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Sample records for early triassic mass

  1. Lethally Hot Temperatures During the Early Triassic Greenhouse

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    Sun, Yadong; Joachimski, Michael M.; Wignall, Paul B.; Yan, Chunbo; Chen, Yanlong; Jiang, Haishui; Wang, Lina; Lai, Xulong

    2012-10-01

    Global warming is widely regarded to have played a contributing role in numerous past biotic crises. Here, we show that the end-Permian mass extinction coincided with a rapid temperature rise to exceptionally high values in the Early Triassic that were inimical to life in equatorial latitudes and suppressed ecosystem recovery. This was manifested in the loss of calcareous algae, the near-absence of fish in equatorial Tethys, and the dominance of small taxa of invertebrates during the thermal maxima. High temperatures drove most Early Triassic plants and animals out of equatorial terrestrial ecosystems and probably were a major cause of the end-Smithian crisis.

  2. Early archosauromorph remains from the Permo-Triassic Buena Vista Formation of north-eastern Uruguay.

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    Ezcurra, Martín D; Velozo, Pablo; Meneghel, Melitta; Piñeiro, Graciela

    2015-01-01

    The Permo-Triassic archosauromorph record is crucial to understand the impact of the Permo-Triassic mass extinction on the early evolution of the group and its subsequent dominance in Mesozoic terrestrial ecosystems. However, the Permo-Triassic archosauromorph record is still very poor in most continents and hampers the identification of global macroevolutionary patterns. Here we describe cranial and postcranial bones from the Permo-Triassic Buena Vista Formation of northeastern Uruguay that contribute to increase the meagre early archosauromorph record from South America. A basioccipital fused to both partial exoccipitals and three cervical vertebrae are assigned to Archosauromorpha based on apomorphies or a unique combination of characters. The archosauromorph remains of the Buena Vista Formation probably represent a multi-taxonomic assemblage composed of non-archosauriform archosauromorphs and a 'proterosuchid-grade' animal. This assemblage does not contribute in the discussion of a Late Permian or Early Triassic age for the Buena Vista Formation, but reinforces the broad palaeobiogeographic distribution of 'proterosuchid grade' diapsids in Permo-Triassic beds worldwide.

  3. Early archosauromorph remains from the Permo-Triassic Buena Vista Formation of north-eastern Uruguay

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    Martín D. Ezcurra

    2015-02-01

    Full Text Available The Permo-Triassic archosauromorph record is crucial to understand the impact of the Permo-Triassic mass extinction on the early evolution of the group and its subsequent dominance in Mesozoic terrestrial ecosystems. However, the Permo-Triassic archosauromorph record is still very poor in most continents and hampers the identification of global macroevolutionary patterns. Here we describe cranial and postcranial bones from the Permo-Triassic Buena Vista Formation of northeastern Uruguay that contribute to increase the meagre early archosauromorph record from South America. A basioccipital fused to both partial exoccipitals and three cervical vertebrae are assigned to Archosauromorpha based on apomorphies or a unique combination of characters. The archosauromorph remains of the Buena Vista Formation probably represent a multi-taxonomic assemblage composed of non-archosauriform archosauromorphs and a ‘proterosuchid-grade’ animal. This assemblage does not contribute in the discussion of a Late Permian or Early Triassic age for the Buena Vista Formation, but reinforces the broad palaeobiogeographic distribution of ‘proterosuchid grade’ diapsids in Permo-Triassic beds worldwide.

  4. Early Triassic marine biotic recovery: the predators' perspective.

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    Scheyer, Torsten M; Romano, Carlo; Jenks, Jim; Bucher, Hugo

    2014-01-01

    Examining the geological past of our planet allows us to study periods of severe climatic and biological crises and recoveries, biotic and abiotic ecosystem fluctuations, and faunal and floral turnovers through time. Furthermore, the recovery dynamics of large predators provide a key for evaluation of the pattern and tempo of ecosystem recovery because predators are interpreted to react most sensitively to environmental turbulences. The end-Permian mass extinction was the most severe crisis experienced by life on Earth, and the common paradigm persists that the biotic recovery from the extinction event was unusually slow and occurred in a step-wise manner, lasting up to eight to nine million years well into the early Middle Triassic (Anisian) in the oceans, and even longer in the terrestrial realm. Here we survey the global distribution and size spectra of Early Triassic and Anisian marine predatory vertebrates (fishes, amphibians and reptiles) to elucidate the height of trophic pyramids in the aftermath of the end-Permian event. The survey of body size was done by compiling maximum standard lengths for the bony fishes and some cartilaginous fishes, and total size (estimates) for the tetrapods. The distribution and size spectra of the latter are difficult to assess because of preservation artifacts and are thus mostly discussed qualitatively. The data nevertheless demonstrate that no significant size increase of predators is observable from the Early Triassic to the Anisian, as would be expected from the prolonged and stepwise trophic recovery model. The data further indicate that marine ecosystems characterized by multiple trophic levels existed from the earliest Early Triassic onwards. However, a major change in the taxonomic composition of predatory guilds occurred less than two million years after the end-Permian extinction event, in which a transition from fish/amphibian to fish/reptile-dominated higher trophic levels within ecosystems became apparent.

  5. Early Triassic marine biotic recovery: the predators' perspective.

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    Torsten M Scheyer

    Full Text Available Examining the geological past of our planet allows us to study periods of severe climatic and biological crises and recoveries, biotic and abiotic ecosystem fluctuations, and faunal and floral turnovers through time. Furthermore, the recovery dynamics of large predators provide a key for evaluation of the pattern and tempo of ecosystem recovery because predators are interpreted to react most sensitively to environmental turbulences. The end-Permian mass extinction was the most severe crisis experienced by life on Earth, and the common paradigm persists that the biotic recovery from the extinction event was unusually slow and occurred in a step-wise manner, lasting up to eight to nine million years well into the early Middle Triassic (Anisian in the oceans, and even longer in the terrestrial realm. Here we survey the global distribution and size spectra of Early Triassic and Anisian marine predatory vertebrates (fishes, amphibians and reptiles to elucidate the height of trophic pyramids in the aftermath of the end-Permian event. The survey of body size was done by compiling maximum standard lengths for the bony fishes and some cartilaginous fishes, and total size (estimates for the tetrapods. The distribution and size spectra of the latter are difficult to assess because of preservation artifacts and are thus mostly discussed qualitatively. The data nevertheless demonstrate that no significant size increase of predators is observable from the Early Triassic to the Anisian, as would be expected from the prolonged and stepwise trophic recovery model. The data further indicate that marine ecosystems characterized by multiple trophic levels existed from the earliest Early Triassic onwards. However, a major change in the taxonomic composition of predatory guilds occurred less than two million years after the end-Permian extinction event, in which a transition from fish/amphibian to fish/reptile-dominated higher trophic levels within ecosystems became

  6. Footprints pull origin and diversification of dinosaur stem lineage deep into Early Triassic.

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    Brusatte, Stephen L; Niedźwiedzki, Grzegorz; Butler, Richard J

    2011-04-07

    The ascent of dinosaurs in the Triassic is an exemplary evolutionary radiation, but the earliest phase of dinosaur history remains poorly understood. Body fossils of close dinosaur relatives are rare, but indicate that the dinosaur stem lineage (Dinosauromorpha) originated by the latest Anisian (ca 242-244 Ma). Here, we report footprints from the Early-Middle Triassic of Poland, stratigraphically well constrained and identified using a conservative synapomorphy-based approach, which shifts the origin of the dinosaur stem lineage back to the Early Olenekian (ca 249-251 Ma), approximately 5-9 Myr earlier than indicated by body fossils, earlier than demonstrated by previous footprint records, and just a few million years after the Permian/Triassic mass extinction (252.3 Ma). Dinosauromorph tracks are rare in all Polish assemblages, suggesting that these animals were minor faunal components. The oldest tracks are quadrupedal, a morphology uncommon among the earliest dinosauromorph body fossils, but bipedality and moderately large body size had arisen by the Early Anisian (ca 246 Ma). Integrating trace fossils and body fossils demonstrates that the rise of dinosaurs was a drawn-out affair, perhaps initiated during recovery from the Permo-Triassic extinction.

  7. Early Triassic alternative ecological states driven by anoxia, hyperthermals, and erosional pulses following the end-Permian mass extinction

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    Pietsch, C.; Petsios, E.; Bottjer, D. J.

    2015-12-01

    The end-Permian mass extinction, 252 million years ago, was the most devastating loss of biodiversity in Earth's history. Massive volcanic eruptions of the Siberian Traps and the concurrent burning of coal, carbonate, and evaporite deposits emplaced greenhouse and toxic gasses. Hyperthermal events of the surface ocean, up to 40°C, led to reduced gradient-driven ocean circulation which yielded extensive equatorial oxygen minimum zones. Today, anthropogenic greenhouse gas production is outpacing carbon input modeled for the end-Permian mass extinction, which suggests that modern ecosystems may yet experience a severe biotic crisis. The Early Triassic records the 5 million year aftermath of the end-Permian mass extinction and is often perceived as an interval of delayed recovery. We combined a new, high resolution carbon isotope record, sedimentological analysis, and paleoecological collections from the Italian Werfen Formation to fully integrate paleoenvironmental change with the benthic ecological response. We find that the marine ecosystem experienced additional community restructuring events due to subsequent hyperthermal events and pulses of erosion. The benthic microfauna and macrofauna both contributed to disaster communities that initially rebounded in the earliest Triassic. 'Disaster fauna' including microbialites, microconchids, foraminifera, and "flat clams" took advantage of anoxic conditions in the first ~500,000 years, dominating the benthic fauna. Later, in the re-oxygenated water column, opportunistic disaster groups were supplanted by a more diverse, mollusc-dominated benthic fauna and a complex ichnofauna. An extreme temperature run-up beginning in the Late Dienerian led to an additional hyperthermal event in the Late-Smithian which co-occurred with increased humidity and terrestrial run-off. Massive siliciclastic deposits replaced carbonate deposition which corresponds to the infaunalization of the benthic fauna. The disaster taxa dominated

  8. Recovery vs. Restructuring: Establishing Ecologic Patterns in Early and Middle Triassic Paleocommunities (Invited)

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    Fraiser, M.; Dineen, A.; Sheehan, P.

    2013-12-01

    Published data has been interpreted as indicating that marine ecological devastation following the end-Permian mass extinction was protracted and may have lasted 5 million years into the Middle Triassic (Anisian). However, a review of previous literature shows that understanding of biotic recovery is typically based on only a few components of the ecosystem, such as on taxonomic diversity, a single genus/phylum, or facies. Typically, paleocommunities are considered fully recovered when dominance and diversity are regained and normal ecosystem functioning has resumed. However, in addition to the biodiversity crash at the end of the Permian, taxonomic and ecologic structure also changed,with the extinction marking the faunal shift from brachiopod-rich Paleozoic Evolutionary Fauna (EF) to the mollusc-rich Modern EF. This suggests that the extreme reorganizational nature of the Triassic does not adhere to the standard definition of recovery, which is a return to previous conditions. Thus, we propose the term 'restructuring' to describe this interval, as Early and Middle Triassic communities might not exhibit the typical characteristics of a 'normal' Permian one. To more fully characterize Triassic ecologic restructuring, paleoecologists should take into account functional diversity and redundancy. We quantified functional richness and regularity in four different paleocommunities from classic Permian and Triassic sections. Functional richness was low in paleocommunities after the end-Permian mass extinction, but increased to high levels by the Middle Triassic. In contrast, functional regularity was low in the Middle Permian, but high in all the Triassic paleocommunities. The change from low to high functional regularity/redundancy at the P/T boundary may be a factor of the highly stressful Triassic environmental conditions (i.e. anoxia, hypercapnia), as high regularity in a community can boost survival in harsh environments. Parameters such as these will more

  9. Exceptional vertebrate biotas from the Triassic of China, and the expansion of marine ecosystems after the Permo-Triassic mass extinction

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    Benton, Michael J.; Zhang, Qiyue; Hu, Shixue; Chen, Zhong-Qiang; Wen, Wen; Liu, Jun; Huang, Jinyuan; Zhou, Changyong; Xie, Tao; Tong, Jinnan; Choo, Brian

    2013-10-01

    The Triassic was a time of turmoil, as life recovered from the most devastating of all mass extinctions, the Permo-Triassic event 252 million years ago. The Triassic marine rock succession of southwest China provides unique documentation of the recovery of marine life through a series of well dated, exceptionally preserved fossil assemblages in the Daye, Guanling, Zhuganpo, and Xiaowa formations. New work shows the richness of the faunas of fishes and reptiles, and that recovery of vertebrate faunas was delayed by harsh environmental conditions and then occurred rapidly in the Anisian. The key faunas of fishes and reptiles come from a limited area in eastern Yunnan and western Guizhou provinces, and these may be dated relative to shared stratigraphic units, and their palaeoenvironments reconstructed. The Luoping and Panxian biotas, both from the Guanling Formation, are dated as Anisian (Pelsonian) on the basis of conodonts and radiometric dates, the former being slightly older than the latter. The Xingyi biota is from the Zhuganpo Formation, and is Ladinian or early Carnian, while the Guanling biota is from the overlying Xiaowa Formation, dated as Carnian. The first three biotas include extensive benthos and burrowing in the sediments, and they were located in restricted basins close to shore. Further, even though the Luoping and Panxian biotas are of similar age, their faunas differ significantly, reflecting perhaps palaeogeographically isolated basins. Between the time of the Xingyi and Guanling biotas, there was a major transgression, and the Guanling biota is entirely different in character from the other three, being dominated by pelagic forms such as large floating crinoids attached to logs, very large ichthyosaurs and thalattosaurs, and pseudoplanktonic bivalves, with no benthos and no burrowing. Phylogenetic study of the fishes and marine reptiles shows apparently explosive diversification among 20 actinopterygian lineages very early in the Early Triassic

  10. Astronomical tuning of the end-Permian extinction and the Early Triassic Epoch of South China and Germany

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    Li, Mingsong; Ogg, James; Zhang, Yang; Huang, Chunju; Hinnov, Linda; Chen, Zhong-Qiang; Zou, Zhuoyan

    2016-05-01

    The timing of the end-Permian mass extinction and subsequent prolonged recovery during the Early Triassic Epoch can be established from astronomically controlled climate cycles recorded in continuous marine sedimentary sections. Astronomical-cycle tuning of spectral gamma-ray logs from biostratigraphically-constrained cyclic stratigraphy through marine sections at Meishan, Chaohu, Daxiakou and Guandao in South China yields an integrated time scale for the Early Triassic, which is consistent with scaling of magnetostratigraphy from climatic cycles in continental deposits of the Germanic Basin. The main marine mass extinction interval at Meishan is constrained to less than 40% of a 100-kyr (kilo-year) cycle (i.e., marine reptiles in the Mesozoic at Chaohu that are considered to represent a significant recovery of marine ecosystems did not appear until 4.7 myr (million years) after the end-Permian extinction. The durations of the Griesbachian, Dienerian, Smithian and Spathian substages, including the uncertainty in placement of widely used conodont biostratigraphic datums for their boundaries, are 1.4 ± 0.1, 0.6 ± 0.1, 1.7 ± 0.1 and 1.4 ± 0.1 myr, implying a total span for the Early Triassic of 5.1 ± 0.1 myr. Therefore, relative to an assigned 251.902 ± 0.024 Ma for the Permian-Triassic boundary from the Meishan GSSP, the ages for these substage boundaries are 250.5 ± 0.1 Ma for base Dienerian, 249.9 ± 0.1 Ma for base Smithian (base of Olenekian stage), 248.2 ± 0.1 Ma for base Spathian, and 246.8 ± 0.1 Ma for the base of the Anisian Stage. This astronomical-calibrated timescale provides rates for the recurrent carbon isotope excursions and for trends in sedimentation accumulation through the Early Triassic of studied sections in South China.

  11. A gigantic nothosaur (Reptilia: Sauropterygia) from the Middle Triassic of SW China and its implication for the Triassic biotic recovery

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    Liu, Jun; Hu, Shi-Xue; Rieppel, Olivier; Jiang, Da-Yong; Benton, Michael J.; Kelley, Neil P.; Aitchison, Jonathan C.; Zhou, Chang-Yong; Wen, Wen; Huang, Jin-Yuan; Xie, Tao; Lv, Tao

    2014-11-01

    The presence of gigantic apex predators in the eastern Panthalassic and western Tethyan oceans suggests that complex ecosystems in the sea had become re-established in these regions at least by the early Middle Triassic, after the Permian-Triassic mass extinction (PTME). However, it is not clear whether oceanic ecosystem recovery from the PTME was globally synchronous because of the apparent lack of such predators in the eastern Tethyan/western Panthalassic region prior to the Late Triassic. Here we report a gigantic nothosaur from the lower Middle Triassic of Luoping in southwest China (eastern Tethyan ocean), which possesses the largest known lower jaw among Triassic sauropterygians. Phylogenetic analysis suggests parallel evolution of gigantism in Triassic sauropterygians. Discovery of this gigantic apex predator, together with associated diverse marine reptiles and the complex food web, indicates global recovery of shallow marine ecosystems from PTME by the early Middle Triassic.

  12. A new stem group echinoid from the Triassic of China leads to a revised macroevolutionary history of echinoids during the end-Permian mass extinction

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    Thompson, Jeffrey R.; Hu, Shi-xue; Zhang, Qi-Yue; Petsios, Elizabeth; Cotton, Laura J.; Huang, Jin-Yuan; Zhou, Chang-yong; Wen, Wen; Bottjer, David J.

    2018-01-01

    The Permian-Triassic bottleneck has long been thought to have drastically altered the course of echinoid evolution, with the extinction of the entire echinoid stem group having taken place during the end-Permian mass extinction. The Early Triassic fossil record of echinoids is, however, sparse, and new fossils are paving the way for a revised interpretation of the evolutionary history of echinoids during the Permian-Triassic crisis and Early Mesozoic. A new species of echinoid, Yunnanechinus luopingensis n. sp. recovered from the Middle Triassic (Anisian) Luoping Biota fossil Lagerstätte of South China, displays morphologies that are not characteristic of the echinoid crown group. We have used phylogenetic analyses to further demonstrate that Yunnanechinus is not a member of the echinoid crown group. Thus a clade of stem group echinoids survived into the Middle Triassic, enduring the global crisis that characterized the end-Permian and Early Triassic. Therefore, stem group echinoids did not go extinct during the Palaeozoic, as previously thought, and appear to have coexisted with the echinoid crown group for at least 23 million years. Stem group echinoids thus exhibited the Lazarus effect during the latest Permian and Early Triassic, while crown group echinoids did not.

  13. The Sail-Backed Reptile Ctenosauriscus from the Latest Early Triassic of Germany and the Timing and Biogeography of the Early Archosaur Radiation

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    Butler, Richard J.; Brusatte, Stephen L.; Reich, Mike; Nesbitt, Sterling J.; Schoch, Rainer R.; Hornung, Jahn J.

    2011-01-01

    Background Archosaurs (birds, crocodilians and their extinct relatives including dinosaurs) dominated Mesozoic continental ecosystems from the Late Triassic onwards, and still form a major component of modern ecosystems (>10,000 species). The earliest diverse archosaur faunal assemblages are known from the Middle Triassic (c. 244 Ma), implying that the archosaur radiation began in the Early Triassic (252.3–247.2 Ma). Understanding of this radiation is currently limited by the poor early fossil record of the group in terms of skeletal remains. Methodology/Principal Findings We redescribe the anatomy and stratigraphic position of the type specimen of Ctenosauriscus koeneni (Huene), a sail-backed reptile from the Early Triassic (late Olenekian) Solling Formation of northern Germany that potentially represents the oldest known archosaur. We critically discuss previous biomechanical work on the ‘sail’ of Ctenosauriscus, which is formed by a series of elongated neural spines. In addition, we describe Ctenosauriscus-like postcranial material from the earliest Middle Triassic (early Anisian) Röt Formation of Waldhaus, southwestern Germany. Finally, we review the spatial and temporal distribution of the earliest archosaur fossils and their implications for understanding the dynamics of the archosaur radiation. Conclusions/Significance Comprehensive numerical phylogenetic analyses demonstrate that both Ctenosauriscus and the Waldhaus taxon are members of a monophyletic grouping of poposauroid archosaurs, Ctenosauriscidae, characterised by greatly elongated neural spines in the posterior cervical to anterior caudal vertebrae. The earliest archosaurs, including Ctenosauriscus, appear in the body fossil record just prior to the Olenekian/Anisian boundary (c. 248 Ma), less than 5 million years after the Permian–Triassic mass extinction. These earliest archosaur assemblages are dominated by ctenosauriscids, which were broadly distributed across northern Pangea and which

  14. The sail-backed reptile Ctenosauriscus from the latest Early Triassic of Germany and the timing and biogeography of the early archosaur radiation.

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    Richard J Butler

    Full Text Available BACKGROUND: Archosaurs (birds, crocodilians and their extinct relatives including dinosaurs dominated Mesozoic continental ecosystems from the Late Triassic onwards, and still form a major component of modern ecosystems (>10,000 species. The earliest diverse archosaur faunal assemblages are known from the Middle Triassic (c. 244 Ma, implying that the archosaur radiation began in the Early Triassic (252.3-247.2 Ma. Understanding of this radiation is currently limited by the poor early fossil record of the group in terms of skeletal remains. METHODOLOGY/PRINCIPAL FINDINGS: We redescribe the anatomy and stratigraphic position of the type specimen of Ctenosauriscus koeneni (Huene, a sail-backed reptile from the Early Triassic (late Olenekian Solling Formation of northern Germany that potentially represents the oldest known archosaur. We critically discuss previous biomechanical work on the 'sail' of Ctenosauriscus, which is formed by a series of elongated neural spines. In addition, we describe Ctenosauriscus-like postcranial material from the earliest Middle Triassic (early Anisian Röt Formation of Waldhaus, southwestern Germany. Finally, we review the spatial and temporal distribution of the earliest archosaur fossils and their implications for understanding the dynamics of the archosaur radiation. CONCLUSIONS/SIGNIFICANCE: Comprehensive numerical phylogenetic analyses demonstrate that both Ctenosauriscus and the Waldhaus taxon are members of a monophyletic grouping of poposauroid archosaurs, Ctenosauriscidae, characterised by greatly elongated neural spines in the posterior cervical to anterior caudal vertebrae. The earliest archosaurs, including Ctenosauriscus, appear in the body fossil record just prior to the Olenekian/Anisian boundary (c. 248 Ma, less than 5 million years after the Permian-Triassic mass extinction. These earliest archosaur assemblages are dominated by ctenosauriscids, which were broadly distributed across northern Pangea and

  15. The sail-backed reptile Ctenosauriscus from the latest Early Triassic of Germany and the timing and biogeography of the early archosaur radiation.

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    Butler, Richard J; Brusatte, Stephen L; Reich, Mike; Nesbitt, Sterling J; Schoch, Rainer R; Hornung, Jahn J

    2011-01-01

    Archosaurs (birds, crocodilians and their extinct relatives including dinosaurs) dominated Mesozoic continental ecosystems from the Late Triassic onwards, and still form a major component of modern ecosystems (>10,000 species). The earliest diverse archosaur faunal assemblages are known from the Middle Triassic (c. 244 Ma), implying that the archosaur radiation began in the Early Triassic (252.3-247.2 Ma). Understanding of this radiation is currently limited by the poor early fossil record of the group in terms of skeletal remains. We redescribe the anatomy and stratigraphic position of the type specimen of Ctenosauriscus koeneni (Huene), a sail-backed reptile from the Early Triassic (late Olenekian) Solling Formation of northern Germany that potentially represents the oldest known archosaur. We critically discuss previous biomechanical work on the 'sail' of Ctenosauriscus, which is formed by a series of elongated neural spines. In addition, we describe Ctenosauriscus-like postcranial material from the earliest Middle Triassic (early Anisian) Röt Formation of Waldhaus, southwestern Germany. Finally, we review the spatial and temporal distribution of the earliest archosaur fossils and their implications for understanding the dynamics of the archosaur radiation. Comprehensive numerical phylogenetic analyses demonstrate that both Ctenosauriscus and the Waldhaus taxon are members of a monophyletic grouping of poposauroid archosaurs, Ctenosauriscidae, characterised by greatly elongated neural spines in the posterior cervical to anterior caudal vertebrae. The earliest archosaurs, including Ctenosauriscus, appear in the body fossil record just prior to the Olenekian/Anisian boundary (c. 248 Ma), less than 5 million years after the Permian-Triassic mass extinction. These earliest archosaur assemblages are dominated by ctenosauriscids, which were broadly distributed across northern Pangea and which appear to have been the first global radiation of archosaurs.

  16. Resetting the evolution of marine reptiles at the Triassic-Jurassic boundary.

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    Thorne, Philippa M; Ruta, Marcello; Benton, Michael J

    2011-05-17

    Ichthyosaurs were important marine predators in the Early Jurassic, and an abundant and diverse component of Mesozoic marine ecosystems. Despite their ecological importance, however, the Early Jurassic species represent a reduced remnant of their former significance in the Triassic. Ichthyosaurs passed through an evolutionary bottleneck at, or close to, the Triassic-Jurassic boundary, which reduced their diversity to as few as three or four lineages. Diversity bounced back to some extent in the aftermath of the end-Triassic mass extinction, but disparity remained at less than one-tenth of pre-extinction levels, and never recovered. The group remained at low diversity and disparity for its final 100 Myr. The end-Triassic mass extinction had a previously unsuspected profound effect in resetting the evolution of apex marine predators of the Mesozoic.

  17. Synchrotron Reveals Early Triassic Odd Couple: Injured Amphibian and Aestivating Therapsid Share Burrow.

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    Vincent Fernandez

    Full Text Available Fossorialism is a beneficial adaptation for brooding, predator avoidance and protection from extreme climate. The abundance of fossilised burrow casts from the Early Triassic of southern Africa is viewed as a behavioural response by many tetrapods to the harsh conditions following the Permo-Triassic mass-extinction event. However, scarcity of vertebrate remains associated with these burrows leaves many ecological questions unanswered. Synchrotron scanning of a lithified burrow cast from the Early Triassic of the Karoo unveiled a unique mixed-species association: an injured temnospondyl amphibian (Broomistega that sheltered in a burrow occupied by an aestivating therapsid (Thrinaxodon. The discovery of this rare rhinesuchid represents the first occurrence in the fossil record of a temnospondyl in a burrow. The amphibian skeleton shows signs of a crushing trauma with partially healed fractures on several consecutive ribs. The presence of a relatively large intruder in what is interpreted to be a Thrinaxodon burrow implies that the therapsid tolerated the amphibian's presence. Among possible explanations for such unlikely cohabitation, Thrinaxodon aestivation is most plausible, an interpretation supported by the numerous Thrinaxodon specimens fossilised in curled-up postures. Recent advances in synchrotron imaging have enabled visualization of the contents of burrow casts, thus providing a novel tool to elucidate not only anatomy but also ecology and biology of ancient tetrapods.

  18. Discovery of Early Triassic conodonts in western Gangdisê and the establishment of the Tangnale Formation

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    ZHENG; YouYe; XU; RongKe; WANG; ChengYuan; MA; GuoTao; LAI; XuLong; YE; DeJin; CAO; Liang; LIANG; JiWei

    2007-01-01

    Most geologists believe that there are no Early and Middle Triassic strata in the W. Gandisê stratigraphic subregion, but the present authors have found Early Triassic conodonts for the first time in the Shiquanhe area, including five conodonts genera (Form genera): Pachycladina, Neohindeodella, Cornudina, Hadrodontina and Hibbardella sp. etc. Then we affirm that Early Triassic deposits exist in the Gandisê stratigraphic subregion, and establish the Tangnale Formation. The conclusion is new important complementary basal data for Triassic stratigraphy division of Gangdisê, reconstructing palaogeography and studying Gangdisê from Paleozoic to Mesozoic island-arc evolution and transition.

  19. Atmospheric carbon injection linked to end-Triassic mass extinction.

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    Ruhl, Micha; Bonis, Nina R; Reichart, Gert-Jan; Sinninghe Damsté, Jaap S; Kürschner, Wolfram M

    2011-07-22

    The end-Triassic mass extinction (~201.4 million years ago), marked by terrestrial ecosystem turnover and up to ~50% loss in marine biodiversity, has been attributed to intensified volcanic activity during the break-up of Pangaea. Here, we present compound-specific carbon-isotope data of long-chain n-alkanes derived from waxes of land plants, showing a ~8.5 per mil negative excursion, coincident with the extinction interval. These data indicate strong carbon-13 depletion of the end-Triassic atmosphere, within only 10,000 to 20,000 years. The magnitude and rate of this carbon-cycle disruption can be explained by the injection of at least ~12 × 10(3) gigatons of isotopically depleted carbon as methane into the atmosphere. Concurrent vegetation changes reflect strong warming and an enhanced hydrological cycle. Hence, end-Triassic events are robustly linked to methane-derived massive carbon release and associated climate change.

  20. Atmospheric Carbon Injection Linked to End-Triassic Mass Extinction

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    Ruhl, M.; Bonis, N.R.; Reichart, G.J.; Sinninghe Damsté, J.S.; Kürschner, W.M.

    2011-01-01

    The end-Triassic mass extinction (similar to 201.4 million years ago), marked by terrestrial ecosystem turnover and up to similar to 50% loss in marine biodiversity, has been attributed to intensified volcanic activity during the break-up of Pangaea. Here, we present compound-specific carbon-isotope

  1. The Triassic dicynodont Kombuisia (Synapsida, Anomodontia) from Antarctica, a refuge from the terrestrial Permian-Triassic mass extinction.

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    Fröbisch, Jörg; Angielczyk, Kenneth D; Sidor, Christian A

    2010-02-01

    Fossils from the central Transantarctic Mountains in Antarctica are referred to a new species of the Triassic genus Kombuisia, one of four dicynodont lineages known to survive the end-Permian mass extinction. The specimens show a unique combination of characters only present in this genus, but the new species can be distinguished from the type species of the genus, Kombuisia frerensis, by the presence of a reduced but slit-like pineal foramen and the lack of contact between the postorbitals. Although incomplete, the Antarctic specimens are significant because Kombuisia was previously known only from the South African Karoo Basin and the new specimens extend the taxon's biogeographic range to a wider portion of southern Pangaea. In addition, the new finds extend the known stratigraphic range of Kombuisia from the Middle Triassic subzone B of the Cynognathus Assemblage Zone into rocks that are equivalent in age to the Lower Triassic Lystrosaurus Assemblage Zone, shortening the proposed ghost lineage of this taxon. Most importantly, the occurrence of Kombuisia and Lystrosaurus mccaigi in the Lower Triassic of Antarctica suggests that this area served as a refuge from some of the effects of the end-Permian extinction. The composition of the lower Fremouw Formation fauna implies a community structure similar to that of the ecologically anomalous Lystrosaurus Assemblage Zone of South Africa, providing additional evidence for widespread ecological disturbance in the extinction's aftermath.

  2. Whirling in the late Permian: ancestral Gyrinidae show early radiation of beetles before Permian-Triassic mass extinction.

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    Yan, Evgeny V; Beutel, Rolf G; Lawrence, John F

    2018-03-16

    Gyrinidae are a charismatic group of highly specialized beetles, adapted for a unique lifestyle of swimming on the water surface. They prey on drowning insects and other small arthropods caught in the surface film. Studies based on morphological and molecular data suggest that gyrinids were the first branch splitting off in Adephaga, the second largest suborder of beetles. Despite its basal position within this lineage and a very peculiar morphology, earliest Gyrinidae were recorded not earlier than from the Upper Triassic. Tunguskagyrus. with the single species Tunguskagyrus planus is described from Late Permian deposits of the Anakit area in Middle Siberia. The genus is assigned to the stemgroup of Gyrinidae, thus shifting back the minimum age of this taxon considerably: Tunguskagyrus demonstrates 250 million years of evolutionary stability for a very specialized lifestyle, with a number of key apomorphies characteristic for these epineuston predators and scavengers, but also with some preserved ancestral features not found in extant members of the family. It also implies that major splitting events in this suborder and in crown group Coleoptera had already occurred in the Permian. Gyrinidae and especially aquatic groups of Dytiscoidea flourished in the Mesozoic (for example Coptoclavidae and Dytiscidae) and most survive until the present day, despite the dramatic "Great Dying" - Permian-Triassic mass extinction, which took place shortly (in geological terms) after the time when Tunguskagyrus lived. Tunguskagyrus confirms a Permian origin of Adephaga, which was recently suggested by phylogenetic "tip-dating" analysis including both fossil and Recent gyrinids. This also confirms that main splitting events leading to the "modern" lineages of beetles took place before the Permian-Triassic mass extinction. Tunguskagyrus shows that Gyrinidae became adapted to swimming on the water surface long before Mesozoic invasions of the aquatic environment took place

  3. Early Triassic development of a foreland basin in the Canadian high Arctic: Implications for a Pangean Rim of Fire

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    Hadlari, Thomas; Dewing, Keith; Matthews, William A.; Alonso-Torres, Daniel; Midwinter, Derrick

    2018-06-01

    Following the amalgamation of Laurasia and Gondwana to form Pangea, some Triassic tectonic models show an encircling arc system called the "Pangean Rim of Fire". Here we show that the stratigraphy and Early Triassic detrital zircon provenance of the Sverdrup Basin in the Canadian Arctic is most consistent with deposition in a retro-arc foreland basin. Late Permian and Early Triassic volcanism was accompanied by relatively high rates of subsidence leading to a starved basin with volcanic input from a magmatic arc to the northwest. The mostly starved basin persisted through the Middle and Late Triassic with nearly continuous input of volcanic ash recorded as bentonites on the northwestern edge of the basin. In the latest Triassic it is interpreted that decreasing subsidence and a significant influx of sand-grade sediment when the arc was exhumed led to filling of the basin at the end of an orogenic cycle. Combined with other hints of Early Triassic arc activity along the western margin of Laurentia we propose that the Pangean Rim of Fire configuration spanned the entire Triassic. This proposed configuration represents the ring of external subduction zones that some models suggest are necessary for the breakup of supercontinents such as Pangea.

  4. High precision time calibration of the Permo-Triassic boundary mass extinction by U-Pb geochronology

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    Baresel, Björn; Bucher, Hugo; Brosse, Morgane; Schaltegger, Urs

    2014-05-01

    U-Pb dating using Chemical Abrasion, Isotope Dilution Thermal Ionization Mass Spectrometry (CA-ID-TIMS) is the analytical method of choice for geochronologists, who are seeking highest temporal resolution and a high degree of accuracy for single grains of zircon. The use of double-isotope tracer solutions, cross-calibrated and assessed in different EARTHTIME labs, coinciding with the reassessment of the uranium decay constants and further improvements in ion counting technology led to unprecedented precision better than 0.1% for single grain, and 0.05% for population ages, respectively. These analytical innovations now allow calibrating magmatic and biological timescales at resolution adequate for both groups of processes. To construct a revised and high resolution calibrated time scale for the Permian-Triassic boundary (PTB) we use (i) high-precision U-Pb zircon age determinations of a unique succession of volcanic ash beds interbedded with shallow to deep water fossiliferous sediments in the Nanpanjiang Basin (South China) combined with (ii) accurate quantitative biochronology based on ammonoids and conodonts and (iii) carbon isotope excursions across the PTB. Using these alignments allows (i) positioning the PTB in different depositional environments and (ii) solving age/stratigraphic contradictions generated by the index, water depth-controlled conodont Hindeodus parvus, whose diachronous first occurrences are arbitrarily used for placing the base of the Triassic. This new age framework provides the basis for a combined calibration of chemostratigraphic records with high-resolution biochronozones of the Late Permian and Early Triassic. Besides the general improvement of the radio-isotopic calibration of the PTB at the ±100 ka level, this will also lead to a better understanding of cause and effect relations involved in this mass extinction.

  5. Timing of global regression and microbial bloom linked with the Permian-Triassic boundary mass extinction: implications for driving mechanisms

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    Baresel, Bjoern; Bucher, Hugo; Bagherpour, Borhan; Brosse, Morgane; Guodun, Kuang; Schaltegger, Urs

    2017-04-01

    High-precision U-Pb dating of single-zircon crystals by chemical abrasion-isotope dilution-thermal ionization mass spectrometry (CA-ID-TIMS) is applied to volcanic beds that are intercalated in sedimentary sequences across the Permian-Triassic boundary (PTB). By assuming that the zircon crystallization age closely approximate that of the volcanic eruption and subsequent deposition, U-Pb zircon geochronology is the preferred approach for dating abiotic and biotic events, such as the formational PTB and the Permian-Triassic boundary mass extinction (PTBME). We will present new U-Pb zircon dates for a series of volcanic ash beds in shallow-marine Permian-Triassic sections in the Nanpanjiang Basin, South China. These high-resolution U-Pb dates indicate a duration of 90 ± 38 kyr for the Permian sedimentary hiatus and a duration of 13 ± 57 kyr for the overlying Triassic microbial limestone in the shallow water settings of the Nanpanjiang pull apart Basin. The age and duration of the hiatus coincides with the formational PTB and the extinction interval in the Meishan Global Stratotype Section and Point, thus strongly supporting a glacio-eustatic regression, which best explains the genesis of the worldwide hiatus straddling the PTB in shallow water records. In adjacent deep marine troughs, rates of sediment accumulation display a six-fold decrease across the PTB compatible with a dryer and cooler climate during the Griesbachian as indicated by terrestrial plants. Our model of the PTBME hinges on the synchronicity of the hiatus with the onset of the Siberian Traps volcanism. This early eruptive phase likely released sulfur-rich volatiles into the stratosphere, thus simultaneously eliciting a short-lived ice age responsible for the global regression and a brief but intense acidification. Abrupt cooling, shrunk habitats on shelves and acidification may all have synergistically triggered the PTBME. Subsequently, the build-up of volcanic CO2 induced this transient cool

  6. Timing of global regression and microbial bloom linked with the Permian-Triassic boundary mass extinction: implications for driving mechanisms.

    Science.gov (United States)

    Baresel, Björn; Bucher, Hugo; Bagherpour, Borhan; Brosse, Morgane; Guodun, Kuang; Schaltegger, Urs

    2017-03-06

    New high-resolution U-Pb dates indicate a duration of 89 ± 38 kyr for the Permian hiatus and of 14 ± 57 kyr for the overlying Triassic microbial limestone in shallow water settings of the Nanpanjiang Basin, South China. The age and duration of the hiatus coincides with the Permian-Triassic boundary (PTB) and the extinction interval in the Meishan Global Stratotype Section and Point, and strongly supports a glacio-eustatic regression, which best explains the genesis of the worldwide hiatus straddling the PTB in shallow water records. In adjacent deep marine troughs, rates of sediment accumulation display a six-fold decrease across the PTB compatible with a dryer and cooler climate as indicated by terrestrial plants. Our model of the Permian-Triassic boundary mass extinction (PTBME) hinges on the synchronicity of the hiatus with the onset of the Siberian Traps volcanism. This early eruptive phase released sulfur-rich volatiles into the stratosphere, thus simultaneously eliciting a short-lived ice age responsible for the global regression and a brief but intense acidification. Abrupt cooling, shrunk habitats on shelves and acidification may all have synergistically triggered the PTBME. Subsequently, the build-up of volcanic CO 2 induced a transient cool climate whose early phase saw the deposition of the microbial limestone.

  7. A NEW SAURICHTHYS (ACTINOPTERYGII FROM THE SPATHIAN (EARLY TRIASSIC OF CHAOHU (ANHUI PROVINCE, CHINA

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    ANDREA TINTORI

    2014-07-01

    Full Text Available A new species of Saurichthys, S. majiashanensis , is described. It comes from a series of marine vertebrate-bearing beds of the upper member of the Nanlinghu Formation outcropping in the large quarry at the top of Majiashan (Majia Hill near Chaohu (Hefei City, Anhui Province, China. Its age is Middle Spathian (Olenekian, Early Triassic. This new species deeply differs from the several pre-Spathian species of Saurichthys mainly for having only two longitudinal scale rows together with a reduced grid-like structure for the neural elements in the vertebral column. Further derived characters are in the endoskeleton of the dorsal and anal fin, where radials articulate only to anterior lepidotrichia, the posterior ones being supported by the first scale from the caudal pedicle mid-dorsal and mid-ventral rows, deeply imbedded in the body. In addition, the haemal spines of the caudal region are much enlarged and reversed, with their distal parts pointing forwards. Though the skull is lacking, postcranial characters are enough to justify the erection of a new species. This new Saurichthys species, together with other few actinopterygians, can be considered as the oldest assemblage of the Triassic Middle Fish Fauna, which bloomed probably in the early Anisian and widespread especially all over the Tethys for the Middle Triassic and at least the Carnian in the Late Triassic. This new fish assemblage, together with some of the oldest marine reptiles, is considered as the beginning of the actual Triassic recovery among marine vertebrates.

  8. Long-term oceanic changes prior the end-Triassic mass extinction

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    Clémence, Marie-Emilie; Mette, Wolfgang; Thibault, Nicolas; Korte, Christoph

    2014-05-01

    A number of potential causes and kill mechanisms have been proposed for the end-Triassic mass extinction such as palaeoclimatic and sea-level variations, massive volcanism and ocean acidification. Recent analysis of the stomatal index and density of fossil leaves and geochemical research on pedogenic carbonate nodules are suggestive of rising atmospheric CO2 concentration and fluctuating climate in the Rhaetian. It seems therefore probable that the end-Triassic event was preceded by large climatic fluctuations and environmental perturbations in the Rhaetian which might have partly affected the composition and diversity of the terrestrial and marine biota prior to the end-Triassic interval. The Northern Calcareous Alps (NCA) has long been favored for the study of the Rhaetian, since the GSSP of the Triassic/Jurassic (T/J) boundary and other important T/J sections are situated in this region. However, the most famous Rhaetian sections in the NCA are composed of carbonates from the Koessen Formation and were situated in a large isolated intraplatform Basin (the Eiberg Basin), bordered to the south-east by a well-developed coral reef in the NW of the Tethys border. Several Rhaetian sections composed of marls and shales of the Zlambach Formation were deposited at the same time on the other side of this reef, in the oceanic Halstatt Basin, which was in direct connection to the Tethys. Here, we present new results on sedimentology, stable isotope and trace element analysis of both intraplatform and oceanic basin deposits in the NCA. Intraplatform Rhaetian sections from the Koessen Formation bear a few minor intervals of shales with enrichments in organic matter, some of which are associated to carbon isotopic excursions. Oceanic sections from the Hallstatt Basin are characterized at the base by very cyclic marl-limestone alternations. Higher up in the section, sediments progressively turn into pure shale deposits and the top of the Formation is characterized by organic

  9. Trajectories of Late Permian – Jurassic radiolarian extinction rates: no evidence for an end-Triassic mass extinction

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    W. Kiessling

    2011-02-01

    Full Text Available The hypothesis that ocean acidification was a proximate trigger of the marine end-Triassic mass extinction rests on the assumption that taxa that strongly invest in the secretion of calcium-carbonate skeletons were significantly more affected by the crisis than other taxa. An argument against this hypothesis is the great extinction toll of radiolarians that has been reported from work on local sections. Radiolarians have siliceous tests and thus should be less affected by ocean acidification. We compiled taxonomically vetted occurrences of late Permian and Mesozoic radiolarians and analyzed extinction dynamics of radiolarian genera. Although extinction rates were high at the end of the Triassic, there is no evidence for a mass extinction in radiolarians but rather significantly higher background extinction in the Triassic than in the Jurassic. Although the causes for this decline in background extinction levels remain unclear, the lack of a major evolutionary response to the end-Triassic event, gives support for the hypothesis that ocean acidification was involved in the dramatic extinctions of many calcifying taxa. doi:10.1002/mmng.201000017

  10. Isotopic evidence bearing on Late Triassic extinction events, Queen Charlotte Islands, British Columbia, and implications for the duration and cause of the Triassic/Jurassic mass extinction

    Science.gov (United States)

    Ward, P.D.; Garrison, G.H.; Haggart, J.W.; Kring, D.A.; Beattie, M.J.

    2004-01-01

    Stable isotope analyses of Late Triassic to earliest Jurassic strata from Kennecott Point in the Queen Charlotte Islands, British Columbia, Canada shows the presence of two distinct and different organic carbon isotope anomalies at the Norian/Rhaetian and Rhaetian/Hettangian (=Triassic/Jurassic) stage boundaries. At the older of these boundaries, which is marked by the disappearance of the bivalve Monotis, the isotope record shows a series of short-lived positive excursions toward heavier values. Strata approaching this boundary show evidence of increasing anoxia. At the higher boundary, marked by the disappearance of the last remaining Triassic ammonites and over 50 species of radiolarians, the isotopic pattern consists of a series of short duration negative anomalies. The two events, separated by the duration of the Rhaetian age, comprise the end-Triassic mass extinction. While there is no definitive evidence as to cause, the isotopic record does not appear similar to that of the impact-caused Cretaceous/Tertiary boundary extinction. ?? 2004 Published by Elsevier B.V.

  11. Contrasting microbial community changes during mass extinctions at the Middle/Late Permian and Permian/Triassic boundaries

    Science.gov (United States)

    Xie, Shucheng; Algeo, Thomas J.; Zhou, Wenfeng; Ruan, Xiaoyan; Luo, Genming; Huang, Junhua; Yan, Jiaxin

    2017-02-01

    Microbial communities are known to expand as a result of environmental deterioration during mass extinctions, but differences in microbial community changes between extinction events and their underlying causes have received little study to date. Here, we present a systematic investigation of microbial lipid biomarkers spanning ∼20 Myr (Middle Permian to Early Triassic) at Shangsi, South China, to contrast microbial changes associated with the Guadalupian-Lopingian boundary (GLB) and Permian-Triassic boundary (PTB) mass extinctions. High-resolution analysis of the PTB crisis interval reveals a distinct succession of microbial communities based on secular variation in moretanes, 2-methylhopanes, aryl isoprenoids, steranes, n-alkyl cyclohexanes, and other biomarkers. The first episode of the PTB mass extinction (ME1) was associated with increases in red algae and nitrogen-fixing bacteria along with evidence for enhanced wildfires and elevated soil erosion, whereas the second episode was associated with expansions of green sulfur bacteria, nitrogen-fixing bacteria, and acritarchs coinciding with climatic hyperwarming, ocean stratification, and seawater acidification. This pattern of microbial community change suggests that marine environmental deterioration was greater during the second extinction episode (ME2). The GLB shows more limited changes in microbial community composition and more limited environmental deterioration than the PTB, consistent with differences in species-level extinction rates (∼71% vs. 90%, respectively). Microbial biomarker records have the potential to refine our understanding of the nature of these crises and to provide insights concerning possible outcomes of present-day anthropogenic stresses on Earth's ecosystems.

  12. A new Lower Triassic ichthyopterygian assemblage from Fossil Hill, Nevada

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    Neil P. Kelley

    2016-01-01

    Full Text Available We report a new ichthyopterygian assemblage from Lower Triassic horizons of the Prida Formation at Fossil Hill in central Nevada. Although fragmentary, the specimens collected so far document a diverse fauna. One partial jaw exhibits isodont dentition with blunt tipped, mesiodistally compressed crowns and striated enamel. These features are shared with the Early Triassic genus Utatsusaurus known from coeval deposits in Japan and British Columbia. An additional specimen exhibits a different dentition characterized by relatively small, rounded posterior teeth resembling other Early Triassic ichthyopterygians, particularly Grippia. This Nevada assemblage marks a southward latitudinal extension for Early Triassic ichthyopterygians along the eastern margin of Panthalassa and indicates repeated trans-hemispheric dispersal events in Early Triassic ichthyopterygians.

  13. High precision time calibration of the Permian-Triassic boundary mass extinction event in a deep marine context

    Science.gov (United States)

    Baresel, Björn; Bucher, Hugo; Brosse, Morgane; Bagherpour, Borhan; Schaltegger, Urs

    2015-04-01

    To construct a revised and high resolution calibrated time scale for the Permian-Triassic boundary (PTB) we use (1) high-precision U-Pb zircon age determinations of a unique succession of volcanic ash layers interbedded with deep water fossiliferous sediments in the Nanpanjiang Basin (South China) combined with (2) accurate quantitative biochronology based on ammonoids, conodonts, radiolarians, and foraminifera and (3) tracers of marine bioproductivity (carbon isotopes) across the PTB. The unprecedented precision of the single grain chemical abrasion isotope-dilution thermal ionization mass spectrometry (CA-ID-TIMS) dating technique at sub-per mil level (radio-isotopic calibration of the PTB at the groups of processes. Using these alignments allows (1) positioning the PTB in different depositional setting and (2) solving the age contradictions generated by the misleading use of the first occurrence (FO) of the conodont Hindeodus parvus, whose diachronous first occurrences are arbitrarily used for placing the base of the Triassic. This new age framework provides the basis for a combined calibration of chemostratigraphic records with high-resolution biochronozones of the Late Permian and Early Triassic. Here, we present new single grain U-Pb zircon data of volcanic ash layers from two deep marine sections (Dongpan and Penglaitan) revealing stratigraphic consistent dates over several volcanic ash layers bracketing the PTB. These analyses define weighted mean 206Pb/238U ages of 251.956±0.033 Ma (Dongpan) and 252.062±0.043 Ma (Penglaitan) for the last Permian ash bed. By calibration with detailed litho- and biostratigraphy new U-Pb ages of 251.953±0.038 Ma (Dongpan) and 251.907±0.033 Ma (Penglaitan) are established for the onset of the Triassic.

  14. Dobrogeria aegyssensis, a new early Spathian (Early Triassic) coelacanth from North Dobrogea (Romania)

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    Cavin, Lionel; Grădinaru, Eugen

    2014-06-01

    The Early Triassic witnessed the highest taxic diversity of coelacanths (or Actinistia), a clade with a single living genus today. This peak of diversity is accentuated here with the description of a new coelacanth discovered in the lower Spathian (Upper Olenekian, Lower Triassic) cropping out in the Tulcea Veche (Old Tulcea) promontory, in the city of Tulcea, in North Dobrogea, Romania. The bone remains were preserved in a block of limestone, which was chemically dissolved. The resulting 3D and matrix-free ossifications correspond mostly to elements of the skull and branchial apparatus. Posterior parietals, postparietal with associated prootic and basisphenoid allow a precise description of the neurocranium. Ossifications of the lower jaw, together with branchial and pectoral elements, complete the description of this coelacanth and support the coining of a new generic and specific name, Dobrogeria aegyssensis. A phylogenetic analysis of actinistians with the new species recovers clades which were found in most recent analyses, i.e. the Sasseniidae, the Laugiidae, the Coelacanthiformes, the Latimerioidei, the Mawsoniidae and the Latimeriidae, and identifies the new taxon as a non-latimerioid coelacanthiform.

  15. Testing the limits in a greenhouse ocean: Did low nitrogen availability limit marine productivity during the end-Triassic mass extinction?

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    Schoepfer, Shane D.; Algeo, Thomas J.; Ward, Peter D.; Williford, Kenneth H.; Haggart, James W.

    2016-10-01

    The end-Triassic mass extinction has been characterized as a 'greenhouse extinction', related to rapid atmospheric warming and associated changes in ocean circulation and oxygenation. The response of the marine nitrogen cycle to these oceanographic changes, and the extent to which mass extinction intervals represent a deviation in nitrogen cycling from other ice-free 'greenhouse' periods of Earth history, remain poorly understood. The well-studied Kennecott Point section in Haida Gwaii, British Columbia, Canada, was deposited in the open Panthalassic Ocean, and is used here as a test case to better understand changes in the nitrogen cycle and marine productivity from the pre-crisis greenhouse of the Rhaetian to the latest-Rhaetian crisis interval. We estimated marine productivity from the late Norian to the early Hettangian using TOC- and P-based paleoproductivity transform equations, and then compared these estimates to records of sedimentary nitrogen isotopes, redox-sensitive trace elements, and biomarker data. Major negative excursions in δ15N (to ≤ 0 ‰) correspond to periods of depressed marine productivity. During these episodes, the development of a stable pycnocline below the base of the photic zone suppressed vertical mixing and limited N availability in surface waters, leading to low productivity and increased nitrogen fixation, as well as ecological stresses in the photic zone. The subsequent shoaling of euxinic waters into the ocean surface layer was fatal for most Triassic marine fauna, although the introduction of regenerated NH4+ into the photic zone may have allowed phytoplankton productivity to recover. These results indicate that the open-ocean nitrogen cycle was influenced by climatic changes during the latest Triassic, despite having existed in a greenhouse state for over 50 million years previously, and that low N availability limited marine productivity for hundreds of thousands of years during the end-Triassic crisis.

  16. Repeated Carbon-Cycle Disturbances at the Permian-Triassic Boundary Separate two Mass Extinctions

    Science.gov (United States)

    Nicol, J. A.; Watson, L.; Claire, M.; Buick, R.; Catling, D. C.

    2004-12-01

    Non-marine organic matter in Permian-Triassic sediments from the Blue Mountains, eastern Australia shows seven negative δ13C excursions of up to 7%, terminating with a positive excursion of 4%. Fluctuations start at the late Permian Glossopteris floral extinction and continue until just above the palynological Permian-Triassic boundary, correlated with the peak of marine mass extinction. The isotopic fluctuations are not linked to changes in depositional setting, kerogen composition or plant community, so they evidently resulted from global perturbations in atmospheric δ13C and/or CO2. The pattern was not produced by a single catastrophe such as a meteorite impact, and carbon-cycle calculations indicate that gas release during flood-basalt volcanism was insufficient. Methane-hydrate melting can generate a single -7% shift, but cannot produce rapid multiple excursions without repeated reservoir regeneration and release. However, the data are consistent with repeated overturning of a stratified ocean, expelling toxic gases that promoted sequential mass extinctions in the terrestrial and marine realms.

  17. Investigating A Unique Open Ocean Geochemical Record Of the End Triassic Mass Extinction from Panthalassa

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    Marroquín, S. M.; Gill, B. C.; Them, T. R., II; Trabucho-Alexandre, J. P.; Aberhan, M.; Owens, J. D.; Gröcke, D. R.; Caruthers, A. H.

    2017-12-01

    The end-Triassic mass extinction ( 201 Ma) was a time of intense disturbance for marine communities. This event is estimated to have produced as much as a loss of 80% of known marine species. The protracted interval of elevated extinction rates is also characterized by a major carbon cycle perturbation and potentially widespread oxygen deficiency within the oceans. While the causes of extinction and environmental feedbacks are still debated it is hypothesized to have been triggered by massive volcanism associated with the Central Atlantic Magmatic Province flood basalts. However, our understanding of the Latest Triassic-Earliest Jurassic interval is limited due to the lack of well-preserved stratigraphic successions outside of the Tethys Ocean (present day Europe), with most of the records from epicontinental and marginal marine settings. To expand our understanding of this critical interval, our study seeks to document biological and environmental changes elsewhere. Specifically, we document and reconstruct these changes in the equatorial Panthalassan Ocean. We will present new data from a sedimentary succession preserved in the Wrangell Mountains of Alaska that spans the Late Triassic through Early Jurassic. The sedimentary succession represents a mixed carbonate-siliciclastic ramp that was deposited at tropical latitudes, adjacent to an island arc in the open Panthalassan Ocean. This succession affords a unique view of open marine conditions, and also holds the potential for excellent temporal control as it contains abundant ash layers throughout, as well as, key ammonite and bivalve fossil occurrences that provide biostratigraphic control. We will present an integrated geochemical and paleontological record from this site using several geochemical proxies (carbon, δ13Ccarb and % total organic carbon, sulfur, δ34S, as well as pyrite contents and iron speciation) along with ammonite and bivalve occurrence data to reconstruct the record of environmental and

  18. Proliferation of MISS-related microbial mats following the end-Permian mass extinction in terrestrial ecosystems: Evidence from the Lower Triassic of the Yiyang area, Henan Province, North China

    Science.gov (United States)

    Tu, Chenyi; Chen, Zhong-Qiang; Retallack, Gregory J.; Huang, Yuangeng; Fang, Yuheng

    2016-03-01

    Microbially induced sedimentary structures (MISSs) are commonly present in siliciclastic shallow marine settings following the end-Permian mass extinction, but have been rarely reported in the post-extinction terrestrial ecosystems. Here, we present six types of well-preserved MISSs from the upper Sunjiagou Formation and lower Liujiagou Formation of Induan (Early Triassic) age in the Yiyang area, Henan Province, North China. These MISSs include: polygonal sand cracks, worm-like structures, wrinkle structures, sponge pore fabrics, gas domes, and leveled ripple marks. Microanalysis shows that these MISSs are characterized by thin clayey laminae and filamentous mica grains arranged parallel to bedding plane as well as oriented matrix supported quartz grains, which are indicative of biogenic origin. Facies analysis suggests that the MISS-hosting sediments were deposited in a fluvial sedimentary system during the Early Triassic, including lake delta, riverbeds/point bars, and flood plain paleoenvironments. Abundant MISSs from Yiyang indicate that microbes also proliferated in terrestrial ecosystems in the aftermath of the Permian-Triassic (P-Tr) biocrisis, like they behaved in marine ecosystems. Microbial blooms, together with dramatic loss of metazoans, may reflect environmental stress and degradation of terrestrial ecosystems or arid climate immediately after the severe Permian-Triassic ecologic crisis.

  19. Macropredatory ichthyosaur from the Middle Triassic and the origin of modern trophic networks.

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    Fröbisch, Nadia B; Fröbisch, Jörg; Sander, P Martin; Schmitz, Lars; Rieppel, Olivier

    2013-01-22

    The biotic recovery from Earth's most severe extinction event at the Permian-Triassic boundary largely reestablished the preextinction structure of marine trophic networks, with marine reptiles assuming the predator roles. However, the highest trophic level of today's marine ecosystems, i.e., macropredatory tetrapods that forage on prey of similar size to their own, was thus far lacking in the Paleozoic and early Mesozoic. Here we report a top-tier tetrapod predator, a very large (>8.6 m) ichthyosaur from the early Middle Triassic (244 Ma), of Nevada. This ichthyosaur had a massive skull and large labiolingually flattened teeth with two cutting edges indicative of a macropredatory feeding style. Its presence documents the rapid evolution of modern marine ecosystems in the Triassic where the same level of complexity as observed in today's marine ecosystems is reached within 8 My after the Permian-Triassic mass extinction and within 4 My of the time reptiles first invaded the sea. This find also indicates that the biotic recovery in the marine realm may have occurred faster compared with terrestrial ecosystems, where the first apex predators may not have evolved before the Carnian.

  20. Early Jurassic diversification of pycnodontiform fishes (Actinopterygii, Neopterygii) after the end-Triassic extinction event: evidence from a new genus and species, Grimmenodon aureum.

    Science.gov (United States)

    Stumpf, Sebastian; Ansorge, Jörg; Pfaff, Cathrin; Kriwet, Jürgen

    2017-07-04

    A new genus and species of pycnodontiform fishes, Grimmenodon aureum , from marginal marine, marine-brackish lower Toarcian ( Harpoceras exaratum ammonite subzone) clay deposits of Grimmen in northeastern Germany is described. The single specimen represents a diagnostic left prearticular dentition characterized by unique tooth arrangement and ornamentation patterns. Grimmenodon aureum , gen. et sp. nov., is the second unambiguously identified pycnodontiform species from the Early Jurassic, in addition to Eomesodon liassicus from the early Lower Jurassic of western Europe. We also report an indeterminate pycnodontiform tooth crown from the upper Pliensbachian ( Pleuroceras apyrenum ammonite subzone) of the same site. The material expands the Early Jurassic range of pycnodontiforms significantly northwards and confirms their presence before and immediately following the onset of the Toarcian Oceanic Anoxic Event (T-OAE) in the marginal marine ecosystems south of the Fennoscandian Shield. Moreover, the new records indicate that the Early Jurassic diversity of pycnodontiform fishes was greater than previously assumed and probably equaled that of the Late Triassic. Therefore, it is hypothesized that the Triassic-Jurassic mass extinction event did not affect pycnodontiform fishes significantly. Micro-computed tomography was used to study the internal anatomy of the prearticular of Grimmenodon aureum , gen. et sp. nov. Our results show that no replacement teeth were formed within the tooth-bearing bone but rather were added posteriorly to functional teeth. http://zoobank.org/urn:lsid:zoobank.org:pub:A56BDE9C-40C4-4CFA-9C2E-F5FA35A66F2 Citation for this article: Stumpf, S., J. Ansorge, C. Pfaff, and J. Kriwet. 2017. Early Jurassic diversification of pycnodontiform fishes (Actinopterygii, Neopterygii) after the end-Triassic extinction event: Evidence from a new genus and species, Grimmenodon aureum . Journal of Vertebrate Paleontology. DOI: 10.1080/02724634.2017.1344679.

  1. Hg concentrations from Late Triassic and Early Jurassic sedimentary rocks: first order similarities and second order depositional and diagenetic controls

    Science.gov (United States)

    Yager, J. A.; West, A. J.; Bergquist, B. A.; Thibodeau, A. M.; Corsetti, F. A.; Berelson, W.; Bottjer, D. J.; Rosas, S.

    2016-12-01

    Mercury concentrations in sediments have recently gained prominence as a potential tool for identifying large igneous province (LIP) volcanism in sedimentary records. LIP volcanism coincides with several mass extinctions during the Phanerozoic, but it is often difficult to directly tie LIP activity with the record of extinction in marine successions. Here, we build on mercury concentration data reported by Thibodeau et al. (Nature Communications, 7:11147, 2016) from the Late Triassic and Early Jurassic of New York Canyon, Nevada, USA. Increases in Hg concentrations in that record were attributed to Central Atlantic Magmatic Province (CAMP) activity in association with the end-Triassic mass extinction. We expand the measured section from New York Canyon and report new mercury concentrations from Levanto, Peru, where dated ash beds provide a discrete chronology, as well as St. Audrie's Bay, UK, a well-studied succession. We correlate these records using carbon isotopes and ammonites and find similarities in the onset of elevated Hg concentrations and Hg/TOC in association with changes in C isotopes. We also find second order patterns that differ between sections and may have depositional and diagenetic controls. We will discuss these changes within a sedimentological framework to further understand the controls on Hg concentrations in sedimentary records and their implications for past volcanism.

  2. PALAEOECOLOGICAL CHANGES AFTER THE END-PERMIAN MASS EXTINCTION: EARLY TRIASSIC OSTRACODS FROM NORTHWESTERN GUANGXI PROVINCE,SOUTH CHINA

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    SYLVIE CRASQUIN-SOLEAU

    2006-03-01

    Full Text Available Early Triassic (Griesbachian to Spathian ostracod faunas are here first discovered and described form the Guangxi Province, South China. Thirty-seven species belonging to fourteen genera are recognized. Seven species are new: Bairdia fengshanensis n.sp., Bairdia wailiensis n.sp., Liuzhinia guangxiensis n.sp., Ptychobairdia luciaae n.sp., Ptychobairdia aldaae n.sp., Paracypris jinyaensis n.sp. and Paracypris gaetanii n.sp. The Griesbachian assemblage from the basal microbial limestone is well diversified and does not suggest any abnormal palaeoenvironmental conditions in terms of salinity, temperature or oxygen content. Particularly, the ostracods are typical of well oxygenated water and do not reflect any anoxia. Dienerian and Smithian ostracods are evidenced for the first time and the assemblages suggest less favourable palaeoenviromental conditions. Diversity and abundance of ostracod assemblages recovered from the Spathian on. The main taxonomic turnover among ostracod assemblages occurred seemingly between the Griesbachian and the Spathian.

  3. Globacrochordiceras gen. nov. (Acrochordiceratidae, late Early Triassic and its significance for stress-induced evolutionary jumps in ammonoid lineages (cephalopods

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    C. Monnet

    2013-08-01

    Full Text Available Globacrochordiceras transpacificum gen. et sp. nov. is an ammonoid (Ammonoidea, Cephalopoda with a shell characterized by plicate ribbing (rounded and undulating ribs strengthening on the venter without interruption, increasing involution through ontogeny, overhanging and deep umbilical wall, absence of tuberculation, subtriangular whorl section, globose adult shape with a closed umbilicus followed by an abrupt egressive coiling, and a subammonitic adult suture line. This new taxon occurs in Nevada (USA and in Guangxi (South China. It has its typical occurrence within the Neopopanoceras haugi Zone of late Spathian age (Early Triassic. The plicate ribbing, suture line and general shell shape are diagnostic of the family Acrochordiceratidae. The large adult size, high degree of involution and subammonitic suture line of Globacrochordiceras markedly contrast with the next younger genus of the family (Paracrochordiceras of early Anisian age, Middle Triassic, which is evolute and displays a ceratitic suture shape. Shell coiling and suture line of Globacrochordiceras are closer to that of the youngest member of the family: Acrochordiceras carolinae (late middle Anisian. The latter is the end-member of a long-term morphological evolutionary trend of the family during the early and middle Anisian. This trend composed of classical increases in adult size (Cope's rule, shell involution and suture indentation, lasted ca. four Myr. The sudden morphological evolutionary jump between Globacrochordiceras and Paracrochordiceras at the Spathian/Anisian (Early/Middle Triassic boundary may correspond to a generalized morphological reset of long-term trends, a process that differs from classic paedomorphic transformations. A dramatic global sea level change and carbon isotope positive excursion at the Early/Middle Triassic boundary both indicate stressful environmental changes that may have triggered this evolutionary jump. doi:10.1002/mmng.201300010

  4. End-Triassic nonmarine biotic events

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    Spencer G. Lucas

    2015-10-01

    Full Text Available The Late Triassic was a prolonged interval of elevated extinction rates and low origination rates that manifested themselves in a series of extinctions during Carnian, Norian and Rhaetian time. Most of these extinctions took place in the marine realm, particularly affecting radiolarians, conodonts, bivalves, ammonoids and reef-building organisms. On land, the case for a Late Triassic mass extinction is much more tenuous and has largely focused on tetrapod vertebrates (amphibians and reptiles, though some workers advocate a sudden end-Triassic (TJB extinction of land plants. Nevertheless, an extensive literature does not identify a major extinction of land plants at the TJB, and a comprehensive review of palynological records concluded that TJB vegetation changes were non-uniform (different changes in different places, not synchronous and not indicative of a mass extinction of land plants. Claims of a substantial perturbation of plant ecology and diversity at the TJB in East Greenland are indicative of a local change in the paleoflora largely driven by lithofacies changes resulting in changing taphonomic filters. Plant extinctions at the TJB were palaeogeographically localized events, not global in extent. With new and more detailed stratigraphic data, the perceived TJB tetrapod extinction is mostly an artifact of coarse temporal resolution, the compiled correlation effect. The amphibian, archosaur and synapsid extinctions of the Late Triassic are not concentrated at the TJB, but instead occur stepwise, beginning in the Norian and extending into the Hettangian. There was a disruption of the terrestrial ecosystem across the TJB, but it was more modest than generally claimed. The ecological severity of the end-Triassic nonmarine biotic events are relatively low on the global scale. Biotic turnover at the end of the Triassic was likely driven by the CAMP (Central Atlantic Magmatic Province eruptions, which caused significant environmental

  5. Structural changes of marine communities over the Permian-Triassic transition: Ecologically assessing the end-Permian mass extinction and its aftermath

    Science.gov (United States)

    Chen, Zhong-Qiang; Tong, Jinnan; Liao, Zhuo-Ting; Chen, Jing

    2010-08-01

    The Permian/Triassic (P/Tr) transition is ecologically assessed based on examining 23 shelly communities from five shallow platform, ramp and shelf basin facies Permian-Triassic boundary (PTB) sections in South China. The shelly communities have undergone two major collapses coinciding with the two episodes of the end-Permian mass extinction. The first P/Tr extinction event devastated shelly communities in all types of settings to some extent. The basin communities have been more severely impacted than both platform and ramp communities. The survival faunas have rebounded more rapidly in shallow niches than in relatively deep habitats. The second P/Tr crisis destroyed the survival communities in shallow setting and had little impact on the basin communities in terms of community structures. The early Griesbachian communities are overall low-diversity and high-dominance. The governorship switch from brachiopods to bivalves in marine communities has been facilitated by two pulses of the end-Permian mass extinction and the whole takeover process took about 200 ka across the P/Tr boundary. Bivalve ecologic takeover initially occurred immediately after the first P/Tr extinction in shallow water habitats and was eventually completed in all niches after the second P/Tr event. Some post-extinction communities have the irregular rarefaction curves due to the unusual community structures rather than sampling intensities.

  6. Terrestrial origin of viviparity in mesozoic marine reptiles indicated by early triassic embryonic fossils.

    Science.gov (United States)

    Motani, Ryosuke; Jiang, Da-yong; Tintori, Andrea; Rieppel, Olivier; Chen, Guan-bao

    2014-01-01

    Viviparity in Mesozoic marine reptiles has traditionally been considered an aquatic adaptation. We report a new fossil specimen that strongly contradicts this traditional interpretation. The new specimen contains the oldest fossil embryos of Mesozoic marine reptile that are about 10 million years older than previous such records. The fossil belongs to Chaohusaurus (Reptilia, Ichthyopterygia), which is the oldest of Mesozoic marine reptiles (ca. 248 million years ago, Early Triassic). This exceptional specimen captures an articulated embryo in birth position, with its skull just emerged from the maternal pelvis. Its headfirst birth posture, which is unlikely to be a breech condition, strongly indicates a terrestrial origin of viviparity, in contrast to the traditional view. The tail-first birth posture in derived ichthyopterygians, convergent with the conditions in whales and sea cows, therefore is a secondary feature. The unequivocally marine origin of viviparity is so far not known among amniotes, a subset of vertebrate animals comprising mammals and reptiles, including birds. Therefore, obligate marine amniotes appear to have evolved almost exclusively from viviparous land ancestors. Viviparous land reptiles most likely appeared much earlier than currently thought, at least as early as the recovery phase from the end-Permian mass extinction.

  7. Geochemical and palynological records for the end-Triassic Mass-Extinction Event in the NE Paris Basin (Luxemburg)

    Science.gov (United States)

    Kuhlmann, Natascha; van de Schootbrugge, Bas; Thein, Jean; Fiebig, Jens; Franz, Sven-Oliver; Hanzo, Micheline; Colbach, Robert; Faber, Alain

    2016-04-01

    The End-Triassic mass-extinction event is one of the "big five" mass extinctions in Earth's history. Large scale flood basalt volcanism associated with the break-up of Pangaea, which resulted in the opening of the central Atlantic Ocean, is considered as the leading cause. In addition, an asteroid impact in Rochechouart (France; 201 ± 2 Ma) may have had a local influence on ecosystems and sedimentary settings. The Luxembourg Embayment, in the NE Paris Basin, offers a rare chance to study both effects in a range of settings from deltaic to lagoonal. A multidisciplinary study (sedimentology, geochemistry, palynology) has been carried out on a number of outcrops and cores that span from the Norian to lower Hettangian. Combined geochemical and palynological records from the Boust core drilled in the NE Paris Basin, provide evidence for paleoenvironmental changes associated with the end-Triassic mass-extinction event. The Triassic-Jurassic stratigraphy of the Boust core is well constrained by palynomorphs showing the disappaerance of typical Triassic pollen taxa (e.g. Ricciisporites tuberculates) and the occurrence of the marker species Polypodiisporites polymicroforatus within the uppermost Rhaetian, prior to the Hettangian dominance of Classopollis pollen. The organic carbon stable isotope record (δ13Corg) spanning the Norian to Hettangian, shows a series of prominent negative excursions within the middle Rhaetian, followed by a trend towards more positive values (approx -24 per mille) within the uppermost Rhaetian Argiles de Levallois Member. The lowermost Hettangian is characterized by a major negative excursion, reaching - 30 per mille that occurs in organic-rich sediments. This so-called "main negative excursion" is well-known from other locations, for example from Mariental in Northern Germany and from St Audrie's Bay in England, and Stenlille in Denmark. Based on redox-sensitive trace element records (V, Cr, Ni, Co, Th, U) the lowermost Hettangian in most of

  8. Early Triassic wrinkle structures on land: stressed environments and oases for life

    Science.gov (United States)

    Chu, Daoliang; Tong, Jinnan; Song, Haijun; Benton, Michael J.; Bottjer, David J.; Song, Huyue; Tian, Li

    2015-06-01

    Wrinkle structures in rocks younger than the Permian-Triassic (P-Tr) extinction have been reported repeatedly in marine strata, but rarely mentioned in rocks recording land. Here, three newly studied terrestrial P-Tr boundary rock succession in North China have yielded diverse wrinkle structures. All of these wrinkles are preserved in barely bioturbated shore-shallow lacustrine siliciclastic deposits of the Liujiagou Formation. Conversely, both the lacustrine siliciclastic deposits of the underlying Sunjiagou Formation and the overlying Heshanggou Formation show rich bioturbation, but no wrinkle structures or other microbial-related structures. The occurrence of terrestrial wrinkle structures in the studied sections reflects abnormal hydrochemical and physical environments, presumably associated with the extinction of terrestrial organisms. Only very rare trace fossils occurred in the aftermath of the P-Tr extinction, but most of them were preserved together with the microbial mats. This suggests that microbial mats acted as potential oases for the surviving aquatic animals, as a source of food and oxygen. The new finds suggests that extreme environmental stresses were prevalent both in the sea and on land through most of the Early Triassic.

  9. Multiple episodes of extensive marine anoxia linked to global warming and continental weathering following the latest Permian mass extinction.

    Science.gov (United States)

    Zhang, Feifei; Romaniello, Stephen J; Algeo, Thomas J; Lau, Kimberly V; Clapham, Matthew E; Richoz, Sylvain; Herrmann, Achim D; Smith, Harrison; Horacek, Micha; Anbar, Ariel D

    2018-04-01

    Explaining the ~5-million-year delay in marine biotic recovery following the latest Permian mass extinction, the largest biotic crisis of the Phanerozoic, is a fundamental challenge for both geological and biological sciences. Ocean redox perturbations may have played a critical role in this delayed recovery. However, the lack of quantitative constraints on the details of Early Triassic oceanic anoxia (for example, time, duration, and extent) leaves the links between oceanic conditions and the delayed biotic recovery ambiguous. We report high-resolution U-isotope (δ 238 U) data from carbonates of the uppermost Permian to lowermost Middle Triassic Zal section (Iran) to characterize the timing and global extent of ocean redox variation during the Early Triassic. Our δ 238 U record reveals multiple negative shifts during the Early Triassic. Isotope mass-balance modeling suggests that the global area of anoxic seafloor expanded substantially in the Early Triassic, peaking during the latest Permian to mid-Griesbachian, the late Griesbachian to mid-Dienerian, the Smithian-Spathian transition, and the Early/Middle Triassic transition. Comparisons of the U-, C-, and Sr-isotope records with a modeled seawater PO 4 3- concentration curve for the Early Triassic suggest that elevated marine productivity and enhanced oceanic stratification were likely the immediate causes of expanded oceanic anoxia. The patterns of redox variation documented by the U-isotope record show a good first-order correspondence to peaks in ammonoid extinctions during the Early Triassic. Our results indicate that multiple oscillations in oceanic anoxia modulated the recovery of marine ecosystems following the latest Permian mass extinction.

  10. Ocean redox change at the Permian-Triassic mass extinction

    DEFF Research Database (Denmark)

    Ruhl, Micha; Bjerrum, Christian J.; Canfield, Donald Eugene

    2013-01-01

    Earth’s history is marked by multiple events of ocean anoxia developing along continental margins and po¬tentially into the open ocean realm. These events of¬ten coincide with the emplacement of large igneous provinces (LIPs) on continents, major perturbations of global geochemical cycles and mar...... these oceanographic changes to similar observations for the Triassic-Jurassic mass extinction and discuss environmental forcing, poten¬tially inherent to major volcanic events and leading to global environmental change and extinction...... ocean redox change over the largest mass extinction event in Earth history, at the Permian-Tri¬assic boundary (at ~252 Ma). This event is marked by a major perturbation in the global exogenic carbon cycle (and associated major negative carbon isotope excursion (CIE)), likely initiated by carbon...... (anoxic but not euxinic) coinciding with the main extinction event. Molybdenum enrichments, often indicative for freely available sulfide in the water-column, only occur dur¬ing the second phase of euxinia. This pattern of ocean redox-change in Svalbard direct¬ly reflects similar trends in Greenland...

  11. Triassic arc-derived detritus in the Triassic Karakaya accretionary complex was not derived from either the S Eurasian margin (Istanbul terrane) or the N Gondwana margin (Taurides)

    Science.gov (United States)

    Ustaömer, Timur; Ayda Ustaömer, Petek; Robertson, Alastair H. F.; Gerdes, Axel; Zulauf, Gernold

    2014-05-01

    We present new U-Pb zircon source age data for Upper Triassic sandstones of the Istanbul Terrane (S Eurasian margin) and also for Triassic sandstones of the Taurides (N Gondwana margin). The main aim is to detect and quantify the contribution of Triassic magmatism as detritus to either of these crustal blocks. This follows the recent discovery of a Triassic magmatic arc source for the Triassic sandstones of the Palaeotethyan Karakaya subduction-accretion complex (Ustaömer et al. 2013; this meeting). Carboniferous (Variscan) zircon grains also form a significant detrital population, plus several more minor populations. Six sandstone samples were studied, two from the İstanbul Terrane (Bakırlıkıran Formation of the Kocaeli Triassic Basin) and four from the Tauride Autochthon (latest Triassic Üzümdere Formation and Mid-Triassic Kasımlar Formations; Beyşehir region). Detrital zircon grains were dated by the laser ablation-inductively coupled plasma-mass spectrometer (LA-ICP-MS) U-Pb method at Goethe University, Frankfurt. Our results do not reveal Triassic detritus in the Üzümdere Formation. The U-Pb age of the analysed zircon grains ranges from 267 Ma to 3.2 Ga. A small fraction of Palaeozoic zircons are Permian (267 to 296 Ma), whereas the remainder are Early Palaeozoic. Ordovician grains (4%) form two age clusters, one at ca. 450 Ma and the other at ca. 474 Ma. Cambrian-aged grains dominate the zircon population, while the second largest population is Ediacaran (576 to 642 Ma). Smaller populations occur at 909-997 Ma, 827-839 Ma, 1.8-2.0 Ga and 2.4-2.6 Ga. The sandstones of the Kasımlar Formation have similar zircon age cluster to those of the somewhat younger Üzümdere Formation, ranging from 239 Ma to 2.9 Ga. A few grains gave Anisian ages. Cambrian zircon grains are less pronounced than in the Kasımlar Formation compared to the Üzümdere Formation. The detrital zircon record of Tauride sandstones, therefore, not indicates significant contribution

  12. Intense and widespread seismicity during the end-Triassic mass extinction due to emplacement of a large igneous province

    DEFF Research Database (Denmark)

    Lindström, Sofie; Pedersen, Gunver Krarup; Schootbrugge, Bas van de

    2015-01-01

    Multiple levels of earthquake-induced soft-sediment deformations (seismites) are concentrated in the end-Triassic mass extinction interval across Europe. The repetitive nature of the seismites rules out an origin by an extraterrestrial impact. Instead, this intense seismic activity is linked...

  13. Terrestrial Origin of Viviparity in Mesozoic Marine Reptiles Indicated by Early Triassic Embryonic Fossils

    OpenAIRE

    Motani, Ryosuke; Jiang, Da-yong; Tintori, Andrea; Rieppel, Olivier; Chen, Guan-bao

    2014-01-01

    Viviparity in Mesozoic marine reptiles has traditionally been considered an aquatic adaptation. We report a new fossil specimen that strongly contradicts this traditional interpretation. The new specimen contains the oldest fossil embryos of Mesozoic marine reptile that are about 10 million years older than previous such records. The fossil belongs to Chaohusaurus (Reptilia, Ichthyopterygia), which is the oldest of Mesozoic marine reptiles (ca. 248 million years ago, Early Triassic). This exc...

  14. Terrestrial origin of viviparity in mesozoic marine reptiles indicated by early triassic embryonic fossils.

    Directory of Open Access Journals (Sweden)

    Ryosuke Motani

    Full Text Available Viviparity in Mesozoic marine reptiles has traditionally been considered an aquatic adaptation. We report a new fossil specimen that strongly contradicts this traditional interpretation. The new specimen contains the oldest fossil embryos of Mesozoic marine reptile that are about 10 million years older than previous such records. The fossil belongs to Chaohusaurus (Reptilia, Ichthyopterygia, which is the oldest of Mesozoic marine reptiles (ca. 248 million years ago, Early Triassic. This exceptional specimen captures an articulated embryo in birth position, with its skull just emerged from the maternal pelvis. Its headfirst birth posture, which is unlikely to be a breech condition, strongly indicates a terrestrial origin of viviparity, in contrast to the traditional view. The tail-first birth posture in derived ichthyopterygians, convergent with the conditions in whales and sea cows, therefore is a secondary feature. The unequivocally marine origin of viviparity is so far not known among amniotes, a subset of vertebrate animals comprising mammals and reptiles, including birds. Therefore, obligate marine amniotes appear to have evolved almost exclusively from viviparous land ancestors. Viviparous land reptiles most likely appeared much earlier than currently thought, at least as early as the recovery phase from the end-Permian mass extinction.

