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Sample records for eastern spruce structural

  1. CLIMATE CHANGE AND ORIENTAL SPRUCE (PICEA ORIENTALIS ECOSYSTEMS IN EASTERN BLACKSEA REGION OF TURKEY

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    Aydın Tüfekçioğlu

    2008-04-01

    Full Text Available Climate change has been getting more attention from scientific community recently. Eastern Black Sea Region of Turkey will get significant influences from the climate change according to regional climate model (RegCM3. Oriental spruce (Picea orientalis L. is an important tree species of Turkey and it only grows in the Eastern Black Sea Region of Turkey. With the increase in global warming, spruce forests started to have serious bark beetle problems. More than 200 000 trees died in the region recently due to bark beetle attack. We used existing literature related to oriental spruce and future climate of the region and field observations done in the different times to assess current status of the spruce stands. Future climate of the region has been predicted using RegCM3 regional climate model. Climate change could significantly influence distribution, diversity, structure and stability of the oriental spruce ecosystems. According to RegCM3 regional climate model, the temperatures will increase 2-4 °C in the region in the next century. Future climate scenarios predict 200-300 mm increases in precipitation in the eastern part of the region while the western part won't have any increase in precipitation in the next century. Temperature increases in the western part of the region can cause more stress on spruce trees and would probably increase bark beetle attacks. Also, fire could become an important threat in the western part of the region. It is possible to observe 400-800 m upward shift in the spruce belt in the western part. Treeline of spruce stands would probably move upward both in western and eastern part of the North-eastern Blacksea Region.

  2. Animal vectors of eastern dwarf mistletoe of black spruce.

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    Michael E. Ostry; Thomas H. Nicholls; D.W. French

    1983-01-01

    Describes a study to determine the importance of animals in the spread of eastern dwarf mistletoe of black spruce. Radio telemetry, banding, and color-marking techniques were used to study vectors of this forest pathogen.

  3. Effects of a western spruce budworm outbreak on private lands in eastern Oregon, 1980-1994.

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    David L. Azuma; David L. Overhulser

    2008-01-01

    Forest Inventory and Analysis data from three inventory periods were used to examine the effects of a western spruce budworm outbreak on private lands in eastern Oregon. Growth was negatively related to defoliation with differences between crown ratio and species. The mortality and salvage harvesting caused changes in stand structure on private lands. Although many...

  4. Phytosociological studies of the forests with sessile oak and Norway spruce from South-Eastern Transylvania

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    Adrian Indreica

    2011-02-01

    Full Text Available The forests with sessile oak (Quercus petraea and Norway spruce (Picea abies from south-eastern Transylvania represent a peculiar type of phytocenoses, rather unusual for the present-day vegetation of Romania’s territory. Aim of the study is to provide a detailed description of the vegetation and to identify the phytosociological and typological units to which it could belong. Beside this, stand structure and regeneration status of the main tree species are illustrated. The studied area is located around Carpathian intermountain depressions Braşov and Ciuc, where vegetation had a peculiar history and today sessile oak forests on high altitude exists, interfering with spruce forests. The hypothesis of the process naturalness is supported by vegetation history in the area, climate, stand structure and peculiarities of herb layer composition (the mixture of relic of both mountain-boreal origin and south-European origin, like Vaccinium vitis-idaea, Pyrola rotundifolia and respectively Potentilla micrantha, Lathyrus venetus respectively. Sintaxonomically, studied phytocenoses with sessile oak and spruce belong mainly to acidophilus oak forests (Luzulo luzuloidis-Quercetum petraeae, but some of them resemble oak-hornbeam forests (Carici pilosae-Carpinetum, indicating a more recent change in stand structure and suggesting that not the soil, but the climate is the driving force of succession. Regeneration of sessile oak is at least satisfactory, but the expansion of spruce in such stands could seriously restrict the survival of sessile oak. A new typological unit will be appropriate,for a better management of sessile oak forests with spruce admixture.

  5. Phytosociological studies of the forests with sessile oak and Norway spruce from South-Eastern Transylvania

    Directory of Open Access Journals (Sweden)

    Adrian Indreica

    2011-06-01

    Full Text Available The forests with sessile oak (Quercus petraea and Norway spruce (Picea abies from south-eastern Transylvania represent a peculiar type of phytocenoses, rather unusual for the present-day vegetation of Romania’s territory. Aim of the study is to provide a detailed description of the vegetation and to identify the phytosociological and typological units to which it could belong. Beside this, stand structure and regeneration status of the main tree species are illustrated. The studied area is located around Carpathian intermountain depressions Brasov and Ciuc, where vegetation had a peculiar history and today sessile oak forests on high altitude exists, interfering with spruce forests. The hypothesis of the process naturalness is supported by vegetation history in the area, climate, stand structure and peculiarities of herb layer composition (the mixture of relic of both mountain-boreal origin and south-European origin, like Vaccinium vitis-idaea, Pyrola rotundifolia and respectively Potentilla micrantha, Lathyrus venetus respectively. Sintaxonomically, studied phytocenoses with sessile oak and spruce belong mainly to acidophilus oak forests (Luzulo luzuloidis-Quercetum petraeae, but some of them resemble oak-hornbeam forests (Carici pilosae-Carpinetum, indicating a more recent change in stand structure and suggesting that not the soil, but the climate is the driving force of succession. Regeneration of sessile oak is at least satisfactory, but the expansion of spruce in such stands could seriously restrict the survival of sessile oak. A new typological unit will be appropriate, for a better management of sessile oak forests with spruce admixture.

  6. Ecology and silviculture of the spruce-fir forests of eastern North America

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    Marinus. Westveld

    1953-01-01

    Using the climax forest as a guide to growing the species best suited to the climate and the site, the author offers a silvicultural system for managing the spruce-fir forests of eastern North America. Based on ecological principles, such silviculture is aimed to bring about forests that are inherently healthy and have a natural resistance to insects and disease.

  7. Chemical Basis of Host Plant Selection by Eastern Spruce Budworm, Choristoneura fumiferana Clem. (Lepidoptera:Tortricidae)

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    Robert L. Talerico; Michael Montgomery; [Tech. Coords

    1983-01-01

    The epicuticular waxes from four host plants of the eastern spruce budworm are examined with respect to their influence on the feeding behavior of the sixth-instar larva. Both current and one-year old needles contain stimulating chemicals in their epicuticular wax layer. Some pure fatty acids known to occur in balsam fir wax are stimulatory, and may serve to enhance...

  8. Availability of ectomycorrhizal fungi to black spruce above the present treeline in Eastern Labrador.

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    Laura Reithmeier

    Full Text Available Ectomycorrhizal fungi (ECMF are an important biotic factor in the survival of conifer seedlings under stressful conditions and therefore have the potential to facilitate conifer establishment into alpine and tundra habitats. In order to assess patterns of ectomycorrhizal availability and community structure above treeline, we conducted soil bioassays in which Picea mariana (black spruce seedlings were grown in field-collected soils under controlled conditions. Soils were collected from distinct alpine habitats, each dominated by a different ectomycorrhizal host shrub: Betula glandulosa, Arctostaphylos alpina or Salix herbacaea. Within each habitat, half of the soils collected contained roots of ectomycorrhizal shrubs (host (+ and the other half were free of host plants (host(-. Forest and glacial moraine soils were also included for comparison. Fungi forming ectomycorrhizae during the bioassays were identified by DNA sequencing. Our results indicate that ECMF capable of colonizing black spruce are widespread above the current tree line in Eastern Labrador and that the level of available inoculum has a significant influence on the growth of seedlings under controlled conditions. Many of the host(- soils possessed appreciable levels of ectomycorrhizal inoculum, likely in the form of spore banks. Inoculum levels in these soils may be influenced by spore production from neighboring soils where ectomycorrhizal shrubs are present. Under predicted temperature increases, ectomycorrhizal inoculum in soils with host shrubs as well as in nearby soils without host shrubs have the potential to facilitate conifer establishment above the present tree line.

  9. Availability of ectomycorrhizal fungi to black spruce above the present treeline in Eastern Labrador.

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    Reithmeier, Laura; Kernaghan, Gavin

    2013-01-01

    Ectomycorrhizal fungi (ECMF) are an important biotic factor in the survival of conifer seedlings under stressful conditions and therefore have the potential to facilitate conifer establishment into alpine and tundra habitats. In order to assess patterns of ectomycorrhizal availability and community structure above treeline, we conducted soil bioassays in which Picea mariana (black spruce) seedlings were grown in field-collected soils under controlled conditions. Soils were collected from distinct alpine habitats, each dominated by a different ectomycorrhizal host shrub: Betula glandulosa, Arctostaphylos alpina or Salix herbacaea. Within each habitat, half of the soils collected contained roots of ectomycorrhizal shrubs (host (+)) and the other half were free of host plants (host(-)). Forest and glacial moraine soils were also included for comparison. Fungi forming ectomycorrhizae during the bioassays were identified by DNA sequencing. Our results indicate that ECMF capable of colonizing black spruce are widespread above the current tree line in Eastern Labrador and that the level of available inoculum has a significant influence on the growth of seedlings under controlled conditions. Many of the host(-) soils possessed appreciable levels of ectomycorrhizal inoculum, likely in the form of spore banks. Inoculum levels in these soils may be influenced by spore production from neighboring soils where ectomycorrhizal shrubs are present. Under predicted temperature increases, ectomycorrhizal inoculum in soils with host shrubs as well as in nearby soils without host shrubs have the potential to facilitate conifer establishment above the present tree line.

  10. Changes of Soil Enzyme Activities in Different Restoration Ages of Spruce Forests on the Eastern Qinghai-Tibet Plateau

    Institute of Scientific and Technical Information of China (English)

    ZHANG Yong-mei; BAO Wei-kai; PANG Xue-yong; WU Ning; ZHOU Guo-yi

    2005-01-01

    Six soil enzymes (invertase, acid phosphatase, proteinase, catalase, peroxidase and polyphenoloxidase ) were chosen for investigation under different spruce forests with restoration ages of 4,10,16 years and an old-growth spruce forest over 400 years old in the eastern Qinghai-Tibet Plateau, China. Results showed that the activities of invertase, phosphatase, proteinase, catalase and peroxidase decreased in newly restored forests except for pholyphenoloxidase. With the development of forests after restoration, the activities of invertase, acid phosphadase, proteinase increased gradually. Our study also indicated that the soil enzyme activities were associated with surface soils and decreased with depths. This result suggested that in the earlier restoration stage the application of organic fertilizer may be more effective by surface addition to soils than deep addition.

  11. Structure and productivity of mixed spruce and fir forests on Mt. Kopaonik

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    Šljukić Biljana

    2017-01-01

    Full Text Available The subject of this research are mixed forests of spruce and fir in the area of NP Kopaonik, which belong to the community of spruce and fir - Abieti-Piceetum abietis Mišić et Popović, 1978. The basis for the study of the structural development and production potential of these forests are data from 12 sample plots, with the average size of 0.18 ha. In terms of coenoecological affiliation all the sample plots belong to the group of ecological units - forests of spruce and fir (Abieti-Piceetum abietis, Mišić et Popović, 1978 on acid brown and brown podzolic soils, which are differentiated into 5 ecological units: Abieti-Piceetum abietis oxalidetosum on brown podzolic soil, Abieti-Piceetum abietis oxalidetosum on acid brown soil, Abieti-Piceetum abietis vaccinietosum on brown podzolic soil, Abieti-Piceetum abietis typicum on brown podzolic soil and Abieti-Piceetum abietis Dr.ymetosum on brown pozolic soil. In structural terms, these forests are characterized by very diverse structural forms, ranging from the structure of even-aged stands to typical multi-storey, unevenaged-aged stands. The form of cumulative curves of tree distribution is in most cases determined by spruce as the dominant species. At the same time, thin and medium-thick trees dominate, while the presence of stems with large dimensions is minimal. The average volume of these forestse is 777 m3•ha-1, with a mixture ratio of 0.7: 0.3 in favor of spruce. The average value of the current volume increment is 14 m3•ha-1, with a 68% share of spruce and 32% of fir. The percentage of increment ranges from 1.6% to 2.5% in all sample plots and is somewhat higher for fir. The site potential, stand characteristics and relations among the tree species have resulted in structural complexity, high productivity and ecological stability of these forests. Therefore, future forest management should avoid radical measures and procedures that would violate the established relationships and

  12. The effects of a western spruce budworm outbreak on the dead wood component in relation to ownership in forests of eastern Oregon

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    David. Azuma

    2010-01-01

    Forest Inventory and Analysis data were used to investigate the effects of a severe western spruce budworm outbreak on the dead wood component of forests in 11 counties of eastern Oregon for two time periods. The ownership and the level of damage (as assessed by aerial surveys) affected the resulting down woody material and standing dead trees. The pattern of coarse...

  13. Acidification of a white spruce ecosystem in eastern Cape Breton Island

    International Nuclear Information System (INIS)

    Bouman, O.T.

    2005-01-01

    A study was conducted in 2003 at an ecosystem monitoring plot near Sydney, Nova Scotia, in a mature white spruce stand on a Shulie soil. The objective was to examine how spruce forests filter atmospheric sulfur dioxide and become destabilized by the resulting soil acidification. The acid rain problem at the level of input, top soil, sub soil and run off was assessed following results from 4 monitoring stations equipped for bulk sampling of throughfall water and two lysimeters for soil water extraction at a depth of 15 cm and 45 cm, respectively. Rainwater was collected in 2 open areas outside the forest along with samples from a stream draining the forest and surrounding wetland. Water samples were collected 8 times between April 2003 and November 2004. Results show that the problem of acid rain is present in eastern Cape Breton Island. Canopy passage was found to lower the average rainwater pH from 4.7 to 4.2 with a related increase of sulfate from 2.2 ppm to 8.3 ppm. Top soil solution pH was 3.9 increasing to 4.5 in the sub soil. Aluminum was found to increase significantly in the soil solution when pH dropped below 4.2. This demonstrated that soil acidification due to acid rain frees the aluminum in the top soil. However, the concentration of metal was reduced at lower soil depth due to base cation exchange. High sodium concentrations in rainwater and throughfall were closely associated with sulfate values, indicating high inputs of saline oceanic spray with the potential to cause a salt effect in the top soil chemistry. Most water samples had very low nitrate concentrations. The water chemistry in the stream fluctuated with the pH, often dropping below 5 when sulfate contents increased during high run off events

  14. Multi-Cohort Stand Structural Classification: Ground- and LiDAR-based Approaches for Boreal Mixedwood and Black Spruce Forest Types of Northeastern Ontario

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    Kuttner, Benjamin George

    Natural fire return intervals are relatively long in eastern Canadian boreal forests and often allow for the development of stands with multiple, successive cohorts of trees. Multi-cohort forest management (MCM) provides a strategy to maintain such multi-cohort stands that focuses on three broad phases of increasingly complex, post-fire stand development, termed "cohorts", and recommends different silvicultural approaches be applied to emulate different cohort types. Previous research on structural cohort typing has relied upon primarily subjective classification methods; in this thesis, I develop more comprehensive and objective methods for three common boreal mixedwood and black spruce forest types in northeastern Ontario. Additionally, I examine relationships between cohort types and stand age, productivity, and disturbance history and the utility of airborne LiDAR to retrieve ground-based classifications and to extend structural cohort typing from plot- to stand-levels. In both mixedwood and black spruce forest types, stand age and age-related deadwood features varied systematically with cohort classes in support of an age-based interpretation of increasing cohort complexity. However, correlations of stand age with cohort classes were surprisingly weak. Differences in site productivity had a significant effect on the accrual of increasingly complex multi-cohort stand structure in both forest types, especially in black spruce stands. The effects of past harvesting in predictive models of class membership were only significant when considered in isolation of age. As an age-emulation strategy, the three cohort model appeared to be poorly suited to black spruce forests where the accrual of structural complexity appeared to be more a function of site productivity than age. Airborne LiDAR data appear to be particularly useful in recovering plot-based cohort types and extending them to the stand-level. The main gradients of structural variability detected using Li

  15. Proceedings of the US/FRG research symposium: effects of atmospheric pollutants on the spruce-fir forests of the Eastern United States and the Federal Republic of Germany

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    Gerard, tech. coord. Hertel; Gerard Hertel

    1988-01-01

    Includes 66 papers presented at the US/FRG research symposium: effects of atmospheric pollutants on the spruce-fir forests of the Eastern United States and the Federal Republic of Germany, which was held October 19-23, 1987, in Burlington, Vermont.

  16. BVOC emission in Norway spruce: the effect of stand structure, high temperature and ozone levels.

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    Pallozzi, Emanuele; Guidolotti, Gabriele; Večeřová, Kristýna; Esposito, Raffaela; Lusini, Ilaria; Juráň, Stanislav; Urban, Otmar; Calfapietra, Carlo

    2015-04-01

    Norway spruce (Picea abies L.) is a widely distributed conifer species in the boreal zone and mountain areas of central Europe and is a moderate emitter of volatile organic compounds (BVOC). Although the vaporization and diffusion processes from resin ducts were generally considered to be the main processes for monoterpene emissions in conifers, recently it has been showed that a significant portion (up to one third) of monoterpene emissions of Norway spruce can originate from novel biosynthesis, thus depending on photosynthetic processes. For this reason, both biosynthesis and emission are strongly influenced by the environment and the stand structure. They increase with both increasing light and temperature during the warmer periods, although those are the periods with the higher ozone concentration that usually act as an inhibitor of both assimilation and isoprenoids synthesis and emission. On the other hand, stand structure can play an important role, because the photosynthetic capacity is influenced by temperature and light conditions through the canopy. In order to assess the effects of stand structure, temperature and ozone on isoprenoids emission of Norway spruce we carried out field and laboratory experiments. In the experimental field campaigns we measured: assimilation and BVOC emission from needles of sun and shade layers within the canopy of the spruce forest present at the Bily Kriz experimental research site (Moravian-Silesian Beskydy Mountains, 49° 33' N, 18° 32' E, NE of Czech Republic, 908 m a.s.l.). Moreover in the same layers we measured continuously concentration of BVOCs in the air using a PTR-TOF-MS. In laboratory we analyzed the effects of short-term exposure to high temperature and high ozone concentrations on branches of spruce trees collected at the Bily Kriz experimental research site. Preliminary results show that in Norway spruce both stand structure and environmental conditions influenced the gas exchange and BVOC emission rates

  17. Cold tolerance and photosystem function in a montane red spruce population: physiological relationships with foliar carbohydrates

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    P.G. Shaberg; G.R. Strimbeck; G.J. Hawley; D.H. DeHayes; J.B. Shane; P.F. Murakami; T.D. Perkins; J.R. Donnelly; B.L. Wong

    2000-01-01

    Red spruce (Picea rubens Sarg.) growing in northern montane forests of eastern North America appears to be distinctive with respect to at least two aspects of winter physiology. First, red spruce attains only a modest level of midwinter cold tolerance compared to other north temperate conifers and appears barely capable of avoiding freezing injury at...

  18. Ecosystem structure and function in the SPRUCE chambers at fine resolution

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    Glenn, N. F.; Graham, J.; Spaete, L.; Hanson, P. J.

    2017-12-01

    The Spruce and Peatland Responses Under Climatic and Environmental change (SPRUCE; operated by DOE's Oak Ridge National Laboratory) aims to assess biological and ecological responses in a peat bog to a range of increased temperatures and the presence of elevated atmospheric CO2 concentrations. We are using terrestrial laser scanning (TLS) to monitor vegetation productivity and hummock-hollow structure at cm-scale in the SPRUCE plots to complement in-situ measurements of gross and net primary production. The hummock-hollow peatland microtopography is associated with fluctuating water levels and sphagnum mosses, and ultimately controls C and methane cycling. We estimate tree growth by calculating increases in tree height and canopy voxel volume between years with the TLS data. Microtopography is also characterized over time with TLS but by using gridded cells to classify regions into hummocks or hollows. Spectroscopy to quantify water content in the sphagnum is used to further classify these microtopographic regions. As multiple years of data collection occur, we will couple our fine-scale remote sensing measurements with in-situ measurements of CO2 and CH4 flux measures to capture species-specific productivity responses to warming and increased CO2.

  19. Fire severity unaffected by spruce beetle outbreak in spruce-fir forests in southwestern Colorado.

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    Andrus, Robert A; Veblen, Thomas T; Harvey, Brian J; Hart, Sarah J

    2016-04-01

    Recent large and severe outbreaks of native bark beetles have raised concern among the general public and land managers about potential for amplified fire activity in western North America. To date, the majority of studies examining bark beetle outbreaks and subsequent fire severity in the U.S. Rocky Mountains have focused on outbreaks of mountain pine beetle (MPB; Dendroctonus ponderosae) in lodgepole pine (Pinus contorta) forests, but few studies, particularly field studies, have addressed the effects of the severity of spruce beetle (Dendroctonus rufipennis Kirby) infestation on subsequent fire severity in subalpine Engelmann spruce (Picea engelmannii) and subalpine fir (Abies lasiocarpa) forests. In Colorado, the annual area infested by spruce beetle outbreaks is rapidly rising, while MPB outbreaks are subsiding; therefore understanding this relationship is of growing importance. We collected extensive field data in subalpine forests in the eastern San Juan Mountains, southwestern Colorado, USA, to investigate whether a gray-stage (fire) spruce beetle infestation affected fire severity. Contrary to the expectation that bark beetle infestation alters subsequent fire severity, correlation and multivariate generalized linear regression analysis revealed no influence of pre-fire spruce beetle severity on nearly all field or remotely sensed measurements of fire severity. Findings were consistent across moderate and extreme burning conditions. In comparison to severity of the pre-fire beetle outbreak, we found that topography, pre-outbreak basal area, and weather conditions exerted a stronger effect on fire severity. Our finding that beetle infestation did not alter fire severity is consistent with previous retrospective studies examining fire activity following other bark beetle outbreaks and reiterates the overriding influence of climate that creates conditions conducive to large, high-severity fires in the subalpine zone of Colorado. Both bark beetle outbreaks and

  20. Understory Structure and Vascular Plant Diversity in Naturally Regenerated Deciduous Forests and Spruce Plantations on Similar Clear-Cuts: Implications for Forest Regeneration Strategy Selection

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    ZhiQiang Fang

    2014-04-01

    Full Text Available The active effect of natural regeneration on understory vegetation and diversity on clear-cut forestlands, in contrast to conifer reforestation, is still controversial. Here we investigated differences in understory vegetation by comparing naturally regenerated deciduous forests (NR and reforested spruce plantations (SP aged 20–40 years on 12 similar clear-cuts of subalpine old-growth spruce-fir forests from the eastern Tibetan Plateau. We found that 283 of the 334 vascular plant species recorded were present in NR plots, while only 264 species occurred in SP plots. This was consistent with richer species, higher cover, and stem (or shoot density of tree seedlings, shrubs, and ferns in the NR plots than in the SP plots. Moreover, understory plant diversity was limited under dense canopy cover, which occurred more frequently in the SP plots. Our findings implied that natural deciduous tree regeneration could better preserve understory vegetation and biodiversity than spruce reforestation after clear-cutting. This result further informed practices to reduce tree canopy cover for spruce plantations or to integrate natural regeneration and reforestation for clear-cuts in order to promote understory vegetation and species diversity conservation.

  1. Climate-induced mortality of spruce stands in Belarus

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    Kharuk, Viacheslav I.; Im, Sergei T.; Dvinskaya, Maria L.; Golukov, Alexei S.; Ranson, Kenneth J.

    2015-12-01

    The aim of this work is an analysis of the causes of spruce (Picea abies L.) decline and mortality in Belarus. The analysis was based on forest inventory and Landsat satellite (land cover classification, climate variables (air temperature, precipitation, evaporation, vapor pressure deficit, SPEI drought index)), and GRACE-derived soil moisture estimation (equivalent of water thickness anomalies, EWTA). We found a difference in spatial patterns between dead stands and all stands (i.e., before mortality). Dead stands were located preferentially on relief features with higher water stress risk (i.e., higher elevations, steeper slopes, south and southwestern exposure). Spruce mortality followed a series of repeated droughts between 1990 and 2010. Mortality was negatively correlated with air humidity (r = -0.52), and precipitation (r = -0.57), and positively correlated with the prior year vapor pressure deficit (r = 0.47), and drought increase (r = 0.57). Mortality increased with the increase in occurrence of spring frosts (r = 0.5), and decreased with an increase in winter cloud cover (r = -0.37). Spruce mortality was negatively correlated with snow water accumulation (r = -0.81) and previous year anomalies in water soil content (r = -0.8). Weakened by water stress, spruce stands were attacked by pests and phytopathogens. Overall, spruce mortality in Belarussian forests was caused by drought episodes and drought increase in synergy with pest and phytopathogen attacks. Vast Picea abies mortality in Belarus and adjacent areas of Russia and Eastern Europe is a result of low adaptation of that species to increased drought. This indicates the necessity of spruce replacement by drought-tolerant indigenous (e.g., Pinus sylvestris, Querqus robur) or introduced (e.g., Larix sp. or Pseudotsuga menzieslii) species to obtain sustainable forest growth management.

  2. Performance of the Forest Vegetation Simulator in managed white spruce plantations influenced by eastern spruce budworm in northern Minnesota

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    Matthew B. Russell; Anthony W. D' Amato; Michael A. Albers; Christopher W. Woodall; Klaus J. Puettmann; Michael R. Saunders; Curtis L. VanderSchaaf

    2015-01-01

    Silvicultural strategies such as thinning may minimize productivity losses from a variety of forest disturbances, including forest insects. This study analyzed the 10-year postthinning response of stands and individual trees in thinned white spruce (Picea glauca [Moench] Voss) plantations in northern Minnesota, USA, with light to moderate defoliation...

  3. In vivo function of Pgβglu-1 in the release of acetophenones in white spruce

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    Melissa H. Mageroy

    2017-07-01

    Full Text Available Eastern spruce budworm (Choristoneura fumiferiana Clemens (ESBW is a major forest pest which feeds on young shoots of white spruce (Picea glauca and can cause landscape level economic and ecological losses. Release of acetophenone metabolites, piceol and pungenol, from their corresponding glycosides, picein and pungenin, can confer natural resistance of spruce to ESBW. A beta-glucosidase gene, Pgβglu-1, was recently discovered and the encoded enzyme was characterized in vitro to function in the release of the defensive acetophenone aglycons. Here we describe overexpression of Pgβglu-1 in a white spruce genotype whose metabolome contains the glucosylated acetophenones, but no detectable amounts of the aglycons. Transgenic overexpression of Pgβglu-1 resulted in release of the acetophenone aglycons in planta. This work provides in vivo evidence for the function of Pgβglu-1.

  4. Four millennia of woodland structure and dynamics at the Arctic treeline of eastern Canada.

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    Auger, Sarah; Payette, Serge

    2010-05-01

    Paleoecological analysis using complementary indicators of vegetation and soil can provide spatially explicit information on ecological processes influencing trajectories of long-term ecosystem change. Here we document the structure and dynamics of an old-growth woodland before and after its inception 1000 years ago. We infer vegetation and soil characteristics from size and age distributions of black spruce (Picea mariana (Mill.) B.S.P.), soil properties, plant fossils, and paleosols. Radiocarbon ages of charcoal on the ground and in the soil indicate that the fire return interval was approximately 300 years between 2750 and 1000 cal. yr BP. No fire evidence was found before and after this period despite the presence of spruce since 4200 cal. yr BP. The size structures of living and dead spruce suggest that the woodland is in equilibrium with present climate in absence of fire. Tree establishment and mortality occurred regularly since the last fire event around 950 cal. yr BP. Both layering and occasional seeding have contributed to stabilize the spatial distribution of spruce over the past 1000 years. Since initial afforestation, soil development has been homogenized by the changing spatial distribution of spruce following each fire. We conclude that the history of the woodland is characterized by vegetation shifts associated with fire and soil disturbances and by millennial-scale maintenance of the woodland's structure despite changing climatic conditions.

  5. Soil chemistry and nutrition of North American spruce-fir stands: Evidence of recent change

    International Nuclear Information System (INIS)

    Joslin, J.D.; Kelly, J.M.; Van Miegroet, H.

    1992-01-01

    One set of hypotheses offered to explain the decline of red spruce (Picea rubens Sarg.) in eastern North America focuses on the effect of acidic deposition on soil chemistry changes that may affect nutrient availability and root function. Long-term soils data suggests that soil acidification has occurred in some spruce stands over the past 50 yr, with plant uptake and cation leaching both contributing to the loss of cations. Studies of tree ring chemistry also have indicated changes in Ca/Al and Mg/Al ratios in red spruce wood, suggesting increases in the ionic strength of soil solution. Irrigation studies using strong acid inputs have demonstrated accelerated displacement of base cations from upper horizons. Spruce-fir (Abies spp.) nutrient budgets indicate that current net Ca and Mg leaching loss rates are of the same order of magnitude as losses to whole tree harvest removals, spread out over a 50-yr rotation. For most cations, red spruce foliar nutrient levels decline with elevation, but it is difficult to assess the contribution of the elevational gradient in atmospheric deposition to this pattern. Compared to northeastern sites, spruce-fir soil solutions in the southern Appalachians have higher nitrate levels and higher Al concentrations, which at times approach the Al toxicity threshold for red spruce seedlings and frequently are at levels known to interfere with cation uptake. There is little evidence that either nutrient deficiencies or Al toxicity are primary causes of red spruce decline in the Northeast, though both may play a role in the Southeast

  6. GENETIC STRUCTURE OF NORWAY SPRUCE (PICEA ABIES): CONCORDANCE OF MORPHOLOGICAL AND ALLOZYMIC VARIATION.

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    Lagercrantz, Ulf; Ryman, Nils

    1990-02-01

    This study describes the population structure of Norway spruce (Picea abies) as revealed by protein polymorphisms and morphological variation. Electrophoretically detectable genetic variability was examined at 22 protein loci in 70 populations from the natural range of the species in Europe. Like other conifers, Norway spruce exhibits a relatively large amount of genetic variability and little differentiation among populations. Sixteen polymorphic loci (73%) segregate for a total of 51 alleles, and average heterozygosity per population is 0.115. Approximately 5% of the total genetic diversity is explained by differences between populations (G ST = 0.052), and Nei's standard genetic distance is less than 0.04 in all cases. We suggest that the population structure largely reflects relatively recent historical events related to the last glaciation and that Norway spruce is still in a process of adaptation and differentiation. There is a clear geographic pattern in the variation of allele frequencies. A major part of the allelefrequency variation can be accounted for by a few synthetic variables (principal components), and 80% of the variation of the first principal component is "explained" by latitude and longitude. The central European populations are consistently depauperate of genetic variability, most likely as an effect of severe restrictions of population size during the last glaciation. The pattern of differentiation at protein loci is very similar to that observed for seven morphological traits examined. This similarity suggests that the same evolutionary forces have acted upon both sets of characters. © 1990 The Society for the Study of Evolution.

  7. Interactions between Bacillus thuringiensis and parasitoids of late-instar larvae of the spruce budworm (Lepidoptera: Tortricidae)

    NARCIS (Netherlands)

    Schoenmaker, A.; Cusson, M.; Frankenhuyzen, van K.

    2001-01-01

    We investigated interactions between Bacillus thuringiensis Berliner var. kurstaki and parasitoids that attack late instars of the eastern spruce budworm, Choristoneura fumiferana (Clemens). In a petri-dish arena, females of Tranosema rostrale rostrale (Brishke) (Hymenoptera: Ichneumonidae) were

  8. Insect-induced tree mortality of boreal forests in eastern Canada under a changing climate.

    Science.gov (United States)

    Zhang, Xiongqing; Lei, Yuancai; Ma, Zhihai; Kneeshaw, Dan; Peng, Changhui

    2014-06-01

    Forest insects are major disturbances that induce tree mortality in eastern coniferous (or fir-spruce) forests in eastern North America. The spruce budworm (SBW) (Choristoneura fumiferana [Clemens]) is the most devastating insect causing tree mortality. However, the relative importance of insect-caused mortality versus tree mortality caused by other agents and how this relationship will change with climate change is not known. Based on permanent sample plots across eastern Canada, we combined a logistic model with a negative model to estimate tree mortality. The results showed that tree mortality increased mainly due to forest insects. The mean difference in annual tree mortality between plots disturbed by insects and those without insect disturbance was 0.0680 per year (P eastern Canada but that tree mortality induced by insect outbreaks will decrease in eastern Canada under warming climate.

  9. TALL HERB SPRUCE FORESTS AS CLIMAX COMMUNITIES ON LOWLAND SWAMPS OF BRYANSK POLESIE

    Directory of Open Access Journals (Sweden)

    O. I. Evstigneev

    2017-09-01

    Full Text Available Nettle grey alder forests are a dominant forest type on lowland swamps in the Bryansk Polesie. They are formed as a result of repeated cuttings in the place of tall herb spruce forests. Tall herb spruce forests are very rare communities in the vegetation cover in this area due to clear cutting, melioration and peat extraction. An assessment of the succession status of tall herb spruce forests and nettle grey alder forests was carried out in this paper. The criteria of climax state and succession state of communities, developed for Eastern European forests, were used. These criteria are based on the degree of intensity of the following signs in the community: 1 the completeness of species composition of tree synusia; 2 the ontogenetic structure of tree species cenopopulation; 3 the gap-mosaic stand structure; 4 the diversity of microsites in soil cover; 5 the completeness of species composition and ecological-coenotic diversity of vascular species. We showed that tall herb spruce forest, as opposed to black alder forest, is close to communities of the climax type. This is evidenced by the following features of cenosis: firstly, all tree species in the area that covers the Bryansk Polesie and that are able to grow on lowland swamps are represented in the spruce forest (Alnus glutinosa, Betula pubescens, Fraxinus excelsior, Padus avium, Picea abies, Salix pentandra, Sorbus aucuparia, Ulmus glabra. Secondly, a steady turnover of generations is carried out in the cenopopulations of main edificators (Picea abies and Alnus glutinosa. This is evidenced by the complete and left-sided structure of their ontogenetic spectrum. Thirdly, a system of asynchronously developing gaps (parcels, which are formed on the site of old tree falls, is formed in the community. This ensures the continuous renewal of spruce and alder populations and creates conditions for the regeneration of other tree species. Fourthly, the structure of biogenic microsites has been formed

  10. Third year effects of cloudwater and ozone on red spruce seedlings

    International Nuclear Information System (INIS)

    Pier, P.A.; Thornton, F.C.; McDuffie, C. Jr.

    1991-01-01

    The reduction in growth of high elevation red spruce in the eastern US has been attributed in part to greater exposure to atmospheric pollution which occurs at high elevation. The authors objective was to evaluate the impact of ambient ozone and cloudwater deposition on the growth of red spruce seedlings at a high elevation site. Potted native and Phyton-grown (Phyton Technologies) red spruce seedlings were exposed in open-top field chambers at Whitetop Mountain, Virginia (elevation 1,680) for the third season to treatments of: (1) exclusion of clouds and 50% reduction in ambient O 3 (COE), (2) O 3 with clouds excluded (CO), (3) exposure to clouds and O 3 , as control chambers (CC), and (4) open plots (AA). Plant biomass components and diameter increment growth for both seedling types were not affected by treatments. Photosynthesis was not enhanced by removal of cloudwater and O 3 . Respiration (R d ) generally was not affected by treatments; however, R d in native seedling needles of previous year and two-year previous growth was significantly greater in CC than CO and COE on several sampling dates, indicating that cloudwater and O 3 may be causing higher R d

  11. Indicators of Population Viabllity in Red Spruce, Picea rubens. I. Reproductive Traits and Fecundity

    Science.gov (United States)

    A. Mosseler; J.E. Major; J.D. Simpson; B. Daigle; K. Lange; Y.S. Park; K.H Johnsen; O.P. Rajora

    2000-01-01

    Red spruce (Picea rubens Sarg.) has experienced a substantial decline across most of its range in eastern North America over the past centmy and probably also in the disjunct Ontario populations where it now occurs only in small isolated stands. Measurements of cone and seed traits from natural populations were used as indicators of the...

  12. Does the orientation of Norway spruce (Picea abies /L./ Karst.) branches within sunlit crown part influence the heterogeneity of biochemical, structural and spectral characteristics of needles?

    NARCIS (Netherlands)

    Lhotáková, Z.; Albrechtová, J.; Malenovsky, Z.; Rock, B.N.; Polák, T.; Cudlín, P.

    2007-01-01

    The goal of this study was to determine if selected biochemical, structural and spectral properties of Norway spruce needles are influenced by the azimuth orientation of the branch. Three youngest needle age classes from 20 mature (100 years old or older) Norway spruce trees were sampled from upper

  13. Bioecology of the conifer swift moth, Korscheltellus gracilis, a root feeder associated with spruce-fir decline

    Science.gov (United States)

    William E. Wallner; David L. Wagner; Bruce L. Parker; Donald L. Tobi

    1991-01-01

    During the past two decades, the decline of red spruce, Picea rubens Sargent, and balsam fir, Abies balsamea (L), at high elevations (900-1200 m) in eastern North America has evoked concern about the effects of anthropogenic deposition upon terrestrial ecosystems. In many high-elevation forests across New England, as many as 50...

  14. Laser microprobe mass analysis (LAMMA) of aluminum and lead in fine roots and their ectomycorrhizal mantles of Norway spruce (Picea abies (L.) Karst.).

    Science.gov (United States)

    Eeckhaoudt, S; Vandeputte, D; Van Praag, H; Van Grieken, R; Jacob, W

    1992-03-01

    Fine roots and ectomycorrhizal root tips were sampled in a Norway spruce (Picea abies (L.) Karst.) stand in the eastern part of the Belgian Ardennes. The cellular and partly subcellular localizations of aluminum and lead were identified by the micro-analytical laser microprobe mass analysis (LAMMA) technique. In fine roots with secondary structure, localization of aluminum was limited to the peripheral cell layers. Lead was found in the outer layers, and also in the primary phloem. Aluminum penetrated the mycorrhizal mantle, but lead was seldom detected in ectomycorrhizae.

  15. Transmittance of young Norway spruce stand canopy for photosynthetically active radiation during the growing season

    International Nuclear Information System (INIS)

    Markova, I.; Kubasek, J.

    2013-01-01

    Analysis of transmittance of young Norway spruce stand canopy for photosynthetically active radiation (PAR) was made at the study site of Bily Kriz (the Moravian-Silesian Beskids Mts., the Czech Republic) at different sky conditions during the growing season in 2010. For the description of PAR transmittance different phenological phases of the spruce stand development in clear and overcast days were chosen. The mean daily PAR transmittance of the spruce canopy was significantly higher in overcast days compared with clear ones. Diffuse PAR thus penetrated into lower parts of the canopy more efficiently than direct one. PAR transmittance of young Norway spruce stand canopy was different in individual phenological phases of the spruce stand canopy which was caused by changes in the stand structure during the growing season. Thus monitoring of transmittance of young Norway spruce stand canopy for PAR can help to describe the development of spruce stand canopy

  16. Damage by the Sitka spruce weevil (Pissodes strobi) and growth patterns for 10 spruce species and hybrids over 26 years in the Pacific Northwest.

    Science.gov (United States)

    Russel G. Mitchell; Kenneth H. Wright; Norman E. Johnson

    1990-01-01

    Ten species and hybrids of spruce (Picea spp.) were planted and observed annually for 26 years at three coastal locations in Oregon and Washington to evaluate growth rates and susceptibility to the Sitka spruce weevil (= white pine weevil), Pissodes strobi The 10 spruce were: Sitka spruce, Norway spruce, Lutz spruce, black...

  17. Effects of climate change on fire and spruce budworm disturbance regimes and consequences on forest biomass production in eastern Canada

    International Nuclear Information System (INIS)

    Gauthier, S.

    2004-01-01

    The dynamics of spruce budworm (SBW) outbreaks and wildfires are expected to change as climatic change progresses. The effects of an altered, combined interaction between SBW and fire may be of greater importance than the individual effect of either on forest biomass production. The objectives of this study are to define current fire and SBW regimes in eastern Canada and relate the characteristics of each regime based upon climate model outputs for 2050 and 2100. The study also attempts to evaluate the impact of predicted changes in SBW and fire disturbance regimes on forest dynamics. The methodology used in the study included data from the Canadian Large Fire Database and historical records of SBW outbreaks. Spatial and environmental variables were presented along with climate models. The analysis was conducted using constrained ordination techniques, and canonical correspondence and redundancy analysis. Projected disturbance regimes were presented for both fire and SBW. The effects of the regimes on biomass productivity were also examined, using a Landscape Disturbance Simulator (LAD). It was concluded that this model will help evaluate the consequences of changes imposed by climatic change on both disturbances individually, as well as their interaction. 10 refs., 1 tab., 2 figs

  18. Cytophotometric differentiation of high elevation spruces: physiological and ecological implications

    International Nuclear Information System (INIS)

    Berlyn, G.P.; Royte, J.L.; Anoruo, A.O.

    1990-01-01

    Red and black spruce and their hybrids can be determined by morphological indices; however, the criteria are somewhat subjective and increasingly difficult to use at higher elevations. Although the chromosome number is identical (2n = 24), red spruce has twice as much nuclear DNA (48 pg) than black spruce (24 pg) and thus the species and their hybrids can also be separated by cytophotometry. This is relevant to spruce decline studies because black spruce is much more resistant to high elevation environmental stresses, both natural and anthropogenic. It also has implications for the effect of climatic changes on the composition of high elevation spruce-fir forests because red spruce can outcompete black spruce under more mesic conditions. Four elevation transects sampling spruce on the east and west sides of Mount Washington (New Hampshire) and Camels Hump (Vermont) and a single transect on the southwest side of Whiteface Mountain (New York) were made to investigate the degree of hybridization and introgression between these two species. A positive correlation was found between increased elevation and increased black spruce genes on Mount Washington and Camels Hump. Pure black spruce was found on Mount Washington from 1356 m to 1582 m. No pure black or red spruce was found on Camels Hump although the proportion of red spruce alleles was significantly greater on Camels Hump. All trees sampled at all elevations on Whiteface Mountain were pure red spruce. Thus the proportion of black spruce alleles in high elevation spruce populations decreases from east to west. This closely parallels the increase in spruce decline which increases from east to west. (author)

  19. Forest Floor Carbon Exchange of a Boreal Black Spruce Forest in Eastern Canada

    Science.gov (United States)

    Bergeron, O.; Margolis, H. A.; Coursolle, C.

    2009-06-01

    This study reports continuous automated measurements of forest floor carbon (C) exchange over feathermoss, lichen, and sphagnum micro-sites in a black spruce forest in eastern North America during snow-free periods over three years. The response of soil respiration (Rs-auto) and forest floor photosynthesis (Pff) to environmental factors was determined. The seasonal contributions of scaled up Rs-auto adjusted for spatial representativeness (Rs-adj) and Pff (Pff-eco) relative to that of total ecosystem respiration (Re) and photosynthesis (Peco), respectively, were also quantified. Shallow soil temperature explained 67-86% of the variation in Rs-auto for all ground cover types, while deeper soil temperatures were related to Rs-auto only for the feathermoss micro-sites. Base respiration was consistently lower under feathermoss, intermediate under sphagnum, and higher under lichen during all three years. The Rs-adj/Re ratio increased from spring through autumn and ranged from 0.85 to 0.87 annually for the snow-free period. The Rs-adj/Re ratio was negatively correlated with the difference between air and shallow soil temperature and this correlation was more pronounced in autumn than summer and spring. Maximum photosynthetic capacity of the forest floor (Pffmax) saturated at low irradiance levels (~200 μmol m-2 s-1) and decreased with increasing air temperature and vapor pressure deficit for all three ground cover types, suggesting that Pff was more limited by desiccation than by light availability. Pffmax was lowest for sphagnum, intermediate for feathermoss, and highest for lichen for two of the three years. Pff normalized for light peaked at air temperatures of 5-8°C, suggesting that this is the optimal temperature range for Pff. The Pff-eco/Peco ratio varied seasonally from 13 to 24% and reached a minimum in mid-summer when both air temperature and Peco were at their maximum. On an annual basis, Pff-eco accounted for 17-18% of Peco depending on the year and the

  20. Effectiveness of polyethylene sheeting in controlling spruce beetles (Coleoptera: Scolytidae) in infested stacks of spruce firewood in Alaska.

    Science.gov (United States)

    Edward H. Holsten; Richard A. Werner

    1993-01-01

    Covering stacks of spruce firewood with either clear or black polyethylene sheeting does not raise log temperatures high enough to kill spruce beetle brood in the logs. Based on the results of this study, we do not recommend the use of polyethylene sheeting as a remedial measure for the reduction of spruce beetle brood in infested firewood or log decks in south-central...

  1. Pelletizing properties of torrefied spruce

    DEFF Research Database (Denmark)

    Stelte, Wolfgang; Clemons, Craig; Holm, Jens K.

    2011-01-01

    analysis revealed a cohesive failure mechanism due to strong inter-particle bonding in spruce pellets as a resulting from a plastic flow of the amorphous wood polymers, forming solid polymer bridges between adjacent particles. Fracture surfaces of pellets made from torrefied spruce possessed gaps and voids...

  2. Space sequestration below ground in old-growth spruce-beech forests-signs for facilitation?

    Science.gov (United States)

    Bolte, Andreas; Kampf, Friederike; Hilbrig, Lutz

    2013-01-01

    Scientists are currently debating the effects of mixing tree species for the complementary resource acquisition in forest ecosystems. In four unmanaged old-growth spruce-beech forests in strict nature reserves in southern Sweden and northern Germany we assessed forest structure and fine rooting profiles and traits (≤2 mm) by fine root sampling and the analysis of fine root morphology and biomass. These studies were conducted in selected tree groups with four different interspecific competition perspectives: (1) spruce as a central tree, (2) spruce as competitor, (3) beech as a central tree, and (4) beech as competitor. Mean values of life fine root attributes like biomass (FRB), length (FRL), and root area index (RAI) were significantly lower for spruce than for beech in mixed stands. Vertical profiles of fine root attributes adjusted to one unit of basal area (BA) exhibited partial root system stratification when central beech is growing with spruce competitors. In this constellation, beech was able to raise its specific root length (SRL) and therefore soil exploration efficiency in the subsoil, while increasing root biomass partitioning into deeper soil layers. According to relative values of fine root attributes (rFRA), asymmetric below-ground competition was observed favoring beech over spruce, in particular when central beech trees are admixed with spruce competitors. We conclude that beech fine rooting is facilitated in the presence of spruce by lowering competitive pressure compared to intraspecific competition whereas the competitive pressure for spruce is increased by beech admixture. Our findings underline the need of spatially differentiated approaches to assess interspecific competition below ground. Single-tree approaches and simulations of below-ground competition are required to focus rather on microsites populated by tree specimens as the basic spatial study area.

  3. Spruce needles used as radioecological biotracers

    International Nuclear Information System (INIS)

    Seidel, C.; Gruber, V.; Baumgartner, A.

    2009-01-01

    In a two years project spruce needle samples of the Austrian Bioindicator Grid were analysed by gamma-ray spectrometry to investigate the spatial and temporal distribution of radionuclides in spruce needles of the last 25 years with the main focus on the radioactive contamination before and after the Chernobyl fallout 1986. More than 600 spruce needle samples at selected locations of the Bioindicator Grid were analysed for different natural and anthropogenic radionuclides: 137 Cs, 40 K, 210 Pb, 226 Ra, 228 Ra, 238 U. Additionally, soil samples were taken at selected sites to study the soil-to-plant transfer. This radioecological evaluation is an important part of an existing environmental surveillance programme in Upper Austria in order to gain basic information on the impact of environmental changes on the radioecological behaviour of spruce trees. (orig.)

  4. Spruce aphid (Elatobium abietinum Walker) (Hemiptera: Aphididae) [Chapter XXIV

    Science.gov (United States)

    Ann M. Lynch

    2014-01-01

    Elatobium abietinum Walker is a spruce-feeding aphid that in Europe is referred to as the green spruce aphid (Day et al., 1998a) (Fig. 1). However, in North America E. abietinum is known simply as the spruce aphid, while the common name "green spruce aphid" refers to a different species, Cinara fornacula Hottes (Hemiptera: Aphididae) (http://www.entsoc.org/...

  5. Space sequestration below ground in old-growth spruce-beech forests – signs for facilitation?

    Directory of Open Access Journals (Sweden)

    Andreas eBolte

    2013-08-01

    Full Text Available Scientists are currently debating the effects of mixing tree species for the complementary resource acquisition in forest ecosystems. In four unmanaged old-growth spruce-beech forests in strict nature reserves in southern Sweden and northern Germany we assessed forest structure and fine rooting profiles and traits (≤ 2 mm by fine root sampling and the analysis of fine root morphology and biomass. These studies were conducted in selected tree groups with four different interspecific competition perspectives: (1 spruce as a central tree, (2 spruce as competitor, (3 beech as a central tree, and (4 beech as competitor. Mean values of life fine root attributes like biomass (FRB, length (FRL, and root area index (RAI were significantly lower for spruce than for beech in mixed stands. Vertical profiles of fine root attributes adjusted to one unit of basal area (BA exhibited partial root system stratification when central beech is growing with spruce competitors. In this constellation, beech was able to raise its specific root length (SRL and therefore soil exploration efficiency in the subsoil, while increasing root biomass partitioning into deeper soil layers. According to relative values of fine root attributes (rFRA, asymmetric below-ground competition was observed favoring beech over spruce, in particular when central beech trees are admixed with spruce competitors. We conclude that beech fine rooting is facilitated in the presence of spruce by lowering competitive pressure compared to intraspecific competition whereas the competitive pressure for spruce is increased by beech admixture. Our findings underline the need of spatially differentiated approaches to assess interspecific competition below ground. Single-tree approaches and simulations of below-ground competition are required to focus rather on microsites populated by tree specimens as the basic spatial study area.

  6. Densities of breeding birds and changes in vegetation in an alaskan boreal forest following a massive disturbance by spruce beetles

    Science.gov (United States)

    Matsuoka, S.M.; Handel, C.M.; Ruthrauff, D.R.

    2001-01-01

    We examined bird and plant communities among forest stands with different levels of spruce mortality following a large outbreak of spruce beetles (Dendroctonus rufipennis (Kirby)) in the Copper River Basin, Alaska. Spruce beetles avoided stands with black spruce (Picea mariana) and selectively killed larger diameter white spruce (Picea glauca), thereby altering forest structure and increasing the dominance of black spruce in the region. Alders (Alnus sp.) and crowberry (Empetrum nigrum) were more abundant in areas with heavy spruce mortality, possibly a response to the death of overstory spruce. Grasses and herbaceous plants did not proliferate as has been recorded following outbreaks in more coastal Alaskan forests. Two species closely tied to coniferous habitats, the tree-nesting Ruby-crowned Kinglet (Regulus calendula) and the red squirrel (Tamiasciurus hudsonicus), a major nest predator, were less abundant in forest stands with high spruce mortality than in low-mortality stands. Understory-nesting birds as a group were more abundant in forest stands with high levels of spruce mortality, although the response of individual bird species to tree mortality was variable. Birds breeding in stands with high spruce mortality likely benefited reproductively from lower squirrel densities and a greater abundance of shrubs to conceal nests from predators.

  7. Establishing trees on cut-over peatlands in eastern Canada

    Directory of Open Access Journals (Sweden)

    J. Bussières

    2008-12-01

    Full Text Available Four major tree-planting trials on cut-over peatlands in eastern Canada were surveyed in 2002, in order to evaluate the potential use of trees in rehabilitation following horticultural peat extraction. At one of the sites, an experiment to determine the appropriate fertilisation rate for trees planted on cut-over peatlands was also conducted over several years. Tree performance was assessed by measuring survival, total height and annual growth of red maple (Acer rubrum L., tamarack (Larix laricina (Du Roi Koch., black spruce (Picea mariana (Mill. B.S.P., jack pine (Pinus banksiana Lamb. and hybrid poplar (Populus spp.. Establishment and growth of tamarack and black spruce in cut-over peatlands showed good potential when compared to performance in conventional forestry plantations. Red maple and jack pine gave poor productivity but promising survival, whilst hybrid poplar plantings failed. Adding nutrients was essential for growth but dosages above 122.5 g of 3.4N-8.3P-24.2K per tree gave no further improvement. Therefore, several different tree species can be planted to reclaim cut-over peatlands in eastern Canada, so long as the appropriate species are chosen and nutrients are provided.

  8. Old lower stem bark lesions apparently caused by unsuccessful spruce beetle attacks still evident on live spruce trees years later

    Science.gov (United States)

    John S. Hard; Ken P. Zogas

    2010-01-01

    We examined old bark lesions on Lutz spruce in young stands on the Kenai Peninsula, Alaska, to determine their cause. Distribution of these lesions along lower stems was similar to the distribution of spruce beetle attacks during epidemics. These lesions apparently resulted from unsuccessful attacks by spruce beetles during the late 1980s and early 1990s and appear to...

  9. A comparison of structural characteristics and ecological factors between forest reserves and managed silver fir - Norway spruce forests in Slovenia

    International Nuclear Information System (INIS)

    Marinšek, A.; Diaci, J.

    2011-01-01

    In order to examine ecological, floristic and structural differences between the forest stands of managed and unmanaged silver fir - Norway spruce forests (Bazzanio trilobatae-Abietetum albae), twelve sample plots (25x25 m) were established in forest reserves and managed forests. Within the plots, subplots and microplots we conducted phytosociological and pedological surveys, analyses of the stand structure, natural regeneration and estimation of solar radiation. We determined that there are no significant differences in floristic composition and ecological factors between managed forest and forest reserve stands. The only variables that were significantly different were the solar radiation variables (ISF; TSF; DSF), vertical structure (cover indexes (CI)) and stand basal area. Small differences in the composition and the structure of the vegetation indicate that, as far as ecosystematic changes are concerned, managing these forests is not as significant as the soil conditions. Solar radiation had a major influence on natural regeneration. Indirect solar radiation seemed to be more important than direct solar radiation. We found a statistically significant positive correlation between silver fir and Norway spruce regeneration and indirect solar radiation and confirmed that the management of light is a significant factor in the management of regeneration. Another trend that was detected was an increase in the number of beech, which will have quite a large proportion in the upper tree layer of the next generation, especially in forest reserves

  10. Soil and soil cover changes in spruce forests after final logging

    Directory of Open Access Journals (Sweden)

    E. M. Lapteva

    2015-10-01

    Full Text Available Soil cover transformation and changes of morphological and chemical properties of Albeluvisols in clear-cuttings of middle taiga spruce forests were studied. The observed changes in structure and properties of podzolic texturally-differentiated soils at cuttings of spruce forests in the middle taiga subzone do not cause their transition to any other soil type. Soil cover of secondary deciduous-coniferous forests which replace cut forests are characterized with a varied soil contour and a combination of the main type of podzolic soils under undisturbed spruce forests. The increased surface hydromorphism in cut areas causes formation of complicated sub-types of podzolic texturally differentiated soils (podzolic surface-gley soils with microprofile of podzol and enlarges their ratio (up to 35–38 % in soil cover structure. Temporary soil over-wetting at the initial (5–10 years stage of after-cutting self-restoring vegetation succession provides for soil gleyzation, improves yield and segregation of iron compounds, increases the migratory activity of humic substances. Low content and resources of total nitrogen in forest litters mark anthropogenic transformation processes of podzolic soils at this stage. Later (in 30–40 years after logging, soils in cut areas still retain signs of hydromorphism. Forest litters are denser, less acidic and thick with a low weight ratio of organic carbon as compared with Albeluvisols of undisturbed spruce forest. The upper mineral soil horizons under secondary deciduous-coniferous forests contain larger amounts of total iron, its mobile (oxalate-dissolvable components, and Fe-Mn-concretions.

  11. Concentrations of dissolved organic carbon along an altitudinal gradient from Norway spruce forest to the mountain birch/alpine ecotone in Norway

    Energy Technology Data Exchange (ETDEWEB)

    Clarke, N.; Roesberg, I.; Aamlid, D.

    2005-07-01

    Concentrations of dissolved organic carbon (DOC) in soil water from the base of the soil organic layer were determined at three forest plots along an altitudinal gradient in eastern Norway. The lowest plot, at 830 m above sea level (a.s.l.), was in Norway spruce forest and there were additional plots at the ecotone between Norway spruce and mountain birch at 925 m a.s.l. and at the forest line (1000 m a.s.l.). DOC concentrations in soil water did not decrease uniformly with altitude although tree biomass, above-ground litterfall and the soil C pool all did so. Significant correlations between DOC and (H{sup +}) or electrical conductivity may reflect the contribution of DOC to solution acidity and the anionic charge, respectively. If mean temperature during the growing season increases, tree growth at any given altitude will tend to increase and the spruce-birch ecotone may move to a higher altitude than at present. Increased C inputs as litter to the soil might then lead to increasing DOC concentrations and fluxes in surface waters. (orig.)

  12. Structure, development and health status of spruce forests affected by air pollution in the western Krkonoše Mts. in 1979–2014

    Directory of Open Access Journals (Sweden)

    Král Jan

    2015-09-01

    Full Text Available The structure and health status of waterlogged or peaty spruce (Picea abies [L.] Karst. forests in the summit parts of the Krkonoše Mts. in the Czech Republic were studied in 1979–2014. The objective was to evaluate the stand structure, dead wood, trend of the health status and productivity on four permanent research plots (PRP in relation to air pollution (SO2 and NOx concentrations and climatic conditions (temperatures and precipitation amounts. Stand structure was evaluated on the base of the measured parameters of individual trees on PRP. The health status of trees was evaluated according to foliage, and their vitality was assessed according to their radial growth documented by dendrochronological analyses. The radial growth was negatively correlated with SO2 and NOx concentrations. Stand dynamics during the observation period was characterised by increased tree mortality, the presence of dead wood and reduction of stand density from 1983 to 1992, while the most severe impairment of health status and stand stability occurred in 1982–1987. The foliage mass of living trees has been gradually increasing since 1988, but no pronounced improvement of tree vitality was documented after the decrease in SO2 concentration. However, particularly physiologically weakened spruce trees were attacked by the European spruce bark beetle (Ips typographus. The process of forest damage is manifested not only by foliage reduction but also by symptoms of various necroses on the assimilatory organs. In terms of climatic data, the weather in April had the most important effect on radial growth. Diameter increment showed positive statistically significant correlation with temperature in growing season, but the precipitation effect was low.

  13. Ammonium assmilation in spruce ectomycorrhizas

    International Nuclear Information System (INIS)

    Chalot, M.; Brun, A.; Botton, B.; Stewart, G.

    1990-01-01

    Assimilation of labelled NH 4 + into amino acids has been followed in ectomycorrhizal roots of spruce. Over an 18 h period of NH 4 + feeding, Gln, Glu and Ala became the most abundant amino acids. Gln was also the most highly labelled amino acid during the experiment, followed by Glu and Ala. This result indicates that Gln synthesis is an important ammonium utilization reaction in spruce mycorrhizas. Addition of MSX to NH 4 + fed mycorrhizas caused an inhibition of Gln accumulation with a corresponding increase in Glu, Ala and Asn levels. The supply of MSX induced a sharp diminution of 15 N enrichment in both amino and amido groups of glutamine. In contrast, the 15 N incorporation into Glu and derivatives (Ala and Asp) remained very high. This study demonstrates that the fungal glutamate dehydrogenase is quite operative in spruce ectomycorrhizas since it is able to sustain ammonium assimilation when glutamine synthetase is inhibited

  14. Forest floor carbon exchange of a boreal black spruce forest in eastern North America

    Science.gov (United States)

    Bergeron, O.; Margolis, H. A.; Coursolle, C.

    2009-09-01

    This study reports continuous automated measurements of forest floor carbon (C) exchange over feathermoss, lichen, and sphagnum micro-sites in a black spruce forest in eastern North America during snow-free periods over three years. The response of soil respiration (Rs-auto) and forest floor photosynthesis (Pff) to environmental factors was determined. The seasonal contributions of scaled up Rs-auto adjusted for spatial representativeness (Rs-adj) and Pff (Pff-eco) relative to that of total ecosystem respiration (Re) and photosynthesis (Peco), respectively, were also quantified. Shallow (5 cm) soil temperature explained 67-86% of the variation in Rs-auto for all ground cover types, while deeper (50 and 100 cm) soil temperatures were related to Rs-auto only for the feathermoss micro-sites. Base respiration was consistently lower under feathermoss, intermediate under sphagnum, and higher under lichen during all three years. The Rs-adj/Re ratio increased from spring through autumn and ranged from 0.85 to 0.87 annually for the snow-free period. The Rs-adj/Re ratio was negatively correlated with the difference between air and shallow soil temperature and this correlation was more pronounced in autumn than summer and spring. Maximum photosynthetic capacity of the forest floor (Pff-max) saturated at low irradiance levels (~200 μmol m-2 s-1) and decreased with increasing air temperature and vapor pressure deficit for all three ground cover types, suggesting that Pff was more limited by desiccation than by light availability. Pff-max was lowest for sphagnum, intermediate for feathermoss, and highest for lichen for two of the three years. Pff normalized for light peaked at air temperatures of 5-8°C, suggesting that this is the optimal temperature range for Pff. The Pff-eco/Peco ratio varied from 13 to 24% over the snow-free period and reached a minimum in mid-summer when both air temperature and Peco were at their maximum. On an annual basis, Pff-eco accounted for 17

  15. Monitoring larval populations of the Douglas-fir tussock moth and the western spruce budworm on permanent plots: sampling methods and statistical properties of data

    Science.gov (United States)

    A.R. Mason; H.G. Paul

    1994-01-01

    Procedures for monitoring larval populations of the Douglas-fir tussock moth and the western spruce budworm are recommended based on many years experience in sampling these species in eastern Oregon and Washington. It is shown that statistically reliable estimates of larval density can be made for a population by sampling host trees in a series of permanent plots in a...

  16. Black spruce growth forms as a record of a changing winter environment at treeline, Quebec, Canada

    International Nuclear Information System (INIS)

    Lavoie, C.; Payette, S.

    1992-01-01

    The environmental conditions prevailing at treeline in subarctic Quebec have been reconstructed over the past 400 yr through a comparative analysis of tree rings and growth forms of black spruce (Picea mariana [Mill.] B.S.P.). Because black spruce growth forms are closely associated with the winter environment, they are a direct response to conditions of low temperature and windblown snow abrasion affecting living tissues at the snow-air interface. The age structure of supranival shoot populations was closely associated with periods of higher stem survival in winter most likely under snowier and windless conditions. Spruce growth on slopes and in the valley revealed periods of low tree-ring growth between 1601 and 1663 and between 1700 and 1904, respectively. A long-lasting period of low radial growth 1697 and 1939 prevailed in the hilltop site. During the 20th century, spruce height increased from 0.8 to 1.6 m on slopes and in the valley, while the basal level of abrasion from windblown snow increased from 0.1 to 0.5 m, suggesting an increasing trend towards warmer and snowier conditions. Abraded spruces growing during the Little Ice Age (1570-1880) were replaced by symmetrical trees during the 20th century. Supranival skirted and whorled spruces which dominated on the hilltop site during the 16th century reverted to infranival cushion and mat growth forms during the Little Ice Age. These stunted spruces were unable to recover during the recent warming because of their inability to catch enough drifting snow to allow vertical growth

  17. Nesting ecology of boreal forest birds following a massive outbreak of spruce beetles

    Science.gov (United States)

    Matsuoka, S.M.; Handel, C.M.

    2007-01-01

    We studied breeding dark-eyed juncos (Junco hyemalis), yellow-rumped warblers (Dendroica coronata), and spruce-nesting birds from 1997 to 1998 among forests with different levels of spruce (Picea spp.) mortality following an outbreak of spruce beetles (Dendroctonus rufipennis) in Alaska, USA. We identified species using live and beetle-killed spruce for nest sites and monitored nests to determine how the outbreak influenced avian habitat selection and reproduction. We tested predictions that 1) nesting success of ground-nesting juncos would increase with spruce mortality due to proliferation of understory vegetation available to conceal nests from predators, 2) nesting success of canopy-nesting warblers would decrease with spruce mortality due to fewer live spruce in which to conceal nests, and 3) both species would alter nest-site selection in response to disturbance. Juncos did not benefit from changes in understory vegetation; nesting success in highly disturbed stands (46%) was comparable to that in undisturbed habitats throughout their range. In stands with low spruce mortality, nesting success of juncos was low (5%) and corresponded with high densities of red squirrels (Tamiasciurus hudsonicus). Yellow-rumped warblers nested exclusively in spruce, but success did not vary with spruce mortality. As disturbance increased, nesting warblers switched from selecting forest patches with high densities of live white spruce (Picea glauca) to patches with beetle-killed spruce. Warblers also placed nests in large-diameter live or beetle-killed spruce, depending on which was more abundant in the stand, with no differences in nesting success. Five of the 12 other species of spruce-nesting birds also used beetle-killed spruce as nest sites. Because beetle-killed spruce can remain standing for >50 years, even highly disturbed stands provide an important breeding resource for boreal forest birds. We recommend that boreal forest managers preserve uncut blocks of infested

  18. Predictive Modeling of Black Spruce (Picea mariana (Mill. B.S.P. Wood Density Using Stand Structure Variables Derived from Airborne LiDAR Data in Boreal Forests of Ontario

    Directory of Open Access Journals (Sweden)

    Bharat Pokharel

    2016-12-01

    Full Text Available Our objective was to model the average wood density in black spruce trees in representative stands across a boreal forest landscape based on relationships with predictor variables extracted from airborne light detection and ranging (LiDAR point cloud data. Increment core samples were collected from dominant or co-dominant black spruce trees in a network of 400 m2 plots distributed among forest stands representing the full range of species composition and stand development across a 1,231,707 ha forest management unit in northeastern Ontario, Canada. Wood quality data were generated from optical microscopy, image analysis, X-ray densitometry and diffractometry as employed in SilviScan™. Each increment core was associated with a set of field measurements at the plot level as well as a suite of LiDAR-derived variables calculated on a 20 × 20 m raster from a wall-to-wall coverage at a resolution of ~1 point m−2. We used a multiple linear regression approach to identify important predictor variables and describe relationships between stand structure and wood density for average black spruce trees in the stands we observed. A hierarchical classification model was then fitted using random forests to make spatial predictions of mean wood density for average trees in black spruce stands. The model explained 39 percent of the variance in the response variable, with an estimated root mean square error of 38.8 (kg·m−3. Among the predictor variables, P20 (second decile LiDAR height in m and quadratic mean diameter were most important. Other predictors describing canopy depth and cover were of secondary importance and differed according to the modeling approach. LiDAR-derived variables appear to capture differences in stand structure that reflect different constraints on growth rates, determining the proportion of thin-walled earlywood cells in black spruce stems, and ultimately influencing the pattern of variation in important wood quality attributes

  19. SPRUCE experiment data infrastructure

    Science.gov (United States)

    Krassovski, M.; Hanson, P. J.; Boden, T.; Riggs, J.; Nettles, W. R.; Hook, L. A.

    2013-12-01

    The Carbon Dioxide Information Analysis Center (CDIAC) at Oak Ridge National Laboratory (ORNL), USA has provided scientific data management support for the US Department of Energy and international climate change science since 1982. Among the many data activities CDIAC performs are design and implementation of the data systems. One current example is the data system and network for SPRUCE experiment. The SPRUCE experiment (http://mnspruce.ornl.gov) is the primary component of the Terrestrial Ecosystem Science Scientific Focus Area of ORNL's Climate Change Program, focused on terrestrial ecosystems and the mechanisms that underlie their responses to climatic change. The experimental work is to be conducted in a bog forest in northern Minnesota, 40 km north of Grand Rapids, in the USDA Forest Service Marcell Experimental Forest (MEF). The site is located at the southern margin of the boreal peatland forest. Experimental work in the 8.1-ha S1 bog will be a climate change manipulation focusing on the combined responses to multiple levels of warming at ambient or elevated CO2 (eCO2) levels. The experiment provides a platform for testing mechanisms controlling the vulnerability of organisms, biogeochemical processes and ecosystems to climatic change (e.g., thresholds for organism decline or mortality, limitations to regeneration, biogeochemical limitations to productivity, the cycling and release of CO2 and CH4 to the atmosphere). The manipulation will evaluate the response of the existing biological communities to a range of warming levels from ambient to +9°C, provided via large, modified open-top chambers. The ambient and +9°C warming treatments will also be conducted at eCO2 (in the range of 800 to 900 ppm). Both direct and indirect effects of these experimental perturbations will be analyzed to develop and refine models needed for full Earth system analyses. SPRUCE provides wide range continuous and discrete measurements. To successfully manage SPRUCE data flow

  20. Long-term landscape changes in a subalpine spruce-fir forest in central Utah, USA

    Directory of Open Access Journals (Sweden)

    Jesse L. Morris1

    2015-12-01

    promoted a legacy of simplified stand structure and composition such that, when climate became favorable for accelerated beetle population growth, the result was a landscape-scale spruce beetle outbreak. The lasting impacts of settlement-era landscape history from the Wasatch Plateau, UT may be relevant for other areas of western North America and Europe where sufficient host carrying capacity is important in managing for resistance and resilience to outbreaks. Keywords: Spruce beetle, Climate change, Dendroctonus, Engelmann spruce, Logging, Western North America, Wildfire

  1. Distribution of Lepidopteran Larvae on Norway Spruce: Effects of Slope and Crown Aspect.

    Science.gov (United States)

    Kulfan, Ján; Dvořáčková, Katarína; Zach, Peter; Parák, Michal; Svitok, Marek

    2016-04-01

    Lepidoptera associated with Norway spruce, Picea abies (L.) Karsten, play important roles in ecosystem processes, acting as plant pests, prey for predators, and hosts for parasites and parasitoids. Their distribution patterns in spruce crowns and forests are only poorly understood. We examined how slope and crown aspect affect the occurrence and abundance of moth larvae on solitary spruce trees in a montane region in Central Europe. Moth larvae were collected from southern and northern crowns of trees growing on south- and north-facing slopes (four treatments) using emergence boxes at the end of winter and by the beating method during the growing season. Species responses to slope and crown aspect were not uniform. Treatment effects on moth larvae were stronger in the winter than during the growing season. In winter, the abundance of bud-boring larvae was significantly higher in northern than in southern crowns regardless of the slope aspect, while both slope and aspect had marginally significant effects on abundance of miners. During the growing season, the occurrence of free-living larvae was similar among treatments. Emergence boxes and beating spruce branches are complementary techniques providing valuable insights into the assemblage structure of moth larvae on Norway spruce. Due to the uneven distribution of larvae detected in this study, we recommend adoption of a protocol that explicitly includes sampling of trees from contrasting slopes and branches from contrasting crown aspect in all seasons. © The Authors 2016. Published by Oxford University Press on behalf of Entomological Society of America. All rights reserved. For Permissions, please email: journals.permissions@oup.com.

