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Sample records for cambrian period

  1. Tubicolous enteropneusts from the Cambrian period.

    Science.gov (United States)

    Caron, Jean-Bernard; Morris, Simon Conway; Cameron, Christopher B

    2013-03-28

    Hemichordates are a marine group that, apart from one monospecific pelagic larval form, are represented by the vermiform enteropneusts and minute colonial tube-dwelling pterobranchs. Together with echinoderms, they comprise the clade Ambulacraria. Despite their restricted diversity, hemichordates provide important insights into early deuterostome evolution, notably because of their pharyngeal gill slits. Hemichordate phylogeny has long remained problematic, not least because the nature of any transitional form that might serve to link the anatomically disparate enteropneusts and pterobranchs is conjectural. Hence, inter-relationships have also remained controversial. For example, pterobranchs have sometimes been compared to ancestral echinoderms. Molecular data identify enteropneusts as paraphyletic, and harrimaniids as the sister group of pterobranchs. Recent molecular phylogenies suggest that enteropneusts are probably basal within hemichordates, contrary to previous views, but otherwise provide little guidance as to the nature of the primitive hemichordate. In addition, the hemichordate fossil record is almost entirely restricted to peridermal skeletons of pterobranchs, notably graptolites. Owing to their low preservational potentials, fossil enteropneusts are exceedingly rare, and throw no light on either hemichordate phylogeny or the proposed harrimaniid-pterobranch transition. Here we describe an enteropneust, Spartobranchus tenuis (Walcott, 1911), from the Middle Cambrian-period (Series 3, Stage 5) Burgess Shale. It is remarkably similar to the extant harrimaniids, but differs from all known enteropneusts in that it is associated with a fibrous tube that is sometimes branched. We suggest that this is the precursor of the pterobranch periderm, and supports the hypothesis that pterobranchs are miniaturized and derived from an enteropneust-like worm. It also shows that the periderm was acquired before size reduction and acquisition of feeding tentacles, and

  2. The evolutionary and genetic origins of consciousness in the Cambrian Period over 500 million years ago

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    Todd E Feinberg

    2013-10-01

    Full Text Available Vertebrates evolved in the Cambrian Period before 520 million years ago, but we do not know when or how consciousness arose in the history of the vertebrate brain. Here we propose multiple levels of isomorphic or somatotopic neural representations as an objective marker for sensory consciousness. All extant vertebrates have these, so we deduce that consciousness extends back to the group’s origin. The first conscious sense may have been vision. Then vision, coupled with additional sensory systems derived from ectodermal placodes and neural crest, transformed primitive reflexive systems into image forming brains that map and perceive the external world and the body’s interior. We posit that the minimum requirement for sensory consciousness and qualia is a brain including a forebrain (but not necessarily a developed cerebral cortex/pallium, midbrain and hindbrain. This brain must also have 1 hierarchical systems of intercommunicating, isomorphically organized, processing nuclei that extensively integrate the different senses into representations that emerge in upper levels of the neural hierarchy; and 2 a widespread reticular formation that integrates the sensory inputs and contributes to attention, awareness, and neural synchronization. We propose a two-step evolutionary history, in which the optic tectum was the original center of multi-sensory conscious perception (as in fish and amphibians: step 1, followed by a gradual shift of this center to the dorsal pallium or its cerebral cortex (in mammals, reptiles, birds: step 2. We address objections to the hypothesis and call for more studies of fish and amphibians. In our view, the lamprey has all the neural requisites and is likely the simplest extant vertebrate with sensory consciousness and qualia. Genes that pattern the proposed elements of consciousness (isomorphism, neural crest, placodes have been identified in all vertebrates. Thus, consciousness is in the genes, some of which are

  3. Cambrian fans

    OpenAIRE

    Reading, Nathan; Speyer, David E.

    2006-01-01

    For a finite Coxeter group W and a Coxeter element c of W, the c-Cambrian fan is a coarsening of the fan defined by the reflecting hyperplanes of W. Its maximal cones are naturally indexed by the c-sortable elements of W. The main result of this paper is that the known bijection cl_c between c-sortable elements and c-clusters induces a combinatorial isomorphism of fans. In particular, the c-Cambrian fan is combinatorially isomorphic to the normal fan of the generalized associahedron for W. Th...

  4. Three-step modernization of the ocean:Modeling of carbon cycles and the revolution of ecological systems in the Ediacaran/Cambrian periods

    Institute of Scientific and Technical Information of China (English)

    Miyuki Tahata; Yusuke Sawaki; Yuichiro Ueno; Manabu Nishizawa; Naohiro Yoshida; Toshikazu Ebisuzaki; Tsuyoshi Komiya; ,Shigenori Maruyama

    2015-01-01

    Important ecological changes of the Earth (oxidization of the atmosphere and the ocean) increase in nutrient supply due to the break-up of the super continent (Rodinia) and the appearance of multi-cellular organisms (macroscopic algae and metazoan) took place in the Ediacaran period, priming the Cambrian explosion. The strong perturbations in carbon cycles in the ocean are recorded as excursions in carbonate and organic carbon isotope ratio (d13Ccarb and d13Corg) from the Ediacaran through early Cambrian periods. The Ediacaraneearly Cambrian sediment records of d13Ccarb and d13Corg, obtained from the drill-core samples in Three Gorges in South China, are compared with the results of numerical simulation of a sim-ple one-zone model of the carbon cycle of the ocean, which has two reservoirs (i.e., dissolved organic carbon (DOC) and dissolved inorganic carbon (DIC). The fluxes from the reservoirs are assumed to be proportional to the mass of the carbon reservoirs. We constructed a model, referred to here as the Best Fit Model (BFM), which reproduce d13Ccarb and d13Corg records in the Ediacaraneearly Cambrian period noted above. BFM reveals that the Shuram excursion is related to three major changes in the carbon cycle or the global ecological system of the Earth:(1) an increase in the coefficient of remineralization by a factor of ca. 100, possibly corresponding to a change in the dominant metabolism from anaerobic respiration to aerobic respiration, (2) an increase of carbon fractionation index from 25&to 33&, possibly corresponding to the change in the primary producer from rock-living cyanobacteria to free-living macro algae, and (3) an in-crease in the coefficient of the organic carbon burial by a factor of ca. 100, possibly corresponding to the onset of a biological pump driven by the flourishing metazoan and zooplankton. The former two changes took place at the start of the Shuram excursion, while the third occurred at the end of the Shuram excursion. The other

  5. Is a Cambrian Explosion Coming for Robotics?

    OpenAIRE

    Gill A. Pratt

    2015-01-01

    About half a billion years ago, life on earth experienced a short period of very rapid diversification called the "Cambrian Explosion." Many theories have been proposed for the cause of the Cambrian Explosion, one of the most provocative being the evolution of vision, allowing animals to dramatically increase their ability to hunt and find mates. Today, technological developments on several fronts are fomenting a similar explosion in the diversification and applicability of robotics. Many of ...

  6. The Cambrian impact hypothesis

    CERN Document Server

    Zhang, Weijia

    2008-01-01

    After a thorough research on the circumstantial changes and the great evolution of life in the Cambrian period, the author propounds such a hypothesis: During the Late Precambrian, about 500-600Ma, a celestial body impacted the Earth. The high temperature ended the great glaciation, facilitated the communication of biological information. The rapid change of Earth environment enkindled the genesis-control system, and released the HSP-90 variations. After the impact, benefited from the protection of the new ozone layer and the energy supplement of the aerobic respiration, those survived underground life exploded. They generated carapaces and complex metabolism to adjust to the new circumstance of high temperature and high pressure. This article uses a large amount of analyses and calculations, and illustrates that this hypothesis fits well with most of the important incidences in astronomic and geologic discoveries.

  7. The Cambrian “explosion”: Slow-fuse or megatonnage?

    OpenAIRE

    Conway Morris, Simon

    2000-01-01

    Clearly, the fossil record from the Cambrian period is an invaluable tool for deciphering animal evolution. Less clear, however, is how to integrate the paleontological information with molecular phylogeny and developmental biology data. Equally challenging is answering why the Cambrian period provided such a rich interval for the redeployment of genes that led to more complex bodyplans.

  8. Tectonic Events May Have Triggered the Cambrian Explosion

    Science.gov (United States)

    Wendel, JoAnna

    2014-11-01

    Major geological changes causing sea level rise at the start of the Cambrian period (540-490 million years ago) could have kick-started the Cambrian Explosion—a geological time period when most major phyla of life suddenly appeared in the fossil record. A paper published in the November issue of Geology (doi:10.1130/G35886.1) proposes a new geological mechanism for this event.

  9. Nomenclature of Cambrian epochs and series based on GSSPs-Comments on an alternative proposal by Rowland and Hicks

    Institute of Scientific and Technical Information of China (English)

    Shanchi Peng; Loren E. Babcock; Gerd Geyer; Malgorzata Moczydlowska

    2006-01-01

    @@ In this issue, Rowland & Hicks advocate a nomenclature for the four epochs of the Cambrian Period that carries over the terms Early, Middle, and Late Cambrian from previous usage (but with new definitions) and introduces a non-geographically-based term, Ichnocambrian, for the earliest Cambrian epoch.

  10. The Cambrian explosion.

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    Briggs, Derek E G

    2015-10-01

    The sudden appearance of fossils that marks the so-called 'Cambrian explosion' has intrigued and exercised biologists since Darwin's time. In On the Origin of Species, Darwin made it clear that he believed that ancestral forms 'lived long before' their first fossil representatives. While he considered such an invisible record necessary to explain the level of complexity already seen in the fossils of early trilobites, Darwin was at a loss to explain why there were no corresponding fossils of these earlier forms. In chapter 9 of the Origin, entitled 'On the imperfection of the geological record', he emphasized the 'poorness of our palaeontological collections' and stated categorically that 'no organism wholly soft can be preserved'. Fortunately much has been discovered in the last 150 years, not least multiple examples of Cambrian and Precambrian soft-bodied fossils. We now know that the sudden appearance of fossils in the Cambrian (541-485 million years ago) is real and not an artefact of an imperfect fossil record: rapid diversification of animals coincided with the evolution of biomineralized shells. And although fossils in earlier rocks are rare, they are not absent: their rarity reflects the low diversity of life at this time, as well as the low preservation potential of Precambrian organisms (see Primer by Butterfield, in this issue). PMID:26439348

  11. At the Origin of Animals: The Revolutionary Cambrian Fossil Record

    OpenAIRE

    Budd, Graham E.

    2013-01-01

    The certain fossil record of animals begins around 540 million years ago, close to the base of the Cambrian Period. A series of extraordinary discoveries starting over 100 years ago with Walcott’s discovery of the Burgess Shale has accelerated in the last thirty years or so with the description of exceptionally-preserved Cambrian fossils from around the world. Such deposits of “Burgess Shale Type” have been recently complemented by other types of exceptional preservation. Together with a rema...

  12. Developmental sequence of Cambrian embryo Markuelia

    Institute of Scientific and Technical Information of China (English)

    DONG XiPing

    2007-01-01

    Based on more exquisitely preserved specimens of Markuelia hunanensis recently recovered from Middle and Upper Cambrian in western Hunan and in the light of Synchrotron radiation X-ray tomographic microscopy, the developmental sequence from cleavage through organogenesis to the pre-hatching of Cambrian embryo Markuelia, especially the developmental sequence during the pre-hatching stage, i.e. from the earliest period when the scalids and tail spines only took shape to the latest period (just about hatching), is established. This developmental sequence provides a pattern of embryonic development during the pre-hatching stage, which has not been established in the living scalidophorans (priapulids, Ioriciferans and kinorhynchs). Thus, it not only enriches our knowledge on the embryonic development of the extant descendants of Markuelia, but also opens a new window to the evolution and development of the animal.

  13. Did Gamma Ray Burst Induce Cambrian Explosion?

    CERN Document Server

    Chen, Pisin

    2014-01-01

    One longstanding mystery in bio-evolution since Darwin's time is the origin of the Cambrian explosion that happened around 540 million years ago (Mya), where an extremely rapid increase of species occurred. Here we suggest that a nearby GRB event ~500 parsecs away, which should occur about once per 5 Gy, might have triggered the Cambrian explosion. Due to a relatively lower cross section and the conservation of photon number in Compton scattering, a substantial fraction of the GRB photons can reach the sea level and would induce DNA mutations in organisms protected by a shallow layer of water or soil, thus expediting the bio-diversification. This possibility of inducing genetic mutations is unique among all candidate sources for major incidents in the history of bio-evolution. A possible evidence would be the anomalous abundance of certain nuclear isotopes with long half-lives transmuted by the GRB photons in geological records from the Cambrian period. Our notion also imposes constraints on the evolution of ...

  14. Did gamma ray burst induce Cambrian explosion?

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    Chen, Pisin; Ruffini, R.

    2015-06-01

    One longstanding mystery in bio-evolution since Darwin's time is the origin of the Cambrian explosion that happened around 540 million years ago (Mya), where an extremely rapid increase of species occurred. Here we suggest that a nearby GRB event 500 parsecs away, which should occur about once per 5 Gy, might have triggered the Cambrian explosion. Due to a relatively lower cross section and the conservation of photon number in Compton scattering, a substantial fraction of the GRB photons can reach the sea level and would induce DNA mutations in organisms protected by a shallow layer of water or soil, thus expediting the bio-diversification. This possibility of inducing genetic mutations is unique among all candidate sources for major incidents in the history of bio-evolution. A possible evidence would be the anomalous abundance of certain nuclear isotopes with long half-lives transmuted by the GRB photons in geological records from the Cambrian period. Our notion also imposes constraints on the evolution of exoplanet organisms and the migration of panspermia.

  15. Sortable elements and Cambrian lattices

    OpenAIRE

    Reading, Nathan

    2005-01-01

    We show that the Coxeter-sortable elements in a finite Coxeter group W are the minimal congruence-class representatives of a lattice congruence of the weak order on W. We identify this congruence as the Cambrian congruence on W, so that the Cambrian lattice is the weak order on Coxeter-sortable elements. These results exhibit W-Catalan combinatorics arising in the context of the lattice theory of the weak order on W.

  16. Tropical shoreline ice in the late Cambrian: Implications for earth's climate between the Cambrian Explosion and the Great Ordovician Biodiversification Event

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    Runkel, Anthony C.; MacKey, T.J.; Cowan, Clinton A.; Fox, David L.

    2010-01-01

    Middle to late Cambrian time (ca. 513 to 488 Ma) is characterized by an unstable plateau in biodiversity, when depauperate shelf faunas suffered repeated extinctions. This poorly understood interval separates the Cambrian Explosion from the Great Ordovician Biodiversification Event and is generally regarded as a time of sustained greenhouse conditions. We present evidence that suggests a drastically different climate during this enigmatic interval: Features indicative of meteoric ice are well preserved in late Cambrian equatorial beach deposits that correspond to one of the shelf extinction events. Thus, the middle to late Cambrian Earth was at least episodically cold and might best be considered a muted analogue to the environmental extremes that characterized the Proterozoic, even though cooling in the two periods may have occurred in response to different triggers. Such later Cambrian conditions may have significantly impacted evolution preceding the Ordovician radiation.

  17. A suspension-feeding anomalocarid from the Early Cambrian.

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    Vinther, Jakob; Stein, Martin; Longrich, Nicholas R; Harper, David A T

    2014-03-27

    Large, actively swimming suspension feeders evolved several times in Earth's history, arising independently from groups as diverse as sharks, rays and stem teleost fishes, and in mysticete whales. However, animals occupying this niche have not been identified from the early Palaeozoic era. Anomalocarids, a group of stem arthropods that were the largest nektonic animals of the Cambrian and Ordovician periods, are generally thought to have been apex predators. Here we describe new material from Tamisiocaris borealis, an anomalocarid from the Early Cambrian (Series 2) Sirius Passet Fauna of North Greenland, and propose that its frontal appendage is specialized for suspension feeding. The appendage bears long, slender and equally spaced ventral spines furnished with dense rows of long and fine auxiliary spines. This suggests that T. borealis was a microphagous suspension feeder, using its appendages for sweep-net capture of food items down to 0.5 mm, within the size range of mesozooplankton such as copepods. Our observations demonstrate that large, nektonic suspension feeders first evolved during the Cambrian explosion, as part of an adaptive radiation of anomalocarids. The presence of nektonic suspension feeders in the Early Cambrian, together with evidence for a diverse pelagic community containing phytoplankton and mesozooplankton, indicate the existence of a complex pelagic ecosystem supported by high primary productivity and nutrient flux. Cambrian pelagic ecosystems seem to have been more modern than previously believed. PMID:24670770

  18. A vanished history of skeletonization in Cambrian comb jellies.

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    Ou, Qiang; Xiao, Shuhai; Han, Jian; Sun, Ge; Zhang, Fang; Zhang, Zhifei; Shu, Degan

    2015-07-01

    Ctenophores are traditionally regarded as "lower" metazoans, sharing with cnidarians a diploblastic grade of organization. Unlike cnidarians, where skeletonization (biomineralization and sclerotization) evolved repeatedly among ecologically important taxa (for example, scleractinians and octocorals), living ctenophores are characteristically soft-bodied animals. We report six sclerotized and armored ctenophores from the early Cambrian period. They have diagnostic ctenophore features (for example, an octamerous symmetry, oral-aboral axis, aboral sense organ, and octaradially arranged ctene rows). Unlike most modern counterparts, however, they lack tentacles, have a sclerotized framework, and have eight pairs of ctene rows. They are resolved as a monophyletic group (Scleroctenophora new class) within the ctenophores. This clade reveals a cryptic history and sheds new light on the early evolution of this basal animal phylum. Skeletonization also occurs in some other Cambrian animal groups whose extant members are exclusively soft-bodied, suggesting the ecological importance of skeletonization in the Cambrian explosion. PMID:26601209

  19. The geobiology of the extremely enriched polymetallic sulfides in the black shale of the lower Cambrian Niutitang formation, Southwestern China

    OpenAIRE

    Xu, Jun; 徐俊

    2014-01-01

    The Precambrian-Cambrian transition is a period with enormous geological and biological changes. There is a wide distribution of black shale sequence in the Late Sinian and Early Cambrian strata along the passive southern margin of the Yangtze Platform in South China. The remarkable polymetallic sulfide extremely enriched ore layer is embedded at the bottom of the Lower Cambrian Niutitang Formation, but its genesis remains highly disputable. Known mechanisms can hardly explain the extreme enr...

  20. A vanished history of skeletonization in Cambrian comb jellies

    OpenAIRE

    Ou, Qiang; Xiao, Shuhai; Han, Jian; Sun, Ge; Zhang, Fang; Zhang, Zhifei; Shu, Degan

    2015-01-01

    Ctenophores are traditionally regarded as “lower” metazoans, sharing with cnidarians a diploblastic grade of organization. Unlike cnidarians, where skeletonization (biomineralization and sclerotization) evolved repeatedly among ecologically important taxa (for example, scleractinians and octocorals), living ctenophores are characteristically soft-bodied animals. We report six sclerotized and armored ctenophores from the early Cambrian period. They have diagnostic ctenophore features (for exam...

  1. Fossils, molecules and embryos: new perspectives on the Cambrian explosion

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    Valentine, J. W.; Jablonski, D.; Erwin, D. H.

    1999-01-01

    The Cambrian explosion is named for the geologically sudden appearance of numerous metazoan body plans (many of living phyla) between about 530 and 520 million years ago, only 1.7% of the duration of the fossil record of animals. Earlier indications of metazoans are found in the Neoproterozic; minute trails suggesting bilaterian activity date from about 600 million years ago. Larger and more elaborate fossil burrows appear near 543 million years ago, the beginning of the Cambrian Period. Evidence of metazoan activity in both trace and body fossils then increased during the 13 million years leading to the explosion. All living phyla may have originated by the end of the explosion. Molecular divergences among lineages leading to phyla record speciation events that have been earlier than the origins of the new body plans, which can arise many tens of millions of years after an initial branching. Various attempts to date those branchings by using molecular clocks have disagreed widely. While the timing of the evolution of the developmental systems of living metazoan body plans is still uncertain, the distribution of Hox and other developmental control genes among metazoans indicates that an extensive patterning system was in place prior to the Cambrian. However, it is likely that much genomic repatterning occurred during the Early Cambrian, involving both key control genes and regulators within their downstream cascades, as novel body plans evolved.

  2. Causes of the Cambrian Explosion.

    OpenAIRE

    Smith, M P; Harper, D.A.T.

    2013-01-01

    In the last decade, at least thirty individual hypotheses have been invoked to explain the Cambrian Explosion, ranging from starbursts in the Milky Way to intrinsic genomic reorganization and developmental patterning. It has been noted (1) that recent hypotheses fall into three categories: a) developmental/genetic, b) ecologic and c) abiotic environmental, with geochemical hypotheses forming an abundant and distinctive subset of the last. With a few notable exceptions, a significant majority ...

  3. The base of the Cambrian in Morocco

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    Sdzuy, K.; Geyer, G.

    The paper summarizes problems of datings of Latest Precambrian to Early Cambrian rocks of Morocco. The known palaeontological data are reviewed briefly. Characteristics and correlatability of the earliest trilobites of Morocco are discussed. Additional remarks concern the appearance of hard part animals at the beginning of the Cambrian.

  4. Hydrogen sulphide release to surface waters at the Precambrian/Cambrian boundary.

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    Wille, Martin; Nägler, Thomas F; Lehmann, Bernd; Schröder, Stefan; Kramers, Jan D

    2008-06-01

    Animal-like multicellular fossils appeared towards the end of the Precambrian, followed by a rapid increase in the abundance and diversity of fossils during the Early Cambrian period, an event also known as the 'Cambrian explosion'. Changes in the environmental conditions at the Precambrian/Cambrian transition (about 542 Myr ago) have been suggested as a possible explanation for this event, but are still a matter of debate. Here we report molybdenum isotope signatures of black shales from two stratigraphically correlated sample sets with a depositional age of around 542 Myr. We find a transient molybdenum isotope signal immediately after the Precambrian/Cambrian transition. Using a box model of the oceanic molybdenum cycle, we find that intense upwelling of hydrogen sulphide-rich deep ocean water best explains the observed Early Cambrian molybdenum isotope signal. Our findings suggest that the Early Cambrian animal radiation may have been triggered by a major change in ocean circulation, terminating a long period during which the Proterozoic ocean was stratified, with sulphidic deep water. PMID:18509331

  5. Armored kinorhynch-like scalidophoran animals from the early Cambrian.

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    Zhang, Huaqiao; Xiao, Shuhai; Liu, Yunhuan; Yuan, Xunlai; Wan, Bin; Muscente, A D; Shao, Tiequan; Gong, Hao; Cao, Guohua

    2015-01-01

    Morphology-based phylogenetic analyses support the monophyly of the Scalidophora (Kinorhyncha, Loricifera, Priapulida) and Nematoida (Nematoda, Nematomorpha), together constituting the monophyletic Cycloneuralia that is the sister group of the Panarthropoda. Kinorhynchs are unique among living cycloneuralians in having a segmented body with repeated cuticular plates, longitudinal muscles, dorsoventral muscles, and ganglia. Molecular clock estimates suggest that kinorhynchs may have diverged in the Ediacaran Period. Remarkably, no kinorhynch fossils have been discovered, in sharp contrast to priapulids and loriciferans that are represented by numerous Cambrian fossils. Here we describe several early Cambrian (~535 million years old) kinorhynch-like fossils, including the new species Eokinorhynchus rarus and two unnamed but related forms. E. rarus has characteristic scalidophoran features, including an introvert with pentaradially arranged hollow scalids. Its trunk bears at least 20 annuli each consisting of numerous small rectangular plates, and is armored with five pairs of large and bilaterally placed sclerites. Its trunk annuli are reminiscent of the epidermis segments of kinorhynchs. A phylogenetic analysis resolves E. rarus as a stem-group kinorhynch. Thus, the fossil record confirms that all three scalidophoran phyla diverged no later than the Cambrian Period. PMID:26610151

  6. The Cambrian Evolutionary Explosion: Novel Evidence from Fossils Studied by X-ray Tomography

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    Chen, Jun-Yuan [Nanjing University, China

    2011-06-01

    The Cambrian explosion (from 542 million years to 488 million years ago) is one of the greatest mysteries in evolutionary biology. It wasn't until this period that complex organisms became common and diverse. the magnitude of the event can be understood based on the contrast between the biota and the degree of diversity of the fossils from both sides. great advances have been made in Cambrian palaeontology over the past century, especially the discovery of the well-preserved soft-bodied fauna from the Middle Cambrian Burgess Shale and the Lower Cambrian Maotianshan Shale deposits. The Cambrian side of the "Cambrian explosion" is richly illustrated and contrasts greatly with the Precambrian side. The study of these extraordinarily preserved fossil biota is extremely difficult. A major challenge is 3-D reconstruction and determining the patter of the cell organization in Weng'an embryos and their buried structures in Maotianshan Shale fossils. This talk will show that two recent technological approaches, propagation phase contrast synchrotron x-ray microtomography and microtomography, provide unique analytical tools that permit the nondestructive computational examination and visualization of the internal and buried characters in virtual sections in any plane, and virtual 3-D depictions of internal structures.

  7. The Ordovician Radiation: A Follow-up to the Cambrian Explosion?

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    Droser, Mary L; Finnegan, Seth

    2003-02-01

    There was a major diversification known as the Ordovician Radiation, in the period immediately following the Cambrian. This event is unique in taxonomic, ecologic and biogeographic aspects.While all of the phyla but one were established during the Cambrian explosion, taxonomic increases during the Ordovician were manifest at lower taxonomic levels although ordinal level diversity doubled. Marine family diversity tripled and within clade diversity increases occurred at the genus and species levels. The Ordovician radiation established the Paleozoic Evolutionary Fauna; those taxa which dominated the marine realm for the next 250 million years. Community structure dramatically increased in complexity. New communities were established and there were fundamental shifts in dominance and abundance.Over the past ten years, there has been an effort to examine this radiation at different scales. In comparison with the Cambrian explosion which appears to be more globally mediated, local and regional studies of Ordovician faunas reveal sharp transitions with timing and magnitudes that vary geographically. These transitions suggest a more episodic and complex history than that revealed through synoptic global studies alone.Despite its apparent uniqueness, we cannot exclude the possibility that the Ordovician radiation was an extension of Cambrian diversity dynamics. That is, the Ordovician radiation may have been an event independent of the Cambrian radiation and thus requiring a different set of explanations, or it may have been the inevitable follow-up to the Cambrian radiation. Future studies should focus on resolving this issue. PMID:21680422

  8. Compilation and network analyses of cambrian food webs.

    Directory of Open Access Journals (Sweden)

    Jennifer A Dunne

    2008-04-01

    Full Text Available A rich body of empirically grounded theory has developed about food webs--the networks of feeding relationships among species within habitats. However, detailed food-web data and analyses are lacking for ancient ecosystems, largely because of the low resolution of taxa coupled with uncertain and incomplete information about feeding interactions. These impediments appear insurmountable for most fossil assemblages; however, a few assemblages with excellent soft-body preservation across trophic levels are candidates for food-web data compilation and topological analysis. Here we present plausible, detailed food webs for the Chengjiang and Burgess Shale assemblages from the Cambrian Period. Analyses of degree distributions and other structural network properties, including sensitivity analyses of the effects of uncertainty associated with Cambrian diet designations, suggest that these early Paleozoic communities share remarkably similar topology with modern food webs. Observed regularities reflect a systematic dependence of structure on the numbers of taxa and links in a web. Most aspects of Cambrian food-web structure are well-characterized by a simple "niche model," which was developed for modern food webs and takes into account this scale dependence. However, a few aspects of topology differ between the ancient and recent webs: longer path lengths between species and more species in feeding loops in the earlier Chengjiang web, and higher variability in the number of links per species for both Cambrian webs. Our results are relatively insensitive to the exclusion of low-certainty or random links. The many similarities between Cambrian and recent food webs point toward surprisingly strong and enduring constraints on the organization of complex feeding interactions among metazoan species. The few differences could reflect a transition to more strongly integrated and constrained trophic organization within ecosystems following the rapid

  9. Ichnofossil from the Cambrian succession of Parahio Valley, Spiti Basin, India: Their stratigraphic and paleoenvironmental significance

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    Pandey, Shivani; Parcha, Suraj Kumar

    2013-04-01

    The Spiti Basin exposes an excellent section of Neoproterozoic- Cretaceous rocks in the Tethyan Himalaya of Himachal Pradesh. The diverse assemblage of ichnofossils is present in the Cambrian succession of Parahio Valley in the Spiti Basin. In the present study nineteen ichnofossils are reported from the Cambrian succession of Parahio Valley. The ichnofossils includes Bergaueria, Chondrites, Cruziana, Didymaulichnus, Dimorphichnus, Diplichnites, Helminthorhaphe, Merostomichnites, ?Monocraterion, Monomorphichnus, Nereites, Palaeopascichnus, Palaeophycus, Phycodes, Planolites, Rusophycus, Skolithos, Scolicia, Treptichnus etc. along with annelid worm, burrow and scratch marks. The described ichnofossil assemblage indicates that the ichnocenosis is dominated by a high behavioral diversity ranging from suspension to deposit feeders. It seems that the ichnofauna present in the Cambrian succession of this section were mostly produced by trilobite and arthropods, whereas some of them were produced by crustacean, priapulid worm, polychaetes and polyphyletic vermiforms. The distribution pattern of ichnofossils shows increase in taxonomic and morphological diversity up in the section. It further indicates that the availability of nutrients significantly increased their abundance as well as spatial distribution during Cambrian. The presence of Chondrites, Treptichnus, and Phycodes at the basal part of the Cambrian indicates shallow to deep environment with anaerobic condition. Whereas, the complex forms like Rusophycus, Cruziana, Monomorphichnus and Nereites represent shelf to slope environment. The appearance of Skolithos in the upper part reflects well oxygenated high energy condition. The environmental changes in the Parahio Valley during Cambrian period was distinctly marked by an anaerobic to aerobic condition and by a faunal change from endobenthic, soft - bodied, deposit feeders to epibenthic grazers. The present ichnofossils indicates that these sediments were

  10. Phosphate Biomineralization of Cambrian Microorganisms

    Science.gov (United States)

    McKay, David S.; Rozanov, Alexei Yu.; Hoover, Richard B.; Westall, Frances

    1998-01-01

    As part of a long term study of biological markers (biomarkers), we are documenting a variety of features which reflect the previous presence of living organisms. As we study meteorites and samples returned from Mars, our main clue to recognizing possible microbial material may be the presence of biomarkers rather than the organisms themselves. One class of biomarkers consists of biominerals which have either been precipitated directly by microorganisms, or whose precipitation has been influenced by the organisms. Such microbe-mediated mineral formation may include important clues to the size, shape, and environment of the microorganisms. The process of fossilization or mineralization can cause major changes in morphologies and textures of the original organisms. The study of fossilized terrestrial organisms can help provide insight into the interpretation of mineral biomarkers. This paper describes the results of investigations of microfossils in Cambrian phosphate-rich rocks (phosphorites) that were found in Khubsugul, Northern Mongolia.

  11. Did Gamma Ray Burst Induce Cambrian Explosion?

    OpenAIRE

    Chen, Pisin; Ruffini, Remo

    2014-01-01

    One longstanding mystery in bio-evolution since Darwin's time is the origin of the Cambrian explosion that happened around 540 million years ago (Mya), where an extremely rapid increase of species occurred. Here we suggest that a nearby GRB event ~500 parsecs away, which should occur about once per 5 Gy, might have triggered the Cambrian explosion. Due to a relatively lower cross section and the conservation of photon number in Compton scattering, a substantial fraction of the GRB photons can...

  12. Diversity partitioning during the Cambrian radiation

    OpenAIRE

    Na, Lin; Kiessling, Wolfgang

    2015-01-01

    Many mechanisms of the Cambrian Explosion have been proposed, but rigorous quantitative analyses of biodiversity dynamics are scarce, although they may shed light on important factors. Using a comprehensive database and sampling standardization, we dissect global diversity patterns. The trajectories of within-community, between-community, and global diversity during the main phase of the Cambrian radiation revealed a low-competition model, which was probably governed by niche contraction and ...

  13. Relationship between the Neoproterozoic snowball Earth and Cambrian explosion

    Science.gov (United States)

    Maruyama, S.; Yoshihara, A.; Isozaki, Y.

    2007-12-01

    Origin of snowball Earth has been debated in terms of greenhouse gas (e.g., Hoffman and Schrag), obliqueness of Earth's rotation axis (Williams, 1975), true polar wander (Evans, 2003), Galactic cosmic ray radiation (Shaviv and Veizer, 2003; Svensmark, 2006), or weakened geomagnetism (Maruyama and Yoshihara, 2003). A major difficulty for the greenhouse gas hypothesis is the on-off switch causing decrease and increase of appropriate amounts of CO2 by plume- and plate tectonics, and also in available amount of CO2 in atmosphere to be consistent with the observations. In contrast, the cosmic ray radiation models due to the star burst peaked at 2.5- 2.1 Ga and 1.4-0.8 Ga can explain on-off switch more easily than the greenhouse gas model. Cosmic ray radiations, however, must be modified by the geomagnetic intensity, fluctuating 150"% to cause the snowball Earth. Time difference between the Neoproterozoic snowball Earth and Cambrian explosion is as large as 250 millions years, and this refuses their direct close-relationship. Role of frequent mass extinctions, i.e., 8 times during 100 m.y. from 585 Ma to 488 Ma, during the Ediacaran and Cambrian, has been proposed (Zhu et al., 2007). This frequency is one order of magnitude higher compared to that in the post-Ordovician time. Yet, the Cambrian explosion cannot be explained by mass extinction which replaced the vacant niches shortly after the mass extinction and never created a new animal with a new body plan. A new model proposed herein is derived from weakened geomagnetism and resultant extensive cosmic radiation to alter gene and genome for a long period over advancement of low magnetic intensity and cosmic radiations (Svensmark, 2006) from 1.2-0.8Ga. As to the new body plans of animals, it took an appreciably long time to prepare all 34 genometypes before the apparent Cambrian explosion. Geochemically extreme conditions and widened shallow marine environment on continental shelf by the return-flow of sweater into

  14. The onset of the 'Ordovician Plankton Revolution' in the late Cambrian

    DEFF Research Database (Denmark)

    Servais, Thomas; Perrier, Vincent; Danelian, Taniel;

    2016-01-01

    The 'Great Ordovician Biodiversification Event' comprises the rapid diversification of marine organisms during the Ordovician Period. It is now clear that this adaptive radiation started for some organisms already in the Cambrian and continued for others beyond the end of the Ordovician, making...... the 'Great Ordovician Biodiversification Event' part of a long-term late Proterozoic and Early Palaeozoic radiation, that in part is expressed by the fossil record as the 'Cambrian Explosion.' A significant diversification of different groups of the plankton is observed in the late Cambrian-Early Ordovician...... interval, leading to the subsequent 'Ordovician Plankton Revolution.' The possible causes of this 'plankton revolution' are currently debated. They include changes in palaeoclimate, palaeogeography or tectonic and volcanic activity, as well as a modified nutrient supply. In this context, the Steptoean...

  15. Possible impact event in the Late Cambrian

    Institute of Scientific and Technical Information of China (English)

    2007-01-01

    The repeated investigation of the Batyrbay section has been done by the authors. Today we see increasing of interest to revising of volumes and boundaries of Stages and Series in the Cambrian and Ordovician,as well as to the events of this time. In this work the description of iron spherules and particles, found in the deep-water limestone layer of the conodont Cordylodus primitivus Zone in the middle Upper Cambrian of the Batyrbay section, Malyi Karatau of South Kazakhstan, and formed during the time of global sea-level changes known as a world-wide Lange Ranch Eustatic event, have been done. From our point of view, finds of iron spherules and particles may evidence about new unknown yet Event of cosmic origin, i.e. falling of a meteorite on Earth in the middle Late Cambrian.

  16. An Early Cambrian tunicate from China.

    Science.gov (United States)

    Shu, D G; Chen, L; Han, J; Zhang, X L

    2001-05-24

    Like the Burgess Shales of Canada, the Chengjiang Lagerstätte from the Lower Cambrian of China is renowned for the detailed preservation as fossils of delicate, soft-bodied creatures, providing an insight into the Cambrian explosion. The fossils of possible hemichordate chordates and vertebrates have attracted particular attention. Tunicates, or urochordates, comprise the most basal chordate clade, and details of their evolution could be important in understanding the sequence of character acquisition that led to the emergence of chordates and vertebrates. However, definitive fossils of tunicates from the Cambrian are scarce or debatable. Here we report a probable tunicate Cheungkongella ancestralis from the Chengjiang fauna. It resembles the extant ascidian tunicate genus Styela whose morphology could be useful in understanding the origin of the vertebrates. PMID:11373678

  17. Cambrian paleomagnetism of the Llano Uplift, Texas

    Science.gov (United States)

    Watts, Doyle R.; van der Voo, Rob; Reeve, Scott C.

    1980-10-01

    Late Cambrian sandstones and limestones sampled from various members of the Riley and Wilberns formations of the Llano uplift show a progression of paleomagnetic pole positions as a function of age. The members, ages, and poles are the following for the Riley Formation: the Hickory Sandstone, Lower Dresbachian, 34°N, 145°E; the Cap Mountain Limestone, Dresbachian, 33°N, 140°E; the Lion Mountain Limestone, Upper Dresbachian, 24°N, 146°E. For, the Wilberns Formation they are the following: the Welge Sandstone/Morgan Creek Limestone, Lower and Middle Franconian, 24°N, 151°E; and the Point peak Shale, Upper Franconian, 6°N, 159°E. These poles are based on thermal, chemical, and alternating field demagnetizations and on vector analysis. Most directions are interpreted to be of reversed polarity, but the Cap Mountain Limestone also yielded normal polarity directions; all directions resided in hematite with blocking temperatures up to 680°C. Almost all Cambrian poles appear to fall in a broad streak between the equator at about 155°E (e.g., the poles from the Tapeats Sandstone and from the Late Cambrian Point Peak Shale of the Wilberns Formation) and about 60°N, 90°E, where Cambrian poles have been obtained by Elston and Bressler (1977) and French et al. (1977). Although, at the present time, partial to complete remagnetization or nondipole behavior of the geomagnetic field are adequate ad hoc hypotheses to explain some of the data, it is suggested that the simplest and most unifying hypothesis to explain all data involves a Cambrian loop of apparent polar wander with respect to North America. This loop occurs before the middle Late Cambrian, with the poles from the Llano uplift falling on the return track.

  18. Collective behavior in an early Cambrian arthropod.

    Science.gov (United States)

    Hou, Xian-Guang; Siveter, Derek J; Aldridge, Richard J; Siveter, David J

    2008-10-10

    Examples that indicate collective behavior in the fossil record are rare. A group association of specimens that belong to a previously unknown arthropod from the Chengjiang Lagerstätte, China, provides evidence that such behavior was present in the early Cambrian (about 525 million years ago), coincident with the earliest extensive diversification of the Metazoa, the so-called Cambrian explosion event. The chainlike form of these specimens is unique for any arthropod, fossil or living, and most likely represents behavior associated with migration. PMID:18845748

  19. Cambrian Research in the New Century—the Fourth International Symposium on the Cambrian System

    Institute of Scientific and Technical Information of China (English)

    Peng Shanchi; Zhu Maoyan; L.E.Babcock

    2006-01-01

    @@ The International Subcomission on Cambrian Stratigraphy (ISCS) of the International Commission on Stratigraphy (ICS)held its Fourth International Symposium on the Cambrian System in Nanjing, the capital city of Jiangsu Province, China, on 18-24August, 2005. The Symposium was held in conjunction with the Tenth Field Conference of the Cambrian Stage Subdivision Working Group of the ISCS. Technical sessions were held in the conference center of Southeast University, and hosted by the Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, the Geological Institute, Chinese Academy of Geological Sciences, and Guizhou University. Associated with the meeting were five field excursions to classical Cambrian localities in North China and South China.

  20. Geochemistry of the Ediacaran-Early Cambrian transition in Central Iberia: Tectonic setting and isotopic sources

    Science.gov (United States)

    Fuenlabrada, José Manuel; Pieren, Agustín P.; Díez Fernández, Rubén; Sánchez Martínez, Sonia; Arenas, Ricardo

    2016-06-01

    A complete Ediacaran-Early Cambrian stratigraphic transition can be observed in the southern part of the Central Iberian Zone (Iberian Massif). Two different stratigraphic units, underlying Ordovician series, display geochemical and Sm-Nd isotopic features in agreement with an evolving geodynamic setting. Pusa Shales (Early Cambrian) rest unconformably on greywackes of the Lower Alcudian Formation (Late Ediacaran). Both sequences present minor compositional variations for major and trace element contents and similar REE patterns, close to those of PAAS (Post Archean Australian Shale). Trace element contents and element ratios suggest mixed sources, with intermediate to felsic igneous contributions for both units. Tectonic setting discrimination diagrams for the Ediacaran greywackes indicate that these turbiditic series were deposited in a sedimentary basin associated with a mature active margin (volcanic arc). However, the compositions of the Cambrian shales fit better with a more stable context, a back-arc or retro-arc setting. εNd(T) and TDM ages are compatible with dominance of a similar cratonic source for both sequences, probably the West Africa Craton. εNd565 values for the Ediacaran greywackes (- 3.0 to - 1.4) along with TDM ages (1256-1334 Ma) imply a significant contribution of juvenile material, probably derived from the erosion of the volcanic arc. However, εNd530 values in the Cambrian shales (- 5.2 to - 4.0) together with older TDM ages (1444-1657 Ma), suggest a higher contribution of cratonic isotopic sources, probably derived from erosion of the adjacent mainland. Coeval with the progressive cessation of arc volcanism along the peri-Gondwanan realm during the Cambrian, there was a period of more tectonic stability and increasing arrival of sediments from cratonic areas. The geochemistry of the Ediacaran-Cambrian transition in Central Iberia documents a tectonic switch in the periphery of Gondwana, from an active margin to a more stable context

  1. Timing and evolution of ocean anoxic event during Early Cambrian in south China

    Science.gov (United States)

    Yang, J.; Jiang, S.; Pi, D.; Ling, H.

