicinal fern)

    Science.gov (United States)

    Jasim, Hind S.; Idris, Mushrifah; Abdullah, Aminah; Kadhum, A. A. H.

    2014-09-01

    Diplazium esculentum is a widely used medicinal fern in Malaysia and other regions worldwide. Heavy metals in plants should be determined because prolonged human intake of toxic trace elements, even at low doses, results in organ malfunction and causes chronic toxicity. Hence, substantial information should be obtained from plants that grow on soils containing high concentrations of heavy metals. This study aimed to determine the physicochemical characteristics of soil and heavy metal concentrations (Pb, Cr, Mn, Cu, and Zn) in different parts of D. esculentum and soil, which were collected from the fern garden of Universiti Kebangsaan Malaysia. Results showed that heavy metals were highly accumulated in D. esculentum roots.

  19. Fern extracts potentiate fluconazole activity and inhibit morphological changes in Candida species

    Directory of Open Access Journals (Sweden)

    Maria A. Freitas

    2017-11-01

    Conclusions: The extracts obtained from the fern species L. venustum and P. calomelanos dose not present significant antifungal activity. However, P. calomelanos potentiates the activity of fluconazole and both extracts inhibits the morphological changes in Candida species, indicating that they have potential pharmacological activity as modulators of fungal biology. Therefore, novel studies are required to characterize the interference of these extracts in the virulence and pathogenicity of Candida species as well as the potential of fern species to treat fungal infections.

  20. Tamarugite from Diana Cave (SW Romania) -first true karst occurrence

    Science.gov (United States)

    Pušcaš, C. M.; Onac, B. P.; Effenberger, H. S.; Povarǎ, I.

    2012-04-01

    Diana Cave is located within the town limits of Baile Herculane (SW Romania) and develops as a 14 m long, westward oriented, unique passage guided by the Diana fault [1]. At the far end of the cave, the thermo-mineral Diana Spring wells forth. In the early 1970s a mine gallery that intersected the cave was created to drain the water into a pumping station and the original cave passage was somewhat altered and reinforced with concrete. Today the concrete and the silty limestone cave walls are heavily corroded by H2SO4 outgassing from the hot water (ca. 50°C) and display abundant gypsum crusts, soggy aggregates of native S, and a variety of more exotic sulfates. Among them, a mineral that has been previously identified in caves only in connection to volcanic activity, either as thermal springs or fumaroles [2]: tamarugite [NaAl(SO4)26H2O]. It was [3] that first mentioned the occurrence of this Na and Al sulfate in Diana Cave, our research aiming to give a detailed description of this mineral, its paragenesis, and mechanisms of precipitation. Recently, tamarugite has also been identified in a sulfuric acid cave from Greece [4]. Along with powder X-ray diffractions coupled with Rietveld refinement, scanning electron microscope, and electron probe micro-analysis, δ18O and δ34S compositions of the sulfate mineral as well as precipitates from the water were analyzed to identify and better constrain the genesis of this rare sulfate. Regrettably, the crystal size of our specimens is inappropriate for identification by means of single crystal X-ray diffraction. Physical and chemical parameters of Diana Spring were as well measured on several occasions. Geochemical analysis suggests that the minute, white tamarugite flakes precipitated in Diana Cave as a result of the interactions between the thermo-mineral water or water vapor and the original limestone bedrock and concrete that blankets the mine gallery. [1] Povara, I., Diaconu, G., Goran, C. (1972). Observations pr

  1. Cave exploitation by an usual epigean species: a review on the current knowledge on fire salamander breeding in cave

    Directory of Open Access Journals (Sweden)

    Raoul Manenti

    2017-11-01

    Full Text Available The fire salamander (Salamandra salamandra is a relatively common epigean amphibian, widely distributed throughout Europe, which usually gives birth to aquatic larvae. Even if epigean streams represent the most common places in which the species breeds, in some countries caves with underground waters are also used. To improve our understanding of the habitat features allowing successful breeding of salamanders in underground sites, we combined an exhaustive review of the available literature, especially the grey one, with direct observations performed from 2008 to 2017 in several natural and artificial caves of Lombardy, Liguria and Tuscany (Italy, Ariège and Provence (France. We provide a synthesis of published and unpublished caves in which the fire salamander breeding has been observed, along with a synthesis of the investigated ecological, behavioural and morphological traits. The use of underground sites is reported in several published papers and appears to be a common phenomenon not limited to single karst areas. The absence of predators, the relative stability of the aquatic habitats and the possibility to exploit new ecological resources are environmental factors that favour the breeding of the fire salamander. Our synthesis suggests that breeding of fire salamanders in caves is not a random event, but a widespread phenomenon that may be linked to specific biogeographical factors. Further insights may be obtained by performing genetic analyses on both cave and epigean populations, and considering larger landscape scales for ecological studies as well. Gene flow between salamanders that breed in caves and in streams probably occurs, but on the other hand, assortative mating might limit it, thus allowing the conservation of local adaptations driving successful cave colonisation.

  2. The CAVE (TM) automatic virtual environment: Characteristics and applications

    Science.gov (United States)

    Kenyon, Robert V.

    1995-01-01

    Virtual reality may best be defined as the wide-field presentation of computer-generated, multi-sensory information that tracks a user in real time. In addition to the more well-known modes of virtual reality -- head-mounted displays and boom-mounted displays -- the Electronic Visualization Laboratory at the University of Illinois at Chicago recently introduced a third mode: a room constructed from large screens on which the graphics are projected on to three walls and the floor. The CAVE is a multi-person, room sized, high resolution, 3D video and audio environment. Graphics are rear projected in stereo onto three walls and the floor, and viewed with stereo glasses. As a viewer wearing a location sensor moves within its display boundaries, the correct perspective and stereo projections of the environment are updated, and the image moves with and surrounds the viewer. The other viewers in the CAVE are like passengers in a bus, along for the ride. 'CAVE,' the name selected for the virtual reality theater, is both a recursive acronym (Cave Automatic Virtual Environment) and a reference to 'The Simile of the Cave' found in Plato's 'Republic,' in which the philosopher explores the ideas of perception, reality, and illusion. Plato used the analogy of a person facing the back of a cave alive with shadows that are his/her only basis for ideas of what real objects are. Rather than having evolved from video games or flight simulation, the CAVE has its motivation rooted in scientific visualization and the SIGGRAPH 92 Showcase effort. The CAVE was designed to be a useful tool for scientific visualization. The Showcase event was an experiment; the Showcase chair and committee advocated an environment for computational scientists to interactively present their research at a major professional conference in a one-to-many format on high-end workstations attached to large projection screens. The CAVE was developed as a 'virtual reality theater' with scientific content and

  3. Eastern Cave- and Crevice-Dwelling Bats Potentially Impacted by USACE Reservoir Operations

    National Research Council Canada - National Science Library

    Martin, C.O

    2002-01-01

    ...; Kasul, Martin, and Allen 2000). This note provides information on six species of forest bats that roost in caves or cave-like structures and potentially occur on Corps projects in the Eastern United States...

  4. The Evaluation of a Motion Base Driving Simulator in a Cave at TACOM

    National Research Council Canada - National Science Library

    Mollenhauer, M. A; Romano, R. A; Brumm, B

    2004-01-01

    The purpose of this presentation is to describe the highlights of a research program designed to investigate the feasibility of creating a motion base driving simulator in a Cave Automatic Virtual Environment (CAVE...

  5. The potential of perennial cave ice in isotope palaeoclimatology

    International Nuclear Information System (INIS)

    Yonge, Charles J.; MacDonald, William D.

    1999-01-01

    Perennial ice from caves on and to the east of the Canadian Great Divide yield delta O 18 and delta D values which are usually high measurements where compared with the average precipitation for the region. Furthermore, these ice data fall below and along lines of lower slope than the Global Meteoric Water Line. To explain the observed relationships, we propose the following process. a vapour-ice isotopic fractionation mechanism operates on warm season vapour when it precipitates as hoar ice on entering the caves. The subsequent fall of hoar to the cave floor through mechanical overloading along with ice derived from ground-water seepage (with a mean annual isotopic composition), results in massive ice formation of a mixed composition. This mixed composition is what is observed in the characteristic relationships found here. Such findings suggest that a warm versus cold climate interpretation for ancient cave ice may be the opposite of that found in the more familiar polar and glacial ice caves. (Author) 3 figs., 1 tab., 12 refs

  6. Differential preservation of vertebrates in Southeast Asian caves

    Directory of Open Access Journals (Sweden)

    Julien Louys

    2017-09-01

    Full Text Available Caves have been an important source of vertebrate fossils for much of Southeast Asia, particularly for the Quaternary. Despite this importance, the mechanisms by which vertebrate remains accumulate and preserve in Southeast Asian caves has never been systematically reviewed or examined. Here, we present the results of three years of cave surveys in Indonesia and Timor-Leste, describing cave systems and their attendant vertebrate accumulations in diverse geological, biogeographical, and environmental settings. While each cave system is unique, we find that the accumulation and preservation of vertebrate remains are highly dependent on local geology and environment. These factors notwithstanding, we find the dominant factor responsible for faunal deposition is the presence or absence of biological accumulating agents, a factor directly dictated by biogeographical history. In small, isolated, volcanic islands, the only significant accumulation occurs in archaeological settings, thereby limiting our understanding of the palaeontology of those islands prior to human arrival. In karstic landscapes on both oceanic and continental islands, our understanding of the long-term preservation of vertebrates is still in its infancy. The formation processes of vertebrate-bearing breccias, their taphonomic histories, and the criteria used to determine whether these represent syngenetic or multiple deposits remain critically understudied. The latter in particular has important implications for arguments on how breccia deposits from the region should be analysed and interpreted when reconstructing palaeoenvironments.

  7. FURTHER NOTES ON THE FERN-GENUS HETEROGONIUM PRESL

    Directory of Open Access Journals (Sweden)

    R. E. HOLTTUM

    2015-11-01

    Full Text Available In the "Sarawak Museum Journal," volume V (1949, pages 156-166,I gave a revised account of the genus Heterogonium Presl, based on specimens in the Singapore herbarium. Dr M. A. Donk wrote subsequently pointing out that specimens at Bogor (Buitenzorg add materially to the information contained in that paper. The present paper gives the result of a study of the Bogor specimens. I am glad to express my gratitude to Dr Donk for calling my attention to species which I had overlooked when searching literature on Malaysian ferns for indications of affinity to Heterogonium. The Bogor material includes the type specimens of Acrostichum teysmannianum Bak., Phegopteris schizoloma v. A. v. R. Dryopteris sagenoides forma contracta v. A. v. R., Polybotrya nieuwenhuisii Racib. and Polybotrya nieuwenhuisii var. brooksii v. A. v. R., also many sheets of H. giganteum, and material of H. sagenoides from a wider geographic range than I had previously seen. Summarizing the results of the present paper, I have united H.nieuwenhuisii and H. stenosemioides of my former paper, and have also united H. saxicola with H. giganteum. Further, I now recognize a second exindusiate species allied to H. sagenoides; but the variation in pubescence among exindusiate specimens, as within the species H. sagenoides proper, is very considerable, and I find it very difficult to draw specific limits. It is likely that there are local races, but much more field work in many localities is necessary before one can speak with certainty about this or define their status taxonomically. One interesting fact is that no exindusiate specimens of this alliance have been found in the Malay Peninsula, whereas Peninsular collections of indusiate H. sagenoides aremore abundant than from any other area. It is especially the variation in pubescence among specimens of H. sagenoides, and the lack of clear-cut varieties or subspecies within the Malay Peninsula, that deters me from distinguishing more

  8. Structural and hydrological controls on the development of a river cave in marble (Tapagem Cave - SE Brazil

    Directory of Open Access Journals (Sweden)

    William Sallun Filho

    2015-01-01

    Full Text Available Tapagem Cave (or Devil’s Cave is a river cave developed in the dolomite marble karst of the Serra do André Lopes (State of São Paulo, southeastern Brazil. Although this region is a plateau with significant variation in elevation and a humid subtropical climate, the cave is an anomalous feature in the André Lopes karst because there are few other caves. The marble, which is in a synclinal structure with subjacent phyllites, is a karst aquifer perched above the regional base level (Ribeira River and has little allogenic recharge. The cave developed on a secondary anticline on the northwest flank of the marble synform forming a blind valley, the Tapagem River sink, that is an underground tributary of Ostras River. Development of the cave is due to the entrenchment of the Ostras through-valley and the large allogenic catchment area of the sink. In plan view, the morphology of the cave can be divided into three different sectors. The first sector, known as the Tourist Sector, has extensive collapse rooms, fossil passages and a variety of speleothems of notable dimensions. The second and most extensive sector is the river passage, which is a sinuous gallery controlled by marble banding with NE-SW cleavage and NW-SE fractures. In cross-section, the passages are vadose canyons up to 70 m in height, controlled by the marble banding. Four NW-SE diabase dykes in this passage do not affect its direction in plan view. The third sector is an extensive network of passages and collapse rooms, which are interlaced in plan view and on different levels, forming a maze pattern. Initially, the Tapagem and Ostras Rivers developed on a gentle surface and flowed into the Ribeira River. With the entrenchment of the Ostras through-valley, the Tapagem River partially infiltrated via a paleosink into the upper passage of the “Erectus Room," remaining a half-blind valley. Following a series of collapses and obstructions, the River next infiltrated via the current

  9. Ancient DNA reveals differences in behaviour and sociality between brown bears and extinct cave bears.

    Science.gov (United States)

    Fortes, Gloria G; Grandal-d'Anglade, Aurora; Kolbe, Ben; Fernandes, Daniel; Meleg, Ioana N; García-Vázquez, Ana; Pinto-Llona, Ana C; Constantin, Silviu; de Torres, Trino J; Ortiz, Jose E; Frischauf, Christine; Rabeder, Gernot; Hofreiter, Michael; Barlow, Axel

    2016-10-01

    Ancient DNA studies have revolutionized the study of extinct species and populations, providing insights on phylogeny, phylogeography, admixture and demographic history. However, inferences on behaviour and sociality have been far less frequent. Here, we investigate the complete mitochondrial genomes of extinct Late Pleistocene cave bears and middle Holocene brown bears that each inhabited multiple geographically proximate caves in northern Spain. In cave bears, we find that, although most caves were occupied simultaneously, each cave almost exclusively contains a unique lineage of closely related haplotypes. This remarkable pattern suggests extreme fidelity to their birth site in cave bears, best described as homing behaviour, and that cave bears formed stable maternal social groups at least for hibernation. In contrast, brown bears do not show any strong association of mitochondrial lineage and cave, suggesting that these two closely related species differed in aspects of their behaviour and sociality. This difference is likely to have contributed to cave bear extinction, which occurred at a time in which competition for caves between bears and humans was likely intense and the ability to rapidly colonize new hibernation sites would have been crucial for the survival of a species so dependent on caves for hibernation as cave bears. Our study demonstrates the potential of ancient DNA to uncover patterns of behaviour and sociality in ancient species and populations, even those that went extinct many tens of thousands of years ago. © 2016 John Wiley & Sons Ltd.

  10. A review of factors affecting cave climates for hibernating bats in temperate North America

    Science.gov (United States)

    Roger W. Perry

    2013-01-01

    The fungal pathogen Geomyces destructans, which causes white-nose syndrome in bats, thrives in the cold and moist conditions found in caves where bats hibernate. To aid managers and researchers address this disease, an updated and accessible review of cave hibernacula and cave microclimates is presented. To maximize energy savings and reduce...

  11. A Framework for Aligning Instructional Design Strategies with Affordances of CAVE Immersive Virtual Reality Systems

    Science.gov (United States)

    Ritz, Leah T.; Buss, Alan R.

    2016-01-01

    Increasing availability of immersive virtual reality (IVR) systems, such as the Cave Automatic Virtual Environment (CAVE) and head-mounted displays, for use in education contexts is providing new opportunities and challenges for instructional designers. By highlighting the affordances of IVR specific to the CAVE, the authors emphasize the…

  12. Guanophilic fungi in three caves of southwestern Puerto Rico

    Directory of Open Access Journals (Sweden)

    Nieves-Rivera Angel M.

    2009-01-01

    Full Text Available Fifty species of guanophilic (bat guano-loving fungi were isolated from field-collected samples within three caves in southwesternPuerto Rico; most were mitosporic fungi (23 species. The caves studied were Cueva La Tuna (Cabo Rojo, Cueva de Malano(Sistema de Los Chorros, San Germán, and Cueva Viento (El Convento Cave-Spring System, Guayanilla-Peñuelas. The mostconspicuous fungus by far was the zygomycete Circinella umbellata (Mucorales. Circinella umbellata dominated the bat guanoincubation chambers (Petri dishes lined with sterile filter paper moistened with sterile water at ambient laboratory conditions.Nineteen species of basidiomycetes (e.g., Ganoderma cf. resinaceum, Geastrum cf. minimum, Lepiota sp., Polyporus sp., Ramariasp. and three species of ascomycetes (Hypoxylon sp., Xylaria anisopleura, and X. kegeliana were also recorded. They were foundon soil, rotting leaves, bark and rotting wood, buried in bat guano located below natural skylights or sinkholes.

  13. The chronology of Moncks Cave, Canterbury, New Zealand

    International Nuclear Information System (INIS)

    Jacomb, C.

    2008-01-01

    Moncks Cave is a key site in understanding the nature and course of change to Maori culture during the early period of New Zealand prehistory because of the range of both perishable and non-perishable artefacts found there in 1889. Understood to have been completely excavated at that time, the interpretation of the material culture of the site has been rendered difficult by the absence of chronological or stratigraphic provenance data. Recent investigations at Moncks Cave revealed several intact cultural deposits, including both faunal and artefactual remains. Eleven radiocarbon determinations on marine shell suggest that the cave was occupied some time between the mid-fourteenth and mid-fifteenth centuries AD. Although the dates cannot be directly correlated with any particular artefact, the results have important implications for the interpretation of the place of the site and its contents as a whole in the context of the New Zealand prehistoric sequence. (author). 22 refs., 5 figs., 1 tab

  14. Thermophysical properties of the Lipari lavas (Southern Tyrrhenian Sea

    Directory of Open Access Journals (Sweden)

    D. Russo

    1997-06-01

    Full Text Available Results of thermophysical investigations into the lavas of the island of Lipari (Southern Tyrrhenian Sea are presented. Samples selected for laboratory measurements belong to four main magmatic cycles, which produced basaltic-andesitic, andesitic and rhyolitic lavas. The wet-bulk density and the thermal conductivity measured on 69 specimens range from 1900 to 2760 kg m-3 and from 1.02 to 2.88 W m-1 K-1, respectively. Porosity is never negligible and its influence on density is maximum in rhyolites of the third cycle. The thermal conductivity is also influenced by the amount of glass. Rhyolitic obsidians show values lower than other rhyolites, although the latter rocks have a larger average porosity. The radioactive heat production determined on 36 specimens varies with the rock type, depending on the amount of U, Th and K. In basic lavas of the first cycle its value is 0.95°± 0.30 mW m-3, while in rhyolites of the fourth cycle it attains 6.68°±0.61 mW m-3. A comparison between results of g-ray spectrometry and X-ray fluorescence points out that the assumption of equilibrium in the decay series of the isotopic elements seems fulfilled. The information obtained is useful not only for the interpretation of geophysical surveys but also for the understanding of the geochemical characteristics of lavas.

  15. Radon and thoron in cave dwellings (Yan'an, China)

    International Nuclear Information System (INIS)

    Wiegand, J.; Feige, S.; Xie Quingling; Schreiber, U.; Wieditz, K.; Wittmann, C.; Luo Xiarong

    2000-01-01

    222 Rn and 220 Rn concentrations were measured in cave dwellings and brick houses in the region of Yan'an (China) during summer 1997. The underground dwellings are built into Quaternary loess, and all investigated houses are founded on it. The median values of indoor 222 Rn and 220 Rn concentrations are 42 (n = 18) and 77Bq m -3 (n = 15) for brick houses and 92 (n = 23) and 215 (n = 17) Bq m -3 for cave dwellings. To classify the dwellings in respect to their cave-character, the fraction of walls having a direct contact to the loess is calculated for each dwelling. While the 222 Rn concentrations are increasing with higher fractions, the 220 Rn concentrations are not correlated with this fraction. On the other hand, due to the short half-life of 220 Rn the distance from the measuring point to the walls is negatively correlated with the 220 Rn concentration, while there is no correlation with the 222 Rn concentration. Therefore, concentric isolines of 220 Rn concentrations showing a strong gradient were detected in cave dwellings. An influence of the ventilation rate is distinct for 222 Rn but weak for 220 Rn. The effective dose rates for 222 Rn and 220 Rn and their progenies are calculated for brick houses (2.7 mSv y -1 ), cave dwellings (7.1 mSv y -1 ), and for traditional cave dwellings with a bed foundation built with loess (16.7 mSv y -1 ). These calculations are based on summer measurements only. It is expected that the true effective dose rates will be significantly higher

  16. Secondary sulfate minerals from Alum Cave Bluff: Microscopy and microanalysis

    Energy Technology Data Exchange (ETDEWEB)

    Lauf, R.J.

    1997-07-01

    Microcrystals of secondary sulfate minerals from Alum Cave Bluff, Great Smoky Mountains National Park, were examined by scanning electron microscopy and identified by X-ray fluorescence (XRF) in the SEM. Among the samples the author discovered three new rare-earth sulfates: coskrenite-(Ce), levinsonite-(Y), and zugshunstite-(Ce). Other minerals illustrated in this report include sulfur, tschermigite, gypsum, epsomite, melanterite, halotrichite, apjohnite, jarosite, slavikite, magnesiocopiapite, and diadochite. Additional specimens whose identification is more tentative include pickeringite, aluminite, basaluminite, and botryogen. Alum Cave is a ``Dana locality`` for apjohnite and potash alum, and is the first documented North American occurrence of slavikite.

  17. A radon survey performed in caves in Slovenia

    International Nuclear Information System (INIS)

    Jovanovic, P.

    2002-01-01

    A survey of radon and radon decay product concentrations in several caves in a limestone region in Slovenia was initiated in 1986. In the period from 1989 to 1998, monthly surveys were undertaken in several caves which are open to tourists or used for speleotherapy purposes. The reason for carrying out these surveys, were dose estimates obtained for the guides and medical staff working in the caves. Daily average radon gas concentration determined ranged from several 100 Bq/m 3 up to 27 kBq/m 3 . Higher values were measured in the summer period. The equilibrium factors derived ranged from 0.05 to 0.89, with the higher values being measured in the winter period in vertical caves. In horizontal caves (with two entrances located opposite one another) these values ranged between 0.55 and 0.89. Annual doses estimated on the basis of various lung models ranged from 10 mSv to 85 mSv per year and per 2000 working hours. A significant difference was observed between the doses estimated by means of dosimetric models, and those estimated on the basis of the epidemiological model presented in ICRP 65. The value for the unattached fraction indicated in ICRP 65 is about 3%, but our measurements performed in the caves yielded higher values of up to 15%, with this highest value being determined in the Postojna cave. In the coming years, we will perform measurements to obtain the values for concentrations of unattached particles of radon daughters and values for particle-size distribution in the 3 different caves with the highest occupancy times for visitors. There are no regulations in force in Slovenia affecting exposures to elevated radon and radon daughter concentrations among underground workers. The health inspectorate can impose radiation monitoring measures for the purposes of performing dose calculations for underground workers. The results from such monitoring measures will contribute to the establishment of an ordinance regulating the performance of measurements at low

  18. Alsophila weidenbrueckii (Cyatheaceae), a new scaly tree fern from Papua New Guinea

    NARCIS (Netherlands)

    Lehnert, M.

    2016-01-01

    The scaly tree fern Alsophila weidenbrueckii is described and illustrated as new to science. It occurs in the Bismarck range in north-eastern New Guinea at 1200–2100 m in evergreen wet mountain forest. The species reaches maturity at a comparatively old age and large size and regenerates only in

  19. Novel fungi from an ancient niche: lachnoid and chalara-like fungi on ferns

    NARCIS (Netherlands)

    Guatimosim, E.; Schwartsburd, P. B.; Crous, P. W.; Barreto, R. W.

    2016-01-01

    A survey was conducted in Brazil to collect fungi on ferns. Based on morphology and inferred phylogeny from DNA sequences of two loci, namely the internal transcribed spacer (ITS) regions and the large subunit nuclear ribosomal RNA gene (LSU), several species belonging to chalara-like genera and

  20. Coeval Eocene blooms of the freshwater fern Azolla in and around Arctic and Nordic seas

    NARCIS (Netherlands)

    Barke, J.; Burgh, A.H.P. van der; Konijnenburg-van Cittert, J.H.A. van; Collinson, M.E.; Pearce, M.A.; Bujak, J.; Heilman-Clausen, C.; Lotter, A.F.; Speelman, E.N.; Kempen, M.M.L. van; Reichart, G.-J.; Brinkhuis, H.

    2012-01-01

    For a short time interval (c. 1.2 Myr) during the early middle Eocene (~ 49 Myr), the central Arctic Ocean was episodically densely covered by the freshwater fern Azolla, implying sustained freshening of surface waters. Coeval Azolla fossils in neighboring Nordic seas were thought to have been

  1. A review of the fern genus Hypolepis (Dennstardtiaceae) in the Malesian and Pacific regions

    NARCIS (Netherlands)

    Brownsey, P.J.

    1987-01-01

    Fourteen species and one subspecies of the fern genus Hypolepis Bernh. are recognised in the Malesian and Pacific regions, excluding Australia and New Zealand. Three species, H. hawaiiensis, H. malesiana and H. scabristipes, and one subspecies, H. elegans subsp. carolinensis, are described for the

  2. Western sword fern avoids the extreme drought of 2012-2014

    Science.gov (United States)

    Emily E. Burns; Peter Cowan; Wendy Baxter; Deborah Zierten; Jarmilla Pittermann

    2017-01-01

    The California drought of 2012 to 2014 was the most severe drought on record for the last century and likely millennium. Warm temperatures with below-average precipitation compounded over the three-year period, creating significant and sustained aridity over the course of three growing seasons throughout the coast redwood ecosystem. The citizen science project, Fern...

  3. Jasmonic and salicylic acid response in the fern Azolla filiculoides and its cyanobiont.

    Science.gov (United States)

    de Vries, Sophie; de Vries, Jan; Teschke, Hendrik; von Dahlen, Janina K; Rose, Laura E; Gould, Sven B

    2018-01-03

    Plants sense and respond to microbes utilizing a multilayered signalling cascade. In seed plants, the phytohormones jasmonic and salicylic acid (JA and SA) are key denominators of how plants respond to certain microbes. Their interplay is especially well-known for tipping the scales in plants' strategies of dealing with phytopathogens. In non-angiosperm lineages, the interplay is less well understood, but current data indicate that it is intertwined to a lesser extent and the canonical JA/SA antagonism appears to be absent. Here, we used the water fern Azolla filiculoides to gain insights into the fern's JA/SA signalling and the molecular communication with its unique nitrogen fixing cyanobiont Nostoc azollae, which the fern inherits both during sexual and vegetative reproduction. By mining large-scale sequencing data, we demonstrate that Azolla has most of the genetic repertoire to produce and sense JA and SA. Using qRT-PCR on the identified biosynthesis and signalling marker genes, we show that Azolla is responsive to exogenously applied SA. Furthermore, exogenous SA application influenced the abundance and gene expression of Azolla's cyanobiont. Our data provide a framework for JA/SA signalling in ferns and suggest that SA might be involved in Azolla's communication with its vertically inherited cyanobiont. © 2018 John Wiley & Sons Ltd.

  4. The cyanobiont in an Azolla fern is neither Anabaena nor Nostoc.

    Science.gov (United States)

    Baker, Judith A; Entsch, Barrie; McKay, David B

    2003-12-05

    The cyanobacterial symbionts in the fern Azolla have generally been ascribed to either the Anabaena or Nostoc genera. By using comparisons of the sequences of the phycocyanin intergenic spacer and a fragment of the 16S rRNA, we found that the cyanobiont from an Azolla belongs to neither of these genera.

  5. Notes on the scholarly trajectory of Mª. Dolores Fernández-Posse

    Directory of Open Access Journals (Sweden)

    Gilman, Antonio

    2007-12-01

    Full Text Available The scholarly trajectory of Mª Dolores Fernández- Posse involved an ongoing process of critical observation, reflection, and change that is quite rare in Archaeology. Her change from normativism to a historically informed functionalism is representative of the general evolution of Spanish archaeological studies over the course of the past thirty years. What is unusual about Fernández-Posse -and attests to her straightforward and unassuming intelligence- is that she herself changed her mind and voiced that change with clarity and humor.

    La trayectoria científica de Mª Dolores Fernández-Posse se caracterizó por un proceso continuo de observación crítica, reflexión y cambio, algo poco común en Arqueología. Su cambio del normativismo a un funcionalismo de orientación histórica es representativo de la evolución de los estudios arqueológicos en España a lo largo de los últimos treinta años. Lo realmente inusual de Fernández-Posse -y lo que atestigua su directa pero no pretenciosa inteligencia- es que ella misma cambió de ideas y publicitó ese cambio con claridad y humor.

  6. New records, names and combinations of scaly tree ferns (Cyatheaceae) in eastern Malesia

    NARCIS (Netherlands)

    Coritico, F.P.; Amoroso, V.B.; Lehnert, M.

