Sample records for early permian rio

  1. Early Permian bipedal reptile.

    Berman, D S; Reisz, R R; Scott, D; Henrici, A C; Sumida, S S; Martens, T


    A 290-million-year-old reptilian skeleton from the Lower Permian (Asselian) of Germany provides evidence of abilities for cursorial bipedal locomotion, employing a parasagittal digitigrade posture. The skeleton is of a small bolosaurid, Eudibamus cursoris, gen. et sp. nov. and confirms the widespread distribution of Bolosauridae across Laurasia during this early stage of amniote evolution. E. cursoris is the oldest known representative of Parareptilia, a major clade of reptiles.

  2. Palaeobotanical evidence of wildfires in the Late Palaeozoic of South America - Early Permian, Rio Bonito Formation, Paraná Basin, Rio Grande do Sul, Brazil

    Jasper, André; Uhl, Dieter; Guerra-Sommer, Margot; Mosbrugger, Volker


    Fossil charcoal, as direct evidence of palaeowildfires, has repeatedly been reported from several plant-bearing deposits from the Late Palaeozoic of the Northern Hemisphere. In contrast charcoal reports from the Late Palaeozoic deposits of the Southern Hemisphere are relatively rare in comparison to the Northern Hemisphere. Although the presence of pyrogenic coal macerals has repeatedly been reported from Late Palaeozoic coals from South America, no detailed anatomical investigations of such material have been published so far. Here is presented an anatomical analysis of charcoal originating from Early Permian sediments of the Quitéria Outcrop, Rio Bonito Formation, Paraná Basin, located in the central-eastern portion of Rio Grande do Sul, Brazil. This charcoal comes from two different coaly facies, and it was possible to scrutinize between three types, based on anatomical characters of the charcoal. Two of these charcoal types can be correlated to gymnosperm woods, and the other type corresponds to strongly permineralized bark with characteristic features of lycopsids. The presence of charcoal in different facies, ranging from parautochtonous to allochtonous origin, indicates that different vegetation types, i.e. plants which grew under wet conditions in the lowland as well as in the more dry hinterland, have experienced wildfires. Taking into account previous petrographic and lithological analyses from the facies in which the charcoal occurs and from the conditions of the wood and bark fragments, it was possible to speculate that the intensity of such wildfires most probably corresponds to forest-crown fires. Moreover, it is possible to state that wildfires have been a more or less common element in distinct Late Palaeozoic terrestrial ecosystems in the South American part of Gondwana. The data support previous assumptions on the occurrence of wildfires in the Early Permian of the Paraná Basin which were based solely on coal-petrographic data.

  3. Early Permian Pangea `B' to Late Permian Pangea `A'

    Muttoni, Giovanni; Kent, Dennis V.; Garzanti, Eduardo; Brack, Peter; Abrahamsen, Niels; Gaetani, Maurizio


    The pre-drift Wegenerian model of Pangea is almost universally accepted, but debate exists on its pre-Jurassic configuration since Ted Irving introduced Pangea 'B' by placing Gondwana farther to the east by ˜3000 km with respect to Laurasia on the basis of paleomagnetic data. New paleomagnetic data from radiometrically dated Early Permian volcanic rocks from parts of Adria that are tectonically coherent with Africa (Gondwana), integrated with published coeval data from Gondwana and Laurasia, again only from igneous rocks, fully support a Pangea 'B' configuration in the Early Permian. The use of paleomagnetic data strictly from igneous rocks excludes artifacts from sedimentary inclination error as a contributing explanation for Pangea 'B'. The ultimate option to reject Pangea 'B' is to abandon the geocentric axial dipole hypothesis by introducing a significant non-dipole (zonal octupole) component in the Late Paleozoic time-averaged geomagnetic field. We demonstrate, however, by using a dataset consisting entirely of paleomagnetic directions with low inclinations from sampling sites confined to one hemisphere from Gondwana as well as Laurasia that the effects of a zonal octupole field contribution would not explain away the paleomagnetic evidence for Pangea 'B' in the Early Permian. We therefore regard the paleomagnetic evidence for an Early Permian Pangea 'B' as robust. The transformation from Pangea 'B' to Pangea 'A' took place during the Permian because Late Permian paleomagnetic data allow a Pangea 'A' configuration. We therefore review geological evidence from the literature in support of an intra-Pangea dextral megashear system. The transformation occurred after the cooling of the Variscan mega-suture and lasted ˜20 Myr. In this interval, the Neotethys Ocean opened between India/Arabia and the Cimmerian microcontinents in the east, while widespread lithospheric wrenching and magmatism took place in the west around the Adriatic promontory. The general

  4. Progress on the palynostratigraphy of the Permian strata in Rio Grande do Sul State, Paraná Basin, Brazil

    Souza Paulo A.


    Full Text Available A review of published papers and results of analysis of new material have allowed improvements on the palynostratigraphy of the Permian strata of the Paraná Basin in Rio Grande do Sul State. Based on first and last occurrences of certain species of pollen taxa, two palynozones are formalized, these are the Vittatina costabilis and Lueckisporites virkkiae Interval Zones, in ascending order. The Vittatina costabilis Interval Zone is subdivided into two units, in ascending order the Protohaploxypinus goraiensis and the Hamiapollenites karrooensis Subzones, and is recognized from the glacial (Itararé Group and post-glacial sequence (Rio Bonito Formation and the base of the Palermo Formation. The Lueckisporites virkkiae Interval Zone occurs from the uppermost Rio Bonito Formation, through the Palermo and Irati formations, and into the Serra Alta and Teresina formations. The main characteristics and reference sections are established, as well as additional criteria to recognize biostratigraphical units, in accordance with the International Stratigraphic Guide. Palynostratigraphical correlation suggests that the Vittatina costabilis Zone concerns the Early Permian (early Cisuralian, while the Lueckisporites virkkiae is regarded as late Early Permian to early Middle Permian (late Cisularian to early Guadalupian.

  5. The flora of Early Permian coal measures from the Parana Basin in Brazil: A review

    Iannuzzi, Roberto [Centro de Investigacoes do Gondwana, Instituto de Geociencias, Universidade Federal do Rio Grande do Sul, Av. Bento Goncalves, 9500, Porto Alegre, RS, 91.509-900 (Brazil)


    This paper presents an updated overview integrating both previous and newly published data on the most important floras found associated with Early Permian coal seams in the Parana Basin, Brazil. These floras occur within the Rio Bonito Formation and correspond to the Gondwana ''Glossopteris Flora.'' For this review, five floras are selected, in ascending stratigraphic order: the ''Sao Joao do Triunfo,'' ''Figueira,'' ''Quiteria,'' ''Morro do Papaleo'' and ''Irapua Bed'' floras. They are the best-known floras of the basin in terms of taxonomic composition, paleoecology and environments of deposition. An early-mid Sakmarian to earliest Artinskian age is indicated for the Rio Bonito Formation based on absolute radiometric and relative biostratigraphic ages. Integration of available information about the selected floras allows evaluation of taphonomic and paleoecological factors to provide a more comprehensive understanding of the Early Permian floral record in the Parana Basin. The variation observed in both the taxonomic composition of individual floras and in the frequency of occurrence of different plant groups is due to the broad range of environmental/edaphic conditions that prevailed in the many different depositional settings represented in the Rio Bonito Formation. A more precise age determination obtained for the plant-bearing deposits permits the establishment of a more confident correlation between the Early Permian floral succession in the Parana Basin and other Early Permian floral successions in other basins. The Sakmarian global warming favored the appearance of pecopterid and sphenopterid ferns amongst the spore-producing plants, and the glossopterids amongst the pollen-producing plants. (author)

  6. A statistical test on diversity changes of Early and Middle Permian fusulinacean fauna in South China

    SHI Yukun; YANG Xiangning


    The rarefaction analysis has been conducted to test the species diversity changes of the Early and Middle Permian fusulinacean fauna in South China. The results reveal that the number of species dramatically increased since the earliest Permian and quickly reached the maximum value in the early Zisongian representing the highest species diversity for the whole Early and Middle Permian. The species diversity stabilized in the plateau through the Zisongian;however, it started to decline in the following Longlinian and sustained a longstanding low level during the mid-Early Permian. With the appearance of new fusulinacean taxa with septulum structures, the number of species raised again in the late-Early Permian, followed by a decline in the Middle Permian Neoschwagerina simplex zone. Although the species diversity increased apparently in the Kuhfengian, it never rebounded back to the same level as in the Early Permian.In the mid-Middle Permian, species diversity began to decrease continuously and led to the disappearance of most fusulinacean species by the end of the Middle Permian.

  7. Panjal Paleomagnetism: Implications for Early Permian Gondwana break-up

    Stojanovic, D.; Aitchison, J.; Ali, J. R.; Ahmad, T.; Ahmad Dar, R.; Agarwal, A.; Roeder, T.


    The mid-Early Permian represents an important phase in Pangaea's development marking the time when the >13,000 km-long string of terranes that are collectively known as 'Cimmeria' separated from Gondwana's Tethyan margin (northern Africa-NE Arabia-northern India-NW and northern Australia). The ~289 Ma Panjal Traps of NW India (Kashmir) are one of a number of mafic suites (Abor, Sikkim etc.) that were erupted onto the Indian block possibly during the separation of the Lhasa/SE Qiangtang block. Herein, we report data from the first modern paleomagnetic study of the unit. Results from four quarry sections (15 individual cooling units) from a locality close to Srinagar together form a tectonically coherent sequence spanning 2-3 km of stratigraphy. The derived direction and paleopole yield key new information concerning (1) the Early Permian location of India, and by inference that of central Gondwana, and (2) inform debates related to Cimmeria's breakup from eastern Gondwana. Moreover, they provide a new independent control for assessing NW Greater India's extent prior to its collision with Asia and the amount of vertical-axis rotation this sector of the Himalayan range experienced in the mid to late Cenozoic.

  8. The paleoclimate of the Kazanian (early Late Permian) world

    Moore, G.T. (Chevron Oil Field Research Co., La Habra, CA (USA)); Peoples, C.J. (Texas A and M Univ., College Station (USA))


    The Kazanian (early Late Permian, 258-253 m.y.) marked the onset of a unique interval in the Phanerozoic, distinguished by a classic end-member case of continental assembly, the megacontinent Pangaea. Compilation of biostratigraphic and lithofacies data indicate a warm, extensively arid world, largely ice free, and characterized by the onset of atmospheric conditions that were exceedingly stressful to the biosphere, the worst of the eon. Using Chevron's version of the Community Climate model, the authors report here on two Kazanian paleoclimate seasonal simulations, one using 200 ppm CO{sub 2} and the other with 2,000 ppm CO{sub 2}. The authors consider the knowledge of plate assembly back to the Permian accurate enough to allow employment of a seasonal model. Simulation of a warmer Earth with an elevated greenhouse effect (modeled as CO{sub 2}) fits the observed geology and isotope signals. The increased CO{sub 2} experiment warmed the entire planet with the greatest increases north of 50{degree} latitude and least changes in the tropics. The warming caused the poleward retreat of sea ice in both hemispheres. Precipitation and evaporation increased, but runoff was confined to areas of very intense rainfall. Monsoons are limited to the southern hemisphere, associated with the western Tethys sea and the eastern equatorial Panthalassa ocean. Extreme southeastern Pangaea (northern Australia) was a focus of precipitation throughout the year. Precipitation occurred in the higher latitudes (50-55{degree}) on the western coast, of Pangaea where storm tracks make landfall. High evaporation rates characterized the restricted Permian (US), Zechstein, and Perm (Soviet Union) basins, a time of evaporite, deposition. Interior Pangaea at middle to high latitudes endured frigid winters ({minus}40{degree}C) and torrid summers (60{degree}C).

  9. Late Carboniferous and Early Permian algal microflora (Liuzhou, South China)

    Li Chun (Nanjing Inst. of Geology and Paleontology, Nanjing (China)); Mamet, B. (Montreal Univ., PQ (Canada))


    Paleozoic algae are important contributors to the formation of carbonate sediments. A study was conducted to compare an Upper Carboniferous-Lower Permian carbonate succession of South China with what is generally observed in the Tethys Sea and North America. The sequence studied was in the Maping Limestone at the Loutishan section in Liuzhou, with some additional information derived from the Maping Limestone at Shangxinping and from the Chuanshan Limestone at Yuhuangshan and in the Nanjing Hills. In the Late Carboniferous, the beresellid algae are important builders and the Tubiphytes-Archaeolithophyllum community is locally replaced by a Tubiphytes-Ungdarella association. Early Permian oncolites were formed by complex Osagia, then by Esmerella. The Asselian Sphaeroschwagerina fauna is underlined by the outburst of epimastoporid dasycladales and phylloid Anchicodiaceae. Microcodium is observed at the contact of the Sphaeroschwagerina moelleri/sphaerica Zones. The overlying and paraconformable Chihsia Limestone is characterized by a Gymnocodium-Gyroporella-Mizzia flora. In most cases, the base of the fusulinid zones, therefore, coincides with an important local floristic change. 72 refs., 4 figs.

  10. Dispersal syndromes of fossil seeds from the Lower Permian of Paraná Basin, Rio Grande do Sul, Brazil.

    Souza, Juliane M; Iannuzzi, Roberto


    The morphological analysis of seeds has been an important subject in modern ecological studies, once it provides evidence about the biology and adaptations of the parent plant. However, this kind of study has been restricted to the ecology of modern plants and is rarely used in interpretations of Paleozoic data. From the understanding of dispersal syndromes analysis as an important tool to paleoecological reconstruction, this study provides a first approach using this tool with seeds from the Lower Permian strata of southern Paraná Basin in Rio Grande do Sul. Based on previously classified seeds and using their biological and taphonomic data, the syndrome of dispersal was interpreted, and their placement in successional groups (pioneer, early-successional and later-successional) was suggested. Seven morphospecies were analyzed: Samaropsis gigas, representing a later-successional species living in water bodies with hydrochory as its dispersal syndrome; Samaropsis kurtzii, typical of early-successional species showing anemochory as its dispersal syndrome and living in distal areas in relation to water bodies; Samaropsis aff. S. millaniana, Cordaicarpus aff. C. brasilianus, Cordaicarpus cerronegrensis and Cordaicarpus truncata have typical characteristics of pioneer plants, exhibiting barochory as their primary dispersal syndrome with other syndromes associated.

  11. Siderite deposits in northern Italy: Early Permian to Early Triassic hydrothermalism in the Southern Alps

    Martin, Silvana; Toffolo, Luca; Moroni, Marilena; Montorfano, Carlo; Secco, Luciano; Agnini, Claudia; Nimis, Paolo; Tumiati, Simone


    We present a minero-petrographic, geochemical and geochronological study of siderite orebodies from different localities of the Southern Alps (northern Italy). Siderite occurs as veins cutting the Variscan basement and the overlying Lower Permian volcano-sedimentary cover (Collio Fm.), and as both veins and conformable stratabound orebodies in the Upper Permian (Verrucano Lombardo and Bellerophon Fms.) and Lower Triassic (Servino and Werfen Fms.) sedimentary sequences of the Lombardian and the Venetian Alps. All types of deposits show similar major- and rare-earth (REE)-element patterns, suggesting a common iron-mineralizing event. The compositions of coexisting siderite, Fe-rich dolomite and calcite suggest formation from hydrothermal fluids at relatively high temperature conditions (≥ 250 °C). Geochemical modelling, supported by REE analyses and by literature and new δ13C and δ18O isotopic data, suggests that fluids responsible for the formation of siderite in the Variscan basement and in the overlying Lower Permian cover were derived from dominant fresh water, which leached Fe and C from volcanic rocks (mainly rhyolites/rhyodacites) and organic carbon-bearing continental sediments. On the basis of U-Th-Pb microchemical dating of uraninite associated with siderite in the Val Vedello and Novazza deposits (Lombardian Alps), the onset of hydrothermalism is constrained to 275 ± 13 Ma (Early-Mid Permian), i.e., it was virtually contemporaneous to the plutonism and the volcanic-sedimentary cycle reported in the same area (Orobic Basin). The youngest iron-mineralizing event is represented by siderite veins and conformable orebodies hosted in Lower Triassic shallow-marine carbonatic successions. In this case, the siderite-forming fluids contained a seawater component, interacted with the underlying Permian successions and eventually replaced the marine carbonates at temperatures of ≥ 250 °C. The absence of siderite in younger rocks suggests an Early Triassic

  12. Discovery of Early Permian Reefs in Xichang Basin, Southwestern China

    Qin Jianxiong; Zeng Yunfu; Wu Yong


    @@ In 1992, Lower Permian reefs were found in various stratigraphic sections in such counties as Ganluo,Puge, and Butuo, in Xichang Basin. This discovery not only promotes the development of Lower Permain sedimentology in Xichang Basin, but also is of great significance for oil-gas exploration along the western margin of Yangtze platform.

  13. Qasimia gen. nov., an early Marattia-like fern from the Permian of Saudi Arabia

    Hill, C.R.; Wagner, R.H.; El-Khayal, A. A.


    The marattialean fern Qasimia schyfsmae (Lemoigne) gen. et comb. nov. is described from the Late Permian plant bed at Unayzah in central Saudi Arabia. Although no organic matter is preserved, impregnation of the compressions by iron minerals at an early stage of diagenesis has partly mineralised the

  14. Stratigraphic and palaeoenvironmental framework of the Early Permian sequence in the Salt Range, Pakistan

    Shahid Ghazi; Nigel P Mountney; Aftab Ahmad Butt; Sadaf Sharif


    The Early Permian Gondwana regime succession of the Nilawahan Group is exposed only in the Salt Range of Pakistan. After a prolonged episode of non-deposition that spanned much of the Palaeozoic, the 350 m thick predominantly clastic sequence of the Nilawahan Group records a late glacial and post-glacial episode in which a range of glacio-fluvial, marine and fluvial environments evolved and accumulated. The Early Permian succession of the Salt Range has been classified into four formations, which together indicates a changing climatic regime during the Early Permian in the Salt Range region. The lower-most, Tobra Formation unconformably overlies a Cambrian sequence and is composed of tillite, diamictite and fresh water facies, which contain a floral assemblage (Gangamopteris and Glossopteris) that confirms an Asselian age. The Tobra Formation is overlain by marginal marine deposits of the Dandot Formation (Sakmarian), which contain an abundant brachiopods assemblage (Eurydesma and Conularia). Accumulation of the Dandot Formation was terminated by a regional sea-level fall and a change to the deposition of the fluvial deposits of the Warchha Sandstone (Artinskian). The Warchha Sandstone was deposited by high sinuosity meandering, avulsion prone river with well developed floodplains. This episode of fluvial sedimentation was terminated by a widespread marine transgression, as represented by the abrupt upward transition to the overlying shallow marine Sardhai Formation (Kungurian). The Early Permian Gondwana sequence represented by the Nilawahan Group is capped by predominantly shallow shelf carbonate deposits of the Tethyan realm. The sedimentologic and stratigraphic relationship of these four lithostratigraphic units in the Salt Range reveals a complex stratigraphic history for the Early Permian, which is mainly controlled by eustatic sea-level change due to climatic variation associated with climatic amelioration at the end of the major Gondwana glacial episode

  15. Permian-Early Triassic tectonics and stratigraphy of the Karoo Supergroup in northwestern Mozambique

    Bicca, Marcos Müller; Philipp, Ruy Paulo; Jelinek, Andrea Ritter; Ketzer, João Marcelo Medina; dos Santos Scherer, Claiton Marlon; Jamal, Daúd Liace; dos Reis, Adriano Domingos


    The Gondwana continent was the base of great basin inception, sedimentation and magmatism throughout the Cambrian to Middle Jurassic periods. The northwestern Mozambique igneous and metamorphic basement assemblages host the NW-trending Moatize Minjova Basin, which has great economic potential for coal and gas mining. This rift basin was activated by an S-SW stress field during the Early Permian period, as constrained by regional and field scale structural data. Tectonically induced subsidence in the basin, from the reactivation of NW-SE and NNE-SSW regional structures is well recorded by faults, folds and synsedimentary fractures within the Early Late Permian Moatize Formation. NW-SE, N-S and NE-SW field structures consist of post-Karoo reactivation patterns related to a NNE-SSW extension produced by the Pangea breakup and early inception stages of the Great East African Rift System. The Early Late Permian sequences of the Moatize-Minjova Basin are composed of fluvial meandering, coal-bearing beds of the Moatize Formation, which comprises mostly floodplain, crevasse splay and fluvial channel lithofacies associations, deposited in a cyclic pattern. This sequence was overlapped by a multiple-story, braided fluvial plain sequence of the Matinde Formation (Late Permian - Early Triassic). Lithofacies associations in the Matinde Formation and its internal relationships suggest deposition of poorly channelized braided alluvial plain in which downstream and probably lateral accretion macroforms alternate with gravity flow deposits. NW paleoflow measurements suggest that Permian fluvial headwaters were located somewhere southeast of the study area, possibly between the African and Antarctic Precambrian highlands.

  16. Integrated Sr isotope variations and global environmental changes through the Late Permian to early Late Triassic

    Song, Haijun; Wignall, Paul B.; Tong, Jinnan; Song, Huyue; Chen, Jing; Chu, Daoliang; Tian, Li; Luo, Mao; Zong, Keqing; Chen, Yanlong; Lai, Xulong; Zhang, Kexin; Wang, Hongmei


    New 87Sr/86Sr data based on 127 well-preserved and well-dated conodont samples from South China were measured using a new technique (LA-MC-ICPMS) based on single conodont albid crown analysis. These reveal a spectacular climb in seawater 87Sr/86Sr ratios during the Early Triassic that was the most rapid of the Phanerozoic. The rapid increase began in Bed 25 of the Meishan section (GSSP of the Permian-Triassic boundary, PTB), and coincided closely with the latest Permian extinction. Modeling results indicate that the accelerated rise of 87Sr/86Sr ratios can be ascribed to a rapid increase (>2.8×) of riverine flux of Sr caused by intensified weathering. This phenomenon could in turn be related to an intensification of warming-driven runoff and vegetation die-off. Continued rise of 87Sr/86Sr ratios in the Early Triassic indicates that continental weathering rates were enhanced >1.9 times compared to those of the Late Permian. Continental weathering rates began to decline in the middle-late Spathian, which may have played a role in the decrease of oceanic anoxia and recovery of marine benthos. The 87Sr/86Sr values decline gradually into the Middle Triassic to an equilibrium values around 1.2 times those of the Late Permian level, suggesting that vegetation coverage did not attain pre-extinction levels thereby allowing higher runoff.

  17. Palaeomagnetism of the Early Permian Mount Leyshon Intrusive Complex and Tuckers Igneous Complex, North Queensland, Australia

    Clark, D. A.; Lackie, M. A.


    This study provides reliable, precisely defined and well-dated Early Permian (286 +/- 6 Ma) palaeomagnetic poles for Australia from the Mount Leyshon Intrusive Complex (MLIC) and the Tuckers Igneous Complex (TIC). Both complexes are associated with prominent negative magnetic anomalies, indicating the presence of rocks carrying stable remanence of reverse polarity, with a Koenigsberger ratio greater than unity. The characteristic remanence carried by the intrusive phases and by locally remagnetized, contact-metamorphosed host rocks is always of reverse polarity, consistent with acquisition during the Permo-Carboniferous (Kiaman) Reverse Superchron. The corresponding palaeopoles confirm that Australia occupied high latitudes in the Early Permian. The pole positions are: MLIC: lat. = 43.2 °S, long. = 137.3 °E dp = 6.0°, dm = 6.4° Q= 6; TIC: lat. = 47.5 °S, long. = 143.0 °E, dp = 6.0°, dm = 6.6° Q= 6. Permian palaeomagnetic overprinting is detectable at considerable distances from the MLIC (2-3 km), well beyond the zone of visible alteration. The primary nature of the Early Permian palaeomagnetic signature is established by full baked contact/aureole tests at both localities. Other new data from Australia are consistent with the poles reported here. Comparison of the Australian, African and South American Apparent Polar Wander Paths (APWP) suggests that mean Permian and Triassic poles from West Gondwana, particularly from South America, are biased by remagnetization in the Jurassic-Cretaceous and that the Late Palaeozoic-Mesozoic APWP for Gondwana is best defined by Australian data. The Australian APWP exhibits substantial movement through the Mesozoic. Provided only that the time-averaged palaeofield was zonal, the Early Triassic palaeomagnetic data from Australia provide an important palaeogeographic constraint that the south geographic pole was within, or very close to, SE Australia around 240 Ma. The new Early Permian poles are apparently more consistent

  18. Depositional model of Early Permian reef-island ocean inEastern Kunlun


    Many fusulinid fossils have been found in thin- to middle-bedded limestones which aredistributed between the Early Permian limestone hills and formerly considered as Early Triassic.The fusulinid fossils, identified as Neoshwagerina sp., Verbeekina sp. and Schwagerina sp., canalso be found in massive limestone hills. At the same time, Early Permian radiolarian chert of deepbasin facies was discovered in Animaqing. All the above show that the massive limestone hills,thin- to middle-bedded limestones and radiolarian chert belong to syndeposits in Early Permianocean. The sediments in the study area can roughly be divided into three types: shallow facies,basin facies and transitional facies. The carbonate buildup can be subdivided into massive bioclas-tic limestone and reef framestone. Basin facies contains thin- or middle-bedded limestone, abyssalred mudstone or ooze, blue-green mudstone and radiolarian chert. Transitional facies includes reeftalus and platformal skirt facies. The Early Permian ocean in Eastern Kunlun is recognized as akind of reef-island ocean environment according to distribution and composition of different facies.The reef-island ocean in Eastern Kunlun is characterized by reef islands (or carbonate buildups)alternating with basins, complicated sea-floor topography, sharp facial change and well-developedreefs

  19. Dispersal Syndromes of fossil Seeds from the Lower Permian of Paraná Basin, Rio Grande do Sul, Brazil

    Juliane M. Souza


    Full Text Available The morphological analysis of seeds has been an important subject in modern ecological studies, once it provides evidence about the biology and adaptations of the parent plant. However, this kind of study has been restricted to the ecology of modern plants and is rarely used in interpretations of Paleozoic data. From the understanding of dispersal syndromes analysis as an important tool to paleoecological reconstruction, this study provides a first approach using this tool with seeds from the Lower Permian strata of southern Paraná Basin in Rio Grande do Sul. Based on previously classified seeds and using their biological and taphonomic data, the syndrome of dispersal was interpreted, and their placement in successional groups (pioneer, early-successional and later-successional was suggested. Seven morphospecies were analyzed: Samaropsis gigas, representing a later-successional species living in water bodies with hydrochory as its dispersal syndrome; Samaropsis kurtzii, typical of early-successional species showing anemochory as its dispersal syndrome and living in distal areas in relation to water bodies; Samaropsis aff. S. millaniana, Cordaicarpus aff. C. brasilianus, Cordaicarpus cerronegrensis and Cordaicarpus truncata have typical characteristics of pioneer plants, exhibiting barochory as their primary dispersal syndrome with other syndromes associated.A Análise morfológica das sementes tem sido um importante objeto em estudos ecológicos modernos, uma vez que fornece evidências sobre a biologia e as adaptações das plantas-mãe de sementes. Entretanto, este tipo de estudo tem sido restrito a ecologia de plantas modernas e é raramente utilizado em interpretações de dados paleozóicos. A partir do entendimento da análise das sindromes de dispersão como uma importante ferramenta para reconstruções paleoecológicas, este estudo oferece uma primeira abordagem utilizando essa ferramenta com sementes do Permiano Inferior do Sul da




    Full Text Available Representatives of the Syringothyrididae (Spiriferinida are a significant component of the Early Permian brachiopod faunas, even if usually subordinate in number of specimens, except for the Interior of Oman, where they dominate the faunal assemblage. A new genus Pachycyrtella - with type-species P. omanensis n. sp. - is here established in order to include gondawanan species - previously assigned to the genus Cyrtella - characterized by high and apsacline to orthocline ventral interarea, thick umbo and deep dorsal median furrow. 

  1. Abrupt environmental and climatic change during the deposition of the Early Permian Haushi limestone, Oman

    Stephenson, Michael; Angiolini, Lucia; Leng, Melanie; Brewer, T. S.; Berra, F.; F. Jadoul; G. Gambacorta; Verna, V.; Al Beloushi, B.


    During the late Sakmarian (Early Permian), the Haushi limestone was deposited in a shallow embayment of the Neotethys Ocean covering what is now north Oman and parts of southeast Saudi Arabia. The sea persisted through the late Sakmarian, but by the time of the deposition of the ?Artinskian Middle Gharif Member, limestone deposition had ceased and generally arid fluvial and minor lacustrine palaeonvironments in a low accommodation space setting had become established. Analysis of three subsur...

  2. Permian, Jurassic and Early Cretaceous palynofloral assemblages from subsurface sedimentary rocks in Chuperbhita Coalfield, Rajmahal Basin, India.

    Tripathi, A


    The results of a palynological analysis of the sedimentary sequence of Borehole RCH-151, Chuperbhita Coalfield, Rajmahal Basin, Bihar are presented here. The borehole penetrated the Rajmahal Formation (comprising two traps sandwiching an intertrappean bed), the thinly represented Dubrajpur Formation and in its lower part, the Coal Measures. The coal-bearing interval is associated with Scheuringipollenites barakarensis, Faunipollenites varius, Densipollenites indicus, Gondisporites raniganjensis and Densipollenites magnicorpus Assemblage Zones. The presence of these biostratigraphic units indicates correlation with the Barakar Formation (Early Permian) and the Barren Measures and Raniganj Formations (both Late Permian). This is the first record, in the Chuperbhita Coalfield, of Late Permian strata, which appear to represent a condensed sequence. Prior to the present study, the Permian succession was thought to have been associated entirely with the Barakar Formation. The overlying Dubrajpur Formation yielded a distinct spore-pollen assemblage (in association with the first report of dinoflagellate, Phallocysta), which is assigned to the newly identified Callialasporites turbatus palynozone of latest Early to early Middle Jurassic age. The diverse spore-pollen flora of the intertrappean bed (Rajmahal Formation) incorporates several age marker taxa, viz. Undulatisporites, Leptolepidites, Klukisporites, Ruffordiaspora, and Coptospora. The assemblages from intertrappean beds are correlated with the Ruffordiaspora australiensis palynozone of Australia. Thus the palynodating indicates Permian, latest Early to early Mid-Jurassic and Early Cretaceous age for the strata studied. This is the first record of definite Jurassic microfossils from the non-marine sequence of Rajmahal Basin, India.

  3. Late Pennsylvanian and early permian chondrichthyan microremains from San Salvador Patlanoaya (Puebla, Mexico)

    Derycke-Khatir, C.; Vachard, D.; Degardin, J.-M.; Flores de Dios, A.; Buitron, B.; Hansen, M.


    The San Salvador Patlanoaya section (Puebla State, Mexico) is known for its richness of many fossil groups. Among them, the calcareous shells have been principally investigated. This paper deals with Missourian-Virgilian (Late Pennsylvanian) and Leonardian (late Early Permian) Mexican fish remains. A discussion about Helicoprion and related genera, is followed by the systematic description of the revised or discovered taxa: Cooperella typicalis, Moreyella cf. M. typicalis, M. (?) sp., "Sturgeonella" quinqueloba, Hybodontidae gen. sp. 1 and 2, scale indet. Palaeobiogeographic implications are suggested. ?? 2005 Elsevier SAS. All rights reserved.

  4. The Permian system in Kansas

    US Fish and Wildlife Service, Department of the Interior — Rocks of Permian age in Kansas were first recognized in 1895, and by the early 21st century the internationally accepted boundary between the Permian and the...

  5. Response of Late Carboniferous and Early Permian Plant Communities to Climate Change

    Dimichele, William A.; Pfefferkorn, Hermann W.; Gastaldo, Robert A.

    Late Carboniferous and Early Permian strata record the transition from a cold interval in Earth history, characterized by the repeated periods of glaciation and deglaciation of the southern pole, to a warm-climate interval. Consequently, this time period is the best available analogue to the Recent in which to study patterns of vegetational response, both to glacial-interglacial oscillation and to the appearance of warm climate. Carboniferous wetland ecosystems were dominated by spore-producing plants and early gymnospermous seed plants. Global climate changes, largely drying, forced vegetational changes, resulting in a change to a seed plant-dominated world, beginning first at high latitudes during the Carboniferous, reaching the tropics near the Permo-Carboniferous boundary. For most of this time plant assemblages were very conservative in their composition. Change in the dominant vegetation was generally a rapid process, which suggests that environmental thresholds were crossed, and involved little mixing of elements from the wet and dry floras.

  6. Response of Late Carboniferous and Early Permian plant communities to climate change

    DiMichele, W.A.; Pfefferkorn, H.W.; Gastaldo, R.A. [Smithsonian Institute, Washington, DC (USA). National Museum of National History


    Late Carboniferous and Early Permian strata record the transition from a cold interval in Earth history, characterized by the repeated periods of glaciation and deglaciation of the southern pole, to a warm-climate interval. Consequently, this time period is the best available analogue to the Recent in which to study patterns of vegetational response, both to glacial-interglacial oscillation and to the appearance of warm climate. Carboniferous wetland ecosystems were dominated by spore-producing plants and early gymnospermous seed plants. Global climate changes, largely drying, forced vegetational changes, resulting in a change to a seed plant-dominated world, beginning first at high latitudes during the Carboniferous, reaching the tropics near the Permo-Carboniferous boundary. For most of this time plant assemblages were very conservative in their composition. Change in the dominant vegetation was generally a rapid process, which suggests that environmental thresholds were crossed, and involved little mixing of elements from the wet and dry floras.

  7. An early geikiid dicynodont from the Tropidostoma Assemblage Zone (late Permian) of South Africa

    Smith, Roger M.H.


    Based on specimens previously identified as Tropidostoma, a new taxon of dicynodont (Bulbasaurus phylloxyron gen. et sp. nov.) from the Karoo Basin of South Africa is described. Bulbasaurus is a medium-sized dicynodont (maximum dorsal skull length 16.0 cm) restricted to the Tropidostoma Assemblage Zone (early Lopingian) of the Beaufort Group. Bulbasaurus can be distinguished from Tropidostoma by an array of characters including the presence of a tall, sharp premaxillary ridge, large, rugose, nearly-confluent nasal bosses, a nasofrontal ridge, massive tusks, robust pterygoids, prominently twisted subtemporal bar, and absence of a distinct postfrontal. Inclusion of Bulbasaurus in a phylogenetic analysis of anomodont therapsids recovers it as a member of Geikiidae, a clade of otherwise later Permian dicynodonts such as Aulacephalodon and Pelanomodon. Bulbasaurus exhibits many of the characters typical of adult Aulacephalodon, but at substantially smaller skull size (these characters are absent in comparably-sized Aulacephalodon juveniles), suggesting that the evolution of typical geikiid morphology preceded gigantism in the clade. Bulbasaurus is the earliest known geikiid and the only member of the group known from the Tropidostoma Assemblage Zone; discovery of this taxon shortens a perplexing ghost lineage and indicates that abundant clades from the later Permian of South Africa (e.g., Geikiidae, Dicynodontoidea) may have originated as rare components of earlier Karoo assemblage zones. PMID:28168104

  8. Palynological records of the Early Permian icehouse-greenhouse transition (Ecca Group, South Africa)

    Götz, Annette E.; Ruckwied, Katrin


    The Permian coal-bearing formations of the South African Karoo Basin play a crucial role in the study and interpretation of Gondwana's climate history and biodiversity in this time of major global changes in terrestrial and marine ecosystems. Here, we report on new palynological data from the No. 2 coal seam of the northern Witbank coal field, documenting the switch from Icehouse to Greenhouse conditions in the Early Permian (Lower Ecca Group). The studied postglacial fluvio-deltaic deposits of a highly proximal setting comprise coarse-grained to pebbly sandstones, partially with an abrupt upward transition into fine-grained sediments and coal, trough cross-stratified medium- to coarse-grained sandstones, and horizontally laminated fine- to medium-grained sandstones and siltstones. The sedimentary organic matter content clearly documents stratal changes in the palynomorph assemblage and variations in the amount and in the type, size and shape of plant debris. Generally, palynofacies is characterized by a high amount of opaque phytoclasts. Amorphous organic matter is characteristic of laminated siltstones and coals. The palynological record indicates a cold climate, fern wetland community, characteristic of lowland alluvial plains, and an upland conifer community in the lower part of the coal seam. Up section, these communities are replaced by a cool-temperate cycad-like lowland vegetation and gymnospermous upland flora. Ongoing studies focus on the cyclic architecture of the coal seam, applying palynofacies analysis as high-resolution correlation tool with respect to decipher signatures of prominent climate amelioration on basin-wide, intercontinental and intra-Gondwanic scales.

  9. Depositional model of Early Permian reef-island ocean inEastern Kunlun

    WANG; Yongbiao


    [1]Geological and Mineral Resources Bureau of Qinghai Province, Lithological Stratigraphy of Qinghai Province (in Chinesel. Wuhan: China University of Geosciences Press, 1997, 135-137.[2]Jiang Chunfa, Yang Jingsui, Feng Binggui, Opening-closing tectonics of Kunlun Mountains (in Chinese), Geological Memoirs, Series 5, No. 12, Beijing: Geological Publishing House, 1992, 8.[3]Bian Qiantao. Luo Xiaoquan, Li Hongsheng et al., Discovery of early Paleozoic and early Carboniferous to early Permian ophiolite. Scientia Geologica Sinica (in Chinese), 1999, 34(4): 523-524.[4]Yin Hongfu. Zhang Kexin. Characteristics of eastern Kunlun orogenic belt, Earth Science, 1997, 22(4): 339-342.[5]Wang Guocan, Zhang Kexin, Liang Bin et al., Texture and tectonic slices of the eastern Kunlun orogenic belt, Earth Science lin Chinese), 1997, 22(4): 352-356.[6]Zhang Kexin, Huang Jichun, Luo Mansheng et al., Sedimentary geochemical features of Animaqing, Earth Science (in Chinese), 1999, 24(2): 111-115.[7]Wang Yongbiao, Zhang Kexin, Gong Yiming et al., The discovery of early Permian reef belt in east Kunlun and its significance, Chinese Science Bulletin, 1998, 43(11): 947-950.[8]Kuzinizov, B. T., Reef Geology and Its Hydroncarbon-bearing Possibility (translated by Li Jianwen), Beijing: Oil Industry Publishing House, 1983, 24-39.[9]Zhao Qiyuan, Marine Geochemistry (in Chinese), Beijing: Geological Publishing House, 1989, 155.[10]Feng Qinglai, A preliminary study on the radiolarian palaeoecology, Geological Science and Technology Information (in Chinese), 1992, 11,41-46.[11]Chen Zongyuan, Introduction of marine sciences (in Chinese), Qingdao: Qingdao Marine University Press, 1992, 282-284.[12]Jia Zhenyuan, Li Zhiqi, Facies and Sedimentary Environment of Carbonate (in Chinese), Wuhan: China University of Geosciences Press, 1989, 41.[13]Ryuichi Sugisaki, Koshi Yamamoto, Manoru Adachi, Triassic bedded cherts in central Japan are not Pelagic, Nature

  10. Late Early Permian continental ichnofauna from Lake Kemp, north-central Texas, USA

    Lucas, S.G.; Voigt, S.; Lerner, A.J.; Nelson, W.J.


    Continental trace fossils of Early Permian age are well known in the western United States from Wolfcampian (~. Asselian to Artinskian) strata, but few examples are known from Leonardian (~. Kungurian) deposits. A substantial ichnofauna from strata of the lower part of the Clear Fork Formation at Lake Kemp, Baylor County, Texas, augments the meager North American record of Leonardian continental trace fossil assemblages. Ichnofossils at Lake Kemp occur in the informally-named Craddock dolomite member of the Clear Fork Formation, which is 12-15. m above the local base of the Clear Fork. The trace-bearing stratum is an up-to-0.3. m thick, laminated to flaser-bedded, dolomitic siltstone that also contains mud cracks, raindrop impressions, microbially induced mat structures, and some land-plant impressions. We interpret the Craddock dolomite member as the feather-edge of a marine transgressive carbonate deposit of an irregular coastline marked by shallow bays or estuaries on the eastern shelf of the Midland basin, and the trace-fossil-bearing stratum at Lake Kemp is an unchannelized flow deposit on a muddy coastal plain. The fossil site at Lake Kemp yields a low to moderately diverse fauna of invertebrate and vertebrate traces. A sparse invertebrate ichnofauna consists of arthropod feeding and locomotion traces assigned to Walpia cf. W. hermitensis White, 1929 and Diplichnites gouldi Gevers in Gevers et al., 1971. Tetrapod footprints are most common and assigned to Batrachichnus salamandroides (Geinitz, 1861), cf. Amphisauropus kablikae (Geinitz and Deichm??ller, 1882), and Dromopus lacertoides (Geinitz, 1861), which represent small temnospondyl, seymouriamorph, and basal sauropsid trackmakers. Both the traces and sedimentary features of the fossil horizon indicate a freshwater setting at the time of track formation, and the trace assemblage represents the Scoyenia ichnofacies and the Batrachichnus ichnofacies in an overbank environment with sheet flooding and shallow

  11. Early Permian coal-forming floras preserved as compressions from the Wuda District (Inner Mongolia, China)

    Pfefferkorn, Hermann W. [Department of Earth and Environmental Science, University of Pennsylvania, Philadelphia, PA 19104-6316 (United States); Wang, Jun [Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008 (China)


    Four different compression/impression floras are preserved in only 4.32 m of the geologic section in the Early Permian Shanxi Formation of the Wuda District of Inner Mongolia, northwestern China. These floras represent four different plant communities and landscapes that followed each other in time. The oldest flora was rooted in sandy clay and initiated peat accumulation that lead to the formation of the lower coal seam. This seam is 230-cm thick and overlain by a 66-cm thick volcanic tuff that preserves a second different flora that grew on the peat at the time of the ash-fall. Standing stems and large plant parts are present. The upper part of the tuff is rooted by a single species of lycopsid (the third flora) again initiating peat accumulation. On top of this second seam of 120 cm thickness rests a roof-shale, deposited as mud in a shallow lake, the formation of which was responsible for the cessation of peat deposition. This fourth flora represents the plants growing around the lake on clastic substrate. Four different environments followed each other in this locality over a geologically short time span and each time conditions prevailed to preserve plant macrofossils. Three of these floras represent peat-forming plant communities of essentially the same time interval. This demonstrates the great variability of vegetation and landscapes in the tropical Cathaysian realm of the Late Paleozoic. (author)

  12. The armoured dissorophid Cacops from the Early Permian of Oklahoma and the exploitation of the terrestrial realm by amphibians

    Reisz, Robert R.; Schoch, Rainer R.; Anderson, Jason S.


    Cacops, one of the most distinctive Paleozoic amphibians, is part of a clade of dissorophoid temnospondyls that diversified in the equatorial region of Pangea during the Late Carboniferous and Early Permian, persisting into the Late Permian in Central Russia and China. Dissorophids were a successful group of fully terrestrial, often spectacularly armoured predators, the only amphibians apparently able to coexist with amniotes when the latter started to dominate terrestrial ecosystems. In this paper, we describe excellent new skulls from the Early Permian of Oklahoma attributed to Cacops, Cacops morrisi sp. nov. and provide for the first time detailed information about this iconic dissorophid. These specimens show anatomical and ontogenetic features that will impact on future studies on the evolution of terrestriality in tetrapods. For example, the large, posteriorly closed tympanic embayment has fine striations on an otherwise smooth surface, documenting the oldest known clear evidence for the presence of a tympanic membrane in the fossil record, a structure that is used for hearing airborne sound in extant tetrapods. The skull of C. morrisi also has several features associated with predatory behaviour, indicating that this dissorophid may have been one of the top terrestrial predators of its time.

  13. [Insects at the borderline between the Permian and the early triassic (Urzhum - Olenek age) and the problem of Permian-Triassic biodiversity crisis].

    Rasnitsyn, A P; Aristov, D S; Rasnitsyn, D A


    Distribution of 115 insect families is considered in 15 local assemblages of European Russia, Siberia, Australia and South Africa. The assemblage ages embrace the Urzhum stage of the Middle Permian, the Late Permian, and the transitional Permian-Triassic interval. The assemblages are ordered statistically using two criteria. Ordination after the appearance of a fauna, that is, relation of the number of younger vs. older families, is found to be generally consistent with the stratigraphic data. The method of minimizing the gaps (ghost ranges) in distribution of the families is useful in interpreting the results. Urzhum time is characterized by the balance of emergence and extinction of families (counted as their first and latest appearances, respectively). In Severodvinsk and particularly in Vyatka time, the number of first appearances was decreasing resulted in prevailing extinction. In the transitional Permian-Triassic interval, the emergence of new families accelerated. Initially, the appearance of assemblages was typically Paleozoic (with older families prevailed). It changed gradually, so as by the end of Vyatka time it turned to be quite post-Paleozoic. Diversity was the highest in Severodvinsk time, and it halved at Vyatka time and at the transition interval. However, if we consider transitional families (those not found on a particular interval, but known before and after), the extinction rate reduces to one-third. And when normalized after the material volume, the diversity drop decreases up to a quarter. There was no mass extinction found at the end of the Permian, and the less so at the Permian-Triassic boundary and during the Lower Triassic. Structure of the Permian-Triassic diversity crisis is similar to that of the Cretaceous crisis in many respects. Since the Middle Triassic and up to now, the biodiversity kept increasing quickly and continuously. This implies that the Permian-Triassic crisis resulted in profound modification of the biosphere

  14. Astronomical tuning of the end-Permian extinction and the Early Triassic Epoch of South China and Germany

    Li, Mingsong; Ogg, James; Zhang, Yang; Huang, Chunju; Hinnov, Linda; Chen, Zhong-Qiang; Zou, Zhuoyan


    The timing of the end-Permian mass extinction and subsequent prolonged recovery during the Early Triassic Epoch can be established from astronomically controlled climate cycles recorded in continuous marine sedimentary sections. Astronomical-cycle tuning of spectral gamma-ray logs from biostratigraphically-constrained cyclic stratigraphy through marine sections at Meishan, Chaohu, Daxiakou and Guandao in South China yields an integrated time scale for the Early Triassic, which is consistent with scaling of magnetostratigraphy from climatic cycles in continental deposits of the Germanic Basin. The main marine mass extinction interval at Meishan is constrained to less than 40% of a 100-kyr (kilo-year) cycle (i.e., Permian extinction. The durations of the Griesbachian, Dienerian, Smithian and Spathian substages, including the uncertainty in placement of widely used conodont biostratigraphic datums for their boundaries, are 1.4 ± 0.1, 0.6 ± 0.1, 1.7 ± 0.1 and 1.4 ± 0.1 myr, implying a total span for the Early Triassic of 5.1 ± 0.1 myr. Therefore, relative to an assigned 251.902 ± 0.024 Ma for the Permian-Triassic boundary from the Meishan GSSP, the ages for these substage boundaries are 250.5 ± 0.1 Ma for base Dienerian, 249.9 ± 0.1 Ma for base Smithian (base of Olenekian stage), 248.2 ± 0.1 Ma for base Spathian, and 246.8 ± 0.1 Ma for the base of the Anisian Stage. This astronomical-calibrated timescale provides rates for the recurrent carbon isotope excursions and for trends in sedimentation accumulation through the Early Triassic of studied sections in South China.

  15. Large Early Permian eruptive complexes in northern Saxony, Germany: Volcanic facies analysis and geochemical characterization

    Hübner, Marcel; Breitkreuz, Christoph; Repstock, Alexander; Heuer, Franziska


    In the course of formation of extensional basins during the Early Permian a widespread volcanic activity led to the deposition of volcanic and volcanosedimentary units in Saxony (Walter 2006, Hoffmann et al. 2013). Situated east of Leipzig, the North Saxonian Volcanic Complex (NSVC) hosts two large caldera complexes, the Rochlitz and Wurzen Volcanic Systems, with diameters of 90 and 52 km, respectively. Volume estimates (> 1000 km3) qualify these as supereruptions according to Mason et al. (2004). In addition to the large caldera systems, the NSVC hosts several small pyroclastic flow deposits ranging from crystal-poor (e.g. Cannewitz and vitrophyric Ebersbach ignimbrites) to crystal-rich units (Wermsdorf and Dornreichenbach ignimbrites). Additionally rhyolitic lava and subvolcanic units are present. The Chemnitz basin (Schneider et al. 2012), located to the south of the NSVC, harbours caldera-outflow facies deposits of the Rochlitz eruption (Fischer 1991), i.e. the partially vitrophyric Planitz ignimbrite. The Rochlitz and Wurzen caldera-fill ignimbrites exhibit relatively high crystal contents with maxima up to 52 and 58 vol.-%, for corresponding 66 and 68 wt.-% SiO2. This is comparable with the 'monotonous intermediates' (Hildreth 1981) in the Cenozoic western USA investigated by Huber et al. (2012). In contrast, the Planitz ignimbrite in the Chemnitz basin reveals predominantly crystal-poor pyroclastics ( 70 analyses), and mineral geochemistry to reconstruct the eruption history and magma genesis of this large Late Paleozoic magmatic complex in Central Europe. Volcanic textures and geochemical trends indicate magma mingling and mixing to have been important during the formation of the Wurzen caldera system. Geothermometric and -barometric calculations based on composition of pyroxene and feldspar suggest deeply seated crustal magma chambers for the NSVC and the Planitz ignimbrite. Fischer, F. (1991): Das Rotliegende des ostthüringisch-westsächsischen Raumes

  16. New Species of the Isolated Psaroniaceous Rachis from the Early Permian in China


    A new kind of marattlalean raches are reported from the coal balls in Coal Seam No.7 in the upper part of the Taiyuan Formation (early Early Permian) from Taiyuan, Shanxi, China and are assigned to the genus Stipitopteris Grand'Eury(Psaroniaceae). The present specimens are different from all six reported species of the genus, and are therefore proposed as a new species: Stipitopteris shanxiensis. The raches of the new species are generally dorsi-ventrally flattened. The main raches usually exhibit scales of different forms on their surface. Beneath the epidermis is a zone of parenchymatous cells, some of which contain tannin-like contents. Inside this is a zone of small sclerenchymatous cells. Inward are the ground tissue and vascular bundles. The vascular bundles are continuous and are in two circles: the outer circle assumes a transversely elliptical shape with the gap and pinna trace, and the inner circle assumes a shallow C-shape with inrolled ends. The ground tissue located at the inner side of the vascular bundle is composed of thicker-walled parenchymatous cells. The cells of the ground tissue are vertically elongated in longitudinal sections. Subordered raches are smaller and have simpler structures than the main raches. The parenchyma zone beneath the epidermis is thinner, usually one to two cells wide and the sclerenchyma zone is usually absent. The scales are poorly developed and there is only one C-shaped vascular bundle. The new species is comparable to the crosiers of Psaronlaceae of the EurameHcan Flora in some aspects,for example, it has a dorsi-ventrally flattened rachis and scales on the surface of the rachis. However, the other features and the preservative conditions of the present specimens indicate that they are not crosiers, but fully developed or mature raches. The new species is the first well-studied anatomically-preserved rachis of Psaroniaceae from the Cathaysian Flora and bears significance not only in understanding the anatomy and

  17. Early Triassic alternative ecological states driven by anoxia, hyperthermals, and erosional pulses following the end-Permian mass extinction

    Pietsch, C.; Petsios, E.; Bottjer, D. J.


    The end-Permian mass extinction, 252 million years ago, was the most devastating loss of biodiversity in Earth's history. Massive volcanic eruptions of the Siberian Traps and the concurrent burning of coal, carbonate, and evaporite deposits emplaced greenhouse and toxic gasses. Hyperthermal events of the surface ocean, up to 40°C, led to reduced gradient-driven ocean circulation which yielded extensive equatorial oxygen minimum zones. Today, anthropogenic greenhouse gas production is outpacing carbon input modeled for the end-Permian mass extinction, which suggests that modern ecosystems may yet experience a severe biotic crisis. The Early Triassic records the 5 million year aftermath of the end-Permian mass extinction and is often perceived as an interval of delayed recovery. We combined a new, high resolution carbon isotope record, sedimentological analysis, and paleoecological collections from the Italian Werfen Formation to fully integrate paleoenvironmental change with the benthic ecological response. We find that the marine ecosystem experienced additional community restructuring events due to subsequent hyperthermal events and pulses of erosion. The benthic microfauna and macrofauna both contributed to disaster communities that initially rebounded in the earliest Triassic. 'Disaster fauna' including microbialites, microconchids, foraminifera, and "flat clams" took advantage of anoxic conditions in the first ~500,000 years, dominating the benthic fauna. Later, in the re-oxygenated water column, opportunistic disaster groups were supplanted by a more diverse, mollusc-dominated benthic fauna and a complex ichnofauna. An extreme temperature run-up beginning in the Late Dienerian led to an additional hyperthermal event in the Late-Smithian which co-occurred with increased humidity and terrestrial run-off. Massive siliciclastic deposits replaced carbonate deposition which corresponds to the infaunalization of the benthic fauna. The disaster taxa dominated

  18. An Early Permian plant assemblage from the Taiyuan Formation of northern China with compression/impression and permineralized preservation.

    Hilton, J; Wang, S -J.; Galtier, J; Li, C -S.


    A small but diverse fossil flora is described from the Early Permian Taiyuan Formation occurring at the Yangshuling mine in Pingquan district of Hebei Province, northern China. Fossils occur as compression/impressions within mudrocks and fine-grained sandstones and also as carbonate permineralizations within volcaniclastic tuffs. All are fragmentary and contain lycopsids, sphenopsids, ferns and seed plants, and include several new species. In the compression assemblage sphenopsid and pteridosperm foliage accounts for the majority of the fossils recognised with only a few other kinds of plant organs present. In contrast, the permineralized assemblage is dominated by cordaitaleans with a composition similar to that occurring in coal-ball assemblages elsewhere in the Taiyuan Formation. From the taxonomic synthesis presented it is apparent that the Yangshuling permineralized assemblage contains many of the plant taxa diagnostic of the northern realm of the Early Permian Cathaysian flora, and preserves a representative sample of the wetland coal-swamp vegetation of this time. The permineralized assemblage at Yangshuling represents the first example of anatomically preserved plants from volcaniclastic lithologies from the Palaeozoic of China, raising the possibility of similarly preserved plant-fossil assemblages elsewhere in the Cathaysian realm.

  19. Taphonomy of Condensed Shell Storm beds of the Teresina Formation in Rio Preto (State of Paraná, Middle Permian, Paraná Basin: Paleoenvironmental Implications

    Jacqueline Peixoto Neves


    Full Text Available The deposits of the Permian Teresina Formation are mainly characterized by fi ne-grained siliciclastic rocks and centimetricintercalations of tempestites (bioclastic sandstones and coquinas. Despite the relevance of the bivalve-rich carbonate bedsof the Teresina Formation to paleoenvironmental studies, their taphonomy is still poorly studied. The fossil concentrationstudied in this work was found in a quarry in the city of Irati, Rio Preto district, Paraná State. The fossil concentration is locatedin the middle/upper portion of the unit, far from the top. The studied bed is a bioclastic, intraclastic, peloidal, grainstone/packstone, with abundant bivalve shell fragments, pelitic and micritic intraclasts, peloids, rare ooids and oncoids, as well aspermineralized of Lycophyta microphylles and fi sh scales. The grains of this carbonate concentration show: high degree oftime-averaging, variable degree of packing (dense to disperse, no sorting and chaotic orientation. Notably, the concentrationincludes a mixture of elements which are indicative of: a restrictive, low energy, carbonate environment (peloids, ooidsand oncoids; b subaerial environment surrounding the main body of water (Lycophyta microphylles and c quiet-waterenvironment punctuated by storm events, where the suspension-feeding bivalves thrived. At least four depositional eventscaused by storm fl ows were recorded. The amalgamated nature of the bed is a result of storm events in an intracratonic basinwith very low seafl oor slope and low rates of sedimentation and subsidence.

  20. Redox chemistry changes in the Panthalassic Ocean linked to the end-Permian mass extinction and delayed Early Triassic biotic recovery.

    Zhang, Guijie; Zhang, Xiaolin; Hu, Dongping; Li, Dandan; Algeo, Thomas J; Farquhar, James; Henderson, Charles M; Qin, Liping; Shen, Megan; Shen, Danielle; Schoepfer, Shane D; Chen, Kefan; Shen, Yanan


    The end-Permian mass extinction represents the most severe biotic crisis for the last 540 million years, and the marine ecosystem recovery from this extinction was protracted, spanning the entirety of the Early Triassic and possibly longer. Numerous studies from the low-latitude Paleotethys and high-latitude Boreal oceans have examined the possible link between ocean chemistry changes and the end-Permian mass extinction. However, redox chemistry changes in the Panthalassic Ocean, comprising ∼85-90% of the global ocean area, remain under debate. Here, we report multiple S-isotopic data of pyrite from Upper Permian-Lower Triassic deep-sea sediments of the Panthalassic Ocean, now present in outcrops of western Canada and Japan. We find a sulfur isotope signal of negative Δ(33)S with either positive δ(34)S or negative δ(34)S that implies mixing of sulfide sulfur with different δ(34)S before, during, and after the end-Permian mass extinction. The precise coincidence of the negative Δ(33)S anomaly with the extinction horizon in western Canada suggests that shoaling of H2S-rich waters may have driven the end-Permian mass extinction. Our data also imply episodic euxinia and oscillations between sulfidic and oxic conditions during the earliest Triassic, providing evidence of a causal link between incursion of sulfidic waters and the delayed recovery of the marine ecosystem.

  1. Evolution of strontium isotopic composition of sea-water from Late Permian to Early Triassic based on study of marine carbonates,Zhongliang Mountain,Chongqing,China


    Collected from a Late Permian to Early Triassic sedimentary section in the Zhongliang Mountain of Chongqing, Southwest China, sixty marine carbonate samples were measured for the 87Sr/86Sr ratios, and corresponding evolution curve was constructed. The concentrations of SiO2, CaO, MgO, Mn and Sr are used to evaluate reservation of strontium isotopic composition for original seawater and the credi-bility of the dissolution method for sample preparation. The results show that most of the samples (except seven samples with the Mn/Sr ratios higher than 2) contain the original geochemistry signa-tures of ancient seawater. Compared to the published 87Sr/86Sr ratios from the Late Permian to Early Triassic, our database reported here is the largest and the curve constructed is the most complete. The strontium isotopic curve from Late Permian to Early Triassic is consistent globally and exhibits a gen-eral trend of steady increase during this period. The minimum of 87Sr/86Sr ratios (0.707011) occurs in the Late Permian (30 m in thickness below the Permian-Triassic boundary), and the maximum (0.708281), near the Early-Middle Triassic boundary. The lack of land plants and the rapid continental weathering result in the increase of 87Sr/86Sr ratios during the interval. The Permian-Triassic boundary in Zhongli-ang Mountain Section has been accepted internationally. The 87Sr/86Sr ratios of six samples near the boundary vary from 0.70714 to 0.70715 with an average of 0.70714, which is consistent with the value of 0.70715 (samples are from articulate brachiopod shells) from Korte et al. published in 2006 (within the error range in experiment). Accordingly, the strontium isotope composition in the Permian-Triassic boundary in this paper is of global significance. It can be confirmed that the 87Sr/86Sr ratios of the sea-water in the Permian-Triassic transition are in the range of 0.70714―0.70715.

  2. The Montalet granite, Montagne Noire, France: An Early Permian syn-extensional pluton as evidenced by new U-Th-Pb data on zircon and monazite

    Poilvet, Jean-Charles; Poujol, Marc; Pitra, Pavel; Van Den Driessche, Jean; Paquette, Jean-Louis


    Dating the magmatism in the Montagne Noire gneiss dome in the southern French Massif Central is a key point for understanding the Late Palaeozoic evolution of this part of the Variscan belt, which is characterised by compressive tectonics during the Carboniferous and extensional tectonics during Stephanian-Permian times. The Montalet granite crops out in the north-western part of the dome and was first considered as an early syntectonic intrusion related to compressive deformation. More recently, it has been dated at 327 Ma and considered as contemporaneous with the diapiric ascent of the Montagne Noire gneiss dome before the Stephanian-Permian extension. We show that in fact, this pluton was emplaced 294 ± 1 Ma ago and is therefore contemporaneous with the Stephanian-Permian extension. This age is consistent with the interpretation of the Montagne Noire Massif as an extensional gneiss dome.

  3. Sedimentary evolution of Rio do Rastro formation (permian-triassic of the Parana Basin) at central south portion of Santa Catarina State, Brazil; Evolucao sedimentar da Formacao Rio do Rastro (Permo-Triassico da Bacia do Parana) na porcao centro sul do estado de Santa Catarina, Brasil

    Warren, Lucas Verissimo; Roldan, Luis Fernando; Steiner, Samar dos Santos; Chamani, Marlei Antonio Carrari [Universidade de Sao Paulo (USP), SP (Brazil). Inst. de Geociencias. Programa de Pos Graduacao em Geoquimica e Geotectonica]. E-mail:; Almeida, Renato Paes de; Hachiro, Jorge; Machado, Romulo [Universidade de Sao Paulo (USP), SP (Brazil). Inst. de Geociencias. Dept. de Geologia Sedimentar e Ambiental


    Between the Late Carboniferous and the Early Triassic, the south portion of the Gondwana Supercontinent witnessed the development of intracratonic basins related with the coeval Sanrafaelic Orogeny. The continuous subsidence and consequent generation accommodation space resulted in the formation of a large confined water body and the accumulation of a transgressive regressive sequence between the Late Permian and the Early Triassic. The progradational nature of the upper portion of this sequence, represented by de Rio do Rasto e Piramboia formations, culminated with the complete filling of the water body. In the south-eastern portion of Santa Catarina State (Southern Brazil), the Rio do Rasto Formation overlies the Teresina formation and is overlain by the Piramboia formation, both contacts being characterized by lithological transition. The lower portion of the Rio do Rasto Formation is characterized by architectural elements deposited in offshore environments subject to storm action. A marked change of the color of the pelitic facies, from gray to red and purple occurs at the top of the unit. At this stratigraphic level, there is also a predominance of deltaic and eolian architectural elements. The intercalation of near shore and offshore architectural elements can be explained by the strong auto cyclic character expected in deltaic depositional systems and by the variation of relative rates of base-level rise. The occurrence of eolian architectural elements interbedded with subaquatically deposited sediments on the second third of the unit can be considered an evidence of continentalization to the top, materialized as the transition to the Piramboia desert system. (author)

  4. Architecture and evolution of an Early Permian carbonate complex on a tectonically active island in east-central California

    Stevens, Calvin H.; Magginetti, Robert T.; Stone, Paul


    The newly named Upland Valley Limestone represents a carbonate complex that developed on and adjacent to a tectonically active island in east-central California during a brief interval of Early Permian (late Artinskian) time. This lithologically unique, relatively thin limestone unit lies within a thick sequence of predominantly siliciclastic rocks and is characterized by its high concentration of crinoidal debris, pronounced lateral changes in thickness and lithofacies, and a largely endemic fusulinid fauna. Most outcrops represent a carbonate platform and debris derived from it and shed downslope, but another group of outcrops represents one or possibly more isolated carbonate buildups that developed offshore from the platform. Tectonic activity in the area occurred before, probably during, and after deposition of this short-lived carbonate complex.

  5. Early Permian intrusions in the Paleozoic sediments of the Eastern North Sea area

    Clausen, O.R.; Andresen, Katrine Juul; Rasmussen, Jens Andreas

    of hydrocarbons within the Paleozoic sediments, the Northern Permian Basin has gained much less interest outside the hydrocarbon producing Mesozoic graben systems. This is mainly due to an apparent lack of potential source rocks. A major E-W striking northward dipping fault system characterizes the study area......This study presents the geometry of Paleozoic intrusions in the Skagerrak area located at the northern flank of the Ringkøbing-Fyn High and suggests factors controlling the formation of the intrusions. The intrusions have here been mapped in detail using 3D seismic data. The study area is located...... is a potential source rock, has local depositional maxima associated to the basement faults. Salt structures which have been periodically active during the post Paleozoic dominate the northern part of the study area. The Paleozoic intrusions observed in the hanging-wall segment of the E-W striking fault system...

  6. Regional implications of new chronostratigraphic and paleogeographic data from the Early Permian Darwin Basin, east-central California

    Stevens, Calvin H.; Stone, Paul; Magginetti, Robert T.


    The Darwin Basin developed in response to episodic subsidence of the western margin of the Cordilleran continental shelf from Late Pennsylvanian (Gzhelian) to Early Permian (late Artinskian) time. Subsidence of the basin was initiated in response to continental truncation farther to the west and was later augmented by thrust emplacement of the Last Chance allochthon. This deep-water basin was filled by voluminous fine-grained siliciclastic turbidites and coarse-grained limestone-gravity-flow deposits. Most of this sediment was derived from the Bird Spring carbonate shelf and cratonal platform to the northeast or east, but some came from an offshore tectonic ridge (Conglomerate Mesa Uplift) to the west that formed at the toe of the Last Chance allochthon. At one point in the late Artinskian the influx of extrabasinal sediment was temporarily cut off, resulting in deposition of a unique black limestone that allows precise correlation throughout the basin. Deep-water sedimentation in the Darwin Basin ended by Kungurian time when complex shallow-water to continental sedimentary facies spread across the region. Major expansion of the Darwin Basin occurred soon after the middle Sakmarian emplacement of the Last Chance allochthon. This tectonic event was approximately coeval with deformation in northeastern Nevada that formed the deep-water Dry Mountain Trough. We herein interpret the two basins to have been structurally continuous. Deposition of the unique black limestone is interpreted to mark a eustatic sea level rise that also can be recognized in Lower Permian sections in east-central Nevada and central Arizona.

  7. Stratigraphy and paleogeographic significance of a Late Pennsylvanian to Early Permian channeled slope sequence in the Darwin Basin, southern Darwin Hills, east-central California

    Stevens, Calvin H.; Stone, Paul; Magginetti, Robert T.; Ritter, Scott M.


    The complex stratigraphy of late Paleozoic rocks in the southern Darwin Hills consists of regionally extensive Mississippian and Early to Middle Pennsylvanian rocks overlain by latest Pennsylvanian to Early Permian rocks, herein called the Darwin Hills sequence. Deposition of this latter sequence marked the beginning of the Darwin Basin. In Mississippian time, a carbonate platform prograded westward over slightly older slope deposits. In the Late Mississippian this platform was exposed to erosion and siliciclastic sediments were deposited. In Early to Middle Pennsylvanian time the area subsided, forming a west-facing ramp that was subjected to deformation and erosion in Middle or early Late Pennsylvanian time. Later this area was tilted westward and deep-water sediments were deposited on this slope. In latest Pennsylvanian to earliest Permian time, a major channel was cut through the older Pennsylvanian rocks and into the Upper Mississippian strata. This channel was gradually filled with increasingly finer grained, deep-water sediment as the area evolved into a basin floor by Early Permian (Sakmarian) time. Expansion of the Darwin Basin in Artinskian time led to a second phase of deposition represented by strata of the regionally extensive Darwin Canyon Formation. The geology in this small area thus documents tectonic events occurring during the early development of the Darwin Basin.

  8. Late Permian to Early Oligocene granitic magmatism of the Phan Si Pan uplift area, NW Vietnam: their relationship to Phanerozoic crustal evolution of Southwest China

    Pham, T. T.; Shellnutt, G.


    The Phan Si Pan uplift area of NW Vietnam is a part of the Archean to Paleoproterozoic Yangtze Block, Southwest China. This area is of particular interest because it experienced a number of Phanerozoic crustal building events including the Emeishan Large Igneous Province, the India-Eurasia collision and Ailaoshan - Red River Fault displacement. In the Phan Si Pan uplift area, there are at least three different geochronological complexes, including: (1) Late Permian, (2) Eocene and (3) Early Oligocene. (1) The Late Permian silicic rocks are alkali ferroan A1-type granitic rocks with U/Pb ages of 251 ± 3 to 254 ± 3 Ma. The Late Permian silicic rocks of Phan Si Pan uplift area intrude the upper to middle crust and are considered to be part of the ELIP that was displaced during the India-Eurasian collision along the Ailaoshan-Red River Fault shear zone and adjacent structures (i.e. Song Da zone). Previous studies suggest the Late Permian granitic rocks were derived by fractional crystallization of high - Ti basaltic magma. (2) The Eocene rocks are alkali ferroan A1-type granites (U/Pb ages 49 ± 0.9 Ma) and are spatially associated with the Late Permian granitic rocks. The trace element ratios of this granite are similar to the Late Permian rocks (Th/Nb=0.2, Th/Ta = 2.5, Nb/U = 24, Nb/La =1.2, Sr/Y=1). The origin of the Eocene granite is uncertain but it is possible that it formed by fractional crystallization of a mafic magma during a period of extension within the Yangtze Block around the time of the India-Eurasia collision. (3) The Early Oligocene granite is characterized as a peraluminous within-plate granite with U/Pb ages of 31.3 ± 0.4 to 34 ± 1 Ma. The Early Oligocene granite has trace element ratios (Th/Nb = 2.1, Th/Ta = 22.6, Nb/U = 4.4, Nb/La = 0.4, Sr/Y = 60.4) similar to crust melts. The high Sr/Y ratio (Sr/Y = 20 - 205) indicates a lower crust source that was garnet-bearing. The Phan Si Pan uplift was neither a subduction zone nor an arc environment

  9. Stratigraphy sequence analysis application for multi scalar characterization of paralic reservoirs - an example in the Guata group (E O-Permian) of the Parana Basin, in the state of Rio Grande do Sul, Brazil; Aplicacao da estratigrafia de sequencias para caracterizacao multiescalar de reservatorios paralicos - um exemplo no Grupo Guata (EO-Permiano) da Bacia do Parana, no Rio Grande do Sul, Brasil

    Kuechle, Juliano [Rio Grande do Sul Univ., Porto Alegre, RS (Brazil). Inst. de Geociencias. Curso de Pos-graduacao em Geociencias]. E-mail:; Holz, Michael [Rio Grande do Sul Univ., Porto Alegre, RS (Brazil). Inst. de Geociencias]. E-mail:


    Detailed stratigraphic analysis of the Early Permian Rio Bonito and Palermo Formations of the Parana Basin in the region of Sao Gabriel County has been used for a study on reservoir characterization of paralic sandstones. Two main depositional systems were recognized, a fluvial-dominated delta system and a wave-dominated barrier island system. The succession is divided in two third-order depositional sequences, enclosing thirteen fourth-order para sequences. This high-resolution stratigraphic framework was the base for a multi scale approach on reservoir characterization of the sandstone bodies. Reservoir heterogeneities are discussed, staring from the scale of depositional sequence (heterogeneity level 1), passing down to heterogeneity at the scale of systems tracts (heterogeneity level 2) and finalizing with an approach at para sequence scale (heterogeneity level 3). Main control on heterogeneity at the first level is base-level variation as generating mechanism for un conformities at the sediment type and rate. At the second level, the reservoir heterogeneity is controlled by the lateral and vertical variations in thickness of particular systems within the different systems tracts, and at the third level, the heterogeneity occurs controlling reservoir continuity and connectivities between reservoirs. The study supplies a model which is useful as a predictive tool for similar geologic settings in producing oil fields. (author)

  10. Paleomagnetic data support Early Permian age for the Abor Volcanics in the lower Siang Valley, NE India: Significance for Gondwana-related break-up models

    Ali, Jason R.; Aitchison, Jonathan C.; Chik, Sam Y. S.; Baxter, Alan T.; Bryan, Scott E.


    Confusion exists as to the age of the Abor Volcanics of NE India. Some consider the unit to have been emplaced in the Early Permian, others the Early Eocene, a difference of ˜230 million years. The divergence in opinion is significant because fundamentally different models explaining the geotectonic evolution of India depend on the age designation of the unit. Paleomagnetic data reported here from several exposures in the type locality of the formation in the lower Siang Valley indicate that steep dipping primary magnetizations (mean = 72.7 ± 6.2°, equating to a paleo-latitude of 58.1°) are recorded in the formation. These are only consistent with the unit being of Permian age, possibly Artinskian based on a magnetostratigraphic argument. Plate tectonic models for this time consistently show the NE corner of the sub-continent >50°S; in the Early Eocene it was just north of the equator, which would have resulted in the unit recording shallow directions. The mean declination is counter-clockwise rotated by ˜94°, around half of which can be related to the motion of the Indian block; the remainder is likely due local Himalayan-age thrusting in the Eastern Syntaxis. Several workers have correlated the Abor Volcanics with broadly coeval mafic volcanic suites in Oman, NE Pakistan-NW India and southern Tibet-Nepal, which developed in response to the Cimmerian block peeling-off eastern Gondwana in the Early-Middle Permian, but we believe there are problems with this model. Instead, we suggest that the Abor basalts relate to India-Antarctica/India-Australia extension that was happening at about the same time. Such an explanation best accommodates the relevant stratigraphical and structural data (present-day position within the Himalayan thrust stack), as well as the plate tectonic model for Permian eastern Gondwana.

  11. Late Carboniferous to early Permian sedimentary–tectonic evolution of the north of Alxa, Inner Mongolia, China: Evidence from the Amushan Formation

    Haiquan Yin


    Full Text Available The late Paleozoic evolution of the Wulijishanhen (WSH-Shangdan (SD area near to the Chaganchulu Ophiolite belt is reinterpreted. Analysis of the upper Carboniferous to lower Permian sedimentary sequence, biological associations, detrital materials, sandstone geochemistry and volcanic rocks indicates that the SD area was an epicontinental sea and rift during the late Paleozoic rather than a large-scale ocean undergoing spreading and closure. This study reveals that the actual evolution of the study area is from the late Carboniferous to the early Permian. The fusulinids Triticites sp. and Pseudoschwagerina sp. in the limestones demonstrate that the Amushan Formation develops during the late Carboniferous to the early Permian. The limestones at the base of the SD section indicate that it is a stable carbonate platform environment, the volcanic rocks in the middle of the sequence support a rift tectonic background, and the overlying conglomerates and sandstones are characteristic of an epicontinental sea or marine molasse setting. The rift volcanism made the differences in the fossil content of the SD and WSH sections and led to two sections expose different levels within the Amushan Formation and different process of tectonic evolution. Moreover, the geochemical characteristics and detrital materials of the sandstones show that the provenance and formation of the sandstones were related to the setting of active continental margin. The quartz-feldspar-lithic fragments distribution diagram indicates that the material source for the sandstones was a recycled orogenic belt. Thus, the source area of the sandstones may have been an active continental margin before the late Carboniferous–early Permian. The characteristics of the regional tectonic evolution of the area indicate that the region may form a small part of the Gobi–Tianshan rift of southern Mongolia.

  12. Record of Permian-Early Triassic continental arc magmatism in the western margin of the Jiamusi Block, NE China: petrogenesis and implications for Paleo-Pacific subduction

    Yang, Hao; Ge, Wenchun; Dong, Yu; Bi, Junhui; Wang, Zhihui; Ji, Zheng; Yang, H.; Ge, W. C.; Dong, Y.; Bi, J. H.; Wang, Z. H.; Ji, Z.


    In this paper, we report zircon U-Pb ages, Hf isotopes and whole-rock geochemical data for the Permian to Early Triassic granitoids from the western margin of the Jiamusi Block (WJB), NE China. The intermediate to felsic (SiO2 = 59.67-74.04 wt%) granitoids belong to calc-alkaline series and are characterized by enrichments in light rare earth elements and large ion lithophile elements with pronounced negative Nb, Ta and Ti anomalies, revealing typical continental magmatic arc geochemical signatures. The zircon U-Pb determinations on the granodiorite, monzogranite, syenogranite and quartz diorite samples yielded ages between ca. 275-245 Ma, which, together with the published coeval intrusive rocks, indicates that Permian to Early Triassic continental arc magmatism occurred extensively in the WJB. The low and mainly negative zircon ɛ Hf(t) values between -7.6 and +1.6 and the zircon Hf model ages of 1.2-1.8 Ga, which are significantly older than their crystallization ages, suggest that they were mainly derived from reworking of ancient crustal materials with a limited input of juvenile components. The geochemical systematics and petrogenetic considerations indicate that the studied granitoids were generated from a zone of melting, assimilation, storage, and homogenization, i.e., a MASHed zone at the base of Paleo- to Mesoproterozoic continental crust, where large portions of igneous rocks and minor clay-poor sediments involved in the source region. In combination with regional geological data, we argue that the Jiamusi Block was unlikely the rifted segment of the Songliao Block and two possible geodynamical models were proposed to interpret the formation of the ca. 275-245 Ma granitoids in the WJB. In the context of Permian global plate reconstruction, we suggest that Paleo-Pacific plate subduction was initiated in the Permian to Early Triassic beneath the Jiamusi Block, and even whole eastern NE China.

  13. Contrasting origin of two A-type rhyolite series from the Early Permian Nomgon bimodal volcanic association (Southern Mongolia)

    Kozlovsky, A. M.; Yarmolyuk, V. V.; Savatenkov, V. M.; Kudryashova, E. A.


    A-type rhyolites of contrasting compositions and eruption characters were revealed among two volcanic series of the Early Permian bimodal association in the Nomgon graben. Rhyolites of the lower volcanic series formed extrusions, lava domes, and tuff horizons. They had low FeOt, Zr, Hf, Nb, Ta, Y, and REE concentrations and also a moderately depleted Nd isotope composition (ɛNd( T) = 6.7-7.1). Their formation was related to anatexis of the juvenile continental crust, triggered by the thermal effect of mafic magmas. Rhyolites of the upper volcanic series formed extensive lava flows and dikes. Their composition was characterized by high FeOt, Zr, Hf, Nb, Ta, Y, and REE concentrations, and also depleted Nd isotope characteristics (ɛNd( T) = 7.7-9.0). These rhyolite melts formed under long-term crystallizational differentiation of basaltoids in the intracrustal magmatic chambers, with limited participation of crustal contamination. The source of magmas for the upper volcanic series was the sublithospheric mantle.

  14. Geological Controls on Mineralogy and Geochemistry of an Early Permian Coal from the Songshao Mine, Yunnan Province, Southwestern China

    Ruixue Wang


    Full Text Available This paper discusses the content, distribution, modes of occurrence, and enrichment mechanism of mineral matter and trace elements of an Early Permian coal from Songshao (Yunnan Province, China by means of coal-petrological, mineralogical, and geochemical techniques. The results show that the Songshao coal is characterized by high total and organic sulfur contents (3.61% and 3.87%, respectively. Lithium (170.39 μg/g and Zr (184.55 μg/g are significantly enriched in the Songshao coal, and, to a lesser extent, elements such as Hg, La, Ce, Nd, Th, Sr, Nb, Sn, Hf, V, and Cr are also enriched. In addition to Hg and Se that are enriched in the roof and floor strata of the coal seam, Li, La, Ce, Pr, Nd, Sm, Gd, Y, Cd, and Sb are slightly enriched in these host rocks. Compared to the upper continental crust, rare earth elements and yttrium in the host rocks and coal samples are characterized by a light-REE enrichment type and have negative Eu, positive Ce and Gd anomalies. Major minerals in the samples of coal, roof, and floor are boehmite, clay minerals (kaolinite, illite, and mixed layer illite-smectite, pyrite, and anatase. Geochemical and mineralogical anomalies of the Songshao coal are attributed to hydrothermal fluids, seawater, and sediment-source rocks.

  15. Late Carboniferous to early Permian sedimentaryetectonic evolution of the north of Alxa, Inner Mongolia, China:Evidence from the Amushan Formation

    Haiquan Yin; Hongrui Zhou; Weijie Zhang; Xiaoming Zheng; Shengyu Wang


    The late Paleozoic evolution of the Wulijishanhen (WSH)-Shangdan (SD) area near to the Chaganchulu Ophiolite belt is reinterpreted. Analysis of the upper Carboniferous to lower Permian sedimentary sequence, biological associations, detrital materials, sandstone geochemistry and volcanic rocks indicates that the SD area was an epicontinental sea and rift during the late Paleozoic rather than a large-scale ocean undergoing spreading and closure. This study reveals that the actual evolution of the study area is from the late Carboniferous to the early Permian. The fusulinids Triticites sp. and Pseudoschwagerina sp. in the limestones demonstrate that the Amushan Formation develops during the late Carboniferous to the early Permian. The limestones at the base of the SD section indicate that it is a stable carbonate platform environment, the volcanic rocks in the middle of the sequence support a rift tectonic back-ground, and the overlying conglomerates and sandstones are characteristic of an epicontinental sea or marine molasse setting. The rift volcanism made the differences in the fossil content of the SD and WSH sections and led to two sections expose different levels within the Amushan Formation and different process of tectonic evolution. Moreover, the geochemical characteristics and detrital materials of the sandstones show that the provenance and formation of the sandstones were related to the setting of active continental margin. The quartz-feldspar-lithic fragments distribution diagram indicates that the material source for the sandstones was a recycled orogenic belt. Thus, the source area of the sandstones may have been an active continental margin before the late Carboniferouseearly Permian. The charac-teristics of the regional tectonic evolution of the area indicate that the region may form a small part of the GobieTianshan rift of southern Mongolia.