  15. A new archosauriform (Reptilia: Diapsida) from the Manda beds (Middle Triassic) of southwestern Tanzania.

    Science.gov (United States)

    Nesbitt, Sterling J; Butler, Richard J; Gower, David J

    2013-01-01

    Archosauria and their closest relatives, the non-archosaurian archosauriforms, diversified in the Early and Middle Triassic, soon after the end-Permian extinction. This diversification is poorly documented in most Lower and Middle Triassic rock sequences because fossils of early groups of archosauriforms are relatively rare compared to those of other amniotes. The early Middle Triassic (? late Anisian) Manda beds of southwestern Tanzania form an exception, with early archosaur skeletons being relatively common and preserved as articulated or associated specimens. The Manda archosaur assemblage is exceptionally diverse for the Middle Triassic. However, to date, no non-archosaurian archosauriforms have been reported from these rocks. Here, we name a new taxon, Asperoris mnyama gen. et sp. nov., from the Manda beds and thoroughly describe the only known specimen. The specimen consists of a well-preserved partial skull including tooth-bearing elements (premaxilla, maxilla), the nasal, partial skull roof, and several incomplete elements. All skull elements are covered in an autapomorphic highly rugose sculpturing. A unique combination of character states indicates that A. mnyama lies just outside Archosauria as a stem archosaur within Archosauriformes, but more precise relationships of A. mnyama relative to other early archosauriform clades (e.g., Erythrosuchidae) cannot be determined currently. Asperoris mnyama is the first confirmed non-archosaurian archosauriform from the Manda beds and increases the morphological and taxonomic diversity of early archosauriforms known from the Middle Triassic. The direct association of A. mnyama with species referable to Archosauria demonstrates that non-archosaurian archosauriforms were present during the rise and early diversification of Archosauria. Non-archosaurian archosauriforms and archosaurs co-occur in fossil reptile assemblages across Pangaea from the late Early Triassic to the end of the Late Triassic.

  16. The Early to Middle Triassic continental–marine transition of NW Bulgaria: sedimentology, palynology and sequence stratigraphy

    Directory of Open Access Journals (Sweden)

    Ajdanlijsky George

    2018-04-01

    Full Text Available Sedimentary facies and cycles of the Triassic continental–marine transition of NW Bulgaria are documented in detail from reference sections along the Iskar river gorge between the villages of Tserovo and Opletnya. The depositional environments evolved from anastomosing and meandering river systems in the Petrohan Terrigenous Group to mixed fluvial and tidal settings in the Svidol Formation, and to peritidal and shallow-marine conditions in the Opletnya Member of the Mogila Formation. For the first time, the palynostratigraphic data presented here allow for dating the transitional interval and for the precise identification of a major sequence boundary between the Petrohan Terrigenous Group and the Svidol Formation (Iskar Carbonate Group. This boundary most probably corresponds to the major sequence boundary Ol4 occurring in the upper Olenekian of the Tethyan realm and thus enables interregional correlation. The identification of regionally traceable sequence boundaries based on biostratigraphic age control is a first step towards a more accurate stratigraphic correlation and palaeogeographic interpretation of the Early to early Middle Triassic in NW Bulgaria.

  17. The Early to Middle Triassic continental-marine transition of NW Bulgaria: sedimentology, palynology and sequence stratigraphy

    Science.gov (United States)

    Ajdanlijsky, George; Götz, Annette E.; Strasser, André

    2018-04-01

    Sedimentary facies and cycles of the Triassic continental-marine transition of NW Bulgaria are documented in detail from reference sections along the Iskar river gorge between the villages of Tserovo and Opletnya. The depositional environments evolved from anastomosing and meandering river systems in the Petrohan Terrigenous Group to mixed fluvial and tidal settings in the Svidol Formation, and to peritidal and shallow-marine conditions in the Opletnya Member of the Mogila Formation. For the first time, the palynostratigraphic data presented here allow for dating the transitional interval and for the precise identification of a major sequence boundary between the Petrohan Terrigenous Group and the Svidol Formation (Iskar Carbonate Group). This boundary most probably corresponds to the major sequence boundary Ol4 occurring in the upper Olenekian of the Tethyan realm and thus enables interregional correlation. The identification of regionally traceable sequence boundaries based on biostratigraphic age control is a first step towards a more accurate stratigraphic correlation and palaeogeographic interpretation of the Early to early Middle Triassic in NW Bulgaria.

  18. Possible climate effects of the CAMP intrusive and extrusive activity and its influence on the end-Triassic mass extinction

    Science.gov (United States)

    Marzoli, A.; Davies, J.; Valeriani, L.; Preto, N.; Cirilli, S.; Panfili, G.; Dal Corso, J.; Vasconcellos, E.; Ernesto, M.; Youbi, N.; Callegaro, S.

    2017-12-01

    The end-Triassic global climate changes were probably triggered by the emplacement of the CAMP (Central Atlantic magmatic province). Here we explore the possibility that CAMP intrusions triggered global warming, while CAMP eruptions triggered short-lived cooling events. The main phase of the end-Triassic environmental changes and mass extinction was marked by two carbon isotopic excursions (CIEs). Based on stratigraphic and geochronologic data, we show that the earliest CAMP intrusions were emplaced at ca. 201.6 Ma prior to the first CIE (Davies et al., 2017). The main phase of CAMP magmatism started during the first CIE at ca. 201.5 Ma and continued until the second CIE and the Triassic-Jurassic boundary (at ca. 201.3 Ma). In particular, intrusion of the over 1 million cubic km of basaltic sills in Amazonia (Brazil) and of widespread sills from North America and Africa occurred within this interval. Multidisciplinary analyses show that organic matter rich sediments close to the sills from Brazil, Morocco, and the USA underwent contact metamorphism and organic carbon depletion. Such process may have released large amounts of thermogenic gases (CO2 and CH4) leading to global perturbation of the carbon cycle and to global warming. The timing of CAMP volcanic eruptions is well constrained by combined geochronologic, stratigraphic and palynologic data. In Morocco, newly observed palynological assemblages for sediments at the top of the lava piles are nearly identical to those found at the base of the volcanic sequences. These new data combined with carbon isotopic data indicate that over 95% of the CAMP lava flows in Morocco erupted during a short time interval at the very beginning of the end-Triassic extinction interval. A similar scenario applies possibly to the lava flows from North America. CAMP basalts are quite sulfur rich (up to 1800 ppm) suggesting that CAMP eruptions emitted large amounts of SO2. Such emissions lead possibly to short-lived cooling events

  19. The Middle Triassic insect radiation revealed by isotopic age and iconic fossils from NW China

    Science.gov (United States)

    Zheng, Daran; Chang, Su-Chin; Wang, He; Fang, Yan; Wang, Jun; Feng, Chongqing; Xie, Guwei; Jarzembowski, Edmund A.; Zhang, Haichun; Wang, Bo

    2017-04-01

    Following the end-Permian mass extinction, the Triassic represented an important period witnessing the recovery and radiation of marine and terrestrial ecosystems. Terrestrial plants and vertebrates have been widely investigated; however the insects, the most diverse organisms on earth, remain enigmatic due to the rarity of Early-Middle Triassic fossils. Here we report new fossils from a Ladinian deposit dated at 238-237 Ma and a Carnian deposit in northwestern China, including the earliest definite caddisfly cases (Trichoptera) and water boatmen (Hemiptera), diverse polyphagan beetles (Coleoptera) and scorpionflies (Mecoptera). Our findings suggest that the Holometabola, comprising the majority of modern-day insect species, experienced an extraordinary diversification in the Middle Triassic and was already been dominant in some Middle and Late Triassic insect faunas, after the extinction of several ecologically dominant, Paleozoic insect groups in the latest Permian and earliest Triassic. This turnover is perhaps related to notable episodes of extreme warming and drying, leading to the eventual demise of coal-swamp ecosystems, evidenced by floral turnover during this interval. The forest revival during the Middle Triassic probably stimulated the rapid radiation and evolution of insects including some key aquatic lineages which built new associations that persist to the present day. Our results provide not only new insights into the early evolution of insect diversity and ecology, but also robust evidence for the view that the Triassic is the "Dawn of the Modern World". Besides, LA-ICP-MS U-Pb dating initially gave a late Ladinian age for the Tongchuan entomnfauna after the results: 237.41 ± 0.91 Ma and 238 ± 0.97 Ma. The age is in agreement with that of the marine Ladinian-Carnian boundary, representing a novel age constraint for the terrestrial strata near this boundary. This age can provide a calibration for marine and terrestrial correlation near Ladinian

  20. Delayed recovery of non-marine tetrapods after the end-Permian mass extinction tracks global carbon cycle

    OpenAIRE

    Irmis, Randall B.; Whiteside, Jessica H.

    2011-01-01

    During the end-Permian mass extinction, marine ecosystems suffered a major drop in diversity, which was maintained throughout the Early Triassic until delayed recovery during the Middle Triassic. This depressed diversity in the Early Triassic correlates with multiple major perturbations to the global carbon cycle, interpreted as either intrinsic ecosystem or external palaeoenvironmental effects. In contrast, the terrestrial record of extinction and recovery is less clear; the effects and magn...

  1. Middle Triassic chirotherid trackways on earthquake influenced intertidal limulid reproduction flats of the European Germanic Basin coasts

    Science.gov (United States)

    Diedrich, Cajus G.

    2012-09-01

    Chirotherid footprints of Synaptichnium, Chirotherium and Isochirotherium appeared during the late Early (Aegean) to early Late (Carnian) Triassic in central Europe. These taxa are partly revised herein, using both perfect and variably preserved tracks, and very long trackways from an upper Pelsonian intertidal-flat megatracksite of the Germanic Basin coast Pelsonian (Karlstadt Formation). The global Middle Triassic distribution of those footprints suggests seasonal migrations across Pangaea of possible archosauriform reptile trackmakers, such as Euparkeria, Ticinosuchus, Arizonasaurus and Batrachotomus, caused by horseshoe-crab mass migrations into tidal-flat beach reproductive zones in the Germanic Basin. Such seasonal migrations may even suggest a Pangaea-wide food-chain reaction, possibly including the mobilization of fish, marine and terrestrial reptiles, and of which situation the Germanic Basin intertidal-flats is a globally unique example.

  2. High sedimentation rates in the Early Triassic after latest Permian mass extinction: Carbonate production is main factor in non-Arctic regions

    Science.gov (United States)

    Horacek, Micha; Brandner, Rainer

    2016-04-01

    A substantial change in sedimentation rates towards higher values has been documented from the Late Permian to the Lower Triassic. Although it is assumed and also has been shown that the deposition of siliciclastic material increased in the Lower Triassic due to stronger erosion because of loss of land cover and increased chemical and physical weathering with extreme climate warming, the main sediment production occurred by marine carbonate production. Still, carbonate production might have been significantly influenced by weathering and erosion in the hinterland, as the transport of dust by storms into the ocean water probably was a main nutrient source for microbial carbonate producers, because "normal" nutrient supply by ocean circulation, i. e. upwelling was strongly reduced due to the elevated temperatures resulting in water-column stratification . Sediment accumulation was also clearly influenced by the paleo-geographic and latitudinal position, with lower carbonate production and sedimentation rates in moderate latitudes. The existence of a "boundary clay" and microbial carbonate mounds and layers in the immediate aftermath of the latest Permian mass extinction points towards a development from a short-timed acid ocean water - resulting in a carbonate production gap and the deposition of the boundary clay towards the deposition of the microbial mounds and layers due to the microbial production of micro-environments with higher alkalinity allowing the production of carbonate. After the return of the ocean water to normal alkalinity planktic production of carbonate resulted in a very high sedimentation rate, especially taking into account the absence of carbonate producing eukaryotic algae and animals.

  3. Duration of and decoupling between carbon isotope excursions during the end-Triassic mass extinction and Central Atlantic Magmatic Province emplacement

    Science.gov (United States)

    Yager, Joyce A.; West, A. Joshua; Corsetti, Frank A.; Berelson, William M.; Rollins, Nick E.; Rosas, Silvia; Bottjer, David J.

    2017-09-01

    Changes in δ13Ccarb and δ13Corg from marine strata occur globally in association with the end-Triassic mass extinction and the emplacement of the Central Atlantic Magmatic Province (CAMP) during the break up of Pangea. As is typical in deep time, the timing and duration of these isotopic excursions has remained elusive, hampering attempts to link carbon cycle perturbations to specific processes. Here, we report δ13Ccarb and δ13Corg from Late Triassic and Early Jurassic strata near Levanto, Peru, where intercalated dated ash beds permit temporal calibration of the carbon isotope record. Both δ13Ccarb and δ13Corg exhibit a broad positive excursion through the latest Triassic into the earliest Jurassic. The first order positive excursion in δ13Corg is interrupted by a negative shift noted in many sections around the world coincident with the extinction horizon. Our data indicate that the negative excursion lasts 85 ± 25 kyrs, longer than inferred by previous studies based on cyclostratigraphy. A 260 ± 80 kyr positive δ13Corg shift follows, during which the first Jurassic ammonites appear. The overall excursion culminates in a return to pre-perturbation carbon isotopic values over the next 1090 ± 70 kyrs. Via chronologic, isotopic, and biostratigraphic correlation to other successions, we find that δ13Ccarb and δ13Corg return to pre-perturbation values as CAMP volcanism ceases and in association with the recovery of pelagic and benthic biota. However, the initiation of the carbon isotope excursion at Levanto predates the well-dated CAMP sills from North America, indicating that CAMP may have started earlier than thought based on these exposures, or that the onset of carbon cycle perturbations was not related to CAMP.

  4. Unappreciated diversification of stem archosaurs during the Middle Triassic predated the dominance of dinosaurs.

    Science.gov (United States)

    Foth, Christian; Ezcurra, Martín D; Sookias, Roland B; Brusatte, Stephen L; Butler, Richard J

    2016-09-15

    Archosauromorpha originated in the middle-late Permian, radiated during the Triassic, and gave rise to the crown group Archosauria, a highly successful clade of reptiles in terrestrial ecosystems over the last 250 million years. However, scientific attention has mainly focused on the diversification of archosaurs, while their stem lineage (i.e. non-archosaurian archosauromorphs) has often been overlooked in discussions of the evolutionary success of Archosauria. Here, we analyse the cranial disparity of late Permian to Early Jurassic archosauromorphs and make comparisons between non-archosaurian archosauromorphs and archosaurs (including Pseudosuchia and Ornithodira) on the basis of two-dimensional geometric morphometrics. Our analysis recovers previously unappreciated high morphological disparity for non-archosaurian archosauromorphs, especially during the Middle Triassic, which abruptly declined during the early Late Triassic (Carnian). By contrast, cranial disparity of archosaurs increased from the Middle Triassic into the Late Triassic, declined during the end-Triassic extinction, but re-expanded towards the end of the Early Jurassic. Our study indicates that non-archosaurian archosauromorphs were highly diverse components of terrestrial ecosystems prior to the major radiation of archosaurs, including dinosaurs, while disparity patterns of the Ladinian and Carnian indicate a gradual faunal replacement of stem archosaurs by the crown group, including a short interval of partial overlap in morphospace during the Ladinian.

  5. Middle-Upper Triassic and Middle Jurassic tetrapod track assemblages of southern Tunisia, Sahara Platform

    Science.gov (United States)

    Niedźwiedzki, Grzegorz; Soussi, Mohamed; Boukhalfa, Kamel; Gierliński, Gerard D.

    2017-05-01

    Three tetrapod track assemblages from the early-middle Mesozoic of southern Tunisia are reported. The strata exposed at the Tejra 2 clay-pit near the Medenine and Rehach site, located in the vicinity of Kirchaou, contain the first tetrapod tracks found in the Triassic of Tunisia. The Middle Jurassic (early Aalenian) dinosaur tracks are reported from the Mestaoua plain near Tataouine. In the Middle Triassic outcrop of the Tejra 2 clay-pit, tridactyl tracks of small and medium-sized dinosauromorphs, were discovered. These tracks represent the oldest evidence of dinosaur-lineage elements in the Triassic deposits of Tunisia. Similar tracks have been described from the Middle Triassic of Argentina, France and Morocco. An isolated set of the manus and pes of a quadrupedal tetrapod discovered in Late Triassic Rehach tracksite is referred to a therapsid tracemaker. The Middle Jurassic deposits of the Mestaoua plain reveal small and large tridactyl theropod dinosaur tracks (Theropoda track indet. A-C). Based on comparison with the abundant record of Triassic tetrapod ichnofossils from Europe and North America, the ichnofauna described here indicates the presence of a therapsid-dinosauromorph ichnoassociation (without typical Chirotheriidae tracks) in the Middle and Late Triassic, which sheds light on the dispersal of the Middle-Upper Triassic tetrapod ichnofaunas in this part of Gondwana. The reported Middle Jurassic ichnofauna show close similarities to dinosaur track assemblages from the Lower and Middle Jurassic of northwestern Africa, North America, Europe and also southeastern Asia. Sedimentological and lithostratigraphic data of each new tracksite have been defined on published data and new observations. Taken together, these discoveries present a tantalizing window into the evolutionary history of tetrapods from the Triassic and Jurassic of southern Tunisia. Given the limited early Mesozoic tetrapod record from the region, these discoveries are of both temporal and

  6. EARLIEST TRIASSIC CONODONTS FROM CHITRAL, NORTHERNMOST PAKISTAN

    Directory of Open Access Journals (Sweden)

    MARIA CRISTINA PERRI

    2004-07-01

    Full Text Available Extensive tracts of very shallow water carbonates in the valleys of the Yarkhun and Mastuj rivers of Chitral (northernmost Pakistan previously though to be Permian (or Cretaceous are shown by conodonts from two horizons in sequences 110 km apart—near Torman Gol (Mastuj valley and near Sakirmul (upper Yarkhun valley—to include earliest Triassic (Scythian—Induan horizons. Both faunas have Isarcicella staeschei Dai & Zhang, Is. lobata Perri, Is. turgida (Kozur et al. and Hindeodus parvus (Kozur & Pjatakova, whereas Is. Isarcica (Huckriede has been recognised only in the Torman Gol occurrence. The presence, respectively, of Is. staeschei in the Sakirmul and Is. isarcica in the Torman Gol occurrences, allows discrimination of the staeschei and isarcica zones respectively the third and the fourth conodont biozones of the Early Triassic conodont biozonation of Perri (in Perri & Farabegoli 2003. Such faunas, consisting mainly of isarcicellids and hindeodids but lacking gondolellids, are characteristic of restricted sea environments across the Permian–Triassic boundary and in the earliest Triassic in other Tethyan areas. The conodont faunas from these two occurrences are remarkably similar, nearly contemporaneous, and indicate shallow water biofacies. They are inferred to equate with the Ailak Dolomite, a sequence of Late Permian–?Late Triassic dolostones discriminated farther up the Yarkhun valley and extending eastwards into the upper Hunza region of northernmost Pakistan. The Zait Limestone and Sakirmul carbonate sequence are consistent with extension of the previously inferred Triassic carbonate platform at least 110 km farther to the SW than previously supposed.

  7. Evidence for Late Permian-Upper Triassic ocean acidification from calcium isotopes in carbonate of the Kamura section in Japan

    Science.gov (United States)

    Ye, F.; Zhao, L., Sr.; Chen, Z. Q.; Wang, X.

    2017-12-01

    Calcium and carbon cycles are tightly related in the ocean, for example, through continental weathering and deposition of carbonate, thus, very important for exploring evolutions of marine environment during the earth history. The end-Permian mass extinction is the biggest biological disaster in the Phanerozoic and there are several studies talking about variations of calcium isotopes across the Permian-Triassic boundary (PTB). However, these studies are all from the Tethys regions (Payne et al., 2010; Hinojosa et al., 2012), while the Panthalassic Ocean is still unknown to people. Moreover, evolutions of the calcium isotopes during the Early to Late Triassic is also poorly studied (Blattler et al., 2012). Here, we studied an Uppermost Permian to Upper Triassic shallow water successions (Kamura section, Southwest Japan) in the Central Panthalassic Ocean. The Kamura section is far away from the continent without any clastic pollution, therefore, could preserved reliable δ44/40Cacarb signals. Conodont zonation and carbonate carbon isotope also provide precious time framework which is necessary for the explaining of the δ44/40Cacarb profile. In Kamura, δ44/40Cacarb and δ13Ccarb both exhibit negative excursions across the PTB, the δ44/40Cacarb value in the end-Permian is 1.0398‰ then abrupt decrease to the minimum value of 0.1524‰. CO2-driven global ocean acidification best explains the coincidence of the δ44/40Cacarb excursion with negative excursions in the δ13Ccarb of carbonates until the Early Smithian(N1a, N1b, N1c, P1, N2, P2). In the Middle and the Late Triassic, the δ44/40 Cacarb average approximately 1.1‰. During the Middle and Late Triassic, strong relationships between δ44/40Cacarb and δ13Ccarb are collapsed, indicating a normal pH values of the seawater in those time. The Siberian Trap volcanism probably played a significant role on the δ44/40Cacarb until the late Early Triassic. After that, δ44/40Cacarb was mostly controlled by carbonate

  8. Discovery of a Triassic magmatic arc source for the Permo-Triassic Karakaya subduction complex, NW Turkey

    Science.gov (United States)

    Ayda Ustaömer, Petek; Ustaömer, Timur; Gerdes, Axel; Robertson, Alastair H. F.; Zulauf, Gernold

    2014-05-01

    The Permo-Triassic Karakaya Complex is well explained by northward subduction of Palaeotethys but until now no corresponding magmatic arc has been identified in the region. With the aim of determining the compositions and ages of the source units, ten sandstone samples were collected from the mappably distinct Ortaoba, Hodul, Kendirli and Orhanlar Units. Zircon grains were extracted from these sandstones and >1300 were dated by the U-Pb method and subsequently analysed for the Lu-Hf isotopic compositions by LA-MC-ICPMS at Goethe University, Frankfurt. The U-Pb-Hf isotope systematics are indicative of two different sediment provenances. The first, represented by the Ortaoba, Hodul and Kendirli Units, is dominated by igneous rocks of Triassic (250-220 Ma), Early Carboniferous-Early Permian (290-340 Ma) and Early to Mid-Devonian (385-400 Ma) ages. The second provenance, represented by the Orhanlar Unit, is indicative of derivation from a peri-Gondwanan terrane. In case of the first provenance, the Devonian and Carboniferous source rocks exibit intermediate eHf(t) values (-11 to -3), consistent with the formation at a continental margin where juvenile mantle-derived magmas mixed with (recycled) old crust having Palaeoproterozoic Hf model ages. In contrast, the Triassic arc magma exhibits higher eHf(t) values (-6 to +6), consistent with the mixing of juvenile mantle-derived melts with (recycled) old crust perhaps somewhat rejuvanated during the Cadomian period. We have therefore identified a Triassic magmatic arc as predicted by the interpretation of the Karakaya Complex as an accretionary complex related to northward subduction (Carboniferous and Devonian granites are already well documented in NW Turkey). Possible explanations for the lack of any outcrop of the source magmatic arc are that it was later subducted or the Karakaya Complex was displaced laterally from its source arc (both post 220 Ma). Strike-slip displacement (driven by oblique subduction?) can also

  9. Geochemical characteristics of natural gas in the hydrocarbon accumulation history, and its difference among gas reservoirs in the Upper Triassic formation of Sichuan Basin, China

    Directory of Open Access Journals (Sweden)

    Peng Wang

    2016-08-01

    Full Text Available The analysis of hydrocarbon generation, trap formation, inclusion homogenization temperature, authigenic illite dating, and ESR dating were used to understand the history of hydrocarbon accumulation and its difference among gas reservoirs in the Upper Triassic formation of Sichuan Basin. The results show the hydrocarbon accumulation mainly occurred during the Jurassic and Cretaceous periods; they could also be classified into three stages: (1 early hydrocarbon generation accumulation stage, (2 mass hydrocarbon generation accumulation stage before the Himalayan Epoch, (3 and parts of hydrocarbon adjustment and re-accumulation during Himalayan Epoch. The second stage is more important than the other two. The Hydrocarbon accumulation histories are obviously dissimilar in different regions. In western Sichuan Basin, the gas accumulation began at the deposition period of member 5 of Xujiahe Formation, and mass accumulation occurred during the early Middle Jurassic up to the end of the Late Cretaceous. In central Sichuan Basin, the accumulation began at the early Late Jurassic, and the mass accumulation occurred from the middle Early Cretaceous till the end of the Late Cretaceous. In southern Sichuan Basin, the accumulation began at the middle Late Jurassic, and the mass accumulation occurred from the middle of the Late Cretaceous to the end of the Later Cretaceous. The accumulation history of the western Sichuan Basin is the earliest, and the southern Sichuan Basin is the latest. This paper will help to understand the accumulation process, accumulation mechanism, and gas reservoir distribution of the Triassic gas reservoirs in the Sichuan Basin better. Meanwhile, it is found that the authigenic illite in the Upper Triassic formation of Sichuan Basin origin of deep-burial and its dating is a record of the later accumulation. This suggests that the illite dating needs to fully consider illite origin; otherwise the dating results may not accurately

  10. Mass-wasting triggered by the end-Triassic mass-extinction

    Science.gov (United States)

    van de Schootbrugge, Bas; Vecoli, Marco; Strother, Paul; Lindstrom, Sofie; Oschmann, Wolfgang

    2014-05-01

    The end-Triassic dieback of tree-forming vegetation across NW Europe and the proliferation of a low-growing herbaceous pioneer vegetation composed of ferns and fern allies, likely had a major impact on weathering and erosion of emerged land masses. In a recently drilled core from northern Germany (Schandelah), palynological analyses provide evidence for this scenario. The uppermost Rhaetian Triletes Beds show increasing amounts of reworked Palaeozoic acritarchs and prasinophytes of up to 30% of the palynomorph fraction. Most of the acritarchs are singletons and can be assigned to Ordovician and Silurian species, such as Ankyrotrochus crispum, Oppilatala eoplanktonica, and Evittia spp. The average age of the reworked acritarch assemblages is observed to increase during the latest Rhaetian, leading to an inverted stratigraphy among Palaeozoic species. Further North, in the Stenlille cores from the Danish Basin, reworked Palaeozoic palynomorphs appear to constitute mainly sphaeromorphic prasinophytes and other Palaeozoic microfossils such as chitinozoans and carboniferous spores. Further south, at Mingolsheim (S Germany) the Triletes Beds contain a clear sign of soil reworking, including mycorrhizal fungal remains and cysts from probable soil organisms. These peculiar changes in palynological assemblages go hand-in-hand with important changes in sedimentology. The reworking of soil and bedrock is occurring in an interval that also contains evidence for earthquake activity in the form of widespread seismites. All these observations may be attributed to a number of mutually non-exclusive mechanisms, including decreased plant cover, an intensified hydrological cycle due to greenhouse warming, and the doming of the Central Atlantic Magmatic Province leading to continental-scale tectonic steepening of basin margins.

  11. Boreal earliest Triassic biotas elucidate globally depauperate hard substrate communities after the end-Permian mass extinction.

    Science.gov (United States)

    Zatoń, Michał; Niedźwiedzki, Grzegorz; Blom, Henning; Kear, Benjamin P

    2016-11-08

    The end-Permian mass extinction constituted the most devastating biotic crisis of the Phanerozoic. Its aftermath was characterized by harsh marine conditions incorporating volcanically induced oceanic warming, widespread anoxia and acidification. Bio-productivity accordingly experienced marked fluctuations. In particular, low palaeolatitude hard substrate communities from shallow seas fringing Western Pangaea and the Tethyan Realm were extremely impoverished, being dominated by monogeneric colonies of filter-feeding microconchid tubeworms. Here we present the first equivalent field data for Boreal hard substrate assemblages from the earliest Triassic (Induan) of East Greenland. This region bordered a discrete bio-realm situated at mid-high palaeolatitude (>30°N). Nevertheless, hard substrate biotas were compositionally identical to those from elsewhere, with microconchids encrusting Claraia bivalves and algal buildups on the sea floor. Biostratigraphical correlation further shows that Boreal microconchids underwent progressive tube modification and unique taxic diversification concordant with changing habitats over time. We interpret this as a post-extinction recovery and adaptive radiation sequence that mirrored coeval subequatorial faunas, and thus confirms hard substrate ecosystem depletion as a hallmark of the earliest Triassic interval globally.

  12. Carbon cycle changes during the Triassic-Jurassic transition

    NARCIS (Netherlands)

    Ruhl, M.|info:eu-repo/dai/nl/304838357

    2010-01-01

    The end-Triassic is regarded as one of the five major mass extinction events of the Phanerozoic. This time interval is marked by up to 50% of marine biodiversity loss and major changes in terrestrial ecosystems. Mass extinction events are often marked by changes in the global carbon cycle. The

  13. Anisian (Middle Triassic) marine ichnocoenoses from the eastern and western margins of the Kamdian Continent, Yunnan Province, SW China: Implications for the Triassic biotic recovery

    Science.gov (United States)

    Feng, Xueqian; Chen, Zhong-Qiang; Woods, Adam; Pei, Yu; Wu, Siqi; Fang, Yuheng; Luo, Mao; Xu, Yaling

    2017-10-01

    Two Anisian (Middle Triassic) marine ichnocoenoses are reported from the Boyun and Junmachang (JMC) sections located along the eastern and western margins of the Kamdian Continent, Yunnan Province, Southwest China, respectively. The Boyun ichnoassemblage is middle Anisian in age and is dominated by robust Rhizocorallium, while the JMC ichnoassemblage is of an early Anisian age and is characterized by the presence of Zoophycos. The ichnoassemblage horizons of the Boyun section represent an inner ramp environment, while the JMC section was likely situated in a mid-ramp setting near storm wave base as indicated by the presence of tempestites. The ichnofossil-bearing successions are usually highly bioturbated in both the Boyun (BI 3-5, BPBI 5) and JMC (BI 3-4, BPBI 3-4) sections. Three large, morphologically complicated ichnogenera: 1) Rhizocorallium; 2) Thalassinoides; and, 3) Zoophycos characterize the Anisian ichnocoenoses. Of these, Rhizocorallium has mean and maximum tube diameters up to 20.4 mm and 28 mm, respectively, while Thalassinoides mean and maximum tube diameters are 14.2 mm and 22 mm, respectively. Zoophycos is present in the early Anisian strata of the JMC section, and represents the oldest known occurrence of this ichnogenus following the latest Permian mass extinction. Similar to coeval ichnoassemblages elsewhere in the world, the Yunnan ichnocoenoses embrace a relatively low ichnodiversity, but their burrows usually penetrate deeply into the sediment, and include large and complex Rhizocorallium and Thalassinoides. All of these ichnologic features are indicative of recovery stage 4 after the latest Permian crisis. Anisian ichnoassemblages occur globally in six different habitat settings, and all show similar ecologic characteristics except for slightly different degrees of ichnotaxonomic richness, indicating that depositional environment is not a crucial factor shaping the recovery of the trace-makers, but may have an impact on their ichnodiversity

  14. A carapace-like bony 'body tube' in an early triassic marine reptile and the onset of marine tetrapod predation.

    Science.gov (United States)

    Chen, Xiao-hong; Motani, Ryosuke; Cheng, Long; Jiang, Da-yong; Rieppel, Olivier

    2014-01-01

    Parahupehsuchus longus is a new species of marine reptile from the Lower Triassic of Yuan'an County, Hubei Province, China. It is unique among vertebrates for having a body wall that is completely surrounded by a bony tube, about 50 cm long and 6.5 cm deep, comprising overlapping ribs and gastralia. This tube and bony ossicles on the back are best interpreted as anti-predatory features, suggesting that there was predation pressure upon marine tetrapods in the Early Triassic. There is at least one sauropterygian that is sufficiently large to feed on Parahupehsuchus in the Nanzhang-Yuan'an fauna, together with six more species of potential prey marine reptiles with various degrees of body protection. Modern predators of marine tetrapods belong to the highest trophic levels in the marine ecosystem but such predators did not always exist through geologic time. The indication of marine-tetrapod feeding in the Nanzhang-Yuan'an fauna suggests that such a trophic level emerged for the first time in the Early Triassic. The recovery from the end-Permian extinction probably proceeded faster than traditionally thought for marine predators. Parahupehsuchus has superficially turtle-like features, namely expanded ribs without intercostal space, very short transverse processes, and a dorsal outgrowth from the neural spine. However, these features are structurally different from their turtle counterparts. Phylogeny suggests that they are convergent with the condition in turtles, which has a fundamentally different body plan that involves the folding of the body wall. Expanded ribs without intercostal space evolved at least twice and probably even more among reptiles.

  15. A carapace-like bony 'body tube' in an early triassic marine reptile and the onset of marine tetrapod predation.

    Directory of Open Access Journals (Sweden)

    Xiao-hong Chen

    Full Text Available Parahupehsuchus longus is a new species of marine reptile from the Lower Triassic of Yuan'an County, Hubei Province, China. It is unique among vertebrates for having a body wall that is completely surrounded by a bony tube, about 50 cm long and 6.5 cm deep, comprising overlapping ribs and gastralia. This tube and bony ossicles on the back are best interpreted as anti-predatory features, suggesting that there was predation pressure upon marine tetrapods in the Early Triassic. There is at least one sauropterygian that is sufficiently large to feed on Parahupehsuchus in the Nanzhang-Yuan'an fauna, together with six more species of potential prey marine reptiles with various degrees of body protection. Modern predators of marine tetrapods belong to the highest trophic levels in the marine ecosystem but such predators did not always exist through geologic time. The indication of marine-tetrapod feeding in the Nanzhang-Yuan'an fauna suggests that such a trophic level emerged for the first time in the Early Triassic. The recovery from the end-Permian extinction probably proceeded faster than traditionally thought for marine predators. Parahupehsuchus has superficially turtle-like features, namely expanded ribs without intercostal space, very short transverse processes, and a dorsal outgrowth from the neural spine. However, these features are structurally different from their turtle counterparts. Phylogeny suggests that they are convergent with the condition in turtles, which has a fundamentally different body plan that involves the folding of the body wall. Expanded ribs without intercostal space evolved at least twice and probably even more among reptiles.

  16. Late Triassic paleolatitude of the Qiangtang block: Implications for the closure of the Paleo-Tethys Ocean

    Science.gov (United States)

    Song, Peiping; Ding, Lin; Li, Zhenyu; Lippert, Peter C.; Yang, Tianshui; Zhao, Xixi; Fu, Jiajun; Yue, Yahui

    2015-08-01

    To better constrain the Late Triassic paleolatitude of the Qiangtang block and the closure of the Paleo-Tethys Ocean, a combined paleomagnetic and zircon U/Pb geochronological study has been conducted on the Upper Triassic Jiapila Formation volcanic rocks on the northern edge of the Qiangtang block of Central Tibet (34.1°N, 92.4°E). These rocks are dated to 204-213 Ma. Progressive thermal or alternating field demagnetization successfully isolated stable characteristic remanent magnetizations (ChRM) that pass both the fold and reversal tests, consistent with a primary magnetization. These are the first volcanic-based paleomagnetic results from pre-Cretaceous rocks of the Qiangtang block that appear to average secular variation well enough to yield a reliable paleolatitude estimate. Based on our new paleomagnetic data from Upper Triassic lavas, we conclude that the Late Triassic pole of the Qiangtang block was located at 64.0°N, 174.7°E, with A95 = 6.6 ° (N = 29). We compile published paleomagnetic data from the Qiangtang block to calculate a Late Triassic latitude for the Qiangtang block at 31.7 ± 3.0°N. The central Paleo-Tethys Ocean basin was located between the North China (NCB) and Tarim blocks to the north and the Qiangtang block to the south during Late Paleozoic-Early Mesozoic. A comparison of published Early Triassic paleopole from the Qiangtang block with the coeval paleopoles from the NCB and Tarim indicates that the Paleo-Tethys Ocean could not have closed during the Early Triassic and that its width was approximately ∼32-38° latitude (∼3500-4200 km). However, the comparison of our new combined Late Triassic paleomagnetic result with the Late Triassic poles of the NCB and Tarim, as well as numerous geological observations, indicates that the closure of the Paleo-Tethys Ocean at the longitude of the Qiangtang block most likely occurred during the Late Triassic.

  17. Evolution of the Early Triassic marine depositional environment in the Croatian Dinarides

    Science.gov (United States)

    Aljinović, Dunja; Smirčić, Duje; Horacek, Micha; Richoz, Sylvain; Krystyn, Leopold; Kolar-Jurkovšek, Tea; Jurkovšek, Bogdan

    2014-05-01

    In the central part of the Dinarides in Croatia, the Early Triassic depositional sequence was investigated by means of litho-, bio- and chemostratigraphy at locality Plavno (ca. 1.000m thick). Conodont and δ13C-isotope analysis were a powerfull tool to determine stage and substage boundaries. The succession begins with the second conodont zone of the Griesbachian Isarcicella staeschei and I. isarcica with low δ13C-values and a steadily increase towards the Griesbachian-Dienerian boundary. Around that boundary a minor, short, negative excursion occurs. In the Dienerian the δ13C-values increase with a steepening of the slope towards the Dienerian-Smithian boundary. Around that boundary a maximum of +5o in shallow water carbonate occurs followed by a steep and continuous drop to low, often negative values in the Smithian. Just before the Smithian-Spathian boundary a steep rise to a second maximum is documented. It is followed by decline in the Spathian and a gentle increase to a rounded peak at the Spathian-Anisian boundary. In lithological sense Plavno succession has threefold division: 1) carbonates representing the oldest Early Triassic strata (early Griesbachian); 2) dominantly red clastics (shales, siltstones and sandstones) with intercalation of oncoid/ooid or bioclast rich grainstones (uppermost Griesbachian, Dienerian and Smithian) and 3) dominantly grey carbonaceous lime mudstones, marls and calcisiltites with ammonoids representing Spathian strata. In the oldest strata (Griesbachian) in macrocrystalline subhedral dolomites rare microspheres and foraminifers Earlandia and Cornuspira point to the stressful conditions related to the end Permian mass extinction. In the uppermost Griesbachian and Dienerian strata, within dominantly clastic deposition, rare coarse oncoliths with typical microbial cortices occur. Their presence fits to the interpretation of biotical-induced precipitation related to PTB extinction and can suggest still stressful condition. The

  18. A new sphenodontian (Lepidosauria: Rhynchocephalia) from the Late Triassic of Argentina and the early origin of the herbivore opisthodontians

    Science.gov (United States)

    Martínez, Ricardo N.; Apaldetti, Cecilia; Colombi, Carina E.; Praderio, Angel; Fernandez, Eliana; Malnis, Paula Santi; Correa, Gustavo A.; Abelin, Diego; Alcober, Oscar

    2013-01-01

    Sphenodontians were a successful group of rhynchocephalian reptiles that dominated the fossil record of Lepidosauria during the Triassic and Jurassic. Although evidence of extinction is seen at the end of the Laurasian Early Cretaceous, they appeared to remain numerically abundant in South America until the end of the period. Most of the known Late Cretaceous record in South America is composed of opisthodontians, the herbivorous branch of Sphenodontia, whose oldest members were until recently reported to be from the Kimmeridgian–Tithonian (Late Jurassic). Here, we report a new sphenodontian, Sphenotitan leyesi gen. et sp. nov., collected from the Upper Triassic Quebrada del Barro Formation of northwestern Argentina. Phylogenetic analysis identifies Sphenotitan as a basal member of Opisthodontia, extending the known record of opisthodontians and the origin of herbivory in this group by 50 Myr. PMID:24132307

  19. Systematics of putative euparkeriids (Diapsida: Archosauriformes from the Triassic of China

    Directory of Open Access Journals (Sweden)

    Roland B. Sookias

    2014-11-01

    Full Text Available The South African species Euparkeria capensis is of great importance for understanding the early radiation of archosauromorphs (including archosaurs following the Permo–Triassic mass extinction, as most phylogenetic analyses place it as the sister taxon to crown group Archosauria within the clade Archosauriformes. Although a number of species from Lower–Middle Triassic deposits worldwide have been referred to the putative clade Euparkeriidae, the monophyly of Euparkeriidae is controversial and has yet to be demonstrated by quantitative phylogenetic analysis. Three Chinese taxa have been recently suggested to be euparkeriids: Halazhaisuchus qiaoensis, ‘Turfanosuchus shageduensis’, and Wangisuchus tzeyii, all three of which were collected from the Middle Triassic Ermaying Formation of northern China. Here, we reassess the taxonomy and systematics of these taxa. We regard Wangisuchus tzeyii as a nomen dubium, because the holotype is undiagnostic and there is no convincing evidence that the previously referred additional specimens represent the same taxon as the holotype. We also regard ‘Turfanosuchus shageduensis’ as a nomen dubium as we are unable to identify any diagnostic features. We refer the holotype to Archosauriformes, and more tentatively to Euparkeriidae. Halazhaisuchus qiaoensis and the holotype of ‘Turfanosuchus shageduensis’ are resolved as sister taxa in a phylogenetic analysis, and are in turn the sister taxon to Euparkeria capensis, forming a monophyletic Euparkeriidae that is the sister to Archosauria+Phytosauria. This is the first quantitative phylogenetic analysis to recover a non-monospecific, monophyletic Euparkeriidae, but euparkeriid monophyly is only weakly supported and will require additional examination. Given their similar sizes, stratigraphic positions and phylogenetic placement, the holotype of ‘Turfanosuchus shageduensis’ may represent a second individual of Halazhaisuchus qiaoensis, but no

  20. A Carapace-Like Bony ‘Body Tube’ in an Early Triassic Marine Reptile and the Onset of Marine Tetrapod Predation

    Science.gov (United States)

    Chen, Xiao-hong; Motani, Ryosuke; Cheng, Long; Jiang, Da-yong; Rieppel, Olivier

    2014-01-01

    Parahupehsuchus longus is a new species of marine reptile from the Lower Triassic of Yuan’an County, Hubei Province, China. It is unique among vertebrates for having a body wall that is completely surrounded by a bony tube, about 50 cm long and 6.5 cm deep, comprising overlapping ribs and gastralia. This tube and bony ossicles on the back are best interpreted as anti-predatory features, suggesting that there was predation pressure upon marine tetrapods in the Early Triassic. There is at least one sauropterygian that is sufficiently large to feed on Parahupehsuchus in the Nanzhang-Yuan’an fauna, together with six more species of potential prey marine reptiles with various degrees of body protection. Modern predators of marine tetrapods belong to the highest trophic levels in the marine ecosystem but such predators did not always exist through geologic time. The indication of marine-tetrapod feeding in the Nanzhang-Yuan’an fauna suggests that such a trophic level emerged for the first time in the Early Triassic. The recovery from the end-Permian extinction probably proceeded faster than traditionally thought for marine predators. Parahupehsuchus has superficially turtle-like features, namely expanded ribs without intercostal space, very short transverse processes, and a dorsal outgrowth from the neural spine. However, these features are structurally different from their turtle counterparts. Phylogeny suggests that they are convergent with the condition in turtles, which has a fundamentally different body plan that involves the folding of the body wall. Expanded ribs without intercostal space evolved at least twice and probably even more among reptiles. PMID:24718682

  1. Changing palaeoenvironments and tetrapod populations in the Daptocephalus Assemblage Zone (Karoo Basin, South Africa) indicate early onset of the Permo-Triassic mass extinction

    Science.gov (United States)

    Viglietti, Pia A.; Smith, Roger M. H.; Rubidge, Bruce S.

    2018-02-01

    Important palaeoenvironmental differences are identified during deposition of the latest Permian Daptocephalus Assemblage Zone (DaAZ) of the South African Beaufort Group (Karoo Supergoup), which is also divided into a Lower and Upper subzone. A lacustrine floodplain facies association showing evidence for higher water tables and subaqueous conditions on the floodplains is present in Lower DaAZ. The change to well-drained floodplain facies association in the Upper DaAZ is coincident with a faunal turnover as evidenced by the last appearance of the dicynodont Dicynodon lacerticeps, the therocephalian Theriognathus microps, the cynodont Procynosuchus delaharpeae, and first appearance of the dicynodont Lystrosaurus maccaigi within the Ripplemead member. Considering the well documented 3-phased extinction of Karoo tetrapods during the Permo-Triassic Mass Extinction (PTME), the facies transition between the Lower and Upper DaAZ represents earlier than previously documented palaeoenvironmental changes associated with the onset of this major global biotic crisis.

  2. Taphonomy of Early Triassic fish fossils of the Vega-Phroso Siltstone Member of the Sulphur Mountain Formation near Wapiti Lake, British Columbia, Canada

    Directory of Open Access Journals (Sweden)

    Karen Anderson

    2013-10-01

    Full Text Available The taphonomy of fishes living in lacustrine environments has been extensively studied in both the laboratory and the fossil record; the taphonomy of marine fishes, however, is poorly known. Triassic marine fishes with heavy ganoid and cosmoid scales, which provided protection from rapid taphonomic loss, offer a means to examine marine fish taphonomy in the fossil record. Four genera of Early Triassic fishes (the ray-finned actinopterygians Albertonia, Bobasatrania, Boreosomus, and the lobe-finned coelacanth (sarcopterygian, Whiteia from the Wapiti Lake, British Columbia locality of the Lower Triassic Sulphur Mountain Formation were examined in order to gain a better understanding of the taphonomy of fish in marine environments, determine ambient environmental conditions in the region during the Early Triassic, and ascertain the habitat and mode of life of the fish. Results indicate that environmental conditions that contributed to the preservation of the fossil fishes of the current study included deposition in deep, quiet waters, which reduced the odds of disarticulation, colder waters under higher pressure, which slowed decay and limited postmortem floatation, and waters that were anoxic, which discouraged predators and scavengers. In addition, the thickness of the primitive ganoid and cosmoid scales of the fossil fishes also increased their preservation potential. Taphonomic, physiological and environmental indicators suggest that Whiteia, Albertonia, and possibly Bobasatrania lived in deep, cold waters near the oxygen minimum zone, while Boreosomus lived higher in the water column. While the anatomical and physiological characteristics of modern fishes will likely continue to inhibit marine taphonomy studies, examination of ancient fish, particularly those with ganoid or cosmoid scales, may provide future avenues of research to gain a better understanding of marine fish taphonomy and provide a powerful tool to examine ancient fish behavior

  3. Absence of suction feeding ichthyosaurs and its implications for triassic mesopelagic paleoecology.

    Science.gov (United States)

    Motani, Ryosuke; Ji, Cheng; Tomita, Taketeru; Kelley, Neil; Maxwell, Erin; Jiang, Da-yong; Sander, Paul Martin

    2013-01-01

    Mesozoic marine reptiles and modern marine mammals are often considered ecological analogs, but the extent of their similarity is largely unknown. Particularly important is the presence/absence of deep-diving suction feeders among Mesozoic marine reptiles because this would indicate the establishment of mesopelagic cephalopod and fish communities in the Mesozoic. A recent study suggested that diverse suction feeders, resembling the extant beaked whales, evolved among ichthyosaurs in the Triassic. However, this hypothesis has not been tested quantitatively. We examined four osteological features of jawed vertebrates that are closely linked to the mechanism of suction feeding, namely hyoid corpus ossification/calcification, hyobranchial apparatus robustness, mandibular bluntness, and mandibular pressure concentration index. Measurements were taken from 18 species of Triassic and Early Jurassic ichthyosaurs, including the presumed suction feeders. Statistical comparisons with extant sharks and marine mammals of known diets suggest that ichthyosaurian hyobranchial bones are significantly more slender than in suction-feeding sharks or cetaceans but similar to those of ram-feeding sharks. Most importantly, an ossified hyoid corpus to which hyoid retractor muscles attach is unknown in all but one ichthyosaur, whereas a strong integration of the ossified corpus and cornua of the hyobranchial apparatus has been identified in the literature as an important feature of suction feeders. Also, ichthyosaurian mandibles do not narrow rapidly to allow high suction pressure concentration within the oral cavity, unlike in beaked whales or sperm whales. In conclusion, it is most likely that Triassic and Early Jurassic ichthyosaurs were 'ram-feeders', without any beaked-whale-like suction feeder among them. When combined with the inferred inability for dim-light vision in relevant Triassic ichthyosaurs, the fossil record of ichthyosaurs does not suggest the establishment of modern

  4. Source and Extent of Volcanic Ashes at the Permian-Triassic Boundary in South China and Its implications

    Science.gov (United States)

    Wang, M.; Zhong, Y. T.; Hou, Y. L.; He, B.

    2017-12-01

    Highly correlated with the Permian-Triassic Boundary (PTB) Mass Extinction in stratigraphic section, volcanic ashes around the P-T Boundary in South China have been suggested to be a likely cause of the PTB Mass Extinction. So the nature, source and extent of these volcanic ashes have great significance in figuring out the cause of the PTB Mass Extinction. In this study, we attempt to constrain the source and extent of the PTB volcanic ashes in South China by studying pyroclastic sedimentary rocks and the spatial distribution of tuffs and ashes in South China. The detrital zircons of tuffaceous sandstones from Penglaitan section yield an age spectrum peaked at 252Ma, with ɛHf(t) values varying from -20 to -5 ,and have Nb/Hf, Th/Nb and Hf/Th ratios similar to those from arc/orogenic-related settings. Coarse tuffaceous sandstones imply that their source is in limited distance. Those pyroclastic sedimentary rocks in Penglaitan are well correlated with the PTB volcanic ashes in Meishan GSSP section in stratigraphy. In the spatial distribution, pyroclastic sedimentary rocks and tuffs distribute only in southwest of South China, while finer volcanic ashes are mainly in the northern part. This spatial distribution suggests the source of tuffs and ashes was to the south or southwest of South China. Former studies especially that of Permian-Triassic magmatism in Hainan Island have supported the existence of a continental arc related to the subduction and closure of Palaeo-Tethys on the southwestern margin of South China during Permian to early Triassic. It is suggested that the PTB ashes possibly derived from this Paleo-Tethys continental arc. The fact that volcanic ashes haven't been reported or found in PTB stratum in North China or Northwest China implies a limited extent of the volcanism, which thus is too small to cause the PTB mass extinction.

  5. A Middle Triassic thoracopterid from China highlights the evolutionary origin of overwater gliding in early ray-finned fishes.

    Science.gov (United States)

    Xu, Guang-Hui; Zhao, Li-Jun; Shen, Chen-Chen

    2015-01-01

    Gliding adaptations in thoracopterid flying fishes represent a remarkable case of convergent evolution of overwater gliding strategy with modern exocoetid flying fishes, but the evolutionary origin of this strategy was poorly known in the thoracopterids because of lack of transitional forms. Until recently, all thoracopterids, from the Late Triassic of Austria and Italy and the Middle Triassic of South China, were highly specialized 'four-winged' gliders in having wing-like paired fins and an asymmetrical caudal fin with the lower caudal lobe notably larger than the upper lobe. Here, we show that the new genus Wushaichthys and the previously alleged 'peltopleurid' Peripeltopleurus, from the Middle Triassic (Ladinian, 235-242 Ma) of South China and near the Ladinian/Anisian boundary of southern Switzerland and northern Italy, respectively, represent the most primitive and oldest known thoracopterids. Wushaichthys, the most basal thoracopterid, shows certain derived features of this group in the skull. Peripeltopleurus shows a condition intermediate between Wushaichthys and Thoracopterus in having a slightly asymmetrical caudal fin but still lacking wing-like paired fins. Phylogenetic studies suggest that the evolution of overwater gliding of thoracopterids was gradual in nature; a four-stage adaption following the 'cranial specialization-asymmetrical caudal fin-enlarged paired fins-scale reduction' sequence has been recognized in thoracopterid evolution. Moreover, Wushaichthys and Peripeltopleurus bear hooklets on the anal fin of supposed males, resembling those of modern viviparious teleosts. Early thoracopterids probably had evolved a live-bearing reproductive strategy. © 2015 The Author(s) Published by the Royal Society. All rights reserved.

  6. Assessing the record and causes of Late Triassic extinctions

    Science.gov (United States)

    Tanner, L.H.; Lucas, S.G.; Chapman, M.G.

    2004-01-01

    Accelerated biotic turnover during the Late Triassic has led to the perception of an end-Triassic mass extinction event, now regarded as one of the "big five" extinctions. Close examination of the fossil record reveals that many groups thought to be affected severely by this event, such as ammonoids, bivalves and conodonts, instead were in decline throughout the Late Triassic, and that other groups were relatively unaffected or subject to only regional effects. Explanations for the biotic turnover have included both gradualistic and catastrophic mechanisms. Regression during the Rhaetian, with consequent habitat loss, is compatible with the disappearance of some marine faunal groups, but may be regional, not global in scale, and cannot explain apparent synchronous decline in the terrestrial realm. Gradual, widespread aridification of the Pangaean supercontinent could explain a decline in terrestrial diversity during the Late Triassic. Although evidence for an impact precisely at the boundary is lacking, the presence of impact structures with Late Triassic ages suggests the possibility of bolide impact-induced environmental degradation prior to the end-Triassic. Widespread eruptions of flood basalts of the Central Atlantic Magmatic Province (CAMP) were synchronous with or slightly postdate the system boundary; emissions of CO2 and SO2 during these eruptions were substantial, but the contradictory evidence for the environmental effects of outgassing of these lavas remains to be resolved. A substantial excursion in the marine carbon-isotope record of both carbonate and organic matter suggests a significant disturbance of the global carbon cycle at the system boundary. Release of methane hydrates from seafloor sediments is a possible cause for this isotope excursion, although the triggering mechanism and climatic effects of such a release remain uncertain. ?? 2003 Elsevier B.V. All rights reserved.

  7. Geographic range did not confer resilience to extinction in terrestrial vertebrates at the end-Triassic crisis.

    Science.gov (United States)

    Dunhill, Alexander M; Wills, Matthew A

    2015-08-11

    Rates of extinction vary greatly through geological time, with losses particularly concentrated in mass extinctions. Species duration at other times varies greatly, but the reasons for this are unclear. Geographical range correlates with lineage duration amongst marine invertebrates, but it is less clear how far this generality extends to other groups in other habitats. It is also unclear whether a wide geographical distribution makes groups more likely to survive mass extinctions. Here we test for extinction selectivity amongst terrestrial vertebrates across the end-Triassic event. We demonstrate that terrestrial vertebrate clades with larger geographical ranges were more resilient to extinction than those with smaller ranges throughout the Triassic and Jurassic. However, this relationship weakened with increasing proximity to the end-Triassic mass extinction, breaking down altogether across the event itself. We demonstrate that these findings are not a function of sampling biases; a perennial issue in studies of this kind.

  8. The timing and pattern of biotic recovery following the end-Permian mass extinction

    Science.gov (United States)

    Chen, Zhong-Qiang; Benton, Michael J.

    2012-06-01

    The aftermath of the great end-Permian period mass extinction 252 Myr ago shows how life can recover from the loss of >90% species globally. The crisis was triggered by a number of physical environmental shocks (global warming, acid rain, ocean acidification and ocean anoxia), and some of these were repeated over the next 5-6 Myr. Ammonoids and some other groups diversified rapidly, within 1-3 Myr, but extinctions continued through the Early Triassic period. Triassic ecosystems were rebuilt stepwise from low to high trophic levels through the Early to Middle Triassic, and a stable, complex ecosystem did not re-emerge until the beginning of the Middle Triassic, 8-9 Myr after the crisis. A positive aspect of the recovery was the emergence of entirely new groups, such as marine reptiles and decapod crustaceans, as well as new tetrapods on land, including -- eventually -- dinosaurs. The stepwise recovery of life in the Triassic could have been delayed either by biotic drivers (complex multispecies interactions) or physical perturbations, or a combination of both. This is an example of the wider debate about the relative roles of intrinsic and extrinsic drivers of large-scale evolution.

  9. Absence of suction feeding ichthyosaurs and its implications for triassic mesopelagic paleoecology.

    Directory of Open Access Journals (Sweden)

    Ryosuke Motani

    Full Text Available Mesozoic marine reptiles and modern marine mammals are often considered ecological analogs, but the extent of their similarity is largely unknown. Particularly important is the presence/absence of deep-diving suction feeders among Mesozoic marine reptiles because this would indicate the establishment of mesopelagic cephalopod and fish communities in the Mesozoic. A recent study suggested that diverse suction feeders, resembling the extant beaked whales, evolved among ichthyosaurs in the Triassic. However, this hypothesis has not been tested quantitatively. We examined four osteological features of jawed vertebrates that are closely linked to the mechanism of suction feeding, namely hyoid corpus ossification/calcification, hyobranchial apparatus robustness, mandibular bluntness, and mandibular pressure concentration index. Measurements were taken from 18 species of Triassic and Early Jurassic ichthyosaurs, including the presumed suction feeders. Statistical comparisons with extant sharks and marine mammals of known diets suggest that ichthyosaurian hyobranchial bones are significantly more slender than in suction-feeding sharks or cetaceans but similar to those of ram-feeding sharks. Most importantly, an ossified hyoid corpus to which hyoid retractor muscles attach is unknown in all but one ichthyosaur, whereas a strong integration of the ossified corpus and cornua of the hyobranchial apparatus has been identified in the literature as an important feature of suction feeders. Also, ichthyosaurian mandibles do not narrow rapidly to allow high suction pressure concentration within the oral cavity, unlike in beaked whales or sperm whales. In conclusion, it is most likely that Triassic and Early Jurassic ichthyosaurs were 'ram-feeders', without any beaked-whale-like suction feeder among them. When combined with the inferred inability for dim-light vision in relevant Triassic ichthyosaurs, the fossil record of ichthyosaurs does not suggest the

  10. Kerogen morphology and geochemistry at the Permian-Triassic transition in the Meishan section, South China: Implication for paleoenvironmental variation

    Science.gov (United States)

    Sawada, Ken; Kaiho, Kunio; Okano, Kazuki

    2012-08-01

    Detailed fluorescent microscopic observations and organic geochemical analyses for insoluble sedimentary organic matter (kerogens) are conducted on the end-Permian to earliest Triassic sediments in the Meishan section A of South China. The main objectives of the present study are to reconstruct variations of marine and terrestrial environments, and to evaluate bulk characteristics of terrestrial input in the palaeo-Tethys ocean for the Permian-Triassic boundary (PTB). Most of kerogens in the Meishan section are mainly composed of marine algae-derived amorphous organic matter, while terrestrial plant-derived amorphous organic matter is remarkably dominant in the mass extinction horizon reported previously. The relative abundances of marine organic matter may vary depending on marine production rather than terrestrial input in the palaeo-Tethys associated with changing terrestrial vegetation. We also identified aromatic furans as major compounds in kerogen pyrolysate of all layers. It is possible that sources of aromatic furans with alkyl group, fungi and lichen, proliferated as disaster biota in terrestrial ecosystem through the PTB. Higher abundances of herbaceous organic matter are observed in the layers above the mass extinction horizon. However, the conifer biomarker retene can be identified in kerogen pyrolysates of all layers. These results imply that the productions of herbaceous plants increased as dominant pioneer biota in early stage of recovery for terrestrial ecosystem after its collapse, but also that woody plant potentially continued to be produced in land area throughout the end-Permian and earliest-Triassic.

  11. A basal ichthyosauriform with a short snout from the Lower Triassic of China.

    Science.gov (United States)

    Motani, Ryosuke; Jiang, Da-Yong; Chen, Guan-Bao; Tintori, Andrea; Rieppel, Olivier; Ji, Cheng; Huang, Jian-Dong

    2015-01-22

    The incompleteness of the fossil record obscures the origin of many of the more derived clades of vertebrates. One such group is the Ichthyopterygia, a clade of obligatory marine reptiles that appeared in the Early Triassic epoch, without any known intermediates. Here we describe a basal ichthyosauriform from the upper Lower Triassic (about 248 million years ago) of China, whose primitive skeleton indicates possible amphibious habits. It is smaller than ichthyopterygians and had unusually large flippers that probably allowed limited terrestrial locomotion. It also retained characteristics of terrestrial diapsid reptiles, including a short snout and body trunk. Unlike more-derived ichthyosauriforms, it was probably a suction feeder. The new species supports the sister-group relationships between ichthyosauriforms and Hupehsuchia, the two forming the Ichthyosauromorpha. Basal ichthyosauromorphs are known exclusively from south China, suggesting that the clade originated in the region, which formed a warm and humid tropical archipelago in the Early Triassic. The oldest unequivocal record of a sauropterygian is also from the same stratigraphic unit of the region.

  12. Time-scale calibration by U-Pb geochronology: Examples from the Triassic Period

    Science.gov (United States)

    Mundil, R.

    2009-05-01

    U-Pb zircon geochronology, pioneered by Tom Krogh, is a cornerstone for the calibration of the time scale. Before Krogh's innovations, U-Pb geochronology was essentially limited by laboratory blank Pb (typically hundreds of nanograms) inherent in the then existing zircon dissolution and purification methods. The introduction of high pressure HF dissolution combined with miniature ion exchange columns (1) reduced the blank by orders of magnitude and allowed mass-spectrometric analyses of minute amounts of material (picograms of Pb and U). Krogh also recognized the need for minimizing the effects of Pb loss, and the introduction of the air-abrasion technique was the method of choice for two decades (2), until the development of the combined annealing and chemical abrasion technique resulted in essentially closed system zircons (3). These are the prerequisite for obtaining precise (permil-level) and accurate radio-isotopic ages of individual zircons contained in primary volcanic ash deposits, which are primary targets for the calibration of the time scale if they occur within fossil bearing sediments. A prime example is the calibration of the Triassic time scale which improved significantly using these techniques. The ages for the base and the top of the Triassic are constrained by U-Pb ages to 252.3 (4) and 201.5 Ma (5), respectively. These dates also constrain the ages of major extinction events at the Permian-Triassic and Triassic-Jurassic boundaries, and are statistically indistinguishable from ages obtained for the Siberian Traps and volcanic products from the Central Atlantic Magmatic Province, respectively, suggesting a causal link. Ages for these continental volcanics, however, are mostly from the K-Ar (40Ar/39Ar) system which requires accounting and correcting for a systematic bias of ca 1 % between U-Pb and 40Ar/39Ar isotopic ages (the 40Ar/39Ar ages being younger) (6). Robust U-Pb age constraints also exist for the Induan- Olenekian boundary (251.2 Ma, (7

  13. Record of Permian-Early Triassic continental arc magmatism in the western margin of the Jiamusi Block, NE China: petrogenesis and implications for Paleo-Pacific subduction

    Science.gov (United States)

    Yang, Hao; Ge, Wenchun; Dong, Yu; Bi, Junhui; Wang, Zhihui; Ji, Zheng; Yang, H.; Ge, W. C.; Dong, Y.; Bi, J. H.; Wang, Z. H.; Ji, Z.

    2017-09-01

    In this paper, we report zircon U-Pb ages, Hf isotopes and whole-rock geochemical data for the Permian to Early Triassic granitoids from the western margin of the Jiamusi Block (WJB), NE China. The intermediate to felsic (SiO2 = 59.67-74.04 wt%) granitoids belong to calc-alkaline series and are characterized by enrichments in light rare earth elements and large ion lithophile elements with pronounced negative Nb, Ta and Ti anomalies, revealing typical continental magmatic arc geochemical signatures. The zircon U-Pb determinations on the granodiorite, monzogranite, syenogranite and quartz diorite samples yielded ages between ca. 275-245 Ma, which, together with the published coeval intrusive rocks, indicates that Permian to Early Triassic continental arc magmatism occurred extensively in the WJB. The low and mainly negative zircon ɛ Hf( t) values between -7.6 and +1.6 and the zircon Hf model ages of 1.2-1.8 Ga, which are significantly older than their crystallization ages, suggest that they were mainly derived from reworking of ancient crustal materials with a limited input of juvenile components. The geochemical systematics and petrogenetic considerations indicate that the studied granitoids were generated from a zone of melting, assimilation, storage, and homogenization, i.e., a MASHed zone at the base of Paleo- to Mesoproterozoic continental crust, where large portions of igneous rocks and minor clay-poor sediments involved in the source region. In combination with regional geological data, we argue that the Jiamusi Block was unlikely the rifted segment of the Songliao Block and two possible geodynamical models were proposed to interpret the formation of the ca. 275-245 Ma granitoids in the WJB. In the context of Permian global plate reconstruction, we suggest that Paleo-Pacific plate subduction was initiated in the Permian to Early Triassic beneath the Jiamusi Block, and even whole eastern NE China.

  14. New hybodontiform and neoselachian sharks from the Lower Triassic of Oman

    DEFF Research Database (Denmark)

    Koot, Martha B.; Cuny, Gilles Guy Roger; Orchard, Michael J.

    2015-01-01

    Elasmobranchs are reported for the first time from Lower Triassic deposits in Oman. The well-preserved remains consist of isolated teeth, dermal denticles and fin spines, recovered from conodont residues. The low-palaeolatitude sections consist of Lopingian-Olenekian shallow and pelagic carbonates...... in exotics, olistoliths and breccia blocks that have been redeposited in younger allochthonous strata of the Hawasina Basin throughout the Oman Mountains at Jabal Safra (olistoliths within the Jurassic Guwayza Formation, Olenekian), as well as at Wadi Alwa (exotic Alwa Formation, Lopingian...... of the early stages of their evolution. All described taxa are new to the Oman fossil record and that of western Neotethys, apart from Omanoselache and Amelacanthus, which have been recognized from Wordian deposits, and Omanoselache is the second genus from Oman known to have survived the late Permian mass...

  15. A basal sauropodomorph (Dinosauria: Saurischia from the Ischigualasto Formation (Triassic, Carnian and the early evolution of Sauropodomorpha.

    Directory of Open Access Journals (Sweden)

    Ricardo N Martinez

    Full Text Available BACKGROUND: The earliest dinosaurs are from the early Late Triassic (Carnian of South America. By the Carnian the main clades Saurischia and Ornithischia were already established, and the presence of the most primitive known sauropodomorph Saturnalia suggests also that Saurischia had already diverged into Theropoda and Sauropodomorpha. Knowledge of Carnian sauropodomorphs has been restricted to this single species. METHODOLOGY/PRINCIPAL FINDINGS: We describe a new small sauropodomorph dinosaur from the Ischigualsto Formation (Carnian in northwest Argentina, Panphagia protos gen. et sp. nov., on the basis of a partial skeleton. The genus and species are characterized by an anteroposteriorly elongated fossa on the base of the anteroventral process of the nasal; wide lateral flange on the quadrate with a large foramen; deep groove on the lateral surface of the lower jaw surrounded by prominent dorsal and ventral ridges; bifurcated posteroventral process of the dentary; long retroarticular process transversally wider than the articular area for the quadrate; oval scars on the lateral surface of the posterior border of the centra of cervical vertebrae; distinct prominences on the neural arc of the anterior cervical vertebra; distal end of the scapular blade nearly three times wider than the neck; scapular blade with an expanded posterodistal corner; and medial lamina of brevis fossa twice as wide as the iliac spine. CONCLUSIONS/SIGNIFICANCE: We regard Panphagia as the most basal sauropodomorph, which shares the following apomorphies with Saturnalia and more derived sauropodomorphs: basally constricted crowns; lanceolate crowns; teeth of the anterior quarter of the dentary higher than the others; and short posterolateral flange of distal tibia. The presence of Panphagia at the base of the early Carnian Ischigualasto Formation suggests an earlier origin of Sauropodomorpha during the Middle Triassic.

  16. Short-snouted toothless ichthyosaur from China suggests Late Triassic diversification of suction feeding ichthyosaurs.

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    P Martin Sander

    Full Text Available BACKGROUND: Ichthyosaurs were an important group of Mesozoic marine reptiles and existed from the Early Triassic to the early Late Cretaceous. Despite a great diversity in body shapes and feeding adaptations, all share greatly enlarged eyes, an elongated rostrum with numerous conical teeth, and a streamlined body. METHODOLOGY/PRINCIPAL FINDINGS: Based on new material from China and the restudy of Shastasaurus pacificus, we here reinterpret the classical large-bodied Late Triassic ichthyosaur genus Shastasaurus to differ greatly from the standard ichthyosaurian body plan, indicating much greater morphological diversity and range of feeding adaptations in ichthyosaurs than previously recognized. Phylogenetic analysis indicates a monophyletic clade consisting of the giant Shonisaurus sikanniensis, Guanlingsaurus liangae, and Shastasaurus pacificus to which the genus name Shastasaurus is applied. Shastasaurus liangae comb. nov. is from the Late Triassic (Carnian Xiaowa Formation of Guizhou Province, southwestern China. The species combines a diminutive head with an entirely toothless and greatly reduced snout. The species also has by far the highest vertebral count among ichthyosaurs (86 presacral vertebrae and >110 caudal vertebrae, a count that is also very high for tetrapods in general. A reduced toothless snout and a diminutive head is also apparently present in the giant S. sikanniensis and presumably in S. pacificus. CONCLUSIONS/SIGNIFICANCE: In analogy to many modern odontocetes, Shastasaurus is interpreted as a specialized suction feeder on unshelled cephalopods and fish, suggesting a unique but widespread Late Triassic diversification of toothless, suction-feeding ichthyosaurs. Suction feeding has not been hypothesized for any of the other diverse marine reptiles of the Mesozoic before, but in Shastasaurus may be linked to the Late Triassic minimum in atmospheric oxygen.

  17. δ 13C evidence that high primary productivity delayed recovery from end-Permian mass extinction

    Science.gov (United States)

    Meyer, K. M.; Yu, M.; Jost, A. B.; Kelley, B. M.; Payne, J. L.

    2011-02-01

    Euxinia was widespread during and after the end-Permian mass extinction and is commonly cited as an explanation for delayed biotic recovery during Early Triassic time. This anoxic, sulfidic episode has been ascribed to both low- and high-productivity states in the marine water column, leaving the causes of euxinia and the mechanisms underlying delayed recovery poorly understood. Here we use isotopic analysis to examine the changing chemical structure of the water column through the recovery interval and thereby better constrain paleoproductivity. The δ 13C of limestones from 5 stratigraphic sections in south China displays a negative gradient of approximately 4‰ from shallow-to-deep water facies within the Lower Triassic. This intense gradient declines within Spathian and lowermost Middle Triassic strata, coincident with accelerated biotic recovery and carbon cycle stabilization. Model simulations show that high nutrient levels and a vigorous biological pump are required to sustain such a large gradient in δ 13C, indicating that Early Triassic ocean anoxia and delayed recovery of benthic animal ecosystems resulted from too much productivity rather than too little.

  18. Mass extinctions drove increased global faunal cosmopolitanism on the supercontinent Pangaea.

    Science.gov (United States)

    Button, David J; Lloyd, Graeme T; Ezcurra, Martín D; Butler, Richard J

    2017-10-10

    Mass extinctions have profoundly impacted the evolution of life through not only reducing taxonomic diversity but also reshaping ecosystems and biogeographic patterns. In particular, they are considered to have driven increased biogeographic cosmopolitanism, but quantitative tests of this hypothesis are rare and have not explicitly incorporated information on evolutionary relationships. Here we quantify faunal cosmopolitanism using a phylogenetic network approach for 891 terrestrial vertebrate species spanning the late Permian through Early Jurassic. This key interval witnessed the Permian-Triassic and Triassic-Jurassic mass extinctions, the onset of fragmentation of the supercontinent Pangaea, and the origins of dinosaurs and many modern vertebrate groups. Our results recover significant increases in global faunal cosmopolitanism following both mass extinctions, driven mainly by new, widespread taxa, leading to homogenous 'disaster faunas'. Cosmopolitanism subsequently declines in post-recovery communities. These shared patterns in both biotic crises suggest that mass extinctions have predictable influences on animal distribution and may shed light on biodiversity loss in extant ecosystems.Mass extinctions are thought to produce 'disaster faunas', communities dominated by a small number of widespread species. Here, Button et al. develop a phylogenetic network approach to test this hypothesis and find that mass extinctions did increase faunal cosmopolitanism across Pangaea during the late Palaeozoic and early Mesozoic.

  19. Osteology of Pseudochampsa ischigualastensis gen. et comb. nov. (Archosauriformes: Proterochampsidae) from the Early Late Triassic Ischigualasto Formation of Northwestern Argentina

    Science.gov (United States)

    Trotteyn, M. Jimena; Ezcurra, Martín D.

    2014-01-01

    Proterochampsids are crocodile-like, probably semi-aquatic, quadrupedal archosauriforms characterized by an elongated and dorsoventrally low skull. The group is endemic from the Middle-Late Triassic of South America. The most recently erected proterochampsid species is “Chanaresuchus ischigualastensis”, based on a single, fairly complete skeleton from the early Late Triassic Ischigualasto Formation of northwestern Argentina. We describe here in detail the non-braincase cranial and postcranial anatomy of this species and revisit its taxonomy and phylogenetic relationships. The phylogenetic analysis recovered ‘Chanaresuchus ischigualastensis’ as part of a trichotomy together with Gualosuchus reigi and Chanaresuchus bonapartei. Accordingly, “Chanaresuchus ischigualastensis” can be potentially more closely related to Gualosuchus reigi, or even Rhadinosuchus gracilis, than to Chanaresuchus bonapartei. In addition, after discussing previously claimed synapomorphies of Chanaresuchus, we could not find unambiguous support for the monophyly of the genus. As a result, we propose here the erection of the new genus Pseudochampsa for ‘Chanaresuchus ischigualastensis’, which results in the new combination Pseudochampsa ischigualastensis. The information provided here about the anatomy and taxonomy of Pseudochampsa ischiguaslastensis will be useful for future quantitative analyses focused on the biogeography and macroevolutionary history of proterochampsids. PMID:25426846

  20. Osteology of Pseudochampsa ischigualastensis gen. et comb. nov. (Archosauriformes: Proterochampsidae from the early late triassic ischigualasto formation of Northwestern Argentina.

    Directory of Open Access Journals (Sweden)

    M Jimena Trotteyn

    Full Text Available Proterochampsids are crocodile-like, probably semi-aquatic, quadrupedal archosauriforms characterized by an elongated and dorsoventrally low skull. The group is endemic from the Middle-Late Triassic of South America. The most recently erected proterochampsid species is "Chanaresuchus ischigualastensis", based on a single, fairly complete skeleton from the early Late Triassic Ischigualasto Formation of northwestern Argentina. We describe here in detail the non-braincase cranial and postcranial anatomy of this species and revisit its taxonomy and phylogenetic relationships. The phylogenetic analysis recovered 'Chanaresuchus ischigualastensis' as part of a trichotomy together with Gualosuchus reigi and Chanaresuchus bonapartei. Accordingly, "Chanaresuchus ischigualastensis" can be potentially more closely related to Gualosuchus reigi, or even Rhadinosuchus gracilis, than to Chanaresuchus bonapartei. In addition, after discussing previously claimed synapomorphies of Chanaresuchus, we could not find unambiguous support for the monophyly of the genus. As a result, we propose here the erection of the new genus Pseudochampsa for 'Chanaresuchus ischigualastensis', which results in the new combination Pseudochampsa ischigualastensis. The information provided here about the anatomy and taxonomy of Pseudochampsa ischiguaslastensis will be useful for future quantitative analyses focused on the biogeography and macroevolutionary history of proterochampsids.

  1. A palaeoequatorial ornithischian and new constraints on early dinosaur diversification.

    Science.gov (United States)

    Barrett, Paul M; Butler, Richard J; Mundil, Roland; Scheyer, Torsten M; Irmis, Randall B; Sánchez-Villagra, Marcelo R

    2014-09-22

    Current characterizations of early dinosaur evolution are incomplete: existing palaeobiological and phylogenetic scenarios are based on a fossil record dominated by saurischians and the implications of the early ornithischian record are often overlooked. Moreover, the timings of deep phylogenetic divergences within Dinosauria are poorly constrained owing to the absence of a rigorous chronostratigraphical framework for key Late Triassic-Early Jurassic localities. A new dinosaur from the earliest Jurassic of the Venezuelan Andes is the first basal ornithischian recovered from terrestrial deposits directly associated with a precise radioisotopic date and the first-named dinosaur from northern South America. It expands the early palaeogeographical range of Ornithischia to palaeoequatorial regions, an area sometimes thought to be devoid of early dinosaur taxa, and offers insights into early dinosaur growth rates, the evolution of sociality and the rapid tempo of the global dinosaur radiation following the end-Triassic mass extinction, helping to underscore the importance of the ornithischian record in broad-scale discussions of early dinosaur history. © 2014 The Author(s) Published by the Royal Society. All rights reserved.

  2. High-precision U-Pb zircon geochronological constraints on the End-Triassic Mass Extinction, the late Triassic Astronomical Time Scale and geochemical evolution of CAMP magmatism

    Science.gov (United States)

    Blackburn, T. J.; Olsen, P. E.; Bowring, S. A.; McLean, N. M.; Kent, D. V.; Puffer, J. H.; McHone, G.; Rasbury, T.

    2012-12-01

    Mass extinction events that punctuate Earth's history have had a large influence on the evolution, diversity and composition of our planet's biosphere. The approximate temporal coincidence between the five major extinction events over the last 542 million years and the eruption of Large Igneous Provinces (LIPs) has led to the speculation that climate and environmental perturbations generated by the emplacement of a large volume of magma in a short period of time triggered each global biologic crisis. Establishing a causal link between extinction and the onset and tempo of LIP eruption has proved difficult because of the geographic separation between LIP volcanic deposits and stratigraphic sequences preserving evidence of the extinction. In most cases, the uncertainties on available radioisotopic dates used to correlate between geographically separated study areas often exceed the duration of both the extinction interval and LIP volcanism by an order of magnitude. The "end-Triassic extinction" (ETE) is one of the "big five" and is characterized by the disappearance of several terrestrial and marine species and dominance of Dinosaurs for the next 134 million years. Speculation on the cause has centered on massive climate perturbations thought to accompany the eruption of flood basalts related to the Central Atlantic Magmatic Province (CAMP), the most aerially extensive and volumetrically one of the largest LIPs on Earth. Despite an approximate temporal coincidence between extinction and volcanism, there lacks evidence placing the eruption of CAMP prior to or at the initiation of the extinction. Estimates of the timing and/or duration of CAMP volcanism provided by astrochronology and Ar-Ar geochronology differ by an order of magnitude, precluding high-precision tests of the relationship between LIP volcanism and the mass extinction, the causes of which are dependent upon the rate of magma eruption. Here we present high precision zircon U-Pb ID-TIMS geochronologic data

  3. CONODONT BIOSTRATIGRAPHY AND LITHOSTRATIGRAPHY ACROSS THE PERMIAN-TRIASSIC BOUNDARY AT THE LUKAC SECTION IN WESTERN SLOVENIA

    Directory of Open Access Journals (Sweden)

    TEA KOLAR-JURKOVSEK

    2011-03-01

    Full Text Available Detailed conodont biostratigraphy and lithostratigraphy of the Late Permian and Early Triassic beds were studied at the LukaC section in western Slovenia. The analyzed section is composed of the Bellerophon Formation ("evaporite-dolomite member" and the newly introduced Lukaè Formation ("transitional beds", "streaky limestone member" and "carbonate-clastic beds member". The Permian-Triassic boundary interval is represented by "transitional beds" of carbonate facies deposited in shallow restricted marine conditions. The presence of H. parvus in sample L1 in the "transitional beds" marks the systemic boundary between Permian and Triassic. The studied interval is characterized by a diverse microfauna that contain conodonts, foraminifers, ostracods and gastropods. Six conodont zones have been recognized, in ascending order, the latest Changhsingian (uppermost Permian praeparvus Zone, and the Griesbachian (lowermost Triassic parvus, lobata, staeschei-isarcica, postparvus and anceps zones. This faunal succession represents the first known and the most complete conodont biozonation across the Permian-Triassic interval from the entire Dinaric region. The recognized conodont biozones can be correlated with the biozonation of the Southern Alps and of the GSSP Meishan D section. 

  4. Ichthyosaurs from the French Rhaetian indicate a severe turnover across the Triassic-Jurassic boundary

    Science.gov (United States)

    Fischer, Valentin; Cappetta, Henri; Vincent, Peggy; Garcia, Géraldine; Goolaerts, Stijn; Martin, Jeremy E.; Roggero, Daniel; Valentin, Xavier

    2014-12-01

    Mesozoic marine reptiles went through a severe turnover near the end of the Triassic. Notably, an important extinction event affected ichthyosaurs, sweeping a large part of the group. This crisis is, however, obscured by an extremely poor fossil record and is regarded as protracted over the entire Norian-earliest Jurassic interval, for the lack of a more precise scenario. The iconic whale-sized shastasaurid ichthyosaurs are regarded as early victims of this turnover, disappearing by the middle Norian. Here we evaluate the pattern of this turnover among ichthyosaurs by analysing the faunal record of two Rhaetian localities. One locality is Autun, eastern France; we rediscovered in this material the holotypes or partial `type' series of Rachitrema pellati, Actiosaurus gaudryi, Ichthyosaurus rheticus, Ichthyosaurus carinatus and Plesiosaurus bibractensis; a revised taxonomic scheme is proposed. The second assemblage comes from a new locality: Cuers, southeastern France. Both these assemblages provide several lines of evidence for the presence of shastasaurid-like ichthyosaurs in the Rhaetian of Europe. These occurrences suggest that both the demise of shastasaurids and the sudden radiation of neoichthyosaurians occurred within a short time window; this turnover appears not only more abrupt but also more complex than previously postulated and adds a new facet of the end-Triassic mass extinction.

  5. A tale of two extinctions : converging end-Permian and end-Triassic scenarios

    NARCIS (Netherlands)

    van de Schootbrugge, Bas; Wignall, Paul B.

    The end-Permian (c. 252 Ma) and end-Triassic (c. 201 Ma) mass-extinction events are commonly linked to the emplacement of the large igneous provinces of the Siberia Traps and Central Atlantic Magmatic Province, respectively. Accordingly, scenarios for both extinctions are increasingly convergent and

  6. Sodium storage in deep paleoweathering profiles beneath the Paleozoic-Triassic unconformity

    Science.gov (United States)

    Thiry, M.; Parcerisa, D.; Ricordel-Prognon, C.; Schmitt, J.-M.