    2017-01-01

    Several scaly tree ferns (Cyatheaceae) from eastern Malesia are transferred from Cyathea s.lat. to other genera: Alsophila roroka from Sulawesi; Alsophila lamoureuxii from New Guinea; Alsophila binayana, A. bisquamata, A. mapahuwensis, A. murkelensis, A. ohaensis, and Sphaeropteris pukuana from

  7. Grass or fern competition reduce growth and survival of planted tree seedlings

    Science.gov (United States)

    Larry H. McCormick; Todd W. Bowersox

    1997-01-01

    Bareroot seedlings of northern red oak, white ash, yellow-poplar and white pine were planted into herbaceous communities at three forested sites in central Pennsylvania that were clearcut 0 to 1 year earlier. Seedlings were grown 4 years in the presence and absence of either an established grass or hay-scented fern community. Survival and height growth were measured...

  8. Population biology of two rare fern species: long-life and long-lasting stability

    Czech Academy of Sciences Publication Activity Database

    Bucharová, Anna; Münzbergová, Z.; Tájek, P.

    2010-01-01

    Roč. 97, č. 8 (2010), s. 1260-1271 ISSN 0002-9122 R&D Projects: GA MŽP SP/2D4/112/08 Institutional research plan: CEZ:AV0Z60050516 Keywords : ferns * life cycle * naturally rare species Subject RIV: EF - Botanics Impact factor: 3.052, year: 2010

  9. Differences between caves with and without bats in a Brazilian karst habitat

    Directory of Open Access Journals (Sweden)

    Camila G. Torquetti

    2017-10-01

    Full Text Available ABSTRACT. Since bats shelter in roosts during their period of diurnal inactivity, the quality and availability of roosts are important aspects of their ecology. Karst areas have great potential for the availability of day roosts, since they form caves, which serve as bat shelters. Here we characterize the caves used by bats in a preserved karst area of Southeastern Brazil. Using logistic regression analysis we identified the cave characteristics that influence bat occupation. Sixty-six caves were characterized based on measurements of internal height and width, height and width of the entrance(s of the cave, number of entrances, maximum horizontal development of cave, and internal temperature and humidity. In nineteen months we found 14 species in 32 caves. Most species were eventually recorded in multiple caves, with the exception of D. rotundus, G. soricina and A. planirostris, which were always found in the same caves. Desmodus rotundus showed maternity roost fidelity. We found no differences in microclimate between the caves that are occupied and those that are not. In other words, the microclimate of the caves studied herein can be characterized as stable over the years. The only predictor affecting the presence of bats in the study area was the cave’s maximum horizontal development: the caves that are occupied have greater horizontal development. Based on our results, we conclude that bats occupy many of the caves and that some species are more frequent in certain caves than in others, including some roosts that are used as maternity roosts. These findings indicate that these caves are important resources for the bats in the karst environment studied, and should be preserved.

  10. Formation of perched lava ponds on basaltic volcanoes: Interaction between cooling rate and flow geometry allows estimation of lava effusion rates

    Science.gov (United States)

    Wilson, L.; Parfitt, E. A.

    1993-01-01

    Perched lava ponds are infrequent but distinctive topographic features formed during some basaltic eruptions. Two such ponds, each approximately 150 m in diameter, formed during the 1968 eruption at Napau Crater and the 1974 eruption of Mauna Ulu, both on Kilauea Volcano, Hawaii. Each one formed where a channelized, high volume flux lava flow encountered a sharp reduction of slope: the flow spread out radially and stalled, forming a well-defined terminal levee enclosing a nearly circular lava pond. We describe a model of how cooling limits the motion of lava spreading radially into a pond and compare this with the case of a channelized flow. The difference in geometry has a major effect, such that the size of a pond is a good indicator of the volume flux of the lava forming it. Lateral spreading on distal shallow slopes is a major factor limiting the lengths of lava flows.

  11. Non-linear growth in tree ferns, Dicksonia antarctica and Cyathea australis.

    Science.gov (United States)

    Blair, David P; Blanchard, Wade; Banks, Sam C; Lindenmayer, David B

    2017-01-01

    Tree ferns are an important structural component of forests in many countries. However, because their regeneration is often unrelated to major disturbances, their age is often difficult to determine. In addition, rates of growth may not be uniform, which further complicates attempts to determine their age. In this study, we measured 5 years of growth of Cyathea australis and Dicksonia antarctica after a large wildfire in 2009 in south-eastern Australia. We found growth rates of these two species were unaffected by aspect and elevation but slope had a minor effect with D. antarctica growing 0.3mm faster for each additional degree of slope. Geographic location influenced growth in both species by up to 12 - 14mm/yr. The most consistent factor influencing growth rate, however, was initial height at the time of the 2009 fire; a finding consistent in both species and all geographic locations. For both tree fern species, individuals that were taller at the commencement of the study had greater overall growth for the duration of the study. This effect did not decrease even among the tallest tree ferns in our study (up to 6 metres tall). Overall, Cyathea australis averaged 73 (± 22)mm/year of growth (± 1SD), with the rate increasing 5mm/yr per metre of additional height. Dicksonia antarctica averaged 33 (± 13)mm/year, increasing by 6mm/yr/m. Growth rates dependent on initial height were unexpected and we discuss possible reasons for this finding. Variable growth rates also suggest that common age estimation methods of dividing height by average growth rate are likely to underestimate the age of short tree ferns, while overestimating the age of tall tree ferns, particularly if they have been subject to a fire.

  12. Non-linear growth in tree ferns, Dicksonia antarctica and Cyathea australis.

    Directory of Open Access Journals (Sweden)

    David P Blair

    Full Text Available Tree ferns are an important structural component of forests in many countries. However, because their regeneration is often unrelated to major disturbances, their age is often difficult to determine. In addition, rates of growth may not be uniform, which further complicates attempts to determine their age. In this study, we measured 5 years of growth of Cyathea australis and Dicksonia antarctica after a large wildfire in 2009 in south-eastern Australia. We found growth rates of these two species were unaffected by aspect and elevation but slope had a minor effect with D. antarctica growing 0.3mm faster for each additional degree of slope. Geographic location influenced growth in both species by up to 12 - 14mm/yr. The most consistent factor influencing growth rate, however, was initial height at the time of the 2009 fire; a finding consistent in both species and all geographic locations. For both tree fern species, individuals that were taller at the commencement of the study had greater overall growth for the duration of the study. This effect did not decrease even among the tallest tree ferns in our study (up to 6 metres tall. Overall, Cyathea australis averaged 73 (± 22mm/year of growth (± 1SD, with the rate increasing 5mm/yr per metre of additional height. Dicksonia antarctica averaged 33 (± 13mm/year, increasing by 6mm/yr/m. Growth rates dependent on initial height were unexpected and we discuss possible reasons for this finding. Variable growth rates also suggest that common age estimation methods of dividing height by average growth rate are likely to underestimate the age of short tree ferns, while overestimating the age of tall tree ferns, particularly if they have been subject to a fire.

  13. Using Lava Tube Skylight Thermal Emission Spectra to Determine Lava Composition on Io: Quantitative Constraints for Observations by Future Missions to the Jovian System.

    Science.gov (United States)

    Davies, A. G.

    2008-12-01

    Deriving the composition of Io's dominant lavas (mafic or ultramafic?) is a major objective of the next missions to the jovian system. The best opportunities for making this determination are from observations of thermal emission from skylights, holes in the roof of a lava tube through which incandescent lava radiates, and Io thermal outbursts, where lava fountaining is taking place [1]. Allowing for lava cooling across the skylight, the expected thermal emission spectra from skylights of different sizes have been calculated for laminar and turbulent tube flow and for mafic and ultramafic composition lavas. The difference between the resulting mafic and ultramafic lava spectra has been quantified, as has the instrument sensitivity needed to acquire the necessary data to determine lava eruption temperature, both from Europa orbit and during an Io flyby. A skylight is an excellent target to observe lava that has cooled very little since eruption (California Institute of Technology, under contract to NASA. AGD is supported by a grant from the NASA OPR Program. References: [1] Davies, A. G., 1996, Icarus, 124, 45-61. [2] Keszthelyi, L., et al., 2006, JGS, 163, 253-264. [3] Davies, A. G., 2007, Volcanism on Io, Cambridge University Press. [4] Keszthelyi, L., et al., 2007, Icarus, 192, 491-502. [5] Davies, A. G., et al., 2006, Icarus, 184, 460-477.

  14. Study of aerosols collected in a speleotherapeutic cave situated below Budapest, Hungary

    Energy Technology Data Exchange (ETDEWEB)

    Kertesz, Zs. E-mail: zsofi@moon.atomki.hu; Borbely-Kiss, I.; Hunyadi, I

    1999-04-02

    The Szemlohegy-cave is one of the well-known hydrothermal caves of the Rozsadomb area of Budapest, which have been used for speleotherapy of respiratory diseases for years. It is known from the periodically changing airborne radon activity concentration data, that airflow of seasonally reversed direction are formed along the cave passages and fissures due to the temperature difference between the surface and cave air. This means that an intensive interaction takes place between the cave and its environment. The pollution of nearby waters and the urban atmospheric air represents a real danger for these caves below Buda, which recently became the part of the UNESCO World Heritage. The study of cave aerosols should be very important from the point of view of either the control possibilities of the environmental impact or speleotherapy, and probably helps in getting acquainted with the cave-forming processes. In this work we applied our standard aerosol sampling method to the high-humidity environment of the caves, and we studied the elemental composition, size fractionation as well as the spatial distribution and the seasonal variation of cave aerosols. Thanks to the sensitivity of PIXE traces of anthropogenic pollution of the Budapest air are shown in the Szemlohegy-cave. Measured elemental concentrations remained less than one-tenth the air quality standard valid for the increasingly protected areas.

  15. Study of aerosols collected in a speleotherapeutic cave situated below Budapest, Hungary

    International Nuclear Information System (INIS)

    Kertesz, Zs.; Borbely-Kiss, I.; Hunyadi, I.

    1999-01-01

    The Szemlohegy-cave is one of the well-known hydrothermal caves of the Rozsadomb area of Budapest, which have been used for speleotherapy of respiratory diseases for years. It is known from the periodically changing airborne radon activity concentration data, that airflow of seasonally reversed direction are formed along the cave passages and fissures due to the temperature difference between the surface and cave air. This means that an intensive interaction takes place between the cave and its environment. The pollution of nearby waters and the urban atmospheric air represents a real danger for these caves below Buda, which recently became the part of the UNESCO World Heritage. The study of cave aerosols should be very important from the point of view of either the control possibilities of the environmental impact or speleotherapy, and probably helps in getting acquainted with the cave-forming processes. In this work we applied our standard aerosol sampling method to the high-humidity environment of the caves, and we studied the elemental composition, size fractionation as well as the spatial distribution and the seasonal variation of cave aerosols. Thanks to the sensitivity of PIXE traces of anthropogenic pollution of the Budapest air are shown in the Szemlohegy-cave. Measured elemental concentrations remained less than one-tenth the air quality standard valid for the increasingly protected areas

  16. Profiling bacterial diversity in a limestone cave of the western Loess Plateau of China

    Directory of Open Access Journals (Sweden)

    Yucheng eWu

    2015-03-01

    Full Text Available Bacteria and archaea sustain subsurface cave ecosystems by dominating primary production and fueling biogeochemical cyclings, despite the permanent darkness and shortage of nutrients. However, the heterogeneity and underlying mechanism of microbial diversity in caves, in particular those well connect to surface environment are largely unexplored. In this study, we examined the bacterial abundance and composition in Jinjia Cave, a small and shallow limestone cave located on the western Loess Plateau of China, by enumerating and pyrosequencing small subunit (SSU rRNA genes. The results clearly reveal the contrasting bacterial community compositions in relation to cave habitat types, i.e., rock wall deposit, aquatic sediment and sinkhole soil, which are differentially connected to the surface environment. The deposits on the cave walls were dominated by putative cave-specific bacterial lineages within the -Proteobacteria or Actinobacteria that are routinely found on cave rocks around the world. In addition, sequence identity with known functional groups suggests enrichments of chemolithotrophic bacteria potentially involved in autotrophic C fixation and inorganic N transformation on rock surfaces. By contrast, bacterial communities in aquatic sediments were more closely related to those in the overlying soils. This is consistent with the similarity in elemental composition between the cave sediment and the overlying soil, implicating the influence of mineral chemistry on cave microhabitat and bacterial composition. These findings provide compelling molecular evidence of the bacterial community heterogeneity in an East Asian cave, which might be controlled by both subsurface and surface environments.

  17. Characterization and management trends of negative and positive impacts of tourism in show caves

    Directory of Open Access Journals (Sweden)

    Heros Augusto Santos Lobo

    2015-08-01

    Full Text Available Caves are one of the most fragile environments on Earth ’s surface, in function of physical, atmospheric and biological specificities. The natural characteristics associated with cultural aspects of local people are the reasons to tourist potential of caves. However, the tourist use of caves can generate negative impacts on the environment. This consequence happens when inappropriate procedures of planning and management of tourism are used. Negative impacts are described in this review, as also indications of management techniques to avoid, decrease or eliminate the negative consequences of tourism in caves. Positive impacts are also presented, in order to achieve a sustainable tourism in show caves. The conclusion argues that the negative impacts should not be considered as impediments to tourism in caves. It is important to know these impacts and use it as a key to get previous answers which allow to raise the sustainability of show caves.

  18. The architectural form of Qikou Cave dwellings in Chinese "Earth" culture

    Science.gov (United States)

    Chen, Xuanchen; Feng, Xinqun

    2018-03-01

    Cave building is not only a kind of architecture with unique style, but also a manifestation of Chinese traditional culture. Cave culture is an important part of Chinese traditional culture. The main purpose of this thesis which studies the architectural form of Qikou Cave, is to analyze how the cave building plays a positive role in promoting the development and application of modern resources and in cultural transmission. Based on a large amount of literature material, and taking Qikou Cave as an example, by studying the morphological characteristics of cave building, the paper takes an optimistic outlook on its future development and the sustainable development of the resources. It is expected that the cave culture can be further explored to promote the traditional Chinese culture and to drive the development of modern construction industry and resource conservation.

  19. Main karst and caves of Switzerland; El karst y las cuevas mas importantes de Suiza

    Energy Technology Data Exchange (ETDEWEB)

    Jeannin, P. Y.

    2016-07-01

    This paper presents an overview of the main karst areas and cave systems in Switzerland. The first part encloses descriptions of the main geological units that hold karst and caves in the country and summarizes a brief history of research and protection of the cave environments. The second part presents three regions enclosing large cave systems. Two regions in the Alps enclose some of the largest limestone caves in Europe: Siebenhengste (Siebenhengste cave system with ∼160 km and Barenschacht with 70 km) and Bodmeren-Silberen (Holloch cave system with 200 km and Silberen System with 39 km). These systems are also among the deepest with depths ranging between 880 and 1340 m. The third example is from the Jura Mountains (northern Switzerland). (Author)

  20. Blow Hole Cave: An unroofed cave on San Salvador Island, the Bahamas, and its importance for detection of paleokarst caves on fossil carbonate platforms

    Czech Academy of Sciences Publication Activity Database

    Bosák, Pavel; Mylroie, J. E.; Hladil, Jindřich; Carew, J. L.; Slavík, Ladislav

    2002-01-01

    Roč. 31, č. 3 (2002), s. 51-74 ISSN 0583-6050. [Karstological School - Classical Karst: Types of Karst /10./. Pstojna, 26.06.2002-28.06.2002] R&D Projects: GA AV ČR IAA3013809; GA AV ČR IAA3013209 Keywords : carbonate platforms * unroofed caves * gamma-ray spectrometry and wellogging Subject RIV: DB - Geology ; Mineralogy http://carsologica.zrc-sazu.si/downloads/313/slavik.pdf

  1. Bioaccumulation of eight heavy metals in cave animals from Dashui ...

    African Journals Online (AJOL)

    The results showed that zinc contents in Porcellio scaber from Dashui and Malang caves were 448.80 and 598.00 mg/kg, respectively, which is the highest among all these 8 metals, while Pb was not detected in Diestrammena marmorata and Rhinolophidae pearsoni, suggesting that both animals were incapable of or poor ...

  2. Mineral magnetic environmental record in clastic cave deposits

    Czech Academy of Sciences Publication Activity Database

    Šlechta, Stanislav; Kadlec, Jaroslav

    2008-01-01

    Roč. 38, special issue (2008), s. 134-134 ISSN 1335-2806. [Paleo, Rock and Environmental Magnetism. Castle Meeting /11./. 22.06.2008-28.06.2008, Bojnice] Institutional research plan: CEZ:AV0Z30130516 Keywords : mineral magnetic environmental reconstruction * cave sediments * Moravian Karst Subject RIV: DE - Earth Magnetism, Geodesy, Geography

  3. Study of firedamp release in sub-level caving

    Energy Technology Data Exchange (ETDEWEB)

    Castano, F.S.; Sanz Delgado, M.A.; Iturriaga Trenor, S.; Alberdi Vinas, C.

    1988-01-01

    The field work developed in the pits of 'Hullera Vasco-Leonesa' in Santa Lucia (Leon) to control firedamp release is described. Conclusions obtained concerning gas emission, irregularity and firedamp balance in sub-level caving winning faces are discussed. Some definitions of basic concepts in firedamp studies are also included. 7 figs.

  4. Full Immersive Virtual Environment Cave[TM] in Chemistry Education

    Science.gov (United States)

    Limniou, M.; Roberts, D.; Papadopoulos, N.

    2008-01-01

    By comparing two-dimensional (2D) chemical animations designed for computer's desktop with three-dimensional (3D) chemical animations designed for the full immersive virtual reality environment CAVE[TM] we studied how virtual reality environments could raise student's interest and motivation for learning. By using the 3ds max[TM], we can visualize…

  5. Inside the neutrino cave, close to the target complex

    CERN Multimedia

    CERN PhotoLab

    1976-01-01

    The photo shows on the left the shielding of the target complex, T9 and T11 for the wide and narrow beams. The direction of the primary proton beam faces the camera. Between the shielding and the cave wall are housed the magnets cooling pipes. The pulley block allows displacements inside the shielding.

  6. Incidence of chromosomal aberrations and micronuclei in cave tour guides.

    Science.gov (United States)

    Bilban, M; Bilban-Jakopin, C; Vrhovec, S

    2001-01-01

    An analysis of structural chromosomal aberrations (SCA) and micronucleus tests (MN) were performed in 38 subjects, cave tour guides and in appropriate control group. The dominant type of chromosomal aberrations in tourist guides were chromosomal breaks (0.013 per cell) and acentric fragments (0.011 per cell). In the control group, these aberrations were present up to 0.008 on cells. Considering the analysed cells of the guides in total (33,556), the incidence of dicentric and rings range is below 0.0008 on cells, even though three dicentric and ring chromosoms were found already in the first 1000 in vitro metaphases of some guides. Only 0.0003 dicentrics and neither other translocations were found in control group (ambiental exposure). The incidence of micronuclei in cytokinesis blocked lymphocytes ranged from 12-32 per 500 CB cells in the cave tour guides and from 4-11 per 500 CB cells in control group. Measurements of radon and its daughters were performed at different locations in the cave. Annual doses from 40-60 mSv were estimated per 2000 work hours for cave guides. The changes found in the genome of somatic cells may be related to the exposure doses of radon and its daughters, although smoking should not be ignored.

  7. Radon in Austrian tourist mines and show caves

    International Nuclear Information System (INIS)

    Ringer, W.; Graeser, J.

    2009-01-01

    The radon situation in tourist mines and show caves is barely investigated in Austria. This paper investigates the influence of its determining factors, such as climate, structure and geology. For this purpose, long-term time-resolved measurements over 6 to 12 months in 4 tourist mines and 2 show caves - with 5 to 9 measuring points each - have been carried out to obtain the course of radon concentration throughout the year. In addition, temperature and air-pressure were measured and compared to the data outside where available. Results suggest that the dominating factors of the average radon concentration are structure and location (geology) of the tunnel-system, whereas the diurnal and annual variation is mainly caused by the changing airflow, which is driven by the difference in temperature inside and outside. Downcast air is connected with very low radon concentrations, upcast air with high concentrations. In some locations the maximum values appear when the airflow ceases. But airflow can be different in different parts of mines and caves. Systems close to the surface show generally lower radon levels than the ones located deeper underground. Due to variation of structure, geology and local climate, the radon situation in mines and caves can only be described by simultaneous measurements at several measuring points. (orig.)

  8. Preliminary study of radioactive waste disposal in granitic underground caves

    International Nuclear Information System (INIS)

    Carvalho, J.F. de; Carajilescov, P.

    1984-01-01

    To date, the disposal of radioactive wastes is one of the major problems faced by the nuclear industry. The utilization of granitic underground caves surrounded by a clay envelope is suggested as a safe alternative for such disposal. A preliminary analysis of the dimensions of those deposits is done. (Author) [pt

  9. Energy and speleogenesis: Key determinants of terrestrial species richness in caves.

    Science.gov (United States)

    Jiménez-Valverde, Alberto; Sendra, Alberto; Garay, Policarp; Reboleira, Ana Sofia P S

    2017-12-01

    The aim of this study was to unravel the relative role played by speleogenesis (i.e., the process in which a cave is formed), landscape-scale variables, and geophysical factors in the determination of species richness in caves. Biological inventories from 21 caves located in the southeastern Iberian Peninsula along with partial least square (PLS) regression analysis were used to assess the relative importance of the different explanatory variables. The caves were grouped according to the similarity in their species composition; the effect that spatial distance could have on similarity was also studied using correlation between matrices. The energy and speleogenesis of caves accounted for 44.3% of the variation in species richness. The trophic level of each cave was the most significant factor in PLS regression analysis, and epigenic caves (i.e., those formed by the action of percolating water) had significantly more species than hypogenic ones (i.e., those formed by the action of upward flows in confined aquifers). Dissimilarity among the caves was very high (multiple-site β sim  = 0.92). Two main groups of caves were revealed through the cluster analysis, one formed by the western caves and the other by the eastern ones. The significant-but low-correlation found between faunistic dissimilarity and geographical distance ( r  =   .16) disappeared once the caves were split into the two groups. The extreme beta-diversity suggests a very low connection among the caves and/or a very low dispersal capacity of the species. In the region under study, two main factors are intimately related to the richness of terrestrial subterranean species in caves: the amount of organic material (trophic level) and the formation process (genesis). This is the first time that the history of a cave genesis has been quantitatively considered to assess its importance in explaining richness patterns in comparison with other factors more widely recognized.

  10. Palaeoclimate Research in Villars Cave (Dordogne, SW-France

    Directory of Open Access Journals (Sweden)

    Genty Dominique

    2008-10-01

    Full Text Available Villars Cave is a typical shallow cave from South-West France (45.44°N; 0.78°E; 175 m asl that has provided several speleothempalaeoclimatic records such as the millennial scale variability of the Last Glacial period and the Last Deglaciation. Monitoring theVillars cave environment over a 13-year period has helped in the understanding of the stable isotopic speleothem content and inthe hydrology. For example, it was demonstrated that most of the calcite CaCO3 carbon comes from the soil CO2, which explainsthe sensitivity of the δ13C to any vegetation and climatic changes. Drip rate monitoring, carried out under four stalactites from thelower and upper galleries, has shown a well marked seasonality of the seepage water with high flow rates during winter and spring.A time delay of about two months is observed between the water excess (estimated from outside meteorological stations and thedrip rate in the cave. A great heterogeneity in the flow rate amplitude variations and in the annual quantity of water between twonearby stalactites is observed, confirming the complexity of the micro-fissure network system in the unsaturated zone. At a dailyscale, the air pressure and drip rates are anti-correlated probably because of pressure stress on the fissure network. Cave air CO2concentration follows soil CO2 production and is correlated with its δ13C content. Since the beginning of the monitoring, the cave airtemperature, in both lower and upper galleries, displays a warming trend of ~+0.4°C±0.1/10yrs. This might be the consequence ofthe outside temperature increase that reaches the Villars Cave galleries through thermal wave conduction. Chemistry monitoringover a few years has shown that the seepage water of the lower gallery stations is significantly more concentrated in trace and minorelements (i.e. Sr, Mg, Ba, U than the upper stations, probably due to the 10-20 m depth difference between these galleries, whichimplies a different seepage pathway

  11. Lava flooding of ancient planetary crusts: geometry, thickness, and volumes of flooded lunar impact basins

    International Nuclear Information System (INIS)

    Head, J.W.

    1982-01-01

    Estimates of lava volumes on planetary surfaces provide important data on the lava flooding history and thermal evolution of a planet. Lack of information concerning the configuration of the topography prior to volcanic flooding requires the use of a variety of techniques to estimate lava thicknesses and volumes. A technique is described and developed which provides volume estimates by artificially flooding unflooded lunar topography characteristic of certain geological environments, and tracking the area covered, lava thicknesses, and lava volumes. Comparisons of map patterns of incompletely buried topography in these artificially flooded areas are then made to lava-flooded topography on the Moon in order to estimate the actual lava volumes. This technique is applied to two areas related to lunar impact basins; the relatively unflooded Orientale basin, and the Archimedes-Apennine Bench region of the Imbrium basin. (Auth.)

  12. Mapping planetary caves with an autonomous, heterogeneous robot team

    Science.gov (United States)

    Husain, Ammar; Jones, Heather; Kannan, Balajee; Wong, Uland; Pimentel, Tiago; Tang, Sarah; Daftry, Shreyansh; Huber, Steven; Whittaker, William L.

    Caves on other planetary bodies offer sheltered habitat for future human explorers and numerous clues to a planet's past for scientists. While recent orbital imagery provides exciting new details about cave entrances on the Moon and Mars, the interiors of these caves are still unknown and not observable from orbit. Multi-robot teams offer unique solutions for exploration and modeling subsurface voids during precursor missions. Robot teams that are diverse in terms of size, mobility, sensing, and capability can provide great advantages, but this diversity, coupled with inherently distinct low-level behavior architectures, makes coordination a challenge. This paper presents a framework that consists of an autonomous frontier and capability-based task generator, a distributed market-based strategy for coordinating and allocating tasks to the different team members, and a communication paradigm for seamless interaction between the different robots in the system. Robots have different sensors, (in the representative robot team used for testing: 2D mapping sensors, 3D modeling sensors, or no exteroceptive sensors), and varying levels of mobility. Tasks are generated to explore, model, and take science samples. Based on an individual robot's capability and associated cost for executing a generated task, a robot is autonomously selected for task execution. The robots create coarse online maps and store collected data for high resolution offline modeling. The coordination approach has been field tested at a mock cave site with highly-unstructured natural terrain, as well as an outdoor patio area. Initial results are promising for applicability of the proposed multi-robot framework to exploration and modeling of planetary caves.

  13. Reproductive Seasonality in Nesticus (Araneae: Nesticidae) Cave Spiders.

    Science.gov (United States)

    Carver, Linnea M; Perlaky, Patricia; Cressler, Alan; Zigler, Kirk S

    2016-01-01

    Spiders of the family Nesticidae are members of cave communities around the world with cave-obligate (troglobiotic) species known from North America, Europe, Asia and the Indo-Pacific. A radiation of Nesticus (Araneae: Nesticidae) in the southern Appalachians includes ten troglobiotic species. Many of these species are of conservation interest due to their small ranges, with four species being single-cave endemics. Despite conservation concerns and their important role as predators in cave communities, we know little about reproduction and feeding in this group. We addressed this knowledge gap by examining populations of two species on a monthly basis for one year. We made further observations on several other species and populations, totaling 671 individual spider observations. This more than doubled the reported observations of reproduction and feeding in troglobiotic Nesticus. Female Nesticus carry egg sacs, facilitating the determination of the timing and frequency of reproduction. We found that Nesticus exhibit reproductive seasonality. Females carried egg sacs from May through October, with a peak in frequency in June. These spiders were rarely observed with prey; only 3.3% (22/671) of individuals were observed with prey items. The frequency at which prey items were observed did not vary by season. Common prey items were flies, beetles and millipedes. Troglobiotic species constituted approximately half of all prey items observed. This result represents a greater proportion of troglobiotic prey than has been reported for various troglophilic spiders. Although our findings shed light on the life history of troglobiotic Nesticus and on their role in cave ecosystems, further work is necessary to support effective conservation planning for many of these rare species.

  14. Pāhoehoe, `a`ā, and block lava: an illustrated history of the nomenclature

    Science.gov (United States)

    Harris, Andrew J. L.; Rowland, Scott K.; Villeneuve, Nicolas; Thordarson, Thor

    2017-01-01

    Lava flows occur worldwide, and throughout history, various cultures (and geologists) have described flows based on their surface textures. As a result, surface morphology-based nomenclature schemes have been proposed in most languages to aid in the classification and distinction of lava surface types. One of the first to be published was likely the nine-class, Italian-language description-based classification proposed by Mario Gemmellaro in 1858. By far, the most commonly used terms to describe lava surfaces today are not descriptive but, instead, are merely words, specifically the Hawaiian words `a`ā (rough brecciated basalt lava) and pāhoehoe (smooth glassy basalt lava), plus block lava (thick brecciated lavas that are typically more silicic than basalt). `A`ā and pāhoehoe were introduced into the Western geological vocabulary by American geologists working in Hawai`i during the 1800s. They and other nineteenth century geologists proposed formal lava-type classification schemes for scientific use, and most of them used the Hawaiian words. In 1933, Ruy Finch added the third lava type, block lava, to the classification scheme, with the tripartite system being formalized in 1953 by Gordon Macdonald. More recently, particularly since the 1980s and based largely on studies of lava flow interiors, a number of sub-types and transitional forms of all three major lava types have been defined. This paper reviews the early history of the development of the pāhoehoe, `a`ā, and block lava-naming system and presents a new descriptive classification so as to break out the three parental lava types into their many morphological sub-types.