    2008-12-01

    The Precambrian/Cambrian (PC-C) interval is one of the most interesting intervals in the evolution of life because of the sudden diversification of animals with mineralized skeletons, known as "Cambrian Explosion". The Yangtze Platform in south China is one of the best occurrences that can provide excellent insights into the palaeo-environmental and biological changes across the PC-C boundary. Our study show that the ocean anoxia were widespread during the Early Cambrian period, however, the start of this anoxic event was not from the PC-C boundary (i.e., 542 Ma), but some 7 Ma later (~535 Ma) when the Niutitang Formation black rock series (black phosphorite, chert, and black shale) deposited along a thousand kilometer long NEE zone in the transitional facies in the Yangtze Platform, while the major Cambrian radiation (Changjiang fauna) took place during 521-511 Ma. During the Niutitang period, the depositional environment of the Early Cambrian sedimentary sequence in south China have evolved from an initial oxic/dysoxic to a major anoxic/euxinic environment, and then back to dysoxic/oxic environment. A Ni-Mo sulfide layer occurred in the lower part of the Niutitang black shales which contains extremely enrichments of many metals, and can serve as a marker layer in south China when the depositional environment turned into euxinic condition. Re-Os isotope study of the sulfide ores and host black shales show an age of 535 Ma. Initial Os isotopic compositions, Mo isotopic compositions, and rare earth elements and Pt group element geochemistry suggest involvement of submarine hydrothermal fluids during the metal enrichments in black shale.

  2. The sudden appearance of diverse animal body plansduring the Cambrian explosion.

    Science.gov (United States)

    Chen, Jun-Yuan

    2009-01-01

    Beautifully preserved organisms from the Lower Cambrian Maotianshan Shale in central Yunnan, southern China, document the sudden appearance of diverse metazoan body plans at phylum or subphylum levels, which were either short-lived or have continued to the present day. These 530 million year old fossil representatives of living animal groups provide us with unique insight into the foundations of living animal groups at their evolutionary roots. Among these diverse animal groups, many are conservative, changing very little since the Early Cambrian. Others, especially Panarthropoda (superphylum), however, evolved rapidly, with origination of novel body plans representing different evolutionary stages one after another in a very short geological period of Early Cambrian time. These nested body plans portray a novel big picture of pararthropod evolution as a progression of step-wise changes both in the head and the appendages. The evolution of the pararthropods displays how the head/trunk boundary progressively shifted to the posterior, and how the simple annulated soft uniramous appendages progressively changed into stalked eyes in the first head appendages, into whip-like sensorial and grasping organs in the second appendage, and into jointed and biramous bipartite limbs in the post-antennal appendages. Haikouella is one of most remarkable fossils representing the origin body plan of Cristozoa, or crest animals (procraniates+craniates). The anatomy of Early Cambrian crest animals, including Haikouella and Yunnanozoon, contributes to novel understanding and discussion for the origins of the vertebrate brain, neural crest cells, branchial system and vertebrae. PMID:19557680

  3. A new formal perspective on 'Cambrian explosions'.

    Science.gov (United States)

    Wallace, Rodrick

    2014-01-01

    The 'Cambrian explosion' 500 Myr ago saw a relatively sudden proliferation of organism Bauplan and ecosystem niche structure that continues to haunt evolutionary biology. Here, adapting standard methods from information theory and statistical mechanics, we model the phenomenon as a noise-driven phase transition, in the context of deep-time relaxation of current path-dependent evolutionary constraints. The result is analogous to recent suggestions that multiple 'explosions' of increasing complexity in the genetic code were driven by rising intensities of available metabolic free energy. In the absence of severe path-dependent lock-in, 'Cambrian explosions' are standard features of blind evolutionary process, representing outliers in the ongoing routine of evolutionary punctuated equilibrium. PMID:24439546

  4. The notion of the Cambrian pananimalia genome.

    OpenAIRE

    Ohno, S.

    1996-01-01

    The toil by photosynthesizing cyanobacteria and blue-green algae of nearly three billion years appeared to have finally resulted in the sufficient accumulation of molecular oxygen. So, the stage was set for the emergence, at the ocean bottom, of diverse animals that were consumers of molecular oxygen. It now appears that this Cambrian explosion, during which nearly all the extant animal phyla have emerged, was of an astonishingly short duration, lasting only 6-10 million years. Inasmuch as on...

  5. Oxygen Requirements for the Cambrian Explosion

    Institute of Scientific and Technical Information of China (English)

    Xingliang Zhang; Linhao Cui

    2016-01-01

    Hypoxic tolerance experiments may be helpful to constrain the oxygen requirement for animal evolution. Based on literature review, available data demonstrate that fishes are more sensitive to hypoxia than crustaceans and echinoderms, which in turn are more sensitive than annelids, whilst mollusks are the least sensitive. Mortalities occur where O2 concentrations are below 2.0 mg/L, equivalent to saturation with oxygen content about 25% PAL (present atmospheric level). Therefore, the minimal oxygen requirement for maintaining animal diversity since Cambrian is determined as 25% PAL. The traditional view is that a rise in atmospheric oxygen concentrations led to the oxygenation of the ocean, thus triggering the evolution of animals. Geological and geochemical studies suggest a constant increase of the oxygen level and a contraction of anoxic oceans during Ediacaran–Cambrian transition when the world oceans experienced a rapid diversification of metazoan lineages. However, fossil first appearances of animal phyla are obviously asynchronous and episodic, showing a sequence as:basal metazoans>lophotrochozoans>ecdysozoans and deuterostomes. According to hitherto known data of fossil record and hypoxic sensitivity of animals, the appearance sequence of different animals is broadly consistent with their hypoxic sensitivity:animals like molluscs and annelids that are less sensitive to hypoxia appeared earlier, while animals like echinoderms and fishes that are more sensitive to hypoxia came later. Therefore, it is very likely that the appearance order of animals is corresponding to the increasing oxygen level and/or the contraction of anoxic oceans during Ediacaran–Cambrian transition.

  6. Yochelcionella (Mollusca, Helcionelloida) from the lower Cambrian of North America

    OpenAIRE

    Atkins C J; Peel J S

    2008-01-01

    Five named species of the helcionelloid mollusc genus Yochelcionella Runnegar & Pojeta, 1974 are recognized from the lower Cambrian (Cambrian Series 2) of North America: Yochelcionella erecta (Walcott, 1891), Y. americana Runnegar &Pojeta, 1980, Y. chinensis Pei, 1985, Y. greenlandica Atkins & Peel, 2004 and Y. gracilis Atkins & Peel, 2004, linking lower Cambrian o...

  7. Periodization

    OpenAIRE

    Lorenz, Daniel S.; Reiman, Michael P.; Walker, John C.

    2010-01-01

    Background: Clinicians are constantly faced with the challenge of designing training programs for injured and noninjured athletes that maximize healing and optimize performance. Periodization is a concept of systematic progression—that is, resistance training programs that follow predictable patterns of change in training variables. The strength training literature is abundant with studies comparing periodization schemes on uninjured, trained, and untrained athletes. The rehabilitation litera...

  8. Decoding the fossil record of early lophophorates : Systematics and phylogeny of problematic Cambrian Lophotrochozoa

    OpenAIRE

    Butler, Aodhán D.

    2015-01-01

    The evolutionary origins of animal phyla are intimately linked with the Cambrian explosion, a period of radical ecological and evolutionary innovation that begins approximately 540 Mya and continues for some 20 million years, during which most major animal groups appear. Lophotrochozoa, a major group of protostome animals that includes molluscs, annelids and brachiopods, represent a significant component of the oldest known fossil records of biomineralised animals, as disclosed by the enigmat...

  9. Adaptive walks in a gene network model of morphogenesis: insights into the Cambrian explosion

    OpenAIRE

    Sole, Ricard V.; Fernandez, Pau; Kauffman, Stuart A.

    2003-01-01

    The emergence of complex patterns of organization close to the Cambrian boundary is known to have happened over a (geologically) short period of time. It involved the rapid diversification of body plans and stands as one of the major transitions in evolution. How it took place is a controversial issue. Here we explore this problem by considering a simple model of pattern formation in multicellular organisms. By modeling gene network-based morphogenesis and its evolution through adaptive walks...

  10. High continental weathering rate during Early Cambrian: Evidence from Os isotopic composition of Early Cambrian Ocean

    Science.gov (United States)

    Jiang, S.-Y.; Yang, J.-H.; Ling, H.-F.; Feng, H.-Z.; Chen, Y.-Q.; Chen, J.-H.

    2003-04-01

    The paleo-ocean environmental change during the Precambrian-Cambrian transition is a key issue related to the causes for an explosive radiation of different metazoan phyla during Early Cambrian. The chemical and isotopic compositions of marine sediments and chemical precipitates such as carbonates, phosphorites, siliceous rocks, and black shales record the changing composition and physical conditions of the seawater in which these rocks accumulated. Organic carbon-rich black shales from marine environments are commonly enriched in a number of trace elements such as Ni, Mo, V, Co, Cr, Au, U, As, Pb, Zn, Cu, Re, and platinum-group-elements (PGE). Recent researches have demonstrated that Re-Os isotopes and PGE contents in black shales are useful proxies for seawater chemistry. It is believed that Re and Os in orgainc-carbon rich black shales are mostly hydrogeneous in origin which were largely sequestered from seawater at the time of deposition. In South China, the Lower Cambrian black shale sequence of the Niutitang Formation (and lateral equivalents) exists broadly several thousands kilometers. The lowermost sequence of this formation contain a thin sulfide ore horizon with an apparently unique and extreme case of metal enrichments such as Mo, Ni, Se, Re, Os, As, Hg, Sb, Ag, Au, Pt, and Pd. In this study, we conducted a preliminary investigation of Re-Os isotopes and Plantium Group Element (PGE) distribution patterns of the balck shales and intercalated Ni-Mo polymetallic sulfide bed from Guizhou and Hunan Provinces. The high rOs(t) values of the black shales indicate that the Early Cambrian ocean in Yangtze Platform had a highly radiogenic Os, possibly as a result of high continental weathering rate at that time. The Ni-Mo polymetallic sulfide ores within the black shales have lower rOs(t) values than the black shales, and they show similar REE and PGE patterns as the hydrothermal siliceous rocks within the Lower Cambrian strata, which suggest that the Ni

  11. Trilobite biogeography and Cambrian tectonic models

    Science.gov (United States)

    Burrett, Clive; Richardson, Robert

    1980-03-01

    The biogeographic patterns of Cambrian trilobites are almost impossible to explain on an Early Palaeozoic Pangaea but may be explained by relative movements of several continental blocks separated by wide ocean basins. Realms, regions, provinces and sub-provinces are recognised by progressively agglomerating 450 faunal lists, based on 1371 genera, distributed through eleven time segments. The agglomerating method proposed, is a simple one that may be performed by hand or computer and provides Medial Cambrian results similar to those of Jell (1974) who used principal components and cluster analyses. Simplified results are obtained when time segments are combined or when only the largest list from each 10° (lat. × long.) tessera is used or when only illustrated papers are used or when only lists containing ⩾ 3, 5 or 10 genera are used. The program was also run with all of the presumably planktonic miomerids removed resulting in smaller, better defined provinces with few links between tectonic blocks. The number of polymerid-only realms is three in Albertella Zone times, four in Glossopleura Zone and Eluinia/Conaspis Zone times and five in all other time segments. The average five realms recognised tend to be restricted to one (or rarely two) of the major tectonic blocks: America (excluding Boston and maritime Canada), Europe (including Morocco), Siberia, China and Australia. The existence of the American realm in Argentina through most of the Cambrian and the unlikelihood of a dismembered Gondwanaland necessitates using the latitudinal control hypothesis of Palmer (1972). This hypothesis places the European realm in high palaeolatitudes, the American realm in low palaeolatitudes in the Southern Hemisphere, the Australian realm in low palaeolatitudes in the Northern Hemisphere and the Siberian realm in temperate palaeolatitudes in the Southern Hemisphere. When the miomerids are included, connections between the peripheral ocean-facing regions of one tectonic block

  12. Cambrian Fossil Embryos from Western Hunan,South China

    Institute of Scientific and Technical Information of China (English)

    DONG Xiping

    2009-01-01

    The exquisitely preserved fossil embryos of Markuelia recovered from the limestones of the Middle Cambrian Haoqiao Formation and Upper Cambrian Bitiao formation in western Hunan,South China are described and illustrated in detail for the first time.A new species Markuelia elegans sp.nov.is established based mainly on embryos from the Upper Cambrian.A few of animal's resting eggs,which are comparable with those of the Ediacaran Doushantuo Formation,have been also found in the Upper Cambrian of western Hunan.The membrane of oue egg from the uppermost Cambrian has been replaced by pyrite and the overgrowth of the pyrite crystals exhibits a unique inorganic reducing conditions promoted the excellent preservation for the Markuelia specimens.The study of Markuelia provides not only constraint on the anatomy,affinity,embryonic development and phylogenetic significance of this wormlike animal and but also opens a new window onto the evolution and development of the earliest animals.

  13. Remagnetized cratonic Cambrian strata from southern Nevada

    Science.gov (United States)

    Gillett, Stephen L.

    1982-08-01

    Stratigraphic sections of Cambrian strata in southeastern Nevada are akin to thin, cratonic facies exposed in the Grand Canyon; their structural setting is much more complicated, however, from Mesozoic and Cenozoic tectonism. Paleomagnetic samples from two sections through these strata appear to have been completely remagnetized. Coarse-grained, lightcolored sandstone from the Tapeats Sandstone yields scattered magnetizations, residing in hematite, that appear to reflect protracted diagenetic acquisition of remanence. A hematitic sandstone in the Bright Angel Shale yields relatively consistent `Paleozoic' directions of magnetization, but petrographic study shows that the hematite results from diagenetic oxidation, and stratigraphic arguments suggest that the oxidation was not penecontemporaneous. Gray limestones of the Jangle and Muav Limestones yield a magnetization, residing in magnetite, that may reflect late Tertiary remagnetization, being imposed during uplift related to the onset of Basin and Range deformation. In any case, this magnetization differs greatly in direction from a hematite magnetization reported from slightly younger Muav Limestone in the Grand Canyon. Both the sampling sites also appear to have been tectonically rotated, but whether this rotation is true or a geometric artifact of the tilt correction cannot be determined from the present data. These results suggest that paleomagnetic data from rocks as old as Cambrian must be scrutinized very carefully before their magnetizations can be accepted as penecontemporaneous, and such scrutiny must include geologic data.

  14. Segmentation, metamerism and the Cambrian explosion.

    Science.gov (United States)

    Couso, Juan Pablo

    2009-01-01

    Data on the molecular and genetic basis of animal development, and on genome sequences, have been challenging our established assumptions about animal evolution for the last decade. Recent such data in animals of particular phylogenetic importance beg us to take another look at whether similarities in developmental and genetic mechanisms in current animals are the product of a common inheritance (homology) or convergent evolution (analogy). The evolution of segmentation, in particular whether segmentation and metameric bodies have arisen just once or several times in evolution, is a prime concern. Segmentation and metamerism are striking developmental and body organisations that exist, in varying degrees, in many complex animals, but the traditional view holds that this is the result of convergent evolution. Here, I review recent palenotological and developmental information and conclude that a metameric body plan is not only a likely ancestral character of bilaterian animals, but also a possible trigger for the Cambrian explosion in body morphology and complexity. This conclusion is supported by the phylogenetic distribution and prevalence of metameric phyla in the Cambrian, and the similarity of the genomes and segmentation mechanisms across current bilaterian phyla. PMID:19247939

  15. The significance of 24-norcholestanes, triaromatic steroids and dinosteroids in oils and Cambrian-Ordovician source rocks from the cratonic region of the Tarim Basin, NW China

    Science.gov (United States)

    Li, Meijun; Wang, T.-G.; Lillis, Paul G.; Wang, Chunjiang; Shi, Shengbao

    2012-01-01

    Two oil families in Ordovician reservoirs from the cratonic region of the Tarim Basin are distinguished by the distribution of regular steranes, triaromatic steroids, norcholestanes and dinosteroids. Oils with relatively lower contents of C28 regular steranes, C26 20S, C26 20R + C27 20S and C27 20R regular triaromatic steroids, dinosteranes, 24-norcholestanes and triaromatic dinosteroids originated from Middle–Upper Ordovician source rocks. In contrast, oils with abnormally high abundances of the above compounds are derived from Cambrian and Lower Ordovician source rocks. Only a few oils have previously been reported to be of Cambrian and Lower Ordovician origin, especially in the east region of the Tarim Basin. This study further reports the discovery of oil accumulations of Cambrian and Lower Ordovician origin in the Tabei and Tazhong Uplifts, which indicates a potential for further discoveries involving Cambrian and Lower Ordovician sourced oils in the Tarim Basin. Dinosteroids in petroleum and ancient sediments are generally thought to be biomarkers for dinoflagellates and 24-norcholestanes for dinoflagellates and diatoms. Therefore, the abnormally high abundance of these compounds in extracts from the organic-rich sediments in the Cambrian and Lower Ordovician and related oils in the cratonic region of the Tarim Basin suggests that phytoplankton algae related to dinoflagellates have appeared and might have flourished in the Tarim Basin during the Cambrian Period. Steroids with less common structural configurations are underutilized and can expand understanding of the early development history of organisms, as well as define petroleum systems.

  16. Formation of the 'Great Unconformity' as a trigger for the Cambrian explosion.

    Science.gov (United States)

    Peters, Shanan E; Gaines, Robert R

    2012-04-19

    The transition between the Proterozoic and Phanerozoic eons, beginning 542 million years (Myr) ago, is distinguished by the diversification of multicellular animals and by their acquisition of mineralized skeletons during the Cambrian period. Considerable progress has been made in documenting and more precisely correlating biotic patterns in the Neoproterozoic-Cambrian fossil record with geochemical and physical environmental perturbations, but the mechanisms responsible for those perturbations remain uncertain. Here we use new stratigraphic and geochemical data to show that early Palaeozoic marine sediments deposited approximately 540-480 Myr ago record both an expansion in the area of shallow epicontinental seas and anomalous patterns of chemical sedimentation that are indicative of increased oceanic alkalinity and enhanced chemical weathering of continental crust. These geochemical conditions were caused by a protracted period of widespread continental denudation during the Neoproterozoic followed by extensive physical reworking of soil, regolith and basement rock during the first continental-scale marine transgression of the Phanerozoic. The resultant globally occurring stratigraphic surface, which in most regions separates continental crystalline basement rock from much younger Cambrian shallow marine sedimentary deposits, is known as the Great Unconformity. Although Darwin and others have interpreted this widespread hiatus in sedimentation on the continents as a failure of the geologic record, this palaeogeomorphic surface represents a unique physical environmental boundary condition that affected seawater chemistry during a time of profound expansion of shallow marine habitats. Thus, the formation of the Great Unconformity may have been an environmental trigger for the evolution of biomineralization and the 'Cambrian explosion' of ecologic and taxonomic diversity following the Neoproterozoic emergence of animals. PMID:22517163

  17. A molecular and isotopic study of palaeoenvironmental conditions through the middle Cambrian in the Georgina Basin, central Australia

    Science.gov (United States)

    Pagès, Anais; Schmid, Susanne; Edwards, Dianne; Barnes, Stephen; He, Nannan; Grice, Kliti

    2016-08-01

    The Cambrian period marks an important point in Earth's history with profound changes in the ocean's biogeochemistry and the occurrence of the most significant evolutionary event in the history of life, the Cambrian explosion. The Cambrian explosion is described as a succession of complex cycles of extinctions and radiations. This study integrates biomarkers and their compound-specific stable carbon isotopes to investigate the palaeoenvironmental depositional conditions in middle Cambrian (Series 3) sedimentary rocks (Thorntonia Limestone, Inca Formation and Currant Bush Limestone) from two drillholes in the Undilla Sub-basin in the eastern Georgina Basin, central Australia. The occurrence of photic zone euxinia (PZE) was detected throughout these three formations by the identification of green sulfur bacteria Chlorobiaceae-derived biomarkers, including a series of 2,3,6-aryl isoprenoids and the intact biomarker isorenieratane. Pulses of enhanced PZE conditions were detected in two core intervals (90-110 mKB, Currant Bush Limestone and 170-200 mKB, Inca Formation) by an increase in the 2,3,6-aryl isoprenoids and C19 biphenyl concentrations. These enhanced PZE conditions were followed by blooms of phytoplankton, as demonstrated by the increase in algal-derived biomarker (i.e. pristane, phytane and the C19n-alkane) concentrations and compound-specific isotopes. These observations confirm that palaeoenvironmental conditions were similar to those reported for the Permian/Triassic and Triassic/Jurassic mass extinction events. The sterane distributions varied across the three formations reflecting possible changes in the phytoplanktonic communities through time. Although a rise in atmospheric oxygen during the Cambrian has been previously associated with the rapid evolution of metazoans, the ecological challenges related to widespread anoxia must have had a major influence on the evolution of life in Cambrian oceans.

  18. Upper Cambrian (Furongian) biostratigraphy in Scandinavia revisited

    DEFF Research Database (Denmark)

    Nielsen, Arne Thorshøj; Weidner, Thomas; Terfelt, Fredrik;

    2014-01-01

    , 73–87], are resurrected and elevated to superzonal rank. These superzones are usually readily recognized in the field, even by non-specialists, and for general correlation and mapping the more wide-ranging biozones are considerably more practicable than the very detailed zonation (formerly......, Parabolina and Leptoplastus (super)zones as well as the recently introduced Acerocarina Superzone ( = Acerocare Zone of older literature) are maintained and formalized. The Protopeltura praecursor, Peltura minor and Peltura scarabaeoides zones are abandoned and replaced by two new units, named the...... Protopeltura and Peltura superzones, respectively. Accordingly, all Furongian superzones have a uniform naming style referring to a characteristic genus. The six Furongian superzones currently comprise 27 trilobite zones. The Paradoxides forchhammeri Superzone (Cambrian Series 3) is extended upwards to the...

  19. An Early Cambrian problematic fossil: Vetustovermis and its possible affinities

    OpenAIRE

    Chen, Jun-Yuan; Huang, Di-Ying; Bottjer, David J.

    2005-01-01

    The Early Cambrian problematic fossil Vetustovermis (Glaessner 1979 Alcheringa 3, 21–31) was described as an annelid or arthropod. Anatomical analysis of 17 new specimens from the Lower Cambrian Maotianshan Shale at Anning, Kunming (South China) does not support its affinities with annelids or arthropods. Anatomical features instead resemble other animal groups including modern flatworms, nemertines and molluscs. The presence of a pelagic slug-like form and ventral foot, as well as a head wit...

  20. Global standard names for the Lowermost Cambrian Series and Stage

    Institute of Scientific and Technical Information of China (English)

    Ed Landing; Shanchi Peng; Loren E.Babcock; Gerd Geyer; Malgorzata Moczydlowska-Vidal

    2007-01-01

    @@ The GSSP marking the base of the Cambrian System was ratified by the IUGS in 1992.Ratification of the GSSP point at the base of the Trichophycus pedum Ichnozone in the Fortune Head section,eastern Newfoundland,Canada,automatically defined the conterminant base of the lowermost series and stage of the Cambrian although names for those subdivisions were not proposed at the time of the decision.

  1. Reconstruction of early Cambrian ocean chemistry from Mo isotopes

    Science.gov (United States)

    Wen, Hanjie; Fan, Haifeng; Zhang, Yuxu; Cloquet, Christophe; Carignan, Jean

    2015-09-01

    The Neoproterozoic-Cambrian transition was a key time interval in the history of the Earth, especially for variations in oceanic and atmospheric chemical composition. However, two conflicting views exist concerning the nature of ocean chemistry across the Precambrian-Cambrian boundary. Abundant geochemical evidence suggests that oceanic basins were fully oxygenated by the late Ediacaran, while other studies provide seemingly conflicting evidence for anoxic deep waters, with ferruginous conditions [Fe(II)-enriched] persisting into the Cambrian. Here, two early Cambrian sedimentary platform and shelf-slope sections in South China were investigated to trace early Cambrian ocean chemistry from Mo isotopes. The results reveal that early Cambrian sediments deposited under oxic to anoxic/euxinic conditions have δ98/95Mo values ranging from -0.28‰ to 2.29‰, which suggests that early Cambrian seawater may have had δ98/95Mo values of at least 2.29‰, similar to modern oceans. The heaviest and relatively homogeneous δ98/95Mo values were recorded in siltstone samples formed under completely oxic conditions, which is considered that Mn oxide-free shuttling was responsible for such heavy δ98/95Mo value. Further, combined with Fe species data and the accumulation extent of Mo and U, the variation of δ98/95Mo values in the two studied sections demonstrate a redox-stratified ocean with completely oxic shallow water and predominantly anoxic (even euxinic) deeper water having developed early on, which eventually became completely oxygenated. This suggests that oceanic circulation at the time became reorganized, and such changes in oceanic chemistry may have been responsible for triggering the "Cambrian Explosion" of biological diversity.

  2. Pulse of atmospheric oxygen during the late Cambrian

    OpenAIRE

    Saltzman, Matthew R.; Young, Seth A.; Kump, Lee R.; Gill, Benjamin C.; Lyons, Timothy W.; Runnegar, Bruce

    2011-01-01

    A rise in atmospheric O2 has been linked to the Cambrian explosion of life. For the plankton and animal radiation that began some 40 million yr later and continued through much of the Ordovician (Great Ordovician Biodiversification Event), the search for an environmental trigger(s) has remained elusive. Here we present a carbon and sulfur isotope mass balance model for the latest Cambrian time interval spanning the globally recognized Steptoean Positive Carbon Isotope Excursion (SPICE) that i...

  3. Primitive soft-bodied cephalopods from the Cambrian.

    Science.gov (United States)

    Smith, Martin R; Caron, Jean-Bernard

    2010-05-27

    The exquisite preservation of soft-bodied animals in Burgess Shale-type deposits provides important clues into the early evolution of body plans that emerged during the Cambrian explosion. Until now, such deposits have remained silent regarding the early evolution of extant molluscan lineages-in particular the cephalopods. Nautiloids, traditionally considered basal within the cephalopods, are generally depicted as evolving from a creeping Cambrian ancestor whose dorsal shell afforded protection and buoyancy. Although nautiloid-like shells occur from the Late Cambrian onwards, the fossil record provides little constraint on this model, or indeed on the early evolution of cephalopods. Here, we reinterpret the problematic Middle Cambrian animal Nectocaris pteryx as a primitive (that is, stem-group), non-mineralized cephalopod, based on new material from the Burgess Shale. Together with Nectocaris, the problematic Lower Cambrian taxa Petalilium and (probably) Vetustovermis form a distinctive clade, Nectocarididae, characterized by an open axial cavity with paired gills, wide lateral fins, a single pair of long, prehensile tentacles, a pair of non-faceted eyes on short stalks, and a large, flexible anterior funnel. This clade extends the cephalopods' fossil record by over 30 million years, and indicates that primitive cephalopods lacked a mineralized shell, were hyperbenthic, and were presumably carnivorous. The presence of a funnel suggests that jet propulsion evolved in cephalopods before the acquisition of a shell. The explosive diversification of mineralized cephalopods in the Ordovician may have an understated Cambrian 'fuse'. PMID:20505727

  4. Diversification of skeletal microstructures of organisms through the interval from the latest Precambrian to the Early Cambrian

    Institute of Scientific and Technical Information of China (English)

    FENG; Weimin(冯伟民); CHEN; Zhe(陈哲); SUN; Weiguo(孙卫国)

    2003-01-01

    Research on early skeletal fossils of southern Shaanxi Province and eastern Yunnan Province shows that the interval from the latest Precambrian to the Early Cambrian was an important period of significant changes in biomineralization of various organisms. The latest Precambrian skeletal fossils, represented by Cloudina, are characterized by cone-in-cone structures, relatively high content of organic matter, and various shell shapes with obvious plastic deformation. In the Early Cambrian the skeletal organisms, consisting of phosphatic, carbonate and siliceous minerals, began to appear in great abundance and to distribute widely. Moreover, microstructure of these skeletal organisms appeared to have been diversified, for example, main types of recent molluscan microstructures, such as prismatic, lamello-fibrillar and regular foliated structures, were already developed at that time; fibrous structures further diversified, such as lamello-fibrillas indicative of mollusks and fibre bundles of hyoliths; there were at least two layers in a shell. This shows that biologically controlled biomineralization had played the main role in the Early Cambrian and it laid a key foundation for formation of the true shell of diverse invertebrates, giving rise to the "Cambrian explosion".

  5. Marine Redox Conditions in the Early Cambrian Ocean:Insights from the Lower Cambrian Phosphorite Deposits, South China

    Institute of Scientific and Technical Information of China (English)

    Haifeng Fan; Hanjie Wen; Xiangkun Zhu

    2016-01-01

    ABSTRACT:It is generally considered that a significant change in oceanic redox conditions occurred during the Ediacaran–Cambrian transition. However, there are currently two major conflicting views on the degree of oxygenation of deep water (oxic vs. ferruginous) during this interval. To date, the oxygenation conditions of the Early Cambrian ocean have not been well constrained. The oxygenation magnitude and mechanism of the Early Cambrian ocean could be critical to the significant biological evolution of the“Cambrian Explosion”. To constrain the Early Cambrian oceanic redox environment, we conducted an integrated study on iron and sulfur isotopes and redox-sensitive elements (Mo, U, and V) of Lower Cambrian phosphorite deposits from two shallow sections (Meishucun and Gezhongwu) and a deeper water section (Zunyi) from the Yangtze Platform, South China. The near zero δ56Fe values from the two shallow sections studied here reflect oxic conditions in the lower phosphorite deposition. An obvious positive shift inδ56Fe and redox-sensitive element content was observed in the middle parts of the two shallow water sections, which might reflect loss of light iron by dissimilatory iron reduction during early diagenesis under suboxic shallow water in the platform. However, the highly positiveδ56Fe values in the deep section could reflect a lower oxidation degree of dissolved Fe(II) under anoxic deep water. The data suggest redox-stratified oceanic conditions during the Early Cambrian, in which completely oxygenated shallow water (platform) coexisted with anoxic deep water (slope). We propose that prolonged upwelling of dissolved organic carbon (DOC)-, Fe(II)- and phosphorus-rich anoxic deep water in a redox-stratified ocean could have increased exchange with the open ocean, resulting in major phosphorite deposition in oxic-suboxic conditions. The progressive oxygenation of the ocean may have facilitated the Early Cambrian biotic diversification.

  6. Why is construction of an Ediacaran-Cambrian global paleogeography so difficult?

    Science.gov (United States)

    Evans, D. A.; Raub, T. D.; Maloof, A. C.

    2004-05-01

    The foremost answer to the above question is that high-precision global biostratigraphy extends only to Early Cambrian time. A more challenging issue concerns paleomagnetic data from the Ediacaran (proposed terminal Proterozoic period) and earliest Cambrian intervals: each paleocontinent's apparent polar wander path shows a large spread in poles, intriguingly distributed around a great circle (excepting Baltica, see below), and implying rates of paleolatitude translation or rotation far exceeding typical values from the Cenozoic. Oscillatory true polar wander (TPW) has been proposed to explain these great-circle distributions, but as more data are gathered the number of required episodes increases steadily. Another mechanism to explain such jumpy datasets is a persistently nonaxial geomagnetic field, with greatly amplified paleosecular variation; however, such an interpretation is unattractive to many geodynamicists. A third approach to explain the anomalous data is to discount them altogether; this is unsatisfactory because some of the most reliable results (e.g., Grenville dykes B component and Bunyeroo Formation pole) must be discarded in order to simplify the paths. We propose a new technique for Ediacaran-Cambrian continental reconstruction, which uses these intriguing great circle pole paths regardless of how they were generated. Each paleocontinent's path yields a best-fit great circle and corresponding pole. In the TPW interpretation, that pole describes the long-lived, common (equatorial) axis to the oscillatory rotations. Given that age constraints on the poles is generally so poor, we may reconstruct each continent to this axis with azimuthal freedom. This is topologically identical to the longitudinal degeneracy of standard paleomagnetic reconstructions, but in the case of TPW uncertainties in positions along the great circle represent uncertainties in paleolatitudes, hence climate zones. Analysis of the global Ediacaran-Cambrian paleomagnetic database

  7. Lithofacies palaeogeography of the Cambrian in Tarim area

    Institute of Scientific and Technical Information of China (English)

    FENG Zengzhao; BAO Zhidong; WU Maobing; JIN Zhenkui; SHI Xiaozhang

    2007-01-01

    The Tarim area in this paper refers to the present Tarim Basin and its surrounding mountains, bounded on the north by the Middle Tianshan Mountains, on the south by the Kunlun Mountains, on the west by the national boundary of China, and on the east by the Altun Mountains. It covers an area of more than 1 000 000 km2. Based on the quantitative and qualitative data from the stratigraphy and petrology of outcrop and well sections, and according to the single factor analysis and multifactor comprehensive mapping method,the single factor maps of the Lower, Middle and Upper Cambrian, and the lithofacies palaeogeography maps of the Early, Middle and Late Cambrian of the Tarim area, are com-piled. The most important character of these lithofacies pal-aeogeography maps is quantification, I.e. The determination of each palaeogeography unit based on accurate quantitativedata and the quantitative single factor maps. The study marksthe first time that this quantitative method has been appliedin the Tarim area. In the Early Cambrian, the present TarimBasin was mainly a carbonate platform, in which there werepenebank, dolostone flats, and gypsum-halite lake. The South Tianshan Basin and East Tarim Basin were siliceous rock-limestone-mudstone basins. The Kunlun Platform was situ-ated in the south, where there were two small lands. The AltunLand was situated in the southeast. In the Middle and Late Cambrian, the framework of lithofacies palaeogeography was similar to that in the Early Cambrian; the land area was gradually reduced; the basin area was enlarged; while the Tarim Carbonate Platform changed little. The transgression continued from the Early Cambrian to the Late Cambrian.

  8. Explaining the Cambrian "Explosion" of Animals

    Science.gov (United States)

    Marshall, Charles R.

    2006-05-01

    The Cambrian "explosion" is a unique episode in Earth history, when essentially all the animal phyla first appear in the fossil record. A variety of environmental, developmental (genetic), and ecological explanations for this complex and somewhat protracted event are reviewed, with a focus on how well each explains the observed increases in disparity and diversity, the time of onset of the radiation, its duration, and its uniqueness. The increase in disparity (the origin of the phyla) and diversity are best understood as being the result of the interplay of the combinatorial bilaterian developmental system and the increase in the number of needs the first bilaterians had to meet as complex ecological interactions developed. The time of onset is constrained by the evolution of the environment, whereas its duration appears to be controlled primarily by rates of developmental innovation. The uniqueness of the event is either due to ensuing developmental limitation, to ecological saturation, or simply to the exhaustion of ecologically viable morphologies that could be produced by the nascent bilaterian developmental system.

  9. Neo proterozoic Cambrian in Uruguay: Dom Feliciano belt against Cuchilla Dionisio Terrane

    International Nuclear Information System (INIS)

    After an exhaustive bibliographic research about the subject, its was detected two opinion groups that beleive different evolution processes during Neoproterozoic Cambrian period in Uruguay.One of them beleives that litho-structural arrangement was generated in an unique Wilson cycle, the other one, supports that the genesis of nowaday structure is a tangential collision near 520 Ma; this point of view coincides with authors thinking 62 papers were consulted and many geological as some geochronological unpublished data were used in the analysis of the subject

  10. Rates of phenotypic and genomic evolution during the Cambrian explosion.

    Science.gov (United States)

    Lee, Michael S Y; Soubrier, Julien; Edgecombe, Gregory D

    2013-10-01

    The near-simultaneous appearance of most modern animal body plans (phyla) ~530 million years ago during the Cambrian explosion is strong evidence for a brief interval of rapid phenotypic and genetic innovation, yet the exact speed and nature of this grand adaptive radiation remain debated. Crucially, rates of morphological evolution in the past (i.e., in ancestral lineages) can be inferred from phenotypic differences among living organisms-just as molecular evolutionary rates in ancestral lineages can be inferred from genetic divergences. We here employed Bayesian and maximum likelihood phylogenetic clock methods on an extensive anatomical and genomic data set for arthropods, the most diverse phylum in the Cambrian and today. Assuming an Ediacaran origin for arthropods, phenotypic evolution was ~4 times faster, and molecular evolution ~5.5 times faster, during the Cambrian explosion compared to all subsequent parts of the Phanerozoic. These rapid evolutionary rates are robust to assumptions about the precise age of arthropods. Surprisingly, these fast early rates do not change substantially even if the radiation of arthropods is compressed entirely into the Cambrian (~542 mega-annum [Ma]) or telescoped into the Cryogenian (~650 Ma). The fastest inferred rates are still consistent with evolution by natural selection and with data from living organisms, potentially resolving "Darwin's dilemma." However, evolution during the Cambrian explosion was unusual (compared to the subsequent Phanerozoic) in that fast rates were present across many lineages. PMID:24035543

  11. Magmatism in a Cambrian Laurentian Plate Rift

    Science.gov (United States)

    Gilbert, M. C.

    2008-12-01

    Evidences of the Cambrian Southern Oklahoma Aulacogen extend over 1000km from about Dallas out to the Uncompahgre Plateau in SW Colorado. The signature of this originally extensional feature can be traced geophysically, and in some places at the present surface, petrologically and temporally, by the presence of mafic rock. It appears to have been the intracontinental third arm of a plume-generated? triple junction which helped to dismember the southern part of Laurentia on the final break-up of a Neoproterozoic supercontinent. Other parts of Laurentia rifted away and are now found in the Precordillera of Argentina. Rift magmatism appears to have been concentrated nearer the plate edge during the breakup. Perhaps as much as 40,000 km3 of mostly subaerial silicic volcanics and shallow-seated granites overlay and filled the top of the rift in the area of SW Oklahoma. The rift fill below the silicic rocks is large, layered mafic complexes and smaller, layered, hydrous gabbros, the whole set appearing as a shallow AMCG complex. Unusually, direct rift sediments are not obvious. Furthermore, silicic and mafic rocks have identical Nd signatures. Finally, about 20 Ma after rifting ceased and later into the Paleozoic during sea incursion, overlying sediments are thickened 4X compared to equivalent units 100's of kms to the rift sides. This rift appears distinct from most modern rifts. Conclusions are 1) This was a hot, narrow rift; 2) Basaltic magmatism , not sedimentation, filled the rift; 3) Magmatic intensity varied along the rift strike; 4) Silicic rocks were generated mostly directly from new mantle-derived basalt liquids through fractionation, not melting of older crustal rocks; 5) Laurentian lithosphere was weak allowing centering of the Early/Middle Paleozoic large "Oklahoma" basin (pre-Anadarko) over the rift.

  12. Proterozoic and early Cambrian protists: evidence for accelerating evolutionary tempo

    Science.gov (United States)

    Knoll, A. H.

    1994-01-01

    In rocks of late Paleoproterozoic and Mesoproterozoic age (ca. 1700-1000 million years ago), probable eukaryotic microfossils are widespread and well preserved, but assemblage and global diversities are low and turnover is slow. Near the Mesoproterozoic-Neoproterozoic boundary (1000 million years ago), red, green, and chromophytic algae diversified; molecular phylogenies suggest that this was part of a broader radiation of "higher" eukaryotic phyla. Observed diversity levels for protistan microfossils increased significantly at this time, as did turnover rates. Coincident with the Cambrian radiation of marine invertebrates, protistan microfossils again doubled in diversity and rates of turnover increased by an order of magnitude. Evidently, the Cambrian diversification of animals strongly influenced evolutionary rates, within clades already present in marine communities, implying an important role for ecology in fueling a Cambrian explosion that extends across kingdoms.