    2009-04-01

    A major sodium accumulation has been recognized for long and by numerous authors in the Permo-Triassic salt deposits (Hay et al., 2006). Beside these basinal deposits, important masses of sodium were stored on the continents within deep palaeoweathering profiles in form of albite. Indeed, wide surfaces and huge volumes of granito-gneissic basements of the Hercynian massifs are albitized from North-Africa up to Scandinavia. These albitized rocks have usually been considered as related to tardi-magmatic metasomatic processes (Cathelineau 1986; Petersson and Eliasson 1997). Geometrical arrangement and dating of these alterations point out that these albitizations, or at least a part of them, developed under low temperature subsurface conditions in relation with the Triassic palaeosurface (Ricordel et al., 2007; Parcerisa et al., 2009). Petrology The albitized igneous rocks show a strong alteration with pseudomorphic replacement of the primary plagioclases into albite, replacement of primary biotite by chlorite and minor precipitation of neogenic minerals like albite, chlorite, apatite, haematite, calcite and titanite. Albitized rocks are characterized by their pink coloration due to the presence of minute haematite inclusions in the albite. The development and distribution of the albitization and related alterations above the unaltered basement occurs in three steps that define a vertical profile, up to 100-150 m depth. 1) In the lower part of the profile, albitization occurs within pink-colored patches in the unaltered rock, giving a pink-spotted aspect to the rock. 2) In the middle part of the profile, rocks have an overall pink coloration due to the albitization of the primary Ca-bearing igneous plagioclases. Usually, this facies develops in a pervasive manner, affecting the whole rock, but it may also be restricted to joints, giving a sharp-pink coloration to the fracture wall. 3) Finally, the top of the profile is defined by the same mineral paragenesis as in the

  7. The Park Volcanics Group : field relations of an igneous suite emplaced in the Triassic-Jurassic Murihiku Terrane, South Island, New Zealand

    International Nuclear Information System (INIS)

    Coombs, D.S.; Cook, N.D.J.; Campbell, J.D.

    1992-01-01

    Park Volcanics Group is proposed for igneous rocks, either shallow intrusive or extrusive, emplaced in the Murihiku Terrane during Triassic-Jurassic times. The term replaces Park Intrusives of Mutch, some members of which are shown to be extrusive rather than intrusive. Formation status within the group is given to Gowan Andesite and Pinney Volcanics (new names) in western Southland, Glenham Porphyry in eastern Southland, and Barnicoat Andesite (new) in the Richmond area, Nelson. Gowan Andesite is a porphyritic feldspar two-pyroxene andesite with a glassy or microcrystalline groundmass. A suite of low-grade metavolcanic rocks which forms the main mass of Malakoff Hill and which has formerly been included in the 'Park Intrusives' is here excluded and ascribed to the Takitimu Group; representative chemical data are given. Glenham Porphyry is typically a porphyritic feldspar two-pyroxene andesite texturally similar to the Gowan Andesite but with significant geochemical differences. Two volumetrically minor members are recognised, Habukinini Trachydacite and Kenilworth Rhyolite. In the north of its outcrop area, Glenham Porphyry is emplaced on or into Late Triassic terrestrial beds; in the middle it overlies Kaihikuan (Middle Triassic) and is overlain by Otapirian (latest Triassic) marine beds; and in the southeast it is directly overlain by Ururoan (late Early to early Middle Jurassic) conglomerates and marine sandstones. Pinney Volcanics are restricted to a very few, probably one, massive conglomeratic horizon in the Oretian Stage. The commonest rock type is a two-pyroxene trachydacite, modified by very-low-grade burial metamorphism. Auto-brecciation is characteristic and rock types change over short distances. Hornblende-rich variants occur as well as more felsic varieties including rhyolite ignimbrite. These may have been erupted onto a bouldery floodplain or shallow-marine surface, but alternatively may have been mass-emplaced by debris avalanche resulting from

  8. Carbon isotope evidence for a vigorous biological pump in the wake of end-Permian mass extinction

    Science.gov (United States)

    Meyer, K. M.; Yu, M.; Jost, A. B.; Payne, J.

    2009-12-01

    Ocean anoxia and euxinia have long been linked to the end-Permian mass extinction and the subsequent Early Triassic interval of delayed biotic recovery. This anoxic, sulfidic episode has been ascribed to both low- and high-productivity states in the marine water column, leaving the causes of euxinia and the mechanisms underlying delayed recovery poorly understood. To examine the nature of the end-Permian and Early Triassic biological production, we measured the carbon isotopic composition of carbonates from an exceptionally preserved carbonate platform in the Nanpanjiang Basin of south China. 13C of limestones from 5 stratigraphic sections displays a gradient of approximately 4‰ from shallow to deep water within the Lower Triassic. The limestones are systematically enriched in the platform interior relative to coeval slope and basin margin deposits by 2-4‰ at the peaks of correlative positive and negative δ13C excursions. This gradient subsequently collapses to less than 1‰ in the Middle Triassic, coincident with accelerated biotic recovery and cessation of δ13C excursions. Based on the relationship between δ18O and δ13C, trace metal analyses, and lithostratigraphic context, we conclude that the carbon isotope gradient is unlikely to reflect meteoric diagenesis, organic matter remineralization, or changes in the mixing ratio of sediment sources and minerals across the platform. Instead, we interpret the relatively depleted δ13C values toward the basin as reflecting DIC input from 13C-depleted deep waters during early diagenesis in a nutrient-rich, euxinic ocean. These observations suggest that a vigorous prokaryote-driven biological pump sustained Early Triassic ocean anoxia and inhibited recovery of animal ecosystems.

  9. Triassic-Jurassic pteridosperms of Australasia: speciation, diversity and decline

    Energy Technology Data Exchange (ETDEWEB)

    Pattemore, G. A.; Rigby, J. F.; Playford, G.

    2015-07-01

    Pteridosperms are preserved abundantly in the Gondwanan Triassic, with many species exhibiting consider- able morphological variation that has been attributed to a hybridization model of speciation. This is an improbable explanation given that hybridization is very rare in gymnosperms. Allopatric speciation resulting from geographic and climatic provincialism is a more likely explanation for the morphological diversity which is well represented in Anisian Norian (Middle and Upper Triassic) floras of Australasia and elsewhere in Gondwana. Most specimens are distributed among three families: Umkomasiaceae, Peltaspermaceae and Matatiellaceae. These families, together with other possibly pteridospermous genera, are reviewed herein. Diversity in these families apparently declined by the Rhaetian and they did not persist into the Gondwanan post-Triassic. Australasian post-Triassic strata contain remarkably different floral assemblages to those of the Triassic. No fructifications are clearly pteridospermous and no remains show any obvious relationship with pteridosperms of the Gondwanan Triassic. Caytonialean fructifications are not known in Australasian strata; however, associated foliage has been reported from the Eastern Gondwanan Upper Triassic through Middle Jurassic including Australia. Much fern-like foliage, claimed to be pteridospermous from the Lower Jurassic through Eocene of Eastern Gondwana, lacks supporting evidence of such affiliation. (Author)

  10. Biostratigraphic reappraisal of the Lower Triassic Sanga do Cabral Supersequence from South America, with a description of new material attributable to the parareptile genus Procolophon

    Science.gov (United States)

    Dias-da-Silva, Sérgio; Pinheiro, Felipe L.; Stock Da-Rosa, Átila Augusto; Martinelli, Agustín G.; Schultz, Cesar L.; Silva-Neves, Eduardo; Modesto, Sean P.

    2017-11-01

    The Sanga do Cabral Supersequence (SCS), comprises the Brazilian Sanga do Cabral Formation (SCF) and the Uruguayan Buena Vista Formation (BVF). So far, the SCS has yielded temnospondyls, parareptiles, archosauromorphs, putative synapsids, and a number of indeterminate specimens. In the absence of absolute dates for these rocks, a biostratigraphic approach is necessary to establish the ages of the SCF and the BVF. It is well established that the SCF is Early Triassic mainly due to the presence of the widespread Gondwanan reptile Procolophon trigoniceps. Conversely, the age of the BVF is subject of great controversy, being regarded alternatively as Permian, Permo-Triassic, and Early Triassic. The BVF has yielded the definite procolophonid Pintosaurus magnidentis. Procolophonoidea is one of the most diverse and conspicuous terrestrial tetrapod groups of the Lower Triassic Lystrosaurus Assemblage Zone in the Karoo Basin of South Africa, which preserves tetrapods from the aftermath of the end-Permian extinction event. Based on a previous interpretation that the fauna of the BVF is Permian, and in the reinterpretation of disarticulated vertebrae from SCF with 'swollen' neural arches as belonging to either seymouriamorphs or diadectomorphs, it was recently suggested that at least part of the SCF is Permian in age, which prompted this comprehensive reevaluation of both SCS's faunal content and geology. Moreoever, new, strikingly large procolophonid specimens (skull, vertebra, and a mandibular fragment) from the SCF are described and referred to the genus Procolophon. The large procolophonid vertebra described here contradicts the recent hypothesis that similar specimens from the SCF belong to seymouriamorphs or diadectomorphs, because its morphology is consistent with that found in Procolophon. There is not a single diagnostic specimen that supports the inference of Permian levels in the SCS. Accordingly, because all diagnostic and biostratigraphically informative fossils

  11. Permian and Triassic microfloral assemblages from the Blue Nile Basin, central Ethiopia

    Science.gov (United States)

    Dawit, Enkurie L.

    2014-11-01

    Palynological investigation was carried out on surface samples from up to 400 m thick continental siliciclastic sediments, here referred to as “Fincha Sandstone”, in the Blue Nile Basin, central Ethiopia. One hundred sixty species were identified from 15 productive samples collected along a continuous road-cut exposure. Six informal palynological assemblage zones have been identified. These assemblage zones, in ascending order, are: “Central Ethiopian Permian Assemblage Zone - CEPAZ I”, earliest Permian (Asselian-Sakmarian); “CEPAZ II”, late Early Permian (Artinskian-Kungurian); CEPAZ III - Late Permian (Kazanian-Tatarian); “CETAZ IV”, Lower Triassic (Olenekian Induan); “CETAZ V”, Middle Triassic (Anisian Ladinian); “CETAZ VI”, Late Triassic (Carnian Norian). Tentative age ranges proposed herein are compared with faunally calibrated palynological zones in Gondwana. The overall composition and vertical distribution of miospores throughout the studied section reveals a wide variation both qualitatively and quantitatively. The high frequency of monosaccate pollen in CEPAZ I may reflect a Glossopterid-dominated upland flora in the earliest Permian. The succeeding zone is dominated by straite/taeniate disaccate pollen and polyplicates, suggesting a notable increase in diversity of glossopterids. The decline in the diversity of taeniate disaccate pollen and the concomitant rise in abundance of non-taeniate disaccates in CEPAZ III may suggest the decline in Glossopteris diversity, though no additional evidence is available to equate this change with End-Permian extinction. More diverse and dominant non-taeniate, disaccate, seed fern pollen assignable to FalcisporitesAlisporites in CETAZ IV may represent an earliest Triassic recovery flora. The introduction of new disaccate forms with thick, rigid sacci, such as Staurosaccites and Cuneatisporites, in CETAZ V and VI may indicate the emergence of new gymnospermous plants that might have favourably

  12. Sedimentary record of subsidence pulse at the Triassic/Jurassic boundary interval in the Slovenian Basin (eastern Southern Alps

    Directory of Open Access Journals (Sweden)

    Rožič Boštjan

    2017-08-01

    Full Text Available In the Alpine Realm the Early Jurassic is characterized by the disintegration and partial drowning of vast platform areas. In the eastern part of the Southern Alps (present-day NW Slovenia, the Julian Carbonate Platform and the adjacent, E-W extending Slovenian Basin underwent partial disintegration, drowning and deepening from the Pliensbachian on, whereas only nominal environmental changes developed on the large Dinaric (Friuli, Adriatic Carbonate Platform to the south (structurally part of the Dinarides. These events, however, were preceded by an earlier - and as yet undocumented extensional event - that took place near the Triassic/Jurassic boundary. This paper provides evidence of an accelerated subsidence from four selected areas within the Slovenian Basin, which show a trend of eastwardly-decreasing deformation. In the westernmost (Mrzli vrh section - the Upper Triassic platform-margin - massive dolomite is overlain by the earliest Jurassic toe-of-slope carbonate resediments and further, by basin-plain micritic limestone. Further east (Perbla and Liščak sections the Triassic-Jurassic transition interval is marked by an increase in resedimented carbonates. We relate this to the increasing inclination and segmentation of the slope and adjacent basin floor. The easternmost (Mt. Porezen area shows a rather monotonous, latest Triassic-Early Jurassic basinal sedimentation. However, changes in the thickness of the Hettangian-Pliensbachian Krikov Formation point to a tilting of tectonic blocks within the basin area. Lateral facies changes at the base of the formation indicate that the tilting occurred at and/or shortly after the Triassic/Jurassic boundary

  13. Sedimentary record of subsidence pulse at the Triassic/Jurassic boundary interval in the Slovenian Basin (eastern Southern Alps)

    Science.gov (United States)

    Rožič, Boštjan; Jurkovšek, Tea Kolar; Rožič, Petra Žvab; Gale, Luka

    2017-08-01

    In the Alpine Realm the Early Jurassic is characterized by the disintegration and partial drowning of vast platform areas. In the eastern part of the Southern Alps (present-day NW Slovenia), the Julian Carbonate Platform and the adjacent, E-W extending Slovenian Basin underwent partial disintegration, drowning and deepening from the Pliensbachian on, whereas only nominal environmental changes developed on the large Dinaric (Friuli, Adriatic) Carbonate Platform to the south (structurally part of the Dinarides). These events, however, were preceded by an earlier - and as yet undocumented extensional event - that took place near the Triassic/Jurassic boundary. This paper provides evidence of an accelerated subsidence from four selected areas within the Slovenian Basin, which show a trend of eastwardly-decreasing deformation. In the westernmost (Mrzli vrh) section - the Upper Triassic platform-margin - massive dolomite is overlain by the earliest Jurassic toe-of-slope carbonate resediments and further, by basin-plain micritic limestone. Further east (Perbla and Liščak sections) the Triassic-Jurassic transition interval is marked by an increase in resedimented carbonates. We relate this to the increasing inclination and segmentation of the slope and adjacent basin floor. The easternmost (Mt. Porezen) area shows a rather monotonous, latest Triassic-Early Jurassic basinal sedimentation. However, changes in the thickness of the Hettangian-Pliensbachian Krikov Formation point to a tilting of tectonic blocks within the basin area. Lateral facies changes at the base of the formation indicate that the tilting occurred at and/or shortly after the Triassic/Jurassic boundary

  14. Restoration of marine ecosystems following the end-Permian mass extinction: pattern and dynamics

    Science.gov (United States)

    Chen, Z.

    2013-12-01

    Life came closest to complete annihilation during the end-Permian mass extinction (EPME). Pattern and cause of this great dying have long been disputed. Similarly, there is also some debate on the recovery rate and pattern of marine organisms in the aftermath of the EPME. Some clades recovered rapidly, within the first 1-3 Myr of the Triassic. For instance, foraminiferal recovery began 1 Myr into the Triassic and was not much affected by Early Triassic crises. Further, some earliest Triassic body and trace fossil assemblages are also more diverse than predicted. Others, ie. Brachiopods, corals etc., however, did not rebound until the Middle Triassic. In addition, although ammonoids recovered fast, reaching a higher diversity by the Smithian than in the Late Permian, much of this Early Triassic radiation was within a single group, the Ceratitina, and their morphological disparity did not expand until the end-Spathian. Here, I like to broaden the modern ecologic network model to explore the complete trophic structure of fossilized ecosystems during the Permian-Triassic transition as a means of assessing the recovery. During the Late Permian and Early Triassic, primary producers, forming the lowest trophic level, were microbes. The middle part of the food web comprises primary and meso-consumer trophic levels, the former dominated by microorganisms such as foraminifers, the latter by opportunistic communities (i.e. disaster taxa), benthic shelly communities, and reef-builders. They were often consumed by invertebrate and vertebrate predators, the top trophic level. Fossil record from South China shows that the post-extinction ecosystems were degraded to a low level and typified by primary producers or opportunistic consumers, which are represented by widespread microbialites or high-abundance, low-diversity communities. Except for some opportunists, primary consumers, namely foraminifers, rebounded in Smithian. Trace-makers recovered in Spathian, which also saw

  15. Determining the Central Atlantic Magmatic Province (CAMPS)'s Role in the Increased Flux of CO2 in the end-Triassic Mass Extinction

    Science.gov (United States)

    Srinivasan, P. S.; Bachan, A.; Stanford School of Earth Sciences Department of Paleobiology

    2011-12-01

    The Central Atlantic Magmatic Province (CAMP) is one of the largest flood basalt provinces known. Its empacement coincided with a period of major plant and animal extinctions-the end-Triassic mass extinction. It is postulated that the release of large amounts of carbon dioxide into the atmosphere from the volcanics was one of the causes of this mass extinction. However,the magnitude of impact on ocean chemistry, and timescales involved remain unclear. To determine CAMP's role in this increased flux of CO2, we studied the geochemistry of samples of rock from the Triassic-Jurassic boundary, in northern Italy. Specifically, by observing the ratios of carbon isotopes 12 and 13 in the organic carbon found in these limestone sedimentary rocks, we could determine the ratio of carbonate to organic burial fluxes globally. We drilled limestone rocks from two different sections in the Southern Alps-- Pozzo Glaciale and Val Adrara. Once they were drilled to a fine powder-like form, we acidified the CaCO3 with HCl to isolate the organic carbon. Then, the organic matter was cleaned to rid the acid, and eventually was placed into tin foil to be placed into the Elemental Analyzer, which determined the percent Carbon in each sample. We tested about 200 samples, and placed them into the Mass Spectrometer machine to determine the isotopic ratios of C12 and C13. According to the data, there was a positive excursion for both sample sets, which means that there was an increase in the amount of C13 in the organic matter. The duration of this excursion was at least a few hundred thousand years. This suggests a protracted increase in the burial flux of organic carbon globally, which is consistent with the hypothesized volcanically driven increase in CO2. This further bolsters the contention that CAMP was responsible, in part, for this mass extinction. By studying the earth's recovery from increased carbon fluxes in the past, we can predict the recovery path that our anthropogenically

  16. Permian Triassic palynofloral transition in Chintalapudi area, Godavari Graben, Andhra Pradesh, India

    Science.gov (United States)

    Jha, Neerja; Chary, M. Basava; Aggarwal, Neha

    2012-10-01

    The entire 606 m-thick sedimentary sequence in borecore MCP-7 from Chintalapudi area, Chintalapudi sub-basin has been lithologically designated as Kamthi Formation. However, the palynological investigation revealed five distinct palynoassemblages, which essentially fall under two groups, one group (Palynoassemblage-I, II and III) having dominance of striate disaccates along with presence of some stratigraphically significant taxa, belongs to Late Permian (Raniganj) palynoflora, while the other group (Palynoassemblages IV and V) shows sharp decline in percentage of characteristic taxa of first group, i.e., striate disaccates, and consequent rise or dominance of taeniate and cingulate cavate spores, belongs to Early Triassic (Panchet) palynoflora. Palynoassemblage-I, II and III (Group I) are characterized by dominance of striate disaccates chiefly, Striatopodocarpites spp. and Faunipollenites spp. along with presence of rare but stratigraphically significant taxa, viz., Gondisporites raniganjensis, Falcisporites nuthaliensis, Klausipollenites schaubergeri, Chordasporites sp., Striomonosaccites, ovatus, Crescentipollenites multistriatus, Verticipollenites debiles, Strotersporites crassiletus, Guttulapollenites hannonicus, G. gondwanensis, Hamiapollenites insolitus, Corisaccites alutus, Lunatisporites ovatus, Weylandites spp. and Vitreisporites pallidus. Palynoassemblage-I is distinguished by significant presence of Densipollenites spp. while Palynoassemblage-II shows significant presence of Crescentipollenites spp. and Palynoassemblage-III differs from the above two assemblages in having significant presence of Guttulapollenites hannonicus. Palynoassemblage-IV (Group II) is characterized by high percentage of taeniate disaccates chiefly Lunatisporites spp., while Palynoassemblage-V (Group II) is characterized by cingulate-cavate trilete spores chiefly, Lundbladispora spp. and Densoisporites spp. Striate disaccates show a sharp decline in these two assemblages. In

  17. A NEW PERLEIDID FROM THE SPATHIAN (OLENEKIAN, EARLY TRIASSIC OF CHAOHU, ANHUI PROVINCE, CHINA

    Directory of Open Access Journals (Sweden)

    ZUOYU SUN

    2013-11-01

    Full Text Available A new actinopterygian genus and species, Chaohuperleidus  primus gen. n. n. sp., is described from the Upper Member of the Nanlinghu Formation (Spathian, Olenekian, Early Triassic at Majiashan (Majia Hill, Chaohu City, Anhui Province, Southeast China. The new taxon is assigned to the order Perleidiformes on the base of a combination of features: large wedge-like preoperculum with expanded infraorbital process and sensory canal running almost vertical along the posterior margin of the preoperculum itself, flank scalessomewhat higher than longer in the trunk region and presence of epaxial rays in the caudal fin.AmongPerleidiformes, the new taxonbelongsto thePerleididae, being very similar to Perleidus in the general body shape, pattern of the skull bones, the dentitionmade of peg-like marginal teeth,and number of epaxial rays. The new genus is characterized by the presence of a very large antero-dorsal process on the suboperculum, a high number (about 55 of transversal scale rows and dorsal and anal fin quite posterior. Chaohuperleidus gen. n., beingSpathianin age,represents the oldest record of the order Perleidiformes. 

  18. Trouble Upstairs: Reconstructing Permian-Triassic Climate during Siberian Traps Magmatism

    Science.gov (United States)

    Black, B. A.; Neely, R. R., III; Lamarque, J. F.; Elkins-Tanton, L. T.; Mills, M. J.

    2014-12-01

    The eruption of large igneous provinces can transfer significant masses of volatiles from Earth's interior to the atmosphere. What are the consequences of this degassing for habitability and extinction? In this presentation, we consider this question in the context of Siberian Traps magmatism, which has been shown to overlap within geochronologic uncertainty with catastrophic deterioration of Permian-Triassic marine and terrestrial ecosystems. To investigate the impacts of endogenic gases on climate, atmospheric chemistry, and ocean circulation, we conducted a series of numerical experiments with a comprehensive global model for the Permian-Triassic. Our simulations predict the intensity and distribution of acid rain and ozone depletion, with implications for terrestrial biota. We further explore feedbacks between sulfur emissions, transient cooling, and shifts in ocean circulation. We suggest that Siberian Traps magmatism may have triggered several distinct kill mechanisms in the oceans and on land, contributing to a complex combined pattern of environmental stress and latest Permian ecological failure.

  19. Depositional setting and vertebrate biostratigraphy of the Triassic ...

    Indian Academy of Sciences (India)

    The Dockum tetrapod diversity is reviewed in a hierarchical phylogeny with remarks on the history of ...... Triassic ecosystem. A review of ... form of fish from the Triassic beds of Texas, with remarks ..... ships and functional implications; J. Vert.

  20. MIDDLE TRIASSIC AUTOCLASTIC DEPOSITS FROM SOUTHWESTERN BOSNIA AND HERZEGOVINA

    Directory of Open Access Journals (Sweden)

    Duje Smirčić

    2016-06-01

    Full Text Available Middle Triassic volcanic and volcaniclastic rocks have been recognized at several localities near Bosansko Grahovo, in southwestern part of Bosnia and Herzegovina in the External Dinarides. Peculiar types of autoclastic rocks were investigated. These are peperites and hyaloclastites. Regarding specific structures, mineral composition and micropetrographic characteristics it was possible to further differentiate hyaloclastites into in situ hyaloclastites and slightly resedimented hyaloclastites that represent genetic succession. All rock types occurred in a deep sea troughs that formed as a consequence of Middle Triassic extensional tectonic and rift related wrench faulting. In situ hyaloclastites and slightly resedimented hyaloclastites were formed due to quenching at the contact of lava effusions with sea water. Genesis of peperites is related to lava emplacement in unconsolidated water saturated lime mudstones that were deposited in deep sea basin. All investigated rock types represent first findings of autoclastic deposits in the External Dinarides. Biostratigraphic constraints achieved by means of conodont species Neogondolella excentrica, Paragondolella excelsa, Paragondolella trammeri and Gladigondolella tethydis indicate Late Anisian to Early Ladinian interval of the autoclastic deposits from Bosansko Grahovo.

  1. Mayer Kangri metamorphic complexes in Central Qiangtang (Tibet, western China): implications for the Triassic-early Jurassic tectonics associated with the Paleo-Tethys Ocean

    Science.gov (United States)

    Wang, Yixuan; Liang, Xiao; Wang, Genhou; Yuan, Guoli; Bons, Paul D.

    2018-03-01

    The Mesozoic orogeny in Central Qiangtang Metamorphic Belt, northern Tibet, provides important insights into the geological evolution of the Paleo-Tethys Ocean. However, the Triassic-early Jurassic tectonics, particularly those associated with the continental collisionstage, remains poorly constrained. Here we present results from geological mapping, structural analysis, P-T data, and Ar-Ar geochronology of the Mayer Kangri metamorphic complex. Our data reveal an E-W-trending, 2 km wide dome-like structure associated with four successive tectonic events during the Middle Triassic and Early Jurassic. Field observations indicate that amphibolite and phengite schist complexes in this complex are separated from the overlying lower greenschist mélange by normal faulting with an evident dextral shearing component. Open antiform-like S2 foliation of the footwall phengite schist truncates the approximately north-dipping structures of the overlying mélange. Microtextures and mineral chemistry of amphibole reveal three stages of growth: Geothermobarometric estimates yield temperatures and pressures of 524 °C and 0.88 GPa for pargasite cores, 386 °C and 0.34 GPa for actinolite mantles, and 404 °C and 0.76 GPa for winchite rims. Peak blueschist metamorphism in the phengite schist occurred at 0.7-1.1 GPa and 400 °C. Our Ar-Ar dating of amphibole reveals rim-ward decreasing in age bands, including 242.4-241.2 Ma, ≥202.6-196.8, and 192.9-189.8 Ma. The results provide evidence for four distinct phases of Mesozoic tectonic evolution in Central Qiangtang: (1) northward oceanic subduction beneath North Qiangtang ( 244-220 Ma); (2) syn-collisional slab-break off (223-202 Ma); (3) early collisional extension driven by buoyant extrusion flow from depth ( 202.6-197 Ma); and (4) post-collision contraction and reburial (195.6-188.7 Ma).

  2. Adaptations for marine habitat and the effect of Triassic and Jurassic predator pressure on development of decompression syndrome in ichthyosaurs

    Science.gov (United States)

    Rothschild, B. M.; Xiaoting, Z.; Martin, L. D.

    2012-06-01

    Decompression syndrome (caisson disease or the "the bends") resulting in avascular necrosis has been documented in mosasaurs, sauropterygians, ichthyosaurs, and turtles from the Middle Jurassic to Late Cretaceous, but it was unclear that this disease occurred as far back as the Triassic. We have examined a large Triassic sample of ichthyosaurs and compared it with an equally large post-Triassic sample. Avascular necrosis was observed in over 15 % of Late Middle Jurassic to Cretaceous ichthyosaurs with the highest occurrence (18 %) in the Early Cretaceous, but was rare or absent in geologically older specimens. Triassic reptiles that dive were either physiologically protected, or rapid changes of their position in the water column rare and insignificant enough to prevent being recorded in the skeleton. Emergency surfacing due to a threat from an underwater predator may be the most important cause of avascular necrosis for air-breathing divers, with relative frequency of such events documented in the skeleton. Diving in the Triassic appears to have been a "leisurely" behavior until the evolution of large predators in the Late Jurassic that forced sudden depth alterations contributed to a higher occurrence of bends.

  3. A bird-like skull in a Triassic diapsid reptile increases heterogeneity of the morphological and phylogenetic radiation of Diapsida

    Science.gov (United States)

    Pritchard, Adam C.; Nesbitt, Sterling J.

    2017-10-01

    The Triassic Period saw the first appearance of numerous amniote lineages (e.g. Lepidosauria, Archosauria, Mammalia) that defined Mesozoic ecosystems following the end Permian Mass Extinction, as well as the first major morphological diversification of crown-group reptiles. Unfortunately, much of our understanding of this event comes from the record of large-bodied reptiles (total body length > 1 m). Here we present a new species of drepanosaurid (small-bodied, chameleon-like diapsids) from the Upper Triassic Chinle Formation of New Mexico. Using reconstructions of micro-computed tomography data, we reveal the three-dimensional skull osteology of this clade for the first time. The skull presents many archaic anatomical traits unknown in Triassic crown-group reptiles (e.g. absence of bony support for the external ear), whereas other traits (e.g. toothless rostrum, anteriorly directed orbits, inflated endocranium) resemble derived avian theropods. A phylogenetic analysis of Permo-Triassic diapsids supports the hypothesis that drepanosaurs are an archaic lineage that originated in the Permian, far removed from crown-group Reptilia. The phylogenetic position of drepanosaurids indicates the presence of archaic Permian clades among Triassic small reptile assemblages and that morphological convergence produced a remarkably bird-like skull nearly 100 Myr before one is known to have emerged in Theropoda.

  4. A Triassic to Cretaceous Sundaland-Pacific subduction margin in West Sarawak, Borneo

    Science.gov (United States)

    Breitfeld, H. Tim; Hall, Robert; Galin, Thomson; Forster, Margaret A.; BouDagher-Fadel, Marcelle K.

    2017-01-01

    Metamorphic rocks in West Sarawak are poorly exposed and studied. They were previously assumed to be pre-Carboniferous basement but had never been dated. New 40Ar/39Ar ages from white mica in quartz-mica schists reveal metamorphism between c. 216 to 220 Ma. The metamorphic rocks are associated with Triassic acid and basic igneous rocks, which indicate widespread magmatism. New U-Pb dating of zircons from the Jagoi Granodiorite indicates Triassic magmatism at c. 208 Ma and c. 240 Ma. U-Pb dating of zircons from volcaniclastic sediments of the Sadong and Kuching Formations confirms contemporaneous volcanism. The magmatic activity is interpreted to represent a Triassic subduction margin in westernmost West Sarawak with sediments deposited in a forearc basin derived from the magmatic arc at the Sundaland-Pacific margin. West Sarawak and NW Kalimantan are underlain by continental crust that was already part of Sundaland or accreted to Sundaland in the Triassic. One metabasite sample, also previously assumed to be pre-Carboniferous basement, yielded Early Cretaceous 40Ar/39Ar ages. They are interpreted to indicate resumption of subduction which led to deposition of volcaniclastic sediments and widespread magmatism. U-Pb ages from detrital zircons in the Cretaceous Pedawan Formation are similar to those from the Schwaner granites of NW Kalimantan, and the Pedawan Formation is interpreted as part of a Cretaceous forearc basin containing material eroded from a magmatic arc that extended from Vietnam to west Borneo. The youngest U-Pb ages from zircons in a tuff layer from the uppermost part of the Pedawan Formation indicate that volcanic activity continued until c. 86 to 88 Ma when subduction terminated.

  5. Characteristics of the Triassic Source Rocks of the Aitutu Formation in the (West Timor Basin

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    Asep Kurnia Permana

    2014-12-01

    Full Text Available DOI:10.17014/ijog.v1i3.192The Triassic rocks of the (West Timor Basin have been identified that was mainly deposited in the  marine environment. The fine grained clastics and carbonate  rocks of this Triassic marine  facies are considered to be the most promising source rocks potential in this basin. In this paper we present geochemical and petrographic data from outcrop samples of the Triassic carbonate Aitutu Formation, due to emphasized the organic maturation, kerogen type of the organic matter and the origin of the organic matter.  A representative of selected sample were subjected to the Rock-Eval Pyrolisis, vitrinite reflectance and thermal alteration index, bitumen extraction, were analyzed on the GC-MS. The samples were collected from marine deposit of the Triassic Sequence. The TOC values of the analyzed sample range between rich and rich organic richness (0.51% - 9.16%, wt.%, TOC, which consists mainly of type II and III kerogen and the organic matter consider to be predominantly oil/gas prone and gas prone potential. The thermal maturity assessed from Tmax, TAI, and vitrinite reflectance shows an immature to early peak mature stage of the organic matter. The GC-MS analyses of the biomarkers indicate mainly the organic matter derived from mixed source rocks facies containing alga debris and higher plant terrestrial origin.

  6. The oldest dinosaur? A Middle Triassic dinosauriform from Tanzania.

    Science.gov (United States)

    Nesbitt, Sterling J; Barrett, Paul M; Werning, Sarah; Sidor, Christian A; Charig, Alan J

    2013-02-23

    The rise of dinosaurs was a major event in vertebrate history, but the timing of the origin and early diversification of the group remain poorly constrained. Here, we describe Nyasasaurus parringtoni gen. et sp. nov., which is identified as either the earliest known member of, or the sister-taxon to, Dinosauria. Nyasasaurus possesses a unique combination of dinosaur character states and an elevated growth rate similar to that of definitive early dinosaurs. It demonstrates that the initial dinosaur radiation occurred over a longer timescale than previously thought (possibly 15 Myr earlier), and that dinosaurs and their immediate relatives are better understood as part of a larger Middle Triassic archosauriform radiation. The African provenance of Nyasasaurus supports a southern Pangaean origin for Dinosauria.

  7. Volatile earliest Triassic sulfur cycle

    DEFF Research Database (Denmark)

    Schobben, Martin; Stebbins, Alan; Algeo, Thomas J.

    2017-01-01

    model experiment. Exposure of evaporite deposits having a high δ 34S may account for the source change, with a possible role for the Siberian Traps volcanism by magmatic remobilization of Cambrian rock salt. A high sulfur cycle turnover rate would have left the ocean system vulnerable to development......Marine biodiversity decreases and ecosystem destruction during the end-Permian mass extinction (EPME) have been linked to widespread marine euxinic conditions. Changes in the biogeochemical sulfur cycle, microbial sulfate reduction (MSR), and marine dissolved sulfate concentrations during...... fractionation and point to a more universal control, i.e., contemporaneous seawater sulfate concentration.The MSR-trend transfer function yielded estimates of seawater sulfate of 0.6-2.8mM for the latest Permian to earliest Triassic, suggesting a balanced oceanic S-cycle with equal S inputs and outputs...

  8. Biomechanical comments about Triassic dinosaurs from Brazil

    Directory of Open Access Journals (Sweden)

    Rafael Delcourt

    2012-01-01

    Full Text Available Triassic dinosaurs of Brazil are found in Santa Maria and Caturrita formations, Rio Grande do Sul state, Brazil. There are three species known from the Santa Maria Formation (Staurikosaurus pricei, Saturnalia tupiniquim and Pampadromaeus barberenai, and two from Caturrita Formation (Guaibasaurus candelariensis and Unaysaurus tolentinoi. These dinosaur materials are, for the most part, well preserved and allow for descriptions of musculature and biomechanical studies. The lateral rotation of the Saturnalia femur is corroborated through calculations of muscle moment arms. The enhanced supracetabular crest of Saturnalia, Guaibasaurus, Staurikosaurus, Herrerasaurus ischigualastensis, Efraasia minor and Chormogisaurus novasi suggests that basal dinosaurs may have maintained an inclination of the trunk at least 20º on the horizontal axis. The pectoral girdle articulation of basal sauropodomorphs (Saturnalia and Unaysaurus was established using a new method, the Clavicular Ring, and the scapular blade remains near 60º on the horizontal axis. This is a plesiomorphic condition among sauropodomorphs and is also seen in the articulated plateosauridae Seitaad ruessi. The Brazilian basal dinosaurs were lightweight with a body mass estimated around 18.5 kg for Staurikosaurus, 6.5 kg for Saturnalia, and 17 kg for Guaibasaurus. Pampadromaeus probably weighed 2.5 kg, but measures of its femur are necessary to confirm this hypothesis. The Triassic dinosaurs from Brazil were diversified but shared some functional aspects that were important in an evolutionary context.

  9. Paleomagnetism and magnetic fabric of the Triassic rocks from Spitsbergen

    Science.gov (United States)

    Dudzisz, K.; Szaniawski, R.; Michalski, K.; Manby, G.

    2017-12-01

    Understanding the origin and directions of the natural remanent magnetization and the tectonic deformation pattern reflected in magnetic fabric is of importance for investigation of the West Spitsbergen Fold and Thrust Belt (WSFTB) and its foreland. Previous research carried out on Triassic rocks from the study area concluded that these rocks record a composite magnetization of both, normal and reverse polarity, consisting of a primary Triassic remanence that is overlapped by a secondary post-folding component. Standard paleomagnetic procedures were conducted in order to determine the remanence components and a low-field AMS was applied to assess the degree and pattern of deformation. The AMS results from the WSFTB reveal a magnetic foliation that parallels the bedding planes and a dominantly NNW-SSE oriented magnetic lineation that is sub-parallel to the regional fold axial trend. These results imply a low to moderate degree of deformation and a maximum strain orientation parallel to that of the fold belt. These data are consistent with an orthogonal convergence model for the WSFTB formation. In turn, the magnetic fabric on the undeformed foreland displays a distinct NNE-SSW orientation that we attribute to the paleocurrent direction. Rock-magnetic analyses reveal that the dominant ferrimagnetic carriers are magnetite and titanomagnetite. The Triassic rocks are characterised by complicated NRM patterns often with overlapping unblocking temperature spectra of particular components. The dominant magnetisation is characterised, however, by a steep inclination of 70-80º. The derived paleomagnetic direction from the WSFTB falls on the Jurassic - recent sector of the apparent polar wander path (APWP) of Baltica after tectonic unfolding. These data imply that at least some of the identified secondary components could have originated before the Eurekan folding event (K/Pg), for example, in Early Cretaceous time which corresponds to the period of rifting events on Barents

  10. 57Fe Mössbauer analysis of the Upper Triassic-Lower Jurassic deep-sea chert: Paleo-redox history across the Triassic-Jurassic boundary and the Toarcian oceanic anoxic event

    International Nuclear Information System (INIS)

    Sato, Tomohiko; Isozaki, Yukio; Shozugawa, Katsumi; Seimiya, Kimiko; Matsuo, Motoyuki

    2012-01-01

    We investigated the paleo-redox change across the Triassic-Jurassic (T-J) boundary (∼200 Ma) and the Early Toarcian oceanic anoxic event (T-OAE; ∼183 Ma) recorded in the Upper Triassic to Lower Jurassic pelagic deep-sea cherts in the Inuyama area, Central Japan. The present 57 Fe Mössbauer spectroscopic analysis for these cherts identified five iron species, i.e., hematite (α-Fe 2 O 3 ), pyrite (FeS 2 ), paramagnetic Fe 3 +  , and two paramagnetic Fe 2 +  with different quadrupole splittings. The occurrence of hematite and pyrite in deep-sea cherts essentially indicates primary oxidizing and reducing depositional conditions, respectively. The results confirmed that oxidizing conditions persisted in deep-sea across the T-J boundary. In contrast, across the T-OAE, deep-sea environment shifted to reducing conditions. The first appearance of the gray pyrite-bearing chert marked the onset of the deep-sea oxygen-depletion in the middle Pliensbachian, i.e., clearly before the shallow-sea T-OAE.

  11. Late Triassic sedimentary records in the northern Tethyan Himalaya: Tectonic link with Greater India

    Directory of Open Access Journals (Sweden)

    Huawen Cao

    2018-01-01

    strata. The Tethyan Himalaya was part of the northern passive margin and was not an exotic terrane separated from India during the Permian to Early Cretaceous. This evidence suggests that the Neo-Tethyan ocean opened prior to the Late Triassic and that the Upper Triassic deposits were derived from continental crustal fragments adjacent to the northern passive continental margin of Greater India.

  12. Permo-triassic volcanism in the San Rafael Block (Mendoza province) and its uraniferous potential

    International Nuclear Information System (INIS)

    Kleiman, L.E.

    1993-01-01

    This paper describes the Permo-triassic volcanism in the San Rafael Block, Mendoza, Argentina, which forms part of the Choiyoi province and it represents by large volumes of intermediate to silicic ignimbrites with minor sub volcanic bodies of rhyolites, andesites and basandesites. Three different suites can be distinguished: the first one (Lower Section) of Early Permian age, is composed of dacites and rhyolites (SiO 2 up to 71 %) with minor andesites, the second one (Upper Section) of Late Permian-Early Triassic age is made up of rhyolites (SiO 2 up to 77 %) with some basandesites and andesites, and the third one, of Triassic age is composed of rhyolites (SiO 2 > 75 %) and basandecites. These suites are easily distinguished by means of trace element data and are believed to represent the transition between a subduction-related magmatic arc and an extensional tectonic regime. This tectonic setting is similar to the prevalent during the Cenozoic in the Sierra Occidental of Mexico and is favourable for the development of long-lived hydrothermal systems which lead to economic U concentrations (i.e. Sierra de Pena Blanca). In the San Rafael Block, the Dr. Baulies-Los Reyunos U deposit, which is hosted in volcanic sediments, is associated to the first suite (Lower Section). Although minor U concentrations are known, up to date, to be related to the second and third suites, these rocks are fertile and seen to be potential source for the formation of uranium deposits within a volcanic caldera environment. (Author)

  13. Post-Triassic thermal history of the Tazhong Uplift Zone in the Tarim Basin, Northwest China: Evidence from apatite fission-track thermochronology

    Directory of Open Access Journals (Sweden)

    Caifu Xiang

    2013-11-01

    Full Text Available The Tarim Basin is a representative example of the basins developed in the northwest China that are characterized by multiple stages of heating and cooling. In order to better understand its complex thermal history, apatite fission track (AFT thermochronology was applied to borehole samples from the Tazhong Uplift Zone (TUZ. Twelve sedimentary samples of Silurian to Triassic depositional ages were analyzed from depths coinciding with the apatite partial annealing zone (∼60–120 °C. The AFT ages, ranging from 132 ± 7 Ma (from a Triassic sample to 25 ± 2 Ma (from a Carboniferous sample, are clearly younger than their depositional ages and demonstrate a total resetting of the AFT thermometer after deposition. The AFT ages vary among different tectonic belts and decrease from the No. Ten Faulted Zone (133–105 Ma in the northwest, the Central Horst Zone in the middle (108–37 Ma, to the East Buried Hill Zone in the south (51–25 Ma. Given the low magnitude of post-Triassic burial heating evidenced by low vitrinite reflectance values (Ro < 0.7%, the total resetting of the AFT system is speculated to result from the hot fluid flow along the faults. Thermal effects along the faults are well documented by younger AFT ages and unimodal single grain age distributions in the vicinity of the faults. Permian–early Triassic basaltic volcanism may be responsible for the early Triassic total annealing of those samples lacking connectivity with the fault. The above arguments are supported by thermal modeling results.

  14. Lunge feeding in early marine reptiles and fast evolution of marine tetrapod feeding guilds.

    Science.gov (United States)

    Motani, Ryosuke; Chen, Xiao-hong; Jiang, Da-yong; Cheng, Long; Tintori, Andrea; Rieppel, Olivier

    2015-03-10

    Traditional wisdom holds that biotic recovery from the end-Permian extinction was slow and gradual, and was not complete until the Middle Triassic. Here, we report that the evolution of marine predator feeding guilds, and their trophic structure, proceeded faster. Marine reptile lineages with unique feeding adaptations emerged during the Early Triassic (about 248 million years ago), including the enigmatic Hupehsuchus that possessed an unusually slender mandible. A new specimen of this genus reveals a well-preserved palate and mandible, which suggest that it was a rare lunge feeder as also occurs in rorqual whales and pelicans. The diversity of feeding strategies among Triassic marine tetrapods reached their peak in the Early Triassic, soon after their first appearance in the fossil record. The diet of these early marine tetrapods most likely included soft-bodied animals that are not preserved as fossils. Early marine tetrapods most likely introduced a new trophic mechanism to redistribute nutrients to the top 10 m of the sea, where the primary productivity is highest. Therefore, a simple recovery to a Permian-like trophic structure does not explain the biotic changes seen after the Early Triassic.

  15. An early Late Triassic long-necked reptile with a bony pectoral shield and gracile appendages

    Directory of Open Access Journals (Sweden)

    Jerzy Dzik

    2016-12-01

    Full Text Available Several partially articulated specimens and numerous isolated bones of Ozimek volans gen. et sp. nov., from the late Carnian lacustrine deposits exposed at Krasiejów in southern Poland, enable a reconstruction of most of the skeleton. The unique character of the animal is its enlarged plate-like coracoids presumably fused with sterna. Other aspects of the skeleton seem to be comparable to those of the only known specimen of Sharovipteryx mirabilis from the latest Middle Triassic of Kyrgyzstan, which supports interpretation of both forms as protorosaurians. One may expect that the pectoral girdle of S. mirabilis, probably covered by the rock matrix in its only specimen, was similar to that of O. volans gen. et sp. nov. The Krasiejów material shows sharp teeth, low crescent scapula, three sacrals in a generalized pelvis (two of the sacrals being in contact with the ilium and curved robust metatarsal of the fifth digit in the pes, which are unknown in Sharovipteryx. Other traits are plesiomorphic and, except for the pelvic girdle and extreme elongation of appendages, do not allow to identify any close connection of the sharovipterygids within the Triassic protorosaurians.

  16. Provenance analysis of the Late Triassic Yichuan Basin: constraints from zircon U-Pb geochronology

    Directory of Open Access Journals (Sweden)

    Meng Xianghong

    2018-03-01

    Full Text Available Laser ablation inductively coupled plasma mass spectrometry (LA-ICP-MS U-Pb dating has been performed on detrital zircons from the Chunshuyao Formation sandstone of Yichuan Basin. The ages of 85 detrital zircon grains are divided into three groups: 252-290 Ma, 1740-2000 Ma, and 2400-2600 Ma. The lack of Early Paleozoic and Neoproterozoic U-Pb ages indicates that there is no input from the Qinling Orogen, because the Qinling Orogen is characterized by Paleozoic and Neoproterozoic material. In combination with previous research, we suggest that the source of the Chunshuyao Formation is most likely recycled from previous sedimentary rocks from the North China Craton. In the Late Triassic, the Funiu ancient land was uplifted which prevented source material from the Qinling Orogen. Owing to the Indosinian orogeny, the strata to the east of the North China Craton were uplifted and eroded. The Yichuan Basin received detrital material from the North China Craton.

  17. Multiple sulfur-isotopic evidence for a shallowly stratified ocean following the Triassic-Jurassic boundary mass extinction

    Science.gov (United States)

    Luo, Genming; Richoz, Sylvain; van de Schootbrugge, Bas; Algeo, Thomas J.; Xie, Shucheng; Ono, Shuhei; Summons, Roger E.

    2018-06-01

    The cause of the Triassic-Jurassic (Tr-J) boundary biotic crisis, one of the 'Big Five' mass extinctions of the Phanerozoic, remains controversial. In this study, we analyzed multiple sulfur-isotope compositions (δ33S, δ34S and δ36S) of pyrite and Spy/TOC ratios in two Tr-J successions (Mariental, Mingolsheim) from the European Epicontinental Seaway (EES) in order to better document ocean-redox variations during the Tr-J transition. Our results show that upper Rhaetian strata are characterized by 34S-enriched pyrite, low Spy/TOC ratios, and values of Δ33Spy (i.e., the deviation from the mass-dependent array) lower than that estimated for contemporaneous seawater sulfate, suggesting an oxic-suboxic depositional environment punctuated by brief anoxic events. The overlying Hettangian strata exhibit relatively 34S-depleted pyrite, high Δ33Spy, and Spy/TOC values, and the presence of green sulfur bacterial biomarkers indicate a shift toward to euxinic conditions. The local development of intense marine anoxia thus postdated the Tr-J mass extinction, which does not provide support for the hypothesis that euxinia was the main killing agent at the Tr-J transition. Sulfur and organic carbon isotopic records that reveal a water-depth gradient (i.e., more 34S-, 13C-depleted with depth) in combination with Spy/TOC data suggest that the earliest Jurassic EES was strongly stratified, with a chemocline located at shallow depths just below storm wave base. Shallow oceanic stratification may have been a factor for widespread deposition of black shales, a large positive shift in carbonate δ13C values, and a delay in the recovery of marine ecosystems following the Tr-J boundary crisis.

  18. Geochronologic evidence of a large magmatic province in northern Patagonia encompassing the Permian-Triassic boundary

    Science.gov (United States)

    Luppo, Tomás; López de Luchi, Mónica G.; Rapalini, Augusto E.; Martínez Dopico, Carmen I.; Fanning, Christopher M.

    2018-03-01

    The Los Menucos Complex (northern Patagonia) consists of ∼6 km thick succession of acidic and intermediate volcanic and pyroclastic products, which has been traditionally assigned to the Middle/Late Triassic. New U/Pb (SHRIMP) zircon crystallization ages of 257 ± 2 Ma at the base, 252 ± 2 Ma at an intermediate level and 248 ± 2 Ma near the top of the sequence, indicate that this volcanic event took place in about 10 Ma around the Permian-Triassic boundary. This volcanism can now be considered as the effusive terms of the neighboring and coeval La Esperanza Plutono-Volcanic Complex. This indicates that the climax of activity of a large magmatic province in northern Patagonia was coetaneous with the end-Permian mass extinctions. Likely correlation of La Esperanza- Los Menucos magmatic province with similar volcanic and plutonic rocks across other areas of northern Patagonia suggest a much larger extension than previously envisaged for this event. Its age, large volume and explosive nature suggest that the previously ignored potential role that this volcanism might have played in climatic deterioration around the Permian-Triassic boundary should be investigated.

  19. Recovery from the most profound mass extinction of all time.

    Science.gov (United States)

    Sahney, Sarda; Benton, Michael J

    2008-04-07

    The end-Permian mass extinction, 251 million years (Myr) ago, was the most devastating ecological event of all time, and it was exacerbated by two earlier events at the beginning and end of the Guadalupian, 270 and 260 Myr ago. Ecosystems were destroyed worldwide, communities were restructured and organisms were left struggling to recover. Disaster taxa, such as Lystrosaurus, insinuated themselves into almost every corner of the sparsely populated landscape in the earliest Triassic, and a quick taxonomic recovery apparently occurred on a global scale. However, close study of ecosystem evolution shows that true ecological recovery was slower. After the end-Guadalupian event, faunas began rebuilding complex trophic structures and refilling guilds, but were hit again by the end-Permian event. Taxonomic diversity at the alpha (community) level did not recover to pre-extinction levels; it reached only a low plateau after each pulse and continued low into the Late Triassic. Our data showed that though there was an initial rise in cosmopolitanism after the extinction pulses, large drops subsequently occurred and, counter-intuitively, a surprisingly low level of cosmopolitanism was sustained through the Early and Middle Triassic.

  20. Anachronistic facies from a drowned Lower Triassic carbonate platform: Lower member of the Alwa Formation (Ba'id Exotic), Oman Mountains

    Science.gov (United States)

    Woods, Adam D.; Baud, Aymon

    2008-09-01

    The lower member of the Alwa Formation (Lower Olenekian), found within the Ba'id Exotic in the Oman Mountains (Sultanate of Oman), consists of ammonoid-bearing, pelagic limestones that were deposited on an isolated, drowned carbonate platform on the Neotethyan Gondwana margin. The strata contain a variety of unusual carbonate textures and features, including thrombolites, Frutexites-bearing microbialites that contain synsedimentary cements, matrix-free breccias surrounded by isopachous calcite cement, and fissures and cavities filled with large botryoidal cements. Thrombolites are found throughout the study interval, and occur as 0.5-1.0 m thick lenses or beds that contain laterally laterally-linked stromatactis cavities. The Frutexites-bearing microbialites occur less frequently, and also form lenses or beds, up to 30 cm thick; the microbialites may be laminated, and often developed on hardgrounds. In addition, the Frutexites-bearing microbialites also contain synsedimentary calcite cement crusts and botryoids (typically fracturing of the limestone and the precipitation of large, botryoidal aragonite cements in fissures that cut across the primary fabric. Environmental conditions, specifically palaeoxygenation and the degree of calcium carbonate supersaturation, likely controlled whether the thrombolites (high level of calcium carbonate supersaturation associated with vertical mixing of water masses and dysoxic conditions) or Frutexites-bearing microbialites (low level of calcium carbonate supersaturation associated with anoxic conditions and deposition below a stable chemocline) formed. The results of this study point to continued environmental stress in the region during the Early Triassic that likely contributed to the uneven recovery from the Permian-Triassic mass extinction.

  1. Breeding Young as a Survival Strategy during Earth’s Greatest Mass Extinction

    Science.gov (United States)

    Botha-Brink, Jennifer; Codron, Daryl; Huttenlocker, Adam K.; Angielczyk, Kenneth D.; Ruta, Marcello

    2016-04-01

    Studies of the effects of mass extinctions on ancient ecosystems have focused on changes in taxic diversity, morphological disparity, abundance, behaviour and resource availability as key determinants of group survival. Crucially, the contribution of life history traits to survival during terrestrial mass extinctions has not been investigated, despite the critical role of such traits for population viability. We use bone microstructure and body size data to investigate the palaeoecological implications of changes in life history strategies in the therapsid forerunners of mammals before and after the Permo-Triassic Mass Extinction (PTME), the most catastrophic crisis in Phanerozoic history. Our results are consistent with truncated development, shortened life expectancies, elevated mortality rates and higher extinction risks amongst post-extinction species. Various simulations of ecological dynamics indicate that an earlier onset of reproduction leading to shortened generation times could explain the persistence of therapsids in the unpredictable, resource-limited Early Triassic environments, and help explain observed body size distributions of some disaster taxa (e.g., Lystrosaurus). Our study accounts for differential survival in mammal ancestors after the PTME and provides a methodological framework for quantifying survival strategies in other vertebrates during major biotic crises.

  2. Compound-specific carbon isotopes from Earth’s largest flood basalt eruptions directly linked to the end-Triassic mass extinction

    Science.gov (United States)

    Whiteside, Jessica H.; Olsen, Paul E.; Eglinton, Timothy; Brookfield, Michael E.; Sambrotto, Raymond N.

    2010-01-01

    A leading hypothesis explaining Phanerozoic mass extinctions and associated carbon isotopic anomalies is the emission of greenhouse, other gases, and aerosols caused by eruptions of continental flood basalt provinces. However, the necessary serial relationship between these eruptions, isotopic excursions, and extinctions has never been tested in geological sections preserving all three records. The end-Triassic extinction (ETE) at 201.4 Ma is among the largest of these extinctions and is tied to a large negative carbon isotope excursion, reflecting perturbations of the carbon cycle including a transient increase in CO2. The cause of the ETE has been inferred to be the eruption of the giant Central Atlantic magmatic province (CAMP). Here, we show that carbon isotopes of leaf wax derived lipids (n-alkanes), wood, and total organic carbon from two orbitally paced lacustrine sections interbedded with the CAMP in eastern North America show similar excursions to those seen in the mostly marine St. Audrie’s Bay section in England. Based on these results, the ETE began synchronously in marine and terrestrial environments slightly before the oldest basalts in eastern North America but simultaneous with the eruption of the oldest flows in Morocco, a CO2 super greenhouse, and marine biocalcification crisis. Because the temporal relationship between CAMP eruptions, mass extinction, and the carbon isotopic excursions are shown in the same place, this is the strongest case for a volcanic cause of a mass extinction to date. PMID:20308590

  3. Late Triassic tropical climate of Pangea: Carbon isotopic and other insights into the rise of dinosaurs

    Science.gov (United States)

    Whiteside, J. H.; Lindström, S.; Irmis, R. B.; Glasspool, I.; Schaller, M. F.; Dunlavey, M.; Nesbitt, S. J.; Smith, N. D.; Turner, A. H.

    2015-12-01

    The rarity and species-poor nature of early dinosaurs and their relatives at low paleolatitudes persisted for 30 million years after their origin and 10-15 million years after they became abundant and speciose at higher latitudes. New environmental reconstructions from stable carbon isotope ratios of preserved organic matter (δ13Corg), atmospheric pCO2 data based on the δ13C of soil carbonate, palynological, and wildfire data from charcoal from early dinosaur-bearing strata at low paleolatitudes in western North America show that variations in δ13Corg and palynomorph ecotypes are tightly correlated, displaying large and high-frequency excursions. These variations occurred within an environment characterized by elevated and increasing atmospheric pCO2, pervasive wildfires, and rapidly fluctuating extreme climatic conditions. Whereas pseudosuchian archosaur-dominated communities were able to persist in these same regions until the end-Triassic, the large-bodied, fast-growing tachymetabolic dinosaurian herbivores were not. We hypothesize that the greater resources required by the herbivores made it difficult from them to adapt to the unstable conditions at low paleolatitudes in the Late Triassic.

  4. Nitrogen isotope and trace metal analyses from the Mingolsheim core (Germany): Evidence for redox variations across the Triassic-Jurassic boundary

    Science.gov (United States)

    Quan, Tracy M.; van de Schootbrugge, Bas; Field, M. Paul; Rosenthal, Yair; Falkowski, Paul G.

    2008-06-01

    The Triassic-Jurassic (T-J) boundary was one of the largest but least understood mass extinction events in the Phanerozoic. We measured bulk organic nitrogen and carbon isotopes and trace metal concentrations from a core near Mingolsheim (Germany) to infer paleoenvironmental conditions associated with this event. Poorly fossiliferous claystones across the boundary have relatively low δ15N values and low concentrations of redox-sensitive elements, characteristic of an oxic environment with significant terrestrial input. The Early Jurassic features enrichment in δ15N coincident with high redox-sensitive element concentrations, indicating an increase in water column denitrification and decreased oxygen concentrations. These redox state variations are concordant with shifts in abundance and species composition in terrestrial and marine microflora. We propose that the mass extinction at the T-J boundary was caused by a series of events resulting in a long period of stratification, deep-water hypoxia, and denitrification in this region of the Tethys Ocean basin.

  5. Wildfire Activity Across the Triassic-Jurassic Boundary in the Polish Basin: Evidence from New Fossil Charcoal & Carbon-isotope Data

    Science.gov (United States)

    Pointer, R.; Belcher, C.; Hesselbo, S. P.; Hodbod, M.; Pieńkowski, G.

    2017-12-01

    New fossil charcoal abundance and carbon-isotope data from two sedimentary cores provide new evidence of extreme environmental conditions in the Polish Basin during the Latest Triassic to Earliest Jurassic. Sedimentary cores from the Polish Basin provide an excellent record of terrestrial environmental conditions across the Triassic-Jurassic Boundary, a time of climatic extremes. Previous work has shown that the marine realm was affected by a large perturbation to the carbon cycle across the Triassic-Jurassic Boundary (manifested by large negative and positive carbon-isotope excursions) and limited records of charcoal abundance and organic geochemistry have indicated important changes in fire regime in the coeval ecosystems. Here we present two new carbon-isotope records generated from fossil plant matter across the Triassic-Jurassic boundary, and present new charcoal records. The charcoal abundance data confirm that there was variation in wildfire activity during the Late Triassic-Early Jurassic in the Polish Basin. Peaks in the number of fossil charcoal fragments present occur in both sedimentary cores, and increases in fossil charcoal abundance are linked to wildfires, signalling a short-lived rise in wildfire activity. Fossil charcoal abundance does not appear to be fully controlled by total organic matter content, depositional environment or bioturbation. We argue that increased wildfire activity is likely caused by an increase in ignition of plant material as a result of an elevated number of lightning strikes. Global warming (caused by a massive input of carbon into the atmosphere, as indicated by carbon-isotope data) can increase storm activity, leading to increased numbers of lightning strikes. Previous Triassic-Jurassic Boundary wildfire studies have found fossil charcoal abundance peaks at other northern hemisphere sites (Denmark & Greenland), and concluded that they represent increases in wildfire activity in the earliest Jurassic. Our new charcoal and

  6. Age and provenance of Triassic to Cenozoic sediments of West and Central Sarawak, Malaysia

    Science.gov (United States)

    Breitfeld, H. Tim; Galin, Thomson; Hall, Robert

    2015-04-01

    Sarawak is located on the northern edge of Sundaland in NW Borneo. West and Central Sarawak include parts of the Kuching and Sibu Zones. These contain remnants of several sedimentary basins with ages from Triassic to Cenozoic. New light mineral, heavy mineral and U-Pb detrital zircon ages show differences in provenance reflecting the tectonic evolution of the region. The oldest clastic sediments are Triassic (Sadong Formation and its deep marine equivalent Kuching Formation). They were sourced by a Triassic (Carnian to Norian) volcanic arc and reworked Paleoproterozoic detritus derived from Cathaysialand. The Upper Jurassic to Cretaceous Pedawan Formation is interpreted as forearc basin fill with distinctive zircon populations indicating subduction beneath present-day West Sarawak which initiated in the Late Jurassic. Subsequent subduction until the early Late Cretaceous formed the Schwaner Mountains magmatic arc. After collision of SW Borneo and other microcontinental fragments with Sundaland in the early Late Cretaceous, deep marine sedimentation (Pedawan Formation) ceased, and there was uplift forming the regional Pedawan-Kayan unconformity. Two episodes of extension followed and were responsible for basin development on land in West Sarawak from the latest Cretaceous onwards, probably in a pull-apart setting. The first episode is associated with sediments of the Kayan Group, deposited in the Latest Cretaceous (Maastrichtian) to Eocene, and the second episode with Upper Eocene sediments of the Ketungau Basin. Zircon ages indicate volcanic activity throughout the Early Cenozoic in NW Borneo, and inherited zircon ages indicate reworking of Triassic and Cretaceous rocks. A large deep marine basin, the Rajang Basin, was north of the Lupar Line Fault in Central Sarawak (Sibu Zone) from the Late Cretaceous to the Late Eocene. Zircons from sediments of the Rajang Basin indicate they have similar ages and provenance to contemporaneous terrestrial sediments of the Kayan

  7. Late Permian to Triassic intraplate orogeny of the southern Tianshan and adjacent regions, NW China

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    Wei Ju

    2014-01-01

    Based on previous studies and recent geochronogical data, we suggest that the final collision between the Tarim Craton and the North Asian continent occurred during the late Carboniferous. Therefore, the Permian was a period of intracontinental environment in the southern Tianshan and adjacent regions. We propose that an earlier, small-scale intraplate orogenic stage occurred in late Permian to Triassic time, which was the first intraplate process in the South Tianshan Orogen and adjacent regions. The later large-scale and well-known Neogene to Quaternary intraplate orogeny was induced by the collision between the India subcontinent and the Eurasian plate. The paper presents a new evolutionary model for the South Tianshan Orogen and adjacent regions, which includes seven stages: (I late Ordovician–early Silurian opening of the South Tianshan Ocean; (II middle Silurian–middle Devonian subduction of the South Tianshan Ocean beneath an active margin of the North Asian continent; (III late Devonian–late Carboniferous closure of the South Tianshan Ocean and collision between the Kazakhstan-Yili and Tarim continental blocks; (IV early Permian post-collisional magmatism and rifting; (V late Permian–Triassic the first intraplate orogeny; (VI Jurassic–Palaeogene tectonic stagnation and (VII Neocene–Quaternary intraplate orogeny.

  8. A theropod tooth from the Late Triassic of southern Africa

    Indian Academy of Sciences (India)

    An isolated, large recurved and finely serrated tooth found associated with the prosauropod Euskelosaurus from the Late Triassic part of the Elliot Formation is described here. It is compared to the Triassic thecodonts and carnivorous dinosaurs and its possible affinity is discussed. The tooth possibly belongs to a basal ...

  9. A high resolution magnetostratigraphic profile across the Permian-Triassic boundary in the Southern Sydney Basin, eastern Australia

    Science.gov (United States)

    Belica, M. E.; Tohver, E.; Nicoll, R.; Denyszyn, S. W.; Pisarevsky, S.; George, A. D.

    2016-12-01

    The Permian-Triassic boundary (PTB) is associated with the largest mass extinction in Phanerozoic geologic history. Despite several decades of intense study, there is ongoing debate regarding the exact timing of extinction and the global correlation of marine and terrestrial P-T sections. The terrestrial record is hampered by a lack of index fossils; however, magnetostratigraphy offers an opportunity for correlation because it relies on the global synchronicity of magnetic reversals. A magnetostratigraphic profile across the Permian-Triassic boundary has been obtained from a stratigraphically continuous terrestrial section in the Southern Sydney Basin of eastern Australia. The 60 m section is located within the Narrabeen Group, which consists of fluvial to lacustrine sandstones and mudstones. Paleomagnetic samples were collected at one meter intervals to determine a detailed reversal record. Samples were stepwise thermally demagnetized to isolate a primary remanence, and magnetic susceptibility was measured in the field at 30 cm intervals with values ranging from -0.047-2.50 (10-3 SI units). Three normal and three reverse magnetozones were detected after removal of a low temperature overprint, and the results show good agreement with the Global Magnetic Polarity Timescale as well as marine Permian-Triassic sections where the PTB is well constrained. Furthermore, a reverse polarity subchron has been identified within the normal magnetozone spanning the PTB similar to results published from the Netherlands and China. The magnetic stratigraphy suggests that the Narrabeen Group was deposited during the late Changhsingian to early Induan, and provides a revised placement of the PTB in the lower Wombarra Claystone. Integration of the magnetostratigraphy with existing isotopic datasets suggests that the terrestrial extinction in eastern Australia occurred 7.5 m below the PTB in the Changhsingian Coalcliff Sandstone. A tuff within a coal seam underlying the Coalcliff

  10. Soft-sediment deformation structures in cores from lacustrine slurry deposits of the Late Triassic Yanchang Fm. (central China

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    Yang Renchao

    2016-09-01

    Full Text Available The fine-grained autochthonous sedimentation in the deep part of a Late Triassic lake was frequently interrupted by gravity-induced mass flows. Some of these mass flows were so rich in water that they must have represented slurries. This can be deduced from the soft-sediment deformation structures that abound in cores from these lacustrine deposits which constitute the Yanchang Fm., which is present in the Ordos Basin (central China.

  11. The Early to Middle Triassic continental–marine transition of NW Bulgaria: sedimentology, palynology and sequence stratigraphy

    OpenAIRE

    Ajdanlijsky George; Götz Annette E.; Strasser André

    2018-01-01

    Sedimentary facies and cycles of the Triassic continental–marine transition of NW Bulgaria are documented in detail from reference sections along the Iskar river gorge between the villages of Tserovo and Opletnya. The depositional environments evolved from anastomosing and meandering river systems in the Petrohan Terrigenous Group to mixed fluvial and tidal settings in the Svidol Formation, and to peritidal and shallow-marine conditions in the Opletnya Member of the Mogila Formation. For the ...

  12. The end-Triassic mass extinction: A new correlation between extinction events and δ13C fluctuations from a Triassic-Jurassic peritidal succession in western Sicily

    Science.gov (United States)

    Todaro, Simona; Rigo, Manuel; Randazzo, Vincenzo; Di Stefano, Pietro

    2018-06-01

    A new δ13Ccarb curve was obtained from an expanded peritidal succession in western Sicily and was used to investigate the relationships between isotopic signatures and biological events on carbonate platforms across the Triassic-Jurassic boundary (TJB). The resulting curve shows two main negative carbon isotopic excursions (CIEs) that fit well with the "Initial" and "Main" CIEs that are recognized worldwide and linked to the End-Triassic Extinction (ETE). In the studied section, the first negative CIE marks the disappearance of the large megalodontids, which were replaced by small and thin-shelled specimens, while the "Main" CIE corresponds to the last occurrence (LO) of the megalodontids and, approximately 50 m upsection, to the total demise of the Rhaetian benthic foraminifer community. Upward, the carbon curve shows a positive trend (ca. +1‰) and a gradual recovery of the benthic communities after an approximately 10 m-thick barren interval populated only by the problematic alga Thaumatoporella parvovesiculifera. A comparison between the Mt. Sparagio δ13Ccarb curve and other coeval Ccarb and Corg curves from carbonate platform, ramp and deep basin successions indicates similar isotopic trends; however, the diverse magnitudes and responses of benthic communities confirm that the carbon cycle perturbations have been globally significant, and were influenced by external forces such as CAMP volcanism. The multiphase nature of the extinction pulses could have been caused by local environmental changes related to transgression/regression phenomena. Overall, this study adds new data and a new timing to the effect of the acidification process on carbon productivity and benthic communities in different environments across the TJB.

  13. New clade of enigmatic early archosaurs yields insights into early pseudosuchian phylogeny and the biogeography of the archosaur radiation.

    Science.gov (United States)

    Butler, Richard J; Sullivan, Corwin; Ezcurra, Martín D; Liu, Jun; Lecuona, Agustina; Sookias, Roland B

    2014-06-10

    The origin and early radiation of archosaurs and closely related taxa (Archosauriformes) during the Triassic was a critical event in the evolutionary history of tetrapods. This radiation led to the dinosaur-dominated ecosystems of the Jurassic and Cretaceous, and the high present-day archosaur diversity that includes around 10,000 bird and crocodylian species. The timing and dynamics of this evolutionary radiation are currently obscured by the poorly constrained phylogenetic positions of several key early archosauriform taxa, including several species from the Middle Triassic of Argentina (Gracilisuchus stipanicicorum) and China (Turfanosuchus dabanensis, Yonghesuchus sangbiensis). These species act as unstable 'wildcards' in morphological phylogenetic analyses, reducing phylogenetic resolution. We present new anatomical data for the type specimens of G. stipanicicorum, T. dabanensis, and Y. sangbiensis, and carry out a new morphological phylogenetic analysis of early archosaur relationships. Our results indicate that these three previously enigmatic taxa form a well-supported clade of Middle Triassic archosaurs that we refer to as Gracilisuchidae. Gracilisuchidae is placed basally within Suchia, among the pseudosuchian (crocodile-line) archosaurs. The approximately contemporaneous and morphologically similar G. stipanicicorum and Y. sangbiensis may be sister taxa within Gracilisuchidae. Our results provide increased resolution of the previously poorly constrained relationships of early archosaurs, with increased levels of phylogenetic support for several key early pseudosuchian clades. Moreover, they falsify previous hypotheses suggesting that T. dabanensis and Y. sangbiensis are not members of the archosaur crown group. The recognition of Gracilisuchidae provides further support for a rapid phylogenetic diversification of crown archosaurs by the Middle Triassic. The disjunct distribution of the gracilisuchid clade in China and Argentina demonstrates that early

  14. Records of Triassic volcanism in Pangean Great Lakes, and implications for reconstructing the distal effects of Large Igneous Provinces

    Science.gov (United States)

    Whiteside, J. H.; Percival, L.; Kinney, S.; Olsen, P. E.; Mather, T. A.; Philpotts, A.

    2017-12-01

    Documentation of the precise timing of volcanic eruptions in sedimentary records is key for linking volcanic activity to both historical and geological episodes of environmental change. Deposition of tuffs in sediments, and sedimentary enrichment of trace metals linked to igneous processes, are both commonly used for such correlations. In particular, sedimentary mercury (Hg) enrichments have been used as a marker for volcanic activity from Large Igneous Provinces (LIPs) to support their link to episodes of major climate change and mass extinction in the geological record. However, linking such enrichments to a specific eruption or eruption products is often challenging or impossible. In this study, the mercury records from two exactly contemporaneous latest Triassic-earliest Jurassic rift lakes are presented. Both sedimentary records feature igneous units proposed to be related to the later (Early Jurassic) stages of volcanism of the Central Atlantic Magmatic Province (CAMP). These CAMP units include a small tuff unit identified by thin-section petrology and identified at 10 localities over a distance of over 200 km, and a major CAMP basalt flow overlying this tuff (and dated at 200.916±0.064 Ma) which is also known across multiple sedimentary basins in both North America and Morocco and is thought to have been emplaced about 120 kyr after the tuff. A potential stratigraphic correlation between Hg enrichments and the igneous units is considered, and compared to the established records of mercury enrichments from the latest Triassic that are thought to be coeval with the earlier stages of CAMP volcanism. Investigating the Hg records of sedimentary successions containing tuffs and basalt units is an important step for demonstrating whether the mercury emissions from specific individual volcanic eruptions in the deep past can be identified in the geological record, and are thus important tools for interpreting the causes of associated past geological events, such as

  15. Permian-Triassic maturation and multistage migration of hydrocarbons in the Assistência Formation (Irati Subgroup, Paraná Basin, Brazil: implications for the exploration model

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    António Mateus

    Full Text Available New lines of geological evidence strongly suggest that the main period of hydrocarbon maturation within Assistência Formation should be Permian-Triassic, stimulated by a high geothermal gradient that also sustained various manifestations of hydrothermal activity. Three main stages of fluid/hydrocarbon migration can also be inferred on the basis of multiscale observations: confined flow in late Permian to Triassic times, depending on the local build-up of fluid pressures; heterogeneous flow in Lower Cretaceous, triggered by a rejuvenated temperature gradient assisted by the early developed permeability conditions; and a late flow possibly driven by local pressure gradients, after complete cooling of dolerite dykes/sills. The early maturation and multistage migration of hydrocarbons have significant consequences in the design of exploration models to be applied in Paraná Basin.

  16. The Triassic metabasalts of Dudin Krš, near Kosovska Mitrovica, Serbia

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    Memović Emin

    2003-01-01

    Full Text Available This paper presents the geochemical characteristics of the metabasalts of Dudin Krš, near Kosovska Mitrovica. The Dudin Krš is the easternmost occurrence of Middle Triassic rift-related volcanic rocks in the Dinarides. Generally, these rocks show similarities to other Triassic volcanic rocks of the Dinarides. They are high-magnesian, ol and ne-normative basalts, with low Zr/Nb and medium/low Ti/Zr and Ti/Y ratios. They exhibit transitional geochemical characteristics, between E-MORB and subalkaline basalts of continental rift zones. Their presence and geochemical affinity is evidence of rifting processes along the continental slope of the Adria block during Middle Triassic.

  17. Precession-driven monsoon variability at the Permian-Triassic boundary — Implications for anoxia and the mass extinction

    Science.gov (United States)

    Winguth, Arne; Winguth, Cornelia

    2013-06-01

    By the end of the Late Permian, most continents had collided to form the supercontinent of Pangea. The associated climatic changes at the Permian-Triassic boundary coincided with the most severe mass extinction in the Phanerozoic. One extinction hypothesis favors a climatic response to an increase in large-scale volcanism resulting in ocean stagnation and widespread anoxia with fatal consequences for marine and land organisms. Recent interpretations of geochemical data suggest that orbitally-driven periodic upwelling of toxic hydrogen-sulfide rich water masses contributed to the extinction of species. In this paper, we use the Community Climate System Model (CCSM3) in order to explore the effect of eccentricity-modulated changes of the precession on the strength of Pangean megamonsoons and their impact on productivity and oxygen distribution. The climate model simulates high variability in monsoonal precipitation, trade winds and equatorial upwelling in response to precessional extremes, leading to remarkable fluctuations in the export of carbon from the euphotic zone and hence reduction in dissolved oxygen concentrations in subsurface layers. These findings are in general agreement with increased primary productivity, intensified euxinia within the oxygen-minimum zone, and decimation of the radiolarian zooplankton community as inferred from Japanese marine sections. Strong changes in river run-off linked to precipitation oscillations possibly led to a high variability in the nutrient supply to the Tethys Ocean, thus affecting regional productivity and oxygen distribution. The model results suggest that orbital variability in the sedimentary record and the associated extinction of species are related rather to periodic anoxia in near surface-to-intermediate depth than to widespread anoxic events in the Panthalassic deep-sea.

  18. Recovery of Carbonate Ecosystems Following the End-Triassic Mass Extinction: Insights from Mercury Anomalies and Their Relationship to the Central Atlantic Magmatic Province

    Science.gov (United States)

    Corsetti, F. A.; Thibodeau, A. M.; Ritterbush, K. A.; West, A. J.; Yager, J. A.; Ibarra, Y.; Bottjer, D. J.; Berelson, W.; Bergquist, B. A.

    2015-12-01

    Recent high-resolution age dating demonstrates that the end-Triassic mass extinction overlapped with the eruption of the Central Atlantic Magmatic Province (CAMP), and the release of CO2 and other volatiles to the atmosphere has been implicated in the extinction. Given the potentially massive release of CO2, ocean acidification is commonly considered a factor in the extinction and the collapse of shallow marine carbonate ecosystems. However, the timing of global marine biotic recovery versus the CAMP eruptions is more uncertain. Here, we use Hg concentrations and Hg/TOC ratios as indicators of CAMP volcanism in continental shelf sediments, the primary archive of faunal data. In Triassic-Jurassic strata, Muller Canyon, Nevada, Hg and Hg/TOC levels are low prior to the extinction, rise sharply in the extinction interval, peak just prior to the appearance of the first Jurassic ammonite, and remain above background in association with a depauperate (low diversity) earliest Jurassic fauna. The return of Hg to pre-extinction levels is associated with a significant pelagic and benthic faunal recovery. We conclude that significant biotic recovery did not begin until CAMP eruptions ceased. Furthermore, the initial benthic recovery in the Muller Canyon section involves the expansion of a siliceous sponge-dominated ecosystem across shallow marine environments, a feature now known from other sections around the world (e.g., Peru, Morocco, Austria, etc.). Carbonate dominated benthic ecosystems (heralded by the return of abundant corals and other skeletal carbonates) did not recover for ~1 million years following the last eruption of CAMP, longer than the typical duration considered for ocean acidification events, implying other factors may have played a role in carbonate ecosystem dynamics after the extinction.

  19. A new stem-neopterygian fish from the Middle Triassic of China shows the earliest over-water gliding strategy of the vertebrates.

    Science.gov (United States)

    Xu, Guang-Hui; Zhao, Li-Jun; Gao, Ke-Qin; Wu, Fei-Xiang

    2013-01-07

    Flying fishes are extraordinary aquatic vertebrates capable of gliding great distances over water by exploiting their enlarged pectoral fins and asymmetrical caudal fin. Some 50 species of extant flying fishes are classified in the Exocoetidae (Neopterygii: Teleostei), which have a fossil record no older than the Eocene. The Thoracopteridae is the only pre-Cenozoic group of non-teleosts that shows an array of features associated with the capability of over-water gliding. Until recently, however, the fossil record of the Thoracopteridae has been limited to the Upper Triassic of Austria and Italy. Here, we report the discovery of exceptionally well-preserved fossils of a new thoracopterid flying fish from the Middle Triassic of China, which represents the earliest evidence of an over-water gliding strategy in vertebrates. The results of a phylogenetic analysis resolve the Thoracopteridae as a stem-group of the Neopterygii that is more crown-ward than the Peltopleuriformes, yet more basal than the Luganoiiformes. As the first record of the Thoracopteride in Asia, this new discovery extends the geographical distribution of this group from the western to eastern rim of the Palaeotethys Ocean, providing new evidence to support the Triassic biological exchanges between Europe and southern China. Additionally, the Middle Triassic date of the new thoracopterid supports the hypothesis that the re-establishment of marine ecosystems after end-Permian mass extinction is more rapid than previously thought.

  20. U-Pb Geochronology of non-marine Upper Triassic strata of the Colorado Plateau (western North America): implications for stratigraphic correlation and paleoenvironmental reconstruction

    Science.gov (United States)

    Rasmussen, C.; Mundil, R.; Irmis, R. B.; Keller, C. B.; Giesler, D.; Gehrels, G. E.

    2017-12-01

    The Triassic is a critical period in Earth history that witnessed the origin of modern ecosystems and frequent climate fluctuations, as well as major environmental events such as flood basalt volcanism and bolide impacts. The Chinle Formation contains a primary non-marine archive for past ecosystems in North America due to its fossil richness and well-studied sedimentology. Moreover, within these highly fossiliferous strata, a biotic turnover has been reported that has been hypothesized to coincide with one or more of the aforementioned environmental events. Unfortunately, few radioisotopic ages have been published for the Late Triassic, limiting our ability for lithological and paleoenvironmental correlations. In addition, the superposition of the Chinle Formation remains illusive due to frequent lateral facies changes and discontinuous outcrops across the Colorado Plateau. The 520 m long core 1A of the Colorado Plateau Coring Project from Petrified Forest National Park (PFNP) (Arizona) provides, for the first time, a continuous section of these early Mesozoic sedimentary strata. Many of the sand- and siltstones from this continuous succession throughout most of the Upper Triassic Chinle Formation contain euhedral zircons suitable for U-Pb analyses. We analyzed >300 crystals each from 10 samples using LA-ICPMS; these results indicated abundant Late Triassic crystals that appear to be closely associated with the depositional age of the host rock. We then selected the youngest grains from these samples to obtain precise CA-TIMS U-Pb single zircon ages in order to constrain the maximum depositional ages (using quantitative methods) of these formations. We are able to revise the proposed time scale (based on outcrop samples) for Upper Triassic strata at PFNP and evaluate whether the biotic turnover observed within the Sonsela Member of these strata coincides with the Manicouagan bolide impact event. This revised chronostratigraphic framework allows intercalibration

  1. Tectono-sedimentary evolution of the Permian-Triassic extension event in the Zagros basin (Iran): results from analogue modelling

    Science.gov (United States)

    Madani-kivi, M.; Zulauf, G.

    2015-12-01

    Since the 1970s, the largest oil and gas reservoirs have been discovered in the Permian-Early Triassic formationsin Saudi Arabia. Thus, this time period is important for the discovery of new oil reserves in Iran. The Arabian passivecontinental margin has undergone lithospheric extension during the Permian-Triassic, which led to the formation of theNeo-Tethys. The aim of this paper is to describe the development of the continental rift basin in the Zagros region basedon the tectono-sedimentological evolution. We have studied well-log data to specify the distribution of synrift depositsin the Zagros and have related this information to the modelling. Environmental changes indicated by various sedimentarysequences, from a siliciclastic basin to a carbonate platform setting, are described. The Cambrian Hormuz salt, whichoverlies the metamorphosed Precambrian basement, becomes effective as a basal detachment layer influencing the styleof overburden deformation during the Permian-Triassic extension event. We have investigated the formation of variousstructures linked to the presence or absence of the Hormuz layer by analogue modelling and relating these structures to theLate Palaeozoic sedimentation. Based on results of the analogue modelling, we argue that the basal detachment layer (Hormuzseries) has contributed to the various structural styles of the extensional basin development in the Fars domain and theLorestan domain.

  2. Vegetation history across the Permian-Triassic boundary in Pakistan (Amb section, Salt Range)

    NARCIS (Netherlands)

    Schneebeli-Hermann, E.; Kurschner, W.M.; Kerp, H.; Bomfleur, B.; Hochuli, P.A.; Bucher, H.; Ware, D.; Roohi, G.

