Lindow, Bent Erik Kramer
The marine Lillebaelt Clay Formation of central Denmark is of early-middle Eocene age (late Ypresian - middle Lutetian; microfossil zones NP 13-NP 15). Over 20 bird fossils collected by amateur palaeontologists have been acquired through the Danish national ‘Danekrae' fossil treasure trove...
Lindow, Bent Erik Kramer
The marine Lillebaelt Clay Formation of central Denmark is of early-middle Eocene age (late Ypresian - middle Lutetian; microfossil zones NP 13-NP 15). Over 20 bird fossils collected by amateur palaeontologists have been acquired through the Danish national ‘Danekrae' fossil treasure trove...... legislation. The fossils are preserved in clay ironstone concretions and almost two-thirds are isolated skulls preserved three-dimensionally. Bird fossils of this age and degree of preservation are rare in an international context. The fossils indicate a very diverse assemblage consisting of both marine...... and terrestrial forms. These include at least one pelagornithid or 'pseudo-toothed bird'; two or three taxa with charadriiform affinities (shorebirds and allies); a massive, narrow-beaked psittaciform (parrots and allies); a large rallid (rail) and one lithornithid (extinct, volant palaeognaths). The Lillebaelt...
Lindow, Bent Erik Kramer
A number of hitherto undescribed fossil bird remains have been recovered from the Lillebælt Clay Formation of central Denmark, which is early-middle Eocene in age (~50 to 43 mya). The core of the material consists of fossils acquired through the Danish ‘Danekræ' fossil treasure trove legislation....... Almost two-thirds of the fossils are isolated skulls preserved three-dimensionally in clay ironstone concretions; bird fossils of this age and degree of preservation are extremely rare in an international context. A preliminary investigation has revealed the presence of at least one odontopterygid......, a member of the extinct 'pseudo-toothed birds' and the first representative of this group known from Denmark. Other taxa present include remains of Lithornithidae and a new taxon possessing a massive, psittacid-like beak. The Lillebælt Clay Formation birds are temporally placed just after the Early Eocene...
Cappelli, Carlotta; Agnini, Claudia; Yamamoto, Yuhji
The early-middle Eocene interval documents the shift from the warmest greenhouse conditions occurred during the Early Eocene Climatic Optimum (EECO, 52-50 Ma) to the beginning of the cooling phase which led to the Oligocene icehouse regime. This important transition is well expressed as a reversal in the global oxygen and carbonate isotope trends (Zachos et al., 2001). Moreover, this interval was a time of remarkable transformation in the marine biosphere. Communities of calcareous nannoplankton, marine calcifying algae at the base of the oceans food chain, experienced transient and permanent profound changes. Calcareous nannofossil are regarded as remarkable tools both in biostratigraphy and paleoecology, with several taxa that show different responses to changes in physical parameters of surface waters. Here, we aim to document calcareous nannoplankton assemblage changes across the early-middle Eocene transition, in order to upset the biostratigraphic framework and to increase comprehension of how phytoplankton communities responded to paleoenvironmental changes at that time. The sedimentary successions recovered at IODP Site U1410 (Exp. 342; 41˚ 19.6987'N; 49˚ 10.1995'W, Norris et al., 2012) on the Southeast Newfoundland Ridge (NW Atlantic) offer an expanded record of the early-middle Eocene interval that is marked by an increase in accumulation rate related to sedimentation of clay-rich nannofossil oozes. Quantitative analysis of calcareous nannofossil assemblages was conducted, encompassing calcareous nannofossil Zones NP12 -NP15 or CNE4-CNE10 (Martini, 1971; Agnini et al., 2014). The study interval records the appearance and proliferation of Noelaerhabdaceae family (i.e, Reticulofenestra/Dictyococcites group), which can be considered one of the most significant shifts in the assemblage structure of the Paleogene. This change was probably favored by modifications in surface water chemistry. The middle Eocene clay-rich sediments contain well preserved
Full Text Available The early Eocene locality of La Borie is located in the village of Saint-Papoul, in southern France. These Eocene flu-vio-lacustrine clay deposits have yielded numerous vertebrate remains. Mammalian taxa found in the fossiliferous levels indicate an age near the reference level MP 8–9, which corresponds to the middle Ypresian, early Eocene. Here we provide a detailed description of the avian remains that were preliminarily reported in a recent study of the vertebrate fauna from La Borie. A maxilla, a quadrate, cervical vertebrae, a femur and two tibiotarsi are assigned to the giant ground bird Gastornis parisiensis (Gastornithidae. These new avian remains add to the fossil record of Gastornis, which is known from the late Paleocene to middle Eocene of Europe, early Eocene of Asia and early Eocene of North America. Gastornis parisiensis differs from the North American Gastornis giganteus in several features, including the more ventral position of the narial openings and the slender orbital process of quadrate. Two tibiotarsi and one tarsometatarsus are assigned to a new genus and species of Geranoididae, Galligeranoides boriensis gen. et sp. nov. So far, this family was known only from the early and middle Eocene of North America. The fossils from La Borie constitute the first record of the Geranoididae in Europe. We show that Gastornis coexisted with the Geranoididae in the early Eocene of both Europe (La Borie and North America (Willwood Formation. The presence of Geranoididae and the large flightless bird Gastornis on either side of the present-day North Atlantic provides further evidence that a high-latitude land connection existed between Europe and North America in the early Eocene.
Bourdon, Estelle; Mourer-Chauviré, Cécile; Laurent, Yves
The Early Eocene locality of La Borie is located in the village of Saint-Papoul, in southern France. These Eocene fluvio-lacustrine clay deposits have yielded numerous vertebrate remains. Mammalian taxa found in the fossiliferous levels indicate an age near the reference level MP 8–9, which...
Dyke, Gareth; Lindow, Bent Erik Kramer
The pattern, pace and extent of the evolutionary radiation of modern birds (Neornithes) by the end-Cretaceous (65 Ma) has long been debated. Well-dated, taphonomically understood and phylogenetically constrained fossil birds from both sides of the Cretaceous-Paleogene (K-Pg) boundary are required...... to quantify the shape of this radiation, but have largely been lacking. Here we report on a large collection of fossil birds from the Lower Eocene of Denmark (ca. 54 Ma) that includes three-dimensionally preserved, articulated specimens from carbonate concretions as well as skeletal imprints and feathers....... These birds are from a marine diatomite sequence (the Fur Formation), a low-energy deep-water preservational environment unique to the Cretaceous and Paleogene avian fossil record. We present taphonomic and palaeoecological information gleaned from these birds that in combination with phylogenetic data have...
Roberto E Yury-Yáñez
Full Text Available Paleogene records of birds in the Eastern margin of the Pacific Ocean have increased in recent years, being almost exclusively restricted to fossil Sphenisciformes (penguins. New avian remains (Ornithurae, Neornithes from Middle-to-Late Eocene levels of the Estratos de Algarrobo unit, in Algarrobo, central Chile, are disclosed in the present work. These new finds are significant in representing the first non-spheniscid bird remains of Middle to Late Eocene age, recovered in mid-latitudes of the eastern Pacific and probably belonging to a procelarid. It complements the regional record of Eocene birds, previously known only at high-latitudes such as Seymour Island (Antarctica and Magallanes (Chile, and low-latitude locations in Peru.Los registros de aves paleógenas en el margen oriental del Océano Pacífico se han incrementado en los últimos años, estando casi exclusivamente restringidos a Sphenisciformes fósiles (pingüinos. Nuevos fósiles avianos (Ornithurae, Neornithes provenientes de niveles del Eoceno Medio a Tardío de los Estratos de Algarrobo, en Algarrobo, Chile central, son dados a conocer en el presente trabajo. Estos nuevos hallazgos son significativos en representar los primeros restos de aves no-spheníscidos de edad Eoceno Medio a Tardío, recuperadas en latitudes intermedias del Pacífico oriental y probablemente pertenezcan a un procelárido. Este hallazgo complementa el registro regional del grupo durante el Eoceno, anteriormente conocido en localidades de altas latitudes como isla Seymour (Antártica y Magallanes (Chile, y localidades de bajas latitudes en Perú.
Birds play an important role in studies addressing the diversity and species richness of tropical ecosystems, but because of the poor avian fossil record in all extant tropical regions, a temporal perspective is mainly provided by divergence dates derived from calibrated molecular analyses. Tropical ecosystems were, however, widespread in the Northern Hemisphere during the early Cenozoic, and the early Eocene German fossil site Messel in particular has yielded a rich avian fossil record. The Messel avifauna is characterized by a considerable number of flightless birds, as well as a high diversity of aerial insectivores and the absence of large arboreal birds. With about 70 currently known species in 42 named genus-level and at least 39 family-level taxa, it approaches extant tropical biotas in terms of species richness and taxonomic diversity. With regard to its taxonomic composition and presumed ecological characteristics, the Messel avifauna is more similar to the Neotropics, Madagascar, and New Guinea than to tropical forests in continental Africa and Asia. Because the former regions were geographically isolated during most of the Cenozoic, their characteristics may be due to the absence of biotic factors, especially those related to the diversification of placental mammals, which impacted tropical avifaunas in Africa and Asia. The crown groups of most avian taxa that already existed in early Eocene forests are species-poor. This does not support the hypothesis that the antiquity of tropical ecosystems is key to the diversity of tropical avifaunas, and suggests that high diversification rates may be of greater significance. © 2016 Cambridge Philosophical Society.
Eberle, Jaelyn J.; Gottfried, Michael D.; Hutchison, J. Howard; Brochu, Christopher A.
BACKGROUND: Discovery of Eocene non-marine vertebrates, including crocodylians, turtles, bony fishes, and mammals in Canada's High Arctic was a critical paleontological contribution of the last century because it indicated that this region of the Arctic had been mild, temperate, and ice-free during the early - middle Eocene (∼53-50 Ma), despite being well above the Arctic Circle. To date, these discoveries have been restricted to Canada's easternmost Arctic - Ellesmere and Axel Heiberg Island...
For the production of glass three ingredients are necessary: sand, a flux to reduce the melting-temperature and calcium to reduce the danger of glass corrosion. The first objects of glass were made with calcium-rich ashes of halophytic plants, until, in the first millennium BC, the glassmakers began to use natron as a flux adding calcium deliberately or choosing a calcium-rich sand. Natron, a mineral applied to fertilize or to preserve, as a spice, a detergent or part of medical and cosmetic articles, was exploited in the regions south and east of the Mediterranean, so the Central European glassmakers had to import natron or the prefabricated raw glass for their work. Beginning in the 8th century AD in Central Europe the flux changed again: The glassmakers increasingly used ashes from wood growing in their native regions so becoming independent of the necessity to import the raw materials. There are various reasons for this change: First, the Mediterranean was no longer the trade area it had been at the time of the antique Roman Empire due to the activities of the Byzantine navy. Then, the climatic change in the 8th century and political upheavals during the 9th century in Egypt--being the main supplier of natron--caused a decrease in exploitation and trade with this good. Finally, the Egyptian state established a monopoly on the natron production, causing a permanent price increase. Nevertheless, during the Early Middle Ages natron was imported into Europe, although not necessarily for glass production. The article shows that glassmakers of Central Europe were able to produce glass since the end of the Western Roman Empire on the basis of the transfer of raw materials and know-how from the East. From the 8th century onwards they emancipated themselves from the dependency on imports by discovering and using native materials for glass production.
The high abundances and cyclic distribution of remains of the freshwater fern Azolla in early middle Eocene sediments from the Arctic Ocean have previously been related to episodic surface water freshening, which was speculated to be orbitally modulated. Our integrated palynological and
Jaelyn J Eberle
Full Text Available BACKGROUND: Discovery of Eocene non-marine vertebrates, including crocodylians, turtles, bony fishes, and mammals in Canada's High Arctic was a critical paleontological contribution of the last century because it indicated that this region of the Arctic had been mild, temperate, and ice-free during the early - middle Eocene (∼53-50 Ma, despite being well above the Arctic Circle. To date, these discoveries have been restricted to Canada's easternmost Arctic - Ellesmere and Axel Heiberg Islands (Nunavut. Although temporally correlative strata crop out over 1,000 km west, on Canada's westernmost Arctic Island - Banks Island, Northwest Territories - they have been interpreted as predominantly marine. We document the first Eocene bony fish and crocodyliform fossils from Banks Island. PRINCIPAL FINDINGS: We describe fossils of bony fishes, including lepisosteid (Atractosteus, esocid (pike, and amiid, and a crocodyliform, from lower - middle Eocene strata of the Cyclic Member, Eureka Sound Formation within Aulavik National Park (∼76°N. paleolat.. Palynology suggests the sediments are late early to middle Eocene in age, and likely spanned the Early Eocene Climatic Optimum (EECO. CONCLUSIONS/SIGNIFICANCE: These fossils extend the geographic range of Eocene Arctic lepisosteids, esocids, amiids, and crocodyliforms west by approximately 40° of longitude or ∼1100 km. The low diversity bony fish fauna, at least at the family level, is essentially identical on Ellesmere and Banks Islands, suggesting a pan-High Arctic bony fish fauna of relatively basal groups around the margin of the Eocene Arctic Ocean. From a paleoclimatic perspective, presence of a crocodyliform, gar and amiid fishes on northern Banks provides further evidence that mild, year-round temperatures extended across the Canadian Arctic during early - middle Eocene time. Additionally, the Banks Island crocodyliform is consistent with the phylogenetic hypothesis of a Paleogene divergence
Eberle, Jaelyn J; Gottfried, Michael D; Hutchison, J Howard; Brochu, Christopher A
Discovery of Eocene non-marine vertebrates, including crocodylians, turtles, bony fishes, and mammals in Canada's High Arctic was a critical paleontological contribution of the last century because it indicated that this region of the Arctic had been mild, temperate, and ice-free during the early - middle Eocene (∼53-50 Ma), despite being well above the Arctic Circle. To date, these discoveries have been restricted to Canada's easternmost Arctic - Ellesmere and Axel Heiberg Islands (Nunavut). Although temporally correlative strata crop out over 1,000 km west, on Canada's westernmost Arctic Island - Banks Island, Northwest Territories - they have been interpreted as predominantly marine. We document the first Eocene bony fish and crocodyliform fossils from Banks Island. We describe fossils of bony fishes, including lepisosteid (Atractosteus), esocid (pike), and amiid, and a crocodyliform, from lower - middle Eocene strata of the Cyclic Member, Eureka Sound Formation within Aulavik National Park (∼76°N. paleolat.). Palynology suggests the sediments are late early to middle Eocene in age, and likely spanned the Early Eocene Climatic Optimum (EECO). These fossils extend the geographic range of Eocene Arctic lepisosteids, esocids, amiids, and crocodyliforms west by approximately 40° of longitude or ∼1100 km. The low diversity bony fish fauna, at least at the family level, is essentially identical on Ellesmere and Banks Islands, suggesting a pan-High Arctic bony fish fauna of relatively basal groups around the margin of the Eocene Arctic Ocean. From a paleoclimatic perspective, presence of a crocodyliform, gar and amiid fishes on northern Banks provides further evidence that mild, year-round temperatures extended across the Canadian Arctic during early - middle Eocene time. Additionally, the Banks Island crocodyliform is consistent with the phylogenetic hypothesis of a Paleogene divergence time between the two extant alligatorid lineages Alligator
The archaeology of Britain during the early Middle Pleistocene (MIS 19-12) is represented by a number of key sites across eastern and southern England. These sites include Pakefield, Happisburgh 1, High Lodge, Warren Hill, Waverley Wood, Boxgrove, Kent's Cavern, and Westbury-sub-Mendip, alongside a 'background scatter' lithic record associated with the principal river systems (Bytham, pre-diversion Thames, and Solent) and raised beaches (Westbourne-Arundel). Hominin behaviour can be characterised in terms of: preferences for temperate or cool temperate climates and open/woodland mosaic habitats (indicated by mammalian fauna, mollusca, insects, and sediments); a biface-dominated material culture characterised by technological diversity, although with accompanying evidence for distinctive core and flake (Pakefield) and flake tool (High Lodge) assemblages; probable direct hunting-based subsistence strategies (with a focus upon large mammal fauna); and generally locally-focused spatial and landscape behaviours (principally indicated by raw material sources data), although with some evidence of dynamic, mobile and structured technological systems. The British data continues to support a 'modified short chronology' to the north of the Alps and the Pyrenees, with highly sporadic evidence for a hominin presence prior to 500-600 ka, although the ages of key assemblages are subject to ongoing debates regarding the chronology of the Bytham river terraces and the early Middle Pleistocene glaciations of East Anglia.
Grogan, D. S.; Whiteside, J. H.; Musher, D.; Rosengard, S. Z.; Vankeuren, M. A.; Pancost, R. D.
The lacustrine Green River Formation is known to span ≥15 million years through the early-middle Eocene, and recent work on radioisotopic dating has provided a framework on which to build ties to the orbitally-tuned marine Eocene record. Here we present a spliced stack of Fischer assay data from drilled cores of the Green River Formation that span both an East-West and a North-South transect of the Uinta Basin of Utah. Detailed work on two cores demonstrate that Fischer assay measurements covary with total organic carbon and bulk carbon isotopes, allowing us to use Fisher assay results as a representative carbon cycling proxy throughout the stack. We provide an age model for this core record by combining radioisotopic dates of tuff layers with frequency analysis of Fischer assay measurements. Identification of orbital frequencies tied directly to magnetochrons through radioisotopic dates allows for a direct comparison of the terrestrial to the marine Eocene record. Our analysis indicates that the marker beds used to correlate the stack cores represent periods of enhanced lake productivity and extreme carbon burial; however, unlike the hyperthermal events that are clearly marked in the marine Eocene record, the hydrocarbon-rich "Mahogany Bed" period of burial does not correspond to a clear carbon isotope excursion. This suggests that the terrestrial realm may have experienced extreme ecological responses to relatively small perturbations in the carbon cycle during the Early Eocene Climatic Optimum. To investigate the ecological responses to carbon cycle perturbations through the hydrocarbon rich beds, we analyzed a suite of microbial biomarkers, finding evidence for cyanobacteria, dinoflagellates, and potentially green sulfur bacteria. These taxa indicate fluctuating oxic/anoxic conditions in the lake during abrupt intervals of carbon burial, suggesting a lake biogeochemical regime with no modern analogues.
Judd, E. J.; Ivany, L. C.; Miklus, N. M.; Uveges, B. T.; Junium, C. K.
The Eocene Epoch was a time of large-scale global climate change, experiencing both the warmest temperatures of the Cenozoic and the onset of southern hemisphere glaciation. The record of average global temperatures throughout this transition is reasonably well constrained, however considerably less is known about the accompanying changes in seasonality. Seasonally resolved temperature data provide a wealth of information not readily available from mean annual temperature data alone. These data are particularly important in the climatically sensitive high latitudes, as they can elucidate the means by which climate changes and the conditions necessary for the growth of ice sheets. Several recent studies, however, have suggested the potential for monsoonal precipitation regimes in the early-middle Eocene high latitudes, which complicates interpretation of seasonally resolved oxygen isotope records in shallow nearshore marine settings. Seasonal precipitation and runoff could create a brackish, isotopically depleted lens in these environments, depleting summertime δ18Ocarb and thereby inflating the inferred mean and range of isotope-derived temperatures. Here, we assess intra-annual variations in temperature in shallow nearshore Antarctic waters during the middle and late Eocene, inferred from high-resolution oxygen isotope profiles from accretionary bivalves of the La Meseta Formation, Seymour Island, Antarctica. To address concerns related to precipitation and runoff, we also subsample exceptionally preserved fossil driftwood from within the formation and use seasonal differences in δ13Corg values to estimate the ratio of summertime to wintertime precipitation. Late Eocene oxygen isotope profiles exhibit strongly attenuated seasonal amplitudes and more enriched mean annual values in comparison with data from the middle Eocene. Preliminary fossil wood data are not indicative of a strongly seasonal precipitation regime, implying that intra-annual variation in oxygen
Juan C García-R
Full Text Available Central to our understanding of the timing of bird evolution is debate about an apparent conflict between fossil and molecular data. A deep age for higher level taxa within Neoaves is evident from molecular analyses but much remains to be learned about the age of diversification in modern bird families and their evolutionary ecology. In order to better understand the timing and pattern of diversification within the family Rallidae we used a relaxed molecular clock, fossil calibrations, and complete mitochondrial genomes from a range of rallid species analysed in a Bayesian framework. The estimated time of origin of Rallidae is Eocene, about 40.5 Mya, with evidence of intrafamiliar diversification from the Late Eocene to the Miocene. This timing is older than previously suggested for crown group Rallidae, but fossil calibrations, extent of taxon sampling and substantial sequence data give it credence. We note that fossils of Eocene age tentatively assigned to Rallidae are consistent with our findings. Compared to available studies of other bird lineages, the rail clade is old and supports an inference of deep ancestry of ground-dwelling habits among Neoaves.
Waddell, Lindsey M.; Moore, Theodore C.
Stable isotope analysis was performed on the structural carbonate of fish bone apatite from early and early middle Eocene samples (˜55 to ˜45 Ma) recently recovered from the Lomonosov Ridge by Integrated Ocean Drilling Program Expedition 302 (the Arctic Coring Expedition). The δ18O values of the Eocene samples ranged from -6.84‰ to -2.96‰ Vienna Peedee belemnite, with a mean value of -4.89‰, compared to 2.77‰ for a Miocene sample in the overlying section. An average salinity of 21 to 25‰ was calculated for the Eocene Arctic, compared to 35‰ for the Miocene, with lower salinities during the Paleocene Eocene thermal maximum, the Azolla event at ˜48.7 Ma, and a third previously unidentified event at ˜47.6 Ma. At the Azolla event, where the organic carbon content of the sediment reaches a maximum, a positive δ13C excursion was observed, indicating unusually high productivity in the surface waters.
Skoglund, G; Nockert, M; Holst, B
Nowadays most plant textiles used for clothing and household are made of cotton and viscose. Before the 19th century however, plant textiles were mainly made from locally available raw materials, in Scandinavia these were: nettle, hemp and flax. It is generally believed that in Viking and early Middle Ages Scandinavia hemp was used only for coarse textiles (i.e. rope and sailcloth). Here we present an investigation of 10 Scandinavian plant fibre textiles from the Viking and Early Middle Ages, believed to be locally produced. Up till now they were all believed to be made of flax. We show that 4 textiles, including two pieces of the famous Överhogdal Viking wall-hanging are in fact made with hemp (in three cases hemp and flax are mixed). This indicates that hemp was important, not only for coarse but also for fine textile production in Viking and Early Middle Ages in Scandinavia.
Students use a dead bird to learn about bird life, anatomy, and death. Students examine a bird body and discuss what happened to the bird. Uses outdoor education as a resource for learning about animals. (SAH)
Paraskevas, George K; Koutsouflianiotis, Konstantinos N; Nitsa, Zoi; Demesticha, Theano; Skandalakis, Panagiotis
The evolution of knowledge regarding the anatomy and physiology of the spleen throughout Antiquity and the Early Middle Ages is described, and general perceptions about this organ during different eras along this time line are presented. The original words of great physicians from the period of time stretching from Ancient Egypt to the Avicennan era are quoted and discussed to demonstrate how knowledge of the spleen has evolved and to present the theories that dominated each era. Furthermore, theories about illnesses relating to the spleen are reported, which show how this organ was perceived-in terms of its function and anatomy-during each era.
Dallanave, E.; Agnini, C.; Pascher, K. M.; Maurizot, P.; Bachtadse, V.; Hollis, C. J.; Dickens, G. R.; Collot, J.; Sevin, B.; Strogen, D.; Monesi, E.
Published seismic profiles acquired from the Tasman Sea and northern Zealandia area (southwest Pacific) point to a widespread Eocene convergent deformation of oceanic and continental crust, with reverse faults and uplift (Tectonic Event of the Cenozoic in the Tasman Area; TECTA). The TECTA is interpreted as the precursor of the Tonga-Kermadec subduction initiation. Grande Terre is the main island of the New Caledonia archipelago and the largest emergent portion of northern Norfolk Ridge (part of northern Zealandia). Eocene sedimentary records exposed in Grande Terre contain a transition from pelagic micrite to terrigenous-rich calciturbidites, marking a shift from passive margin to convergent tectonic regime. This could represent the local expression of the convergence inception observed on a regional scale. We conducted an integrated magneto-biostratigraphic study, based on calcareous nannofossil and radiolaria, of two early-middle Eocene records cropping out near Noumea (southwest Grande Terre) and Koumac (northwest Grande Terre). The natural remanent magnetization of the sediments is complicated by multiple vector components, likely related to the late Eocene obduction, but a characteristic remanent magnetization has been successfully isolated. Overall the record spans from magnetic polarity Chron C23n to C18n, i.e. from 51 to 39 Ma. In this robust magnetic polarity-based chronological frame, the pelagic micrite to terrigenous-rich calciturbidites occurred near the top of Chron C21n and is dated 46 Ma. Furthermore, the magnetic mineral assemblage within part of the calciturbidites consists of hematite associated with maghemite. This association indicates emergent land as source of the terrigenous, suggesting a considerable uplift. Because 94% of the Zealandia continent is submerged, ocean drilling is needed to gauge the full extent and timing of Eocene compressive deformation revealed by the seismic profiles acquired in the Tasman area. This is a primary aim of
Waterhouse, David; Lindow, Bent Erik Kramer; Zelenkov, Nikita
Two new fossil psittaciform birds from the Lower Eocene ‘Mo Clay' (Fur Formation) of Denmark (c. 54 Ma) are described. An unnamed specimen is assigned to the extinct avian family of stem-group parrots, Pseudasturidae (genus and species incertae sedis), while a second (Mopsitta tanta gen. et sp. nov.......) is the largest fossil parrot yet known. Both specimens are the first fossil records of these birds from Denmark. Although the phylogenetic position of Mopsitta is unclear (it is classified as family incertae sedis), this form is phylogenetically closer to Recent Psittacidae than to other known Palaeogene...
Elsik, W.C. (MycoStrat Connection, Houston, TX (United States))
Palynomorph sequences of several orders of magnitude were found in the Queen City and Weches formations respectively at Six Mile and Burleson bluffs on the Brazos River, Milam and Burleson counties, Texas. The long term development of the subtropical to tropical Claibornian palynoflora included Engelhardtia spp., Friedrichipollis claibornensis, Nudopollis terminalis, Pollenites laesius and Symplocoipollenites spp. Shorter term fluctuations in sea level were reflected by common herbaceous pollen in the Queen City, and common mangrove pollen in the Weches. Paleoenvironments were marginally to fully marine; dinocysts occurred throughout. The Wetzeliella group of dinocysts were present only in the Queen City at Six Mile Bluff. Late Paleocene to Early Eocene pollen, and Early Middle Eocene pollen with last effective occurrences near the Queen City and Weches boundary included Aesculiidites circumstriatus, Annona foveoreticulata and a new species of Platycarya. Five short term warmer-cooler couplet events were represented by successive abundance peaks of Juglandaceae followed by Ulmus; Alnus supports the three upper Ulmus peaks. One deep water event was recorded by an abundance of fresh water Pediastrum at the Queen City and Weches boundary. That boundary event was bracketed by two of the Alnus and Ulmus peaks.
Jens Lorenz Franzen
Full Text Available The early Middle Eocene locality of Grube Messel, near Darmstadt (Germany, is famous for its complete vertebrate skeletons. The degree of preservation of soft tissues, such as body silhouettes, internal organs and gut contents, is frequently remarkable. The present specimen was analyzed for remnants of the reproductive system. Classic anatomy and osteology and high-resolution micro-x-ray were applied to describe the fetus of the European Eocene equoid Eurohippus messelensis. Scanning electronic microscopy (SEM was used for determination of soft tissue remnants. The fetus is the earliest and best-preserved fossil specimen of its kind. The postcranial fetal skeleton is almost complete and largely articulated, allowing the conclusion that the pregnant mare was in late gestation. The apparent intrauterine position of the fetus is normal for the phase of pregnancy. Death of mare and fetus were probably not related to problems associated with parturition. Soft tissue interpreted as the uteroplacenta and a broad uterine ligament are preserved due to bacterial activity and allow considerations on the evolutionary development of the structures.
Khramov, Alexander V; Makarkin, Vladimir N
Four genera and four species of Osmylopsychopidae are described from the Jurassic of Kyrgyzstan, Central Asia: Oligophlebiopsis biramosa gen. et sp. nov. (Early Jurassic of Sogyuty); Osmylopsychoides anteromedialis gen. et sp. nov., Psychostoechotes undulatus gen. et sp. nov. and Osmylopsychostoechus sogulensis gen. et sp. nov. (all from the late Early to early Middle Jurassic of Sai-Sagul). By their poorly-developed outer gradate series of crossveins, these taxa (except O. anteromedialis gen. et sp. nov.) are more similar to Triassic genera than to the Middle/Late Jurassic Osmylopsychopidae (particularly from Daohugou, China). Two isolated hind wings from Sai-Sagul (i.e., Osmylopsychostoechus sp. and Osmylopsychopidae gen. et sp. indet.) are preliminarily assigned to this family.
National Oceanic and Atmospheric Administration, Department of Commerce — This data set contains sensitive biological resource data for wading birds, shorebirds, waterfowl, raptors, diving birds, seabirds, passerine birds, and gulls and...
National Oceanic and Atmospheric Administration, Department of Commerce — This data set contains sensitive biological resource data for wading birds, shorebirds, waterfowl, raptors, diving birds, seabirds, passerine birds, gulls, and terns...
Engelbrecht, Andrea; Mörs, Thomas; Reguero, Marcelo A.; Kriwet, Jürgen
Rare remains of predominantly deep-water sharks of the families Hexanchidae, Squalidae, Dalatiidae, Centrophoridae, and Squatinidae are described from the Eocene La Meseta Formation, Seymour Island, Antarctic Peninsula, which has yielded the most abundant chondrichthyan assemblage from the Southern Hemisphere to date. Previously described representatives of Hexanchus sp., Squalus weltoni, Squalus woodburnei, Centrophorus sp., and Squatina sp. are confirmed and dental variations are documented. Although the teeth of Squatina sp. differ from other Palaeogene squatinid species, we refrain from introducing a new species. A new dalatiid taxon, Eodalatias austrinalis gen. et sp. nov. is described. This new material not only increases the diversity of Eocene Antarctic elasmobranchs but also allows assuming that favourable deep-water habitats were available in the Eocene Antarctic Ocean off Antarctica in the Eocene. The occurrences of deep-water inhabitants in shallow, near-coastal waters of the Antarctic Peninsula agrees well with extant distribution patterns.
van den Biggelaar, D.F.A.M.; van Balen, R.T.; Kluiving, S.J.; Verpoorte, A.; Alink, A
The artefact size of the Early Middle Palaeolithic (EMP) assemblages in ice-pushed Rhine–Meuse deposits in the central Netherlands decreases northwestward. This trend correlates to the downstream fining direction of the Rhine–Meuse fluvial system, the source of the rock material, showing that
Adnet, S.; Antoine, P.-O.; Hassan Baqri, S. R.; Crochet, J.-Y.; Marivaux, L.; Welcomme, J.-L.; Métais, G.
New selachians (sharks and rays) have been collected from several late Eocene and early Oligocene marine localities in the Bugti Hills (Balochistan, Pakistan). Two new species of Requiem sharks (close to the Recent "Bull shark") are described : Carcharhinus balochensis and Carcharhinus perseus. The rest of the fauna is notable for the strong representation of Carcharhiniformes. These selachian faunas represent a unique tropical association for the Oligocene period and one of the first modern tropical selachian faunas, with modern taxa such as the two new species of "Bull sharks", Negaprion sp. and one of the first occurrences of Sphyrna sp. Moreover, these faunas permit paleoenvironmental interpretation of adjacent land masses. The relatively modern aspect of these faunas, compared with other contemporaneous and younger selachian associations from Atlantic and Mediterranean seas, suggests biogeographic isolation of selachian communities living in eastern and western parts of the Tethys before its final closure during the early-middle Miocene.
Tarby, Sara; Cermak, Daniel
Sara Tarby, Daniel Cermak-Sassenrath. Screamy Bird. Digital game. Kulturnatten 2016, Danish Science Ministry, Copenhagen, DK, Oct 14, 2016.......Sara Tarby, Daniel Cermak-Sassenrath. Screamy Bird. Digital game. Kulturnatten 2016, Danish Science Ministry, Copenhagen, DK, Oct 14, 2016....
Lenz, Olaf K.; Wilde, Volker; Riegel, Walter
The Palaeogene was the most recent greenhouse period on Earth. Especially for the Late Palaeocene and Early Eocene, several superimposed short-term hyperthermal events have been described, including extremes such as the Palaeocene-Eocene Thermal Maximum. Major faunal and floral turnovers in the marine and terrestrial realms were recorded in association with these events. High-resolution palynological analysis of the early Middle Eocene maar lake sediments at Messel, near Darmstadt, Germany, provides an insight into the dynamics of a climax vegetation during the Middle Eocene greenhouse climate in a time span without significant climatic excursions. Numerical techniques like detrended correspondence analysis and wavelet analysis have been applied to recognize cyclic fluctuations and long-term trends in the vegetation through a time interval of approximately 640 kyr. Based on the numerical zoning of the pollen diagram, three phases in the development of the vegetation may be distinguished. Throughout these phases, the climax vegetation did not change substantially in qualitative composition, but a trend towards noticeably less humid conditions probably in combination with a drop of the water level in the lake may be recognized. A shift in algal population from the freshwater dinoflagellate cyst Messelodinium thielepfeifferae to a dominance of Botryococcus in the uppermost part of the core is interpreted as a response to changes in acidity and nutrient availability within the lake. Time series analyses of pollen assemblages show that variations in the Milankovitch range of eccentricity, obliquity and precession can be distinguished. In addition, fluctuations in the sub-Milankovitch range are indicated. This demonstrates that floral changes during steady depositional conditions in the Middle Eocene of Messel were controlled by orbital forcing.
Soriano, Sylvain; Grün, Rainer; Marra, Fabrizio; Nomade, Sebastien; Pereira, Alison; Boschian, Giovanni; Pollarolo, Luca; Fang, Fang; Bahain, Jean-Jacques
We present here the results of a technological and typological analysis of the Acheulian and early Middle Paleolithic assemblages from Torre in Pietra (Latium, Italy) together with comparisons with the Acheulian small tools of Castel di Guido. The assemblages were never chronometrically dated before. We have now 40Ar/39Ar dates and ESR-U-series dates, within a geomorphological framework, which support correlations to marine isotope stages. The Acheulian (previously correlated to MIS 9) is now dated to MIS 10 while the Middle Paleolithic is dated to MIS 7. Lithic analyses are preceded by taphonomic evaluations. The Levallois method of the Middle Paleolithic assemblage is an innovation characterized by the production of thin flake blanks without cortex. In contrast, the small tool blanks of the Acheulian were either pebbles or thick flakes with some cortex. They provided a relatively easy manual prehension. The choice of Levallois thin flake blanks in the Middle Paleolithic assemblage suggest that the new technology is most likely related to the emergence of hafting. Accordingly, the oldest direct evidence of hafting technology is from the site of Campitello Quarry in Tuscany (Central Italy) where birch-bark tar, found on the proximal part of two flint flakes, is dated to the end of MIS 7. Nevertheless, a peculiar feature of the Middle Paleolithic at Torre in Pietra is the continuous presence of small tool blanks on pebbles and cores and on thick flake albeit at a much lower frequency than in the older Acheulian industries. The adoption of the new technology is thus characterized by innovation combined with a degree of stability. The persistence of these habits in spite of the introduction of an innovative technique underlies the importance of cultural transmission and conformity in the behavior of Neandertals. PMID:27525705
van den Bergh, Gerrit D; Kaifu, Yousuke; Kurniawan, Iwan; Kono, Reiko T; Brumm, Adam; Setiyabudi, Erick; Aziz, Fachroel; Morwood, Michael J
The evolutionary origin of Homo floresiensis, a diminutive hominin species previously known only by skeletal remains from Liang Bua in western Flores, Indonesia, has been intensively debated. It is a matter of controversy whether this primitive form, dated to the Late Pleistocene, evolved from early Asian Homo erectus and represents a unique and striking case of evolutionary reversal in hominin body and brain size within an insular environment. The alternative hypothesis is that H. floresiensis derived from an older, smaller-brained member of our genus, such as Homo habilis, or perhaps even late Australopithecus, signalling a hitherto undocumented dispersal of hominins from Africa into eastern Asia by two million years ago (2 Ma). Here we describe hominin fossils excavated in 2014 from an early Middle Pleistocene site (Mata Menge) in the So'a Basin of central Flores. These specimens comprise a mandible fragment and six isolated teeth belonging to at least three small-jawed and small-toothed individuals. Dating to ~0.7 Ma, these fossils now constitute the oldest hominin remains from Flores. The Mata Menge mandible and teeth are similar in dimensions and morphological characteristics to those of H. floresiensis from Liang Bua. The exception is the mandibular first molar, which retains a more primitive condition. Notably, the Mata Menge mandible and molar are even smaller in size than those of the two existing H. floresiensis individuals from Liang Bua. The Mata Menge fossils are derived compared with Australopithecus and H. habilis, and so tend to support the view that H. floresiensis is a dwarfed descendent of early Asian H. erectus. Our findings suggest that hominins on Flores had acquired extremely small body size and other morphological traits specific to H. floresiensis at an unexpectedly early time.
Abdul Aziz, H.; di Stefano, A.; Foresi, L. M.; Hilgen, Frederik J.; Iaccarino, S. M.; Kuiper, K. F.; Lirer, F.; Salvatorini, G.; Turco, E.
An integrated magneto-biostratigraphic framework is presented for Middle Miocene sediments of DSDP Site 372 located in the Western Mediterranean. Detailed biostratigraphic analysis shows a nearly complete sequence of early Middle Miocene calcareous plankton bioevents in the Mediterranean, including
Collinson, Margaret; Barke, Judith; van der Burgh, Johan; van Konijnenburg-van Cittert, Johanna; Pearce, Martin; Bujak, Jonathan; Brinkhuis, Henk
Azolla is a free-floating freshwater fern that is renowned for its rapid vegetative spread and invasive biology, being one of the world's fastest growing aquatic macrophytes. Two species of this plant have been shown to have bloomed and reproduced in enormous numbers in the latest Early to earliest Middle Eocene of the Arctic Ocean and North Sea based on samples from IODP cores from the Lomonosov Ridge (Arctic) and from outcrops in Denmark (Collinson et al 2009 a,b Review of Palaeobotany and Palynology 155,1-14; and doi:10.1016/j.revpalbo.2009.12.001). To determine the geographic and temporal extent of this Azolla phenomenon, and the spatial distribution of the different species, we have examined samples from 15 additional sites using material from ODP cores and commercial exploration wells. The sites range from the Sub-Arctic (Northern Alaska and Canadian Beaufort Mackenzie Basin) to the Nordic Seas (Norwegian-Greenland Sea and North Sea Basin). Our data show that the Azolla phenomenon involved at least three species. These are distinguished by characters of the megaspore apparatus (e.g. megaspore wall, floats, filosum) and the microspore massulae (e.g. glochidia fluke tips). The Lomonosov Ridge (Arctic) and Danish occurrences are monotypic but in other sites more than one species co-existed. The attachment to one another and the co-occurrence of megaspore apparatus and microspore massulae, combined with evidence that these spores were shed at the fully mature stage of their life cycle, shows that the Azolla remains were not transported over long distances, a fact which could not be assumed from isolated massula fragments alone. Our evidence, therefore, shows that Azolla plants grew on the ocean surfaces for approximately 1.2 million years (from 49.3 to 48.1 Ma) and that the Azolla phenomenon covered the area from Denmark northwards across the North Sea Basin and the whole of the Arctic and Nordic seas. Apparently, early Middle Eocene Northern Hemisphere middle
Fairchild, Dana M [Armour, SD
The bird guard provides a device to protect electrical insulators comprising a central shaft; a clamp attached to an end of the shaft to secure the device to a transmission tower; a top and bottom cover to shield transmission tower insulators; and bearings to allow the guard to rotate in order to frighten birds away from the insulators.
National Oceanic and Atmospheric Administration, Department of Commerce — This data set contains sensitive biological resource data for endangered waterbirds and passerine birds, migratory shorebirds and waterfowl, gulls and terns,...
Samanta, Arpita; Sarkar, Anindya
A warm greenhouse climate, punctuated by a series of rapid warming events (known as hyperthermals), is characteristic of the Late Paleocene to Early Eocene period. Rapid addition of 13C depleted carbon to the exogenic carbon cycle, in an otherwise overall higher atmospheric CO2 level, is thought to set off the hyperthermal events. For understanding the fate of ongoing global warming and response of the climate system and biota, researchers for past few decades are paying more attention to comprehend this climatic enigma. Existing proxies from the most distinct hyperthermal event i.e., PETM indicate that the mean annual sea surface temperature (MASST) was comparatively higher (by ˜8 ˚ C) at high latitude and to a lesser extent towards the equator. Apart from the prominent hyperthermal events the rest of the Early Eocene was significantly warmer and thought to be more equable compare to present. Terrestrial proxy records from the mid-latitude regions indicated that the Mean Annual Temperature (MAT) and Minimum Winter Temperature (MWT) was high, thus reducing the seasonality or difference between MWT and Maximum Summer Temperature (MST). In absence of proxy data from the low latitude region, a ≥40 ˚ C summer temperature was predicted assuming a mild Eocene temperature gradient of ˜0.4 ˚ C/ ˚ latitude and mid-latitude temperature data. Even question was raised about the existence of the tropical rain forest in such climatic extreme. Recent pollen census data, on contrary, suggest proliferation of the tropical rain forest during this climatic extreme. Important in this context is that there is a very few direct evidence of Late Paleocene-Early Eocene MAT and seasonality data from the low latitude/equatorial regions. To resolve this issue, oxygen and carbon isotope ratios of larger benthic foraminifera (Nummulites burdigalensis) were measured in laser based carbonate device attached with the Delta V advantage continuous flow stable isotope ratio mass spectrometer
Permanent link: http://www.ias.ac.in/article/fulltext/jess/124/06/1201-1221. Keywords. Mammalia; Rodentia; Chapattimyidae; Eocene; Subathu Formation; India. Abstract. Based on isolated upper cheek teeth, two new early Eocene rodents (Subathumys solanorius gen. et sp. nov. and Subathumys globulus gen. et sp. nov.) ...
Kiel, Steffen; Kahl, Wolf-Achim; Goedert, James L.
The bone-eating marine annelid Osedax consumes mainly whale bones on the deep-sea floor, but recent colonization experiments with cow bones and molecular age estimates suggesting a possible Cretaceous origin of Osedax indicate that this worm might be able grow on a wider range of substrates. The suggested Cretaceous origin was thought to imply that Osedax could colonize marine reptile or fish bones, but there is currently no evidence that Osedax consumes bones other than those of mammals. We provide the first evidence that Osedax was, and most likely still is, able to consume non-mammalian bones, namely bird bones. Borings resembling those produced by living Osedax were found in bones of early Oligocene marine flightless diving birds (family Plotopteridae). The species that produced these boreholes had a branching filiform root that grew to a length of at least 3 mm, and lived in densities of up to 40 individuals per square centimeter. The inclusion of bird bones into the diet of Osedax has interesting implications for the recent suggestion of a Cretaceous origin of this worm because marine birds have existed continuously since the Cretaceous. Bird bones could have enabled this worm to survive times in the Earth's history when large marine vertebrates other than fish were rare, specifically after the disappearance of large marine reptiles at the end-Cretaceous mass extinction event and before the rise of whales in the Eocene.
National Oceanic and Atmospheric Administration, Department of Commerce — This data set contains sensitive biological resource data for wading birds, shorebirds, waterfowl, diving birds, seabirds, passerine birds, gulls, and terns in...
... of pet birds. Because of the risk of avian influenza (bird flu), USDA restricts the importation of pet birds from ... or look dirty may be ill. Learn the signs of illness in a bird, which include appearing ...
CAROLINA ACOSTA HOSPITALECHE
Full Text Available Penguins have peculiar modifications in their skeletal anatomy as a consequence of their extremely specialized diving habit. Morphological specialization is particularly evident in the forelimb. However, the kinematics of the pectoral girdle appears to be key to the locomotion of penguins. Penguin scapulae have an unusual morphology among birds. Modern penguins have a very large (especially broad scapula, whereas this bone is long but narrower in basal fossil species. The recent finding of an incomplete scapula with a singular acromion in the Upper Eocene Submeseta Allomember of the La Meseta Formation in the Antarctic Peninsula reveals a scapula proportionally narrower than those of modern penguins but similar to that of Waimanu and possibly other Eocene species. Osteological comparisons and muscular dissections of modern penguins show that the most striking feature is the curvature of the acromion, and the consequent enlargement of the facies articularis clavicularis. The configuration of the acromion and the corpus scapula reflects a lack of functional optimization in terms of the resistance to forces transverse to the body axis. The scapula´s general morphology suggests it belonged to a medium to large-sized penguin species with no so specialized diving skills.
The palaeoclimatic significance of Eurasian Giant Salamanders (Cryptobranchidae: Zaissanurus, Andrias) - indications for elevated humidity in Central Asia during global warm periods (Eocene, late Oligocene warming, Miocene Climate Optimum)
Vasilyan, Davit; Böhme, Madelaine; Winklhofer, Michael
Cryptobranchids represent a group of large sized (up to 1.8 m) tailed amphibians known since the Middle Jurassic (Gao & Shubin 2003). Two species are living today in eastern Eurasia: Andrias davidianus (China) and A. japonicus (Japan). Cenozoic Eurasian fossil giant salamanders are known with two genera and two or three species from over 30 localities, ranging from the Late Eocene to the Early Pliocene (Böhme & Ilg 2003). The Late Eocene species Zaissanurus beliajevae is restricted to the Central Asian Zaissan Basin (SE-Kazakhstan, 50°N, 85°E), whereas the Late Oligocene to Early Pliocene species Andrias scheuchzeri is distributed from Central Europe to the Zaissan Basin. In the latter basin the species occur during two periods; the latest Oligocene and the late Early to early Middle Miocene (Chkhikvadse 1982). Andrias scheuchzeri is osteological indistinguishable from both recent species, indicating a similar ecology (Westfahl 1958). To investigate the palaeoclimatic significance of giant salamanders we analyzed the climate within the present-day distribution area and at selected fossil localities with independent palaeoclimate record. Our results indicate that fossil and recent Andrias species occur in humid areas where the mean annual precipitation reach over 900 mm (900 - 1.300 mm). As a working hypothesis (assuming a similar ecology of Andrias and Zaissanurus) we interpret occurrences of both fossil Eurasian giant salamanders as indicative for humid palaeoclimatic conditions. Based on this assumption the Late Eocene, the latest Oligocene (late Oligocene warming) and the late Early to early Middle Miocene (Miocene Climatic Optimum) of Central Asia (Zaissan Basin) are periods of elevated humidity, suggesting a direct (positive) relationship between global climate and Central Asian humidity evolution. Böhme M., Ilg A. 2003: fosFARbase, www.wahre-staerke.com/ Chkhikvadze V.M. 1982. On the finding of fossil Cryptobranchidae in the USSR and Mongolia. Vertebrata
Thomas, Ellen; D’haenens, Simon; Speijer, Robert P.; Alegret, Laia
The early Eocene greenhouse world was marked by multiple transient hyperthermal events. The most extreme was the Paleocene-Eocene Thermal Maximum (PETM, ~56 Ma), linked to the extinction of the globally recognised deep-sea benthic foraminiferal Velasco fauna, which led to the development of early Eocene assemblages. This turnover has been studied at high resolution, but faunal development into the later early Eocene is poorly documented. There is no widely accepted early Eocene equivalent of the Late Cretaceous-Paleocene Velasco fauna, mainly due to the use of different taxonomic concepts. We compiled Ypresian benthic foraminiferal data from 17 middle bathyal-lower abyssal ocean drilling sites in the Pacific, Atlantic and Indian Oceans, in order to characterise early Eocene deep-sea faunas by comparing assemblages across space, paleodepth and time. Nuttallides truempyi, Oridorsalis umbonatus, Bulimina trinitatensis, the Bulimina simplex group, the Anomalinoides spissiformis group, pleurostomellids, uniserial lagenids, stilostomellids and lenticulinids were ubiquitous during the early Eocene (lower-middle Ypresian). Aragonia aragonensis, the Globocassidulina subglobosa group, the Cibicidoides eocaenus group and polymorphinids became ubiquitous during the middle Ypresian. The most abundant early Ypresian taxa were tolerant to stressed or disturbed environments, either by opportunistic behavior (Quadrimorphina profunda, Tappanina selmensis, Siphogenerinoides brevispinosa) and/or the ability to calcify in carbonate-corrosive waters (N. truempyi). Nuttallides truempyi, T. selmensis and other buliminids (Bolivinoides cf. decoratus group, Bulimina virginiana) were markedly abundant during the middle Ypresian. Contrary to the long-lived, highly diverse and equitable Velasco fauna, common and abundant taxa reflect highly perturbed assemblages through the earliest Ypresian, with lower diversity and equitability following the PETM extinction. In contrast, the middle Ypresian
Philip D Gingerich
Full Text Available BACKGROUND: Protocetidae are middle Eocene (49-37 Ma archaeocete predators ancestral to later whales. They are found in marine sedimentary rocks, but retain four legs and were not yet fully aquatic. Protocetids have been interpreted as amphibious, feeding in the sea but returning to land to rest. METHODOLOGY/PRINCIPAL FINDINGS: Two adult skeletons of a new 2.6 meter long protocetid, Maiacetus inuus, are described from the early middle Eocene Habib Rahi Formation of Pakistan. M. inuus differs from contemporary archaic whales in having a fused mandibular symphysis, distinctive astragalus bones in the ankle, and a less hind-limb dominated postcranial skeleton. One adult skeleton is female and bears the skull and partial skeleton of a single large near-term fetus. The fetal skeleton is positioned for head-first delivery, which typifies land mammals but not extant whales, evidence that birth took place on land. The fetal skeleton has permanent first molars well mineralized, which indicates precocial development at birth. Precocial development, with attendant size and mobility, were as critical for survival of a neonate at the land-sea interface in the Eocene as they are today. The second adult skeleton is the most complete known for a protocetid. The vertebral column, preserved in articulation, has 7 cervicals, 13 thoracics, 6 lumbars, 4 sacrals, and 21 caudals. All four limbs are preserved with hands and feet. This adult is 12% larger in linear dimensions than the female skeleton, on average, has canine teeth that are 20% larger, and is interpreted as male. Moderate sexual dimorphism indicates limited male-male competition during breeding, which in turn suggests little aggregation of food or shelter in the environment inhabited by protocetids. CONCLUSIONS/SIGNIFICANCE: Discovery of a near-term fetus positioned for head-first delivery provides important evidence that early protocetid whales gave birth on land. This is consistent with skeletal
Ding, Dianshi; Liu, Guijian; Sun, Xiaohui; Sun, Ruoyu
To investigate the magnitude to which the carbon isotopic ratio (δ13C) varies in coals in response to their contemporary terrestrial environment, the Early-Middle Permian Huainan coals (including coals from the Shanxi Formation, Lower Shihezi Formation and Upper Shihezi Formation) in North China were systematically sampled. A 2.5‰ variation range of δ13C values (-25.15‰ to -22.65‰) was observed in Huainan coals, with an average value of -24.06‰. As coal diagenesis exerts little influence on carbon isotope fractionation, δ13C values in coals were mainly imparted by those of coal-forming flora assemblages which were linked to the contemporary climate. The δ13C values in coals from the Shanxi and Lower Shihezi Formations are variable, reflecting unstable climatic oscillations. Heavy carbon isotope is enriched in coals of the Capitanian Upper Shihezi Formation, implying a shift to high positive δ13C values of coeval atmospheric CO2. Notably, our study provides evidence of the Kamura event in the terrestrial environment for the first time.
Sánchez Yustos, Policarpo; Diez Martín, Fernando
The Northwest of Iberia has yielded one of the most complete European Middle Paleolithic records. Despite this wealth of information, very little is known about population dynamics during this period. For that reason, the main concern of this paper is to provide socio-environmental models that may help explain Early Middle Paleolithic (EMP) population dynamics in NW Iberia, assessing to what extent they were shaped by climate forces. The archaeological record is analyzed on the basis of the heuristics of ecological models, already employed in the European Pleistocene record but never at a regional scale, in order to detect long-term changes in the composition of EMP populations, and the environmental, biological and sociocultural process influencing those changes. According to the models proposed, we have detected a long-term population dynamic between MIS 11 and MIS 6, characterized by low environmental stress, high biological productivity, interaction among populations and sociocultural complexity. Eventually, this population dynamic was broken due to an extreme climate phase in late MIS 6 that had a profound impact on populations and sociocultural structures. As a result, the Upper Pleistocene population of NW Iberia was concentrated in the Cantabrian region. This area became an isolated Neanderthal glacial refugium that hosted a population with different origins and fragile long-term demographic stability.
... 9 Animals and Animal Products 1 2010-01-01 2010-01-01 false Certificate for pet birds, commercial birds, zoological birds, and research birds. 93.104 Section 93.104 Animals and Animal Products ANIMAL... (INCLUDING POULTRY) AND ANIMAL PRODUCTS IMPORTATION OF CERTAIN ANIMALS, BIRDS, FISH, AND POULTRY, AND CERTAIN...
Ruslanova Rida Raisovna
Full Text Available Early Middle Ages in the Southern Urals is the time of the tumultuous ethnocultural processes, that is an echo of the era of the Great Migration. At this time, the bakhmutinskaya culture was formed (3rd-8th centuries A.D.. The Birsk burial ground is one of the unique monuments of this period – it appeared in the second third of the 1st millennium B.C. The Birsk burial ground is a fiducial monument for studying history, ethno-cultural, migration and trade processes occurring in the Southern Urals, and the content in the composition of grave goods makes it supplies an important source in the study of early medieval history of East European forest. A variety of types of beads from the Birsk burial ground allows suggesting that the necropolis was one of the major points on the caravan trade and exchange path. According to it, the exchange could take place on imports of products (furs, honey, metals. The article describes a set of beads from the Birsk burials – evidence of a monument in the system of early medieval antiquities (3rd-8th centuries A.D.. The complex morpho-technological research dealt with 218 complexes containing 6705 instances of beads and jewelry. The feature of the monument is the presence of necklaces jewelry from all the selected materials along with the material. The Birsk burial ground demonstrates various forms of products, colors used glass for monochrome and polychrome decorations. The presented work can be used in the study of material culture and trade exchange operations of the medieval population of the Urals.
Full Text Available In this paper we examine the relation between the loss of formal gender and Case features on simple demonstratives and the topic shifting property they manifest. The examination period spans between Old English and Early Middle English. While we argue that this loss has important discourse-pragmatic and derivational effects on demonstratives, we also employ the Strong Minimalist Hypothesis approach (Chomsky 2001 and feature valuation, as defined in Pesetsky & Torrego (2007, to display how their syntactic computation and pragmatic properties have come about. To account for the above innovations yielding the Early Middle English ϸe (‘the’, we first discuss the formal properties of the Old English demonstratives which distinguish number, gender, and Case features. This inflectional variety of forms allows the Old English demonstratives to be used independently and to show the anaphoric and discourse-linking properties of topics. Crucially, the same properties characterise also German and Dutch demonstratives that manifest Case and/or gender morphology overtly, which shows that the syntactic distribution of LIs and their morphological richness should be considered as intertwined. The above properties are then confronted with the determiner system in Early Middle English, whose forms undergo inflectional levelling producing the invariant ϸe/ðe form that loses its distributional independence and acquires the article status. The levelling process in question is argued to stimulate the shift of the [+ref/spec] feature from the formal to the semantic pole. This suggests that the Early Middle English ϸe form no longer counts as an appropriate anaphor in topic shift contexts owing to its indeterminacy of Case, gender, and φ-features, which means that it cannot satisfy the Full Interpretation requirement at the interfaces.
Bourisly, Ali K
The aim of this study was to identify age-related changes of P300 peak amplitude and P300 latency between closely separated nonsenile age groups (late young-aged adults and early middle-aged adults) and to investigate whether or not P300 has the potential to be used as a measure of cognitive aging even among nonsenile age groups. Twenty-eight adults (25-55 years old) completed an event-related potential oddball task. The elicitation of both P300 peak amplitude and P300 latency indicated age-related changes of P300. The results of the study showed that the P300 target peak amplitude was significantly larger in late young age compared with early middle age and that P300 target latency was also significantly delayed in early middle age compared with late young age. The results of this work contribute toward research efforts on a consensus on how aging affects event-related potential and/or P300. The main conclusions are that there exist significant age-related P300 changes even between closely separated, relatively younger, and nonsenile age groups, and that P300 has the potential to be used as a measure for cognitive aging even in nonsenile adults.
Beraldi-Campesi, Hugo; Cevallos-Ferriz, Sergio R. S.; Centeno-García, Elena; Arenas-Abad, Concepción; Fernández, Luis Pedro
A depositional model of the Eocene-Oligocene Coatzingo Formation in Tepexi de Rodríguez (Puebla, Mexico) is proposed, based on facies analysis of one of the best-preserved sections, the Axamilpa Section. The sedimentary evolution is interpreted as the retrogradation of an alluvial system, followed by the progressive expansion of an alkaline lake system, with deltaic, palustrine, and evaporitic environments. The analysis suggests a change towards more arid conditions with time. Fossils from this region, such as fossil tracks of artiodactyls, aquatic birds and cat-like mammals, suggest that these animals traversed the area, ostracods populated the lake waters, and plants grew on incipient soils and riparian environments many times throughout the history of the basin. The inferred habitat for some fossil plants coincides with the sedimentological interpretation of an arid to semiarid climate for that epoch. This combined sedimentological-paleontological study of the Axamilpa Section provides an environmental context in which fossils can be placed and brings into attention important biotic episodes, like bird and camelid migrations or the origin of endemic but extinct plants in this area.
Full Text Available One new genus and three new species of Lygistorrhinidae in early Eocene Cambay amber from India are described, which significantly increases our knowledge about this group in the Eocene. Lygistorrhina indica n. sp. is the oldest fossil known from this extant genus. Indorrhina sahnii n. gen. et sp. shows morphological similarities to each of the two extant genera Lygistorrhina and Asiorrhina. Palaeognoriste orientale is the third species known from a group that has only been recorded from Eocene Baltic amber before. The latter finding reveals faunal links between Cambay amber and the probably slightly younger Baltic amber, adding further evidence that faunal exchange between Europe/Asia and India took place before the formation of Cambay amber.
Lyu, X.; Liu, Z.
The formation of oceanic red beds that usually present oxic and oligotrophic conditions with low sedimentation rate has been used to trace depositional paleoenvironment and paleoclimate change. Red beds overlying oceanic basalts were drilled at two adjacent Sites U1433 and U1434 of IODP Expedition 349 in the Southwest Subbasin of the South China Sea. The occurrence of early-middle Miocene red beds may indicate that at that time there was oxic and quiet marine environment in the deep South China Sea. To understand their formation of red-color, local depositional condition, and potential paleoceanographic significance, major elements (XRF), trace and rare earth elements (ICP-MS), Fe chemical speciation (modified sequential iron extraction procedure), and Fe oxic minerals (CBD and DRS) were analyzed. Geochemical and mineralogical data reveal that hematite and goethite are responsible for the reddish color and red beds were deposited under highly oxic, oligotrophic conditions with a little later hydrothermal influence in the South China Sea. Our results indicate that: (1) after treatment using the CBD procedure, the red samples presented a change in color to greenish, showing the iron oxides being responsible for the sediment color; (2) enriched Mn, depleted U, S enrichment factors, and negative Ce anomaly show that the water mass was pre-oxidized before transported to the study location; (3) low primary productivity was inferred from the lower P, Ba enrichment factors in red beds compared to non-red beds; (4) the excess Mo influx at the bottom may come from the later hydrothermal input; (5) the diverse Ca enrichment factors and correlations between Fe and Al suggest different allogenic sources for red beds at our two sites. We conclude that the red beds at Sites U1433 and U1434 despite their diverse sources both developed in externally oxidized water mass and low primary productivity conditions, and partially altered by hydrothermal fluids after their pelagic
Slotnick, B.S.; Dickens, G.R.; Nicolo, M.J.; Hollis, C.J.; Crampton, J.S.; Strong, C.P.; Zachos, J.C.; Sluijs, A.; Lourens, L.; Lauretano, V.
Global temperatures rose ~6°C from the late Paleocene ca. 58 Ma to the Early Eocene Climatic Optimum (EECO) ca. 52–50 Ma. Superimposed, were certainly two geologically brief (< 200 kyr) intervals of extreme warming, the Paleocene-Eocene thermal maximum (PETM) and Eocene thermal maximum 2 (ETM-2
Harrington, Carolyn Lang
Describes a lesson for third-grade students that begins with an examination of bird prints done by John James Audubon and moves into the students creating their own torn paper birds. Introduces the students to the beauty of birds and focuses on the environmental issues that face birds and their habitats. (CMK)
Přikryl, Tomáš; Vodrážka, R.