  16. A large aberrant stem ichthyosauriform indicating early rise and demise of ichthyosauromorphs in the wake of the end-Permian extinction.

    Jiang, Da-Yong; Motani, Ryosuke; Huang, Jian-Dong; Tintori, Andrea; Hu, Yuan-Chao; Rieppel, Olivier; Fraser, Nicholas C; Ji, Cheng; Kelley, Neil P; Fu, Wan-Lu; Zhang, Rong


    Contrary to the fast radiation of most metazoans after the end-Permian mass extinction, it is believed that early marine reptiles evolved slowly during the same time interval. However, emerging discoveries of Early Triassic marine reptiles are questioning this traditional view. Here we present an aberrant basal ichthyosauriform with a hitherto unknown body design that suggests a fast radiation of early marine reptiles. The new species is larger than coeval marine reptiles and has an extremely small head and a long tail without a fluke. Its heavily-built body bears flattened and overlapping gastral elements reminiscent of hupehsuchians. A phylogenetic analysis places the new species at the base of ichthyosauriforms, as the sister taxon of Cartorhynchus with which it shares a short snout with rostrally extended nasals. It now appears that ichthyosauriforms evolved rapidly within the first one million years of their evolution, in the Spathian (Early Triassic), and their true diversity has yet to be fully uncovered. Early ichthyosauromorphs quickly became extinct near the Early-Middle Triassic boundary, during the last large environmental perturbation after the end-Permian extinction involving redox fluctuations, sea level changes and volcanism. Marine reptile faunas shifted from ichthyosauromorph-dominated to sauropterygian-dominated composition after the perturbation.




    Full Text Available The transitional faunas of the Permian Huqf succession of Oman make it one of the key-sections for the intercalibration of Early to Middle Permian biostratigraphical scales. The abundance of fossils improved the knowledge of some marine faunas which populated the North-Eastern Gondwanan fringe during times of climatic changes in the Permian. A Sterlitamakian (upper Sakmarian, Lower Permian bivalve fauna from the Saiwan Formation in the Huqf area, informally named "Dickinsartella Fauna", is described in the present paper. The specimens examined were collected from the "Pachycyrtella Bed" (Auctorum, the basal bed of the Formation in its type locality. The Dickinsartella Fauna can be identified for the presence of the new genus Dickinsartella, which dominates the bivalve thanatocoenosis with D. pistacina sp. n. (type species. The bivalve fauna from the Pachycyrtella Bed includes the new species Stutchburia sangallii and Promytilus  mazzolenii, and also Astartella obliqua Dickins, 1963, Nuculopsis cf. bangarraensis Dickins, 1963, ?Oriocrassatella sp., and indeterminable aviculopectinids. This fauna shows a low taxonomic diversity. Nevertheless, some species are represented by a high number of generally well-preserved specimens, i.e. some specimens of S. sangallii sp. n. and A. obliqua show part of the ligament.  The good preservation of the shells permitted the microstructural analysis of D. pistacina sp. n. and S. sangallii sp. n. The microstructure of S. sangallii sp. n. supports the close phylogenetical link between modiomorphids and crassatelloids recognized by some previous authors.The new genus Dickinsartella includes the more recent species belonging to the important Paleozoic Order Cyrtodontida Scarlato & Starobogatov, 1971. The discovery of Dickinsartella gen. n. and other taxa of the Pachycyrtella Bed, present also in the Sakmarian levels of the Carnarvon and Perth Basins in Western Australia,  indicates a wider distribution of the

  18. Support for an “A-type” Pangea reconstruction from high-fidelity Late Permian and Early to Middle Triassic paleomagnetic data from Argentina

    Domeier, Mathew; van der Voo, Rob; Tomezzoli, Renata N.; Tohver, Eric; Hendriks, Bart W. H.; Torsvik, Trond H.; Vizan, Haroldo; Dominguez, Ada


    A major disparity is observed between the late Paleozoic-early Mesozoic apparent polar wander paths (APWPs) of Laurussia and Gondwana when the landmasses are re-assembled in a conventional "A-type" Pangea. This discrepancy has endured from the earliest paleomagnetic reconstructions of the supercontinent, and has prompted discussions of non-dipole paleomagnetic fields and alternative paleogeographic models. Here we report on a joint paleomagnetic-geochronologic study of Late Permian and Early to Middle Triassic volcanic and volcaniclastic rocks from Argentina, which demonstrates support for an A-type model, without requiring modification to the geocentric axial dipole hypothesis. New SHRIMP U-Pb and 40Ar-39Ar isotopic dating has reinforced the inferred age of the sequences, which we estimate at ˜264 Ma (Upper Choiyoi Group) and ˜245 Ma (Puesto Viejo Group). Field-stability tests demonstrate that the volcanic rocks are carrying early/primary magnetizations, which yield paleopoles: 73.7°S, 315.6°E, A95: 4.1°, N: 40 (Upper Choiyoi) and 76.7°S, 312.4°E, A95: 7.3°, N: 14 (Puesto Viejo). A comprehensive magnetic fabric analysis is used to evaluate structural restorations and to correct for magnetization anisotropy. The paleomagnetic results derived from volcaniclastic rocks are interpreted to be affected by inclination shallowing, and corrections are discussed. A comparison of these new results with the existing Permian-Triassic paleomagnetic data from Gondwana suggests the presence of widespread bias in the latter. We contend that such bias can explain the observed APWP disparity, at least for Late Permian-Middle Triassic time, and that alternative paleogeographic reconstructions or non-dipole paleomagnetic fields do not need to be invoked to resolve the discrepancy.

  19. Climatic - biotic continuum - a few examples from the Pennsylvanian - Early Permian

    Kossovaya, O.


    13C 0,5-1,0 ‰ in the overlay limestone. The taxonomic diversity is very low. The recovery of bentic biota is characteristic for the Asselian -Lower Artinskian and ended by abrupt extinction at the Late Artinskian or Kungurian. The paleoclimatic affinities of this event are rather controversial. In the Southern hemisphere the restriction of the glacial cover is supported by the appearance first temperate biota - forams, small solitary rugosa, bryozoan, rare bivalves and brachiopods, that are characteristic for the temperate water. This level coincides with colonial corals flourishing in the subtropical area. The average value within Asselian - Lower Artinskian in the Most section (Central Urals) is δ 13C 4- 5‰. The end of Early Artinskian coincides with the minimum value both δ13C and 18O probably mirroring Late Sakmarian deglaciation (Kossovaya et al., 2011). The obtained data in spite of the difference in the absolute value show the similar trend with published data (Korte et al., 2005). Following "small biotic event" fixed in the Northern Hemisphere is characterized by disappearance of Kleopatrinidae and Durhamihidae (Rugosa), other reef-building organisms, diversification of ostracods and replacement of the brachiopod genera composition. Possible trigger could be collision processes in the Eastern part of Pangea challenged the circulation change.

  20. Juvenile crustal recycling in an accretionary orogen: Insights from contrasting Early Permian granites from central Inner Mongolia, North China

    Yuan, Lingling; Zhang, Xiaohui; Xue, Fuhong; Liu, Fulin


    Coeval high-K calc-alkaline to alkaline granites constitute important components of post-collisional to post-orogenic igneous suites in most orogenic belts of various ages on Earth and their genesis harbors a key to ascertaining critical geodynamic controls on continental crustal formation and differentiation. This zircon U-Pb dating and geochemical study documents three contrasting Early Permian granites from Erenhot of central Inner Mongolia, eastern Central Asian Orogenic Belt (CAOB) and reveals concurrent high-K calc-alkaline to alkaline granite association derived from successive partial melting of distinct protoliths. The ca. 280 Ma Gancihuduge (GCG) pluton shows a calc-alkaline I-type character, with initial 87Sr/86Sr ratios of 0.7035 to 0.7039, εNd(t) of + 1.87 to + 4.70, zircon εHf(t) of + 8.0 to + 13.2 and δ18O from 7.4 to 8.7‰. The ca. 276 Ma Cailiwusu (CLS) pluton is magnesian and peraluminous, with initial 87Sr/86Sr ratios of 0.7036 to 0.7040, εNd(t) of + 1.9 to + 2.4, zircon εHf(t) of + 6.5 to + 12.1 and δ18O from 9.7 to 10.9‰. These features are consistent with partial melts of mixed sources composed of newly underplated meta-basaltic to -andesitic protoliths and variable supracrustal components, with distinctively higher proportion of the latter in the CLS pluton. By contrast, the ca. 279 Ma Kunduleng (KDL) suite exhibits an A-type magmatic affinity, with typical enrichment in alkalis, Ga, Zr, Nb and Y, εNd(t) of + 2.39 to + 3.55, zircon εHf(t) from + 8.3 to + 12.3 and δ18O values from 6.8 to 7.5‰. These features suggest that they stem from high-temperature fusion of dehydrated K-rich mafic to intermediate protoliths. Besides presenting a snapshot into a stratified crustal architecture in δ18O, these contrasting granites could not only serve as a temporal marker for monitoring post-collisional extension in the aftermath of a retreating subduction zone, but also present spatial magmatic proxy for tracing crustal formation and

  1. Circum-Pacific accretion of oceanic terranes to continental blocks: accretion of the Early Permian Dun Mountain ophiolite to the E Gondwana continental margin, South Island, New Zealand

    Robertson, Alastair


    Accretionary orogens, in part, grow as a result of the accretion of oceanic terranes to pre-existing continental blocks, as in the circum-Pacific and central Asian regions. However, the accretionary processes involved remain poorly understood. Here, we consider settings in which oceanic crust formed in a supra-subduction zone setting and later accreted to continental terranes (some, themselves of accretionary origin). Good examples include some Late Cretaceous ophiolites in SE Turkey, the Jurassic Coast Range ophiolite, W USA and the Early Permian Dun Mountain ophiolite of South Island, New Zealand. In the last two cases, the ophiolites are depositionally overlain by coarse clastic sedimentary rocks (e.g. Permian Upukerora Formation of South Island, NZ) that then pass upwards into very thick continental margin fore-arc basin sequences (Great Valley sequence, California; Matai sequence, South Island, NZ). Field observations, together with petrographical and geochemical studies in South Island, NZ, summarised here, provide evidence of terrane accretion processes. In a proposed tectonic model, the Early Permian Dun Mountain ophiolite was created by supra-subduction zone spreading above a W-dipping subduction zone (comparable to the present-day Izu-Bonin arc and fore arc, W Pacific). The SSZ oceanic crust in the New Zealand example is inferred to have included an intra-oceanic magmatic arc, which is no longer exposed (other than within a melange unit in Southland), but which is documented by petrographic and geochemical evidence. An additional subduction zone is likely to have dipped westwards beneath the E Gondwana margin during the Permian. As a result, relatively buoyant Early Permian supra-subduction zone oceanic crust was able to dock with the E Gondwana continental margin, terminating intra-oceanic subduction (although the exact timing is debatable). The amalgamation ('soft collision') was accompanied by crustal extension of the newly accreted oceanic slab, and

  2. A new Permian bivalve-dominated assemblage in the Rio do Rasto Formation, Paraná Basin, Brazil: Faunal turnover driven by regional-scale environmental changes in a vast epeiric sea

    Simões, Marcello Guimarães; Matos, Suzana Aparecida; Anelli, Luiz Eduardo; Rohn, Rosemarie; Warren, Lucas Veríssimo; David, Juliana Machado


    The basal portion of the Permian Rio do Rasto Formation (Serrinha Member), Passa Dois Group, Paraná Basin, Brazil, records an entirely new bivalve fauna intercalated between the underlying Pinzonella neotropica assemblage (uppermost portion of the Teresina Formation) and the overlying Leinzia similis assemblage (Rio do Rasto Formation). Mollusks of these assemblages lived in marginal shallow-water habitats of an immense epeiric sea and were dominated by endemic bivalve species. Taxonomic analysis revealed the presence of Terraia curvata (60.61%), Astartellopsis prosoclina (19.70%), Cowperesia emerita (10.61%), Leinzia curta (4.55%), Terraia bipleura (3.03%) and Beurlenella elongatella (1.52%), which are associated with conchostracans and plant remains. Species composition, abundance, and dominance in this novel assemblage differ notably from the preceding ones, suggesting a substantial evolutionary turnover. Regional-scale environmental changes recognized based on taphonomy, facies analysis, and geochemical data consist of progressive freshening of the marginal habitats of the Paraná Basin and taxic changes that include the following: (a) loss of genera, (b) decrease in bivalve abundance and ecological guilds, (c) disappearance of the dominant bivalve group (Pinzonellinae) and (d) diversification of Terrainae bivalves. The ecological signature also changed notably because only infaunal suspension-feeding bivalves are present, indicating a significant loss of functional diversity at the regional scale. Likely stressor factors (among others) are tied to freshening events, suggesting profound changes in (a) salinity, (b) primary productivity and (c) a lack of coarse, stable substrates coupled with high bioturbation rates. Hence, our regional example could offer valuable clues to benthic (bivalve) community responses in a habitat subjected to (a) rapid climate changes and (b) freshening events in shallow-water settings. Finally, the stratigraphic range of the

  3. Reassessment of petrogenesis of Carboniferous–Early Permian rift-related volcanic rocks in the Chinese Tianshan and its neighboring areas

    Linqi Xia


    Full Text Available The Carboniferous−Early Permian rift-related volcanic successions, covering large areas in the Chinese Tianshan and its adjacent areas, make up a newly recognized important Phanerozoic large igneous province in the world, which can be further divided into two sub-provinces: Tianshan and Tarim. The regional unconformity of Lower Carboniferous upon basement or pre-Carboniferous rocks, the ages (360–351 Ma of the youngest ophiolite and the peak of subduction metamorphism of high pressure–low temperature metamorphic belt and the occurrence of Ni-Cu-bearing mafic-ultramafic intrusion with age of ∼352 Ma and A-type granite with age of ∼358 Ma reveal that the final closure of the Paleo-Asian Ocean might take place in the Early Mississippian. Our summation shows that at least four criteria, being normally used to identify ancient asthenosphere upwelling (or mantle plumes, are met for this large igneous province: (1 surface uplift prior to magmatism; (2 being associated with continental rifting and breakup events; (3 chemical characteristics of asthenosphere (or plume derived basalts; (4 close links to large-scale mineralization and the uncontaminated basalts, being analogous to those of many “ore-bearing” large igneous provinces, display Sr-Nd isotopic variations between plume and EM1 geochemical signatures. These suggest that a Carboniferous asthenosphere upwelling and an Early Permian plume played the central role in the generation of the Tianshan–Tarim (central Asia large igneous province.

  4. Confirmation of Sigillaria Brongniart as a coal-forming plant in Cathaysia: occurrence from an Early Permian autochthonous peat-forming flora in Inner Mongolia

    Wang, J.; Feng, Z.; Zhang, Y.; Wang, S.J. [Chinese Academy of Sciences, Nanjing (China)


    A common lycopsid genus, Sigillaria Brongniart, has been recorded most frequently in peat-forming forests in Europe and North America, but rarely in China. Although Sigillaria, in China, has been found in coal balls and used as evidence that it was a coalforming element, it has never been recorded as compression/impressions in peat-forming settings. Recent investigation of an Early Permian autochthonous peat-forming flora of the Taiyuan Formation near Wuda, Inner Mongolia, has provided evidence that Sigillaria could be a major element of peat-forming vegetation in China. Sigillaria is the only arborescent lycopsid acting as a major contributor to peat/coal formation. The genus Sigillaria is another element that China has in common with the Palaeozoic low-land floras of Europe and North America, enhancing the common floral aspects between the tropical swamp vegetations of the east and west regions of the Palaeotethys Ocean.

  5. Application of Markov chain and entropy analysis to lithologic succession – an example from the early Permian Barakar Formation, Bellampalli coalfield, Andhra Pradesh, India

    Ram Chandra Tewari; D P Singh; Z A Khan


    A statistical approach by a modified Markov process model and entropy function is used to prove that the early Permian Barakar Formation of the Bellampalli coal field developed distinct cyclicities during deposition.From results,the transition path of lithological states typical for the Bellampalli basin is as:coarse to medium-grained sandstone → interbedded fine-grained sandstone/shale → shale → coal and again shale.The majority of cycles are symmetrical but asymmetrical cycles are present as well.The chi-square stationarity test implies that these cycles are stationary in space and time.The cycles are interpreted in terms of in-channel,point bar and overbank facies association in a fluvial system.The randomness in the occurrence of facies within a cycle is evaluated in terms of entropy,which can be calculated from the Markov matrices.Two types of entropies are calculated for every facies state;entropy after deposition (post)and entropy before deposition (pre),which together form entropy set;the entropy for the whole system is also calculated.These values are plotted and compared with Hattori ’s idealized plots,which indicate that the sequence is essentially a symmetrical cycle (type-B of Hattroi). The symmetrical cyclical deposition of early Permian Barakar Formation is explained by the lateral migration of stream channels in response to varying discharge and rate of deposition across the alluvial plain.In addition,the fining upward cycles in the upper part enclosing thick beds of fine clastics,as well as coal may represent differential subsidence of depositional basin.

  6. Caltepec fault zone: An Early Permian dextral transpressional boundary between the Proterozoic Oaxacan and Paleozoic Acatlán complexes, southern Mexico, and regional tectonic implications

    Elías-Herrera, Mariano; Ortega-Gutiérrez, Fernando


    The tectonic boundary between the Grenville-age Oaxacan and Paleozoic Acatlán crystalline complexes in southern Mexico, named the Caltepec fault zone (CFZ), is characterized for the first time as a dextral transpressional, NNW trending and ENE dipping ductile fault zone of Early Permian age. The complexes are welded by a syntectonic magmatic epidote-bearing granite along the entire length of the CFZ. From east to west, the 2-6 km wide CFZ consists of disrupted and retrograded banded gneisses of the Oaxacan complex, quartz-feldspar mylonite, and the syntectonic magmatic epidote-bearing Cozahuico granite (CZG) with huge xenoliths (up to several kilometers long and up to 600 m wide) of the Proterozoic gneisses, thrust westward over metasedimentary tectonites of the Acatlán complex. The CZG shows magmatic fabrics that represent a transition to solid-state deformation characterized by subvertical foliation, subhorizontal NNE and SSE dipping mineral stretching lineation and dextral kinematics. The megaxenoliths underwent partial melting developing banded migmatites with layers of epidote-bearing granitic neosome. The parallelism of fabrics in these anatexitic rocks and in the enclosing deformed granite suggests that ductile deformation, migmatization of xenolithic gneisses, and granite emplacement along the CFZ were coeval. The neosome yielded a U-Pb zircon concordant age of 275.6 +/- 1 Ma probably dating the peak of the tectonothermal event. We interpret the CFZ as a major terrane boundary accommodating transpressional interaction between the Acatlán and Oaxaquia blocks, which were amalgamated in an oblique convergent setting by Early Permian time, as the leading edge of Gondwana impinged onto the southern margin of Laurentia along the Marathon-Ouachita suture to form Pangea.

  7. Discovery of organic connection of Chiropteris Kurr and Nystroemia Halle from Early Permian of western Henan, China

    WANG Jun; H. W. Pfefferkorn; SUN Bainian; LIU Lujun


    Nystroemia reniformis (Kawasaki) comb. nov. is proposed based on specimens showing organic connections between Chiropteris reniformis Kawasaki and Nystroemia pectiniformis Halle from the Lower Permian of Dengfeng in Henan. Vegetative leaf has a long petiole, reniformed, cordate at the base, entire or slightly undulate at the margin; veins are fine, bifurcating repeatedly and finally becoming reticulate and forming polygonal or rhomboidal meshes. Fertile shoot arises from bract axils. The form of the bract is identical to that of a vegetative leaf. Ovuliferous organ arises from the axil of leafy organ, or in other case, both ovuliferous and leafy organs irregularly arise from the axis of the fertile shoot respectively. Numerous bicornute platyspermic seeds are arranged in two rows, obconical, with micropyle pointing upwards. It may represent a new extinct gymnosperms.

  8. Petrography and geochemistry of the Permian-Triassic boundary interval, Yangou section, South China: Implications for early Griesbachian seawater δ13CDIC gradient with depth

    Li, Rong


    The carbon isotopic composition (δ13Ccarb) recorded in shelf carbonates has been widely used as a proxy for the isotopic composition (δ13CDIC) of surface ocean water to establish paleocean chemistry and circulation patterns. However, δ13Ccarb values do not necessarily preserve the δ13CDIC, due to post-depositional diagenetic alteration. In order to examine the early Griesbachian surface-to-deep δ13CDIC gradient with depth, the diagenetic features of the Permian-Triassic boundary interval (beds 18 to 35) from Yangou section, located in the Yangtze carbonate platform interior, South China, are delineated to compare with those of the slope GSSP Meishan section. The petrographic and geochemical observations show that the early Griesbachian carbonates in the Yangou section underwent pervasive dolomitization in its early diagenetic history. Three types of early replacement dolomites and one type of dolomite cement are present. The dolomite crystals display internal zonation, with high-Ca calcian dolomite (HCD) core being encased successively by calcite and an outermost Fe-rich HCD cortex. The initial dolomitization took place in anoxic seawater, and underwent subsequent diagenetic system involved with meteoric water. The two most negative δ13C values in claystones of Beds 21-3 and 35 are probably related to meteoric diagenesis. Above and/or below the meteorically influenced beds, the dolomite and calcite have uniformly positive δ13C values. The primary carbon isotopic compositions are probably preserved in the early Griesbachian carbonate from the platform Yangou section, which could probably be related to the poor formation of the outermost Fe-rich HCD cortex. Compared to the slope carbonate from the Meishan section, the platform carbonate from the Yangou section has lower primary δ13Ccarb values. It is estimated that the δ13CDIC gradient with depth between Yangou and Meishan is less than the previously suggested. The results highlight the need for evaluation

  9. An early bird from Gondwana: Paleomagnetism of Lower Permian lavas from northern Qiangtang (Tibet) and the geography of the Paleo-Tethys

    Song, Peiping; Ding, Lin; Li, Zhenyu; Lippert, Peter C.; Yue, Yahui


    The origin of the northern Qiangtang block and its Late Paleozoic-Early Mesozoic drift history remain controversial, largely because paleomagnetic constraints from pre-Mesozoic units are sparse and of poor quality. In this paper, we provide a robust and well-dated paleomagnetic pole from the Lower Permian Kaixinling Group lavas on the northern Qiangtang block. This pole suggests that the northern Qiangtang block had a paleolatitude of 21.9 ± 4.7 °S at ca. 296.9 ± 1.9 Ma. These are the first volcanic-based paleomagnetic results from pre-Mesozoic rocks of the Qiangtang block that appear to average secular variation accurately enough to yield a well-determined paleolatitude estimate. This new pole corroborates the hypothesis, first noted on the basis of less rigorous paleomagnetic data, the presence of diamictites, detrital zircon provenance records, and faunal assemblages, that the northern Qiangtang block rifted away from Gondwana prior to the Permian. Previous studies have documented that the northern Qiangtang block accreted to the Tarim-North China continent by Norian time. We calculate a total northward drift of ca. 7000 km over ca. 100 myr, which corresponds to an average south-north plate velocities of ∼7.0 cm/yr. Our results do not support the conclusion that northern Qiangtang has a Laurasian affinity, nor that the central Qiangtang metamorphic belt is an in situ Paleo-Tethys suture. Our analysis, however, does not preclude paleogeographies that interpret the central Qiangtang metamorphic belt as an intra-Qiangtang suture that developed at southernly latitudes outboard of the Gondwanan margin. We emphasize that rigorous paleomagnetic data from Carboniferous units of northern Qiangtang and especially upper Paleozoic units from southern Qiangtang can test and further refine these paleogeographic interpretations.

  10. A low diversity, seasonal tropical landscape dominated by conifers and peltasperms: Early Permian Abo Formation, New Mexico

    DiMichele, W.A.; Chaney, D.S.; Nelson, W.J.; Lucas, S.G.; Looy, C.V.; Quick, K.; Jun, W.


    Walchian conifers (Walchia piniformis Sternberg, 1825) and peltasperms similar to Supaia thinnfeldioides White and cf. Supaia anomala White dominate floodplain deposits of a narrow stratigraphic interval of the middle Abo Formation, Lower Permian of central New Mexico. The plant fossils occur in thinly bedded units up to two meters thick, consisting of coarse siltstone to very fine sandstone with clay partings. Bedding is primarily tabular, thin, and bears rare ripple marks and trough cross beds. Bedding surfaces display mud cracks, raindrop imprints, horizontal and vertical burrows of invertebrates, and footprints of terrestrial vertebrates. These features indicate intermittent and generally unchannelized stream flow, with repeated exposure to air. Channels appear to have cannibalized one another on a slowly subsiding coastal plain. Conifers are dominant at three collecting sites and at three others Supaia dominates. Although each of these genera occurs in assemblages dominated by the other, there are no truly co-dominant assemblages. This pattern suggests alternative explanations. Landscapes could have consisted of a small-scale vegetational patchwork dominated almost monospecifically in any one patch, meaning that these plants could have coexisted across the landscape. On the other hand, conifer and supaioid dominance could have been temporally distinct, occurring during different episodes of sedimentation; although in the field there are no noticeable sedimentological differences between conifer-dominated and Supaia-dominated channel deposits, they may represent slightly different climatic regimes. The considerable morphological differences between conifers and Supaia suggest that the floristic patterns are not a taphonomic effect of the loss of a significant part of the original biodiversity. In general, the climate under which this vegetation developed appears to have been relatively warm and arid, based on the geology (pervasive red color [oxidation

  11. Early Permian extensional shearing of an Ordovician granite: The Saint-Eutrope "C/S-like" orthogneiss (Montagne Noire, French Massif Central)

    Pitra, Pavel; Poujol, Marc; Van Den Driessche, Jean; Poilvet, Jean-Charles; Paquette, Jean-Louis


    Dating the magmatic events in the Montagne Noire gneiss dome is a key point to arbitrate between the different interpretations of the Late Carboniferous-Early Permian tectonics in this southern part of the Variscan belt. The Saint-Eutrope orthogneiss crops out along the northern flank of the dome. We show that the protolith of this orthogneiss is an Ordovician granite dated at 455 ± 2 Ma (LA-ICP-MS U-Pb dating on zircon). This age is identical to that previously obtained on the augen orthogneiss of the southern flank, strongly suggesting that both orthogneiss occurrences have the same Ordovician protolith. The Saint-Eutrope orthogneiss experienced intense shearing along the Espinouse extensional detachment at ca. 295 Ma (LA-ICP-MS U-Pb-Th on monazite), an age close to that determined previously on mica by the 39Ar-40Ar method and contemporaneous with the emplacement age of the syntectonic Montalet granite farther to the west. This normal sense shearing reworked previous fabrics related to Variscan thrusting that can be still observed in the augen orthogneiss of the southern flank, and is responsible for the spectacular "C/S-like" pattern of the Saint-Eutrope orthogneiss. This work also shows that care is needed when dealing with C/S-type structures, since they can develop not only in syntectonic intrusions, but also in orthogneisses affected by an intense secondary deformation, at decreasing temperature.

  12. A Late Carboniferous-Early Permian slab window in the West Junggar of NW China: Geochronological and geochemical evidence from mafic to intermediate dikes

    Yin, Jiyuan; Long, Xiaoping; Yuan, Chao; Sun, Min; Zhao, Guochun; Geng, Hongyan


    The Late Carboniferous to Early Permian is a critical period for the formation of particular high-temperature magmatic associations in West Junggar, NW China and the geodynamic processes of ridge subduction and related slab windows in the Central Asian Orogenic Belt. Mafic to intermediate dikes, exposed in the central and southeastern West Junggar extensively, mainly consist of Nb-enriched and magnesian suites. The Nb-enriched dikes (305 Ma) are characterized by high Nb contents (5.63-9.08 ppm) and large variations of major element contents, exhibiting high-K to medium-K calc-alkaline characteristics with high Na2O/K2O (1.33-3.85) and low Sr/Y (9-32) ratios. The dikes have moderate Mg# (44.6-52.9), LREE-rich and sub-horizontal HREE patterns ((La/Yb)N = 3.05-7.18; (Gd/Yb)N = 1.43-1.78) with weak Eu anomalies (Eu*/Eu = 0.81-1.10) and positive εNd (t) values (+ 5.2 to + 8.0), and likely originate from partial melting of mantle peridotite that was metasomatized by subducted sediment-derived melts and slab fluids. The medium-K magnesian dikes (305 Ma) are characterized by high MgO (3.05-7.28 wt.%), Cr (22.3-311 ppm) and Ni (32.6-199 ppm) contents and all plot in the field of high-Mg andesite. Some have very high Mg numbers (Mg# > 60), suggesting a primitive melt composition. These samples display positive Ba, Sr, K and negative Nb-Ta-Ti anomalies as well as strongly-depleted HREE patterns with weak positive Eu anomalies. They have high Sr/Y (27-126) and relatively high εNd (t) values (+ 6.0 to + 8.3), which suggest a similar petrogenesis as the Nb-enriched dikes, but were metasomatized by adakitic melts. The high-K magnesian dikes (284 Ma) are high-K calc-alkaline and have lower MgO (3.63-5.57 wt.%) compared with the medium-K suite. They have high Sr (422-819 ppm), Sr/Y (26-62) ratios and εNd (t) values (+ 7.0 to + 7.1) with relatively flat REE patterns ((La/Yb)N = 2.4-5.5), probably originating from partial melting of K-enriched mantle wedge peridotite metasomatized

  13. Petrogenesis of the Early Permian volcanic rocks in the Chinese South Tianshan: Implications for crustal growth in the Central Asian Orogenic Belt

    Huang, He; Zhang, Zhaochong; Santosh, M.; Zhang, Dongyang; Wang, Tao


    The Paleozoic and Early Mesozoic magmatic suites in the Central Asian Orogenic Belt (CAOB) provide important insights on the crustal growth and reworking process associated with the construction of the largest Phanerozoic orogen on the Earth. Among the tectonic blocks of the CAOB, the South Tianshan Terrane (STT) occupies the southwestern margin and is located adjacent to the Tarim Craton. Here we investigate the Early Permian Xiaotikanlike Formation in the central part of the Chinese STT in Xinjiang in Northwest China. The formation is composed of a series of terrestrial volcanic lava flows and volcanic breccia, interbedded with siltstones, sandstones and sandy conglomerates. Zircon U-Pb and Lu-Hf isotopic analysis, whole-rock major oxide, trace element and Sr-Nd isotopic data are presented for the volcanic lava flows of the Xiaotikanlike Formation exposed in the Boziguo'er, Laohutai and Wensu regions. The new zircon ages from our study, together with those reported in previous investigations on the rhyolitic lava flow from the Wensu region, suggest that the volcanic rocks of the Xiaotikanlike Formation simultaneously erupted at ca. 285 Ma. The lavas of the formation show a wide range of SiO2 (49.88 to 78.56 wt.%). The basaltic rocks show SiO2 from 49.88 to 53.78 wt.%, MgO from 3.73 to 7.01 wt.% and Mg# from 41 to 61. They possess slightly enriched Sr-Nd isotope signature [(87Sr/86Sr)t = 0.70495-0.70624 and εNd(t) = - 0.5 to + 0.6], and have trace and rare earth element patterns similar to those of oceanic island basalts (OIBs). Petrographic and whole-rock chemical characteristics indicate that the basaltic lava flows are dominantly tholeiitic, and were likely derived from a spinel-dominated peridotite asthenospheric mantle source. The felsic lavas of the Xiaotikanlike Formation show SiO2 in the range of 60.71 to 78.56 wt.% and display overall similar immobile element pattern characterized by notable troughs at Nb-Ta, P and Ti and gently sloping REEs. Zircon Lu

  14. A Re-Description of 'Mycterosaurus' smithae, an Early Permian Eothyridid, and Its Impact on the Phylogeny of Pelycosaurian-Grade Synapsids.

    Neil Brocklehurst

    Full Text Available 'Mycterosaurus' smithae, from the Cisuralian (early Permian of Colorado, was first described in 1965 as a second species of the genus Mycterosaurus. While the type species of this genus, M. longiceps, has been shown by multiple cladistic analyses to belong to the basal synapsid family Varanopidae, 'M.' smithae has been largely ignored since its original description. Additional preparation and synchrotron scanning has revealed new significant information that supports the assignment of this species to a new genus: Vaughnictis gen. nov. Vaughnictis lacks many of the characteristics of mycterosaurines and varanopids in general: it lacks the slender femur, the linguo-labially compressed and strongly recurved teeth, and the lateral boss on the postorbital characteristic of this family. Instead, it possesses coronoid teeth, a large supratemporal, and a large pineal foramen positioned midway along the length of the parietal, features that support its assignment to Eothyrididae. Moreover, the postcranium shares many characters with the eothyridid Oedaleops. An expanded version of a recently published phylogenetic analysis of pelycosaurian-grade synapsids positions Vaughnictis as the sister taxon of Eothyris within the clade Eothyrididae. The addition of data on the postcranium of eothyridids and the inclusion of the recently-described basal caseid Eocasea confirms the recently-disputed position of caseasaurs as the most basal synapsids. As the parsimony analysis produced low support values and a lack of resolution due to missing data, additional analyses were undertaken using Bayesian and Implied Weights methods, which produced better resolution and relationships with higher support values. While the results are similar, alternative positions for the enigmatic Moscovian age (Carboniferous synapsid Echinerpeton are suggested by Bayesian analysis; the parsimony analysis found it to be an ophiacodontid, while the Bayesian and Implied Weights analysis found

  15. Spectral gamma-ray signature of fluvial deposits: a case study from the Late Permian Rio do Rasto Formation, Parana Basin, Brazil; Assinatura gamaespectrometrica de depositos fluviais: estudo de caso na Formacao do Rio do Rasto, Permiano Superior da Bacia do Parana

    Sowek, Guilherme Arruda, E-mail: [Universidade Federal do Parana (UFPR), Curitiba, PR (Brazil). Programa de Pos-graduacao em Geologia; Ferreira, Francisco Jose Fonseca; Vesely, Fernando Farias, E-mail:, E-mail: [Universidade Federal do Parana (UFPR), Curitiba, PR (Brazil). Dept. de Geologia. Setor de Ciencias da Terra; Berton, Fabio, E-mail: [Universidade Federal do Parana (UFPR), Curitiba, PR (Brazil)


    Fluvial channel-fill deposits form highly heterogeneous hydrocarbon reservoirs. The study of outcrop analogs can help in the characterization of these heterogeneities, which are usually not detected by subsurface geophysical methods. The aim of this research is to compare outcrop log signatures with grain size trends and depositional elements of the fluvial deposits of the Late Permian Rio do Rasto Formation. A series of vertical gamma-ray logs were assembled in two outcrops in order to: 1) characterize log-facies in a succession composed of alternated flood plain, channel fill and eolian strata; 2) define within-channel spectral gamma-ray variability of a mixed-load composite point bar deposit and its relationship with grain size trends and lithofacies; 3) correlate log signatures observed in the outcrop sections with deep exploratory wells drilled several tens of kilometers from the study area. The results of this study show that gamma-ray logs have good correlation with grain size trends and that different depositional elements have distinct signatures. On the other hand, point bar deposits exhibit strong lateral changes in log signature due variations in grain size and mud content within lateral accretion strata. Although frequent, the classic bell-shaped log motif was not always detected, which means that the amount of fluvial channel-fill deposits recognized in subsurface can be underestimated. Similar log signatures were detected in the boreholes, at least in the closest ones, helping in paleoenvironmental interpretation in the subsurface. (author)

  16. Subduction-related Late Carboniferous to Early Permian Magmatism in the Eastern Pontides, the Camlik and Casurluk plutons: Insights from geochemistry, whole-rock Sr-Nd and in situ zircon Lu-Hf isotopes, and U-Pb geochronology

    Karsli, Orhan; Dokuz, Abdurrahman; Kandemir, Raif


    Late Carboniferous to early Permian granitoid rocks represent a volumetrically minor component of the Eastern Pontide lithosphere, but they preserve useful information about the region's tectonomagmatic history. The Casurluk and Camlik plutons primarily consist of gabbro, gabbroic diorite, diorite, monzogabbro, monzodiorite and monzonite, which intrude early to middle Carboniferous granitic basement rocks in the region. In this study, we use in situ zircon U-Pb ages and Lu-Hf isotopic values, whole-rock Sr-Nd isotopic values, and mineral chemistry and geochemistry of these plutons to determine petrogenesis and crustal evolution; we also discuss geodynamic implications. LA-ICP-MS zircon U-Pb dating of magmatic zircons from the rocks suggests that the plutons were emplaced during the late Carboniferous to early Permian (302 Ma). The metaluminous and I-type intrusive rocks belong to the high-K calc-alkaline series. In addition, they are relatively enriched in light rare earth elements (LREEs) and large-ion lithophile elements (LILEs); they are depleted in heavy rare earth elements (HREEs) and high field strength elements (HFSEs), such as Nb and Ti. All of the samples have homogeneous initial ISr values (0.70675 to 0.70792) and low εNd (t) values (- 5.1 to - 3.3). Zircons from the rocks of both plutons have uniform negative to slightly positive εHf (t) values (- 3.5 to 1.4) and old Hf two-stage model ages (1323 to 1548 Ma), implying that they have the same source, as well as suggesting the involvement of old enriched lithospheric mantle materials during their magma genesis. These results, combined with the εHf (t) values and two-stage model ages, demonstrate that the primary magmas were derived from partial melting of old lithospheric mantle material metasomatized by subduction-related fluids. Considering other regional geological data from the Sakarya Zone where these plutons formed, we conclude that late Carboniferous to early Permian magmatism in the area




    Full Text Available Early Triassic (Griesbachian to Spathian ostracod faunas are here first discovered and described form the Guangxi Province, South China. Thirty-seven species belonging to fourteen genera are recognized. Seven species are new: Bairdia fengshanensis n.sp., Bairdia wailiensis n.sp., Liuzhinia guangxiensis n.sp., Ptychobairdia luciaae n.sp., Ptychobairdia aldaae n.sp., Paracypris jinyaensis n.sp. and Paracypris gaetanii n.sp. The Griesbachian assemblage from the basal microbial limestone is well diversified and does not suggest any abnormal palaeoenvironmental conditions in terms of salinity, temperature or oxygen content. Particularly, the ostracods are typical of well oxygenated water and do not reflect any anoxia. Dienerian and Smithian ostracods are evidenced for the first time and the assemblages suggest less favourable palaeoenviromental conditions. Diversity and abundance of ostracod assemblages recovered from the Spathian on. The main taxonomic turnover among ostracod assemblages occurred seemingly between the Griesbachian and the Spathian.

  18. Petrogenesis and tectonic implications of the Early Carboniferous to the Late Permian Barleik plutons in the West Junggar (NW China)

    Liu, Bo; Han, Bao-Fu; Ren, Rong; Chen, Jia-Fu; Wang, Zeng-Zhen; Zheng, Bo


    The Paleozoic accretionary orogenesis and continental crustal growth in Central Asia are thought to have close relationship with the evolution of the Paleo-Asian Ocean (PAO). The well-exposed plutons in the northern Barleik Mountains of the West Junggar region, NW China, may provide essential insights into the evolution of the Junggar Ocean, a branch of the PAO, and mechanism of continental crustal growth. Our work on the Barleik plutons indicates an early suite of 324-320 Ma diorite and a late suite of 314-259 Ma quartz syenite and granitic porphyry. All the plutons are characterized by high-K calc-alkaline to shoshonitic signatures, varying depletion in Nb, Ta, Sr, P, Eu, and Ti, low initial 87Sr/86Sr ratios (0.70241-0.70585), strongly positive εNd(t) values (+ 5.7-+7.7), and young one-stage Nd model ages (390-761 Ma), suggesting that they resulted from different batches of magma that were produced by fractional crystallization of a metasomatized mantle source with minor crustal contamination. The diorite is coeval with the youngest arc magmatic rocks, indicating a subduction-related origin. By contrast, the quartz syenite and granitic porphyry are geochemically similar to A2-type granites, with high Zr, Ga, and FeOT/[FeOT + MgO], and are coeval with the widespread plutons in the West Junggar. This, together with the occurrence of Late Carboniferous fluvial deposits and the lack of < 320 Ma ophiolitic and subduction-related metamorphic lithologies, definitively indicates a post-collisional setting after the closure of the Junggar Ocean. Slab breakoff accompanied by asthenospheric upwelling and basaltic underplating is a possible geodynamic process that is responsible for the post-collisional magmatism and vertical crustal growth in the region. Thus a tectonic switch from subduction to post-collision started at the end of the Early Carboniferous ( 320 Ma), probably as a result of the final closure of the Junggar Ocean.

  19. Palynostratigraphy of Permian succession in the Mand–Raigarh Coalfield, Chhattisgarh, India and phytogeographical provincialism

    Srikanta Murthy; Ram-Awatar; Saurabh Gautam


    Palynofloras have been recorded from the Barakar Formation in the Borehole MBKW-3, Barpali–Karmitikra Block, Mand–Raigarh Coalfield, Chhattisgarh. Three distinct palynoassemblages have been identified and referred to the following palynoassemblage zones – Gondisporites raniganjensis (Latest Permian); Faunipollenites varius (latest Early Permian), and Scheuringipollenites barakarensis (late Early Permian). It is inferred that these deposits contain the representative palynoassamblages of Early to Late Permian in age. The First Appearance Datum (FAD)s of Arcuatipollenites pellucidus, A. ovatus, Guttulapollenites hannonicus, Lundbladispora microconata, Alisporites opii, Klausipollenites sp., and Goubinispora indica (at 41.95, 45.90, 98.35 m depths), indicate the closing phase of Permian, as these elements are the key species that mark a transition from Permian to the Lower Triassic. An attempt has been made here to reconstruct the phytogeographical provincialism on the basis of Guttulapollenites recorded in this basin.

  20. Tectonic transition from Late Carboniferous subduction to Early Permian post-collisional extension in the Eastern Tianshan, NW China: Insights from geochronology and geochemistry of mafic-intermediate intrusions

    Zhang, Xiaoran; Zhao, Guochun; Eizenhöfer, Paul R.; Sun, Min; Han, Yigui; Hou, Wenzhu; Liu, Dongxing; Wang, Bo; Liu, Qian; Xu, Bing; Zhu, Chloe Yanlin


    The closure of the Junggar Ocean between the Central Tianshan and Junggar terranes is essential in understanding the final assembly of the southern Central Asian Orogenic Belt. This study presents new whole-rock geochemical, Sr-Nd and zircon U-Pb-Hf isotopic data for mafic-intermediate intrusions from the Central Tianshan block to provide robust constraints on the final closure of the ocean. LA-ICP-MS U-Pb dating on magmatic-type zircons yields weighted mean 206Pb/238U ages of ca. 310 Ma and ca. 290 Ma, which are interpreted as the crystallization ages of the intrusions. Petrographic and geochemical analyses of the Late Carboniferous mafic-intermediate rocks, characterized by typical subduction-related signatures, low Sm/Yb (plate. In contrast, the Early Permian mafic magmatism exhibits typical within-plate basalt affinities, such as high TiO2 (2.7-3.2 wt.%) contents, elevated Ti/V (86.0-115.1) and Zr/Y (4.9-9.3) ratios, OIB-like trace element patterns and high εNd(t) (+ 1.1 to + 4.5) and zircon εHf(t) (+ 3.0 to + 9.8) values. In association with previous investigations, we suggest that their protoliths were most probably derived from the partial melting of an asthenospheric mantle source in the garnet stability field, plausibly induced by asthenosphere upwelling during the slab break-off of the Junggar oceanic plate, which agrees well with the linear distributions of Permian mafic-ultramafic rocks in the Eastern Tianshan. Collectively, our data pinpoint a tectonic transition from oceanic subduction to post-collisional extension during Late Carboniferous to Early Permian time, probably triggered by the closure of the Junggar Ocean and subsequent arc-continent collision between the Central Tianshan and Junggar terranes that gave rise to the final assembly of the Eastern Tianshan.

  1. Permian stratigraphy and correlation of Northeast China: A review

    Shen, S.-Z.; Zhang, H.; Shang, Q. H.; Li, W.-Z.


    Palaeontological, lithostratigraphical data from the Permian strata and correlation of the Permian successions for different tectonic units in Northeast China are reviewed and summarized in this paper. Permian strata in Northeast China are dominated by brachiopods, fusulinoideans and land plants, with limited ammonoids, conodonts and bivalves. The Cisuralian (Early Permian) in the northern margin of the North China Block and in the Manchuride Belt is composed mostly of marine massive limestone with the characteristic Pseudoschwagerina Zone in the Asselian and Sakmarian and the Misellina claudiae Zone in the Kungurian. The Cisuralian in the Xing'an Block and the northeastern part of Inner Mongolia is dominated by huge terrestrial deposits with fossil plants. The Guadalupian (Middle Permian) in the Manchuride, Altaid and Yanbian Belts are characterized by bi-temperate Roadian or early Wordian Monodiexodina fauna and the late Wordian-Capitanian Codonofusiella- Schwagerina or Neoschwagerina- Yabeina faunas, the mixed brachiopod faunas between the Boreal/antitropical and the Palaeoequatorial Cathaysian forms, the Roadian or early Wordian solitary coral faunas, and the late Wordian-Capitanian compound Waagenophyllum- Wentzelella fauna. The Nadanhada Terrane contains some exotic limestone blocks with a typical Cathaysian Neoschwagerina- Yabeina fauna in a Late Jurassic-Early Cretaceous mélange, which is related to Mesozoic subduction in the western Circum-Pacific region. The Lopingian (Late Permian) in Northeast China is mostly characterized by terrestrial molasse deposits with a mixed flora between the Boreal Angaran and the palaeoequatorial Cathaysian Provinces, indicating the final closure of the Palaeo-Asian Ocean.

  2. Extinguishing a Permian World

    Schneebeli-Hermann, E.


    At the end of the Permian, ca. 252 Ma ago, marine and terrestrial fauna were facing the most extensive mass extinction in Earth history (Raup and Sepkoski, 1982). 80%–95% of all species on Earth, on land and in the oceans, became extinct (Benton et al., 2004) within an estimated time interval of les

  3. Extinguishing a Permian World

    Schneebeli-Hermann, E.


    At the end of the Permian, ca. 252 Ma ago, marine and terrestrial fauna were facing the most extensive mass extinction in Earth history (Raup and Sepkoski, 1982). 80%–95% of all species on Earth, on land and in the oceans, became extinct (Benton et al., 2004) within an estimated time interval of

  4. Petrogenesis of the flood basalts from the Early Permian Panjal Traps, Kashmir, India: Geochemical evidence for shallow melting of the mantle

    Shellnutt, J. Gregory; Bhat, Ghulam M.; Wang, Kuo-Lung; Brookfield, Michael E.; Jahn, Bor-Ming; Dostal, Jaroslav


    The Early Permian Panjal Traps of northern India represent a significant eruption of volcanic rocks which occurred during the opening of the Neotethys Ocean. Basaltic, basaltic-andesites, dacitic and rhyolitic rocks collected from Guryal Ravine and Pahalgam show evidence for subaerial and subaqueous eruptions indicating that they are contemporaneous with the formation of a shallow marine basin. The major and trace element geochemistry of the basalts is consistent with a within-plate setting and there are basalts which have high-Ti (TiO2 > 2.0 wt.%) and low-Ti (TiO2 < 1.8 wt.%) compositions. The ‘high-Ti’ basalts are similar to OIB whereas the ‘low-Ti’ basalts are similar to continental tholeiites. The identification of ‘high- and low-Ti’ basalts within the Panjal Traps is analogous to other large igneous provinces (e.g. Karoo, Deccan, Parana, Emeishan). The Sr-Nd isotopic values (εNd(T) = - 5.3 to + 1.3; ISr = 0.70432 to 0.71168) of both types of basalts overlap indicating that the rocks may have originated from the same ancient subcontinental lithospheric (i.e. EMII-like) mantle source (TDM = ~ 2000 Ma). The two groups of basalts can be modeled by using a primitive mantle source and different degrees of partial melting where the high-Ti rocks are produced by ~ 1% partial melting of a spinel peridotite source whereas the low-Ti rocks are produced by ~ 8% partial melting. Trace elemental and isotope modeling indicates that some of the basalts assimilated ≤ 10% crustal material. In contrast, the basaltic-andesites are likely formed by mixing between basaltic magmas and crustal melts which produced rocks with higher SiO2 (~ 55 wt.%) content and enriched isotopic signatures (εNd(T) = - 6.1; ISr = 0.70992). The Panjal Trap volcanism was likely due to partial melting of the SCLM within a passive extensional setting related to the rifting of Cimmeria from Gondwana. Contemporaneous volcanic and plutonic granitic rocks throughout the Himalaya are probably

  5. The petrogenesis of the Early Permian Variscan granites of the Cornubian Batholith: Lower plate post-collisional peraluminous magmatism in the Rhenohercynian Zone of SW England

    Simons, B.; Shail, Robin K.; Andersen, Jens C. Ø.


    The Early Permian Cornubian Batholith was generated during an extensional regime following Variscan convergence within the Rhenohercynian Zone of SW England. Its component granites can be classified, using mineralogical, textural and geochemical criteria, into five main types, all of which are peraluminous (A/CNK > 1.1): G1 (two-mica), G2 (muscovite), G3 (biotite), G4 (tourmaline) and G5 (topaz). G1 granites formed through up to 20% muscovite and minor biotite dehydration melting of a metagreywacke source at moderate temperatures and pressures (731-806 °C, > 5 kbar). Younger G3 granites formed through higher temperature, lower pressure (768-847 °C, < 4 kbar) biotite-dominated melting of a similar source. Partial melting was strongly influenced by the progressive lower-mid crustal emplacement of mafic igneous rocks during post-Variscan extension and a minor (< 5%-10%) mantle-derived component in the granites is possible. Two distinct fractionation series, G1-G2 and G3-G4, are defined using whole-rock geochemical and mineral chemical data. Variations in the major elements, Ba, Sr and Rb indicate that G1 and G3 granites underwent 15%-30% fractionation of an assemblage dominated by plagioclase, alkali feldspar and biotite to form more evolved G2 and G4 granites, respectively. Decreasing whole-rock abundances of Zr, Th and REE support the fractionation of zircon, monazite, apatite and allanite. Subsolidus alteration in G2 and G4 granites is indicated by non-primary muscovite and tourmaline and modification of major and trace element trends for G3-G4 granites, particularly for P2O5 and Rb. Topaz (G5) granites show low Zr, REE and extreme enrichment in Rb (up to 1530 ppm) and Nb (79 ppm) that cannot be related in a straightforward manner to continued differentiation of the G1-G2 or G3-G4 series. Instead, they are considered to represent partial melting, mediated by granulite facies fluids, of a biotite-rich restite following extraction of G1 and/or G3 magmas; they do

  6. Carboniferous-Permian rugose coral Cyathaxonia faunas in China


    The Cyathaxonia faunas are widely distributed in the Carboniferous and Permian strata in China.The fauna in China contains 12 families and 40 genera,and can be recognized as three episodes from Early Carboniferous to the end-Permian in terms of their occurrence and composition:1) Mississippian;2) Pennsylvanian-Early Permian;3) Middle Permian-Late Permian.They were relatively abundant in episodes 1 and 2.A decrease of family Antiphyllidae and an increase of family Hapsiphyllidae can be observed from Early Carboniferous to the end-Permian.Two case studies are given to illustrate the bio-facies of the Cyathaxonia faunas,which are from the Mississippian of Baoshan,West Yunnan and the Mississippian of Chaohu,South Anhui.In both areas,Cyathaxonia faunas occur just below the large dissepimented solitary and compound coral assemblages in a continuous sequence in a single section,which implies that the occurrence of these small,non-dissepimented solitary corals is not strictly related with Gondwanan or Peri-gondwanan cold water environment,but might be controlled by such benthic conditions as deeper,mud-rich,quieter,and darker sedimentary environments.

  7. 柴达木盆地及其邻区早—中二叠世构造-岩相古地理格局%Early-middle Permian Tectonic-lithofacies Palaeogeographic Characteristics in Qaidam Basin and Its Adjacent Areas

    陈守建; 陈奋宁; 计文化; 李荣社; 洛长义; 刘荣利; 李国栋


    通过大量野外观察、分析测试和综合研究,结合覆盖全区不同比例尺的区域地质调查资料,对柴达木盆地及其邻区早—中二叠世构造-岩相古地理格局进行了研究,并探讨了其形成机制.结果表明:柴达木盆地及其邻区中、北部早—中二叠世为陆内盆山-裂谷系统,主体处于伸展构造背景,总体表现为堑垒相间的构造古地理格局;南部为巴颜喀拉洋盆,早—中二叠世是巴颜喀拉洋盆扩张最为剧烈且规模最大的时期,巴颜喀拉洋盆中洋岛、海山遍布;早—中二叠世晚期强烈的华力西构造运动使古板块间的相对运动加剧,巴颜喀拉洋盆大洋岩石圈板块向北强烈俯冲,形成东昆仑陆缘岩浆弧及其南部增生带,东昆仑陆缘岩浆弧岩浆活动十分强烈,宗务隆山—西秦岭陆源裂谷盆地进一步发育,形成十分复杂的构造-岩相古地理格局.早—中二叠世是研究区地球动力学机制从强烈扩张到强烈汇聚的转折时期,加强其构造-岩相古地理研究对建立该区地层序列、探讨其地质演化历史以及指导找矿等均具有重要意义.%Through the field observation, the analysis of test data, the comprehensive regional geological survey and the previous research result, the forming mechanism and tectonic-lithofacies paleogeography during the Early-middle Permian in Qaidam Basin and its adjacent areas were discussed. The results showed that the Early-middle Permian in the central and northern parts of Qaidam Basin and its adjacent areas was intracontinental basin-and-range and rift valley system, and mainly located on extensional tectonic setting, and the tectonic paleogeography was graben and horst alternately! the Early-middle Permian in the southern part was the period with most violent expansion cosmically in Bayankala ocean basin, and ocean-island and seamount were all over the ocean basinj the strong Hualixi tectonic movement in the late of Early

  8. Palaeotopography-governed sediment distribution—a new predictive model for the Permian Upper Rotliegend in the Dutch sector of the Southern Permian Basin

    Mijnlieff, H.F.; Geluk, M.


    The Southern Permian Basin (SPB) formed a large, elongated saucer-shaped inland depression extending from the UK to Poland and from Belgium to Denmark. During the Middle to early Late Permian the SPB was filled progressively by playa sediments from its centre in northwestern Germany before it was fl

  9. Palaeotopography-governed sediment distribution—a new predictive model for the Permian Upper Rotliegend in the Dutch sector of the Southern Permian Basin

    Mijnlieff, H.F.; Geluk, M.


    The Southern Permian Basin (SPB) formed a large, elongated saucer-shaped inland depression extending from the UK to Poland and from Belgium to Denmark. During the Middle to early Late Permian the SPB was filled progressively by playa sediments from its centre in northwestern Germany before it was fl

  10. Radiometric age determination of tonsteins and stratigraphic constraints for the Lower Permian coal succession in southern Parana Basin, Brazil

    Guerra-Sommer, Margot; Cazzulo-Klepzig, Miriam; Hartmann, Leo Afraneo; Formoso, Milton Luis Laquintinie [Instituto de Geociencias, Universidade Federal do Rio Grande do Sul, Avenida Bento Goncalves, 9500, 91501-970 Porto Alegre, Rio Grande do Sul (Brazil); Santos, Joao Orestes Schneider [Centre for Global Targeting, University of Western Australia, Crawley, Perth 6009, WA (Australia); Ketzer, Joao Marcelo [Instituto de Meio Ambiente, Pontificia Universidade Catolica, Avenida Ipiranga, Porto Alegre, Rio Grande do Sul (Brazil)


    Ion microprobe (SHRIMP II) dating of zircons from tonsteins interbedded with coal seams from the Candiota and Faxinal coalfields (Early Permian, Rio Bonito Formation, Parana Basin, Brazil) is presented. The mean ages obtained (290.6 {+-} 1.5 Ma) are more precise than previously published intervals. Calibrations of chronostratigraphic data with radiometric ages show that the main coal succession from the southern Basin is constrained to the Middle Sakmarian. The {+-} 2 Ma time interval of deposition supports the hypothesis that the coal-generating process was quite rapid in terms of geological time. In a general context, Faxinal and Candiota coals are assigned, into the Protohaploxypinus goraiensis Subzone, besides some paleocological differences evidenced by palynological studies. This bio-interval does not correspond to a consistent palynostratigraphic tool and more accurate biostratigraphic zonation for the Carboniferous-Permian interval must be delineated. The new results have far-reaching significance for correlations of the Basin with sequences of the Argentinian Paganzo Basin (302 {+-} 6 Ma and 288 {+-} 7 Ma) and also with the Karoo Basin, with the lowermost Ecca Group (288 {+-} 3 Ma and 289.6 {+-} 3.8 Ma). This new evidence supports the presence of an active and widespread Lower Permian explosive volcanic event in western Gondwana, which is interpreted as the same volcanism which produced the Choiyoi Group in western Argentina. According to this correlation the ash-fall source is located about 1400 km to the southwest of their area of deposition. (author)

  11. Early Permian arc-related volcanism and sedimentation at the western margin of Gondwana:Insight from the Choiyoi Group lower section

    Leonardo Strazzere; Daniel A. Gregori; Leonardo Benedini


    Permian sedimentary and basic to intermediate volcanic rocks assigned to the Conglomerado del Río Blanco and Portezuelo del Cenizo Formation, lower part of the Choiyoi Group, crop out between the Cordon del Plata, Cordillera Frontal and Precordillera of Mendoza Province, Argentina. The sedimentary rocks are represented by six lithofacies grouped in three facies associations. They were deposited by mantled and gravitational flows modified by high-energy fluvial currents that evolved to low-energy fluvial and lacustrine environments. They constitute the Conglomerado del Río Blanco, which cover unconformably marine Carboniferous sequences. Five volcanic and volcaniclastic facies make up the beginning of volcanic activity. The first volcanic event in the Portezuelo del Cenizo is basaltic to andesitic lava-flows emplaced in the flanks of volcanoes. Lava collapse produced thick block and ash flows. Interbedding in the intermediate volcanic rocks, there are dacites of different geochemical signature, which indicate that the development of acidic volcanism was coetaneous with the first volcanic activity. The geochemistry of these rocks induces to consider that the Choiyoi Group Lower section belongs to a magmatic arc on continental crust. The age of this section is assigned to the lower Permian (277 ? 3.0 Ma, Kungurian age).

  12. Germanidade e banhos medicinais nos primórdios dos balneários no Rio Grande do Sul Germanism and medicinal bathing in the early days of health resorts in Rio Grande do Sul

    Sílvio Marcus de Souza Correa


    Full Text Available Nos primórdios dos balneários no Rio Grande do Sul, encontravam-se alguns imigrantes alemães, não apenas entre os banhistas, mas também entre os empresários do incipiente ramo do curismo-turismo. Era um grupo pequeno de imigrantes de origem urbana que, em geral, já conhecia as vantagens curativas ou revigorantes dos banhos em balneários europeus. Entre eles destacavam-se os médicos, importantes emissores de um discurso científico em prol dos balneários. As práticas terapêuticas de banhos de mar chegaram ao Brasil meridional pela imigração européia da segunda metade do século XIX, embora sua difusão só tenha ocorrido na primeira metade do século seguinte, quando se desenvolveram as primeiras praias balneárias no Rio Grande do Sul.In the early days of bathing resorts some German immigrants were found not only among the bathers, but also among the entrepreneurs of the incipient branch of 'curism-tourism'. It was a small group of immigrants of urban origin who, in general, already knew the curative or reinvigorating advantages of the baths in European bathing resorts. Among them, doctors were prominent, important emissaries of a scientific discourse in favor of bathing resorts. The therapeutic practices of bathing in the sea arrived to meridional Brazil with the European immigration of the second half of the nineteenth century, although its diffusion only took place in the first half of the following century, when the first bathing beaches in Rio Grande do Sul were developed.

  13. High sedimentation rates in the Early Triassic after latest Permian mass extinction: Carbonate production is main factor in non-Arctic regions

    Horacek, Micha; Brandner, Rainer


    A substantial change in sedimentation rates towards higher values has been documented from the Late Permian to the Lower Triassic. Although it is assumed and also has been shown that the deposition of siliciclastic material increased in the Lower Triassic due to stronger erosion because of loss of land cover and increased chemical and physical weathering with extreme climate warming, the main sediment production occurred by marine carbonate production. Still, carbonate production might have been significantly influenced by weathering and erosion in the hinterland, as the transport of dust by storms into the ocean water probably was a main nutrient source for microbial carbonate producers, because "normal" nutrient supply by ocean circulation, i. e. upwelling was strongly reduced due to the elevated temperatures resulting in water-column stratification . Sediment accumulation was also clearly influenced by the paleo-geographic and latitudinal position, with lower carbonate production and sedimentation rates in moderate latitudes. The existence of a "boundary clay" and microbial carbonate mounds and layers in the immediate aftermath of the latest Permian mass extinction points towards a development from a short-timed acid ocean water - resulting in a carbonate production gap and the deposition of the boundary clay towards the deposition of the microbial mounds and layers due to the microbial production of micro-environments with higher alkalinity allowing the production of carbonate. After the return of the ocean water to normal alkalinity planktic production of carbonate resulted in a very high sedimentation rate, especially taking into account the absence of carbonate producing eukaryotic algae and animals.

  14. Permian magmatism, Permian detachment faulting, and Alpine thrusting in the Orobic Anticline, southern Alps, Italy

    Pohl, Florian; Froitzheim, Niko; Geisler-Wierwille, Thorsten; Schlöder, Oliver


    The Grassi Detachment Fault is located in the Orobic Alps east of Lake Como and was described by Froitzheim et al. (2008) as an Early Permian extensional structure. Many issues still remained unclear, like the exact timing of faulting and the extension from the well-exposed part of the detachment towards west. The Grassi Detachment Fault separates the Variscan Basement in its footwall from the volcanic and sedimentary rocks of the Early Permian Collio Formation within its hanging wall, marked by a mylonitic and cataclastic layer whose textures indicate top-to-the-southeast displacement. The footwall basement is formed by the Variscan Morbegno Gneiss and two granitic intrusions, the Val Biandino Quarz Diorite (VBQD) and the Valle Biagio Granite (VBG). The former is syntectonic with respect to the detachment, whereas for the latter, the relation to the detachment is unknown. The age of the VBQD is poorly defined as 312 Ma ± 48 Ma (Thöni et al. 1992); the VBG has not been dated. Volcanic rocks of the Collio Formation in the hanging wall may represent the extrusive part of the magmatic system. In our study area west of Val Biandino, several faults and shear zones are exposed: (1) The Grassi Detachment Fault is represented by mylonites and cataclasites with top-SE shear sense, between basement rocks and the Collio Volcanics. Towards NW, it is truncated by the unconformably overlying Late Permian Verrucano Lombardo. This may reflect the eroded culmination of a Permian metamorphic core complex. (2) A steeply NW-dipping, brittle normal fault is found further west in the footwall between VBQD and VBG. It is sealed by the basal unconformity of the Verrucano Lombardo and therefore should also be of Early Permian age (Sciunnach, 2001). It may represent an antithetic fault with respect to the detachment, accommodating the uplift of the magmatically inflated core complex. (3) The Biandino Fault is a steeply SE-dipping reverse fault, affecting also the Late Permian Verrucano

  15. Interpretation of "fungal spikes" in Permian-Triassic boundary sections

    Hochuli, Peter A.


    Abundant occurrences of the palynomorph Reduviasporonites have been described as ;fungal spike; from several Permian/Triassic boundary sections and related to the supposed destruction of woody vegetation by fungal pathogens during the Permian/Triassic extinction event. The biological affinity of this taxa considered by some authors of fungal origin is still controversially discussed since there is geochemical evidence that it is most probably related to algae. The abundance peak of this species is used by some authors as a stratigraphic marker, notably in terrestrial Permian/Triassic boundary sections from South China. Illustrations of the reported fungal remains however show potentially erroneous taxonomic identification of Reduviasporonites, and, based on differences in thermal maturation, they may represent recent contamination. Here Reduviasporonites chalastus of Early Triassic age is illustrated together with recent fungal remains originating from a strongly weathered and otherwise barren sample from a Middle Triassic section.

  16. Permian brachiopods from new localities in northeast Thailand: Implications for paleobiogeographic analysis

    Pérez-Huerta, Alberto; Chonglakmani, Chongpan; Chitnarin, Anisong


    A small Permian brachiopod fauna is described from new localities in northeastern Thailand. Brachiopods were collected from early Permian (Asselian) limestones of the Nam Maholan Formation and middle Permian (Murgabian) sandstones of the Nam Duk Formation and limestones of the Khao Khwang Formation. Analyses of taxa confirm preliminary hypotheses of Cathaysian affinities for brachiopods and fusulinids found in this part of Thailand. Fossils found in sandstones of the Nam Duk molasse facies, however, also show possible Gondwanan relationships with brachiopod taxa described in Australia. This has to be further tested with ongoing research in a better understanding of the paleobiogeography of this part of Southeast Asia.

  17. Permian biogeography of the Indian subcontinent with special reference to the marine fauna

    Singh, Trilochan

    Permian biogeography of the Indian subcontinent is discussed in the light of brachiopods and associated fossils from different localities. The discussion is based primarily on the Permian "biome" concept of Waterhouse and Bonham-Carter (1975), wherein three biomes are proposed: group A of subpolar, group B of temperate, and group C of tropical character. Data on the occurrence of Permian brachiopods and associated fossils are given for the Salt Range, Karakoram, and Himalayan regions of India, Nepal, Bhutan, Tibet, and Peninsular India with respect to the age of the fauna. Marine Permian localities of the Himalayan region include those of Ladakh, Zanskar, Lahaul and Spiti, Kashmir, Bhadarwah-Bhallesh-Chamba, Kinnaur, Garhwal, Kumaun, Darjeeling, Sikkim, and Arunachal Pradesh. Permian marine localities of Peninsular India, which forms a part of central Gondwanaland, include those of Bap, Badhaura, Umaria, Manendragarh, and Daltonganj, where marine transgression occurred in Early Permian time. The faunas of these localities are discussed with respect to their age, which falls into two groups, Early and Late Permian. It is suggested that widespread colder climatic conditions prevailed in the Indian subcontinent during the early Early Permian. Similar conditions continued in most of the localities until the late Early Permian, except at west Karakoram (Shaksgam valley), Zanskar, north Tibet (central and western part), and the Salt Range. However, during the Late Permian, climatic conditions were varied. Cold climatic conditions prevailed in north Tibet (central part), Kumaun Tethyan Himalaya, and south Tibet; temperate conditions occurred in west Karakoram (Shaksgam valley), Zanskar, Lahaul and Spiti, Bhadarwah-Bhallesh-Chamba, north Nepal, and north Sikkim; and tropical conditions occurred in the Salt Range, east Karakoram, Ladakh, Kashmir, and north Tibet (western and eastern parts). At a few localities there appear to be some anomalies that might be due to lack of

  18. Extinction pattern of reef ecosystems in latest Permian

    WU YaSheng; FAN JiaSong; JIANG HongXia; YANG Wan


    Studies of two Permian-Triassic boundary (PTB) sections on top of a Changhsingian reef in Ziyun, Guizhou Province, southwestern China indicate that the end-Permian mass extinction of reef ecosystems occurred in two steps. The first step is the extinction of all stenotropic organisms such as calcisponges and fusulinids in the latest Permian (in the Clarkina yini conodont zone). The biota after the first extinction is simple, comprising eurytropic organisms including microgastropods, ostracods, and some small burrowing organisms, or only algal mats. At the beginning of the Early Triassic (I.e. the beginning of the Hindeodus parvus zone), the environments became anoxic, and the microgastropod dominated biota or algal mats disappeared, which constituted the second episode of the mass extinction. The biota after the second extinction comprises small spherical microproblematica, some kinds of specialized organisms tolerant of anoxic or oxygen-poor conditions. As the environments became oxygenated, the specialized biota was replaced by a microgastropod-dominated simple biota. When the environmental conditions improved further, the simple biota was replaced by a diverse biota with normal-sized ammonoids, bivalves, and gastropods, representing restoration of normal oceanic conditions. Comparison with PTB sections in Dolomites, Italy and Meishan, Zhejiang Province shows that non-reef ecosystems had a similar first episode of mass extinction in the latest Permian. In the case that oceanic anoxia happened, non-reef ecosystems had a second extinction episode similar to that of reef ecosystems.

  19. Early Evidence for Zika Virus Circulation among Aedes aegypti Mosquitoes, Rio de Janeiro, Brazil

    Ayllón, Tania; Campos, Renata de Mendonça; Brasil, Patrícia; Morone, Fernanda Cristina; Câmara, Daniel Cardoso Portela; Meira, Guilherme Louzada Silva; Tannich, Egbert; Yamamoto, Kristie Aimi; Carvalho, Marilia Sá; Pedro, Renata Saraiva; Cadar, Daniel; Ferreira, Davis Fernandes; Honório, Nildimar Alves


    During 2014–2016, we conducted mosquito-based Zika virus surveillance in Rio de Janeiro, Brazil. Results suggest that Zika virus was probably introduced into the area during May–November 2013 via multiple in-country sources. Furthermore, our results strengthen the hypothesis that Zika virus in the Americas originated in Brazil during October 2012–May 2013. PMID:28628464

  20. Geochemistry of Sedimentation and Diagenesis in Qixia Formation (Early Permian) of Badong,Hubei Province, China: Implications for T-R Cycle Interpretation


    Combined with sedimentary observation and mineralogical research on a type of drusy celestite of early diagenesis in origin, this geochemical study on the Qixia Formation at the Shuibuya Section, Badong County, Hubei Province of south China evaluates the contribution of sedimentation and diagenesis to the mass fractions of CaO, Al2O3, MgO, SiO2, Fe2O3, Na2O, K2O, Mn, Sr, P and Ba in the carbonates. The Sr, initially released from the stabilization of carbonate calcium mineral, precipitated with sulfate ion as celestite in early diagenetic environment, where sulfate reduction was the minimum. Then it redistributed in burial diagenetic environment, where celestite was replaced by calcite. The fractions of the MgO and SiO2 in the carbonates were mainly modified by the early dolomitization and silicification respectively. That of the Na2O was overprinted by the burial diagenesis. Multivariate statistical analysis on data of sixty bulk rocks indicates that the mass fractions of the Al2O3, K2O, Fe2O3, Mn, CaO, P and Ba were affiliated with the sedimentary factor despite the obvious decrease of CaO during early dolomitization and silicification. Among them, the Al2O3 and K2O mainly represent the detrital components. In addition, a dysaerobic sedimentary background was confirmed by trace element measurement and information derived from the formation of the celestite. This genetic discrimination facilitates the interpretation of the transgressive-regressive (T-R) cycle and specialities featured this unusual carbonate unit.

  1. U/Pb zircon geochronology and tempo of the end-permian mass extinction

    Bowring; Erwin; Jin M W Martin YG; Davidek; Wang


    The mass extinction at the end of the Permian was the most profound in the history of life. Fundamental to understanding its cause is determining the tempo and duration of the extinction. Uranium/lead zircon data from Late Permian and Early Triassic rocks from south China place the Permian-Triassic boundary at 251.4 +/- 0.3 million years ago. Biostratigraphic controls from strata intercalated with ash beds below the boundary indicate that the Changhsingian pulse of the end-Permian extinction, corresponding to the disappearance of about 85 percent of marine species, lasted less than 1 million years. At Meishan, a negative excursion in delta13C at the boundary had a duration of 165,000 years or less, suggesting a catastrophic addition of light carbon.

  2. Hydrous parental magmas of Early to Middle Permian gabbroic intrusions in western Inner Mongolia, North China: New constraints on deep-Earth fluid cycling in the Central Asian Orogenic Belt

    Pang, Chong-Jin; Wang, Xuan-Ce; Xu, Bei; Luo, Zhi-Wen; Liu, Yi-Zhi


    The role of fluids in the formation of the Permian-aged Xigedan and Mandula gabbroic intrusions in western Inner Mongolia was significant to the evolution of the Xing'an Mongolia Orogenic Belt (XMOB), and the active northern margin of the North China Craton (NCC). Secondary Ion Mass Spectroscopy (SIMS) U-Pb zircon geochronology establishes that the Xigedan gabbroic intrusion in the northern NCC was emplaced at 266 Ma, and is therefore slightly younger than the ca 280 Ma Mandula gabbroic intrusion in the XMOB. Along with their felsic counterparts, the mafic igneous intrusions record extensive bimodal magmatism along the northern NCC and in the XMOB during the Early to Middle Permian. The Mandula gabbroic rocks have low initial 87Sr/86Sr ratios (0.7040-0.7043) and positive εNd(t) (+6.2 to +7.3) and εHf(t) values (+13.4 to +14.5), resembling to those of contemporaneous Mandula basalts. These features, together with the presence of amphibole and the enrichment of large ion lithophile elements (LILE, e.g., Rb, Ba, U and Sr) and depletion of Nb-Ta suggest that the parental magmas of the Mandula mafic igneous rocks were derived from a depleted mantle source metasomatized by water-rich fluids. In contrast, the Xigedan gabbroic rocks are characterised by high 87Sr/86Sr ratios (0.7078-0.7080) and zircon δ18O values (5.84-6.61‰), but low εNd(t) (-9.3 to -10.2) and εHf(t) values (-8.76 to -8.54), indicative of a long-term enriched subcontinental lithosphere mantle source that was metasomatized by recycled, high δ18O crustal materials prior to partial melting. The high water contents (4.6-6.9 wt%) and arc-like geochemical signature (enrichment of fluid-mobile elements and depletion of Nb-Ta) of the parental magmas of the Xigedan gabbroic rocks further establish the existence of a mantle hydration event caused by fluid/melts released from hydrated recycled oceanic crust. Incompatible element modelling shows that 5-10% partial melting of an enriched mantle source by

  3. Retention in Early Care at an HIV Outpatient Clinic in Rio de Janeiro, Brazil, 2000-2013.

    Silva, Daniel S; De Boni, Raquel B; Lake, Jordan E; Cardoso, Sandra W; Ribeiro, Sayonara; Moreira, Ronaldo I; Clark, Jesse L; Veloso, Valdilea G; Grinsztejn, Beatriz; Luz, Paula M


    Retention in early HIV care has been associated with virologic suppression and improved survival, but remains understudied in Brazil. We estimated retention in early HIV care for the period 2000-2013, and identified socio-demographic and clinical factors associated with good retention in an urban cohort from Rio de Janeiro, Brazil. Antiretroviral therapy-naïve, HIV-infected persons ≥18 years old linked to care between 2000 and 2011 were included. Retention in the first 2 years post-linkage (i.e. early care) was defined by the proportion of 6-month intervals with ≥1 HIV laboratory result. "Good" retention was defined as ≥1 HIV laboratory result recorded in at least three intervals. Overall, 80 % of participants met criteria for good retention and retention significantly improved over the study period. Older age, higher education level and early antiretroviral therapy initiation were associated with good retention. Efforts to improve retention in early care in this population should target younger and less-educated HIV-infected persons.

  4. Feasibility study of heavy oil recovery in the Permian Basin (Texas and New Mexico)

    Olsen, D.K.; Johnson, W.I.


    This report is one of a series of publications assessing the feasibility of increasing domestic heavy oil production. Each report covers select areas of the United States. The Permian Basin of West Texas and Southeastern New Mexico is made up of the Midland, Delaware, Val Verde, and Kerr Basins; the Northwestern, Eastern, and Southern shelves; the Central Basin Platform, and the Sheffield Channel. The present day Permian Basin was one sedimentary basin until uplift and subsidence occurred during Pennsylvanian and early Permian Age to create the configuration of the basins, shelves, and platform of today. The basin has been a major light oil producing area served by an extensive pipeline network connected to refineries designed to process light sweet and limited sour crude oil. Limited resources of heavy oil (10`` to 20`` API gravity) occurs in both carbonate and sandstone reservoirs of Permian and Cretaceous Age. The largest cumulative heavy oil production comes from fluvial sandstones of the Cretaceous Trinity Group. Permian heavy oil is principally paraffinic and thus commands a higher price than asphaltic California heavy oil. Heavy oil in deeper reservoirs has solution gas and low viscosity and thus can be produced by primary and by waterflooding. Because of the nature of the resource, the Permian Basin should not be considered a major heavy oil producing area.

  5. Feasibility study of heavy oil recovery in the Permian Basin (Texas and New Mexico)

    Olsen, D.K.; Johnson, W.I.


    This report is one of a series of publications assessing the feasibility of increasing domestic heavy oil production. Each report covers select areas of the United States. The Permian Basin of West Texas and Southeastern New Mexico is made up of the Midland, Delaware, Val Verde, and Kerr Basins; the Northwestern, Eastern, and Southern shelves; the Central Basin Platform, and the Sheffield Channel. The present day Permian Basin was one sedimentary basin until uplift and subsidence occurred during Pennsylvanian and early Permian Age to create the configuration of the basins, shelves, and platform of today. The basin has been a major light oil producing area served by an extensive pipeline network connected to refineries designed to process light sweet and limited sour crude oil. Limited resources of heavy oil (10'' to 20'' API gravity) occurs in both carbonate and sandstone reservoirs of Permian and Cretaceous Age. The largest cumulative heavy oil production comes from fluvial sandstones of the Cretaceous Trinity Group. Permian heavy oil is principally paraffinic and thus commands a higher price than asphaltic California heavy oil. Heavy oil in deeper reservoirs has solution gas and low viscosity and thus can be produced by primary and by waterflooding. Because of the nature of the resource, the Permian Basin should not be considered a major heavy oil producing area.

  6. The relationship of the Indian and western Australian Permian marine faunas

    Dickins, J. M.; Shah, S. C.

    At Gondwana Five, Wellington, New Zealand, 1980, the relationship of the Permian Peninsular and Himalayan India fauna and flora was explored. The authors concluded that Peninsular India could not have been far from southern Asia during the Permian. This conclusion has been confirmed by data presented at the Symposium on the Tibet Plateau, Beijing, China, 1980, and the recent discovery of Eurydesma and the Eurydesma fauna in northwest Tibet. The relationship of the marine faunas of Peninsular and Himalayan India with those of the western Australian region and the climatic implications have now been considered by the authors. Although the Lower Permian Gondwana cold- and cold-temperate-water faunas of both areas have important similarities, significant differences can be tabulated. Such differences are not apparent in the faunas from within the Peninsular-Himalayan Indian region, so that the faunas of this region are closer to each other than they are to the faunas of the western Australian region. In India the early faunas are overlain by warm-water faunas, whereas in the western Australian region temperate-water conditions prevail. Only in the Upper Permian, when the world as a whole became warmer, are closer faunal links established between the two areas. It is concluded that Western Australia was unlikely to have been close to India during the Permian, and that the faunal relationships indicate India cannot be placed alongside Western Australia. This conclusion is supported by the different geological development in the two areas during the Permian.

  7. Permian geology of Gondwana countries: An overview

    Dickins, J.M. (Australian Bureau of Mineral Resources, Canberra (Australia))


    Earliest Permian sequences of Antarctica, southern and east-central Africa, the southern part of the Arabian Peninsula, Pakistan, peninsular and Himalayan India, Tibet, western and eastern Australia, New Zealand, and South America are all characterized by glacial deposits and cold-water marine faunas. In the course of the Permian, considerable faunal (and floral) and climatic divergence occurred. Although folding is not necessarily present, the effects of the strong compressive tectonic phase (Hunter-Bowen Orogenic Folding Phase of Dickins) beginning in the mid-Permian (traditional two-fold subdivision) and of acidic and intermediate volcano-magmatic activity are apparent in all these regions as in other parts of the world. The progressive continentality of the Upper Permian (worldwide regression) culminates at the Permian-Triassic (Changxingian-Griesbachian) boundary.