    2015-01-01

    Hypotheses about the Permian–Triassic floral turnover range from a catastrophic extinction of terrestrial plant communities to a gradual change in floral composition punctuated by intervals indicating dramatic changes in the plant communities. The shallow marine Permian–Triassic succession in the

  3. The terminal Permian in European Russia: Vyaznikovian Horizon, Nedubrovo Member, and Permian-Triassic boundary

    Science.gov (United States)

    Lozovsky, V. R.; Balabanov, Yu. P.; Karasev, E. V.; Novikov, I. V.; Ponomarenko, A. G.; Yaroshenko, O. P.

    2016-07-01

    The comprehensive analysis of the data obtained on terrestrial vertebrata, ostracods, entomologic fauna, megaflora, and microflora in deposits of the Vyaznikovian Horizon and Nedubrovo Member, as well as the paleomagnetic data measured in enclosing rocks, confirms heterogeneity of these deposits. Accordingly, it is necessary to distinguish these two stratons in the terminal Permian of the East European Platform. The combined sequence of Triassic-Permian boundary deposits in the Moscow Syneclise, which is considered to be the most complete sequence in the East European Platform, is as follows (from bottom upward): Vyatkian deposits; Vyaznikovian Horizon, including Sokovka and Zhukovo members; Nedubrovo Member (Upper Permian); Astashikha and Ryabi members of the Vokhmian Horizon (Lower Triassic). None of the sequences of Permian-Triassic boundary deposits known in the area of study characterizes this sequence in full volume. In the north, the Triassic deposits are underlain by the Nedubrovo Member; in the south (the Klyazma River basin), the sections are underlain by the Vyaznikovian Horizon. The Permian-Triassic boundary adopted in the General Stratigraphic Scale of Russia for continental deposits of the East European platform (the lower boundary of the Astashikha Member) is more ancient than the one adopted in the International Stratigraphic Chart. The same geological situation is observed in the German Basin and other localities where Triassic continental deposits are developed. The ways of solving this problem are discussed in this article.

  4. NEW TRIASSIC ASTEROIDEA (ECHINODERMATA SPECIMENS AND THEIR EVOLUTIONARY SIGNIFICANCE

    Directory of Open Access Journals (Sweden)

    DANIEL B. BLAKE

    2017-04-01

    Full Text Available The Paleozoic-Mesozoic transition saw the disappearance of asteroid stem groups and the ascent of the crown group, but late Paleozoic and Triassic asteroids are rare and transition events are poorly documented. Three new Middle and Late Triassic specimens augment existing data; included are a specimen of Trichasteropsis weissmanni from Germany, a specimen of Trichasteropsis? sp. indet. from Italy, and a possible member of the extant Poraniidae from Slovenia. Presence of a small ossicle at the interbrachial midline and adjacent to the marginal series of the new T. weissmanni specimen is consistent with similar expressions not only of other trichasteropsids but also occurrence of two interbrachial ossicles in Paleozoic, stem-group asterozoans; presence is in turn consistent with a hypothesis of derivation of the axillary/odontophore coupling from two ossicles rather than direct derivation of the crown-group odontophore from a single stem-group axillary. Morphology of Trichasteropsis? sp. indet., including, for example, the evenly-tapering arms are reminiscent of those of diverse crown-group asteroids whereas the enlarged distal arms of T. weissmanni are unique, the morphology of T? sp. indet. thereby potentially indicative of a plesiomorphic, stemward positioning within the Trichasteropsiidae. The range of the Poraniidae is tentatively extended to the Carnian. Similarities shared by the Poraniidae and the Trichasteropsiidae suggest stemward positioning within crown-group diversification; however, known Triassic fossils do not appear closely related to extant taxa identified in recent molecular studies as basal within the crown-group. A temperate climate is suggested as preferred by the Triassic asteroids rather than a tropical, warmer one.

  5. Neutron activation analysis of Permian-Triassic boundary layer samples at the Selong Site in China

    International Nuclear Information System (INIS)

    Miyamoto, Y.; Sakamoto, K.; Mingqing, W.

    1997-01-01

    Thirty samples from a limestone stratum across the Permian-Triassic (P-Tr) boundary layer in China were analyzed for 30 elements by instrumental neutron activation analysis, wavelength dispersive X-ray fluorescence and ICP-MS, and also for mineral compositions with a powder X-ray diffractometer. The depth profile was found to indicate a sudden change of elemental and mineral compositions across the P-Tr boundary. Also the profile showed several peaks in elemental concentrations in the lower Permian layered samples as well as in the overlying Triassic strata, which are associated with the change of mineral compositions. Elemental profiles were found to be classified into 4 groups and to give some insights in the geochemical records. Ir is far less abundant (0.1 ppt) compared with that of the K-T boundaries (10 ppb), and the Ir/Co ratio is outside the K-T and Cl chondrite trends. This change of elementary profile is suggestive of the internal causes rather than the external ones such as an asteroid impact for the mass extinction at the P-Tr boundary. (author)

  6. Revision of the Dysmorphoptilidae (Hemiptera: Cicadomorpha: Prosboloidea) of the Queensland Triassic-Part 2.

    Science.gov (United States)

    Lambkin, Kevin J

    2016-03-15

    The extinct hemipteran family Dysmorphoptilidae was a major component of the Triassic insect fauna of Queensland preserved at the Denmark Hill, Dinmore, Mount Crosby and Gayndah fossil insect sites. A total of 13 species have now been identified, of which eight species in five genera were examined in the first part of this revision. This second part revises the remaining five species in three genera. Eoscartoides Evans, 1956 (= Mesonirvana Evans, 1956, syn. nov.), comprising Eoscartoides bryani Evans, 1956 (= Mesonirvana abrupta Evans, 1956, syn. nov.) (Mount Crosby), Eoscartoides orthocladus (Tillyard, 1922) comb. nov. (Denmark Hill), and Eoscartoides dmitryi sp. nov. (Dinmore), is distinguished by a strongly developed arc-like strigil in the basal costal space, a very short stem of RA, and a deeply forked M1+2. The monotypic Eoscarterella Evans, 1956, with type species Eoscarterella media Evans, 1956 (Mount Crosby), has a strongly lobate tegmen with peculiar surface sculpture and M1+2 simple. Eoscartoides and Eoscarterella differ from most dysmorphoptilids in having more or less lobate tegmina with even margins (without the antero-apical emargination so typical of the family), as well as the early entry of RA1 into the costal margin and the associated extensive and antero-apically positioned RA2. These characters are also shared with two other dysmorphoptilids, the Australian Permian Belmontocarta Evans and an unnamed Triassic species from Kyrgyzstan, and the four thus form a distinct subgroup within the family. On the other hand, the monotypic Trifidella Evans, 1956 (= Alotrifidus Evans, 1956, syn. nov.), with type species Trifidella perfecta Evans, 1956 (= Alotrifidus interruptus Evans, 1956, syn. nov.) (Mount Crosby), is a more typical dysmorphoptilid with a distinct emargination, RA entering the margin much more apically, and RA2 of limited extent. Trifidella is presumably the sister of the Queensland Triassic Dysmorphoptiloides Evans, sharing the basal

  7. Upper Triassic spongiomorph and coral association dredged off the northwestern Australian shelf

    Science.gov (United States)

    Stanley, G. D.

    Upper Triassic corals and spongiomorphs dredged during BMR Cruise 95 from the Rowley Terrace, offshore Canning Basin of northwestern Australia, indicate possible new occurrences of reef facies. These are comparable to counterparts known from the Norhtern Limestone Alps of central Europe. A branching spongiomorph, represented by the genus Spongiomorpha sp. and two coral taxa, Pamiroseris rectilamellosa (Winkler) and Retiophyllia tellae (Stoppani), are reported herein. Collectively, these fossils indicate a Late Triassic (Noria-Rhaetian) age. Although different in taxonomic composition, the fauna compares with one previously reported from a Late Triassic Ocean Drilling Project reef site (site 764) on the Wombat Plateau, some 350 km to the west. The Rowley Terrace occurrences may represent an eastward extension of the Wombat reefs, developed along the rifted margin of Gondwana.

  8. Aggregations and parental care in the Early Triassic basal cynodonts Galesaurus planiceps and Thrinaxodon liorhinus

    Directory of Open Access Journals (Sweden)

    Sandra C. Jasinoski

    2017-01-01

    Full Text Available Non-mammaliaform cynodonts gave rise to mammals but the reproductive biology of this extinct group is still poorly known. Two exceptional fossils of Galesaurus planiceps and Thrinaxodon liorhinus, consisting of juveniles closely associated with an adult, were briefly described more than 50 years ago as examples of parental care in non-mammaliaform cynodonts. However, these two Early Triassic fossils have largely been excluded from recent discussions of parental care in the fossil record. Here we re-analyse these fossils in the context of an extensive survey of other aggregations found in these two basal cynodont taxa. Our analysis revealed six other unequivocal cases of aggregations in Thrinaxodon, with examples of same-age aggregations among immature or adult individuals as well as mixed-age aggregations between subadult and adult individuals. In contrast, only one additional aggregation of Galesauruswas identified. Taking this comprehensive survey into account, the two previously described cases of parental care in Galesaurus and Thrinaxodon are substantiated. The juveniles are the smallest specimens known for each taxon, and the size difference between the adult and the two associated juveniles is the largest found for any of the aggregations. The juveniles of Thrinaxodon are approximately only 37% of the associated adult size; whereas in Galesaurus, the young are at least 60% of the associated adult size. In each case, the two juvenile individuals are similar in size, suggesting they were from the same clutch. Even though parental care was present in both Galesaurus and Thrinaxodon, intraspecific aggregations were much more common in Thrinaxodon, suggesting it regularly lived in aggregations consisting of both similar and different aged individuals.

  9. Death in Guizhou — Late Triassic drowning of the Yangtze carbonate platform

    Science.gov (United States)

    Enos, Paul; Wei Jiayong; Lehrmann, Daniel J.

    1998-06-01

    The Yangtze platform in south China formed a stable palaeogeographic element from the Late Proterozoic to the end of the Middle Triassic with deposition of shallow-water carbonates during much of this time. A portion of the Yangtze platform in south-central Guizhou drowned at the transition from Permian to Triassic, as the south-adjacent Nanpanjiang basin encroached about 100 km northward, but a new, stable platform margin was established that persisted through the Early and Middle Triassic. This long history as a stable carbonate platform ended at the transition from the Ladinian to the Carnian. The latest Ladinian rocks, the Yangliujing Formation, are 490 m of shoaling-upward carbonate cycles of grapestone and bioclastic grainstone, fenestral limestone, and stromatolitic dolomudstone, commonly overprinted by extensive subaerial diagenesis. The beginning of the Carnian is marked by a rapid transition to medium-dark-grey, nodular lime mudstones containing ammonoids, conodonts and thin-shelled bivalves, the Zhuganpo Formation. The upper part of this thin pelagic limestone contains many muddy intraclasts, some slightly bored and encrusted, indicating incipient cementation. The overlying Wayao Formation is a condensed black shale with thin interbeds of dark-grey, manganiferous lime mudstone near the base. Ammonoids, conodonts, thin-shelled bivalves, and articulated crinoid stems are abundant. Fine-grained greywacke with sole marks forms prominent bundles within grey, calcareous shale in the overlying Laishike Formation. Ammonoids and thin-shelled bivalves occur sporadically in this 810-m-thick unit. Calcareous shale with thicker-shelled bivalves and packages of cleaner, coarser-grained sandstone characterize the Banan Formation, 460 m thick. The sandstone units generally coarsen and thicken upward, with ripples, medium-scale trough cross-beds, and rare U-tube burrows. Quartzose, coal-bearing siliciclastics 690 m thick form the overlying Huobachong Formation. Thick

  10. The Colorado Plateau Coring Project: A Continuous Cored Non-Marine Record of Early Mesozoic Environmental and Biotic Change

    Science.gov (United States)

    Irmis, Randall; Olsen, Paul; Geissman, John; Gehrels, George; Kent, Dennis; Mundil, Roland; Rasmussen, Cornelia; Giesler, Dominique; Schaller, Morgan; Kürschner, Wolfram; Parker, William; Buhedma, Hesham

    2017-04-01

    The early Mesozoic is a critical time in earth history that saw the origin of modern ecosystems set against the back-drop of mass extinction and sudden climate events in a greenhouse world. Non-marine sedimentary strata in western North America preserve a rich archive of low latitude terrestrial ecosystem and environmental change during this time. Unfortunately, frequent lateral facies changes, discontinuous outcrops, and a lack of robust geochronologic constraints make lithostratigraphic and chronostratigraphic correlation difficult, and thus prevent full integration of these paleoenvironmental and paleontologic data into a regional and global context. The Colorado Plateau Coring Project (CPCP) seeks to remedy this situation by recovering a continuous cored record of early Mesozoic sedimentary rocks from the Colorado Plateau of the western United States. CPCP Phase 1 was initiated in 2013, with NSF- and ICDP-funded drilling of Triassic units in Petrified Forest National Park, northern Arizona, U.S.A. This phase recovered a 520 m core (1A) from the northern part of the park, and a 240 m core (2B) from the southern end of the park, comprising the entire Lower-Middle Triassic Moenkopi Formation, and most of the Upper Triassic Chinle Formation. Since the conclusion of drilling, the cores have been CT scanned at the University of Texas - Austin, and split, imaged, and scanned (e.g., XRF, gamma, and magnetic susceptibility) at the University of Minnesota LacCore facility. Subsequently, at the Rutgers University Core Repository, core 1A was comprehensively sampled for paleomagnetism, zircon geochronology, petrography, palynology, and soil carbonate stable isotopes. LA-ICPMS U-Pb zircon analyses are largely complete, and CA-TIMS U-Pb zircon, paleomagnetic, petrographic, and stable isotope analyses are on-going. Initial results reveal numerous horizons with a high proportion of Late Triassic-aged primary volcanic zircons, the age of which appears to be a close

  11. The first Triassic dipteran (Insecta) from South America, with review of Hennigmatidae.

    Science.gov (United States)

    Lara, María Belén; Lukashevich, Elena D

    2013-01-01

    The first Triassic dipteran from South America is described based on an isolated wing from the lower Upper Triassic deposits of Argentina (Mendoza Province, Potrerillos Formation, Quebrada del Durazno locality). Trihennigma zavattierii gen. et sp. nov. is a member of the Mesozoic family Hennigmatidae, previously recorded only from Eurasia. A key for the genera and species of Hennigmatidae is provided and systematic position of the taxa is discussed.

  12. Review of potential host rocks for radioactive wasste disposal in the southeast United States: Triassic basin subregion

    International Nuclear Information System (INIS)

    1980-10-01

    Based on an evaluation of existing information, areas were identified within the Triassic basins of the southeastern United States with geologic properties considered favorable for containment of radioactive waste. The study region included both exposed and buried Triassic basins from Maryland to Georgia. These basins are long, narrow northeast-trending troughs filled with continental deposits derived from Paleozoic and Precambrian metamorphic and igneous rocks bordering the basins. The rocks are predominantly red in color and consist mainly of fanglomerates, conglomerates, arkosic sandstones, siltstones, claystones, shales, and argillites. The investigation identified 14 exposed and 5 buried basins within the study region. Candidate areas for further investigation were identified which meet the broad general criteria for tectonic stability, slow ground water movement, and long flow paths to the biosphere. These include: the Danville Triassic Basin in Virginia; the Dan River, Durham, and Wadesboro Triassic Basins in North Carolina; and the buried Florence and Dunbarton Triassic Basins in South Carolina. Other rock types in the southeast may prove more or less suitable as host rocks for a repository, but the available data suggest that the argillaceous Triassic rocks offer sufficient promise to be considered for additional study

  13. The new Permian-Triassic paleomagnetic pole for the East European Platform corrected for inclination shallowing

    Science.gov (United States)

    Fetisova, A. M.; Veselovskiy, R. V.; Scholze, F.; Balabanov, Yu. P.

    2018-01-01

    The results of detailed paleomagnetic studies in seven Upper Permian and Lower Triassic reference sections of East Europe (Middle Volga and Orenburg region) and Central Germany are presented. For each section, the coefficient of inclination shallowing f (King, 1955) is estimated by the Elongation-Inclination (E-I) method (Tauxe and Kent, 2004) and is found to vary from 0.4 to 0.9. The paleomagnetic directions, corrected for the inclination shallowing, are used to calculate the new Late Permian-Early Triassic paleomagnetic pole for the East European Platform (N = 7, PLat = 52.1°, PLong = 155.8°, A95 = 6.6°). Based on this pole, the geocentric axial dipole hypothesis close to the Paleozoic/Mesozoic boundary is tested by the single plate method. The absence of the statistically significant distinction between the obtained pole and the average Permian-Triassic (P-Tr) paleomagnetic pole of the Siberian Platform and the coeval pole of the North American Platform corrected for the opening of the Atlantic (Shatsillo et al., 2006) is interpreted by us as evidence that 250 Ma the configuration of the magnetic field of the Earth was predominantly dipolar; i.e., the contribution of nondipole components was at most 10% of the main magnetic field. In our opinion, the hypothesis of the nondipolity of the geomagnetic field at the P-Tr boundary, which has been repeatedly discussed in recent decades (Van der Voo and Torsvik, 2001; Bazhenov and Shatsillo, 2010; Veselovskiy and Pavlov, 2006), resulted from disregarding the effect of inclination shallowing in the paleomagnetic determinations from sedimentary rocks of "stable" Europe (the East European platform and West European plate).

  14. Basin geodynamics and sequence stratigraphy of Upper Triassic to Lower Jurassic deposits of Southern Tunisia

    Science.gov (United States)

    Carpentier, Cédric; Hadouth, Suhail; Bouaziz, Samir; Lathuilière, Bernard; Rubino, Jean-Loup

    2016-05-01

    Aims of this paper are to propose a geodynamic and sequential framework for the late Triassic and early Jurassic of and south Tunisia and to evidence the impact of local tectonics on the stratigraphic architecture. Facies of the Upper Triassic to Lower Jurassic of Southern Tunisia have been interpreted in terms of depositional environments. A sequential framework and correlation schemes are proposed for outcrops and subsurface transects. Nineteen middle frequency sequences inserted in three and a half low frequency transgression/regression cycles were evidenced. Despite some datation uncertainties and the unknown durations of Lower Jurassic cycles, middle frequency sequences appear to be controlled by eustasy. In contrast the tectonics acted as an important control on low frequency cycles. The Carnian flooding was certainly favored by the last stages of a rifting episode which started during the Permian. The regression accompanied by the formation of stacked angular unconformities and the deposition of lowstand deposits during the late Carnian and Norian occured during the uplift and tilting of the northern basin margins. The transpressional activity of the Jeffara fault system generated the uplift of the Tebaga of Medenine high from the late Carnian and led to the Rhaetian regional angular Sidi Stout Unconformity. Facies analysis and well-log correlations permitted to evidence that Rhaetian to Lower Jurassic Messaoudi dolomites correspond to brecciated dolomites present on the Sidi Stout unconformity in the North Dahar area. The Early-cimmerian compressional event is a possible origin for the global uplift of the northern African margin and Western Europe during the late Carnian and the Norian. During the Rhaetian and the early Jurassic a new episode of normal faulting occured during the third low frequency flooding. This tectonosedimentary evolution ranges within the general geodynamic framework of the north Gondwana margin controlled by the opening of both

  15. Conodont zonation of Lower Triassic strata in Slovenia

    Directory of Open Access Journals (Sweden)

    Tea Kolar-Jurkovšek

    2015-12-01

    Full Text Available The paper presents the results of a conodont study carried out in the Triassic strata in the area of the Slovenian part of the Southern Alps, External Dinarides and the Transition region between the External and Internal Dinarides. The following conodont zones have been distinguished: Hindeodus praeparvus Z., H. parvus Z., Isarcicella lobata Z., I. staeschei – I. isarcica Z., H. postparvus Z., Hadrodontina aequabilis Z., Ha. anceps Z., Eurygnathodus costatus Z., Neospathodus planus Z., N. robustus Z., Platyvillosus corniger Z., Pl. regularis Z., Pachycladina obliqua Z., Foliella gardenae Z., Triassospathodus hungaricus Z., T. symmetricus Z., N. robustispinus - T. homeri Z. and T. triangularis Z. The introduced conodont zonation spans from the Induan, including the Permian-Triassic boundary interval to the late Olenekian and is valid for the shallow shelf environments of western Tethys.

  16. The quest for chron E23r at Partridge Island, bay of Fundy, Canada: CAMP emplacement postdates the end-Triassic extinction event at the North American craton

    NARCIS (Netherlands)

    Deenen, M.H.L.; Krijgsman, W.; Ruhl, M.

    2011-01-01

    The Partridge Island stratigraphic section at the Bay of Fundy, Maritime Canada, reveals a continental sedimentary succession with the end-Triassic mass extinction level closely followed by basalts of the Central Atlantic Magmatic Province (CAMP). New Paleomagnetic data show that a short reverse

  17. Development of a high resolution chemostratigraphy for the Late Triassic-Early Jurassic Newark Basin

    Science.gov (United States)

    Kinney, S.; Olsen, P. E.; Chang, C.

    2017-12-01

    The 6.7 km of continuous core recovered from the paleo-tropical Triassic-Jurassic Newark rift basin during the Newark Basin Coring Project (NBCP) has provided a wealth of data since the conclusion of drilling 25 years ago. These cores comprise the longest ( 30 Myr) continuously-cored record of orbitally-paced environmental change and have informed our understanding in several different areas including tropical climate change, history of CO­2, mass extinctions, the geological time scale, and solar system dynamics. Despite the utility of NBCP cores for these endeavors, a critical missing dataset is a comprehensive characterization of their geochemical variations relevant to paleoenvironmental and paleoclimatic interests, largely a consequence of the cost of analyses at an appropriate resolution using conventional techniques. With the advent of new technology permitting the rapid acquisition of reliable geochemical data, such limitations may no longer be an obstacle for constructing a high-resolution chemostratigraphic record for the NBCP. We present the results of a proof-of-concept study using both ICP-MS-calibrated scanning ITRAX XRF and handheld Laser Induced Breakdown Spectroscopy (LIBS) using the SciAps Z-300. We will show elemental abundances at resolutions as high as 500 mm obtained using these methods from correlative sections of the Titusville and Nursery cores (Lockatong Fm.). These sections are sufficiently long to capture orbital variations and include the range of lithologies present throughout the entire section. Our preliminary results are consistent with previous, semi-quantitative means (e.g., depth ranks) of assessing Milankovitch-scale orbital variations and are also consistent with core and hole geophysical data, demonstrating that these methods can acquire meaningful geochemical data from the entire NBCP. With continued work, we aim to provide an objective characterization of orbitally-paced lake level cyclicity using geochemical proxy

  18. OLDEST STEM TELEOSTEI FROM THE LATE LADINIAN (MIDDLE TRIASSIC OF SOUTHERN CHINA

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    ANDREA TINTORI

    2015-11-01

    Full Text Available The origin of the largest modern vertebrate group, the Teleostei, saw major refinements in the last decades, thanks to newly discovered and stratigraphically closely spaced Triassic Lagerstätten. Here we report the oldest Pholidophoriformes (stem teleosts that were collected during a large scale yet detailed excavation of Upper Ladinian (Middle Triassic marine deposits in Xingyi City, Guizhou Province, China. Taxonomic comparisons support the erection of a new pholidophorid genus, Malingichthys gen. nov., with two species. The new genus shows a partially fused skull roof, a preopercular bone with a hockey-stick shape and, for the first time in Pholidophoridae, supraneural elements. Most Triassic marine vertebrate clades (fishes and reptiles, Malingichthys included first emerged in the South China Block, with Late Ladinian most showing an important faunal transformation that was strengthened by our last findings. The material here described is about 2 million years older than the previous records for pholidophorids. 

  19. Comments on Triassic pterosaurs with discussion about ontogeny and description of new taxa

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    Alexander W.A. Kellner

    2015-06-01

    Full Text Available Eudimorphodon ranzii was the first Triassic pterosaur to be described and several specimens have been referred to this taxon mainly based on the presence of multicuspid teeth. Since this dental feature has been observed in several other pterosaurs, the revision of some specimens assigned to Eudimorphodon shows that they represent new taxa as follows: Arcticodactyluscromptonellus (comb. nov., Austriadraco dallavecchiai (gen. et sp. nov. and Bergamodactyluswildi (gen. et sp. nov.. A preliminary analysis of pterosaur ontogeny resulted in the recognition of six distinct ontogenetic stages (OS1-6. According to this classification, the holotype of Arcticodactyluscromptonellus has reached OS2, and although being ontogenetically much younger than others, the conspicuous anatomical differences lead to its exclusion from Eudimorphodon. The holotypes of Austriadraco dallavecchiai,Bergamodactyluswildi and Carniadactylus rosenfeldi have reached at least OS5, which demonstrates that the anatomical differences among them cannot be explained by ontogeny. Moreover, Bergamodactyluswildi reaches about 60% of the maximized wingspan of Carniadactylus rosenfeldi and further concurs that these specimens collected in distinct Triassic Islands of Europe are not conspecific. The present study increases the diversity of Triassic flying reptiles and further pushes the origins of this clade back to at least the Middle Triassic.

  20. Was it the CO2 or the sulfur that did it? The mechanism of mass extinction of continental vertebrates at the end-Triassic extinction.

    Science.gov (United States)

    Olsen, P. E.; Schaller, M. F.; Kent, D. V.; Et-Touhami, M.

    2012-04-01

    Eruption of the giant Central Atlantic Magmatic Province (CAMP) is temporally linked to the end-Triassic extinction (ETE) event (1,2,3,4,5). Proposed killing mechanisms for plants and marine biota have included both CO2 (4) and sulfur aerosols (3,6). Here we examine the kill mechanisms of the CAMP for land animals where we seek to explain the selectivity of the extinctions. One striking aspect of Late Triassic continental communities is the prominent latitudinal provinciality with diverse crurotarsians (crocodile-relatives) and other non-dinosaurs in the tropics and much higher dinosaur diversity in the higher latitudes (7). Only a very few crurotarsian lineages survived the ETE, and a near-homogenization of continental vertebrate assemblages ensued. Background high CO2 concentration in the Late Triassic resulted in lack of polar ice and probably very high-temperature tropical continental interiors. While a doubling of CO2 associated with CAMP eruptions produced a few degree increase in average temperatures (depending on the sensitivity), plausibly leading to some tropical lethality, how this produced a mass extinction in higher, cooler latitudes is harder to explain. While successive CAMP eruptions contributed to CO2 doublings over hundreds to thousands of years, taking two orders of magnitude longer to return to background, EACH major CAMP eruption produced an abrupt sulfur aerosol cooling lasting several years or decades, depending eruption duration. But there were many of these coolings as opposed to the few super-greenhouse warmings suggested by the CO2 proxy data (4,5). The scale of these sulfur injections would have been orders of magnitude larger than anything seen historically (8) possibly leading to freezing tropical temperatures. Both cruotarsians and dinosaurs as well as most other "reptiles", with their presumably uricotelic nitrogen waste systems (9) were been relatively resistant to heat induced water stress, but the former lack insolation, while the

  1. Pre-Cretaceous Agaricomycetes yet to be discovered: Reinvestigation of a putative Triassic bracket fungus from southern Germany

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    A. P. Kiecksee

    2012-08-01

    Full Text Available Agaricomycetes are major components of extant terrestrial ecosystems; however, their fruiting bodies are exceedingly rare as fossils. Reinvestigation of a peculiar fossil from Late Triassic sediments of southern Germany interpreted as a bracket fungus revealed that this fossil in fact represents a wood abnormality, resulting from injury to the cambium and subsequent callus growth in a Baieroxylon -like ginkgoalean wood. As a result, the fossil record of the Agaricomycetes does not yet pre-date the Early Cretaceous, suggesting a late diversification of basidiomycetes possessing large fruiting bodies. doi:10.1002/mmng.201200006

  2. Astronomical timescale calibration for the Permian-Triassic boundary transition interval from global correlation of cyclic marine sequences

    Science.gov (United States)

    Huang, C.; Hinnov, L. A.; Tong, J.; Chen, Z.

    2011-12-01

    The mass extinctions near the Permian-Triassic boundary (PTB) resulted in the greatest dying of life on Earth. The cause of this catastrophe remains enigmatic. High-resolution chronology is crucial to understanding the recorded pattern of biotic evolution and possible causes for the extinctions. Magnetic susceptibility (MS) data from Shangsi, South China shows evidence for astronomical forcing through the PTB interval, with strong 405-kyr cycling. This allows development of an astrochronology for the PTB interval based on the 405-kyr orbital eccentricity metronome that has been proposed for the Mesozoic timescale. Radioisotope dating combined with the 405-kyr tuned MS series from Shangsi shows that the 405-kyr-cycle predominates throughout the PTB interval. In the Permian segment, ~100-kyr cyclicity dominates, and the 100-kyr-scale MS maxima correlate with high-amplitude precession-scale MS variations. Minima in the ~1.5-Myr, 405-kyr and ~100-kyr cycles converge at 252.6 Ma, approximately 200 kyr before the onset of the main mass extinction near the PTB. In the Triassic aftermath, the recorded astronomical signal is different, with predominant 405-kyr cycles and loss of 100 kyr cyclicity, and appearance of ~33 kyr (obliquity scale) cyclicity; 100-kyr cyclicity strengthens again 2 Myr later. This pattern indicates a change in the response of the depositional environment (or magnetic susceptibility) to astronomical forcing before and after the mass extinction interval. The astrochronology interpolates the timescale between the radioisotopically determined absolute dates; this facilitates estimation of ages for specific events in the PTB crisis, including magnetic reversals, biozone boundaries, and the mass extinctions. An estimated ~700 kyr duration for the Mass Extinction Interval (MEI) at Shangsi based on the 405-kyr tuning is supported by eccentricity-tuned estimates of three other sections in China (Meishan, Huangzhishan, and Heping), and two Alpine sections

  3. Lithofacies palaeogeography and biostratigraphy of the lowermost horizons of the Middle Triassic Hallstatt Limestones (Argolis Peninsula, Greece

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    Fotini A. Pomoni

    2013-07-01

    Taking into consideration the present location of the Hallstatt Formation, in the context of the Hellenides, an area suitable for the deposition of the Hallstatt Limestones, should be located between the sub-Pelagonian (western part of the Pelagonian zone and Pindos geotectonic zones, which during the Triassic corresponded to a platform slope and a deep ocean, respectively. The widespread Middle Triassic Han Bulog Limestones (ammonoid-bearing pelagic limestones from Triassic successions of the Eastern Alps (Dinarides, Hellenides may have formed partly in similar slope environments.

  4. The origin and early evolution of dinosaurs.

    Science.gov (United States)

    Langer, Max C; Ezcurra, Martin D; Bittencourt, Jonathas S; Novas, Fernando E

    2010-02-01

    The oldest unequivocal records of Dinosauria were unearthed from Late Triassic rocks (approximately 230 Ma) accumulated over extensional rift basins in southwestern Pangea. The better known of these are Herrerasaurus ischigualastensis, Pisanosaurus mertii, Eoraptor lunensis, and Panphagia protos from the Ischigualasto Formation, Argentina, and Staurikosaurus pricei and Saturnalia tupiniquim from the Santa Maria Formation, Brazil. No uncontroversial dinosaur body fossils are known from older strata, but the Middle Triassic origin of the lineage may be inferred from both the footprint record and its sister-group relation to Ladinian basal dinosauromorphs. These include the typical Marasuchus lilloensis, more basal forms such as Lagerpeton and Dromomeron, as well as silesaurids: a possibly monophyletic group composed of Mid-Late Triassic forms that may represent immediate sister taxa to dinosaurs. The first phylogenetic definition to fit the current understanding of Dinosauria as a node-based taxon solely composed of mutually exclusive Saurischia and Ornithischia was given as "all descendants of the most recent common ancestor of birds and Triceratops". Recent cladistic analyses of early dinosaurs agree that Pisanosaurus mertii is a basal ornithischian; that Herrerasaurus ischigualastensis and Staurikosaurus pricei belong in a monophyletic Herrerasauridae; that herrerasaurids, Eoraptor lunensis, and Guaibasaurus candelariensis are saurischians; that Saurischia includes two main groups, Sauropodomorpha and Theropoda; and that Saturnalia tupiniquim is a basal member of the sauropodomorph lineage. On the contrary, several aspects of basal dinosaur phylogeny remain controversial, including the position of herrerasaurids, E. lunensis, and G. candelariensis as basal theropods or basal saurischians, and the affinity and/or validity of more fragmentary taxa such as Agnosphitys cromhallensis, Alwalkeria maleriensis, Chindesaurus bryansmalli, Saltopus elginensis, and

  5. High diversity, low disparity and small body size in plesiosaurs (Reptilia, Sauropterygia from the Triassic-Jurassic boundary.

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    Roger B J Benson

    Full Text Available Invasion of the open ocean by tetrapods represents a major evolutionary transition that occurred independently in cetaceans, mosasauroids, chelonioids (sea turtles, ichthyosaurs and plesiosaurs. Plesiosaurian reptiles invaded pelagic ocean environments immediately following the Late Triassic extinctions. This diversification is recorded by three intensively-sampled European fossil faunas, spanning 20 million years (Ma. These provide an unparalleled opportunity to document changes in key macroevolutionary parameters associated with secondary adaptation to pelagic life in tetrapods. A comprehensive assessment focuses on the oldest fauna, from the Blue Lias Formation of Street, and nearby localities, in Somerset, UK (Earliest Jurassic: 200 Ma, identifying three new species representing two small-bodied rhomaleosaurids (Stratesaurus taylori gen et sp. nov.; Avalonnectes arturi gen. et sp. nov and the most basal plesiosauroid, Eoplesiosaurus antiquior gen. et sp. nov. The initial radiation of plesiosaurs was characterised by high, but short-lived, diversity of an archaic clade, Rhomaleosauridae. Representatives of this initial radiation were replaced by derived, neoplesiosaurian plesiosaurs at small-medium body sizes during a more gradual accumulation of morphological disparity. This gradualistic modality suggests that adaptive radiations within tetrapod subclades are not always characterised by the initially high levels of disparity observed in the Paleozoic origins of major metazoan body plans, or in the origin of tetrapods. High rhomaleosaurid diversity immediately following the Triassic-Jurassic boundary supports the gradual model of Late Triassic extinctions, mostly predating the boundary itself. Increase in both maximum and minimum body length early in plesiosaurian history suggests a driven evolutionary trend. However, Maximum-likelihood models suggest only passive expansion into higher body size categories.

  6. Multiple generations of high salinity formation water in the Triassic Sherwood Sandstone: Wytch Farm oilfield, onshore UK

    International Nuclear Information System (INIS)

    Worden, R.H.; Manning, D.A.C.; Bottrell, S.H.

    2006-01-01

    The origin and heterogeneity of oilfield formation water in the Lower Triassic Sherwood Sandstone at Wytch Farm in the Wessex Basin, UK, have been investigated using production data, detailed water geochemistry and O, S and H stable isotope data. The formation waters are highly saline, NaCl-type brines with TDS values of up to 230,000mg/L. There is a general decrease in salinity from the flanks of the field to the crest with Cl - decreasing from about 136,000 to 109,000mg/L. The Cl/Br ratio of the water shows that salinity was largely derived from the dissolution of Upper Triassic continental evaporites found off-structure to the west and north of the field. The water in the field had a meteoric source although variation in δ 2 H values suggests that there may be meteoric waters of different ages in the oilfield, reflecting recharge under different palaeoclimatic conditions. At the crest of the field, aqueous SO 4 2- resulted from dissolution of anhydrite in the reservoir. In contrast, in other parts of the field there is an indication that some of the dissolved SO 4 2- was derived from oxidation of pyrite at some point on the recharge path of meteoric water to the field. There were two meteoric influx events bringing different Cl - concentrations and different δ 2 H values. The first was probably before the Eocene oil influx and could have occurred in the Lower Cretaceous or early Tertiary. The second meteoric influx event probably occurred after or during oil migration into the Wytch Farm structure since the second meteoric water is found at the flanks of the field adjacent to the regions where salt is found in the stratigraphy. The preservation of heterogeneities in oilfield formation water compositions suggests that there has been little aqueous fluid movement or diffusive flux for over 40 million years. Mass flux has been restricted by density stratification within the aquifer and the very low effective permeability for the aqueous phase in the oil

  7. A NEW LEPIDOSAUROMORPH REPTILE FROM THE MIDDLE TRIASSIC OF THE DOLOMITES (NORTHERN ITALY

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    SILVIO RENESTO

    2003-11-01

    Full Text Available A new genus and species of diapsid reptile is described. The specimen was collected from the Anisian (Middle Triassic succession of Monte Prà della Vacca (Kühwiesenkopf in the Dolomites of Braies (Bolzano/Bozen, Northern Italy. Despite being incomplete, the specimen shows enough characters to allow its placement within Lepidosauriformes, close to the Middle Jurassic genus Marmoretta. The importance of this find lies in the great rarity of lepidosauriforms of Anisian age, an epoch approximating the appearance of the first members of the true Lepidosauria (rhyncocephalians and squamates. The new genus may thus add knowledge to the diversity of early lepidosaurians. Some characters of the skeleton, mainly of the forelimb, suggest a terrestrial, perhaps arboreal life style. 

  8. Mesozoic-Cenozoic evolution of the Zoige depression in the Songpan-Ganzi flysch basin, eastern Tibetan Plateau: Constraints from detrital zircon U-Pb ages and fission-track ages of the Triassic sedimentary sequence

    Science.gov (United States)

    Tang, Yan; Zhang, Yunpeng; Tong, Lili

    2018-01-01

    The Zoige depression is an important depocenter within the northeast Songpan-Ganzi flysch basin, which is bounded by the South China, North China and Qiangtang Blocks and forms the northeastern margin of the Tibetan Plateau. This paper discusses the sediment provenance and Mesozoic-Cenozoic evolution of the Zoige depression in the Songpan-Ganzi flysch basin, eastern Tibetan Plateau, using the detrital zircon U-Pb ages and apatite fission-track data from the Middle to Late Triassic sedimentary rocks in the area. The U-Pb ages of the Middle to Late Triassic zircons range from 260-280 Ma, 429-480 Ma, 792-974 Ma and 1800-2500 Ma and represent distinct source region. Our new results demonstrate that the detritus deposited during the Middle Triassic (Ladinian, T2zg) primarily originated from the Eastern Kunlun and North Qinling Orogens, with lesser contributions from the North China Block. By the Late Triassic (early Carnian, T3z), the materials at the southern margin of the North China Block were generally transported westward to the basin along a river network that flowed through the Qinling region between the North China and South China Blocks: this interpretation is supported by the predominance of the bimodal distribution of 1.8 Ga and 2.5 Ga age peaks and a lack of significant Neoproterozoic zircon. Since the Late Triassic (middle Carnian, T3zh), considerable changes have occurred in the source terranes, such as the cessation of the Eastern Kunlun Orogen and North China Block sources and the rise of the northwestern margin of the Yangtze Block and South Qinling Orogen. These drastic changes are compatible with a model of a sustained westward collision between the South China and North China Blocks during the late Triassic and the clockwise rotation of the South China Block progressively closed the basin. Subsequently, orogeny-associated folds have formed in the basin since the Late Triassic (late Carnian), and the study area was generally subjected to uplifting and

  9. Triassic Sequence Geological Development of the Arctic with focus on Svalbard and the Barents Shelf

    Energy Technology Data Exchange (ETDEWEB)

    Moerk, Atle

    1998-12-31

    Triassic rocks are of great interest for exploration in Arctic areas as they have proved to include both good hydrocarbon source rocks and potential hydrogen reservoir rocks. In this thesis, the stratigraphy and sedimentology of the Arctic Triassic successions are studied within a sequence stratigraphical framework. Inter-regional comparisons throughout the Arctic are based on comparisons of transgressive-regressive sequences. Improved dating of the studied sequences, and the recognition and correlation of sequence boundaries of second and third order, facilitate interpretation of facies distribution and the geological development both within and between the studied areas. Main emphasis is given to the Triassic succession of Svalbard and the Barents Shelf, which through this study is integrated within a circum-Arctic sequence stratigraphical framework. Good correspondence of the Triassic sequence boundaries between the different Arctic areas indicate that they are mainly controlled by eustacy, while decreasing correspondence of the sequence boundaries in the Jurassic and Cretaceous periods indicate that local and large scale tectonism becomes progressively more dominant in the circum-Arctic Realm through the Mesozoic Era. These hypotheses are further discussed. 701 refs., 110 figs., 12 tabs.

  10. Lunge feeding in early marine reptiles and fast evolution of marine tetrapod feeding guilds

    OpenAIRE

    Motani, R; Chen, XH; Jiang, DY; Cheng, L; Tintori, A; Rieppel, O

    2015-01-01

    Traditional wisdom holds that biotic recovery from the end-Permian extinction was slow and gradual, and was not complete until the Middle Triassic. Here, we report that the evolution of marine predator feeding guilds, and their trophic structure, proceeded faster. Marine reptile lineages with unique feeding adaptations emerged during the Early Triassic (about 248 million years ago), including the enigmatic Hupehsuchus that possessed an unusually slender mandible. A new specimen of this genus ...

  11. Models for the rise of the dinosaurs.

    Science.gov (United States)

    Benton, Michael J; Forth, Jonathan; Langer, Max C

    2014-01-20

    Dinosaurs arose in the early Triassic in the aftermath of the greatest mass extinction ever and became hugely successful in the Mesozoic. Their initial diversification is a classic example of a large-scale macroevolutionary change. Diversifications at such deep-time scales can now be dissected, modelled and tested. New fossils suggest that dinosaurs originated early in the Middle Triassic, during the recovery of life from the devastating Permo-Triassic mass extinction. Improvements in stratigraphic dating and a new suite of morphometric and comparative evolutionary numerical methods now allow a forensic dissection of one of the greatest turnovers in the history of life. Such studies mark a move from the narrative to the analytical in macroevolutionary research, and they allow us to begin to answer the proposal of George Gaylord Simpson, to explore adaptive radiations using numerical methods. Copyright © 2014 The Authors. Published by Elsevier Inc. All rights reserved.

  12. Rock magnetism and magnetic fabric of the Triassic rocks from the West Spitsbergen Fold-and-Thrust Belt and its foreland

    Science.gov (United States)

    Dudzisz, Katarzyna; Szaniawski, Rafał; Michalski, Krzysztof; Chadima, Martin

    2018-03-01

    Magnetic fabric and magnetomineralogy of the Early Triassic sedimentary rocks, collected along the length of the West Spitsbergen Fold-and-Thrust Belt (WSFTB) and from subhorizontal beds on its foreland, is presented with the aim to compare magnetic mineralogy of these areas, determine the carriers of magnetic fabric and identify tectonic deformation reflected in the magnetic fabric. Magnetic mineralogy varies and only in part depends on the lithology. The magnetic fabric at all sampling sites is controlled by paramagnetic minerals (phyllosilicates and Fe-carbonates). In the fold belt, it reflects the low degree of deformation in a compressional setting with magnetic lineation parallel to fold axis (NW-SE). This is consistent with pure orthogonal compression model of the WSFTB formation, but it also agrees with decoupling model. Inverse fabric, observed in few sites, is carried by Fe-rich carbonates. In the WSFTB foreland, magnetic lineation reflects the Triassic paleocurrent direction (NE-SW). The alternation between normal and inverse magnetic fabric within the stratigraphic profile could be related to sedimentary cycles.

  13. Upper Triassic limestones from the northern part of Japan: new insights on the Panthalassa Ocean and Hokkaido Island

    Science.gov (United States)

    Peyrotty, Giovan; Peybernes, Camille; Ueda, Hayato; Martini, Rossana

    2017-04-01

    In comparison with the well-known Tethyan domain, Upper Triassic limestones from the Panthalassa Ocean are still poorly known. However, these carbonates represent a unique opportunity to have a more accurate view of the Panthalassa Ocean during the Triassic. Their study will allow comparison and correlation of biotic assemblages, biostratigraphy, diagenesis, and depositional settings of different Triassic localities from Tethyan and Panthalassic domains. Moreover, investigation of these carbonates will provide data for taxonomic revisions and helps to better constrain palaeobiogeographic models. One of the best targets for the study of these carbonates is Hokkaido Island (north of Japan). Indeed, this island is a part of the South-North continuity of Jurassic to Paleogene accretionary complexes, going from the Philippines to Sakhalin Island (Far East Russia). Jurassic and Cretaceous accretionary complexes of Japan and Philippines contain Triassic mid-oceanic seamount carbonates from the western Panthalassa Ocean (Onoue & Sano, 2007; Kiessling & Flügel, 2000). They have been accreted either as isolated limestone slabs or as clasts and boulders, and are associated with mudstones, cherts, breccias and basaltic rocks. Two major tectonic units forming Hokkaido Island and containing Triassic limestones have been accurately explored and extensively sampled: the Oshima Belt (west Hokkaido) a Jurassic accretionary complex, and the Cretaceous Sorachi-Yezo Belt (central Hokkaido). The Sorachi-Yezo Belt is composed of Cretaceous accretionary complexes in the east and of Cretaceous clastic basin sediments deposited on a Jurassic basement in the west (Ueda, 2016), both containing Triassic limestones. The origin of this belt is still matter of debate especially because of its western part which is not in continuity with any other accretionary complex known in the other islands of Japan and also due to the lack of data in this region. One of the main goals of this study is to

  14. Triassic rejuvenation of unexposed Archean-Paleoproterozoic deep crust beneath the western Cathaysia block, South China

    Science.gov (United States)

    Li, Xi-Yao; Zheng, Jian-Ping; Xiong, Qing; Zhou, Xiang; Xiang, Lu

    2018-01-01

    Jurassic (ca. 150 Ma) Daoxian basalts from the western Cathaysia block (South China) entrained a suite of deep-seated crustal xenoliths, including felsic schist, gneiss and granulite, and mafic two-pyroxene granulite and metagabbro. Zircon U-Pb-Hf isotopic, whole-rock elemental and Sr-Nd-Pb isotopic compositions have been determined for these valuable xenoliths to reveal the poorly-known, unexposed deep crust beneath South China. Detrital zircons from the garnet-biotite schists show several populations of ages at 0.65-0.5 Ga, 1.1-0.75 Ga, 1.6-1.4 Ga, 1.8-1.7 Ga, 2.5-2.4 Ga, 2.8 Ga, and 3.5 Ga, representing a multi-sourced, meta-sedimentary origin with deposition time at the early Cambrian. One mafic granulite contains zircons with concordant U-Pb ages of Neoarchean ( 2520 Ma), as well as Hf model ages of 2.8-2.6 Ga and positive εHf(t) values (up to 6.3), suggesting an accretion of juvenile crust in Neoarchean, probably as the main framework of the lower crust. Geochemical and geochronological evidence shows the mafic granulite and metagabbro were produced by underplating of magmas derived from the depleted asthenosphere and mixed with EM2-type materials during the Late Triassic (205-196 Ma). This magmatic underplating also resulted in the widespread metamorphism of the mafic lower crust and felsic middle crust (e.g., the felsic granulite and gneiss) at 202-201 Ma. We suggest the existence of a highly evolved Archean-Paleoproterozoic basement beneath the western Cathaysia block, which experienced episodic accretion and reworking and the strong rejuvenation during the Triassic. A three-layered structure of the lower crust could exist beneath the Daoxian area during the Jurassic time: its upper layer is an evolved Archean-Paleoproterozoic basement; the middle hybrid layer represents a mixture of Archean-Paleoproterozoic basement with newly accreted/reworked Proterozoic to Phanerozoic materials; and the deeper layer consists of mafic granulites derived from the

  15. Organic geochemical characterization of terrestrial source rocks of the Triassic Madygen formation (Southern Tien Shan, Kyrgyzstan)

    Energy Technology Data Exchange (ETDEWEB)

    Berner, U.; Scheeder, G.; Kus, J. [Section Geochemistry of Petroleum and Coal, BGR, Hannover (Germany); Voigt, S.; Schneider, J.W. [Geological Inst., TU Bergakademic Freiberg (Germany)

    2009-09-15

    Along the northern foothills of the Turkestan-Alai Range (SW Kyrgyzstan), a 1000 to 1500m thick succession of Mesozoic deposits is exposed recording regional changes of the paleo-landscape during Triassic to Cretaceous times. Detailed litho- and biofacies analyses, conducted by the TU Bergakademie Freiberg since 2006, provided for the first time a nearly complete columnar section of the continental Triassic Madygen Formation of Kyrgyzstan. Organic petrographical and organic geochemical methods (including RockEval pyrolyses, and biomarker analyses) have been applied to a suite of terrestrial sedimentary rocks of Triassic age with the intention to identify the depositional environment. Our investigations suggest that the potential source rocks of the terrestrial pluvial Madygen Formation might generate predominantly gaseous hydrocarbons at higher maturities. (orig.)

  16. A new Triassic shortening-extrusion tectonic model for Central-EasternAsia: Structural, geochronological and paleomagnetic investigations in the Xilamulun Fault (North China)

    Science.gov (United States)

    Zhao, Pan; Faure, Michel; Chen, Yan; Xu, Bei

    2017-04-01

    At the northern margin of the North China Block (NCB), the Xilamulun Fault (XMF) is a key belt to decipher the tectonic evolution of Central-Eastern Asia, as it records the Paleozoic final closure of the Paleo-Asian Ocean, and localizes a Late Triassic intracontinental deformation. In this study, structural analysis, 40Ar-39Ar dating, and paleomagnetic studies were performed to investigate the kinematics of the XMF and to further discuss its Triassic geodynamic significance in the Central-Eastern Asia framework after the Paleozoic Central Asian Orogenic evolution. The structural analyses reveal two phases of ductile deformation. The first one (D1), which displays N-verging and E-W trending folds, is related to the Early Paleozoic collisional event between the NCB and the Songliao-Hunshandake Block (SHB). The second phase (D2) displays a high-angle foliation and a pervasive sub-horizontalE-W stretching lineation with kinematic criteria indicative of dextral strike-slip shearing. The 40Ar-39Ar dating on mylonitic granite places the main shearing event around 227-209 Ma. This D2 shearing is coeval with that of the dextral strike-slip Bayan Obo-Chifeng Fault (BCF) and the Chicheng-Fengning-Longhua Fault to the south, which together constitute a dextral shearing fault system on the northern margin of the NCB during the Late Triassic. The paleomagnetic study performed on the Middle Permian Guangxingyuan pluton, located between the XMF and BCF, documents a local clockwise rotation of this pluton with respect to the NCB and SHB. Our multidisciplinary study suggests anNNW-SSE shortening and strike-slip shearing dominated tectonic setting on the northern margin of the NCB during the Late Triassic. Combining the contemporaneous dextral strike-slip movements of the XMF and BCF in northern China and the sinistral strike-slip movement of East Gobi Fault (EGF) in southeastern Mongolia with the large-scale tectonic framework, a Late Triassic NNW-SSE shortening-eastward extrusion

  17. Integrated multi-stratigraphic study of the Coll de Terrers late Permian-Early Triassic continental succession from the Catalan Pyrenees (NE Iberian Peninsula): A geologic reference record for equatorial Pangaea

    Science.gov (United States)

    Mujal, Eudald; Fortuny, Josep; Pérez-Cano, Jordi; Dinarès-Turell, Jaume; Ibáñez-Insa, Jordi; Oms, Oriol; Vila, Isabel; Bolet, Arnau; Anadón, Pere

    2017-12-01

    The most severe biotic crisis on Earth history occurred during the Permian-Triassic (PT) transition around 252 Ma. Whereas in the marine realm such extinction event is well-constrained, in terrestrial settings it is still poorly known, mainly due to the lack of suitable complete sections. This is utterly the case along the Western Tethys region, located at Pangaea's equator, where terrestrial successions are typically build-up of red beds often characterised by a significant erosive gap at the base of the Triassic strata. Henceforth, documenting potentially complete terrestrial successions along the PT transition becomes fundamental. Here, we document the exceptional Coll de Terrers area from the Catalan Pyrenees (NE Iberian Peninsula), for which a multidisciplinary research is conducted along the PT transition. The red-bed succession, located in a long E-W extended narrow rift system known as Pyrenean Basin, resulted from a continuous sedimentary deposition evolving from meandering (lower Upper Red Unit) to playa-lake/ephemeral lacustrine (upper Upper Red Unit) and again to meandering settings (Buntsandstein facies). Sedimentary continuity is suggested by preliminary cyclostratigraphic analysis that warrants further analysis. Our combined sedimentological, mineralogical and geochemical data infer a humid-semiarid-humid climatic trend across the studied succession. The uppermost Permian strata, deposited under an orbitally controlled monsoonal regime, yields a relatively diverse ichnoassemblage mainly composed of tetrapod footprints and arthropod trace fossils. Such fossils indicate appropriate life conditions and water presence in levels that also display desiccation structures. These levels alternate with barren intervals formed under dry conditions, being thus indicative of strong seasonality. All these features are correlated with those reported elsewhere in Gondwana and Laurasia, and suggest that the Permian-Triassic boundary might be recorded somewhere around

  18. Permian-Triassic Tethyan realm reorganization: Implications for the outward Pangea margin

    Science.gov (United States)

    Riel, Nicolas; Jaillard, Etienne; Martelat, Jean-Emmanuel; Guillot, Stéphane; Braun, Jean

    2018-01-01

    We present a new conceptual model to explain the first order Permian-Triassic evolution of the whole > 30 000 km long Pangea margin facing the Panthalassa ocean. Compilation of available geological, geochemical, geochronogical and paleomagnetic data all along this system allowed us to distinguish three part of the margin: western Laurentia, western Gondwana and eastern Gondwana. These segments record distinct tectonic and magmatic events, which all occur synchronously along the whole margin and correlate well with the main geodynamic events of this period, i.e. subduction of the Paleotethys mid-ocean ridge at 310-280 Ma, opening of the Neotethys at 280-260 Ma, counterclockwise rotation of Pangea at 260-230 Ma and closure of the Paleotethys at 230-220 Ma. Between 260 and 230 Ma, the reorganization of the Tethyan realm triggered the up to 35° rotation of Pangea around an Euler pole located in northernmost South America. This implied both an increase and a decrease of the convergence rate between the margin and the Panthalassa ocean, north and south of the Euler pole, respectively. Thus, the Permian-Triassic Pangean margin was marked: in western Laurentia by marginal sea closure, in western Gondwana by widespread bimodal magmatic and volcanic activity, in eastern Gondwana by transpressive orogenic phase. Therefore, we propose that the Permian-Triassic evolution of the outward margin of Pangea was controlled by the Tethyan realm reorganization.

  19. Prominent occurrence of iron oxides at KTB mass extinction: a review

    African Journals Online (AJOL)

    Out of the five major mass extinction, which have taken place in the history of life, Cretaceous–Tertiary boundary (KTB) mass extinction was the 2nd most disastrous, the most severe being that at Permian–Triassic boundary (PTB). On the basis of iridium anomaly at a number of KTB sites it has been established that the ...

  20. Provenance of Permian-Triassic Gondwana Sequence Units Accreted to the Banda Arc: Constraints from U/Pb and Hf Analysis of Zircons and Igneous Geochemistry

    Science.gov (United States)

    Flores, J. A.; Spencer, C. J.; Harris, R. A.; Hoiland, C.

    2011-12-01

    Analysis of zircons from Australian affinity Permo-Triassic units of the Timor region yield age distributions with large peaks at 230-400 Ma and 1750-1900 Ma (n=435). Similar zircon age peaks are also found in rocks from NE Australia and the eastern Cimmerian block. It is likely that these terranes, which are now widely separated, were once part of the northern edge of Gondwana near what is now the NW margin of Australia. The Cimmerian Block was removed from Gondwana during Early Permian rifting and initiation of the Neo-Tethys Ocean. Hf analysis of zircon from the Aileu Complex in Timor and Kisar shows bimodal (juvenial and evolved) magmatism in the Gondwana Sequence of NW Australia at ~300 Ma. The magmatic event produced basalt with rift valley and ocean floor geochemical affinities, and rhyolite. Similar rock types and isotopic signatures are also found in Permo-Triassic igneous units throughout the Cimmerian continental block. The part of the Cimmerian Block with zircon distributions most like the Gondwana Sequence of NW Australia is the terranes of northern Tibet and Malaysia. The large 1750-1900 Ma zircon peak is much more wide spread, and appears in terranes from Baoshan (SW China) to Borneo. The Permo-Triassic rocks of the Timor region fill syn-rift intracratonic basins that successfully rifted in the Jurassic to form the NW margin of Australia. This passive continental margin first entered the Sunda Trench in the Timor region at around 8 Ma causing the Permo-Triassic rocks to accrete to the edge of the Asian Plate and emerge as a series of mountainous islands in the young collision zone. Eventually, the Australian continental margin will collide with the southern edge of the Asian plate and these Gondwana terranes will rejoin. However, it may be difficult to reconstruct the various ventures of they made over the past 300 Ma.

  1. Global taxonomic diversity of anomodonts (tetrapoda, therapsida and the terrestrial rock record across the Permian-Triassic boundary.

    Directory of Open Access Journals (Sweden)

    Jörg Fröbisch

    Full Text Available The end-Permian biotic crisis (~252.5 Ma represents the most severe extinction event in Earth's history. This paper investigates diversity patterns in Anomodontia, an extinct group of therapsid synapsids ('mammal-like reptiles', through time and in particular across this event. As herbivores and the dominant terrestrial tetrapods of their time, anomodonts play a central role in assessing the impact of the end-Permian extinction on terrestrial ecosystems. Taxonomic diversity analysis reveals that anomodonts experienced three distinct phases of diversification interrupted by the same number of extinctions, i.e. an end-Guadalupian, an end-Permian, and a mid-Triassic extinction. A positive correlation between the number of taxa and the number of formations per time interval shows that anomodont diversity is biased by the Permian-Triassic terrestrial rock record. Normalized diversity curves indicate that anomodont richness continuously declines from the Middle Permian to the Late Triassic, but also reveals all three extinction events. Taxonomic rates (origination and extinction indicate that the end-Guadalupian and end-Permian extinctions were driven by increased rates of extinction as well as low origination rates. However, this pattern is not evident at the final decline of anomodont diversity during the Middle Triassic. Therefore, it remains unclear whether the Middle Triassic extinction represents a gradual or abrupt event that is unique to anomodonts or more common among terrestrial tetrapods. The end-Permian extinction represents the most distinct event in terms of decline in anomodont richness and turnover rates.

  2. Understanding Late Triassic low latitude terrestrial ecosystems: new insights from the Colorado Plateau Coring Project (CPCP)

    Science.gov (United States)

    Irmis, R. B.; Olsen, P. E.; Parker, W.; Rasmussen, C.; Mundil, R.; Whiteside, J. H.

    2017-12-01

    The Chinle Formation of southwestern North America is a key paleontological archive of low paleolatitude non-marine ecosystems that existed during the Late Triassic hothouse world. These strata were deposited at 5-15°N latitude, and preserve extensive plant, invertebrate, and vertebrate fossil assemblages, including early dinosaurs; these organisms lived in an unpredictably fluctuating semi-arid to arid environment with very high atmospheric pCO2. Despite this well-studied fossil record, a full understanding of these ecosystems and their integration with other fossil assemblages globally has been hindered by a poor understanding of the Chinle Formation's age, duration, and sedimentation rates. Recently, the CPCP recovered a 520m continuous core through this formation from the northern portion of Petrified Forest National Park (PEFO) in northern Arizona, USA. This core has provided a plethora of new radioisotopic and magnetostratigraphic data from fresh, unweathered samples in unambiguous stratigraphic superposition. These constraints confirm that virtually all fossil-bearing horizons in Chinle outcrops in the vicinity of PEFO are Norian in age. Furthermore, they indicate that the palynomorph zone II and Adamanian vertebrate biozone are at least six million years long, whereas the overlying palynomorph zone III and Revueltian vertebrate biozone persisted for at least five million years, with the boundary between 216-214 Ma. This confirms that the rich late Adamanian-early Revueltian vertebrate fossil assemblages, where dinosaurs are exclusively rare, small-bodied carnivorous theropods, are contemporaneous with higher latitude assemblages in Europe, South America, and Africa where large-bodied herbivorous sauropodomorph dinosaurs are common. The age constraints also confirm that several palynomorph biostratigraphic ranges in the Chinle Formation differ from those of the same taxa in eastern North American (Newark Supergroup) and Europe. These data are consistent

  3. Arsenic, barium, strontium and uranium geochemistry and their utility as tracers to characterize groundwaters from the Espadán–Calderona Triassic Domain, Spain

    Energy Technology Data Exchange (ETDEWEB)

    Giménez-Forcada, Elena, E-mail: e.gimenez@igme.es [Instituto Geológico y Minero de España — IGME, Unidad de Salamanca, Azafranal 48, 37001 Salamanca (Spain); Vega-Alegre, Marisol [University of Valladolid — UVA, Department of Analytical Chemistry, Campus Miguel Delibes, Paseo Belén 7, 47011 Valladolid (Spain)

    2015-04-15

    A set of analytical data from the Espadán–Calderona Triassic Domain aquifers was processed using hierarchical agglomerative cluster analysis (HCA) and principal component analysis (PCA), to achieve a quantitative and independent approach to investigate the characteristics of groundwater composition and possible differences between groundwater flows from Triassic aquifers from the Espadán–Calderona Triassic Domain (Spain). Mineralization in the Triassic series has led to the presence of several metals and metalloids in groundwater, including As, Mn, Fe and U. These are associated with fresher bicarbonate groundwaters, characterized by lower Sr/Ba ratios. Levels containing sulfate evaporitic salt, which are interbedded through the Triassic series, seem to exert a strong influence on the chemistry of several groundwaters, characterized by calcium sulfate facies with high Sr concentration and high Sr/Ba ratios. The application of multivariate statistical techniques to the interpretation of analytical results allows the differentiation of groundwater types occurring in the Triassic aquifers and identification of the role of a number of minor or trace elements and their ratios that can be treated as hydrogeochemical tracers. With them it was possible to correlate the different recharge waters with the tectonic morphology of the Espadán–Calderona Triassic Domain. - Highlights: • Groundwater hydrochemistry of Espadán–Calderona Triassic Domain was investigated. • Multivariate statistical analysis and metal tracers differentiated groundwater types. • The Sr/Ba ratio and As and U concentrations are good indicators of groundwater flows. • The chemistry of groundwater flows correlates with structural morphology of recharge areas.

  4. Arsenic, barium, strontium and uranium geochemistry and their utility as tracers to characterize groundwaters from the Espadán–Calderona Triassic Domain, Spain

    International Nuclear Information System (INIS)

    Giménez-Forcada, Elena; Vega-Alegre, Marisol

    2015-01-01

    A set of analytical data from the Espadán–Calderona Triassic Domain aquifers was processed using hierarchical agglomerative cluster analysis (HCA) and principal component analysis (PCA), to achieve a quantitative and independent approach to investigate the characteristics of groundwater composition and possible differences between groundwater flows from Triassic aquifers from the Espadán–Calderona Triassic Domain (Spain). Mineralization in the Triassic series has led to the presence of several metals and metalloids in groundwater, including As, Mn, Fe and U. These are associated with fresher bicarbonate groundwaters, characterized by lower Sr/Ba ratios. Levels containing sulfate evaporitic salt, which are interbedded through the Triassic series, seem to exert a strong influence on the chemistry of several groundwaters, characterized by calcium sulfate facies with high Sr concentration and high Sr/Ba ratios. The application of multivariate statistical techniques to the interpretation of analytical results allows the differentiation of groundwater types occurring in the Triassic aquifers and identification of the role of a number of minor or trace elements and their ratios that can be treated as hydrogeochemical tracers. With them it was possible to correlate the different recharge waters with the tectonic morphology of the Espadán–Calderona Triassic Domain. - Highlights: • Groundwater hydrochemistry of Espadán–Calderona Triassic Domain was investigated. • Multivariate statistical analysis and metal tracers differentiated groundwater types. • The Sr/Ba ratio and As and U concentrations are good indicators of groundwater flows. • The chemistry of groundwater flows correlates with structural morphology of recharge areas

  5. Moessbauer effect study of the diagenesis on the southern Brazilian Triassic paleoherpetofauna

    International Nuclear Information System (INIS)

    Da Costa, M.I. Jr.; Holz, M.; Schultz, C.L.

    1994-01-01

    Moessbauer spectroscopy is applied to identify the iron contents of bony elements of southern Brazilian Triassic reptile remains, and the question of the paragenetic mineral assemblage is discussed. (orig.)

  6. Prelude of benthic community collapse during the end-Permian mass extinction in siliciclastic offshore sub-basin: Brachiopod evidence from South China

    Science.gov (United States)

    Wu, Huiting; He, Weihong; Weldon, Elizabeth A.

    2018-04-01

    Analysis of the Permian-Triassic palaeocommunities from basinal facies in South China provides an insight into the environmental deterioration occurring in the prelude to the mass extinction event. Quantitative and multivariate analyses on three brachiopod palaeocommunities from the Changhsingian to the earliest Triassic in basinal facies in South China have been undertaken in this study. Although the end-Permian extinction has been proved to be a one-stepped event, ecological warning signals appeared in the palaeocommunities long before the main pulse of the event. A brachiopod palaeocommunity turnover occurred in the upper part of the Clarkina changxingensis Zone, associated with a significant decrease of palaeocommunity diversity and brachiopod body size. During this turnover the dominant genera changed from Fusichonetes and Crurithyris (or/and Paracrurithyris) to the more competitive genus Crurithyris (or/and Paracrurithyris). The brachiopod palaeocommunity turnover was supposed to be triggered by the decreased marine primary productivity and increased volcanic activity. Moreover, such early warning signals are found not only in the deep-water siliceous facies, but also in the shallow-water clastic facies and carbonate rock facies in South China.

  7. Depositional environments and cyclo- and chronostratigraphy of uppermost Carboniferous-Lower Triassic -lacustrine deposits, southern Bogda Mountains, NW China - A terrestrfluvialial paleoclimatic record of mid-latitude NE Pangea

    Science.gov (United States)

    Yang, W.; Feng, Q.; Liu, Yajing; Tabor, N.; Miggins, D.; Crowley, J.L.; Lin, J.; Thomas, S.

    2010-01-01

    Two uppermost Carboniferous–Lower Triassic fluvial–lacustrine sections in the Tarlong–Taodonggou half-graben, southern Bogda Mountains, NW China, comprise a 1834 m-thick, relatively complete sedimentary and paleoclimatic record of the east coast of mid-latitude NE Pangea. Depositional environmental interpretations identified three orders (high, intermediate, and low) of sedimentary cycles. High-order cycles (HCs) have five basic types, including fluvial cycles recording repetitive changes of erosion and deposition and lacustrine cycles recording repetitive environmental changes associated with lake expansion and contraction. HCs are grouped into intermediate-order cycles (ICs) on the basis of systematic changes of thickness, type, and component lithofacies of HCs. Nine low-order cycles (LCs) are demarcated by graben-wide surfaces across which significant long-term environmental changes occurred. A preliminary cyclostratigraphic framework provides a foundation for future studies of terrestrial climate, tectonics, and paleontology in mid-latitude NE Pangea.Climate variabilities at the intra-HC, HC, IC, and LC scales were interpreted from sedimentary and paleosol evidence. Four prominent climatic shifts are present: 1) from the humid–subhumid to highly-variable subhumid–semiarid conditions at the beginning of Sakamarian; 2) from highly-variable subhumid–semiarid to humid–subhumid conditions across the Artinskian-Capitanian unconformity; 3) from humid–subhumid to highly-variable subhumid–semiarid conditions at early Induan; and 4) from the highly-variable subhumid–semiarid to humid–subhumid conditions across the Olenekian-Anisian unconformity. The stable humid–subhumid condition from Lopingian to early Induan implies that paleoclimate change may not have been the cause of the end-Permian terrestrial mass extinction. A close documentation of the pace and timing of the extinction and exploration of other causes are needed. In addition, the

  8. ON THE SIZE AND COMOVING MASS DENSITY EVOLUTION OF EARLY-TYPE GALAXIES

    International Nuclear Information System (INIS)

    Van der Wel, Arjen; Bell, Eric F.; Van den Bosch, Frank C.; Gallazzi, Anna; Rix, Hans-Walter

    2009-01-01

    We present a simple, empirically motivated model that simultaneously predicts the evolution of the mean size and the comoving mass density of massive (>10 11 M sun ) early-type galaxies from z = 2 to the present. First, we demonstrate that some size evolution of the population can be expected simply due to the continuous emergence of early-type galaxies. The Sloan Digital Sky Survey (SDSS) data reveal that in the present-day universe more compact early-type galaxies with a given dynamical mass have older stellar populations. This implies that with increasing look-back time, the more extended galaxies will be more and more absent from the population. In contrast, at a given stellar velocity dispersion, SDSS data show that there is no relation between size and age, which implies that the velocity dispersion can be used to estimate the epoch at which galaxies stopped forming stars, turning into early-type galaxies. Based on this, we define an empirically motivated, redshift-dependent velocity dispersion threshold above which galaxies do not form stars at a significant rate, which we associate with the transformation into early-type galaxies. Applying this 'formation' criterion to a large sample of nearby early-type galaxies, we predict the redshift evolution in the size distribution and the comoving mass density. The resulting evolution in the mean size is roughly half of the observed evolution. Then we include a prescription for the merger histories of galaxies between the 'formation' redshift and the present, based on cosmological simulations of the assembly of dark matter halos. Such mergers after the transformation into an early-type galaxy are presumably dissipationless ('dry'), where the increase in size is expected to be approximately proportional to the increase in mass. This model successfully reproduces the observed evolution since z ∼ 2 in the mean size and in the comoving mass density of early-type galaxies with mass M > 10 11 M sun . We conclude that

  9. A new theropod dinosaur from India with remarks on the Gondwana-Laurasia connection in the Late Triassic

    Science.gov (United States)

    Chatterjee, S.

    Walkeria maleriensis (n. g., n. sp.) from the Late Triassic Maleri Formation of the Godavari Valley of India is the earliest known dinosaur from Asia. It is a small podokesaurid theropod, very similar to Procompsognathus of Germany, Coelophysis of North America, and Syntarsus from Zimbabwe and North America. The podokesaurs are of particular interest to students of organic evolution because they are the earliest known theropods from which Archaeopteryx, the oldest known fossil bird, was probably evolved. Traditionally, India has been regarded as a part of Gondwana. It is generally believed that Gondwana remained an integral geographic unit throughout the Triassic. If this is so, a strong faunal correlation between India and other Gondwana continents should be expected in Late Triassic time. Contrary to this, the Maleri fauna is overwhelmingly "northern". Walkeria occurs in association with metoposaurs, parasuchids, protorosaurs, aetosaurs, rhynchosaurs, and traversodonts. Most of these taxa have been identified in the Dockum fauna of North America, indicating a close paleontologic link between India and Laurasia. Possibly the route of faunal migration between India and North America during the Late Triassic was via northern Africa.

  10. Disentangling Diagenesis From the Rock Record: An Example From the Permo-Triassic Wordie Creek Formation, East Greenland

    Science.gov (United States)

    Roberts, J.; Turchyn, A. V.; Wignall, P. B.; Newton, R. J.; Vane, C. H.

    2018-01-01

    The measurement of isotope ratios in sedimentary rocks deposited over geological time can provide key insights to past environmental change over important intervals in the past. However, it is important to be aware that secondary alteration can overprint the original isotopic records. We demonstrate this principle using high-resolution carbon, sulfur, and oxygen isotope measurements in organic carbon, pyrite, and carbonate minerals (δ13Corg, δ34Spyr, δ34SCAS, δ13Ccarb, and δ18Ocarb) and kerogen analyses (HI and OI) from the Wordie Creek Formation, East Greenland. These sediments were initially deposited across the Permo-Triassic transition, but as we will show, the carbonate record has been altered by interaction with meteoric water significantly after initial deposition. Comparison of the better preserved organic carbon and pyrite records with a proximal Permo-Triassic sequence reveals significant pyrite-sulfur isotope variability across the Permo-Triassic transition. This regional heterogeneity argues against basin-wide euxinia and instead suggests localized changes in sulfur fractionation in response to variations in organic carbon flux. This hypothesis can be used to explain seemingly inconsistent regional trends in other sulfur isotopes across the Permo-Triassic transition.

  11. Conodont succession and reassessment of major events around the Permian-Triassic boundary at the Selong Xishan section, southern Tibet, China

    Science.gov (United States)

    Yuan, Dong-Xun; Zhang, Yi-Chun; Shen, Shu-Zhong

    2018-02-01

    A major discrepancy for the age of the Selong Group from middle Cisuralian (Early Permian) to Changhsingian resulted from previous reports of Sakmarian, Kungurian and Guadalupian (Middle Permian) conodonts and Lopingian (Late Permian) brachiopods. Recently, Cisuralian and Guadalupian conodonts were reported again from the Selong Group and the basal part of the Kangshare Formation at the Selong section, but the age discrepancy remains. We present our conodont materials based on large samples collected from the Selong Group and our interpretation based on identifications using a sample population approach. Three conodont zones are recognized in our re-investigation of the upper part of the Selong Group. They include the Vjalovognathus sp., the Mesogondolella hendersoni, and the M. sheni zones, in ascending order. These zones are overlain by the basal Triassic Hindeodus parvus Zone and the Otoceras woodwardi Zone. Our reassessment of conodonts reported by previous studies from Selong and nearby sections suggest that all specimens consistently point to a Lopingian age; the upper part of the Selong Group is latest Changhsingian in age based on the presence of Clarkina orchardi and Mesogondolella sheni. Previously reported early Cisuralian and Guadalupian conodonts are misidentified using a form species concept. A hiatus may be present at the erosional surface between the Selong Group and the Waagenites Bed of the basal part of the Kangshare Formation. However, the hiatus is minimal because conodont and brachiopod assemblages above and below this surface are very similar, and it results from a latest Changhsingian transgression just before the extinction that follows a global latest Changhsingian regression. There is a distinct rapid end-Permian mass extinction at Selong within the Waagenites Bed, as indicated by the disappearances of all benthic brachiopods, rugose corals and Permian bryozoans. The burst of Clarkina species in the Waagenites Bed and throughout the

  12. Triassic volcanic units in coastal region of Antofagasta, northern Chile

    International Nuclear Information System (INIS)

    Basso, M.; Cortes, J.A.; Marinovic, N

    2001-01-01

    U-Pb geochronological evidence of a Middle to Late Triassic volcanic event was found in the coastal region of Antofagasta, northern Chile (23 o -23 o 30 ). Two new ages were obtained from rhyolitic tuffs and an associated dome, which have classically been attributed to the Jurassic La Negra Formation (au)

  13. Spectroscopic Studies on Organic Matter from Triassic Reptile Bones, Upper Silesia, Poland

    Science.gov (United States)

    Surmik, Dawid; Boczarowski, Andrzej; Balin, Katarzyna; Dulski, Mateusz; Szade, Jacek; Kremer, Barbara; Pawlicki, Roman

    2016-01-01

    Fossil biomolecules from an endogenous source were previously identified in Cretaceous to Pleistocene fossilized bones, the evidence coming from molecular analyses. These findings, however, were called into question and an alternative hypothesis of the invasion of the bone by bacterial biofilm was proposed. Herewith we report a new finding of morphologically preserved blood-vessel-like structures enclosing organic molecules preserved in iron-oxide-mineralized vessel walls from the cortical region of nothosaurid and tanystropheid (aquatic and terrestrial diapsid reptiles) bones. These findings are from the Early/Middle Triassic boundary (Upper Roetian/Lowermost Muschelkalk) strata of Upper Silesia, Poland. Multiple spectroscopic analyses (FTIR, ToF-SIMS, and XPS) of the extracted "blood vessels" showed the presence of organic compounds, including fragments of various amino acids such as hydroxyproline and hydroxylysine as well as amides, that may suggest the presence of collagen protein residues. Because these amino acids are absent from most proteins other than collagen, we infer that the proteinaceous molecules may originate from endogenous collagen. The preservation of molecular signals of proteins within the "blood vessels" was most likely made possible through the process of early diagenetic iron oxide mineralization. This discovery provides the oldest evidence of in situ preservation of complex organic molecules in vertebrate remains in a marine environment. PMID:26977600

  14. Spectroscopic Studies on Organic Matter from Triassic Reptile Bones, Upper Silesia, Poland.

    Directory of Open Access Journals (Sweden)

    Dawid Surmik

    Full Text Available Fossil biomolecules from an endogenous source were previously identified in Cretaceous to Pleistocene fossilized bones, the evidence coming from molecular analyses. These findings, however, were called into question and an alternative hypothesis of the invasion of the bone by bacterial biofilm was proposed. Herewith we report a new finding of morphologically preserved blood-vessel-like structures enclosing organic molecules preserved in iron-oxide-mineralized vessel walls from the cortical region of nothosaurid and tanystropheid (aquatic and terrestrial diapsid reptiles bones. These findings are from the Early/Middle Triassic boundary (Upper Roetian/Lowermost Muschelkalk strata of Upper Silesia, Poland. Multiple spectroscopic analyses (FTIR, ToF-SIMS, and XPS of the extracted "blood vessels" showed the presence of organic compounds, including fragments of various amino acids such as hydroxyproline and hydroxylysine as well as amides, that may suggest the presence of collagen protein residues. Because these amino acids are absent from most proteins other than collagen, we infer that the proteinaceous molecules may originate from endogenous collagen. The preservation of molecular signals of proteins within the "blood vessels" was most likely made possible through the process of early diagenetic iron oxide mineralization. This discovery provides the oldest evidence of in situ preservation of complex organic molecules in vertebrate remains in a marine environment.

  15. Integrated geophysical study of the Triassic salt bodies' geometry and evolution in central Tunisia

    Science.gov (United States)

    Azaiez, Hajer; Amri, Dorra Tanfous; Gabtni, Hakim; Bedir, Mourad; Soussi, Mohamed

    2008-01-01

    A comprehensive study, integrating gravity, magnetic and seismic reflection data, has been used to resolve the complex Triassic salt body geometry and evolution in central Tunisia. Regional seismic lines across the study area show a detachment level in the Upper Triassic evaporites, associated with chaotic seismic facies below the Souinia, Majoura, and Mezzouna structures. The Jurassic and Lower Cretaceous seismic horizons display pinching-outs and onlapping around these structures. A stack-velocity section confirms the existence of a high-velocity body beneath the Souinia Mountain. Regional gravity and magnetic profiles in this area were elaborated from ETAP (the Tunisian Firm of Petroleum Activities) measure stations. These profiles were plotted following the same layout from the west (Souinia) to the east (Mezzouna), across the Majoura and Kharrouba mountains. They highlight associated gravity and magnetic negative anomalies. These gravity and magnetic data coupled to the reflection seismic data demonstrate that, in the Souinia, Majoura, and El Hafey zones, the Triassic salt reaches a salt pillow and a salt-dome stage, without piercing the cover. These stages are expressed by moderately low gravity anomalies. On the other hand, in the Mezzouna area (part of the North-South Axis), the Triassic salt had pierced its cover during the Upper Cretaceous and the Tertiary, reaching a more advanced stage as a salt diapir and salt wall. These stages express important low gravity and magnetic anomalies. These results confirm the model of Tanfous et al. (2005) of halokinetic movements by fault intrusions inducing, from the west to the east, structures at different stages of salt pillow, salt dome, and salt diapir.

  16. THE TRIASSIC/JURASSIC BOUNDARY IN THE ANDES OF ARGENTINA

    Directory of Open Access Journals (Sweden)

    ALBERTO C. RICCARDI

    2004-03-01

    Full Text Available The Arroyo Malo Formation at Alumbre Creek, on the northern bank of the Atuel River, west central Argentina, comprises a c. 300 m thick continuous marine succession across the Triassic-Jurassic System boundary, consisting of massive and laminated pelites indicative of a slope depositional environment. Late Triassic invertebrates, including ammonoids, nautiloids, bivalves, gastropods, brachiopods and corals are restricted to the lower 150 m. Beds between 125-135 m from the bottom yield Choristoceras cf. marshi Hauer, a species found in the Marshi/Crickmayi Zone of Europe and North America, together with loose fragments of Psiloceras cf. pressum Hillebrandt, coeval with the lower to middle part of the Hettangian Planorbis Zone. About 80 m higher are beds yielding Psiloceras cf. rectocostatum Hillebrandt, a species that gives name to an Andean biozone partially coeval with the Johnstoni and Plicatulum Subzones, upper Planorbis Zone. Other fossils recorded in the Rhaetian strata of this section are foraminifers, ostracods and plant remains identified as Zuberia cf. zuberi (Szaj. Freng. and Clathropteris sp. The section was also sampled for conodonts and radiolarians, thus far with negative results. A palaeomagnetic study is underway.

  17. Geomorphological stability of Permo-Triassic albitized profiles - case study of the Montseny-Guilleries High (NE Iberia)

    Science.gov (United States)

    Parcerisa, D.; Casas, L.; Franke, C.; Gomez-Gras, D.; Lacasa, G.; Nunez, J. A.; Thiry, M.