  2. Managing Understory Vegetation for Maintaining Productivity in Black Spruce Forests: A Synthesis within a Multi-Scale Research Model

    Directory of Open Access Journals (Sweden)

    Gilles Joanisse

    2013-07-01

    Full Text Available Sustainable management of boreal ecosystems involves the establishment of vigorous tree regeneration after harvest. However, two groups of understory plants influence regeneration success in eastern boreal Canada. Ericaceous shrubs are recognized to rapidly dominate susceptible boreal sites after harvest. Such dominance reduces recruitment and causes stagnant conifer growth, lasting decades on some sites. Additionally, peat accumulation due to Sphagnum growth after harvest forces the roots of regenerating conifers out of the relatively nutrient rich and warm mineral soil into the relatively nutrient poor and cool organic layer, with drastic effects on growth. Shifts from once productive black spruce forests to ericaceous heaths or paludified forests affect forest productivity and biodiversity. Under natural disturbance dynamics, fires severe enough to substantially reduce the organic layer thickness and affect ground cover species are required to establish a productive regeneration layer on such sites. We succinctly review how understory vegetation influences black spruce ecosystem dynamics in eastern boreal Canada, and present a multi-scale research model to understand, limit the loss and restore productive and diverse ecosystems in this region. Our model integrates knowledge of plant-level mechanisms in the development of silvicultural tools to sustain productivity. Fundamental knowledge is integrated at stand, landscape, regional and provincial levels to understand the distribution and dynamics of ericaceous shrubs and paludification processes and to support tactical and strategic forest management. The model can be adapted and applied to other natural resource management problems, in other biomes.

  3. Long-term trends in radial growth of Siberian spruce and Scots pine in Komi Republic (northwestern Russia)

    Energy Technology Data Exchange (ETDEWEB)

    Lopatin, E. (Univ. of Joensuu (Finland)); Kolstroem, T. (Russian Academy of Sciences, Syktyvkar (Russian Federation)); Spiecker, H. (Univ. of Freiburg (Germany))

    2008-07-01

    Komi is situated on the eastern boundary of the European part of Russia, in the boreal region where large areas of natural forest still exist. Using radial growth measurements it was possible to attain positive long-term trends of growth in Scots pine (Pinus sylvestris) and Siberian spruce (Picea obovata) in the Komi Republic. Increases in the radial growth of Siberian spruce in the forest-tundra were 134% and in the northern taiga zone 35% over successive 50-year periods from 1901 to 1950 and from 1951 to 2000. Respectively, in the middle taiga zone a 76% increase in radial growth was found (over 100 years), whilst in the southern taiga zone the changes were not statistically significant. The increase in radial growth of Scots pine in the northern taiga zone was 32%. In the middle taiga zone the radial growth increase in Scots pine was 55% and in the southern taiga zone the changes were not statistically significant. The long-term growth trends of Komi were compared with those in other parts of Europe. (orig.)

  4. Predicting Effects of Climate Change on Habitat Suitability of Red Spruce (Picea rubens Sarg. in the Southern Appalachian Mountains of the USA: Understanding Complex Systems Mechanisms through Modeling

    Directory of Open Access Journals (Sweden)

    Kyung Ah Koo

    2015-04-01

    Full Text Available Alpine, subalpine and boreal tree species, of low genetic diversity and adapted to low optimal temperatures, are vulnerable to the warming effects of global climate change. The accurate prediction of these species’ distributions in response to climate change is critical for effective planning and management. The goal of this research is to predict climate change effects on the distribution of red spruce (Picea rubens Sarg. in the Great Smoky Mountains National Park (GSMNP, eastern USA. Climate change is, however, conflated with other environmental factors, making its assessment a complex systems problem in which indirect effects are significant in causality. Predictions were made by linking a tree growth simulation model, red spruce growth model (ARIM.SIM, to a GIS spatial model, red spruce habitat model (ARIM.HAB. ARIM.SIM quantifies direct and indirect interactions between red spruce and its growth factors, revealing the latter to be dominant. ARIM.HAB spatially distributes the ARIM.SIM simulations under the assumption that greater growth reflects higher probabilities of presence. ARIM.HAB predicts the future habitat suitability of red spruce based on growth predictions of ARIM.SIM under climate change and three air pollution scenarios: 10% increase, no change and 10% decrease. Results show that suitable habitats shrink most when air pollution increases. Higher temperatures cause losses of most low-elevation habitats. Increased precipitation and air pollution produce acid rain, which causes loss of both low- and high-elevation habitats. The general prediction is that climate change will cause contraction of red spruce habitats at both lower and higher elevations in GSMNP, and the effects will be exacerbated by increased air pollution. These predictions provide valuable information for understanding potential impacts of global climate change on the spatiotemporal distribution of red spruce habitats in GSMNP.

  5. Nutrient imbalance in Norway spruce

    International Nuclear Information System (INIS)

    Thelin, Gunnar

    2000-11-01

    The studies presented in my thesis indicate that growing Norway spruce in monoculture does not constitute sustainable forest management in a high N and S deposition environment, such as in southern Sweden. The combination of N-induced high growth rates and leaching due to soil acidification causes soil reserves of nutrients to decrease. This will increase the risk of nutrient imbalance within the trees when nutrient demands are not met. The development of nutrient imbalance in Scania, southern Sweden, was shown as negative trends in needle and soil nutrient status from the mid-80s to the present in Norway spruce and Scots pine stands. This imbalance appears to be connected to high levels of N and S deposition. Clear negative effects on tree vitality were found when using a new branch development method. Today, growth and vitality seems to be limited by K, rather than N, in spruce stands older than 40 years. However, younger stands appear to be able to absorb the deposited N without negative effects on growth and vitality. When investigating effects of nutrient stress on tree vitality, indicators such as branch length and shoot multiplication rate, which include effects accumulated over several years, are suitable. Countermeasures are needed in order to maintain the forest production at a high level. Positive effects on tree nutrient status after vitality fertilization (N-free fertilization) was shown in two micronutrient deficient stands in south-central Sweden. In addition, tree vitality was positively affected after the application of a site-adapted fertilizer to the canopy. Site-adaption of fertilizers will most likely improve the possibilities of a positive response on tree growth and vitality in declining stands. In a survey of Norway spruce in mixtures with beech, birch, or oak compared to monocultures it was shown that spruce nutrient status was higher in mixtures with deciduous species than in monocultures. By using mixed-species stands the need for

  6. Nutrient imbalance in Norway spruce

    Energy Technology Data Exchange (ETDEWEB)

    Thelin, Gunnar

    2000-11-01

    The studies presented in my thesis indicate that growing Norway spruce in monoculture does not constitute sustainable forest management in a high N and S deposition environment, such as in southern Sweden. The combination of N-induced high growth rates and leaching due to soil acidification causes soil reserves of nutrients to decrease. This will increase the risk of nutrient imbalance within the trees when nutrient demands are not met. The development of nutrient imbalance in Scania, southern Sweden, was shown as negative trends in needle and soil nutrient status from the mid-80s to the present in Norway spruce and Scots pine stands. This imbalance appears to be connected to high levels of N and S deposition. Clear negative effects on tree vitality were found when using a new branch development method. Today, growth and vitality seems to be limited by K, rather than N, in spruce stands older than 40 years. However, younger stands appear to be able to absorb the deposited N without negative effects on growth and vitality. When investigating effects of nutrient stress on tree vitality, indicators such as branch length and shoot multiplication rate, which include effects accumulated over several years, are suitable. Countermeasures are needed in order to maintain the forest production at a high level. Positive effects on tree nutrient status after vitality fertilization (N-free fertilization) was shown in two micronutrient deficient stands in south-central Sweden. In addition, tree vitality was positively affected after the application of a site-adapted fertilizer to the canopy. Site-adaption of fertilizers will most likely improve the possibilities of a positive response on tree growth and vitality in declining stands. In a survey of Norway spruce in mixtures with beech, birch, or oak compared to monocultures it was shown that spruce nutrient status was higher in mixtures with deciduous species than in monocultures. By using mixed-species stands the need for

  7. Labeling Feral Spruce Budworm (Lepidoptera: Tortricidae) Populations With Rubidium.

    Science.gov (United States)

    MacKinnon, Wayne; Eveleigh, Eldon; Silk, Peter; Forbes, Glen

    2016-04-01

    Rubidium (Rb) is a trace element that occurs naturally in low concentrations and is easily absorbed by plants, making it a useful tool for labeling insect defoliators, such as spruce budworm (Choristoneura fumiferana Clemens). Balsam fir trees (Abies balsamea (L.) Miller) injected with either 8 or 16 g per tree of rubidium chloride (RbCl) showed quick uptake and distribution throughout the crown, with no negative effects on tree shoot growth or spruce budworm survival and development. Adult spruce budworm that fed as larvae on trees injected with RbCl were clearly labeled, with significantly higher Rb concentrations than the background levels found in adults that fed as larvae on control trees. Rb concentrations in feral spruce budworm adults for both the 8 g (9 µg/g) and 16 g (25 µg/g) per tree treatments were at least five times lower than those in laboratory-reared adults on 1,000 µg/g RbCl diet (125 µg/g); survival, development, pupal weight, sex ratio, and mating status of spruce budworm were not adversely affected by Rb treatment. Egg masses laid by feral females that fed as larvae on Rb-labeled trees were also labeled with Rb. Injecting trees with RbCl is a viable technique for labeling feral spruce budworm populations to help distinguish local populations from immigrants to better evaluate the success of early intervention strategies such as mating disruption. © Crown copyright 2016.

  8. Composition and Elevation of Spruce Forests Affect Susceptibility to Bark Beetle Attacks: Implications for Forest Management

    Directory of Open Access Journals (Sweden)

    Massimo Faccoli

    2014-01-01

    Full Text Available The spruce bark beetle, Ips typographus (L. (Coleoptera: Curculionidae, Scolytinae, is one of the most destructive insects infesting spruce forests in Europe. Data concerning infestations of I. typographus occurring over the last 19 years (1994–2012 on the Southern Alps were analyzed in seven spruce forest types: (1 pure spruce plantations; (2 pure spruce reforestations; (3 pure spruce mountain forests; (4 pure spruce alpine forests; (5 spruce-conifer mixed forests; (6 spruce-broadleaf mixed forests; and (7 spruce-conifer-broadleaf mixed forests. The collected data included the amount of I. typographus damage and the location and composition of the infested forests. The results indicate that different forest types are differently susceptible to I. typographus. Plantations, reforestations and mountain spruce forests show mean damage and mean number of infestations higher than other forest types. Within pure spruce forests, alpine forests growing at high elevations (>1300 m suffer low damage. Furthermore, the mean number of infestation spots recorded annually in the different spruce forest types is negatively correlated with a Naturality Index value. The results suggest that forest composition and elevation are the main factors driving the risk of I. typographus damage. A new management strategy for some spruce forest types is needed, with a progressive reduction of pure spruce forests at low altitude and an increase of broadleaf composition.

  9. Growth strategy of Norway spruce under air elevated [CO2

    Science.gov (United States)

    Pokorny, R.; Urban, O.; Holisova, P.; Sprtova, M.; Sigut, L.; Slipkova, R.

    2012-04-01

    Plants will respond to globally increasing atmospheric CO2 concentration ([CO2]) by acclimation or adaptation at physiological and morphological levels. Considering the temporal onset, physiological responses may be categorized as short-term and morphological ones as long-term responses. The degree of plant growth responses, including cell division and cell expansion, is highly variable. It depends mainly on the specie's genetic predisposition, environment, mineral nutrition status, duration of CO2 enrichment, and/or synergetic effects of other stresses. Elevated [CO2] causes changes in tissue anatomy, quantity, size, shape and spatial orientation and can result in altered sink strength. Since, there are many experimental facilities for the investigation of elevated [CO2] effects on trees: i) closed systems or open top chambers (OTCs), ii) semi-open systems (for example glass domes with adjustable lamella windows - DAWs), and iii) free-air [CO2] enrichments (FACE); the results are still unsatisfactory due to: i) relatively short-term duration of experiments, ii) cultivation of young plants with different growth strategy comparing to old ones, iii) plant cultivation under artificial soil and weather conditions, and iv) in non-representative stand structure. In this contribution we are discussing the physiological and morphological responses of Norway spruce trees cultivated in DAWs during eight consecutive growing seasons in the context with other results from Norway spruce cultivation under air-elevated [CO2] conditions. On the level of physiological responses, we discuss the changes in the rate of CO2 assimilation, assimilation capacity, photorespiration, dark respiration, stomatal conductance, water potential and transpiration, and the sensitivity of these physiological processes to temperature. On the level of morphological responses, we discuss the changes in bud and growth phenology, needle and shoot morphology, architecture of crown and root system, wood

  10. The vegetation of spruce forests in the Pinega State Reserve

    Directory of Open Access Journals (Sweden)

    Sergey Yu. Popov

    2016-08-01

    Full Text Available The Pinega Natural State Reserve is located in the Arkhangelsk Province in the northern taiga subzone. Spruce forests represent the dominant vegetation formation of its territory. The vegetation of this forest is classified, based on 192 phytosociological descriptions. It reveals 12 associations, which represent 7 groups of associations. Detailed characteristics of these syntaxa, including analysis of their biodiversity, are provided. The revealed syntaxa differ both in species composition and environmental conditions: moisture, nutrition, nitrogen availability and acidity. Most poor conditions in terms of mineral nutrition occupy sphagnous spruce forests and bilberry-dominated spruce forests, while under the richest conditions varioherbaceous, humidoherbaceous and nemoral-herbaceous spruce forests occur. The Pinega Reserve is the only locality, where the Piceetum rubo saxatilis-vacciniosum association occurs in the northern taiga subzone.

  11. Some observations on age relationships in spruce-fir regeneration

    Science.gov (United States)

    Barton M. Blum

    1973-01-01

    Measurement of the ages of seedlings of balsam fir (Abies balsamea (L) Mill.), red spruce (Picea rubens Sarg.), and white spruce (Picea glauca (Moench) Voss) 15 years after the first harvest of a two-cut shelterwood operation revealed that very few potential crop-tree seedlings in the sample occurred as advance...

  12. Spruce needles used as radioecological biotracers; Fichtennadeln als radiooekologische Bioindikatoren

    Energy Technology Data Exchange (ETDEWEB)

    Seidel, C.; Gruber, V.; Baumgartner, A. [BOKU - Univ. fuer Bodenkultur Wien (Austria). LLC-Labor Arsenal; Idinger, J. [Technische Univ. Wien (Austria). Atominst.; Fuerst, A. [BFW - Bundesforschungs- und Ausbildungszentrum fuer Wald, Naturgefahren und Landschaft, Wien (Austria). Inst. fuer Waldschutz, Pflanzenanalyse; Maringer, F.J. [BOKU - Univ. fuer Bodenkultur Wien (Austria). LLC-Labor Arsenal; BEV - Bundesamt fuer Eich- und Vermessungswesen, Wien (Austria)

    2009-07-01

    In a two years project spruce needle samples of the Austrian Bioindicator Grid were analysed by gamma-ray spectrometry to investigate the spatial and temporal distribution of radionuclides in spruce needles of the last 25 years with the main focus on the radioactive contamination before and after the Chernobyl fallout 1986. More than 600 spruce needle samples at selected locations of the Bioindicator Grid were analysed for different natural and anthropogenic radionuclides: {sup 137}Cs, {sup 40}K, {sup 210}Pb, {sup 226}Ra, {sup 228}Ra, {sup 238}U. Additionally, soil samples were taken at selected sites to study the soil-to-plant transfer. This radioecological evaluation is an important part of an existing environmental surveillance programme in Upper Austria in order to gain basic information on the impact of environmental changes on the radioecological behaviour of spruce trees. (orig.)

  13. Negative impacts of high temperatures on growth of black spruce forests intensify with the anticipated climate warming.

    Science.gov (United States)

    Girardin, Martin P; Hogg, Edward H; Bernier, Pierre Y; Kurz, Werner A; Guo, Xiao Jing; Cyr, Guillaume

    2016-02-01

    An increasing number of studies conclude that water limitations and heat stress may hinder the capacity of black spruce (Picea mariana (Mill.) B.S.P.) trees, a dominant species of Canada's boreal forests, to grow and assimilate atmospheric carbon. However, there is currently no scientific consensus on the future of these forests over the next century in the context of widespread climate warming. The large spatial extent of black spruce forests across the Canadian boreal forest and associated variability in climate, demography, and site conditions pose challenges for projecting future climate change responses. Here we provide an evaluation of the impacts of climate warming and drying, as well as increasing [CO2 ], on the aboveground productivity of black spruce forests across Canada south of 60°N for the period 1971 to 2100. We use a new extensive network of tree-ring data obtained from Canada's National Forest Inventory, spatially explicit simulations of net primary productivity (NPP) and its drivers, and multivariate statistical modeling. We found that soil water availability is a significant driver of black spruce interannual variability in productivity across broad areas of the western to eastern Canadian boreal forest. Interannual variability in productivity was also found to be driven by autotrophic respiration in the warmest regions. In most regions, the impacts of soil water availability and respiration on interannual variability in productivity occurred during the phase of carbohydrate accumulation the year preceding tree-ring formation. Results from projections suggest an increase in the importance of soil water availability and respiration as limiting factors on NPP over the next century due to warming, but this response may vary to the extent that other factors such as carbon dioxide fertilization, and respiration acclimation to high temperature, contribute to dampening these limitations. © 2015 Her Majesty the Queen in Right of Canada. Reproduced with

  14. Does the azimuth orientation of Norway spruce (Picea abies/L./Karst.) branches within sunlit crown part influence the heterogeneity of biochemical, structural and spectral characteristics of needles?

    Czech Academy of Sciences Publication Activity Database

    Lhotáková, Z.; Albrechtová, J.; Malenovský, Zbyněk; Rock, B. N.; Polák, T.; Cudlín, Pavel

    2007-01-01

    Roč. 59, č. 3 (2007), s. 283-297 ISSN 0098-8472 R&D Projects: GA MŠk ME 658 Institutional research plan: CEZ:AV0Z60870520 Keywords : branch azimuth orientation * Norway spruce * chlorophyll * phenolic compounds * needle structure * spectral reflectance indices * remote sensing Subject RIV: EF - Botanics Impact factor: 1.810, year: 2007

  15. Concentrations of Ca and Mg in early stages of sapwood decay in red spruce, eastern hemlock, red maple, and paper birch

    Science.gov (United States)

    Kevin T. Smith; Walter C. Shortle; Jody Jellison; Jon Connolly; Jonathan Schilling

    2007-01-01

    The decay of coarse woody debris is a key component in the formation of forest soil and in the biogeochemical cycles of Ca and Mg. We tracked changes in density and concentration of Ca and Mg in sapwood of red maple (Acer rubrum L.), red spruce (Picea rubens Sarg.), paper birch (Betula papyrifera Marsh.), and...

  16. Two-dimensional wavelet analysis of spruce budworm host basal area in the Border Lakes landscape

    Science.gov (United States)

    Patrick M. James; Brian R. Sturtevant; Phil Townsend; Pete Wolter; Marie-Josee. Fortin

    2011-01-01

    Increases in the extent and severity of spruce budworm (Choristoneura fumiferana Clem.) outbreaks over the last century are thought to be the result of changes in forest structure due to forest management. A corollary of this hypothesis is that manipulations of forest structure and composition can be used to reduce future forest vulnerability....

  17. Decadal Variations in Eastern Canada's Taiga Wood Biomass Production Forced by Ocean-Atmosphere Interactions.

    Science.gov (United States)

    Boucher, Etienne; Nicault, Antoine; Arseneault, Dominique; Bégin, Yves; Karami, Mehdi Pasha

    2017-05-26

    Across Eastern Canada (EC), taiga forests represent an important carbon reservoir, but the extent to which climate variability affects this ecosystem over decades remains uncertain. Here, we analyze an extensive network of black spruce (Picea mariana Mill.) ring width and wood density measurements and provide new evidence that wood biomass production is influenced by large-scale, internal ocean-atmosphere processes. We show that while black spruce wood biomass production is primarily governed by growing season temperatures, the Atlantic ocean conveys heat from the subtropics and influences the decadal persistence in taiga forests productivity. Indeed, we argue that 20-30 years periodicities in Sea Surface Temperatures (SSTs) as part of the the Atlantic Multi-decadal Oscillation (AMO) directly influence heat transfers to adjacent lands. Winter atmospheric conditions associated with the North Atlantic Oscillation (NAO) might also impact EC's taiga forests, albeit indirectly, through its effect on SSTs and sea ice conditions in surrounding seas. Our work emphasizes that taiga forests would benefit from the combined effects of a warmer atmosphere and stronger ocean-to-land heat transfers, whereas a weakening of these transfers could cancel out, for decades or longer, the positive effects of climate change on Eastern Canada's largest ecosystem.

  18. Connection between the decline of spruce and occurrence of animal pests, especially nematodes

    Energy Technology Data Exchange (ETDEWEB)

    Timans, U.

    1986-12-01

    In various regions of Bavaria, affected by the decline of spruce, attack by insects and especially nematodes was examined on diseased and healthy spruces. A connection between harmful forest insects and the decline of spruce did not become evident, neither over wide areas nor by examination of single trees. Attack by nematodes was examined in soil and wood samples and also in fine feeder roots of diseased and healthy trees. Plant-parasitic nematodes were not found in the wood and in feeder roots. Although root-parasitic nematodes were present in soil samples, their density was too little to account for a direct damage to spruce. They occurred likewise in samples from healthy and diseased trees. Plant-parasitic nematodes can thus be excluded as a possible causal agent for the decline of spruce.

  19. Evaluation of the antiaggregation pheromone, 3-methylcyclohex-2-en-1-one (MCH), to protect live spruce from spruce beetle (Coleoptera: Scolytidae) infestation in sourthern Utah.

    Science.gov (United States)

    Darrell W. Ross; Gary E. Daterman; A. Steven Munson

    2004-01-01

    The spruce beetle, Dendroctonus rufipennis (Kirby), produces the antiaggregation pheromone 3-methylcyclohex-2-en- 1-one (MCH) (Rudinsky et al. 1974). MCH has reduced the numbers of spruce beetles attracted to infested logs and synthetic semiochemical lures or reduced colonization rates throughout the beetles range (Kline

  20. Factors affecting spruce establishment and recruitment near western treeline, Alaska

    Science.gov (United States)

    Miller, A. E.; Sherriff, R.; Wilson, T. L.

    2015-12-01

    Regional warming and increases in tree growth are contributing to increased productivity near the western forest margin in Alaska. The effects of warming on seedling recruitment has received little attention, in spite of forecasted forest expansion near western treeline. Here, we used stand structure and environmental data from white spruce (Picea glauca) stands (n = 95) sampled across a longitudinal gradient to explore factors influencing white spruce growth, establishment and recruitment in southwest Alaska. Using tree-ring chronologies developed from a subset of the plots (n = 30), we estimated establishment dates and basal area increment (BAI) for trees of all age classes across a range of site conditions. We used GLMs (generalized linear models) to explore the relationship between tree growth and temperature in undisturbed, low elevation sites along the gradient, using BAI averaged over the years 1975-2000. In addition, we examined the relationship between growing degree days (GDD) and seedling establishment over the previous three decades. We used total counts of live seedlings, saplings and live and dead trees, representing four cohorts, to evaluate whether geospatial, climate, and measured plot covariates predicted abundance of the different size classes. We hypothesized that the relationship between abundance and longitude would vary by size class, and that this relationship would be mediated by growing season temperature. We found that mean BAI for trees in undisturbed, low elevation sites increased with July maximum temperature, and that the slope of the relationship with temperature changed with longitude (interaction significant with 90% confidence). White spruce establishment was positively associated with longer summers and/or greater heat accumulation, as inferred from GDD. Seedling, sapling and tree abundance were also positively correlated with temperature across the study area. The response to longitude was mixed, with smaller size classes

  1. Release of suppressed red spruce using canopy gap creation—Ecological restoration in the Central Appalachians

    Science.gov (United States)

    Rentch, J.S.; Ford, W. Mark; Schuler, T.S.; Palmer, J.; Diggins, Corinne A.

    2016-01-01

    Red spruce (Picea rubens) and red spruce-northern hardwood mixed stands once covered as much as 300,000 ha in the Central Appalachians, but now comprise no more than 21,000 ha. Recently, interest in restoration of this forest type has increased because red spruce forests provide habitat for a number of rare animal species. Our study reports the results of an understory red spruce release experiment in hardwood-dominated stands that have a small component of understory red spruce. In 2005, 188 target spruce were identified in sample plots at six locations in central West Virginia. We projected a vertical cylinder above the crown of all target spruces, and in 2007, we performed a release treatment whereby overtopping hardwoods were treated with herbicide using a stem injection technique. Release treatments removed 0–10% (Control), 11–50% (Low), 51–89% (Medium), and ≤90% (High) of the basal area of overtopping trees. We also took canopy photographs at the time of each remeasurement in 2007, 2010, and 2013, and compared basal removal treatments and resulting 2010 canopy openness and understory light values. The high treatment level provided significantly greater six-year dbh and height growth than the other treatment levels. Based on these results, we propose that a tree-centered release approach utilizing small canopy gaps that emulate the historical, gap-phase disturbance regime provides a good strategy for red spruce restoration in hardwood forests where overstory spruce are virtually absent, and where red spruce is largely relegated to the understory.

  2. Development of soil water regime under spruce stands

    Directory of Open Access Journals (Sweden)

    Tužinský Ladislav

    2017-06-01

    Full Text Available The aim of this paper is to analyse the water regime of soils under spruce ecosystems in relation to long-lasting humid and drought periods in the growing seasons 1991-2013. The dominant interval humidity in observing growing seasons is semiuvidic interval with soil moisture between hydro-limits maximal capillary capacity (MCC and point of diminished availability (PDA. Gravitationally seepage concentrated from accumulated winter season, water from melting snow and existing atmospheric precipitation occurs in the soil only at the beginning of the growing season. The supplies of soil water are significantly decreasing in the warm climate and precipitant deficient days. The greatest danger from drought threatens Norway spruce during the summer months and it depends on the duration of dry days, water supply at the beginning of the dry days, air temperature and the intensity of evapotranspiration. In the surface layers of the soil, with the maximum occurrence of active roots, the water in semiarid interval area between hydro-limits PDA and wilting point (WP decreases during the summer months. In the culminating phase occurs the drying to moisture state with capillary stationary and the insufficient supply of available water for the plants. Physiological weakening of Norway spruce caused by set of outlay components of the water balance is partially reduced by delivering of water by capillary action from deeper horizons. In extremely dry periods, soil moisture is decreasing also throughout the soil profile (0-100 cm into the bottom third of the variation margin hydro-limits MCC-PDA in the category of capillary less moving and for plants of low supply of usable water (60-90 mm. The issue of deteriorated health state of spruce ecosystems is considered to be actual. Changes and developments of hydropedological conditions which interfere the mountain forests represent the increasing danger of the drought for the spruce.

  3. The structural style of foot wall shortcuts along the eastern foothills of the Colombian eastern cordillera. Differences with other inversion related structures

    International Nuclear Information System (INIS)

    Mora, Andres; Parra, Mauricio

    2008-01-01

    For the first time we show geological evidence of unambiguously documented foot wall shortcuts adjacent to the trace of inverted master normal faults, in the Eastern Cordillera of Colombia. The Eastern Cordillera is an orogen whose width and location are traced by a Mesozoic Graben. However, few structures related with the Graben have been documented up to the date. In this study we propose the Ariari-Guatiquia region as a type location for a unique observation of foot wall shortcuts. The master normal faults in the Ariari-Guatiquia region, and documented in this manuscript, were active during the Lower Cretaceous, partially inverted during the Andean orogenesis (since the Oligocene at least) and active still nowadays. In the hanging wall basins of those master normal faults, like the Servita fault, all the Cretaceous syn-rift sequence has been deposited and maximum paleo-temperatures in the lowermost Cretaceous rocks are higher than those for the Zircon FT partial annealing zone (250 Celsius degrade; 23,15 K). In contraction, the inverted master normal faults are high angle basement involved features that generated the main topographic contrast and exposing Lower Cretaceous units or older. In contrast, in the adjacent foot wall shortcuts only part of the syn-rift Lower Cretaceous sequence was deposited or more commonly was not deposited at all. Maximum paleo-temperatures reached by the basal Cretaceous units exposed in the hanging wall blocks of the foot wall shortcuts are always less than those of the Zircon FT partial annealing zone (250 Celsius degrade; 23,15 K). Finally we use AFT data to document that the foot wall shortcuts originated during the Late Miocene and later as shallowly dipping faults generating low elevation hanging wall areas. All the described features are present in the Ariari-Guatiquia region. However, northwards and along strike in the Eastern foothills there is a lot of partially analogue scenarios with respect to those described in the

  4. Preservation of beech and spruce wood by allyl alcohol-based copolymers

    International Nuclear Information System (INIS)

    Solpan, Dilek; Gueven, Olgun

    1999-01-01

    Allyl alcohol (AA), acrylonitrile (AN), methyl methacrylate (MMA), monomers and monomer mixtures AA+AN, AA+MMA were used to conserve and consolidate Beech and Spruce. After impregnation, copolymerisation and polymerisation were accomplished by gamma irradiation. The fine structure of wood+polymer(copolymer) composites was investigated by Scanning Electron Microscopy (SEM). It was observed that copolymer obtained from AA+MMA monomer mixture showed the optimum compatibility. The compressional strength and Brinell Hardness Numbers determined for untreated and treated wood samples indicated that the mechanical strength of wood+copolymer composites was increased. It was found that the mechanical strength of the wood samples containing the AA+MMA copolymer was higher than the others. In the presence of P(AA/MMA), at highest conversion, the compressive strength perpendicular to the fibres in Beech and Spruce increased approximately 100 times. The water uptake capacity of wood+copolymer composites was observed to decrease by more than 50% relative to the original samples, and biodegradation did not take place

  5. Biomass of Sacrificed Spruce/Aspen (SNF)

    Data.gov (United States)

    National Aeronautics and Space Administration — Dimension analysis (diameter at breast high, tree height, depth of crown), estimated leaf area, and total aboveground biomass for sacrificed spruce and aspens in...

  6. Soil structure interactions of eastern U.S. type earthquakes

    International Nuclear Information System (INIS)

    Chang Chen; Serhan, S.

    1991-01-01

    Two types of earthquakes have occurred in the eastern US in the past. One of them was the infrequent major events such as the 1811-1812 New Madrid Earthquakes, or the 1886 Charleston Earthquake. The other type was the frequent shallow earthquakes with high frequency, short duration and high accelerations. Two eastern US nuclear power plants, V.C Summer and Perry, went through extensive licensing effort to obtain fuel load licenses after this type of earthquake was recorded on sites and exceeded the design bases beyond 10 hertz region. This paper discusses the soil-structure interactions of the latter type of earthquakes

  7. Interspecific variation in resistance of two host tree species to spruce budworm

    Science.gov (United States)

    Fuentealba, Alvaro; Bauce, Éric

    2016-01-01

    Woody plants regularly sustain biomass losses to herbivorous insects. Consequently, they have developed various resistance mechanisms to cope with insect attack. However, these mechanisms of defense and how they are affected by resource availability are not well understood. The present study aimed at evaluating and comparing the natural resistance (antibiosis and tolerance) of balsam fir (Abies balsamea [L.] Mill.) and white spruce (Picea glauca [Moench) Voss] to spruce budworm, Choristoneura fumiferana (Clem.), and how drainage site quality as a component of resource availability affects the expression of resistance over time (6 years). Our results showed that there are differences in natural resistance between the two tree species to spruce budworm, but it was not significantly affected by drainage quality. Balsam fir exhibited higher foliar toxic secondary compounds concentrations than white spruce in all drainage classes, resulting in lower male pupal mass, survival and longer male developmental time. This, however, did not prevent spruce budworm from consuming more foliage in balsam fir than in white spruce. This response suggests that either natural levels of measured secondary compounds do not provide sufficient toxicity to reduce defoliation, or spruce budworm has developed compensatory mechanisms, which allow it to utilize food resources more efficiently or minimize the toxic effects that are produced by its host's defensive compounds. Larvae exhibited lower pupal mass and higher mortality in rapidly drained and subhygric sites. Drainage class also affected the amount of foliage destroyed but its impact varied over the years and was probably influenced by climatic variables. These results demonstrate the complexity of predicting the effect of resource availability on tree defenses, especially when other confounding environmental factors can affect tree resource allocation and utilization.

  8. Ethanol exposure can inhibit red spruce ( Picea rubens ) seed germination

    Science.gov (United States)

    John R. Butnor; Brittany M. Verrico; Victor Vankus; Stephen R. Keller

    2018-01-01

    Flotation of seeds in solvents is a common means of separating unfilled and filled seeds. While a few protocols for processing red spruce (Picea rubens) seeds recommend ethanol flotation, delayed and reduced germination have been reported. We conducted an ethanol bioassay on seeds previously stored at -20°C to quantify the concentration required to separate red spruce...

  9. High-Titer Methane from Organosolv-Pretreated Spruce and Birch

    Directory of Open Access Journals (Sweden)

    Leonidas Matsakas

    2017-02-01

    Full Text Available The negative impact of fossil fuels and the increased demand for renewable energy sources has led to the use of novel raw material sources. Lignocellulosic biomass could serve as a possible raw material for anaerobic digestion and production of biogas. This work is aimed at using forest biomass, both softwood (spruce and hardwood (birch, as a raw material for anaerobic digestion. We examined the effect of different operational conditions for the organosolv pretreatment (ethanol content, duration of treatment, and addition of acid catalyst on the methane yield. In addition, we investigated the effect of addition of cellulolytic enzymes during the digestion. We found that inclusion of an acid catalyst during organosolv pretreatment improved the yields from spruce, but it did not affect the yields from birch. Shorter duration of treatment was advantageous with both materials. Methane yields from spruce were higher with lower ethanol content whereas higher ethanol content was more beneficial for birch. The highest yields obtained were 185 mL CH4/g VS from spruce and 259.9 mL CH4/g VS from birch. Addition of cellulolytic enzymes improved these yields to 266.6 mL CH4/g VS and 284.2 mL CH4/g VS, respectively.

  10. Isoprenoid emission variation of Norway spruce across a European latitudinal transect

    DEFF Research Database (Denmark)

    van Meeningen, Ylva; Wang, Min; Karlsson, Tomas

    2017-01-01

    to the formation and growth of secondary organic aerosols (SOA) in the atmosphere. Isoprenoid emissions were measured from Norway spruce trees at seven different sites, distributed from Ljubljana in Slovenia to Piikkiö in Finland. Four of the sites were part of a network of genetically identical spruce trees...

  11. Comparison of different ground techniques to map leaf area index of Norway spruce forest canopy

    NARCIS (Netherlands)

    Homolova, L.; Malenovsky, Z.; Hanus, J.; Tomaskova, I.; Dvoráková, M.; Pokorny, R.

    2007-01-01

    The leaf area index (LAI) of three monocultures of Norway spruce (Picea abies (L.) Karst), different in age and structure, was measured by means of two indirect optical techniques of LAI field mapping: 1/ plant canopy analyser LAI-2000, and 2/ digital hemispherical photographs (DHP). The supportive

  12. Building Resilience into Sitka Spruce (Picea sitchensis (Bong. Carr. Forests in Scotland in Response to the Threat of Climate Change

    Directory of Open Access Journals (Sweden)

    Andrew D. Cameron

    2015-02-01

    Full Text Available It is expected that a warming climate will have an impact on the future productivity of European spruce forests. In Scotland, Sitka spruce (Picea sitchensis (Bong. Carr. dominates the commercial forestry sector and there is growing pressure to develop alternative management strategies to limit potential economic losses through climate change. This review considers management options to increase the resilience of Sitka spruce dominated forests in Scotland. Given the considerable uncertainty over the potential long-term impacts of climate change, it is recommended that Sitka spruce should continue to be planted where it already grows well. However, new planting and restocking should be established in mixtures where silviculturally practicable, even if no-thin regimes are adopted, to spread future risks of damage. Three potentially compatible species with Sitka spruce are western hemlock (Tsuga heterophylla (Raf. Sarg., grand fir (Abies grandis (Lamb. Lindl. and Douglas fir (Pseudotsuga menziesii (Mirb. Franco and all form natural mixtures in its native range in North America. The predicted windier climate will require a range of management inputs, such as early cutting of extraction racks and early selective thinning, to improve stability. The potential to improve resilience to particularly abiotic damage through transforming even-aged stands into irregular structures and limiting the overall size of the growing stock is discussed.

  13. Crustal structure of the Eastern Alps and their foreland

    DEFF Research Database (Denmark)

    Grad, M.; Brückl, E.; Majdanski, M.

    2009-01-01

    The subject of this paper concerns the seismic modelling of the crustal structure in the transition zone from the Bohemian Massif, across the Molasse basin and the Eastern Alps to the Southern Alps, mainly on the territory of Austria. The CEL10/Alp04 profile crosses the triple point of the European......) are distinct up to 60-90 km offset and are characterized by large variations in apparent velocity and amplitude. The contact between the Molasse basin and the Eastern Alps represents a barrier for seismic waves. Mid-crustal reflections (Pc) are usually recorded at short distance intervals (20-50 km......, was undertaken using a ray-tracing technique. The P-wave velocity in the crystalline upper crust of the Bohemian Massif and Molasse basin is about 6.15 km s-1, which is slightly higher than in the Alpine area (about 6.0 km s-1). Below the northern accretionary wedge of the Eastern Alps low-velocity sediments...

  14. Predicting Effects of Climate Change on Habitat Suitability of Red Spruce (Picea rubens Sarg.) in the Southern Appalachian Mountains of the USA: Understanding Complex Systems Mechanisms through Modeling

    OpenAIRE

    Koo, Kyung; Patten, Bernard; Madden, Marguerite

    2015-01-01

    Alpine, subalpine and boreal tree species, of low genetic diversity and adapted to low optimal temperatures, are vulnerable to the warming effects of global climate change. The accurate prediction of these species’ distributions in response to climate change is critical for effective planning and management. The goal of this research is to predict climate change effects on the distribution of red spruce (Picea rubens Sarg.) in the Great Smoky Mountains National Park (GSMNP), eastern USA. Clim...

  15. Contrasting development of declining and living larch-spruce stands after a disturbance event: A case study from the High Tatra Mts.

    Directory of Open Access Journals (Sweden)

    Šebeň Vladimír

    2015-09-01

    Full Text Available The decline of spruce stands caused by bark beetle outbreaks is a serious economic and ecological problem of forestry in Slovakia. In the preceding period, the decline affected mainly secondary spruce forests. Over the last decade, due to large bark-beetle outbreaks this problem has been observed also in natural spruce forests, even at high elevations. We dealt with this issue in a case study of short-term development of larch-spruce stands in the High Tatras (at a site called Štart. We compared the situation in the stand infested by bark beetles several years after the wind-throw in 2004 with the stand unaffected by bark beetles. We separately analysed the development of the mature (parent stands and the regeneration. The results indicated that forest decline caused by bark beetles significantly depended on the stand structure (mainly tree species composition, which affected the period of stand disintegration. Mortality of spruce trees slowed down biomass accumulation (and thus carbon sequestration in the forest ecosystem. In the new stand, pioneer tree species dominated (in the conditions of the High Tatras it is primarily rowan, although their share in the parent stand was negligible. The results showed different trends in the accumulation of below-ground and above-ground biomass in the declined and living stands. In the first years after the stand decline, rowan accumulated significantly more biomass than the main tree species, i.e. spruce. The reverse situation was under the surviving stand, where spruce trees accumulated more biomass than rowan. The different share of spruce and pioneer tree species, mainly rowan, affected the ratio between fixed (in woody parts of trees and rotating (in foliage carbon in the undergrowth. Forest die-back is a big source of carbon emissions from dead individuals, and the compensation of these losses in the form of carbon sequestration by future stands is a matter of several decades.

  16. TopoGreenland: Lithospheric structure and topography in Central-Eastern Greenland

    Science.gov (United States)

    Thybo, H.; Shulgin, A.; Kraft, H. A.; Vinnik, L. P.

    2017-12-01

    We present models of the seismic structure of the crust and upper mantle in the interior of Greenland based on new seismological data from the TopoGreenland experiment. Until this experiment, all seismic data in Greenland was acquired close to the coast, where the crustal structure is affected by oceanic break-up. The TopoGreenland data acquisition programme in central-eastern Greenland included the first controlled source seismic experiment in interior Greenland and deployment of 24 broadband (BB) onshore stations for 3 years, partly on the ice cap. The 320 km long seismic refraction/wide-angle reflection profile was acquired on the ice cap by a team of six people during two-months in summer of 2011. We present a 2D velocity model of the crust based on tomographic inversion and forward ray tracing modelling of the controlled source data. It shows a decrease of crustal thickness from 47 km below the centre of Greenland in the western to 40 km in its eastern part of the profile. High lower crustal velocities (Vp 6.8 - 7.3 km/s) below central Greenland may result from past collision tectonics or be related to the passage of the Iceland mantle plume. Crustal receiver functions in the surrounding area demonstrate constant structure along the coast and pronounced, relatively sharp variation in crustal thickness around the mountains at the edge of the ice cap. Surprisingly the thickest crust is observed below the lowest topography under the ice cap, whereas the crust is thin below the high mountains at its edge, and thins further below elevated topography out to the coast. Receiver Function interpretation of the mantle and transition zone structure shows a complicated mosaic variation that cannot be correlated to the variation in topography. The origin of the pronounced mountain ranges around the North Atlantic Ocean with average elevation above 1500 m and peak elevations of more than 3.5 km near Scoresby Sund in Eastern Greenland, is unknown. Our new results demonstrate

  17. Developing silvicultural systems based on partial cutting in western hemlock–Sitka spruce stands of southeast Alaska.

    Science.gov (United States)

    Robert L. Deal; J.C. Tappeiner; Paul E. Hennon

    2002-01-01

    The effects of partial cutting on species composition, stand structure and growth, tree size distribution, and tree disease and mortality were evaluated on 73 plots in 18 stands that were harvested 12–96 years ago in southeast Alaska. Partially-cut stands had diverse and highly complex stand structures similar to uncut stands. Sitka spruce was maintained in mixed...

  18. The effect of the environment on the structure, quantity and composition of spruce needle wax

    International Nuclear Information System (INIS)

    Guenthardt-Goerg, M.S.

    1994-01-01

    The tubular structure (10-nonacosanol), as formed in spring on the wax surface of new spruce needles (Picea abies (L.)Karst.), or as regenerated on previous-year needles, becomes gradually fused and flattened in relation to needle exposure, particularly wind and rain. Structural flattening does not necessarily imply changes in wax quantity, composition or lead to changes in needle transpiration or photosynthesis, and was approximately reproduced by bathing excised twigs in water (with pH having little effect). In 4-year-old plants of one clone planted out at a Swiss plateau and alpine sites, changes in wax structure were similar to those found in mature trees. No such changes were found in plants with O 3 , SO 2 , ambient air, charcoal-filtered air, or in plants grown outside the chambers but shielded from rain. Area-related needle wax quantity in mature trees differed between the two sites, but did not differ in young plants under different treatments (fumigation or planted out at the sites). Minor differences in wax composition, however, were found to be related to the ozone dose of the fumigation or the ambient ozone dose at the sites. In each needle wax sample, 68 compounds grouped into 12 constituent classes were quantified. The quantity of the individual substituent classes varied among wax samples from genetically different mature trees at the two sites in a tree-specific way. Variation of these quantities was not larger than among young cloned plants after different treatments. (orig.)

  19. Response of Lutz, Sitka, and white spruce to attack by Dendroctonus rufipennis (Coleoptera: Scolytidae) and blue stain fungi

    Science.gov (United States)

    Richard A. Werner; Barbara L. Illman

    1994-01-01

    Mechanical wounding and wounding plus inoculation with a blue-stain fungus, Leptographium abietinum (Peck), associated with the spruce beetle, Dendroctonus rufipennis (Kirby), caused an induced reaction zone or lesion around the wound sites in Lutz spruce, Picea lutzii Little, Sitka spruce, P. sitchensis (Bong.) Carr., and white spruce, P. glauca (Moench) Voss, in...

  20. Morphogenetic Litter Types of Bog Spruce Forests

    Directory of Open Access Journals (Sweden)

    T. T. Efremova

    2015-02-01

    Full Text Available For the first time the representation of moss litter morphogenetic structure of valley-riverside and streamside spruce forests was determined for the wetland intermountain area of Kuznetsk Alatau. In general, the litter of (green moss-hypnum spruce forest can be characterized as medium thickness (9–17 cm with high storage of organic matter (77–99 t/ha, which differs in neutral environmental conditions pH 6.8–7.0 and high percentage of ash 11–28 %. Formation litter types were identified, which depend on the content of mineral inclusions in organogenic substrate and the degree of its drainage. The differentiation of litter subhorizons was performed, visual diagnostic indicators of fermentative layers were characterized, and additional (indexes to indicate their specificity were developed. Peat- and peaty-fermentative, humified-fermentative and (black mold humus-fermentative layers were selected. Peat- and peaty-fermentative layers are characterized by content of platy peat macroaggregates of coarse vegetable composition, the presence of abundant fungal mycelium and soil animals are the primary decomposers – myriopoda, gastropoda mollusks. Humified-fermentative layers are identified by including the newly formed amorphous humus-like substances, nutty-granular structural parts of humus nature and soil animals’ humificators – enchytraeids and earthworms. (Black mold humus-fermentative layers are diagnosed by indicators with similar humified-fermentative, but differ from them in clay-humus composition of nutty-granular blue-grey parts. The nomenclature and classification of moss litter were developed on the basis of their diagnostic characteristics of fermentative layers – peat, peaty, reduced peaty, (black mold humus-peaty, reduced (black mold humus-peaty. Using the method of discriminant analysis, we revealed that the physical-chemical properties, mainly percentage of ash and decomposition degree of plant substrate, objectively

  1. The timing and nature of Late Quaternary vegetation changes in the northern Great Plains, USA and Canada: a re-assessment of the spruce phase

    Science.gov (United States)

    Yansa, Catherine H.

    2006-02-01

    This paper revises the chronology for the northward migration of Picea glauca (white spruce) across the northern Great Plains, following the recession of the Laurentide Ice Sheet, and reinterprets the species composition and structure of the late-glacial vegetation on the basis of pollen and plant-macrofossil analysis. The timing of spruce migration is based on 26 14C ages obtained from Picea macrofossils. The date for the appearance of white spruce in southern South Dakota, USA, remains unchanged, 12,600 14C yr BP (ca 15,000 cal yr BP), but its arrival in southern Saskatchewan, Canada, by 10,300 14C yr BP (ca 12,100 cal yr BP) is about 1500 years later than previously estimated based on an organic sediment date. Picea glauca thus migrated northwards at an average rate of 0.38 km/ 14C year (0.30 km/calendar year), significantly slower than the previously published rate of 2 km/ 14C year. White spruce trees probably inhabited lake shorelines, whereas prairie, parkland, and boreal plants occupied both lowlands and uplands, forming an open white spruce parkland. This interpretation differs from a previous reconstruction of a boreal-type spruce forest and thus offers another paleoclimatic interpretation. Precipitation was probably low and summer temperatures relatively mild, averaging about 19 °C.

  2. Growth responses of Norway spruce (Picea abies (L.) Karst.) to the climate in the south-eastern part of the Ceskomoravska Upland (Czech Republic)

    Czech Academy of Sciences Publication Activity Database

    Rybníček, M.; Čermák, P.; Kolář, Tomáš; Žid, T.

    2012-01-01

    Roč. 39, č. 2 (2012), s. 149-157 ISSN 1733-8387 R&D Projects: GA MŠk(CZ) ED1.1.00/02.0073; GA MŽP(CZ) SP/2D1/93/07 Institutional support: RVO:67179843 Keywords : Ceskomoravska Upland * Norway spruce * precipitation * temperature * tree ring Subject RIV: DB - Geology ; Mineralogy Impact factor: 1.653, year: 2012

  3. Release of Suppressed Red Spruce Using Canopy Gap Creation--Ecological Restoration in the Central Appalachians

    Science.gov (United States)

    J.S. Rentch; W.M. Ford; Thomas Schuler; Jeff Palmer; C.A. Diggins

    2016-01-01

    Red spruce (Picea rubens) and red spruce-northern hardwood mixed stands once covered as much as 300,000 ha in the Central Appalachians, but now comprise no more than 21,000 ha. Recently, interest in restoration of this forest type has increased because red spruce forests provide habitat for a number of rare animal species. Our study reports the...

  4. Cenozoic structures and the tectonic evolution of the eastern North Sea

    DEFF Research Database (Denmark)

    Clausen, O.R.; Nielsen, S.B.; Egholm, D.L.

    2011-01-01

    Abundant seismic sections and well data from the Cenozoic succession in the eastern North Sea area generally reveal normal faulting, salt tectonics and localized tectonic inversion. However, inferences on the Cenozoic dynamic evolution of the region require thorough analysis of interactions between...... or cover tectonism took place. Our objectives are thus 1) to analyze the interaction between basement and cover structures, and if possible 2) to relate the structures to the regional tectonic evolution. The Zechstein evaporites pinch out onto the Ringkøbing-Fyn High, which in the eastern North Sea...... influencede.g. Miocene deposition and controlled the generation of second order faults. The latter detached along the top Chalk Group due to the topography generated during faulting, i.e. they are second order detachment surfaces. We conclude that the regional tectonic significance of the Cenozoic structures...

  5. Minimal approaches to genetic improvement of growth rates in white spruce

    Science.gov (United States)

    D.T. Lester

    1973-01-01

    Several features of central importance to genetic improvement of white spruce have been demonstrated by tree breeders. First, white spruce is genetically a highly variable species and much of the existent variation can be readily incorporated in planting stock (Jeffers 1969, Holst and Teich 1969). Second, local seed often is not the best for rapid growth (Nienstaedt...

  6. Organic halogens in spruce forest throughfall

    DEFF Research Database (Denmark)

    Öberg, G.; Johansen, C.; Grøn, C.

    1998-01-01

    . No relationship between the position of the collectors and the forest edge or dominating wind-direction was found, suggesting that dry deposition was not a major source. The concentration of organic halogens was related to that of organic carbon and decreased from the tree-trunk and outwards. In addition......, the concentrations were higher during the growing season than during the dormant season. This indicates that the major part of the organic carbon and organic halogens in spruce forest throughfall originates from canopy leachates or other internal sources. (C) 1998 Elsevier Science Ltd.......Deposition of dissolved organic halogens by throughfall was determined in a small spruce forest site in Denmark (56 degrees 28'N, 8 degrees 24'E). The mean annual deposition of dissolved organic halogens was 377 g ha(-1)yr(-1), and larger than the general deposition by precipitation...

  7. Relationship between norway spruce status and soil water base cations/aluminum rations in the Czech republic

    Czech Academy of Sciences Publication Activity Database

    Hruška, J.; Cudlín, Pavel; Krám, P.

    2001-01-01

    Roč. 130, - (2001), s. 983-988 ISSN 0049-6979 R&D Projects: GA ČR GA206/94/0832 Institutional research plan: CEZ:AV0Z6087904 Keywords : BC/Al molar ratio * norway spruce * crown structure Subject RIV: GK - Forestry Impact factor: 0.772, year: 2001

  8. Forest floor carbon exchange of a boreal black spruce forest in eastern North America

    Directory of Open Access Journals (Sweden)

    O. Bergeron

    2009-09-01

    Full Text Available This study reports continuous automated measurements of forest floor carbon (C exchange over feathermoss, lichen, and sphagnum micro-sites in a black spruce forest in eastern North America during snow-free periods over three years. The response of soil respiration (Rs-auto and forest floor photosynthesis (Pff to environmental factors was determined. The seasonal contributions of scaled up Rs-auto adjusted for spatial representativeness (Rs-adj and Pff (Pff-eco relative to that of total ecosystem respiration (Re and photosynthesis (Peco, respectively, were also quantified.

    Shallow (5 cm soil temperature explained 67–86% of the variation in Rs-auto for all ground cover types, while deeper (50 and 100 cm soil temperatures were related to Rs-auto only for the feathermoss micro-sites. Base respiration was consistently lower under feathermoss, intermediate under sphagnum, and higher under lichen during all three years. The Rs-adj/Re ratio increased from spring through autumn and ranged from 0.85 to 0.87 annually for the snow-free period. The Rs-adj/Re ratio was negatively correlated with the difference between air and shallow soil temperature and this correlation was more pronounced in autumn than summer and spring.

    Maximum photosynthetic capacity of the forest floor (Pff-max saturated at low irradiance levels (~200 μmol m−2 s−1 and decreased with increasing air temperature and vapor pressure deficit for all three ground cover types, suggesting that Pff was more limited by desiccation than by light availability. Pff-max was lowest for sphagnum, intermediate for feathermoss, and highest for lichen for two

  9. Norway spruce crown structure changes under long-term multiple stress impact in Central European Mts

    Czech Academy of Sciences Publication Activity Database

    Moravec, Ivo; Cudlín, Pavel; Polák, T.

    2003-01-01

    Roč. 22, Suppl. 1 (2003), s. 252-255 ISSN 1335-342X. [Long Term Air Pollution Effect on Forest Ecosystems (International Meeting for Specialists in Air Pollution Effects on Forest Ecosystems)/20./. Zvolen, 30.08.2002-01.09.2002] R&D Projects: GA MŠk OK 355 Institutional research plan: CEZ:AV0Z6087904 Keywords : Norway spruce * crown transformation * tree status Subject RIV: EH - Ecology, Behaviour Impact factor: 0.100, year: 2003

  10. Transcriptome mining, functional characterization, and phylogeny of a large terpene synthase gene family in spruce (Picea spp.

    Directory of Open Access Journals (Sweden)

    Dullat Harpreet K

    2011-03-01

    Full Text Available Abstract Background In conifers, terpene synthases (TPSs of the gymnosperm-specific TPS-d subfamily form a diverse array of mono-, sesqui-, and diterpenoid compounds, which are components of the oleoresin secretions and volatile emissions. These compounds contribute to defence against herbivores and pathogens and perhaps also protect against abiotic stress. Results The availability of extensive transcriptome resources in the form of expressed sequence tags (ESTs and full-length cDNAs in several spruce (Picea species allowed us to estimate that a conifer genome contains at least 69 unique and transcriptionally active TPS genes. This number is comparable to the number of TPSs found in any of the sequenced and well-annotated angiosperm genomes. We functionally characterized a total of 21 spruce TPSs: 12 from Sitka spruce (P. sitchensis, 5 from white spruce (P. glauca, and 4 from hybrid white spruce (P. glauca × P. engelmannii, which included 15 monoterpene synthases, 4 sesquiterpene synthases, and 2 diterpene synthases. Conclusions The functional diversity of these characterized TPSs parallels the diversity of terpenoids found in the oleoresin and volatile emissions of Sitka spruce and provides a context for understanding this chemical diversity at the molecular and mechanistic levels. The comparative characterization of Sitka spruce and Norway spruce diterpene synthases revealed the natural occurrence of TPS sequence variants between closely related spruce species, confirming a previous prediction from site-directed mutagenesis and modelling.

  11. Near Real-time Ecological Forecasting of Peatland Responses to Warming and CO2 Treatment through EcoPAD-SPRUCE

    Science.gov (United States)

    Huang, Y.; Jiang, J.; Stacy, M.; Ricciuto, D. M.; Hanson, P. J.; Sundi, N.; Luo, Y.

    2016-12-01

    Ecological forecasting is critical in various aspects of our coupled human-nature systems, such as disaster risk reduction, natural resource management and climate change mitigation. Novel advancements are in urgent need to deepen our understandings of ecosystem dynamics, boost the predictive capacity of ecology, and provide timely and effective information for decision-makers in a rapidly changing world. Our Ecological Platform for Assimilation of Data (EcoPAD) facilitates the integration of current best knowledge from models, manipulative experimentations, observations and other modern techniques and provides both near real-time and long-term forecasting of ecosystem dynamics. As a case study, the web-based EcoPAD platform synchronizes real- or near real-time field measurements from the Spruce and Peatland Responses Under Climatic and Environmental Change Experiment (SPRUCE), a whole ecosystem warming and CO2 enrichment treatment experiment, assimilates multiple data streams into process based models, enhances timely feedback between modelers and experimenters, and ultimately improves ecosystem forecasting and makes best utilization of current knowledge. In addition to enable users to (i) estimate model parameters or state variables, (ii) quantify uncertainty of estimated parameters and projected states of ecosystems, (iii) evaluate model structures, (iv) assess sampling strategies, and (v) conduct ecological forecasting, EcoPAD-SPRUCE automated the workflow from real-time data acquisition, model simulation to result visualization. EcoPAD-SPRUCE promotes seamless feedback between modelers and experimenters, hand in hand to make better forecasting of future changes. The framework of EcoPAD-SPRUCE (with flexible API, Application Programming Interface) is easily portable and will benefit scientific communities, policy makers as well as the general public.

  12. INVESTIGATION OF THE ESSENTIAL OIL OF EUROPEAN SPRUCE ROOTS (LAT. PICEA ABIES H. KARST, PINACEAE FAMILY

    Directory of Open Access Journals (Sweden)

    D. K. Guljaev

    2017-01-01

    Full Text Available The article is devoted to the study of essential oil extracted from the roots of European Spruce (Lat. Picea abies.The aim is to establish the component composition of the essential oil and the peculiarities of its localization in the roots of European Spruce.Materials and methods. The objects of the study are the roots of European Spruce not longer than two centimeters in diameter, peeled and dried. The study of their anatomical signs was carried out according to the methodology of the State Pharmacopoeia of the Russian Federation (the XIII-th edition with the”Biomed-6” microscope using the DCN 510 nozzle. The essential oil was obtained by hydrodistillation of European Spruce roots using the device of Clevenger by Method 2 of the State Pharmacopoeia of the Russian Federation (the XIII-th edition. The component composition of the essential oil was determined by an Agilent 7890A gas-liquid chromatograph with an Agilent 5975C mass-selective detector.Results and discussion. As a result of the microscopic examination of the roots of European Spruce, it was established that the essential oil is localized mainly in resinous courses located in the wood of the root. In the central part of the root, resin moves are of a larger diameter. More than 18 components were found in the essential oil of European Spruce roots, 14 of them were identifi ed. The main component of the essential oil is sesquiterpene lactone – tanbergol.Conclusion. The essential oil of European Spruce roots has a unique component composition that includes components not characteristic for the essential oil of spruce greenery. The difference in the component composition indicates the difference in properties and pharmacological activity. Further studies are of interest for determining the prospects of using European Spruce roots.

  13. Impact of environmental change on the radioecology of spruce trees in Upper Austria

    International Nuclear Information System (INIS)

    Seidel, Claudia; Gruber, V.; Baumgartner, A.; Fuerst, A.

    2008-01-01

    In a two years project spruce needle samples of the Austrian Bioindicator Grid were analysed by gamma spectrometry to detect the geographical and temporal distribution of radionuclides in spruce needles of the last 25 years with the main focus on the radioactive contamination before and after the Chernobyl fallout in 1986. This radioecological evaluation is an important part of an existing environmental surveillance programme in Upper Austria in order to gain basic information on the impact of environmental changes on the radioecological behaviour of spruce trees. Moreover, the results of the current studies can be an important input for the discussion of using whole trees for biomass energy. Every year spruce needle samples of the two youngest needle sprouts are taken from two spruce trees at each location of the Bioindicator Grid. For this study samples of selected locations evenly spaced out among the area of Upper Austria were analysed for different natural and anthropogenic radionuclides: 137 Cs, 40 K, 210 Pb, 226 Ra, 228 Ra, 238 U. Additionally, soil samples were taken at selected sites to study the relationship between 137 Cs and 40 K-activity concentrations in soils and spruce needles and to estimate transfer factors. Another important question of the study is the correlation between anthropogenic pollutants and radionuclides. To date more than 500 spruce needle samples were analysed. 137 Cs- activity concentrations reach D.L. (2 Bq/kg)-5150 Bq/kg, whereas the maximum was measured after the Chernobyl fallout in 1986. In time series of 137 Cs-activity concentrations Caesium cycling was observed. The activity concentrations of 40 K reached D.L. (15 Bq/kg)- 294 Bq/kg and 210 Pb reach D.L. (5 Bq/kg)-45 Bq/kg. The measured 226 Ra, 228 Ra and 238 U concentrations were mostly below detection limits. Most samples of younger sprouts revealed higher 137 Cs activity concentrations than older sprouts. 40 K-activity concentrations showed nearly the same level in

  14. Radioecological investigations on tree rings of spruce

    International Nuclear Information System (INIS)

    Haas, G.; Mueller, A.

    1995-01-01

    Tree ring analysis contributes essentially to the explanation of physiological and element-specific transport phenomena in trees. After the accident in Chernobyl the behaviour of Cs-134 and Cs-137 in trees is most informative for the prediction of the future development of the distribution of these elements. In this study the uptake and the long term behaviour of Cs-134, Cs-137, Pb-210, Ra-226, Ra-228, K-40, Th-228, Th-230, Th-232, U-234 and U-238 in tree rings of spruce are examined by α- and γ-spectrometry. All samples are dried at 105C and ashed at 450C in a muffle furnace. The distributions found in the tree rings vary for different radionuclides. A soil profile from the spruce stand provides additional information

  15. Adaptive Evolution and Demographic History of Norway Spruce (Picea Abies)

    OpenAIRE

    Källman, Thomas

    2009-01-01

    One of the major challenges in evolutionary biology is to determine the genetic basis of adaptive variation. In Norway spruce (Picea abies) the timing of bud set shows a very strong latitudinal cline despite a very low genetic differentiation between populations. The timing of bud set in Norway spruce is under strong genetic control and triggered by changes in photoperiod, but no genes controlling this response have so far been described. In this thesis we used a combination of functional stu...

  16. Effect of stand edge on the natural regeneration of spruce, beech and Douglas-fir

    Directory of Open Access Journals (Sweden)

    Lumír Dobrovolný

    2012-01-01

    Full Text Available Our work aimed at studying the strategy of woody plants regeneration during the regeneration of a spruce stand with the admixture of beech and Douglas-fir by border cutting (NW-SE aspect on acidic sites of higher elevations in the Bohemian-Moravian Upland. Spruce is better adapted to bear shade than Douglas-fir. Nevertheless, in optimal light conditions up to a distance of ca. 35 m (about 16% DIFFSF from the stand edge, the Douglas-fir can put the spruce into danger as to height growth. By contrast to beech, the density of spruce is significantly higher within the distance of 45 m (about 15% DIFFSF from the stand edge but further on the situation would change to the benefit of beech. The density of Douglas-fir significantly dominates over beech within a distance of 35 m from the stand edge; from 55 m (less than 15% DIFFSF, the situation changes in favour of beech. Beech can survive in full shade deep in the stand core waiting for its opportunity to come. As compared to spruce and Douglas-fir, the height growth of beech was at all times significantly greater at a distance of 25 m from the stand edge. Converted to practical conditions, spruce and Douglas-fir with individually admixed beech seedlings showed good prosperity approximately up to a distance of one stand height from the edge. A mixture of spruce and beech did well at a greater distance but good prosperity at a distance of 2–3 stand heights was shown only by beech. Thus, border regeneration eliminates disadvantages of the climatic extremes of clear-cutting and specifics of shelterwood felling during which one – usually shade-tolerant tree species dominates in the natural regeneration (e.g. beech.

  17. Characteristics and modeling of spruce wood under dynamic compression load

    International Nuclear Information System (INIS)

    Eisenacher, Germar

    2014-01-01

    Spruce wood is frequently used as an energy absorbing material in impact limiters of packages for the transportation of radioactive material. A 9m drop test onto an unyielding target is mandatory for the packages. The impact results in a dynamic compression load of the spruce wood inside the impact limiter. The lateral dilation of the wood is restrained thereby due to encasing steel sheets. This work's objective was to provide a material model for spruce wood based on experimental investigations to enable the calculation of such loading conditions. About 600 crush tests with cubical spruce wood specimens were performed to characterize the material. The compression was up to 70% and the material was assumed to be transversely isotropic. Particularly the lateral constraint showed to have an important effect: the material develops a high lateral dilation without lateral constraint. The force-displacement characteristics show a comparably low force level and no or only slight hardening. Distinctive softening occurs after the linear-elastic region when loaded parallel to the fiber. On the other hand, using a lateral constraint results in significantly higher general force levels, distinctive hardening and lateral forces. The softening effect when loaded parallel to the fiber is less distinctive. Strain rate and temperature raise or lower the strength level, which was quantified for the applicable ranges of impact limiters. The hypothesis of an uncoupled evolution of the yield surface was proposed based on the experimental findings. It postulates an independent strength evolution with deviatoric and volumetric deformation. The hypothesis could be established using the first modeling approach, the modified LS-DYNA material model MAT075. A transversely isotropic material model was developed based thereupon and implemented in LS-DYNA. The material characteristics of spruce wood were considered using a multi-surface yield criterion and a non-associated flow rule. The yield

  18. Influence of nutrition and various substrates on spruce seedling growth

    Directory of Open Access Journals (Sweden)

    Đukić Matilda

    2004-01-01

    Full Text Available The results of the influence of main macronutrients (N, P, and K on growth and development of spruce (Picea abies L. Karst one-year old seedlings are presented. They were grown in containers, in nursery conditions, on four different substrates. There is a good influence on biogenous element contents, height, root collar diameter, needle length and mass, root mass as well as physiological vitality of spruce seedlings. It was observed that the effect of nutrition depends also on the type of substrate.

  19. Experimental Whole-Ecosystem Warming Alters Vegetation Phenology in a Boreal Spruce Bog: Initial Results from the SPRUCE Experiment

    Science.gov (United States)

    Richardson, A. D.

    2016-12-01

    Phenology is one of the most robust indicators of the biological impacts of global change. However, the response of phenology to future environmental conditions still remains highly uncertain because of the challenges associated with conducting realistic manipulative experiments. At the SPRUCE (Spruce and Peatland Responses Under Climatic and Environmental Change) experiment in the north-central United States, experimental temperature (0 to +9°C above ambient) and CO2 (ambient and elevated) treatments are being applied to mature, and intact, Picea mariana-Sphagnum spp. bog communities in their native habitat through the use of ten large (approximately 12 m wide, 10 m high) open-topped enclosures. We are tracking vegetation green-up and senescence in these chambers using repeat digital photography. Within each chamber, images are recorded every 30 minutes and uploaded to PhenoCam (http://phenocam.sr.unh.edu), where processed to yield quantitative measures of canopy color. These data are complemented by on-the-ground phenological data collected by human observers. Air warming treatments at SPRUCE began in August 2015. We observed a delay in senescence during autumn 2015 (2-5 days per degree of warming) and an advance in onset during spring 2016 (1-4 days per degree of warming). These patterns are robust across species and methods of phenological observation (i.e. camera-based vs. human observer). And, our results show very little evidence for photoperiod acting as a constraint on the response to warming. Early spring onset and consequent loss of frost hardiness in the warmest chambers proved disadvantageous when a brief period of extreme cold (to -12°C in the control chambers, to -3°C in the +9°C chambers) followed a month of generally mild weather. Foliage mortality for both Larix and Picea was immediate and severe, although both species subsequently re-flushed. These results give support for the hypothesis that warming may enhance the likelihood of spring frost

  20. Structural evolution of Halaban Area, Eastern Arabian Shield, Saudi Arabia

    Science.gov (United States)

    Al-Amri, Yousef; Kassem1, Osama M. K.

    2017-04-01

    Neoproterozoic basement complex comprises a metamorphic/igneous suite (Abt schist and sheared granitoids) with syn-accretionary transpressive structures, unconformably overlain by a post-amalgamation volcanosedimentary sequence. This study aims to attempt to exposed post-accretionary thrusting and thrust-related structures at Halaban area, Eastern Arabian Shield. The Rf/ϕ and Fry methods are utilized on quartz and feldspar porphyroclasts, as well as on mafic crystals, such as hornblende and biotite, in eighteen samples. The X/Z axial ratios range from 1.12 to 4.99 for Rf/ϕ method and from 1.65 to 4.00 for Fry method. The direction of finite strain for the long axes displays clustering along the WNW trend (occasionally N) with slight plunging. Finite strain accumulated without any significant volume change contemporaneously with syn-accretionary transpressive structures. It indicates that the contacts between various lithological units in the Halaban area were formed under brittle to semi-ductile deformation conditions. The penetrative subhorizontal foliation was concurrent with thrusting and shows nearly the same attitudes of tectonic contacts with the overlying nappes. Keywords: Finite strain analysis, volcanosedimentary sequence, Halaban area, Eastern Arabian Shield, Saudi Arabia.

  1. Experimental warming delays autumn senescence in a boreal spruce bog: Initial results from the SPRUCE experiment

    Science.gov (United States)

    Richardson, Andrew; Furze, Morgan; Aubrecht, Donald; Milliman, Thomas; Nettles, Robert; Krassovski, Misha; Hanson, Paul

    2016-04-01

    Phenology is considered one of the most robust indicators of the biological impacts of global change. In temperate and boreal regions, long-term data show that rising temperatures are advancing spring onset (e.g. budburst and flowering) and delaying autumn senescence (e.g. leaf coloration and leaf fall) in a wide range of ecosystems. While warm and cold temperatures, day length and insolation, precipitation and water availability, and other factors, have all been shown to influence plant phenology, the future response of phenology to rising temperatures and elevated CO2 still remains highly uncertain because of the challenges associated with conducting realistic manipulative experiments to simulate future environmental conditions. At the SPRUCE (Spruce and Peatland Responses Under Climatic and Environmental Change) experiment in the north-central United States, experimental temperature (0 to +9° C above ambient) and CO2 (ambient and elevated) treatments are being applied to mature, and intact, Picea mariana-Sphagnum spp. bog communities in their native habitat through the use of ten large (approximately 12 m wide, 10 m high) open-topped enclosures. We are tracking vegetation green-up and senescence in these chambers, at both the individual and whole-community level, using repeat digital photography. Within each chamber, digital camera images are recorded every 30 minutes and uploaded to the PhenoCam (http://phenocam.sr.unh.edu) project web page, where they are displayed in near-real-time. Image processing is conducted nightly to extract quantitative measures of canopy color, which we characterize using Gcc, the green chromatic coordinate. Data from a camera mounted outside the chambers (since November 2014) indicate strong seasonal variation in Gcc for both evergreen shrubs and trees. Shrub Gcc rises steeply in May and June, and declines steeply in September and October. By comparison, tree Gcc rises gradually from March through June, and declines gradually from

  2. Crustal structure and active tectonics in the Eastern Alps

    DEFF Research Database (Denmark)

    Brückl, E.; Behm, M.; Decker, K.