Roč. 34, č. 4 (2012), s. 895-908 ISSN 1280-9659 Institutional research plan: CEZ:AV0Z30130516 Institutional support: RVO:67985831 Keywords : Teleostei * scales * lepidology * Eocene * La Meseta Formation Subject RIV: DB - Geology ; Mineralogy Impact factor: 0.927, year: 2012
characteristic of warm tropical climate suggesting the prevalence of such climate during early Eocene in northeast India. 1. Introduction ... family (Dutta et al. 2009, 2011b). These are also known as dammar resins and have been classified as Class II resins (Anderson et al. 1992; Dutta et al. 2009). These have been reported ...
M.L.J. Baatsen; A.S. von der Heydt; M. Kliphuis; J.P. Viebahn (Jan); H.A. Dijkstra
htmlabstractThe Eocene-Oligocene Transition (EOT) marks a major step within the Cenozoic climate in going from a greenhouse into an icehouse state, with the formation of a continental-scale Antarctic ice sheet. The roles of steadily decreasing CO2 concentrations versus changes in ocean
Smith, Thierry; Rana, Rajendra S.; Missiaen, Pieter; Rose, Kenneth D.; Sahni, Ashok; Singh, Hukam; Singh, Lachham
The geographic origin of bats is still unknown, and fossils of earliest bats are rare and poorly diversified, with, maybe, the exception of Europe. The earliest bats are recorded from the Early Eocene of North America, Europe, North Africa and Australia where they seem to appear suddenly and simultaneously. Until now, the oldest record in Asia was from the Middle Eocene. In this paper, we report the discovery of the oldest bat fauna of Asia dating from the Early Eocene of the Cambay Formation at Vastan Lignite Mine in Western India. The fossil taxa are described on the basis of well-preserved fragments of dentaries and lower teeth. The fauna is highly diversified and is represented by seven species belonging to seven genera and at least four families. Two genera and five species are new. Three species exhibit very primitive dental characters, whereas four others indicate more advanced states. Unexpectedly, this fauna presents strong affinities with the European faunas from the French Paris Basin and the German Messel locality. This could result from the limited fossil record of bats in Asia, but could also suggest new palaeobiogeographic scenarios involving the relative position of India during the Early Eocene.
) witnessed a series of short-lived hyperthermal events (Paleocene–Eocene thermal maximum (PETM), H1/ETM2/ELMO, H2, I1, and I2), superimposed on the long-term global warming trend (Cramer et al. 2003; Nicolo et al. 2007; Zachos et al.
Hoeken-Klinkenberg, van P.M.J.
In order to continue the investigation of the Nigerian Maastrichtian pollenflora (van Hoeken-Klinkenberg, 1964) with Lower Tertiary material, a study has been made of some boreholesamples of the Maastrichtian, Paleocene and Eocene in Nigeria. A comparison is made with the South American pollen and
S. J. Gibbs
Full Text Available Late Paleocene and early Eocene hyperthermals are transient warming events associated with massive perturbations of the global carbon cycle, and are considered partial analogues for current anthropogenic climate change. Because the magnitude of carbon release varied between the events, they are natural experiments ideal for exploring the relationship between carbon cycle perturbations, climate change and biotic response. Here we quantify marine biotic variability through three million years of the early Eocene that include five hyperthermals, utilizing a method that allows us to integrate the records of different plankton groups through scenarios ranging from background to major extinction events. Our long time-series calcareous nannoplankton record indicates a scaling of biotic disruption to climate change associated with the amount of carbon released during the various hyperthermals. Critically, only the three largest hyperthermals, the Paleocene–Eocene Thermal Maximum (PETM, Eocene Thermal Maximum 2 (ETM2 and the I1 event, show above-background variance, suggesting that the magnitude of carbon input and associated climate change needs to surpass a threshold value to cause significant biotic disruption.
Boulila, Slah; Vahlenkamp, Maximilian; De Vleeschouwer, David; Laskar, Jacques; Yamamoto, Yuhji; Pälike, Heiko; Kirtland Turner, Sandra; Sexton, Philip F.; Westerhold, Thomas; Röhl, Ursula
Until now, the middle Eocene has remained a poorly constrained interval of efforts to produce an astrochronological timescale for the entire Cenozoic. This has given rise to a so-called "Eocene astronomical timescale gap" (Vandenberghe et al., 2012). A high-resolution astrochronological calibration for this interval has proven to be difficult to realize, mainly because carbonate-rich deep-marine sequences of this age are scarce. In this paper, we present records from middle Eocene carbonate-rich sequences from the North Atlantic Southeast Newfoundland Ridge (IODP Exp. 342, Sites U1408 and U1410), of which the cyclical sedimentary patterns allow for an orbital calibration of the geologic timescale between ∼38 and ∼48 Ma. These carbonate-rich cyclic sediments at Sites U1408 and U1410 were deposited as drift deposits and exhibit prominent lithological alternations (couplets) between greenish nannofossil-rich clay and white nannofossil ooze. The principal lithological couplet is driven by the obliquity of Earth's axial tilt, and the intensity of their expression is modulated by a cyclicity of about 173 kyr. This cyclicity corresponds to the interference of secular frequencies s3 and s6 (related to the precession of nodes of the Earth and Saturn, respectively). This 173-kyr obliquity amplitude modulation cycle is exceptionally well recorded in the XRF (X-ray fluorescence)-derived Ca/Fe ratio. In this work, we first demonstrate the stability of the (s3-s6) cycles using the latest astronomical solutions. Results show that this orbital component is stable back to at least 50 Ma, and can thus serve as a powerful geochronometer in the mid-Eocene portion of the Cenozoic timescale. We then exploit this potential by calibrating the geochronology of the recovered middle Eocene timescale between magnetic polarity Chrons C18n.1n and C21n. Comparison with previous timescales shows similarities, but also notable differences in durations of certain magnetic polarity chrons. We
Bijl, Peter K; Bendle, James A P; Bohaty, Steven M; Pross, Jörg; Schouten, Stefan; Tauxe, Lisa; Stickley, Catherine E; McKay, Robert M; Röhl, Ursula; Olney, Matthew; Sluijs, Appy; Escutia, Carlota; Brinkhuis, Henk
The warmest global temperatures of the past 85 million years occurred during a prolonged greenhouse episode known as the Early Eocene Climatic Optimum (52-50 Ma). The Early Eocene Climatic Optimum terminated with a long-term cooling trend that culminated in continental-scale glaciation of Antarctica from 34 Ma onward. Whereas early studies attributed the Eocene transition from greenhouse to icehouse climates to the tectonic opening of Southern Ocean gateways, more recent investigations invoked a dominant role of declining atmospheric greenhouse gas concentrations (e.g., CO2). However, the scarcity of field data has prevented empirical evaluation of these hypotheses. We present marine microfossil and organic geochemical records spanning the early-to-middle Eocene transition from the Wilkes Land Margin, East Antarctica. Dinoflagellate biogeography and sea surface temperature paleothermometry reveal that the earliest throughflow of a westbound Antarctic Counter Current began ~49-50 Ma through a southern opening of the Tasmanian Gateway. This early opening occurs in conjunction with the simultaneous onset of regional surface water and continental cooling (2-4 °C), evidenced by biomarker- and pollen-based paleothermometry. We interpret that the westbound flowing current flow across the Tasmanian Gateway resulted in cooling of Antarctic surface waters and coasts, which was conveyed to global intermediate waters through invigorated deep convection in southern high latitudes. Although atmospheric CO2 forcing alone would provide a more uniform middle Eocene cooling, the opening of the Tasmanian Gateway better explains Southern Ocean surface water and global deep ocean cooling in the apparent absence of (sub-) equatorial cooling.
Eberle, J. J.; Fricke, H. C.; Humphrey, J.; Hackett, L.; Newbrey, M.; Hutchison, H.
As a deep time analog for today’s rapidly warming Arctic region, early Eocene (~53 Ma) rocks on Ellesmere Island, Arctic Canada (~79° N.) preserve evidence of lush swamp forests inhabited by turtles, alligators, primates, tapirs, and hippo-like Coryphodon. Although the rich flora and fauna of the early Eocene Arctic imply warmer, wetter conditions that at present, quantitative estimates of Eocene Arctic climate are rare. By analyzing oxygen isotope ratios of biogenic phosphate from mammal, fish, and turtle fossils from a single locality on central Ellesmere Island, we provide estimates of early Eocene Arctic temperature, including mean annual temperature (MAT) of ~ 8° C, mean annual range in temperature (MART) of ~ 16.5° C, warm month mean temperature (WMMT) of 16 - 19° C, and cold month mean temperature (CMMT) of 0 - 1° C. Our seasonal range in temperature is similar to the range in estimated MAT obtained using different proxies. In particular, unusually high estimates of early Eocene Arctic MAT and sea surface temperature (SST) by others that are based upon the distribution of branched glycerol dialkyl glycerol tetraether (GDGT) membrane lipids in terrestrial soil bacteria and marine Crenarchaeota fall within our range of WMMT, suggesting a bias towards summer values. Consequently, caution should be taken when using these methods to infer MAT and SST that, in turn, are used to constrain climate models. From a paleontologic perspective, our temperature estimates verify that alligators and tortoises, by way of nearest living relative-based climatic inference, are viable paleoclimate proxies for mild, above-freezing year-round temperatures. Although in both of these reptiles, past temperature tolerances were greater than in their living descendants.
Bijl, Peter K.; Bendle, James A. P.; Bohaty, Steven M.; Pross, Jörg; Schouten, Stefan; Tauxe, Lisa; Stickley, Catherine E.; McKay, Robert M.; Röhl, Ursula; Olney, Matthew; Sluijs, Appy; Escutia, Carlota; Brinkhuis, Henk; Klaus, Adam; Fehr, Annick; Williams, Trevor; Carr, Stephanie A.; Dunbar, Robert B.; Gonzàlez, Jhon J.; Hayden, Travis G.; Iwai, Masao; Jimenez-Espejo, Francisco J.; Katsuki, Kota; Kong, Gee Soo; Nakai, Mutsumi; Passchier, Sandra; Pekar, Stephen F.; Riesselman, Christina; Sakai, Toyosaburo; Shrivastava, Prakash K.; Sugisaki, Saiko; Tuo, Shouting; van de Flierdt, Tina; Welsh, Kevin; Yamane, Masako
The warmest global temperatures of the past 85 million years occurred during a prolonged greenhouse episode known as the Early Eocene Climatic Optimum (52–50 Ma). The Early Eocene Climatic Optimum terminated with a long-term cooling trend that culminated in continental-scale glaciation of Antarctica from 34 Ma onward. Whereas early studies attributed the Eocene transition from greenhouse to icehouse climates to the tectonic opening of Southern Ocean gateways, more recent investigations invoked a dominant role of declining atmospheric greenhouse gas concentrations (e.g., CO2). However, the scarcity of field data has prevented empirical evaluation of these hypotheses. We present marine microfossil and organic geochemical records spanning the early-to-middle Eocene transition from the Wilkes Land Margin, East Antarctica. Dinoflagellate biogeography and sea surface temperature paleothermometry reveal that the earliest throughflow of a westbound Antarctic Counter Current began ∼49–50 Ma through a southern opening of the Tasmanian Gateway. This early opening occurs in conjunction with the simultaneous onset of regional surface water and continental cooling (2–4 °C), evidenced by biomarker- and pollen-based paleothermometry. We interpret that the westbound flowing current flow across the Tasmanian Gateway resulted in cooling of Antarctic surface waters and coasts, which was conveyed to global intermediate waters through invigorated deep convection in southern high latitudes. Although atmospheric CO2 forcing alone would provide a more uniform middle Eocene cooling, the opening of the Tasmanian Gateway better explains Southern Ocean surface water and global deep ocean cooling in the apparent absence of (sub-) equatorial cooling. PMID:23720311
Full Text Available Three new or from this site previously unrecorded birds are described from the Middle Eocene of Messel (Hessen, Germany. Serudaptus pohli n. gen. n. sp. is a new taxon of the Pseudasturidae and is distinguished from the other species of this family by its short and stout tarsometatarsus and the long and strong claws. An isolated foot of a gruiform bird is assigned to Idiornis cf. itardiensis (Idiornithidae and would, if this assignment can be confirmed, demonstrate the existence of this species for a period of 23 million years. The proportions of the toes suggest that Idiornis cf. itardiensis was less cursorial than its next recent relatives, the Cariamidae. In addition thereto, an exceptionally well preserved skeleton of a sandcoleid bird (Coliiformes: Sandcoleidae gives new information on the osteology and feathering of sandcoleid birds. The specimen resembles the genera Eoglaucidium and Anneavis but it has not been possible to assign it to one of these two genera with certainty. Drei neue oder von dieser Fundstelle bisher nicht nachgewiesene Vögel werden aus dem Mittel-Eozän von Messel (Hessen, Deutschland beschrieben. Serudaptus pohli n. gen. n. sp. ist ein neues Taxon der Pseudasturidae und unterscheidet sich von den anderen Arten dieser Familie durch den kurzen und gedrungenen Tarsometatarsus und die langen, kräftigen Krallen. Der isolierte Fuß eines gruiformen Vogels wird Idiornis cf. itardiensis (Idiornithidae zugeordnet und würde, wenn sich diese Zuordnung bestätigt, die Existenz dieser Art über einen Zeitraum von 23 Millionen Jahren zeigen. Die Zehenproportionen legen nahe, dass Idiornis cf. itardiensis weniger an eine laufende Lebensweise angepasst war als seine nächsten rezenten Verwandten, die Cariamidae. Darüber hinaus zeigt ein außergewöhnlich gut erhaltenes Skelett eines Vertreters der Sandcoleidae bisher unbekannte Einzelheiten der Osteologie und Befiederung dieser Familie. Das Exemplar ähnelt den Gattungen
Spicer, Robert A.; Yang, Jian; Herman, Alexei B.; Kodrul, Tatiana; Maslova, Natalia; Spicer, Teresa E. V.; Aleksandrova, Galina; Jin, Jianhua
The onset and development of the Asian monsoon systems is a topic that has attracted considerable research effort but proxy data limitations, coupled with a diversity of definitions and metrics characterizing monsoon phenomena, have generated much debate. Failure of geological proxies to yield metrics capable of distinguishing between rainfall seasonality induced by migrations of the Inter-tropical Convergence Zone (ITCZ) from that attributable to topographically modified seasonal pressure reversals has frustrated attempts to understand mechanisms underpinning monsoon development and dynamics. Here we circumvent the use of such single climate parameter metrics in favor of detecting directly the distinctive attributes of different monsoon regimes encoded in leaf fossils. Leaf form adapts to the prevailing climate, particularly under the extreme seasonal stresses imposed by monsoons, so it is likely that fossil leaves carry a unique signature of past monsoon regimes. Leaf form trait spectra obtained from fossils from Eocene basins in southern China were compared with those seen in modern leaves growing under known climate regimes. The fossil leaf trait spectra, including those derived from previously published fossil floras from northwestern India, were most similar to those found in vegetation exposed to the modern Indonesia-Australia Monsoon (I-AM), which is largely a product of seasonal migrations of the ITCZ. The presence of this distinctive leaf physiognomic signature suggests that although a monsoon climate existed in Eocene time across southern Asia the characteristics of the modern topographically-enhanced South Asia Monsoon had yet to develop. By the Eocene leaves in South Asia had become well adapted to an I-AM type regime across many taxa and points to the existence of a pervasive monsoon climate prior to the Eocene. No fossil trait spectra typical of exposure to the modern East Asia monsoon were seen, suggesting the effects of this system in southern
Winguth, A. M. E.; Hughlett, T. M.; Brown, M.; Rothstein, M.; Shields, C. A.; Winguth, C.
A series of DeepMIP climate sensitivity experiments have been carried out with the Community Earth System Model CESM1.2 to evaluate how changes in the radiative forcing could have contributed to explain Eocene hyperthermal events. A rise in Eocene greenhouse gas forcing could have been linked to an increase in volcanism and associated destabilization of marine carbon reservoirs by dissociation of clathrathes, reorganization of the marine microbial loop, or terrestrial sources from e.g. wetlands. Such environmental changes could potentially have led to additional biophysical feedbacks altering the cloud aerosol optical depth for example by alteration of marine plankton productivity and DMS emissions to the atmosphere. The analysis of our simulations suggests a substantial warming from 3x to 12x CO2 PAL, reaching moderate temperatures of up to 20 °C over Antarctica and in the Article realm in the most extreme scenario, consistent to proxy estimates in a high CO2 world. The lower equator-to-pole temperature gradient compared to present-day is due to the lack of an ice sheet, an increase in greenhouse gases, and a lower cloud optical depth. The climate simulations suggest an intensified hydrological cycle with higher precipitation in the tropics, particularly over the Indian Eocene continent, and in mid-latitudes, whereas mega-droughts are prominent in the subtropics, particularly in Africa and South America. The Eocene geography (the closure of the Drake Passage and the more southern location of Australia) and a lower-than-present meridional temperature gradient contribute to a much weaker surface ocean circulation near the Antarctic continent as compared to the current pronounced Antarctic Circumpolar Current.
Baatsen, M. L. J.; von der Heydt, A. S.; Kliphuis, M.; Viebahn, J.; Dijkstra, H. A.
The Eocene-Oligocene Transition (EOT) marks a major step within the Cenozoic climate in going from a greenhouse into an icehouse state, with the formation of a continental-scale Antarctic ice sheet. The roles of steadily decreasing CO2 concentrations versus changes in ocean circulation at the EOT are still debated and the threshold for Antarctic glaciation is obscured by uncertainties in global geometry. Here, a detailed study of the late Eocene ocean circulation is carried out using an ocean general circulation model under two slightly different geography reconstructions of the middle-to-late Eocene (38 Ma). Using the same atmospheric forcing, both geographies give a profoundly different equilibrium ocean circulation state. The underlying reason for this sensitivity is the presence of multiple equilibria characterised by either North or South Pacific deep water formation. A possible shift from a southern towards a northern overturning circulation would result in significant changes in the global heat distribution and consequently make the Southern Hemisphere climate more susceptible for significant cooling and ice sheet formation on Antarctica.
The plant fossil record for the neotropics is still sparse and temporally discontinuous. The location and description of new fossil material are fundamental for understanding evolutionary and biogeographic patterns of lineages. A new fossil record of Passifloraceae from the late Eocene of Colombia is described in this study. Plant fossils were collected from a new locality from the Eocene Esmeraldas Formation. Eighteen fossil seeds were selected, described, and compared with fossil and extant angiosperm seeds based on the literature and herbarium collections. Taxonomic affinities of the fossil seeds within Passifloraceae s.l. were evaluated by comparing morphological characters of the seeds in a phylogenetic context. Stratigraphic information associated with the fossil locality was used to interpret the environment and taphonomic processes associated with fossil deposition. A new seed fossil genus and species, Passifloroidesperma sogamosense gen. and sp. nov., is described and associated with the subfamily Passifloroideae based on the presence of a foveolate seed surface, ruminate endosperm, and a seed coat with prismatic palisade cells. The depositional environment of the locality is described as a floodplain associated with river channels. A detailed review of the Passifloraceae fossil record indicates that P. sogamosense is the oldest confirmed record of Passifloraceae. Its late Eocene age provides a minimum age that can be used as a calibration point for the crown Passifloroideae node in future dating analyses that together with its neotropical geographic location can shed light on the origin and diversification of the subfamily. © 2017 Botanical Society of America.
National Oceanic and Atmospheric Administration, Department of Commerce — This data set contains sensitive biological resource data for diving birds, gulls and terns, seabirds, shorebirds, and waterfowl for the North Slope of Alaska....
Khozyem, H. M.; Adatte, T.; Keller, G.; Spangenberg, J.; Bajpai, S.; Samant, B.
The Paleocene Eocene Thermal Maximum (PETM, 55.9Ma)) is globally related with the extinction of deep benthic foraminifera, the diversification of both planktic foraminifera and modern mammals. In India, the tempo and timing of modern mammal dispersion, their association with the PETM or EECO (Early Eocene Climatic Optimum) and the India-Asia collision remain uncertain. Four Indian sections (Giral, Bhavnagar, Vastan and Tadkeshware lignite mines) have been studied using sedimentology, micropaleontology, mineralogy (bulk and clay mineralogy) and geochemistry (stable isotopes, major and trace elements, organic matter). Both PETM and ETM2 (second Eocene Thermal Maximum, 53.7Ma), a short-lived warming episode that followed the PETM, are globally marked by a pronounced δ13Ccarb and δ13Corg negative excursion. Both isotopic excursions have been recognized in the Vastan and Tarkeswhar lignite mines (Cambay basin, Gujarat) associated with the main mammal bearing level. The lower shift is located above the first lignite seam and corresponds to the transition from continental to shallow marine conditions. The upper excursion appears to be linked to the ETM2 and corresponds to a second marine incursion containing bivalves, benthic (Nummulites burdigalensis) and planktic foraminifera located below the second lignite seam. A very pronounced δ13Corg peak has been detected in the Giral lignite mine (Barmer, Rajhastan) around 6m above the vertebrate bearing level and may correspond to the PETM. This correlation is confirmed by palynological data and more particularly by a dinoflagellate acme that globally characterizes the PETM interval. Our micropaleontological data combined with stable carbone isotopes indicate the presence of both PETM and ETM2 events and constrain the age of the early mammals in northwestern India in between these two thermal events in the early Eocene. These new data will significantly improve the ongoing debate on whether mammals originated in or out of
Feb 25, 2005 ... covers approximately 250ha.The area encompassed by this study extends from Mambrui to the north, the sea to the east, the opposite bank of the estuary to the south and the Sabaki bridge and Malindi-Garsen road to the west. The area is defined as an Important Bird Area(IBA) by BirdLife International in ...
Halford, A. J.
When space computers first started listening into space radio, they noticed that there were radio noises that happened on the morning side of the Earth. Because these waves sounded like noises birds make in the morning, we named these waves after them. These bird sounding waves can move around the Earth, flying up and down, and sometimes move into an area where there is more stuff. This area is also much colder than where these bird noises are first made. When the waves move into this cold area where there is more stuff, they start to sound like angry birds instead of happy birds. Both of these waves, the happy and angry bird sounding waves, are very important to our understanding of how the tiny things in space move and change. Sometimes the waves which sound like birds can push these tiniest of things into the sky. The happy bird sounding waves can push the tiniest things quickly while the angry bird sounding waves push the tinest of things more slowly. When the tiny things fall into the sky, they create beautiful space lights and light that burns which can hurt people in up goers and not so up goers as well as our things like phones, and space computers. We study these waves that sound like birds to better understand when and where the tiny things will fall. That way we can be prepared and enjoy watching the pretty space lights at night with no worries.
Breda, Marzia; Lister, Adrian M.
The ancestry of the modern fallow deer, Dama dama, has been tentatively traced back to Pliocene/Early Pleistocene forms referred to 'Pseudodama', characterized by unpalmated three- or four-point antlers. By the late Middle Pleistocene, Dama with palmated antlers appears, as Dama dama clactoniana. However, fallow deer from the interim period, the early Middle Pleistocene, are poorly-known. A new specimen from Pakefield (Suffolk, UK), represented by a portion of cranium with a substantial part of both antlers plus a mandible and scapula, is the most complete medium-sized deer specimen from the British early Middle Pleistocene (ca 700 ka). The position and orientation of the basal tine, together with dental characters and mandibular morphology, are typical of fallow deer. The narrow palmation is reminiscent of D. dama clactoniana, but the lack of palmation tines is unique. Moreover, the lack of second (and third) tines in an adult specimen differs from both D. dama dama and D. d. clactoniana, being a primitive character shared with the last representatives of 'Pseudodama' which, on the other hand, has a circular beam lacking any palmation. This combination of features justifies the erection of a new species provisionally placed within the genus Dama, Dama roberti n. sp. Another specimen, from Soleilhac (Auvergne, France), represented by portions of the two antlers, a mandible and a tibia, shares antler morphology with the Pakefield specimen and can be ascribed to the same new species. Isolated antler and dental remains from coeval British sites are tentatively ascribed to D. roberti n. sp. The new species has implications for the ancestry of modern fallow deer.
Levison, Matthew E
Although many people these days actually work very hard at leisure time activities, diseases are most commonly acquired from birds during the course of work in the usual sense of the term, not leisure. However, travel for pleasure to areas where the diseases are highly endemic puts people at risk of acquiring some of these bird-related diseases (for example, histoplasmosis and arbovirus infections), as does ownership of birds as pets (psittacosis).
National Oceanic and Atmospheric Administration, Department of Commerce — This data set contains sensitive biological resource data for pelagic birds, shorebirds, wading birds, waterfowl, gulls, terns, and passerine birds in Guam and the...
National Oceanic and Atmospheric Administration, Department of Commerce — This data set contains sensitive biological resource data for wading birds, shorebirds, waterfowl, raptors, diving birds, seabirds, passerine birds, and gulls and...
Sideritic lacustrine mudstone was found in drill core from a uranium deposit in the Death Valley area in the eastern part of the Seward Peninsula, Alaska. The precursor sediments for this rock were deposited in an unusual iron-meromictic Eocene lake, herein named Lake Tubutulik, which occupied part of the Boulder Creek basin, a graben that is probably a southern extension of the larger Death Valley basin. The Boulder Creek basin is bounded on the west by granite of the Upper Cretaceous Darby pluton and on the east by Precambrian to Paleozoic metasedimentary rocks. The lake basin was formed by basaltic flows that dammed the valley of the ancestral Tubutulik River in early Eocene time. The lake sediments included a nearshore facies of fine-grained organic mud and an offshore facies of laminated sideritic mud. The offshore (profundal) laminated mudstone consists of alternating layers of authigenic siderite and detrital grains, mostly quartz and clay minerals. Both lacustrine facies contain turbidites. The lacustrine rocks graded laterally into an onshore facies of colluvial and fluvial sandstone, paludal mudstone, and coal. The ancient lake occupied a small, deep basin in a tectonically active area of high relief. Meromixis was apparently stabilized by reduced iron and bicarbonate dissolved in the monimolimnion. The intensity of meromixis decreased as the lake became shallower from sediment filling. The source of the dissolved iron in the monoimolimnion was probably the Eocene basalt. Carbon isotope analysis of the siderite suggests that the dissolved bicarbonate in the profundal facies was largely inorganic. Sideritic carbon in one sample from the onshore paludal facies has an isotopic signature (δ 13 C = +16.9) consistent with residual carbonate formed during methanogenic fermentation
Zeebe, Richard E.; Westerhold, Thomas; Littler, Kate; Zachos, James C.
Multimillion-year proxy records across the Paleocene and Eocene show prominent variations on orbital time scales. The cycles, which have been identified at various sites across the globe, preferentially concentrate spectral power at eccentricity and precessional frequencies. It is evident that these cycles are an expression of changes in global climate and carbon cycling paced by astronomical forcing. However, little is currently known about the link between orbital forcing and the carbon cycle-climate system and the amplitude of associated atmospheric CO2 variations. Here we use simple and complex carbon cycle models to explore the basic effect of different orbital forcing schemes and noise on the carbon cycle. Our primary modeling target is the high-resolution, ˜7.7 Myr long, benthic isotope record at Ocean Drilling Program Site 1262 in the South Atlantic. For direct insolation forcing (as opposed to artificial eccentricity-tilt-precession), one major challenge is understanding how the system transfers spectral power from high to low frequencies. We discuss feasible solutions, including insolation transformations analogous to electronic AC-DC conversion (DC'ing). Regarding mechanisms, we focus on tropical insolation and a long-term carbon imbalance in terrestrial organic burial/oxidation but do not rule out other scenarios. Our analysis shows that high-latitude mechanisms are unlikely drivers of orbitally paced changes in the late Paleocene-early Eocene (LPEE) Earth system. Furthermore, we provide constraints on the origin and isotopic composition of a possible LPEE cyclic carbon imbalance/source responding to astronomical forcing. Our simulations also reveal a mechanism for the large δ13C-eccentricity lag at the 400 kyr period observed in Paleocene, Oligocene, and Miocene sections. We present the first estimates of orbital-scale variations in atmospheric CO2 during the late Paleocene and early Eocene.
Spiegel, Cornelia; Lisker, Frank; Piepjohn, Karsten; Estrada, Solveig; Lorenz, Henning
The structural evolution of the Arctic Ocean and its surrounding continental areas is still poorly constrained, due to ice coverage and inaccessibility. The only scientific coring campaign within the central Arctic Ocean (the ACEX project) was positioned on the Lomonosov Ridge. This is a pronounced geomorphic structure of presumably continental origin, which stretches across the entire Arctic Ocean between the northernmost margin of the North American continent (Ellesmere Island) and the Siberian Shelf, bordering the New Siberian Islands. Geophysical data suggest that the Lomonosov Ridge may be continuous with the Siberian and Ellesmerian continental margins (e.g., Poselov et al., 2011). Rather unexpectedly, the ACEX project revealed that the Lomonosov Ridge was in very shallow water or even exposed to erosion between 44 and 18 Ma. As an explanation, it was suggested that the Lomonosov Ridge experienced compressional tectonics at that time, which may have affected the entire central Arctic Ocean, propagating from North America towards the Siberian shelf (ÓRegan et al., 2008). Here we present the first low-temperature thermochronological data from northern Ellesmere Island and from the New Siberian Islands, recording the erosion and exhumation history of these areas. Our apatite (U-Th)/He data show that while southern and central Ellesmere Island was characterized by very slow erosion during the Cenozoic, northern Ellesmere Island bordering the Arctic Ocean experienced km-scale erosion during the Eocene, contemporaneously with the stalled subsidence / uplift period of the Lomonosov Ridge. The thermochronology data from the New Siberian Islands reflect a complex erosion history: the eastern part of the North Siberian Islands, the DeLong Island Group, experienced rather limited erosion during the Cenozoic and most of the Mesozoic. By contrast, data from the western New Siberian Islands - the Lyakhov Island Group - in direct continuation of the Lomonosov Ridge are
Julián F. Petrulevičius
Full Text Available A new burmagomphid anisopteran, Satelitala soberana gen. et sp. nov. is described from the lower Eocene of Laguna del Hunco, Patagonia, Argentina. The new genus is characterised by hindwing characters such as the subdiscoidal triangle not elongated; anal loop divided longitudinally; paranal cell divided longitudinally; five terminal cells between RP and MA; five terminal cells between MP and CuA; and obtuse angle between PsA and CuP+AA. Burmagomphid dragonflies were represented so far only by one specimen from the middle Cretaceous of Southeast Asia. This new record extends the distribution to Patagonia, to the Cenozoic, and also to paleolake deposits.
Otto, Angelika; White, James D; Simoneit, Bernd R T
Numerous saturated and aromatic hydrocarbons, but not polar compounds, originating from plants and microorganisms (biomarkers) have been reported in sediments, coals, and petroleum. Here we describe natural product terpenoids found in two fossil conifers, Taxodium balticum (Eocene) and Glyptostrobus oregonensis (Miocene). A similar terpenoid pattern is also observed in extant Taxodium distichum. The preservation of characteristic terpenoids (unaltered natural products) in the fossil conifers supports their systematic assignment to the Cypress family (Cupressaceae sensu lato). The results also show that fossil conifers can contain polar terpenoids, which are valuable markers for (paleo)chemosystematics and phylogeny.
Bosboom, R.E.|info:eu-repo/dai/nl/322947359; Abels, H.A.|info:eu-repo/dai/nl/304848018; Hoorn, C.; van den Berg, B.C.J.; Guo, Z.; Dupont-Nivet, G.|info:eu-repo/dai/nl/313092559
Global climate cooling from greenhouse to icehouse conditions occurred across an enigmatic transitional interval during the Eocene epoch characterized by incipient polar ice-sheet formation as well as short-lived warming events, of which the Middle Eocene Climatic Optimum (MECO) is most noticeable.
Kristensen, Hans Viborg; Cuny, Gilles; Redsted Rasmussen, Arne
Abstract. – The earliest record of Palaeophis ever found in Denmark is here based on vertebrae described from the Paleocene/Eocene Stolleklint Clay of the Isle of Mors (northern Denmark). Although much smaller, they appear quite similar to the Eocene vertebra described from the Fur Formation in t...
Douglas, P.M.J.; Affek, H.P.; Ivany, L.C.; Houben, A.J.P.; Sijp, W.P.; Sluijs, A.; Schouten, S.; Pagani, M.
Paleoclimate studies suggest that increased global warmth during the Eocene epoch was greatly amplified at high latitudes, a state that climate models cannot fully reproduce. However, proxy estimates of Eocene near-Antarctic sea surface temperatures (SSTs) have produced widely divergent results at
Sluijs, A.; Schouten, S.; Donders, T.H.; Schoon, P.L.; Röhl, U.; Reichart, G.-J.; Sangiorgi, F.; Kim, J.-H.; Sinninghe Damsté, J.S.; Brinkhuis, H.
Several episodes of abrupt and transient warming, each lasting between 50,000 and 200,000 years, punctuated the long-term warming during the Late Palaeocene and Early Eocene (58 to 51 Myr ago) epochs1,2. These hyperthermal events, such as the Eocene Thermal Maximum 2 (EMT2) that took place about
Barron, John A.; Stickley, Catherine E.; Bukry, David
Eocene diatom and silicoflagellate biostratigraphy are summarized and correlated with the most recent geologic time scale as well as with the global oxygen isotope and eustatic sea level curves. The global distribution of Eocene diatom/silicoflagellate-bearing sediments varies considerably, reflecting changing oceanic gateways and paleoceanography with changing patterns that are punctuated by four major depositional events.
Solov'yov, Ilia; Schulten, Klaus
A proposed model for migrating birds' magnetic sense can withstand moderate orientational disorder of a key protein in the eye.......A proposed model for migrating birds' magnetic sense can withstand moderate orientational disorder of a key protein in the eye....
Solov'yov, Ilia; Greiner, Walter
The Earth's magnetic field provides an important source of directional information for many living organisms, especially birds, but the sensory receptor responsible for magnetic field detection still has to be identified. Recently, magnetic iron oxide particles were detected in dendritic endings...... field, by a bird....
... However, some ducks can be infected without any signs of illness. Top of Page Avian Influenza in Wild Birds Avian influenza A viruses have ... hours. Some ducks can be infected without any signs of illness. Avian influenza outbreaks are of concern in domesticated birds for ...
Snep, Robbert P.H.; Kooijmans, Jip Louwe; Kwak, Robert G.M.; Foppen, Ruud P.B.; Parsons, Holly; Awasthy, Monica; Sierdsema, Henk L.K.; Marzluff, John M.; Fernandez-Juricic, Esteban; Laet, de Jenny
Following the call from the United Nations Convention on Biological Diversity “Cities & Biodiversity Outlook” project to better preserve urban biodiversity, this paper presents stakeholder-specific statements for bird conservation in city environments. Based upon the current urban bird
Fish and Wildlife Service (Dept. of Interior), Washington, DC.
The materials in this educational packet are designed for use with students in grades 4 through 7. They consist of an overview, teaching guides and student data sheets for three activities, and a poster. The overview discusses why, how, where, and when birds migrate as well as problems birds encounter while migrating; the importance of research…
Greek mythology it was a nondescript bird but in the medieval bestiaries it became pure white. The caladrius is used in the coats of arms of the South African Medical and Dental Council and also the Medical University of Soufhern Africa. These appear to be the first use of this medically significant bird in modern heraldry.
Vizán, H.; Geuna, S.; Melchor, R.; Bellosi, E. S.; Lagorio, S. L.; Vásquez, C.; Japas, M. S.; Ré, G.; Do Campo, M.
The red bed succession cropping out in the Quebrada Santo Domingo in northwestern Argentina had been for long considered as Upper Triassic-Lower Jurassic in age based on weak radiometric and paleontological evidence. Preliminary paleomagnetic data confirmed the age and opened questions about the nature of fossil footprints with avian features discovered in the section. Recently the stratigraphic scheme was reviewed with the identification of previously unrecognized discontinuities, and a radiometric dating obtained in a tuff, indicated an Eocene age for the Laguna Brava Formation and the fossil bird footprints, much younger than the previously assigned. We present a detailed paleomagnetic study interpreted within a regional tectonic and stratigraphic framework, looking for an explanation for the misinterpretation of the preliminary paleomagnetic data. The characteristic remanent magnetizations pass a tilt test and a reversal test. The main magnetic carrier is interpreted to be low Ti titanomagnetites and to a lesser extent hematite. The characteristic remanent magnetization would be essentially detrital. The obtained paleomagnetic pole (PP) for the Laguna Brava Formation has the following geographic coordinates and statistical parameters: N = 29, Lon. = 184.5° E, Lat. = 75.0° S, A95 = 5.6° and K = 23.7. When this PP is compared with another one with similar age obtained in an undeformed area, a declination anomaly is recognized. This anomaly can be interpreted as Laguna Brava Formation belonging to a structural block that rotated about 16° clockwise along a vertical axis after about 34 Ma. This block rotation is consistent with the regional tectonic framework, and would have caused the fortuitous coincidence of the PP with Early Jurassic poles. According to the interpreted magnetostratigraphic correlation, the Laguna Brava Formation would have been deposited during the Late Eocene with a mean sedimentation rate of about 1.4 cm per thousand years, probably in
Jałoszyński, Paweł; Perkovsky, Evgeny
Among nearly 1270 inclusions of Coleoptera found in Upper Eocene Rovno amber, 69 were identified as ant-like stone beetles (Scydmaeninae); 34 were possible to unambiguously determine to the tribal level and were studied in detail. Rovnoleptochromus ableptonoides gen. & sp. n. (Mastigitae: Clidicini), Vertheia quadrisetosa gen. & sp. n. (Cephenniitae: Eutheiini), Cephennomicrus giganteus sp. n. (Cephenniitae: Cephenniini), Glaesoconnus unicus gen. & sp. n. (Scydmaenitae: Glandulariini), Rovnoscydmus frontalis gen. & sp. n. (Scydmaenitae: Glandulariini; type species of Rovnoscydmus), Rovnoscydmus microscopicus sp. n., Euconnus (incertae sedis, near Cladoconnus) palaeogenus sp. n. (Scydmaenitae: Glandulariini), and Stenichnus (s. str.) proavus sp. n. (Scydmaenitae: Glandulariini) are described. Additionally, specimens representing one undescribed species of Vertheia, one of Cephennodes, five of Cephennomicrus, one of Euconnus, one of Microscydmus are recorded, and nine specimens representing an unknown number of species of Rovnoscydmus (and two putative Rovnoscydmus), one Euconnus (and one putative Euconnus), two putative Microscydmus and one putative Scydmoraphes were found in the studied material. The composition of Scydmaeninae fauna in Rovno amber is discussed in the context of ecological preferences and distribution of extant taxa. It is concluded that subtropical and tropical taxa were present in the region where Rovno amber has formed, most notably the second genus and species of the extant tribe Clidicini known from the Eocene of Europe, and six species of the extant genus Cephennomicrus, for the first time found in the fossil record. An annotated catalog of nominal species of Scydmaeninae known in the fossil record is given.
In this paper a mathematical model for the transmission dynamics of bird flu among birds and humans is presented. The model assumes that there is no migration of birds in the susceptible bird population immediately the disease starts. The model formulated is analyzed using dynamical systems theory . The analysis of the ...
In this paper a mathematical model for the transmission dynamics of bird flu among birds and humans is presented. The model assumes that there is no migration of birds in the susceptible bird population immediately the disease starts. The model formulated is analyzed using dynamical systems theory. The analysis of the ...
For many animals, the effort to rear their young is considerable. In birds, this often includes building nests, incubating eggs, feeding the chicks, and protecting them from predators. Perhaps for this reason, about 1% of birds (around 100 species) save themselves the effort and cheat instead. They are obligate brood parasites, laying their eggs in the nests of other species and leaving the hosts or foster parents to rear the foreign chicks for them. Some birds also cheat on individuals of the same species (intraspecific brood parasitism). Intraspecific brood parasitism has been reported in around 200 species, but is likely to be higher, as it can often only be detected by genetic analyses.
This bachelor thesis is about hatching synchrony in birds. Generally, among birds there are two types of hatching - asynchronous and synchronous- and the type of hatching is primarily determined by the time of the onset of incubation. In many bird species, including most precocial ones, incubation does not begin until the last egg has been laid, which results in hatching of all the eggs within a few hours. In synchronously-hatched broods, all the chicks are about the same age. Thus no single ...
Garun-Rashid Abdul-Kadyrovich Guseynov
Full Text Available Founded in 438 and included on the World Heritage List in 2003, the ancient city of Derbent emerged in the area which, by the time of its foundation, was known under various names which belong, as it is shown in the paper, to different Bulgar (Turkic languages: province of Chor, Ζουάρ-ος fortress, Chora / Djora way, Τζοΰρ, city of Chora, Chola / Choga / Chol, gate of Chola / Choga, province of Sul. These dialectal place names meaning ʻsteep bank, precipiceʼ meet the localization of the city in a narrow passageway between the Greater Caucasus and the Caspian Sea. The author argues that all etymologies of the considered oikonyms and horonyms based on linguistic data other than Bulgar should be recognized as inadequate. Their Bulgar origin is supported by the toponymic landscape and the names of the peoples who lived within its borders in the early Middle Ages. The existence of toponymic parallels outside the region in question is explained by the migration, in the first centuries AD, of the Bulgars from the North-Eastern Caucasus and Dagestan to Eastern and Southern Europe — the Volga region, the Balkans and Crimea.
Qian, Tao; Liu, Shaofeng; Wang, Zongxiu; Li, Wangpeng; Chen, Xinlu
The Mesozoic northern Yangtze foreland basin system was formed by continental collision between the North China and South China plates along the Mianlue suture. Synorogenic stratigraphic sequences of Late Triassic to Early-Middle Jurassic age were developed in the northern Yangtze foreland basin. The upper Middle Jurassic Shaximiao Formation consists mainly of thick-bedded terrestrial successions that serve as the main body of the basin-filling sequences, suggesting intense tectonism in the peripheral orogeny of the foreland basin. Laser-ICP-MS U-Pb analysis of 254 detrital zircon grains from sandstone samples and several published Lower-Middle Jurassic samples, detrital compositions, petrofacies, and paleocurrent reconstructions in the northern Yangtze foreland basin indicate that discrete source areas included the Qinling-Dabieshan ranges and the Mianlue suture zone to the north, and the South China plate to the south. The stratigraphic succession and sediment provenance of the foreland basin imply that the early Mianlue oceanic basin, magmatic arc, and nonmarine molasse foreland basin during the period of deposition were modified or buried by the subsequent continent-continent collision between the North China-Qinling-Dabieshan plate and the Yangtze plate during the Jurassic, which followed the oblique amalgamation between these plates during the Middle-Late Triassic.
Morado-Urbina, Carlos Eduardo; Alvarado-Vázquez, Perla Abigail; Montiel-Ruiz, Rosa Mariana; Acosta-González, Rosa Issel; Castañeda-Corral, Gabriela; Jiménez-Andrade, Juan Miguel
This study was performed to evaluate whether early, middle, or late treatment of zoledronate, an approved bisphosphonate that blocks bone resorption, can reduce nociceptive behaviors in a mouse arthritis model. Arthritis was produced by repeated intra-articular knee injections of complete Freund's adjuvant (CFA). A dose-response curve with zoledronate (3, 30, 100, and 300 μg/kg, i.p., day 4 to day 25, twice weekly for 3 weeks) was performed, and the most effective dose of zoledronate (100 μg/kg, i.p.) was initially administered at different times of disease progression: day 4 (early), day 15 (middle), or day 21 (late) and continued until day 25 after the first CFA injection. Flinching of the injected extremity (spontaneous nociceptive behavior), vertical rearings and horizontal activity (functional outcomes), and knee edema were assessed. Zoledronate improved both functional outcomes and reduced flinching behavior. At day 25, the effect of zoledronate on flinching behavior and vertical rearings was greater in magnitude when it was given early or middle rather than late in the treatment regimen. Chronic zoledronate did not reduce knee edema in CFA-injected mice nor functional outcomes in naïve mice by itself. These results suggest that zoledronate may have a positive effect on arthritis-induced nociception and functional disabilities. © 2014 Wiley Periodicals, Inc.
All over the nation, birds of all shapes and sizes attempt to make schools a their favorite hangout. Their arrival can lead to sanitation issues, added facility degradation, distracted students and health problems.
California Natural Resource Agency — This data set provides access to information gathered on annual breeding bird surveys in California using a map layer developed by the Department. This data layer...
National Aeronautics and Space Administration — A recent breakthrough in MWIR detector design, has resulted in a high operating temperature (HOT) barrier infrared detector (BIRD) that is capable of spectral...
... in People Spread of Bird Flu Viruses Between Animals and People Examples of Human Infections with Avian Influenza A ... Subtypes Transmission of Avian Influenza A Viruses Between Animals and People Related Links Research Glossary of Influenza (Flu) Terms ...
U.S. Environmental Protection Agency — The data are maintained by the USGS (https://www.pwrc.usgs.gov/bbs/RawData/) and provides information on the trends and status of North American bird populations...
In this article, the author describes a watercolor art lesson on Audubon birds. She also discusses how science, technology, writing skills, and the elements and principles of art can be incorporated into the lesson.
Queiroz, Ana Isabel; Soares, Filipa
UID/ELT/00657/2013 WOS:000374914600004 IF/00222/2013 Birds are emblematic natural elements of landscapes. Readily noticeable and appreciated due to their songs and flight, they have been thoroughly used as components of literary scenarios. This paper analyses their representations in a broad corpus (144 writings by 67 writers) since the nineteenth century, divided in three time-periods. It aims to understand which wild birds are represented in Portuguese literature, how those represe...
Aydoğdu, A; Frasca, P; D'Apice, C; Manzo, R; Thornton, J M; Gachomo, B; Wilson, T; Cheung, B; Tariq, U; Saidel, W; Piccoli, B
In this paper we introduce a mathematical model to study the group dynamics of birds resting on wires. The model is agent-based and postulates attraction-repulsion forces between the interacting birds: the interactions are "topological", in the sense that they involve a given number of neighbors irrespective of their distance. The model is first mathematically analyzed and then simulated to study its main properties: we observe that the model predicts birds to be more widely spaced near the borders of each group. We compare the results from the model with experimental data, derived from the analysis of pictures of pigeons and starlings taken in New Jersey: two different image elaboration protocols allow us to establish a good agreement with the model and to quantify its main parameters. We also discuss the potential handedness of the birds, by analyzing the group organization features and the group dynamics at the arrival of new birds. Finally, we propose a more refined mathematical model that describes landing and departing birds by suitable stochastic processes. Copyright © 2016 Elsevier Ltd. All rights reserved.
Full Text Available Two fragments of white marble are embedded in the apsis’ wall outside the parish church of St. Agatha in Santhià (near Vercelli, in Piedmont: the first one is engraved with the final part of a dedicatory inscription to a martyr by a clergyman named Vvalpertus (or Alpertus; the second one is a part of a small pillar or a lintel coming from a presbytery enclosure, and is decorated with a motif of intertwined gallons. The inscription stayed in the middle of a pluteus bordered by a carved frame, of which only a part of the lower band survives: the letters and the style of the decorations allow a chronology within the 8th century, while the fragment of pillar is to be traced back to a period between the 8th and the first half of the 9th. Of special interest are the Lombard onomastic of the dedicator and, in the final surviving text, the “signature” of a sculptor, which is present in a second inscription of Lombard age found in Piedmont, adorned with carved decorations as well. The analysis of these artefacts - hitherto unpublished - offered the opportunity to restudy two other inscriptions once existing in the church of St. Agatha, long lost and of uncertain authenticity: all these data allow various deductions about the history of Santhià over the centuries of the early Middle ages, the presence of a clergy in the countryside of the diocese of Vercelli, and the work of sculptors and engravers active between the 8th and 9th century in an area between Lombardy, Piedmont, Liguria and Provence, labelled in the past under the unifying name of “bottega delle Alpi Marittime”.
Assel, Melissa J; Gerdtsson, Axel; Thorek, Daniel L J; Carlsson, Sigrid V; Malm, Johan; Scardino, Peter T; Vickers, Andrew; Lilja, Hans; Ulmert, David
To evaluate whether anthropometric parameters add to PSA measurements in middle-aged men for risk assessment of prostate cancer (PCa) diagnosis and death. After adjusting for PSA, both BMI and weight were significantly associated with an increased risk of PCa death with the odds of a death corresponding to a 10 kg/m2 or 10 kg increase being 1.58 (95% CI 1.10, 2.28; p = 0.013) and 1.14 (95% CI 1.02, 1.26; p = 0.016) times greater, respectively. AUCs did not meaningfully increase with the addition of weight or BMI to prediction models including PSA. In 1974 to 1986, 22,444 Swedish men aged 44 to 50 enrolled in Malmö Preventive Project, Sweden, and provided blood samples and anthropometric data. Rates of PSA screening in the cohort were very low. Documentation of PCa diagnosis and disease-specific death up to 2014 was retrieved through national registries. Among men with anthropometric measurements available at baseline, a total of 1692 men diagnosed with PCa were matched to 4190 controls, and 464 men who died of disease were matched to 1390 controls. Multivariable conditional logistic regression was used to determine whether diagnosis or death from PCa were associated with weight and body mass index (BMI) at adulthood after adjusting for PSA. Men with higher BMI and weight at early middle age have an increased risk of PCa diagnosis and death after adjusting for PSA. However, in a multi-variable numerical statistical model, BMI and weight do not importantly improve the predictive accuracy of PSA. Risk-stratification of screening should be based on PSA without reference to anthropometrics.
Vlad D. Ghimpu
Full Text Available Based on the research of many rock-hewn churches of the Romanian region, located in the Dobruja andaround the Carpathian mountain range, the author notes the existence there of a number of Christian churches belonging to the early Christian period and the early Middle Ages. According to the author, the religious buildingswere created for the most part of very strong material because of the highly volatile external environment, due to the pressure of the migratory and nomadic peoples. On the basis of new archaeological data it can be argued that in the territory of the future Moldova there was a diocese under the guidance of a chorbishop (bishop of rural settlements. The presence and discovery of a very valuable exhibit, which is a rare example of art, iconography and folklore heritage of Romanians in early Middle Ages (red brick with depictions, gives us the first written evidence of the presence of Christianity in this area.
Abels, H.A.; Xu, Y.; Utescher, T.; Dupont-Nivet, G.
Climate models suggest that Asian paleoenvironments, monsoons and continental aridification were primarily governed by tectonic uplift and sea retreat since the Eocene with potential contribution of global climate changes. However, the cause and timing of these paleoenvironmental changes remain
Kvaček, Z.; Teodoridis, V.; Mach, K.; Přikryl, Tomáš; Dvořák, Z.
Roč. 89, č. 1 (2014), s. 21-66 ISSN 1214-1119 Keywords : boundary * climate * Early Oligocene * fauna * flora * Late Eocene * vegetation Subject RIV: DB - Geology ; Mineralogy Impact factor: 1.515, year: 2014
Pollen data show that in southeastern North America, the Eocene angiosperm flora attained its maximum relative diversity some 8 m.y. after the late early Eocene to earliest middle Eocene to earliest middle Eocene climatic maximum. Increasing diversity resulted in part from the flora's position on a large continent which allowed easy migration. In western Europe, the floral diversity began decreasing even before the climatic maximum. Paleogeography played large roles in this diversity decrease. In western Europe, terrestrial floras were on islands and peninsulas in the sea, so that the floras underwent increasing isolation and partial local extermination. Temperate plants generally did not migrate to western Europe, because of a lack of nearby uplands, lack of northern terrestrial source areas for these plants, and presence of the Turgai Straights barrier. -from Authors
Full Text Available Unlike airplanes birds must have either flapping or oscillating wings (the hummingbird. Only such wings can produce both lift and thrust – two sine qua non attributes of flying.The bird wings have several possibilities how to obtain the same functions as airplane wings. All are realized by the system of flight feathers. Birds have also the capabilities of adjusting the shape of the wing according to what the immediate flight situation demands, as well as of responding almost immediately to conditions the flow environment dictates, such as wind gusts, object avoidance, target tracking, etc. In bird aerodynamics also the tail plays an important role. To fly, wings impart downward momentum to the surrounding air and obtain lift by reaction. How this is achieved under various flight situations (cruise flight, hovering, landing, etc., and what the role is of the wing-generated vortices in producing lift and thrust is discussed.The issue of studying bird flight experimentally from in vivo or in vitro experiments is also briefly discussed.
Major, Andrew T; Smith, Craig A
Sexual differentiation in birds is controlled genetically as in mammals, although the sex chromosomes are different. Males have a ZZ sex chromosome constitution, while females are ZW. Gene(s) on the sex chromosomes must initiate gonadal sex differentiation during embryonic life, inducing paired testes in ZZ individuals and unilateral ovaries in ZW individuals. The traditional view of avian sexual differentiation aligns with that expounded for other vertebrates; upon sexual differentiation, the gonads secrete sex steroid hormones that masculinise or feminise the rest of the body. However, recent studies on naturally occurring or experimentally induced avian sex reversal suggest a significant role for direct genetic factors, in addition to sex hormones, in regulating sexual differentiation of the soma in birds. This review will provide an overview of sex determination in birds and both naturally and experimentally induced sex reversal, with emphasis on the key role of oestrogen. We then consider how recent studies on sex reversal and gynandromorphic birds (half male:half female) are shaping our understanding of sexual differentiation in avians and in vertebrates more broadly. Current evidence shows that sexual differentiation in birds is a mix of direct genetic and hormonal mechanisms. Perturbation of either of these components may lead to sex reversal. © 2016 S. Karger AG, Basel.
Suzuki, Shigeyuki; Peña, Rolando E.; Tam, Tomas A.; Yumul, Graciano P.; Dimalanta, Carla B.; Usui, Mayumi; Ishida, Keisuke
The origin of the Philippine Archipelago is characterized by the combination of the oceanic Philippine Mobile Belt (PMB) and the Palawan Continental Block (PCB). This paper is focused on the geologic evolution of the PMB in northern Luzon from Eocene to Pliocene. The study areas (northern Luzon) are situated in the central part of the PMB which is occupied by its typical components made up of a pre-Paleocene ophiolitic complex, Eocene successions, Eocene to Oligocene igneous complex and late Oligocene to Pliocene successions. Facies analysis of the middle Eocene and late Oligocene to early Pliocene successions was carried out to understand the depositional environment of their basins. Modal sandstone compositions, which reflect the basement geology of the source area, were analyzed. Major element geochemistry of sediments was considered to reconstruct the tectonic settings. The following brief history of the PMB is deduced. During the middle Eocene, the PMB was covered by mafic volcanic rocks and was a primitive island arc. In late Eocene to late Oligocene time, the intermediate igneous complex was added to the mafic PMB crust. By late Oligocene to early Miocene time, the PMB had evolved into a volcanic island arc setting. Contributions from alkalic rocks are detected from the rock fragments in the sandstones and chemical composition of the Zigzag Formation. During the middle Miocene to Pliocene, the tectonic setting of the PMB remained as a mafic volcanic island arc.
Larsen, Frank Wugt; Bladt, Jesper Stentoft; Balmford, Andrew
1. Most biodiversity is still unknown, and therefore, priority areas for conservation typically are identified based on the presence of surrogates, or indicator groups. Birds are commonly used as surrogates of biodiversity owing to the wide availability of relevant data and their broad popular...... appeal. However, some studies have found birds to perform relatively poorly as indicators. We therefore ask how the effectiveness of this approach can be improved by supplementing data on birds with information on other taxa. 2. Here, we explore two strategies using (i) species data for other taxa...... and (ii) genus- and family-level data for invertebrates (when available). We used three distinct species data sets for sub-Saharan Africa, Denmark and Uganda, which cover different spatial scales, biogeographic regions and taxa (vertebrates, invertebrates and plants). 3. We found that networks of priority...
Angst, D.; Lécuyer, C.; Amiot, R.; Buffetaut, E.; Fourel, F.; Martineau, F.; Legendre, S.; Abourachid, A.; Herrel, A.
The mode of life of the early Tertiary giant bird Gastornis has long been a matter of controversy. Although it has often been reconstructed as an apex predator feeding on small mammals, according to other interpretations, it was in fact a large herbivore. To determine the diet of this bird, we analyze here the carbon isotope composition of the bone apatite from Gastornis and contemporaneous herbivorous mammals. Based on 13C-enrichment measured between carbonate and diet of carnivorous and herbivorous modern birds, the carbonate δ13C values of Gastornis bone remains, recovered from four Paleocene and Eocene French localities, indicate that this bird fed on plants. This is confirmed by a morphofunctional study showing that the reconstructed jaw musculature of Gastornis was similar to that of living herbivorous birds and unlike that of carnivorous forms. The herbivorous Gastornis was the largest terrestrial tetrapod in the Paleocene biota of Europe, unlike the situation in North America and Asia, where Gastornis is first recorded in the early Eocene, and the largest Paleocene animals were herbivorous mammals. The structure of the Paleocene terrestrial ecosystems of Europe may have been similar to that of some large islands, notably Madagascar, prior to the arrival of humans.
National Oceanic and Atmospheric Administration, Department of Commerce — This data set contains biological resource data for alcids, shorebirds, waterfowl, diving birds, pelagic birds, gulls and terns in Cook Inlet and Kenai Peninsula,...
Tabatabaei, Seyed Ali; Soleimani, Mohammad; Behrouz, Mahmoud Jabbarvand
To report 30 patients with bird attack-related eye injuries. This study was performed among patients coming to Farabi Eye Hospital, Tehran, Iran, from 2010 to 2015 with a history of bird attack causing eye injury. The inclusion criteria were a history of bird attack by pecking causing eye injury and having treatment and follow-up record for at least 6 months after treatment. The primary eye examinations included a full ophthalmic examination including evaluation of uncorrected visual acuity and best-corrected visual acuity (BCVA), anterior segment slit lamp biomicroscopy, and photography. For all patients with penetrating injury, primary repair was undertaken. Thirty patients (10 females and 20 males) with a mean age of 23.3 ± 18.5 years entered the study. The most common zone of injury was zone 1 (P < 0.001), and lensectomy was not needed in majority of patients (P < 0.001). The most common bird causing the injury was mynah (P < 0.001). Those patients with baseline BCVA of less than 20/200 or those with endophthalmitis had statistically worse final BCVA after treatment. Patients attacked by mynah bird had significantly better pretreatment uncorrected visual acuity and BCVA. The most common bird causing the eye injury among the sample of patients from Iran was mynah, which differs with previous studies indicating the rooster attack as the most common cause of eye injury. The authors also found that the most common zone of injury was zone 1, and the presence of endophthalmitis and lower baseline BCVA were significant risk factors for worse visual outcomes.
Tobalske, Bret W
Power output is a unifying theme for bird flight and considerable progress has been accomplished recently in measuring muscular, metabolic and aerodynamic power in birds. The primary flight muscles of birds, the pectoralis and supracoracoideus, are designed for work and power output, with large stress (force per unit cross-sectional area) and strain (relative length change) per contraction. U-shaped curves describe how mechanical power output varies with flight speed, but the specific shapes and characteristic speeds of these curves differ according to morphology and flight style. New measures of induced, profile and parasite power should help to update existing mathematical models of flight. In turn, these improved models may serve to test behavioral and ecological processes. Unlike terrestrial locomotion that is generally characterized by discrete gaits, changes in wing kinematics and aerodynamics across flight speeds are gradual. Take-off flight performance scales with body size, but fully revealing the mechanisms responsible for this pattern awaits new study. Intermittent flight appears to reduce the power cost for flight, as some species flap-glide at slow speeds and flap-bound at fast speeds. It is vital to test the metabolic costs of intermittent flight to understand why some birds use intermittent bounds during slow flight. Maneuvering and stability are critical for flying birds, and design for maneuvering may impinge upon other aspects of flight performance. The tail contributes to lift and drag; it is also integral to maneuvering and stability. Recent studies have revealed that maneuvers are typically initiated during downstroke and involve bilateral asymmetry of force production in the pectoralis. Future study of maneuvering and stability should measure inertial and aerodynamic forces. It is critical for continued progress into the biomechanics of bird flight that experimental designs are developed in an ecological and evolutionary context.