    2010-05-01

    Massif paleoalteration profiles (≥ 200 m) occur in the upper parts of the Montseny-Guilleries High (NE Catalan Coastal Ranges). The profiles consist of hard albitized-chloritized-hematized facies in the lower part and softer kaolinized-hematized facies in the upper part of the section. Preliminary paleomagnetic data show Triassic ages for both, the albitized and the kaolinized parts, and point to a surficial formation altered under oxidising conditions. Similar paleoalteration profiles have already been described and dated to Triassic ages elsewhere in Europe [Schmitt, 1992; Ricordel et al., 2007; Parcerisa et al., 2009]. These Permian-Triassic alterations are following a succession of different mineral transformations from the top to the base of the profile: 1) Red facies are defined by an increase in the amount and size of haematite crystals leading to the red colour of the rocks. The increase on haematite content is pervasively affecting the whole rock and is accompanied by the kaolinitization of the feldspars. 2) Pink facies: here, the granite shows an uniform pink colouration, which is mainly due to the albitization of the primary Ca-bearing plagioclases, accompanied by a precipitation of minute haematite, sericite, and calcite crystals inside the albite. Additionally primary biotite is fully chloritized. The pink granites are much more resistant to the present-day weathering than the "unaltered" facies at the base of the profile. 3) Spotted facies is characterized by a partial alteration of the rock, which caused a pink-screened aspect to the rock. The alteration developed along the fractures and is less well developed or absent in the non-fractured zones. In the pink-screened facies, the plagioclases are partially albitized and contain numerous hematite inclusions. Biotites are usually almost entirely chloritized. 4) Unaltered facies: These granites are coloured white to greyish, containing plagioclase and K-feldspar that do not show any trace of

  18. Palaeomagnetism of the Early Permian Mount Leyshon Intrusive Complex and Tuckers Igneous Complex, North Queensland, Australia

    Science.gov (United States)

    Clark, D. A.; Lackie, M. A.

    2003-06-01

    This study provides reliable, precisely defined and well-dated Early Permian (286 +/- 6 Ma) palaeomagnetic poles for Australia from the Mount Leyshon Intrusive Complex (MLIC) and the Tuckers Igneous Complex (TIC). Both complexes are associated with prominent negative magnetic anomalies, indicating the presence of rocks carrying stable remanence of reverse polarity, with a Koenigsberger ratio greater than unity. The characteristic remanence carried by the intrusive phases and by locally remagnetized, contact-metamorphosed host rocks is always of reverse polarity, consistent with acquisition during the Permo-Carboniferous (Kiaman) Reverse Superchron. The corresponding palaeopoles confirm that Australia occupied high latitudes in the Early Permian. The pole positions are: MLIC: lat. = 43.2 °S, long. = 137.3 °E dp = 6.0°, dm = 6.4° Q= 6; TIC: lat. = 47.5 °S, long. = 143.0 °E, dp = 6.0°, dm = 6.6° Q= 6. Permian palaeomagnetic overprinting is detectable at considerable distances from the MLIC (2-3 km), well beyond the zone of visible alteration. The primary nature of the Early Permian palaeomagnetic signature is established by full baked contact/aureole tests at both localities. Other new data from Australia are consistent with the poles reported here. Comparison of the Australian, African and South American Apparent Polar Wander Paths (APWP) suggests that mean Permian and Triassic poles from West Gondwana, particularly from South America, are biased by remagnetization in the Jurassic-Cretaceous and that the Late Palaeozoic-Mesozoic APWP for Gondwana is best defined by Australian data. The Australian APWP exhibits substantial movement through the Mesozoic. Provided only that the time-averaged palaeofield was zonal, the Early Triassic palaeomagnetic data from Australia provide an important palaeogeographic constraint that the south geographic pole was within, or very close to, SE Australia around 240 Ma. The new Early Permian poles are apparently more consistent

  19. Mercury anomalies as a proxy for large igneous province volicanism and effects on the carbon cycle in a U-Pb age-constrained section spanning the end-Triassic mass extinction, Levanto, Peru

    Science.gov (United States)

    Yager, J. A.; West, A. J.; Bergquist, B. A.; Thibodeau, A. M.; Corsetti, F. A.; Berelson, W.; Rosas, S.; Bottjer, D. J.

    2017-12-01

    Understanding the causes of the end-Triassic extinction and their potential relationship to Central Atlantic Magmatic Province (CAMP) volcanism necessitates careful correlation of carbon cycle records (largely from marine sections) and volcanism (largely from terrestrial successions) in a robust chronological framework. Here, we report stable carbon isotopes and mercury concentrations and isotopes from the Levanto section in Northern Peru as a putative proxy for CAMP (a large igneous province) in a marine section. Levanto represents deposition well below storm wave base and is lithologically homogenous before, during, and after the end-Triassic extinction interval, making it ideal for detailed chemostratigraphy. Furthermore, abundant intercalated ash beds allow us to correlate mercury concentrations in the marine record directly with CAMP basalt ages, providing a test of the correspondence of mercury anomalies with the eruption of CAMP volcanics. Age dating and C isotope analyses provide an opportunity to explore orbital tuning of the carbon isotope record and ground truth it with existing U-Pb ages from the section, a feature not available in any other marine sections examined to date. The abundance of U-Pb dated ashes in the Levanto section allows us to correlate orbital tuning with other basins, which lack absolute age control, providing a better understanding for the C cycle changes associated with the Triassic-Jurassic boundary.

  20. Utilizing borehole electrical images to interpret lithofacies of fan-delta: A case study of Lower Triassic Baikouquan Formation in Mahu Depression, Junggar Basin, China

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    Yuan Rui

    2017-11-01

    Full Text Available Large-scale conglomerate fan-delta aprons were typical deposits on the slope of Mahu Depression during the Early Triassic. Without outcrops, it is difficult to study the lithofacies only by examining the limited cores from the main oil-bearing interval of the Baikouquan Formation. Borehole electrical imaging log provides abundant high-resolution geologic information that is obtainable only from real rocks previously. Referring to the lithology and sedimentary structure of cores, a case study of fan-deltas in the Lower Triassic Baikouquan Formation of the Mahu Depression presents a methodology for interpreting the complicated lithofacies utilizing borehole electrical images. Eleven types of lithologies and five types of sedimentary structures are summarized in borehole electrical images. The sediments are fining upward from gravel to silt and clay in the Baikouquan Formation. Fine-pebbles and granules are the main deposits in T1b1 and T1b2, but sandstones, siltstones and mudstones are more developed in T1b3. The main sedimentary textures are massive beddings, cross beddings and scour-and-fill structures. Parallel and horizontal beddings are more developed in T1b3 relatively. On integrated analysis of the lithology and sedimentary structure, eight lithofacies from electrical images, referred to as image lithofacies, is established for the fan-deltas. Granules to coarse-pebbles within massive beddings, granules to coarse-pebbles within cross and parallel beddings, siltstones within horizontal and massive beddings are the most developed lithofacies respectively in T1b1, T1b2 and T1b3. It indicates a gradual rise of the lake level of Mahu depression during the Early Triassic, with the fan-delta aprons retrograding towards to the margin of the basin. Therefore, the borehole electrical imaging log compensate for the limitation of cores of the Baikouquan Formation, providing an effective new approach to interpret the lithofacies of fan-delta.

  1. Utilizing borehole electrical images to interpret lithofacies of fan-delta: A case study of Lower Triassic Baikouquan Formation in Mahu Depression, Junggar Basin, China

    Science.gov (United States)

    Yuan, Rui; Zhang, Changmin; Tang, Yong; Qu, Jianhua; Guo, Xudong; Sun, Yuqiu; Zhu, Rui; Zhou, Yuanquan (Nancy)

    2017-11-01

    Large-scale conglomerate fan-delta aprons were typical deposits on the slope of Mahu Depression during the Early Triassic. Without outcrops, it is difficult to study the lithofacies only by examining the limited cores from the main oil-bearing interval of the Baikouquan Formation. Borehole electrical imaging log provides abundant high-resolution geologic information that is obtainable only from real rocks previously. Referring to the lithology and sedimentary structure of cores, a case study of fan-deltas in the Lower Triassic Baikouquan Formation of the Mahu Depression presents a methodology for interpreting the complicated lithofacies utilizing borehole electrical images. Eleven types of lithologies and five types of sedimentary structures are summarized in borehole electrical images. The sediments are fining upward from gravel to silt and clay in the Baikouquan Formation. Fine-pebbles and granules are the main deposits in T1b1 and T1b2, but sandstones, siltstones and mudstones are more developed in T1b3. The main sedimentary textures are massive beddings, cross beddings and scour-and-fill structures. Parallel and horizontal beddings are more developed in T1b3 relatively. On integrated analysis of the lithology and sedimentary structure, eight lithofacies from electrical images, referred to as image lithofacies, is established for the fan-deltas. Granules to coarse-pebbles within massive beddings, granules to coarse-pebbles within cross and parallel beddings, siltstones within horizontal and massive beddings are the most developed lithofacies respectively in T1b1, T1b2 and T1b3. It indicates a gradual rise of the lake level of Mahu depression during the Early Triassic, with the fan-delta aprons retrograding towards to the margin of the basin. Therefore, the borehole electrical imaging log compensate for the limitation of cores of the Baikouquan Formation, providing an effective new approach to interpret the lithofacies of fan-delta.

  2. Development of lower Triassic wrinkle structures: implications for the search for life on other planets.

    Science.gov (United States)

    Mata, Scott A; Bottjer, David J

    2009-11-01

    Wrinkle structures are microbially mediated sedimentary structures that are a common feature of Proterozoic and earliest Phanerozoic siliciclastic seafloors on Earth and occur only rarely in post-Cambrian strata. These macroscopic microbially induced sedimentary structures are readily identifiable at the outcrop scale, and their recognition on other planetary bodies by landed missions may suggest the presence of past microbial life. Wrinkle structures of the Lower Triassic (Spathian) Virgin Limestone Member of the Moenkopi Formation in the western United States record an occurrence of widespread microbialite formation in the wake of the end-Permian mass extinction, the largest biotic crisis of the Phanerozoic. Wrinkle structures occur on proximal sandy tempestites deposited within the offshore transition. Storm layers appear to have been rapidly colonized by microbial mats and were subsequently buried by mud during fair-weather conditions. Wrinkle structures exhibit flat-topped crests and sinuous troughs, with associated mica grains oriented parallel to bedding, suggestive of trapping and binding activity. Although Lower Triassic wrinkle structures postdate the widespread occurrence of these features during the Proterozoic and Cambrian, they exhibit many of the same characteristics and environmental trends, which suggests a conservation of microbial formational and preservational processes in subtidal siliciclastic settings on Earth from the Precambrian into the Phanerozoic. In the search for extraterrestrial life, it may be these conservative characteristics that prove to be the most useful and robust for recognizing microbial features on other planetary bodies, and may add to an ever-growing foundation of knowledge for directing future explorations aimed at seeking out macroscopic microbial signatures.

  3. MIDDLE TRIASSIC FORAMINIFERA FROM THE SECEDA CORE (DOLOMITES, NORTHERN ITALY

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    FLORIAN MAURER

    2002-11-01

    Full Text Available The assemblage of foraminifera in turbidite beds in Middle Triassic basinal deposits straddling the Anisian/Ladinian boundary interval was studied in 224 thin sections. The fauna consists mainly of shallow-water inhabitants, associated with calcareous hyaline foraminifera (Lagenina of open marine environment. Due to a well established  biostratigraphy in the studied interval, the first and last appearance of some shallow water, benthic foraminifera can be assigned to the Mid Triassic ammonoid stratigraphy. The species Meandrospira dinarica Kochansky-Devidè & Pantic and Arenovidalina chialingchiagensis Ho are limited to the Reitzi ammonoid zone. The species Variostoma alta Kristan and Hoyenella gr. sinensis both do not superate the Curionii zone in age in the studied succession.  The biostratigraphic most important event occurs at the base of the Gredleri zone,  with the appearance of the family Involutinidae Bütschli, represented by the genera Lamelliconus and Aulotortus. The faunal composition is similar to those of neighbouring paleoprovinces, but generally a lower faunistical diversification compared to foraminiferal assemblages in the Anisian or Carnian is observed.   

  4. Comparison of mantle lithosphere beneath early Triassic kimberlite fields in Siberian craton reconstructed from deep-seated xenocrysts

    Directory of Open Access Journals (Sweden)

    I.V. Ashchepkov

    2016-07-01

    Kharamai mantle clinopyroxenes represent three geochemical types: (1 harzburgitic with inclined linear REE, HFSE troughs and elevated Th, U; (2 lherzolitic or pyroxenitic with round TRE patterns and decreasing incompatible elements; (3 eclogitic with Eu troughs, Pb peak and high LILE content. Calculated parental melts for garnets with humped REE patterns suggest dissolution of former Cpx and depression means Cpx and garnets extraction. Clinopyroxenes from Ary-Mastakh fields show less inclined REE patterns with HMREE troughs and an increase of incompatible elements. Clinopyroxenes from Kuranakh field show flatter spoon-like REE patterns and peaks in Ba, U, Pb and Sr, similar to those in ophiolitic harzburgites. The PT diagrams for the mantle sections show high temperature gradients in the uppermost SCLM accompanied by an increase of P-Fe#Ol upward and slightly reduced thickness of the mantle keel of the Siberian craton, resulting from the influence of the Permian–Triassic superplume, but with no signs of delamination.

  5. Paleomagnetic and magnetostratigraphic investigations of the whitehorse group/quartermaster (Dewey Lake) formation (upper permian-lowermost triassic) in the Palo Duro basin, northwest Texas, USA

    Science.gov (United States)

    Collins, Dylan R.

    In northwest Texas, upper Permian to lowermost Triassic hematite-cemented detrital sedimentary rocks, which include a small number of regionally extensive ash beds, were deposited during the time interval of the greatest mass extinction event sequences in Earth history. The magnetic polarity stratigraphy, as well as key rock magnetic properties, of the upper Whitehorse Group (WH) and Quartermaster formations (QM) at selected sections in the Palo Duro Basin, have been determined using thermal, and chemical demagnetization approaches and anisotropy of magnetic susceptibility, acquisition of isothermal remanent magnetization (IRM) and backfield demagnetization, and thermal demagnetization of three component IRM methods. Demagnetization results show that the WH/QM contains a primary/near-primary characteristic remanent magnetization at each level sampled and thus the magnetic polarity stratigraphy for each section can be compared with existing polarity time scales across the Permian-Triassic boundary. Estimated site mean directions yield a paleomagnetic pole for the latest Permian for North America of 57.8°N, 130.6°E from 38 sampled sites.

  6. Patterns of fossil distributions within their environmental context from the Middle Triassic in South Canyon, Central Nevada, USA

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    Pedro M. Monarrez

    2014-01-01

    Full Text Available The Middle Triassic records the return of diverse marine communities after the severe effects of the end-Permian mass extinction. This diversification leads to the Mesozoic/modern adaptive radiation resulting in substantial changes in marine communities in comparison to their Paleozoic predecessors. This analysis focuses on the faunal abundance, ecological patterns, and environmental interpretation of a Middle Triassic section in Central Nevada. Twelve bulk samples were collected. Visible fossils were identified and tallied from hand samples and thin-sections were used to aid in environmental interpretation. Beginning in the Late Anisian, we observed an ammonoid dominated to flat-clam, epifaunal dominated benthic community within a muddy, quiet, inner shelf depositional environment. Through time, epifaunal bivalves dominate within a middle shelf environment followed by an increase in infaunalization and shell-thickness. During this time the presence of oncoids and the reported finding of corals suggest the middle shelf environment gave way to a higher energy patch reef shelf edge environment. Finally, we observe epifaunal brachiopods communities at the top of our section deposited in a middle shelf environment. In sum, we observe the dominance of modern taxa (i.e., bivalves with Paleozoic ecologies (i.e., epifaunal, followed by the dominance of modern taxa with Modern ecologies (i.e., infaunal, thick shells and then a return to Paleozoic taxa (i.e., brachiopods and Paleozoic ecologies within an overall transgressive environment.

  7. Early crocodylomorph increases top tier predator diversity during rise of dinosaurs.

    Science.gov (United States)

    Zanno, Lindsay E; Drymala, Susan; Nesbitt, Sterling J; Schneider, Vincent P

    2015-03-19

    Triassic predatory guild evolution reflects a period of ecological flux spurred by the catastrophic end-Permian mass extinction and terminating with the global ecological dominance of dinosaurs in the early Jurassic. In responding to this dynamic ecospace, terrestrial predator diversity attained new levels, prompting unique trophic webs with a seeming overabundance of carnivorous taxa and the evolution of entirely new predatory clades. Key among these was Crocodylomorpha, the largest living reptiles and only one of two archosaurian lineages that survive to the present day. In contrast to their existing role as top, semi-aquatic predators, the earliest crocodylomorphs were generally small-bodied, terrestrial faunivores, occupying subsidiary (meso) predator roles. Here we describe Carnufex carolinensis a new, unexpectedly large-bodied taxon with a slender and ornamented skull from the Carnian Pekin Formation (~231 Ma), representing one of the oldest and earliest diverging crocodylomorphs described to date. Carnufex bridges a problematic gap in the early evolution of pseudosuchians by spanning key transitions in bauplan evolution and body mass near the origin of Crocodylomorpha. With a skull length of >50 cm, the new taxon documents a rare instance of crocodylomorphs ascending to top-tier predator guilds in the equatorial regions of Pangea prior to the dominance of dinosaurs.

  8. Palaeogeographic evolution of the marine Middle Triassic marine Germanic Basin changements - With emphasis on the carbonate tidal flat and shallow marine habitats of reptiles in Central Pangaea

    Science.gov (United States)

    Diedrich, Cajus G.

    2009-01-01

    began to diversify at exactly this time, and it seems likely that this was a direct reaction to these environmental changes. It can be inferred that the earliest dinosaurs must have started to evolve in the late Early Triassic, because in Europe it can be demonstrated that at least two main dinosaur groups already were present and clearly differentiated by the middle part of the Middle Triassic, and all three of the major groups of dinosaurs (theropods, sauropods and ornithischians) had diversified and spread globally throughout terrestrial habitats by the end of the Triassic Period. Six new palaeogeographic maps, representing time intervals from the Aegean to the Illyrian (Anisian) stages, show these important environmental changes in detail and explain the direction and timing of terrestrial reptile exchanges between the Central Massif, Rhenish Massif, and Bohemian Massif, and also the direction and timing of marine reptile exchanges between the Alps of Central Pangaea and the ancient northern Tethys Ocean and Germanic Basin Sea.

  9. Spondarthritis in the triassic.

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    Juan Carlos Cisneros

    Full Text Available BACKGROUND: The evidence of several forms of arthritis has been well documented in the fossil record. However, for pre-Cenozoic vertebrates, especially regarding reptiles, this record is rather scarce. In this work we present a case report of spondarthritis found in a vertebral series that belonged to a carnivorous archosaurian reptile from the Lower Triassic (∼245 million years old of the South African Karoo. METHODOLOGY/PRINCIPAL FINDINGS: Neutron tomography confirmed macroscopic data, revealing the ossification of the entire intervertebral disc space (both annulus fibrosus and nucleus pulposus, which supports the diagnosis of spondarthritis. CONCLUSIONS/SIGNIFICANCE: The presence of spondarthritis in the new specimen represents by far the earliest evidence of any form of arthritis in the fossil record. The present find is nearly 100 million years older than the previous oldest report of this pathology, based on a Late Jurassic dinosaur. Spondarthritis may have indirectly contributed to the death of the animal under study.

  10. N and C Isotopic Compositions of the Lower Triassic of Southern Primorye and Reconstruction of Habitat Conditions of Marine Organisms after Mass Extinction at the End of the Permian

    Science.gov (United States)

    Zakharov, Y. D.; Horacek, M.; Shigeta, Y.; Popov, A. M.; Maekawa, T.

    2018-02-01

    Data on the N and C isotopic composition are presented for the Lower Triassic claystones of the Abrek section of southern Primorye (Far East). The results showed five N isotopic intervals and several negative C isotopic excursions of the Induan-lower Olenekian stages of the Abrek section.

  11. Investigation of mineral composition of differently treated devonian, quaternary and triassic clays of Latvia

    International Nuclear Information System (INIS)

    Kosorukovs, A.; Viss, R.

    1999-01-01

    Clayey fractions (particle size less than 5 μm )of the Latvian Devonian (Kuprava and Liepa deposits), Quaternary (Laza and Ugale deposits) and Triassic (Akmene deposit, Republic of Lithuania) clays have been obtained. The clayey fractions were converted in the form in which all the cations were exchanged for magnesium ions. After the ion exchange the fractions were treated with dimethyl sulfoxide or glycerol in the course for two days, one sample being subjected to thermal treatment at 550±110 C for two hours. Diffractograms for the treated samples have been obtained using a DRON-2,0 diffractometer (Co-radiation). Analysis of the obtained diffractograms show that: 1) the main clayey minerals of the Devonian clays occur to be hydromicas (mainly hydromuscovite) containing admixtures of kaolinite and quartz; 2) the main clayey minerals of the Quarternary clays also occur to be hydromicas - mixtures of hydrobiotite and hydromuscovite containing admixtures of kaolinite and iron-containing chlorite; 3) smectite occurs to be the main mineral of the Triassic clay; it contains admixtures of hydromica and chlorite; 4) the Triassic and Quaternary clays contain fine- and coarse-grained carbonates, mainly calcite, in quantities of 10-16%; 5) iron and titanium are included in fine grained minerals. (author)

  12. A hyper-robust sauropodomorph dinosaur ilium from the Upper Triassic-Lower Jurassic Elliot Formation of South Africa: Implications for the functional diversity of basal Sauropodomorpha

    Science.gov (United States)

    McPhee, Blair W.; Choiniere, Jonah N.

    2016-11-01

    It has generally been held that the locomotory habits of sauropodomorph dinosaurs moved in a relatively linear evolutionary progression from bipedal through "semi-bipedal" to the fully quadrupedal gait of Sauropoda. However, there is now a growing appreciation of the range of locomotory strategies practiced amongst contemporaneous taxa of the latest Triassic and earliest Jurassic. Here we present on the anatomy of a hyper-robust basal sauropodomorph ilium from the Late Triassic-Early Jurassic Elliot Formation of South Africa. This element, in addition to highlighting the unexpected range of bauplan diversity throughout basal Sauropodomorpha, also has implications for our understanding of the relevance of "robusticity" to sauropodomorph evolution beyond generalized limb scaling relationships. Possibly representing a unique form of hindlimb stabilization during phases of bipedal locomotion, the autapomorphic morphology of this newly rediscovered ilium provides additional insight into the myriad ways in which basal Sauropodomorpha managed the inherited behavioural and biomechanical challenges of increasing body-size, hyper-herbivory, and a forelimb primarily adapted for use in a bipedal context.

  13. Osteology of Carnufex carolinensis (Archosauria: Psuedosuchia from the Pekin Formation of North Carolina and Its Implications for Early Crocodylomorph Evolution.

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    Susan M Drymala

    Full Text Available Crocodylomorphs originated in the Late Triassic and were the only crocodile-line archosaurs to survive the end-Triassic extinction. Recent phylogenetic analyses suggest that the closest relatives of these generally gracile, small-bodied taxa were a group of robust, large-bodied predators known as rauisuchids implying a problematic morphological gap between early crocodylomorphs and their closest relatives. Here we provide a detailed osteological description of the recently named early diverging crocodylomorph Carnufex carolinensis from the Upper Triassic Pekin Formation of North Carolina and assess its phylogenetic position within the Paracrocodylomorpha. Carnufex displays a mosaic of crocodylomorph, rauisuchid, and dinosaurian characters, as well as highly laminar cranial elements and vertebrae, ornamented dermal skull bones, a large, subtriangular antorbital fenestra, and a reduced forelimb. A phylogenetic analysis utilizing a comprehensive dataset of early paracrocodylomorphs and including seven new characters and numerous modifications to characters culled from the literature recovers Carnufex carolinensis as one of the most basal members of Crocodylomorpha, in a polytomy with two other large bodied taxa (CM 73372 and Redondavenator. The analysis also resulted in increased resolution within Crocodylomorpha and a monophyletic clade containing the holotype and two referred specimens of Hesperosuchus as well as Dromicosuchus. Carnufex occupies a key transition at the origin of Crocodylomorpha, indicating that the morphology typifying early crocodylomorphs appeared before the shift to small body size.

  14. U-Th-Geochemistry of Permian and Triassic sediments of the Drauzug, Carinthia, Austria

    International Nuclear Information System (INIS)

    Kurat, G.; Korkisch, J.; Niedermayr, G.; Seemann, R.

    1976-05-01

    Chemical analysis of samples of Triassic and Permian rocks from the Drauzug, Carinthia and Austria was carried out. U concentration was measured by flurimetry, Th and Cu by spectrophotometry using Thoronol method or Arsenazo III method for Th, Fe by titrimetry and V, Ba, Sr by atomic absorption spectrophotometry. The average U concentration ranged from 0.8 to 4.6 ppm and the Th concentration from 3.2 to 15.6 ppm depending upon the mineral material. The quartzporphyries contained the highest concentration of both. It was concluded that the Permian-Triassic series are very similar to the equivalent deposits in Northern Italy. Assuming a lateral displacement, the former represent the Northern marginal part of the latter and therefore are inferior in thickness and thus not favourable for larger U mineralization

  15. Depositional environments of Late Triassic lake, east-central New Mexico

    Energy Technology Data Exchange (ETDEWEB)

    Hester, P.M. (Bureau of Land Management, Albuquerque, NM (USA))

    1989-09-01

    The Redonda Member of the Chinle Formation represents deposition in a large, polymictic lake during the Late Triassic (Norian) in east-central New Mexico. This study documents and defines an extensive lacustrine system situated in western Pangaea which was influenced by both tectonic and climatic events. Areal extent of the lake may have been as much as 5,000 km{sup 2}.

  16. Triassic regolithization: A major stage of pre-enrichment in the formation of unconformity related deposits in Southern France

    International Nuclear Information System (INIS)

    Schmitt, J.M.; Clement, J.Y.

    1989-01-01

    The formation of unconformity related uranium deposits in Canada and Australia is currently thought to have involved some stage of preconcentration within the Proterozoic regolith. Uranium deposits in the southern Massif central (France) are spatially linked to the Mesozoic unconformity. Under this unconformity, rocks of the Hercynian basement as well as Permo-Carboniferous sediments show a regolithic alteration dating back to the Late Permian to Late Triassic period. On the elevated parts of the Triassic landscape, 30 to 50 m deep weathering profiles are preserved. Three main zones can be distinguished: a lower pink coloured zone, showing partly albitized and chloritized rocks: a middle bleached zone with neogenic clays; and an upper reddish zone with Fe-Mn oxyhydroxides. Towards the Triassic basin, much deeper (200-300 m) alteration profiles are observed on Permo-Carboniferous sediments. The two upper regolithic zones are present, but the lower albitized one is very developed with two subzones: analcite/albite at the top, and K-feldspar/albite at the bottom. Geochemical data show that potassium is fixed in the uppermost horizons of the regolith, whereas sodium is transported towards the lower horizons and basin areas and fixed in analcite/albite zones. Uranium, vanadium, copper and less mobile elements such as titanium and zirconium are strongly leached from the weathering profiles in elevated parts of the landscape but are enriched in basin zones of albitization up to 5-15 times. Thus, solutions generated from the weathering profiles have brought about a major redistribution of uranium in the Triassic landscape. The remarkable applanation and tectonic stability of the area as well as a subarid climate seem to have favoured this 'regolithization' process. In southern France, geochemical differentiation during the Triassic has thus given rise to important uranium pre-enrichment, and the regolith is a major control for later uranium deposits. (author). 16 refs, 8

  17. Preliminary study of the uranium potential of the Wadesboro Triassic basin, North Carolina

    International Nuclear Information System (INIS)

    Thayer, P.A.; Harris, W.B.

    1981-11-01

    This report presents results of a four-channel spectrometric survey of the Wadesboro Triassic basin and adjacent Piedmont, North and South Carolina. A total of 216 gamma-ray spectrometric measurements was taken at 165 sites in the Wadesboro Basin and at 37 sites in the adjacent Piedmont. The normal sampling density in the Wadesboro Basin is one site per 4.5 km 2 . Surface radiometric surveys reveal no anomalous radioactivity. Estimated uranium concentrations (eU) from the area are from 0.1 to 4.9 ppM. Average eU content of Triassic sandstones is 1.5 ppM, which is nearly the average for adjacent metavolcanic rocks (1.6 ppM). Granitic intrusives of the Pageland and Lilesville plutons display the highest eU concentrations in the area, averaging 3.3 ppM. The uranium/thorium ratio is consistently low throughout the area, from 0.01 to 0.54; the log uranium/log thorium ratio is from -1.11 to a maximum of 0.66. Triassic sandstones have the highest values, averaging 0.25. Granites of the Pageland and Lilesville plutons have an average uranium/thorium ratio of 0.21, and Slate Belt metavolcanic rocks average 0.15. On the basis of surface radiometric surveys and geologic studies, it is believed that sedimentary strata of the Wadesboro Basin are poor targets for further uranium exploration. This conclusion is based on the lack of many favorable characteristics associated with fluvial uranium deposits

  18. Stratigraphic distribution, taphonomy and paleoenvironments of Spinicaudata in the Triassic and Jurassic of the Paraná Basin

    Science.gov (United States)

    Jenisch, Alan Gregory; Lehn, Ilana; Gallego, Oscar Florencio; Monferran, Mateo Daniel; Horodyski, Rodrigo Scalise; Faccini, Ubiratan Ferrucio

    2017-12-01

    exoskeletons is likely a function of parameters, e.g., the transport duration, the distance from life position, and the magnitude of events causing their final burial. Within the observed species, the recognition of Eustheria minuta in the stratigraphic level of the Passo das Tropas creek corroborates an age for these deposits between the late Middle Triassic and early Upper Triassic. The presence of a new form, likely related to the family Fushunograptidae in sediments from the Caturrita Formation, suggests a Jurassic age for these deposits.

  19. An integrated perspective on the Permian-Triassic "Great Dying"

    Science.gov (United States)

    Algeo, T. J.

    2017-12-01

    The 252-Ma end-Permian mass extinction (EPME), marked by the loss of 90% of marine invertebrate species, was the largest biocrisis in Earth history. Intensive study of this "Great Dying" has led to major insights and a broad consensus regarding many aspects of this event. The ultimate trigger is regarded as eruption of the Siberian Traps Large Igneous Province (STLIP), which released large quantities of greenhouse gases (CO2 and CH4) and sulfate aerosols, triggering a catastrophic global warming of 10°C and acidification of both land surfaces and the surface ocean. On land, a massive die-off of vegetation led to a transient episode of rapid soil erosion and a longer-term increase in weathering rates linked to elevated temperatures. In the ocean, widespread anoxia developed concurrently with the EPME, triggered by ocean-surface warming that reduced dissolved oxygen solubility in seawater and that intensified vertical stratification. Expanded anoxia led to massive burial of organic matter and reduced sulfur, although the evidence for this is indirect (C, U and S isotopes); few organic-rich deposits of Early Triassic age have been found, suggesting that organic sedimentation occurred mainly on continental slopes or in the deep ocean. Other aspects of the end-Permian crisis remain under debate. For example, there is no consensus regarding changes in marine productivity levels in the aftermath of the EPME, which would have been stimulated by enhanced subaerial weathering but depressed by reduced overturning circulation-the evidence to date may favor localized positive and negative changes in productivity. Also under scrutiny is evidence for volcanic eruptions and environmental perturbations during the 100 kyr prior to the EPME, which are likely to have occurred but remain poorly dated and quantified. The greatest uncertainty, however, may surround the nature of the proximate kill mechanism(s) during the EPME. Many hypotheses have been advanced including mechanisms

  20. UPPER OLENEKIAN (SPATHIAN AMMONOIDS FROM CHIOS (LOWER TRIASSIC, GREECE: TAXONOMY AND STRATIGRAPHIC POSITIONS

    Directory of Open Access Journals (Sweden)

    DOROTHEE MERTMANN

    2003-11-01

    Full Text Available  On the Greek island of Chios, Lower Triassic sections bave delivered rich ammonoid faunas from red limestones of the Hallstatt facies. Here, we present a palaeontological description of Spathian ammonoids, which have been collected during two field campaigns by Italian-German teams (Assereto et al. 1980; Gaetani et al. 1992. The ammonoid associations are composed of Ceratitida and Phylloceratida, each order containing species of the Noritaceae, Hedenstroemiaceae, Xenodiscaceae, Dinaritaceae, Pinacocerataceae, Phyllocerataceae, Ptychitaceae, Ussuritaceae and Megaphyllitaceae. Long ranging genera, e.g. Procarnites and Leiophyllites, are present as well as endemic ones, e.g. Chiotites, which occurs exclusively in the Lower Triassic of Chios. The ammonoid association is indicative of the Prohungarites-Subcolumbites zone sensu Kummel (1973a. The haugi zone, installed as uppermost Olenekian in North America, is not represented by ammonoids on Chios. 

  1. Preliminary study of the uranium potential of the northern part of the Durham Triassic Basin, North Carolina

    International Nuclear Information System (INIS)

    Harris, W.B.; Thayer, P.A.

    1981-09-01

    This report presents results of a four-channel spectrometric survey of the northern part of the Durham Triassic basin and adjacent Piedmont, North Carolina. Gamma-ray spectrometric measurements were obtained at 112 localities from 136 different lithologies. The nominal sampling density in the Durham Basin is one site per 2 mi 2 . Surface radiometric surveys reveal no anomalous radioactivity in the northern part of the Durham Basin. Uranium concentrations in Triassic rocks are from 0.6 to 9.7 ppM and average 2.9 ppM. Mudrocks contain from 1.3 to 9.7 ppM, and the average is 4.5 ppM. Sandstones contain from 0.6 to 8.8 ppM, and the average is 2.5 ppM. Fanglomerates contain the lowest concentrations of uranium, from 1.4 to 2.0 ppM, for an average of 1.8 ppM. Uranium/thorium ratios average 0.27 for Triassic rocks and are from 0.04 to 1.85. The mean log uranium/log thorium for Triassic rocks is 0.37. Mudrock has the highest average uranium/thorium ratio (0.32), and the range is 0.09 to 0.66. Sandstones have an average uranium/thorium ratio of 0.26, and the range is 0.04 to 1.85. Fanglomerates have the lowest range uranium/thorium ratio (0.19), and the range is 0.12 to 0.19. On the basis of surface radiometric surveys and geologic studies, it is believed that sedimentary strata in the northern part of the Durham Basin are poor targets for further uranium exploration. This conclusion is based on the lack of favorable characteristics commonly present in fluvial uranium deposits. Among these are: (1) carbonaceous material is absent in Triassic rocks of the northern basin, (2) indicators of a reduzate facies in sandstones are not present, and (3) no tuffaceous beds are associated with sediments in the northern Durham Basin

  2. An ancestral turtle from the Late Triassic of southwestern China.

    Science.gov (United States)

    Li, Chun; Wu, Xiao-Chun; Rieppel, Olivier; Wang, Li-Ting; Zhao, Li-Jun

    2008-11-27

    The origin of the turtle body plan remains one of the great mysteries of reptile evolution. The anatomy of turtles is highly derived, which renders it difficult to establish the relationships of turtles with other groups of reptiles. The oldest known turtle, Proganochelys from the Late Triassic period of Germany, has a fully formed shell and offers no clue as to its origin. Here we describe a new 220-million-year-old turtle from China, somewhat older than Proganochelys, that documents an intermediate step in the evolution of the shell and associated structures. A ventral plastron is fully developed, but the dorsal carapace consists of neural plates only. The dorsal ribs are expanded, and osteoderms are absent. The new species shows that the plastron evolved before the carapace and that the first step of carapace formation is the ossification of the neural plates coupled with a broadening of the ribs. This corresponds to early embryonic stages of carapace formation in extant turtles, and shows that the turtle shell is not derived from a fusion of osteoderms. Phylogenetic analysis places the new species basal to all known turtles, fossil and extant. The marine deposits that yielded the fossils indicate that this primitive turtle inhabited marginal areas of the sea or river deltas.

  3. Ostracods (Crustacea associated with microbialites across the Permian-Triassic boundary in Dajiang (Guizhou Province, South China

    Directory of Open Access Journals (Sweden)

    Marie-Beatrice FOREL

    2012-08-01

    Full Text Available 26 samples were processed for a taxonomic study of ostracods from the Upper Permian (Changhsingian - Lower Triassic (Griesbachian interval of the Dajiang section, Guizhou Province, South China. 112 species belonging to 27 genera are recognized. Five new species are described: Acratia candyae sp. nov, Bairdia adelineae sp. nov., Bairdia? huberti sp. nov., Bairdia jeromei sp. nov., Orthobairdia jeanlouisi sp. nov. The unexpected survival faunas associated with microbial formations in the aftermath of the end-Permian extinction are documented for the first time. Ostracod biodiversity variations and palaeo-environmental modifications associated with microbial growth through the Permian-Triassic boundary (PTB are discussed.

  4. Rapid enhancement of chemical weathering recorded by extremely light seawater lithium isotopes at the Permian-Triassic boundary

    Science.gov (United States)

    Sun, He; Xiao, Yilin; Gao, Yongjun; Zhang, Guijie; Casey, John F.; Shen, Yanan

    2018-04-01

    Lithium (Li) isotope analyses of sedimentary rocks from the Meishan section in South China reveal extremely light seawater Li isotopic signatures at the Permian–Triassic boundary (PTB), which coincide with the most severe mass extinction in the history of animal life. Using a dynamic seawater lithium box model, we show that the light seawater Li isotopic signatures can be best explained by a significant influx of riverine [Li] with light δ7Li to the ocean realm. The seawater Li isotope excursion started ≥300 Ky before and persisted up to the main extinction event, which is consistent with the eruption time of the Siberian Traps. The eruption of the Siberian Traps exposed an enormous amount of fresh basalt and triggered CO2 release, rapid global warming, and acid rains, which in turn led to a rapid enhancement of continental weathering. The enhanced continental weathering delivered excessive nutrients to the oceans that could lead to marine eutrophication, anoxia, acidification, and ecological perturbation, ultimately resulting in the end-Permian mass extinction.

  5. Large-Diameter Burrows of the Triassic Ischigualasto Basin, NW Argentina: Paleoecological and Paleoenvironmental Implications

    Science.gov (United States)

    Colombi, Carina E.; Fernández, Eliana; Currie, Brian S.; Alcober, Oscar A.; Martínez, Ricardo; Correa, Gustavo

    2012-01-01

    Large-diameter ichnofossils comprising three morphotypes have been identified in the Upper Triassic Ischigualasto and Los Colorados formations of northwestern Argentina. These burrows add to the global record of the early appearance of fossorial behavior during early Mesozoic time. Morphotypes 1 and 2 are characterized by a network of tunnels and shafts that can be assigned to tetrapod burrows given similarities with previously described forms. However, differences in diameter, overall morphology, and stratigraphic occurrence allow their independent classification. Morphotype 3 forms a complex network of straight branches that intersect at oblique angles. Their calcareous composition and surface morphology indicate these structures have a composite biogenic origin likely developed due to combined plant/animal interactions. The association of Morphotypes 1 and 2 with fluvial overbank lithologies deposited under an extremely seasonal arid climate confirms interpretations that the early appearance of burrowing behavior was employed by vertebrates in response to both temperature and moisture-stress associated with seasonally or perpetually dry Pangean paleoclimates. Comparisons of burrow morphology and biomechanical attributes of the abundant paleovertebrate fauna preserved in both formations permit interpretations regarding the possible burrow architects for Morphotypes 1 and 2. In the case of the Morphotype 1, the burrow constructor could be one of the small carnivorous cynodonts, Ecteninion or Probelesodon. Assigning an architect for Morphotype 2 is more problematic due to mismatches between the observed burrow morphology and the size of the known Los Colorados vertebrates. PMID:23227195

  6. The geology and hydrocarbon possibilities of the Triassic-Jurassic Fundy Basin, eastern Canada

    Energy Technology Data Exchange (ETDEWEB)

    Wade, J.A.; Fensome, R.A. [Geological Survey of Canada, Dartmouth, NS (Canada). Atlantic Geoscience Centre; Brown, D.E. [Canada-Nova Scotia Offshore Petroleum Board, Halifax, NS (Canada)

    1997-09-01

    The development of the Mesozoic sedimentary basins beneath the waters of the eastern coast of North America was discussed. These basins have been linked to the rifting of the central part of Pangaea during Mid and Late Triassic time that ended in the formation of a series of grabens extending from Florida to The Grand Banks of Newfoundland, one of them being the Bay of Fundy Basin which is about 16,500 square kilometres in size. Onshore and offshore geologic mapping and seismic interpretations have shown their age range to be from the Mid Triassic Anisian or Ladinian to Mid Jurassic. Up to 12 km of Mesozoic rocks were deposited in the basin with up to 9 km still present. The depositional history of the area was described. The two areas with greatest hydrocarbon potential are the Bay of Fundy and the Chignecto subbasins.

  7. Paleomagnetism and Magnetostratigraphy of Upper Permian to Lower Triassic (?) Beaufort Group Strata at Bethulie, Karoo Basin, Free State Province, South Africa

    Science.gov (United States)

    Geissman, J. W.; Gastaldo, R. A.; Neveling, J.; Makubalo, S.

    2017-12-01

    A multifaceted approach to understand the timing of interpreted environmental changes during the Late Permian to possibly Early Triassic (?) time in the Beaufort Group strata of the Karoo Basin includes work to establish robust magnetic polarity records for sections previously interpreted to encompass end-Permian extinction events. Demonstrating the preservation of an early-acquired remanence (RM) in Karoo strata is required for a robust magnetostratigraphy. Yet, this is challenging due to thermochemical effects related to the Early Jurassic (ca. 183 Ma) Karoo Large Igneous Province (LIP), and the NE to SW increase in burial diagenesis attending Cape Fold Belt tectonism. Documentation of a primary RM in these strata, which appears to be preserved in some areas, requires careful laboratory- and field-based assessment. We report data from 53 sites collected at the well-studied Bethulie section, Free State Province, in which several samples per individual horizon to assess ChRM uniformity. Strata well-removed from dikes yield both normal and reverse polarity ChRM. It is always the case that the first-removed RM is a NNW seeking, moderate to steep negative-inclination ChRM (normal polarity); NRM intensities are typically 1 to 5 mA/m. Sites BT15 and BT21, which are located in strata lying some 4 m below the often-cited "event bed" interval inferred to be the terrestrial expression of the Permian/Triassic boundary, is dominated by a well-defined reverse RM with a normal overprint RM unblocked below 400oC, implying elevated temperatures (i.e., 100 to 250oC+) for ca. 1 Ma (+/-). Contact tests are positive but complicated. Host strata collected in distances equal to or less than 1-2 dike widths from the intrusions have been thermally remagnetized and demonstrate high NRM intensities (> 50 mA/m). Collectively, we interpret these data to indicate that any ChRM, with the exception of those from host strata in baked contacts, that persists above Tlub of 425oC is pre-Karoo LIP

  8. Mass loss in early stages of stellar evolution

    International Nuclear Information System (INIS)

    Dearborn, D.S.P.; Kozlowski, M.; Schramm, D.

    1976-01-01

    It is known that stars lose mass between their birth on the main sequence and their death as white dwarfs, or as neutron stars or black holes for more massive stars. Solar wind considerations indicate that not all of the mass loss occurs in a 'last gasp' to form a planetary nebula. Mass loss has be observed in O-B stars and in red giants, but the mass loss observed in these stages do not appear sufficient to account for all the loss required for stars to become white dwarfs. The problem is at what stages and in what manner the excess mass is lost. Suggestions have tended to favour He flashes for stars with M < 2.25 M. and possibly double shell instability flashes for stars in the range from 2.25 M. to 8 M. Although is is possible that significant mass is lost during these stages, there appears to be no quantitative support for this suggestion, and in particular none for the He flash. It is shown here that there is increasing evidence for substantial mass loss during the immediate post main sequence stages when the star is moving from the main sequence to the giant branch. Mass loss at these early stages of evolution may have implications on the subsequent evolution of the star and the amount of nuclearly processed material that is ejected into the interstellar-medium. The behaviour of Arcturus is discussed in these connections. (U.K.)

  9. Detection of Enhanced Central Mass-to-light Ratios in Low-mass Early-type Galaxies: Evidence for Black Holes?

    Science.gov (United States)

    Pechetti, Renuka; Seth, Anil; Cappellari, Michele; McDermid, Richard; den Brok, Mark; Mieske, Steffen; Strader, Jay

    2017-11-01

    We present dynamical measurements of the central mass-to-light ratio (M/L) of a sample of 27 low-mass early-type {{ATLAS}}3{{D}} galaxies. We consider all {{ATLAS}}3{{D}} galaxies with 9.7 text{}}M/L{{s}} are higher than dynamical {\\text{}}M/L{{s}} derived at larger radii and stellar population estimates of the galaxy centers in ˜80% of galaxies, with a median enhancement of ˜14% and a statistical significance of 3.3σ. We show that the enhancement in the central M/L is best described either by the presence of black holes in these galaxies or by radial initial mass function variations. Assuming a black hole model, we derive black hole masses for the sample of galaxies. In two galaxies, NGC 4458 and NGC 4660, the data suggest significantly overmassive black holes, while in most others only upper limits are obtained. We also show that the level of M/L enhancements we see in these early-type galaxy nuclei are consistent with the larger enhancements seen in ultracompact dwarf galaxies (UCDs), supporting the scenario where massive UCDs are created by stripping galaxies of these masses.

  10. The osteology of the basal archosauromorph Tasmaniosaurus triassicus from the Lower Triassic of Tasmania, Australia.

    Directory of Open Access Journals (Sweden)

    Martín D Ezcurra

    Full Text Available Proterosuchidae are the most taxonomically diverse archosauromorph reptiles sampled in the immediate aftermath of the Permo-Triassic mass extinction and represent the earliest radiation of Archosauriformes (archosaurs and closely related species. Proterosuchids are potentially represented by approximately 15 nominal species collected from South Africa, China, Russia, Australia and India, but the taxonomic content of the group is currently in a state of flux because of the poor anatomic and systematic information available for several of its putative members. Here, the putative proterosuchid Tasmaniosaurus triassicus from the Lower Triassic of Hobart, Tasmania (Australia, is redescribed. The holotype and currently only known specimen includes cranial and postcranial remains and the revision of this material sheds new light on the anatomy of the animal, including new data on the cranial endocast. Several bones are re-identified or reinterpreted, contrasting with the descriptions of previous authors. The new information provided here shows that Tasmaniosaurus closely resembles the South African proterosuchid Proterosuchus, but it differed in the presence of, for example, a slightly downturned premaxilla, a shorter anterior process of maxilla, and a diamond-shaped anterior end of interclavicle. Previous claims for the presence of gut contents in the holotype of Tasmaniosaurus are considered ambiguous. The description of the cranial endocast of Tasmaniosaurus provides for the first time information about the anatomy of this region in proterosuchids. The cranial endocast preserves possibly part of the vomero-nasal ( = Jacobson's system laterally to the olfactory bulbs. Previous claims of the absence of the vomero-nasal organs in archosaurs, which is suggested by the extant phylogenetic bracket, are questioned because its absence in both clades of extant archosaurs seems to be directly related with the independent acquisition of a non-ground living

  11. Stratigraphy and tectonics of Permo-Triassic basins in the Netherlands and surrounding areas

    NARCIS (Netherlands)

    Geluk, M.C.

    2005-01-01

    This thesis addresses different aspects of the geological development during the Permian and Triassic (300 to 200 Ma) of the Netherlands and surrounding areas. The study area encompasses the Southern Permian Basin (SPB), a large intracratonic basin stretched out from the United Kingdom in the west

  12. Triassic oils and related hydrocarbon kitchens in the Adriatic basin

    Energy Technology Data Exchange (ETDEWEB)

    Novelli, L.; Demaison, G. (AGIP, Milan (Italy))

    1988-08-01

    Without exception, the oils from both the Abruzzi basin and Albanian foredeep are of lower Liassic to Upper Triassic origin. This is demonstrated by biological marker-based correlations between the oils and stratigraphically controlled, carbonate-rich source rocks. The biomarker studies also provided proof to conclude that many of the oils possess low API gravities and high sulfur contents because they are immature rather than biodegraded. Following the geochemical investigations, a computer-aided, basinwise maturation simulation of the hydrocarbon kitchens was carried out, with backstripping in geologic time. The simulations, performed with the Tissot-Espitalie kinetic model, used basin-specific kerogen activation energies obtained by the optimum method. These simulated values were calibrated with observed values in deep wells. Two characteristics diverge from normal petroleum basin situations (e.g., the North Sea basin): sulfur-rich kerogens in the source rocks, featuring relatively low activation energy distributions, and low geothermal gradients in the subsurface. The geographic outlines of simulated Triassic-lower Liassic hydrocarbon kitchens closely coincide with the zones of petroleum occurrence and production in the Adriatic basin. Furthermore, API gravities of the oils are broadly predicted by the mathematical simulations. This methodology has once again shown its ability to rationally high-grade the petroleum-rich sectors of sedimentary basin while identifying those areas where chances of success are extremely low regardless of the presence of structures.

  13. Uranium isotope evidence for an expansion of marine anoxia during the end-Triassic extinction

    NARCIS (Netherlands)

    Jost, Adam B.; Bachan, Aviv; van de Schootbrugge, Bas; Lau, Kimberly V.; Weaver, Karrie L.; Maher, Kate; Payne, Jonathan L.

    The end-Triassic extinction coincided with an increase in marine black shale deposition and biomarkers for photic zone euxinia, suggesting that anoxia played a role in suppressing marine biodiversity. However, global changes in ocean anoxia are difficult to quantify using proxies for local anoxia.

  14. Prolonged Permian Triassic ecological crisis recorded by molluscan dominance in Late Permian offshore assemblages.

    Science.gov (United States)

    Clapham, Matthew E; Bottjer, David J

    2007-08-07

    The end-Permian mass extinction was the largest biotic crisis in the history of animal life, eliminating as many as 95% of all species and dramatically altering the ecological structure of marine communities. Although the causes of this pronounced ecosystem shift have been widely debated, the broad consensus based on inferences from global taxonomic diversity patterns suggests that the shift from abundant brachiopods to dominant molluscs was abrupt and largely driven by the catastrophic effects of the end-Permian mass extinction. Here we analyze relative abundance counts of >33,000 fossil individuals from 24 silicified Middle and Late Permian paleocommunities, documenting a substantial ecological shift to numerical dominance by molluscs in the Late Permian, before the major taxonomic shift at the end-Permian mass extinction. This ecological change was coincident with the development of fluctuating anoxic conditions in deep marine basins, suggesting that numerical dominance by more tolerant molluscs may have been driven by variably stressful environmental conditions. Recognition of substantial ecological deterioration in the Late Permian also implies that the end-Permian extinction was the climax of a protracted environmental crisis. Although the Late Permian shift to molluscan dominance was a pronounced ecological change, quantitative counts of 847 Carboniferous-Cretaceous collections from the Paleobiology Database indicate that it was only the first stage in a stepwise transition that culminated with the final shift to molluscan dominance in the Late Jurassic. Therefore, the ecological transition from brachiopods to bivalves was more protracted and complex than their simple Permian-Triassic switch in diversity.

  15. Prolonged Permian–Triassic ecological crisis recorded by molluscan dominance in Late Permian offshore assemblages

    Science.gov (United States)

    Clapham, Matthew E.; Bottjer, David J.

    2007-01-01

    The end-Permian mass extinction was the largest biotic crisis in the history of animal life, eliminating as many as 95% of all species and dramatically altering the ecological structure of marine communities. Although the causes of this pronounced ecosystem shift have been widely debated, the broad consensus based on inferences from global taxonomic diversity patterns suggests that the shift from abundant brachiopods to dominant molluscs was abrupt and largely driven by the catastrophic effects of the end-Permian mass extinction. Here we analyze relative abundance counts of >33,000 fossil individuals from 24 silicified Middle and Late Permian paleocommunities, documenting a substantial ecological shift to numerical dominance by molluscs in the Late Permian, before the major taxonomic shift at the end-Permian mass extinction. This ecological change was coincident with the development of fluctuating anoxic conditions in deep marine basins, suggesting that numerical dominance by more tolerant molluscs may have been driven by variably stressful environmental conditions. Recognition of substantial ecological deterioration in the Late Permian also implies that the end-Permian extinction was the climax of a protracted environmental crisis. Although the Late Permian shift to molluscan dominance was a pronounced ecological change, quantitative counts of 847 Carboniferous–Cretaceous collections from the Paleobiology Database indicate that it was only the first stage in a stepwise transition that culminated with the final shift to molluscan dominance in the Late Jurassic. Therefore, the ecological transition from brachiopods to bivalves was more protracted and complex than their simple Permian–Triassic switch in diversity. PMID:17664426

  16. Palaeoclimatic trends deduced from the hydrochemistry of a Triassic sandstone aquifer, U.K

    International Nuclear Information System (INIS)

    Bath, A.H.; Edmunds, W.M.; Andrews, J.N.

    1978-01-01

    A detailed geochemical study (elemental, isotopic and dissolved inert gases) of unconfined and confined sections of the Triassic non-marine sandstone aquifer in Eastern England has been undertaken. Aspects of the recharge history of this aquifer over the past 40 000 years are revealed by examination of the data. (orig./HK) [de

  17. Origin and tectonic evolution of upper Triassic Turbidites in the Indo-Burman ranges, West Myanmar

    Science.gov (United States)

    Yao, Wei; Ding, Lin; Cai, Fulong; Wang, Houqi; Xu, Qiang; Zaw, Than

    2017-11-01

    The Pane Chaung Formation is exposed in the Indo-Burman Ranges, and has been involved in collision between the Indian Plate and West Burma Block. However, controversies exist over the provenance and paleogeographic reconstruction of the Pane Chaung Formation. This study presents new petrographical and detrital zircon Usbnd Pb ages and Hf isotopic data from the Pane Chaung Formation in Rakhine Yoma and Chin Hills, west Myanmar. The depositional age of the Pane Chaung Formation is Late Triassic, indicated by the Carnian-Norian Halobia fossils and maximum depositional ages between 233.0 ± 2.5 Ma and 206.2 ± 1.8 Ma. Upper Triassic sandstones contain 290-200 Ma detrital zircons, εHf(t) values of - 6 to 11 and TDMC of 1.6 to 0.6 Ga, interpreted to be derived from West Papua region. The most abundant zircon age population of 750-450 Ma is derived from Pan-African orogenic belts in Australia. Zircons of 1250-900 Ma age were derived from the Grenvillian orogen in Australia. Archean zircons are interpreted to be derived from the Yilgarn and Pilbara cratons in Western Australia. Detrital zircon ages of the Pane Chaung Formation are distinct from similar aged strata in Indochina and Sibumasu, but comparable to NW Australia (Carnarvon Basin) and Greater India (Langjiexue Formation). It is suggested that the Pane Chaung Formation was deposited in a Late Triassic submarine fan along the northern margin of Australia.

  18. Effects of soil erosion and anoxic–euxinic ocean in the Permian–Triassic marine crisis

    Directory of Open Access Journals (Sweden)

    Kunio Kaiho

    2016-08-01

    Full Text Available The largest mass extinction of biota in the Earth’s history occurred during the Permian–Triassic transition and included two extinctions, one each at the latest Permian (first phase and earliest Triassic (second phase. High seawater temperature in the surface water accompanied by euxinic deep-intermediate water, intrusion of the euxinic water to the surface water, a decrease in pH, and hypercapnia have been proposed as direct causes of the marine crisis. For the first-phase extinction, we here add a causal mechanism beginning from massive soil and rock erosion and leading to algal blooms, release of toxic components, asphyxiation, and oxygen-depleted nearshore bottom water that created environmental stress for nearshore marine animals. For the second-phase extinction, we show that a soil and rock erosion/algal bloom event did not occur, but culmination of anoxia–euxinia in intermediate waters did occur, spanning the second-phase extinction. We investigated sedimentary organic molecules, and the results indicated a peak of a massive soil erosion proxy followed by peaks of marine productivity proxy. Anoxic proxies of surface sediments and water occurred in the shallow nearshore sea at the eastern and western margins of the Paleotethys at the first-phase extinction horizon, but not at the second-phase extinction horizon. Our reconstruction of ocean redox structure at low latitudes indicates that a gradual increase in temperature spanning the two extinctions could have induced a gradual change from a well-mixed oxic to a stratified euxinic ocean beginning immediately prior to the first-phase extinction, followed by culmination of anoxia in nearshore surface waters and of anoxia and euxinia in the shallow-intermediate waters at the second-phase extinction over a period of approximately one million years or more. Enhanced global warming, ocean acidification, and hypercapnia could have caused the second-phase extinction approximately 60 kyr after

  19. Permian-Triassic thermal anomaly of the active margin of South America as a result of plate kinematics reorganization

    Science.gov (United States)

    Riel, Nicolas; Jaillard, Etienne; Guillot, Stéphane; Martelat, Jean-Emmanuel; Braun, Jean

    2013-04-01

    From Permian to Triassic times, tectonic plate reorganization provoked Pangaea breakup, counterclockwise rotation of Gondwana, closing of the Paleo-Tethys Ocean and opening of the Neo-Tethys oceanic realm. Meanwhile, the switch from arc volcanism to widespread S-type magmatism along the western South American active margin around 275-265 Ma is symptomatic of the onset of a large-scale Permian-Triassic thermal anomaly (PTTA)affecting the whole margin. Here we report metamorphic and U-Pb geochronological results from the El Oro metamorphic complex in the forearc zone of southwestern Ecuador, which recorded the last step, at 230-225 Ma, of the PTTA. The change in the drift direction of Gondwana from north to east at ca. 270 Ma was related to plate reorganization and provoked the verticalization of the subducted Panthalassa slab. As the slab verticalized, strong heat advection produced a high heat flow beneath the active margin inducing the development of a huge thermal anomaly responsible for the PTTA, which lasted 30 Ma. This voluminous magmatic activity culminated at the Permian-Triassic boundary, and may have contributed to the degradation of life conditions on the Earth surface.

  20. Chromium isotope record of the Otavi Group, Namibia

    DEFF Research Database (Denmark)

    Rodler, Alexandra; Frei, Robert; Gaucher, Claudio

    2013-01-01

    -Triassic (~252 Ma) boundaries, which coincide with Central Atlantic Magmatic Province (CAMP) and Siberian Trap volcanism, respectively. The Triassic-Jurassic mass extinction is often contributed to carbon release driven ocean acidification while the Permian-Triassic mass extinction is suggested to be related...... to widespread ocean anoxia. We compare Permian-Triassic and Triassic-Jurassic ocean redox change along continental margins in different geographic regions (Permian-Triassic: Greenland, Svalbard, Iran; Triassic-Jurassic: UK, Austria) and discuss its role in marine mass extinction. Speciation of iron [(FeHR/ Fe...... extinctions however shows 2 phases of euxinia along continental margins, with an initial short peak at the onset of volcanism followed by a shift to ferruginous conditions, possibly due to a strongly diminished ocean sulphate reservoir because of massive initial pyrite burial. D34Spyrite suggests...

  1. Astronomical tuning and magnetostratigraphy of the Upper Triassic Xujiahe Formation of South China and Newark Supergroup of North America: Implications for the Late Triassic time scale

    Science.gov (United States)

    Li, Mingsong; Zhang, Yang; Huang, Chunju; Ogg, James; Hinnov, Linda; Wang, Yongdong; Zou, Zhuoyan; Li, Liqin

    2017-10-01

    The time scale of the Late Triassic Epoch has a divergence of age models, especially for the durations of competing definitions for its Rhaetian Stage (uppermost Triassic). The astrochronology derived from relative depth of lacustrine-bearing clastic successions and astronomically tuned geomagnetic polarity time scale (APTS) of the Newark Supergroup of eastern North America provides a basis for the Late Triassic time scale. However, the Newark APTS has been challenged regarding its age scale and completeness; therefore an independent astronomical-tuned magnetic polarity zonation is required to verify the upper Newark APTS reference scale. We compiled a 6.5 million year (myr) APTS with magnetic stratigraphy from four sections of the lacustrine-fluvial, dinosaur-track-bearing Xujiahe Formation in the Sichuan Basin of South China that also has dating from detrital zircons and regional biostratigraphy. Variations in natural gamma-ray and magnetic susceptibility that reflect variable continental weathering in the source regions of the Xujiahe Formation are paced by Milankovitch cycles, especially the 100-kyr short eccentricity and 405-kyr long eccentricity. The cycle-tuned magnetostratigraphy of the Xujiahe Formation is compared directly via the magnetic-polarity zones to the depth ranks of the Newark Supergroup that are indicative of relative depths of lacustrine facies. The Sichuan APTS indicates that there is no significant hiatus between the sedimentary succession and the basalt flows at the top of the Newark Supergroup. The Sichuan APTS is compatible with the magnetostratigraphy from the candidate Global Boundary Stratotype Section and Point (GSSP) for the Norian-Rhaetian boundary interval at the Pignola-Abriola of South Italy, but does not extend downward to the proposed GSSP in Austria associated with the longer Rhaetian option. The earliest dinosaur tracks in China are from the middle of this Xujiahe Formation, therefore are implied to be middle Rhaetian in age

  2. Initial mass function for early-type stars in starburst galaxies

    International Nuclear Information System (INIS)

    Sekiguchi, K.; Anderson, K.S.

    1987-01-01

    The IMF slope of early-type stars in starburst galaxies is investigated using IUE observations and a technique that utilizes mass-linewidth relations for early-type stars. Fourteen low-resolution IUE spectra of eight starburst galaxies and three H II region galaxies are used to obtain line-strength ratios Si IV(1400 A)/C IV(1550 A). These are compared to model line ratios, and indicate that the average IMF slope for OB stars in these intense star-formation regions is appreciably flatter than that of the solar neighborhood. 46 references

  3. Mineralogy and petrography of samples of Permo-Triassic sedimentary strata from Cumbria

    International Nuclear Information System (INIS)

    Kemp, S.J.; Milodowski, A.E.; Bloodworth, A.J.

    1989-03-01

    The mineralogy and petrography of Permo-Triassic sediments from Cumbria are described. The samples are duplicates of those selected for sorption experiments by Harwell Laboratory and are considered representative of Permo-Triassic strata underlying the BNFL Sellafield site. The regional geological context of the samples is briefly discussed. X-ray diffraction analysis indicates that the St Bees Sandstone and Eden Shales samples are predominantly composed of quartz with minor feldspar, gypsum and hematite. The clay assemblage is formed of dioctahedral mica and trioctahedral chloride with minor kaolinite. Diagnostic testing also reveals the presence of corrensite, a regularly interstratified chlorite-smectite, in one of the samples. Possible diagenetic processes responsible for corrensite formation, including the alteration of originally deposited smectite by magnesium-rich pore waters, are discussed. Modelling of the diffraction profile indicates that the chlorite and corrensite are iron-rich species. Back scattered electron microscopy shows the samples to be typical red bed sediments exhibiting framework grain dissolution, the breakdown and replacement of unstable ferromagnesian minerals being accompanied by the precipitation of hematite and anatase. Particle-size analysis shows similar distributions for the St Bees Sandstone samples but a greater proportion of clay material in the Eden Shales. (author)

  4. Magnetic microsperules in Permian and Triassic bedded chert from Southwest Japan

    OpenAIRE

    Iwahashi,Junko; Yoshida,Masaru; Miono,Shigeyoshi; Santosh,Geetha; Santosh,M.

    1991-01-01

    Magnetic microspherules collected from the Permian and Triassic bedded cherts in Southwest Japan were studied. The size range, as estimated from 801 microspherules, vary from 3 to 100μm, with a peak size distribution between 10-20μm. Three broad shapes are recognized : spherical-, drop- and rocket-shaped, all categories including hollow particles. The surface textures as studied from scanning electron microscope show five dominant varieties : random mosaic, dendritic mosaic, feather crystal, ...

  5. The Triassic upwelling system of Arctic Alaska

    Science.gov (United States)

    Yurchenko, I.; Graham, S. A.

    2017-12-01

    The Middle to Upper Triassic Shublik Formation of Arctic Alaska is a laterally and vertically heterogeneous rock unit that has been analyzed both in outcrop and in the subsurface. The Shublik Formation sediments are distinguished by a characteristic set of lithologies that include glauconitic, phosphatic, organic-rich, and cherty facies consistent with a coastal upwelling zone deposition interpretation. It is often recognized by abundance of impressions and shells of distinctive Triassic bivalves. To understand main controls on lithofacies distributions, this study reviews and refines lithologic and paleoenvironmental interpretations of the Shublik Formation, and incorporates the newly acquired detailed geochemical analyses of two complete Shublik cores. This work focuses on organic geochemistry (analyses of biomarkers and diamondoids), chemostratigraphy (hand-held XRF), and iron speciation analysis to reconstruct paleoproductivity and redox conditions. Based on the available evidence, during Shublik deposition, an upwelling-influenced open shelf resulted in high nutrient supply that stimulated algal blooms leading to high net organic productivity, reduced water transparency, oxygen deficiency, and water column stratification. Evidence of such eutrophic conditions is indicated by the lack of photic benthic organisms, bioturbation and trace fossils, and dominance of the monospecific light-independent epibenthic bivalves. The flat, subcircular, thin shells of these carbonate-secreting organisms allowed them to adapt to dysoxic conditions, and float on soft, soupy, muddy substrate. The distinctive clay- and organic-rich facies with abundant bivalves occurred on the mid to outer stable broad shelf, and were deposited when organic productivity at times overlapped with periods of increased siliciclastic input controlled by sea level and changes in local sediment dispersal systems, and therefore are more spatially and temporally localized than the widespread clay

  6. Detrital zircon U-Pb geochronology of Cambrian to Triassic miogeoclinal and eugeoclinal strata of Sonora, Mexico

    Science.gov (United States)

    Gehrels, G.E.; Stewart, John H.

    1998-01-01

    One hundred and eighty two individual detrital zircon grains from Cambrian through Permian miogeoclinal strata, Ordovician eugeoclinal rocks, and Triassic post-orogenic sediments in northwestern Sonora have been analyzed. During Cambrian, Devonian, Permian, and Triassic time, most zircons accumulating along this part of the Cordilleran margin were shed from 1.40-1.45 and 1.62-1.78 Ga igneous rocks that are widespread in the southwestern United States and northwestern Mexico. Zircons with ages of approximately 1.11 Ga are common in Cambrian strata and were apparently shed from granite bodies near the sample site. The sources of 225-280 Ma zircons in our Triassic sample are more problematic, as few igneous rocks of these ages are recognized in northwestern Mexico. Such sources may be present but unrecognized, or the grains could have been derived from igneous rocks of the appropriate ages to the northwest in the Mojave Desert region, to the east in Chihuahua and Coahuila, or to the south in accreted(?) arc-type terranes. Because the zircon grains in our Cambrian and Devonian to Triassic samples could have accumulated in proximity to basement rocks near their present position or in the Death Valley region of southern California, our data do not support or refute the existence of the Mojave-Sonora megashear. Ordovician strata of both miogeoclinal and eugeoclinal affinity are dominated by >1.77 Ga detrital zircons, which are considerably older than most basement rocks in the region. Zircon grains in the miogeoclinal sample were apparently derived from the Peace River arch area of northwestern Canada and transported southward by longshore currents. The eugeoclinal grains may also have come from the Peace River arch region, with southward transport by either sedimentary or tectonic processes, or they may have been shed from off-shelf slivers of continents (perhaps Antarctica?) removed from the Cordilleran margin during Neoproterozoic rifting. It is also possible that the

  7. Age and microfacies of oceanic Upper Triassic radiolarite components from the Middle Jurassic ophiolitic mélange in the Zlatibor Mountains (Inner Dinarides, Serbia and their provenance

    Directory of Open Access Journals (Sweden)

    Gawlick Hans-Jürgen

    2017-08-01

    Full Text Available Oceanic radiolarite components from the Middle Jurassic ophiolitic mélange between Trnava and Rožanstvo in the Zlatibor Mountains (Dinaridic Ophiolite Belt west of the Drina–Ivanjica unit yield Late Triassic radiolarian ages. The microfacies characteristics of the radiolarites show pure ribbon radiolarites without crinoids or thin-shelled bivalves. Beside their age and the preservation of the radiolarians this points to a deposition of the radiolarites on top of the oceanic crust of the Neo-Tethys, which started to open in the Late Anisian. South of the study area the ophiolitic mélange (Gostilje–Ljubiš–Visoka–Radoševo mélange contains a mixture of blocks of 1 oceanic crust, 2 Middle and Upper Triassic ribbon radiolarites, and 3 open marine limestones from the continental slope. On the basis of this composition we can conclude that the Upper Triassic radiolarite clasts derive either from 1 the younger parts of the sedimentary succession above the oceanic crust near the continental slope or, more convincingly 2 the sedimentary cover of ophiolites in a higher nappe position, because Upper Triassic ribbon radiolarites are only expected in more distal oceanic areas. The ophiolitic mélange in the study area overlies different carbonate blocks of an underlying carbonate-clastic mélange (Sirogojno mélange. We date and describe three localities with different Upper Triassic radiolarite clasts in a mélange, which occurs A on top of Upper Triassic fore-reef to reefal limestones (Dachstein reef, B between an Upper Triassic reefal limestone block and a Lower Carnian reef limestone (Wetterstein reef, and C in fissures of an Upper Triassic lagoonal to back-reef limestone (Dachstein lagoon. The sedimentary features point to a sedimentary and not to a tectonic emplacement of the ophiolitic mélange (= sedimentary mélange filling the rough topography of the topmost carbonate-clastic mélange below. The block spectrum of the underlying and

  8. Age and microfacies of oceanic Upper Triassic radiolarite components from the Middle Jurassic ophiolitic mélange in the Zlatibor Mountains (Inner Dinarides, Serbia) and their provenance

    Science.gov (United States)

    Gawlick, Hans-Jürgen; Djerić, Nevenka; Missoni, Sigrid; Bragin, Nikita Yu.; Lein, Richard; Sudar, Milan; Jovanović, Divna

    2017-08-01

    Oceanic radiolarite components from the Middle Jurassic ophiolitic mélange between Trnava and Rožanstvo in the Zlatibor Mountains (Dinaridic Ophiolite Belt) west of the Drina-Ivanjica unit yield Late Triassic radiolarian ages. The microfacies characteristics of the radiolarites show pure ribbon radiolarites without crinoids or thin-shelled bivalves. Beside their age and the preservation of the radiolarians this points to a deposition of the radiolarites on top of the oceanic crust of the Neo-Tethys, which started to open in the Late Anisian. South of the study area the ophiolitic mélange (Gostilje-Ljubiš-Visoka-Radoševo mélange) contains a mixture of blocks of 1) oceanic crust, 2) Middle and Upper Triassic ribbon radiolarites, and 3) open marine limestones from the continental slope. On the basis of this composition we can conclude that the Upper Triassic radiolarite clasts derive either from 1) the younger parts of the sedimentary succession above the oceanic crust near the continental slope or, more convincingly 2) the sedimentary cover of ophiolites in a higher nappe position, because Upper Triassic ribbon radiolarites are only expected in more distal oceanic areas. The ophiolitic mélange in the study area overlies different carbonate blocks of an underlying carbonate-clastic mélange (Sirogojno mélange). We date and describe three localities with different Upper Triassic radiolarite clasts in a mélange, which occurs A) on top of Upper Triassic fore-reef to reefal limestones (Dachstein reef), B) between an Upper Triassic reefal limestone block and a Lower Carnian reef limestone (Wetterstein reef), and C) in fissures of an Upper Triassic lagoonal to back-reef limestone (Dachstein lagoon). The sedimentary features point to a sedimentary and not to a tectonic emplacement of the ophiolitic mélange (= sedimentary mélange) filling the rough topography of the topmost carbonate-clastic mélange below. The block spectrum of the underlying and slightly older

  9. Molecular Evidence for Radical Changes in Ocean Chemistry Across the Permian Triassic Boundary at Meishan in South China

    Science.gov (United States)

    Love, G. D.; Cao, C.; Jin, Y.; Summons, R. E.

    2004-12-01

    Samples from outcrop and from a new core drilled through the Permian Triassic Boundary at the type section at Meishan have been examined for biomarker and isotopic evidence of biotic and associated environmental change. Late Permian sediments from Meishan Beds 22-27 are characterized by indicators of anoxia including low Pr/Ph ratios and abundant aryl isoprenoids and isoreneieratane derived from the precursor carotenoid isorenieratene. The latter compounds are biomarkers for green sulfur bacteria (Chlorobiaceae) and are considered reliable indicators of euxinic water columns where sulfide extends to the photic zone. The peak of Chlorobiaceae biomarker abundance coincides with a rapid and synchronous drop in the ƒO13C and ƒO15N values of kerogen. On passing up into the Early Triassic, the biomarker signal for Chlorobiaceae wanes and is almost absent by Bed 30 where it is replaced by one for cyanobacteria including abundant hopanes and 2-methylhopanes and accompanying methyl and dimethyl alkanes. A very high value for the hopane/sterane ratio from Beds 30-38 indicates continuing dominant cyanobacterial productivity and only minor inputs from an algal plankton. The prevalence of aryl isoprenoids in P-Tr sediments at the Meishan section of South China is also recorded in a recently cored borehole, Hovea-3, of the Perth Basin, Western Australia. This suggests similar paleoenvironmental conditions prevailed across the Tethys Ocean during and immediately after the P-Tr Boundary. In particular, the presence of biomarkers for Chlorobiaceae at two separate locations indicates that water column euxinia was pervasive during the extinction event and suggests that sulfide may have been a key toxic agent.

  10. The continental Permian-Triassic boundary in the Netherlands: Implications for the geomagnetic polarity time scale

    NARCIS (Netherlands)

    Szurlies, M.; Geluk, M.C.; Krijgsman, W.; Kürschner, W.M.

    2012-01-01

    In Central and NW Europe, the transition from the Permian to the Triassic (i.e., the Zechstein–Buntsandstein boundary interval) is developed mainly in red bed facies. This continental sedimentary succession is marked by relatively high sedimentation rates providing a high temporal resolution

  11. Mass Extinctions of Pangea (Jean Baptiste Lamarck Medal Lecture)

    Science.gov (United States)

    Wignall, Paul B.

    2017-04-01

    The 80 million years of Earth history from middle of the Permian to the early Jurassic were some of the worst life ever experienced. The interval includes two mass extinctions that bracket the Triassic period and several lesser crises. It was to be nearly another 120 million years before another major crisis was to strike (this time it was the famous one that removed the dinosaurs). So what was so bad about the 80 million years and why was it so good afterwards? My talk will try to provide at least some of the answers. There are plenty of clues. Notably, the interval coincides with the presence of the Pangea supercontinent and all the extinctions coincided with the eruption of large igneous provinces (LIPs). Indeed, every LIP of this interval coincides with an extinction crisis, a perfect correlation that completely breaks down afterwards. However, getting from correlation to causation is far from straight forward. There are many unknowns - how much gas was released by the volcanism, how quickly and what type of gases were they? These are all questions under investigation. Most of the extinctions of Pangean time coincide with rapid global warming and extensive marine anoxia suggesting that greenhouse gas emissions linked to volcanism were an important extinction driver. For the most severe crises (Permo-Triassic and end-Triassic) losses occurred throughout the food chain all the way down to the primary producers of the oceans and across all habitats including terrestrial ecosystems. At the other end of the spectrum of disaster, the lesser extinctions (Toarcian, Smithian/Spathian) only affected marine invertebrates. The full panoply of catastrophe was played out during the Permo-Triassic mass extinction and has received the most attention. The record in South China shows that there were two phases of extinction. These straddle the boundary and show selective losses initially for shallow-water organisms that were susceptible to high temperatures and then for deeper

  12. High influx of carbon in walls of agglutinated foraminifers during the Permian-Triassic transition in global oceans

    Science.gov (United States)

    Nestell, Galina P.; Nestell, Merlynd K.; Ellwood, Brooks B.; Wardlaw, Bruce R.; Basu, Asish R.; Ghosh, Nilotpal; Phuong Lan, Luu Thi; Rowe, Harry D.; Hunt, Andrew G.; Tomkin, Jonathan H.; Ratcliffe, Kenneth T.

    2015-01-01

    The Permian–Triassic mass extinction is postulated to be related to the rapid volcanism that produced the Siberian flood basalt (Traps). Unrelated volcanic eruptions producing several episodes of ash falls synchronous with the Siberian Traps are found in South China and Australia. Such regional eruptions could have caused wildfires, burning of coal deposits, and the dispersion of coal fly ash. These eruptions introduced a major influx of carbon into the atmosphere and oceans that can be recognized in the wallstructure of foraminiferal tests present in survival populations in the boundary interval strata. Analysis of free specimens of foraminifers recovered from residues of conodont samples taken at aPermian–Triassic boundary section at Lung Cam in northern Vietnam has revealed the presence of a significant amount of elemental carbon, along with oxygen and silica, in their test wall structure, but an absence of calcium carbonate. These foraminifers, identified as Rectocornuspira kalhori, Cornuspira mahajeri, and Earlandia spp. and whose tests previously were considered to be calcareous, are confirmed to be agglutinated, and are now referred to as Ammodiscus kalhori and Hyperammina deformis. Measurement of the 207Pb/204Pb ratios in pyrite clusters attached to the foraminiferal tests confirmed that these tests inherited the Pb in their outer layer from carbon-contaminated seawater. We conclude that the source of the carbon could have been either global coal fly ash or forest fire-dispersed carbon, or a combination of both, that was dispersed into the Palaeo-Tethys Ocean immediately after the end-Permian extinction event.

  13. The precise temporal calibration of dinosaur origins.

    Science.gov (United States)

    Marsicano, Claudia A; Irmis, Randall B; Mancuso, Adriana C; Mundil, Roland; Chemale, Farid

    2016-01-19

    Dinosaurs have been major components of ecosystems for over 200 million years. Although different macroevolutionary scenarios exist to explain the Triassic origin and subsequent rise to dominance of dinosaurs and their closest relatives (dinosauromorphs), all lack critical support from a precise biostratigraphically independent temporal framework. The absence of robust geochronologic age control for comparing alternative scenarios makes it impossible to determine if observed faunal differences vary across time, space, or a combination of both. To better constrain the origin of dinosaurs, we produced radioisotopic ages for the Argentinian Chañares Formation, which preserves a quintessential assemblage of dinosaurian precursors (early dinosauromorphs) just before the first dinosaurs. Our new high-precision chemical abrasion thermal ionization mass spectrometry (CA-TIMS) U-Pb zircon ages reveal that the assemblage is early Carnian (early Late Triassic), 5- to 10-Ma younger than previously thought. Combined with other geochronologic data from the same basin, we constrain the rate of dinosaur origins, demonstrating their relatively rapid origin in a less than 5-Ma interval, thus halving the temporal gap between assemblages containing only dinosaur precursors and those with early dinosaurs. After their origin, dinosaurs only gradually dominated mid- to high-latitude terrestrial ecosystems millions of years later, closer to the Triassic-Jurassic boundary.

  14. Paleoclimate cycles and tectonic controls on fluvial, lacustrine, and eolian strata in upper Triassic Chinle Formation, San Juan basin

    Energy Technology Data Exchange (ETDEWEB)

    Dubiel, R.F. (Geological Survey, Denver, CO (USA))

    1989-09-01

    Sedimentologic study of the Upper Triassic Chinle Formation in the San Juan basin (SJB) indicates that Late Triassic paleoclimate and tectonic movements influenced the distribution of continental lithofacies. The Shinarump, Monitor Butte, and Petrified Forest Members in the lower part of the Chinle consist of complexly interfingered fluvial, floodplain, marsh, and lacustrine rocks; the Owl Rock and Rock Point Members in the upper part consists of lacustrine-basin and eolian sandsheet strata. Facies analysis, vertebrate and invertebrate paleontology, and paleoclimate models demonstrate that the Late Triassic was dominated by tropical monsoonal circulation, which provided abundant precipitation interspersed with seasonally dry periods. Owl Rock lacustrine strata comprise laminated limestones that reflect seasonal monsoonal precipitation and larger scale, interbedded carbonates and fine-grained clastics that represent longer term, alternating wet and dry climatic cycles. Overlying Rock Point eolian sand-sheet and dune deposits indicate persistent alternating but drier climatic cyclicity. Within the Chinle, upward succession of lacustrine, alternating lacustrine/eolian sand-sheet, and eolian sand-sheet/dune deposits reflects an overall decrease in precipitation due to the northward migration of Pangaea out of low latitudes dominated by monsoonal circulation.

  15. The hELENa project - I. Stellar populations of early-type galaxies linked with local environment and galaxy mass

    NARCIS (Netherlands)

    Sybilska, A.; Lisker, T.; Kuntschner, H.; Vazdekis, A.; van de Ven, G.; Peletier, R.; Falcón-Barroso, J.; Vijayaraghavan, R.; Janz, J.

    2017-01-01

    We present the first in a series of papers in The role of Environment in shaping Low-mass Early-type Nearby galaxies (hELENa) project. In this paper, we combine our sample of 20 low-mass early types (dEs) with 258 massive early types (ETGs) from the ATLAS3D survey - all observed with the SAURON

  16. Remagnetization mechanisms in Triassic red beds from South China

    Science.gov (United States)

    Jiang, Zhaoxia; Liu, Qingsong; Dekkers, Mark J.; Zhao, Xiang; Roberts, Andrew P.; Yang, Zhenyu; Jin, Chunsheng; Liu, Jianxing

    2017-12-01

    Paleogeographic reconstructions based on paleomagnetic data rely on the reliability of the natural remanent magnetization (NRM) as a primary geomagnetic signal. Remagnetizations, however, can be common in many rock types, including late Paleozoic and Mesozoic red beds, and they complicate paleogeographic interpretations. Extracting the primary NRM from partially remagnetized rocks, and understanding the remagnetization mechanism are important in these contexts. We carried out a systematic paleomagnetic study of red bed samples from the Triassic Huangmaqing Formation, Nanjing (32.0°N, 118.9°E), South China. Two NRM components carried by secondary and primary hematite are isolated in 47 of the 94 samples studied, where the latter component has a direction in stratigraphic coordinates of D = 29.2 °, I = 34.6 ° (α95 = 10.9 °, 47 samples from 6 sites) that yields a paleopole of λ = 60.8°N, ϕ = 228.1°E, dp / dm = 12.5 / 7.2, which is consistent with Triassic pole positions for the South China Block. A secondary chemical remanent magnetization (CRM) (D = 227.1 °, I = 80.8 °, α95 = 7.3 °) is documented in all 94 samples from 10 sites and is carried by pigmentary hematite that is inferred to have been generated by magnetite oxidation during orogenic activity. This secondary component has steep inclinations and is interpreted to have been influenced by a combination of the remanence carried by original parent magnetite, the orogenic stress field, and the prevailing geomagnetic field direction during deformation. This CRM direction is recorded commonly by red beds from the South China Block, and is significant for regional tectonic studies in the area.