  13. Cambrian explosion triggered by geosphere-biosphere feedbacks

    Science.gov (United States)

    von Bloh, Werner; Bounama, Christine; Franck, Siegfried

    2003-09-01

    A new hypothesis for the cause of the Cambrian explosion is presented. For that the evolution of the planet Earth is described by the co-evolution of the geosphere-biosphere system. Here we specify our previously published Earth system model for the long-term carbon cycle by introducing three different types of biosphere: procaryotes, eucaryotes, and complex multicellular life. They are characterized by different global temperature tolerance windows. The biotic enhancement of silicate weathering by complex multicellular life adds an additional feedback to the system and triggers the Cambrian explosion. The Cambrian explosion is characterized by a sudden increase of biomass and a rapid cooling, which amplified the spread of complex multicellular life. Cooling events in the Neoproterozoic, however, could force a premature appearance of complex multicellular life.

  14. Discussions on the sedimentary-tectonic event and tectonic setting of the North Tarim Basin in Cryogenian-Cambrian

    Science.gov (United States)

    Zhou, X. B.; Li, J. H.; Li, W. S.

    2012-04-01

    Across the Tarim Basin, limited surface outcrops of Cryogenian to Cambrian sedimentary succession are completely exposed in the vicinity of Aksu area(Northwest Tarim), Kuruktag(Northeast Tarim)and Southwest Tarim, thus provides a unique, well preserved and accessible means by which to study the early development of the north Tarim Basin. Based on the field geological investigation in the northwestern and northeastern of Tarim Basin, with the referencing of paleomagnetism mapping and previous research, basin evolution process in Cryogenian-Cambrian is discussed according to sedimentary-tectonic event and other evidences. The major lithological types of Cryogenian-Cambrian system in Northeast Tarim are: tillite, clastic rocks(rich in organic matter) and carbonate ,with interbeds of volcanic rocks while in Northwest Tarim, the calstic rocks and carbonate are the common rock type, with tillite and volcanic interbeds in a small amount. The north margin of Tarim Block, which was a part of Rodinia supercontinent, neighboring the northwestern margin of Australia, was deeply rifted in Cryogenian-Ediacaran and developed into two rifts in the northwestern and northeastern margin, while formed a thick layer of the rift-passive margin deposits and the layer in the northwestern rift was not completely developed as the northeastern. The deepest rift-passive magin sediment which can be observed is Cryogenian-Middle Ordovician strata, and the period can be divided into Cryogenian faulted period (supercontinent rifting stage) and Ediacaran-Middle Ordovician subsidence period (plate drifting stage).

  15. Darwin's dilemma: the realities of the Cambrian ‘explosion’

    OpenAIRE

    Conway Morris, Simon

    2006-01-01

    The Cambrian ‘explosion’ is widely regarded as one of the fulcrum points in the history of life, yet its origins and causes remain deeply controversial. New data from the fossil record, especially of Burgess Shale-type Lagerstätten, indicate, however, that the assembly of bodyplans is not only largely a Cambrian phenomenon, but can already be documented in fair detail. This speaks against a much more ancient origin of the metazoans, and current work is doing much to reconcile the apparent dis...

  16. Provenance shift in Cambrian mid-Baltica: detrital zircon chronology of Ediacaran–Cambrian sandstones in Estonia

    Directory of Open Access Journals (Sweden)

    Yukio Isozaki

    2014-12-01

    Full Text Available In order to clarify the tectono-sedimentary history of Paleozoic Baltica, age spectra of detrital zircon grains from the Ediacaran (Kotlin Regional Stage and Lower Cambrian sandstones (lowermost Lontova and Lükati formations in western Estonia in central Baltica were analyzed by LA-ICPMS. The abundant occurrence of Archean to Mesoproterozoic (2800–1000 Ma zircon grains was confirmed in all samples. The new data provided the following information on the provenance of siliciclastic material as well as a major change in the sedimentary regime of the Paleo-Baltic basin during the Early Cambrian: (1 the Ediacaran–Lower Cambrian Paleo-Baltic basin received abundant terrigenous clastics from the core of Baltica underlain by the Archean–Mesoproterozoic crystalline crust, (2 the exposed surface area of the 1600 Ma Rapakivi granites apparently was more extensive during the Ediacaran–Early Cambrian than at present, (3 a major re-organization of the basin geometry occurred in the middle Early Cambrian (ca 530–515 Ma in central Baltica, inducing a change in the sediment supply system, (4 in contrast to the total absence of Neoproterozoic detrital zircon grains before the middle Early Cambrian, their sudden appearance at this time, together with consistent occurrence at least until the mid-Devonian, suggests a significant uplift event located in southeast Baltica and/or in a more easterly land domain (e.g., in Sarmatia, (5 possible sources for the Neoproterozoic zircon grains include the peripheral mobile belts with pan-African signatures around Baltica, e.g., the so-called Gondwanan fragments along the Tornquist margin to the southwest and the Timanian belt along the northeastern margin.

  17. Dr. Cooper Curtice - Unknown worker in interpreting the Cambrian of Alabama

    Science.gov (United States)

    Yochelson, E.L.; Osborne, W.E.

    1999-01-01

    Cooper Curtice was an assistant to C. D. Walcott from 1883-1886. In 1885, he spent four months, mostly in Alabama, measuring sections of Paleozoic rocks and searching for fossils, mainly in the Cambrian. In 1888, Walcott concurred with foreign authorities that the rocks called Middle Cambrian in North America were Early Cambrian in age and vice versa, requiring a new interpretation of Cambrian strata. Curtice returned to Alabama for geologic investigations in 1892, and again briefly with Walcott in 1895. Since that time Cambrian stratigraphy in the southeastern United States has remained virtually unchanged.

  18. Cambrian trace fossil Zoophycos from the Czech Republic

    Czech Academy of Sciences Publication Activity Database

    Doucek, J.; Mikuláš, Radek

    2014-01-01

    Roč. 65, č. 6 (2014), s. 403-409. ISSN 1335-0552 R&D Projects: GA ČR GA205/09/1521 Institutional support: RVO:67985831 Keywords : Cambrian * Bohemian Massif * Železné hory * ichnology * marine settings * Zoophycos Subject RIV: DB - Geology ; Mineralogy Impact factor: 0.761, year: 2014

  19. New Evidence for Pre-Cambrian Development of Complex Animals

    Institute of Scientific and Technical Information of China (English)

    2006-01-01

    @@ Much mystery surrounds the sudden and mushrooming appearance of animals in the fossil record about 540 million years ago. Known as the "Cambrian explosion," the change took place from zero evidence to all the basic types of creatures in today's world within a few million years.

  20. Nonbiomineralized carapaces in Cambrian seafloor landscapes (Sirius Passet, Greenland)

    DEFF Research Database (Denmark)

    Mangano, M. Gabriela; Bromley, Richard Granville; Harper, David A.T.; Nielsen, Arne Thorshøj; Smith, M. Paul; Vinther, Jakob

    2012-01-01

    Widespread microbial mats and the absence of significant bioturbation resulted in a poorly developed mixed layer and extensive cohesive substrates during the Ediacaran-Cambrian transition. Large nonbiomineralized arthropod carapaces overprinted with trails, interconnected burrow systems, narrow-c...... served as the primary food for small macrofauna and meiofauna....

  1. A New Landmark for Exploration of the Cambrian Explosion

    Institute of Scientific and Technical Information of China (English)

    Zhao Baohua; Song Jianlan; Guo Haiyan

    2002-01-01

    @@ New studies suggest that the Kaili Biota in southwestern China's Guizhou Province could be the third revealing evidence for the spectacular Cambrian Explosion of life about 530 million years ago, next to the Burgess Shale Biota of Canada, found in 1909,and the Chengjiang Biota, brought to light in Yunnan Province, China,in 1984.

  2. Oceanic anoxia at the Precambrian-Cambrian boundary

    Science.gov (United States)

    Kimura, Hiroto; Watanabe, Yoshio

    2001-11-01

    The Precambrian-Cambrian (PC-C) boundary separates fossils representing two discrete evolutionary phases: the Neoproterozoic soft-bodied Ediacarian biotas and Cambrian small shelly faunas. The biological discontinuity is suspected to have been a result of mass extinction; however, recent discoveries of the Ediacarian biotas in Cambrian sediments have led to an understanding that the faunal change was gradual through the PC-C transition. Th/U ratios, which are high in oxidizing conditions and low in reducing conditions, show a considerable positive correlation with δ13C values at all studied sites of the PC-C boundary. This correlation indicates that reported δ13C variation across the PC-C boundary from numerous localities corresponds to redox variation in the depositional environment. The negative δ13C anomaly that occurs worldwide at the PC-C boundary, therefore, corresponds to the widespread development of an oxygen-deficient shallow marine environment. This finding suggests that widespread oceanic oxygen deficiency, which has been interpreted to reflect Phanerozoic mass extinction events, also occurred immediately before the Cambrian explosion.

  3. New Vetulicoliids from the Lower Cambrian Guanshan Fauna,Kunming

    Institute of Scientific and Technical Information of China (English)

    LUO Huilin; FU Xiaoping; HU Shixue; LI Yong; CHEN Liangzhong; YOU Ting; LIU Qi

    2005-01-01

    The Guanshan Fauna, a soft-bodied fauna intermediate between the Chengjiang Fauna and the Kaili Fauna and also the Burgess Shale Fauna stratigraphically, consists of trilobites, trilobitoides, Tuzoia, Vetulicola, Paleoscolex, brachiopods and sponges. The discovery and research of this fauna is of great significance in understading the "Cambrian Explosion" and the evolution of early life. The occurrence of vetulicoliids from the Guanshan Fauna not only adds new members to the taxonomic list, but also provides new information to the evolution of this animal group. This paper describe Vetulicola gantoucunensis Luo, Fu et Hu sp. nov. from the Lower Cambrian Wulongqing Formation in the Kunming area. Also presented are the amended description of Vetulicola and the comparisons with related genera within Vetulicoliids. The affinity, distribution, as well as evolution of vetulicoliids are discussed.

  4. New and extraordinary Early Cambrian sponge spicule assemblage from China

    Science.gov (United States)

    Zhang, Xi-Guang; Pratt, Brian R.

    1994-01-01

    The fossil record of siliceous sponges, compared with that of other skeleton- secreting Metazoa, is poorly known, based as it is on disarticulated spicules and sporadically preserved body fossils. Abundant spicules recovered from Lower Cambrian strata in Shaanxi, China, essentially double the known morphological diversity of siliceous sponges for that interval of geologic time. These fossils, along with a comparable coeval fauna from South Australia, have a remarkably modern aspect, thereby demonstrating that the principal siliceous sponge groups and styles of body architecture were established quickly in the earliest Phanerozoic as part of the Cambrian "explosion" and that they inhabited a variety of low-energy, relatively deep water settings. The similarity of spicule shape and variation to that of younger assemblages reflects a conservative architecture for the siliceous sponges.

  5. Cambrian extensional tectonics and magmatism within the Southern Oklahoma aulacogen

    Science.gov (United States)

    McConnell, D. A.; Gilbert, M. C.

    1990-03-01

    The tectonic evolution of the Southern Oklahoma aulacogen is partially constrained by the petrological consequences of a bimodal igneous suite associated with the Cambrian rift. Mineralogical and compositional layers and erosional surfaces are recognized as initially subplanar, subhorizontal markers. The progressive rotation of these horizons can be explained by uniform-sense normal faulting. Magmatism, confined to the aulacogen trend, elevated the thermal gradient producing a crustal strength-anisotropy. This ensured the localization of the extension throughout the Cambrian rifting event. The presence of a substantial volume of mafic igneous rocks within the crust along the aulacogen's trend suggests that crustal attenuation was compensated for by the addition of mantle derived material during extension.

  6. Upper Cambrian chitons (Mollusca, polyplacophora) from Missouri, USA

    Science.gov (United States)

    Pojeta, J., Jr.; Vendrasco, M.J.; Darrough, G.

    2010-01-01

    Numerous new specimens reveal a greater presence of chitons in Upper Cambrian rocks than previously suspected. Evidence is presented showing that the chiton esthete sensory system is present in all chiton species in this study at the very beginning of the known polyplacophoran fossil record. The stratigraphic occurrences and paleobiogeography of Late Cambrian chitons are documented. The 14 previously-named families of Cambrian and Ordovician chitons are reviewed and analyzed. Aulochitonidae n. fam. is defined, based on Aulochiton n. gen.; A. sannerae n. sp. is also defined. The long misunderstood family Preacanthochitonidae and its type genus Preacanthochiton Bergenhayn, 1960, are placed in synonymy with Mattheviidae and Chelodes Davidson & King, 1874, respectively; Eochelodes Marek, 1962, also is placed in synonymy with Chelodes, and Elongata Stinchcomb & Darrough, 1995, is placed in synonymy with Hemithecella Ulrich & Bridge, 1941. At the species level, H. elongata Stinchcomb & Darrough, 1995, and Elongata perplexa Stinchcomb & Darrough, 1995, are placed in synonymy with H. eminensis Stinchcomb & Darrough, 1995. The Ordovician species H. abrupta Stinchcomb & Darrough, 1995, is transferred to the genus Chelodes as C. abrupta (Stinchcomb & Darrough, 1995). The Ordovician species Preacanthochiton baueri Hoare & Pojeta, 2006, is transferred to the genus Helminthochiton as H. ? baueri (Hoare & Pojeta, 2006). The Ordovician species H. marginatus Hoare & Pojeta, 2006, is transferred to the genus Litochiton as L. marginatus (Hoare & Pojeta, 2006). Matthevia walcotti Runnegar, Pojeta, Taylor, & Collins, 1979, is treated as a synonym of Hemithecella expansa Ulrich & Bridge, 1941. In addition, other multivalved Cambrian mollusks are discussed; within this group, Dycheiidae n. fam. is defined, as well as Paradycheia dorisae n. gen. and n. sp. Cladistic analysis indicates a close relationship among the genera here assigned to the Mattheviidae, and between Echinochiton Pojeta

  7. Contributions to the Proterozoic and Cambrian Evolution of Eukaryotes

    OpenAIRE

    Dong, Lin

    2007-01-01

    This thesis makes several contributions to improve our understanding of Proterozoic-Cambrian evolution of eukaryote life. Chapter 1 provides, for the first time, a quantitative characterization of the evolutionary trends of Proterozoic macroalgae. The analysis reveals that morphological disparity of Paleoproterozoic macroalgae was low but increased in the Mesoproterozoic and Ediacaran, with a plateau in between. There was also a significant increase in thallus surface/volume ratio and maxi...

  8. Did homeobox gene duplications contribute to the Cambrian explosion?

    OpenAIRE

    Holland, Peter W. H.

    2015-01-01

    The Cambrian explosion describes an apparently rapid increase in the diversity of bilaterian animals around 540–515 million years ago. Bilaterian animals explore the world in three-dimensions deploying forward-facing sense organs, a brain, and an anterior mouth; they possess muscle blocks enabling efficient crawling and burrowing in sediments, and they typically have an efficient ‘through-gut’ with separate mouth and anus to process bulk food and eject waste, even when burrowing in sediment. ...

  9. Early-Middle Cambrian Palynomorph Microfossils and Related Geochemical Events in South China

    Institute of Scientific and Technical Information of China (English)

    Leiming Yin; Chunjiang Wang; Yuanlong Zhao; Zhiji Ou

    2016-01-01

    The earliest Cambrian acritarch ‘Asteridium–Heliosphaeridium–Comasphaeridium’ assemblage and the Early Cambrian (‘Chiungchussuan’) acritarch ‘Skiagia ornate–Fimbriaglomerella membranacea’ assemblage in South China can be correlated with assemblages from synchronous strata elsewhere in the world. Recent geochemical study and biomarker evidence further confirm a biostratigraphic change between Cambrian Series 2 and Series 3 and support the recognition of a major geological and biotic event during this time interval.

  10. First report of Crumillospongia (Demospongea) from the Cambrian of Europe (Murero biota, Spain)

    OpenAIRE

    García-Bellido D C; Dies Álvarez M E; Gámez Vintaned J A Liñán E; Gozalo R

    2011-01-01

    The demosponge genus Crumillospongia, originally described from the Burgess Shale (middle Cambrian of Canada), has only been cited from lower and middle Cambrian localities of North America and China. The taxon is now also described from uppermost lower Cambrian rocks of the Murero Lagerstätte (Zaragoza Province, NE Spain). Crumillospongia mureroensis sp. nov. is a small to medium sized sack-shaped to elongate demosponge characterized by the presence of d...

  11. Lower Cambrian polychaete from China sheds light on early annelid evolution

    OpenAIRE

    Liu, Jianni; Ou, Qiang; Han, Jian; Li, Jinshu; Wu, Yichen; Jiao, Guoxiang; He, Tongjiang

    2015-01-01

    We herein report a fossilized polychaete annelid, Guanshanchaeta felicia gen. et sp. nov., from the Lower Cambrian Guanshan Biota (Cambrian Series 2, stage 4). The new taxon has a generalized polychaete morphology, with biramous parapodia (most of which preserve the evidence of chaetae), an inferred prostomium bearing a pair of appendages, and a bifid pygidium. G. felicia is the first unequivocal annelid reported from the Lower Cambrian of China. It represents one of the oldest annelids among...

  12. Cambrian small shelly fossils from the Çal Tepe Formation, Taurus Mountains, Turkey

    OpenAIRE

    Sarmiento, Graciela N.; Fernández Remolar, David Carlos; Göncüoglu, M. Cemal

    2001-01-01

    [EN] Lower and Middle Cambrian carbonate rocks of the Çal Tepe Formation, cropping out in the western Taurus Mountains, yielded a large number of microfossil remains. Small shelly fossils from a single level in the upper Lower Cambrian represent a high diversity biota that could be related to the «Cambrian explosion». Microfossil association from the lower Middle Cambrian sediments of the Çal Tepe Formation is taxonomically very reduced and a dominant taxon is Hadimopanella GEDIK. Th...

  13. Cambrian small shelly fossils from the Çal Tepe Formation, Taurus Mountains, Turkey

    OpenAIRE

    Sarmiento, Graciela; Fernández Remolar, David Carlos; Göncüoglu, M. Cemal

    2001-01-01

    Lower and Middle Cambrian carbonate rocks of the Çal Tepe Formation, cropping out in the western Taurus Mountains, yielded a large number of microfossil remains. Small shelly fossils from a single level in the upper Lower Cambrian represent a high diversity biota that could be related to the «Cambrian explosion». Microfossil association from the lower Middle Cambrian sediments of the Çal Tepe Formation is taxonomically very reduced and a dominant taxon is Hadimopanella GEDIK. This sudden chan...

  14. Antarctica, supercontinents and the palaeogeography of the Cambrian 'explosion'

    Science.gov (United States)

    Dalziel, Ian

    2014-05-01

    Laurentia is bordered by latest Precambrian-Cambrian rifted margins and must therefore have been located within a Precambrian supercontinent. Geochronologic and geochemical evidence indicates that it was attached to parts of the East Antarctic craton within the Rodinian supercontinent in the late Mesoproterozoic. The Mawson craton of Antarctica rifted from the proto-Pacific margin of Laurentia during the Neooproterozoic, colliding with the present 'southern cone' of Laurentia at ~600 Ma along the Shackleton Range suture zone as Gondwana and Laurentia amalgamated to form the ephemeral Pannotia supercontinental assembly at the end of the Precambrian. The abrupt appearance of almost all animal phyla in the fossil record is often colloquially referred to as the Cambrian 'explosion' of life on Earth. It is also named 'Darwin's dilemma,' as he appreciated that this seemingly mysterious event posed a major problem for his theory of evolution by natural selection. It coincided with a time of major marine transgression over all the continents. Although the metazoan 'explosion' is now seen as more protracted than formerly recognized, it is still regarded one of the most critical events in the history of the biosphere. One of the most striking aspects of the earliest Cambrian fossils is geographic differentiation. In particular, the first benthic trilobite faunas on Laurentia, ancestral North America, and the newly amalgamated southern supercontinent of Gondwana are distinctly different. This has led to the suggestion of an unknown vicariant event intervening between an ancestral trilobite clade and higher members that are represented in the fossil record, possibly one related to the breakup of a supercontinent. Igneous rocks along the Panthalassic margin of Gondwana, including South America, southernmost Africa and the Ellsworth-Whitmore crustal block of Antarctica, and along the proto-Appalachian margin of Laurentia indicate that final separation of Laurentia from

  15. Lower Cambrian Black Rock Series and Associated Stratiform Deposits in Southern China

    Institute of Scientific and Technical Information of China (English)

    陈南生; 范德廉; 等

    1990-01-01

    Widespread in more than ten provinces of southern China are the Lower Cambrian black argillaceous-arenaceous rock series consisting of black carbonaceous shales,black carbonaceousargillo-siliceous rocks,black carbonaceous silicolite and black silicolite and black carbonaceous argillaceous siltstone.The Ni,Mo,V,Cu,U,Ba,Ag,P contents of these series are usually several to tens of times higher than their average values in shales.As viewed from lithological sequence,element association and evolution,these black series can be diveided into two types.There occur a variety of deposits in the black rock series,such as phosphorite deposits,Ni-Mo-V polymetal deposits and "Stone coal" seams.According to ore composition,texture and structure,the phosphorite deposits fall into three types,i.e.,thick-bedded,lenticular and nodular,The Ni-Mo-V polymetal deposits,V deposits and reworked sedimentary CU-U-Cd polymetal deposits in terms of their element assiciation and ore genesis.As for the stratiform deposits in the black rock series,three mineralization stages have been recognized,i.e.,the formation of phosphorite deposits,of Ni-Mo polymetal deposits and of V-Cu-U-Cd polymetal deposits as well as of ":stone coal" seams.Evidence strongly suggests that lower organisms have played an important role in mineralization. Our studies indicate that the Lower Cambrian black argillaceous-arenaceous rock series in southern China and their associated stratiform deposits occur generally in the basal parts of a big sedimentary cycle,i.e.,the initial period of a transgression,or in the fine detrital-siliceous formation between two carbonate formatons,formed in a humid climatic zone between two arid ones.As for the environment of deposition,it is suggested that the Lower Cambrian black argillaceous-arenaceous rock series and associated stratiform deposits(excluding thick-bedded phosphorite deposits)were formed mainly in a restricted and poorly-fed sea basin of stagnant shallow water containing

  16. Self-recognition and Ca2+-dependent carbohydrate-carbohydrate cell adhesion provide clues to the cambrian explosion.

    Science.gov (United States)

    Fernàndez-Busquets, Xavier; Körnig, André; Bucior, Iwona; Burger, Max M; Anselmetti, Dario

    2009-11-01

    The Cambrian explosion of life was a relatively short period approximately 540 Ma that marked a generalized acceleration in the evolution of most animal phyla, but the trigger of this key biological event remains elusive. Sponges are the oldest extant Precambrian metazoan phylum and thus a valid model to study factors that could have unleashed the rise of multicellular animals. One such factor is the advent of self-/non-self-recognition systems, which would be evolutionarily beneficial to organisms to prevent germ-cell parasitism or the introduction of deleterious mutations resulting from fusion with genetically different individuals. However, the molecules responsible for allorecognition probably evolved gradually before the Cambrian period, and some other (external) factor remains to be identified as the missing triggering event. Sponge cells associate through calcium-dependent, multivalent carbohydrate-carbohydrate interactions of the g200 glycan found on extracellular proteoglycans. Single molecule force spectroscopy analysis of g200-g200 binding indicates that calcium affects the lifetime (+Ca/-Ca: 680 s/3 s) and bond reaction length (+Ca/-Ca: 3.47 A/2.27 A). Calculation of mean g200 dissociation times in low and high calcium within the theoretical framework of a cooperative binding model indicates the nonlinear and divergent characteristics leading to either disaggregated cells or stable multicellular assemblies, respectively. This fundamental phenomenon can explain a switch from weak to strong adhesion between primitive metazoan cells caused by the well-documented rise in ocean calcium levels at the end of Precambrian time. We propose that stronger cell adhesion allowed the integrity of genetically uniform animals composed only of "self" cells, facilitating genetic constitutions to remain within the metazoan individual and be passed down inheritance lines. The Cambrian explosion might have been triggered by the coincidence in time of primitive animals

  17. High-Precision U-Pb Geochronology and Correlation: An example Using the Neoproterozic-Cambrian Transition

    Science.gov (United States)

    Bowring, S. A.; Grotzinger, J. P.; Amthor, J.; Martin, M. E.

    2001-05-01

    The precise, global correlation of Precambrian and Paleozoic sedimentary rocks can be achieved using temporally calibrated chemostratigraphic records. This approach is essential for determining rates and causes of environmental and faunal change, including mass extinctions. For example, The Neoproterozoic is marked by major environmental change, including periods of global glaciation, large fluctuations in the sequestration of carbon and major tectonic reorganization followed by the explosive diversification of animals in the earliest Cambrian. The extreme climatic change associated with these glaciations have been implicated as a possible trigger for the Cambrian explosion. The recognition of thin zircon-bearing air-fall ash in Neoproterozoic and Cambrian rocks has allowed the establishment of a high-precision temporal framework for animal evolution and is helping to untangle the history of glaciations. In some cases analytical uncertainties translate to age uncertainties of less than 1 Ma and when integrated with chemostratigraphy, the potential for global correlations at even higher resolution. Progress in the global correlation of Neoproterozoic strata has been achieved through the use of C and Sr isotope chemostratigraphy although it has been hampered by a lack of precise geochronological and faunal control. For example, the period from ca 800-580 Ma is characterized by at least two and perhaps as many as four glacial events that are interpreted by many to be global glaciations on a "Snowball Earth". A lack of precise chronological constraints on the number and duration of glaciations, multiple large excursions in the carbon isotopic record, and an absence of detailed biostratigraphy have complicated global correlation and hindered our understanding of this important period of Earth history. However, the ongoing integration of chemostratigraphic and geochronological data are improving temporal resolution and detailed correlations. These data are critical for

  18. Moulting in the lobopodian Onychodictyon from the lower Cambrian of Greenland

    DEFF Research Database (Denmark)

    Topper, Timothy P.; Skovsted, Christian B.; Peel, John S.; Harper, David A. T.

    2013-01-01

    A number of lobopodian taxa from the Cambrian display pairs of sclerotized plates symmetrically positioned along the dorsum of the animal, predominantly above the walking appendages. Most genera were described from complete body fossils exquisitely preserved in the famous Cambrian Lagerstatten, b...

  19. Beyond the Burgess Shale: Cambrian microfossils track the rise and fall of hallucigeniid lobopodians.

    Science.gov (United States)

    Caron, Jean-Bernard; Smith, Martin R; Harvey, Thomas H P

    2013-09-22

    Burgess Shale-type deposits are renowned for their exquisite preservation of soft-bodied organisms, representing a range of animal body plans that evolved during the Cambrian 'explosion'. However, the rarity of these fossil deposits makes it difficult to reconstruct the broader-scale distributions of their constituent organisms. By contrast, microscopic skeletal elements represent an extensive chronicle of early animal evolution--but are difficult to interpret in the absence of corresponding whole-body fossils. Here, we provide new observations on the dorsal spines of the Cambrian lobopodian (panarthropod) worm Hallucigenia sparsa from the Burgess Shale (Cambrian Series 3, Stage 5). These exhibit a distinctive scaly microstructure and layered (cone-in-cone) construction that together identify a hitherto enigmatic suite of carbonaceous and phosphatic Cambrian microfossils--including material attributed to Mongolitubulus, Rushtonites and Rhombocorniculum--as spines of Hallucigenia-type lobopodians. Hallucigeniids are thus revealed as an important and widespread component of disparate Cambrian communities from late in the Terreneuvian (Cambrian Stage 2) through the 'middle' Cambrian (Series 3); their apparent decline in the latest Cambrian may be partly taphonomic. The cone-in-cone construction of hallucigeniid sclerites is shared with the sclerotized cuticular structures (jaws and claws) in modern onychophorans. More generally, our results emphasize the reciprocal importance and complementary roles of Burgess Shale-type fossils and isolated microfossils in documenting early animal evolution. PMID:23902914

  20. Community structure and composition of the Cambrian Chengjiang biota

    Institute of Scientific and Technical Information of China (English)

    2010-01-01

    Based on previously published species data(228 species in over 18 phyla) and field sampling(114 species and 18406 individuals) in the Chengjiang-Haikou-Anning area,we analyzed quantitatively the paleocommunity composition and structure of the Cambrian Chengjiang biota(Cambrian Series 2,eastern Yunnan,China).Arthropods dominate the community both in species diversity(species:37%) and in abundance(individuals:51.8%).Priapulids(individuals:22.6%) and brachiopods(individuals:16.3%) follow in abundance rank.The arthropod Kunmingella douvillei(26.2%),the priapulid Cricocosmia jinning-ensis(15.4%),and the brachiopod Diandongia pista(11%) are the three most abundant species.Ecological analyses show that the community was dominated by epifaunal organisms(species:63%,individuals:68.4%) followed by infaunal organisms(species:11.9%,individuals:25.9%),nektobenthic organisms(species:11.5%,individuals:2.6%),and pelagic organisms(species:5.3%,individuals:3.1%).The diverse feeding strategies,dominated by suspension feeders(species:35.6%,individuals:26.1%) and hunter/scavengers(species:31.1%,individuals:40.4%),indicate the former existence of a complex food chain and intense competition.Epifaunal vagrant omnivores(28.2%),infaunal vagrant hunter/scavengers(19.8%),epifaunal sessile suspension feeders(17.7%),and epifaunal vagrant hunter/scavengers(15.3%) were the most abundant ecological groups,represented primarily by arthropods,poriferans,priapulids,and brachiopods.Ecological group analyses reveal that the early Cambrian Chengjiang biota is similar in community patterns and functional relations to modern biotas in shallow marine settings.

  1. Did homeobox gene duplications contribute to the Cambrian explosion?

    Science.gov (United States)

    Holland, Peter W H

    2015-01-01

    The Cambrian explosion describes an apparently rapid increase in the diversity of bilaterian animals around 540-515 million years ago. Bilaterian animals explore the world in three-dimensions deploying forward-facing sense organs, a brain, and an anterior mouth; they possess muscle blocks enabling efficient crawling and burrowing in sediments, and they typically have an efficient 'through-gut' with separate mouth and anus to process bulk food and eject waste, even when burrowing in sediment. A variety of ecological, environmental, genetic, and developmental factors have been proposed as possible triggers and correlates of the Cambrian explosion, and it is likely that a combination of factors were involved. Here, I focus on a set of developmental genetic changes and propose these are part of the mix of permissive factors. I describe how ANTP-class homeobox genes, which encode transcription factors involved in body patterning, increased in number in the bilaterian stem lineage and earlier. These gene duplications generated a large array of ANTP class genes, including three distinct gene clusters called NK, Hox, and ParaHox. Comparative data supports the idea that NK genes were deployed primarily to pattern the bilaterian mesoderm, Hox genes coded position along the central nervous system, and ParaHox genes most likely originally specified the mouth, midgut, and anus of the newly evolved through-gut. It is proposed that diversification of ANTP class genes played a role in the Cambrian explosion by contributing to the patterning systems used to build animal bodies capable of high-energy directed locomotion, including active burrowing. PMID:26605046

  2. A Precambrian-Cambrian oil play in southern Utah

    Energy Technology Data Exchange (ETDEWEB)

    Lillis, P.G.; Palacas, J.G.; Warden, A. [Geological Survey, Denver, CO (United States)

    1995-06-01

    The potential of the Precambrian Chuar Group as a petroleum source rock in southern Utah and northern Arizona resulted in the drilling of two wildcat wells in 1994. Both wells penetrated the Cambrian Tapeats Sandstone (the target reservoir rock) and presumably Precambrian rocks. The first well, Burnett Federal 36-1, was drilled east of Kanab, Utah (sec.36, T.34S., R.3W.) to a total depth of 5,365 ft and encountered Precambrian (?) reddish-brown sedimentary rocks at 4,790 ft. The Tapeats Sandstone had live oil shows and minor CO{sub 2} (?) gas shows. The second well, BHP Federal 28-1, was drilled near Capitol Reef (sec.28, T.33S., R.7E.) to a total depth of 6,185 ft and encountered the Tapeats Sandstone at 5,922 ft and Precambrian (?) phyllite at 6,125 ft. The upper Paleozoic rocks had abundant live oil/tar shows, and the Cambrian Bright Angel Shale and Tapeats Sandstone had numerous oil shows. There were no gas shows in the well except for a large CO{sub 2} gas kick in the Tapeats Sandstone. A drill-stem test from 5,950 to 6,185 ft yielded mostly CO{sub 2} (92%) and nitrogen gas (6%) and minor amounts of helium, argon, hydrogen, and methane. The {delta}{sup 13}C of the CO{sub 2} is -3.9 per mil PDB. The chemical composition of the extracted oil in the Cambrian sandstones is significantly different than oils produced from the Upper Valley field (upper Paleozoic reservoirs) and the tar sands that are widespread throughout southern and central Utah. However, the oil composition is similar in several aspects to the composition of some of the Precambrian Chuar Group bitumen extracts from the Grand Canyon area in Arizona. The encouraging features of both wells are the good reservoir characteristics and oil shows in the Tapeats Sandstone. In the BHP well the oil appears to be a new oil type, possibly derived from Precambrian or Cambrian source rocks.

  3. The Cambrian explosion triggered by critical turning point in genome size evolution.

    Science.gov (United States)

    Li, Dirson Jian; Zhang, Shengli

    2010-02-01

    The Cambrian explosion is a grand challenge to science today and involves multidisciplinary study. This event is generally believed as a result of genetic innovations, environmental factors and ecological interactions, even though there are many conflicts on nature and timing of metazoan origins. The crux of the matter is that an entire roadmap of the evolution is missing to discern the biological complexity transition and to evaluate the critical role of the Cambrian explosion in the overall evolutionary context. Here, we calculate the time of the Cambrian explosion by a "C-value clock"; our result quite fits the fossil records. We clarify that the intrinsic reason of genome evolution determined the Cambrian explosion. A general formula for evaluating genome size of different species has been found, by which the genome size evolution can be illustrated. The Cambrian explosion, as a major transition of biological complexity, essentially corresponds to a critical turning point in genome size evolution. PMID:20074549

  4. Syn-rift unconformities punctuating the lower-middle Cambrian transition in the Atlas Rift, Morocco

    Science.gov (United States)

    Álvaro, J. Javier; Ezzouhairi, Hassan; Clausen, Sébastien; Ribeiro, M. Luisa; Solá, Rita

    2015-04-01

    The Cambrian Tamdroust and Bab n'Ali Volcanic Complexes represent two magmatic episodes developed in the latest Ediacaran-Cambrian Atlas Rift of Morocco. Their rifting pulses were accompanied by accumulation of volcanosedimentary edifices (dominated by effusive lava flows in the former and explosive acidic aprons in the latter) associated with active tilting and uplift. Sealing of their peneplaned horst-and-graben palaeotopographies led to the onset of distinct onlapping geometries and angular discordances capping eroded basements ranging from the Ediacaran Ouarzazate Supergroup to the Cambrian Asrir Formation. Previous interpretations of these discordances as pull-apart or compressive events are revised here and reinterpreted in an extensional (rifting) context associated with active volcanism. The record of erosive unconformities, stratigraphic gaps, condensed beds and onlapping patterns across the traditional "lower-middle Cambrian" (or Cambrian Series 2-3) transition of the Atlas Rift must be taken into consideration for global chronostratigraphic correlation based on their trilobite content.

  5. Development, triploblastism, physics of wetting and the Cambrian explosion.

    Science.gov (United States)

    Fleury, Vincent

    2013-09-01

    The Cambrian explosion is characterized by the sudden outburst of organized animal plans, which occurred circa 530 M years ago. Around that time, many forms of animal life appeared, including several which have since disappeared. There is no general consensus about "why" this happened, and why it had any form of suddenness. However, all organized animal plans share a common feature: they are triploblastic, i.e., composed of 3 layers of tissue, endoderm, ectoderm and mesoderm. I show here that, within simple hypotheses, the formation of the mesoderm has intrinsically a physical exponential dynamics, leading rapidly to triploblastism, and eventually, to animal formation. A novel physico-mathematical framework including epithelium-mesenchyme transition, visco-elastic constitutive equations, and conservation laws, is presented which allows one to describe gastrulation as a self-wetting phenomenon of a soft solid onto itself. This phenomenon couples differentiation and migration during gastrulation, and leads in a closed form to an exponential scaling law for the formation of the mesoderm. Therefore, the Cambrian explosion might have started, actually, by a true viscoelastic "explosion": the exponential run-away of mesenchymal cells. PMID:23959076

  6. Preservational Pathways of Corresponding Brains of a Cambrian Euarthropod.

    Science.gov (United States)

    Ma, Xiaoya; Edgecombe, Gregory D; Hou, Xianguang; Goral, Tomasz; Strausfeld, Nicholas J

    2015-11-16

    The record of arthropod body fossils is traceable back to the "Cambrian explosion," marked by the appearance of most major animal phyla. Exceptional preservation provides crucial evidence for panarthropod early radiation. However, due to limited representation in the fossil record of internal anatomy, particularly the CNS, studies usually rely on exoskeletal and appendicular morphology. Recent studiesshow that despite extreme morphological disparities, euarthropod CNS evolution appears to have been remarkably conservative. This conclusion is supported by descriptions from Cambrian panarthropods of neural structures that contribute to understanding early evolution of nervous systems and resolving controversies about segmental homologies. However, the rarity of fossilized CNSs, even when exoskeletons and appendages show high levels of integrity, brought into question data reproducibility because all but one of the aforementioned studies were based on single specimens. Foremost among objections is the lack of taphonomic explanation for exceptional preservation of a tissue that some see as too prone to decay to be fossilized. Here we describe newly discovered specimens of the Chengjiang euarthropod Fuxianhuia protensa with fossilized brains revealing matching profiles, allowing rigorous testing of the reproducibility of cerebral structures. Their geochemical analyses provide crucial insights of taphonomic pathways for brain preservation, ranging from uniform carbon compressions to complete pyritization, revealing that neural tissue was initially preserved as carbonaceous film and subsequently pyritized. This mode of preservation is consistent with the taphonomic pathways of gross anatomy, indicating that no special mode is required for fossilization of labile neural tissue. PMID:26526373

  7. Ouachita trough: Part of a Cambrian failed rift system

    Science.gov (United States)

    Lowe, Donald R.

    1985-11-01

    Pre-flysch (Cambrian-Mississippian) strata of the Ouachita Mountains of Arkansas and Oklahoma include two main sandstone lithofacies: (1) a craton-derived lithofacies made up largely of mature medium- to coarse-grained quartzose and carbonate detritus and, in some units, sediment eroded from exposed basement rocks and (2) an orogen-derived facies made up mainly of fine-grained quartzose sedimentary and metasedimentary debris and possibly, in lower units, a volcaniclastic component. Paleocurrent and distribution patterns indicate that detritus of facies I in the Benton uplift was derived from north and detritus of facies II throughout the Ouachitas was derived from south and east of the depositional basin. Overall sedimentological results suggest that the Ouachita trough was a relatively narrow, two-sided basin throughout most and probably all of its existence and never formed the southern margin of the North American craton. Regional comparisons suggest that it was one of several basins, including the Southern Oklahoma aulacogen, Reelfoot Rift, Illinois Basin, and Rome trough, that formed as a Cambrian failed rift system 150 to 250 m.y. after initial rifting along the Appalachian margin of the North American craton.

  8. Fossil Association from the Lower Cambrian Yanjiahe Formation in the Yangtze Gorges Area, Hubei, South China

    Institute of Scientific and Technical Information of China (English)

    GUO Junfeng; Tsuyoshi KOMIYA; LI Yong; HAN Jian; ZHANG Xingliang; ZHANG Zhifei; OU Qiang; LIU Jianni; SHU Degan; Shigenori MARUYAMA

    2008-01-01

    Apart from previously reported Small Shelly Fossils (SSFs), a macroscopic fossil assemblage, comprising abundant algae, cone-shaped tubular fossil forms, and probable impressions of a megascopic metazoan, comes from the Lower Cambrian Yanjiahe Formation in the Yangtze Gorges area of western Hubei Province, south China. The visible fossils are preserved in thin-laminated siltstone or muddy siltstone intercalated between 8-15 mm-thick carbonate deposits, probably representing sedimentary settings of a constrained local depression in the shallow water carbonate platform during the Early Cambrian Meishucunian Stage. The macroscopic fossil association provides significant fossil evidence about the evolution of life from the late Precambrian to the 'Cambrian explosion' interval.

  9. Sponge fossil assemblage from the Early Cambrian Hetang Formation in southern Anhui

    Institute of Scientific and Technical Information of China (English)

    CHEN Zhe; HU Jie; ZHOU Chuanming; XIAO Shuhai; YUAN Xunlai

    2004-01-01

    Abundant well-preserved large articulated sponge fossils and isolated spicules have been reported from the Early Cambrian Hetang Formation, southern Anhui Province. This unique epifaunal fossil assemblage dominated by articulated sponge fossils is called the Xidi Sponge Fauna.The sponge fauna lived in a quiet oxygenic environment below the storm wave base. Bloom of phytoplankton and rapid sedimentation rate resulted in the deposition of the black shales. Sufficient food supply, lack of other competitors,abundant ecological niches, and demand for oxygen during early Cambrian were in favor of the diversification and evolution of large sponges in the Early Cambrian.

  10. Phylogenomic insights into the cambrian explosion, the colonization of land and the evolution of flight in arthropoda.