  15. Lava Tubes as Martian Analog sites on Hawaii Island

    Science.gov (United States)

    Andersen, Christian; Hamilton, J. C.; Adams, M.

    2013-10-01

    The existence of geologic features similar to skylights seen in Mars Reconnaissance Orbiter HIRISE imagery suggest Martian lava tube networks. Along with pit craters, these features are evidence of a past era of vulcanism. If these were contemporary with the wet Mars eras, then it is suggestive that any Martian life may have retreated into these subsurface oases. Hawaii island has numerous lava tubes of differing ages, humidity, lengths and sizes that make ideal analog test environments for future Mars exploration. PISCES has surveyed multiple candidate sites during the past summer with a team of University of Hawaii at Hilo student interns. It should be noted that Lunar features have also been similarly discovered via Lunar Reconnaissance Orbiter LROC imagery.

  16. Numerical simulation of lava flow using a GPU SPH model

    Directory of Open Access Journals (Sweden)

    Eugenio Rustico

    2011-12-01

    Full Text Available A smoothed particle hydrodynamics (SPH method for lava-flow modeling was implemented on a graphical processing unit (GPU using the compute unified device architecture (CUDA developed by NVIDIA. This resulted in speed-ups of up to two orders of magnitude. The three-dimensional model can simulate lava flow on a real topography with free-surface, non-Newtonian fluids, and with phase change. The entire SPH code has three main components, neighbor list construction, force computation, and integration of the equation of motion, and it is computed on the GPU, fully exploiting the computational power. The simulation speed achieved is one to two orders of magnitude faster than the equivalent central processing unit (CPU code. This GPU implementation of SPH allows high resolution SPH modeling in hours and days, rather than in weeks and months, on inexpensive and readily available hardware.

  17. Phylogeny of Marsileaceous Ferns and Relationships of the Fossil Hydropteris pinnata Reconsidered.

    Science.gov (United States)

    Pryer

    1999-09-01

    Recent phylogenetic studies have provided compelling evidence that confirms the once disputed hypothesis of monophyly for heterosporous leptosporangiate ferns (Marsileaceae and Salviniaceae). Hypotheses for relationships among the three genera of Marsileaceae (Marsilea, Regnellidium, and Pilularia), however, have continued to be in conflict. The phylogeny of Marsileaceae is investigated here using information from morphology and rbcL sequence data. In addition, relationships among all heterosporous ferns, including the whole-plant fossil Hydropteris pinnata are reconsidered. Data sets of 71 morphological and 1239 rbcL characters for 23 leptosporangiate ferns, including eight heterosporous ingroup taxa and 15 homosporous outgroup taxa, were subjected to maximum parsimony analysis. Morphological analyses were carried out both with and without the fossil Hydropteris, and it was excluded from all analyses with rbcL data. An annotated list of the 71 morphological characters is provided in the appendix. For comparative purposes, the Rothwell and Stockey (1994) data set was also reanalyzed here. The best estimate of phylogenetic relationships for Marsileaceae in all analyses is that Pilularia and Regnellidium are sister taxa and Marsilea is sister to that clade. Morphological synapomorphies for various nodes are discussed. Analyses that included Hydropteris resulted in two most-parsimonious trees that differ only in the placement of the fossil. One topology is identical to the relationship found by Rothwell and Stockey (1994), placing the fossil sister to the Azolla plus Salvinia clade. The alternative topology places Hydropteris as the most basal member of the heterosporous fern clade. Equivocal interpretations for character evolution in heterosporous ferns are discussed in the context of these two most-parsimonious trees. Because of the observed degree of character ambiguity, the phylogenetic placement of Hydropteris is best viewed as unresolved, and recognition of the

  18. Isolation of polymorphic microsatellite markers and tests of cross-amplification in four widespread European calcicole ferns

    NARCIS (Netherlands)

    Groot, de G.A.; Korpelainen, H.; Wubs, E.R.J.; Erkens, R.H.J.

    2011-01-01

    Premise of the study: Studies on the biogeography and population genetics of the widespread European rock ferns Asplenium scolopendrium , A. trichomanes subsp. quadrivalens , Polystichum setiferum , and P. aculeatum would potentially yield interesting new insights into the colonization capacities of

  19. Calculated viscosity-distance dependence for some actively flowing lavas

    International Nuclear Information System (INIS)

    Pieri, D.

    1987-01-01

    The importance of viscosity as a gauge of the various energy and momentum dissipation regimes of lava flows has been realized for a long time. Nevertheless, despite its central role in lava dynamics and kinematics, it remains among the most difficult of flow physical properties to measure in situ during an eruption. Attempts at reconstructing the actual emplacement viscosities of lava flows from their solidified topographic form are difficult. Where data are available on the position of an advancing flow front as a function of time, it is possible to calculate the effective viscosity of the front as a function of distance from the vent, under the assumptions of a steady state regime. As an application and test of an equation given, relevant parameters from five recent flows on Mauna Loa and Kilauea were utilized to infer the dynamic structure of their aggregate flow front viscosity as they advanced, up to cessation. The observed form of the viscosity-distance relation for the five active Hawaiian flows examined appears to be exponential, with a rapid increase just before the flows stopped as one would expect

  20. Wisata Bencana : Sebuah Studi Kasus Lava Tour Gunung Merapi

    Directory of Open Access Journals (Sweden)

    Zein Mufarrih Muktaf

    2017-09-01

    ABSTRACK The emergence of ecotourism trends as part of nature tourism to be an offer for tourists who want to feel the sensation of different tourist. In addition to the emergence of ecotourism, also appeared many other sort of tourism, such as dark tourism and disaster tourism. Dark tourism and disaster tourism is interesting enough to be discussion. The quention of this research is how the phenomenon of disaster tourism on Lava Tour in Mount Merapi? The purpose of this research is to know the practice of disaster tour “Lava Tour” Mount Merapi. The object of research is community-based tourism in Lava Tour area located in Disaster Prone Area (Kawasan Rawan Bencana III. Research method using case study approach. The conclusion of this research is, first, disaster tour is educational tour which destruction, death and back a life as tourist attraction. Secondly, that disaster tour presents a trip or tour because tourists can direct to see the disaster site. Third, the role of communication between the community-based tourism to the tourists are very important, such as telling the chronology of events to the tourists. It is better if the source of information teller is a direct victim or a direct eye witness, because it is more authentic and convincing. Fourth, disaster tourism prefers the interaction between witnesses and tourists. Fifth, disaster tours can be part of disaster literacy, as witnesses or victims explain a lot about disaster. Keywod : disaster tourism; tour; Mount Merapi; Tourism Communication; disaster literacy

  1. Modeling risk assessment for nuclear processing plants with LAVA

    International Nuclear Information System (INIS)

    Smith, S.T.; Tisinger, R.M.

    1988-01-01

    Using the Los Alamos Vulnerability and Risk Assessment (LAVA) methodology, the authors developed a model for assessing risks associated with nuclear processing plants. LAVA is a three-part systematic approach to risk assessment. The first part is the mathematical methodology; the second is the general personal computer-based software engine; and the third is the application itself. The methodology provides a framework for creating applications for the software engine to operate upon; all application-specific information is data. Using LAVA, the authors build knowledge-based expert systems to assess risks in applications systems comprising a subject system and a safeguards system. The subject system model is sets of threats, assets, and undesirable outcomes. The safeguards system model is sets of safeguards functions for protecting the assets from the threats by preventing or ameliorating the undesirable outcomes, sets of safeguards subfunctions whose performance determine whether the function is adequate and complete, and sets of issues, appearing as interactive questionnaires, whose measures (in both monetary and linguistic terms) define both the weaknesses in the safeguards system and the potential costs of an undesirable outcome occurring

  2. Palæomagnetism of Hawaiian lava flows

    Science.gov (United States)

    Doell, Richard R.; Cox, Allan

    1961-01-01

    PALÆOMAGNETIC investigations of volcanic rocks extruded in various parts of the world during the past several million years have generally revealed a younger sequence of lava flows magnetized nearly parallel to the field of a theoretical geocentric axial dipole, underlain by a sequence of older flows with exactly the opposite direction of remanent magnetization. A 180-degree reversal of the geomagnetic field, occurring near the middle of the Pleistocene epoch, has been inferred by many workers from such results1–3. This is a preliminary report of an investigation of 755 oriented samples collected from 152 lava flows on the island of Hawaii, selected to represent as many stratigraphic horizons as possible. (Sampling details are indicated in Table 1.) This work was undertaken because Hawaii's numerous thick sequences of lava flows, previously mapped as Pliocene to Historic by Stearns and Macdonald4, and afterwards assigned ages ranging from later Tertiary to Recent, by Macdonald and Davis5, appeared to offer an ideal opportunity to examine the most recent reversal of Earth's field.

  3. LAVA: a conceptual framework for automated risk assessment

    International Nuclear Information System (INIS)

    Smith, S.T.; Brown, D.C.; Erkkila, T.H.; FitzGerald, P.D.; Lim, J.J.; Massagli, L.; Phillips, J.R.; Tisinger, R.M.

    1986-01-01

    At the Los Alamos National Laboratory we are developing the framework for generating knowledge-based systems that perform automated risk analyses on an organization's assets. An organization's assets can be subdivided into tangible and intangible assets. Tangible assets include facilities, materiel, personnel, and time, while intangible assets include such factors as reputation, employee morale, and technical knowledge. The potential loss exposure of an asset is dependent upon the threats (both static and dynamic), the vulnerabilities in the mechanisms protecting the assets from the threats, and the consequences of the threats successfully exploiting the protective systems vulnerabilities. The methodology is based upon decision analysis, fuzzy set theory, natural-language processing, and event-tree structures. The Los Alamos Vulnerability and Risk Assessment (LAVA) methodology has been applied to computer security. LAVA is modeled using an interactive questionnaire in natural language and is fully automated on a personal computer. The program generates both summary reports for use by both management personnel and detailed reports for use by operations staff. LAVA has been in use by the Nuclear Regulatory Commission and the National Bureau of Standards for nearly two years and is presently under evaluation by other governmental agencies. 7 refs

  4. Study of the thermoluminescent properties of lava from different origins

    International Nuclear Information System (INIS)

    Molina, D.; Correcher, V.; Delgado, A.; Garcia G, J.

    2002-01-01

    In this work has been studied the thermoluminescent signal (Tl) of lava from different geographical area (Costa Rica, the Canary Islands, Hawaii, Iceland and Italy) and originating in distinct eruptions, for its possible use such as in the dating field (geological and archaeological) as in retrospective dosimetry. Due that the light emission is intimately related with the punctual defects existent in the structure of material associated to the presence of different mineral phases, it was realized a study by X-ray diffraction for determining the main components of the lava observing the presence, in distinct proportions of cristobalite, plagioclases (chalcosodic feldspars) and philosilicates (augite, montmorillonite, forsterite and actinolite). All the detected mineral components present Tl emission in the blue region. Each one of the lava were artificially irradiated for proving the dependence of the luminescent signal with the dose in the range 1 to 25 Gy, observing a linear response with the dose in all the cases and not appreciating saturation in the Tl emission. Such the appropriate signal of natural samples (TLN) as the irradiated samples in the laboratory (TLI) show a complex structure associated with a continuous distribution of traps at temperature higher than 100 C which could be explained as consequence of the dynamic formation-annihilation of centers [AlO 4 /alkali] + and [AlO 4 ] 0 . In TLI was observed that a nearer to 85 C appeared a maximum whose structure correspond a discrete distribution of traps, coexisting therefore the two types of traps structure. (Author)

  5. Evolution and classification of Elaphoglossum and Asplenium ferns on Cuba, and discovery of a Miocene Elaphoglossum in Dominican amber

    OpenAIRE

    Lóriga Piñero, Josmaily

    2018-01-01

    This dissertation deals with the systematics and evolution of Neotropical ferns of the genera Elaphoglossum and Asplenium, with particular focus on the species of Cuba and the West Indies. It also includes an analysis and description of an Elaphoglossum frond fragment preserved in Miocene Dominican amber. The worldwide genera Elaphoglossum with 600 species and Asplenium with 685 species are the most species-rich groups of leptosporangiate ferns. On Cuba, Elaphoglossum has 34 species and Asple...

  6. Ferns are less dependent on passive dilution by cell expansion to coordinate leaf vein and stomatal spacing than angiosperms.

    Directory of Open Access Journals (Sweden)

    Madeline R Carins Murphy

    Full Text Available Producing leaves with closely spaced veins is a key innovation linked to high rates of photosynthesis in angiosperms. A close geometric link between veins and stomata in angiosperms ensures that investment in enhanced venous water transport provides the strongest net carbon return to the plant. This link is underpinned by "passive dilution" via expansion of surrounding cells. However, it is not known whether this 'passive dilution' mechanism is present in plant lineages other than angiosperms and is another key feature of the angiosperms' evolutionary success. Consequently, we sought to determine whether the 'passive dilution' mechanism is; (i exclusive to the angiosperms, (ii a conserved mechanism that evolved in the common ancestor of ferns and angiosperms, or (iii has evolved continuously over time. To do this we first we assessed the plasticity of vein and stomatal density and epidermal cell size in ferns in response to light environment. We then compared the relationships between these traits found among ferns with modelled relationships that assume vein and stomatal density respond passively to epidermal cell expansion, and with those previously observed in angiosperms. Vein density, stomatal density and epidermal cell size were linked in ferns with remarkably similar relationships to those observed in angiosperms, except that fern leaves had fewer veins per stomata. However, plasticity was limited in ferns and stomatal spacing was dependent on active stomatal differentiation as well as passive cell expansion. Thus, ferns (like angiosperms appear to coordinate vein and stomatal density with epidermal cell expansion to some extent to maintain a constant ratio between veins and stomata in the leaf. The different general relationships between vein density and stomatal density in ferns and angiosperms suggests the groups have different optimum balances between the production of vein tissue dedicated to water supply and stomatal tissue for gas

  7. A comparison of alpha and beta diversity patterns of ferns, bryophytes and macrolichens in tropical montane forests of southern Ecuador

    OpenAIRE

    Mandl, N A; Lehnert, M; Kessler, M; Gradstein, S R

    2010-01-01

    We present a first comparison of patterns of alpha and beta diversity of ferns, mosses, liverworts and macrolichens in neotropical montane rainforests, and explore the question whether specific taxa may be used as surrogates for others. In three localities in southern Ecuador, we surveyed terrestrial and epiphytic species assemblages in ridge and slope forests in 28 plots of 400 m² each. The epiphytic habitat was significantly richer in ferns, liverworts, and macrolichens than the terrestrial...

  8. Historical reconstruction of climatic and elevation preferences and the evolution of cloud forest-adapted tree ferns in Mesoamerica

    OpenAIRE

    Sosa, Victoria; Ornelas, Juan Francisco; Ram?rez-Barahona, Santiago; G?ndara, Etelvina

    2016-01-01

    Background Cloud forests, characterized by a persistent, frequent or seasonal low-level cloud cover and fragmented distribution, are one of the most threatened habitats, especially in the Neotropics. Tree ferns are among the most conspicuous elements in these forests, and ferns are restricted to regions in which minimum temperatures rarely drop below freezing and rainfall is high and evenly distributed around the year. Current phylogeographic data suggest that some of the cloud forest-adapted...

  9. Magnetic Susceptibility and Heavy Metals in Guano from South Sulawesi Caves

    Science.gov (United States)

    Rifai, H.; Putra, R.; Fadila, M. R.; Erni, E.; Wurster, C. M.

    2018-04-01

    Measurement of some magnetic properties have been performed on vertical profile from South Sulawesi caves (Mampu and Bubau) by using low cost, rapid, sensitive and non destructive magnetic method. The aim is to attempt to use magnetic characters as a fingerprint for anthropogenic pollution in the caves. Guano samples were collected every 5 cm at a certain section of Mampu and Bubau cave, South Sulawesi, starting from surface through 300 cm in depth of mampu Cave and 30 cm of Bubau Cave. The magnetic parameters such as magnetic susceptibility and percentage frequency dependence susceptibility were measured using the Bartington MS2-MS2B instruments and supported by X-Ray Fluoroscence (XRF) to know their element composition. The results show that the samples had variations in magnetic susceptibility from 3.5 to 242.6 x 10‑8 m3/kg for Mampu Cave and from 8.6 to 106.5 x 10‑8 m3/kg for Bubau Cave and also magnetic domain. Then, the XRF results show that the caves contain several heavy metals. Magnetic and heavy metal analyses showing that the magnetic minerals in caves are lithogenic (Fe-bearing minerals) in origin and anthropogenic (Zn content) in the caves.

  10. Observations of obsidian lava flow emplacement at Puyehue-Cordón Caulle, Chile

    Science.gov (United States)

    Tuffen, H.; Castro, J. M.; Schipper, C. I.; James, M. R.

    2012-04-01

    The dynamics of obsidian lava flow emplacement remain poorly understood as active obsidian lavas are seldom seen. In contrast with well-documented basaltic lavas, we lack observational data on obsidian flow advance and temporal evolution. The ongoing silicic eruption at Puyehue-Cordón Caulle volcanic complex (PCCVC), southern Chile provides an unprecedented opportunity to witness and study obsidian lava on the move. The eruption, which started explosively on June 4th 2011, has since June 20 generated an active obsidian flow field that remains active at the time of writing (January 2012), with an area of ~6 km2, and estimated volume of ~0.18 km3. We report on observations, imaging and sampling of the north-western lava flow field on January 4th and 10th 2012, when vent activity was characterised by near-continuous ash venting and Vulcanian explosions (Schipper et al, this session) and was simultaneously feeding the advancing obsidian flow (Castro et al, this session). On January 4th the north-western lava flow front was characterised by two dominant facies: predominant rubbly lava approximately 30-40 m thick and mantled by unstable talus aprons, and smoother, thinner lobes of more continuous lava ~50 m in length that extended roughly perpendicular to the overall flow direction, forming lobes that protrude from the flow margin, and lacked talus aprons. The latter lava facies closely resembled squeeze-up structures in basaltic lava flows[1] and appeared to originate from and overlie the talus apron of the rubbly lava. Its upper surface consisted of smooth, gently folded lava domains cut by crevasse-like tension gashes. During ~2 hours of observation the squeeze-up lava lobe was the most frequent location of small-volume rockfalls, which occurred at ~1-10 minute intervals from the flow front and indicated a locus of lava advance. On January 10th the squeeze-up lava lobes had evolved significantly, with disruption and breakage of smooth continuous lava surfaces to form

  11. Speleogenesis, geometry, and topology of caves: A quantitative study of 3D karst conduits

    Science.gov (United States)

    Jouves, Johan; Viseur, Sophie; Arfib, Bruno; Baudement, Cécile; Camus, Hubert; Collon, Pauline; Guglielmi, Yves

    2017-12-01

    Karst systems are hierarchically spatially organized three-dimensional (3D) networks of conduits behaving as drains for groundwater flow. Recently, geostatistical approaches proposed to generate karst networks from data and parameters stemming from analogous observed karst features. Other studies have qualitatively highlighted relationships between speleogenetic processes and cave patterns. However, few studies have been performed to quantitatively define these relationships. This paper reports a quantitative study of cave geometries and topologies that takes the underlying speleogenetic processes into account. In order to study the spatial organization of caves, a 3D numerical database was built from 26 caves, corresponding to 621 km of cumulative cave passages representative of the variety of karst network patterns. The database includes 3D speleological surveys for which the speleogenetic context is known, allowing the polygenic karst networks to be divided into 48 monogenic cave samples and classified into four cave patterns: vadose branchwork (VB), water-table cave (WTC), looping cave (LC), and angular maze (AM). Eight morphometric cave descriptors were calculated, four geometrical parameters (width-height ratio, tortuosity, curvature, and vertical index) and four topological ones (degree of node connectivity, α and γ graph indices, and ramification index) respectively. The results were validated by statistical analyses (Kruskal-Wallis test and PCA). The VB patterns are clearly distinct from AM ones and from a third group including WTC and LC. A quantitative database of cave morphology characteristics is provided, depending on their speleogenetic processes. These characteristics can be used to constrain and/or validate 3D geostatistical simulations. This study shows how important it is to relate the geometry and connectivity of cave networks to recharge and flow processes. Conversely, the approach developed here provides proxies to estimate the evolution of

  12. Reconciling Mining with the Conservation of Cave Biodiversity: A Quantitative Baseline to Help Establish Conservation Priorities.

    Science.gov (United States)

    Jaffé, Rodolfo; Prous, Xavier; Zampaulo, Robson; Giannini, Tereza C; Imperatriz-Fonseca, Vera L; Maurity, Clóvis; Oliveira, Guilherme; Brandi, Iuri V; Siqueira, José O

    2016-01-01

    Caves pose significant challenges for mining projects, since they harbor many endemic and threatened species, and must therefore be protected. Recent discussions between academia, environmental protection agencies, and industry partners, have highlighted problems with the current Brazilian legislation for the protection of caves. While the licensing process is long, complex and cumbersome, the criteria used to assign caves into conservation relevance categories are often subjective, with relevance being mainly determined by the presence of obligate cave dwellers (troglobites) and their presumed rarity. However, the rarity of these troglobitic species is questionable, as most remain unidentified to the species level and their habitats and distribution ranges are poorly known. Using data from 844 iron caves retrieved from different speleology reports for the Carajás region (South-Eastern Amazon, Brazil), one of the world's largest deposits of high-grade iron ore, we assess the influence of different cave characteristics on four biodiversity proxies (species richness, presence of troglobites, presence of rare troglobites, and presence of resident bat populations). We then examine how the current relevance classification scheme ranks caves with different biodiversity indicators. Large caves were found to be important reservoirs of biodiversity, so they should be prioritized in conservation programs. Our results also reveal spatial autocorrelation in all the biodiversity proxies assessed, indicating that iron caves should be treated as components of a cave network immersed in the karst landscape. Finally, we show that by prioritizing the conservation of rare troglobites, the current relevance classification scheme is undermining overall cave biodiversity and leaving ecologically important caves unprotected. We argue that conservation efforts should target subterranean habitats as a whole and propose an alternative relevance ranking scheme, which could help simplify the

  13. Seasonal Temperature Variations controlling Cave Ventilation Processes in Cueva Larga, Puerto Rico

    Science.gov (United States)

    Winter, A.; Vieten, R.; Warken, S. F.; Schrӧder-Ritzrau, A.; Miller, T. E.; Scholz, D.

    2016-12-01

    Two years of monthly monitoring result in much better understanding of ventilation processes in caves. Cueva Larga, a tropical cave in Puerto Rico is 1440 m long with a large main passage (about 116,000 m3). Cave air pCO2 in the main passage varied seasonally, between 600 ppm in winter and 1800 ppm in summer. The seasonal variability in cave pCO2 made it possible to estimate a cave air exchange time of 36±5 days and a winter ventilation rate of 3,200±800 m3/day for the main cave passage. Calculations of virtual temperature and differences between cave and surface temperature show that the seasonal temperature cycle is the main driver of the alternation between a well-ventilated winter mode and a near-stagnant summer mode. The winter mode is characterized by a positive buoyancy contrast at night leading to maximal cave ventilation, while during summer ventilation is at a minimum. Between winter and summer, a transitional mode of partial cave ventilation is observed. On shorter time scales (diurnal to weekly), cave pCO2 is also influenced by atmospheric pressure but this variation is one order of magnitude lower than the seasonal pCO2 change. The cave morphology of Cueva Larga including its large volume, tubular shape and the obstructed cave entrance geometry are important boundary conditions for the observed ventilation patterns. Our findings emphasize that cave systems with varying morphology have to be studied individually in order to correctly describe ventilation processes.

  14. Ancient lineage, young troglobites: recent colonization of caves by Nesticella spiders.

    Science.gov (United States)

    Zhang, Yuanyuan; Li, Shuqiang

    2013-09-04

    The evolution and origin of cave organisms is a recurring issue in evolutionary studies, but analyses are often hindered by the inaccessibility of caves, morphological convergence, and complex colonization processes. Here we investigated the evolutionary history of Nesticella cave spiders, which are mainly distributed in the Yunnan-Guizhou Plateau, China. With comprehensive sampling and phylogenetic and coalescent-based analyses, we investigated the tempo and mode of diversification and the origins of these troglobites. We also aimed to determine which factors have influenced the diversification of this little-known group. Coalescent-based species delimitation validated the 18 species recognized by morphological inspection and also suggested the existence of cryptic lineages. Divergence time estimates suggested that Nesticella cave spiders in the Yunnan-Guizhou Plateau constituted a monophyletic troglobite clade that originated in the middle Miocene (11.1-18.6 Ma). Although the Yunnan-Guizhou Plateau clade was composed exclusively of troglobite species, suggesting an ancient common subterranean ancestor, we favor multiple, independent cave colonizations during the Pleistocene over a single ancient cave colonization event to explain the origin of these cave faunas. The diversification of plateau Nesticella has been greatly influenced by the sequential uplift of the plateau and likely reflects multiple cave colonizations over time by epigean ancestors during Pleistocene glacial advances. We concluded that plateau cave Nesticella represent an ancient group of spiders, but with young troglobite lineages that invaded caves only recently. The absence of extant epigean relatives and nearly complete isolation among caves supported their relict status. Our work highlights the importance of comprehensive sampling for studies of subterranean diversity and the evolution of cave organisms. The existence of potentially cryptic species and the relict status of Nesticella

  15. Key variables influencing patterns of lava dome growth and collapse

    Science.gov (United States)

    Husain, T.; Elsworth, D.; Voight, B.; Mattioli, G. S.; Jansma, P. E.

    2013-12-01

    Lava domes are conical structures that grow by the infusion of viscous silicic or intermediate composition magma from a central volcanic conduit. Dome growth can be characterized by repeated cycles of growth punctuated by collapse, as the structure becomes oversized for its composite strength. Within these cycles, deformation ranges from slow long term deformation to sudden deep-seated collapses. Collapses may range from small raveling failures to voluminous and fast-moving pyroclastic flows with rapid and long-downslope-reach from the edifice. Infusion rate and magma rheology together with crystallization temperature and volatile content govern the spatial distribution of strength in the structure. Solidification, driven by degassing-induced crystallization of magma leads to the formation of a continuously evolving frictional talus as a hard outer shell. This shell encapsulates the cohesion-dominated soft ductile core. Here we explore the mechanics of lava dome growth and failure using a two-dimensional particle-dynamics model. This meshless model follows the natural evolution of a brittle carapace formed by loss of volatiles and rheological stiffening and avoids difficulties of hour-glassing and mesh-entangelment typical in meshed models. We test the fidelity of the model against existing experimental and observational models of lava dome growth. The particle-dynamics model follows the natural development of dome growth and collapse which is infeasible using simple analytical models. The model provides insight into the triggers that lead to the transition in collapse mechasnism from shallow flank collapse to deep seated sector collapse. Increase in material stiffness due to decrease in infusion rate results in the transition of growth pattern from endogenous to exogenous. The material stiffness and strength are strongly controlled by the magma infusion rate. Increase in infusion rate decreases the time available for degassing induced crystallization leading to a

  16. Sources of sulphate minerals in limestone cave-a possible evidence of anthropogenic activity: a case study in Črna Jama Cave (Slovenia).

    Science.gov (United States)

    Jarc, Simona; Miler, Miloš; Šebela, Stanka; Zupančič, Nina

    2017-12-01

    In the caves, the formation of cave minerals is a consequence of a variety of chemical reactions, some of them also due to human activity. There are many caves in Slovenia, but sulphate minerals are not very often reported and analysed. In this study, the presence of sulphate minerals is detected by SEM/EDS analysis of speleothems from Črna Jama, a cave near Kočevje (southern Slovenia). The cave is characterised by its dark, almost black colour on cave walls, floor and speleothems. Anthropogenic influence in the cave is still visible, including the remains of a fireplace, some inscriptions on the walls and wooden containers. The analyses of some of the black-coated speleothems reveal the presence of calcium sulphate, confirmed by XRD as gypsum. Gypsum crystals are around 50 μm in size, and they occur in thin crusts. Additionally, some rare authigenic baryte crystals a few micrometres in size are detected. The sulphates δ 34 S value in gypsum found on dark coloured speleothems is + 10.4‰ Vienna Canyon Diablo Troilite (VCDT), while the sulphate δ 34 S of the bedrock is + 8.6‰ VCDT. The more likely source of sulphate ions is thus biomass burning rather than bedrock. Also, bedrock and biomass ash are a very probable source of calcium and barium. The highly probable pyrogenous origin of sulphates draws attention to human impact on cave mineralogy.

  17. Disruption of tephra fall deposits caused by lava flows during basaltic eruptions

    Science.gov (United States)

    Brown, R. J.; Thordarson, T.; Self, S.; Blake, S.