  17. Inclination Shallowing in the Permian/Triassic Boundary Sedimentary Sections of the East European Platform: the New Paleomagnetic Pole and its Significance for GAD Hypothesis

    Science.gov (United States)

    Veselovskiy, R. V.; Fetisova, A. M.; Balabanov, Y.

    2017-12-01

    One of the key challenges which are traditionally encountered in studying the paleomagnetism of terrigenous sedimentary strata is the necessity to allow for the effect of shallowing of paleomagnetic inclinations which takes place under the compaction of the sediment at the early stages of diagenesis and most clearly manifests itself in the case of midlatitude sedimentation. Traditionally, estimating the coefficient of inclination flattening (f) implies routine re-deposition experiments and studying their magnetic anisotropy (Kodama, 2012), which is not possible in every standard paleomagnetic laboratory. The Elongation-Inclination (E/I) statistical method for estimating the coefficient of inclination shallowing, which was recently suggested in (Tauxe and Kent, 2004), does not require the investigation of the rock material in a specially equipped laboratory but toughens the requirements on the paleomagnetic data and, primarily, regarding the volume of the data, which significantly restricts the possibilities of the post factum estimation and correction for inclination shallowing. We present the results of the paleomagnetic reinvestigation of the some key sections of the Upper Permian and Lower Triassic rocks located on the East European Platform. The obtained paleomagnetic data allowed us to estimate the coefficient of inclination shallowing by the E/I method and calculate a new P-Tr paleomagnetic pole for Europe. The absence of a statistically significant difference between the mean Siberian, European and North American Permian-Triassic paleomagnetic poles allow us to conclude that 252 Ma the configuration of the Earth's magnetic field was predominantly dipole. We believe that the assumption of the non-dipolarity of the geomagnetic field at the Permian-Triassic boundary, which has been repeatedly discussed in recent decades (Van der Voo and Torsvik, 2001; Bazhenov and Shatsillo, 2010; Veselovskiy and Pavlov, 2006), arose due to the failure to take into account the

  18. FIRST RECORD OF ICHTHYOSAURS IN SICILY(UPPER TRIASSIC OF MONTE SCALPELLO, CATANIA PROVINCE

    Directory of Open Access Journals (Sweden)

    CRISTIANO DAL SASSO

    2014-03-01

    Full Text Available Here we report for the first time on the presence of ichthyosaurs in Sicily, southern Italy. The deposit of origin (Mufara Formation can be dated to the upper Carnian (Tuvalian substage based on a typical association of ammonites, one of which (Shastites sp. is embedded in the sediment still encrusting one of the bone specimens recently found. The latter consist of two isolated vertebral centra that are referred to the Ichthyosauria thanks to their disk-like shape (i.e. they are much taller than long combined with the amphicelous condition, lack of transverse processes, and presence of rib articulations on the central sides. The largest specimen is more precisely an anterior dorsal vertebra from an adult individual, ascribed to Shastasauridae indet. By the presence of elongated reniform diapophyseal facets, cranially not truncated, and absence of parapophyses. The smaller specimen represents an anterior cervical element from an immature individual of a likely smaller-sized, indeterminate taxon. These finds indicate that the biodiversity of the Monte Scalpello Triassic fauna is higher than previously known, but above all represent the southernmost occurrence of Triassic ichthyosaurs in the western Tethys basin.

  19. Early detection of breast cancer mass lesions by mammogram segmentation images based on texture features

    International Nuclear Information System (INIS)

    Mahmood, F.H.

    2012-01-01

    Mammography is at present one of the available method for early detection of masses or abnormalities which is related to breast cancer.The calcifications. The challenge lies in early and accurate detection to overcome the development of breast cancer that affects more and more women throughout the world. Breast cancer is diagnosed at advanced stages with the help of the digital mammogram images. Masses appear in a mammogram as fine, granular clusters, which are often difficult to identify in a raw mammogram. The incidence of breast cancer in women has increased significantly in recent years. This paper proposes a computer aided diagnostic system for the extraction of features like mass lesions in mammograms for early detection of breast cancer. The proposed technique is based on a four-step procedure: (a) the preprocessing of the image is done, (b) regions of interest (ROI) specification, (c) supervised segmentation method includes two to stages performed using the minimum distance (M D) criterion, and (d) feature extraction based on Gray level Co-occurrence matrices GLC M for the identification of mass lesions. The method suggested for the detection of mass lesions from mammogram image segmentation and analysis was tested over several images taken from A L-llwiya Hospital in Baghdad, Iraq.The proposed technique shows better results.

  20. Late Paleozoic to Triassic magmatism in the north-central High Andes, Chile: New insights from SHRIMP U-Pb geochronology and O-Hf isotopic signatures in zircon

    Science.gov (United States)

    Hernández González, Álvaro; Deckart, Katja; Fanning, Mark; Arriagada, César

    2014-05-01

    The Chilean High Andes (28o- 31oS) comprises a vast number of late Paleozoic - Triassic granitoids which give information about the last stages of Gondwana assemblage. Particularly, previous studies determined two tectonic configurations during this time: subduction related compressional setting (late Carboniferous - Late Permian) and non-subduction post-collisional extensional setting (Late Permian - Triassic), as the last stage of Gondwana assemblage. However, new O-Hf isotopic data along new U-Pb SHRIMP ages in zircon have shown that this model should be modified and updated to the new analytical data available. δ18O values indicate a strong change in the tectonic configuration approximately 270 Ma (earliest middle Permian) and thus, units can be divided into 2 mayor groups: late Carboniferous to earliest middle Permian and middle Permian to Triassic. The oldest group shows slightly low values of ɛHfi (ca. +1 to -4) with high δ18O (ca. >6.5 o/oo), indicating an elevated supracrustal component and the addition of less radiogenic continental-like material, which along significant residence time (TDM2: Mesoproterozoic) can be interpreted as magmas formed at depth in a subduction-related continental arc, and contaminated with supracrustal material and/or oceanic sediments transported through the subducted slab to the mantle-wedge. Subsequently, middle Permian - Triassic rocks show a wider range of ɛHfi values (ca. +3 to -3) with relatively low, mantle-like δ18O (ca. 4.5-6.5 o/oo), indicating a source of magmas without the addition of supracrustal material for some plutons, whilst for others, a slight input. The higher positive values of ɛHfi can be related to the influence of new juvenile material in the source of some magmas. This isotopic data can be interpreted as rocks formed as the result of melting of an old thinned mafic crust (with mantle-like δ18O values characteristic of this type of rocks) with limited addition of supracrustal material; in

  1. Theropoda dinosaurs tracks from Triassic basin nd Ischigualasto - Villa Union, western Argentina

    International Nuclear Information System (INIS)

    Contreras, V.; Bracco, A.

    2004-01-01

    The Anchisauripus isp. and Theropoda indet. dinosaurs tracks from the Triassic Los Rastros and Ischigualasto formations, at the Ischigualasto-Villa Union Basin, Western Argentina, are described in this paper. This record completes the paleontological data provided by rest of bones and increases the stratigraphic range of some taxa in which bones are either missing or scarse. On the other hand, this report also allows us to enlarge the faunistic assemblage of some vertebrate assemblages [es

  2. Evolution of Northeastern Mexico during the early Mesozoic: potential areas for research and exploration José Rafael Barboza-Gudiño

    Science.gov (United States)

    Barboza-Gudiño, R.

    2013-05-01

    The lower Mesozoic succession of central and northeastern Mexico was deposited in a late Paleozoic-early Mesozoic remnant basin, formed at the westernmost culmination of the Ouachita-Marathon geosuture, after closure of the Rheic Ocean. Triassic fluvial deposits of El Alamar Formation (El Alamar River) are distributed in Tamaulipas and Nuevo Leon as remnants of a continental succession deposited close to the western margin of equatorial Pangea, such fluvial systems flowed to the ocean, located to the west and contributed to construction of the so-called Potosí submarine fan (Zacatecas Formation). Petrographic, geochemical, and detrital zircon geochronology studies indicate that both, marine and continental Triassic successions, come from a continental block and partially from a recycled orogen, showing grenvillian (900-1300 Ma) and Pan-African (500-700 Ma) zircon age populations, typical for peri-gondwanan blocks, in addition to zircons from the Permo-Triassic East Mexico arc (240-280 Ma). The absence of detrital zircons from the southwestern North American craton, represent a strong argument against left lateral displacement of Mexico to the southwest during the Jurassic up to their actual position, as proposed by the Mojave-Sonora megashear hypothesis. Towards the end of the Triassic or in earliest Jurassic time, began the subduction along the western margin of Pangea, which causes deformation of the Late Triassic Zacatecas Formation and subsequent magmatism in the continental Jurassic arc known as "Nazas Arc ", whose remnants are now exposed in central- to northeastern Mexico. Wide distributed in northern Mexico occurred also deposition of a red bed succession, overlying or partially interstratified with the Early to Middle Jurassic volcanic rocks of the Nazas Formation. To the west and southwest, such redbeds change transitionally to marine and marginal sedimentary facies which record sedimentation at the ancient paleo-pacific margin of Mexico (La Boca and

  3. The African cynodont Aleodon (Cynodontia, Probainognathia in the Triassic of southern Brazil and its biostratigraphic significance.

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    Agustín G Martinelli

    Full Text Available In this contribution we report the first occurrence of the enigmatic African probainognathian genus Aleodon in the Middle-early Late Triassic of several localities from the state of Rio Grande do Sul in southern Brazil. Aleodon is unusual among early probainognathians in having transversely-expanded postcanine teeth, similar to those of gomphodont cynognathians. This genus was previously known from the Manda Beds of Tanzania and the upper Omingonde Formation of Namibia. The Brazilian record of this genus is based upon multiple specimens representing different ontogenetic stages, including three that were previously referred to the sectorial-toothed probainognathian Chiniquodon theotonicus. We propose a new species of Aleodon (A. cromptoni sp. nov. based on the specimens from Brazil. Additionally, we tentatively refer one specimen from the upper Omingonde Formation of Namibia to this new taxon, strengthening biostratigraphic correlations between these strata. Inclusion of A. cromptoni in a phylogenetic analysis of eucynodonts recovers it as the sister-taxon of A. brachyrhamphus within the family Chiniquodontidae. The discovery of numerous specimens of Aleodon among the supposedly monospecific Chiniquodon samples of Brazil raises concerns about chiniquodontid alpha taxonomy, particularly given the extremely broad geographic distribution of Chiniquodon. The discovery of Brazilian Aleodon and new records of the traversodontid Luangwa supports the hypothesis that at least two subzones can be recognized in the Dinodontosaurus Assemblage Zone.

  4. Hydrogeological study of the Triassic series in the JeffaraDahar region (Southern part of Tunisia): Contribution of well logs data and seismic reflection

    Energy Technology Data Exchange (ETDEWEB)

    Ben Lasmar, R.; Guellala, R.; Zouhri, L.; Sarsar Naouali, B.; Garrach, M.; Inoubli, M.H.

    2016-07-01

    The present study concentrates on the interpretation of well logs and seismic reflection data in the JeffaraDahar region (Southeast part of Tunisia) for a better characterization of the Triassic aquifer, a potential target of water supply. Lithological columns and their corresponding well logs reveal that Sidi Stout, Kirchaou and Touareg. sandstones as well as Mekraneb and Rehach dolomites are the main reservoirs of the Triassic aquifer. Well log analysis highlights many permeable and fractured layers that could play an important role in the groundwater circulation. The interpreted seismic sections and the resulting isochrone maps show a tectonic influence on the Triassic aquifer geometry in the Jeffara-Dahar region. The normal faulting of E-W and NW-SE accidents created an aquifer compartmentalized by raised and tilted blocks. Seismic cross-sections reveal that this structure controls the depth of permeable formations and the circulation of groundwater. These results will be useful for rationalising the future hydrogeological research that will be undertaken in the Jeffara-Dahar area. (Author)

  5. Magnetic Polarity Stratigraphy and Rock Magnetic Data From the Continuous Cored Record of Triassic Continental Environmental Change, the Colorado Plateau Coring Project

    Science.gov (United States)

    Geissman, J. W.; McIntosh, J.; Buhedma, H. M. A.

    2017-12-01

    Despite the fact that the Triassic Period (ca. 251.9-201.3 Ma) is bound by two of Earth's largest mass extinctions, experienced giant bolide impacts and eruption of three large igneous provinces, and witnessed evolution of the main components of modern tetrapod communities, the time interval has sparse geochronologic calibration. The US NSF- and ICDP-funded coring of Phase 1 of the CPCP was completed in 2013, with the recovery of two major cores (6.35 cm diameter: 1A, 518m length and 2B, 253m; 31km apart) from the Petrified Forest National Park spanning the Chinle and Moenkopi fms. Core 1A has been fully sampled, with specimens obtained either by drilling or by extraction of core fragments and packing in ceramic boxes. Specimens are subjected to progressive thermal demagnetization or a combination of alternating field (AF) followed by thermal treatment. In several cases, specimens were extracted from each core segment to test for internal consistency. Chinle hematitic mudstones and siltstones have NRM intensities between 130 to 0.5 mA/m, with bulk susceptibilities from 2 x 10-2 to 5 x 10-5 SI units. More indurated hematitic siltstones/ medium sandstones of the Moenkopi Fm have NRM intensities and bulk susceptibilities that are far less variable (NRM: 9.0 to 1.2 mA/m, MS: 3.0 X 10-4 and 0.5 x 10-5 SI vol). Thermal demagnetization typically isolates magnetizations of N declination and shallow inclination (interpreted as normal polarity) and antipodes (reverse) (image), a polarity stratigraphy is being compiled for much of the section. Response is typically more interpretable for very hematitic Chinle mudstone sections and most Moenkopi rocks. Coarser grained, less hematitic Chinle strata rarely yield interpretable results, likely due to coarse-grained detrital magnetite, and it is likely that these intervals will not yield robust polarity information. Some core segments yield well-resolved magnetizations that are inconsistent with a Triassic field and we suspect

  6. Relationship between oxidative stress and muscle mass loss in early postmenopause: an exploratory study.

    Science.gov (United States)

    Zacarías-Flores, Mariano; Sánchez-Rodríguez, Martha A; García-Anaya, Oswaldo Daniel; Correa-Muñoz, Elsa; Mendoza-Núñez, Víctor Manuel

    2018-04-09

    Endocrine changes due to menopause have been associated to oxidative stress and muscle mass loss. The study objective was to determine the relationship between both variables in early postmenopause. An exploratory, cross-sectional study was conducted in 107 pre- and postmenopausal women (aged 40-57 years). Levels of serum lipid peroxides and uric acid and enzymes superoxide dismutase and glutathione peroxidase, as well as total plasma antioxidant capacity were measured as oxidative stress markers. Muscle mass using bioelectrical impedance and muscle strength using dynamometry were also measured. Muscle mass, skeletal muscle index, fat-free mass, and body mass index were calculated. More than 90% of participants were diagnosed with overweight or obesity. Postmenopausal women had lower values of muscle mass and strength markers, with a negative correlation between lipid peroxide level and skeletal muscle index (r= -0.326, p<.05), and a positive correlation between uric acid and skeletal muscle index (r=0.295, p<.05). A multivariate model including oxidative stress markers, age, and waist circumference showed lipid peroxide level to be the main contributor to explain the decrease in skeletal muscle mass in postmenopause, since for every 0.1μmol/l increase in lipid peroxide level, skeletal muscle index decreases by 3.03 units. Our findings suggest an association between increased oxidative stress and muscle mass loss in early postmenopause. Copyright © 2018 SEEN y SED. Publicado por Elsevier España, S.L.U. All rights reserved.

  7. Geochemical evidences for palaeoclimatic fluctuations at the Triassic-Jurassic boundary: southwestern margin of the Neotethys in the Salt Range, Pakistan

    Science.gov (United States)

    Iqbal, Shahid; Wagreich, Michael; Jan, Irfanullah; Kürschner, Wolfram Michael; Gier, Susanne

    2017-04-01

    The Triassic-Jurassic boundary interval reveals a change from warm-arid to a warm and humid climate in the Tethyan domain. Sea-level reconstruction records across the European basins during this interval reveal an end-Triassic global regression event and is linked to the Central Atlantic Magmatic Province (CAMP) activity and Pangaea breakup. In the Tethyan Salt Range of Pakistan a succession of Upper Triassic dolomites/green-black mudstones (Kingriali Formation), overlying quartzose sandstone, mudstones, laterites and Lower Jurassic conglomerates/pebbly sandstones (Datta Formation) provides information on the palaeoclimatic evolution of the area. Preliminary palynological results from the mudstones indicate a Rhaetian age for the Kingriali Formation and a Hettangian age for the Datta Formation. X-ray diffraction (XRD) analysis of the mudstones (upper part of the Kingriali Formation) indicates the presence of mainly illite while kaolinite is a minor component. The kaolinite content, a reflection of the advanced stage of chemical weathering and hence warm-humid conditions, increases up-section in the overlying sandstone-mudstone succession. The overlying laterite-bauxite horizons lack illite/smectite and are entirely composed of kaolinite, boehmite and haematite. At places these kaolinite rich horizons are mined in the area (Western Salt Range). The bulk rock geochemistry of the succession confirms a similar trend. The Chemical Index of Alteration (CIA) displays an increasing trend from the Upper Triassic shales (CIA 75-80) through the overlying sandstones/mudstones-laterites to the overlying quartz rich sandstones and mudstones (CIA 90-97). The overall results for the succession reveal an increasing chemical maturity trend (increase in the intensity of chemical weathering) from Rhaetian to Hettangian thereby supporting a change from warm-arid to a warm-humid palaeoclimate, probably extreme greenhouse conditions.

  8. SOME INOZOID SPONGES FROM UPPER TRIASSIC (NORIAN-RHAETIAN NAYBAND FORMATION OF CENTRAL IRAN

    Directory of Open Access Journals (Sweden)

    BABA SENOWBARI-DARYAN

    1997-11-01

    Full Text Available Some small-scaled Norian-Rhaetian reefs and reef mounds are imbedded within the shales, siltstones and siliciclastic-carbonate deposits of the Nayband Formation in central Iran. These deposits belong to the central Iranian plate as part of the Cimmerian Continent. Most of the biogenic rocks have a biostromal geometry, biohermal constructions are rare. Inozoid, sphinctozoid, and chaetetid sponges are, beside of corals and other reef builders, the most important reef organisms within these bioconstructions. In some reefs a variety of hexactinellid sponges also occur. The following inozoid sponges are described in this paper: Radiofibra norica n. sp., Permocorynella maxima n. sp., ?Sestrostomella robusta, Marawandia iranica, n. gen., n. sp. and Enaulofungia? triassica n. sp. The first four taxa are among the most abundant sponges within the Nayband Formation where it is exposed in several localities in central Iran. Enaulofungia? triassica, however, is not an abundant sponge there. These inozoid sponges have never been reported from the Triassic deposits of this region. Radiofibra, until now known only from the Upper Permian of Djebel Tebaga (Tunisia, is reported here for the first time from Triassic rocks. The stratigraphic as well as the paleogeographic distribution of all the genera are discussed.   

  9. The Pangaean megamonsoon - evidence from the Upper Triassic Chinle Formation, Colorado Plateau

    Science.gov (United States)

    Dubiel, R.F.; Totman, Parrish J.; Parrish, J.M.; Good, S.C.

    1991-01-01

    The Chinle was deposited between about 5?? to 15??N paleolatitude in the western equatorial region of Pangaea, a key area for documenting the effects of the monsoonal climate. This study summarizes sedimentological and paleontologic data from the Chinle Formation on the Colorado Plateau and integrates that data with paleoclimatic models. The evidence for abundant moisture and seasonality attest to the reversal of equatorial flow and support the hypothesis that the Triassic Pangaean climate was dominated by monsoonal circulation. -from Authors

  10. The Total Mass of the Early-Type Galaxy NGC 4649 (M60

    Directory of Open Access Journals (Sweden)

    Ćirković, M. M.

    2008-12-01

    Full Text Available In this paper the problem of the total mass and the total mass-to-light ratio of the early-type galaxy NGC~4649 (M60 is analyzed. Use is made of two independent techniques: the X-ray methodology which is based on the temperature of the X-ray halo of NGC~4649 and the tracer mass estimator (TME which uses globular clusters (GCs observed in this galaxy. The mass is calculated in Newtonian and MOdified Newtonian Dynamics (MOND approaches and it is found that inside 3 effective radii ($R_e$ there is no need for large amounts of dark matter. Beyond $3R_e$ the dark matter starts to play important dynamical role. The possible reasons for the discrepancy between the estimates of the total mass based on X-rays and TME in the outer regions of NGC~4649 are also discussed.

  11. A middle Permian ophiolite fragment in Late Triassic greenschist- to blueschist-facies rocks in NW Turkey: An earlier pulse of suprasubduction-zone ophiolite formation in the Tethyan belt

    Science.gov (United States)

    Topuz, Gültekin; Okay, Aral I.; Schwarz, Winfried H.; Sunal, Gürsel; Altherr, Rainer; Kylander-Clark, Andrew R. C.

    2018-02-01

    The Eastern Mediterranean region within the Tethyan belt is characterised by two main pulses of suprasubduction-zone ophiolite formation during the Early-Middle Jurassic and Late Cretaceous. Despite vast exposures of the Permo-Triassic accretionary complexes, related suprasubduction-zone ophiolites and the timing of subduction initiation leading to the formation of Permo-Triassic accretionary complexes are unknown so far. Here we report on a 40 km long and 0.3 to 1.8 km wide metaophiolite fragment within transitional greenschist- to blueschist-facies oceanic rocks from NW Turkey. The metaophiolite fragment is made up mainly of serpentinite and minor dykes or stocks of strongly sheared metagabbro with mineral assemblages involving actinolite/winchite, chlorite, epidote, albite, titanite and phengite. The metagabbro displays (i) variable CaO and MgO contents, (ii) anomalously high Mg# (= 100 ∗ molar MgO/(MgO + FeOtot)) of 75-88, and (iii) positive Eu anomalies, together with low contents of incompatible elements such as Ti, P and Zr, suggesting derivation from former plagioclase cumulates. The serpentinites comprise serpentine, ± chlorite, ± talc, ± calcite and relict Cr-Al spinel surrounded by ferrichromite to magnetite. Relict Cr-Al spinels are characterised by (i) Cr/(Cr + Al) ratios of 0.45-0.56 and Mg/(Mg + Fe2 +) ratio of 0.76-0.22, (ii) variable contents of ZnO and MnO, and (iii) extremely low TiO2 contents. Zn and Mn contents are probably introduced into Cr-Al spinels during greenschist- to blueschist metamorphism. Compositional features of the serpentinite such as (i) Ca- and Al-depleted bulk compositions, (ii) concave U-shaped, chondrite-normalised rare earth element patterns (REE) with enrichment of light and heavy REEs, imply that serpentinites were probably derived from depleted peridotites which were refertilised by light rare earth element enriched melts in a suprasubduction-zone mantle wedge. U-Pb dating on igneous zircons from three metagabbro

  12. Tectonic controls on preservation of Middle Triassic Halfway reservoir facies, Peejay Field, northeastern British Columbia: a new hydrocarbon exploration model

    Energy Technology Data Exchange (ETDEWEB)

    Caplan, M. L. [British Columbia Univ., Vancouver, BC (Canada). Dept. of Geological Sciences; Moslow, T. F. [Calgary Univ., AB (Canada). Dept. of Geology and Geophysics

    1997-12-01

    The Peejay Field in northeastern British Columbia was chosen as the site of a detailed study to establish the paleogeography, geological history and genesis of reservoir facies of Middle Triassic strata. A total of 132 cores and well logs from 345 wells were examined to establish the depositional model, to identify the origin of all reservoir facies and to construct an exploration model to improve the prediction of reservoir facies. Results show that the Middle Triassic Halfway Formation of northeastern British Columbia is comprised of at least four west-southwest prograding paleoshorelines. The Lithofacies Succession One quartz-arenites paleoshore faces have less porosity and permeability and are laterally discontinuous. For these reasons shoreface facies have minimal reservoir quality. The tidal inlet fill successions were found to have the greatest observed porosity, permeability and lateral continuity in the Peejay Field. The geometry and orientation of these tidal inlet fill deposits are controlled by tectonic processes. It was suggested that the success of hydrocarbon exploration in this structurally complex area of northeastern British Columbia and west-central Alberta depends on further stratigraphic and sedimentological examination of Middle Triassic strata on a regional scale to obtain a complete understanding of the geological history of the area. 39 refs., 13 refs.

  13. Marine anoxia and delayed Earth system recovery after the end-Permian extinction.

    Science.gov (United States)

    Lau, Kimberly V; Maher, Kate; Altiner, Demir; Kelley, Brian M; Kump, Lee R; Lehrmann, Daniel J; Silva-Tamayo, Juan Carlos; Weaver, Karrie L; Yu, Meiyi; Payne, Jonathan L

    2016-03-01

    Delayed Earth system recovery following the end-Permian mass extinction is often attributed to severe ocean anoxia. However, the extent and duration of Early Triassic anoxia remains poorly constrained. Here we use paired records of uranium concentrations ([U]) and (238)U/(235)U isotopic compositions (δ(238)U) of Upper Permian-Upper Triassic marine limestones from China and Turkey to quantify variations in global seafloor redox conditions. We observe abrupt decreases in [U] and δ(238)U across the end-Permian extinction horizon, from ∼3 ppm and -0.15‰ to ∼0.3 ppm and -0.77‰, followed by a gradual return to preextinction values over the subsequent 5 million years. These trends imply a factor of 100 increase in the extent of seafloor anoxia and suggest the presence of a shallow oxygen minimum zone (OMZ) that inhibited the recovery of benthic animal diversity and marine ecosystem function. We hypothesize that in the Early Triassic oceans-characterized by prolonged shallow anoxia that may have impinged onto continental shelves-global biogeochemical cycles and marine ecosystem structure became more sensitive to variation in the position of the OMZ. Under this hypothesis, the Middle Triassic decline in bottom water anoxia, stabilization of biogeochemical cycles, and diversification of marine animals together reflect the development of a deeper and less extensive OMZ, which regulated Earth system recovery following the end-Permian catastrophe.

  14. Heterogeneous Rates of Molecular Evolution and Diversification Could Explain the Triassic Age Estimate for Angiosperms.

    Science.gov (United States)

    Beaulieu, Jeremy M; O'Meara, Brian C; Crane, Peter; Donoghue, Michael J

    2015-09-01

    Dating analyses based on molecular data imply that crown angiosperms existed in the Triassic, long before their undisputed appearance in the fossil record in the Early Cretaceous. Following a re-analysis of the age of angiosperms using updated sequences and fossil calibrations, we use a series of simulations to explore the possibility that the older age estimates are a consequence of (i) major shifts in the rate of sequence evolution near the base of the angiosperms and/or (ii) the representative taxon sampling strategy employed in such studies. We show that both of these factors do tend to yield substantially older age estimates. These analyses do not prove that younger age estimates based on the fossil record are correct, but they do suggest caution in accepting the older age estimates obtained using current relaxed-clock methods. Although we have focused here on the angiosperms, we suspect that these results will shed light on dating discrepancies in other major clades. ©The Author(s) 2015. Published by Oxford University Press, on behalf of the Society of Systematic Biologists. All rights reserved. For Permissions, please email: journals.permissions@oup.com.

  15. Sedimentology of the Essaouira Basin (Meskala Field) in context of regional sediment distribution patterns during upper Triassic pluvial events

    Science.gov (United States)

    Mader, Nadine K.; Redfern, Jonathan; El Ouataoui, Majid

    2017-06-01

    Upper Triassic continental clastics (TAGI: Trias Argilo-Greseux Inferieur) in the Essaouira Basin are largely restricted to the subsurface, which has limited analysis of the depositional environments and led to speculation on potential provenance of the fluvial systems. Facies analysis of core from the Meskala Field onshore Essaouira Basin is compared with tentatively time-equivalent deposits exposed in extensive outcrops in the Argana Valley, to propose a process orientated model for local versus regional sediment distribution patterns in the continuously evolving Moroccan Atlantic rift during Carnian to Norian times. The study aims to unravel the climatic overprint and improve the understanding of paleo-climatic variations along the Moroccan Atlantic margin to previously recognised Upper Triassic pluvial events. In the Essaouira Basin, four facies associations representing a progressive evolution from proximal to distal facies belts in a continental rift were established. Early ephemeral braided river systems are succeeded by a wet aeolian sandflat environment with a strong arid climatic overprint (FA1). This is followed by the onset of perennial fluvial deposits with extensive floodplain fines (FA2), accompanied by a distinct shift in fluvial style, suggesting increase in discharge and related humidity, either locally or in the catchment area. The fluvial facies transitions to a shallow lacustrine or playa lake delta environment (FA3), which exhibits cyclical abandonment. The delta is progressively overlain by a terminal playa with extensive, mottled mudstones (FA4), interpreted to present a return from cyclical humid-arid conditions to prevailing aridity in the basin. In terms of regional distribution and sediment source provenance, paleocurrent data from Carnian to Norian deposits (T5 to T8 member) in the Argana Valley suggest paleoflow focused towards the S and SW, not directed towards the Meskala area in the NW as previously suggested. A major depo

  16. Magnetostratigraphy of a Marine Triassic-Jurassic Boundary Section, Kennecott Point, Queen Charlotte Islands: Implications for the Temporal Correlation of a 'Big Five' Mass Extinction Event.

    Science.gov (United States)

    Hilburn, I. A.; Kirschvink, J. L.; Ward, P. D.; Haggart, J. W.; Raub, T. D.

    2008-12-01

    Several causes have been proposed for Triassic-Jurassic (T-J) boundary extinctions, including global ocean anoxia/euxinia, an impact event, and/or eruption of the massive Central Atlantic Magmatic Province (CAMP), but poor intercontinental correlation makes testing these difficult. Sections at Kennecott Point, Queen Charlotte Islands, British Columbia span the late Norian through Rhaetian (Triassic) and into the earliest Hettangian (Jurassic) and provide the best integrated magneto- and chemostratigraphic framework for placing necessary temporal constraints upon the T-J mass extinctions. At Kennecott Point, turnover of radiolaria and ammonoids define the T-J boundary marine extinction and are coincident with a 2 ‰ negative excursion in δ13Corg similar in magnitude to that observed at Ferguson Hill (Muller Canyon), Nevada (1, 2). With Conodont Alteration Index values in the 1-2 range, Kennecott Point provides the ideal setting for use of magnetostratigraphy to tie the marine isotope excursion into the chronostratigraphic framework of the Newark, Hartford, and Fundy Basins. In the summer of 2005, we collected a ~1m resolution magnetostratigraphic section from 105 m of deep marine, silt- and sandstone turbidites and interbedded mudstones, spanning the T-J boundary at Kennecott Point. Hybrid progressive demagnetization - including zero-field, low-temperature cycling; low-field AF cleaning; and thermal demagnetization in ~25°C steps to 445°C under flowing N2 gas (3) - first removed a Northerly, steeply inclined component interpreted to be a Tertiary overprint, revealing an underlying dual-polarity component of moderate inclination. Five major polarity zones extend through our section, with several short, one-sample reversals interspersed amongst them. Comparison of this pattern with other T-J boundary sections (4-6) argues for a Northern hemisphere origin of our site, albeit with large vertical-axis rotations. A long normal chron bounds the T-J boundary punctuated

  17. Spatial coincidence and similar geochemistry of Late Triassic and Eocene-Oligocene magmatism in the Andes of northern Chile: evidence from the MMH porphyry type Cu-Mo deposit, Chuquicamata District

    Science.gov (United States)

    Zentilli, Marcos; Maksaev, Victor; Boric, Ricardo; Wilson, Jessica

    2018-04-01

    The MMH porphyry type copper-molybdenum deposit in northern Chile is the newest mine in the Chuquicamata District, one of largest copper concentrations on Earth. Mineralized Eocene-Oligocene porphyry intrusions are hosted by essentially barren Triassic granodiorites. Despite a century of exploitation, geologists still have problems in the mine distinguishing the Triassic granodiorite from the most important ore-carrying Eocene porphyries in the district. To resolve the problem, internally consistent high-quality geochemical analyses of the Triassic and Tertiary intrusives were carried out: explaining the confusion, they show that the rock units in question are nearly identical in composition and thus respond equally to hydrothermal alteration. In detail, the only difference in terms of chemical composition is that the main Eocene-Oligocene porphyries carry relatively less Fe and Ni. Unexpectedly, the mineralized Eocene-Oligocene porphyries have consistently less U and Th than other Tertiary intrusions in the district, a characteristic that may be valuable in exploration. The supergiant copper-molybdenum deposits in the Central Andes were formed within a narrow interval between 45 and 31 Ma, close to 7% of the 200 My duration of "Andean" magmatism, which resulted from subduction of oceanic lithosphere under South America since the Jurassic. Although recent work has shown that subduction was active on the margin since Paleozoic times, pre-Andean (pre-Jurassic) "Gondwanan" magmatism is often described as being very different, having involved crustal melting and the generation of massive peraluminous rhyolites and granites. This study shows that the indistinguishable Late Triassic and Eocene-Oligocene intrusions occupy the same narrow NS geographic belt in northern Chile. If it is accepted that magma character may determine the potential to generate economic Cu-Mo deposits, then Late Triassic volcano-plutonic centres in the same location in the South American margin

  18. A Review on Permian to Triassic Active or Convergent Margin in Southeasternmost Gondwanaland: Possibility of Exploration Target for Tin and Hydrocarbon Deposits in the Eastern Indonesia

    Directory of Open Access Journals (Sweden)

    Amiruddin Amiruddin

    2014-06-01

    Full Text Available http://dx.doi.org/10.17014/ijog.vol4no1.20094An active convergence of continental margin is probably generated in Gondwanaland during Permian to Triassic period which is characterized by the presence of magmatic and volcanic belts and back-arc ba- sins occupied respectively by Permian to Triassic rocks. The magmatic belt is occupied by peraluminous granitic plutons showing characteristics of S- type granite and is considered as tin-bearing granites. The back-arc basins are occupied by the Southern Papua and Galille-Bowen-Gunnedah-Sydney Basins. Those large basins are respectivelly filled by fluvial, fluvio- deltaic to marine Permian-Triassic sediments, which are unconformably overlain by the Jurrassic-Cretaceous marine succession. The paleomagnetic data, confirmed by flora content found in Australia and Papua, indicate that those areas initially belong to the Gondwanaland before part of them were drifted and rotated into the present day position. Tectonically, the presence of those Permian-Triassic magmatic-volcanic belts and back-arc basins in behind, indicates that at the time there were huge compressive activities: convergence of paleo-oceanic Pasific Plate moving westward, collided and subducted into the Southeastern Gondwana Continental Plate, moved relatively eastwards. This phenomenon resembles to the formation of Sumatera Tertiary tectonic zones producing back-arc basins, i.e. South Sumatera, Central, and North Sumatera Basins including the Tertiary Magmatic Arc. Concerning the similarity of Permian-Triassic geological condition of the magmatic arc and back-arc basins in Eastern Indonesia and Eastern Australia including paleoposition, paleotectonic setting, strati- graphic succession, and lithologic composition, it is suggested to carry out an increase in a more intens- ive tin exploration in the Eastern Indonesia, e.g. Bird Head area and Banggai Sula Island, and also for hydrocarbon target (coal, coalbed methane, oil and gas, and oil

  19. The hELENa project - I. Stellar populations of early-type galaxies linked with local environment and galaxy mass

    OpenAIRE

    Sybilska, A.; Lisker, T.; Kuntschner, H.; Vazdekis, A.; van de Ven, G.; Peletier, R.; Falcón-Barroso, J.; Vijayaraghavan, R.; Janz, J.

    2017-01-01

    We present the first in a series of papers in T$h$e role of $E$nvironment in shaping $L$ow-mass $E$arly-type $N$earby g$a$laxies (hELENa) project. In this paper we combine our sample of 20 low-mass early types (dEs) with 258 massive early types (ETGs) from the ATLAS$^{\\mathrm{3D}}$ survey - all observed with the SAURON integral field unit (IFU) - to investigate early-type galaxies' stellar population scaling relations and the dependence of the population properties on local environment, exten...

  20. A Middle Triassic pachypleurosaur (Diapsida: Eosauropterygia) from a restricted carbonate ramp in the Western Carpathians (Gutenstein Formation, Fatric Unit): paleogeographic implications

    Science.gov (United States)

    Čerňanský, Andrej; Klein, Nicole; Soták, Ján; Olšavský, Mário; Šurka, Juraj; Herich, Pavel

    2018-02-01

    An eosauropterygian skeleton found in the Middle Triassic (upper Anisian) Gutenstein Formation of the Fatric Unit (Demänovská dolina Valley, Low Tatra Mountains, Slovakia) represents the earliest known occurrence of marine tetrapods in the Western Carpathians. The specimen represents a partly articulated portion of the postcranial skeleton (nine dorsal vertebrae, coracoid, ribs, gastral ribs, pelvic girdle, femur and one zeugopodial element). It is assigned to the Pachypleurosauria, more precisely to the Serpianosaurus-Neusticosaurus clade based on the following combination of features: (1) small body size; (2) morphology of vertebrae, ribs and femur; (3) tripartite gastral ribs; and (4) microanatomy of the femur as revealed by μCT. Members of this clade were described from the epicontinental Germanic Basin and the Alpine Triassic (now southern Germany, Switzerland, Italy), and possibly from Spain. This finding shows that pachypleurosaur reptiles attained a broader geographical distribution during the Middle Triassic, with their geographical range reaching to the Central Western Carpathians. Pachypleurosaurs are often found in sediments formed in shallow, hypersaline carbonate-platform environments. The specimen found here occurs in a succession with vermicular limestones in a shallow subtidal zone and stromatolitic limestones in a peritidal zone, indicating that pachypleurosaurs inhabited hypersaline, restricted carbonate ramps in the Western Carpathians.

  1. A Middle Triassic pachypleurosaur (Diapsida: Eosauropterygia from a restricted carbonate ramp in the Western Carpathians (Gutenstein Formation, Fatric Unit: paleogeographic implications

    Directory of Open Access Journals (Sweden)

    Čerňanský Andrej

    2018-02-01

    Full Text Available An eosauropterygian skeleton found in the Middle Triassic (upper Anisian Gutenstein Formation of the Fatric Unit (Demänovská dolina Valley, Low Tatra Mountains, Slovakia represents the earliest known occurrence of marine tetrapods in the Western Carpathians. The specimen represents a partly articulated portion of the postcranial skeleton (nine dorsal vertebrae, coracoid, ribs, gastral ribs, pelvic girdle, femur and one zeugopodial element. It is assigned to the Pachypleurosauria, more precisely to the Serpianosaurus–Neusticosaurus clade based on the following combination of features: (1 small body size; (2 morphology of vertebrae, ribs and femur; (3 tripartite gastral ribs; and (4 microanatomy of the femur as revealed by μCT. Members of this clade were described from the epicontinental Germanic Basin and the Alpine Triassic (now southern Germany, Switzerland, Italy, and possibly from Spain. This finding shows that pachypleurosaur reptiles attained a broader geographical distribution during the Middle Triassic, with their geographical range reaching to the Central Western Carpathians. Pachypleurosaurs are often found in sediments formed in shallow, hypersaline carbonate-platform environments. The specimen found here occurs in a succession with vermicular limestones in a shallow subtidal zone and stromatolitic limestones in a peritidal zone, indicating that pachypleurosaurs inhabited hypersaline, restricted carbonate ramps in the Western Carpathians.

  2. New Paleomagnetic Data from Upper Permian and Lower Triassic Volcanic Sequences from Hua Binh, Quynh Nhai, and Thuan Chau Localities, Northwest Veitnam and Their Bearing on the Accretion History of Southeast Asia

    Science.gov (United States)

    Geissman, J. W.; Chi, C. T.

    2015-12-01

    New paleomagnetic data from Upper Permian to Lower Triassic volcanic rocks sampled in NW Vietnam provide more quantitative constraints on the paleogeographic setting of crustal elements that comprise the Song Da Terrane, east of the Song Ma suture, between the South China block (SCB) and north Indochina. These include results from 12 sites (125 samples) from basalts of the Vien Nam Formation, exposed at Hoa Binh Dam; eight sites (74 samples) from basalts of the Cam Thuy Formation near Thuan Chau; and 19 sites (198 samples) from andesites and basalts of the Vien Nam Formation near Quynh Nhai. The collection is limited by the quality of exposures and quantity of independent flows. Most sites yield interpretable magnetizations in progressive demagnetization, and the response implies that characteristic remanent magnetization (ChRM) components are carried by low-titanium magnetite or hematite, or a combination of both; these are isolated from secondary components. Rock magnetic data and petrography support the retention of an early-acquired thermoremanent magnetization in most sites. The Vien Nam Formation mafic volcanic rocks yield a grand mean, in geographic coordinates, of D=33.8o, I=-28.4o ( a95 = 9.5o, k =30.3, N=9 accepted sites), and a pole position at Lat=41.1N, Long=239.8E and a paleolatitude at ~15o S during the Late Permian to Early Triassic. Permian basalts of the Cam Thuy Formation provide a grand mean, corrected for structural tilt, of D=216.1o, I=+10.5o, a95=8.9o, k=107.8, and N= 4, with a pole position at Lat=45.6N, Long=226.8E. Volcanic rocks at the Quynh Nhai locality likely yield the most robust paleofield determination, as the data set is of dual polarity and passes a reversal test. The tilt corrected grand mean (normal polarity) is D=48.3o, I=-10.0o, a95=8.0o, k=27.7, N = 13, and this in turn yields an inferred paleomagnetic pole at Lat=35.7N, Long=217.4E, and a paleolatitude of 5.1oS for the late Permian. Compared with the Late Permian-Early

  3. A Middle Triassic stem-turtle and the evolution of the turtle body plan.

    Science.gov (United States)

    Schoch, Rainer R; Sues, Hans-Dieter

    2015-07-30

    The origin and early evolution of turtles have long been major contentious issues in vertebrate zoology. This is due to conflicting character evidence from molecules and morphology and a lack of transitional fossils from the critical time interval. The ∼220-million-year-old stem-turtle Odontochelys from China has a partly formed shell and many turtle-like features in its postcranial skeleton. Unlike the 214-million-year-old Proganochelys from Germany and Thailand, it retains marginal teeth and lacks a carapace. Odontochelys is separated by a large temporal gap from the ∼260-million-year-old Eunotosaurus from South Africa, which has been hypothesized as the earliest stem-turtle. Here we report a new reptile, Pappochelys, that is structurally and chronologically intermediate between Eunotosaurus and Odontochelys and dates from the Middle Triassic period (∼240 million years ago). The three taxa share anteroposteriorly broad trunk ribs that are T-shaped in cross-section and bear sculpturing, elongate dorsal vertebrae, and modified limb girdles. Pappochelys closely resembles Odontochelys in various features of the limb girdles. Unlike Odontochelys, it has a cuirass of robust paired gastralia in place of a plastron. Pappochelys provides new evidence that the plastron partly formed through serial fusion of gastralia. Its skull has small upper and ventrally open lower temporal fenestrae, supporting the hypothesis of diapsid affinities of turtles.

  4. Structural complexity at and around the Triassic-Jurassic GSSP at Kuhjoch, Northern Calcareous Alps, Austria

    Science.gov (United States)

    Palotai, M.; Pálfy, J.; Sasvári, Á.

    2017-10-01

    One of the key requirements for a Global Stratotype Section and Point (GSSP) is the absence of tectonic disturbance. The GSSP for the Triassic-Jurassic system boundary was recently defined at Kuhjoch, Northern Calcareous Alps, Austria. New field observations in the area of the Triassic-Jurassic boundary GSSP site demonstrate that the overturned, tight, and almost upright Karwendel syncline was formed at semibrittle deformation conditions, confirmed by axial planar foliation. Tight to isoclinal folds at various scales were related to a tectonic transport to the north. Brittle faulting occurred before and after folding as confirmed by tilt tests (the rotation of structural data by the average bedding). Foliation is ubiquitous in the incompetent units, including the Kendlbach Formation at the GSSP. A reverse fault (inferred to be formed as a normal fault before folding) crosscuts the GSSP sections, results in the partial tectonic omission of the Schattwald Beds, and thus makes it impossible to measure a complete and continuous stratigraphic section across the whole Kendlbach Formation. Based on these observations, the Kuhjoch sections do not fulfil the specific requirement for a GSSP regarding the absence of tectonic disturbances near boundary level.

  5. Impact of diagenesis and low grade metamorphosis on Triassic sabkha dolomite δ26Mg

    Science.gov (United States)

    Immenhauser, A.; Geske, A.; Richter, D.; Buhl, D.; Niedermayr, A.

    2012-12-01

    Dolomite is a common rock forming mineral in the geological record but its value as archive of ancient seawater δ26Mg signatures and their variations in time are at present underexplored. Unknown factors include the sensitivity of δ26Mg ratio to processes in the diagenetic and low grade metamorphic domain. Here, we document and discusses the first detailed δ26Mg data set from early diagenetic and burial dolomites. Samples come from the Upper Triassic Hauptdolomit (Dolomia Principale; The Dolomites, Italy) and include coeval dolmicrites that underwent differential burial diagenesis in a temperature range between about 100 and more than 350°C. As indicated by dolmicrite 87/86Sr ratios, sabkha calcian D1 dolomites precipitated from evaporated seawater and stabilized at an early diagenetic stage to D2 dolomites analysed here. With increasing burial temperature, dolomite δ26Mg ratio scatter in the data set decreases with increasing Mg/Ca ratio and degree of order. Specifically, δ26Mg ratio variability is reduced from ~0.7‰ at burial temperatures beneath 100°C to about ~0.2‰ at temperatures in excess of 350°C, respectively, with mean δ26Mg values ranging constantly near -1.9‰. This suggests that, at least for the rock buffered system investigated here, dolmicrite δ26Mg proxy data are conservative and preserve near pristine values even at elevated burial temperatures. At present, the main element of uncertainty is the Mg-isotope fractionation factor between (evaporated) seawater and dolomite. A possible solution to this problem includes the compilation of a data from modern sabkha environments including pore water and calcian dolomite δ26Mg isotope signatures.

  6. Osteology of a new specimen of Macrocnemus aff. M. fuyuanensis (Archosauromorpha, Protorosauria) from the Middle Triassic of Europe: potential implications for species recognition and paleogeography of tanystropheid protorosaurs

    Science.gov (United States)

    Jaquier, Vivien P.; Fraser, Nicholas C.; Furrer, Heinz; Scheyer, Torsten M.

    2017-11-01

    Over the past two decades, a wealth of marine and terrestrial reptiles, including protorosaurian archosauromorphs, has been described from Triassic shales and limestone layers in southern China. Recovered from the eastern margin of the Tethys Ocean, these forms often show remarkable similarities to taxa that were previously known and described from Europe, i.e., the western Tethyan margin. One protorosaurian that is known from the western and the eastern Tethyan province is the genus Macrocnemus, with currently three recognized species: 1) M. bassanii from the Middle Triassic Besano Formation and Meride Limestone (late Anisian – early Ladinian), UNESCO World Heritage Site Monte San Giorgio, Ticino, Switzerland; 2) M. obristi from the Prosanto Formation (early Ladinian) of the Ducan area, Grisons, Switzerland; and 3) M. fuyuanensis from the Falang Formation (Ladinian), Yunnan Province, southern China. Recently a new specimen, PIMUZ T 1559, from the upper Besano Formation at Meride, Ticino, Switzerland, was prepared, revealing a disarticulated skeleton which includes most of the cranium and lower jaw, pre-caudal vertebral column and ribs, the forelimbs, and girdle elements. Unambiguously assignable to the genus Macrocnemus, it evinces particularly gracile elongated cervical ribs, as well as a humerus/radius ratio that is comparable only to that of M. fuyuanensis from southern China. Based on this feature we tentatively recognize the new specimen as M. aff. fuyuanensis from Europe. The position and exquisite preservation of the clavicle and interclavicle in this specimen allows a revision of the shoulder girdle of Macrocnemus when articulated, which also has implications for closely related protorosaurian taxa, such as the long-necked Tanystropheus. Furthermore, differences in the shape and morphology of the interclavicle including pointed wing-like lateral processes and a short, fusiform caudal process represent rare discrete characters that allow separation of the

  7. Osteology of a New Specimen of Macrocnemus aff. M. fuyuanensis (Archosauromorpha, Protorosauria from the Middle Triassic of Europe: Potential Implications for Species Recognition and Paleogeography of Tanystropheid Protorosaurs

    Directory of Open Access Journals (Sweden)

    Vivien P. Jaquier

    2017-11-01

    Full Text Available Over the past two decades, a wealth of marine and terrestrial reptiles, including protorosaurian archosauromorphs, has been described from Triassic shales and limestone layers in southern China. Recovered from the eastern margin of the Tethys Ocean, these forms often show remarkable similarities to taxa that were previously known and described from Europe, i.e., the western Tethyan margin. One protorosaurian that is known from the western and the eastern Tethyan province is the genus Macrocnemus, with currently three recognized species: (1 Macrocnemus bassanii from the Middle Triassic Besano Formation and Meride Limestone (late Anisian–early Ladinian, UNESCO World Heritage Site Monte San Giorgio, Ticino, Switzerland; (2 Macrocnemus obristi from the Prosanto Formation (early Ladinian of the Ducan area, Grisons, Switzerland; and (3 Macrocnemus fuyuanensis from the Falang Formation (Ladinian, Yunnan Province, southern China. Recently a new specimen, PIMUZ T 1559, from the upper Besano Formation at Meride, Ticino, Switzerland, was prepared, revealing a disarticulated skeleton which includes most of the cranium and lower jaw, pre-caudal vertebral column and ribs, the forelimbs, and girdle elements. Unambiguously assignable to the genus Macrocnemus, it evinces particularly gracile elongated cervical ribs, as well as a humerus/radius ratio that is comparable only to that of M. fuyuanensis from southern China. Based on this feature we tentatively recognize the new specimen as M. aff. fuyuanensis from Europe. The position and exquisite preservation of the clavicle and interclavicle in this specimen allows a revision of the shoulder girdle of Macrocnemus when articulated, which also has implications for closely related protorosaurian taxa, such as the long-necked Tanystropheus. Furthermore, differences in the shape and morphology of the interclavicle including pointed wing-like lateral processes and a short, fusiform caudal process represent rare

  8. Absolute Plate Motion Control Since the Triassic from the Cocos Slab and its Associated Subduction Record in Mexico

    Science.gov (United States)

    Boschman, L.; Van Hinsbergen, D. J. J.; Langereis, C. G.; Molina-Garza, R. S.; Kimbrough, D. L.; Spakman, W.

    2017-12-01

    A positive wave speed anomaly interpreted as the Cocos slab stretches from the uppermost mantle at the Middle America trench in the west, to the lowermost mantle below the Atlantic in the east. The length and continuity of this slab indicates long-lived, uninterrupted eastward subduction of the attached Cocos Plate and its predecessor, the Farallon Plate. The geological record of Mexico contains Triassic to present day evidence of subduction, of which the post-Late Cretaceous phase is of continental margin-style. Interpretations of the pre-Upper Cretaceous subduction-related rock assemblages are under debate, and vary from far-travelled exotic intra-oceanic island arc character to in-situ extended continental margin origin. We present new paleomagnetic data that show that Triassic, Jurassic and Cretaceous subduction-related rocks from the Vizcaíno Peninsula and the Guerrero terrane have a paleolatitudinal plate motion history that is equal to that of the North American continent. This suggests that these rock assemblages were part of the overriding plate and were perhaps only separated from the North American continent by temporal fore- or back-arc spreading. The entire Triassic-present day subduction record, and hence, reconstructed trench location, can therefore be linked to the Cocos slab, which provides control on longitudinal plate motion of North America since the time of Pangea. Compared to the latest state of the art mantle frames, in which longitudes are essentially unconstrained for pre-Cretaceous times, our reconstructed absolute position of North America requires a significant westward longitudinal shift for Mesozoic times.

  9. Evidence for Different Disk Mass Distributions between Early- and Late-type Be Stars in the BeSOS Survey

    Energy Technology Data Exchange (ETDEWEB)

    Arcos, C.; Kanaan, S.; Curé, M. [Instituto de Física y Astronomía, Facultad de Ciencias, Universidad de Valparaíso. Av. Gran Bretana 1111, Valparaíso (Chile); Jones, C. E.; Sigut, T. A. A. [Department of Physics and Astronomy, The University of Western Ontario, London, ON N6A 3K7 (Canada)

    2017-06-10

    The circumstellar disk density distributions for a sample of 63 Be southern stars from the BeSOS survey were found by modeling their H α emission line profiles. These disk densities were used to compute disk masses and disk angular momenta for the sample. Average values for the disk mass are 3.4 × 10{sup −9} and 9.5 × 10{sup −10} M {sub ⋆} for early (B0–B3) and late (B4–B9) spectral types, respectively. We also find that the range of disk angular momentum relative to the star is (150–200) J {sub ⋆}/ M {sub ⋆} and (100–150) J {sub ⋆}/ M {sub ⋆}, again for early- and late-type Be stars, respectively. The distributions of the disk mass and disk angular momentum are different between early- and late-type Be stars at a 1% level of significance. Finally, we construct the disk mass distribution for the BeSOS sample as a function of spectral type and compare it to the predictions of stellar evolutionary models with rapid rotation. The observed disk masses are typically larger than the theoretical predictions, although the observed spread in disk masses is typically large.

  10. The Maximum Mass Solar Nebula and the early formation of planets

    Science.gov (United States)

    Nixon, C. J.; King, A. R.; Pringle, J. E.

    2018-03-01

    Current planet formation theories provide successful frameworks with which to interpret the array of new observational data in this field. However, each of the two main theories (core accretion, gravitational instability) is unable to explain some key aspects. In many planet formation calculations, it is usual to treat the initial properties of the planet forming disc (mass, radius, etc.) as free parameters. In this paper, we stress the importance of setting the formation of planet forming discs within the context of the formation of the central stars. By exploring the early stages of disc formation, we introduce the concept of the Maximum Mass Solar Nebula (MMSN), as opposed to the oft-used Minimum Mass Solar Nebula (here mmsn). It is evident that almost all protoplanetary discs start their evolution in a strongly self-gravitating state. In agreement with almost all previous work in this area, we conclude that on the scales relevant to planet formation these discs are not gravitationally unstable to gas fragmentation, but instead form strong, transient spiral arms. These spiral arms can act as efficient dust traps allowing the accumulation and subsequent fragmentation of the dust (but not the gas). This phase is likely to populate the disc with relatively large planetesimals on short timescales while the disc is still veiled by a dusty-gas envelope. Crucially, the early formation of large planetesimals overcomes the main barriers remaining within the core accretion model. A prediction of this picture is that essentially all observable protoplanetary discs are already planet hosting.

  11. A preliminary report on coprolites from the Late Triassic part of the Kap Stewart Formation, Jameson Land, East Greenland

    DEFF Research Database (Denmark)

    Milàn, Jesper; Clemmensen, Lars B; Adolphsen, Jan Schulz

    2012-01-01

    The basal part of the Triassic-Jurassic (Rhaetian-Sinemurian) Kap Stewart Formation, exposed at Jameson Land, East Greenland, yields an extensive coprolite collection from black, parallel-laminated mudstone (“paper shale”), representing an open lacustrine system. Preliminary investigations show t...

  12. Chronology of Fluctuating Sea Levels since the Triassic

    Science.gov (United States)

    Haq, Bilal U.; Hardenbol, Jan; Vail, Peter R.

    1987-03-01

    Advances in sequence stratigraphy and the development of depositional models have helped explain the origin of genetically related sedimentary packages during sea level cycles. These concepts have provided the basis for the recognition of sea level events in subsurface data and in outcrops of marine sediments around the world. Knowledge of these events has led to a new generation of Mesozoic and Cenozoic global cycle charts that chronicle the history of sea level fluctuations during the past 250 million years in greater detail than was possible from seismic-stratigraphic data alone. An effort has been made to develop a realistic and accurate time scale and widely applicable chronostratigraphy and to integrate depositional sequences documented in public domain outcrop sections from various basins with this chronostratigraphic frame-work. A description of this approach and an account of the results, illustrated by sea level cycle charts of the Cenozoic, Cretaceous, Jurassic, and Triassic intervals, are presented.

  13. Picrite "Intelligence" from the Middle-Late Triassic Stikine arc: Composition of mantle wedge asthenosphere

    Science.gov (United States)

    Milidragovic, D.; Zagorevski, A.; Weis, D.; Joyce, N.; Chapman, J. B.

    2018-05-01

    Primitive, near-primary arc magmas occur as a volumetrically minor ≤100 m thick unit in the Canadian Cordillera of northwestern British Columbia, Canada. These primitive magmas formed an olivine-phyric, picritic tuff near the base of the Middle-Late Triassic Stuhini Group of the Stikine Terrane (Stikinia). A new 40Ar/39Ar age on hornblende from a cross-cutting basaltic dyke constrains the tuff to be older than 221 ± 2 Ma. An 87Sr/86Sr isochron of texturally-unmodified tuff samples yields 212 ± 25 Ma age, which is interpreted to represent syn-depositional equilibration with sea-water. Parental trace element magma composition of the picritic tuff is strongly depleted in most incompatible trace elements relative to MORB and implies a highly depleted ambient arc mantle. High-precision trace element and Hf-Nd-Pb isotopic analyses indicate an origin by mixing of a melt of depleted ambient asthenosphere with ≤2% of subducted sediment melt. Metasomatic addition of non-conservative incompatible elements through melting of subducted Panthalassa Ocean floor sediments accounts for the arc signature of the Stuhini Group picritic tuff, enrichment of light rare earth elements (LREE) relative to heavy rare earth elements (HREE) and high field strength elements (HFSE), and anomalous enrichment in Pb. The inferred Panthalassan sediments are similar in composition to the Neogene-Quaternary sediments of the modern northern Cascadia Basin. The initial Hf isotopic composition of the picritic tuff closely approximates that of the ambient Middle-Late Triassic asthenosphere beneath Stikinia and is notably less radiogenic than the age-corrected Hf isotopic composition of the Depleted (MORB) Mantle reservoir (DM or DMM). This suggests that the ambient asthenospheric mantle end-member experienced melt depletion (F ≤ 0.05) a short time before picrite petrogenesis. The mantle end-member in the source of the Stuhini Group picritic tuff is isotopically similar to the mantle source of

  14. Preliminary study of the uranium potential of the Triassic Sanford basin and Colon cross structure, North Carolina

    International Nuclear Information System (INIS)

    Lee, C.H.

    1978-01-01

    A preliminary geologic investigation was conducted to determine if Triassic sedimentary rocks of the Sanford basin and Colon cross structure in North Carolina are favorable hosts for uranium deposits. Rocks of adjacent Carolina slate belt were also examined as a potential source of uranium. On the basis of favorability criteria for sandstone-type uranium deposits, and geologic and geophysical investigations of the study area, the most favorable sites for further investigation are (1) at the contacts between the Pekin and Cumnock and between the Pekin and Sanford Formations near the Colon cross structure and (2) at the base of the Jonesboro fault, which lies below the Sanford Formation, northwest of Sanford. The highly weathered granites southeast of the Jonesboro fault were a source of the detritus deposited on the cross structure and may have been a primary source of uranium. Uranium leached from the coarse sediment (Pekin Formation) of the cross structure may have been transported downdip and may have been precipitated by the carbonaceous shales of the Cumnock Formation on the western side of the cross structure or at the Pekin-Sanford contact to the east. The Jonesboro fault may provide an impermeable barrier to ground-water migration in the metamorphosed basement rocks below the Triassic sediments. Such a barrier would constitute a favorable site for the precipitation and retention of uranium. Scintillometer surveys and laboratory analyses indicate no anomalous surface radioactivity in the study area. However, deep surface weathering may have caused the uranium to be leached from the exposed rocks and redeposited at depth. Geologic investigations show that conditions which have proven favorable for deposition of uranium in other areas are present in the Triassic rocks of the Sanford basin and Colon cross structure. However, because of deep surface weathering, further subsurface studies are necessary to confirm the favorability of the rocks as hosts for uranium

  15. Permo-Triassic vertebrate extinctions: A program

    Science.gov (United States)

    Olson, E. C.

    1988-01-01

    Since the time of the Authors' study on this subject, a great deal of new information has become available. Concepts of the nature of extinctions have changed materially. The Authors' conclusion that a catastrophic event was not responsible for the extinction of vertebrates has modified to the extent that hypotheses involving either the impact of a massive extra-terrestrial body or volcanism provide plausible but not currently fully testable hypotheses. Stated changes resulted in a rapid decrease in organic diversity, as the ratio of origins of taxa to extinctions shifted from strongly positive to negative, with momentary equilibrium being reached at about the Permo-Triassic boundary. The proximate causes of the changes in the terrestrial biota appear to lie in two primary factors: (1) strong climatic changes (global mean temperatures, temperature ranges, humidity) and (2) susceptibility of the dominant vertebrates (large dicynodonts) and the glossopteris flora to disruption of the equlibrium of the world ecosystem. The following proximate causes have been proposed: (1) rhythmic fluctuations in solar radiation, (2) tectonic events as Pangea assembled, altering land-ocean relationships, patterns of wind and water circulation and continental physiography, (3) volcanism, and (4) changes subsequent to impacts of one or more massive extra terrestrial objects, bodies or comets. These hypotheses are discussed.

  16. Two types of bone necrosis in the Middle Triassic Pistosaurus longaevus bones: the results of integrated studies

    Science.gov (United States)

    Surmik, Dawid; Rothschild, Bruce M.; Dulski, Mateusz; Janiszewska, Katarzyna

    2017-07-01

    Avascular necrosis, diagnosed on the basis of either a specific pathological modification of the articular surfaces of bone or its radiologic appearance in vertebral centra, has been recognized in many Mesozoic marine reptiles as well as in present-day marine mammals. Its presence in the zoological and paleontologic record is usually associated with decompression syndrome, a disease that affects secondarily aquatic vertebrates that could dive. Bone necrosis can also be caused by infectious processes, but it differs in appearance from decompression syndrome-associated aseptic necrosis. Herein, we report evidence of septic necrosis in the proximal articular surface of the femur of a marine reptile, Pistosaurus longaevus, from the Middle Triassic of Poland and Germany. This is the oldest recognition of septic necrosis associated with septic arthritis in the fossil record so far, and the mineralogical composition of pathologically altered bone is described herein in detail. The occurrence of septic necrosis is contrasted with decompression syndrome-associated avascular necrosis, also described in Pistosaurus longaevus bone from Middle Triassic of Germany.

  17. Early-type galaxies: mass-size relation at z ˜ 1.3 for different environments

    Science.gov (United States)

    Raichoor, A.; Mei, S.; Stanford, S. A.; Holden, B. P.; Nakata, F.; Rosati, P.; Shankar, F.; Tanaka, M.; Ford, H.; Huertas-Company, M.; Illingworth, G.; Kodama, T.; Postman, M.; Rettura, A.; Blakeslee, J. P.; Demarco, R.; Jee, M. J.; White, R. L.

    2011-12-01

    We combine multi-wavelength data of the Lynx superstructure and GOODS/CDF-S to build a sample of 75 visually selected early-type galaxies (ETGs), spanning different environments (cluster/group/field) at z ˜ 1.3. By estimating their mass, age (SED fitting, with a careful attention to the stellar population model used) and size, we are able to probe the dependence on the environment of the mass-size relation. We find that, for ETGs with 10^{10} < M / M_⊙ < 10^{11.5}, (1) the mass-size relation in the field did not evolve overall from z ˜ 1.3 to present; (2) the mass-size relation in cluster/group environments at z ˜ 1.3 lies at smaller sizes than the local mass-size relation (R_{e,z ˜ 1.3}/R_{e,z = 0} ˜ 0.6-0.8).

  18. A new marine reptile from the Triassic of China, with a highly specialized feeding adaptation

    Science.gov (United States)

    Cheng, Long; Chen, Xiao-Hong; Shang, Qing-Hua; Wu, Xiao-Chun

    2014-03-01

    The Luoping fauna (Anisian, Middle Triassic) is probably the oldest of Triassic faunas in Guizhou-Yunnan area, China. The reptilian assemblage is comprised of ichthyosaurs, a number of sauropterygians (pachypleurosaur-like forms), saurosphargids, protorosaurs, and archosauriforms. Here, we report on a peculiar reptile, newly found in this fauna. Its dentition is fence or comb-like and bears more than 175 pleurodont teeth in each ramus of the upper and lower jaws, tooth crown is needle-like distally and blade-shaped proximally; its rostrum strongly bends downward and the anterior end of its mandible expands both dorsally and ventrally to form a shovel-headed structure; and its ungual phalanges are hoof-shaped. The specializations of the jaws and dentition indicate that the reptile may have been adapted to a way of bottom-filter feeding in water. It is obvious that such delicate teeth are not strong enough to catch prey, but were probably used as a barrier to filter microorganisms or benthic invertebrates such as sea worms. These were collected by the specialized jaws, which may have functioned as a shovel or pushdozer (the mandible) and a grasper or scratcher (the rostrum). Our preliminary analysis suggests that the new reptile might be more closely related to the Sauropterygia than to other marine reptiles.

  19. A new marine reptile from the Triassic of China, with a highly specialized feeding adaptation.

    Science.gov (United States)

    Cheng, Long; Chen, Xiao-Hong; Shang, Qing-Hua; Wu, Xiao-Chun

    2014-03-01

    The Luoping fauna (Anisian, Middle Triassic) is probably the oldest of Triassic faunas in Guizhou-Yunnan area, China. The reptilian assemblage is comprised of ichthyosaurs, a number of sauropterygians (pachypleurosaur-like forms), saurosphargids, protorosaurs, and archosauriforms. Here, we report on a peculiar reptile, newly found in this fauna. Its dentition is fence or comb-like and bears more than 175 pleurodont teeth in each ramus of the upper and lower jaws, tooth crown is needle-like distally and blade-shaped proximally; its rostrum strongly bends downward and the anterior end of its mandible expands both dorsally and ventrally to form a shovel-headed structure; and its ungual phalanges are hoof-shaped. The specializations of the jaws and dentition indicate that the reptile may have been adapted to a way of bottom-filter feeding in water. It is obvious that such delicate teeth are not strong enough to catch prey, but were probably used as a barrier to filter microorganisms or benthic invertebrates such as sea worms. These were collected by the specialized jaws, which may have functioned as a shovel or pushdozer (the mandible) and a grasper or scratcher (the rostrum). Our preliminary analysis suggests that the new reptile might be more closely related to the Sauropterygia than to other marine reptiles.

  20. Multiple populations within globular clusters in Early-type galaxies Exploring their effect on stellar initial mass function estimates

    Science.gov (United States)

    Chantereau, W.; Usher, C.; Bastian, N.

    2018-05-01

    It is now well-established that most (if not all) ancient globular clusters host multiple populations, that are characterised by distinct chemical features such as helium abundance variations along with N-C and Na-O anti-correlations, at fixed [Fe/H]. These very distinct chemical features are similar to what is found in the centres of the massive early-type galaxies and may influence measurements of the global properties of the galaxies. Additionally, recent results have suggested that M/L variations found in the centres of massive early-type galaxies might be due to a bottom-heavy stellar initial mass function. We present an analysis of the effects of globular cluster-like multiple populations on the integrated properties of early-type galaxies. In particular, we focus on spectral features in the integrated optical spectrum and the global mass-to-light ratio that have been used to infer variations in the stellar initial mass function. To achieve this we develop appropriate stellar population synthesis models and take into account, for the first time, an initial-final mass relation which takes into consideration a varying He abundance. We conclude that while the multiple populations may be present in massive early-type galaxies, they are likely not responsible for the observed variations in the mass-to-light ratio and IMF sensitive line strengths. Finally, we estimate the fraction of stars with multiple populations chemistry that come from disrupted globular clusters within massive ellipticals and find that they may explain some of the observed chemical patterns in the centres of these galaxies.

  1. Determinants of Growth, Adiposity and Bone Mass in Early Life : The Generation R Study

    NARCIS (Netherlands)

    D.H.M. Heppe (Denise)

    2016-01-01

    markdownabstractAbstract Environmental influences during fetal life and early infancy have been suggested to influence body composition throughout the life-course. Especially poor fetal nutrition and fetal growth restriction have been designated important risk factors for gaining high fat mass

  2. VALLESAURUS CENENSIS WILD, 1991, A DREPANOSAURID (REPTILIA DIAPSIDA FROM THE LATE TRIASSIC OF NORTHERN ITALY

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    SILVIO RENESTO

    2006-03-01

    Full Text Available Vallesaurus cenensis, a small drepanosaurid reptile from the Norian (Late Triassic beds of the Calcare di Zorzino (Zorzino Limestone is described. The holotype and only known specimen represents the first drepanosaurid in which the skull is preserved articulated with the postcranial skeleton. The study of Vallesaurus anatomy confirms previous data about arboreal adaptation in all known drepanosaurids and permits more firm hypotheses about the phylogenetic relationships of the Drepanosauridae to be proposed.

  3. The elemental geochemistry of Lower Triassic shallow-marine carbonates from central Saudi Arabia: Implications for redox conditions in the immediate aftermath of the latest Permian mass extinction

    Science.gov (United States)

    Eltom, Hassan A.; Abdullatif, Osman M.; Babalola, Lamidi O.

    2018-03-01

    The southern margin of the Tethys Ocean was occupied by a broad, shallow continental shelf during the Permian-Triassic boundary interval, with the area of present-day Saudi Arabia located from 10° to 30° south of the paleo-equator. The strata deposited in modern Saudi Arabia in the aftermath of the latest Permian mass extinction (LPME) are dominated by oolitic microbialite limestone (OML), which are overlain by skeletal oolitic limestones (SOL) capped by dolostones and dolomitic limestones (DDL). This succession reflects changes in depositional setting, which can be potentially tied to redox conditions using redox sensitive trace elements and rare earth elements (REEs). Statistical analyses reveals that trace elements and REEs are associated with detrital material, and possibly with diagenetic minerals as well. Proxies such as the Y/Ho, Pr/Pr*, Smn/Ybn, Lan/Smn and Lan/Ybn ratios indicate that REEs do not record a seawater-like pattern, and cannot be used as redox indicator. The presence of a normal marine fauna implies oxic conditions during deposition of the DDL and SOL units. However, the OML unit, which represents the immediate aftermath of LPME, lacks both a normal marine fauna and reliable geochemical signals, making it difficult to infer redox conditions in the depositional environment. Similar to published data from sections that reflect shallow marine condition in the LPME of the Tethys Ocean, chemical index of alteration values are consistently high throughout the study succession, suggesting globally intense chemical weathering in the aftermath of the LPME. As a result, geochemical redox proxies in shallow marine carbonates of the Tethys Ocean are likely to be contaminated by detrital material that have been generated by chemical weathering, and thus, other methods are required to determine depositional redox conditions.

  4. Carbon cycling and mass extinctions: the Permo-Triassic of the Arabian Margin.

    OpenAIRE

    Clarkson, Matthew Oliver

    2014-01-01

    The end-Permian extinction at 252 Ma is widely regarded as the most severe of the Phanerozoic mass-extinctions and enabled the evolution of the modern carbon cycle and ecosystem structure. The cause of the extinction is still debated but the synergistic pressures of global climate change, such as anoxia and ocean acidification, were clearly important. The extinction occurred in two phases and is marked by a uniquely protracted recovery period of ~ 5 Myrs where diversity fails to reach pre-ext...

  5. Geochemistry of the triassic-Jurassic alpine continental deposits: origin and geodynamic implications

    International Nuclear Information System (INIS)

    Poinssot, Ch.; Goffe, B.; Toulhoat, P.

    1997-01-01

    Mid-Triassic to mid-Jurassic Alpine continental deposits are known all along the former Brianconnais peninsula. They constitutes small karstic pockets on the thick Triassic calcareous series and their chemistry evolves between bauxites s.s. and aluminous argilites. Most of them were deeply buried during the Alpine orogenesis as recorded by HP-LT metamorphism. Only the deposits of the Pre-Alps were submitted to lower PT conditions (diagenesis-anchizone boundary) during their incorporation in the thrust wedge of the 'Prealpes Medianes'. These formations are known for containing traces of light elements (Li, F) and heavy elements (Zn, REE...). In order to understand the possible origin of these elements, we studied the geochemistry (major and trace elements) of two representative deposits, one in Vanoise which underwent a HP-LT metamorphism, the other one in the Pre-Alps, which was only submitted to diagenesis. Trace elements patterns allow us to preclude an autochthonous origin for these formations as well as the intervention of metasomatism, and demonstrate a granitic origin. Moreover, discrimination diagrams for granites indicate an obvious alkaline granitic origin for these deposits. In the framework of the Alpine palaeogeography, we then discuss the possible granitic sources. Two main sources can be invoked: either a Brianconnais s.s. formation (crystalline or sediments), which supposes a more intense erosion as classically admitted, or more distant sources such as the Corso-Sardinian alkaline acid-rocks, which supposes a complex palaeo-hydrography. This confirms the sedimentary origin of the light elements in these rocks and precludes the intervention of light elements-rich hydrothermal fluids migrating through Alpine metamorphic units. (author)

  6. The archaeology of early medieval violence: the mass grave at Budeč, Czech Republic

    Czech Academy of Sciences Publication Activity Database

    Štefan, I.; Stránská, Petra; Vondrová, H.

    2016-01-01

    Roč. 90, č. 351 (2016), s. 759-776 ISSN 0003-598X Institutional support: RVO:67985912 Keywords : Czech Republic * Bohemia * Budeč * tenth-eleventh centuries * early medieval * mass grave * violence * warfare Subject RIV: AC - Archeology, Anthropology, Ethnology Impact factor: 1.536, year: 2016

  7. The first 50Myr of dinosaur evolution: macroevolutionary pattern and morphological disparity.

    Science.gov (United States)

    Brusatte, Stephen L; Benton, Michael J; Ruta, Marcello; Lloyd, Graeme T

    2008-12-23

    The evolutionary radiation of dinosaurs in the Late Triassic and Early Jurassic was a pivotal event in the Earth's history but is poorly understood, as previous studies have focused on vague driving mechanisms and have not untangled different macroevolutionary components (origination, diversity, abundance and disparity). We calculate the morphological disparity (morphospace occupation) of dinosaurs throughout the Late Triassic and Early Jurassic and present new measures of taxonomic diversity. Crurotarsan archosaurs, the primary dinosaur 'competitors', were significantly more disparate than dinosaurs throughout the Triassic, but underwent a devastating extinction at the Triassic-Jurassic boundary. However, dinosaur disparity showed only a slight non-significant increase after this event, arguing against the hypothesis of ecological release-driven morphospace expansion in the Early Jurassic. Instead, the main jump in dinosaur disparity occurred between the Carnian and Norian stages of the Triassic. Conversely, dinosaur diversity shows a steady increase over this time, and measures of diversification and faunal abundance indicate that the Early Jurassic was a key episode in dinosaur evolution. Thus, different aspects of the dinosaur radiation (diversity, disparity and abundance) were decoupled, and the overall macroevolutionary pattern of the first 50Myr of dinosaur evolution is more complex than often considered.

  8. THALAMID SPONGES FROM THE UPPER TRIASSIC (NORIAN-RHAETIAN NAYBAND FORMATION NEAR WALI-ABAD, SE ABADEH, CENTRAL IRAN(CONTRIBUTION TO TRIASSIC PALEONTOLOGY OF IRAN 4

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    BABA SENOWBARI-DARYAN

    1999-03-01

    Full Text Available An association of thalamid sponges including Amblysiphonella, Nevadathalamia, Stylothalamia, and Neoguadalupia, with the hexactinellid Casearia is described from the Upper Triassic (Norian-Rhaetian Nayband Formation of Hambast-Mts. near the small town of Wali-Abad (Abadeh region, central Iran. These Norian-Rhaetian (Neoguadalupia, Amblysiphonella, Nevadathalamia and Liassic (Stylothalamia columnaris Le Maitre thalamid sponge assemblages are exceptional associations, not previously reported from Iran nor from other localities in the world. The microfacies and organism associations in the sponge-bearing carbonates is discussed. Differences of macro- and microfaunal composition, as well as flora, support the recognition of Kristan-Tollmann et al. to classify the Nayband Formation in the Abadeh region as of a distinct unit, termed the "Wali-Abad-Faziesregion". Following thalamid sponge species are described as new: Nevadathalamia waliabadensis n. sp. and Stylothalamia hambastensis n. sp. 

  9. Triassic North American paleodrainage networks and sediment dispersal of the Chinle Formation: A quantitative approach utilizing detrital zircons

    Science.gov (United States)

    Blum, M. D.; Umbarger, K.

    2017-12-01

    The Triassic Chinle Formation is a fluvial succession deposited in a backarc setting across the present-day Colorado Plateau of the southwestern United States. Existing studies have proposed various mechanisms responsible for the unique stratigraphic architecture and depositional sequences of the Chinle. However, these studies lack necessary age control to correlate stratigraphic patterns with contemporaneous mechanisms. This study will collect new samples for detrital zircon analysis, as well as upgrade existing samples (to n=300) from Dickinson and Gehrels (2008), to improve the resolution of Triassic sediment provenance from source-to-sink. The improved dataset allows appraisal of the multiple provenance terranes that contributed to the Chinle depositional system to delineate and reconstruct paleodrainage patterns. The additional samples will be collected systematically from the base of the Chinle, and vertically throughout the section to capture a regional story of how the continental scale drainage reorganized through time. U-Pb ages of detrital zircons will be utilized to provide quantitative fingerprinting information to constrain interpretations for the origin and transport history of the Chinle fluvial succession in time and space.

  10. Je ne menge poinct de porcq: Orlando di Lasso’s Early Parody Mass

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    Klemen Grabnar

    2013-07-01

    Full Text Available One of the most remarkable 16th-century composers was Orlando di Lasso (1530/1532–1594, perhaps the most prominent musician of his time. In the second half of the 16th and the early 17th centuries, his works were widespread across Europe, especially in its central and western part. Some of the surviving contemporary sources of Lasso’s music – both printed and manuscript – are preserved in Slovenian libraries and archives. Among them are two incompletely preserved manuscripts, dating from ca. 1600, today kept at the National and University Library in Ljubljana (Ms 232 and Ms 285. They both contain Lasso’s Missa super Je ne menge poinct de porcq, first published in Lasso’s second book of Masses, Quinque missae, by Claudio Merulo in 1570. Interestingly, this early Munich-period Mass is based on a Parisian chanson by Claudin de Sermisy (ca. 1490–1562, whose content is unequivocally scatological. It was quite common for 16th-century composers to base their settings of the Mass Ordinary on pre-existent polyphonic compositions, whether sacred or secular. One of the rare descriptions of this technique – widely known as parody technique – is given in Pietro Cerone’s El melopeo y maestro (1613. It can serve as the basis for an elementary analysis of how Lasso employed Sermisy’s chanson Je ne menge point de porc. The analysis shows that Lasso, for various reasons, does not always follow the established practices as they can be observed from Cerone’s treatise. Nevertheless he demonstrates significant skill in utilising borrowed material to compose his parody Mass. Lasso’s Missa super Je ne menge poinct de porcq is particularly intriguing due to its use of Sermisy’s chanson; what could be the reason for Lasso’s decision to use Sermisy’s profoundly secular chanson in order to compose a Mass setting? Although the answer can be multifaceted (such as an intention to establish a special textual relation between the Mass and the

  11. Triassic marine reptiles gave birth to live young.

    Science.gov (United States)

    Cheng, Yen-Nien; Wu, Xiao-Chun; Ji, Qiang

    2004-11-18

    Sauropterygians form the largest and most diverse group of ancient marine reptiles that lived throughout nearly the entire Mesozoic era (from 250 to 65 million years ago). Although thousands of specimens of this group have been collected around the world since the description of the first plesiosaur in 1821 (ref. 3), no direct evidence has been found to determine whether any sauropterygians came on shore to lay eggs (oviparity) like sea turtles, or gave birth in the water to live young (viviparity) as ichthyosaurs and mosasauroids (marine lizards) did. Viviparity has been proposed for plesiosaur, pachypleurosaur and nothosaur sauropterygians, but until now no concrete evidence has been advanced. Here we report two gravid specimens of Keichousaurus hui Young from the Middle Triassic of China. These exquisitely preserved specimens not only provide the first unequivocal evidence of reproductive mode and sexual dimorphism in sauropterygians, but also indicate that viviparity could have been expedited by the evolution of a movable pelvis in pachypleurosaurs. By extension, this has implications for the reproductive pattern of other sauropterygians and Mesozoic marine reptiles that possessed a movable pelvis.

  12. Sedimentary evolution of the Mesozoic continental redbeds using geochemical and mineralogical tools: the case of Upper Triassic to Lowermost Jurassic Monte di Gioiosa mudrocks (Sicily, southern Italy)

    Science.gov (United States)

    Perri, Francesco; Critelli, Salvatore; Mongelli, Giovanni; Cullers, Robert L.