    2010-01-01

    fragment (PA), was interpreted and a triple junction was inferred. The goal of this study has been to relate these deep crustal structures to active tectonics. We used elastic plate modeling to reconsider the Moho fragmentation. We interpret subduction of EU below AD and PA from north to south......During the last decade, a series of controlled source seismic experiments brought new insight into the crustal and lithospheric structure of the Eastern Alps and their adjacent tectonic provinces. A fragmentation of the lithosphere into three blocks, Europe (EU), Adria (AD), and the new Pannonian...

  3. Relationship between deep structure and oil-gas in the eastern Tarim Basin

    Science.gov (United States)

    Yu, Changqing; Qu, Chen; Han, Jianguang

    2017-04-01

    The Tarim Basin is a large composite superimposed basin which developed in the Presinian continental basement. It is an important area for oil and gas replacement in China. In the eastern part of Tarim Basin, the exploration and research degree is very low and less system, especially in the study of tectonic evolution and physical property change. Basing on the study of geophysics, drilling and regional geological data in this area, analysis of comprehensive geophysical, geological and geophysical analysis comparison are lunched by new methods and new technology of geophysical exploration. Fault, tectonic evolution and change of deep character in the eastern Tarim Basin are analyzed in system. Through in-depth study and understanding of the deep structure and physical changes of the eastern region, we obtain the fault characteristics in the study area and the deep structure and physical change maps to better guide the oil and gas exploration in this area. The east area is located in the eastern Tarim Basin, west from the Garr Man depression, Well Kunan 1 - Well Gucheng 4 line to the East, north to Kuruketage uplift group near Qunke 1 wells, south to Cherchen fault zone, east to Lop Nor depression, an area of about 9 * 104 square kilometres, Including the East of Garr Man sag, Yingjisu depression, Kongquehe slope, Tadong low uplift and the Lop Nor uplift, five two grade tectonic units. The east area of Tarim is belonging to Tarim plate. It changes with the evolution of the Tarim plate. The Tarim plate is closely related to the collision between the Yining - the Junggar plate, the Siberia plate and the southern Qiangtang - the central Kunlun plate. Therefore, it creates a complex tectonic pattern in the eastern Tarim basin. Earth electromagnetic, gravity, deep seismic and other geophysical data are processed by a new generation of geophysical information theory and method, including multi-scale inversion of potential field inversion (Hou and Yang, 2011), 3D

  4. Radial Growth Response of Black Spruce Stands Ten Years after Experimental Shelterwoods and Seed-Tree Cuttings in Boreal Forest

    Directory of Open Access Journals (Sweden)

    Miguel Montoro Girona

    2016-10-01

    Full Text Available Partial cutting is thought to be an alternative to achieve sustainable management in boreal forests. However, the effects of intermediate harvest intensity (45%–80% on growth remain unknown in black spruce (Picea mariana (Mill. B.S.P. stands, one of the most widely distributed boreal species with great commercial interest. In this study, we analysed the effect of three experimental shelterwood and one seed-tree treatments on tree radial growth in even-aged black spruce stands, 10 years after intervention. Our results show that radial growth response 8–10 years after cutting was 41% to 62% higher than in untreated plots, with stand structure, treatment, tree position relative to skidding trails, growth before cutting and time having significant interactions. The stand structure conditioned tree growth after cutting, being doubled in younger and denser stands. Tree spatial position had a pronounced effect on radial growth; trees at the edge of the skidding trails showed twice the increase in growth compared to interior trees. Dominant trees before cutting located close to the skidding trails manifested the highest growth response after cutting. This research suggests that the studied treatments are effective to enhance radial wood production of black spruce especially in younger stands, and that the edge effect must be considered in silvicultural management planning.

  5. Microbial Activity in Forest Soil Under Beech, Spruce, Douglas Fir and Fir

    Directory of Open Access Journals (Sweden)

    Hajnal-Jafari Timea

    2016-08-01

    Full Text Available The aim of this research was to investigate the microbial activity in forest soil from different sites under deciduous and coniferous trees in Serbia. One site on Stara planina was under beech trees (Fagus sp. while another under mixture of spruce (Picea sp. and Douglas fir (Pseudotsuga sp.. The site on Kopaonik was under mixture of beech (Fagus sp. and spruce (Picea sp. trees. The site on Tara was dominantly under fir (Abies sp., beech (Fagus sp. and spruce (Picea sp.. The total number of bacteria, the number of actinobacteria, fungi and microorganisms involved in N and C cycles were determined using standard method of agar plates. The activities of dehydrogenase and ß-glucosidase enzymes were measured by spectrophotometric methods. The microbial activity was affected by tree species and sampling time. The highest dehydrogenase activity, total number of bacteria, number of actinobacteria, aminoheterotrophs, amylolytic and cellulolytic microorganisms were determined in soil under beech trees. The highest total number of fungi and number of pectinolytic microorganisms were determined in soil under spruce and Douglas fir trees. The correlation analyses proved the existence of statistically significant interdependency among investigated parameters.

  6. Biosynthesis of the major tetrahydroxystilbenes in spruce, astringin and isorhapontin, proceeds via resveratrol and is enhanced by fungal infection.

    Science.gov (United States)

    Hammerbacher, Almuth; Ralph, Steven G; Bohlmann, Joerg; Fenning, Trevor M; Gershenzon, Jonathan; Schmidt, Axel

    2011-10-01

    Stilbenes are dibenzyl polyphenolic compounds produced in several unrelated plant families that appear to protect against various biotic and abiotic stresses. Stilbene biosynthesis has been well described in economically important plants, such as grape (Vitis vinifera), peanut (Arachis hypogaea), and pine (Pinus species). However, very little is known about the biosynthesis and ecological role of stilbenes in spruce (Picea), an important gymnosperm tree genus in temperate and boreal forests. To investigate the biosynthesis of stilbenes in spruce, we identified two similar stilbene synthase (STS) genes in Norway spruce (Picea abies), PaSTS1 and PaSTS2, which had orthologs with high sequence identity in sitka (Picea sitchensis) and white (Picea glauca) spruce. Despite the conservation of STS sequences in these three spruce species, they differed substantially from angiosperm STSs. Several types of in vitro and in vivo assays revealed that the P. abies STSs catalyze the condensation of p-coumaroyl-coenzyme A and three molecules of malonyl-coenzyme A to yield the trihydroxystilbene resveratrol but do not directly form the dominant spruce stilbenes, which are tetrahydroxylated. However, in transgenic Norway spruce overexpressing PaSTS1, significantly higher amounts of the tetrahydroxystilbene glycosides, astringin and isorhapontin, were produced. This result suggests that the first step of stilbene biosynthesis in spruce is the formation of resveratrol, which is further modified by hydroxylation, O-methylation, and O-glucosylation to yield astringin and isorhapontin. Inoculating spruce with fungal mycelium increased STS transcript abundance and tetrahydroxystilbene glycoside production. Extracts from STS-overexpressing lines significantly inhibited fungal growth in vitro compared with extracts from control lines, suggesting that spruce stilbenes have a role in antifungal defense.

  7. Lophodermium piceae and Tryblidiopsis pinastri. Two latent colonizers of Norway spruce

    Energy Technology Data Exchange (ETDEWEB)

    Lehtijaervi, A. [Swedish Univ. of Agricultural Sciences, Uppsala (Sweden). Dept. of Forest Mycology and Pathology

    1998-12-31

    Among the endophytic microfungi colonizing Norway spruce, the non-pathogenic species Lophodermium piceae and Tryblidiopsis pinastri are ubiquitous. Most Norway spruce in Sweden are colonized by these fungi. L. piceae colonizes healthy needles of various ages, while T pinastri colonizes the bark of branches. New shoots become infected around the time of their emergence. Fruit bodies are formed after the needles and branches die. It was found that L. piceae colonized needles during the summer and early autumn, about six weeks after a prolonged period of intensive rain. The number of individual fungal mycelia per needle increased linearly with needle age. In experiments with trees, irrigation resulted in lower total colonization and delayed colonization of the current-year needles, regardless of whether fertilizer had been added. Colonization by L. piceae was lowest in the ammonium sulfate treatment, which simulated the deposition of air pollutants. Colonization was highest in the control and drought treatments, which did not differ significantly from each other. Only a few pathogenic fungi can damage Norway spruce needles. Needles damaged by the rust fungus Chrysomyxa abietis were investigated to study interactions between fungi. The frequency and intensity of L. piceae colonization were found to be similar for C. abietis infected and healthy needles. However, in needles partially infected with rust, L. piceae seemed to establish itself easier in the rust-infected part than in the green part. The genetic structures of populations of T. pinastri in southern Sweden and Finland were investigated using DNA markers produced by means of arbitrarily primed PCR. Single spore isolates from apothecia were used in the analysis. A considerable amount of variation was detected. No geographical differentiation was found among the populations studied

  8. Long-term development of nursing mixtures of Sitka spruce and larch species in an experiment in northern Scotland

    Directory of Open Access Journals (Sweden)

    William L. Mason

    2014-12-01

    Full Text Available Aim of the study: An experiment was established in 1966 to compare the growth and development of 50: 50 mixtures of Sitka spruce (Picea sitchensis with either Japanese larch (Larix kaempferi or tamarack (L. laricina with that found in pure plots of Sitka spruce. The site was one of moderate nitrogen availability where the presence of heather (Calluna vulgaris could be expected to limit the growth ofSitka spruce.Area of the study: North-east Scotland.Material and methods: There were different patterns of spruce growth in the pure plots and in the mixtures, with faster spruce growth in mixture in the years approaching and immediately following canopy closure (i.e. ages 15-25. Foliage analysis suggested that this was linked with improved nitrogen status of spruce trees in the mixed compared to the pure plots.Main results: At years 20 and 25 there were significant differences in height, diameter, and basal area between treatments, with the largest basal area being found in the Japanese larch/Sitka spruce mixtures, indicative of overyielding in the mixed plots. However, when the experiment was clearfelled at 41 years of age, all treatments had self-thinned to produce spruce dominated stands of similar height with only an occasional larch tree surviving in plots that were originally 50:50 mixtures.Research highlights: There were no differences between treatments in basal area, harvested volume or sawlog outturn after 41 years. These results can be interpreted as showing facilitation between the larch and the spruce during the establishment phase followed by competition for light once canopy closure had occurred.Keywords: Mixed stand dynamics; facilitation; nitrogen status; product outturn.

  9. Belowground effects of enhanced tropospheric ozone and drought in a beech/spruce forest (Fagus sylvatica L./Picea abies [L.] Karst)

    International Nuclear Information System (INIS)

    Nikolova, Petia S.; Andersen, Christian P.; Blaschke, Helmut; Matyssek, Rainer; Haeberle, Karl-Heinz

    2010-01-01

    The effects of experimentally elevated O 3 on soil respiration rates, standing fine-root biomass, fine-root production and δ 13 C signature of newly produced fine roots were investigated in an adult European beech/Norway spruce forest in Germany during two subsequent years with contrasting rainfall patterns. During humid 2002, soil respiration rate was enhanced under elevated O 3 under beech and spruce, and was related to O 3 -stimulated fine-root production only in beech. During dry 2003, the stimulating effect of O 3 on soil respiration rate vanished under spruce, which was correlated with decreased fine-root production in spruce under drought, irrespective of the O 3 regime. δ 13 C signature of newly formed fine-roots was consistent with the differing g s of beech and spruce, and indicated stomatal limitation by O 3 in beech and by drought in spruce. Our study showed that drought can override the stimulating O 3 effects on fine-root dynamics and soil respiration in mature beech and spruce forests. - Drought has the capacity to override the stimulating ozone effect on soil respiration in adult European beech/Norway spruce forest.

  10. Cloning and characterization of chitinases from interior spruce and lodgepole pine.

    Science.gov (United States)

    Kolosova, N; Breuil, C; Bohlmann, J

    2014-05-01

    Chitinases have been implicated in the defence of conifers against insects and pathogens. cDNA for six chitinases were cloned from interior spruce (Picea glauca x engelmannii) and four from lodgepole pine (Pinus contorta). The cloned interior spruce chitinases were annotated class I PgeChia1-1 and PgeChia1-2, class II PgeChia2-1, class IV PgeChia4-1, and class VII PgeChia7-1 and PgeChia7-2; lodgepole pine chitinases were annotated class I PcChia1-1, class IV PcChia4-1, and class VII PcChia7-1 and PcChia7-2. Chitinases were expressed in Escherichia coli with maltose-binding-protein tags and soluble proteins purified. Functional characterization demonstrated chitinolytic activity for the three class I chitinases PgeChia1-1, PgeChia1-2 and PcChia1-1. Transcript analysis established strong induction of most of the tested chitinases, including all three class I chitinases, in interior spruce and lodgepole pine in response to inoculation with bark beetle associated fungi (Leptographium abietinum and Grosmannia clavigera) and in interior spruce in response to weevil (Pissodes strobi) feeding. Evidence of chitinolytic activity and inducibility by fungal and insect attack support the involvement of these chitinases in conifer defense. Copyright © 2014 Elsevier Ltd. All rights reserved.

  11. MICROSTRUCTURE MODIFICATIONS INDUCED IN SPRUCE WOOD BY FREEZING

    Directory of Open Access Journals (Sweden)

    Maria Bernadett SZMUTKU

    2011-12-01

    Full Text Available Scanning Electron Microscopy (SEM is amodern, non-invasive method for objective andspecialized image analysis of anatomical materialfeatures at microscopic level. Referring to wood, itoffers the possibility to view in 3D a bunch ofneighboring cells, in all three grain directions.This allows the imaging of modifications thatmight appear in the structure of the wood cellmembrane (e.g. micro-fissures caused by differentfactors, including temperature variations. This paperpresents the results of the SEM analysis performedon European spruce (Picea abies samples, cut fromboards which were subjected to freezing and thawingunder different conditions of temperature variationand time of exposure.The main aim of this research was to reveal theconditions which determine the occurrence of microfissuresin the cell wall and consequently lead tostrength losses in wood.

  12. Occurrence of spruce bark beetles in forest stands at different levels of air pollution stress

    International Nuclear Information System (INIS)

    Grodzki, Wojciech; McManus, Michael; Knizek, Milos; Meshkova, Valentina; Mihalciuc, Vasile; Novotny, Julius; Turcani, Marek; Slobodyan, Yaroslav

    2004-01-01

    The spruce bark beetle, Ips typographus (L.) is the most serious pest of mature spruce stands, mainly Norway spruce, Picea abies (L.) Karst. throughout Eurasia. A complex of weather-related events and other environmental stresses are reported to predispose spruce stands to bark beetle attack and subsequent tree mortality; however the possible role of industrial pollution as a predisposing factor to attack by this species is poorly understood. The abundance and dynamics of I. typographus populations was evaluated in 60-80 year old Norway spruce stands occurring on 10x50 ha sites in five countries within the Carpathian range that were selected in proximity to established ozone measurement sites. Data were recorded on several parameters including the volume of infested trees, captures of adult beetles in pheromone traps, number of attacks, and the presence and relative abundance of associated bark beetle species. In several cases, stands adjacent to sites with higher ozone values were associated with higher bark beetle populations. The volume of sanitary cuttings, a reflection of tree mortality, and the mean daily capture of beetles in pheromone traps were significantly higher at sites where the O 3 level was higher. However, the mean infestation density on trees was higher in plots associated with lower O 3 levels. Captures of beetles in pheromone traps and infestation densities were higher in the zone above 800 m. However, none of the relationships was conclusive, suggesting that spruce bark beetle dynamics are driven by a complex interaction of biotic and abiotic factors and not by a single parameter such as air pollution. - Air pollution (ozone) can be one of predisposing factors that increases the susceptibility of mountain Norway spruce stands to attack by Ips typographus and associated bark beetle species

  13. Biological and Energy Productivity of Natural Spruce Forests in the Ukrainian Carpathians

    Directory of Open Access Journals (Sweden)

    R. D. Vasilishyn

    2014-10-01

    Full Text Available The modern practice of forestry production in Ukraine, which is in the process of implementing the conceptual changes in forest management and harmonization of its basic approaches to the basics of sustainable development, requires a significant expansion of the current regulatory and informational tools used to assess the ecological functions of forests. For this purpose, during the 2012–2014, as part of an international project GESAPU, models and tables of bioproductivity for forest tree species in Ukraine were completed. The article presents the results of modeling the dynamics of the conversion coefficients for the main components of phytomass of modal natural spruce forests of the Carpathian region of Ukraine based on information from 32 plots in the database of «Forest Phytomass of Ukraine». According to the state forest accounting of Ukraine as of January 1, 2011, the spruce forests in the Ukrainian Carpathians cover an area of 426.2 thousand ha, 45 % of which are spruce of natural origin. To evaluate the productivity of modal dynamics of pure and mixed spruce stands, the study developed models of the stock and overall productivity, derived by Bertalanffy growth function. On the basis of these models, normative reference tables of biological productivity of natural modal spruce forests of the Ukrainian Carpathians were developed. To successfully meet the challenges of evaluating the energy possibilities of forestry of Ukraine, the study used tables of energetic productivity of investigated stands. Built on the basis of the tables of bioproductivity, they reflect the dynamic processes of energy storage in the phytomass components and can be used in forest management to predict volumes of energetic woods.

  14. Disentangling the effects of acidic air pollution, atmospheric CO2 , and climate change on recent growth of red spruce trees in the Central Appalachian Mountains.

    Science.gov (United States)

    Mathias, Justin M; Thomas, Richard B

    2018-05-20

    In the 45 years after legislation of the Clean Air Act, there has been tremendous progress in reducing acidic air pollutants in the eastern United States, yet limited evidence exists that cleaner air has improved forest health. Here, we investigate the influence of recent environmental changes on the growth and physiology of red spruce (Picea rubens Sarg.) trees, a key indicator species of forest health, spanning three locations along a 100 km transect in the Central Appalachian Mountains. We incorporated a multiproxy approach using 75-year tree ring chronologies of basal tree growth, carbon isotope discrimination (∆ 13 C, a proxy for leaf gas exchange), and δ 15 N (a proxy for ecosystem N status) to examine tree and ecosystem level responses to environmental change. Results reveal the two most important factors driving increased tree growth since ca. 1989 are reductions in acidic sulfur pollution and increases in atmospheric CO 2 , while reductions in pollutant emissions of NO x and warmer springs played smaller, but significant roles. Tree ring ∆ 13 C signatures increased significantly since 1989, concurrently with significant declines in tree ring δ 15 N signatures. These isotope chronologies provide strong evidence that simultaneous changes in C and N cycling, including greater photosynthesis and stomatal conductance of trees and increases in ecosystem N retention, were related to recent increases in red spruce tree growth and are consequential to ecosystem recovery from acidic pollution. Intrinsic water use efficiency (iWUE) of the red spruce trees increased by ~51% across the 75-year chronology, and was driven by changes in atmospheric CO 2 and acid pollution, but iWUE was not linked to recent increases in tree growth. This study documents the complex environmental interactions that have contributed to the recovery of red spruce forest ecosystems from pervasive acidic air pollution beginning in 1989, about 15 years after acidic pollutants started to

  15. Lessons from native spruce forests in Alaska: managing Sitka spruce plantations worldwide to benefit biodiversity and ecosystem services

    Science.gov (United States)

    Robert L. Deal; Paul Hennon; Richard O' Hanlon; David D' Amore

    2014-01-01

    There is increasing interest worldwide in managing forests to maintain or improve biodiversity, enhance ecosystem services and assure long-term sustainability of forest resources. An important goal of forest management is to increase stand diversity, provide wildlife habitat and improve forest species diversity. We synthesize results from natural spruce forests in...

  16. Assessing the suitability of Norway spruce wood as an environmental archive for sulphur

    International Nuclear Information System (INIS)

    Barrelet, T.; Ulrich, A.; Rennenberg, H.; Zwicky, C.N.; Kraehenbuehl, U.

    2008-01-01

    The aim of this study was to assess the suitability of Norway spruces (Picea abies L. Karst.) as an environmental archive for sulphur. For this purpose spruce trees were sampled in two distinct regions of Switzerland: the Alps and the Swiss Plateau, which differ significantly with respect to S immission. Wood samples were measured using two methods: LASER Ablation high resolution inductively coupled plasma mass spectrometry (LA-HR-ICP-MS) and inductively coupled plasma optical emission spectrometry (ICP-OES) after acid digestion. Independently corroborated by previous measurements of sulphur in peat bogs, the rise and fall of sulphur dioxide pollution in Switzerland appears to be reflected in spruce wood sulphur content. While the wood sulphur content profile of trees sampled in the Alps is relatively flat, the profiles of trees located on the Swiss Plateau display a characteristic sulphur peak. This corresponds to air pollution data in the different regions and indicates that the trees reacted on the changing S supply and recorded a pollution signal in the wood. - Norway spruce is a suitable archive for sulphur when physiological aspects and limitations are taken into account

  17. Specific impacts of beech and Norway spruce on the structure and diversity of the rhizosphere and soil microbial communities.

    Science.gov (United States)

    Uroz, S; Oger, P; Tisserand, E; Cébron, A; Turpault, M-P; Buée, M; De Boer, W; Leveau, J H J; Frey-Klett, P

    2016-06-15

    The impacts of plant species on the microbial communities and physico-chemical characteristics of soil are well documented for many herbs, grasses and legumes but much less so for tree species. Here, we investigate by rRNA and ITS amplicon sequencing the diversity of microorganisms from the three domains of life (Archaea, Bacteria and Eukaryota:Fungi) in soil samples taken from the forest experimental site of Breuil-Chenue (France). We discovered significant differences in the abundance, composition and structure of the microbial communities associated with two phylogenetically distant tree species of the same age, deciduous European beech (Fagus sylvatica) and coniferous Norway spruce (Picea abies Karst), planted in the same soil. Our results suggest a significant effect of tree species on soil microbiota though in different ways for each of the three microbial groups. Fungal and archaeal community structures and compositions are mainly determined according to tree species, whereas bacterial communities differ to a great degree between rhizosphere and bulk soils, regardless of the tree species. These results were confirmed by quantitative PCR, which revealed significant enrichment of specific bacterial genera, such as Burkholderia and Collimonas, known for their ability to weather minerals within the tree root vicinity.

  18. Micro- and macro-geographic scale effect on the molecular imprint of selection and adaptation in Norway spruce.

    Directory of Open Access Journals (Sweden)

    Marta Scalfi

    Full Text Available Forest tree species of temperate and boreal regions have undergone a long history of demographic changes and evolutionary adaptations. The main objective of this study was to detect signals of selection in Norway spruce (Picea abies [L.] Karst, at different sampling-scales and to investigate, accounting for population structure, the effect of environment on species genetic diversity. A total of 384 single nucleotide polymorphisms (SNPs representing 290 genes were genotyped at two geographic scales: across 12 populations distributed along two altitudinal-transects in the Alps (micro-geographic scale, and across 27 populations belonging to the range of Norway spruce in central and south-east Europe (macro-geographic scale. At the macrogeographic scale, principal component analysis combined with Bayesian clustering revealed three major clusters, corresponding to the main areas of southern spruce occurrence, i.e. the Alps, Carpathians, and Hercynia. The populations along the altitudinal transects were not differentiated. To assess the role of selection in structuring genetic variation, we applied a Bayesian and coalescent-based F(ST-outlier method and tested for correlations between allele frequencies and climatic variables using regression analyses. At the macro-geographic scale, the F(ST-outlier methods detected together 11 F(ST-outliers. Six outliers were detected when the same analyses were carried out taking into account the genetic structure. Regression analyses with population structure correction resulted in the identification of two (micro-geographic scale and 38 SNPs (macro-geographic scale significantly correlated with temperature and/or precipitation. Six of these loci overlapped with F(ST-outliers, among them two loci encoding an enzyme involved in riboflavin biosynthesis and a sucrose synthase. The results of this study indicate a strong relationship between genetic and environmental variation at both geographic scales. It also

  19. Micro- and macro-geographic scale effect on the molecular imprint of selection and adaptation in Norway spruce.

    Science.gov (United States)

    Scalfi, Marta; Mosca, Elena; Di Pierro, Erica Adele; Troggio, Michela; Vendramin, Giovanni Giuseppe; Sperisen, Christoph; La Porta, Nicola; Neale, David B

    2014-01-01

    Forest tree species of temperate and boreal regions have undergone a long history of demographic changes and evolutionary adaptations. The main objective of this study was to detect signals of selection in Norway spruce (Picea abies [L.] Karst), at different sampling-scales and to investigate, accounting for population structure, the effect of environment on species genetic diversity. A total of 384 single nucleotide polymorphisms (SNPs) representing 290 genes were genotyped at two geographic scales: across 12 populations distributed along two altitudinal-transects in the Alps (micro-geographic scale), and across 27 populations belonging to the range of Norway spruce in central and south-east Europe (macro-geographic scale). At the macrogeographic scale, principal component analysis combined with Bayesian clustering revealed three major clusters, corresponding to the main areas of southern spruce occurrence, i.e. the Alps, Carpathians, and Hercynia. The populations along the altitudinal transects were not differentiated. To assess the role of selection in structuring genetic variation, we applied a Bayesian and coalescent-based F(ST)-outlier method and tested for correlations between allele frequencies and climatic variables using regression analyses. At the macro-geographic scale, the F(ST)-outlier methods detected together 11 F(ST)-outliers. Six outliers were detected when the same analyses were carried out taking into account the genetic structure. Regression analyses with population structure correction resulted in the identification of two (micro-geographic scale) and 38 SNPs (macro-geographic scale) significantly correlated with temperature and/or precipitation. Six of these loci overlapped with F(ST)-outliers, among them two loci encoding an enzyme involved in riboflavin biosynthesis and a sucrose synthase. The results of this study indicate a strong relationship between genetic and environmental variation at both geographic scales. It also suggests that an

  20. Structurally controlled 'teleconnection' of large-scale mass wasting (Eastern Alps)

    Science.gov (United States)

    Ostermann, Marc; Sanders, Diethard

    2015-04-01

    In the Brenner Pass area (Eastern Alps) , closely ahead of the most northward outlier ('nose') of the Southern-Alpine continental indenter, abundant deep-seated gravitational slope deformations and a cluster of five post-glacial rockslides are present. The indenter of roughly triangular shape formed during Neogene collision of the Southern-Alpine basement with the Eastern-Alpine nappe stack. Compression by the indenter activated a N-S striking, roughly W-E extensional fault northward of the nose of the indenter (Brenner-normal fault; BNF), and lengthened the Eastern-Alpine edifice along a set of major strike-slip faults. These fault zones display high seismicity, and are the preferred locus of catastrophic rapid slope failures (rockslides, rock avalanches) and deep-seated gravitational slope deformations. The seismotectonic stress field, earthquake activity, and structural data all indicate that the South-Alpine indenter still - or again - exerts compression; in consequence, the northward adjacent Eastern Alps are subject mainly to extension and strike-slip. For the rockslides in the Brenner Pass area, and for the deep-seated gravitational slope deformations, the fault zones combined with high seismic activity predispose massive slope failures. Structural data and earthquakes mainly record ~W-E extension within an Eastern Alpine basement block (Oetztal-Stubai basement complex) in the hangingwall of the BNF. In the Northern Calcareous Alps NW of the Oetztal-Stubai basement complex, dextral faults provide defacement scars for large rockfalls and rockslides. Towards the West, these dextral faults merge into a NNW-SSE striking sinistral fault zone that, in turn, displays high seismic activity and is the locus of another rockslide cluster (Fern Pass cluster; Prager et al., 2008). By its kinematics dictated by the South-Alpine indenter, the relatively rigid Oetztal-Stubai basement block relays faulting and associated mass-wasting over a N-S distance of more than 60

  1. Seasonal flight patterns of the Spruce bark beetle (Ips typographus) in Sweden

    OpenAIRE

    Öhrn, Petter

    2012-01-01

    The major bark beetle threat to Norway spruce (Picea abies (L.) Karst.) in Eurasia is the spruce bark beetle Ips typographus. Beetles cause damage after population build-up in defenseless trees. To minimize attacks, timely removal of these trees is important. This is practiced by clearing of wind throws and sanitation felling. Thus, knowledge about the region-specific flight pattern and voltinism of I. typographus is necessary for efficient pest management. This thesis focuses on the ...

  2. Ash-forming elements in four Scandinavian wood species part 3: Combustion of five spruce samples

    Energy Technology Data Exchange (ETDEWEB)

    Werkelin, Johan; Lindberg, Daniel; Skrifvars, Bengt-Johan; Hupa, Mikko [Aabo Akademi Process Chemistry Centre, Piispankatu 8, FI-20500 Turku (Finland); Bostroem, Dan [Energy Technology and Thermal Process Chemistry, Umeaa University, SE-901 87 Umeaa (Sweden)

    2011-01-15

    Forest residue is the remaining fraction after the outtake of timber, which comprises the tree tops and branches. It may as fuel cause damage to the combustion device through ash slagging and fouling. The objective of this work was to model the ash composition from well-specified samples of a spruce tree: wood, bark, twigs, needles, and shoots. Their ash at 1000 C was modelled using global chemical equilibrium calculations, and laboratory-made ash of the five samples was analyzed by XRD and SEM-EDXA. According to the results, the risk of slagging arises from the spruce foliage: molten alkali silicates from spruce needles and probably molten alkali phosphates from spruce shoots may cause problems in the furnace. Fouling caused by condensing alkali vapours can be produced by all five samples. The amount of alkali vapours in the flue gas was in the same order of magnitude for all five samples, in spite of large differences in their original alkali contents. (author)

  3. A comparison of pine and spruce in recovery from winter stress; changes in recovery kinetics, and the abundance and phosphorylation status of photosynthetic proteins during winter.

    Science.gov (United States)

    Merry, Ryan; Jerrard, Jacob; Frebault, Julia; Verhoeven, Amy

    2017-09-01

    During winter evergreens maintain a sustained form of thermal energy dissipation that results in reduced photochemical efficiency measured using the chlorophyll fluorescence parameter Fv/Fm. Eastern white pine (Pinus strobus L.) and white spruce [Picea glauca (Moench) Voss] have been shown to differ in their rate of recovery of Fv/Fm from winter stress. The goal of this study was to monitor changes in photosynthetic protein abundance and phosphorylation status during winter recovery that accompany these functional changes. An additional goal was to determine whether light-dependent changes in light harvesting complex II (LHCII) phosphorylation occur during winter conditions. We used a combination of field measurements and recovery experiments to monitor chlorophyll fluorescence and photosynthetic protein content and phosphorylation status. We found that pine recovered three times more slowly than spruce, and that the kinetics of recovery in spruce included a rapid and slow component, while in pine there was only a rapid component to recovery. Both species retained relatively high amounts of the light harvesting protein Lhcb5 (CP26) and the PsbS protein during winter, suggesting a role for these proteins in sustained thermal dissipation. Both species maintained high phosphorylation of LHCII and the D1 protein in darkness during winter. Pine and spruce differed in the kinetics of the dephosphorylation of LHCII and D1 upon warming, suggesting the rate of dephosphorylation of LHCII and D1 may be important in the rapid component of recovery from winter stress. Finally, we demonstrated that light-dependent changes in LHII phosphorylation do not continue to occur on subzero winter days and that needles are maintained in a phosphorylation pattern consistent with the high light conditions to which those needles are exposed. Our results suggest a role for retained phosphorylation of both LHCII and D1 in maintenance of the photosynthetic machinery in a winter conformation

  4. Characterization of variable EST SSR markers for Norway spruce (Picea abies L.

    Directory of Open Access Journals (Sweden)

    Spiess Nadine

    2011-10-01

    Full Text Available Abstract Background Norway spruce is widely distributed across Europe and the predominant tree of the Alpine region. Fast growth and the fact that timber can be harvested cost-effectively in relatively young populations define its status as one of the economically most important tree species of Northern Europe. In this study, EST derived simple sequence repeat (SSR markers were developed for the assessment of putative functional diversity in Austrian Norway spruce stands. Results SSR sequences were identified by analyzing 14,022 publicly available EST sequences. Tri-nucleotide repeat motifs were most abundant in the data set followed by penta- and hexa-nucleotide repeats. Specific primer pairs were designed for sixty loci. Among these, 27 displayed polymorphism in a testing population of 16 P. abies individuals sampled across Austria and in an additional screening population of 96 P. abies individuals from two geographically distinct Austrian populations. Allele numbers per locus ranged from two to 17 with observed heterozygosity ranging from 0.075 to 0.99. Conclusions We have characterized variable EST SSR markers for Norway spruce detected in expressed genes. Due to their moderate to high degree of variability in the two tested screening populations, these newly developed SSR markers are well suited for the analysis of stress related functional variation present in Norway spruce populations.

  5. Relating structural growth environment to white spruce sapling establishment at the Forest-Tundra Ecotone

    Science.gov (United States)

    Maguire, A.; Boelman, N.; Griffin, K. L.; Jensen, J.; Hiers, E.; Johnson, D. M.; Vierling, L. A.; Eitel, J.

    2017-12-01

    The effect of climate change on treeline position at the latitudinal Forest-Tundra ecotone (FTE) is poorly understood. While the FTE is expansive (stretching 13,000 km acros the panarctic), understanding relationships between climate and tree function may depend on very fine scale processes. High resolution tools are therefore needed to appropriately characterize the leading (northernmost) edge of the FTE. We hypothesized that microstructural metrics obtainable from lidar remote sensing may explain variation in the physical growth environment that governs sapling establishment. To test our hypothesis, we used terrestrial laser scanning (TLS) to collect highly spatially resolved 3-D structural information of white spruce (Picea glauca) saplings and their aboveground growth environment at the leading edge of a FTE in northern Alaska and Northwest Territories, Canada. Coordinates of sapling locations were extracted from the 3-D TLS data. Within each sampling plot, 20 sets of coordinates were randomly selected from regions where no saplings were present. Ground roughness, canopy roughness, average aspect, average slope, average curvature, wind shelter index, and wetness indexwere extracted from point clouds within a variable radius from all coordinates. Generalized linear models (GLM) were fit to determine which microstructural metrics were most strongly associated with sapling establishment. Preliminary analyses of three plots suggest that vegetation roughness, wetness index, ground roughness, and slope were the most important terrain metrics governing sapling presence (Figure 1). Comprehensive analyses will include eight plots and GLMs optimized for scale at which structural parameters affect sapling establishment. Spatial autocorrelation of sample locations will be accounted for in models. Because these analyses address how the physical growth environment affects sapling establishment, model outputs will provide information for improving understanding of the

  6. Estimation of forest structural parameters using 5 and 10 meter SPOT-5 satellite data

    Science.gov (United States)

    Peter T. Wolter; Phillip A. Townsend; Brian R. Sturtevant

    2009-01-01

    Large areas of forest in the US and Canada are affected by insects and disease each year. Over the past century, outbreaks of the Eastern spruce budworm have become more frequent and severe. The notion of designing a more pest resistant landscape through prescriptive management practices hinges on our ability to effectively model forest?insect dynamics at regional...

  7. Formation of chloroform in spruce forest soil - results from laboratory incubation studies

    DEFF Research Database (Denmark)

    Haselmann, K.F.; Laturnus, F.; Svensmark, B.

    2000-01-01

    The release of chloroform, 1,1,1-trichloroethane, tetrachloromethane, trichloroethene and tetrachloroethene from an organic rich spruce forest soil was studied in laboratory incubation experiments by dynamic headspace analysis, thermodesorption and gas chromatography. Performance parameters...... are presented for the dynamic headspace system. For spruce forest soil, the results showed a significant increase in chloroform concentration in the headspace under aerobic conditions over a period of seven days, whereas the concentration of the other compounds remained fairly constant. A biogenic formation...

  8. Optimal uneven-aged stocking guides: an application to spruce-fir stands in New England

    Science.gov (United States)

    Jeffrey H. Gove; Mark J. Ducey

    2014-01-01

    Management guides for uneven-aged forest stands periodically need to be revisited and updated based on new information and methods. The current silvicultural guide for uneven-aged spruce-fir management in Maine and the northeast (Frank, R.M. and Bjorkbom, J.C. 1973 A silvicultural guide for spruce-fir in the northeast. General Technical Report NE-6, Forest Service. U.S...

  9. Natural and anthropogenic stress in spruce and beech ecosystems in the Solling project

    International Nuclear Information System (INIS)

    Murach, D.

    1989-01-01

    The investigations show that low Ca/Al-ratios in mineral soil horizons cause a shallow root system with spruce. But obviously other soil conditions expect acidification can also be responsible for a shallow root system. Additional investigations and field experiments have to find out the driving forces. A new phenomenon in fine root investigations are the low fine root concentrations in the uppermost mineral soil horizons. It is to be observed with spruce as well as with beech in stands with strong acidification of the top soil. Further experiments are needed to evaluate what are the main chemical variables (pH, heavy metals etc.) In this way soil acidification can cause reductions of fine root biomasses in subsoils as well as in uppermost soil horizons. These reductions can influence the response of trees to water stress. Relations to the symptoms of needle losses with spruce are likely. The close correlations between the chemistry of the soil, the roots and the needles indicate, that the yellowing symptoms of spruce trees, which were related to Mg-deficiency, are caused by Mg-deficiency in the soil. It is to be emphazised that it is not a pure Mg-deficiency. Ion antagonism to Al and especially to H in connection with fine root distribution are of main importance. (orig.)

  10. Carbon and Nitrogen Pools and Fluxes in Adjacent Mature Norway Spruce and European Beech Forests

    Directory of Open Access Journals (Sweden)

    Filip Oulehle

    2016-11-01

    Full Text Available We compared two adjacent mature forest ecosystem types (spruce vs. beech to unravel the fate of assimilated carbon (C and the cycling of organic and inorganic nitrogen (N without the risk of the confounding influences of climatic and site differences when comparing different sites. The stock of C in biomass was higher (258 t·ha−1 in the older (150 years beech stand compared to the younger (80 years planted spruce stand (192 t·ha−1, whereas N biomass pools were comparable (1450 kg·ha−1. Significantly higher C and N soil pools were measured in the beech stand, both in forest floor and mineral soil. Cumulative annual CO2 soil efflux was similar among stands, i.e., 9.87 t·ha−1·year−1 of C in the spruce stand and 9.01 t·ha−1·year−1 in the beech stand. Soil temperature explained 78% (Q10 = 3.7 and 72% (Q10 = 4.2 of variability in CO2 soil efflux in the spruce and beech stand, respectively. However, the rather tight N cycle in the spruce stand prevented inorganic N losses, whereas losses were higher in the beech stand and were dominated by nitrate in the mineral soil. Our results highlighted the long-term consequences of forest management on C and N cycling.

  11. Heat Resistance of Glued Finger Joints in Spruce Wood Constructions

    Directory of Open Access Journals (Sweden)

    Martin Sviták

    2014-10-01

    Full Text Available The heat resistance of glued spruce wood was evaluated for different joint types and adhesives. Bending strength, modulus of elasticity, and also fracture evaluation were investigated on glued spruce samples made by the finger-jointed principle. Finger-jointed samples were glued with polyurethane (PUR and melamine-urea-formaldehyde (MUF adhesives. Heat loading was realized at temperatures 60, 80, and 110 °C and compared with wood with 20 °C. A static bending test with four-point flexural test was used. Elevated temperature and adhesive type had an important influence on the bending strength. On the other hand, adhesive type had a significant influence on the modulus of elasticity, but elevated temperature had no substantial influence.

  12. Using maximum entropy modeling to identify and prioritize red spruce forest habitat in West Virginia

    Science.gov (United States)

    Nathan R. Beane; James S. Rentch; Thomas M. Schuler

    2013-01-01

    Red spruce forests in West Virginia are found in island-like distributions at high elevations and provide essential habitat for the endangered Cheat Mountain salamander and the recently delisted Virginia northern flying squirrel. Therefore, it is important to identify restoration priorities of red spruce forests. Maximum entropy modeling was used to identify areas of...

  13. Eocene to Miocene Out-of-Sequence Deformation in the Eastern Tibetan Plateau: Insights From Shortening Structures in the Sichuan Basin

    Science.gov (United States)

    Tian, Yuntao; Kohn, Barry P.; Qiu, Nansheng; Yuan, Yusong; Hu, Shengbiao; Gleadow, Andrew J. W.; Zhang, Peizhen

    2018-02-01

    A distinctive NNE trending belt of shortening structures dominates the topography and deformation of the eastern Sichuan Basin, 300 km east of the Tibetan Plateau. Debate continues as to whether the structures resulted from Cenozoic eastward growth of the Tibetan Plateau. A low-temperature thermochronology (AFT and AHe) data set from four deep boreholes and adjacent outcrops intersecting a branch of the shortening structures indicates distinctive differential cooling at 35-28 Ma across the structure, where stratigraphy has been offset vertically by 0.8-1.3 km. This result forms the first quantitative evidence for the existence of a late Eocene-Oligocene phase of shortening in the eastern Sichuan Basin, synchronous with the early phase of eastward growth and extrusion of the Tibetan Plateau. Further, a compilation of regional Cenozoic structures reveals a Miocene retreat of deformation from the foreland basin to the hinterland areas. Such a tectonic reorganization indicates that Eocene to Miocene deformation in the eastern Tibetan Plateau is out-of-sequence and was probably triggered by enhanced erosion in the eastern Tibetan Plateau.

  14. Climatic Sensitivity of a Mixed Forest Association of White Spruce and Trembling Aspen at Their Southern Range Limit

    Directory of Open Access Journals (Sweden)

    Sophan Chhin

    2016-10-01

    Full Text Available Climatic sensitivity of white spruce (Picea glauca (Moench Voss was examined growing in association with trembling aspen (Populus tremuloides Michx. at their southern limit of distribution in a transitional ecotone between the southern boreal forest and northern prairie region. The study was carried out in the Spruce Woods Provincial Park (SWPP located in southwestern Manitoba, Canada. The dry regional climate restricted trembling aspen growth during the growing season via moisture deficiency and temperature induced drought stress. Warm, mild winters also negatively affected radial growth of trembling aspen. Growth of white spruce was moderated by conditions within the aspen stands as radial growth patterns showed low variability from year to year, a low common growth signal, and a stronger response to temperature than to precipitation. Nonetheless, the dry regional climate still restricted growth of white spruce during the growing season via temperature induced drought stress. The findings of the study for white spruce support the stress gradient hypothesis in which facilitative interactions between tree species are expected under harsher environmental conditions.

  15. Development of epicormic sprouts in Sitka spruce following thinning and pruning in south-east Alaska.

    Science.gov (United States)

    Robert L. Deal; R. James Barbour; Michael H. McClellan; Dean L. Parry

    2003-01-01

    The frequency and size of epicormic sprouts in Sitka spruce (Picea sitchensis (Bong.) Carr.) were assessed in five 23-29 year-old mixed Sitka spruce-western hemlock (Tsuga heterophylla (Raf.) Sarg.) stands that were uniformly thinned and pruned to 2.4, 3.7 and 5.2 m lift heights. Six to nine years after treatment sprouts were...

  16. A Norway spruce FLOWERING LOCUS T homolog is implicated in control of growth rhythm in conifers.

    Science.gov (United States)

    Gyllenstrand, Niclas; Clapham, David; Källman, Thomas; Lagercrantz, Ulf

    2007-05-01

    Growth in perennial plants possesses an annual cycle of active growth and dormancy that is controlled by environmental factors, mainly photoperiod and temperature. In conifers and other nonangiosperm species, the molecular mechanisms behind these responses are currently unknown. In Norway spruce (Picea abies L. Karst.) seedlings, growth cessation and bud set are induced by short days and plants from southern latitudes require at least 7 to 10 h of darkness, whereas plants from northern latitudes need only 2 to 3 h of darkness. Bud burst, on the other hand, is almost exclusively controlled by temperature. To test the possible role of Norway spruce FLOWERING LOCUS T (FT)-like genes in growth rhythm, we have studied expression patterns of four Norway spruce FT family genes in two populations with a divergent bud set response under various photoperiodic conditions. Our data show a significant and tight correlation between growth rhythm (both bud set and bud burst), and expression pattern of one of the four Norway spruce phosphatidylethanolamine-binding protein gene family members (PaFT4) over a variety of experimental conditions. This study strongly suggests that one Norway spruce homolog to the FT gene, which controls flowering in angiosperms, is also a key integrator of photoperiodic and thermal signals in the control of growth rhythms in gymnosperms. The data also indicate that the divergent adaptive bud set responses of northern and southern Norway spruce populations, both to photoperiod and light quality, are mediated through PaFT4. These results provide a major advance in our understanding of the molecular control of a major adaptive trait in conifers and a tool for further molecular studies of adaptive variation in plants.

  17. Forest insects and diseases in Kouchibouguac National Park in 1994. Technical note No. 306

    Energy Technology Data Exchange (ETDEWEB)

    Lemieux, G.R.

    1995-11-01

    Personnel of the Forest Insect and Disease Survey regularly survey national parks for forest insect and disease conditions. This document presents some of the conditions encountered in Kouchibouguac National Park in 1992, including balsam twig aphids, gypsy moth, whitespotted sawyer bettle, white pine weevil, frost damage, Eastern tent caterpiller, uglynest caterpillar, hypoxylon canker, spruce budmoth, Eastern spruce gall adelgid, and other pests encountered.

  18. Forest insects and diseases in Kouchibouguac National Park in 1992. Technical note No. 275. Annual publication

    Energy Technology Data Exchange (ETDEWEB)

    Lemieux, G.R.

    1993-01-01

    Personnel of the Forest Insect and Disease Survey regularly survey national parks for forest insect and disease conditions. This document presents some of the conditions encountered in Kouchibouguac National Park in 1992, including balsam twig aphids, gypsy moth, whitespotted sawyer bettle, white pine weevil, frost damage, Eastern tent caterpiller, uglynest caterpillar, hypoxylon canker, spruce budmoth, Eastern spruce gall adelgid, and other pests encountered.

  19. Forest insects and diseases in Kouchibouguac National Park in 1993. Technical note No. 295

    Energy Technology Data Exchange (ETDEWEB)

    Lemieux, G.R.

    1994-01-01

    Personnel of the Forest Insect and Disease Survey regularly survey national parks for forest insect and disease conditions. This document presents some of the conditions encountered in Kouchibouguac National Park in 1992, including balsam twig aphids, gypsy moth, whitespotted sawyer bettle, white pine weevil, frost damage, Eastern tent caterpiller, uglynest caterpillar, hypoxylon canker, spruce budmoth, Eastern spruce gall adelgid, and other pests encountered.

  20. Spruce Lake Dam reconstruction

    Energy Technology Data Exchange (ETDEWEB)

    Snyder, G. [SGE Acres Ltd., Fredericton, NB (Canada); Barnard, J. [SGE Acres Ltd., St. John' s, NF (Canada); Vriezen, C. [City of Saint John, NF (Canada); Stephenson, M. [Jacques Whitford Environment Ltd., Fredericton, NB (Canada)

    2004-09-01

    Spruce Lake Dam was constructed in 1898 as part of the water supply system for Saint John, New Brunswick. The original dam was a 6 meter high, 140 meter long concrete gravity dam with an intake structure at its mid point and an overflow spillway at the left abutment. A rehabilitation project was launched in 2001 to bring the deteriorated dam into conformance with the dam safety guidelines of the Canadian Dam Association. The project criteria included minimal disruption to normal operation of water supply facilities and no negative effect on water quality. The project involved installation of a new low level outlet, removal of a gate house and water intake pipes, replacement of an access road culvert in the spillway channel, and raising the earth dam section by 1.8 meters to allow for increased water storage. The new raised section has an impervious core. The project also involved site and geotechnical investigations as well as hydrotechnical and environmental studies. This presentation described the final design of the remedial work and the environmental permitting procedures. Raising the operating level of the system proved successful as demonstrated by the fewer number of pumping days required after dam rehabilitation. The dam safety assessment under the Canadian Environmental Assessment Act began in April 2001, and the rehabilitation was completed by the end of 2002. 1 tab., 8 figs.

  1. Warming drives a front of white spruce establishment near western treeline, Alaska.

    Science.gov (United States)

    Miller, Amy E; Wilson, Tammy L; Sherriff, Rosemary L; Walton, James

    2017-12-01

    Regional warming has led to increased productivity near the boreal forest margin in Alaska. To date, the effects of warming on seedling recruitment have received little attention, in spite of forecasted forest expansion. Here, we used stand structure and environmental data from 95 white spruce (Picea glauca) plots sampled across a longitudinal gradient in southwest Alaska to explore factors influencing spruce establishment and recruitment near western treeline. We used total counts of live seedlings, saplings, and trees, representing five life stages, to evaluate whether geospatial, climate, and measured plot covariates predicted abundance, using current abundance distributions as a surrogate for climate conditions in the past. We used generalized linear models to test the null hypothesis that conditions favorable for recruitment were similar along the environmental gradient represented by longitude, by exploring relationships between per-plot counts of each life stage and the covariates hypothesized to affect abundance. We also examined the relationship between growing degree days (GDD) and seedling establishment over a period of three decades using tree-ring chronologies obtained from cores taken at a subset of our sites (n = 30). Our results indicated that seedling, sapling, and tree abundance were positively correlated with temperature across the study area. The response to longitude was mixed, with earlier life stages (seedlings, saplings) most abundant at the western end of the gradient, and later life stages (trees) most abundant to the east. The differential relationship between longitude and life-stage abundance suggests a moving front of white spruce establishment through time, driven by changes in environmental conditions near the species' western range limit. Likewise, we found a positive relationship between periods of seedling establishment and GDD, suggesting that longer summers and/or greater heat accumulation might enhance establishment

  2. Fire Severity Controlled Susceptibility to a 1940s Spruce Beetle Outbreak in Colorado, USA.

    Science.gov (United States)

    Kulakowski, Dominik; Veblen, Thomas T; Bebi, Peter

    2016-01-01

    The frequency, magnitude, and size of forest disturbances are increasing globally. Much recent research has focused on how the occurrence of one disturbance may affect susceptibility to subsequent disturbances. While much has been learned about such linked disturbances, the strength of the interactions is likely to be contingent on the severity of disturbances as well as climatic conditions, both of which can affect disturbance intensity and tree resistance to disturbances. Subalpine forests in western Colorado were affected by extensive and severe wildfires in the late 19th century and an extensive and severe outbreak of spruce beetle (Dendroctonus rufipennis) in the 1940s. Previous research found that most, but not all, of the stands that burned and established following the late 19th century fires were not susceptible to the 1940s outbreak as beetles preferentially attack larger trees and stands in advanced stages of development. However, previous research also left open the possibility that some stands that burned and established following the 19th century fires may have been attacked during the 1940s outbreak. Understanding how strongly stand structure, as shaped by disturbances of varying severity, affected susceptibility to past outbreaks is important to provide a baseline for assessing the degree to which recent climate change may be relaxing the preferences of beetles for larger trees and for stands in latter stages of structural development and thereby changing the nature of linked disturbances. Here, dendroecological methods were used to study disturbance history and tree age of stands in the White River National Forest in Western Colorado that were identified in historical documents or remotely-sensed images as having burned in the 19th century and having been attacked by spruce beetle in the 1940s. Dendroecological reconstructions indicate that in young post-fire stands only old remnant trees that survived the otherwise stand-replacing fires were

  3. Plasticity in variation of xylem and phloem cell characteristics of Norway spruce under different local conditions

    Directory of Open Access Journals (Sweden)

    Jozica eGricar

    2015-09-01

    Full Text Available There is limited information on intra-annual plasticity of secondary tissues of tree species growing under different environmental conditions. To increase the knowledge about the plasticity of secondary growth, which allows trees to adapt to specific local climatic regimes, we examined climate–radial growth relationships of Norway spruce (Picea abies (L. H. Karst. from three contrasting locations in the temperate climatic zone by analyzing tree-ring widths for the period 1932–2010, and cell characteristics in xylem and phloem increments formed in the years 2009–2011. Variation in the structure of xylem and phloem increments clearly shows that plasticity in seasonal dynamics of cambial cell production and cell differentiation exists on xylem and phloem sides. Anatomical characteristics of xylem and phloem cells are predominantly site-specific characteristics, because they varied among sites but were fairly uniform among years in trees from the same site. Xylem and phloem tissues formed in the first part of the growing season seemed to be more stable in structure, indicating their priority over latewood and late phloem for tree performance. Long-term climate and radial growth analyses revealed that growth was in general less dependent on precipitation than on temperature; however, growth sensitivity to local conditions differed among the sites. Only partial dependence of radial growth of spruce on climatic factors on the selected sites confirms its strategy to adapt the structure of wood and phloem increments to function optimally in local conditions.

  4. Shallow crustal structure of eastern-central Trans-Mexican Volcanic Belt.

    Science.gov (United States)

    Campos-Enriquez, J. O.; Ramón, V. M.; Lermo-Samaniego, J.

    2015-12-01

    Central-eastern Trans-Mexican Volcanic Belt (TMVB) is featured by large basins (i.e., Toluca, Mexico, Puebla-Tlaxcala, Libres-Oriental). It has been supposed that major crustal faults limit these basins. Sierra de Las Cruces range separates the Toluca and Mexico basins. The Sierra Nevada range separates Mexico basin from the Puebla-Tlaxcala basin. Based in gravity and seismic data we inferred the Toluca basin is constituted by the Ixtlahuaca sub-basin, to the north, and the Toluca sub-basin to the south, which are separated by a relative structural high. The Toluca depression is more symmetric and bounded by sub-vertical faults. In particular its eastern master fault controlled the emplacement of Sierra de Las Cruces range. Easternmost Acambay graben constitutes the northern and deepest part of the Ixtlahuaca depression. The Toluca-Ixtlahuaca basin is inside the Taxco-San Miguel de Allende fault system, and limited to the west by the Guerrero terrane which continues beneath the TMVB up to the Acambay graben. Mexico basin basement occupies an intermediate position and featured by a relative structural high to the north-east, as established by previous studies. This relative structural high is limited to the west by the north-south Mixhuca trough, while to the south it is bounded by the east-west Copilco-Xochimilco-Chalco sub-basin. The Puebla-Tlaxcala basin basement is the shallowest of these 3 tectonic depressions. In general, features (i.e., depth) and relationship between these basins, from west to east, are controlled by the regional behavior of the Sierra Madre Oriental fold and thrust belt basement (i.e., Oaxaca Complex?). This study indicates that an active east-west regional fault system limits to the south the TMVB (from the Nevado de Toluca volcano through the Popocatepetl volcano and eastward along southern Puebla-Tlaxcala basin). The Tenango and La Pera fault systems constituting the western part of this regional fault system coincide with northern

  5. Comparison of different ground techniques to map leaf area index of Norway spruce forest canopy

    OpenAIRE

    Homolova, L.; Malenovsky, Z.; Hanus, J.; Tomaskova, I.; Dvoráková, M.; Pokorny, R.

    2007-01-01

    The leaf area index (LAI) of three monocultures of Norway spruce (Picea abies (L.) Karst), different in age and structure, was measured by means of two indirect optical techniques of LAI field mapping: 1/ plant canopy analyser LAI-2000, and 2/ digital hemispherical photographs (DHP). The supportive measurements with the TRAC instrument were conducted to produce mainly the element clumping index. The aim of the study was to compare the performances of LAI-2000 and DHP and to evaluate effect of...

  6. Eastern rim of the Chesapeake Bay impact crater: Morphology, stratigraphy, and structure

    Science.gov (United States)

    Poag, C.W.

    2005-01-01

    This study reexamines seven reprocessed (increased vertical exaggeration) seismic reflection profiles that cross the eastern rim of the Chesapeake Bay impact crater. The eastern rim is expressed as an arcuate ridge that borders the crater in a fashion typical of the "raised" rim documented in many well preserved complex impact craters. The inner boundary of the eastern rim (rim wall) is formed by a series of raterfacing, steep scarps, 15-60 m high. In combination, these rim-wall scarps represent the footwalls of a system of crater-encircling normal faults, which are downthrown toward the crater. Outboard of the rim wall are several additional normal-fault blocks, whose bounding faults trend approximately parallel to the rim wall. The tops of the outboard fault blocks form two distinct, parallel, flat or gently sloping, terraces. The innermost terrace (Terrace 1) can be identified on each profile, but Terrace 2 is only sporadically present. The terraced fault blocks are composed mainly of nonmarine, poorly to moderately consolidated, siliciclastic sediments, belonging to the Lower Cretaceous Potomac Formation. Though the ridge-forming geometry of the eastern rim gives the appearance of a raised compressional feature, no compelling evidence of compressive forces is evident in the profiles studied. The structural mode, instead, is that of extension, with the clear dominance of normal faulting as the extensional mechanism. 

  7. Drought-triggered western spruce budworm outbreaks in the Interior Pacific Northwest: A multi-century dendrochronological record

    Science.gov (United States)

    A. Flower; D. G. Gavin; E. K. Heyerdahl; R. A. Parsons; G. M. Cohn

    2014-01-01

    Douglas-fir forests in the interior Pacific Northwest are subject to sporadic outbreaks of the western spruce budworm, a species widely recognized as the most destructive defoliator in western North America. Outbreaks of the western spruce budworm often occur synchronously over broad regions and lead to widespread loss of leaf area and decrease in growth rates in...

  8. Middle Eastern rhinoplasty.

    Science.gov (United States)

    Azizzadeh, Babak; Mashkevich, Grigoriy

    2010-02-01

    The ethnic appearance of the Middle Eastern nose is defined by several unique visual features, particularly a high radix, wide overprojecting dorsum, and an amorphous hanging nasal tip. These external characteristics reflect distinct structural properties of the osseo-cartilaginous nasal framework and skin-soft tissue envelope in patients of Middle Eastern extraction. The goal, and the ultimate challenge, of rhinoplasty on Middle Eastern patients is to achieve balanced aesthetic refinement, while avoiding surgical westernization. Detailed understanding of the ethnic visual harmony in a Middle Eastern nose greatly assists in preserving native nasal-facial relationships during rhinoplasty on Middle Eastern patients. Esthetic alteration of a Middle Eastern nose follows a different set of goals and principles compared with rhinoplasties on white or other ethnic patients. This article highlights the inherent nasal features of the Middle Eastern nose and reviews pertinent concepts of rhinoplasty on Middle Eastern patients. Essential considerations in the process spanning the consultation and surgery are reviewed. Reliable operative techniques that achieve a successful aesthetic outcome are discussed in detail. Copyright 2010 Elsevier Inc. All rights reserved.

  9. Pathogenicity of Neonectria fuckeliana on Norway Spruce Clones in Sweden and Potential Management Strategies

    OpenAIRE

    Martin Pettersson; Venche Talgø; John Frampton; Bo Karlsson; Jonas Rönnberg

    2018-01-01

    The fungus Neonectria fuckeliana has become an increasing problem on Norway spruce (Picea abies) in the Nordic countries during recent years. Canker wounds caused by the pathogen reduce timber quality and top-dieback is a problem for the Christmas tree industry. In this study, four inoculation trials were conducted to examine the ability of N. fuckeliana to cause disease on young Norway spruce plants and determine how different wound types would affect the occurrence and severity of the disea...

  10. PEI detoxification of pretreated spruce for high solids ethanol fermentation

    DEFF Research Database (Denmark)

    Cannella, David; Sveding, Per Viktor; Jørgensen, Henning

    2014-01-01

    .e. spruce) this has been difficult to reach. The main reason behind this difference is the higher recalcitrance of woody substrates which require harsher pretreatment conditions, thus generating higher amounts of inhibitory compounds, ultimately lowering fermentation performances. In this work we studied...... ethanol production from spruce performing the whole process, from pretreatment to hydrolysis and fermentation, at 30% dry matter (equivalent to similar to 20% WIS). Hydrolysis and fermentation was performed in a horizontal free fall mixing reactor enabling efficient mixing at high solids loadings....... In batch simultaneous saccharification and fermentation (SSF), up to 76% cellulose to ethanol conversion was achieved resulting in a concentration of 51 g/kg of ethanol. Key to obtaining this high ethanol yield at these conditions was the use of a detoxification technology based on applying a soluble...

  11. Spruce budworm core B.t. test - 1980 combined summary

    Science.gov (United States)

    Gerald S. Walton; Franklin B. Lewis

    1982-01-01

    Two commercial preparations of Bacillus thuringiensis (Bet.) were aerially applied in 1980 to populations of spruce budworm (Choristoneura fumiferana (Clem.)) in Arizona, Maine, New Hampshire, and Wisconsin. Operations were conducted under the auspices of CANUSA-East with standardized procedures for spray application and population...

  12. Applicability of the PROSPECT model for Norway spruce needles

    NARCIS (Netherlands)

    Malenovsky, Z.; Albrechtova, J.; Lhotakova, Z.; Zurita Milla, R.; Clevers, J.G.P.W.; Schaepman, M.E.; Cudlin, P.

    2006-01-01

    The potential applicability of the leaf radiative transfer model PROSPECT (version 3.01) was tested for Norway spruce (Picea abies (L.) Karst.) needles collected from stress resistant and resilient trees. Direct comparison of the measured and simulated leaf optical properties between 450¿1000 nm

  13. The institutional structure and political economy of food distribution systems: A comparative analysis of six Eastern European countries

    DEFF Research Database (Denmark)

    Esbjerg, Lars; Skytte, Hans

    This paper discusses the food distribution systems of six Eastern European countries. It considers the macro and task environments of distribution systems, discussing the constraints and opportunities the environments present to companies. The institutional structure of retailing and wholesaling...... are analysed and important developments in the institutional structure are noted. The internal political economy of distribution channels in Eastern Europe is analysed and the modernisation of distribution systems discussed. Finally, some conclusions are offered and areas for future research suggested....

  14. Damages and causes of death in plantations with containerised seedlings of Scots pine and Norway spruce in the central of Sweden

    Energy Technology Data Exchange (ETDEWEB)

    Naumburg, Jan

    2000-07-01

    In 1972, 94 forest areas were planted with containerised seedlings, 83 with Scots pine (Pinus sylvestris L.) and 11 with Norway spruce (Picea abies (L.) Karst.), in the central of Sweden. In the first season after planting, 99% of the Scots pine and 98% of the Norway spruce seedlings survived. Three seasons after plantation, 67% of Scots pine and 62% of Norway spruce were alive. The most common type of known damages causing mortality were mammals and insects. Vegetation was registered as the cause of mortality at some occasions in Scots pine plantations, whereas vegetation never was considered as the cause of death in Norway spruce plantations. The average size of the scarification patches were 0.25 m{sup 2} and 0.4 m{sup 2} in Scots pine and Norway spruce respectively. In Scots pine plantations there were 1600 planted seedlings ha{sup -1} and in Norway spruce there were 1550 ha{sup -1}. After the third growing season, the numbers of main crop plants, including naturally regenerated hardwood and softwood plants, were 1500 ha{sup -1} for Scots pine and 1350 ha{sup -1} for Norway spruce. The studied plantings had been approved if the recommended number of seedlings had been planted. As there always is some mortality among planted seedlings, in the present study 35-40%, this phenomenon has to be taken into consideration when dimensioning the number of seedlings which are to be planted.

  15. Structure and resilience of fungal communities in Alaskan boreal forest soils

    Science.gov (United States)

    D. Lee Taylor; Ian C. Herriott; Kelsie E. Stone; Jack W. McFarland; Michael G. Booth; Mary Beth Leigh

    2010-01-01

    This paper outlines molecular analyses of soil fungi within the Bonanza Creek Long Term Ecological Research program. We examined community structure in three studies in mixed upland, black spruce (Picea mariana (Mill.) BSP), and white spruce (Picea glauca (Moench) Voss) forests and examined taxa involved in cellulose...

  16. Photolysis of polycyclic aromatic hydrocarbons adsorbed on spruce [Picea abies (L.) Karst.] needles under sunlight irradiation

    International Nuclear Information System (INIS)

    Niu Junfeng; Chen Jingwen; Martens, D.; Quan Xie; Yang Fenglin; Kettrup, A.; Schramm, K.-W.

    2003-01-01

    Photolysis of PAHs on surfaces may determine their ultimate fate in the environment. - Photolysis of polycyclic aromatic hydrocarbons (PAHs) sorbed on surfaces of spruce [Picea abies (L.) Karst.] needles under sunlight irradiation was investigated. PAHs were produced by combustion of polyvinyl chloride (PVC), wood, high-density polyethylene (HDPE), and styrene in a stove. The factors of sunlight irradiation on the surfaces of spruce needles were taken into consideration when investigating the kinetic parameters. The photolysis of the 18 PAHs under study follows first-order kinetics. The photolysis half-lives range from 15 h for dibenzo(a,h)anthracene to 75 h for phenanthrene. Photolysis of some PAHs on surfaces of spruce needles may play an important role on the fate of PAHs in the environment

  17. Establishment and growth of white spruce on a boreal forest floodplain: interactions between microclimate and mammalian herbivory

    Science.gov (United States)

    Amy C. Angell; Knut. Kielland

    2009-01-01

    White spruce (Picea glauca (Moench) Voss) is a dominant species in late-successional ecosystems along the Tanana River, interior Alaska, and the most important commercial timber species in these boreal floodplain forests. Whereas white spruce commonly seed in on young terraces in early primary succession, the species does not become a conspicuous...

  18. Assembly of the Complete Sitka Spruce Chloroplast Genome Using 10X Genomics' GemCode Sequencing Data.

    Directory of Open Access Journals (Sweden)

    Lauren Coombe

    Full Text Available The linked read sequencing library preparation platform by 10X Genomics produces barcoded sequencing libraries, which are subsequently sequenced using the Illumina short read sequencing technology. In this new approach, long fragments of DNA are partitioned into separate micro-reactions, where the same index sequence is incorporated into each of the sequencing fragment inserts derived from a given long fragment. In this study, we exploited this property by using reads from index sequences associated with a large number of reads, to assemble the chloroplast genome of the Sitka spruce tree (Picea sitchensis. Here we report on the first Sitka spruce chloroplast genome assembled exclusively from P. sitchensis genomic libraries prepared using the 10X Genomics protocol. We show that the resulting 124,049 base pair long genome shares high sequence similarity with the related white spruce and Norway spruce chloroplast genomes, but diverges substantially from a previously published P. sitchensis- P. thunbergii chimeric genome. The use of reads from high-frequency indices enabled separation of the nuclear genome reads from that of the chloroplast, which resulted in the simplification of the de Bruijn graphs used at the various stages of assembly.

  19. Determination of the moisture content of Nordic spruce wood through cone heater experiments and an integral model

    DEFF Research Database (Denmark)

    Mindykowski, Pierrick Anthony; Jørgensen, M.; Svensson, Staffan

    2015-01-01

    The combination of cone heater experiments and an integral model was used to determine the moisture content of Nordic spruce with varying degree of drying. Nine specimens of Nordic spruce were pre-heated to 105°C in a convective oven for durations ranging from 0 days (no drying) and up to 63 days...

  20. Structural changes in Central and Eastern European economies: breaking news or breaking the ice?

    Czech Academy of Sciences Publication Activity Database

    Égert, B.; Jiménez-Rodríguez, R.; Kočenda, Evžen; Morales-Zumaquero, A.

    2006-01-01

    Roč. 39, 1-2 (2006), s. 85-103 ISSN 1573-9414 R&D Projects: GA MŠk LC542 Institutional research plan: CEZ:MSM0021620846 Keywords : Central and Eastern European (CEE) countries * multiple structural breaks * volatility Subject RIV: AH - Economics

  1. Community Structure Of Reef Fish In Eastern Luwu Water Territory

    Directory of Open Access Journals (Sweden)

    Henny Tribuana Cinnawara

    2015-01-01

    Full Text Available Abstract One bio-indicators the condition of coral reefs is a presence of reef fish. The purpose of research is to determine species composition abundance distribution and structure of reef fish communities in these waters. Data collection was conducted in April at six locations in the north and the south eastern Luwu. Mechanical Underwater Visual Cencus UVC and transect method Line intercept Transec LIT with SCUBA equipment used for research data collection. Total reef fish species collected as many as 366 species belonging to 31 families consisting of 150 species of fish target fish consumption 10 species of indicator fish indicator species 206 types of major fissh. The most dominant indicator type of fish is Chaetodon octofasciatus while the major dominant family Pomacentridae Labridae and Apogonidae. Diversity index values ranged from 2.145 to 3.408. Dominance index C is in the range of 0.056 to 0.298. The result is expected to be a reference literature as basic data for the management of reef fish especially in the waters of eastern Luwu.

  2. A Norway Spruce FLOWERING LOCUS T Homolog Is Implicated in Control of Growth Rhythm in Conifers1[OA

    Science.gov (United States)

    Gyllenstrand, Niclas; Clapham, David; Källman, Thomas; Lagercrantz, Ulf

    2007-01-01

    Growth in perennial plants possesses an annual cycle of active growth and dormancy that is controlled by environmental factors, mainly photoperiod and temperature. In conifers and other nonangiosperm species, the molecular mechanisms behind these responses are currently unknown. In Norway spruce (Picea abies L. Karst.) seedlings, growth cessation and bud set are induced by short days and plants from southern latitudes require at least 7 to 10 h of darkness, whereas plants from northern latitudes need only 2 to 3 h of darkness. Bud burst, on the other hand, is almost exclusively controlled by temperature. To test the possible role of Norway spruce FLOWERING LOCUS T (FT)-like genes in growth rhythm, we have studied expression patterns of four Norway spruce FT family genes in two populations with a divergent bud set response under various photoperiodic conditions. Our data show a significant and tight correlation between growth rhythm (both bud set and bud burst), and expression pattern of one of the four Norway spruce phosphatidylethanolamine-binding protein gene family members (PaFT4) over a variety of experimental conditions. This study strongly suggests that one Norway spruce homolog to the FT gene, which controls flowering in angiosperms, is also a key integrator of photoperiodic and thermal signals in the control of growth rhythms in gymnosperms. The data also indicate that the divergent adaptive bud set responses of northern and southern Norway spruce populations, both to photoperiod and light quality, are mediated through PaFT4. These results provide a major advance in our understanding of the molecular control of a major adaptive trait in conifers and a tool for further molecular studies of adaptive variation in plants. PMID:17369429

  3. Competition modifies effects of enhanced ozone/carbon dioxide concentrations on carbohydrate and biomass accumulation in juvenile Norway spruces and European beech

    International Nuclear Information System (INIS)

    Liu, X.; Rennenberg, H.; Kozovits, A. R.; Grams, T. E.; Blaschke, H.; Matyssek, R.