The accidental killing of birds by wind generators, and design improvements in the towers that support the turbines that might cut down on the bird killings were discussed. The first problem for the industry began in the late 1980s when the California Energy Commission reported as many as 160 birds (the majority being raptors, including the protected golden eagle) killed in one year in the vicinity of wind power plants. The key factor identified was the design of the towers as birds of prey are attracted to lattice towers as a place to hunt from. Tubular towers do not provide a place for the birds to perch, therefore they reduce the potential for bird strikes. Bird strikes also have been reported in Spain and the siting of the towers have been considered as the principal cause of the bird strikes. In view of these incidents, the wind power industry is developing standards for studying the potential of bird strikes and is continuing to study bird behaviour leading to collisions, the impact of topography, cumulative impacts and new techniques to reduce bird strikes. Despite the reported incidents, the risk of bird strikes by wind turbines, compared to other threats to birds such as pollution, oil spills, and other threats from fossil and nuclear fuels, is considered to be negligible. With continuing efforts to minimize incidents by proper design and siting, wind power can continue to grow as an environmentally sound and efficient source of energy
Rana, R.S.; Kumar, K.; Escarguel, G.; Sahni, A.; Rose, K.D.; Smith, T.; Singh, H.; Singh, L. [Wadia Institute of Himalayan Geology, Dehra Dun (India)
A new ailuravine rodent, Meldimys musak sp. nov. (Mammalia: Rodentia, Ischyromyidae), is recorded from the lower Eocene lignites of western India. It is the oldest record of Rodentia from India. M. musak is more derived than the earliest Eocene ailuravine Euromys cardosoi from Portugal and more generalized than late early Eocene E. inexpectatus and Ailuravus michauxi from France. Its dental morphology closely corresponds to the middle early Eocene species M. louisi, which lived about 52 Ma in Western Europe. Meldimys was previously known only from Europe, and ailuravines were previously reported only from Europe and North America. Its occurrence in India allows the first direct correlation between the early Eocene land mammal horizons of Europe and India, and raises the possibility of a terrestrial faunal exchange between India and Eurasia close to the Palaeocene-Eocene transition.
Scomazzon, Ana Karina; Moutinho, Luciane Profs; Nascimento, Sara; Lemos, Valesca Brasil; Matsuda, Nilo Siguehiko
increasingly abundant to the top of carbonates, also shales and siltstones intercalated with the sabkha plain. The carbonate strata have similar marine fossils to those of Itaituba Formation, however, the fauna becomes impoverished in abundance and diversity. The conodont fauna of Early-Middle Pennsylvanian of Amazonas Basin suggests similarities of species with the North American Midcontinent region and the Illinois Basin indicating their probable cosmopolitism; however in a slightly different paleogeographic context, probably due to latitudinal differences, coal deposits are present in North America, whereas widespread evaporitic deposits are registered to the Tapajós Group, Amazonas Basin.
Cluzel, Dominique; Jolivet, Laurent; Cadet, Jean-Paul
The polycyclic Ogcheon belt of South Korea represents the boundary between two Precambrian blocks. Investigation on the timing and kinematics of the block agglomeration is of considerable bearing on the understanding of the constitution of the Asian continent. We report here structural evidence showing that the earlier tectonic event dates back to early middle Paleozoic (the "Ogcheon tectonism" or "orogeny"). The ductile piling up of nappes of the Ogcheon belt corresponds to an intracontinental orogeny involving the opening and the subsequent closure of an aborted rift and not of a wide oceanic area. The nappes contain unequivocal evidence for ductile shearing with a northwest over southeast sense of movement. Field evidence includes the presence of S1 and S1-2 foliations bearing an extensive stretching lineation perpendicular to the belt, associated with a strongly noncoaxial deformation regime. The F1 isoclinal folds are commonly "A type" and sheath folds whose axes are parallel to the stretching lineation, whereas the southeast facing F2 recumbent folds have subhorizontal axes parallel to the belt. All lines of evidence (unconformity, superposed deformations, age of the metamorphism, etc…) imply that the D1-2 "Ogcheon tectonism" is pre-middle Carboniferous (Late Silurian-Early Devonian). The subsequent Indosinian (Middle Triassic) F3 upright synfolial folds that trend NNE-SSW and later structures clearly overprint the middle Paleozoic ductile thrusts and isoclinal folds. The early Paleozoic Ogcheon Supergroup is formed of a thick volcanosedimentary pile deposited above an early platform sequence in a rift basin. On the basis of the geochemical characteristics of the metavolcanics, it appears that oceanization did not occur and Ogcheon rift aborted soon after it was formed. We suggest that middle Paleozoic ductile nappe structures have been formed by the structural inversion of the rift extensional features. During the ductile stacking episode, the early formed
Tomotani, B.M.; Ramakers, J.J.C.; Gienapp, P.
Climate change can affect populations and species in various ways. Rising temperatures can shift geographical distributions and lead to (phenotypic or genetic) changes in traits, mostly phenology, which may affect demography. Most of these effects are well documented in birds. For example, the
Brian Roy Lockhart
Many years ago, I had an epiphany that I would like to share. Several students and I were installing research plots in the forests on Pittman Island, Issaquena County, Mississippi, an island adjacent to the Mississippi River, near the borders of Arkansas, Mississippi, and Louisiana. While eating lunch, we watched a bird, more specifically a prothonotary warbler (
... domesticated birds. If possible, avoid rural areas, small farms and open-air markets. Wash your hands. This is one of the simplest and best ways to prevent infections of all kinds. Use an alcohol-based hand sanitizer containing at least 60 percent alcohol ...
http://www.ias.ac.in/article/fulltext/reso/008/07/0022-0032. Keywords. Birds. nesting. territory; coloniality; heronries. ecology; nesting strategies. Author Affiliations. Abdul Jamil Urfi1. Department of Environmental Biology, School of Environmental Studies, University of Delhi, Delhi 110007. Resonance – Journal of Science ...
Dyke, Gareth J.; Dortangs, Rudi W.; Jagt, John W.; Mulder, Eric W. A.; Schulp, Anne S.; Chiappe, Luis M.
Birds known from more than isolated skeletal elements are rare in the fossil record, especially from the European Mesozoic. This paucity has hindered interpretations of avian evolution immediately prior to, and in the aftermath of, the Cretaceous-Tertiary (K-T) extinction event. We report on a
Crandall, Lee S.
When CHARLES CORDIER arrived from Costa Rica on October 9, 1942, bringing with him, among other great rarities, three Bare-necked Umbrella Birds (Cephalopterus ornatus glabricollis), it seemed to us that the mere possession of such fabulous creatures was satisfaction enough. True, they were not
Introducing students to different hawks and owls found in Wisconsin and building a basis for appreciation of these birds in their own environment is the purpose of this teacher's guide. Primarily geared for upper elementary and junior high grades, the concepts presented could be used in conjunction with the study of ecology. A filmstrip is…
The extremely long migration flights of some birds are carried out in one hop, necessitating a substantial prior build-up of fat fuel. We summarize the basic elements of bird flight physics with a simple model, and show how the fat reserves influence flight distance, flight speed and the power expended by the bird during flight. (paper)
This feature article presents the basics of birding, or bird watching, and discusses its appeal, especially to serious birders. A section on "citizen scientists" explains organizations that collect data on birds and describes projects they organize. Other sections discuss the legacy of John James Audubon and the bald eagle.
This copiously illustrated document is designed to be a field quide to birds of prey that are common to Wisconsin, as well as to some that enter the state occasionally. An introduction discusses birds of prey with regard to migration patterns, the relationship between common names and the attitudes of people toward certain birds, and natural signs…
The extremely long migration flights of some birds are carried out in one hop, necessitating a substantial prior build-up of fat fuel. We summarize the basic elements of bird flight physics with a simple model, and show how the fat reserves influence flight distance, flight speed and the power expended by the bird during flight.
Griffith, E.M.; Paytan, A.; Eisenhauer, A.; Bullen, T.D.; Thomas, E.
During the Eocene-Oligocene transition (EOT, ca. 34 Ma), Earth's climate cooled significantly from a greenhouse to an icehouse climate, while the calcite (CaCO3) compensation depth (CCD) in the Pacific Ocean increased rapidly. Fluctuations in the CCD could result from various processes that create an imbalance between calcium (Ca) sources to, and sinks from, the ocean (e.g., weathering and CaCO3 deposition), with different effects on the isotopic composition of dissolved Ca in the oceans due to differences in the Ca isotopic composition of various inputs and outputs. We used Ca isotope ratios (??44/40Ca) of coeval pelagic marine barite and bulk carbonate to evaluate changes in the marine Ca cycle across the EOT. We show that the permanent deepening of the CCD was not accompanied by a pronounced change in seawater ??44/40Ca, whereas time intervals in the Neogene with smaller carbonate depositional changes are characterized by seawater ??44/40Ca shifts. This suggests that the response of seawater ??44/40Ca to changes in weathering fluxes and to imbalances in the oceanic alkalinity budget depends on the chemical composition of seawater. A minor and transient fluctuation in the Ca isotope ratio of bulk carbonate may reflect a change in isotopic fractionation associated with CaCO3 precipitation from seawater due to a combination of factors, including changes in temperature and/or in the assemblages of calcifying organisms. ?? 2011 Geological Society of America.
Ricardo Pérez-de la Fuente
Full Text Available A new fossil fly species, Acartophthalmites willii sp. n. (Diptera: Acalyptratae: Opomyzoidea from Baltic amber (Eocene, 56−33.9 Ma, is described based on a male originally assigned by Hennig (1969 to A. tertiaria Hennig, 1965, who erroneously also referred to it in the same work as “A. electrica Hennig” (unavailable name. The new species, representing the third named species of the extinct genus with unclear familial relationships Acartophthalmites Hennig, 1965, is herein described and illustrated in detail, and its systematic implications and relationships are discussed. From the morphological standpoint, the new species represents an intermediate form between the two formerly described species within the genus, therefore expanding the character combination diversity in this lineage of acalyptrate flies. The genus Acartophthalmites is considered to be most closely related to Clusiidae and, therefore, it is herein tentatively classified within the superfamily Opomyzoidea. The current work takes part of an effort to review the Acartophthalmites diversity in order to gain knowledge on the morphological data from the specimens described within the genus and ultimately enable a reliable analysis of its phylogenetic relationships with other acalyptrates.
Full Text Available The upper Eocene Youganwo Formation in Maoming Province, Guangdong Province, China, produced abundant remains of turtles and crocodiles, but mammalian remains are rare. The previously found mammals from Youganwo Formation include a nimravid carnivore and an amynodontid perissodactyl. Here we report on a new brontotheriid perissodactyl from the Youganwo Formation. Maobrontops paganus gen. et sp. nov. is described based on a maxillary fragment with P4 and M1–2 (SYSU-M-4. Maobrontops paganus gen. et sp. nov. is characterized by the combination of a simple P4 without a hypocone with derived molars having large molar fossae and large anterolingual cingular cusp. A parsimony implied weighting character analysis places Maobrontops gen. nov. in the clade Embolotheriita as a sister taxon to the terminal subclade containing Nasamplus, Protembolotherium, and Embolotherium. Maobrontops paganus gen. et sp. nov. is one of the largest Asian brontotheres. The brontotheriid fauna of South China is endemic and includes at least three valid taxa: Dianotitan from Brontotheriita and Pygmaetitan and Maobrontops gen. nov. from Embolotheriita.
... single bird, the single largest medium bird which can enter the inlet, and the large flocking bird must...) (d) Large flocking bird. An engine test will be performed as follows: (1) Large flocking bird engine.... (4) Ingestion of a large flocking bird under the conditions prescribed in this paragraph must not...
Coxall, Helen; Bohaty, Steve; Wilson, Paul; Liebrand, Diederik; Nyberg, Anna; Holmström, Max
Integrated Ocean Drilling Program (IODP) Expedition 342 drilled sediment drifts on the Newfoundland margin to recover high-resolution records of North Atlantic ocean-climate history and track the evolution of the modern climate system through the Late Cretaceous and Early Cenozoic. An early Paleogene deep-sea benthic stable isotope composite record from multiple Exp. 342 sites is currently in development and will provide a key reference section for investigations of Atlantic and global climate dynamics. This study presents initial results for the late Eocene slice of the composite from Site U1411, located at mid depth (˜2850m Eocene paleodepth) on the Southeast Newfoundland Ridge. Stable oxygen (δ18O) and carbon (δ13C) isotope ratios were measured on 640 samples hosting exceptionally well-preserved epifaunal benthic foraminifera obtained from the microfossil-rich uppermost Eocene clays at 4cm spacing. Sedimentation rates average 2-3 cm/kyr through the late Eocene, such that our sampling resolution is sufficient to capture the dominant Milankovitch frequencies. Late Eocene Site U1411 benthic δ18O values (1.4 to 0.5‰ VPDB) are comparable to the Pacific and elsewhere in the Atlantic at similar depths; however, δ13C is lower by ˜0.5 ‰ with values intermediate between those of the Southern Labrador Sea to the north (-1 to 0) and mid latitude/South Atlantic (0.5 to 1.5) to the south, suggesting poorly ventilated bottom waters in the late Eocene North Atlantic and limited production of North Atlantic deep water. Applying the initial shipboard magneto-biostratigraphic age framework, the Site U1411 benthic δ13C and δ18O records display clear cyclicity on orbital timescales. Spectral analysis of the raw unfiltered datasets identifies eccentricity (400 and 100 kyr), obliquity (40 kyr) and precession (˜20 kyr) signals imprinted on our time series, revealing distinct climatic heart beats in the late Eocene prior to the transition into the 'ice house'.
Steinthorsdottir, Margret; Porter, Amanda S.; Holohan, Aidan; Kunzmann, Lutz; Collinson, Margaret; McElwain, Jennifer C.
A unique stratigraphic sequence of fossil leaves of Eotrigonobalanus furcinervis (extinct trees of the beech family, Fagaceae) from central Germany has been used to derive an atmospheric pCO2 record with multiple data points spanning the late middle to late Eocene, two sampling levels which may be earliest Oligocene, and two samples from later in the Oligocene. Using the inverse relationship between the density of stomata and pCO2, we show that pCO2 decreased continuously from the late middle to late Eocene, reaching a relatively stable low value before the end of the Eocene. Based on the subsequent records, pCO2 in parts of the Oligocene was similar to latest Eocene values. These results suggest that a decrease in pCO2 preceded the large shift in marine oxygen isotope records that characterizes the Eocene-Oligocene transition and that when a certain threshold of pCO2 change was crossed, the cumulative effects of this and other factors resulted in rapid temperature decline, ice build up on Antarctica and hence a change of climate mode.
Full Text Available This study of the Paleocene–Eocene boundary within a foreland basin of southern Tibet, which was dominated by a carbonate ramp depositional environment, documents more complex environmental conditions than can be derived from studies of the deep oceanic environment. Extinction rates for larger foraminiferal species in the Zongpu-1 Section apply to up to 46% of the larger foraminiferal taxa. The extinction rate in southern Tibet is similar to rates elsewhere in the world, but it shows that the Paleocene fauna disappeared stepwise through the Late Paleocene, with Eocene taxa appearing abruptly above the boundary. A foraminifera turnover was identified between Members 3 and 4 of the Zongpu Formation—from the Miscellanea–Daviesina assemblage to an Orbitolites–Alveolina assemblage. The Paleocene and Eocene boundary is between the SBZ 4 and SBZ 5, where it is marked by the extinction of Miscellanea miscella and the first appearance of Alveolina ellipsodalis and a large number of Orbitolites. Chemostratigraphically, the δ13C values from both the Zongpu-1 and Zongpu-2 Sections show three negative excursions in the transitional strata, one in Late Paleocene, one at the boundary, and one in the early Eocene. The second negative excursion of δ13C, which is located at the P–E boundary, coincides with larger foraminifera overturn. These faunal changes and the observed δ13C negative excursions provide new evidence on environmental changes across the Paleocene–Eocene boundary in Tibet.
Migratory movements of birds has always fascinated man and led to many questions concerning the ecological drivers behind, the necessary adaptations and the navigational abilities required. However, especially for the long-distance migrants, basic descriptions of their movements are still lacking...... and a forest reserve. In the degraded habitat all species used more space, although the consequence on bird density is less clear. Two manuscripts relate the migratory movements of a long-distance migrant with models of navigation. One compares model predictions obtained by simulation with actual movements......, and conclude that the currently believed theoretical framework is insufficient to explain the observed performance. The other study investigates the ability of a displaced experienced migrant to navigate back to the normal migration route. It documents the capability, but also finds interesting patterns...
Frieling, Joost; Peterse, Francien; Lunt, Daniel; Bohaty, Steven; Sinninghe Damste, Jaap; Reichart, Gert-Jan; Sluijs, Appy
The Paleocene-Eocene Thermal Maximum (PETM; 56 Ma) was a period of rapid 4-5°C global warming and a global negative carbon isotope excursion (CIE) of 3-4.5‰, signaling the input of at least 1500 Gt of δ13C-depleted carbon into the ocean-atmosphere system. Carbon cycle modeling has indicated that the shape and magnitude of this CIE are generally explained by a large and rapid initial pulse, followed by 50 kyr of 13C-depleted carbon injection. Crucially, some of suggested carbon sources, e.g. submarine methane hydrates and permafrost thawing, may respond to warming and act as positive carbon cycle feedbacks on millennial timescales. Previous analyses showed that warming started prior to the CIE at some high and mid-latitude sites, but is still unknown whether this is a global signal and timing and magnitude of such a warming remain poorly constrained. We generated a new high-resolution TEX86 and δ13C record from Ocean Drilling Program Site 959 in the eastern tropical Atlantic and find that initial warming preceded the PETM CIE by 10 kyr. Moreover, cross-correlation functions on these new and published temperature-δ13C data imply that substantial (2-3 °C) warming lead 13C-depleted carbon injection by an average of 2-3 kyr globally. Finally, a data compilation shows that global burial fluxes of non-detrital Ba approximately tripled across all depths of the ocean studied, which on PETM time scales can only be explained by significant Ba addition to the oceans. Submarine hydrates are Ba-rich and require warming to dissociate. The simplest explanation for the temperature lead and Ba addition to the ocean is that methane hydrates dissociated as a response to initial warming and acted as a positive carbon cycle feedback during the PETM. The attribution of the CIE to a carbon cycle feedback naturally leads to the question what caused the early warming. The absence of a 13C change during the early warming implies a 13C-neutral CO2 source, such as volcanism, is the most
Full Text Available The Eocene flysh in the region of Split (Dalmatia, Croatia is char¬acterized by the presence of layers with different characteristics. It mainly includes thin-layered marls, clayey marls, calcareous marls, clastic lay¬ered limestones, calcarenites and breccias. Those parts that can be de¬scribed as the soft rocks or hard clays by the mechanical means, exposed to weathering reduce the durability within "an engineering time scale". The paper deals with the factors that influence the weathering process. The analyzed weathering is a combination of processes acting simulta¬neously. Most of these processes depend on the change of the water con¬tent, thus the weathering process mainly develops when a material is subjected to the wetting-drying process, On the base of these results form of degradation process is modeled. The weathering process can be main¬ly described as physical weathering combined with chemical weathering on the free surfaces and on the cracks walls. Erosion as a result of weath¬ering, is the dominant geomorphic process on analyzed flysch terrain. According to the analysis, as the most appropriate due to the characteris¬tics the tests are chosen as index properties. Some of these tests are modified in order to adapt them to the determined characteristics of ma¬terials from flysch layers. The correlations between the measured values are used as the basis for the classification proposal of the analyzed mate¬rial, according to its resistance to weathering processes. Roughly, three main groups of samples are recognizable: the first one with carbonate content more then 90% is not weathered at the engineers time scale; the second group with carbonate content from 75% to 90% include samples susceptible to weathering in engineers time scale; the third group with carbonate content less then 75% include samples in which the weather¬ing occurs immediately after the exposition to the weathering factors.
Weijers, Johan W. H.; Schouten, Stefan; Sluijs, Appy; Brinkhuis, Henk; Sinninghe Damsté, Jaap S.
The Palaeocene-Eocene Thermal Maximum (PETM; ˜55.5 Ma) is a geologically relatively brief episode of extreme warmth. Both deep and surface ocean temperatures increased by up to 5 °C in equatorial waters and up to 8 °C in mid and high latitude waters. From the continents, the annual mean air temperature response during the PETM is still largely unknown, mainly due to a lack of quantitative temperature proxies and sufficient suitable, continuous high resolution records. Recently, a new proxy for continental temperature reconstructions has been proposed, based on the distribution of membrane lipids of bacteria in present-day soils [ J.W.H. Weijers, S. Schouten, J.C. van den Donker, E.C. Hopmans, J.S. Sinninghe Damsté (2007) Environmental controls on bacterial tetraether membrane lipid distribution in soils, Geochimica et Cosmochimica Acta 71, 703-713] and shown to reconstruct annual mean air temperature. In this study we applied this new proxy in an attempt to reconstruct the air temperature in high latitude continental areas during the PETM by analysis of a marine sedimentary sequence obtained from the Lomonosov Ridge in the central Arctic Ocean (Integrated Ocean Drilling Program Expedition 302, Site 004, Hole A). The results indicate a warming of ˜8 °C above background values of ˜17 °C. This warming is coincident with a similar rise in sea surface temperatures documented earlier. Our results thus further confirm the warm conditions in the Arctic, and point to a strongly reduced latitudinal temperature gradient during the PETM.
Kerstin K Zander
Full Text Available Threatened species programs need a social license to justify public funding. A contingent valuation survey of a broadly representative sample of the Australian public found that almost two thirds (63% supported funding of threatened bird conservation. These included 45% of a sample of 645 respondents willing to pay into a fund for threatened bird conservation, 3% who already supported bird conservation in another form, and 15% who could not afford to pay into a conservation fund but who nevertheless thought that humans have a moral obligation to protect threatened birds. Only 6% explicitly opposed such payments. Respondents were willing to pay about AUD 11 annually into a conservation fund (median value, including those who would pay nothing. Highest values were offered by young or middle aged men, and those with knowledge of birds and those with an emotional response to encountering an endangered bird. However, the prospect of a bird going extinct alarmed almost everybody, even most of those inclined to put the interests of people ahead of birds and those who resent the way threatened species sometimes hold up development. The results suggest that funding for threatened birds has widespread popular support among the Australian population. Conservatively they would be willing to pay about AUD 14 million per year, and realistically about AUD 70 million, which is substantially more than the AUD 10 million currently thought to be required to prevent Australian bird extinctions.
Roč. 67, 3/4 (2011), s. 149-156 ISSN 0036-5343 Institutional research plan: CEZ:AV0Z30130516 Keywords : fossil fish fauna * Osteichthyes * scales * morphology * Paleogene * Upper Eocene * Upper Eocene (Czech Republic) Subject RIV: DB - Geology ; Mineralogy http://www.nm.cz/publikace/archiv-en.php?id=4&rok=67&kcislu=3-4&f_=Show
Douglas, Peter M J; Affek, Hagit P; Ivany, Linda C; Houben, Alexander J P; Sijp, Willem P; Sluijs, Appy; Schouten, Stefan; Pagani, Mark
Paleoclimate studies suggest that increased global warmth during the Eocene epoch was greatly amplified at high latitudes, a state that climate models cannot fully reproduce. However, proxy estimates of Eocene near-Antarctic sea surface temperatures (SSTs) have produced widely divergent results at similar latitudes, with SSTs above 20 °C in the southwest Pacific contrasting with SSTs between 5 and 15 °C in the South Atlantic. Validation of this zonal temperature difference has been impeded by uncertainties inherent to the individual paleotemperature proxies applied at these sites. Here, we present multiproxy data from Seymour Island, near the Antarctic Peninsula, that provides well-constrained evidence for annual SSTs of 10-17 °C (1σ SD) during the middle and late Eocene. Comparison of the same paleotemperature proxy at Seymour Island and at the East Tasman Plateau indicate the presence of a large and consistent middle-to-late Eocene SST gradient of ∼7 °C between these two sites located at similar paleolatitudes. Intermediate-complexity climate model simulations suggest that enhanced oceanic heat transport in the South Pacific, driven by deep-water formation in the Ross Sea, was largely responsible for the observed SST gradient. These results indicate that very warm SSTs, in excess of 18 °C, did not extend uniformly across the Eocene southern high latitudes, and suggest that thermohaline circulation may partially control the distribution of high-latitude ocean temperatures in greenhouse climates. The pronounced zonal SST heterogeneity evident in the Eocene cautions against inferring past meridional temperature gradients using spatially limited data within given latitudinal bands.
Slotnick, B.S.; Dickens, G.R.; Nicolo, M.J.; Hollis, C.J.; Crampton, J.S.; Zachos, J.C.; Sluijs, A.
The late Paleocene to early Eocene was marked by major changes in Earth surface temperature and carbon cycling. This included at least two, and probably more, geologically brief (<200-k.yr.) intervals of extreme warming, the Paleocene-Eocene thermal maximum (PETM) and the Eocene thermal maximum-2
Wood, A E
The earliest known representative of the fundamentally South American and African hystricognathous rodents has recently been found in the middle or late Eocene of southwestern Texas; this discovery supports the postulate of a northern and independent origin for the two southern groups and increases the evidence against mid-Tertiary trans-Atlantic migration of these rodents at a time when the South Atlantic was narrower than it is at present. The fossil seems to be related to the North American Eocene family Sciuravidae.
A general deterioration of terrestrial climate took place during middle Eocene to earliest Oligocene time in southern California and in the Gulf Coast. Pollen data, calibrated by calcareous nannofossil ages, indicate four events of rapid floral and/or vegetational change among angiosperms during this time interval. The events can be correlated between the two regions even though these regions lay within different floristic provinces, and each event of angiosperm change is interpreted to indicate a pulse of rapid climatic shift. The most distinct of these events is the Middle Eocene Diversity Decline, which resulted from a peak in last appearances (extinctions, emigrations) centered in the early Bartonian. -from Author
Williams, T. C.; Williams, J. M.
Observations of bird migration with NASA radars were made at Wallops Island, Va. Simultaneous observations were made at a number of radar sites in the North Atlantic Ocean in an effort to discover what happened to those birds that were observed leaving the coast of North America headed toward Bermuda, the Caribbean and South America. Transatlantic migration, utilizing observations from a large number of radars is discussed. Detailed studies of bird movements at Wallops Island are presented.
Doyle, P.A.T.; Gorman, M.L.; Patterson, I.J.; Howe, S.
This paper reports that the creation of a club for the purpose of encouraging oil and gas workers to watch birds may not at first seem a viable proposition. To the layperson, birds offshore conjures up an image of hundreds of seagulls following fishing boats, and very little else. Also, the act of birdwatching is not seen as a typical offshore worker's activity. Anyone who has worked on an installation offshore and who has any interest in wildlife will be aware of the occasional presence of land-birds. Two decades ago, prompted by some keen offshore workers, a single oil company set up a monitoring program, which quickly became popular with a number of its employees. Birds seem offshore were recorded on data forms and collected together. At this stage the club was purely another recreation facility; however, when the data were collated it was soon realized that installations offshore were being used as staging posts by birds on migration, and that the information being collected would be of great interest in the study of bird movements. All over Britain, at strategic points on the coastline, there are bird observatories which record the arrival and departure of migrating birds. The presence of several hundred solid structures up and down the North Sea, which are used by birds en route, represents a huge, unique bird observatory, capable of uncovering facts about bird migration which have long eluded land-based scientists. Eleven years ago, the North Sea Bird Club began, composed of eight member companies, a recorder from Aberdeen University and a representative from the Nature Conservancy Council. The club received data from 41 installations, and the recorder collated these on Aberdeen University's computer and produced an annual report of sightings
Anders Pape Møller; Wei Liang
The life history of tropical birds differs from that of their temperate counterparts by late start of reproduction, small clutch sizes, and high rates of adult survival. Thus, tropical species should have greater residual reproductive value than temperate species. Therefore, tropical birds can be predicted to take smaller risks than closely related temperate birds in order not to jeopardize their prospects of survival, which is the single most important component of fitness, and which is grea...
National Oceanic and Atmospheric Administration, Department of Commerce — This data set contains sensitive biological resource data for wading birds, shorebirds, waterfowl, raptors, diving birds, pelagic birds, passerine birds, gulls and...
National Oceanic and Atmospheric Administration, Department of Commerce — This data set contains sensitive biological resource data for alcids, diving birds, gulls, terns, passerine birds, pelagic birds, raptors, shorebirds, wading birds,...
National Oceanic and Atmospheric Administration, Department of Commerce — This data set contains sensitive biological resource data for diving birds, gulls, terns, passerine birds, pelagic birds, raptors, shorebirds, wading birds,...
National Oceanic and Atmospheric Administration, Department of Commerce — This data set contains sensitive biological resource data for diving birds, gulls, terns, passerine birds, pelagic birds, raptors, shorebirds, wading birds, and...
Schumann, G.; Manz, W.; Reitner, J.; Lustrino, M.
Little is known about the manifold life forms of the deep biosphere although there is increasing scientific evidence that an extensive biosphere does exist in extreme environments such as the rocks below the seafloor. The ODP Leg 200 was devoted to the study of Eocene oceanic crust of the North Pacific Ocean. Within a massive tholeiitic lava flow unit, at depth of 51 mbsf underneath a water column of about 5000 m, we found unique filamentous structures. Based on morphological traits like branching, septa and central pores the filaments are interpreted as fungi. These filaments were found within carbonate-filled vesicles ranging in size from 0.5 to 3 mm in diameter. The net of fungal hyphae completely fills the whole pore space from the basalt-carbonate boundary towards the center of the pores. The cross section dimension of these filaments is about 5-10 micrometer and the length differ from 50 to several hundreds micrometer. Thereby the cell septa of the hyphae are clearly visible. The number of hyphae ranges from some tenth to some hundreds per particular pore. The presence of pyrite within the carbonate cements points out anaerobic conditions in this habitat. After removing the carbonate by etching the vesicles with diluted formic acid, the 3-dimensional structure of the fungus could be clearly visualized. Fine structure analysis of the hyphae obtained by field emission scanning electron microscopy (FE-SEM) revealed a network of tiny small fibers coating the surface of the hyphae. Semi-quantitative chemical analyses of the etched hyphae were conducted with an energy dispersive spectrometer system (EDS) coupled with the FE-SEM. The results evidence a chemical composition of the hyphae different from the surrounding carbonate matrix. Undisturbed filamentous growth through different calcite crystals within the vesicles and small open space between the fungi and matrix indicate endolithic fungal growth after the calcium carbonate filling of the vesicles. To the best
Minwer-Barakat, Raef; Marigó, Judit; Femenias-Gual, Joan; Moyà-Solà, Salvador
The Eocene was one of the warmest epochs of the Cenozoic and documented the first occurrence of several orders of modern mammals. Among them, Euprimates underwent a very important radiation favored by the development of dense forests throughout the Northern Hemisphere. Two main groups reached a great abundance and diversity during the Eocene, Adapiformes and Omomyiformes, which are related to the main clades of living primates (strepsirrhines and haplorhines, respectively). In the Iberian Peninsula, Eocene primates have been known since the 1960s, when several fossil sites containing prosimian remains were discovered. Nevertheless, it was not until 2010 that the research on Eocene primates from Spain has increased strikingly, and the results achieved in this last stage have surpassed those of the whole past century in terms of number of publications. Besides some interesting findings in the Ebro, Almazán and Miranda-Trebiño basins, the Pyrenees have yielded the most abundant record of Eocene primates from the Iberian Peninsula, constituting therefore an excellent region for evaluating the evolution of primates through this epoch. In the early Eocene continental deposits of the Àger area, adapiforms are well represented, with three species of the genus Agerinia. Besides, the only record of Plesiadapiformes (archaic primates) from Spain has been documented in this zone. The middle Eocene is particularly well represented in the Eastern Pyrenees. In the section of Sant Jaume de Frontanyà, three primate species have been described in the last years. The adapiform Anchomomys frontanyensis and the omomyiform Pseudoloris pyrenaicus, found in the oldest levels of the section, and the omomyiform Necrolemur anadoni, identified in the youngest levels, have allowed reconstructing the relationships of these taxa with their correlatives found in other parts of Europe. Late Eocene deposits with mammal remains crop out in the area of La Pobla de Segur. The most relevant fossil
... resultant intensive agricultural projects that follow. Such impacts have resulted in a decline in both bird habitats and biodiversity. Of particular concern are the areas important to all our endemic bird species, and already we are witnessing a series of very worrying developments. Taking each. Scopus 32: 27–34, June 2013 ...
Walter, Ingo; Briess, Klaus; Baerwald, Wolfgang; Skrbek, Wolfgang; Schrandt, Fredrich
For hot spot events as forest fires, volcanic activity or burning oil spills and coal seams a dedicate dspace instrumentation does not exist. With its successful launch end of October 2001 with the Indian Polar Satellite Launch Vehicle the German Aerospace Center starts closing this gap with the micro-satellite mission BIRD. As space segment serves a three-axis stabilized satellite of 92 kg including a contingent of over 30% for the scientific instruments. The main payload of the BIRD micro-satellite is the newly developed Hot Spot Recognition System. It's a dual-channel instrument for middle and thermal IR imagery based on cooled MCT line detectors. The miniaturization by integrated detector/cooler assemblies provides a highly efficient design. A complement for the hot spot detection is the wide-angle stereo-scanner WAOSS-B. It is a hardware re-use dedicated to vegetation and cloud assessment in the visible spectral range. Besides the main objective of hot spot detection the mission has to answer several technological questions of the operation of cooled detectors in space, special aspects of their adaptation to the satellite platform as well as their calibration.
Antoine, Pierre-Olivier; Marivaux, Laurent; Croft, Darin A.; Billet, Guillaume; Ganerød, Morgan; Jaramillo, Carlos; Martin, Thomas; Orliac, Maëva J.; Tejada, Julia; Altamirano, Ali J.; Duranthon, Francis; Fanjat, Grégory; Rousse, Sonia; Gismondi, Rodolfo Salas
The long-term isolation of South America during most of the Cenozoic produced a highly peculiar terrestrial vertebrate biota, with a wide array of mammal groups, among which caviomorph rodents and platyrrhine primates are Mid-Cenozoic immigrants. In the absence of indisputable pre-Oligocene South American rodents or primates, the mode, timing and biogeography of these extraordinary dispersals remained debated. Here, we describe South America's oldest known rodents, based on a new diverse caviomorph assemblage from the late Middle Eocene (approx. 41 Ma) of Peru, including five small rodents with three stem caviomorphs. Instead of being tied to the Eocene/Oligocene global cooling and drying episode (approx. 34 Ma), as previously considered, the arrival of caviomorphs and their initial radiation in South America probably occurred under much warmer and wetter conditions, around the Mid-Eocene Climatic Optimum. Our phylogenetic results reaffirm the African origin of South American rodents and support a trans-Atlantic dispersal of these mammals during Middle Eocene times. This discovery further extends the gap (approx. 15 Myr) between first appearances of rodents and primates in South America. PMID:21993503
Frieling, J.|info:eu-repo/dai/nl/338017909; Huurdeman, Emiel; Rem, Charlotte; Donders, T.H.|info:eu-repo/dai/nl/290469872; Pross, Jorg; Bohaty, Steven M.; Holdgate, Guy; Gallagher, Stephen; McGowran, Brian; Bijl, P.K.|info:eu-repo/dai/nl/314028110
Detailed, stratigraphically well-constrained environmental reconstructions are available for Paleocene and Eocene strata at a range of sites in the southwest Pacific Ocean (New Zealand and East Tasman Plateau; ETP) and Integrated Ocean Discovery Program (IODP) Site U1356 in the south of the
Garassino, Alessandro; Angeli, De Antonio
Recently, Beschin et al. (2001) have described two specimens of palinurids, discovered at Chiampo (“Albanello” quarry, Vicenza, northern Italy). Preserved three dimensionally, these were collected from limestones of Lutetian (middle Eocene) age. The peculiar ornament of the dorsal carapace surface,
Sahy, D.; Condon, D.J.; Terry, D.O., Jr.; Fischer, A.U.; Kuiper, K.F.
Records of terrestrial environmental change indicate that continental cooling and/or aridification may have predated the greenhouse-icehouse climate shift at the Eocene-Oligocene transition (EOT) by ca. 600 kyr. In North America, marine-terrestrial environmental change asynchronicity is inferred
Bieńkowska-Wasiluk, Małgorzata; Bonde, Niels Christensøn; Møller, Peter Rask
to the perciform suborder Notothenioidei. This group is almost unknown as fossils. Similarities to the living, ‘primitive’ nototheniid Dissostichus eleginoides are indicated in the dentition. Gadiform evolution in the Paleocene-Eocene, and the possibility of a correlation between the origin and evolution...
Přikryl, Tomáš; Skupien, P.
Roč. 268, č. 1 (2013), s. 113-123 ISSN 0077-7749 Institutional research plan: CEZ:AV0Z30130516 Institutional support: RVO:67985831 Keywords : Elasmobranchii * Leidybatis jugossus * Centrophorus * Eocene * Western Carpathian Subject RIV: DB - Geology ; Mineralogy Impact factor: 0.541, year: 2013
Studying past intervals of abrupt global warming and massive carbon release can improve our knowledge in ways relevant to understanding future climate change. Possible paleo-analogues for future climate change are the early Paleogene hyperthermal events, such as the Paleocene-Eocene Thermal Maximum
Smith, V.; Warny, S.; Bralower, T. J.; Jones, H.; Lowery, C. M.; Smit, J.; Vajda, V.; Vellekoop, J.; 364 Scientists, E.
International Ocean Discovery Program (IODP) Expedition 364, with support from the International Continental Scientific Drilling Program, cored through Paleocene and Eocene sediments and into the impact structure of the Chicxulub impact crater. Three palynological studies of the post-impact section are currently underway. The two other studies are investigating the dinoflagellate palynology and terrestrial palynology of the K/Pg boundary section, while this study focuses on the early Eocene terrestrial palynology of the IODP 364 core, which has yielded a diverse and well preserved pollen assemblage. A few samples from the Early Paleocene have also been examined but organic microfossil preservation is quite poor. Samples from this core are the oldest palynological record from the Yucatan peninsula. Sample preparation and detailed abundance counts of sixty samples throughout the post-impact section are in progress, with a particular focus on the Paleocene-Eocene Thermal Maximum (PETM) and the Early Eocene Climatic Optimum (EECO). Terrestrial palynomorph assemblages will be used to reconstruct paleoclimatological conditions throughout this time period. Floral response to hyperthermal events in the IODP 364 core will be compared with records from other Gulf of Mexico and Caribbean sections. In addition to the biological and paleoclimatological implications of this research, age control from foraminiferal and nannofossil biostratigraphy, paleomagnetism, and radiometric dating will provide a chronological framework for the terrestrial pollen biostratigraphy, with applications to hydrocarbon exploration in the Wilcox Formation and age equivalent sections in the Gulf of Mexico.
Advokaat, Eldert; van Hinsbergen, D.J.J.|info:eu-repo/dai/nl/269263624; Maffione, M.; Langereis, C.G.|info:eu-repo/dai/nl/073584223; Vissers, R.L.M.|info:eu-repo/dai/nl/068789203; Cherchi, A.; Schroeder, R.; Madani, H.; Columbu, S.
Key to understanding the complex Mediterranean subduction history is the kinematic reconstruction of its paleogeography after Jurassic extension between Iberia, Eurasia, and Africa. While post-Eocene Liguro-Provençal back-arc extension, and associated Miocene ~50° counterclockwise (ccw) rotation of
The early Eocene represents an ideal case study to analyse the impact of enhanced global warming on the ocean-atmosphere system and the relationship between carbon cycling and climate. During this time interval, the Earth’s surface experienced a long-term warming trend that culminated in a period of
Barron, John A.; Bukry, David B.; Gersonde, Rainer
Abundant and well-preserved diatoms and silicofl agellate assemblages are documented through a complete late Eocene sequence, ODP Hole 1090B, recovered from the southern Agulhas Ridge in the sub-Antarctic South Atlantic. A sequence of Cestodiscus (diatom) species occurrence events involving C. pulchellus var. novazealandica, C. fennerae, C. antarcticus, C. convexus, C. trochus, and C. robustus is tied with paleomagnetic stratigraphy and provides the basis of proposing a new diatom zonation for the latest middle Eocene to early Oligocene (~37.6–33.4 Ma) of the sub-Antarctic South Atlantic. Comparison with previously published diatom occurrence charts suggested this zonation should be applicable throughout the low latitude regions of the world’s oceans. Silicofl agellates belong to the Dictyocha hexacantha and the overlying Corbisema apiculata Zones. The late Eocene succession of silicofl agellate species is dominated by Naviculopsis (20–60%). Naviculopsis constricta and N. foliacea dominate the D. hexacantha Zone, followed by the N. constricta, then N. biapiculata in the C. apiculata Zone. Cold-water Distephanus is most abundant in the latest Eocene along with N. biapiculata. The tops of zonal guide fossils Dictyocha hexacantha and Hannaites quadria (both 36.6 Ma) and Dictyocha spinosa (37.1 Ma) are tied with paleomagnetic stratigraphy.
The Paleocene and Eocene are characterized by strong greenhouse climates. Atmospheric CO2 concentrations and global temperatures were much higher than today. The period from 60 to 50 million years ago (Ma) is marked by a gradual warming trend of ~8 ºC in the deep ocean. The interval from 56 to 50 Ma
Lauretano, V.; Littler, K.; Polling, M.; Zachos, J.C.; Lourens, L.J.
Recent studies have shown that the Early Eocene Climatic Optimum (EECO) was preceded by a series of short-lived global warming events, known as hyperthermals. Here we present high-resolution benthic stable carbon and oxygen isotope records from ODP Sites 1262 and 1263 (Walvis Ridge, SE Atlantic)
The results described in this thesis provide a rather complex picture of climatic, environmental and biotic changes preceding and arising from the onset of Antarctic glaciation. This period is commonly known as the greenhouse to icehouse transition across the Eocene-Oligocene Transition (EOT, 34-33
Stickley, C.E.; Koç, N.; Pearce, R.B.; Kemp, A.E.S.; Jordan, R.W.; Sangiorgi, F.; St. John, K.
Finely laminated Middle Eocene sediments from the central Arctic contain high abundances of the delicate, sea ice–dwelling fossil diatoms Synedropsis spp. and sea ice–rafted debris (sea ice–IRD), establishing an offshore seasonal sea ice regime ca. 47 Ma. Synedropsis spp. co-occur with other
Sun, Jimin; Ni, Xijun; Bi, Shundong; Wu, Wenyu; Ye, Jie; Meng, Jin; Windley, Brian F
The Eocene-Oligocene Boundary (~34 million years ago) marks one of the largest extinctions of marine invertebrates in the world oceans and of mammalian fauna in Europe and Asia in the Cenozoic era. A shift to a cooler climate across this boundary has been suggested as the cause of this extinction in the marine environment, but there is no manifold evidence for a synchronous turnover of flora, fauna and climate at the Eocene-Oligocene Boundary in a single terrestrial site in Asia to support this hypothesis. Here we report new data of magnetostratigraphy, pollen and climatic proxies in the Asian interior across the Eocene-Oligocene Boundary; our results show that climate change forced a turnover of flora and fauna, suggesting there was a change from large-size perissodactyl-dominant fauna in forests under a warm-temperate climate to small rodent/lagomorph-dominant fauna in forest-steppe in a dry-temperate climate across the Eocene-Oligocene Boundary. These data provide a new terrestrial record for this significant Cenozoic environmental event.
Handley, L.; O'Halloran, A.; Pearson, P.N.; Hawkins, E.; Nicholas, C.J.; Schouten, S.; McMillan, I.K.; Pancost, R.D.
The Paleocene-Eocene Thermal Maximum (PETM), at ca. 55.8 Ma, is one of the most studied instances of past greenhouse gas-induced global warming. As such, it provides a rich opportunity to examine the impact of such global change on local climates. The effects of increased continental and sea surface
MINDA, Haruya; FURUZAWA, Fumie A.; SATOH, Shinsuke; NAKAMURA, Kenji
A C-band polarimetric radar on Okinawa Island successfully observed large-scale bird migrations over the western Pacific Ocean. The birds generated interesting polarimetric signatures. This paper describes the signatures and speculates bird behavior.
Our knowledge concerning the periodical movements in animals called migrations is chiefly based on observations on birds. By and by, however, a number of facts concerning migration in other animal groups have been assembled and it seems worth while to compare them with those known for birds. There
Loptson, Claire; Lunt, Dan; Francis, Jane
The climates during the Cretaceous (~144 to 66 Ma) and the early Eocene (~56 to 48 Ma) were much warmer than the present day. Atmospheric CO2 levels for these past climates have a large uncertainty associated with them, but were possibly as high as 2000 to 3000 ppm for the early Eocene (Beerling and Royer, 2011; Lowenstein and Demicco, 2006) and maximum values are thought to range from 800 to 1800 ppm during the Cretaceous (Royer et al., 2012). Current modelling efforts have had great difficulty in replicating the shallow latitudinal temperature gradient indicated by proxy data for these time periods (e.g. Heinemann et al., 2009; Winguth et al., 2010; Shellito et al., 2009). Mechanisms that can result in such a low temperature gradient have not been found (Winguth et al., 2010; Beerling et al., 2011; Sloan and Morrill, 1998), but a contributing factor could be that not all climate feedbacks are included in these models. Vegetation feedbacks have been shown to be especially important (e.g. Otto-Bliesner and Upchurch, 1997; Bonan, 2008) so by including a more accurate representation of vegetation in the climate model, the model-data discrepancies may be reduced. A fully coupled atmosphere-ocean GCM, HadCM3L, coupled to a dynamic global vegetation model (TRIFFID), was used to simulate the climate and the predicted vegetation distributions for and the early Eocene and 12 different time slices representing different ages throughout the Cretaceous at 4x pre-industrial CO2. The only difference in the way these simulations were set up are different boundary conditions that are specific to that time period, e.g. different solar constants and paleogeographies. This allows a direct comparison between the time slices. We present the changes in climate, and therefore vegetation, during the Cretaceous due to changes in these boundary conditions alone, with a focus on Antarctica. Additional Eocene simulations were also carried out with a) fixed globally-uniform vegetation and b
Mehay, S.; Jahren, A.; Schubert, B.; Eberle, J. J.; Summons, R. E.
The Early to Middle Eocene (~56-45 Ma) was a “greenhouse” interval with average global temperatures warmer than any other time in the Cenozoic. This period was characterized by warm climates at high latitude leading to lush forests and the arrival of new mammal groups north of the Arctic Circle (>73°N). Glycerol dialkyl glycerol tetraethers (GDGTs) are membrane-spanning lipids characteristic of certain archaea and bacteria and it has been demonstrated that branched and cyclic GDGTs derived from soil bacteria vary in structure as a function of environmental factors. Proxies based on the relative abundances of methyl branched and cyclopentyl bacterial tetraethers are hypothesized to correlate with mean annual air temperature and soil pH. Here we present the occurrence and distribution of GDGTs in a range of paleosol and sediment samples from Axel Heiberg Island and Ellesmere Island, Nunavut (eastern Canadian Arctic) and Banks Island in the Northwest Territories (western Canadian Arctic). Preliminary results on 11 paleosol samples from the middle Eocene-aged Geodetic Hills Fossil Forest on Axel Heiberg Island indicate a mean annual air temperature of about 9°C. Earlier paleotemperature estimates for Axel Heiberg Island led to values ranging from 9°C to 15°C for the Middle Eocene. Recent temperature prediction for Ellesmere Island (Early Eocene) based upon oxygen isotope ratios of biogenic phosphate from mammal and fish fossils led to ~8°C. In contrast, GDGTs from a marine sedimentary sequence from Lomonosov Ridge in the central Arctic Ocean led to much higher Early Eocene temperature. Thus, the evaluation of the paleotemperature for the Early to Middle Eocene is still a subject of controversy. Ongoing GDGTs analysis of samples from Ellesmere and Banks Islands should give a more comprehensive paleoenvironmental description of the Eocene Arctic. Differences observed between the various paleotemperature estimates will also be discussed. GDGTs distributions are
Frieling, Joost; Huurdeman, Emiel P.; Rem, Charlotte C. M.; Donders, Timme H.; Pross, Jörg; Bohaty, Steven M.; Holdgate, Guy R.; Gallagher, Stephen J.; McGowran, Brian; Bijl, Peter K.
Detailed, stratigraphically well-constrained environmental reconstructions are available for Paleocene and Eocene strata at a range of sites in the southwest Pacific Ocean (New Zealand and East Tasman Plateau; ETP) and Integrated Ocean Discovery Program (IODP) Site U1356 in the south of the Australo-Antarctic Gulf (AAG). These reconstructions have revealed a large discrepancy between temperature proxy data and climate models in this region, suggesting a crucial error in model, proxy data or both. To resolve the origin of this discrepancy, detailed reconstructions are needed from both sides of the Tasmanian Gateway. Paleocene-Eocene sedimentary archives from the west of the Tasmanian Gateway have unfortunately remained scarce (only IODP Site U1356), and no well-dated successions are available for the northern sector of the AAG. Here we present new stratigraphic data for upper Paleocene and lower Eocene strata from the Otway Basin, southeast Australia, on the (north)west side of the Tasmanian Gateway. We analyzed sediments recovered from exploration drilling (Latrobe-1 drill core) and outcrop sampling (Point Margaret) and performed high-resolution carbon isotope geochemistry of bulk organic matter and dinoflagellate cyst (dinocyst) and pollen biostratigraphy on sediments from the regional lithostratigraphic units, including the Pebble Point Formation, Pember Mudstone and Dilwyn Formation. Pollen and dinocyst assemblages are assigned to previously established Australian pollen and dinocyst zonations and tied to available zonations for the SW Pacific. Based on our dinocyst stratigraphy and previously published planktic foraminifer biostratigraphy, the Pebble Point Formation at Point Margaret is dated to the latest Paleocene. The globally synchronous negative carbon isotope excursion that marks the Paleocene-Eocene boundary is identified within the top part of the Pember Mudstone in the Latrobe-1 borehole and at Point Margaret. However, the high abundances of the
Austermann, Jacqueline; Carter, Laura B.; Middleton, Jennifer; Stellmann, Jessica; Pyle, Lacey
Oxygen isotope records reveal that deep-sea temperatures were relatively stable in the early and mid Paleocene before they rose by approx. 4°C to peak in the early Eocene. This Early Eocene Climate Optimum was followed by a 17 Myr cooling trend that led to the onset of Antarctic glaciation at the end of the Eocene. Several studies have examined the potential influence of perturbations to the sinks and sources of atmospheric carbon as mechanisms for the temperature drawdown over the Eocene. Examination of the changing magnitude of carbon sinks has focused on the importance of increased weathering associated with the uplift of the Tibetan plateau (Raymo and Ruddiman, 1992), the continental drift of basaltic provinces through the equatorial humid belt (Kent and Muttoni, 2013), or the emplacement of ophiolites during arc-continent collision in the face of a closing Tethys ocean (Jagoutz et al., 2016). With respect to carbon sources, the shutdown of Tethys subduction and related arc volcanism has been argued to significantly decrease carbon emissions and consequently global temperatures (Hoareau et al., 2015). In this study, we re-assess and quantify proposed atmospheric carbon sinks and sources to obtain an integrated picture of carbon flux changes over the Paleocene and Eocene and to estimate the relative importance of different mechanisms. To constrain carbon sources, we attempt to calculate the outgassing associated with large igneous provinces, mid-ocean ridges and volcanic arcs. We use plate reconstructions to track changes in length and divergence / convergence rates at plate boundaries as well as account for the onset and extinction of volcanic arcs. To constrain carbon sinks, we account for the sequestering of carbon due to silicate weathering and organic carbon burial. We again make use of plate reconstructions to trace highly weatherable arc systems and basaltic extrusions through the tropical humid belt and to assess the interplay between warmer Eocene
Garnit, Hechmi; Bouhlel, Salah; Jarvis, Ian
The Late Paleocene-Early Eocene phosphorites of the Metlaoui Group in Tunisia are a world-class phosphate resource. We review the characteristics of phosphorites deposited in three areas: the Northern Basins; Eastern Basins; and Gafsa-Metlaoui Basin. Comprehensive new bulk rock elemental data are presented, together with complementary mineralogical and mineral chemical results. Carbonate fluorapatite (francolite) constitutes the dominant mineral phase in the deposits. Phosphorite samples are enriched in Cd, Sr, U, rare-earth elements and Y, together with environmentally diagnostic trace elements that provide detrital (Cr, Zr), productivity (Cu, Ni, Zn) and redox (Mo, V) proxies. Suboxic bottom-water conditions predominated, with suboxic to anoxic porewaters accompanying francolite precipitation. Phosphorite deposition occurred under increasingly arid climate conditions, accompanying global Paleocene-Eocene warming. The Northern Basins show the strongest Tethys Ocean influence, with surface seawater rare-earth element signatures consistently developed in the phosphorites. Bed-scale compositional variation indicates relatively unstable environmental conditions and episodes of sediment redeposition, with varying detrital supply and a relatively wet local climate. Glauconitic facies in the Northern Basins and the more isolated evaporite-associated phosphorites in the dryer Eastern Basins display the greatest diagenetic influences. The phosphorite - organic-rich marl - diatom-bearing porcelanite facies association in the Gafsa-Metlaoui Basin represents the classic coastal upwelling trinity. Modified Tethyan waters occurred within the Basin during phosphorite deposition, with decreasing marine productivity from NW to SE evidenced by systematically falling enrichment factors for Cu, Ni, Cd and Zn in the phosphorites. Productivity declined in concert with increasing basin isolation during the deposition of the commercial phosphorite beds in the latest Paleocene to earliest
Zachary, W.; Keating-Bitonti, C.
The Eocene-Oligocene transition is a significant global cooling event with the first growth of continental ice on Antarctica. In the geologic record, the size of fossils can be used to indirectly observe how organisms respond to climate change. For example, organisms tend to be larger in cooler environments as a physiological response to temperature. This major global cooling event should influence organism physiology, resulting in significant size trends observed in the fossil record. Benthic foraminifera are protists and those that grow a carbonate shell are both well-preserved and abundant in marine sediments. Here, we used the foraminiferal fossil record to study the relationship between their size and global cooling. We hypothesize that cooler temperatures across the Eocene-Oligocene boundary promoted shell size increase. To test this hypothesis, we studied benthic foraminifera from 10 deep-sea cores drilled at Ocean Drilling Program Site 744, located in the southern Indian Ocean. We washed sediment samples over a 63-micron sieve and picked foraminifera from a 125-micron sieve. We studied the benthic foraminiferal genus Cibicidoides and its size change across this cooling event. Picked specimens were imaged and we measured the diameter of their shells using "imageJ". Overall, we find that Cibicidoides shows a general trend of increasing size during this transition. In particular, both the median and maximum sizes of Cibicidoides increase from the Eocene into the Oligocene. We also analyzed C. pachyderma and C. mundulus for size trends. Although both species increase in median size across the boundary, only C. pachyderma shows a consistent trend of increasing maximum, median, and minimum shell diameter. After the Eocene-Oligocene boundary, we observe that shell diameter decreases following peak cooling and that foraminiferal sizes remain stable into the early Oligocene. Therefore, the Eocene-Oligocene cooling event appears to have strong influence on shell size.
Muttoni, G.; Dallanave, E.
The magnetostratigraphy and rock-magnetism of the Paleocene-Eocene interval has been studied in the recent years in several Tethyan marine sections of the Belluno Basin of NE Italy (Possagno, Cicogna, South Ardo, Alano). The paleomagnetic results, integrated with calcareous nannofossil biostratigraphy, allowed the recovery of a virtually continuous ~27 Myr-long interval of time spanning from the K/Pg boundary (~65 Ma) at the South Ardo section up to the middle-late Eocene boundary (~38 Ma) at the Alano GSSP candidate, and bracketing some of the most extreme climate conditions of the Cenozoic such the Early Eocene climatic optimum (EECO) and the Paleocene-Eocene thermal maximum (PETM). The rock-magnetic data indicate that the magnetic mineralogy of the sediments generally consists of variable proportions of magnetite-maghemite-hematite, which are iron oxides characterized by different oxidation states and crystal structures. We reconstructed the rock-magnetic variability across the investigated interval, and placed it on a temporal reference frame using a CK95-based age-depth function for comparison with oxygen isotope data from the literature. The rock-magnetic data indicate that relatively warmer climate periods (i.e. the PETM and the early Eocene warming trend leading to EECO) are associated with high contents of detrital hematite relative to magnetite-maghemite, while relatively cooler climates (i.e. the Paleocene) are associated with a relative increase in magnetite-maghemite. We speculate that the increase of detrital hematite observed during warm periods is due to intensified chemical weathering rates of land silicates under warm and humid climates. We therefore show that rock-magnetic properties can be useful proxies to study the efficiency of the silicate weathering negative feedback mechanism to stabilize long-term Earth's surface temperatures.
Bohaty, S. M.; Zachos, J. C.
The late middle-to-late Eocene (~45 to 34 Ma) is a critical period of Cenozoic climatic evolution, characterized by long-term cooling of deep waters and small-scale growth of ice sheets on Antarctica. This time interval, however, is poorly represented in deep-sea sediment archives, and high-resolution paleoceanographic records have not been developed for many of the sections that are available. Several Southern Ocean sites have adequate stratigraphic coverage through the middle-to-late Eocene and provide an opportunity to investigate regional paleoceanographic change through this time interval. In order to supplement the existing database, we have generated high-resolution stable isotope records from fine-fraction carbonates and foraminifera at Site 748, located on the southern Kerguelen Plateau. Integration of middle-to-late Eocene oxygen isotope records from Sites 748 and 738 (Kerguelen Plateau) and Site 689 (Maud Rise) reveals a prominent, short-term warming event at ~41.5 Ma, designated as the Middle Eocene Climatic Optimum (MECO). At Site 748, both surface-dwelling (Acarinina spp.) and benthic (Cibicidoides spp.) exhibit two δ 18O minima, indicating that this event is characterized by two brief intervals of peak warming in both surface and deep waters. Benthic foraminiferal δ 18O data further indicate that middle-bathyal waters warmed ~ 4°C on both the Kerguelen Plateau and Maud Rise during the MECO. Accordingly, this transient event (isotope excursion, as observed during the Paleocene-Eocene Thermal Maximum, suggests that this event was not triggered by methane dissociation. Instead, the MECO may possibly be linked to increased pCO2 associated with the Chron C20-19 plate reorganization. Possible sources of CO2 during this period of global tectonic change are metamorphic decarbonation in the Himalayan orogen and inter-plate volcanism.
Sagoo, Navjit; Valdes, Paul; Flecker, Rachel; Gregoire, Lauren J
Geological data for the Early Eocene (56-47.8 Ma) indicate extensive global warming, with very warm temperatures at both poles. However, despite numerous attempts to simulate this warmth, there are remarkable data-model differences in the prediction of these polar surface temperatures, resulting in the so-called 'equable climate problem'. In this paper, for the first time an ensemble with a perturbed climate-sensitive model parameters approach has been applied to modelling the Early Eocene climate. We performed more than 100 simulations with perturbed physics parameters, and identified two simulations that have an optimal fit with the proxy data. We have simulated the warmth of the Early Eocene at 560 ppmv CO2, which is a much lower CO2 level than many other models. We investigate the changes in atmospheric circulation, cloud properties and ocean circulation that are common to these simulations and how they differ from the remaining simulations in order to understand what mechanisms contribute to the polar warming. The parameter set from one of the optimal Early Eocene simulations also produces a favourable fit for the last glacial maximum boundary climate and outperforms the control parameter set for the present day. Although this does not 'prove' that this model is correct, it is very encouraging that there is a parameter set that creates a climate model able to simulate well very different palaeoclimates and the present-day climate. Interestingly, to achieve the great warmth of the Early Eocene this version of the model does not have a strong future climate change Charney climate sensitivity. It produces a Charney climate sensitivity of 2.7(°)C, whereas the mean value of the 18 models in the IPCC Fourth Assessment Report (AR4) is 3.26(°)C±0.69(°)C. Thus, this value is within the range and below the mean of the models included in the AR4.
Cui, Ying; Schubert, Brian A.
Multiple short-lived global warming events, known as hyperthermals, occurred during the early Eocene (56-52 Ma). Five of these events - the Paleocene-Eocene Thermal Maximum (PETM or ETM1), H1 (or ETM2), H2, I1, and I2 - are marked by a carbon isotope excursion (CIE) within both marine and terrestrial sediments. The magnitude of CIE, which is a function of the amount and isotopic composition of carbon added to the ocean-atmosphere system, varies significantly between marine versus terrestrial substrates. Here we use the increase in carbon isotope fractionation by C3 land plants in response to increased pCO2 to reconcile this difference and reconstruct a range of background pCO2 and peak pCO2 for each CIE, provided two potential carbon sources: methane hydrate destabilization and permafrost-thawing/organic matter oxidation. Although the uncertainty on each pCO2 estimate using this approach is low (e.g., median uncertainty = + 23% / - 18%), this work highlights the potential for significant systematic bias in the pCO2 estimate resulting from sampling resolution, substrate type, diagenesis, and environmental change. Careful consideration of each of these factors is required especially when applying this approach to a single marine-terrestrial CIE pair. Given these limitations, we provide an upper estimate for background early Eocene pCO2 of 463 +248/-131 ppmv (methane hydrate scenario) to 806 +127/-104 ppmv (permafrost-thawing/organic matter oxidation scenario). These results, which represent the first pCO2 proxy estimates directly tied to the Eocene hyperthermals, demonstrate that early Eocene warmth was supported by background pCO2 less than ∼3.5× preindustrial levels and that pCO2 > 1000 ppmv may have occurred only briefly, during hyperthermal events.