    2011-10-01

    The continental redbeds from the Internal Domains of the central-western Mediterranean Chains have an important role in the palaeogeographic and palaeotectonic reconstructions of the Alpine circum-Mediterranean orogen evolution since these redbeds mark the Triassic-Jurassic rift-valley stage of Tethyan rifting. The composition and the sedimentary evolution of the Middle Triassic to Lowermost Jurassic continental redbeds of the San Marco d'Alunzio Unit (Peloritani Mountains, Southern Italy), based on mineralogical and chemical analyses, suggests that the studied mudrock sediments share common features with continental redbeds that constitute the Internal Domains of the Alpine Mediterranean Chains. Phyllosilicates are the main components in the mudrocks. The 10 Å-minerals (illite and micas), the I-S mixed layers, and kaolinite are the most abundant phyllosilicates. The amount of illitic layers in I-S mixed layers coupled with the illite crystallinity values (IC) are typical of high degree of diagenesis, corresponding to a lithostatic/tectonic loading of about 4-5 km. The mineralogical assemblage coupled with the A-CN-K plot suggest post-depositional K-enrichments. Palaeoweathering proxies (PIA and CIW) record intense weathering at the source area. Further, the studied sediments are affected by reworking and recycling processes and, as consequence, it is likely these proxies monitor cumulative effect of weathering. The climate in the early Jurassic favoured recycling and weathering occurred under hot, episodically humid climate with a prolonged dry season. The source-area is the low-grade Paleozoic metasedimentary basement. Mafic supply is minor but not negligible as suggested by provenance proxies.

  13. Early peroxisome proliferator-activated receptor gamma regulated genes involved in expansion of pancreatic beta cell mass

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    Vivas Yurena

    2011-12-01

    Full Text Available Abstract Background The progression towards type 2 diabetes depends on the allostatic response of pancreatic beta cells to synthesise and secrete enough insulin to compensate for insulin resistance. The endocrine pancreas is a plastic tissue able to expand or regress in response to the requirements imposed by physiological and pathophysiological states associated to insulin resistance such as pregnancy, obesity or ageing, but the mechanisms mediating beta cell mass expansion in these scenarios are not well defined. We have recently shown that ob/ob mice with genetic ablation of PPARγ2, a mouse model known as the POKO mouse failed to expand its beta cell mass. This phenotype contrasted with the appropriate expansion of the beta cell mass observed in their obese littermate ob/ob mice. Thus, comparison of these models islets particularly at early ages could provide some new insights on early PPARγ dependent transcriptional responses involved in the process of beta cell mass expansion Results Here we have investigated PPARγ dependent transcriptional responses occurring during the early stages of beta cell adaptation to insulin resistance in wild type, ob/ob, PPARγ2 KO and POKO mice. We have identified genes known to regulate both the rate of proliferation and the survival signals of beta cells. Moreover we have also identified new pathways induced in ob/ob islets that remained unchanged in POKO islets, suggesting an important role for PPARγ in maintenance/activation of mechanisms essential for the continued function of the beta cell. Conclusions Our data suggest that the expansion of beta cell mass observed in ob/ob islets is associated with the activation of an immune response that fails to occur in POKO islets. We have also indentified other PPARγ dependent differentially regulated pathways including cholesterol biosynthesis, apoptosis through TGF-β signaling and decreased oxidative phosphorylation.

  14. A new chronology for the late Triassic to early Jurassic. Geologica Ultraiectina (323)

    NARCIS (Netherlands)

    Deenen, M.H.L.

    2010-01-01

    Throughout geological history of life, a small number of mass extinctions have forever changed the path of evolution. The two main mechanisms that account for these dramatic events are asteroid impacts and massive volcanic eruptions of Large Igneous Provinces (LIP’s). The interplay between

  15. Permo-Triassic arc-like granitoids along the northern Lancangjiang zone, eastern Tibet: Age, geochemistry, Sr-Nd-Hf isotopes, and tectonic implications

    Science.gov (United States)

    Wang, Xinyu; Wang, Shifeng; Wang, Chao; Tang, Wenkun

    2018-05-01

    Large volumes of Permo-Triassic granitoids are exposed along the Northern Lancangjiang zone, eastern Tibet, and these rocks provide insights into the tectonic evolution of the Paleo-Tethys Ocean. We conducted detailed geological fieldwork and geochemical analysis of the Xiaochangdu and Kagong plutons that crop out along the Northern Lancangjiang magmatic belt. Zircon U-Pb data constrain the emplacement of the Xiaochangdu quartz diotites to between 263 and 257 Ma, and the Kagong granites and diorites to between 234 and 232 Ma. The Xiaochangdu quartz diorites are enriched in light rare earth (LREE) and large ion lithophile elements (LILE), depleted in high field strength elements (HFSE), have low (87Sr/86Sr)i ratios, and near-positive εNd(t) (-0.26 to 1.58) and εHf(t) (0.68-8.83) values, similar to typical subduction- related mantle-derived arc magmas. They are also characterized by high Al2O3 concentrations and low Nb/U (3.48-7.59) and Ce/Pb (3.22-4.86) ratios, indicating that their mantle source was modified by subducted pelagic sediments; Coeval granites and diorites from the Kagong pluton exhibit low A/CNK values, high LREE/HREE (heavy rare earth element) ratios, enrichment in LILE, and depletion in HFSE, also characteristic of typical arc magmas. Their variable SiO2 contents (57%- 75%), (87Sr/86Sr)i ratios, and εNd(t) (1.02-4.49) and εHf(t) (2.52-6.93) values, and relatively high zircon saturation temperatures (721-827 °C), suggest underplating of mantle-derived mafic melts beneath the lower crust. Their magmatic evolution can be explained using a MASH model. In combination with regional geological studies, our geochemical and geochronological results suggest that the late Permian Xiaochangdu and Late Triassic Kagong arc-like granitoids represent a section of a Permo-Triassic magmatic arc that was associated with the eastward subduction of the Paleo-Tethys oceanic slab beneath the Northern Qiangtang-Changdu terrane. Combined with other geological evidence

  16. Re-Os Geochronology Pins Age and Os Isotope Composition of Middle Triassic Black Shales and Seawater, Barents Sea and Spitsbergen (Svalbard)

    Science.gov (United States)

    Xu, G.; Hannah, J. L.; Bingen, B.; Stein, H. J.; Yang, G.; Zimmerman, A.; Weitschat, W.; Weiss, H. M.

    2008-12-01

    Absolute age control throughout the Triassic is extraordinarily sparse. Two "golden spikes" have been added recently (http://www.stratigraphy.org/cheu.pdf) within the otherwise unconstrained Triassic, but ages of stage boundaries remain controversial. Here we report two Re-Os isochrons for Anisian (Middle Triassic) black shales from outcrop in western Svalbard and drill core from the Svalis Dome about 600 km to the SE in the Barents Sea. Black shales of the Blanknuten Member, Botneheia Formation, from the type section at Botneheia, western Spitsbergen (Svalbard), have total organic carbon (TOC) contents of 2.6 to 6.0 wt%. Rock-Eval data suggest moderately mature (Tmax = 440-450° C) Type II-III kerogens (Hydrogen Index (HI) = 232-311 mg HC/g TOC). Re-Os data yield a well-constrained Model 3 age of 241 Ma and initial 187Os/188Os (Osi) of 0.83 (MSWD = 16, n = 6). Samples of the possibly correlative Steinkobbe Formation from IKU core hole 7323/07-U-04 into the Svalis Dome in the Barents Sea (at about 73°30'N, 23°15'E) have TOC contents of 1.4 to 2.4%. Rock-Eval data suggest immature (Tmax = 410-430°) Type II-III kerogens (HI = 246-294 mg HC/g TOC). Re-Os data yield a precise Model 1 age of 239 Ma and Osi of 0.776 (MSWD = 0.2, n = 5). The sampled section of Blanknuten shale underlies a distinctive Frechitas (formerly Ptychites) layer, and is therefore assumed to be middle Anisian. The Steinkobbe core was sampled at 99-100 m, just above the Olenekian-Anisian transition. It is therefore assumed to be lower Anisian. The two isochron ages overlap within uncertainty, and fall within constraints provided by biozones and the current ICS-approved stage boundary ages. The Re-Os ages support the correlation of the Botneheia and Steinkobbe formations. The nearly identical Osi ratios suggest regional homogeneity of seawater and provide new information for the Os seawater curve, marking a relatively high 187Os/188Os ratio during profound ocean anoxia in the Middle Triassic.

  17. Feedback in low-mass galaxies in the early Universe.

    Science.gov (United States)

    Erb, Dawn K

    2015-07-09

    The formation, evolution and death of massive stars release large quantities of energy and momentum into the gas surrounding the sites of star formation. This process, generically termed 'feedback', inhibits further star formation either by removing gas from the galaxy, or by heating it to temperatures that are too high to form new stars. Observations reveal feedback in the form of galactic-scale outflows of gas in galaxies with high rates of star formation, especially in the early Universe. Feedback in faint, low-mass galaxies probably facilitated the escape of ionizing radiation from galaxies when the Universe was about 500 million years old, so that the hydrogen between galaxies changed from neutral to ionized-the last major phase transition in the Universe.

  18. The Lower Triassic Sorkh Shale Formation of the Tabas Block, east central Iran: Succesion of a failed-rift basin at the Paleotethys margin

    Science.gov (United States)

    Lasemi, Y.; Ghomashi, M.; Amin-Rasouli, H.; Kheradmand, A.

    2008-01-01

    The Lower Triassic Sorkh Shale Formation is a dominantly red colored marginal marine succession deposited in the north-south trending Tabas Basin of east central Iran. It is correlated with the unconformity-bounded lower limestone member of the Elika Formation of the Alborz Mountains of northern Iran. The Sorkh Shale is bounded by the pre-Triassic and post-Lower Triassic interregional unconformities and consists mainly of carbonates, sandstones, and evaporites with shale being a minor constituent. Detailed facies analysis of the Sorkh Shale Formation resulted in recognition of several genetically linked peritidal facies that are grouped into restricted subtidal, carbonate tidal flat, siliciclastic tidal flat, coastal plain and continental evaporite facies associations. These were deposited in a low energy, storm-dominated inner-ramp setting with a very gentle slope that fringed the Tabas Block of east central Iran and passed northward (present-day coordinates) into deeper water facies of the Paleotethys passive margin of northern Cimmerian Continent. Numerous carbonate storm beds containing well-rounded intraclasts, ooids and bioclasts of mixed fauna are present in the Sorkh Shale Formation of the northern Tabas Basin. The constituents of the storm beds are absent in the fair weather peritidal facies of the Sorkh Shale Formation, but are present throughout the lower limestone member of the Elika Formation. The Tabas Block, a part of the Cimmerian continent in east central Iran, is a rift basin that developed during Early Ordovician-Silurian Paleotethys rifting. Facies and sequence stratigraphic analyses of the Sorkh Shale Formation has revealed additional evidence supporting the Tabas Block as a failed rift basin related to the Paleotethys passive margin. Absence of constituents of the storm beds in the fair weather peritidal facies of the Sorkh Shale Formation, presence of the constituents of the storm beds in the fair weather facies of the Elika Formation (the

  19. The development of the Middle Triassic tectonical controlled Germanic Basin of Central Europe and the palaeoenvironmental related distribution of marine and terrestrial reptiles

    Science.gov (United States)

    Diedrich, Cajus G.

    2010-05-01

    -Italy). Bolletino della Società Paleontologica Italiana, 41 (1), 37-40. Bachmann, G.H. and Aref, M.A.M., 2005. A seismite in Triassic gypsum deposits (Grabfeld Formation, Ladinian), Southwest Germany. Sedimentary Geology 180, 75-89. De Zanche, V. and Farabegoli, E. 1988. Anisian paleogeographic evolution in the Central-Western Southern Alps. Memoirs Scientifique Geologique 40, 399-411. Demathieu, G.R. 1985. Trace fossil assemblages in Middle Triassic marginal marine deposits, eastern border of the Massif Central, France. Societe Economie Paléontologie et Mineralogie, Special Publications, 35, 53-66. Diedrich, C. 2005. Actuopalaeontological trackway experiments with Iguana on intertidal flat carbonates of the Arabian Gulf - a comparison to fossil Rhynchosauroides tracks of Triassic carbonate tidal flat megatracksites in the European Germanic Basin. Senckenbergiana maritime, 35 (2), 203-220. Diedrich, C. 2008a. Millions of reptile tracks - Early to Middle Triassic carbonate tidal flat migration bridges of Central Europe. Palaeogeography, Palaeoclimatology, Palaeoecology, 259, 410-423. Diedrich, C. 2008b. Palaeogeographic evolution of the marine Middle Triassic marine Germanic Basin changements - with emphasis on the carbonate tidal flat and shallow marine habitats of reptiles in Central Pangaea. Global and Planetary Change, 65 (2009), 27-55. Diedrich, C. 2009a. The vertebrates of the Anisian/Ladinian boundary (Middle Triassic) from Bissendorf (NW Germany) and their contribution to the anatomy, palaeoecology, and palaeobiogeography of the Germanic Basin reptiles. Palaeogeography, Palaeoclimatology, Palaeoecology, 273 (2009), 1-16. Diedrich, C. 2009b. Die Saurierspuren-Grabung im basalen Mittleren Muschelkalk (Anis, Mitteltrias) von Bernburg (Sachsen-Anhalt). Archäologie in Sachsen-Anhalt, Sonderband 2009, 1-62. Diedrich, 2010a. Palaeoecology of Placodus gigas (Reptilia) and other placodontids - macroalgae feeder of the Middle Triassic in the Germanic Basin of Central Europe and

  20. Paleomagnetic evidence for a Tertiary not Triassic age for rocks in the lower part of the Grober-Fuqua #1 well, southeastern Albuquerque Basin, New Mexico

    Science.gov (United States)

    Hudson, M.R.; Grauch, V.J.S.

    2003-01-01

    A sedimentary sequence penetrated in the lower part of the Grober-Fuqua #1 well in the southeastern Albuquerque Basin has previously been interpreted as either Triassic or Eocene in age. Paleomagnetic study of three specimens from two core fragments yielded a 54.5?? mean inclination of remanent magnetization relative to bedding. This inclination is like that expected in Tertiary time and is distinct from an expected low-angle Triassic inclination. Although the data are very few, when considered in combination with stratigraphic relations and the presence of a gravity low in this southeastern part of the basin, the paleomagnetic evidence favors a Tertiary age for strata in the lower part of the Grober-Fuqua #1 well.

  1. Boron isotopes in brachiopods during the end-Permian mass extinction: constraints on pH evolution and seawater chemistry

    Science.gov (United States)

    Jurikova, Hana; Gutjahr, Marcus; Liebetrau, Volker; Brand, Uwe; Posenato, Renato; Garbelli, Claudio; Angiolini, Lucia; Eisenhauer, Anton

    2017-04-01

    The global biogeochemical cycling of carbon is fundamental for life on Earth with the ocean playing a key role as the largest and dynamically evolving CO2 reservoir. The boron isotope composition (commonly expressed in δ11B) of marine calcium carbonate is considered to be one of the most reliable paleo-pH proxies, potentially enabling us to reconstruct past ocean pH changes and understand carbon cycle perturbations along Earth's geological record (e.g. Foster et al., 2008; Clarkson et al., 2015). Brachiopods present an advantageous and largely underutilised archive for Phanerozoic carbon cycle reconstructions considering their high abundance in the geological record and its origin dating back to the early Cambrian. Moreover, their shell made of low-magnesium calcite makes these marine calcifiers more resistant to post-depositional diagenetic alteration of primary chemical signals. We have investigated the δ11B using MC-ICP-MS (Neptune Plus) and B/Ca and other elemental ratios (Mg/Ca, Sr/Ca, Al/Ca, Li/Ca, Ba/Ca, Na/Ca and Fe/Ca) using ICP-MS-Quadrupole (Agilent 7500cx) from the same specimens in pristine brachiopod shells from two sections from northern Italy during the Late Permian. These sections cover the δ13C excursion in excess of ˜4 ‰ (Brand et al., 2012) and are associated with major climate and environmental perturbations that lead to the mass extinction event at the Permian-Triassic boundary. Particular emphasis will be placed on the implications of our new paleo-pH estimates on the seawater chemistry during the Late Permian. Brand, U., Posenato, R., Came, R., Affek, H., Angiolini, L., Azmy, K. and Farabegoli, E.: The end-Permian mass extinction: A rapid volcanic CO2 and CH4-climatic catastrophe, Chemical Geology 323, 121-144, doi:10.1016/j.chemgeo.2012.06.015, 2012. Clarkson, M.O., Kasemann, S.A., Wood, R.A., Lenton, T.M., Daines, S.J., Richoz, S., Ohnemueller, F., Meixner, A., Poulton, S.W. and Tipper, E.T.: Ocean acidification and the Permo-Triassic

  2. Microbialites and global environmental change across the Permian-Triassic boundary: a synthesis.

    Science.gov (United States)

    Kershaw, S; Crasquin, S; Li, Y; Collin, P-Y; Forel, M-B; Mu, X; Baud, A; Wang, Y; Xie, S; Maurer, F; Guo, L

    2012-01-01

    Permian-Triassic boundary microbialites (PTBMs) are thin (0.05-15 m) carbonates formed after the end-Permian mass extinction. They comprise Renalcis-group calcimicrobes, microbially mediated micrite, presumed inorganic micrite, calcite cement (some may be microbially influenced) and shelly faunas. PTBMs are abundant in low-latitude shallow-marine carbonate shelves in central Tethyan continents but are rare in higher latitudes, likely inhibited by clastic supply on Pangaea margins. PTBMs occupied broadly similar environments to Late Permian reefs in Tethys, but extended into deeper waters. Late Permian reefs are also rich in microbes (and cements), so post-extinction seawater carbonate saturation was likely similar to the Late Permian. However, PTBMs lack widespread abundant inorganic carbonate cement fans, so a previous interpretation that anoxic bicarbonate-rich water upwelled to rapidly increase carbonate saturation of shallow seawater, post-extinction, is problematic. Preliminary pyrite framboid evidence shows anoxia in PTBM facies, but interbedded shelly faunas indicate oxygenated water, perhaps there was short-term pulsing of normally saturated anoxic water from the oxygen-minimum zone to surface waters. In Tethys, PTBMs show geographic variations: (i) in south China, PTBMs are mostly thrombolites in open shelf settings, largely recrystallised, with remnant structure of Renalcis-group calcimicrobes; (ii) in south Turkey, in shallow waters, stromatolites and thrombolites, lacking calcimicrobes, are interbedded, likely depth-controlled; and (iii) in the Middle East, especially Iran, stromatolites and thrombolites (calcimicrobes uncommon) occur in different sites on open shelves, where controls are unclear. Thus, PTBMs were under more complex control than previously portrayed, with local facies control playing a significant role in their structure and composition. © 2011 Blackwell Publishing Ltd.

  3. Tectonic contact beetwen Paleozoic and Triassic rocks south of Podolševa (Slovenia

    Directory of Open Access Journals (Sweden)

    Bogomir Celarc

    2002-12-01

    Full Text Available Tectonic contact between Paleozoic and Triassic rocks south of Podolševa, which passes prominent cliffs over the left bank of Savinja river (Klemenča peč, Jamnikova peč, Golerjeva peč, Strevčeva peč and Huda peč, is on the basis of the new mapping a steep fault, dipping to the north. In the last phase, the fault was dextral strike – slip active, before was probably reverse, and can represent accompanying structure of Periadriatic lineament. It was under transpresive tectonic regime and complies with model of positive flower structure.We named it Podol{evski prelom (Podol{eva fault.

  4. Evidence for a constant initial mass function in early-type galaxies based on their X-ray binary populations

    International Nuclear Information System (INIS)

    Peacock, Mark B.; Zepf, Stephen E.; Maccarone, Thomas J.; Kundu, Arunav; Gonzalez, Anthony H.; Lehmer, Bret D.; Maraston, Claudia

    2014-01-01

    A number of recent studies have proposed that the stellar initial mass function (IMF) of early type galaxies varies systematically as a function of galaxy mass, with higher mass galaxies having bottom-heavy IMFs. These bottom-heavy IMFs have more low-mass stars relative to the number of high mass stars, and therefore naturally result in proportionally fewer neutron stars (NSs) and black holes (BHs). In this paper, we specifically predict the variation in the number of BHs and NSs based on the power-law IMF variation required to reproduce the observed mass-to-light ratio trends with galaxy mass. We then test whether such variations are observed by studying the field low-mass X-ray binary (LMXB) populations of nearby early-type galaxies. In these binaries, an NS or BH accretes matter from a low-mass donor star. Their number is therefore expected to scale with the number of BHs and NSs present in a galaxy. We find that the number of LMXBs per K-band light is similar among the galaxies in our sample. These data therefore demonstrate the uniformity of the slope of the IMF from massive stars down to those now dominating the K-band light and are consistent with an invariant IMF. Our results are inconsistent with an IMF which varies from a Kroupa/Chabrier like IMF for low-mass galaxies to a steep power-law IMF (with slope x = 2.8) for high mass galaxies. We discuss how these observations constrain the possible forms of the IMF variations and how future Chandra observations can enable sharper tests of the IMF.

  5. Evidence for a constant initial mass function in early-type galaxies based on their X-ray binary populations

    Energy Technology Data Exchange (ETDEWEB)

    Peacock, Mark B.; Zepf, Stephen E. [Department of Physics and Astronomy, Michigan State University, East Lansing, MI 48824 (United States); Maccarone, Thomas J. [Texas Tech University, Physics Department, Box 41051, Lubbock, TX 79409 (United States); Kundu, Arunav [Eureka Scientific, Inc., 2452 Delmer Street, Suite 100 Oakland, CA 94602 (United States); Gonzalez, Anthony H. [Department of Astronomy, University of Florida, Gainesville, FL 32611 (United States); Lehmer, Bret D. [The Johns Hopkins University, Homewood Campus, Baltimore, MD 21218 (United States); Maraston, Claudia, E-mail: mpeacock@msu.edu [Institute of Cosmology and Gravitation, Dennis Sciama Building, Burnaby Road, Portsmouth PO1 3FX (United Kingdom)

    2014-04-01

    A number of recent studies have proposed that the stellar initial mass function (IMF) of early type galaxies varies systematically as a function of galaxy mass, with higher mass galaxies having bottom-heavy IMFs. These bottom-heavy IMFs have more low-mass stars relative to the number of high mass stars, and therefore naturally result in proportionally fewer neutron stars (NSs) and black holes (BHs). In this paper, we specifically predict the variation in the number of BHs and NSs based on the power-law IMF variation required to reproduce the observed mass-to-light ratio trends with galaxy mass. We then test whether such variations are observed by studying the field low-mass X-ray binary (LMXB) populations of nearby early-type galaxies. In these binaries, an NS or BH accretes matter from a low-mass donor star. Their number is therefore expected to scale with the number of BHs and NSs present in a galaxy. We find that the number of LMXBs per K-band light is similar among the galaxies in our sample. These data therefore demonstrate the uniformity of the slope of the IMF from massive stars down to those now dominating the K-band light and are consistent with an invariant IMF. Our results are inconsistent with an IMF which varies from a Kroupa/Chabrier like IMF for low-mass galaxies to a steep power-law IMF (with slope x = 2.8) for high mass galaxies. We discuss how these observations constrain the possible forms of the IMF variations and how future Chandra observations can enable sharper tests of the IMF.

  6. Mass spectrometry-based serum proteome pattern analysis in molecular diagnostics of early stage breast cancer

    Directory of Open Access Journals (Sweden)

    Stobiecki Maciej

    2009-07-01

    Full Text Available Abstract Background Mass spectrometric analysis of the blood proteome is an emerging method of clinical proteomics. The approach exploiting multi-protein/peptide sets (fingerprints detected by mass spectrometry that reflect overall features of a specimen's proteome, termed proteome pattern analysis, have been already shown in several studies to have applicability in cancer diagnostics. We aimed to identify serum proteome patterns specific for early stage breast cancer patients using MALDI-ToF mass spectrometry. Methods Blood samples were collected before the start of therapy in a group of 92 patients diagnosed at stages I and II of the disease, and in a group of age-matched healthy controls (104 women. Serum specimens were purified and the low-molecular-weight proteome fraction was examined using MALDI-ToF mass spectrometry after removal of albumin and other high-molecular-weight serum proteins. Protein ions registered in a mass range between 2,000 and 10,000 Da were analyzed using a new bioinformatic tool created in our group, which included modeling spectra as a sum of Gaussian bell-shaped curves. Results We have identified features of serum proteome patterns that were significantly different between blood samples of healthy individuals and early stage breast cancer patients. The classifier built of three spectral components that differentiated controls and cancer patients had 83% sensitivity and 85% specificity. Spectral components (i.e., protein ions that were the most frequent in such classifiers had approximate m/z values of 2303, 2866 and 3579 Da (a biomarker built from these three components showed 88% sensitivity and 78% specificity. Of note, we did not find a significant correlation between features of serum proteome patterns and established prognostic or predictive factors like tumor size, nodal involvement, histopathological grade, estrogen and progesterone receptor expression. In addition, we observed a significantly (p = 0

  7. Triassic tholeiitic dolerites («ophites» of the El Grado diapir (Pyrenees, Huesca, Spain: emplacement and composition

    Directory of Open Access Journals (Sweden)

    Lago San José, M.

    2000-04-01

    Full Text Available Mesozoic dolerites in the south Pyrenean sector of El Grado (Huesca, Spain preserve emplacement structures (fluidity structures at the top and load structures at the base developed during their intrusion into unconsolidated marly-evaporitic Triassic sediments (Keuper facies. By analogy with other dolerites in the south Pyrenean domain, their emplacement age is equivalent to the uppermost Keuper facies terms, but prior to the final Triassic-early Liassic carbonated sediments. Radiometric ages (187-197±7 Ma show that the emplacement occurred during the lower Liassic. The petralogical differentiation from the chilled margin facies to the central facies, and also to the late pegmatitoids, is consistent with that obtained from major elements, trace elements and REE. Their tholeiitic affinity, as defined by their geochemical composition, is equivalent to that of similar racks in the Pyrenean domain. However, the rocks analyzed here, which are located at the external sector of this domain, display a greater petralogical and geochemical differentiation as compared to similar rocks in the central sectors of the Pyrenean domain.Las doleritas mesozoicas del sector surpirenaico de El Grado (Huesca conservan estructuras de emplazamiento (con desarrollo del movimiento de lava fluida al techo y de carga en su base desarrolladas al instruir en los sedimentos margo-evaporíticos en facies Keuper, todavía inconsolidados. Por similitud con otras doleritas del dominio surpirenaico, la edad del emplazamiento es equivalente a la de los términos superiores de la facies Keuper y previa a la sedimentación carbonatada del Trías terminal-Lías inferior. Las determinaciones de edades radiométricas (187-197±7 Ma indican que el emplazamiento debió tener lugar durante el Lías inferior. La diferenciación petralógica, desde la facies del borde enfriado a la central y, también, al posterior diferenciado pegmatoide concuerda con la obtenida con elementos mayores

  8. The metallogeny of Late Triassic rifting of the Alexander terrane in southeastern Alaska and northwestern British Columbia

    Science.gov (United States)

    Taylor, C.D.; Premo, W.R.; Meier, A.L.; Taggart, J.E.

    2008-01-01

    A belt of unusual volcanogenic massive sulfide (VMS) occurrences is located along the eastern margin of the Alexander terrane throughout southeastern Alaska and northwestern British Columbia and exhibits a range of characteristics consistent with a variety of syngenetic to epigenetic deposit types. Deposits within this belt include Greens Creek and Windy Craggy, the economically most significant VMS deposit in Alaska and the largest in North America, respectively. The occurrences are hosted by a discontinuously exposed, 800-km-long belt of rocks that consist of a 200- to 800-m-thick sequence of conglomerate, limestone, marine elastic sedimentary rocks, and tuff intercalated with and overlain by a distinctive unit of mafic pyroclastic rocks and pillowed flows. Faunal data bracket the age of the host rocks between Anisian (Middle Triassic) and late Norian (late Late Triassic). This metallogenic belt is herein referred to as the Alexander Triassic metallogenic belt. The VMS occurrences show systematic differences in degree of structural control, chemistry, and stratigraphic setting along the Alexander Triassic metallogenic belt that suggest important spatial or temporal changes in the tectonic environment of formation. At the southern end of the belt, felsic volcanic rocks overlain by shallow-water limestones characterize the lower part of the sequence. In the southern and middle portion of the belt, a distinctive pebble conglomerate marks the base of the section and is indicative of high-energy deposition in a near slope or basin margin setting. At the northern end of the belt the conglomerates, limestones, and felsic volcanic rocks are absent and the belt is composed of deep-water sedimentary and mafic volcanic rocks. This northward change in depositional environment and lithofacies is accompanied by a northward transition from epithermal-like structurally controlled, discontinuous, vein- and pod-shaped, Pb-Zn-Ag-Ba-(Cu) occurrences with relatively simple mineralogy

  9. Subduction of Proterozoic to Late Triassic continental basement in the Guatemala suture zone: A petrological and geochronological study of high-pressure metagranitoids from the Chuacús complex

    Science.gov (United States)

    Maldonado, Roberto; Ortega-Gutiérrez, Fernando; Ortíz-Joya, Guillermo A.

    2018-05-01

    Many continental subduction complexes contain abundant granitic rocks coexisting with minor volumes of eclogite-facies rocks. Characterization of granitic protoliths is crucial to decipher the origin of subducted continental crust, whereas knowledge of its metamorphic evolution is required to constrain the mechanisms of burial and exhumation. In this work we present geochronological and petrological evidence that demonstrate the occurrence of a subducted Proterozoic to Late Triassic granitic basement in the Chuacús complex of central Guatemala. Metagranitoids exposed in this area are interlayered with eclogite and other high-pressure rocks, and their structure is considerably variable due to strain partitioning during deformation. Laser ablation-inductively coupled plasma-mass spectrometry U-Pb zircon data from two ferroan metagranites yield protolith crystallization ages of ca. 1.1 Ga and their trace-element abundances suggest an origin related to intraplate magmatism, while a high-silica, peraluminous metagranite is dated at 1.0 Ga and was probably originated by partial melting of a high-grade continental crust. On the other hand, two megacrystic to augen metagranitoids yield protolith crystallization ages of ca. 224 Ma, which are identical within errors to the protolith age of hosted eclogitic metabasites. Their high incompatible trace element abundances together with the observed spatial-temporal relationships with mafic protoliths suggest that Late Triassic bimodal magmatism in the Chuacús complex was probably originated in a within-plate setting. Regardless of their age or structure, the studied metagranites preserve evidences for high-pressure metamorphic equilibration, such as the occurrence of Ca-rich garnet (XCa up to 0.52) in association with phengite (Si contents of up to 3.4 pfu) and rutile. The integration of Zr-in-rutile thermometry and phengite barometry allows the peak metamorphic conditions to be constrained at 640-680 °C and 13 kbar. This

  10. Deciphering the diagenetic alteration degree in thrombolites across the Permian-Triassic boundary and the evaluation of REY as a proxy of palaeoseawater

    Science.gov (United States)

    Li, Rong

    2017-10-01

    The thrombolites across the Permian-Triassic boundary (PTB) are widely distributed in South China. In order to examine the utility of rare earth element and yttrium (REY) in the thrombolites as a proxy of palaeoseawater, the petrographic and geochemical (Sr, Mn, Fe, REY, δ13C, δ18O) features of thrombolites from Cili and Taiping sections are comparatively studied to determine the diagenetic alteration degrees of the thrombolites and the impact of diagenesis on REY concentrations and distribution patterns. The thrombolites from Cili section, digitate in mesoscopic morphology, are latest Permian in age. In contrast, most of the thrombolites in Taiping section are mottled and formed in the earliest Triassic. The variation in thrombolite morphology across the PTB is probably related to increasing amount of metazoan and increasing intensity of bioturbation after the end-Permian mass extinction. Compared to the thrombolites from Taiping section, those from Cili section underwent more extensive diagenetic alteration, which are characterized by more dolomitic content, lower Sr concentrations, more negative δ18O values, and higher Mn/Sr ratios. The REY concentrations are higher in the thrombolites from Taiping section (5-303 ppm, average 48 ppm) than in the thrombolites from Cili section (17-34 ppm, average 25 ppm). Shale-normalized REY distribution patterns of the thrombolites from both sites are similar to those of oxygenated seawater, which are characterized by positive La anomalies (Pr/Pr∗ = 1.1-1.5), negative Ce anomalies (Ce/Ce∗ = 0.2-0.9), and light rare earth element (LREE) depletion. The preserved seawater like REY distribution pattern indicates that diagenesis did not alter the REY distribution patterns. The thrombolite samples from Cili section, compared to the counterparts from Taiping, have less positive La anomaly and less negative Ce anomaly. For the thrombolites from Cili section, a positive correlation exists between Ce anomaly and siliciclastic

  11. A Dome-Headed Stem Archosaur Exemplifies Convergence among Dinosaurs and Their Distant Relatives.

    Science.gov (United States)

    Stocker, Michelle R; Nesbitt, Sterling J; Criswell, Katharine E; Parker, William G; Witmer, Lawrence M; Rowe, Timothy B; Ridgely, Ryan; Brown, Matthew A

    2016-10-10

    Similarities in body plan evolution, such as wings in pterosaurs, birds, and bats or limblessness in snakes and amphisbaenians, can be recognized as classical examples of convergence among animals [1-3]. We introduce a new Triassic stem archosaur that is unexpectedly and remarkably convergent with the "dome-headed" pachycephalosaur dinosaurs that lived over 100 million years later. Surprisingly, numerous additional taxa in the same assemblage (the Otis Chalk assemblage from the Dockum Group of Texas) demonstrate the early acquisition of morphological novelties that were later convergently evolved by post-Triassic dinosaurs. As one of the most successful clades of terrestrial vertebrates, dinosaurs came to occupy an extensive morphospace throughout their diversification in the Mesozoic Era [4, 5], but their distant relatives were first to evolve many of those "dinosaurian" body plans in the Triassic Period [6-8]. Our analysis of convergence between archosauromorphs from the Triassic Period and post-Triassic archosaurs demonstrates the early and extensive exploration of morphospace captured in a single Late Triassic assemblage, and we hypothesize that many of the "novel" morphotypes interpreted to occur among archosaurs later in the Mesozoic already were in place during the initial Triassic archosauromorph, largely non-dinosaurian, radiation and only later convergently evolved in diverse dinosaurian lineages. Copyright © 2016 Elsevier Ltd. All rights reserved.

  12. Teleseismic P-wave tomography and the upper mantle structure of the Sulu orogenic belt (China): implications for Triassic collision and exhumation mechanism

    Science.gov (United States)

    Peng, Miao; Tan, Handong; Jiang, Mei; Xu, Zhiqin; Li, Zhonghai; Xu, Lehong

    2016-12-01

    As the largest ultrahigh-pressure (UHP) metamorphic tectonic unit outcropping in the world, the Dabie-Sulu UHP metamorphic belt is considered to be one of the best areas for studying the continental dynamics. However, their continental collision and exhumation mechanism are still debated. We performed a 3D teleseismic P-wave tomography beneath the Sulu orogen for the purpose of understanding the deep structure. The tomographic results show that there is a prominently near-SN-trending low-velocity zone (LVZ) close to the Tanlu fault (TLF), indicating a slab tear of the subducted Yangtze plate (YZP) during the initial Early Triassic collision. Our results also suggest that both the Yangze crustal slab and the North China lithospheric slab were dragged downwards by the subducted oceanic slab, which constituted a ‘two-sided’ subduction mode. A conceptual geodynamic model is proposed to explain the exhumation of Sulu high- to UHP rocks and imply a polyphase exhumation driven by buoyancy of continental materials at different depth and upward extrusion of crustal partial melting rocks to the surface at the later stage.

  13. Early detection of tumor masses by in vivo hematoporphyrin-mediated fluorescence imaging

    International Nuclear Information System (INIS)

    Autiero, Maddalena; Celentano, Luigi; Cozzolino, Rosanna; Laccetti, Paolo; Marotta, Marcello; Mettivier, Giovanni; Cristina Montesi, Maria; Quarto, Maria; Riccio, Patrizia; Roberti, Giuseppe; Russo, Paolo

    2007-01-01

    We investigated the capability of fluorescence reflectance imaging (FRI) for the early detection of surface tumors in mice. We used a hematoporphyrin (HP) compound (HP dichlorohydrate) as a red fluorescent marker and a low noise, high sensitivity, digital CCD camera for fluorescence imaging. In this preliminary study, highly malignant anaplastic human thyroid carcinoma cells were implanted subcutaneously in one mouse and their growth was monitored daily for 5 days by FRI. The selective HP uptake by the tumor tissues was successfully observed: we observed the fluorescence of tumor only 3 days after cancer cells injection, i.e. when the tumor mass was neither visible (to the naked eye) or palpable. These measurements indicate that FRI is a suitable technique to detect minute subcutaneous tumor masses. This FRI system will be coupled to a radionuclide imaging system based on a CdTe detector for in vivo multimodal imaging in mice

  14. Paleoclimatology of Upper Triassic Playa Cycles. New Insights Into an Orbital Controlled Monsoon System (Norian, German Basin)

    OpenAIRE

    Vollmer, Thorsten

    2005-01-01

    Paleoclimatology of Upper Triassic Playa Cycles: New Insights Into an Orbital Controlled Monsoon System (Norian, German Basin) Abstract The main purpose of the project was to study the rhythmic sediments of the Steinmergel Keuper playa system in the North and South German basin in order to test the hypothesis of possible climate control on sedimentation. Furthermore, in case of there being orbital control on sedimentation, the North/South correlation was tested based on high-resolution cyclos...

  15. The post collisional metamorphic evolution from Ultra High Temperature to Amphibolite facies metamorphism in the Odesan area during the Triassic collision between the North and South China cratons.

    Science.gov (United States)

    Lee, Byung Choon; Oh, Chang Whan; Kim, Tae Sung; Yi, Kee Wook

    2015-04-01

    The Odaesan Gneiss Complex (OGC) is the eastern end of the Hongseong-Odesan collision belt in Korean Peninsula which is the extension of the Dabie-Sulu collision belt between the North and South China blocks. The OGC mainly consists of banded and migmatitic gneiss with porphyritic granitoid and amphibolite. The banded gneiss can be subdivided into garnet-biotite and garnet-orthopyroxene banded gneisses. The highest metamorphic P/T conditions of the migmatitic and garnet-biotite banded gneiss were 760-820°C/6.3-7.2kbar and 810-840°C/7.2-7.8kbar respectively. On the other hand, the garnet-orthopyroxene banded gneiss records 940-950°C/10.5-10.7kbar that is corresponded to UHT metamorphic condition. These data indicate that the peak UHT metamorphic condition of the study area was preserved only within the garnet-orthopyroxene banded gneiss because its lower water content than other gneisses and UHT metamorphic mineral assemblage was completely replaced by the granulite facies metamorphism in other gneisses due to their higher water content than the garnet-orthopyroxene banded gneiss. Finally all gneisses experienced amphibolite facies retrograde metamorphism which is observed locally within rocks, such as garnet rim and surrounding area. The peak UHT metamorphism is estimated to occur at ca. 250-230 Ma using SHRIMP zircon U-Pb age dating and was caused by the heat supplied from asthenospheric mantle through the opening formed by slab break-off during early post collision stage. The calculated metamorphic conditions represent that geothermal gradient of the study area during the post collision stage was 86°C/kbar indicating the regional low-P/T metamorphic event. Besides the Triassic metamorphic age, two Paleoproterozoic metamorphic ages of ca. 1930 and 1886 Ma are also recognized by the SHRIMP age dating from the banded gneisses and Paleoproterozoic emplacement age of ca. 1847 Ma is identified from the porphyritic granitoid which formed in the within plate tectonic

  16. Toward Understanding the Early Stags of an Impulsively Accelerated Coronal Mass Ejection

    Science.gov (United States)

    2010-08-09

    B. E., & Howard, R. A . 2009, ApJ, 702, 901 Wood, B. E., Karovska , M., Chen, J., Brueckner, G. E., Cook, J. W., & Howard, R. A . 1999, ApJ, 512, 484...ar X iv :1 00 8. 11 71 v1 [ as tr o- ph .S R ] 6 A ug 2 01 0 Astronomy & Astrophysics manuscript no. bubble c© ESO 2010 August 9, 2010 Toward...understanding the early stages of an impulsively accelerated coronal mass ejection SECCHI observations S. Patsourakos1, A . Vourlidas2, and B. Kliem3,4

  17. Allogenic controls on the fluvial architecture and fossil preservation of the Upper Triassic Ischigualasto Formation, NW Argentina

    Science.gov (United States)

    Colombi, Carina E.; Limarino, Carlos O.; Alcober, Oscar A.

    2017-12-01

    The Upper Triassic Ischigualasto Formation in NW Argentina was deposited in a fluvial system during the synrift filling of the extensional Ischigualasto-Villa Unión Basin. The expansive exposures of the fluvial architecture and paleosols provide a framework to reconstruct the paleoenvironmental evolution of this basin during the Upper Triassic using continental sequence stratigraphy. The Ischigualasto Formation deposition can be divided into seven sequential sedimentary stages: the 1) Bypass stage; 2) Confined low-accommodation stage; 3) Confined high accommodation stage; 4) Unstable-accommodation stage; 5) Unconfined high-accommodation stage; 6) Unconfined low-accommodation stage; and finally, 7) Unconfined high-accommodation stage. The sedimentary evolution of the Ischigualasto Formation was driven by different allogenic controls such as rises and falls in lake levels, local tectonism, subsidence, volcanism, and climate, which also produced modifications of the equilibrium profile of the fluvial systems. All of these factors result in different accommodations in central and flank areas of the basin, which led to different architectural configurations of channels and floodplains. Allogenic processes affected not only the sequence stratigraphy of the basin but also the vertebrate and plant taphocenosis. Therefore, the sequence stratigraphy can be used not only as a predictive tool related to fossil occurrence but also to understand the taphonomic history of the basin at each temporal interval.

  18. CHAMBERED HEXACTINELLID SPONGES FROM UPPER TRIASSIC(NORIAN-RHAETIAN? REEFS OF NAYBAND FORMATION IN CENTRAL IRAN

    Directory of Open Access Journals (Sweden)

    B. SENOWBARI-DARYAN

    2012-07-01

    Full Text Available This paper describes several chambered hexactinellid sponges, including Casearia iranica n.sp., C. vezvanensis n. sp., C. delijanensis n. sp., Esfahanella magna gen. n. n. sp., and E. parva gen. n. n. sp. from reefs of the Upper Triassic (Norian-Rhaetian Nayband Formation exposed south of the town of Delijan in central Iran. The relative abundance of chambered and non-chambered hexactinellid sponges at this locality - as compared to hypercalcified representatives - highlight the importance of this group of sponges in reef and reefal limestones in central and east Tethys (China, Caucasia, Iran. 

  19. Relict zircon U-Pb age and O isotope evidence for reworking of Neoproterozoic crustal rocks in the origin of Triassic S-type granites in South China

    Science.gov (United States)

    Gao, Peng; Zheng, Yong-Fei; Chen, Yi-Xiang; Zhao, Zi-Fu; Xia, Xiao-Ping

    2018-02-01

    Granites derived from partial melting of sedimentary rocks are generally characterized by high δ18O values and abundant relict zircons. Such relict zircons are valuable in tracing the source rocks of granites and the history of crustal anatexis. Here we report in-situ U-Pb ages, O isotopes and trace elements in zircons from Triassic granites in the Zhuguangshan and Jiuzhou regions, which are located in the Nanling Range and the Darongshan area, respectively, in South China. Zircon U-Pb dating yields magma crystallization ages of 236 ± 2 Ma for the Zhuguangshan granites and 246 ± 2 Ma to 252 ± 3 Ma for the Jiuzhou granites. The Triassic syn-magmatic zircons are characterized by high δ18O values of 10.1-11.9‰ in Zhuguangshan and 8.5-13.5‰ in Jiuzhou. The relict zircons show a wide range of U-Pb ages from 315 to 2185 Ma in Zhuguangshan and from 304 to 3121 Ma in Jiuzhou. Nevertheless, a dominant age peak of 700-1000 Ma is prominent in both occurrences, demonstrating that their source rocks were dominated by detrital sediments weathered from Neoproterozoic magmatic rocks. Taking previous results for regional granites together, Neoproterozoic relict zircons show δ18O values in a small range from 5 to 8‰ for the Nanling granites but a large range from 5 to 11‰ for the Darongshan granites. In addition, relict zircons of Paleozoic U-Pb age occur in the two granitic plutons. They exhibit consistently high δ18O values similar to the Triassic syn-magmatic zircons in the host granites. These Paleozoic relict zircons are interpreted as the peritectic product during transient melting of the metasedimentary rocks in response to the intracontinental orogenesis in South China. Therefore, the relict zircons of Neoproterozoic age are directly inherited from the source rocks of S-type granites, and those of Paleozoic age record the transient melting of metasedimentary rocks before intensive melting for granitic magmatism in the Triassic.

  20. The influence of galaxy environment on the stellar initial mass function of early-type galaxies

    Science.gov (United States)

    Rosani, Giulio; Pasquali, Anna; La Barbera, Francesco; Ferreras, Ignacio; Vazdekis, Alexandre

    2018-06-01

    In this paper, we investigate whether the stellar initial mass function (IMF) of early-type galaxies depends on their host environment. To this purpose, we have selected a sample of early-type galaxies from the SPIDER catalogue, characterized their environment through the group catalogue of Wang et al., and used their optical Sloan Digital Sky Survey (SDSS) spectra to constrain the IMF slope, through the analysis of IMF-sensitive spectral indices. To reach a high enough signal-to-noise ratio, we have stacked spectra in velocity dispersion (σ0) bins, on top of separating the sample by galaxy hierarchy and host halo mass, as proxies for galaxy environment. In order to constrain the IMF, we have compared observed line strengths and predictions of MIUSCAT/EMILES synthetic stellar population models, with varying age, metallicity, and `bimodal' (low-mass tapered) IMF slope (Γ _b). Consistent with previous studies, we find that Γ _b increases with σ0, becoming bottom-heavy (i.e. an excess of low-mass stars with respect to the Milky Way like IMF) at high σ0. We find that this result is robust against the set of isochrones used in the stellar population models, as well as the way the effect of elemental abundance ratios is taken into account. We thus conclude that it is possible to use currently state-of-the-art stellar population models and intermediate resolution spectra to consistently probe IMF variations. For the first time, we show that there is no dependence of Γb on environment or galaxy hierarchy, as measured within the 3 arcsec SDSS fibre, thus leaving the IMF as an intrinsic galaxy property, possibly set already at high redshift.

  1. Late Palaeozoic to Triassic formations unconformably deposited over the Ronda peridotites (Betic Cordilleras): Evidence for their Variscan time of crustal emplacement

    Energy Technology Data Exchange (ETDEWEB)

    Sanz de Galdeano, C.; Ruiz Cruz, M.D.

    2016-07-01

    The age of the emplacement of the Ronda Peridotites has been widely debated during recent decades, and ages ranging from the Palaeozoic to the early Miocene have been proposed, although most of the current interpretations suggest an Oligocene-Miocene age. In this article, we describe two meta-sedimentary formations (the lower one formed by detrital sediments and the upper one by marbles) that were unconformably deposited over the Ronda peridotites and now record low-grade metamorphism. The detrital formation contains layers of acidic rocks with an age of 269±9 Ma and the overlying marbles are assumed to be Triassic. The existence of these unconformable formations over the peridotites is crucial for the dating of the exhumation of the latter. The presence of peridotite clasts in the detrital formation indicates that peridotites were exposed during the Permian and other data suggest that peridotites were exhumed during the late Carboniferous. During the Alpine cycle, the peridotites operated as an element situated at the bottom of the tectonically higher Alpujarride/Sebtide unit (the Jubrique unit) and forming part of it, then being incorporated to the Alpine thrusts of this unit. (Author)

  2. Early Mycoplasma pneumoniae infection presenting as multiple pulmonary masses: an unusual presentation in a child

    Energy Technology Data Exchange (ETDEWEB)

    Yang, Edward; Altes, Talissa; Anupindi, Sudha A. [The Children' s Hospital of Philadelphia, Department of Radiology, Philadelphia, PA (United States)

    2008-04-15

    Mycoplasma pneumoniae is a major cause of community-acquired pneumonia. Because most children are not imaged prior to onset of clinical symptoms, the appearance of early Mycoplasma infection has not been extensively studied. We present the case of an 11-year-old boy with large pulmonary masses incidentally detected during spine MRI evaluation for scoliosis. Eight days later, the patient developed acute respiratory symptoms, and the masses seen previously had evolved into a diffuse bronchiolitis. Diagnostic testing identified Mycoplasma pneumoniae as the likely etiology. We briefly review chest CT findings of infection by Mycoplasma and compare them to this unusual presentation of Mycoplasma pneumonia with subclinical imaging findings. (orig.)

  3. Early Mycoplasma pneumoniae infection presenting as multiple pulmonary masses: an unusual presentation in a child

    International Nuclear Information System (INIS)

    Yang, Edward; Altes, Talissa; Anupindi, Sudha A.

    2008-01-01

    Mycoplasma pneumoniae is a major cause of community-acquired pneumonia. Because most children are not imaged prior to onset of clinical symptoms, the appearance of early Mycoplasma infection has not been extensively studied. We present the case of an 11-year-old boy with large pulmonary masses incidentally detected during spine MRI evaluation for scoliosis. Eight days later, the patient developed acute respiratory symptoms, and the masses seen previously had evolved into a diffuse bronchiolitis. Diagnostic testing identified Mycoplasma pneumoniae as the likely etiology. We briefly review chest CT findings of infection by Mycoplasma and compare them to this unusual presentation of Mycoplasma pneumonia with subclinical imaging findings. (orig.)

  4. Ground-water conditions in the Triassic aquifer in Deaf Smith and Swisher Counties

    International Nuclear Information System (INIS)

    Duffin, G.L.

    1984-12-01

    In April 1984, the Director of the Nuclear Waste Programs of the Governor's Office requested a study be undertaken by the Texas Department of Water Resources on the ground-water conditions in the Triassic aquifer in Deaf Smith and Swisher Counties. The need for the study was prompted by the U.S. Department of Energy's (DOE) announcement that consideration was being given to locating high-level nuclear waste repository sites in these counties and by the concern over what impacts operation of such sites might have on the ground-water resources in the area. The results of the study, including a discussion of the occurrence of ground water and a tabulation of basic data obtained during the investigation are presented in this report

  5. Sr evolution in the Upper Permian and Lower Triassic carbonates, northeast Sichuan basin, China: Constraints from chemistry, isotope and fluid inclusions

    International Nuclear Information System (INIS)

    Li, Kaikai; Cai, Chunfang; Jiang, Lei; Cai, Liulu; Jia, Lianqi; Zhang, Bing; Xiang, Lei; Yuan, Yuyang

    2012-01-01

    Petrographic features, C, O and Sr isotopes, rare earth and trace elements were determined, and fluid inclusions were analyzed on various stages of interparticle cements and vug-fillings from the Upper Permian and Lower Triassic sour reservoirs in northeastern Sichuan basin. The aim was to assess the origin and evolution of palaeo-waters in the carbonates. The original water was contemporary seawater, from which marine cements precipitated with slightly high Sr contents (mean 1911 ppm), 87 Sr/ 86 Sr ratios from 0.7067 to 0.7082 and nonluminescent CL. The palaeo-seawater was diluted by meteoric water, as indicated by bright cathodoluminescence (CL) and Sr-depletion (0–516 ppm) in low-temperature calcite. When buried to temperatures of about 60–90 °C during Middle to Late Triassic, the palaeo-water was enriched in Sr released from the transformation of precursor aragonite and calcite to dolomite, resulting in precipitation of substantial pre-bitumen Sr-rich minerals (SrSO 4 and SrCO 3 ). For un-dolomitized limestone sections, aragonite neomorphism may have contributed Sr to the precipitation of small amounts of Sr-bearing minerals and calcite crystals with elevated homogenization temperatures (HTs, mainly from 90 to 130 °C) and wide Sr contents (from 34 to 3825 ppm), as recorded in stage III calcite. Since the Middle Jurassic, almost all of the early stage celestite and significant amounts of solid CaSO 4 have been consumed by reactions with hydrocarbons (i.e., TSR), resulting in water enriched in isotopically light CO 2 and HCO 3 - ,Sr 2+ ,Ba 2+ and Eu 2+ , as recorded in calcite with low δ 13 C values (down to −18.9‰), 87 Sr/ 86 Sr ratios from 0.7072 to 0.7076, high HTs (mainly 110–198 °C), positive Eu anomalies and high Sr and Ba contents. Subsequently, the water was uplifted and cooled down to about 115 °C, celestite and strontianite were precipitated with the occurrence of natural elemental S immiscible inclusions. TSR may have produced

  6. EVIDENCE FOR LADINIAN (MIDDLE TRIASSIC PLATFORM PROGRADATION IN THE GYULAKESZI AREA, TAPOLCA BASIN, WESTERN HUNGARY: MICROFACIES ANALYSIS AND BIOSTRATIGRAPHY

    Directory of Open Access Journals (Sweden)

    ZSOLT RÓBERT NAGY

    2014-07-01

    Full Text Available A shallowing-upward carbonate sequence was studied from the outcrop at Gyulakeszi, Tapolca Basin (western Hungary, and it is interpreted as a Middle Triassic (Curionii or younger platform progradation. Two lithostratigraphic units are distinguished. Microfacies analysis and micropaleontological investigation conducted on the red nodular, cherty limestone (Vászoly and Buchenstein formations suggest that the lower unit was deposited during the Reitzi and the Secedensis ammonoid zones. The overlying white platform limestone (upper unit is typical of a prograding platform and includes gravity-driven deposits at the base followed by periplatform facies deposited in shallow marine warm waters around the fair-weather wave base. The section at Gyulakeszi was unaffected by fabric-destructive dolomitization, which is uncharacteristic of similar platform facies in the Balaton Highland. Isopachous and radiaxial fibrous calcite cement found in the grainstone and boundstone facies are indicative of early lithification and diagenesis in the marine phreatic zone. “Evinospongiae”-type cement is described for the first time from the Balaton Highland and it is similar to the outer platform cements published previously from the Alps (Italy and Austria. The progradation could have advanced over the pelagic limestones as early as the Curionii zone, which is an undocumented event in the Veszprém Plateau. Similar event, however, is well known from the Western Dolomites, where aggradation was followed by intense progradation during the Gredleri and Archelaus ammonoid zones. The length of this progradation event at Gyulakeszi, however, is ambiguous since proven Ladinian (Longobardian rocks are not exposed in the study area and were not penetrated by boreholes in the Tapolca Basin.

  7. Sedimentology of the Upper Triassic-Lower Jurassic (?) Mosolotsane Formation (Karoo Supergroup), Kalahari Karoo Basin, Botswana

    Science.gov (United States)

    Bordy, Emese M.; Segwabe, Tebogo; Makuke, Bonno

    2010-08-01

    The Mosolotsane Formation (Lebung Group, Karoo Supergroup) in the Kalahari Karoo Basin of Botswana is a scantly exposed, terrestrial red bed succession which is lithologically correlated with the Late Triassic to Early Jurassic Molteno and Elliot Formations (Karoo Supergroup) in South Africa. New evidence derived from field observations and borehole data via sedimentary facies analysis allowed the assessment of the facies characteristics, distribution and thickness variation as well as palaeo-current directions and sediment composition, and resulted in the palaeo-environmental reconstruction of this poorly known unit. Our results show that the Mosolotsane Formation was deposited in a relatively low-sinuosity meandering river system that drained in a possibly semi-arid environment. Sandstone petrography revealed mainly quartz-rich arenites that were derived from a continental block provenance dominated by metamorphic and/or igneous rocks. Palaeo-flow measurements indicate reasonably strong, unidirectional current patterns with mean flow directions from southeast and east-southeast to northwest and west-northwest. Regional thickness and facies distributions as well as palaeo-drainage indicators suggest that the main depocenter of the Mosolotsane Formation was in the central part of the Kalahari Karoo Basin. Separated from this main depocenter by a west-northwest - east-southeast trending elevated area, an additional depocenter was situated in the north-northeast part of the basin and probably formed part of the Mid-Zambezi Karoo Basin. In addition, data also suggests that further northeast-southwest trending uplands probably existed in the northwest and east, the latter separating the main Kalahari Karoo depocenter from the Tuli Basin.

  8. Petrogenesis and tectonic implication of the Late Triassic post-collisional volcanic rocks in Chiang Khong, NW Thailand

    Science.gov (United States)

    Qian, Xin; Wang, Yuejun; Feng, Qinglai; Zi, Jian-Wei; Zhang, Yuzhi; Chonglakmani, Chongpan

    2016-04-01

    The volcanic rocks exposed within the Chiang Khong-Lampang-Tak igneous zone in NW Thailand provide important constraints on the tectonic evolution of the eastern Paleotethys ocean. An andesite sample from the Chiang Khong area yields a zircon U-Pb age of 229 ± 4 Ma, significantly younger than the continental-arc and syn-collisional volcanic rocks (ca. 238-241 Ma). The Chiang Khong volcanic rocks are characterized by low MgO (1.71-6.72 wt.%) and high Al2O3 (15.03-17.76 wt.%). They are enriched in LILEs and LREEs and depleted in HFSEs, and have 87Sr/86Sr (i) ratios of 0.7050-0.7065, εNd (t) of - 0.32 to - 1.92, zircon εHf (t) and δ18O values of 3.5 to - 11.7 and 4.30-9.80 ‰, respectively. The geochemical data for the volcanic rocks are consistent with an origin from the enriched lithospheric mantle that had been modified by slab-derived fluid and recycled sediments. Based on available geochronological and geochemical evidences, we propose that the Late Triassic Chiang Khong volcanic rocks are equivalent to the contemporaneous volcanic rocks in the Lancangjiang igneous zone in SW China. The formation of these volcanic rocks was possibly related to the upwelling of the asthenospheric mantle during the Late Triassic, shortly after slab detachment, which induced the melting of the metasomatized mantle wedge.

  9. Reassessment of the evidence for postcranial skeletal pneumaticity in Triassic archosaurs, and the early evolution of the avian respiratory system.

    Directory of Open Access Journals (Sweden)

    Richard J Butler

    Full Text Available Uniquely among extant vertebrates, birds possess complex respiratory systems characterised by the combination of small, rigid lungs, extensive pulmonary air sacs that possess diverticula that invade (pneumatise the postcranial skeleton, unidirectional ventilation of the lungs, and efficient crosscurrent gas exchange. Crocodilians, the only other living archosaurs, also possess unidirectional lung ventilation, but lack true air sacs and postcranial skeletal pneumaticity (PSP. PSP can be used to infer the presence of avian-like pulmonary air sacs in several extinct archosaur clades (non-avian theropod dinosaurs, sauropod dinosaurs and pterosaurs. However, the evolution of respiratory systems in other archosaurs, especially in the lineage leading to crocodilians, is poorly documented. Here, we use µCT-scanning to investigate the vertebral anatomy of Triassic archosaur taxa, from both the avian and crocodilian lineages as well as non-archosaurian diapsid outgroups. Our results confirm previous suggestions that unambiguous evidence of PSP (presence of internal pneumatic cavities linked to the exterior by foramina is found only in bird-line (ornithodiran archosaurs. We propose that pulmonary air sacs were present in the common ancestor of Ornithodira and may have been subsequently lost or reduced in some members of the clade (notably in ornithischian dinosaurs. The development of these avian-like respiratory features might have been linked to inferred increases in activity levels among ornithodirans. By contrast, no crocodile-line archosaur (pseudosuchian exhibits evidence for unambiguous PSP, but many of these taxa possess the complex array of vertebral laminae and fossae that always accompany the presence of air sacs in ornithodirans. These laminae and fossae are likely homologous with those in ornithodirans, which suggests the need for further investigation of the hypothesis that a reduced, or non-invasive, system of pulmonary air sacs may be have

  10. Body Mass Index Trajectories from Adolescence to Early Young Adulthood : Do Adverse Life Events Play a Role?

    NARCIS (Netherlands)

    Elsenburg, Leonie K.; Smidt, Nynke; Hoek, Hans W.; Liefbroer, Aart C.

    2017-01-01

    Objective: The aim of this study was to investigate whether there are different classes of body mass index (BMI) development from early adolescence to young adulthood and whether these classes are related to the number of adverse life events children experienced. Methods: Data were from the TRAILS

  11. Evolutionary reversion of editing sites of ndh genes suggests their origin in the Permian-Triassic, before the increase of atmospheric CO2

    Directory of Open Access Journals (Sweden)

    Mercedes eMartin

    2015-07-01

    Full Text Available The plastid ndh genes have hovered frequently on the edge of dispensability. They are absent in the plastid DNA of many algae and certain higher plants and present editing sites requiring C-to-U corrections of primary transcripts. The evolutionary origin of editing sites and their loss due to C-to-T reversions at the DNA level are unknown and must be related to the dispensability of the ndh genes in specific environments. In order to better understand the evolution of ndh gene editing sites, we have created expandable data banks with the 12 editing sites of the ndhB gene (600 GenBank sequences and both editing sites of the ndhF gene (1,600 GenBank sequences. Since their origin via T-to-C mutations that probably occurred between 300 and 200 Myr BP (Permian-Triassic, ndh editing sites have undergone independent and random C-to-T reversions in the different angiosperm lineages. Some of these reversions appear early in angiosperm diversification. Old C-to-T reversions can be traced back to radiation steps that gave origin to main classes, orders and some families.

  12. Dark matter contraction and stellar-mass-to-light ratio gradients in massive early-type galaxies

    Science.gov (United States)

    Oldham, Lindsay J.; Auger, Matthew W.

    2018-05-01

    We present models for the dark and luminous mass structure of 12 strong lensing early-type galaxies. We combine pixel-based modelling of multiband Hubble Space Telescope imaging with Jeans modelling of kinematics obtained from Keck/ESI spectra to disentangle the dark and luminous contributions to the mass. Assuming a generalised NFW (gNFW) profile for the dark matter halo and a spatially constant stellar-mass-to-light ratio ϒ⋆ for the baryonic mass, we infer distributions for ϒ⋆ consistent with initial mass functions (IMFs) that are heavier than the Milky Way's (with a global mean mismatch parameter relative to a Chabrier IMF μαc = 1.80 ± 0.14) and halo inner density slopes that span a large range but are generally cuspier than the dark-matter-only prediction (μ _{γ ^' }} = 2.01_{-0.22}^{+0.19}). We investigate possible reasons for overestimating the halo slope, including the neglect of spatially varying stellar-mass-to-light ratios and/or stellar orbital anisotropy, and find that a quarter of the systems prefer radially declining stellar-mass-to-light ratio gradients, but that the overall effect on our inference on the halo slope is small. We suggest a coherent explanation of these results in the context of inside-out galaxy growth, and that the relative importance of different baryonic processes in shaping the dark halo may depend on halo environment.

  13. Early efficacy of the ketogenic diet is not affected by initial body mass index percentile.

    Science.gov (United States)

    Shull, Shastin; Diaz-Medina, Gloria; Wong-Kisiel, Lily; Nickels, Katherine; Eckert, Susan; Wirrell, Elaine

    2014-05-01

    Predictors of the ketogenic diet's success in treating pediatric intractable epilepsy are not well understood. The aim of this study was to determine whether initial body mass index and weight percentile impact early efficacy of the traditional ketogenic diet in children initiating therapy for intractable epilepsy. This retrospective study included all children initiating the ketogenic diet at Mayo Clinic, Rochester from January 2001 to December 2010 who had body mass index (children ≥2 years of age) or weight percentile (those diet initiation and seizure frequency recorded at diet initiation and one month. Responders were defined as achieving a >50% seizure reduction from baseline. Our cohort consisted of 48 patients (20 male) with a median age of 3.1 years. There was no significant correlation between initial body mass index or weight percentile and seizure frequency reduction at one month (P = 0.72, r = 0.26 and P = 0.91, r = 0.03). There was no significant association between body mass index or weight percentile quartile and responder rates (P = 0.21 and P = 0.57). Children considered overweight or obese at diet initiation (body mass index or weight percentile ≥85) did not have lower responder rates than those with body mass index or weight percentiles ketogenic diet. Copyright © 2014 Elsevier Inc. All rights reserved.

  14. The importance of Hindeodus parvus (Conodonta for the definition of the Permian-Triassic boundary and evaluation of the proposed sections for a global stratotype section and point (GSSP for the base of the Triassic

    Directory of Open Access Journals (Sweden)

    Yurij D. Zakharov

    1995-12-01

    Full Text Available The biostratigraphic Permian/Triassic (P/T boundary is defined by the first appearance of H. parvus. The first appearance of H. parvus within the dine H. latidentatus-H. parvus is a globally recognizable event in the conodont evolution.The first appearance of H. parvus is not facies related and can be observed both in ammonoid-free shallow-water deposits and in ammonoid-bearing pelagic deposits. H. parvus is a common, easily determinable species known so far from the entire Tethys, Japan, western North America, Boreal realm (Greenlandand the Tethyan margin of Gondwana. H. parvus is the first species withworld-wide distribution to appear after the absolute minimum in the faunal diversity indicated by the minimum in The Meishan section (South China contains a continuous, pelagic sedimentaryrecord across the P/T boundary without stratigraphie gaps. It is nearly unaltered thermally (CAI = 1-1.5. Its fossil content (ammonoids, conodonts, foraminifers,bivalves, brachiopods, sporomorphs etc. and event succession have been thoroughlystudied. Absolute age and magnetostratigraphy have also been subjected to intensive studies. The section is readily accessible and under protection of the government. This section is best suitable as a global stratotype section and point (GSSP for the base of the Triassic. No other section in the world is known tobe qualified for defining the P/T boundary in a GSSP. H. parvus made its earliest appearence in the middle part of Boundary Bed 2 (Bed 27 at Meishan. It evolved within Bed 27 from H. latidentatus within a phylomorphogenetic continuum in a continuous and monofacial stratum. The biostratigraphic P/T boundary lies very close to the event boundary (15 cm above the event boundary at the baseof Boundary Bed 1 = Bed 25, and a few centimetres above the minimum in Ô'13C in the lower Boundary Bed 2.

  15. Synchrotron microtomography of a Nothosaurus marchicus skull informs on nothosaurian physiology and neurosensory adaptations in early Sauropterygia.

    Directory of Open Access Journals (Sweden)

    Dennis F A E Voeten

    Full Text Available Nothosaurs form a subclade of the secondarily marine Sauropterygia that was well represented in late Early to early Late Triassic marine ecosystems. Here we present and discuss the internal skull anatomy of the small piscivorous nothosaur Nothosaurus marchicus from coastal to shallow marine Lower Muschelkalk deposits (Anisian of Winterswijk, The Netherlands, which represents the oldest sauropterygian endocast visualized to date. The cranial endocast is only partially encapsulated by ossified braincase elements. Cranial flattening and lateral constriction by hypertrophied temporal musculature grant the brain a straight, tubular geometry that lacks particularly well-developed cerebral lobes but does potentially involve distinguishable optic lobes, suggesting vision may have represented an important sense during life. Despite large orbit size, the circuitous muscular pathway linking the basisphenoidal and orbital regions indicates poor oculomotor performance. This suggests a rather fixed ocular orientation, although eye placement and neck manoeuvrability could have enabled binocular if not stereoscopic vision. The proportionally large dorsal projection of the braincase endocast towards the well-developed pineal foramen advocates substantial dependence on the corresponding pineal system in vivo. Structures corroborating keen olfactory or acoustic senses were not identified. The likely atrophied vomeronasal organ argues against the presence of a forked tongue in Nothosaurus, and the relative positioning of external and internal nares contrasts respiratory configurations proposed for pistosauroid sauropterygians. The antorbital domain furthermore accommodates a putative rostral sensory plexus and pronounced lateral nasal glands that were likely exapted as salt glands. Previously proposed nothosaurian 'foramina eustachii' arose from architectural constraints on braincase development rather than representing functional foramina. Several modifications to

  16. Sedimentary noise and sea levels linked to land-ocean water exchange and obliquity forcing.

    Science.gov (United States)

    Li, Mingsong; Hinnov, Linda A; Huang, Chunju; Ogg, James G

    2018-03-08

    In ancient hothouses lacking ice sheets, the origins of large, million-year (myr)-scale sea-level oscillations remain a mystery, challenging current models of sea-level change. To address this mystery, we develop a sedimentary noise model for sea-level changes that simultaneously estimates geologic time and sea level from astronomically forced marginal marine stratigraphy. The noise model involves two complementary approaches: dynamic noise after orbital tuning (DYNOT) and lag-1 autocorrelation coefficient (ρ 1 ). Noise modeling of Lower Triassic marine slope stratigraphy in South China reveal evidence for global sea-level variations in the Early Triassic hothouse that are anti-phased with continental water storage variations in the Germanic Basin. This supports the hypothesis that long-period (1-2 myr) astronomically forced water mass exchange between land and ocean reservoirs is a missing link for reconciling geological records and models for sea-level change during non-glacial periods.

  17. A basal thunnosaurian from Iraq reveals disparate phylogenetic origins for Cretaceous ichthyosaurs

    Science.gov (United States)

    Fischer, Valentin; Appleby, Robert M.; Naish, Darren; Liston, Jeff; Riding, James B.; Brindley, Stephen; Godefroit, Pascal

    2013-01-01

    Cretaceous ichthyosaurs have typically been considered a small, homogeneous assemblage sharing a common Late Jurassic ancestor. Their low diversity and disparity have been interpreted as indicative of a decline leading to their Cenomanian extinction. We describe the first post-Triassic ichthyosaur from the Middle East, Malawania anachronus gen. et sp. nov. from the Early Cretaceous of Iraq, and re-evaluate the evolutionary history of parvipelvian ichthyosaurs via phylogenetic and cladogenesis rate analyses. Malawania represents a basal grade in thunnosaurian evolution that arose during a major Late Triassic radiation event and was previously thought to have gone extinct during the Early Jurassic. Its pectoral morphology appears surprisingly archaic, retaining a forefin architecture similar to that of its Early Jurassic relatives. After the initial latest Triassic radiation of early thunnosaurians, two subsequent large radiations produced lineages with Cretaceous representatives, but the radiation events themselves are pre-Cretaceous. Cretaceous ichthyosaurs therefore include distantly related lineages, with contrasting evolutionary histories, and appear more diverse and disparate than previously supposed. PMID:23676653

  18. Early-life bisphenol a exposure and child body mass index: a prospective cohort study.

    Science.gov (United States)

    Braun, Joseph M; Lanphear, Bruce P; Calafat, Antonia M; Deria, Sirad; Khoury, Jane; Howe, Chanelle J; Venners, Scott A

    2014-11-01

    Early-life exposure to bisphenol A (BPA) may increase childhood obesity risk, but few prospective epidemiological studies have investigated this relationship. We sought to determine whether early-life exposure to BPA was associated with increased body mass index (BMI) at 2-5 years of age in 297 mother-child pairs from Cincinnati, Ohio (HOME Study). Urinary BPA concentrations were measured in samples collected from pregnant women during the second and third trimesters and their children at 1 and 2 years of age. BMI z-scores were calculated from weight/height measures conducted annually from 2 through 5 years of age. We used linear mixed models to estimate BMI differences or trajectories with increasing creatinine-normalized BPA concentrations. After confounder adjustment, each 10-fold increase in prenatal (β = -0.1; 95% CI: -0.5, 0.3) or early-childhood (β = -0.2; 95% CI: -0.6, 0.1) BPA concentrations was associated with a modest and nonsignificant reduction in child BMI. These inverse associations were suggestively stronger in girls than in boys [prenatal effect measure modification (EMM) p-value = 0.30, early-childhood EMM p-value = 0.05], but sex-specific associations were imprecise. Children in the highest early-childhood BPA tercile had lower BMI at 2 years (difference = -0.3; 95% CI: -0.6, 0.0) and larger increases in their BMI slope from 2 through 5 years (BMI increase per year = 0.12; 95% CI: 0.07, 0.18) than children in the lowest tercile (BMI increase per year = 0.07; 95% CI: 0.01, 0.13). All associations were attenuated without creatinine normalization. Prenatal and early-childhood BPA exposures were not associated with increased BMI at 2-5 years of age, but higher early-childhood BPA exposures were associated with accelerated growth during this period.

  19. SDSS-IV MaNGA: global stellar population and gradients for about 2000 early-type and spiral galaxies on the mass-size plane

    Science.gov (United States)

    Li, Hongyu; Mao, Shude; Cappellari, Michele; Ge, Junqiang; Long, R. J.; Li, Ran; Mo, H. J.; Li, Cheng; Zheng, Zheng; Bundy, Kevin; Thomas, Daniel; Brownstein, Joel R.; Roman Lopes, Alexandre; Law, David R.; Drory, Niv

    2018-05-01

    We perform full spectrum fitting stellar population analysis and Jeans Anisotropic modelling of the stellar kinematics for about 2000 early-type galaxies (ETGs) and spiral galaxies from the MaNGA DR14 sample. Galaxies with different morphologies are found to be located on a remarkably tight mass plane which is close to the prediction of the virial theorem, extending previous results for ETGs. By examining an inclined projection (`the mass-size' plane), we find that spiral and early-type galaxies occupy different regions on the plane, and their stellar population properties (i.e. age, metallicity, and stellar mass-to-light ratio) vary systematically along roughly the direction of velocity dispersion, which is a proxy for the bulge fraction. Galaxies with higher velocity dispersions have typically older ages, larger stellar mass-to-light ratios and are more metal rich, which indicates that galaxies increase their bulge fractions as their stellar populations age and become enriched chemically. The age and stellar mass-to-light ratio gradients for low-mass galaxies in our sample tend to be positive (centre Spiral galaxies with large mass and size have the steepest gradients, while the most massive ETGs, especially above the critical mass Mcrit ≳ 2 × 1011 M⊙, where slow rotator ETGs start dominating, have much flatter gradients. This may be due to differences in their evolution histories, e.g. mergers.

  20. The Triassic eucynodont Candelariodon barberenai revisited and the early diversity of stem prozostrodontians

    Directory of Open Access Journals (Sweden)

    Agustín G. Martinelli

    2017-09-01

    Full Text Available The dental anatomy of Candelariodon barberenai from the Dinodontosaurus Assemblage Zone (Pinheiros-Chiniquá Sequence, Santa Maria Supersequence, late Ladinian–early Carnian of south Brazil, is redescribed. Candelariodon was originally classified as Eucynodontia incertae sedis and our analysis recovered this taxon deeply nested within Probainognathia, as the sister taxon of Potheriodon plus Prozostrodontia. The lower postcanine dentition of Candelariodon has several apomorphies shared with Prozostrodon, Santacruzgnathus, Brasilodon/Brasilitherium, and some basal mammaliaforms (Morganucodon, Megazostrodon, such as a lingual cingulum with discrete cusps e and g and two distinct morphologies in the tooth row. The reinterpretation of Candelariodon as a probainognatian cynodont more derived than Probainognathus and the rich Brazilian fossil record document an important adaptive radiation of non-mammaliaform prozostrodontians and closely related forms prior to the origin of the mammaliaform clade.

  1. The phylogenetic relationships of basal archosauromorphs, with an emphasis on the systematics of proterosuchian archosauriforms

    Science.gov (United States)

    2016-01-01

    The early evolution of archosauromorphs during the Permo-Triassic constitutes an excellent empirical case study to shed light on evolutionary radiations in deep time and the timing and processes of recovery of terrestrial faunas after a mass extinction. However, macroevolutionary studies of early archosauromorphs are currently limited by poor knowledge of their phylogenetic relationships. In particular, one of the main early archosauromorph groups that need an exhaustive phylogenetic study is “Proterosuchia,” which as historically conceived includes members of both Proterosuchidae and Erythrosuchidae. A new data matrix composed of 96 separate taxa (several of them not included in a quantitative phylogenetic analysis before) and 600 osteological characters was assembled and analysed to generate a comprehensive higher-level phylogenetic hypothesis of basal archosauromorphs and shed light on the species-level interrelationships of taxa historically identified as proterosuchian archosauriforms. The results of the analysis using maximum parsimony include a polyphyletic “Prolacertiformes” and “Protorosauria,” in which the Permian Aenigmastropheus and Protorosaurus are the most basal archosauromorphs. The enigmatic choristoderans are either found as the sister-taxa of all other lepidosauromorphs or archosauromorphs, but consistently placed within Sauria. Prolacertids, rhynchosaurs, allokotosaurians and tanystropheids are the major successive sister clades of Archosauriformes. The Early Triassic Tasmaniosaurus is recovered as the sister-taxon of Archosauriformes. Proterosuchidae is unambiguosly restricted to five species that occur immediately after and before the Permo-Triassic boundary, thus implying that they are a short-lived “disaster” clade. Erythrosuchidae is composed of eight nominal species that occur during the Early and Middle Triassic. “Proterosuchia” is polyphyletic, in which erythrosuchids are more closely related to Euparkeria and more

  2. Elevated Extinction Rates as a Trigger for Diversification Rate Shifts: Early Amniotes as a Case Study

    Science.gov (United States)

    Brocklehurst, Neil; Ruta, Marcello; Müller, Johannes; Fröbisch, Jörg

    2015-01-01

    Tree shape analyses are frequently used to infer the location of shifts in diversification rate within the Tree of Life. Many studies have supported a causal relationship between shifts and temporally coincident events such as the evolution of “key innovations”. However, the evidence for such relationships is circumstantial. We investigated patterns of diversification during the early evolution of Amniota from the Carboniferous to the Triassic, subjecting a new supertree to analyses of tree balance in order to infer the timing and location of diversification shifts. We investigated how uneven origination and extinction rates drive diversification shifts, and use two case studies (herbivory and an aquatic lifestyle) to examine whether shifts tend to be contemporaneous with evolutionary novelties. Shifts within amniotes tend to occur during periods of elevated extinction, with mass extinctions coinciding with numerous and larger shifts. Diversification shifts occurring in clades that possess evolutionary innovations do not coincide temporally with the appearance of those innovations, but are instead deferred to periods of high extinction rate. We suggest such innovations did not cause increases in the rate of cladogenesis, but allowed clades to survive extinction events. We highlight the importance of examining general patterns of diversification before interpreting specific shifts. PMID:26592209

  3. Early to Middle Jurassic tectonic evolution of the Bogda Mountains, Northwest China: Evidence from sedimentology and detrital zircon geochronology

    Science.gov (United States)

    Ji, Hongjie; Tao, Huifei; Wang, Qi; Qiu, Zhen; Ma, Dongxu; Qiu, Junli; Liao, Peng

    2018-03-01

    The Bogda Mountains, as an important intracontinental orogenic belt, are situated in the southern part of the Central Asian Orogenic Belt (CAOB), and are a key area for understanding the Mesozoic evolution of the CAOB. However, the tectonic evolution of the Bogda Mountains remains controversial during the Mesozoic Era, especially the Early to Middle Jurassic Periods. The successive Lower to Middle Jurassic strata are well preserved and exposed along the northern flank of the Western Bogda Mountains and record the uplift processes of the Bogda Mountains. In this study, we analysed sedimentary facies combined with detrital zircon U-Pb geochronology at five sections of Lower to Middle Jurassic strata to detect the tectonic evolution and changes of provenance in the Bogda area. During Early to Middle Jurassic times, the fluvial, deltaic and lacustrine environments dominated in the western section of the Bogda area. The existence of Early Triassic peak age indicates that the Bogda Mountains did not experience uplift during the period of early Badaowan Formation deposition. The Early Triassic to Late Permian granitoid plutons and Carboniferous volcanic rocks from the Barkol and Santanghu areas were the main provenances. The significant change in the U-Pb age spectrum implies that the Eastern Bogda Mountains initiated uplift in the period of late Badaowan Formation deposition, and the Eastern Junggar Basin and the Turpan-Hami Basin were partially partitioned. The Eastern Bogda Mountains gradually became the major provenance. From the period of early Sangonghe to early Toutunhe Formations deposition, the provenance of the sediments and basin-range frame were similar to that of late Badaowan. However, the Eastern Bogda Mountains suffered intermittent uplift three times, and successive denudation. The uplifts respectively happened in early Sangonghe, late Sangonghe to early Xishanyao, and late Xishanyao to early Toutunhe. During the deposition stage of Toutunhe Formation, a

  4. Siliciclastics in the Upper Triassic dolomite formations of the Krizna Unit (Maid Fatra Mountains, Western Carpathians): constraints for the Carnian Pluvial Event in the Fatric Basin

    Czech Academy of Sciences Publication Activity Database

    Sýkora, M.; Siblík, Miloš; Soták, J.

    2011-01-01

    Roč. 62, č. 2 (2011), s. 121-138 ISSN 1335-0552 Institutional research plan: CEZ:AV0Z30130516 Keywords : Carnian event * Upper Triassic dolomites * Western Carpathians * Krizna Unit * shaly interbeds * lingulids * conchostracans Subject RIV: DB - Geology ; Mineralogy Impact factor: 0.787, year: 2011

  5. DARK MATTER CONTRACTION AND THE STELLAR CONTENT OF MASSIVE EARLY-TYPE GALAXIES: DISFAVORING 'LIGHT' INITIAL MASS FUNCTIONS

    International Nuclear Information System (INIS)

    Auger, M. W.; Treu, T.; Gavazzi, R.; Bolton, A. S.; Koopmans, L. V. E.; Marshall, P. J.

    2010-01-01

    We use stellar dynamics, strong lensing, stellar population synthesis models, and weak lensing shear measurements to constrain the dark matter (DM) profile and stellar mass in a sample of 53 massive early-type galaxies. We explore three DM halo models (unperturbed Navarro, Frenk, and White (NFW) halos and the adiabatic contraction models of Blumenthal and Gnedin) and impose a model for the relationship between the stellar and virial mass (i.e., a relationship for the star formation efficiency as a function of halo mass). We show that, given our model assumptions, the data clearly prefer a Salpeter-like initial mass function (IMF) over a lighter IMF (e.g., Chabrier or Kroupa), irrespective of the choice of DM halo. In addition, we find that the data prefer at most a moderate amount of adiabatic contraction (Blumenthal adiabatic contraction is strongly disfavored) and are only consistent with no adiabatic contraction (i.e., an NFW halo) if a mass-dependent IMF is assumed, in the sense of a more massive normalization of the IMF for more massive halos.