    基于一些业已报道的Kungurian阶与Roadian阶界线附近的菊石材料进行重新研究,认为中国中、下二叠统界线附近的菊石生物群主要分布在华南和中国西部两个地区.通过与俄罗斯地区同时代的菊石动物群比较后认为中国在这一界线附近的菊石群以地方性的分子为主,主要包括:假海罗菊石亚科(Pseudohaloritinae)和寿昌菊石亚科(Shouchangoceratinae).通过再研究,修订多个菊石动物群的组成,并认为双堡塘组下部的时代为空谷阶,中上部为罗德阶,范家屯组的时代为罗德阶或沃德阶,西藏日喀则的加拉组和堆龙德庆的乌鲁龙组为罗德阶.通过菊石动物群研究认为中国最有潜力解决Kungurian阶和Roadian阶界线的剖面为甘肃的双堡塘剖面和西藏加拉剖面.%The Early-Middle Permian (Cisuralian-Guada-lupian) boundary is marked globally by tremendous changes in terms of the development of am-monoid faunas. At that time the new Order Cera-titida appeared and their first known representatives are found in the lowest part of the Middle Permian Roadian Stage. The earliest Ceratitida therefore serve as reliable markers for the boundary between the Lower and Middle Permian in the Tethyan region and North America in terms of am-monoids. In other biogeographic regions, however, where Ceratitida have not been found, representatives of the families Pseudogastrioceratidae and Spirolegoceratidae play an important role in biostratigraphical subdivision. The GSSP (Global Stratotype Section and Point) between the Lower and Middle Permian is defined in the Glass Mountains of Texas based on the first appearance of the conodonts species Jinogondolella nanking-ensis. However, the first Middle Permian representatives ammonoids appear slightly below this level in the Texas section. As the Roadian Stage was originally defined by Furnish based on am-monoid data. Many ammonoid experts do not agree with a definition based on conodonts in

  9. Palynology of Permian Gondwana sequence of Umrer coalfield, Maharashtra

    Jha, N.; Tewari, R.; Rajanikanth, A. [Birbal Sahni Institute of Paleobotany, Lucknow (India)


    Quantitative and qualitative palynological analysis of Early Permian sediments, Umrer Coalfield, Wardha Basin has been carried out. The microspore assemblage consists of 22 genera and 40 species. It is characterized by dominance of radial monosaccates chiefly, Parasacciles and subdominance of non-striate disaccates chiefly, Scheuringipollenites. Presence of Crucisaccites and Caheniasaccites suggests Upper Karharbari (Late Sakmarian-Early Artinskian) affinity. The presence of the Karharbari palynozone has been demarcated in lithologically designated Barakar Formation. The present finding corroborates the earlier studies by Bharadwaj and Anand Prakash (1974).

  10. Large perturbations of the carbon cycle during recovery from the end-permian extinction.

    Payne, Jonathan L; Lehrmann, Daniel J; Wei, Jiayong; Orchard, Michael J; Schrag, Daniel P; Knoll, Andrew H


    High-resolution carbon isotope measurements of multiple stratigraphic sections in south China demonstrate that the pronounced carbon isotopic excursion at the Permian-Triassic boundary was not an isolated event but the first in a series of large fluctuations that continued throughout the Early Triassic before ending abruptly early in the Middle Triassic. The unusual behavior of the carbon cycle coincides with the delayed recovery from end-Permian extinction recorded by fossils, suggesting a direct relationship between Earth system function and biological rediversification in the aftermath of Earth's most devastating mass extinction.




    Full Text Available Additional findings of fusulinids and corals from Permian rocks of Western Karakorum and Eastern Hindu Kush are reported. The Lashkargaz Fm. of Western Karakorum (Sakmarian - Kubergandian in age, is redefined in its uppermost part. A new unit of sucrosic dolostone, the Ini Sar Fm., of early Murgabian age in its basal part, is established for a level previously attributed to the Ailak Fm. The Ailak Fm. is confirmed to be Late Permian in its lower part. A rich fusulinid fauna of early Kubergandian age was found in a carbonate nappe previously unidentified, along the Pakistan-Afghanistan border in the Eastern Hindu Kush. 

  12. Tectonic implications of post-folding Permian magnetizations in the Carapacha Basin, La Pampa province, Argentina

    Tomezzoli, Renata N.; Melchor, Ricardo N.; MacDonald, William D.


    Paleomagnetic results from Permian clastic and igneous rocks of the Carapacha Basin of the Gondwáides orogenic zone of central Argentina are mainly consistent with results reported previously from the same zone further east, e.g., in the Sierra de la Ventana. Three lithologic entities were analysed: the lower member and upper member of the Carapacha Formation, and an andesite intrusive into the upper member. The directions of their characteristic remanences are similar, differ significantly from the present field direction, and are post-folding. The in situ magnetization directions are moreover consistent with directions expected for late Permian poles of the APWP for South America. The in situ pole for the lower Carapacha Formation is 70°S, 049°E, A95=11° (San Roberto pole); the pole for the upper Carapacha, combined with similar directions from the intrusive, is 64°S, 005°E, A95=5° (Río Curacó pole). These magnetizations imply that the Carapacha Formation, with a minimum age of early Late Permian (about 260 Ma.), was deformed before the end of the Permian. Structural evidence, as well as paleomagnetic, IRM, and AMS experimental results, support the interpretations. The paleogeographic implications of these results are interpreted as a significant counterclockwise movement of Gondwana between the early and the late Permian.

  13. Permian and Triassic microfloral assemblages from the Blue Nile Basin, central Ethiopia

    Dawit, Enkurie L.


    Palynological investigation was carried out on surface samples from up to 400 m thick continental siliciclastic sediments, here referred to as “Fincha Sandstone”, in the Blue Nile Basin, central Ethiopia. One hundred sixty species were identified from 15 productive samples collected along a continuous road-cut exposure. Six informal palynological assemblage zones have been identified. These assemblage zones, in ascending order, are: “Central Ethiopian Permian Assemblage Zone - CEPAZ I”, earliest Permian (Asselian-Sakmarian); “CEPAZ II”, late Early Permian (Artinskian-Kungurian); CEPAZ III - Late Permian (Kazanian-Tatarian); “CETAZ IV”, Lower Triassic (Olenekian Induan); “CETAZ V”, Middle Triassic (Anisian Ladinian); “CETAZ VI”, Late Triassic (Carnian Norian). Tentative age ranges proposed herein are compared with faunally calibrated palynological zones in Gondwana. The overall composition and vertical distribution of miospores throughout the studied section reveals a wide variation both qualitatively and quantitatively. The high frequency of monosaccate pollen in CEPAZ I may reflect a Glossopterid-dominated upland flora in the earliest Permian. The succeeding zone is dominated by straite/taeniate disaccate pollen and polyplicates, suggesting a notable increase in diversity of glossopterids. The decline in the diversity of taeniate disaccate pollen and the concomitant rise in abundance of non-taeniate disaccates in CEPAZ III may suggest the decline in Glossopteris diversity, though no additional evidence is available to equate this change with End-Permian extinction. More diverse and dominant non-taeniate, disaccate, seed fern pollen assignable to FalcisporitesAlisporites in CETAZ IV may represent an earliest Triassic recovery flora. The introduction of new disaccate forms with thick, rigid sacci, such as Staurosaccites and Cuneatisporites, in CETAZ V and VI may indicate the emergence of new gymnospermous plants that might have favourably

  14. Photic Zone Euxinia During the Permian-Triassic Superanoxic Event

    Grice, Kliti; Cao, Changqun; Love, Gordon D.; Böttcher, Michael E.; Twitchett, Richard J.; Grosjean, Emmanuelle; Summons, Roger E.; Turgeon, Steven C.; Dunning, William; Jin, Yugan


    Carbon and sulfur isotopic data, together with biomarker and iron speciation analyses of the Hovea-3 core that was drilled in the Perth Basin, Western Australia, indicate that euxinic conditions prevailed in the paleowater column during the Permian-Triassic superanoxic event. Biomarkers diagnostic for anoxygenic photosynthesis by Chlorobiaceae are particularly abundant at the boundary and into the Early Triassic. Similar conditions prevailed in the contemporaneous seas off South China. Our evidence for widespread photic-zone euxinic conditions suggests that sulfide toxicity was a driver of the extinction and a factor in the protracted recovery.

  15. Silicified wood from the Permian and Triassic of Antarctica: Tree rings from polar paleolatitudes

    Ryberg, P.E.; Taylor, E.L.


    The mass extinction at the Permian-Triassic boundary produced a floral turnover in Gondwana in which Paleozoic seed ferns belonging to the Glossopteridales were replaced by corystosperm seed ferns and other seed plant groups in the Mesozoic. Secondary growth (wood production) in both plant groups provides information on plant growth in relation to environment in the form of permineralized tree rings. Techniques utilized to analyze extant wood can be used on fossil specimens to better understand the climate from both of these periods. Late Permian and early Middle Triassic tree rings from the Beardmore Glacier area indicate an environment where extensive plant growth occurred at polar latitudes (~80–85°S, Permian; ~75°S, Triassic). A rapid transition to dormancy in both the Permian and Triassic woods suggests a strong influence of the annual light/dark cycle within the Antarctic Circle on ring production. Latewood production in each ring was most likely triggered by the movement of the already low-angled sun below the horizon. The plants which produced the wood have been reconstructed as seasonally deciduous, based on structural and sedimentologic evidence. Although the Late Permian climate has been reconstructed as cold temperate and the Middle Triassic as a greenhouse, these differences are not reflected in tree ring anatomy or wood production in these plant fossils from the central Transantarctic Mountains.

  16. From source-to-sink: The Late Permian SW Gondwana paleogeography and sedimentary dispersion unraveled by a multi-proxy analysis

    Alessandretti, Luciano; Machado, Rômulo; Warren, Lucas Veríssimo; Assine, Mario Luis; Lana, Cristiano


    The Late Permian sedimentary succession of the Paraná Basin, southern Brazil, provide a valuable source of information about sediment provenance, tectonic processes and, consequently, the paleogeography of the southwestern Gondwana supercontinent. In order to understand the patterns of sedimentary dispersal and reconstruct the Late Permian source-to-sink dynamic, we report a complete series of U-Pb ages and Hf isotopic compositions of detrital zircons from the Rio do Rasto Formation sandstones allied with detailed paleocurrent and sedimentologic data. Our integrated provenance study reveals a consistent sediment transport from the south to the north and northwest. According to the evaluation of zircon ages and Hf isotopes, it was possible to determine four distinct source areas: (i) a distant Late Paleozoic active magmatic arc located in the southwestern Gondwana margin (i.e. Gondwanides Orogen), corresponding to the North Patagonian Massif; (ii) recycling of orthoquartzites from the uplifted Paleozoic Ventania Fold Belt and immature sandstones from the Claromecó Foreland Basin in central-eastern Argentina and the Silurian-Devonian successions of the southern Paraná Basin (central-northern Uruguay) and North Patagonian Massif; (iii) exhumed areas of the Archean-Paleoproterozoic basement and Neoproterozoic to Early Paleozoic mobile belts of the Damara in southwestern Africa and Ribeira Fold Belt in Uruguay and southern Brazil; and (iv) southeastward provenance of Grenvillian (1.2-1.0 Ga) zircons coming from the mafic to intermediate Mesoproterozoic igneous units of the Namaqua-Natal Belt in South Africa and Namibia. These data allow us to argue that sediments deposited in the Paraná Basin during the Late Permian come from both short- and long-distance source areas. In this context, an important population of Permian detrital zircons comes from the Gondwanides Orogen in the south, probably carried by transcontinental alluvial systems. Close to the source area

  17. Permian-Triassic Osteichthyes (bony fishes): diversity dynamics and body size evolution.

    Romano, Carlo; Koot, Martha B; Kogan, Ilja; Brayard, Arnaud; Minikh, Alla V; Brinkmann, Winand; Bucher, Hugo; Kriwet, Jürgen


    The Permian and Triassic were key time intervals in the history of life on Earth. Both periods are marked by a series of biotic crises including the most catastrophic of such events, the end-Permian mass extinction, which eventually led to a major turnover from typical Palaeozoic faunas and floras to those that are emblematic for the Mesozoic and Cenozoic. Here we review patterns in Permian-Triassic bony fishes, a group whose evolutionary dynamics are understudied. Based on data from primary literature, we analyse changes in their taxonomic diversity and body size (as a proxy for trophic position) and explore their response to Permian-Triassic events. Diversity and body size are investigated separately for different groups of Osteichthyes (Dipnoi, Actinistia, 'Palaeopterygii', 'Subholostei', Holostei, Teleosteomorpha), within the marine and freshwater realms and on a global scale (total diversity) as well as across palaeolatitudinal belts. Diversity is also measured for different palaeogeographical provinces. Our results suggest a general trend from low osteichthyan diversity in the Permian to higher levels in the Triassic. Diversity dynamics in the Permian are marked by a decline in freshwater taxa during the Cisuralian. An extinction event during the end-Guadalupian crisis is not evident from our data, but 'palaeopterygians' experienced a significant body size increase across the Guadalupian-Lopingian boundary and these fishes upheld their position as large, top predators from the Late Permian to the Late Triassic. Elevated turnover rates are documented at the Permian-Triassic boundary, and two distinct diversification events are noted in the wake of this biotic crisis, a first one during the Early Triassic (dipnoans, actinistians, 'palaeopterygians', 'subholosteans') and a second one during the Middle Triassic ('subholosteans', neopterygians). The origination of new, small taxa predominantly among these groups during the Middle Triassic event caused a




    .... The Davaly Formation corresponds to the Bolorian Stage of the Lower Permian (Cisuralian); the Asni Formation corresponds to the Kubergandian Stage, and to the lower half of the Murgabian Stage of the Middle Permian (Guadalupian...

  19. Prolonged Permian Triassic ecological crisis recorded by molluscan dominance in Late Permian offshore assemblages.

    Clapham, Matthew E; Bottjer, David J


    The end-Permian mass extinction was the largest biotic crisis in the history of animal life, eliminating as many as 95% of all species and dramatically altering the ecological structure of marine communities. Although the causes of this pronounced ecosystem shift have been widely debated, the broad consensus based on inferences from global taxonomic diversity patterns suggests that the shift from abundant brachiopods to dominant molluscs was abrupt and largely driven by the catastrophic effects of the end-Permian mass extinction. Here we analyze relative abundance counts of >33,000 fossil individuals from 24 silicified Middle and Late Permian paleocommunities, documenting a substantial ecological shift to numerical dominance by molluscs in the Late Permian, before the major taxonomic shift at the end-Permian mass extinction. This ecological change was coincident with the development of fluctuating anoxic conditions in deep marine basins, suggesting that numerical dominance by more tolerant molluscs may have been driven by variably stressful environmental conditions. Recognition of substantial ecological deterioration in the Late Permian also implies that the end-Permian extinction was the climax of a protracted environmental crisis. Although the Late Permian shift to molluscan dominance was a pronounced ecological change, quantitative counts of 847 Carboniferous-Cretaceous collections from the Paleobiology Database indicate that it was only the first stage in a stepwise transition that culminated with the final shift to molluscan dominance in the Late Jurassic. Therefore, the ecological transition from brachiopods to bivalves was more protracted and complex than their simple Permian-Triassic switch in diversity.

  20. Aerobic Marine Habitat Loss During the Late Permian Extinction

    Penn, J. L.; Deutsch, C.; Payne, J.; Sperling, E. A.


    Rapid climate change at the end of the Permian is thought to have triggered the most severe mass extinction in Earth's history, but the precise mechanism of biodiversity loss is unknown. Geological evidence points to lethally hot equatorial temperatures and an expansion of anoxic ocean waters as likely culprits. However, previous climate model simulations of the warm Early Triassic exhibit weak tropical warming, and anoxic conditions require a massive and unconstrained increase in the ocean nutrient reservoir. Reconciling model predictions with the geologic record remains a key challenge to identifying the kill-mechanism, which must also take into account the role of animal physiology. Here we apply a recently developed index for the metabolic scope of marine animals to the first global climate simulations of the Permian-Triassic transition to quantify the effects of ocean warming and oxygen (O2) depletion on aerobic habitat availability. Forcing with extreme CO2 concentrations warms the surface ocean by over 10oC, consistent with paleoproxies for upper ocean temperature change. Warming depletes global O2, with greatest losses occuring in tropical deep waters as a result of their reduced ventilation. Together warming and deoxygenation would have constricted the occurrence of marine habitat by 80% globally, by decreasing the metabolic index of the Permian ocean. These changes are most pronounced in the tropics where the fossil record suggests recovery was severely inhibited. Fossil deposits also record changes in animal body size across the extinction. We find that adaptation via body size reductions can compensate for increasing hypoxia at high latitudes, and even prevent extinction there, but cannot maintain the habitability of the tropics.

  1. Terrestrial paleoenvironment characterization across the Permian-Triassic boundary in South China

    Bercovici, Antoine; Cui, Ying; Forel, Marie-Béatrice; Yu, Jianxin; Vajda, Vivi


    Well-preserved marine fossils in carbonate rocks permit detailed studies of the end-Permian extinction event in the marine realm. However, the rarity of fossils in terrestrial depositional environments makes it more challenging to attain a satisfactory degree of resolution to describe the biotic turnover on land. Here we present new sedimentological, paleontological and geochemical (X-ray fluorescence) analysis from the study of four terrestrial sections (Chahe, Zhejue, Mide and Jiucaichong) in Western Guizhou and Eastern Yunnan (Yangtze Platform, South China) to evaluate paleoenvironmental changes through the Permian-Triassic transition. Our results show major differences in the depositional environments between the Permian Xuanwei and the Triassic Kayitou formations with a change from fluvial-lacustrine to coastal marine settings. This change is associated with a drastic modification of the preservation mode of the fossil plants, from large compressions to small comminuted debris. Plant fossils spanning the Permian-Triassic boundary show the existence of two distinct assemblages: In the Xuanwei Formation, a Late Permian (Changhsingian) assemblage with characteristic Cathaysian wetland plants (mainly Gigantopteris dictyophylloides, Gigantonoclea guizhouensis, G. nicotianaefolia, G. plumosa, G. hallei, Lobatannularia heinanensis, L. cathaysiana, L. multifolia, Annularia pingloensis, A. shirakii, Paracalamites stenocostatus, Cordaites sp.) is identified. In the lowermost Kayitou Formation, an Early Triassic (Induan) Annalepis-Peltaspermum assemblage is shown, associated with very rare, relictual gigantopterids. Palynological samples are poor, and low yield samples show assemblages almost exclusively represented by spores. A ∼1 m thick zone enriched in putative fungal spores was identified near the top of the Xuanwei Formation, including diverse multicellular forms, such as Reduviasporonites sp. This interval likely corresponds to the PTB "fungal spike

  2. Marine anoxia and delayed Earth system recovery after the end-Permian extinction.

    Lau, Kimberly V; Maher, Kate; Altiner, Demir; Kelley, Brian M; Kump, Lee R; Lehrmann, Daniel J; Silva-Tamayo, Juan Carlos; Weaver, Karrie L; Yu, Meiyi; Payne, Jonathan L


    Delayed Earth system recovery following the end-Permian mass extinction is often attributed to severe ocean anoxia. However, the extent and duration of Early Triassic anoxia remains poorly constrained. Here we use paired records of uranium concentrations ([U]) and (238)U/(235)U isotopic compositions (δ(238)U) of Upper Permian-Upper Triassic marine limestones from China and Turkey to quantify variations in global seafloor redox conditions. We observe abrupt decreases in [U] and δ(238)U across the end-Permian extinction horizon, from ∼3 ppm and -0.15‰ to ∼0.3 ppm and -0.77‰, followed by a gradual return to preextinction values over the subsequent 5 million years. These trends imply a factor of 100 increase in the extent of seafloor anoxia and suggest the presence of a shallow oxygen minimum zone (OMZ) that inhibited the recovery of benthic animal diversity and marine ecosystem function. We hypothesize that in the Early Triassic oceans-characterized by prolonged shallow anoxia that may have impinged onto continental shelves-global biogeochemical cycles and marine ecosystem structure became more sensitive to variation in the position of the OMZ. Under this hypothesis, the Middle Triassic decline in bottom water anoxia, stabilization of biogeochemical cycles, and diversification of marine animals together reflect the development of a deeper and less extensive OMZ, which regulated Earth system recovery following the end-Permian catastrophe.

  3. Carnivorous dinocephalian from the Middle Permian of Brazil and tetrapod dispersal in Pangaea

    Cisneros, Juan Carlos; Abdala, Fernando; Atayman-Güven, Saniye; Rubidge, Bruce S.; Celâl Şengör, A. M.; Schultz, Cesar L.


    The medial Permian (∼270-260 Ma: Guadalupian) was a time of important tetrapod faunal changes, in particular reflecting a turnover from pelycosaurian- to therapsid-grade synapsids. Until now, most knowledge on tetrapod distribution during the medial Permian has come from fossils found in the South African Karoo and the Russian Platform, whereas other areas of Pangaea are still poorly known. We present evidence for the presence of a terrestrial carnivorous vertebrate from the Middle Permian of South America based on a complete skull. Pampaphoneus biccai gen. et sp. nov. was a dinocephalian "mammal-like reptile" member of the Anteosauridae, an early therapsid predator clade known only from the Middle Permian of Russia, Kazakhstan, China, and South Africa. The genus is characterized, among other features, by postorbital bosses, short, bulbous postcanines, and strongly recurved canines. Phylogenetic analysis indicates that the Brazilian dinocephalian occupies a middle position within the Anteosauridae, reinforcing the model of a global distribution for therapsids as early as the Guadalupian. The close phylogenetic relationship of the Brazilian species to dinocephalians from South Africa and the Russian Platform suggests a closer faunistic relationship between South America and eastern Europe than previously thought, lending support to a Pangaea B-type continental reconstruction.

  4. The Upper Permian in the Netherlands

    Visser, W.A.


    The Upper Permian in the Netherlands, as known from borehole data, is deposited in a mainly evaporitic facies north of the Brabant and Rhenish Massifs. In the extreme south (Belgian Campine, de Peel) a near-shore facies of reef dolomites and elastics occurs. In the western and central Netherlands th

  5. The Upper Permian in the Netherlands

    Visser, W.A.


    The Upper Permian in the Netherlands, as known from borehole data, is deposited in a mainly evaporitic facies north of the Brabant and Rhenish Massifs. In the extreme south (Belgian Campine, de Peel) a near-shore facies of reef dolomites and elastics occurs. In the western and central Netherlands

  6. Mechanism on bipolar distribution of Permian brachiopods

    Chengwen WANG; Songmei ZHANG


    By reasearch on geographic distribution, nine genera in bipolar distribution are selected from Permian brachiopods. These taxa originated from middle-high latitude areas in the boreal realm, of which five genera were derived from Late Carboniferous, and other four genera originated from Permian. They were all in bipolar distribution during some different stages in Permian. Specific diversity for each genus was high in the boreal realm, whereas in the Gondwana realm was very low. Perdurability was long in the boreal realm, and short in the Gondwana realm. It was the time when these nine genera came to their maximum diversity that these genera appeared in the Gondwana and formed bipolar distribution; while they also migrated to the low latitude from high latitude. This shows very close relationship between several main cooling events in Permian and the migration of genera from the boreal realm to the Gondwana realm through the Tethys. Therefore, the cooling events might be the main drive which caused these cold-water-type brachiopods migrated to the Gondwana realm and being bipolar distribution. In this process, the planula tolerance to warm water would be another important factor.

  7. Contrasting microbial community changes during mass extinctions at the Middle/Late Permian and Permian/Triassic boundaries

    Xie, Shucheng; Algeo, Thomas J.; Zhou, Wenfeng; Ruan, Xiaoyan; Luo, Genming; Huang, Junhua; Yan, Jiaxin


    Microbial communities are known to expand as a result of environmental deterioration during mass extinctions, but differences in microbial community changes between extinction events and their underlying causes have received little study to date. Here, we present a systematic investigation of microbial lipid biomarkers spanning ∼20 Myr (Middle Permian to Early Triassic) at Shangsi, South China, to contrast microbial changes associated with the Guadalupian-Lopingian boundary (GLB) and Permian-Triassic boundary (PTB) mass extinctions. High-resolution analysis of the PTB crisis interval reveals a distinct succession of microbial communities based on secular variation in moretanes, 2-methylhopanes, aryl isoprenoids, steranes, n-alkyl cyclohexanes, and other biomarkers. The first episode of the PTB mass extinction (ME1) was associated with increases in red algae and nitrogen-fixing bacteria along with evidence for enhanced wildfires and elevated soil erosion, whereas the second episode was associated with expansions of green sulfur bacteria, nitrogen-fixing bacteria, and acritarchs coinciding with climatic hyperwarming, ocean stratification, and seawater acidification. This pattern of microbial community change suggests that marine environmental deterioration was greater during the second extinction episode (ME2). The GLB shows more limited changes in microbial community composition and more limited environmental deterioration than the PTB, consistent with differences in species-level extinction rates (∼71% vs. 90%, respectively). Microbial biomarker records have the potential to refine our understanding of the nature of these crises and to provide insights concerning possible outcomes of present-day anthropogenic stresses on Earth's ecosystems.

  8. On a new stereospondylomorph temnospondyl from the Middle-Late Permian of Southern Brazil

    Adriana Strapasson


    Full Text Available A new temnospondyl is described from the Middle–Upper Permian sequence of the Paraná Basin (Rio do Rasto Formation in southern Brazil. The material consists of disarticulated cranial and postcranial elements, preserved in association. The cranial elements include part of the orbital region of the skull roof, the basicranium, a number of endocranial elements, stapes and a right hemimandible. The postcranial elements include vertebrae, ribs, pectoral girdle elements, a right femur and a cluster of scales. The new species displays a rhinesuchid pattern, which is similar to the South African rhinesuchids from the Upper Permian Beaufort Group of the Karoo Basin, but differs from them by the presence of a robust and elongated epipterygoid with a blade-like anterior process in addition to elongated and deeper muscular pockets on the parasphenoid, which allow the assignment of this specimen to a new species. However, the phylogenetic analysis grouped the material described herein and Australerpeton cosgriffi inside Stereospondylomorpha, in a transitional position between the Laurasian assemblages and South African temnospondyls. This result supports a connection between the Brazilian and Eastern European Permian fauna and provides important data for future biostratigraphic studies.

  9. Provincialization of terrestrial faunas following the end-Permian mass extinction.

    Sidor, Christian A; Vilhena, Daril A; Angielczyk, Kenneth D; Huttenlocker, Adam K; Nesbitt, Sterling J; Peecook, Brandon R; Steyer, J Sébastien; Smith, Roger M H; Tsuji, Linda A


    In addition to their devastating effects on global biodiversity, mass extinctions have had a long-term influence on the history of life by eliminating dominant lineages that suppressed ecological change. Here, we test whether the end-Permian mass extinction (252.3 Ma) affected the distribution of tetrapod faunas within the southern hemisphere and apply quantitative methods to analyze four components of biogeographic structure: connectedness, clustering, range size, and endemism. For all four components, we detected increased provincialism between our Permian and Triassic datasets. In southern Pangea, a more homogeneous and broadly distributed fauna in the Late Permian (Wuchiapingian, ∼257 Ma) was replaced by a provincial and biogeographically fragmented fauna by Middle Triassic times (Anisian, ∼242 Ma). Importantly in the Triassic, lower latitude basins in Tanzania and Zambia included dinosaur predecessors and other archosaurs unknown elsewhere. The recognition of heterogeneous tetrapod communities in the Triassic implies that the end-Permian mass extinction afforded ecologically marginalized lineages the ecospace to diversify, and that biotic controls (i.e., evolutionary incumbency) were fundamentally reset. Archosaurs, which began diversifying in the Early Triassic, were likely beneficiaries of this ecological release and remained dominant for much of the later Mesozoic.

  10. Permian continental paleoenvironments in Southeastern Asia: New insights from the Luang Prabang Basin (Laos)

    Bercovici, Antoine; Bourquin, Sylvie; Broutin, Jean; Steyer, Jean-Sébastien; Battail, Bernard; Véran, Monette; Vacant, Renaud; Khenthavong, Bounxou; Vongphamany, Sotsy


    In Laos (Southeastern Asia), Late Paleozoic sediments were identified by early French explorations across Indochina during the late 19th century (Pavie missions), but little work was undertaken to characterize the sedimentological and stratigraphical context until now. From detailed sedimentological and paleontological studies, we propose an interpretation of the depositional environment and of the stratigraphic context of series located on the right bank of the Mekong River in the Luang Prabang Basin where three main formations were described. The silicoclastic Red Claystone Formation, attributed to alluvial plain environment, contains large fragments of unidentified dicynodonts. The Limestones and Sandstones Formation preserves a new macrofloral assemblage displaying affinities with Middle to Late Permian Cathaysian floras of South China. This assemblage occurs as an intercalation within marine calcareous sandstones that have yielded a marine fauna, including the ammonoid Pseudotirolites sp. which indicates a Late Permian (Changhsingian) age. The well-developed Purple Claystones Formation yielded an abundant and well preserved Late Permian fauna composed of a carnivorous amphibian and numerous Dicynodon cranial and postcranial elements. This formation shows a vertical evolution from braided river to alluvial plain with sheet-flood sand bed and bed-load rivers, with a constant supply of volcanic clasts. Results from the analysis of the paleontological associations in the Luang Prabang Basin suggest that a continental communication between Laurussia and the Indochina Block existed during the Permian, allowing for migration of the terrestrial Dicynodon fauna.

  11. Geochemical characteristics and tectonic implications of the early Permian volcanic rocks from Ongniud Banner,Inner Mongolia%内蒙古翁牛特旗地区早二叠世火山岩地球化学特征及其构造意义

    曹代勇; 赵发; 刘登; 刘新华; 杨荣丰


    The Early Permian volcano-sedimentary sequences of Ongniud Banner consist mainly of andesite, rhyolite,perlite,volcanic breccia,tuff,tuffaceous sandstone,siliceous rock.Rock assemblage and sedimentary formations indicate that are of fore-arc basin sedimentary feature between subduction zone and island arc in Early Permian.The volcanic rocks from Elitu Formation have SiO2 =50.23%~73.56%,Mg#=6.21 ~49.54,Na2O+K2O=5.27% ~10.73%,Na2O/K2O =0.36 ~4.17,belonging to high-K cal-alkaline (HK CA) ~shoshonite (SHO)series.The volcanic rocks are characterized with(La/Yb)N =5.52 ~9.89,mid-intensive negative Eu abnormality,LILE enrichment such as Rb,K,La,Th and HFSE depletion such as P,Ti,Ta,Nb, and indicating that magma was formed in the tectonic setting of the island arc and active continental margin related to the plate subduction.R1-R2 diagram also indicates that volcanic rocks are generated at syn-collision or post-orogenic period,perhaps representing the mid-later subduction stage of the Palaeo-Asian Ocean Plate and North China Plate.Taken together,it suggests that the region located still in the Palaeo-Asian Ocean,rather than the intracontinental taphrogenic trough in Early Permian.%研究区早二叠世火山-沉积岩系主要由火山角砾岩、凝灰质砂岩、凝灰岩、硅质岩、流纹岩、安山岩、珍珠岩等组成.其沉积建造和岩性组合表明:早二叠世,研究区位于岛弧和俯冲带之间的弧前盆地环境.二叠系下统额里图组火山岩的SiO2=50.23%~73.56%,Mg#=6.21~49.54,Na2O+K2O=5.27%~10.73%,Na2O/K2O=0.36~4.17,其中安山岩Mg#=49.54~62.61,归属为高钾钙碱性-钾玄岩系列.微量元素(La/Yb)N 介于5.52~9.89之间,δEu=0.09~0.90,具有中等-强烈负铕异常,相对富集大离子亲石元素包括Rb,K,La,Th等,而等高场强元素P,Ta,Nb、Ti则表现亏损,指示岩浆形成于与俯冲带有关的岛弧或

  12. 西藏申扎早二叠世碳酸盐岩的极低碳稳定同位素及其冷泉指示意义%Extremely 13 C-depleted Carbonate Indicating Cold Seepage in Early Permian from Xainza,Tibet

    安显银; 张予杰; 朱同兴


    Extremely 1 3 C-depleted isotope is the key evidence to identify ancient cold seepage and methane re-lease event in Earth history.There has been no report of direct evidence of cold seepage in Early Permian period so far.Here we unprecedentedly present the carbonate carbon and oxygen isotopes with extreme 1 3 C depletion in Early Permian of Lasha Block,Tibet.The negative correlation of carbon and oxygen isotopes is pronounced and the lowest carbon isotope is to -34.69‰(VPDB),exclusively indicating anaerobic oxidation of methane. These carbonates occur in the upper member of the Lanfjie Formation in Xainza,Tibet as carbonate lens and carbonate-cemented concretion,and calcite crystal fans are more depleted in 1 3 C than carbonate cements in the concretions.We infer the occurrence of ancient cold seepage in Early Permian in the study area and attribute this event to the postglacial destabilization of gas hydrate reservoir.This study is significant to investigate glac-ioeustatic change,oceanographic geochemistry,paleoecology and climate change.%极低碳稳定同位素是识别地质历史时期古冷泉及其甲烷释放事件的关键证据,目前,世界范围内尚无有关二叠纪冷泉活动的直接证据报道。本研究首次报道了西藏地区拉萨地块早二叠世碳酸盐岩的极低碳同位素数据,δ13 CV -PDB值最低可达-34.69‰,碳、氧稳定同位素呈现较好的负相关性,指示为甲烷厌氧氧化作用成因。该碳酸盐岩发育于西藏申扎地区下二叠统昂杰组上段(碎屑岩段),以透镜状碳酸盐岩夹层及碳酸盐质胶结结核产出,可见方解石结晶扇,其碳同位素普遍较结核中碳酸盐胶结物更贫13 C。推测研究区存在早二叠世古甲烷冷泉渗漏活动,其成因可能与冈瓦纳冰期后温暖海水导致的海洋甲烷水合物藏的失稳释放有关。作为研讨地质历史时期水合物-冰期反馈模型的重要实例,本研究对进一步

  13. Análise espacial da mortalidade neonatal precoce no Município do Rio de Janeiro, 1995-1996 Spatial analysis of early neonatal mortality in the municipality of Rio de Janeiro, 1995-1996

    Carla Lourenço Tavares de Andrade


    Full Text Available O objetivo deste trabalho foi estabelecer a distribuição espacial da mortalidade neonatal precoce (0-3 dias no Município do Rio de Janeiro de 1995 a 1996, visando identificar os fatores mais explicativos das variações espaciais. Considerando os bairros como unidade ecológica de análise, foram analisados indicadores relativos às condições sócio-econômicas e às características das mães dos recém-nascidos por bairro de residência. A análise estatística espacial dos dados foi realizada utilizando as técnicas de Cliff & Ord, apropriadas para "dados de áreas". Por meio do mapa temático da mortalidade de 0-3 dias, observou-se nitidamente a presença de dois aglomerados de taxas elevadas. A dependência espacial foi igualmente constatada pelos resultados da análise estatística. As variáveis que melhor explicaram os aglomerados espaciais foram: "proporção de mães adolescentes", "proporção de pessoas residentes em favelas em 1996" e "proporção de chefes com rendimento até um salário mínimo". A metodologia de estatística espacial permitiu maior compreensão da distribuição geográfica da mortalidade neonatal precoce, e deu margem a outros tipos de investigações, que poderão subsidiar os programas preventivos e contribuir ao declínio da mortalidade infantil.The objective of this study was to establish the spatial distribution of the early neonatal mortality rate (0-3 days in the municipality of Rio de Janeiro for 1995-1996, identifying the best explanatory factors for spatial variations. By considering Rio de Janeiro's 153 neighborhoods as ecological units of analysis, socioeconomic and maternal indicators were analyzed according to place of residence. Spatial statistical analysis was performed using the Cliff & Ord methodology, appropriate for lattice data. From the 0-3 day mortality thematic map, we clearly identified two clusters of high early neonatal mortality rates. Spatial dependence was also confirmed by the

  14. The Late Permian-Early Triassic cyclostratigraphy in Shangsi section of Guangyuan area, Sichuan Province: Implications for P-T geological event based on wavelet analysis%四川广元上寺剖面晚二叠世-早三叠世旋回地层:基于小波分析的P-T界线地质事件探讨

    乔彦国; 时志强; 王艳艳; 张彪


    . The rapid rate of deposition displayed a strong influence on marine ecosystems, that is, it restrained the diversity of marine fauna and delayed the resuscitating of marine ecosystems. The cyclicity of the latest Permian sedimentary record was more obvious and stable in Shangsi section. However, the periodicity turned to be labile when it came to the Early Triassic. We concluded that the accelerated deposition rate and deteriorated sedimentary cyclicity were triggered by the turbulent paleoclimate and mussy marine environment. The perturbation of the terrestrial realm contributed to the change of marine conditions, which might be one of the major reasons for the marine ecosystem recovering relatively slowly in the Early Triassic.

  15. Permian plate margin volcanism and tuffs in adjacent basins of west Gondwana: Age constraints and common characteristics

    López-Gamundí, Oscar


    Increasing evidence of Permian volcanic activity along the South American portion of the Gondwana proto-Pacific margin has directed attention to its potential presence in the stratigraphic record of adjacent basins. In recent years, tuffaceous horizons have been identified in late Early Permian-through Middle Permian (280-260 Ma) sections of the Paraná Basin (Brazil, Paraguay, and Uruguay). Farther south and closer to the magmatic tract developed along the continental margin, in the San Rafael and Sauce Grande basins of Argentina, tuffs are present in the Early to Middle Permian section. This tuff-rich interval can be correlated with the appearance of widespread tuffs in the Karoo Basin. Although magmatic activity along the proto-Pacific plate margin was continuous during the Late Paleozoic, Choiyoi silicic volcanism along the Andean Cordillera and its equivalent in Patagonia peaked between the late Early Permian and Middle Permian, when extensive rhyolitic ignimbrites and consanguineous airborne tuffaceous material erupted in the northern Patagonian region. The San Rafael orogenic phase (SROP) interrupted sedimentation along the southwestern segment of the Gondwana margin (i.e., Frontal Cordillera, San Rafael Basin), induced cratonward thrusting (i.e., Ventana and Cape foldbelts), and triggered accelerated subsidence in the adjacent basins (Sauce Grande and Karoo) located inboard of the deformation front. This accelerated subsidence favored the preservation of tuffaceous horizons in the syntectonic successions. The age constraints and similarities in composition between the volcanics along the continental margin and the tuffaceous horizons in the San Rafael, Sauce Grande, Paraná, and Karoo basins strongly suggest a genetic linkage between the two episodes. Radiometric ages from tuffs in the San Rafael, Paraná, and Karoo basins indicate an intensely tuffaceous interval between 280 and 260 Ma.

  16. Crinoids from Svalbard in the aftermath of the end−Permian mass extinction

    Salamon Mariusz A.


    Full Text Available The end-Permian mass extinction constituted a major event in the history of crinoids. It led to the demise of the major Paleozoic crinoid groups including cladids, disparids, flexibles and camerates. It is widely accepted that a single lineage, derived from a late Paleozoic cladid ancestor (Ampelocrinidae, survived this mass extinction. Holocrinid crinoids (Holocrinus, Holocrinida along with recently described genus Baudicrinus (Encrinida, the only crinoid groups known from the Early Triassic, are considered the stem groups for the post-Paleozoic monophyletic subclass Articulata. Here, we report preliminary data on unexpectedly diverse crinoid faunas comprising at least four orders from the Lower Triassic (Induan and Olenekian of Svalbard, extending their stratigraphic ranges deeper into the early Mesozoic. These findings strongly imply that the recovery of crinoids in the aftermath of the end-Permian extinction began much earlier at higher palaeolatitudes than in the central Tethys.

  17. Cretaceous stem chondrichthyans survived the end-Permian mass extinction.

    Guinot, Guillaume; Adnet, Sylvain; Cavin, Lionel; Cappetta, Henri


    Cladodontomorph sharks are Palaeozoic stem chondrichthyans thought to go extinct at the end-Permian mass extinction. This extinction preceded the diversification of euselachians, including modern sharks. Here we describe an outer-platform cladodontomorph shark tooth assemblage from the Early Cretaceous of southern France, increasing the fossil record of this group by circa 120 million years. Identification of this material rests on new histological observations and morphological evidence. Our finding shows that this lineage survived mass extinctions most likely by habitat contraction, using deep-sea refuge environments during catastrophic events. The recorded gap in the cladodontomorph lineage represents the longest gap in the fossil record for an extinct marine vertebrate group. This discovery demonstrates that the deep-sea marine diversity, poorly known during most of the fish evolutionary history, contains essential data for a complete understanding of the long-term evolution of marine fish paleobiodiversity.

  18. The terminal Permian in European Russia: Vyaznikovian Horizon, Nedubrovo Member, and Permian-Triassic boundary

    Lozovsky, V. R.; Balabanov, Yu. P.; Karasev, E. V.; Novikov, I. V.; Ponomarenko, A. G.; Yaroshenko, O. P.


    The comprehensive analysis of the data obtained on terrestrial vertebrata, ostracods, entomologic fauna, megaflora, and microflora in deposits of the Vyaznikovian Horizon and Nedubrovo Member, as well as the paleomagnetic data measured in enclosing rocks, confirms heterogeneity of these deposits. Accordingly, it is necessary to distinguish these two stratons in the terminal Permian of the East European Platform. The combined sequence of Triassic-Permian boundary deposits in the Moscow Syneclise, which is considered to be the most complete sequence in the East European Platform, is as follows (from bottom upward): Vyatkian deposits; Vyaznikovian Horizon, including Sokovka and Zhukovo members; Nedubrovo Member (Upper Permian); Astashikha and Ryabi members of the Vokhmian Horizon (Lower Triassic). None of the sequences of Permian-Triassic boundary deposits known in the area of study characterizes this sequence in full volume. In the north, the Triassic deposits are underlain by the Nedubrovo Member; in the south (the Klyazma River basin), the sections are underlain by the Vyaznikovian Horizon. The Permian-Triassic boundary adopted in the General Stratigraphic Scale of Russia for continental deposits of the East European platform (the lower boundary of the Astashikha Member) is more ancient than the one adopted in the International Stratigraphic Chart. The same geological situation is observed in the German Basin and other localities where Triassic continental deposits are developed. The ways of solving this problem are discussed in this article.

  19. Coring and High-Resolution Imaging of the Permian-Triassic Boundary in Deltadalen, Svalbard

    Planke, Sverre


    The Permian-Triassic extinction was likely triggered by the voluminous igneous activity of the Siberian Traps. There are, however, limited information available about the effects of the Siberian Traps volcanism in Permian to Triassic sedimentary successions outside Siberia. We drilled two boreholes across the Permian-Triassic boundary (PTB) in Deltadalen, Svalbard, in August 2014, to better document the extinction event and environmental changes in the sedimentary succession in this region. Even though the PTB event is studied in several outcrops to varying degree if scrutiny on Svalbard, a scientific core is important for understanding the details of the PTB event, as outcrops may not permit studies at the appropriate levels of resolution, exposure or confidence. The Deltadalen site was chosen as an optimal drilling location after field work in the Isfjorden area in 2013 and early 2014. The almost 100 m deep holes were fully cored and penetrated the Triassic Vikinghøgda Formation (about 85 m recovered) and the Permian Kapp Starostin Formation (about 15 m). The main lithologies include Permian chert-rich green glauconitic sandstones and Triassic mudstones. A 10 m thick sedimentary reference outcrop section across the PTB was further logged and sampled in a nearby river valley. A comprehensive core analysis program is now underway. High-resolution XRF, MST and hyperspectral core scanning has been completed along the entire core surface, while high-resolution CT scanning has been carried out for whole core sections. The DD-1 core has subsequently been split, photographed, described, and sampled for geochemical, micropaleontological, petrological and magnetic analyses. Almost 10 bentonite layers have furthermore been sampled for volcanological and geochronological studies. Our goal is that the cores will become an important future reference section of the PTB in Svalbard and the high-Arctic, and help constrain the extent of the regional effects of explosive volcanism

  20. Inter-taxa differences in extinction process of Maokouan (Middle Permian) fusulinaceans


    Analyses of the extinction process of fusulinacean in the Maokouan (Middle Permian) biotic crisis have revealed remarkable differences between taxa with various morphological features. Nankinellinids suffered a heavy loss of species in the early stage of the Maokouan event. Schwagerinids and neoschwagerinids both showed a stepwise decrease in species diversity, but the pulses of species extinction occurred in different stages of the extinction process. The species extinction of verbeekinids happened primarily in the Late Maokouan.

  1. Behavior of lophophorates during the end-Permian mass extinction and recovery

    Powers, Catherine M.; Bottjer, David J.


    The end-Permian mass extinction devastated most marine communities and the recovery was a protracted event lasting several million years into the Early Triassic. Environmental and biological processes undoubtedly controlled patterns of recovery for marine invertebrates in the aftermath of the extinction, but are often difficult to single-out. The global diversity and distribution of marine lophophorates during the aftermath of the end-Permian mass extinction indicates that stenolaemate bryozoans, rhynchonelliform brachiopods, and lingulid brachiopods displayed distinct recovery patterns. Bryozoans were the most susceptible of the lophophorates, experiencing relatively high rates of extinction at the end of the Permian, and becoming restricted to the Boreal region during the Early Triassic. The recovery of bryozoans was also delayed until the Late Triassic and characterized by very low diversity and abundance. Following the final disappearance of Permian rhynchonelliform brachiopod survivors, Early Triassic rhynchonelliform brachiopod abundance remained suppressed despite a successful re-diversification and a global distribution, suggesting a decoupling between global taxonomic and ecological processes likely driven by lingering environmental stress. In contrast with bryozoans and rhynchonelliforms, lingulid brachiopods rebounded rapidly, colonizing shallow marine settings left vacant by the extinction. Lingulid dominance, characterized by low diversity but high numerical abundance, was short-lived and they were once again displaced back into marginal settings as environmental stress changed through the marine recovery. The presence in lingulid brachiopods of the respiratory pigment hemerythrin, known to increase the efficacy of oxygen storage and transport, when coupled with other morphological and physiological adaptations, may have given lingulids a survival advantage in environmentally stressed Early Triassic settings.




    Full Text Available Detailed conodont biostratigraphy and lithostratigraphy of the Late Permian and Early Triassic beds were studied at the LukaC section in western Slovenia. The analyzed section is composed of the Bellerophon Formation ("evaporite-dolomite member" and the newly introduced Lukaè Formation ("transitional beds", "streaky limestone member" and "carbonate-clastic beds member". The Permian-Triassic boundary interval is represented by "transitional beds" of carbonate facies deposited in shallow restricted marine conditions. The presence of H. parvus in sample L1 in the "transitional beds" marks the systemic boundary between Permian and Triassic. The studied interval is characterized by a diverse microfauna that contain conodonts, foraminifers, ostracods and gastropods. Six conodont zones have been recognized, in ascending order, the latest Changhsingian (uppermost Permian praeparvus Zone, and the Griesbachian (lowermost Triassic parvus, lobata, staeschei-isarcica, postparvus and anceps zones. This faunal succession represents the first known and the most complete conodont biozonation across the Permian-Triassic interval from the entire Dinaric region. The recognized conodont biozones can be correlated with the biozonation of the Southern Alps and of the GSSP Meishan D section. 

  3. Thecamoebians (Testate Amoebae) Straddling the Permian-Triassic Boundary in the Guryul Ravine Section, India: Evolutionary and Palaeoecological Implications.

    Singh, Vartika; Pandita, Sundeep K; Tewari, Rajni; van Hengstum, Peter J; Pillai, Suresh S K; Agnihotri, Deepa; Kumar, Kamlesh; Bhat, G D


    Exceptionally well-preserved organic remains of thecamoebians (testate amoebae) were preserved in marine sediments that straddle the greatest extinction event in the Phanerozoic: the Permian-Triassic Boundary. Outcrops from the Late Permian Zewan Formation and the Early Triassic Khunamuh Formation are represented by a complete sedimentary sequence at the Guryul Ravine Section in Kashmir, India, which is an archetypal Permian-Triassic boundary sequence. Previous biostratigraphic analysis provides chronological control for the section, and a perspective of faunal turnover in the brachiopods, ammonoids, bivalves, conodonts, gastropods and foraminifera. Thecamoebians were concentrated from bulk sediments using palynological procedures, which isolated the organic constituents of preserved thecamoebian tests. The recovered individuals demonstrate exceptional similarity to the modern thecamoebian families Centropyxidae, Arcellidae, Hyalospheniidae and Trigonopyxidae, however, the vast majority belong to the Centropyxidae. This study further confirms the morphologic stability of the thecamoebian lineages through the Phanerozoic, and most importantly, their apparent little response to an infamous biological crisis in Earth's history.

  4. Petrogenesis of Late Carboniferous to Early Permian Granitoid Plutons in the Chinese South Tianshan: Implications for Crustal Accretion%中国南天山晚石炭世一早二叠世花岗质侵入岩的岩石成因与地壳增生

    黄河; 张招崇; 张东阳; 杜红星; 马乐天; 康建丽; 薛春纪


    晚石炭世—早二叠世在中亚南天山造山带形成了大量的花岗质侵入岩.中国境内这些岩体以晚石炭世Ⅰ型花岗岩、早二叠世Ⅰ型和S型花岗岩以及最晚期的A型花岗岩为代表.不同类型的岩石在源区特征和岩浆形成的温压条件上存在一定的差异,而这些差异性反映了该时期内构造环境的演化过程.本文选取铁列克岩体、盲起苏岩体、英买来岩体、川乌鲁杂岩体、巴雷公岩体、霍什布拉克岩体和麻扎山岩体作为典型岩体进行了地质年代学、岩石学和地球化学的研究.铁列克岩体和盲起苏岩体为晚石炭世Ⅰ型花岗岩,岩浆形成于地壳下部变质火成岩在同碰撞构造背景下发生的部分熔融作用;英买来岩体为早二叠世S型花岗岩,是变质沉积岩为主的中元古代古老地层在放射性物质蜕变产生的热源作用下重熔的产物;川乌鲁杂岩体中的长英质岩石为早二叠世Ⅰ型花岗岩,并具有岩浆混合成因,代表了由于碰撞导致的加厚地壳底部物质部分熔融产生的花岗质岩浆.而杂岩体中的铁镁质岩石代表了富集交代的岩石圈地幔部分熔融产生的底侵幔源岩浆.英买来岩体和川乌鲁杂岩体均形成于构造体制由同碰撞挤压背景向后碰撞伸展环境转化的阶段;形成于275 Ma前后的A型花岗岩代表了软流圈物质与下地壳在低压条件下相互作用形成的高温花岗质岩浆.其中巴雷公岩体为A2型花岗岩,代表了后碰撞环境的尾声;霍什布拉克岩体和麻扎山岩体都为A1型花岗岩,指示了板内裂谷环境.%A large volume of granitoid plutons formed in the South Tianshan orogenic belt during Late Carboniferous to Early Permian. These rocks can be generally classified as Late Carboniferous I-type granites, Early Permian I and S-type granites, and A-type granites that formed at -275 Ma. The different rocks have some differences in features of source

  5. Late Permian to Triassic intraplate orogeny of the southern Tianshan and adjacent regions, NW China

    Wei Ju


    Based on previous studies and recent geochronogical data, we suggest that the final collision between the Tarim Craton and the North Asian continent occurred during the late Carboniferous. Therefore, the Permian was a period of intracontinental environment in the southern Tianshan and adjacent regions. We propose that an earlier, small-scale intraplate orogenic stage occurred in late Permian to Triassic time, which was the first intraplate process in the South Tianshan Orogen and adjacent regions. The later large-scale and well-known Neogene to Quaternary intraplate orogeny was induced by the collision between the India subcontinent and the Eurasian plate. The paper presents a new evolutionary model for the South Tianshan Orogen and adjacent regions, which includes seven stages: (I late Ordovician–early Silurian opening of the South Tianshan Ocean; (II middle Silurian–middle Devonian subduction of the South Tianshan Ocean beneath an active margin of the North Asian continent; (III late Devonian–late Carboniferous closure of the South Tianshan Ocean and collision between the Kazakhstan-Yili and Tarim continental blocks; (IV early Permian post-collisional magmatism and rifting; (V late Permian–Triassic the first intraplate orogeny; (VI Jurassic–Palaeogene tectonic stagnation and (VII Neocene–Quaternary intraplate orogeny.

  6. Severest crisis overlooked-Worst disruption of terrestrial environments postdates the Permian-Triassic mass extinction.

    Hochuli, Peter A; Sanson-Barrera, Anna; Schneebeli-Hermann, Elke; Bucher, Hugo


    Generally Early Triassic floras are believed to be depauperate, suffering from protracted recovery following the Permian-Triassic extinction event. Here we present palynological data of an expanded East Greenland section documenting recovered floras in the basal Triassic (Griesbachian) and a subsequent fundamental floral turnover, postdating the Permian-Triassic boundary extinction by about 500 kyrs. This event is marked by a swap in dominating floral elements, changing from gymnosperm pollen-dominated associations in the Griesbachian to lycopsid spore-dominated assemblages in the Dienerian. This turnover coincides with an extreme δ(13)Corg negative shift revealing a severe environmental crisis, probably induced by volcanic outbursts of the Siberian Traps, accompanied by a climatic turnover, changing from cool and dry in the Griesbachian to hot and humid in the Dienerian. Estimates of sedimentation rates suggest that this environmental alteration took place within some 1000 years. Similar, coeval changes documented on the North Indian Margin (Pakistan) and the Bowen Basin (Australia) indicate the global extent of this crisis. Our results evidence the first profound disruption of the recovery of terrestrial environments about 500kyrs after the Permian-Triassic extinction event. It was followed by another crisis, about 1myrs later thus, the Early Triassic can be characterised as a time of successive environmental crises.

  7. Permian vegetational Pompeii from Inner Mongolia and its implications for landscape paleoecology and paleobiogeography of Cathaysia.

    Wang, Jun; Pfefferkorn, Hermann W; Zhang, Yi; Feng, Zhuo


    Plant communities of the geologic past can be reconstructed with high fidelity only if they were preserved in place in an instant in time. Here we report such a flora from an early Permian (ca. 298 Ma) ash-fall tuff in Inner Mongolia, a time interval and area where such information is filling a large gap of knowledge. About 1,000 m(2) of forest growing on peat could be reconstructed based on the actual location of individual plants. Tree ferns formed a lower canopy and either Cordaites, a coniferophyte, or Sigillaria, a lycopsid, were present as taller trees. Noeggerathiales, an enigmatic and extinct spore-bearing plant group of small trees, is represented by three species that have been found as nearly complete specimens and are presented in reconstructions in their plant community. Landscape heterogenity is apparent, including one site where Noeggerathiales are dominant. This peat-forming flora is also taxonomically distinct from those growing on clastic soils in the same area and during the same time interval. This Permian flora demonstrates both similarities and differences to floras of the same age in Europe and North America and confirms the distinct character of the Cathaysian floral realm. Therefore, this flora will serve as a baseline for the study of other fossil floras in East Asia and the early Permian globally that will be needed for a better understanding of paleoclimate evolution through time.

  8. Permian Radiolarians from the A’nyemaqen Mélange Zone in the Huashixia Area of Madoi County, Qinghai Province, Western China, and Their Implications on Regional Tectonism

    Zhigang Zhao; Xing Huang; Xionghua Zhang; Bing Yang; Zhong-Qiang Chen; Fuhao Xiong


    The Huashixia area in the Madoi County, Qinghai Province, western China is one of the most important areas for the studies of the formation and evolution of the A’nyemaqen mélange zone due to the well exposed, relatively complete tectonostratigraphic successions. However, the timing and evolu-tionary process of the A’nyemaqen paleo-ocean has long been poorly understood. Recently, we obtained abundant radiolarians from the siliceous rocks of the Maerzheng Formation of this mélange zone in Huashixia. Total 19 species in 10 radiolarian genera have been identified. They belong mainly to the Al-baillellaria, Spumellaria, Entactinaria, and Latentifistularia. Four radiolarian assemblage zones are es-tablished, including: thePseudoalbaillella rhombothoracata,Pseudoalbaillella scalprata,Ruzhencevispongus uralicus,andFollicucullus bipartitus-Follicucullussp. cf.F. ventricosus zones. Radiolarian faunal correla-tions suggest a late Artinskian–early Kungurian age and late Capitanian–early Wuchiapingian age for the first two zones from the lower part and the last two zones from the upper parts of the Maerzheng Forma-tion, respectively. All radiolarians are typical of deep water elements living in deep oceans in low latitudes during the Permian. Previously, the radiolarians of Early Carboniferous to Early Permian age and fusu-linids of Early–Middle Permian age have also been reported from the A’nyemaqen mélange successions. The combination of the newly found radiolarians with the previously reported faunal and microfloral as-semblages as well as radiometric ages obtained from the same stratigraphic units indicates that the A’nyemaqen paleo-ocean may have opened initially during the Early Carboniferous, expanded from the Early–Middle Permian, reached its acme in the early Late Permian, shrunk during the latest Permian to Early Triassic, and eventually was closed in the Middle Triassic.




    Full Text Available The diversity curve of Permian fusulinid genera shows two peaks corresponding to the Asselian-Sakmarian and Midian times. The minimal generic diversity is recorded in the late Bolorian. The most significant extinction events occurred at the end of the Midian (71% of all genera, Bolorian (48%, Asselian (27% and Sakmarian (23% ages. The fusulinid assemblage was most notably changed by the appearance of new genera (52% of the total number in the Kubergandian age. These data  identify two main stages (Asselian-Bolorian and Kubergandian-Dorashamian and four second-order stages (Asselian-Sakmarian, Yakhtashian-Bolorian, Kubergandian-Midian, and Dzhulfian-Dorashamian in the Permian history of fusulinids. The main stages correspond to two Permian series of the East European scale, which can be considered as subsystems named Cisuralian and Tethysian, respectively. The latter are subdivided into the Uralian, Darvasian, Yanghsingian and Lopingian series which correspond to the second-order stages. The scale suggested does not contradict the traditional two-member subdivision and has an advantage over the accepted global three-member chronostratigraphic scale because the series suggested are more proportional to each other in scope and reflect natural evolutionary processes of the marine biota. In addition, the application of the global scale to the Tethyan sequences is hampered by a limited number of criteria used in the drawing of series and stage boundaries, as evidenced by the existing different views on the position of the lower Guadalupian boundary in the Tethyan sections. 

  10. Significance of new breakthrough in and favorable targets of gas exploration in the Middle Permian system, Sichuan Basin

    Ping Shen


    Full Text Available The Middle Permian system in the Sichuan Basin represents favorable conditions for the formation of natural gas reservoirs. However, only fracture-vuggy limestone gas reservoirs were discovered in early exploration, whose scales and results were limited. In 2014, a high-yield industrial gas flow was produced there in Well Shuangtan 1, the major risk-exploration well, which means a great breakthrough in natural gas exploration in the Middle Permian system, Sichuan Basin. This plays a strategically important and guiding role in oil and gas exploration in ultra-deep marine formations. This paper illustrates the deployment background of Well Shuangtan 1, major exploration achievements and its significance in the gas exploration in the Middle Permian system, Sichuan Basin. Moreover, exploration potential and subsequent exploration targets of Middle Permian are analyzed in terms of its hydrocarbon source conditions, sedimentary characteristics, reservoir characteristics and areal extension, resources and favorable exploration zones. It is concluded that the Middle Permian system in the Sichuan Basin is a continent-connected carbonate platform with carbonate deposits being dominant. There are three types of reservoirs, including pore dolomite, karst fracture-vuggy, and fracture, which present considerable natural gas resources and huge exploration potential. Favorable exploration zones classification of Middle Permian in the Sichuan Basin indicates that the most favorable exploration zones are mainly distributed in the central-western Sichuan Basin. In particular, the northwestern Sichuan Basin is the most favorable exploration area which is characterized by the most excellent hydrocarbon source conditions, developed dolomite reservoirs and zonal distribution of structures.

  11. Life crises on land across the Permian-Triassic boundary in South China

    Peng, Yuanqiao; Shi, G. R.


    The western Guizhou and eastern Yunnan area of southwest China commands a unique and significant position globally in the study of Permian-Triassic boundary (PTB) events as it contains well and continuously exposed PTB sections of marine, non-marine and marginal-marine origin in the same area. By using a range of high-resolution stratigraphic methods including biostratigraphy, eventostratigraphy, chronostratigraphy and chemostratigraphy, not only are the non-marine PTB sections correlated with their marine counterparts in the study area with high-resolution, the non-marine PTB sections of the study area can also be aligned with the PTB Global Stratotype Section and Point (GSSP) at Meishan in eastern China. Plant megafossils ("megaplants") in the study area indicate a major loss in abundance and diversity across the PTB, and no coal beds and/or seams have been found in the non-marine Lower Triassic although they are very common in the non-marine Upper Permian. The megaplants, however, did not disappear consistently across the whole area, with some elements of the Late Permian Cathaysian Gigantopteris flora surviving the PTB mass extinction and locally even extending up to the Lower Triassic. Palynomorphs exhibit a similar temporal pattern characterized by a protracted stepwise decrease from fern-dominated spores in the Late Permian to pteridosperm and gymnosperm-dominated pollen in the Early Triassic, which was however punctuated by an accelerated loss in both abundance and diversity across the PTB. Contemporaneous with the PTB crisis in the study area was the peculiar prevalence and dominance of some fungi and/or algae species. The temporal patterns of megaplants and palynomorphs across the PTB in the study area are consistent with the regional trends of plant changes in South China, which also show a long-term decrease in species diversity from the Late Permian Wuchiapingian through the Changhsingian to the earliest Triassic, with about 48% and 77% losses of

  12. Chemo- and palyno-stratigraphy of the Permian-Triassic transition in the Boreal region

    van Soelen, Els; Planke, Sverre; Svensen, Henrik; Twitchett, Richard; Polozov, Alexander; Kürschner, Wolfram


    Late Permian and early Triassic sediments from Boreal regions are studied using palynological and organic geochemical tools. We present preliminary results from two sites: a Norwegian site which is represented by a 100-m long borehole core and outcrops from Deltadalen on Spitsbergen, and a Russian site which is represented by outcrops and short cores collected near Norilsk in northern Siberia. The main goals of the project are to improve the stratigraphy and to study the environmental changes at high resolution. There is a growing scientific consensus that end Permian biotic crisis was linked to the Siberian Traps Large Igneous Province (LIP) event. However, direct evidence for a stratigraphic correlation of the marine and terrestrial extinction events, with the volcanic successions in the Siberian basin, is rather limited. The Permian-Triassic boundary successions in the Arctic are crucial for direct correlation eastwards to the Siberian Traps. The magnitude and timing of a carbon isotope excursion near the Permian-Triassic boundary is an important stratigraphical tool that may help to unravel the sequence of the events happening during this important period. Preliminary results from the Deltadalen core near the base of the Vikinghøgda Formation show shifts in δ13C from -24.5 to -32.7‰ in the interval expected to span the Permian/Triassic boundary. New Rock-Eval pyrolysis data will shed further light on the origin of the organic matter (e.g. marine versus terrestrial) and may help to understand how much of the δ13C signal can be explained by changes in organic matter source and how much may be attributed to a global change in the carbon isotope signature. Furthermore, compound specific isotope analysis will be done on terrestrial derived lipids (long chain n-alkanes) to reconstruct changes in atmospheric carbon isotopes. In addition to chemostratigraphy, the palynological record will be used for biostratigraphical studies at both Deltadalen and Norilsk

  13. Palynofloral associations before and after the Permian-Triassic mass extinction, Kap Stosch, East Greenland

    Schneebeli-Hermann, Elke; Hochuli, Peter A.; Bucher, Hugo


    The Permian-Triassic boundary (PTB) interval is known to document a major biodiversity crisis in the history of life. It is generally accepted that this crisis had a significant impact on marine invertebrates. The consequences for terrestrial ecosystems are still controversially discussed. Based on palynological analysis we present a high time resolution microfloral succession of the expanded Late Permian (Wuchiapingian)-Early Triassic (Dienerian) section from Kap Stosch, East Greenland. The quantitative distribution of palynomorphs (range charts and relative abundance data) allows for the differentiation of six distinct palynofloral associations. Ammonoids and conodonts provide independent age control for these assemblages. The Wuchiapingian association I, documented from the Ravnefjeld Formation, shows a typical Late Permian assemblage dominated by bisaccate and monosaccate pollen grains and Vittatina spp. It is separated from association II, present in the basal part of the Wordie Creek Formation, by an important hiatus. This association of Changhsingian or earliest Griesbachian age is characterised by the common occurrence of Ephedripites spp. and reduced abundance and diversity of Vittatina spp. Association III, dated as Griesbachian by the presence of ammonoids, is marked by high relative abundances of taeniate bisaccate pollen grains and high spore diversity. A distinct floral break occurs between the gymnosperm dominated Permian and Griesbachian floras and the lycopsid spore dominated Dienerian associations IV-VI. Ammonoid occurrences verify a Dienerian age for the latter associations. Association V is marked by the absence of non-taeniate bisaccate, striate monosaccate pollen grains, and Vittatina spp. Aratrisporites spp. a typical Triassic lycopsid spore occur consistently from this level onwards. Association VI is characterised by highest lycopsid spore abundances. Cluster analysis demonstrates that Griesbachian assemblages (associations II?-III) are

  14. Return to Coalsack Bluff and the Permian Triassic boundary in Antarctica

    Retallack, Gregory J.; Greaver, Tara; Jahren, A. Hope


    Coalsack Bluff was the first discovery site in Antarctica for the latest Permian to earliest Triassic reptile Lystrosaurus. This together with discovery of Permian Glossopteris leaves during the heroic age of Antarctic exploration, indicated not only that Antarctica was part of Gondwanaland, but also that Antarctic rocks recorded faunas from the greatest of all mass extinctions at the Permian-Triassic boundary. Pinpointing the exact stratigraphic level of this life crisis has recently become possible using δ 13C values in terrestrial organic matter. Multiple, short-lived events of 13C depletion may reflect carbon cycle crises, with the isotopic change a measure of terrestrial and atmospheric disequilibrium. Additional evidence for ecosystem reorganization came from changes in paleosol types and their root traces. Such studies previously completed at the Antarctic localities of Graphite Peak, Mount Crean, Portal Mountain, Shapeless Mountain and Allan Hills, are here extended to Coalsack Bluff. Carbon isotopic values in Permian rocks at Coalsack Bluff average - 23.08 ± 0.25‰, but begin to decline within the last coal with leaves ( Glossopteris), roots ( Vertebraria) and permineralized stumps ( Araucarioxylon) of glossopterids. The low point in ä 13C values is - 27.19‰ at 5.6 m above the last coal, which is capped by unusually abundant pyrite, and a claystone breccia with common clasts of redeposited clayey soils. Above this are massive quartz-rich sandstones of braided streams, considered a geomorphic response to deforestation and soil erosion following the mass extinction. Distinctive berthierine-bearing paleosols (Dolores pedotype) within these sandstones have unoxidized iron taken as evidence of severe groundwater hypoxia. Other paleosols at this stratigraphic level are like those in other Early Triassic rocks of Antarctica, which indicate unusually warm and humid conditions for such high paleolatitude lowlands. Waterlogging is also indicated by newly




    Full Text Available The Permian sequence of the Transcaucasia, which comprises the Davaly, Asni, Gnishik, Arpa, Khachik, and Akhura Formations, is described. A precise dating of the formations based on the analysis of fusulinid assemblages is also provided. The Davaly Formation corresponds to the Bolorian Stage of the Lower Permian (Cisuralian; the Asni Formation corresponds to the Kubergandian Stage, and to the lower half of the Murgabian Stage of the Middle Permian (Guadalupian; the Gnishik Formation corresponds approximately to the upper half of the Murgabian Stage; and the Arpa and Khachik Formations represent the entire Midian Stage of the Middle Permian. Only the uppermost Chanakhchy Beds of the Khachik Formation are referred to the upper series of the Permian (Lopingian, based on the latest proposal of the International Subcommission on Permian Stratigraphy concerning series boundaries. The Akhura Formation encompasses the Dzhulfian and Dorashamian Stages of the Upper Permian. Refined correlations between the Permian sequence of the Transcaucasia and the most complete Permian sequences of the Tethyan region (Central Iran, southestern Pamirs, South China, Japan are presented. The transgressive overlapping of the Permian deposits is recorded in these areas, similar to the Transcaucasia. Although the transgression began during the Bolorian, it reached the widest extent in the Kubergandian and therefore, in most sections the transgressive series begins with deposits of Kubergandian age. The Middle and Upper Permian deposits of the Transcaucasia, Iran, and southern Turkey belonged to a single carbonate platform. Similar extensive carbonate platforms are recorded in the southern parts of Afghanistan and China. A change in the sedimentary regime was recorded in all these platforms at the boundary between the Guadalupian and Lopingian epochs. The Guadalupian time was marked by the intensive accumulation of biolitithic and detrital limestones produced by benthic

  16. Further paleomagnetic results for lower Permian basalts of the Baoshan Terrane, southwestern China, and paleogeographic implications

    Xu, Yingchao; Yang, Zhenyu; Tong, Ya-Bo; Wang, Heng; Gao, Liang; An, Chunzhi


    The Baoshan Terrane of southwestern China is considered to have been part of the Cimmerian block during the late Paleozoic; consequently, knowledge of its paleoposition and geological evolution can provide constraints on the Permian breakup of northern East Gondwana. Therefore, we conducted paleomagnetic and rockmagnetic studies on lower Permian basalts from four localities in the Baoshan Terrane. The basalts hold a stable characteristic remanent magnetization (ChRM) at high temperatures (300-680 °C) that is carried by magnetite, maghemite, and hematite with both pseudo-single and multiple domains. To test the reliability of data from these volcanic rocks, we analyzed the geomagnetic secular variation (GSV) and reliability of both the present data and previous paleomagnetic data. The results from 23 sites yield a single reversed polarity directed downwards to the southwest, giving a site-mean direction of Dg/Ig = 156.7°/56.6° (kg = 8.0, α95 = 11.4°) before tilt correction, and Ds/Is = 218.3°/60.1° (ks = 14.1, α95 = 8.4°) after tilt correction. The result passed the fold test, but the GSV was able to be averaged out in only two sections. All available data were examined section-by-section using the angular dispersion (SB) of virtual geomagnetic poles (VGPs) to ensure that the GSV was completely averaged out. Because the dispersion in declinations is likely to have been affectedby subsequent tectonic deformation, the paleosecular variation (PSV) could not be evaluated from all the data amassed from different sections, and the PSV was able to be removed from only four (combined) sections. A small-circle fit of these VGPs gives an averaged paleocolatitude of 51.9° ± 3.7° (N = 31 sites) centered on 24°N, 99°E. The result indicates that the sampled area of the Baoshan Terrane was located at a latitude of 38°S ± 3.7° during the late early Permian. A comparison of this result with early Permian data from Gondwanan blocks suggests that the Baoshan Terrane

  17. New Late Permian paleomagnetic data from Argentina: Refinement of the apparent polar wander path of Gondwana

    Domeier, Mathew; van der Voo, Rob; Tohver, Eric; Tomezzoli, Renata N.; Vizan, Haroldo; Torsvik, Trond H.; Kirshner, Jordan


    The Late Paleozoic-Early Mesozoic apparent polar wander path of Gondwana is largely constructed from relatively old paleomagnetic results, many of which are considered unreliable by modern standards. Paleomagnetic results derived from sedimentary sequences, which are generally poorly dated and prone to inclination shallowing, are especially common. Here we report the results of a joint paleomagnetic-geochronologic study of a volcanic complex in central Argentina. U-Pb dating of zircons has yielded a robust age estimate of 263.0 +1.6/-2.0 Ma for the complex. Paleomagnetic analysis has revealed a pretilting (primary Permian) magnetization with dual polarities. Rock magnetic experiments have identified pseudo-single domain (titano)magnetite and hematite as the mineralogic carriers of the magnetization. Lightning-induced isothermal remagnetizations are widespread in the low-coercivity magnetic carriers. The resulting paleomagnetic pole is 80.1°S, 349.0°E, A95 = 3.3°, N = 35, and it improves a Late Permian mean pole calculated from a filtered South American paleomagnetic data set. More broadly, this new, high-quality, igneous-based paleomagnetic pole falls between the previously distinct Late Permian segments of the Laurussian and Gondwanan apparent polar wander paths, suggesting that the long-recognized disparity between these large paleomagnetic data sets may be primarily due to the inclusion of low-quality or systemically biased data.

  18. Stability of salt in the Permian salt basin of Kansas, Oklahoma, Texas and New Mexico, with a section on dissolved salts in surface water

    Bachman, George Odell; Johnson, Ross Byron


    The Permian salt basin in the Western Interior of the United States is defined as that region comprising a series of sedimentary basins in which halite and associated salts accumulated during Permian time. The region includes the western parts of Kansas, Oklahoma, and Texas, and eastern parts of Colorado and New Mexico. Following a long period of general tectonic stability throughout the region during most of early Paleozoic time, there was much tectonic activity in the area of the Permian salt basin during Late Pennsylvanian and Early Permian time just before bedded salt was deposited. The Early Permian tectonism was followed by stabilization of the basins in which the salt was deposited. These salt basins were neither contemporaneous nor continuous throughout the region, so that many salt beds are also discontinuous. In general, beds in the northern part of the basin (Kansas and northern Oklahoma) are older and the salt is progressively younger towards the south. Since Permian time the Permian salt basin has been relatively stable tectonically. Regionally, the area of the salt basin has been tilted and warped, has undergone periods of erosion, and has been subject to a major incursion of the sea; but deep-seated faults or igneous intrusions that postdate Permian salt are rare. In areas of the salt basin where salt is near the surface, such as southeastern New Mexico and central Kansas, there are no indications of younger deep-seated faulting and only a few isolated igneous intrusives of post-Permian age. On the other hand, subsidence or collapse of the land surface resulting from dissolution has been commonplace in the Permian salt basin. Some dissolution of salt deposits has probably been taking place ever since deposition of the salt more than 230 million years ago. Nevertheless, the subsurface dissolution fronts of the thick bedded-salt deposits of the Permian basin have retreated at a very slow average rate during that 230 million years. The preservation of

  19. Late Permian to Triassic intraplate orogeny of the southern Tianshan and adjacent regions, NW China

    Wei Ju; Guiting Hou


    The South Tianshan Orogen and adjacent regions of Central Asia are located in the southwestern part of the Central Asian Orogenic Belt. The formation of South Tianshan Orogen was a diachronous, scissors-like process, which took place during the Palaeozoic, and its western segment was accepted as a site of the final collision between the Tarim Craton and the North Asian continent, which occurred in the late Palaeozoic. However, the post-collisional tectonic evolution of the South Tianshan Orogen and adjacent regions remains debatable. Based on previous studies and recent geochronogical data, we suggest that the final collision between the Tarim Craton and the North Asian continent occurred during the late Carboniferous. Therefore, the Permian was a period of intracontinental environment in the southern Tianshan and adjacent regions. We propose that an earlier, small-scale intraplate orogenic stage occurred in late Permian to Triassic time, which was the first intraplate process in the South Tianshan Orogen and adjacent regions. The later large-scale and well-known Neogene to Quaternary intraplate orogeny was induced by the collision between the India subcontinent and the Eurasian plate. The paper presents a new evolutionary model for the South Tianshan Orogen and adjacent regions, which includes seven stages: (I) late Ordovicianeearly Silurian opening of the South Tianshan Ocean;(II) middle Silurianemiddle Devonian subduction of the South Tianshan Ocean beneath an active margin of the North Asian continent; (III) late Devonianelate Carboniferous closure of the South Tianshan Ocean and collision between the Kazakhstan-Yili and Tarim continental blocks;(IV) early Permian post-collisional magmatism and rifting;(V) late PermianeTriassic the first intraplate orogeny;(VI) JurassicePalaeogene tectonic stagnation and (VII) NeoceneeQuaternary intraplate orogeny.

  20. Rio carnival

    Alex Miller


      With the Rio 2016 Olympic and Paralympic Games due to start in August, Brazil is hoping that a financial crisis and a dramatic cost-cutting of the Games budget will all be forgotten once the two-week...

  1. Rio+20

    Robert Horn


    Full Text Available This reflection on Rio+20 examines many of the major social institutions and how they fulfilled their functions during the United Nations Conference on Sustainable Development at Rio. The institutions are: 1. Nation-states as a collective. 2. Individual nation-states. 3. Vanguard institutions (some NGOs. 4. Action and convening NGOs. 5. Global media. 6. Governments of nation-states acting domestically 7. Individual governments in bilateral and multilateral situations. 8. Similar institutions in different countries acting together. 9. Businesses. 10. Global science. Each is considered within the assumptions of what the society expects them to deliver (in general, what is possible for them to deliver, and what they did deliver at Rio. In approaching Rio+20, our account differs considerably from much of the reportage by the mainstream media.

  2. Grain-rimming kaolinite in Permian Rotliegend reservoir rocks

    Waldmann, Svenja; Gaupp, Reinhard


    Upper Rotliegend sediments of Permian age from the northeast Netherlands show moderate to good reservoir qualities. The predominant control is by the presence of authigenic grain-rimming kaolinite, which has a negative, but in some parts also a positive, effect on reservoir quality. To better understand the formation and distribution of grain-rimming kaolinite, reservoir rocks were studied in terms of composition and diagenetic processes. Petrographic evidence, summarized as a paragenetic sequence, is integrated with geochemical modeling results to identify early mesodiagenetic water-rock interactions under the participation of gases, i.e., CO2 and H2S, released from underlying Carboniferous source rocks. The sediments investigated were deposited at varying distance from the southern flank of the Southern Permian Basin. Sediments near the basin margin are mainly attributed to a fluvial environment and comprise medium to coarse-grained sandstones and conglomerates. There, vermicular kaolinite occurs with a lath-like structure. Distal to the basin margin, mainly in sandstones intercalated with fine-grained playa sediments, comparatively high amounts of grain-rimming kaolinite occur. There, the presence of this mineral has a significant influence on the rock properties and the reservoir quality. Geochemical modeling suggests that the formation of such kaolinites cannot be explained exclusively by in situ feldspar dissolution. The modeling results support evidence that kaolinite can be formed from precursor clay minerals under the presence of CO2-rich formation waters. Such clay minerals could be corrensite, smectite-chlorite mixed-layer minerals, or chlorite that is potentially present in Rotliegend sediments during early diagenesis. Furthermore, the geochemical modeling can reflect several mineral reactions that were identified from petrographic analysis such as the formation of illite and kaolinite at the expense of feldspar dissolution and consequent silica

  3. Comparative Earth history and Late Permian mass extinction

    Knoll, A. H.; Bambach, R. K.; Canfield, D. E.; Grotzinger, J. P.


    The repeated association during the late Neoproterozoic Era of large carbon-isotopic excursions, continental glaciation, and stratigraphically anomalous carbonate precipitation provides a framework for interpreting the reprise of these conditions on the Late Permian Earth. A paleoceanographic model that was developed to explain these stratigraphically linked phenomena suggests that the overturn of anoxic deep oceans during the Late Permian introduced high concentrations of carbon dioxide into surficial environments. The predicted physiological and climatic consequences for marine and terrestrial organisms are in good accord with the observed timing and selectivity of Late Permian mass extinction.

  4. Review of the Permian Camptoneuritidae (Insecta:Grylloblattida)

    Danil S.ARISTOV; Sergey Yu.STOROZHENKO; CUI Yingying


    The Permian family Camptoneuritidae (Insecta:Grylloblattida) is reviewed.New synonymies are proposed:Camptoneuritidae Martynov,1930=Demopteridae Carpenter,1950,syn.nov.,= Jabloniidae Kukalova,1964,syn.nov.Key to genera of Camptoneuritidae is given.All genera are redescribed and illustrated.Camptoneurites soyanensis sp.nov.from Soyana locality (Middle Permian,Kazanian Stage;Arkhangelsk Region,Russia) and Tyulkinia bashkuevi sp.nov.from Tyulkino locality (Lower Permian,Kungurian Stage;Perm Region,Russia) are described.

  5. The carbon and sulfur cycles and atmospheric oxygen from middle Permian to middle Triassic

    Berner, Robert A.