    Science.gov (United States)

    Wheat, Christopher W; Wahlberg, Niklas

    2013-01-01

    The timing of the origin of arthropods in relation to the Cambrian explosion is still controversial, as are the timing of other arthropod macroevolutionary events such as the colonization of land and the evolution of flight. Here we assess the power of a phylogenomic approach to shed light on these major events in the evolutionary history of life on earth. Analyzing a large phylogenomic dataset (122 taxa, 62 genes) with a Bayesian-relaxed molecular clock, we simultaneously reconstructed the phylogenetic relationships and the absolute times of divergences among the arthropods. Simulations were used to test whether our analysis could distinguish between alternative Cambrian explosion scenarios with increasing levels of autocorrelated rate variation. Our analyses support previous phylogenomic hypotheses and simulations indicate a Precambrian origin of the arthropods. Our results provide insights into the 3 independent colonizations of land by arthropods and suggest that evolution of insect wings happened much earlier than the fossil record indicates, with flight evolving during a period of increasing oxygen levels and impressively large forests. These and other findings provide a foundation for macroevolutionary and comparative genomic study of Arthropoda. PMID:22949483

  11. Pulse of atmospheric oxygen during the late Cambrian.

    Science.gov (United States)

    Saltzman, Matthew R; Young, Seth A; Kump, Lee R; Gill, Benjamin C; Lyons, Timothy W; Runnegar, Bruce

    2011-03-01

    A rise in atmospheric O(2) has been linked to the Cambrian explosion of life. For the plankton and animal radiation that began some 40 million yr later and continued through much of the Ordovician (Great Ordovician Biodiversification Event), the search for an environmental trigger(s) has remained elusive. Here we present a carbon and sulfur isotope mass balance model for the latest Cambrian time interval spanning the globally recognized Steptoean Positive Carbon Isotope Excursion (SPICE) that indicates a major increase in atmospheric O(2). We estimate that this organic carbon and pyrite burial event added approximately 19 × 10(18) moles of O(2) to the atmosphere (i.e., equal to change from an initial starting point for O(2) between 10-18% to a peak of 20-28% O(2)) beginning at approximately 500 million years. We further report on new paired carbon isotope results from carbonate and organic matter through the SPICE in North America, Australia, and China that reveal an approximately 2‰ increase in biological fractionation, also consistent with a major increase in atmospheric O(2). The SPICE is followed by an increase in plankton diversity that may relate to changes in macro- and micronutrient abundances in increasingly oxic marine environments, representing a critical initial step in the trophic chain. Ecologically diverse plankton groups could provide new food sources for an animal biota expanding into progressively more ventilated marine habitats during the Ordovician, ultimately establishing complex ecosystems that are a hallmark of the Great Ordovician Biodiversification Event. PMID:21368152

  12. From snowball earth to the cambrian explosion: the interpretative potential of the isotope record

    Science.gov (United States)

    Shields, G.

    2003-04-01

    The interval in Earth history between 750 Ma and 530 Ma represents the erratic transition between the distinctly different "Earth systems" of the Proterozoic and the Phanerozoic. Geologists have developed several indirect methods to study this key interval but perhaps none with as much wanton enthusiasm as isotope geochemistry. Here I attempt a review of the isotope record across this key transition period, with special emphasis on the interpretative role that isotopes can play in understanding events such as the "Snowball Earth" glaciations and the "Cambrian Explosion". The marine isotope record reveals the Proterozoic-Phanerozoic transition to have been a unique period in Earth history with first-order features in all three of the major isotope proxies: 87Sr/86Sr, 34S/32S and 13C/12C ratios. Seawater 87Sr/86Sr, increased sharply between about 900 Ma and 500 Ma from 0.704 to 0.709, the largest rise ever. Although attributed to increasing rates of continental erosion, this remains only the most plausible explanation because few aspects of the rise have been satisfactorily linked to particular tectonic events. Much of the uncertainty surrounding the Proterozoic 87Sr/86Sr record is due to poor age dating and discrepancies in stratigraphic correlation schemes, which together with diagenetic alteration have led to the current inability to realise the potential of the 87Sr/86Sr record for global stratigraphic correlation. Carbon isotopes have played a key role in recent palaeoenvironmental interpretations of the Proterozoic-Phanerozoic transition, most of which have already been discarded or merely forgotten. Nevertheless, it is clear that the extraordinary δ13C values of the later Neoproterozoic (and the earliest Cambrian) demand extraordinary explanations, for which there is still much scope for future invention. Carbon isotopes may also play an important supporting role in correlation, however, the role that δ13C may play in global stratigraphic correlation has

  13. A review of the Late Cambrian (Furongian) palaeogeography in the western Mediterranean region, NW Gondwana

    Science.gov (United States)

    Álvaro, J. Javier; Ferretti, Annalisa; González-Gómez, Cristina; Serpagli, Enrico; Tortello, M. Franco; Vecoli, Marco; Vizcaïno, Daniel

    2007-11-01

    The Cambrian-Ordovician transition of the western Mediterranean region (NW Gondwana) is characterized by the record of major erosive unconformities with gaps that range from a chronostratigraphic stage to a series. The hiatii are diachronous and involved progressively younger strata along the Gondwanan margin, from SW (Morocco) to NE (Montagne Noire). They can be related to development of a multi-stage rifting (further North), currently connected to the opening of the Rheic Ocean, and concomitant erosion on southern rift shoulders. The platforms of this margin of Gondwana occupied temperate-water, mid latitudes and were dominated by siliciclastic sedimentation, while carbonate factories were only episodically active in the Montagne-Noire platform. The Upper Cambrian is devoid of significant gaps in the southern Montagne Noire and the Iberian Chains. There, the sedimentation took place in a transgressive-dominated depositional system, with common offshore deposits and clayey substrates, and was bracketed by two major regressive trends. The Late Cambrian is also associated with the record of volcanic activity ( e.g., in the Cantabrian and Ossa-Morena zones, and the northern Montagne Noire), and widespread development of a tectonic instability that led to the episodic establishment of palaeotopographies and record of slope-related facies associations. Several immigration events are recognized throughout the latest Middle Cambrian, Late Cambrian and Tremadocian. The trilobites show a stepwise replacement of Acado-Baltic-type families ( e.g., the conocoryphid-paradoxidid-solenopleurid assemblage) characterized by: (i) a late Languedocian (latest Middle Cambrian) co-occurrence of Middle Cambrian trilobite families with the first anomocarid, dorypygid and proasaphiscid invaders; (ii) a Late Cambrian immigration replacing previous faunas, composed of trilobites (aphelaspidids, catillicephalids, ceratopygids, damesellids, eulomids, idahoiids, linchakephalids, lisariids

  14. Discovery of hydrothermal venting community at the base of Cambrian barite in Guizhou Province, Western China:Implication for the Cambrian biological explosion

    Institute of Scientific and Technical Information of China (English)

    2008-01-01

    The barite deposit in the Niutitang Formation of Lower Cambrian, in Tianzhu County, Guizhou Province, western China, is a superlarge barite deposit with about 2 × 109 tons of reserves: Mineral, petrological and geochemical studies reveal that this barite deposit belongs to a hydrothermal sedimentary deposit. Microscopic observations indicate that a lot of algae, sponge spicules and tube-type fossils are well-preserved in this barite section, and moreover, those fossils share most characteristics of the deep-sea hydrothermal venting community in the modern Pacific. We suggest that the hydrothermal venting community was thriving in hydrothermal vent in early Cambrian, and it is of great significance for elucidating the geological background of the "Cambrian explosion".

  15. Early Cambrian origin of modern food webs: evidence from predator arrow worms

    OpenAIRE

    Vannier, J; Steiner, M.; Renvoisé, E; Hu, S.-X.; Casanova, J.-P

    2006-01-01

    Although palaeontological evidence from exceptional biota demonstrates the existence of diverse marine communities in the Early Cambrian (approx. 540–520 Myr ago), little is known concerning the functioning of the marine ecosystem, especially its trophic structure and the full range of ecological niches colonized by the fauna. The presence of a diverse zooplankton in Early Cambrian oceans is still an open issue. Here we provide compelling evidence that chaetognaths, an important element of mo...

  16. The Precambrian–Cambrian nonconformity at the Ercall Quarries, The Wrekin, Shropshire, UK

    OpenAIRE

    Pringle, JK; Stimpson, IG

    2013-01-01

    The Precambrian–Cambrian boundary is iconic, marking the first appearance of shelly fauna in the fossil record, and opening the Phanerozoic Eon. In England and Wales, the transition from predominantly Precambrian igneous rocks to Cambrian sedimentary strata is generally unconformable. An exceptional exposure of this transition can be observed in the Ercall Quarries in Shropshire, a classic locality in all senses of the word.

  17. Exceptionally preserved brachiopods from the Chengjiang Lagerstatte (Yunnan, China) : Perspectives on the Cambriane explosion of metazoans

    OpenAIRE

    Zhang, Zhifei; Holmer, L. E.

    2013-01-01

    The Cambrian explosion was coined to describe the geologically sudden appearance of numerous bilaterian body plans (Phyla) around the Ediacaran-Cambrian transition, around 565-520 million years ago. Many explanations and conjectures have been postulated in order to explain the pattern and duration of this explosive radiation of many different phyla of early metazoans. Here, we focus on the evolution of a phylum of marine suspension-feeding animals – the brachiopods, as exemplified by the exce...

  18. The Genetic Response to Snowball Earth: Role of HSP90 in the Cambrian Explosion

    OpenAIRE

    Baker, Michael E.

    2006-01-01

    The events that shaped the Cambrian explosion from 545 to 530 myr ago, when multicellular animals suddenly appeared in the fossil record, are not fully understood. It is likely that the evolution of new transcription factors and other signal transduction proteins that regulated developmental networks were important in the emergence of diverse animal phyla seen in the Cambrian. I propose that one or both extensive glaciations that ended about 670 myr and 635 myr ago were important in the evo...

  19. The C-value enigma and timing of the Cambrian explosion

    OpenAIRE

    Li, Dirson Jian; Zhang, Shengli

    2008-01-01

    The Cambrian explosion is a grand challenge to science today and involves multidisciplinary study. This event is generally believed as a result of genetic innovations, environmental factors and ecological interactions, even though there are many conflicts on nature and timing of metazoan origins. The crux of the matter is that an entire roadmap of the evolution is missing to discern the biological complexity transition and to evaluate the critical role of the Cambrian explosion in the overall...

  20. A sclerite-bearing stem group entoproct from the early Cambrian and its implications

    OpenAIRE

    Zhifei Zhang; Holmer, Lars E.; Skovsted, Christian B; Glenn A. Brock; Graham E. Budd; Dongjing Fu; Xingliang Zhang; Degan Shu; Jian Han; Jianni Liu; Haizhou Wang; Aodhán Butler; Guoxiang Li

    2013-01-01

    The Lophotrochozoa includes disparate tentacle-bearing sessile protostome animals, which apparently appeared in the Cambrian explosion, but lack an uncontested fossil record. Here we describe abundant well preserved material of Cotyledion tylodes Luo et Hu, 1999, from the Cambrian (Series 2) Chengjiang deposits, reinterpreted here as a stem-group entoproct. The entoproct affinity is supported by the sessile body plan and interior soft anatomy. The body consists of an upper calyx and a lower e...

  1. Lower Cambrian polychaete from China sheds light on early annelid evolution

    Science.gov (United States)

    Liu, Jianni; Ou, Qiang; Han, Jian; Li, Jinshu; Wu, Yichen; Jiao, Guoxiang; He, Tongjiang

    2015-06-01

    We herein report a fossilized polychaete annelid, Guanshanchaeta felicia gen. et sp. nov., from the Lower Cambrian Guanshan Biota (Cambrian Series 2, stage 4). The new taxon has a generalized polychaete morphology, with biramous parapodia (most of which preserve the evidence of chaetae), an inferred prostomium bearing a pair of appendages, and a bifid pygidium. G. felicia is the first unequivocal annelid reported from the Lower Cambrian of China. It represents one of the oldest annelids among those from other early Paleozoic Lagerstätten including Sirius Passet from Greenland (Vinther et al., Nature 451: 185-188, 2008) and Emu Bay from Kangaroo island (Parry et al., Palaeontology 57: 1091-1103, 2014), and adds to our increasing roll of present-day animal phyla recognized in the early Cambrian Guanshan Biota. This finding expands the panorama of the Cambrian `explosion' exemplified by the Guanshan Biota, suggesting the presence of many more fossil annelids in the Chengjiang Lagerstätte and the Kaili Biota. In addition, this new taxon increases our knowledge of early polychaete morphology, which suggests that polychaete annelids considerably diversified in the Cambrian.

  2. A Euxinic-Wedge Model for Mo Cycling in the Early Cambrian Oceans

    Science.gov (United States)

    Li, C.; Cheng, M.; Zhou, L.; Algeo, T. J.; Jin, C. S.; Lei, L. D.; Feng, L. J.; Jiang, S. Y.

    2015-12-01

    Ocean redox conditions during the early Cambrian, a critical period in Earth history marked by rapid evolution of animals, are uncertain. Previous studies have suggested well oxic conditions based on Mo isotopes and stratified, anoxic deep waters based on Fe-S-C and trace metal data. The Niutitang Formation (529-514 Ma) at Yangjiaping (South China) consists of a lower black shale member (LM, 0-18 m) and an upper gray silty shale member (UM, 24-80 m). The LM is moderately to strongly enriched in Mo, U and V, and heterogeneous in Mo-isotopic compositions (δ98Mo = ‒0.65‰ to +2.14‰), while the UM shows lesser enrichment of Mo, U and V, higher Mo/U ratios, and intermediate and more uniform δ98Mo (+1.16‰ to +1.71‰). The metal data and previously reported Fe-speciation data suggest a redox change from dominantly euxinic conditions for the LM to suboxic or weakly anoxic-euxinic conditions for the UM, consistent with a shift of the chemocline from the water column to the sediment during a eustatic fall. We propose an "euxinic wedge" model for the δ98Mo patterns observed here. In this model, the early Cambrian ocean exhibited a lateral transition from ferruginous conditions in nearshore areas to euxinic conditions at mid-depths and back to ferruginous conditions in offshore areas. This pattern was the result of terrigenous Fe-Mnox reduction, sulfate resupply via rivers, and hydrothermal inputs of reactive Fe into the deep basin. Within this framework, sedimentary δ98Mo variation in the study units can be explained through the combined effects of Mo isotopic fractionation associated with Fe-Mnox adsorption and depth-dependent H2S concentrations. Importantly, the heaviest δ98Mo values, which were previously interpreted to reflect full oxygenation, can be explained by the operation of a globally widespread Fe-Mn particulate shuttle that removed large quantities of isotopically light Mo to the sediment, leaving seawater enriched in 98Mo.

  3. Reconstructing redox conditions of mesoproterozoic-early Cambrian seawater using δ98/95Mo

    International Nuclear Information System (INIS)

    Full text: Mo isotope composition of seawater is determined by its relative removal in oxic and anoxic sinks as Mo isotope fractionates during its removal from seawater in oxic sediments whereas it remains invariable during its precipitation in reducing sediments. Hence, Mo isotope composition of reducing sediments can be used to track the temporal variation in redox condition of the global ocean. In this study an attempt is being made to reconstruct the redox conditions during Mesoproterozoic-Early Cambrian period using δ98/95∼Mo in black shales from Vindhyan and Outer belt of Lesser Himalaya. This time span witnessed the important evolutionary changes in the early earth such as rise in atmospheric oxygen and changes in the biosphere. Samples processed for this study were collected from Bijaigarh Shales of Kaimur formation of the Vindhyan and Lower Tal formation of the Lesser Himalaya. The depositional ages of these formations was estimated to be -1200 Ma and -540 Ma for Bijaigarh and Tal formations respectively. Mo isotope composition of black shales were determined by their complete acid digestion in presence of 97Mo-100Mo double spike followed by column chromatography separation. Mo isotope composition in purified samples were analysed using Finnigan Neptune MC-ICP-MS at Physical Research Laboratory. δ98/95% Mo measured in Bijaigarh Shales indicate variation from 0.4 to 1.35% with concentration ranging from 1.6-34 ppm whereas the Tal black shales display variability of 0.3-1.5%0 with concentration 16-80 ppm. In general, higher 09%sMo in Bijaigarh shales are associated with higher Mo concentration indicating the dominance of seawater Mo. Lower δ99/95%Mo in some samples could result from detrital component. Based on these data it can be inferred that Mesoproterozoic ocean was more anoxic compared to that of Early Cambrian and the contemporary ocean. More work is underway to track and further quantify the anoxicity of the ocean with time

  4. Carbon isotope fluctuations through the neoproterozoic-lower cambrian Birmania basin, Rajasthan, India

    International Nuclear Information System (INIS)

    The paleogeographic reconstruction of the end phase of Neoproterozoic assumes a significance due to the fact that the interactions between the biosphere and the atmosphere, hydrosphere and geosphere during this period of time strongly influenced the evolution of life and initiated a remarkable phase of organic evolution well documented in the rocks of this age (Banerjee and Majumdar, 1999). Time related changes in these carbon reservoirs and processes during the end Proterozoic have played an important role in the evolution of the environment and of life (Des Marais, 1997). Brasier (1992) discussed at length various factors that pushed shallow marine ecosystem of the late Neoproterozoic-Cambrian towards P-limitation and suggested that nitrate fixation, density stratification and massive removal of P in the sediments were responsible for the formation of phosphatic deposits within this time interval. East Gondwana assembly of continents provide fine examples of stratigraphic successions in which the ideas related to oceanic stratification, phosphate deposition, enhanced organic production and carbon isotope fluctuations can be verified and correlated from continent to continent (Banerjee and Majumdar, 1999). The Birmania basin is an oval shaped isolated remnant of the Marwar basin (Neo-proterozoic-Early Palaeozoic) located in the heart of the Thar desert of western Rajasthan, India. It is underlain by Malani Igneous Suite of rocks which range in age from 780 to 680Ma (Rathore et al., 1999).The Birmania basin comprises around 900 metre thick sedimentary sequence of siliciclastic, carbonate and phosphorite facies. These sequences are unconformably overlain by La-thi conglomerate of Jurassic age in north-ern flank of the basin. Global Neoproterozoic glacial activity in the western Rajasthan is also reported and is represented by Pokaran Boulder Bed, few kms away from study area. The Pokaran Boulder Bed overlies the Malani Igneous Suite and is mainly consist of boulders

  5. Body Size Evolution in Conodonts from the Cambrian through the Triassic

    Science.gov (United States)

    Schaal, E. K.; Morgan, D. J.; Payne, J.

    2013-12-01

    The size of an organism exercises tremendous control over its physiology, life history, and ecology, yet the factors that influence body size evolution remain poorly understood. One major limitation is the lack of appropriate datasets spanning long intervals of evolutionary time. Here, we document size trends in conodonts (tooth-like microfossils from marine chordates) because they evolved rapidly and are known to change size during intervals of environmental change. By measuring photographs from the Catalogue of Conodonts (Ziegler 1982), we compiled a database of conodont P1 element measurements for 575 species and subspecies from the Cambrian through Triassic periods. Because tooth size correlates with body size in conodont animals and their extant relatives, conodont element length can serve as a proxy for the size of the conodont animal. We find that mean and maximum size across species increased during the early Paleozoic, peaked during the Devonian-Mississippian, and then generally decreased until conodonts went extinct at the end of the Triassic. We used regression analyses to compare conodont mean size trends to potential environmental predictors, such as changing atmospheric pO2, atmospheric pCO2, and sea level. Conodont size exhibited poor correlation with these environmental factors, suggesting that conodont evolution may have been more strongly influenced by other environmental covariates or ecological variables such as predation and competition.

  6. Nonlinear geosphere-biosphere interactions and the Cambrian explosion

    Science.gov (United States)

    von Bloh, W.; Bounama, C.; Franck, S.

    2003-04-01

    A conceptual model for the global carbon cycle of the Earth containing the reservoirs mantle, ocean floor, continental crust, continental biosphere, the kerogen , as well as the aggregated reservoir ocean and atmosphere is presented. In this study the evolution of the mean global surface temperature, the biomass, and reservoir sizes over the whole history and future of the Earth under a maturing Sun is investigated. Reasonable values for the present distribution of carbon in the surface reservoirs of the Earth are obtained and a pronounced global minimum of mean surface temperature at the present state of the Earth is found. Furthermore, three different biosphere types are introduced: procaryotes, eucaryotes, and higher metazoa. They all differ in their temperature tolerance interval and their biogenic enhancement of silicate rock weathering. Around 500 Myr in the past we find a rise of higher metazoa caused by the nonlinear feedback between biosphere and climate. Biotic amplifying of weathering provides and maintains the environment of higher life forms. Such a mechanism may explain the so-called Cambrian explosion.

  7. Literary Crisis or Cambrian Period? Theory, Practice, and Public Policy Implications.

    Science.gov (United States)

    Manzo, Anthony V.

    2003-01-01

    Contends that there is no literacy crisis in the United States. Notes that research on college-level proficient readers has indicated that reading educators are doing a better job of educating students to become mature readers and that even at the elementary level the news is better than most people realize. Suggests that the conception of reading…

  8. Ichnology of the Early Cambrian Tal Group, Mussoorie Syncline, Lesser Himalaya, India

    Indian Academy of Sciences (India)

    Meera Tiwari; S K Parcha; Rajita Shukla; Harshita Joshi

    2013-12-01

    The Lesser Himalayan sequence is considered as one of the best developed sections of Cambrian successions, exposed in five different synclines. Mussoorie Syncline, being one of the five synclines, exposes the Cambrian Tal Group. This paper describes nine ichnotaxa, viz., Dimorphichnus isp., ?Diplichnites isp., Monomorphichnus isp., Nereites isp., Palaeopasichnus isp., Palaeophycus isp., Planolites montanus, Planolites isp., Skolithos isp., Treptichnus isp. from the Dhaulagiri Formation of the Tal Group. A detailed analysis of the ichnofossils indicates that the entire succession of Tal Group reflects shallow marine conditions in general and Mussoorie Syncline in particular. The above ichnofossil assemblage along with earlier ichnofossils and other faunal occurrences substantiates the assignment of Early Cambrian age to the Dhaulagiri Formation.

  9. Ichnology of the Early Cambrian Tal Group, Mussoorie Syncline, Lesser Himalaya, India

    Science.gov (United States)

    Tiwari, Meera; Parcha, S. K.; Shukla, Rajita; Joshi, Harshita

    2013-12-01

    The Lesser Himalayan sequence is considered as one of the best developed sections of Cambrian successions, exposed in five different synclines. Mussoorie Syncline, being one of the five synclines, exposes the Cambrian Tal Group. This paper describes nine ichnotaxa, viz., Dimorphichnus isp., ? Diplichnites isp., Monomorphichnus isp., Nereites isp., Palaeopasichnus isp., Palaeophycus isp., Planolites montanus, Planolites isp., Skolithos isp., Treptichnus isp. from the Dhaulagiri Formation of the Tal Group. A detailed analysis of the ichnofossils indicates that the entire succession of Tal Group reflects shallow marine conditions in general and Mussoorie Syncline in particular. The above ichnofossil assemblage along with earlier ichnofossils and other faunal occurrences substantiates the assignment of Early Cambrian age to the Dhaulagiri Formation.

  10. Cambrian Sequence Stratigraphy and Sea Level Cycles of North China Platform

    Institute of Scientific and Technical Information of China (English)

    1999-01-01

    The Cambrian of the North China platform consists chiefly of shallow water deposits and shows the sedimentary characters of an epicontinental sea basin. Controlled mainly by global sea level changes and sedimentary influx, the depositional sequences all exhibit as composite sequences. From bottom upward, 14 sequences (3rd order) are recognized, which may be grouped into 5 sequence sets and further into 2 mesosequences (2nd order). It is suggested herein that the Cambrian/Ordovician boundary may better be set at the MFS (maximum flooding surface) of the sequence OSq1, above which the conodont Cordylodus lindstroemi occurs. This position is about 40 m above the traditional Cambrian/Ordovician boundary and is within the Yeli Formation.

  11. Sponge spicules in phosphorites of the Early Cambrian Gezhongwu Formation, Zhijin, Guizhou

    Institute of Scientific and Technical Information of China (English)

    YANG Ruidong; QIAN Yi; ZHANG Jie; ZHANG Weihua; JIANG Lijun; GAO Hui

    2004-01-01

    Phosphorites occurring at the bottom of the Cambrian system contain abundant small shelly fossils, which are the product of the first episode of life explosion in the Cambrian. It was previously reported that the small shelly fossils are dominated by hyolithids, olivooids, zhijinitids, conodontomorphs, yubelichitids, camenitids and algae, with minor amounts of sponge fossils. Large amounts of sponge spicules, diverse in form, have been found for the first time in the Gezhongwu Formation phosphorites at Shixing, Zhijin County, Guizhou Province, of which such spicules as diaxon-triactins, diaxon-tetractins, pentaxon-pentactins and hexon-hexactins account for 30%. These spicules constitute the sponge clastic phosphorites made up of sponge clastics. Meanwhile, it is also expected that the radiation and diversity of sponge animals started as early as in the earliest Early Cambrian. Habit and burying environment of sponge animal are discussed in the paper.

  12. MicroRNAs and metazoan macroevolution: insights into canalization, complexity, and the Cambrian explosion.

    Science.gov (United States)

    Peterson, Kevin J; Dietrich, Michael R; McPeek, Mark A

    2009-07-01

    One of the most interesting challenges facing paleobiologists is explaining the Cambrian explosion, the dramatic appearance of most metazoan animal phyla in the Early Cambrian, and the subsequent stability of these body plans over the ensuing 530 million years. We propose that because phenotypic variation decreases through geologic time, because microRNAs (miRNAs) increase genic precision, by turning an imprecise number of mRNA transcripts into a more precise number of protein molecules, and because miRNAs are continuously being added to metazoan genomes through geologic time, miRNAs might be instrumental in the canalization of development. Further, miRNAs ultimately allow for natural selection to elaborate morphological complexity, because by reducing gene expression variability, miRNAs increase heritability, allowing selection to change characters more effectively. Hence, miRNAs might play an important role in shaping metazoan macroevolution, and might be part of the solution to the Cambrian conundrum. PMID:19472371

  13. Articulated Wiwaxia from the Cambrian Stage 3 Xiaoshiba Lagerstätte

    Science.gov (United States)

    Yang, Jie; Smith, Martin R.; Lan, Tian; Hou, Jin-Bo; Zhang, Xi-Guang

    2014-04-01

    Wiwaxia is a bizarre metazoan that has been interpreted as a primitive mollusc and as a polychaete annelid worm. Extensive material from the Burgess Shale provides a detailed picture of its morphology and ontogeny, but the fossil record outside this lagerstätte is scarce, and complete wiwaxiids are particularly rare. Here we report small articulated specimens of Wiwaxia foliosa sp. nov. from the Xiaoshiba fauna (Cambrian Stage 3, Hongjingshao Formation, Kunming, south China). Although spines are absent, the fossils' sclerites - like those of W. corrugata - are symmetrically arranged in five distinct zones. They form rows across the body, and were individually added and shed throughout growth to retain an approximately symmetrical body shape. Their development pattern suggests a molluscan affinity. The basic body plan of wiwaxiids is fundamentally conserved across two continents through Cambrian Stages 3-5 - revealing morphological stasis in the wake of the Cambrian explosion.

  14. A sclerite-bearing stem group entoproct from the early Cambrian and its implications.

    Science.gov (United States)

    Zhang, Zhifei; Holmer, Lars E; Skovsted, Christian B; Brock, Glenn A; Budd, Graham E; Fu, Dongjing; Zhang, Xingliang; Shu, Degan; Han, Jian; Liu, Jianni; Wang, Haizhou; Butler, Aodhán; Li, Guoxiang

    2013-01-01

    The Lophotrochozoa includes disparate tentacle-bearing sessile protostome animals, which apparently appeared in the Cambrian explosion, but lack an uncontested fossil record. Here we describe abundant well preserved material of Cotyledion tylodes Luo et Hu, 1999, from the Cambrian (Series 2) Chengjiang deposits, reinterpreted here as a stem-group entoproct. The entoproct affinity is supported by the sessile body plan and interior soft anatomy. The body consists of an upper calyx and a lower elongate stalk with a distal holdfast. The soft anatomy includes a U-shaped gut with a mouth and aboral anus ringed by retractable marginal tentacles. Cotyledion differs from extant entoprocts in being larger, and having the calyx and the stalk covered by numerous loosely-spaced external sclerites. The description of entoprocts from the Chengjiang biota traces the ancestry of yet another lophotrochozoan phylum back to the Cambrian radiation, and has important implications for the earliest evolution of lophotrochozoans. PMID:23336066

  15. Biodiversity and taphonomy of the Early Cambrian Guanshan biota,eastern Yunnan

    Institute of Scientific and Technical Information of China (English)

    Michael; STEINER

    2010-01-01

    The Guanshan biota from eastern Yunnan(Cambrian Series 2,early Stage 4) is a Burgess Shale-type fossil biota with abundant exceptionally preserved soft-bodied fossils after the discovery of the well-known Cambrian Chengjiang fauna and Kaili biota in South China.The geological settings,sedimentology,taphonomy,and the fossil assemblage of the Guanshan biota are briefly summarized here.The Guanshan biota consists of about 60 taxa belonging to more than 10 metazoan groups and algae,among which the lobopods,eldonids,hyolithids with helens,and green algae are reported for the first time.The common occurrence of soft-bodied preservation in many groups,notably the trilobites and brachiopods,makes the Guanshan biota a significant fossil lagersttte for understanding the metazoan evolution during Cambrian explosion and taphonomy of the Burgess Shale-type fossils.

  16. Comparative analysis of methods used to define eustatic variations in outcrop: Late Cambrian interbasinal sequence development

    Energy Technology Data Exchange (ETDEWEB)

    Osleger, D. (Univ. of California, Riverside (United States)); Read, J.F. (Virginia Polytechnic Inst. and State Univ., Blacksburg (United States))

    1993-03-01

    Interbasinal correlation of Late Cambrian cyclic carbonates from the Appalachian and Cordilleran passive margins, the Texas craton, and the southern Oklahoma aulacogen defines six major third-order depositional sequences. Graphic correlation of biostratigraphically-constrained strata was used to establish equivalency of stratigraphic sequences between the individual sections. Relatively isochronous biomere boundaries were used as time datums for lithostratigraphic correlation. Although the individual sections are composed of different types of meter-scale cycles and component lithofacies that reflect the various environmental settings of the localities, the overall upward-shallowing character of individual sequences is evident. The sequences are: late Cedaria, mid-Crepicephalus, late Crepicephalus, Aphelaspis to earliest Elvinia, Elvinia to early Saukia, and Saukia to the Cambrian-Ordovician boundary. Interbasinal correlation of stratigraphic sequences permits an evaluation of quantitative techniques for determining accommodation history. Correlation of Fischer plots of cyclic successions from separate basins supports a eustatic control of Late Cambrian sequence development. R2/R3 curves derived from subsidence analysis of the Late Cambrian sections provide good resolution of the second- and third-order scales of accommodation change, and interbasinal correlations of R2/R3 curves also support eustatic control on sequence development. Comparing the accomodation curves and subsidence analysis with paleobathymetric trends of Late Cambrian cyclic strata suggests that the curves may approximate the form of the eustatic sealevel signal. A composite eustatic sealevel curve for Late Cambrian time in North America was created by qualitatively combining the accommodation curves defined by the different techniques for each of the four localities. 129 refs., 16 figs., 3 tabs.

  17. Biological and Molecular Geochemical Evidence for Dinoflagellate Ancestors in the Upper Sinian-Cambrian

    Institute of Scientific and Technical Information of China (English)

    2000-01-01

    Dinoflagellates are single celled organisms that reflect the ecological conditions in modem oceans and lakes. Their earliest undisputed fossil record suggests that dinoflagellates originated from the Middle Triassic (c.240 Ma ago). However, the presence of molecular biomarkers (dinosterane, 4α-methyl-24-ethylcholestane and triaromatic dinosteroids) in rock extracts and coccoid dinoflagellate fossils from the upper Sinian to Cambrian of the Tarim basin confirms the hypothesis that dinoflagellates have an ancient origin, and predate the oldest undisputed dinoflagellate fossils at least by 300 Ma, as early as the late Sinian-Cambrian.

  18. Discovery of sponge body fossils from the late Meishucunian (Cambrian) at Jinsha, Guizhou, south China

    Institute of Scientific and Technical Information of China (English)

    YANG Xinglian; ZHAO Yuanlong; WANG Yue; WANG Pingli

    2005-01-01

    Here we report discovery of a sponge body fossil Triticispongia sp. from the base of lower Cambrian Niutitang Formation at Jinsha, Guizhou. Stratigraphically, the fossil horizon is located below Ni-Mo ore layer with the Niutitang Biota above, and is equivalent to the late Meishucunian. The species is global in shape with skeletons composed of stauractins and monaxons. Triticispongia sp. reported here may be the earliest sponge body fossils of Cambrian, which provides new informationfor understanding early evolution and radiation of sponge animals.

  19. Extraordinary fossils reveal the nature of Cambrian life: a commentary on Whittington (1975) 'The enigmatic animal Opabinia regalis, Middle Cambrian, Burgess Shale, British Columbia'.

    Science.gov (United States)

    Briggs, Derek E G

    2015-04-19

    Harry Whittington's 1975 monograph on Opabinia was the first to highlight how some of the Burgess Shale animals differ markedly from those that populate today's oceans. Categorized by Stephen J. Gould as a 'weird wonder' (Wonderful life, 1989) Opabinia, together with other unusual Burgess Shale fossils, stimulated ongoing debates about the early evolution of the major animal groups and the nature of the Cambrian explosion. The subsequent discovery of a number of other exceptionally preserved fossil faunas of Cambrian and early Ordovician age has significantly augmented the information available on this critical interval in the history of life. Although Opabinia initially defied assignment to any group of modern animals, it is now interpreted as lying below anomalocaridids on the stem leading to the living arthropods. This commentary was written to celebrate the 350th anniversary of the journal Philosophical Transactions of the Royal Society. PMID:25750235

  20. Extraordinary fossils reveal the nature of Cambrian life: a commentary on Whittington (1975) ‘The enigmatic animal Opabinia regalis, Middle Cambrian, Burgess Shale, British Columbia’

    Science.gov (United States)

    Briggs, Derek E. G.

    2015-01-01

    Harry Whittington's 1975 monograph on Opabinia was the first to highlight how some of the Burgess Shale animals differ markedly from those that populate today's oceans. Categorized by Stephen J. Gould as a ‘weird wonder’ (Wonderful life, 1989) Opabinia, together with other unusual Burgess Shale fossils, stimulated ongoing debates about the early evolution of the major animal groups and the nature of the Cambrian explosion. The subsequent discovery of a number of other exceptionally preserved fossil faunas of Cambrian and early Ordovician age has significantly augmented the information available on this critical interval in the history of life. Although Opabinia initially defied assignment to any group of modern animals, it is now interpreted as lying below anomalocaridids on the stem leading to the living arthropods. This commentary was written to celebrate the 350th anniversary of the journal Philosophical Transactions of the Royal Society. PMID:25750235

  1. Combined Microfacies-Log-Analysis of Cambrian and Ordovician Carbonate Rocks (Upper Cambrian, Western Hills, Bejing; TZ-162 well, Tarim Basin, Western China)

    OpenAIRE

    Wang, Anjia

    2014-01-01

    The Tarim Basin in northwestern China is the second largest petroleum field in China. The reservoirs are predominantly Palaeozoic (Devonian, Ordovician, and Cambrian) carbonate rocks buried at a depth of 5000 m – 6000 m. Ordovician carbonate rocks are one of the main topics in recent hydrocarbon drilling projects. This great depth and a complex inhomogeneous geological and tectonic development results in many problems in hydrocarbon exploration and production. The Ordovician sequence is do...

  2. Extraordinary fossils reveal the nature of Cambrian life: a commentary on Whittington (1975) ‘The enigmatic animal Opabinia regalis, Middle Cambrian, Burgess Shale, British Columbia’

    OpenAIRE

    Briggs, Derek E G

    2015-01-01

    Harry Whittington's 1975 monograph on Opabinia was the first to highlight how some of the Burgess Shale animals differ markedly from those that populate today's oceans. Categorized by Stephen J. Gould as a ‘weird wonder’ (Wonderful life, 1989) Opabinia, together with other unusual Burgess Shale fossils, stimulated ongoing debates about the early evolution of the major animal groups and the nature of the Cambrian explosion. The subsequent discovery of a number of other exceptionally preserved ...

  3. On the evolution and histology of some Cambrian protoconodonts, paraconodonts and primitive euconodonts

    Institute of Scientific and Technical Information of China (English)

    DONG Xiping

    2004-01-01

    Previous reports on the morphological evolution of protoconodonts and paraconodonts are reviewed. The evolutionary trends exhibited by species of the protoconodont genus Gapparodus and the paraconodont genus Westergaadodina are discussed. Based on the present study on histology, genus Paibiconus is protoconodont, while genus Yongshunella is paraconodont (fig. 1 (a)). However, the nature of Huayuanodontus has not been fully understood so far.Recently, the transitional form New Gen. Sensu Miller, 1980 of late Late Cambrian has been found in Hunan Province, South China. It is considered as the Sister-Group of Proconodontus.Moreover, another kind of transitional form of middle Late Cambrian and late Late Cambrian between Prooneotodus rotundatus and Proconodontus, which looks like Prooneotodus rotundatus in gross morphology, but has anterior and posterior costae, has also been found in Hunan,South China. This kind of transitional form is believed to be probably the real intermediate link between Prooneotodus rotundatus and Proconodontus, and Prooneotodus rotundatus is most probably the ancestor of Proconodontus. The two transitional forms noted above are all euconodonts by the study of their histology. Unlike the euconodonts of Post-Cambrian, the evolutionary relationships among the species of protoconodonts, paraconodonts and the primitive euconodonts could not be made clear without the histological study on them.

  4. Gut contents as direct indicators for trophic relationships in the Cambrian marine ecosystem.

    Science.gov (United States)

    Vannier, Jean

    2012-01-01

    Present-day ecosystems host a huge variety of organisms that interact and transfer mass and energy via a cascade of trophic levels. When and how this complex machinery was established remains largely unknown. Although exceptionally preserved biotas clearly show that Early Cambrian animals had already acquired functionalities that enabled them to exploit a wide range of food resources, there is scant direct evidence concerning their diet and exact trophic relationships. Here I describe the gut contents of Ottoia prolifica, an abundant priapulid worm from the middle Cambrian (Stage 5) Burgess Shale biota. I identify the undigested exoskeletal remains of a wide range of small invertebrates that lived at or near the water sediment interface such as hyolithids, brachiopods, different types of arthropods, polychaetes and wiwaxiids. This set of direct fossil evidence allows the first detailed reconstruction of the diet of a 505-million-year-old animal. Ottoia was a dietary generalist and had no strict feeding regime. It fed on both living individuals and decaying organic matter present in its habitat. The feeding behavior of Ottoia was remarkably simple, reduced to the transit of food through an eversible pharynx and a tubular gut with limited physical breakdown and no storage. The recognition of generalist feeding strategies, exemplified by Ottoia, reveals key-aspects of modern-style trophic complexity in the immediate aftermath of the Cambrian explosion. It also shows that the middle Cambrian ecosystem was already too complex to be understood in terms of simple linear dynamics and unique pathways. PMID:23300612

  5. Progress and significance in research on the early Middle Cambrian Kaili Biota, Guizhou Province,China

    Institute of Scientific and Technical Information of China (English)

    2002-01-01

    Important progress in research on the Kaili Biota has been made recently. Many interesting components from Chengjiang Biota and Burgess Shale Biota have been discovered, e.g. Microdictyon of lobopodia; Ottoia, Palaeoscolex of worms; Naraoia, Marrella of Trilobitioidea, Mollisonia, anamalocarids and other non-trilobite arthropods; and new sorts of echinoder-mas, macroalage fossils and so on. Recent work on the Kaili Biota has resulted in the following developments: (i) an increase in the number of animal genera, up to more than 100 genera in total, so that the Kaili Biota has become the third most diverse of the Burgess Shale-type Biota after the Burgess Shale and Chengjiang Biotas; and (ii) the most noteworthy fossils in the Kaili Biota are echinoderms, non-trilobite arthropods and soft-bodied medusiform fossils, especially the most diverse echinoderms. The progress provides envidence for the biodiversity of marine organisms presented after the "Cambrian Explosion" and serves as a link between the earlist Cambrian Chengjiang Biota and late early Middle Cambrian Burgess Shale Biota. It is of great significance in the reconstruction of the Cambrian palaeoplate, palaeongeography and in research on taphonomy.