    2015-10-01

    Observations in the USA, Iceland and Tenerife, Canary Islands reveal how processes occurring during basaltic eruptions can result in complex physical and stratigraphic relationships between lava and proximal tephra fall deposits around vents. Observations illustrate how basaltic lavas can disrupt, dissect (spatially and temporally) and alter sheet-form fall deposits. Complexity arises through synchronous and alternating effusive and explosive activity that results in intercalated lavas and tephra deposits. Tephra deposits can become disrupted into mounds and ridges by lateral and vertical displacement caused by movement (including inflation) of underlying pāhoehoe lavas and clastogenic lavas. Mounds of tephra can be rafted away over distances of 100 s to 1,000 s m from proximal pyroclastic constructs on top of lava flows. Draping of irregular topography by fall deposits and subsequent partial burial of topographic depressions by later lavas can result in apparent complexity of tephra layers. These processes, deduced from field relationships, have resulted in considerable stratigraphic complexity in the studied proximal regions where fallout was synchronous or alternated with inflation of subjacent lava sheets. These mechanisms may lead to diachronous contact relationships between fall deposits and lava flows. Such complexities may remain cryptic due to textural and geochemical quasi-homogeneity within sequences of interbedded basaltic fall deposits and lavas. The net effect of these processes may be to reduce the usefulness of data collected from proximal fall deposits for reconstructing basaltic eruption dynamics.

  18. Effect of pH on lead removal from water using tree fern as the sorbent.

    Science.gov (United States)

    Ho, Yuh-Shan

    2005-07-01

    The sorption of lead from water onto an agricultural by-product, tree fern, was examined as a function of pH. The sorption processes were carried out using an agitated and baffled system. Pseudo-second-order kinetic analyses were performed to determine the rate constant of sorption, the equilibrium sorption capacity, and the initial sorption rate. Application of the pseudo-second-order kinetics model produced very high coefficients of determination. Results showed the efficiency of tree fern as a sorbent for lead. The optimum pH for lead removal was between 4 and 7, with pH 4.9 resulting in better lead removal. Ion exchange occurred in the initial reaction period. In addition, a relation between the change in the solution hydrogen ion concentration and equilibrium capacity was developed and is presented.

  19. Nona Fernández's Mapocho: Spirits in a Material Wasteland

    Directory of Open Access Journals (Sweden)

    Resha S Cardone

    2015-01-01

    Full Text Available This analysis of Mapocho (2002, a novel by Chilean writer and performer Nona Fernández, explores the significance of the author’s environmentalist representation of Chilean history as the accumulation of spiritual and material contaminants—ghosts and trash—that the victorious from throughout the country’s history have attempted to erase to further various economic and political agendas, particularly the neoliberal model installed during the Pinochet dictatorship. Fernández’s depiction of Chile as an ecological and spiritual wasteland in which the female protagonist (recollects, recycles and reuses the specters and detritus of past conflicts represents the author’s own literary project of advocating for human rights and environmental justice during the Chilean transition while modeling a way to counter the collective amnesia promoted under the newly established democracy.

  20. Rhizospheric metagenome of the terrestrial mangrove fern Acrostichum from Indian Sunderbans

    Directory of Open Access Journals (Sweden)

    Sayak Ganguli

    2017-12-01

    Full Text Available This study reports the analyses of the rhizospheric microbiome of the terrestrial mangrove fern Acrostichum aureum Linn. from the Indian Sunderbans. Samples were collected using standard protocols and 16S rRNA gene V3–V4 region amplicon sequencing was performed to identify the microbial communities prevalent in the rhizosphere. A total of 1,931,252 quality checked reads were assembled into 204,818 contigs and were analysed using QIIME to reveal the abundance of Proteobacteria, Acidobacteria and Planctomycetes. The data is available at the NCBI - Sequence Read Archive with accession number: SRX2660456. This is the first report of the rhizospheric microbiome belonging to a fern species.

  1. Response of nitrogen-fixing water fern Azolla biofertilization to rice crop.

    Science.gov (United States)

    Bhuvaneshwari, K; Singh, Pawan Kumar

    2015-08-01

    The water fern Azolla harbors nitrogen-fixing cyanobacterium Anabaena azollae as symbiont in its dorsal leaves and is known as potent N 2 fixer. Present investigation was carried out to study the influence of fresh Azolla when used as basal incorporation in soil and as dual cropped with rice variety Mahsoori separately and together with and without chemical nitrogen fertilizer in pots kept under net house conditions. Results showed that use of Azolla as basal or dual or basal plus dual influenced the rice crop positively where use of fern as basal plus dual was superior and served the nitrogen requirement of rice. There was marked increase in plant height, number of effective tillers, dry mass and nitrogen content of rice plants with the use of Azolla and N-fertilizers alone and other combinations. The use of Azolla also increased organic matter and potassium contents of the soil.

  2. Fern inventorization in Laiwangi-Wanggameti National Park, East Sumba, Waingapu, NTT

    Directory of Open Access Journals (Sweden)

    I NYOMAN PENENG

    2007-07-01

    Full Text Available The inventory and the collection of the ferns in Laiwangi-Wanggameti National Park will be plamted as collection plants in Bali Botanical Garden. In this research used the explorative method. The result of the research has collected 70 numbers of the ferns and 229 speciment. They consist of 21 family, 30 genus.and 70 species. From 70 species. There are 3 species such as Licopodium sernuum, Lygodium javanicum (Tumb. Sw. and Ophioglossum pendulum L. are predicted as new collection for the Bali Botanic Garden. The dominant genus in Laiwangi- Wanggameti National Park was Cyclosorus, Asplenium, Athyrium, and Pteris. Which are growing to cover the basic of the forest in damp places at the river bank.

  3. Modelling phytoremediation by the hyperaccumulating fern, Pteris vittata, of soils historically contaminated with arsenic.

    Science.gov (United States)

    Shelmerdine, Paula A; Black, Colin R; McGrath, Steve P; Young, Scott D

    2009-05-01

    Pteris vittata plants were grown on twenty-one UK soils contaminated with arsenic (As) from a wide range of natural and anthropogenic sources. Arsenic concentration was measured in fern fronds, soil and soil pore water collected with Rhizon samplers. Isotopically exchangeable soil arsenate was determined by equilibration with (73)As(V). Removal of As from the 21 soils by three sequential crops of P. vittata ranged between 0.1 and 13% of total soil As. Ferns grown on a soil subjected to long-term sewage sludge application showed reduced uptake of As because of high available phosphate concentrations. A combined solubility-uptake model was parameterised to enable prediction of phytoremediation success from estimates of soil As, 'As-lability' and soil pH. The model was used to demonstrate the remediation potential of P. vittata under different soil conditions and with contrasting assumptions regarding re-supply of the labile As pool from unavailable forms.

  4. Screening for Ptaquiloside in Ferns: Using Herbarium Specimens for Qualitative Mapping Purposes.

    Science.gov (United States)

    Rasmussen, Lars Holm; Pedersen, Henrik AErenlund

    2017-11-01

    Ptaquiloside (PTA) is a noxious carcinogen found widespread in Bracken (Pteridium sp.) but with scattered and unresolved distribution outside the genus. The carcinogen causes Bovine Enzootic Haematuria among cattle all-over the World and is under suspicion of causing human cancers. To set-up a methodology for large-scale qualitative studies on the distribution of PTA in ferns using already available herbarium specimens as source. PTA and the main degradation product pterosin B (PtB) were quantified in aqueous frond extracts by HPLC-DAD. PTA was quantified after forced reaction into PtB. Optimal reaction conditions were tested using hydrochloric acid, sulphuric acid, trifluoroacetic acid (TFA), acetic acid and temperature as the experimental variables. A pair-wise test for PTA degradation in plant-press were used to explore the effect of this preservation regime. A selection of up to 50 years old Bracken herbarium specimens were tested for PTA and PtB. The methodology was applied on 21 fern species from Denmark. An optimised TFA-based method results in 30.7% higher conversion and a 1:1 reaction between PTA and PtB. Full three-dimensional resolution of the analyte was obtained. Preservation of fronds in a plant press increase formation of PtB. Hence, the method is only suitable for qualitative studies. Presence of PTA and PtB were found in samples up to 50 years old. Among 21 ferns tested, the compounds were only found in Pteridium aquilinum. Herbarium specimens up to 50 years old can be used for explorative risk assessment of ferns using HPLC-DAD for quantification and identification. Copyright © 2017 John Wiley & Sons, Ltd. Copyright © 2017 John Wiley & Sons, Ltd.

  5. High gene flow in epiphytic ferns despite habitat loss and fragmentation

    OpenAIRE

    Winkler, Manuela; Koch, Marcus; Hietz, Peter

    2011-01-01

    Tropical montane forests suffer from increasing fragmentation and replacement by other types of land-use such as coffee plantations. These processes are known to affect gene flow and genetic structure of plant populations. Epiphytes are particularly vulnerable because they depend on their supporting trees for their entire life-cycle. We compared population genetic structure and genetic diversity derived from AFLP markers of two epiphytic fern species differing in their ability to colonize sec...

  6. Don Andrés Bello y don José Fernández Madrid

    Directory of Open Access Journals (Sweden)

    Gabriel Giraldo Jaramillo

    1959-05-01

    Full Text Available Breve pero intensa fue la amistad de estos dos ilustres grancolombianos durante su residencia en Londres. Fernández Madrid desempeñaba el cargo de Ministro Plenipotenciario de Colombia ante el Gobierno de Su Majestad Británica y Bello estuvo por algún tiempo al frente de la secretaría de dicha misión diplomática.

  7. Non-linear growth in tree ferns, Dicksonia antarctica and Cyathea australis

    OpenAIRE

    Blair, David P.; Blanchard, Wade; Banks, Sam C.; Lindenmayer, David B.

    2017-01-01

    Tree ferns are an important structural component of forests in many countries. However, because their regeneration is often unrelated to major disturbances, their age is often difficult to determine. In addition, rates of growth may not be uniform, which further complicates attempts to determine their age. In this study, we measured 5 years of growth of Cyathea australis and Dicksonia antarctica after a large wildfire in 2009 in south-eastern Australia. We found growth rates of these two spec...

  8. Breeding Systems of Three Tree Ferns : Alsophila firma (Cyatheaceae), Cyathea stipularis (Cyatheaceae), and Lophosoria quadripinnata (Lophosoriaceae)

    OpenAIRE

    DOUGLAS E., SOLTIS; PAMELA S., SOLTIS; ALAN R., SMITH; Department of Botany, Washington State University; Department of Botany, Washington State University; University Herbarium, University of California

    1991-01-01

    Breeding-system data have been available for a large number and diverse array of angiosperms for a relatively long time. In contrast, breeding systems of ferns and their allies (pteridophytes) have only recently been examined, and breeding-system data from natural populations of sporophytes are still lacking for pteridophytes representing many life-history strategies. Few studies, for example, have examined breeding systems of tropical pteridophytes, and no breeding-system data are available ...

  9. Moonraker and Tetris: Japanese Microrovers for Lunar Cave Exploration

    Science.gov (United States)

    Yoshida, K.; Britton, N.; Walker, J.; Shimizu, T.; Tanaka, T.; Hakamada, T.

    2015-10-01

    A Japanese team HAKUTO is developing a robotic system for exploration of Lunar lava tubes. Motivated by Google Lunar XPRIZE that requires 500 m travel on any surface of Moon, but the team plans to go down into a skylight in Lacus Mortis.

  10. Neoendemism in Madagascan scaly tree ferns results from recent, coincident diversification bursts.

    Science.gov (United States)

    Janssen, Thomas; Bystriakova, Nadia; Rakotondrainibe, France; Coomes, David; Labat, Jean-Noël; Schneider, Harald

    2008-08-01

    More than 80% of Madagascar's 12,000 plant species are endemic with the degree of endemism reaching as much as 95% in the scaly tree ferns, an important species rich component of Madagascar's evergreen rainforests. Predominantly African or Asian ancestry and divergence times usually postdating the separation of Madagascar from the Gondwanan landmasses have been demonstrated for several Madagascan animal and angiosperm groups. However, evolutionary studies of rainforest-specific lineages are scarce and the ecological context of radiation events has rarely been investigated. Here, we examine the evolution of Madagascan tree ferns as a rainforest-specific model family, integrate results from bioclimatic niche analysis with a dated phylogenetic framework, and propose an evolutionary scenario casting new light on our knowledge of the evolution of large island endemic clades. We show that Madagascar's extant tree fern diversity springs from three distinct ancestors independently colonizing Madagascar in the Miocene and that these three monophyletic clades diversified in three coincident radiation bursts during the Pliocene, reaching exceptionally high diversification rates and most likely responding to a common climatic trigger. Recent diversification bursts may thus have played a major role in the evolution of the extant Madagascan rainforest biome, which hence contains a significant number of young, neoendemic taxa.

  11. Phototropic leaf movements and photosynthetic performance in an amphibious fern, Marsilea quadrifolia.

    Science.gov (United States)

    Kao, Wen-Yuan; Lin, Bai-Ling

    2010-09-01

    Diurnal phototropism has not been reported in ferns. In this study we found that the four leaflets of the amphibious fern Marsilea quadrifolia are capable of adjusting their leaflet angle and leaflet azimuth in response to changes in the position of the sun's direct beam, exhibiting more diaphototropic movements (orienting the plane of the lamina perpendicular to incident light) in the morning and late afternoon, and more paraphototropic movements (orienting the plane of the lamina parallel to incident light) at noon. In addition, by cutting off the leaflet lamina and covering portions of leaflets with black tape, the junction between the leaflet and petiole was found to be responsible for light reception. Among the light spectrum investigated, blue light was the most effective at inducing diaphototropism. The role of diurnal phototropism in enhancing carbon return and ameliorating photoinhibition was also evaluated. It was concluded that diurnal phototropic leaf movement represents one of the plastic responses enabling this amphibious fern to grow under terrestrial conditions.

  12. Mitigating arsenic contamination in rice plants with an aquatic fern, Marsilea minuta.

    Science.gov (United States)

    Hassi, Ummehani; Hossain, Md Tawhid; Huq, S M Imamul

    2017-10-10

    Dangers of arsenic contamination are well known in human civilization. The threat increases when arsenic is accumulated in food and livestock through irrigated crops or animal food. Hence, it is important to mitigate the effects of arsenic as much as possible. This paper discusses a process for reducing the level of arsenic in different parts of rice plants with an aquatic fern, Marsilea minuta L. A pot experiment was done to study the possibility of using Marsilea minuta as a phytoremediator of arsenic. Rice and Marsilea minuta were allowed to grow together in soils. As a control, Marsilea minuta was also cultured alone in the presence and absence of arsenic (applied at 1 mg/L as irrigation water). We did not find any significant change in the growth of rice due to the association of Marsilea minuta, though it showed a reduction of approximately 58.64% arsenic accumulation in the roots of rice grown with the association of fern compared to that grown without fern. We measured a bioaccumulation factor (BF) of > 5.34, indicating that Marsilea minuta could be a good phytoremediator of arsenic in rice fields.

  13. The accumulation and subcellular distribution of arsenic and antimony in four fern plants.

    Science.gov (United States)

    Feng, R; Wang, X; Wei, C; Tu, S

    2015-01-01

    In the present study, Pteris cretica 'Albo-Lineata' (PC), Pteris fauriei (PF), Humata tyermanii Moore (HT), and Pteris ensiformis Burm (PE), were selected to explore additional plant materials for the phytoremediation of As and Sb co-contamination. To some extent, the addition of As and Sb enhanced the growth of HT, PE, and PF. Conversely, the addition of As and Sb negatively affected the growth of PC and was accompanied with the accumulation of high levels of As and Sb in the roots. The highest concentration of Sb was recorded as 6405 mg kg(-1) in the roots of PC, and that for As was 337 mg kg(-1) in the rhizome of PF. To some degree, As and Sb stimulated the uptake of each other in these ferns. Arsenic was mainly stored in the cytoplasmic supernatant (CS) fraction, followed by the cell wall (CW) fraction. In contrast, Sb was mainly found in the CW fraction and, to a lesser extent, in the CS fraction, suggesting that the cell wall and cytosol play different roles in As and Sb accumulation by fern plants. This study demonstrated that these fern plants show a good application potential in the phytoremediation of As and Sb co-contaminated environments.

  14. The Enrichment and Transfer of Heavy Metals for Two Ferns in Pb-Zn Tailing

    Directory of Open Access Journals (Sweden)

    Mai Jiajie

    2017-01-01

    Full Text Available The enrichment and transfer of 8 heavy metals of Equisetum ramosissimum and Pteris vittata growing naturally close to edge of the sewage pool in Bencun Pb-Zn Tailing, Eastern Guangdong were investigated. The results indicated that the pollution of Cd, Pb, Hg, Zn was very severe in this tailing, followed by Cu and Mn. The potential ecological risk of heavy metals was assessed to be very strong based on soil background values of Guangdong Province and at high risk according to criteria of the second grade State Soil Environmental Quality Standard, and Cd, Hg, Pb were the main factors leading to potential ecological risk. The content of 8 heavy metals in the two ferns did not reach critical content of hyperaccumulator, so neither of them was typical hyperaccumulator, but both had a certain tolerance to these heavy metal pollution. Underground parts of Pteris vittata had an enrichment coefficient above 1 and that of Equisetum ramosissimum had a value near 1, therefore the two ferns could be utilized as potential enrichment plants. The two ferns have strong adaptability to the tailing habitat and can be used as pioneers in ecological restoration of Pb-Zn tailings.

  15. Determination of the Radon Concentration and Radioactivity Level in Karaca Cave

    International Nuclear Information System (INIS)

    Kara, A.

    2008-01-01

    In this study, the radon gas concentration in the Karaca cave which is open to tourism has been determined and the negative effects of radon gas on people were discussed. Karaca cave (Gumushane) is visited by many tourists every year. The measurements of radon gas concentration which affects the health of human beings negatively and even causes the lung cancer when it reaches high points have been done for the summer and winter season. LR-115 passive radon detector was used to determine radon concentrations in the cave both winter and summer season. The average radon concentration in the Karaca cave were determined as 823 Bq/m 3 and 1023 Bq/m 3 for the summer and winter season, respectively. Moreover, to find out the natural radioactivity in the cave, the gamma spectroscopic analysis of soil, stone and stalagmite samples were carried out and their relations with the radon gas in the cave atmosphere was analyzed

  16. Mediterranean Recluse Spider, Loxosceles rufescens (Araneae: Sicariidae from Charkhab Cave, Southern Iran

    Directory of Open Access Journals (Sweden)

    Saber Sadeghi

    2017-04-01

    Full Text Available Background: The best-known dangerous spiders belong to the six genera. The genus Loxosceles or violin spiders are well known for their ability to cause skin necrosis or loxoscelism. All Loxosceles species have medical im­portance due to their necrotizing venom. The present article reports the occurrence of L. rufescens in Charkhab Cave, south of Iran (Larestan.Methods: The specimens were collected from the Charkhab Cave using handling forceps, paintbrush and aspirator and preserved in 96% ethanol.Results: Loxosceles rufescens, a medically important spider, is recorded from Charkhab Cave in Fars Province (southwest of Iran. Identification of L. rufescens was performed based on external morphology and the features of male genitalia.Conclusion: Presence of L. rufescens in south of Iran especially in a cave confirmed that this species is a widely distributed species in Iran. Therefore, cavers or cave visitors should be aware of this poisonous spider in caves.

  17. Geophysical exploration in vicinity of the Unicorn Cave, South Harz Mountains, Germany

    Science.gov (United States)

    Kaufmann, Georg; Romanov, Douchko; Nielbock, Ralf

    2010-05-01

    The Unicorn Cave in the southern Harz Mountains in Germany is located in an outcrop of dolomite from the Zechstein formation, which is underlain by Grauwacke rocks. The cave, about 600 meters long, consists of several large chambers, which are connected by a gallery following the main fault alignments in E/W, NE/SW, and NW/SE direction. The overburden of the cave is shallow, between 10 and 30 m. We have used this cave site to perform a sensitivity test for both gravimetric and geoelectic methods above the cave. Additionally, geoelectic mapping has been used to assess the thickness of the cave sediments in one of the chambers. Our results show a clear signal in the Bouguer anomaly, which can only be explained by a combined model of the void space and the sediment filling. Geoelectric results are less clear, but support the gravimetry.

  18. Evolution and adaptation of marine annelids in interstitial and cave habitats

    DEFF Research Database (Denmark)

    Martinez Garcia, Alejandro

    relatives in cave subterranean ecological refugia. Active colonization and ecological speciation to particular cave niches has been alternatively suggested, but the evaluation of that scenario is obscured by the dominance of crustaceans in anchialine habitats, ecologically similar out and inside caves....... The main goal of this thesis is to explore the evolutionary processes behind colonization and adaptation to submarine cave ecosystems in the Atlantic Ocean using annelids as a model, mainly when they involved ancestrally interstitial forms. In order to do that, we studied selected lineages of annelids...... with cave and interstitial representatives, mainly the families Protodrilidae, Nerillidae, Saccocirridae and Scalibregmatidae. The studies combined the characterization of ancestral and cave habitats, with morphological investigations and phylogenetic analyses founded on extensive taxon coverage...

  19. Measurements of seasonal and daily radon daughter concentration fluctuations in National Park Service caves

    International Nuclear Information System (INIS)

    Yarborough, K.A.

    1977-01-01

    The National Park Service (NPS) is studying levels of airborne alpha radiation from radon and thoron in all NPS administered caves in which tours for visitors are regularly conducted. The NPS research has the dual but complementary objectives of safeguarding health at the NPS administered caves, and to develop data on alpha radiation levels and on natural airflows in NPS caves. The results reported here for NPS caves describe concepts hypothesized for the objectives. In addition the data can be used by various agencies to clarify health standards for exposures to low airborne alpha radiation levels in cave environments. These results show daily and seasonal trends and the influence of natural ventilation by air circulation for each cave investigated

  20. A diverse intrinsic antibiotic resistome from a cave bacterium.

    Science.gov (United States)

    Pawlowski, Andrew C; Wang, Wenliang; Koteva, Kalinka; Barton, Hazel A; McArthur, Andrew G; Wright, Gerard D

    2016-12-08

    Antibiotic resistance is ancient and widespread in environmental bacteria. These are therefore reservoirs of resistance elements and reflective of the natural history of antibiotics and resistance. In a previous study, we discovered that multi-drug resistance is common in bacteria isolated from Lechuguilla Cave, an underground ecosystem that has been isolated from the surface for over 4 Myr. Here we use whole-genome sequencing, functional genomics and biochemical assays to reveal the intrinsic resistome of Paenibacillus sp. LC231, a cave bacterial isolate that is resistant to most clinically used antibiotics. We systematically link resistance phenotype to genotype and in doing so, identify 18 chromosomal resistance elements, including five determinants without characterized homologues and three mechanisms not previously shown to be involved in antibiotic resistance. A resistome comparison across related surface Paenibacillus affirms the conservation of resistance over millions of years and establishes the longevity of these genes in this genus.

  1. Exposure to radon in caves and abandoned mines

    International Nuclear Information System (INIS)

    Dixon, D.W.

    1996-01-01

    The significance for health of exposure to radon daughters during leisure excursions in discussed mines or caves is considered for visitors with interests in speleology, mineral collecting, mining history and youth training. General members of the public might also enter tunnels. Potential annual exposures based on radon measurements in various mines and caves are estimated and it is concluded that the annual exposure of individuals who undertake frequent and prolonged trips to underground systems might exceed 10 6 Bq h m -3 . Exposures to general members of the public are likely to be much lower. The National Radiological Protection Board has developed proposals for a coherent and comprehensive scheme to promote dose limitation in a wide range of circumstances. These are described, with current initiatives on consultation with National Associations, local Government and many special interest groups. (author)

  2. Decommissioning of fuel PIE caves at Berkeley Nuclear Laboratories

    International Nuclear Information System (INIS)

    Brant, A.W.

    1990-01-01

    This paper describes the first major contract awarded to private industry to carry out decommissioning of a facility with significant radiation levels. The work required operatives to work in pressurised suits, entry times were significantly affected by sources of radiation in the Caves, being as low as thirty minutes per day initially. The Caves at Berkeley Nuclear Laboratories carry out post irradiation examination of fuel elements support units and reactor core components from CEGB power stations. The decommissioning work is part of an overall refurbishment of the facility to allow the receipt of AGR Fuel Stringer Component direct from power stations. The paper describes the decommissioning and decontamination of the facility from the remote removal and clean up work carried out by the client to the hands-on work. It includes reference to entry times, work patterns, interfaces with the client and the operations of the laboratory. Details of a specially adapted size reduction method are given. (Author)

  3. The fossil Osmundales (Royal Ferns)—a phylogenetic network analysis, revised taxonomy, and evolutionary classification of anatomically preserved trunks and rhizomes

    OpenAIRE

    Benjamin Bomfleur; Guido W. Grimm; Stephen McLoughlin

    2017-01-01

    The Osmundales (Royal Fern order) originated in the late Paleozoic and is the most ancient surviving lineage of leptosporangiate ferns. In contrast to its low diversity today (less than 20 species in six genera), it has the richest fossil record of any extant group of ferns. The structurally preserved trunks and rhizomes alone are referable to more than 100 fossil species that are classified in up to 20 genera, four subfamilies, and two families. This diverse fossil record constitutes an exce...

  4. The Anatomy of the Blue Dragon: Changes in Lava Flow Morphology and Physical Properties Observed in an Open Channel Lava Flow as a Planetary Analogue

    Science.gov (United States)

    Sehlke, A.; Kobs-Nawotniak, S. E.; Hughes, S. S.; Sears, D. W. G.; Downs, M.; Whittington, A. G.; Lim, D. S. S.; Heldmann, J. L.

    2017-12-01

    Lava terrains on other planets and moons exhibit morphologies similar to those found on Earth, such as smooth pāhoehoe transitioning to rough `a`ā terrains based on the viscosity - strain rate relationship of the lava. Therefore, the morphology of lava flows is governed by eruptive conditions such as effusion rate, underlying slope, and the fundamental thermo-physical properties of the lava, including temperature (T), composition (X), viscosity (η), fraction of crystals (φc) and vesicles (φb), as well as bulk density (ρ). These textural and rheological changes were previously studied for Hawaiian lava, where the lava flow started as channelized pāhoehoe and transitioned into `a`ā, demonstrating a systematic trend in T, X, η, φc, φb, and ρ. NASA's FINESSE focuses on Science and Exploration through analogue research. One of the field sites is Craters of the Moon, Idaho. We present field work done at a 3.0 km long lava flow belonging to the Blue Dragon lavas erupted from a chain of spatter cones, which then coalesced into channelized flows. We acquired UAV imagery along the entire length of the flow, and generated a high resolution DTM of 5 cm/pixel, from which we derived height profiles and surface roughness values. Field work included mapping the change in surface morphology and sample collection every 150 meters. In the laboratory, we measured φc, φb, and ρ for all collected samples. Viscosity measurements were carried out by concentric cylinder viscometry at subliquidus temperatures between 1310ºC to 1160ºC to study the rheology of the lava, enabling us to relate changes in flow behavior to T and φc. Our results are consistent with observations made for Hawaiian lava, including increasing bulk density downflow, and porosity changing from connected to isolated pore space. Crystallinity increases downflow, and the transition from pāhoehoe to `a`ā occurs between 1230ºC to 1150ºC, which is prompted by nucleation and growth of plagioclase

  5. Natural-Scale Lava Flow Experiments on Video: Variations with Temperature, Slope, and Effusion Rate

    Science.gov (United States)

    Karson, J. A.; Wysocki, R.; Edwards, B. R.; Lev, E.

    2013-12-01

    Investigations of active basaltic lava flows and analog materials show that flow dynamics and final flow morphology are strongly determined by the rapidly evolving rheology of the lava crust which constrains the downslope advance of the lava flow. The non-dimensional factor Ψ (ratio of the time scale of crust formation to advective heat loss) provides a useful means of comparing different flows. The key parameters that control Ψ include the melt viscosity, temperature, effusion rate, and slope. Experimental lava flows, up to several meters long created in the Syracuse University Lava Project permit these variables to be investigated independently and in combination in volume-limited flows (Pele), that provide additional information on lava crust development. New, continuous flow (cooling-limited) experiments show downslope variations under constant flow conditions.

  6. A meta-analysis of aneurysm formation in laser assisted vascular anastomosis (LAVA)

    Science.gov (United States)

    Chen, Chen; Peng, Fei; Xu, Dahai; Cheng, Qinghua

    2009-08-01

    Laser assisted vascular anastomosis (LAVA) is looked as a particularly promising non-suture method in future. However, aneurysm formation is one of the main reasons delay the clinical application of LAVA. Some scientists investigated the incidence of aneurysms in animal model. To systematically analyze the literature on reported incidence of aneurysm formation in LAVA therapy, we performed a meta-analysis comparing LAVA with conventional suture anastomosis (CSA) in animal model. Data were systematically retrieved and selected from PUBMED. In total, 23 studies were retrieved. 18 studies were excluded, and 5 studies involving 647 animals were included. Analysis suggested no statistically significant difference between LAVA and CSA (OR 1.24, 95%CI 0.66-2.32, P=0.51). Result of meta analysis shows that the technology of LAVA is very close to clinical application.