Thorup, Kasper; Holland, Richard A.; Tøttrup, Anders P.
orient during migration. Despite the difficulties associated with following free-flying birds over long distances, a number of possibilities currently exist for tracking the long distance, sometimes even globe-spanning, journeys undertaken by migrating birds. Birds fitted with radio transmitters can......For many years, orientation in migratory birds has primarily been studied in the laboratory. Although a laboratory-based setting enables greater control over environmental cues, the laboratory-based findings must be confirmed in the wild in free-flying birds to be able to fully understand how birds...... system that enables experienced birds to navigate and guide inexperienced, young birds to their species-specific winter grounds...
Kovalev, Alexander; Filippov, Alexander E; Gorb, Stanislav N
The bird feather vane can be separated into two parts by pulling the barbs apart. The original state can be re-established easily by lightly stroking through the feather. Hooklets responsible for holding vane barbs together are not damaged by multiple zipping and unzipping cycles. Because numerous microhooks keep the integrity of the feather, their properties are of great interest for understanding mechanics of the entire feather structure. This study was undertaken to estimate the separation force of single hooklets and their arrays using force measurement of an unzipping feather vane. The hooklets usually separate in some number synchronously (20 on average) with the highest observed separation force of 1.74 mN (average force 0.27 mN), whereas the single hooklet separation force was 14 μN. A simple numerical model was suggested for a better understanding of zipping and unzipping behaviour in feathers. The model demonstrates features similar to those observed in experiments.
The conservation and management of wading birds has received considerable attention over the past twenty years, through research, population monitoring, habitat protection, and through activities of specialist groups devoted to all three groups, the herons, ibises and allies, and flamingos. While populations are best known in North America, greatest advances in knowledge may have come in Australasia. The status of most species and many populations is now sufficiently known to allow assessment of risk. Conservation and management techniques allow creation of global and regional action plans for conservation of many species. Global action plans are being developed, but few regional plans have been undertaken. Management of nesting sites is now particularly well appreciated. Although known in broad stroke, much remains to be learned about managing feeding habitat. Problems related to disturbance, conflict with humans, habitat loss, contaminants and other environmental stresses remain for some species and many populations. New challenges lie in creating conservation action that account for genetic stocks.
National Oceanic and Atmospheric Administration, Department of Commerce — This data set contains sensitive biological resource data for shorebirds, waterfowl, raptors, diving birds, and seabirds in the Bristol Bay Subarea. The Subarea...
Eugene N. Anderson
Full Text Available The Mongol Empire, the largest contiguous empire the world has ever known, had, among other things, a goodly number of falconers, poultry raisers, birdcatchers, cooks, and other experts on various aspects of birding. We have records of this, largely in the Yinshan Zhengyao, the court nutrition manual of the Mongol empire in China (the Yuan Dynasty. It discusses in some detail 22 bird taxa, from swans to chickens. The Huihui Yaofang, a medical encyclopedia, lists ten taxa used medicinally. Marco Polo also made notes on Mongol bird use. There are a few other records. This allows us to draw conclusions about Mongol ornithology, which apparently was sophisticated and detailed.
Introduction to Avian Medicine: Companion Birds and Wild Birds. T W deMaar. Abstract. No abstract. The Kenya Veterinarian Vol. 21 2001: pp. 20-22. Full Text: EMAIL FULL TEXT EMAIL FULL TEXT · DOWNLOAD FULL TEXT DOWNLOAD FULL TEXT · http://dx.doi.org/10.4314/kenvet.v21i1.39504 · AJOL African Journals ...
Carmichael, Matthew J.; Lunt, Daniel J.; Huber, Matthew; Heinemann, Malte; Kiehl, Jeffrey; LeGrande, Allegra; Loptson, Claire A.; Roberts, Chris D.; Sagoo, Navjit; Shields, Christine
A range of proxy observations have recently provided constraints on how Earth's hydrological cycle responded to early Eocene climatic changes. However, comparisons of proxy data to general circulation model (GCM) simulated hydrology are limited and inter-model variability remains poorly characterised. In this work, we undertake an intercomparison of GCM-derived precipitation and P - E distributions within the extended EoMIP ensemble (Eocene Modelling Intercomparison Project; Lunt et al., 2012), which includes previously published early Eocene simulations performed using five GCMs differing in boundary conditions, model structure, and precipitation-relevant parameterisation schemes. We show that an intensified hydrological cycle, manifested in enhanced global precipitation and evaporation rates, is simulated for all Eocene simulations relative to the preindustrial conditions. This is primarily due to elevated atmospheric paleo-CO2, resulting in elevated temperatures, although the effects of differences in paleogeography and ice sheets are also important in some models. For a given CO2 level, globally averaged precipitation rates vary widely between models, largely arising from different simulated surface air temperatures. Models with a similar global sensitivity of precipitation rate to temperature (dP=dT ) display different regional precipitation responses for a given temperature change. Regions that are particularly sensitive to model choice include the South Pacific, tropical Africa, and the Peri-Tethys, which may represent targets for future proxy acquisition. A comparison of early and middle Eocene leaf-fossil-derived precipitation estimates with the GCM output illustrates that GCMs generally underestimate precipitation rates at high latitudes, although a possible seasonal bias of the proxies cannot be excluded. Models which warm these regions, either via elevated CO2 or by varying poorly constrained model parameter values, are most successful in simulating a
Meng, F. W.; Wang, X.-Q.; Ni, P.; Kletetschka, Günther; Li, Y-P.; Yang, Ch.-H.
Roč. 30, č. 3 (2015), s. 321-330 ISSN 0891-2556 Institutional support: RVO:67985831 Keywords : ancient microorganism * primary fluid inclusion * halite * Eocene Subject RIV: DB - Geology ; Mineralogy Impact factor: 0.440, year: 2015
... 21 Food and Drugs 8 2010-04-01 2010-04-01 false Psittacine birds. 1240.65 Section 1240.65 Food and... DISEASES Specific Administrative Decisions Regarding Interstate Shipments § 1240.65 Psittacine birds. (a) The term psittacine birds shall include all birds commonly known as parrots, Amazons, Mexican double...
Tajovský, Karel; Mock, A.; Krumpál, M.
Roč. 37, - (2001), s. 321-323 ISSN 1164-5563 Institutional research plan: CEZ:AV0Z6066911 Keywords : birds nests * microsites * millipedes Subject RIV: EH - Ecology, Behaviour Impact factor: 0.317, year: 2001
Ligabue-Braun, Rodrigo; Carlini, Célia Regina
Until very recently, toxicity was not considered a trait observed in birds, but works published in the last two decades started to shed light on this subject. Poisonous birds are rare (or little studied), and comprise Pitohui and Ifrita birds from Papua New Guinea, the European quail, the Spoor-winged goose, the Hoopees, the North American Ruffed grouse, the Bronzewings, and the Red warbler. A hundred more species are considered unpalatable or malodorous to humans and other animals. The present review intends to present the current understanding of bird toxicity, possibly pointing to an ignored research field. Whenever possible, biochemical characteristics of these poisons and their effects on humans and other animals are discussed, along with historical aspects of poison discovery and evolutionary hypothesis regarding their function. Copyright © 2015 Elsevier Ltd. All rights reserved.
Among 2589 captive wild birds, examined between 1974 and 1983, we found leukosis in 26 birds belonging to 13 different species and five orders. We diagnosed lymphoid leukosis in 11 birds (two Melopsittacus undulatus, two Psittacus erithacus one Platycerus eximius, one Columba livia, one Streptopelia decaocto, one Polyplectron bicalcaratum, one Pavo cristatus, one Aptenodytes patachonia and one finch, species unknown), myeloid leukosis in 14 (nine Melopsittacus undulatus, two Agapomis personata fischeri, two Urgeainthus bengalus and one Neophemia pulchella) and stem cell leukosis in one bird (Serinus canaria). Among the cases with lymphoid leukosis we distinguished between lymphoblastic (four cases) and prolymphocytic forms (seven). Myeloid leukosis was subdivided into poorly differentiated (12 cases) and well differentiated myeloblastosis (two).
... resources including migratory birds and their habitats. Large-scale efforts to influence bird migration and... timing and speed of bird migrations. It is possible that re-distribution of birds at smaller scales could...-0040; 91200-1231-9BPP-L2] RIN 1018-AX06 Migratory Bird Hunting; Final Frameworks for Early-Season...
... Service 50 CFR Part 20 Migratory Bird Hunting; Migratory Bird Hunting Regulations on Certain Federal...-L2] RIN 1018-AX34 Migratory Bird Hunting; Migratory Bird Hunting Regulations on Certain Federal..., Interior. ACTION: Final rule. SUMMARY: This rule prescribes special early-season migratory bird hunting...
... Service 50 CFR Part 20 Migratory Bird Hunting; Proposed Migratory Bird Hunting Regulations on Certain...-FXMB1231099BPP0L2] RIN 1018-AX97 Migratory Bird Hunting; Proposed Migratory Bird Hunting Regulations on Certain...) proposes special migratory bird hunting regulations for certain Tribes on Federal Indian reservations, off...
... Service 50 CFR Part 20 Migratory Bird Hunting; Supplemental Proposals for Migratory Game Bird Hunting...] RIN 1018-AX97 Migratory Bird Hunting; Supplemental Proposals for Migratory Game Bird Hunting... in an earlier document to establish annual hunting regulations for certain migratory game birds for...
... Service 50 CFR Part 20 Migratory Bird Hunting; Proposed Migratory Bird Hunting Regulations on Certain...-FXMB1231099BPP0] RIN 1018-AY87 Migratory Bird Hunting; Proposed Migratory Bird Hunting Regulations on Certain...) proposes special migratory bird hunting regulations for certain Tribes on Federal Indian reservations, off...
Gordon F. Bennett
Full Text Available Linear measurements and derived indices of trypanosomes from species of Fennoscandian birds were compared to those reported form Trypanosoma avium, T. everetti, T. ontarioensis and T. paddae. The trypanosomes encountered in the Fennoscandian birds were identified as T. avium from Tengmalm's owl Aegolius funereus and the pied flycatcher Ficedula hypoleuca, T. everetti from the great tit Parus major and collared flycatcher F. albicollis and T. ontarioensis from the collared flycatcher; T. paddae was not seen.
Calcareous plankton (calcareous nannofossils, planktic foraminifera) is currently studied from DSDP Site 401 during the latest Paleocene to Middle Eocene greenhouse episode. This site is situated on the edge of a tilted fault block underlying the southern edge of the Meriadzek terrace on the North Biscay margin and comprises a 100-m-thick, nearly complete sedimentary record of Paleocene to middle Eocene sediments covering the Paleocene-Eocene Thermal Maximum as well as the Early Eocene Climatic Optimum. Detailed age control is achieved by calcareous nannofossil and planktic foraminiferal biostratigraphy suggesting that the study interval covers the NP9 to NP16 and P5 to E13 biozones, respectively, with a pronounced condensed interval across the Early to Middle Eocene boundary (NP13/NP14). Besides DSDP Site 550 this site represents one of the most northern locations which consists of Paleogene carbonates and will provide us a more complete picture by considering also long-term records from the northern hemisphere. Here we present first assemblage data from the two plankton groups which are accompanied by planktic and benthic foraminiferal oxygen and carbon isotope data in order to unravel the impact of the long-term climate change on the planktic ecosystem during the Eocene greenhouse.
Cooper, Lisa Noelle; Seiffert, Erik R; Clementz, Mark; Madar, Sandra I; Bajpai, Sunil; Hussain, S Taseer; Thewissen, J G M
Anthracobunidae is an Eocene family of large mammals from south Asia that is commonly considered to be part of the radiation that gave rise to elephants (proboscideans) and sea cows (sirenians). We describe a new collection of anthracobunid fossils from Middle Eocene rocks of Indo-Pakistan that more than doubles the number of known anthracobunid fossils and challenges their putative relationships, instead implying that they are stem perissodactyls. Cranial, dental, and postcranial elements allow a revision of species and the recognition of a new anthracobunid genus. Analyses of stable isotopes and long bone geometry together suggest that most anthracobunids fed on land, but spent a considerable amount of time near water. This new evidence expands our understanding of stem perissodactyl diversity and sheds new light on perissodactyl origins.
Lisa Noelle Cooper
Full Text Available Anthracobunidae is an Eocene family of large mammals from south Asia that is commonly considered to be part of the radiation that gave rise to elephants (proboscideans and sea cows (sirenians. We describe a new collection of anthracobunid fossils from Middle Eocene rocks of Indo-Pakistan that more than doubles the number of known anthracobunid fossils and challenges their putative relationships, instead implying that they are stem perissodactyls. Cranial, dental, and postcranial elements allow a revision of species and the recognition of a new anthracobunid genus. Analyses of stable isotopes and long bone geometry together suggest that most anthracobunids fed on land, but spent a considerable amount of time near water. This new evidence expands our understanding of stem perissodactyl diversity and sheds new light on perissodactyl origins.
Alerstam, Thomas; Bäckman, Johan; Gudmundsson, Gudmundur A; Hedenström, Anders; Henningsson, Sara S; Karlsson, Håkan; Rosén, Mikael; Strandberg, Roine
Studies of bird migration in the Beringia region of Alaska and eastern Siberia are of special interest for revealing the importance of bird migration between Eurasia and North America, for evaluating orientation principles used by the birds at polar latitudes and for understanding the evolutionary implications of intercontinental migratory connectivity among birds as well as their parasites. We used tracking radar placed onboard the ice-breaker Oden to register bird migratory flights from 30 ...
Bougeois, Laurie; de Rafélis, Marc; Tindall, Julia; Proust, Jean-Noël; Reichart, Gert-Jan; de Nooijer, Lennart; Guo, ZhaoJie; Ormukov, Cholponbek; Dupont-Nivet, Guillaume
Climate models suggest that the onset of Asian monsoons and aridification have been governed by Tibetan plateau uplift, global climate changes and the retreat to the west of the vast epicontinental Proto-Paratethys sea during the warm Eocene greenhouse period (55-34 million years ago). However, the role of the Proto-Paratethys sea on climate remains to be quantified by accurate and precise reconstructions. By applying a novel intra-annual geochemical multi-proxy methodology on Eocene oyster shells of the Proto-Paratethys sea and comparing results to climate simulations and sedimentology analyses, we show that the Central Asian region was generally arid with a high seasonal contrast characterized by hot and arid summers and wetter winters. Hotter and more arid summers despite the presence of the Proto-Paratethys may be explained by warmer Eocene global conditions with a strong anticyclonic Hadley cell descending at Central Asian latitudes and a stronger Foehn effect from the emerging Tibetan Plateau to the south. This implies that the shallow sea did not have a strong dampening thermal effect on the monsoonal circulation in contrast to previous circulation models results but in agreement with recent evidence for Eocene summer monsoons. Enhanced winter precipitations, relative to modern, is linked to a westerly moisture source coming from the Proto-Paratethys sea at that time. Additional bulk sediment stable isotope data from marine limestones and pedogenic carbonates suggest a gradual decrease in this westerly moisture source, which is in line with the retreat of the Proto-Paratethys followed by the Oligo-Miocene orogeny of the Central Asian ranges (Tian Shan and Pamir) shielding the westerlies.
Huck, Claire E.; van de Flierdt, Tina; Bohaty, Steven M.; Hammond, Samantha J.
We assess early-to-middle Eocene seawater neodymium (Nd) isotope records from seven Southern Ocean deep-sea drill sites to evaluate the role of Southern Ocean circulation in long-term Cenozoic climate change. Our study sites are strategically located on either side of the Tasman Gateway and are positioned at a range of shallow (fish teeth at intermediate/deep Indian Ocean pelagic sites (Ocean Drilling Program (ODP) Sites 738 and 757 and Deep Sea Drilling Project (DSDP) Site 264), indicate a dominant Southern Ocean-sourced contribution to regional deep waters (ɛNd(t) = -9.3 ± 1.5). IODP Site U1356 off the coast of Adélie Land, a locus of modern-day Antarctic Bottom Water production, is identified as a site of persistent deep water formation from the early Eocene to the Oligocene. East of the Tasman Gateway an additional local source of intermediate/deep water formation is inferred at ODP Site 277 in the SW Pacific Ocean (ɛNd(t) = -8.7 ± 1.5). Antarctic-proximal shelf sites (ODP Site 1171 and Site U1356) reveal a pronounced erosional event between 49 and 48 Ma, manifested by 2 ɛNd unit negative excursions in seawater chemistry toward the composition of bulk sediments at these sites. This erosional event coincides with the termination of peak global warmth following the Early Eocene Climatic Optimum and is associated with documented cooling across the study region and increased export of Antarctic deep waters, highlighting the complexity and importance of Southern Ocean circulation in the greenhouse climate of the Eocene.
M.H. Hermiyanto Zajuli
Full Text Available DOI: 10.17014/ijog.v6i2.119Indonesia contains many Paleogene and Neogene basins which some of them have been proven to be a very prolific producer of oil and gas. A study on the result of Rock-Eval pyrolysis and biomarker undertaken on the Eocene Mandai Group was able to assess hydrocarbon potential of the Paleogene fine sediments in the frontier basin, especially West Kalimantan area. East Ketungau Basin is located in the western Kalimantan, bounded with Melawi Basin by the Semitau High in the south and West Ketungau Basin in the west. The Mandai Group was deposited in the East Ketungau Basin during Eocene, consisting of sandstone and mudstone facies. Mudstone facies comprises shale, claystone, and coal. Seven samples of Eocene fine sediments collected from East Ketungau Basin were analyzed by Rock-Eval pyrolisis and three samples for biomarker to evaluate their hydrocarbon potential. The Rock-Eval pyrolisis result of Mandai Group shows that TOC value of this facies ranges from 0.34 % to 5.16 %, Potential Yield (PY between 0.06 and 4.78 mg HC/g rock, and Hydrogen Index (HI from 12 to 89. Based on that result, the fine sediments of Mandai Group are included into a gas prone source rock potential with poor to fair categories. Moreover Tmax values vary from 426o C to 451o C. The Eocene fine sediments of Mandai Group fall under kerogen type III. Based on Tmax and biomarker analyses, the maturity of the sediments is situated within immature to mature level. The fine sediments of Mandai Group were deposited in a terrestrial to marine environment under anoxic to sub-oxic condition.
Stickley, Catherine E; St John, Kristen; Koç, Nalân; Jordan, Richard W; Passchier, Sandra; Pearce, Richard B; Kearns, Lance E
Oceanic sediments from long cores drilled on the Lomonosov ridge, in the central Arctic, contain ice-rafted debris (IRD) back to the middle Eocene epoch, prompting recent suggestions that ice appeared in the Arctic about 46 million years (Myr) ago. However, because IRD can be transported by icebergs (derived from land-based ice) and also by sea ice, IRD records are restricted to providing a history of general ice-rafting only. It is critical to differentiate sea ice from glacial (land-based) ice as climate feedback mechanisms vary and global impacts differ between these systems: sea ice directly affects ocean-atmosphere exchanges, whereas land-based ice affects sea level and consequently ocean acidity. An earlier report assumed that sea ice was prevalent in the middle Eocene Arctic on the basis of IRD, and although somewhat preliminary supportive evidence exists, these data are neither comprehensive nor quantified. Here we show the presence of middle Eocene Arctic sea ice from an extraordinary abundance of a group of sea-ice-dependent fossil diatoms (Synedropsis spp.). Analysis of quartz grain textural characteristics further supports sea ice as the dominant transporter of IRD at this time. Together with new information on cosmopolitan diatoms and existing IRD records, our data strongly suggest a two-phase establishment of sea ice: initial episodic formation in marginal shelf areas approximately 47.5 Myr ago, followed approximately 0.5 Myr later by the onset of seasonally paced sea-ice formation in offshore areas of the central Arctic. Our data establish a 2-Myr record of sea ice, documenting the transition from a warm, ice-free environment to one dominated by winter sea ice at the start of the middle Eocene climatic cooling phase.
Studying past intervals of abrupt global warming and massive carbon release can improve our knowledge in ways relevant to understanding future climate change. Possible paleo-analogues for future climate change are the early Paleogene hyperthermal events, such as the Paleocene-Eocene Thermal Maximum (PETM; ~56 Ma), during which large amounts of carbon were released. More recently, another distinct period of global warming was discovered, similar in nature to the PETM, occurring approximately t...
Rubincam, David Parry
Drag from hydrogen in the interstellar cloud which formed Gould's Belt may have sent small meteoroids with embedded helium to the Earth, perhaps explaining part or all of the (sup 3) He spike seen in the sedimentary record at the Eocene-Oligocene transition. Assuming the Solar System passed through part of the cloud, meteoroids in the asteroid belt up to centimeter size may have been dragged to the resonances, where their orbital eccentricities were pumped up into Earth-crossing orbits.
William C. Clyde
Full Text Available Analysis of lithofacies, paleoflow directions, and sandstone petrography of upper Paleocene-lower Eocene paralic and continental sediments exposed along the transpressional suture zone of the western margin of the Indian plate indicate that the process of deformation and uplift of the carbonate shelf in this area had started by late Paleocene time. This tectonic uplift and deformation is documented by: (1 an overall shallowing upward synorogenic sequence of sediments, (2 proximal conglomerate facies (consisting of lower Paleocene and Mesozoic clasts dominating in the western part of the study area and distal facies of sandstone and shale dominating in the eastern part of the study area, (3 the existence of an unconformity of late Paleocene-early Eocene age in the Quetta and Kalat regions, (4 paleocurrent directions in deltaic and fluvial deposits indicating southeastward flowing sediment dispersal paths during late Paleocene-early Eocene time, which is opposite to that found in the late Cretaceous, suggesting a reversal in the depositional slope of the Cretaceous shelf, and (5 petrographic study of sandstones indicating a collision suture/fold thrust belt provenance. This episode of uplift and deformation could be the result of India-Arabian transpression with associated ophiolite obduction or, more likely, to represent the local response to initial India-Asia contact. The unroofing pattern and uplift geometry of the western Indian shelf suggests that this tectonism first started in the southern part of the study area (Kalat-Khuzdar area during the late Paleocene-early Eocene and proceeded northward in a time-transgressive fashion.
Full Text Available Pleuraspidotheriids are a group of primitive ungulate mammals that, until recently, were thought to be restricted to the late Paleocene of Western Europe. It has been hypothesized that this family actually survived in Central Anatolia until at least the middle Eocene. However, these anachronistically young Anatolian “survivors”, including the genus Hilalia, were previously documented mainly by dental remains. Here, we describe the previously unknown astragalus of Hilalia saribeya, which confirms the pleuraspidotheriid affinities of the genus, and supports phylogenetic reconstructions that place Hilalia as the sister group of Pleuraspidotherium. The morphology of the astragalus suggests sub-cursorial plantigrade locomotion for H. saribeya, although its tarsal morphology remains generalized enough that scansorial capabilities cannot be ruled out. The evolution of Hilalia is addressed in the context of the apparent geographic isolation of Central Anatolia during the Eocene. The endemic character of the mammalian fauna of Central Anatolia during the middle Eocene emphasizes how the complex paleogeography of the northern margin of Neotethys impacted local biotas in a region situated at the crossroads of very distinctive biogeographic zones.
Full Text Available Cambay amber originates from the warmest period of the Eocene, which is also well known for the appearance of early angiosperm-dominated megathermal forests. The humid climate of these forests may have triggered the evolution of epiphytic lineages of bryophytes; however, early Eocene fossils of bryophytes are rare. Here, we present evidence for lejeuneoid liverworts and pleurocarpous mosses in Cambay amber. The preserved morphology of the moss fossil is inconclusive for a detailed taxonomic treatment. The liverwort fossil is, however, distinctive; its zig-zagged stems, suberect complicate-bilobed leaves, large leaf lobules, and small, deeply bifid underleaves suggest a member of Lejeuneaceae subtribe Lejeuneinae (Harpalejeunea, Lejeunea, Microlejeunea. We tested alternative classification possibilities by conducting divergence time estimates based on DNA sequence variation of Lejeuneinae using the age of the fossil for corresponding age constraints. Consideration of the fossil as a stem group member of Microlejeunea or Lejeunea resulted in an Eocene to Late Cretaceous age of the Lejeuneinae crown group. This reconstruction is in good accordance with published divergence time estimates generated without the newly presented fossil evidence. Balancing available evidence, we describe the liverwort fossil as the extinct species Microlejeunea nyiahae, representing the oldest crown group fossil of Lejeuneaceae.
Minwer-Barakat, Raef; Marigó, Judit; Becker, Damien; Costeur, Loïc
Primates reached a great abundance and diversity during the Eocene, favored by warm temperatures and by the development of dense forests throughout the Northern Hemisphere. Here we describe new primate material from La Verrerie de Roches, a Middle Eocene karstic infill situated in the Jura Region (Switzerland), consisting of more than 80 dental remains. The primate assemblage from La Verrerie de Roches includes five different taxa. The best represented primate is Necrolemur aff. anadoni, similar in size and overall morphology to Necrolemur anadoni but resembling in some features the younger species Necrolemur antiquus. Microchoerines are also represented by two species of Pseudoloris, P. pyrenaicus and Pseudoloris parvulus, constituting the unique joint record of these two species known up to now. Remains of Adapiformes are limited to one isolated tooth of a large anchomomyin and another tooth belonging to the small adapine Microadapis cf. sciureus. The studied primate association allows assigning La Verrerie de Roches to the Robiacian Land Mammal Age. More specifically, this site can be confidently situated between the MP15 and MP16 reference levels, although the primate assemblage probably indicates some degree of temporal mixing. This is the first record of P. pyrenaicus and a form closely related to N. anadoni out of the Iberian Peninsula. The identification of these microchoerines in Switzerland gives further support to the connection of NE Spain and Central Europe during the Middle Eocene. Copyright © 2017 Elsevier Ltd. All rights reserved.
Graham, A.; Cozadd, D.; Areces-Mallea, A.; Frederiksen, N.O.
An assemblage of 46 fossil pollen and spore types is described from a core drilled through the middle Eocene Saramaguacan Formation, Camaguey Province, eastern Cuba. Many of the specimens represent unidentified or extinct taxa but several can be identified to family (Palmae, Bombacaceae, Gramineae, Moraceae, Myrtaceae) and some to genus (Pteris, Crudia, Lymingtonia?). The paleo-climate was warm-temperate to subtropical which is consistent with other floras in the region of comparable age and with the global paleotemperature curve. Older plate tectonic models show a variety of locations for proto-Cuba during Late Cretaceous and later times, including along the norther coast of South America. More recent models depict western and central Cuba as two separate parts until the Eocene, and eastern Cuba (joined to northern Hispaniola) docking to central Cuba also in the Eocene. All fragments are part of the North American Plate and none were directly connected with northern South America in late Mesozoic or Cenozoic time. The Saramaguacan flora supports this model because the assemblage is distinctly North American in affinities, with only one type (Retimonocolpites type 1) found elsewhere only in South America.
Rust, Jes; Singh, Hukam; Rana, Rajendra S.; McCann, Tom; Singh, Lacham; Anderson, Ken; Sarkar, Nivedita; Nascimbene, Paul C.; Stebner, Frauke; Thomas, Jennifer C.; Solórzano Kraemer, Monica; Williams, Christopher J.; Engel, Michael S.; Sahni, Ashok; Grimaldi, David
For nearly 100 million years, the India subcontinent drifted from Gondwana until its collision with Asia some 50 Ma, during which time the landmass presumably evolved a highly endemic biota. Recent excavations of rich outcrops of 50–52-million-year-old amber with diverse inclusions from the Cambay Shale of Gujarat, western India address this issue. Cambay amber occurs in lignitic and muddy sediments concentrated by near-shore chenier systems; its chemistry and the anatomy of associated fossil wood indicates a definitive source of Dipterocarpaceae. The amber is very partially polymerized and readily dissolves in organic solvents, thus allowing extraction of whole insects whose cuticle retains microscopic fidelity. Fourteen orders and more than 55 families and 100 species of arthropod inclusions have been discovered thus far, which have affinities to taxa from the Eocene of northern Europe, to the Recent of Australasia, and the Miocene to Recent of tropical America. Thus, India just prior to or immediately following contact shows little biological insularity. A significant diversity of eusocial insects are fossilized, including corbiculate bees, rhinotermitid termites, and modern subfamilies of ants (Formicidae), groups that apparently radiated during the contemporaneous Early Eocene Climatic Optimum or just prior to it during the Paleocene-Eocene Thermal Maximum. Cambay amber preserves a uniquely diverse and early biota of a modern-type of broad-leaf tropical forest, revealing 50 Ma of stasis and change in biological communities of the dipterocarp primary forests that dominate southeastern Asia today. PMID:20974929
Kiehl, Jeffrey T; Shields, Christine A
The Palaeocene-Eocene Thermal Maximum (PETM) was a significant global warming event in the Earth's history (approx. 55 Ma). The cause for this warming event has been linked to increases in greenhouse gases, specifically carbon dioxide and methane. This rapid warming took place in the presence of the existing Early Eocene warm climate. Given that projected business-as-usual levels of atmospheric carbon dioxide reach concentrations of 800-1100 ppmv by 2100, it is of interest to study past climates where atmospheric carbon dioxide was higher than present. This is especially the case given the difficulty of climate models in simulating past warm climates. This study explores the sensitivity of the simulated pre-PETM and PETM periods to change in cloud condensation nuclei (CCN) and microphysical properties of liquid water clouds. Assuming lower levels of CCN for both of these periods leads to significant warming, especially at high latitudes. The study indicates that past differences in cloud properties may be an important factor in accurately simulating past warm climates. Importantly, additional shortwave warming from such a mechanism would imply lower required atmospheric CO2 concentrations for simulated surface temperatures to be in reasonable agreement with proxy data for the Eocene.
Weiss, A.; Martindale, R. C.; Kosir, A.; Oefinger, J.
The Paleocene-Eocene Thermal Maximum (PETM) event ( 56.3 Ma) was a period of massive carbon release into the Earth system, resulting in significant shifts in ocean chemistry. It has been proposed that ocean acidification - a decrease in the pH and carbonate saturation state of the water as a result of dissolved carbon dioxide in sea water - occurred in both the shallow and deep marine realms. Ocean acidification would have had a devastating impact on the benthic ecosystem, and has been proposed as the cause of decreased carbonate deposition in marine sections and coral reef collapse during the late Paleocene. To date, however, the only physical evidence of Paleocene-Eocene ocean acidification has been shown for offshore sites (i.e., a shallow carbonate compensation depth), but isotope analysis (i.e. B, I/Ca) suggests that acidification occurred in the shallow shelves as well. Several sites in the Kras region of Slovenia, has been found to contain apparent erosion surfaces coeval with the Paleocene-Eocene Boundary. We have investigated these potentially acidified horizons using petrography, stable carbon isotopes, cathodoluminescence, and elemental mapping. These datasets will inform whether the horizons formed by seafloor dissolution in an acidified ocean, or are due to subaerial exposure, or burial diagenesis (i.e. stylotization). Physical erosion and diagenesis can easily be ruled out based on field relationships and petrography, but the other potential causes must be analyzed more critically.
Zanazzi, Alessandro; Kohn, Matthew J; MacFadden, Bruce J; Terry, Dennis O
The Eocene-Oligocene transition towards a cool climate (approximately 33.5 million years ago) was one of the most pronounced climate events during the Cenozoic era. The marine record of this transition has been extensively studied. However, significantly less research has focused on continental climate change at the time, yielding partly inconsistent results on the magnitude and timing of the changes. Here we use a combination of in vivo stable isotope compositions of fossil tooth enamel with diagenetic stable isotope compositions of fossil bone to derive a high-resolution (about 40,000 years) continental temperature record for the Eocene-Oligocene transition. We find a large drop in mean annual temperature of 8.2 +/- 3.1 degrees C over about 400,000 years, the possibility of a small increase in temperature seasonality, and no resolvable change in aridity across the transition. The large change in mean annual temperature, exceeding changes in sea surface temperatures at comparable latitudes and possibly delayed in time with respect to marine changes by up to 400,000 years, explains the faunal turnover for gastropods, amphibians and reptiles, whereas most mammals in the region were unaffected. Our results are in agreement with modelling studies that attribute the climate cooling at the Eocene-Oligocene transition to a significant drop in atmospheric carbon dioxide concentrations.
Full Text Available Background We revisit the palaeopalynological record of Loranthaceae, using pollen ornamentation to discriminate lineages and to test molecular dating estimates for the diversification of major lineages. Methods Fossil Loranthaceae pollen from the Eocene and Oligocene are analysed and documented using scanning-electron microscopy. These fossils were associated with molecular-defined clades and used as minimum age constraints for Bayesian node dating using different topological scenarios. Results The fossil Loranthaceae pollen document the presence of at least one extant root-parasitic lineage (Nuytsieae and two currently aerial parasitic lineages (Psittacanthinae and Loranthinae by the end of the Eocene in the Northern Hemisphere. Phases of increased lineage diversification (late Eocene, middle Miocene coincide with global warm phases. Discussion With the generation of molecular data becoming easier and less expensive every day, neontological research should re-focus on conserved morphologies that can be traced through the fossil record. The pollen, representing the male gametophytic generation of plants and often a taxonomic indicator, can be such a tracer. Analogously, palaeontological research should put more effort into diagnosing Cenozoic fossils with the aim of including them into modern systematic frameworks.
Zeebe, R. E.; Westerhold, T.; Littler, K.; Zachos, J. C.
Multi-million-year proxy records (d13C, d18O, %CaCO3, Fe, etc.) show prominent variations on orbital time scale during the Paleocene and Eocene. The cycles have been identified at various sites across the globe and preferentially concentrate spectral power at eccentricity and precessional frequencies. It is almost certain that these cycles are an expression of changes in global climate and carbon cycling paced by orbital variations. However, little is currently known about (1) the driving mechanism linking orbital forcing to changes in climate and carbon cycling and (2) the amplitude of atmospheric CO2 variations associated with these cycles. We have used simple and complex carbon cycle models to explore the basic effect of different orbital forcing schemes and noise on the carbon cycle by forcing different carbon cycle parameters. For direct insolation forcing (opposed to eccentricity - tilt - precession), one major challenge is understanding how the system transfers spectral power from high to low frequencies. We will discuss feasible solutions to this problem, including insolation transformations analogous to electronic AC-DC conversion (DC'ing). Our results show that high-latitude mechanisms are unlikely drivers of orbitally paced changes in the Paleocene-Eocene Earth system. Based on a synthesis of modeling and proxy data analysis, we present the first estimates of orbital-scale variations in atmospheric CO2 during the Paleocene and Eocene.
Lauretano, Vittoria; Lourens, Lucas J.; Zachos, James C.; Sluijs, Appy
During the Early Eocene, the Earth experienced a prolonged warming trend (55-50 Ma), punctuated by a series of short-lived global warming events known as "hyperthermals", of which the Paleocene-Eocene Thermal Maximum, (PETM) is the most extreme, followed by the ETM2 (Elmo) and the ETM3 ("X-event"). Along with these events, a number of less pronounced negative carbon isotope excursions, termed A to L by Cramer et al (2003), occurred throughout the Paleogene up to the Early Eocene Climatic Optimum (EECO). The drastic increase in temperature during the hyperthermals is associated with the release of large amounts of isotopically light carbon into the ocean-atmosphere system. Several hypotheses have been proposed to determine the sources and mechanisms of these carbon inputs, in particular the dissociation of marine gas hydrates. However, what still remains unclear is whether or not these events share a common source. To address this question we compare the quantitative relationship between high-resolution carbon and oxygen stable isotope records from benthic foraminiferal tests. A similar correspondence between changes in the carbon cycle and global temperature among the events, regardless of their magnitudes, would imply a similarity in their origins. Following Stap et al. 2010, we present the results of the comparison between our data and previously published data, using deep-sea material to extend the record to the I1-I2 events.
Gumus, Abdurrahman; Lee, Seoho; Ahsan, Syed S; Karlsson, Kolbeinn; Gabrielson, Richard; Guglielmo, Christopher G; Winkler, David W; Erickson, David
The metabolism of birds is finely tuned to their activities and environments, and thus research on avian systems can play an important role in understanding organismal responses to environmental changes. At present, however, the physiological monitoring of bird metabolism is limited by the inability to take real-time measurements of key metabolites during flight. In this study, we present an implantable biosensor system that can be used for continuous monitoring of uric acid levels of birds during various activities including flight. The system consists of a needle-type enzymatic biosensor for the amperometric detection of uric acid in interstitial fluids. A lightweight two-electrode potentiostat system drives the biosensor, reads the corresponding output current and wirelessly transfers the data or records to flash memory. We show how the device can be used to monitor, in real time, the effects of short-term flight and rest cycles on the uric acid levels of pigeons. In addition, we demonstrate that our device has the ability to measure uric acid level increase in homing pigeons while they fly freely. Successful application of the sensor in migratory birds could open up a new way of studying birds in flight which would lead to a better understanding of the ecology and biology of avian movements.
National Oceanic and Atmospheric Administration, Department of Commerce — This data set contains sensitive biological resource data for wading birds, shorebirds, waterfowl, raptors, diving birds, seabirds, passerine birds, and gulls and...
.... This annotated bibliography of bird hazards to aircraft, termed ABBHA, is a compilation of citations with abstracts on a wide range of related topics such as bird strike tolerance engineering, bird...
Self-Trail, J. M.; Robinson, M. M.; Edwards, L. E.; Powars, D. S.; Wandless, G. A.; Willard, D. A.
An exceptional Paleocene-Eocene boundary section occurs in a cluster of six short (Paleocene Aquia Formation and silty clay of the lower Eocene Marlboro Clay. Sediment samples were analyzed for carbon and oxygen isotopes, percent calcium carbonate, calcareous nannofossils, planktic and benthic foraminifera, dinoflagellates, pollen, and lithology. A well-defined carbon isotope excursion (CIE) documents a gradual negative shift in δ13C values that starts below the lithologic break between the Aquia Formation and the Marlboro Clay. A benthic foraminifer extinction event, reduction of calcareous nannofossil assemblages, and change in core color from gray to alternating gray and pink also occurs within the CIE transition. These alternating changes in color coincide with cyclic peaks in the carbon isotope and percent calcium carbonate curves, where gray color corresponds to a positive shift in carbon isotope values and to a corresponding increase in percent benthic and planktic foraminifera. The upper third of the Marlboro Clay is barren of all calcareous microfossil material, although the presence of foraminiferal molds and linings proves that deposition occurred in a marine environment. Co-occurrence of the dinoflagellates Apectodinium augustum and Phthanoperidinium crenulatum at the top of the Marlboro Clay suggests that the Marlboro Clay at Mattawoman Creek is truncated. This is corroborated by the absence in the Marlboro of specimens of the calcareous nannofossil Rhomboaster-Discoaster assemblage, which is restricted to early Eocene Zone NP9b. Based on planktic/benthic foraminifera ratios, deposition of sediments at Mattawoman Creek occurred predominantly in an inner neritic environment, at water depths between 25-50 m. Occasional deepening to approximately 75m (middle neritic environment) occurred in the early Eocene, as represented by the basal Marlboro Clay. The planktic/benthic ratio, however, could also be affected by surface productivity and/or river runoff
Vo, Hong D.; Schiffner, Ingo; Srinivasan, Mandyam V.
It is essential for birds to be agile and aware of their immediate environment, especially when flying through dense foliage. To investigate the type of visual signals and strategies used by birds while negotiating cluttered environments, we presented budgerigars with vertically oriented apertures of different widths. We find that, when flying through narrow apertures, birds execute their maneuvers in an anticipatory fashion, with wing closures, if necessary, occurring well in advance of the aperture. When passing through an aperture that is narrower than the wingspan, the birds close their wings at a specific, constant distance before the aperture, which is independent of aperture width. In these cases, the birds also fly significantly higher, possibly pre-compensating for the drop in altitude. The speed of approach is largely constant, and independent of the width of the aperture. The constancy of the approach speed suggests a simple means by which optic flow can be used to gauge the distance and width of the aperture, and guide wing closure. PMID:27270506
Peterjohn, Bruce G.
Located along Delaware Bay and the Atlantic coast, the state of Delaware’s significance for bird conservation has been well established for decades. The extensive tidal habitats and marshes bordering Delaware Bay host shorebird and waterbird populations of hemispheric importance, and protecting these populations has become an urgent conservation priority in recent years. Other habitats found in the state vary from barrier beaches to dry coniferous woods on the coastal plain and mesophytic communities along the Piedmont in the north, allowing a diverse avifauna to prosper within a small geographic area. Ornithologists and birders have actively studied birds within the state for more than a century, but surprisingly, no single reference has provided a complete summary of the status and distribution of the state’s birds until publication of the Birds of Delaware.Review info: Birds of Delaware. By Gene K. Hess, Richard L. West, Maurice V. Barnhill III, and Lorraine M. Fleming, 2000. ISBN: 0-8229-4069-8, 635 pp.
Polycyclic aromatic hydrocarbons (PAH) are present throughout the global environment and are produced naturally and by activities of humans. Effects of PAH on birds have been determined by studies employing egg injection, egg immersion, egg shell application, single and multiple oral doses, subcutaneous injection, and chemical analysis of field-collected eggs and tissue. The four-to six-ring aromatic compounds are the most toxic to embryos, young birds, and adult birds. For embryos, effects include death, developmental abnormalities, and a variety of cellular and biochemical responses. For adult and young birds, effects include reduced egg production and hatching, increased clutch or brood abandonment, reduced growth, increased organweights, and a variety of biochemical responses. Trophic level accumulation is unlikely. Environmental exposure to PAH in areas of high human population or habitats affected by recent petroleum spills might be sufficient to adversely affect reproduction. Evidence of long-term effects of elevated concentrations of environmental PAH on bird populations is very limited and the mechanisms of effect are unclear.
Ramachandran, Reshma; McDaniel, Cd
Parthenogenesis or "virgin birth" is embryonic development in unfertilized eggs. It is a routine means of reproduction in many invertebrates. However even though parthenogenesis occurs naturally in even more advanced vertebrates, like birds, it is mostly abortive in nature. In fact, multiple limiting factors, such as delayed and unorganized development as well as unfavorable conditions developing within the unfertilized egg upon incubation, are associated with termination of progressive development of parthenogenetic embryos. In birds, diploid parthenogenesis is automictic and facultative producing only males. However, the mechanisms controlling parthenogenesis in birds are not clearly elucidated. Additionally, it appears from even very recent research that these mechanisms may hinder the normal fertilization process and subsequent embryonic development. For instance, virgin quail and turkey hens exhibiting parthenogenesis have reduced reproductive performance following mating. Also, genetic selection and environmental factors, such as live virus vaccinations, are known to trigger the process of parthenogenesis in birds. Therefore, parthenogenesis has a plausible negative impact on the poultry industry. Hence, a better understanding of parthenogenesis and the mechanisms that control it could benefit commercial poultry production. In this context, the aim of this review is to provide a complete overview of the process of parthenogenesis in birds.
Vo, Hong D; Schiffner, Ingo; Srinivasan, Mandyam V
It is essential for birds to be agile and aware of their immediate environment, especially when flying through dense foliage. To investigate the type of visual signals and strategies used by birds while negotiating cluttered environments, we presented budgerigars with vertically oriented apertures of different widths. We find that, when flying through narrow apertures, birds execute their maneuvers in an anticipatory fashion, with wing closures, if necessary, occurring well in advance of the aperture. When passing through an aperture that is narrower than the wingspan, the birds close their wings at a specific, constant distance before the aperture, which is independent of aperture width. In these cases, the birds also fly significantly higher, possibly pre-compensating for the drop in altitude. The speed of approach is largely constant, and independent of the width of the aperture. The constancy of the approach speed suggests a simple means by which optic flow can be used to gauge the distance and width of the aperture, and guide wing closure.
The manifest success of birds in flight over small and large distances, in confined quarters and also in gusty conditions has inspired admiration, investigation and sometimes imitation from the earthbound human. Birds occupy a range of scales (2 g - 12 kg in mass, and 0.05 - 3 m in wingspan) that overlaps certain micro air vehicle (MAV) designs and there is interest in whether some bird-like properties (flapping wings, deformable feathers, movable tails) might be useful or even necessary for successful MAVs. A bird with 5 cm mean chord flying at 8 m/s has a nominal Reynolds number of 2 - 3 x 10^4. This is an extremely inconvenient range for design, operation and analysis of lifting surfaces, even in steady motion, because their properties are very sensitive to boundary layer separation. The moderate- to high-amplitude flapping motions, together with the complex surface geometry and mechanical properties of the wings themselves lead to yet further challenges. This talk will review some of the theoretical and practical approaches towards understanding and analyzing the aerodynamics of various types of bird flight, including some recent research results that suggest that this effort is far from complete.
Fox, Anthony David; Heldbjerg, Henning; Nyegaard, Timme
Avian Introduced Alien Species (IAS) constitute a threat to the integrity of native biodiversity, the economy and human health, so here we briefly review some of the problems posed by such species around the world in relation to such bird species in Denmark. A new European Union Regulation...... on Invasive Alien Species implemented in January 2015 establishes a framework for actions to combat alien species, which requires Member States to prevent the spread of alien species, provide early warning and rapid responses to their presence and management of established alien species where they occur. We...... show the importance of mechanisms such as DOF’s (Dansk Ornitologisk Forening, BirdLife Denmark) Atlas project, Common Bird Census (breeding and wintering species) and DOFbasen to contribute data on the current geographical and numerical distribution of the few serious alien avian species already...
Nyegaard, Timme; Heldbjerg, Henning; Fox, Anthony David
Avian Introduced Alien Species (IAS) constitute a threat to the integrity of native biodiversity, the economy and human health, so here we briefly review some of the problems posed by such species around the world in relation to bird species in Denmark. A new European Union Regulation on Invasive...... Alien Species implemented in January 2015 requires a framework for actions to combat alien species, which requires Member States to prevent the spread of alien species, provide early warning and rapid responses to their presence and management of established alien species where they occur. We show...... the importance of mechanisms such as DOFs (Danish Ornithological Society, BirdLife Denmark) Atlas project, Common Bird Monitoring (breeding and wintering species) and DOFbasen to contribute data on the current geographical and numerical distribution of the few serious alien avian species already present...
Some occurrences of rare or unusual bird species reported by us in a previous paper (Ostrich 86(3): 267–276, 2015) are considered to be doubtful by Hogg and Vande weghe (Ostrich 88(1): 83–88, 2017). We believe that some of the problems raised are taxonomic. The main objective of our study was to obtain reliable ...
California Natural Resource Agency — The objective of this project was to digitally map the boundaries of Audubon California's Important Bird Areas (IBA). Existing Important Bird Areas identify critical...
The Wadden Sea region is one of the most important breeding areas in Western Europe for coastal breeding bird species. It is expected that management of salt marshes is important for successful conservation of breeding bird populations but the impact of management on the habitat quality for breeding birds is still not fully understood. In this study the effects of management on the three crucial habitat characteristics for breeding birds were studied on mainland salt marshes: effects of manag...
Sprague, Terry; Harrington, M. Elizabeth; Krogh, Carmen M. E.
Bird kill rate and disruption of habitat has been reported when industrial wind turbines are introduced into migratory bird paths or other environments. While the literature could be more complete regarding the documentation of negative effects on birds and bird habitats during the planning, construction, and operation of wind power projects,…
...-1231-9BPP] RIN 1018-AX09 Migratory Bird Permits; Changes in the Regulations Governing Migratory Bird... governing migratory bird rehabilitation in the United States. Before creation of those regulations.... FOR FURTHER INFORMATION CONTACT: Dr. George T. Allen, Division of Migratory Bird Management, U.S. Fish...
... 50 CFR Part 20 Migratory Bird Hunting; Proposed 2011-12 Migratory Game Bird Hunting Regulations (Preliminary) With Requests for Indian Tribal Proposals and Requests for 2013 Spring and Summer Migratory Bird... 50 CFR Part 20 [Docket No. FWS-R9-MB-2011-0014; 91200-1231-9BPP-L2] RIN 1018-AX34 Migratory Bird...
... DEPARTMENT OF THE INTERIOR Fish and Wildlife Service Migratory Birds; Take of Migratory Birds by... Forces to incidentally take migratory birds during approved military readiness activities without violating the Migratory Bird Treaty Act (MBTA). The Authorization Act provided this interim authority to...
Home; Journals; Resonance – Journal of Science Education; Volume 1; Issue 10. Microbiology as if Bird Watching. Milind G Watve. Classroom Volume 1 Issue 10 October 1996 pp 78-81. Fulltext. Click here to view fulltext PDF. Permanent link: http://www.ias.ac.in/article/fulltext/reso/001/10/0078-0081. Author Affiliations.
When high school students are asked what's the best part of science class, many will say it's the field trips. Students enjoy engaging in authentic, community-based science outside the classroom. To capitalize on this, Patrick Willis created the Bird Box Survey Project for his introductory field biology class. The project takes students…
nesting in colonies.· Some field tests of the model are discussed in Box 2. Some interesting ideas have been advanced by ecologists to explore the advantages which birds could derive from nesting in colonies and the most persuasive of these is the geometric model. Box 2. Field Tests of the Economic Defendability Model.
During the war I was able to identify some collections of birds from Sumatra, present in the Leiden Museum. These collections were brought together by E. Jacobson and W. C. van Heurn in the Padang Highlands in 1013; by W. Groeneveldt in the same area in 1914 and 1915; bij L. P. Cosquino de Bussy and
One day out of the great wilderness of the water there came an albatross. Circling the ship on .... and mathematicians. Conventional aerodynamic ... suspended in air has inspired over a dozen aerodynamic models. GENERAL I ARTICLE scaling concepts fail to explain subtle features of bird flight, as debated by over fifty ...
Williams, JB; Tieleman, BI; Williams, Joseph B.
We call into question the idea that birds have not evolved unique physiological adaptations to desert environments. The rate at which desert larks metabolize energy is lower than in mesic species within the same family, and this lower rate of living translates into a lower overall energy requirement
business opportunities for universities, laboratories and corpo- rations. Modern scientists seem to be getting increasingly ob- sessed with seasonal and sensational .... around SHAR and the Nelapattu Sanctuary, Professor Dhawan distilled the essence of bird flight for the expert and the neo- phyte alike. The sheer biometric ...
Solov'yov, Ilia; Hore, Peter J.; Ritz, Thorsten
increased interest following the proposal in 2000 that free radical chemistry could occur in the bird's retina initiated by photoexcitation of cryptochrome, a specialized photoreceptor protein. In the present paper we review the important physical and chemical constraints on a possible radical...
... 14 Aeronautics and Space 1 2010-01-01 2010-01-01 false Bird impact. 35.36 Section 35.36... STANDARDS: PROPELLERS Tests and Inspections § 35.36 Bird impact. The applicant must demonstrate, by tests or... 4-pound bird at the critical location(s) and critical flight condition(s) of a typical installation...
... 14 Aeronautics and Space 1 2010-01-01 2010-01-01 false Bird strike. 29.631 Section 29.631... STANDARDS: TRANSPORT CATEGORY ROTORCRAFT Design and Construction General § 29.631 Bird strike. The... safe landing (for Category B) after impact with a 2.2-lb (1.0 kg) bird when the velocity of the...
Metz, I.C.; Ellerbroek, J.; Muhlhausen, Thorsten; Kügler, D.; Hoekstra, J.M.
This paper presents a fast-time simulation environment for assessing the risk of bird strikes in aviation. An existing air traffic simulator was enhanced in order to simulate air and bird traffic simultaneously and to recognize collisions between birds and aircraft. Furthermore, a method was
Mihail, Michael D.; George, Thomas F.; Feldman, Bernard J.
This article describes an experiment that measures the forces acting on a flying bird during takeoff. The experiment uses a minimum of equipment and only an elementary knowledge of kinematics and Newton's second law. The experiment involves first digitally videotaping a bird during takeoff, analyzing the video to determine the bird's position as a…
van der Boon, A.; Kuiper, K.; van der Ploeg, R.; Cramwinckel, M.; Honarmand, M.; Sluijs, A.; Krijgsman, W.; Langereis, C. G.
A 500 kyr episode of 3-5 °C gradual global climate warming, some 40 Myr ago, has been termed the Middle Eocene climatic optimum (MECO). It has been associated with a rise in atmospheric CO2 concentrations, but the source of this carbon remains enigmatic. We show, based on new Ar-Ar ages of volcanic rocks in Iran and Azerbaijan, that the time interval spanning the MECO was associated with a massive increase in continental arc volcanism. We also collected almost 300 Ar-Ar and U-Pb ages from literature. Typically, U-Pb ages from the Eocene are slightly younger, by 3 Myr, than Ar-Ar ages. We observed that U-Pb ages are obtained mostly from intrusive rocks and therefore must reflect an intrusive stage that post-dated extrusive volcanism. Combining all ages for extrusive rocks, we show that they cluster around 40.2 Ma, exactly within the time span of the MECO (40.5-40.0 Ma). We estimate volumes of volcanism based on a shapefile of outcrops and average thickness of the sequences. We calculate CO2 estimates using a relation volcanism-CO2 that was earlier used for the Deccan traps (Tobin et al., 2017). Our calculations indicate that the volume of the Iranian middle Eocene volcanic rocks (estimated at 37000 km3) is sufficient to explain the CO2 rise during the MECO. We conclude that continental arc flare-up in the Neotethys subduction zone is a plausible candidate for causing the MECO.
Lossada, Ana C.; Giambiagi, Laura; Hoke, Gregory D.; Fitzgerald, Paul G.; Creixell, Christian; Murillo, Ismael; Mardonez, Diego; Velásquez, Ricardo; Suriano, Julieta
The Andes between 28° and 30°S represent a transition between the Puna-Altiplano Plateau and the Frontal/Principal Cordillera fold-and-thrust belts to the south. While significant early Cenozoic deformation documented in the Andean Plateau, deciphering the early episodes of deformation during Andean mountain building in the transition area is largely unstudied. Apatite fission track (AFT) and (U-Th-Sm)/He (AHe) thermochronology from a vertical and a horizontal transect reveal the exhumation history of the High Andes at 30°S, an area at the heart of this major transition. Interpretation of the age-elevation profile, combined with inverse thermal modeling, indicates that the onset of rapid cooling was underway by 35 Ma, followed by a significant decrease in cooling rate at 30-25 Ma. AFT thermal models also reveal a second episode of rapid cooling in the early Miocene ( 18 Ma) related to rock exhumation to its present position. Low exhumation between the rapid cooling events allowed for the development of a partial annealing zone. We interpret the observed Eocene rapid exhumation as the product of a previously unrecognized compressive event in this part of the Andes that reflects a southern extension of Eocene orogenesis recognized in the Puna/Altiplano. Renewed early-Miocene exhumation indicates that the late Cenozoic compressional stresses responsible for the main phase of uplift of the South Central Andes also impacted the core of the range in this transitional sector. The major episode of Eocene exhumation suggests the creation of significant topographic relief in the High Andes earlier than previously thought.
Lauretano, V.; Littler, K.; Polling, M.; Zachos, J. C.; Lourens, L. J.
Recent studies have shown that the Early Eocene Climatic Optimum (EECO) was preceded by a series of short-lived global warming events, known as hyperthermals. Here we present high-resolution benthic stable carbon and oxygen isotope records from ODP Sites 1262 and 1263 (Walvis Ridge, SE Atlantic) between ~ 54 and ~ 52 million years ago, tightly constraining the character, timing, and magnitude of six prominent hyperthermal events. These events, which include Eocene Thermal Maximum (ETM) 2 and 3, are studied in relation to orbital forcing and long-term trends. Our findings reveal an almost linear relationship between δ13C and δ18O for all these hyperthermals, indicating that the eccentricity-paced covariance between deep-sea temperature changes and extreme perturbations in the exogenic carbon pool persisted during these events towards the onset of the EECO, in accordance with previous observations for the Paleocene Eocene Thermal Maximum (PETM) and ETM2. The covariance of δ13C and δ18O during H2 and I2, which are the second pulses of the "paired" hyperthermal events ETM2-H2 and I1-I2, deviates with respect to the other events. We hypothesize that this could relate to a relatively higher contribution of an isotopically heavier source of carbon, such as peat or permafrost, and/or to climate feedbacks/local changes in circulation. Finally, the δ18O records of the two sites show a systematic offset with on average 0.2 ‰ heavier values for the shallower Site 1263, which we link to a slightly heavier isotopic composition of the intermediate water mass reaching the northeastern flank of the Walvis Ridge compared to that of the deeper northwestern water mass at Site 1262.
Akkiraz, M.S.; Kayseri, M.S.; Akgun, F. [Dokuz Eylul University, Izmir (Turkey). Dept. of Geological Engineering
In this study, the lignite-bearing Yoncali formation between Yozgat and Sorgun, in central Anatolia has been palynologically examined. Based on 37 outcrop samples, quantitative palynological studies recognized 64 genera and 136 palynoflora species in the palynological assemblage, which indicated a Middle-Late Eocene age. This paper also presents a quantitative palaeovegetation and palaeoclimate reconstruction for the Middle-Upper Eocene coal occurrences of Central Anatolia on the basis of palynomorph assemblages. The diversified floral and ecological characteristics of the pollen taxa indicates that the Middle-Upper Eocene formations in central Anatolia were characterized by the presence of a complex mangrove swamp with contributions by Nypa, Pelliciera, Avicennia, Diporites tszkaszentgydrgyi and dinoflagellate cysts which reflect warm climatic conditions. Lowland-riparian and montane elements are characterized by the dominance of Myricaceae, Symplocaceae, Icacinaceae, Quercus, Pinus and Castanea. Swamp-freshwater elements are represented by Sparganjaceae, Nymphaceae, Taxodjaceae, Cupressaceae and Nyssa as well as fern spores such as Osmundaceae and Gleicheniaceae. The calculations were performed with the help of the 'Coexistence Approach' method to climatically evaluate palynoflora from the Yozgat-Sorgun area. The obtained results have been compared to data derived from the application of the Coexistence Approach to other, already published Central Anatolian palynofloras of the same age. The results of the climatic inferences suggest that the palaeoclimatic conditions were in the megathermal zone, megatherm/mesotherm intermediate zone whereas mesothermic conditions prevailed in the montane region. Likewise, the results of mean annual range of temperatures indicate the influence of the Indian ocean, which enabled the development of the mangroves.
Villegas-Martín, Jorge; Netto, Renata Guimarães
The trace fossil Bichordites monastiriensis is found in early Eocene turbiditic sandstones of the upper-slope deposits from the Capdevila Formation in Los Palacios Basin, Pinar del Río region, western Cuba. The potential tracemakers of B. monastiriensis include fossil spatangoids from the family Eupatagidae. The record of Bichordites in the deposits from Cuba allows to suppose that Eupatagidae echinoids were the oldest potential tracemakers of Bichordites isp. and reinforce the hypothesis that the ichnological record are relevant in envisaging the evolutionary history of the spatangoids.
Full Text Available In this paper are described the coralgal facies identified in the Upper Eocene-Lower Oligocene limestone succession (Cozla Formation outcropping in two quarries at Letca and Rastoci (Sălaj district, Romania. In the studied profiles the coral and algae limestones are interlayered with bioclastic limestones with foraminifera. On the top of relatively deep water deposits, coral and algae crusts and dendritic corals coated by algae were deposited. The environment registered a gradual deepening, the deposits being completely immersed, while bioclastic limestones with foraminifera were recurrently formed. This cycle is repeated, the whole succession being caracterized by several such “parasequences”.
Khozyem, H.; Adatte, T.; Keller, G.; Spangenberg, J. E.; Bajpai, S.; Samant, B.; Mathur, S.