    2004-01-01

    Potential interactions of carbon dioxide and ozone on carbohydrate concentrations and contents were studied in Norway spruce and European beech saplings to test the hypotheses that (1) prolonged exposure to elevated carbon dioxide does not compensate for the limiting effects of ozone on the accumulation of sugars and starches, or biomass partitioning to the root; and (2) growth of mixed-species planting will repress plant responses to elevated ozone and carbon dioxide. Norway spruce and European beech saplings were acclimated for one year to ambient and elevated carbon dioxide, followed by exposure to factorial combinations of ambient and elevated ozone and carbon dioxide during the next two years. In spruce trees, sugar and starch content was greater in saplings exposed to elevated carbon dioxide; in beech, the response was the opposite. The overall conclusion was that the results did not support Hypothesis One, because the adverse effects were counteracted by elevated carbon dioxide. Regarding Hypothesis Two, it was found to be supportive for beech but not for spruce. In beech, the reduction of sugars and starch by elevated ozone and stimulation by elevated carbon dioxide were repressed by competitive interaction with spruce, whereas in spruce saplings elevated concentrations of carbon dioxide resulted in higher concentrations of sugar and starch, but only in leaves and coarse roots and only when grown in combination with beech. Elevated ozone in spruce saplings produced no significant effect on sugar or starch content either in intra- or interspecific competition. 57 refs., 1 tab., 5 figs

  4. Competition modifies effects of enhanced ozone/carbon dioxide concentrations on carbohydrate and biomass accumulation in juvenile Norway spruces and European beech

    Energy Technology Data Exchange (ETDEWEB)

    Liu, X.; Rennenberg, H. [University of Freiburg, Inst. of Forest Botany and Tree Physiology, Freiburg (Germany); Kozovits, A. R.; Grams, T. E.; Blaschke, H.; Matyssek, R. [Technische Universitat Munchen, Dept. of Ecology and Ecophysiology of Plants, Freising (Germany)

    2004-09-01

    Potential interactions of carbon dioxide and ozone on carbohydrate concentrations and contents were studied in Norway spruce and European beech saplings to test the hypotheses that (1) prolonged exposure to elevated carbon dioxide does not compensate for the limiting effects of ozone on the accumulation of sugars and starches, or biomass partitioning to the root; and (2) growth of mixed-species planting will repress plant responses to elevated ozone and carbon dioxide. Norway spruce and European beech saplings were acclimated for one year to ambient and elevated carbon dioxide, followed by exposure to factorial combinations of ambient and elevated ozone and carbon dioxide during the next two years. In spruce trees, sugar and starch content was greater in saplings exposed to elevated carbon dioxide; in beech, the response was the opposite. The overall conclusion was that the results did not support Hypothesis One, because the adverse effects were counteracted by elevated carbon dioxide. Regarding Hypothesis Two, it was found to be supportive for beech but not for spruce. In beech, the reduction of sugars and starch by elevated ozone and stimulation by elevated carbon dioxide were repressed by competitive interaction with spruce, whereas in spruce saplings elevated concentrations of carbon dioxide resulted in higher concentrations of sugar and starch, but only in leaves and coarse roots and only when grown in combination with beech. Elevated ozone in spruce saplings produced no significant effect on sugar or starch content either in intra- or interspecific competition. 57 refs., 1 tab., 5 figs.

  5. Proportion of various dendromass components of spruce (Picea abies), and partial models for modification of wind speed and radiation by pure spruce stands

    International Nuclear Information System (INIS)

    Wollmerstädt, J.; Sharma, S.C.; Marsch, M.

    1992-01-01

    Means for quantifying dendromass components of spruce stands have been discussed, and partial models for modification of radiation and wind by the pure spruce stand were developed. By means of a sampling procedure, the components needle dry mass and branchwood dry mass without needles of individual trees are recorded. Using the relationship between branch basal diameter and needle respectively branchwood dry mass, the total needle and branchwood dry mass of trees is estimated. Based on that, stand or regional parameters for the allometric function between diameter breast height and needle respectively branchwood dry mass can be determined for defined H/D-clusters. Published data from various sources were used in this paper. The lowest coefficients of determination were found in H/D-cluster 120 (H/D-values over 114). Therefore, further differentiation within this range seems to be necessary. For assimilation models, there should be quantification of needle dry mass separately for needle age classes and morphological characteristics of needles. Basis for the estimate of tree-bole volume is the relationship between H/D-value and oven-dry weight. There are problems as far as methods for quantifying the subterranean dendromass (e.g. dynamics of fine roots) are concerned; this is requiring considerable efforts, too. Spatial structure was also described by allometric functions (crown length and crown cover in relation to diameter breast height). For the partial model to express wind modification by the stand, standardized wind profiles as related to crown canopy density were used. The modification of radiation by the stand is closely related with the vertical needle mass distribution (sum curves). These two partial models have to be considered as an approach for the description of the modifying effect by the stocking [de

  6. Spruce colonization at treeline: where do those seeds come from?

    Science.gov (United States)

    Piotti, A; Leonardi, S; Piovani, P; Scalfi, M; Menozzi, P

    2009-08-01

    At treeline, selection by harsh environmental conditions sets an upward limit to arboreal vegetation. Increasing temperatures and the decline of traditional animal raising have favoured an upward shift of treeline in the last decades. These circumstances create a unique opportunity to study the balance of the main forces (selection and gene flow) that drive tree migration. We conducted a parentage analysis sampling and genotyping with five microsatellite markers in all Norway spruce individuals (342 juveniles and 23 adults) found in a recently colonized treeline area (Paneveggio forest, Eastern Alps, Italy). Our goal was to evaluate local reproductive success versus gene flow from the outside. We were able to identify both parents among local adults for only 11.1% of the juveniles. In the gamete pool we sampled, two-thirds were not produced locally. Effective seed dispersal distance distribution was characterized by a peak far from the seed source (mean 344.66 m+/-191.02 s.d.). Reproductive success was skewed, with six local adults that generated almost two-thirds (62.4%) of juveniles with local parents. Our findings indicate that, although a few local adults seem to play an important role in the colonization process at treeline, large levels of gene flow from outside were maintained, suggesting that the potential advantages of local adults (such as local adaptation, proximity to the colonization area, phenological synchrony) did not prevent a large gamete immigration.

  7. Experiences with the Haertel-Truebungstest at diagnosing injuries to spruce caused by atmospheric pollution

    Energy Technology Data Exchange (ETDEWEB)

    Pelz, E

    1958-01-01

    The Haertel-Truebungstest is based on the observation that spruce needles secrete more wax at the stomata in areas affected by atmospheric pollution than in areas without. The increased secretion of wax is taken for a symptom of injury caused by atmospheric pollution. The wax is extracted in boiling water and, after cooling down, the turbidity of the extract is measured quantitatively in a photometer. Investigations on spruce in Saxony and Thuringia confirm the tendency to increasing turbidity towards the source of smoke. A considerable dispersion of turbidity values however impairs the results of this method, so that a great number of samples must be used in order to find out a trend. The dispersion of turbidity values in spruce forests that are not directly influenced by atmospheric pollution substantially exceeds the dispersion of the +/- 1% of the turbidity value that was found by Haertel. As was stated by Haertel and Papesch, the dispersion of individual turbidity values grows with increasing doses of SO/sub 2/. In Saxony the individual smoke emissions so densely cover one another, that there are scarcely any areas without air pollution and, probably, a relation exists between this continuous influence of smoke and SO/sub 2/ and the dispersion of turbidity values. It may be, however, that the environmental factors have also a certain influence, as, on vast areas of Saxony, spruce is growing on sites that are not its natural habitat.

  8. Carbon sources in vertical profile of Norway spruce stand

    Czech Academy of Sciences Publication Activity Database

    Pavelka, Marian; Janouš, Dalibor; Urban, Otmar; Acosta, Manuel; Pokorný, Radek; Havránková, Kateřina; Formanek, P.

    2003-01-01

    Roč. 2, č. 30 (2003), s. 199-206 ISSN 1336-5266 R&D Projects: GA MŠk(CZ) LN00A141; GA ČR(CZ) GA526/03/1021 Institutional research plan: CEZ:AV0Z6087904 Keywords : Carbon stock * respiration * Norway spruce Subject RIV: EH - Ecology, Behaviour

  9. Evidence That Drought-Induced Stomatal Closure Is Not an Important Constraint on White Spruce Performance Near the Arctic Treeline in Alaska

    Science.gov (United States)

    Sullivan, P.; Brownlee, A.; Ellison, S.; Sveinbjornsson, B.

    2014-12-01

    Tree cores collected from trees growing at high latitudes have long been used to reconstruct past climates, because of close positive correlations between temperature and tree growth. However, in recent decades and at many sites, these relationships have deteriorated and have even become negative in some instances. The observation of declining tree growth in response to rising temperature has prompted many investigators to suggest that high latitude trees may be increasingly exhibiting drought-induced stomatal closure. In the Brooks Range of northern Alaska, the observation of low and declining growth of white spruce is more prevalent in the central and eastern parts of the range, where precipitation is lower, providing superficial support for the drought stress hypothesis. In this study, we investigated the occurrence of white spruce drought-induced stomatal closure in four watersheds along a west to east gradient near the Arctic treeline in the Brooks Range. We obtained a historical perspective on tree growth and water relations by collecting increment cores for analysis of ring widths and carbon isotopes in tree-ring alpha-cellulose. Meanwhile, we made detailed assessments of contemporary water relations at the scales of the whole canopy and the needle. All of our data indicate that drought-induced stomatal closure is probably not responsible for low and declining growth in the central and eastern Brooks Range. Carbon isotope discrimination has generally increased over the past century and our calculations indicate that needle inter-cellular CO2 concentration is much greater now than it was in the early 1900's. Measurements of needle gas exchange are consistent with the tree core record, in the sense that instances of low photosynthesis at our sites are not coincident with similarly low stomatal conductance and low inter-cellular CO2 concentration. Finally, hourly measurements of xylem sap flow indicate that trees at our study sites are able to maintain near

  10. Influence of road salting on the adjacent Norway spruce (Picea abies) forest

    International Nuclear Information System (INIS)

    Forczek, S.T.; Benada, O.; Kofronova, O.; Sigler, K.; Matucha, M.

    2011-01-01

    Winter deicing and traffic spreads salt to road-adjacent Norway spruce trees in the form of spraying and salt slops. Our use of Na36Cl revealed roots as the main pathway of salt uptake. One-shot application of a concentrated Na36Cl solution to spruce saplings by both irrigation and spraying causes macroscopic damage to the needles and affects the needle phyllosphere. Irrigation affects the trees more than spraying because Cl uptake through roots is faster and eventually leads to higher chloride content in the plant. Along with the root-needle route, spray-deposited chloride from the needles is re-transported back into the soil and again taken up by roots to needles

  11. SPECIES DIVERSITY AND COMMUNITY STRUCTURE OF FRUIT-FEEDING BUTTERFLIES (LEPIDOPTERA: NYMPHALIDAE IN AN EASTERN AMAZONIAN FOREST

    Directory of Open Access Journals (Sweden)

    LUCAS PEREIRA MARTINS

    Full Text Available ABSTRACT Deforestation has negative impacts on diversity and community patterns of several taxa. In the eastern Amazon, where much deforestation is predicted for the coming years, forests patches may be essential to maintain the local biodiversity. Despite increasing concerns about the conservation of threatened areas, few studies have been performed to analyze the communities of diversified groups, such as insects, in the eastern Amazon. Here, we investigated species diversity and community structure of fruit-feeding butterflies, a well-known bioindicator group, in a threatened remnant of an eastern Amazonian forest located on Maranhão Island, northeastern Brazil. Fruit-feeding butterflies were sampled monthly for one year. Diversity and evenness indices, richness estimators, rarefaction curve, and rank-abundance plot were used to describe community structure in the study area. We captured 529 fruit-feeding butterflies in four subfamilies, 23 genera and 34 species. The three most abundant species, Hamadryas februa, Hamadryas feronia, and Hermeuptychia cf. atalanta are indicators of disturbed habitats and represented more than half of the collected individuals. Richness estimators revealed that between 87 and 94% of the fruit-feeding butterfly species were sampled, suggesting few additional records would be made for the area. Our results indicate that human-caused disturbances have altered local community patterns and provide baseline data for future research in threatened regions of the eastern Amazon.

  12. Vulnerability of white spruce tree growth in interior Alaska in response to climate variability: dendrochronological, demographic, and experimental perspectives

    Science.gov (United States)

    A.D. McGuire; R.W. Ruess; A. Lloyd; J. Yarie; J.S. Clein; G.P. Juday

    2010-01-01

    This paper integrates dendrochronological, demographic, and experimental perspectives to improve understanding of the response of white spruce (Picea glauca (Moench) Voss) tree growth to climatic variability in interior Alaska. The dendrochronological analyses indicate that climate warming has led to widespread declines in white spruce growth...

  13. Solar radiation as a factor influencing the raid spruce bark beetle (Ips typographus) during spring swarming

    International Nuclear Information System (INIS)

    Mezei, P.

    2011-01-01

    Monitoring of spruce bark beetle in nature reserve Fabova hola Mountain in the Slovenske Rudohorie Mountains at an altitude of 1.100-1.440 meters was conducted from 2006 to 2009. Slovenske Rudohorie Mountains was affected by two windstorms (2004 and 2007) followed by a gradation of bark beetles. This article has examined the dependence between amount of solar radiation and trapping of spruce bark beetle into pheromone traps.

  14. Fire Severity Controlled Susceptibility to a 1940s Spruce Beetle Outbreak in Colorado, USA.

    Directory of Open Access Journals (Sweden)

    Dominik Kulakowski

    Full Text Available The frequency, magnitude, and size of forest disturbances are increasing globally. Much recent research has focused on how the occurrence of one disturbance may affect susceptibility to subsequent disturbances. While much has been learned about such linked disturbances, the strength of the interactions is likely to be contingent on the severity of disturbances as well as climatic conditions, both of which can affect disturbance intensity and tree resistance to disturbances. Subalpine forests in western Colorado were affected by extensive and severe wildfires in the late 19th century and an extensive and severe outbreak of spruce beetle (Dendroctonus rufipennis in the 1940s. Previous research found that most, but not all, of the stands that burned and established following the late 19th century fires were not susceptible to the 1940s outbreak as beetles preferentially attack larger trees and stands in advanced stages of development. However, previous research also left open the possibility that some stands that burned and established following the 19th century fires may have been attacked during the 1940s outbreak. Understanding how strongly stand structure, as shaped by disturbances of varying severity, affected susceptibility to past outbreaks is important to provide a baseline for assessing the degree to which recent climate change may be relaxing the preferences of beetles for larger trees and for stands in latter stages of structural development and thereby changing the nature of linked disturbances. Here, dendroecological methods were used to study disturbance history and tree age of stands in the White River National Forest in Western Colorado that were identified in historical documents or remotely-sensed images as having burned in the 19th century and having been attacked by spruce beetle in the 1940s. Dendroecological reconstructions indicate that in young post-fire stands only old remnant trees that survived the otherwise stand

  15. Extraction of antioxidants from spruce (Picea abies) bark using eco-friendly solvents.

    Science.gov (United States)

    Co, Michelle; Fagerlund, Amelie; Engman, Lars; Sunnerheim, Kerstin; Sjöberg, Per J R; Turner, Charlotta

    2012-01-01

    Antioxidants are known to avert oxidation processes and they are found in trees and other plant materials. Tree bark is a major waste product from paper pulp industries; hence it is worthwhile to develop an extraction technique to extract the antioxidants. To develop a fast and environmentally sustainable extraction technique for the extraction of antioxidants from bark of spruce (Picea abies) and also to identify the extracted antioxidants that are abundant in spruce bark. A screening experiment that involved three different techniques was conducted to determine the best technique to extract antioxidants. The antioxidant capacity of the extracts was determined with DPPH (2,2-diphenyl-1-picrylhydrazyl) assay. Pressurised fluid extraction (PFE) turned out to be the best technique and a response surface design was therefore utilised to optimise PFE. Furthermore, NMR and HPLC-DAD-MS/MS were applied to identify the extracted antioxidants. PFE using water and ethanol as solvent at 160 and 180°C, respectively, gave extracts of the highest antioxidant capacity. Stilbene glucosides such as isorhapontin, piceid and astringin were identified in the extracts. The study has shown that PFE is a fast and environmentally sustainable technique, using water and ethanol as solvent for the extraction of antioxidants from spruce bark. Copyright © 2011 John Wiley & Sons, Ltd.

  16. Non-spray spruce budworm management alternatives: Will thinning second-growth balsam fir in northwestern Ontario alter the stands` vulnerability to spruce budworm damage. NWOFTDU technical report No. 53

    Energy Technology Data Exchange (ETDEWEB)

    Van Schip, W.M.; Towill, W.D.; Zago, T.

    1990-12-31

    The objective of this study was to determine whether balsam fir stands, thinned to different levels of basal area, differ in their susceptibility and vulnerability to damage created by infestations of spruce budworm. In 1976-77, the investigators thinned two relatively pure, juvenile, second-growth balsam fir stands in northwestern Ontario to four levels of basal area by girdling or felling. The stands were assessed in 1988. Annual aerial and ground monitoring of the study sites indicated that one area experienced a moderate to severe defoliation by spruce budworm during 1984-86, while the other area experienced moderate to severe defoliation by budworm during 1980-86. This report presents results of the study and discusses the relation between thinning, budworm damage, and stand productivity.

  17. Crown structure, radiation absorption, photosynthesis and transpiration

    OpenAIRE

    Wang, Yingping

    1988-01-01

    A complex simulation model, MAESTRO, has been developed and validated against field measurements in plantation in both Scotland and Australia. It has been shown that MAESTRO can reasonably predict the daily course of PAR (photosynetically active radiation) transmittance at points below the canopies of radiata pine and Sitka spruce plantations. 1. Four structural properties of the Sitka spruce tree crown have been identified and evaluation in relation to PAR absorption, photosynthesis and ...

  18. Soil surface CO2 fluxes in a Norway spruce stand

    Czech Academy of Sciences Publication Activity Database

    Acosta, Manuel; Janouš, Dalibor; Marek, Michal V.

    2004-01-01

    Roč. 12, č. 50 (2004), s. 573-578 ISSN 1212-4834 R&D Projects: GA AV ČR(CZ) KJB3087301 Institutional research plan: CEZ:AV0Z6087904 Keywords : Norway spruce * Soil CO2 efflux * Q10 Subject RIV: EH - Ecology, Behaviour

  19. Component biomass equations for black spruce in Maine

    Science.gov (United States)

    M. M. Czapowskyj; D. J. Robison; R. D. Briggs; E. H. White; E. H. White

    1985-01-01

    Component biomass prediction equations are presented for young black spruce (Picea mariana B.S.P. (Mill,:)) in northern Maine. A weighted least squares model was used to construct the eq~iationsfo r small trees from 1 to 15 cm d.b.h., and an ordinary least squares model for trees less than 2 m in height. A linearized allometric model was also tested but was not used....

  20. Determination of the terpene flux from orange species and Norway spruce by relaxed eddy accumulation

    DEFF Research Database (Denmark)

    Christensen, C.S.; Hummelshøj, P.; Jensen, N.O.

    2000-01-01

    Terpene fluxes from a Norway spruce (Picea abies) forest and an orange orchard (Citrus clementii and Citrus sinensis) were measured by relaxed eddy accumulation (REA) during summer 1997. alpha-pinene and beta-pinene were the most abundant terpenes emitted from Norway spruce and constituted approx...... rate by using two precision pumps operated at approximately 60 mi min(-1). The terpenes collected on the adsorbent tubes were significantly decomposed by ozone during sampling unless ozone scrubbers were applied. (C) 2000 Elsevier Science Ltd. All rights reserved....

  1. Controls on moss evaporation in a boreal black spruce forest

    NARCIS (Netherlands)

    Heijmans, M.M.P.D.; Arp, W.J.; Chapin, F.S.

    2004-01-01

    [1] Mosses are an important component of the boreal forest, but little is known about their contribution to ecosystem carbon, water, and energy exchange. We studied the role of mosses in boreal forest evapotranspiration by conducting two experiments in a black spruce forest in Fairbanks, Alaska.

  2. Animal damage to young spruce and fir in Maine

    Science.gov (United States)

    Barton M. Blum

    1977-01-01

    The loss of terminal buds on small balsam fir (Abies balsamea (L.) Mill.) and spruce (Picea spp.) trees because of nipping by mammals or birds has increased on the Penobscot Experimental Forest in recent years. The cut stem is smooth and slightly angled; there is no sign of tearing. Unnipped trees grew about 13 percent more than...

  3. Estimation of arboreal lichen biomass available to woodland caribou in Hudson Bay lowland black spruce sites

    Directory of Open Access Journals (Sweden)

    Sarah K. Proceviat

    2003-04-01

    Full Text Available An arboreal lichen index to be utilized in assessing woodland caribou habitat throughout northeastern Ontario was developed. The "index" was comprised of 5 classes, which differentiated arboreal lichen biomass on black spruce trees, ranging from maximal quantities of arboreal lichen (class 5 to minimal amounts of arboreal lichen (class 1. This arboreal lichen index was subsequently used to estimate the biomass of arboreal lichen available to woodland caribou on lowland black spruce sites ranging in age from 1 year to 150 years post-harvest. A total of 39 sites were assessed and significant differences in arboreal lichen biomass were found, with a positive linear relationship between arboreal lichen biomass and forest age. It is proposed that the index be utilized by government and industry as a means of assessing the suitability of lowland black spruce habitat for woodland caribou in this region.

  4. Raman imaging to investigate ultrastructure and composition of plant cell walls : distribution of lignin and cellulose in black spruce wood (Picea mariana)

    Science.gov (United States)

    Umesh P. Agarwal

    2006-01-01

    A detailed understanding of the structural organization of the cell wall of vascular plants is important from both the perspectives of plant biology and chemistry and of commercial utilization. A state-of-the-art 633-nm laser-based confocal Raman microscope was used to determine the distribution of cell wall components in the cross section of black spruce wood in situ...

  5. A 4-year record of sitka spruce and western hemlock seed fall on the Cascade Head Experimental Forest.

    Science.gov (United States)

    Robert H. Ruth; Carl M. Berntsen

    1955-01-01

    Four years' measurement of seed fall in the spruce-hemlock type on the Cascade Head Experimental Forest indicates that an ample supply of seed is distributed over clear-cut areas under staggered-setting cutting. The largest tract sampled was 81 acres; in spite of a seed crop failure in 1950, it received an average of 243,000 viable spruce and hemlock seeds per...

  6. Conifer R2R3-MYB transcription factors: sequence analyses and gene expression in wood-forming tissues of white spruce (Picea glauca

    Directory of Open Access Journals (Sweden)

    Grima-Pettenati Jacqueline

    2007-03-01

    Full Text Available Abstract Background Several members of the R2R3-MYB family of transcription factors act as regulators of lignin and phenylpropanoid metabolism during wood formation in angiosperm and gymnosperm plants. The angiosperm Arabidopsis has over one hundred R2R3-MYBs genes; however, only a few members of this family have been discovered in gymnosperms. Results We isolated and characterised full-length cDNAs encoding R2R3-MYB genes from the gymnosperms white spruce, Picea glauca (13 sequences, and loblolly pine, Pinus taeda L. (five sequences. Sequence similarities and phylogenetic analyses placed the spruce and pine sequences in diverse subgroups of the large R2R3-MYB family, although several of the sequences clustered closely together. We searched the highly variable C-terminal region of diverse plant MYBs for conserved amino acid sequences and identified 20 motifs in the spruce MYBs, nine of which have not previously been reported and three of which are specific to conifers. The number and length of the introns in spruce MYB genes varied significantly, but their positions were well conserved relative to angiosperm MYB genes. Quantitative RTPCR of MYB genes transcript abundance in root and stem tissues revealed diverse expression patterns; three MYB genes were preferentially expressed in secondary xylem, whereas others were preferentially expressed in phloem or were ubiquitous. The MYB genes expressed in xylem, and three others, were up-regulated in the compression wood of leaning trees within 76 hours of induction. Conclusion Our survey of 18 conifer R2R3-MYB genes clearly showed a gene family structure similar to that of Arabidopsis. Three of the sequences are likely to play a role in lignin metabolism and/or wood formation in gymnosperm trees, including a close homolog of the loblolly pine PtMYB4, shown to regulate lignin biosynthesis in transgenic tobacco.

  7. Mountain Norway spruce forests: Needle supply and its nutrient content

    Czech Academy of Sciences Publication Activity Database

    Kovářová, Marcela; Vacek, S.

    2003-01-01

    Roč. 49, - (2003), s. 327-332 ISSN 1212-4834 R&D Projects: GA AV ČR KSK6005114; GA ČR GA206/99/1416 Institutional research plan: CEZ:AV0Z6005908 Keywords : Šumava Mts. * Mountain Norway spruce forest * needle mass Subject RIV: EF - Botanics

  8. Wood-thermoplastic composites manufactured using beetle-killed spruce from Alaska

    Science.gov (United States)

    V. Yadama; Eini Lowell; N. Petersen; D. Nicholls

    2009-01-01

    The primary objectives of the study were to characterize the critical properties of wood flour produced using highly deteriorated beetle-killed spruce for wood-plastic composite (WPC) production and evaluate important mechanical and physical properties of WPC extruded using an industry standard formulation. Chemical composition analysis indicated no significant...

  9. The variability of wood density and compression strength of Norway spruce

    Czech Academy of Sciences Publication Activity Database

    Horáček, Petr; Fajstavr, Marek; Stojanović, Marko

    2017-01-01

    Roč. 10, 1-2 (2017), s. 17-26 ISSN 1803-2451 R&D Projects: GA MŠk(CZ) LO1415 Institutional support: RVO:86652079 Keywords : Norway spruce * wood density * compression strength * variability Subject RIV: GK - Forestry OBOR OECD: Forestry

  10. Distinct genecological patterns in seedlings of Norway spruce and silver fir from a mountainous landscape.

    Science.gov (United States)

    Frank, Aline; Sperisen, Christoph; Howe, Glenn Thomas; Brang, Peter; Walthert, Lorenz; St Clair, John Bradley; Heiri, Caroline

    2017-01-01

    Understanding the genecology of forest trees is critical for gene conservation, for predicting the effects of climate change and climate change adaptation, and for successful reforestation. Although common genecological patterns have emerged, species-specific details are also important. Which species are most vulnerable to climate change? Which are the most important adaptive traits and environmental drivers of natural selection? Even though species have been classified as adaptive specialists vs. adaptive generalists, large-scale studies comparing different species in the same experiment are rare. We studied the genecology of Norway spruce (Picea abies) and silver fir (Abies alba), two co-occurring but ecologically distinct European conifers in Central Europe. For each species, we collected seed from more than 90 populations across Switzerland, established a seedling common-garden test, and developed genecological models that associate population variation in seedling growth and phenology to climate, soil properties, and site water balance. Population differentiation and associations between seedling traits and environmental variables were much stronger for Norway spruce than for silver fir, and stronger for seedling height growth than for bud phenology. In Norway spruce, height growth and second flushing were strongly associated with temperature and elevation, with seedlings from the lowlands being taller and more prone to second flush than seedlings from the Alps. In silver fir, height growth was more weakly associated with temperature and elevation, but also associated with water availability. Soil characteristics explained little population variation in both species. We conclude that Norway spruce has become an adaptive specialist because trade-offs between rapid juvenile growth and frost avoidance have subjected it to strong diversifying natural selection based on temperature. In contrast, because silver fir has a more conservative growth habit, it has

  11. The immigration and spread of spruce forest in Norway, traced by biostratigraphical studies and radiocarbon datings. A preliminary report

    International Nuclear Information System (INIS)

    Hafsten, U.

    1985-01-01

    Pollen-analytic studies and radiocarbon datings from 86 sites, mostly ombrotrophic peatbogs, situated within the Norwegian spruce domain, show that the occupation of the areas by spruce forest was the result of a protracted spread from cast, or northeast, to west and south, which started in late pre-Christian time and was completed mainly during the Middle Ages

  12. Cadmium and zinc toxicity in white pine, red maple, and Norway spruce

    Energy Technology Data Exchange (ETDEWEB)

    Mitchell, C.D.; Fretz, T.A.

    1977-01-01

    Excessive accumulation of Cd or Zn in white pine (Pinus strobus L.), red maple (Acer rubrum L.), and Norway spruce (Picea abies L.), resulted in reduced root initiation, poor development of root laterals, chlorosis, dwarfism, early leaf drop, wilting, and necrosis of current season's growth. Positive correlations existed between nutrient culture levels and tissue accumulation when plants were grown in sand. White pine seddlings accumulated more Cd and Zn than either red maple or Norway spurce when compared in similar experiments. White pine appeared to be more tolerant of Cd and Zn accumulation than either red maple or Norway spruce since visual phytotoxicity symptoms were observed only at the higher: 1 treatment levels. Accumulation of Cd and Zn from a medium of 2 sand:1 soil:1 perlite by volume was also observed.

  13. Problems in interpreting effects of air pollutants on spruce epicuticular waxes

    International Nuclear Information System (INIS)

    Bermadinger-Stabentheiner, E.

    1994-01-01

    Spruce needles are covered with rod-like crystals, which also fill the antechambers of the stomata with a dense meshwork. The scanning electron microscope (SEM) is very useful for studying epicuticular wax structure; with no intricate or laborious preparation, it is possible to obtain valuable information about the needle surface. Because the epicuticular wax layer forms a barrier between the plant and its environment, all influences that reach the surface from outside impact on this layer and, therefore, changes in epicuticular wax structure serve as diagnostic criteria for damage caused by air pollutants. This pollution influence begins as fusion of wax rods at the tips and results finally in total loss of the crystalline structure. Despite the simplicity of SEM investigations, alterations (artefacts) can occur to wax structures that may be confused with alterations caused by air pollutants (i.e., a too dense layer of twigs and needles, or careless handling with tweezers, results in mechanical damage that often influences the entire surface). Overheating occurring during transport or preparation and/or incorrect storage also produce artefacts. If the occurrence of such artefacts is taken into consideration, several contradictory interpretations of effects of air pollutants on epicuticular waxes can be explained. (orig.)

  14. Isoprenoid emission variation of Norway spruce across a European latitudinal transect

    Science.gov (United States)

    van Meeningen, Ylva; Wang, Min; Karlsson, Tomas; Seifert, Ana; Schurgers, Guy; Rinnan, Riikka; Holst, Thomas

    2017-12-01

    Norway spruce (Picea abies) is one of the dominant tree species in the European boreal zone with the capacity to grow over large areas within Europe. It is an important emitter of biogenic volatile organic compounds (BVOCs), which can act as precursors of photochemical smog and ozone and contribute to the formation and growth of secondary organic aerosols (SOA) in the atmosphere. Isoprenoid emissions were measured from Norway spruce trees at seven different sites, distributed from Ljubljana in Slovenia to Piikkiö in Finland. Four of the sites were part of a network of genetically identical spruce trees and contained two separate provenances. The remaining three sites were part of other networks which have been used to conduct studies in the European boreal zone. There were minimal differences in the standardized emission rates between sites and across latitudes. The emission profile differed between provenances and sites, but there were not any distinct patterns which could be connected to a change in latitude. By using genetically identical trees and comparing the emission rates between sites and with genetically different trees, it was observed that the emission patterns were mostly influenced by genetics. But in order to confirm this possible stability of the relative emission profile based on genetics, more studies need to be performed. The effects of branch height, season and variation between years on observed emission pattern variations were also investigated. There were indications of potential influences of all three factors. However, due to different experimental setups between measurement campaigns, it is difficult to draw any robust conclusions.

  15. Effects of soil calcium and aluminum on the physiology of balsam fir and red spruce saplings in northern New England

    Science.gov (United States)

    Richard L. Boyce; Paul G. Schaberg; Gary J. Hawley; Joshua M. Halman; Paula F. Murakami

    2013-01-01

    We examined the influence of calcium (Ca) and aluminum (Al) nutrition on the foliar physiology of red spruce (Picea rubens Sarg.) and balsam fir [Abies balsamea (L.) Mill.] in northern New England, USA. At the Hubbard Brook Experimental Forest (NH, USA), spruce and fir saplings were sampled from control, Al-, and Ca-supplemented...

  16. Quantifying Boreal Forest Structure and Composition Using UAV Structure from Motion

    Directory of Open Access Journals (Sweden)

    Michael Alonzo

    2018-03-01

    Full Text Available The vast extent and inaccessibility of boreal forest ecosystems are barriers to routine monitoring of forest structure and composition. In this research, we bridge the scale gap between intensive but sparse plot measurements and extensive remote sensing studies by collecting forest inventory variables at the plot scale using an unmanned aerial vehicle (UAV and a structure from motion (SfM approach. At 20 Forest Inventory and Analysis (FIA subplots in interior Alaska, we acquired overlapping imagery and generated dense, 3D, RGB (red, green, blue point clouds. We used these data to model forest type at the individual crown scale as well as subplot-scale tree density (TD, basal area (BA, and aboveground biomass (AGB. We achieved 85% cross-validation accuracy for five species at the crown level. Classification accuracy was maximized using three variables representing crown height, form, and color. Consistent with previous UAV-based studies, SfM point cloud data generated robust models of TD (r2 = 0.91, BA (r2 = 0.79, and AGB (r2 = 0.92, using a mix of plot- and crown-scale information. Precise estimation of TD required either segment counts or species information to differentiate black spruce from mixed white spruce plots. The accuracy of species-specific estimates of TD, BA, and AGB at the plot scale was somewhat variable, ranging from accurate estimates of black spruce TD (+/−1% and aspen BA (−2% to misallocation of aspen AGB (+118% and white spruce AGB (−50%. These results convey the potential utility of SfM data for forest type discrimination in FIA plots and the remaining challenges to develop classification approaches for species-specific estimates at the plot scale that are more robust to segmentation error.

  17. The Social Consequences of Postcommunist Structural Change: An Analysis of Suicide Trends in Eastern Europe

    Science.gov (United States)

    Minagawa, Yuka

    2013-01-01

    Guided by Durkheim's classic theory of suicide, this article examines suicide trends and determinants in Eastern European countries for the period of 1989-2006, with particular attention given to the association between postcommunist social change and suicide mortality. I find that countries characterized by more drastic structural change…

  18. Death of spruce needles due to air-borne ash

    Energy Technology Data Exchange (ETDEWEB)

    Maran, B

    1959-01-01

    This paper discusses the damage caused by the deposition of ash, with a high content of sulfur dioxide, on spruce trees. The data in this paper covers the source of the pollution, the effects of weather on the transport of the pollution, and the type of damage caused by the pollution. Other types of trees included in the data are pine, larch, and 5 broadleaf species.

  19. Downward Migration of Coastal Conifers as a Response to Recent Land Emergence in Eastern Hudson Bay, Québec

    Science.gov (United States)

    Bégin, Yves; Bérubé, Dominique; Grégoire, Martin

    1993-07-01

    Postglacial uplift in the eastern Hudson Bay area is among the most rapid in the world (300 m during the last 8000 yr). Although emergence curves based on 14 C-dated raised shorelines give a consistent basis for modeling relative sea-level changes, such a low-resolution dating method is inappropriate for estimating trends over recent decades. A major downward displacement of white spruce ( Picea glauca (Moench) Voss) and tamarack ( Larix laricina (DuRoi) K. Koch) occurred on protected shores as a response to shoreline retreat during this century. Analysis of the age distribution of trees indicates a progradation of white spruce and tamarack on gently sloping terrain ranging from 1.3 and 2.6 cm/yr, respectively, toward the sea. Improvement of climatic conditions during the 20th century favored such expansion which was probably faster than the real land emergence rates, but recent episodes of high water levels caused regression of forest margins over the highly exposed shores. Nevertheless, the downward trend of the treeline over this century substantiates the projections of 14C-dated coastal emergence curves during the modern period (1.0 to 1.3 cm/yr) by providing an estimate of the maximum rates of shoreline retreat.

  20. Development of a comprehensive analytical method for the determination of chlorinated paraffins in spruce needles applied in passive air sampling

    International Nuclear Information System (INIS)

    Iozza, Saverio; Schmid, Peter; Oehme, Michael

    2009-01-01

    Conifer needles are used for the monitoring of atmospheric persistent organic pollutants. The objective of the present study was to develop a method for the detection of airborne chlorinated paraffins (CPs) using spruce needles as a passive sampler. The method is based on liquid extraction of the cuticular wax layer followed by chromatographic fractionation and detection of CPs using two different GCMS techniques. Total CP concentrations (sum of short (SCCP), medium (MCCP) and long chain CPs (LCCP)) were determined by EI-MS/MS. SCCP and MCCP levels as well as congener group patterns (n-alkane chain length, chlorine content) could be evaluated using ECNI-LRMS. For the first time, data on environmental airborne CPs on spruce needles taken within the Monitoring Network in the Alpine Region for Persistent and other Organic Pollutants (MONARPOP) are presented providing evidence that spruce needles are a suitable passive sampling system for the monitoring of atmospheric CPs. - A developed method for chlorinated paraffins (CPs) provided evidence that spruce needles are a suitable passive sampling system for the monitoring of atmospheric CPs.

  1. Biochemical diagnosis of spruce trees in regions polluted by smoke

    Energy Technology Data Exchange (ETDEWEB)

    Cvrkal, H

    1959-01-01

    The material showing different degrees of damage caused by gases in smoke was sampled from 24 trees (spruces) in two smoke polluted regions - the Svatonice and Krusne Hory regions of Czechoslovakia. The essential oils were distilled out in the usual manner and chromatographed. The following terpenes were investigated: santene, ..cap alpha..-pinene, camphene, ..beta..-pinene, ..beta..-phelandrene, substance A, which could not be identified. The following relationships were determined on the basis of the compiled per cent content tables and the degree of damage caused by gases in smoke: the highest degree of damage occurs in spruce trees in whose essential oils camphene is represented in minimal amounts; smoke damages are not observed when the dipentene content is increased even at the cost of a lower camphene content; the degree of damage is also influenced to some extent by the presence of ..beta..-pinene. A higher degree of damage caused by gases in smoke presupposes a high ..beta..-pinene content in essential oils. Results suggest that the terpene changes do not occur during the damage, but are the consequence of specific species characteristics.

  2. Experimental Study Regarding the Freezing and Thawing Dynamics of Spruce Wood

    Directory of Open Access Journals (Sweden)

    Maria - Bernadett SZMUTKU

    2013-03-01

    Full Text Available The article presents the results regarding theevolution of the temperature field in spruce wood(Picea abies L. during freezing at two differentfreezing rates: -100C/h (rapid freezing and -10C/h(slow freezing and then during thawing at +50C,+30°C, +50°C temperature.This approach aimed at simulating thetemperature variations which occur inside timberduring open air storage in winter in two situations(sudden vs. gradual drop of temperature, and thenwhen the timber enters the drying kiln, depending onthe temperature applied in the initial heating phase.The results clearly show that the freezing ratesignificantly influences the thawing time and speed,which increase by 13-17% in the case of slowly frozenwood (at -10C/min compared to rapidly frozen wood(at -100C/min. It was also established that theoptimum temperature in the heating-up phase whendrying frozen spruce is 300C instead of the usual500C. This value leads to much better dryinguniformity without significantly prolonging the dryingtime.

  3. A study by non-isothermal thermal methods of spruce wood bark materialss after their application for dye removal

    Directory of Open Access Journals (Sweden)

    VIORICA DULMAN

    2005-11-01

    Full Text Available This paper deals with a study of some materials obtained from spruce bark (Picea abies, Romania, after retention of some dyes frequently used in dyeing processes in the textile industry and waste water treatment. These materials obtained by dye retention exhibit a particular thermal behavior which is different from that of the blank sample (spruce bark. The characteristic temperatures, weight losses, the residue remaining after thermo-oxidative degradation, as well as the activation energies of the significant thermo-destruction stages, estimated from non-isothermal thermogravimetric data, together with the thermal quantities calculated from DTAdata support the conclusion presented in a previous study on dye retention from aqueous solution. The obtained results made evident that, under optimal retention conditions, spruce bark shows the highest retention capacity for the Basic Blue dye, followed by Direct Brown 95 and Direct Brown 2.

  4. TopoGreenland: crustal structure in central-eastern Greenland along a new refraction profile

    Science.gov (United States)

    Shulgin, Alexey; Thybo, Hans; Field Team TopoGreenland

    2013-04-01

    We present the seismic structure in the interior of Greenland based on the first measurements by the seismic refraction/wide angle reflection method. Previous seismic surveys have only been carried out offshore and near the coast of Greenland, where the crustal structure is affected by oceanic break-up and may not be representative of the interior of the island. Acquisition of geophysical data in onshore Greenland is logistically complicated by the presence of an up to 3.4 km thick ice sheet, permanently covering most of the land mass. The seismic data was acquired by a team of six people during a two-month long experiment in summer of 2011 on the ice cap in the interior of central-eastern Greenland. The EW-trending profile extends 310 km inland from the approximate edge of the stable ice cap near Scoresby Sund across the center of the ice cap. The planned extension of the profile by use of OBSs and air gun shooting in Scoresbysund Fjord to the east coast of Greenland was unfortunately canceled, because navigation was prevented by ice drift. 350 Reftek Texan receivers recorded high-quality seismic data from 8 equidistant shots along the profile. Explosive charge sizes were 1 ton at the ends and ca. 500 kg along the profile, loaded with about 125 kg at 35-85 m depth in individual boreholes. Two-dimensional velocity model based on tomographic inversion and forward ray tracing modeling shows a decrease of crustal thickness from 47 km below the center of Greenland in the western part to 40 km in the eastern part of the profile. Earlier studies show that crustal thickness further decreases eastward to ca. 30 km below the fjord system, but details of the changes are unknown. Relatively high lower crustal velocities (Vp 6.8 - 7.3) in the western part of the TopoGreenland profile may indicate past collision tectonics or may be related or to the passage of the Iceland mantle plume. The origin of the pronounced circum-Atlantic mountain ranges in Norway and eastern Greenland

  5. Testing two novel stump-lifting heads in a final felling Norway spruce stand

    Energy Technology Data Exchange (ETDEWEB)

    Kaerhae, K. (Metsaeteho Oy, Helsinki (Finland)), Email: kalle.karha@metsateho.fi; Mutikainen, A. (TTS Research, Rajamaeki (Finland)), Email: arto.mutikainen@tts.fi

    2009-07-01

    The use of stump and root wood chips has increased very rapidly in the 21st century in Finland: in the year 2000, the total consumption of stump wood chips for energy generation was 10 GWh, while in 2008 it was around 1.2 TWh. Metsaeteho Oy and TTS Research tested two new stump-lifting devices for lifting stumps in a final felling Norway spruce (picea abies) stand. In the time study with the Vaekevae Stump Processor lifting head, the productivity of stump lifting was 7,5 m3 / E{sub 0}-hour when lifting spruce stumps with a diameter of 30 cm from clayey soil, and 8.3 m3 /E{sub 0}-hour when lifting spruce stumps from sandy soil. When lifting stumps with a diameter of 40 cm, the stump-lifting productivity was 9.0 m3 /E{sub 0}-h (clay) and 10,5 m3 / E{sub 0}-h (sand). The results of this relatively restricted test indicated that the Vaekevae Stump Processor is s reliable and effective stump-lifting head that enables the harvesting of high-quality stump raw material for energy generation. The stump lifting productivity of the other lifting head (Jaervinen) was lower than that of the Vaekevae Strump Processor. Some development suggestions for the Jaervinen lifting head were presented and discussed. (orig.)

  6. Ozone flux over a Norway spruce forest and correlation with net ecosystem production

    International Nuclear Information System (INIS)

    Zapletal, Milos; Cudlin, Pavel; Chroust, Petr; Urban, Otmar; Pokorny, Radek; Edwards-Jonasova, Magda; Czerny, Radek; Janous, Dalibor; Taufarova, Klara; Vecera, Zbynek; Mikuska, Pavel; Paoletti, Elena

    2011-01-01

    Daily ozone deposition flux to a Norway spruce forest in Czech Republic was measured using the gradient method in July and August 2008. Results were in good agreement with a deposition flux model. The mean daily stomatal uptake of ozone was around 47% of total deposition. Average deposition velocity was 0.39 cm s -1 and 0.36 cm s -1 by the gradient method and the deposition model, respectively. Measured and modelled non-stomatal uptake was around 0.2 cm s -1 . In addition, net ecosystem production (NEP) was measured by using Eddy Covariance and correlations with O 3 concentrations at 15 m a.g.l., total deposition and stomatal uptake were tested. Total deposition and stomatal uptake of ozone significantly decreased NEP, especially by high intensities of solar radiation. - Highlights: → We estimate ozone deposition flux to a Norway spruce forest using the gradient method and model. → The mean stomatal uptake of ozone is approximately 47% of the total deposition. → We measure net ecosystem production (NEP) using Eddy Covariance. → We test whether elevated total deposition and stomatal uptake of O 3 imply a reduction of NEP. → Deposition and stomatal uptake of O 3 decrease NEP, especially by high intensities of solar radiation. - Net ecosystem production of a Norway spruce forest decreases with increasing deposition and stomatal uptake of ozone.

  7. Deep physical structure and geotectonic implications of the eastern margin of the Qinghai–Tibet Plateau

    Directory of Open Access Journals (Sweden)

    Jun Li

    2016-10-01

    Full Text Available The eastern margin of the Qinghai–Tibet Plateau (QTP is the focus of studies on eastward lateral extrusion of the latter’s crustal material. This study aims to explore the structural response of the QTP’s eastern crust–mantle to the extrusion, and the basis for the latter’s geological structure. Data on long-period magnetotelluric sounding of cross-tectonic units and Bouguer gravity were used to determine the physical structure of the crust–mantle at the plateau’s eastern margin. The findings are as follows: (i the apparent density structure indicates extensive presence of a low-density material in the middle–lower crusts of the Songpan and Sichuan–Yunnan blocks at the QTP’s eastern margin. On the other hand, the Yangtze cratonic block (Sichuan Basin contains a material with a significantly higher density. To the west of the Longmenshan–Panxi tectonic zones, and along the lower crust at 40–50 km depth, is an obvious low-density zone aligned in a northeast–southwest orientation; (ii the electrical structural model spanning Songpan block–Longmenshan tectonic zone–Yangtze block reveals three distinct electrical structural units along the cross-section bounded by the Longmenshan tectonic zone. The first is the Songpan block, which has high and low resistivity at the shallow layer and middle–lower crusts, respectively. Next is the Yangtze craton, which has low and relatively higher resistivity at the shallow layer and middle–lower crusts, respectively. Third is the Longmenshan transitional tectonic zone, whose shallow layer and deep structure are characterized by an electrical structure with a thrust nappe towards the east, and a high-conductivity material extending to the lithospheric mantle, respectively; (iii the apparent density and electrical structures indicate that the Panxi tectonic zone has a weakened structure in the lower crust; and (iv physical properties of the QTP’s deep structure indicate that its

  8. Impact of climate change on radial growth of Siberian spruce and Scots pine in North-western Russia

    Directory of Open Access Journals (Sweden)

    Lopatin E

    2007-01-01

    Full Text Available When adapting forest management practices to a changing environment, it is very important to understand the response of an unmanaged natural forest to climate change. The method used to identify major climatic factors influencing radial growth of Siberian spruce and Scots pine along a latitudinal gradient in north-western Russia is dendroclimatic analysis. A clear increasing long-term trend was identified in air temperature and precipitation. During the last 20 years, all meteorological stations experienced temperature increases, and 40 years ago precipitation began to increase. This is shown by the radial increment of Siberian spruce and Scots pine. Therefore, climate change could partly explain the increased forest productivity. The total variance explained by temperature varied from 22% to 41% and precipitation from 19% to 38%. The significant climatic parameters for radial increment in Komi Republic were identified, and the relation between temperature and precipitation in explained variance changes over time for Siberian spruce.

  9. Age and size effects on seed productivity of northern black spruce

    Science.gov (United States)

    J. N. Viglas; C. D. Brown; J. F. Johnstone

    2013-01-01

    Slow-growing conifers of the northern boreal forest may require several decades to reach reproductive maturity, making them vulnerable to increases in disturbance frequency. Here, we examine the relationship between stand age and seed productivity of black spruce (Picea mariana (Mill.) Britton, Sterns & Poggenb.) in Yukon Territory and Alaska....

  10. Model of experimental clonal seed orchard for the production of Serbian spruce šPicea omorika /Panc./Purkyne intraspecific hybrids

    Directory of Open Access Journals (Sweden)

    Šijačić-Nikolić Mirjana

    2002-01-01

    Full Text Available The presented model for the establishment of an experimental clonal seed orchard of Serbian spruce was designed based on the results of the analysis and assessment of the genetic potential of Serbian spruce seedling seed orchard at Godovik. Based on the results of the analyses, eight superior half-sib lines of Serbian spruce were selected, of which 24 genotypes were selected. Their hybridisation, by the model of incomplete diallel cross resulted in 21 combinations at the level of half-sib lines, i.e. 48 combinations at the level of parent genotypes. The applied study methods identified the potentially valuable genotypes-cone producers i.e. pollinators, which will be fixed by cloning in the seed orchard of the second generation for the production of the promising hybrids.

  11. Norway spruce (Picea abies Karst. variability in progeny tests in Bosnia and Herzegovina

    Directory of Open Access Journals (Sweden)

    Cvjetković Branislav

    2017-01-01

    Full Text Available Norway spruce is one of the most important economic species is Bosnia and Herzegovina. It is the species at the edge of its natural range; nevertheless it attains significant growth and yield results. The species is often used for afforestation purposes. In the previous period, 4 progeny tests of Norway spruce were established in Bosnia and Herzegovina. The offspring from 6 natural populations: Han Pijesak 1, Han Pijesak 2, Foča, Potoci, Olovo and Kneževo were build-in the progeny tests. In 2016, the samples of Norway spruce from 2 progeny tests: Srebrenica and Drinić were collected. In total, 360 samples were collected. DNA isolation was done according to Dumoline et al. (1990. For assessment of genetic differences among populations, co-dominant nSSR microsatellite system had been used. The number of effective alleles ranged from 7.78 in the population Potoci up to 15 in the population Kneževo, the average number of alleles was 13. The observed heterozygosity ranged from 0.61 for the population Han Pijesak 2 up to 0.68 for population Kneževo. The average observed heterozygosity was 0.65. Fixation index was in the range from -0.073 in the population Potoci, to 0.030 for the population Han Pijesak 2. The average value of Wright fixation index is -0.007. The average fixation index indicates the existence of a very small number of homozygotes. Concerning the variability among populations it has been concluded that the total level of genetic differentiation among populations was very low (FST= 0.026. The result of Nei’s genetic distance shows that the populations Olovo and Potoci are separated from other populations. The results obtained by genetic markers, in addition with other, morphological and physiological markers will be the basis for the further investigation of Norway spruce adaptability and possibility for the transfer of genetic material in light of climate changes.

  12. The diversity of microhabitats and their impact on the regeneration of spruce and rowan in the mountain forests of the Low Tatras

    International Nuclear Information System (INIS)

    Gloncak, P.

    2010-01-01

    In this paper the authors describe the questions: what is proportion of different types of microhabitats in natural spruce?; which types of microhabitats prefers spruce (Picea abies) and rowan (Sorbus aucuparia) in the early stages of their development?; what role do ground vegetation and dead wood play?

  13. The diversity of microhabitats and their impact on the regeneration of spruce and rowan in the mountain forests of the Low Tatras

    International Nuclear Information System (INIS)

    Gloncak, P.

    2010-01-01

    In this presentation the authors describe the questions: what is proportion of different types of microhabitats in natural spruce?; which types of microhabitats prefers spruce (Picea abies) and rowan (Sorbus aucuparia) in the early stages of their development?; what role do ground vegetation and dead wood play?

  14. Near real time/low latency data collection for climate warming manipulations and an elevated CO2 SPRUCE experiment

    Science.gov (United States)

    Krassovski, M.; Hanson, P. J.; Riggs, J. S.; Nettles, W. R., IV

    2017-12-01

    Climate change studies are one of the most important aspects of modern science and related experiments are getting bigger and more complex. One such experiment is the Spruce and Peatland Responses Under Climatic and Environmental Change experiment (SPRUCE, http://mnspruce.ornl.gov) conducted in in northern Minnesota, 40 km north of Grand Rapids, in the USDA Forest Service Marcell Experimental Forest (MEF). The SPRUCE experimental mission is to assess ecosystem-level biological responses of vulnerable, high carbon terrestrial ecosystems to a range of climate warming manipulations and an elevated CO2 atmosphere. This manipulation experiment generates a lot of observational data and requires a reliable onsite data collection system, dependable methods to transfer data to a robust scientific facility, and real-time monitoring capabilities. This presentation shares our experience of establishing near real time/low latency data collection and monitoring system using satellite communication.

  15. Conservation of element concentration in xylem sap of red spruce

    Science.gov (United States)

    Kevin T. Smith; Walter C. Shortle

    2001-01-01

    We investigated the chemistry of xylem sap as a marker of red spruce metabolism and soil chemistry at three locations in northern New England. A Scholander pressure chamber was used to extract xylem sap from roots and branches cut from mature trees in early June and September. Root sap contained significantly greater concentrations of K, Ca, Mg, Mn, and A1 than branch...

  16. Spruce monoculture establishment affects functional traits of soil microarthropod communities

    Czech Academy of Sciences Publication Activity Database

    Farská, Jitka; Prejzková, Kristýna; Rusek, Josef

    2013-01-01

    Roč. 68, č. 3 (2013), s. 479-486 ISSN 0006-3088 R&D Projects: GA ČR GA526/03/1259; GA MŠk LC06066 Grant - others:SGA BF JU(CZ) 30-0004; GAJU(CZ) 143/2010/P Institutional support: RVO:60077344 Keywords : Oribatida * Collembola * spruce * beech Subject RIV: EH - Ecology, Behaviour Impact factor: 0.696, year: 2013

  17. Photosynthesis in Norway spruce seedlings infected by the needle rust Chrysomyxa rhododendri.

    Science.gov (United States)

    Bauer, Helmut; Plattner, Karin; Volgger, Waltraud

    2000-02-01

    Chrysomyxa rhododendri (DC.) De Bary is a needle rust with a host shift between Rhododendron sp. and Norway spruce (Picea abies (L.) Karst.), penetrating only the new developing flushes of the conifer. Because little is known about its effects on trees, we investigated several parameters related to photosynthesis in artificially infected 3-year-old Norway spruce seedlings. The potential efficiency of photosystem II (PSII; derived from chlorophyll fluorescence measurements) was reduced in infected current-year needles as soon as disease symptoms were visible, about three weeks after inoculation. Two weeks later, photosynthetic O(2) evolution (P(max)) of infected needles was less than 20% of control needles, whereas respiratory O(2) uptake (R(D)) was about three times higher than that of control needles. Nonstructural carbohydrate concentrations were about 60% of control values in all parts of the shoots of infected trees. Photosynthetic inhibition was associated with marked decreases in chlorophyll concentration and chlorophyll a/b ratio but only a small reduction in carotenoid concentration. In infected trees, P(max) of noninfected 1-year-old and 2-year-old needles was 50 and 80% higher than in the corresponding age class of needles of control trees. Estimation of potential daily net dry mass production, based on P(max), R(D), specific leaf area, carbon content and needle biomass, indicated that seedlings infected once were able to produce 60%, and those infected twice only 25%, of the dry mass of controls. We conclude that afforestation and regeneration of Norway spruce is seriously impaired in regions where seedlings are frequently attacked by Chrysomyxa.

  18. The Electrical Resistivity Structure of the Eastern Anatolian Collision Zone, Northeastern Anatolia

    Science.gov (United States)

    Cengiz, Özlem; Tuǧrul Başokur, Ahmet; Tolak Çiftçi, Elif

    2016-04-01

    The Northeastern Anatolia is located at the intensely deformed Eastern Anatolian Collision Zone (EACZ), and its tectonic framework is characterized by the collision of the Arabian plate with Eurasian. Although extensive attention is given to understand the crustal and upper mantle processes at this convergent boundary, there is still an ongoing debate over the geodynamic processes of the region. In this study, we were specifically interested in the geoelectric properties and thus geodynamics of the crust beneath the EACZ. Magnetotelluric (MT) measurements were made on two profiles across the north of the EACZ in 1998 as part of a national project undertaken by the Turkish Petroleum Corporation (TPAO). MT data in the frequency range of 300-0.001 Hz were collected from 168 stations located along 78 km north to south and 47 km west to east profiles where direct convergence occurs between Arabian and Eurasian plates. Two and three-dimensional inversion algorithms were used to obtain resistivity models of the study area. According to these models, the upper crust consists of low resistivity sedimentary rocks (basement rocks of the Eastern Anatolian Accretionary Complex and Pontides. While the upper and lower crustal resistivity at the northern part of the study area shows a layered structure, significant horizontal and vertical variations for the rest of the EACZ exists on resistivity models. The broad low resistivity zones (structure supports the southward subduction model with the resistive continental block and the deep conductive zones presumably corresponding to the oceanic crust.

  19. Effects of artificial defoliation of pines on the structure and physiology of the soil fungal community of a mixed pine-spruce forest

    Science.gov (United States)

    Cullings, Ken; Raleigh, Christopher; New, Michael H.; Henson, Joan

    2005-01-01

    Loss of photosynthetic area can affect soil microbial communities by altering the availability of fixed carbon. We used denaturing gradient gel electrophoresis (DGGE) and Biolog filamentous-fungus plates to determine the effects of artificial defoliation of pines in a mixed pine-spruce forest on the composition of the fungal community in a forest soil. As measured by DGGE, two fungal species were affected significantly by the defoliation of pines (P soil fungus increased. The decrease in the amount of Cenococcum organisms may have occurred because of the formation of extensive hyphal networks by species of this genus, which require more of the carbon fixed by their host, or because this fungus is dependent upon quantitative differences in spruce root exudates. The defoliation of pines did not affect the overall composition of the soil fungal community or fungal-species richness (number of species per core). Biolog filamentous-fungus plate assays indicated a significant increase (P soil fungi and the rate at which these substrates were used, which could indicate an increase in fungal-species richness. Thus, either small changes in the soil fungal community give rise to significant increases in physiological capabilities or PCR bias limits the reliability of the DGGE results. These data indicate that combined genetic and physiological assessments of the soil fungal community are needed to accurately assess the effect of disturbance on indigenous microbial systems.

  20. Techno-economic and ex-ante environmental assessment of C6 sugars production from spruce and corn. Comparison of organosolv and wet milling technologies

    NARCIS (Netherlands)

    Moncada, Jonathan; Vural Gursel, Iris; Huijgen, Wouter J J; Dijkstra, Jan Wilco; Ramírez, Andrea

    2018-01-01

    This study assesses the techno-economic and environmental performance of C6 sugars production from softwood (spruce) and corn. Two technologies were considered in the assessment: organosolv of spruce woodchips (2nd generation) and corn wet milling (1st generation). Process models were developed to

  1. Aerodynamic parameter changes above a young spruce forest stand during five growing seasons

    Czech Academy of Sciences Publication Activity Database

    Hurtalová, T.; Matejka, F.; Rožnovský, J.; Marková, Irena; Janouš, Dalibor

    2004-01-01

    Roč. 34, č. 2 (2004), s. 131-146 ISSN 1335-2806 R&D Projects: GA ČR(CZ) GA526/00/0485 Keywords : aerodynamic parameters * roughness length * young spruce stand Subject RIV: DG - Athmosphere Sciences, Meteorology

  2. Aerodynamic resistance of spruce forest stand in relation to roughness length and airflow

    Czech Academy of Sciences Publication Activity Database

    Hurtalová, T.; Matějka, F.; Rožnovský, J.; Janouš, Dalibor

    2003-01-01

    Roč. 33, č. 3 (2003), s. 147-160 ISSN 1335-2806 R&D Projects: GA ČR(CZ) GA526/03/1104 Keywords : aerodynamic resistance * spruce forest stand * roughness length Subject RIV: DG - Athmosphere Sciences, Meteorology

  3. EVALUATION OF THE IMPACT OF THE ECKLONIA MAXIMA EXTRACT ON SELECTED MORPHOLOGICAL FEATURES OF YELLOW PINE, SPRUCE AND THUJA STABBING

    Directory of Open Access Journals (Sweden)

    Jacek Sosnowski Sosnowski

    2016-07-01

    Full Text Available The study was focused on the impact of an extract of Ecklonia maxima on selected morphological features of yellow pine (Pinus ponderosa Dougl. ex C. Lawson, prickly spruce (Picea pungens Engelm. Variety Glauca, thuja (Thuja occidentalis variety Smaragd. The experiment was established in April 12, 2012 on the forest nursery in Ceranów. April 15, 2013 was introduced research agent in the form of a spraying an aqueous solution extract of Ecklonia maxima with trade name Kelpak SL. Biologically active compounds in the extract are plant hormones: auxin and cytokinin. There were studied increment in plant height, needle length of yellow pine, twigs length in prickly spruce and thuja. The measurements of increment in length of twigs and needles were made in each case on the same, specially marked parts of plants and have carried them on the 27th of each month beginning in May and ending in September. The results were evaluated statistically using the analysis of variance. Medium differentiations were verified by Tukey's test at a significance level p ≤ 0.05. The study showed that the diversity of traits features in the experiment was depended on the extract, the tree species and the measurement time. The best results after the extract using showed a pine and spruce. Seaweed preparation contributed to increment increased of trees height for in the pine and spruce and the needles length of pine and twigs of spruce. The species showing no reaction to the extract was thuja.

  4. Wood variables affecting the friction coefficient of spruce pine on steel

    Science.gov (United States)

    Truett J. Lemoine; Charles W. McMillin; Floyd G. Manwiller

    1970-01-01

    Wood of spruce pine, Pinus glabra Walk., was factorially segregated by moisture content (0, 10, and 18 percent), specific gravity (less than 0.45 and more than 0.45), and extractive content (unextracted and extractive-freE), and the kinetic coefficient of friction on steel (having surface roughness of 9 microinches RMS) determined for tangential...

  5. Multipartite Symbioses Among Fungi, Mites, Nematodes, and the Spruce Beetle, Dendroctonus rufipennis.

    Science.gov (United States)

    Yasmin Cardoza; John Moser; Kier Klepzizg; Raffa Kenneth

    2008-01-01

    The spruce beetle, Dendroctonus rufipennis, is an eruptive forest pest of signifcant economic and ecological importance. D. rufipennis has symbiotic associations with a number of microorganisms, especially the ophiostomatoid fungus Leptographium abietinum. The nature of this interaction is only partially understood. Additionally, mite and nematode associates can...

  6. The complete nitrogen cycle of an N-saturated spruce forest ecosystem.

    Science.gov (United States)

    Kreutzer, K; Butterbach-Bahl, K; Rennenberg, H; Papen, H

    2009-09-01

    Long-term nitrogen deposition into forest ecosystems has turned many forests in Central Europe and North America from N-limited to N-saturated systems, with consequences for climate as well as air and groundwater quality. However, complete quantification of processes that convert the N deposited and contributed to ecosystem N cycling is scarce. In this study, we provide the first complete quantification of external and internal N fluxes in an old-growth spruce forest, the Höglwald, Bavaria, Germany, exposed to high chronic N deposition. In this forest, N cycling is dominated by high rates of mineralisation of soil organic matter, nitrification and immobilisation of ammonium and nitrate into microbial biomass. The amount of ammonium available is sufficient to cover the entire N demand of the spruce trees. The data demonstrate the existence of a highly dynamic internal N cycle within the soil, driven by growth and death of the microbial biomass, which turns over approximately seven times each year. Although input and output fluxes are of high environmental significance, they are low compared to the internal fluxes mediated by microbial activity.

  7. Afforestation of Boreal Open Woodlands: Early Performance and Ecophysiology of Planted Black Spruce Seedlings

    Directory of Open Access Journals (Sweden)

    Daniel Lord

    2013-06-01

    Full Text Available Open lichen woodlands (LWs are degraded stands that lack the ability to regenerate naturally due to a succession of natural and/or anthropogenic disturbances. As they represent both interesting forest restoration and carbon sequestration opportunities, we tested disc scarification and planting of two sizes of containerized black spruce (Picea mariana Mill. (BSP seedlings for their afforestation. We compared treatment of unproductive LWs to reforestation of harvested, closed-crown black spruce-feathermoss (BSFM stands. After one year, seedling survival and nutritional status were equivalent among stand types but despite higher root elongation index (REI, planted seedlings in LWs had lower relative growth rate, smaller total biomass and stem diameter than those in BSFM stands. Soil fertility variables, soil temperature, nor seedling water potential, helped at explaining this early growth response. Disc scarification significantly improved seedling first-year survival, biomass and foliar nutrient concentrations of P, Ca, and Mg. Smaller planting stock showed higher REI, higher shoot water potential, and higher foliar nutrient concentration of all but one of the measured nutrients (N, P, K and Mg. Hence, preliminary results suggest that planting of smaller containerized black spruce stock, combined with disc scarification, shows potential for afforestation of unproductive LWs. The impact of the lichen mat and other potential growth limiting factors on afforestation of these sites requires further investigation.

  8. Electromagnetic characterization of white spruce at different moisture contents using synthetic aperture radar imaging

    Science.gov (United States)

    Ingemi, Christopher M.; Owusu Twumasi, Jones; Yu, Tzuyang

    2018-03-01

    Detection and quantification of moisture content inside wood (timber) is key to ensuring safety and reliability of timber structures. Moisture inside wood attracts insects and fosters the development of fungi to attack the timber, causing significant damages and reducing the load bearing capacity during their design life. The use of non-destructive evaluation (NDE) techniques (e.g., microwave/radar, ultrasonic, stress wave, and X-ray) for condition assessment of timber structures is a good choice. NDE techniques provide information about the level of deterioration and material properties of timber structures without obstructing their functionality. In this study, microwave/radar NDE technique was selected for the characterization of wood at different moisture contents. A 12 in-by-3.5 in-by-1.5 in. white spruce specimen (picea glauca) was imaged at different moisture contents using a 10 GHz synthetic aperture radar (SAR) sensor inside an anechoic chamber. The presence of moisture was found to increase the SAR image amplitude as expected. Additionally, integrated SAR amplitude was found beneficial in modeling the moisture content inside the wood specimen.

  9. Characteristics and modeling of spruce wood under dynamic compression load; Charakteristik und Modellierung von Fichtenholz unter dynamischer Druckbelastung

    Energy Technology Data Exchange (ETDEWEB)

    Eisenacher, Germar

    2014-01-28

    Spruce wood is frequently used as an energy absorbing material in impact limiters of packages for the transportation of radioactive material. A 9m drop test onto an unyielding target is mandatory for the packages. The impact results in a dynamic compression load of the spruce wood inside the impact limiter. The lateral dilation of the wood is restrained thereby due to encasing steel sheets. This work's objective was to provide a material model for spruce wood based on experimental investigations to enable the calculation of such loading conditions. About 600 crush tests with cubical spruce wood specimens were performed to characterize the material. The compression was up to 70% and the material was assumed to be transversely isotropic. Particularly the lateral constraint showed to have an important effect: the material develops a high lateral dilation without lateral constraint. The force-displacement characteristics show a comparably low force level and no or only slight hardening. Distinctive softening occurs after the linear-elastic region when loaded parallel to the fiber. On the other hand, using a lateral constraint results in significantly higher general force levels, distinctive hardening and lateral forces. The softening effect when loaded parallel to the fiber is less distinctive. Strain rate and temperature raise or lower the strength level, which was quantified for the applicable ranges of impact limiters. The hypothesis of an uncoupled evolution of the yield surface was proposed based on the experimental findings. It postulates an independent strength evolution with deviatoric and volumetric deformation. The hypothesis could be established using the first modeling approach, the modified LS-DYNA material model MAT075. A transversely isotropic material model was developed based thereupon and implemented in LS-DYNA. The material characteristics of spruce wood were considered using a multi-surface yield criterion and a non-associated flow rule. The

  10. Intermediate-scale community-level flux of CO2 and CH4 in a Minnesota peatland: putting the SPRUCE project in a global context

    Science.gov (United States)

    P. J. Hanson; A. L. Gill; X. Xu; J. R. Phillips; D. J. Weston; Randy Kolka; J. S. Riggs; L. A. Hook

    2016-01-01

    Peatland measurements of CO2 and CH4 flux were obtained at scales appropriate to the in situ biological community below the tree layer to demonstrate representativeness of the spruce and peatland responses under climatic and environmental change (SPRUCE) experiment. Surface flux measurements were made using dual open-path...

  11. Effects of early cone collection and artificial ripening on white spruce and red pine germination

    Energy Technology Data Exchange (ETDEWEB)

    Winston, D.A.; Haddon, B.D.

    1981-12-01

    Two experiments were conducted at the Petawawa National Forestry Institute. Chalk River, Ontario in 1978 to test the feasibility of early cone collection and to determine the optimum conditions for the artificial ripening of white spruce and red pine cones. Cones were collected at periodic intervals, commencing 7 weeks before natural cone ripeness, and stored under four storage conditions and three storage periods. White spruce cones collected August 1, 4 weeks before natural seed dispersal, and stored on open, screened trays for 12 weeks at 5 degrees Celcius and 75-95% relative humidity yielded seeds of high germinability. Seeds extracted from cones immediately after this collection failed to germinate. Cold storage of white spruce cones at 5 degrees Celcius for as little as 4 weeks eliminated dormancy and the subsequent need for seed stratification after extraction. Good germination of red pine seeds was obtained from cones collected August 16, 7 weeks before natural seed dispersal, and stored on screened trays in a well-ventilated, unheated building, for 4 weeks. Completion of embryo growth must be attained before artificial ripening can be successfully applied; it may be used as an index for commencement of cone collections provided subsequent cone handling includes artificial ripening. (Refs. 39).