  7. Asymmetric Meckel Cave Enlargement: A Potential Marker of PHACES Syndrome.

    Science.gov (United States)

    Wright, J N; Wycoco, V

    2017-06-01

    PHACES syndrome is a complex of morphologic abnormalities of unknown cause and includes posterior fossa abnormalities; head and neck infantile hemangiomas; arterial, cardiac, and eye anomalies; and sternal or abdominal wall defects. Accurate identification of the syndrome is important for optimal treatment. The purpose of this study was to investigate the incidence of asymmetric Meckel cave enlargement, a potential novel imaging marker, in a population of patients referred for evaluation of possible PHACES syndrome. Eighty-five patients referred for neuroimaging evaluation of possible PHACES syndrome were identified and stratified on the basis of their ultimate clinical PHACES diagnosis categorization into PHACES, possible PHACES, or not PHACES. MR imaging studies were subsequently reviewed for the presence or absence of unilateral Meckel cave enlargement, with the reviewer blinded to the ultimate PHACES syndrome categorization. Twenty-five of 85 patients (29%) were ultimately categorized as having PHACES or possible PHACES according to consensus guidelines. Asymmetric Meckel cave enlargement was present in 76% (19/25) of these patients and in 82% (19/23) of only those patients with definite PHACES. This finding was present in none of the 60 patients determined not to have PHACES syndrome. In 7/19 patients (37%) with this finding, subtle MR imaging abnormalities consistent with PHACES were missed on the initial MR imaging interpretation. Asymmetric Meckel cave enlargement was a common feature of patients with PHACES in our cohort and may serve as a novel imaging marker. Increased awareness of this imaging feature has the potential to increase the diagnostic accuracy of PHACES. © 2017 by American Journal of Neuroradiology.

  8. Post-speleogenetic biogenic modification of Gomantong Caves, Sabah, Borneo

    Science.gov (United States)

    Lundberg, Joyce; McFarlane, Donald A.

    2012-07-01

    The Gomantong cave system of eastern Sabah, Malaysia, is well-known as an important site for harvesting edible bird-nests and, more recently, as a tourist attraction. Although the biology of the Gomantong system has been repeatedly studied, very little attention has been given to the geomorphology. Here, we report on the impact of geobiological modification in the development of the modern aspect of the cave, an important but little recognized feature of tropical caves. Basic modeling of the metabolic outputs from bats and birds (CO2, H2O, heat) reveals that post-speleogenetic biogenic corrosion can erode bedrock by between ~ 3.0 mm/ka (1 m/~300 ka) and ~ 4.6 mm/ka (1 m/~200 ka). Modeling at high densities of bats yields rates of corrosion of ~ 34 mm/ka (or 1 m/~30 ka). Sub-aerial corrosion creates a previously undescribed speleological feature, the apse-flute, which is semicircular in cross-section and ~ 80 cm wide. It is vertical regardless of rock properties, developing in parallel but apparently completely independently, and often unbroken from roof to floor. They end at a blind hemi-spherical top with no extraneous water source. Half-dome ceiling conch pockets are remnants of previous apse-fluting. Sub-cutaneous corrosion creates the floor-level guano notch formed by organic acid dissolution of bedrock in contact with guano. Speleogenetic assessment suggests that as much as 70-95% of the total volume of the modern cave may have been opened by direct subaerial biogenic dissolution and biogenically-induced collapse, and by sub-cutaneous removal of limestone, over a timescale of 1-2 Ma.

  9. Reconstruction of bomb 14C time history recorded in the stalagmite from Postojna Cave

    International Nuclear Information System (INIS)

    Vokal, B.; Genty, D.; Obelic, B.

    2002-01-01

    The karstic caves provide valuable resources for reconstruction of environmental conditions on the continent in the past. This is possible due to the great stability of climatic conditions within a cave. Secondary minerals deposited in caves, known as speleothems, preserve records of long-term climatic and environmental changes at the site of their deposition and in the vicinity. The purity of speleothems and their chemical and physical stability make them exceptionally well suited for detailed geochemical and isotopic analysis

  10. Insectivorous bat reproduction and human cave visitation in Cambodia: A perfect conservation storm?

    Science.gov (United States)

    Cappelle, Julien; Hoem, Thavry

    2018-01-01

    Cave roosting bats represent an important component of Southeast Asian bat diversity and are vulnerable to human disturbance during critical reproductive periods (pregnancy, lactation and weaning). Because dramatic growth of cave tourism in recent decades has raised concerns about impacts on cave bats in the region, we assessed the reproductive phenology of two insectivorous species (Hipposideros larvatus sensu lato and Taphozous melanopogon) at three caves in Cambodia for 23 months in 2014–2016 and evaluated human visitation to these sites between 2007 and 2014. Despite the differing foraging strategies employed by the two taxa, the temporal consistency observed in proportions of pregnant, lactating and juvenile bats indicates that their major birth peaks coincide with the time of greatest cave visitation annually, particularly for domestic visitors and namely during the Cambodian new year in April. They also reflect rainfall patterns and correspond with the reproductive phenology of insectivorous cave bats in Vietnam. These findings were predictable because 1) insect biomass and thus food availability for insectivorous bats are optimal for ensuring survival of young following this period, and 2) the Khmer new year is the most significant month for religious ceremonies and thus domestic cave visitation nationally, due to the abundance of Buddhist shrines and temples in Cambodian caves. While the impact of visitor disturbance on bat population recruitment cannot be empirically assessed due to lack of historical data, it is nonetheless likely to have been considerable and raises a conservation concern. Further, because growing evidence suggests that insectivorous cave bats exhibit reproductive synchrony across continental Southeast Asia where countless cave shrines are heavily frequented during April in Theravada Buddhist countries (e.g., Myanmar, Thailand, Cambodia and Laos), our results may have wider applicability in the region. We consequently advocate for

  11. Technical devices of powered roof support for the top coal caving as automation objects

    Science.gov (United States)

    Nikitenko, M. S.; Kizilov, S. A.; Nikolaev, P. I.; Kuznetsov, I. S.

    2018-05-01

    In the paper technical devices for the top coal caving as automation objects in the composition of the longwall mining complex (LTCC) are considered. The proposed concept for automation of the top coal caving process allows caving efficiency to be ensured, coal dilution to be prevented, conveyor overloading to be prevented, the shearer service personnel to be unloaded, the influence of the “human factor” to be reduced.

  12. Late quaternary brown bear (Ursidae: Ursus cf. arctos) from a cave in the Huachuca Mountains, Arizona

    Science.gov (United States)

    Nicholas J. Czaplewski; Steve Willsey

    2013-01-01

    In 2008, Steve Willsey discovered the fragmentary cranium of a bear loose on the floor of a cave at about 2270 m elevation near the crest of the Huachuca Mountains. In 2009, we revisited the cave to examine the specimen with the intention of identifying the species. We photographed and measured the main pieces and left them in the cave. The skull is from an adult,...

  13. Insectivorous bat reproduction and human cave visitation in Cambodia: A perfect conservation storm?

    Science.gov (United States)

    Lim, Thona; Cappelle, Julien; Hoem, Thavry; Furey, Neil

    2018-01-01

    Cave roosting bats represent an important component of Southeast Asian bat diversity and are vulnerable to human disturbance during critical reproductive periods (pregnancy, lactation and weaning). Because dramatic growth of cave tourism in recent decades has raised concerns about impacts on cave bats in the region, we assessed the reproductive phenology of two insectivorous species (Hipposideros larvatus sensu lato and Taphozous melanopogon) at three caves in Cambodia for 23 months in 2014-2016 and evaluated human visitation to these sites between 2007 and 2014. Despite the differing foraging strategies employed by the two taxa, the temporal consistency observed in proportions of pregnant, lactating and juvenile bats indicates that their major birth peaks coincide with the time of greatest cave visitation annually, particularly for domestic visitors and namely during the Cambodian new year in April. They also reflect rainfall patterns and correspond with the reproductive phenology of insectivorous cave bats in Vietnam. These findings were predictable because 1) insect biomass and thus food availability for insectivorous bats are optimal for ensuring survival of young following this period, and 2) the Khmer new year is the most significant month for religious ceremonies and thus domestic cave visitation nationally, due to the abundance of Buddhist shrines and temples in Cambodian caves. While the impact of visitor disturbance on bat population recruitment cannot be empirically assessed due to lack of historical data, it is nonetheless likely to have been considerable and raises a conservation concern. Further, because growing evidence suggests that insectivorous cave bats exhibit reproductive synchrony across continental Southeast Asia where countless cave shrines are heavily frequented during April in Theravada Buddhist countries (e.g., Myanmar, Thailand, Cambodia and Laos), our results may have wider applicability in the region. We consequently advocate for

  14. Insectivorous bat reproduction and human cave visitation in Cambodia: A perfect conservation storm?

    Directory of Open Access Journals (Sweden)

    Thona Lim

    Full Text Available Cave roosting bats represent an important component of Southeast Asian bat diversity and are vulnerable to human disturbance during critical reproductive periods (pregnancy, lactation and weaning. Because dramatic growth of cave tourism in recent decades has raised concerns about impacts on cave bats in the region, we assessed the reproductive phenology of two insectivorous species (Hipposideros larvatus sensu lato and Taphozous melanopogon at three caves in Cambodia for 23 months in 2014-2016 and evaluated human visitation to these sites between 2007 and 2014. Despite the differing foraging strategies employed by the two taxa, the temporal consistency observed in proportions of pregnant, lactating and juvenile bats indicates that their major birth peaks coincide with the time of greatest cave visitation annually, particularly for domestic visitors and namely during the Cambodian new year in April. They also reflect rainfall patterns and correspond with the reproductive phenology of insectivorous cave bats in Vietnam. These findings were predictable because 1 insect biomass and thus food availability for insectivorous bats are optimal for ensuring survival of young following this period, and 2 the Khmer new year is the most significant month for religious ceremonies and thus domestic cave visitation nationally, due to the abundance of Buddhist shrines and temples in Cambodian caves. While the impact of visitor disturbance on bat population recruitment cannot be empirically assessed due to lack of historical data, it is nonetheless likely to have been considerable and raises a conservation concern. Further, because growing evidence suggests that insectivorous cave bats exhibit reproductive synchrony across continental Southeast Asia where countless cave shrines are heavily frequented during April in Theravada Buddhist countries (e.g., Myanmar, Thailand, Cambodia and Laos, our results may have wider applicability in the region. We consequently

  15. Dietary ecology of the extinct cave bear: Evidence of omnivory as inferred from dental microwear textures

    OpenAIRE

    D. Brent Jones; Larisa R.G. DeSantis

    2016-01-01

    The diet of the extinct European cave bear, Ursus spelaeus, has widely been debated. Diverging from the extant brown bear (Ursus arctos) approximately 1.2 million years ago, the cave bear is one of the most ubiquitous fossil bears occurring in Europe during the middle and Late Pleistocene. Early morphological studies suggested that the cave bear was likely specialized on processing tough and/or abrasive foods, while later two-dimensional low-magnification microwear studies suggested that they...

  16. Direct measurement of present-day tectonic movement and associated radon flux in Postojna Cave, Slovenia

    Czech Academy of Sciences Publication Activity Database

    Šebela, S.; Vaupotič, J.; Košťák, Blahoslav; Stemberk, Josef

    2010-01-01

    Roč. 72, č. 1 (2010), s. 21-34 ISSN 1090-6924 R&D Projects: GA ČR GA205/09/2024; GA MŠk MEB091005 Institutional research plan: CEZ:AV0Z30460519 Keywords : fault displacements * radon flux * Postojna Cave Subject RIV: DB - Geology ; Mineralogy Impact factor: 0.842, year: 2010 www.caves.org/pub/journal/Journal_of_Cave_and_Karst_Studies_volume_72.htm

  17. Investigation of Sediment Pathways and Concealed Sedimentological Features in Hidden River Cave, Kentucky

    Science.gov (United States)

    Feist, S.; Maclachlan, J. C.; Reinhardt, E. G.; McNeill-Jewer, C.; Eyles, C.

    2016-12-01

    Hidden River Cave is part of a cave system hydrogeologically related to Mammoth Cave in Kentucky and is a multi-level active cave system with 25km of mapped passages. Upper levels experience flow during flood events and lower levels have continuously flowing water. Improper industrial and domestic waste disposal and poor understanding of local hydrogeology lead to contamination of Hidden River Cave in the early 1940s. Previously used for hydroelectric power generation and as a source of potable water the cave was closed to the public for almost 50 years. A new sewage treatment plant and remediation efforts since 1989 have improved the cave system's health. This project focuses on sedimentological studies in the Hidden River Cave system. Water and sediment transport in the cave are being investigated using sediment cores, surface sediment samples and water level data. An Itrax core scanner is used to analyze sediment cores for elemental concentrations, magnetic susceptibility, radiography, and high resolution photography. Horizons of metal concentrations in the core allow correlation of sedimentation events in the cave system. Thecamoebian (testate amoebae) microfossils identified in surface samples allow for further constraint of sediment sources, sedimentation rates, and paleoclimatic analysis. Dive recorders monitor water levels, providing data to further understand the movement of sediment through the cave system. A general time constraint on the sediment's age is based on the presence of microplastic in the surface samples and sediment cores, and data from radiocarbon and lead-210 dating. The integration of various sedimentological data allows for better understanding of sedimentation processes and their record of paleoenvironmental change in the cave system. Sediment studies and methodologies from this project can be applied to other karst systems, and have important applications for communities living on karst landscapes and their water management policies.

  18. A cave population of Isbrueckerichthys alipionis (Gosline, 1947 in the Upper Ribeira karst area, southeastern Brazil (Siluriformes: Loricariidae

    Directory of Open Access Journals (Sweden)

    Eleonora Trajano

    Full Text Available A cave population of the armored catfish Isbrueckerichthys alipionis is reported from the Santana Cave, in the rio Betari watershed, Upper Ribeira karst area, Iporanga, São Paulo State, southeastern Brazil. The cave population was compared to an epigean population of I. alipionis and no significant differences where found in morphology or degree of pigmentation. As the cave population is known for at least 30 years and is apparently isolated from epigean streams, it is classified as troglophilic. The discovery of this troglophilic species in the Santana Cave is an additional strong argument for the conservation of that cave.

  19. The Caves of Naica: a decade of research

    International Nuclear Information System (INIS)

    Gazquez, F.; Calaforra, J. M.; Porti, P.; Badino, G.

    2016-01-01

    The caves of the Naica Mine have been the subject of study by scientists from up to seven counties over the past decade. Up to fifty research works have published to date, most relating to the origin of the giant selenite crystals of the Cueva de los Cristales. Nevertheless, a great deal of knowledge has been generated about other relevant aspects of the Naica system. This paper puts together the vast information available about the Naica caves, from the discovery of the Cueva de los Cristales in 2000 to the more recent investigations addressing mineralogy, microclimatology and the use of gypsum speleothems as a palaeo-environmental proxy. Special attention has been paid to novel research lines that have started to use the speleothems of Naica as a study case, particularly in fields such as Astrobiology and Planetary geology. Moreover, the conservation challenges which these caves will face in the near future as consequence of the end of mining activities have also been addressed in this article. (Author)

  20. Geology of the Tyrrhenus Mons Lava Flow Field, Mars

    Science.gov (United States)

    Crown, David A.; Mest, Scott C.

    2014-11-01

    The ancient, eroded Martian volcano Tyrrhenus Mons exhibits a central caldera complex, layered flank deposits dissected by radial valleys, and a 1000+ km-long flow field extending to the southwest toward Hellas Planitia. Past studies suggested an early phase of volcanism dominated by large explosive eruptions followed by subsequent effusive activity at the summit and to the southwest. As part of a new geologic mapping study of northeast Hellas, we are examining the volcanic landforms and geologic evolution of the Tyrrhenus Mons flow field, including the timing and nature of fluvial activity and effects on volcanic units. New digital geologic mapping incorporates THEMIS IR (100 m/pixel) and CTX (5 m/pixel) images as well as constraints from MOLA topography.Mapping results to-date include delineation of the boundaries of the flow field, identification and mapping of volcanic and erosional channels within the flow field, and mapping and analysis of lava flow lobes. THEMIS IR and CTX images allow improved discrimination of the numerous flow lobes that are observed in the flow field, including refinement of the margins of previously known flows and identification of additional and smaller lobes. A prominent sinuous rille extending from Tyrrhenus Mons’ summit caldera is a major feature that supplied lava to the flow field. Smaller volcanic channels are common throughout the flow field; some occur in segments along crests of local topographic highs and may delineate lava tubes. In addition to volcanic channels, the flow field surface is characterized by several types of erosional channels, including wide troughs with scour marks, elongate sinuous channels, and discontinuous chains of elongate pits and troughs. High-resolution images reveal the widespread and significant effects of fluvial activity in the region, and further mapping studies will examine spatial and temporal interactions between volcanism and fluvial processes.

  1. Cave-dwelling bats (Mammalia: Chiroptera) and conservation concerns in South central Mindanao, Philippines

    OpenAIRE

    Krizler C. Tanalgo; John Aries G. Tabora

    2015-01-01

    The stable microclimate in caves provides a relatively constant habitat for many bat species in the Philippines, but human encroachment continues to disrupt this habitat and imperil many of the species roosting in the caves.  In South central Mindanao, the diversity and conservation status of cave bats remain undocumented and unexplored.  We employed mist-netting to capture bats from five different caves within the town of Kabacan, northern Cotabato, Philippines.  A total of 14 bat species we...

  2. Occurrence of organic wastewater and other contaminants in cave streams in northeastern Oklahoma and northwestern Arkansas

    Science.gov (United States)

    Bidwell, Joseph R.; Becker, C.; Hensley, S.; Stark, R.; Meyer, M.T.

    2010-01-01

    The prevalence of organic wastewater compounds in surface waters of the United States has been reported in a number of recent studies. In karstic areas, surface contaminants might be transported to groundwater and, ultimately, cave ecosystems, where they might impact resident biota. In this study, polar organic chemical integrative samplers (POCISs) and semipermeable membrane devices (SPMDs) were deployed in six caves and two surface-water sites located within the Ozark Plateau of northeastern Oklahoma and northwestern Arkansas in order to detect potential chemical contaminants in these systems. All caves sampled were known to contain populations of the threatened Ozark cavefish (Amblyopsis rosae). The surface-water site in Oklahoma was downstream from the outfall of a municipal wastewater treatment plant and a previous study indicated a hydrologic link between this stream and one of the caves. A total of 83 chemicals were detected in the POCIS and SPMD extracts from the surface-water and cave sites. Of these, 55 chemicals were detected in the caves. Regardless of the sampler used, more compounds were detected in the Oklahoma surface-water site than in the Arkansas site or the caves. The organic wastewater chemicals with the greatest mass measured in the sampler extracts included sterols (cholesterol and ??-sitosterol), plasticizers [diethylhexylphthalate and tris (2-butoxyethyl) phosphate], the herbicide bromacil, and the fragrance indole. Sampler extracts from most of the cave sites did not contain many wastewater contaminants, although extracts from samplers in the Oklahoma surfacewater site and the cave hydrologically linked to it had similar levels of diethylhexyphthalate and common detections of carbamazapine, sulfamethoxazole, benzophenone, N-diethyl-3-methylbenzamide (DEET), and octophenol monoethoxylate. Further evaluation of this system is warranted due to potential ongoing transport of wastewaterassociated chemicals into the cave. Halogenated organics

  3. The significance of the second cave episode in Jerome’s Vita Malchi

    Directory of Open Access Journals (Sweden)

    Jacobus P. Kritzinger

    2014-02-01

    Full Text Available The authors argue that the second cave episode in Jerome’s Vita Malchi Monachi Captivi should, in view of the similarities with the first cave episode and the high incidence of literary devices employed in it, be recognised for its value in the interpretation of this vita. The book was intended as a defence of, and an exhortation to a life of celibacy and this dual purpose is clearly demonstrated in both episodes in which a cave is used as the setting. The second cave episode has been neglected in the scholarly debate about the purpose of the book and this article attempts to set the record straight.

  4. A conservation status index, as an auxiliary tool for the management of cave environments

    Directory of Open Access Journals (Sweden)

    Christiane Ramos Donato

    2014-09-01

    Full Text Available The conservation of the Speleological Heritage involves bioecological, geomorphological and anthropogenic studies, both from inside the caves and from the external environments that surround them. This study presents a method to rank caves according to their priority for conservation and restoration. Nine caves were evaluated: indicators related to the environmental impacts and the vulnerability status presented by those caves (intrinsic features and the values scored in a ‘Cave Conservation Index’ (CCI were established. We also used a rapid assessment protocol to measure cave vulnerability for prioritization of conservation/restoration actions (RAP-cr comparing natural cavities with the same lithology, due to “strictu sensu” peculiarities. Based on the protocols applied in caves of the municipality of Laranjeiras, Sergipe, Northeastern Brazil, we concluded that the present method attended to the needs for the classification of the caves into categories of conservation/restoration status, using little time and financial effort, through rapid diagnostics that facilitate the comparisons. In this perspective, the CCI can be used to indicate areas that should be protected and caves that should be prioritized to have initiated activities of conservation and restoration.

  5. NVP melt/magma viscosity: insight on Mercury lava flows

    Science.gov (United States)

    Rossi, Stefano; Morgavi, Daniele; Namur, Olivier; Vetere, Francesco; Perugini, Diego; Mancinelli, Paolo; Pauselli, Cristina

    2016-04-01

    After more than four years of orbiting Mercury, NASA's MESSENGER spacecraft came to an end in late April 2015. MESSENGER has provided many new and surprising results. This session will again highlight the latest results on Mercury based on MESSENGER observations or updated modelling. The session will further address instrument calibration and science performance both retrospective on MESSENGER and on the ESA/JAXA BepiColombo mission. Papers covering additional themes related to Mercury are also welcomed. Please be aware that this session will be held as a PICO session. This will allow an intensive exchange of expertise and experience between the individual instruments and mission. NVP melt/magma viscosity: insight on Mercury lava flows S. Rossi1, D. Morgavi1, O. Namur2, D. Perugini1, F.Vetere1, P. Mancinelli1 and C. Pauselli1 1 Dipartimento di Fisica e Geologia, Università di Perugia, piazza Università 1, 06123 Perugia, Italy 2 Uni Hannover Institut für Mineralogie, Leibniz Universität Hannover, Callinstraβe 3, 30167 Hannover, Germany In this contribution we report new measurements of viscosity of synthetic komatitic melts, used the behaviour of silicate melts erupted at the surface of Mercury. Composition of Mercurian surface magmas was calculated using the most recent maps produced from MESSENGER XRS data (Weider et al., 2015). We focused on the northern hemisphere (Northern Volcanic Province, NVP, the largest lava flow on Mercury and possibly in the Solar System) for which the spatial resolution of MESSENGER measurements is high and individual maps of Mg/Si, Ca/Si, Al/Si and S/Si were combined. The experimental starting material contains high Na2O content (≈7 wt.%) that strongly influences viscosity. High temperature viscosity measurements were carried out at 1 atm using a concentric cylinder apparatus equipped with an Anton Paar RheolabQC viscometer head at the Department of Physics and Geology (PVRG_lab) at the University of Perugia (Perugia, Italy

  6. Multifractal characterization of Vesuvio lava-flow margins and its implications

    Science.gov (United States)

    Luongo, G.; Mazzarella, A.; Di Donna, G.

    2000-09-01

    The digitized lava-flow margins of well-defined extended eruptions occurring at Vesuvio in 1760, 1794, 1861, 1906, 1929 and 1944 are found to follow fractal behaviours inside a scaling region enclosed between 50 and 400 m. Although the invariance region is well respected, the fractal dimension D varies from one lava flow to another: the more irregular the lava-flow margin, the larger the value of D. The ascertained dependence of D on the duration of premonitory activity, preceding the emission of lavas, might provide some insight into the inner volcanic processes before the eruption and into the dynamical processes operating during flow emplacement.

  7. Study of the thermoluminescent properties of lava from different origins; Estudio de las propiedades termoluminiscentes de lavas de diferentes origenes

    Energy Technology Data Exchange (ETDEWEB)

    Molina, D. [Centro de Proteccion e Higiene de las Radiaciones, A.P. 6195, C.P. 10600, La Habana (Cuba); Correcher, V.; Delgado, A. [CIEMAT. Dosimetria de Radiaciones, Av. Complutense 22, Madrid 28040 (Spain); Garcia G, J. [CSIC. Museo Nacional de Ciencias Naturales, C/Jose Gutierrez Abascal 2. Madrid 28006 (Spain)

    2002-07-01

    In this work has been studied the thermoluminescent signal (Tl) of lava from different geographical area (Costa Rica, the Canary Islands, Hawaii, Iceland and Italy) and originating in distinct eruptions, for its possible use such as in the dating field (geological and archaeological) as in retrospective dosimetry. Due that the light emission is intimately related with the punctual defects existent in the structure of material associated to the presence of different mineral phases, it was realized a study by X-ray diffraction for determining the main components of the lava observing the presence, in distinct proportions of cristobalite, plagioclases (chalcosodic feldspars) and philosilicates (augite, montmorillonite, forsterite and actinolite). All the detected mineral components present Tl emission in the blue region. Each one of the lava were artificially irradiated for proving the dependence of the luminescent signal with the dose in the range 1 to 25 Gy, observing a linear response with the dose in all the cases and not appreciating saturation in the Tl emission. Such the appropriate signal of natural samples (TLN) as the irradiated samples in the laboratory (TLI) show a complex structure associated with a continuous distribution of traps at temperature higher than 100 C which could be explained as consequence of the dynamic formation-annihilation of centers [AlO{sub 4}/alkali]{sup +} and [AlO{sub 4}]{sup 0}. In TLI was observed that a nearer to 85 C appeared a maximum whose structure correspond a discrete distribution of traps, coexisting therefore the two types of traps structure. (Author)

  8. The role of natural ventilation in the exposure to radon in the Postojna Cave

    Science.gov (United States)

    Gregorič, A.; Smerajec, M.; Vaupotič, J.

    2012-04-01

    Postojna Cave is the biggest of 21 show caves in Slovenia and one of present day's most visited show caves in the world. Long and branched out cave system, large entrances at different levels, inflow of the Pivka river, and large variation of the outdoor air temperature and precipitation, make the Postojna Cave also a very complex climatic system in which each part shows different conditions. The cave is only naturally ventilated and it is therefore characterised by high radon concentration, which depends on the ventilation regime in different seasons, resulting in typical annual cycles of radon levels in the cave air. Postojna Cave is a typical horizontal cave, where the difference between outside and cave air temperature represents the main driving force for air circulation. In winter, when the cave temperature is higher than outside, cave air is released from the cave into the outdoor atmosphere due to the air draught caused by the 'chimney effect', thus allowing fresh and cold outdoor air to enter the cave through low lying openings. This effect is not operative in summer, when the outside temperature is higher than in the cave, and air draught is minimal or reversed. In addition, air circulation can be locally altered due to other processes, like changing level of Pivka river during the rainy season and local geomorphologic characteristics of cave passages. High radon concentration in the Postojna Cave is the reason for thorough studies of the methodology for dose estimates of the personnel working in the cave. Due to high relative humidity and low air circulation, the cave air is characterised by very low particle concentration, which play an important role in radon dosimetry. Therefore parallel monitoring of radioactive aerosols of radon decay products (RnDP) and general (non-radioactive) aerosols in the particle size range of 10-1100 nm was performed in the air of Postojna Cave at the lowest point of tourist path in summer, winter and both transitional

  9. Volcanic eruptions on Io: Heat flow, resurfacing, and lava composition

    Science.gov (United States)

    Blaney, Diana L.; Johnson, Torrence V.; Matson, Dennis L.; Veeder, Glenn J.

    1995-01-01

    We model an infrared outburst on Io as being due to a large, erupting lava flow which increased its area at a rate of 1.5 x 10(exp 5)/sq m and cooled from 1225 to 555 K over the 2.583-hr period of observation. The inferred effusion rate of 3 x 10(exp 5) cu m/sec for this eruption is very high, but is not unprece- dented on the Earth and is similar to the high eruption rates suggested for early lunar volcanism. Eruptions occur approxi- mately 6% of the time on Io. These eruptions provide ample resurfacing to explain Io's lack of impact craters. We suggest that the large total radiometric heat flow, 10(exp 14) W, and the size and temperature distribution of the thermal anomalies (McEwen et al. 1992; Veeder et al. 1994) can be accounted for by a series of silicate lava flows in various stages of cooling. We propose that the whole suite of Io's currently observed thermal anomalies was produced by multiple, high-eruptive-rate silicate flows within the past century.