  6. Gut contents as direct indicators for trophic relationships in the Cambrian marine ecosystem.

    Directory of Open Access Journals (Sweden)

    Jean Vannier

    Full Text Available Present-day ecosystems host a huge variety of organisms that interact and transfer mass and energy via a cascade of trophic levels. When and how this complex machinery was established remains largely unknown. Although exceptionally preserved biotas clearly show that Early Cambrian animals had already acquired functionalities that enabled them to exploit a wide range of food resources, there is scant direct evidence concerning their diet and exact trophic relationships. Here I describe the gut contents of Ottoia prolifica, an abundant priapulid worm from the middle Cambrian (Stage 5 Burgess Shale biota. I identify the undigested exoskeletal remains of a wide range of small invertebrates that lived at or near the water sediment interface such as hyolithids, brachiopods, different types of arthropods, polychaetes and wiwaxiids. This set of direct fossil evidence allows the first detailed reconstruction of the diet of a 505-million-year-old animal. Ottoia was a dietary generalist and had no strict feeding regime. It fed on both living individuals and decaying organic matter present in its habitat. The feeding behavior of Ottoia was remarkably simple, reduced to the transit of food through an eversible pharynx and a tubular gut with limited physical breakdown and no storage. The recognition of generalist feeding strategies, exemplified by Ottoia, reveals key-aspects of modern-style trophic complexity in the immediate aftermath of the Cambrian explosion. It also shows that the middle Cambrian ecosystem was already too complex to be understood in terms of simple linear dynamics and unique pathways.

  7. The distribution of the oil derived from Cambrian source rocks in Lunnan area, the Tarim Basin, China

    Institute of Scientific and Technical Information of China (English)

    2007-01-01

    There are great differences in biomarks between Cambrian oil and Middle-Upper Ordoviclan oil. In this stuty, the authors analyzed 40 oils found in Lunnan area by GC-MS and calculated the content of Cambrian oil in the 40 oils according to the steroid indexes of typical oil mixture and match experiment. The results show that it is a general phenomenon in Ordovician reservoir that the oil derived from Cambrian source rock mixed with the oil derived from Middle-Upper Ordovician source rock in Lunnan area, the mixture degree of the two oils is lower in Carboniferous reservoir than in Ordovician reservoir, and the oils kept in Triassic reservoir have single source, Middle-Upper Ordovician source rock. The mixture oils mainly composed of Cambrian oil (>50%) distributed in Sangtamu fault zone, and the oils found in Lunnan fault zone are Middle-Upper Ordovician oil. This distribution of oils in Lunnan area is owing to that Lunnan fault zone is located in anticline axis part, Lunnan fault zone underwent serious erosion, and the oils from Cambrian source rock accumulated in Lunnan fault zone were degraded completely during Caledonian-Hercynian movement. But the Cambrian oil accumulated in Sangtamu fault zone was not degraded completely and some of them were left for the location of Sangtamu fault zone is lower than Lunnan fault zone. Later, the oil derived from Middle-Upper Ordovician source rock mixed with the remained Cambrian oil, and the mixture oil formed in Sangtamu fault zone.

  8. Geological features of Sinian-Early Cambrian intracratonic rift of the Sichuan Basin

    Directory of Open Access Journals (Sweden)

    Guoqi Wei

    2015-01-01

    Full Text Available With the major breakthrough in Sinian-Cambrian gas exploration in the Sichuan Basin, the geologic understanding on this formation has changed. One change concerns the discovery of a nearly SN negative structure in the west of Gaoshiti-Moxi area, but there is controversy over the nature, boundary features and distribution scope, sedimentary features, formation stage and evolution of the structure [11–14]. Since the negative structure belt is closely related to the formation of the Anyue Sinian-Cambrian giant gas field, it is necessary to get a better understanding of its geological features in order to guide the gas exploration in Sinian-Cambrian of the Sichuan Basin in the future. Through an analysis of the drilling data of Sinian-Cambrian in this area, an observation of outcrop profiles and the use of 27000 km 2D seismic data and latest drilling data (Well Gaoshi 17, and in combination with regional structure background, the boundary and distribution scope, sedimentary features and evolution of the Mianzhu-Changning rift in craton were investigated. The results showed that: ① the rift in overall SN direction had a steep and fairly stable eastern boundary, and gentler western boundary with variable positions in different stages; ② at the end of Sinian Deng2 member, the rift was a nearly symmetrical depression with an area of 3 × 104 km2; ③ at the end of Deng4 member, it was a fault depression which was steep in the east and gentle in the west controlled by its eastern boundary fault with an area of 8 × 104 km2; and ④ the evolution of the rift can be divided into four stages: embryonic stage in Deng1-Deng2, formation stage in Deng3-Deng4, filling and subsidence stage in Early Cambrian Maidingping-Qiongzhusi Formation, shrinkage and disappearing stage in Early Cambrian Canglangpu-Longwangmiao Formation. Through the study, it was concluded that both sides of the rift are in good Sinian-Cambrian gas-reservoir-forming conditions

  9. An Early Cambrian archaeocyath–trilobite fauna in limestone erratics from the Upper Carboniferous Fitzroy Tillite Formation, Falkland Islands

    OpenAIRE

    Stone, P; Thomson, M.R.A; Rushton, A.W.A.

    2012-01-01

    Rare clasts of limestone contained in the uppermost Carboniferous Fitzroy Tillite Formation of the Falkland Islands contain a rich Cambrian fauna of archaeocyaths together with a radiocyath and a few trilobites. Neither Cambrian strata nor limestone are present in the indigenous rock succession and the clasts are regarded as exotic erratics, introduced during the Permo-Carboniferous glaciation of southern Gondwana, prior to its Mesozoic break-up. Nineteen archaeocyath taxa have been identifie...

  10. Geochronological Constraints on Neoproterozoic Glaciations, the first appearance of Metazoans, and the Cambrian Explosion.

    Science.gov (United States)

    Bowring, S.; Condon, D.; Ramezani, J.; Myrow, P.; Landing, E.

    2004-05-01

    Studies of Neoproterozoic climate fluctuations, plate reconstructions, biological evolution and their interrelationships have been hindered by a lack of high-precision geochronological constraints. The correlation and estimates of duration for Neoproterozoic glaciations has relied on physical/chemo-stratigraphy, and thermal subsidence models respectively. New geochronological constraints from Neoproterozoic successions worldwide have sharpened the debate as to the number, synchroneity, and duration of glacial episodes and the relationship, if any, between Metazoan evolution and global glaciation(s). Crucial to the debate are correct interpretation of geochronological data that range from U-Pb zircon studies of intercalated volcanic ash-beds, U-Pb detrital zircon studies, Re-Os from black shales, Rb-Sr from clay-rich rocks, U-Pb and Pb-Pb from carbonates and phosphates, and Lu-Hf from phosphates. Development of a highly resolved Neoproterozoic timescale will require integration and cross-calibration of multiple dating techniques and consideration of what is actually being recorded by each chronometer. A review of available geological and geochronological data indicate that there were at least three and perhaps as many as five periods of Neoproterozoic glacial deposition including rocks from United States (Idaho and Virginia), Newfoundland and the Northwest Territories of Canada, Namibia, and Oman. What must be evaluated is how the paleogeographic distribution of glaciated regions varied with time during the Neoproterozoic. Do Neoproterozoic glacial successions distributed worldwide record a small number of globally synchronous, long-lived glaciations, or numerous diachronous glacial epochs, or a combination of both? At present, the duration of only one glacial deposit, the ca 581 Ma Gaskiers Formation (Newfoundland), is known and it is on the order of 1 Ma, at odds with a long-lived global glaciation predicted by the snowball Earth hypothesis. Other major issues are

  11. Eocrinoid echinoderms from the Lower Cambrian Guanshan Fauna in Wuding, Yunnan, China

    Institute of Scientific and Technical Information of China (English)

    HU ShiXue; LUO HuiLin; HOU ShuGuang; Bernd-Dietrich ERDTMANN

    2007-01-01

    This is a brief report of a new occurrence of eocrinoids from the Early Cambrian Wulongqing Formation in Yunnan, China. The eocrinoids from the Guanshan fauna are among the earliest known eocrinoids. Different from many other Early and Middle Cambrian eocrinoids, the Guanshan eocrinoids are characterized by the absence of sutural pores and epispires, the long and spiral brachioles, the extremely long stalk, and the ratio of the length of the stalk versus that of the calyx. The discovery of the eocrinoids from the Guanshan fauna not only provides new information to the investigation of the early evolution of this animal group, but also shed new light on the occurrence and migration of early eocrinoids.

  12. Lower Cambrian yolk-pyramid embryos from Southern Shaanxi,China

    Institute of Scientific and Technical Information of China (English)

    CHEN Junyuan; Andreas BRAUN; Dieter WALOSZEK; PENG Qingqing; Andreas MAAS

    2004-01-01

    Phosphatized globules with radially arranged pillars were recently recorded from the Lower Cambrian phosphate deposit, Ningqiang, Shaanxi by Yue and Bengtson. These authors interpreted the globules as blastula stage of embryos and the pillars as blastomeres. On the basis of new additional material, we reinterpret these structures as yolk-pyramid stages of possible arthropod eggs. The 20 embryos under present study range from 380 μm to 600 μm in diameter and contain about 120 pyramids. Some embryos having a higher number of pyramids are tentatively interpreted as slightly later developmental stages of the same animal. These 543-million-year-old embryos may push back the evolutionary history of the arthropods to a deeper time and also suggest that one important pattern of arthropod development was already present at the beginning of the Cambrian.

  13. Experimental mineralization of crustacean eggs: new implications for the fossilization of Precambrian-Cambrian embryos

    Science.gov (United States)

    Hippler, D.; Hu, N.; Steiner, M.; Scholtz, G.; Franz, G.

    2012-05-01

    Phosphatized globular microfossils from the Ediacaran and lower Cambrian of South China represent an impressive record of early animal evolution and development. However, their phylogenetic affinity is strongly debated. Understanding key processes and conditions that cause exceptional egg and embryo preservation and fossilization are crucial for a reliable interpretation of their phylogenetic position. We conducted phosphatization experiments on eggs of the marbled crayfish Procambarus that indicate a close link between early mineralization and rapid anaerobic decay of the endochorional envelope. Our experiments replicated the different preservational stages of degradation observed in the fossil record. Stabilization of the spherical morphology was achieved by pre-heating of the eggs. Complete surface mineralization occurred under reduced conditions within one to two weeks, with fine-grained brushite (CaHPO4 · 2H2O) and calcite. The mechanisms of decay, preservation of surface structures, and mineral replacement in the experiment were likely similar during fossilization of Cambrian embryos.

  14. Rise to modern levels of ocean oxygenation coincided with the Cambrian radiation of animals.

    Science.gov (United States)

    Chen, Xi; Ling, Hong-Fei; Vance, Derek; Shields-Zhou, Graham A; Zhu, Maoyan; Poulton, Simon W; Och, Lawrence M; Jiang, Shao-Yong; Li, Da; Cremonese, Lorenzo; Archer, Corey

    2015-01-01

    The early diversification of animals (∼ 630 Ma), and their development into both motile and macroscopic forms (∼ 575-565 Ma), has been linked to stepwise increases in the oxygenation of Earth's surface environment. However, establishing such a linkage between oxygen and evolution for the later Cambrian 'explosion' (540-520 Ma) of new, energy-sapping body plans and behaviours has proved more elusive. Here we present new molybdenum isotope data, which demonstrate that the areal extent of oxygenated bottom waters increased in step with the early Cambrian bioradiation of animals and eukaryotic phytoplankton. Modern-like oxygen levels characterized the ocean at ∼ 521 Ma for the first time in Earth history. This marks the first establishment of a key environmental factor in modern-like ecosystems, where animals benefit from, and also contribute to, the 'homeostasis' of marine redox conditions. PMID:25980960

  15. Cambrian spiral-plated echinoderms from Gondwana reveal the earliest pentaradial body plan.

    Science.gov (United States)

    Smith, Andrew B; Zamora, Samuel

    2013-08-22

    Echinoderms are unique among animal phyla in having a pentaradial body plan, and their fossil record provides critical data on how this novel organization came about by revealing intermediate stages. Here, we report a spiral-plated animal from the early Cambrian of Morocco that is the most primitive pentaradial echinoderm yet discovered. It is intermediate between helicoplacoids (a bizarre group of spiral-bodied echinoderms) and crown-group pentaradiate echinoderms. By filling an important gap, this fossil reveals the common pattern that underpins the body plans of the two major echinoderm clades (pelmatozoans and eleutherozoans), showing that differential growth played an important role in their divergence. It also adds to the striking disparity of novel body plans appearing in the Cambrian explosion. PMID:23804624

  16. A superarmored lobopodian from the Cambrian of China and early disparity in the evolution of Onychophora.

    Science.gov (United States)

    Yang, Jie; Ortega-Hernández, Javier; Gerber, Sylvain; Butterfield, Nicholas J; Hou, Jin-Bo; Lan, Tian; Zhang, Xi-guang

    2015-07-14

    We describe Collinsium ciliosum from the early Cambrian Xiaoshiba Lagerstätte in South China, an armored lobopodian with a remarkable degree of limb differentiation including a pair of antenna-like appendages, six pairs of elongate setiferous limbs for suspension feeding, and nine pairs of clawed annulated legs with an anchoring function. Collinsium belongs to a highly derived clade of lobopodians within stem group Onychophora, distinguished by a substantial dorsal armature of supernumerary and biomineralized spines (Family Luolishaniidae). As demonstrated here, luolishaniids display the highest degree of limb specialization among Paleozoic lobopodians, constitute more than one-third of the overall morphological disparity of stem group Onychophora, and are substantially more disparate than crown group representatives. Despite having higher disparity and appendage complexity than other lobopodians and extant velvet worms, the specialized mode of life embodied by luolishaniids became extinct during the Early Paleozoic. Collinsium and other superarmored lobopodians exploited a unique paleoecological niche during the Cambrian explosion. PMID:26124122

  17. Mineralogy and Chemistry of Coronadite from Middle Cambrian Manganese Deposits at Wadi Dana, Southern Jordan

    OpenAIRE

    Ahmad Al-Malabeh; Tayel El-Hasan

    2009-01-01

    Coronadite was reported from the upper-most horizons of the Middle Cambrian sediments at two locations in Wadi Dana, central Wadi Araba and Jordan. The unit is composed of dolomite, limestone and shale. Geochemical investigations show appreciable variations in Mn, Pb, Fe, K and Ba contents in the coronadite between the two studied sites. Pb was found to increase downward in both sites in spite of the lateral distance between them. Fe does not vary vertically, but its concentration decreases e...

  18. Precambrian-Cambrian Sedimentology, Stratigraphy, and Paleontology in the Great Basin (Western United States)

    OpenAIRE

    Sappenfield, Aaron Dale

    2015-01-01

    Thick accumulations of Neoproterozoic and early Phanerozoic strata are distributed throughout much of the arid continental interior of western North America, providing an expansive and well-exposed archive of this important time in Earth’s history. The information presented herein supplements evaluations regarding the utility and limitations of this archive by providing an integrated sedimentological, paleontological, and geochronological description for Precambrian-Cambrian strata exposed i...

  19. The Middle Cambrian fossil Pikaia and the evolution of chordate swimming

    OpenAIRE

    Lacalli Thurston

    2012-01-01

    Abstract Conway Morris and Caron (2012) have recently published an account of virtually all the available information on Pikaia gracilens, a well-known Cambrian fossil and supposed basal chordate, and propose on this basis some new ideas about Pikaia’s anatomy and evolutionary significance. Chief among its chordate-like features are the putative myomeres, a regular series of vertical bands that extends the length of the body. These differ from the myomeres of living chordates in that boundari...

  20. Cambrian Series 3 carbonate platform of Korea dominated by microbial-sponge reefs

    Science.gov (United States)

    Hong, Jongsun; Lee, Jeong-Hyun; Choh, Suk-Joo; Lee, Dong-Jin

    2016-07-01

    Metazoans have been considered as negligible components of Cambrian Series 3 and Furongian microbial-dominated reefs, in contrast to their presence in earlier Terreneuvian-Cambrian Series 2 microbial-archaeocyath reefs. However, recent discoveries of sponges in Cambrian Series 3-Furongian reefs of Australia, China, Iran, USA, and Korea have raised question regarding their contribution in terms of carbonate platform development, which have never been assessed. This study examines Cambrian Series 3 deposits of the Daegi Formation, Korea to elucidate this question. The 100-m-thick middle part of the Daegi Formation is dominated by boundstone facies, which occupies 45% of the study interval, as well as bioclastic wackestone to packstone, bioclastic grainstone, and ooid packstone to grainstone facies. The Daegi reefs are primarily thrombolitic in composition, with 90% (n = 26/29) of the reefs containing an average of 9% sponges in aerial percentage calculated from thin sections. Lithistid sponges composed of peloidal fabrics, some desma spicules, and spicule networks commonly occupy the interstitial space in microbial clusters, are encrusted by mesoclots and Epiphyton, and are surrounded by micrite. Subordinate non-lithistid demosponges occur within clusters of microbial elements. The middle Daegi Formation can be largely subdivided into shoal environment dominated by grainstone to packstone facies and shallow subtidal platform interior environment located behind shoal with wackestone to packstone facies. The microbial-sponge reefs mainly developed around platform interior as patch reefs. The current study indicates that metazoans in the form of lithistid and non-lithistid demosponges are nearly ubiquitously incorporated in Daegi reefs and contributed greatly to the formation of microbial-sponge reefs as well as carbonate platform during the time. Study of these microbial-sponge reefs and their distribution within the carbonate platform may help us to understand how

  1. Cambrian and Ordovician Fossil-Lagerstätten in the Barrandian area

    Czech Academy of Sciences Publication Activity Database

    Fatka, O.; Budil, P.; Kraft, P.; Mergl, M.; Mikuláš, Radek; Valent, M.; Lajblová, K.; Rak, Š.; Steinová, M.; Szabad, M.; Micka, V.; Aubrechtová, M.; Lajbl, L.; Nohejlová, M.; Vodička, J.

    2012-01-01

    Roč. 18, č. 1 (2012), s. 22-25. ISSN 1212-6209. [Congress of CGS and SGS. Moninec, 2011.09.22-2011.09.25] R&D Projects: GA ČR GA205/06/0395; GA ČR GA205/09/1521 Institutional research plan: CEZ:AV0Z30130516 Keywords : Cambrian * Ordovician * Fossil-Lagerstätten Subject RIV: DB - Geology ; Mineralogy http://www.sci.muni.cz/gap/casop/

  2. Cambrian spiral-plated echinoderms from Gondwana reveal the earliest pentaradial body plan

    OpenAIRE

    Smith, Andrew B.; Zamora, Samuel

    2013-01-01

    Echinoderms are unique among animal phyla in having a pentaradial body plan, and their fossil record provides critical data on how this novel organization came about by revealing intermediate stages. Here, we report a spiral-plated animal from the early Cambrian of Morocco that is the most primitive pentaradial echinoderm yet discovered. It is intermediate between helicoplacoids (a bizarre group of spiral-bodied echinoderms) and crown-group pentaradiate echinoderms. By filling an important ga...

  3. Beyond the Burgess Shale: Cambrian microfossils track the rise and fall of hallucigeniid lobopodians

    OpenAIRE

    Caron, Jean-Bernard; Smith, Martin R; Harvey, Thomas H. P.

    2013-01-01

    Burgess Shale-type deposits are renowned for their exquisite preservation of soft-bodied organisms, representing a range of animal body plans that evolved during the Cambrian ‘explosion’. However, the rarity of these fossil deposits makes it difficult to reconstruct the broader-scale distributions of their constituent organisms. By contrast, microscopic skeletal elements represent an extensive chronicle of early animal evolution—but are difficult to interpret in the absence of corresponding w...

  4. Testing the Cambrian explosion hypothesis by using a molecular dating technique

    OpenAIRE

    Bromham, L.; Rambaut, A; Fortey, R; Cooper, A.; Penny, D

    1998-01-01

    Molecular studies have the potential to shed light on the origin of the animal phyla by providing independent estimates of the divergence times, but have been criticized for failing to account adequately for variation in rate of evolution. A method of dating divergence times from molecular data addresses the criticisms of earlier studies and provides more realistic, but wider, confidence intervals. The data are not compatible with the Cambrian explosion hypothesis as an explanation for the or...

  5. Articulated Wiwaxia from the Cambrian Stage 3 Xiaoshiba Lagerstätte.

    OpenAIRE

    Yang, J.; Smith, M. R.; Lan, T.; Hou, J.-B.; Zhang, X.-G.

    2014-01-01

    Wiwaxia is a bizarre metazoan that has been interpreted as a primitive mollusc and as a polychaete annelid worm. Extensive material from the Burgess Shale provides a detailed picture of its morphology and ontogeny, but the fossil record outside this lagerstätte is scarce, and complete wiwaxiids are particularly rare. Here we report small articulated specimens of Wiwaxia foliosa sp. nov. from the Xiaoshiba fauna (Cambrian Stage 3, Hongjingshao Formation, Kunming, south China). Although spines ...

  6. A superarmored lobopodian from the Cambrian of China and early disparity in the evolution of Onychophora

    OpenAIRE

    Yang, Jie; Ortega-Hernández, Javier; Gerber, Sylvain; Butterfield, Nicholas J.; Hou, Jin-bo; Lan, Tian; Zhang, Xi-guang

    2015-01-01

    Paleozoic lobopodians constitute a diverse assemblage of worm-like organisms that are known from various exceptional fossil deposits and were among the earliest animals to develop skeletonized body parts for protection. Here, we describe Collinsium ciliosum gen. et sp. nov., an armored lobopodian from the early Cambrian Xiaoshiba Lagerstätte (South China). Collinsium belongs to an extinct clade of superarmored lobopodians characterized by supernumerary dorsal spines, and specialized limbs for...

  7. Gut Contents as Direct Indicators for Trophic Relationships in the Cambrian Marine Ecosystem

    OpenAIRE

    Vannier, Jean

    2012-01-01

    Present-day ecosystems host a huge variety of organisms that interact and transfer mass and energy via a cascade of trophic levels. When and how this complex machinery was established remains largely unknown. Although exceptionally preserved biotas clearly show that Early Cambrian animals had already acquired functionalities that enabled them to exploit a wide range of food resources, there is scant direct evidence concerning their diet and exact trophic relationships. Here I describe the gut...

  8. Response to "Discussion on the systematic position of the Early Cambrian priapulomorph worms"

    Institute of Scientific and Technical Information of China (English)

    HAN Jian; HU Shixue

    2006-01-01

    @@ Due to the combination of complex states of preservation and taphonomies in Chengjiang fossils, Huang et al.[1] expressed different opinions to those of Han et al.[2] on the morphology and systematic position of the fossil priapulomorph worms (Priapulomorpha) based on their additional materials. These diverse opinions are critical in any investigation of the diversity of the Introverta in the Cambrian explosion. However, some of their ideas need to be discussed in further detail.

  9. Can fast early rates reconcile molecular dates with the Cambrian explosion?

    OpenAIRE

    Bromham, L D; Hendy, M D

    2000-01-01

    Molecular dates consistently place the divergence of major metazoan lineages in the Precambrian, leading to the suggestion that the 'Cambrian explosion' is an artefact of preservation which left earlier forms unrecorded in the fossil record. While criticisms of molecular analyses for failing to deal with variation in the rate of molecular evolution adequately have been countered by analyses which allow both site-to-site and lineage-specific rate variation, no analysis to date has allowed the ...

  10. Modern optics in exceptionally preserved eyes of Early Cambrian arthropods from Australia

    Science.gov (United States)

    Lee, Michael S. Y.; Jago, James B.; García-Bellido, Diego C.; Edgecombe, Gregory D.; Gehling, James G.; Paterson, John R.

    2011-06-01

    Despite the status of the eye as an ``organ of extreme perfection'', theory suggests that complex eyes can evolve very rapidly. The fossil record has, until now, been inadequate in providing insight into the early evolution of eyes during the initial radiation of many animal groups known as the Cambrian explosion. This is surprising because Cambrian Burgess-Shale-type deposits are replete with exquisitely preserved animals, especially arthropods, that possess eyes. However, with the exception of biomineralized trilobite eyes, virtually nothing is known about the details of their optical design. Here we report exceptionally preserved fossil eyes from the Early Cambrian (~515 million years ago) Emu Bay Shale of South Australia, revealing that some of the earliest arthropods possessed highly advanced compound eyes, each with over 3,000 large ommatidial lenses and a specialized `bright zone'. These are the oldest non-biomineralized eyes known in such detail, with preservation quality exceeding that found in the Burgess Shale and Chengjiang deposits. Non-biomineralized eyes of similar complexity are otherwise unknown until about 85 million years later. The arrangement and size of the lenses indicate that these eyes belonged to an active predator that was capable of seeing in low light. The eyes are more complex than those known from contemporaneous trilobites and are as advanced as those of many living forms. They provide further evidence that the Cambrian explosion involved rapid innovation in fine-scale anatomy as well as gross morphology, and are consistent with the concept that the development of advanced vision helped to drive this great evolutionary event.

  11. Paleobiology of the Early Cambrian Yanjiahe Formation in Hubei Province of South China

    OpenAIRE

    Broce, Jesse

    2013-01-01

    Fossils recovered from limestones of the lower Cambrian (Stage 2-3) Yanjiahe Formation in Hubei Province, South China, recovered using acetic acid maceration, fracturing, and thin sectioning techniques were examined using a combination of analytical techniques, including energy dispersive spectroscopic (EDS) elemental mapping and micro-focus X-ray computed tomography (micro-CT). One important fossil recovered and analyzed with these techniques is a fossilized embryo. Fossilized animal embryos...

  12. Cyclostratigraphy of Late Cambrian cyclic carbonates: an interbasinal field and modelling study, U.S.A.

    OpenAIRE

    Osleger, David Allen

    1995-01-01

    An interbasinal study of Late Cambrian cyclic carbonate successions in the Appalachian and Cordilleran passive margins, the Texas cratonic embayment and the southern Oklahoma aulacogen has provided controls on the simultaneous development of peritidal to subtidal meter-scale cycles and the larger scale depositional sequences on which they are superimposed. Fining-upward peritidal cycles grade seaward into coarsening-upward, shallow to deep subtidal cycles that form a continuum across...

  13. Geosphere-Biosphere Feedbacks as a Trigger for the Cambrian explosion

    Science.gov (United States)

    von Bloh, W.; Bounama, C.; Franck, S.

    2003-12-01

    A new hypothesis for the cause of the Cambrian explosion is presented. The Cambrian explosion 542 Myr ago is characterized by a sudden increase of biomass and a rapid cooling, which amplified the spread of complex multicellular life. The evolution of the planet Earth is described by the co-evolution of the geosphere-biosphere system. A conceptual model for the global carbon cycle of the Earth containing the reservoirs mantle, ocean floor, continental crust, continental biosphere, the kerogen, as well as the aggregated reservoir ocean and atmosphere is presented. In this study the evolution of the mean global surface temperature, the biomass, and reservoir sizes over the whole history and future of the Earth under a maturing Sun is investigated. Here we specify our previously published Earth system model (Franck et al., 2002) by introducing three different types of biosphere: procaryotes, eucaryotes, and complex multicellular life. They are characterized by different temperature tolerance windows and biotic amplification factors of silicate weathering. The biotic enhancement of weathering by complex multicellular life adds an additional feedback to the system and triggers the Cambrian explosion. The Cambrian explosion was mainly driven by extrinsic environmental causes, i.e. a gradual cooling of the Earth. The Cambrian explosion was so rapid because of a positive feedback between the spread of biosphere, increased silicate weathering, and a consequent cooling of the climate. The environment itself has been actively changed by the biosphere maintaining the temperature conditions for its existence. Furthermore, it can be shown that the coupled geosphere-biosphere system exhibit s bistability: there exist two stable model solutions for a certain interval of the biotic enhancement factor, one solution with and one without complex multicellular life. In particular, cooling events in the Neoproterozoic, could force a premature appearance of complex multicellular life

  14. A new U/Pb date for the basal Meishucun section and implications for the age of the Cambrian explosion

    Science.gov (United States)

    Brooks, B. J.; Crowley, J. L.; Bowring, S. A.; Cervato, C.; Jin, Y.

    2006-12-01

    The Early Cambrian of southern China has long been recognized to record the spectacular transition from microscopic small shelly fossils to a fauna characterized by large, diverse higher bilaterians during part of the Early Cambrian evolutionary explosion. Understanding the timing and rate of this evolutionary transition has been aided through the integration of U/Pb geochronology into Cambrian fossil biozones, which has established strong tie-points between platforms. The Meishucun section (Yunnan, China) is one of the units that best preserves what could be considered the prelude to the Cambrian explosion. Several attempts at dating volcanic horizons within this section have resulted in U/Pb zircon dates that range from 538 to 525 Ma, making correlations to sections further afield problematic. This new high-precision U/Pb date of 533 Ma for Meishucun's Bed 5 is in broad agreement but considerably more precise than previous U/Pb ages. This indicates that the low diversity fauna of the Anabarites trisulcatus--Protohertzina anabarica zone persisted for no less than the first 9 m.y. of the Cambrian in China, and sets an additional lower bracket age for the emergence of higher bilaterians (e.g. trilobites, soft-bodied fauna).

  15. A Cambrian micro-lobopodian and the evolution of arthropod locomotion and reproduction

    Institute of Scientific and Technical Information of China (English)

    2007-01-01

    The evolutionary success of arthropods, the most abundant and diverse animal group, is mainly based on their segmented body and jointed appendages, features that had evolved most likely already before the Cambrian. The first arthropod-like animals, the lobopodians from the Early Cambrian, were unsclerotized and worm-like, and they had unjointed tubular legs. Here we describe the first three-dimensionally preserved Cambrian lobopodian. The material presented of Orstenotubulus evamuellerae gen. et sp. nov. is the smallest and youngest of a lobopodian known. O. evamuellerae shows strikingly detailed similarities to Recent tardigrades and/or onychophorans in its cellular-structured cuticle and the telescopic spines. It also shows similarities to other, longer known lobopodians, but which are ten times as large as the new form. These similarities include the finely annulated body and legs, which is characteristic also for Recent onychophorans, and paired humps continuing into spines situated dorsally to the leg insertions, a feature lacking in the extant forms. The morphology of O. evamuellerae not only elucidates our knowledge about lobopodians, but also aids in a clearer picture of the early evolution of arthropods. An example is the single ventral gonopore between a limb pair of O. evamuellerae, which indicates that a single gonopore, as developed in onychophorans, tardigrades, pentastomids, myriapods and insects, might represent the plesiomorphic state for Arthropoda, while the paired state in chelicerates and crustaceans was convergently achieved. Concerning life habits, the lateral orientation of the limbs and their anchoring spines of the new lobopodian imply that early arthropods were crawlers rather than walkers.

  16. Molecular Fossils for Understanding Biodiversity During the Neoproterozoic-Cambrian Transition in China

    Science.gov (United States)

    Pi, Y.; Tuo, J.; McFadden, K.; Xiao, S.; Zhang, C. L.

    2005-12-01

    Neoproterozoic-Cambrian rocks in South China contain an extraordinary fossil record, including exceptionally well preserved animal embryos, acritarchs, and multi-cellular algae. The goal of this study was to evaluate the microbial diversity associated with these remarkably preserved fossil assemblages at the Neoproterozoic-Cambrian transition. Rock samples of 520-632 Ma in age were collected in the Yangtze Gorges area and southern Anhui Province, China. Samples were powdered and extracted for organic biomarkers. The content of bitumen A accounted for 4-16% of the rock material and most of it (49-79%) was asphaltenes. Saturated and aromatic hydrocarbons accounted for 2-6% and 1-3%, respectively. Analysis using GC-MS indicated the predominance of n-alkanes and less abundant isoprenoid alkanes in the saturated fractions. The n-alkanes were characterized by homologues dominated by C15-C17, which is consistent with the result of high thermal-evolution (Brocks et al., 2003. GCA 67:4321-4335). Hopanoids were present in less abundance and ranged from C29 to C32. A smaller amount of heavy-molecular-weight n-alkanes (C23-C39) was also detected, which indicated a source of high plants and must be contamination from younger organic matter. Still, patterns of variation can be detected among these samples. For example, the ratio of pristane to phytane was all greater than 1.0 except for one sample (JLW9.3) from Yangtze Gorges area. The results indicate that sample JLW9.3 might have been deposited in a reducing environment whereas the other samples might have been formed in relatively oxidative environments. The overall results, however, suggest that rock samples from the Neoproterozoic-Cambrian transition in China have gone through significant metamorphism; thus, understanding of microbial communities using molecular biomarkers in such altered rocks needs to be cautiously executed.

  17. The Urbilateria: Insights From Comparative Developmental Biology Into The Cambrian Radiation

    Science.gov (United States)

    Erwin, D. H.

    2004-12-01

    A critical issue in understanding the patterns of morphological and developmental evolution during the Cambrian radiation of animal body plans is the morphological complexity of the last common ancestor of the two great bilaterian clades, the protostomes and deuterostomes. Often known as the Urbilateria or the protostome-deuterostome ancestor (PDA) this node in metazoan phylogeny distinguishes the sponges and cnidarians from the bilaterian clades. Highly conserved developmental genes and complexes suggests that the PDA was a relatively complex organism, with dorsal-ventral and anterior-posterior patterning, eyes, appendages, heart, central nervous system, segmentation and other relatively complex morphological features. If this view of the PDA is correct, it has important implications for understanding the Cambrian radiation, for such a complex animal is likely to leave traces in the sedimentary record even if it lacked a durable skeleton. However, this interpretation of conserved developmental sequences may overestimate the extent to which homologies of developmental sequences implies morphological conservation. In other words, does the presence of the Nkx 2.5 gene in the PDA necessitate the presence of a heart, or simple the appearance of contractile muscles? If the latter, then the PDA may have been a much less complex organism. This latter view of developmental evolution suggests the PDA possessed a developmental toolkit, but that the specific implementation of these tools in specific body plans occurred after the PDA, within specific clades. Moreover, this view also strongly suggests that changes in the physical environment, or in ecological relationships, was primarily responsible for the evolutionary innovations seen in the Cambrian, rather than developmental innovations.

  18. Preliminary basin analysis of late Proterozoic-Cambrian post-rift strata, southeast Idaho thrust belt

    Energy Technology Data Exchange (ETDEWEB)

    Link, P.K.; Jansen, S.T.; Halimdihardja, P.; Lande, A.C.; Zahn, P.D.

    1987-08-01

    Strata of the Brigham Group in the Paris-Putnam plate of the southeastern Idaho thrust belt span the late Proterozoic-Cambrian boundary and consist of quartzose sandstone with subordinate pebble conglomerate and siltstone. The Brigham Group is overlain by fossiliferous Cambrian carbonate units that represent the transition from siliciclastic to carbonate deposition in the Cordilleran miogeocline. The Brigham Group contains four stratigraphic sequences bounded by regional disconformities. The lower sequence includes strata below the Brigham group (upper member, Pocatello Formation), plus the Papoose Creek Formation and most of the overlying Caddy Canyon Quartzite. This sequence is dominantly marine with shoreface and braided fluvial strata at the top. The first sequence is overlain disconformably by offshore sub-wave base marine strata of the upper Caddy Canyon Quartzite and Inkom Formation. This second sequence is entirely marine and is composed dominantly of siltstone with sandstone-filled channels. The third sequence comprises the Mutual Formation, an entirely braided fluvial and lacustrine unit. The fourth sequence (Sauk sequence) locally overlies the Mutual Formation with an erosional unconformity and consists of dominantly marine strata of the Camelback Mountain Quartzite, Gibson Jack Formation, Windy Pass Argillite, Twin Knobs Formation, and Sedgwick peak Quartzite. Correlations of these sequences to the McCoy Creek Group of eastern Nevada suggests uniform conditions of sea level and subsidence across the late Proterozoic-Cambrian Cordilleran miogeocline. This preliminary synthesis suggests the Brigham and McCoy Creek Groups are post-rift deposits, as indicated by regional persistence of facies, paleocurrents, and quartzose petrology.

  19. The Lower Cambrian of Scandinavia: Depositional environment, sequence stratigraphy and palaeogeography

    Science.gov (United States)

    Nielsen, Arne Thorshøj; Schovsbo, Niels Hemmingsen

    2011-08-01

    Lower Cambrian successions described from Scandinavia are reviewed and subjected to sequence stratigraphical analysis; comparisons are also made with successions described from northeast Poland, Estonia, Latvia and Lithuania. The treated stratigraphic interval is bounded upwards by a regional unconformity ascribed to the Hawke Bay Event. The East European regional stage classification, comprising the Rovnian, Lontovan, Dominopolian, Ljubomlian, Vergalian, Rausvian and Kibartian, is adopted for the Lower Cambrian of Scandinavia. These units are approximately equivalent to the Terreneuvian and Cambrian provisional series 2. The Rovnian and Lontovan stages are pre-trilobitic. The Dominopolian and 'Ljubomlian' stages encompass the ' Rusophycus' and Schmidtiellus mickwitzi zones; whether the former zone is of pre-trilobitic age is uncertain but possible. The 'Ljubomlian' is treated informally because the definition adopted in this paper does not correspond to the original concept of the stage. The Vergalian and Rausvian are for the time being classified as one combined stage. The lower main part of the Vergalian-Rausvian corresponds to the new informal Holmia kjerulfi- 'Ornamentaspis' linnarssoni zone, whereas the upper part is separated as the new informal Comluella?-Ellipsocephalus lunatus zone. This zone also includes the Kibartian Stage. Volborthella and poorly known olenellid trilobites range into the Kibartian and the stage is considered of Early Cambrian age. The Holmia inusitata Zone is abandoned; it is contemporaneous with the traditional ' O.' linnarssoni Zone. The autochthonous strata underlying the Hawke Bay unconformity in the Laisvall sector, Swedish Lapland, are assigned to the Laisberg and Grammajukku formations and it is proposed to abandon the Laisvall and Såvvare formations. The Laisberg Fm can locally be divided into the Ackerselet, Saivatj, Maiva, Kautsky Ore, Tjalek, Nadok Ore and Assjatj members. The Vakkejokk Breccia near Luopakte is likely

  20. Vermicular fossils in the Early Cambrian Xidashan Formation in the Quruqtagh region of Xinjiang, China

    Institute of Scientific and Technical Information of China (English)

    YANG Ruidong; ZHANG Chuanlin; SONG Guoqi; LUO Xinrong

    2006-01-01

    Large amounts of vermicular fossils and a minor amount of sponge animal fossils have been found in the Early Cambrian Xidashan Formation in the region of Quruqtagh, Xinjiang. Vermicular fossils are generally more than 50 mm long and 0.3-1.4 mm wide; their wrinkled lamellae are microfine with 3-10 pieces within the length of every each millimeter. The fossils are considered to be Sabellidites cambriensis Sokolov (1965). The Xidashan Formation is the highest stratum in which Sabellidites occur, as has been so far reported.

  1. Organic carbon isotopes of the Sinian and Early Cambrian black shales on Yangtze Platform, China

    Institute of Scientific and Technical Information of China (English)

    李任伟; 卢家烂; 张淑坤; 雷加锦

    1999-01-01

    Organic matter of the Sinian and early Cambrian black shales on the Yangtze Platform belongs to the light carbon group of isotopes with the δ13C values from - 27 % to -35 % , which are lower than those of the contemporaneously deposited carbonates and phosphorites. A carbon isotope-stratified paleooceanographic model caused by upwelling is proposed, which can be used not only to interpret the characteristics of organic carbon isotopic compositions of the black shales, but also to interpret the paleogeographic difference in the organic carbon isotope compositions of various types of sedimentary rocks.

  2. Early Cambrian phosphatized blastula-and gastrula-stage animal fossils from southern Shaanxi

    Institute of Scientific and Technical Information of China (English)

    HUA Hong; CHEN Zhe; ZHANG Luyi

    2004-01-01

    Marine invertebrate animal embryos and their early developmental products are of great significance to the study of taxonomy and phylogeny of early animals. A great number of phosphatized globular fossils were collected from the early Cambrian Kuanchuanpu Member (upper Dengying Formation), southern Shaanxi, and a nearly complete developmental sequence--from a fertilized egg, via blastodisc formation, blastula development, blastodisc enlargement toward gastrulae, to tissue differentiation--can be discerned in this collection. This discovery provides unmatchable material for studies on the origin, taxonomy, radiation,and ontogeny of early metazoans.

  3. Origin and variability of the late Precambrian-Cambrian Athel Silicilyte, South Oman Salt Basin

    OpenAIRE

    Al Rajaibi, Ibrahim Mohamed amur

    2011-01-01

    The Precambrian-Cambrian Athel Silicilyte is an enigmatic chert unit of up to 390 m thick found as slabs (each slab typically 2 × 6 km across) entrapped within salt domes at a depth of 4-5 km in the South Oman Salt Basin. This formation is a prolific self-charged reservoir with high porosity (up to 34 %) and high oil saturation (80 %). Despite its economic value, the origin and the variability of this formation are not fully understood. This study therefore aims to investigate the variability...

  4. Rise to modern levels of ocean oxygenation coincided with the Cambrian radiation of animals.

    OpenAIRE

    Chen, X.; Ling, HF; Vance, D.; Shields-Zhou, GA; Zhu, M.; Poulton, SW; Och, LM; Jiang, SY; Li, D.; Cremonese, L; Archer, C.