The Paleocene Eocene Thermal Maximum (PETM, 55.5Ma) is globally related with the extinction of deep benthic foraminifera, the diversification of both plancktic foraminifera and mammals. In India, the tempo and timing of mammals dispersion, their association with the PETM or EECO (Early Eocene Climatic Optimum) and the India- Asia collision remain uncertain (Smith et al., 2006 Clementz, 2010). Three sections located in north and northwest India have been studied using sedimentology, micropaleontology, mineralogy (bulk and clay mineralogy) and geochemistry (stable isotopes, major and trace elements, organic matter). Both PETM and ETM2 (second Eocene Thermal Maximum, 53.7Ma), a short-lived warming episode that followed the PETM, are globally marked by a pronounced δ13Ccarb and org negative peak. Both isotopic excursions have been recognized in the Vastan and Tarkeswhar lignite mines (Cambay basin, Gujarat), above the main mammals bearing level. The lower shift is located above the first lignite seam (=lignite 2 of Sahni et al, 2004, 2009) and corresponds to the transition from continental to shallow marine conditions marked by benthic foraminifera and bivalves. The upper excursion appears to be linked to the ETM2 and corresponds to a second marine incursion containing bivalves, benthic (Nummulites burdigalensis) and planktic foraminifera located below the second lignite seam (lignite 1 of Sahni et al, 2004, 2009). A single but very pronounced δ13Corg peak has been detected in the Giral Lignite mine (Barmer, Rajhastan), around 6m above the vertebrates bearing level and may correspond to the PETM. This correlation is confirmed by palynological data (Tripathi et al., 2009, Sahni et al., 2004, 2009) and more particularly by an acme in the dinoflagellate Apectodinium that globally characterizes the PETM interval (Sluijs et al. 2007). Our micropaleontological data combined with stable carbone isotopes indicate the presence of both PETM and ETM2 events and constrain the
Carriol, René-Pierre; Bonde, Niels Christensøn; Jakobsen, Sten Lennart
New taxa of thoracican cirripedes are recorded from the Eocene Mo Clay of northwest Jutland, Denmark, namely Stipilepas molerensis Carriol n. gen., n. sp., a scalpelliform gooseneck barnacle, and Plesiobrachylepas jutlandica Carriol n. gen., n. sp., a brachylepadomorph sessile form. This material...... is of importance in that the former not only represents the first extinct example of the tribe Ashinkailepadini, it also is the second fossil neolepadine, while the latter adds to the diversity of the family Brachylepadidae, which previously embraced only three genera. The geological context of these taxa...
Tanaka, Gengo; Parker, Andrew R.; Siveter, David J.; Maeda, Haruyoshi; Furutani, Masumi
The exceptionally preserved eyes of an Eocene dolichopodid fly contained in Baltic amber show remarkable detail, including features at micrometre and submicrometre levels. Based on this material, we establish that it is likely that the neural superposition compound eye existed as far back as 45 Ma. The ommatidia have an open rhabdom with a trapezoidal arrangement of seven rhabdomeres. Such a structure is uniquely characteristic of the neural superposition compound eye of present-day flies. Optical analysis reveals that the fossil eyes had a sophisticated and efficient optical system. PMID:19129103
A revision of the genus Helius Lepeletier & Serville, 1828 (Diptera: Limoniidae) from Baltic amber (Eocene) is presented. Redescriptions of 5 species, Helius formosus Krzemiński, 1993, Helius linus Podenas, 2002, Helius minutus (Loew, 1850), Helius mutus Podenas, 2002, Helius pulcher (Loew, 1850) of this genus from Baltic amber are given and documented by photographs and drawings. Four new species of the genus Helius from Baltic amber are described: Helius gedanicus sp. nov., Helius hoffeinsorum sp. nov., Helius similis sp. nov., Helius fossilis sp. nov. A key to species of Helius from Baltic amber is provided. Patterns morphological evolution and the aspects evolutionary history of Helius are discussed.
Smit, Matthijs Arjen; Hacker, Bradley; Lee, Jeffrey
Tectonic reconstructions of the Himalayan orogeny depend on the age at which crustal thickening commenced. To investigate this age, we analyzed garnet from middle crustal rocks exposed in the north Himalayan Mabja and Kangmar gneiss domes of Tibet using Lu-Hf geochronology. Garnet yielded Lu-Hf a...... recrystallization during subsequent high-temperature ductile extension. The new Lu-Hf ages are the first to confirm that crustal thickening and contraction in the Tibetan Himalaya was broadly synchronous with the early Eocene collision between Greater India and the Eurasian plate...
The Paleocene and Eocene are characterized by strong greenhouse climates. Atmospheric CO2 concentrations and global temperatures were much higher than today. The period from 60 to 50 million years ago (Ma) is marked by a gradual warming trend of ~8 ºC in the deep ocean. The interval from 56 to 50 Ma is further characterized by several transient perturbations of the carbon cycle. Essentially, these perturbations, or “hyperthermal” events mark phases of rapid (103 – 104 years) warming, associat...
Rubincam, David Parry
Drag from hydrogen in the interstellar cloud which formed Gould's Belt may have sent interplanetary dust particle (IDPs) and small meteoroids with embedded helium to the Earth, perhaps explaining part the helium-3 flux increase seen in the sedimentary record near the Eocene-Oligocene transition. Assuming the Solar System passed through part of the cloud, IDPs in the inner Solar System may have been dragged to Earth, while dust and small meteoroids in the asteroid belt up to centimeter size may have been dragged to the resonances, where their orbital eccentricities were pumped up into Earth-crossing orbits; however, this hypotheses does not explain the Popigai and Chesapeake Bay impacts.
Full Text Available Valid publication of new names of fossil plant taxa published since 1 January 1996 requires a diagnosis or description in English, besides other requirements included in the International Code of Nomenclature for algae, fungi, and plants (Melbourne Code adopted by the Eighteenth International Botanical Congress, Melbourne, Australia, July 2011 (McNeill et al. 2012. In order to validate names published from the late Eocene flora of the Staré Sedlo Formation, North Bohemia, diagnosed only in German (Knobloch et al. 1996, English translations are provided, including references to the type material and further relevant information.
The rodent fauna of Fuenferrada contains two species of Theridomyidae and three species of Gliridae. It is compared with the Early Oligocene fauna of Olalla 4A, and with the Late Eocene fauna of Aguatón 2D. The absence of Cricetidae serves to determine its age as latest Eocene.
Fan, Majie; Constenius, Kurt N.; Dettman, David L.
The paleoelevation and size of the North America Cordilleran orogen during the late Cretaceous-Paleogene contractional and subsequent extensional tectonics remain enigmatic. We present new estimates of paleorelief of the northern Cordilleran orogenic front during the middle and late Eocene using oxygen isotope compositions of unaltered molluscan fossils and paleosol carbonates in the Kishenehn basin. Bounded by several mountains ranges to the east, the Kishenehn basin was a half graben developed during middle Eocene to early Miocene collapse of the Cordilleran orogen. These mollusk taxa include three sympatric groups with affinities to wet tropical, semi-arid subtropical, and temperate environments. Our reconstructed surface water δ18O values vary between -19.8‰ and -6.3‰ (VSMOW) during the middle and late Eocene. The large differences in paleoenvironments and surface water δ18O values suggest that the catchment of the Kishenehn basin was at variable elevation. The estimated paleorelief between the basin and the surrounding mountains, based on both Rayleigh condensation model and predictions of Eocene precipitation isotope values using an isotope-enabled global climate model, is ∼4 km, and the basin floor was <1.5 km high. This high topography and high relief paleogeography suggest that the Cordilleran orogenic front reached an elevation of at least 4 km, and the crust thickness may have reached more than 55 km before Eocene gravitational collapse. We attribute the maintenance of high Eocene topography to the combination of an inherited thick crust, thermal uplift caused by mantle upwelling, and isostatic uplift caused by removing lower lithosphere or oceanic slab.
Dallanave, Edoardo; Tauxe, Lisa; Muttoni, Giovanni; Rio, Domenico
We describe a scenario of climate forcing on sedimentation recorded in the late Paleocene-early Eocene Cicogna marine section from the Belluno Basin (NE Italy). Previously published magneto-biostratigraphic data revealed that the ˜81 m Cicogna section extends from Chron C25r to Chron C23r spanning the NP7/NP8-NP12 nannofossil zones (˜52.2-56.6 Ma). Using previously published rock magnetic data, augmented by data from this study, we describe and thoroughly discuss a pronounced increase of hematite (relative to maghemite or magnetite) between ˜54.9 and 54.6 Ma immediately above the Paleocene-Eocene boundary, followed by a second, long-term increasing trend from ˜54 Ma up to ˜52.2 Ma in the early Eocene. This hematite is essentially of detrital origin, insofar as it is associated with a strong shallow bias of paleomagnetic inclinations, and is interpreted to have formed on land by the weathering of Fe-bearing silicates and other primary minerals. We speculate that the warm and humid climate typical of the Paleocene-Eocene thermal maximum (PETM, ˜54.9 Ma) as well as of the warming trend leading to the early Eocene climatic optimum (EECO; ˜52-50 Ma) enhanced continental weathering of silicate rocks with the consequent production, transport, and sedimentation of detrital hematite grains. This hypothesis is confirmed by a statistical correlation between the rock magnetic properties and global climate as revealed by a standard benthic oxygen isotope record from the literature. Our temporal coupling between oxidation state of sedimentary magnetic phases and global climate is therefore consistent with the existence in the Paleocene-Eocene of the silicate weathering negative feedback mechanism for the long-term stabilization of the Earth's surface temperature.
Dipineto, Ludovico; De Luca Bossa, Luigi Maria; Russo, Tamara Pasqualina; Cutino, Eridania Annalisa; Gargiulo, Antonio; Ciccarelli, Francesca; Raia, Pasquale; Menna, Lucia Francesca; Fioretti, Alessandro
A total of 170 birds of prey admitted to two Wildlife Rescue and Rehabilitation Centers of Italy were examined. Birds were divided by diurnal (n = 15) and nocturnal (n = 7) species, sampled by cloacal swabs, and examined for Campylobacter spp. by cultural and molecular methods. Campylobacter spp. were isolated in 43 out of the 170 (25.3%) birds of prey examined. Among these, 43/43 (100%) were identified as Campylobacter jejuni and 10/43 (23.3%) were identified as Campylobacter coli recovered from mixed infections. Diurnal birds of prey showed a significantly higher prevalence value (P = 0.0006) for Campylobacter spp. than did nocturnal birds of prey.
Full Text Available Nearly one in five bird species has separate breeding and overwintering distributions, and the regular migrations of these species cause a substantial seasonal redistribution of avian diversity across the world. However, despite its ecological importance, bird migration has been largely ignored in studies of global avian biodiversity, with few studies having addressed it from a macroecological perspective. Here, we analyse a dataset on the global distribution of the world's birds in order to examine global spatial patterns in the diversity of migratory species, including: the seasonal variation in overall species diversity due to migration; the contribution of migratory birds to local bird diversity; and the distribution of narrow-range and threatened migratory birds. Our analyses reveal a striking asymmetry between the Northern and Southern hemispheres, evident in all of the patterns investigated. The highest migratory bird diversity was found in the Northern Hemisphere, with high inter-continental turnover in species composition between breeding and non-breeding seasons, and extensive regions (at high latitudes where migratory birds constitute the majority of the local avifauna. Threatened migratory birds are concentrated mainly in Central and Southern Asia, whereas narrow-range migratory species are mainly found in Central America, the Himalayas and Patagonia. Overall, global patterns in the diversity of migratory birds indicate that bird migration is mainly a Northern Hemisphere phenomenon. The asymmetry between the Northern and Southern hemispheres could not have easily been predicted from the combined results of regional scale studies, highlighting the importance of a global perspective.
Cox, Daniel T C; Gaston, Kevin J
At a time of unprecedented biodiversity loss, researchers are increasingly recognizing the broad range of benefits provided to humankind by nature. However, as people live more urbanized lifestyles there is a progressive disengagement with the natural world that diminishes these benefits and discourages positive environmental behaviour. The provision of food for garden birds is an increasing global phenomenon, and provides a readily accessible way for people to counter this trend. Yet despite its popularity, quite why people feed birds remains poorly understood. We explore three loosely defined motivations behind bird feeding: that it provides psychological benefits, is due to a concern about bird welfare, and/or is due to a more general orientation towards nature. We quantitatively surveyed households from urban towns in southern England to explore attitudes and actions towards garden bird feeding. Each household scored three Likert statements relating to each of the three motivations. We found that people who fed birds regularly felt more relaxed and connected to nature when they watched garden birds, and perceived that bird feeding is beneficial for bird welfare while investing time in minimising associated risks. Finally, feeding birds may be an expression of a wider orientation towards nature. Overall, we found that the feelings of being relaxed and connected to nature were the strongest drivers. As urban expansion continues both to threaten species conservation and to change peoples' relationship with the natural world, feeding birds may provide an important tool for engaging people with nature to the benefit of both people and conservation.
Laboratory experiments have shown the magnetic compass to be wavelength dependent: migratory birds require light from the blue-green part of the spectrum for magnetic compass orientation, whereas red light (visible long-wavelength disrupts magnetic orientation. We designed a field study to test if and how changing light color influenced migrating birds under field conditions. We found that nocturnally migrating birds were disoriented and attracted by red and white light (containing visible long-wavelength radiation, whereas they were clearly less disoriented by blue and green light (containing less or no visible long-wavelength radiation. This was especially the case on overcast nights. Our results clearly open perspective for the development of bird-friendly artificial lighting by manipulating wavelength characteristics. Preliminary results with an experimentally developed bird-friendly light source on an offshore platform are promising. What needs to be investigated is the impact of bird-friendly light on other organisms than birds.
Full Text Available This article explores the waning days of the Ottoman Empire and emergence of the modern state system in the early twentieth century Levant from the purview of Kenizé Mourad's self-narrative Regards from the Dead Princess: Novel of a Life. A work of history and literary fiction, Mourad's novel is an account of the last remnants of a secular Levantine culture, the story of a crumbling empire, and the personal tale of a young woman and her exiled imperial family strewn about the continents, torn between Lebanon, Europe, and the Indian subcontinent.
Morgan, C.D.; Bereskin, S.R.
The oil-productive Eocene Green River Formation in the central Uinta Basin of northeastern Utah is divided into five distinct intervals. In stratigraphically ascending order these are: 1) Uteland Butte, 2) Castle Peak, 3) Travis, 4) Monument Butte, and 5) Beluga. The reservoir in the Uteland Butte interval is mainly lacustrine limestone with rare bar sandstone beds, whereas the reservoirs in the other four intervals are mainly channel and lacustrine sandstone beds. The changing depositional environments of Paleocene-Eocene Lake Uinta controlled the characteristics of each interval and the reservoir rock contained within. The Uteland Butte consists of carbonate and rare, thin, shallow-lacustrine sandstone bars deposited during the initial rise of the lake. The Castle Peak interval was deposited during a time of numerous and rapid lake-level fluctuations, which developed a simple drainage pattern across the exposed shallow and gentle shelf with each fall and rise cycle. The Travis interval records a time of active tectonism that created a steeper slope and a pronounced shelf break where thick cut-and-fill valleys developed during lake-level falls and rises. The Monument Butte interval represents a return to a gentle, shallow shelf where channel deposits are stacked in a lowstand delta plain and amalgamated into the most extensive reservoir in the central Uinta Basin. The Beluga interval represents a time of major lake expansion with fewer, less pronounced lake-level falls, resulting in isolated single-storied channel and shallow-bar sandstone deposits.
van Kempen, Monique M L; Smolders, Alfons J P; Lamers, Leon P M; Roelofs, Jan G M
In order to understand the physicochemical mechanisms that could explain the massive growth of Azolla arctica in the Eocene Arctic Ocean, we carried out a laboratory experiment in which we studied the interacting effects of rain and wind on the development of salinity stratification, both in the presence and in the absence of a dense Azolla cover. Additionally, we carried out a mesocosm experiment to get a better understanding of the nutrient cycling within and beneath a dense Azolla cover in both freshwater and brackish water environments. Here we show that Azolla is able to create a windproof, small-scale salinity gradient in brackish waters, which allows for efficient recycling of nutrients. We suggest that this mechanism ensures the maintenance of a large standing biomass in which additional input of nutrients ultimately result in a further expansion of an Azolla cover. As such, it may not only explain the extent of the Azolla event during the Eocene, but also the absence of intact vegetative Azolla remains and the relatively low burial efficiency of organic carbon during this interval.
Monique M L van Kempen
Full Text Available In order to understand the physicochemical mechanisms that could explain the massive growth of Azolla arctica in the Eocene Arctic Ocean, we carried out a laboratory experiment in which we studied the interacting effects of rain and wind on the development of salinity stratification, both in the presence and in the absence of a dense Azolla cover. Additionally, we carried out a mesocosm experiment to get a better understanding of the nutrient cycling within and beneath a dense Azolla cover in both freshwater and brackish water environments. Here we show that Azolla is able to create a windproof, small-scale salinity gradient in brackish waters, which allows for efficient recycling of nutrients. We suggest that this mechanism ensures the maintenance of a large standing biomass in which additional input of nutrients ultimately result in a further expansion of an Azolla cover. As such, it may not only explain the extent of the Azolla event during the Eocene, but also the absence of intact vegetative Azolla remains and the relatively low burial efficiency of organic carbon during this interval.
Goldner, A; Herold, N; Huber, M
Two main hypotheses compete to explain global cooling and the abrupt growth of the Antarctic ice sheet across the Eocene-Oligocene transition about 34 million years ago: thermal isolation of Antarctica due to southern ocean gateway opening, and declining atmospheric CO2 (refs 5, 6). Increases in ocean thermal stratification and circulation in proxies across the Eocene-Oligocene transition have been interpreted as a unique signature of gateway opening, but at present both mechanisms remain possible. Here, using a coupled ocean-atmosphere model, we show that the rise of Antarctic glaciation, rather than altered palaeogeography, is best able to explain the observed oceanographic changes. We find that growth of the Antarctic ice sheet caused enhanced northward transport of Antarctic intermediate water and invigorated the formation of Antarctic bottom water, fundamentally reorganizing ocean circulation. Conversely, gateway openings had much less impact on ocean thermal stratification and circulation. Our results support available evidence that CO2 drawdown--not gateway opening--caused Antarctic ice sheet growth, and further show that these feedbacks in turn altered ocean circulation. The precise timing and rate of glaciation, and thus its impacts on ocean circulation, reflect the balance between potentially positive feedbacks (increases in sea ice extent and enhanced primary productivity) and negative feedbacks (stronger southward heat transport and localized high-latitude warming). The Antarctic ice sheet had a complex, dynamic role in ocean circulation and heat fluxes during its initiation, and these processes are likely to operate in the future.
Full Text Available The Eocene volcanic rocks from the southwest of the Jandaq (Kuh-e-Godar-e-Siah, Central-East Iran micro-continent are andesitic basalt and andesite in composition. These rocks contain xenoliths with granulitic mineralogy. Mineral assemblage of these xenoliths is plagioclase + phlogopite + corundum + sillimanite + chlorite + phengite with granublastic, poiklioblastic and foliated textures in the pick metamorphic condition. Thermometry of phlogopite in these xenoliths suggests the average temperature 782oC. The characteristics of the xenoliths are consistent with the granolitic facies metamorphism of the Al-saturated Si-undersaturated crustal sediments at the lower crust condition. Melting of these granulites forms the magma which crystallized the S-type granitoids. Differentiation and crystallization of this magma causes the S-type granite formation. Therefore, the S-type granites in the study area are probably generated from melting of the granulites parts of which brought to the surface as xenoliths by Eocene magmatism in south of the Jandaq (Kuh-e-Godar-e-Siah. S-type granites in the study area are located along the Doruneh, Chupanan and Aeirakan faults in the Aeirakan area and Jandaq ophiolite. These granites are the source of uranium, thorium and uranium ore in southwest of the Aeirakan mountain.
Bond, Mariano; Tejedor, Marcelo F.; Campbell, Kenneth E.; Chornogubsky, Laura; Novo, Nelson; Goin, Francisco
The platyrrhine primates, or New World monkeys, are immigrant mammals whose fossil record comes from Tertiary and Quaternary sediments of South America and the Caribbean Greater Antilles. The time and place of platyrrhine origins are some of the most controversial issues in primate palaeontology, although an African Palaeogene ancestry has been presumed by most primatologists. Until now, the oldest fossil records of New World monkeys have come from Salla, Bolivia, and date to approximately 26 million years ago, or the Late Oligocene epoch. Here we report the discovery of new primates from the ?Late Eocene epoch of Amazonian Peru, which extends the fossil record of primates in South America back approximately 10 million years. The new specimens are important for understanding the origin and early evolution of modern platyrrhine primates because they bear little resemblance to any extinct or living South American primate, but they do bear striking resemblances to Eocene African anthropoids, and our phylogenetic analysis suggests a relationship with African taxa. The discovery of these new primates brings the first appearance datum of caviomorph rodents and primates in South America back into close correspondence, but raises new questions about the timing and means of arrival of these two mammalian groups.
Gammon, Paul R.; James, Noel P.; Pisera, Andrjez
Siliceous spongolite and spiculite are generally interpreted as deep- and/or cold-water deposits, largely because modern analogs are found mostly in cold- and/or deep-water environments. However, extensive shallow-water, biosiliceous accumulations are present in late Eocene age deposits of southern Western Australia. The attributes and paleoenvironmental setting of these accumulations frame the specific climatic and/or oceanographic constraints that allow such shallow neritic sediments to form. The generally fine grained siliciclastic-biosiliceous sediments accumulated in a nearshore, shallow-water setting, around numerous islands, and offshore from a warm-temperate, rainforest-covered upland. Sponges compose a lithistid-hexactinellid assemblage of Mesozoic Tethyan affinity, confined to deep water in the modern ocean, that migrated into neritic paleoenvironments during late Eocene time. Lithistids dominated shoreface environments and were reworked into sponge-conglomerate beaches, while hexactinellids flourished offshore in muddy subshoreface environments. These deposits also contain common soft demosponge spicules, and a rare but otherwise normal marine calcareous biota. The biosiliceous sediments grade offshore into bryozoan-rich marls and limestones. Sponges outcompeted the calcareous benthos in these inner shelf environments because of elevated amounts of land-derived nutrients that included high dissolved silica and a calm hydrodynamic setting. These deposits suggest that neither temperature nor water depth are critical factors for prolific sponge colonization. The rarity of such deposits in the rock record likely reflects the necessity for the coincidence of all of these factors, or they are misinterpreted.
Full Text Available We analyse systematic fractures occurring in sandstone beds in Eocene flsch of the Slovenian coastal area. Two nearly perpendicular fracture sets were identifid: fractures F1 are generally NW-SE oriented, wellexpressed and predominately planar, whereas fractures F2 are NE-SW-striking, shorter, more irregular in shape, and terminate against the F1 set. The average orientation of both sets does not change signifiantly in a coastal transect crossing all principal structural domains of the area. We analysed fracture spacing with respect to layer thickness and determined fracture spacing index for both fracture sets. We interpret both fracture sets as tensional (Mode I joints originating in two distinct extensional episodes. Set F1 is older and formed in NE-SW directed tension which we correlate with the well-documented regional post-Dinaric orogen-perpendicular extension of presumably mid-Miocene age. Set F2 formed in NW-SE oriented tension, which is compatible with previously documented NE-SW-striking normal faults occurring in the area, but was so far not documented elsewhere. We interpret that F1 fractures predate folding and thrusting in the coastal belt. Earlier, Eocene-Oligocene Dinaric thrusting therefore did not signifiantly affect the coastal area, whereas post-F1 shortening, associated with northward indentation and underthrusting of the Adria microplate, did not commence before late Miocene.
Chen, Iju; Manchester, Steven R; Chen, Zhiduan
Well-preserved seeds from the early Eocene of Wutu, Shandong, China are assigned to the genus Nuphar (Nymphaeaceae) based on morphology and anatomy. The seeds of Nuphar wutuensis sp. nov. are ellipsoidal to ovoid, 4-5 mm long with a clearly visible raphe ridge, and a truncate apex capped by a circular operculum ca. 1 mm in diameter bearing a central micropylar protrusion. These features, along with the testa composed of a uniseriate outer layer of equiaxial pentagonal to hexagonal surface cells and a middle layer 4-6 cells thick composed of thick-walled, periclinally elongate sclereids, correspond to the morphology and anatomy of extant Nuphar and distinguish this fossil species from all other extant and extinct genera of Nymphaeales. These seeds provide the oldest record for the genus in Asia and are supplemented by a similar well-preserved specimen from the Paleocene of North Dakota, USA. These data, together with the prior recognition of Brasenia (Cabombaceae) in the middle Eocene, indicate that the families Nymphaeaceae and Cabombaceae had differentiated by the early Tertiary.
Squires, R.L.; Wilson, M.
A 600-m thick unnamed marine, predominantly transition-zone siltstone unit along the south flank of the Mount Pinos uplift, in the northern Lockwood Valley area, previously has been suggested to be early Eocene (Capay Stage) in age at its base. This present study shows the entire unit to be this age. Unconformably overlying the pre-Tertiary granite basement is 30 m of unfossiliferous muddy siltstone that grades upward into 50 m of very fine sandstone with rarely fossiliferous lenses of medium to coarse sandstone. Gradationally above the sandstone is 100 m of muddy siltstone with less rarely fossiliferous lenses of conglomeratic sandstone. Macrofossil collections made at 10 localities in these lower 180 m yielded a sparse fauna of subtropical shallow-marine gastropods and bivalves, as well as rare specimens of discocyclinid foraminifera. from 180 to 500 m above the base of the section is unfossiliferous siltstone with local occurrences of lower shoreface, alternating laminated and bioturbated very fine sandstone. The uppermost 100 m of the section is siltstone with rarely fossiliferous lenses of fine to medium sandstone. Collections made at five localities yielded subtropical shallow-marine mollusks. Evidence of a West Coast provincial molluscan Capay Stage (early Eocene) age for all the fossiliferous beds of the siltstone unit is the presence of Turritella andersoni, a species diagnostic of this stage. Commonly associated mollusks are Cryptoconus cooperi, Cylichnina tantilla, Ectinochilus (Macilentos) macilentus, and Turritella buwaldana. Unconformably overlying the unit is the Oligocene-lower Miocene nonmarine Plush Ranch Formation.
Sluijs, A.; Schouten, S.; Pagani, M.; Woltering, M.; Brinkhuis, H.; Damste, J.S.S.; Dickens, G.R.; Huber, M.; Reichart, G.-J.; Stein, R.; Matthiessen, J.; Lourens, L.J.; Pedentchouk, N.; Backman, J.; Moran, K.; Clemens, S.; Cronin, T.; Eynaud, F.; Gattacceca, J.; Jakobsson, M.; Jordan, R.; Kaminski, M.; King, J.; Koc, N.; Martinez, N.C.; McInroy, D.; Moore, T.C.; O'Regan, M.; Onodera, J.; Palike, H.; Rea, B.; Rio, D.; Sakamoto, T.; Smith, D.C.; St John, K.E.K.; Suto, I.; Suzuki, N.; Takahashi, K.; Watanabe, M. E.; Yamamoto, M.
The Palaeocene/Eocene thermal maximum, ???55 million years ago, was a brief period of widespread, extreme climatic warming, that was associated with massive atmospheric greenhouse gas input. Although aspects of the resulting environmental changes are well documented at low latitudes, no data were available to quantify simultaneous changes in the Arctic region. Here we identify the Palaeocene/Eocene thermal maximum in a marine sedimentary sequence obtained during the Arctic Coring Expedition. We show that sea surface temperatures near the North Pole increased from ???18??C to over 23??C during this event. Such warm values imply the absence of ice and thus exclude the influence of ice-albedo feedbacks on this Arctic warming. At the same time, sea level rose while anoxic and euxinic conditions developed in the ocean's bottom waters and photic zone, respectively. Increasing temperature and sea level match expectations based on palaeoclimate model simulations, but the absolute polar temperatures that we derive before, during and after the event are more than 10??C warmer than those model-predicted. This suggests that higher-than-modern greenhouse gas concentrations must have operated in conjunction with other feedback mechanisms-perhaps polar stratospheric clouds or hurricane-induced ocean mixing-to amplify early Palaeogene polar temperatures. ?? 2006 Nature Publishing Group.
Tang, Maoyun; Liu-Zeng, Jing; Hoke, Gregory D.; Xu, Qiang; Wang, Weitao; Li, Zhanfei; Zhang, Jinyu; Wang, Wei
The topography evolution of the southeastern Tibetan Plateau provides a constraint for evaluating various geodynamic models of plateau formation. We reconstruct the Paleocene to Eocene paleoelevation of the Cenozoic Gonjo basin in the Qiangtang terrane, using oxygen and carbon stable isotopic results from pedogenic carbonates of the lower Ranmugou Formation. Lithofacies associations indicate that the lower Ranmugou Formation was deposited in alluvial fan and fluvial floodplain environments. U-Pb dating of volcanics within the middle Ranmugou Formation constrains the deposition of the lower Ranmugou Formation as prior to 43.2 Ma. Paleoelevations are calculated using both a thermodynamic model and an empirical relationship. The empirical relationship of elevation-δ18O is determined from a series of modern water samples. Calculated paleoelevations indicate that the Gonjo basin attained a minimum average elevation of 2100-2500 m in the early Eocene. Together with recent paleoaltimetry studies in the region, it can be concluded that the crust of southeastern Tibetan Plateau was already thickened by that time. The calculation is based on the section in northern Gonjo basin, where δ18O values of paleosol nodules appear to be unaltered, but we cannot rule out the possibility that δ18O values of the pedogenic carbonates had been partially reset. Our preliminary results favor tectonic models compatible with pre-Miocene uplift of the SE-central Tibetan Plateau.
Hovan, Steven A.; Rea, David K.
Deep Sea Drilling Project (DSDP) Site 215 provides an expanded section across the Paleocene/Eocene boundary, the most complete mid-latitude sequence from a Southern Hemisphere location in the Indo-Pacific area. The events of this transition occurred during a span of about 1.2 m.y. Oxygen isotope values derived from benthic foraminiferal calcite decrease by about 1.0‰, a decrease most likely related to warming of deep ocean waters. Turnovers of benthic foraminifera accompany δ18O changes and culminate in the predominant extinction event at the end of the Paleocene Epoch. Carbon isotope ratios also shift dramatically toward lighter values near the end of the Paleocene, beginning about 0.45 m.y. after oxygen isotope values start to change. The intensity of Southern Hemisphere atmospheric circulation as recorded by grain sizes of eolian particles shows a large and rapid reduction beginning another 0.45 m.y. later. A significant reduction of zonal wind strength at the Paleocene/Eocene boundary, until now observed only at Northern Hemisphere locations, appears to have been a global phenomenon related to decreased latitudinal thermal gradients occasioned by more effective poleward heat transport via the deep ocean.
Full Text Available Original biostratigraphic and sedimentologic data concerning the type-section of the Albidona Formation (Liguride Complex, Southern Apennines, Italy are presented and discussed. Since its definition in 1962, this lithostratigraphic unit has been the object of controversial interpretations in terms of age attribution and paleotectonic significance. Based on cross-observations performed on calcareous nannofossils and palynomorphs, we conclude that the Albidona Formation must be assigned to the Eocene and, based on these evidences, we make a review of the previous biostratigraphic literature. We further recognise four different turbidite systems (named A to D, bounded by minor stratigraphic hiatuses, that are characterised by different sedimentary facies assemblages and petrofacies. The overall vertical arrangement demonstrates that the Albidona Formation was deposited in a tectonically mobile basin during a phase of deformation that is older than the apenninic deformation and must be likely referred to the alpine tectonics of the Calabrian arc. Moreover, the relationship with the underlying folded unit suggests for the Albidona Formation the significance of an episutural deposit relevant to a Paleogene deformation that affected the older units of the Liguride Complex. By considering stratigraphic and sedimentological features, we suggest a correlation of the Albidona Formation with analogue turbidite suites cropping out in the Apennines, discussing their significance in the context of the Eocene tectonic paroxysm in the Mediterranean area.
Hook, Robert W.; Warwick, Peter D.; Swanson, Sharon M.; Hackley, Paul C.; Warwick, Peter D.; Karlsen, Alexander K.; Merrill, Matthew D.; Valentine, Brett J.
The lignite deposits within the upper Eocene Yegua Formation (Claiborne Group) and the overlying Jackson Group are among the coal resources that were not quantitatively assessed as part of the U.S. Geological Survey's (USGS) National Coal Resource Assessment (NCRA) program in the Gulf Coastal Plain coal province. In the past, these lignite-bearing stratigraphic units often have been evaluated together because of their geographic and stratigraphic proximity (Fisher, 1963; Kaiser, 1974; Kaiser et al., 1980; Jackson and Garner, 1982; Kaiser, 1996) (Figures 1, 2). The term “Yegua-Jackson trend“ is used informally herein for the lignite-bearing outcrops of these Late Eocene deposits in Texas. Lignite beds in the Yegua-Jackson trend generally are higher both in ash yield and sulfur content than those of the underlying Wilcox Group (Figure 2). Recent studies (Senkayi et al., 1987; Ruppert et al., 1994; Warwick et al., 1996, 1997) have shown that some lignite beds within the Yegua-Jackson trend contain partings of volcanic ash and host elevated levels of trace elements that have been identified as potentially hazardous air pollutants (HAPs) in the United States Clean Air Amendments of 1990. Lignite beds within the Yegua Formation are thin (less than or equal to 6 ft) and laterally discontinuous in comparison with most Wilcox Group deposits (Ayers, 1989a); in contrast, the Jackson Group lignite beds range up to 12 ft in total thickness and are relatively continuous laterally, extending nearly 32 mi along strike.
Fırat Demirkaya, Demirkan; Aksoy, Necmettin; Üner, Serkan
Continental collision zones are sites of intense crustal deformation resulting in different geological processes such as crustal thickening, regional contraction, and volcanic activity. The East Anatolian Plateau emerges from the compression and following collision between the Eurasian and Arabian plates. Numerous basins were formed by this compressional tectonism, including the Pasinler, Muş, and Lake Van basins. The E-W trending Aǧrı Basin is located at north of the collision zone formed by compressional tectonism in Eocene and shaped by the volcanism that was active since Miocene. The basins fill initiates with Middle Eocene to Lower Miocene shallow marine deposits composing of reefal limestones, mudstones, and sandstones. Ongoing compressional regime and related regional uplift caused regression and latter transition from shallow marine to terrestrial environments in Middle Miocene took place. Marine units are unconformably overlain by Upper Miocene to Pliocene lacustrine and fluvial deposits. The Quaternary sedimentation is represented by alluvial fan deposits derived from the northern margin of the basin and young fluvial sediments. Volcanic and volcanoclastic products are frequently observed in Miocene to Recent basin deposits. Currently, E-W trending thrust faults, forming the Aǧrı Basin, continue to control the shape and depositional processes of the basin. Deformation of alluvial fans and channel shifting of the Murat and Seryan rivers which longitudinally passing the basin are the prominent geomorphological indicators of this tectonic activity.
Vahlenkamp, Maximilian; Niezgodzki, Igor; De Vleeschouwer, David; Bickert, Torsten; Harper, Dustin; Lohmann, Gerrit; Pälike, Heiko; Zachos, James C.
The role of the Atlantic Meridional Overturning Circulation (AMOC) as a key player for abrupt climatic changes (e.g. Heinrich Stadials) during the Pleistocene is relatively well constrained. However, the timing of the onset of a „modern" North Atlantic Deepwater (NADW) formation are still debated: Recent estimates range from the middle Miocene to the Early Eocene [Davies et al., 2001, Stoker et al., 2005, Hohbein et al., 2012] and are mainly based on the seismic interpretation contourite drifts. Another understudied aspect of the AMOC is its behavior during climatic variations on orbital time scales and under different climatic boundary conditions (icehouse vs hothouse). IODP Expedition 342 drilled carbonate-rich sequences from sediment drifts offshore Newfoundland that cover the middle Eocene with high sedimentation rates ( 3 cm/ kyr). We present a 2 Myr long stable carbon and oxygen isotope record of benthic foraminifera nuttalides truempyi spanning magnetochron C20r in unprecedented resolution (Nielsen, T., and Shannon, P. M., 2005, Neogene stratigraphy and the sedimentary and oceanographic development of the NW European Atlantic margin: Marine and Petroleum Geology, v. 22, no. 9, p. 977-1005. Hohbein, M. W., Sexton, P. F., and Cartwright, J. A., 2012, Onset of North Atlantic Deep Water production coincident with inception of the Cenozoic global cooling trend: Geology, v. 40, no. 3, p. 255-258.
Iwajomo, Soladoye Babatola
Bird migration has attracted a lot of interests over past centuries and the methods used for studying this phenomenon has greatly improved in terms of availability, dimension, scale and precision. In spite of the advancements, relatively more is known about the spring migration of trans......-Saharan migrants than autumn migration. Information about the behavior and interactions of migrants during the nonbreeding season in sub-Saharan Africa is also scarce for many species. Furthermore, very little is known about intra-African migration. This thesis summarizes my research on the autumn migration...... of birds from Europe to Africa and opens up the possibility of studying intra-African migration. I have used long-term, standardized autumn ringing data from southeast Sweden to investigate patterns in biometrics, phenology and population trends as inferred from annual trapping totals. In addition, I...
The earliest Antarctic Cenozoic (late early Eocene) bryozoan fossil records, recognized in the La Meseta Fm. on Seymour Island (Antarctic Peninsula) are connected with the major K/T phase of the cheilostome evolution, clear preponderance of cerioporoidean cyclostomes along with abundant occurrence of microporoideans, umbonulomorphs and lepraliomorphs. The presence of the loose, small zooecia of the cheilostome bryozoans in the lowermost part of this formation, systematically includes the buguloids and catenicelloideans such as e.g. Beanidae, Catenicellidae, Savignyellidae and Calwelliidae families, which in the present day are widely distributed in the tropical-warm latitudes mostly in the shallow-marine settings (Hara 2015). Undoubtedly, the occurrence of over 90% of the warm-loving multilamellar cyclostomes with the relatively slow growing rate is connected with a short-term episode in the lower part of the La Meseta Formation (Telm1), during their long in situ evolution. The recently recognized biota in the middle part of the La Meseta Formation (Telm4 and Telm5) on the NE side of the Seymour Island, reveal a presence of the microporoideans of the Micropora as well as free-living lunulitiforms belonging to the Lunulites and Otionellina genera, which developed disc-shaped colonies (Hara et al. 2015, in review). They are dominated in the Telm5, along with the new umbonulomorph of the family Brydonellidae Uharella seymourensis, found as an epilithozoic, encrusting bryozoan occurring in a loose residuum of the siliciclastic sediments. Environmentally, Recent, free-living lunulitids are known to occur in warm, shallow-shelf conditions, at temperatures of 10-29˚C, on coarse, sandy to muddy bottom with low to moderate deposition in fairly high velocity current regime and they are overwhelmingly associated with sand fauna environments. The dominance of the lunulitiform colonies in the Telm4-5 may suggests the shallow-water setting in this middle part of the formation
Zhao, Xuebing; Chen, Mingyan; Wu, Zhaolu; Wang, Zijiang
Many migratory bird species fly during the night (nocturnal migrants) and have been shown to display some phototaxis to artificial light. During 2006 to 2009, we investigated phototaxis in nocturnal migrants at Jinshan Yakou in Xinping County (N23°56', E101°30'; 2400 m above sea-level), and at the Niaowang Mountain in Funing County (N23°30', E105°35'; 1400 m above sea-level), both in the Yunnan Province of Southwest China. A total of 5069 birds, representing 129 species, were captured by mist-netting and artificial light. The extent of phototaxis effect on bird migration was examined during all four seasons, three phases of the moon, and under two weather conditions (mist and wind). Data were statistically analyzed to determine the extent to which these factors may impact phototaxis of nocturnal migrants. The results point to phototaxis in birds migrating in the spring and autumn, especially in the autumn. Furthermore, migrating birds were more readily attracted to artificial lights during nights with little moonlight, mist, and a headwind. Regardless of the initial orientation in which birds flew, either following the wind or against the wind, birds would always fly against the wind when flying towards the light. This study broadens our understanding of the nocturnal bird migration, potentially resulting in improved bird ringing practices, increased awareness, and better policies regarding bird protection.
Abels, H.A.; Lauretano, V.; van Yperen, Anna E.; Hopman, Tarek; Zachos, J.C.; Lourens, L.J.; Gingerich, P.D.; Bowen, G.J.
Transient greenhouse warming events in the Paleocene and Eocene were associated with the addition of isotopically light carbon to the exogenic atmosphere–ocean carbon system, leading to substantial environmental and biotic change. The magnitude of an accompanying carbon isotope excursion (CIE) can
With the realization that the Arctic Ocean was covered with enormous quantities of the aquatic floating fern Azolla 49 Myrs ago, new questions regarding the Eocene conditions facilitating these blooms arose. This dissertation describes the reconstruction of paleo-environmental conditions
Nielsen, Jan Kresten; Milàn, Jesper; Mesfun, Daniel
A cliff exposure of the Eocene Lillebælt Clay Formation, on the Røsnæs peninsula of Zealand, Denmark, has yielded a diverse trace-fossil assemblage. The trace fossils are described formally for the first time and assigned to Phymatoderma melvillensis, unnamed clusters of small burrows, Ophiomorph...
Donald R. Prothero
Full Text Available The Eocene-Oligocene Makah Formation and subjacent middle Eocene Hoko River Formation of the northwestern Olympic Peninsula, Washington, yield mollusks, crustaceans, foraminifera, and early neocete whales; their age has never been precisely established. We sampled several sections; most samples showed a stable single-component remanence held largely in magnetite and passed a Class I reversal test. The upper Refugian (late Eocene and lower Zemorrian (early Oligocene rocks at Baada Point correlate with Chron C13r (33.7–34.7 Ma and Chron C12r (30–33 Ma. The Ozette Highway section of the Makah Formation spanned the early Refugian to late Refugian, with a sequence that correlates with Chrons C15r-C13r (33.7–35.3 Ma, and a long reversed early Zemorrian section that correlates with Chron C12r (30–33 Ma. The type section of the Hoko River Formation correlates with Chron C18r (40.0–41.2 Ma. The area sampled shows about 45∘ of post-Oligocene counterclockwise tectonic rotation, consistent with results obtained from the Eocene-Oligocene rocks in the region.
Wade, B.S.; Houben, A.J.P.; Quaijtaal, W.; Schouten, S.; Rosenthal, Y.; Miller, K.G.; Katz, M.E.; Wright, J.D.; Brinkhuis, H.
The Eocene-Oligocene transition (EOT; ca. 33–34 Ma) was a time of pronounced climatic change, marked by the establishment of continental-scale Antarctic ice sheets. The timing and extent of temperature change associated with the EOT is controversial. Here we present multiproxy EOT climate records
Short-term episodes of global warming (hyperthermal events) were superimposed on the warming trend into the Early Eocene Climate Optimum (EECO). The Paleocene-Eocene Thermal Maximum (PETM; 56 Ma) was the most extreme, followed by Eocene Thermal Maximum-2 and -3 (ETM2: 1.8 myr, ETM3: 3.1 myr post-PETM). Hyperthermals are characterized by negative carbon isotope excursions (CIEs, emission of isotopically light carbon in the ocean-atmosphere), negative oxygen isotope excursions (global warming) and carbonate dissolution (ocean acidification). Sensitivity of biota to environmental changes due to carbon emissions can be evaluated by studying their response to hyperthermals of different magnitude. Deep-sea benthic foraminiferal records across PETM, ETM2 and -3 are available for Site 1262 (3600 m) and 1263 (1500m) on SE Atlantic Walvis Ridge. Benthic foraminifera (carbonate and agglutinated) are absent in the carbonate-free PETM clay-layer (Site 1262: 65 kyr; Site1263: 10 kyr). Deep-sea benthic foraminifera suffered extinction and diversity loss at the start of the PETM, as they did globally, with diversity recovering only partially. Stable isotope records show a larger PETM-CIE and amount of warming at Site 1263 than global average (McCarren et al., 2008), and warming was more pronounced at Site 1263 than at 1262 during ETM2 (Jennions et al., 2015) and ETM3 (Roehl et al., 2005). During ETM2 and -3, carbonate dissolution affected the sites, both remaining between CCD and lysocline. Assemblages were more severely affected (larger drop in benthic foraminiferal accumulation rates, BFAR) at the shallower site, opposite to expected if caused mainly by carbonate corrosivity. The large decrease in BFAR indicates a decline in food arrival at the sea floor, more pronounced at the shallower site, as supported by changes in relative and absolute abundance of species, and more pronounced at ETM2 than at ETM3. Greater warming at intermediate depths could have been caused by ocean
Smith, M. Elliot; Carroll, Alan R.; Scott, Jennifer J.; Singer, Brad S.
Repeated global reorganizations of carbon cycling and biotic, oceanic and terrestrial processes occurred during the Early Eocene, and appear to have been paced by cyclic variations in the eccentricity of the Earth's orbit. The phase relationship(s) between insolation variation, terrestrial paleoclimate, and atmospheric pCO2 during these events remains enigmatic however, due to their poorly constrained timing relative to specific orbital configurations. Here we use tiered interpolation between radioisotopic ages and paleomagnetic polarity chrons to compare high-resolution δ13C and lithofacies records from the Wilkins Peak Member of the Green River Formation of western North America to a Fe-intensity XRF record from the western Atlantic Ocean, and to numerical solutions for Earth's orbital configuration. Wilkins Peak Member lithofacies stacking patterns record cyclic geomorphic responses to insolation and climate fluctuations, spanning an interval of 1.8 Ma. Previous macrostratigraphic analyses using 40Ar/39Ar and U-Pb ash bed ages indicate that these cycles reflect long and short eccentricity modulation of precession. Hydrologic variance appears to have occurred inversely with intervals of maximum sediment advection, with carbonate- and evaporite-dominated lacustrine modes during eccentricity maxima, and siliciclastic-dominated alluvial modes during eccentricity minima. Stable carbon isotope analyses of 126 meters of Wilkins Peak Member strata reveal a regular ∼5 per mil oscillation between high-δ13C lacustrine modes and low-δ13C alluvial modes. Tiered interpolation between paleomagnetically characterized terrestrial ash beds facilitates the integration of 11 radioisotopic ages with the geomagnetic polarity timescale, resulting in significant expansion of chron C23 and shortening of chron C22 relative to timescales based on seafloor magnetic anomaly profiles. The new proposed timescale permits direct comparison of terrestrial and marine climate proxy records
Breitfeld, H. T.; Hennig, J.; BouDagher-Fadel, M.; Hall, R.
The offshore Sarawak Basin NW of North Sarawak is a major hydrocarbon province in SE Asia. A very thick sedimentary sequence of Oligocene to ?Early Miocene age, named Cycle 1, is an important hydrocarbon source and reservoir. Despite numerous wells the stratigraphy and tectonic history is not very well understood. The Nyalau Formation of onshore North Sarawak is the supposed equivalent of the offshore Cycle 1 sequence. The Nyalau Formation is a thick sedimentary sequence of mainly tidal to deltaic deposits. The formation is dominated by well-bedded sandstone-mudstone alternations and thicker sandstones with abundant bioturbation. The sandstones are predominantly arenaceous. Various lithic fragments and feldspar indicate multiple sources and fresh input from igneous and metamorphic rocks. Interbedded thin limestone beds and marls yielded Early Miocene foraminifera for the upper part of the succession. Zircons separated from the sandstones yielded mainly Cretaceous and Triassic ages. The Triassic is the dominant age population. The Nyalau Formation conformably overlies the Buan Shale and the Tatau Formation, and in places unconformably overlies the Belaga Formation. The Belaga Formation is part of the Rajang Group that represents remnants of a large submarine fan deposited in the Late Cretaceous to Eocene in Central Sarawak. In contrast to the Nyalau Formation, the majority of zircons from the Rajang Group have Cretaceous ages. This marks an important change in provenance at the major unconformity separating the Belaga and Nyalau Formations. This unconformity was previously interpreted as the result of an orogeny in the Late Eocene. However, there is no evidence for a subduction or collision event at this time in Sarawak. We interpret it to mark plate reorganisation in the Middle Eocene and name it the Rajang Unconformity. Borneo is the principal source of Cretaceous zircons which were derived from the Schwaner Mountains and West Sarawak. The dominant Triassic zircon
Burke, K. D.; Williams, J. W.
As the world warms due to rising greenhouse gas concentrations, the climate system is moving toward a state without precedent in the historical record. Various past climate states have been proposed as potential analogues or model systems for the coming decades, including the early to middle Holocene, the last interglacial, the middle Pliocene, and the early Eocene. However, until now, such comparisons have been qualitative. To compare these time periods to the projected climate states for the 21st and 22nd centuries, we conduct a climate similarity analysis using the standardized Euclidean distance metric (SED) and seasonal means of surface air temperature and precipitation. We make this future-to-past comparison using 30-year mean climatologies, for every decade between 2020 and 2280 AD (27 total comparisons). The list of past earth system states includes the historical period (1940-1970 AD), a pre-industrial control (ca. 1850), the middle Holocene (ca. 6 ka), the last glacial maximum (ca. 21 ka), the last interglacial (ca. 125 ka), the middle Pliocene (ca. 3 Ma), and the early Eocene (ca. 50-55 Ma). To reduce uncertainties resulting from choice of earth system model, analyses are based on simulations from three earth system models (HadCM, CCSM, NASA/GISS Model-E), using in part experiments from PMIP2, PMIP3/CMIP5, EoMIP, and PlioMIP. Results are presented for two representative concentration pathways (RCP's 4.5, 8.5). By 2050 AD, the most common past climate analogue is sourced from the Pliocene for RCP 8.5, while by 2190 AD, the Eocene becomes the source of the most common past climate analogue. For RCP 4.5, in which radiative forcings stabilize this century, the Pliocene becomes the most important past climate analogue by 2100 AD. Low latitude climates are the first to most closely resemble these past earth warm periods. The mid-latitudes then follow this pattern by the end of the 22nd century. Although no past state of the earth system is a perfect analogue
Willard, D. A.
Proxy records from Eocene hyperthermals provide evidence for rates and magnitudes of environmental changes associated with these events, as well as their impacts on terrestrial and marine ecosystems. The Middle Eocene Climatic Optimum (MECO) occurred ~40 Million years ago (Ma) and lasted ~500,000 years, and previous research has documented changes in marine and terrestrial biotas during and after this event. Cores collected in sediment drifts on Newfoundland Ridge off the coast of eastern Canada during IODP Expedition 342 recovered expanded sections of the middle Eocene, with sedimentation rates of 1-2 cm/kyr. We present results from pollen, palynofacies, and dinocyst analyses from Sites 1408 (41.438'N, 49.786'W, 3022 mwd), 1409 (41.296'N, 49.233'W, 3501 mwd), and 1410 (41.328'N, 49.170'W, 3387 mwd), spanning an interval that includes the MECO. Palynological assemblages are well preserved throughout the middle Eocene at Sites 1408 and 1410, whereas carbonate-rich samples from Site 1409 were completely barren of palynomorphs. Substantial variability in dinocyst species composition and diversity, pollen assemblages, and organic palynofacies were observed within the MECO event. Fluctuations in concentrations of opaque organic matter, insect fragments, and pollen and spores from terrestrial vegetation reflect changes in terrigenous influx, intensity of the hydrologic cycle, and source vegetation. Dinocyst assemblage shifts are correlated with changes in productivity, nutrient supply, and salinity. Integration of palynological data with other proxies will provide further insights into correlations between increased terrigenous input and eutrophication, leads and lags between terrestrial and marine responses to climate fluctuations, and environmental stability during the Middle Eocene.
... birds in those parts of the United States adapted thereto where the same have become scarce or extinct, and also to regulate the introduction of American or foreign birds or animals in localities where they...
Established in 2009, the Green Energy Act aims to increase the use of renewable energy sources including wind, water, solar and bioenergy in Ontario. The development of these resources is a major component of the province's plan, which aims to mitigate the contribution to climate change and to involve the Ontario's economy in the improvement of the quality of the environment. The Green Energy Act also considers as important the implementation of a coordinated provincial approval process, suggesting the integration of all Ministry requirements into a unique process during the evaluation of newly proposed renewable energy projects. The Ministry of the Environment's Renewable Energy Approval Regulation details the requirements for wind power projects involving significant natural features. Birds are an important part of Ontario's biodiversity and, according to the Ministry of Natural Resources, their habitats are considered as significant wildlife habitat (SWH). The Renewable Energy Approval Regulation and this guideline are meant to provide elements and guidance in order to protect bird SWH during the selection of a location of wind power facilities. . 27 refs., 1 tab., 2 figs.
Conway, Courtney J.
Little is known about the population status of many marsh-dependent birds in North America but recent efforts have focused on collecting more reliable information and estimates of population trends. As part of that effort, a standardized survey protocol was developed in 1999 that provided guidance for conducting marsh bird surveys throughout North America such that data would be consistent among locations. The original survey protocol has been revised to provide greater clarification on many issues as the number of individuals using the protocol has grown. The Standardized North American Marsh Bird Monitoring Protocol instructs surveyors to conduct an initial 5-minute passive point-count survey followed by a series of 1-minute segments during which marsh bird calls are broadcast into the marsh following a standardized approach. Surveyors are instructed to record each individual bird from the suite of 26 focal species that are present in their local area on separate lines of a datasheet and estimate the distance to each bird. Also, surveyors are required to record whether each individual bird was detected within each 1-minute subsegment of the survey. These data allow analysts to use several different approaches for estimating detection probability. The Standardized North American Marsh Bird Monitoring Protocol provides detailed instructions that explain the field methods used to monitor marsh birds in North America.
Steiner, Gerald; Bartels, Thomas; Krautwald-Junghanns, Maria-Elisabeth; Koch, Edmund
Birds are traditionally classified as male or female based on their anatomy and plumage color as judged by the human eye. Knowledge of a bird's gender is important for the veterinary practitioner, the owner and the breeder. The accurate gender determination is essential for proper pairing of birds, and knowing the gender of a bird will allow the veterinarian to rule in or out gender-specific diseases. Several biochemical methods of gender determination have been developed for avian species where otherwise the gender of the birds cannot be determined by their physical appearances or characteristics. In this contribution, we demonstrate that FT-IR spectroscopy is a suitable tool for a quick and objective determination of the bird's gender. The method is based on differences in chromosome size. Male birds have two Z chromosomes and female birds have a W-chromosome and a Z-chromosome. Each Z-chromosome has approx. 75.000.000 bps whereas the W-chromosome has approx. 260.00 bps. This difference can be detected by FT-IR spectroscopy. Spectra were recorded from germ cells obtained from the feather pulp of chicks as well as from the germinal disk of fertilized but non-bred eggs. Significant changes between cells of male and female birds occur in the region of phosphate vibrations around 1080 and 1120 cm-1.
Full Text Available We review characteristics of birds in boreal forests in the context of their ecological sustainability under both natural and anthropogenic disturbances. We identify the underlying ecological factors associated with boreal bird populations and their variability, review the interactions between boreal bird populations and disturbance, and describe some tools on how boreal bird populations may be conserved in the future. The boreal system has historically been an area with extensive disturbance such as fire, insect outbreaks, and wind. In addition, the boreal system is vulnerable to global climate change as well as increasing pressure on forest and water resources. Current knowledge indicates that birds play an important role in boreal forests, and sustaining these populations affords many benefits to the health of boreal forests. Many issues must be approached with caution, including the lack of knowledge on our ability to mimic natural disturbance regimes with management, our lack of understanding on fragmentation due to logging activity, which is different from permanent conversion to other land uses such as agriculture or residential area, and our lack of knowledge on what controls variability in boreal bird populations or the linkage between bird population fluctuations and productivity. The essential role that birds can provide is to clarify important ecological concerns and variables that not only will help to sustain bird populations, but also will contribute to the long-term health of the boreal forest for all species, including humans.
BIRDS, PARASITES, CLASSIFICATION, ARTHROPODA , DISTRIBUTION, FLIGHT, MITES, ECOLOGY, LICE, INDIA, JAPAN, TAIWAN, CHINA, PHILIPPINES, THAILAND, BORNEO, INDONESIA, SINGAPORE, ASIA, TABLES(DATA), HANDBOOKS, ARMY RESEARCH, DIPTERA.
The time and rate of bird strike incidences, species composition and number of birds involved in strikes, and the behaviour of birds were recorded at Bole International Airport from December 1994 to December 1995. The gut contents of the birds struck by aircraft were analyzed. A total of 33 bird strikes were recorded during ...
The Royal Library owns one of the most exceptional works in book history, an original edition of John James Audubon Birds of America. This edition, in a format called “double elephant folio” was published from 1827 to 1838. On basis of existing literature, this article briefly describes the work...... is now owned by the Royal Library. The acquisition of the Danish set by the Classen’s Library is examined by analyzing previously unpublished letters and is described for the first time, although not comprehensively, in this article. The provenance of this work, as described by Waldemar Fries in 1973...
Bralower, Timothy J.; Self-Trail, Jean
The Paleocene-Eocene Thermal Maximum (PETM) is characterized by a transient group of nannoplankton, belonging to the genus Discoaster. Our investigation of expanded shelf sections provides unprecedented detail of the morphology and phylogeny of the transient Discoasterduring the PETM and their relationship with environmental change. We observe a much larger range of morphological variation than previously documented suggesting that the taxa belonged to a plexus of highly gradational morphotypes rather than individual species. We propose that the plexus represents malformed ecophenotypes of a single species that migrated to a deep photic zone refuge during the height of PETM warming and eutrophication. Anomalously, high rates of organic matter remineralization characterized these depths during the event and led to lower saturation levels, which caused malformation. The proposed mechanism explains the co-occurrence of malformed Discoaster with pristine species that grew in the upper photic zone; moreover, it illuminates why malformation is a rare phenomenon in the paleontological record.
Kyte, F. T.; Shukolyukov, A.; Hildebrand, A. R.; Lugmair, G. W.; Hanova, J.
The late Eocene (approx. 35 Ma) was a time of multiple large body impacts superimposed within an interval of dust accretion. At least two spherule layers are preserved in deep sea sediments: North American microtektites and the slightly older cpx spherules. The two largest impact structures in the Cenozoic, the 45 km Chesapeake Bay structure and the 100 km Popigai structure, are indicated as the respective spherule sources. Enhanced 3He concentrations extending across a 3 m.y. duration in upper Eocene sediments from the Massignano quarry in Italy indicate accretion of dust over this time interval. To characterize one of the impactors we have analyzed splits from the 125-250 micron cpx spherules from ODP 709C, and two fractions from the Massignano layer. Splits of each sample were analyzed for minor and trace elements by instrumental activation analysis (INAA) including Ir, Cr, Fe, Ni, and Co. Additional splits were analyzed for their Cr-isotopic composition, using thermal ionization mass spectrometry. Significant concentrations of Ir were found in all samples, with the highest levels in the Massignano coarse sample and the lowest in the 709C sample. In all cases, element/Ir ratios are much higher than in chondritic meteorites; this may reflect elemental fractionation due to preferential concentration of Cr in spinel growing in the impact fireball. The Cr-isotopic compositions of the 709C and Massignano coarse samples are both non-terrestrial with a positive epsilon 53, indicating a 53Cr/52Cr ratio higher than in terrestrial materials. Microprobe surveys showed that the Massignano samples had significant fine grained oxide grains (mixed with the spherules in the coarse sample) that were not Ni- or Cr-rich, including Ti-rich spinels (likely terrestrial contaminants). In contrast, the ODP 709C sample is a pure extract of generally well-preserved cpx spherules composed of clinopyroxene in a glass matrix. Both the Massignano coarse and the 709C cpx samples have
Grønbech, Gitte Lyng; Ibsen, Lars Bo; Nielsen, Benjaminn Nordahl
with a constant rate of strain were conducted on Søvind Marl in the Danish Continuous Loading Oedometer Apparatus, which enables test water to be applied at 200 kPa, and measures the horizontal stresses via pressure gauges placed in a stiff oedometer ring. The horizontal and vertical stresses were measured from......The present study evaluated earth pressure at rest, K0, in highly overconsolidated Eocene clay called Søvind Marl, which exhibits extremely high plasticity indices of up to 300%, a highly fissured structure, and preconsolidation stresses up to 6,800 kPa. Continuous Loading Oedometer (CLO) tests...... in situ stresses to various stress levels to estimate continuous K0 development in this highly overconsolidated clay. The normally consolidated earth pressure at rest was found for two different sample ages of Søvind Marl to be between 0.42 and 0.68. Results indicated the overconsolidated K0 reached...
Rose, Kenneth D; Holbrook, Luke T; Rana, Rajendra S; Kumar, Kishor; Jones, Katrina E; Ahrens, Heather E; Missiaen, Pieter; Sahni, Ashok; Smith, Thierry
Cambaytheres (Cambaytherium, Nakusia and Kalitherium) are recently discovered early Eocene placental mammals from the Indo-Pakistan region. They have been assigned to either Perissodactyla (the clade including horses, tapirs and rhinos, which is a member of the superorder Laurasiatheria) or Anthracobunidae, an obscure family that has been variously considered artiodactyls or perissodactyls, but most recently placed at the base of Proboscidea or of Tethytheria (Proboscidea+Sirenia, superorder Afrotheria). Here we report new dental, cranial and postcranial fossils of Cambaytherium, from the Cambay Shale Formation, Gujarat, India (~54.5 Myr). These fossils demonstrate that cambaytheres occupy a pivotal position as the sister taxon of Perissodactyla, thereby providing insight on the phylogenetic and biogeographic origin of Perissodactyla. The presence of the sister group of perissodactyls in western India near or before the time of collision suggests that Perissodactyla may have originated on the Indian Plate during its final drift toward Asia.