  12. Increased spruce tree growth in Central Europe since 1960s.

    Science.gov (United States)

    Cienciala, Emil; Altman, Jan; Doležal, Jiří; Kopáček, Jiří; Štěpánek, Petr; Ståhl, Göran; Tumajer, Jan

    2018-04-01

    Tree growth response to recent environmental changes is of key interest for forest ecology. This study addressed the following questions with respect to Norway spruce (Picea abies, L. Karst.) in Central Europe: Has tree growth accelerated during the last five decades? What are the main environmental drivers of the observed tree radial stem growth and how much variability can be explained by them? Using a nationwide dendrochronological sampling of Norway spruce in the Czech Republic (1246 trees, 266 plots), novel regional tree-ring width chronologies for 40(±10)- and 60(±10)-year old trees were assembled, averaged across three elevation zones (break points at 500 and 700m). Correspondingly averaged drivers, including temperature, precipitation, nitrogen (N) deposition and ambient CO 2 concentration, were used in a general linear model (GLM) to analyze the contribution of these in explaining tree ring width variability for the period from 1961 to 2013. Spruce tree radial stem growth responded strongly to the changing environment in Central Europe during the period, with a mean tree ring width increase of 24 and 32% for the 40- and 60-year old trees, respectively. The indicative General Linear Model analysis identified CO 2 , precipitation during the vegetation season, spring air temperature (March-May) and N-deposition as the significant covariates of growth, with the latter including interactions with elevation zones. The regression models explained 57% and 55% of the variability in the two tree ring width chronologies, respectively. Growth response to N-deposition showed the highest variability along the elevation gradient with growth stimulation/limitation at sites below/above 700m. A strong sensitivity of stem growth to CO 2 was also indicated, suggesting that the effect of rising ambient CO 2 concentration (direct or indirect by increased water use efficiency) should be considered in analyses of long-term growth together with climatic factors and N

  13. Observations on the effects of acid rain treatment on needle surfaces of scots pine and Norway spruce seedlings

    International Nuclear Information System (INIS)

    Turunen, M.; Huttunen, S.; Back, J.

    1994-01-01

    Scots pine (Pinus sylvestris L.) and Norway spruce (Picea abies L. Karst.) seedlings were subjected to acid rain treatment at pH 3, pH 4 and pH 7 in a field experiment during 1986-1989. SEM+EDS, TEM, and measurements of wax quantity were used to detect changes in needle surfaces. After 5 weeks at pH 3 and pH 4 acid rain treatment, CaSO 4 -crystallites were observed on visibly undamaged pine and spruce needle surfaces. Direct acid rain damage in conjunction with CaSO 4 -crystallites was observed only occasionally in wax structures. Two-month-old pine needles had 50% less wax in early August after exposure at pH 3 and pH 4 than water controls. The occurrence of CaSO 4 -crystallites on acid rain-treated needle surfaces, and more abundant deposition of Ca oxalate crystallites in the inner walls of epi- and hypodermal cells could be involved with acid rain-induced calcium leaching. Calcium sulphate is probably a result of the disturbed wax and cuticle biosynthesis resulting in undeveloped, permeable cuticles. At the end of experiment, no CaSO 4 -crystallites were seen on needle surfaces. Soil analysis revealed an increase in the soluble Ca concentrations at pH 3. (orig.)

  14. Winter photosynthesis in red spruce (Picea rubens Sarg.): limitations, potential benefits, and risks

    Science.gov (United States)

    P.G. Schaberg

    2000-01-01

    Numerous cold-induced changes in physiology limit the capacity of northern conifers to photosynthesize during winter. Studies of red spruce (Picea rubens Sarg.) have shown that rates of field photosynthesis (Pfield) and laboratory measurements of photosynthetic capacity (Pmax) generally parallel seasonal...

  15. The effects of fire on the thermal stability of permafrost in lowland and upland black spruce forests of interior Alaska in a changing climate

    International Nuclear Information System (INIS)

    Jafarov, E E; Romanovsky, V E; Marchenko, S S; Genet, H; McGuire, A D

    2013-01-01

    Fire is an important factor controlling the composition and thickness of the organic layer in the black spruce forest ecosystems of interior Alaska. Fire that burns the organic layer can trigger dramatic changes in the underlying permafrost, leading to accelerated ground thawing within a relatively short time. In this study, we addressed the following questions. (1) Which factors determine post-fire ground temperature dynamics in lowland and upland black spruce forests? (2) What levels of burn severity will cause irreversible permafrost degradation in these ecosystems? We evaluated these questions in a transient modeling–sensitivity analysis framework to assess the sensitivity of permafrost to climate, burn severity, soil organic layer thickness, and soil moisture content in lowland (with thick organic layers, ∼80 cm) and upland (with thin organic layers, ∼30 cm) black spruce ecosystems. The results indicate that climate warming accompanied by fire disturbance could significantly accelerate permafrost degradation. In upland black spruce forest, permafrost could completely degrade in an 18 m soil column within 120 years of a severe fire in an unchanging climate. In contrast, in a lowland black spruce forest, permafrost is more resilient to disturbance and can persist under a combination of moderate burn severity and climate warming. (letter)

  16. Non-functional plastid ndh gene fragments are present in the nuclear genome of Norway spruce (Picea abies L. Karsch): insights from in silico analysis of nuclear and organellar genomes.

    Science.gov (United States)

    Ranade, Sonali Sachin; García-Gil, María Rosario; Rosselló, Josep A

    2016-04-01

    Many genes have been lost from the prokaryote plastidial genome during the early events of endosymbiosis in eukaryotes. Some of them were definitively lost, but others were relocated and functionally integrated to the host nuclear genomes through serial events of gene transfer during plant evolution. In gymnosperms, plastid genome sequencing has revealed the loss of ndh genes from several species of Gnetales and Pinaceae, including Norway spruce (Picea abies). This study aims to trace the ndh genes in the nuclear and organellar Norway spruce genomes. The plastid genomes of higher plants contain 11 ndh genes which are homologues of mitochondrial genes encoding subunits of the proton-pumping NADH-dehydrogenase (nicotinamide adenine dinucleotide dehydrogenase) or complex I (electron transport chain). Ndh genes encode 11 NDH polypeptides forming the Ndh complex (analogous to complex I) which seems to be primarily involved in chloro-respiration processes. We considered ndh genes from the plastidial genome of four gymnosperms (Cryptomeria japonica, Cycas revoluta, Ginkgo biloba, Podocarpus totara) and a single angiosperm species (Arabidopsis thaliana) to trace putative homologs in the nuclear and organellar Norway spruce genomes using tBLASTn to assess the evolutionary fate of ndh genes in Norway spruce and to address their genomic location(s), structure, integrity and functionality. The results obtained from tBLASTn were subsequently analyzed by performing homology search for finding ndh specific conserved domains using conserved domain search. We report the presence of non-functional plastid ndh gene fragments, excepting ndhE and ndhG genes, in the nuclear genome of Norway spruce. Regulatory transcriptional elements like promoters, TATA boxes and enhancers were detected in the upstream regions of some ndh fragments. We also found transposable elements in the flanking regions of few ndh fragments suggesting nuclear rearrangements in those regions. These evidences

  17. Hartig' net formation of Tricholoma vaccinum-spruce ectomycorrhiza in hydroponic cultures.

    Science.gov (United States)

    Henke, Catarina; Jung, Elke-Martina; Kothe, Erika

    2015-12-01

    For re-forestation of metal-contaminated land, ectomycorrhizal trees may provide a solution. Hence, the study of the interaction is necessary to allow for comprehensive understanding of the mutually symbiotic features. On a structural level, hyphal mantle and the Hartig' net formed in the root apoplast are essential for plant protection and mycorrhizal functioning. As a model, we used the basidiomycete Tricholoma vaccinum and its host spruce (Picea abies). Using an optimized hydroponic cultivation system, both features could be visualized and lower stress response of the tree was obtained in non-challenged cultivation. Larger spaces in the apoplasts could be shown with high statistical significance. The easy accessibility will allow to address metal stress or molecular responses in both partners. Additionally, the proposed cultivation system will enable for other experimental applications like addressing flooding, biological interactions with helper bacteria, chemical signaling, or other biotic or abiotic challenges relevant in the natural habitat.

  18. SOIL CO2 EFFLUX FROM ISOTOPICALLY LABELED BEECH AND SPRUCE IN SOUTHERN GERMANY

    Science.gov (United States)

    • Carbon acquisition and transport to roots in forest trees is difficult to quantify and is affected by a number of factors, including micrometeorology and anthropogenic stresses. The canopies of mature European beech (Fagus sylvatica) and Norway spruce (Picea abies) were expose...

  19. Soil saprotrophic micromycetes in Norway spruce forests in the Šumava National Park

    Czech Academy of Sciences Publication Activity Database

    Nováková, Alena

    2001-01-01

    Roč. 7, - (2001), s. 177-184 ISSN 1211-7420 R&D Projects: GA ČR GA206/99/1416 Institutional research plan: CEZ:AV0Z6066911 Keywords : soil saprotrophic micromycetes * Norway spruce forest * bark beetle Subject RIV: EH - Ecology, Behaviour

  20. Contribution of black spruce (Picea mariana) transpiration to growing season evapotranspiration in a subarctic discontinuous permafrost peatland complex

    Science.gov (United States)

    Helbig, M.; Warren, R. K.; Pappas, C.; Sonnentag, O.; Berg, A. A.; Chasmer, L.; Baltzer, J. L.; Quinton, W. L.; Patankar, R.

    2016-12-01

    Partitioning the components of evapotranspiration (ET), evaporation and transpiration, has been increasingly important for the better understanding and modeling of carbon, water, and energy dynamics, and for reliable water resources quantification and management. However, disentangling its individual processes remains highly uncertain. Here, we quantify the contribution of black spruce transpiration, the dominant overstory, to ET of a boreal forest-wetland landscape in the southern Taiga Plains. In these ecosystems, thawing permafrost induces rapid landscape change, whereby permafrost-supported forested plateaus are transformed into bogs or fens (wetlands), resulting in tree mortality. Using historical and projected rates of forest-wetland changes, we assess how the contribution of black spruce transpiration to landscape ET might be altered with continued permafrost loss, and quantify the resulting water balance changes. We use two nested eddy covariance flux towers and a footprint model to quantify ET over the entire landscape. Sap flux density of black spruce is measured using the heat ratio method during the 2013 (n=22) and 2014 (n=3) growing seasons, and is used to estimate tree-level transpiration. Allometric relations between tree height, diameter at breast height and sapwood area are derived to upscale tree-level transpiration to overstory transpiration within the eddy covariance footprint. Black spruce transpiration accounts for <10% of total landscape ET. The largest daily contribution of overstory transpiration to landscape ET is observed shortly after the landscape becomes snow-free, continually decreasing throughout the progression of the growing season. Total transpiration is notably lower in 2014 (2.34 mm) than 2013 (2.83 mm) over the same 40-day period, corresponding to 3% of cumulative landscape ET in both years. This difference is likely due to the antecedent moisture conditions, where the 2014 growing season was proceeded by lower than average

  1. An experimental study on the effects of exhaust gas on spruce (Picea abies L. Karst.)

    Energy Technology Data Exchange (ETDEWEB)

    Hautala, E.L.; Holopainen, J.; Kaerenlampi, L. [Kuopio Univ. (Finland). Dept. of Ecology and Environmental Science; Surakka, J.; Ruuskanen, J. [Kuopio Univ. (Finland). Dept. of Environmental Sciences

    1995-12-31

    Motor vehicle exhausts are significant contributors to air pollution. Besides fine particles and inorganic gases, like CO, SO{sub 2} and NO{sub x}, exhaust gas contains a large group of aromatic hydrocarbon compounds, many of which are phytotoxic. In field studies, exhausts are found to have both direct and indirect harmful effects on roadside plants. However, only few experimental studies have been made about the effects of exhaust gas emissions on coniferous trees. The aim of this study was to survey the effects of exhausts on spruce (Picea abies L. Karst.) in standardized conditions. The concentrations of major exhaust gas components in the chamber atmosphere were detected simultaneously. The effects of exhaust on epistomatal waxes of first-year spruce needles are described. (author)

  2. An experimental study on the effects of exhaust gas on spruce (Picea abies L. Karst.)

    Energy Technology Data Exchange (ETDEWEB)

    Hautala, E L; Holopainen, J; Kaerenlampi, L [Kuopio Univ. (Finland). Dept. of Ecology and Environmental Science; Surakka, J; Ruuskanen, J [Kuopio Univ. (Finland). Dept. of Environmental Sciences

    1996-12-31

    Motor vehicle exhausts are significant contributors to air pollution. Besides fine particles and inorganic gases, like CO, SO{sub 2} and NO{sub x}, exhaust gas contains a large group of aromatic hydrocarbon compounds, many of which are phytotoxic. In field studies, exhausts are found to have both direct and indirect harmful effects on roadside plants. However, only few experimental studies have been made about the effects of exhaust gas emissions on coniferous trees. The aim of this study was to survey the effects of exhausts on spruce (Picea abies L. Karst.) in standardized conditions. The concentrations of major exhaust gas components in the chamber atmosphere were detected simultaneously. The effects of exhaust on epistomatal waxes of first-year spruce needles are described. (author)

  3. Processing of spruce and larch resins to obtain balsam for the confection industry

    Energy Technology Data Exchange (ETDEWEB)

    Buinova, E.F.; Yaremchenko, N.G.; Izotova, L.V.; Lepeiko, A.G.; Meteshkina, L.P.; Zhuravlev, P.I.; Chumakov, V. A.; Nikiforova, V.N.; Krylova, E.N.

    1981-01-01

    Modified balsam was prepared by esterification of purified and precipitated larch or spruce oleoresins with glycerol at 255-265 degrees for 4-6 hours. The modified balsams, which were obtained in 55-65% yield, can be used as a base for chewing gum manufacture.

  4. ESTIMATING ROOT RESPIRATION IN SPRUCE AND BEECH: DECREASES IN SOIL RESPIRATION FOLLOWING GIRDLING

    Science.gov (United States)

    A study was undertaken to follow seasonal fluxes of CO2 from soil and to estimate the contribution of autotrophic (root + mycorrhizal) to total soil respiration (SR) in a mixed stand of European beech (Fagus sylvatica) and Norway spruce (Picea abies) near Freising, Germany. Matu...

  5. Management of spruce-fir in even-aged stands in the central Rocky Mountains

    Science.gov (United States)

    Robert R. Alexander; Carleton B. Edminster

    1980-01-01

    Potential production of Engelmann spruce and subalpine fir in the central Rocky Mountains is simulated for vario.us combinations of stand density, site quality, ages, and thinning schedules. Such estimates are needed to project future development of stands managed in different ways for various uses.

  6. Evaluation of IRS-1C LISS-3 satellite data for Norway spruce defoliation assessment

    Energy Technology Data Exchange (ETDEWEB)

    Falkenstroem, H.

    1999-02-01

    Satellite based remote sensing supported by air photo and field surveys, provide a means to area covering forest health assessment on a regional scale. Landsat TM data has been extensively used in studies of spruce and fir defoliation in Europe and North America. The temporal coverage of Landsat TM in combination with cloudiness however restrict the availability of data. In this study the LISS-3 sensor onboard the Indian Resource Satellite, IRS-1C, was evaluated for defoliation assessments in Norway spruce (Picea abies) in the central part of Sweden. The near infrared wavelength band proved to be best correlated with mean stand defoliation. After normalisation of satellite data for topographic conditions, the correlation coefficient increased from -0,19 to -0,83. Normalising satellite data for species composition did not improve the results though. The correction coefficients involved in the procedure were originally developed for Landsat TM, and proved to be inadequate for the LISS-3 data set. A thorough examination of the effects of species composition on LISS-3 data is needed to yield better results. The correlation between observed defoliation in the verification stands and predicted (based on the inverse regression function between corrected NIR values and defoliation in reference stands) was 0,70, despite a very limited range of defoliation in the verification set. IRS-1C LISS-3 is fully comparable to Landsat TM for spruce defoliation studies, although the results would probably not be significantly improved 49 refs, 7 figs, 10 tabs

  7. Sequestration of soil nitrogen as tannin-protein complexes may improve the competitive ability of sheep laurel (Kalmia angustifolia) relative to black spruce (Picea mariana).

    Science.gov (United States)

    Joanisse, G D; Bradley, R L; Preston, C M; Bending, G D

    2009-01-01

    The role of litter tannins in controlling soil nitrogen (N) cycling may explain the competitive ability of Kalmia relative to black spruce (Picea mariana), although this has not been demonstrated experimentally. Here, the protein-precipitation capacities of purified tannins and leaf extracts from Kalmia and black spruce were compared. The resistance to degradation of tannin-protein precipitates from both species were compared by monitoring carbon (C) and N dynamics in humus amended with protein, purified tannins or protein-tannin precipitates. The purity of the precipitates was verified using solid-state (13)C nuclear magnetic resonance (NMR) spectra. The ability of mycorrhizal fungi associated with both species to grow on media amended with tannin-protein complexes as the principal N source was also compared. The protein precipitation capacity of Kalmia tannins was superior to those of black spruce. Humus amended with protein increased both mineral and microbial N, whereas humus amended with tannin-protein precipitates increased dissolved organic N. Mycorrhizal fungi associated with Kalmia showed better growth than those associated with black spruce when N was provided as tannin-protein precipitates. These data suggest that Kalmia litter increases the amount of soil N sequestered as tannin-protein complexes, which may improve the competitive ability of Kalmia relative to black spruce by favouring N uptake by mycorrhizas associated with the former.

  8. Size-dependence of tree growth response to drought for Norway spruce and European beech individuals in monospecific and mixed-species stands.

    Science.gov (United States)

    Ding, H; Pretzsch, H; Schütze, G; Rötzer, T

    2017-09-01

    Climate anomalies have resulted in changing forest productivity, increasing tree mortality in Central and Southern Europe. This has resulted in more severe and frequent ecological disturbances to forest stands. This study analysed the size-dependence of growth response to drought years based on 384 tree individuals of Norway spruce [Picea abies (L.) Karst.] and European beech [Fagus sylvatica ([L.)] in Bavaria, Germany. Samples were collected in both monospecific and mixed-species stands. To quantify the growth response to drought stress, indices for basal area increment, resistance, recovery and resilience were calculated from tree ring measurements of increment cores. Linear mixed models were developed to estimate the influence of drought periods. The results show that ageing-related growth decline is significant in drought years. Drought resilience and resistance decrease significantly with growth size among Norway spruce individuals. Evidence is also provided for robustness in the resilience capacity of European beech during drought stress. Spruce benefits from species mixing with deciduous beech, with over-yielding spruce in pure stands. The importance of the influence of size-dependence within tree growth studies during disturbances is highlighted and should be considered in future studies of disturbances, including drought. © 2017 German Botanical Society and The Royal Botanical Society of the Netherlands.

  9. Using silviculture to influence carbon sequestration in southern Appalachian spruce-fir forests

    Science.gov (United States)

    Patrick T. Moore; R. Justin DeRose; James N. Long; Helga. van Miegroet

    2012-01-01

    Enhancement of forest growth through silvicultural modification of stand density is one strategy for increasing carbon (C) sequestration. Using the Fire and Fuels Extension of the Forest Vegetation Simulator, the effects of even-aged, uneven-aged and no-action management scenarios on C sequestration in a southern Appalachian red spruce-Fraser fir forest were modeled....

  10. Effects of sewage sludge addition to Norway spruce seedlings on nitrogen availability and soil fauna in clear-cut areas

    International Nuclear Information System (INIS)

    Nieminen, Jouni K.; Räisänen, Mikko

    2013-01-01

    Anaerobically digested and composted sewage sludge (CSS) has been suggested to be a slow-release fertilizer in forestry and an alternative to quick-release inorganic fertilizers. The effects of CSS with or without added carbohydrate on inorganic nitrogen availability and on soil animals were tested in two Norway spruce plantations. Half of the seedlings were individually fertilized with CSS, and the rest were left as controls. Solid sucrose was added to half of the fertilized and untreated seedlings. Soil samples were taken in the autumn in the first and the second year after the treatments. CSS increased soil NH 4 –N (2100%), the proportion of soil NO 3 –N, and the N concentration of spruce needles. CSS greatly reduced the abundances of enchytraeids, tardigrades and collembolans, but increased the proportion and abundance of bacterial-feeding nematodes irrespective of carbohydrate addition. A better stabilization method needs to be developed before CSS can be used as a forest fertilizer. -- Highlights: •Spruces were fertilized with anaerobically digested and composted sewage sludge (CSS). •CSS increased soil N, proportion of NO 3 –N, and N concentration of spruce needles. •CSS reduced the abundances of enchytraeids, tardigrades and collembolans. •CSS increased the proportion and abundance of bacterial-feeding nematodes. •Sucrose did not reduce N pools or counteract negative CSS effects on soil animals. -- Composting and carbohydrate addition do not mitigate the harmful effects of anaerobically digested sewage sludge in boreal forest soil

  11. Crustal structure of the Central-Eastern Greenland: results from the TopoGreenland refraction profile

    Science.gov (United States)

    Shulgin, Alexey; Thybo, Hans

    2014-05-01

    Until present, seismic surveys have only been carried out offshore and near the coasts of Greenland, where the crustal structure is affected by oceanic break-up. We present the deep seismic structure of the crust of the interior of Greenland, based on the new and the only existing so far seismic refraction/wide-angle reflection profile. The seismic data was acquired by a team of six people during a two-month long experiment in summer of 2011 on the ice cap in the interior of central-eastern Greenland. The presence of an up to 3.4 km thick ice sheet, permanently covering most of the land mass, made acquisition of geophysical data logistically complicated. The profile extends 310 km inland in E-W direction from the approximate edge of the stable ice cap near the Scoresby Sund across the center of the ice cap. 350 Reftek Texan receivers recorded high-quality seismic data from 8 equidistant shots along the profile. Explosive charge sizes were 1 ton at the ends and ca. 500 kg along the profile, loaded with about 125 kg at 35-85 m depth in individual boreholes. Given that the data acquisition was affected by the thick ice sheet, we questioned the quality of seismic records in such experiment setup. We have developed an automatic routine to check the amplitudes and spectra of the selected seismic phases and to check the differences/challenges in making seismic experiments on ice and the effects of ice on data interpretation. Using tomographic inversion and forward ray tracing modelling we have obtained the two-dimensional velocity model down to a 50 km depth. The model shows a decrease of crustal thickness from 47 km below the centre of Greenland in the western part of the profile to 40 km in its eastern part. Relatively high lower crustal velocities (Vp 6.8 - 7.3 km/s) in the western part of the TopoGreenland profile may result from past collision tectonics or, alternatively, may be related to the speculated passage of the Iceland mantle plume. Comparison of our results

  12. Diet compositions and trophic guild structure of the eastern Chukchi Sea demersal fish community

    Science.gov (United States)

    Whitehouse, George A.; Buckley, Troy W.; Danielson, Seth L.

    2017-01-01

    Fishes are an important link in Arctic marine food webs, connecting production of lower trophic levels to apex predators. We analyzed 1773 stomach samples from 39 fish species collected during a bottom trawl survey of the eastern Chukchi Sea in the summer of 2012. We used hierarchical cluster analysis of diet dissimilarities on 21 of the most well sampled species to identify four distinct trophic guilds: gammarid amphipod consumers, benthic invertebrate generalists, fish and shrimp consumers, and zooplankton consumers. The trophic guilds reflect dominant prey types in predator diets. We used constrained analysis of principal coordinates (CAP) to determine if variation within the composite guild diets could be explained by a suite of non-diet variables. All CAP models explained a significant proportion of the variance in the diet matrices, ranging from 7% to 25% of the total variation. Explanatory variables tested included latitude, longitude, predator length, depth, and water mass. These results indicate a trophic guild structure is present amongst the demersal fish community during summer in the eastern Chukchi Sea. Regular monitoring of the food habits of the demersal fish community will be required to improve our understanding of the spatial, temporal, and interannual variation in diet composition, and to improve our ability to identify and predict the impacts of climate change and commercial development on the structure and functioning of the Chukchi Sea ecosystem.

  13. Quantifying sources of variation in the frequency of fungi associated with spruce beetles: implications for hypothesis testing and sampling methodology in bark beetle-symbiont relationships.

    Science.gov (United States)

    Brian H. Aukema; Richard A. Werner; Kirsten E. Haberkern; Barbara L. Illman; Murray K. Clayton; Kenneth F. Raffa

    2005-01-01

    The spruce beetle, Dendroctonus rufipennis (Kirby), causes landscape level mortality to mature spruce (Picea spp.) throughout western and northern North America. As with other bark beetles, this beetle is associated with a variety of fungi, whose ecological functions are largely unknown. It has been proposed that the relative...

  14. Dynamics of calcium concentration in stemwood of red spruce and Siberian fir

    Science.gov (United States)

    Kevin T. Smith; Walter C. Shortle; Rakesh Minocha; Vladislav A. Alexeyev

    1996-01-01

    The atmospheric deposition of strong acid anions such as sulfate and nitrate shifts the ion exchange equilibrium in the rooting zone of sensitive forests. Red spruce and other northern coniferous forests are especially sensitive to deposition due to the shallow rooting of trees in a mor-type forest floor. Initially, the deposition of strong acid ions mobilizes...

  15. Red spruce stand dynamics, simulations, and restoration opportunities in the central Appalachians

    Science.gov (United States)

    James S. Rentch; Thomas M. Schuler; W. Mark Ford; Gergory J. Nowacki

    2007-01-01

    Red spruce (Picea rubens)-dominated forests occupied as much as 600,000 ha in West Virginia prior to exploitive logging era of the late nineteenth and early twentieth centuries. Subsequently, much of this forest type was converted to northern hardwoods. As an important habitat type for a number of rare or sensitive species, only about 12,000 ha of...

  16. Two spruce shoot candidate reference materials from the German environmental specimen bank

    International Nuclear Information System (INIS)

    Backhaus, F.; Bagschik, U.; Burow, M.; Froning, M.; Mohl, C.; Ostapczuk, P.; Rossbach, M.; Schladot, J.D.; Stoeppler, M.; Waidmann, E.; Byrne, A.R.; Zeisler, R.

    1994-01-01

    Two new materials are introduced that might serve as useful aids for the harmonisation of analytical results. Spruce shoots, cryogenically homogenized and characterized for 50 elements from two sampling sites of the German Environmental Specimen Bank (ESB) are presented as possible third generation reference materials that might also act as calibrating materials in speciation analysis. (author)

  17. Juvenility and serial vegetative propagation of Norway spruce clones (Picea abies Karst.).

    Science.gov (United States)

    J.B. St. Clair; J. Kleinschmit; J. Svolba

    1985-01-01

    Effects associated with progressive maturation of clones are of greatest concern in clonal tree improvement programs. Serial propagation has been in use at the Lower Saxony Forest Research Institute since 1968 to arrest maturation in Norway spruce clones. By 1980 cuttings were established in the nursery that had been serially propagated from one to five cycles. This...

  18. Stable carbon isotope analysis reveals widespread drought stress in boreal black spruce forests.

    Science.gov (United States)

    Walker, Xanthe J; Mack, Michelle C; Johnstone, Jill F

    2015-08-01

    Unprecedented rates of climate warming over the past century have resulted in increased forest stress and mortality worldwide. Decreased tree growth in association with increasing temperatures is generally accepted as a signal of temperature-induced drought stress. However, variations in tree growth alone do not reveal the physiological mechanisms behind recent changes in tree growth. Examining stable carbon isotope composition of tree rings in addition to tree growth can provide a secondary line of evidence for physiological drought stress. In this study, we examined patterns of black spruce growth and carbon isotopic composition in tree rings in response to climate warming and drying in the boreal forest of interior Alaska. We examined trees at three nested scales: landscape, toposequence, and a subsample of trees within the toposequence. At each scale, we studied the potential effects of differences in microclimate and moisture availability by sampling on northern and southern aspects. We found that black spruce radial growth responded negatively to monthly metrics of temperature at all examined scales, and we examined ∆(13)C responses on a subsample of trees as representative of the wider region. The negative ∆(13)C responses to temperature reveal that black spruce trees are experiencing moisture stress on both northern and southern aspects. Contrary to our expectations, ∆(13)C from trees on the northern aspect exhibited the strongest drought signal. Our results highlight the prominence of drought stress in the boreal forest of interior Alaska. We conclude that if temperatures continue to warm, we can expect drought-induced productivity declines across large regions of the boreal forest, even for trees located in cool and moist landscape positions. © 2015 John Wiley & Sons Ltd.

  19. MILD ALKALINE TREATMENT ACTIVATES SPRUCE WOOD FOR ENZYMATIC PROCESSING: A POSSIBLE STAGE IN BIO-REFINERY PROCESSES

    Directory of Open Access Journals (Sweden)

    Yan Wang

    2011-05-01

    Full Text Available The structure of wood is so compact that enzymes are too large to penetrate into the structure and thereby attack the wood components for modifications that can be valuable for various purposes. Here we present a pretreatment method based on traditional kraft pulping, which opens the wood structure, so that enzymes are able to attack the wood components. To study this kind of chemical pretreatment, spruce wood samples were treated at similar conditions used in kraft cooking at varying intensities (H-factors. To verify if the structure was “opened” for enzymes, the pretreated wood samples were incubated with a cellulolytic culture filtrate, and the released reducing sugar concentration after the enzymatic hydrolysis was measured. The results indicated that un-pretreated wood fibers could not be attacked by the enzymes, but already relatively mild pretreatment was sufficient for letting the culture filtrate attack wood polysaccharides, and more intensive treatments opened the structure further. The mildest treatments did not cause any significant yield losses of lignin (Klason lignin. Some galactogluco-mannans were however lost during the pretreatments. The mechanisms behind the effect and the technical significance of the method are discussed.

  20. Effects of artificial defoliation of pines on the structure and physiology of the soil fungal community of a mixed pine-spruce forest

    Science.gov (United States)

    Cullings, Ken; Raleigh, Christopher; New, Michael H.; Henson, Joan

    2005-01-01

    Loss of photosynthetic area can affect soil microbial communities by altering the availability of fixed carbon. We used denaturing gradient gel electrophoresis (DGGE) and Biolog filamentous-fungus plates to determine the effects of artificial defoliation of pines in a mixed pine-spruce forest on the composition of the fungal community in a forest soil. As measured by DGGE, two fungal species were affected significantly by the defoliation of pines (P the frequency of members of the ectomycorrhizal fungus genus Cenococcum decreased significantly, while the frequency of organisms of an unidentified soil fungus increased. The decrease in the amount of Cenococcum organisms may have occurred because of the formation of extensive hyphal networks by species of this genus, which require more of the carbon fixed by their host, or because this fungus is dependent upon quantitative differences in spruce root exudates. The defoliation of pines did not affect the overall composition of the soil fungal community or fungal-species richness (number of species per core). Biolog filamentous-fungus plate assays indicated a significant increase (P the number of carbon substrates utilized by the soil fungi and the rate at which these substrates were used, which could indicate an increase in fungal-species richness. Thus, either small changes in the soil fungal community give rise to significant increases in physiological capabilities or PCR bias limits the reliability of the DGGE results. These data indicate that combined genetic and physiological assessments of the soil fungal community are needed to accurately assess the effect of disturbance on indigenous microbial systems.

  1. Application of Raman spectroscopy to analyse lignin/cellulose ratio in Norway spruce tree rings

    Czech Academy of Sciences Publication Activity Database

    Vítek, Petr; Klem, Karel; Urban, Otmar

    2017-01-01

    Roč. 20, 1-2 (2017), s. 41-48 ISSN 1803-2451 R&D Projects: GA MŠk(CZ) LO1415 Institutional support: RVO:86652079 Keywords : dendrochronology * vibrational spectroscopy * lignification * Norway spruce * abiotic stress Subject RIV: EF - Botanics OBOR OECD: Plant sciences, botany

  2. Internal Structure of a Lobate Debris Apron Complex in Eastern Hellas: Evidence for Multiple Mid-Latitude Glaciations on Mars

    Science.gov (United States)

    Quartini, E.; Holt, J. W.; Brothers, T. C.

    2011-03-01

    The past depositional history of a lobate debris apron complex in eastern Hellas has been investigated by conducting a combined analysis of its surface morphology and subsurface structure using a CTX mosaic and orbital radar sounding data from SHARAD.

  3. Foliar and soil chemistry at red spruce sites in the Monongahela National Forest

    Science.gov (United States)

    Stephanie J. Connolly

    2010-01-01

    In 2005, soil and foliar chemistry were sampled from 10 sites in the Monongahela National Forest which support red spruce. Soils were sampled from hand-dug pits, by horizon, from the O-horizon to bedrock or 152 cm, and each pit was described fully. Replicate, archived samples also were collected.

  4. Ecological Factors Influencing Norway Spruce Regeneration on Nurse Logs in a Subalpine Virgin Forest

    Directory of Open Access Journals (Sweden)

    Sophie Stroheker

    2018-03-01

    Full Text Available Regeneration of Picea abies in high-elevation mountain forests often depends on the presence of coarse woody debris (CWD, as logs provide sites with more favorable conditions for spruce regeneration compared to the forest floor. However, there is little quantitative knowledge on the factors that are conducive to or hindering spruce establishment on CWD. We examined spruce regeneration on CWD by sampling 303 plots (50 cm × 50 cm each on 56 downed logs in a virgin forest in the Swiss Alps. Variables describing microsite conditions were measured, and fungi were isolated from wood samples. To investigate the relationship between the ecological factors and establishment success, two models were fitted with seedling and sapling density as response variables, respectively. Besides log diameter, the models identified different ecological factors as significant for seedling and sapling establishment, i.e., regeneration depends on different factors in different development stages. Seedling density depended on the type of rot, log inclination, and decay stage. Sapling density depended mainly on light availability, cover by bark and moss, the time of tree fall, and the distance between the log surface and the forest floor. A total of 22 polypore fungi were isolated from the wood samples, four of them being threatened species. White- and brown-rot fungi were found in all decay stages. The visual assessment of the type of rot in the field corresponded in only 15% of cases to the type of rot caused by the isolated fungi; hence caution is needed when making field assessments of rot types.

  5. 3D Crustal Velocity Structure Model of the Middle-eastern North China Craton

    Science.gov (United States)

    Duan, Y.; Wang, F.; Lin, J.; Wei, Y.

    2017-12-01

    Lithosphere thinning and destruction in the middle-eastern North China Craton (NCC), a region susceptible to strong earthquakes, is one of the research hotspots in solid earth science. Up to 42 wide-angle reflection/refraction deep seismic sounding (DSS) profiles have been completed in the middle-eastern NCC, we collect all the 2D profiling results and perform gridding of the velocity and interface depth data, and build a 3D crustal velocity structure model for the middle-eastern NCC, named HBCrust1.0, using the Kriging interpolation method. In this model, four layers are divided by three interfaces: G is the interface between the sedimentary cover and crystalline crust, with velocities of 5.0-5.5 km/s above and 5.8-6.0 km/s below. C is the interface of the upper and lower crust, with velocity jump from 6.2-6.4 km/s to 6.5-6.6 km/s. M is the interface between the crust and upper mantle, with velocity 6.7-7.0 km/s at the crust bottom and 7.9-8.0 km/s on mantle top. Our results show that the first arrival time calculated from HBCust1.0 fit well with the observation. It also demonstrates that the upper crust is the main seismogenic layer, and the brittle-ductile transition occurs at depths near interface C. The depth of interface Moho varies beneath the source area of the Tangshan earth-quake, and a low-velocity structure is found to extend from the source area to the lower crust. Based on these observations, it can be inferred that stress accumulation responsible for the Tangshan earthquake may have been closely related to the migration and deformation of the mantle materials. Comparisons of the average velocities of the whole crust, the upper and the lower crust show that the average velocity of the lower crust under the central part of the North China Basin (NCB) in the east of the craton is obviously higher than the regional average, this high-velocity probably results from longterm underplating of the mantle magma. This research is founded by the Natural Science

  6. In Silico Analysis of Small RNAs Suggest Roles for Novel and Conserved miRNAs in the Formation of Epigenetic Memory in Somatic Embryos of Norway Spruce.

    Science.gov (United States)

    Yakovlev, Igor A; Fossdal, Carl G

    2017-01-01

    Epigenetic memory in Norway spruce affects the timing of bud burst and bud set, vitally important adaptive traits for this long-lived forest species. Epigenetic memory is established in response to the temperature conditions during embryogenesis. Somatic embryogenesis at different epitype inducing (EpI) temperatures closely mimics the natural processes of epigenetic memory formation in seeds, giving rise to epigenetically different clonal plants in a reproducible and predictable manner, with respect to altered bud phenology. MicroRNAs (miRNAs) and other small non-coding RNAs (sRNAs) play an essential role in the regulation of plant gene expression and may affect this epigenetic mechanism. We used NGS sequencing and computational in silico methods to identify and profile conserved and novel miRNAs among small RNAs in embryogenic tissues of Norway spruce at three EpI temperatures (18, 23 and 28°C). We detected three predominant classes of sRNAs related to a length of 24 nt, followed by a 21-22 nt class and a third 31 nt class of sRNAs. More than 2100 different miRNAs within the prevailing length 21-22 nt were identified. Profiling these putative miRNAs allowed identification of 1053 highly expressed miRNAs, including 523 conserved and 530 novels. 654 of these miRNAs were found to be differentially expressed (DEM) depending on EpI temperature. For most DEMs, we defined their putative mRNA targets. The targets represented mostly by transcripts of multiple-repeats proteins, like TIR, NBS-LRR, PPR and TPR repeat, Clathrin/VPS proteins, Myb-like, AP2, etc. Notably, 124 DE miRNAs targeted 203 differentially expressed epigenetic regulators. Developing Norway spruce embryos possess a more complex sRNA structure than that reported for somatic tissues. A variety of the predicted miRNAs showed distinct EpI temperature dependent expression patterns. These putative EpI miRNAs target spruce genes with a wide range of functions, including genes known to be involved in epigenetic

  7. In Silico Analysis of Small RNAs Suggest Roles for Novel and Conserved miRNAs in the Formation of Epigenetic Memory in Somatic Embryos of Norway Spruce

    Directory of Open Access Journals (Sweden)

    Igor A. Yakovlev

    2017-09-01

    Full Text Available Epigenetic memory in Norway spruce affects the timing of bud burst and bud set, vitally important adaptive traits for this long-lived forest species. Epigenetic memory is established in response to the temperature conditions during embryogenesis. Somatic embryogenesis at different epitype inducing (EpI temperatures closely mimics the natural processes of epigenetic memory formation in seeds, giving rise to epigenetically different clonal plants in a reproducible and predictable manner, with respect to altered bud phenology. MicroRNAs (miRNAs and other small non-coding RNAs (sRNAs play an essential role in the regulation of plant gene expression and may affect this epigenetic mechanism. We used NGS sequencing and computational in silico methods to identify and profile conserved and novel miRNAs among small RNAs in embryogenic tissues of Norway spruce at three EpI temperatures (18, 23 and 28°C. We detected three predominant classes of sRNAs related to a length of 24 nt, followed by a 21–22 nt class and a third 31 nt class of sRNAs. More than 2100 different miRNAs within the prevailing length 21–22 nt were identified. Profiling these putative miRNAs allowed identification of 1053 highly expressed miRNAs, including 523 conserved and 530 novels. 654 of these miRNAs were found to be differentially expressed (DEM depending on EpI temperature. For most DEMs, we defined their putative mRNA targets. The targets represented mostly by transcripts of multiple-repeats proteins, like TIR, NBS-LRR, PPR and TPR repeat, Clathrin/VPS proteins, Myb-like, AP2, etc. Notably, 124 DE miRNAs targeted 203 differentially expressed epigenetic regulators. Developing Norway spruce embryos possess a more complex sRNA structure than that reported for somatic tissues. A variety of the predicted miRNAs showed distinct EpI temperature dependent expression patterns. These putative EpI miRNAs target spruce genes with a wide range of functions, including genes known to be

  8. THE GEOMORPHOLOGIC FEATURES OF INTRUSIVE MAGMATIC STRUCTURES FROM BÂRGĂU MOUNTAINS (EASTERN CARPATHIANS, ROMANIA

    Directory of Open Access Journals (Sweden)

    Ioan Bâca

    2016-08-01

    Full Text Available Igneous intrusive structures from Bârgău Mountains belong to the group of central Neogene volcanic chain of the Eastern Carpathians of Romania. The evolution of the relief developed on these structures are three main stages: the stage of injection of structures (Pannonian, the stage of uncovering of igneous intrusive bodies from Oligo-Miocene sedimentary cover (Pliocene, and the stage of subaerial modeling of magmatic bodies (Pliocene-current.In those circumstances, the geodiversity of intrusive magmatic structures from Bârgău Mountains is represented by several types of landforms such as: polycyclic landforms (erosional levels, structural landforms (the configuration of igneous intrusive structures, petrographic landforms (andesites, lithological contact, fluvial landforms (valleys, slopes, ridges, periglacial landforms (cryogenic and crionival landforms, biogenic and anthropogenic landforms. This study highlights certain features of the landforms modeled on igneous intrusive bodies with the aim of developing some strategy for tourism recovery by local and county authorities.

  9. Effects of warming on the structure and function of a boreal black spruce forest

    Energy Technology Data Exchange (ETDEWEB)

    Stith T.Gower

    2010-03-03

    A strong argument can be made that there is a greater need to study the effect of warming on boreal forests more than on any other terrestrial biome. Boreal forests, the second largest forest biome, are predicted to experience the greatest warming of any forest biome in the world, but a process-based understanding of how warming will affect the structure and function of this economically and ecologically important forest biome is lacking. The effects of warming on species composition, canopy structure and biogeochemical cycles are likely to be complex; elucidating the underlying mechanisms will require long-term whole-ecosystem manipulation to capture all the complex feedbacks (Shaver et al. 2000, Rustad et al. 2001, Stromgren 2001). The DOE Program for Ecosystem Research funded a three year project (2002-2005) to use replicated heated chambers on soil warming plots in northern Manitoba to examine the direct effects of whole-ecosystem warming. We are nearing completion of our first growing season of measurements (fall 2004). In spite of the unforeseen difficulty of installing the heating cable, our heating and irrigation systems worked extremely well, maintaining environmental conditions within 5-10% of the specified design 99% of the time. Preliminary data from these systems, all designed and built by our laboratory at the University of Wisconsin, support our overall hypothesis that warming will increase the carbon sink strength of upland boreal black spruce forests. I request an additional three years of funding to continue addressing the original objectives: (1) Examine the effect of warming on phenology of overstory, understory and bryophyte strata. Sap flux systems and dendrometer bands, monitored by data loggers, will be used to quantify changes in phenology and water use. (2) Quantify the effects of warming on nitrogen and water use by overstory, understory and bryophytes. (3) Compare effects of warming on autotrophic respiration and above- and belowground

  10. Transformation of solar radiation in Norway spruce stands into produced biomass - the effect of stand density

    International Nuclear Information System (INIS)

    Marková, I.; Marek, M.V.; Pokorný, R.

    2011-01-01

    The present paper is focused on the assessment of the effects of stand density and leaf area development on radiation use efficiency in the mountain cultivated Norway spruce stand. The young even-aged (17-years-old in 1998) plantation of Norway spruce was divided into two experimental plots differing in their stand density in 1995. During the late spring of 2001 next cultivating high-type of thinning of 15% intensity in a reduction of stocking density was performed. The PAR regime of investigated stands was continually measured since 1992. Total aboveground biomass (TBa) and TBa increment were obtained on the basis of stand inventory. The dynamic of LAI development showed a tendency to be saturated, i.e. the LAI value close to 11 seems to be maximal for the local conditions of the investigated mountain cultivated Norway spruce stand in the Beskids Mts. Remarkable stimuli (up to 17%) of LAI formation were started in 2002, i.e. as an immediate response to thinning. Thus, the positive effect of thinning on LAI increase was confirmed. The data set of absorbed PAR and produced TBa in the period 1998-2003 was processed by the linear regression of Monteith's model, which provided the values of the coefficient of solar energy conversion efficiency into biomass formation. The differences in biomass formation values between the dense and sparse plot after thinning amounted to 18%

  11. Stem volume losses in grand firs topkilled by western spruce budworm in Idaho

    Science.gov (United States)

    George T. Ferrell; Robert F. Scharpf

    1982-01-01

    Mature grand firs (Abies grandis [Dougl. ex D. Don] Lindl.) were sampled in two stands, one cutover and one virgin, in the Little Salmon River drainage in west-central Idaho, to estimate stem volume losses associated with topkilling. Damage to the stands resulted from three outbreaks of western spruce budworm (Choristoneura occidentalis...

  12. Root uptake of lead by Norway spruce grown on Pb-210 spiked soils

    DEFF Research Database (Denmark)

    Hovmand, M.F.; Nielsen, Sven Poul; Johnsen, I.

    2009-01-01

    The root uptake of lead (Pb) by trees and the transfer of Pb by leaf litter deposition to the forest floor were investigated through a pot experiment with Norway spruce. Natural Pb and radio isotopic lead (210Pb) were determined in needles and twigs and in the pot soil spiked with 210Pb...

  13. Genetic variability and health of Norway spruce stands in the Regional Directorate of the State Forests in Krosno

    Directory of Open Access Journals (Sweden)

    Gutkowska Justyna

    2017-03-01

    Full Text Available The study was conducted in 2015 in six spruce stands situated in different forest districts administratively belonging to the Regional Directorate of State Forests in Krosno. Each spruce population was represented by 30 trees and assessed in terms of their current health status. Genetic analyses were performed on shoot samples from each tree using nine nuclear DNA markers and one mitochondrial DNA marker (nad1. The health status of the trees was described according to the classification developed by Szczepkowski and Tarasiuk (2005 and the correlation between health classes and the level of genetic variability was computed with STATISTICA (α = 0.05.

  14. Seasonal courses of photosynthetic parameters in sun- and shade-acclimated spruce shoots

    Czech Academy of Sciences Publication Activity Database

    Urban, Otmar; Holub, Petr; Klem, Karel

    2017-01-01

    Roč. 10, 1-2 (2017), s. 49-56 ISSN 1803-2451 R&D Projects: GA MŠk(CZ) LO1415 Institutional support: RVO:86652079 Keywords : CO2 assimilation * gas-exchange * Norway spruce * ontogeny * temperature * vapour pressure deficit * vertical canopy profile Subject RIV: EH - Ecology, Behaviour OBOR OECD: Environmental sciences (social aspects to be 5.7) https://beskydy.mendelu.cz/10/1/0049/

  15. Management intensity affects traits of soil microarthropod community in montane spruce forest

    Czech Academy of Sciences Publication Activity Database

    Farská, Jitka; Prejzková, Kristýna; Rusek, Josef

    2014-01-01

    Roč. 75, March (2014), s. 71-79 ISSN 0929-1393 R&D Projects: GA ČR GA526/03/1259; GA ČR GAP504/12/1218; GA MŠk LC06066 Grant - others:GAJU(CZ) 143/2010/P Institutional support: RVO:60077344 Keywords : Oribatida * Collembola * spruce forest * trait * management intensity Subject RIV: EH - Ecology, Behaviour Impact factor: 2.644, year: 2014

  16. Phenotypic evidence suggests a possible major-gene element to weevil resistance in Sitka spruce

    Science.gov (United States)

    John N. King; René I. Alfaro; Peter Ott; Lara vanAkker

    2012-01-01

    The weevil resistance breeding program against the white pine weevil, Pissodes strobi Peck (Coleoptera: Curculionidae), particularly for Sitka spruce (Picea sitchensis (Bong.) Carr), is arguably one of the most successful pest resistance breeding programs for plantation forest species, and it has done a lot to rehabilitate...

  17. Soluble and cell wall-bound phenolics and lignin in Ascocalyx abietina infected Norway spruces

    Czech Academy of Sciences Publication Activity Database

    Cvikrová, Milena; Malá, J.; Hrubcová, Marie; Eder, Josef

    2006-01-01

    Roč. 170, č. 3 (2006), s. 563-570 ISSN 0168-9452 R&D Projects: GA ČR GA206/04/0999 Institutional research plan: CEZ:AV0Z50380511 Keywords : Ascocalyx abietina (Lagerberg) * Lignin * Norway spruce Subject RIV: GK - Forestry Impact factor: 1.631, year: 2006

  18. Temporal and spatial heterogeneity of soil CO2 efflux in a Norway spruce stand

    Czech Academy of Sciences Publication Activity Database

    Kurajdová, Jana; Acosta, Manuel; Pavelka, Marian

    2006-01-01

    Roč. 2006, č. 19 (2006), s. 1 ISSN 1803-1013 R&D Projects: GA MŠk OC 627.001 Institutional research plan: CEZ:AV0Z60870520 Keywords : soil CO2 efflux * Norway spruce stand * temperature * spatial and temporal heterogeneity * stand density Subject RIV: ED - Physiology

  19. Seismic structure of the crust and upper mantle in central-eastern Greenland

    DEFF Research Database (Denmark)

    Kraft, Helene Anja

    Geophysical and geological knowledge of the interior of Greenland is very limited. The lack of knowledge arises mainly due to the logistical challenges related to conducting geophysical fieldwork on the up to 3400 m thick ice sheet, which covers around 80% of the land area. This PhD thesis is based...... on the very first regional passive seismic study in central-Eastern Greenland, focusing on the area between Scoresby Sund and Summit. The study aims to image the structure of subsurface Greenland starting from the crust and down to the mantle transition zone. Furthermore, the thesis links these observations....... The receiver functions were jointly inverted for the velocity structure of the crust and delay times, and shapes of signals originating at the mantle transition zone discontinuities, P410s and P660s, were analysed. The crustal models show a deepening of the Moho from east to west from less than 20 km depth...

  20. Differences in the photosynthetic UV-B response between European beech (Fagus sylvatica L.) and Norway spruce (Picea abies (L.) Karst.) saplings

    International Nuclear Information System (INIS)

    Šprtová, M.; Marek, M.V.; Urban, O.; Kalina, J.; Špunda, V.

    2008-01-01

    Cloned saplings of Norway spruce (Picea abies (L) Karst.) and beech (Fagus sylvatica L.) (7 years old) were exposed to enhanced UV-B irradiation (+25%) continuously over three growing seasons (1999-2001). Selected parameters of variable chlorophyll alpha fluorescence and pigment composition were analysed in the late summer of the third growing season to evaluate the influence of long-term elevated UV-B irradiation on broadleaf and conifer tree species. To obtain information on the xanthophyll cycle, the de-epoxidation state (DEPS) was calculated. These tree species responded differentially to the long-term effects of enhanced UV-B radiation, Norway spruce was more sensitive compared to the European beech. The results show that in Norway spruce long-term exposure to enhanced UV-B radiation under field conditions caused negative changes at the level of primary photosynthetic reactions. Contrary to the beech, this had higher degree of UV-B protective responses. UV-B radiation is not effective stressor to its primary photosynthetic reaction

  1. Ecosystem warming does not affect photosynthesis or aboveground autotrophic respiration for boreal black spruce

    Energy Technology Data Exchange (ETDEWEB)

    Bronson, D.R. [Wyoming Univ., Laramie, WY (United States). Dept. of Renewable Resources; Gower, S.T. [Wisconsin Univ., Madison, WI (United States). Dept. of Forest Ecology and Management

    2010-04-15

    Substantial increases in climatic temperatures may cause boreal forests to become a carbon source. An improved understanding of the effect of climatic warming on photosynthesis and autotrophic respiration is needed in order to determine the impact of temperature increases on net carbon balances. This study measured the light-saturated photosynthesis foliage respiration and stem respiration of black spruce in heated and control plots during a 3-year period at a site located in Thompson, Manitoba. Greenhouses and soil-heating cables were used to maintain air and soil temperatures at 5 degrees C above ambient air and soil temperatures. Studies were conducted to determine the influence of soil and air warming; soil-only warming; and greenhouses maintained at ambient temperatures. The study showed that treatment differences for photosynthesis, foliage respiration, and stem respiration were not significant over the 3-year period. Results suggested that black spruce may not have significant changes in photosynthesis or respiration rates in warmer climates. 38 refs., 3 tabs., 4 figs.

  2. Environmental protection in Eastern Europe

    International Nuclear Information System (INIS)

    Kabala, S.

    1990-01-01

    There is a need for the development of institutional structures to make the transfer of technology work to improve the ability of Eastern Europe to find solutions to its environmental problems. Envisioned is technical assistance: U.S. experts who will work on-site with Eastern European experts. The idea is to technically train individuals in pollution-prevention methods. Trained experts could then upgrade processes to save input, energy, and materials. In the exchange of environmental information, discussions have led to four issues: it is expensive to transfer equipment and make qualified personnel available for a long period; information is comparatively inexpensive to convey; in Eastern Europe there are trained and competent personnel; the theoretical knowledge, academic knowledge, and education are at a very high level but little of this knowledge has been put into practice. The technology transfer goal is to develop a response to needs identified by partner institutions and counterpart professionals that will enable Eastern Europe to tap resources that do exist- scientific, managerial, and economic resources and tap information resources in the US in order to address the environmental problems that exist in Eastern Europe

  3. Monitoring western spruce budworm with pheromone- baited sticky traps to predict subsequent defoliation

    Science.gov (United States)

    Christine G. Niwa; David L. Overhulser

    2015-01-01

    A detailed procedure is described for monitoring western spruce budworm with pheromone-baited sticky traps and interpreting the results to predict defoliation the following year. Information provided includes timing of the survey, how to obtain traps and baits, how many traps are needed, trap assembly, field placement of traps, and how to evaluate the catches.

  4. Higher fine-scale genetic structure in peripheral than in core populations of a long-lived and mixed-mating conifer--eastern white cedar (Thuja occidentalis L.).

    Science.gov (United States)

    Pandey, Madhav; Rajora, Om P

    2012-04-05

    Fine-scale or spatial genetic structure (SGS) is one of the key genetic characteristics of plant populations. Several evolutionary and ecological processes and population characteristics influence the level of SGS within plant populations. Higher fine-scale genetic structure may be expected in peripheral than core populations of long-lived forest trees, owing to the differences in the magnitude of operating evolutionary and ecological forces such as gene flow, genetic drift, effective population size and founder effects. We addressed this question using eastern white cedar (Thuja occidentalis) as a model species for declining to endangered long-lived tree species with mixed-mating system. We determined the SGS in two core and two peripheral populations of eastern white cedar from its Maritime Canadian eastern range using six nuclear microsatellite DNA markers. Significant SGS ranging from 15 m to 75 m distance classes was observed in the four studied populations. An analysis of combined four populations revealed significant positive SGS up to the 45 m distance class. The mean positive significant SGS observed in the peripheral populations was up to six times (up to 90 m) of that observed in the core populations (15 m). Spatial autocorrelation coefficients and correlograms of single and sub-sets of populations were statistically significant. The extent of within-population SGS was significantly negatively correlated with all genetic diversity parameters. Significant heterogeneity of within-population SGS was observed for 0-15 m and 61-90 m between core and peripheral populations. Average Sp, and gene flow distances were higher in peripheral (Sp = 0.023, σg = 135 m) than in core (Sp = 0.014, σg = 109 m) populations. However, the mean neighborhood size was higher in the core (Nb = 82) than in the peripheral (Nb = 48) populations. Eastern white cedar populations have significant fine-scale genetic structure at short distances. Peripheral populations have several

  5. Oxyfluorfen safe to use engelmann spruce seedbeds. Forest Service research note

    Energy Technology Data Exchange (ETDEWEB)

    Sloan, J.P.

    1993-06-01

    Oxyfluorfen, a diphenylether herbicide, can be applied to Engelmann spruce nursery beds without significant damage to seedlings. Oxyfluorfen, applied at 0.5 lb/acre with preemergence timing for two years, reduced seedling dry mass. When the rate of herbicide application was reduced, the timing was delayed, or the applications were discontinued the second year, there was little damage to seedlings. Five of 10 herbicide treatments significantly reduced seedling densities compared to the no-treatment plots.

  6. Interaction of soil filamentous fungi affects needle composition and nutrition of Norway spruce seedlings

    Czech Academy of Sciences Publication Activity Database

    Mrnka, Libor; Tokárová, H.; Vosátka, Miroslav; Matějka, P.

    2009-01-01

    Roč. 23, č. 5 (2009), s. 887-897 ISSN 0931-1890 R&D Projects: GA ČR GA206/05/0269 Institutional research plan: CEZ:AV0Z60050516 Keywords : mycelial interaction s * norway spruce * nutrient cycling Subject RIV: EF - Botanics Impact factor: 1.603, year: 2009

  7. Finite-Frequency Seismic Tomography of Body Waves and Surface Waves from Ambient Seismic Noise: Crustal and Mantle Structure Beneath Eastern Eurasia

    National Research Council Canada - National Science Library

    Ren, Yong; Zhang, Wei; Yang, Ting; Shen, Yang; Yang, Xiaoping

    2008-01-01

    To improve seismic calibration for nuclear explosion monitoring, we use 3D sensitivity kernels of finite-frequency body and surface waves to develop models of the crustal and mantle structures beneath eastern Eurasia...

  8. H and Al ionic toxicity in seedlings of spruce and beech trees

    Energy Technology Data Exchange (ETDEWEB)

    Rost-Siebert, K.

    1985-01-01

    The influence of aluminium and hydrogen on seedlings of spruce and beech trees was tested. Parameters as root growth, state of health, minerals content and production of dry matter were measured. The results are discussed and show that ion concentrations of hydrogen and aluminium, average for todays forest soils, reduce root growth, damage roots and disturb the uptake of Ca/sup 2+/ and Mg/sup 2+/.

  9. The rheological structure of the lithosphere in the Eastern Marmara region, Turkey

    Science.gov (United States)

    Oruç, Bülent; Sönmez, Tuba

    2017-05-01

    The aim of this work is to propose the geometries of the crustal-lithospheric mantle boundary (Moho) and lithosphere-asthenosphere boundary (LAB) and the 1D thermal structure of the lithosphere, in order to establish a rheological model of the Eastern Marmara region. The average depths of Moho and LAB are respectively 35 km and 51 km from radially averaged amplitude spectra of EGM08 Bouguer anomalies. The geometries of Moho and LAB interfaces are estimated from the Parker-Oldenburg gravity inversion algorithm. Our results show the Moho depth varies from 31 km at the northern part of North Anatolian Fault Zone (NAFZ) to 39 km below the mountain belt in the southern part of the NAFZ. The depth to the LAB beneath the same parts of the region ranges from 45 km to 55 km. Having lithospheric strength and thermal boundary layer structure, we analyzed the conditions of development of lithosphere thinning. A two-dimensional strength profile has been estimated for rheology model of the study area. Thus we suggest that the rheological structure consists of a strong upper crust, a weak lower crust, and a partly molten upper lithospheric mantle.

  10. New method for diagnosis of smoke damage to spruce

    Energy Technology Data Exchange (ETDEWEB)

    Haertel, O

    1953-01-01

    Spruce needles excrete more wax in smoke regions than in smoke-free areas. This increased wax excretion can be used as a measure to determine the effects of smoke on the plant. The wax is extracted with hot water and is preserved after cooling in the form of turbidity which can be determined photometrically. Good correlations between the degree of turbidity and the distance from the source of smoke are obtained. The low scatter of values, especially in smoke-free regions (+/-1%) makes the turbidity most useful in outlining the limits of the areas affected by smoke. 11 references, 5 tables.

  11. Crustal Structure and Evolution of the Eastern Himalayan Plate Boundary System, Northeast India

    Science.gov (United States)

    Mitra, S.; Priestley, K. F.; Borah, Kajaljyoti; Gaur, V. K.

    2018-01-01

    We use data from 24 broadband seismographs located south of the Eastern Himalayan plate boundary system to investigate the crustal structure beneath Northeast India. P wave receiver function analysis reveals felsic continental crust beneath the Brahmaputra Valley, Shillong Plateau and Mikir Hills, and mafic thinned passive margin transitional crust (basement layer) beneath the Bengal Basin. Within the continental crust, the central Shillong Plateau and Mikir Hills have the thinnest crust (30 ± 2 km) with similar velocity structure, suggesting a unified origin and uplift history. North of the plateau and Mikir Hills the crustal thickness increases sharply by 8-10 km and is modeled by ˜30∘ north dipping Moho flexure. South of the plateau, across the ˜1 km topographic relief of the Dawki Fault, the crustal thickness increases abruptly by 12-13 km and is modeled by downfaulting of the plateau crust, overlain by 13-14 km thick sedimentary layer/rocks of the Bengal Basin. Farther south, beneath central Bengal Basin, the basement layer is thinner (20-22 km) and has higher Vs (˜4.1 km s-1) indicating a transitional crystalline crust, overlain by the thickest sedimentary layer/rocks (18-20 km). Our models suggest that the uplift of the Shillong Plateau occurred by thrust faulting on the reactivated Dawki Fault, a continent margin paleorift fault, and subsequent back thrusting on the south dipping Oldham Fault, in response to flexural loading of the Eastern Himalaya. Our estimated Dawki Fault offset combined with timing of surface uplift of the plateau reveals a reasonable match between long-term uplift and convergence rate across the Dawki Fault with present-day GPS velocities.

  12. Sulfur Speciation in Peat: a Time-zero Signature for the " Spruce and Peatland Responses Under Climate and Environmental Change" Experiment

    Science.gov (United States)

    Furman, O.; Toner, B. M.; Sebestyen, S. D.; Kolka, R. K.; Nater, E. A.

    2014-12-01

    As part of the "Spruce and Peatland Responses Under Climate and Environmental Change" (SPRUCE) experiment, we made initial measurements of sulfur speciation in peat. These observations represent a "time-zero" relative to the intended soil warming experiment which begins in 2015. Total sulfur and sulfur speciation were measured in peat cores (solid phase) from nine plots (hollows and hummocks) to a depth of 2 m. Peat samples were packed under nitrogen and frozen in the field immediately after collection. All subsequent sample storage, handling, and processing were conducted under inert gas. Sulfur speciation was measured using bulk sulfur 1s X-ray absorption near edge structure (XANES) spectroscopy at the SXRMB instrument at the Canadian Light Source, Saskatoon, SK, Canada and at the 9-BM instrument, Advanced Photon Source, Argonne National Laboratory, IL, USA. Total sulfur concentrations ranged from 968 to 4077 mg sulfur / kg dry peat. Sulfur content increased with depth from 2 g sulfur / m2 in the 0-10 cm increment to a maximum value of 38 g sulfur / m2 in the 50-60 cm increment. These sulfur loadings produced high quality XANES spectra. The nine cores exhibited reproducible trends with depth in both total sulfur and specific sulfur species; however, variability in sulfur speciation was greatest in the top 40 cm. All sulfur detected within the peat solids was in an organic form. The most abundant sulfur species group was composed of organic mono-sulfide and thiol forms, representing approximately half of the total sulfur at all depths. Sulfonate and ester-sulfate species were 10-15 mol% of sulfur and exhibited low variability with depth. A subsurface maximum in organic di-sulfide was observed in the 20-30 cm depth increment, which is the transition zone between transiently oxidized acrotelm and permanently saturated anaerobic catotelm. Quantification of major sulfur pools is important for the SPRUCE experiment as they are likely to be indicators of changes in the

  13. Modelling long-term water yield effects of forest management in a Norway spruce forest

    Czech Academy of Sciences Publication Activity Database

    Yu, X.; Lamačová, A.; Duffy, Ch.; Krám, P.; Hruška, Jakub; White, T.; Bhatt, G.

    2015-01-01

    Roč. 60, č. 2 (2015), s. 174-191 ISSN 0262-6667 R&D Projects: GA MŠk(CZ) ED1.1.00/02.0073 Institutional support: RVO:67179843 Keywords : Lysina critical zone observatory * PIHM * Norway spruce * forest management Subject RIV: EH - Ecology, Behaviour Impact factor: 2.182, year: 2015

  14. Haloperoxidase-like activity in spruce forest soil. A source of volatile halogenated organic compounds?

    DEFF Research Database (Denmark)

    Laturnus, F.; Mehrtens, G.; Grøn, C.

    1995-01-01

    Haloperoxidase-like activity was monitored in samples from a podzol soil in an uncontaminated spruce forest at Klosterhede, Denmark. Activity for the oxidation of chloride and bromide was found. The pH optima for chlorination and bromination ranged between pH 2.5 and 4: Very high activity, up to 4...

  15. The influence of moisture content on the water vapour resistance of surface coated spruce

    DEFF Research Database (Denmark)

    Engelund, E.T.; Ulriksen, L.; Hansen, Kurt Kielsgaard

    2005-01-01

    Two series of cup tests are carried out. The first series is performed on spruce specimens having moisture transport in either radial direction (R-direction) or in tangential direction (T-direction). The T-direction tests are made as wet cup tests having 93 %RH inside the cups, while the R-direct...

  16. The performance of moss, grass, and 1- and 2-year old spruce needles as bioindicators of contamination: a comparative study at the scale of the Czech Republic.

    Science.gov (United States)

    Suchara, Ivan; Sucharova, Julie; Hola, Marie; Reimann, Clemens; Boyd, Rognvald; Filzmoser, Peter; Englmaier, Peter

    2011-05-01

    Moss (Pleurozium schreberi), grass (Avenella flexuosa), and 1- and 2-year old spruce (Picea abies) needles were collected over the territory of the Czech Republic at an average sample density of 1 site per 290km(2). The samples were analysed for 39 elements (Ag, Al, As, Ba, Be, Bi, Ca, Cd, Ce, Co, Cr, Cs, Cu, Fe, Ga, Hg, K, La, Li, Mg, Mn, Mo, Na, Nd, Ni, Pb, Pr, Rb, S, Sb, Se, Sn, Sr, Th, Tl, U, V, Y and Zn) using ICP-MS and ICP-AES techniques (the major nutrients Ca, K, Mg and Na were not analysed in moss). Moss showed by far the highest element concentrations for most elements. Exceptions were Ba (spruce), Mn (spruce), Mo (grass), Ni (spruce), Rb (grass) and S (grass). Regional distribution maps and spatial trend analysis were used to study the suitability of the four materials as bioindicators of anthropogenic contamination. The highly industrialised areas in the north-west and the far east of the country and several more local contamination sources were indicated in the distribution maps of one or several sample materials. At the scale of the whole country moss was the best indicator of known contamination sources. However, on a more local scale, it appeared that spruce needles were especially well suited for detection of urban contamination. Copyright © 2010 Elsevier B.V. All rights reserved.

  17. The ''red'' decline of Norway spruce or ''røde rødgraner'' - is it ammonium overload or top-dying?

    DEFF Research Database (Denmark)

    Engvild, K.C.

    2005-01-01

    by nitrogen overload was investigated. A sensitive biochemical indicator of nitrogen status of conifer trees is the content of free amino acids of the urea cycle,arginine and ornithine. In pot experiments with four times overload of N as ammonium nitrate the free arginine content increased more than 100 times....... There was no indication of nitrogen overload. The “red” Norway spruce maysuffer from “top-dying” a common disorder of Norway spruce in Great Britain, believed to be caused by several mild winters in a row. In that case the symptoms should diminish after the very cold winter 1995-96....

  18. Tomographic Imaging of the Seismic Structure Beneath the East Anatolian Plateau, Eastern Turkey

    Science.gov (United States)

    Gökalp, Hüseyin

    2012-10-01

    The high level of seismic activity in eastern Turkey is thought to be mainly associated with the continuing collision of the Arabian and Eurasian tectonic plates. The determination of a detailed three-dimensional (3D) structure is crucial for a better understanding of this on-going collision or subduction process; therefore, a body wave tomographic inversion technique was performed on the region. The tomographic inversion used high quality arrival times from earthquakes occurring in the region from 1999 to 2001 recorded by a temporary 29 station broadband IRIS-PASSCAL array operated by research groups from the Universities of Boğaziçi (Turkey) and Cornell (USA). The data was inverted and consisted of 3,114 P- and 2,298 S-wave arrival times from 252 local events with magnitudes ( M D) ranging from 2.5 to 4.8. The stability and resolution of the results were qualitatively assessed by two synthetic tests: a spike test and checkerboard resolution test and it was found that the models were well resolved for most parts of the imaged domain. The tomographic inversion results reveal significant lateral heterogeneities in the study area to a depth of ~20 km. The P- and S-wave velocity models are consistent with each other and provide evidence for marked heterogeneities in the upper crustal structure beneath eastern Turkey. One of the most important features in the acquired tomographic images is the high velocity anomalies, which are generally parallel to the main tectonic units in the region, existing at shallow depths. This may relate to the existence of ophiolitic units at shallow depths. The other feature is that low velocities are widely dispersed through the 3D structure beneath the region at deeper crustal depths. This feature can be an indicator of the mantle upwelling or support the hypothesis that the Anatolian Plateau is underlain by a partially molten uppermost mantle.

  19. Calcium addition at the Hubbard Brook Experimental Forest increases the capacity for stress tolerance and carbon capture in red spruce (Picea rubens) trees during the cold season

    Science.gov (United States)

    Paul G. Schaberg; Rakesh Minocha; Stephanie Long; Joshua M. Halman; Gary J. Hawley; Christopher. Eagar

    2011-01-01

    Red spruce (Picea rubens Sarg.) trees are uniquely vulnerable to foliar freezing injury during the cold season (fall and winter), but are also capable of photosynthetic activity if temperatures moderate. To evaluate the influence of calcium (Ca) addition on the physiology of red spruce during the cold season, we measured concentrations of foliar...

  20. Wood Anatomy and Insect Defoliator Systems: Is there an anatomical response to sustained feeding by the western spruce budworm (Choristoneura occidentalis) on Douglas-fir (Pseudotusga menziesii)?

    Science.gov (United States)

    Axelson, Jodi; Gärtner, Holger; Alfaro, René; Smith, Dan

    2013-04-01

    The western spruce budworm (Choristoneura occidentalis Freeman) is the most widespread and destructive defoliator of coniferous forests in western North America, and has a long-term coexistence with its primary host tree, Douglas-fir (Pseudotsuga menziesii Franco). Western spruce budworm (WSB) outbreaks usually last for several years, and cause reductions in annual growth, stem defects, and regeneration delays. In British Columbia, the WSB is the second most damaging insect after the mountain pine beetle, and sustained and/or severe defoliation can result in the mortality of host trees. Numerous studies have used tree rings to reconstruct WSB outbreaks across long temporal scales, to evaluate losses in stand productivity, and examine isotope ratios. Although some studies have looked at the impacts of artificial defoliation on balsam fir in eastern North America, there has been no prior research on how WSB outbreaks affect the anatomical structure of the stem as described by intra-annual wood density and potential cell size variations. The objective of this study was to anatomically examine the response of Douglas-fir to sustained WSB outbreaks in two regions of southern British Columbia. We hypothesize that the anatomical intra-annual characteristics of the tree rings, such as cell wall thickness, latewood cell size, and/or lumen area changes during sustained WSB outbreaks. To test this hypothesis we sampled four permanent sample plots in coastal and dry interior sites, which had annually resolved defoliation data collected over a 7-12 year period. At each site diameter-at-breast height (cm), height (m), and crown position were recorded and three increment cores were extracted from 25 trees. Increment cores were prepared to permit anatomical and x-ray density analyses. For each tree, a 15µm thick micro section was cut from the radial plane. Digital images of the micro sections were captured and processed. In each annual ring, features such as cell lumen area (µm2

  1. Intercontinental and intracontinental biogeography of the eastern Asian - Eastern North American disjunct Panax (the ginseng genus, Araliaceae), emphasizing its diversification processes in eastern Asia.