    2015-01-01

    The early diversification of animals (∼630 Ma), and their development into both motile and macroscopic forms (∼575–565 Ma), has been linked to stepwise increases in the oxygenation of Earth's surface environment. However, establishing such a linkage between oxygen and evolution for the later Cambrian ‘explosion' (540–520 Ma) of new, energy-sapping body plans and behaviours has proved more elusive. Here we present new molybdenum isotope data, which demonstrate that the areal extent of oxygenat...

  5. Cambrian Evolutionary Radiation: Context, correlation, and chronostratigraphy—Overcoming deficiencies of the first appearance datum (FAD) concept

    Science.gov (United States)

    Landing, Ed; Geyer, Gerd; Brasier, Martin D.; Bowring, Samuel A.

    2013-08-01

    Use of the first appearance datum (FAD) of a fossil to define a global chronostratigraphic unit's base can lead to intractable correlation and stability problems. FADs are diachronous—they reflect species' evolutionary history, dispersal, biofacies, preservation, collection, and taxonomy. The Cambrian Evolutionary Radiation is characterised by diachronous FADs, biofacies controls, and provincialism of taxa and ecological communities that confound a stable Lower Cambrian chronostratigraphy. Cambrian series and stage definitions require greater attention to assemblage zone successions and non-biostratigraphic, particularly carbon isotope, correlation techniques such as those that define the Ediacaran System base. A redefined, basal Cambrian Trichophycus pedum Assemblage Zone lies above the highest Ediacaran-type biotas (vendobionts, putative metazoans, and calcareous problematica such as Cloudina) and the basal Asteridium tornatum-Comasphaeridium velvetum Zone (acritarchs). This definition and the likely close correspondence of evolutionary origin and local FAD of T. pedum preserves the Fortune Head, Newfoundland, GSSP of the Cambrian base and allows the presence of sub-Cambrian, branched ichnofossils. The sub-Tommotian-equivalent base of Stage 2 (a suggested "Laolinian Stage") should be defined by the I'/L4/ZHUCE δ13C positive peak, bracketed by the lower ranges of Watsonella crosbyi and Aldanella attleborensis (molluscs) and the Skiagia ornata-Fimbrioglomerella membranacea Zone (acritarchs). The W. crosbyi and A. attleborensis FADs cannot define a Stage 2 base as they are diachronous even in the Newfoundland "type" W. crosbyi Zone. The Series 2 base cannot be based on a species' FAD owing to the provincialism of skeletalised metazoans in the Terreneuvian-Series 2 boundary interval and global heterochrony of the oldest trilobites. A Series 2 and Stage 3 (a suggested "Lenaldanian Series" and "Zhurinskyan Stage," new) GSSP base is proposed at the Siberian lower

  6. Isotopic compositions of small shelly fossil Anabarites from Lower Cambrian in Yangtze Platform of South China: Implications for palaeocean temperature

    Institute of Scientific and Technical Information of China (English)

    2007-01-01

    Anabarites belong to small shelly fossils (SSF), which occur widely in the Lower Cambrian strata of Yangtze Platform in South China. They are phosphate shell in composition and represent the earliest stage of the Cambrian bioradiation of Bilateria, the socalled "Cambrian Explosion". In this study, we attempted to separate Anabarites fossils from Lower Cambrian dolostones, and we obtained samples of both the fossils (SSF) and the granular phosphates (GP). Isotopic analyses were performed on samples of SSF, GP, and matrix dolostone (DH-23). The results showed that the δ30 Si values of the quartz filling in fossils celoms, and the siliceous materials in granular phosphates are -0.6‰ and -0.7‰, which is different from normal sedimentary siliceous rocks from the Lower Cambrian strata (0-0.7‰) as reported by Li et al., but is consistent with the data for siliceous rocks and cherts of submarine hydrothermal origin. It is likely that a later hydrothermal replacement could have taken place in the SSF-bearing sedimentary rocks. The oxygen isotope values of the phosphate of SSF and GP are 16.8‰ and 17.0‰, respectively. These are significantly higher than the Neoproterozoic phosphate ores (10.9‰-13.9‰) as reported by Ling et al., hence, late diagenesis and hydrothermal replacement may not have caused a significant change in the oxygen isotope compositions of the small shelly fossils, and the calculated temperatures (25.4-26.3 ℃ ) for palaeo-seawater using a SSF phosphate oxygen isotope thermometer are therefore considered here as the upper limit of seawater temperature in the Early Cambrian ocean of the Yangtze Platform.

  7. Arachnostega Bertling, 1992 in the Drumian (Cambrian) sediments of the Teplá-Barrandian region (Czech Republic)

    Czech Academy of Sciences Publication Activity Database

    Fatka, O.; Mikuláš, Radek; Szabad, M.; Micka, V.; Valent, M.

    2011-01-01

    Roč. 61, č. 4 (2011), s. 367-381. ISSN 0001-5709 R&D Projects: GA ČR GA205/06/0395; GA ČR GA205/09/1521 Institutional research plan: CEZ:AV0Z30130516 Keywords : Cambrian * Teplá-Barrandian region * ichnofossils * Jince Formation * Buchava Formation * Arachnostega * Cambrian (Czech Republic) Subject RIV: DB - Geology ; Mineralogy Impact factor: 0.565, year: 2011 https://geojournals.pgi.gov.pl/agp/article/view/9705

  8. Early colonization of metazoans in the deep-water: Evidences from the lowermost Cambrian black shales of South China

    Science.gov (United States)

    Zhu, M.-Y.; Yang, A.-H.; Zhang, J.-M.; Li, G.-X.; Yang, X.-L.

    2003-04-01

    Diversity of metazoans is high in the deep-water of the present ocean. But it is unknown that when the metazoans began to colonize in the deep-water and what kinds of metazoans first colonized in the deep-water since origin and radiation of metazoans during the Precambrian-Cambrian transition interval. Up to the present, colonization of the deep-sea began in the Ordovician. Although it is suggested that animals were penetrated into the intermediate water depth during the Precambrian, evidences support such suggestion are based on the problematic Ediacaran-grade fossils. However, almost fossil materials that support the Cambrian Explosion hypothesis were discovered from the lowermost Cambrian shallow-water deposits. The abundant earliest Cambrian mineralized small shelly fossils (SSF) are globally from the shallow-water deposits, and the well-known Chengjiang fauna that may records most complete features of metazoans in the ocean after the Cambrian Explosion, occurs as well in the shallow basin near an old land on the Yangtze Platform. In order to understand ecology of the Cambrian Explosion time interval and how happened of the onshore-offshore trends of metazoans, we focused our attention on collecting fossils in the lowermost Cambrian deposits under the varied facies on the Yangtze Platformm during recent years. Investigations of the shallow-water carbonate facies and the oxygen-depleted deep-water black shale facies revealed additional biological and ecological information that are not recorded in the Chengjiang fauna in the siliclastic shallow-water facies. Here we report our discovery of a particular fossil association from more than 10 sections in the deep-water black shales (Qiongzhusian) in the out shelf and slope area of the Yangtze Platform. The fossil association is composed of pelagic and sessile organisms, including abundant sponges, 3 types of bivalved arthropods, 3 types of tubular animals and few problematic organisms. The fossils have either

  9. Sinian-Cambrian stratigraphic framework for shallow- to deep-water environments of the Yangtze Platform: an integrated approach

    Institute of Scientific and Technical Information of China (English)

    ZHU Maoyan; ZHANG Junming; Michael STEINER; YANG Aihua; LI Guoxiang; Bernd D. ERDTMANN

    2003-01-01

    The Sinian (Terminal Proterozoic) and Early Cambrian shallow- to deep-water sequences of the Yangtze Platform were investigated. Based on integrated lithostratigraphic, biostratigraphic, and other approaches, the shallow-water sequence from the base of the Sinian (base of the Doushantuo Formation) to the top of the Qiongzhusian (top of the Yu'anshan Formation) was subdivided into 12 stratigraphic intervals. These 12 intervals were applied in turn to the subdivision and correlation of the sequences present in various facies of the Yangtze Platform. The high-resolution stratigraphic framework here developed can serve as a time frame for ongoing multidisciplinary analyses of the "Cambrian explosion".

  10. Experimental mineralization of crustacean eggs: new implications for the fossilization of Precambrian–Cambrian embryos

    Directory of Open Access Journals (Sweden)

    D. Hippler

    2012-05-01

    Full Text Available Phosphatized globular microfossils from the Ediacaran and lower Cambrian of South China represent an impressive record of early animal evolution and development. However, their phylogenetic affinity is strongly debated. Understanding key processes and conditions that cause exceptional egg and embryo preservation and fossilization are crucial for a reliable interpretation of their phylogenetic position. We conducted phosphatization experiments on eggs of the marbled crayfish Procambarus that indicate a close link between early mineralization and rapid anaerobic decay of the endochorional envelope. Our experiments replicated the different preservational stages of degradation observed in the fossil record. Stabilization of the spherical morphology was achieved by pre-heating of the eggs. Complete surface mineralization occurred under reduced conditions within one to two weeks, with fine-grained brushite (CaHPO4 · 2H2O and calcite. The mechanisms of decay, preservation of surface structures, and mineral replacement in the experiment were likely similar during fossilization of Cambrian embryos.

  11. Brachiopods hitching a ride: an early case of commensalism in the middle Cambrian Burgess Shale

    Science.gov (United States)

    Topper, Timothy P.; Holmer, Lars E.; Caron, Jean-Bernard

    2014-10-01

    Ecological interactions, including symbiotic associations such as mutualism, parasitism and commensalism are crucial factors in generating evolutionary novelties and strategies. Direct examples of species interactions in the fossil record generally involve organisms attached to sessile organisms in an epibiont or macroboring relationship. Here we provide support for an intimate ecological association between a calcareous brachiopod (Nisusia) and the stem group mollusc Wiwaxia from the Burgess Shale. Brachiopod specimens are fixed to Wiwaxia scleritomes, the latter showing no signs of decay and disarticulation, suggesting a live association. We interpret this association as the oldest unambiguous example of a facultative ectosymbiosis between a sessile organism and a mobile benthic animal in the fossil record. The potential evolutionary advantage of this association is discussed, brachiopods benefiting from ease of attachment, increased food supply, avoidance of turbid benthic conditions, biofoul and possible protection from predators, suggesting commensalism (benefiting the symbiont with no impact for the host). While Cambrian brachiopods are relatively common epibionts, in particular on sponges, the association of Nisusia with the motile Wiwaxia is rare for a brachiopod species, fossil or living, and suggests that symbiotic associations were already well established and diversified by the ``middle'' (Series 3, Stage 5) Cambrian.

  12. Mantle insulation beneath the West African craton during the Precambrian-Cambrian transition

    Science.gov (United States)

    Doblas, Miguel; López-Ruiz, José; Cebriá, José-María; Youbi, Nasrrddine; Degroote, Eugenio

    2002-09-01

    At the time of the Precambrian-Cambrian transition, the West African craton underwent widespread magmatism, hydrothermal activity, and thermal rejuvenation. This tectonothermal event gave rise to an anorogenic “ring of fire” along the rim of this craton, following the Pan-African Brasiliano belt that was reactivated by extension and transtension. The thermal phenomena were due to the progressive peripheral release of mantle heat that had built up beneath this craton because of strong insulating conditions. The West African craton at the Precambrian-Cambrian transition can thus be envisioned in terms of a gigantic pressure-cooker with a thick blanketing lithospheric lid. These insulation processes triggered an unusually hot mantle that was channeled by edge-driven convection toward the peri West African craton extensional corridors and released through magmatic pressure-relief valves. Massive ice melting and outgassing of volcanic CO2 gave rise to a planet-scale sea-level rise, a greenhouse effect, and the end of the icehouse snowball Earth. These processes played an important role in the Phanerozoic explosion of life on Earth.

  13. The evolution of associative learning: A factor in the Cambrian explosion.

    Science.gov (United States)

    Ginsburg, Simona; Jablonka, Eva

    2010-09-01

    The Cambrian explosion is probably the most spectacular diversification in evolutionary history, and understanding it has been a challenge for biologists since the time of Darwin. We propose that one of the key factors that drove this great diversification was associative learning. Although the evolutionary emergence of associative learning required only small modifications in already existing memory mechanisms and may have occurred in parallel in several groups, once this type of learning appeared on the evolutionary scene, it led to extreme diversifying selection at the ecological level: it enabled animals to exploit new niches, promoted new types of relations and arms races, and led to adaptive responses that became fixed through genetic accommodation processes. This learning-based diversification was accompanied by neurohormonal stress, which led to an ongoing destabilization and re-patterning of the epigenome, which, in turn, enabled further morphological, physiological, and behavioral diversification. Our hypothesis combines several previous ideas about the dynamics of the Cambrian explosion and provides a unifying framework that includes both ecological and genomic factors. We conclude by suggesting research directions that would clarify the timing and manner in which associative learning evolved, and the effects it had on the evolution of nervous systems, genomes, and animal morphology. PMID:20558182

  14. Preliminary estimate of crustal extension during Cambrian rifting in the southern midcontinent

    Energy Technology Data Exchange (ETDEWEB)

    McConnell, D.A.; Gilbert, M.

    1985-01-01

    The Southern Oklahoma Aulacogen is a prominent rift extending 500+ km NW into the southern Midcontinent from the probably Cambrian plate margin. The biomodal igneous floor of the rift has been strongly structurally inverted and is now partially exposed in the core of the Wichita Mountains. Utilizing structural and stratigraphic patterns, and character of the igneous sequence, the style of tectonism and magnitude of displacement can be inferred for the initiation of the rift. Five possible extensional pulses can be related to specific igneous features although these could be part of one continuous episode lasting from about 565 to 525 mybp. Pennsylvanian transpressive faults are assumed to be reactivated Cambrian normal faults (and they may even have an earlier parentage). Using known thickness of continuous rhyolite (1.4 km), an initial width of 60 km, and a half-graben configuration, an estimate of extension is possible independent of bounding fault dip. For a brittle-ductile transition between 10 to 15 km, the brittle extension in the upper crust varies between 8-5%. Using the minimum mafic volume needed within the ductile lower crust (40,000 km/sup 3/), and a total crustal thickness of 35 to 45 km, the lower crust was extended 8-6% to 10-7%. The upper and lower crustal estimates of extension are in good agreement confirming a relatively shallow brittle-ductile transition. This is consistent with concomitant igneous activity and an enhanced geothermal gradient.

  15. The paleomagnetism of the salt pseudomorph beds of middle cambrian age from the salt range, West Pakistan

    NARCIS (Netherlands)

    Wensink, H.

    1972-01-01

    Oriented cores for a paleomagnetic investigation were collected from ten sites in the sedimentary redbeds of the Salt Pseudomorph Beds of Middle Cambrian age m the Salt Range near Khcwra. All samples were subjected to progressive, thermal demagnetization procedures which revealed the characteristic

  16. The paleomagnetism of the Salt Pseudomorph Beds of Middle Cambrian age from the Salt Range, West Pakistan

    NARCIS (Netherlands)

    Wensink, H.

    1972-01-01

    Oriented cores for a paleomagnetic investigation were collected from ten sites in the sedimentary redbeds of the Salt Pseudomorph Beds of Middle Cambrian age in the Salt Range near Khewra. All samples were subjected to progressive, thermal demagnetization procedures which revealed the characteristic

  17. Estimation of porosity of Khewra sandstone of cambrian age by using helium porosimeter and its application in reservoir evaluation

    International Nuclear Information System (INIS)

    Estimation of petrophysical properties of the rock formations played decisive role in all the processes of petroleum exploration. The Cambrian sequence is well established as reservoir rocks in the various parts of the world from where petroleum is being tapped. The Cambrian sequence has been encountered in the Potwar area and limited petroleum is being produced from the Adhi Oil Field. The Khewra Sandstone of the Cambrian sequence is outcropped in the Khewra Gorge, Salt Range Pakistan below an unconformity qf the Tobra Formation. For the assessment of porosity and reservoir characterization Helium Porosimeter has been used, six samples Qf the upper horizon were collected from various locations of the Khewra Gorge and the Khewra Choha Sadden Shah road side section; cores were prepared from these samples according to the instrument standard. The results of this study revealed that the upper horizon Qf the Khewra Sandstone Formation has good porosity ranging from 18.76% to 21.07%, porosity varied in different parts of the formation. These results are in good agreement with the internationally reported values for petroleum reservoir of the Cambrian sandstone. (author)

  18. The structure and sedimentary sequence of intracratonic rift from Late Sinian to Early Cambrian in the Sichuan Basin, South China

    Science.gov (United States)

    Gu, Zhidong; Zhang, Baomin; Lu, Weihu; Zhai, Xiufen; Jiang, Hua

    2016-04-01

    Sichuan Basin is located in the northwest of Upper Yangtze craton of South China, and there is developed an intracratonic rift from Late Sinian to Early Cambrian in the middle of Sichuan Basin, and the paper systematically discusses the structure and sedimentary sequence of the intracratonic rift based on the fields, drilling and seismic data, and so on. Detailed structural interpretation of 2D and 3D seismic profiles displays the development of two stages of intracratonic rift due to regional extension with the depth of 2000m, and plane distribution of intracratonic rift presents the V-pattern from the northwest to the southeast in the middle of Sichuan Basin with the width from 100km to 20km. The drilling data from the intracratonic rift shows the obvious thinning of Upper Sinian and thickening of Lower Cambrian. And field outcrops situated in the intracratonic rift reveal that the Upper Sinian is mainly composed of siliceous rock, shale and carbonate, with the thickness of less than 100m, but the thickness of Upper Sinian on the platform reaches 1000m by contrast; They also reveals that Lower Cambrian is mainly composed of shale, mudstone, and siltstone with the development of gravity current, and the thickness of Lower Cambrian reaches 2000m. The formation of intracratonic rift may be initiated by pre-existing basement weakness zone and deep mantle dynamics.

  19. Glauconitic deposits at Julegård on the south coast of Bornholm, Denmark dated to the Cambrian

    DEFF Research Database (Denmark)

    Clemmensen, Lars B; Bromley, Richard Granville; Holm, Paul Martin

    2011-01-01

    the deposits belong to the Lower Cambrian Norretorp Member of the Læså Formation. The shallow marine deposits are strongly bioturbated, but only a single ichnoassociation is represented. The ichnogenus is referable to either Trichophycus Miller and Dyer, 1878 or Teichichnus Seilacher, 1955. Rare...

  20. Paleoecologic Implications of Ichnofossils Associated with Slightly Skeletonized Body Fossils, Middle Cambrian of the Barrandian Area, Czech Republic

    Czech Academy of Sciences Publication Activity Database

    Mikuláš, Radek; Fatka, O.; Szabad, M.

    2012-01-01

    Roč. 19, č. 4 (2012), s. 199-210. ISSN 1042-0940 R&D Projects: GA ČR GA205/09/1521 Institutional support: RVO:67985831 Keywords : Middle Cambrian * ichnofossils * slightly skeletonized fauna * Barrandian area Subject RIV: DB - Geology ; Mineralogy Impact factor: 0.727, year: 2012

  1. Decoupled evolution of soft and hard substrate communities during the Cambrian Explosion and Great Ordovician Biodiversification Event.

    Science.gov (United States)

    Buatois, Luis A; Mángano, Maria G; Olea, Ricardo A; Wilson, Mark A

    2016-06-21

    Contrasts between the Cambrian Explosion (CE) and the Great Ordovician Biodiversification Event (GOBE) have long been recognized. Whereas the vast majority of body plans were established as a result of the CE, taxonomic increases during the GOBE were manifested at lower taxonomic levels. Assessing changes of ichnodiversity and ichnodisparity as a result of these two evolutionary events may shed light on the dynamics of both radiations. The early Cambrian (series 1 and 2) displayed a dramatic increase in ichnodiversity and ichnodisparity in softground communities. In contrast to this evolutionary explosion in bioturbation structures, only a few Cambrian bioerosion structures are known. After the middle to late Cambrian diversity plateau, ichnodiversity in softground communities shows a continuous increase during the Ordovician in both shallow- and deep-marine environments. This Ordovician increase in bioturbation diversity was not paralleled by an equally significant increase in ichnodisparity as it was during the CE. However, hard substrate communities were significantly different during the GOBE, with an increase in ichnodiversity and ichnodisparity. Innovations in macrobioerosion clearly lagged behind animal-substrate interactions in unconsolidated sediment. The underlying causes of this evolutionary decoupling are unclear but may have involved three interrelated factors: (i) a Middle to Late Ordovician increase in available hard substrates for bioerosion, (ii) increased predation, and (iii) higher energetic requirements for bioerosion compared with bioturbation. PMID:27247396

  2. Decoupling of body-plan diversification and ecological structuring during the Ediacaran-Cambrian transition: evolutionary and geobiological feedbacks.

    Science.gov (United States)

    Mángano, M Gabriela; Buatois, Luis A

    2014-04-01

    The rapid appearance of bilaterian clades at the beginning of the Phanerozoic is one of the most intriguing topics in macroevolution. However, the complex feedbacks between diversification and ecological interactions are still poorly understood. Here, we show that a systematic and comprehensive analysis of the trace-fossil record of the Ediacaran-Cambrian transition indicates that body-plan diversification and ecological structuring were decoupled. The appearance of a wide repertoire of behavioural strategies and body plans occurred by the Fortunian. However, a major shift in benthic ecological structure, recording the establishment of a suspension-feeder infauna, increased complexity of the trophic web, and coupling of benthos and plankton took place during Cambrian Stage 2. Both phases were accompanied by different styles of ecosystem engineering, but only the second one resulted in the establishment of the Phanerozoic-style ecology. In turn, the suspension-feeding infauna may have been the ecological drivers of a further diversification of deposit-feeding strategies by Cambrian Stage 3, favouring an ecological spillover scenario. Trace-fossil information strongly supports the Cambrian explosion, but allows for a short time of phylogenetic fuse during the terminal Ediacaran-Fortunian. PMID:24523279

  3. Results of high temperature processing of high-carbon materials from the lower Cambrian period of the Earth's history

    Science.gov (United States)

    Maslov, O. D.

    2016-07-01

    The paper reports on the observation of spontaneous fission of nuclides, concentrated in fly ash during the combustion of high-carbon (graphite) material, chemogenic siliceous-carbonaceous rocks and carbonaceous shale in the mixture with brown coal. In the samples obtained, the spontaneous fission was measured by track method. The zones of precipitation of spontaneous fission of nuclides and their lighter homologues on thermochromatographic column were determined. A nuclide with a half-life of 62 days was detected in the alkaline trap. The chemical treatment procedure included co-precipitation with iron hydroxide, dissolution in NH4OH + H2O2 solution and distillation by heating up to 100°C followed by AgI co-precipitation. Based on the chemical behavior it can be concluded that the detected radionuclide belongs to the halides. The content of the parent nuclide in high-carbon (graphite) material and chemogenic siliceous-carbonaceous rock corresponds to 10-14 g/g.

  4. The First Paleomagnetic data from the Cambrian basalts of Henrietta Island (De Long Archipelago, Arctic Ocean)

    Science.gov (United States)

    Metelkin, D. V.; Zhdanova, A.; Vernikovskiy, V. A.; Matushkin, N. Y.

    2015-12-01

    Henrietta Island in De Long archipelago (East-Siberian sea) still remains poorly studied geologically but last investigations show that its volcano-sedimentary sequences can help reconstruct tectonic evolution of East Russian Arctic in Early Paleozoic stage. The deposits lying on Precambrian basements are deformed to varying degrees and intruded by mafic dykes.The study was carried out on two basaltic lava flows whose 40Ar/39Ar age is 520.6±9.5 Ma. Previously the age of these basalts was assumed Cretaceous. According to available data the underlaying sediments contain zircons with Cambrian and Ordovician ages but all boundaries between these basalts and other strata are tectonic. So we suppose the age of basalts as Middle Cambrian but more precise geochronological data are required. All magnetic measurements were performed at the Laboratory of Geodynamics and Paleomagnetism of Institute of Petroleum Geology and Geophysics (Novosibirsk). Basalt samples has relatively high magnetic susceptibility values varying from 5x10-4 to 180x10-4SI units. NRM values range is from 3 to 170 mA/m. Petromagnetic parameters including also coercive characteristics point at the good potentially preserving primary magnetization. Stepwise thermal demagnetization permits to isolate characteristic components of magnetization and calculate mean directions in two lava flows: 1. Ds=294.3°, Is=29.1°, K=81.1, α95=5.1; 2. Ds=301.0°, Is=28.3°, K=34.4, α95=7.9). The mean paleomagnetic pole has coordinates: Plat=20.9°, Plong = 42.6°, dp/dm=14.3/7.9. Paleolatitude was defined as 15.3° but the question of the hemisphere for De Long Islands is open yet. In case of south hemisphere in Middle Cambrian according to available paleomagnetic data De Long islands could be placed close to Taimyr margin of Siberia and in case of northern hemisphere they may be located near south (in present-day coordinates) margin of Siberia. The work was supported by grant RFBR 14-05-31399 and Russian Research Fund

  5. A highly redox-heterogeneous ocean in South China during the early Cambrian (˜529-514 Ma): Implications for biota-environment co-evolution

    Science.gov (United States)

    Jin, Chengsheng; Li, Chao; Algeo, Thomas J.; Planavsky, Noah J.; Cui, Hao; Yang, Xinglian; Zhao, Yuanlong; Zhang, Xingliang; Xie, Shucheng

    2016-05-01

    The "Cambrian Explosion" is known for rapid increases in the morphological disparity and taxonomic diversity of metazoans. It has been widely proposed that this biological event was a consequence of oxygenation of the global ocean, but this hypothesis is still under debate. Here, we present high-resolution Fe-S-C-Al-trace element geochemical records from the Jinsha (outer shelf) and Weng'an (outer shelf) sections of the early Cambrian Yangtze Platform, integrating these results with previously published data from six correlative sections representing a range of water depths (Xiaotan, Shatan, Dingtai, Yangjiaping, Songtao, and Longbizui). The integrated iron chemistry and redox-sensitive trace element data suggest that euxinic mid-depth waters dynamically coexisted with oxic surface waters and ferruginous deep waters during the earliest Cambrian, but that stepwise expansion of oxic waters commenced during Cambrian Stage 3 (∼ 521- 514 Ma). Combined with data from lower Cambrian sections elsewhere, including Oman, Iran and Canada, we infer that the global ocean exhibited a high degree of redox heterogeneity during the early Cambrian, consistent with low atmospheric oxygen levels (∼ 10- 40% of present atmospheric level, or PAL). A large spatial gradient in pyrite sulfur isotopic compositions (δ34Spy), which vary from a mean of - 12.0 ‰ in nearshore areas to + 22.5 ‰ in distal deepwater sections in lower Cambrian marine units of South China imply low concentrations and spatial heterogeneity of seawater sulfate, which is consistent with a limited oceanic sulfate reservoir globally. By comparing our reconstructed redox chemistry with fossil records from the lower Cambrian of South China, we infer that a stepwise oxygenation of shelf and slope environments occurred concurrently with a gradual increase in ecosystem complexity. However, deep waters remained anoxic and ferruginous even as macrozooplankton and suspension-feeding mesozooplankton appeared during

  6. The anatomy, taphonomy, taxonomy and systematic affinity of Markuelia: Early Cambrian to Early Ordovician scalidophorans

    Science.gov (United States)

    Dong, X.-P.; Bengtson, S.; Gostling, N.J.; Cunningham, J.A.; Harvey, T.H.P.; Kouchinsky, A.; Val'Kov, A.K.; Repetski, J.E.; Stampanoni, M.; Marone, F.; Donoghue, P.C.J.

    2010-01-01

    Markuelia is a vermiform, annulated introvertan animal known as embryonic fossils from the Lower Cambrian to Lower Ordovician. Analysis of an expanded and revised dataset for Introverta shows that the precise position of Markuelia within this clade is dependent on the taxa included. As a result, Markuelia is assigned to the scalidophoran total group to reflect uncertainty as to whether it is a stem-scalidophoran or a stem-priapulid. The taxonomy of the genus is revised to provide an improved taxonomic framework for material assigned to Markuelia. Five species are recognized: M. secunda Val'kov, M. hunanensis Dong and Donoghue, M. lauriei Haug et al., M. spinulifera sp. nov. and M. waloszeki sp. nov. Finally, the preservation of Markuelia is evaluated in the light of both the taphonomy of the fossil embryos themselves and the experimental taphonomy of the priapulid Priapulus caudatus, which has been proposed as both a close relative and an anatomical analogue of Markuelia. ?? The Palaeontological Association.

  7. Ediacarian sponge spicule clusters from southwestern Mongolia and the origins of the Cambrian fauna

    Science.gov (United States)

    Brasier, Martin; Green, Owen; Shields, Graham

    1997-04-01

    Carbon and strontium isotopic data are used to show that the earliest sponge spicule clusters and associated phosphatic sediments (with Anabarites) from southwestern Mongolia are of Ediacarian age. Spicule morphologies include bundles of oxeas arranged in three-dimensional quadrules, linked together at junctions by tetracts, pentacts, hexacts, or polyactines. All are referred to the Phylum Porifera, Class Hexactinellida. These sponge spicules provide the oldest remains that can be assigned without question to an extant phylum, and also the first firm evidence for filter feeding and metazoan silica biomineralization in the fossil record. It is suggested that siliceous and phosphatic members of the “Cambrian fauna” may have had their origins in eutrophic and outer shelf facies of the Late Proterozoic.

  8. Cephalic and appendage morphology of the Cambrian arthropod Sidneyia inexpectans Walcott, 1911

    DEFF Research Database (Denmark)

    Stein, Martin

    2013-01-01

    evidence for variability of head segment counts in Cambrian arthropods, and to falsify the hypothesis of a head with three postantennular segments in the euarthropod ground pattern. Restudy of a substantial amount of material of S. inexpectans shows that previous interpretations of a short head were based...... understood, but the exopod seemed to differ from that of other artiopodans, except for the shared presence of lamellae. The head was considered to comprise only the ocular and antennular segments, these being covered entirely on the ventral side by a large doublure. This short head was often taken as an...... at least two postantennular appendage bearing segments. The appendage morphology is shown to be consistent with artiopodan affinities. The exopod is of the bilobate flap-like type with lamellae inserting on the proximal portion, earlier proposed as a potential autapomorphy of Artiopoda. Reinforcement...

  9. Preliminary notes on soft-bodied fossil concentrations from the Early Cambrian Chengjiang deposits

    Institute of Scientific and Technical Information of China (English)

    HAN Jian; SHU Degan; ZHANG Zhifei; LIU Jianni; ZHANG Xingliang; YAO Yang

    2006-01-01

    The efforts of labor-intensive collecting in the Early Cambrian Chengjiang deposits in eastern Yunnan Province, China led to the discovery of many horizons containing exceptionally well preserved soft-bodied fossil concentrations, many of which can be assigned to either monospecific concentrations or paucispecific concentrations. The features of these fossil concentrations support the hypothesis that frequent storm events producing tempestites mainly contributed to the preservation of abundant softbodied fossils in the Chengjiang deposits, and indicate that the balance of the ecological web in this region was probably frequently destroyed or upset by such storm events during that geological time. Animals in a fossil assemblage in such fossil concentrations probably occupied similar ecological biotopes.

  10. Petrographic and geochemical composition of kerogen in the Furongian (U. Cambrian) Alum Shale, central Sweden

    DEFF Research Database (Denmark)

    Sanei, H.; Petersen, H. I.; Schovsbo, N. H.; Jiang, C.; Goodsite, M. E.

    2014-01-01

    degradation of labile kerogen by radiation from uranium-rich minerals. The higher degree of aromaticity in Alum shale suggests lower than expected oil-proneness. The results of this study suggest that the Alum Shale is a gas-prone source producing aromatic hydrocarbon mixtures with an unexpectedly low......This paper presents an integrated geochemical and petrological study of the Hällekis-1 core from the Furongian (upper Cambrian) Alum shales in central Sweden to characterise organic matter composition, depositional environment, and potential hydrocarbon generation capability. The results show that...... organic-rich Alum Shale (TOC: 8.9-28.0. wt.%) contains mainly immature, predominantly algal-derived kerogen with unusually reduced hydrocarbon generation potential as suggested by relatively low Hydrogen Index (HI) values (HI: 251-471. mg HC/g TOC) and high degree of aromaticity. In the absence of thermal...

  11. Ferroan dolomite cement in Cambrian sandstones: burial history and hydrocarbon generation of the Baltic sedimentary basin

    DEFF Research Database (Denmark)

    Sliaupa, S.; Cyziene, J.; Molenaar, Nicolaas;

    2008-01-01

    The conditions and timing of carbonate cementation in Cambrian sandstones of the Baltic sedimentary basin were determined by oxygen and carbon stable isotope and chemical data in combination with optical and cathodoluminescence petrographic studies. Studied samples represent a range in present...... burial depth from 340 to 2150 m. The carbonate cement is dominantly ferroan dolomite that occurs as dispersed patches of poikilotopic crystals. Temperatures of dolomite precipitation, based on delta O-18 values, range from 27 degrees C in the shallow buried to 95 degrees C in the deep buried sandstones....... The burial history modelling points to development of most of the dolomite cement during rapid Silurian-Devonian subsidence and Carboniferous-early Permian uplift. A wide range of precipitation temperatures indicate that temperature was not a major factor in triggering the carbonate cementation. Dolomite...

  12. Extensive Cambrian braidplain sedimentation: Insights from the southwestern Cordillera, U. S. A

    Energy Technology Data Exchange (ETDEWEB)

    Fedo, C.M. (Virginia Polytechnic Inst. and State Univ., Blacksburg (United States)); Prave, A.R. (City College of New York, NY (United States))

    1991-02-01

    The Lower to Middle Cambrian time-transgressive coarse terrigenous clastic rocks that form the basal part of the Cordilleran cratonal (Tapeats Sandstone) to miogeoclinal (Wood Canyon Formation, Zabriskie Quartzite) succession have long been though of as products of marine sediment dispersion processes. The authors' recent facies analyses and reinterpretation suggest, instead, that those strata record an extensive continental braidplain which interacted with marine environments. Marine deposits are distinguished by the presence of trace fossils and abundant wave- and tide-influenced sedimentary structures. The most abundant braidplain facies are those assigned a fluvial origin. Coarser, more poorly sorted deposits are feldspathic, whereas finer, better sorted deposits are quartzitic. Beds typically are wedge- and channel-shaped and their width ranges inversely to grain size; channels in conglomeratic layers are relatively narrow (< 20 m wide), whereas those in sand-dominated intervals are shallow (< 1 m) and wide (tens of meters) with broadly concave-up bases. Trough cross-strata and flat laminations are common with planar cross-strata less common. Basal lags and cyclic arrangement of sedimentary structures are rare, although thinning- and fining-upward intervals are developed locally. Paleocurrent indicators across the entire region are west directed and of low variance. They suggest that the dominant mode of sediment transport was by down-channel migrating dunes and that subordinate processes included sheet flow (flat laminated sandstones) and eolian deflation (textural bimodality). These consistent paleocurrent directions attest to the remarkable longevity (Early through Middle Cambrian) of an extensive, westward-draining fluvial system.

  13. Evidence for microbial dissolution of pyrite from the Lower Cambrian oolitic limestone, South China

    Directory of Open Access Journals (Sweden)

    W. Liu

    2011-03-01

    Full Text Available The oxidative dissolution of the sulphide mineral pyrite (FeS2 has been of significant interest since it affects global geochemical cycles, generates acid mine drainage, and is used in industrial metal extraction. Several different groups of prokaryotes are known to catalyze the dissolution of pyrite and use the free energy generated from the oxidation, which may result in the dissolution of the mineral and the precipitation of the secondary ferric iron minerals either on the cell surface or is separated from the cells. However, straightforward evidence for such metabolic process in the ancient sediments is rare. Here we report pyrite crystals from the Lower Cambrian oolitic limestones that show indications of microbial erosion in various degrees. Erosion pits and tubular micro-tunnels with characteristic shapes and sizes in our samples are generally similar to those obtained from the laboratory studies on the oxidative dissolution of pyrite by iron-oxidizing bacteria. Diagenetic examination demonstrates that the bioerosion predates the consolidation of the limestone. In addition, bacillus-sized and -shaped microfossils encrusted with iron oxides are present in our samples, which are very likely to be fossilized sheaths produced by iron-oxidizing bacteria. Our findings indicate that the microbial oxidative dissolution of pyrite existed in the Cambrian shallow marine carbonate sediments. Furthermore, we suggest that characteristic pitting patterns on the pyrite crystals from ancient sediments are an important clue to trace the evolution of life, in particular, the evolution of metabolism like microbial iron oxidation in the remote past on our planet, independent of biomarkers, isotopic signals and body fossils as well.

  14. Origin of northern Gondwana Cambrian sandstone revealed by detrital zircon SHRIMP dating

    Science.gov (United States)

    Avigad, D.; Kolodner, K.; McWilliams, M.; Persing, H.; Weissbrod, T.

    2003-01-01

    Voluminous Paleozoic sandstone sequences were deposited in northern Africa and Arabia following an extended Neoproterozoic orogenic cycle that culminated in the assembly of Gondwana. We measured sensitive high-resolution ion microprobe (SHRIMP) U-Pb ages of detrital zircons separated from several Cambrian units in the Elat area of southern Israel in order to unravel their provenance. This sandstone forms the base of the widespread siliciclastic section now exposed on the periphery of the Arabian-Nubian shield in northeastern Africa and Arabia. Most of the detrital zircons we analyzed yielded Neoproterozoic concordant ages with a marked concentration at 0.55–0.65 Ga. The most likely provenance of the Neoproterozoic detritus is the Arabian-Nubian shield; 0.55–0.65 Ga was a time of posttectonic igneous activity, rift-related volcanism, and strike-slip faulting there. Of the zircons, 30% yielded pre-Neoproterozoic ages grouped at 0.9–1.1 Ga (Kibaran), 1.65–1.85 Ga, and 2.45–2.7 Ga. The majority of the pre-Neoproterozoic zircons underwent Pb loss, possibly as a consequence of the Pan-African orogeny resetting their provenance. Rocks of the Saharan metacraton and the southern Afif terrane in Saudi Arabia (∼1000 km south of Elat) are plausible sources of these zircons. Kibaran basement rocks are currently exposed more than 3000 km south of Elat (flanking the Mozambique belt), but the shape of the detrital zircons of that age and the presence of feldspar in the host sandstone are not fully consistent with such a long-distance transport. Reworking of Neoproteorozoic glacial detritus may explain the presence of Kibaran detrital zircons in the Cambrian of Elat, but the possibility that the Arabian-Nubian shield contains Kibaran rocks (hitherto not recognized) should also be explored.

  15. Detrital zircon geochronology of Neoproterozoic to Middle Cambrian miogeoclinal and platformal strata: Northwest Sonora, Mexico

    Science.gov (United States)

    Gross, E.L.; Stewart, John H.; Gehreis, G.E.

    2000-01-01

    Eighty-five detrital zircon grains from Mesoproterozoic and/or Neoproterozoic to Middle Cambrian sedimentary strata in northwest Sonora, Mexico, have been analyzed to determine source terranes and provide limiting depositional ages of the units. The zircon suites from the Mesoproterozoic and/or Neoproterozoic El Alamo Formation and El Aguila unit yield ages between 1.06 Ga and 2.67 Ga, with predominant ages of 1.1 to 1.2 Ga. Zircons from the Lower? and Middle Cambrian Bolsa Quartzite show age groups from 525 Ma to 1.63 Ga, with a dominant population of 1.1 to 1.2 Ga grains. Grains older than 1.2 Ga in the samples were most likely derived from basement terranes and ???1.4 Ga granitic bodies of the southwest U.S. and northwest Mexico. It is also possible that the sediments were transported from the south, although source rocks of the appropriate age are not presently exposed south of the study area in northern Mexico. Three possibilities for the dominant 1.1 to 1.2 Ga grains include derivation from: (I) exposures of the Grenville belt in southern North America, (2) local 1.1-1.2 Ga granite bodies, or (3) a southern source, such as the Oaxaca terrane, that was subsequently rifted away. Sampling of additional units in the western U.S. and northern Mexico may help resolve the ambiguity surrounding the source of the 1.1 to 1.2 Ga grains.

  16. The Meers Fault in Southern Oklahoma: Holocene Movements on a Fault with Pennsylvanian and Cambrian Linages

    Science.gov (United States)

    Keller, G. R.; Holland, A. A.; Luza, K.; Oldow, J. S.; Crain, K.

    2011-12-01

    The Meers fault and subparallel fault strands in southern Oklahoma is the southernmost element of the complex and massive (>10 km of throw) frontal fault zone that forms the boundary between the Anadarko basin, which is the deepest intra-continental basin in the United States, and the uplifted Cambrian igneous rocks of the Wichita Mountains. The Wichita uplift is evidence of extraordinary Pennsylvanian intra-plate deformation along the trend of the Southern Oklahoma aulacogen, which is a classic example of a failed and massively inverted rift. The Meers Fault is the best-documented Holocene fault scarp east of Colorado and probably represents reactivation of a Pennsylvanian oblique thrust that in turn is likely to be an inverted Cambrian normal fault. The magnitude of these structures is shown on images from 3-D industry seismic reflection data ~25 km northwest of the northwestern mapped extent of the Meers fault that indicate the Pennsylvanian structure, or a northern strand of it, has a reverse throw of ~6km at depth. The fault displays a conspicuous and continuous scarp that is at least 25 km long and is evident in air photos and 1:100,000 scale geologic mapping, but this feature is not well mapped in detail beyond the area of trenching studies conducted in the 1980's. In the Holocene, 3-5 m of vertical surface displacement has been documented and left-lateral strike slip displacement on the fault is 2-3 times greater than the vertical displacement. During this movement, Quaternary soils along the fault were folded and ruptured, and the scarp has dammed small gullies where fine-grained alluvium has collected and has been used in the dating efforts. The most recent movement occurred (1100-1300 y ago) with a variety of earlier events having been proposed. As such, this fault represents one of the highest potential seismic hazards in the central/eastern United States.