  10. Remagnetization of lava flows spanning the last geomagnetic reversal

    Science.gov (United States)

    Vella, Jérôme; Carlut, Julie; Valet, Jean-Pierre; Goff, Maxime Le; Soler, Vicente; Lopes, Fernando

    2017-08-01

    Large directional changes of remanent magnetization within lava flows that cooled during geomagnetic reversals have been reported in several studies. A geomagnetic scenario implies extremely rapid geomagnetic changes of several degrees per day, thus difficult to reconcile with the rate of the earth's core liquid motions. So far, no complete rock magnetic model provides a clear explanation. We revisited lava flows sandwiched between an underlying reverse and an overlying normal polarity flow marking the last reversal in three distinct volcanic sequences of the La Palma Island (Canary archipelago, Spain) that are characterized by a gradual evolution of the direction of their remanent magnetization from bottom to top. Cleaning efficiency of thermal demagnetization was not improved by very rapid heating and cooling rates as well as by continuous demagnetization using a Triaxe magnetometer. We did not observe partial self-reversals and minor changes in magnetic grain sizes are not related to the within-flow directional changes. Microscopic observations indicate poor exsolution, which suggests post-cooling thermochemical remagnetization processes. This scenario is strongly reinforced by laboratory experiments that show large resistance to thermal demagnetization when thermoremanence was acquired over a long time period. We speculate that in the present situation exsolution was reactivated during in field reheating and yielded formation of new magnetite, yet magnetic domain state rearrangements could also play a role. Initial reheating when the overlying flow took place, albeit moderate (less than 200-300 °C), was enough to produce overlying components with significantly higher unblocking temperatures.

  11. Mapping gullies, dunes, lava fields, and landslides via surface roughness

    Science.gov (United States)

    Korzeniowska, Karolina; Pfeifer, Norbert; Landtwing, Stephan

    2018-01-01

    Gully erosion is a widespread and significant process involved in soil and land degradation. Mapping gullies helps to quantify past, and anticipate future, soil losses. Digital terrain models offer promising data for automatically detecting and mapping gullies especially in vegetated areas, although methods vary widely measures of local terrain roughness are the most varied and debated among these methods. Rarely do studies test the performance of roughness metrics for mapping gullies, limiting their applicability to small training areas. To this end, we systematically explored how local terrain roughness derived from high-resolution Light Detection And Ranging (LiDAR) data can aid in the unsupervised detection of gullies over a large area. We also tested expanding this method for other landforms diagnostic of similarly abrupt land-surface changes, including lava fields, dunes, and landslides, as well as investigating the influence of different roughness thresholds, resolutions of kernels, and input data resolution, and comparing our method with previously published roughness algorithms. Our results show that total curvature is a suitable metric for recognising analysed gullies and lava fields from LiDAR data, with comparable success to that of more sophisticated roughness metrics. Tested dunes or landslides remain difficult to distinguish from the surrounding landscape, partly because they are not easily defined in terms of their topographic signature.

  12. Disclosing the temperature of columnar jointing in lavas.

    Science.gov (United States)

    Lamur, Anthony; Lavallée, Yan; Iddon, Fiona E; Hornby, Adrian J; Kendrick, Jackie E; von Aulock, Felix W; Wadsworth, Fabian B

    2018-04-12

    Columnar joints form by cracking during cooling-induced contraction of lava, allowing hydrothermal fluid circulation. A lack of direct observations of their formation has led to ambiguity about the temperature window of jointing and its impact on fluid flow. Here we develop a novel thermo-mechanical experiment to disclose the temperature of columnar jointing in lavas. Using basalts from Eyjafjallajökull volcano (Iceland) we show that contraction during cooling induces stress build-up below the solidus temperature (980 °C), resulting in localised macroscopic failure between 890 and 840 °C. This temperature window for incipient columnar jointing is supported by modelling informed by mechanical testing and thermal expansivity measurements. We demonstrate that columnar jointing takes place well within the solid state of volcanic rocks, and is followed by a nonlinear increase in system permeability of <9 orders of magnitude during cooling. Columnar jointing may promote advective cooling in magmatic-hydrothermal environments and fluid loss during geothermal drilling and thermal stimulation.

  13. Evolution of the rpoB-psbZ region in fern plastid genomes: notable structural rearrangements and highly variable intergenic spacers.

    Science.gov (United States)

    Gao, Lei; Zhou, Yuan; Wang, Zhi-Wei; Su, Ying-Juan; Wang, Ting

    2011-04-13

    The rpoB-psbZ (BZ) region of some fern plastid genomes (plastomes) has been noted to go through considerable genomic changes. Unraveling its evolutionary dynamics across all fern lineages will lead to clarify the fundamental process shaping fern plastome structure and organization. A total of 24 fern BZ sequences were investigated with taxon sampling covering all the extant fern orders. We found that: (i) a tree fern Plagiogyria japonica contained a novel gene order that can be generated from either the ancestral Angiopteris type or the derived Adiantum type via a single inversion; (ii) the trnY-trnE intergenic spacer (IGS) of the filmy fern Vandenboschia radicans was expanded 3-fold due to the tandem 27-bp repeats which showed strong sequence similarity with the anticodon domain of trnY; (iii) the trnY-trnE IGSs of two horsetail ferns Equisetum ramosissimum and E. arvense underwent an unprecedented 5-kb long expansion, more than a quarter of which was consisted of a single type of direct repeats also relevant to the trnY anticodon domain; and (iv) ycf66 has independently lost at least four times in ferns. Our results provided fresh insights into the evolutionary process of fern BZ regions. The intermediate BZ gene order was not detected, supporting that the Adiantum type was generated by two inversions occurring in pairs. The occurrence of Vandenboschia 27-bp repeats represents the first evidence of partial tRNA gene duplication in fern plastomes. Repeats potentially forming a stem-loop structure play major roles in the expansion of the trnY-trnE IGS.

  14. Evolution of the rpoB-psbZ region in fern plastid genomes: notable structural rearrangements and highly variable intergenic spacers

    Directory of Open Access Journals (Sweden)

    Su Ying-Juan

    2011-04-01

    Full Text Available Abstract Background The rpoB-psbZ (BZ region of some fern plastid genomes (plastomes has been noted to go through considerable genomic changes. Unraveling its evolutionary dynamics across all fern lineages will lead to clarify the fundamental process shaping fern plastome structure and organization. Results A total of 24 fern BZ sequences were investigated with taxon sampling covering all the extant fern orders. We found that: (i a tree fern Plagiogyria japonica contained a novel gene order that can be generated from either the ancestral Angiopteris type or the derived Adiantum type via a single inversion; (ii the trnY-trnE intergenic spacer (IGS of the filmy fern Vandenboschia radicans was expanded 3-fold due to the tandem 27-bp repeats which showed strong sequence similarity with the anticodon domain of trnY; (iii the trnY-trnE IGSs of two horsetail ferns Equisetum ramosissimum and E. arvense underwent an unprecedented 5-kb long expansion, more than a quarter of which was consisted of a single type of direct repeats also relevant to the trnY anticodon domain; and (iv ycf66 has independently lost at least four times in ferns. Conclusions Our results provided fresh insights into the evolutionary process of fern BZ regions. The intermediate BZ gene order was not detected, supporting that the Adiantum type was generated by two inversions occurring in pairs. The occurrence of Vandenboschia 27-bp repeats represents the first evidence of partial tRNA gene duplication in fern plastomes. Repeats potentially forming a stem-loop structure play major roles in the expansion of the trnY-trnE IGS.

  15. Different slopes of a mountain can determine the structure of ferns and lycophytes communities in a tropical forest of Brazil

    Directory of Open Access Journals (Sweden)

    FELIPE C. NETTESHEIM

    2014-03-01

    Full Text Available A community of Ferns and Lycophytes was investigated by comparing the occurrence of species on different slopes of a paleoisland in Southeastern Brazil. Our goal was to evaluate the hypothesis that slopes with different geographic orientations determine a differentiation of Atlantic Forest ferns and lycophytes community. We recorded these plants at slopes turned towards the continent and at slopes turned towards the open sea. Analysis consisted of a preliminary assessment on fern beta diversity, a Non Metric Multidimensional Scaling (NMDS and a Student t-test to confirm if sites sampling units ordination was different at each axis. We further used the Pearson coefficient to relate fern species to the differentiation pattern and again Student's t-test to determine if richness, plant cover and abundance varied between the two sites. There was a relatively low number of shared species between the two sites and ferns and lycophytes community variation was confirmed. Some species were detected as indicators of the community variation but we were unable to detect richness, plant cover or abundance differences. Despite the evidence of this variation between the slopes, further works are needed to evaluate which processes are contributing to determine this pattern.

  16. Owyhee River intracanyon lava flows: does the river give a dam?

    Science.gov (United States)

    Ely, Lisa L.; Brossy, Cooper C.; House, P. Kyle; Safran, Elizabeth B.; O'Connor, Jim E.; Champion, Duane E.; Fenton, Cassandra R.; Bondre, Ninad R.; Orem, Caitlin A.; Grant, Gordon E.; Henry, Christopher D.; Turrin, Brent D.

    2013-01-01

    Rivers carved into uplifted plateaus are commonly disrupted by discrete events from the surrounding landscape, such as lava flows or large mass movements. These disruptions are independent of slope, basin area, or channel discharge, and can dominate aspects of valley morphology and channel behavior for many kilometers. We document and assess the effects of one type of disruptive event, lava dams, on river valley morphology and incision rates at a variety of time scales, using examples from the Owyhee River in southeastern Oregon. Six sets of basaltic lava flows entered and dammed the river canyon during two periods in the late Cenozoic ca. 2 Ma–780 ka and 250–70 ka. The dams are strongly asymmetric, with steep, blunt escarpments facing up valley and long, low slopes down valley. None of the dams shows evidence of catastrophic failure; all blocked the river and diverted water over or around the dam crest. The net effect of the dams was therefore to inhibit rather than promote incision. Once incision resumed, most of the intracanyon flows were incised relatively rapidly and therefore did not exert a lasting impact on the river valley profile over time scales >106 yr. The net long-term incision rate from the time of the oldest documented lava dam, the Bogus Rim lava dam (≤1.7 Ma), to present was 0.18 mm/yr, but incision rates through or around individual lava dams were up to an order of magnitude greater. At least three lava dams (Bogus Rim, Saddle Butte, and West Crater) show evidence that incision initiated only after the impounded lakes filled completely with sediment and there was gravel transport across the dams. The most recent lava dam, formed by the West Crater lava flow around 70 ka, persisted for at least 25 k.y. before incision began, and the dam was largely removed within another 35 k.y. The time scale over which the lava dams inhibit incision is therefore directly affected by both the volume of lava forming the dam and the time required for sediment

  17. Environmental drivers of phototrophic biofilms in an Alpine show cave (SW-Italian Alps)

    International Nuclear Information System (INIS)

    Piano, E.; Bona, F.; Falasco, E.; La Morgia, V.; Badino, G.; Isaia, M.

    2015-01-01

    The proliferation of lampenflora is a major threat for the conservation of show caves, since phototrophic organisms cause physical, chemical and aesthetic damage to speleothems. In this paper we examine the environmental factors influencing the presence and the growth of the three main photosynthetic groups composing phototrophic biofilms in the Bossea show cave (SW-Italian Alps). The presence and the primary production of cyanobacteria, diatoms and green algae were detected with BenthoTorch®, an instrument for in situ measurement of chlorophyll a concentration that has never been used before in caves. By means of different techniques of regression analysis, we highlighted the response of the three photosynthetic groups to different environmental factors. Illuminance proved to be the main factor influencing positively both the probability of the presence and the productivity of the three groups. The presence of seeping water on the substrate and the distance from the cave entrance proved to play an important role in determining patterns of colonization. By means of GIS techniques, we provide thematic maps of the cave, providing a representation of pattern of the density of the three examined photosynthetic groups within different areas of the cave. The same approach may apply to other show caves, aiming at providing suggestions for the cave management (i.e. cleaning of the cave walls and positioning of artificial lights) and reduce impact caused by tourism. - Highlights: • We used a PAM fluorimeter on autotrophic biofilms in a show cave for the first time. • We modelled the environmental factors influencing phototrophic biofilms. • Illuminance, moisture and distance from the entrance proved to be significant. • We produced thematic maps illustrating our results. • We provide suggestions for cave management

  18. Environmental drivers of phototrophic biofilms in an Alpine show cave (SW-Italian Alps)

    Energy Technology Data Exchange (ETDEWEB)

    Piano, E., E-mail: elena.piano@unito.it [Department of Life Sciences and Systems Biology, University of Turin, Via Accademia Albertina 13, 10123 Turin (Italy); Bona, F.; Falasco, E. [Department of Life Sciences and Systems Biology, University of Turin, Via Accademia Albertina 13, 10123 Turin (Italy); La Morgia, V. [ISPRA, via Ca' Fornacetta, 9, 40064 Ozzano dell' Emilia (Italy); Badino, G. [Department of Physics, University of Turin, Via P. Giuria 1, 10125 Turin (Italy); Isaia, M. [Department of Life Sciences and Systems Biology, University of Turin, Via Accademia Albertina 13, 10123 Turin (Italy)

    2015-12-01

    The proliferation of lampenflora is a major threat for the conservation of show caves, since phototrophic organisms cause physical, chemical and aesthetic damage to speleothems. In this paper we examine the environmental factors influencing the presence and the growth of the three main photosynthetic groups composing phototrophic biofilms in the Bossea show cave (SW-Italian Alps). The presence and the primary production of cyanobacteria, diatoms and green algae were detected with BenthoTorch®, an instrument for in situ measurement of chlorophyll a concentration that has never been used before in caves. By means of different techniques of regression analysis, we highlighted the response of the three photosynthetic groups to different environmental factors. Illuminance proved to be the main factor influencing positively both the probability of the presence and the productivity of the three groups. The presence of seeping water on the substrate and the distance from the cave entrance proved to play an important role in determining patterns of colonization. By means of GIS techniques, we provide thematic maps of the cave, providing a representation of pattern of the density of the three examined photosynthetic groups within different areas of the cave. The same approach may apply to other show caves, aiming at providing suggestions for the cave management (i.e. cleaning of the cave walls and positioning of artificial lights) and reduce impact caused by tourism. - Highlights: • We used a PAM fluorimeter on autotrophic biofilms in a show cave for the first time. • We modelled the environmental factors influencing phototrophic biofilms. • Illuminance, moisture and distance from the entrance proved to be significant. • We produced thematic maps illustrating our results. • We provide suggestions for cave management.

  19. Census of biodiversity in marine caves of the eastern Mediterranean Sea

    Directory of Open Access Journals (Sweden)

    V. GEROVASILEIOU

    2015-03-01

    Full Text Available Scientific information on the biodiversity of marine caves in the eastern Mediterranean is limited, especially when considering the extensively studied caves of the north-western and central Mediterranean. Aiming to enhance current knowledge regarding cave communities, this study represents a first assessment of the marine cave biota of the eastern Mediterranean, as this has been defined by the European Union’s Marine Strategy Framework Directive (MSFD. Information retrieved from an extensive overview of relevant scientific documents was combined with original data recorded from 23 marine caves of the north-eastern Mediterranean. Our results report a total of 520 taxa recorded in eastern Mediterranean marine caves to date, the majority of which are sponges, polychaetes, rhodophytes, bivalves, fishes, and gastropods. These include several protected, endemic, and alien species. However, not all taxonomic groups have been equally studied among different areas and future studies are expected to raise the number of endemic and alien species. The overall observed trend is that the reported species number is generally related to sampling effort and scientific expertise. The most well-studied marine cave communities of the eastern Mediterranean are those of the Aegean Sea (especially its northern sector, which presented the highest number of species, followed by those of the Levantine. Furthermore, our research in Aegean caves revealed numerous new records for the marine cave fauna of the eastern basin, while several species are reported for the first time in the marine cave habitat. The critical need for further scientific research, monitoring, and conservation of this unique ecosystem was highlighted by (i the presence of certain species endemic to the eastern Mediterranean coupled with a high proportion of alien species, especially in the Levantine, and (ii the marine cave habitat availability in isolated insular areas of the eastern Mediterranean.

  20. Chemical Ecology of Cave-Dwelling Millipedes: Defensive Secretions of the Typhloiulini (Diplopoda, Julida, Julidae).

    Science.gov (United States)

    Makarov, Slobodan E; Bodner, Michaela; Reineke, Doris; Vujisić, Ljubodrag V; Todosijević, Marina M; Antić, Dragan Ž; Vagalinski, Boyan; Lučić, Luka R; Mitić, Bojan M; Mitov, Plamen; Anđelković, Boban D; Lucić, Sofija Pavković; Vajs, Vlatka; Tomić, Vladimir T; Raspotnig, Günther

    2017-04-01

    Cave animals live under highly constant ecological conditions and in permanent darkness, and many evolutionary adaptations of cave-dwellers have been triggered by their specific environment. A similar "cave effect" leading to pronounced chemical interactions under such conditions may be assumed, but the chemoecology of troglobionts is mostly unknown. We investigated the defensive chemistry of a largely cave-dwelling julid group, the controversial tribe "Typhloiulini", and we included some cave-dwelling and some endogean representatives. While chemical defense in juliform diplopods is known to be highly uniform, and mainly based on methyl- and methoxy-substituted benzoquinones, the defensive secretions of typhloiulines contained ethyl-benzoquinones and related compounds. Interestingly, ethyl-benzoquinones were found in some, but not all cave-dwelling typhloiulines, and some non-cave dwellers also contained these compounds. On the other hand, ethyl-benzoquinones were not detected in troglobiont nor in endogean typhloiuline outgroups. In order to explain the taxonomic pattern of ethyl-benzoquinone occurrence, and to unravel whether a cave-effect triggered ethyl-benzoquinone evolution, we classed the "Typhloiulini" investigated here within a phylogenetic framework of julid taxa, and traced the evolutionary history of ethyl-benzoquinones in typhloiulines in relation to cave-dwelling. The results indicated a cave-independent evolution of ethyl-substituted benzoquinones, indicating the absence of a "cave effect" on the secretions of troglobiont Typhloiulini. Ethyl-benzoquinones probably evolved early in an epi- or endogean ancestor of a clade including several, but not all Typhloiulus (basically comprising a taxonomic entity known as "Typhloiulus sensu stricto") and Serboiulus. Ethyl-benzoquinones are proposed as novel and valuable chemical characters for julid systematics.

  1. Bermuda: Search for Deep Water Caves 2009 on the R/V Endurance between 20090905 and 20090930

    Data.gov (United States)

    National Oceanic and Atmospheric Administration, Department of Commerce — Deep-water marine caves are one of the Earth's last largely unexplored frontiers of undiscovered fauna (animal life). More than 150 limestone caves are known to...

  2. Global biogeography of scaly tree ferns (Cyatheaceae): evidence for Gondwanan vicariance and limited transoceanic dispersal.

    Science.gov (United States)

    Korall, Petra; Pryer, Kathleen M

    2014-02-01

    Scaly tree ferns, Cyatheaceae, are a well-supported group of mostly tree-forming ferns found throughout the tropics, the subtropics and the south-temperate zone. Fossil evidence shows that the lineage originated in the Late Jurassic period. We reconstructed large-scale historical biogeographical patterns of Cyatheaceae and tested the hypothesis that some of the observed distribution patterns are in fact compatible, in time and space, with a vicariance scenario related to the break-up of Gondwana. Tropics, subtropics and south-temperate areas of the world. The historical biogeography of Cyatheaceae was analysed in a maximum likelihood framework using Lagrange. The 78 ingroup taxa are representative of the geographical distribution of the entire family. The phylogenies that served as a basis for the analyses were obtained by Bayesian inference analyses of mainly previously published DNA sequence data using MrBayes. Lineage divergence dates were estimated in a Bayesian Markov chain Monte Carlo framework using beast. Cyatheaceae originated in the Late Jurassic in either South America or Australasia. Following a range expansion, the ancestral distribution of the marginate-scaled clade included both these areas, whereas Sphaeropteris is reconstructed as having its origin only in Australasia. Within the marginate-scaled clade, reconstructions of early divergences are hampered by the unresolved relationships among the Alsophila , Cyathea and Gymnosphaera lineages. Nevertheless, it is clear that the occurrence of the Cyathea and Sphaeropteris lineages in South America may be related to vicariance, whereas transoceanic dispersal needs to be inferred for the range shifts seen in Alsophila and Gymnosphaera . The evolutionary history of Cyatheaceae involves both Gondwanan vicariance scenarios as well as long-distance dispersal events. The number of transoceanic dispersals reconstructed for the family is rather few when compared with other fern lineages. We suggest that a causal

  3. Population genetic variation in the tree fern Alsophila spinulosa (Cyatheaceae): effects of reproductive strategy.

    Science.gov (United States)

    Wang, Ting; Su, Yingjuan; Li, Yuan

    2012-01-01

    Essentially all ferns can perform both sexual and asexual reproduction. Their populations represent suitable study objects to test the population genetic effects of different reproductive systems. Using the diploid homosporous fern Alsophila spinulosa as an example species, the main purpose of this study was to assess the relative impact of sexual and asexual reproduction on the level and structure of population genetic variation. Inter-simple sequence repeats analysis was conducted on 140 individuals collected from seven populations (HSG, LCH, BPC, MPG, GX, LD, and ZHG) in China. Seventy-four polymorphic bands discriminated a total of 127 multilocus genotypes. Character compatibility analysis revealed that 50.0 to 70.0% of the genotypes had to be deleted in order to obtain a tree-like structure in the data set from populations HSG, LCH, MPG, BPC, GX, and LD; and there was a gradual decrease of conflict in the data set when genotypes with the highest incompatibility counts were successively deleted. In contrast, in population ZHG, only 33.3% of genotypes had to be removed to achieve complete compatibility in the data set, which showed a sharp decline in incompatibility upon the deletion of those genotypes. All populations examined possessed similar levels of genetic variation. Population ZHG was not found to be more differentiated than the other populations. Sexual recombination is the predominant source of genetic variation in most of the examined populations of A. spinulosa. However, somatic mutation contributes most to the genetic variation in population ZHG. This change of the primary mode of reproduction does not cause a significant difference in the population genetic composition. Character compatibility analysis represents an effective approach to separate the role of sexual and asexual components in shaping the genetic pattern of fern populations.

  4. Marine Biodiversity in Juan Fernández and Desventuradas Islands, Chile: Global Endemism Hotspots.

    Science.gov (United States)

    Friedlander, Alan M; Ballesteros, Enric; Caselle, Jennifer E; Gaymer, Carlos F; Palma, Alvaro T; Petit, Ignacio; Varas, Eduardo; Muñoz Wilson, Alex; Sala, Enric

    2016-01-01

    The Juan Fernández and Desventuradas islands are among the few oceanic islands belonging to Chile. They possess a unique mix of tropical, subtropical, and temperate marine species, and although close to continental South America, elements of the biota have greater affinities with the central and south Pacific owing to the Humboldt Current, which creates a strong biogeographic barrier between these islands and the continent. The Juan Fernández Archipelago has ~700 people, with the major industry being the fishery for the endemic lobster, Jasus frontalis. The Desventuradas Islands are uninhabited except for a small Chilean military garrison on San Félix Island. We compared the marine biodiversity of these islands across multiple taxonomic groups. At San Ambrosio Island (SA), in Desventuradas, the laminarian kelp (Eisenia cokeri), which is limited to Desventuradas in Chile, accounted for >50% of the benthic cover at wave exposed areas, while more sheltered sites were dominated by sea urchin barrens. The benthos at Robinson Crusoe Island (RC), in the Juan Fernández Archipelago, comprised a diverse mix of macroalgae and invertebrates, a number of which are endemic to the region. The biomass of commercially targeted fishes was >2 times higher in remote sites around RC compared to sheltered locations closest to port, and overall biomass was 35% higher around SA compared to RC, likely reflecting fishing effects around RC. The number of endemic fish species was extremely high at both islands, with 87.5% of the species surveyed at RC and 72% at SA consisting of regional endemics. Remarkably, endemics accounted for 99% of the numerical abundance of fishes surveyed at RC and 96% at SA, which is the highest assemblage-level endemism known for any individual marine ecosystem on earth. Our results highlight the uniqueness and global significance of these biodiversity hotspots exposed to very different fishing pressures.

  5. Antioxidative responses to arsenic in the arsenic-hyperaccumulator Chinese brake fern (Pteris vittata L.)

    International Nuclear Information System (INIS)

    Cao Xinde; Ma, Lena Q.; Tu Cong

    2004-01-01

    This study measured antioxidative responses of Chinese brake fern (Pteris vittata L.) upon exposure to arsenic (As) of different concentrations. Chinese brake fern was grown in an artificially-contaminated soil containing 0 to 200 mg As kg -1 (Na 2 HAsO 4 ) for 12 weeks in a greenhouse. Soil As concentrations at ≤20 mg kg -1 enhanced plant growth, with 12-71% biomass increase compared to the control. Such beneficial effects were not observed at >20 mg As kg -1 . Plant As concentrations increased with soil As concentrations, with more As being accumulated in the fronds (aboveground biomass) than in the roots and with maximum frond As concentration being 4675 mg kg -1 . Arsenic uptake by Chinese brake enhanced uptake of nutrient elements K, P, Fe, Mn, and Zn except Ca and Mg, whose concentrations mostly decreased. The contents of non-enzymatic antioxidants (glutathione, acid-soluble thiol) followed similar trends as plant As concentrations, increasing with soil As concentrations, with greater contents in the fronds than in the roots especially when exposed to high As concentrations (>50 mg kg -1 ). The activities of enzymatic antioxidants (superoxide dismutase, catalase, ascorbate peroxidase, guaiacol peroxidase) in Chinese brake followed the same trends as plant biomass, increasing with soil As up to 20 mg kg -1 and then decreased. The results indicated though both enzymatic and non-enzymatic antioxidants played significant roles in As detoxification and hyperaccumulation in Chinese brake, the former is more important at low As exposure (≤20 mg kg -1 ), whereas the latter is more critical at high As exposure (50-200 mg kg -1 ). - At low levels of arsenic exposure, enzymatic antioxidants are important for arsenic detoxification and accumulation in Chinese brake fern, while non-enzymatic antioxidants were more important at high arsenic exposure

  6. Marine Biodiversity in Juan Fernández and Desventuradas Islands, Chile: Global Endemism Hotspots

    Science.gov (United States)

    Friedlander, Alan M.; Ballesteros, Enric; Caselle, Jennifer E.; Gaymer, Carlos F.; Palma, Alvaro T.; Petit, Ignacio; Varas, Eduardo; Muñoz Wilson, Alex; Sala, Enric

    2016-01-01

    The Juan Fernández and Desventuradas islands are among the few oceanic islands belonging to Chile. They possess a unique mix of tropical, subtropical, and temperate marine species, and although close to continental South America, elements of the biota have greater affinities with the central and south Pacific owing to the Humboldt Current, which creates a strong biogeographic barrier between these islands and the continent. The Juan Fernández Archipelago has ~700 people, with the major industry being the fishery for the endemic lobster, Jasus frontalis. The Desventuradas Islands are uninhabited except for a small Chilean military garrison on San Félix Island. We compared the marine biodiversity of these islands across multiple taxonomic groups. At San Ambrosio Island (SA), in Desventuradas, the laminarian kelp (Eisenia cokeri), which is limited to Desventuradas in Chile, accounted for >50% of the benthic cover at wave exposed areas, while more sheltered sites were dominated by sea urchin barrens. The benthos at Robinson Crusoe Island (RC), in the Juan Fernández Archipelago, comprised a diverse mix of macroalgae and invertebrates, a number of which are endemic to the region. The biomass of commercially targeted fishes was >2 times higher in remote sites around RC compared to sheltered locations closest to port, and overall biomass was 35% higher around SA compared to RC, likely reflecting fishing effects around RC. The number of endemic fish species was extremely high at both islands, with 87.5% of the species surveyed at RC and 72% at SA consisting of regional endemics. Remarkably, endemics accounted for 99% of the numerical abundance of fishes surveyed at RC and 96% at SA, which is the highest assemblage-level endemism known for any individual marine ecosystem on earth. Our results highlight the uniqueness and global significance of these biodiversity hotspots exposed to very different fishing pressures. PMID:26734732

  7. Edge effects on understory epiphytic ferns and epiphyllous bryophytes in moist afromontane forests of Ethiopia

    Directory of Open Access Journals (Sweden)

    Hylander Kristoffer

    2013-12-01

    Full Text Available Most studies on edge effects in tropical forests have been conducted in landscapes with low human population density and in situations where the edges have been left unused after logging of the adjacent area. Here we studied forest margins heavily used by local farmers in a forest/agriculture mosaic landscape in Ethiopia. We compared forest structure and plant species composition across 41 forest-agriculture ecotones from 200 m out into the agricultural area to 200 m into the forest. There are strong edge effects from the edge and into the forest on canopy cover and number of stumps and apparently these forest-agricultural edges are intensively used by humans. They are penetrated by paths, beehives are found in the trees, timber of various dimensions is harvested and there is sometimes substantial cover of perennial wild (or semi-wild crops such as coffee and spices. The number of understory epiphytic fern species as well as number of epiphyllous (i.e., growing on leaves bryophyte species was lower at 20 m than at 75 m from the edge. The number of fern species was higher in newly created edges and thereafter they declined, which indicates an extinction debt. This pattern was not seen for the epiphyllous bryophytes. It is likely that different human management activities are responsible for many of the found edge effects besides wind and sun effects from the edge. Tropical forest margins provide important resources for people in many landscapes. It is important to understand how such use affects the biota of the forests. This study shows that there are substantial edge effects, but that the edge effects do not seem to become worse over time for epiphyllous bryophytes and only slightly so for ferns.