    The results of a theoretical isotope mass balance model are presented for the time dependence of burial and weathering-plus-degassing fluxes within the combined long-term carbon and sulfur cycles. Averaged data for oceanic δ 13C and δ 34S were entered for every million years from 270 to 240 Ma (middle Permian to middle Triassic) to study general trends across the Permian-Triassic boundary. Results show a drop in the rate of global organic matter burial during the late Permian and a predominance of low values during the early-to-middle Triassic. This overall decrease with time is ascribed mainly to epochs of conversion of high biomass forests to low biomass herbaceous vegetation resulting in a decrease in the production of terrestrially derived organic debris. Additional contributions to lessened terrestrial carbon burial were increased aridity and a drop in sea level during the late Permian which led to smaller areas of low-lying coastal wetlands suitable for coal and peat deposition. Mirroring the drop in organic matter deposition was an increase in the burial of sedimentary pyrite, and a dramatic increase in the calculated global mean ratio of pyrite-S to organic-C. High S/C values resulted from an increase of deposition in marine euxinic basins combined with a decrease in the burial of low-pyrite associated terrestrial organic matter. The prediction of increased oceanic anoxia during the late Permian and early Triassic agrees with independent studies of the composition of sedimentary rocks. Weathering plus burial fluxes for organic carbon and pyrite sulfur were used to calculate changes in atmospheric oxygen. The striking result is a continuous drop in O 2 concentration from ˜30% to ˜13% over a twenty million year period. This drop was brought about mainly by a decrease in the burial of terrestrially derived organic matter. but with a possible contribution from the weathering of older organic matter on land. It must have exerted a considerable influence on

  6. Social anxiety and negative early life events in university students Eventos negativos na infância e ansiedade social em estudantes universitários

    Cynthia Binelli


    Full Text Available INTRODUCTION: There is substantial evidence regarding the impact of negative life events during childhood on the aetiology of psychiatric disorders. We examined the association between negative early life events and social anxiety in a sample of 571 Spanish University students. METHODS: In a cross-sectional survey conducted in 2007, we collected data through a semistructured questionnaire of sociodemographic variables, personal and family psychiatric history, and substance abuse. We assessed the five early negative life events: (i the loss of someone close, (ii emotional abuse, (iii physical abuse, (iv family violence, and (v sexual abuse. All participants completed the Liebowitz Social Anxiety Scale. RESULTS: Mean (SD age was 21 (4.5, 75% female, LSAS score was 40 (DP = 22, 14.2% had a psychiatric family history and 50.6% had negative life events during childhood. Linear regression analyses, after controlling for age, gender, and family psychiatric history, showed a positive association between family violence and social score (p = 0.03. None of the remaining stressors produced a significant increase in LSAS score (p > 0.05. CONCLUSION: University students with high levels of social anxiety presented higher prevalence of negative early life events. Thus, childhood family violence could be a risk factor for social anxiety in such a population.INTRODUÇÃO: Existem evidências substanciais sobre o impacto de eventos negativos da vida durante a infância na etiologia dos transtornos psiquiátricos. Examinamos a associação entre os eventos negativos ocorridos na infância e a ansiedade social em uma amostra de 571 estudantes universitários espanhóis. MÉTODOS: Em um estudo transversal realizado em 2007, foram coletados os dados de variáveis sociodemográficas, história psiquiátrica pessoal e familiar e abuso de substâncias por meio de um questionário semiestruturado e avaliamos cinco eventos negativos ocorridos na infância: (i a perda de

  7. New data on Upper Carboniferous–Lower Permian deposits of Bol'shevik Island, Severnaya Zemlya Archipelago

    Victoria B. Ershova


    Full Text Available We present here a detailed study of the Upper Carboniferous–Lower Permian stratigraphy of Bol'shevik Island in the Severnaya Zemlya Archipelago, consisting of the analysis of sedimentary structures and lithostratigraphy, U/Pb detrital zircon dating and structural studies. The preserved sedimentary structures suggest that the studied strata were deposited in a relatively small meandering fluvial system. U/Pb dating of detrital zircons reveals that the Upper Carboniferous–Lower Permian sandstones contain a primary age population ranging from 450 to 570 millions of years, with a predominance of Early–Middle Ordovician zircons. This detrital zircon distribution indicates that the studied formations were derived locally from the erosion of Lower Ordovician deposits of Bol'shevik Island or elsewhere in the archipelago. Our structural studies suggest that Upper Carboniferous–Lower Permian deposits are deformed into a series of west–north-west verging open asymmetric folds, suggesting a west–north-west direction of tectonic transport and that deformation across the island is post-Early Permian in age.

  8. The Luoping biota: exceptional preservation, and new evidence on the Triassic recovery from end-Permian mass extinction.

    Hu, Shi-xue; Zhang, Qi-yue; Chen, Zhong-Qiang; Zhou, Chang-yong; Lü, Tao; Xie, Tao; Wen, Wen; Huang, Jin-yuan; Benton, Michael J


    The timing and nature of biotic recovery from the devastating end-Permian mass extinction (252 Ma) are much debated. New studies in South China suggest that complex marine ecosystems did not become re-established until the middle-late Anisian (Middle Triassic), much later than had been proposed by some. The recently discovered exceptionally preserved Luoping biota from the Anisian Stage of the Middle Triassic, Yunnan Province and southwest China shows this final stage of community assembly on the continental shelf. The fossil assemblage is a mixture of marine animals, including abundant lightly sclerotized arthropods, associated with fishes, marine reptiles, bivalves, gastropods, belemnoids, ammonoids, echinoderms, brachiopods, conodonts and foraminifers, as well as plants and rare arthropods from nearby land. In some ways, the Luoping biota rebuilt the framework of the pre-extinction latest Permian marine ecosystem, but it differed too in profound ways. New trophic levels were introduced, most notably among top predators in the form of the diverse marine reptiles that had no evident analogues in the Late Permian. The Luoping biota is one of the most diverse Triassic marine fossil Lagerstätten in the world, providing a new and early window on recovery and radiation of Triassic marine ecosystems some 10 Myr after the end-Permian mass extinction.

  9. Timing of the terrestrial Permian-Triassic boundary biotic crisis: Implications from U-Pb dating of authigenic zircons

    YU JianXin; LI HuiMin; ZHANG SuXin; YANG FengQing; FENG QingLai


    The Late Permian to Early Triassic transition represents one of the most important Phanerozoic mass extinction episodes.The cause of this event is still in debate between catastrophic and gradual mechanisms.This study uses the U-Pb method on zircons from the uppermost Permian/lowermost Triassic clay deposits at Chahe (Guizhou Province,SW China) to examine time constraints for this event.The results of both this and previous studies show that the ages of Bed 68a and 68c (the upper clay bed of the terrestrial Permian-Triassic boundary (PTB)) respectively are 252.6~2.8 and 247.5±2.8 Ma.This age (within the margin of error) almost accords with the upper clay bed (Bed 28) age of Meishan and the eruption age of Tunguss Basalt,and is so far the most accurate age obtained from terrestrial PTB.The claystone of Bed 68 was formed in the earliest Triassic.The biotic crisis occurred at nearly the same time in terrestrial and marine environments during Permian-Triassic interval; however the extinction patterns and processes are different.The extinction pattern of the terrestrial plants shows a major decline at the PTB after long-term evolution,followed by a retarded extinction of the relicts in the earliest Triassic.

  10. Timing of the terrestrial Permian-Triassic boundary biotic crisis:Implications from U-Pb dating of authigenic zircons


    The Late Permian to Early Triassic transition represents one of the most important Phanerozoic mass extinction episodes. The cause of this event is still in debate between catastrophic and gradual mechanisms. This study uses the U-Pb method on zircons from the uppermost Permian/lowermost Triassic clay deposits at Chahe (Guizhou Province, SW China) to examine time constraints for this event. The results of both this and previous studies show that the ages of Bed 68a and 68c (the upper clay bed of the terrestrial Permian-Triassic boundary (PTB)) respectively are 252.6±2.8 and 247.5±2.8 Ma. This age (within the margin of error) almost accords with the upper clay bed (Bed 28) age of Meishan and the eruption age of Tunguss Basalt, and is so far the most accurate age obtained from terrestrial PTB. The claystone of Bed 68 was formed in the earliest Triassic. The biotic crisis occurred at nearly the same time in terrestrial and marine environments during Permian-Triassic interval; however the extinction patterns and processes are different. The extinction pattern of the terrestrial plants shows a major decline at the PTB after long-term evolution, followed by a retarded extinction of the relicts in the earliest Triassic.

  11. Stratigraphy, biostratigraphy and C-isotopes of the Permian-Triassic non-marine sequence at Dalongkou and Lucaogou, Xinjiang Province, China

    Metcalfe, I.; Foster, C. B.; Afonin, S. A.; Nicoll, R. S.; Mundil, R.; Xiaofeng, Wang; Lucas, S. G.


    Measured lithostratigraphic sections of the classic Permian-Triassic non-marine transitional sequences covering the upper Quanzijie, Wutonggou, Guodikeng and lower Jiucaiyuan Formations at Dalongkou and Lucaogou, Xinjiang Province, China are presented. These measured sections form the framework and reference sections for a range of multi-disciplinary studies of the P-T transition in this large ancient lake basin, including palynostratigraphy, vertebrate biostratigraphy, chemostratigraphy and magnetostratigraphy. The 121 m thick Wutonggou Formation at Dalongkou includes 12 sandstone units ranging in thickness from 0.5 to 10.5 m that represent cyclical coarse terrigenous input to the lake basin during the Late Permian. The rhythmically-bedded, mudstone-dominated Guodikeng Formation is 197 m and 209 m thick on the north and south limbs of the Dalongkou anticline, respectively, and 129 m thick at Lucaogou. Based on limited palynological data, the Permian-Triassic boundary was previously placed approximately 50 m below the top of this formation at Dalongkou. This boundary does not coincide with any mappable lithologic unit, such as the basal sandstones of the overlying Jiucaiyuan Formation, assigned to the Early Triassic. The presence of multiple organic δ13C-isotope excursions, mutant pollen, and multiple algal and conchostracan blooms in this formation, together with Late Permian palynomorphs, suggests that the Guodikeng Formation records multiple climatic perturbation signals representing environmental stress during the late Permian mass extinction interval. The overlap between the vertebrates Dicynodon and Lystrosaurus in the upper part of this formation, and the occurrence of late Permian spores and the latest Permian to earliest Triassic megaspore Otynisporites eotriassicus is consistent with a latest Permian age for at least part of the Guodikeng Formation. Palynostratigrahic placement of the Permian-Triassic boundary in the Junggar Basin remains problematic

  12. Calibrating the end-Permian mass extinction.

    Shen, Shu-zhong; Crowley, James L; Wang, Yue; Bowring, Samuel A; Erwin, Douglas H; Sadler, Peter M; Cao, Chang-qun; Rothman, Daniel H; Henderson, Charles M; Ramezani, Jahandar; Zhang, Hua; Shen, Yanan; Wang, Xiang-dong; Wang, Wei; Mu, Lin; Li, Wen-zhong; Tang, Yue-gang; Liu, Xiao-lei; Liu, Lu-jun; Zeng, Yong; Jiang, Yao-fa; Jin, Yu-gan


    The end-Permian mass extinction was the most severe biodiversity crisis in Earth history. To better constrain the timing, and ultimately the causes of this event, we collected a suite of geochronologic, isotopic, and biostratigraphic data on several well-preserved sedimentary sections in South China. High-precision U-Pb dating reveals that the extinction peak occurred just before 252.28 ± 0.08 million years ago, after a decline of 2 per mil (‰) in δ(13)C over 90,000 years, and coincided with a δ(13)C excursion of -5‰ that is estimated to have lasted ≤20,000 years. The extinction interval was less than 200,000 years and synchronous in marine and terrestrial realms; associated charcoal-rich and soot-bearing layers indicate widespread wildfires on land. A massive release of thermogenic carbon dioxide and/or methane may have caused the catastrophic extinction.

  13. Consumo de refrigerantes entre escolares de séries iniciais da cidade de Pelotas, Rio Grande do Sul = Consumption of soft drinks among students from early grades in the city of Pelotas, Rio Grande do Sul

    Flores, Thaynã Ramos


    Full Text Available Objetivo: Descrever o consumo diário de refrigerantes entre escolares do 1° ao 4° ano da rede municipal de ensino fundamental da cidade de Pelotas, Rio Grande do Sul. Materiais e Métodos: Foi realizado um estudo transversal, de abordagem quantitativa, com 356 escolares do 1° ao 4° ano matriculados em escolas da zona urbana do município, no ano de 2011. O desfecho deste estudo foi o consumo diário de refrigerantes, avaliado através do Formulário de Marcadores de Consumo Alimentar proposto pelo Sistema de Vigilância Alimentar e Nutricional. A frequência de consumo foi descrita segundo gênero, idade, escolaridade materna, nível socioeconômico e estado nutricional do escolar. Foram consideradas as prevalências e os seus respectivos intervalos de confiança de 95%, adotando-se o nível de significância estatística de 5%. Resultados: Dos 411 escolares considerados elegíveis para o estudo, foram avaliados 356, sendo de 13,4% as perdas e recusas. A prevalência de consumo diário de refrigerantes foi de 21,6%. Não foram encontradas diferenças estatisticamente significativas de consumo diário de refrigerantes entre os sexos, faixa etária, classes de bens de consumo e estado nutricional. Conclusão: Os escolares da rede municipal de ensino fundamental de Pelotas apresentaram elevada prevalência de consumo diário de refrigerantes. Diante esse quadro, ressalta-se a importância da educação nutricional no ambiente escolar


    Monica; J.Campi; 石光荣


    A new genus and species, Linshuichonetes elfinis, belonging to the Rugosochonetidae, is described from the Early Permian (Late Artinskian or early Kungurian) Liangshan Formation of the Yangtze block. The new genus is defined externally by the presence of fine, but delayed, capillation and a weak or absent median sulcus and fold and by the presence of a distinct posteromedian sinus on the ventral umbo; and internally by a lack of median, lateral and accessory septa in the dorsal interior; absence of vascular trunks in the ventral interior and the presence of distinct radiating rows of papillae in the interior of both valves, particularly an unusual clustered arrangement of papillae on the posteromedian portion of the dorsal interior. The local environment during the deposition of the Liangshan Formation appears to have been a restricted tidal flat or lagoon which experienced frequent sealevel fluctuations associated with the onset of the Yanghsingian transgression. The new species, L. elfinis, appears to have several morphological adaptations enabling successful exploitation of this environment. It was typically a very small and thin-valved species with a high surface area to volume ratio, an advantage in an oxygen restricted environment. The small size and numerous body spinules would have aided individuals to remain suspended at the top of the fine, soft substrate. It also dominated the brachiopod assemblage in the Liangshan Formation, comprising up to 94% of specimens within a bed. These factors indicate that the new species appears to be an opportunistic species.%描述了产于四川邻水地区早二叠世梁山组的一新属新种,Linshuichonetes elfinis gen. et sp.nov.。新属以个体特别微小,背内无任何隔板,壳表饰有细瘤和细刺为特征。新种代表了华南地区阳新期海侵开始阶段生活于局限的潮坪或湖环境下的一种机会种。

  15. Assessment of Hydrocarbon Generation Potential of Permian Gondwana Coals, Bangladesh

    H. M. Zakir Hossain


    Full Text Available This paper represents the geochemical characteristics of Gondwana coals from the Barapukuria coal mine, Bangladesh in order to investigate the potential for hydrocarbon generation. A total number of twenty three coal samples were analyzed Rock-Eval pyrolysis, CHNS elemental analyses, maceral analysis and vitrinite reflectance. The samples were collected from drill hole GDH-40 of the Barapukuria coal mine encountered within Gondwana succession of Permian age. The TOC contents of the coal samples range between ~50 and 76 wt.% and the organic matter consists predominantly of type III and type IV kerogen with respect to hydrocarbon generation. The GP, HI, PI and Tmax values range between 7 and 35 mg HC/g rock, 20 and 62 mg HC/g TOC, 0.02 and 0.04, and 430 and 437oC, respectively. The organic matter is mainly gas prone and thermally immature to early mature level. The potential coal bed methane (CBM generation of the Barapukuria basin is estimated to be 11 Gm3. Thus, underground coal gasification (UCG is helpful for better development of subsurface coals at the Barapukuria basin, Bangladesh.

  16. Reply to the comments on: ;From source-to-sink: The Late Permian SW gondwana paleogeography and sedimentary dispersion unraveled by a multi-proxy analysis; [journal of South American earth sciences 70 (2016) 368-382] by Vesely & Schemiko

    Alessandretti, Luciano; Warren, Lucas Veríssimo; Assine, Mario Luis; Machado, Rômulo; Lana, Cristiano


    The publication of the comments by Vesely & Schemiko (Comment on ;From source-to-sink: The Late Permian SW Gondwana paleogeography and sedimentary dispersion unraveled by a multi-proxy analysis; by L. Alessandretti, R. Machado, L.V. Warren, M.L. Assine and C. Lana [Journal of South American Earth Sciences 70 (2016) 368-382], Journal of South American Earth Sciences, this issue) on our paper entitled ;From source-to-sink: The Late Permian SW Gondwana paleogeography and sedimentary dispersion unraveled by a multi-proxy analysis; (L. Alessandretti, R. Machado, L.V. Warren, M.L. Assine and C. Lana (Journal of South American Earth Sciences 70 (2016) 368-382) provides a worthy opportunity to further clarify our observations and interpretations regarding the provenance of the Late Permian Rio do Rasto Formation and its implications on SW Gondwana paleogeography and sedimentary dispersion.

  17. Thuringopteryx- eine ,,permische" Eintagsfliege im Buntsandstein (Insecta, Pterygota) Thuringopteryx, a "Permian" mayfly in the Buntsandstein (Insecta, Pterygota)


    The Misthodotida have been known from the Permian only. However, Thuringopteryx gimmi KUHN, 1937 from the middle Buntsandstein of Thuringia, Germany, usually considered to be a representative of the Odonatoptera, Palaeodictyptera or Saltatoria-Ensifera, belongs in the stem group of the mayflies, sharing with the Misthodotida a one-branched CuA and with Misthodotes a short Sc. Hence, the Misthodotida survived at least until the early Mesozoic. The phylogenetic relationships of Mist...

  18. Coricladus quiteriensis gen. et sp. nov., a new conifer in Southern-Brazil Gondwana (Lower Permian, Paraná Basin

    Jasper André


    Full Text Available A new taxon of conifers (Coricladus quiteriensis is described based on megafloristic remains from the roofshale level at the Quitéria Outcrop (Rio Bonito Formation - Lower Permian - Southern Paraná Basin - Rio Grande do Sul - Brazil. This megafloristic community is included in the Botrychiopsis Zone - Botrychiopsis valida Sub-Zone (Kungurian/Roadian. The assemblage, preserved as impressions, do not present remains of epidermic characters, and is composed mainly of isolated vegetative branches with spirally disposed acicular leaves, presenting a conspicuous central vein and also isolated fertile branches with sparse and irregular leaves and terminal cones. Leafless principal branches, organically connected with sterile and fertile branches, are rare. Reproductive feminine scales, disposed in a plane, are organized in lax terminal cones on branches, composed by 4 (four distal ovuliferous scales, and 8 (eight elliptical-elongated anatropous seeds. Paleoecological data pointed out to a mesophylous to higrophylous habitat in swampy environments.

  19. Delayed recovery of non-marine tetrapods after the end-Permian mass extinction tracks global carbon cycle.

    Irmis, Randall B; Whiteside, Jessica H


    During the end-Permian mass extinction, marine ecosystems suffered a major drop in diversity, which was maintained throughout the Early Triassic until delayed recovery during the Middle Triassic. This depressed diversity in the Early Triassic correlates with multiple major perturbations to the global carbon cycle, interpreted as either intrinsic ecosystem or external palaeoenvironmental effects. In contrast, the terrestrial record of extinction and recovery is less clear; the effects and magnitude of the end-Permian extinction on non-marine vertebrates are particularly controversial. We use specimen-level data from southern Africa and Russia to investigate the palaeodiversity dynamics of non-marine tetrapods across the Permo-Triassic boundary by analysing sample-standardized generic richness, evenness and relative abundance. In addition, we investigate the potential effects of sampling, geological and taxonomic biases on these data. Our analyses demonstrate that non-marine tetrapods were severely affected by the end-Permian mass extinction, and that these assemblages did not begin to recover until the Middle Triassic. These data are congruent with those from land plants and marine invertebrates. Furthermore, they are consistent with the idea that unstable low-diversity post-extinction ecosystems were subject to boom-bust cycles, reflected in multiple Early Triassic perturbations of the carbon cycle.

  20. The youngest trigonotarbid Permotarbus schuberti n. gen., n. sp. from the Permian Petrified Forest of Chemnitz in Germany

    J. A. Dunlop


    Full Text Available A new trigonotarbid (Arachnida: Trigonotarbida is described as Permotarbus schuberti n. gen., n. sp. from the Early Permian Petrified Forest (Rotliegend of Chemnitz in Saxony (Germany. At ca. 290 Ma it represents the youngest record of this extinct arachnid order discovered to date. Its familial affinities are uncertain, but may lie close to the Aphantomartidae. The distribution of the trigonotarbid genera through time is summarised, together with a list of their seventy-seven fossil-yielding localities. Together they offer a broad overview of the group's fossil record, which is heavily biased towards the Moscovian Stage (ca. 307–312 Ma of the Late Carboniferous in Europe and North America. This is due in no small part to numerous localities associated with coal mining districts, and trigonotarbids are found less frequently after this stage. While it is tempting to associate this with biological events – such as a putative "Carboniferous Rainforest Collapse" dating to ca. 305 Ma – it is difficult to differentiate the effects of genuine extinction patterns from artefacts caused by fewer appropriate localities in the economically less relevant latest Carboniferous and Early Permian strata. Nevertheless, trigonotarbids became extinct at some point after the Early Permian and loss of the Coal Measures forests remains one of the most likely possible causes. doi:10.1002/mmng.201300012

  1. Trace fossil evidence for late Permian shallow water condition in Guryul ravine, Kashmir, India

    Parcha, Suraj; Horacek, Micha; Krystyn, Leopold; Pandey, Shivani


    The present study is focused on the Late Permian (Changhsingian) succession, present in the Guryul ravine, Kashmir Basin. The basin has a complete Cambro-Triassic sequence and thus contains a unique position in the geology of Himalaya. The Guryul Ravine Permian mainly comprises of mixed siliciclastic-carbonate sediments deposited in a shallow-shelf or ramp setting. The present assemblage of Ichnofossils is the first significant report of trace fossils in the Guryul ravine since early reports in the 1970s. The Ichnofossils reported from this section include: Diplichnites, Dimorphichnus, Monomorphichnus, Planolites, Skolithos along with burrow, scratch marks and annelid worm traces?. The ichnofossils are mainly preserved in medium grain sandstone-mudstone facies. The Ichnofossils are widely distributed throughout the section and are mostly belonging to arthropods and annelid origin, showing behavioral activity, mainly dwelling and feeding, and evidence the dominant presence of deposit feeders. The vertical to slightly inclined biogenic structures are commonly recognized from semi-consolidated substrate which are characteristic features of the near shore/foreshore marine environment, with moderate to high energy conditions. The topmost layer of silty shale contains trace fossils like Skolithos and poorly preserved burrows. The burrow material filled is same as that of host rock. The studied Zewan C and D sequence represents the early to late part of the Changhsingian stage, from 40 to 5 m below the top of Zewan D member with bioturbation still evident in some limestone layers till 2 metres above. No trace fossils could be recognized in the topmost 3 m beds of Zewan D due to their gliding related amalgamated structure. The widespread distribution of traces and their in situ nature will be useful for interpretation of the paleoecological and paleoenvironmental conditions during the late Permian in the Guryul ravine of Kashmir.




    Full Text Available The Permian to Triassic transition in Jordan is characterised by a sequence boundary underlain by red-bed, alluvial lithofacies deposited in a humid-tropical climate by low-sinuosity rivers, and overlain by shallow marine siliciclastics with thin carbonates. The low-gradient alluvial floodplain was repeatedly subjected to the development of ferralitic and pisolitic paleosols on the interfluves. In contrast, dysaerobic environments in the fluvial channels and abandoned lakes resulted in the preservation of a prolific flora of macro-plants and palynomorphs that indicate a probable range from Mid- to Late Permian age, though the abundant presence of the distinctive pollen Pretricolpipollenites bharadwajii  indicates the youngest part of that range.  Above the sequence boundary, reddened shallow-marine beds characterised by ripple cross-laminated, siltstones/sandstone with desiccation cracks and sparse surface burrows mark the initial Triassic marine transgression in the region (Arabian Plate Tr 10. These are followed by two thin limestone (packstone beds with shallow scours and bivalve shell lags, that have yielded a low diversity assemblage of conodonts (e.g. Hadrodontina aequabilis and foraminifera (“Cornuspira” mahajeri that are interpreted as euryhaline  taxa characterising the early Induan (Early Triassic. Thus the absence of body fossils and vertical infaunal burrows in the lowest marine beds may reflect low-diversity ecosystems following the Permian-Triassic extinction event, or be a result of stressed shallow marine environments. A gradational upward increase in grey, green and yellow siltstones beds accompanied by a concomitant increase in bioturbation (and infaunal vertical burrows and thin-shelled bivalves about 15 m above the boundary indicates colonisation of the substrate under more normal shallow marine conditions perhaps indicating recovery phase following the extinction event.

  3. Two episodic changes of trace fossils through the Permian-Triassic transition in the Meishan cores,Zhejiang Province


    Six ichnogenera,Chondrites,Palaeophycus,Planolites,Rhizocorallium,Skolithos,and Thalassinoides,were identified from the drilling cores near the Global Stratotype Section and Point(GSSP) of Permian-Triassic boundary at Meishan,Zhejiang Province.The ichnological indices,including abundance,bioturbation index,and disturbed depth of trace fossils,show two episodes of intense changes near the Permian-Triassic Boundary(PTB).Episode I occurred in Beds 25-27b when the ecologically complicate forms such as Chondrites,Skolithos,Rhizocorallium,and Thalassinoides disappeared hereafter,the bioturbation index reduced from 1-5 to 1-3,and the disturbed depth declined from 5-66 to 2-5 cm.Episode II took place at the base of Bed 33 with the disappearance of Palaeophycus and Planolites,and subsequent absence of trace fossils and bioturbation till the middle-upper part of Bed 41 when the disturbed structures reoccurred,but they are only tiny Planolites and the bioturbation index was never higher than 3 and the disturbed depth less than 4 mm.Episode I shows an intense change,corresponding to the main stage of the end-Permian mass extinction,whereas Episode II is relatively weak,corresponding to the epilogue of the mass extinction of trace makers in the Early Triassic.Subsequently,ichnofossils were dominated by surface tracks in simple ecological habit and structures.This phenomenon indicates that the Early Triassic benthonic fauna is changed from sessile benthic system to mobile benthic system after the end-Permian mass extinction.In other words,the evolution of the trace fossils across the Permian-Triassic transition had an episodic process similar to the body fossils.In addition,the change of ichnofabrics is well coincided with the negative excursion of carbon isotopes and the expansion of cyanobacteria.As the results of physical and biogenic processes,trace fossils provided unique materials for the study of the biotic and environmental events,as well as their coupling evolution




    Full Text Available An amphibian fauna from Permo-Carboniferous boundary beds is recorded for the first time in Italy. A thin fossiliferous level has been found in the Perdasdefogu Basin in southeastern Sardinia;it yields several speciments of Branchiosaurus cf."B." petrolei Gaudry 1875, often in mass mortality assemblages.Repeated mass mortality events testify to sudden changes in the environment of the basin, possibly due to seasonal variations. The finding of speciments very close to Branchiosaurus petrolei,which is a common species in the Central France basins,confirms that Sardinia at the time belonged to the same hydrographic basin of continental Europe, with no seaway in between.Furthermore, though not the primary focus of this note, we report the first discovery of the xenacanth teeth and acanthodian spines in Italy.  

  5. Permian paleoclimate data from fluid inclusions in halite

    Benison, K.C.; Goldstein, R.H.


    This study has yielded surface water paleotemperatures from primary fluid inclusions in mid Permian Nippewalla Group halite from western Kansas. A 'cooling nucleation' method is used to generate vapor bubbles in originally all-liquid primary inclusions. Then, surface water paleotemperatures are obtained by measuring temperatures of homogenization to liquid. Homogenization temperatures ranged from 21??C to 50??C and are consistent along individual fluid inclusion assemblages, indicating that the fluid inclusions have not been altered by thermal reequilibration. Homogenization temperatures show a range of up to 26??C from base to top of individual cloudy chevron growth bands. Petrographic and fluid inclusion evidence indicate that no significant pressure correction is needed for the homogenization temperature data. We interpret these homogenization temperatures to represent shallow surface water paleotemperatures. The range in temperatures from base to top of single chevron bands may reflect daily temperatures variations. These Permian surface water temperatures fall within the same range as some modern evaporative surface waters, suggesting that this Permian environment may have been relatively similar to its modern counterparts. Shallow surface water temperatures in evaporative settings correspond closely to local air temperatures. Therefore, the Permian surface water temperatures determined in this study may be considered proxies for local Permian air temperatures.

  6. Middle-Late Permian mass extinction on land

    Retallack, G.J.; Metzger, C.A.; Greaver, T.; Jahren, A.H.; Smith, R.M.H.; Sheldon, N.D. [University of Oregon, Eugene, OR (United States). Dept. of Geological Science


    The end-Permian mass extinction has been envisaged as the nadir of biodiversity decline due to increasing volcanic gas emissions over some 9 million years. We propose a different tempo and mechanism of extinction because we recognize two separate but geologically abrupt mass extinctions on land, one terminating the Middle Permian (Guadalupian) at 260.4 Ma and a later one ending the Permian Period at 251 Ma. Our evidence comes from new paleobotanical, paleopedological, and carbon isotopic studies of Portal Mountain, Antarctica, and comparable studies in the Karoo Basin, South Africa. Extinctions have long been apparent among marine invertebrates at both the end of the Guadalupian and end of the Permian, which were also times of warm-wet greenhouse climatic transients, marked soil erosion, transition from high- to low-sinuosity and braided streams, soil stagnation in wetlands, and profound negative carbon isotope anomalies. Both mass extinctions may have resulted from catastrophic methane outbursts to the atmosphere from coal intruded by feeder dikes to flood basalts, such as the end-Guadalupian Emeishan Basalt and end-Permian Siberian Traps.

  7. Timing of global regression and microbial bloom linked with the Permian-Triassic boundary mass extinction: implications for driving mechanisms

    Baresel, Björn; Bucher, Hugo; Bagherpour, Borhan; Brosse, Morgane; Guodun, Kuang; Schaltegger, Urs


    New high-resolution U-Pb dates indicate a duration of 89 ± 38 kyr for the Permian hiatus and of 14 ± 57 kyr for the overlying Triassic microbial limestone in shallow water settings of the Nanpanjiang Basin, South China. The age and duration of the hiatus coincides with the Permian-Triassic boundary (PTB) and the extinction interval in the Meishan Global Stratotype Section and Point, and strongly supports a glacio-eustatic regression, which best explains the genesis of the worldwide hiatus straddling the PTB in shallow water records. In adjacent deep marine troughs, rates of sediment accumulation display a six-fold decrease across the PTB compatible with a dryer and cooler climate as indicated by terrestrial plants. Our model of the Permian-Triassic boundary mass extinction (PTBME) hinges on the synchronicity of the hiatus with the onset of the Siberian Traps volcanism. This early eruptive phase released sulfur-rich volatiles into the stratosphere, thus simultaneously eliciting a short-lived ice age responsible for the global regression and a brief but intense acidification. Abrupt cooling, shrunk habitats on shelves and acidification may all have synergistically triggered the PTBME. Subsequently, the build-up of volcanic CO2 induced a transient cool climate whose early phase saw the deposition of the microbial limestone.

  8. Timing of global regression and microbial bloom linked with the Permian-Triassic boundary mass extinction: implications for driving mechanisms

    Baresel, Björn; Bucher, Hugo; Bagherpour, Borhan; Brosse, Morgane; Guodun, Kuang; Schaltegger, Urs


    New high-resolution U-Pb dates indicate a duration of 89 ± 38 kyr for the Permian hiatus and of 14 ± 57 kyr for the overlying Triassic microbial limestone in shallow water settings of the Nanpanjiang Basin, South China. The age and duration of the hiatus coincides with the Permian-Triassic boundary (PTB) and the extinction interval in the Meishan Global Stratotype Section and Point, and strongly supports a glacio-eustatic regression, which best explains the genesis of the worldwide hiatus straddling the PTB in shallow water records. In adjacent deep marine troughs, rates of sediment accumulation display a six-fold decrease across the PTB compatible with a dryer and cooler climate as indicated by terrestrial plants. Our model of the Permian-Triassic boundary mass extinction (PTBME) hinges on the synchronicity of the hiatus with the onset of the Siberian Traps volcanism. This early eruptive phase released sulfur-rich volatiles into the stratosphere, thus simultaneously eliciting a short-lived ice age responsible for the global regression and a brief but intense acidification. Abrupt cooling, shrunk habitats on shelves and acidification may all have synergistically triggered the PTBME. Subsequently, the build-up of volcanic CO2 induced a transient cool climate whose early phase saw the deposition of the microbial limestone. PMID:28262815

  9. The oldest post-Palaeozoic Crinoid and Permian-Triassic origins of the Articulata (Echinodermata).

    Oji, Tatsuo; Twitchett, Richard J


    The Crinoidea are the most primitive class of living echinoderms, and suffered a severe crisis during the Late Permian mass extinction event. All post-Palaeozoic crinoids, including living species, belong to the Articulata, and morphological and recent molecular studies demonstrate that they form a monophyletic clade. The Articulata originated from Palaeozoic cladid crinoids, but the nature and timing of their origination remains obscure. Problems with understanding the origin and early evolution of the Articulata have arisen because the Permian-Triassic crinoid fossil record is particularly poor. We report on a new genus and species from the earliest Triassic, which is the oldest known post-Palaeozoic articulate crinoid and fundamentally alters our understanding of the early evolution of the Articulata. Prior to this study, the most primitive post-Palaeozoic articulate was thought to be Holocrinus of the order Isocrinida. Unexpectedly, the new taxon belongs to the order Encrinida, which reveals a previously hidden diversity of crinoids in the earliest Triassic. Its discovery implies either a dramatic radiation of crinoids in the immediate post-extinction aftermath, when environmental conditions were at their most severe, or a pre-extinction origin of the crown group articulates and survival of multiple lineages.

  10. Diagenesis and Fluid Flow History in Sandstones of the Upper Permian Black Jack Formation, Gunnedah Basin, Eastern Australia

    BAI Guoping; John B. KEENE


    The fluid flow history during diagenesis of sandstones in the Upper Permian Black Jack Formation of the Gunnedah Basin has been investigated through integrated petrographic observations, fluid inclusion investigations and stable isotope analyses. The early precipitation of mixed-layer illite/smectite, siderite, calcite, ankerite and kaolin proceeded at the presence of Late Permian connate meteoric waters at temperatures of up to 60℃. These evolved connate pore waters were also parental to quartz, which formed at temperatures of up to 87℃. The phase of maximum burial was characterized by development of filamentous illite and late calcite at temperatures of up to ~90℃. Subsequent uplifting and cooling led to deep meteoric influx from surface, which in turn resulted in dissolution of labile grains and carbonate cements, and formation of second generation of kaolin. Dawsonite was the last diagenetic mineral precipitated and its formation is genetically related to deep-seated mamagtic sourced CO2.

  11. 吉林延边地区汪清县大兴沟早二叠世华夏植物群的发现及其地质意义%Discovery of Early Permian Cathysia Flora from Daxinggou of Wangqing County, Yanbian Area, Jilin Province, China and Its Geological Significance

    李明松; 孙跃武; 赵国伟


    nauxii, Pecopteris huichensis and P. marginata; whereas there are no typical Angara elements, such as Angaridiurn, Cardioneura, Rufloria and "Neoggerathiopsis". Therefore, the Hesheng Flora is definitely of Cathaysia floras in nature. Since most species of the Hesheng Flora are common in the Taiyuan and Shanxi formations in northern China, and some are even only found in the Shanxi and Xiashihezi formations in northern China such as Sphenopteris rotunda, Pecopteris marginata , P. yabei , Odontopteris subcrenulata , Taeniopteris yernauxii and Paracalamites stenocostatus, the Hesheng Flora is considered as late Cisuralian of Permian (Artinskian-early Kungurian) in age.The discovery of this Cisuralian flora shows that the distribution of Cathaysia flora should extend northward to the Daxinggou district in Wangqing County, and that the boundary between the Cathaysia flora and the Angara flora is probably located in a narrow northwestward zone geographically ranged from the Kaishantun of Yanji City-Daxinggou of Wangqing County to the Jiefangcun of Hunchun City-Shiliping of Wangqing County. This is beneficial not only for the study on the boundary between North China Plate and Jiamusi-Mongolia Block, but also for the research on the eastern part of Xar Moron River suture zone in Yanbian area on the paleobotanical basis.

  12. An unusual, structurally preserved ovule from the Permian of Antarctica.

    Klavins, S D.; Taylor, E L.; Krings, M; Taylor, T N.


    Anatomically preserved ovules are described from silicified peat of Late Permian age collected from Skaar Ridge in the central Transantarctic Mountains, Antarctica. The small ovules are significant in possessing fleshy apical appendages and a funnel-shaped micropylar extension formed by the sarcotestal layer of the integument, by which they differ from all other Permian ovules described to date. The apical modifications may have functioned in pollination and/or seed dispersal. Similarity with the apical organization of earlier Paleozoic ovules is shown to be superficial, since the analogous structures are developmentally derived from different tissues. Although the ovules occur in rocks in which glossopterids are the only gymnosperms represented, there is insufficient evidence to assign them to a taxonomic group. These ovules are of particular importance because there are so few anatomically preserved gymnosperm reproductive structures known from the Permian and thus provide new data on the diversity of late Paleozoic gymnosperms.

  13. Generation and expulsion of oils from Permian coals of the Sydney Basin, Australia

    Ahmed, M.; Volk, H.; George, S.C.; Faiz, M.; Stalker, L. [CSIRO Petrology, Sydney, NSW (Australia)


    Organic geochemical and petrological assessment of coals/coaly shales and fine grained sediments, coupled with organic geochemical analyses of oil samples, all from Permo-Triassic sections of the Southern Sydney Basin (Australia), have enabled identification of the source for the widely distributed oil shows and oil seeps in this region. The Permian coals have higher hydrogen indices, higher liptinite contents, and much higher total organic matter extract yields than the fine grained sediments. A variety of source specific parameters obtained from n-alkanes, regular isoprenoids, terpanes, steranes and diasteranes indicate that the oil shows and seeps were generated and expelled predominantly from higher plant derived organic matter deposited in oxic environments. The source and maturity related biomarkers and aromatic hydrocarbon distributions of the oils are similar to those of the coals. The oil-coal relationship also is demonstrated by similarities in the carbon isotopic composition of the total oils, coal extracts, and their individual n-alkanes. Extracts from the Permo-Triassic fine grained sediments, on the other hand, have organic geochemical signatures indicative of mixed terrestrial and prokaryotic organic matter deposited in suboxic environments, which are significantly different from both the oils and coal extracts. The molecular signatures indicating the presence of prokaryotic organic matter in some of the coal extracts and oils may be due to thin sections of possibly calcareous lithologies interbedded within the coal measures. The genetic relationship between the oils and coals provides new evidence for the generation and expulsion of oils from the Permian coals and raises the possibility for commercial oil accumulations in the Permian and Early Triassic sandstones, potentially in the deeper offshore part of the Sydney Basin.

  14. Importance of carbon isotopic data of the Permian-Triassic boundary layers in the Verkhoyansk region for the global correlation of the basal Triassic layer

    Zakharov, Yu. D.; Biakov, A. S.; Richoz, S.; Horacek, M.


    This paper is dedicated to a global correlation of marine Permian-Triassic boundary layers on the basis of partially published and original data on the δ13Corg and δ13Ccarb values of the Suol section (Setorym River, South Verkhoyansk region). The section consists of six carbon isotopic intervals, which are easily distinguishable in the carbon isotopic curves for a series of Permian-Triassic reference sections of Eurasia and Northern America, including paleontologically described sections of Central Iran, Kashmir, and Southern China. This suggests that the Permian-Triassic boundary in the Suol section is close to the carbon isotopic minimum of interval IV. In light of new data, we suggest considering the upper part of the Late Permian Changhsingian Stage and the lower substage of the Early Triassic Induan Stage of Siberia in the volumes of the rank Otoceras concavum zone and the Tompophiceras pascoei and Wordieoceras decipiens zones, respectively. The O. concavum zone of the Verkhoyansk region probably corresponds to the Late Changhsingian Hypophiceras triviale zone of Greenland. The carbon isotopic intervals II, III, IV, and V in the Permian-Triassic boundary layers of the Verkhoyansk region traced in a series of the reference sections of Eurasia correspond, most likely, to intensification of volcanic activity at the end of the Late Changhsingian and to the first massive eruptions of Siberian traps at the end of the Changhsingian and the beginning of the Induan Stages. New data indicate the possible survival of ammonoids of the Otoceratoidea superfamily at the species level after mass extinction of organisms at the end of the Permian.




    Full Text Available Late Wordian (Guadalupian brachiopods from Member 2 of the Panjshah Formation in the Karakorum (N Pakistan are described. The brachiopod assemblage, dated by the associated fusulinids and conodonts, consists of 29 genera (3 of which are questionable and 1 unidentifiable of the orders Productida, Orthida, Rhynchonellida, Athyridida, Spiriferida and Terebratulida. Hunzininae, a new subfamily of the Spiriferellidae is proposed; it includes Darbandia n. gen., with type species D. vagabunda n. sp. and Elivina chapursani n. sp. A third new species is assigned to the genus Anchorhynchia of the family Wellerellidae: A. cimmerica n. sp.A quantitative biostratigraphic analysis demonstrates two major faunal changes in the Elivina chapursani- Chapursania tatianae Assemblage Zone of the upper part of Member 2, which are not strictly linked to lithological changes. This biozone is correlated with the brachiopod faunas of the Gnishik Formation of Armenia and those of the basal Takhtabulak Formation of SE Pamir.The faunal elements of the Elivina chapursani- Chapursania tatianae Assemblage Zone are an admixture of wide-ranging, Tethyan (particularly abundant, Gondwanan and endemic (Cimmerian genera, representing a transitional fauna and a biostratigraphic tool for intercontinental correlation, which are particularly problematic in this time interval. The Panjshah transitional fauna demonstrates the persistence of the Transhimalayan Province of the Cimmerian Region into the late Guadalupian, which originated at the end of the Cisuralian and occupied Armenia, Central Afghanistan, Karakorum and SE Pamir. It provides also some insights into the biodiversity pattern before the mass extinction at the end of the Guadalupian, and suggests that this event was as rapid as the end- Permian mass extinction, at least in Central Asia. 

  16. An accurately delineated Permian-Triassic Boundary in continental successions


    The Permian-Triassic Boundary Stratigraphic Set (PTBST), characteristic of the GSSP section of Meishan and widespread in marine Permian-Triassic Boundary (PTB) sequences of South China, is used to trace and recognize the PTB in a continental sequence at Chahe (Beds 66f―68c). Diversified Permian plant fossils extended to the PTBST, and a few relicts survived above that level. Sporomorphs are dominated by fern spores of Permian nature below the PTBST, above which they are replaced by gymnosperm pollen of Triassic aspect. In the nearby Zhejue Section, the continental PTBST is characterized by the fungal 'spike' recorded in many places throughout the world. The boundary claybeds (66f and 68a,c) of the PTBST are composed of mixed illite-montmorillonite layers analogous with those at Meishan. They contain volcanogenic minerals such as β quartz and zircon. U/Pb dating of the upper claybed gives ages of 247.5 and 252.6 Ma for Beds 68a and 68c respectively, averaging 250 Ma. In contrast to the situation in Xinjiang and South Africa, the sediment sequence of the Permian-Triassic transition in the Chahe section (Beds 56―80) become finer upward. Shallowing and coarsening upward is not, therefore, characteristic of the Permian-Triassic transition everywhere. The occurrence of relicts of the Gigantopteris Flora in the Kayitou Fm. indicates that, unlike most marine biota, relicts of this paleophytic flora survived into the earliest Triassic. It is concluded that Bed 67 at Chahe corresponds to Bed 27 at Meishan, and that the PTB should be put within the 60-cm-thick Bed 67b④, now put at its base tentatively. This is the most accurate correlation of the PTB in continental facies with that in the marine GSSP.

  17. Hypoxia, global warming, and terrestrial late Permian extinctions.

    Huey, Raymond B; Ward, Peter D


    A catastrophic extinction occurred at the end of the Permian Period. However, baseline extinction rates appear to have been elevated even before the final catastrophe, suggesting sustained environmental degradation. For terrestrial vertebrates during the Late Permian, the combination of a drop in atmospheric oxygen plus climate warming would have induced hypoxic stress and consequently compressed altitudinal ranges to near sea level. Our simulations suggest that the magnitude of altitudinal compression would have forced extinctions by reducing habitat diversity, fragmenting and isolating populations, and inducing a species-area effect. It also might have delayed ecosystem recovery after the mass extinction.

  18. Faunal migration into the Late Permian Zechstein Basin

    Sørensen, Anne Mehlin; Håkansson, Eckart; Stemmerik, Lars


    the northern margin of Pangea is used to test hypotheses concerning Late Palaeozoic evolution of the North Atlantic region. During the Permian, the Atlantic rift system formed a seaway between Norway and Greenland from the boreal Barents Shelf to the warm and arid Zechstein Basin. This seaway is considered...... to be the only marine connection to the Zechstein Basin and therefore the only possible migration route for bryozoans to enter the basin. The distribution of Permian bryozoans is largely in keeping with such a connection from the cool Barents Shelf past the East Greenland Basin to the warm Zechstein Basin...

  19. Stratigraphy and paleontology of Lower Permian rocks north of Cananea, northern Sonora, Mexico

    Blodgett, R.B.; Moore, T.E.; Gray, F.


    Lower Permian carbonate and overlying red bed clastic rocks are present in a 2 km2 stratigraphic window in the vicinity of Rancho La Cueva, Santa Cruz sheet (scale 1:50,000), northern Sonora, Mexico. This exposure lies unconformably beneath predominantly intermediate Upper Cretaceous volcanics yielding 40Ar/39Ar ages of 73.4?? 0.18 and 71.1 ?? 0.35 Ma. The lower part of the Permian succession consists of light- to medium-gray colored limestones of the Colina Limestone, with a minimum thickness of 235 m. Sedimentary features suggest shallow water, slightly restricted depositional environments. Although lacking observable fossils for the most part, two intervals of richly fossiliferous, silicified shell beds are present near the base and top of the Colina Limestone. The lower fauna consist mostly of gastropods and bivalves. The presence of a new microdomatid gastropod species. Glyptospira sonorensis n. sp., close to Glytospira arelela Plas, suggests a late Wolfcampian age for this horizon. The upper fauna are predominantly molluscan dominated (gastropods and bivalves), but some brachiopods (productids and the rhynchonellid genus Pontisia) are also present. Gastropod genera include Bellerophon, Warthia, Euomphalus (represented by the species, Euomphalus kaibabensis Chronic), Baylea, Worthenia, Naticopsis, Goniasma, Kinishbia, Cibecuia, and Glyptospira. The gastropods suggest a Leonardian (late Early Permian) age for this horizon, and many of the species have previously been recorded from the Supai Group and Kaibab Formation of northern and central Arizona. The Colina Limestone is conformably overlain by 11.2 m of light-gray lime mudstone and dolostone, assigned here to the Epitaph Dolomite, which in turn is succeeded by 58.8 m of red-colored sandstone and gray lime mudstone, assigned here to the Scherrer Formation. This Lower Permian succession is significant because it further strengthens the stratigraphic ties of southeastern Arizona rocks with those of northern

  20. The formation of early settled villages and the emergence of leadership: A test of three theoretical models in the Rio Ilave, Lake Titicaca Basin, southern Peru

    Craig, Nathan Mcdonald

    Despite obvious necessities for understanding later developments in the region, Preceramic cultures in the Titicaca Basin are poorly understood due to lacking empirical data. This dissertation focuses on examination of two processes occurring in the Rio Ilave drainage during the Late 5000--3100 B.C.E. (6000--4400 B.P.) and Terminal 3100--1500 B.C.E. Archaic (4400--3200 B.P.): the formation of settled villages and the emergence of leadership. These processes are tested against theoretical predictions derived from: population resource imbalance, human behavioral ecology, and agency theories. Resource imbalance theory predicts appearance of density dependent adaptive problems prior to major cultural transformations. Human behavioral ecology predicts increased dietary diversity and incorporation of greater processing costs occuring coincidentally with reduced residential mobility. Symbolic mate value advertising is a predicted leadership strategy. Agency theory predicts internal sources of social tension, potentially independent of external forces, as primary causes of cultural change. Feast hosting and the formation political economy are expected to be central to both social processes investigated from this perspective. Local paleoclimatic proxies suggest aridity from ca 4000--2000 B.C.E. Reanalysis of pedestrian survey data indicate population growth during the Late Archaic. Excavation and ground penetrating radar survey at Pirco and Jiskairumoko demonstrate reduced residential mobility and increased evidence for social differentiation appearing during the Late Archaic's (before 3000 B.C.E.) end. New cultural patterns include: pithouses, more groundstone, internal storage, costly non-local artifacts, domesticated plants and animals, costly grave goods, and use of ochre in symbolic contexts. No clear cut evidence for inter-community promotional feasting was identified but some finds suggest intra-community alliance feasts. By the end of the Terminal Archaic ca. 1500 B

  1. Early development of Auchenipterus osteomystax (Osteichthyes, Auchenipteridae from the Paraná river basin, Brazil Caracterização do desenvolvimento inicial de Auchenipterus osteomystax (Osteichthyes, Auchenipteridae da bacia do rio Paraná, Brasil

    Maristela Cavicchioli Makrakis


    Full Text Available The aim of the present study was to characterize the early development of Auchenipterus osteomystax (Ribeiro, 1918 (Osteichthyes, Auchenipteridae. Specimens were taken from plankton samples in the Upper Paraná river, Ivaí river, Upper Ivinhema river and Itaipu Reservoir. Seventy-four larvae and seven juveniles (with standard length ranging from 5.25 mm to 39.0 mm were analyzed. Specimens were identified and separated according to their development and morphometric and meristic analyses were performed. The early development of A. osteomystax follows the pattern of other Siluriformes. However, individuals presented more pigmentation on the head and on fore part of the body. Number of myomeres ranged from 44 to 50O objetivo deste trabalho é caracterizar o desenvolvimento inicial de Auchenipterus osteomystax (Ribeiro, 1918 (Osteichthyes, Auchenipteridae. O material analisado é proveniente de amostras coletadas no plâncton no alto rio Paraná, rio Ivaí, alto rio Ivinhema e reservatório de Itaipu. Foram analisadas 74 larvas e 7 juvenis, entre 5,25 mm e 39 mm de comprimento padrão. Os espécimes foram identificados e separados em diferentes estágios, e foram obtidas as variáveis morfométricas e merísticas. As larvas de A. osteomystax apresentam o desenvolvimento inicial semelhante ao de outros Siluriformes, no entanto são caracterizadas principalmente por apresentar um aglomerado de pigmentos na cabeça e na região anterior do corpo e número de miômeros variando de 44 a 50

  2. Discovery of Late Permian Jianshanzi volcanic rocks in Qinghe valley of northern Liaoning and its geologic significance

    Yuejun CHEN; Chunlin SUN; Yuewu SUN


    Jianshanzi volcanic rocks at Qinghe valley in the northern Liaoning were considered belonging to Early Proterozoic Gaojiayu Formation of Liaohe Group, or to Early Cambrian Beidagou Formation of Qinghezhen Group, or Middle-Late Proterozoic Shenjiapu Formation-Complex of Kaiyuan Group-complex. Dating the zircons from the dacite with schistosity is 2 506 Ma in method of U-Pb (SHRIMP). This evidence indicates the rocks may be referred to Late Permian in age. Discovery of the rocks is significant to re-recognize stratigraphic property of "Liaohe Group", regional geotectonic location and revolution of orogenic zone in Qinghe valley of the northern Liaoning.

  3. South Primorye, Far East Russia—A key region for global Permian correlation

    Kotlyar, Galina V.; Belyansky, Gennady C.; Burago, Valentina I.; Nikitina, Antonina P.; Zakharov, Yuri D.; Zhuravlev, Andrey V.


    Significant differences between faunal and floral associations existing in different paleogeographic realms in the Kungurian-Late Permian interval make it difficult to correlate the Permian deposits of the world. Resolving this problem is one of the main tasks of Permian stratigraphy. The global significance of Permian strata of the Primorye region of Far East Russia is enhanced by the specific Middle Permian mixed Tethyan, Boreal and Gondwanan-type brachiopod fauna, mixed Angara-Euromerican-Cathaysian flora, and their close spatial and stratigraphical association with fusulinids, bryozoans, ammonoids, conodonts. These facts permit tracing of global correlational levels of some Permian sequences within the different paleobiogeographical realms: for example, the Monodiexodina sutschanica- Metadoliolina dutkevichi fusulinid zone of the Wordian age and Parafusulina stricta fusulinid zone of the Capitanian age. The Late Permian fauna of the Primorye is mainly Tethyan in origin and provides correlation with similar aged sequences from South China.

  4. High precision time calibration of the Permian-Triassic boundary mass extinction event in a deep marine context

    Baresel, Björn; Bucher, Hugo; Brosse, Morgane; Bagherpour, Borhan; Schaltegger, Urs


    To construct a revised and high resolution calibrated time scale for the Permian-Triassic boundary (PTB) we use (1) high-precision U-Pb zircon age determinations of a unique succession of volcanic ash layers interbedded with deep water fossiliferous sediments in the Nanpanjiang Basin (South China) combined with (2) accurate quantitative biochronology based on ammonoids, conodonts, radiolarians, and foraminifera and (3) tracers of marine bioproductivity (carbon isotopes) across the PTB. The unprecedented precision of the single grain chemical abrasion isotope-dilution thermal ionization mass spectrometry (CA-ID-TIMS) dating technique at sub-per mil level (radio-isotopic calibration of the PTB at the Permian and Early Triassic. Here, we present new single grain U-Pb zircon data of volcanic ash layers from two deep marine sections (Dongpan and Penglaitan) revealing stratigraphic consistent dates over several volcanic ash layers bracketing the PTB. These analyses define weighted mean 206Pb/238U ages of 251.956±0.033 Ma (Dongpan) and 252.062±0.043 Ma (Penglaitan) for the last Permian ash bed. By calibration with detailed litho- and biostratigraphy new U-Pb ages of 251.953±0.038 Ma (Dongpan) and 251.907±0.033 Ma (Penglaitan) are established for the onset of the Triassic.

  5. Thecamoebians (Testate Amoebae Straddling the Permian-Triassic Boundary in the Guryul Ravine Section, India: Evolutionary and Palaeoecological Implications.

    Vartika Singh

    Full Text Available Exceptionally well-preserved organic remains of thecamoebians (testate amoebae were preserved in marine sediments that straddle the greatest extinction event in the Phanerozoic: the Permian-Triassic Boundary. Outcrops from the Late Permian Zewan Formation and the Early Triassic Khunamuh Formation are represented by a complete sedimentary sequence at the Guryul Ravine Section in Kashmir, India, which is an archetypal Permian-Triassic boundary sequence. Previous biostratigraphic analysis provides chronological control for the section, and a perspective of faunal turnover in the brachiopods, ammonoids, bivalves, conodonts, gastropods and foraminifera. Thecamoebians were concentrated from bulk sediments using palynological procedures, which isolated the organic constituents of preserved thecamoebian tests. The recovered individuals demonstrate exceptional similarity to the modern thecamoebian families Centropyxidae, Arcellidae, Hyalospheniidae and Trigonopyxidae, however, the vast majority belong to the Centropyxidae. This study further confirms the morphologic stability of the thecamoebian lineages through the Phanerozoic, and most importantly, their apparent little response to an infamous biological crisis in Earth's history.

  6. Late Permian to recent magmatic activity on the African-Arabian margin of Tethys

    Wilson, M. [Leeds University (United Kingdom). School of Earth Sciences; Guiraud, R. [Universite de Montpellier II (France). Laboratoire de Geophysique et Tectonique; Moreau, C. [Universite de la Rochelle (France). Departement des Sciences de la Terre; Bellion, Y.J.-C. [Laboratoire de Geologie, Avignon (France). Faculte des Science


    Magmatic activity on the African-Arabian margin of Tethys has fluctuated significantly during the past 250 Ma in response to major phases of mantle plume activity and to extensional stresses within the African plate related to periods of continental break-up. A series of maps have been compiled, based upon stratigraphic constraints and K-Ar and {sup 40}Ar-{sup 39}Ar age determinations, to illustrate the changing patterns of igneous activity since the Late Permian. These are divided into five stages: I, Late Permian (256 Ma)-Latest Triassic (208 MA); II, Early Jurassic (208 Ma, Liassic)-Mid Jurassic (157 Ma, Dogger); III, Late Jurassic (157 Ma)-Earliest Aptian (120 Ma); IV, Mid Aptian (119 Ma)-Mid Eocene (42 Ma); V, Late Eocene (42 Ma)-Recent. Magmatism appears to be extremely long-lived in a number of areas (e.g. Sudan, Air), with volcanic centres located along major basement fault zones, such as the Guinean-Nubian lineaments. (author)

  7. Dynamic anoxic ferruginous conditions during the end-Permian mass extinction and recovery

    Clarkson, M. O.; Wood, R. A.; Poulton, S. W.; Richoz, S.; Newton, R. J.; Kasemann, S. A.; Bowyer, F.; Krystyn, L.


    The end-Permian mass extinction, ~252 million years ago, is notable for a complex recovery period of ~5 Myr. Widespread euxinic (anoxic and sulfidic) oceanic conditions have been proposed as both extinction mechanism and explanation for the protracted recovery period, yet the vertical distribution of anoxia in the water column and its temporal dynamics through this time period are poorly constrained. Here we utilize Fe-S-C systematics integrated with palaeontological observations to reconstruct a complete ocean redox history for the Late Permian to Early Triassic, using multiple sections across a shelf-to-basin transect on the Arabian Margin (Neo-Tethyan Ocean). In contrast to elsewhere, we show that anoxic non-sulfidic (ferruginous), rather than euxinic, conditions were prevalent in the Neo-Tethys. The Arabian Margin record demonstrates the repeated expansion of ferruginous conditions with the distal slope being the focus of anoxia at these times, as well as short-lived episodes of oxia that supported diverse biota.

  8. Functional diversity of marine ecosystems after the Late Permian mass extinction event

    Foster, William J.; Twitchett, Richard J.


    The Late Permian mass extinction event about 252 million years ago was the most severe biotic crisis of the past 500 million years and occurred during an episode of global warming. The loss of around two-thirds of marine genera is thought to have had substantial ecological effects, but the overall impacts on the functioning of marine ecosystems and the pattern of marine recovery are uncertain. Here we analyse the fossil occurrences of all known benthic marine invertebrate genera from the Permian and Triassic periods, and assign each to a functional group based on their inferred lifestyle. We show that despite the selective extinction of 62-74% of these genera, all but one functional group persisted through the crisis, indicating that there was no significant loss of functional diversity at the global scale. In addition, only one new mode of life originated in the extinction aftermath. We suggest that Early Triassic marine ecosystems were not as ecologically depauperate as widely assumed. Functional diversity was, however, reduced in particular regions and habitats, such as tropical reefs; at these smaller scales, recovery varied spatially and temporally, probably driven by migration of surviving groups. We find that marine ecosystems did not return to their pre-extinction state, and by the Middle Triassic greater functional evenness is recorded, resulting from the radiation of previously subordinate groups such as motile, epifaunal grazers.

  9. Terrestrial Permian - Triassic boundary sections in South China

    Bercovici, Antoine; Vajda, Vivi


    The Permian-Triassic boundary interval in China comprises a significant record of faunal and floral changes during this important extinction event. Here we discuss the details of palynomorph preservation at the classical Western Guizhou and Eastern Yunnan sections in an effort to expand the stratigraphy and paleontology from these earlier studies.

  10. Oceanic Anoxia and the End Permian Mass Extinction

    Wignall; Twitchett


    Data on rocks from Spitsbergen and the equatorial sections of Italy and Slovenia indicate that the world's oceans became anoxic at both low and high paleolatitudes in the Late Permian. Such conditions may have been responsible for the mass extinction at this time. This event affected a wide range of shelf depths and extended into shallow water well above the storm wave base.

  11. Late Carboniferous to Late Permian carbon isotope stratigraphy

    Buggisch, Werner; Krainer, Karl; Schaffhauser, Maria


    An integrated study of the litho-, bio-, and isotope stratigraphy of carbonates in the Southern Alps was undertaken in order to better constrain δ13C variations during the Late Carboniferous to Late Permian. The presented high resolution isotope curves are based on 1299 δ13Ccarb and 396 δ13Corg a...

  12. Terrestrial acidification during the end-Permian biosphere crisis?

    Sephton, Mark A.; Jiao, Dan; Engel, Michael H.; Looy, Cindy V.; Visscher, Henk


    Excessive acid rainfall associated with emplacement of the Siberian Traps magmatic province is increasingly accepted as a major contributing factor to the end-Permian biosphere crisis. However, direct proxy evidence of terrestrial acidification is so far not available. In this paper, we seek to dete

  13. Study on the Late Permian Claraia in South China

    YANG; Fengqing


    [1]Nakazawa, K., On Claraia of Kashmir and Iran, J. Paleont. Soc. Idia., 1977, 20: 194-204.[2]Kulikov, M. V., Tkachuk, G. A., On finding of Claraia (Bivalvia) in Upper Permian deposits of Northern Caucasus,Doklady Akad. Nauk. USSR, 1979, 245(4): 905-908.[3]Yin. H. F.. Uppermost Permian (Changxingian) Pectinacea from South China, Riv. Ital. Paleont., 1982, 88(3): 337-386.[4]Yin. H. F.. Bivalves near the Permian-Triassic boundary in South China, J. Paleo.,1985, 59(3): 572-600.[5]Yang. Z. Y.. Yin H. F., Wu, S. B. et al., Permian-Triassic Boundary Stratigraphy and Fauna of South China (in Chinese).Geological Memoirs, Ser. 2, No. 6, Beijing: Geological Publishing House, 1987, 56-57, 236-237.[6]Guo. F. X.. Bivalves Fossil of Yunnan (in Chinese), Kunming: Yunnan Technology Press, 1985.[7]Fang. Z. J., Discussion on Late Permian “Claraia”, Act. Palaeo. Sini. (in Chinese),1993, 2(6):653-661.[8]Zhang. Z. M.. Discussion on the ligament structure of Claraia and its evolution, Act. Palaeo. Sini. (in Chinese), 1980,19(6): 433-443.[9]Nanjing Institute of Geology and Paleontology, Academia Sinica, The Bivalves of China (in Chinese), Beijing: Science Press, 1976.[10]Zhou. Z. R.. Two ecological types of Permian Ammonoids, Science in China (in Chinese), Ser. B, 1985, (7): 648-657.[11] Stanley. S. M., Relation of shell form to life habitats of the Bivalvia (Mollusca), Geol. Soc. Amer., 1970, 125: 1-496.[12]Stanley, S. M.. tunctional morphology and evolution of byssally attached bivalve mollusks, J. Paleont, 1972, 46(2): 165-207.[13]Yin, H. F.. Ding, M. H., Zhang, K. X. et al., Late Permian-Middle Triassic Ecostratigraphy of the Yangtze Platform and Its Margins, Beijing: Science Press, 1995, 69-76.[14]Leonardi, E. Trias inferiore delle Venezie, Mem. Inst. Geol. Univ. Padova., 1935, II, 1-136.[15]Ichikawa. K. Zur Taxonomie und Phylogenie der Triadischen Pteriidae (Lamellibranch), Palaeotog., 1958, III, A, 132-212.

  14. A influência da traqueostomia precoce no desmame ventilatório de pacientes com traumatismo craniencefálico grave The influence of early tracheostomy in the weaning of patients with severe traumatic brain injury

    Renata Lenize Pasini


    Full Text Available JUSTIFICATIVA E OBJETIVOS: Atualmente, ainda é questionável se a traqueostomia precoce (TP pode influenciar no desmame ventilatório ou no tempo de internação hospitalar de pacientes com traumatismo cranioencefálico (TCE. O objetivo primário deste estudo foi verificar se a TP influencia o tempo de ventilação mecânica (VM em pacientes com TCE grave. MÉTODO: Estudo prospectivo, observacional, incluindo 33 pacientes com pontuação admissional na escala de coma de Glasgow (ECG 12 dias e o desmame ventilatório. RESULTADOS: O tempo total de VM foi menor no grupo TP (n = 10; p BACKGROUND AND OBJECTIVES: Actually, It’s doubtful if early tracheostomy (ET can influence mechanical ventilation (MV weaning time or the hospital length of stay in patients with traumatic brain injury (TBI. The main objective of this trial was to verify the influence of ET on weaning time of patients with severe TBI. METHODS: Prospective, observational study, including 33 patients with severe TBI (GCS 12 days and the weaning from MV. RESULTS: Total ventilation mechanical time has been reduced in the early tracheostomy group (n = 10; p < 0.0001. A lower GCS punctuation (mean 5.3 ± 2.5 in ET group has been negatively correlated with hospital length of stay (LOS (p = 0.02. CONCLUSIONS: Early tracheostomy can decrease mechanical ventilation time, but does not influence hospital LOS in patients with severe traumatic brain injury.

  15. Amphibian and Paleoisciforms from the Lower Part of the Taquaral Member of the Permian Irati Formation, São Paulo State, Brazil

    Artur Chahud


    Full Text Available In the east-central region of the State of São Paulo, Brazil, good exposures of Permo-Carboniferous sequences of theParaná intracratonic basin are found. These sequences start with the Tubarão Supergroup deposits, the PermocarboniferousItararé Group, and the earliest Permian Guatá Group, of which the Tatuí Formation is the only unit occurring in the Stateof São Paulo. The Permian Passa Dois Group overlies the Tubarão Supergroup, with the Early Permian Irati Formation andthe Middle Permian Corumbataí Formation. Two members are recognized in the Irati Formation, Taquaral and Assistência.Most beds of the Taquaral Member are gray laminated siltic-argillaceous, but sandstone beds are present in lower outcropsof this member. One of these sandstones, 9.5 cm thick unconformably overlying sedimentos of the Tatuí Formation, exhibitsdiversifi ed vertebrate remains. The present paper deals with a Palaeonisciformes mandible fragment, one amphibianTemnospondyli tooth, and one Temnospondyli mandible fragment. These fossils are associated to Chondrichthyes andOsteichthyes. It is an open question whether the Parnaíba Basin Palaeonisciformes Brasilichthys macrognathus and theTemnospondyli Prionosuchus plummery belong to the same taxa of the Paraná Basin fossils. Only better preserved fossilsmight solve this question.

  16. Paleomagnetism of Permian rocks of the Subpolar Urals, Kozhim River: To the history of evolution of the thrust structures in the Subpolar Urals

    Iosifidi, A. G.; Popov, V. V.


    The collections of Permian rocks from sections of the Kozhim River (Asselian, Kungurian, and Ufimian stages) and the Kama River (Ufimian and Kazanian stages) are studied. The paleomagnetic directions determined on the studied structures closely agree with the existing data for the Subpolar Urals and Russian Platform (RP). In the Middle Permian red clays of the Kama River region, the paleomagnetic pole N/n = 28/51, Φ = 47° N, Λ = 168° E, dp = 3°, and dm = 5° is obtained. The analysis of the existing paleomagnetic determinations for the Early and Middle Permian of the Russian and Siberian platforms and Kazakhstan blocks (KBs) is carried out. For the Subpolar Ural sections, the estimates are obtained for the local rotations during the collision of the Uralian structures with the Russian and Siberian platforms and KBs. The amplitudes of the horizontal displacements of the studied structures are, on average, 170 ± 15 km per Middle Permian. The scenario describing the evolution of the horizontal rotations of the structures of Subpolar Urals is suggested.

  17. A multistratigraphic approach to pinpoint the Permian-Triassic boundary in continental deposits: The Zechstein-Lower Buntsandstein transition in Germany

    Scholze, Frank; Wang, Xu; Kirscher, Uwe; Kraft, Johannes; Schneider, Jörg W.; Götz, Annette E.; Joachimski, Michael M.; Bachtadse, Valerian


    The Central European Basin is very suitable for high-resolution multistratigraphy of Late Permian to Early Triassic continental deposits. Here the well exposed continuous transition of the lithostratigraphic Zechstein and Buntsandstein Groups of Central Germany was studied for isotope-chemostratigraphy (δ13Corg, δ13Ccarb, δ18Ocarb), major and trace element geochemistry, magnetostratigraphy, palynology, and conchostracan biostratigraphy. The analysed material was obtained from both classical key sections (abandoned Nelben clay pit, Caaschwitz quarries, Thale railway cut, abandoned Heinebach clay pit) and a recent drill core section (Caaschwitz 6/2012) spanning the Permian-Triassic boundary. The Zechstein-Buntsandstein transition of Central Germany consists of a complex sedimentary facies comprising sabkha, playa lake, aeolian, and fluvial deposits of predominantly red-coloured siliciclastics and intercalations of lacustrine oolitic limestones. The new data on δ13Corg range from - 28.7 to - 21.7 ‰ showing multiple excursions. Most prominent negative shifts correlate with intercalations of oolites and grey-coloured clayey siltstones, while higher δ13Corg values correspond to an onset of palaeosol overprint. The δ13Ccarb values range from - 9.7 to - 1.3 ‰ with largest variations recorded in dolomitic nodules from the Zechstein Group. In contrast to sedimentary facies shifts across the Zechstein-Buntsandstein boundary, major element values used as a proxy (CIA, CIA*, CIA-K) for weathering conditions indicate climatic stability. Trace element data used for a geochemical characterization of the Late Permian to Early Triassic transition in Central Germany indicate a decrease in Rb contents at the Zechstein-Buntsandstein boundary. New palynological data obtained from the Caaschwitz quarry section reveal occurrences of Late Permian palynomorphs in the Lower Fulda Formation, while Early Triassic elements were recorded in the upper part of the Upper Fulda Formation

  18. Permian Triassic palynofloral transition in Chintalapudi area, Godavari Graben, Andhra Pradesh, India

    Neerja Jha; M Basava Chary; Neha Aggarwal


    The entire 606 m-thick sedimentary sequence in borecore MCP-7 from Chintalapudi area, Chintalapudi sub-basin has been lithologically designated as Kamthi Formation. However, the palynological investigation revealed five distinct palynoassemblages, which essentially fall under two groups, one group (Palynoassemblage-I, II and III) having dominance of striate disaccates along with presence of some stratigraphically significant taxa, belongs to Late Permian (Raniganj) palynoflora, while the other group (Palynoassemblages IV and V) shows sharp decline in percentage of characteristic taxa of first group, i.e., striate disaccates, and consequent rise or dominance of taeniate and cingulate cavate spores, belongs to Early Triassic (Panchet) palynoflora. Palynoassemblage-I, II and III (Group I) are characterized by dominance of striate disaccates chiefly, Striatopodocarpites spp. and Faunipollenites spp. Along with presence of rare but stratigraphically significant taxa, viz., Gondisporites raniganjensis, Falcisporites nuthaliensis, Klausipollenites schaubergeri, Chordasporites sp., Striomonosaccites, ovatus, Crescentipollenites multistriatus, Verticipollenites debiles, Strotersporites crassiletus, Guttulapollenites hannonicus, G. gondwanensis, Hamiapollenites insolitus, Corisaccites alutus, Lunatisporites ovatus, Weylandites spp. and Vitreisporites pallidus. Palynoassemblage-I is distinguished by significant presence of Densipollenites spp. while Palynoassemblage-II shows significant presence of Crescentipollenites spp. and Palynoassemblage-III differs from the above two assemblages in having significant presence of Guttulapollenites hannonicus. Palynoassemblage-IV (Group II) is characterized by high percentage of taeniate disaccates chiefly Lunatisporites spp., while Palynoassemblage-V (Group II) is characterized by cingulate-cavate trilete spores chiefly, Lundbladispora spp. and Densoisporites spp. Striate disaccates show a sharp decline in these two assemblages. In

  19. Nystroemiaceae, a new family of Permian gymnosperms from China with an unusual combination of features.

    Wang, Jun; Pfefferkorn, Hermann W


    Nystroemiaceae is proposed as a new family of gymnosperms from the Permian of Cathaysia that adds to the diversity of gymnosperms known from this critical time in seed plant evolution. This family is characterized by bifurcating and highly branched pinnate ovuliferous organs bearing bicornute ovules (seeds) and entire leaves with anastomozing veins that are born on complex and modern-looking branching systems with clear axillary branching. The reconstruction is based on numerous large specimens from two localities in North China, in which the different plant parts are attached to each other. The ovulate structures show some apparently plesiomorphic (primitive) character states more typical of early seed plants, whereas the leaves and branches show the clearly apomorphic (derived) character states of broad-leaved gymnosperms.

  20. Nystroemiaceae, a new family of Permian gymnosperms from China with an unusual combination of features

    Wang, Jun; Pfefferkorn, Hermann W.


    Nystroemiaceae is proposed as a new family of gymnosperms from the Permian of Cathaysia that adds to the diversity of gymnosperms known from this critical time in seed plant evolution. This family is characterized by bifurcating and highly branched pinnate ovuliferous organs bearing bicornute ovules (seeds) and entire leaves with anastomozing veins that are born on complex and modern-looking branching systems with clear axillary branching. The reconstruction is based on numerous large specimens from two localities in North China, in which the different plant parts are attached to each other. The ovulate structures show some apparently plesiomorphic (primitive) character states more typical of early seed plants, whereas the leaves and branches show the clearly apomorphic (derived) character states of broad-leaved gymnosperms. PMID:19656793

  1. Illite/smectite clays preserving porosity at depth in Lower Permian Reservoirs, Northern Perth Basin

    D.D. Ferdinando; J.C. Baker; A. Gongora; B.A. Pidgeon


    The appraisal well Hovea-2, drilled in July 2002, was the first well in the onshore Perth Basin to discover commercial volumes of gas in sandstone reservoirs of the Artinskian (Early Permian) High Cliff Sandstone sealed beneath basal shales and siltstones of the Irwin River Coal Measures. A drill stem test in this formation, in the interval 2,370-419 mMDRT (measured depth below rotary table), flowed gas to surface at 16.5 MMcfd. Thereafter, the High Cliff Sandstone became an important play for exploration in the basin; however, no additional discoveries have been made in this reservoir, even though it has been tested in another seven wells (Jingemia-1, Eremia-1, Kunzia-1, Corybas-1, Yardarino-6, Hakia-2 and Bunjong-1).

  2. Groundwater ferricretes from the Silurian of Ireland and Permian of the Spanish Pyrenees

    Wright, V. P.; Sloan, R. J.; Valero Garcés, B.; Garvie, L. A. J.


    Haematite nodule horizons occur in alluvial-lacustrine deposits from the Mill Cove Formation (Silurian) of Dingle, Ireland and the Unidad Roja Superior (Permian) of the Spanish Pyrenees. These horizons, generally less than 1 m thick, are not associated with palaeosol profiles but are hosted by laminated or structureless, bioturbated mudstones to sandstones. The sequences contain weakly developed calcrete profiles. The haematite nodules are interpreted as having formed around fluctuating water-tables and are analogous to present-day groundwater ferricretes (laterites or plinthites). Fe 2+, mobilised during periods of reducing groundwaters, was segregated into ferric oxide nodules during falls in the water-table or during influxes of oxygenated meteoric waters. The prevailing climate during deposition of both sequences was probably semi-arid and the presence of ferricrete-like horizons is not evidence of humid climates. The horizons represent early "diagenetic" effects, not related to prevailing soil moisture regime.

  3. Shallow lacustrine system of the Permian Pedra de Fogo Formation, Western Gondwana, Parnaíba Basin, Brazil

    Araújo, Raphael Neto; Nogueira, Afonso César Rodrigues; Bandeira, José; Angélica, Rômulo Simões


    contact with the Motuca Formation, are considered here as excellent biostratigraphic markers. Fish remains, ostracods, bryozoans and scolecodonts represent other fossils that are present in the succession. Mudflat deposits developed in an arid and hot climate probably in the Early Permian. Semi-arid conditions prevailed in the Middle Permian allowing the proliferation of fauna and flora in adjacent humid regions and onto the lake margin. The climate variation was responsible for the contraction and expansion phases of the lake, fed by sporadic sheet floods carrying plant remains. The reestablishment of the arid climate, at the end of Permian, marks the final sedimentation of the Pedra de Fogo Formation, linked to the consolidation of the Pangaea Supercontinent. This last event was concomitant with the deposition of the Motuca Formation red beds and the development of extensive ergs related to the Triassic Sambaíba Formation in Western Gondwana.

  4. Permian plants from the Chutani Formation (Titicaca Group, Northern Altiplano of Bolivia: II. The morphogenus Glossopteris

    Roberto Iannuzzi


    Full Text Available Fossil plants belonging to the morphogenera Glossopteris, Pecopteris and Asterotheca were collected from the upper part of the Chutani Formation (Titicaca Group, near the town of San Pablo de Tiquina, on the southeastern shore of Lake Titicaca (northern Altiplano, Bolivia. This paper presents the first description of specimens of the morphogenus Glossopteris from Bolivia. The Bolivian specimens of Glossopteris consist of poorly-preserved impressions, although they present the diagnostic features of this morphogenus. They are fragments of leaves with secondary veins of taeniopterid-type, typical of glossopterids from Late Permian deposits of Gondwana. The only species of Pecopteris confirmed in the first part of this study, i.e. P. dolianitii Rösler and Rohn (see Vieira et al. 2004, was previously reported from the Late Permian beds of the Rio do Rasto and Estrada Nova formations in the Paraná Basin (southern Brazil. Therefore, a Late Permian age is proposed for the fossil plant-bearing beds of the Chutani Formation based on the analyzed assemblage. The phytogeographic implications of this new find are briefly analyzed.Plantas fósseis, pertencentes aos morfo-gêneros Glossopteris, Pecopteris e Asterotheca, foram coletadas na porção superior da seção aflorante da Formação Chutani, próxima ao povoado de San Pablo de Tiquina, sudeste do lago Titicaca (Altiplano norte, Bolívia. Este trabalho apresenta a primeira descrição de espécimes do morfo-gênero Glossopteris provenientes da Bolívia. Os espécimes estudados de Glossopteris consistem em impressões foliares pobremente preservadas nas quais feições diagnósticas estão presentes. Os fragmentos foliares apresentam venação secundária do tipo teniopteróide, uma característica típica de glossopterídeas encontradas em depósitos do Permiano Superior do Gondwana. Por sua vez, a única espécie de Pecopteris confirmada para estes níveis da Formação Chutani, i.e. P. dolianitii

  5. Insect mimicry of plants dates back to the Permian

    Garrouste, Romain; Hugel, Sylvain; Jacquelin, Lauriane; Rostan, Pierre; Steyer, J.-Sébastien; Desutter-Grandcolas, Laure; Nel, André


    In response to predation pressure, some insects have developed spectacular plant mimicry strategies (homomorphy), involving important changes in their morphology. The fossil record of plant mimicry provides clues to the importance of predation pressure in the deep past. Surprisingly, to date, the oldest confirmed records of insect leaf mimicry are Mesozoic. Here we document a crucial step in the story of adaptive responses to predation by describing a leaf-mimicking katydid from the Middle Permian. Our morphometric analysis demonstrates that leaf-mimicking wings of katydids can be morphologically characterized in a non-arbitrary manner and shows that the new genus and species Permotettigonia gallica developed a mimicking pattern of forewings very similar to those of the modern leaf-like katydids. Our finding suggests that predation pressure was already high enough during the Permian to favour investment in leaf mimicry.