    Science.gov (United States)

    Zuo, Yun-Juan; Wen, Jun; Zhou, Shi-Liang

    2017-12-01

    The intercontinental biogeography between eastern Asia and eastern North America has attracted much attention from evolutionary biologists. Further insights into understanding the evolution of the intercontinental disjunctions have been hampered by the lack of studies on the intracontinental biogeography in eastern Asia, a region with complex geology, geography, climates and habitats. Herein we studied the biogeographic history of the eastern Asian-eastern North American disjunct genus Panax with special emphasis on the investigation of its uneven diversification in Asia. This study reconstructs the diversification history of Panax and also emphasizes a large clade of Panax taxa, which has a wide distribution in eastern Asia, but was unresolved in previous studies. We examined the noncoding plastid DNA fragments of trnH-psbA, rps16, and psbM-trnD, the mitochondrial b/c intron of NAD1, and the nuclear ribosomal internal transcribed spacer (ITS) region of 356 samples from 47 populations. The results revealed the subtropical Northern Hemisphere origin (Asia or Asia and North America) of Panax in the Paleocene. Intercontinental disjunctions between eastern Asia and eastern North America formed twice in Panax, once estimated in early Eocene for the split of P. trifolius and another in mid-Miocene for the divergence of P. quinquefolius. Intercontinental diversifications in Panax showed temporal correlation with the increase of global temperature. The evolutionary radiation of the P. bipinnatifidus species complex occurred around the boundary of Oligocene and Miocene. Strong genetic structure among populations of the species complex was detected and the populations may be isolated by distance. The backbone network and the Bayesian clustering analysis revealed a major evolutionary radiation centered in the Hengduan Mountains of western China. Our results suggested that the evolutionary radiation of Panax was promoted by geographic barriers, including mountain ranges

  2. Efflux of CO2 from soil in Norway Spruce stands of different ages: a case study

    Czech Academy of Sciences Publication Activity Database

    Dařenová, Eva; Fabiánek, Tomáš; Pavelka, Marian

    2016-01-01

    Roč. 6, č. 2 (2016), s. 98-102 ISSN 1805-0174 R&D Projects: GA MŠk(CZ) LO1415 Institutional support: RVO:67179843 Keywords : spruce forest * Picea abies * soil temperatures * moisture * respiration Subject RIV: EH - Ecology, Behaviour

  3. Structural characteristic of the Eastern Plains soils of Colombia, submitted to several handling systems

    International Nuclear Information System (INIS)

    Amezquita, E; Saenz J I; Thomas, R J; Vera, R R; Hoyos, P; Molina, D L; Chavez, L F

    1997-01-01

    Soil productivity and sustainability depends on the building and/or conservation of an adequate and dynamic equilibrium between physical, chemical and biological properties and processes in the volume of soil explored by roots, so that there is no constraints in the availability of water and nutrients to plants. Soil structure is one of the soil properties that are more vulnerable to the intensity of use in tropical soils. Aggregate size distribution, aggregate stability and pore size distribution are some of the attributes that are usually used to describe structural changes and can act as indicators of structural sustainability. This paper presents and discusses the behavior of these attributes under different soil management treatments (native savanna, Brachiaria alone; Brachiaria + legume and monocrop) in the Colombian Eastern plains soil classified as Typic haplustox Kaolinitic iso-hyperthermic. These results showed highly statistical significant differences between treatments in the parameters studied and allow concluding that aggregate size distribution and stability could be indicators of susceptibility to degradation

  4. Individual variation of sap-flow rate in large pine and spruce trees and stand transpiration: a pilot study at the central NOPEX site

    Science.gov (United States)

    Čermák, J.; Cienciala, E.; Kučera, J.; Lindroth, A.; Bednářová, E.

    1995-06-01

    Transpiration in a mixed old stand of sub-boreal forest in the Norunda region (central Sweden) was estimated on the basis of direct measurement of sap flow rate in 24 large Scots pine and Norway spruce trees in July and August 1993. Sap flow rate was measured using the trunk tissue heat balance method based on internal (electric) heating and sensing of temperature. Transpiration was only 0.7 mm day -1 in a relatively dry period in July (i.e. about 20% of potential evaporation) and substantially higher after a rainy period in August. The error of the estimates of transpiration was higher during a dry period (about 13% and 22% in pine and spruce, respectively) and significantly lower (about 9% in both species) during a period of sufficient water supply. Shallow-rooted spruce trees responded much faster to precipitation than deeply rooted pines.

  5. Seasonal changes in camera-based indices from an open canopy black spruce forest in Alaska, and comparison with indices from a closed canopy evergreen coniferous forest in Japan

    Science.gov (United States)

    Nagai, Shin; Nakai, Taro; Saitoh, Taku M.; Busey, Robert C.; Kobayashi, Hideki; Suzuki, Rikie; Muraoka, Hiroyuki; Kim, Yongwon

    2013-06-01

    Evaluation of the carbon, water, and energy balances in evergreen coniferous forests requires accurate in situ and satellite data regarding their spatio-temporal dynamics. Daily digital camera images can be used to determine the relationships among phenology, gross primary productivity (GPP), and meteorological parameters, and to ground-truth satellite observations. In this study, we examine the relationship between seasonal variations in camera-based canopy surface indices and eddy-covariance-based GPP derived from field studies in an Alaskan open canopy black spruce forest and in a Japanese closed canopy cedar forest. The ratio of the green digital number to the total digital number, hue, and GPP showed a bell-shaped seasonal profile at both sites. Canopy surface images for the black spruce forest and cedar forest mainly detected seasonal changes in vegetation on the floor of the forest and in the tree canopy, respectively. In contrast, the seasonal cycles of the ratios of the red and blue digital numbers to the total digital numbers differed between the two sites, possibly due to differences in forest structure and leaf color. These results suggest that forest structural characteristics, such as canopy openness and seasonal forest-floor changes, should be considered during continuous observations of phenology in evergreen coniferous forests.

  6. Higher acid-chlorite reactivity of cell corner middle lamella lignin in black spruce

    Science.gov (United States)

    Umesh P. Agarwal

    2007-01-01

    To determine if there was a delignification behavior difference between secondary wall (S2) and middle lamella (cell corner or CC) lignin, black spruce cross-sections were acid-chlorite delignified and the tissue was evaluated in-situ by Raman imaging. Lignin concentration in S2 and CC was determined in numerous latewood cell areas in the two hour delignified cross...

  7. Impact of elevated carbon dioxide concentration on carbohydrate content in Norway spruce needles

    Czech Academy of Sciences Publication Activity Database

    Cabálková, Jana; Chmelík, Josef

    2005-01-01

    Roč. 99, S (2005), s552-s553 ISSN 0009-2770. [Meeting on Chemistry and Life /3./. Brno, 20.09.2005-22.09.2005] R&D Projects: GA ČR GA526/03/1182 Institutional research plan: CEZ:AV0Z40310501 Keywords : Norway spruce * needles * carbohydrates Subject RIV: CB - Analytical Chemistry, Separation Impact factor: 0.445, year: 2005

  8. Ash recycling to spruce and beech stands effects on nutrients, growth, nitrogen dynamics and carbon balance; Askaaterfoering till gran- och bokbestaand - effekter paa naering, tillvaext, kvaevedynamik och kolbalans

    Energy Technology Data Exchange (ETDEWEB)

    Thelin, Gunnar

    2006-03-15

    Ash recycling is an important part in a modern, sustainable forestry, especially in whole-tree harvest systems. Nutrients lost at harvest are returned to the forest with the wood-ash. In the project the effects of ash treatment on needle and leaf chemistry, tree growth, soil chemistry, soil water chemistry, and carbon and nitrogen dynamics were studied on 23 Norway spruce sites in south-western Sweden and in ten European beech sites in Scania, southern Sweden. On some of the sites there were previously established ash recycling experiments, but on a majority of the sites ash recycling was performed without experimental lay-out and ash and control plots were established afterwards. The most common dose was two tons of self hardened crushed wood-ash and two tons of Mg-lime. On average seven to eight years after ash recycling the results were 1. increased exchangeable stores of base cations in the soil in the beech and the spruce stands 2. increased base saturation in the beech and the spruce stands and increased BC/Al in the spruce stands 3. increased concentrations and ratios to N of P, Ca, Zn, and S in the needles, the increased P-values are especially important since P is close to or below deficiency levels in a majority of the spruce stands 4. decreased K-concentration in the beech leaves 5. increased tree growth with on average 14 % in the ash treated spruce stands compared to the control plots 6. increased carbon and nitrogen amounts in the biomass in the spruce stands 7. tendencies towards increased amounts of carbon and nitrogen in the soil in the beech stands and no effect in the soil in the spruce stands 8. increased concentrations of Ca, Mg, and SO{sub 4} and no effect on ANC in the soil water 9. no effect on potential net mineralization but increased potential nitrification rates 10. decreased concentration of nitrate in the soil water in the beech stands and no effect in the spruce stands 11. lower system N losses in the beech stands and possibly in the

  9. User's guide to the weather model: a component of the western spruce budworm modeling system.

    Science.gov (United States)

    W. P. Kemp; N. L. Crookston; P. W. Thomas

    1989-01-01

    A stochastic model useful in simulating daily maximum and minimum temperature and precipitation developed by Bruhn and others has been adapted for use in the western spruce budworm modeling system. This document describes how to use the weather model and illustrates some aspects of its behavior.

  10. Natural regeneration ecology of a secondary altimontane spruce forests at Jelendol

    International Nuclear Information System (INIS)

    Rozman, E.; Diaci, J.

    2008-01-01

    Natural regeneration of altimontane spruce forests at Jelendol is retarded due to many factors. In autumn 2003, gaps of different size and parts of the surrounding stand were covered with a 5 x 5 grid m to define sampling plots. A total of 227 plots with 1,5 x 1,5 m in size were installed to analyse general regeneration conditions and inhibitors. The following ecological parameters were estimated on each plot: micro relief, inclination, soil depth,ground cover, direct and diffuse solar radiation. Woody regeneration (density, height, height increment) and ground vegetation were recorded at each plot. Considering that N-S and E-W radiation asymmetry was explicit, the distribution of direct and diffuse radiation was divided into four groups among the plots. Spruce regeneration (28.,605 per ha) was mainly found at the edge of the large gap, though total regeneration density and radiation were not correlated. However, in both the stand and the small gaps, the lack of radiation hindered further development. This study showed that light conditions were not the only factors affecting the regeneration success at an altitude of 1,500 m. The presence of woody debris was important, while the influence of the herb layer (predominant species were Festuca altissima All. and Calamagrostis arundinacea (L.) Roth.) and soil depth proved to be negative. The impact of browsing, however, remained the main problem. (author)

  11. Carbon sequestration by young Norway spruce monoculture

    Science.gov (United States)

    Pokorny, R.; Rajsnerova, P.; Kubásek, J.

    2012-04-01

    Many studies have been focused on allometry, wood-mass inventory, carbon (C) sequestration, and biomass expansion factors as the first step for the evaluation of C sinks of different plant ecosystems. To identify and quantify these terrestrial C sinks, and evaluate CO2 human-induced emissions on the other hand, information for C balance accounting (for impletion of commitment to Kyoto protocol) are currently highly needed. Temperate forest ecosystems have recently been identified as important C sink. Carbon sink might be associated with environmental changes (elevated [CO2], air temperature, N deposition etc.) and large areas of managed fast-growing young forests. Norway spruce (Pice abies L. Karst) is the dominant tree species (35%) in Central European forests. It covers 55 % of the total forested area in the Czech Republic, mostly at high altitudes. In this contribution we present C sequestration by young (30-35 year-old) Norway spruce monocultures in highland (650-700 m a.s.l., AT- mean annual temperature: 6.9 ° C; P- annual amount of precipitation: 700 mm; GL- growing season duration: 150 days) and mountain (850-900 m a.s.l.; AT of 5.5 ° C; P of 1300 mm; and GL of 120 days) areas and an effect of a different type of thinning. However, the similar stem diameter at the breast height and biomass proportions among above-ground tree organs were obtained in the both localities; the trees highly differ in their height, above-ground organ's biomass values and total above ground biomass, particularly in stem. On the total mean tree biomass needle, branch and stem biomass participated by 22 %, 24 % and 54 % in highland, and by 19 %, 23 % and 58 % in mountain area, respectively. Silvicultural management affects mainly structure, density, and tree species composition of the stand. Therefore, dendrometric parameters of a tree resulted from genotype, growth conditions and from management history as well. Low type of thinning (LT; common in highland) stimulates rather tree

  12. Shear wave velocity structure of northern and North-Eastern Ethiopia

    International Nuclear Information System (INIS)

    Kebede, F.; Mammo, T.; Panza, G.F.; Vuan, A.; Costa, G.

    1995-10-01

    The non-linear inversion technique known as hedgehog is utilized to define the average crustal structure of North and North-Eastern Ethiopia. To accomplish the task a two dimensional frequency-time analysis is performed to obtain Rayleigh wave group velocity dispersion curves. Six earthquakes recorded by the broad-band digital seismograph installed at the Geophysical Observatory of Addis Ababa University are utilized. The crustal structure between the Gulf of Tadjura (western Gulf of Aden) and Addis Ababa crossing southern Afar (path I) can be approximated by a total thickness of about 22 km with average S-wave velocity in the range 2.3 - 3.9 km/s. The crust-mantle transition is poorly developed at greater depths and the shear wave velocity ranges from 4.0 km/s to 4.3 km/s. If the effect of the plateau part is taken into account the average total crustal thickness is found to be less than 18 km and the average S-wave velocity varies in the range 2.4 - 3.9 km/s. The low shear wave velocity under the Afar crust is consistent with the result of other geophysical studies. For path II, which passes through the border of the Western Ethiopian plateau, the average crustal structure is found to be approximated by a thickness of about 40 km and average S-wave velocity between 3.0 km/s and 3.9 km/s. The crust overlies a lithospheric mantle with a shear wave velocity in the range 4.1-4.4 km/s. (author). 37 refs, 11 figs, 4 tabs

  13. Higher fine-scale genetic structure in peripheral than in core populations of a long-lived and mixed-mating conifer - eastern white cedar (Thuja occidentalis L.)

    Science.gov (United States)

    2012-01-01

    Background Fine-scale or spatial genetic structure (SGS) is one of the key genetic characteristics of plant populations. Several evolutionary and ecological processes and population characteristics influence the level of SGS within plant populations. Higher fine-scale genetic structure may be expected in peripheral than core populations of long-lived forest trees, owing to the differences in the magnitude of operating evolutionary and ecological forces such as gene flow, genetic drift, effective population size and founder effects. We addressed this question using eastern white cedar (Thuja occidentalis) as a model species for declining to endangered long-lived tree species with mixed-mating system. Results We determined the SGS in two core and two peripheral populations of eastern white cedar from its Maritime Canadian eastern range using six nuclear microsatellite DNA markers. Significant SGS ranging from 15 m to 75 m distance classes was observed in the four studied populations. An analysis of combined four populations revealed significant positive SGS up to the 45 m distance class. The mean positive significant SGS observed in the peripheral populations was up to six times (up to 90 m) of that observed in the core populations (15 m). Spatial autocorrelation coefficients and correlograms of single and sub-sets of populations were statistically significant. The extent of within-population SGS was significantly negatively correlated with all genetic diversity parameters. Significant heterogeneity of within-population SGS was observed for 0-15 m and 61-90 m between core and peripheral populations. Average Sp, and gene flow distances were higher in peripheral (Sp = 0.023, σg = 135 m) than in core (Sp = 0.014, σg = 109 m) populations. However, the mean neighborhood size was higher in the core (Nb = 82) than in the peripheral (Nb = 48) populations. Conclusion Eastern white cedar populations have significant fine-scale genetic structure at short distances. Peripheral

  14. Higher fine-scale genetic structure in peripheral than in core populations of a long-lived and mixed-mating conifer - eastern white cedar (Thuja occidentalis L.

    Directory of Open Access Journals (Sweden)

    Pandey Madhav

    2012-04-01

    Full Text Available Abstract Background Fine-scale or spatial genetic structure (SGS is one of the key genetic characteristics of plant populations. Several evolutionary and ecological processes and population characteristics influence the level of SGS within plant populations. Higher fine-scale genetic structure may be expected in peripheral than core populations of long-lived forest trees, owing to the differences in the magnitude of operating evolutionary and ecological forces such as gene flow, genetic drift, effective population size and founder effects. We addressed this question using eastern white cedar (Thuja occidentalis as a model species for declining to endangered long-lived tree species with mixed-mating system. Results We determined the SGS in two core and two peripheral populations of eastern white cedar from its Maritime Canadian eastern range using six nuclear microsatellite DNA markers. Significant SGS ranging from 15 m to 75 m distance classes was observed in the four studied populations. An analysis of combined four populations revealed significant positive SGS up to the 45 m distance class. The mean positive significant SGS observed in the peripheral populations was up to six times (up to 90 m of that observed in the core populations (15 m. Spatial autocorrelation coefficients and correlograms of single and sub-sets of populations were statistically significant. The extent of within-population SGS was significantly negatively correlated with all genetic diversity parameters. Significant heterogeneity of within-population SGS was observed for 0-15 m and 61-90 m between core and peripheral populations. Average Sp, and gene flow distances were higher in peripheral (Sp = 0.023, σg = 135 m than in core (Sp = 0.014, σg = 109 m populations. However, the mean neighborhood size was higher in the core (Nb = 82 than in the peripheral (Nb = 48 populations. Conclusion Eastern white cedar populations have significant fine-scale genetic structure at short

  15. Climate driven changes in Engelmann spruce stands at timberline in the La Sal Mountains

    Science.gov (United States)

    James F. Fowler; Steven Overby; Barb Smith

    2012-01-01

    Due to global warming spruce-fir forest and associated vegetation may experience elevational displacement and altered species composition at the timberline-treeline ecotone. These forests and their component species are predicted to migrate upslope and thus landscape features such as timberline and treeline may move upslope as well. Prior to this study, baseline data...

  16. Enhanced thermal stability of the thylakoid membranes from spruce. A comparison with selected angiosperms

    Czech Academy of Sciences Publication Activity Database

    Karlický, Václav; Kurasová, Irena; Ptáčková, B.; Večeřová, Kristýna; Urban, Otmar; Špunda, Vladimír

    2016-01-01

    Roč. 130, 1-3 (2016), s. 357-371 ISSN 0166-8595 R&D Projects: GA MŠk(CZ) LO1415; GA ČR GA13-28093S Institutional support: RVO:67179843 Keywords : Norway spruce * Thermal stability * Circular dichroism * Photosystem II organization * Thylakoid membrane Subject RIV: ED - Physiology Impact factor: 3.864, year: 2016

  17. The epigenetic memory of temperature during embryogenesis modifies the expression of bud burst-related genes in Norway spruce epitypes.

    Science.gov (United States)

    Carneros, Elena; Yakovlev, Igor; Viejo, Marcos; Olsen, Jorunn E; Fossdal, Carl Gunnar

    2017-09-01

    Epigenetic memory affects the timing of bud burst phenology and the expression of bud burst-related genes in genetically identical Norway spruce epitypes in a manner usually associated with ecotypes. In Norway spruce, a temperature-dependent epigenetic memory established during embryogenesis affects the timing of bud burst and bud set in a reproducible and predictable manner. We hypothesize that the clinal variation in these phenological traits, which is associated with adaptation to growth under frost-free conditions, has an epigenetic component. In Norway spruce, dehydrins (DHNs) have been associated with extreme frost tolerance. DHN transcript levels decrease gradually prior to flushing, a time when trees are highly sensitive to frost. Furthermore, EARLY BUD BREAK 1 genes (EBB1) and the FT-TFL1-LIKE 2-gene (PaFTL2) were previously suggested to be implied in control of bud phenology. Here we report an analysis of transcript levels of 12 DHNs, 3 EBB1 genes and FTL2 in epitypes of the same genotype generated at different epitype-inducing temperatures, before and during spring bud burst. Earlier flushing of epitypes originating from embryos developed at 18 °C as compared to 28 °C, was associated with differential expression of these genes between epitypes and between buds and last year's needles. The majority of these genes showed significantly different expressions between epitypes in at least one time point. The general trend in DHN expression pattern in buds showed the expected reduction in transcript levels when approaching flushing, whereas, surprisingly, transcript levels peaked later in needles, mainly at the moment of bud burst. Collectively, our results demonstrate that the epigenetic memory of temperature during embryogenesis affects bud burst phenology and expression of the bud burst-related DHN, EBB1 and FTL2 genes in genetically identical Norway spruce epitypes.

  18. Feedbacks of windthrow for Norway spruce and Scots pine stands under changing climate

    International Nuclear Information System (INIS)

    Panferov, O; Rauch, E; Doering, C; Ahrends, B; Sogachev, A

    2009-01-01

    Wind damage is one of the major natural disturbances that can occur worldwide in most types of forests. Enhanced management using adequate decision support systems (DSS) can considerably reduce the risk of windthrow. The decision support system 'Forest and Climate Change' (DSS-WuK) which is currently being developed at Goettingen University aims at providing a tool for the quantitative assessment of biotic and abiotic risks for forest ecosystems under the conditions of changing climate. In order to assess the future risks of wind damage the system employs a coupled modelling approach combining the turbulence model SCAlar DIStribution (SCADIS) with the soil-vegetation-atmosphere-transfer (SVAT) model BROOK 90. The present study investigates projections of wind damage in Solling, Germany under climate scenarios A1B and B1, taking into account the windthrow feedbacks-changes of microclimate as a result of tree fall and consequent stabilization or destabilization of a forest stand. The results of the study indicate that in Solling the risk of windthrow for spruce and pine forest stands is likely to increase considerably during the 21st century. The general tendencies indicate that under A1B the probability of damage would be higher than under B1 and that under the same climate and soil conditions the risk for spruce stands would be higher than for pine stands of equal age. The degree of damage and feedback contribution as well as a sign of feedback in each particular case will strongly depend on the particular local or regional combination of climatic and soil factors with tree species, age and structure. For Solling the positive feedback to local climatic forcing is found. The feedback contributes considerably (up to 6% under given conditions) to the projected forest damage and cannot be neglected. Therefore, the adequate projection of future damage probabilities can be performed only with a process-based coupled soil-atmosphere model with corresponding high spatial

  19. Determination of stress bioindicators in three Norway spruce stands in northern Germany

    International Nuclear Information System (INIS)

    Godbold, D.L.; Feig, R.; Cremer-Herms, A.; Huettermann, A.

    1993-01-01

    At three sites in northern Germany (Witzenhausen, Egge-Mountains and Hamburg-Eissendorf) biochemical stress bioindicators (chlorophyll, starch, proline and acid phosphatase and peroxidase activity) were determined in the needles of healthy and damaged Norway spruce (Picea abies L. Karst.). Peroxidase activity was higher in 1 and 2 yr old needles of damaged Norway spruce at the Witzenhausen and Hamburg-Eissendorf sites. Phosphatase activity was significantly higher in needles of damaged trees in all needle ages at the Egge-mountains site in 1983, and at the Witzenhausen site in current and 1 yr old needles. At the Hamburg-Eissendorf site lower phosphatase activity in needless of damaged trees was determined. At the Witzenhausen site lower levels of chlorophyll were determined in the 1 and 2 yr old needles of damaged trees, whereas at the Hamburg-Eissendorf and Egge-mountains sites lower chlorophyll levels were found in current needles. Thus between healthy and damaged trees at a specific site differences in the stress bioindicators could be found, however no common pattern between the sites could be determined. The study indicates that these biochemical bioindicators may be used to show a general stress, but it is difficult to relate them to a specific stress factor. 21 refs., 1 fig., 2 tabs

  20. Natural regeneration ecology of a secondary altimontane spruce forests at Jelendol

    OpenAIRE

    Rozman, Elizabeta; Diaci, Jurij

    2008-01-01

    Natural regeneration of altimontane spruce forests at Jelendol is retarded dueto many factors. In autumn 2003, gaps of different size and parts of the surrounding stand were covered with a 5 x 5 grid m to define sampling plots. Atotal of 227 plots with 1,5 x 1,5 m in size were installed to analyse generalregeneration conditions and inhibitors. The following ecological parameters were estimated on each plot: micro relief, inclination, soil depth,ground cover, direct and diffuse solar radiation...

  1. Structural protein relationships among eastern equine encephalitis viruses.

    Science.gov (United States)

    Strizki, J M; Repik, P M

    1994-11-01

    We have re-evaluated the relationships among the polypeptides of eastern equine encephalitis (EEE) viruses using SDS-PAGE and peptide mapping of individual virion proteins. Four to five distinct polypeptide bands were detected upon SDS-PAGE analysis of viruses: the E1, E2 and C proteins normally associated with alphavirus virions, as well as an additional more rapidly-migrating E2-associated protein and a high M(r) (HMW) protein. In contrast with previous findings by others, the electrophoretic profiles of the virion proteins of EEE viruses displayed a marked correlation with serotype. The protein profiles of the 33 North American (NA)-serotype viruses examined were remarkably homogeneous, with variation detected only in the E1 protein of two isolates. In contrast, considerable heterogeneity was observed in the migration profiles of both the E1 and E2 glycoproteins of the 13 South American (SA)-type viruses examined. Peptide mapping of individual virion proteins using limited proteolysis with Staphylococcus aureus V8 protease confirmed that, in addition to the homogeneity evident among NA-type viruses and relative heterogeneity among SA-type viruses, the E1 and E2 proteins of NA- and SA-serotype viruses exhibited serotype-specific structural variation. The C protein was highly conserved among isolates of both virus serotypes. Endoglycosidase analyses of intact virions did not reveal substantial glycosylation differences between the glycoproteins of NA- and SA-serotype viruses. Both the HMW protein and the E2 protein (doublet) of EEE virus appeared to contain, at least in part, high-mannose type N-linked oligosaccharides. No evidence of O-linked glycans was found on either the E1 or the E2 glycoprotein. Despite the observed structural differences between proteins of NA- and SA-type viruses, Western blot analyses utilizing polyclonal antibodies indicated that immunoreactive epitopes appeared to be conserved.

  2. Climatic control of stand thinning in unmanaged spruce forests of the southern taiga in European Russia

    International Nuclear Information System (INIS)

    Vygodskaya, N.N.; Schulze, E.D.; Tchebakova, N.M.

    2002-01-01

    The demography of Picea abies trees was studied over a period of about 30 yr on permanent plots in six forest types of an unmanaged forest located in a forest reserve of the Southern Taiga, NW of Moscow. This study encompassed a broad range of conditions that are typical for old growth spruce forests in the boreal region, including sites with a high water table and well drained sites, podzolic soils, acidic soils and organic soils. At all sites stand density, tree height, breast height diameter and age has been periodically recorded since 1968. Tree density ranged between 178 and 1035 trees/ha for spruce and between 232 and 1168 trees/ha for the whole stand, including mainly Betula and Populus. Biomass ranged between 5.4 and 170 tdw/ha for spruce and between 33 to 198 tdw/ha for the whole stand. Averaged over a long period of time, biomass did not change with stand density according to the self-thinning rule. In fact, on most sites biomass remained almost constant in the long term, while stand density decreased. The study demonstrates that the loss of living trees was not regulated by competitive interactions between trees, but by disturbances caused by climatic events. Dry years caused losses of minor and younger trees without affecting biomass. In contrast, periodic storms resulted in a loss of biomass without affecting density, except for extreme events, where the whole stand may fall. Dry years followed by wet years enhance the effect on stand density. Since mainly younger trees were lost, the apparent average age of the stand increased more than real time (20% for Picea). Average mortality was 2.8 ± 0.5% yr 1 for spruce. Thus, the forest is turned over once every 160-180 yr by disturbances. The demography of dead trees shows that the rate of decay depends on the way the tree died. Storm causes uprooting and stem breakage, where living trees fall to the forest floor and decay with a mean residence time (t1/2) of about 16 yr (decomposition rate constant k d = 0

  3. Seasonal Changes of Coefficient Q10 in CO2 Flux from Soil Under Spruce Stand

    Czech Academy of Sciences Publication Activity Database

    Pavelka, Marian; Janouš, Dalibor

    2002-01-01

    Roč. 15, č. 15 (2002), s. 43-48. ISBN 80-7157-297-7 R&D Projects: GA ČR GA526/00/0485 Grant - others:EVK2(XE) CT-1999-00032 Keywords : soil CO2 efflux * Norway spruce * Q10 * respiration * soil Subject RIV: EH - Ecology, Behaviour

  4. Accumulation of logging residue in first thinnings of Scots pine and Norway spruce. Impact of top bucking diameter of roundwood

    Energy Technology Data Exchange (ETDEWEB)

    Raeisaenen, T.; Nurmi, J. (Finnish Forest Research Inst., Kannus (Finland)), e-mail: tommi.raisanen@metla.fi

    2010-07-01

    The aim of this study was to examine the impacts of changes in the minimum top diameter of roundwood on the accumulation of logging residue. The aim was also to compare estimates of residue accumulation calculated by tree-specific biomass models with field measurements from thinnings. Felling experiments were performed in first thinnings of pine and spruce to evaluate the model calculations. In the felling, mean relative masses of the tree tops of spruce were nearly doubled with each increment of 2 cm in the top diameter. Respectively in pine, the mean relative tree top mass was increased by 50-60 % when the top diameter was increased by 2 cm. The mass of total residue (tree top and all delimbed branches) was similarly increased, but the differences were not as large. Compared to pine, a lesser variation in the crown mass of the spruce sample resulted in a more accurate model prediction of masses of tree tops and total residue. The results indicate that the residue accumulation from a small group of trees cannot be predicted very reliably, but when a larger tree population or area is considered, the model predictions are enhanced to a more practicable level. (orig.)

  5. Geologic structure of the eastern mare basins. [lunar basalts

    Science.gov (United States)

    Dehon, R. A.; Waskom, J. D.

    1976-01-01

    The thickness of mare basalts in the eastern maria are estimated and isopachs of the basalts are constructed. Sub-basalt basin floor topography is determined, and correlations of topographic variations of the surface with variations in basalt thickness or basin floor topography are investigated.

  6. Mechanical properties of timber from wind damaged Norway spruce

    DEFF Research Database (Denmark)

    Hoffmeyer, Preben

    2003-01-01

    A storm may subject a tree to such bending stresses that extensive compression damage develops in the lee side. The tree may survive the wind load or it may be thrown. However, the damage is inherent and it may be of a magnitude to influence the mechanical properties of boards sawn from the stem....... The paper reports on a investigation of the relation between degree of damage and mechanical proper-ties of sawn timber from wind damaged Norway spruce. The project included about 250 bolts from wind damaged trees. The majority of bolts were cut to deliver a full-diameter plank containing the pith...

  7. Interspecific Plant Interactions Reflected in Soil Bacterial Community Structure and Nitrogen Cycling in Primary Succession.

    Science.gov (United States)

    Knelman, Joseph E; Graham, Emily B; Prevéy, Janet S; Robeson, Michael S; Kelly, Patrick; Hood, Eran; Schmidt, Steve K

    2018-01-01

    Past research demonstrating the importance plant-microbe interactions as drivers of ecosystem succession has focused on how plants condition soil microbial communities, impacting subsequent plant performance and plant community assembly. These studies, however, largely treat microbial communities as a black box. In this study, we sought to examine how emblematic shifts from early successional Alnus viridus ssp. sinuata (Sitka alder) to late successional Picea sitchensis (Sitka spruce) in primary succession may be reflected in specific belowground changes in bacterial community structure and nitrogen cycling related to the interaction of these two plants. We examined early successional alder-conditioned soils in a glacial forefield to delineate how alders alter the soil microbial community with increasing dominance. Further, we assessed the impact of late-successional spruce plants on these early successional alder-conditioned microbiomes and related nitrogen cycling through a leachate addition microcosm experiment. We show how increasingly abundant alder select for particular bacterial taxa. Additionally, we found that spruce leachate significantly alters the composition of these microbial communities in large part by driving declines in taxa that are enriched by alder, including bacterial symbionts. We found these effects to be spruce specific, beyond a general leachate effect. Our work also demonstrates a unique influence of spruce on ammonium availability. Such insights bolster theory relating the importance of plant-microbe interactions with late-successional plants and interspecific plant interactions more generally.

  8. Population structure and phylogeography reveal pathways of colonization by a migratory marine reptile (Chelonia mydas) in the central and eastern Pacific.

    Science.gov (United States)

    Dutton, Peter H; Jensen, Michael P; Frey, Amy; LaCasella, Erin; Balazs, George H; Zárate, Patricia; Chassin-Noria, Omar; Sarti-Martinez, Adriana Laura; Velez, Elizabeth

    2014-11-01

    Climate, behavior, ecology, and oceanography shape patterns of biodiversity in marine faunas in the absence of obvious geographic barriers. Marine turtles are an example of highly migratory creatures with deep evolutionary lineages and complex life histories that span both terrestrial and marine environments. Previous studies have focused on the deep isolation of evolutionary lineages (>3 mya) through vicariance; however, little attention has been given to the pathways of colonization of the eastern Pacific and the processes that have shaped diversity within the most recent evolutionary time. We sequenced 770 bp of the mtDNA control region to examine the stock structure and phylogeography of 545 green turtles from eight different rookeries in the central and eastern Pacific. We found significant differentiation between the geographically separated nesting populations and identified five distinct stocks (F ST = 0.08-0.44, P eastern Pacific Chelonia mydas form a monophyletic group containing 3 subclades, with Hawaii more closely related to the eastern Pacific than western Pacific populations. The split between sampled central/eastern and western Pacific haplotypes was estimated at around 0.34 mya, suggesting that the Pacific region west of Hawaii has been a more formidable barrier to gene flow in C. mydas than the East Pacific Barrier. Our results suggest that the eastern Pacific was colonized from the western Pacific via the Central North Pacific and that the Revillagigedos Islands provided a stepping-stone for radiation of green turtles from the Hawaiian Archipelago to the eastern Pacific. Our results fit with a broader paradigm that has been described for marine biodiversity, where oceanic islands, such as Hawaii and Revillagigedo, rather than being peripheral evolutionary "graveyards", serve as sources and recipients of diversity and provide a mechanism for further radiation.

  9. Utilization of hyperspectral image optical indices to assess the Norway spruce forest health status

    Czech Academy of Sciences Publication Activity Database

    Mišurec, J.; Kopáčková, V.; Lhotáková, Z.; Hanuš, Jan; Weyermann, J.; Entcheva-Campbel, P.; Albrechtová, J.

    2012-01-01

    Roč. 6, JUN 2012 (2012), 63545-1-63545-25 ISSN 1931-3195 R&D Projects: GA ČR GA205/09/1989 Institutional research plan: CEZ:AV0Z60870520 Keywords : chlorophyll * optical indices * Norway spruce * continuum removal * HyMap * actual physiological status * Sokolov basin * forest management Subject RIV: CE - Biochemistry Impact factor: 0.876, year: 2012

  10. Physiological implications of seasonal variation in membrane-associated calcium in red spruce mesophyll cells

    Science.gov (United States)

    D.H. DeHayes; P.G. Schaberg; G.J. Hawley; C.H. Borer; J.R. Cumming; J.R. Strimbeck

    1997-01-01

    We examined the pattern of seasonal variation in total foliar calcium (Ca) pools and plasma membrane-associated Ca (mCa) in mesophyll cells of current-year and 1-year-old needles of red spruce (Picea rubens Sarg.) and the relationship between mCa and total foliar Ca on an individual plant and seasonal basis. Foliar samples were collected from...

  11. Determination and analysis of uptake of gaseous hydrogen peroxide by red spruce seedlings, determined by CSTR-type chamber experiments

    International Nuclear Information System (INIS)

    Claiborn, C.S.; Aneja, V.P.; Carbonell, R.G.

    1991-01-01

    In order to better understand the pathways for damage, the fate of gaseous hydrogen peroxide in red spruce needles was examined. The uptake of gaseous hydrogen peroxide by red spruce trees was determined from controlled exposure chamber experiments in which the chamber behaved as a Continuous Stirred Tank Reactor (CSTR). The results from these experiments were analyzed using a detailed transport model developed from fundamental principles, in order to determine the fate of hydrogen peroxide in the needles and characterize the exposure. The chamber was specially designed to accommodate highly reactive gases. All inner surfaces were Teflon-coated to minimize wall losses. Fluxes of hydrogen peroxide, carbon dioxide, and water vapor were determined. Both daytime and nighttime conditions were examined. Although other investigators have reported that the flux of other, less water-soluble pollutants to red spruce decreases at night when the stomata closes, the hydrogen peroxide flux did not exhibit this behavior. The results of these studies suggest that, at the concentrations observed in the atmosphere, hydrogen peroxide does not reach the inner, mesophyll tissues, but is lost in water condensing in the cuticular wax residing in the stomatal antechamber, above the stomata. The implications of the condensation in the stomatal antechamber and subsequent reactions occurring in this water for forest damage are discussed

  12. Physiological responses of Norway spruce trees to elevated CO2 and SO2

    NARCIS (Netherlands)

    Tausz, M.; De Kok, L.J.; Stulen, I.

    Young Norway spruce (Picea abies (L.) Karst.) trees were exposed to elevated CO2 (0.8 mL L(-1)), SO2 (0.06 mu L L(-1)), and elevated CO2 and SO2 (0.8 mL L(-1) and 0.06 mu L L(-1), respectively) for three months. Exposure to elevated CO2 resulted in an increased biomass production of the needles,

  13. Height-Diameter Models for Mixed-Species Forests Consisting of Spruce, Fir, and Beech

    Directory of Open Access Journals (Sweden)

    Petráš Rudolf

    2014-06-01

    Full Text Available Height-diameter models define the general relationship between the tree height and diameter at each growth stage of the forest stand. This paper presents generalized height-diameter models for mixed-species forest stands consisting of Norway spruce (Picea abies Karst., Silver fir (Abies alba L., and European beech (Fagus sylvatica L. from Slovakia. The models were derived using two growth functions from the exponential family: the two-parameter Michailoff and three-parameter Korf functions. Generalized height-diameter functions must normally be constrained to pass through the mean stand diameter and height, and then the final growth model has only one or two parameters to be estimated. These “free” parameters are then expressed over the quadratic mean diameter, height and stand age and the final mathematical form of the model is obtained. The study material included 50 long-term experimental plots located in the Western Carpathians. The plots were established 40-50 years ago and have been repeatedly measured at 5 to 10-year intervals. The dataset includes 7,950 height measurements of spruce, 21,661 of fir and 5,794 of beech. As many as 9 regression models were derived for each species. Although the “goodness of fit” of all models showed that they were generally well suited for the data, the best results were obtained for silver fir. The coefficient of determination ranged from 0.946 to 0.948, RMSE (m was in the interval 1.94-1.97 and the bias (m was -0.031 to 0.063. Although slightly imprecise parameter estimation was established for spruce, the estimations of the regression parameters obtained for beech were quite less precise. The coefficient of determination for beech was 0.854-0.860, RMSE (m 2.67-2.72, and the bias (m ranged from -0.144 to -0.056. The majority of models using Korf’s formula produced slightly better estimations than Michailoff’s, and it proved immaterial which estimated parameter was fixed and which parameters

  14. Impacts of non-native Norway spruce plantation on abundance and species richness of ground beetles (Coleoptera: Carabidae

    Directory of Open Access Journals (Sweden)

    Z. Elek

    2001-06-01

    Full Text Available The impacts of non-native Norway spruce plantation on the abundance and species richness of carabids were studied in the Bükk National Park in Hungary, central Europe. Pitfall catches from recently established (5 yr old, young (15 yr after planting, middle-aged (30 yr after planting, old Norway spruce Picea abies plantation (50 yr after planting, and a native submontane beech forest (Fagetum sylvaticae as a control stand were compared.

    Our results showed that deciduous forest species decreased significantly in abundance in the plantations, and appeared in high abundance only in the native beech forest. Furthermore, open habitat species increased remarkably in abundance in the recently established plantation. Carabids were significantly more abundant and species rich in the native forest than in the plantations, while differences were not significant among the plantations. Multiple regression between the abundance and species richness of carabids and twelve environmental measurements showed that pH of the soil, herb cover and density of the carabids’ prey had a significant effect in determining abundance and species richness.

    Our results showed that plantation of non-native Norway spruce species had a detrimental effect on the composition of carabid communities and no regeneration could be observed during the growth of plantations even 50 yr after the establishment. This emphasises the importance of an active nature management practice to facilitate the recolonization of the native species.

  15. Left Dislocation in North-Eastern Neo-Aramaic dialects | Khan ...

    African Journals Online (AJOL)

    The North-Eastern Neo-Aramaic (NENA) dialects, which are the focus of this paper, were spoken across a wide area encompassing northern Iraq, north-west Iran, south-eastern Turkey, Armenia and Georgia. In these spoken dialects a distinction should be made between two major types of Left Dislocation (LD) structures.

  16. Use of gadolinium chloride as a contrast agent for imaging spruce knots by magnetic resonance

    Science.gov (United States)

    Thomas L. Eberhardt; Chi-Leung So; Amy H. Herlihy; Po-Wah So

    2006-01-01

    Treatments of knot-containing spruce wood blocks with a paramagnetic salt, gadolinium (III) chloride, in combination with solvent pretreatments, were evaluated as strategies to enhance the visualization of wood features by magnetic resonance imaging (MRI). Initial experiments with clear wood and excised knot samples showed differences in moisture uptake after...

  17. Relation between uranium mineralization and structural features, Gebel Gattar, north eastern desert, Egypt

    International Nuclear Information System (INIS)

    Salman, A.B.; Shalaby, M.H.; Abuzaid, M.M.; Ragab, A.

    1998-01-01

    Gebel Gattar area is situated in the northern Eastern Desert of Egypt, SW Hurghada city and is considered as an area of high potentialities for uranium deposits. The area is covered by Hammamat sediments and Gattarian granites. The Hammamat sediments are dissected by different types of dykes, while Gebel Gattar granites are cut only by basic dykes. These granites are mentioned as the younger pink granites, perthitic leucogranites, calc-alkaline and within plate granites. The structural deformations of the study area are represented by primary structures and secondary ones. The most prevailing structures are folding, faulting and jointing. The faults, especially those trending in the NNE-SSW and N-S directions played as pass ways to the ascending uranium-bearing hydrothermal solutions carrying uranium mineralizations. Most of them are located within a large pull apart basin. It is found from the relation between structures and uranium mineralization within the highly pro missing shear zones that uranium mineralizations are located within a large pull-apart basin, having about 2 km length and 0.5 km width. This idea is based up on the distribution of uranium mineralized lenses as shown in a block diagram. This conclusion is based on the structural framework of the area, the shape of mineralization and its distribution and their mutual relationships of Gl, Gll and GVl shear zones

  18. Tolerance of Norway spruce (Picea abies [L.] Karst.) embryogenic tissue to penicillin, carbapenem and aminoglycoside antibiotics

    Czech Academy of Sciences Publication Activity Database

    Malá, J.; Pavingerová, Daniela; Cvrčková, H.; Bříza, Jindřich; Dostál, J.; Šíma, P.

    2009-01-01

    Roč. 55, č. 4 (2009), s. 156-161 ISSN 1212-4834 R&D Projects: GA MZe QH71290 Institutional research plan: CEZ:AV0Z50510513 Keywords : somatic embryogenesis * Norway spruce * penicillin antibiotics * Agrobacterium tumefaciens * carbapenem antibiotics Subject RIV: EB - Genetics ; Molecular Biology

  19. BASIC CONCEPTS AND METHODS OF RESTORATION OF NATURAL FORESTS IN EASTERN EUROPE

    Directory of Open Access Journals (Sweden)

    V. N. Korotkov

    2017-03-01

    Full Text Available The modern forest in coniferous-broadleaf (hemiboreal and broadleaf zones of Eastern Europe were formed as a result of long-term human impact. This led to the loss of natural forests and total dominance of secondary forests combined with monocultures of spruce and pine that were created in clearings, burned areas and fallow lands. The reforestation model that was common in the late XIX and first half of the XX century and that was focused on the establishment of monocultures commercially valuable coniferous tree species (spruce and pine over large areas has resulted in declining biological diversity, increasing risk of tree damage due to outbreaks of pathogens and phytophagous insects, decreasing soil fertility, worsening soil and water conservation functions of forests. When restoring the prototypes of natural forests it is necessary to be guided by the modern concepts of synecology and model reconstructions of forest cover in pre-anthropogenic period that are briefly discussed in the paper. Based on the analysis of literature and research experience the author proposes the concept of natural forest restoration that can be applied primarily to the coniferous-broadleaf and broadleaf forests. The main goal is to create multiple-aged and polydominant near-natural forest ecosystems with higher resistance to fungal diseases and outbreaks of phytophagous insects. The field of concept application is specially protected natural areas (national parks, natural parks, wildlife sanctuaries, etc., different categories of protective forests located within the zones of coniferous-broadleaf and broadleaf forests on the East European Plain. The formation of multiple-aged forests is possible when group felling and group-clear felling that largely imitate the natural gap-mosaic stand are implemented. The formation of new generations of trees is possible both due to the natural regeneration and the development of forest cultures. The article provides the full set

  20. Imaging of Norway spruce early somatic embryos with the ESEM, Cryo-SEM and laser scanning microscope.

    Science.gov (United States)

    Neděla, Vilém; Hřib, Jiří; Havel, Ladislav; Hudec, Jiří; Runštuk, Jiří

    2016-05-01

    This article describes the surface structure of Norway spruce early somatic embryos (ESEs) as a typical culture with asynchronous development. The microstructure of extracellular matrix covering ESEs were observed using the environmental scanning electron microscope as a primary tool and using the scanning electron microscope with cryo attachment and laser electron microscope as a complementary tool allowing our results to be proven independently. The fresh samples were observed in conditions of the air environment of the environmental scanning electron microscope (ESEM) with the pressure from 550Pa to 690Pa and the low temperature of the sample from -18°C to -22°C. The samples were studied using two different types of detector to allow studying either the thin surface structure or material composition. The scanning electron microscope with cryo attachment was used for imaging frozen extracellular matrix microstructure with higher resolution. The combination of both electron microscopy methods was suitable for observation of "native" plant samples, allowing correct evaluation of our results, free of error and artifacts. Copyright © 2016 Elsevier Ltd. All rights reserved.

  1. The Genetic Structure of Wild Orobanche cumana Wallr. (Orobanchaceae Populations in Eastern Bulgaria Reflects Introgressions from Weedy Populations

    Directory of Open Access Journals (Sweden)

    Rocío Pineda-Martos

    2014-01-01

    Full Text Available Orobanche cumana is a holoparasitic plant naturally distributed from central Asia to south-eastern Europe, where it parasitizes wild Asteraceae species. It is also an important parasitic weed of sunflower crops. The objective of this research was to investigate genetic diversity, population structure, and virulence on sunflower of O. cumana populations parasitizing wild plants in eastern Bulgaria. Fresh tissue of eight O. cumana populations and mature seeds of four of them were collected in situ on wild hosts. Genetic diversity and population structure were studied with SSR markers and compared to weedy populations. Two main gene pools were identified in Bulgarian populations, with most of the populations having intermediate characteristics. Cross-inoculation experiments revealed that O. cumana populations collected on wild species possessed similar ability to parasitize sunflower to those collected on sunflower. The results were explained on the basis of an effective genetic exchange between populations parasitizing sunflower crops and those parasitizing wild species. The occurrence of bidirectional gene flow may have an impact on wild populations, as new physiological races continuously emerge in weedy populations. Also, genetic variability of wild populations may favour the ability of weedy populations to overcome sunflower resistance mechanisms.

  2. The Genetic Structure of Wild Orobanche cumana Wallr. (Orobanchaceae) Populations in Eastern Bulgaria Reflects Introgressions from Weedy Populations

    Science.gov (United States)

    Pineda-Martos, Rocío; Pujadas-Salvà, Antonio J.; Fernández-Martínez, José M.; Stoyanov, Kiril; Pérez-Vich, Begoña

    2014-01-01

    Orobanche cumana is a holoparasitic plant naturally distributed from central Asia to south-eastern Europe, where it parasitizes wild Asteraceae species. It is also an important parasitic weed of sunflower crops. The objective of this research was to investigate genetic diversity, population structure, and virulence on sunflower of O. cumana populations parasitizing wild plants in eastern Bulgaria. Fresh tissue of eight O. cumana populations and mature seeds of four of them were collected in situ on wild hosts. Genetic diversity and population structure were studied with SSR markers and compared to weedy populations. Two main gene pools were identified in Bulgarian populations, with most of the populations having intermediate characteristics. Cross-inoculation experiments revealed that O. cumana populations collected on wild species possessed similar ability to parasitize sunflower to those collected on sunflower. The results were explained on the basis of an effective genetic exchange between populations parasitizing sunflower crops and those parasitizing wild species. The occurrence of bidirectional gene flow may have an impact on wild populations, as new physiological races continuously emerge in weedy populations. Also, genetic variability of wild populations may favour the ability of weedy populations to overcome sunflower resistance mechanisms. PMID:25143963

  3. Physical structure and algae community of summer upwelling off eastern Hainan

    Science.gov (United States)

    Xu, H.; Liu, S.; Xie, Q.; Hong, B.; Long, T.

    2017-12-01

    The upwelling system is the most productive ecosystem along the continental shelf of the northern South China Sea Shelf. It brings nutrient from bottom to surface and blooms biotic community driven by summer monsoon. In this study, we present observed results of physical and biotic community structures during August, 2015 in the upwelling system along Hainan eastern coast, which is one the strongest upwelling systems in the northern South China Sea. By using hydrological data collected by CTD, we found a significant cold water tongue with high salinity which extended from offshore to 100 m isobaths. However, dissolved oxygen (DO) showed a sandwich structure in which high core of DO concentration appeared at the layer from 5 m to 30 m. It possibly was caused by the advection transport of high DO from adjacent area. Basically, this upwelling system was constrained at northern area of 18.8ºN in horizontal due to the weakening summer monsoon in August. In addition, we collected water sample at the upwelling area and measured algae categories and concentration by high performance liquid chromatography (HPLC). Results show the biotic community was dominated by five types of algae mainly, they were diatoms, dinoflagellates, green algae, prokaryotes and prochlorococcus. And different patterns of different algae were demonstrated. In the upwelling area, diatoms and prokaryotes show opposite structures, and more complex pattern for the rest three algae indicating an active biotic community in the upwelling system.

  4. Two New Epiphyllous Leptolejeunea (Hepaticae: Lejeuneaceae from Eastern Himalaya, India

    Directory of Open Access Journals (Sweden)

    Monalisa Dey

    2010-11-01

    Full Text Available Two new epiphyllous species of the genus Leptolejeunea (Spruce Schiffn., viz. L. mirikana M. Dey et D. K. Singh sp. nov. and L. udarii M. Dey et D. K. Singh sp. nov., are described from Darjeeling district of West Bengal and East district of Sikkim respectively in Eastern Himalaya, India. While, the former is characterized by obovate leaf lobes with acute – subacute leaf apices and entire margin; rectangular – trapezoid underleaves with rectangular – lanceolate lobes which are 3-4 cells long, 2-3 cells wide, or rarely 1 cell wide at apex, and crenulate – denticulate margin; presence of ocelli in female bracts and bracteoles and obconical perianth with truncate apices and 5 obliquely – horizontally spreading horn-like keels extending from apex to 1/2-3/5 of perianth length, the latter is distinct in having small length-breadth ratio of the leaves ranging between 1.2:1-1.3:1; obovate leaf lobes with subacute apices; terminal as well as intercalary androecia with male bracteoles present throughout; numerous ocelli scattered on female bracts, bracteoles and perianth and obconical – campanulate perianth with 5 smooth, erect – obliquely spreading horn-like keels. A key to the Indian species of the genus has been provided.

  5. Soil Warming: Consequences for Foliar Litter Decay in a Spruce-Fir Forest in Maine, USA

    Science.gov (United States)

    Lindsey E. Rustad; Ivan J. Fernandez

    1998-01-01

    Increased rates of litter decay due to projected global warming could substantially alter the balance between C assimilation and release in forest soils, with consequent feedbacks to climate change. This study was conducted to investigate the effects of soil warming on the decomposition of red spruce (Picea rubens Sarg.) and red maple (...

  6. Influence of sulfur dioxide on the mineral composition of needles from spruces

    Energy Technology Data Exchange (ETDEWEB)

    Materna, J

    1961-01-01

    Until recently all the authors knew about changes in the mineral composition of plants exposed to air pollution was that the sulfur content increases considerably. The question arises whether other mineral substances, too, accumulate in the assimilating organs of smoke injured plants, particularly cations such as calcium, potassium and magnesium. Results of analyses of spruce needles from an air polluted forest in the Erzebirge in Czechoslovakia yielded no relationship between the accumulation of sulfates and the mentioned cations.

  7. Measurement methods and variability assessment of the Norway spruce total leaf area: Implications for remote sensing

    NARCIS (Netherlands)

    Homolova, L.; Lukes, P.; Malenovsky, Z.; Lhotakova, Z.; Kaplan, V.; Hanus, J.

    2013-01-01

    Estimation of total leaf area (LAT) is important to express biochemical properties in plant ecology and remote sensing studies. A measurement of LAT is easy in broadleaf species, but it remains challenging in coniferous canopies. We proposed a new geometrical model to estimate Norway spruce LAT and

  8. The incidence of dwarf mistletoe in Minnesota black spruce stands detected by operational inventories

    Science.gov (United States)

    Fred Baker; Mark Hansen; John D. Shaw; Manfred Mielke; Dixon Shelstad

    2012-01-01

    We surveyed black spruce stands within 0.5 miles of US Forest Service Forest Inventory and Analysis (FIA) plots and compared dwarf mistletoe status with that of the FIA and Minnesota Department of Natural Resources (DNR) forest inventories. Our results differed from FIA results in 3 of 16 stands with FIA plots, with FIA most often not recording dwarf mistletoe in...

  9. Balsam fir conservation and red spruce ecosystem restoration initiatives in the West Virginia highlands

    Science.gov (United States)

    Corey A. Bonasso; David W. Saville

    2010-01-01

    The West Virginia Highlands Conservancy has been working for more than a decade to protect, conserve, and restore the spruce-fir forests in West Virginia. Beginning in the mid 1990s an effort was initiated to conserve balsam fir in West Virginia where it reaches its southern most extent in North America. This work led to further efforts which have focused on the...

  10. Revised Site Index Curves for Balsam Fir and White Spruce in the Lake States

    Science.gov (United States)

    Willard H. Carmean; Jerold T. Hahn

    1981-01-01

    The original site index curves for balsam fir and white spruce are revised from a breast height age to a total age basis. Site index values from these revised curves are thus comparable to index values for other species that are based upon total tree age. This note also includes formulations for estimating site index by using computers or programmable, hand-...

  11. Stratigraphy and structure of eastern Syria across the Euphrates depression

    Science.gov (United States)

    Sawaf, Tarif; Al-Saad, Damen; Gebran, Ali; Barazangi, Muawia; Best, John A.; Chaimov, Thomas A.

    1993-04-01

    A N-S crustal-scale geotransect across the northern Arabian platform in eastern Syria reveals an alternating series of basement uplifts and basins separated by predominantly transpressional fault zones above an effectively uniform crust. Four major tectonic provinces are crossed along a 325 × 100 km corridor that extends from the Iraqi border in the south to the Turkish border in the north: the Rutbah uplift, the Euphrates depression, the Abd el Aziz structural zone, and the Qamichli uplift. These features are the manifestations of reactivated pre-Cenozoic structures that responded to forces acting along nearby Arabian plate boundaries, particularly Cenozoic convergence and collision along the margins of the northern Arabian platform i.e., the Bitlis suture and the East Anatolian fault in southern Turkey and the Zagros suture in Iran and Iraq. The database for this study consists of 3000 km of industry seismic reflection data, 28 exploratory wells, and geologic and Bouguer gravity maps. The deep crustal structure and, in part, the basement geometry along this transect are inferred from two-dimensional modeling of Bouguer gravity, whereas the shallow (about 8 km) structure is constrained primarily by well and seismic data. Features of the geotransect reveal: (1) A relatively uniform crustal column approximately 37 km thick with only minor crustal thinning beneath the Euphrates. Crustal thinning may be slightly more pronounced beneath the Euphrates (about 35 km) to the southeast of the transect where the Bouguer gravity anomaly is slightly higher. (2) Along the Euphrates depression, ongoing subsidence, which began during the Late Cretaceous, resulted in the deposition of at least 3 km of Late Cretaceous and Cenozoic rocks. The structural complexity of the Paleozoic and most of the Mesozoic sedimentary sections along the transect contrasts markedly with a relatively simple, flat-lying Cenozoic section along most of the transect. A notable exception is the Abd el Aziz

  12. Seismic response and fragility evaluation for an Eastern US NPP including soil-structure interaction effects

    International Nuclear Information System (INIS)

    Ghiocel, Dan M.; Wilson, Paul R.; Thomas, Gary G.; Stevenson, John D.

    1998-01-01

    The paper discusses methodological aspects involved in a probabilistic seismic soil-structure interaction (SSI) analysis for a Seismic Probabilistic Risk Assessment (SPRA) review. An example of an Eastern US nuclear power plant (NPP) is presented. The approach presented herein follows the current practice of the Individual Plant Examination for External Events (IPEEE) program in the US. The NPP is founded on a relatively soft soil deposit, and thus the SSI effects on seismic responses are significant. Probabilistic models used for the idealization of the seismic excitation and the surrounding soil deposit are described. Using a lognormal format, computed random variability effects were combined with those proposed in the SPRA methodology guidelines. Probabilistic floor response spectra and structural fragilities for different NPP buildings were computed. Structural capacities were determined following the current practice which assumes independent median safety factors for strength and inelastic absorption. Limitations of the IPEEE practice for performing SPRA are discussed and alternate procedures, more rigorous and simple to implement, are suggested

  13. Time since death and decay rate constants of Norway spruce and European larch deadwood in subalpine forests determined using dendrochronology and radiocarbon dating

    Science.gov (United States)

    Petrillo, M.; Cherubini, P.; Fravolini, G.; Ascher, J.; Schärer, M.; Synal, H.-A.; Bertoldi, D.; Camin, F.; Larcher, R.; Egli, M.

    2015-09-01

    Due to the large size and highly heterogeneous spatial distribution of deadwood, the time scales involved in the coarse woody debris (CWD) decay of Picea abies (L.) Karst. and Larix decidua Mill. in Alpine forests have been poorly investigated and are largely unknown. We investigated the CWD decay dynamics in an Alpine valley in Italy using the five-decay class system commonly employed for forest surveys, based on a macromorphological and visual assessment. For the decay classes 1 to 3, most of the dendrochronological samples were cross-dated to assess the time that had elapsed since tree death, but for decay classes 4 and 5 (poorly preserved tree rings) and some others not having enough tree rings, radiocarbon dating was used. In addition, density, cellulose and lignin data were measured for the dated CWD. The decay rate constants for spruce and larch were estimated on the basis of the density loss using a single negative exponential model. In the decay classes 1 to 3, the ages of the CWD were similar varying between 1 and 54 years for spruce and 3 and 40 years for larch with no significant differences between the classes; classes 1-3 are therefore not indicative for deadwood age. We found, however, distinct tree species-specific differences in decay classes 4 and 5, with larch CWD reaching an average age of 210 years in class 5 and spruce only 77 years. The mean CWD rate constants were 0.012 to 0.018 yr-1 for spruce and 0.005 to 0.012 yr-1 for larch. Cellulose and lignin time trends half-lives (using a multiple-exponential model) could be derived on the basis of the ages of the CWD. The half-lives for cellulose were 21 yr for spruce and 50 yr for larch. The half-life of lignin is considerably higher and may be more than 100 years in larch CWD.

  14. Analysis of neotectonic structures in the Eastern Precordillera of Argentina in relation to seismic hazard by the application of integrated geophysical methods

    Science.gov (United States)

    Correa-Otto, Sebastián; Ariza, Juan; Lince Klinger, Federico; Giménez, Mario; López Hidalgo, Andrés

    2018-03-01

    The city of San Juan, in the Central-Western region of Argentina, has been the target of very destructive superficial earthquakes, some of which have not been associated to a clear structural source up to this date. The city is constantly growing outside the valley where it is located, towards the area of Eastern Precordillera which is currently having an increased socio-cultural activity. Thus, this study is focused on increasing the geological knowledge of the latter by studying the eastern flank of Sierra Chica de Zonda (Eastern Precordillera) whose proved neotectonic activity represents a geohazard. On the basis of the general geological setting the neotectonic structures in the study area are related to a major active synclinal folding located just under the western sector of the San Juan city. Geophysical potential methods (gravimetric and magnetometric surveys) were used to recognize contacts by contrast of density and magnetic susceptibility. In order to reduce the ambiguity of these methods the gravi-magnetometric results were constrained by using seismic and electrical tomographies. These contacts where geophysical properties abruptly change, were interpreted as faults despite many of them not having a superficial expression. The latter being of great importance to asses the seismic hazard of the study area.

  15. Antioxidant defences of Norway spruce bark against bark beetles and its associated blue-stain fungus

    Directory of Open Access Journals (Sweden)

    Felicijan Mateja

    2015-12-01

    Full Text Available Bark beetles and their fungal associates are integral parts of forest ecosystems, the European spruce bark beetle (Ips typographus Linnaeus, 1758 and the associated pathogenic blue stain fungus Ceratocystis polonica (SIEM. C. MOREAU, are the most devastating pests regarding Norway spruce [Picea abies (L. H. KARST.]. Bark beetles commonly inhabit weakened and felled trees as well as vital trees. They cause physiological disorders in trees by destroying a phloem and cambium or interrupt the transpiration -ow in the xylem. Conifers have a wide range of effective defence mechanisms that are based on the inner bark anatomy and physiological state of the tree. The basic function of bark defences is to protect the nutrient-and energy-rich phloem, the vital meristematic region of the vascular cambium, and the transpiration -ow in the sapwood. The main area of defence mechanisms is secondary phloem, which is physically and chemically protected by polyphenolic parenchyma (PP cells, sclerenchyma, calcium oxalate crystals and resin ducts. Conifer trunk pest resistance includes constitutive, inducible defences and acquired resistance. Both constitutive and inducible defences may deter beetle invasion, impede fungal growth and close entrance wounds. During a successful attack, systemic acquired resistance (SAR becomes effective and represents a third defence strategy. It gradually develops throughout the plant and provides a systemic change within the whole tree’s metabolism, which is maintained over a longer period of time. The broad range of defence mechanisms that contribute to the activation and utilisation of SAR, includes antioxidants and antioxidant enzymes, which are generally linked to the actions of reactive oxygen species (ROS. The presented review discusses the current knowledge on the antioxidant defence strategies of spruce inner bark against the bark beetle (Ips typographus and associated blue stain fungus (Ceratocystis polonica.

  16. Taxonomic identity of a galling adelgid (Hemiptera: Adelgidae) from three spruce species in central Japan

    Science.gov (United States)

    Masakazu Sano; Nathan P. Havill; Kenichi. Ozaki

    2011-01-01

    Gall-forming insects are commonly highly host-specific, and galling species once thought to be oligo- or polyphagous are often found to represent a complex of host-specific races or cryptic species. A recent DNA barcoding study documented that an unidentified species of the genus Adelges is a gall-former associated with four spruce species (...

  17. Beech and spruce under the influence of electromagnetic radiation by radar

    International Nuclear Information System (INIS)

    Götz, G.; Matyssek, R.; Käs, G.

    2001-01-01

    Throughout a three-year study period beech and spruce trees were examined for potential effects of electromagnetic radiation by radar on the morphological and physiological performance at the crown level. No effects of radar on photosynthesis, transpiration, stomatal regulation as well as twig and foliage differentiation were found in late summer after seasonal exposure to this kind of radiation, when comparing radar-exposed with shielded crown parts. Adverse effects caused by radar on forest trees appear to be unlikely on a short-term scale, given conditions similar to those of this case study [de

  18. Analyzing tree cores to detect petroleum hydrocarbon-contaminated groundwater at a former landfill site in the community of Happy Valley-Goose Bay, eastern Canadian subarctic

    DEFF Research Database (Denmark)

    Fonkwe, Merline L D; Trapp, Stefan

    2016-01-01

    -gas chromatography-mass spectrometry. BTEX compounds were detected in tree cores, corroborating known groundwater contamination. A zone of anomalously high concentrations of total BTEX constituents was identified and recommended for monitoring by groundwater wells. Tree cores collected outside the landfill site......This research examines the feasibility of analyzing tree cores to detect benzene, toluene, ethylbenzene, and m, p, o-xylene (BTEX) compounds and methyl tertiary-butyl ether (MTBE) in groundwater in eastern Canada subarctic environments, using a former landfill site in the remote community of Happy...... Valley-Goose Bay, Labrador. Petroleum hydrocarbon contamination at the landfill site is the result of environmentally unsound pre-1990s disposal of households and industrial solid wastes. Tree cores were taken from trembling aspen, black spruce, and white birch and analyzed by headspace...

  19. Pathogenicity of Neonectria fuckeliana on Norway Spruce Clones in Sweden and Potential Management Strategies

    Directory of Open Access Journals (Sweden)

    Martin Pettersson

    2018-02-01

    Full Text Available The fungus Neonectria fuckeliana has become an increasing problem on Norway spruce (Picea abies in the Nordic countries during recent years. Canker wounds caused by the pathogen reduce timber quality and top-dieback is a problem for the Christmas tree industry. In this study, four inoculation trials were conducted to examine the ability of N. fuckeliana to cause disease on young Norway spruce plants and determine how different wound types would affect the occurrence and severity of the disease. Symptom development after 8–11 months was mainly mild and lesion lengths under bark were generally minor. However, N. fuckeliana could still be reisolated and/or molecularly detected. Slow disease development is in line with older studies describing N. fuckeliana as a weak pathogen. However, the results do not explain the serious increased damage by N. fuckeliana registered in Nordic forests and Christmas tree plantations. Potential management implications, such as shearing Christmas trees during periods of low inoculum pressure, cleaning secateurs between trees, and removal and burning of diseased branches and trees to avoid inoculum transfer and to keep disease pressure low, are based on experiments presented here and experiences with related pathogens.

  20. Alkaline intrusion in a granulite ensemble in the Eastern Ghats belt

    Indian Academy of Sciences (India)

    Alkaline magmatism; Eastern Ghats belt; pull-apart structure; plume tectonics. ... is one of several bodies in the high-grade Eastern Ghats belt, but this one is an ... Geological Studies Unit, Indian Statistical Institute, 203 B. T. Road, Kolkata 700 ...

  1. Seedling establishment and distribution of direct radiation in slit-shaped openings of Norway spruce forests in the intermediate Alps

    International Nuclear Information System (INIS)

    Brang, P.

    1996-01-01

    Direct radiation is crucial for Norway spruce (Picea abies (L.) Karst.) seedling establishment in high-montane and subalpine spruce forests. Fisheye photography was used to estimate the daily distribution of direct radiation in small forest openings on a north-northwest and a south facing slope near Sedrun (Grisons, Switzerland). In slit-shaped openings on the north-northwest facing slope long sunflecks mostly occurred in the afternoon, when the sun shines parallel to the slit axis. This is in accordance to the silvicultural intention. However, since the stands are clumpy and therefore pervious to sunlight, the daily sunfleck distribution is fairly even notwithstanding the slit orientation, and direct radiation at noon is the dominant form of incident energy. In small circular to rectangular openings on the south facing slope direct radiation peaks at noontide. A seeding trial imitating natural seedling establishment was set in place in openings on both slopes. Based on this trial, the relations among seedling establishment, aspect, slit shape, size, and orientation are discussed for Norway spruce forests in the intermediate Alps. The directional weather factors such as radiation and precipitation can be highly influenced by slits, which is why suitable microclimate for seedling establishment can be promoted provided the slits are oriented appropriately. Slits in which the most insolated edges are oriented windward are especially favourable

  2. Interspecific Plant Interactions Reflected in Soil Bacterial Community Structure and Nitrogen Cycling in Primary Succession

    Directory of Open Access Journals (Sweden)

    Joseph E. Knelman

    2018-02-01

    Full Text Available Past research demonstrating the importance plant–microbe interactions as drivers of ecosystem succession has focused on how plants condition soil microbial communities, impacting subsequent plant performance and plant community assembly. These studies, however, largely treat microbial communities as a black box. In this study, we sought to examine how emblematic shifts from early successional Alnus viridus ssp. sinuata (Sitka alder to late successional Picea sitchensis (Sitka spruce in primary succession may be reflected in specific belowground changes in bacterial community structure and nitrogen cycling related to the interaction of these two plants. We examined early successional alder-conditioned soils in a glacial forefield to delineate how alders alter the soil microbial community with increasing dominance. Further, we assessed the impact of late-successional spruce plants on these early successional alder-conditioned microbiomes and related nitrogen cycling through a leachate addition microcosm experiment. We show how increasingly abundant alder select for particular bacterial taxa. Additionally, we found that spruce leachate significantly alters the composition of these microbial communities in large part by driving declines in taxa that are enriched by alder, including bacterial symbionts. We found these effects to be spruce specific, beyond a general leachate effect. Our work also demonstrates a unique influence of spruce on ammonium availability. Such insights bolster theory relating the importance of plant–microbe interactions with late-successional plants and interspecific plant interactions more generally.

  3. Faulting in eastern New Mexico: Revision 1

    International Nuclear Information System (INIS)

    Murphy, P.J.