  17. Geodatabase of the datasets used to represent the four aquifer subunits of the Cambrian-Ordovician aquifer system, Illinois, Iowa, Michigan, Minnesota, and Wisconsin

    Data.gov (United States)

    U.S. Geological Survey, Department of the Interior — This geodatabase includes spatial datasets that represent the Cambrian-Ordovician aquifer system in the States of Wisconsin, Michigan, Minnesota, Iowa, and...

  18. "The Chengjiang Fauna and the Cambrian Explosion" Sponsord by NSFC won the first-class award of National Natural Sciences Award

    Institute of Scientific and Technical Information of China (English)

    Liu Y.; Yao Y.P.

    2004-01-01

    @@ "The Chengjiang Fauna and the Cambrian explosion'' sponsord by NSFC won the first-class award of National Natural Sciences Award. The Awarding ceremony was held in the Great Hall of the People in Beijing on February 20, 2004. "The Chengjiang Fauna and the Cambrian explosion" was sponsord by NSFC and carried out by several groups led by Prof. Chen Junyuan of the Nanjing Institute of Geology and Palaeontology,CAGS, Prof. Hou Xianguang of Yunnan University and Prof.

  19. Periodic functions with variable period

    OpenAIRE

    Pryjmak, M. V

    2010-01-01

    The examples of rhythmical signals with variable period are considered. The definition of periodic function with the variable period is given as a model of such signals. The examples of such functions are given and their variable periods are written in the explicit form. The system of trigonometric functions with the variable period is considered and its orthogonality is proved. The generalized system of trigonometric functions with the variable period is also suggested; some conditions of it...

  20. Le volcanisme cambrien du Maroc central : implications géodynamiquesThe Central Morocco Cambrian volcanism: geodynamic implications

    Science.gov (United States)

    Ouali, Houssa; Briand, Bernard; Bouchardon, Jean-Luc; Capiez, Paul

    2003-05-01

    In southeastern Central Morocco, the Bou-Acila volcanic complex is considered of Cambrian age. In spite of low-grade metamorphic effect, initial volcanic texture and mineralogy can be recognized and volcanic rocks are dominated by dolerites and porphyric dolerites. The initial mineralogy is composed of plagioclases, pyroxenes and dark minerals. A secondary mineral assemblage is composed of albite, epidote, chlorite and calcite. According to their immobile elements compositions, the southeastern central Morocco metavolcanites are of within-plate continental tholeiites. This volcanism and those recognized in many other areas in Morocco confirm a Cambrian extensive episode within the Gondwana supercontinent. To cite this article: H. Ouali et al., C. R. Geoscience 335 (2003).To cite this article: H. Ouali et al., C. R. Geoscience 335 (2003).

  1. Paleomagnetism of the Cambrian Sediments in Bornholm (Denmark) and Southern Sweden and Implications for Paleogeography of Baltica

    Science.gov (United States)

    Lewandowski, M.; Abrahamsen, N.

    2001-12-01

    In Bornholm the Phanerozoic sedimentary succession overlying Precambrian basement starts with the Lower Cambrian (ca. 545 Ma) Nekso Sandstone Fm (NSF) that, in the earlier paleomagnetic analyses, have yielded characteristic magnetization (ChRM) similar to the Permian direction for Baltica. Here, we present paleomagnetic results from other Lower Cambrian formations in Bornholm (Balka Sandstone, Broens Odde siltstone) and in southern Sweden (Hardebega, Mickwitzia and Lingula Sandstones). Lithologically, most of these formations are very similar to the NSF and they are all considered stratigraphically younger. The intention was to check, whether a Permian paleomagnetic overprint occured on a regional scale and could it be attributed to secondary magnetic phases originating from fluids, which possibly traveled laterally along the contact between the basement and the Lower Cambrian sediments. In this report we show results of this regional test, followed by a comprehensive re-analysis of our Nekso data, including new results obtained after supplementary sampling and new petrologic information, which appeared recently. We have found that the NSF possessed a unique ChRM. The well-grouped and stable characteristic magnetization of the Nekso Sandstone contrasts with fairly chaotic, soft and badly preserved magnetization of the Balka, the Hardeberga, the Mickwitzia and the Lingulid sandstones of Bornholm and southern Sweden. A regional geological context, including the inferred diagenetic evolution of Lower Cambrian sediments, points to a syndepositional/early diagenetic origin of the characteristic remanence of the Nekso Sandstone, revealing a stable remanence applicable for plate tectonic interpretations. Similarity of the characteristic remanence of the NSF to the Permian direction for Baltica has been confirmed, but it is supposed to be casual, because of lack of any sign of a regional Permian remagnetization within the other Cambrian deposits of Southern Scandinavia

  2. A New Arthropod, Guangweicaris Luo, Fu et Hu gen. nov.from the Early Cambrian Guanshan Fauna, Kunming, China

    Institute of Scientific and Technical Information of China (English)

    LUO Huilin; FU Xiaoping; HU Shixue; LI Yong; HOU Shuguang; YOU Ting; PANG Jiyuan; LIU Qi

    2007-01-01

    The Guanshan Fauna is a soft-bodied fauna dominated by arthropods (including trilobites,trilobitoides, Tuzoia, Isoxys, and bradorids) in association with priapulids, brachiopods,anomalocaridids, vetulicoliids, sponges, chancellorids, and echinoderms. This paper reports and describes a new arthropod from the yellowish green mudstone at the lower part of the Wulongqing Formation, Canglangpuan Stage, Lower Cambrian in Kunming, Yunnan, China. The stratigraphic and geographic distribution, classification, fossil preservation, life style of this new arthropod and comparisons with other fossil arthropods are also discussed in details. The discovery and research of the non-mineralized arthropod, Guangweicaris Luo, Fu et Hu gen. nov. from the Guanshan Fauna adds new members to the taxonomic list and provides new information to the evolution of early arthropods.Furthermore, this study would shed new light into the "Cambrian Explosion" and the evolution of early life.

  3. The demosponge Leptomitus cf. L. lineatus, first occurence from the Middle Cambrian of Spain (Murero Formation, Western Iberian Chain)

    OpenAIRE

    García-Bellido, Diego

    2003-01-01

    The Middle Cambrian Murero Formation in the Iberian Chain (NE Spain) has yielded a small collection of softbodied fossils (palaeoscolecid worms, onychophorans, algae), a host of fossils from organisms with mineralized skeletons such as trilobites and brachiopods, and some trace fossils. This paper deals with the description of the only specimen of the demosponge Leptomitus known so far from Spain and its association with the brachiopod Micromitra, showing a probable case of commensalism by th...

  4. Decoupling of body-plan diversification and ecological structuring during the Ediacaran–Cambrian transition: evolutionary and geobiological feedbacks

    OpenAIRE

    Mángano, M. Gabriela; Luis A. Buatois

    2014-01-01

    The rapid appearance of bilaterian clades at the beginning of the Phanerozoic is one of the most intriguing topics in macroevolution. However, the complex feedbacks between diversification and ecological interactions are still poorly understood. Here, we show that a systematic and comprehensive analysis of the trace-fossil record of the Ediacaran–Cambrian transition indicates that body-plan diversification and ecological structuring were decoupled. The appearance of a wide repertoire of behav...

  5. On gamma-ray bursts and their biological effects :a case for an extrinsic trigger of the Cambrian explosion ?

    CERN Document Server

    Horváth, J E

    2003-01-01

    We discuss some the effects of local gamma-ray bursts on the earth's atmosphere. A rough calculation of the fraction of ozone destruction by catalytic $NO_{x}$ cycles is given, which in turn serves to argue how the large flux of gammas from these events would have indirectly provoked major extinction of living organisms. We give specific examples of these features, and tentatively identify the Cambrian explosion seen in the actual fossil record as an event caused by a GRB.

  6. A new vetulicolian from Australia and its bearing on the chordate affinities of an enigmatic Cambrian group

    OpenAIRE

    García-Bellido, Diego C.; Lee, Michael S. Y.; Gregory D. Edgecombe; Jago, James B; Gehling, James G; Paterson, John R

    2014-01-01

    Background Vetulicolians are one of the most problematic and controversial Cambrian fossil groups, having been considered as arthropods, chordates, kinorhynchs, or their own phylum. Mounting evidence suggests that vetulicolians are deuterostomes, but affinities to crown-group phyla are unresolved. Results A new vetulicolian from the Emu Bay Shale Konservat-Lagerstätte, South Australia, Nesonektris aldridgei gen. et sp. nov., preserves an axial, rod-like structure in the posterior body region ...

  7. The Sirius Passet Lagerstatte : silica death masking opens the window on the earliest matground community of the Cambrian explosion.

    OpenAIRE

    Strang, K.M.; Armstrong, H.A.; Harper, D.A.T.; Trabucho-Alexandre, João P.

    2016-01-01

    The Sirius Passet Lagerstätte (SP), Peary Land, North Greenland, occurs in black slates deposited at or just below storm wave base. It represents the earliest Cambrian microbial mat community with exceptional preservation, predating the Burgess Shale by 10 million years. Trilobites from the SP are preserved as complete, three-dimensional, concave hyporelief external moulds and convex epirelief casts. External moulds are shown to consist of a thin veneer of authigenic silica. The casts are for...

  8. Molecular fossils and sources of Cambrian heavy oil of Well Tadong-2 in theTarim Basin, Xinjiang, China

    Institute of Scientific and Technical Information of China (English)

    TANG Youjun; ZAN Ling

    2009-01-01

    Research on the molecular fossil characteristics of heavy oil from Well Tadong-2 is of great importance to constrain the source of marine crude oils in the Tarim Basin, Xinjiang, China. The authors synthetically applied the isotope mass spectrograph, chromatography and chromatography-mass spectrography to the studies of molecular fossil characteristics of heavy oil from Well Tadong-2 in the Tarim Basin, and the results obtained revealed that heavy oil from Well Tadong-2 is characterized by high gammacerane, high C28 sterane, low rearranged sterane and high C27-triaromatic steroid, these characteristics are similar to those of Cambrian-Lower Ordovician source rocks, demonstrating that Cambrian crude oils came from Cambrian-Lower Ordovician source rocks; condensed compounds (fluoranthene, pyrene, benzo[a]anthracene, bow, benzo fluoranthene, benzopyrene) of high abundance were detected in heavy oil from Well Tadong-2, and the carbon isotopic values of whole oil are evidently heavy, all the above characteristics revealed that hydrocarbons in the crude oils became densified in response to thermal alteration.

  9. Sm-Nd isotopic compositions of Cambrian-Ordovician strata at the Jinggangshan area in Jiangxi Province:Tectonic implications

    Institute of Scientific and Technical Information of China (English)

    SHEN WeiZhou; LING HongFei; SHU LiangShu; ZHANG FangRong; XIANG Lei

    2009-01-01

    Whole-rock Sm-Nd Isotopes were investigated for Cambrian-Ordovician strata from the Jinggangshan area beTween the Yangtze Block and Cathysla Block in South China. These strata were deposited as a greatly thick unit of muddy-sandy laminas with intercalated carbonate and organic carbon-bearing layers. They have low ε(t) values of -13.9 to -7.9 and old Nd model ages of 1842 to 2375 Ma. In t-tdiagram, they are far away from the concordant line but fall within the evolution zone of the Proterozoic crust of South China. This indicates that the Cambrian-Ordovician strata are mainly composed of mat-ters eroded from ancient Paleoproterozoic crust that may mainly consist of continental-derived detrital sediments with high maturity in the Cathyaia Block. However, the Ordovician Jueshangou Formation and Dui'erehi Formation have ε(t) values of -10.5 and -7.9 at the higher end of the above range and Nd model ages of 1842 to 2059 Ma at the lower end of the above range. This suggests involvement of more detritus that were eroded from the relatively juvenile crust from Late Paleoproterozoic to the Early Neoproterozoic. All the Nd model ages for the Cambrian-Ordovlcian sedimentary rocks in the Cathysia Block and the southeastern margin of the Yangtze Block are older than 1800 Ma, suggesting that no material from the Early Paleozoic depleted mantle-derived magmas was involved in these regions.

  10. Ichnofossils and their significance in the Cambrian successions of the Parahio Valley in the Spiti Basin, Tethys Himalaya, India

    Science.gov (United States)

    Parcha, S. K.; Pandey, Shivani

    2011-11-01

    The Spiti Basin exposes well preserved Cambrian successions in the Tethys Himalaya. The present ichnofossil assemblage is reported from the Debsakhad Member of the Kunzum La Formation. The ichnofossils includes the ichnogenera Bergaueria, Chondrites, Cruziana, Didymaulichnus, Dimorphichnus, Diplichnites, Helminthorhaphe, Merostomichnites, ?Monocraterion, Monomorphichnus, Nereites, Palaeopascichnus, Palaeophycus, Phycodes, Planolites, Rusophycus, Skolithos, Scolicia, Treptichnus, etc. along with annelid worm, burrow and scratch marks. These ichnogenera can be assigned to cubichnial, repichnial, pascichnial to fodinichnial behaviors. The ichnofossils reported from this section provide evidence regarding the developmental patterns during the early phase of life. In absence of trilobites, the present assemblage of ichnofossils is very significant in assigning the age of the Debsakhad Member. The abundance of ichnofossils in sandstone, siltstone and in shale beds indicate that the ichnocenosis is dominated by a high behavioral diversity ranging from the suspension to deposit feeders. Three lithofacies were observed in this section, they show a vertical disposition, which further reflects general upward coarsening trend. Ichnofossils are mostly produced by arthropods along with crustacean, polychaetes and polyphyletic vermiforms. Due to the paucity of body fossil, as well as microbiota in the lowermost beds of the Debsakhad Member, the Precambrian-Cambrian boundary could not be demarcated. However, the presence of Treptichnus and Phycodes can be considered as a horizon marker for the beginning of Lower Cambrian in this section.

  11. Quartz cementation mechanisms between adjacent sandstone and shale in Middle Cambrian, West Lithuania

    Science.gov (United States)

    Zhou, Lingli; Friis, Henrik

    2013-04-01

    Quartz is an important cementing material in siliciclastic sandstones that can reduce porosity and permeability severely. For efficiently predicting and extrapolating petrophysical properties such as porosity and permeability, the controls on the occurrence and the degree of quartz cementation need to be better understood. Because it is generally difficult to identify specific sources for quartz cement, many models attempting to explain quartz cementation conclude that external sources of silica are needed to explain the observed quantity of quartz cement, such as the mass transfer between sandstone and shale. Cambrian sandstones in Lithuania have abundant quartz cement which has significant effect on reservoir properties. The detrital quartz grains have been dissolved extensively along the shale-quartz contacts zones, making it a natural laboratory to study the influence of mass transfer between sandstone and shale to quartz cementation on petrophysical properties and reservoir quality. Our Cambrian shale samples in west Lithuania are mainly silty shale or siltstone (sample locations vary from 330 to 2090 m of burial depth). They are composed of quartz, clay and traces of feldspars, sericite, calcite, and pyrite. The clay minerals are mainly illite, with variable content of kaolinite and traces of chlorite. In the sandstone lamina, authigenic illite occurs as pore-filling cement which was composed of fibrous illite; pore-filling kaolinite is generally well crystallized and occurs as hexagonal plates arranged in booklets; quartz overgrowth are obvious in these sandstone laminas, especially in the contact zones between sandstone and shale. Dolomite and pyrite cementation are also present in some sandstone laminas but with few quartz overgrowth. Depositional facies and architecture played an important role in the precipitation of silica. Three different possible sources are distinguished for the quartz overgrowths in the intercalated sandstones: 1) Pressure

  12. Replacement of benthic communities in two Neoproterozoic-Cambrian subtropical-to-temperate rift basins, High Atlas and Anti-Atlas, Morocco

    Science.gov (United States)

    Clausen, Sébastien; Álvaro, J. Javier; Zamora, Samuel

    2014-10-01

    The ‘Cambrian explosion’ is often introduced as a major shift in benthic marine communities with a coeval decline of microbial consortia related to the diversification of metazoans and development of bioturbation (‘Agronomic Revolution’). Successive community replacements have been reported along with ecosystem diversification and increase in guild complexity from Neoproterozoic to Cambrian times. This process is recorded worldwide but with regional diachroneities, some of them directly controlled by the geodynamic conditions of sedimentary basins. The southern High Atlas and Anti-Atlas of Morocco record development of two rifts, Tonian (?) - early Cryogenian and latest Ediacarian-Cambrian in age, separated by the onset of the Pan-African Orogeny. This tectonically controlled, regional geodynamic change played a primary control on pattern and timing of benthic ecosystem replacements. Benthic communities include microbial consortia, archaeocyathan-thromboid reefal complexes, chancelloriid-echinoderm-sponge meadows, and deeper offshore echinoderm-dominated communities. Microbial consortia appeared in deeper parts of the Tonian (?) - early Cryogenian fluvio-deltaic progradational rift sequences, lacustrine environments of the Ediacaran Volcanic Atlasic Chain (Ouarzazate Supergroup) and the Ediacaran-Cambrian boundary interval, characterized by the peritidal-dominated Tifnout Member (Adoudou Formation). They persisted and were largely significant until Cambrian Age 3, as previous restricted marine conditions precluded the immigration of shelly metazoans in the relatively shallow epeiric parts of the Cambrian Atlas Rift. Successive Cambrian benthic communities were replaced as a result of distinct hydrodynamic and substrate conditions, which allow identification of biotic (e.g., antagonistic relationships between microbial consortia and echinoderms, and taphonomic feedback patterns in chancelloriid-echinoderm-sponge meadows) and abiotic (e.g., rifting

  13. Early Cambrian eodiscoid trilobites of the Yangtze Platform and their stratigraphic implications

    Institute of Scientific and Technical Information of China (English)

    YANG Aihua; ZHU Maoyan; ZHANG Junming; LI Guoxiang

    2003-01-01

    Analysis of the taxonomy and stratigraphic distribution of the Early Cambrian eodiscoid trilobites of the Yangtze Platform indicates that species of Tsunyidiscus and Hupeidiscus can be used for biostratigraphic correlation. T. aclis occurs only in the lower Qiongzhusian, T. armatus occurs in the lower and middle Qiongzhusian, while T. niutitangensis and T. tingi occur mainly in the middle and upper Qiongzhusian, with only T. tingi extending upward into the lower Canglangpuian. H. orientalis first appears in the upper Qiongzhusian, reaching peak abundance in the lower Canglangpuian and becoming extinct above the Drepanuroides Zone. Biostratigraphic and lithostratigraphic correlations indicate that the limestone in the lower part of the Mingxinsi Formation, the Upper Member of Jiumenchong Formation, and the upper part of the Middle Member of the Huangbailing Formation, can be correlated with each other as Hupediscus, an index fossil of the lower Canglangpuian, is abundant in all these units. The lower part of the Niutitang and Jiumenchong formations in eastern Guizhou, which mainly represent Qiongzhusian strata, can be correlated with the Lower Member of the Huangbailing Formation. The basal part of the Niutitang Formation, which may be equivalent to the Upper Meishucunian Shiyantou Formation in eastern Yunnan, is very condensed. Well-preserved sponge faunas and non-mineralized fossils in the lower part of the Niutitang and Hetang formations are not older than Qiongzhusian in age.

  14. Experimental taphonomy and the anatomy and diversity of the earliest fossil vertebrates (Chengjiang Biota, Cambrian, China)

    Science.gov (United States)

    Purnell, Mark; Gabbott, Sarah; Murdock, Duncan; Cong, Peiyun

    2016-04-01

    The oldest fossil vertebrates are from the Lower Cambrian Chengjiang biota of China, which contains four genera of fish-like, primitive vertebrates: Haikouichthys, Myllokunmingia, Zhongjianichthys and Zhongxiniscus. These fossils play key roles in calibrating molecular clocks and informing our view of the anatomy of animals close to the origin of vertebrates, potentially including transitional forms between vertebrates and their nearest relatives. Despite the evident importance of these fossils, the degree to which taphonomic processes have affected their anatomical completeness has not been investigated. For example, some or all might have been affected by stemward slippage - the pattern observed in experimental decay of non-biomineralised chordates in which preferential decay of synapomorphies and retention of plesiomorphic characters would cause fossil taxa to erroneously occupy more basal positions than they should. This hypothesis is based on experimental data derived from decay of non-biomineralised chordates under laboratory conditions. We have expanded this analysis to include a broader range of potentially significant environmental variables; we have also compared and combined the results of experiments from several taxa to identify general patterns of chordate decay. Examination of the Chengjiang vertebrates in the light of these results demonstrates that, contrary to some assertions, experimentally derived models of phylogenetic bias are applicable to fossils. Anatomical and phylogenetic interpretations of early vertebrates that do not take taphonomic biases into account risk overestimating diversity and the evolutionary significance of differences between fossil specimens.

  15. The newly-developed Cambrian biostratigraphic succession and chronostratigraphic scheme for South China

    Institute of Scientific and Technical Information of China (English)

    PENG ShanChi

    2009-01-01

    Biostratigraphy and chronostratigraphy are common means of subdividing rock strata.The biostratigraphic succession and chronostratigraphic scheme are regarded as the basis for the subdivision and correlation of regional and global stratigraphic units.This paper reviews research on the biostratigraphy and chronostratigraphy carried out in the Jiangnan Slope Belt,South China since 1978,and,in particular,the biostratigraphic succession and chronostratigraphic scheme developed in the last decade.The updated biostratigraphic succession of South China includes 36 biozones,the base of each of which is defined by the first appearance of a single taxon (index fossil),and a poorly fossiliferous interzone.The updated chronostratigraphic scheme of South China comprises 4 series and 10 stages,embracing 3 global chronostratigraphic units,the Guzhangian Stage,Furongian Series,and Paibian Stage,and 2 GSSPa,all of which are erected in northwestern Hunan and 2 International Tie Points recognized in sections also in northwestern Hunan and each was accepted by the International Subcommission on Cambrian Stratigraphy for defining the base of the global stage.The biostratigraphic and chronostratigraphic systems are recognized as Chinese standards both domestically and worldwide.Future research is suggested for improving the two sets of stratigraphic standards.

  16. Phylogeny and evolutionary significance of vermiform animals from the Early Cambrian Chengjiang Lagersttte

    Institute of Scientific and Technical Information of China (English)

    David; BAINES

    2010-01-01

    The Early Cambrian Chengjiang Biota of China has preserved many species of vermiform animals in exquisite detail,which provide a unique window into the origin and early evolution of different vermiform phyla.This article reviews and discusses the forty-five Chengjiang vermiform animals discovered so far,and specifically comments on the phylogeny,modes of life,evolutionary significance as well as existing debates concerning the Chengjiang priapulid and lobopodian phyla.This article analyzes existing problems with Chengjiang vermiform morphological research and highlights various factors that affect the final preserved fossil morphology,including taphonomy and shape change caused by vermiform movement,as well as proposing that Anningvermis multispinosa and Corynetis brevis are synonymous.Chengjiang vermiform animals displayed a great diversity of morphologies and ecological roles,which is significant for understanding the origin and evolution of early metazoans;the potential for future research is enormous but needs to be broadened by interdisciplinary work involving taphonomy,geochemistry,cladistics,and genetics.

  17. Preservation and phylogeny of Cambrian ecdysozoans tested by experimental decay of Priapulus.

    Science.gov (United States)

    Sansom, Robert S

    2016-01-01

    The exceptionally preserved Cambrian fossil record provides unique insight into the early evolutionary history of animals. Understanding of the mechanisms of exceptional soft tissue preservation frames all interpretations of the fauna and its evolutionary significance. This is especially true for recent interpretations of preserved nervous tissues in fossil ecdysozoans. However, models of soft tissue preservation lack empirical support from actualistic studies. Here experimental decay of the priapulid Priapulus reveal consistent bias towards rapid loss of internal non-cuticular anatomy compared with recalcitrant cuticular anatomy. This is consistent with models of Burgess Shale-type preservation and indicates that internal tissues are unlikely to be preserved with fidelity if organically preserved. This pattern, along with extreme body margin distortion, is consistent with onychophoran decay, and is therefore resolved as general for early ecdysozoans. Application of these patterns to phylogenetic data finds scalidophoran taxa to be very sensitive to taphonomically informed character coding, but not panarthropodan taxa. Priapulid decay also have unexpected relevance for interpretation of myomeres in fossil chordates. The decay data presented serve not only as a test of models of preservation but also a framework with which to interpret ecdysozoan fossil anatomies, and the subsequent evolutionary inferences drawn from them. PMID:27595908

  18. Geochemical investigations on some of the Infra Cambrian Acid intrusive and volcanic rocks in Iran

    International Nuclear Information System (INIS)

    Geochemical investigations on some of the acid intrusive and volcanic rocks of Iran, which are attributed to the Infra cambrian, have been carried out to reveal their possible genetic relationships. These igneous rocks include: Do ran, Shah-Bolaghi, Sarve-Jahan, NE Ardakan - Yazd (Darbastegoon), south Mahabad, Bornavard (Tak nar), Zarigan, Narigan, Chardormaloo granites, Rizo and De zoo micro granites and volcanic rocks, Ga chin rhyolites (Bandar-Abass), and Ghareh-Dash (east of Sae en-De j). These intrusive s, except south Mahabad granodiorite, belong to the class of the alkali feldspathic granite to granitic rocks. Most of these rocks are hololeucocratic and devoid of ferro magnesian minerals. The volcanic rocks are mainly felsic (Rhyolite and Quartz porphyra) associated with vacillation's rocks. These suites of rocks plot on the chemical diagrams in sub alkaline field. Regarding to variations of Na2O, K2O and Ca O contents, they can further be subdivided into sodi c, potas sic, sodi-potas sic and sodi-calci c subgroups. The normalized values (MORB and chondritic) of their trace elements on the spider diagrams overlap each other indicates that some genetic relations ships exist among samples of white Doran Granite, Shahbolaghi, Sarve-Jahan and some Darbastegoon, in one hand, and Taknar, Narigan Granites, Gachin, Rizoo and Dezoo Rhyolites, on the other hand. The overlap also shows that the mentioned igneous bodies have been originated from continental crust and the samples of each groups, at least belong to a distinct tectonic al regime

  19. Original molluscan radula: comparisons among Aplacophora, Polyplacophora, Gastropoda, and the Cambrian fossil Wiwaxia corrugata.

    Science.gov (United States)

    Scheltema, Amélie H; Kerth, Klaus; Kuzirian, Alan M

    2003-08-01

    As the original molluscan radula is not known from direct observation, we consider what the form of the original radula may have been from evidence provided by neomenioid Aplacophora (Solenogastres), Gastropoda, Polyplacophora, and the Cambrian fossil Wiwaxia corrugata (Matthews). Conclusions are based on direct observation of radula morphology and its accessory structures (salivary gland ducts, radular sac, anteroventral radular pocket) in 25 species and 16 genera of Aplacophora; radula morphogenesis in Aplacophora; earliest tooth formation in Gastropoda (14 species among Prosobranchia, Opisthobranchia, and Pulmonata); earliest tooth formation in four species of Polyplacophora; and the morphology of the feeding apparatus in W. corrugata. The existence of a true radula membrane and of membranoblasts and odontoblasts in neomenioids indicates that morphogenesis of the aplacophoran radula is homologous to that in other radulate Mollusca. We conclude from p redness of salivary gland ducts, a divided radular sac, and a pair of anteroventral pockets that the plesiomorphic state in neomenioids is bipartite, formed of denticulate bars that are distichous (two teeth per row) on a partially divided or fused radula membrane with the largest denticles lateral, as occurs in the genus Helicoradomenia. The tooth morphology in Helicoradomenia is similar to the feeding apparatus in W. corrugata. We show that distichy also occurs during early development in several species of gastropods and polyplacophorans. Through the rejection of the null hypothesis that the earliest radula was unipartite and had no radula membrane, we conclude that the original molluscan radula was similar to the radula found in Helicoradomena species. PMID:12833382

  20. The importance of preadapted genomes in the origin of the animal bodyplans and the Cambrian explosion.

    Science.gov (United States)

    Marshall, Charles R; Valentine, James W

    2010-05-01

    The genomes of taxa whose stem lineages branched early in metazoan history, and of allied protistan groups, provide a tantalizing outline of the morphological and genomic changes that accompanied the origin and early diversifications of animals. Genome comparisons show that the early clades increasingly contain genes that mediate development of complex features only seen in later metazoan branches. Peak additions of protein-coding regulatory genes occurred deep in the metazoan tree, evidently within stem groups of metazoans and eumetazoans. However, the bodyplans of these early-branching clades are relatively simple. The existence of major elements of the bilaterian developmental toolkit in these simpler organisms implies that these components evolved for functions other than the production of complex morphology, preadapting the genome for the morphological differentiation that occurred higher in metazoan phylogeny. Stem lineages of the bilaterian phyla apparently required few additional genes beyond their diploblastic ancestors. As disparate bodyplans appeared and diversified during the Cambrian explosion, increasing complexity was accommodated largely through changes in cis-regulatory networks, accompanied by some additional gene novelties. Subsequently, protein-coding genic richness appears to have essentially plateaued. Some genomic evidence suggests that similar stages of genomic evolution may have accompanied the rise of land plants. PMID:19930449

  1. Neoproterozoic Land Colonisation, Rising Oxygen, Global Cooling and the Cambrian Explosion

    Science.gov (United States)

    Lenton, T. M.; Watson, A. J.

    2004-12-01

    The Neoproterozoic (1000-542 Ma BP) was a time of severe glaciations and a major transition from microscopic to macroscopic life forms. We develop the hypothesis that a rise in atmospheric oxygen in the Neoproterozoic was driven by the biological colonization of the land surface. If early forms of photosynthetic land life selectively weathered continental rock in order to extract nutrients, this would have led to an increase in the flux of biologically available phosphorus to the ocean. We show that recent models for coupled biogeochemical cycles, despite differences in the feedback mechanisms represented, predict this would lead to a rise in atmospheric oxygen concentration, consistent with biological and geochemical evidence. Increased weathering of silicate rocks would also have caused a decline in atmospheric carbon dioxide, which could have been a causal factor in the Neoproterozoic glaciations. A rise in oxygen may have provided a necessary condition for the evolution of animals with hard skeletons seen in the Cambrian explosion. Furthermore, an increase in phosphorus supply to the ocean may have driven an increase in the phosphorus content of marine primary producers. This would have represented an increase in food quality for grazing animals, which have a high phosphorus requirement, and may thus have removed a further limitation on their evolutionary radiation.

  2. Population Structure, Life Strategies and Systematics of Phosphatocope Ostracods from the Middle Cambrian of Bornholm

    Directory of Open Access Journals (Sweden)

    I. Hinz-Schallreuter

    1998-01-01

    Full Text Available She Middle Cambrian Borregård Member of Bornholm which is the stratigraphical equivalent to the Baltoscandian Exsulans Limestone yielded a rich and comparatively diverse ostracod fauna in its upper part. It comprises eight species out of four genera. They belong to three known subfamilies that are redefined on the basis of special characters of the contact margin. An ontogenetic character unknown from typical ostracods is documented in several species: during early ontogeny the gestalt (l:h ratio develops constantly in becoming increasingly longer until the so-called ontogenetical turning point (OTP from which the direction of growth focusses on carapace height. This phenomenon may be explained by changes in body morphology. The Borregård association represents an ecologic community type differing from other yet recorded Middle Cambrian communities in the Baltoscandic region. Apart from Vestrogothia longispinosa which is a common faunal element in Baltoscandian ostracod faunas, the yet recorded species of Bidimorpha are unknown from other Baltoscandian occurrences. Vice versa, the known species of Bidimorpha described from Swedish occurrences have not been recognized in the rich Borregård community. Similarly, Falites insula and Hesslandona abdominalis may be of local significance, too, but the respective records from contemporaneous Swedish localities are insufficient in this respect, yet. Due to specific morphological adaptations, the four genera are assumed to represent different benthic life strategies. New taxa are: Bidimorpha arator n. sp., Bidimorpha labiator n. sp., Bidimorpha sexspinosa n. sp., Falites insula n. sp., Hesslandona abdominalis n. sp., Vestrogothia herrigi n. sp. and Vestrogothia minilaterospinata n. sp. Aus dem mittelkambrischen Borregård Member von Bornholm, einem zeitlichen Äquivalent des baltoskandischen Exsulanskalkes, wurde eine reiche und vergleichsweise diverse Ostrakodenfauna gewonnen. Sie umfaßt 8 Arten aus

  3. From Cycles to Sequences: Sequence Stratigraphy and Relative Sea Level Change for the Late Cambrian of the North China Platform

    Institute of Scientific and Technical Information of China (English)

    MEI Mingxiang; MA Yongsheng; DENG Jun; CHEN Huijun

    2005-01-01

    In the Late Cambrian, the North China Platform was a typical carbonate ramp platform. The Upper Cambrian of the northern part of the North China Platform is famous for the development of bioherm limestones and storm calcirudites and can be divided from bottom to top into the Gushan, Changshan and Fengshan formations. In this set of strata, the deep-ramp mudstone and marls and the shallow-ramp packstones and grainstones constitute many carbonate meter-scale cycles of subtidal type. More tidal-flat dolomites are developed in the Upper Cambrian of the southern margin of the North China platform, in which limestone and dolomite beds also constitute many carbonate meter-scale cycles of the peritidal type. These cycles are marked by a variety of litho-facies successions. There are regularly vertical stacking patterns of meter-scale cycles in long-term third-order sequences, which is the key to discerning such sequences. Thirdorder sequence is marked by a particular sedimentary-facies succession that is the result of the environment-changing process of deepening and shoaling, which is genetically related to third-order sea level changes. Furthermore, four thirdorder sequences can be grouped in the Upper Cambrian of the North China Platform. The main features of these four thirdorder sequences in the northern part of the platform can be summarized as follows: firstly, sequence-boundaries are characterized by drowning unconformities; secondly, the sedimentary-facies succession is generally constituted by one from deep-ramp facies to shallow-ramp facies; thirdly, a succession of "CS (?)+HST" (i.e., "condensed section and highstand system") forms these four third-order sequences. The chief features for the third-order sequences in the southern part of the North China Platform comprises: more dolomites are developed in the HSTs of third-order sequences and also developed more carbonate meter-scale cycles of peritidal types; the sedimentary-facies succession of the third

  4. Effects of fracture reactivation and diagenesis on fracture network evolution: Cambrian Eriboll Formation, NW Scotland

    Science.gov (United States)

    Hooker, J. N.; Eichhubl, P.; Xu, G.; Ahn, H.; Fall, A.; Hargrove, P.; Laubach, S.; Ukar, E.

    2011-12-01

    The Cambrian Eriboll Formation quartzarenites contain abundant fractures with varying degrees of quartz cement infill. Fractures exist that are entirely sealed; are locally sealed by bridging cements but preserve pore space among bridges; are mostly open but lined with veneers of cement; or are devoid of cement. Fracture propagation in the Eriboll Formation is highly sensitive to the presence of pre-existing fractures. Fracture reactivation occurs in opening mode as individual fractures repeatedly open and are filled or bridged by syn-kinematic cements. As well, reactivation occurs in shear as opening of one fracture orientation coincides with shear displacement along pre-existing fractures of different orientations. The tendency for pre-existing fractures to slip varies in part by the extent of cement infill, yet we observe shear and opening-mode reactivation even among sealed fractures. Paleotemperature analysis of fluid inclusions within fracture cements suggests some fractures now in outcrop formed deep in the subsurface. Fractures within the Eriboll Formation may therefore affect later fracture propagation throughout geologic time. With progressive strain, fault zones develop within fracture networks by a sequence of opening-mode fracture formation, fracture reactivation and linkage, fragmentation, cataclasis, and the formation of slip surfaces. Cataclasite within fault zones is commonly more thoroughly cemented than fractures in the damage zone or outside the fault zone. This variance of cement abundance is likely the result of (1) continued exposure of freshly broken quartz surfaces within cataclasite, promoting quartz precipitation, and (2) possibly more interconnected pathways for mass transfer within the fault zone. Enhanced cementation of cataclasite results in strengthening or diagenetic strain hardening of the evolving fault zone. Further slip is accommodated by shear localization along discrete slip surfaces. With further linkage of fault segments

  5. The Middle Cambrian fossil Pikaia and the evolution of chordate swimming

    Directory of Open Access Journals (Sweden)

    Lacalli Thurston

    2012-07-01

    Full Text Available Abstract Conway Morris and Caron (2012 have recently published an account of virtually all the available information on Pikaia gracilens, a well-known Cambrian fossil and supposed basal chordate, and propose on this basis some new ideas about Pikaia’s anatomy and evolutionary significance. Chief among its chordate-like features are the putative myomeres, a regular series of vertical bands that extends the length of the body. These differ from the myomeres of living chordates in that boundaries between them (the myosepta are gently curved, with minimal overlap, whereas amphioxus and vertebrates have strongly overlapping V- and W-shaped myomeres. The implication, on biomechanical grounds, is that myomeres in Pikaia exerted much less tension on the myosepta, so the animal would have been incapable of swimming as rapidly as living chordates operating in the fast-twitch mode used for escape and attack. Pikaia either lacked the fast-twitch fibers necessary for such speeds, having instead only slow-twitch fibers, or it had an ancestral fiber type with functional capabilities more like modern slow fibers than fast ones. The first option is supported by the sequence of development in zebrafish, where both myoseptum formation and fast fiber deployment show a dependence on slow fibers, which develop first. For Pikaia, the absence of fast fibers has both behavioral and anatomical implications, which are discussed. Among the latter is the possibility that a notochord may not have been needed as a primary stiffening device if other structures (for example, the dorsal organ could perform that role.

  6. Depositional and diagenetic variability within the Cambrian Mount Simon Sandstone: Implications for carbon dioxide sequestration

    Science.gov (United States)

    Bowen, B.B.; Ochoa, R.I.; Wilkens, N.D.; Brophy, J.; Lovell, T.R.; Fischietto, N.; Medina, C.R.; Rupp, J.A.

    2011-01-01

    The Cambrian Mount Simon Sandstone is the major target reservoir for ongoing geologic carbon dioxide (CO2) sequestration demonstrations throughout the midwest United States. The potential CO2 reservoir capacity, reactivity, and ultimate fate of injected CO2 depend on textural and compositional properties determined by depositional and diagenetic histories that vary vertically and laterally across the formation. Effective and efficient prediction and use of the available pore space requires detailed knowledge of the depositional and diagenetic textures and mineralogy, how these variables control the petrophysical character of the reservoir, and how they vary spatially. Here, we summarize the reservoir characteristics of the Mount Simon Sandstone based on examination of geophysical logs, cores, cuttings, and analysis of more than 150 thin sections. These samples represent different parts of the formation and depth ranges of more than 9000 ft (>2743 m) across the Illinois Basin and surrounding areas. This work demonstrates that overall reservoir quality and, specifically, porosity do not exhibit a simple relationship with depth, but vary both laterally and with depth because of changes in the primary depositional facies, framework composition (i.e., feldspar concentration), and diverse diagenetic modifications. Diagenetic processes that have been significant in modifying the reservoir include formation of iron oxide grain coatings, chemical compaction, feldspar precipitation and dissolution, multiple generations of quartz overgrowth cementation, clay mineral precipitation, and iron oxide cementation. These variables provide important inputs for calculating CO2 capacity potential, modeling reactivity, and are also an important baseline for comparisons after CO2 injection. Copyright ??2011. The American Association of Petroleum Geologists/Division of Environmental Geosciences. All rights reserved.

  7. Enigmatic Connection Between Rhyolites and Shallow Granites in a Cambrian Rift

    Science.gov (United States)

    Gilbert, M. C.

    2007-12-01

    The Cambrian Southern Oklahoma Aulacogen (SOA) exposes shallow-seated, A-type sheet granites emplaced on top of layered mafic bodies and under cover of A-type rhyolites, this gabbro-rhyolite boundary being a crustal magma trap. Granites and rhyolites appear to overlap in age, although rhyolites built up first, with the rhyolite pile thickening as later granites coarsened texturally to medium-grained types from earlier fine-grained sheets. Outcrop relations + widespread subsurface data, including regional gravity, indicate rhyolite volume exceeds granite volume. Both silicic and mafic members of the SOA sequence have the same positive Nd signature showing a fundamental genetic relationship between the rifting process and the development of new crust in the SOA. Major element geochemistry of rhyolites (~76% SiO2) and granites (73-76% SiO2) is similar, as are many trace element abundances and signatures. However, some indices are different: Zr in rhyolites is ~600-700ppm, in granites ~300-500; Zr/Nb in rhyolites 8-11, in granites 4-8; K/Rb in rhyolites 300's, in granites 200's. In only one locality can a continuous connection between fine-grained granite and a texturally rhyolitic lobe be seen. Thus, it is clear that none of the rhyolites and granites are directly related at their emplacement level. Conclusions: 1)Silicic magma, both rhyolitic and granitic, was fractionated from mafic sources in the middle rift crust. 2)Magma driving pressures for silicic liquids must have been higher for those becoming rhyolites than for those that became granites. 3)These differences imply that rhyolitic magma generally came from a deeper depth in the mid-crust, although the ultimate mafic sources may have been similar. 4)More generally as this case shows, spatially near, and chemically similar, rhyolite and granite bodies in the same tectonic setting may not be directly related.