  6. Abrupt and gradual extinction among Late Permian land vertebrates in the Karoo basin, South Africa.

    Ward, Peter D; Botha, Jennifer; Buick, Roger; De Kock, Michiel O; Erwin, Douglas H; Garrison, Geoffrey H; Kirschvink, Joseph L; Smith, Roger


    The Karoo basin of South Africa exposes a succession of Upper Permian to Lower Triassic terrestrial strata containing abundant terrestrial vertebrate fossils. Paleomagnetic/magnetostratigraphic and carbon-isotope data allow sections to be correlated across the basin. With this stratigraphy, the vertebrate fossil data show a gradual extinction in the Upper Permian punctuated by an enhanced extinction pulse at the Permian-Triassic boundary interval, particularly among the dicynodont therapsids, coinciding with negative carbon-isotope anomalies.

  7. 准噶尔盆地西北缘下二叠统火山岩岩性、岩相及其与储层的关系%Early Permian Volcanic Lithology, Lithofacies and Their Relations to Reservoir in Northwestern Margin of the Junggar Basin

    鲜本忠; 牛花朋; 朱筱敏; 董国栋; 朱世发; 安思奇


    Early Permian volcanic rocks in northwestern margin of the Junggar Basin have different rock types and lithofacies. In this paper, data from core, thin sections and element chemistry were used to study volcanic lithology and their relations to reservoir quality. The Jiamuhe Formation is characterized by lava and volcanoclastic rocks, which are mainly basaltic andesites, andesites and quartz andesites. The lithology of volcanic rocks in the Fengcheng Formation are different in different areas. Lava including basaltic andesitoid and tephrite was the main type in Kebai area, while welded pyroclastic rock and volcaniclastic lava of rhyolite, pollenite and andesitoid are the main types in Wuxia area. The high-quality reservoir developed in eruptive breccias and lava such as andesite in the Jiamuhe Formation and in the Fengcheng Formation of the Kebai area, and in welded pyroclastic rock and breccias lava with stone bubbles in the Fengcheng Formation of the Wuxia area. Besides, same volcanic rock would have different reservoir quality because of different volcanic environments. In the studied area, although diagenesis was diverse in different parts, high-quality volcanic rock reservoir developed mainly in the upper effusive sub-facies, air fall, and pyroclastic subfacies in explosive facies. Effusive and explosive facies were common in the Jiamuhe Formation, while effusive facies and pyroclastic flow subfacies in explosive facies were the main facies in Fengcheng Formation of Kebai area and of Wuxia area, respectively. Therefore, distinct explorative targets should be chosen in different layers in different areas. Lithofacies was another important controlling factor for volcanic reservoir besides fracturing in fault zones and leaching under the regional unconformable surface.%  与石炭系相比,准噶尔盆地西北缘下二叠统火山岩岩性、岩相类型丰富,研究程度低。本文利用岩心、薄片及元素地球化学资料,开展了火山岩

  8. The Permian-Triassic boundary & mass extinction in China

    Ian Metcalfe; Lance Black; Qu Xun; Mao Xiaodong; Robert S. Nicoll; Roland Mundil; Clinton Foster; Jonathan Glen; John Lyons; Wang Xiaofeng; Wang Cheng-yuan; Paul R. Renne


    @@ The first appearance of Hindeodus parvus (Kozur & Pjatakova) at the Permian-Triassic (P-T) GSSP level (base of Bed 27c) at Meishan is here confirmed. Hindeodus changxingensis Wang occurs from Beds 26 to 29 at Meishan and appears to be restricted to the narrow boundary interval immediately above the main mass extinction level in Bed 25. It is suggested that this species is therefore a valuable P-T boundary interval index taxon.

  9. New SHRIMP U-Pb and 40Ar/39Ar constraints on the crustal stabilization of southern South America, from the margin of the Rio de Plata (Sierra de Ventana) craton to northern Patagonia

    Tohver, E.; Cawood, P. A.; Rossello, E.; Lopez de Luchi, M. G.; Rapalini, A.; Jourdan, F.


    Two models exist to explain the late Paleozoic tectonic history for southern South America: an accretionary model of crustal growth through magmatism and a collisional model involving pre-existing continental elements, namely, the Rio de Plata craton and the possibly allochthonous terrane(s) of Patagonia, the Northern Patagonia Massif and the Deseado Massif. We report new U-Pb and 40Ar/39Ar results from rocks within a posited collision zone between the SW edge of the Rio de Plata craton and the northern margin of the Northern Patagonia Massif. Igneous basement samples from the Sierra de Ventana region, Buenos Aires province, were dated by ion microprobe (SHRIMP) analysis of zircon. A previously unrecognized occurrence of Paleoproterozoic basement indicates that the Rio de Plata craton extends ca.250 km farther west than considered. The majority of the basement rocks are shallow mid-Cambrian granitoids and rhyolites, including the rocks of the Cerro Colorado granite, which is intrusive into the sediments of the Curamalal Gp, signifying that these mature quartzites and conglomerates are older than early Cambrian in age, possibly correlated with the low-grade sedimentary rocks of the Tandilia Range that includes the La Tinta Fm. The 40Ar/39Ar ages from biotite, muscovite, and sericite from three different sheared basement localities demonstrates deformation in the latest Permian (265-260 Ma), ca. 20 Ma after the foreland deposition of the synorogenic Tunas Fm. in the upper Pilahuinco Gp, constrained by 282.4 ± 2.8 Ma zircon ages in volcanic ashbeds. Farther south, along the northern margin of the Northern Patagonian Massif, late Ordovician 40Ar/39Ar cooling ages of granites intrusive into the Cambro-Ordovician Nahuel Niyeu Fm. are consistent with the presence of Ordovician magmatism along the W edge of the Rio de Plata craton. These ages alternate with late Permian 40Ar/39Ar cooling ages from undeformed granites and pegmatites, as well as early Jurassic cross

  10. Hooked: Habits of the Chinese Permian gigantopterid Gigantonoclea

    Seyfullah, Leyla J.; Glasspool, Ian J.; Hilton, Jason


    Based upon anatomical evidence, Permian aged gigantopterid fossils are in general reconstructed as climbing or scrambling plants. Gigantonoclea, a genus of adpressed gigantopterid foliage from the Permian of northern China, has been reported to co-occur with hook-like organs that were interpreted as indicating a scrambling/climbing habit. We reinvestigated these hook-like structures and re-evaluated the nature of the co-occurrences in context with the flora preserved in each plant-bearing fossil 'bed' in the North China sedimentary succession. New findings show that the species Gigantonoclea hallei probably climbed using specially adapted clusters of compound grappling hook-like shoots borne on the stems. This structural arrangement comprising shoots of hooks is new to the scrambling/climbing concept in gigantopterids. However, a key figured specimen previously reported as showing intermediate hook-tipped leaf morphology on a sole pinnule tip is discounted as such and is reinterpreted as a 'normal' pinnule partially hidden under sediment that results in an unusual appearance to this pinnule tip. Adaptations for climbing or scrambling based upon 'hooked leaves' observed in Gigantonoclea lagrelii are no longer supported and are reinterpreted as incompletely expanded leaves where the vernation process was interrupted. These data weaken prior interpretations of G. lagrelii as a climber/scrambler and raise doubts about the ubiquity of hooks amongst the gigantopterids as structures enabling them to climb or scramble their way through the Permian world.

  11. Thecamoebians from Late Permian Gondwana sediments of peninsular India.

    Farooqui, Anjum; Aggarwal, Neha; Jha, Neerja


    The evolutionary history of thecamoebians (testate amoebae) extends back to the Neoproterozoic Era. However, until now, these have had a restricted, discontinuous and modest record across the world. The studied sediment of Raniganj Formation (Godavari Graben), Andhra Pradesh, India has been assigned as Late Permian on the basis of co-occurring age-diagnostic Late Permian palynomorphs. About sixteen thecamoebian species and one taxon incertae sedis have been recorded here in the palynological slides on the basis of shell morphology and morphometry. Out of these, five belong to the family Arcellidae, seven to Centropyxidae, two to Trigonopyxidae, one to Difflugiidae, one to Plagiopyxidae, and one is regarded incertae sedis. The morphometric characteristics of fossil forms resemble their corresponding extant species studied from ecologically diverse fresh water wetlands in India. In general, the ratio of shell diameter and aperture diameter of Late Permian fossil and extant specimens show significant correlation in all the studied species. Except that, the ratio of shell length and breadth is the distinguishing feature between Centropyxis aerophila and C. aerophila 'sylvatica', rather than the ratio of shell length and longest diameter of the shell aperture in both fossil and extant forms. The study elucidates the minimal morphological evolution in thecamoebians and their survival during mass extinction periods and stressful environmental conditions over the geological timescale.

  12. Hot acidic Late Permian seas stifle life in record time

    Georgiev, Svetoslav; Stein, Holly J.; Hannah, Judith L.; Bingen, Bernard; Weiss, Hermann M.; Piasecki, Stefan


    The end of Permian time (252-251 Ma) hosts the largest mass extinction in Earth history, yet events heralding this global catastrophe remain intensely disputed. We present a chemostratigraphic marker, the 187Re/ 188Os ratio, which soars to unprecedented levels approaching the Permo-Triassic boundary. These ratios are tied to profound trace element changes and a precise Re-Os time record at 252 Ma preserved in black shales from East Greenland and the mid-Norwegian shelf. Within a 36-meter shale section, an 80-fold increase in Re concentrations (two-fold for Os) signals seawater conditions that became increasingly inhospitable to life. Unwavering initial 187Os/ 188Os ratios of 0.6 preclude mafic volcanism and meteorite impact as the direct cause of Late Permian anoxia. We argue that extraordinarily high 187Re/ 188Os ratios are the hallmark of simultaneously rising ocean temperature and acidity, leading to loss of oxygen and the stifling of life in latest Permian time.

  13. Hypercalcified demosponges and the end-Permian extinction

    Finks, Robert M.


    The hypothesis is presented here that the hypercalcified demosponges, characterized by a solid, largely external skeleton of spherulitic aragonite, or of high-Mg calcite, had symbiotic cyanobacteria, from the outset of their existence, that enabled them to precipitate their peculiar skeleton, and that both the skeleton and the cyanobacteria enabled them to survive the late and end-Permian extinctions, almost alone among sponge groups. The cyanobacteria would have provided them with photosynthetic oxygen to survive hypoxia (one of the proposed causes of the extinctions), and perhaps also were cropped for food, if the external bacteria, normally fed upon by sponges, were reduced by the hypoxia. The sponges' external skeleton would also have protected them in their shallow-water reefy environment from excess ultraviolet light that may have resulted from hypoxic loss of the ozone layer of the atmosphere. This is supported by the fact that the genus Guadalupia which has a lightly-calcified flat upper surface, died out by the end of the Permian, while 31 other genera of hypercalcified Demospongea as well as 3 genera of hypercalcified Calcarea were the only genera of sponges of any kind to continue from the Permian into the Triassic.

  14. Strangelove Ocean and Deposition of Unusual Shallow-Water Carbonates After the End-Permian Mass Extinction

    Rampino, Michael R.; Caldeira, Ken


    The severe mass extinction of marine and terrestrial organisms at the end of the Permian Period (approx. 251 Ma) was accompanied by a rapid negative excursion of approx. 3 to 4 per mil in the carbon-isotope ratio of the global surface oceans and atmosphere that persisted for some 500,000 into the Early Triassic. Simulations with an ocean-atmosphere/carbon-cycle model suggest that the isotope excursion can be explained by collapse of ocean primary productivity (a Strangelove Ocean) and changes in the delivery and cycling of carbon in the ocean and on land. Model results also suggest that perturbations of the global carbon cycle resulting from the extinctions led to short-term fluctuations in atmospheric pCO2 and ocean carbonate deposition, and to a long-term (>1 Ma) decrease in sedimentary burial of organic carbon in the Triassic. Deposition of calcium carbonate is a major sink of river-derived ocean alkalinity and for CO2 from the ocean/atmosphere system. The end of the Permian was marked by extinction of most calcium carbonate secreting organisms. Therefore, the reduction of carbonate accumulation made the oceans vulnerable to a build-up of alkalinity and related fluctuations in atmospheric CO2. Our model results suggest that an increase in ocean carbonate-ion concentration should cause increased carbonate accumulation rates in shallow-water settings. After the end-Permian extinctions, early Triassic shallow-water sediments show an abundance of abiogenic and microbial carbonates that removed CaCO3 from the ocean and may have prevented a full 'ocean-alkalinity crisis' from developing.

  15. Recovery and diversification of marine communities following the late Permian mass extinction event in the western Palaeotethys

    Foster, William J.; Sebe, Krisztina


    The recovery of benthic invertebrates following the late Permian mass extinction event is often described as occurring in the Middle Triassic associated with the return of Early Triassic Lazarus taxa, increased body sizes, platform margin metazoan reefs, and increased tiering. Most quantitative palaeoecological studies, however, are limited to the Early Triassic and the timing of the final phase of recovery is rarely quantified. Here, quantitative abundance data of benthic invertebrates were collected from the Middle Triassic (Anisian) succession of the Mecsek Mountains (Hungary), and analysed with univariate and multivariate statistics to investigate the timing of recovery following the late Permian mass extinction. These communities lived in a mixed siliciclastic-carbonate ramp setting on the western margin of the Palaeotethys Ocean. The new data presented here is combined with the previously studied Lower Triassic succession of the Aggtelek Karst (Hungary), which records deposition of comparable facies and in the same region of the Palaeotethys Ocean. The Middle Triassic benthic fauna can be characterised by three distinct ecological states. The first state is recorded in the Viganvár Limestone Formation representing mollusc-dominated communities restricted to above wave base, which are comparable to the lower and mid-Spathian Szin Marl Formation faunas. The second state is recorded in the Lapis Limestone Formation and records extensive bioturbation that is not limited to wave base and is comparable to the upper Spathian Szinpetri Limestone Formation. The third ecological state occurs in the Zuhánya Limestone Formation which was deposited in the Pelsonian Binodosus Zone, and has a more 'Palaeozoic' structure with sessile brachiopods dominating assemblages for the first time in the Mesozoic. The return of community-level characteristics to pre-extinction levels and the diversification of invertebrates suggests that the final stages of recovery and the radiation

  16. Structure of the Anayet Permian basin (Axial Zone, Central Pyrenees)

    Rodriguez, L.; Cuevas, J.; Tubía, J. M.


    The Anayet Permian basin was generated by strike-slip tectonics that opened subsident basins with pull-apart geometries in the western Spanish Axial Zone (between the Aragon and Tena valleys). A continental succession of Permian age, that represents the first post-variscan deposits in the area, fills the basin and covers discordantly Devonian to Carboniferous limestones, sandstones and slates. Permian deposits have been classically divided in four main detrital groups, with three basic volcanic episodes interbedded (Gisbert, 1984, Bixel, 1987): the Grey Unit (50-120 m, Estefanian to Kungurian) with slates, conglomerates, tobaceous slates, coal and pyroclastic deposits, the Transition Unit (50 m maximum) showing grey and red sandstones and lutites with oolitic limestones intercalated, the Lower Red Unit (250 m) composed of cross-bedded red sandstones and andesitic volcanic rocks at the top, and finally the Upper Red Unit (400 m minimum, top eroded) formed by three fining up megasequences of carbonates, red sandstones and lutites with lacustrine carbonates intercalated and alkali basalts at the top. Increasingly older rocks are found towards the western part of the basin, where its depocenter is located. South-vergent angular folds deform the Permian sedimentary succession. Fold axes are N115 °E-trending, almost horizontal and are characterized by a remarkably constant orientation. Folds exhibit a long limb dipping slightly to the north and a short vertical limb, occasionally reversed. In the Anayet basin four main folds, with a wavelength of 400 m, can be distinguished, two anticlines and two synclines, with minor folds associated. Related to the angular folds an axial plane foliation, E-trending and dipping 40 to 60° to the north, is developed in the lutites. The more competent rocks, conglomerates and breccias, only locally show a spaced fracture cleavage. No main thrusts have been detected in Permian rocks. However, minor scale decollements, usually low angle

  17. Early

    Kamel Abd Elaziz Mohamed


    Conclusion: Early PDT is recommended for patients who require prolonged tracheal intubation in the ICU as outcomes like the duration of mechanical ventilation length of ICU stay and hospital stay were significantly shorter in early tracheostomy.

  18. Fluid-rock reactions in an evaporitic melange, Permian Haselgebirge, Austrian Alps

    Spotl, C.; Longstaffe, F.J.; Ramseyer, K.; Kunk, M.J.; Wiesheu, R.


    Tectonically isolated blocks of carbonate rocks present within the anhydritic Haselgebirge melange of the Northern Calcareous Alps record a complex history of deformation and associated deep-burial diagenetic to very low-grade metamorphic reactions. Fluids were hot (up to ~ 250 ??C) and reducing brines charged with carbon dioxide. Individual carbonate outcrops within the melange record different regimes of brine-rock reactions, ranging from pervasive dolomite recrystallization to dedolomitization. Early diagenetic features in these carbonates were almost entirely obliterated. Matrix dolomite alteration was related to thermochemical sulphate reduction (TSR) recognized by the replacement of anhydrite by calcite + pyrite ?? native sulphur. Pyrite associated with TSR is coarsely crystalline and characterized by a small sulphur isotope fractionation relative to the precursor Permian anhydrite. Carbonates associated with TSR show low Fe/Mn ratios reflecting rapid reaction of ferrous iron during sulphide precipitation. As a result, TSR-related dolomite and calcite typically show bright Mn(II)-activated cathodoluminescence in contrast to the dull cathodoluminescence of many (ferroan) carbonate cements in other deep-burial settings. In addition to carbonates and sulphides, silicates formed closely related to TSR, including quartz, K-feldspar, albite and K-mica. 40Ar/39Ar analysis of authigenic K-feldspar yielded mostly disturbed step-heating spectra which suggest variable cooling through the argon retention interval for microcline during the Late Jurassic. This timing coincides with the recently recognized subduction and closure of the Meliata-Hallstatt ocean to the south of the Northern Calcareous Alps and strongly suggests that the observed deep-burial fluid-rock reactions were related to Jurassic deformation and melange formation of these Permian evaporites.

  19. Dolomitized cells within chert of the Permian Assistência Formation, Paraná Basin, Brazil

    Calça, Cléber P.; Fairchild, Thomas R.; Cavalazzi, Barbara; Hachiro, Jorge; Petri, Setembrino; Huila, Manuel Fernando Gonzalez; Toma, Henrique E.; Araki, Koiti


    well-preserved organic-walled cyanobacteria and portions of microbial mat. Clearly, dolomitization began very early in the microbial mats, prior to compaction of the sediment or full obliteration of cellular remains, followed very closely by silicification thereby impeding continued degradation and providing a window onto very well-preserved Permian microbial mats.

  20. Relationships between ocean anoxia, the biological pump, and marine animal life during the Permian-Triassic mass extinction (Invited)

    Meyer, K. M.; Schaal, E. K.; Payne, J.


    Ocean anoxia/euxinia and carbon cycle instability have long been linked to the end-Permian mass extinction and the Early Triassic interval of delayed or interrupted biotic recovery. Many hypotheses to explain this extinction event invoke the release of greenhouse gases during the emplacement of the Siberian Traps, which likely triggered abrupt changes in marine biogeochemical cycling, atmospheric chemistry, and biodiversity. However, the precise ways in which volcanism and these perturbations are linked and how they governed the tempo and mode of biotic recovery remain poorly understood. Here we highlight new C, Ca, and Sr isotopic data that serve to link volcanic CO2 inputs to changes in marine biogeochemistry and environmental change. We then examine the relationship between ocean biogeochemistry, the biological pump, and marine animal ecosystems during the end-Permian mass extinction and Early Triassic recovery. Finally, we use numerical simulations to probe whether these relationships also explain broad Phanerozoic trends in ocean nutrient status, anoxia, and productivity of marine ecosystems.

  1. Structure and regional significance of the Late Permian(?) Sierra Nevada - Death Valley thrust system, east-central California

    Stevens, C.H.; Stone, P.


    An imbricate system of north-trending, east-directed thrust faults of late Early Permian to middle Early Triassic (most likely Late Permian) age forms a belt in east-central California extending from the Mount Morrison roof pendant in the eastern Sierra Nevada to Death Valley. Six major thrust faults typically with a spacing of 15-20 km, original dips probably of 25-35??, and stratigraphic throws of 2-5 km compose this structural belt, which we call the Sierra Nevada-Death Valley thrust system. These thrusts presumably merge into a de??collement at depth, perhaps at the contact with crystalline basement, the position of which is unknown. We interpret the deformation that produced these thrusts to have been related to the initiation of convergent plate motion along a southeast-trending continental margin segment probably formed by Pennsylvanian transform truncation. This deformation apparently represents a period of tectonic transition to full-scale convergence and arc magmatism along the continental margin beginning in the Late Triassic in central California. ?? 2005 Elsevier B.V. All rights reserved.

  2. Peronosporomycetes (Oomycota from a Middle Permian permineralised peat within the Bainmedart Coal Measures, Prince Charles Mountains, Antarctica.

    Ben J Slater

    Full Text Available The fossil record of Peronosporomycetes (water moulds is rather sparse, though their distinctive ornamentation means they are probably better reported than some true fungal groups. Here we describe a rare Palaeozoic occurrence of this group from a Guadalupian (Middle Permian silicified peat deposit in the Bainmedart Coal Measures, Prince Charles Mountains, Antarctica. Specimens are numerous and comprise two morphologically distinct kinds of ornamented oogonia, of which some are attached to hyphae by a septum. Combresomyces caespitosus sp. nov. consists of spherical oogonia bearing densely spaced, long, hollow, slender, conical papillae with multiple sharply pointed, strongly divergent, apical branches that commonly form a pseudoreticulate pattern under optical microscopy. The oogonia are attached to a parental hypha by a short truncated stalk with a single septum. Combresomyces rarus sp. nov. consists of spherical oogonia bearing widely spaced, hollow, broad, conical papillae that terminate in a single bifurcation producing a pair of acutely divergent sharply pointed branches. The oogonium bears a short truncate extension where it attaches to the parental hypha. We propose that similarities in oogonium shape, size, spine morphology and hyphal attachment between the Permian forms from the Prince Charles Mountains and other reported Peronosporomycetes from Devonian to Triassic strata at widely separated localities elsewhere in the world delimit an extinct but once cosmopolitan Palaeozoic to early Mesozoic branch of the peronosporomycete clade. We name this order Combresomycetales and note that it played an important role in late Palaeozoic and early Mesozoic peatland ecosystems worldwide.




    Full Text Available The best preserved Permian-Triassic boundary beds in Turkey are found in the Hadim region of the central Taurides. The succession is exposed in one of the allochthonous units of the Tauride Belt, the Aladag Unit, whose stratigraphy includes beds ranging from the Devonian to the Cretaceous systems. In the Aladag Unit, the Permian-Triassic boundary beds are entirely composed of carbonates. The Permian portion of these beds belongs to the Paradagmarita Zone, whereas the lowermost Triassic contains the Lower Griesbachian marker Rectocornuspira kalhori. The uppermost Permian carbonates, composed of meter-scale upward shallowing subtidal cycles, are characterized by oolitic limestones of regressive character at the top and are overlain sharply by Lower Triassic stromatolites. Cyclic Upper Permian carbonates are interpreted as highstand sytems tract deposits of the last third-order sequence of the Permian System. The Permian-Triassic boundary is an unconformity corresponding to both erosional and non-depositional hiatuses. The gap at the Permian-Triassic boundary partially corresponds to the shelf-margin systems tract and partly to the transgressive systems tract of the overlying third-order sequence. Stromatolites are interpreted as transgressive systems tract deposits. Special issueInternational Conference on Paleozoic Foraminifera, Paleoforams 2001Edited by Demir Altiner (Guest Editor

  4. Olson's Extinction and the latitudinal biodiversity gradient of tetrapods in the Permian.

    Brocklehurst, Neil; Day, Michael O; Rubidge, Bruce S; Fröbisch, Jörg


    The terrestrial vertebrate fauna underwent a substantial change in composition between the lower and middle Permian. The lower Permian fauna was characterized by diverse and abundant amphibians and pelycosaurian-grade synapsids. During the middle Permian, a therapsid-dominated fauna, containing a diverse array of parareptiles and a considerably reduced richness of amphibians, replaced this. However, it is debated whether the transition is a genuine event, accompanied by a mass extinction, or whether it is merely an artefact of the shift in sampling from the palaeoequatorial latitudes to the palaeotemperate latitudes. Here we use an up-to-date biostratigraphy and incorporate recent discoveries to thoroughly review the Permian tetrapod fossil record. We suggest that the faunal transition represents a genuine event; the lower Permian temperate faunas are more similar to lower Permian equatorial faunas than middle Permian temperate faunas. The transition was not consistent across latitudes; the turnover occurred more rapidly in Russia, but was delayed in North America. The argument that the mass extinction is an artefact of a latitudinal biodiversity gradient and a shift in sampling localities is rejected: sampling correction demonstrates an inverse latitudinal biodiversity gradient was prevalent during the Permian, with peak diversity in the temperate latitudes.

  5. 40 CFR 81.242 - Pecos-Permian Basin Intrastate Air Quality Control Region.


    ... Air Quality Control Regions § 81.242 Pecos-Permian Basin Intrastate Air Quality Control Region. The Pecos-Permian Basin Intrastate Air Quality Control Region (New Mexico) consists of the territorial area... Quality Control Region. 81.242 Section 81.242 Protection of Environment ENVIRONMENTAL PROTECTION...

  6. Trace Metals, Rare Earths, Carbon and Pb Isotopes as Proxies of Environmental Catastrophe at the Permian - Triassic Boundary in Spiti Himalayas, India

    Ghosh, N.; Basu, A. R.; Garzione, C. N.; Ghatak, A.; Bhargava, O. N.; Shukla, U. K.; Ahluwalia, A. D.


    Himalayan sediments from Spiti Valley, India preserve geochemical signatures of the Permian - Triassic (P-Tr) mass extinction in the Neo-Tethys Ocean. We integrate new sedimentological and fossil record with high-resolution geochemical-isotopic data from Spiti that reveals an ecological catastrophe of global proportions. Trace elements of U, Th, Nb, Ta, Zr, Hf, the rare earths (REE) and carbon, oxygen and lead isotopes measured across the P-Tr boundary in Spiti are used as proxies for evaluating abrupt changes in this continental shelf environment. δ13Corg excursions of 2.4‰ and 3.1‰ in Atargu and Guling P-Tr sections in Spiti Valley are associated with an abrupt fall of biological productivity while δ13Ccarb and δ18Ocarb record of these sediments shows effects of diagenesis. Here, the P-Tr boundary is compositionally distinct from the underlying Late Permian gray shales, as a partly gypsiferous ferruginous layer that allows additional geochemical-isotopic investigation of sedimentary sources. Conspicuous Ce - Eu anomalies in the light REE-enriched Late Permian shales reflect the source composition of the adjacent Panjal Trap basalts of Kashmir. An abrupt change of this source to continental crust is revealed by Nb - Ta and Zr - Hf anomalies at the P-Tr ferruginous layer and continuing through the overlying Early Triassic carbonate rocks. Pb concentration and isotope ratios of 206Pb/204Pb, 207Pb/204Pb and 208Pb/204Pb identify changes in the sedimentary element flux, distinguishing the Late Permian shales from the distinct siliciclastic continental crustal signature in the Early Triassic carbonates. These geochemical-isotopic constraints on the sedimentary geochemistry of one of the most critical transitions in geological record establish the utility of multi-proxy datasets for paleoenvironmental reconstructions.

  7. Climate warming during and in the aftermath of the End-Permian mass extinction (Arne Richter Award for Outstanding ECSs Lecture)

    Sun, Yadong


    The end-Permian mass extinction saw the most catastrophic diversity loss in the Phanerozoic. The extinction event was accompanied with a rapid temperature raise from 25 °C to 32 °C across the Permian-Triassic boundary, suggesting a warming climate might have played an important role in the extinction event. This high amplitude warming of 8-10 °C is seen in South China, Iran and Armenia, pointing to a true global signature. Oxygen isotope data measured from conodont phosphate in South China suggest that the temperature continued to increase in the Early Triassic and reached the first thermal maximum in the late Griesbachian. The late Griesbachian Thermal Maximum accompanied with the extinction of many Permian holdovers, such as the conodont Hindeodus and the ammonoid Otoceras. The following substage, the Dienerian, saw a 3-4 °C temperature decrease which coincides with a transient recovery pulse in which several groups began to diversify. The early and middle Smithian represent a relatively stable high temperature plateau but the late Smithian saw a further 2°C temperature increase to produce sea surface temperatures that exceeded 40°C. The Late Smithian Thermal Maximum coincided with major diversity loss of marine nektons such as conodont and ammonoid and minor extinctions among many other groups such as bivalves and gastropods. The Spathian saw an initial cooling trend followed by relatively stable temperatures in the middle part and further cooling at the end of this stage and stabilization of temperatures in the earliest Middle Triassic. High amplitude temperature changes may have played a vital role in controlling the pace of recovery in the aftermath of the end Permian mass extinction.

  8. Permian volcanisms in eastern and southeastern margins of the Jiamusi Massif, northeastern China: zircon U-Pb chronology, geochemistry and its tectonic implications

    MENG En; XU WenLiang; YANG DeBin; PEI FuPing; Yu Yang; Zhang XingZhou


    LA-ICP-MS zircon U-Pb dating and geochemical data for the Late Paleozoic volcanic rocks from eastern and southeastern margins of the Jiamusi Massif are presented to understand the regional tectonic evolution. Zircons from eight representative volcanic rocks are euhedral-subhedral in shape and dis-play striped absorption and fine-scale oscillatory growth zoning as well as high Th/U ratios (0.33-2.37), implying a magmatic origin. The dating results show that the Late Paleozoic volcanic rocks in the study area can be divided into two stages, I.e., the Early Permian (a weighted mean 206pb/238U age of 288 Ma) and the Middle Permian volcanisms (a weighted mean 206Pb/238U age of 268 Ma). The former is com-posed mainly of basalt, basaltic-andesite, andesite and minor dacite. They are characterized by low SiO2 contents, high Mg# (0.40-0.59), enrichment in Na (Na2O/K2O = 1.26-4.25) and light rare earth elements (LREEs), relative depletion in heavy rare earth elements (HREEs) and high field strength ele-ments (HFSEs), indicating that an active continental margin setting could exist in the eastern margin of the Jiamusi Massif in the Early Permian. The latter consists mainly of rhyolite and minor dacite with high SiO2 (77.23%-77.52%), low MgO (0.11%-0.14%), enrichment in K2O (Na2O/K2O ratios <0.80) and Rb, Th, U and depletion in Eu, Sr, P and Ti, implying a crust-derived origin. Therefore, it is proposed that the Middle Permian volcanic rocks could have formed under the collision of the Jiamusi and the Khanka Massifs.

  9. Permian oolitic carbonates from the Baoshan Block in western Yunnan, China, and their paleoclimatic and paleogeographic significance

    Huang, Hao; Jin, Xiaochi; Li, Fei; Shen, Yang


    Marine carbonate ooids are environment—sensitive and hence valuable for paleoclimatic and paleogeographic reconstructions. This paper describes Permian ooids from the Baoshan Block in western Yunnan, China, in order to offer a new means to refine the uncertain paleogeographic details of this Gondwana-derived block. Four major types of ooids (micritic ooids, compound ooids, leached ooids and half-moon ooids) are documented from the Hewanjie Formation in the northern and the Shazipo Formation in the southern Baoshan Block. These ooids are dated via biostratigraphic analysis to be Wordian-early Wuchiapingian and signify an ameliorated shallow-marine temperature for the Guadalupian strata of the Baoshan Block. Results of this study, coupled with literature data, reveal diachronous debut of Permian ooids among the Gondwana-derived blocks: mostly Sakmarian in Central Taurides of Turkey, Central Iran, Central Pamir and Karakorum Block versus Wordian-Capitanian in Baoshan Block, Peninsular Thailand and South Qiangtang. In contrast, Asselian-Sakmarian strata of Baoshan Block as well as Peninsular Thailand and South Qiangtang are characterized by glaciomarine diamictites. These observations suggest that the Baoshan Block was probably situated at a considerably higher paleolatitude under distinct influence of Gondwana glaciation during the Asselian-Sakmarian than those blocks yielding Sakmarian ooids. Moreover, marine ooids are virtually absent nearby the equator within the Permian Tethys, similar to the modern situation. The Baoshan Block is accordingly interpreted to drift to warm-water southern mid-latitudes during the Wordian-Capitanian and remain to the south of Central Iran, Karakorum Block and South China, which were equatorially located in the Capitanian.

  10. The Permian-Triassic boundary & mass extinction in China

    Metcalfe, I.; Nicoll, R.S.; Mundil, R.; Foster, C.; Glen, J.; Lyons, J.; Xiaofeng, W.; Cheng-Yuan, W.; Renne, P.R.; Black, L.; Xun, Q.; Xiaodong, M.


    The first appearance of Hindeodus parvus (Kozur & Pjatakova) at the Permian-Triassic (P-T) GSSP level (base of Bed 27c) at Meishan is here confirmed. Hindeodus changxingensis Wang occurs from Beds 26 to 29 at Meishan and appears to be restricted to the narrow boundary interval immediately above the main mass extinction level in Bed 25. It is suggested that this species is therefore a valuable P-T boundary interval index taxon. Our collections from the Shangsi section confirm that the first occurrence of Hindeodus parvus in that section is about 5 in above the highest level from which a typical Permian fauna is recovered. This may suggest that that some section may be missing at Meishan. The age of the currently defined Permian-Triassic Boundary is estimated by our own studies and a reassessment of previous worker's data at c. 253 Ma, slightly older than our IDTIMS 206Pb/238U age of 252.5 ??0.3 Ma for Bed 28, just 8 cm above the GSSP boundary (Mundil et al., 2001). The age of the main mass extinction, at the base of Bed 25 at Meishan, is estimated at slightly older than 254 Ma based on an age of >254 Ma for the Bed 25 ash. Regardless of the absolute age of the boundary, it is evident that the claimed <165,000 y short duration for the negative carbon isotope excursion at the P-T boundary (Bowring et al., 1998) cannot be confirmed. Purportedly extraterrestrial fullerenes at the boundary (Hecker et al., 2001) have equivocal significance due to their chronostratigraphic non-uniqueness and their occurrence in a volcanic ash.

  11. Deep electrical conductivity structure of the Rio Grande Rift in Colorado and New Mexico: Early results from a two-year magnetotelluric study

    Feucht, D. W.; Bedrosian, P.; Sheehan, A. F.


    A wideband and long-period magnetotelluric experiment is underway across the Rio Grande Rift in Colorado and New Mexico in order to provide constraints on the thermal and rheological state of the lithosphere beneath this region of intra-continental extension. Magnetotellurics is a passive source electromagnetic technique that at long periods has depth penetration into the deep crust and upper mantle. Important questions about continental rifting remain unresolved, including the role of magmatism, volatiles and inherited lithospheric structure in the initiation and development of rifting. Recent seismic imaging studies show thinned crust and low seismic wavespeeds in the crust and upper mantle beneath the Rio Grande Rift. New and ongoing geodetic work confirms the low strain-rate environment of the region yet shows surprisingly uniform deformation over an area far wider than the rift's physiographic expression. Electrical conductivity models from this experiment will provide information complementary to these studies and can be used to determine the relative contributions of thermal and compositional heterogeneity in the crust and upper mantle to processes of continental extension. Over the past two years, magnetotelluric data has been collected at ~100 site locations along three 450 km long east-west transects of the rift axis. These three profiles extend across the northern, central, and southern portions of the rift and include sites in the High Plains, Colorado Front Range, southern Rocky Mountains, San Juan Basin, Sangre de Cristo Mountains, and southern Basin and Range along the New Mexico/Mexico border. A comparison of results from these segments will be used to examine along-strike variation in the spatial extent of rifting and associated modification of the lithosphere. Data assessment shows high-quality signal to periods in excess of 10 000 s, which corresponds to upper-mantle depths in this region of high upper-crustal conductivity and low crustal

  12. Avaliação psicométrica de três questionários sobre o historial familiar Evaluation of psychometric properties of three questionnaires addressing early experiences in family context

    Ivandro Soares Monteiro


    Full Text Available CONTEXTO: A família é a arena primária na qual a criança desenvolve a sua sociabilização. As experiências tidas neste sistema podem constituir um factor protector ou de risco, contribuindo para a sua vulnerabilidade ou resiliência diante de desordens mentais quando adulto. A avaliação da história familiar de experiências emocionais relevantes é problemática, e o uso de instrumentos psicométricos pode auxiliar a superar certas limitações metodológicas. OBJECTIVOS: Desenvolver três subquestionários para avaliação das experiências emocionais decorridas no contexto familiar da infância até a adolescência. Esses instrumentos são adaptações para o português do Family Background Questionnaire (FBQ tratando dos cuidados paliativos e do ambiente familiar e são administrados retrospectivamente a adultos. MÉTODOS: Três versões do FBQ que tratam de (1 cuidados paternos, (2 cuidados maternos e (3 ambiente familiar foram produzidas. Os questionários foram aplicados a 280 participantes. A avaliação das propriedades psicométricas desses questionários teve como base a determinação da consistência interna, análise factorial e correlação de Spearman. RESULTADOS: Os três questionários mostraram-se adequados para avaliação de seus respectivos domínios. CONCLUSÕES: A avaliação objetiva das primeiras experiências emocionais no contexto familiar produz insights importantes para a compreensão do desenvolvimento psicológico, com implicações clínicas e de investigação.BACKGROUND: Family is the primary setting where children develop their social skills. Early experiences may represent protective or risk factors that interact with their vulnerability to or resilience against mental disorders in adulthood. The assessment of the family history of relevant emotional experiences is problematic, and the use of psychometric instruments may help overcome certain methodological limitations. OBJECTIVES: To develop three

  13. A sudden end-Permian mass extinction (Invited)

    Shen, S.


    The end-Permian mass extinction is the largest of the Phanerozoic. In the immediate aftermath the marine ecosystem was dominated by microbial and communities with disaster taxa. Plausible kill mechanism includes an extremely rapid, explosive release of gases such as carbon dioxide, methane and hydrogen sulfide. Siberian flood volcanism has been suggested as the most possible mechanism to trigger the massive release of greenhouse gases from volcanic eruptions and interaction of magmas with carbon from thick organic-rich deposits or rapid venting of coal-derived methane or massive combustion of coal. A sharp δ13C isotopic excursion, rapid disappearance of carbonate benthic communities and δ18O data from conodont apatite suggest rapid global warming. The end-Permian mass extinction occurred in less than 200,000 years. This extinction interval is constrained by two ash beds (Beds 25 and 28) at the Meishan section. However, the extinction patterns remain controversial largely due to the condensed nature of the Meishan sections. Geochemical signals and their interpretations are also contentious. Thus, the level of achievable stratigraphic resolution becomes crucial to determine the nature of the event and a detailed study of the extinction interval is essential to unravel the extinction pattern, chemostratigraphy, and the causes. However, the extinction interval at Meishan is only 26 cm thick and contains distinct gaps at the Permian-Triassic boundary (PTB) and possibly the base of Bed 25. Thus, it is impossible to resolve a detailed extinction pattern. Studying expanded sections is crucial to understand the detailed events before, during and after the main extinction. In this report, we show a highly-expanded Permian-Triassic boundary section in Guangxi Province, South China. The last 4.5 m between beds 22 and 28 of the Meishan sections is represented by a sequence of ~560 m at the section and the extinction interval between beds 24e and 28 at Meishan is represented

  14. Factors Controlling Pre-Columbian and Early Historic Maize Productivity in the American Southwest, Part 1: The Southern Colorado Plateau and Rio Grande Regions

    Benson, L.V.


    Maize is the New World's preeminent grain crop and it provided the economic basis for human culture in many regions within the Americas. To flourish, maize needs water, sunlight (heat), and nutrients (e. g., nitrogen). In this paper, climate and soil chemistry data are used to evaluate the potential for dryland (rainon-field) agriculture in the semiarid southeastern Colorado Plateau and Rio Grande regions. Processes that impact maize agriculture such as nitrogen mineralization, infiltration of precipitation, bare soil evaporation, and transpiration are discussed and evaluated. Most of the study area, excepting high-elevation regions, receives sufficient solar radiation to grow maize. The salinities of subsurface soils in the central San Juan Basin are very high and their nitrogen concentrations are very low. In addition, soils of the central San Juan Basin are characterized by pH values that exceed 8.0, which limit the availability of both nitrogen and phosphorous. In general, the San Juan Basin, including Chaco Canyon, is the least promising part of the study area in terms of dryland farming. Calculations of field life, using values of organic nitrogen for the upper 50 cm of soil in the study area, indicate that most of the study area could not support a 10-bushel/acre crop of maize. The concepts, methods, and calculations used to quantify maize productivity in this study are applicable to maize cultivation in other environmental settings across the Americas. ?? 2010 US Government.

  15. Early seafloor spreading in the South Atlantic: new evidence for M-series magnetochrons north of the Rio Grande Fracture Zone

    Bird, Dale E.; Hall, Stuart A.


    Recent tectonic reconstructions of the South Atlantic have partitioned the ocean basin into several segments based upon one or more proposed intraplate South American deformation zones. In several of these reconstructions, opening of the southern segment(s) by seafloor spreading prior to Aptian-Albian time is accompanied by contemporaneous strike-slip motion along an intraplate boundary extending southeastward from the Andean Cochabamba—Santa Cruz bend to the Rio Grande Fracture Zone (RGFZ). We have examined new magnetic data over the Pelotas, Santos and Campos Basins, offshore Argentina and Brazil, acquired by ION-GXT in tandem with long-offset, long record seismic reflection data, and identified seafloor spreading anomalies M4, M3, M2 and M0 (˜131, ˜129, ˜128 and ˜125 Ma). Integrating these results with our earlier work, we have been able to correlate magnetochrons M4, M3, M2 and M0 north and south of the RGFZ on the South American margin, and north and south of the Walvis Ridge on the African side. Our results are therefore inconsistent with diachronous opening models that involve substantial continental strike-slip motion north of RGFZ during M4 to M0 time. Although the ocean basin may have opened from south to north, our results indicate that seafloor spreading began north of the RGFZ earlier than previously proposed.

  16. Trophic network models explain instability of Early Triassic terrestrial communities.

    Roopnarine, Peter D; Angielczyk, Kenneth D; Wang, Steve C; Hertog, Rachel


    Studies of the end-Permian mass extinction have emphasized potential abiotic causes and their direct biotic effects. Less attention has been devoted to secondary extinctions resulting from ecological crises and the effect of community structure on such extinctions. Here we use a trophic network model that combines topological and dynamic approaches to simulate disruptions of primary productivity in palaeocommunities. We apply the model to Permian and Triassic communities of the Karoo Basin, South Africa, and show that while Permian communities bear no evidence of being especially susceptible to extinction, Early Triassic communities appear to have been inherently less stable. Much of the instability results from the faster post-extinction diversification of amphibian guilds relative to amniotes. The resulting communities differed fundamentally in structure from their Permian predecessors. Additionally, our results imply that changing community structures over time may explain long-term trends like declining rates of Phanerozoic background extinction.

  17. Boron isotopes in brachiopods during the end-Permian mass extinction: constraints on pH evolution and seawater chemistry

    Jurikova, Hana; Gutjahr, Marcus; Liebetrau, Volker; Brand, Uwe; Posenato, Renato; Garbelli, Claudio; Angiolini, Lucia; Eisenhauer, Anton


    The global biogeochemical cycling of carbon is fundamental for life on Earth with the ocean playing a key role as the largest and dynamically evolving CO2 reservoir. The boron isotope composition (commonly expressed in δ11B) of marine calcium carbonate is considered to be one of the most reliable paleo-pH proxies, potentially enabling us to reconstruct past ocean pH changes and understand carbon cycle perturbations along Earth's geological record (e.g. Foster et al., 2008; Clarkson et al., 2015). Brachiopods present an advantageous and largely underutilised archive for Phanerozoic carbon cycle reconstructions considering their high abundance in the geological record and its origin dating back to the early Cambrian. Moreover, their shell made of low-magnesium calcite makes these marine calcifiers more resistant to post-depositional diagenetic alteration of primary chemical signals. We have investigated the δ11B using MC-ICP-MS (Neptune Plus) and B/Ca and other elemental ratios (Mg/Ca, Sr/Ca, Al/Ca, Li/Ca, Ba/Ca, Na/Ca and Fe/Ca) using ICP-MS-Quadrupole (Agilent 7500cx) from the same specimens in pristine brachiopod shells from two sections from northern Italy during the Late Permian. These sections cover the δ13C excursion in excess of ˜4 ‰ (Brand et al., 2012) and are associated with major climate and environmental perturbations that lead to the mass extinction event at the Permian-Triassic boundary. Particular emphasis will be placed on the implications of our new paleo-pH estimates on the seawater chemistry during the Late Permian. Brand, U., Posenato, R., Came, R., Affek, H., Angiolini, L., Azmy, K. and Farabegoli, E.: The end-Permian mass extinction: A rapid volcanic CO2 and CH4-climatic catastrophe, Chemical Geology 323, 121-144, doi:10.1016/j.chemgeo.2012.06.015, 2012. Clarkson, M.O., Kasemann, S.A., Wood, R.A., Lenton, T.M., Daines, S.J., Richoz, S., Ohnemueller, F., Meixner, A., Poulton, S.W. and Tipper, E.T.: Ocean acidification and the Permo

  18. Sickle cell disease: acute clinical manifestations in early childhood and molecular characteristics in a group of children in Rio de Janeiro

    da Silva Filho, Isaac Lima; Ribeiro, Georgina Severo; Moura, Patrícia Gomes; Vechi, Monica Longo; Cavalcante, Andréa Cony; de Andrada-Serpa, Maria José


    Objectives To describe clinical events of sickle cell disease and the correlation with β-globin haplotypes and α-thalassemia in under 6-year-old children. Methods A retrospective study was conducted of under 6-year-old children from the neonatal screening program in Rio de Janeiro. Forty-eight male and 48 female children were enrolled in this study, 79 with sickle cell anemia and 17 with hemoglobin SC. The mean age was 29.9 (standard deviation = 20.9) months, 62 (16.2 ± 8.6) were aged between 0-3 years old and 34 (54.9 ± 11.3) were from 3-6 years old. Painful events, acute splenic sequestration, hemolytic crises, hand-foot and acute chest syndromes and infections were evaluated. Results The events were more frequent in under 3-year-old children, 94% of children had at least one episode. Infection was the most common event affecting 88.5% of children. Acute splenic sequestration took place earlier, while painful crises and acute chest syndromes in under 6-year-old children. Thal-α 3.7 was observed in 20.9% of cases. Bantu was the most frequent haplotype found, followed by Benin. No correlation was observed between clinical events and β-globin haplotypes. Children with sickle cell anemia and α-thalassemia have less infectious events. No correlation was found among these polymorphisms and clinical events, however, the majority of children with Bantu/Bantu and without α-thalassemia had more clinical events. PMID:23049419

  19. Paleomagnetism of Permian and Triassic rock, central Chilean Andes

    Forsythe, Randall D.; Kent, Dennis V.; Mpodozis, Constantino; Davidson, John

    The first paleomagnetic data from Permian and Triassic formations west of the Andean divide are presented. Four formations of Permian or Triassic age in the central Chilean Andes have been investigated: two are located in the coastal ranges, and two are in the main cordillera. Of the formations in the main cordillera (Pastos Blancos and Matahuaico formations), only the Pastos Blancos Formation has yielded characteristic directions. While a fold test is absent, magnetizations are most likely secondary and yield pre-tilt corrected concordant inclinations, but yield declinations discordant 30° clockwise in comparison to the South American apparent polar wander path. Both formations from the coastal ranges (Cifuncho and Pichidangui formations) yielded stable directions. Postfolding magnetizations in the Cifuncho Formation also show declinations discordant 30° clockwise and concordant inclinations. The Pichidangui Formation has two stable components: one of postfolding age is concordant to apparent polar wander path data, and one of probable prefolding (Late Triassic) age is concordant in declination, but discordant in inclination. Further work is needed to better define the prefolding magnetizations in the Pichidangui Formation, but at present these preliminary results are the first paleomagnetic signs of displaced terranes along the Pacific margin of Chile. If correct, the results suggest that the Pichidangui Formation was some 15° of latitude farther south during the Late Triassic and had likely moved northward to its present latitudinal position with respect to cratonic South America by Middle to Late Jurassic.

  20. Calcium isotope constraints on the end-Permian mass extinction.

    Payne, Jonathan L; Turchyn, Alexandra V; Paytan, Adina; Depaolo, Donald J; Lehrmann, Daniel J; Yu, Meiyi; Wei, Jiayong


    The end-Permian mass extinction horizon is marked by an abrupt shift in style of carbonate sedimentation and a negative excursion in the carbon isotope (delta(13)C) composition of carbonate minerals. Several extinction scenarios consistent with these observations have been put forward. Secular variation in the calcium isotope (delta(44/40)Ca) composition of marine sediments provides a tool for distinguishing among these possibilities and thereby constraining the causes of mass extinction. Here we report delta(44/40)Ca across the Permian-Triassic boundary from marine limestone in south China. The delta(44/40)Ca exhibits a transient negative excursion of approximately 0.3 per thousand over a few hundred thousand years or less, which we interpret to reflect a change in the global delta(44/40)Ca composition of seawater. CO(2)-driven ocean acidification best explains the coincidence of the delta(44/40)Ca excursion with negative excursions in the delta(13)C of carbonates and organic matter and the preferential extinction of heavily calcified marine animals. Calcium isotope constraints on carbon cycle calculations suggest that the average delta(13)C of CO(2) released was heavier than -28 per thousand and more likely near -15 per thousand; these values indicate a source containing substantial amounts of mantle- or carbonate-derived carbon. Collectively, the results point toward Siberian Trap volcanism as the trigger of mass extinction.

  1. The end-Permian mass extinction: A complex, multicausal extinction

    Erwin, D. H.


    The end-Permian mass extinction was the most extensive in the history of life and remains one of the most complex. Understanding its causes is particularly important because it anchors the putative 26-m.y. pattern of periodic extinction. However, there is no good evidence for an impact and this extinction appears to be more complex than others, involving at least three phases. The first began with the onset of a marine regression during the Late Permian and resulting elimination of most marine basins, reduction in habitat area, and increased climatic instability; the first pulse of tetrapod extinctions occurred in South Africa at this time. The second phase involved increased regression in many areas (although apparently not in South China) and heightened climatic instability and environmental degradation. Release of gas hydrates, oxidation of marine carbon, and the eruption of the Siberian flood basalts occurred during this phase. The final phase of the extinction episode began with the earliest Triassic marine regression and destruction of nearshore continental habitats. Some evidence suggests oceanic anoxia may have developed during the final phase of the extinction, although it appears to have been insufficient to the sole cause of the extinction.

  2. Flourishing ocean drives the end-Permian marine mass extinction.

    Schobben, Martin; Stebbins, Alan; Ghaderi, Abbas; Strauss, Harald; Korn, Dieter; Korte, Christoph


    The end-Permian mass extinction, the most severe biotic crisis in the Phanerozoic, was accompanied by climate change and expansion of oceanic anoxic zones. The partitioning of sulfur among different exogenic reservoirs by biological and physical processes was of importance for this biodiversity crisis, but the exact role of bioessential sulfur in the mass extinction is still unclear. Here we show that globally increased production of organic matter affected the seawater sulfate sulfur and oxygen isotope signature that has been recorded in carbonate rock spanning the Permian-Triassic boundary. A bifurcating temporal trend is observed for the strata spanning the marine mass extinction with carbonate-associated sulfate sulfur and oxygen isotope excursions toward decreased and increased values, respectively. By coupling these results to a box model, we show that increased marine productivity and successive enhanced microbial sulfate reduction is the most likely scenario to explain these temporal trends. The new data demonstrate that worldwide expansion of euxinic and anoxic zones are symptoms of increased biological carbon recycling in the marine realm initiated by global warming. The spatial distribution of sulfidic water column conditions in shallow seafloor environments is dictated by the severity and geographic patterns of nutrient fluxes and serves as an adequate model to explain the scale of the marine biodiversity crisis. Our results provide evidence that the major biodiversity crises in Earth's history do not necessarily implicate an ocean stripped of (most) life but rather the demise of certain eukaryotic organisms, leading to a decline in species richness.

  3. 东北地区二叠纪沉积特征及原型盆地分析%Analysis on Permian Depositional Characteristics and Proto-type Basin of Northeast China

    崔军平; 任战利; 史政; 李金翔; 王文青


    The distribution of Permian in northeast China is large, and its sedimentary facies types includes half deep sea facies, offshore facies, shore shallow lake facies and half-lake facies. The region in northeast China is in a continental-margin and epicontinental sedimentation environment during Lower Permian to Middle Permian, and the deposition center is in the Wendouermiao-Xilamulun river area. In the Later Permian, this region was a large-scale depression basin, and all suture lines already were closed, while the earth crust was lifting and the sea water was withdrawing, thus the most areas became continental depositing, and partial areas formed residual-marine depositing. As a whole, the rule of sedimentary facies of this region presents a gradual transformation from marine facies in the Early Permian to land facies in the Late Permian. The sedimentation of this region in Late Paleozoic was controlled by the suture line of Xilamulun river, and the suture line became to be closed gradually from west to east during Permian and closed completely until the Late Permian, and the closed time is earlier in west and later in east.%东北地区二叠系分布广泛,沉积相类型主要为半深海相、滨浅海相、滨浅湖相和半深湖相等.早二叠世—中二叠世东北地区主要处于大陆边缘浅海沉积环境,面积大,沉积中心位于南部温都尔庙—西拉木伦河东西向条带区.晚二叠世东北地区为大型坳陷型盆地,东北地区各缝合线均已闭合,随着地壳抬升,海水退出,大部分地区为陆相沉积,局部有残留海相沉积,总体上具有从早二叠世到晚二叠世由海相向陆相逐渐转变的规律.西拉木伦河缝合带对晚古生代沉积起控制作用,在二叠纪整个缝合带由西向东逐渐拼合,直至晚二叠世,两板块拼合完成,闭合时间具有西早东晚的特点.

  4. Biospheric perturbations during Gondwanan times: From theNeoproterozoic-Cambrian radiation to the end-Permian crisis

    Erwin, Douglas H.

    The history of marine life during the Palaeozoic is structured by the NeoproterozoicEarly Cambrian radiation, the Ordovician radiation and a diversity plateau from the Late Ordovician into the Permian. This plateau is punctuated by mass extinctions in the end-Ordovician, Late Devonian, and finally the catastrophic end-Permian mass extinction, which brings the Palaeozoic to a close. Underlying these radiations and extinctions is a constant pattern of evolutionary change, as species respond to environmental changes, biological innovations, shifting biogeography and other factors. Are these patterns structured by global processes, or are they the summation of a host of local and regional events? One school of palaeontologists has argued that these patterns are driven by biotic interactions on a global scale, with mass extinctions as perturbations to longer-term processes, but with little lasting effect. Others have accorded the primary influence to radiations and mass extinctions. New studies from of the Ordovician radiation support a third possibility, that global signals are the summation of events at scales from local to global mass extinctions. Variations in biotic diversity, at scales ranging from global evolutionary radiations and mass extinctions to regional fluctuations or the changing fortunes of individual community assemblages, reflects the vicissitudes of environmental circumstance. Determining the appropriate level of analysis (global, regional or local) will require greater biogeographic data, particularly from Gondwanan continents. This will allow the development of explicit, testable models linking geological, geochemical and palaeontological patterns to common processes.

  5. Sedimentology of the cold-climate, coal-bearing, Lower Permian 'Lower Freshwater Sequence' of Tasmania

    Martini, I.P.; Banks, M.R. (University of Guelph, Ontario (Canada). Dept. of Land Resource Science)


    The Lower Permian 'Lower Freshwater Sequence' of Tasmania consists of fluvial and coastal sediments deposited in a cold-climate setting adjacent to ice-ridden, shallow seas. The sequence represents a regressive pulse during the filling of a marine basin, possibly a fjord, up to 150 km wide and more than 270 km long, occupied by glaciers in the late Carboniferous. The climatic conditions during the early Permian in Tasmania are revealed by (1) well-developed continental scree breccias, (2) the flora (Glossopteris-Gangamopteris assemblage), (3) rarity of fossils in tidal deposits, (4) the structure of the trace fossil population (intense bioturbation in offshore muds), (5) the foramol nature of the fauna of the marine beds below and above the Lower Freshwater Sequence, a fauna dominated by brachiopods and bryozoa and containing Eurydesma, and (6) features of the sediments themselves, such as 'lonestones' in muddy offshore marine deposits, and oxygen isotopes of associated marine carbonates. Although the sedimentation in the Lower Freshwater Sequence is qualitatively analogous to that of the Quaternary of Canada, the fluvial and coastal sediments do not show evidence of ice rafting nor of frozen ground. Lack of such evidence suggests a cold-temperate rather than a subpolar setting. 64 refs., 14 figs.

  6. Smithian-Spathian boundary event: Evidence for global climatic change in the wake of the end-Permian biotic crisis

    Galfetti, Thomas; Hochuli, Peter A.; Brayard, Arnaud; Bucher, Hugo; Weissert, Helmut; Vigran, Jorunn Os


    One of the most important carbon cycle perturbations following the end-Permian mass extinction event straddles the Smithian-Spathian boundary (SSB) (Olenekian, Early Triassic). This anomaly is characterized by a prominent positive carbon isotope excursion known from Tethyan marine rocks. Its global significance is established here by a new high paleolatitude record (Spitsbergen). Paleontological evidence, such as Boreal palynological data (Barents Sea, Norway) and global patterns of ammonoid distribution, indicates a synchronous major change in terrestrial and marine ecosystems near the SSB. The reestablishment of highly diverse plant ecosystems, including the rise of woody gymnosperms and decline of the formerly dominating lycopods, is interpreted as an effect of a major climate change. This hypothesis is supported by modeling of ammonoid paleobiogeography, the distribution patterns of which are interpreted as a proxy for sea surface temperatures (SST). The latest Smithian thus appears to have been a time of a warm and equable climate as expressed by an almost flat pole to equator SST gradient. In contrast, the steep Spathian SST gradient suggests latitudinally differentiated climatic conditions. We propose that this drastic climate change and the global carbon cycle perturbation were triggered by a massive end-Smithian CO2 injection. The SSB event could therefore represent one of the causes for stepwise and delayed recovery of marine and terrestrial biotas in the wake of the end-Permian biotic crisis.

  7. AUthigenic feldspar as an indicator of paleo-rock/water interactions in Permian carbonates of the Northern Calcareous Alps, Austria

    Spotl, C.; Kralik, M.; Kunk, M.J.


    Dolostones interbedded with Upper Permian evaporites at the base of the Northern Calcareous Alps contain abundant authigenic K-feldspar. Two petrographically, structurally, and isotopically distinct generations of K-feldspar can be distinguished: crystals composed of an inclusion-rich core and a clear rim, and optically unzoned, transparent crystals. Both feldspar types have essentially identical K-feldspar end-member compositions with ??? 99.5 mole % Or component. Low oxygen isotope ratios (+16.1??? to +18.1??? SMOW) suggest precipitation from 18O-enriched, saline fluids at temperatures in excess of ??? 140??C. 40Ar/39Ar plateau-age spectra of five samples range from 145 ?? 1 to 144 ?? 1 Ma (Early Berriasian) and suggest that both types of feldspar were formed within an interval that did not exceed ??? 2 m.y. Rb/Sr model ages range from 152 to 140 Ma, assuming that the burial diagenetic regime was buffered with respect to strontium by the associated marine Permian evaporites. Authigenic K-feldspar records two distinct events of hot brine flow, most likely triggered by tectonic movements (detachment) and by an increase in the subsurface temperature in response to thrust loading.




    Full Text Available Omanaster imbricatus is a new genu s and species of Sakmarian (Early Permian asteroids collected from the basal Pachycyrtella Bed of the Saiwan Formation of Oman, Arabian Peninsula; the family Omanasteridae is recognized. Late Paleozoic and especially Permian asteroids are rare and O. imbricatus differs significantly from those previously described, thereby providing an important addition to known late Paleozoic diversity. Unfortunately the single available specimen is incomplete with remaining ossicles both leached and partially fused, and available data are limited. Adambulacral form and arrangement of O. imbricatus are both suggestive of corresponding expressions of certain earlier Paleozoic species and unlike those of the crown-group, suggesting an enduring Paleozoic lineage but one not phylogenetically a part of the Mesozoic diversification. The Pachycyrtella Bed has been interpreted as recording a succession of pioneer palaeocommunities colonizing a turbulent, shallow-water settingaffected by oscillatory flows. The apparently flattened appearance of O. imbricatus is suggestive of appearances of certain Cretaceous and extant species recovered from similar environments thus suggesting both homoplasy and the versatility of asteroid evolution across extended spans of geologic time.

  9. The Dynamic Response of Marine Life to Extreme Temperature and Low Oxygen Events Following the End-Permian Mass Extinction.

    Pietsch, C.; Bottjer, D. J.


    The end-Permian mass extinction was the most devastating taxonomic and ecological crisis in the history of life on Earth. The recovery lasted 5 My making it the longest in geologic history, although the cause of the delay is still heavily debated. We find that additional environmental changes during the recovery interval reset the attempts that marine communities made toward ecological complexity, resulting in the overall appearance of a stagnant recovery. The extinction mechanisms during the end-Permian include extreme temperature change and low oxygen environments resulting from the volcanic emission of carbon dioxide and other toxic gasses to the atmosphere. The biotic response to ancient environmental change is a direct analog for the ecological impacts of modern anthropogenic climate change. We applied an ecological recovery rubric to benthic, sea floor dwelling, communities throughout the Early Triassic recovery in two major ocean basins. Newly collected bulk fossil data from the Moenkopi and Thaynes Formations from the Southwest US and the Werfen Formation in Italy were analyzed along with literature data. In Italy, directly following the extinction, low oxygen environments prevented an ecological rebound. Once low oxygen conditions receded, 600 kyr after the extinction, taxonomic diversity, fossil body size, and trace fossil complexity rebounded. A little more than 1 My into the Early Triassic, an extreme temperature event resulted in a reset of community complexity in both Italy and the Southwest US. The body size of gastropods and the repopulation of echinoderms were significantly inhibited as was trace fossil complexity. Low oxygen conditions that developed in the last ~2My of the Early Triassic limited diversity and body size in the Southwest United States. The stagnant recovery is re-interpreted as dynamic resets and rapid rebounds driven by environmental perturbations throughout the Early Triassic.

  10. The Origin and Early Radiation of Archosauriforms: Integrating the Skeletal and Footprint Record.

    Bernardi, Massimo; Klein, Hendrik; Petti, Fabio Massimo; Ezcurra, Martín D


    We present a holistic approach to the study of early archosauriform evolution by integrating body and track records. The ichnological record supports a Late Permian-Early Triassic radiation of archosauriforms not well documented by skeletal material, and new footprints from the Upper Permian of the southern Alps (Italy) provide evidence for a diversity not yet sampled by body fossils. The integrative study of body fossil and footprint data supports the hypothesis that archosauriforms had already undergone substantial taxonomic diversification by the Late Permian and that by the Early Triassic archosauromorphs attained a broad geographical distribution over most parts of Pangea. Analysis of body size, as deduced from track size, suggests that archosauriform average body size did not change significantly from the Late Permian to the Early Triassic. A survey of facies yielding both skeletal and track record indicate an ecological preference for inland fluvial (lacustrine) environments for early archosauromorphs. Finally, although more data is needed, Late Permian chirotheriid imprints suggest a shift from sprawling to erect posture in archosauriforms before the end-Permian mass extinction event. We highlight the importance of approaching palaeobiological questions by using all available sources of data, specifically through integrating the body and track fossil record.

  11. The Origin and Early Radiation of Archosauriforms: Integrating the Skeletal and Footprint Record.

    Massimo Bernardi

    Full Text Available We present a holistic approach to the study of early archosauriform evolution by integrating body and track records. The ichnological record supports a Late Permian-Early Triassic radiation of archosauriforms not well documented by skeletal material, and new footprints from the Upper Permian of the southern Alps (Italy provide evidence for a diversity not yet sampled by body fossils. The integrative study of body fossil and footprint data supports the hypothesis that archosauriforms had already undergone substantial taxonomic diversification by the Late Permian and that by the Early Triassic archosauromorphs attained a broad geographical distribution over most parts of Pangea. Analysis of body size, as deduced from track size, suggests that archosauriform average body size did not change significantly from the Late Permian to the Early Triassic. A survey of facies yielding both skeletal and track record indicate an ecological preference for inland fluvial (lacustrine environments for early archosauromorphs. Finally, although more data is needed, Late Permian chirotheriid imprints suggest a shift from sprawling to erect posture in archosauriforms before the end-Permian mass extinction event. We highlight the importance of approaching palaeobiological questions by using all available sources of data, specifically through integrating the body and track fossil record.

  12. Diagenetic overprint on negative δ13C excursions across the Permian/Triassic boundary: A case study from Meishan section, China

    Li, Rong


    The Permian-Triassic carbonate succession that formed during the biggest mass extinction event in geological history has long been studied to examine the biomass extinction patterns and mechanisms. Such studies have shown that the stable inorganic carbon isotope displays a worldwide significant negative shift in the Permian/Triassic transitional period, which has generally been attributed to a synchronistic shift in δ13C in the global carbon cycle. The assertion, however, is based on the assumption that the δ13C signals recorded in the carbonate succession are primary in origin. In this study, the diagenetic features of Beds 24 to 62 from Meishan Section, China, which is the Global Stratotype Section and Point (GSSP) of the Permian-Triassic boundary, are examined to determine the negative shift in δ13Ccarb across the Permian-Triassic mass extinction (PTME) is diagenetic in origin or not. Bed 24, the topmost unit of Changhsing Formation, is formed of skeletal packstones. Beds 25 to 62, belonging to Yinkeng Formation, are formed of either claystones, or mudstones and/or calcareous mudstones with dolomite being present. Petrographic data indicate that zoned dolomite crystals are widespread in early Triassic beds. Geochemical data including stable isotopic composition, minor and trace elements, and rare-earth elements indicate that the dolomite and coexisting calcite are formed in different depositional/diagenetic environment. The dramatically negative δ13C excursions of calcite in Beds 26 and 28 are related to meteoric diagenesis, while the negative δ13C excursions of calcite in dolomite-bearing beds are ascribed to enriched 12C resulted from dolomitization mediated by sulfate reducing bacterial (SRB) in burial process. The results show that the δ13C signals recorded in the global stratotype section that spans the PTB is not primary in origin. The synchronistic negative shift in δ13C signals across the PTME are partly contributed by diagenesis.

  13. The Cathaysia Flora and the Mixed Late Permian Cathaysian-Angaran Floras in East Asia


    The Cathaysia flora, one of the most prominent floras of the Carboniferous and Permian, was mainly distributed in East Asia and was characterized by numerous endemic elements. China is one of the most important localities of the Cathaysia flora and it is also the center of origin of this flora. This paper reviews and discusses the characteristics of the Cathaysia flora and the biogeographically mixed Permian Cathaysian-Angaran floras of East Asia. In addition, the formative mechanism of the mixed Permian floras is also discussed.

  14. Permian macro- and miofloral diversity, palynodating and palaeoclimate implications deduced from the coal-bearing sequences of Singrauli coalfield, Son-Mahanadi Basin, central India

    Singh, Kamal Jeet; Murthy, Srikanta; Saxena, Anju; Shabbar, Husain


    The coal-bearing sequences of Barakar and Raniganj formations exposed in Bina and Jhingurdah open-cast collieries, respectively, are analysed for their macro- and miofloral content. The sediment successions primarily comprise of sandstones, shales, claystones and coal seams. In addition to the diverse glossopterid assemblage, four palynoassemblage zones, namely Zones I and II in Bina Colliery and Zones III and IV in Jhingurdah Colliery, have also been recorded in the present study. The megafossil assemblage from the Barakar strata of Bina Colliery comprises of three genera, namely Gangamopteris, Glossopteris and cf. Noeggerathiopsis. Palynoassemblage-I is characterised by the dominance of non-striate bisaccate pollen genus Scheuringipollenites and subdominance of striate bisaccate Faunipollenites and infers these strata to be of Early Permian (Artinskian) age (Lower Barakar Formation). The palynoassemblage has also yielded a large number of naked fossil spore tetrads, which is the first record of spore tetrads from any Artinskian strata in the world and has a significant bearing on the climatic conditions. The palynoassemblage-II is characterised with the dominance of Faunipollenites over Scheuringipollenites and is indicative of Kungurian age (Upper Barakar Formation). The megafossil assemblage from the Raniganj Formation of Jhingurdah Colliery comprises of five genera with 26 species representing four orders, viz., Equisetales, Cordaitales, Cycadales and Glossopteridales. The order Glossopteridales is highly diversified with 23 taxa and the genus Glossopteris, with 22 species, dominates the flora. The mioflora of this colliery is represented by two distinct palynoassemblages. The palynoassemblage-III is correlatable with the palynoflora of Early Permian (Artinskian) Lower Barakar Formation. The assemblage suggests the continuity of older biozones into the younger ones. The palynoassemblage-IV equates the beds with composition V: Striatopodocarpites

  15. Permian macro- and miofloral diversity, palynodating and palaeoclimate implications deduced from the coal-bearing sequences of Singrauli coalfield, Son–Mahanadi Basin, central India

    Kamal Jeet Singh; Srikanta Murthy; Anju Saxena; Husain Shabbar


    The coal-bearing sequences of Barakar and Raniganj formations exposed in Bina and Jhingurdah open-cast collieries, respectively, are analysed for their macro- and miofloral content. The sediment successions primarily comprise of sandstones, shales, claystones and coal seams. In addition to the diverseglossopterid assemblage, four palynoassemblage zones, namely Zones I and II in Bina Colliery and Zones III and IV in Jhingurdah Colliery, have also been recorded in the present study. The megafossil assemblage from the Barakar strata of Bina Colliery comprises of three genera, namely Gangamopteris, Glossopterisand cf. Noeggerathiopsis. Palynoassemblage-I is characterised by the dominance of non-striate bisaccate pollen genus Scheuringipollenites and subdominance of striate bisaccate Faunipollenites and infers these strata to be of Early Permian (Artinskian) age (Lower Barakar Formation). The palynoassemblagehas also yielded a large number of naked fossil spore tetrads, which is the first record of spore tetrads from any Artinskian strata in the world and has a significant bearing on the climatic conditions. The palynoassemblage-II is characterised with the dominance of Faunipollenites over Scheuringipollenites and is indicative of Kungurian age (Upper Barakar Formation). The megafossil assemblage from the Raniganj Formation of Jhingurdah Colliery comprises of five genera with 26 species representing four orders, viz., Equisetales, Cordaitales, Cycadales and Glossopteridales. The order Glossopteridales is highly diversifiedwith 23 taxa and the genus Glossopteris, with 22 species, dominates the flora. The mioflora of this colliery is represented by two distinct palynoassemblages. The palynoassemblage-III is correlatable with the palynoflora of Early Permian (Artinskian) Lower Barakar Formation. The assemblage suggests thecontinuity of older biozones into the younger ones. The palynoassemblage-IV equates the beds with composition V: Striatopodocarpites

  16. The Late Permian Ocean: What's the Big Stink?

    Meyer, K. M.; Ridgwell, A.; Kump, L. R.


    Since the ocean is (and has been) sulfate rich, the development of basinal to global anoxia is often associated with the buildup of hydrogen sulfide in anoxic waters. Bacterial sulfate reduction begins to dominate after oxygen and nitrate have been depleted, producing hydrogen sulfide. Hence, low atmospheric oxygen content, warm surface ocean temperatures, and high O2 demand reduce oceanic oxygen content and favor the establishment of euxinia. Biomarker evidence for photic zone sulfide and biogeochemical calculations suggest that the end-Permian mass extinction was one interval during which extreme anoxia may have led to H2S buildup. We hypothesize that H2S release to the atmosphere would be possible if the upward flux of sulfide from deep water in a largely euxinic ocean exceeded the oxygen flux into the surface ocean from wind mixing. In this scenario, destabilization of the chemocline (oxygen-sulfide interface) would cause sulfide poisoning in both the marine and terrestrial realms and contribute to the extinction. We used the end-Permian configuration of GENIE (, an energy-moisture-balance atmosphere model coupled to a 3-D, non-eddy-resolving, frictional geostrophic model to evaluate this hypothesis. This model includes marine biogeochemistry and capably simulates processes associated with the transition to oceanic anoxia. We performed a series of simulations designed to identify the conditions necessary for widespread euxinia and chemocline destabilization. We characterized the magnitude of hydrogen sulfide flux as a function of increasing oceanic phosphate content resulting from P release from sediments in anoxic environments. Significant ocean-atmosphere fluxes of H2S result from 6- to 10-fold increases in ocean phosphate at modern oxygen levels. These fluxes are focused in upwelling regions, although toxic H2S concentrations are also observed in the surface waters of nearshore equatorial regions. Our initial simulations support the

  17. Early period of early Permian tubular cyanobacteria reef in the eastern part of south Qinling Mountains


    A typical reef occurs in the eastern part of south Qinling Mountains in Sanlixia, Xunyang, Shaanxi Province. The reef, extending in north-south direction, was built up by various kinds of calcareous tubes including Girvanella, tubular cyanobacteria and problem microbials. The bioclastic bank, built up by algal clasts, foraminiferous, and fusulinids, formed the reef basement, on which dasyclads baffle bioclasts formed the original bafflestone at a small scale on the reef basement, and then a mass of bindstone was made up by all kinds of calcareous tubes and algae. On lateral part the reef is cryptalgal mudstone. The reef that occurred in the geosynclinal area is so familiar with one of the platforms in many respects, such as in reef structure, morphology and building mold. This indicates that building organisms have amazing adaptability and identity for building capacity in the two kinds of geotectonic background mentioned above.

  18. Early period of early Permian tubular cyanobacteria reef in the eastern part of south Qinling Mountains

    巩恩普; 范嘉松


    A typical reef occurs in the eastern part of south Qinling Mountains in Sanlixia, Xun-yang, Shaanxi Province. The reef, extending in north-south direction, was built up by various kinds of calcareous tubes including Girvanella, tubular cyanobacteria and problem microbials. The bio-clastic bank, built up by algal clasts, foraminiferous, and fusulinids, formed the reef basement, on which dasyclads baffle bioclasts formed the original bafflestone at a small scale on the reef basement, and then a mass of bindstone was made up by all kinds of calcareous tubes and algae. On lateral part the reef is cryptalgal mudstone. The reef that occurred in the geosynclinal area is so familiar with one of the platforms in many respects, such as in reef structure, morphology and building mold. This indicates that building organisms have amazing adaptability and identity for building capacity in the two kinds of geotectonic background mentioned above.

  19. Calculating eustatic amplitude of Middle Permian from reefs

    WU; Yasheng; (吴亚生); FAN; Jiasong; (范嘉松)


    Methods for calculating ancient eustatic change amplitudes according to reef fabric- facies are proposed, with a new method for determining sediment-loading subsidence. Compared with methods based on non-reefal deposits, these methods are more accurate in restoration of original sediment thickness, determination of sediment-loading subsidence, as well as restoration of ancient water depth. According to the reef in Guangxi, China, the amplitude of sea-level rise during Middle Permian (Neoschwagerina-Yabeina zone) is 249.5 m. According to the coeval reef of the Guadalupe Mountains, New Mexico and Texas, the coeval sea-level rise is 247 m. With these effective methods available, it is feasible to establish more accurate eustatic curve of Phanerozoic.

  20. Methanogenic burst in the end-Permian carbon cycle.

    Rothman, Daniel H; Fournier, Gregory P; French, Katherine L; Alm, Eric J; Boyle, Edward A; Cao, Changqun; Summons, Roger E


    The end-Permian extinction is associated with a mysterious disruption to Earth's carbon cycle. Here we identify causal mechanisms via three observations. First, we show that geochemical signals indicate superexponential growth of the marine inorganic carbon reservoir, coincident with the extinction and consistent with the expansion of a new microbial metabolic pathway. Second, we show that the efficient acetoclastic pathway in Methanosarcina emerged at a time statistically indistinguishable from the extinction. Finally, we show that nickel concentrations in South China sediments increased sharply at the extinction, probably as a consequence of massive Siberian volcanism, enabling a methanogenic expansion by removal of nickel limitation. Collectively, these results are consistent with the instigation of Earth's greatest mass extinction by a specific microbial innovation.

  1. Ocean redox change at the Permian-Triassic mass extinction

    Ruhl, Micha; Bjerrum, Christian J.; Canfield, Donald Eugene


    and marine (mass) ex¬tinction. The geographic and temporal extend and the intensity (ferruginous vs. euxinic) of anoxic con¬ditions is, however, strongly debated and not well constraint. This complicates understanding of close coupling between Earth’s physical, chemical and bi¬ological processes. We studied...... ocean redox change over the largest mass extinction event in Earth history, at the Permian-Tri¬assic boundary (at ~252 Ma). This event is marked by a major perturbation in the global exogenic carbon cycle (and associated major negative carbon isotope excursion (CIE)), likely initiated by carbon...... (anoxic but not euxinic) coinciding with the main extinction event. Molybdenum enrichments, often indicative for freely available sulfide in the water-column, only occur dur¬ing the second phase of euxinia. This pattern of ocean redox-change in Svalbard direct¬ly reflects similar trends in Greenland...

  2. Ocean redox change at the Permian-Triassic mass extinction

    Ruhl, Micha; Bjerrum, Christian J.; Canfield, Donald Eugene


    Earth’s history is marked by multiple events of ocean anoxia developing along continental margins and po¬tentially into the open ocean realm. These events of¬ten coincide with the emplacement of large igneous provinces (LIPs) on continents, major perturbations of global geochemical cycles...... ocean redox change over the largest mass extinction event in Earth history, at the Permian-Tri¬assic boundary (at ~252 Ma). This event is marked by a major perturbation in the global exogenic carbon cycle (and associated major negative carbon isotope excursion (CIE)), likely initiated by carbon...... and marine (mass) ex¬tinction. The geographic and temporal extend and the intensity (ferruginous vs. euxinic) of anoxic con¬ditions is, however, strongly debated and not well constraint. This complicates understanding of close coupling between Earth’s physical, chemical and bi¬ological processes. We studied...

  3. Preliminary Study on the PGE Geochemistry of the Permian Basalts in the Jinping Area

    Nian Hong; Shen Wei; Zhang Xueshu


    To disclose and study the magma genesis of the Permian basalts in the Jinping (金平)area, the PGE and Au contents of the Permian basalts in the Jinping area were assayed by use of the ICP-MS method. The PGE data show relatively strong differentiation of basaltic magma in comparison with the primitive mantle. The primitive mantle-normalized pattern is a left-dipping curve of the Pd-Pt enriched type. The Pd/Ir ratios of the PGE of the Permian basalts in the Jinping area are higher than those of the primitive mantle and the upper primitive mantle and the chondrite meteorite. The Pd/Ir ramaterial source. It is concluded that they are derived from the basaltic magma of the upper mantle source with a lower melting degree, which shares the similar magma material source with the Emeishan Permian basalts as a whole.

  4. Brachiopods from the Permian Triassic boundary beds at the Selong Xishan section, Xizang (Tibet), China

    Shen, Shuzhong; Yugan, Jin


    The brachiopods from the Waagenites Bed and the upper part of the Coral Bed above and below the Permian-Triassic sequence boundary at the Selong Xishan section of southern Xizang suggest a rich brachiopod fauna existing in Selong, and a rapid invasion of Tethyan brachiopods into the Himalayas at the very end of the Permian. A transgression beginning from the Waagenites Bed and a rapid flooding at the beginning of the Triassic is reflected by changes in composition of the Permian brachiopod assemblages in Selong. However, the dominant forms in the Permian-Triassic sequence boundary beds, including Krotovia, Spiriferella and Neospirifer, extend from the underlying beds of the Selong Group without significant evolutionary change. A brief discussion of each taxon and descriptions of two new species, Stenoscisma selongensis and Martinia attenuatelloides, are presented.

  5. Altered River Morphology in South Africa Related to the Permian-Triassic Extinction

    Peter D. Ward; David R. Montgomery; Roger Smith


    The Permian-Triassic transition in the Karoo Basin of South Africa was characterized by a rapid and apparently basin-wide change from meandering to braided river systems, as evidenced by preserved sedimentary facies...