Edwards, Lucy E.
Dinocysts recovered from sediments related to the Chesapeake Bay impact structure in Virginia and the earliest Eocene suboxic environment in Maryland show strange and intriguing details of preservation. Features such as curled processes, opaque debris, breakage, microborings and cyst ghosts, among others, invite speculation about catastrophic depositional processes, rapid burial and biological and chemical decay. Selected specimens from seven cores taken in the coastal plain of Virginia and Maryland show abnormal preservation features in various combinations that merit illustration, description, discussion and further study. Although the depositional environments described are extreme, many of the features discussed are known from, or could be found in, other environments. These environments will show both similarities to and differences from the extreme environments here.
Wilf, Peter; Carvalho, Mónica R; Gandolfo, María A; Cúneo, N Rubén
The nightshade family Solanaceae holds exceptional economic and cultural importance. The early diversification of Solanaceae is thought to have occurred in South America during its separation from Gondwana, but the family's sparse fossil record provides few insights. We report 52.2-million-year-old lantern fruits from terminal-Gondwanan Patagonia, featuring highly inflated, five-lobed calyces, as a newly identified species of the derived, diverse New World genus Physalis (e.g., groundcherries and tomatillos). The fossils are considerably older than corresponding molecular divergence dates and demonstrate an ancient history for the inflated calyx syndrome. The derived position of these early Eocene fossils shows that Solanaceae were well diversified long before final Gondwanan breakup. Copyright © 2017, American Association for the Advancement of Science.
Grønbech, Gitte Lyng; Ibsen, Lars Bo; Nielsen, Benjaminn Nordahl
Determination of the preconsolidation stresses is a key tool in geotechnical engineering used to evaluate and estimate the behavior of soils. However, it has been proven difficult to accurately estimate the preconsolidation in highly fissured overconsolidated clays, due to the effect of the fissu......Determination of the preconsolidation stresses is a key tool in geotechnical engineering used to evaluate and estimate the behavior of soils. However, it has been proven difficult to accurately estimate the preconsolidation in highly fissured overconsolidated clays, due to the effect...... of the fissured structure of the clay. This article presents oedometer tests performed on Søvind Marl, a plastic Eocene clay, with a high presence of fissures and slickensides. Four incremental loading oedometer tests and four continuous loading oedometer tests were performed in order to determine...
Wolfe, Alexander P; Csank, Adam Z; Reyes, Alberto V; McKellar, Ryan C; Tappert, Ralf; Muehlenbachs, Karlis
We report exceptional preservation of fossil wood buried deeply in a kimberlite pipe that intruded northwestern Canada's Slave Province 53.3±0.6 million years ago (Ma), revealed during excavation of diamond source rock. The wood originated from forest surrounding the eruption zone and collapsed into the diatreme before resettling in volcaniclastic kimberlite to depths >300 m, where it was mummified in a sterile environment. Anatomy of the unpermineralized wood permits conclusive identification to the genus Metasequoia (Cupressaceae). The wood yields genuine cellulose and occluded amber, both of which have been characterized spectroscopically and isotopically. From cellulose δ(18)O and δ(2)H measurements, we infer that Early Eocene paleoclimates in the western Canadian subarctic were 12-17°C warmer and four times wetter than present. Canadian kimberlites offer Lagerstätte-quality preservation of wood from a region with limited alternate sources of paleobotanical information.
Goussin, Fanny; Cordier, Carole; Boulvais, Philippe; Guillot, Stéphane; Roperch, Pierrick; Replumaz, Anne
Cenozoic post-collisional volcanism of the Tibetan Plateau, emplaced on an accreted continental margin under compression, could bring important information regarding the edification of the Plateau. In this study, we combined petrography, whole rock geochemistry, stable isotopes and in situ mineral analysis to decipher the genesis of Eocene-Oligocene magmatic rocks from the Nangqian basin (35-38 Ma, [Spurlin et al., 2005; Xu et al., 2016]), located at the hinge between Central Tibet and the Eastern Indo-Asia Collision Zone. Our dataset includes potassic trachyandesites; amphibole-bearing potassic trachytes; and rare ultrapotassic (K2O/Na2O ≥ 4) mafic syenites. All samples have high REE abundances (La = 100 - 500 x primitive mantle). Fractionation of heavy REE (Gd/YbN > 3) indicates melting in the garnet stability field, and relative depletion in high-field strength elements (Nb, Ta) indicates a selective enrichment of the source by metasomatic fluids. This metasomatism event is also evidenced by the occurrence of re-equilibrated mantle xenocrysts of phlogopite (Mg# = 88 - 90 and Cr2O3 content = 0.9 - 1.82 wt%) in mafic syenites. Potassic trachyandesites have specific composition, with negative Zr-Hf anomaly and low Hf/Sm (0.2 - 0.4). Indeed, they include xenocrystic aggregates, composed of magmatic clinopyroxene, apatite and subordinate biotite and feldspar, with interstitial calcite and dolomite. δ18OV -SMOW (9.2 - 11.0 ) and δ13CV -PDB (-6.1 - -4.0 ) of these rocks indicate the presence of primary, mantle-derived carbonates. In situ analysis of the major and trace element compositions of the carbonates, clinopyroxenes and apatites further suggest that these aggregates represent cumulates of a carbonate-bearing magma. These xenoliths thus show that the lithospheric mantle was also metasomatized by CO2-rich fluids. Cenozoic carbonatites in China have been identified in Maoniuping in Western Sichuan (31.7 Ma), Lixian in the Western Qinlin (22-23 Ma), and
Schumann, Dirk; Raub, Timothy D.; Kopp, Robert E.; Guerquin-Kern, Jean-Luc; Wu, Ting-Di; Rouiller, Isabelle; Smirnov, Aleksey V.; Sears, S. Kelly; Lücken, Uwe; Tikoo, Sonia M.; Hesse, Reinhard; Kirschvink, Joseph L.; Vali, Hojatollah
We report the discovery of exceptionally large biogenic magnetite crystals in clay-rich sediments spanning the Paleocene–Eocene Thermal Maximum (PETM) in a borehole at Ancora, NJ. Aside from previously described abundant bacterial magnetofossils, electron microscopy reveals novel spearhead-like and spindle-like magnetite up to 4 μm long and hexaoctahedral prisms up to 1.4 μm long. Similar to magnetite produced by magnetotactic bacteria, these single-crystal particles exhibit chemical composition, lattice perfection, and oxygen isotopes consistent with an aquatic origin. Electron holography indicates single-domain magnetization despite their large crystal size. We suggest that the development of a thick suboxic zone with high iron bioavailability—a product of dramatic changes in weathering and sedimentation patterns driven by severe global warming—drove diversification of magnetite-forming organisms, likely including eukaryotes. PMID:18936486
Zachos, James C.; Wara, Michael W.; Bohaty, Steven; Delaney, Margaret L.; Petrizzo, Maria Rose; Brill, Amanda; Bralower, Timothy J.; Premoli-Silva, Isabella
The Paleocene-Eocene Thermal Maximum (PETM) has been attributed to a rapid rise in greenhouse gas levels. If so, warming should have occurred at all latitudes, although amplified toward the poles. Existing records reveal an increase in high-latitude sea surface temperatures (SSTs) (8° to 10°C) and in bottom water temperatures (4° to 5°C). To date, however, the character of the tropical SST response during this event remains unconstrained. Here we address this deficiency by using paired oxygen isotope and minor element (magnesium/calcium) ratios of planktonic foraminifera from a tropical Pacific core to estimate changes in SST. Using mixed-layer foraminifera, we found that the combined proxies imply a 4° to 5°C rise in Pacific SST during the PETM. These results would necessitate a rise in atmospheric pCO2 to levels three to four times as high as those estimated for the late Paleocene.
Francisco Javier Rodríguez-Tovar
Full Text Available Ichnological, sedimentological and geochemical analyses were conducted on the Eocene Frysjaodden Formation in order to interpret palaeoenvironment prodelta sediments in the Central Basin of Spitsbergen. Phycosiphon incertum is the exclusive ichnotaxon showing differences in size, distribution, abundance and density, and relation to laminated/bioturbated intervals. Large P. incertum mainly occur dispersed, isolated and randomly distributed throughout the weakly laminated/non-laminated intervals. Small P. incertum occur occasionally in patches of several burrows within laminated intervals or as densely packed burrows in thin horizons in laminated intervals or constituting fully bioturbated intervals that are several centimetres thick. Ichnological changes are mainly controlled by oxygenation, although the availability of benthic food cannot be discarded. Changes in oxygenation and rate of sedimentation can be correlated with the registered variations in the Bouma sequence of the distal turbiditic beds within prodeltal shelf sediments.
Matthews, Kara J.; Dietmar Müller, R.; Whittaker, Joanne; Flament, Nicolas; Seton, Maria
The late Cretaceous to mid Eocene history of the southwest and southernmost Pacific has been subject to starkly contrasting interpretations, ranging from relative tectonic quiescence with the Lord Howe Rise (LHR) being part of the Pacific plate to a dynamic subduction setting. In the first scenario the Tasman Sea would have formed as a consequence of divergence between the Pacific and Australian plates, whereas in the second scenario it would have formed as a marginal basin associated with subduction. The first scenario is supported by a number of arguments, including a lack of evidence for deformation and tectonic activity in New Zealand during this period and a geodynamic modelling inference, namely that the bend in the Hawaiian-Emperor chain can be better reproduced if the LHR is part of the Pacific plate. The second scenario is supported by regional plate kinematic models reconciling a variety of observations including back-arc basin formation and destruction through time and the history of arc-continent collisions. The primary problem with the first scenario is the use of a plate circuit that leaves relative motion between East and West Antarctica unconstrained, leading to an improbable history of periodic compression and extension. The main problem with the alternative scenario is a lack of sampled late Cretaceous volcanic arc rocks east of the LHR. We analysed available geological and geophysical data to constrain the locations of and movements along the plate boundaries in the southwest and southern Pacific from the late Cretaceous to mid Eocene, and assessed how Pacific plate motion is best quantified during this period. Geological and geophysical evidence suggests that a plate boundary separated the Pacific plate from the LHR. The distribution of lower mantle slab material that is imaged by seismic tomography beneath New Zealand is best explained if subduction occurred to the east of the LHR during the entire late Cretaceous to mid Eocene period. Rocks
Sustainable development requires the management and preservation of water resources indispensable for all human activities. When groundwater constitutes the main water resource, vulnerability maps therefore are an important tool for identifying zones of high pollution risk and taking preventive measures in potential pollution sites. The vulnerability assessment for the Eocene aquifer in the Moroccan basin of Oum Er-Rabia is based on the DRASTIC method that uses seven parameters summarizing climatic, geological, and hydrogeological conditions controlling the seepage of pollutant substances to groundwater. Vulnerability maps were produced by using GIS techniques and applying the “generic” and “agricultural” models according to the DRASTIC charter. Resulting maps revealed that the aquifer is highly vulnerable in the western part of the basin and areas being under high contamination risk are more extensive when the “agricultural” model was applied.
Engelbrecht, Andrea; Mörs, Thomas; Reguero, Marcelo A.; Kriwet, Jürgen
Seymour Island, Antarctic Peninsula, is known for its wealth of fossil remains. This island provides one of the richest fossiliferous Paleogene sequences in the world. Chondrichthyans seemingly dominate this Eocene marine fauna and offer a rare insight into high-latitude faunas during the Palaeogene. So far, only a few isolated teeth of carcharhinid sharks have been reported from Seymour Island. Bulk sampling in the well-exposed La Meseta and Submeseta formations yielded new and abundant chondrichthyan material, including numerous teeth of carcharhinid and triakid sharks. Here, we present a reevaluation of the previously described carcharhinid remains and a description of new taxa: Meridiogaleus cristatus, gen. et sp. nov., Kallodentis rythistemma, gen. et sp. nov., Abdounia richteri, sp. nov., and Abdounia mesetae, sp. nov. The carcharhiniforms Mustelus sp. and Galeorhinus sp. are reported based on rare material, whereas teeth previously assigned to Scoliodon represent a nomen dubium. PMID:29551850
Saber, Shaban G.; Salama, Yasser F.
Three Eocene stratigraphic successions east of the Beni Suef area are described and measured. These successions are made up of four rock units that are from base to top: Qarara (upper Lutetian), El Fashn (Bartonian), Beni Suef, and Maadi (Priabonian) formations. A detailed facies and sequence analysis unravels the stratigraphic framework and constructs a depositional model for the Middle-Upper Eocene succession. Ten microfacies types were grouped into four facies associations on a homoclinal ramp that compose the Upper Lutetian-Priabonian succession exposed in the east Beni Suef area. The depositional environment varied from a shallow to deep ramp setting. Four third-order depositional sequences were identified in the studied sections. The sequence boundaries are paleosoil horizons that can be traced throughout the entire outcrop area. Missing biozones are also evidence of the sequence boundaries. The history of these sequences mirrors the eustatic sea-level changes and the local tectonics in the region. Each sequence comprises facies associations that make up lowstand and/or transgressive and highstand systems tracts. The lowstand systems tract (LST) deposits are mainly sandstone facies and in Sequences 3 and 4 at Gabal Abyiad and Gabal Homret Shaibun respectively. The transgressive systems tract (TST) of Sequence 1 is dominated by nummulitic facies at Gabal Diya. The shale, mudstone and wackestone facies with planktic foraminifera and echinoids dominate the TST of Sequences 2 and 3 at Gabal Abyiad and Gabal Homret Shaibun respectively. The highstand systems tract (HST) of the studied sections is characterized by benthic foraminifera and bryozoan wackestone and packstone facies.
Full Text Available Radiometric and geochemical studies were carried out at Red Hill in the southern part of King George Island (South Shetland Islands, northern Antarctic Peninsula on the Bransfield Strait coast. The rock succession at Red Hill has been determined to represent the Baranowski Glacier Group that was previously assigned a Late Cretaceous age. Two formations were distinguished within this succession: the lower Llano Point Formation and the upper Zamek Formation. These formations have stratotypes defined further to the north on the western coast of Admiralty Bay. On Red Hill the Llano Point Formation consists of terrestrial lavas and pyroclastic breccia; the Zamek Formation consist predominantly of fine to coarse tuff, pyroclastic breccia, lavas, tuffaceous mud-, silt-, and sandstone, locally conglomeratic. The lower part of the Zamek Formation contains plant detritus (Nothofagus, dicotyledonous, thermophilous ferns and numerous coal seams (vitrinitic composition that confirm the abundance of vegetation on stratovolcanic slopes and surrounding lowlands at that time. Selected basic to intermediate igneous rocks from the succession have been analysed for the whole-rock K-Ar age determination. The obtained results indicate that the Red Hill succession was formed in two stages: (1 from about 51–50 Ma; and (2 46–42 Ma, i.e. during the Early to Middle Eocene. This, in combination with other data obtained from other Baranowski Glacier Group exposures on western coast of Admiralty Bay, confirms the recently defined position of the volcano-clastic succession in the stratigraphic scheme of King George Island. The new stratigraphic position and lithofacies development of the Red Hill succession strongly suggest its correlation with other Eocene formations containing fossil plants and coal seams that commonly occur on King George Island.
Keppie, J. D.; Moran-Zenteno, D. J.; Martiny, B. M.; Gonzalez-Torres, E.
Traditional Cenozoic reconstructions of southern Mexico that place the Chortis block (Honduras and northern Nicaragua) adjacent southwestern Mexico using the Motagua Fault Zone violate plate tectonic principles, and are inconsistent with the presence of undeformed upper Cretaceous - Recent sediments in the Gulf of Tehuantepec that straddle the westward projection of the Motagua Fault Zone, the inferred northern boundary of the Chortis block. Rotating the Chortis block about a rotation pole near Santiago (Chile) places it to the southwest of its present position at 45 Ma with the Farallon Plate lying off southwestern Mexico. Examination of the geological record of southwestern Mexico (102-96°W) records several synchronous events in the upper Oligocene- lower Miocene (29-19 Ma): (1) an hiatus in arc magmatism; (2) removal of a wide (~210 km) upper Eocene-lower Oligocene forearc; (3) exhumation of 13-20 km of upper Eocene-lower Oligocene arc along the present day coast; and (4) breakup of the Farallon Plate. We suggest that the magmatic hiatus was caused by subduction of the forearc, which replaced the mantle wedge by relatively cool crust and has since been deeply subducted. Potential causes for subduction of the forearc include collision of an oceanic plateau with the trench, and a change in plate kinematics synchronous with breakup of the Farallon Plate and initiation of the Guadalupe-Nazca spreading ridge. Following removal of the forearc, arc magmatism resumed in two segments offset by the Veracruz-Oaxaca Line: (i) the 19 Ma to Present E-W Trans-Mexican Volcanic Belt; and (ii) the 22-10 Ma WNW part of the Central American Volcanic Arc. Estimates of the dip of the Benioff zone based on Miocene arc/forearc widths indicate that it changed from 11 to 39 degrees (west to east) across the Veracruz-Oaxaca Line, which is interpreted as a transform fault with a ~100 km dextral offset.
Pagani, Mark; Pedentchouk, Nikolai; Huber, Matthew; Sluijs, Appy; Schouten, Stefan; Brinkhuis, Henk; Damsté, Jaap S Sinninghe; Dickens, Gerald R
The Palaeocene/Eocene thermal maximum represents a period of rapid, extreme global warming 55 million years ago, superimposed on an already warm world. This warming is associated with a severe shoaling of the ocean calcite compensation depth and a >2.5 per mil negative carbon isotope excursion in marine and soil carbonates. Together these observations indicate a massive release of 13C-depleted carbon and greenhouse-gas-induced warming. Recently, sediments were recovered from the central Arctic Ocean, providing the first opportunity to evaluate the environmental response at the North Pole at this time. Here we present stable hydrogen and carbon isotope measurements of terrestrial-plant- and aquatic-derived n-alkanes that record changes in hydrology, including surface water salinity and precipitation, and the global carbon cycle. Hydrogen isotope records are interpreted as documenting decreased rainout during moisture transport from lower latitudes and increased moisture delivery to the Arctic at the onset of the Palaeocene/Eocene thermal maximum, consistent with predictions of poleward storm track migrations during global warming. The terrestrial-plant carbon isotope excursion (about -4.5 to -6 per mil) is substantially larger than those of marine carbonates. Previously, this offset was explained by the physiological response of plants to increases in surface humidity. But this mechanism is not an effective explanation in this wet Arctic setting, leading us to hypothesize that the true magnitude of the excursion--and associated carbon input--was greater than originally surmised. Greater carbon release and strong hydrological cycle feedbacks may help explain the maintenance of this unprecedented warmth.
Full Text Available A brief (~150 kyr period of widespread global average surface warming marks the transition between the Paleocene and Eocene epochs, ~56 million years ago. This so-called "Paleocene-Eocene thermal maximum" (PETM is associated with the massive injection of 13C-depleted carbon, reflected in a negative carbon isotope excursion (CIE. Biotic responses include a global abundance peak (acme of the subtropical dinoflagellate Apectodinium. Here we identify the PETM in a marine sedimentary sequence deposited on the East Tasman Plateau at Ocean Drilling Program (ODP Site 1172 and show, based on the organic paleothermometer TEX86, that southwest Pacific sea surface temperatures increased from ~26 °C to ~33°C during the PETM. Such temperatures before, during and after the PETM are >10 °C warmer than predicted by paleoclimate model simulations for this latitude. In part, this discrepancy may be explained by potential seasonal biases in the TEX86 proxy in polar oceans. Additionally, the data suggest that not only Arctic, but also Antarctic temperatures may be underestimated in simulations of ancient greenhouse climates by current generation fully coupled climate models. An early influx of abundant Apectodinium confirms that environmental change preceded the CIE on a global scale. Organic dinoflagellate cyst assemblages suggest a local decrease in the amount of river run off reaching the core site during the PETM, possibly in concert with eustatic rise. Moreover, the assemblages suggest changes in seasonality of the regional hydrological system and storm activity. Finally, significant variation in dinoflagellate cyst assemblages during the PETM indicates that southwest Pacific climates varied significantly over time scales of 103 – 104 years during this event, a finding comparable to similar studies of PETM successions from the New Jersey Shelf.
Light, J. J.; Passchier, S.
The Eocene Oligocene Transition (EOT) 34 million years ago (Ma), marked the global climate change from greenhouse to icehouse, and the full establishment of the East Antarctic Ice Sheet (EAIS). The initiation of the EAIS during the EOT is believed to have been a step-wise transition; however, data resolution is low and merits the need for further study. The purpose of this study is to expand upon existing knowledge of EAIS dynamics spanning the EOT by creating a higher resolution geochemical record of cores taken from continental shelf sites 1166 in Prydz Bay and U1360 from the Wilkes Land margin. We used Inductively Coupled Plasma Optical Emission Spectrometry and Mass Spectrometry (ICP-OES/ ICP-MS) to determine the bulk chemical composition of samples. Results were used to calculate the Chemical Index of Alteration (CIA), Al2O3/TiO2 ratios, and trace elemental variation down core. CIA values for the early Oligocene in Site U1360 indicate an arid colder environment less likely to be chemically weathered. In contrast, Hole 1166A shows values similar to average shales that increase up core and abruptly decrease at the overlying Neogene diamict, suggesting a warmer more humid environment at Prydz Bay during the late Eocene. Al2O3/TiO2 ratios were used to evaluate mud provenance changes at each site. At site 1166 redox sensitive elements (Cr, Ni, and V) show similar down core distributions to one another. The changes in elemental intensities are likely being controlled by factors such as sediment provenance, changes in redox conditions and surficial weathering. We expect the outcomes of this study to allow us to interpret regional depositional environments at a higher resolution, as well as to shed light on the EAIS's step-wise initiation.
Grímsson, FriĐgeir; Zetter, Reinhard; Labandeira, Conrad C.; Engel, Michael S.; Wappler, Torsten
Abstract The middle Eocene Messel and Eckfeld localities are renowned for their excellently preserved faunas and diverse floras. Here we describe for the first time pollen from insect-pollinated plants found in situ on well-preserved ancient bees using light and scanning electron microscopy. There have been 140 pollen types reported from Messel and 162 pollen types from Eckfeld. Here we document 23 pollen types, six from Messel and 18 from Eckfeld (one is shared). The taxa reported here are all pollinated by insects and mostly not recovered in the previously studied dispersed fossil pollen records. Typically, a single or two pollen types are found on each fossil bee specimen, the maximum number of distinct pollen types on a single individual is five. Only five of the 23 pollen types obtained are angiosperms of unknown affinity, the remainder cover a broad taxonomic range of angiosperm trees and include members of several major clades: monocots (1 pollen type), fabids (7), malvids (4), asterids (5) and other core eudicots (1). Seven types each can be assigned to individual genera or infrafamilial clades. Since bees visit only flowers in the relative vicinity of their habitat, the recovered pollen provides a unique insight into the autochthonous palaeo-flora. The coexistence of taxa such as Decodon, Elaeocarpus, Mortoniodendron and other Tilioideae, Mastixoideae, Olax, Pouteria and Nyssa confirms current views that diverse, thermophilic forests thrived at the Messel and Eckfeld localities, probably under a warm subtropical, fully humid climate. Our study calls for increased attention to pollen found in situ on pollen-harvesting insects such as bees, which can provide new insights on insect-pollinated plants and complement even detailed palaeo-palynological knowledge obtained mostly from pollen of wind-pollinated plants in the dispersed pollen record of sediments. In the case of Elaeocarpus, Mortoniodendron, Olax and Pouteria the pollen collected by the middle Eocene
Cleary, Gráinne P; Parsons, Holly; Davis, Adrian; Coleman, Bill R; Jones, Darryl N; Miller, Kelly K; Weston, Michael A
Private gardens provide habitat and resources for many birds living in human-dominated landscapes. While wild bird feeding is recognised as one of the most popular forms of human-wildlife interaction, almost nothing is known about the use of bird baths. This citizen science initiative explores avian assemblages at bird baths in private gardens in south-eastern Australia and how this differs with respect to levels of urbanisation and bioregion. Overall, 992 citizen scientists collected data over two, four-week survey periods during winter 2014 and summer 2015 (43% participated in both years). Avian assemblages at urban and rural bird baths differed between bioregions with aggressive nectar-eating species influenced the avian assemblages visiting urban bird baths in South Eastern Queensland, NSW North Coast and Sydney Basin while introduced birds contributed to differences in South Western Slopes, Southern Volcanic Plains and Victorian Midlands. Small honeyeaters and other small native birds occurred less often at urban bird baths compared to rural bird baths. Our results suggest that differences between urban versus rural areas, as well as bioregion, significantly influence the composition of avian assemblages visiting bird baths in private gardens. We also demonstrate that citizen science monitoring of fixed survey sites such as bird baths is a useful tool in understanding large-scale patterns in avian assemblages which requires a vast amount of data to be collected across broad areas.
...-0014; 91200-1231-9BPP-L2] RIN 1018-AX34 Migratory Bird Hunting; Supplemental Proposals for Migratory Game Bird Hunting Regulations for the 2011-12 Hunting Season; Notice of Meetings AGENCY: Fish and... migratory game birds for the 2011-12 hunting season. This supplement to the proposed rule provides the...
...-0060; FF09M21200-134-XMB123199BPP0] RIN 1018-AX90 Migratory Bird Permits; Definition of ``Hybrid'' Migratory Bird AGENCY: Fish and Wildlife Service, Interior. ACTION: Final rule. SUMMARY: We, the U.S. Fish and Wildlife Service (FWS), revise the definition of ``hybrid'' as it relates to birds protected under...
...-91200 FF09M26000] Proposed Information Collection; Migratory Bird Harvest Information Program and Migratory Bird Surveys AGENCY: Fish and Wildlife Service, Interior. ACTION: Notice; request for comments...) or 703- 358-2482 (telephone). SUPPLEMENTARY INFORMATION: I. Abstract The Migratory Bird Treaty Act...
... Part III Department of the Interior Fish and Wildlife Service 50 CFR Part 20 Migratory Bird Hunting; Proposed 2010-11 Migratory Game Bird Hunting Regulations (Preliminary) With Requests for Indian Tribal Proposals and Requests for 2011 Spring and Summer Migratory Bird Subsistence Harvest Proposals in...
...-0082; 91200-1231-9BPP-L2] RIN 1018-AW67 Migratory Bird Subsistence Harvest in Alaska; Harvest Regulations for Migratory Birds in Alaska During the 2010 Season AGENCY: Fish and Wildlife Service, Interior... Service, are reopening the public comment period on our proposed rule to establish migratory bird...
Cleary, Gráinne P.; Parsons, Holly; Davis, Adrian; Coleman, Bill R.; Jones, Darryl N.; Miller, Kelly K.; Weston, Michael A.
Private gardens provide habitat and resources for many birds living in human-dominated landscapes. While wild bird feeding is recognised as one of the most popular forms of human-wildlife interaction, almost nothing is known about the use of bird baths. This citizen science initiative explores avian assemblages at bird baths in private gardens in south-eastern Australia and how this differs with respect to levels of urbanisation and bioregion. Overall, 992 citizen scientists collected data over two, four-week survey periods during winter 2014 and summer 2015 (43% participated in both years). Avian assemblages at urban and rural bird baths differed between bioregions with aggressive nectar-eating species influenced the avian assemblages visiting urban bird baths in South Eastern Queensland, NSW North Coast and Sydney Basin while introduced birds contributed to differences in South Western Slopes, Southern Volcanic Plains and Victorian Midlands. Small honeyeaters and other small native birds occurred less often at urban bird baths compared to rural bird baths. Our results suggest that differences between urban versus rural areas, as well as bioregion, significantly influence the composition of avian assemblages visiting bird baths in private gardens. We also demonstrate that citizen science monitoring of fixed survey sites such as bird baths is a useful tool in understanding large-scale patterns in avian assemblages which requires a vast amount of data to be collected across broad areas. PMID:26962857
Schumann, D.; Raub, T. D.; Kopp, R. E.; Guerquin-Kern, J. L.; Wu, T. D.; Rouiller, I.; Smirnov, A. V.; Sears, S. K.; Lücken, U.; Tikoo, S. M.; Hesse, R.; Kirschvink, J. L.; Vali, H.
The Paleocene-Eocene Thermal Maximum (PETM) is one of the most severe climatic events of the Cenozoic Era. A massive injection of light carbon into the oceans and atmosphere over a few thousand of years triggered drastic perturbation of Earth's climate resulting in abrupt global warming of ~5-9oC [Sluijs et al., 2007] that persisted for ~180,000 years. This episode is marked by the diversification and radiation of terrestrial plants and mammals while in the marine realm numerous deep-sea benthic foraminifera species disappeared and new forms evolved. Sediments deposited during the PETM are clay-rich and contain distinct evidence of these climatic changes. Kopp et al., (2007) and Lippert & Zachos (2007) report an extraordinary magnetofossil ‘Lagerstätte' in lowermost Eocene kaolinite-rich clay sediments deposited at subtropical paleolatitude in the Atlantic Coastal Plain of New Jersey, USA. Magnetofossils are magnetic particles produced most abundantly by magnetotactic bacteria. Kopp et al. (2007) and Lippert & Zachos (2007) used ferromagnetic resonance (FMR) spectroscopy, other rock magnetic methods, and transmission electron microscopy (TEM) of magnetic separates to characterize sediments from boreholes at Ancora (ODP Leg 174AX) and Wilson Lake, NJ, respectively. These sediments contain abundant ~40- to 300-nm cuboidal, elongate-prismatic and bullet-shaped magnetofossils, sometimes arranged in short chains, resembling crystals in living magnetotactic bacteria. Despite the scarcity of intact magnetofossil chains, the asymmetry ratios of the FMR spectra reflects a profusion of elongate single domain (SD) crystals and/or chains. Here we address both conundrums by reporting the discovery from these same sediments of exceptionally large and novel biogenic magnetite crystals unlike any previously reported from living organisms or from sediments. Aside from abundant bacterial magnetofossils, electron microscopy reveals novel spearhead-like and spindle-like magnetite
Full Text Available To answer the question of whether sensory pleasure exists in birds, I trained an African-gray parrot (Psittacus erythacus named Aristote to speak. Stage 1 of the study consisted in gaining Aristote's affection. In Stage 2 Aristote was taught to speak, following Irene Pepperberg's triangular method: another person and I would talk together and look at Aristote only when it used understandable French words. Thus Aristote learned to say a few words for obtaining toys or getting my attention; e.g. “donne bouchon” (give cork or “donne gratte” (give scratch/tickle, with the appropriate reward. In Stage 3, the word bon (good was added to the short list of words used by Aristote. I said “bon” when giving Aristote the stimuli it requested and which would, presumably, be pleasurable; e.g. gratte bon. Aristote started to use short sentences such as “yaourt bon” (good yogurt. Eventually, Aristote transferred the word bon to new stimuli such as raisin (grape, an association I myself had never made. Such a use of vocabulary, and moreover its transfer, likely shows that this bird experienced sensory pleasure.
Lahner, Lesanna L.; Franson, J. Christian
Lead in its various forms has been used for thousands of years, originally in cooking utensils and glazes and more recently in many industrial and commercial applications. However, lead is a potent, potentially deadly toxin that damages many organs in the body and can affect all animals, including humans. By the mid 1990s, lead had been removed from many products in the United States, such as paint and fuel, but it is still commonly used in ammunition for hunting upland game birds, small mammals, and large game animals, as well as in fishing tackle. Wild birds, such as mourning doves, bald eagles, California condors, and loons, can die from the ingestion of one lead shot, bullet fragment, or sinker. According to a recent study on loon mortality, nearly half of adult loons found sick or dead during the breeding season in New England were diagnosed with confirmed or suspected lead poisoning from ingestion of lead fishing weights. Recent regulations in some states have restricted the use of lead ammunition on certain upland game hunting areas, as well as lead fishing tackle in areas frequented by common loons and trumpeter swans. A variety of alternatives to lead are available for use in hunting, shooting sports, and fishing activities.
Tsai, S S; Park, J H; Hirai, K; Itakura, C
Amongst eye lesions in birds that died in quarantine, cataracts were the most common disorders (37/241, 15.4%), being prevalent in the annular pads of cockatiels (Nymphicus hollandicus), Amazon parrots (Amazona aestiva aestiva) and budgerigars (Melopsittacus undulatus). The incidence in male birds was more than twice that in females. Deposition of crystals, mostly in the cornea, was the second most frequent lesion (21/293, 8.7%), mainly found in cockatiels, parakeets (Psittacula krameri manillensis), Amazon parrots (Amazona aestiva aestiva), budgerigars and finches (Poephila gouldiae gouldiae). These corneal crystals were negative to PAS and Kossa's stains. Six parakeets (Psittacula krameri manillensis) had calcium salts deposited in the inner plexiform layer of the retina and occasionally in the iris and ciliary body. Neither inflammation nor neo-vascularization was observed when cataracts, corneal crystalline deposition, and retinal and ciliary calcification were present. Intranuclear inclusion bodies typical for papovavirus infection were found in the eyelids of six budgerigars (2.5%). Similar inclusions were simultaneously found in the pars ciliaris retinae (4, 1.7%), inner plexiform of retina (1, 0.4%) and anterior epithelium of the cornea (1, 0.4%). Other lesions such as candidial endophthalmitis, conjunctival cryptosporidiosis, corneal dystrophy, keratitis, corneal perforation and iridocyclitis, were occasional findings.
Mora, Thierry; Walczak, Aleksandra M.; Del Castello, Lorenzo; Ginelli, Francesco; Melillo, Stefania; Parisi, Leonardo; Viale, Massimiliano; Cavagna, Andrea; Giardina, Irene
The correlated motion of flocks is an example of global order emerging from local interactions. An essential difference with respect to analogous ferromagnetic systems is that flocks are active: animals move relative to each other, dynamically rearranging their interaction network. This non-equilibrium characteristic has been studied theoretically, but its impact on actual animal groups remains to be fully explored experimentally. Here, we introduce a novel dynamical inference technique, based on the principle of maximum entropy, which accommodates network rearrangements and overcomes the problem of slow experimental sampling rates. We use this method to infer the strength and range of alignment forces from data of starling flocks. We find that local bird alignment occurs on a much faster timescale than neighbour rearrangement. Accordingly, equilibrium inference, which assumes a fixed interaction network, gives results consistent with dynamical inference. We conclude that bird orientations are in a state of local quasi-equilibrium over the interaction length scale, providing firm ground for the applicability of statistical physics in certain active systems.
Wilcoxen, Travis E; Horn, David J; Hogan, Brianna M; Hubble, Cody N; Huber, Sarah J; Flamm, Joseph; Knott, Madeline; Lundstrom, Lisa; Salik, Faaria; Wassenhove, Samantha J; Wrobel, Elizabeth R
Among the most popular reasons that people feed wild birds is that they want to help birds. The extent to which supplemental food helps birds, however, is not well established. From spring 2011 to spring 2014, we examined how feeding of wild birds influences the health of individual birds at forested sites in central Illinois, USA. Specifically, we compared three forested sites where we provided supplemental food with three forested sites for which no supplemental food was available and monitored changes in the individual health of birds. In addition, we determined whether any changes in bird health had occurred after feeders had been removed from sites 10 months before. Generally, the individual health of birds improved with supplemental feeding, including increased antioxidant levels, reduced stress (heterophil-to-lymphocyte ratio) and more rapid feather growth. In some species, we also found improved body condition index scores and innate immune defense. The difference among sites was not present 10 months after feeders were removed, suggesting that the impact on health was indeed related to supplemental feeding. Potential negative effects of supplemental feeding were also found, including an increase in infectious disease prevalence among individual birds at forested sites where supplemental food was offered. Birds with clear signs of pathology showed deficits in most of the physiological metrics in which birds at feeder sites typically showed improved health condition. At the peak of prevalence of infectious disease, 8.3% of all birds at feeders exhibited symptoms of conjunctivitis, pox, dermal disease or cloacal disease. We found both positive and negative impacts of wild bird feeding, and that, in general, birds that had access to supplemental food were in better physiological condition. Moreover, the negative effects we found may be mitigated by hobbyists engaging in safer bird-feeding practices.
Sieges, Mason L.; Smolinsky, Jaclyn A.; Baldwin, Michael J.; Barrow, Wylie C.; Randall, Lori A.; Buler, Jeffrey J.
In response to the Deepwater Horizon oil spill in spring 2010, the Natural Resources Conservation Service implemented the Migratory Bird Habitat Initiative (MBHI) to provide temporary wetland habitat for migrating and wintering waterfowl, shorebirds, and other birds along the northern Gulf of Mexico via managed flooding of agricultural lands. We used weather-surveillance radar to conduct broad regional assessments of bird response to MBHI activities within the Mississippi Alluvial Valley and the West Gulf Coastal Plain. Across both regions, birds responded positively to MBHI management by exhibiting greater relative bird densities within sites relative to pre-management conditions in prior years and relative to surrounding non-flooded agricultural lands. Bird density at MBHI sites was generally greatest during winter for both regions. Unusually high flooding in the years prior to implementation of the MBHI confounded detection of overall changes in remotely sensed soil wetness across sites. The magnitude of bird response at MBHI sites compared to prior years and to non-flooded agricultural lands was generally related to the surrounding landscape context: proximity to areas of high bird density, amount of forested wetlands, emergent marsh, non-flooded agriculture, or permanent open water. However, these relationships varied in strength and direction between regions and seasons, a finding which we attribute to differences in seasonal bird composition and broad regional differences in landscape configuration and composition. We detected greater increases in relative bird use at sites in closer proximity to areas of high bird density during winter in both regions. Additionally, bird density was greater during winter at sites with more emergent marsh in the surrounding landscape. Thus, bird use of managed wetlands could be maximized by enrolling lands located near areas of known bird concentration and within a mosaic of existing wetlands. Weather-radar observations
Shamoun-Baranes, J.; Bouten, W.; Sierdsema, H.; van Belle, J.; van Gasteren, J.R.; van Loon, E.E.
The NL-BAM was developed as a web-based decision support tool to be used by the bird hazard avoidance experts in the ecology unit of the Royal Netherlands Air Force. The NL-BAM will be used together with the ROBIN 4 radar system to provide BirdTAMS, for real time warnings and flight planning and to
Gwinner, E. G.
Several species of warbler birds were observed in an effort to determine what initiates and terminates migration. Environmental and endogenous timing mechanisms were analyzed. The results indicate that endogenous stimuli are dominant factors for bird migration especially for long distances. It was concluded that environmental factors act as an assist mechanism.
Xu, Xing; Zhou, Zhonghe; Dudley, Robert; Mackem, Susan; Chuong, Cheng-Ming; Erickson, Gregory M; Varricchio, David J
Recent discoveries of spectacular dinosaur fossils overwhelmingly support the hypothesis that birds are descended from maniraptoran theropod dinosaurs, and furthermore, demonstrate that distinctive bird characteristics such as feathers, flight, endothermic physiology, unique strategies for reproduction and growth, and a novel pulmonary system originated among Mesozoic terrestrial dinosaurs. The transition from ground-living to flight-capable theropod dinosaurs now probably represents one of the best-documented major evolutionary transitions in life history. Recent studies in developmental biology and other disciplines provide additional insights into how bird characteristics originated and evolved. The iconic features of extant birds for the most part evolved in a gradual and stepwise fashion throughout archosaur evolution. However, new data also highlight occasional bursts of morphological novelty at certain stages particularly close to the origin of birds and an unavoidable complex, mosaic evolutionary distribution of major bird characteristics on the theropod tree. Research into bird origins provides a premier example of how paleontological and neontological data can interact to reveal the complexity of major innovations, to answer key evolutionary questions, and to lead to new research directions. A better understanding of bird origins requires multifaceted and integrative approaches, yet fossils necessarily provide the final test of any evolutionary model. Copyright © 2014, American Association for the Advancement of Science.
Caro, S.P.; Balthazart, J.
Birds are anosmic or at best microsmatic… This misbelief persisted until very recently and has strongly influenced the outcome of communication studies in birds, with olfaction remaining neglected as compared to acoustic and visual channels. However, there is now clear empirical evidence showing
The paper summarizes the current understanding of the evolution and diversification of birds. New insights into this field have mainly come from two fundamentally different, but complementary sources of information: the many newly discovered Mesozoic bird fossils and the wealth of genetic analyses
James G. Dickson
Fire has long been a natural and anthropogenic force shaping southern forests and their fauna. Some species are attracted to recent burns. There is little direct mortality of adult birds by fire, but growing season fires may consume some nests. Fire affects bird communities mainly through effects on vegetation. Fires effective enough to limit understory hardwood...
Jun 1, 2014 ... Keywords: Bird Louse, Kano Sub-urb areas, Menacanthus cornutus, Poultry, Prevalence ... including retarded growth, low egg production and ..... Bush, S. E. and Clayton, D. H.. (2005). Correlated evolution of host and parasite body size: Tests of Harrison's rule using birds and lice. Evolution, 59: 1744-.
Study on the prevalence of bird lice in four selected farms in Kano metropolis was conducted to determine the lice species richness, lice abundance and percent prevalence in the four poultry farms. Two hundred and forty (240) birds were examined from four poultry farms within Kano in Tofa, Fagge, Brigade and Gwarzo ...
Prokop, Pavol; Rodak, Rastislav
A pupil's ability to identify common organisms is necessary for acquiring further knowledge of biology. We investigated how pupils were able to identify 25 bird species following their song, growth habits, or both features presented simultaneously. Just about 19% of birds were successfully identified by song, about 39% by growth habit, and 45% of…
Justine A. Kummer
Full Text Available Feeding wild birds creates an important link between homeowners and conservation. The effects of bird feeders and year-round feeding on birds have not been well studied, however, particularly in relationship to bird-window collisions. We determined effects of bird feeder presence and placement on bird-window collisions at residential homes. Paired month-long trials in which a feeder was either present or absent for one month and then removed or added for the second month were completed at 55 windows at 43 houses. In each trial, homeowners were asked to search their study window daily for evidence of a bird-window collision. During the study there were 51 collisions when there was no bird feeder and 94 when the feeder was present. The season when each trial was set up was the best individual predictor of bird-window collisions. The largest number of collisions was observed during fall migration and the lowest during the winter months. There were no collisions at 26 of the study windows. High variance was observed in the number of collisions at different houses, indicating that effects of bird feeders are context dependent. Changing the occurrence, timing, and placement of feeders can alter collision rates but is only one of many factors that influence whether a residential house is likely to have a bird window-collision or not.
Brusatte, Stephen L; O'Connor, Jingmai K; Jarvis, Erich D
Birds are one of the most recognizable and diverse groups of modern vertebrates. Over the past two decades, a wealth of new fossil discoveries and phylogenetic and macroevolutionary studies has transformed our understanding of how birds originated and became so successful. Birds evolved from theropod dinosaurs during the Jurassic (around 165-150 million years ago) and their classic small, lightweight, feathered, and winged body plan was pieced together gradually over tens of millions of years of evolution rather than in one burst of innovation. Early birds diversified throughout the Jurassic and Cretaceous, becoming capable fliers with supercharged growth rates, but were decimated at the end-Cretaceous extinction alongside their close dinosaurian relatives. After the mass extinction, modern birds (members of the avian crown group) explosively diversified, culminating in more than 10,000 species distributed worldwide today. Copyright © 2015 Elsevier Ltd. All rights reserved.
Giammarino, M; Vaschetti, G; Boano, G
This study aims at evaluating the prevalence and density of haemoparasites in a native population of immature and adult male and female birds of Burkina Faso. Protists from the orders Eucoccidiorida, Kinetoplastida, as well as microfilaria were found. During the period from the 27th to the 29th of November 2003, we trapped 110 birds: on 101 sampled birds, 47 harboured blood parasites, with an overall prevalence of infection of 46.5%; 15 birds (14.8%) harboured mixed infections. Haemoproteus (prevalence of infection 30.7%), Plasmodium (13.9%), Trypanosoma (10.9%), microfilaria (8.9%) and Leucocytozoon (5.9%) were most frequently recorded. Occurrence of blood parasites was very high in comparison to that seen in birds from other areas of Africa.
Alerstam, Thomas; Bäckman, Johan; Gudmundsson, Gudmundur A; Hedenström, Anders; Henningsson, Sara S; Karlsson, Håkan; Rosén, Mikael; Strandberg, Roine
Studies of bird migration in the Beringia region of Alaska and eastern Siberia are of special interest for revealing the importance of bird migration between Eurasia and North America, for evaluating orientation principles used by the birds at polar latitudes and for understanding the evolutionary implications of intercontinental migratory connectivity among birds as well as their parasites. We used tracking radar placed onboard the ice-breaker Oden to register bird migratory flights from 30 July to 19 August 2005 and we encountered extensive bird migration in the whole Beringia range from latitude 64 degrees N in Bering Strait up to latitude 75 degrees N far north of Wrangel Island, with eastward flights making up 79% of all track directions. The results from Beringia were used in combination with radar studies from the Arctic Ocean north of Siberia and in the Beaufort Sea to make a reconstruction of a major Siberian-American bird migration system in a wide Arctic sector between longitudes 110 degrees E and 130 degrees W, spanning one-third of the entire circumpolar circle. This system was estimated to involve more than 2 million birds, mainly shorebirds, terns and skuas, flying across the Arctic Ocean at mean altitudes exceeding 1 km (maximum altitudes 3-5 km). Great circle orientation provided a significantly better fit with observed flight directions at 20 different sites and areas than constant geographical compass orientation. The long flights over the sea spanned 40-80 degrees of longitude, corresponding to distances and durations of 1400-2600 km and 26-48 hours, respectively. The birds continued from this eastward migration system over the Arctic Ocean into several different flyway systems at the American continents and the Pacific Ocean. Minimization of distances between tundra breeding sectors and northerly stopover sites, in combination with the Beringia glacial refugium and colonization history, seemed to be important for the evolution of this major
van der Boon, A.; Beniest, A.; Ciurej, A.; Gaździcka, E.; Grothe, A.; Sachsenhofer, R. F.; Langereis, C. G.; Krijgsman, W.
During the Eocene-Oligocene transition (EOT), a major palaeoenvironmental change took place in the Paratethys Sea of central Eurasia. Restricted connectivity and increased stratification resulted in wide-spread deposition of organic-rich sediments which nowadays make up important hydrocarbon source rocks. The North Alpine Foreland Basin (NAFB) was a major gateway of the Paratethys Sea to the open ocean during the Eocene, but the age of closure of this gateway is still uncertain. The Ammer section in southern Germany documents the shallowing of this connection and subsequent disappearance of marine environments in the NAFB, as reflected in its sedimentary succession of turbidites to marls (Deutenhausen to Tonmergel beds), via coastal sediments (Baustein beds) to continental conglomerates (Weißach beds). Here, we apply organic geochemistry and date the lithological transitions in the Ammer section using integrated stratigraphy, including magnetostratigraphy and biostratigraphy. Nannoplankton and dinocyst results can be reconciled when dinoflagellate species Wetzeliella symmetrica is of late Eocene age. Our magnetostratigraphy then records C13r-C13n-C12r and allows calculation of sediment accumulation rates and estimation of ages of lithological transitions. We show that the shallowing from turbiditic slope deposits (Deutenhausen beds) to shelf sediments (Tonmergel beds) coincides with the Eocene-Oligocene boundary at 33.9 Ma. The transition to continental sediments is dated at ca. 33.15 Ma, significantly older than suggested by previous studies. We conclude that the transition from marine to continental sediments drastically reduced the marine connection through the western part of the NAFB and influenced the oxygen conditions of the Paratethys Sea.
Nielsen, Jan Kresten; Milàn, Jesper; Mesfun, Daniel
A cliff exposure of the Eocene Lillebælt Clay Formation, on the Røsnæs peninsula of Zealand, Denmark, has yielded a diverse trace-fossil assemblage. The trace fossils are described formally for the first time and assigned to Phymatoderma melvillensis, unnamed clusters of small burrows, Ophiomorpha nodosa, Spongeliomorpha isp., Dreginozoum beckumensis, Bichordites isp., Chondrites isp., Atollites zitteli? and ?Rhizocorallium isp. The preservation of the trace fossils is strongly related to ear...
Gaillard, Christian; Olivero, Davide; Chebance, Marie
International audience; Enigmatic biogenic structures are reported from the Palaeogene of south-eastern France in the Castellane-Blieux area (Alpes de Haute-Provence French Department). They occur at the Eocene/Oligocene transition at the top of the Marnes Bleues Formation, which corresponds to a clearly defined regressive sequence, where thick marls are overlain by lacustrine limestones or sandstones. The marls have abundant shallow marine fauna and the upper part of the Formation correspond...
Vahlenkamp, Maximilian; Niezgodzki, Igor; De Vleeschouwer, David; Bickert, Torsten; Harper, Dustin; Kirtland Turner, Sandra; Lohmann, Gerrit; Sexton, Philip; Zachos, James; Pälike, Heiko
North Atlantic Deep Water (NADW) currently redistributes heat and salt between Earth's ocean basins, and plays a vital role in the ocean-atmosphere CO2 exchange. Despite its crucial role in today's climate system, vigorous debate remains as to when deep-water formation in the North Atlantic started. Here, we present datasets from carbonate-rich middle Eocene sediments from the Newfoundland Ridge, revealing a unique archive of paleoceanographic change from the progressively cooling climate of the middle Eocene. Well-defined lithologic alternations between calcareous ooze and clay-rich intervals occur at the ∼41-kyr beat of axial obliquity. Hence, we identify obliquity as the driver of middle Eocene (43.5-46 Ma) Northern Component Water (NCW, the predecessor of modern NADW) variability. High-resolution benthic foraminiferal δ18O and δ13C suggest that obliquity minima correspond to cold, nutrient-depleted, western North Atlantic deep waters. We thus link stronger NCW formation with obliquity minima. In contrast, during obliquity maxima, Deep Western Boundary Currents were weaker and warmer, while abyssal nutrients were more abundant. These aspects reflect a more sluggish NCW formation. This obliquity-paced paleoceanographic regime is in excellent agreement with results from an Earth system model, in which obliquity minima configurations enhance NCW formation.
Zeichner, S. S.; Kim, S.; Colman, A. S.
Early-Mid Eocene (56.0-33.9Mya) is characterized by a temperate Antarctic climate and shallower latitudinal temperature gradients than those in present day. The warmer waters off the coast of the Antarctic Peninsula provided suitable habitats for taxa (i.e., sharks) that live today at lower latitudes. Stable isotope analysis of Eocene shark teeth provides a proxy to understand high latitude temperature gradients. However, shark ecology, in particular migration and occupation of tidal versus pelagic habitats, must be considered in the interpretation of stable isotope data. In this study, we analyze d18OPO4 values from the enameloid of Striatolamia (synonymized with Carcharias) shark teeth from the La Meseta formation (Seymour Island, Antarctica) to estimate paleotemperature in Early-Mid Eocene Antarctica, and assess the impact of ecology versus environmental signals on d18OPO4 values. We compare the ranges and offsets between our measured shark tooth d18OPO4 and published bivalve d18OCO3 values to test whether shark teeth record signals of migration across latitudinal temperature gradients, or instead reflect seasonal and long-term temporal variation across La Meseta stratigraphic units.
Rose, Kenneth D; Chester, Stephen G B; Dunn, Rachel H; Boyer, Doug M; Bloch, Jonathan I
More than 25 new specimens of Teilhardina brandti, one of the oldest known euprimates, are reported from earliest Eocene strata of the southern Bighorn Basin, Wyoming. The new fossils include the first upper dentitions, a dentary showing the lower dental formula for the first time, and the first postcrania ascribed to T. brandti (tarsals and terminal phalanges). The elongated navicular and long talar neck suggest that T. brandti was an active arboreal quadruped, and the terminal phalanges constitute the oldest evidence for nails in Euprimates. Phylogenetic analysis incorporating the new data indicates that T. brandti is more derived than T. belgica but less so than T. americana. The hypothesis that Teilhardina originated in Asia (T. asiatica) and dispersed westward to Europe (T. belgica) and then to North America (T. brandti and T. magnoliana) during the earliest Eocene Paleocene-Eocene Thermal Maximum is most consistent with available evidence, including the relative age of fossil samples and their stage of evolution. Copyright © 2011 Wiley-Liss, Inc.
Lambert, Olivier; Martínez-Cáceres, Manuel; Bianucci, Giovanni; Di Celma, Claudio; Salas-Gismondi, Rodolfo; Steurbaut, Etienne; Urbina, Mario; de Muizon, Christian
Although combined molecular and morphological analyses point to a late middle Eocene (38-39 million years ago) origin for the clade Neoceti (Odontoceti, echolocating toothed whales plus Mysticeti, baleen whales, and relatives), the oldest known mysticete fossil dates from the latest Eocene (about 34 million years ago) of Antarctica [1, 2]. Considering that the latter is not the most stemward mysticete in recent phylogenies and that Oligocene toothed mysticetes display a broad morphological disparity most likely corresponding to contrasted ecological niches, the origin of mysticetes from a basilosaurid ancestor and its drivers are currently poorly understood [1, 3-8]. Based on an articulated cetacean skeleton from the early late Eocene (Priabonian, around 36.4 million years ago) of the Pisco Basin, Peru, we describe a new archaic tooth-bearing mysticete, Mystacodon selenensis gen. et sp. nov. Being the geologically oldest neocete (crown group cetacean) and the earliest mysticete to branch off described so far, the new taxon is interpreted as morphologically intermediate between basilosaurids and later toothed mysticetes, providing thus crucial information about the anatomy of the skull, forelimb, and innominate at these critical initial stages of mysticete evolution. Major changes in the morphology of the oral apparatus (including tooth wear) and flipper compared to basilosaurids suggest that suction and possibly benthic feeding represented key, early ecological traits accompanying the emergence of modern filter-feeding baleen whales' ancestors. Copyright © 2017 Elsevier Ltd. All rights reserved.
Viana, Duarte S; Gangoso, Laura; Bouten, Willem; Figuerola, Jordi
Long-distance dispersal (LDD) promotes the colonization of isolated and remote habitats, and thus it has been proposed as a mechanism for explaining the distributions of many species. Birds are key LDD vectors for many sessile organisms such as plants, yet LDD beyond local and regional scales has never been directly observed nor quantified. By sampling birds caught while in migratory flight by GPS-tracked wild falcons, we show that migratory birds transport seeds over hundreds of kilometres and mediate dispersal from mainland to oceanic islands. Up to 1.2% of birds that reached a small island of the Canary Archipelago (Alegranza) during their migration from Europe to Sub-Saharan Africa carried seeds in their guts. The billions of birds making seasonal migrations each year may then transport millions of seeds. None of the plant species transported by the birds occurs in Alegranza and most do not occur on nearby Canary Islands, providing a direct example of the importance of environmental filters in hampering successful colonization by immigrant species. The constant propagule pressure generated by these LDD events might, nevertheless, explain the colonization of some islands. Hence, migratory birds can mediate rapid range expansion or shifts of many plant taxa and determine their distribution. © 2016 The Author(s).
Blus, L.J.; Wiemeyer, Stanley N.; Kerwin, J.A.; Stendell, R.C.; Ohlendorf, H.M.; Stickel, L.F.
Pollution of estuaries affects bird populations indirectly through changes in habitat and food supply. The multi-factor pollution of Chesapeake Bay has resulted in diminution of submerged aquatic plants and consequent change in food habits of the canvasback duck. Although dredge-spoil operations can improve wildlife habitat, they often result in its demise. Pollution of estuaries also affects birds directly, through chemical toxication, which may result in outright mortality or in reproductive impairment. Lead from industrial sources and roadways enters the estuaries and is accumulated in tissues of birds. Lead pellets deposited in estuaries as a result of hunting are consumed by ducks with sufficient frequency .to result m large annual die-offs from lead poisoning. Fish in certain areas, usually near industrial sources, may contain levels of mercury high enough to be hazardous to birds that consume them. Other heavy metals are present in estuarine birds, but their significance is poorly known. Oil exerts lethal or sublethal effects on birds by oiling their feathers, oiling eggs and young by contaminated parents, and by ingestion of oil-contaminated food. Organochlorine chemicals, of both agricultural and industrial origin, travel through the food chains and reach harmful levels in susceptible species of birds in certain estuarine ecosystems. Both outright mortality and reproductive impairment have occurred.
Full Text Available The Early Eocene Thermal Maximum 2 (ETM2 at ~53.7 Ma is one of multiple hyperthermal events that followed the Paleocene-Eocene Thermal Maximum (PETM, ~56 Ma. The negative carbon excursion and deep ocean carbonate dissolution which occurred during the event imply that a substantial amount (103 Gt of carbon (C was added to the ocean-atmosphere system, consequently increasing atmospheric CO2(pCO2. This makes the event relevant to the current scenario of anthropogenic CO2 additions and global change. Resulting changes in ocean stratification and pH, as well as changes in exogenic cycles which supply nutrients to the ocean, may have affected the productivity of marine phytoplankton, especially calcifying phytoplankton. Changes in productivity, in turn, may affect the rate of sequestration of excess CO2 in the deep ocean and sediments. In order to reconstruct the productivity response by calcareous nannoplankton to ETM2 in the South Atlantic (Site 1265 and North Pacific (Site 1209, we employ the coccolith Sr/Ca productivity proxy with analysis of well-preserved picked monogeneric populations by ion probe supplemented by analysis of various size fractions of nannofossil sediments by ICP-AES. The former technique of measuring Sr/Ca in selected nannofossil populations using the ion probe circumvents possible contamination with secondary calcite. Avoiding such contamination is important for an accurate interpretation of the nannoplankton productivity record, since diagenetic processes can bias the productivity signal, as we demonstrate for Sr/Ca measurements in the fine (<20 μm and other size fractions obtained from bulk sediments from Site 1265. At this site, the paleoproductivity signal as reconstructed from the Sr/Ca appears to be governed by cyclic changes, possibly orbital forcing, resulting in a 20–30% variability in Sr/Ca in dominant genera as obtained by ion probe. The ~13 to 21
Keery, John S.; Holden, Philip B.; Edwards, Neil R.
The early Eocene, from about 56 Ma, with high atmospheric CO2 levels, offers an analogue for the response of the Earth's climate system to anthropogenic fossil fuel burning. In this study, we present an ensemble of 50 Earth system model runs with an early Eocene palaeogeography and variation in the forcing values of atmospheric CO2 and the Earth's orbital parameters. Relationships between simple summary metrics of model outputs and the forcing parameters are identified by linear modelling, providing estimates of the relative magnitudes of the effects of atmospheric CO2 and each of the orbital parameters on important climatic features, including tropical-polar temperature difference, ocean-land temperature contrast, Asian, African and South (S.) American monsoon rains, and climate sensitivity. Our results indicate that although CO2 exerts a dominant control on most of the climatic features examined in this study, the orbital parameters also strongly influence important components of the ocean-atmosphere system in a greenhouse Earth. In our ensemble, atmospheric CO2 spans the range 280-3000 ppm, and this variation accounts for over 90 % of the effects on mean air temperature, southern winter high-latitude ocean-land temperature contrast and northern winter tropical-polar temperature difference. However, the variation of precession accounts for over 80 % of the influence of the forcing parameters on the Asian and African monsoon rainfall, and obliquity variation accounts for over 65 % of the effects on winter ocean-land temperature contrast in high northern latitudes and northern summer tropical-polar temperature difference. Our results indicate a bimodal climate sensitivity, with values of 4.36 and 2.54 °C, dependent on low or high states of atmospheric CO2 concentration, respectively, with a threshold at approximately 1000 ppm in this model, and due to a saturated vegetation-albedo feedback. Our method gives a quantitative ranking of the influence of each of the
Khan, Natasha; Rehman, Khaista; Ahmad, Sajjad; Khokher, Jamil; Hajana, M. Iqbal; Hanif, M.
In this study, seismic data from two wells (Pak G2-1 and Indus Marine-1C) and age diagnostic larger benthic foraminifera (LBF) within drill cuttings has been used for the first time to identify depositional sequences within the carbonates in the Offshore Indus Basin, Pakistan. The Offshore Indus is tectonically categorized as a passive continental margin where carbonates occur as shelf carbonates in the near offshore and on volcanic seamounts in deeper waters. Seismic data analysis has indicated the presence of minor faults and carbonate buildups above the igneous basement in the south. Patterns of the seismic reflections enabled definition of three seismic facies units identified as: Unit 1 basement, represented by chaotic, moderate amplitude reflection configuration; while parallel bedding and the drape of overlying strata is typical character of Unit 2, carbonate mound facies. The younger Miocene channels represent Unit 3. The diagnosis of Alveolina vredenburgi/cucumiformis biozone confirmed the Ilerdian (55-52 Ma) stage constituting a second order cycle of deposition for the Eocene carbonates (identified as Unit 2). The carbonate succession has been mainly attributed to an early highstand system tract (HST). The environmental conditions remained favorable leading to the development of keep-up carbonates similar to pinnacle buildups as a result of aggradation during late transgressive system tract and an early HST. The carbonate sequence in the south (Pak G2-1) is thicker and fossiliferous representing inner to middle shelf depths based on fauna compared to the Indus Marine-1C in the north, which is devoid of fossils. Three biozones (SBZ 5, SBZ 6 and SBZ 8) were identified based on the occurrence of LBF. The base of the SBZ 5 zone marks the larger foraminifera turnover and the Paleocene-Eocene (P-E) boundary. The LBF encountered in this study coincides with earlier findings for the P-E boundary. Our findings indicate that the entire Ilerdian stage ranges from 55
Colombera, Luca; Arévalo, Oscar J.; Mountney, Nigel P.