    1987-08-01

    This area in eastern New Mexico is being studied because of its proximity to a proposed high-level nuclear waste repository site in Deaf Smith County, Texas. Regional and local tectonics must be thoroughly understood in order to be able to predict future tectonic activities in the site region. Hydrogeologic studies indicate that ground-water recharge, for regional deep and shallow aquifer systems, occurs primarily within the study area. Regional, easterly ground-water flow may be significantly affected by the spacing, orientation, and character of faults identified in this report. The tectonic history of eastern New Mexico is developed from information from 660 exploratory wells. A history of recurrent tectonic movements is evident, beginning possibly in the late Precambrian and extending into the late Cenozoic. The nature of the evidence includes the lateral and vertical distribution and the lithology of these deposits. The results are presented mainly as isopach and structure contour maps and as structural cross sections. The Paleozoic tectonic history of eastern New Mexico is similar and related to the history of the Texas Panhandle. Differences occur primarily in the Mesozoic and Cenozoic histories; tectonic uplift persisted for longer periods and faults show evidence of repeated movements in eastern New Mexico. This is probably a result of the proximity of the area to the Laramide and Basin and Range deformational events. 442 refs., 35 figs

  4. Role of pre-existing structures in controlling the Cenozoic tectonic evolution of the eastern Tibetan plateau: New insights from analogue experiments

    Science.gov (United States)

    Sun, Ming; Yin, An; Yan, Danping; Ren, Hongyu; Mu, Hongxu; Zhu, Lutao; Qiu, Liang

    2018-06-01

    Pre-existing weakness due to repeated tectonic, metamorphic, and magmatic events is a fundamental feature of the continental lithosphere on Earth. Because of this, continental deformation results from a combined effect of boundary conditions imposed by plate tectonic processes and heterogeneous and anisotropic mechanical strength inherited from protracted continental evolution. In this study, we assess how this interaction may have controlled the Cenozoic evolution of the eastern Tibetan plateau during the India-Asia collision. Specifically, we use analogue models to evaluate how the pre-Cenozoic structures may have controlled the location, orientation, and kinematics of the northwest-striking Xianshuihe and northeast-striking Longmen Shan fault zones, the two most dominant Cenozoic structures in eastern Tibet. Our best model indicates that the correct location, trend, and kinematics of the two fault systems can only be generated and maintained if the following conditions are met: (1) the northern part of the Songpan-Ganzi terrane in eastern Tibet has a strong basement whereas its southern part has a weak basement, (2) the northern strong basement consists of two pieces bounded by a crustal-scale weak zone that is expressed by the Triassic development of a northwest-trending antiform exposing middle and lower crustal rocks, and (3) the region was under persistent northeast-southwest compression since ∼35 Ma. Our model makes correct prediction on the sequence of deformation in eastern Tibet; the Longmen Shan right-slip transpressional zone was initiated first as an instantaneous response to the northeast-southwest compression, which is followed by the formation of the Xianshuihe fault about a half way after the exertion of northeast-southwest shortening in the model. The success of our model highlights the importance of pre-existing weakness, a key factor that has been largely neglected in the current geodynamic models of continental deformation.

  5. Crustal structure of the Central-Eastern Greenland: results from the Topo Greenland refraction profile

    DEFF Research Database (Denmark)

    Shulgin, Alexey; Thybo, Hans

    2014-01-01

    Until present, seismic surveys have only been carried out offshore and near the coasts of Greenland, where the crustal structure is affected by oceanic break-up. We present the deep seismic structure of the crust of the interior of Greenland, based on the new and the only existing so far seismic...... refraction/wide-angle reflection profile. The seismic data was acquired by a team of six people during a two-month long experiment in summer of 2011 on the ice cap in the interior of central-eastern Greenland. The presence of an up to 3.4 km thick ice sheet, permanently covering most of the land mass, made...... acquisition of geophysical data logistically complicated. The profile extends 310 km inland in E-W direction from the approximate edge of the stable ice cap near the Scoresby Sund across the center of the ice cap. 350 Reftek Texan receivers recorded high-quality seismic data from 8 equidistant shots along...

  6. A spatially explicit hydro-ecological modeling framework (BEPS-TerrainLab V2.0): Model description and test in a boreal ecosystem in Eastern North America

    Science.gov (United States)

    Govind, Ajit; Chen, Jing Ming; Margolis, Hank; Ju, Weimin; Sonnentag, Oliver; Giasson, Marc-André

    2009-04-01

    SummaryA spatially explicit, process-based hydro-ecological model, BEPS-TerrainLab V2.0, was developed to improve the representation of ecophysiological, hydro-ecological and biogeochemical processes of boreal ecosystems in a tightly coupled manner. Several processes unique to boreal ecosystems were implemented including the sub-surface lateral water fluxes, stratification of vegetation into distinct layers for explicit ecophysiological representation, inclusion of novel spatial upscaling strategies and biogeochemical processes. To account for preferential water fluxes common in humid boreal ecosystems, a novel scheme was introduced based on laboratory analyses. Leaf-scale ecophysiological processes were upscaled to canopy-scale by explicitly considering leaf physiological conditions as affected by light and water stress. The modified model was tested with 2 years of continuous measurements taken at the Eastern Old Black Spruce Site of the Fluxnet-Canada Research Network located in a humid boreal watershed in eastern Canada. Comparison of the simulated and measured ET, water-table depth (WTD), volumetric soil water content (VSWC) and gross primary productivity (GPP) revealed that BEPS-TerrainLab V2.0 simulates hydro-ecological processes with reasonable accuracy. The model was able to explain 83% of the ET, 92% of the GPP variability and 72% of the WTD dynamics. The model suggests that in humid ecosystems such as eastern North American boreal watersheds, topographically driven sub-surface baseflow is the main mechanism of soil water partitioning which significantly affects the local-scale hydrological conditions.

  7. Is it possible and necessary to control European spruce bark beetle Ips typographus (L. outbreak in the Białowieża Forest?

    Directory of Open Access Journals (Sweden)

    Hilszczański Jacek

    2017-03-01

    Full Text Available In response to the information published in ‘Forest Research Papers’ (vol. 77(4, 2016, regarding the problem of the European spruce bark beetle Ips typographus (L. in the Białowieża Forest, we present our viewpoint on this issue. The role of the European spruce bark beetle in the Białowieża Forest is discussed based on the experience gained in Europe’s forests. We present the effects of I. typographus outbreaks on forest biodiversity as well as outbreak mitigation in the context of the processes taking place in semi-natural forests.

  8. Historical and projected carbon balance of mature black spruce ecosystems across north america: The role of carbon-nitrogen interactions

    Science.gov (United States)

    Clein, Joy S.; McGuire, A.D.; Zhang, X.; Kicklighter, D.W.; Melillo, J.M.; Wofsy, S.C.; Jarvis, P.G.; Massheder, J.M.

    2002-01-01

    The role of carbon (C) and nitrogen (N) interactions on sequestration of atmospheric CO2 in black spruce ecosystems across North America was evaluated with the Terrestrial Ecosystem Model (TEM) by applying parameterizations of the model in which C-N dynamics were either coupled or uncoupled. First, the performance of the parameterizations, which were developed for the dynamics of black spruce ecosystems at the Bonanza Creek Long-Term Ecological Research site in Alaska, were evaluated by simulating C dynamics at eddy correlation tower sites in the Boreal Ecosystem Atmosphere Study (BOREAS) for black spruce ecosystems in the northern study area (northern site) and the southern study area (southern site) with local climate data. We compared simulated monthly growing season (May to September) estimates of gross primary production (GPP), total ecosystem respiration (RESP), and net ecosystem production (NEP) from 1994 to 1997 to available field-based estimates at both sites. At the northern site, monthly growing season estimates of GPP and RESP for the coupled and uncoupled simulations were highly correlated with the field-based estimates (coupled: R2= 0.77, 0.88 for GPP and RESP; uncoupled: R2 = 0.67, 0.92 for GPP and RESP). Although the simulated seasonal pattern of NEP generally matched the field-based data, the correlations between field-based and simulated monthly growing season NEP were lower (R2 = 0.40, 0.00 for coupled and uncoupled simulations, respectively) in comparison to the correlations between field-based and simulated GPP and RESP. The annual NEP simulated by the coupled parameterization fell within the uncertainty of field-based estimates in two of three years. On the other hand, annual NEP simulated by the uncoupled parameterization only fell within the field-based uncertainty in one of three years. At the southern site, simulated NEP generally matched field-based NEP estimates, and the correlation between monthly growing season field-based and

  9. Factors affecting the accuracy of genomic selection for growth and wood quality traits in an advanced-breeding population of black spruce (Picea mariana).

    Science.gov (United States)

    Lenz, Patrick R N; Beaulieu, Jean; Mansfield, Shawn D; Clément, Sébastien; Desponts, Mireille; Bousquet, Jean

    2017-04-28

    identified to carry large effects, indicating a minor role for short-range LD in this population. This study supports the integration of GS models in advanced-generation tree breeding programs, given that high genomic prediction accuracy was obtained with a relatively small number of markers due to high relatedness and family structure in the population. In boreal spruce breeding programs and similar ones with long breeding cycles, much larger gain per unit of time can be obtained from genomic selection at an early age than by the conventional approach. GS thus appears highly profitable, especially in the context of forward selection in species which are amenable to mass vegetative propagation of selected stock, such as spruces.

  10. Time since death and decay rate constants of Norway spruce and European larch deadwood in subalpine forests determined using dendrochronology and radiocarbon dating

    Science.gov (United States)

    Petrillo, Marta; Cherubini, Paolo; Fravolini, Giulia; Marchetti, Marco; Ascher-Jenull, Judith; Schärer, Michael; Synal, Hans-Arno; Bertoldi, Daniela; Camin, Federica; Larcher, Roberto; Egli, Markus

    2016-03-01

    Due to the large size (e.g. sections of tree trunks) and highly heterogeneous spatial distribution of deadwood, the timescales involved in the coarse woody debris (CWD) decay of Picea abies (L.) Karst. and Larix decidua Mill. in Alpine forests are largely unknown. We investigated the CWD decay dynamics in an Alpine valley in Italy using the chronosequence approach and the five-decay class system that is based on a macromorphological assessment. For the decay classes 1-3, most of the dendrochronological samples were cross-dated to assess the time that had elapsed since tree death, but for decay classes 4 and 5 (poorly preserved tree rings) radiocarbon dating was used. In addition, density, cellulose, and lignin data were measured for the dated CWD. The decay rate constants for spruce and larch were estimated on the basis of the density loss using a single negative exponential model, a regression approach, and the stage-based matrix model. In the decay classes 1-3, the ages of the CWD were similar and varied between 1 and 54 years for spruce and 3 and 40 years for larch, with no significant differences between the classes; classes 1-3 are therefore not indicative of deadwood age. This seems to be due to a time lag between the death of a standing tree and its contact with the soil. We found distinct tree-species-specific differences in decay classes 4 and 5, with larch CWD reaching an average age of 210 years in class 5 and spruce only 77 years. The mean CWD rate constants were estimated to be in the range 0.018 to 0.022 y-1 for spruce and to about 0.012 y-1 for larch. Snapshot sampling (chronosequences) may overestimate the age and mean residence time of CWD. No sampling bias was, however, detectable using the stage-based matrix model. Cellulose and lignin time trends could be derived on the basis of the ages of the CWD. The half-lives for cellulose were 21 years for spruce and 50 years for larch. The half-life of lignin is considerably higher and may be more than

  11. Phenology of Lymantria monacha (Lepidoptera:Lymantriidae) laboratory reared on spruce foliage or a newly developed artificial diet

    Science.gov (United States)

    Melody A. Keena; Alice Vandel; Oldrich. Pultar

    2010-01-01

    Lymantria monacha (L.) (Lepidoptera: Lymantriidae) is a Eurasian pest of conifers that has potential for accidental introduction into North America. The phenology over the entire life cycle for L. monacha individuals from the Czech Republic was compared on Picea glauca (Moench) Voss (white spruce) and a newly...

  12. Dynamics and composition of litterfall in an unmanaged Norway spruce (Picea abies) forest after bark-beetle outbreak

    Czech Academy of Sciences Publication Activity Database

    Kopáček, Jiří; Cudlín, Pavel; Fluksová, H.; Kaňa, Jiří; Picek, T.; Šantrůčková, H.; Svoboda, M.; Vaněk, D.

    2015-01-01

    Roč. 20, č. 3 (2015), s. 305-323 ISSN 1239-6095 R&D Projects: GA ČR(CZ) GAP504/12/1218 Institutional support: RVO:60077344 ; RVO:67179843 Keywords : bark beetle * litter * Norway spruce Subject RIV: EH - Ecology, Behaviour Impact factor: 1.476, year: 2015

  13. Fluoroquinolone Resistance Mechanisms and population structure of Enterobacter cloacae non-susceptible to Ertapenem in North-Eastern France.

    Directory of Open Access Journals (Sweden)

    Thomas eGuillard

    2015-10-01

    Full Text Available Fluoroquinolone (FQ agents are a potential resort to treat infection due to Enterobacteriaceae producing extended spectrum β-lactamase and susceptible to FQ. In a context of increase of non-susceptibility to carbapenems among Enterobacteriaceae, we characterized FQ resistance mechanisms in 75 Enterobacter cloacae isolates non-susceptible to ertapenem in North-Eastern France in 2012 and describe the population structure by pulsed field gel electrophoresis (PFGE and multilocus sequence typing (MLST.Among them, 14.7% (12/75 carried a carbapenemase-encoding gene. Except one isolate producing VIM-1, the carbapenemase-producing isolates carried the well-known IncL/M pOXA48a plasmid. Most of the isolates (59/75 harbored at least a FQ-R determinant. qnr genes were predominant (40%, 30/75. The MLST study revealed that E. cloacae isolates’ clonality was wide (24 different STs. The more widespread STs were ST74, ST101, ST110, ST114 and ST133. Carbapenem MICs were higher for E. cloacae ST74 than for other E. cloacae isolates. PMQR determinants were more often observed in E. cloacae ST74 isolates. These findings showed that (i pOXA-48a is spreading in North-Eastern France, (ii qnr is preponderant in E. cloacae, (iii E. cloacae comprised a large amount of lineages spreading in North-Eastern France and (iv FQ as an alternative to β-lactams to treat ertapenem non-susceptible Enterobacteriaceae are compromised.

  14. Tensile strength of glulam laminations of Nordic spruce

    DEFF Research Database (Denmark)

    Hoffmeyer, Preben; Bräuner, Lise; Boström, Lars

    1999-01-01

    Design of glulam according to the European timber code Eurocode 5 is based on the standard document prEN1194 , according to which glulam beam strength is to be established either by full scale testing or by calculation. The calculation must be based on a knowledge of lamination tensile strength....... This knowledge may be obtained either by adopting a general rule that the characteristic tensile strength is sixty percent of the characteristic bending strength, or by performing tensile tests on an adequate number of laminations representative of the whole population. The present paper presents...... an investigation aimed at establishing such an adequate experimental background for the assignment of strength classes for glulam made of visually strength graded laminations from Nordic sawmills. The investigation includes more than 1800 boards (laminations) of Norway spruce (Picea abies) sampled from eight...

  15. Growth comparison of northern white-cedar to balsam fir and red spruce by site class

    Science.gov (United States)

    Philip V. Hofmeyer; Laura S. Kenefic; Robert S. Seymour; John C. Brissette

    2006-01-01

    Though northern white-cedar is a common and economically important component of the Acadian Forest of Maine and adjacent Canada, there is little regional data about the growth and development of this species. Sixty sites in northern Maine were used to compare growth of cedar to that of red spruce and balsam fir along a range of site classes and light exposures. On...

  16. Central-European mountain spruce (Picea abies (L.) Karst.) forests: regeneration of tree species after a bark beetle outbreak

    Czech Academy of Sciences Publication Activity Database

    Jonášová, Magda; Prach, Karel

    2004-01-01

    Roč. 23, č. 1 (2004), s. 15-27 ISSN 0925-8574 Institutional research plan: CEZ:AV0Z6087904 Keywords : forest management * mountain spruce forest * natural regeneration Subject RIV: GK - Forestry Impact factor: 0.890, year: 2004 http://www.sciencedirect.com

  17. Afforestation of Boreal Open Woodlands: Early Performance and Ecophysiology of Planted Black Spruce Seedlings

    OpenAIRE

    Tremblay, Pascal; Boucher, Jean-Francois; Tremblay, Marc; Lord, Daniel

    2013-01-01

    Open lichen woodlands (LWs) are degraded stands that lack the ability to regenerate naturally due to a succession of natural and/or anthropogenic disturbances. As they represent both interesting forest restoration and carbon sequestration opportunities, we tested disc scarification and planting of two sizes of containerized black spruce (Picea mariana Mill. (BSP)) seedlings for their afforestation. We compared treatment of unproductive LWs to reforestation of harvested, closed-crown black spr...

  18. Culturable bacterial populations associated with ectomycorrhizae of Norway spruce stands with different degrees of decline in the Czech Republic

    Czech Academy of Sciences Publication Activity Database

    Avidano, L.; Rinaldi, M.; Gindro, R.; Cudlín, Pavel; Martinotti, M G.; Fracchia, L.

    2010-01-01

    Roč. 56, č. 1 (2010), s. 52-64 ISSN 0008-4166 Institutional research plan: CEZ:AV0Z60870520 Keywords : Ectomycorrhizae * Norway spruce * forest decline Subject RIV: EE - Microbiology, Virology Impact factor: 1.235, year: 2010

  19. Susceptibility of burned black spruce (Picea mariana) forests to non-native plant invasions in interior Alaska

    Science.gov (United States)

    Katie V. Spellman; Christa P.H. Mulder; Teresa N. Hollingsworth

    2014-01-01

    As climate rapidly warms at high-latitudes, the boreal forest faces the simultaneous threats of increasing invasive plant abundances and increasing area burned by wildfire. Highly flammable and widespread black spruce (Picea mariana) forest represents a boreal habitat that may be increasingly susceptible to non-native plant invasion. This study assess the role of burn...

  20. Female strobili incidence in a Minnesota population of black spruce: heritability and correlation with height growth

    Science.gov (United States)

    C. Dana Nelson; C. A. Mohn

    1989-01-01

    Significant family variation in female strobili incidence, ripeness-to-flower and production were found in a Minnesota black spruce (Picea mariana (Mill.) B.S.P.) population tested at four locations. Heritability estimates indicated that gain in early flowering from selection would be possible. Height growth through age 12 years was positively correlated (genetic and...

  1. Regulation of Gene Expression during the Onset of Ligninolytic Oxidation by Phanerochaete chrysosporium on Spruce Wood

    Science.gov (United States)

    Premsagar Korripally; Christopher G. Hunt; Carl J. Houtman; Don C. Jones; Peter J. Kitin; Dan Cullen; Kenneth E. Hammel; A. A. Brakhage

    2015-01-01

    Since uncertainty remains about how white rot fungi oxidize and degrade lignin in wood, it would be useful to monitor changes in fungal gene expression during the onset of ligninolysis on a natural substrate. We grew Phanerochaete chrysosporium on solid spruce wood and included oxidant-sensing beads bearing the fluorometric dye BODIPY 581/591 in...

  2. Assessment of weather-associated causes of red spruce winter injury and consequences to aboveground carbon sequestration

    Science.gov (United States)

    Paul G. Schaberg; Brynne E. Lazarus; Gary J. Hawley; Joshua M. Halman; Catherine H. Borer; Christopher F. Hansen

    2011-01-01

    Despite considerable study, it remains uncertain what environmental factors contribute to red spruce (Picea rubens Sarg.) foliar winter injury and how much this injury influences tree C stores. We used a long-term record of winter injury in a plantation in New Hampshire and conducted stepwise linear regression analyses with local weather and regional...

  3. Ground vegetation dynamics in mountain spruce (Picea abies (L.) Karsten) forests recovering after air pollution stress impact

    Czech Academy of Sciences Publication Activity Database

    Vávrová, Eva; Cudlín, O.; Vavříček, D.; Cudlín, Pavel

    2009-01-01

    Roč. 205, č. 2 (2009), s. 305-321 ISSN 1385-0237 R&D Projects: GA MŠk(CZ) OC 141 Institutional research plan: CEZ:AV0Z60870520 Keywords : forest decline * norway spruce * microsite conditions * recovery * understorey layer Subject RIV: GK - Forestry Impact factor: 1.567, year: 2009

  4. Structure, Aboveground Biomass, and Soil Characterization of Avicennia marina in Eastern Mangrove Lagoon National Park, Abu Dhabi

    Science.gov (United States)

    Alsumaiti, Tareefa Saad Sultan

    Mangrove forests are national treasures of the United Arab Emirates (UAE) and other arid countries with limited forested areas. Mangroves form a crucial part of the coastal ecosystem and provide numerous benefits to society, economy, and especially the environment. Mangrove trees, specifically Avicennia marina, are studied in their native habitat in order to characterize their population structure, aboveground biomass, and soil properties. This study focused on Eastern Mangrove Lagoon National Park in Abu Dhabi, which was the first mangrove protected area to be designated in UAE. In situ measurements were collected to estimate Avicennia marina status, mortality rate (%), height (m), crown spread (m), stem number, diameter at breast height (cm), basal area (m), and aboveground biomass (t ha-1 ). Small-footprint aerial light detection and ranging (LIDAR) data acquired by UAE were processed to characterize mangrove canopy height and aboveground biomass density. This included extraction of LIDAR-derived height percentile statistics, segmentation of the forest into structurally homogenous units, and development of regression relationships between in situ reference and remote sensing data using a machine learning approach. An in situ soil survey was conducted to examine the soils' physical and chemical properties, fertility status, and organic matter. The data of soil survey were used to create soil maps to evaluate key characteristics of soils, and their influence on Avicennia marina in Eastern Mangrove Lagoon National Park. The results of this study provide new insights into Avicennia marina canopy population, structure, aboveground biomass, and soil properties in Abu Dhabi, as data in such arid environments is lacking. This valuable information can help in managing and preserving this unique ecosystem.

  5. Structuring and evolution of Neogene transcurrent basins in the Tellian foreland domain, north-eastern Tunisia

    Science.gov (United States)

    Melki, Fetheddine; Zouaghi, Taher; Harrab, Salah; Sainz, Antonio Casas; Bédir, Mourad; Zargouni, Fouad

    2011-07-01

    The Neogene sedimentary basins (Serravallian to Quaternary) of the Tellian tectonic foreland in north-eastern Tunisia formed within the overall NE-SW sinistral strike-slip tectonic framework of the Ras El Korane-Thibar and El Alia-Teboursouk fault systems. From stratigraphic logs, structural cross sections and interpretation of 2D seismic lines and boreholes, the pre-Neogene basement can be interpreted to be structured according to Eocene (NW-SE) compressional and Oligocene extensional phases. This basement comprises structural highs (anticlines and horsts) and subsiding areas (synclines, half-grabens and grabens) formed during the Neogene. The subsiding areas are delineated by faults striking N030E, N-S and N140E, defining (i) narrow, strongly subsiding synclines, (ii) lozenge-shaped basins and (iii) trapezoidal basins. The architecture of their fill results from the sedimentary balance between tectonics and eustatism. Halokinesis and clay diapirism (driven by Triassic and Neogene evaporites and clays) also played an important role in basin evolution, contributing to the formation of domes and diapirs along active faults.

  6. Structural model Soapaga failure from spectral correlation and magnetic gravity anomalies in the eastern cordillera, Colombia

    International Nuclear Information System (INIS)

    Rodriguez Diana Marcela; Hernandez Orlando; Kammer Andreas

    2009-01-01

    The aim of this research is to apply spectral correlation, local favorability indexes and Poisson's theorem as numerical methods for data processing and interpretation of potential field data associated with structural features; these techniques are applied to theoretical and real gravity and magnetic data of the Soapaga fault, located in the Boyaca Department, in the eastern Andean Mountains. Theoretical data of the Soapaga fault was obtained by forward modeling of geological and structural sections. Real data of the Soapaga fault included compiled gravity data and acquired magnetic data along four profiles oriented perpendicular to the fault. As a result, the geometry of the fault and its structural characteristics were obtained by interactive forward and inverse modeling. This methodology allows highlighting anomaly trends associated with density and magnetic susceptibility contrast that occur along the Soapaga fault zone. Additionally, this work provides a quantitative approach to establish the relationship between gravity and magnetic anomalies, supported by a rigorous mathematical methodology rather than isolated data interpretation to better understand the gravity and magnetic signatures of outcropping and hidden structural features.

  7. Effects of deicing salt on the vitality and health of two spruce species, Picea abies Karst., and Picea glehnii Masters planted along roadsides in northern Japan

    International Nuclear Information System (INIS)

    Kayama, M.; Quoreshi, A.M.; Kitaoka, S.; Kitahashi, Y.; Sakamoto, Y.; Maruyama, Y.; Kitao, M.; Koike, T.

    2003-01-01

    Innate physiological characters of conifers may increase uptake of sodium and chloride and result in enhanced tree injury. - In northern Japan, the growth of Picea abies Karst., and Picea glehnii Masters, which have been planted along the highways, is often suppressed due to several environmental stresses. To examine the adverse effects of deicing salt, the primary source of stress, we measured needle life span, photosynthetic capacity, and water potential and transpiration rate of the two spruce species at a site with damaged trees, near the roadside and a site with healthy trees, located far from the highway. Results from the analysis showed large amounts of sodium and chlorine in the soil and snow at the damaged site. These elements had accumulated in the needles of the spruce. Moreover, physiological traits of the spruce, at the damaged site were also affected. Therefore, we concluded that poor physiological traits might be attributed to an accumulation of deicing salt in the needles, resulting in the suppression of tree growth

  8. High-efficiency Agrobacterium-mediated transformation of Norway spruce (Picea abies) and loblolly pine (Pinus taeda)

    Science.gov (United States)

    Wenck, A. R.; Quinn, M.; Whetten, R. W.; Pullman, G.; Sederoff, R.; Brown, C. S. (Principal Investigator)

    1999-01-01

    Agrobacterium-mediated gene transfer is the method of choice for many plant biotechnology laboratories; however, large-scale use of this organism in conifer transformation has been limited by difficult propagation of explant material, selection efficiencies and low transformation frequency. We have analyzed co-cultivation conditions and different disarmed strains of Agrobacterium to improve transformation. Additional copies of virulence genes were added to three common disarmed strains. These extra virulence genes included either a constitutively active virG or extra copies of virG and virB, both from pTiBo542. In experiments with Norway spruce, we increased transformation efficiencies 1000-fold from initial experiments where little or no transient expression was detected. Over 100 transformed lines expressing the marker gene beta-glucuronidase (GUS) were generated from rapidly dividing embryogenic suspension-cultured cells co-cultivated with Agrobacterium. GUS activity was used to monitor transient expression and to further test lines selected on kanamycin-containing medium. In loblolly pine, transient expression increased 10-fold utilizing modified Agrobacterium strains. Agrobacterium-mediated gene transfer is a useful technique for large-scale generation of transgenic Norway spruce and may prove useful for other conifer species.

  9. MULTIFRACTAL STRUCTURE OF CENTRAL AND EASTERN EUROPEAN FOREIGN EXCHANGE MARKETS

    Directory of Open Access Journals (Sweden)

    Cn#259;pun#351;an Rn#259;zvan

    2012-07-01

    Full Text Available It is well known that empirical data coming from financial markets, like stock market indices, commodities, interest rates, traded volumes and foreign exchange rates have a multifractal structure. Multifractals were introduced in the field of economics to surpass the shortcomings of classical models like the fractional Brownian motion or GARCH processes. In this paper we investigate the multifractal behavior of Central and Eastern European foreign exchange rates, namely the Czech koruna, Croatian kuna, Hungarian forint, Polish zlot, Romanian leu and Russian rouble with respect to euro from January 13, 2000 to February 29, 2012. The dynamics of exchange rates is of interest for investors and traders, monetary and fiscal authorities, economic agents or policy makers. The exchange rate movements affect the international balance of payments, trade flows, and allocation of the resources in national and international economy. The empirical results from the multifractal detrending fluctuation analysis algorithm show that the six exchange rate series analysed display significant multifractality. Moreover, generating shuffled and surrogate time series, we analyze the sources of multifractality, long-range correlations and heavy-tailed distributions, and we find that this multifractal behavior can be mainly attributed to the latter. Finally, we propose a foreign exchange market inefficiency ranking by considering the multifractality degree as a measure of inefficiency. The regulators, through policy instruments, aim to improve the informational inefficiency of the markets, to reduce the associated risks and to ensure economic stabilization. Evaluation of the degree of information efficiency of foreign exchange markets, for Central and Eastern Europe countries, is important to assess to what extent these countries are prepared for the transition towards fully monetary integration. The weak form efficiency implies that the past exchange rates cannot help to

  10. Investigation of cadmium pollution in the spruce saplings near the metal production factory.

    Science.gov (United States)

    Hashemi, Seyed Armin; Farajpour, Ghasem

    2016-02-01

    Toxic metals such as lead and cadmium are among the pollutants that are created by the metal production factories and disseminated in the nature. In order to study the quantity of cadmium pollution in the environment of the metal production factories, 50 saplings of the spruce species at the peripheries of the metal production factories were examined and the samples of the leaves, roots, and stems of saplings planted around the factory and the soil of the environment of the factory were studied to investigate pollution with cadmium. They were compared to the soil and saplings of the spruce trees planted outside the factory as observer region. The results showed that the quantity of pollution in the leaves, stems, and roots of the trees planted inside the factory environment were estimated at 1.1, 1.5, and 2.5 mg/kg, respectively, and this indicated a significant difference with the observer region (p metal production factory was estimated at 6.8 mg/kg in the depth of 0-10 cm beneath the level of the soil. The length of roots in the saplings planted around the factory of metal production stood at 11 and 14.5 cm in the observer region which had a significant difference with the observer region (p production processes in the factory. © The Author(s) 2013.

  11. Lead mobility within the xylem of red spruce seedlings: Implications for the development of pollution histories

    Science.gov (United States)

    John R. Donnelly; John B. Shane; Paul G. Schaberg

    1990-01-01

    Development of Pb pollution histories using tree ring analyses has been troubled by possible mobility of Pb within stem xylem. In a 2-yr study, we exposed red spruce (Picea rubens Sarg.) seedlings to Pb during one growing season, with Pb excluded in either the previous or following growing season. Lead levels within xylem rings and bark were...

  12. Shoot-level terpenoids emission in Norway spruce (Picea abies) under natural field and manipulated laboratory conditions

    Czech Academy of Sciences Publication Activity Database

    Esposito, R.; Lusini, I.; Večeřová, Kristýna; Holišová, Petra; Pallozzi, E.; Guidolotti, G.; Urban, Otmar; Calfapietra, Carlo

    2016-01-01

    Roč. 108, nov (2016), s. 530-538 ISSN 0981-9428 R&D Projects: GA MŠk(CZ) LO1415; GA MŠk(CZ) LD13031; GA MŠk(CZ) LM2015061 Institutional support: RVO:67179843 Keywords : Norway spruce * BVOC * Monoterpenes * Temperature * Ozone * Sun-shade * Stress Subject RIV: EF - Botanics Impact factor: 2.724, year: 2016

  13. Enhanced UV-B radiation may lead to the decrease of specific phenolic compounds in Norway spruce needles

    Czech Academy of Sciences Publication Activity Database

    Urban, Otmar; Vrchotová, Naděžda; Tříska, Jan; Marek, Michal V.

    2008-01-01

    Roč. 17, č. 5 (2008), s. 564-567 ISSN 1018-4619 R&D Projects: GA MŠk 2B06068 Institutional research plan: CEZ:AV0Z60870520 Keywords : 4-hydroxyacetophenone * catechin * Norway spruce * picein * screening pigments * ultraviolet radiation Subject RIV: ED - Physiology Impact factor: 0.463, year: 2008

  14. REGIONALIZATION IN EASTERN AND CENTRAL EUROPE: OBSTACLES AND PERSPECTIVES

    Directory of Open Access Journals (Sweden)

    Gyula Horváth

    2012-01-01

    Full Text Available Regional policy depends on efficient administrative systems for designing and implementing strategies, and places considerable demands on Member States’ public administrations in terms of e.g. financial management and monitoring; project selection procedures; ex ante environmental impact assessments and cost-benefit analyses; and the monitoring and evaluation of outputs, results and impacts. EU member states have taken a range of different approaches to the administration of regional policy.The construction of regions in the countries of Eastern and Central Europe became one of the important debate topics for preparation for EU membership. Despite the numerous similarities in the changes that have taken place in the territorial structures of the Eastern and Central European countries, the differences in the responses individual countries gave to the challenges of regional development and the varied results of their development efforts demonstrate that the “Eastern European Bloc” is at least as heterogeneous as the former member states of the European Union. EU accession opened up a Pandora’s Box in the countries of Eastern and Central Europe. The fundamental issue of how unitarily structured states can be set on a decentralised path became the centre of debate. The paper introduces the Central and Eastern European achievements of region building processes and searches for an explanation of the reasons for the difficulties of Eastern and Central Europe in regional construction; it summarises the administrative and political development pre-requisites of the transition to a regional outline of the possible advantages of a regional institutional system in the creation of the Cohesion Policy ensuring a decrease in regional differences.

  15. Mummy Lake: An unroofed ceremonial structure within a large-scale ritual landscape

    Science.gov (United States)

    Benson, Larry V.; Griffin, Eleanor R.; Stein, J.R.; Friedman, R. A.; Andrae, S. W.

    2014-01-01

    The structure at Mesa Verde National Park known historically as Mummy Lake and more recently as Far View Reservoir is not part of a water collection, impoundment, or redistribution system. We offer an alternative explanation for the function of Mummy Lake. We suggest that it is an unroofed ceremonial structure, and that it serves as an essential component of a Chacoan ritual landscape. A wide constructed avenue articulates Mummy Lake with Far View House and Pipe Shrine House. The avenue continues southward for approximately 6 km where it apparently divides connecting with Spruce Tree House and Sun Temple/Cliff Palace. The avenue has previously been interpreted as an irrigation ditch fed by water impounded at Mummy Lake; however, it conforms in every respect to alignments described as Chacoan roads. Tree-ring dates indicate that the construction of Spruce Tree House and Cliff Palace began about A.D. 1225, roughly coincident with the abandonment of the Far View community. This pattern of periodically relocating the focus of an Anasazi community by retiring existing ritual structures and linking them to newly constructed facilities by means of broad avenues was first documented by Fowler and Stein (1992) in Manuelito Canyon, New Mexico. Periods of intense drought appear to have contributed to the relocation of prehistoric Native Americans from the Far View group to Cliff Palace/Spruce Tree House in the mid-13th century and eventually to the abandonment of all Anasazi communities in southwestern Colorado in the late-13th century.

  16. Crustal Structure of the Ionian Basin and Eastern Sicily Margin: Results From a Wide-Angle Seismic Survey

    Science.gov (United States)

    Dellong, David; Klingelhoefer, Frauke; Kopp, Heidrun; Graindorge, David; Margheriti, Lucia; Moretti, Milena; Murphy, Shane; Gutscher, Marc-Andre

    2018-03-01

    In the Ionian Sea (central Mediterranean) the slow convergence between Africa and Eurasia results in the formation of a narrow subduction zone. The nature of the crust of the subducting plate remains debated and could represent the last remnants of the Neo-Tethys ocean. The origin of the Ionian basin is also under discussion, especially concerning the rifting mechanisms as the Malta Escarpment could represent a remnant of this opening. This subduction retreats toward the south-east (motion occurring since the last 35 Ma) but is confined to the narrow Ionian basin. A major lateral slab tear fault is required to accommodate the slab roll-back. This fault is thought to propagate along the eastern Sicily margin but its precise location remains controversial. This study focuses on the deep crustal structure of the eastern Sicily margin and the Malta Escarpment. We present two two-dimensional P wave velocity models obtained from forward modeling of wide-angle seismic data acquired onboard the R/V Meteor during the DIONYSUS cruise in 2014. The results image an oceanic crust within the Ionian basin as well as the deep structure of the Malta Escarpment, which presents characteristics of a transform margin. A deep and asymmetrical sedimentary basin is imaged south of the Messina strait and seems to have opened between the Calabrian and Peloritan continental terranes. The interpretation of the velocity models suggests that the tear fault is located east of the Malta Escarpment, along the Alfeo fault system.

  17. Long-term effects of CO2 enrichment on bud phenology and shoot growth patterns of Norway spruce juvenile trees

    Czech Academy of Sciences Publication Activity Database

    Pokorný, Radek; Tomášková, Ivana; Drápelová, I.; Kulhavý, J.; Marek, Michal V.

    2010-01-01

    Roč. 56, č. 6 (2010), s. 251-257 ISSN 1212-4834 R&D Projects: GA AV ČR IAA600870701 Institutional research plan: CEZ:AV0Z60870520 Keywords : bud * elevated CO2 * Norway spruce Subject RIV: EH - Ecology, Behaviour

  18. Fine root status element contents in three Norway spruce stands in the Krkonose Mts

    Czech Academy of Sciences Publication Activity Database

    Goldbold, D.; Fritz, H.; Cudlín, Pavel; Bonifacio, E.

    2003-01-01

    Roč. 22, Suppl. 1 (2003), s. 91-94 ISSN 1335-342X. [Long Term Air Pollution Effect on Forest Ecosystems (International Meeting for Specialists in Air Pollution Effects on Forest Ecosystems)/20./. Zvolen, 30.08.2002-01.09.2002] R&D Projects: GA MŠk OK 355 Institutional research plan: CEZ:AV0Z6087904 Keywords : Ca:Al ratios, fine roots, spruce Subject RIV: EH - Ecology, Behaviour Impact factor: 0.100, year: 2003

  19. Genetic differentiation and phylogeography of Mediterranean-North Eastern Atlantic blue shark (Prionace glauca, L. 1758 using mitochondrial DNA: panmixia or complex stock structure?

    Directory of Open Access Journals (Sweden)

    Agostino Leone

    2017-12-01

    Full Text Available Background The blue shark (Prionace glauca, Linnaeus 1758 is one of the most abundant epipelagic shark inhabiting all the oceans except the poles, including the Mediterranean Sea, but its genetic structure has not been confirmed at basin and interoceanic distances. Past tagging programs in the Atlantic Ocean failed to find evidence of migration of blue sharks between the Mediterranean and the adjacent Atlantic, despite the extreme vagility of the species. Although the high rate of by-catch in the Mediterranean basin, to date no genetic study on Mediterranean blue shark was carried out, which constitutes a significant knowledge gap, considering that this population is classified as “Critically Endangered”, unlike its open-ocean counterpart. Methods Blue shark phylogeography and demography in the Mediterranean Sea and North-Eastern Atlantic Ocean were inferred using two mitochondrial genes (Cytb and control region amplified from 207 and 170 individuals respectively, collected from six localities across the Mediterranean and two from the North-Eastern Atlantic. Results Although no obvious pattern of geographical differentiation was apparent from the haplotype network, Φst analyses indicated significant genetic structure among four geographical groups. Demographic analyses suggest that these populations have experienced a constant population expansion in the last 0.4–0.1 million of years. Discussion The weak, but significant, differences in Mediterranean and adjacent North-eastern Atlantic blue sharks revealed a complex phylogeographic structure, which appears to reject the assumption of panmixia across the study area, but also supports a certain degree of population connectivity across the Strait of Gibraltar, despite the lack of evidence of migratory movements observed by tagging data. Analyses of spatial genetic structure in relation to sex-ratio and size could indicate some level of sex/stage biased migratory behaviour.

  20. The influence of red wood ants .i.Formica polyctena./i. on nutrient availability ant growth of spruce tree

    Czech Academy of Sciences Publication Activity Database

    Frouz, Jan; Rybníček, M.; Cudlín, Pavel

    2007-01-01

    Roč. 9, - (2007) ISSN 1607-7962. [European Geosciences Union General Assembly 2007. 15.04.2007-20.04.2007, Vienna] Institutional research plan: CEZ:AV0Z60660521; CEZ:AV0Z60870520 Keywords : Formica polyctena * nutrient availability * growth of spruce tree Subject RIV: EH - Ecology, Behaviour

  1. Regional Instability in the Abundance of Open Stands in the Boreal Forest of Eastern Canada

    Directory of Open Access Journals (Sweden)

    Rija Rapanoela

    2016-05-01

    Full Text Available Fires are a key disturbance of boreal forests. In fact, they are the main source of renewal and evolution for forest stands. The variability of fire through space and time results in a diversified forest mosaic, altering their species composition, structure and productivity. A resilient forest is assumed to be in a state of dynamic equilibrium with the fire regime, so that the composition, age structure and succession stages of forests should be consistent with the fire regime. Dense spruce-moss stands tend, however, to diminish in favour of more open stands similar to spruce-lichen stands when subjected to more frequent and recurring disturbances. This study therefore focused on the effects of spatial and temporal variations in burn rates on the proportion of open stands over a large geographic area (175,000 km2 covered by black spruce (Picea mariana (Mill. Britton, Sterns, Poggenb.. The study area was divided into 10 different zones according to burn rates, as measured using fire-related data collected between 1940 and 2006. To test if the abundance of open stands was unstable over time and not in equilibrium with the current fire regime, forest succession was simulated using a landscape dynamics model that showed that the abundance of open stands should increase progressively over time in zones where the average burn rate is high. The proportion of open stands generated during a specific historical period is correlated with the burn rate observed during the same period. Rising annual burn rates over the past two decades have thereby resulted in an immediate increase in the proportion of open stands. There is therefore a difference between the current proportion of open stands and the one expected if vegetation was in equilibrium with the disturbance regime, reflecting an instability that may significantly impact the way forest resources are managed. It is apparent from this study that forestry planning should consider the risks associated

  2. Recent evolution and divergence among populations of a rare Mexican endemic, Chihuahua spruce, following holocene climatic warming

    Science.gov (United States)

    F. Thomas Ledig; Virginia Jacob-Cervantes; Paul D. Hodgskiss

    1997-01-01

    Fragmentation and reduction in population size are expected to reduce genetic diversity. However, examples from natural populations of forest trees are scarce. The range of Chihuahua spruce retreated northward and fragmented coincident with the warming climate that marked the early Holocene. The isolated populations vary from 15 to 2441 trees, which provided an...

  3. Fine and coarse root parameters from mature black spruce displaying genetic x soil moisture interaction in growth

    Science.gov (United States)

    John E. Major; Kurt H. Johnsen; Debby C. Barsi; Moira Campbell

    2012-01-01

    Fine and coarse root biomass, C, and N mass parameters were assessed by root size and soil depths from soil cores in plots of 32-year-old black spruce (Picea mariana (Mill.) Britton, Sterns & Poggenb.) from four full-sib families studied previously for drought tolerance and differential productivity on a dry and wet...

  4. Ecophysiology of seedling establishment in contrasting spruce-fir forests of southern Appalachian and Rocky Mountain ecotones, USA

    Science.gov (United States)

    William K. Smith; Keith N.C. Reinhardt; Daniel M. Johnson

    2010-01-01

    Fraser fir (Abies fraseri [Pursh] Poiret) and red spruce (Picea rubens Sarg.) occur as codominant trees in six relic, mountain-top populations that make up the high-elevation forests of the Southern Appalachian Mountains (SA). These two relic species of the former boreal forest have experienced a significant decline over the past...

  5. Relationships among foliar chemistry, foliar polyamines, and soil chemistry in red spruce trees growing across the northeastern United States

    Science.gov (United States)

    Minocha, R.; Shortle, W.C.; Lawrence, G.B.; David, M.B.; Minocha, S.C.

    1997-01-01

    Forest trees are constantly exposed to various types of natural and anthropogenic stressors. A major long-term goal of our research is to develop a set of early physiological and biochemical markers of stress in trees before the appearance of visual symptoms. Six red spruce (Picea rubens Sarg.) stands from the northeastern United States were selected for collection of soil and foliage samples. All of the chosen sites had soil solution pH values below 4.0 in the Oa horizon but varied in their geochemistry. Some of these sites were apparently under some form of environmental stress as indicated by a large number of dead and dying red spruce trees. Samples of soil and needles (from apparently healthy red spruce trees) were collected from these sites four times during a two-year period. The needles were analyzed for perchloric acid-soluble polyamines and exchangeable inorganic ions. Soil and soil solution samples from the Oa and B horizons were analyzed for their exchange chemistry. The data showed a strong positive correlation between Ca and Mg concentrations in the needles and in the Oa horizon of the soil. However, needles from trees growing on relatively Ca-rich soils with a low exchangeable Al concentration and a low Al:Ca soil solution ratio had significantly lower concentrations of putrescine and spermidine than those growing on Ca-poor soils with a high exchangeable Al concentration and a high Al:Ca soil solution in the Oa horizon. The magnitude of this change was several fold higher for putrescine concentrations than for spermidine concentrations. Neither putrescine nor spermidine were correlated with soil solution Ca, Mg, and Al concentrations in the B horizon. The putrescine concentrations of the needles always correlated significantly with exchangeable Al (r2=0.73, p???0.05) and still solution Al:Ca ratios (r2=0.91, p???0.01) of the Oa horizon. This suggests that in conjunction with soil chemistry, putrescine and/or spermidine may be used as a potential

  6. Role of bark and wood destroying insect pests in drying off of spruce and pines in planting weakened by smoke

    Energy Technology Data Exchange (ETDEWEB)

    Kudela, M; Wolf, R

    1963-01-01

    This paper describes a detailed study made in 1958-62, indicating the part played by smoke and the various groups and individual species of insects, in the mortality in middle-aged and mature Pine and Spruce stands.

  7. Population genetic structure of Monimopetalum chinense (Celastraceae), an endangered endemic species of eastern China.

    Science.gov (United States)

    Xie, Guo-Wen; Wang, De-Lian; Yuan, Yong-Ming; Ge, Xue-Jun

    2005-04-01

    Monimopetalum chinense (Celastraceae) standing for the monotypic genus is endemic to eastern China. Its conservation status is vulnerable as most populations are small and isolated. Monimopetalum chinense is capable of reproducing both sexually and asexually. The aim of this study was to understand the genetic structure of M. chinense and to suggest conservation strategies. One hundred and ninety individuals from ten populations sampled from the entire distribution area of M. chinense were investigated by using inter-simple sequence repeats (ISSR). A total of 110 different ISSR bands were generated using ten primers. Low levels of genetic variation were revealed both at the species level (Isp=0.183) and at the population level (Ipop=0.083). High clonal diversity (D = 0.997) was found, and strong genetic differentiation among populations was detected (49.06 %). Small population size, possible inbreeding, limited gene flow due to short distances of seed dispersal, fragmentation of the once continuous range and subsequent genetic drift, may have contributed to shaping the population genetic structure of the species.

  8. Combined fluorescence, reflectance, and ground measurements of a stressed Norway spruce forest for forest damage assessment

    Science.gov (United States)

    Banninger, C.

    1991-01-01

    The detection and monitoring of stress and damage in forested areas is of utmost importance to forest managers for planning purposes. Remote sensing are the most suitable means to obtain this information. This requires that remote sensing data employed in a forest survey be properly chosen and utilized for their ability to measure canopy spectral features directly related to key tree and canopy properties that are indicators of forest health and vitality. Plant reflectance in the visible to short wave IR regions (400 to 2500 nm) provides information on its biochemical, biophysical, and morphological make up, whereas plant fluorescence in the 400 to 750 nm region is more indicative of the capacity and functioning of its photosynthetic apparatus. A measure of both these spectral properties can be used to provide an accurate assessment of stress and damage within the forest canopy. Foliar chlorophyll and nitrogen are essential biochemical constituents required for the proper functioning and maintenance of a plant's biological processes. Chlorophyll-a is the prime reactive center for photosynthesis, by which a plant converts CO2 and H2O into necessary plant products. Nitrogen forms an important component of the amino-acids, enzymes, proteins, alkaloids, and cyanogenic compounds that make up a plant, including its pigments. Both chlorophyll and nitrogen have characteristic absorption features in the visible to short wave IR region. By measuring the wavelength position and depth of these features and the fluorescence response of the foliage, the health and vitality of a canopy can be ascertained. Examples for a stressed Norway spruce forest in south-eastern Austria are presented.

  9. High rates of solar radiation - an important natural stress factor of the photosynthetic activity of mountainous norway spruce stands

    International Nuclear Information System (INIS)

    Sprtova, M.; Marek, M.V.

    1996-01-01

    Photosynthetic activity can be regarded as the basis of biomass productivity and vitality of forest trees, respectively. Moreover, this activity is under the strong influence of environment. Excess of photosynthetically active radiation (PhAR) can be a harmful factor of environment which is the reason of photoinhibition. Photoinhibition is demonstrated by a decrease of photosynthetic rate. An analysis of the influence of PhAR excess on function of the assimilatory apparatus of Norway spruce during summer days was done. The strong influence of PhAR excess on values of parameters of photosynthesis reflecting changes in the level of quanta capture and electron transport chain was observed. The comprehensive description of the method of chlorophyll a is given. Excess of PhAR caused rapid changes of assimilatory apparatus function and thus this PhAR excess can be regarded as a significant stress of productional activity of Norway spruce stands under field conditions

  10. Actin distribution in mitotic apparatus of somatic embryo cells of Norway spruce (Picea abies (L.) Karst.)

    Czech Academy of Sciences Publication Activity Database

    Cenklová, Věra; Binarová, Pavla; Havel, L.

    2003-01-01

    Roč. 46, č. 2 (2003), s. 167-174 ISSN 0006-3134 R&D Projects: GA ČR GA204/99/D092; GA AV ČR IAA5020803; GA ČR GV522/96/K186 Institutional research plan: CEZ:AV0Z5038910 Keywords : cytoskeleton * embryogenesis of spruce * mitosis Subject RIV: EF - Botanics Impact factor: 0.919, year: 2003

  11. Severe red spruce winter injury in 2003 creates unusual ecological event in the northeastern United States

    Science.gov (United States)

    Brynne E. Lazarus; Paul G. Schaberg; Donald H. DeHayes; Gary J. Hawley

    2004-01-01

    Abundant winter injury to the current-year (2002) foliage of red spruce (Picea rubens Sarg.) became apparent in the northeastern United States in late winter of 2003. To assess the severity and extent of this damage, we measured foliar winter injury at 28 locations in Vermont and surrounding states and bud mortality at a subset of these sites. Ninety percent of all...

  12. Seismically-triggered soft-sediment deformation structures close to a major strike-slip fault system in the Eastern Alps (Hirlatz cave, Austria)

    Science.gov (United States)

    Salomon, Martina Lan; Grasemann, Bernhard; Plan, Lukas; Gier, Susanne; Schöpfer, Martin P. J.

    2018-05-01

    We investigate episodic soft-sediment deformation structures cross-cut by normal faults preserved in unlithified finely laminated calcite rich sediments in the Hirlatz cave in the Northern Calcareous Alps (Austria). These sediments comprise varve-like alternations of brighter carbonate/quartz rich layers, and darker clay mineral rich layers. The deformed sediments contain abundant millimeter to centimeter-scale soft-sediment structures (load casts, ball-and-pillow structures), sheet slumps (thrust faults and folds), erosive channels filled with slides and chaotic slumps. After deposition and soft-sediment deformation normal faults developed within the entire sedimentary succession, an event that probably correlates with an offset of c. 10 cm of the passage wall above the outcrop. Our major conclusions are: (i) The sediments have a glacial origin and were deposited in the Hirlatz cave under phreatic fluvio-lacustrine conditions. The deposition and the soft-sediment deformation occurred most likely during the last glaciation (i.e. around 25 ka ago); (ii) The liquefaction and formation of the soft-sediment structures in water-saturated stratified layers was triggered by episodic seismic events; (iii) The internally deformed sediments were later displaced by normal faults; (iv) A possible source for the seismic events is the active sinistral Salzach-Ennstal-Mariazeller-Puchberger (SEMP) strike-slip fault which is located about 10 km south of the outcrop and plays a major role in accommodating the extrusion of the Eastern Alps towards the Pannonian Basin. To our knowledge, the described structures are the first report of liquefaction and seismically induced soft-sediment deformations in Quaternary sediments in the Eastern Alps.

  13. Observation on the effect of gamma rays on spruce (Picea smithiana) seeds germination

    International Nuclear Information System (INIS)

    Sharma, S.K.

    1997-01-01

    Spruce (Picea smithiana) forests cover extensive areas in western Himalayas and account for 48.80% of the total conifer forest area and growing stock in the region. Baldwin studied the effect of x-rays on the seeds of conifers trees. A little work has been done on the genus Picea in respect of physical mutagens. Keeping this in consideration the present trial has been done at preliminary level to study the effect of gamma rays on the germination behaviour. (author). 3 refs., 1 tab

  14. Influence of fiber degradation and concentration of fermentable sugars on simultaneous saccharification and fermentation of high-solids spruce slurry to ethanol.

    Science.gov (United States)

    Hoyer, Kerstin; Galbe, Mats; Zacchi, Guido

    2013-10-08

    Saccharification and fermentation of pretreated lignocellulosic materials, such as spruce, should be performed at high solids contents in order to reduce the cost of the produced bioethanol. However, this has been shown to result in reduced ethanol yields or a complete lack of ethanol production. Previous studies have shown inconsistent results when prehydrolysis is performed at a higher temperature prior to the simultaneous saccharification and fermentation (SSF) of steam-pretreated lignocellulosic materials. In some cases, a significant increase in overall ethanol yield was reported, while in others, a slight decrease in ethanol yield was observed. In order to investigate the influence of prehydrolysis on high-solids SSF of steam-pretreated spruce slurry, in the present study, the presence of fibers and inhibitors, degree of fiber degradation and initial fermentable sugar concentration has been studied. SSF of whole steam-pretreated spruce slurry at a solids content of 13.7% water-insoluble solids (WIS) resulted in a very low overall ethanol yield, mostly due to poor fermentation. The yeast was, however, able to ferment the washed slurry and the liquid fraction of the pretreated slurry. Performing prehydrolysis at 48°C for 22 hours prior to SSF of the whole pretreated slurry increased the overall ethanol yield from 3.9 to 62.1%. The initial concentration of fermentable sugars in SSF could not explain the increase in ethanol yield in SSF with prehydrolysis. Although the viscosity of the material did not appear to decrease significantly during prehydrolysis, the degradation of the fibers prior to the addition of the yeast had a positive effect on ethanol yield when using whole steam-pretreated spruce slurry. The results of the present study suggest that the increase in ethanol yield from SSF when performing prehydrolysis is a result of fiber degradation rather than a decrease in viscosity. The increased concentration of fermentable sugars at the beginning of the

  15. Partitioning of ecosystem respiration in a paludified shallow-peat spruce forest in the southern taiga of European Russia

    International Nuclear Information System (INIS)

    Kurbatova, J; Tatarinov, F; Varlagin, A; Avilov, V; Molchanov, A; Kozlov, D; Ivanov, D; Valentini, R

    2013-01-01

    Soil, tree stems, and ecosystem carbon dioxide fluxes were measured by chambers and eddy covariance methods in a paludified shallow-peat spruce forest in the southern taiga of European Russia (Tver region, 56° N 33° E) during the growing seasons of 2002–2012. The site was established in 1998 as part of the EUROSIBERIAN CARBONFLUX project, an international field experiment examining atmosphere–biosphere interaction in Siberia and European Russia. In all years the observed annual cumulative net ecosystem flux was positive (the forest was a source of carbon to the atmosphere). Soil and tree stem respiration was a significant part of the total ecosystem respiration (ER) in this paludified shallow-peat spruce forest. On average, 49% of the ER came from soil respiration. We found that the soil fluxes exhibited high seasonal variability, ranging from 0.7 to 10 μmol m −2  s −1 . Generally, the soil respiration depended on the soil temperature and ground water level. In drought conditions, the soil respiration was low and did not depend on temperature. The stem respiration of spruces grew intensively in May, had permanently high values from June to the end of September, and in October it dramatically decreased. The tree stem respiration in midsummer was about 3–5 μmol m −2  s −1 for dominant trees and about 1–2 μmol m −2  s −1 for subdominant trees. The respiration of living tree stems was about 10–20% of the ER. (letter)

  16. Norway spruce needles as bioindicator of air pollution in the area of influence of the Sostanj Thermal Power Plant, Slovenia

    Energy Technology Data Exchange (ETDEWEB)

    Samar Al Sayegh Petkovsek; Franc Batic; Cvetka Ribaric Lasnik [ERICo Velenje, Velenje (Slovenia). Ecological Research and Industrial Cooperation Institute

    2008-01-15

    This paper reports the results of total sulphur content, photosynthetic pigments, ascorbic acid (vitamin C) and {alpha}-tocopherol (vitamin E) analysed in current-year needles of Norway spruce (Picea abies (L.) Karst.) in the area influenced by sulphur emissions from the Sostanj Thermal Power Plant (STPP), Slovenia, in the period 1991-2004. Ten differently polluted sampling sites in the emission area of STPP were selected. After desulphurization of emission gases from STPP total sulphur content in needles decreased and vitality parameters of needles increased. Moreover, a strong correlation between the average annual emissions of SO{sub 2} from STPP and average annual sulphur content (increase) or average annual chlorophyll content (decrease) in current-year needles was found. The results showed that spruce needles may be an useful bioindicator for detecting changes in the emission rates of SO{sub 2}.

  17. Comparing the Costs and Revenues of Transformation to Continuous Cover Forestry for Sitka Spruce in Great Britain

    Directory of Open Access Journals (Sweden)

    Owen Davies

    2015-07-01

    Full Text Available Recently continuous cover forestry (CCF has become an accepted approach to forest management in Britain, but uncertainty about its economic consequences may be a barrier to its wider use. A study was carried out to examine the costs and revenues of transforming a stand of Sitka spruce (Picea sitchensis (Bong. Carr. to CCF. The main conclusion is that transformation to CCF need not be more costly than clearfelling and replanting if natural regeneration is successful and the aim is to produce a simple canopy structure. The long-term value of transformation to a more complex canopy structure, with three or more strata, is lower and the extra costs need to be justified in terms of management objectives. The main output from the study is an analysis spreadsheet that empowers practitioners and policy makers to investigate the effects of costs, revenues and discount rates on estimates of net present value over 20 years, 100 years and in perpetuity, to suit local conditions. This paper summarises the method and results of the study in a British context, sets these in a wider international context, and considers the merits, applications and possible further developments of the approach.

  18. Estimation of Spruce Needle-Leaf Chlorophyll Content Based on DART and PARAS Canopy Reflectance Models

    Czech Academy of Sciences Publication Activity Database

    Yáñez-Rausell, L.; Malenovský, Z.; Rautiainen, M.; Clevers, J G P W.; Lukeš, Petr; Hanuš, Jan; Schaepman, M. E.

    2015-01-01

    Roč. 8, č. 4 (2015), s. 1534-1544 ISSN 1939-1404 R&D Projects: GA MŠk(CZ) ED1.1.00/02.0073 Institutional support: RVO:67179843 Keywords : Chlorophyll a plus b estimation * CHRIS-PROBA * coniferous forest * continuum removal * discrete anisotropic radiative transfer model (DART) * needle-leaf * Norway spruce * optical indices * PARAS * PROSPECT * radiative transfer * recollision probability Subject RIV: EH - Ecology, Behaviour Impact factor: 2.145, year: 2015

  19. Seismic reflection and structuring characterization of deep aquifer system in the Dakhla syncline (Cap Bon, North-Eastern Tunisia)

    Science.gov (United States)

    Bellali, Abir; Jarraya Horriche, Faten; Gabtni, Hakim; Bédir, Mourad

    2018-04-01

    The Dakhla syncline is located in the North-Eastern Tunisia. It is bounded by Abd El Rahmene anticline to the North-West, El Haouaria Graben to the North-East, Grombalia Graben to the South-West and the Mediterranean Sea to the East. The main aquifer reservoirs of Dakhla syncline are constituted by stacks of fluvial to deltaic Neogene sequences and carbonates. The interpretation of eight seismic reflection profiles, calibrated by wire line logging data of three oil wells, hydraulic wells and geologic field sections highlighted the impact of tectonics on the structuring geometry of aquifers and their distribution in elevated structures and subsurface depressions. Lithostratigraphic correlations and seismic profiles analysis through the syncline show that the principal aquifers are thickest within the central and northern part of the study area and thinnest to the southern part of the syncline. Seismic sections shows that the fracture/fault pattern in this syncline is mainly concentrated along corridors with a major direction of NW-SE and secondary directions of N-S, E-W and NE-SW with different release. This is proved by the complexity structure of Eastern Tunisia, resulted from the interaction between the African and Eurasiatic plates. Isochron maps of aquifers systems exhibited the structuring of this syncline in sub-surface characterized by important lateral and vertical geometric and thickness variations. Seismic sections L1, L2, L3, L4, L5 and petroleum wells showed an heterogeneous multilayer aquifers of Miocene formed by the arrangement of ten sandstone bodies, separated by impermeable clay packages. Oligo-Miocene deposits correspond to the most great potential aquifers, with respectively an average transmissivity estimated: Somaa aquifer 6.5 10-4 m2/s, Sandstone level aquifer 2.6 10-3 m2/s, Beglia aquifer 1.1 10-3 m2/s, Ain Ghrab aquifer 1.3 10-4 m2/s and Oligocene aquifer 2 10-3 m2/s. The interpretation of spatial variations of seismic units and the

  20. Radiation Oncology in the Developing Economies of Central and Eastern Europe.

    Science.gov (United States)

    Esiashvili, Natia

    2017-04-01

    Eastern Europe is represented by 22 countries of significant variability in population density and degree of economic development. They have been affected by past geopolitical isolation due to their association with the "Soviet Block." Currently, all Eastern European countries except Slovenia are low- or middle-income level and 10 of them are part of European Union. Health care systems in Central and Eastern Europe have been influenced by the legacy of centralized soviet-era governance; however, most countries, particularly in European Union zone, have gone through health care reforms directed toward modernizing infrastructure and staffing. The level of health financing available through health insurance has increased in the region, although still lags behind the Western European levels. After adjusting for differing population age structures, overall incidence rates in both sexes are lower in Eastern and Central Europe compared with the Northern and Western European countries; however, mortality remains higher. There is an ongoing shortage of oncology services in Eastern Europe, including radiotherapy equipment and personnel. Eastern European radiotherapy field is highly diverse with large differences among countries regarding staffing structure, training, accreditation, and defined roles and responsibilities. The rapid diffusion of technological innovations has been identified as one of the most important factors driving the escalating health care expenses, and the need for better cost-effective solutions applicable to the local health care systems and levels of economic development. Copyright © 2017 Elsevier Inc. All rights reserved.

  1. Laboratory and Field Evaluation of the Entomopathogenic Fungus Beauveria bassiana (Deuteromycotina: Hyphomycetes) for Population Management of Spruce Beetle, Dendroctonus rufipennis (Coleoptera: Scolytinae), in Felled Trees and Factors Limiting Pathogen Success.

    Science.gov (United States)

    Davis, Thomas Seth; Mann, Andrew J; Malesky, Danielle; Jankowski, Egan; Bradley, Clifford

    2018-03-24

    An isolate of the entomopathogenic fungus Beauveria bassiana (Bals.) Vuill. (Deuteromycotina: Hyphomycetes) was tested for its ability to reduce survival and reproduction of spruce beetle, Dendroctonus rufipennis (Kirby) (Coleoptera: Scolytinae), under laboratory and field conditions. Conidial suspension applied directly to adults or to filter papers that adults contacted had a median survival time of 3-4 d in laboratory assays and beetles died more rapidly when exposed to conidial suspension than when treated with surfactant solution only. In the field, conidial suspension was applied to the surface of felled and pheromone-baited Engelmann spruce (Picea engelmannii) trees using a backpack sprayer. Mortality of colonizing parent beetles (F0), reproduction (abundance of F1 offspring in logs), and emergence of F1 beetles from logs was compared between treated and nontreated logs. Application of spore suspension increased mortality of F0 adults by 36% on average. Total F1 reproduction was reduced by 17% and emergence from logs was reduced by 13% in treated logs, but considerable variability in reproduction and emergence was observed. Viable spores were re-isolated from treated logs up to 90 d after application, indicating that spores are capable of long-term persistence on the tree bole microhabitat. Subsequent in vitro tests revealed that temperatures below 15°C and exposure to spruce monoterpenes likely limit performance of B. bassiana under field conditions, but exposure to low-intensity light or interactions with spruce beetle symbiotic fungi were not strongly inhibitory. It is concluded that matching environmental tolerances of biocontrol fungi to field conditions can likely improve their usefulness for control of spruce beetle in windthrown trees.

  2. Rearing Glypta Fumiferanae [hym.:Ichneumonida] on a multivoltine laboratory colony of the Western Spruce Budworm (Choristoneura Occidentalis) [LEP.:Tortricidae

    Science.gov (United States)

    Nancy Rappaport; Marion Page

    1985-01-01

    Methods were devloped for rearing Glypta fumiferanae Viereck on a nondiapausing laboratory colony of the western spruce budworm, Choristoneura occidentalis Freeman. Both host and parasite are univoltine and undergo diapause in nature. In this study, the parasite's voltinism was synchronized with that of a nondiapausing...

  3. Comparing modern and presettlement forest dynamics of a subboreal wilderness: Does spruce budworm enhance fire risk?

    Science.gov (United States)

    Sturtevant, Brian R.; Miranda, Brian R.; Shinneman, Douglas J.; Gustafson, Eric J.; Wolter, Peter T.

    2012-01-01

    Insect disturbance is often thought to increase fire risk through enhanced fuel loadings, particularly in coniferous forest ecosystems. Yet insect disturbances also affect successional pathways and landscape structure that interact with fire disturbances (and vice-versa) over longer time scales. We applied a landscape succession and disturbance model (LANDIS-II) to evaluate the relative strength of interactions between spruce budworm (Choristoneura fumiferana) outbreaks and fire disturbances in the Boundary Waters Canoe Area (BWCA) in northern Minnesota (USA). Disturbance interactions were evaluated for two different scenarios: presettlement forests and fire regimes vs. contemporary forests and fire regimes. Forest composition under the contemporary scenario trended toward mixtures of deciduous species (primarily Betula papyrifera and Populus spp.) and shade-tolerant conifers (Picea mariana, Abies balsamea, Thuja occidentalis), with disturbances dominated by a combination of budworm defoliation and high-severity fires. The presettlement scenario retained comparatively more “big pines” (i.e., Pinus strobus, P. resinosa) and tamarack (L. laricina), and experienced less budworm disturbance and a comparatively less-severe fire regime. Spruce budworm disturbance decreased area burned and fire severity under both scenarios when averaged across the entire 300-year simulations. Contrary to past research, area burned and fire severity during outbreak decades were each similar to that observed in non-outbreak decades. Our analyses suggest budworm disturbances within forests of the BWCA have a comparatively weak effect on long-term forest composition due to a combination of characteristics. These include strict host specificity, fine-scaled patchiness created by defoliation damage, and advance regeneration of its primary host, balsam fir (A. balsamea) that allows its host to persist despite repeated disturbances. Understanding the nature of the three-way interaction

  4. Non-destructive analysis and detection of internal characteristics of spruce logs through X computerized tomography

    International Nuclear Information System (INIS)

    Longuetaud, F.

    2005-10-01

    Computerized tomography allows a direct access to internal features of scanned logs on the basis of density and moisture content variations. The objective of this work is to assess the feasibility of an automatic detection of internal characteristics with the final aim of conducting scientific analyses. The database is constituted by CT images of 24 spruces obtained with a medical CT scanner. Studied trees are representative of several social status and are coming from four stands located in North-Eastern France, themselves are representative of several age, density and fertility classes. The automatic processing developed are the following. First, pith detection in logs dealing with the problem of knot presence and ring eccentricity. The accuracy of the localisation was less than one mm. Secondly, the detection of the sapwood/heart-wood limit in logs dealing with the problem of knot presence (main source of difficulty). The error on the diameter was 1.8 mm which corresponds to a relative error of 1.3 per cent. Thirdly, the detection of the whorls location and comparison with an optical method. Fourthly the detection of individualized knots. This process allows to count knots and to locate them in a log (longitudinal position and azimuth); however, the validation of the method and extraction of branch diameter and inclination are still to be developed. An application of this work was a variability analysis of the sapwood content in the trunk: at the within-tree level, the sapwood width was found to be constant under the living crown; at the between-tree level, a strong correlation was found with the amount of living branches. A great number of analyses are possible from our work results, among others: architectural analysis with the pith tracking and the apex death occurrence; analysis of radial variations of the heart-wood shape; analysis of the knot distribution in logs. (author)

  5. Inheritance of Budbreak and Correlation with Early Height Growth in White Spruce (Picea glauca) from New England

    Science.gov (United States)

    Ronald C. Wilkinson

    1977-01-01

    Variation in budbreak date among 37 half-sib families of white spruce in a replicated one-parent progeny test plantation in southern Maine was only 5 days. Differences in the mean date of budbreak between years were greater than those between families, but the genetic correlation between date of budbreak in different years was .661. Heritability estimates ranged from ....

  6. Calcium fertilization increases the concentration of calcium in sapwood and calcium oxalate in foliage of red spruce

    Science.gov (United States)

    Kevin T. Smith; Walter C. Shortle; Jon H. Connolly; Rakesh Minocha; Jody Jellison

    2009-01-01

    Calcium cycling plays a key role in the health and productivity of red spruce forests in the northeastern US. A portion of the flowpath of calcium within forests includes translocation as Ca2+ in sapwood and accumulation as crystals of calcium oxalate in foliage. Concentrations of Ca in these tree tissues have been used as markers of...

  7. Genotype-environment interaction and stability in ten-year height growth of Norway spruce Clones (Picea abies Karst.).

    Science.gov (United States)

    J.B. St. Clair; J. Kleinschmit

    1986-01-01

    Norway spruce cuttings of 40 clones were tested on seven contrasting sites in northern Germany. Analysis of variance for ten-year height growth indicate a highly significant clone x site interaction. This interaction may be reduced by selection of stable clones. Several measures of stability were calculated and discussed. Characterization of sites by the method of...

  8. High methane emissions from restored Norway spruce swamps in southern Finland over one growing season

    Directory of Open Access Journals (Sweden)

    M. Koskinen

    2016-02-01

    Full Text Available Forestry-drained peatlands in the boreal region are currently undergoing restoration in order to bring these ecosystems closer to their natural (undrained state. Drainage affects the methane (CH4 dynamics of a peatland, often changing sites from CH4 sources to sinks. Successful restoration of a peatland would include restoration of not only the surface vegetation and hydrology, but also the microbial populations and thus CH4 dynamics. As a pilot study, CH4 emissions were measured on two pristine, two drained and three restored boreal spruce swamps in southern Finland for one growing season. Restoration was successful in the sense that the water table level in the restored sites was significantly higher than in the drained sites, but it was also slightly higher than in the pristine sites. The restored sites were surprisingly large sources of CH4 (mean emissions of 52.84 mg CH4 m-2 d-1, contrasting with both the pristine (1.51 mg CH4 m-2 d-1 and the drained sites (2.09 mg CH4 m-2 d-1. More research is needed to assess whether the high CH4 emissions observed in this study are representative of restored spruce mires in general.

  9. Ecophysiology at SPRUCE: Impacts of whole ecosystem warming and elevated CO2 on leaf-level photosynthesis and respiration of two ericaceous shrubs in a boreal peatland

    Science.gov (United States)

    Ward, E. J.; Dusenge, M. E.; Warren, J.; Murphy, B. K.; Way, D.; King, A. W.; McLennan, D.; Montgomery, R.; Stefanski, A.; Reich, P. B.; Cruz Aguilar, M.; Wullschleger, S.; Bermudez Villanueva, R.; Hanson, P. J.