  6. A contribuição dos parques infantis de Mário de Andrade para a construção de uma pedagogia da educação infantil The contribution of Mário de Andrade's 'parques infantis' to the construction of a pedagogy of early childhood education

    Ana Lúcia Goulart de Faria


    Full Text Available Os Parques Infantis do Departamento de Cultura da Prefeitura Municipal de São Paulo durante os seus três primeiros anos de funcionamento (1935-1938 na gestão de Mário de Andrade integravam projeto governamental de educação não-escolar para as crianças pequenas de família operária, por meio do qual lhes foi garantido o direito à infância. A pesquisa sobre o nosso folclore realizada pelo poeta-educador "300-350" e a diversidade étnica brasileira consistiam o principal fundamento dessa experiência educacional na qual as crianças reviviam as tradições populares e, através da arte e dos jogos tradicionais infantis, tinham a possibilidade de ser criança, de viver a especificidade dessa fase da vida. A planta arquitetônica e o regimento interno dos parques estão aqui destacados revelando entre outras características elementos para a construção de uma pedagogia da educação infantil.The "Parques infantis" of the Department of Culture of the city of São Paulo in their first three years (1935-1938 during Mário de Andrade term as director were part of a governmental non-schooling educational project for young children from working-class families, through which their right to childhood was assured. The research on our folklore performed by the poet-educator ("300-350" and the brazilian ethnical diversity made up the fundamentals of this educational experience in which children relived popular traditions, had the chance of being children through art and traditional child play, of experiencing the specifics of that age. The architectural plan and the rules and regulations of the "parques infantis" are pointed out here showing, among other characteristics, elements for the construction of a pedagogy of early childhood education.




    Full Text Available In the course of an investigation on the major mass extintion event near the P/T boundary, in the vicinity of Shahreza (South Isfahan in Iran, a stratigraphically significant new species of foraminifera (Dagmarita shahreaensis n. sp. has been discovered in the Wuchiapingian/Dzhulfian (Late Permian of central Iran (Hambast Formation. This new species is described and emplaced in the phylogeny of the globivalvulinid foraminifera, which evolved rapidly during the Middle/Late Permian

  8. Anoxia duirng the Late Permian Binary Mass Extinction and Dark Matter

    Abbas, Samar; Abbas, Afsar; Mohanty, Shukadev


    Recent evidence quite convincingly indicates that the Late Permian biotic crisis was in fact a binary extinction with a distinct end-Guadalupian extinction pulse preceding the major terminal end-Permian Tartarian event by 5 million years. In addition anoxia appears to be closely associated with each of these end-Paleozoic binary extinctions. Most leading models cannot explain both anoxia and the binary characteristic of this crisis. In this paper we show that the recently proposed volcanogeni...

  9. The Lopingian Series of China and End-Permian Mass Extinction

    JIN Yugan; SHEN Shuzhong; WANG Xiangdong; WANG Yue; CAO Changqun


    The end-Permian mass extinction is the greatest bioevent in the geological history, which wiped out nearly 95% of the marine species and 75% of the terrestrial species,followed by a biological winter for more than five million years.The cause and the process of the end-Permian mass extinction remain an unsolved mystery in earth science and one of the most difficult and interesting scientific problems.

  10. Evolution and extinction of Permian fusulinid fauna in the Khao Tham Yai Limestone in NE Thailand

    Hada, Shigeki; Khosithanont, Somboon; Goto, Hiroya; Fontaine, Henri; Salyapongse, Sirot


    The first detailed biostratigraphic investigation of a single limestone unit within the Khao Tham Yai Limestone shows it was deposited continuously in a shelf setting without any intercalation of clastic beds. It ranges from the Wordian (middle Middle Permian) up to the Wuchiapingian (lower Upper Permian). The limestone unit is divided in ascending order into three fusulinid zones, i.e. Colania, Lepidolina and Codonofusiella zones. The middle zone is characterized by an abundance of large-tested fusulinids characteristic of the Lepidolina Zone. Shell sizes of the fusulinid species in this zone display continuous rapid morphological change along a one-way evolutionary path from small, primitive species with simple structure to large, highly evolved species having a complicated wall structure. Fusulinid biostratigraphy in a single limestone unit elucidates evolution and extinction patterns of Permian fusulinids of the shallow-water Tethyan shelf area in the Indochina Block. Our study reveals that the boundary between the Guadalupian (Middle Permian) and Lopingian (Upper Permian) in the Khao Tham Yai Limestone is clearly defined as an abrupt change in the fusulinid assemblages from the elimination of large-tested Verbeekinids and Schwagerinids to the domination of small-shelled Schubertellids. The Schubertellids underwent slower evolutionary morphological change than earlier fusulinids and were decreasingly dominant through the Permian. A similar pattern of fusulinid evolution and extinction at the Guadalupian-Lopingian boundary occurs in shallow-water Tethyan shelf areas and mid-oceanic shallow-water environments in mid-Panthalassa. Eventually, even smaller fusulinids abruptly become extinct. Clastic deposits finally replace previous carbonate formations characterized by algae-foraminifera biota. It starts in the upper Middle Permian in the southern parts and spreads throughout the whole area in the lower Upper Permian in NE Thailand. These observations suggest

  11. Volcanismo calcoalcalino neopaleozoico en la Precordillera de La Rioja. Petrología y caracterización litoestratigráfica de la Formación Punta del Agua (Carbonífero Superior-Pérmico Inferior Late Palaeozoic calc-alkaline volcanism in the Precordillera of La Rioja: petrology and lithostratigraphical characterization of the Punta del Agua Formation (Late Carboniferous - Early Permian

    M. Remesal


    Pampeanas. The unit consists of interbedded lava flows, pyroclastic flows and clastic sedimentary deposits. The lava flows have been divided according to their composition and textural characteristics; a dacitic sill is also included. Pyroclastic flows consist of block- and ash-flow deposits and reoignimbrites. Between periods of volcanicity, conglomerates, lithic sandstones and scarce mudstones were deposited in lenticular bodies that reflect the irregular morphology of the volcanic area. Twelve samples of the volcanic rocks, obtained from both flanks of the Punta del Agua syncline were geochemically analysed. The variation in chemical composition ranges from basaltic andesites to rhyolites, but andesitic compositions dominate. The calc-alkaline character of these rocks and the abundance of K in the andesites, suggest that these magmas originated by subduction with crystal fractionation and contamination processes. The age of the Punta del Agua Formation was established on the basis of its stratigraphical relations and radiometric dating. Both suggest a Late Carboniferous age, although an Early Permian age for the uppermost part of the sequence cannot be disregarded.

  12. Magnetostratigraphic correlations of Permian-Triassic marine-to-terrestrial sections from China

    Glen, J.M.G.; Nomade, S.; Lyons, J.J.; Metcalfe, I.; Mundil, R.; Renne, P.R.


    We have studied three Permian-Triassic (PT) localities from China as part of a combined magnetostratigraphic, 40Ar/39Ar and U-Pb radioisotopic, and biostratigraphic study aimed at resolving the temporal relations between terrestrial and marine records across the Permo-Triassic boundary, as well as the rate of the biotic recovery in the Early Triassic. The studied sections from Shangsi (Sichuan Province), Langdai (Guihzou Province), and the Junggar basin (Xinjiang Province), span marine, paralic, and terrestrial PT environments, respectively. Each of these sections was logged in detail in order to place geochronologic, paleomagnetic, geochemical, conodont and palynologic samples within a common stratigraphic context. Here we present rock-magnetic, paleomagnetic and magnetostratigraphic results from the three localities. At Shangsi, northern Sichuan Province, we sampled three sections spanning Permo-Triassic marine carbonates. Magnetostratigraphic results from the three sections indicate that the composite section contains at least eight polarity chrons and that the PT boundary occurs within a normal polarity chron a short distance above the mass extinction level and a reversed-to-normal (R-N) polarity reversal. Furthermore, the onset of the Illawarra mixed interval lies below the sampled section indicating that the uppermost Permian Changhsingian and at least part of the Wuchiapingian stages postdate the end of the Kiaman Permo-Carboniferous Reversed Superchron. At Langdai, Guizhou Province, we studied magnetostratigraphy of PT paralic mudstone and carbonate sediments in two sections. The composite section spans an R-N polarity sequence. Section-mean directions pass a fold test at the 95% confidence level, and the section-mean poles are close to the mean PT pole for the South China block. Based on biostratigraphic constraints, the R-N transition recorded at Langdai is consistent with that at Shangsi and demonstrates that the PT boundary occurred within a normal

  13. Biogeochemical evidence for euxinic oceans and ecological disturbance presaging the end-Permian mass extinction event

    Cao, Changqun; Love, Gordon D.; Hays, Lindsay E.; Wang, Wei; Shen, Shuzhong; Summons, Roger E.


    The Permian-Triassic Boundary event at 252.2 Ma marks the largest extinction of marine fauna in the Phanerozoic and there is a wide consensus that the extinction coincided with an intense oceanic anoxic event. The stratotype of the Changhsingian Stage, precisely constrained by the PTB Global Stratotype Section and Point (GSSP) and the GSSP for the Wuchiapingian-Changhsingian Boundary, both at Meishan in southern China, is well-documented in respect to geochronology and the pattern of extinction. Here we report secular trends in bulk isotopic parameters and lipid biomarkers in a core spanning 214 m of stratigraphic section across the PTB and through the entire Changhsingian interval. Our analysis of these data, viewed in the context of relative sea level change and strontium isotopes, reveals distinct shifts in paleoenvironmental conditions and profound changes in plankton ecology well before and following the biological extinction event. Specifically, patterns of steroids and triterpenoids indicate a marine plankton community that was heavily dominated by bacteria during the late Wuchiapingian, middle Changhsingian and early Griesbachian stages. Secular trends in aromatic hydrocarbons diagnostic for anoxygenic green sulphur bacteria (Chlorobiaceae) identify periods when euxinic conditions extended into the photic zone during the entire Changhsingian stage. Here also, the δ15N of organic nitrogen progressively shifted from positive values around + 2 or + 3‰ to - 1‰ coincident with a sharp negative excursion in δ13C org and slightly postdating the sharp minimum in δ13C values of inorganic carbon that occurs at the top of Bed 24. These results, together the published chronology indicate that conditions unfavourable for aerobiosis existed in the marine photic zone at Meishan for 1.5 million years prior to the main phase of the biological extinction. The induction of marine euxinic conditions, worldwide, at the end of the Permian was likely a consequence of the

  14. Organic richness and gas generation potential of Permian Barren Measures from Raniganj field, West Bengal, India

    Annapurna Boruah; S Ganapathi


    The organic geochemistry of shales in terms of its organic richness, hydrocarbon source potential, thermal maturity, depositional environment, etc., are essential stipulations for shale gas resources assessment. In this study, a total of 32 core samples of Permian Barren Measures from four boreholes in Raniganj field of Damodar Basin were analysed to evaluate their gas generation potential using Rock–Eval pyrolysis techniques. Petrographic analysis brings out the lithofacies of Barren Measures as carbonaceous silty shale, iron rich claystone and sand-shale intercalation. The total organic content (TOC) of the shale units of Barren Measures ranges from 3.75 to 20.9 wt%, whereas hydrogen index (HI) ranges from 58.45 to 125.34 mg HC/g TOC. Present study suggests early to late maturated (0.6–1%) organic matters in Barren Measures with gas prone type III kerogen. The study analysed the effect of burial history on the preservation and maturation of organic matters. The organic richness, kerogen type, thermal maturity and petrographic properties of Barren Measures signify fair to excellent gas generation potential.

  15. Anoxia/high temperature double whammy during the Permian-Triassic marine crisis and its aftermath.

    Song, Haijun; Wignall, Paul B; Chu, Daoliang; Tong, Jinnan; Sun, Yadong; Song, Huyue; He, Weihong; Tian, Li


    The Permian-Triassic mass extinction was the most severe biotic crisis in the past 500 million years. Many hypotheses have been proposed to explain the crisis, but few account for the spectrum of extinction selectivity and subsequent recovery. Here we show that selective losses are best accounted for by a combination of lethally warm, shallow waters and anoxic deep waters that acted to severely restrict the habitable area to a narrow mid-water refuge zone. The relative tolerance of groups to this double whammy provides the first clear explanation for the selective extinction losses during this double-pulsed crisis and also the fitful recovery. Thus, high temperature intolerant shallow-water dwellers, such as corals, large foraminifers and radiolarians were eliminated first whilst high temperature tolerant ostracods thrived except in anoxic deeper-waters. In contrast, hypoxia tolerant but temperature intolerant small foraminifers were driven from shallow-waters but thrived on dysoxic slopes margins. Only those mollusc groups, which are tolerant of both hypoxia and high temperatures, were able to thrive in the immediate aftermath of the extinction. Limited Early Triassic benthic recovery was restricted to mid-water depths and coincided with intervals of cooling and deepening of water column anoxia that expanded the habitable mid-water refuge zone.

  16. Subsequent biotic crises delayed marine recovery following the late Permian mass extinction event in northern Italy.

    Foster, William J; Danise, Silvia; Price, Gregory D; Twitchett, Richard J


    The late Permian mass extinction event was the largest biotic crisis of the Phanerozoic and has the longest recovery interval of any extinction event. It has been hypothesised that subsequent carbon isotope perturbations during the Early Triassic are associated with biotic crises that impeded benthic recovery. We test this hypothesis by undertaking the highest-resolution study yet made of the rock and fossil records of the entire Werfen Formation, Italy. Here, we show that elevated extinction rates were recorded not only in the Dienerian, as previously recognised, but also around the Smithian/Spathian boundary. Functional richness increases across the Smithian/Spathian boundary associated with elevated origination rates in the lower Spathian. The taxonomic and functional composition of benthic faunas only recorded two significant changes: (1) reduced heterogeneity in the Dienerian, and (2) and a faunal turnover across the Smithian/Spathian boundary. The elevated extinctions and compositional shifts in the Dienerian and across the Smithian/Spathian boundary are associated with a negative and positive isotope excursion, respectively, which supports the hypothesis that subsequent biotic crises are associated with carbon isotope shifts. The Spathian fauna represents a more advanced ecological state, not recognised in the previous members of the Werfen Formation, with increased habitat differentiation, a shift in the dominant modes of life, appearance of stenohaline taxa and the occupation of the erect and infaunal tiers. In addition to subsequent biotic crises delaying the recovery, therefore, persistent environmental stress limited the ecological complexity of benthic recovery prior to the Spathian.

  17. Lower and middle Guadalupian shelf carbonates, eastern margin of Central Basin platform, Permian basin, west Texas

    Ward, R.F.; Chalcraft, R.G.


    Lower and middle Guadalupian shelf carbonates serve as the reservoir for a nearly continuous band of oil fields extending 100 mi along the eastern margin of the Central Basin platform of west Texas. Approximately 5 billion bbl of oil have been produced from stratigraphic-structural traps within the Upper Permian (Gaudalupian Series) dolomites of the San Andrea and Grayburg Formations in Upton, Crane, Ector, Pecos, and Andrews Counties, Texas. The San Andrea and Grayburg Formations are cyclical shallowing-upward carbonate sequences of open shelf through sabkha facies whose depositional strike parallels the eastern margin of the Central Basin platform. Porosity and permeability of reservoir rock are governed by diagenetic processes such as dolomitization, anhydrite porosity occlusion, leaching, silicification, and authigenic clay formation. Self sediments are primarily burrowed wackestones and packstones that locally contain pelletal, skeletal, and ooid grainstones. Typical subtidal shelf sediments are capped by algal-laminated dolomite, nodular anhydritic dolomite, and bedded anhydrite. The fauna is normally sparse and dominated by foraminifera and algae. Less common faunal components include pelecypods, crinoids, sponges, Bryozoa, brachiopods, gastropods, and coral that are associated with the development of small scattered patch reefs. Lowering the sea level during the early Guadalpian initiated basinward progradation of San Andres carbonate facies with hydrocarbon reservoirs best developed in shallow self fusulinid wackestones to packstone and oolitic grainstone. Reservoir dolomites of the Grayburg formation are present east of San Andres fields with optimal reservoir properties occurring near the San Andreas outer shelf margin.




    Full Text Available The uppermost Carboniferous-Lower Permian Dorud Group of the Gaduk section in Central Alborz (Iran is more than 44 m-thick; it includes thick succession of conglomerates, quartzarenites, calcareous sandstones, oncolitic fusulinid limestones, sandy limestones, sandstones and shales. The Toyeh, Emarat and Shah Zeid formations of this Group were dated from the Gzhelian to the Sakmarian. A review of the uppermost Gzhelian, Asselian and lower Sakmarian smaller foraminifers of the Emarat Formation of the Gaduk section is here presented. Three foraminiferal biozones are defined: the Nodosinelloides shikhanica-Geinitzina primitiva Zone is latest Gzhelian in age; the Nodosinelloides spp.-Geinitzina spp.-Pseudoacutella partoazari Zone is Asselian; the Rectogordius iranicus gadukensis-Endothyra cf. bamberi Zone is early Sakmarian in age. A new subspecies Rectogordius iranicus gadukensis n. subsp. is described, and the genera Pseudovidalina and Grovesella are discussed. The studied assemblages are correlated with those from the Carnic Alps (Austria-Italy, East European Platform of Russia, the Urals (Russia, Darvaz (Uzbekistan, the northern and central Pamirs (Tajikistan, Central Iran, northern Afghanistan and other classical regions of the Tethyan realm.

  19. Abrupt Changes at the Permian/Triassic Boundary: Tempo of Events from High-Resolution Cyclostratigraphy

    Rampino, M. R.; Prokoph, A.; Adler, A. C.


    the nearby Reppwand outcrop section, the same faunal changes occurs over only 0.8 m or about 8,000 years, close to the limit of time-resolution induced by bioturbation and reworking in these sediments. The sharp negative global carbon-isotope shift took place within less than or equal to 40,000 yr, and the isotope excursions persisted for approximately 480,000 yr into the Early Triassic. The results indicate that the severe marine faunal event that marks the P/Tr boundary was very sudden, perhaps less than the resolution window in the GK-1 core, and suggest a catastrophic cause. The wavelet-analysis approach to high-resolution cyclostratigraphy can be applied to other P/Tr boundary sections, and when combined with precise absolute dating and magnetostratigraphic methods promises a significant increase in resolution in determining the correlation and tempo of the end-Permian extinctions and related events worldwide.

  20. Two pulses of oceanic environmental disturbance during the Permian-Triassic boundary crisis

    Shen, Jun; Feng, Qinglai; Algeo, Thomas J.; Li, Chao; Planavsky, Noah J.; Zhou, Lian; Zhang, Mingliang


    Pyrite morphology, iron speciation, and pyrite sulfur isotope data from the Xiakou section (Hubei Province, South China) were integrated to explore oceanic environmental variations through the Permian-Triassic transition and their possible relations to the largest mass extinction in Earth history. High ratios of highly-reactive iron to total iron (FeHR/FeT > 0.6) and pyrite iron to highly-reactive iron (FePy/FeHR > 0.7) together with a high abundance of small (mean diameter Permian (pre-extinction interval). High ratios of FeHR/FeT (mostly >0.6) along with lower values of FePy/FeHR (Permian to earliest Triassic (post-extinction interval). The largest fluctuations of these redox proxies are observed in the ∼1.0 m of strata directly overlying the first (latest Permian) extinction horizon, indicating unsettled marine environmental conditions marked by frequent perturbations during the ∼60-kyr interval immediately following the mass extinction. The two largest redox events at Xiakou coincided with deposition of volcanic ash layers that have been correlated with Beds 25 and 28 of the Meishan D section, each of which was associated with an extinction pulse (the first and second extinction horizons of latest Permian and earliest Triassic age, respectively). Thus, our observations document two pulses of oceanic environmental change during the Permian-Triassic transition, each associated with a faunal crisis and possibly triggered by contemporaneous volcanic activity.

  1. O ensino secundário no império e na primeira república no Brasil: entre as ciências e as humanidades. O caso do Liceu de Humanidades de Campos/RJ (1880-1930 - The secondary teaching in the last decade of the brazilian monarchy and in the early republica

    Silvia Alicia Martínez, Maria Amelia de Almeida Pinto Boynard


    Full Text Available Resumo Vinculado a uma pesquisa maior que estuda a trajetória do Liceu de Humanidades de  Campos  (LHC,  instituição   pública  de  ensino secundário do Estado  do Rio de  Janeiro, o trabalho se centra  no período 1880-1930.  Três aspectos principais articulam o texto: os regulamentos e as normas expedidos pelos órgãos da administração, que  configuram,  no  tempo,  a  estrutura   e  o  funcionamento  da instituição;  as  práticas,  que   operacionalizam  -ou  não-  o  que  a legislação sugere; e a materialidade dessas práticas, que configura os suportes  de circulação do  ideário pedagógico da época. As fontes indicam que até 1930  o LHC preocupou-se com o desenvolvimento das Ciências Físicas e Naturais,  sem,  contudo, abandonar o ensino das humanidades, que se tornou a marca da cultura da instituição e da cidade. Palavras-chave: ensino secundário; cultura material escolar; práticas; organização curricular.   THE SECONDARY TEACHING  IN THE LAST DECADE OF THE BRAZILIAN MONARCHY AND IN THE EARLY REPUBLICAN GOVERNMENT:  BETWEEN  SCIENCES AND HUMANITIES, THE LYCEUM OF HUMANITIES OF CAMPOS/RJ (1880-1930 Abstract The  work  is linked to  a major research focusing the  journey of a public institution  for secondary teaching  in  the  State  of Rio  de Janeiro, the  Lyceum of  Humanities  of Campos (LHC,  from  its origins (1880 until 1930.  Three aspects are articulated in the text: the   regulations   and   ordinances   issued   by   the   administrative authorities,   which  shape  the  structure   and  functioning  of  the institution;  the  practices that  operate what  the  rules  and  assizes suggest; the  materialization  of these practises, shaping the  classic pedagogical idealism of the era. The sources indicate that up to the decade of 1930  the LHC had as its main objective the development of  Physics and  Natural

  2. Paleomagnetism of the Late Proterozoic Sierras Bayas Group and the Ediacaran-Cambrian Apparent Polar Wander Path of the Rio de la Plata Craton

    Rapalini, A. E.; Trindade, R. I.; Poiré, D. G.; Vieira, L.


    The apparent polar wander path (APWP) of the Rio de la Plata craton (RP) for the Late Proterozoic and Early Paleozoic is of significance to reconstruct the tectonic processes that led to the amalgamation of Gondwana. In order to define the Ediacaran-Cambrian APWP for RP, a systematic paleomagnetic study was carried out in the Sierras Bayas Group exposed in the province of Buenos Aires, Argentina. 328 samples from 44 sites were collected in the Cryogenian to Ediacaran Villa Mónica Fm., the Ediacaran Cerro Largo, Loma Negra and Las Aguilas Fms. and the Cambrian (?) Cerro Negro Fm. Sampling included mainly limestone and dolomite, red claystone and marls. Although most sites presented secondary magnetizations of likely recent and Permian (?) age, a few sites from the Cerro Largo and uppermost Villa Mónica Fms. carry a pre-tectonic magnetization that yielded a preliminary paleomagnetic pole at 34.6°S, 211.3° (6.4°/10-9°; n: 18 samples), while a pre-tectonic remanence from the Cerro Negro Fm. yielded a paleomagnetic pole at 10.9°N, 323.5°E (9.7°/13.3°, n:17 samples). These two preliminary paleomagnetic poles for the Sierras Bayas Group together with previously published ones permit to define with confidence the APWP of RP from ca. 600 to ca. 500 Ma.

  3. Paleomagnetism and dating of a thick lava pile in the Permian Bakaly formation of eastern Kazakhstan: Regularities and singularities of the paleomagnetic record in thick lava series

    Bazhenov, Mikhail L.; Van der Voo, Rob; Menzo, Zachary; Dominguez, Ada R.; Meert, Joseph G.; Levashova, Natalia M.


    Paleomagnetic results on thick lava series are among the most important sources of information on the characteristics of ancient geomagnetic fields. Most paleo-secular variation data from lavas (PSVL) are of late Cenozoic age. There are far fewer results from lavas older than 5 Ma. The Central Asia Orogenic Belt that occupies several million square kilometers in Asia is probably the world's largest area of Paleozoic volcanism and is thus an attractive target for PSVL studies. We studied a ca. 1700 m thick lava pile in eastern Kazakhstan of Early Permian age. Magmatic zircons, successfully separated from an acid flow in this predominantly basaltic sequence, yielded an Early Permian age of 286.3 ± 3.5 Ma. Oriented samples were collected from 125 flows, resulting in 88 acceptable quality flow-means (n ⩾ 4 samples, radius of confidence circle α95 ⩽ 15°) of the high-temperature magnetization component. The uniformly reversed component is pre-tilting and arguably of a primary origin. The overall mean direction has a declination = 242.0° and an inclination = -56.2° (k = 71.5, α95 = 1.8°; N = 88 sites; pole at 44.1°N, 160.6°E, A95 = 2.2°). Our pole agrees well with the Early Permian reference data for Baltica, in accord with the radiometric age of the lava pile and geological views on evolution of the western part of the Central Asia Orogenic Belt. The new Early Permian result indicates a comparatively low level of secular variation especially when compared to PSVL data from intervals with frequent reversals. Still, the overall scatter of dispersion estimates that are used as proxies for SV magnitudes, elongation values and elongation orientations for PSVL data is high and cannot be fitted into any particular field model with fixed parameters. Both observed values and numerical simulations indicate that the main cause for the scatter of form parameters (elongation values and elongation orientations) is the too small size of collections. Dispersion estimates

  4. The Pattern and Evolution of the Permian Palaeobiogeography and Tectonic Palaeogeography in Jilin and Heilongjiang Orogenic Belt

    Peng Xiangdong; Li Xiaomin; Liu Pengju


    The orogenic belt locates between the North China plate and Siberia plate. The Permian palaeobiogeography and tectonic palaeogeography changed quickly and clearly. The line from Changchun to Yanji is an important palaeobiogeographic provincing line, which may be the collission suture belt of the North China plate and north middle massifs. The orogenic belt has been divided into 2 regions: the North middle massif region and the North Margin of North China plate, the pattern and evolution of Permian palaeobiogeography in the present area were discussed and the Permian biota mixture and its significants were analysed. Then, Based on the above, the Permian tectonic palaeogeography of the orogenic belt is reconstructed.

  5. Conodont body size and diversity trends after the end-Permian extinction: implications for the recovery of pelagic vs. benthic marine ecosystems

    Schaal, E. K.; Morgan, D. J.; Clapham, M.; Rego, B. L.; Wang, S. C.; Payne, J.


    Many marine clades decreased in size across the end-Permian extinction horizon, but the pattern and timing of subsequent size increase during recovery is poorly constrained. The tempo of recovery from the end-Permian extinction is key to understanding the role of evolutionary and environmental constraints in the recovery of ecosystems. However, not all marine organisms share the same recovery pattern. In this study, we document Late Permian to Late Triassic trends in size distribution of conodonts, pelagic chordates that exhibit much more rapid diversification in the Early Triassic compared to benthic clades. Because tooth size correlates with body weight in chordates over many orders of magnitude and is commonly used to predict body size in fossils, conodont elements can serve as a proxy for the size of the conodont animal. Our dataset includes both specimens from high-resolution samples through an exceptionally exposed carbonate platform in south China and size measurements from the published literature for conodonts and seven other marine clades. In platform slope sections of south China, we observe a size decrease across the P/Tr boundary, but in platform interior sections, large conodonts are prolific in the earliest Triassic. Comparison with global data shows that while there is decrease in median conodont size across the P/Tr boundary, there is little change in maximum conodont size, and pre-extinction size distributions return by the Smithian. While benthic clades show Early Triassic size reduction and slow recovery, pelagic clades such as conodonts and ammonoids show large size and rapid diversification after the extinction event. This decoupling of recovery between benthic and pelagic marine ecosystems could reflect a depth gradient in environmental parameters such as oxygen availability or intrinsic differences in the evolutionary dynamics of these clades.

  6. Morphology and histology of dorsal spines of the xenacanthid shark Orthacanthus platypternus from the Lower Permian of Texas, USA: palaeobiological and palaeoenvironmental implications

    Kimberly G. Beck


    Full Text Available Detailed studies on Carboniferous species of the xenacanth Orthacanthus have shown that the xenacanth dorsal fin spine can be used for skeletochronological analyses and provides valuable information about development, growth and environmental life conditions of those extinct sharks. We report here for the first time the histology and skeletochronology of Permian specimens, dorsal spines of Orthacanthus platypternus from the Craddock Bone Bed (lower Clear Fork Formation; Early Permian, Leonardian age of northern Baylor County (north-central Texas, USA. Twelve dorsal spines of O. platypternus preserve a highly vascularized wall mainly composed of centrifugally growing dentine in a succession of dentine layers, probably deposited with an annual periodicity. As expected, spines of individuals with 1–2 dentine layers, presumably juveniles, present the smallest sizes. However, spines of individuals showing at least 3–4 dentine layers and interpreted to be subadults/young adults, are distributed in two spine-size clusters corresponding to females (probably the largest spines and males, in agreement with the hypothesis of sexual size dimorphism proposed in a previous biometric analysis. Our comparative study of O. platypternus and the Stephanian species O. meridionalis further suggests that spine denticulation can be useful for distinguishing between species of Orthacanthus and sexually dimorphic forms (juvenile to adults in each species. Total body length estimations of O. platypternus from the Craddock Bone Bed point to relatively large juveniles and small subadults/young adults (less than 2 m in total length, living as opportunistic predators in the pond-channel coastal plain environments represented by the bone bed deposits. The com-parative analyses of the ontogenetic stages of the recorded specimens of O. platypternus and their distribution along different facies and localities indicate that this species was euryhaline, diadromous with a

  7. A sixteenth century Portuguese report concerning an early Javanese world map Relatório português do século XVI descreve mapa-múndi antigo de origem javanesa

    Benjamin B. Olshin


    Full Text Available In the sixteenth century, there were numerous voyages of discovery in the oceans of the world, expanding European understanding and influence. These explorations, and in particular the knowledge they created, have tended to overshadow other ventures into the unknown. Not just the Europeans, but other cultures as well, navigated the seas and accumulated geographical information, putting together their own ideas about the distribution of the lands and seas around the globe. Arab and Asian seafarers plied the oceans in trade and exploration, and created maps and geographical texts. These maps and texts, however, are not as numerous as a scholar might wish. The Asian maps, particularly early ones, are few in number, and these are virtually all Chinese and Korean creations, world maps based on traditional religious concepts of a circular landmass. There are few maps based on actual navigation. But an early sixteenth-century Portuguese document gives a brief description of a Javanese map, which apparently showed much of the world. It also depicted the routes of the navigations of the Chinese and other Pacific peoples. The map itself has been lost, but we read of it in a letter from the Portuguese admiral Afonso de Albuquerque to his king, Dom Manuel, dated 1 April 1512.No século XVI, várias viagens de descobrimento expandiram a influência e o reconhecimento europeus. Essas expedições marítimas, e em particular o conhecimento que geraram, ofuscaram outras aventuras pelo desconhecido. Outros povos navegaram pelos oceanos, acumularam informações e organizaram suas próprias idéias sobre a distribuição de terras e mares. Navegadores árabes e asiáticos criaram mapas, cartas náuticas e textos geográficos expondo suas descobertas. Esses mapas, no entanto, não são tão numerosos. Os asiáticos, particularmente os mais antigos, são poucos e quase todos originários da China e Coréia, e baseavam-se em conceitos religiosos tradicionais e na id

  8. Sandstone provenance and U-Pb ages of detrital zircons from Permian-Triassic forearc sediments within the Sukhothai Arc, northern Thailand: Record of volcanic-arc evolution in response to Paleo-Tethys subduction

    Hara, Hidetoshi; Kunii, Miyuki; Miyake, Yoshihiro; Hisada, Ken-ichiro; Kamata, Yoshihito; Ueno, Katsumi; Kon, Yoshiaki; Kurihara, Toshiyuki; Ueda, Hayato; Assavapatchara, San; Treerotchananon, Anuwat; Charoentitirat, Thasinee; Charusiri, Punya


    Provenance analysis and U-Pb dating of detrital zircons in Permian-Triassic forearc sediments from the Sukhothai Arc in northern Thailand clarify the evolution of a missing arc system associated with Paleo-Tethys subduction. The turbidite-dominant formations within the forearc sediments include the Permian Ngao Group (Kiu Lom, Pha Huat, and Huai Thak formations), the Early to earliest Late Triassic Lampang Group (Phra That and Hong Hoi formations), and the Late Triassic Song Group (Pha Daeng and Wang Chin formations). The sandstones are quartzose in the Pha Huat, Huai Thak, and Wang Chin formations, and lithic wacke in the Kiu Lom, Phra That, Hong Hoi and Pha Daeng formations. The quartzose sandstones contain abundant quartz, felsic volcanic and plutonic fragments, whereas the lithic sandstones contain mainly basaltic to felsic volcanic fragments. The youngest single-grain (YSG) zircon U-Pb age generally approximates the depositional age in the study area, but in the case of the limestone-dominant Pha Huat Formation the YSG age is clearly older. On the other hand, the youngest cluster U-Pb age (YC1σ) represents the peak of igneous activity in the source area. Geological evidence, geochemical signatures, and the YC1σ ages of the sandstones have allowed us to reconstruct the Sukhothai arc evolution. The initial Sukhothai Arc (Late Carboniferous-Early Permian) developed as a continental island arc. Subsequently, there was general magmatic quiescence with minor I-type granitic activity during the Middle to early Late Permian. In the latest Permian to early Late Triassic, the Sukhothai Arc developed in tandem with Early to Middle Triassic I-type granitic activity, Middle to Late Triassic volcanism, evolution of an accretionary complex, and an abundant supply of sediments from the volcanic rocks to the trench through a forearc basin. Subsequently, the Sukhothai Arc became quiescent as the Paleo-Tethys closed after the Late Triassic. In addition, parts of sediments of

  9. Late Permian topography at the southern margin of the Northern Permian Basin: Paleogeography inferred from 3D seismic analysis

    Clausen, Ole R.; Andresen, Katrine J.; Rasmussen, Jens A.


    The Top Pre Zechstein (TPZ) surface in the North Sea Basin is often mapped because it reveals the total basement tectonics in the area. In areas where Zechstein salt is present halokinetic processes, differential subsidence, and Mesozoic faulting however significantly alter the TPZ surface. The study area is located at the southern margin of the Northern Permian Basin in the eastern North Sea at the northern flank of the Ringkøbing-Fyn High. This area occurs approximately at the pinch-out line of the late Permian Zechstein salt and constitutes an excellent theater illustrating a range of salt-related problems. The TPZ surface is characterized by an overall NNW-ward dip defining the northern flank of the RFH and is transected by a set of NNW-SSE striking faults, and a E-W striking set of minor faults. Salt structures in the northern part of the study area introduce velocity pull-up (artefacts) at the TPZ surface and furthermore cause intense faulting of the Mesozoic and Cenozoic cover sediments. Pronounced isolated topographic highs similar to hills can be observed in the southern part of the study area where no to very little Zechstein evaporites are present. In the central part where Zechstein evaporites are present, small topographic highs similar to ridges can be observed at the footwall crest of minor faults. The Zechstein evaporites generally onlap towards the south in the study area but in the transitional zone around the hills, onlap from all directions onto the hills is observed. This suggests that the hills reflect paleo-topography developed during sub-aerial exposure before and perhaps during the deposition of the Zechstein sediments. The internal reflections within the hills show that they are composed of southward dipping sediments and very evident erosional truncations can be observed. The hills are aligned parallel to the major E-W striking basement fault, but are not directly associated to faults offsetting the TPZ surface. However, the alignment

  10. Simulated warm polar currents during the middle Permian

    Winguth, A.M.E.; Kutzbach, J.E. [Wisconsin Univ., Madison, WI (USA). Center for Climatic Research; Heinze, C.; Maier-Reimer, E.; Mikolajewicz, U. [Max-Planck-Institut fuer Meteorologie, Hamburg (Germany); Rowley, D.; Rees, A.; Ziegler, A.M. [Chicago Univ., IL (United States). Dept. of Geophysical Sciences


    During Permian Stage 6 (Wordian, Kazanian) the Pangaean supercontinent was surrounded by a superocean - Panthalassa. An ocean general circulation model has been coupled to an atmospheric energy balance model to simulate the sensitivity of the Wordian climate ({proportional_to}265 million years ago) to changes in greenhouse gas concentrations, high latitude geography, and Earth orbital configurations. The model shows a high sensitivity of the ocean circulation to changes in the greenhouse gas forcing, ranging from a forceful southern circulation at low CO{sub 2} concentration (present level) to a more symmetric circulation cell with deep water formation in both hemispheres at high CO{sub 2} concentration (8 x present level). The simulated climate with 4 x present level CO{sub 2} concentration agrees generally well with climate-sensitive sediments and phytogeographic patterns. In this experiment, the model simulates strong subtropical gyres with similarities to the modern South Pacific circulation and moderate surface temperatures on the southern continent Gondwana, resulting from a strong poleward heat transport in the ocean. An even more moderate climate can be generated if high latitude land is removed so that ocean currents can penetrate into the polar regions or if orbital configurations favor high summer insolation over Gondwana. (orig.)

  11. Radon in the Creswell Crags Permian limestone caves

    Gillmore, G.K. E-mail:; Phillips, P.S.; Denman, A.R.; Gilbertson, D.D


    An investigation of radon levels in the caves of Creswell Crags, Derbyshire, an important Site of Special Scientific Interest (SSSI) shows that the Lower Magnesian Limestone (Permian) caves have moderate to raised radon gas levels (27-7800 Bq m{sup -3}) which generally increase with increasing distance into the caves from the entrance regions. This feature is partly explained in terms of cave ventilation and topography. While these levels are generally below the Action Level in the workplace (400 Bq m{sup -3} in the UK), they are above the Action Level for domestic properties (200 Bq m{sup -3}). Creswell Crags has approximately 40,000 visitors per year and therefore a quantification of effective dose is important for both visitors and guides to the Robin Hood show cave. Due to short exposure times the dose received by visitors is low (0.0016 mSv/visit) and regulations concerning exposure are not contravened. Similarly, the dose received by guides is fairly low (0.4 mSv/annum) due in part to current working practice. However, the risk to researchers entering the more inaccessible areas of the cave system is higher (0.06 mSv/visit). This survey also investigated the effect of seasonal variations on recorded radon concentration. From this work summer to winter ratios of between 1.1 and 9.51 were determined for different locations within the largest cave system.

  12. Time-calibrated Milankovitch cycles for the late Permian.

    Wu, Huaichun; Zhang, Shihong; Hinnov, Linda A; Jiang, Ganqing; Feng, Qinglai; Li, Haiyan; Yang, Tianshui


    An important innovation in the geosciences is the astronomical time scale. The astronomical time scale is based on the Milankovitch-forced stratigraphy that has been calibrated to astronomical models of paleoclimate forcing; it is defined for much of Cenozoic-Mesozoic. For the Palaeozoic era, however, astronomical forcing has not been widely explored because of lack of high-precision geochronology or astronomical modelling. Here we report Milankovitch cycles from late Permian (Lopingian) strata at Meishan and Shangsi, South China, time calibrated by recent high-precision U-Pb dating. The evidence extends empirical knowledge of Earth's astronomical parameters before 250 million years ago. Observed obliquity and precession terms support a 22-h length-of-day. The reconstructed astronomical time scale indicates a 7.793-million year duration for the Lopingian epoch, when strong 405-kyr cycles constrain astronomical modelling. This is the first significant advance in defining the Palaeozoic astronomical time scale, anchored to absolute time, bridging the Palaeozoic-Mesozoic transition.

  13. Fullerenes and interplanetary dust at the Permian-Triassic boundary.

    Poreda, Robert J; Becker, Luann


    We recently presented new evidence that an impact occurred approximately 250 million years ago at the Permian-Triassic boundary (PTB), triggering the most severe mass extinction in the history of life on Earth. We used a new extraterrestrial tracer, fullerene, a third carbon carrier of noble gases besides diamond and graphite. By exploiting the unique properties of this molecule to trap noble gases inside of its caged structure (helium, neon, argon), the origin of the fullerenes can be determined. Here, we present new evidence for fullerenes with extraterrestrial noble gases in the PTB at Graphite Peak, Antarctica, similar to PTB fullerenes from Meishan, China and Sasayama, Japan. In addition, we isolated a (3)He-rich magnetic carrier phase in three fractions from the Graphite Peak section. The noble gases in this magnetic fraction were similar to zero-age deep-sea interplanetary dust particles (IDPs) and some magnetic grains isolated from the Cretaceous-Tertiary boundary. The helium and neon isotopic compositions for both the bulk Graphite Peak sediments and an isolated magnetic fraction from the bulk material are consistent with solar-type gases measured in zero-age deep-sea sediments and point to a common source, namely, the flux of IDPs to the Earth's surface. In this instance, the IDP noble gas signature for the bulk sediment can be uniquely decoupled from fullerene, demonstrating that two separate tracers are present (direct flux of IDPs for (3)He vs. giant impact for fullerene).

  14. Radon in the creswell crags Permian limestone caves.

    Gillmore, G K; Phillips, P S; Denman, A R; Gilbertson, D D


    An investigation of radon levels in the caves of Creswell Crags, Derbyshire, an important Site of Special Scientific Interest (SSSI) shows that the Lower Magnesian Limestone (Permian) caves have moderate to raised radon gas levels (27-7800 Bq m(-3)) which generally increase with increasing distance into the caves from the entrance regions. This feature is partly explained in terms of cave ventilation and topography. While these levels are generally below the Action Level in the workplace (400 Bq m(-3) in the UK), they are above the Action Level for domestic properties (200 Bq m(-3)). Creswell Crags has approximately 40,000 visitors per year and therefore a quantification of effective dose is important for both visitors and guides to the Robin Hood show cave. Due to short exposure times the dose received by visitors is low (0.0016 mSv/visit) and regulations concerning exposure are not contravened. Similarly, the dose received by guides is fairly low (0.4 mSv/annum) due in part to current working practice. However, the risk to researchers entering the more inaccessible areas of the cave system is higher (0.06 mSv/visit). This survey also investigated the effect of seasonal variations on recorded radon concentration. From this work summer to winter ratios of between 1.1 and 9.51 were determined for different locations within the largest cave system.

  15. Bioessential element-depleted ocean following the euxinic maximum of the end-Permian mass extinction

    Takahashi, Satoshi; Yamasaki, Shin-ichi; Ogawa, Yasumasa; Kimura, Kazuhiko; Kaiho, Kunio; Yoshida, Takeyoshi; Tsuchiya, Noriyoshi


    We describe variations in trace element compositions that occurred on the deep seafloor of palaeo-superocean Panthalassa during the end-Permian mass extinction based on samples of sedimentary rock from one of the most continuous Permian-Triassic boundary sections of the pelagic deep sea exposed in north-eastern Japan. Our measurements revealed low manganese (Mn) enrichment factor (normalised by the composition of the average upper continental crust) and high cerium anomaly values throughout the section, suggesting that a reducing condition already existed in the depositional environment in the Changhsingian (Late Permian). Other redox-sensitive trace-element (vanadium [V], chromium [Cr], molybdenum [Mo], and uranium [U]) enrichment factors provide a detailed redox history ranging from the upper Permian to the end of the Permian. A single V increase (representing the first reduction state of a two-step V reduction process) detected in uppermost Changhsingian chert beds suggests development into a mildly reducing deep-sea condition less than 1 million years before the end-Permian mass extinction. Subsequently, a more reducing condition, inferred from increases in Cr, V, and Mo, developed in overlying Changhsingian grey siliceous claystone beds. The most reducing sulphidic condition is recognised by the highest peaks of Mo and V (second reduction state) in the uppermost siliceous claystone and overlying lowermost black claystone beds, in accordance with the end-Permian mass extinction event. This significant increase in Mo in the upper Changhsingian led to a high Mo/U ratio, much larger than that of modern sulphidic ocean regions. This trend suggests that sulphidic water conditions developed both at the sediment-water interface and in the water column. Above the end-Permian mass extinction horizon, Mo, V and Cr decrease significantly. On this trend, we provide an interpretation of drawdown of these elements in seawater after the massive element precipitation event

  16. Molecular Evidence for Radical Changes in Ocean Chemistry Across the Permian Triassic Boundary at Meishan in South China

    Love, G. D.; Cao, C.; Jin, Y.; Summons, R. E.


    Samples from outcrop and from a new core drilled through the Permian Triassic Boundary at the type section at Meishan have been examined for biomarker and isotopic evidence of biotic and associated environmental change. Late Permian sediments from Meishan Beds 22-27 are characterized by indicators of anoxia including low Pr/Ph ratios and abundant aryl isoprenoids and isoreneieratane derived from the precursor carotenoid isorenieratene. The latter compounds are biomarkers for green sulfur bacteria (Chlorobiaceae) and are considered reliable indicators of euxinic water columns where sulfide extends to the photic zone. The peak of Chlorobiaceae biomarker abundance coincides with a rapid and synchronous drop in the ƒO13C and ƒO15N values of kerogen. On passing up into the Early Triassic, the biomarker signal for Chlorobiaceae wanes and is almost absent by Bed 30 where it is replaced by one for cyanobacteria including abundant hopanes and 2-methylhopanes and accompanying methyl and dimethyl alkanes. A very high value for the hopane/sterane ratio from Beds 30-38 indicates continuing dominant cyanobacterial productivity and only minor inputs from an algal plankton. The prevalence of aryl isoprenoids in P-Tr sediments at the Meishan section of South China is also recorded in a recently cored borehole, Hovea-3, of the Perth Basin, Western Australia. This suggests similar paleoenvironmental conditions prevailed across the Tethys Ocean during and immediately after the P-Tr Boundary. In particular, the presence of biomarkers for Chlorobiaceae at two separate locations indicates that water column euxinia was pervasive during the extinction event and suggests that sulfide may have been a key toxic agent.

  17. Calcimicrobialite after end-Permian mass extinction in South China and its palaeoenvironmental significance

    WANG Yongbiao; TONG Jinnan; WANG Jiasheng; ZHOU Xiugao


    Calcimicrobialites, which could be correlated to the layer 27 in Meishan section according to the Hindeodus parvus, occur abruptly on the end-Permian mass extinction boundary in South China. Microbialites mainly distribute on the top of reef facies or shallow carbonate platforms, thinning into deep facies. All the microbialites discovered are composed of micrite and coarse crystal digitate carbonate or patch carbonate. Microfossils usually dominate in the microbialites, and small gastropods, bivalves and ostracodes can also be found. This fossil assemblage represents a simple but particular remanent biota after the end-Permian mass extinction on the top of reefs or shallow carbonate platforms.Abrupt occurrence of microbialites above the mass extinction boundary is the ecological response to the end-Permian global event in reef or shallow carbonate facies. Many studies have been done on the Permian-Triassic boundary and event in deep water facies sections or middle to lower shelf facies sections. However, the calcimicrobialites in South China are mainly located above reef facies or shallow carbonate platform facies. It will surely be helpful for people to know more about the different responses in different depths of ancient marine environment during the transition between Permian and Triassic by the study of petrology,palaeontology and palaeoecology of the calcimicrobialites.

  18. Proliferation of MISS-related microbial mats following the end-Permian mass extinction in terrestrial ecosystems: Evidence from the Lower Triassic of the Yiyang area, Henan Province, North China

    Tu, Chenyi; Chen, Zhong-Qiang; Retallack, Gregory J.; Huang, Yuangeng; Fang, Yuheng


    Microbially induced sedimentary structures (MISSs) are commonly present in siliciclastic shallow marine settings following the end-Permian mass extinction, but have been rarely reported in the post-extinction terrestrial ecosystems. Here, we present six types of well-preserved MISSs from the upper Sunjiagou Formation and lower Liujiagou Formation of Induan (Early Triassic) age in the Yiyang area, Henan Province, North China. These MISSs include: polygonal sand cracks, worm-like structures, wrinkle structures, sponge pore fabrics, gas domes, and leveled ripple marks. Microanalysis shows that these MISSs are characterized by thin clayey laminae and filamentous mica grains arranged parallel to bedding plane as well as oriented matrix supported quartz grains, which are indicative of biogenic origin. Facies analysis suggests that the MISS-hosting sediments were deposited in a fluvial sedimentary system during the Early Triassic, including lake delta, riverbeds/point bars, and flood plain paleoenvironments. Abundant MISSs from Yiyang indicate that microbes also proliferated in terrestrial ecosystems in the aftermath of the Permian-Triassic (P-Tr) biocrisis, like they behaved in marine ecosystems. Microbial blooms, together with dramatic loss of metazoans, may reflect environmental stress and degradation of terrestrial ecosystems or arid climate immediately after the severe Permian-Triassic ecologic crisis.

  19. Evolución tectono-magmática durante el Pérmico al Jurásico temprano en la Cordillera del Viento (37°05´S - 37°15´S: nuevas evidencias geológicas y geoquímicas del inicio de la cuenca Neuquina Tectono-magmatic evolution during the Permian to the Early Jurassic in the Cordillera del Viento (37°05´S - 37°15´S: new geological and geochemical evidences for the early stages of the Neuquén basin

    E.J. Llambías


    podría explicar la forma no lineal de la cuenca Neuquina en la provincia del Neuquén.The southern extremity of the Cordillera del Viento is mainly composed of volcanic sequences, which form the basement of the Neuquén basin. The oldest rocks are the sediments in the Huaraco Formation, Andacollo Group, of Upper Carboniferous age. After being folded by the San Rafael orogenic phase during the Lower Permian, these sediments were intruded by granodioritic and monzogranitic plutons along with dikes and domes of rhyolitic nature. They were then covered in angular unconformity by rhyolitic ignimbrites, which are co-genetic with the intrusives. The resulting volcanic-plutonic complex is suggested to be correlative with the Permian Choiyoi Group exposed in the Cordillera Frontal and the Bloque de San Rafael geological provinces. This volcanic-plutonic complex was subjected to strong erosion during the Early to Middle Triassic. The development of an undulated erosion surface (Huarpican unconformity resulted in the exposure of the cupolas of the plutons. This surface support the Cordillera del Viento Formation (ex Groeber's Choiyoilitense, which consists of a volcanic sequence of andesitic to rhyolitic composition up to 1.300 meters thick. The next unit, the Milla Michicó Formation, was also deposited unconformably. This unit is composed of bedded flows of basalts, andesitic basalts and rhyolites with thickness up to 200 meters. The succession continued with the unconformable deposition of the La Primavera Formation, which is the oldest unit of the Cuyo Group. It is composed of marine sediments interbedded with basaltic flows at the base and rhyolitic ignimbrites in the upper levels. The thickness varies from a few meters up to 400 meters. The La Primavera Formation is covered in apparent conformity by the Los Molles Formation, which has limestones at the base and contains pelites with andesitic lahar intercalations deposited in a turbiditic regime. Regionally, the Huarpican

  20. Chondritic meteorite fragments associated with the Permian-Triassic boundary in Antarctica.

    Basu, Asish R; Petaev, Michail I; Poreda, Robert J; Jacobsen, Stein B; Becker, Luann


    Multiple chondritic meteorite fragments have been found in two sedimentary rock samples from an end-Permian bed at Graphite Peak in Antarctica. The Ni/Fe, Co/Ni, and P/Fe ratios in metal grains; the Fe/Mg and Mn/Fe ratios in olivine and pyroxene; and the chemistry of Fe-, Ni-, P-, and S-bearing oxide in the meteorite fragments are typical of CM-type chondritic meteorites. In one sample, the meteoritic fragments are accompanied by more abundant discrete metal grains, which are also found in an end-Permian bed at Meishan, southern China. We discuss the implications of this finding for a suggested global impact event at the Permian-Triassic boundary.

  1. Distribution of the Permian Monodiexodina in Karakorum and Kunlun and its Geological Significance

    YAO Jianxin; WANG Naiwen; XIAO Xuchang; JI Zhansheng; WU Guichun; WU Zhenjie; LI Boqin; WANG Jun; WANG Yong; ZHAI Qingguo


    The Permian fusulinoidean genus Monodiexodina is widely distributed in east Tethys. The genus might be an important indicator for the northern margin of Gondwana in northwestern China, but this is disputed. Monodiexodina.bearing areas can be restored as in either northern or southern middle latitudes with a symmetrical distribution between a high latitudinal, cool/cold water climatic realm and a paleotropical, warm water realm. Permian strata bearing Monodiexodina in Karakornm, Muztag Pear, and Buka Daban Pear of the east Kunlun Mountains can be correlated with each other. Faunal analyses and the stratigraphical position of Monodiexodina.bearing strata indicate that both Karakorum, east Kunlun, and the Pamirs were formed in a cool temperate sea area of the northern hemisphere in middle latitudes during the Permian, rather than at the Gondwana margin.

  2. Analyses of Sequence Stratigraphy and Environments across Permian-Triassic Boundary in Liaotian, Northwestern Jiangxi Province


    Based on the study of lithology, sedimentology and paleontology at the Permian-Triassic boundary in Liaotian, Northwestern Jiangxi Province, the sequence stratigraphy and depositional environments across the boundary are reconstructed. The top part of the Upper Permian Changxing Formation is composed of very thick-bedded light-colored dolomitic limestone formed in high deposition rate on carbonate ramp, which indacates a transgression systems tract (TST). The Lower Triassic Qinglong Formation shows continuous deposition with the underlying Upper Permian. The lower member of Qinglong Formation consists of calcareous shale, shelly limestome and dolomitic limestone with abundant bivalves (Claraia sp. )and trace fossils (Chondrites). The calcareous shale at the bottom of Lower Triassic indicates a calm deep water environment to form the condensed section (CS). The shelly limestome and dolomitic limestone with shell fossils, intraclast, algal ooide show clean but turbulent environment of carbonate ramp, which produce the deposition of highstand systems tract (TST).

  3. Archaeolithoporella and Tubiphytes : Affinties and Paleoecology in Permian Reefs of South China

    王生海; 范嘉松; J.Keith Rigby


    Archaeolithoporella is one of the most important Permian reef-building organisms, but itsaffinity has long been controversial. It was first reported in Japan and was considered as a coralline red alga. Since then it has been recognized in Permian reefs in other parts of the world. Many researchers have studied the materials they had found and the Japanese holotype specimen. Most have concluded that it is a problematic alga. or just a problematica because of weak criteria for definition. Some have even considered it inorganic. However, relict cellular structures and microtubular laminae found in well-preserved specimens from Permian reefs in South China strongly support a coralline red algal affinity for the genus.In situ preserved Archaeolithoporella is restricted within the typical framework facies and it occurs most abundantly in syndepositionally cemented framestones. This suggests that Archaeolithoporella required a firm substrate and clear but agitated sea water. Archaeolithoporella is often found

  4. Evolutionary patterns of Productida (Brachiopoda) morphology during the Permian in South China


    The evolutionary patterns of Productida (brachiopod) morphology throughout the Permian show that while the percentage proportion of Productida (brachiopod) with strongly concentric and radial ornamentation declined from the Cisuralian to the Guadalupian, and then increased towards the Changhsingian via Wuchiapingian, the percentage proportion of Productida (brachiopod) with fine concentric and radial ornamentation distinctly increased from the Cisuralian to the Guadalupian, slightly declined towards the Wuchiapingian, and then increased towards the Changhsingian. From the Cisuralian to the Changhsingian, the percentage proportion of brachiopods with spinose ornamentation shows a persistent declining trend. The shell size generally indicates a miniaturization trend at species level during the Wuchiapingian to Changhsingian (including the transitional bed). These evolutionary patterns of brachiopod ornamentation and size are possibly related to the anoxia, food shortage, sea-level fluctuation, and change of substrate in the Permian (including the Permian-Triassic transitional interval) in South China.

  5. Evolutionary patterns of Productida (Brachiopoda)morphology during the Permian in South China

    ZHANG Yang; HE WeiHong


    The evolutionary patterns of Productida (brachiopod) morphology throughout the Permian show that while the percentage proportion of Productida (brachiopod) with strongly concentric and radial ornamentation declined from the Cisuralian to the Guadalupian,and then increased towards the Changhsingian via Wuchiapingian,the percentage proportion of Productida (brachiopod) with fine concentric and radial ornamentation distinctly increased from the Cisuralian to the Guadalupian,slightly declined towards the Wuchiapingian,and then increased towards the Changhsingian.From the Cisuralian to the Changhsingian,the percentage proportion of brachiopods with spinose ornamentation shows a persistent declining trend.The shell size generally indicates a miniaturization trend at species level during the Wuchiapingian to Changhsingian (including the transitional bed).These evolutionary patterns of brachiopod ornamentation and size are possibly related to the anoxia,food shortage,sea-level fluctuation,and change of substrate in the Permian (including the Permian-Triassic transitional interval) in South China.

  6. Morphological disparity of ammonoids and the mark of Permian mass extinctions.

    Villier, Loïc; Korn, Dieter


    The taxonomic diversity of ammonoids, in terms of the number of taxa preserved, provides an incomplete picture of the extinction pattern during the Permian because of a strongly biased fossil record. The analysis of morphological disparity (the variety of shell shapes) is a powerful complementary tool for testing hypotheses about the selectivity of extinction and permits the recognition of three distinct patterns. First, a trend of decreasing disparity, ranging for about 30 million years, led to a minimum disparity immediately before the Permian-Triassic boundary. Second, the strongly selective Capitanian crisis fits a model of background extinction driven by standard environmental changes. Third, the end-Permian mass extinction operated as a random, nonselective sorting of morphologies, which is consistent with a catastrophic cause.

  7. Good genes and good luck: ammonoid diversity and the end-Permian mass extinction.

    Brayard, Arnaud; Escarguel, Gilles; Bucher, Hugo; Monnet, Claude; Brühwiler, Thomas; Goudemand, Nicolas; Galfetti, Thomas; Guex, Jean


    The end-Permian mass extinction removed more than 80% of marine genera. Ammonoid cephalopods were among the organisms most affected by this crisis. The analysis of a global diversity data set of ammonoid genera covering about 106 million years centered on the Permian-Triassic boundary (PTB) shows that Triassic ammonoids actually reached levels of diversity higher than in the Permian less than 2 million years after the PTB. The data favor a hierarchical rather than logistic model of diversification coupled with a niche incumbency hypothesis. This explosive and nondelayed diversification contrasts with the slow and delayed character of the Triassic biotic recovery as currently illustrated for other, mainly benthic groups such as bivalves and gastropods.

  8. Altered river morphology in south africa related to the permian-triassic extinction

    Ward; Montgomery; Smith


    The Permian-Triassic transition in the Karoo Basin of South Africa was characterized by a rapid and apparently basin-wide change from meandering to braided river systems, as evidenced by preserved sedimentary facies. This radical changeover in river morphology is consistent with geomorphic consequences stemming from a rapid and major die-off of rooted plant life in the basin. Evidence from correlative nonmarine strata elsewhere in the world containing fluvial Permian-Triassic boundary sections suggests that a catastrophic terrestrial die-off of vegetation was a global event, producing a marked increase in sediment yield as well as contributing to the global delta(13)C excursion across the Permian-Triassic boundary.

  9. Trouble Upstairs: Reconstructing Permian-Triassic Climate during Siberian Traps Magmatism

    Black, B. A.; Neely, R. R., III; Lamarque, J. F.; Elkins-Tanton, L. T.; Mills, M. J.


    The eruption of large igneous provinces can transfer significant masses of volatiles from Earth's interior to the atmosphere. What are the consequences of this degassing for habitability and extinction? In this presentation, we consider this question in the context of Siberian Traps magmatism, which has been shown to overlap within geochronologic uncertainty with catastrophic deterioration of Permian-Triassic marine and terrestrial ecosystems. To investigate the impacts of endogenic gases on climate, atmospheric chemistry, and ocean circulation, we conducted a series of numerical experiments with a comprehensive global model for the Permian-Triassic. Our simulations predict the intensity and distribution of acid rain and ozone depletion, with implications for terrestrial biota. We further explore feedbacks between sulfur emissions, transient cooling, and shifts in ocean circulation. We suggest that Siberian Traps magmatism may have triggered several distinct kill mechanisms in the oceans and on land, contributing to a complex combined pattern of environmental stress and latest Permian ecological failure.

  10. Timing of mammal-like reptile extinctions across the Permian-Triassic boundary in South Africa

    MacLeod, Kenneth G.; Smith, Roger M. H.; Koch, Paul L.; Ward, Peter D.


    The rate, timing, and pattern of change in different regions and paleoenvironments are critical for distinguishing among potential causes for the Permian-Triassic (P-T) extinction. Carbon isotopic stratigraphy can provide global chronostratigraphic control. We report a large δ13C excursion at the P-T boundary and no long-term Permian δ13C trends for samples from the interior of Pangea. Stratigraphic gaps between available samples limit the resolution of our δ13C curve, but the excursion is within a 15-m-thick zone of overlap between Permian and Triassic taxa. Sedimentological and taphonomic observations demonstrate that this 15 m interval does not represent geologically instantaneous deposition. Together these data support a rapid and globally synchronous P-T event, but suggest that it occurred over a geologically resolvable interval of time.

  11. Alkane Biomarkers in Permian-Triassic Boundary Strata at Meishan Section, Changxing,Zhejiang Province


    Meishan Section D in Changxing County, Zhejiang Province, China has been selected as the global stratotype of the Permian-Triassic boundary and various studies have been done at the boundary, but the gas chromatographic-mass spectrographic analysis of alkane biomarkers has not been investigated. This paper presents the results of a study of the biomarkers analyzed in a series of samples across the Permian-Triassic boundary at both Meishan Section A and Section D. The results show that the overall concentration of alkane biomarkers in the Permian-Triassic boundary strata is high in Bed 26 while it is low in Bed 27. A variety of biomarker parameters demonstrate that the main sources of organic matter in the sediment are algae and bacteria and that the depositional environment varied from weakly oxidizing to reducing during the studied interval.

  12. Ocean anoxia did not cause the Latest Permian Extinction

    Proemse, Bernadette C.; Grasby, Stephen E.; Wieser, Michael E.; Mayer, Bernhard; Beauchamp, Benoit


    The Latest Permian Extinction (LPE, ~252 million years ago) was a turning point in the history of life on Earth with a loss of ~96% of all marine species and ~70% of all terrestrial species. While, the event undoubtedly shaped the evolution of life its cause remains enigmatic. A leading hypothesis is that the global oceans became depleted in oxygen (anoxia). In order to test this hypothesis we investigated a proxy for marine oxygen levels (molybdenum isotopic composition) in shale across the LPE horizon located on the subtropical northwest margin of Pangea at that time. We studied two sedimentary records in the Sverdrup basin, Canadian High Arctic: Buchanan Lake (eastern Axel Heiberg Island; 79° 26.1'N, 87° 12.6'W), representing a distal deep-water slope environment, and West Blind Fiord (southwest Ellesmere Island; 78° 23.9'N, 85° 57.2'W), representing a deep outer shelf environment (below storm wave base). The molybdenum isotopic composition (δ98/95Mo) of sediments has recently become a powerful tool as a paleo-oceanographic proxy of marine oxygen levels. Sample preparation was carried out in a metal-free clean room facility in the isotope laboratory of the Department of Physics and Astronomy, University of Calgary, Canada, that is supplied by HEPA-filtered air. Molybdenum isotope ratios were determined on a Thermo Scientific multi-collector inductively coupled plasma mass spectrometer (MC-ICP-MS) with an uncertainty better than ±0.10o for δ98/95Mo values. Results from the Buchanan Lake section show a large shift in δ98/95Mo values from 2.02o to +2.23o at the extinction horizon, consistent with onset of euxinic conditions. In contrast, West Blind Fiord shales, representing the sub-storm wave base shelf environment, show little change in the molybdenum isotopic composition (1.34o to +0.05), indicating ongoing oxic conditions across the LPE (Proemse et al., 2013). Our results suggest that areas of the Pangea continental shelf (North West Pangea) experienced

  13. Permian Paleomagnetism of Eastern Kazakhstan: Implications Concerning the Amalgamation of Eurasia

    Collins, A. Q.; Levashova, N. M.; Degtyarev, K. E.; Bazhenov, M. L.; Van der Voo, R.


    Kazakhstan is the central and most complex region of the Ural-Mongol belt of Eurasia. This region is bounded by the Baltica platform to the west, the Siberian platform to the northeast, and Tarim to the south. Many geologists think that the major tectonic units of Kazakhstan had amalgamated by the Silurian, with the possibility of relatively small motions through the middle-late Paleozoic, whereas others have postponed the welding of Kazakhstan until the Late Carboniferous. Furthermore, some authors have suggested that the Siberian block and the adjacent regions of the Ural-Mongol belt were displaced with respect to Baltica, westward and northwestward (in present day co-ordinates) during the Late Permian along strike slip faults by some 1500km. We studied Upper Permian volcanics from eastern Kazakhstan (47.5N, 80.5E). A univectorial characteristic remanent magnetization (ChRM) component was isolated from most samples after the removal of a weak, low-temperature remanence. The fold test (McFadden and Jones, 1981) is positive. The observed pole (PLat: 43, Plong: 172, a95: 3.6) falls close to the Baltica APWP and nearly coincides with the mean pole for middle Permian time (261-275 Ma; PLat: 46, Plong: 166, a95: 3.8). The overall agreement of the European and Siberian APWP's is rather good for the Permian, and Late Permian paleolatitudes from Tarim also agree with the European grid. Thus we conclude that our study area had already been attached to the European, Siberian, and Tarim blocks during the Late Permian. This interpretation disagrees with models that suggest large-scale displacement between the European and Siberian platforms during the final stages of Ural-Mongol fold belt evolution.

  14. Sedimentary features and exploration targets of Middle Permian reservoirs in the SW Sichuan Basin

    Guoming Xu


    Full Text Available The exploration direction and targets for the large-scale Middle Permian gas reservoirs in the Sichuan Basin are hot spots and challenges in current exploration researches. The exploration successes of large gas field of Cambrian Longwangmiao Formation in Gaoshiti-Moxi region, Central Sichuan Basin, indicated that prospective sedimentary facies belt was the basis for the formation of large gas fields. In this paper, based on seismic data, outcrop data and drilling data, the tectonic framework and sedimentary features of the Middle Permian in the SW Sichuan Basin were comprehensively studied. The following conclusions were reached from the perspective of sedimentary facies control: (1 during the Middle Permian, this region was in shallow water gentle slope belts with high energy, where thick reef flat facies were deposited; (2 the basement was uplifted during Middle Permian, resulting in the unconformity weathering crust at the top of Maokou Formation due to erosion; the SW Sichuan Basin was located in the karst slope belt, where epigenic karstification was intense; and (3 reef flat deposits superimposed by karst weathering crust was favorable for the formation of large-scale reef flat karst reservoirs. Based on the combination of the resources conditions and hydrocarbon accumulation conditions in this region, it was pointed out that the Middle Permian has great potential of large-scale reef flat karst gas reservoir due to its advantageous geological conditions; the Middle Permian traps with good hydrocarbon accumulation conditions were developed in the Longmen Mountain front closed structural belt in the SW Sichuan Basin and Western Sichuan Basin depression slope belt, which are favorable targets for large-scale reef flat karst reservoirs.

  15. Petrogenesis of Permian A-type granitoids in the Cihai iron ore district, Eastern Tianshan, NW China: Constraints on the timing of iron mineralization and implications for a non-plume tectonic setting

    Zheng, Jiahao; Mao, Jingwen; Chai, Fengmei; Yang, Fuquan


    The geochronology and geochemistry of granitoids in the Eastern Tianshan, NW China provide important constraints on the timing of iron mineralization, as well as in understanding evolution history of the southern Central Asian Orogenic Belt (CAOB). Here we present results from a detailed study on granitoid rocks from the Cihai iron ore district in the Beishan region, southern part of the Eastern Tianshan. The granitoid rocks are composed of granodiorite, quartz monzonite, granite, and monzonite. Zircon U-Pb analyses yielded the ages of 294.1 ± 2.2 Ma, 286.5 ± 0.7 Ma, 284.3 ± 3.3 Ma, and 265.6 ± 3.0 Ma, respectively, suggesting they were formed in Early-Middle Permian. Among these granitoid rocks, the ages of quartz monzonite and granite are close to the timing of iron mineralization (~ 282 Ma), indicating they may provide a source of iron in the Cihai ore district. Geochemically, the granodiorite, granite, and quartz monzonite samples are characterized by high FeOt/(FeOt + MgO) and Ga/Al ratios (0.84-0.94 and 2.28-3.27, respectively), as well as high zircon saturation temperatures (781-908 °C), similar to those of typical A-type granitoids. Isotopically, they display consistently depleted Hf isotopic compositions (εHf(t) = + 1.18 to + 15.37). Geological, geochemical, and isotopic data suggest that the Cihai A-type granitoids were derived from melting of juvenile lower crust. Some Early Permian A-type granitoids were recently identified in the Tarim and Eastern Tianshan with the ages between 294 and 269 Ma. The A-type granitoids in the Eastern Tianshan formed earlier between 294-284 Ma and exhibit characteristics of A2 type granitoids, whereas the A-type granitoids in the Tarim formed later between 277-269 Ma and show A1 granitoids affinity. We suggest that the Permian Tarim mantle plume does not account for the formation of the A-type granitoids in the Eastern Tianshan area, and the Eastern Tianshan was in a non-plume tectonic setting during Early Permian

  16. Anoxia duirng the Late Permian Binary Mass Extinction and Dark Matter

    Abbas, S; Mohanty, S; Abbas, Samar; Abbas, Afsar; Mohanty, Shukadev


    Recent evidence quite convincingly indicates that the Late Permian biotic crisis was in fact a binary extinction with a distinct end-Guadalupian extinction pulse preceding the major terminal end-Permian Tartarian event by 5 million years. In addition anoxia appears to be closely associated with each of these end-Paleozoic binary extinctions. Most leading models cannot explain both anoxia and the binary characteristic of this crisis. In this paper we show that the recently proposed volcanogenic dark matter scenario succeeds in doing this.

  17. Reexamination of Radiolarian Biostratigraphy in Permian in Pelagic Chert Sequences at Dachongling Section, South China


    Permian radiolarian biostratigraphy was reexamined in the bedded chert section in Qinzhou area of southeast Guangxi Zhuang Autonomous Region, South China. On the basis of the analyses of characteristic radiolarian composition, six radiolarian zones are recognized, namely, Pseudoalbaillella longtanensis Zone, Pseudoalbaillella globosa Zone, Follicucullus monacanthus Zone, Follicucullus scholasticus Zone, Follicucullus charveti Zone and Neoalbaillella ornithoformis Zone, in ascending order. Correlation of these radiolarian zones with those in Japan is presented. The Neoalbaillella ornithoformis Zone was firstly discovered in Dachongling Section, which may indicate that pelagic chert of the late Permian existed there, according to the co-occurring conodonts.

  18. RIO Country Report 2015

    Grimpe, Christoph; Mitchell, Jessica

    The 2015 series of RIO Country Reports analyse and assess the policy and the national research and innovation system developments in relation to national policy priorities and the EU policy agenda with special focus on ERA and Innovation Union. The executive summaries of these reports put forward...

  19. RIO Country Report 2015

    Grimpe, Christoph; Mitchell, Jessica

    The 2015 series of RIO Country Reports analyse and assess the policy and the national research and innovation system developments in relation to national policy priorities and the EU policy agenda with special focus on ERA and Innovation Union. The executive summaries of these reports put forward...

  20. The Concertina Coast: the role of basement inheritance during repeated reactivation events along Australia's northern margin since the Permian

    Keep, Myra; Gartrell, Anthony


    The present day configuration of Australia's northern margin includes a series of Phanerozoic sedimentary basins forming the North West Shelf. Their polyphase history, dominantly extensional, and closely associated with the breakup of Eastern Gondwana, includes the early formation of intracratonic basins (from the mid-Devonian), overprinted by Permo-Carboniferous rifting that generated the dominant NE-trending structural trends that persist to the present-day. Subsequent Mesozoic extension, associated with the formation of abyssal plains, further refined the margin, creating additional depocentres. During this polyphase rift history, a number of periods of inversion have punctuated the margin. These include a Carboniferous event (the Meda Transpression), a late Permian to Early Triassic event, sometimes referred to as the Bedout Movement (possibly transtensional), and two events, one in the Middle to Late Triassic, followed by another in the Late Triassic to Early Jurassic, often referred to as the Fitzroy events. These various events, recorded locally, caused inversion, folding, uplift and erosion where documented, with the Fitzroy events described as transpressional, resulting from right-lateral oblique inversion. Subsequent inversion during the Cretaceous, also attributed to dextral transpression, caused long wavelength folding and fault inversion in some basins. Whereas the effects of earlier inversions are somewhat sporadic across the North West Shelf, the effects of Neogene inversion have been documented across both the active and passive segments of the present day North West Shelf, and also appear to be strongly controlled by right-lateral oblique reactivation mechanisms, with associated seismicity and focal mechanism solutions. The history of the North West Shelf therefore includes 6 discrete episodes of reactivation and inversion, apparently strongly dominated by oblique mechanisms, which punctuate the long, multi-phase extensional history. Whereas

  1. New paleomagnetic results from the Permian and Mesozoic rocks in central and northeast Thailand: their implications for the construction of the Indochina block in Pangea

    Zhang, D.; Yan, Y.; Huang, B.; Zhao, J.


    Paleomagnetic studies of the Indochina block, aiming to reconstruct the paleogeography, have been undertaken for several decades. Since the Indochina block is lack of reliable paleomagnetic data to constraint its paleo-positions during the Middle Permian to Upper Triassic, the paleogeography reconstruction is still in debate between different models reported. Here we present new paleomagnetic data of Middle Permian to Upper Triassic sediment rocks from the Indochina block in Thailand, and recalculate paleomagnetic data reported by different authors. We collected the Permian samples in 20 sites distributed in the central Thailand, and Triassic samples from the Huai Hin Lat and Nam Phong formations in 13 sites in the northern Thailand.The magnetic directions of the 11 sites of Permian limestones are not significantly clustered after tilt correction which implying a remagnetized result. Remarkably, in geographic coordinate, the 11 sites were distributed along a circle showing a similar inclination which is 22.9° implying the paleolatitude to be about 12°. Totally, 13 sites from the Huai Hin Lat formation are included in the calculation of the formation mean direction Dg/Ig = 21.4°/38.1°, kg = 19.5, α95 = 9.6° before and Ds/Is = 43.0°/48.0°, ks = 47.4, α95 = 6.1°, N = 13 after bedding correction. A pre-folding characteristic magnetization is suggested by the positive fold test result derived from the Huai Hin Lat formation, and thus implies a primary remanence of the Norian Stage Upper Triassic rocks. A new Nam Phong formation mean direction derived from 11 sites is Dg/Ig = 36.5°/31.3°, kg = 14.7, α95 = 12.3°before and Ds/Is = 36.4°/37.8°, ks = 68.5, α95 = 5.6°, N = 11 after bedding correction. The two formation mean directions correspond to the magnetic pole positions , Plat./Plon=48.7°N/165.9°E, A95=7.2° and Plat./Plon=55.2°N/178.0°E, A95=5.9°, respectively. A remarkable tectonic movement (~8° southward) of the Indochina block from the age of

  2. Inversion tectonics in the Anayet Permian basin (Axial Zone, Central Pyrenees)

    Rodriguez, L.; Cuevas, J.; Tubía, J. M.


    During Permian times the Pyrenees were characterized by extensional tectonics that opened subsident basins with pull-apart geometries. The Anayet Permian basin crops out in the western Spanish Axial Zone between the Aragon and Tena valleys. It is WNW-trending and it is filled by a continental Permian succession that represents the first post-variscan deposits in the area. Permian deposits rest discordantly over Devonian to Carboniferous limestones, sandstones and slates. In the Anayet basin, Permian deposits have been classically divided in four main detrital groups, mainly composed of sandstones and conglomerates, with three basic volcanic episodes interbedded (Gisbert, 1984, Bixel, 1987). Due to the lithological characteristics of the Permian rocks in this region, there are almost no accurate age constraints for these units. A detailed structural study of the area, including mapping and balanced cross-sections, shows increasingly older rocks to the west of the Anayet basin. Moreover, it can be deduced a mean slope of around 11 % to the west for the basin. These data confirm that the basin depocenter was located to the west and that the Anayet basin was partitioned by N10 °E-trending normal faults. Although the contacts between the Permian and the Devono-Carboniferous rocks are covered by quaternary deposits in most of its extent, a fault contact can also be recognized. The fault contact is a 3 m thick shear zone oriented N120 ° E and dipping 60° to the North. It develops breccias, fault gouges and sigmoidal S-C tectonites indicating a reverse motion. The contact places Permian slates and sandstones over Carboniferous limestones and is almost parallel to the alpine cleavage deforming Permian rocks. The slope of the contact together with the presence of younger rocks in the hangingwall of the reverse fault points out that the original contact was a normal fault reactivated as a high-angle reverse fault during the positive inversion tectonics induced by the

  3. Comment on “Stratigraphy, petrography and dispersion of the lower Permian syn-eruptive deposits in the Viar Basin, Spain” by S. Sierra, C. Moreno and E. Pascual [Sedimentary Geology 217 (2009) 1-29

    Wagner, R. H.; Mayoral, E.


    Comments are provided on a published paper on "Lower Permian" strata in SW Spain [S. Sierra, C. Moreno and E. Pascual, Stratigraphy, petrography and dispersion of the lower Permian syn-eruptive deposits in the Viar Basin, Spain, Sedimentary Geology 217 (2009) 1-29], which failed to take into account up-to-date information on this most southerly occurrence in western Europe. This generally lacustrine basin commenced with valley fill deposits in a deeply incised palaeotopography and contains mainly red beds with two basaltic intervals and a major acidic volcanic episode linked to a nearby volcanic centre to the Northeast. The stratigraphic succession shows southeasterly onlap. Although a syn-sedimentary fault has been postulated on the NE basin margin, the field evidence disproves this notion as well as the assumption that this "Early Permian" basin would have been controlled by a rejuvenated Ossa-Morena/South Portuguese boundary fault. In fact, this basin lies within the (former) area of Ossa-Morena.

  4. The Cordon del Portillo Permian magmatism, Mendoza, Argentina, plutonic and volcanic sequences at the western margin of Gondwana

    Gregori, Daniel; Benedini, Leonardo


    The Cerro Punta Blanca, Cerro Bayo and Cerro Punta Negra stocks, parts of the Cordillera Frontal Composite Batholith, cropping out in the Cordón del Portillo, records the Gondwana magmatic development of the Cordillera Frontal of Mendoza, in western Argentina. In this area, the San Rafael Orogenic phase, that represents the closure of the Late Carboniferous-Early Permian marine basins, begins at 284 Ma, and ceased before 276 Ma. The Cerro Punta Blanca, Cerro Bayo and Cerro Punta Negra stocks represent a post-orogenic magmatism and are equivalents to the Choiyoi Group. The Gondwana magmatic activity in the Cordón del Portillo area can be divided into two stages. The Cerro Punta Blanca stock (c.a. 276 Ma) represents an early post-orogenic, subduction-related magmatism similar to the basic-intermediate section of the Choiyoi Group (c.a. 277 Ma). The late post-orogenic second event was recorded by the Cerro Bayo (262 Ma) and Cerro Punta Negra stocks which represent a transition between subduction-related and intra-plate magmatism. This event represents the intrusive counterpart of the acidic facies of the upper section of the Choiyoi Group (c.a. 273 Ma). This extensional condition continued during the Triassic when the Cacheuta basin developed.

  5. Global taxonomic diversity of anomodonts (tetrapoda, therapsida) and the terrestrial rock record across the Permian-Triassic boundary.

    Fröbisch, Jörg


    The end-Permian biotic crisis (~252.5 Ma) represents the most severe extinction event in Earth's history. This paper investigates diversity patterns in Anomodontia, an extinct group of therapsid synapsids ('mammal-like reptiles'), through time and in particular across this event. As herbivores and the dominant terrestrial tetrapods of their time, anomodonts play a central role in assessing the impact of the end-Permian extinction on terrestrial ecosystems. Taxonomic diversity analysis reveals that anomodonts experienced three distinct phases of diversification interrupted by the same number of extinctions, i.e. an end-Guadalupian, an end-Permian, and a mid-Triassic extinction. A positive correlation between the number of taxa and the number of formations per time interval shows that anomodont diversity is biased by the Permian-Triassic terrestrial rock record. Normalized diversity curves indicate that anomodont richness continuously declines from the Middle Permian to the Late Triassic, but also reveals all three extinction events. Taxonomic rates (origination and extinction) indicate that the end-Guadalupian and end-Permian extinctions were driven by increased rates of extinction as well as low origination rates. However, this pattern is not evident at the final decline of anomodont diversity during the Middle Triassic. Therefore, it remains unclear whether the Middle Triassic extinction represents a gradual or abrupt event that is unique to anomodonts or more common among terrestrial tetrapods. The end-Permian extinction represents the most distinct event in terms of decline in anomodont richness and turnover rates.