  8. Continental scale patterns and predictors of fern richness and phylogenetic diversity

    Directory of Open Access Journals (Sweden)

    Nathalie eNagalingum

    2015-04-01

    Full Text Available Because ferns have a wide range of habitat preferences and are widely distributed, they are an ideal group for understanding how diversity is distributed. Here we examine fern diversity on a broad-scale using standard and corrected richness measures as well as phylogenetic indices; in addition we determine the environmental predictors of each diversity metric. Using the combined records of Australian herbaria, a dataset of over 60,000 records was obtained for 89 genera to infer richness. A phylogenetic tree of all the genera was constructed and combined with the herbarium records to obtain phylogenetic diversity patterns. A hotspot of both taxic and phylogenetic diversity occurs in the Wet Tropics of northeastern Australia. Although considerable diversity is distributed along the eastern coast, some important regions of diversity are identified only after sample-standardization of richness and through the phylogenetic metric. Of all of the metrics, annual precipitation was identified as the most explanatory variable, in part, in agreement with global and regional fern studies. Precipitation was combined with a different variable for each different metric. For corrected richness, precipitation is combined with temperature seasonality, while correlation of phylogenetic diversity to precipitation plus radiation indicates support for the species-energy hypothesis. Significantly high and significantly low phylogenetic diversity were found in geographically separate areas. These areas are correlated with different climatic conditions such as seasonality in precipitation. The use of phylogenetic metrics identifies additional areas of significant diversity, some of which have not been revealed using traditional taxonomic analyses, suggesting that different ecological and evolutionary processes have operated over the continent. Our study demonstrates that it is possible and vital to incorporate evolutionary metrics when inferring biodiversity hotspots

  9. Population genetic variation in the tree fern Alsophila spinulosa (Cyatheaceae: effects of reproductive strategy.

    Directory of Open Access Journals (Sweden)

    Ting Wang

    Full Text Available BACKGROUND: Essentially all ferns can perform both sexual and asexual reproduction. Their populations represent suitable study objects to test the population genetic effects of different reproductive systems. Using the diploid homosporous fern Alsophila spinulosa as an example species, the main purpose of this study was to assess the relative impact of sexual and asexual reproduction on the level and structure of population genetic variation. METHODOLOGY/PRINCIPAL FINDINGS: Inter-simple sequence repeats analysis was conducted on 140 individuals collected from seven populations (HSG, LCH, BPC, MPG, GX, LD, and ZHG in China. Seventy-four polymorphic bands discriminated a total of 127 multilocus genotypes. Character compatibility analysis revealed that 50.0 to 70.0% of the genotypes had to be deleted in order to obtain a tree-like structure in the data set from populations HSG, LCH, MPG, BPC, GX, and LD; and there was a gradual decrease of conflict in the data set when genotypes with the highest incompatibility counts were successively deleted. In contrast, in population ZHG, only 33.3% of genotypes had to be removed to achieve complete compatibility in the data set, which showed a sharp decline in incompatibility upon the deletion of those genotypes. All populations examined possessed similar levels of genetic variation. Population ZHG was not found to be more differentiated than the other populations. CONCLUSIONS/SIGNIFICANCE: Sexual recombination is the predominant source of genetic variation in most of the examined populations of A. spinulosa. However, somatic mutation contributes most to the genetic variation in population ZHG. This change of the primary mode of reproduction does not cause a significant difference in the population genetic composition. Character compatibility analysis represents an effective approach to separate the role of sexual and asexual components in shaping the genetic pattern of fern populations.

  10. Feasibility Study of Padarangin Cave Slogohimo Wonogiri for Eco-Tourism

    Directory of Open Access Journals (Sweden)

    Kuswaji Dwi Priyono

    2004-01-01

    Full Text Available Area of Karst of mount Sewu in countryside of Padarangin own immeasurable of potency to be developed by besides activity of mining namely the tourism activity. Cave of nature found enable to can be developed as tourism, but date not yet been known how big potency of relevant tourism with the cave.  In line with the problems, this research aim to provide basic data of potency of cave and assess elegibility of cave. Padarangin for ecotourism in Wonogiri Regeny. Method used by survey of field and interview resident. Result of research indicate that cave of Padarangin have mount of cave at height 848 m msl which relative narrow, chamber horizontal as long as 63,8 m and vertical in 32,5 m. Needed by a special equipment to enter cave with capasities 10 – 15 people once the visit/incoming, owning value of sakral trusted by local society. The cave competent to be developed as tourism object of special enthusasm with visit limited.

  11. Low regional diversity of late cave bears mitochondrial DNA at the time of Chauvet Aurignacian paintings

    NARCIS (Netherlands)

    Bon, Celine; Berthonaud, Veronique; Fosse, Philippe; Gely, Bernard; Maksud, Frederic; Vitalis, Renaud; Philippe, Michel; van der Plicht, Johannes; Elalouf, Jean-Marc

    The Chauvet-Pont d'Arc and Deux-Ouvertures caves, located along the Ardeche River (France), contain abundant remains of the extinct cave bear (Ursus spelaeus). Because they also display a variety of Palaeolithic anthropogenic evidences, such as the earliest charcoal drawings recorded to date

  12. A world review of fungi, yeasts, and slime molds in caves

    Directory of Open Access Journals (Sweden)

    McAlpine Donald F.

    2013-01-01

    Full Text Available We provide a review of fungi, yeasts, and slime molds that have been found in natural solution caves and mines worldwide. Such habitats provide frequent roost sites for bats, and in eastern North America the environmental conditions that support white-nose syndrome, a lethal fungal disease currently devastating bat populations. A list of 1029 species of fungi, slime moulds, and yeasts in 518 genera have been documented from caves and mines worldwide in 225 articles. Ascomycota dominate the cave environment. Most research has been conducted in temperate climates, especially in Europe. A mean of 17.9±24.4SD fungal species are reported per study. Questions remain about the origin and ecological roles of fungi in caves, and which, if any, are cave-specialists. In the northern hemisphere, caves are generally characterized by relatively stable, low temperatures and a lack of organic substrates. This environment favors communities of oligotrophic, psychrotolerant fungi. Data that may help explain how cave environmental features and faunas inf luence the introduction and transmission of cave fungi remains scant.

  13. Phylogenetic diversity of fungal communities in areas accessible and not accessible to tourists in Naracoorte Caves.

    Science.gov (United States)

    Adetutu, Eric M; Thorpe, Krystal; Bourne, Steven; Cao, Xiangsheng; Shahsavari, Esmaeil; Kirby, Greg; Ball, Andrew S

    2011-01-01

    The fungal diversity in areas accessible and not accessible to tourists at UNESCO World Heritage-listed Naracoorte Caves was investigated with culture-dependent and culture-independent techniques for assistance in cave management protocol development. The caves were selected based on tourist numbers and configurations: Stick Tomato (open, high numbers), Alexandra (lockable openings, high numbers) and Strawhaven (control; no access). Culture-based survey revealed Ascomycota dominance irrespective of sampling area with Microascales (Trichurus sp.) being most frequently isolated. Some Hypocreales-like sequences belonging to Fusarium sp., Trichoderma sp. and Neonectria sp. (Stick Tomato) were cultured only from areas not accessible to tourists. These orders also were detected by DGGE assay irrespective of sampling area. The predominance of Ascomycota (especially Microascales) suggested their important ecological roles in these caves. Culture-independent analysis showed higher Shannon fungal diversity values (from ITS-based DGGE profiles) in tourist-accessible areas of these caves than in inaccessible areas with the fungal community banding patterns being substantially different in Stick Tomato Cave. Further investigations are needed to determine the cause of the differences in the fungal communities of Stick Tomato Cave, although cave-related factors such as use, configuration and sediment heterogeneity might have contributed to these differences.

  14. Identification of Martian Cave Skylights Using the Temperature Change During Day and Night

    Directory of Open Access Journals (Sweden)

    Jongil Jung

    2014-06-01

    Full Text Available Recently, cave candidates have been discovered on other planets besides the Earth, such as the Moon and Mars. When we go to other planets, caves could be possible human habitats providing natural protection from cosmic threats. In this study, seven cave candidates have been found on Pavonis Mons and Ascraeus Mons in Tharsis Montes on Mars. The cave candidates were selected using the images of the Context Camera (CTX on the Mars Reconnaissance Orbiter (MRO. The Context Camera could provide images with the high resolution of 6 meter per pixel. The diameter of the candidates ranges from 50 to 100m. Cushing et al. (2007 have analyzed the temperature change at daytime and nighttime using the Thermal Emission Imaging System (THEMIS for the sites of potential cave candidates. Similarly, we have examined the temperature change at daytime and at nighttime for seven cave candidates using the method of Cushing et al. (2007. Among those, only one candidate showed a distinct temperature change. However, we cannot verify a cave based on the temperature change only and further study is required for the improvement of this method to identify caves more clearly.

  15. Identification of Martian Cave Skylights Using the Temperature Change During Day and Night

    Science.gov (United States)

    Jung, Jongil; Yi, Yu; Kim, Eojin

    2014-06-01

    Recently, cave candidates have been discovered on other planets besides the Earth, such as the Moon and Mars. When we go to other planets, caves could be possible human habitats providing natural protection from cosmic threats. In this study, seven cave candidates have been found on Pavonis Mons and Ascraeus Mons in Tharsis Montes on Mars. The cave candidates were selected using the images of the Context Camera (CTX) on the Mars Reconnaissance Orbiter (MRO). The Context Camera could provide images with the high resolution of 6 meter per pixel. The diameter of the candidates ranges from 50 to 100m. Cushing et al. (2007) have analyzed the temperature change at daytime and nighttime using the Thermal Emission Imaging System (THEMIS) for the sites of potential cave candidates. Similarly, we have examined the temperature change at daytime and at nighttime for seven cave candidates using the method of Cushing et al. (2007). Among those, only one candidate showed a distinct temperature change. However, we cannot verify a cave based on the temperature change only and further study is required for the improvement of this method to identify caves more clearly.

  16. Aerosol particle size distribution in building and caves: impact to the radon-related dose evaluation

    International Nuclear Information System (INIS)

    Berka, Z.; Thinova, L.; Brandejsova, E.; Zdimal, V.; Fronka, A.; Milka, D.

    2004-01-01

    The results of evaluation of the aerosol particle size spectra observed in the Bozkov cave are presented and compared with the spectra observed in residential areas. The radon-to-dose conversion factor is discussed, as is the correction factor referred to as the cave factor. (P.A.)

  17. Metagenomic Analysis from the Interior of a Speleothem in Tjuv-Ante's Cave, Northern Sweden.

    Directory of Open Access Journals (Sweden)

    Marie Lisandra Zepeda Mendoza

    Full Text Available Speleothems are secondary mineral deposits normally formed by water supersaturated with calcium carbonate percolating into underground caves, and are often associated with low-nutrient and mostly non-phototrophic conditions. Tjuv-Ante's cave is a shallow-depth cave formed by the action of waves, with granite and dolerite as major components, and opal-A and calcite as part of the speleothems, making it a rare kind of cave. We generated two DNA shotgun sequencing metagenomic datasets from the interior of a speleothem from Tjuv-Ante's cave representing areas of old and relatively recent speleothem formation. We used these datasets to perform i an evaluation of the use of these speleothems as past biodiversity archives, ii functional and taxonomic profiling of the speleothem's different formation periods, and iii taxonomic comparison of the metagenomic results to previous microscopic analyses from a nearby speleothem of the same cave. Our analyses confirm the abundance of Actinobacteria and fungi as previously reported by microscopic analyses on this cave, however we also discovered a larger biodiversity. Interestingly, we identified photosynthetic genes, as well as genes related to iron and sulphur metabolism, suggesting the presence of chemoautotrophs. Furthermore, we identified taxa and functions related to biomineralization. However, we could not confidently establish the use of this type of speleothems as biological paleoarchives due to the potential leaching from the outside of the cave and the DNA damage that we propose has been caused by the fungal chemical etching.

  18. Sistema Faro, Isla de Mona, Puerto Rico: speleogenesis of the worlds largest flank margin cave

    International Nuclear Information System (INIS)

    Lace, M. J.; Kambesis, P. N.; Mylroie, J. E.

    2016-01-01

    Isla de Mona, a small, uplifted carbonate plateau jutting out of the waters of the Mona Passage, is an incredibly fragile and densely karstic environment. Expedition work was conducted by the Isla de Mona Project in cooperation with the Departamento Recursos Naturales y Ambientales de Puerto Rico (DRNA), including contributions from many researchers and cavers volunteering from across the U.S and Puerto Rico in the course of 12 separate expeditions, spanning a 14 year period (1998 to 2013). Over 200 caves have been documented on the island to date, the majority of this inventory is composed of flank margin caves but also includes sea caves, pit caves and talus caves. The most extensive example of cave development on the island is Sistema Faro - a sprawling maze-like series of chambers formed within the eastern point of the island with over 40 cliffside entrances overlooking the Caribbean Sea. Detailed cartography and analysis of the geomorphology and development of the Sistema Faro has helped form a complex model of carbonate island cave development as a function of tectonic uplift, lithology, sea level changes, karst hydrogeology and cliff retreat. This communication examines the roles these controls have played in the genesis of the world's largest flank margin cave. (Author)

  19. DNA analysis of fecal bacteria to augment an epikarst dye trace study at Crump's Cave, Kentucky

    Science.gov (United States)

    A rainfall simulation experiment was performed to investigate the transport behavior of fecal-derived bacteria through shallow karst soils and through the epikarst. The experiment was conducted at Cave Springs Cavern located just south of Mammoth Cave National Park on the Sinkhole Plain of South Cen...

  20. Geological conditions of origin of the Potočka zijalka cave

    Directory of Open Access Journals (Sweden)

    Stanko Buser

    2002-12-01

    Full Text Available In the Potočka zijalka cave on Mt. Ol{eva in Southern Karavanke during excavation of cave sediments of Würm age that contain cultural remains of the Cromagnon Man also large amounts of gravel were before the Badenian transgression, from the Central Alps during Miocene.

  1. Sediments of Biśnik Cave (Poland): Lithology and stratigraphy of the Middle Palaeolithic site

    Czech Academy of Sciences Publication Activity Database

    Krajcarz, M. T.; Bosák, Pavel; Šlechta, Stanislav; Pruner, Petr; Komar, M.; Dresler, J.; Madeyska, T.

    326/327, April (2014), s. 6-19 ISSN 1040-6182 R&D Projects: GA AV ČR IAA300130701 Institutional research plan: CEZ:AV0Z30130516 Institutional support: RVO:67985831 Keywords : karsology * caves * Biśnik Cave (Poland) Subject RIV: DB - Geology ; Mineralogy Impact factor: 2.062, year: 2014

  2. Mimogonellus dreybrodti sp. n., a new cave-inhabiting Osoriinae from Laos (Coleoptera: Staphylinidae).

    Science.gov (United States)

    Yin, Zi-Wei; Steiner, Helmut

    2017-10-17

    A new osoriine species, Mimogonellus dreybrodti Yin & Steiner, sp. n., collected from a cave in Houaphanh Province, Laos, is described and illustrated. This represents the third Mimogonellus species in Asia, and the first in the genus known to inhabit a cave environment.

  3. Phylogeography of Sardinian cave salamanders (genus Hydromantes) is mainly determined by geomorphology.

    Science.gov (United States)

    Chiari, Ylenia; van der Meijden, Arie; Mucedda, Mauro; Lourenço, João M; Hochkirch, Axel; Veith, Michael

    2012-01-01

    Detecting the factors that determine the interruption of gene flow between populations is key to understanding how speciation occurs. In this context, caves are an excellent system for studying processes of colonization, differentiation and speciation, since they represent discrete geographical units often with known geological histories. Here, we asked whether discontinuous calcareous areas and cave systems represent major barriers to gene flow within and among the five species of Sardinian cave salamanders (genus Hydromantes) and whether intraspecific genetic structure parallels geographic distance within and among caves. We generated mitochondrial cytochrome b gene sequences from 184 individuals representing 48 populations, and used a Bayesian phylogeographic approach to infer possible areas of cladogenesis for these species and reconstruct historical and current dispersal routes among distinct populations. Our results show deep genetic divergence within and among all Sardinian cave salamander species, which can mostly be attributed to the effects of mountains and discontinuities in major calcareous areas and cave systems acting as barriers to gene flow. While these salamander species can also occur outside caves, our results indicate that there is a very poor dispersal of these species between separate cave systems.

  4. Distribution of meiofaunal abundances in a marine cave complex with secondary openings and freshwater filtrations

    DEFF Research Database (Denmark)

    Riera, Rodrigo; Monterroso, Óscar; Núñez, Jorge

    2018-01-01

    Submerged sea caves are priority areas for conservation according to the Habitat Directive 92/43/CEE because of their unique biodiversity. A limited number of publications exist about communities living on sediments inside caves, mostly focused on the macrofaunal fraction (>0.5-mm body size). Mei...

  5. Hydrogeology of the Sterkfontein Cave System, Cradle of Humankind, South Africa

    CSIR Research Space (South Africa)

    Hobbs, Philip J

    2017-09-01

    Full Text Available A water level rise of almost 3 m in the space of two years in the Sterkfontein Cave system since late-2009 necessitated the re-routing of the tourist path through the cave to successively higher elevations on three occasions. It also raisedconcern...

  6. 36 CFR 290.3 - Nomination, evaluation, and designation of significant caves.

    Science.gov (United States)

    2010-07-01

    ..., DEPARTMENT OF AGRICULTURE CAVE RESOURCES MANAGEMENT § 290.3 Nomination, evaluation, and designation of... on National Forest System lands shall possess one or more of the following features, characteristics... information. (4) Hydrologic. The cave is a part of a hydrologic system or contains water which is important to...

  7. Invertebrate fossils from cave sediments: A new proxy for pre-Quaternary paleoenvironments

    Czech Academy of Sciences Publication Activity Database

    Moldovan, O.T.; Mihevc, A.; Miko, L.; Constantin, S.; Meleg, I. N.; Petculescu, A.; Bosák, Pavel

    2011-01-01

    Roč. 8, č. 7 (2011), s. 1825-1837 ISSN 1726-4170 R&D Projects: GA AV ČR IAA300130701 Institutional research plan: CEZ:AV0Z30130516 Keywords : fossil invertebrates * Oribatida * Pliocene/Pleistocene * caves * caves (Slovenia) Subject RIV: DB - Geology ; Mineralogy Impact factor: 3.859, year: 2011

  8. Effect of habitat preference on frond life span in three Cyathea tree ferns

    Science.gov (United States)

    Chiu, Tzu Yun; Wang, Hsiang Hua; Lun Kuo, Yao; Kume, Tomonori

    2013-04-01

    It has been reported that plants living in various geographical areas had different physiological forms, as factors of microenvironment have strong impacts on physiological characters. However, the physiological characters of fronds have been scarcely reported in ferns. In this study, we investigated physiological differences in response to the habitat preference in the three tree ferns in northeast Taiwan, Cyathea lepifera, C. spinulosa, and C. podophylla, prefer to open site, edge of forest, and interior forest, respectively. The canopy openness above the individuals of C. lepifera, C. spinulosa and C. podophylla were 29.2 ± 14.10 , 7.0 ± 3.07 and 5.0 ± 2.24 %, respectively. Among three species, C. podophylla had the longest frond life span (13.0 ± 4.12 months) than the two others (C. lepifera (6.8 ± 1.29 months) and C. spinulosa (7.3 ±1.35 months). Our result supported the general patterns that shade intolerant species have a shorter leaf life span than shade tolerant species. The maximum net CO2 assimilation of C. lepifera, C. spinulosa and C. podophylla were 11.46 ± 1.34, 8.27 ± 0.69, and 6.34 ± 0.54 μmol CO2 m-2 s-1, respectively. As well, C. lepifera had the highest photosynthetic light saturation point (LSP), while C. podophylla had the lowest LSP among these three tree ferns. These suggested that C. lepifera could be more efficient for capturing and utilizing light resources under the larger canopy openness condition than the other two species. We also found that frond C : N ratio were positively correlated with frond life span among species. C. podophylla, with the longest frond life span, had the highest frond C : N ratio (22.17 ± 1.95), which was followed by C. spinulosa (18.58 ± 1.37) and C. lepifera (18.68 ± 2.63) with shorter frond life span. The results were consistent to the theory that the fronds and leaves of shade intolerant species have high photosynthetic abilities with low C : N ratio. Key words: Canopy openness, frond life span

  9. CLASSICAL AND ROMANTIC FEATURES IN BEETHOVEN’S SONG CYCLE AN DIE FERNE GELIEBTE

    Directory of Open Access Journals (Sweden)

    TUREA ELENA

    2016-12-01

    Full Text Available This article examines the cycle of songs An die ferne Geliebte written by Beethoven in 1816 as the fi rst example of a vocal cycle in the history of music. It is investigated in terms of correlation between the classical and romantic features pursued both in the content and structural aspect. It features the presentiment of the romantic lied in the musical language and development principles of the thematic material, in morphogenesis. Th e semantic correlations between the cycle numbers and the melodicrhythmic ties are highlighted, special emphasis is laid on the piano part in the dramaturgy of the whole.

  10. Absorption of foliar-applied arsenic by the arsenic hyperaccumulating fern (Pteris vittata L.)

    Energy Technology Data Exchange (ETDEWEB)

    Bondada, Bhaskar R.; Tu, Shuxin; Ma, Lena Q

    2004-10-01

    The fact that heavy metals can enter various domains of the plant system through foliar pathways spurred us to explore if the fronds of the Chinese brake fern (Pteris vittata L.), a hyperaccumulator of arsenic, a carcinogenic metalloid, was proficient in absorbing arsenic in the form of sprays. The specific objective of this study was to investigate the impact of frond age, form of arsenic, and time of application on the absorption of foliar-applied arsenic by the brake fern; also examined were the effects of foliar sprays on surface ultrastructure and arsenic speciation in the frond following absorption. Foliar sprays of different arsenic concentrations (0, 50, 100, 200, and 400 ppm) were applied to young and fertile fronds. A positive linear relationship existed between arsenic concentration and absorption; the arsenic concentration of fronds increased from 50 to 200 ppm. Time-course analysis with excised pinnae indicated an initial linear increase followed by a plateau at 48 h. The young fronds with immature sori absorbed more arsenic (3100 ppm) than the fertile mature fronds (890 ppm). In the frond, the arsenic absorption was greatest in the lamina of the pinnae followed by the sori and the rachis. Applying arsenic during night (20:00-22:00 h) or afternoon (12:00-14:00 h) resulted in greater absorption of arsenic than the application in the morning (08:00-10:00 h). The arsenic absorption was greater through abaxial surfaces than through adaxial surfaces. The brake fern absorbed more arsenic when it was applied in the form of arsenite. Regardless of the form of arsenic and the surface it was applied to, arsenic occurred as arsenite, the reduced and the most toxic form of arsenic, after having been absorbed by the fronds. Scanning electron microscopy revealed no surface morphological alterations following all arsenic sprays. The study unequivocally illustrated that the Chinese brake fern absorbed foliar-applied arsenic with great efficiency. Consequently, the

  11. Absorption of foliar-applied arsenic by the arsenic hyperaccumulating fern (Pteris vittata L.)

    International Nuclear Information System (INIS)

    Bondada, Bhaskar R.; Tu, Shuxin; Ma, Lena Q.

    2004-01-01

    The fact that heavy metals can enter various domains of the plant system through foliar pathways spurred us to explore if the fronds of the Chinese brake fern (Pteris vittata L.), a hyperaccumulator of arsenic, a carcinogenic metalloid, was proficient in absorbing arsenic in the form of sprays. The specific objective of this study was to investigate the impact of frond age, form of arsenic, and time of application on the absorption of foliar-applied arsenic by the brake fern; also examined were the effects of foliar sprays on surface ultrastructure and arsenic speciation in the frond following absorption. Foliar sprays of different arsenic concentrations (0, 50, 100, 200, and 400 ppm) were applied to young and fertile fronds. A positive linear relationship existed between arsenic concentration and absorption; the arsenic concentration of fronds increased from 50 to 200 ppm. Time-course analysis with excised pinnae indicated an initial linear increase followed by a plateau at 48 h. The young fronds with immature sori absorbed more arsenic (3100 ppm) than the fertile mature fronds (890 ppm). In the frond, the arsenic absorption was greatest in the lamina of the pinnae followed by the sori and the rachis. Applying arsenic during night (20:00-22:00 h) or afternoon (12:00-14:00 h) resulted in greater absorption of arsenic than the application in the morning (08:00-10:00 h). The arsenic absorption was greater through abaxial surfaces than through adaxial surfaces. The brake fern absorbed more arsenic when it was applied in the form of arsenite. Regardless of the form of arsenic and the surface it was applied to, arsenic occurred as arsenite, the reduced and the most toxic form of arsenic, after having been absorbed by the fronds. Scanning electron microscopy revealed no surface morphological alterations following all arsenic sprays. The study unequivocally illustrated that the Chinese brake fern absorbed foliar-applied arsenic with great efficiency. Consequently, the

  12. Understory fern community structure, growth and spore production responses to a large-scale hurricane experiment in a Puerto Rico rainforest

    Science.gov (United States)

    Joanne M. Sharpe; Aaron B. Shiels

    2014-01-01

    Ferns are abundant in most rainforest understories yet their responses to hurricanes have not been well studied. Fern community structure, growth and spore production were monitored for two years before and five years after a large-scale experiment that simulated two key components of severe hurricane disturbance: canopy openness and debris deposition. The canopy was...

  13. Is there foul play in the leaf pocket? The metagenome of floating fern Azolla reveals endophytes that do not fix N2 but may denitrify.

    Science.gov (United States)

    Dijkhuizen, Laura W; Brouwer, Paul; Bolhuis, Henk; Reichart, Gert-Jan; Koppers, Nils; Huettel, Bruno; Bolger, Anthony M; Li, Fay-Wei; Cheng, Shifeng; Liu, Xin; Wong, Gane Ka-Shu; Pryer, Kathleen; Weber, Andreas; Bräutigam, Andrea; Schluepmann, Henriette

    2018-01-01

    Dinitrogen fixation by Nostoc azollae residing in specialized leaf pockets supports prolific growth of the floating fern Azolla filiculoides. To evaluate contributions by further microorganisms, the A. filiculoides microbiome and nitrogen metabolism in bacteria persistently associated with Azolla ferns were characterized. A metagenomic approach was taken complemented by detection of N 2 O released and nitrogen isotope determinations of fern biomass. Ribosomal RNA genes in sequenced DNA of natural ferns, their enriched leaf pockets and water filtrate from the surrounding ditch established that bacteria of A. filiculoides differed entirely from surrounding water and revealed species of the order Rhizobiales. Analyses of seven cultivated Azolla species confirmed persistent association with Rhizobiales. Two distinct nearly full-length Rhizobiales genomes were identified in leaf-pocket-enriched samples from ditch grown A. filiculoides. Their annotation revealed genes for denitrification but not N 2 -fixation. 15 N 2 incorporation was active in ferns with N. azollae but not in ferns without. N 2 O was not detectably released from surface-sterilized ferns with the Rhizobiales. N 2 -fixing N. azollae, we conclude, dominated the microbiome of Azolla ferns. The persistent but less abundant heterotrophic Rhizobiales bacteria possibly contributed to lowering O 2 levels in leaf pockets but did not release detectable amounts of the strong greenhouse gas N 2 O. © 2017 The Authors. New Phytologist © 2017 New Phytologist Trust.

  14. LiDAR-derived surface roughness signatures of basaltic lava types at the Muliwai a Pele Lava Channel, Mauna Ulu, Hawai`i

    Science.gov (United States)

    Whelley, Patrick L.; Garry, W. Brent; Hamilton, Christopher W.; Bleacher, Jacob E.

    2017-11-01

    We used light detection and ranging (LiDAR) data to calculate roughness patterns (homogeneity, mean-roughness, and entropy) for five lava types at two different resolutions (1.5 and 0.1 m/pixel). We found that end-member types (´áā and pāhoehoe) are separable (with 95% confidence) at both scales, indicating that roughness patterns are well suited for analyzing types of lava. Intermediate lavas were also explored, and we found that slabby-pāhoehoe is separable from the other end-members using 1.5 m/pixel data, but not in the 0.1 m/pixel analysis. This suggests that the conversion from pāhoehoe to slabby-pāhoehoe is a meter-scale process, and the finer roughness characteristics of pāhoehoe, such as ropes and toes, are not significantly affected. Furthermore, we introduce the ratio ENT/HOM (derived from lava roughness) as a proxy for assessing local lava flow rate from topographic data. High entropy and low homogeneity regions correlate with high flow rate while low entropy and high homogeneity regions correlate with low flow rate. We suggest that this relationship is not directional, rather it is apparent through roughness differences of the associated lava type emplaced at the high and low rates, respectively.