The Tremp Group of the Tremp-Graus Basin (Southern Pyrenees, Spain) is a succession of predominantly continental origin that records the Paleocene-Eocene Thermal Maximum (PETM), a transient episode of extreme global warming that occurred across the Paleocene-Eocene boundary. For this succession, the stratigraphic position of the PETM is accurately determined, and histories of tectonic and sea-level controls are well constrained. Building upon previous studies, this work assesses changes in sedimentary architecture through the PETM in the Tremp Group, based on quantitative sedimentological analyses documented over a km-scale strike-oriented transect in the Arén area, with the scope to better understand the response of this alluvial system to the hyperthermal event. The analysed features represent a partial record of the geomorphic organization and processes of the system at the time of deposition, and are therefore interpretable in terms of geomorphic change in alluvial landscapes caused by the PETM. The record of the PETM, as previously recognized, begins at a time when erosional palaeotopographic relief was developed and deposition was confined in valleys. A shift between valley back-filling and widespread aggradation is observed at the onset of the PETM interval, which demonstrates uniquely the impact of the hyperthermal on both depositional loci and interfluves. Compared to underlying strata, the interval that embodies the onset and main phase of the PETM is characterized by: (i) higher proportion of channel deposits; (ii) channel complexes of greater average thickness and width; (iii) barforms and channel fills that are slightly thicker; (iv) increased thickness of sets of cross-stratified sandstones; (v) similar values of maximum extraclast size, by architectural element. An evident change in the facies organization of channel deposits is also seen through the stratigraphy, though this appears to predate the PETM. Increased channel-body density in the PETM
Scotchman, J. I.; Pickering, K. T.; Robinson, S. A.
Sub-Milankovitch millennial scale climate variability appears ubiquitous throughout the Quaternary and Pleistocene palaeoenvironmental records (e.g. McManus et al., 1999) yet the driving mechanism remains elusive. Possible mechanisms are generally linked to Quaternary-specific oceanic and cryospheric conditions (e.g. Maslin et al., 2001). An alternative external control, such as solar forcing, however, remains a compelling alternative hypothesis (e.g. Bond et al., 2001). This would imply that millennial-scale cycles are an intrinsic part of the Earth’s climatic system and not restricted to any specific period of time. Determining which of these hypotheses is correct impacts on our understanding of the climate system and its role as a driver of cyclic sedimentation during both icehouse and greenhouse climates. Here we show that Middle Eocene, laminated deep-marine sediments deposited in the Ainsa Basin, Spanish Pyrenees, contain 1,565-year (469 mm) cycles modulated by a 7,141-year (2157 mm) period. Climatic oscillations of 1,565-years recorded by element/Al ratios, are interpreted as representing climatically driven variation in sediment supply (terrigenous run-off) to the Ainsa basin. Climatic oscillations with this period are comparable to Quaternary Bond (~1,500-year), Dansgaard-Oeschger (~1,470-year) and Heinrich (~7,200-year) climatic events. Recognition of similar millennial-scale oscillations in the greenhouse climate of the Middle Eocene would appear inconsistent with an origin dependent upon Quaternary-specific conditions. Our observations lend support for pervasive millennial-scale climatic variability present throughout geologic time likely driven by an external forcing mechanism such as solar forcing. References Bond, G., Kromer, B., Beer, J., Muscheler, R., Evans, M.N., Showers, W., Hoffmann, S., Lotti-Bond, R., Hajdas, I., Bonani, G. 2001. Persistent Solar Influence on North Atlantic Climate During the Holocene. Science, 294, 2130-2136 Maslin, M
Bouchal, J. M.
The Florissant Fossil Beds National Monument is located in Teller County in central Colorado, at approximate latitude 38°54'N and longitude 105°13'. The lithologies of the Florissant Formation consist of coarse-grained arkosic and volcanoclastic sandstones and conglomerates, finer shale, and tuffaceus mudstone and siltstone. It is divided into six units, mostly of lacustrine and fluvial origin with volcanic sediments interfingering and topping the strata. Volcanic units have been dated using the 40Ar/39Ar single-crystal method, giving an absolute age of ca. 34 Ma for the upper fossiliferous sedimentary unit. This pinpoints the formation of the Florissant sediments at the end of the Eocene, providing fruitful insight into the changing palaeoecosystem of the region at the dawn of the Oligocene. The formation is very well known for its rich fossil insect fauna and well preserved plant macrofossils found in the shale units, and the silicified tree stumps occurring in the lower mudstone unit. The sample used for this study originates from the upper shale unit, the fifth unit from the base of the formation. Previous studies on the plant macrofossils, mesofossils and the palynoflora have shown that during the late Eocene the surroundings of Florissant palaeo-lake were covered by diverse mixed broad-leaved evergreen/deciduous and needle-leafed forests. Until now pollen from the Florissant Formation has mostly been described according to conventional morphological nomenclature, using light microscopy (LM) only. In this study the same individual pollen grains are investigated using both LM and scanning electron microscopy (SEM), by means of single grain technique. This provides best exploitable results concerning a more detailed resolution regarding taxonomy and more accurate identifications. The main goal of this study is to compile a well resolved taxonomic species list based on the palynoflora, to clarify the generic and species diversity of selected families (e
Perez-Cruz, L. L.; Fucugauchi, J. U.
Chicxulub crater was formed by an asteroid impact on the Yucatan carbonate platform about 65.5 Ma ago at the Cretaceous/Paleogene (K/Pg) boundary. After crater formation, carbonate deposition gradually covered the structure, preserving a largely undisturbed sedimentary record for the Paleogene. As part of the studies, drilling programs with continuous core recovery have been conducted over the past years. The cores preserve a record of post-impact processes, life recovery in target area, platform evolution and emergence, sea-level changes and the paleoceanographic and paleoclimatic conditions in the region. To contribute to the knowledge of the Paleocene-Eocene warming and changes in oceanic hydrography, we carried out x-ray fluorescence and stable oxygen and carbon isotope studies of the carbonate sections on top of the impact breccias sampled in the Santa Elena and Yaxcopoil-1 boreholes. The Santa Elena borehole is located 110 km radial distance from crater center, outside the crater rim. Yaxcopoil-1 borehole is located 62 km away from crater center in the terrace zone inside the crater rim. The Cenozoic sequence is about 332 m and 796 m thick in the Santa Elena and Yaxcopoil-1 boreholes. The stable isotope records are correlated to data on marine sediment cores from the ocean drilling projects for the Paleocene and Eocene, which permit inferences on the paleoceanographic and paleoclimatic reconstruction as well as additional stratigraphic constraints for the sections. The positive bulk carbon isotope (δ13C) values in the basal Paleocene sediments reflect a return or enhancement of ocean productivity following plankton extinctions following the K/Pg impact. The Paleocene represents a time of global warmth temperatures with low vertical and latitudinal gradients in the oceans, likely resulting from elevated CO2 levels. δ13C values in the Santa Elena borehole show increases between 3 and 4 % by the late Paleocene that decrease to levels prevailing before the K
Fucugauchi, J. U.; Perez-Cruz, L. L.
Chicxulub formed 66 Ma ago by an asteroid impact on the Yucatan carbonate platform, southern Gulf of Mexico. Impact produced a 200 km diameter crater, platform fracturing, deformation and ejecta emplacement. Carbonate sedimentation restarted and crater was covered by up to 1 km of sediments. Drilling programs have sampled the Paleogene sediments, which record the changing sedimentation processes in the impact basin and platform. Here, results of a study of the Paleocene-Eocene sediments cored in the Santa Elena borehole are used to characterize the K/Pg and PETM. The borehole reached a depth of 504 m and was continuously cored, sampling the post-impact sediments and impact breccias, with contact at 332 m. For this study, we analyzed the section from ~230 to ~340 m, corresponding to the upper breccias and Paleocene-Eocene sediments. The lithological column, constructed from macroscopic and thin-section petrographic analyses, is composed of limestones and dolomitized limestones with several thin clay layers. Breccias are melt and basement clast rich, described as a suevitic unit. Section is further investigated using paleomagnetic, rock magnetic, X-ray fluorescence geochemical and stable isotope analyses. Magnetic polarities define a sequence of reverse to normal, which correlate to the geomagnetic polarity time scale from chrons 29r to 26r. The d13 C values in the first 20 m interval range from 1.2 to 3.5 %0 and d18 O values range from -1.4 to -4.8 %0. Isotope values show variation trends that correlate with the marine carbon and oxygen isotope patterns for the K-Pg boundary and early Paleocene. Positive carbon isotopes suggest relatively high productivity, with apparent recovery following the K-Pg extinction event. Geochemical data define characteristic trends, with Si decreasing gradually from high values in the suevites, low contents in Paleocene sediments with intervals of higher variability and then increased values likely marking the PETM. Variation trends are
Agnini, Claudia; Spofforth, David J. A.; Dickens, Gerald R.; Rio, Domenico; Pälike, Heiko; Backman, Jan; Muttoni, Giovanni; Dallanave, Edoardo
We present records of stable carbon and oxygen isotopes, CaCO3 content, and changes in calcareous nannofossil assemblages across an 81 m thick section of upper Paleocene-lower Eocene marine sedimentary rocks now exposed along the Cicogna Stream in northeast Italy. The studied stratigraphic section represents sediment accumulation in a bathyal hemipelagic setting from approximately 57.5 to 52.2 Ma, a multi-million-year time interval characterized by perturbations in the global carbon cycle and changes in calcareous nannofossil assemblages. The bulk carbonate δ13C profile for the Cicogna section, once placed on a common timescale, resembles that at several other locations across the world, and includes both a long-term drop in δ13C and multiple short-term carbon isotope excursions (CIEs). This precise correlation of widely separated δ13C records in marine sequences results from temporal changes in the carbon composition of the exogenic carbon cycle. However, diagenesis has likely modified the δ13C record at Cicogna, an interpretation supported by variations in bulk carbonate δ18O, which do not conform to expectations for a primary signal. The record of CaCO3 content reflects a combination of carbonate dilution and dissolution, as also inferred at other sites. Our detailed documentation and statistical analysis of calcareous nannofossil assemblages show major differences before, during and after the Paleocene-Eocene Thermal Maximum. Other CIEs in our lower Paleogene section do not exhibit such a distinctive change; instead, these events are sometimes characterized by variations restricted to a limited number of taxa and transient shifts in the relative abundance of primary assemblage components. Both long-lasting and short-lived modifications to calcareous nannofossil assemblages preferentially affected nannoliths or holococcoliths such as Discoaster, Fasciculithus, Rhomboaster/Tribrachiatus, Sphenolithus and Zygrhablithus, which underwent distinct variations in
Grein, Michaela; Roth-Nebelsick, Anita; Wilde, Volker; Konrad, Wilfried; Utescher, Torsten
Various proxies (such as pedogenic carbonates, boron isotopes or phytoplankton) and geochemical models were applied in order to reconstruct palaeoatmospheric carbon dioxide, partially providing conflicting results. Another promising proxy is the frequency of stomata (pores on the leaf surface used for gaseous exchange). In this project, fossil plant material from the Messel Pit (Hesse, Germany) is used to reconstruct atmospheric carbon dioxide concentration in the Middle Eocene by analyzing stomatal density. We applied the novel mechanistic-theoretical approach of Konrad et al. (2008) which provides a quantitative derivation of the stomatal density response (number of stomata per leaf area) to varying atmospheric carbon dioxide concentration. The model couples 1) C3-photosynthesis, 2) the process of diffusion and 3) an optimisation principle providing maximum photosynthesis (via carbon dioxide uptake) and minimum water loss (via stomatal transpiration). These three sub-models also include data of the palaeoenvironment (temperature, water availability, wind velocity, atmospheric humidity, precipitation) and anatomy of leaf and stoma (depth, length and width of stomatal porus, thickness of assimilation tissue, leaf length). In order to calculate curves of stomatal density as a function of atmospheric carbon dioxide concentration, various biochemical parameters have to be borrowed from extant representatives. The necessary palaeoclimate data are reconstructed from the whole Messel flora using Leaf Margin Analysis (LMA) and the Coexistence Approach (CA). In order to obtain a significant result, we selected three species from which a large number of well-preserved leaves is available (at least 20 leaves per species). Palaeoclimate calculations for the Middle Eocene Messel Pit indicate a warm and humid climate with mean annual temperature of approximately 22°C, up to 2540 mm mean annual precipitation and the absence of extended periods of drought. Mean relative air
Gadagkar, Raghavendra; Kolatkar, Milind
Birds are remarkable for their extraordinary efforts at nest building and brood care. Given that so many species of birds spend so much time and effort at these activities, there is plenty of room for some species to take it easy, lay their eggs in the nests of other species and hitch-hike on their hosts. The cuckoo that lays its eggs in the nests of a variety of host species is well known. Indeed, over 80 species, i.e., over 1% of bird species are known to be such obligate inter-specific bro...
Mallon, Julie M.; Bildstein, Keith L.; Katzner, Todd E.
Birds use atmospheric updrafts to subsidize soaring flight. We observed highly variable soaring flight by Black Vultures (Coragyps atratus) and Turkey Vultures (Cathartes aura) in Virginia, USA, that was inconsistent with published descriptions of terrestrial avian flight. Birds engaging in this behavior regularly deviated vertically and horizontally from linear flight paths. We observed the soaring flight behavior of these 2 species to understand why they soar in this manner and when this behavior occurs. Vultures used this type of soaring mainly at low altitudes (birds because it permits continuous subsidized flight when other types of updraft are not available.
Waddell, L. M.; Moore, T. C.
Plate tectonic reconstructions indicate that the Arctic was largely isolated from the world ocean during the early and middle Eocene, with exchange limited to shallow, and possibly intermittent, connections to the North Atlantic and Tethys (via the Turgay Strait). Relative isolation, combined with an intensification of the hydrologic cycle under an Eocene greenhouse climate, is suspected to have led to the development of a low- salinity surface water layer in the Arctic that could have affected deep and intermediate convection in the North Atlantic. Sediment cores recently recovered from the Lomonosov Ridge by the IODP 302 Arctic Coring Expedition (ACEX) allow for the first assessment of the salinity of the Arctic Ocean during the early and middle Eocene. Stable isotope analysis performed on the structural carbonate of fish bone apatite from ~30 samples between the ages of ~55 and ~44 myr yielded δ18O values between -6.84‰ and -2.96‰ VPDB, with a mean value of -4.89‰. From the δ18O values we calculate that the Arctic Ocean was probably brackish during most of the early and middle Eocene, with an average salinity of 19 to 24‰. Negative excursions in the δ18O record (<-6‰) indicate three events during which the salinity of the Arctic surface waters was severely lowered: the Paleocene Eocene Thermal Maximum (PETM), the Azolla event at ~49 Ma, and a third previously unidentified event at ~46 Ma. During the PETM, low salinities developed under conditions of increased regional precipitation and runoff associated with extreme high latitude warmth and possible tectonic uplift in the North Atlantic. During the other two low-salinity events, sea level was lowered by ~20-30 m, implying a possible severing of Arctic connections to the world ocean. The most positive δ18O value (-2.96‰) occurs at ~45 Ma, the age of the youngest dropstone discovered in the ACEX sediments, and may therefore correspond to a climatic cooling rather than a high salinity event.
Full Text Available Here we present a high-resolution cyclostratigraphy based on X-ray fluorescence (XRF core scanning data from a new record retrieved from the tropical western Atlantic (Demerara Rise, ODP Leg 207, Site 1258. The Eocene sediments from ODP Site 1258 cover magnetochrons C20 to C24 and show well developed cycles. This record includes the missing interval for reevaluating the early Eocene part of the Geomagnetic Polarity Time Scale (GPTS, also providing key aspects for reconstructing high-resolution climate variability during the Early Eocene Climatic Optimum (EECO. Detailed spectral analysis demonstrates that early Eocene sedimentary cycles are characterized by precession frequencies modulated by short (100 kyr and long (405 kyr eccentricity with a generally minor obliquity component. Counting of both the precession and eccentricity cycles results in revised estimates for the duration of magnetochrons C21r through C24n. Our cyclostratigraphic framework also corroborates that the geochronology of the Eocene Green River Formation (Wyoming, USA is still questionable mainly due to the uncertain correlation of the "Sixth tuff" to the GPTS.
Right at the onset of the long-term Cenozoic cooling trend the dominant eccentricity-modulated precession cycles of ODP Site 1258 are interrupted by strong obliquity cycles for a period of ~800 kyr in the middle of magnetochron C22r. These distinct obliquity cycles at this low latitude site point to (1 a high-latitude driving mechanism on global climate variability from 50.1 to 49.4 Ma, and (2 seem to coincide with a significant drop in atmospheric CO2 concentration below a critical threshold between 2- and 3-times the pre-industrial level (PAL. The here newly identified orbital configuration of low eccentricity in combination with high obliquity amplitudes during this ~800-kyr period and the crossing of a critical pCO2 threshold may have led to the formation of the first ephemeral
Maslova, Natalia P; Kodrul, Tatiana M; Song, Yunsheng; Volkova, Lyudmila D; Jin, Jianhua
• Phylogenetic analysis of DNA sequence data has changed our view on the evolution and systematics of plant taxa. Based on the phylogenetic analysis of several molecular markers, fruit anatomy, and pollen morphology, the genera Altingia and Semiliquidambar were formally transferred to the genus Liquidambar. The new species of Liquidambar from the Eocene of South China significantly extends our knowledge of the variability of the morphological characters of this genus in the geological past. Fossil leaves in conjunction with data on the associated reproductive structures allow us to make inferences about patterns in the evolutionary history of Liquidambar.• Fossil leaves and associated reproductive structures preserved as impressions were described and compared with the corresponding organs of extant and fossil relatives. The morphological variation of numerous leaves was examined by stereomicroscopy.• Liquidambar maomingensis sp. nov. is characterized by polymorphic leaves including both palmately lobed and unlobed leaves. This study presents the first observations of such dimorphism in the fossil record of Liquidambar leaves. Two distinct leaf groups are interpreted as sun and shade leaves.• The fossil leaves and associated infructescences from Maoming probably belong to the same plant. The occurrence of fossil leaves similar to those of extant species previously considered within Semiliquidambar and Liquidambar with the associated infructescences close to those of Altingia provide paleobotanical evidence that justifies combining the genera Liquidambar, Altingia, and Semiliquidambar into the single genus Liquidambar as recently proposed based on molecular markers. © 2015 Botanical Society of America, Inc.
Rao, M. R.; Sahni, Ashok; Rana, R. S.; Verma, Poonam
Early Eocene sedimentary successions of south Asia, are marked by the development of extensive fossil-bearing, lignite-rich sediments prior to the collision of India with Asia and provide data on contemporary equatorial faunal and vegetational assemblages. One such productive locality in western India is the Vastan Lignite Mine representing approximately a 54-52 Ma sequence dated by the presence of benthic zone marker species, Nummulites burdigalensis burdigalensis. The present study on Vastan Lignite Mine succession is based on the spore-pollen and dinoflagellate cyst assemblages and documents contemporary vegetational changes. 86 genera and 105 species belonging to algal remains (including dinoflagellate cysts), fungal remains, pteridophytic spores and angiospermous pollen grains have been recorded. On the basis of first appearance, acme and decline of palynotaxa, three cenozones have been recognized and broadly reflect changing palaeodepositional environments. These are in ascending stratigraphic order (i) Proxapertites Spp. Cenozone, (ii) Operculodinium centrocarpum Cenozone and (iii) Spinizonocolpites Spp. Cenozone. The basal sequence is lagoonal, palm-dominated and overlain by more open marine conditions with dinoflagellate cysts and at the top, mangrove elements are dominant. The succession has also provided a unique record of fish, lizards, snakes, and mammals.
May, Jeffrey A.; Yeo, Ross K.; Warme, John E.
Field studies document an apparent eustatic control on facies patterns along a tectonically active margin. In the San Diego Embayment and northern Baja California, progradational-retrogradational shoreline sequences characterize Late Cretaceous and Eocene forearc stratigraphy. Extensive benthonic foraminifera and nannoplankton data provide control on the age and distribution of facies changes along these depositionally compact, bathymetrically steep-gradient margins. The complete stratigraphic package is arranged into three scales and patterns of depositional sequences. Timing and geometry of the two largest sequences provide relative sea-level curves that correlate exceptionally well with worldwide sea-level curves. The major depositional cycle is asymmetric, hundreds of meters thick, characterized by a thin basal retrogradational sequence overlain by a thick progradational sequence; each cycle correlates to a coastal-onlap "supercycle". Smaller scale stratigraphic rhythms, controlled by global "cycles" and "paracycles", compose depositional cycles. Local depostional pulses overprint these two larger order sequences of sedimentation. Coeval cycles and depositional rhythms in isolated coastal basins from Oregon to Baja California further indicate a primary eustatic control on sedimentation. Field-based facies analysis thus supports the use of the "Vail curve" or other coastal-onlap and global sea-level curves as a predictive tool in basin analysis.
An abundant and diversified assemblage of siliceous loose sponge spicules has been identified in the Late Eocene deposits cropping out along the southern coasts of Australia. Based on the comparison of the obtained spicules with those of living sponges, representatives of at least 43 species within 33 genera, 26 families, and 9 orders of "soft" Demospongiae and Homoscleromorpha have been identified in the assemblage. Within the studied sediments, the spicules representing demosponge orders Poecilosclerida, Hadromerida, and Astrophorida were the most diverse. The rest of the five demosponge orders (Halichondrida, Agelasida, Haplosclerida, Spirophorida, and Chondrosida) are represented by single families. Also, a single family Plakinidae within the class Homoscleromorpha that includes two genera was present. The diversity of spicules is similar in all studied samples and areas, even distant geographically, and there are only minor differences between the sections. That indicates a homogenous character of this rich siliceous sponge assemblage. Most of the studied sponge spicules have Recent equivalents among present-day siliceous spicules. However, the fossil ones are bigger which is most likely due to different environmental conditions. Among the recognized sponge species, at least eleven (Agelas cf. axifera, Agelas cf. wiedenmayeri, Penares sclerobesa, Histodermella australis, Trikentrion flabelliforme, Cliona cf. mucronata, Tethya cf. omanensis, Terpios sp., Placinolopha cf. sarai, Dotona pulchella, and Sigmosceptrella quadrilobata) are noted for the first time in the fossil record.
Fontorbe, Guillaume; Frings, Patrick J.; De La Rocha, Christina L.; Hendry, Katharine R.; Carstensen, Jacob; Conley, Daniel J.
Silicon isotope ratios (expressed as δ30Si) in marine microfossils can provide insights into silica cycling over geologic time. Here we used δ30Si of sponge spicules and radiolarian tests from the Paleogene Equatorial Transect (Ocean Drilling Program Leg 199) spanning the Eocene and Oligocene ( 50-23 Ma) to reconstruct dissolved silica (DSi) concentrations in deep waters and to examine upper ocean δ30Si. The δ30Si values range from -3.16 to +0.18‰ and from -0.07 to +1.42‰ for the sponge and radiolarian records, respectively. Both records show a transition toward lower δ30Si values around 37 Ma. The shift in radiolarian δ30Si is interpreted as a consequence of changes in the δ30Si of source DSi to the region. The decrease in sponge δ30Si is interpreted as a transition from low DSi concentrations to higher DSi concentrations, most likely related to the shift toward a solely Southern Ocean source of deep water in the Pacific during the Paleogene that has been suggested by results from paleoceanographic tracers such as neodymium and carbon isotopes. Sponge δ30Si provides relatively direct information about the nutrient content of deep water and is a useful complement to other tracers of deep water circulation in the oceans of the past.
Greenwalt, Dale E.; Goreva, Yulia S.; Siljeström, Sandra M.; Rose, Tim; Harbach, Ralph E.
Although hematophagy is found in ∼14,000 species of extant insects, the fossil record of blood-feeding insects is extremely poor and largely confined to specimens identified as hematophagic based on their taxonomic affinities with extant hematophagic insects; direct evidence of hematophagy is limited to four insect fossils in which trypanosomes and the malarial protozoan Plasmodium have been found. Here, we describe a blood-engorged mosquito from the Middle Eocene Kishenehn Formation in Montana. This unique specimen provided the opportunity to ask whether or not hemoglobin, or biomolecules derived from hemoglobin, were preserved in the fossilized blood meal. The abdomen of the fossil mosquito was shown to contain very high levels of iron, and mass spectrometry data provided a convincing identification of porphyrin molecules derived from the oxygen-carrying heme moiety of hemoglobin. These data confirm the existence of taphonomic conditions conducive to the preservation of biomolecules through deep time and support previous reports of the existence of heme-derived porphyrins in terrestrial fossils. PMID:24127577
Harding, Ian C.; Charles, Adam J.; Marshall, John E. A.; Pälike, Heiko; Roberts, Andrew P.; Wilson, Paul A.; Jarvis, Edward; Thorne, Robert; Morris, Emily; Moremon, Rebecca; Pearce, Richard B.; Akbari, Shir
Palaeoenvironmental manifestations of the Paleocene-Eocene thermal maximum (PETM; ~ 56 Ma) are relatively well documented in low- to mid-latitude settings and at high southern latitudes, but no documented high northern latitude sites record the entire hyperthermal event. We present high-resolution multi-proxy records from a PETM succession on Spitsbergen in the high Arctic (palaeolatitude ~ 75 °N). By comparing our results with those from Integrated Ocean Drilling Program Site 302-4A, we document regional palaeoenvironmental variations in the expression of the PETM, with evidence for major differences in basin-margin vegetation and water column oxygen depletion. Sedimentological, palynological and geochemical data demonstrate a pre-PETM sea level rise in Spitsbergen before the - 4‰ δ 13C TOC excursion, which culminated in maximum flooding during the peak of the event. The appearance of the dinoflagellate cyst Apectodinium before the onset of the carbon isotope excursion (CIE) corroborates that environmental change in the Arctic had begun prior to the CIE. Sedimentological and palynological evidence indicate that elevated terrestrial runoff resulted in water column stratification, providing further evidence for an intensification of the hydrological cycle during the PETM.
Zeebe, R. E.
Paleorecords show that the Paleocene-Eocene Thermal Maximum (PETM, 56 Ma) was associated with a large carbon isotope excursion and global warming >5 K. Surprisingly, the records indicate little if any decline in, for instance, stable carbon isotope values after 50 kyr. Similar durations of the PETM main phase can be inferred from oxygen isotope records (indicating temperature) and surface ocean carbonate chemistry proxies. The inferred main phase duration of > 50 kyr is also independent of the age model applied. One age model is based on orbital cycle stratigraphy, the other on extraterrestrial 3He concentrations. Despite differences in the age models, they agree that the duration of the PETM main phase lasted for at least 50 kyr, a duration that is also consistent with observations from the most expanded terrestrial sequences. Conventional carbon-cycle/climate models forced with an initial carbon input pulse over ~10 kyr fail to reproduce the long duration of the PETM without invoking additional, slow carbon release over more than 50~kyr (so-called bleeding). However, a potential carbon source for the bleeding, as well as its release mechanism have hitherto remained elusive. In this presentation, I will suggest a plausible mechanism for the long duration of the PETM based on first-principle calculations. If the mechanism is also applicable to the near future, the magnitude and duration of future warming could substantially exceed current projections.
Pälike, Cecily; Delaney, Margaret L.; Zachos, James C.
amounts of 13C-depleted carbon were released to the oceans and atmosphere during a period of abrupt global warming at the Paleocene-Eocene thermal maximum (PETM) (˜55 Ma). Investigations of qualitative sedimentologic and paleontologic redox proxies such as bioturbation and benthic assemblages from pelagic and hemipelagic sections suggest transient reductions in bottom water oxygen during this interval, possibly on a global scale. Here, we present bulk sediment manganese (Mn) and uranium (U) enrichment factors (EF) in Atlantic and Pacific deep-sea cores to constrain relative paleoredox changes across the PETM. Mn EF range from 1 to 9 in Atlantic sites, 1 to 35 in Southern Ocean sites, and are at crustal averages (EF = 1) in Pacific sites. U EF range from 1 to 5 in Atlantic sites, 1 to 90 in Southern Ocean sites, and are at crustal averages in Pacific sites. Our results indicate suboxic conditions prior to, during, and in the recovery from the PETM at intermediate depth sites in the Atlantic and Southern Ocean while the Pacific sites remained relatively oxygenated. The difference in oxygenation between the Atlantic and Pacific sites leads us to suggest the source for isotopically light carbon release during the PETM was in the Atlantic.
Full Text Available The study deals with environmental changes during the Paleocene–Eocene Thermal Maximum (PETM and its background conditions in Spitsbergen through analysis of benthic foraminiferal assemblages (FA in a section drilled in the Paleogene Central Basin. The impact of this extreme global warming occurs here in prodelta shelf mudstones composing the lower part of the Gilsonryggen Member (Frysjaodden Formation. The start of the PETM perturbation is marked by a faunal turnover, in which the medium-diversity circumpolar Reticulophragmium assemblage was replaced by a low-diversity Trochammina fauna. During the hyperthermal period, benthic foraminiferal diversity decreased severely, while the dominance of small-sized taxa with epifaunal morphology strongly increased. This low-diversity fauna occurs in sediments with a reduced thorium/uranium ratio (proxy for oxygenation and kaolinite enrichment (proxy for high humidity. The faunal changes were thus caused by the combined effects of hypoxic and hyposaline conditions in a stratified water column, due to extreme warming with its accompanying intensified hydrologic cycle. The PETM acme coincides with the maximum flooding surface (MFS of the Gilsonryggen depositional sequence, composed of the Gilsonryggen Member and the overlying Battfjellet and Aspelintoppen formations. The transgressive phase of the sequence was initiated by local tectonics, while the eustatic sea-level rise of the PETM was superimposed on this transgression.To access the supplementary material for this article, please see supplementary files under Article Tools online.
Khader, A.; McKee, M.
Value of information (VOI) analysis evaluates the benefit of collecting additional information to reduce or eliminate uncertainty in a specific decision-making context. It makes explicit any expected potential losses from errors in decision making due to uncertainty and identifies the “best” information collection strategy as one that leads to the greatest expected net benefit to the decision-maker. This study investigates the willingness to pay for groundwater quality monitoring in the Eocene Aquifer, Palestine, which is an unconfined aquifer located in the northern part of the West Bank. The aquifer is being used by 128,000 Palestinians to fulfill domestic and agricultural demands. The study takes into account the consequences of pollution and the options the decision maker might face. Since nitrate is the major pollutant in the aquifer, the consequences of nitrate pollution were analyzed, which mainly consists of the possibility of methemoglobinemia (blue baby syndrome). In this case, the value of monitoring was compared to the costs of treating for methemoglobinemia or the costs of other options like water treatment, using bottled water or importing water from outside the aquifer. And finally, an optimal monitoring network that takes into account the uncertainties in recharge (climate), aquifer properties (hydraulic conductivity), pollutant chemical reaction (decay factor), and the value of monitoring is designed by utilizing a sparse Bayesian modeling algorithm called a relevance vector machine.
Patricia G Weaver
Full Text Available BACKGROUND: Preservation of original organic components in fossils across geological time is controversial, but the potential such molecules have for elucidating evolutionary processes and phylogenetic relationships is invaluable. Chitin is one such molecule. Ancient chitin has been recovered from both terrestrial and marine arthropods, but prior to this study had not been recovered from fossil marine mollusks. METHODOLOGY/PRINCIPAL FINDINGS: Organics consistent with β-chitin are recovered in cuttlebones of Mississaepia mississippiensis from the Late Eocene (34.36 million years ago marine clays of Hinds County, Mississippi, USA. These organics were determined and characterized through comparisons with extant taxa using Scanning Electron Microscopy/Energy Dispersive Spectrometry (SEM/EDS, Field Emission Scanning Electron Microscopy (Hyperprobe, Fourier Transmission Infrared Spectroscopy (FTIR and Immunohistochemistry (IHC. CONCLUSIONS/SIGNIFICANCE: Our study presents the first evidence for organics consistent with chitin from an ancient marine mollusk and discusses how these organics have been degraded over time. As mechanisms for their preservation, we propose that the inorganic/organic lamination of the cuttlebone, combined with a suboxic depositional environment with available free Fe(2+ ions, inhibited microbial or enzymatic degradation.
Full Text Available The Paleocene-Eocene Thermal Maximum (PETM occurred approximately 55 million years ago, and is one of the most dramatic abrupt global warming events in the geological record. This warming was triggered by the sudden release of thousands of gigatons of carbon into the atmosphere and is widely perceived to be the best analogue for current anthropogenic climate change. Yet, the mechanism of recovery from this event remains controversial. A massive increase in the intensity of the marine biological pump ("productivity feedback" has been suggested to cause a drawdown of atmospheric CO2 and subsequent carbon sequestration in the ocean. A re-evaluation of the "productivity feedback hypothesis", based on biogenic barium mass accumulation rates (Ba-MARs for a site in the Southern Ocean, finds that any increase in export production lagged the initial carbon release by at least ~70 000 years. This implies that export production did not facilitate rapid removal of excess carbon from the atmosphere. Thus, the most likely mechanism for carbon removal appears to be silicate weathering, which occurred at much slower rates than previously assumed.
Dominici, Stefano; Zuschin, Martin
The middle Eocene, a time of global transition from greenhouse to icehouse climate, was approached through high-resolution stratigraphy at a few classic localities of the Paris Basin. Quantitative data on the distribution of molluscan species abundance, collected at 12 different shell beds representative of the middle Lutetian and the lower Bartonian, formed the basis for a palaeoecological study. The succession can be subdivided into a hierarchy of depositional sequences, interpreted as the product of relative sea-level change. Abundance distributions are better correlated with 5th-order depositional sequences than geographic locality, suggesting that sea-level played an important role in the distribution of palaeocommunities. Rarefied diversities were measured and compared with analogous data from modern tropical and warm-temperate intertidal and subtidal communities. The palaeoecological analysis shows that sea-level variation is responsible for a major change in the upper part of the middle Lutetian, leading from high-diversity subtidal to low-diversity intertidal and shallow subtidal palaeocommunities. The study did not confirm that the stage-level drop in species richness documented in this basin is related to the global climatic deterioration. Instead, the global climatic signal might be obscured in the Paris Basin by facies control.
García, Daniel; Martínez, Daniel; Stouffer, Daniel B; Tylianakis, Jason M
Exotic species are thought to alter the structure of natural communities and disrupt ecosystem functioning through invasion. Nevertheless, exotic species may also provide ecological insurance when they contribute to maintain ecosystem functions after the decline of native species following anthropogenic disturbance. Here, this hypothesis is tested with the assemblage of frugivorous birds and fleshy-fruited plants of New Zealand, which has suffered strong historical declines in native birds while simultaneously gaining new frugivores introduced by European settlers. We studied the plant-frugivore assemblage from measures of fruit and bird abundances and fruit consumption in nine forest patches, and tested how this changed across a gradient of relative abundance of exotic birds. We then examined how each bird species' role in the assemblage (the proportion of fruits and the number of plant species consumed) varied with their relative abundance, body size and native/exotic status. The more abundant and, to a lesser extent, larger birds species consumed a higher proportion of fruits from more plant species. Exotic birds consumed fruits less selectively and more proportionate to the local availability than did native species. Interaction networks in which exotic birds had a stronger role as frugivores had higher generalization, higher nestedness and higher redundancy of plants. Exotic birds maintained frugivory when native birds became rarer, and diversified the local spectrum of frugivores for co-occurring native plants. These effects seemed related to the fact that species abundances, rather than trait-matching constraints, ultimately determined the patterns of interactions between birds and plants. By altering the structure of plant-frugivore assemblages, exotic birds likely enhance the stability of the community-wide seed dispersal in the face of continued anthropogenic impact. © 2014 The Authors. Journal of Animal Ecology © 2014 British Ecological Society.
Lacey, M. E. R.
This thesis presents an interdisciplinary approach towards understanding the ways in which Anglo-Saxons perceived the birds around them and the cultural associations with which we find them endowed in the literature. It focuses on closely examining the entire range of primary sources available to us in order to build as accurate and as complete a picture of Anglo-Saxon bird-lore as possible, and it stresses the indivisibility of observational experiences of birds and their cultural associatio...
the establishment of long grass aisles at both sides of the main runway and a weekly record of the distribution of lapwings. Virtually no birds were...an exhibition was organized in the Air France Communication Showroom to inform aircrews about birdstrike hazards. This exhibition was supported by a...sensors, Virtual (computerized) and artificial birds, and detailed analyses of existing data bases to detect and explain "non-randomness" in bird strikes
Shorter, Angela; Segers, Marcia
How can an engineer design a bird feeder that attracts many birds? This question resulted from kindergarten students' observations of the bird feeders in their school's bird sanctuary. The challenging question is the heart of project-based learning (PBL), a teaching strategy in which students tackle real-world problems and design projects to solve…
... 50 Wildlife and Fisheries 6 2010-10-01 2010-10-01 false Subsistence migratory bird species. 92.22... (CONTINUED) MISCELLANEOUS PROVISIONS MIGRATORY BIRD SUBSISTENCE HARVEST IN ALASKA General Regulations Governing Subsistence Harvest § 92.22 Subsistence migratory bird species. You may harvest birds or gather...
Encinas-Nagel, Nuri; Enderlein, Dirk; Piepenbring, Anne; Herden, Christiane; Heffels-Redmann, Ursula; Felippe, Paulo A.N.; Arns, Clarice; Hafez, Hafez M.
Avian bornavirus (ABV) has been identified as the cause of proventricular dilatation disease in birds, but the virus is also found in healthy birds. Most studies of ABV have focused on captive birds. We investigated 86 free-ranging psittacine birds in Brazil and found evidence for natural, long-term ABV infection. PMID:25417715
... 50 Wildlife and Fisheries 6 2010-10-01 2010-10-01 false Custody of birds of another. 20.37 Section... WILDLIFE AND PLANTS (CONTINUED) MIGRATORY BIRD HUNTING Possession § 20.37 Custody of birds of another. No person shall receive or have in custody any migratory game birds belonging to another person unless such...
... 50 Wildlife and Fisheries 6 2010-10-01 2010-10-01 false Possession of live birds. 20.38 Section 20... WILDLIFE AND PLANTS (CONTINUED) MIGRATORY BIRD HUNTING Possession § 20.38 Possession of live birds. Every migratory game bird wounded by hunting and reduced to possession by the hunter shall be immediately killed...
... 50 Wildlife and Fisheries 6 2010-10-01 2010-10-01 false Transportation of birds of another. 20.42... WILDLIFE AND PLANTS (CONTINUED) MIGRATORY BIRD HUNTING Transportation Within the United States § 20.42 Transportation of birds of another. No person shall transport migratory game birds belonging to another person...
... 50 Wildlife and Fisheries 6 2010-10-01 2010-10-01 false Importation of birds of another. 20.62... WILDLIFE AND PLANTS (CONTINUED) MIGRATORY BIRD HUNTING Importations § 20.62 Importation of birds of another. No person shall import migratory game birds belonging to another person. ...
... 50 Wildlife and Fisheries 6 2010-10-01 2010-10-01 false Gift of migratory game birds. 20.40... WILDLIFE AND PLANTS (CONTINUED) MIGRATORY BIRD HUNTING Possession § 20.40 Gift of migratory game birds. No person may receive, possess, or give to another, any freshly killed migratory game birds as a gift...
... 9 Animals and Animal Products 1 2010-01-01 2010-01-01 false Replacement birds and poultry. 82.15...- EASE (END) AND CHLAMYDIOSIS Exotic Newcastle Disease (END) § 82.15 Replacement birds and poultry. Birds... eradicated and that replacement birds or poultry will not become infected with END. ...
... 19 Customs Duties 1 2010-04-01 2010-04-01 false Game animals and birds. 10.76 Section 10.76... TREASURY ARTICLES CONDITIONALLY FREE, SUBJECT TO A REDUCED RATE, ETC. General Provisions Animals and Birds § 10.76 Game animals and birds. (a) The following classes of live game animals and birds may be...
N.L. Rodenhouse; S.N. Matthews; K.P. McFarland; J.D. Lambert; L.R. Iverson; A. Prasad; T.S. Stillett; R.T. Holmes
We used three approaches to assess potential effects of climate change on birds of the Northeast. First, we created distribution and abundance models for common bird species using climate, elevation, and tree species variables and modeled how bird distributions might change as habitats shift. Second, we assessed potential effects on high-elevation birds, especially...
... promulgating migratory bird permit regulations for a permit to use raptors (birds of prey) in abatement activities. Abatement means the use of trained raptors to flush, scare (haze), or take birds or other.... The policy memorandum and conditions govern current administration of Federal Migratory Bird Special...
Detailed observations were made of Pallid Harrier Circus macrourus behaviour when hunting birds, the bird prey species and bird capture success in northern Cameroon. Four hunting strategies are described to capture birds: fast contour flight, overt approach with rapid acceleration, covert ambush, and stoop from flight.
... and suggestions on migratory bird permit regulations for a permit to use raptors (birds of prey) in abatement activities. Abatement means the use of trained raptors to flush, scare (haze), or take birds or... and held in ``temporary'' caging for extended periods of time, i.e., multiple birds held in a trailer...
Luciani, V.; Agnini, C.; Fornaciari, E.; Giusberti, L.; Rio, D.; Spofforth, D. J. A.; Pälike, H.
The transient (ca. 500 kyr) climatic warming event at ca. 40 Ma, known as Middle Eocene Climatic Optimum (MECO), significantly interrupted the overall cooling trend of the Middle Eocene. Originally documented in several deep sea sites at the Southern Ocean (Bohaty and Zachos, 2003), now it appears to be recorded worldwide by pronounced changes of the ^13C and ^18O values and coeval oscillations in global CCD (Tripati et al, 2005). Information on the planktonic foraminiferal response to this event is so far lacking. Here we present a detailed planktonic foraminiferal analysis of the MECO interval from a marginal basin of the central-western Tethys (Alano di Piave section, northeastern Italy). The expanded and continuous Alano section provides an excellent record of this event and offers an unique opportunity to better understand the role of climate upon calcareous plankton evolution. The initiation of the MECO occurs within magnetochron C18r at ca. 40.5 Ma with minimum ^18O and ^13C values achieved at the base of C18n.2n ca. 40.13 Ma, which are interpreted to represent peak warming conditions. Two sapropel-like, organic-rich intervals coincide with the major change in ^13C record at Alano (Agnini et al., 2007a; Spofforth et al., 2008). The MECO event correlates the E12 (P13) and lower E3 (P14) planktonic foraminiferal zones. The high-resolution quantitative planktonic foraminiferal analysis performed on both >38 m and >63 m fraction reveals pronounced and complex changes indicating a strong environmental perturbation that parallels the variations of the stable isotope curves. These changes are primarily represented by the marked increase in abundance of the eutrophic subbotinids and of the small, low-oxygen tolerant Streptochilus, Chiloguembelina and Pseudohastigerina, by the consistent and significant entrance of the eutrophic opportunist triserial Jenkynsina and of Pseudoglobigerinella bolivariana, typical species of high-productivity, upwelling areas. The
Göçmengil, Gönenç; Karacık, Zekiye; Genç, Ş. Can
Obliteration of the Mesozoic Neo-Tethyan Ocean and succeeding collision of the micro plates along the northern part of Turkey lead the development of the İzmir-Ankara-Erzincan suture zone (IAESZ). The suturing and collision stages terminate with the amalgamation of the three different crustal blocks (Pontides, Central Anatolian Crystalline Complex and Anatolide-Tauride Block) in the Paleocene-Early Eocene period. After the collisional stage; a new phase of extension and magmatism concomitantly developed at the both sides and as well as along the IAESZ during the Middle Eocene period. However, the origin, mechanism and driving force of the post-collisional magmatism is still enigmatic. To understand and better constrain the syn-to post collisional evolutionary stages, we have carried out volcano-stratigraphy and geochemistry based study on the middle Eocene magmatic associations along a transect ( 100 km) from Pontides to the Central Anatolian Crystalline Complex (CACC) at the NE part of the Turkey. Middle Eocene magmatic activity in the region has been represented by calc-alkaline, alkaline, shoshonitic volcanic and granitic rocks together with scarce gabbroic intrusions. We particularly focused on middle Eocene volcano-sedimentary successions (MEVSS) to constrain the tectono-magmatic evolution of the abovementioned transect. The volcano-sedimentary succsessions are coevally developed and cover the crustal blocks (Pontides and CACC) and the IAESZ with a region wide unconformity. We have differentiated three lava series (V1-V2-V3) and their sub-groups (V1a-V1b; V2a-V2b) in MEVSS. Generally, all lava series have middle-K to shoshonitic composition with distinct subduction characteristics. V1 series is marked by presence of hydrous phenocrysts such as amphibole+biotite. V1a sub-group constitute the first volcanic product and characterized by the high Mg# (42-69); alkaline basaltic andesite, and hawaiites. V1b sub-group is represented by calc-alkaline, low Mg# (24
Cardona, A.; Weber, M. B.; Bayona, G.; Jaramillo, C.; Montes, C.; Ojeda, G.; Duque, J. F.; Salazar, C. A.
Geological characteristics from the Cretaceous to Eocene metamorphic and igneous basement rocks from the Guajira and the NW corner of the Sierra Nevada de Santa Marta massifs in the northern Colombian Caribbean region, and the stratigraphy of adjacent basins, reveal different stages of growth of a segmented Late Cretaceous to Eocene accretionary orogen formed by interaction between the NW margin of South America and the Caribbean plate. Extensive intercalations of metavolcano-sedimentary rocks in the Santa Marta region, chemical composition of spinels and pyroxenes from serpentinized mantle tectonites and gabbros, as well as whole rock geochemistry from basaltic dykes from the Guajira massifs, record the evolution of a Cretaceous intra-oceanic arc in a succesion of Mariana- and Chile-type subduction styles. Geological and positive gravity signatures suggest that this arc was accreted and tectonically underplated to the continental margin of South America, creating a Maastrichian-Paleocene syn-orogenic basin, presently cropping out in the Cesar-Rancheria valley. After this collisional event, subduction of the Caribbean plate under South America started, as revealed by intrusion of composite stitched Late Paleocene-Eocene granitoids in the NW corner of the SNSM and Guajira massifs. These plutons show continental calc-alkaline geochemical signatures and more evolved Sr and Nd isotopic compositions than juvenile and older intra-oceanic arc magmatic rocks. This new subduction environment is linked to the convergence between North and South America at this time. Low pressure estimations and fast cooling rates between 450-250° for the Eocene granitoids in the Guajira and Santa Marta regions, indicate shallow < 7 Km depths of intrusion. Local unconformable stratigraphic relationships with Eocene-Miocene sediments at the Guajira region and overimposed post-magmatic low temperature deformation in some of these granitoids suggest an important Late Eocene unroofing and
California Natural Resource Agency — These data are summary statistics of abundances of birds counted within 100-m radius circles with 10-minute point counts at multiple sample points along 36 randomly...
Jones, M P
This article describes concepts of raptor behavior in captive birds of prey with inferences from the behavior of their wild counterparts that will assist the veterinarian in understanding the causes of managing abnormal behaviors.
Williams, B.K.; Nichols, J.D.
Mathematical modeling of migratory bird populations is reviewed in the context of migratory bird management. We focus on dynamic models of waterfowl, since most management-oriented migratory bird models concern waterfowl species. We describe the management context for these modeling efforts, with a focus on large-scale operational data collection programs and on processes by which waterfowl harvest is regulated and waterfowl habitats are protected and managed. Through their impacts on key population parameters such as recruitment and survival rate, these activities can influence population dynamics, thereby providing managers some measure of control over the status of populations. Recent applications of the modeling of waterfowl are described in terms of objectives, mathematical structures, and contributions to management. Finally, we discuss research needs and data limitations in migratory bird modeling, and offer suggestions to increase the value to managers of future modeling efforts.
van Zeeland, Yvonne
The use of behavior modifying drugs may be considered in birds with behavior problems, especially those refractory to behavior modification therapy and environmental management. To accomplish behavior change, a variety of drugs can be used, including psychoactive drugs, hormones, antihistamines,
California Department of Resources — These data are summary statistics of abundances of birds counted within 100-m radius circles with 10-minute point counts at 15 sample points within Spears and Didion...
A European-wide analysis of changing species distributions shows that butterflies outrun birds in the race to move northwards in response to climate change, but that neither group keeps up with increasing temperatures.
Vasić Vojislav F.
Full Text Available Figures of birds and bird heads are frequently represented in the Bronze and Iron Age in Europe. Birds are usually represented as amulets, vases and parts of vases, parts of complex objects or compositions and are among the most wide spread symbols that are connected with cult, magic and mythology of that time. There has been much discussion on this topic up to now. There are many assumptions concerning the role of different bird representations in the spiritual life of man of that period, which are still not firmly based. Our goal, in the study of this topic, is not to talk about these assumptions, more or less acceptable, but to pay more attention to the creation of bird figures and heads and to conclude to what extent it is possible to identify ornithological, among these numerous representations, individual birds, characteristics of their morphology and way of life. Namely, if the premise is correct, that the users of these ornithomorphic objects as well as their craftsmen, were able to differentiate bird types, morphologically and bionomically, or more correctly taxonomic groups of bird types (families, subfamilies, tribes genuses, to the extent to which these differences are expressed lexicological (nomenclatorilly, in the majority of classical languages then the attempt to make a further step towards the ornithological identification of motifs of these objects can be considered possible. In cases when it seems that the details on an object and their combination can be used for a more convincing identification, we take the freedom to speculate about the cultural-economic value of the recognized birds. Here we deal with three examples of bird representations from the Bronze and Iron Age of the Central Balkans. These are: the Dupljaja Cart, the Posamenterie Fibula from Dobrinci and the Bronze Cart from Glasinac, with the help of which we will try to demonstrate this new type of collaboration between archaeology and ornithology. We hope that
Anders Pape Møller
Full Text Available Animals living in areas contaminated by radioactive material from Chernobyl suffer from increased oxidative stress and low levels of antioxidants. Therefore, normal development of the nervous system is jeopardized as reflected by high frequencies of developmental errors, reduced brain size and impaired cognitive abilities in humans. Alternatively, associations between psychological effects and radiation have been attributed to post-traumatic stress in humans.Here we used an extensive sample of 550 birds belonging to 48 species to test the prediction that even in the absence of post-traumatic stress, there is a negative association between relative brain size and level of background radiation. We found a negative association between brain size as reflected by external head volume and level of background radiation, independent of structural body size and body mass. The observed reduction in brain size in relation to background radiation amounted to 5% across the range of almost a factor 5,000 in radiation level. Species differed significantly in reduction in brain size with increasing background radiation, and brain size was the only morphological character that showed a negative relationship with radiation. Brain size was significantly smaller in yearlings than in older individuals.Low dose radiation can have significant effects on normal brain development as reflected by brain size and therefore potentially cognitive ability. The fact that brain size was smaller in yearlings than in older individuals implies that there was significant directional selection on brain size with individuals with larger brains experiencing a viability advantage.
Griffin, D. R.
The use of a tracking radar to measure the flight paths of migrating birds on nights with opaque clouds is discussed. The effects of wind and lack of visual references are examined. The limitations of the radar observations are described, and samples of tracks obtained during radar observations are included. It is concluded that nonvisual mechanisms of orientation make it possible for birds to migrate in opaque clouds, but the exact nature of the sensory information cannot be determined by radar observations.
Birds of prey host a wide variety of endoparasites. The majority of these endoparasites are nematodes. They can be found mainly in the digestive and respiratory system. The current accepted phylogeny of nematodes found in birds of prey is based on morphological traits. In this study molecular data were used to assess phylogenetic relationships in this group of parasitic nematodes. The aim of the study was to evaluate a method for rapid species identification, to construct a phylogeny of paras...
Hedenström, Anders; Rosén, Mikael
Projected body frontal area is used when estimating the parasite drag of bird flight. We investigated the relationship between projected frontal area and body mass among passerine birds, and compared it with an equation based on waterfowl and raptors, which is used as default procedure in a widespread software package for flight performance calculations. The allometric equation based on waterfowl/raptors underestimates the frontal area compared to the passerine equation presented here. Conseq...
Clausager, I.; Nohr, H.
The paper is a review of the present knowledge on impacts of wind turbines on birds, requested by the Danish Ministry of the Environment and Energy. The main conclusions of the review are, that in nearly all the studies so far the numbers of birds recorded colliding with wind turbines have been limited. Some studies indicate that stationary (breeding) birds inside the wind turbine area in the short run habituate to wind turbines, especially the noise and visual impacts, and that the risk for collision becomes low. However, some of the few more long term studies indicate that a negative impact may occur in later generations of breeding birds. In some studies a disturbance effect on bird species, which temporarily stay inside a wind turbine area in order to forage or rest, is observed. The degree of impact is species-specific. An effect is typically recorded inside a zone of up to 250-800 m, with geese and waders as the most sensitive groups of birds. (author)
Full Text Available Our ideas are based on the following assumptions. Egg as a standalone system is formed within another system, which is the body of the female. Both systems are implemented on the basis of a common genetic code. In this regard, for example, the dendrogram constructed by morphological criteria eggs should be approximately equal to those constructed by other molecular or morphological criteria adult birds. It should be noted that the dendrogram show only the degree of genetic similarity of taxa, therefore, the identity of materials depends on the number of analyzed criteria and their quality, ie, they should be the backbone. The greater the number of system-features will be included in the analysis and in one other case, the like are dendrogram. In other cases, we will have a fragmentary similarity, which is also very important when dealing with controversial issues. The main message of our research was to figure out the eligibility of usage the morphological characteristics of eggs as additional information in taxonomy and phylogeny of birds. Our studies show that the shape parameters of bird eggs show a stable attachment to certain types of birds and complex traits are species-specific. Dendrogram and diagrams built by the quantitative value of these signs, exhibit significant similarity with the dendrogram constructed by morphological, comparative anatomy, paleontology and molecular criteria for adult birds. This suggests the possibility of using morphological parameters eggs as additional information in dealing with taxonomy and phylogeny of birds.
Vetterling, J M
Eight species representing 8 genera of gallinaceous birds were used: Alectoris graeca; Colinus virginianus; Coturnix coturnix; Gallus gallus; Meleagris gallopavo; Numidia meleagris; Pavo cristatus; Phasianus colchicus. Three week-old birds were dosed with sporulated oocysts of Eimeria tenella Beltsville strain. At 4, 24, 48, 72, 96, 120 and 144, and 168 hr after inoculation, 1-3 infected birds and uninoculated controls of each species were killed by cardiac exsanguination. Pieces of intestines were fixed and examined for stages of E. tenella as stained paraffin sections or indirect fluorescent antibody preparations. Oocyst counts were made in droppings collected for the first 6 days of the patent period. Sporozoites were found in the lamina propria of some birds of 5 species at 4 hr postinoculation, but no stages were found thereafter except in the breeds of G. gallus and A. gracea. At 144 and 168 hr postinoculation, a few macrogametes were found in the ceca of 2 A. gracea, but no oocysts were found in the feces. No statistical difference was found between the number of oocysts produced/bird in the breeds of G. gallus examined. It is evident from these observations the E. tenella did not complete its life cycle in several close phylogenetic relatives of G. gallus, even though in other studies this parasite was found to complete its life cycle in cell cultures derived from the same birds.
Robert M. Chipley
The idea for the Important Bird Area Program originated in a series of studies in the early 1980s conducted by BirdLife International. Recognizing that these studies could become a powerful tool for conservation, BirdLife International began an effort to identify and gather data regarding the most important areas for birds in Europe and to make this information...
... byproducts of poultry, game birds, or other birds from regions where highly pathogenic avian influenza (HPAI... THE UNITED STATES § 95.30 Restrictions on entry of products and byproducts of poultry, game birds, or other birds from regions where highly pathogenic avian influenza (HPAI) subtype H5N1 exists. (a...
Oleinik, Janaina; de Macedo, Jose Antonio Fernandes; Yuanjian, Wang Zufferey
Climate changes are expected to affect bird migration in several aspects including timing changes, breeding and migration orientation. The correlation analysis of several climate conditions (e.g. temperature, wind, humidity, etc) and bird migration trajectory is the key for explaining bird behavior during migration. Moreover, the resulting correlation can be used for predicting new bird behavior according to climate changes. In this paper we propose an integrated solution for correlating bird...
James G. Dickson; Richard N. Conner; J. Howard Williamson
Birds were censused annually from 4 250-x80-in transects in a young pine plantation from age to 2 to 17 to assess changes in the bird community.Bird abundance was low and the bird communitry was the least diverse when the pine plantation was sparsely vegetated at age 2. As the plantation developed rapidly into the shrub stage, the bird communitry became more abundant...
Bird strikes can lead to permanent deformations, sudden decrease of thrust, even engine failure during the flight. Bird strikes on rotating blades can also cause slices of birds hitting other parts which may lead to greater damages. Bird strikes cannot be completely avoided. However, reduction of bird impacting on jet engines can be achieved by suitable design and manufacturing, through the mathematical modelling, simulation analysis and practical experiment of jet engines.
Sauer, John R.
Conservation concerns, federal mandates to monitor birds, and citizen science programs have spawned a variety of surveys that collect information on bird populations. Unfortunately, all too frequently these surveys are poorly designed and use inappropriate counting methods. Some of the flawed approaches reflect a lack of understanding of statistical design; many ornithologists simply are not aware that many of our most entrenched counting methods (such as point counts) cannot appropriately be used in studies that compare densities of birds over space and time. It is likely that most of the readers of The Condor have participated in a bird population survey that has been criticized for poor sampling methods. For example, North American readers may be surprised to read in Bird Census Techniques that the North American Breeding Bird Survey 'is seriously flawed in its design,' and that 'Analysis of trends is impossible from points that are positioned along roads' (p. 109). Our conservation efforts are at risk if we do not acknowledge these concerns and improve our survey designs. Other surveys suffer from a lack of focus. In Bird Census Techniques, the authors emphasize that all surveys require clear statements of objectives and an understanding of appropriate survey designs to meet their objectives. Too often, we view survey design as the realm of ornithologists who know the life histories and logistical issues relevant to counting birds. This view reflects pure hubris: survey design is a collaboration between ornithologists, statisticians, and managers, in which goals based on management needs are met by applying statistical principles for design to the biological context of the species of interest. Poor survey design is often due to exclusion of some of these partners from survey development. Because ornithologists are too frequently unaware of these issues, books such as Bird Census Techniques take on added importance as manuals for educating ornithologists about
Benson, Roger B J; Starmer-Jones, Ethan; Close, Roger A; Walsh, Stig A
The bony labyrinth of vertebrates houses the semicircular canals. These sense rotational accelerations of the head and play an essential role in gaze stabilisation during locomotion. The sizes and shapes of the semicircular canals have hypothesised relationships to agility and locomotory modes in many groups, including birds, and a burgeoning palaeontological literature seeks to make ecological interpretations from the morphology of the labyrinth in extinct species. Rigorous tests of form-function relationships for the vestibular system are required to support these interpretations. We test the hypothesis that the lengths, streamlines and angles between the semicircular canals are related to body size, wing kinematics and flying style in birds. To do this, we applied geometric morphometrics and multivariate phylogenetic comparative methods to a dataset of 64 three-dimensional reconstructions of the endosseous labyrinth obtained using micro-computed tomography scanning of bird crania. A strong relationship between centroid size of the semicircular canals and body size indicates that larger birds have longer semicircular canals compared with their evolutionary relatives. Wing kinematics related to manoeuvrability (and quantified using the brachial index) explain a small additional portion of the variance in labyrinth size. We also find strong evidence for allometric shape change in the semicircular canals of birds, indicating that major aspects of the shape of the avian labyrinth are determined by spatial constraints. The avian braincase accommodates a large brain, a large eye and large semicircular canals compared with other tetrapods. Negative allometry of these structures means that the restriction of space within the braincase is intense in small birds. This may explain our observation that the angles between planes of the semicircular canals of birds deviate more strongly from orthogonality than those of mammals, and especially from agile, gliding and flying
Pancost, R. D.; Taylor, K. W.; Handley, L.; Huber, M.; Hollis, C. J.