  6. The Permian Rotliegend reservoir architecture of the Dutch Koekoekspolder geothermal doublet

    Mijnlieff, H.F.; Bloemsma, M.R.; Donselaar, M.E.; Henares, S.; Redjosentono, A.E.; Veldkamp, J.G.; Weltje, G.J.


    The Dutch Koekoekspolder geothermal doublet was drilled in 2011 targeting the Permian Rotliegendreservoir. The encountered reservoir properties were less favorable than expected pre-drill. Post-drill integrated evaluation of vintage data and the new data from the geothermal wells resulted in a refin

  7. Assessment of Permian tight oil and gas resources in the Junggar basin of China, 2016

    Potter, Christopher J.; Schenk, Christopher J.; Tennyson, Marilyn E.; Klett, Timothy R.; Gaswirth, Stephanie B.; Leathers-Miller, Heidi M.; Finn, Thomas M.; Brownfield, Michael E.; Pitman, Janet K.; Mercier, Tracey J.; Le, Phuong A.; Drake, Ronald M.


    Using a geology-based assessment methodology, the U.S. Geological Survey estimated undiscovered, technically recoverable mean resources of 764 million barrels of oil and 3.5 trillion cubic feet of gas in tight reservoirs in the Permian Lucaogou Formation in the Junggar basin of northwestern China.

  8. A tale of two extinctions : converging end-Permian and end-Triassic scenarios

    van de Schootbrugge, Bas; Wignall, Paul B.


    The end-Permian (c. 252 Ma) and end-Triassic (c. 201 Ma) mass-extinction events are commonly linked to the emplacement of the large igneous provinces of the Siberia Traps and Central Atlantic Magmatic Province, respectively. Accordingly, scenarios for both extinctions are increasingly convergent and

  9. Phosphatic Permian rocks of the Adobe Range, Nevada, and their environment of deposition

    Ketner, Keith Brindley


    Permian sedimentary rocks in the Adobe range, northern Nevada, are phosphatic, and although the particles of phosphate are relatively more disseminated, they closely resemble the rocks of the Phosphoria Formation. In the northern Adobe Range, where the entire Permian sequence is approximately correlative with the Phosphoria Formation, it is 200 m thick and averages 1.7 percent P2O5 . In the southern Adobe Range, the Permian sequence is more than 1,700 m thick, and the upper half which is roughly correlative with the Phosphoria Formation averages more than 2 percent P2O5. Some thin beds in rocks of Permian age contain more than 20 percent P2O5. Phosphatic rocks of the Adobe Range were deposited in shallow water among islands in the western part of the epicontinental Phosphoria sea. The continental margin and the open ocean lay far to the west. At the same time, the Phosphoria Formation was being deposited in the eastern and central parts of the Phosphoria sea. Theories based on the work of Kasakov done in 1937 relating phosphate deposition directly to sites of upwelling oceanic waters are questioned. Nondeposition of diluent materials such as detritus and carbonate is probably of more importance in producing phosphate in economic concentrations than is geographic position with respect to upwelling waters.

  10. Climatic and biotic upheavals following the end-Permian mass extinction

    Romano, C.; Goudemand, N.; Vennemann, T.W.; Ware, D.; Schneebeli-Hermann, E.; Hochuli, P.A.; Brühwiler, T.; Brinkmann, W.; Bucher, H.


    Recovery from the end-Permian mass extinction is frequently described as delayed, with complex ecological communities typically not found in the fossil record until the Middle Triassic epoch. However, the taxonomic diversity of a number of marine groups, ranging from ammonoids to benthic foraminifer

  11. The continental Permian-Triassic boundary in the Netherlands: Implications for the geomagnetic polarity time scale

    Szurlies, M.; Geluk, M.C.; Krijgsman, W.; Kürschner, W.M.


    In Central and NW Europe, the transition from the Permian to the Triassic (i.e., the Zechstein–Buntsandstein boundary interval) is developed mainly in red bed facies. This continental sedimentary succession is marked by relatively high sedimentation rates providing a high temporal resolution favorab

  12. Pattern of marine mass extinction near the Permian-Triassic boundary in South China.

    Jin, Y G; Wang, Y; Wang, W; Shang, Q H; Cao, C Q; Erwin, D H


    The Meishan section across the Permian-Triassic boundary in South China is the most thoroughly investigated in the world. A statistical analysis of the occurrences of 162 genera and 333 species confirms a sudden extinction event at 251.4 million years ago, coincident with a dramatic depletion of delta13C(carbonate) and an increase in microspherules.

  13. Controls on body size during the Late Permian mass extinction event.

    He, W-H; Twitchett, R J; Zhang, Y; Shi, G R; Feng, Q-L; Yu, J-X; Wu, S-B; Peng, X-F


    This study examines the morphological responses of Late Permian brachiopods to environmental changes. Quantitative analysis of body size data from Permian-Triassic brachiopods has demonstrated significant, directional changes in body size before, during and after the Late Permian mass extinction event. Brachiopod size significantly reduced before and during the extinction interval, increased for a short time in more extinction-resistant taxa in the latter stages of extinction and then dramatically reduced again across the Permian/Triassic boundary. Relative abundances of trace elements and acritarchs demonstrate that the body size reductions which happened before, during and after extinction were driven by primary productivity collapse, whereas declining oxygen levels had less effect. An episode of size increase in two of the more extinction-resistant brachiopod species is unrelated to environmental change and possibly was the result of reduced interspecific competition for resources following the extinction of competitors. Based on the results of this study, predictions can be made for the possible responses of modern benthos to present-day environmental changes.

  14. Biogenic silica microfossils in sediments of the Permian - Carboniferous Unayzah Formation, Saudi Arabia

    Garming, J.F.L.; Franks, S.G.; Cremer, H.; Abbink, O.A.


    Biogenic silica particles (BSPs) have been discovered in sediments of the Permian - Carboniferous Unayzah Formation of Saudi Arabia. The BSPs are extracted from sediments that are generally barren of macro- or microfossils. BSPs have been found in the Basal Khuff Clastics (BKC), and the Unayzah A, B

  15. The prospects of oil occurrence of the upper Permian and Triassic layers in the Michaiusk area

    Alabushin, A.A.; Turchaninov, G.N.


    Oil manifestations in the upper Permian and Triassic deposits in the Michaiusk area are analyzed. In the Triassic deposits, the T-1 layer is identified, and it is recommended as a first order objective for the set up of prospect developing operations in the search for oil.

  16. Sphenophytes, pteridosperms and possible cycads from the Wuchiapingian (Lopingian, Permian) of Bletterbach (Dolomites, Northern Italy)

    Kustatscher, E.; Bauer, K.; Butzmann, R.; Fischer, T.C.; Meller, B.; van Konijnenburg-van Cittert, J.H.A.; Kerp, H.


    The Bletterbach flora is the most important late Permian (Lopingian) flora of the Southern Alps. The study of a new fossiliferous bed stratigraphically below the cephalopod bed yielded almost 500 plant fossils, 28 of which belong to rare Lopingian plant groups: horsetails, seed ferns and possible cy

  17. Taeniopterid lamina on Phasmatocycas megasporophylls (Cycadales) from the Lower Permian of Kansas, U.S.A.

    Gillespie, W.H.; Pfefferkorn, H.W.


    New specimens of Phasmatocycas and Taeniopteris from the original Lower Permian locality in Kansas demonstrate organic attachment of the two and corroborate Mamay's hypothesis that Phasmatocycas and Taeniopteris were parts of the same plant. These forms also suggest that cycads evolved from taxa with entire leaves; i.e. Taeniopteris, rather than from pteridosperms with compound leaves. ?? 1986.

  18. Lower Permian Radiolaria from the Pos Blau area, Ulu Kelantan, Malaysia

    Jasin, Basir; Aziz Ali, Che


    Twenty two species of Radiolaria were identified from a 30 mthick chert sequence exposed at a roadcut near Pos Blau, Ulu Kelantan. The radiolarian assemblage represents the top part of the Pseudoalbaillella lomentaria Zone, upper Wolfcampian which is equivalent to the Sakmarian, Lower Permian. These radiolarian faunas contain elements present in both Southern Urals and Japanese faunas.

  19. A new Permagrionidae from the Middle Permian of the South of France (Odonatoptera: Protozygoptera).

    Fate, Caitlin; Lapeyrie, Jean; Nel, Andre


    The new permagrionid protozygopteran genus and species Salagoulestes wesleyi is described from the Middle Permian of Lodève Basin, Salagou Formation. It seems to be more closely related to the two genera Scytolestes and Permagrion than to any other Permagrionidae. It increases the diversity of the odonatopteran fauna in the Salagou Formation to 14 different species.

  20. The natural remanent magnetizations of the exeter volcanic traps (Permian, Europe)

    Zijderveld, J.D.A.


    The natural remanent magnetizations of 35 samples from five quarries in the Permian traps from the Exeter region were carefully examined. A considerable number of the samples contained a secondary magnetism, the intensity of which was directly proportional to the susceptibility. This secondary magne

  1. Ammonoid Succession of Setorym River (Verkhoyansk Area) and Problem of PermianTriassic Boundary in Boreal Realm


    The presence of a single Otoceras species (O. boreale), morphologically very variable, at the base of the Nekuchan Formation in Verkhoyansk, we believe, is to be obvious. Some morphological evidence leaves no doubt that two described morphs of O. boreale are a strictly corresponding sexual dimorphic pair. It is very likely that Kummel's idea that Canadian O. concavum Tozer is an invalid species is truthful, considering the range of variability seen in larger Siberian and Himalayan Otoceras fauna.Just above the upper Tatarian Imtachan Formation, the six stages of ammonoid succession can be recognized within the lower part of the Nekuchan Formation in the Setorym River Section: (a) Otoceras boreale ; (b) Otoceras boreale - Tom po phiceras pascoei ; (c) Otoceras boreale - Tom po phiceras pascoei Aldanoceras ; (d) Tompophiceras pascoei -Otoceras boreale - Aldanoceras ; ( e) Tompophiceras morpheous - T. pascoei - Aldanoceras ; (f) Tom po phiceras mor pheous - T. pascoei - Wordieoceras domokhotovi -Ophiceras transitorium ; (g) Tompophiceras morpheous - T. pascoei, corresponding to the Otoceras boreale and Tompophiceras morpheous zones. In spite of the domination of Otocerataceae or Xenodiscaceae in both of these zones and the presence of some Permian type conodonts in the lower part of the Otoceras boreale Zone, they seem to be early Induan in age on the basis of the following arguments: (1) in contrast to the underlying regressive type sediments of the Upper Tatarian Imtachan Formation, both the Otoceras boreale and the Tompophiceras morpheous zones of the lowermost part of the Nekuchan Formation correspond to the single transgressive cycle; (2) typical early Induan ammonoids (Ophiceras and Wordieoceras) have been recognized in the Tompophiceras morpheous zone;(3) all described ammonoid succession stages (a-g) are characterized by very gradual changes and therefore correspond to the different parts of the single zone or to the different zones of the same stage,but not

  2. Age, petrogenesis, and tectonic setting of the Permian bimodal volcanic rocks in the eastern Jiamusi Massif, NE China

    Bi, Jun-Hui; Ge, Wen-Chun; Yang, Hao; Wang, Zhi-Hui; Dong, Yu; Liu, Xi-Wen; Ji, Zheng


    We present new in situ zircon U-Pb and Hf isotope, whole-rock geochemical, and Sr-Nd isotopic data for volcanic rocks from the Jiejinkou and Baoqing areas in the eastern Jiamusi Massif. These volcanic rocks are bimodal and consist of basalts, basaltic andesites, rhyolites, and rhyolitic tuffs that can be subdivided into mafic and silicic groups. Zircon U-Pb dating by LA-ICP-MS indicates that these volcanic rocks were erupted between the Early and Middle Permian (290-267 Ma). The mafic rocks in this area have positive εNd(t) (+0.07 to +6.43) values, and are enriched in light rare earth elements (LREEs) and depleted in heavy REE, Nb, and Ta. From these rocks, the meta-basalt of Jinlu and basaltic andesite of Taipinggou and Haojiatun were derived from parental magmas generated by the partial melting of depleted mantle wedge material that was metasomatized by subduction-related melts. These magmas then underwent variable degrees of fractional crystallization and assimilated insignificant amounts of crustal material. The meta-basalt of Liming likely originated from the metasomatized mantle-derived melts hybridized by the convective asthenosphere during the evolution of the magmas. In comparison, the silicic rocks have negative εNd(t) and variable zircon εHf(t) values, are enriched in the large-ion lithophile elements (LILEs) and LREE, and are depleted in high-field-strength elements (e.g., Nb, Ta, and Ti), yielding arc-like geochemical signatures. The geochemical and zircon εHf(t) characteristics of Jiangfeng and Longtouqiao rhyolites are indicative of formation from magmas generated by the partial melting of mafic lower crustal material, whereas the Liming meta-rhyolite was probably produced from a source involving some depleted mantle components. The bimodal volcanic rocks provide convincing evidence that the Early-Middle Permian volcanism in the Jiamusi Massif occurred in an extensional environment probably associated with slab break-off during the westward

  3. Permian/Triassic Boundary and Its Correlation in the Daxian-Xuanhan Area, Northeastern Sichuan Province, China

    MA Yongsheng; MOU Chuanlong; TAN Qinyin; YU Qian; WANG Ruihua


    There are different opinions about the Permian/Triassic boundary and its correlation in the Daxian- Xuanhan area because of lacking typical lithological symbols and fossil criterions. Especially,it is very hard to determine the Permian/Triassic boundary in the Daxian-Xuanhan buried areas. This brings many difficulties to geological research and oil and gas exploration. Based on field investigation and studies of well cores, newly found fossils, lithological characteristics and well log analyses, the authors discussed the Permian/Triassic boundary in the Daxian-Xuanhan area and brought forward a specific scheme for its determination and regional correlation.

  4. Ostracods (Crustacea associated with microbialites across the Permian-Triassic boundary in Dajiang (Guizhou Province, South China

    Marie-Beatrice FOREL


    Full Text Available 26 samples were processed for a taxonomic study of ostracods from the Upper Permian (Changhsingian - Lower Triassic (Griesbachian interval of the Dajiang section, Guizhou Province, South China. 112 species belonging to 27 genera are recognized. Five new species are described: Acratia candyae sp. nov, Bairdia adelineae sp. nov., Bairdia? huberti sp. nov., Bairdia jeromei sp. nov., Orthobairdia jeanlouisi sp. nov. The unexpected survival faunas associated with microbial formations in the aftermath of the end-Permian extinction are documented for the first time. Ostracod biodiversity variations and palaeo-environmental modifications associated with microbial growth through the Permian-Triassic boundary (PTB are discussed.

  5. First report of the fertile plant genus Umkomasia from Late Permian beds in India and its biostratigraphic significance

    Chandra, S.; Singh, K.J.; Jha, N. [Birbal Sahni Institute for Paleobotany, Lucknow (India)


    The genus Umkomasia Thomas, a female fructification of Dicroidium, is reported from Late Permian beds exposed in the Behra Rivulet near the village of Karaonda in the Tatapani-Ramkola Coalfield, Chhattisgarh, India. This is the first record of this genus from the Late Permian, and from Indian Gondwana. Two species have been recognized: Umkomasia polycarpa Holmes and U. uniramia Axsmith, Taylor, Taylor and Cuneo. The find suggests that the genus Dicroidium appeared in the Late Permian before reaching its acme in the Middle-Upper Triassic.




    Full Text Available The systematics of lingulide brachiopods, from the end-Permian mass extinction interval, is here studied and discussed. The material has been collected from upper Permian (Changhsingian beds of Southern Alps and Lower Triassic beds of several Tethyan localities, where the surviving phase following the peak the end-Permian mass extinction is recorded. The study contributes to fill the gap of knowledge regarding the lingulide systematics during a time lapse crucial for the fate of the Mesozoic and Cenozoic marine organisms. The systematics is based both on inner shell morphology and shell microstructure, which are considered to be the most useful taxonomical characters to study the lingulide phylogeny. The specimens have been referred to species of the new genus Trentingula, which is characterized by a shell with a secondary layer virgose fabric and a primitive disposition of the ventral muscle umbonal scar in the Lingulidae phylogeny. Trentingula n. gen. comprises four species: T. lorigae n. gen. n. sp., type-species, T. borealis (Bittner, T. mazzinensis n. gen. n. sp., and T. prinothi n. gen. n. sp. The type-species is late Griesbachian – Dienerian in age and has a wide geographic distribution in the western Tethys (Southern Alps and Hungary. Trentingula prinothi n. gen. n. sp. occurs in the Upper Permian Bellerophon Formation of the Dolomites; it has a large shell with a short mantle cavity. Trentingula mazzinensis n. gen. n. sp. occurs in the Griesbachian Mazzin Member of Werfen Formation and is characterized by a small sized shell, about half of the type species, which records the “Lilliput effect” related to the aftermath of the end-Permian mass extinction.

  7. Geological event sequences of the Permian-Triassic transition recorded in the microfacies in Meishan section

    CAO ChangQun; ZHENG QuanFeng


    The microfacies sequence in the key interval of Beds 24-29 of the Meishan section comprehensively recorded the geological events during the Permian-Triassic transition, including the anoxia, storm disturbance, hard-ground/firm-ground form, volcanic eruption, weathering input, and microbialite de-velopment. This investigation of event sequences on the microfacies provided synthetically some clues to clarifying the previously proposed mechanisms of the end-Permian extinction. The deposit succession in Bed 24 of the Changxing Formation was developed dominantly with organic-rich lamina, but interrupted intermittently by storm disturbances. It indicates characteristically a stagnated, strati-fled and anoxic ocean during the latest Permian. The latest Permian transgression occurred rapidly at the top of Bed 24d and was characterized by the hard-ground interface. Abundant clay mineral of illite indicates the occurrence of persistent weathering of continental volcanics and then buried in ocean throughout this key Permo-Triassic transitional interval. However, the "boundary ash clay" of Bed 25 yielding normal grading of hyaloclastite was the result of precipitation of volcanic dusts. The firm-ground substrate in Unit 27-2 was lithe and excavated intensively by trace fossil Glossifungites; it means also the occurrence of the earliest Triassic rapid transgression and coincides with the first ap-pearance of conodont Hindeodus parvus. Dune-shaped microbialite in Unit 27-5 defines biostrati-graphically that its development, especially in the carbonate platform of South China, should corre-spond to the earliest Triassic; and it also indicates the final episode of the end-Permian extinction event.

  8. Geological event sequences of the Permian-Triassic transition recorded in the microfacies in Meishan section


    The microfacies sequence in the key interval of Beds 24―29 of the Meishan section comprehensively recorded the geological events during the Permian-Triassic transition, including the anoxia, storm disturbance, hard-ground/firm-ground form, volcanic eruption, weathering input, and microbialite development. This investigation of event sequences on the microfacies provided synthetically some clues to clarifying the previously proposed mechanisms of the end-Permian extinction. The deposit succession in Bed 24 of the Changxing Formation was developed dominantly with organic-rich lamina, but interrupted intermittently by storm disturbances. It indicates characteristically a stagnated, stratified and anoxic ocean during the latest Permian. The latest Permian transgression occurred rapidly at the top of Bed 24d and was characterized by the hard-ground interface. Abundant clay mineral of illite indicates the occurrence of persistent weathering of continental volcanics and then buried in ocean throughout this key Permo-Triassic transitional interval. However, the "boundary ash clay" of Bed 25 yielding normal grading of hyaloclastite was the result of precipitation of volcanic dusts. The firm-ground substrate in Unit 27-2 was lithe and excavated intensively by trace fossil Glossifungites; it means also the occurrence of the earliest Triassic rapid transgression and coincides with the first appearance of conodont Hindeodus parvus. Dune-shaped microbialite in Unit 27-5 defines biostrati-graphically that its development, especially in the carbonate platform of South China, should corre-spond to the earliest Triassic; and it also indicates the final episode of the end-Permian extinction event.

  9. Microbialites and global environmental change across the Permian-Triassic boundary: a synthesis.

    Kershaw, S; Crasquin, S; Li, Y; Collin, P-Y; Forel, M-B; Mu, X; Baud, A; Wang, Y; Xie, S; Maurer, F; Guo, L


    Permian-Triassic boundary microbialites (PTBMs) are thin (0.05-15 m) carbonates formed after the end-Permian mass extinction. They comprise Renalcis-group calcimicrobes, microbially mediated micrite, presumed inorganic micrite, calcite cement (some may be microbially influenced) and shelly faunas. PTBMs are abundant in low-latitude shallow-marine carbonate shelves in central Tethyan continents but are rare in higher latitudes, likely inhibited by clastic supply on Pangaea margins. PTBMs occupied broadly similar environments to Late Permian reefs in Tethys, but extended into deeper waters. Late Permian reefs are also rich in microbes (and cements), so post-extinction seawater carbonate saturation was likely similar to the Late Permian. However, PTBMs lack widespread abundant inorganic carbonate cement fans, so a previous interpretation that anoxic bicarbonate-rich water upwelled to rapidly increase carbonate saturation of shallow seawater, post-extinction, is problematic. Preliminary pyrite framboid evidence shows anoxia in PTBM facies, but interbedded shelly faunas indicate oxygenated water, perhaps there was short-term pulsing of normally saturated anoxic water from the oxygen-minimum zone to surface waters. In Tethys, PTBMs show geographic variations: (i) in south China, PTBMs are mostly thrombolites in open shelf settings, largely recrystallised, with remnant structure of Renalcis-group calcimicrobes; (ii) in south Turkey, in shallow waters, stromatolites and thrombolites, lacking calcimicrobes, are interbedded, likely depth-controlled; and (iii) in the Middle East, especially Iran, stromatolites and thrombolites (calcimicrobes uncommon) occur in different sites on open shelves, where controls are unclear. Thus, PTBMs were under more complex control than previously portrayed, with local facies control playing a significant role in their structure and composition.

  10. Uppermost Permian to Lower Triassic Conodont Zonation from Enshi area, western Hubei Province, South China

    Lyu, Z.; Zhao, L.; Chen, Z. Q.; Ma, D.; Yan, P.; Zhan, P.


    The Permian-Triassic transition witnessed the largest biotic turnover of Earth life during the Phanerozoic history. Ecosystems in sea and on land have also experienced the most protected restoration following the end-Permian mass extinction. These biocrises were also associated with climatic and environmental extremes through the latest Permian to Middle Triassic. In order to uncover the links among these extreme events, we need to establish high-resolution biochronostratigraphy, which offers precise timescales for reconstructing event sequences and probing the possible causes. Of these, conodont biostratigraphy is an operational tool in enhancing stratigraphic resolution. Although their ancestors and phylogeny remain unclear, conodonts are a rapid evolutionary lineage and extremely abundant in the Triassic marine carbonate successions. Here, we present recent study results of the Lower Triassic conodont zonation from the Ganxi and Jianshi areas, western Hubei Province, South China, which were situated on a carbonate ramp at the southern northern margin of the Upper Yangtze Platform. Therein, the uppermost Permian to Lower Triassic successions are well exposed and yield abundant conodonts. A total of nine conodont zones was established: (1) Clarkina yini-Clarkina zhangi Zone, (2) Hindeodus changxingensis Zone, (3) Hindeodus parvus Zone, (4) Isarcicella staeschei Zone, (5) Clarkina planata Zone, (6) Neoclarkina discrete Zone, (7) Neospathodus dieneri Zone, (8) Novispathodus waageni Zone, and (9) Triassospathodus homeri Zone. The Ns. dieneri M1, Ns. dieneri M2 and Ns. dieneri M3 subzones have also been distinguished from the Ns. dieneri Zone. Both Nv. waageni eowaageni subzones and Nv. waageni waageni subzones are also recognizable from the Nv. waageni Zone. The first occurrence of H. parvus marks the Permian-Triassic boundary(PTB), while the first occurrence of Nv. waageni eowaageni defines the Induan-Olenekian boundary. These conodont zones correlate well with

  11. An integrated biomarker, isotopic and palaeoenvironmental study through the Late Permian event at Lusitaniadalen, Spitsbergen

    Nabbefeld, Birgit; Grice, Kliti; Twitchett, Richard J.; Summons, Roger E.; Hays, Lindsay; Böttcher, Michael E.; Asif, Muhammad


    The largest extinction of the Phanerozoic occurred near the Permian/Triassic (P/Tr) boundary some 252 Ma ago. Several scenarios and drivers have been proposed for this event. Here we report for the first time an integrated study comprising sedimentological data, biomarker distributions/abundances and selected stable carbon and hydrogen isotopes along with bulk isotopes (δ 34S pyrite, δ 13C carb, δ 13C org) for a Late Permian section from Lusitaniadalen, Spitsbergen, Norway. Sedimentological and geochemical data support a marine transgression and collapse of the marine ecosystem in the Late Permian. Strong evidence for waxing and waning of photic zone euxinia throughout the Late Permian is provided by Chlorobiaceae-derived biomarkers (including δ 13C data) and δ 34S pyrite, implying multiple phases of H 2S outgassing and potentially several pulses of extinction. A rapid decrease in abundance of various land-plant biomarkers prior to the marine collapse event indicates a dramatic decline of land-plants during the Late Permian and/or increasing distance from palaeoshoreline as a consequence of sea level rise. Changes in δD of selected biomarkers also suggest a change in source of organic matter (OM) or sea level rise. We also found biomarker and isotopic evidence for a phytoplanktonic bloom triggered by eutrophication as a consequence of the marine collapse. Compound specific isotope analyses (CSIA) of algal and land-plant-derived biomarkers, as well as δ 13C of carbonate and bulk OM provide strong evidence for synchronous changes in δ 13C of marine and atmospheric CO 2, attributed to a 13C-depleted source. The source could be associated with isotopically depleted methane released from the melting of gas clathrates and/or from respired OM, due to collapse of the marine ecosystem.

  12. The Floral Response to the Permian Tectonic Evolution in Tarim Plate%塔里木板块二叠纪构造演化的古植物群(大植物、孢粉)响应



    The Tarim plate drifted constantly northward and collided with the Kazakhstan Plate during Permo-Carboniferous. In Carboniferous, the Tarim Plate was pieced together with the Yili Terrane of the Kazakhstan Plate. Then the Tarim-Yili Plate collided with the Kazakhstan Plate, and the Junggar Ocean finished subduction during Late Carboniferous and Early Permian. At last, the two plates were united into the southern part of the Angaran Land of the Pangea. The plate drifting resulted in the disappearance of the ocean between the Tarim Plate, the Kazakhstan Plate and the Junggar Plate. The plant distribution is mainly controlled by the climatic condition, and the geographic barriers also have an effect on the plant migration. The northward drift of the Tarim Plate led to the elevation of the plate and the cooling of climate there. As a result, the Euramerican flora of the plate was replaced by the exotic flora during the Permian. The northward drift of theTarim Plate is thought to be the principal impetus to the floral provincial succession of the plate. The Permian dry climate inthe northern hemisphere and the plant migration might have played positive roles in this succession. The whole floral succession of the Tarim Plate from the Euramerican stage to the Angara stage as described in this paper was recorded in the Permian deposits. The Permian floral succession of the Tarim Plate was controlled by the tectonic setting then and reflected the corresponding history of the plate drift. Three developing stages of the floral succession of the Tarim Plate have been recognized, viz.,①the Euramerican flora stage (Asselian-Roadian);②the Euramerican and Angara mixed flora stage (Wordian-Early Wuchiapingian);③the Angara flora stage (Middle-Late Wuchiapingian-Changhsingian).%塔里木板块二叠纪的构造演化导致板块古地理位置、古地貌和古环境的演变(包括气候条件的改变),相应地塔里木板块的植物群在区系性质方面发

  13. A comparison of the biological,geological events and environmental backgrounds between the Neoproterozoic-Cambrian and Permian-Triassic transitions


    The Neoproterozoic-Cambrian(N-C) and Permian-Triassic(P-T) transitions have been regarded the two most critical transitions in earth history because of the explosive biological radiation in the early Cambrian(the Cambrian Explosion) and the largest mass extinction at the end-Permian.Previous studies suggest that these two critical transitions showed certain comparability in major evolutionary events.In other words,a series of biological,geological,and geochemical events that had happened in the N-C transition occurred repeatedly during the P-T transition.Those events included continental re-configuration related to the deep mantle dynamics,global-scale glaciations,large C-,Sr-,and S-isotope perturbations indicating atmospheric and oceanic changes,abnormal precipitation of carbonates,and associated multiple biological radiations and mass extinctions.The coupling of those events in both N-C and P-T transitions suggests that deep mantle dynamics could be a primary mechanism driving dramatic changes of environment on the earth’s surface,which in turn caused major biological re-organizations.A detailed comparison of those events during the two critical transitions indicates that despite their general comparability,significant differences do exist in magnitude,duration,and frequency.The supercontinent Rodinia began to rift before the Snowball Earth time.By contrast,the supercontinent Pangea entered the dispersal stage after the greatest glaciation from the Late Carboniferous to Cisuralian.Quantitative data and qualitative analyses of different fossil groups show a more profound mass extinction during the N-C transition than at the end-Permian in terms of ecosystem disruption.This is indicated by the disappearance of the whole Ediacaran biota at the N-C boundary.The subsequent appearances of many new complex animals at phylum level in the early Cambrian mark the establishment of a brand new ecosystem.However,the end-Permian mass extinction is manifested mainly by the

  14. Sedimentary Environments and Geochemical Characteristics of Early Permian Carbonate Platform in the Western Henan Province%豫西地区早二叠世碳酸盐岩台地沉积环境及地球化学特征

    姚旭; 周瑶琪; 李素


    that of LREE /HREE and Fe.Compared with open carbonate platform,restricted carbonate platform (mainly in Jiaozuo area)is featured by the widespread micrite,which infer that the ancient ocean in the restricted carbonate platform contained less fossils and colloids,and became much more restricted and quieter than that of open carbonate platform.The bottom limestones of the Lower Limestone Section of Taiyuan Forma-tion,which is formed in transgressive systems tract(TST)during the early phase of first transgression,are fea-tured by high content of biodetritus,Sc,Ba and ∑REE,low value of Sr/Ba,low differentiation between LREE and HREE.The top limestones of the Lower Limestone Section formed in highstand systems tract (HST), which reflected the sedimentary environment of the limited carbonate platform in the research area evolving from the previous turbulent water into confined and quite surroundings with biodetritus and collides diminishing and evaporation strengthening.It aims to put implications on the research of the characteristics and evolution by comparing with the sedimentary characteristics of Taiyuan Formation limestones in different stage of the system tracts in restricted carbonate platform.

  15. Evaluating transition-metal catalysis in gas generation from the Permian Kupferschiefer by hydrous pyrolysis

    Lewan, M.D.; Kotarba, M.J.; Wieclaw, D.; Piestrzynski, A.


    Transition metals in source rocks have been advocated as catalysts in determining extent, composition, and timing of natural gas generation (Mango, F. D. (1996) Transition metal catalysis in the generation of natural gas. Org. Geochem.24, 977–984). This controversial hypothesis may have important implications concerning gas generation in unconventional shale-gas accumulations. Although experiments have been conducted to test the metal-catalysis hypothesis, their approach and results remain equivocal in evaluating natural assemblages of transition metals and organic matter in shale. The Permian Kupferschiefer of Poland offers an excellent opportunity to test the hypothesis with immature to marginally mature shale rich in both transition metals and organic matter. Twelve subsurface samples containing similar Type-II kerogen with different amounts and types of transition metals were subjected to hydrous pyrolysis at 330° and 355 °C for 72 h. The gases generated in these experiments were quantitatively collected and analyzed for molecular composition and stable isotopes. Expelled immiscible oils, reacted waters, and spent rock were also quantitatively collected. The results show that transition metals have no effect on methane yields or enrichment. δ13C values of generated methane, ethane, propane and butanes show no systematic changes with increasing transition metals. The potential for transition metals to enhance gas generation and oil cracking was examined by looking at the ratio of the generated hydrocarbon gases to generated expelled immiscible oil (i.e., GOR), which showed no systematic change with increasing transition metals. Assuming maximum yields at 355 °C for 72 h and first-order reaction rates, pseudo-rate constants for methane generation at 330 °C were calculated. These rate constants showed no increase with increasing transition metals. The lack of a significant catalytic effect of transition metals on the extent, composition, and timing of

  16. Molecular carbon isotope variations in core samples taken at the Permian-Triassic boundary layers in southern China

    Wang, Ruiliang; Zhang, Shuichang; Brassell, Simon; Wang, Jiaxue; Lu, Zhengyuan; Ming, Qingzhong; Wang, Xiaomei; Bian, Lizeng


    Stable carbon isotope composition (δ13C) of carbonate sediments and the molecular (biomarker) characteristics of a continuous Permian-Triassic (PT) layer in southern China were studied to obtain geochemical signals of global change at the Permian-Triassic boundary (PTB). Carbonate carbon isotope values shifted toward positive before the end of the Permian period and then shifted negative above the PTB into the Triassic period. Molecular carbon isotope values of biomarkers followed the same trend at and below the PTB and remained negative in the Triassic layer. These biomarkers were acyclic isoprenoids, ranging from C15 to C40, steranes (C27 dominates) and terpenoids that were all significantly more abundant in samples from the Permian layer than those from the Triassic layer. The Triassic layer was distinguished by the dominance of higher molecular weight (waxy) n-alkanes. Stable carbon isotope values of individual components, including n-alkanes and acyclic isoprenoids such as phytane, isop-C25, and squalane, are depleted in δ13C by up to 8-10‰ in the Triassic samples as compared to the Permian. Measured molecular and isotopic variations of organic matter in the PT layers support the generally accepted view of Permian oceanic stagnation followed by a massive upwelling of toxic deep waters at the PTB. A series of large-scale (global) outgassing events may be associated with the carbon isotope shift we measured. This is also consistent with the lithological evidence we observed of white thin-clay layers in this region. Our findings, in context with a generally accepted stagnant Permian ocean, followed by massive upwelling of toxic deep waters might be the major causes of the largest global mass extinction event that occurred at the Permian-Triassic boundary.

  17. Global Taxonomic Diversity of Anomodonts (Tetrapoda, Therapsida) and the Terrestrial Rock Record Across the Permian-Triassic Boundary

    Jörg Fröbisch


    The end-Permian biotic crisis (~252.5 Ma) represents the most severe extinction event in Earth's history. This paper investigates diversity patterns in Anomodontia, an extinct group of therapsid synapsids ('mammal-like reptiles'), through time and in particular across this event. As herbivores and the dominant terrestrial tetrapods of their time, anomodonts play a central role in assessing the impact of the end-Permian extinction on terrestrial ecosystems. Taxonomic diversity analysis reveals...

  18. Shallow marine ecosystem feedback to the Permian/Triassic mass extinction

    Yongbiao WANG; Zheng MENG; Wei LIAO; Zeting WENG; Hao YANG


    Late Permian reefs developed widely on shallow marine carbonate platforms in South China but disappeared far below the main mass extinction level of the latest Permian. The collapse of reef ecosystem may be related to the enhanced volcanism at the end of Late Permian. Notably, some colony corals and reef-building sponges were found to occur near the mass extinction boundary, inferring the eclipse of reef ecosystem is ahead of the disappearance of reef-building organisms, and the triggers would be present long before the main mass extinction. As the primary producers, the calcareous algae are rich in platform limestones of Late Permian and played a very important role in maintaining the shallow benthic ecosystems. The calcareous algae were found to disappear synchronously with the great reduction of foraminifers,which were ecologically associated with these algae. The extinction of Late Permian calcareous algae greatly reduced the biodiversity of primary producers in the shallow marine environment and destroyed in part the structure and the base of the shallow marine ecosystems,which in turn cause the extinction of ecologically associated metazoan. Microbialites developed on carbonate platforms immediately after the end-Permian mass extinction, representing a simple and unique microbial ecosystem. Widespread occurrence of microbialites symbolized the deterioration of marine environmental conditions and the dramatic revolution of marine ecosystems. As the new primary producers instead of the extinguished calcareous algae, cyanobacteria in the microbialites were an important base of this peculiar ecosystem and contributed greatly to the survival of the remnant faunas after the mass extinction. Widespread occurrence of microbialites in shallow marine environment is suggested to be related to the elevated level of volcanism-induced greenhouse gases and enhanced evaporation and hypersaline condition in addition to the decrease of metazoan grazing pressure. The change

  19. Guanling Biota: A Marker of Triassic Biotic Recovery from the end-Permian Extinction in the Ancient Guizhou Sea

    JIANG Dayong; Ryosuke MOTANI; LI Chun; HAO Weicheng; SUN Yuanlin; SUN Zuoyu; Lars SCHMITZ


    After a slow recovery from the end-Permian extinction during the Early Triassic and rapid radiation in the Middle Triassic, evolution of organisms reached a new peak phase in the Late Triassic. The Guanling Biota from the Wayao Member (conodont Paragondolella polygnathiformis Zone), Falang Formation, Xinpu, Guanling County, Guizhou Province, southwestern China corresponds to this peak that marks the full recovery from the end-Permian extinction of marine ecosystems. The biota is of high diversity, containing well preserved and completely articulated skeletons of vertebrates comprising marine reptiles, fishes, and invertebrates including crinoids, ammonites, bivalves, and other fossils,and is one of the best examples of marine ecosystem records in life history. The fossil marine reptiles and crinoids are most significant in this biota, especially the marine reptiles, which provide an important link between the Triassic Pacific and Tethys, and between Triassic basal forms and the Jurassic-Cretaceous marine top predators. The most remarkable fossils are the large completely articulated ichthyosaur skeletons up to and more than 10 m, and the first recorded thalattosaurs and placodonts in China. Following our review, of the 17 named reptilian taxa the eight listed here are considered to be valid: three ichthyosaurs (Qianichthyosaurus zhoui Li, 1999; Guizhouichthyosaurus tangae Cao and Luo in Yin et al.,2000, Guanlingsaurus liangae Yin in Yin et al., 2000), three thalattosaurs (Anshunsaurus huangguoshuensis Liu, 1999,Xinpusaurus suni Yin in Yin et al., 2000, Xinpusaurus kohi Jiang et al., 2004), and two placodonts (Sinocyamodus xinpuensis Li, 2000, Psephochelys polyosteoderma Li and Rieppel, 2002). Mixosaurus guanlingensis Cao in Yin et al.,2000 might be a junior synonym of Qianichthyosaurus zhoui Li, 1999, and Cymbospondylus asiaticus Li and You, 2002and Panjiangsaurus epicharis Chen and Cheng, 2003 might be junior synonyms of Guizhouichthyosaurus tangae Cao and Luo in

  20. Major factors controlling fracture development in the Middle Permian Lucaogou Formation tight oil reservoir, Junggar Basin, NW China

    Zhang, Chen; Zhu, Deyu; Luo, Qun; Liu, Luofu; Liu, Dongdong; Yan, Lin; Zhang, Yunzhao


    Natural fractures in seven wells from the Middle Permian Lucaogou Formation in the Junggar Basin were evaluated in light of regional structural evolution, tight reservoir geochemistry (including TOC and mineral composition), carbon and oxygen isotopes of calcite-filled fractures, and acoustic emission (AE). Factors controlling the development of natural fractures were analyzed using qualitative and/or semi-quantitative techniques, with results showing that tectonic factors are the primary control on fracture development in the Middle Permian Lucaogou Formation of the Junggar Basin. Analyses of calcite, dolomite, and TOC show positive correlations with the number of fractures, while deltaic lithofacies appear to be the most favorable for fracture development. Mineral content was found to be a major control on tectonic fracture development, while TOC content and sedimentary facies mainly control bedding fractures. Carbon and oxygen isotopes vary greatly in calcite-filled fractures (δ13C ranges from 0.87‰ to 7.98‰, while δ18O ranges from -12.63‰ to -5.65‰), indicating that fracture development increases with intensified tectonic activity or enhanced diagenetic alteration. By analyzing the cross-cutting relationships of fractures in core, as well as four Kaiser Effect points in the acoustic emission curve, we observed four stages of tectonic fracture development. First-stage fractures are extensional, and were generated in the late Triassic, with calcite fracture fills formed between 36.51 °C and 56.89 °C. Second-stage fractures are shear fractures caused by extrusion stress from the southwest to the northeast, generated by the rapid uplift of the Tianshan in the Middle and Late Jurassic; calcite fracture fills formed between 62.91 °C and 69.88 °C. Third-stage fractures are NNW-trending shear fractures that resulted from north-south extrusion and thrusting in a foreland depression along the front of the Early Cretaceous Bogda Mountains. Calcite fracture

  1. Variable sediment flux in generation of Permian subduction-related mafic intrusions from the Yanbian region, NE China

    Guo, Feng; Li, Hongxia; Fan, Weiming; Li, Jingyan; Zhao, Liang; Huang, Miwei


    This paper presents petrology, mineralogy, zircon U-Pb ages, and whole-rock major, trace element and Sr-Nd-Hf isotopic compositions of four Permian (273-253 Ma) subduction-related mafic intrusions (including the Qinggoushan and Qianshan gabbros, and the Wangqing and Shuguang diorites) from the Yanbian region, NE China, with aims to understand the role of subducted sediment flux in generation of arc mafic cumulates. These intrusions have mineral assemblages crystallized in water-saturated parental magmas and show variable degrees of crystal accumulation as observed in mafic cumulates in subduction zones. Mass-balance consideration indicates that their parental magmas were calc-alkaline with arc-type trace element features (enrichments in large ion lithophile elements (LILE) and light rare earth elements (LREE) and depletions in Nb-Ta). They also have Sr-Nd-Hf isotopic compositions, i.e., 87Sr/86Sr(i) = 0.7029-0.7047, εNd(t) = + 0.9 ~ + 6.8, εHf(t) = + 5.6 ~ + 14.6, similar to modern arc basalts. The parental magmas were likely derived from a mantle wedge variably metasomatized by sediment melt and fluid from the subducting paleo-Asian Oceanic slab. Combined trace elemental and isotopic modeling results suggest that the parental magma of Qinggoushan gabbro was formed through 5-20% melting of the mantle wedge with 1% and 1.5% additions of sediment fluid and sediment melt, respectively; 5-10% melting of the mantle wedge through inputs of 1% sediment fluid and 2% sediment melt produced the Qianshan gabbro; 10-20% melting of the mantle wedge with additions of 1% sediment fluid and 3% sediment melt formed the Wangqing diorite; whereas 5-20% melting of the mantle wedge through an input of 1.5% sediment melt produced the Shuguang diorite. The Hf-Nd isotopic array of the Yanbian Permian mafic intrusions reflected the existence of an Indian Ocean-type mantle, which was isotopically distinct from the Pacific-type mantle during early Paleozoic in the Central Asian Orogenic

  2. Thermal and cementation histories of Permian shelf-edge carbonate rocks in the Nanpanjiang Basin, South China

    Halley, Robert B.; Parrish, Judith T.; Zahn, Xie; Wenhai, Hu; Scholle, Peter A.; Zhongrui, Su; Yirong, Zhang; Yunming, Huang; Guangxuan, Li


    As of 1984, the Nanpanjiang Basin of South China has had almost no exploration by drilling although oil seeps exist among its margins and producing wells occur in adjacent basins.  however, cooperative studies by petroleum geologists and geochemists of the United States and the People's Republic of China (1982-1984) show that calcite-cemented reef and fore-reef carbonate rocks near Ziyun contain bitumen in Upper Permian shelf-edge sediment.  the cementation history consists of three episodes: (1)precipitation of syndepositional marine cement (formerly botryoidal aragonnite and fibrous magnesian calcite); (2)precipitation of post-depositional early cement (radiaxial calcite); and (3)precipitation of late burial cement (white calcite spar).  Hydrocarbons were introduced into the rocks between cementation episodes 2 and 3.  Fluid inclusion analysis of secondary inclusions in the burial cement indicates that the rocks were heated to nearly 200 C after hydrocarbon migration and cementation episode 3.  Bitumen remains in the rocks as evidence of the earlier presence of liquid hydrocarbons.

  3. Raining lead around 250mya a smoking gun for an Australian impact origin of the Permian Extinction

    Standard, J C


    Recent documentation of extreme atmospheric sulfur and methane contents at the time of the vast Permo-Triassic (P-T) extinction makes it possible to interpret an observation that has lain unnoticed in the geological literature for 40 years. This is the finding of microscopic metallic lead tear drops in the fluvial strata of the early Triassic sandstones that overlie Permian coal beds and other sedimentary deposits in the Sydney basin of Australia. Elemental lead is almost unknown in nature, so its occurrence in these graphite-loaded sandstones is a provocative finding. While climate change and vulcanism could explain the carbon and sulfur anomalies, the only way to account for metallic lead aerodynamic droplets is by massive impact and vaporization of lead mineral-containing formations. Since lead occurs geologically as the sulfide and since lead is an easily reduced element, its occurrence in conjunction with sulfur and carbon count anomalies suggests a bolide impact on carbon-loaded strata in a sulfide mine...

  4. Episodic euxinia in the Changhsingian (late Permian) of South China: Evidence from framboidal pyrite and geochemical data

    Wei, Hengye; Algeo, Thomas J.; Yu, Hao; Wang, Jiangguo; Guo, Chuan; Shi, Guo


    A multiproxy study of a new Upper Permian-Lower Triassic section (Xiaojiaba) in Sichuan Province, China, documents large changes in marine productivity, redox conditions and detrital input prior to the latest Permian mass extinction. Marine productivity, as proxied by total organic carbon content (TOC), biogenic SiO2, and excess barium, displays a long-term decline through most of the Changhsingian stage (late late Permian), culminating in very low values around the Permian-Triassic boundary. Concurrently, redox proxies including pyrite framboid, δ34Spy, Moauth and Uauth, and Corg/P document a shift from suboxic to dysoxic/oxic conditions that was interrupted by several episodes of benthic euxinia, and detrital siliciclastic proxies (Al, Hf, Nb, and REEs) suggest an increased flux of weathered material from land areas. The long-term changes in productivity, redox conditions, and terrigenous detrital fluxes were probably caused by a regional sea-level fall across the South China Craton. On the other hand, the brief euxinic episodes occurring during the late Permian had oceanographic causes, probably related to the transient upward expansion of the chemocline at the top of the oceanic oxygen-minimum zone. These euxinic episodes may have been harbingers of the more widespread anoxia that developed concurrently with the latest Permian mass extinction and that may have played a major role in triggering the largest biotic crisis of the Phanerozoic.

  5. Biostratigraphic correlation and mass extinction during the Permian-Triassic transition in terrestrial-marine siliciclastic settings of South China

    Chu, Daoliang; Yu, Jianxin; Tong, Jinnan; Benton, Michael J.; Song, Haijun; Huang, Yunfei; Song, Ting; Tian, Li


    The Permian-Triassic boundary marks the greatest mass extinction during the Phanerozoic, which was coupled with major global environmental changes, and is known especially from well-preserved marine fossil records and continuous carbonate deposits. However, the placement of the Permian-Triassic boundary in terrestrial sections and accurate correlation with the marine strata are difficult due to the absence of the key marine index fossils in terrestrial-marine siliciclastic settings. Here, we present detailed fossil data from four terrestrial sections, two paralic sections and one shallow marine section in South China. Our data show that the rapid mass disappearance of the Gigantopteris flora in various sections represents the end-Permian mass extinction and the base of the Permian-Triassic transitional beds in terrestrial-marine siliciclastic settings of South China. In particular, we find a mixed marine and terrestrial biota from the coastal transitional sections of the Permian-Triassic transitional Kayitou Formation, which provides a unique intermediate link for biostratigraphic correlation between terrestrial and marine sequences. Accordingly, the Euestheria gutta-bearing conchostracan fauna and the Pteria ussurica variabilis-Towapteria scythica-Eumorphotis venetiana bivalve assemblage are proposed as markers of the Permian-Triassic transitional beds in terrestrial-marine siliciclastic settings of South China.

  6. Depositional model of Permian Luodianian volcanic island and its impact on the distribution of fusulinid assemblage in southern Qinghai, Northwest China

    NIU ZhiJun; XU AnWu; WANG JianXiong; DUAN QiFa; ZHAO XiaoMing; YAO HuaZhou


    resulting from calm shallow water was deposited at the interior of the Qamdo Block from the Devonian to early Early Permian. At the beginning of the peak period of activity of pan-riftzation (Luodianian), alternate volcanic island and shallow marine environment within continent crust came into being. Uniform and stable shallow-water carbonate platform was formed during the Xiangboan. This suggested that the activity of rift basin was evidently weakened. Subsequently the instability of the basin appreciably increased with the occurrence of basalt in late Kuhfengian. At last the whole Qamdo Block turned into the closure period of rift during the Late Permian.

  7. Depositional model of Permian Luodianian volcanic island and its impact on the distribution of fusulinid assemblage in southern Qinghai,Northwest China


    sequence re-sulting from calm shallow water was deposited at the interior of the Qamdo Block from the Devonian to early Early Permian. At the beginning of the peak period of activity of pan-riftzation (Luodianian), al-ternate volcanic island and shallow marine environment within continent crust came into being. Uni-form and stable shallow-water carbonate platform was formed during the Xiangboan. This suggested that the activity of rift basin was evidently weakened. Subsequently the instability of the basin appre-ciably increased with the occurrence of basalt in late Kuhfengian. At last the whole Qamdo Block turned into the closure period of rift during the Late Permian.

  8. Palaeomagnetic studies in the Permian Basin of Largentière and implications for the Late Variscan rotations in the French Massif Central

    Henry, B.; Becq-Giraudon, J. F.; Rouvier, H.


    Detailed geological observations and palaeomagnetic analyses were carried out in the Largentière Stephano-Autunian basin and on the Stephanian deposits of the Alès coalfield, both located at the southeastern margin of the French Massif Central. Because of unfavourable rock types, the Alès Stephanian deposits did not yield any results. The palaeomagnetic pole (164.9 degE, 45.4 degN, K=89, A_95=4.1 deg) deduced from a study of the Autunian sediments of the Largentière Basin agrees very well with the reference pole for stable Europe. The Lodève-Largentière area, that is the southeastern border of the Massif Central, has been stable since Early Permian time with respect to stable Europe, whereas the western part (the Saint-Affrique Rodez Basin and, probably, the Brive Basin) has been rotated counterclockwise.

  9. Paleomagnetic investigation of the Early Permian Panjal Traps of NW India; regional tectonic implications

    Stojanovic, Denis; Aitchison, Jonathan C.; Ali, Jason R.; Ahmad, Talat; Dar, Reyaz Ahmad


    The ∼289 Ma Panjal Traps of NW India (Kashmir) are part of a series of rift-related mafic suites (Abor, Sikkim etc.) that were erupted onto northern India (present-day coordinates) around the same time as separation of the Cimmerian blocks of Qiangtang and Sibumasu. We report new data from only the second paleomagnetic investigation of this unit. Standard alternating field and thermal demagnetization methods were used to isolate characteristic magnetizations from seven outcrops at three locations within the Kashmir Valley, NW India. Analysis of four sections (14 individual cooling units) from close to Srinagar, that together form a tectonically coherent sequence spanning ∼3 km of stratigraphy, yield a single-component, primary magnetization with a mean direction of Dec: 134.8°, Inc: 55.3° (α95 = 8.9°, k = 21.0). An inclination-only mean of 52.5° (α95 = 8.9°, k = 47.2) gives a paleolatitude of ∼33°S (±5°). A paleopole of 110.5°E 8.4°S (A95 = 10.7) is also calculated. Assuming the magnetization records a portion of the reverse polarity Kiaman Superchron, the new result indicates extrusion of the Panjal Traps basalts at mid-latitudes in the southern hemisphere. By inference this constrains the location of central Gondwana, and informs debates related to Cimmeria's detachment from Gondwana.

  10. Early Permian conodont fauna and stratigraphy of the Garden Valley Formation, Eureka County, Nevada

    Wardlaw, Bruce R.; Gallegos, Dora M.; Chernykh, Valery V.; Snyder, Walter S.


    The lower Part of the Garden Valley Formation yields two distinct conodont faunas. One of late Asselian age dominated by Mesogondolella and Streptognathodus and one of Artinskian age dominated by Sweetognathus with Mesogondolella. The Asselian fauna contains the same species as those found in the type area of the Asselian in the southern Urals including Mesogondolella dentiseparata, described for the first time outside of the Urals. Apparatuses for Sweetognathus whitei, Diplognathodus stevensi, and Idioprioniodus sp. are described. The Garden Valley Formation represents a marine pro-delta basin and platform, and marine and shore fan delta complex deposition. The fan-delta complex was most likely deposited from late Artinskian to lateWordian. The Garden Valley Formation records tremendous swings in depositional setting from shallow-water to basin to shore.

  11. Geochemical significance of the relative enrichment of pristane and the negative excursion of δ13CPr across the Permian-Triassic Boundary at Meishan, China

    WANG Chunjiang; LIU Yimei; LIU Hongxiu; ZHU Lei; SHI Quan


    Changes in the distribution of isoprenoid hydrocarbons and in δ13Ckero or δ13CPr across the section from the Changxing Formation to the lower Yinkeng Formation at Meishan (the GSSP of PTB) are studied, in which a relative enrichment of pristane and a sharp shift in both δ13Ckero and δ13Cmole near the boundary are revealed. A possible mechanism for the abnormal accumulation of pristane near PTB is suggested: Lipids in some marine animals and their metabolites, rich in pristane, can be preserved and accumulate in some suitable depositing facies under certain special conditions, such as the catastrophic death of animals and/or developing of an anoxic water column, and thus the pristane enrichment occurs in the sediment. The occurrences of isoprenoid enrichment and high Pr/Ph ratios have been found in other typical marine anoxic sediments accompanied by mass extinction in various extent during the most important geological periods, such as Late Ordovician, end of Devonian, Early Jurassic and Late Cretaceous. These occurrences support to a certain extent the above mechanism that interprets the genesis of a high Pr/Ph ratio in some typical marine anoxic sediments. The synchronous sharp negative shifts in δ13Cmole, δ13Ckero and δ13Ccarb from Beds 24 to 26 of the Meishan profile indicate that a sudden and strong variation of carbon source in the atmosphere-ocean system occurred during the quick but equable evolution from the end-Permian to Early Triassic, and the coupling between the pristane enrichment and negative shift in δ13Ckero and δ13CPr may well indicate the importance of the release of methane hydrates and marine anoxia in the end-Permian mass extinction.

  12. Paleoenvironmental reconstitution of Motuca and Sambaíba formations, Permian-Triassic of the Parnaíba Basin in southwest Maranhão state, Brazil

    Francisco Romério Abrantes Júnior


    Full Text Available The interval between the Late Paleozoic and Early Mesozoic was marked by paleogeographic and paleoclimatic global changes, partly attributed to catastrophic events. The intense continentalization of the supercontinent Pangaea of End-Permian propitiated the development of extensive deserts that succeeded the coastal and platform environments of Early Permian. The records of these events in northern Brazil are found in intracratonic basins, particularly in the Permo-triassic succession of the Parnaíba Basin. The facies and stratigraphic outcrops analysis of this succession allowed the individualization of 14 sedimentary facies grouped into four facies associations (FA: FA1 and FA2 related to deposits of Motuca Formation and, FA3 and FA4, representative of the base of Sambaíba Formation. The FA1 – Shallow lake/Mudflat consists of red laminated mudstone with lenses of gypsum, calcite and marl, besides lobes of sigmoidal sandstones. The FA2 – Saline pan consists of lenticular bodies of laminated gypsum, nodular gypsum and gypsarenite, overlapped by greenish mudstones with dolomite nodules and palygorskite. The FA3 – sand sheet and FA4 – dunes field are formed, respectively, for orange cream sandstones with even parallel stratification and medium- to large-scale cross-bedding. In the contact between Motuca and Sambaíba formations occurs a deformed interval, laterally continuous for hundreds of kilometers. Brecciated and contorted bedded siltstones and mudstone (Motuca Formation and sandstone with sinsedimentary faults/microfaults, convolute lamination and mud-filled injection dykes (Sambaíba Formation are interpreted as seismites triggered by high magnitude earthquakes (> 8 according Richter scale.

  13. The End-Permian mass extinction: What really happened and did it matter?

    Erwin, D H


    Marine communities of the Paleozoic differ markedly from those of the post-Paleozoic, a dichotomy long recognized as the most fundamental change between the Cambrian metazoan radiation and the present. The end-Permian mass extinction of about 54% of marine families eliminated many of the groups that dominated Paleozoic communities. Correlative changes occurred in terrestrial vertebrate and plant communities, but there is no clear evidence that these changes are related to the marine extinction. The marine extinction occurred during a period of physical change, and a variety of extinction mechanisms have been proposed, most related to a major Late Permian marine regression or to climatic changes. Unfortunately, the regression has made it difficult to gather data on the rate, timing and pattern of extinction, and the available data exclude only a few hypotheses. Thus the largest mass extinction, and the one with the greatest evolutionary importance, is also the most poorly understood.

  14. Multiple S-isotopic evidence for episodic shoaling of anoxic water during Late Permian mass extinction.

    Shen, Yanan; Farquhar, James; Zhang, Hua; Masterson, Andrew; Zhang, Tonggang; Wing, Boswell A


    Global fossil data show that profound biodiversity loss preceded the final catastrophe that killed nearly 90% marine species on a global scale at the end of the Permian. Many hypotheses have been proposed to explain this extinction and yet still remain greatly debated. Here, we report analyses of all four sulphur isotopes ((32)S, (33)S, (34)S and (36)S) for pyrites in sedimentary rocks from the Meishan section in South China. We observe a sulphur isotope signal (negative δ(34)S with negative Δ(33)S) that may have resulted from limitation of sulphate supply, which may be linked to a near shutdown of bioturbation during shoaling of anoxic water. These results indicate that episodic shoaling of anoxic water may have contributed to the profound biodiversity crisis before the final catastrophe. Our data suggest a prolonged deterioration of oceanic environments during the Late Permian mass extinction.

  15. Episodic perturbations of end-Permian atmosphere recorded in plant spore chemistry

    Fraser, Wesley; Lomax, Barry; Beerling, David; James, David; Pyle, John; Self, Stephen; Sephton, Mark; Wellman, Charles


    The largest marine Phanerozoic extinction occurred 251 million years ago at the end of the Permian period with a contemporaneous major reorganisation of terrestrial. Previous work suggests the eruption of the Siberian Traps large igneous province could have generated substantial volumes of ozone depleting substances; the result being a partial collapse of the stratospheric ozone layer, and commensurate increase in ultraviolet-B (UV-B, 280-315nm) radiation. Increased UV-B flux would contribute additional pressures to an already stressed environment and flora and fauna. Here we present data utilising a new biogeochemical proxy for UV-B radiation to analyse clubmoss (lycophyta) megaspores to track UV-B radiation across the end-Permian interval. Our biogeochemical data when combined with published work on spore and pollen mutations suggests a highly dynamic global atmospheric system, oscillating between episodes of high and low UV-B flux, most likely driven by pulsed eruptive phases of the Siberian Traps.

  16. Study on REEs as tracers for late permian coal measures in Bijie City, Guizhou Province, China

    WANG Qiang; YANG Ruidong


    Analyses of Rare Earth Elements (REEs) in 13 coal samples collected from Late Permian coal measures of Bijie City in western Guizhon Province were conducted using Inductively Coupled Plasma-Mass-Spectrometry (ICP-MS). The results indicated that REEs patterns were not controlled by materials from the sea, whereas the contribution of land plants was about 1%. The major sources of REEs were from terrigenous material as indicated by negative Eu anomaly. There were similar distribution curves of REEs between Bijie's coal and Emeishan basalt. M12 coal seam, which had the highest ∑REE, appeared near the boundary between Longtan Formation and Changxing Formation, which was closely correlated to the eruption of Emeishan basalt. The Emeishan basalt contributed to REEs enrichment of M12. So the sources of REEs were controlled by terrigenous material, and the Emeishan basalt was the predominant source of terrigenous material, which dominated the enrichment and pattern of REEs in Late Permian coal measure from Bijie.

  17. A latest Permian non-reef calcisponge fauna from Laibin, Guangxi, southern China and its significance

    Ya-Sheng Wu


    Full Text Available A calcisponge fauna occurs in uppermost Permian Conodont Clarkina meishanensis yini zone of the sequence exposed in the vicinity of Laibin, Guangxi Zhuang Autonomous Region, southern China. The fauna is dominated by one thalamid species, Amblysiphonella vesiculosa de Koninck, 1863, and one new sclerosponge genus and species, Radiofibrosclera laibinensis gen. et sp. nov. They are associated with a few other accessory species, including the thalamid sponges Amblysiphonella laibinensis Deng, 1981, Colospongia sp., Polycystocoelia sp., and the inozoan sponge Acoelia discontinua sp. nov. Though the individuals are abundant, the species diversity is very low. Without common calcisponge components of Changhsingian reefal faunas, the assemblage is interpreted as not a reefal fauna. The water depth at which they dwelled was less than 105 m, and more probably less than 40 m. Its occurrence indicates a significant sea-level drop at the end of Late Permian Changhsingian Age.

  18. Carbon Capture and Storage in the Permian Basin, a Regional Technology Transfer and Training Program

    Rychel, Dwight


    The Permian Basin Carbon Capture, Utilization and Storage (CCUS) Training Center was one of seven regional centers formed in 2009 under the American Recovery and Reinvestment Act of 2009 and managed by the Department of Energy. Based in the Permian Basin, it is focused on the utilization of CO2 Enhanced Oil Recovery (EOR) projects for the long term storage of CO2 while producing a domestic oil and revenue stream. It delivers training to students, oil and gas professionals, regulators, environmental and academia through a robust web site, newsletter, tech alerts, webinars, self-paced online courses, one day workshops, and two day high level forums. While course material prominently features all aspects of the capture, transportation and EOR utilization of CO2, the audience focus is represented by its high level forums where selected graduate students with an interest in CCUS interact with Industry experts and in-house workshops for the regulatory community.

  19. Sandrewia, n. gen., a problematical plant from the Lower Permian of Texas and Kansas

    Mamay, S.H.


    Sandrewia, n. gen., monotypified by S. texana, n. sp., is a plant from Lower Permian beds of north-central Texas and east-central Kansas. It is characterized by stout axes with spirally disposed, laxly inserted, petiolate leaves; the laminae are broadly flabelliform with coarse, open venation. The leaves are reminiscent of the vojnovskyalean Nephropsis, of the Permian Petchora Basin, U.S.S.R., but biologic relationships are only speculative because of limited material. However, leaf characteristics render Sandrewia easily identifiable. Its presently limited stratigraphic range, along with floristic associations, indicates it may be a useful guide fossil and supports the author's beliefs regarding important times and places in Paleozoic plant evolution. ?? 1975.

  20. Evidence of Impact at the Permian/Triassic Boundary from Moessbauer Spectroscopy

    Verma, H. C.; Upadhyay, C. [Indian Institute of Technology, Department of Physics (India); Tripathi, R. P. [Jai Narain Vyas University, Department of Physics (India); Shukla, A. D.; Bhandari, N. [Physical Research Laboratory, Navrangpura (India)


    Sediment samples from Spiti valley, India, deposited about 250 Million years ago during the Permian-Triassic transition, have been analysed. {sup 57}Fe Moessbauer spectroscopic studies show that iron occurs mainly as nanoparticles of hematite and goethite with a broad size distribution. The results show a high degree of resemblance with the Cretaceous-Tertiary boundary (65 My) samples where an asteroidal impact is believed to have caused mass extinction.



    The transitional beds between Bellerophon and Werfen Formations, few centimetres thick and latest Permian in age, contain a brachiopod fauna with a relatively rich athyridoid assemblage, among which Janiceps is the most common and characteristic genus. The internal morphology of the type species (J. peracuta) is here described for the first time, and a taxonomical revision of the South Alpine species is proposed. The new subfamily Janicepsinae is proposed, which contains Janiceps and the new ...

  2. Volatile-organic molecular characterization of shale-oil produced water from the Permian Basin

    Khan, Naima A.; Engle, Mark A.; Dungan, Barry; Holguin, F. Omar; Xu, Pei; Carroll, Kenneth C.


    Growth in unconventional oil and gas has spurred concerns on environmental impact and interest in beneficial uses of produced water (PW), especially in arid regions such as the Permian Basin, the largest U.S. tight-oil producer. To evaluate environmental impact, treatment, and reuse potential, there is a need to characterize the compositional variability of PW. Although hydraulic fracturing has caused a significant increase in shale-oil production, there are no high-resolution organic composition data for the shale-oil PW from the Permian Basin or other shale-oil plays (Eagle Ford, Bakken, etc.). PW was collected from shale-oil wells in the Midland sub-basin of the Permian Basin. Molecular characterization was conducted using high-resolution solid phase micro extraction gas chromatography time-of-flight mass spectrometry. Approximately 1400 compounds were identified, and 327 compounds had a >70% library match. PW contained alkane, cyclohexane, cyclopentane, BTEX (benzene, toluene, ethylbenzene, and xylene), alkyl benzenes, propyl-benzene, and naphthalene. PW also contained heteroatomic compounds containing nitrogen, oxygen, and sulfur. 3D van Krevelen and double bond equivalence versus carbon number analyses were used to evaluate molecular variability. Source composition, as well as solubility, controlled the distribution of volatile compounds found in shale-oil PW. The salinity also increased with depth, ranging from 105 to 162 g/L total dissolved solids. These data fill a gap for shale-oil PW composition, the associated petroleomics plots provide a fingerprinting framework, and the results for the Permian shale-oil PW suggest that partial treatment of suspended solids and organics would support some beneficial uses such as onsite reuse and bio-energy production.

  3. Paleoecology of flora from coal measures of Upper Permian in western Guizhou

    Guo, Y. (China University of Mining and Technology (China))


    Based on paleobotany, palynology and sedimentology, the paper discusses the paleoecology of flora from coal measures of Upper Permian in Western Guizhou. The four major floras are: the swamp flora dominated by Lycopodiales, the water front floras dominated by Calamites, the floodplain flora dominated by Gigantopterides, Psaronics and the upland flora dominated by conifers, Ginkgos and Pteridospermus. The features of the different floras are described in detail. 6 refs., 3 figs.

  4. Volatile-organic molecular characterization of shale-oil produced water from the Permian Basin.

    Khan, Naima A; Engle, Mark; Dungan, Barry; Holguin, F Omar; Xu, Pei; Carroll, Kenneth C


    Growth in unconventional oil and gas has spurred concerns on environmental impact and interest in beneficial uses of produced water (PW), especially in arid regions such as the Permian Basin, the largest U.S. tight-oil producer. To evaluate environmental impact, treatment, and reuse potential, there is a need to characterize the compositional variability of PW. Although hydraulic fracturing has caused a significant increase in shale-oil production, there are no high-resolution organic composition data for the shale-oil PW from the Permian Basin or other shale-oil plays (Eagle Ford, Bakken, etc.). PW was collected from shale-oil wells in the Midland sub-basin of the Permian Basin. Molecular characterization was conducted using high-resolution solid phase micro extraction gas chromatography time-of-flight mass spectrometry. Approximately 1400 compounds were identified, and 327 compounds had a >70% library match. PW contained alkane, cyclohexane, cyclopentane, BTEX (benzene, toluene, ethylbenzene, and xylene), alkyl benzenes, propyl-benzene, and naphthalene. PW also contained heteroatomic compounds containing nitrogen, oxygen, and sulfur. 3D van Krevelen and double bond equivalence versus carbon number analyses were used to evaluate molecular variability. Source composition, as well as solubility, controlled the distribution of volatile compounds found in shale-oil PW. The salinity also increased with depth, ranging from 105 to 162 g/L total dissolved solids. These data fill a gap for shale-oil PW composition, the associated petroleomics plots provide a fingerprinting framework, and the results for the Permian shale-oil PW suggest that partial treatment of suspended solids and organics would support some beneficial uses such as onsite reuse and bio-energy production.


    Reinaldo Leopoldino Souza Neto


    at Day 0 (7.0±3.1mm; 8.1±1.9mm e 6.5±1.7mm and at Day 10 (9.8±1.4mm; 12.5±1.2mm e 11.8±2.8mm. The pregnancy rates at AI were 57, 48, 48 and 46% and at the end of the breeding season were 77, 88, 68 and 78%, EW-AI, EW-FTAI, EW-FTAI10 and TW-FTAI respectively, without significance (P>0.05. According to the this study, EW didn’t improve reproductive performance compared to TW associated to estrous synchronization programs to FTAI and 10 days of interval from EW and hormonal therapy didn’t increase fertility in FTAI.

    Key words: Beef cows, early weaning, fixed-time artificial insemination.
    O objetivo deste experimento foi avaliar a eficiência reprodutiva de vacas de corte com cria ao pé em programas de inseminação artificial (IA, utilizando práticas de desmame definitivo antecipado (DA ou temporário (DT e terapia de sincronização de estros para inseminação artificial a tempo fixo (IATF. Utilizaram-se 205 vacas Angus x Nelore, com condição corporal de 2,6±0,4 (escala de 1 a 5 e período pós-parto variando entre 54 e 122 dias. Após o DA, o número de 153 vacas foi dividido em três grupos de acordo com o protocolo utilizado para IA ou IATF. Submeteu-se um grupo de vacas ao controle de estros e IA por um período de trinta dias (DA-IA, n= 53, a partir de dez dias depois do DA. A sincronização de estros para a IATF foi realizada em dois grupos de vacas após o DA, diferindo apenas em relação ao momento do início dos tratamentos. No grupo DA-IATF (n= 50, o tratamento iniciou-se no dia do aparte dos terneiros, enquanto no grupo DA-IATF 10 (n= 50 os hormônios foram aplicados dez dias após o DA. A terapia hormonal consistiu da aplicação de um dispositivo intravaginal contendo 1,9 g de progesterona, por oito dias, e a aplicação de 2 mg de benzoato de estradiol, im. No momento da retirada do implante, foram aplicados 75 mcg de cloprostenol, im, e, 24 horas após, 1mg de benzoato de estradiol, im. Procedeu-se à IATF entre 52 a

  6. Jurassic cooling ages in Paleozoic to early Mesozoic granitoids of northeastern Patagonia: 40Ar/39Ar, 40K-40Ar mica and U-Pb zircon evidence

    Martínez Dopico, Carmen I.; Tohver, Eric; López de Luchi, Mónica G.; Wemmer, Klaus; Rapalini, Augusto E.; Cawood, Peter A.


    U-Pb SHRIMP zircon crystallization ages and Ar-Ar and K-Ar mica cooling ages for basement rocks of the Yaminué and Nahuel Niyeu areas in northeastern Patagonia are presented. Granitoids that cover the time span from Ordovician to Early Triassic constitute the main outcrops of the western sector of the Yaminué block. The southern Yaminué Metaigneous Complex comprises highly deformed Ordovician and Permian granitoids crosscut by undeformed leucogranite dikes (U-Pb SHRIMP zircon age of 254 ± 2 Ma). Mica separates from highly deformed granitoids from the southern sector yielded an Ar-Ar muscovite age of 182 ± 3 Ma and a K-Ar biotite age of 186 ± 2 Ma. Moderately to highly deformed Permian to Early Triassic granitoids made up the northern Yaminué Complex. The Late Permian to Early Triassic (U-Pb SHRIMP zircon age of 252 ± 6 Ma) Cabeza de Vaca Granite of the Yaminué block yielded Jurassic mica K-Ar cooling ages (198 ± 2, 191 ± 1, and 190 ± 2 Ma). At the boundary between the Yaminué and Nahuel Niyeu blocks, K-Ar muscovite ages of 188 ± 3 and 193 ± 5 Ma were calculated for the Flores Granite, whereas the Early Permian Navarrete granodiorite, located in the Nahuel Niyeu block, yielded a K-Ar biotite age of 274 ± 4 Ma. The Jurassic thermal history is not regionally uniform. In the supracrustal exposures of the Nahuel Niyeu block, the Early Permian granitoids of its western sector as well as other Permian plutons and Ordovician leucogranites located further east show no evidence of cooling age reset since mica ages suggest cooling in the wake of crystallization of these intrusive rocks. In contrast, deeper crustal levels are inferred for Permian-Early Triassic granitoids in the Yaminué block since cooling ages for these rocks are of Jurassic age (198-182 Ma). Jurassic resetting is contemporaneous with the massive Lower Jurassic Flores Granite, and the Marifil and Chon Aike volcanic provinces. This intraplate deformational pulse that affected northeastern

  7. Middle Permian palaeobiogeography study in East Kunlun, A'nyêmaqên and Bayan Har

    WANG; Yongbiao; YANG; Hao


    Three regions can easily be identified in the study area according to the Middle Permian palaeobiogeographic distribution of biota, they are the southern slope of East Kunlun, A'nyêmaqên and Bayan Har. Biotic constitution and ecology in the southern slope of East Kunlun and Bayan Har are very similar. Both the diversity and abundance of organisms in these two areas are very high and reefs are widely developed. However, biotic diversity and abundance in A'nyêmaqên which is between the above two areas are obviously low. Differentiation of palaeobiogeographic distribution in these areas should be due to the baring of A'nyêmaqên ocean in the time of Middle Permian. Middle Permian radiolarian chert and thick abyssal red ooze are widely spread in A'nyêmaqên, implying that the A'nyêmaqên ocean had a great scale in size. Vast scale of deep ocean basin became an impassable gulf for some of the benthos, and as a result, only part of the organisms could have the chance to get to the isolated islands situated in ocean basin. Small living space and hard conditions in the islands further limited the abundance and diversity of biota. Tectonic background reflected by the geochemical study of basalt in the three areas is coupling well enough with the palaeobiogeographic division.

  8. Modern Pearl River Delta and Permian Huainan coalfield, China: A comparative sedimentary facies study

    Suping, P.; Flores, R.M.


    Sedimentary facies types of the Pleistocene deposits of the Modern Pearl River Delta in Guangdong Province, China and Permian Member D deposits in Huainan coalfield in Anhui Province are exemplified by depositional facies of anastomosing fluvial systems. In both study areas, sand/sandstone and mud/mudstone-dominated facies types formed in diverging and converging, coeval fluvial channels laterally juxtaposed with floodplains containing ponds, lakes, and topogenous mires. The mires accumulated thin to thick peat/coal deposits that vary in vertical and lateral distribution between the two study areas. This difference is probably due to attendant sedimentary processes that affected the floodplain environments. The ancestral floodplains of the Modern Pearl River Delta were reworked by combined fluvial and tidal and estuarine processes. In contrast, the floodplains of the Permian Member D were mainly influenced by freshwater fluvial processes. In addition, the thick, laterally extensive coal zones of the Permian Member D may have formed in topogenous mires that developed on abandoned courses of anastomosing fluvial systems. This is typified by Seam 13-1, which is a blanket-like body that thickens to as much as 8 in but also splits into thinner beds. This seam overlies deposits of diverging and converging, coeval fluvial channels of the Sandstone D, and associated overbank-floodplain deposits. The limited areal extent of lenticular Pleistocene peat deposits of the Modern Pearl River Delta is due to their primary accumulation in topogenous mires in the central floodplains that were restricted by contemporaneous anastomosing channels.

  9. Late Permian carbonate concretions in the marine siliciclastic sediments of the Ravnefjeld Formation, East Greenland

    Nielsen, J.K. [Univ. of Copenhagen, Geological Inst., Copenhagen (Denmark); Hanken, N.M. [Univ. of Tromsoe, Dept. of Geology, Tromsoe (Norway)


    The investigation of carbonate concretions from the Late Permian Ravnefjeld Formation in East Greenland forms part of the multi-disciplinary research project Research of the Sedimentary Basins of North and East Greenland (TUPOLAR). The TUPOLAR project focuses on investigations and evaluation of potential hydrocarbon and mineral resources of the Upper Permian - Mesozoic sedimentary basins. In this context, the Upper Permian Ravnefjeld Formation occupies a pivotal position because it contains local mineralization and has source rock potential for hydrocarbons adjacent to potential carbonate reservoir rocks of the partly time-equivalent Wegener Halvoe Formation. A better understanding of the sedimentary facies and diagenesis of the Ravnefjeld Formation is therefore crucial for an evaluation of the economic evaluation of East Greenland. The original fieldwork was carried out in 1998, when sampling was undertaken of representative carbonate concretions and surrounding beds from a limited number of well-exposed sections in the Ravnefjeld Formation. The sampled material was subsequently investigated by a combination of petrography and stable isotope chemistry to decipher the relationships between the diagenetic development of the carbonate concretions and the mineralization in the sequence. The sequential precipitation of the cement generations was analysed in cement-filled primary voids in gastropods because these showed the most complete development of the different cement generations. The geochemistry of stable isotopes was also studied. During the petrographic work, we became aware of a hitherto unrecognised biota dominated by calcispheres. The well-developed cement generations in primary cavities in skeletal material were used to elucidate the diagenesis. (BA)

  10. Size variation of the end Permian conodont Neogondolella at Meishan Section, Changxing, Zhejiang and its significance


    This study is based on both a generic and species level investigation of the individual size of the latest Permian conodont Neogondolella Pa elements collected from Meishan Section A, Changxing, Zhejiang Province. In this study, an obvious size reduction of Neogondolella Pa elements within bed 24e of the upper Changxing Limestone is recognized. The size variation of the Neogondolella occurs simultaneously with some important events including the negative shift of δ13C, change in the ratio of the abundance of cyanobacterial biomarkers versus that of other general bacterial biomarkers and the shallowing of the sea water. Through the investigation of the paleoenvironmental changes and the analysis of the paleoecology of the conodont genus Neogondolella, the authors propose that the major factors for the size reduction of species of the conodont genus Neogondolella are food shortages caused by the mass extinction, the shallowing of the sea water as well as the increase in opacity of the sea water during the end Permian. The same phenomenon of Neogondolella size reduction is also observed in preliminary research from the same horizon at Shangsi Section, Sichuan Province. All the evidence suggests that there was a mass extinction that occurred at the horizon of bed 24e, and the evidence supports the viewpoint of a multi-phase mass extinction during the Permian and Triassic transition in South China.

  11. Earth's biggest 'whodunnit': unravelling the clues in the case of the end-Permian mass extinction.

    White, Rosalind V


    The mass extinction that occurred at the end of the Permian period, 250 million years ago, was the most devastating loss of life that Earth has ever experienced. It is estimated that ca. 96% of marine species were wiped out and land plants, reptiles, amphibians and insects also suffered. The causes of this catastrophic event are currently a topic of intense debate. The geological record points to significant environmental disturbances, for example, global warming and stagnation of ocean water. A key issue is whether the Earth's feedback mechanisms can become unstable on their own, or whether some forcing is required to precipitate a catastrophe of this magnitude. A prime suspect for pushing Earth's systems into a critical condition is massive end-Permian Siberian volcanism, which would have pumped large quantities of carbon dioxide and toxic gases into the atmosphere. Recently, it has been postulated that Earth was also the victim of a bolide impact at this time. If further research substantiates this claim, it raises some intriguing questions. The Cretaceous-Tertiary mass extinction, 65 million years ago, was contemporaneous with both an impact and massive volcanism. Are both types of calamity necessary to drive Earth to the brink of faunal cataclysm? We do not presently have enough pieces of the jigsaw to solve the mystery of the end-Permian extinction, but the forensic work continues.

  12. Carbon isotope excursions across the Permian-Triassic boundary in the Meishan section, Zhejiang Province, China


    Both gradual and sharp decrease in organic and carbonate carbon isotope values were detected across the Permian-Triassic boundary in the Meishan section, Changxing, Zhejiang Province, China. The gradual decrease in organic carbon isotope values started at the bottom of Bed 23, coinciding with the strong oscillations of total organic carbon (TOC) contents, indicates increasing fluxes from carbonate to organic carbon reservoir during this interval. A 2.3‰ sharp drop of inorganic carbon isotope values occurred at the uppermost part of Bed 24e. A 3.7‰ sharp drop of organic carbon isotope values occurred in Bed 26. The dramatic drop of inorganic carbon isotope value of 8‰ reported previously is not confirmed from the unweathered carbonate samples in Bed 27. The large-scale fluctuation of organic carbon isotope values in the Yinkeng Formation reflects different extent of mixing of marine and terrestrial organic matters. The gradual depletion and subsequent sharp drop of carbon isotopes near the Permian-Triassic boundary might indicate complex causes of the end-Permian mass extinction.