  15. Paleomagnetism of Holocene lava flows from the Reykjanes Peninsula and the Tungnaá lava sequence (Iceland): implications for flow correlation and ages

    Science.gov (United States)

    Pinton, Annamaria; Giordano, Guido; Speranza, Fabio; Þórðarson, Þorvaldur

    2018-01-01

    The impact of Holocene eruptive events from hot spots like Iceland may have had significant global implications; thus, dating and knowledge of past eruptions chronology is important. However, at high-latitude volcanic islands, the paucity of soils severely limits 14C dating, while the poor K content of basalts strongly restricts the use of K/Ar and Ar/Ar methods. Even tephrochronology, based on 14C age determinations, refers to layers that rarely lie directly above lava flows to be dated. We report on the paleomagnetic dating of 25 sites from the Reykjanes Peninsula and the Tungnaá lava sequence of Iceland. The gathered paleomagnetic directions were compared with the available reference paleosecular variation curves of the Earth magnetic field to obtain the possible emplacement age intervals. To test the method's validity, we sampled the precisely dated Laki (1783-1784 AD) and Eldgjà (934-938 AD) lavas. The age windows obtained for these events encompass the true flow ages. For sites from the Reykjanes peninsula and the Tugnaá lava sequence, we derived multiple possible eruption events and ages. In the Reykjanes peninsula, we propose an older emplacement age (immediately following the 870 AD Iceland Settlement age) for Ogmundarhraun and Kapelluhraun lava fields. For pre-historical (older than the settlement age) Tugnaá eruptions, the method has a dating precision of 300-400 years which allows an increase of the detail in the chronostratigraphy and distribution of lavas in the Tugnaá sequence.

  16. LiDAR-Derived Surface Roughness Signatures of Basaltic Lava Types at the Muliwai a Pele Lava Channel, Mauna Ulu, Hawai'i

    Science.gov (United States)

    Whelley, Patrick L.; Garry, W. Brent; Hamilton, Christopher W.; Bleacher, Jacob E.

    2017-01-01

    We used light detection and ranging (LiDAR) data to calculate roughness patterns (homogeneity, mean-roughness, and entropy) for five lava types at two different resolutions (1.5 and 0.1 m/pixel). We found that end-member types (a a and pahoehoe) are separable (with 95% confidence) at both scales, indicating that roughness patterns are well suited for analyzing types of lava. Intermediate lavas were also explored, and we found that slabby-pahoehoe is separable from the other end-members using 1.5 m/pixel data, but not in the 0.1 m/pixel analysis. This suggests that the conversion from pahoehoe to slabby-pahoehoe is a meter-scale process, and the finer roughness characteristics of pahoehoe, such as ropes and toes, are not significantly affected. Furthermore, we introduce the ratio ENT/HOM (derived from lava roughness) as a proxy for assessing local lava flow rate from topographic data. High entropy and low homogeneity regions correlate with high flow rate while low entropy and high homogeneity regions correlate with low flow rate.We suggest that this relationship is not directional, rather it is apparent through roughness differences of the associated lava type emplaced at the high and low rates, respectively.

  17. Application of Ground Penetrating Radar Supported by Mineralogical-Geochemical Methods for Mapping Unroofed Cave Sediments

    Directory of Open Access Journals (Sweden)

    Teja Čeru

    2018-04-01

    Full Text Available Ground penetrating radar (GPR using a special unshielded 50 MHz Rough Terrain Antenna (RTA in combination with a shielded 250 MHz antenna was used to study the capability of this geophysical method for detecting cave sediments. Allochthonous cave sediments found in the study area of Lanski vrh (W Slovenia are now exposed on the karst surface in the so-called “unroofed caves” due to a general lowering of the surface (denudation of carbonate rocks and can provide valuable evidence of the karst development. In the first phase, GPR profiles were measured at three test locations, where cave sediments are clearly evident on the surface and appear with flowstone. It turned out that cave sediments are clearly visible on GPR radargrams as areas of strong signal attenuation. Based on this finding, GPR profiling was used in several other places where direct indicators of unroofed caves or other indicators for speleogenesis are not present due to strong surface reshaping. The influence of various field conditions, especially water content, on GPR measurements was also analysed by comparing radargrams measured in various field conditions. Further mineralogical-geochemical analyses were conducted to better understand the factors that influence the attenuation in the area of cave sediments. Samples of cave sediments and soils on carbonate rocks (rendzina were taken for X-ray diffraction (XRD and X-ray fluorescence (XRF analyses to compare the mineral and geochemical compositions of both sediments. Results show that cave sediments contain higher amounts of clay minerals and iron/aluminium oxides/hydroxides which, in addition to the thickness of cave sediments, can play an important role in the depth of penetration. Differences in the mineral composition also lead to water retention in cave sediments even through dry periods which additionally contribute to increased attenuation with respect to surrounding soils. The GPR method has proven to be reliable for

  18. LAVA: An Open-Source Approach To Designing LAMP (Loop-Mediated Isothermal Amplification DNA Signatures

    Directory of Open Access Journals (Sweden)

    Gardner Shea N

    2011-06-01

    Full Text Available Abstract Background We developed an extendable open-source Loop-mediated isothermal AMPlification (LAMP signature design program called LAVA (LAMP Assay Versatile Analysis. LAVA was created in response to limitations of existing LAMP signature programs. Results LAVA identifies combinations of six primer regions for basic LAMP signatures, or combinations of eight primer regions for LAMP signatures with loop primers, which can be used as LAMP signatures. The identified primers are conserved among target organism sequences. Primer combinations are optimized based on lengths, melting temperatures, and spacing among primer sites. We compare LAMP signature candidates for Staphylococcus aureus created both by LAVA and by PrimerExplorer. We also include signatures from a sample run targeting all strains of Mycobacterium tuberculosis. Conclusions We have designed and demonstrated new software for identifying signature candidates appropriate for LAMP assays. The software is available for download at http://lava-dna.googlecode.com/.

  19. Historical reconstruction of climatic and elevation preferences and the evolution of cloud forest-adapted tree ferns in Mesoamerica.

    Science.gov (United States)

    Sosa, Victoria; Ornelas, Juan Francisco; Ramírez-Barahona, Santiago; Gándara, Etelvina

    2016-01-01

    Cloud forests, characterized by a persistent, frequent or seasonal low-level cloud cover and fragmented distribution, are one of the most threatened habitats, especially in the Neotropics. Tree ferns are among the most conspicuous elements in these forests, and ferns are restricted to regions in which minimum temperatures rarely drop below freezing and rainfall is high and evenly distributed around the year. Current phylogeographic data suggest that some of the cloud forest-adapted species remained in situ or expanded to the lowlands during glacial cycles and contracted allopatrically during the interglacials. Although the observed genetic signals of population size changes of cloud forest-adapted species including tree ferns correspond to predicted changes by Pleistocene climate change dynamics, the observed patterns of intraspecific lineage divergence showed temporal incongruence. Here we combined phylogenetic analyses, ancestral area reconstruction, and divergence time estimates with climatic and altitudinal data (environmental space) for phenotypic traits of tree fern species to make inferences about evolutionary processes in deep time. We used phylogenetic Bayesian inference and geographic and altitudinal distribution of tree ferns to investigate ancestral area and elevation and environmental preferences of Mesoamerican tree ferns. The phylogeny was then used to estimate divergence times and ask whether the ancestral area and elevation and environmental shifts were linked to climatic events and historical climatic preferences. Bayesian trees retrieved Cyathea, Alsophyla, Gymnosphaera and Sphaeropteris in monophyletic clades. Splits for species in these genera found in Mesoamerican cloud forests are recent, from the Neogene to the Quaternary, Australia was identified as the ancestral area for the clades of these genera, except for Gymnosphaera that was Mesoamerica. Climate tolerance was not divergent from hypothesized ancestors for the most significant

  20. The physiological resilience of fern sporophytes and gametophytes: advances in water relations offer new insights into an old lineage

    Directory of Open Access Journals (Sweden)

    Jarmila ePittermann

    2013-08-01

    Full Text Available Ferns are some of the oldest vascular plants in existence and they are the second most diverse lineage of tracheophytes next to angiosperms. Recent efforts to understand fern success have fo-cused on the physiological capacity and stress tolerance of both the sporophyte and the gameto-phyte generations. In this review, we examine these insights through the lens of plant water rela-tions, focusing primarily on the form and function of xylem tissue in the sporophyte, as well as the tolerance to and recovery from drought and desiccation stress in both stages of the fern life cycle. The absence of secondary xylem in ferns is compensated by selection for efficient primary xylem composed of large, closely arranged tracheids with permeable pit membranes. Protection from drought-induced hydraulic failure appears to arise from a combination of pit membrane traits and the arrangement of vascular bundles. Features such as tracheid-based xylem and vari-ously sized megaphylls are shared between ferns and more derived lineages, and offer an oppor-tunity to compare convergent and divergent hydraulic strategies critical to the success of xylem-bearing plants. Fern gametophytes show a high degree of desiccation tolerance but new evidence shows that morphological attributes in the gametophytes may facilitate water retention, though little work has addressed the ecological significance of this variation. We conclude with an emergent hypothesis that selection acted on the physiology of both the sporophyte and gameto-phyte generations in a synchronous manner that is consistent with selection for drought tolerance in the epiphytic niche, and the increasingly diverse habitats of the mid to late Cenozoic.

  1. Stability numerical analysis of soil cave in karst area to drawdown of underground water level

    Science.gov (United States)

    Mo, Yizheng; Xiao, Rencheng; Deng, Zongwei

    2018-05-01

    With the underground water level falling, the reliable estimates of the stability and deformation characteristics of soil caves in karst region area are required for analysis used for engineering design. Aimed at this goal, combined with practical engineering and field geotechnical test, detail analysis on vertical maximum displacement of top, vertical maximum displacement of surface, maximum principal stress and maximum shear stress were conducted by finite element software, with an emphasis on two varying factors: the size and the depth of soil cave. The calculations on the soil cave show that, its stability of soil cave is affected by both the size and depth, and only when extending a certain limit, the collapse occurred along with the falling of underground water; Additionally, its maximum shear stress is in arch toes, and its deformation curve trend of maximum displacement is similar to the maximum shear stress, which further verified that the collapse of soil cave was mainly due to shear-failure.

  2. Effect of soil-rock system on speleothems weathering in Bailong Cave, Yunnan Province, China*

    Science.gov (United States)

    Wang, Jing; Song, Lin-hua

    2005-01-01

    Bailong Cave with its well-developed Middle Triassic calcareous dolomite’s system was opened as a show cave for visitors in 1988. The speleothem scenery has been strongly weathered as white powder on the outer layers. Study of the cave winds, permeability of soil-rock system and the chemical compositions of the dripping water indicated: (1) The cave dimension structure distinctively affects the cave winds, which were stronger at narrow places. (2) Based on the different soil grain size distribution, clay was the highest in composition in the soil. The response sense of dripping water to the rainwater percolation was slow. The density of joints and other openings in dolomite make the dolomite as mesh seepage body forming piles of thin and high columns and stalactites. (3) Study of 9 dripping water samples by HYDROWIN computer program showed that the major mineral in the water was dolomite. PMID:15682505

  3. Occupational exposure to radon in Australian Tourist Caves an Australian-wide study of radon levels

    Energy Technology Data Exchange (ETDEWEB)

    Solomon, S B; Langroo, R; Peggie, J R [Australian Radiation Laboratory. Yallambie, VIC (Australia); Lyons, R G [University of Auckland, Auckland, (New Zealand). Department of Physics; James, J M [University of Sydney, Sydney, NSW (Australia). Department of Chemisty

    1996-02-01

    The study described in this report sets out to determine which Australian show caves have long- term radon levels in excess of the proposed action level of 1000 Bq m{sup -3}. The collaborative study between the Australian Radiation Laboratory (ARL), the University of Sydney and the University of Auckland, was carried out with the support of a Research Grant from Worksafe Australia. The aims of this study were to measure radon levels for each season over a period of one year, at representative sites in all developed show caves around Australia, to determine yearly average radon levels for each cave tour, based on these site measurements, to estimate the radiation doses to the tour guides employed in these caves, and to identify caves with radon concentrations in excess of the action level. (authors) 7 refs., 10 tabs., 2 figs.

  4. Occupational exposure to radon in Australian Tourist Caves an Australian-wide study of radon levels

    International Nuclear Information System (INIS)

    Solomon, S.B.; Langroo, R.; Peggie, J.R.; Lyons, R.G.; James, J.M.

    1996-02-01

    The study described in this report sets out to determine which Australian show caves have long- term radon levels in excess of the proposed action level of 1000 Bq m -3 . The collaborative study between the Australian Radiation Laboratory (ARL), the University of Sydney and the University of Auckland, was carried out with the support of a Research Grant from Worksafe Australia. The aims of this study were to measure radon levels for each season over a period of one year, at representative sites in all developed show caves around Australia, to determine yearly average radon levels for each cave tour, based on these site measurements, to estimate the radiation doses to the tour guides employed in these caves, and to identify caves with radon concentrations in excess of the action level. (authors)

  5. U-series dating of Paleolithic art in 11 caves in Spain.

    Science.gov (United States)

    Pike, A W G; Hoffmann, D L; García-Diez, M; Pettitt, P B; Alcolea, J; De Balbín, R; González-Sainz, C; de las Heras, C; Lasheras, J A; Montes, R; Zilhão, J

    2012-06-15

    Paleolithic cave art is an exceptional archive of early human symbolic behavior, but because obtaining reliable dates has been difficult, its chronology is still poorly understood after more than a century of study. We present uranium-series disequilibrium dates of calcite deposits overlying or underlying art found in 11 caves, including the United Nations Educational, Scientific, and Cultural Organization (UNESCO) World Heritage sites of Altamira, El Castillo, and Tito Bustillo, Spain. The results demonstrate that the tradition of decorating caves extends back at least to the Early Aurignacian period, with minimum ages of 40.8 thousand years for a red disk, 37.3 thousand years for a hand stencil, and 35.6 thousand years for a claviform-like symbol. These minimum ages reveal either that cave art was a part of the cultural repertoire of the first anatomically modern humans in Europe or that perhaps Neandertals also engaged in painting caves.

  6. The Astrobiology of the Subsurface: Exploring Cave Habitats on Earth, Mars and Beyond

    Science.gov (United States)

    Boston, Penelope Jane

    2016-01-01

    We are using the spectacular underground landscapes of Earth caves as models for the subsurfaces of other planets. Caves have been detected on the Moon and Mars and are strongly suspected for other bodies in the Solar System including some of the ice covered Ocean Worlds that orbit gas giant planets. The caves we explore and study include many extreme conditions of relevance to planetary astrobiology exploration including high and low temperatures, gas atmospheres poisonous to humans but where exotic microbes can flourish, highly acidic or salty fluids, heavy metals, and high background radiation levels. Some cave microorganisms eat their way through bedrock, some live in battery acid conditions, some produce unusual biominerals and rare cave formations, and many produce compounds of potential pharmaceutical and industrial significance. We study these unique lifeforms and the physical and chemical biosignatures that they leave behind. Such traces can be used to provide a "Field Guide to Unknown Organisms" for developing life detection space missions.

  7. Detecting and characterizing unroofed caves by ground penetrating radar

    Science.gov (United States)

    Čeru, Teja; Šegina, Ela; Knez, Martin; Benac, Čedomir; Gosar, Andrej

    2018-02-01

    The bare karst surface in the southeastern part of Krk Island (Croatia) is characterized by different surface karst features, such as valley-like shallow linear depressions and partially or fully sediment-filled depressions of various shapes and sizes. They were noticed due to locally increased thickness of sediment and enhanced vegetation but had not yet been systematically studied and defined. Considering only the geometry of the investigated surface features and the rare traces of cave environments detected by field surveys, it was unclear which processes (surface karstification and/or speleogenesis) contributed most to their formation. The low-frequency ground penetrating radar (GPR) method using a special 50 MHz RTA antenna was applied to study and describe these karst features. Three study sites were chosen and 5 km of GPR profiles were positioned to include various surface features. The results obtained from the GPR investigation lead to the following conclusions: (1) an increased thickness of sediment was detected in all the investigated depressions indicating their considerable depth; (2) areas between different depressions expressed as attenuated zones in GPR images reveal their interconnection; (3) transitions between surface and underground features are characterized by a collapsed passage visible in the GPR data; and (4) an underground continuation of surface valley-like depressions was detected, proving the speleogenetic origin of such features. Subsurface information obtained using GPR indicates that the valley-like depressions, irregular depressions completely or partially filled with sediment, and some dolines are associated with a nearly 4 km-long unroofed cave and developed as a result of karst denudation. In the regional context, these results suggest long-lasting karstification processes in the area, in contrast to the pre-karstic fluvial phase previously assumed to have occurred here. This research is the first application of the GPR method to

  8. Exposure to radon in the Gadime Cave, Kosovo.

    Science.gov (United States)

    Bahtijari, M; Vaupotic, J; Gregoric, A; Stegnar, P; Kobal, I

    2008-02-01

    Air radon concentration was measured in summer and winter at 11 points along the tourist guided route in the Gadime Cave in Kosovo using alpha scintillation cells and etched track detectors. At two points in summer, values higher than 1700Bqm(-3) were observed; they otherwise were in the range 400-1000Bqm(-3). Values were lower in winter. The effective dose received by a person during a 90min visit is 3.7microSv in summer and 2.5microSv in winter. For a tourist guide the annual effective dose is less than 3.5mSv.

  9. Exposure to radon in the Gadime Cave, Kosovo

    International Nuclear Information System (INIS)

    Bahtijari, M.; Vaupotic, J.; Gregoric, A.; Stegnar, P.; Kobal, I.

    2008-01-01

    Air radon concentration was measured in summer and winter at 11 points along the tourist guided route in the Gadime Cave in Kosovo using alpha scintillation cells and etched track detectors. At two points in summer, values higher than 1700 Bq m -3 were observed; they otherwise were in the range 400-1000 Bq m -3 . Values were lower in winter. The effective dose received by a person during a 90 min visit is 3.7 μSv in summer and 2.5 μSv in winter. For a tourist guide the annual effective dose is less than 3.5 mSv

  10. Exposure to radon in the Gadime Cave, Kosovo

    Energy Technology Data Exchange (ETDEWEB)

    Bahtijari, M. [Faculty of Education, University of Prishtina, Prishtina, Kosovo (Country Unknown); Vaupotic, J.; Gregoric, A.; Stegnar, P. [Jozef Stefan Institute, PO Box 3000, 1001 Ljubljana (Slovenia); Kobal, I. [Jozef Stefan Institute, PO Box 3000, 1001 Ljubljana (Slovenia)], E-mail: ivan.kobal@ijs.si

    2008-02-15

    Air radon concentration was measured in summer and winter at 11 points along the tourist guided route in the Gadime Cave in Kosovo using alpha scintillation cells and etched track detectors. At two points in summer, values higher than 1700 Bq m{sup -3} were observed; they otherwise were in the range 400-1000 Bq m{sup -3}. Values were lower in winter. The effective dose received by a person during a 90 min visit is 3.7 {mu}Sv in summer and 2.5 {mu}Sv in winter. For a tourist guide the annual effective dose is less than 3.5 mSv.

  11. 118-C-4 Horizontal Rod Cave Characterization Report

    International Nuclear Information System (INIS)

    Encke, B.D.; Thoren, R.A.

    1998-03-01

    This report addresses the characterization data collected from 118-C- 4 Horizontal Rod Cave in December 1996 and August 1997. The characterization activities evaluated the radiological status and identified hazardous materials locations. The scope of this report is limited to the 118-C-4 Facility Structure. Information in this report can be used to identify the waste streams, provide specific chemical and radiological data to aid in planning decontamination and decommissioning (D ampersand D) activities, and allow proper disposal of the demolition debris, as required by the Comprehensive Environmental Response, Compensation, and Liability Act of 1980

  12. Designing user models in a virtual cave environment

    Energy Technology Data Exchange (ETDEWEB)

    Brown-VanHoozer, S. [Argonne National Lab., Idaho Falls, ID (United States); Hudson, R. [Argonne National Lab., IL (United States); Gokhale, N. [Madge Networks, San Jose, CA (United States)

    1995-12-31

    In this paper, the results of a first study into the use of virtual reality for human factor studies and design of simple and complex models of control systems, components, and processes are described. The objective was to design a model in a virtual environment that would reflect more characteristics of the user`s mental model of a system and fewer of the designer`s. The technology of a CAVE{trademark} virtual environment and the methodology of Neuro Linguistic Programming were employed in this study.

  13. Geological constraints on cave development in the plateau-gorge karst of South China (Wulong, Chongqing)

    Science.gov (United States)

    Szczygieł, Jacek; Golicz, Mateusz; Hercman, Helena; Lynch, Erin

    2018-03-01

    The Houping Tiankeng cluster is a part of the South China Karst UNESCO World Natural Heritage Site. Within the distinctive Wulong plateau-gorge karst, > 200 km of cave passages have been documented to date. This paper focuses on detailed tectonic and morphological research on the Luo Shui Kong cave, enriched with U-series dating of speleothems and complemented by morphometric analysis of the San Wang Dong and Er Wang Dong caves. All of these caves exhibit three regional levels of cave development: 1) 1040-1020 m a.s.l.; 2) 900-840 m a.s.l.; and 3) 740-660 m a.s.l. The Houping Tiankeng area is a carbonate rock sequence several hundred meters thick, overlain by the less soluble Lower Ordovician strata, limiting recharge points to faults exposing underlying easily soluble formations. This leads to the domination of concentrated, high-volume inflow and thus results in caves of large volume in the plateau-gorge karst. Shafts connecting the surface with cave passages located underneath formed along faults, changing the hydrogeological pattern through karst water capture and remodeling of existing conduits, albeit mainly by increasing their overall dimensions rather than by deepening them. The most favorable structures for cave-level development are two sets of joints conjugated with gently inclined bedding. Since these joints are characterized by a small vertical extent, downward development is limited. Hence, most of the passages are wide but not deep canyons and typical of a water-table cave pattern. Places where the fault plane is eroded from the surface and where, at the same time, an underneath cave chamber ceiling expands upwards are particularly predisposed to the formation of a tiankeng.

  14. Microbial communities in dark oligotrophic volcanic ice cave ecosystems of Mt. Erebus, Antarctica

    Directory of Open Access Journals (Sweden)

    Bradley M. Tebo

    2015-03-01

    Full Text Available The Earth’s crust hosts a subsurface, dark, and oligotrophic biosphere that is poorly understood in terms of the energy supporting its biomass production and impact on food webs at the Earth’s surface. Dark oligotrophic volcanic ecosystems (DOVEs are good environments for investigations of life in the absence of sunlight as they are poor in organics, rich in chemical reactants and well known for chemical exchange with Earth’s surface systems. Ice caves near the summit of Mt. Erebus (Antarctica offer DOVEs in a polar alpine environment that is starved in organics and with oxygenated hydrothermal circulation in highly reducing host rock. We surveyed the microbial communities using PCR, cloning, sequencing and analysis of the small subunit (16S ribosomal and Ribulose-1,5-bisphosphate Carboxylase/Oxygenase (RubisCO genes in sediment samples from three different caves, two that are completely dark and one that receives snow-filtered sunlight seasonally. The microbial communities in all three caves are composed primarily of Bacteria and fungi; Archaea were not detected. The bacterial communities from these ice caves display low phylogenetic diversity, but with a remarkable diversity of RubisCO genes including new deeply branching Form I clades, implicating the Calvin-Benson-Bassham cycle as a pathway of CO2 fixation. The microbial communities in one of the dark caves, Warren Cave, which has a remarkably low phylogenetic diversity, were analyzed in more detail to gain a possible perspective on the energetic basis of the microbial ecosystem in the cave. Atmospheric carbon (CO2 and CO, including from volcanic emissions, likely supplies carbon and/or some of the energy requirements of chemoautotrophic microbial communities in Warren Cave and probably other Mt. Erebus ice caves. Our work casts a first glimpse at Mt. Erebus ice caves as natural laboratories for exploring carbon, energy and nutrient sources in the subsurface biosphere and the

  15. LAVA: A conceptual framework for automated risk assessment

    International Nuclear Information System (INIS)

    Smith, S.T.; Brown, D.C.; Erkkila, T.H.; FitzGerald, P.D.; Lim, J.J.; Massagli, L.; Phillips, J.R.; Tisinger, R.M.

    1986-01-01

    At the Los Alamos National Laboratory the authors are developing the framework for generating knowledge-based systems that perform automated risk analyses on an organizations's assets. An organization's assets can be subdivided into tangible and intangible assets. Tangible assets include facilities, material, personnel, and time, while intangible assets include such factors as reputation, employee morale, and technical knowledge. The potential loss exposure of an asset is dependent upon the threats (both static and dynamic), the vulnerabilities in the mechanisms protecting the assets from the threats, and the consequences of the threats successfully exploiting the protective systems vulnerabilities. The methodology is based upon decision analysis, fuzzy set theory, natural language processing, and event tree structures. The Los Alamos Vulnerability and Risk Assessment (LAVA) methodology has been applied to computer security. The program generates both summary reports for use by both management personnel and detailed reports for use by operations staff

  16. Observation of Possible Lava Tube Skylights by SELENE cameras

    Science.gov (United States)

    Haruyama, Junichi; Hiesinger, Harald; van der Bogert, Carolyn

    We have discovered three deep hole-structures on the Moon in the Terrain Camera and Multi-band Imager on the SELENE. These holes are large depth to diameter ratios: Marius Hills Hole (MHH) is 65 m in diameter and 88-90 m in depth, Mare Tranquillitatis Hole (MTH) is 120 x 110 m in diameter and 180 m in depth, and Mare Ingenii Hole (MIH) is 140 x 110 m in diameter and deeper than 90 m. No volcanic material from the holes nor dike-relating pit craters is seen around the holes. They are possible lava tube skylights. These holes and possibly connected tubes have a lot of scientific interests and high potentialities as lunar bases.

  17. Intraflow width variations in Martian and terrestrial lava flows

    Science.gov (United States)

    Peitersen, Matthew N.; Crown, David A.

    1997-03-01

    Flow morphology is used to interpret emplacement processes for lava flows on Earth and Mars. Accurate measurements of flow geometry are essential, particularly for planetary flows where neither compositional sampling nor direct observations of active flows may be possible. Width behavior may indicate a flow's response to topography, its emplacement regime, and its physical properties. Variations in width with downflow distance from the vent may therefore provide critical clues to flow emplacement processes. Flow width is also one of the few characteristics that can be readily measured from planetary mission data with accuracy. Recent analyses of individual flows at two terrestrial and four Martian sites show that widths within an individual flow vary by up to an order of magnitude. Width is generally thought to be correlated to topography; however, recent studies show that this relationship is neither straightforward nor easily quantifiable.

  18. Magnetic property zonation in a thick lava flow

    Science.gov (United States)

    Audunsson, Haraldur; Levi, Shaul; Hodges, Floyd

    1992-04-01

    Intraflow structures and magmatic evolution in an extensive and thick (30-60 m) basaltic lava flow are examined on the basis of grain size and composition-dependent magnetic properties of titanomagnetite materials. Microprobe data indicate that the intraflow oxidation state Fe(3+)/Fe(2+) of the initially precipitated primary titanomagnetites increases with falling equilibrium temperature from the flow margins to a maximum near the center, the position of lowest equilibrium temperature. In contrast, Curie temperature measurements indicate that titanomagnetite oxidation increases with height in the flow. Modification of the initially symmetric equilibrium titanomagnetite compositions was caused by subsolidus high-temperature oxidation possibly due to hydrogen loss produced by dissociation of magmatic water, as well as unknown contributions of circulating air and percolating water from above. The titanomagnetites of the basal layer of the flow remain essentially unaltered.

  19. 230Th-238U disequilibria in historical lavas from Iceland

    International Nuclear Information System (INIS)

    Condomines, M.; Morand, P.; Alleegre, C.J.; Sigvaldason, G.

    1981-01-01

    The 230 Th- 238 U disequilibrium studies on historical lavas from Iceland show a relative homogeneity for Th/U ratios and also a variation for ( 230 Th/ 232 Th) activity ratios at the scale of the island. The ( 230 Th/ 238 U) disequilibrium ratio is always greater than 1 which indicates that partial melting produces magmas with Th/U ratios greater than those of the mantle source. Furthermore, there seems to be a correlation between the variations of ( 230 Th/ 232 Th) (and delta 18 O) ratios and the geographical location of the samples along the active zones of Iceland. We develop and discuss several models in order to explain these variations. (orig.)

  20. Dynamics of a fluid flow on Mars: Lava or mud?

    Science.gov (United States)

    Wilson, Lionel; Mouginis-Mark, Peter J.

    2014-05-01

    A distinctive flow deposit southwest of Cerberus Fossae on Mars is analyzed. The flow source is a ∼20 m deep, ∼12 × 1.5 km wide depression within a yardang associated with the Medusae Fossae Formation. The flow traveled for ∼40 km following topographic lows to leave a deposit on average 3-4 km wide. The surface morphology of the deposit suggests that it was produced by the emplacement of a fluid flowing in a laminar fashion and possessing a finite yield strength. We use topographic data from a digital elevation model (DEM) to model the dynamics of the motion and infer that the fluid had a Bingham rheology with a plastic viscosity of ∼1 Pa s and a yield strength of ∼185 Pa. Although the low viscosity is consistent with the properties of komatiite-like lava, the combination of values of viscosity and yield strength, as well as the surface morphology of the flow, suggests that this was a mud flow. Comparison with published experimental data implies a solids content close to 60% by volume and a grain size dominated by silt-size particles. Comparison of the ∼1.5 km3 deposit volume with the ∼0.03 km3 volume of the source depression implies that ∼98% of the flow material was derived from depth in the crust. There are similarities between the deposit studied here, which we infer to be mud, and other flow deposits on Mars currently widely held to be lavas. This suggests that a re-appraisal of many of these deposits is now in order.