    2017-12-01

    The Spruce and Peatland Responses Under Changing Environments (SPRUCE) project is a large-scale, long-term experiment investigating the effects of warming and elevated CO2 on an ombrotrophic bog in Minnesota, USA. SPRUCE uses 10 large (12.8-m diameter) enclosures to increase air and soil temperatures to a range of targets (+0 °C, +2.25 °C, +4.5 °C, +6.75 °C, +9 °C) under both ambient and elevated (+500 ppm) CO2 concentrations. Whole-ecosystem-warming treatments began in August 2015 and elevated CO2 treatments began in June 2016. This talk will address the photosynthetic and respiratory responses of vascular plants to the treatments as measured with a variety of in-situ and ex-situ measurements conducted throughout the 2016 and 2017 growing seasons. We will focus on the responses of two dominant ericaceous shrubs (Rhododendron groenlandicum and Chamaedaphne calyculata), which account for more 80% of the understory biomass of this open-canopy forest. Such physiological changes are not only leading indicators of changes in plant growth and community structure, but are crucial to understanding carbon cycling of raised bogs and representing boreal peatlands in global dynamic vegetation models. Pre-treatment data collected at this site indicate that the physiologically active season typically begins in late May and extends into the fall until freezing nighttime temperatures are consistently reached, typically in October. Post-treatment measurements made during seasonal transitions indicate a longer active physiological season in warmer treatments. Results from 2016 measurements show some degree of thermal acclimation of photosynthesis in R. groenlandicum and of respiration in both species in the early growing season, but not late season. Late season measurements show a down-regulation of photosynthesis in both shrub species grown under elevated CO2. Taken as a whole, these results indicate complex interactions between phenological changes and treatment effects on

  10. Spatial pattern of structural ageing in eastern Croatia: evolution and explanations

    Directory of Open Access Journals (Sweden)

    Marijan Jukic

    2015-11-01

    Full Text Available This article aims to examine the ageing situation and social policy issues in the Osijek-Baranja County of eastern Croatia. Using historical evidence from census data, research suggests that the evolution of the ageing pattern has been mainly determined by such factors as development of the transport system, changes in political-territorial organisation, supply of jobs in the cities, deagrarianisation and a domestic war in the 1990s. The increased importance of urban centres, through planned industrialisation and administrative centralisation, has accelerated and intensified rural-tourban migration. Consequently, the spatial pattern of structural ageing has been substantially affected. A significant variation was found in urban and rural areas and also within sub-regional units. The findings suggest that the evolution of spatial disparities in the ageing pattern is because of unplanned migration; spatial differences in the level of socio-economic development; the influence of tradition, such as higher fertility rates historically in some areas; and suburbanisation, notably around the city of Osijek. The article concludes that ageing is affecting the country's economic growth and the formal and informal social support systems, including the provision of resources for older citizens in the endangered areas.

  11. Retrieval of spruce leaf chlorophyll content from airborne image data using continuum removal and radiative transfer

    Czech Academy of Sciences Publication Activity Database

    Malenovský, Z.; Homolová, L.; Zurita-Milla, R.; Lukeš, Petr; Kaplan, Věroslav; Hanuš, Jan; Gastellu-Etchegory, J.P.; Schaepman, M.E.

    2013-01-01

    Roč. 131, APR (2013), s. 85-102 ISSN 0034-4257 R&D Projects: GA MŠk(CZ) ED1.1.00/02.0073; GA MŠk(CZ) LM2010007 Institutional support: RVO:67179843 Keywords : Chlorophyll retrieval * Imaging spectroscopy * Continuum removal * Radiative transfer * PROSPECT * DART * Optical indices * Norway spruce * High spatial resolution * AISA Subject RIV: EH - Ecology, Behaviour Impact factor: 4.769, year: 2013

  12. Effects of sewage sludge addition to Norway spruce seedlings on nitrogen availability and soil fauna in clear-cut areas.

    Science.gov (United States)

    Nieminen, Jouni K; Räisänen, Mikko

    2013-07-01

    Anaerobically digested and composted sewage sludge (CSS) has been suggested to be a slow-release fertilizer in forestry and an alternative to quick-release inorganic fertilizers. The effects of CSS with or without added carbohydrate on inorganic nitrogen availability and on soil animals were tested in two Norway spruce plantations. Half of the seedlings were individually fertilized with CSS, and the rest were left as controls. Solid sucrose was added to half of the fertilized and untreated seedlings. Soil samples were taken in the autumn in the first and the second year after the treatments. CSS increased soil NH4-N (2100%), the proportion of soil NO3-N, and the N concentration of spruce needles. CSS greatly reduced the abundances of enchytraeids, tardigrades and collembolans, but increased the proportion and abundance of bacterial-feeding nematodes irrespective of carbohydrate addition. A better stabilization method needs to be developed before CSS can be used as a forest fertilizer. Copyright © 2013 Elsevier Ltd. All rights reserved.

  13. Recent climatic drying leads to age-independent growth reductions of white spruce stands in western Canada.

    Science.gov (United States)

    Hogg, Edward H; Michaelian, Michael; Hook, Trisha I; Undershultz, Michael E

    2017-12-01

    Since 2001, climatic conditions have been notably drier than normal across large areas of the western Canadian interior, leading to widespread impacts on the forests of this region. This poses a major concern for the future, given climate change projections for continued warming and drying. We conducted tree-ring analysis in 75 pure stands of white spruce (Picea glauca) across Alberta and west-central Saskatchewan to examine the effects of recent climatic drying on the growth of this important boreal tree species. Allometric equations were used to calculate annual growth in aboveground tree biomass (G BM ) from ring width measurements. Results showed an increasing trend in G BM from the 1960s to the 1990s, followed by a sharp decline during the severe drought of 2001-2002. Of the 75 stands, only 18 recovered sufficiently to cause an increase in mean G BM from the predrought decade of 1991-2000 to the subsequent decade of 2001-2010. The remaining 57 stands exhibited a decline in mean G BM between these decades. Climatic drying was a major cause of the growth decline, as shown by the significant stand-level relationship between percentage change in decadal mean G BM and the change in decadal mean values of a climate moisture index from 1991-2000 to 2001-2010. Subsequent analyses of boreal stands sampled across Alberta during 2015 revealed that white spruce growth had declined even further as drought conditions intensified during 2014-2015. Overall, there was a 38% decrease in mean G BM between 1997 and 2015, but surprisingly, the percentage decrease was not significantly different for young, productive stands compared with older, less productive stands. Thus, stand ageing cannot explain the observed decline in white spruce growth during the past quarter century, suggesting that these forests are at risk if the trend towards more frequent, severe drought continues in the region. © 2017 Her Majesty the Queen in Right of Canada Global Change Biology ©2017 John Wiley

  14. The wind and fire disturbance in Central European mountain spruce forests: the regeneration after four years

    Directory of Open Access Journals (Sweden)

    Monika Budzáková

    2013-03-01

    Full Text Available A strong windstorm in November 2004 resulted in a huge blown-down spruce forest area in the southern part of the Tatra National Park in the Western Carpathians in Slovakia, Central Europe. The aim of this work is to study the vegetation composition of spruce forest at differently managed sites four years after this disturbance. Four study areas were selected for this purpose: (i an area where the fallen trees were extracted and new seedlings were planted; (ii an area, which was hit by a forest fire after the extraction; (iii an area where no active management was applied; (iv a reference forest unaffected by such disturbance. A total of 100 plots were selected, 25 of each area type. The result of DCA and CCA analyses consistently indicated that after this short period the non-extracted and extracted areas are currently most similar to the reference forest area, while the fire affected area differed. A one-way ANOVA comparing species cover for the different plot sizes indicated some significant differences between the extracted and non-extracted plots. The abundance of certain species commonly occurring in spruce forests, such as Dyopteris carthusiana agg., Vaccinium myrtillus and Avenella flexuosa, correlated weli with the non-extracted plots, compared to the extracted plots. Coverage of these species was lowest on burned plots. The lowest Shannon-Wiener’s diversity values were recorded in burned plots. This was most likely a consequence of mono-dominant competitive species spread, (mainly Chamerion angustifolium which profited from the altered ecological conditions following the fire. Although some differences were also registered in the Shannon-Wiener diversity index between the remaining research plots, however these were not statistically significant. The most important results of our investigations include the extensive influence of fire disturbance on vegetation. Study revealed that the wind-disturbed area is able to regenerate

  15. Life?cycle impacts of ethanol production from spruce wood chips under high-gravity conditions

    OpenAIRE

    Janssen, Matty; Xiros, Charilaos; Tillman, Anne-Marie

    2016-01-01

    Background Development of more sustainable biofuel production processes is ongoing, and technology to run these processes at a high dry matter content, also called high-gravity conditions, is one option. This paper presents the results of a life?cycle assessment (LCA) of such a technology currently in development for the production of bio-ethanol from spruce wood chips. Results The cradle-to-gate LCA used lab results from a set of 30 experiments (or process configurations) in which the main p...

  16. The influence of climate change on stomatal ozone flux to a mountain Norway spruce forest

    Czech Academy of Sciences Publication Activity Database

    Zapletal, M.; Pretel, J.; Chroust, P.; Cudlín, Pavel; Edwards-Jonášová, Magda; Urban, Otmar; Pokorný, Radek; Czerný, Radek; Hůnová, I.

    2012-01-01

    Roč. 169, OCT 2012 (2012), s. 267-273 ISSN 0269-7491 R&D Projects: GA MŠk(CZ) ED1.1.00/02.0073; GA MŠk OC10022; GA MŠk(CZ) LM2010007 Institutional research plan: CEZ:AV0Z60870520 Keywords : Stomatal ozone flux * AOT40 * Phytotoxic Ozone Dose * Norway spruce * Net ecosystem production * Ozone * Climate change Subject RIV: EH - Ecology, Behaviour Impact factor: 3.730, year: 2012

  17. Hydraulic and mechanical properties of young Norway spruce clones related to growth and wood structure

    Science.gov (United States)

    ROSNER, SABINE; KLEIN, ANDREA; MÜLLER, ULRICH; KARLSSON, BO

    2011-01-01

    Summary Stem segments of eight five-year-old Norway spruce (Picea abies (L.) Karst.) clones differing in growth characteristics were tested for maximum specific hydraulic conductivity (ks100), vulnerability to cavitation and behavior under mechanical stress. The vulnerability of the clones to cavitation was assessed by measuring the applied air pressure required to cause 12 and 50% loss of conductivity (Ψ12, Ψ50) and the percent loss of conductivity at 4 MPa applied air pressure (PLC4MPa). The bending strength and stiffness and the axial compression strength and stiffness of the same stem segments were measured to characterize wood mechanical properties. Growth ring width, wood density, latewood percentage, lumen diameter, cell wall thickness, tracheid length and pit dimensions of earlywood cells, spiral grain and microfibril angles were examined to identify structure–function relationships. High ks100 was strongly and positively related to spiral grain angle, which corresponded positively to tracheid length and pit dimensions. Spiral grain may reduce flow resistance of the bordered pits of the first earlywood tracheids, which are characterized by rounded tips and an equal distribution of pits along the entire length. Wood density was unrelated to hydraulic vulnerability parameters. Traits associated with higher hydraulic vulnerability were long tracheids, high latewood percentage and thick earlywood cell walls. The positive relationship between earlywood cell wall thickness and vulnerability to cavitation suggest that air seeding through the margo of bordered pits may occur in earlywood. There was a positive phenotypic and genotypic relationship between ks100 and PLC4MPa, and both parameters were positively related to tree growth rate. Variability in mechanical properties depended mostly on wood density, but also on the amount of compression wood. Accordingly, hydraulic conductivity and mechanical strength or stiffness showed no tradeoff. PMID:17472942

  18. Structural Evolution and Mobile Shale Deformation in the Eastern Niger Delta

    International Nuclear Information System (INIS)

    Wiener, R. W.; Aikhionbare, D. O. L.

    2002-01-01

    Regional cross-sections and restorations of the eastern delta constructed from 2D and 3D seismic data show the structural evolution of paired extensional contractional belts and the kinematic and geometric evolution of mobile shale. The delta consists of an updip extensional belt and downdip zones of transitional and contractional deformation linked by a regional detachment. The extensional belt is characterized by zones of N-dipping (counterregional) and S-dipping (regional) normal faults.In the regional fault trend, sediment accommodation space is created largely by lateral movement of mobile substrate due to sediment loading and gravity. The transitional belt is characterized by low relief, shale-cored detachment folds and normal faults. The contractional belt consists of 2 parts, the high relief shale-cored detachment fold belt (mobile shale) and the fold/thrust belt: In the mobile shale belt, anticlines are generally symmetric and characterized by parallel-folded cover and highly variable thickness in the underlying ductile shale zone.Palinspastic restoration of the mobile shale by area balance shows a high degree of lateral and vertical mobility. Isostatic restoration of the depositional wedge that is the precursor to the mobile shale suggests lateral movement of 10s of kms from the extensional to the contractional domain. The fold and thrust belt is characterized by a train of asymmetric fault-related folds. The zone of ductile substrate is thin in this area, which may account for the change in structural style from high relief detachment folds in the mobile shale belt to a more classic fold/thrust belt style to the south

  19. Similarity of nutrient uptake and root dimensions of Engelmann spruce and subalpine fir at two contrasting sites in Colorado

    Energy Technology Data Exchange (ETDEWEB)

    Yanai, R; McFarlane, K; Lucash, M; Kulpa, S; Wood, D

    2009-10-09

    Nutrient uptake capacity is an important parameter in modeling nutrient uptake by plants. Researchers commonly assume that uptake capacity measured for a species can be used across sites. We tested this assumption by measuring the nutrient uptake capacity of intact roots of Engelmann spruce (Picea engelmanni Parry) and subalpine fir (Abies lasiocarpa (Hook.) Nutt.) at Loch Vale Watershed and Fraser Experimental Forest in the Rocky Mountains of central Colorado. Roots still attached to the tree were exposed to one of three concentrations of nutrient solutions for time periods ranging from 1 to 96 hours, and solutions were analyzed for ammonium, nitrate, calcium, magnesium, and potassium. Surprisingly, the two species were indistinguishable in nutrient uptake within site for all nutrients (P > 0.25), but uptake rates differed by site. In general, nutrient uptake was higher at Fraser (P = 0.01, 0.15, 0.03, 0.18 for NH{sub 4}{sup +}, NO{sub 3}{sup -}, Ca{sup 2+}, and K{sup +}, respectively), which is west of the Continental Divide and has lower atmospheric deposition of N than Loch Vale. Mean uptake rates by site for ambient solution concentrations were 0.12 {micro}mol NH{sub 4}{sup +} g{sub fwt}{sup -1} h{sup -1}, 0.02 {micro}mol NO{sub 3}{sup -} g{sub fwt}{sup -1}, 0.21 {micro}mol Ca{sup 2+} g{sub fwt}{sup -1} h{sup -1}, and 0.01 {micro}mol Mg{sup 2+} g{sub fwt}{sup -1} h{sup -1} at Loch Vale, and 0.21 {micro}mol NH{sub 4}{sup +} f{sub fwt}{sup -1}h{sup -1}, 0.04 {micro}mol NO{sub 3}{sup -} g{sub fwt}{sup -1} h{sup -1}, 0.51 {micro}mol Ca{sup 2+}g{sub fwt}{sup -1}h{sup -1}, and 0.07 {micro}mol Mg{sup 2+} f{sub fwt}{sup -1}h{sup -1} at Fraser. The importance of site conditions in determining uptake capacity should not be overlooked when parameterizing nutrient uptake models. We also characterized the root morphology of these two species and compared them to other tree species we have measured at various sites in the northeastern USA. Engelman spruce and subalpine fir

  20. Tectolinear interpretation of a 1:5,000,000 Landsat-1 mosaic compared with the structure of central and eastern United States

    Science.gov (United States)

    Kutina, Jan; Carter, William D.

    1978-01-01

    The pattern of lineaments and curvilinear features interpreted from a 1:5,000,000 mosaic of satellite images (Landsat-1 was superimposed on a simplified version of the Geological Map of the United States, 1:2,500,000 scale, showing the structural scheme of Central and Eastern United States. A comparison of the above two patterns, shown in Fig. 1, is presented in this paper.

  1. Structural patterns of the Lake Erçek Basin, eastern Anatolia (Turkey): evidence from single-channel seismic interpretation

    Science.gov (United States)

    Toker, Mustafa; Tur, Hüseyin

    2017-11-01

    This study presents an analysis of the single-channel high-resolution shallow seismic reflection data from Lake Erçek, eastern Anatolia, to provide key information on the deformational elements, on the fault patterns and on the overall tectonic structure of the Lake Erçek Basin. High-resolution seismic data reveal major structural and deformational features, including N-S trending normal faults and W-E trending reverse faults bounding the Lake Erçek Basin, basement highs and folded structures along the marginal sections of the lake. The N-S trending normal faults asymmetrically control the steep western margin and the gentle eastern deltaic section, while the W-E trending reverse faults appear at the northern and southern margins. The N-S trending normal faults, half-graben structure, and the gradual thickening of sediments in the Erçek Basin toward the fault scarps strongly suggest an extensional tectonic regime resulting from an N-S compression. The Erçek Basin is an extension-controlled depocenter; it is a relatively undeformed and flat-lying deep Basin, forming a typical example of the half-graben structure. The N-S trending normal faults appear to be currently active and control the lake center and the E-delta section, resulting in subsidence in the lake floor. In the N- and S-margins of the lake, there is evidence of folding, faulting and accompanying block uplifting, suggesting a significant N-S compressional regime that results in the reverse faulting and basement highs along the marginal sections. The folding and faulting caused strong uplift of the basement blocks in the N- and S- margins, subsequently exposing the shelf and slope areas. The exposed areas are evident in the erosional unconformity of the surface of the basement highs and thinned sediments. The tilted basement strata and subsequent erosion over the basement block highs suggest prominent structural inversion, probably long before the formation of the lake. New high-resolution seismic

  2. Sources of Respired Carbon in a Northern Minnesota Ombrotrophic Spruce Bog: Preliminary 14C Results from the SPRUCE Site.

    Science.gov (United States)

    Guilderson, T. P.; McNicol, G.; Machin, A.; Hanson, P. J.; McFarlane, K. J.; Osuna, J. L.; Pett-Ridge, J.; Singleton, M. J.

    2014-12-01

    A significant uncertainty in future land-surface carbon budgets is the response of wetlands to climate change. A corollary and related question is the future net climate (radiative) forcing impact from wetlands. Active wetlands emit both CO2 and CH4 to the atmosphere. CH4 is, over a few decades, a much more potent greenhouse gas than CO2. CO2 has a longer atmospheric lifetime and a longer 'tail' to its radiative influence. Whether wetlands are a net source or sink of atmospheric carbon under future climate change will depend on ecosystem response to rising temperatures and elevated CO2. The largest uncertainty in future wetland C-budgets, and their climate forcing is the stability of the large below-ground carbon stocks, often in the form of peat, and the partitioning of CO2 and CH4 released via ecosystem respiration. In advance of a long-term experimental warming and elevated CO2 manipulation at the DOE Spruce and Peatland Responses Under Climatic and Environmental Change (SPRUCE) site in the Marcell Experimental Forest, we have characterized the source of respired carbon used for both the production of CO2 and CH4. Samples were collected in early June, late July, and will be collected in early September from three large (~1.1 m2, ~0.5m3) chambers from the control plot, and two of the experimental plots selected for heating (+9°C, +4.5°C). Early June fluxes from the three chambers were ~5500 mgC-m-2-d-1 and ~16 mgC-m-2-d-1 for CO2 and CH4 respectively. Radiocarbon analysis of CO2 and CH4 indicate that the source for the respired carbon is for the most part recent, with most 14C values between 30 and 40‰ - i.e., carbon that was photosynthetically fixed in the last few years. In concert with rising air and ground temperatures fluxes in late July increased to ~6500 mgC-m-2-d-1 and ~86 mgC-m-2-d-1. Although deep-heating was initiated in mid to late June we hypothesize that the July respiration signal is dominated by the regular seasonal cycle of natural warming

  3. Carbon pools in a montane old-growth Norway spruce ecosystem in Bohemian Forest: Effects of stand age and elevation

    Czech Academy of Sciences Publication Activity Database

    Seedre, M.; Kopáček, Jiří; Janda, P.; Bače, R.; Svoboda, M.

    2015-01-01

    Roč. 346, June (2015), s. 106-113 ISSN 0378-1127 R&D Projects: GA ČR(CZ) GAP504/12/1218 Institutional support: RVO:60077344 Keywords : carbon dynamics * soil carbon * spruce biomass C * dead root C * unmanaged ecosystem Subject RIV: GK - Forestry Impact factor: 2.826, year: 2015

  4. Investigation on the Scratch Strength of Clear Paints Used in Furniture Industries on the Wood Species Beech, Elm, Alder and Spruce

    Directory of Open Access Journals (Sweden)

    Mohammad Ghofrani

    2014-05-01

    Full Text Available In This research, the scratch strength (Cross-Cut Test of clear paints (nitrocellulose lacquers and acid catalyst lacquers of wood species Beech, Elm, Alder and spruce were studied as a function of moisture content (MC of the samples. For this purpose, lumbers (550×110×12 mm were cut from sapwood in tangential surfaces and were air dried for one month according to wood drying procedures. Then, for pre-conditioning of moisture content, at the levels of 8%, 12% and 15%, the samples were placed in three clima rooms. Then, finish applied and strength tests were performed. The results revealed that for acid catalyst lacquers the highest scratch strength (10.4% belongs to Elm wood with 8% moisture content, and the lowest scratch strength (53.6% belongs to Spruce wood with nitrocellulose lacquers having 15% moisture content.

  5. Fine root dynamics in lodgepole pine and white spruce stands along productivity gradients in reclaimed oil sands sites.

    Science.gov (United States)

    Jamro, Ghulam Murtaza; Chang, Scott X; Naeth, M Anne; Duan, Min; House, Jason

    2015-10-01

    Open-pit mining activities in the oil sands region of Alberta, Canada, create disturbed lands that, by law, must be reclaimed to a land capability equivalent to that existed before the disturbance. Re-establishment of forest cover will be affected by the production and turnover rate of fine roots. However, the relationship between fine root dynamics and tree growth has not been studied in reclaimed oil sands sites. Fine root properties (root length density, mean surface area, total root biomass, and rates of root production, turnover, and decomposition) were assessed from May to October 2011 and 2012 using sequential coring and ingrowth core methods in lodgepole pine (Pinus contorta Dougl.) and white spruce (Picea glauca (Moench.) Voss) stands. The pine and spruce stands were planted on peat mineral soil mix placed over tailings sand and overburden substrates, respectively, in reclaimed oil sands sites in Alberta. We selected stands that form a productivity gradient (low, medium, and high productivities) of each tree species based on differences in tree height and diameter at breast height (DBH) increments. In lodgepole pine stands, fine root length density and fine root production, and turnover rates were in the order of high > medium > low productivity sites and were positively correlated with tree height and DBH and negatively correlated with soil salinity (P < 0.05). In white spruce stands, fine root surface area was the only parameter that increased along the productivity gradient and was negatively correlated with soil compaction. In conclusion, fine root dynamics along the stand productivity gradients were closely linked to stand productivity and were affected by limiting soil properties related to the specific substrate used for reconstructing the reclaimed soil. Understanding the impact of soil properties on fine root dynamics and overall stand productivity will help improve land reclamation outcomes.

  6. Ancient trade routes shaped the genetic structure of horses in eastern Eurasia.

    Science.gov (United States)

    Warmuth, Vera M; Campana, Michael G; Eriksson, Anders; Bower, Mim; Barker, Graeme; Manica, Andrea

    2013-11-01

    Animal exchange networks have been shown to play an important role in determining gene flow among domestic animal populations. The Silk Road is one of the oldest continuous exchange networks in human history, yet its effectiveness in facilitating animal exchange across large geographical distances and topographically challenging landscapes has never been explicitly studied. Horses are known to have been traded along the Silk Roads; however, extensive movement of horses in connection with other human activities may have obscured the genetic signature of the Silk Roads. To investigate the role of the Silk Roads in shaping the genetic structure of horses in eastern Eurasia, we analysed microsatellite genotyping data from 455 village horses sampled from 17 locations. Using least-cost path methods, we compared the performance of models containing the Silk Roads as corridors for gene flow with models containing single landscape features. We also determined whether the recent isolation of former Soviet Union countries from the rest of Eurasia has affected the genetic structure of our samples. The overall level of genetic differentiation was low, consistent with historically high levels of gene flow across the study region. The spatial genetic structure was characterized by a significant, albeit weak, pattern of isolation by distance across the continent with no evidence for the presence of distinct genetic clusters. Incorporating landscape features considerably improved the fit of the data; however, when we controlled for geographical distance, only the correlation between genetic differentiation and the Silk Roads remained significant, supporting the effectiveness of this ancient trade network in facilitating gene flow across large geographical distances in a topographically complex landscape. © 2013 John Wiley & Sons Ltd.

  7. PEI detoxification of pretreated spruce for high solids ethanol fermentation

    International Nuclear Information System (INIS)

    Cannella, David; Sveding, Per Viktor; Jørgensen, Henning

    2014-01-01

    Highlights: • High solids (30% dry matter) pretreatment, enzymatic hydrolysis and fermentation. • Horizontal rotary reactor for hydrolysis and fermentation. • In situ hydrolysates detoxification using inhibitors adsorbing PEI polymer. • 50% of inhibitors recovered as by-product, recyclability of PEI polymer up to 5 times. • 76% of maximum theoretical ethanol was fermented at final concentration of 51 g/kg. - Abstract: Performing the bioethanol production process at high solids loading is a requirement for economic feasibility at industrial scale. So far this has successfully been achieved using wheat straw and other agricultural residues at 30% of water insoluble solids (WIS), but for softwood species (i.e. spruce) this has been difficult to reach. The main reason behind this difference is the higher recalcitrance of woody substrates which require harsher pretreatment conditions, thus generating higher amounts of inhibitory compounds, ultimately lowering fermentation performances. In this work we studied ethanol production from spruce performing the whole process, from pretreatment to hydrolysis and fermentation, at 30% dry matter (equivalent to ∼20% WIS). Hydrolysis and fermentation was performed in a horizontal free fall mixing reactor enabling efficient mixing at high solids loadings. In batch simultaneous saccharification and fermentation (SSF), up to 76% cellulose to ethanol conversion was achieved resulting in a concentration of 51 g/kg of ethanol. Key to obtaining this high ethanol yield at these conditions was the use of a detoxification technology based on applying a soluble polyelectrolyte polymer (polyethylenimine, PEI) to absorb inhibitory compounds in the material. On average 50% removal and recovery of the main inhibitors (HMF, furfural, acetic acid and formic acid) was achieved dosing 1.5% w/w of soluble PEI. The use of PEI was compatible with operating the process at high solids loadings and enabled fermentation of hydrolysates, which

  8. Modular organization of the white spruce (Picea glauca) transcriptome reveals functional organization and evolutionary signatures.

    Science.gov (United States)

    Raherison, Elie S M; Giguère, Isabelle; Caron, Sébastien; Lamara, Mebarek; MacKay, John J

    2015-07-01

    Transcript profiling has shown the molecular bases of several biological processes in plants but few studies have developed an understanding of overall transcriptome variation. We investigated transcriptome structure in white spruce (Picea glauca), aiming to delineate its modular organization and associated functional and evolutionary attributes. Microarray analyses were used to: identify and functionally characterize groups of co-expressed genes; investigate expressional and functional diversity of vascular tissue preferential genes which were conserved among Picea species, and identify expression networks underlying wood formation. We classified 22 857 genes as variable (79%; 22 coexpression groups) or invariant (21%) by profiling across several vegetative tissues. Modular organization and complex transcriptome restructuring among vascular tissue preferential genes was revealed by their assignment to coexpression groups with partially overlapping profiles and partially distinct functions. Integrated analyses of tissue-based and temporally variable profiles identified secondary xylem gene networks, showed their remodelling over a growing season and identified PgNAC-7 (no apical meristerm (NAM), Arabidopsis transcription activation factor (ATAF) and cup-shaped cotyledon (CUC) transcription factor 007 in Picea glauca) as a major hub gene specific to earlywood formation. Reference profiling identified comprehensive, statistically robust coexpressed groups, revealing that modular organization underpins the evolutionary conservation of the transcriptome structure. © 2015 The Authors. New Phytologist © 2015 New Phytologist Trust.

  9. Identification and Functional Characterization of Monofunctional ent-Copalyl Diphosphate and ent-Kaurene Synthases in White Spruce Reveal Different Patterns for Diterpene Synthase Evolution for Primary and Secondary Metabolism in Gymnosperms1[W][OA

    Science.gov (United States)

    Keeling, Christopher I.; Dullat, Harpreet K.; Yuen, Mack; Ralph, Steven G.; Jancsik, Sharon; Bohlmann, Jörg

    2010-01-01

    The biosynthesis of the tetracyclic diterpene ent-kaurene is a critical step in the general (primary) metabolism of gibberellin hormones. ent-Kaurene is formed by a two-step cyclization of geranylgeranyl diphosphate via the intermediate ent-copalyl diphosphate. In a lower land plant, the moss Physcomitrella patens, a single bifunctional diterpene synthase (diTPS) catalyzes both steps. In contrast, in angiosperms, the two consecutive cyclizations are catalyzed by two distinct monofunctional enzymes, ent-copalyl diphosphate synthase (CPS) and ent-kaurene synthase (KS). The enzyme, or enzymes, responsible for ent-kaurene biosynthesis in gymnosperms has been elusive. However, several bifunctional diTPS of specialized (secondary) metabolism have previously been characterized in gymnosperms, and all known diTPSs for resin acid biosynthesis in conifers are bifunctional. To further understand the evolution of ent-kaurene biosynthesis as well as the evolution of general and specialized diterpenoid metabolisms in gymnosperms, we set out to determine whether conifers use a single bifunctional diTPS or two monofunctional diTPSs in the ent-kaurene pathway. Using a combination of expressed sequence tag, full-length cDNA, genomic DNA, and targeted bacterial artificial chromosome sequencing, we identified two candidate CPS and KS genes from white spruce (Picea glauca) and their orthologs in Sitka spruce (Picea sitchensis). Functional characterization of the recombinant enzymes established that ent-kaurene biosynthesis in white spruce is catalyzed by two monofunctional diTPSs, PgCPS and PgKS. Comparative analysis of gene structures and enzyme functions highlights the molecular evolution of these diTPSs as conserved between gymnosperms and angiosperms. In contrast, diTPSs for specialized metabolism have evolved differently in angiosperms and gymnosperms. PMID:20044448

  10. Vegetation and invertebrate community response to eastern hemlock decline in southern new England

    Science.gov (United States)

    Laura L. Ingwell; Mailea Miller-Pierce; R. Talbot Trotter; Evan L. Preisser

    2012-01-01

    The introduction of Adelges tsugae (Hemlock Woolly Adelgid [HWA]) to the eastern United States has had a devastating impact on Tsuga canadensis (Eastern Hemlock). Although much research has been done to assess HWA impacts on ecosystem processes and vegetation structure, few researchers have examined community-level changes in...

  11. Leaf physiological responses of mature Norway Spruce trees exposed to elevated carbon dioxide and temperature

    Science.gov (United States)

    Lamba, Shubhangi; Uddling, Johan; Räntfors, Mats; Hall, Marianne; Wallin, Göran

    2014-05-01

    Leaf photosynthesis, respiration and stomatal conductance exert strong control over the exchange of carbon, water and energy between the terrestrial biosphere and the atmosphere. As such, leaf physiological responses to rising atmospheric CO2 concentration ([CO2]) and temperature have important implications for the global carbon cycle and rate of ongoing global warming, as well as for local and regional hydrology and evaporative cooling. It is therefore critical to improve the understanding of plant physiological responses to elevated [CO2] and temperature, in particular for boreal and tropical ecosystems. In order to do so, we examined physiological responses of mature boreal Norway spruce trees (ca 40-years old) exposed to elevated [CO2] and temperature inside whole-tree chambers at Flakaliden research site, Northern Sweden. The trees were exposed to a factorial combination of two levels of [CO2] (ambient and doubled) and temperature (ambient and +2.8 degree C in summer and +5.6 degree C in winter). Three replicates in each of the four treatments were used. It was found that photosynthesis was increased considerably in elevated [CO2], but was not affected by the warming treatment. The maximum rate of photosynthetic carboxylation was reduced in the combined elevated [CO2] and elevated temperature treatment, but not in single factor treatments. Elevated [CO2] also strongly increased the base rate of respiration and to a lesser extent reduced the temperature sensitivity (Q10 value) of respiration; responses which may be important for the carbon balance of these trees which have a large proportion of shaded foliage. Stomatal conductance at a given VPD was reduced by elevated temperature treatment, to a degree that mostly offset the higher vapour pressure deficit in warmed air with respect to transpiration. Elevated [CO2] did not affect stomatal conductance, and thus increased the ratio of leaf internal to external [CO2]. These results indicate that the large elevated

  12. European spruce bark beetle (Ips typographus, L.) green attack affects foliar reflectance and biochemical properties

    Science.gov (United States)

    Abdullah, Haidi; Darvishzadeh, Roshanak; Skidmore, Andrew K.; Groen, Thomas A.; Heurich, Marco

    2018-02-01

    The European spruce bark beetle Ips typographus, L. (hereafter bark beetle), causes major economic loss to the forest industry in Europe, especially in Norway Spruce (Picea abies). To minimise economic loss and preclude a mass outbreak, early detection of bark beetle infestation (so-called ;green attack; stage - a period at which trees are yet to show visual signs of infestation stress) is, therefore, a crucial step in the management of Norway spruce stands. It is expected that a bark beetle infestation at the green attack stage affects a tree's physiological and chemical status. However, the concurrent effect on key foliar biochemical such as foliar nitrogen and chlorophyll as well as spectral responses are not well documented in the literature. Therefore, in this study, the early detection of bark beetle green attacks is investigated by examining foliar biochemical and spectral properties (400-2000 nm). We also assessed whether bark beetle infestation affects the estimation accuracy of foliar biochemicals. An extensive field survey was conducted in the Bavarian Forest National Park (BFNP), Germany, in the early summer of 2015 to collect leaf samples from 120 healthy and green attacked trees. The spectra of the leaf samples were measured using an ASD FieldSpec3 equipped with an integrating sphere. Significant differences (p < 0.05) between healthy and infested needle samples were found in the mean reflectance spectra, with the most pronounced differences being observed in the NIR and SWIR regions between 730 and 1370 nm. Furthermore, significant differences (p < 0.05) were found in the biochemical compositions (chlorophyll and nitrogen concentration) of healthy versus green attacked samples. Our results further demonstrate that the estimation accuracy of foliar chlorophyll and nitrogen concentrations, utilising partial least square regression model, was lower for the infested compared to the healthy trees. We show that early stage of infestation reduces not only

  13. Interactions between near-ground temperature and radiation, silvicultural treatments and frost damage to Norway spruce seedlings

    OpenAIRE

    Langvall, Ola

    2000-01-01

    Several different silvicultural treatments were studied in two experiments. In the first, mechanical scarification, slash removal, vegetation control, clear-cut age and seedling types were investigated with respect to frost injury to Norway spruce (Picea abies (L.) Karst.) seedlings. Frost damage was also related to near-ground minimum temperature. In the other experiment, the effects of Scots pine (Pinus sylvestris (L.)) shelterwood density gradients, ranging from dense, uncut forest to comp...

  14. Risk evaluation of the climatic change impact on secondary Norway spruce stands as exemplified by the Křtiny Training Forest Enterprise

    Czech Academy of Sciences Publication Activity Database

    Čermák, P.; Jankovský, L.; Cudlín, Pavel

    2004-01-01

    Roč. 50, č. 6 (2004), s. 256-262 ISSN 1212-4834 R&D Projects: GA MŠk OC E27.001 Institutional research plan: CEZ:AV0Z6087904 Keywords : climatic change * Norway spruce * risk assessment Subject RIV: GK - Forestry

  15. Eastern European Transformation and Youth Attitudes toward Violence

    Directory of Open Access Journals (Sweden)

    Berit Haußmann

    2011-12-01

    Full Text Available

    This cross-national examination of the motives behind adolescent approval of violence in major cities in Germany and four Eastern European transformation societies (Czech Republic, Poland, Russia, Slovenia draws on Institutional Anomie Theory (IAT, which leads us to expect higher instrumental motivation for violence among adolescents in the transforming societies. Differences in institutional structure and cultural orientations between Germany and the Eastern European societies are assessed using data from ILO and ESS. Analysis of the different motives for violence is based on data collected by the Criminological Research Institute of Lower Saxony (KFN in Kraków, Volgograd, Ljubljana, Plzen, and Hamburg. Comparability and cross-cultural interpretability of the violence
    attitude measure is assessed by applying confirmatory factor analyses in multi-group comparisons. A lack of sufficient data meant that specific assumed linkages as implied by IAT could not be established, but the results for a specific instrumental/utilitarian motive for violence and for institutional structure and cultural orientations point to the utility of applying insights from IAT to understanding the dynamics of violence within the Eastern European context of transformation.
    We find empirical indications that specific features of the family mediate the “Eastern effect” on the instrumental/utilitarian motive. Further research is needed to discover whether economic dominance affects motives for youth violence indirectly via the socialization provided by non-economic institutions rather than directly via cultural orientations.

  16. Growth and Yield of 15-Year Plantations of Pine, Spruce and Birch in Agricultural Land

    Directory of Open Access Journals (Sweden)

    Daugaviete Mudrite

    2017-07-01

    Full Text Available The growth data and the potential returns from 15-year-old plantations of pine Pinus sylvestris L. (6 trial sites, spruce Picea abies Karst L. (9 trial sites and silver birch Betula pendula Roth (13 trial sites, established in abandoned agricultural lands in a variety of soil types (sod calcareous, anthrosols, podzolic, podzols, gley, podzolic gley, alluvial, using the planting density 2,500 and 3,300 and also 5,000 trees/ha are analysed.

  17. Norway spruce fine root dynamics and carbon input into soil in relation to environmental factors

    OpenAIRE

    Leppälammi-Kujansuu, Jaana

    2014-01-01

    Knowledge of the quantity of belowground litter carbon (C) input is scarce but highly valued in C budget calculations. Specifically, the turnover rate of fine roots is considered to be one of the most important parameters in the estimation of changes in soil C stock. In this thesis Norway spruce (Picea abies L. (Karst.)) fine root lifespan and litter production and their responses to nutrient availability and temperature were examined. Aboveground foliage and understory litter C inputs were a...

  18. Multi-instrumental Investigation of Affecting of Early Somatic Embryos of Spruce by Cadmium(II) and Lead(II) Ions

    Czech Academy of Sciences Publication Activity Database

    Šupálková, V.; Petřek, J.; Baloun, J.; Adam, V.; Bartušek, Karel; Trnková, L.; Beklová, M.; Diopan, V.; Havel, L.; Kizek, R.

    2007-01-01

    Roč. 7, 5 (2007), s. 743-759 ISSN 1424-8220 R&D Projects: GA ČR(CZ) GA102/07/0389 Institutional research plan: CEZ:AV0Z20650511 Keywords : early somatic embryos of spruces * glutathione * heavy metals * plant cells * esterase activity * fluorescence detection Subject RIV: JA - Electronics ; Optoelectronics, Electrical Engineering Impact factor: 1.573, year: 2007

  19. Retrospective determination of 137Cs specific activity distribution in spruce bark and bark aggregated transfer factor in forests on the scale of the Czech Republic ten years after the Chernobyl accident.

    Science.gov (United States)

    Suchara, I; Rulík, P; Hůlka, J; Pilátová, H

    2011-04-15

    The (137)Cs specific activities (mean 32Bq kg(-1)) were determined in spruce bark samples that had been collected at 192 sampling plots throughout the Czech Republic in 1995, and were related to the sampling year. The (137)Cs specific activities in spruce bark correlated significantly with the (137)Cs depositions in areas affected by different precipitation sums operating at the time of the Chernobyl fallout in 1986. The ratio of the (137)Cs specific activities in bark and of the (137)Cs deposition levels yielded bark aggregated transfer factor T(ag) about 10.5×10(-3)m(-2)kg(-1). Taking into account the residual specific activities of (137)Cs in bark 20Bq kg(-1) and the available pre-Chernobyl data on the (137)Cs deposition loads on the soil surface in the Czech Republic, the real aggregated transfer factor after and before the Chernobyl fallout proved to be T*(ag)=3.3×10(-3)m(-2)kg(-1) and T**(ag)=4.0×10(-3)m(-2)kg(-1), respectively. The aggregated transfer factors T*(ag) for (137)Cs and spruce bark did not differ significantly in areas unequally affected by the (137)Cs fallout in the Czech Republic in 1986, and the figures for these aggregated transfer factors were very similar to the mean bark T(ag) values published from the extensively affected areas near Chernobyl. The magnitude of the (137)Cs aggregated transfer factors for spruce bark for the pre-Chernobyl and post-Chernobyl period in the Czech Republic was also very similar. The variability in spruce bark acidity caused by the operation of local anthropogenic air pollution sources did not significantly influence the accumulation and retention of (137)Cs in spruce bark. Increasing elevation of the bark sampling plots had a significant effect on raising the remaining (137)Cs specific activities in bark in areas affected by precipitation at the time when the plumes crossed, because the sums of this precipitation increased with elevation (covariable). Copyright © 2011 Elsevier B.V. All rights reserved.

  20. First results about effects of liming on saprophytic fungal communities in the Ah-horizon of a spruce forest soil in France (Vosges); Erste Resultate ueber den Effekt von Kalkung auf die Pilzpopulation (Saprophyten) im Ah-Horizont eines Fichtenwaldbodens in Frankreich (Vogesen)

    Energy Technology Data Exchange (ETDEWEB)

    Devevre, O [Centre I.N.R.A. de Nancy, Lab. de Microbiologie Forestiere, 54 - Champenoux (France); Roquebert, M F [Musee National d` Histoire Naturelle, Lab. de Cryptogamie, 75 - Paris (France); Garbaye, J [Centre I.N.R.A. de Nancy, Lab. de Microbiologie Forestiere, 54 - Champenoux (France)

    1993-04-01

    Soil fungi, including mycorrhiza, are strongly affected by zoil chemical parameters such as the ratio of calcium and/or magnesium to aluminium and the pH-value. So, it was very interesting to compare the rhizospheric microfungal flora between a declining spruce stand and a healthy spruce stand. The site chosen for this investigation was situated in the Vosges in the northeast of France. The rhizospheric soil, from the Ah-horizon of a sandy loam podzol, limed (the healthy spruce stand) or unlimed (the declining spruce stand) was sampled in a 65-year-old Norway spruce forest. The study was made 7 years after liming. Fungal isolations were performed using the dilution plate method. Pronounced differences in species abundance and composition were found between the limed and the unlimed stands. Of the 49 isolated species (24 from declining spruce plot and 34 from healthy spruce plot) only nine were found at both plots. The greatest diversity is observed at the healthy spruce stand; it may be due to the liming. This study indicates that soil microfungi could be sensitive to increased acidity of the rain with subsequent effects. (orig.) [Deutsch] Bodenpilze einschliesslich der Mykorrhizapilze sind stark von bodenchemischen Parameters wie dem Verhaeltnis von Calcium und/oder Magnesium zu Aluminium sowie vom pH-Wert des Bodens abhaengig. Deshalb wurde die Pilzmikroflora eines geschaedigten Fichtenbestand mit einem gesunden Fichtenbestand in den Vogesen, im Nordosten Frankreichs verglichen. In einem 65jaehrigen Fichtenbestand wurde der durchwurzelte Boden des Ah-Horizontes eines sandig-lehmigen Podsols einer gekalkten (gesunder Fichtenbestand) sowie einer ungekalkten Parzelle (geschaedigter Fichtenbestand) beprobt. Die Studie wurde 7 Jahre nach der Kalkung durchgefuehrt. Die Isolation der Pilze wurde anhand der Verduennungstechnik auf Kulturmedium mit DRBC-Agar durchgefuehrt. Sowohl im Artenvorkommen als auch in den Populationsstaerken bestanden betraechtliche Unterschiede

  1. Studying sulfur functional groups in Norway spruce year rings using S L-edge total electron yield spectroscopy

    International Nuclear Information System (INIS)

    Struis, Rudolf P.W.J.; Ludwig, Christian; Barrelet, Timothee; Kraehenbuehl, Urs; Rennenberg, Heinz

    2008-01-01

    Profiles of the major sulfur functional groups in mature Norway spruce wood tissue have been established for the first time. The big challenge was the development of a method suitable for sulfur speciation in samples with very low sulfur content (< 100 ppm). This became possible by synchrotron X-ray absorption spectroscopy at the sulfur L-edge in total electron yield (TEY) detection mode with thin gold-coated wood slices. Functional groups were identified using sulfur compound spectra as fingerprints. Latewood of single year rings revealed metabolic plausible sulfur forms, particularly inorganic sulfide, organic disulfide, methylthiol, and highly oxidized sulfur. Form-specific profiles with Norway spruces from three different Swiss forest sites revealed high, but hitherto little-noticed, sulfur intensities attributable to natural heartwood formation and a common, but physiologically unexpected maximum around year ring 1986 with trees from the industrialized Swiss Plateau. It is hypothesized whether it may have resulted from the huge reduction in sulfur emissions after 1980 due to Swiss policy. Comparison with total S content profiles from optical emission spectroscopy underlined the more accurate and temporally better resolved TEY data with single wood year rings and it opened novel insights into the wood cell chemistry

  2. Overstory Tree Mortality in Ponderosa Pine and Spruce-Fir Ecosystems Following a Drought in Northern New Mexico

    Directory of Open Access Journals (Sweden)

    Brian P. Oswald

    2016-10-01

    Full Text Available Drought-caused tree dieback is an issue around the world as climates change and many areas become dryer and hotter. A drought from 1998–2004 resulted in a significant tree dieback event in many of the wooded areas in portions of the Jemez Mountains and the adjacent Pajarito Plateau in northern New Mexico. The objectives of this study were to evaluate and quantify the differences in tree mortality before and after a recent drought in ponderosa pine and spruce-fir ecosystems, and to assess the effect of mechanical thinning on ponderosa pine mortality. Significant increases in mortality were observed in the unthinned ponderosa pine ecosystem. Mortality varied significantly between species and within size classes. Mechanical thinning of ponderosa pines reduced overstory mortality to non-significant levels. A lack of rainfall, snowfall, and increases in daily minimum temperature contributed most to the mortality. Adaptive management, including the use of thinning activities, appear to moderate the impact of climate change on ponderosa pine forests in this region, increasing the long-term health of the ecosystem. The impact of climate change on the spruce-fir ecosystems may accelerate successional changes.

  3. Carbon flux to woody tissues in a beech/spruce forest during summer and in response to chronic O3 exposure

    Science.gov (United States)

    The present study compares the dynamics in carbon (C) allocation of adult deciduous beech (Fagus sylvatica) and evergreen spruce (Picea abies) during summer and in response to seven-year-long exposure with twice-ambient ozone (O3) concentrations (2 × O3). Focus was on the respira...

  4. Can nutrient limitations explain low and declining white spruce growth near the Arctic treeline in the eastern Brooks Range, Alaska?

    Science.gov (United States)

    Ellison, S.; Sullivan, P. F.

    2014-12-01

    The position of the Arctic treeline is of critical importance for global carbon cycling and surface energy budgets. However, controls on tree growth at treeline remain uncertain. In the Alaskan Brooks Range, 20th century warming has caused varying growth responses among treeline trees, with trees in the west responding positively, while trees in the east have responded negatively. The prevailing explanation of this trend ascribes the negative growth response to warming-induced drought stress in the eastern Brooks Range. However, recent measurements of carbon isotope discrimination in tree rings, xylem sap flow and needle gas exchange suggest that drought stress cannot explain these regional growth declines. Additionally, evidence from the western Brooks Range suggests that nutrient availability, rather than drought stress, may be the proximate control on tree growth. In this study, we investigated the hypothesis that low and declining growth of eastern Brooks Range trees is due to low and declining soil nutrient availability, which may continue to decrease with climate change as soils become drier and microbial activity declines. We compared microclimate, tree performance, and a wide range of proxies for soil nutrient availability in four watersheds along a west-east transect in the Brooks Range during the growing seasons of 2013 and 2014. We hypothesized that soil nutrient availability would track closely with the strong west-east precipitation gradient, with higher rainfall and greater soil nutrient availability in the western Brooks Range. We expected to find that soil water contents in the west are near optimum for nitrogen mineralization, while those in the east are below optimum. Needle nitrogen concentration, net photosynthesis, branch extension growth, and growth in the main stem are expected to decline with the hypothesized decrease in soil nutrient availability. The results of our study will elucidate the current controls on growth of trees near the

  5. Decomposition of soil organic matter from boreal black spruce forest: Environmental and chemical controls

    Science.gov (United States)

    Wickland, K.P.; Neff, J.C.

    2008-01-01

    Black spruce forests are a dominant covertype in the boreal forest region, and they inhabit landscapes that span a wide range of hydrologic and thermal conditions. These forests often have large stores of soil organic carbon. Recent increases in temperature at northern latitudes may be stimulating decomposition rates of this soil carbon. It is unclear, however, how changes in environmental conditions influence decomposition in these systems, and if substrate controls of decomposition vary with hydrologic and thermal regime. We addressed these issues by investigating the effects of temperature, moisture, and organic matter chemical characteristics on decomposition of fibric soil horizons from three black spruce forest sites. The sites varied in drainage and permafrost, and included a "Well Drained" site where permafrost was absent, and "Moderately well Drained" and "Poorly Drained" sites where permafrost was present at about 0.5 m depth. Samples collected from each site were incubated at five different moisture contents (2, 25, 50, 75, and 100% saturation) and two different temperatures (10??C and 20??C) in a full factorial design for two months. Organic matter chemistry was analyzed using pyrolysis gas chromatography-mass spectrometry prior to incubation, and after incubation on soils held at 20??C, 50% saturation. Mean cumulative mineralization, normalized to initial carbon content, ranged from 0.2% to 4.7%, and was dependent on temperature, moisture, and site. The effect of temperature on mineralization was significantly influenced by moisture content, as mineralization was greatest at 20??C and 50-75% saturation. While the relative effects of temperature and moisture were similar for all soils, mineralization rates were significantly greater for samples from the "Well Drained" site compared to the other sites. Variations in the relative abundances of polysaccharide-derivatives and compounds of undetermined source (such as toluene, phenol, 4-methyl phenol, and

  6. Norway spruce needles as bioindicator of air pollution in the area of influence of the Sostanj Thermal Power Plant, Slovenia

    Energy Technology Data Exchange (ETDEWEB)

    Al Sayegh Petkovsek, Samar [ERICo Velenje, Ecological Research and Industrial Cooperation Institute, Koroska 58, 3320 Velenje (Slovenia)], E-mail: samar.petkovsek@erico.si; Batic, Franc [Agronomy Department, Biotechnical Faculty, University of Ljubljana, Jamnikarjeva 101, 1000 Ljubljana (Slovenia); Ribaric Lasnik, Cvetka [Institute of Environment and Spatial Planning, Ipavceva 18, 3000 Celje (Slovenia)

    2008-01-15

    This paper reports the results of total sulphur content, photosynthetic pigments, ascorbic acid (vitamin C) and {alpha}-tocopherol (vitamin E) analysed in current-year needles of Norway spruce (Picea abies (L.) Karst.) in the area influenced by sulphur emissions from the Sostanj Thermal Power Plant (STPP), Slovenia, in the period 1991-2004. Ten differently polluted sampling sites in the emission area of STPP were selected. After desulphurization of emission gases from STPP total sulphur content in needles decreased and vitality parameters of needles increased. Moreover, a strong correlation between the average annual emissions of SO{sub 2} from STPP and average annual sulphur content (increase) or average annual chlorophyll content (decrease) in current-year needles was found. The results showed that spruce needles may be an useful bioindicator for detecting changes in the emission rates of SO{sub 2}. - Total sulphur content in needles is a well proved indicator of sulphur burden in the environment.

  7. Modelling growth-competition relationships in trembling aspen and white spruce mixed boreal forests of Western Canada.

    Science.gov (United States)

    Huang, Jian-Guo; Stadt, Kenneth J; Dawson, Andria; Comeau, Philip G

    2013-01-01

    We examined the effect of competition on stem growth of Picea glauca and Populus tremuloides in boreal mixedwood stands during the stem exclusion stage. We combined traditional approaches of collecting competition data with dendrochronology to provide retrospective measurements of stem diameter growth. Several competition indices including stand basal area (BA), the sum of stem diameter at breast height (SDBH), and density (N) for the broadleaf and coniferous species, as well as similar indices considering only trees with diameters greater than each subject (BAGR, SDBHGR, and NGR), were evaluated. We used a nonlinear mixed model to characterize the basal area increment over the past 5, 10, 15, 20, 25, 30, and 35 years as a function of growth of nearby dominant trees, the size of the subject trees, deciduous and coniferous competition indices, and ecoregions. SDBHGR and BAGR were better predictors for spruce, and SDBHGR and NGR were better for aspen, respectively, than other indices. Results showed strongest correlations with long-term stem growth, as the best models integrated growth for 10-25 years for aspen and ≥ 25 for spruce. Our model demonstrated a remarkable capability (adjusted R(2)>0.67) to represent this complex variation in growth as a function of site, size and competition.

  8. Modelling growth-competition relationships in trembling aspen and white spruce mixed boreal forests of Western Canada.

    Directory of Open Access Journals (Sweden)

    Jian-Guo Huang

    Full Text Available We examined the effect of competition on stem growth of Picea glauca and Populus tremuloides in boreal mixedwood stands during the stem exclusion stage. We combined traditional approaches of collecting competition data with dendrochronology to provide retrospective measurements of stem diameter growth. Several competition indices including stand basal area (BA, the sum of stem diameter at breast height (SDBH, and density (N for the broadleaf and coniferous species, as well as similar indices considering only trees with diameters greater than each subject (BAGR, SDBHGR, and NGR, were evaluated. We used a nonlinear mixed model to characterize the basal area increment over the past 5, 10, 15, 20, 25, 30, and 35 years as a function of growth of nearby dominant trees, the size of the subject trees, deciduous and coniferous competition indices, and ecoregions. SDBHGR and BAGR were better predictors for spruce, and SDBHGR and NGR were better for aspen, respectively, than other indices. Results showed strongest correlations with long-term stem growth, as the best models integrated growth for 10-25 years for aspen and ≥ 25 for spruce. Our model demonstrated a remarkable capability (adjusted R(2>0.67 to represent this complex variation in growth as a function of site, size and competition.

  9. Modeling Wood Fibre Length in Black Spruce (Picea mariana (Mill. BSP Based on Ecological Land Classification

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    Elisha Townshend

    2015-09-01

    Full Text Available Effective planning to optimize the forest value chain requires accurate and detailed information about the resource; however, estimates of the distribution of fibre properties on the landscape are largely unavailable prior to harvest. Our objective was to fit a model of the tree-level average fibre length related to ecosite classification and other forest inventory variables depicted at the landscape scale. A series of black spruce increment cores were collected at breast height from trees in nine different ecosite groups within the boreal forest of northeastern Ontario, and processed using standard techniques for maceration and fibre length measurement. Regression tree analysis and random forests were used to fit hierarchical classification models and find the most important predictor variables for the response variable area-weighted mean stem-level fibre length. Ecosite group was the best predictor in the regression tree. Longer mean fibre-length was associated with more productive ecosites that supported faster growth. The explanatory power of the model of fitted data was good; however, random forests simulations indicated poor generalizability. These results suggest the potential to develop localized models linking wood fibre length in black spruce to landscape-level attributes, and improve the sustainability of forest management by identifying ideal locations to harvest wood that has desirable fibre characteristics.

  10. Adaptation of lodgepole pine and interior spruce to climate: implications for reforestation in a warming world.

    Science.gov (United States)

    Liepe, Katharina J; Hamann, Andreas; Smets, Pia; Fitzpatrick, Connor R; Aitken, Sally N

    2016-02-01

    We investigated adaptation to climate in populations of two widespread tree species across a range of contrasting environments in western Canada. In a series of common garden experiments, bud phenology, cold hardiness, and seedling growth traits were assessed for 254 populations in the interior spruce complex (Picea glauca, P. engelmannii, and their hybrids) and for 281 populations of lodgepole pine (Pinus contorta). Complex multitrait adaptations to different ecological regions such as boreal, montane, coastal, and arid environments accounted for 15-20% of the total variance. This population differentiation could be directly linked to climate variables through multivariate regression tree analysis. Our results suggest that adaptation to climate does not always correspond linearly to temperature gradients. For example, opposite trait values (e.g., early versus late budbreak) may be found in response to apparently similar cold environments (e.g., boreal and montane). Climate change adaptation strategies may therefore not always be possible through a simple shift of seed sources along environmental gradients. For the two species in this study, we identified a relatively small number of uniquely adapted populations (11 for interior spruce and nine for lodgepole pine) that may be used to manage adaptive variation under current and expected future climates.

  11. The historical role of Ips hauseri (Coleoptera: Curculionidae) in the spruce forest of Ile-Alatausky and Medeo National Parks

    Science.gov (United States)

    N. Mukhamadiev; A. Lynch; C. O' Connor; A. Sagitov; N. Ashikbaev; I. Panyushkina

    2014-01-01

    On 17 May and 27 June 2011 severe cyclonic storms damaged several hundred hectares of spruce forest (Picea schrenkiana) in the Tian Shan Mountains. Bark beetle populations increased rapidly in dead and damaged trees, particularly Ips hauseri, I. typographus, I. sexdentatus, and Piiyogenesperfossus (all Coleoptera: Curculionidae), and there is concern about the...

  12. Spread of Heterobasidion annosum in European fir and Norway spruce after 4 years of treatment with pollutants

    International Nuclear Information System (INIS)

    Raddi, P.; Mugnal, L.; Capretti, P.

    1993-01-01

    By means of a series of inoculations with S- and F-isolates of Heterobasidion annosum on plants treated with pollutants, it was observed that the treatments with ABS and H 2 SO 4 caused an increase in silver fir in the colonization by group F and decrease in the growth of the plants. In Norway spruce, both isolates were favoured by the pollutants. (orig.) [de

  13. Patterns of nucleotide diversity at photoperiod related genes in Norway spruce [Picea abies (L.) Karst].

    Science.gov (United States)

    Källman, Thomas; De Mita, Stéphane; Larsson, Hanna; Gyllenstrand, Niclas; Heuertz, Myriam; Parducci, Laura; Suyama, Yoshihisa; Lagercrantz, Ulf; Lascoux, Martin

    2014-01-01

    The ability of plants to track seasonal changes is largely dependent on genes assigned to the photoperiod pathway, and variation in those genes is thereby important for adaptation to local day length conditions. Extensive physiological data in several temperate conifer species suggest that populations are adapted to local light conditions, but data on the genes underlying this adaptation are more limited. Here we present nucleotide diversity data from 19 genes putatively involved in photoperiodic response in Norway spruce (Picea abies). Based on similarity to model plants the genes were grouped into three categories according to their presumed position in the photoperiod pathway: photoreceptors, circadian clock genes, and downstream targets. An HKA (Hudson, Kreitman and Aquade) test showed a significant excess of diversity at photoreceptor genes, but no departure from neutrality at circadian genes and downstream targets. Departures from neutrality were also tested with Tajima's D and Fay and Wu's H statistics under three demographic scenarios: the standard neutral model, a population expansion model, and a more complex population split model. Only one gene, the circadian clock gene PaPRR3 with a highly positive Tajima's D value, deviates significantly from all tested demographic scenarios. As the PaPRR3 gene harbours multiple non-synonymous variants it appears as an excellent candidate gene for control of photoperiod response in Norway spruce.

  14. Moisture performance properties of exterior sheathing products made of spruce plywood or OSB

    Energy Technology Data Exchange (ETDEWEB)

    Ojanen, T.; Ahonen, J. [VTT Building and Transport, Espoo (Finland)

    2005-02-01

    Plywood and OSB (Oriented Strand Board) are building boards that are made of timber and glue and that have good structural strength. OSB is a relatively new product that is produced from various timber materials and even some wastewood can be used as raw material for this product. Due to the easy access of raw material and economical reasons, the use of OSB has been highly increasing. The moisture safety of the building envelope depends on the properties of the building products. The moisture performance properties of the sheathing boards have not got enough attention, too often these wood based materials are equated without considering the real properties and their effect on the overall moisture performance. This research was carried out to experimentally determine the material properties and performance characteristics that have an effect on the moisture performance of building structures where OSB or spruce plywood are applied as exterior sheathing boards. A relatively representative amount of samples were used to study the moisture properties of European OSB and plywood products and a comparison to one Canadian OSB and one plywood product was done. All the products used in this research were meant to be used also as exterior sheathing boards. The results shows clearly the differences between OSB and plywood products. The products are not interchangeable. The main differences can be found from the vapour permeability levels that have an effect on the drying efficiency of building structures. The products have different performance criteria, and the climate conditions and moisture loads have to be studied to evaluate their suitability and moisture safety aspects in different applications. Water repellent features and drying efficiency are somewhat opposite properties of the products and the dimensional changes under varying moisture contents can set some boundary conditions when the optimum solution for the exterior sheathing is considered. (orig.)

  15. Evolutionary engineering strategies to enhance tolerance of xylose utilizing recombinant yeast to inhibitors derived from spruce biomass

    Directory of Open Access Journals (Sweden)

    Koppram Rakesh

    2012-05-01

    Full Text Available Abstract Background One of the crucial factors for a sustainable and economical production of lignocellulosic based bioethanol is the availability of a robust fermenting microorganism with high tolerance to inhibitors generated during the pretreatment of lignocellulosic raw materials, since these inhibitors are known to severely hinder growth and fermentation. Results A long-term adaptation in repetitive batch cultures in shake flasks using a cocktail of 12 different inhibitors and a long-term chemostat adaptation using spruce hydrolysate were used as evolutionary engineering strategies to improve the inhibitor tolerance in the metabolically engineered xylose utilizing Saccharomyces cerevisiae strain, TMB3400. The yeast was evolved for a period of 429 and 97 generations in repetitive batch cultures and chemostat cultivation, respectively. During the evolutionary engineering in repetitive batch cultures the maximum specific growth rate increased from 0.18 h-1 to 0.33 h-1 and the time of lag phase was decreased from 48 h to 24 h. In the chemostat adaptation, after 97 generations, the specific conversion rates of HMF and furfural were found to be 3.5 and 4 folds higher respectively, compared to rates after three generations. Two evolved strains (RK60-5, RKU90-3 and one evolved strain (KE1-17 were isolated from evolutionary engineering in repetitive batches and chemostat cultivation, respectively. The strains displayed significantly improved growth performance over TMB3400 when cultivated in spruce hydrolysate under anaerobic conditions, the evolved strains exhibited 25 to 38% increase in specific consumption rate of sugars and 32 to 50% increased specific ethanol productivity compared to TMB3400. The evolved strains RK60-5 and RKU90-3 were unable to consume xylose under anaerobic conditions, whereas, KE1-17 was found to consume xylose at similar rates as TMB3400. Conclusion Using evolutionary engineering strategies in batch and chemostat

  16. Host Genetics and Environment Drive Divergent Responses of Two Resource Sharing Gall-Formers on Norway Spruce: A Common Garden Analysis.

    Science.gov (United States)

    Axelsson, E Petter; Iason, Glenn R; Julkunen-Tiitto, Riitta; Whitham, Thomas G

    2015-01-01

    A central issue in the field of community genetics is the expectation that trait variation among genotypes play a defining role in structuring associated species and in forming community phenotypes. Quantifying the existence of such community phenotypes in two common garden environments also has important consequences for our understanding of gene-by-environment interactions at the community level. The existence of community phenotypes has not been evaluated in the crowns of boreal forest trees. In this study we address the influence of tree genetics on needle chemistry and genetic x environment interactions on two gall-inducing adelgid aphids (Adelges spp. and Sacchiphantes spp.) that share the same elongating bud/shoot niche. We examine the hypothesis that the canopies of different genotypes of Norway spruce (Picea abies L.) support different community phenotypes. Three patterns emerged. First, the two gallers show clear differences in their response to host genetics and environment. Whereas genetics significantly affected the abundance of Adelges spp. galls, Sacchiphantes spp. was predominately affected by the environment suggesting that the genetic influence is stronger in Adelges spp. Second, the among family variation in genetically controlled resistance was large, i.e. fullsib families differed as much as 10 fold in susceptibility towards Adelges spp. (0.57 to 6.2 galls/branch). Also, the distribution of chemical profiles was continuous, showing both overlap as well as examples of significant differences among fullsib families. Third, despite the predicted effects of host chemistry on galls, principal component analyses using 31 different phenolic substances showed only limited association with galls and a similarity test showed that trees with similar phenolic chemical characteristics, did not host more similar communities of gallers. Nonetheless, the large genetic variation in trait expression and clear differences in how community members respond to host

  17. Host Genetics and Environment Drive Divergent Responses of Two Resource Sharing Gall-Formers on Norway Spruce: A Common Garden Analysis.

    Directory of Open Access Journals (Sweden)

    E Petter Axelsson

    Full Text Available A central issue in the field of community genetics is the expectation that trait variation among genotypes play a defining role in structuring associated species and in forming community phenotypes. Quantifying the existence of such community phenotypes in two common garden environments also has important consequences for our understanding of gene-by-environment interactions at the community level. The existence of community phenotypes has not been evaluated in the crowns of boreal forest trees. In this study we address the influence of tree genetics on needle chemistry and genetic x environment interactions on two gall-inducing adelgid aphids (Adelges spp. and Sacchiphantes spp. that share the same elongating bud/shoot niche. We examine the hypothesis that the canopies of different genotypes of Norway spruce (Picea abies L. support different community phenotypes. Three patterns emerged. First, the two gallers show clear differences in their response to host genetics and environment. Whereas genetics significantly affected the abundance of Adelges spp. galls, Sacchiphantes spp. was predominately affected by the environment suggesting that the genetic influence is stronger in Adelges spp. Second, the among family variation in genetically controlled resistance was large, i.e. fullsib families differed as much as 10 fold in susceptibility towards Adelges spp. (0.57 to 6.2 galls/branch. Also, the distribution of chemical profiles was continuous, showing both overlap as well as examples of significant differences among fullsib families. Third, despite the predicted effects of host chemistry on galls, principal component analyses using 31 different phenolic substances showed only limited association with galls and a similarity test showed that trees with similar phenolic chemical characteristics, did not host more similar communities of gallers. Nonetheless, the large genetic variation in trait expression and clear differences in how community members

  18. Effects of ABA application on cessation of shoot elongation in long-day grown Norway spruce seedlings.

    Science.gov (United States)

    Heide, O M

    1986-06-01

    Abscisic acid (ABA) was applied in lanolin to apical buds of Norway spruce (Picea abies (L.) Karst.) seedlings actively growing in a 24 h photoperiod. At a rate of 100 microg per plant, ABA suspended shoot elongation for about three weeks in the majority of plants but failed to induce normal winter buds. The role of ABA in the induction of dormancy is thus uncertain in conifers as well as in deciduous woody plants.

  19. Assessing health literacy in the eastern and middle-eastern cultures

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    Satish Chandrasekhar Nair

    2016-08-01

    Full Text Available Abstract Background Health literacy is a term employed to assess the ability of people to meet the increasing demands related to health in a rapidly evolving society. Low health literacy can affect the social determinants of health, health outcomes and the use of healthcare services. The purpose of the study was to develop a survey construct to assess health literacy within the context of regional culture. Different socioeconomic status among the Eastern and Middle Eastern countries may restrict, health information access and utilization for those with low literacy. Methods By employing expert panel, Delphi technique, focus group methodologies, and pre-testing using participants (N = 900 from the UAE and India, a survey construct to the Eastern-Middle Eastern cultures was developed. Reliability was assessed using Cronbach’s α and validity using Factor analysis. Kiaser-Meyer-Olkin (KMO sampling adequacy and Bartlett’s tests were used to assess the strength of the relationship among the variables. Results Inclusion of non-health related items were found to be critical in the authentic assessment of health literacy in the Eastern and Middle Eastern population given the influence of social desirability. Thirty-two percentage of the original 19-item construct was eliminated by the focus group for reasons of relevance and impact for the local culture. Field pretesting participants from two countries, indicated overall construct reliability (Cronbach’s α =0.85, validity and consistency (KMO value of 0.92 and Bartlett’s test of sphericity was significant. Conclusion The Eastern-Middle Eastern Adult Health Literacy (EMAHL13, screening instrument is brief, simple, a useful indicator of whether or not a patient can read. It assessespatients’ ability to comprehend by distinguishing between health and non-health related items. The EMAHL13 will be a useful too for the reliable assessment of health literacy in countries, where culture plays a

  20. Assessing health literacy in the eastern and middle-eastern cultures.

    Science.gov (United States)

    Nair, Satish Chandrasekhar; Satish, Karthyayani Priya; Sreedharan, Jayadevan; Ibrahim, Halah

    2016-08-19

    Health literacy is a term employed to assess the ability of people to meet the increasing demands related to health in a rapidly evolving society. Low health literacy can affect the social determinants of health, health outcomes and the use of healthcare services. The purpose of the study was to develop a survey construct to assess health literacy within the context of regional culture. Different socioeconomic status among the Eastern and Middle Eastern countries may restrict, health information access and utilization for those with low literacy. By employing expert panel, Delphi technique, focus group methodologies, and pre-testing using participants (N = 900) from the UAE and India, a survey construct to the Eastern-Middle Eastern cultures was developed. Reliability was assessed using Cronbach's α and validity using Factor analysis. Kiaser-Meyer-Olkin (KMO) sampling adequacy and Bartlett's tests were used to assess the strength of the relationship among the variables. Inclusion of non-health related items were found to be critical in the authentic assessment of health literacy in the Eastern and Middle Eastern population given the influence of social desirability. Thirty-two percentage of the original 19-item construct was eliminated by the focus group for reasons of relevance and impact for the local culture. Field pretesting participants from two countries, indicated overall construct reliability (Cronbach's α =0.85), validity and consistency (KMO value of 0.92 and Bartlett's test of sphericity was significant). The Eastern-Middle Eastern Adult Health Literacy (EMAHL13), screening instrument is brief, simple, a useful indicator of whether or not a patient can read. It assessespatients' ability to comprehend by distinguishing between health and non-health related items. The EMAHL13 will be a useful too for the reliable assessment of health literacy in countries, where culture plays a significant impact. This will be the first steptowards providing