  13. Palynostratigraphy of Permian succession from Binja Block, South Karanpura Coalfield, Jharkhand, India

    Srikanta Murthy; Archana Tripathi; B Chakraborti; U P Singh


    Palynological investigations are carried out on approximately 538.00-m thick Gondwana strata from borehole SKB-1, Binja Block, South Karanpura Coalfield in Jharkhand. Based on the distribution pattern of age marker palynotaxa, two distinct palynoassemblages are identified. Palynoassemblage-I in the lithologically designated Barren Measures and Barakar formations, between 552.00 and 53.20 m depth show dominance of striate bisaccate (Striatopodocarpites, Crescentipollenites, and Faunipollenites) and abundance of nonstriate bisaccate (Scheuringipollenites). Upward the Palynoassemblage-II (39.50–13.80 m depth) is rich in striate bisaccate (Striatopodocarpites and Crescentipollenites) and significant enveloping monosaccate Densipollenites magnicorpus pollen. These strata have been equated with Raniganj Formation of Latest Permian age. The First Appearance Datum (FAD) of Arcuatipollenites pellucidus, Playfordiaspora cancellosa, Alisporites sp., Falcisporites sp. and Krempipollenites indicus observed at 13.80 and 39.50 m depth, mark the transition of Permian into the Lower Triassic. The FADs of Guttulapollenites spp. at 49.10 and 504.70 m, Goubinispora morondavensis at 415.90 m, Alisporites ovalis and Arcuatipollenites sp. at 526.70 m is observed and suggest that these sediments are equivalent to Raniganj Formation, Late Permian in age.

  14. A paleoclimatic simulation of the Late Permian greenhouse world and its consequences

    Moore, G.T.; Jacobson, S.R.; Hayashida, D.N. (Chevron Oil Field Research Co., La Habra, CA (United States))


    Sea-floor spreading assembled all the major cratonic blocks into a single supercontinent once in the Phanerozoic Eon. This unique Late Permian crustal tectonic event produced Pangaea and an enormous oceanic basin volume that dropped sea level to a global lowstand unrivaled in the Phanerozoic. Two paleoclimatic simulations using a numerical three-dimensional general circulation model tested changes in the greenhouse effect. The authors conclude that for a simulation to fit the Late Permian geologic record, the paleoatmosphere must contain an enhanced greenhouse gas effect. A third simulation tested changes of paleogeography in southern Pangaea (Gondwana) that did not appreciably alter the harsh continental paleoclimate. The simulated paleoclimatic changes provide extraordinarily warm ocean and atmosphere, and a significant reduction in continental rainfall and runoff. These conditions inevitably lead to more aridity and less vegetation on land, gradually reduce the delivery of vital nutrients from continental sources to marine margins, systematically liberate CO{sub 2} dissolved in ocean water, and incrementally increase stress on marine and terrestrial biotas. These consequences severely disrupted rates of oxygen and carbon cycling. Their quantitative paleoclimatic simulation is consistent with distributions of red beds, evaporites, coals, marine shelf areas, seawater isotope trends, and paleontologic originations and extinctions. Thus, the Pangaean plate assembly probably triggered an inexorable sequence of geophysical, geochemical, and biological events that forced an elevated greenhouse effect in the Late Permian, nearly annihilating the Phanerozoic biota.

  15. The Permian seamount stratigraphic sequence in Chiang Mai, North Thailand and its tectogeographic significance

    CHONGLAKMANI; Chongpan


    The widespread Permian carbonate strata outcropped in northwestern Thailand are considered as the evidence for the Late Paleozoic shallow Tethys. Our investigation, however, shows that basalt can be discovered usually under the Permian carbonate sequence in Chiang Mai-Fang area, northwestern Thailand. The basalt belongs to subalkalic basalt and potassic trachybasalt. They are characterized by high P and Ti in major elements, by high content, enriching LREE, lacking δEu anomaly in rare earth elements, and by enrichment of the large ion lithophile element (LILE) (K, Rb, Ba) and high field strength element (HFSE) (Nb, Ta, Zr) in trace elements, which can be compared with the characters of the oceanic island basalt in Three Rivers (Jinsha River, Lancang River, and Nujiang River) area, southwestern China. Therefore, the Permian carbonate in the studied area was deposited on a sea-mount, rather than on a stable carbonate platform. The oceanic basin is correlated to the Late Paleozoic ocean represented by the Changning-Menglian Belt in southwestern China and they are a major basin of the Paleo-Tethyan Archipelagoes Ocean. The result indicates nonexistence of a Shan-Thai Block in the Late Paleozoic.

  16. The Permian seamount stratigraphic sequence in Chiang Mai, North Thailand and its tectogeographic sisnificance

    FENG QingLai; YANG WenQiang; SHEN ShangYue; CHONGLAKMANI Chongpan; MALILA Kitsana


    The widespread Permian carbonate strata outcropped in northwestern Thailand are considered as the evidence for the Late Paleozoic shallow Tethys. Our investigation, however, shows that basalt can be discovered usually under the Permian carbonate sequence in Chiang Mai-Fang area, northwestern Thailand. The basalt belongs to subalkalic basalt and potassic trachybasalt. They are characterized by high P and Ti in major elements, by high content, enriching LREE, lacking o"Eu anomaly in rare earth elements, and by enrichment of the large ion lithophile element (LILE) (K, Rb, Ba) and high field strength element (HFSE) (Nb, Ta, Zr) in trace elements, which can be compared with the characters of the oceanic island basalt in Three Rivers (Jinsha River, Lancang River, and Nuiiang River) area,southwestern China. Therefore, the Permian carbonate in the studied area was deposited on a seamount, rather than on a stable carbonate platform. The oceanic basin is correlated to the Late Paleozoic ocean represented by the Changning-Menglian Belt in southwestern China and they are a major basin of the Paleo-Tethyan Archipelagoes Ocean. The result indicates nonexistence of a Shan-Thai Block in the Late Paleozoic.

  17. Schematic designs for penetration seals for a repository in the Permian Basin. [Deaf Smith County, Texas

    Kelsall, P.C.; Case, J.B.; Coons, W.E.; Franzone, J.G.; Meyer, D.


    The isolation of radioactive wastes in geologic repositories requires that human-made penetrations such as shafts, tunnels, or boreholes are adequately sealed. This report describes schematic seal designs for a repository in bedded salt referenced to the stratigraphy of the Permian Basin. The designs are presented for extensive peer review and will be updated as conceptual designs if the Permian Basin is selected as a candidate repository site. The principal material used in the seal system in the repository-level rooms and tunnels is crushed salt obtained from excavating the repository. It is anticipated that crushed salt will consolidate in response to closure of the repository rooms, to the degree that mechanical and hydrologic properties will eventually match those of undisturbed, intact salt. For Permian Basin Unit 4 salt, analyses indicate that this process will require approximately 700 years for a seal located at the base of one of the repository shafts (where there is little increase in temperature due to waste emplacement) and approximately 200 years for a seal located in a main passageway within the repository. These analyses are based on uncertain laboratory data regarding intact salt creep rates and crushed salt consolidation characteristics, and must be regarded as preliminary. Bulkheads composed of concrete, as well as bentonite-rich earth fill, are also included in the seal system as components which will have low permeability during the period required for salt consolidation.

  18. Notes on the discovery of Permian Acipenseriformes in China

    LU Liwu; LI Daqing; YANG Liangfeng


    @@ Acipenseriformes is a primitive group of Osteichthyan fishes. Most of its living representatives, such as Psephurus gladius and Acipenser sinensis of the Yangtze River, are fishes facing extinction. Up to now, the earliest fossil Acipenseriformes is Chondrosteus found in the Early Jurassic of England[1].

  19. Permian-Triassic Conodonts from Dajiang (Guizhou, South China) and Their Implication for the Age of Microbialite Deposition in the Aftermath of the End-Permian Mass Extinction

    Haishui Jiang; Xulong Lai; Yadong Sun; Paul B Wignall; Jianbo Liu; Chunbo Yan


    The widespread microbialites deposition that followed the End-Permian mass extinction in the Tethyan realm have been intensively studied because of the evidence they provide on the nature of this crisis and its aftermath. However, the age of the microbialite event remains controversial. New conodont collection across the Permian-Triassic (P-T) transition from Dajiang (Guizhou Province, South China) in this study enable us to discriminate four conodont zones, in ascending order, they are:Hindeodus parvus zone, Isarcicella lobata zone, Isarcicella isarcica zone and Hindeodus sosioensis zone. The age of microbialite in the P-T transition at the Dajiang Section is considered to be within the Hindeodus parvus zone and thus to clearly post-date the main extinction crisis. Reviewing the age of onset of microbialites throughout the Tethyan regions reveals two different ages: a Hindeodus changxingensis zone age is dominant in south-western and westernmost Tethys, whilst most other re-gions show microbialite deposition began in the Hindeodus parvus zone. Our investigation also indicates that two conodont changes occur at this time:an increase of hindeodid species immediately following a sequence boundary and the mass extinction, and a phase of extinction losses in the earliest Triassic Isarcicella isarcica zone during highstand development.

  20. The early larval development of the tropical reef lobster Enoplometopus antillensis Lütken (Astacidea, Enoplometopidae reared in the laboratory Desenvolvimento dos estágios iniciais da lagosta de recife tropical Enoplometopus antillensis Lütken (Astacidea, Enoplometopidae cultivada em laboratório

    Fernando A. Abrunhosa


    Full Text Available The early stages of the tropical reef lobster Enoplometopus antillensis Lütken, 1865 were described and illustrated in detail from specimens reared in the laboratory. Ovigerous females were captured in their habitat, at a depth of about 15 meters and transported to the laboratory. The larvae were reared in a recirculation water tank for approximately 15 days and then transferred to four 10 liters aquariums. The larvae were fed on Artemia sp. nauplii. Microalgae Dunaliella viridis was added daily to the culture. The larvae moulted seven times progressing through the zoea VIII. Megalopa stage was not achieved. The intermoulting period of each stage averaged from eight to 12 days. Morphological comparisons with previous reports are briefly discussed.Os estágios iniciais da lagosta tropical Enoplometopus antillensis Lütken, 1865 foram descritos e ilustrados a partir de espécimes cultivados em laboratório. Fêmeas ovígeras foram capturadas em seu habitat, na profundidade cerca de 15 metros e transportadas para o laboratório. As larvas foram cultivadas em tanques de água recirculante por aproximadamente 15 dias e, então transferidas para quatro aquários (capacidade 10 litros. As larvas foram alimentadas com náuplios de Artemia sp. recém eclodidos. A microalga Dunaliella viridis AUTOR foi diariamente adicionada no cultivo. As larvas mudaram sete vezes alcançando o zoea VIII. O estágio megalopa não foi obtido. O período de intermuda de cada estágio variou de cerca de oito a 12 dias. Comparações morfológicas com trabalhos anteriores são brevemente discutidas.

  1. Coal petrology of coal seams from the Leao-Butia Coalfield, Lower Permian of the Parana Basin, Brazil - Implications for coal facies interpretations

    Silva, M.B. [Laboratorio de Oceanografia Geologica, Departamento de Geociencias, Fundacao Universidade Federal do Rio Grande, FURG, Av. Italia km 08, Campus Carreiros, 96201-900, Rio Grande, RS (Brazil); Kalkreuth, W.; Holz, M. [Instituto de Geociencias, UFRGS, Av. Bento Goncalves, 9500 91501-970 Porto Alegre, RS (Brazil)


    In the Leao-Butia Coalfield, Rio Grande do Sul the coal seams occur in the Rio Bonito Formation, Guata Group, Tubarao Supergroup of the Parana Basin, Brazil and are of Permian (Artinskian-Kungurian) age. This study is the first detailed investigation on the coal petrographic characterization of the coal-bearing sequence in relation to the depositional settings of the precursor mires, both in terms of whole seam characterization and in-seam variations. The study is based on the analyses of nine coal seams (I2, CI, L4, L3, L2, L1, S3, S2, S1), which were selected from core of borehole D-193, Leao-Butia and represent the entire coal-bearing sequence. The interpretation of coal facies and depositional environment is based on lithotype, maceral and microlithotype analyses using different facies-critical petrographic indices, which were displayed in coal facies diagrams. The seams are characterized by the predominance of dull lithotypes (dull, banded dull). The dullness of the coal is attributed to relatively high mineral matter, inertinite and liptinite contents. The petrographic composition is dominated by vitrinite (28-70 vol.% mmf) and inertinite (> 30 vol.% mmf) groups. Liptinite contents range from 7 to 30 vol.% (mmf) and mineral matter from 4-30 vol.%. Microlithotypes associations are dominated by vitrite, duroclarite, carbominerite and inertite. It is suggested that the observed vertical variations in petrographic characteristics (lithotypes, microlithotypes, macerals, vitrinite reflectance) were controlled by groundwater level fluctuations in the ancient mires due to different accommodation/peat accumulation rates. Correlation of the borehole strata with the general sequence-stratigraphical setting suggests that the alluvial fan system and the coal-bearing mudstone succession are linked to a late transgressive systems tract of sequence 2. Based on average compositional values obtained from coal facies diagrams, a deposition in a limno-telmatic to limnic coal

  2. Modeling streamflow from snowmelt in the upper Rio Grande

    Annual snowpack in the high elevation snowsheds of the Upper Rio Grande (URG) Basin is a vital source of surface water for irrigated agriculture in New Mexico. Maximum streamflow from the annual snowpack usually occurs in early May for the southernmost snowsheds (e.g., Ojo Caliente) and at the end o...

  3. Depositional setting, petrology and chemistry of Permian coals from the Paraná Basin: 2. South Santa Catarina Coalfield, Brazil

    Kalkreuth, W.; Holz, M.; Mexias, A.; Balbinot, M.; Levandowski, J.; Willett, J.; Finkelman, R.; Burger, H.


    In Brazil economically important coal deposits occur in the southern part of the Paran?? Basin, where coal seams occur in the Permian Rio Bonito Formation, with major coal development in the states of Rio Grande de Sul and Santa Catarina. The current paper presents results on sequence stratigraphic interpretation of the coal-bearing strata, and petrological and geochemical coal seam characterization from the South Santa Catarina Coalfield, Paran?? Basin.In terms of sequence stratigraphic interpretation the precursor mires of the Santa Catarina coal seams formed in an estuarine-barrier shoreface depositional environment, with major peat accumulation in a high stand systems tract (Pre-Bonito and Bonito seams), a lowstand systems tract (Ponta Alta seam, seam A, seam B) and a transgressive systems tract (Irapu??, Barro Branco and Treviso seams).Seam thicknesses range from 1.70 to 2.39. m, but high proportions of impure coal (coaly shale and shaley coal), carbonaceous shale and partings reduce the net coal thickness significantly. Coal lithoypes are variable, with banded coal predominant in the Barro Branco seam, and banded dull and dull coal predominantly in Bonito and Irapu?? seams, respectively. Results from petrographic analyses indicate a vitrinite reflectance range from 0.76 to 1.63 %Rrandom (HVB A to LVB coal). Maceral group distribution varies significantly, with the Barro Branco seam having the highest vitrinite content (mean 67.5 vol%), whereas the Irapu?? seam has the highest inertinite content (33.8. vol%). Liptinite mean values range from 7.8. vol% (Barro Branco seam) to 22.5. vol% (Irapu?? seam).Results from proximate analyses indicate for the three seams high ash yields (50.2 - 64.2wt.%). Considering the International Classification of in-Seam Coals, all samples are in fact classified as carbonaceous rocks (>50wt.% ash). Sulfur contents range from 3.4 to 7.7 wt.%, of which the major part occurs as pyritic sulfur. Results of X-ray diffraction indicate the

  4. Petrographic approach to the study of organic microfossils from the Irati Subgroup (Permian, Paraná Basin, Brazil)

    Calça, Cléber Pereira; Fairchild, Thomas R.


    Early diagenetic chert, infrequently exploited in Phanerozoic micropaleontology, was examined for organic-walled microfossils in petrographic thin sections of silicified dolostones from diverse levels and localities of the Assistência Formation (Permian, Paraná Basin) in the state of São Paulo, Brazil. In contrast to previous palynological studies of this formation, the use of thin sections allowed direct observation in three dimensions of common palynomorphs, as well as benthic microbial mats preserved in situ in various stages of their life cycles and degradation. As in palynological residues from the more well-known shale of this formation, the chert contains wind-dispersed pollen grains and phytoclasts derived from terrestrial sources and planktonic cryptarchs (unornamented coccoidal unicellular or colonial palynomorphs). However, only in the chert is it possible to see much more delicate microfossils, such as abundant cyanobacteria of the in situ benthic microbiota as well as chlorophycean microalgae of the microphytoplankton. Post-depositional processes affecting the formation have destroyed all but the most resistant organic remains in the other lithologies, such that only rare, degraded pollen grains are seen in the unsilicified dolostone of the formation, and in the shale the vast majority of microfossils have been compacted to flattened disks. On the other hand, early silicification not only preserved organic remains at an incipient stage of decomposition but also impeded significant further degradation due to compaction, recrystallization, and oxidation. Thus, the petrographic study of such chert can complement traditional palynological investigations in Phanerozoic rocks by furnishing hitherto unavailable information, especially with regard to benthic organic microfossils and fragile organic-walled phytoplankton normally absent from organic residues.

  5. Global taxonomic diversity of anomodonts (tetrapoda, therapsida and the terrestrial rock record across the Permian-Triassic boundary.

    Jörg Fröbisch

    Full Text Available The end-Permian biotic crisis (~252.5 Ma represents the most severe extinction event in Earth's history. This paper investigates diversity patterns in Anomodontia, an extinct group of therapsid synapsids ('mammal-like reptiles', through time and in particular across this event. As herbivores and the dominant terrestrial tetrapods of their time, anomodonts play a central role in assessing the impact of the end-Permian extinction on terrestrial ecosystems. Taxonomic diversity analysis reveals that anomodonts experienced three distinct phases of diversification interrupted by the same number of extinctions, i.e. an end-Guadalupian, an end-Permian, and a mid-Triassic extinction. A positive correlation between the number of taxa and the number of formations per time interval shows that anomodont diversity is biased by the Permian-Triassic terrestrial rock record. Normalized diversity curves indicate that anomodont richness continuously declines from the Middle Permian to the Late Triassic, but also reveals all three extinction events. Taxonomic rates (origination and extinction indicate that the end-Guadalupian and end-Permian extinctions were driven by increased rates of extinction as well as low origination rates. However, this pattern is not evident at the final decline of anomodont diversity during the Middle Triassic. Therefore, it remains unclear whether the Middle Triassic extinction represents a gradual or abrupt event that is unique to anomodonts or more common among terrestrial tetrapods. The end-Permian extinction represents the most distinct event in terms of decline in anomodont richness and turnover rates.

  6. Late Permian Tsunamites in Guryul Ravine (Kashmir, India) - revisited and rejected

    Krystyn, Leopold; Horacek, Micha; Brandner, Rainer; Parcha, Suraj


    Recent claims for tsunami-related event beds induced by the Siberian Trap basalts in this section (Brookfield et al., 2013) have to be questioned. Identical storm generated carbonate beds occur not only during a short interval close to the Permian-Triassic (P-T) boundary but through a major part of the late Permian (Changhsingian) succession there - as low as 26 m below the so-called tsunami beds. Moreover, during our recent study in a closely neighbouring place called Mandakpal (less than 10 km to the southeast), no signs of tsunamites have been detected in time-correlative finegrained sediments. Based on sedimentary and trace fossil evidence we interpret the late Permian of Guryul as relatively shallow, neritic and delta-influenced. The so-called tsunamites are shelly-enriched discontinuous carbonate lenses fed downslope through local channels. Judging from the distinct facies change from the storm related "tsunamites" to thinly bedded mud turbidites above, the sudden deepening may be explained by local and still rift-related tectonics along the NIM (North-Indian Gondwana Margin) which led to episodic seismic induced sediment redeposition in the area of Guryul. Synsedimentary tectonic activity with tilting and eventual Horst and Graben structure building along the large NIM is indicated by margin inversion during the P-T boundary interval leading to sedimentary breaks and 20 times thinner, condensed limestone deposits far offshore from Guryul in Spiti (Krystyn et al., 2004) and Tibet (Orchard et al., 1994). Thus, local seismic activity seems to be a far more logic explanation of the Guryul "tsunamites" than the eruption of the Siberian Traps more than 6000 km away. References Brookfield, M. E., Algeo, T. J., Hannigan, R., Williams, J and Bhat, G. M., 2013: Shaken and Stirred: Seismites and Tsunamites at the Permian-Triassic boundary, Guryul Ravine, Kashmir, India. Palaios, v. 28, 568-582. Krystyn, L., Balini, M. and Nicora, A., 2004: Lower and Middle Triassic

  7. Geochemical characteristics of the Permian basins and their provenances across the Solonker Suture Zone: Assessment of net crustal growth during the closure of the Palaeo-Asian Ocean

    Eizenhöfer, Paul R.; Zhao, Guochun; Zhang, Jian; Han, Yigui; Hou, Wenzhu; Liu, Dongxing; Wang, Bo


    The Solonker Suture Zone is commonly recognised as the location of the Late Permian to Early Triassic closure of the Palaeo-Asian Ocean in the southeastern segment of the Central Asian Orogenic Belt. However, the absence of typical suture-related features, as a consequence of uncommon collisional geometries, gave it a cryptic nature. Thus, the tectonic setting, which led to suturing, still remains enigmatic. A geochemical characterisation of Permian sedimentary and volcanic rocks across the suture was carried out. Supplemented with Hf and Nd isotopic analyses, this approach enables not only a better definition of such regional suture, but also estimates on the long-controversial issue of net crustal growth in accretionary tectonic environments. The results indicate short sedimentary transport distances between the arc basins and their provenances, of which the studied volcanic rocks were a major contributor. Similar enrichment and depletion patterns with respect to N-MORB and average continental crust further corroborate a close source-sediment relationship. Immobile element provenance analyses indicate that the active continental northern margin of the North China Craton was a major source for arc basins to the south of the Solonker Suture Zone. To its north, arc basins are interpreted to be sourced by a more complex mixture of provenances, e.g., the Baolidao volcanic arc suite and the heterogenous Precambrian basement of southern Mongolia. An overall collisional tectonic setting across the suture is recognised. The geochemical signature of sedimentary rocks to the south of the suture points at an active continental arc setting, whereas the bimodal geochemical distribution of the samples to the north shows a contemporaneous active oceanic island arc as well as a passive margin environment. These features favour a double-sided subduction of the Palaeo-Asian Ocean beneath the North China Craton and the Mongolian Arcs throughout the Palaeozoic, including back

  8. New Paleomagnetic Data from Upper Permian and Lower Triassic Volcanic Sequences from Hua Binh, Quynh Nhai, and Thuan Chau Localities, Northwest Veitnam and Their Bearing on the Accretion History of Southeast Asia

    Geissman, J. W.; Chi, C. T.


    New paleomagnetic data from Upper Permian to Lower Triassic volcanic rocks sampled in NW Vietnam provide more quantitative constraints on the paleogeographic setting of crustal elements that comprise the Song Da Terrane, east of the Song Ma suture, between the South China block (SCB) and north Indochina. These include results from 12 sites (125 samples) from basalts of the Vien Nam Formation, exposed at Hoa Binh Dam; eight sites (74 samples) from basalts of the Cam Thuy Formation near Thuan Chau; and 19 sites (198 samples) from andesites and basalts of the Vien Nam Formation near Quynh Nhai. The collection is limited by the quality of exposures and quantity of independent flows. Most sites yield interpretable magnetizations in progressive demagnetization, and the response implies that characteristic remanent magnetization (ChRM) components are carried by low-titanium magnetite or hematite, or a combination of both; these are isolated from secondary components. Rock magnetic data and petrography support the retention of an early-acquired thermoremanent magnetization in most sites. The Vien Nam Formation mafic volcanic rocks yield a grand mean, in geographic coordinates, of D=33.8o, I=-28.4o ( a95 = 9.5o, k =30.3, N=9 accepted sites), and a pole position at Lat=41.1N, Long=239.8E and a paleolatitude at ~15o S during the Late Permian to Early Triassic. Permian basalts of the Cam Thuy Formation provide a grand mean, corrected for structural tilt, of D=216.1o, I=+10.5o, a95=8.9o, k=107.8, and N= 4, with a pole position at Lat=45.6N, Long=226.8E. Volcanic rocks at the Quynh Nhai locality likely yield the most robust paleofield determination, as the data set is of dual polarity and passes a reversal test. The tilt corrected grand mean (normal polarity) is D=48.3o, I=-10.0o, a95=8.0o, k=27.7, N = 13, and this in turn yields an inferred paleomagnetic pole at Lat=35.7N, Long=217.4E, and a paleolatitude of 5.1oS for the late Permian. Compared with the Late Permian-Early

  9. Rock in Rio: forever young

    Ricardo Ferreira Freitas


    Full Text Available The purpose of this article is to discuss the role of Rock in Rio: The Musical, as herald of megafestival Rock in Rio. Driven by the success that musicals have reached in Brazil, we believe that the design of this spectacle of music, dance and staging renews the brand of the rock festival, once it adds the force of young and healthy bodies to its concept. Moreover, the musical provides Rock in Rio with some distance from the controversal trilogy of sex, drugs and rock and roll, a strong mark of past festivals around the world. Thus, the musical expands the possibilities of growth for the brand.

  10. Rock in Rio: eternamente jovem

    Ricardo Ferreira Freitas; Flávio Lins Rodrigues


    The purpose of this article is to discuss the role of Rock in Rio: The Musical, as herald of megafestival Rock in Rio. Driven by the success that musicals have reached in Brazil, we believe that the design of this spectacle of music, dance and staging renews the brand of the rock festival, once it adds the force of young and healthy bodies to its concept. Moreover, the musical provides Rock in Rio with some distance from the controversal trilogy of sex, drugs and rock and roll, a strong mark ...

  11. Paleomagnetic results from the Upper Permian of the eastern Qiangtang Terrane of Tibet and their tectonic implications

    Huang, Kainian; Opdyke, Neil D.; Peng, Xingjie; Li, Jiguang, Li


    Paleomagnetic samples were collected from red intercalations within the Upper Permian Tuoba Formation from the eastern Qiangtang Terrane of Tibet. Progressive thermal demagnetization has revealed a prefolding, possibly primary, characteristic remanent magnetization (ChRM), which yields a tilt-corrected mean direction ofD/I=25.2°6.7° (α 95 = 8.6°). The equatorial paleolatitude for the Qiangtang Terrane indicated by this result and the similarity of Late Permian fauna and flora shared by the Qiangtang Terrane and South China Block (SCB) suggest that the two terranes were in close proximity during the Late Permian. A Late Permian paleogeographic reconstruction involving the major terranes of the present-day East Asia is constructed, in which the Songpan-Garze Sea is surrounded by Laurasia, the North China Block (NCB), the SCB and the Qiangtang Terrane. The Permian development of the Songpan-Garze Sea appears to be related to the rifting of the Qiangtang Terrane off the SCB. Its Triassic evolution may resemble that of the Solomon Sea today in the southwestern Pacific.

  12. The end-Permian regression in the western Tethys: sedimentological and geochemical evidence from offshore the Persian Gulf, Iran

    Tavakoli, Vahid; Naderi-Khujin, Mehrangiz; Seyedmehdi, Zahra


    Detailed sedimentological and geochemical records across the Permian-Triassic boundary (PTB) in five offshore wells of the central Persian Gulf served to interpret the end-Permian sea-level change in this region. A decrease in sea level at the PTB was established by petrographical and geochemical study of the boundary. Thin sections showed that Upper Permian strata are composed of dolomite with minor anhydrite, changing into limestone in Lower Triassic sediments. Brine dilution toward the boundary supports sea-level fall in the Permian-Triassic transition, reflected by a decrease in anhydrite content and a shallowing-upward trend from lagoonal to peritidal facies. Isotopic changes at the boundary are in favor of sea-level fall. Changes in both carbon (from about 4 to -1‰) and oxygen (from 2 to -5‰) stable isotopes show negative excursions. The shift in carbon isotope values is a global phenomenon and is interpreted as resulting from carbonate sediment interaction with 12C-rich waters at the end-Permian sea-level fall. However, the oxygen isotope shift is attributed to the effect of meteoric waters with negative oxygen isotope values. The increase in strontium isotope ratios is also consistent with the high rate of terrestrial input at the boundary. The effect of meteoric conditions during diagenesis is evident from vuggy and moldic porosities below the PTB. The following transgression at the base of the Triassic is evident from the presence of reworked fossils and intraclasts resulting from deposition from agitated water.

  13. Rapid Carbonate Depositional Changes Following the Permian-Triassic Mass Extinction:Sedimentary Evidence from South China

    Li Tian; Jinnan Tong; David Bottjer; Daoliang Chu; Lei Liang; Huyue Song; Haijun Song


    Various environmental changes were associated with the Permian-Triassic mass extinc-tion at 252.2 Ma. Diverse unusual sediments and depositional phenomena have been uncovered as re-sponses to environmental and biotic changes. Lithological and detailed conodont biostratigraphic cor-relations within six Permian-Triassic boundary sections in South China indicate rapid fluctuations in carbonate deposition. Four distinct depositional phases can be recognized:(1) normal carbonate depo-sition on the platform and slope during the latest Permian;(2) reduced carbonate deposition at the on-set of the main extinction horizon; (3) expanded areas of carbonate deposition during the Hindeodus changxingsensis Zone to the H. parvus Zone;and (4) persistent mud-enriched carbonate deposition in the aftermath of the Permian-Triassic transition. Although availability of skeletal carbonate was sig-nificantly reduced during the mass extinction, the increase in carbonate deposition did not behave the same way. The rapid carbonate depositional changes, presented in this study, suggest that diverse envi-ronmental changes played key roles in the carbonate deposition of the Permian-Triassic mass extinction and onset of its aftermath. An overview of hypotheses to explain these changes implies enhanced terres-trial input, abnormal ocean circulation and various geobiological processes contributed to carbonate saturation fluctuations, as the sedimentary response to large volcanic eruptions.

  14. Magnetic properties of the remagnetized Middle-Ordovician limestones of the Ponón Trehué Formation (San Rafael Block, central-western Argentina): Insights into the Permian widespread Sanrafaelic overprint

    Fazzito, Sabrina Y.; Rapalini, Augusto E.


    The widespread Sanrafaelic remagnetization reset most of the early Cambrian to mid-Ordovician carbonate platform of the Argentine Precordillera and the calcareous units of the San Rafael Block. We conducted a detailed rock-magnetic study on the Middle-Ordovician limestones of the Ponón Trehué Formation at both limbs of a tight anticline exposed in the San Rafael Block (Mendoza province, central-western Argentina) that are carriers of a syntectonic magnetization of Permian age. We found that the magnetic overprint in the Ponón Trehué Formation is carried by both pyrrhotite and magnetite, with goethite and subordinate haematite likely related to weathering. Hysteresis parameters, frequency dependence of magnetic susceptibility, Cisowski and modified Lowrie-Fuller tests suggest the presence of ultrafine particles of chemical origin. Demagnetization of natural remanent magnetization and of three-axis isothermal remanence confirm pyrrhotite and magnetite as important contributors to the remanence. Both minerals carry the same magnetic syntectonic component suggesting a coeval or nearly coeval remanence acquisition and therefore mineral formation. This and the results of the magnetic fabric analyses indicate an authigenic origin of the magnetic minerals during folding associated with the Sanrafaelic tectonic phase (ca. 280 Ma). Although the chemically active (oxidizing?) fluids expelled from the orogen as it developed in the early Permian is a viable explanation for the Sanrafaelic remagnetization, the role of the nearly coeval magmatism in Precordillera and the San Rafael Block remains to be properly evaluated.

  15. An abrupt extinction in the Middle Permian (Capitanian) of the Boreal Realm with a causal link to anoxia, acidification and mercury poisoning

    Bond, David; Wignall, Paul; Joachimski, Michael; Sun, Yadong; Savov, Ivan; Grasby, Stephen; Beauchamp, Benoit; Blomeier, Dierk


    The controversial Capitanian (Middle Permian, 262 Ma) extinction event is mostly known from equatorial latitudes and consequently its global extent is poorly resolved. We demonstrate that there were two, severe extinctions amongst brachiopods in northern Boreal latitudes (Spitsbergen), in the Middle to Late Permian, separated by a recovery phase. New age dating of the Kapp Starostin Formation of Spitsbergen using strontium and carbon isotopic trends suggests that the first crisis occurred in the Capitanian. This age assignment indicates that this Middle Permian extinction is manifest at higher latitudes. Redox proxies (pyrite framboids and trace metals) show that the Boreal crisis coincided with an intensification of oxygen depletion, implicating anoxia in the extinction scenario. The highly toxic metal mercury becomes enriched in strata at the Middle Permian extinction level implicating death-by-toxicity (and a possible link to volcanism). Finally, the near-total loss of carbonates across the Boreal Realm in the Middle to Late Permian also suggests a role for acidification. New in prep. data from Ellesmere Island, Arctic Canada (samples collected July 2015) tentatively suggests that this potent "three strikes and you're out" extinction mechanism was a Boreal-wide phenomenon. The Late Permian recovery interval saw the appearance of new brachiopod and bivalve taxa alongside survivors, and an increased mollusk dominance, resulting in an assemblage reminiscent of younger Mesozoic assemblages. The subsequent end-Permian mass extinction terminated this Late Permian radiation.

  16. Chemical signature of two Permian volcanic ash deposits within a bentonite bed from Melo, Uruguay

    Liane M. Calarge


    Full Text Available A Permian bentonite deposit at Melo, Uruguay is composed of a calcite-cemented sandstone containing clay pseudomorphs of glass shards (0-0.50 m overlying a pink massive clay deposit (0.50-2.10m. The massive bed is composed of two layers containing quartz and smectite or pure smectite respectively. The smectite is remarkably homogeneous throughout the profile: it is a complex mixed layer composed of three layer types whose expandability with ethylene glycol (2EG 1EG or 0EG sheets in the interlayer zone which correspond to low-, medium- and high-charge layers respectively varies with the cation saturating the interlayer zone. The smectite homogeneity through the profile is the signature of an early alteration process in a lagoonal water which was over saturated with respect to calcite. Compaction during burial has made the bentonite bed a K-depleted closed system in which diagenetic illitization was inhibited. Variations in major, REE and minor element abundances throughout the massive clay deposit suggest that it originated from two successive ash falls. The incompatible element abundances are consistent with that of a volcanic glass fractionated from a rhyolite magma formed in a subduction/collision geological context.Um depósito Permiano de bentonita em Melo, Uruguai,é composto por um arenito com cimento calcítico contendo pseudomorfos de argila sobre detritos vítreos(0-0.50 m superpostos a um deposito maciço de argila rosado (0.50-2.10 m. A camada maciça é composta por dois níveis contendo quartzo e esmectita ou esmectita pura, respectivamente. A homogeneidade de esmectita ao longo do perfil é notável: trata-se de um interestratificado composto de três tipos de camadas, cuja expansibilidade com etileno-glicol (folhas 2EG, 1EG ou 0EG na zona interfoliar correspondentes a camadas com baixa, média e alta carga, respectivamente variam com o tipo de cátion que satura a zona interfoliar. A homogeneidade da esmectita ao longo do perfil

  17. End-Permian Mass Extinction in the Oceans: An Ancient Analog for the Twenty-First Century?

    Payne, Jonathan L.; Clapham, Matthew E.


    The greatest loss of biodiversity in the history of animal life occurred at the end of the Permian Period (˜252 million years ago). This biotic catastrophe coincided with an interval of widespread ocean anoxia and the eruption of one of Earth's largest continental flood basalt provinces, the Siberian Traps. Volatile release from basaltic magma and sedimentary strata during emplacement of the Siberian Traps can account for most end-Permian paleontological and geochemical observations. Climate change and, perhaps, destruction of the ozone layer can explain extinctions on land, whereas changes in ocean oxygen levels, CO2, pH, and temperature can account for extinction selectivity across marine animals. These emerging insights from geology, geochemistry, and paleobiology suggest that the end-Permian extinction may serve as an important ancient analog for twenty-first century oceans.

  18. New Middle Permian palaeopteran insects from Lodève Basin in southern France (Ephemeroptera, Diaphanopterodea, Megasecoptera

    Jakub Prokop


    Full Text Available Three new palaeopteran insects are described from the Middle Permian (Guadalupian of Salagou Formation in the Lodève Basin (South of France, viz. the diaphanopterodean Alexrasnitsyniidae fam. n., based on Alexrasnitsynia permiana gen. et sp. n., the Parelmoidae Permelmoa magnifica gen. et sp. n., and Lodevohymen lapeyriei gen. et sp. n. (in Megasecoptera or Diaphanopterodea, family undetermined. In addition the first record of mayflies attributed to family Syntonopteridae (Ephemeroptera is reported. These new fossils clearly demonstrate that the present knowledge of the Permian insects remains very incomplete. They also confirm that the Lodève entomofauna was highly diverse providing links to other Permian localities and also rather unique, with several families still not recorded in other contemporaneous outcrops.

  19. Sickle cell disease: acute clinical manifestations in early childhood and molecular characteristics in a group of children in Rio de Janeiro Manifestações clínicas agudas na primeira e segunda infâncias e características moleculares da doença falciforme em um grupo de crianças do Rio de Janeiro

    Isaac Lima da Silva Filho


    Full Text Available OBJECTIVE: To describe clinical events of sickle cell disease and the correlation with β-globin haplotypes and α-thalassemia in under 6-year-old children. METHODS: A retrospective study was conducted of under 6-year-old children from the neonatal screening program in Rio de Janeiro. Forty-eight male and 48 female children were enrolled in this study, 79 with sickle cell anemia and 17 with hemoglobin SC. The mean age was 29.9 (standard deviation = 20.9 months, 62 (16.2 ± 8.6 were aged between 0-3 years old and 34 (54.9 ± 11.3 were from 3-6 years old. Painful events, acute splenic sequestration, hemolytic crises, hand-foot and acute chest syndromes and infections were evaluated. RESULTS: The events were more frequent in under 3-year-old children, 94% of children had at least one episode. Infection was the most common event affecting 88.5% of children. Acute splenic sequestration took place earlier, while painful crises and acute chest syndromes in under 6-year-old children. Thal-α 3.7 was observed in 20.9% of cases. Bantu was the most frequent haplotype found, followed by Benin. No correlation was observed between clinical events and β-globin haplotypes. Children with sickle cell anemia and α-thalassemia have less infectious events. No correlation was found among these polymorphisms and clinical events, however, the majority of children with Bantu/Bantu and without α-thalassemia had more clinical events.OBJETIVO: Avaliar a expressão clínica da anemia falciforme (AF e sua relação com os haplótipos da globina beta e talassemia alfa (Tal em crianças na primeira e segunda infâncias. MÉTODOS: Foi realizado um estudo descritivo e retrospectivo de série de casos em crianças na primeira e segunda infâncias provenientes da triagem neonatal do Rio de Janeiro. Um total de 96 crianças, 79 com AF e 17 com HbSC, 48 homens e 48 mulheres, média de idade de 29,9 (DP+/-20,9 meses, sendo 62 (16,2+/-8,6 na primeira e 34 (54,9+/-11,3 na segunda

  20. Early infection and asymptomatic spread of hepatitis A virus in a public child care center in Rio de Janeiro, Brazil: should attending children under two years of age be vaccinated?

    Liliane M Morais


    Full Text Available A cross-sectional study was conducted in order to identify hepatitis A virus (HAV serological markers in 418 individuals (mean age, 16.4 years; range, 1 month-80 years at a public child care center in Rio de Janeiro, Brazil, as well as to analyze risk factors and determine circulating genotypes. Serum samples were tested using an enzyme immunoassay. Reverse transcription polymerase chain reaction (RT-PCR was used to detect and characterize HAV RNA, and sequencing was performed. Anti-HAV antibodies and IgM anti-HAV antibodies were detected, respectively, in 89.5% (374/418 and 10.5% (44/418 of the individuals tested. Acute HAV infection in children was independently correlated with crawling (p < 0.05. In 56.8% (25/44 of the IgM anti-HAV-positive individuals and in 33.3% (5/15 of the IgM anti-HAV-negative individuals presenting clinical symptoms, HAV RNA was detected. Phylogenetic analysis revealed co-circulation of subgenotypes IA and IB in 93.3% (28/30 of the amplified samples. In present study, we verify that 79% (30/38 of children IgM anti-HAV-positive were asymptomatic. In child care centers, this asymptomatic spread is a more serious problem, promoting the infection of young children, who rarely show signs of infection. Therefore, vaccinating children below the age of two might prevent the asymptomatic spread of hepatitis A.

  1. Inorganic chemistry, petrography and palaeobotany of Permian coals in the Prince Charles Mountains, East Antarctica

    Holdgate, G.R. [School of Earth Sciences, The University of Melbourne, Victoria, 3010 (Australia); McLoughlin, S. [School of Natural Resource Sciences, Queensland University of Technology, P.O. Box 2434, Brisbane, Queensland 4001 (Australia); Drinnan, A.N. [School of Botany, The University of Melbourne, Victoria, 3010 (Australia); Finkelman, R.B.; Willett, J.C.; Chiehowsky, L.A. [U.S. Geological Survey, Mail Stop 956, Reston, VA 20092 (United States)


    Sampled outcrops of Permian coal seams of the Bainmedart Coal Measures in the Lambert Graben, eastern Antarctica, have been analysed for their proximates, ultimates, ash constituents and trace elements. A similar series of samples has been analysed for their principle maceral and microlithotype components and vitrinite reflectance. The coals are sub-bituminous to high volatile bituminous in rank; maturity increases markedly in southern exposures around Radok Lake where the oldest part of the succession is exposed and some strata have been intruded by mafic dykes and ultramafic sills. The coal ash is mostly silica and aluminium oxides, indicating that the mineral ash component is mostly quartz and various clay minerals. The ratio of silica to aluminium oxides appears to increase in an upward stratigraphic direction. The coal macerals include a relatively high liptinite content (mainly sporinite) that is significantly higher than for typical Gondwana coals. Greater degrees of weathering within the floodbasin/peat mire environments associated with climatic drying towards the end of the Permian might account for both preferential sporopollenin preservation and increased silica:aluminium oxide ratios up-section. Correlation of the coal maceral components to adjacent peninsula India coals indicates the closest comparative coals of similar age and rank occur within the Godavari Basin, rather then the Mahanadi Basin, which is traditionally interpreted to have been contiguous with the Lambert Graben before Gondwanan breakup. The petrological characteristics suggest that either previous interpretations of Palaeozoic basin alignments between Antarctica and India are incorrect, or that environmental settings and post-Permian burial histories of these basins were strongly independent of their tectonic juxtaposition. A permineralized peat bed within the succession reveals that the coals predominantly comprise wood- and leaf-rich debris derived from low-diversity forest

  2. Rare earth elements stratigraphic significance in late Permian coal measure from Bijie City, Guizhou Province, China

    WANG Qiang; YANG Ruidong; BAO Miao


    Rare earth elements (REEs) are good geological indicators. In order to understand REEs stratigraphic significance, REEs m Late Permian coal measure from Bijie City, western Guizhou Province, China were studied. The results showed that the contents of both light rare earth element (LREE) and ∑ REE were sharply increased in the boundary between Longtan Formation and Changxing Formation, which resulted from the gyration and discontinuity eruption of Emeishan basalt (REEs source) and frequent transgression-regression during forming coal. The coal measure and strata could be subdivided and correlate, and the sea-level change could be under stood by studying REEs content variation in coal measure.

  3. Sedimentology and distribution of Upper Permian lowstand fans, Bredehorn Member, Schuchert Dal Formation, East Greenland

    Kreiner-Moeller, M.; Piasecki, S.; Stemmerik, L.


    The Upper Permian basin of central East Greenland was characterized by shallow marine platform carbonates along the basin margins and over fault crests, and deeper water siliciclastics in the basin centres during the later stages of deposition. The basinal, siliciclastic-dominated succession includes the organic rich, source prone shales of the Ravnefjeld Formation and sandy and silty sediments of the Schuchert Dal Formation. The traditional view of these formations is that they represent discrete stages of basin evolution. However, renewed fieldwork indicates that the lower part of the sandy Bredehorn Member of the Schuchert Dal Formation is time equivalent to the upper part of the Ravnefjeld Formation. The presentation discusses the details.

  4. A new tubeworm of possible serpulid affinity from the Permian of Sicily

    Rossana Sanfilippo


    Full Text Available A new tubeworm, Palaeotubus sosiensis gen. et sp. nov., is described from the Permian Pietra di Salomone limestone (Sosio Valley, W Sicily. The new species is characterized by a thick tube ornamented with slightly flared peristomes and numerous longitudinal keels. The internal structure of the tube was obliterated during diagenesis, which prevents the tube unequivocal systematic attribution. Nevertheless, all preserved morphological characters strongly suggest that the new tube belongs to serpulids. When confirmed by further findings, the positive attribution of this new species to serpulids will imply that it represents the ancestor of the Serpulidae, and the earliest evidence of calcareous tubeworm polychaetes from the Palaeozoic.

  5. Impact event at the Permian-Triassic boundary: evidence from extraterrestrial noble gases in fullerenes.

    Becker, L; Poreda, R J; Hunt, A G; Bunch, T E; Rampino, M


    The Permian-Triassic boundary (PTB) event, which occurred about 251.4 million years ago, is marked by the most severe mass extinction in the geologic record. Recent studies of some PTB sites indicate that the extinctions occurred very abruptly, consistent with a catastrophic, possibly extraterrestrial, cause. Fullerenes (C60 to C200) from sediments at the PTB contain trapped helium and argon with isotope ratios similar to the planetary component of carbonaceous chondrites. These data imply that an impact event (asteroidal or cometary) accompanied the extinction, as was the case for the Cretaceous-Tertiary extinction event about 65 million years ago.

  6. Nitrogen isotope chemostratigraphy across the Permian-Triassic boundary at Chaotian, Sichuan, South China

    Saitoh, M.; Ueno, Y.; Nishizawa, M.; Isozaki, Y.; Takai, K.; Yao, J.; Ji, Z.


    Nitrogen isotopic compositions of upper Permian to lowermost Triassic rocks were analyzed at Chaotian, Sichuan, China, to clarify changes in the oceanic N cycle around the Permian-Triassic boundary (P-TB) including the entire Changhsingian (Late Late Permian) prior to the extinction. The analyzed ca. 40 m thick interval across the P-TB at Chaotian consists of three stratigraphic units: the upper Wujiaping Formation, the Dalong Formation, and the lowermost Feixianguan Formation, in ascending order. The upper Wujiaping Formation is composed of dark gray limestone with diverse shallow-marine fossils, such as calcareous algae, deposited on the shallow shelf. In contrast, the overlying Dalong Formation, ca. 25 m thick, is mainly composed of thinly bedded black mudstone and siliceous mudstone containing abundant radiolarians, deposited on the relatively deep slope/basin. Absence of bioturbation, substantially high TOC contents, and abundant occurrence of pyrite framboids in the Dalong Formation indicate deposition under anoxic conditions. The lowermost Feixianguan Formation is composed of thinly bedded gray marl and micritic limestone with minor fossils, deposited on the relatively shallow slope. δ15N values are in positive values around +1 to +2‰ in the upper Wujiaping Formation implying denitrification and/or anammox in the ocean. δ15N values gradually decrease to -1‰ in the lower Dalong Formation and are consistently low (around 0‰) in the middle Dalong to lowermost Feixianguan Formation. No clear δ15N shift is recognized across the extinction horizon. The consistently low δ15N values suggest the enhanced N fixation in the ocean during the Changhsingian at Chaotian. Composite profiles based on previous and the present studies demonstrate the substantial δ15N variation on a global scale in the late Permian to earliest Triassic; a systematic δ15N difference by low and high latitudes is particularly clarified. Although the enhanced N fixation throughout the

  7. Community stability and selective extinction during the Permian-Triassic mass extinction

    Roopnarine, Peter D.; Angielczyk, Kenneth D.


    The fossil record contains exemplars of extreme biodiversity crises. Here, we examined the stability of terrestrial paleocommunities from South Africa during Earth's most severe mass extinction, the Permian-Triassic. We show that stability depended critically on functional diversity and patterns of guild interaction, regardless of species richness. Paleocommunities exhibited less transient instability—relative to model communities with alternative community organization—and significantly greater probabilities of being locally stable during the mass extinction. Functional patterns that have evolved during an ecosystem's history support significantly more stable communities than hypothetical alternatives.

  8. Evaluating Non-potable Water Usage for Oil and Gas Purposes in the Permian Basin

    Marsac, K.; Pedrazas, M.; Suydam, S.; Navarre-Sitchler, A.


    Oil and gas company water usage is currently an area of extreme concern in the water stressed Western United States. 87% of the wells in Permian Basin are being drilled in areas of high or extreme water stress. Using recycled produced water or groundwater that does not meet the USDW drinking water standards for oil and gas purposes could assist in relieving both water stress and tension between oil and gas companies and the public. However, non-USDW drinking water (TDS over 10,000 ppm), has the potential to react with formation water causing mineral precipitation, reducing the permeability of the producing formation. To evaluate the potential of non-potable water usage in the Permian Basin, available groundwater chemistry data was compiled into a database. Data was collected from the NETL-run NATCARB database, the USGS Produced Water Database, and the Texas Railroad Commission. The created database went through a system of quality assurance and control for pH, TDS, depth and charge balance. Data was used to make a set of waters representative of Permian Basin groundwater based on TDS, Ca/Mg ratio and Cl/SO 4 ratio. Low, medium and high of these three characteristics; representing the 25 th , 50 th and 75 th percentile respectively; was used to make a matrix of 27 waters. Low TDS is 64,660 ppm, medium TDS is 98,486 ppm, and high TDS is 157,317 ppm. Ca/Mg ratios range from 1.98 to 7.26, and Cl/SO 4 ratios range from 32.96 to 62.34. Geochemical models of the mixing of these 27 waters with an average water were used to evaluate for possible precipitation. Initial results are positive, with the highest total precipitation being 2.371 cm 3 of dolomite and anhydrite in 2000 cm 3 of water with high TDS, high Ca/Mg ratio and low Cl/SO 4 ratio. This indicates a maximum of approximately 0.12% of porosity would be filled with mineral precipitation during the mixing of chosen Permian Basin waters.

  9. The use of perioperative serial blood lactate levels, the APACHE II and the postoperative MELD as predictors of early mortality after liver transplantation O uso da dosagem seriada do lactato sérico no perioperatório, do APACHE II e do MELD pós-operatório como preditores de mortalidade precoce após transplante hepático

    Anibal Basile-Filho


    Full Text Available PURPOSE: To evaluate the accuracy of different parameters in predicting early (one-month mortality of patients submitted to orthotopic liver transplantation (OLT. METHODS: This is a retrospective study of forty-four patients (38 males and 10 females, mean age of 52.2 ± 8.9 years admitted to the Intensive Care Unit of a tertiary hospital. Serial lactate blood levels, APACHE II, MELD post-OLT, creatinine, bilirubin and INR parameters were analyzed by receiver-operator characteristic (ROC curves as evidenced by the area under the curve (AUC. The level of significance was set at 0.05. RESULTS: The mortality of OLT patients within one month was 17.3%. Differences in blood lactate levels became statistically significant between survivors and nonsurvivors at the end of the surgery (pOBJETIVO: Avaliar qual parâmetro é o mais eficiente na predição de mortalidade precoce (um mês de pacientes submetidos a transplante ortotópico de fígado (OLT. MÉTODOS: Estudo retrospectivo em cinqüenta e oito pacientes adultos (44 homens e 14 mulheres, com uma idade média de 51,7 ± 10,1 anos admitidos na Unidade de Terapia Intensiva de um hospital terciário. Os parâmetros como a dosagem seriada de lactato no sangue, APACHE II, MELD pós-OLT, creatinina, bilirrubina e INR foram analisados por curvas ROC (Receiver-operator characteristic, evidenciado pela área abaixo da curva (AUC. O nível de significância foi definido em 0,05. RESULTADOS: A mortalidade dos pacientes OLT em até um mês foi de 17,3%. As diferenças no nível de lactato no sangue tornaram-se estatisticamente significantes entre sobreviventes e não sobreviventes no final da cirurgia (p < 0,05. A AUC foi de 0,726 (95%CI = 0,593-0,835 para APACHE II (p = 0,02; 0,770 (95%CI = 0,596-0,849 para o lactato sérico (L7-L8 (p = 0,03; 0,814 (95%CI = 0,690-0,904 para MELD post-OLT (p < 0,01; 0,550 (95%CI = 0,414-0,651 de creatinina (p = 0,64; 0,705 (95%CI = 0,571-0,818 de bilirrubina (p = 0,05 e 0

  10. Experimental use of alcohol in early adolescence: the 11-year follow-up of the 1993 Pelotas (Brazil birth cohort study Uso experimental de álcool no início da adolescência: a visita de 11 anos da coorte de nascimentos de Pelotas, Rio Grande do Sul, Brasil, 1993

    Ricardo B. Noal


    Full Text Available The aim of this study was to estimate the prevalence of experimental alcohol intake and associated factors in early adolescence. The overall sample consisted of 4,452 adolescents (mean age = 11.3 years; SD = 0.3 from the 1993 birth cohort in Pelotas, Rio Grande do Sul State, Brazil. Experimental use of alcohol was reported by 17.5% of the interviewees (95%CI: 16.3-18.6, and 5% reported having tried alcohol at nine years of age or younger. Prevalence of experimental alcohol use was higher among adolescents whose mothers had consumed alcohol during pregnancy, whose parents consumed alcohol, who worked outside the home themselves, and who had ever tried smoking. Family strife, parental alcohol intake, and adolescent smoking were strong predictors of experimental alcohol use in early adolescence. Special attention should be targeted to these groups in order to avoid heavy and premature alcohol use in early adolescence.Este trabalho teve como objetivo avaliar a prevalência de uso experimental de bebidas alcoólicas e fatores associados no início da adolescência. Compuseram a amostra 4.452 adolescentes (média = 11,3 anos; DP = 0,3 da coorte de nascimentos de 1993 de Pelotas, Rio Grande do Sul, Brasil. O uso experimental de álcool foi referido por 17,5% dos entrevistados (IC95%: 16,3-18,6 e 5% referiram ter experimentado com nove anos ou menos. O risco de uso experimental de álcool foi maior nos adolescentes cujas mães referiram ter bebido na gestação, quando ambos os pais usavam bebidas alcoólicas, entre os adolescentes que trabalhavam fora e naqueles que já experimentaram cigarro ou outras drogas. As relações familiares, o uso de álcool pelos pais, e o tabagismo do adolescente são os fatores fortemente associados ao uso experimental de álcool no início da adolescência. Mais atenção deveria ser dada especialmente a esses grupos, a fim de combater o consumo precoce de álcool.

  11. Geochronology and geochemistry of Permian bimodal volcanic rocks from central Inner Mongolia, China: Implications for the late Palaeozoic tectonic evolution of the south-eastern Central Asian Orogenic Belt

    Zhang, Zhicheng; Chen, Yan; Li, Ke; Li, Jianfeng; Yang, Jinfu; Qian, Xiaoyan


    Zircon U-Pb ages, geochemical data and Sr-Nd isotopic data are presented for volcanic rocks from the lower Permian Dashizhai Formation. These rocks are widely distributed in the south-eastern Central Asian Orogenic Belt in central Inner Mongolia, China. The volcanic rocks mainly consist of basaltic andesite and rhyolite, subordinate dacite and local andesite, and exhibit bimodal geochemical features. The results of zircon U-Pb dating indicate that the volcanic rocks formed during the early Permian (292-279 Ma). The mafic volcanic rocks belong to low-K tholeiitic to medium-K calc-alkaline series. These mafic volcanic rocks are also characterised by moderately enriched light rare earth element (LREE) patterns; high abundances of Th, U, Zr and Hf; negative Nb, Ta and Ti anomalies; initial 87Sr/86Sr ratios of 0.70514-0.70623; and positive εNd(t) values (+1.9 to +3.8). These features indicate that the mafic volcanic rocks were likely derived from the high-percentage partial melting of subduction-related metasomatised asthenospheric mantle. The felsic rocks show an A-type affinity, with enrichments in alkalis, Th, U and LREEs. The felsic rocks are depleted in Ba, Sr, Nb, Ta and Ti and exhibit moderately LREE-enriched patterns (LaN/YbN = 2.09-6.45) and strongly negative Eu anomalies (Eu/Eu∗ = 0.04-0.25). These features, along with the positive εNd(t) values (+2.6 to +7.7) and young TDM2 ages (TDM2 = 435-916 Ma), indicate that the felsic rocks were likely derived from a juvenile crustal source that mainly consisted of juvenile mid-ocean ridge basalt-related rocks. The volcanic association in this study and in previously published work widely distributed in central Inner Mongolia. The observations in this study suggest that the lower Permian volcanic rocks formed in an identical tectonic environment. The regional geological data indicate that the bimodal volcanic rocks from the lower Permian Dashizhai Formation in the study area formed in an extensional setting that was

  12. Magnetostratigraphy and high-resolution cyclic stratigraphy of the continental Permian-Triassic boundary interval (Central Germany)

    Szurlies, M.; Bachmann, G. H.; Menning, M.; Nowaczyk, N. R.; Käding, K.-C.


    The Buntsandstein represents the lower group of the tripartite classic Germanic Triassic supergroup. It is traditionally subdivided by lithological criteria into three subgroups. In its type area of Central Germany, the Lower Buntsandstein subgroup is approximately 300 m thick and consists of red beds, predominantly fine-grained sandstones, siltstones and claystones with interbedded oolitic limestones. The clastic sediments were deposited during Late Permian to Early Triassic times in fluvio-lacustrine environments of a large intracratonic basin. The oolites were formed in hypersaline playa lakes. The Lower Buntsandstein is further subdivided into the Calvörde Formation and the Bernburg Formation. The clastic-carbonatic succession shows a distinct cyclicity, pragmatically subdivided into 20 approximately 10--25 m thick small-scale fining-upward cycles. Commonly, these cycles have a sandy-oolitic base and a shaly upper part. The small-scale fining-upward cycles are correlation cycles that are obvious in wireline logs, cores and outcrops. With wireline logs (e.g., gamma-ray logs) they can be correlated over almost the entire Central European Basin from the Netherlands in the west via Germany to Poland in the east. The oolites form well correlatable litho-markers, indicating a great consistency of facies development. The thickness of the small-scale fining-upward cycles correspond to genetic small-scale cycles, which are considered to reflect climatic fluctuation of alternating drier and wetter periods due to solar-induced 100 ka Milankovitch eccentricity cycles. With a combined litho- and log-stratigraphic investigation, a precise stratigraphic positioning of paleomagnetic results has been realized. In the surroundings of the Harz Mountains nine sections (eight outcrops, one well) were investigated. For the paleomagnetic investigations, the fine-grained sandstones, siltstones and claystones as well as the oolitic limestones were sampled in 1--2 m intervals, yielding

  13. Research on Outcrop Sequence Stratigraphy of Permian in Middle-Lower Yangtze Region


    According to the latest International Chronostratigraphic Scheme (ICS, 2000), the Permian in the Middle-Lower Yangtze region of South China can be divided into three series and nine stages relevant to the traditional six stages of South China. From Assellian to Changxingian of Permian, 44 Ma in age range, the strata are composed of 14 third-order sequences, each of which is 3.14 Ma in average age range. There is one third-order sequence of Zisongian, equivalent to middle and upper Chuanshan Formation or equal to Asselian and two thirds of Sakmarian. There are two third-order sequences, corre sponding to Liang-shan Formation or Zhenjiang Formation and upper Chuanshan Formation, which are assigned to Longlingian, coinciding with Artinskian and one third of Sakmarian. In addition, three third-order sequences, equal to Qixia Formation, are attributed to Chihsian, corresponding to Kubergandian and one third of Roadian. Four third-order sequences, comprising Gufeng, Maokou, Yanqiao,Yinping and Wuxue formations, are assigned to Maokouan, equivalent to two thirds of Roadian, Word ian and Capitanian. Two third-order sequences, equal to Longtan Formation or Wujiaping Formation,are included in Wuchiapingian. Other two third-order sequences, corresponding to Changxing Formation or Dalong Formation, are assigned to Changhsingian. In brief, these above third-order sequences can be incorporated into 4 sequences sets.

  14. Bedout: a possible end-Permian impact crater offshore of northwestern Australia.

    Becker, L; Poreda, R J; Basu, A R; Pope, K O; Harrison, T M; Nicholson, C; Iasky, R


    The Bedout High, located on the northwestern continental margin of Australia, has emerged as a prime candidate for an end-Permian impact structure. Seismic imaging, gravity data, and the identification of melt rocks and impact breccias from drill cores located on top of Bedout are consistent with the presence of a buried impact crater. The impact breccias contain nearly pure silica glass (SiO2), fractured and shock-melted plagioclases, and spherulitic glass. The distribution of glass and shocked minerals over hundreds of meters of core material implies that a melt sheet is present. Available gravity and seismic data suggest that the Bedout High represents the central uplift of a crater similar in size to Chicxulub. A plagioclase separate from the Lagrange-1 exploration well has an Ar/Ar age of 250.1 +/- 4.5 million years. The location, size, and age of the Bedout crater can account for reported occurrences of impact debris in Permian-Triassic boundary sediments worldwide.

  15. Geochemistry of environmentally sensitive trace elements in Permian coals from the Huainan coalfield, Anhui, China

    Chen, J.; Liu, Gaisheng; Jiang, M.; Chou, C.-L.; Li, H.; Wu, B.; Zheng, Lingyun; Jiang, D.


    To study the geochemical characteristics of 11 environmentally sensitive trace elements in the coals of the Permian Period from the Huainan coalfield, Anhui province, China, borehole samples of 336 coals, two partings, and four roof and floor mudstones were collected from mineable coal seams. Major elements and selected trace elements were determined by inductively coupled plasma optical emission spectrometry (ICP-OES), inductively coupled plasma mass spectrometry (ICP-MS), and hydride generation atomic absorption spectrometry (HAAS). The depositional environment, abundances, distribution, and modes of occurrence of trace elements were investigated. Results show that clay and carbonate minerals are the principal inorganic constituents in the coals. A lower deltaic plain, where fluvial channel systems developed successively, was the likely depositional environment of the Permian coals in the Huainan coalfield. All major elements have wider variation ranges than those of Chinese coals except for Mg and Fe. The contents of Cr, Co, Ni, and Se are higher than their averages for Chinese coals and world coals. Vertical variations of trace elements in different formations are not significant except for B and Ba. Certain roof and partings are distinctly higher in trace elements than underlying coal bench samples. The modes of occurrence of trace elements vary in different coal seams as a result of different coal-forming environments. Vanadium, Cr, and Th are associated with aluminosilicate minerals, Ba with carbonate minerals, and Cu, Zn, As, Se, and Pb mainly with sulfide minerals. ?? 2011 Elsevier B.V.

  16. Extension of Cathaysian flora during the Permian. Climatic and paleogeographic constraints

    Fluteau, F.; Besse, J.; Broutin, J.; Berthelin, M.


    Recent paleobotanic investigations have been described during the Late Permian intermingled Cathaysian (South China block), Gondwanian and Euramerian flora assemblage in Oman (Arabian plate). By analogy with present rainforest flora, Cathaysian floral elements in northeastern Gondwana require permanent precipitation, even if a short dry season may be endured. On the contrary, Eurasian flora is most probably adapted to dryness. We investigate the temporal and spatial extension of this surprising flora assemblage using numerical climatic simulations and paleomagnetic reconstructions. We suggest that the settlement of Cathaysian flora was possible as soon as this area reached about 20°S. According to paleomagnetic data, the southeastern Arabian plate reached this latitude during Late Permian, which exactly corresponds to the age of the oldest discovered Cathaysian flora elements in Oman. Moreover, the paleomagnetic configuration of plates allows for easy Chinese to Gondwana connections using migrating blocks in the Tethys or island arcs. However, the climatic simulations suggest the presence of a dry season lasting about 5 months in southeastern Arabia. Such precipitation conditions would probably prevent the settlement of Cathaysian plants. We thus propose that the intermingled assemblage reflected the paleoenvironmental conditions: Cathaysian plants grew in a flood plain with perennial water supply, whereas Euramerian plants lived out of the flood plain over adjacent hills where water was only supplied by seasonal precipitation. Migration paths of these different floras are also discussed.

  17. Rapid expansion of oceanic anoxia immediately before the end-Permian mass extinction.

    Brennecka, Gregory A; Herrmann, Achim D; Algeo, Thomas J; Anbar, Ariel D


    Periods of oceanic anoxia have had a major influence on the evolutionary history of Earth and are often contemporaneous with mass extinction events. Changes in global (as opposed to local) redox conditions can be potentially evaluated using U system proxies. The intensity and timing of oceanic redox changes associated with the end-Permian extinction horizon (EH) were assessed from variations in (238)U/(235)U (δ(238)U) and Th/U ratios in a carbonate section at Dawen in southern China. The EH is characterized by shifts toward lower δ(238)U values (from -0.37‰ to -0.65‰), indicative of an expansion of oceanic anoxia, and higher Th/U ratios (from 0.06 to 0.42), indicative of drawdown of U concentrations in seawater. Using a mass balance model, we estimate that this isotopic shift represents a sixfold increase in the flux of U to anoxic facies, implying a corresponding increase in the extent of oceanic anoxia. The intensification of oceanic anoxia coincided with, or slightly preceded, the EH and persisted for an interval of at least 40,000 to 50,000 y following the EH. These findings challenge previous hypotheses of an extended period of whole-ocean anoxia prior to the end-Permian extinction.

  18. Methane release from igneous intrusion of coal during Late Permian extinction events

    Retallack, G.J.; Jahren, A.H. [University of Oregon, Eugene, OR (USA). Dept. of Geological Science


    Unusually large and locally variable carbon isotope excursions coincident with mass extinctions at the end of the Permian Period (253 Ma) and Guadalupian Epoch (260 Ma) can be attributed to methane outbursts to the atmosphere. Methane has isotopic values {delta}{sup 13}C low enough to reduce to feasible amounts the carbon required for isotopic mass balance. The duration of the carbon isotopic excursions and inferred methane releases are here constrained to < 10,000 yr by counting annual varves in lake deposits and by estimating peat accumulation rates. On paleogeographic maps, the most marked carbon isotope excursions form linear arrays back to plausible methane sources: end-Permian Siberian Traps and Longwood-Bluff intrusions of New Zealand and end-Guadalupian Emeishan Traps of China. Intrusion of coal seams by feeder dikes to flood basalts could create successive thermogenic methane outbursts of the observed timing and magnitude, but these are unreasonably short times for replenishment of marine or permafrost sources of methane. Methane released by fracturing and heating of coal during intrusion of large igneous provinces may have been a planetary hazard comparable with bolide impact.

  19. Filamentous cyanobacteria fossils and their significance in the Permian-Triassic boundary section at Laolongdong, Chongqing

    JIANG HongXia; WU YaSheng; CAI ChunFang


    The microbial communities blooming immediately after the end-Permian mass extinction represent abnormally extreme environments, and vary in different areas. In this study, filamentous cyanobacterial biota was found in the strata after the extinction in the famous Permian-Triassic boundary section at Laolongdong, Chongqing, southwest China. In thin sections, the filamentous cyanobacterial fossils are below 1 mm in length, and generally taper to one end, with the widest diameter up to 0.08 mm. Some of them are curved, indicating that they are soft in life. Their walls are composed of cryptocrystalline to microcrystalline calcites. The filaments have round cross section, and are internally filled with micrites and fine sparry calcites, which indicate that the filaments are originally empty. They are randomly dis-tributed in the rocks, but in some places, they tend to be distributed in radial pattern. The filamentous organisms are morphologically similar to Rivularia of Rivulariaceae, Cyanobacteria Phylum, but with calcified sheaths, and are tentatively regarded as an indeterminate new species in Rivularia: Rivularia sp. Cyanobacteria are photosynthetic autotrophic, and can survive in dysoxic condition. The blooming of this organism and the absence of other organisms may indicate that the environment was oxygen-deficient and shallow, since this photosynthetic autotrophic organism needed to live within photic zone.

  20. Catastrophic dispersion of coal fly ash into oceans during the latest Permian extinction

    Grasby, S.E.; Sanei, H.; Beauchamp, B. [Geological Survey Canada Calgary, Calgary, AB (Canada)


    During the latest Permian extinction about 250 Myr ago, more than 90% of marine species went extinct, and biogeochemical cycles were disrupted globally. The cause of the disruption is unclear, but a link between the eruption of the Siberian Trap flood basalts and the extinction has been suggested on the basis of the rough coincidence of the two events. The flood basalt volcanism released CO{sub 2}. In addition, related thermal metamorphism of Siberian coal measures and organic-rich shales led to the emission of methane, which would have affected global climate and carbon cycling, according to model simulations. This scenario is supported by evidence for volcanic eruptions and gas release in the Siberian Tunguska Basin, but direct indicators of coal combustion have not been detected. Here we present analyses of terrestrial carbon in marine sediments that suggest a substantial amount of char was deposited in Permian aged rocks from the Canadian High Arctic immediately before the mass extinction. Based on the geochemistry and petrology of the char, we propose that the char was derived from the combustion of Siberian coal and organic-rich sediments by flood basalts, which was then dispersed globally. The char is remarkably similar to modern coal fly ash, which can create toxic aquatic conditions when released as slurries. We therefore speculate that the global distribution of ash could have created toxic marine conditions.

  1. Microbial response to limited nutrients in shallow water immediately after the end-Permian mass extinction.

    Jia, C; Huang, J; Kershaw, S; Luo, G; Farabegoli, E; Perri, M C; Chen, L; Bai, X; Xie, S


    Previous work indicates that a variety of microbes bloomed in the oceans after the end-Permian faunal mass extinction, but evidence is sporadically documented. Thus, the nature and geographic distribution of such microbes and their associations are unclear, addressed in this study using a series of biomarker groups. On the basis of microbial biomarker records of the 2-methylhopane index, evidence is presented for cyanobacterial blooms in both the western and eastern Tethys Sea and in both shallow and deep waters, after the mass extinction. The enhanced relative abundance of C(28) (expressed by the C(28) /C(29) ratio of) regular steranes suggests a bloom of prasinophyte algae occurred immediately after the end-Permian faunal extinction, comparable with those observed in some other mass extinctions in Phanerozoic. Significantly, cyanobacteria and prasinophyte algae show a synchronized onset of bloom in the shallow water Bulla section, north Italy, inferring for the first time their coupled response to the biotic crisis and the associated environmental conditions. However, in Meishan of Zhejiang Province in south China, the bloom declined earlier than in Bulla. The association of increased 2-methylhopane index with a negative shift in the nitrogen isotope composition infers a scenario of enhanced nitrogen fixation by cyanobacteria immediately after the faunal mass extinction. N(2) fixation by cyanobacteria is here interpreted to have provided prasinophyte algae with ammonium in nutrient-limited shallow waters, and thus caused their associated blooms.

  2. The Siberian Traps and the End-Permian mass extinction: a critical review



    The association between the Siberian Traps,the largest continental flood basalt province,and thelargest-known mass extinction event at the end of the Permian period,has been strengthened by re-cently-published high-precision 40Ar/39Ar dates from widespread localities across the Siberian prov-ince[i].We argue that the impact of the volcanism was amplified by the prevailing late Permian envi-ronmental conditions-in particular,the hothouse climate,with sluggish oceanic circulation,that wasleading to widespread oceanic anoxia.Volcanism released large masses of sulphate aerosols andcarbon dioxide,the former triggering short-duration volcanic winters,the latter leading to long-termwarming.Whilst the mass of CO2 released from individual eruptions was small compared with the totalmass of carbon in the atmosphere-ocean system,the long 'mean lifetime' of atmospheric C02,com-pared with the eruption flux and duration,meant that significant accumulation could occur over periodsof 10s years.Compromise of the carbon sequestration systems (by curtailment of photosynthesis,de-struction of biomass,and warming and acidification of the oceans) probably led to rapid atmosphericCO2 build-up,warming,and shallow-water anoxia,leading ultimately to mass extinction.

  3. Calcium and calcium isotope changes during carbon cycle perturbations at the end-Permian

    Komar, Nemanja; Zeebe, Richard


    Negative carbon and calcium isotope excursions, as well as climate shifts, took place during the most severe mass extinction event in Earth's history, the end-Permian (˜252 Ma). Investigating the connection between carbon and calcium cycles during transient carbon cycle perturbation events, such as the end-Permian, may help resolve the intricacies between the coupled calcium-carbon cycles, as well as provide a tool for constraining the causes of mass extinction. Here, we identify the deficiencies of a simplified calcium model employed in several previous studies and we demonstrate the importance of a fully coupled carbon-cycle model when investigating the dynamics of carbon and calcium cycling. Simulations with a modified version of the LOSCAR model, which includes a fully coupled carbon-calcium cycle, indicate that increased weathering rates and ocean acidification (potentially caused by Siberian Trap volcanism) are not capable of producing trends observed in the record, as previously claimed. Our model results suggest that combined effects of carbon input via Siberian Trap volcanism (12,000 Pg C), the cessation of biological carbon export, and variable calcium isotope fractionation (due to a change in the seawater carbonate ion concentration) represents a more plausible scenario. This scenario successfully reconciles δ13C and δ44Ca trends observed in the sediment record, as well as the proposed warming of >6oC.

  4. A Proposed Area for Study of Accessory Section and Point of Terrestrial Permian-Triassic Boundary


    After the establishment of the global stratotype section and point (GSSP) of the PermianTriassic boundary (PTB), the definition of the accessory section and point (ASP) of the terrestrial Permian-Triassic boundary (TPTB) is now on the agenda. However, all good TPTB sections so far known have the following shortcomings: (1) the exact TPTB horizon is difficult to define paleontologically with high-resolution, and (2) accurate correlation between marine and terrestrial PTBs is hard to attain. In order to enhance the understanding of the nature of the global life crisis in both the marine and terrestrial environments across the Paleozoic-Mesozoic transition, these shortcomings need to be addressed. In western Guizhou and eastern Yunnan, Southwest China, some fossiliferous PTB sections which include marine, paralic and terrestrial are well-developed, allowing bed-to-bed correlation of the PTB sequences.Fortunately, the marine PTB sequence in this area is almost the same as found at the Meishan Section,where the GSSP of the PTB is located, which may provide a reliable auxiliary marker for high-resolution demarcation of the TPTB. These features found in western Guizhou and eastern Yunnan make this area a good place to study the ASP of the TPTB, so we propose to study the ASP of the TPTB in this area.

  5. Geochemical characteristics of the Permian sedimentary rocks from Qiangtang Basin: Constraints for paleoenvironment and paleoclimate

    Junjie Hu


    Full Text Available Qiangtang Basin is expected to become important strategic petroleum exploitation area in China. However, little research has been done on the Permian strata in this area. This paper presents Lower Permian Zhanjin Formation geochemical data from the Jiaomuri area, reconstructing the paleo-depositional environment and providing information for further petroleum exploration. The geochemical characteristics of 19 samples were investigated. These geochemical samples show a developed mud flat characteristic with light rich clay content. The geological data were used to constrain the paleoredox environment, which proved that these sediments were deposited mainly beneath a slightly oxic water column with relatively low paleoproductivity as evidenced by the P/Ti (mean of 0.07 and Ba/Al (mean of 20.5. Palaeoclimate indexes such as the C-value (0.24-1.75 and Sr/Cu (1.28-11.58 reveal a humid climatic condition during Zhanjin Formation sediment deposition. The ω(LaN/ω(YbN ratio values indicate a fast sedimentary rate during the deposition period.

  6. REE Geochemical Study of the Permian-Triassic Marine Sedimentary Environment in Guizhou Province

    南君亚; 刘丛强; 周德全; 王筑明


    This paper deals with the REE geochemical characteristics of Permian-Triassic marine carbonate rocks in Guizhou Province. It is found that there are two broad categories of stratigraphic units in the region studied in accordance with their ∑ REE contents and REE distribution patterns: one is characterized by LREE enrichment and slight Ce depletion, with the REE distribution patterns similar to those of North American shales, and the other features relative HREE enrichment and relatively remarkable Ce depletion, with the REE distribution patterns close to those of pelagic sediments. In terms of their different ∑ REE contents, five types of stratigraphic units can be distinguished. Incorporation of detrital minerals, REE complexing capability, oxidation-reduction conditions of the media are the main factors affecting the REE composition and REE distribution patterns of marine carbonate rocks in the region studied. In the light of REE geochemical characteristics of carbonate rocks, coupled with sedimentary facies analysis, this paper discusses the characteristics of the Permian-Triassic marine sedimentary environment in Guizhou Province and its evolutional rules.

  7. Permian Mengkarang coal facies and environment, based on organic petrology study

    Nana Suwarna


    Full Text Available Permian Mengkarang Coal Measures is situated in the middle part of Sumatera Island. Some fresh outcrop samples of the Permian Mengkarang coals have been analyzed both macroscopically and microscopically, to asses their depositional environment. On the basis of organic-petrological analysis, the coal seams show variation in the predominance of some macerals, indicating successions of environmental changes. The dominant maceral group is vitrinite, present in very low to very high values; whilst the minor one is inertinite showing low amount. Environmental information derived from the organic facies study shows that the coals were deposited in wet zone of mire, ranges from wet limnic-telmatic zone to telmatic wet forest swamp under rapid burial condition, due to rapid basin subsidence. The organic facies concept is thus applicable in basin studies context and has potential to become an additional tool for interpretation of depositional environment.    

  8. Application of seismic facies and attributes analysis on the identification of Permian igneous rock

    Xu Yongzhong; Yang Haijun; Liu Yongfu; Wang Shuangshuang; Yang Peng; Zhao Jixiang


    Seismic facies and attributes analysis techniques are introduced.The geological characteristics of some oil fields in western China are used in conjunction with drilling results and logging data to identify the lithology,intrusion periods,and distribution range of the Permian igneous rocks in this area.The lithologic classification,the vertical and horizontal distribution,and the intrusion periods of igneous rock were deduced through this study.Combining seismic facies and attributes analysis based on optimization can describe the igneous rock in detail.This is an efficient way to identify lithology and intrusion periods.Using geological data and GR-DT logging cross-plots the Permian igneous rock from TP to TT was divided into three periods.The lithology of the first period is tuff and clasolite with a thickness ranging from 18 to 80 ms.The second is basalt with a thickness ranging from 0 to 20 ms.The third is tuff and clasolite and dacite whose thickness ranges from 60 to 80 ms.These results can help understand the clasolite trap with low amplitude and the lithologic trap of the Carboniferous and Silurian.They can also guide further oil and/or gas exploration.

  9. Research on Genesis of Pyrite near the Permian-Triassic Boundary in Meishan, Zhejiang, China

    JIANG Yao-fa; TANG Yue-gang; CHOU Chen-lin


    The content and crystal forms of pyrite and sulfur isotope composition of pyrite sulfur as well as its vertical distribution near the Permian-Triassic (P/T) boundary in the Meishan section, Changxing county, Zhejiang province, China were studied using geological, petrological, mineralogical and geochemical methods (techniques). The result showed that the genesis of abundant pyrites in bed 24e2 at the uppermost part of the Changxing Formation in the Meishan section may be related to volcanic activity. In bed 24e2 of the Meishan section, pyrite has its highest content of 1.84% and the sulfur isotope composition has the highest δ34S value at +2.2‰ which is very similar to that of the average value of volcanic gas. There are some volcanic products such as β-quartz, siliceous cylinders and siliceous spherules which coexisted with pyrites in beds 24e2 and 24f. It can be concluded that a large quantity of volcanic ash fell into the South China Sea and was incorporated into marine sediments during the formation of limestone at the uppermost part of the Changxing Formation. The volcanic eruption with massive amounts of H2S and SO2 gas at the end of the Permian period resulted in the enrichment of H2S in the South China Sea areas. The reaction of H2S with reactive iron minerals formed the mass of abundant pyrites.

  10. Structure and evolution of Middle Permian palaeo-seamounts in Bayan Har and its adjacent area