  6. Implications of diapir-derived detritus and gypsic paleosols in Lower Triassic strata near the Castle Valley salt wall, Paradox Basin, Utah

    Science.gov (United States)

    Lawton, Timothy F.; Buck, Brenda J.

    2006-10-01

    Gypsum-bearing growth strata and sedimentary facies of the Moenkopi Formation on the crest and NE flank of the Castle Valley salt wall in the Paradox Basin record salt rise, evaporite exposure, and salt-withdrawal subsidence during the Early Triassic. Detrital gypsum and dolomite clasts derived from the middle Pennsylvanian Paradox Formation were deposited in strata within a few kilometers of the salt wall and indicate that salt rise rates roughly balanced sediment accumulation, resulting in long-term exposure of mobile evaporite. Deposition took place primarily in flood-basin or inland sabkha settings that alternated between shallow subaqueous and subaerial conditions in a hyperarid climate. Matrix-supported and clast-supported conglomerates with gypsum fragments represent debris-flow deposits and reworked debris-flow deposits, respectively, interbedded with flood-basin sandstone and siltstone during development of diapiric topography. Mudstone-rich flood-basin deposits with numerous stage I to III gypsic paleosols capped by eolian gypsum sand sheets accumulated during waning salt-withdrawal subsidence. Association of detrital gypsum, eolian gypsum, and gypsic paleosols suggests that the salt wall provided a common source for gypsum in the surrounding strata. This study documents a previously unrecognized salt weld with associated growth strata containing diapir-derived detritus and gypsic palesols that can be used to interpret halokinesis.

  7. Osteology of the Late Triassic aetosaur Scutarx deltatylus (Archosauria: Pseudosuchia

    Directory of Open Access Journals (Sweden)

    William G. Parker

    2016-08-01

    Full Text Available Aetosaurians are some of the most common fossils collected from the Upper Triassic Chinle Formation of Arizona, especially at the Petrified Forest National Park (PEFO. Aetosaurians collected from lower levels of the park include Desmatosuchus spurensis, Paratypothorax, Adamanasuchus eisenhardtae, Calyptosuchus wellesi, and Scutarx deltatylus. Four partial skeletons collected from the park between 2002 and 2009 represent the holotype and referred specimens of Scutarx deltatylus. These specimens include much of the carapace, as well as the vertebral column, and shoulder and pelvic girdles, and a new naming convention proposed for osteoderms descriptions better differentiates portions of the carapace and ventral armor. A partial skull from the holotype specimen represents the first aetosaur skull recovered and described from Arizona since the 1930s. The key morphological feature distinguishing Scutarx deltatylus is the presence of a prominent, triangular boss located in the posteromedial corner of the dorsal surface of the dorsal paramedian osteoderms. Scutarx deltatylus can be distinguished from closely related forms Calyptosuchus wellesi and Adamanasuchus eisenhardtae not only morphologically, but also stratigraphically. Thus, Scutarx deltatylus is potentially an index taxon for the upper part of the Adamanian biozone.

  8. MIDDLE TRIASSIC PLATFORM AND BASIN EVOLUTION OF THE SOUTHERN BAKONY MOUNTAINS (TRANSDANUBIAN RANGE, HUNGARY

    Directory of Open Access Journals (Sweden)

    TAMÁS BUDAI

    2006-11-01

    Full Text Available Middle Triassic history of the Southern Bakony Mts. is outlined on the base of horizontal and vertical facies changes of the formations. During the Pelsonian (Balatonicus Chron the evolution of the basins and platforms was determined basically by synsedimentary tectonics. The Felsõörs basin of the Balaton Highland opened due to the block-faulting of the Bithynian carbonate ramp (Megyehegy Dolomite. Above the drowning blocks „halfgraben” basins were formed (Felsõörs Formation, while isolated platforms developed on the uplifted ones in the middle part of the Balaton Highland and on the Veszprém plateau (Tagyon Formation. Due to the relative sea-level fall in the early Illyrian, the platforms became subaerially exposed and karstified. As a consequence of the late Illyrian tectonic subsidence (manifested by neptunian dykes the central platform of the Balaton Highland has been drowned (Camunum Subchron. On the contrary, the Anisian platform of the Veszprém plateau was totally flooded only during the latest Illyrian (Reitzi Subchron due to eustatic sea-level rise. It was followed by a short highstand period (Secedensis Chron, characterised by the first progradation of the Budaörs platform on the Veszprém plateau and highstand shedding in the basins and on the submarine high (Vászoly Limestone in the centre of the Balaton Highland basin. Due to the following rapid sea-level rise, carbonate sedimentation continued in eupelagic basin from the Fassanian (Buchenstein Formation. At the beginning of the late Longobardian highstand period (Regoledanus Chron the Budaörs platform intensively prograded from the Veszprém plateau to the southwest, causing highstand shedding in the Balaton Highland basin (Füred Limestone. 

  9. Early changes in socioeconomic status do not predict changes in body mass in the first decade of life.

    Science.gov (United States)

    Starkey, Leighann; Revenson, Tracey A

    2015-04-01

    Many studies link childhood socioeconomic status (SES) to body mass index (BMI), but few account for the impact of socioeconomic mobility throughout the lifespan. This study aims to investigate the impact of socioeconomic mobility on changes in BMI in childhood. Analyses tested whether [1] socioeconomic status influences BMI, [2] changes in socioeconomic status impact changes in BMI, and [3] timing of socioeconomic status mobility impacts BMI. Secondary data spanning birth to age 9 were analyzed. SES and BMI were investigated with gender, birth weight, maternal race/ethnicity, and maternal nativity as covariates. Autoregressive structural equation modeling and latent growth modeling were used. Socioeconomic status in the first year of life predicted body mass index. Child covariates were consistently associated with body mass index. Rate of change in socioeconomic status did not predict change in body mass index. The findings suggest that early socioeconomic status may most influence body mass in later childhood.

  10. Mass extinctions: Ecological selectivity and primary production

    Science.gov (United States)

    Rhodes, Melissa Clark; Thayer, Charles W.

    1991-09-01

    If mass extinctions were caused by reduced primary productivity, then extinctions should be concentrated among animals with starvation-susceptible feeding modes, active lifestyles, and high-energy budgets. The stratigraphic ranges (by stage) of 424 genera of bivalves and 309 genera of articulate brachiopods suggest that there was an unusual reduction of primary productivity at the Cretaceous/Tertiary (K/T) boundary extinction. For bivalves at the K/T, there were (1) selective extinction of suspension feeders and other susceptible trophic categories relative to deposit feeders and other resistant categories, and (2) among suspension feed-ers, selective extinction of bivalves with active locomotion. During the Permian-Triassic (P/Tr) extinction and Jurassic background time, extinction rates among suspension feeders were greater for articulate brachiopods than for bivalves. But during the K/T event, extinction rates of articulates and suspension-feeding bivalves equalized, possibly because the low-energy budgets of articulates gave them an advantage when food was scarce.

  11. Palynostratigraphy of Permian succession in the Mand–Raigarh ...

    Indian Academy of Sciences (India)

    Lucknow. References. Bharadwaj D C 1976 Palaeogeograhy of India during Gond- wana time and its bearing on the climate; Geophytology. 6 153–161. Chakraborti B 2001 New palaeobotanical evidence for Early. Triassic to Middle Triassic age for the lower most red bed of Kampthi Formation, Raigarh Gondwana Basin, ...

  12. A Low Mass for Mars from Jupiter's Early Gas-Driven Migration

    Science.gov (United States)

    Walsh, Kevin J.; Morbidelli, Alessandro; Raymond, Sean N.; O'Brien, David P.; Mandell, Avi M.

    2011-01-01

    Jupiter and Saturn formed in a few million years from a gas-dominated protoplanetary disk, and were susceptible to gas-driven migration of their orbits on timescales of only approximately 100,000 years. Hydrodynamic simulations show that these giant planets can undergo a two-stage, inward-then-outward, migration. The terrestrial planets finished accreting much later and their characteristics, including Mars' small mass, are best reproduced by starting from a planetesimal disk with an outer edge at about one astronomical unit from the Sun (1 AU is the Earth-Sun distance). Here we report simulations of the early Solar System that show how the inward migration of Jupiter to 1.5 AU, and its subsequent outward migration, lead to a planetesimal disk truncated at 1 AU; the terrestrial planets then form from this disk over the next 30-50 million years, with an Earth/Mars mass ratio consistent with observations. Scattering by Jupiter initially empties but then repopulates the asteroid belt, with inner-belt bodies originating between 1 and 3 AU and outer-belt bodies originating between and beyond the giant planets. This explains the significant compositional differences across the asteroid belt. The key aspect missing from previous models of terrestrial planet formation is the substantial radial migration of the giant planets, which suggests that their behaviour is more similar to that inferred for extrasolar planets than previously thought.

  13. The SLUGGS survey: a comparison of total-mass profiles of early-type galaxies from observations and cosmological simulations, to ˜4 effective radii

    Science.gov (United States)

    Bellstedt, Sabine; Forbes, Duncan A.; Romanowsky, Aaron J.; Remus, Rhea-Silvia; Stevens, Adam R. H.; Brodie, Jean P.; Poci, Adriano; McDermid, Richard; Alabi, Adebusola; Chevalier, Leonie; Adams, Caitlin; Ferré-Mateu, Anna; Wasserman, Asher; Pandya, Viraj

    2018-06-01

    We apply the Jeans Anisotropic Multi-Gaussian Expansion dynamical modelling method to SAGES Legacy Unifying Globulars and GalaxieS (SLUGGS) survey data of early-type galaxies in the stellar mass range 1010 physical processes shaping the mass distributions of galaxies in cosmological simulations are still incomplete. For galaxies with M* > 1010.7 M⊙ in the Magneticum simulations, we identify a significant anticorrelation between total-mass density profile slopes and the fraction of stellar mass formed ex situ (i.e. accreted), whereas this anticorrelation is weaker for lower stellar masses, implying that the measured total-mass density slopes for low-mass galaxies are less likely to be determined by merger activity.

  14. The Longitudinal Relation Between Accumulation of Adverse Life Events and Body Mass Index From Early Adolescence to Young Adulthood

    NARCIS (Netherlands)

    Elsenburg, Leonie K.; Smidt, Nynke; Liefbroer, Aart C.

    Objective: Stressors, such as adverse life events, can cause weight changes through behavioral and biological mechanisms. Whether the accumulation of adverse life events is related to body mass index (BMI) across multiple time points from early adolescence to young adulthood has not been

  15. The longitudinal relation between accumulation of adverse life events and body mass index from early adolescence to young adulthood

    NARCIS (Netherlands)

    Elsenburg, Leonie K.; Smidt, Nynke; Liefbroer, Aart C.

    2017-01-01

    Objective: Stressors, such as adverse life events, can cause weight changes through behavioral and biological mechanisms. Whether the accumulation of adverse life events is related to body mass index (BMI) across multiple time points from early adolescence to young adulthood has not been

  16. The Longitudinal Relation Between Accumulation of Adverse Life Events and Body Mass Index From Early Adolescence to Young Adulthood

    NARCIS (Netherlands)

    Elsenburg, L.; Smidt, N.; Liefbroer, A.C.

    2017-01-01

    Objective: Stressors, such as adverse life events, can cause weight changes through behavioral and biological mechanisms. Whether the accumulation of adverse life events is related to body mass index (BMI) across multiple time points from early adolescence to young adulthood has not been

  17. THE LATE TRIASSIC ARCHOSAUROMORPH TRILOPHOSAURUS AS AN ARBOREAL CLIMBER

    Directory of Open Access Journals (Sweden)

    JUSTIN A. SPIELMANN

    2005-11-01

    Full Text Available Two species of the unusual archosauromorph Trilophosaurus, T. buettneri Case and T. jacobsi Murry, are known from diverse localities in the Upper Triassic Chinle Group in the southwestern USA. Both species likely occupied similar ecological niches, based on morphological similarities in the postcrania, which are essentially identical. Trilophosaurus occurrences in the Chinle Group are relatively rare, but individual sites are exceptionally rich, suggesting that Trilophosaurus lived in a different paleoenvironment than more typical Chinle vertebrates, which lived in or near streams (phytosaurs, metoposaurs or on floodplains (aetosaurs, rauisuchians, and dinosaurs. Two potential interpretations are that Trilophosaurus was either an arboreal climber or a fossorial digger. However, the gross skeletal features of Trilophosaurus are not compatible with a fossorial mode of life: the limbs are too long and gracile, proximal limb elements are longer than distal ones, and the claws are laterally compressed, not transversely broadened. The intermittent study of Trilophosaurus has caused the theory of it being arboreal, originally proposed by Gregory, to receive little mention in subsequent studies. We reexamined the functional morphology of Trilophosaurus using a qualitative functional morphological analysis of the skeleton, a quantitative examination of claw curvature, and a quantitative examination of manus/trunk and pes/trunk ratios. Claw morphology of Trilophosaurus shows similarities to the arboreal drepanosaurs Drepanosaurus and Megalancosaurus. Our analysis provides ample evidence to suggest that Trilophosaurus was arboreal. 

  18. Under the armor: X-ray computed tomographic reconstruction of the internal skeleton of Coahomasuchus chathamensis (Archosauria: Aetosauria) from the Upper Triassic of North Carolina, USA, and a phylogenetic analysis of Aetosauria.

    Science.gov (United States)

    Hoffman, Devin K; Heckert, Andrew B; Zanno, Lindsay E

    2018-01-01

    Aetosauria is a clade of heavily armored, quadrupedal omnivorous to herbivorous archosaurs known from the Late Triassic across what was the supercontinent of Pangea. Their abundance in many deposits relative to the paucity of other Triassic herbivores indicates that they were key components of Late Triassic ecosystems. However, their evolutionary relationships remain contentious due, in large part, to their extensive dermal armor, which often obstructs observation of internal skeletal anatomy and limits access to potentially informative characters. In an attempt to address this problem we reanalyzed the holotype of a recently described species of Coahomasuchus , C. chathamensis , from the Sanford sub-basin of North Carolina using computed tomography (CT). CT scans of the holotype specimen clarify preservation of the skeleton, revealing several articulated vertebrae and ribs, an isolated vertebra, left ulna, left scapula, and the right humerus, though none of the material resulted in updated phylogenetic scorings. Reexamination of aetosaur materials from the holotype locality also indicates that several isolated osteoderms and elements of the appendicular skeleton are newly referable. Based on these results, we update the Coahomasuchus chathamensis hypodigm and conduct a revised phylogenetic analysis with improved character scorings for Coahomasuchus and several other aetosaurs. Our study recovers Coahomasuchus in a polytomy with Aetosaurus and the Typothoracinae, in contrast with a recent analysis that recovered Coahomasuchus as a wild-card taxon.

  19. Plant–insect interactions from Middle Triassic (late Ladinian of Monte Agnello (Dolomites, N-Italy—initial pattern and response to abiotic environmental perturbations

    Directory of Open Access Journals (Sweden)

    Torsten Wappler

    2015-04-01

    Full Text Available The Paleozoic–Mesozoic transition is characterized by the most massive extinction of the Phanerozoic. Nevertheless, an impressive adaptive radiation of herbivorous insects occurred on gymnosperm-dominated floras not earlier than during the Middle to Late Triassic, penecontemporaneous with similar events worldwide, all which exhibit parallel expansions of generalized and mostly specialized insect herbivory on plants, expressed as insect damage on a various plant organs and tissues. The flora from Monte Agnello is distinctive, due to its preservation in subaerially deposited pyroclastic layers with exceptionally preserved details. Thus, the para-autochthonous assemblage provides insights into environmental disturbances, caused by volcanic activity, and how they profoundly affected the structure and composition of herbivory patterns. These diverse Middle Triassic biota supply extensive evidence for insect herbivore colonization, resulting in specific and complex herbivory patterns involving the frequency and diversity of 20 distinctive damage types (DTs. These DT patterns show that external foliage feeders, piercer-and-suckers, leaf miners, gallers, and oviposition culprits were intricately using almost all tissue types from the dominant host plants of voltzialean conifers (e.g., Voltzia, horsetails, ferns (e.g., Neuropteridium, Phlebopteris, Cladophlebis and Thaumatopteris, seed ferns (e.g., Scytophyllum, and cycadophytes (e.g., Bjuvia and Nilssonia.

  20. 40Ar/39Ar dates in the Central Cordillera of Colombia: Evidence for an upper triassic regional tecnomagmatic event

    International Nuclear Information System (INIS)

    Vinasco, C.J.; Cordani, U.G.; Vasconcelos, P

    2001-01-01

    The Coastal, Western, Central and the Eastern Cordilleras constitute four independent mountain ranges in the Northern Andes. The Coastal and Western Cordillera comprises allochthonous oceanic sequences of basic volcanic rocks and marine sediments of Cenozoic and Upper Cretaceous age (Aspden et al., 1987). The so-called Central Cordillera Polymetamorphic Complex (Restrepo and Toussaint, 1982), made up the pre-Mesozoic Central Cordillera basement, consists of low through high-grade metamorphic rocks intruded by Meso-Cenozoic batholiths. The Eastern Cordillera is a polydeformed continental mountain range consisting of Precambrian and Paleozoic metamorphic and igneous rocks overlain by Paleozoic to Mesozoic sedimentary covers. The northern part of the Central Cordillera has been described as a suspect terrain affected by several tectonometamorphic episodes. The most important of them have been always considered those related to the Hercynian orogeny, in Devonian and Carboniferous times, and those occurring at the beginning of the Andean Cycle in the Cretaceous (Restrepo and Toussaint, 1982). A possible Lower Paleozoic and even Precambrian age for the basement of the terrain has also been proposed. Although some indications of a Permo-Triassic event are already available, this period was not considered as of great importance for the geologic evolution of the region. The available radiometric dates in the 200-250 Ma interval, for the Central Cordillera, are as follows: 1. One K-Ar whole rock age for a Granulite of 251±21 Ma. (Restrepo et al, 1991) of the El Retiro group SE of Medellin. However, a K-Ar biotite age of 110±10 Ma. (Toussaint et al., 1978) was obtained for a migmatite rock associated with the same group. 2. A Sm/Nd garnet and whole rock age of 226±17 Ma in rocks associated to granulites, SE of Medellin, interpreted as metamorphic ages (Correa and Martins, 2001). 3. K-Ar biotite dates for the Puqui metamorphic complex, occurring north of Medellin, ranging

  1. Rates of dinosaur body mass evolution indicate 170 million years of sustained ecological innovation on the avian stem lineage.

    Science.gov (United States)

    Benson, Roger B J; Campione, Nicolás E; Carrano, Matthew T; Mannion, Philip D; Sullivan, Corwin; Upchurch, Paul; Evans, David C

    2014-05-01

    Large-scale adaptive radiations might explain the runaway success of a minority of extant vertebrate clades. This hypothesis predicts, among other things, rapid rates of morphological evolution during the early history of major groups, as lineages invade disparate ecological niches. However, few studies of adaptive radiation have included deep time data, so the links between extant diversity and major extinct radiations are unclear. The intensively studied Mesozoic dinosaur record provides a model system for such investigation, representing an ecologically diverse group that dominated terrestrial ecosystems for 170 million years. Furthermore, with 10,000 species, extant dinosaurs (birds) are the most speciose living tetrapod clade. We assembled composite trees of 614-622 Mesozoic dinosaurs/birds, and a comprehensive body mass dataset using the scaling relationship of limb bone robustness. Maximum-likelihood modelling and the node height test reveal rapid evolutionary rates and a predominance of rapid shifts among size classes in early (Triassic) dinosaurs. This indicates an early burst niche-filling pattern and contrasts with previous studies that favoured gradualistic rates. Subsequently, rates declined in most lineages, which rarely exploited new ecological niches. However, feathered maniraptoran dinosaurs (including Mesozoic birds) sustained rapid evolution from at least the Middle Jurassic, suggesting that these taxa evaded the effects of niche saturation. This indicates that a long evolutionary history of continuing ecological innovation paved the way for a second great radiation of dinosaurs, in birds. We therefore demonstrate links between the predominantly extinct deep time adaptive radiation of non-avian dinosaurs and the phenomenal diversification of birds, via continuing rapid rates of evolution along the phylogenetic stem lineage. This raises the possibility that the uneven distribution of biodiversity results not just from large-scale extrapolation of

  2. Analysis of a Permo-Triassic polarity transition in different absolute reconstructions of Pangaea, considering a model with features of the present Earth magnetic field

    Directory of Open Access Journals (Sweden)

    M. A. Van Zele

    2007-06-01

    Full Text Available The main objective of this paper is to show that the distribution of transitional palaeomagnetic data recorded at 250 Ma are in agreement with simulated data that depend on the sampling site, using a model that considers features of the Present Earth magnetic field. The analysis was performed comparing simulated reversals with the Permo-Triassic polarity transition recorded in the Siberian Trap Basalts. The palaeomagnetic data were corrected according to the Palaeo-latitude and Palaeo-longitude of Siberia (absolute reconstruction at 250 Ma using hotspot tracks. To obtain the motion of Siberia relative to hotspots from the Present time back to 250 Ma, three different Pangaea models were considered (Pangaea A, Pangaea A2, Pangaea B. In spite of the uncertainties associated with the use of hotspot frameworks and Pangaea configurations, both the modelled and recorded data show a remarkable fit when absolute reconstructions of Pangaea A and A2 configurations are performed. The agreement between both simulated and recorded data suggests that similar features to that of the Present Earth magnetic field could have been involved in reversals since the Permo-Triassic.

  3. The hELENa project - I. Stellar populations of early-type galaxies linked with local environment and galaxy mass

    Science.gov (United States)

    Sybilska, A.; Lisker, T.; Kuntschner, H.; Vazdekis, A.; van de Ven, G.; Peletier, R.; Falcón-Barroso, J.; Vijayaraghavan, R.; Janz, J.

    2017-09-01

    We present the first in a series of papers in The role of Environment in shaping Low-mass Early-type Nearby galaxies (hELENa) project. In this paper, we combine our sample of 20 low-mass early types (dEs) with 258 massive early types (ETGs) from the ATLAS3D survey - all observed with the SAURON integral field unit - to investigate early-type galaxies' stellar population scaling relations and the dependence of the population properties on local environment, extended to the low-σ regime of dEs. The ages in our sample show more scatter at lower σ values, indicative of less massive galaxies being affected by the environment to a higher degree. The shape of the age-σ relations for cluster versus non-cluster galaxies suggests that cluster environment speeds up the placing of galaxies on the red sequence. While the scaling relations are tighter for cluster than for the field/group objects, we find no evidence for a difference in average population characteristics of the two samples. We investigate the properties of our sample in the Virgo cluster as a function of number density (rather than simple clustrocentric distance) and find that dE ages correlate with the local density such that galaxies in regions of lower density are younger, likely because they are later arrivals to the cluster or have experienced less pre-processing in groups, and consequently used up their gas reservoir more recently. Overall, dE properties correlate more strongly with density than those of massive ETGs, which was expected as less massive galaxies are more susceptible to external influences.

  4. A Mass Spectrometric Analysis Method Based on PPCA and SVM for Early Detection of Ovarian Cancer.

    Science.gov (United States)

    Wu, Jiang; Ji, Yanju; Zhao, Ling; Ji, Mengying; Ye, Zhuang; Li, Suyi

    2016-01-01

    Background. Surfaced-enhanced laser desorption-ionization-time of flight mass spectrometry (SELDI-TOF-MS) technology plays an important role in the early diagnosis of ovarian cancer. However, the raw MS data is highly dimensional and redundant. Therefore, it is necessary to study rapid and accurate detection methods from the massive MS data. Methods. The clinical data set used in the experiments for early cancer detection consisted of 216 SELDI-TOF-MS samples. An MS analysis method based on probabilistic principal components analysis (PPCA) and support vector machine (SVM) was proposed and applied to the ovarian cancer early classification in the data set. Additionally, by the same data set, we also established a traditional PCA-SVM model. Finally we compared the two models in detection accuracy, specificity, and sensitivity. Results. Using independent training and testing experiments 10 times to evaluate the ovarian cancer detection models, the average prediction accuracy, sensitivity, and specificity of the PCA-SVM model were 83.34%, 82.70%, and 83.88%, respectively. In contrast, those of the PPCA-SVM model were 90.80%, 92.98%, and 88.97%, respectively. Conclusions. The PPCA-SVM model had better detection performance. And the model combined with the SELDI-TOF-MS technology had a prospect in early clinical detection and diagnosis of ovarian cancer.

  5. Recognition of the Kaweka Terrane in northern South Island, New Zealand : preliminary evidence from Rb-Sr metamorphic and U-Pb detrital zircon ages

    International Nuclear Information System (INIS)

    Adams, C.J.; Mortimer, N.; Campbell, H.J.; Griffin, W.L.

    2011-01-01

    Detrital zircon U-Pb ages and Rb-Sr metamorphic ages from low-grade Torlesse Supergroup metasedimentary rocks from North Canterbury and Marlborough provide preliminary evidence for a continuation of the Kaweka Terrane of the central North Island into the Torlesse Composite Terrane of the South Island. This would extend from the south side of the Wairau Fault in the upper Wairau River valley southwards to the Lake Tennyson and Lake Sumner areas, and as far as Hawarden. Rb-Sr ages indicate Jurassic metamorphism, 170±24 Ma, with initial 87 Sr/ 86 Sr ratios at that time 0.7073 ± 0.0007 i.e. similar to their North Island counterparts. These Kaweka Terrane rocks have detrital zircon ages that follow the distinctive pattern of the Torlesse rocks in general, i.e. substantial (>30%) Permian-Triassic and Precambrian-Early Palaeozoic groupings, but they also have minor youngest age components c. 175-165 Ma which constrain a maximum Early-Middle Jurassic depositional age. In detail, significant, older zircon components are Late Permian-Early Triassic (260-240 Ma) and Late Triassic (220-210 Ma), but the Precambrian-Early Palaeozoic detrital zircon ages are more scattered. The Kaweka Terrane thus forms a linear belt (c. 500 km) along the length of the Torlesse Composite Terrane in both North and South Islands. In the South Island it coincides with the Esk Head Belt, i.e. east of the Rakaia Terrane (Triassic-Late Permian) and west of the Pahau Terrane (Early Cretaceous-Late Jurassic). (author). 61 refs., 4 figs., 2 tabs.

  6. Suitability of Gypsum for the Production of Gypsum Plaster an Example from the Abu-Ruweis Evaporites (Upper Triassic), as Subayhi Area, Northwest Jordan

    International Nuclear Information System (INIS)

    Saqqa, W.; Arikat, M.

    2003-01-01

    The gypsum of Abu Ruweis Formation (Upper Triassic) of the northwestern Jordan was chemically, physically and mechanically assessed for likely industrial uses. Chemical results indicate that the investigated gypsum (calcium sulfate dihydrate) is highly pure. This is confirmed by the high ratios of CaO (32-33%) andSO 3 (41-43%) and the very low content of other elements such as Mg, Al and Si. Trace elements are also negligible. The CaSO 4 % (86-88%), purity ratio (CaO/SO 3 ) (0.85) , and combined water (≅5%) for β-hemihydrate gypsum agree well with the jordanian standards for gypsum plaster. Thermal analyses indicate that the temperature required to obtain hemihydrate gypsum falls between 175deg-200deg. A significant mass variation or maximum loss on weight (-20%) was achieved after initial dehydration by heating to 200deg. Heating to 225deg is far enough to evolve all combined water molecules. The results of fineness, consistency, setting time, compressive strength and flexural strength for β-hemilhydrate agree, in most of the case, with the local and international standards intended for gypsum building components. The study showed that the absorption and consistency ratios for final gypsum product are proportional to porosity, whilst the consistency itself decreases with the increase of applied stresses. (author)

  7. SDSS-IV MaNGA: Variation of the Stellar Initial Mass Function in Spiral and Early-type Galaxies

    Science.gov (United States)

    Li, Hongyu; Ge, Junqiang; Mao, Shude; Cappellari, Michele; Long, R. J.; Li, Ran; Emsellem, Eric; Dutton, Aaron A.; Li, Cheng; Bundy, Kevin; Thomas, Daniel; Drory, Niv; Lopes, Alexandre Roman

    2017-04-01

    We perform Jeans anisotropic modeling (JAM) on elliptical and spiral galaxies from the MaNGA DR13 sample. By comparing the stellar mass-to-light ratios estimated from stellar population synthesis and from JAM, we find a systematic variation of the initial mass function (IMF) similar to that in the earlier {{ATLAS}}3{{D}} results. Early-type galaxies (elliptical and lenticular) with lower velocity dispersions within one effective radius are consistent with a Chabrier-like IMF, while galaxies with higher velocity dispersions are consistent with a more bottom-heavy IMF such as the Salpeter IMF. Spiral galaxies have similar systematic IMF variations, but with slightly different slopes and larger scatters, due to the uncertainties caused by the higher gas fractions and extinctions for these galaxies. Furthermore, we examine the effects of stellar mass-to-light ratio gradients on our JAM modeling, and we find that the trends become stronger after considering the gradients.

  8. Sommerxylon spiralosus from Upper Triassic in southernmost Paraná Basin (Brazil: a new taxon with taxacean affinity

    Directory of Open Access Journals (Sweden)

    Etiene F. Pires

    2004-09-01

    Full Text Available The anatoical description of silici?ed Gymnospermae woods from Upper Triassic sequences of southernmost Paraná Basin (Brazil has allowed the identi?cation of a new taxon: Sommerxylon spiralosus n.gen. et n.sp. Diagnostic parameters, such as heterocellular medulla composed of parenchymatous and sclerenchymatous cells, primary xylem endarch, secondary xylem with dominant uniseriate bordered pits, spiral thickenings in the radial walls of tracheids, medullar rays homocellular, absence of resiniferous canals and axial parenchyma, indicate its relationship with the family Taxaceae, reporting on the first recognition of this group in the Triassic on Southern Pangea. This evidence supports the hypothesis that the Taxaceae at the Mesozoic were not con?ned to the Northern Hemisphere.A descrição anatômica de lenhos silicificados de Gymnospermae em seqüência do Triássico Superior no sul da Bacia do Paraná (Brasil, possibilitou a identificação de um novo taxon: Sommerxylon spiralosus n.gen. et n.sp. Parâmetros diagnósticos tais como medula heterocelular, composta por células parenquimáticas e esclerenquimáticas, xilema primário endarco, xilema secundário com pontoações areoladas unisseriadas dominantes, espessamentos espiralados nas paredes radiais dos traqueídeos, raios lenhosos homocelulares, ausência de canais resiníferos e de parênquima axial, indicam a sua vinculação à família Taxaceae, constituindo-se em reconhecimento inédito da presença deste grupo no Triássico Superior no sul do Pangea. Esta evidência suporta a hipótese de que a família Taxaceae não estava confinada ao Hemisfério Norte durante o Mesozóico.

  9. THE SL2S GALAXY-SCALE LENS SAMPLE. IV. THE DEPENDENCE OF THE TOTAL MASS DENSITY PROFILE OF EARLY-TYPE GALAXIES ON REDSHIFT, STELLAR MASS, AND SIZE

    Energy Technology Data Exchange (ETDEWEB)

    Sonnenfeld, Alessandro; Treu, Tommaso; Suyu, Sherry H. [Physics Department, University of California, Santa Barbara, CA 93106 (United States); Gavazzi, Raphaël [Institut d' Astrophysique de Paris, UMR7095 CNRS-Université Pierre et Marie Curie, 98bis bd Arago, F-75014 Paris (France); Marshall, Philip J. [Kavli Institute for Particle Astrophysics and Cosmology, Stanford University, 452 Lomita Mall, Stanford, CA 94305 (United States); Auger, Matthew W. [Institute of Astronomy, University of Cambridge, Madingley Road, Cambridge CB3 0HA (United Kingdom); Nipoti, Carlo, E-mail: sonnen@physics.ucsb.edu [Astronomy Department, University of Bologna, via Ranzani 1, I-40127 Bologna (Italy)

    2013-11-10

    We present optical and near-infrared spectroscopy obtained at Keck, Very Large Telescope, and Gemini for a sample of 36 secure strong gravitational lens systems and 17 candidates identified as part of the Strong Lensing Legacy Survey. The deflectors are massive early-type galaxies in the redshift range z{sub d} = 0.2-0.8, while the lensed sources are at z{sub s} = 1-3.5. We combine these data with photometric and lensing measurements presented in the companion paper III and with lenses from the Sloan Lens Advanced Camera for Surveys and Lènses Structure and Dynamics surveys to investigate the cosmic evolution of the internal structure of massive early-type galaxies over half the age of the universe. We study the dependence of the slope of the total mass density profile, γ' (ρ(r)∝r{sup -γ{sup '}}), on stellar mass, size, and redshift. We find that two parameters are sufficient to determine γ' with less than 6% residual scatter. At fixed redshift, γ' depends solely on the surface stellar mass density ∂γ'/∂Σ{sub *} = 0.38 ± 0.07, i.e., galaxies with denser stars also have steeper slopes. At fixed M{sub *} and R{sub eff}, γ' depends on redshift, in the sense that galaxies at a lower redshift have steeper slopes (∂γ'/∂z = –0.31 ± 0.10). However, the mean redshift evolution of γ' for an individual galaxy is consistent with zero dγ'/dz = –0.10 ± 0.12. This result is obtained by combining our measured dependencies of γ' on z, M{sub *},R{sub eff} with the evolution of the R{sub eff}-M{sub *} taken from the literature, and is broadly consistent with current models of the formation and evolution of massive early-type galaxies. Detailed quantitative comparisons of our results with theory will provide qualitatively new information on the detailed physical processes at work.

  10. THE SL2S GALAXY-SCALE LENS SAMPLE. IV. THE DEPENDENCE OF THE TOTAL MASS DENSITY PROFILE OF EARLY-TYPE GALAXIES ON REDSHIFT, STELLAR MASS, AND SIZE

    International Nuclear Information System (INIS)

    Sonnenfeld, Alessandro; Treu, Tommaso; Suyu, Sherry H.; Gavazzi, Raphaël; Marshall, Philip J.; Auger, Matthew W.; Nipoti, Carlo

    2013-01-01

    We present optical and near-infrared spectroscopy obtained at Keck, Very Large Telescope, and Gemini for a sample of 36 secure strong gravitational lens systems and 17 candidates identified as part of the Strong Lensing Legacy Survey. The deflectors are massive early-type galaxies in the redshift range z d = 0.2-0.8, while the lensed sources are at z s = 1-3.5. We combine these data with photometric and lensing measurements presented in the companion paper III and with lenses from the Sloan Lens Advanced Camera for Surveys and Lènses Structure and Dynamics surveys to investigate the cosmic evolution of the internal structure of massive early-type galaxies over half the age of the universe. We study the dependence of the slope of the total mass density profile, γ' (ρ(r)∝r -γ ' ), on stellar mass, size, and redshift. We find that two parameters are sufficient to determine γ' with less than 6% residual scatter. At fixed redshift, γ' depends solely on the surface stellar mass density ∂γ'/∂Σ * = 0.38 ± 0.07, i.e., galaxies with denser stars also have steeper slopes. At fixed M * and R eff , γ' depends on redshift, in the sense that galaxies at a lower redshift have steeper slopes (∂γ'/∂z = –0.31 ± 0.10). However, the mean redshift evolution of γ' for an individual galaxy is consistent with zero dγ'/dz = –0.10 ± 0.12. This result is obtained by combining our measured dependencies of γ' on z, M * ,R eff with the evolution of the R eff -M * taken from the literature, and is broadly consistent with current models of the formation and evolution of massive early-type galaxies. Detailed quantitative comparisons of our results with theory will provide qualitatively new information on the detailed physical processes at work

  11. Astronomical cycle origin of bedded chert: A middle Triassic bedded chert sequence, Inuyama, Japan

    Science.gov (United States)

    Ikeda, Masayuki; Tada, Ryuji; Sakuma, Hironobu

    2010-09-01

    Astronomical forcing is one of the main drivers of climate change, and astronomical cyclicity recorded in sediments provides a clue to understand the dynamics of the global climate system. Bedded cherts consist of rhythmic alternations of chert and shale beds. Although previous studies have hypothesized that the origin of bedded chert is related to astronomical cycles (e.g. Fischer, 1976; Hori et al., 1993), conclusive proof remains elusive. To explore this possibility, we established a continuous, high-resolution lithostratigraphy of middle Triassic bedded chert in Central Japan. The average duration of each chert-shale couplet is 20 kyr, similar to that of the precession cycle. Spectral analysis of a bed number series of thickness variations in chert beds was performed assuming that each chert-shale couplet represents a 20-kyr precession cycle. The results reveal cycles involving approximately 200, 20, 5, and 2-3 beds, corresponding to periodicities of approximately 4000, 400, 100, and 40-60 kyr, respectively. By further assuming that the 20-bed cycle represents a 405-kyr eccentricity cycle of constant and stable periodicity, we converted the bed number series to a time series. Spectral analysis of the time series revealed distinct periodicities of 3600, 117, 97, and 38 kyr, in addition to 405 kyr. Besides 3600 kyr, these periodicities agree well with the 120, 95, and 37 kyr periodicities for eccentricity cycles and the obliquity cycle during the Triassic. Moreover, we detected amplitude modulation of the approximately 100-kyr cycle of thickness variations in chert beds with a 405-kyr periodicity, which may correspond to amplitude modulation of 100-kyr climatic precession cycle with the 405-kyr periodicity. The approximately 3600-kyr periodicity described above and 1800-kyr periodicity manifested as the amplitude modulation of the 405-kyr cycle are correlated to present-day long-term eccentricity cycles of 2400 and 4800 kyr evolved by chaotic behavior of solar

  12. The Middle Triassic evolution of the Bangong-Nujiang Tethyan Ocean: evidence from analyses of OIB-type basalts and OIB-derived phonolites in northern Tibet

    Science.gov (United States)

    Fan, Jian-Jun; Li, Cai; Liu, Jin-Heng; Wang, Ming; Liu, Yi-Ming; Xie, Chao-Ming

    2017-12-01

    In this paper, we present new major and trace element chemical data for the basalts and phonolites of the Nare ocean island fragment (NaOI), as well as zircon U-Pb age data and Hf isotope compositions for the NaOI phonolites in the middle segment of the Bangong-Nujiang Suture Zone, northern Tibet. Our aim is to assess the genesis of these rocks and to reconstruct the Middle Triassic evolution of the Bangong-Nujiang Tethyan Ocean (BNTO). The NaOI retains an ocean island-type double-layered structure comprising a basaltic basement and an oceanic sedimentary cover sequence (conglomerate and limestone, the latter accompanied by layers of erupted phonolite near the top of the sequence). The basalts in the NaOI are enriched in light rare earth elements and high field strength elements (Nb, Ta, Zr, Hf, and Ti), and they exhibit chondrite-normalized REE patterns and primitive mantle-normalized trace element patterns similar to those of ocean island basalts. Taking into consideration their high Dy/Yb, Sm/Yb, and La/Sm ratios, we conclude that the NaOI basalts were derived from the partial melting of garnet peridotite in the mantle. The NaOI phonolites have LREE-enriched chondrite-normalized REE patterns with negative Eu anomalies (Eu/Eu* = 0.41-0.43) and primitive mantle-normalized trace element patterns with enrichments in Nb, Ta, Zr, and Hf, and depletions in Ba, U, Sr, P, and Ti. Given the high contents of Nb (172-256 ppm), Ta (11.8-16.0 ppm), Zr (927-1117 ppm), and Hf (20.8-26.9 ppm), and the very low contents of MgO (0.11-0.25 wt%), the very low Mg# values (5-10), and the near-zero contents of Cr (1.27-7.59 ppm), Ni (0.43-7.19 ppm), and Co (0.11-0.38 ppm), and the small and homogeneously positive ɛ Hf(t) values (+ 4.9 to + 9.5), we infer that the NaOI phonolites were formed by the fractional crystallization of an OIB-derived mafic parent magma. The phonolites of the NaOI contain zircons that yielded U-Pb ages of 239 and 242 Ma, indicating that the NaOI formed during

  13. On origin of radium aureoles around Triassic uranium mineralization zones in the Peribaltic Syneclize

    International Nuclear Information System (INIS)

    Szewczyk, J.

    1985-01-01

    In the second half of the seventies, the Geological Institute began search for sandstone uranium deposits in the Triassic of the Peribaltic Syneclize. Detailed analysis of both laboratory and geophysical data showed presence of radium (Ra-226) aureoles around uranium ore bodies hitherto found by drillings. The mechanism of origin of the aureoles is explained and methodological proposal of their use in further search for uranium deposits is given. Theoretical modelling showed that origin of the aureoles is mainly related to movement of deposit waters percolating through uranium ore body. The influence of shape and dimensions of radium aureole-generating ore bodies on extent of the aureoles appears subordinate. Aureoles interesting from the point of view of prospecting may originate when velocity of percolating waters falls within the range from 10 -8 to 10 -6 cm/s. 23 refs., 7 figs., 1 tab. (author)

  14. New results of microfaunal and geochemical investigations in the Permian-Triassic boundary interval from the Jadar Block (NW Serbia)

    Science.gov (United States)

    Sudar, Milan N.; Kolar-Jurkovšek, Tea; Nestell, Galina P.; Jovanović, Divna; Jurkovšek, Bogdan; Williams, Jeremy; Brookfield, Michael; Stebbins, Alan

    2018-04-01

    Detail results of microfaunal, sedimentological and geochemical investigations are documented from a newly discovered section of the Permian-Triassic boundary (PTB) interval in the area of the town of Valjevo (northwestern Serbia). The presence of various and abundant microfossils (conodonts, foraminifers, and ostracodes) found in the Upper Permian "Bituminous limestone" Formation enabled a determination of the Changhsingian Hindeodus praeparvus conodont Zone. This paper is the first report of latest Permian strata from the region, as well as from all of Serbia, where the PTB interval sediments have been part of a complex/integrated study by means of biostratigraphy and geochemistry.

  15. Body dissatisfaction and body mass in girls and boys transitioning from early to mid-adolescence: additional role of self-esteem and eating habits

    Directory of Open Access Journals (Sweden)

    Mäkinen Mauno

    2012-06-01

    Full Text Available Abstract Background In the transition from early to mid-adolescence, gender differences in pubertal development become significant. Body dissatisfaction is often associated with body mass, low self-esteem and abnormal eating habits. The majority of studies investigating body dissatisfaction and its associations have been conducted on female populations. However, some evidence suggests that males also suffer from these problems and that gender differences might already be observed in adolescence. Aims To examine body dissatisfaction and its relationship with body mass, as well as self-esteem and eating habits, in girls and boys in transition from early to mid-adolescence. Methods School nurses recorded the heights and weights of 659 girls and 711 boys with a mean age of 14.5 years. The Rosenberg Self-Esteem Scale and the Body Dissatisfaction subscale of the Eating Disorder Inventory were used as self-appraisal scales. Eating data were self-reported. Results The girls were less satisfied with their bodies than boys were with theirs (mean score (SD: 30.6 (SD 12.2 vs. 18.9 (SD 9.5; p  Conclusions Body mass, self-esteem and eating habits revealed a significant relationship with body dissatisfaction in the transitional phase from early to mid-adolescence in girls and boys, but significant gender differences were also found.

  16. Global nickel anomaly links Siberian Traps eruptions and the latest Permian mass extinction.

    Science.gov (United States)

    Rampino, Michael R; Rodriguez, Sedelia; Baransky, Eva; Cai, Yue

    2017-09-29

    Anomalous peaks of nickel abundance have been reported in Permian-Triassic boundary sections in China, Israel, Eastern Europe, Spitzbergen, and the Austrian Carnic Alps. New solution ICP-MS results of enhanced nickel from P-T boundary sections in Hungary, Japan, and Spiti, India suggest that the nickel anomalies at the end of the Permian were a worldwide phenomenon. We propose that the source of the nickel anomalies at the P-T boundary were Ni-rich volatiles released by the Siberian volcanism, and by coeval Ni-rich magma intrusions. The peaks in nickel abundance correlate with negative δ 13 C and δ 18 O anomalies, suggesting that explosive reactions between magma and coal during the Siberian flood-basalt eruptions released large amounts of CO 2 and CH 4 into the atmosphere, causing severe global warming and subsequent mass extinction. The nickel anomalies may provide a timeline in P-T boundary sections, and the timing of the peaks supports the Siberian Traps as a contributor to the latest Permian mass extinction.

  17. Permian-Triassic palynofacies and chemostratigraphy in a core recovered from central Spitsbergen.

    Science.gov (United States)

    van Soelen, Els; Hasic, Edi; Planke, Sverre; Svensen, Henrik; Sleveland, Arve; Midtkandal, Ivar; Twitchett, Richard; Kürschner, Wolfram

    2017-04-01

    The Late Permian biotic crisis is one of the most severe extinction events in the history of the Earth, affecting both terrestrial and marine environments. A large igneous province (LIP) in Siberia is thought to be linked with this global event; however, correlation between the volcanic event and the biotic crisis is difficult and requires well dated and high resolution Permian-Triassic boundary successions from the Arctic region. The Svalbard end-Permian drilling project is aimed at improved correlation of the Permian-Triassic sections in Svalbard with the Siberian Traps. The core was collected from Deltadalen, in central Spitsbergen, with additional samples collected from an outcrop close to the drilling site. As part of this collaborative project, carbon isotope geochemistry, palynofacies and palynomorphs were studied in order to learn more about the biostratigraphy and to understand changes in the source(s) of organic matter. Objectives were to reconstruct the paleo-environment; to correlate the core with other sites on Svalbard, and with global records; and to identify and characterize the Late Permian extinction event in the core. A carbon isotope shift is an important global stratigraphic marker in the latest Permian and occurs near the base of the Vikinghøgda Formation in the Deltadalen core, where bulk rock values change from -24.5 to -32.7‰. Palynomorph preservation was generally poor in both core and outcrop samples which prevented detailed examination of species and limited their usefulness for biostratigraphy. Still, palynofacies were useful for correlative purposes. AOM (amorphous organic matter) in the core increases at the lithological change from sandstones to siltstones, and is indicative of anoxic conditions. Similar high levels of AOM in the outcrop samples can be correlated with the core. Palynological analyses show that the spore/pollen ratio starts to increase before the negative shift in the isotope curve. Such an increase in spore

  18. Some contrasting biostratigraphic links between the Baker and Olds Ferry Terranes, eastern Oregon

    Science.gov (United States)

    Nestell, Merlynd K.; Blome, Charles D.

    2016-01-01

    New stratigraphic and paleontologic data indicate that ophiolitic melange windows in the Olds Ferry terrane of eastern Oregon contain limestone blocks and chert that are somewhat different in age than those present in the adjacent Baker terrane melange. The melange windows in the Olds Ferry terrane occur as inliers in the flyschoid Early and Middle Jurassic age Weatherby Formation, which depositionally overlies the contact between the melange-rich Devonian to Upper Triassic rocks of the Baker terrane on the north, and Upper Triassic and Early Jurassic volcanic arc rocks of the Huntington Formation on the south. The Baker terrane and Huntington Formation represent fragments of a subduction complex and related volcanic island arc, whereas the Weatherby Formation consists of forearc basin sedimentary deposits. The tectonic blocks in the melange windows of the Weatherby Formation (in the Olds Ferry terrane) are dated by scarce biostratigraphic evidence as Upper Pennsylvanian to Lower Permian and Upper Triassic. In contrast, tectonic blocks of limestone in theBaker terrane yield mostly fusulinids and small foraminifers of Middle Pennsylvanian Moscovian age at one locality.Middle Permian (Guadalupian) Tethyan fusulinids and smaller foraminifers (neoschwagerinids and other Middle Permian genera) are present at a few other localities. Late Triassic conodonts and bryozoans are also present in a few of the Baker terrane tectonic blocks. These limestone blocks are generally embedded in Permian and Triassic radiolarian bearing chert or argillite. Based on conodont, radiolarian and fusulinid data, the age limits of the meange blocks in the Weatherby Formation range from Pennsylvanian to Late Triassic.

  19. BIRD-LIKE HEAD ON A CHAMELEON BODY: NEW SPECIMENS OF THE ENIGMATIC DIAPSID REPTILE MEGALANCOSAURUS FROM THE LATE TRIASSIC OF NORTHERN ITALY

    Directory of Open Access Journals (Sweden)

    SILVIO RENESTO

    2000-07-01

    Full Text Available Two new Megalancosaurus specimens collected from the Norian (Late Triassic Calcare di Zorzino (Zorzino Limestone Formation (Bergamo, Lombardy, Northern Italy, and from the coeval Dolomia di Forni (Forni Dolostone Formation (Udine, Friuli, North-eastern Italy improve our knowledge of skeletal anatomy and mode of life of this genus. Morphology of observable skull elements and cervical vertebrae in one of the new specimens shows some resemblance to the possible Triassic bird Protoavis, while the postcranial skeleton of Megalancosaurus is completely non-avian. This may suggest that either Megalancosaurus and Protoavis developed a similar neck structure as a response to the same functional requirement, or that part of the disarticulated material ascribed to Protoavis may indeed belong to a Megalancosaurus-like reptile. Megalancosaurus shows a very high adaptation to arboreal life and a peculiar feeding strategy. Recent suggestions that Megalancosaurus may have been a glider and a possible model for bird ancestry are discussed. Some skeletal features of Megalancosaurus may indeed be interpreted as gliding adaptation, but evidence is weak, and if this reptile was a glider, however, its gliding bauplan should have been completely different from the one usually accepted for bird ancestors, showing instead more morphological similarities with gliding squirrels, phalangeriids and putative ancestors of bats and pterosaurs (according to a non cursorial model for these latter. 

  20. A new saurichthyiform (Actinopterygii with a crushing feeding mechanism from the Middle Triassic of Guizhou (China.

    Directory of Open Access Journals (Sweden)

    Feixiang Wu

    Full Text Available Equipped with an effective predatory feeding mechanism enhanced by large and sharp teeth, pointed snout and elongate body, saurichthyiform fishes are considered common fish-eaters in the early Mesozoic aquatic ecosystems. Additionally, because of the similar body plan across species, saurichthyiforms are also regarded evolutionally conservative, with few morphological and ecological changes during their long history. However, their phylogenetic affinity remains unclear as to whether they are chondrostean, neopterygian or stem-actinopteran, and likewise the intrarelationships of the group have rarely been explored.Here we report a new saurichthyiform from the Middle Triassic of Guizhou, China, based on the well-preserved specimens including a 3-D braincase. The new taxon, Yelangichthys macrocephalus gen. et sp. nov., is unique among saurichthyiforms in having a peculiar neurocranium with a broad orbital tectum, paired posterior myodomes, a deep, transverse fossa in the posterodorsal part of the orbit, and a feeding mechanism structured for durophagy. Phylogenetic analysis places Yelangichthys gen. nov. at the most basal position in the Saurichthyiformes as the sister to Saurichthyidae, and a new family Yelangichthyidae is erected to include only Y. macrocephalus gen. et sp. nov. The monophyly of the Chondrostei comprising [Saurichthyiformes + Acipenseriformes] Birgeriiformes is supported, but not the monophyly of Saurichthys, the type genus of Saurichthyidae. With its outstanding osteological details, Yelangichthys gen. nov. greatly increases the neurocranial variations in saurichthyiforms, and its novel feeding structure suggests the consumption of hard-preys instead of fishes.Our findings highlight the detailed osteology of a saurichthyiform braincase and its feeding design. We suggest that saurichthyiforms are closely allied to the Acipenseriformes. Saurichthyiforms were very diverse in the cranial osteology and they might have undergone a

  1. A New Saurichthyiform (Actinopterygii) with a Crushing Feeding Mechanism from the Middle Triassic of Guizhou (China)

    Science.gov (United States)

    Wu, Feixiang; Chang, Mee-mann; Sun, Yuanlin; Xu, Guanghui

    2013-01-01

    Background Equipped with an effective predatory feeding mechanism enhanced by large and sharp teeth, pointed snout and elongate body, saurichthyiform fishes are considered common fish-eaters in the early Mesozoic aquatic ecosystems. Additionally, because of the similar body plan across species, saurichthyiforms are also regarded evolutionally conservative, with few morphological and ecological changes during their long history. However, their phylogenetic affinity remains unclear as to whether they are chondrostean, neopterygian or stem-actinopteran, and likewise the intrarelationships of the group have rarely been explored. Methodology/Principal Findings Here we report a new saurichthyiform from the Middle Triassic of Guizhou, China, based on the well-preserved specimens including a 3-D braincase. The new taxon, Yelangichthys macrocephalus gen. et sp. nov., is unique among saurichthyiforms in having a peculiar neurocranium with a broad orbital tectum, paired posterior myodomes, a deep, transverse fossa in the posterodorsal part of the orbit, and a feeding mechanism structured for durophagy. Phylogenetic analysis places Yelangichthys gen. nov. at the most basal position in the Saurichthyiformes as the sister to Saurichthyidae, and a new family Yelangichthyidae is erected to include only Y. macrocephalus gen. et sp. nov. The monophyly of the Chondrostei comprising [Saurichthyiformes + Acipenseriformes] Birgeriiformes is supported, but not the monophyly of Saurichthys, the type genus of Saurichthyidae. With its outstanding osteological details, Yelangichthys gen. nov. greatly increases the neurocranial variations in saurichthyiforms, and its novel feeding structure suggests the consumption of hard-preys instead of fishes. Conclusions/Significance Our findings highlight the detailed osteology of a saurichthyiform braincase and its feeding design. We suggest that saurichthyiforms are closely allied to the Acipenseriformes. Saurichthyiforms were very diverse in the

  2. The evolution of the manus of early theropod dinosaurs is characterized by high inter- and intraspecific variation.

    Science.gov (United States)

    Barta, Daniel E; Nesbitt, Sterling J; Norell, Mark A

    2018-01-01

    The origin of the avian hand, with its reduced and fused carpals and digits, from the five-fingered hands and complex wrists of early dinosaurs represents one of the major transformations of manus morphology among tetrapods. Much attention has been directed to the later part of this transition, from four- to three-fingered taxa. However, earlier anatomical changes may have influenced these later modifications, possibly paving the way for a later frameshift in digit identities. We investigate the five- to four-fingered transition among early dinosaurs, along with changes in carpus morphology. New three-dimensional reconstructions from computed tomography data of the manus of the Triassic and Early Jurassic theropod dinosaurs Coelophysis bauri and Megapnosaurus rhodesiensis are described and compared intra- and interspecifically. Several novel findings emerge from these reconstructions and comparisons, including the first evidence of an ossified centrale and a free intermedium in some C. bauri specimens, as well as confirmation of the presence of a vestigial fifth metacarpal in this taxon. Additionally, a specimen of C. bauri and an unnamed coelophysoid from the Upper Triassic Hayden Quarry, New Mexico, are to our knowledge the only theropods (other than alvarezsaurs and birds) in which all of the distal carpals are completely fused together into a single unit. Several differences between the manus of C. bauri and M. rhodesiensis are also identified. We review the evolution of the archosauromorph manus more broadly in light of these new data, and caution against incorporating carpal characters in phylogenetic analyses of fine-scale relationships of Archosauromorpha, in light of the high degree of observed polymorphism in taxa for which large sample sizes are available, such as the theropod Coelophysis and the sauropodomorph Plateosaurus. We also find that the reduction of the carpus and ultimate loss of the fourth and fifth digits among early dinosaurs did not

  3. K-Ar Geochronology and isotopic composition of the late oligocene- early miocene Ancud volcanic complex, Chiloe

    International Nuclear Information System (INIS)

    Munoz B, Jorge; Duhart O, Paul; Farmer, G. Lang; Stern, Charles R

    2001-01-01

    The Ancud Volcanic Complex (Gally and Sanchez , 1960) forms a portion of the Mid-Tertiary Coastal Magmatic Belt which outcrops in the area of northern Chiloe island. Main exposures occur at Ancud, Punta Polocue, Punihuil, Pumillahue, Tetas de Teguaco and Bahia Cocotue. The Ancud Volcanic Complex consists of basaltic to basaltic andesites lava flows and volcanic necks and rhyolitic pyroclastic flows and vitric domes. Previous studies indicate a Late Oligocene-Early Miocene age (Garcia et al., 1988; Stern and Vergara, 1992; Munoz et al., 2000). The Ancud Volcanic Complex covers and intrudes Palaeozoic-Triassic metamorphic rocks and is partially covered by an early to middle Miocene marine sedimentary sequence known as Lacui Formation (Valenzuela, 1982) and by Pleistocene glacial deposits (Heusser, 1990). At Punihuil locality, lava flows are interbedded with the lower part of the marine sedimentary sequence, which includes significant amounts of redeposited pyroclastic components. Locally, the presence of hyaloclastic breccias suggests interaction of magma with marine water (au)

  4. Dietary energy density affects fat mass in early adolescence and is not modified by FTO variants.

    Directory of Open Access Journals (Sweden)

    Laura Johnson

    Full Text Available Dietary energy density (DED does not have a simple linear relationship to fat mass in children, which suggests that some children are more susceptible than others to the effects of DED. Children with the FTO (rs9939609 variant that increases the risk of obesity may have a higher susceptibility to the effects of DED because their internal appetite control system is compromised. We tested the relationship between DED and fat mass in early adolescence and its interaction with FTO variants.We carried out a prospective analysis on 2,275 children enrolled in the Avon Longitudinal Study of Parents and Children (ALSPAC. Diet was assessed at age 10 y using 3-day diet diaries. DED (kJ/g was calculated excluding drinks. Children were genotyped for the FTO (rs9939609 variant. Fat mass was estimated at age 13 y using the Lunar Prodigy Dual-energy X-ray Absorptiometry scanner. There was no evidence of interaction between DED at age 10 y and the high risk A allele of the FTO gene in relation to fat mass at age 13 y (beta = 0.005, p = 0.51, suggesting that the FTO gene has no effect on the relation between DED at 10 y and fat mass at 13 y. When DED at 10 y and the A allele of FTO were in the same model they were independently related to fat mass at 13 y. Each A allele of FTO was associated with 0.35+/-0.13 kg more fat mass at 13 y and each 1 kJ/g DED at 10 y was associated with 0.16+/-0.06 kg more fat mass at age 13 y, after controlling for misreporting of energy intake, gender, puberty, overweight status at 10 y, maternal education, TV watching, and physical activity.This study reveals the multi-factorial origin of obesity and indicates that although FTO may put some children at greater risk of obesity, encouraging a low dietary energy density may be an effective strategy to help all children avoid excessive fat gain.

  5. Carbonate Minerals with Magnesium in Triassic Terebratula Limestone in the Term of Limestone with Magnesium Application as a Sorbent in Desulfurization of Flue Gases

    Science.gov (United States)

    Stanienda-Pilecki, Katarzyna

    2017-09-01

    This article presents the results of studies of Triassic (Muschelkalk) carbonate rock samples of the Terebratula Beds taken from the area of the Polish part of the Germanic Basin. It is the area of Opole Silesia. The rocks were studied in the term of possibility of limestone with magnesium application in desulfurization of flue gases executed in power plants. Characteristic features of especially carbonate phases including magnesium-low-Mg calcite, high-Mg calcite, dolomite and huntite were presented in the article. They were studied to show that the presence of carbonate phases with magnesium, especially high-Mg calcite makes the desulfurization process more effective. Selected rock samples were examined using a microscope with polarized, transmitted light, X-ray diffraction, microprobe measurements and FTIR spectroscopy. The results of studies show a domination of low magnesium calcite in the limestones of the Terebratula Beds. In some samples dolomite and lower amounts of high-Mg calcite occurred. Moreover, huntite was identified. The studies were very important, because carbonate phases like high-Mg calcite and huntite which occurred in rocks of the Triassic Terebratula Beds were not investigated in details by other scientists but they presence in limestone sorbent could influence the effectiveness of desulfurization process.

  6. A novel compact mass detection platform for the open access (OA) environment in drug discovery and early development.

    Science.gov (United States)

    Gao, Junling; Ceglia, Scott S; Jones, Michael D; Simeone, Jennifer; Antwerp, John Van; Zhang, Li-Kang; Ross, Charles W; Helmy, Roy

    2016-04-15

    A new 'compact mass detector' co-developed with an instrument manufacturer (Waters Corporation) as an interface for liquid chromatography (LC), specifically Ultra-high performance LC(®) (UPLC(®) or UHPLC) analysis was evaluated as a potential new Open Access (OA) LC-MS platform in the Drug Discovery and Early Development space. This new compact mass detector based platform was envisioned to provide increased reliability and speed while exhibiting significant cost, noise, and footprint reductions. The new detector was evaluated in batch mode (typically 1-3 samples per run) to monitor reactions and check purity, as well as in High Throughput Screening (HTS) mode to run 24, 48, and 96 well plates. The latter workflows focused on screening catalysis conditions, process optimization, and library work. The objective of this investigation was to assess the performance, reliability, and flexibility of the compact mass detector in the OA setting for a variety of applications. The compact mass detector results were compared to those obtained by current OA LC-MS systems, and the capabilities and benefits of the compact mass detector in the open access setting for chemists in the drug discovery and development space are demonstrated. Copyright © 2016 Elsevier B.V. All rights reserved.

  7. New absolute paleointensity determinations for the Permian-Triassic boundary from the Kuznetsk Trap Basalts.

    Science.gov (United States)

    Kulakov, E.; Metelkin, D. V.; Kazansky, A.

    2015-12-01

    We report the results of a pilot absolute paleointensity study of the ~250 Ma basalts of Kuznetsk traps (Kuznetsk Basin, Altai-Sayan folded area). Studied samples are characterized by a reversed polarity of natural remanent magnetization that corresponds to the lower part of Siberian Trap basalts sequence. Geochemical similarity of Kuznets basalts with those from Norilsk region supports this interpretation. Primary origin of thermal remanence in our sample is confirmed by a positive backed contact test. Rock magnetic analyses indicate that the ChRM is carried by single-domain titanomagnetite. The Coe-version of the Thellier-Therllier double-heating method was utilized for the paleointensity determinations. In contrast to the previous studies of the Permian-Triassic Siberian trap basalts, our data indicate that by the P-T boundary the paleofield intensity was relatively high and comparable with geomagnetic field strength for the last 10 millions of years. New results question the duration of the "Mesozoic dipole-low".

  8. Mass Society

    DEFF Research Database (Denmark)

    Borch, Christian

    2017-01-01

    the negative features usually ascribed by late nineteenth-century crowd psychology to spontaneous crowds, and attributes these to the entire social fabric. However, in contrast to crowd psychology, theorists of mass society often place greater emphasis on how capitalism, technological advances, or demographic......Mass society is a societal diagnosis that emphasizes – usually in a pejorative, modernity critical manner – a series of traits allegedly associated with modern society, such as the leveling of individuality, moral decay, alienation, and isolation. As such, the notion of mass society generalizes...... developments condition such negative features, and some theorists argue that mass society produces a propensity to totalitarianism. Discussions of mass society culminated in the early and mid-twentieth century....

  9. A Unique Late Triassic Dinosauromorph Assemblage Reveals Dinosaur Ancestral Anatomy and Diet.

    Science.gov (United States)

    Cabreira, Sergio Furtado; Kellner, Alexander Wilhelm Armin; Dias-da-Silva, Sérgio; Roberto da Silva, Lúcio; Bronzati, Mario; Marsola, Júlio Cesar de Almeida; Müller, Rodrigo Temp; Bittencourt, Jonathas de Souza; Batista, Brunna Jul'Armando; Raugust, Tiago; Carrilho, Rodrigo; Brodt, André; Langer, Max Cardoso

    2016-11-21

    Dinosauromorpha includes dinosaurs and other much less diverse dinosaur precursors of Triassic age, such as lagerpetids [1]. Joint occurrences of these taxa with dinosaurs are rare but more common during the latest part of that period (Norian-Rhaetian, 228-201 million years ago [mya]) [2, 3]. In contrast, the new lagerpetid and saurischian dinosaur described here were unearthed from one of the oldest rock units with dinosaur fossils worldwide, the Carnian (237-228 mya) Santa Maria Formation of south Brazil [4], a record only matched in age by much more fragmentary remains from Argentina [5]. This is the first time nearly complete dinosaur and non-dinosaur dinosauromorph remains are found together in the same excavation, clearly showing that these animals were contemporaries since the first stages of dinosaur evolution. The new lagerpetid preserves the first skull, scapular and forelimb elements, plus associated vertebrae, known for the group, revealing how dinosaurs acquired several of their typical anatomical traits. Furthermore, a novel phylogenetic analysis shows the new dinosaur as the most basal Sauropodomorpha. Its plesiomorphic teeth, strictly adapted to faunivory, provide crucial data to infer the feeding behavior of the first dinosaurs. Copyright © 2016 Elsevier Ltd. All rights reserved.

  10. Tetrapod tracks in Permo–Triassic eolian beds of southern Brazil (Paraná Basin

    Directory of Open Access Journals (Sweden)

    Heitor Francischini

    2018-05-01

    Full Text Available Tetrapod tracks in eolianites are widespread in the fossil record since the late Paleozoic. Among these ichnofaunas, the ichnogenus Chelichnus is the most representative of the Permian tetrapod ichnological record of eolian deposits of Europe, North America and South America, where the Chelichnus Ichnofacies often occurs. In this contribution, we describe five sets of tracks (one of which is preserved in cross-section, representing the first occurrence of Dicynodontipus and Chelichnus in the “Pirambóia Formation” of southern Brazil. This unit represents a humid desert in southwestern Pangea and its lower and upper contacts lead us to consider its age as Lopingian–Induan. The five sets of tracks studied were compared with several ichnotaxa and body fossils with appendicular elements preserved, allowing us to attribute these tracks to dicynodonts and other indeterminate therapsids. Even though the “Pirambóia Formation” track record is sparse and sub-optimally preserved, it is an important key to better understand the occupation of arid environments by tetrapods across the Permo–Triassic boundary.

  11. Origin of the Permian-Triassic komatiites, northwestern Vietnam

    Science.gov (United States)

    Hanski, Eero; Walker, Richard J.; Huhma, Hannu; Polyakov, Gleb V.; Balykin, Pavel A.; Tran Trong Hoa; Ngo Thi Phuong

    Rare examples of Phanerozoic komatiites are found in the Song Da zone, NW Vietnam. These komatiites were erupted through continental crust and may belong to the SE extension of the Permo-Triassic Emeishan volcanic province located in SW China. They provide a good opportunity to study the source characteristics of starting plume magmas in a continental flood basalt province. Erupted on late-Permian carbonate rocks, the komatiitic rocks are interbedded with low-Ti olivine basalts. Basaltic komatiites display pyroxene spinifex textures, while more magnesian rocks (MgO up to 32 wt.%) are porphyritic, containing a single, cognate population of euhedral to elongated olivine phenocrysts with Fo up to 93.0%. This suggests a highly magnesian parental magma with 22-23 wt.% MgO. In terms of major and minor elements, the komatiites are similar to the ca. 89 Ma old Gorgona Island komatiites of Colombia. The Song Da komatiites are also strongly light-rare-earth-element- (LREE) depleted (CeN/YbN 0.30-0.62) and have unfractionated heavy rare earth element (HREE) patterns. The komatiites have high Os concentrations (up to 7.0 ppb), low but variable Re/Os ratios, and define an isochron with an age of 270+/-21 Ma, and an initial 188Os/187Os ratio of 0.12506+/- 0.00041 (γOs=+0.02+/-0.40). The Os isotopic systematics of the komatiites show no effects of crustal contamination. In contrast, their initial ɛNd values range from +3 to +8, reflecting varying but generally small degrees of contamination with Proterozoic sialic basement material. Associated low-Ti basalts have low initial ɛNd values (-0.8 to -7.5), high initial γOs values (>=15), flat or LREE-enriched REE patterns, and Nb-Ta depletion. These characteristics are also attributed to variable extents of crustal contamination.

  12. New results of microfaunal and geochemical investigations in the Permian–Triassic boundary interval from the Jadar Block (NW Serbia

    Directory of Open Access Journals (Sweden)

    Sudar Milan N.

    2018-04-01

    Full Text Available Detail results of microfaunal, sedimentological and geochemical investigations are documented from a newly discovered section of the Permian–Triassic boundary (PTB interval in the area of the town of Valjevo (northwestern Serbia. The presence of various and abundant microfossils (conodonts, foraminifers, and ostracodes found in the Upper Permian “Bituminous limestone” Formation enabled a determination of the Changhsingian Hindeodus praeparvus conodont Zone. This paper is the first report of latest Permian strata from the region, as well as from all of Serbia, where the PTB interval sediments have been part of a complex/integrated study by means of biostratigraphy and geochemistry.

  13. The X-Ray Luminosity Functions of Field Low-Mass X-Ray Binaries in Early-Type Galaxies: Evidence for a Stellar Age Dependence

    Science.gov (United States)

    Lehmer, B. D.; Berkeley, M.; Zezas, A.; Alexander, D. M.; Basu-Zych, A.; Bauer, F. E.; Brandt, W. N.; Fragos, T.; Hornschemeier, A. E.; Kalogera, V.; hide

    2014-01-01

    We present direct constraints on how the formation of low-mass X-ray binary (LMXB) populations in galactic fields depends on stellar age. In this pilot study, we utilize Chandra and Hubble Space Telescope (HST) data to detect and characterize the X-ray point source populations of three nearby early-type galaxies: NGC 3115, 3379, and 3384. The luminosity-weighted stellar ages of our sample span approximately equal to 3-10 Gyr. X-ray binary population synthesis models predict that the field LMXBs associated with younger stellar populations should be more numerous and luminous per unit stellar mass than older populations due to the evolution of LMXB donor star masses. Crucially, the combination of deep Chandra and HST observations allows us to test directly this prediction by identifying and removing counterparts to X-ray point sources that are unrelated to the field LMXB populations, including LMXBs that are formed dynamically in globular clusters, Galactic stars, and background AGN/galaxies. We find that the "young" early-type galaxy NGC 3384 (approximately equals 2-5 Gyr) has an excess of luminous field LMXBs (L(sub x) approximately greater than (5-10) × 10(exp 37) erg s(exp -1)) per unit K-band luminosity (L(sub K); a proxy for stellar mass) than the "old" early-type galaxies NGC 3115 and 3379 (approximately equals 8-10 Gyr), which results in a factor of 2-3 excess of L(sub X)/L(sub K) for NGC 3384. This result is consistent with the X-ray binary population synthesis model predictions; however, our small galaxy sample size does not allow us to draw definitive conclusions on the evolution field LMXBs in general. We discuss how future surveys of larger galaxy samples that combine deep Chandra and HST data could provide a powerful new benchmark for calibrating X-ray binary population synthesis models.

  14. Early socioeconomic adversity and young adult physical illness: the role of body mass index and depressive symptoms.

    Science.gov (United States)

    Wickrama, K A S; Kwon, Josephine A; Oshri, Assaf; Lee, Tae Kyoung

    2014-10-01

    The present study investigated the psychophysiological inter- and intra-individual processes that mediate the linkage between childhood and/or adolescent socioeconomic adversities and adult health outcomes. Specifically, the proposed model examined the roles of youth depressive symptoms and body mass index (BMI) trajectories as mediators that explain the link between early adversity and young adults' general health and physical illnesses after controlling for gender, race or ethnicity, and earlier general health reports. Using a nationally representative sample of 12,424 from National Longitudinal Study of Adolescent Health (Add Health), this study used growth curve modeling to consider both the severity (initial level) and the change over time (deterioration or elevation) as psychophysiological mediators, thereby acknowledging multiple facets of depressive symptoms and BMI trajectories as psychophysiological mediators of early adversity to adult health. Results provide evidence for (1) the influence of early childhood and early adolescent cumulative socioeconomic adversity on both the initial levels and changes over time of depressive symptoms and BMI and (2) the independent influences depressive symptoms and BMI trajectories on the general health and the physical illnesses of young adults. These findings contribute valuable knowledge to existing research by elucidating how early adversity exerts an enduring long-term influence on physical health problems in young adulthood; furthermore, this information suggests that effective intervention and prevention programs should incorporate multiple facets (severity and change over time) of multiple mechanisms (psychological and physiological). Published by Elsevier Inc.

  15. Muddy and dolomitic rip-up clasts in Triassic fluvial sandstones: Origin and impact on potential reservoir properties (Argana Basin, Morocco)

    Science.gov (United States)

    Henares, Saturnina; Arribas, Jose; Cultrone, Giuseppe; Viseras, Cesar

    2016-06-01

    The significance of rip-up clasts as sandstone framework grains is frequently neglected in the literature being considered as accessory components in bulk sandstone composition. However, this study highlights the great value of muddy and dolomitic rip-up clast occurrence as: (a) information source about low preservation potential from floodplain deposits and (b) key element controlling host sandstone diagenetic evolution and thus ultimate reservoir quality. High-resolution petrographic analysis on Triassic fluvial sandstones from Argana Basin (T6 and T7/T8 units) highlights the significance of different types of rip-up clasts as intrabasinal framework components of continental sediments from arid climates. On the basis of their composition and ductility, three main types are distinguished: (a) muddy rip-up clasts, (b) dolomitic muddy rip-up clasts and (c) dolomite crystalline rip-up clasts. Spatial distribution of different types is strongly facies-related according to grain size. Origin of rip-up clasts is related to erosion of coeval phreatic dolocretes, in different development stages, and associated muddy floodplain sediments. Cloudy cores with abundant inclusions and clear outer rims of dolomite crystals suggest a first replacive and a subsequent displacive growth, respectively. Dolomite crystals are almost stoichiometric. This composition is very similar to that of early sandstone dolomite cement, supporting phreatic dolocretes as dolomite origin in both situations. Sandstone diagenesis is dominated by mechanical compaction and dolomite cementation. A direct correlation exists between: (1) muddy rip-up clast abundance and early reduction of primary porosity by compaction with irreversible loss of intergranular volume (IGV); and (2) occurrence of dolomitic rip-up clasts and dolomite cement nucleation in host sandstone, occluding adjacent pores but preserving IGV. Both processes affect reservoir quality by generation of vertical and 3D fluid flow baffles and

  16. Provenance and detrital zircon geochronologic evolution of lower Brookian foreland basin deposits of the western Brooks Range, Alaska, and implications for early Brookian tectonism

    Science.gov (United States)

    Moore, Thomas; O'Sullivan, Paul B.; Potter, Christopher J.; Donelick, Raymond A.

    2015-01-01

    The Upper Jurassic and Lower Cretaceous part of the Brookian sequence of northern Alaska consists of syntectonic deposits shed from the north-directed, early Brookian orogenic belt. We employ sandstone petrography, detrital zircon U-Pb age analysis, and zircon fission-track double-dating methods to investigate these deposits in a succession of thin regional thrust sheets in the western Brooks Range and in the adjacent Colville foreland basin to determine sediment provenance, sedimentary dispersal patterns, and to reconstruct the evolution of the Brookian orogen. The oldest and structurally highest deposits are allochthonous Upper Jurassic volcanic arc–derived sandstones that rest on accreted ophiolitic and/or subduction assemblage mafic igneous rocks. These strata contain a nearly unimodal Late Jurassic zircon population and are interpreted to be a fragment of a forearc basin that was emplaced onto the Brooks Range during arc-continent collision. Synorogenic deposits found at structurally lower levels contain decreasing amounts of ophiolite and arc debris, Jurassic zircons, and increasing amounts of continentally derived sedimentary detritus accompanied by broadly distributed late Paleozoic and Triassic (359–200 Ma), early Paleozoic (542–359 Ma), and Paleoproterozoic (2000–1750 Ma) zircon populations. The zircon populations display fission-track evidence of cooling during the Brookian event and evidence of an earlier episode of cooling in the late Paleozoic and Triassic. Surprisingly, there is little evidence for erosion of the continental basement of Arctic Alaska, its Paleozoic sedimentary cover, or its hinterland metamorphic rocks in early foreland basin strata at any structural and/or stratigraphic level in the western Brooks Range. Detritus from exhumation of these sources did not arrive in the foreland basin until the middle or late Albian in the central part of the Colville Basin.These observations indicate that two primary provenance areas provided

  17. Early Jurassic extensional inheritance in the Lurestan region of the Zagros fold-and-thrust belt, Iran.

    Science.gov (United States)

    Tavani, Stefano; Parente, Mariano; Vitale, Stefano; Puzone, Francesco; Erba, Elisabetta; Bottini, Cinzia; Morsalnejad, Davoud; Mazzoli, Stefano

    2017-04-01

    It has long been recognized that the tectonic architecture of the Zagros mountain belt was strongly controlled by inherited structures previously formed within the Arabian plate. These preexisting features span in age from the pre-Cambrian to the Mesozoic, showing different trends and deformation styles. Yet, these structures are currently not fully understood. This uncertainty is partly related with the paucity of exposures, which rarely allows a direct observation of these important deformation features. The Lurestan Province of Iran provides a remarkable exception, since it is one of the few places of the Zagros mountain belt where exposures of Triassic and Jurassic rocks are widespread. In this area we carried out structural observations on Mesozoic extensional structures developed at the southern margin of the Neo-Tethyan basin. Syn-sedimentary extensional faults are hosted within the Triassic-Cretaceous succession, being particularly abundant in the Jurassic portion of the stratigraphy. Early to Middle Jurassic syn-sedimentary faults are observed in different paleogeographic domains of the area, and their occurrence is coherent with the subsequent transition from shallow-water to deep-sea basin environments, observed in a wide portion of the area. Most of the thrusts exposed in the area may indeed be interpreted as reactivated Jurassic extensional faults, or as reverse faults whose nucleation was controlled by the location of preexisting normal faults, as a result of positive inversion during crustal shortening and mountain building.

  18. Sedimentary facies and depositional environments of early Mesozoic Newark Supergroup basins, eastern North America

    Science.gov (United States)

    Smoot, J.P.

    1991-01-01

    The early Mesozoic Newark Supergroup consists of continental sedimentary rocks and basalt flows that occupy a NE-trending belt of elongate basins exposed in eastern North America. The basins were filled over a period of 30-40 m.y. spanning the Late Triassic to Early Jurassic, prior to the opening of the north Atlantic Ocean. The sedimentary rocks are here divided into four principal lithofacies. The alluvial-fan facies includes deposits dominated by: (1) debris flows; (2) shallow braided streams; (3) deeper braided streams (with trough crossbeds); or (4) intense bioturbation or hyperconcentrated flows (tabular, unstratified muddy sandstone). The fluvial facies include deposits of: (1) shallow, ephemeral braided streams; (2) deeper, flashflooding, braided streams (with poor sorting and crossbeds); (3) perennial braided rivers; (4) meandering rivers; (5) meandering streams (with high suspended loads); (6) overbank areas or local flood-plain lakes; or (7) local streams and/or colluvium. The lacustrine facies includes deposits of: (1) deep perennial lakes; (2) shallow perennial lakes; (3) shallow ephemeral lakes; (4) playa dry mudflats; (5) salt-encrusted saline mudflats; or (6) vegetated mudflats. The lake margin clastic facies includes deposits of: (1) birdfoot deltas; (2) stacked Gilbert-type deltas; (3) sheet deltas; (4) wave-reworked alluvial fans; or (5) wave-sorted sand sheets. Coal deposits are present in the lake margin clastic and the lacustrine facies of Carnian age (Late Triassic) only in basins of south-central Virginia and North and South Carolina. Eolian deposits are known only from the basins in Nova Scotia and Connecticut. Evaporites (and their pseudomorphs) occur mainly in the northern basins as deposits of saline soils and less commonly of saline lakes, and some evaporite and alkaline minerals present in the Mesozoic rocks may be a result of later diagenesis. These relationships suggest climatic variations across paleolatitudes, more humid to the

  19. The Triassic-Liassic volcanic sequence and rift evolution in the Saharan Atlas basins (Algeria). Eastward vanishing of the Central Atlantic magmatic province

    Energy Technology Data Exchange (ETDEWEB)

    Meddah, A.; Bertrand, H.; Seddiki, A.; Tabeliouna, M.

    2017-11-01

    We investigate the Triassic-Liassic sequence in ten diapirs from the Saharan Atlas (Algeria). Based on detailed mapping, two episodes are identified. The first one consists of a volcano-sedimentary sequence in which three volcanic units were identified (lower, intermediate and upper units). They are interlayered and sometimes imbricated with siliciclastic to evaporitic levels which record syn-sedimentary tectonics. This sequence was deposited in a lagoonal-continental environment and is assigned to the Triassic magmatic rifting stage. The second episode, lacking lava flows (post magmatic rifting stage), consists of carbonate levels deposited in a lagoonal to marine environment during the Rhaetian-Hettangian. The volcanic units consist of several thin basaltic flows, each 0.5 to 1m thick, with a total thickness of 10–15m. The basalts are low-Ti continental tholeiites, displaying enrichment in large ion lithophile elements and light rare earth elements [(La/Yb)n= 2.5-6] with a negative Nb anomaly. Upwards decrease of light-rare-earth-elements enrichment (e.g. La/Yb) is modelled through increasing melting rate of a spinel-bearing lherzolite source from the lower (6–10wt.%) to the upper (15–20wt.%) unit. The lava flows from the Saharan Atlas share the same geochemical characteristics and evolution as those from the Moroccan Atlas assigned to the Central Atlantic magmatic province. They represent the easternmost witness of this large igneous province so far known.

  20. The Triassic-Liassic volcanic sequence and rift evolution in the Saharan Atlas basins (Algeria). Eastward vanishing of the Central Atlantic magmatic province

    International Nuclear Information System (INIS)

    Meddah, A.; Bertrand, H.; Seddiki, A.; Tabeliouna, M.

    2017-01-01

    We investigate the Triassic-Liassic sequence in ten diapirs from the Saharan Atlas (Algeria). Based on detailed mapping, two episodes are identified. The first one consists of a volcano-sedimentary sequence in which three volcanic units were identified (lower, intermediate and upper units). They are interlayered and sometimes imbricated with siliciclastic to evaporitic levels which record syn-sedimentary tectonics. This sequence was deposited in a lagoonal-continental environment and is assigned to the Triassic magmatic rifting stage. The second episode, lacking lava flows (post magmatic rifting stage), consists of carbonate levels deposited in a