The application of glycerol dialkyl glycerol tetraether (GDGT) based sea surface temperature proxies (eg TEX-86) has brought about major advances in our understanding of Cenozoic climate. However, TEX-86 approaches yield high temperatures at high latitudes during the EECO, and the resulting low latitudinal temperature gradients challenge our understanding of the physical climate system. We examine existing Paleogene TEX-86 data using three approaches. First, we have compared TEX-86 SST estimates to those derived from oxygen isotopes or Mg/Ca ratios of co-occurring well-preserved planktonic foraminifera. This reveals a strong correlation between foraminiferal SSTs and those obtained from linear TEX-86 calibrations, but the latter are systematically 4-6°C warmer. SSTs derived from non-linear TEX-86 calibrations are similar to inorganic SST estimates at high temperatures, but an offset still persists at lower temperatures. We have also compared TEX-86 SST estimates from marginal settings to associated continental mean air temperatures (MAT) derived from soil bacterial GDGTs (the MBT/CBT index). Again, the two approaches exhibit a strong correlation but with the former yielding 4-6°C warmer temperatures. Second, we have developed SST records from New Zealand, in order to place the EECO SSTs into a longer-term Paleogene context. Our temperature estimates indicate that sea floor and sea surface temperatures increased by ~10°C from late Paleocene to early Eocene times. Late Paleocene TEX-86-derived SSTs for the Canterbury Basin range from 18 to 23°C, consistent with coeval TEX-86 records from the Campbell Plateau and south Tasman Sea. During the EECO, TEX-86 derived SSTs from the Canterbury Basin and south Tasman Sea indicate tropical conditions, with temperatures peaking at 30-32°C at 50 Ma and then declining to 24-26°C into the Middle Eocene. MBT/CBT-derived MATs broadly parallel the SST trends, albeit with slightly lower values. These records suggest that
...-9BPP-L2] RIN 1018-AX06 Migratory Bird Hunting; Proposed Frameworks for Late-Season Migratory Bird... between real and perceived changes, and the absence of adequate experimental controls. Consequently, we... options and a study plan to evaluate the effect of the proposed action in achieving those objectives. It...
I. S. Mityay
Full Text Available Our ideas are based on the following assumptions. Egg as a standalone system is formed within another system, which is the body of the female. Both systems are implemented on the basis of a common genetic code. In this regard, for example, the dendrogram constructed by morphological criteria eggs should be approximately equal to those constructed by other molecular or morphological criteria adult birds. It should be noted that the dendrogram show only the degree of genetic similarity of taxa, therefore, the identity of materials depends on the number of analyzed criteria and their quality, ie, they should be the backbone. The greater the number of system-features will be included in the analysis and in one other case, the like are dendrogram. In other cases, we will have a fragmentary similarity, which is also very important when dealing with controversial issues. The main message of our research was to figure out the eligibility of usage the morphological characteristics of eggs as additional information in taxonomy and phylogeny of birds. Our studies show that the shape parameters of bird eggs show a stable attachment to certain types of birds and complex traits are species-specific. Dendrogram and diagrams built by the quantitative value of these signs, exhibit significant similarity with the dendrogram constructed by morphological, comparative anatomy, paleontology and molecular criteria for adult birds. This suggests the possibility of using morphological parameters eggs as additional information in dealing with taxonomy and phylogeny of birds. Keywords: oology, geometrical parameters of eggs, bird systematics
C. F. Dawber
Full Text Available Culturing studies and empirically based core top calibrations have been used to infer that elemental ratios in benthic foraminifera can be used as proxies to reconstruct past variations in bottom water temperature and saturation state (Δ [CO32−]. However the mechanisms linking elemental ratios to these parameters are poorly constrained. Here, we explore the environmental parameters influencing the incorporation of B, Li, Sr and Mg in Oridorsalis umbonatus in early Cenozoic sediments from Ocean Drilling Program Site 1209. We investigate the influence of middle Eocene variations in intermediate water Δ [CO32−] using relationships developed from core top samples. The fidelity of bottom water Δ[CO32−] reconstructions based on single element ratios is assessed by comparing the X/Ca-based reconstructions to each other and to carbon cycle proxy records (benthic foraminifera δ13C, organic carbon content, foraminifera dissolution indices, and a seawater δ18O reconstruction for Site 1209. Discrepancies in the reconstructed Δ[CO32−] values based on these different metal ratios suggest that there are still gaps in our understanding of the parameters influencing X/Ca and demonstrate that caution is required when interpreting palaeo-reconstructions that are derived from a single elemental ratio. The downcore record of O. umbonatus Mg/Ca does not exhibit any similarities with the Li/Ca, B/Ca and Sr/Ca records, suggesting that the environmental parameters influencing Mg/Ca may be different for this species, consistent with temperature as the strongest control on this elemental ratio. This hypothesis is supported by the coefficients of multiple linear regression models on published Mg/Ca data. An incomplete understanding of the controls on elemental incorporation into benthic foraminifera hinders our ability to confidently quantify changes in saturation state using single X/Ca reconstructions over a range of timescales.
Full Text Available Cetacea are secondarily aquatic amniotes that underwent their land-to-sea transition during the Eocene. Primitive forms, called archaeocetes, include five families with distinct degrees of adaptation to an aquatic life, swimming mode and abilities that remain difficult to estimate. The lifestyle of early cetaceans is investigated by analysis of microanatomical features in postcranial elements of archaeocetes. We document the internal structure of long bones, ribs and vertebrae in fifteen specimens belonging to the three more derived archaeocete families--Remingtonocetidae, Protocetidae, and Basilosauridae--using microtomography and virtual thin-sectioning. This enables us to discuss the osseous specializations observed in these taxa and to comment on their possible swimming behavior. All these taxa display bone mass increase (BMI in their ribs, which lack an open medullary cavity, and in their femora, whereas their vertebrae are essentially spongious. Humeri and femora show opposite trends in microanatomical specialization in the progressive independence of cetaceans from a terrestrial environment. Humeri change from very compact to spongious, which is in accordance with the progressive loss of propulsive role for the forelimbs, which were used instead for steering and stabilizing. Conversely, hind-limbs in basilosaurids became strongly reduced with no involvement in locomotion but display strong osteosclerosis in the femora. Our study confirms that Remingtonocetidae and Protocetidae were almost exclusively aquatic in locomotion for the taxa sampled, which probably were shallow water suspended swimmers. Basilosaurids display osseous specializations similar to those of modern cetaceans and are considered more active open-sea swimmers. This study highlights the strong need for homologous sections in comparative microanatomical studies, and the importance of combining information from several bones of the same taxon for improved functional
Stevens, Nancy J.; Hill, Robert V.; Al-Wosabi, Mohammed; Schulp, Anne; As-Saruri, Mustafa; Al-Nimey, Fuad; Jolley, Lea Ann; Schulp-Stuip, Yvonne; O'Connor, Patrick
During the Cenozoic, the Arabian Plate separated from continental Africa and assumed a closer geographical relationship with Eurasia. As such, the vertebrate fossil record of the Arabian Peninsula has great potential for documenting faunal interchanges that occurred as a result of such tectonic events, with a shift from a primarily Afro-Arabian fauna in the Palaeogene to a more cosmopolitan fauna in the Neogene. Understanding of the sequence and timing of this faunal interchange has long been hampered by a lack of palaeontological data. Recently recovered fossils from the Middle Eocene Kaninah Formation of Yemen constitute the earliest Palaeogene record of continental vertebrates from the Arabian Peninsula, thereby offering a rare glimpse at the region's post- -Cretaceous fauna. Here we describe fossil materials from the Kaninah Formation, a collection of dental and postcranial elements representing a mesoeucrocodylian crocodyliform of unclear affinities. The specimen exhibits ziphodont tooth morphology along with a biserial paravertebral shield and polygonal gastral osteoderms, consistent with certain mesoeucrocodylians (e.g., ziphodontan notosuchians). Yet the associated fragmentary anterior caudal vertebra, although badly abraded, preserves morphology suggestive of procoely. This vertebral type in combination with the dental and osteoderm morphology is much more taxonomically restrictive and consistent with the suite of characters exhibited by atoposaurids, a finding that would significantly extend that clade through the Cretaceous/Palaeogene boundary. Alternatively, given the relative paucity of information from the region during the Palaeogene, the combination of characteristics of the Kaninah crocodyliform may reflect a novel or poorly known form exhibiting previously unrecognised character mosaicism. We take a conservative approach, and refer the Kaninah specimen to Mesoeucrocodylia, Atoposauridae (?) pending discovery of more complete material. New fossils
Ferràndez-Cañadell, Carles; Briguglio, Antonino; Hohenegger, Johann; Wöger, Julia
In foraminifera, so-called “double tests” usually arise due to abnormal growth originating mainly from twinning, but may also be caused by irregularities in the early chambers and by regeneration after test injury that modifies the direction of growth. A fourth cause of double tests has only rarely been reported: the fusion of the tests of two adult individuals. We studied an early Eocene Nummulites double test consisting of two adult individuals that fused after an extended period of independent growth. The specimen was studied using computed tomography with micrometric resolution (micro-CT) that allowed bi- and three-dimensional visualization of the internal structure. Before fusion each individual test had 30–36 chambers, which, by comparison with growth rates in recent nummulitids, implies at least three months of independent growth. After fusion, the compound test grew in two spirals that fused after about one whorl and then continued in a single spiral. To fuse their tests, either adult individuals have to be forced to do so or the allorecognition (ability to distinguish between self and another individual) mechanisms must fail. A possible explanation for the merged Nummulites tests in this study is forced fusion in attached individuals after surviving ingestion and digestion by a metazoan. Alternatively, environmental stress could lead to a failure of allorecognition mechanisms and/or foraminiferal motility. Once fused, subsequent growth seems to be determined mainly by the relative orientation of individual tests. In any case, the frequency in which adult fusion occurs remains unknown. PMID:26166916
Leonardo M. Carneiro
Full Text Available Zeusdelphys complicatus is one of the most enigmatic metatherians from the Itaboraí Basin. The type and only known specimen was previously regarded as the upper dentition of Eobrasilia; an M4 of a new taxon; an M3 of a Kollpaniidae (now regarded as a group of “condylarths”; a probable M1 of an incertae sedis taxon; and as an M1 of a Protodidelphidae. Herein, we present a morphological review of the dental structures of Zeusdelphys complicatus, presenting new interpretations and comparing it with other North and South American taxa. We also perform a phylogenetic analysis in order to test the affinities of Zeusdelphys and the validity of most studied characters. The results recovered Zeusdelphys complicatus as more closely related to Hatcheritherium alpha than to any other metatherian. Glasbiidae were recovered as the sister lineage of Protodidelphidae within Didelphimorphia, as true marsupials. Ectocentrocristus was recovered as the sister taxon of Zeusdelphys + Hatcheritherium, as a Hatcheriformes. The analysis recovered this suborder as an independent lineage from Polydolopimorphia, being more closely related to “Alphadontidae”. The affinities with Protodidelphidae are a result of convergent evolution, as Zeusdelphys is more closely related to Hatcheritherium alpha from the Late Cretaceous of North America. The results support a North American origin for Hatcheriformes. The presence of strong sea-level lowstands and islands in the Caribbean Plate during the Late Cretaceous provide valid data to support a faunal interchange between Americas during the latest Late Cretaceous. Based on the results, Zeusdelphys represents a South American early Eocene surviving Hatcheriformes.
Carmichael, Matthew J.; Inglis, Gordon N.; Badger, Marcus P. S.; Naafs, B. David A.; Behrooz, Leila; Remmelzwaal, Serginio; Monteiro, Fanny M.; Rohrssen, Megan; Farnsworth, Alexander; Buss, Heather L.; Dickson, Alexander J.; Valdes, Paul J.; Lunt, Daniel J.; Pancost, Richard D.
The Paleocene-Eocene Thermal Maximum (PETM) hyperthermal, 56 million years ago (Ma), is the most dramatic example of abrupt Cenozoic global warming. During the PETM surface temperatures increased between 5 and 9 °C and the onset likely took < 20 kyr. The PETM provides a case study of the impacts of rapid global warming on the Earth system, including both hydrological and associated biogeochemical feedbacks, and proxy data from the PETM can provide constraints on changes in warm climate hydrology simulated by general circulation models (GCMs). In this paper, we provide a critical review of biological and geochemical signatures interpreted as direct or indirect indicators of hydrological change at the PETM, explore the importance of adopting multi-proxy approaches, and present a preliminary model-data comparison. Hydrological records complement those of temperature and indicate that the climatic response at the PETM was complex, with significant regional and temporal variability. This is further illustrated by the biogeochemical consequences of inferred changes in hydrology and, in fact, changes in precipitation and the biogeochemical consequences are often conflated in geochemical signatures. There is also strong evidence in many regions for changes in the episodic and/or intra-annual distribution of precipitation that has not widely been considered when comparing proxy data to GCM output. Crucially, GCM simulations indicate that the response of the hydrological cycle to the PETM was heterogeneous - some regions are associated with increased precipitation - evaporation (P - E), whilst others are characterised by a decrease. Interestingly, the majority of proxy data come from the regions where GCMs predict an increase in PETM precipitation. We propose that comparison of hydrological proxies to GCM output can be an important test of model skill, but this will be enhanced by further data from regions of model-simulated aridity and simulation of extreme precipitation
Cristianini T. Bergue
Full Text Available The Eocene-Pliocene deep sea ostracodes from the ODP site 744A (Kerguelen Plateau are herein studied under the taxonomic and paleoecologic aspects. 28 species are identified, being the genera Krithe, Cytherella and Dutoitella the most diversified. A faunal threshold was recorded in the Early Oligocene, which is tentatively explained under the knowledge of the paleoceanographical studies carried out not only in the Kerguelen Plateau but also in adjacent areas. The faunal turnover and variations in both richness and abundance possibly reflect the inception of psychrosphere and the influence of hydrological changes in the preservation of carapaces. Moreover, the influence of those changes on carbonate preservation is discussed as the cause of faunal impoverishment in the upper portion of the core.Ostracodes do intervalo Eoceno-Plioceno do sítio 744A do ODP (Platô Kerguelen são aqui estudados sob o aspecto taxonômico e paleoecológico. 28 espécies são identificadas, sendo os gêneros Krithe, Cytherella e Dutoitella os mais diversificados. Uma transição faunística registrada no Eoligoceno é investigada com base em estudos paleoceanográficos realizados no Platô Kerguelen e em áreas adjacentes. A transição e as variações de riqueza e abundância possivelmente refletem o estabelecimento da psicrosfera e mudanças hidrológicas associadas, na composição da fauna. Além disso, a influência destas mudanças na preservação do carbonato é discutida comopossível causa do empobrecimento da fauna na porção superior do testemunho.
Hou, Zhonge; Sket, Boris; Fišer, Cene; Li, Shuqiang
Current theory predicts that a shift to a new habitat would increase the rate of diversification, while as lineages evolve into multiple species, intensified competition would decrease the rate of diversification. We used Holarctic amphipods of the genus Gammarus to test this hypothesis. We sequenced four genes (5,088 bp) for 289 samples representing 115 Gammarus species. A phylogenetic analysis showed that Gammarus originated from the Tethyan region with a saline ancestry in the Paleocene, and later colonized the freshwater habitat in the Middle Eocene. Ancestral range reconstruction and diversification mode analysis combined with paleogeological and paleoclimatic evidence suggested that the habitat shift from saline to freshwater led to an increased diversification rate. The saline lineage of Gammarus dispersed to both sides of the Atlantic at 55 million years ago (Ma), because of the few barriers between the Tethys and the Atlantic, and diversified throughout its evolutionary history with a constant diversification rate [0.04 species per million years (sp/My)]. The freshwater Gammarus, however, underwent a rapid diversification phase (0.11 sp/My) until the Middle Miocene, and lineages successively diversified across Eurasia via vicariance process likely driven by changes of the Tethys and landmass. In particular, the freshwater Gammarus lacustris and Gammarus balcanicus lineages had a relatively high diversification shift, corresponding to the regression of the Paratethys Sea and the continentalization of Eurasian lands during the Miocene period. Subsequently (14 Ma), the diversification rate of the freshwater Gammarus decreased to 0.05 and again to 0.01 sp/My. The genus Gammarus provides an excellent aquatic case supporting the hypothesis that ecological opportunities promote diversification. PMID:21844362
Zimmer, M.; Cai, Y.; Corley, A.; Liang, J. A.; Powars, D.; Goldstein, S. L.; Kent, D. V.; Broecker, W. S.
The Paleocene-Eocene Thermal Maximum (PETM) was a global hyperthermal ( 5ºC warming) event marked by a rapid carbon isotope excursion (CIE) of >1‰ in the marine carbonate record (e.g. Zeebe et al. Nature Geoscience 2009). Possible explanations for the CIE include intrusion of a sill complex into organic carbonate (Aarnes et al. J. Geol. Soc. 2015), dissolution of methane hydrates (Thomas et al. Geology 2002), and a comet impact event (Schaller et al. Science 2016). Here we present new data across the PETM from the VirginiaDEQ-USGS Surprise Hill (SH) core, Northumberland Co., VA. We analyzed the Marlboro Clay, a thick, kaolinite-rich clay unit that marks the initiation of the PETM in the mid-Atlantic Coastal Plain of North America, as well as units above and below it. Bulk sediment records a δ13C excursion of approximately -5‰ across the CIE, while benthic foraminifera (Cibicidoides spp.) record a synchronous excursion of approximately -4.5‰. These results are consistent with other records from the New Jersey Coastal Plain (Makarova et al. Paleoceanography 2017). The excursion coincides with an increase in magnetic susceptibility, a decrease in bulk CaCO3 content, and an 2.5‰ decrease of δ18O in both the bulk sediment and benthic foraminifera of the SH core. Pb isotope analyses of the <63 μm fraction sediments indicate a unique provenance make-up for the Marlboro Clay. The results of the study thus indicate that PETM Marlboro Clay was not generated simply by intensified weathering of the same source area as the underlying Aquia Formation and overlying Nanjemoy Formation. Any hypothesis that aims to explain the mechanism that triggered the PETM must also account for the observed distinct provenance make-up of the Marlboro Clay.
Full Text Available We investigate the late Paleocene/early Eocene (PE climate using the coupled atmosphere-ocean-sea ice model ECHAM5/MPI-OM. The surface in our PE control simulation is on average 297 K warm and ice-free, despite a moderate atmospheric CO2 concentration of 560 ppm. Compared to a pre-industrial reference simulation (PR, low latitudes are 5 to 8 K warmer, while high latitudes are up to 40 K warmer. This high-latitude amplification is in line with proxy data, yet a comparison to sea surface temperature proxy data suggests that the Arctic surface temperatures are still too low in our PE simulation.
To identify the mechanisms that cause the PE-PR surface temperature differences, we fit two simple energy balance models to the ECHAM5/MPI-OM results. We find that about 2/3 of the PE-PR global mean surface temperature difference are caused by a smaller clear sky emissivity due to higher atmospheric CO2 and water vapour concentrations in PE compared to PR; 1/3 is due to a smaller planetary albedo. The reduction of the pole-to-equator temperature gradient in PE compared to PR is due to (1 the large high-latitude effect of the higher CO2 and water vapour concentrations in PE compared to PR, (2 the lower Antarctic orography, (3 the smaller surface albedo at high latitudes, and (4 longwave cloud radiative effects. Our results support the hypothesis that local radiative effects rather than increased meridional heat transports were responsible for the "equable" PE climate.
Larisa A Doguzhaeva
Full Text Available The Subclass Coleoidea (Class Cephalopoda accommodates the diverse present-day internally shelled cephalopod mollusks (Spirula, Sepia and octopuses, squids, Vampyroteuthis and also extinct internally shelled cephalopods. Recent Spirula represents a unique coleoid retaining shell structures, a narrow marginal siphuncle and globular protoconch that signify the ancestry of the subclass Coleoidea from the Paleozoic subclass Bactritoidea. This hypothesis has been recently supported by newly recorded diverse bactritoid-like coleoids from the Carboniferous of the USA, but prior to this study no fossil cephalopod indicative of an endochochleate branch with an origin independent from subclass Bactritoidea has been reported.Two orthoconic conchs were recovered from the Early Eocene of Seymour Island at the tip of the Antarctic Peninsula, Antarctica. They have loosely mineralized organic-rich chitin-compatible microlaminated shell walls and broadly expanded central siphuncles. The morphological, ultrustructural and chemical data were determined and characterized through comparisons with extant and extinct taxa using Scanning Electron Microscopy/Energy Dispersive Spectrometry (SEM/EDS.Our study presents the first evidence for an evolutionary lineage of internally shelled cephalopods with independent origin from Bactritoidea/Coleoidea, indicating convergent evolution with the subclass Coleoidea. A new subclass Paracoleoidea Doguzhaeva n. subcl. is established for accommodation of orthoconic cephalopods with the internal shell associated with a broadly expanded central siphuncle. Antarcticerida Doguzhaeva n. ord., Antarcticeratidae Doguzhaeva n. fam., Antarcticeras nordenskjoeldi Doguzhaeva n. gen., n. sp. are described within the subclass Paracoleoidea. The analysis of organic-rich shell preservation of A. nordenskjoeldi by use of SEM/EDS techniques revealed fossilization of hyposeptal cameral soft tissues. This suggests that a depositional environment
Gandolfo, María A; Hermsen, Elizabeth J
Radially symmetrical, five-winged fossil fruits from the highly diverse early Eocene Laguna del Hunco flora of Chubut Province, Patagonia, Argentina, are named, described and illustrated. The main goals are to assess the affinities of the fossils and to place them in an evolutionary, palaeoecological and biogeographic context. Specimens of fossil fruits were collected from the Tufolitas Laguna del Hunco. They were prepared, photographed and compared with similar extant and fossil fruits using published literature. Their structure was also evaluated by comparing them with that of modern Ceratopetalum (Cunoniaceae) fruits through examination of herbarium specimens. The Laguna del Hunco fossil fruits share the diagnostic features that characterize modern and fossil Ceratopetalum (symmetry, number of fruit wings, presence of a conspicuous floral nectary and overall venation pattern). The pattern of the minor wing (sepal) veins observed in the Patagonian fossil fruits is different from that of modern and previously described fossil Ceratopetalum fruits; therefore, a new fossil species is recognized. An apomorphy (absence of petals) suggests that the fossils belong within crown-group Ceratopetalum . The Patagonian fossil fruits are the oldest known record for Ceratopetalum . Because the affinities, provenance and age of the fossils are so well established, this new Ceratopetalum fossil species is an excellent candidate for use as a calibration point in divergence dating studies of the family Cunoniaceae. It represents the only record of Ceratopetalum outside Australasia, and further corroborates the biogeographic connection between the Laguna del Hunco flora and ancient and modern floras of the Australasian region. © The Author 2017. Published by Oxford University Press on behalf of the Annals of Botany Company.
Denis, Elizabeth H.; Pedentchouk, Nikolai; Schouten, Stefan; Pagani, Mark; Freeman, Katherine H.
Fire has been an important component of ecosystems on a range of spatial and temporal scales. Fire can affect vegetation distribution, the carbon cycle, and climate. The relationship between climate and fire is complex, in large part because of a key role of vegetation type. Here, we evaluate regional scale fire-climate relationships during a past global warming event, the Paleocene-Eocene Thermal Maximum (PETM), in order to understand how vegetation influenced the links between climate and fire occurrence in the Arctic region. To document concurrent changes in climate, vegetation, and fire occurrence, we evaluated biomarkers, including polycyclic aromatic hydrocarbons (PAHs), terpenoids, and alkanes, from the PETM interval at a marine depositional site (IODP site 302, the Lomonosov Ridge) in the Arctic Ocean. Biomarker, fossil, and isotope evidence from site 302 indicates that terrestrial vegetation changed during the PETM. The abundance of the C29n-alkanes, pollen, and the ratio of leaf-wax n-alkanes relative to diterpenoids all indicate that proportional contributions from angiosperm vegetation increased relative to that from gymnosperms. These changes accompanied increased moisture transport to the Arctic and higher temperatures, as recorded by previously published proxy records. We find that PAH abundances were elevated relative to total plant biomarkers throughout the PETM, and suggest that fire occurrence increased relative to plant productivity. The fact that fire frequency or prevalence may have increased during wetter Arctic conditions suggests that changes in fire occurrence were not a simple function of aridity, as is commonly conceived. Instead, we suggest that the climate-driven ecological shift to angiosperm-dominated vegetation was what led to increased fire occurrence. Potential increases in terrestrial plant biomass that arose from warm, wet, and high CO2 conditions were possibly attenuated by biomass burning associated with compositional changes
Doguzhaeva, Larisa A; Bengtson, Stefan; Reguero, Marcelo A; Mörs, Thomas
The Subclass Coleoidea (Class Cephalopoda) accommodates the diverse present-day internally shelled cephalopod mollusks (Spirula, Sepia and octopuses, squids, Vampyroteuthis) and also extinct internally shelled cephalopods. Recent Spirula represents a unique coleoid retaining shell structures, a narrow marginal siphuncle and globular protoconch that signify the ancestry of the subclass Coleoidea from the Paleozoic subclass Bactritoidea. This hypothesis has been recently supported by newly recorded diverse bactritoid-like coleoids from the Carboniferous of the USA, but prior to this study no fossil cephalopod indicative of an endochochleate branch with an origin independent from subclass Bactritoidea has been reported. Two orthoconic conchs were recovered from the Early Eocene of Seymour Island at the tip of the Antarctic Peninsula, Antarctica. They have loosely mineralized organic-rich chitin-compatible microlaminated shell walls and broadly expanded central siphuncles. The morphological, ultrustructural and chemical data were determined and characterized through comparisons with extant and extinct taxa using Scanning Electron Microscopy/Energy Dispersive Spectrometry (SEM/EDS). Our study presents the first evidence for an evolutionary lineage of internally shelled cephalopods with independent origin from Bactritoidea/Coleoidea, indicating convergent evolution with the subclass Coleoidea. A new subclass Paracoleoidea Doguzhaeva n. subcl. is established for accommodation of orthoconic cephalopods with the internal shell associated with a broadly expanded central siphuncle. Antarcticerida Doguzhaeva n. ord., Antarcticeratidae Doguzhaeva n. fam., Antarcticeras nordenskjoeldi Doguzhaeva n. gen., n. sp. are described within the subclass Paracoleoidea. The analysis of organic-rich shell preservation of A. nordenskjoeldi by use of SEM/EDS techniques revealed fossilization of hyposeptal cameral soft tissues. This suggests that a depositional environment favoring soft
Smith, J.J.; Hasiotis, S.T.; Kraus, M.J.; Woody, D.T.
Soil organisms, as recorded by trace fossils in paleosols of the Willwood Formation, Wyoming, show significant body-size reductions and increased abundances during the Paleocene-Eocene Thermal Maximum (PETM). Paleobotanical, paleopedologic, and oxygen isotope studies indicate high temperatures during the PETM and sharp declines in precipitation compared with late Paleocene estimates. Insect and oligochaete burrows increase in abundance during the PETM, suggesting longer periods of soil development and improved drainage conditions. Crayfish burrows and molluscan body fossils, abundant below and above the PETM interval, are significantly less abundant during the PETM, likely because of drier floodplain conditions and lower water tables. Burrow diameters of the most abundant ichnofossils are 30-46% smaller within the PETM interval. As burrow size is a proxy for body size, significant reductions in burrow diameter suggest that their tracemakers were smaller bodied. Smaller body sizes may have resulted from higher subsurface temperatures, lower soil moisture conditions, or nutritionally deficient vegetation in the high-CO2 atmosphere inferred for the PETM. Smaller soil fauna co-occur with dwarf mammal taxa during the PETM; thus, a common forcing mechanism may have selected for small size in both above- and below-ground terrestrial communities. We predict that soil fauna have already shown reductions in size over the last 150 years of increased atmospheric CO2 and surface temperatures or that they will exhibit this pattern over the next century. We retrodict also that soil fauna across the Permian-Triassic and Triassic-Jurassic boundary events show significant size decreases because of similar forcing mechanisms driven by rapid global warming.
Birdwell, Justin E.
Concentrations of the lanthanide series or rare earth elements and yttrium were determined for lacustrine oil shale samples from the Eocene Green River Formation in the Piceance Basin of Colorado and the Uinta Basin of Utah. Unprocessed oil shale, post-pyrolysis (spent) shale, and leached shale samples were examined to determine if oil-shale processing to generate oil or the remediation of retorted shale affects rare earth element concentrations. Results for unprocessed Green River oil shale samples were compared to data published in the literature on reference materials, such as chondritic meteorites, the North American shale composite, marine oil shale samples from two sites in northern Tibet, and mined rare earth element ores from the United States and China. The Green River oil shales had lower rare earth element concentrations (66.3 to 141.3 micrograms per gram, μg g-1) than are typical of material in the upper crust (approximately 170 μg g-1) and were also lower in rare earth elements relative to the North American shale composite (approximately 165 μg g-1). Adjusting for dilution of rare earth elements by organic matter does not account for the total difference between the oil shales and other crustal rocks. Europium anomalies for Green River oil shales from the Piceance Basin were slightly lower than those reported for the North American shale composite and upper crust. When compared to ores currently mined for rare earth elements, the concentrations in Green River oil shales are several orders of magnitude lower. Retorting Green River oil shales led to a slight enrichment of rare earth elements due to removal of organic matter. When concentrations in spent and leached samples were normalized to an original rock basis, concentrations were comparable to those of the raw shale, indicating that rare earth elements are conserved in processed oil shales.
Ksepka, Daniel T; Clarke, Julia A; DeVries, Thomas J; Urbina, Mario
We present the first detailed description of the giant Eocene penguin Icadyptes salasi. The species is characterized by a narrow skull with a hyper-elongate spear-like beak, a robust cervical column and a powerful flipper. The bony beak tip of Icadyptes is formed by fusion of several elements and is unique among penguins, differing markedly from previously described giant penguin beaks. Vascular canal patterning similar to that of boobies, frigatebirds and albatrosses suggests I. salasi may have had a thin, sheet-like rhamphotheca unlike the thick rugose rhamphotheca of modern penguins. Together, these features suggest a novel ecology for I. salasi, most likely involving the capture of larger prey items via spearing. As the first described giant penguin specimen to preserve a complete wing skeleton, the I. salasi holotype yields significant insight into the shape, proportions and orientation of the wing in giant penguins. In articulation, the forelimb of I. salasi is straighter, permitting less manus and antibrachium flexion, than previous depictions of giant penguin wings. Cross-sections of the humerus and ulna reveal a level of osteosclerosis equalling or surpassing that of extant penguins. Based on ontogenetic data from extant penguins and the morphology of the carpometacarpus of I. salasi, we infer the retention of a free alular phalanx in basal penguins. Previously, the status of this element in penguins was disputed. Differences in the proportions of the manual phalanges contribute to a more abruptly tapering wingtip in I. salasi compared with crown penguins. Fossils from Peru, including the I. salasi holotype specimen, document that penguins expanded to nearly the whole of their extant latitudinal range early in their evolutionary history and during one of the warmest intervals in the Cenozoic.
Ding, Lin; Xu, Qiang; Yue, Yahui; Wang, Houqi; Cai, Fulong; Li, Shun
Paleoelevation reconstruction using oxygen isotopes is making a significant contribution to understanding the Cenozoic uplift of the Himalayas and the Tibetan Plateau. This paper presents new oxygen and carbon isotopic compositions from well dated Tertiary paleosols, lacustrine calcareous carbonates, and marls from the Nianbo (60-54 Ma) and upper Pana Formations (51-48 Ma) of the Linzizong Group in the Linzhou (Penbo) Basin. The sediments of the Nianbo Formation, which are >180 m-thick, were deposited in alluvial fans, braided rivers, fan deltas, and on nearshore to offshore lacustrine settings, whereas those of the upper Pana Formation are >100 m-thick and are comprised predominantly of proximal alluvial fan and braided river deposits. Correlations between the lithofacies and stable isotopic compositions suggest that the basin was mainly a hydrologically open environment. It is confirmed that the δ18Oc and δ13Cc values from Nianbo and Pana Formations have not yet been reset by late-stage diagenesis based on petrographic examination, oxygen isotope of the fossil ostracodes, and tectonic deformation of strata. The paleoelevations are reconstructed using the corrected most negative paleosurface water δ18Opsw values. These imply that the Linzhou area had attained an elevation of 4500±400 m during the period of the Indo-Asian collision, i.e., achieved a near-present elevation, and may form an Andean-type mountain range stretching the Gangdese arc before collision. The Gangdese Mountains probably maintained high elevations since at least the Paleocene and could play a crucial role in the climate change in the interior of the Tibetan Plateau during the Early Cenozoic. The paleogeomorphic scenario of the Eocene Tibet is proposed to exist at two high mountains in excess of 4500 m that sandwiched a low elevation basin.
Smith, Jon J.; Hasiotis, Stephen T.; Kraus, Mary J.; Woody, Daniel T.
Soil organisms, as recorded by trace fossils in paleosols of the Willwood Formation, Wyoming, show significant body-size reductions and increased abundances during the Paleocene–Eocene Thermal Maximum (PETM). Paleobotanical, paleopedologic, and oxygen isotope studies indicate high temperatures during the PETM and sharp declines in precipitation compared with late Paleocene estimates. Insect and oligochaete burrows increase in abundance during the PETM, suggesting longer periods of soil development and improved drainage conditions. Crayfish burrows and molluscan body fossils, abundant below and above the PETM interval, are significantly less abundant during the PETM, likely because of drier floodplain conditions and lower water tables. Burrow diameters of the most abundant ichnofossils are 30–46% smaller within the PETM interval. As burrow size is a proxy for body size, significant reductions in burrow diameter suggest that their tracemakers were smaller bodied. Smaller body sizes may have resulted from higher subsurface temperatures, lower soil moisture conditions, or nutritionally deficient vegetation in the high-CO2 atmosphere inferred for the PETM. Smaller soil fauna co-occur with dwarf mammal taxa during the PETM; thus, a common forcing mechanism may have selected for small size in both above- and below-ground terrestrial communities. We predict that soil fauna have already shown reductions in size over the last 150 years of increased atmospheric CO2 and surface temperatures or that they will exhibit this pattern over the next century. We retrodict also that soil fauna across the Permian-Triassic and Triassic-Jurassic boundary events show significant size decreases because of similar forcing mechanisms driven by rapid global warming. PMID:19805060
Chun, C. O.; Delaney, M. L.; Zachos, J. C.
Manganese and Uranium enrichment factors (EF) relative to crustal averages from nine Deep Sea Drilling Project and Ocean Drilling Program sites are presented to examine paleo-redox changes across the Paleocene-Eocene Thermal Maximum (PETM). We use geochemical tracers to infer the bottom-water oxygen concentrations during this abrupt warming event. Determining redox changes will test the hypothesis that low oxygen concentrations caused the benthic foraminifera extinction event as well as contribute to our understanding of where the source of isotopically light carbon was released. The comparison of Mn EF before and after a reductive cleaning procedure allows us to determine the presence of Mn-oxides or Mn-carbonates. Mn EF representing Mn oxides and no U EF suggest oxygenated bottom-waters. Mn-carbonates and U EF suggest reducing conditions and an early diagenetic phase. We will present our results relative to the orbitally tuned age model of Site 1263. Mn EF range from 1-9 in Atlantic sites, 1-35 Southern Ocean sites, and 5-400 in Pacific sites. U EF range from 1-5 in Atlantic sites, 1-90 in Southern Ocean sites, and crustal averages EF =1 in Pacific sites. Our initial results indicate reducing conditions prior, during, and in the recovery of the PETM at intermediate depth sites in the Atlantic and Southern Ocean while the Pacific sites remain oxygenated. Reducing bottom- water conditions in the Atlantic and Southern Ocean may have contributed to the benthic foraminifera extinction event. Release of methane in the oceans oxidizes to carbon dioxide, depleting bottom-water oxygen concentrations. Because our sites in the Atlantic have the most reducing bottom-waters, we suggest the Atlantic Ocean could have been the source for isotopically light carbon release during the PETM.
McIntosh, Jennifer C.; Warwick, Peter D.; Martini, Anna M.; Osborn, Stephen G.
Thirty-six formation waters, gas, and microbial samples were collected and analyzed from natural gas and oil wells producing from the Paleocene to Eocene Wilcox Group coal beds and adjacent sandstones in north-central Louisiana, USA, to investigate the role hydrology plays on the generation and distribution of microbial methane. Major ion chemistry and Cl−Br relations of Wilcox Group formation waters suggest mixing of freshwater with halite-derived brines. High alkalinities (up to 47.8 meq/L), no detectable SO4, and elevated δ13C values of dissolved inorganic carbon (up to 20.5‰ Vienna Peedee belemnite [VPDB]) and CO2 (up to 17.67‰ VPDB) in the Wilcox Group coals and adjacent sandstones indicate the dominance of microbial methanogenesis. The δ13C and δD values of CH4, and carbon isotope fractionation of CO2 and CH4, suggest CO2 reduction is the major methanogenic pathway. Geochemical indicators for methanogenesis drop off significantly at chloride concentrations above ∼1.7 mol/L, suggesting that high salinities inhibit microbial activity at depths greater than ∼1.6 km. Formation waters in the Wilcox Group contain up to 1.6% modern carbon (A14C) to at least 1690 m depth; the covariance of δD values of co-produced H2O and CH4 indicate that the microbial methane was generated in situ with these Late Pleistocene or younger waters. The most enriched carbon isotope values for dissolved inorganic carbon (DIC) and CO2, and highest alkalinities, were detected in Wilcox Group sandstone reservoirs that were CO2 flooded in the 1980s for enhanced oil recovery, leading to the intriguing hypothesis that CO2 sequestration may actually enhance methanogenesis in organic-rich formations.
Quick, J.V.; Thomas, N.G.; Brogdon, L.D.; Jones, C.A.; Martin, T.S.
The results of a subsurface uranium favorability study of Tertiary rocks (late Eocene through Pliocene) in the Coastal Plain of South Texas are given. In ascending order, these rock units include the Yegua Formation, Jackson Group, Frio Clay, Catahoula Tuff, Oakville Sandstone, and Goliad Sand. The Vicksburg Group, Anahuac Formation, and Fleming Formation were not considered because they have unfavorable lithologies. The Yegua Formation, Jackson Group, Frio Clay, Catahoula Tuff, Oakville Sandstone, and Goliad Sand contain sandstones that may be favorable uranium hosts under certain environmental and structural conditions. All except the Yegua are known to contain ore-grade uranium deposits. Yegua and Jackson sandstones are found in strand plain-barrier bar systems that are aligned parallel to depositional and structural strike. These sands grade into shelf muds on the east, and lagoonal sediments updip toward the west. The lagoonal sediments in the Jackson are interrupted by dip-aligned fluvial systems. In both units, favorable areas are found in the lagoonal sands and in sands on the updip side of the strand-plain system. Favorable areas are also found along the margins of fluvial systems in the Jackson. The Frio and Catahoula consist of extensive alluvial-plain deposits. Favorable areas for uranium deposits are found along the margins of the paleo-channels where favorable structural features and numerous optimum sands are present. The Oakville and Goliad Formations consist of extensive continental deposits of fluvial sandstones. In large areas, these fluvial sandstones are multistoried channel sandstones that form very thick sandstone sequences. Favorable areas are found along the margins of the channel sequences. In the Goliad, favorable areas are also found on the updip margin of strand-plain sandstones where there are several sandstones of optimum thickness.
Quick, J.V.; Thomas, N.G.; Brogdon, L.D.; Jones, C.A.; Martin, T.S.
The results of a subsurface uranium favorability study of Tertiary rocks (late Eocene through Pliocene) in the Coastal Plain of South Texas are given. In ascending order, these rock units include the Yegua Formation, Jackson Group, Frio Clay, Catahoula Tuff, Oakville Sandstone, and Goliad Sand. The Vicksburg Group, Anahuac Formation, and Fleming Formation were not considered because they have unfavorable lithologies. The Yegua Formation, Jackson Group, Frio Clay, Catahoula Tuff, Oakville Sandstone, and Goliad Sand contain sandstones that may be favorable uranium hosts under certain environmental and structural conditions. All except the Yegua are known to contain ore-grade uranium deposits. Yegua and Jackson sandstones are found in strand plain-barrier bar systems that are aligned parallel to depositional and structural strike. These sands grade into shelf muds on the east, and lagoonal sediments updip toward the west. The lagoonal sediments in the Jackson are interrupted by dip-aligned fluvial systems. In both units, favorable areas are found in the lagoonal sands and in sands on the updip side of the strand-plain system. Favorable areas are also found along the margins of fluvial systems in the Jackson. The Frio and Catahoula consist of extensive alluvial-plain deposits. Favorable areas for uranium deposits are found along the margins of the paleo-channels where favorable structural features and numerous optimum sands are present. The Oakville and Goliad Formations consist of extensive continental deposits of fluvial sandstones. In large areas, these fluvial sandstones are multistoried channel sandstones that form very thick sandstone sequences. Favorable areas are found along the margins of the channel sequences. In the Goliad, favorable areas are also found on the updip margin of strand-plain sandstones where there are several sandstones of optimum thickness
Denis, E. H.; Pedentchouk, N.; Schouten, S.; Pagani, M.; Freeman, K. H.
Fire, an important component of ecosystems at a range of spatial and temporal scales, affects vegetation distribution, the carbon cycle, and climate. In turn, climate influences fuel composition (e.g., amount and type of vegetation), fuel availability (e.g., vegetation that can burn based on precipitation and temperature), and ignition sources (e.g., lightning). Climate studies predict increased wildfire activity in future decades, but mechanisms that control the relationship between climate and fire are complex. Reconstructing environmental conditions during past warming events (e.g., the Paleocene-Eocene Thermal Maximum (PETM)) will help elucidate climate-vegetation-fire relationships that are expressed over long durations (1,000 - 10,000 yrs). The abrupt global warming during the PETM dramatically altered vegetation and hydrologic patterns, and, possibly, fire occurrence. To investigate coincident changes in climate, vegetation, and fire occurrence, we studied biomarkers, including polycyclic aromatic hydrocarbons (PAHs), terpenoids, and alkanes from the PETM interval at IODP site 302 (the Lomonosov Ridge) in the Arctic Ocean. Both pollen and biomarker records indicate angiosperms abundance increased during the PETM relative to gymnosperms, reflecting a significant ecological shift to angiosperm-dominated vegetation. PAH abundances increased relative to plant biomarkers throughout the PETM, which suggests PAH production increased relative to plant productivity. Increased PAH production associated with the angiosperm vegetation shift indicates a greater prevalence of more fire-prone species. A time lag between increased moisture transport (based on published δD of n-alkanes data) to the Arctic and increased angiosperms and PAH production suggests wetter conditions, followed by increased air temperatures, favored angiosperms and combined to enhance fire occurrence.
Foreman, Brady Z; Heller, Paul L; Clementz, Mark T
Climate strongly affects the production of sediment from mountain catchments as well as its transport and deposition within adjacent sedimentary basins. However, identifying climatic influences on basin stratigraphy is complicated by nonlinearities, feedback loops, lag times, buffering and convergence among processes within the sediment routeing system. The Palaeocene/Eocene thermal maximum (PETM) arguably represents the most abrupt and dramatic instance of global warming in the Cenozoic era and has been proposed to be a geologic analogue for anthropogenic climate change. Here we evaluate the fluvial response in western Colorado to the PETM. Concomitant with the carbon isotope excursion marking the PETM we document a basin-wide shift to thick, multistoried, sheets of sandstone characterized by variable channel dimensions, dominance of upper flow regime sedimentary structures, and prevalent crevasse splay deposits. This progradation of coarse-grained lithofacies matches model predictions for rapid increases in sediment flux and discharge, instigated by regional vegetation overturn and enhanced monsoon precipitation. Yet the change in fluvial deposition persisted long after the approximately 200,000-year-long PETM with its increased carbon dioxide levels in the atmosphere, emphasizing the strong role the protracted transmission of catchment responses to distant depositional systems has in constructing large-scale basin stratigraphy. Our results, combined with evidence for increased dissolved loads and terrestrial clay export to world oceans, indicate that the transient hyper-greenhouse climate of the PETM may represent a major geomorphic 'system-clearing event', involving a global mobilization of dissolved and solid sediment loads on Earth's surface.
Prasad, V; Farooqui, A; Tripathi, S K M; Garg, R; Thakur, B
Equatorial rain forests that maintain a balance between speciation and extinction are hot-spots for studies of biodiversity. Western Ghats in southern India have gained attention due to high tropical biodiversity and endemism in their southern most area. We attempted to track the affinities of the pollen fl ora of the endemic plants of Western Ghat area within the fossil palynoflora of late Palaeocene-early Eocene (approximately 55-50 Ma) sedimentary deposits of western and northeastern Indian region. The study shows striking similarity of extant pollen with twenty eight most common fossil pollen taxa of the early Palaeogene. Widespread occurrences of coal and lignite deposits during early Palaeogene provide evidence of existence of well diversified rain forest community and swampy vegetation in the coastal low lying areas all along the western and northeastern margins of the Indian subcontinent. Prevalence of excessive humid climate during this period has been seen as a result of equatorial positioning of Indian subcontinent, superimposed by a long term global warming phase (PETM and EECO) during the early Palaeogene. The study presents clear evidence that highly diversifi ed equatorial rain forest vegetation once widespread in the Indian subcontinent during early Palaeogene times, are now restricted in a small area as a refugia in the southernmost part of the Western Ghat area. High precipitation and shorter periods of dry months seem to have provided suitable environment to sustain lineages of ancient tropical vegetation in this area of Western Ghats in spite of dramatic climatic changes subsequent to the post India-Asia collision and during the Quaternary and Recent times.
Baddouh, M'bark; Meyers, Stephen R.; Carroll, Alan R.; Beard, Brian L.; Johnson, Clark M.
The Green River Formation (GRF) provides one of the premier paleoclimate archives of the Early Eocene Climatic Optimum (∼50 Ma), representing the apex of the early Cenozoic greenhouse climate. Rhythmic lake-level variability expressed in the GRF has inspired numerous hypotheses for the behavior of the Eocene hydrologic cycle, including its linkage to astronomical forcing, solar variability, and the El Niño Southern Oscillation (ENSO). However, the lack of sufficient proxy data to document atmospheric water-mass transport and the geographic pattern of evaporation/precipitation/runoff has made it difficult to discriminate between different models for astronomical forcing. Variable 87Sr/86Sr ratios of bedrock that encompass the GRF provide an opportunity to reconstruct the spatial expression of the Eocene hydrologic cycle and its linkage to lake level. Here Sr isotope data from the Wilkins Peak Member, a rhythmic succession that has been demonstrated to record Milankovitch forcing of lake levels, indicate that high lake levels reflect an increased proportion of runoff from less radiogenic rocks west of the basin, eliminating a number of the existing astronomical-forcing hypotheses. The 87Sr/86Sr variability is consistent with a change in mean ENSO state, which is predicted by climate models to be linked to orbital-insolation. Thus, the 87Sr/86Sr data reveal a coupling of high frequency (ENSO) and low frequency (astronomical) climate variability, and also predict the existence of sizable astronomically-forced alpine snowpack during the last greenhouse climate. More broadly, this study demonstrates the utility of 87Sr/86Sr as a powerful tool for reconstructing the deep-time hydrologic cycle.
Dusel-Bacon, C.; Murphy, J.M.
We present an apatite fission-track (AFT) study of five plutonic rocks and seven metamorphic rocks across 310 km of the Yukon-Tanana Upland in east-central Alaska. Samples yielding ???40 Ma AFT ages and mean confined track lengths > 14 ??m with low standard deviations cooled rapidly from >120??C to 40 Ma suggest partial annealing and, therefore, lower maximum temperatures (???90-105??C). A few samples with single-grain ages of ???20 Ma apparently remained above ???50??C after initial cooling. Although the present geothermal gradient in the western Yukon-Tanana Upland is ???32??C/km, it could have been as high as 45??C/km during a widespread Eocene intraplate magmatic episode. Prior to rapid exhumation, samples with ???40 Ma AFT ages were >3.8-2.7 km deep and samples with >50 Ma AFT ages were >3.3-2.0 km deep. We calculate a 440-320 m/Ma minimum rate for exhumation of all samples during rapid cooling. Our AFT data, and data from rocks north of Fairbanks and from the Eielson deep test hole, indicate up to 3 km of post-40 Ma vertical displacement along known and inferred northeast-trending high-angle faults. The predominance of 40-50 Ma AFT ages throughout the Yukon-Tanana Upland indicates that, prior to the post-40 Ma relative uplift along some northeast-trending faults, rapid regional cooling and exhumation closely followed the Eocene extensional magmatism. We propose that Eocene magmatism and exhumation were somehow related to plate movements that produced regional-scale oroclinal rotation, northward translation of outboard terranes, major dextral strike-slip faulting, and subduction of an oceanic spreading ridge along the southern margin of Alaska.
Luciani, Valeria; D'Onofrio, Roberta; Dickens, Gerald R.; Wade, Bridget S.
At low latitude locations in the northern hemisphere, striking changes in the relative abundances and diversity of the two dominant planktic foraminifera genera, Morozovella and Acarinina, are known to have occurred close to the Early Eocene Climatic Optimum (EECO; 49-53 Ma). Lower Eocene carbonate-rich sediments at Ocean Drilling Program (ODP) Site 1263 were deposited on a bathymetric high (Walvis Ridge) at 40° S, and afford an opportunity to examine such planktic foraminiferal assemblage changes in a temperate southern hemisphere setting. We present here quantified counts of early Eocene planktic foraminiferal assemblages from Hole 1263B, along with bulk sediment stable isotope analyses and proxy measurements for carbonate dissolution. The bulk sediment δ13C record at Site 1263 resembles similar records generated elsewhere, such that known and inferred hyperthermal events can be readily identified. Although some carbonate dissolution has occurred, the well-preserved planktic foraminiferal assemblages mostly represent primary changes in environmental conditions. Our results document the permanent decrease in Morozovella abundance and increase in Acarinina abundance at the beginning of the EECO, although this switch occurred 165 kyr after that at low-latitude northern hemisphere locations. This suggests that unfavourable environmental conditions for morozovellids at the start of the EECO, such as sustained passage of a temperature threshold or other changes in surface waters, occurred at lower latitudes first. The remarkable turnover from Morozovella to Acarinina was widely geographically widespread, although the causal mechanism remains elusive. In addition, at Site 1263, we document the virtual disappearance within the EECO of the biserial chiloguembelinids, commonly considered as inhabiting intermediate water depths, and a reduction in abundance of the thermocline-dwelling subbotinids. We interpret these changes as signals of subsurface water properties
Full Text Available The Ypresian Cambay Shale Formation at Vastan and Mangrol lignite mines in Gujarat, western India, has yielded a rich vertebrate fauna with numerous taxa of European affinities. Here we report a new, approximately contemporary vertebrate assemblage from two fossiliferous layers in the nearby mine of Tadkeshwar. These layers have yielded a similar mammal fauna with the co-occurrence of the perissodactyl-like cambaytheriid Cambaytherium thewissi, the adapoid primates Marcgodinotius indicus and cf. Asiadapis cambayensis, and the hyaenodontid Indohyaenodon raoi. The presence of these species in both Vastan and Tadkeshwar mines and at different levels suggests that the deposits between the two major lignite seams represent a single land mammal age. Apart from the aforementioned species there is a new, smaller species of Cambaytherium, and a new genus and species of esthonychid tillodont. This fauna also contains the first large early Eocene vertebrates from India, including an unidentified Coryphodon-like pantodont, a dyrosaurid crocodyliform and a new giant madtsoiid snake. Among the Tadkeshwar vertebrates several taxa are of Gondwana affinities, such as Pelomedusoides turtles, dyrosaurids, and large madtsoiids, attesting that the early Eocene was a crucial period in India during which Laurasian taxa of European affinities co-existed with relict taxa from Gondwana before the India-Asia collision. Our results suggest that terrestrial faunas could have dispersed to or from Europe during episodes of contact between the Indian subcontinent and different island blocks along the northern margin of the Neotethys, such as the Kohistan–Ladakh island-arc system. Gondwana taxa might represent remnants of ghost lineages shared with Madagascar, which reached the Indian subcontinent during the late Cretaceous; alternatively they might have come from North Africa and passed along the southern margin of the Neotethys to reach the Indian subcontinent. These
Schouten, Stefan; Woltering, Martijn; Rijpstra, W. Irene C.; Sluijs, Appy; Brinkhuis, Henk; Sinninghe Damsté, Jaap S.
A study of upper Paleocene-lower Eocene (P-E) sediments deposited on the Lomonosov Ridge in the central Arctic Ocean reveals relatively high abundances of terrestrial biomarkers. These include dehydroabietane and simonellite derived from conifers (gymnosperms) and a tetra-aromatic triterpenoid derived from angiosperms. The relative percentage of the angiosperm biomarker of the summed angiosperm + conifer biomarkers was increased at the end of the Paleocene-Eocene thermal maximum (PETM), different when observed with pollen counts which showed a relative decrease in angiosperm pollen. Stable carbon isotopic analysis of these biomarkers shows that the negative carbon isotope excursion (CIE) during the PETM amounts to 3‰ for both conifer biomarkers, dehydroabietane and simonellite, comparable to the magnitude of the CIE inferred from marine carbonates, but significantly lower than the 4.5‰ of the terrestrial C 29n-alkane [M. Pagani, N. Pedentchouk, M. Huber, A. Sluijs, S. Schouten, H. Brinkhuis, J.S. Sinninghe Damsté, G.R. Dickens, and the IODP Expedition 302 Expedition Scientists (2006), Arctic's hydrology during global warming at the Paleocene-Eocene thermal maximum. Nature, 442, 671-675.], which is a compound sourced by both conifers and angiosperms. Conspicuously, the angiosperm-sourced aromatic triterpane shows a much larger CIE of 6‰ and suggests that angiosperms increased in their carbon isotopic fractionation during the PETM. Our results thus indicate that the 4.5‰ C 29n-alkane CIE reported previously represents the average CIE of conifers and angiosperms at this site and suggest that the large and variable CIE observed in terrestrial records may be partly explained by the variable contributions of conifers and angiosperms. The differential response in isotopic fractionation of angiosperms and conifers points to different physiological responses of these vegetation types to the rise in temperature, humidity, and greenhouse gases during the PETM.
Methner, Katharina; Mulch, Andreas; Teyssier, Christian; Wells, Michael L.; Cosca, Michael A.; Gottardi, Raphael; Gebelin, Aude; Chamberlain, C. Page
Metamorphic core complexes (MCCs) in the North American Cordillera reflect the effects of lithospheric extension and contribute to crustal adjustments both during and after a protracted subduction history along the Pacific plate margin. While the Miocene-to-recent history of most MCCs in the Great Basin, including the Raft River-Albion-Grouse Creek MCC, is well documented, early Cenozoic tectonic fabrics are commonly severely overprinted. We present stable isotope, geochronological (40Ar/39Ar), and microstructural data from the Raft River detachment shear zone. Hydrogen isotope ratios of syntectonic white mica (δ2Hms) from mylonitic quartzite within the shear zone are very low (−90‰ to −154‰, Vienna SMOW) and result from multiphase synkinematic interaction with surface-derived fluids. 40Ar/39Ar geochronology reveals Eocene (re)crystallization of white mica with δ2Hms ≥ −154‰ in quartzite mylonite of the western segment of the detachment system. These δ2Hms values are distinctively lower than in localities farther east (δ2Hms ≥ −125‰), where 40Ar/39Ar geochronological data indicate Miocene (18–15 Ma) extensional shearing and mylonitic fabric formation. These data indicate that very low δ2H surface-derived fluids penetrated the brittle-ductile transition as early as the mid-Eocene during a first phase of exhumation along a detachment rooted to the east. In the eastern part of the core complex, prominent top-to-the-east ductile shearing, mid-Miocene 40Ar/39Ar ages, and higher δ2H values of recrystallized white mica, indicate Miocene structural and isotopic overprinting of Eocene fabrics.
Westerhold, Thomas; Röhl, Ursula; Wilkens, Roy H.; Gingerich, Philip D.; Clyde, William C.; Wing, Scott L.; Bowen, Gabriel J.; Kraus, Mary J.
A consistent chronostratigraphic framework is required to understand the effect of major paleoclimate perturbations on both marine and terrestrial ecosystems. Transient global warming events in the early Eocene, at 56-54 Ma, show the impact of large-scale carbon input into the ocean-atmosphere system. Here we provide the first timescale synchronization of continental and marine deposits spanning the Paleocene-Eocene Thermal Maximum (PETM) and the interval just prior to the Eocene Thermal Maximum 2 (ETM-2). Cyclic variations in geochemical data come from continental drill cores of the Bighorn Basin Coring Project (BBCP, Wyoming, USA) and from marine deep-sea drilling deposits retrieved by the Ocean Drilling Program (ODP). Both are dominated by eccentricity-modulated precession cycles used to construct a common cyclostratigraphic framework. Integration of age models results in a revised astrochronology for the PETM in deep-sea records that is now generally consistent with independent 3He age models. The duration of the PETM is estimated at ˜ 200 kyr for the carbon isotope excursion and ˜ 120 kyr for the associated pelagic clay layer. A common terrestrial and marine age model shows a concurrent major change in marine and terrestrial biota ˜ 200 kyr before ETM-2. In the Bighorn Basin, the change is referred to as Biohorizon B and represents a period of significant mammalian turnover and immigration, separating the upper Haplomylus-Ectocion Range Zone from the Bunophorus Interval Zone and approximating the Wa-4-Wa-5 land mammal zone boundary. In sediments from ODP Site 1262 (Walvis Ridge), major changes in the biota at this time are documented by the radiation of a second generation of apical spine-bearing sphenolith species (e.g., S. radians and S. editus), the emergence of T. orthostylus, and the marked decline of D. multiradiatus.
Muijres, Florian T; Johansson, L Christoffer; Bowlin, Melissa S; Winter, York; Hedenström, Anders
Flight is one of the energetically most costly activities in the animal kingdom, suggesting that natural selection should work to optimize flight performance. The similar size and flight speed of birds and bats may therefore suggest convergent aerodynamic performance; alternatively, flight performance could be restricted by phylogenetic constraints. We test which of these scenarios fit to two measures of aerodynamic flight efficiency in two passerine bird species and two New World leaf-nosed bat species. Using time-resolved particle image velocimetry measurements of the wake of the animals flying in a wind tunnel, we derived the span efficiency, a metric for the efficiency of generating lift, and the lift-to-drag ratio, a metric for mechanical energetic flight efficiency. We show that the birds significantly outperform the bats in both metrics, which we ascribe to variation in aerodynamic function of body and wing upstroke: Bird bodies generated relatively more lift than bat bodies, resulting in a more uniform spanwise lift distribution and higher span efficiency. A likely explanation would be that the bat ears and nose leaf, associated with echolocation, disturb the flow over the body. During the upstroke, the birds retract their wings to make them aerodynamically inactive, while the membranous bat wings generate thrust and negative lift. Despite the differences in performance, the wake morphology of both birds and bats resemble the optimal wake for their respective lift-to-drag ratio regimes. This suggests that evolution has optimized performance relative to the respective conditions of birds and bats, but that maximum performance is possibly limited by phylogenetic constraints. Although ecological differences between birds and bats are subjected to many conspiring variables, the different aerodynamic flight efficiency for the bird and bat species studied here may help explain why birds typically fly faster, migrate more frequently and migrate longer distances
Florian T Muijres
Full Text Available Flight is one of the energetically most costly activities in the animal kingdom, suggesting that natural selection should work to optimize flight performance. The similar size and flight speed of birds and bats may therefore suggest convergent aerodynamic performance; alternatively, flight performance could be restricted by phylogenetic constraints. We test which of these scenarios fit to two measures of aerodynamic flight efficiency in two passerine bird species and two New World leaf-nosed bat species. Using time-resolved particle image velocimetry measurements of the wake of the animals flying in a wind tunnel, we derived the span efficiency, a metric for the efficiency of generating lift, and the lift-to-drag ratio, a metric for mechanical energetic flight efficiency. We show that the birds significantly outperform the bats in both metrics, which we ascribe to variation in aerodynamic function of body and wing upstroke: Bird bodies generated relatively more lift than bat bodies, resulting in a more uniform spanwise lift distribution and higher span efficiency. A likely explanation would be that the bat ears and nose leaf, associated with echolocation, disturb the flow over the body. During the upstroke, the birds retract their wings to make them aerodynamically inactive, while the membranous bat wings generate thrust and negative lift. Despite the differences in performance, the wake morphology of both birds and bats resemble the optimal wake for their respective lift-to-drag ratio regimes. This suggests that evolution has optimized performance relative to the respective conditions of birds and bats, but that maximum performance is possibly limited by phylogenetic constraints. Although ecological differences between birds and bats are subjected to many conspiring variables, the different aerodynamic flight efficiency for the bird and bat species studied here may help explain why birds typically fly faster, migrate more frequently and migrate