  8. Proterozoic sequences and their implications for precamorian and cambrian geologic evolution of Western Kentucky: Evidence from seismic-reflection data

    Science.gov (United States)

    Drahovzal, J.A.

    1997-01-01

    Analyses of two seismic-reflection lines in western Kentucky indicate the presence of two Proterozoic, unconformity-bounded sequences. One is autochthonous and of probable Late Proterozoic age; the other is allochthonous and of probable Middle Proterozoic age. Reflector patterns and apparent relationships to similar sequences elsewhere in the region suggest that the two sequences are of continental-rift origin. The two Proterozoic sequences lie beneath and adjacent to rocks of the Cambrian rift sequence in the Rough Creek Graben. The oldest sequence, the pre-Grenville sequence, was apparently folded and thrust faulted by the Grenville compressional event, implying that it is older than ???0.975 Ga (Middle Proterozoic). Two seismic-reflection pattern types are present in the western Kentucky data that may relate to the Middle Run (lithic arenite) and volcanic sequences defined farther east near the Grenville Front. The presence of imbricate, thrust-belt geometries in the pre-Grenville sequence extends the known westward limit of Grenville compressional structures into western Kentucky. The younger, post-Grenville sequence is less deformed and was apparently formed after the Grenville compressional event; several lines of evidence indicate that it is Late Proterozoic (0.7 to 0.6 Ga) in age. This probable siliciclastic and volcanic-rift sequence is represented by only thin remnants in western Kentucky and has no equivalent near the Grenville Front in southwestern Ohio and central Kentucky. Rocks of the better documented Cambrian rifting event belong to the thick, pre-Knox sequence in the Rough Creek Graben of western Kentucky and lie unconformably above these earlier sequences. A previously undocumented, northward-thickening interval within the lower part of the Cambrian pre-Knox sequence is recognized north of the Rough Creek Graben.

  9. Geochemical comparison of a Cambrian alginite potential source rock, and hydrocarbons from the Colville/Tweed Lake area, Northwest Territories

    Energy Technology Data Exchange (ETDEWEB)

    Wielens, J.B.W.; Fowler, M.G.; Brooks, P.W. (Inst. of Sedimentary and Petroleum Geology, Calgary, AB (Canada)); Dick, H. Von der (Canadian Hunter Exploration Ltd., Calgary, AB (Canada)); Monnier, F. (Chevron Oil Field Research, CA (USA))

    1990-06-01

    The reservoir for gas discoveries in the Colville/Tweed Lake area is the Lower Cambrian Mount Clark sandstone. Heavy oils are also found in this area in Cretaceous formations for which no source rock has previously been assigned. A comparison and correlation of reservoir and oil seep hydrocarbons is presented, and a potential source rock discussed. The gasoline range composition of Tweed Lake condensate shows an unusual distribution, distinguished by a very high abundance of n-alkanes and iso-alkanes and an extremely low abundance of benzene and toluene. The C{sub 15+} saturate fraction gas chromatograms of the condensate and of bitumens found with the Mount Clark reservoir also show many similarities to Ordovidian oils sourced from kukersite intervals. Geochemical screening of samples from several wells indicates that only the Middle Cambrian Mount Cap Formation is sufficiently organic-rich to be a probable source rock. Microscopic examination of one particularly rich interval shows dominance by alginite of a species resembling Gloeocapsomorpha prisca, the predominant contributor to kukersites. The organic matter is Type I, known to generate hydrocarbons at higher maturities. Organic geochemical investigations indicate that a source rock similar to this sample is responsible for the hydrocarbons in the Mount Clark reservoirs. The actual source rocks must be present in areas which have not been drilled to date. Bitumens recovered from surface seeps in the area are very biodegraded and cannot be correlated with any specific source rock, but they were not derived from the Cambrian source rock. 24 refs., 7 figs., 3 tabs.

  10. Paleomagnetic results from the Cambrian and Ordovician sediments of Bornholm (Denmark) and Southern Sweden and paleogeographical implications for Baltica

    Science.gov (United States)

    Lewandowski, Marek; Abrahamsen, Niels

    2003-11-01

    If apparent polar wander paths (APWP) cross, the question arises how to prove the older magnetization to be primary and not just a younger overprint. This problem is typically met in areas affected by percolating mineralizing fluids and/or heating due to a younger regional igneous activity. The Permian magnetic overprint is the classical example. Earlier paleomagnetic studies over the Lowermost Cambrian Nekso Sandstone Fm (NSF) of Bornholm (Denmark) yielded a characteristic remanent magnetization (ChRM) similar to the Permian directions for Baltica. Since a possible reason could be a chemical overprint, we checked whether this phenomenon did take place on a regional scale. Some samples therefore were collected from other Lower Cambrian clastics of Bornholm and Southern Scandinavia. In result we show that the well-grouped and stable ChRM of the NSF contrasts with fairly chaotic, soft, and badly preserved magnetizations of the Balka, Hardeberga, Mickwitzia, and Lingulid sandstones of Bornholm and Southern Sweden, thus not indicating widespread paleomagnetic overprint. We demonstrate that the ChRM of the NSF is most probably of syndepositional/early diagenetic origin and its similarity to the Permian direction for Baltica is only casual. We propose a normal polarity and a near-equatorial position on the Southern Hemisphere for Baltica in the early Cambrian time, as well as a more complicated trend of the APWP for this paleocontinent than envisaged by other authors. Paleomagnetic results from the Arenigian limestones of the Laesaa Formation (Bornholm) that yield excellently defined but most probably only secondary components are also presented.

  11. Trilobites from the Çal Tepe Formation (Cambrian), Near Seydişehir, Central Taurides, Southwestern Turkey

    OpenAIRE

    DEAN, WILLIAM T.

    2005-01-01

    The type section of the Çal Tepe Formation, near Seydişehir, is reviewed. The basal dolomite member is unfossiliferous, but the succeeding black limestone member (24 m), light-grey limestone member (10.15 m), and red nodular limestone member (46.7 m) are subdivided into thirty-seven numbered, often fossiliferous beds. Trilobites from the black limestone member, late Early Cambrian, exhibit affinities with Morocco, Spain and northwestern Europe; they include one new genus and species (Pamphyli...

  12. Detrital zircon U-Pb geochronology of Cambrian to Triassic miogeoclinal and eugeoclinal strata of Sonora, Mexico

    Science.gov (United States)

    Gehrels, G.E.; Stewart, John H.

    1998-01-01

    One hundred and eighty two individual detrital zircon grains from Cambrian through Permian miogeoclinal strata, Ordovician eugeoclinal rocks, and Triassic post-orogenic sediments in northwestern Sonora have been analyzed. During Cambrian, Devonian, Permian, and Triassic time, most zircons accumulating along this part of the Cordilleran margin were shed from 1.40-1.45 and 1.62-1.78 Ga igneous rocks that are widespread in the southwestern United States and northwestern Mexico. Zircons with ages of approximately 1.11 Ga are common in Cambrian strata and were apparently shed from granite bodies near the sample site. The sources of 225-280 Ma zircons in our Triassic sample are more problematic, as few igneous rocks of these ages are recognized in northwestern Mexico. Such sources may be present but unrecognized, or the grains could have been derived from igneous rocks of the appropriate ages to the northwest in the Mojave Desert region, to the east in Chihuahua and Coahuila, or to the south in accreted(?) arc-type terranes. Because the zircon grains in our Cambrian and Devonian to Triassic samples could have accumulated in proximity to basement rocks near their present position or in the Death Valley region of southern California, our data do not support or refute the existence of the Mojave-Sonora megashear. Ordovician strata of both miogeoclinal and eugeoclinal affinity are dominated by >1.77 Ga detrital zircons, which are considerably older than most basement rocks in the region. Zircon grains in the miogeoclinal sample were apparently derived from the Peace River arch area of northwestern Canada and transported southward by longshore currents. The eugeoclinal grains may also have come from the Peace River arch region, with southward transport by either sedimentary or tectonic processes, or they may have been shed from off-shelf slivers of continents (perhaps Antarctica?) removed from the Cordilleran margin during Neoproterozoic rifting. It is also possible that the

  13. Trace fossils accompanying possible "Ediacaran organisms" in the Middle Cambrian sediments of the St. Petersburg Region, Russia

    Czech Academy of Sciences Publication Activity Database

    Natalin, N. M.; Mikuláš, Radek; Dronov, A. V.

    2010-01-01

    Roč. 60, č. 1 (2010), s. 71-75+[2]. ISSN 0001-5709 R&D Projects: GA ČR GA205/04/0151; GA ČR GA205/09/1521 Institutional research plan: CEZ:AV0Z30130516 Keywords : Skolithos ichnofacies * Edicarian-like organism * Middle Cambrian * shallow-marine * Baltic Shied * ichnology (Russia) Subject RIV: DB - Geology ; Mineralogy Impact factor: 0.779, year: 2010 https://geojournals.pgi.gov.pl/agp/article/view/9821/8355

  14. From Basin Black Shales to Platform Carbonate Rocks: A Study on Sequence Stratigraphy for the Lower Cambrian of the Upper-Yangtze Region in South China

    Institute of Scientific and Technical Information of China (English)

    2007-01-01

    In the Upper-Yangtze region, especially in Guizhou Province and its adjacent areas, the Lower Cambrian is well developed and is marked by a succession from black shales of the basin facies to carbonate rocks of the platform facies. The drowning event of the platform occurring at the turn from Sinian to Cambrian resulted in a set of black shales, i.e. the Niutitang Formation, which makes up the bottom part of the Lower Cambrian. With the shoaling of the sedimentary environment, a set of carbonate rocks, i.e. the Qingxudong Formation, was formed in the top part of the Lower Cambrian. Thus, the Lower Cambrian in the study area makes up one second-order sequence that can be further subdivided into five third-order sequences, and forms a regularly cyclic succession of transgression-regression. There is a regularly vertical stacking pattern for the third-order sequences in the second-order sequence. From bottom to top, the succession of the "CS (condensed section)+HST (high-stand system tract)" of the third-order sequences is changed into the succession of the "TST (transgressive system tract)+CS+HST". Correspondingly, the drowning-type sequence boundary is changed into the exposure-type one. Therefore, both the second-order and the third-order sequences have similar sedimentary-facies architectures. A concomitant with these temporal changes,the Lower Cambrian with a thickness of 1000 m that contains five third-order sequences is changed into a condensed succession that cannot identify third-order sequences toward the southeast with the deepening of the sedimentary environment. According to the elementary features of the third-order sequences, i.e. the regularity o sedimentary-facies successions in space and the synchronism of sedimentary-environment changes in time, the detailed division of the third-order sequences at main logged sections in different paleogeographical background becomes the basis to establish the sequence-stratigraphic framework that can demonstrate

  15. Paleoceangraphic significance of redox-sensitive metals of black shales in the basal Lower Cambrian Niutitang Formation in Guizhou Province,South China

    Institute of Scientific and Technical Information of China (English)

    YANG Jinghong; JIANG Shaoyong; LING Hongfei; FENG Hongzhen; CHEN Yongquan; CHEN Jianhua

    2004-01-01

    Black shales of the Lower Cambrian Niutitang Formation occur widely on the Yangtze Platform. In this study, we analyzed black shales from two sections (Zhongnan and Zhijin) in Guizhou Province. The redox-sensitive metal concentration and distribution of black shales from the two sections provide good evidence for the anoxic conditions of the basal Cambrian ocean on the Yangtze Platform. Our geochemical data suggest that the black shales from the Zhongnan section may have deposited in a more strongly anoxic environment than the black shales in the Zhijin section.

  16. "The Chengjiang Fauna and the Cambrian Explosion" Sponsord by NSFC won the first-class award of National Natural Sciences Award

    Institute of Scientific and Technical Information of China (English)

    Liu; Y.; Yao; Y.P.

    2004-01-01

      "The Chengjiang Fauna and the Cambrian explosion'' sponsord by NSFC won the first-class award of National Natural Sciences Award. The Awarding ceremony was held in the Great Hall of the People in Beijing on February 20, 2004. "The Chengjiang Fauna and the Cambrian explosion" was sponsord by NSFC and carried out by several groups led by Prof. Chen Junyuan of the Nanjing Institute of Geology and Palaeontology,CAGS, Prof. Hou Xianguang of Yunnan University and Prof.……

  17. A normalised seawater strontium isotope curve: possible implications for Neoproterozoic-Cambrian weathering rates and the further oxygenation of the Earth

    Directory of Open Access Journals (Sweden)

    G. A. Shields

    2007-07-01

    Full Text Available The strontium isotope composition of seawater is strongly influenced on geological time scales by changes in the rates of continental weathering relative to ocean crust alteration. However, the potential of the seawater 87Sr/86Sr curve to trace globally integrated chemical weathering rates has not been fully realised because ocean 87Sr/86Sr is also influenced by the isotopic evolution of Sr sources to the ocean. A preliminary attempt is made here to normalise the seawater 87Sr/86Sr curve to plausible trends in the 87Sr/86Sr ratios of the three major Sr sources: carbonate dissolution, silicate weathering and submarine hydrothermal exchange. The normalised curve highlights the Neoproterozoic-Phanerozoic transition as a period of exceptionally high continental influence, indicating that this interval was characterised by a transient increase in global weathering rates and/or by the weathering of unusually radiogenic crustal rocks. Close correlation between the normalised 87Sr/86Sr curve, a published seawater δ34S curve and atmospheric pCO2 models is used here to argue that elevated chemical weathering rates were a major contributing factor to the steep rise in seawater 87Sr/86Sr from 650 Ma to 500 Ma. Elevated weathering rates during the Neoproterozoic-Cambrian interval led to increased nutrient availability, organic burial and to the further oxygenation of Earth's surface environment. Use of normalised seawater 87Sr/86Sr curves will, it is hoped, help to improve future geochemical models of Earth System dynamics.

  18. Organic Carbon Isotopic Evolution during the Ediacaran-Cambrian Transition Interval in Eastern Guizhou, South China: Paleoenvironmental and Stratigraphic Implications

    Institute of Scientific and Technical Information of China (English)

    YANG Xinglian; ZHU Maoyan; GUO Qingjun; ZHAO Yuanlong

    2007-01-01

    Secular variations of carbon isotopic composition of organic carbon can be used in the study of global environmental variation, the carbon cycle, stratigraphic delimitation, and biological evolution, etc. Organic carbon isotopic analysis of the Nangao and Zhalagou sections in eastern Guizhou reveals a negative excursion near the Precambrian-Cambrian boundary that correlates with a distinct carbonate carbon isotopic negative excursion at this boundary globally. Our results also demonstrate that several alternating positive and negative shifts occur in the Meishucunian, and an obvious negative anomaly appears at the boundary between the Meishucunian and Qiongzhusian. The isotope values are stable in the middle and lower parts but became more positive in the upper part of the Qiongzhusian. Evolution of organic carbon isotopes from the two sections in the deepwater facies can be well correlated with that of the carbonate carbon isotopes from the section in the shallow water facies. Integrated with other stratigraphic tools, we can precisely establish a lower Cambrian stratigraphic framework from shallow shelf to deep basin of the Yangtze Platform.

  19. Oxygen-isotope trends and seawater temperature changes across the Late Cambrian Steptoean positive carbon-isotope excursion (SPICE event)

    Science.gov (United States)

    Elrick, M.; Rieboldt, S.; Saltzman, M.; McKay, R.M.

    2011-01-01

    The globally recognized Late Cambrian Steptoean positive C-isotope excursion (SPICE) is characterized by a 3???-5??? positive ??13C shift spanning <4 m.y. Existing hypotheses suggest that the SPICE represents a widespread ocean anoxic event leading to enhanced burial/preservation of organic matter (Corg) and pyrite. We analyzed ??18O values of apatitic inarticulate brachiopods from three Upper Cambrian successions across Laurentia to evaluate paleotemperatures during the SPICE. ??18O values range from ~12.5??? to 16.5???. Estimated seawater temperatures associated with the SPICE are unreasonably warm, suggesting that the brachiopod ??18O values were altered during early diagenesis. Despite this, all three localities show similar trends with respect to the SPICE ??13C curve, suggesting that the brachiopod apatite preserves a record of relative ??18O and temperature changes. The trends include relatively high ??18O values at the onset of the SPICE, decreasing and lowest values during the main event, and an increase in values at the end of the event. The higher ??18O values during the global extinction at the onset of the SPICE suggests seawater cooling and supports earlier hypotheses of upwelling of cool waters onto the shallow shelf. Decreasing and low ??18O values coincident with the rising limb of the SPICE support the hypothesis that seawater warming and associated reduced thermohaline circulation rates contributed to decreased dissolved O2 concentrations, which enhanced the preservation/burial of Corg causing the positive ??13C shift. ?? 2011 Geological Society of America.

  20. Experimental mineralization of crustacean eggs leads to surprising tissue conservation: new implications for the fossilization of Precambrian-Cambrian embryos

    Directory of Open Access Journals (Sweden)

    D. Hippler

    2011-12-01

    Full Text Available Phosphatized globular microfossils from the Ediacaran and Lower Cambrian of South China represent an impressive record of early animal evolution and development, however their affinity based on putative embryonic metazoan, bacterial and inorganic features is strongly debated. Understanding key processes and conditions that cause exceptional egg and embryo preservation and fossilization are therefore crucial for a reliable interpretation of their phylogenetic position. Taphonomic experiments on eggs of the marbled crayfish indicate a close link between early mineralization and rapid anaerobic decay of the endochorional envelope, producing different preservational stages of degradation resembling the various decay stages observed in the fossil record. Stabilization of the spherical morphology was achieved by pre-heating of the eggs. Complete surface mineralization occurred under reduced conditions within one to two weeks, with fine-grained brushite (CaHPO4·2H2O over calcite as the dominating mineral phase. Although the endochorional envelope was not preserved, experiments resulted in exceptional preservation of the embryonic tissue at the cellular level. Thus our findings suggest that the mechanisms of decay, preservation of surface structures, and mineral replacement in the experiment and during fossilization of Cambrian embryos were likely operating at a similar rationale.

  1. Cambrian magmatism, Variscan high-grade metamorphism and imposed greenschist facies shearing in the Central Sredna Gora basement units (Bulgaria

    Directory of Open Access Journals (Sweden)

    Lazarova Anna

    2015-12-01

    Full Text Available Gneisses from the deep structural levels of the European Variscan Belt are well exposed in the Central Sredna Gora in Bulgaria. In general, migmatites predominate, but unmigmatized domains (or domains with incipient migmatization are also documented in this area. This paper presents new structural, petrographic and U-Pb isotope geochronological data from such an unmigmatized part of the Variscan high-grade metamorphic basement (the Koprivshtitsa Unit. A predominant part of this unit represents an alternation of metagranitoids and metabasites. The protolith crystallization age of the metagranitoids is constrained at 491.5 ± 7.6 Ma by U-Pb laser ablation method on zircons. This age coincides with the previously available Late Cambrian protolith ages of metabasic rocks that crop out within the adjacent migmatitic unit. The Korpivshtitsa Unit comprises also lesser orthogneisses with Late Neoproterozoic protoliths. Based on the available local and regional paleogeographic reconstruction schemes, we suggest that the Upper Cambrian magmatic rocks intruded Upper Neoproterozoic crust during the initial opening stages of the Rheic Ocean or a related basin. Subsequently, both were involved in the Variscan high-grade deformation. The contact of the Koprivshitsa Unit with the migmatitic part of the metamorphic complex coincides with a north-vergent greenschist facies thrust zone – the Chuminska Shear Zone. The exact time of the shearing is still not well constrained but it clearly postdates the Variscan high-temperature metamorphism of the gneisses.

  2. Experimental mineralization of crustacean eggs leads to surprising tissue conservation: new implications for the fossilization of Precambrian-Cambrian embryos

    Science.gov (United States)

    Hippler, D.; Hu, N.; Steiner, M.; Scholtz, G.; Franz, G.

    2011-12-01

    Phosphatized globular microfossils from the Ediacaran and Lower Cambrian of South China represent an impressive record of early animal evolution and development, however their affinity based on putative embryonic metazoan, bacterial and inorganic features is strongly debated. Understanding key processes and conditions that cause exceptional egg and embryo preservation and fossilization are therefore crucial for a reliable interpretation of their phylogenetic position. Taphonomic experiments on eggs of the marbled crayfish indicate a close link between early mineralization and rapid anaerobic decay of the endochorional envelope, producing different preservational stages of degradation resembling the various decay stages observed in the fossil record. Stabilization of the spherical morphology was achieved by pre-heating of the eggs. Complete surface mineralization occurred under reduced conditions within one to two weeks, with fine-grained brushite (CaHPO4·2H2O) over calcite as the dominating mineral phase. Although the endochorional envelope was not preserved, experiments resulted in exceptional preservation of the embryonic tissue at the cellular level. Thus our findings suggest that the mechanisms of decay, preservation of surface structures, and mineral replacement in the experiment and during fossilization of Cambrian embryos were likely operating at a similar rationale.

  3. Isotopic evolution of the terminal Neoproterozoic and early Cambrian carbon cycle on the northern Yangtze Platform, South China

    Institute of Scientific and Technical Information of China (English)

    GUO Qingjun; LIU Congqiang; Harald STRAUSS; Tatiana GOLDBERG

    2003-01-01

    Profound geotectonic, climatic and biological changes occur during the terminal Neoproterozoic and its transition into the early Cambrian. These are reflected in temporal variations of the chemical and isotopic composition of seawater. We are studying a sequence of sedimentary rocks at the Shatan section, northern Yangtze Platform, Sichuan Province of China. This succession comprises, in ascending stratigraphic order, predominantly calcareous sediments of the Sinian upper Dengying Formation and black shales of the lower Cambrian Guojiaba Formation (time equivalent of Niutitang Fm.). Paleoenvironmental setting represents shallow-water shelf deposits. The objective of our study is to provide temporal records for the isotopic compositions of organic and carbonate carbon throughout this time interval. Organic carbon isotope values display a range between -35.8‰ and -30.1‰ with clear stratigraphic variations. Carbonate carbon isotope data vary between -3.5‰ and +0.5‰. These secular variations are interpreted to reflect perturbations of the global carbon cycle, specifically changes in the fractional burial of organic carbon. However, local conditions have further affected the isotopic signals.

  4. Proterozoic-Cambrian detrital zircon and monazite ages from the Anakie lnlier, central Queensland: Grenville and Pacific-Gondwana signatures

    International Nuclear Information System (INIS)

    The Anakie Metamorphic Group is a complexly deformed, dominantly metasedimentary succession in central Queensland. Metamorphic cooling is constrained to ca 500 Ma by previously published K-Ar ages. Detrital-zircon SHRIMP U-Pb ages from three samples of greenschist facies quartz-rich psammites (Bathampton Metamorphics), west of Clermont, are predominantly in the age range 1300-1000Ma (65-75%). They show that a Grenville-aged orogenic belt must have existed in northeastern Australia, which is consistent with the discovery of a potential Grenville source farther north. The youngest detrital zircons in these samples are ca 580 Ma, indicating that deposition may have been as old as latest Neoproterozoic. Two samples have been analysed from amphibolite facies pelitic schist from the western part of the inlier (Wynyard Metamorphics). One sample contains detrital monazite with two age components of ca 580-570 Ma and ca 540 Ma. The other sample only has detrital zircons with the youngest component between 510Ma and 700Ma (Pacific-Gondwana component), which is consistent with a Middle Cambrian age for these rocks. These zircons were probably derived from igneous activity associated with rifting events along the Gondwanan passive margin. These constraints confirm correlation of the Anakie Metamorphic Group with latest Neoproterozoic - Cambrian units in the Adelaide Fold Belt of South Australia and the Wonominta Block of western New South Wales. Copyright (2001) Geological Society of Australia

  5. 40Ar/39Ar dating of exceptional concentration of metals by weathering of Precambrian rocks at the Precambrian–Cambrian boundary

    DEFF Research Database (Denmark)

    Parnell, John; Mark, Darren F.; Frei, Robert; Fallick, Anthony E.; Ellam, Rob M.

    2014-01-01

    peneplanation. An 40Ar/39Ar age of 542.62 ± 0.38 Ma (1 sigma, full external precision, Renne et al., 2011) for metalliferous alteration clays in Scotland shows that this event occurred immediately prior to the Precambrian–Cambrian boundary. A negative δ53Cr isotopic signature for the clay is consistent with...

  6. Substrate as test of behaviour and as the ichnotaxonomical problem: predation and scavenging traces on trilobite exosceletons, Middle Cambrian, Czech Republic

    Czech Academy of Sciences Publication Activity Database

    Mikuláš, Radek; Fatka, O.; Szabad, M.

    Moscow : PIN RAN, 2010. s. 39-40. [Workshop on Ichnotaxonomy /4./. 21.06.2010-26.06.2010, Moscow - St. Petersburg] R&D Projects: GA ČR GA205/09/1521 Institutional research plan: CEZ:AV0Z30130516 Keywords : Cambrian * ichnology * ichnotaxonomy * ichnofossils * predation traces * scavening traces * palaeontology Subject RIV: DB - Geology ; Mineralogy

  7. U-Pb zircon ages and structural development of metagranitoids of the Teplá crystalline complex: evidence for pervasive Cambrian plutonism within the Bohemian massif (Czech Republic)

    Czech Academy of Sciences Publication Activity Database

    Dörr, W.; Fiala, Jiří; Vejnar, Zdeněk; Zulauf, G.

    1998-01-01

    Roč. 87, č. 1 (1998), s. 135-149. ISSN 0016-7835 R&D Projects: GA AV ČR KSK1012601 Grant ostatní: DFG(DE) Zu73/1-3 Keywords : Cambrian plutonism * U-Pb analyses * Bohemian massif Subject RIV: DB - Geology ; Mineralogy Impact factor: 0.841, year: 1998

  8. Germanium/silicon of the Ediacaran-Cambrian Laobao cherts: Implications for the bedded chert formation and paleoenvironment interpretations

    Science.gov (United States)

    Dong, Lin; Shen, Bing; Lee, Cin-Ty A.; Shu, Xu-jie; Peng, Yang; Sun, Yuanlin; Tang, Zhuanhong; Rong, Hong; Lang, Xianguo; Ma, Haoran; Yang, Fan; Guo, Wen

    2015-03-01

    Sedimentary strata of the terminal Ediacaran (635-542 Ma) to early Cambrian (542-488 Ma) Laobao-Liuchapo bedded cherts in the South China Block include the Ediacaran Oxidation Event and the Cambrian explosion. Understanding the origin and depositional environment of the bedded cherts may provide insight into how the Earth's surface environment changed between the Proterozoic and Phanerozoic. We measured major and trace element compositions and Ge/Si ratios of the Laobao cherts from northern Guangxi province. The Laobao cherts were deposited in the deep basinal environment of the South China Block. We show that the composition of the Laobao cherts is determined by a mixture of four components: quartz, clay, carbonate, and pyrite/iron-oxide. The quartz component is the dominant component of the Laobao cherts. The maximum estimated Ge/Si of the quartz component is between 0.4 and 0.5 μmol/mol, which is close to the Ge/Si of modern seawater and biogenic silica but 1 order of magnitude lower than that of hydrothermal fluids. These Ge/Si systematics suggest that normal seawater rather than mid-ocean ridge hydrothermal fluids is the primary Si source for the Laobao cherts. The Ge/Si of the clay component varies between 1 and 10 μmol/mol, which is comparable to the Ge/Si of typical marine clays, but 10-100 times lower than that of chert nodules from early Ediacaran beds (the Doushantuo Formation) predating the terminal Ediacaran Laobao cherts studied here. Our observations indicate that the clay component Ge/Si ratio decreased from the early Ediacaran to the late Ediacaran. We speculate that high Ge/Si ratios in clays reflect the preferential chelation of Ge by dissolved organic compounds adsorbed onto clays. If so, this suggests that the decrease in Ge/Si ratio of the clay component in the Ediacaran signifies a decrease in the total dissolved organic carbon content of seawater toward the Ediacaran-Cambrian transition, consistent with oxidation of the oceans during the

  9. The Kalkarindji Large Igneous Province and the Early-Middle Cambrian Extinction.

    Science.gov (United States)

    Jourdan, Fred; Evins, Lena

    2010-05-01

    Despite being one of the largest large igneous provinces (LIP) on Earth, the recently discovered Kalkarindji LIP, Australia, has received only very little attention (e.g., Glass & Philips, 2005; Evins et al., 2009). This province is located across the Northern Territories and Western Australia and covers a currently known area of ≥ 2.1x106 km² (Evins et al., 2009) with hints that it stretches as far as the southernmost part of South Australia (equivalent to a total area of ≥ 3x106 km²). The province includes flows, intrusions and volcanic tuffs. The age of the province is currently based only on three robust 40Ar/39Ar apparent ages ranging from 504.6 ± 2.5 to 507.5 ± 1.6 Ma (Glass & Philips, 2005; Evins et al., 2009). This province is of particular interest as it seems to coincide with a global anoxic event (Hough et al., 2006) and more importantly, the Early-Middle Cambrian (EMC) extinction where 50% of the genera (e.g. Archeocyaths, Trilobites) went extinct. The age of the EMC extinction has been estimated at ~510 Ma according to the revised timescale of 2009 but so far, attempts to date the extinction level have not yielded any precise date. Considering the now well-established 40K decay constant bias (Min et al., 2000), the 40Ar/39Ar ages obtained so far suggest an emplacement age of the Kalkarindji province between ~509 and ~512 Ma, thus apparently synchronous with the EMC extinction. However, due to the paucity of available ages, it is still not clear if the Kalkarindji LIP was emplaced during only few hundred ka, or rather was more sluggish over few Ma. The Kalkarindji basalts are overlapping thick sulfate-rich evaporite layers and carbonate rocks of the Precambrian central Australia basins. Evaporite and carbonate metamorphism might have been responsible for a significant source of CO2, SO2 and halocarbons as demonstrated for the Siberian traps (Svensen et al., 2009). In addition, the occurrence of pyroclastics eruptions as indicated by the presence

  10. Detrital zircon provenance of Mesoproterozoic to Cambrian arenites in the Western United States and Northwestern Mexico

    Science.gov (United States)

    Stewart, John H.; Gehrels, G.E.; Barth, A.P.; Link, P.K.; Christie-Blick, N.; Wrucke, C.T.

    2001-01-01

    U-Pb isotopic dating of detrital zircon from supracrustal Proterozoic and Cambrian arenites from the western United States and northern Mexico reveal three main age groups, 1.90 to 1.62 Ga, 1.45 to 1.40 Ga, and 1.2 to 1.0 Ga. Small amounts of zircons with ages of 3.1 to 2.5 Ga, 1.57 Ga, 1.32 Ga, 1.26 Ga, 0.7 Ga, and 0.5 Ga are also present. Detrital zircons ranging in age from 1.90 to 1.62 Ga and from 1.45 to 1.40 Ga are considered to have been derived from Proterozoic crystalline basement rocks of these known ages, and probably in part from reworked Proterozoic supracrustal sedimentary rocks, of the western United States. The 1.2 to 1.0 Ga detrital zircon ages from California, Arizona, and Sonora are characterized by distinct spikes (1.11 Ga, in particular) in the age-probability plots. These spikes are interpreted to indicate the influx of zircon from major silicic volcanic fields. Igneous rocks such as the Pikes Peak Granite (1.093 Ga) of Colorado, and the Aibo Granite (1.110 Ga) of Sonora, Mexico, may represent the deeply eroded roots of such volcanic fields. Samples from farther north along the Cordilleran margin that contain abundant 1.2-1.0 Ga detrital zircons do not show spikes in the age distribution, but rather ages spread out across the entire 1.2-1.0 Ga range. These age spectra resemble those for detrital zircons from the Grenville province, which is considered their source. Less common detrital zircons had a variety of sources. Zircons ranging in age from 3.36 to 2.31 Ga were apparently derived from inland parts of the North American continent from Wyoming to Canada. Zircons of about 1.577 Ga are highly unusual and may have had an exotic source; they may have come from Australia and been deposited in North America when Australia and North America were juxtaposed as part of the hypothetical Rodinian supercontinent. Detrital zircon of ??1.320 Ga apparently had the same source as that for tuff (1.320 Ga) in the Pioneer Shale of the Apache Group in Arizona

  11. Lack of inhibiting effect of oil emplacement on quartz cementation: Evidence from Cambrian reservoir sandstones, Paleozoic Baltic Basin

    DEFF Research Database (Denmark)

    Molenaar, Nicolaas; Cyziene, Jolanta; Sliaupa, Saulius;

    2008-01-01

    cementation is derived from internal sources. Rather, in spite of large variation in porosity and quartz cement content, a regular pattern of porosity decrease is related to increasing temperature or depth. The observed heterogeneity is due to local factors that influence the precipitation of quartz cement......Currently, the question of whether or not the presence of oil in sandstone inhibits quartz cementation and preserves porosity is still debated. Data from a number of Cambrian sandstone oil fields and dry fields have been studied to determine the effects of oil emplacement on quartz cementation. The......, including sandstone architecture, i.e., distribution of shales within the sandstone bodies, and sandstone thickness. Heterogeneity is inherent to sandstone architecture and to the fact that silica for quartz cementation is derived from heterogeneously distributed local pressure solution. Models predicting...

  12. Speciation of the genus Arthricocephalus Bergeron, 1899 (Trilobita) from the late Early Cambrian and its stratigraphic significance

    Institute of Scientific and Technical Information of China (English)

    YUAN Jinliang; ZHAO Yuanlong; YANG Xinglian

    2006-01-01

    The genus Arthricocephalus Bergeron,1899 is revised, and Halipanktos Balker & Peel, 1997 is suggested here as a senior synonym. The subgenus Arthricocephalus (Arthricocephalites) Chien & Lin in Lu et al., 1974 is considered as a separate genus.Of the 20 previously assigned species of Arthricocephalus ( Arthricocephalus ) Bergeron, 1899, Arthricocephalus ( Arthricocephalites )Chien & Lin in Lu et al., 1974, Arthricocephalus (Euarthricocephalus) Ju, 1983 are lumped into eight species. The speciation trend of Arthricocephalus and Arthricocephalites is demonstrated based on their stratigraphic occurrences. It not only enhances the resolution of the biostatigraphic zonation in the uppermost Lower Cambrian, but also represents a potential candidate to define the Duyunian stage. The base of the stage is suggested at the first appearance datum (FAD) of Arthricocephalus chauveaui Bergeron, 1899 within the evolutionary lineage from Ar. jiangkouensis Yin in Yin & Li, 1978 to Ar. chauveaui in a global scale.

  13. Late Ediacaran-Cambrian structures and their reactivation during the Variscan and Alpine cycles in the Anti-Atlas (Morocco)

    Science.gov (United States)

    Soulaimani, A.; Michard, A.; Ouanaimi, H.; Baidder, L.; Raddi, Y.; Saddiqi, O.; Rjimati, E. C.

    2014-10-01

    The post-Pan-African evolution of the northern border of the West African Craton is largely controlled by the remobilisation of Late Neoproterozoic basement faults. The Upper Ediacaran volcanic and volcano-sedimentary sequences of the Ouarzazate Group show dramatic and rapid thickness changes, consistent with active extensional faulting associated with post-orogenic collapse and incipient continental rifting. The geometry and kinematics of these faults differ from west to east in the Anti-Atlas. N- to NE-trending faults dominate in western Anti-Atlas in response to E-W to NW-SE pure extension, while a transtensive opening regime characterize the central (Bou Azzer) and eastern (Saghro-Ougnate) Anti-Atlas. The marine incursion in the west-central Anti-Atlas during the late Ediacaran-Early Cambrian occurred without major geodynamical break between the continental Ouarzazate Group and marine sediments of the Adoudou Fm. Extensional tectonics went on during the Early Cambrian, being concentrated in the western and central parts of the belt. From Middle Cambrian to Lower Devonian and mainly due to thermal subsidence, the Anti-Atlas basement was buried under marine sediments with dominant south-derived detrital input. Basement faults control the distribution of subsiding versus shallow areas. During the Middle-Late Devonian, the dislocation of the Saharan platform occurred, mainly in the eastern Anti-Atlas where Precambrian faults were also remobilized during the Early Carboniferous. During the Variscan orogeny, the Paleozoic series of the Anti-Atlas basin were involved in folding tectonics, concomitant with the uplift of Proterozoic basement blocks bounded by inherited basement faults. The pre-existing rift-related faults were variably inverted across the Anti-Atlas. In the westernmost part of the belt, Variscan shortening induced positive inversions along the remobilized basement faults, but in some cases, some faults preserved an apparently normal throw. Some hidden

  14. Source tracing of noble metal elements in Lower Cambrian black rock series of Guizhou-Hunan Provinces, China

    Institute of Scientific and Technical Information of China (English)

    2000-01-01

    The Lower Cambrian black rock series of South China is abnormally rich in noble metal elements. According to the concentrations, the ratios, the relations, the distribution and partition patterns of noble metal elements, the authors think that the noble metals and other elements are neither directly from extraterrestrial materials, nor from the products of normal marine sedimentation. The abnormal enrichment of noble metal elements is closely related with hydrothermal fluid that flew out on the sea floor through deep cycling and reaction with Proterozoic ultramafic-mafic igneous rocks forming noble metal rich fluid. It is possible to form industrial multiple-element- ore-deposits, especially hydrothermal type platinum-group-element-ore-deposits in the region with strong hydrothermal action.

  15. Source tracing of noble metal elements in Lower Cambrian black rock series of Guizhou-Hunan Provinces, China

    Institute of Scientific and Technical Information of China (English)

    李胜荣; 高振敏

    2000-01-01

    The Lower Cambrian black rock series of South China is abnormally rich in noble metal elements. According to the concentrations, the ratios, the relations, the distribution and partition patterns of noble metal elements, the authors think that the noble metals and other elements are neither directly from extraterrestrial materials, nor from the products of normal marine sedimentation. The abnormal enrichment of noble metal elements is closely related with hydrothermal fluid that flew out on the sea floor through deep cycling and reaction with Proterozoic ultramafic-mafic igneous rocks forming noble metal rich fluid. It is possible to form industrial multiple-element-ore-deposits, especially hydrothermal type platinum-group-element-ore-deposits in the region with strong hydrothermal action.

  16. Chengjiang fauna & the Cambrian taxonomic explosion%澄江动物群与寒武纪大爆发

    Institute of Scientific and Technical Information of China (English)

    赵生才

    1998-01-01

    @@ 1984年,中国学者在云南澄江寒武纪地层发现一批丰富多彩的古化石群,迅速在世界上引起强烈反响,美国据世界一些著名科学家联合建议,将此列入本世纪最惊人的发现之一.德国古生物学家赛拉赫(Seilacher)等著名学者认为,"寒武纪大爆发(Cambrian Explosion)"是生命历史中最伟大、但也是了解最少的一次生物事件,而中国澄江动物群拥有解开这个谜的线索.

  17. A New Ichnogenus Kailidiscus Produced by Attachment from the Middle Cambrian Kaili Formation in Taijiang, Guizhou Province, China

    Institute of Scientific and Technical Information of China (English)

    2006-01-01

    Over 190 specimens of an attachment ichnofossil Kailidiscus (n. ichnogen.) have been found in the Kaili Bitoa, a Burgess Shale-type biota, from the middle Kaili Formation (early Middle Cambrian) in Taijiang County, Guizhou Province, southern China. Kailidiscus is an epichnia with a circular to oval attachment platform, on which there is a carbonaceous film probably formed by the body of the living organism. Structures such as a marginal furrow, an inner and outer marginal ride, many convex fold-ridges, and a large caved peripheral furrow on the attachment platform may be the result of basal tissues of a sessile organism that wrinkled to increase the attachment's surface area. The Kailidiscus organism may have been a sessile cnidarian attached to the muddy seafloor. Kailidiscus organisms lived in relatively quiet water,and were buried by a sudden influx of sediment.

  18. Phosphatized Punctatus with mouthparts and its embryo fossils from the Lower Cambrian of Ningqiang, south Shaanxi, China

    Institute of Scientific and Technical Information of China (English)

    LIU Yunhuan; LI Yong; SHAO Tiequan; ZHU Zhixin; YU Bo; WANG Zhuo; ZHANG Wanqian; LI Rongxi

    2007-01-01

    The appearance of coelenterates marks the real beginning ofmetazoan evolution. It therefore has a prominent position in the origin and evolutionary history of organisms,and is also a pivotal question of evolutionary biology.Punctatus is an extinct, ancient marine animal from the early stage of the Cambrian explosion, occurring at. The lowermost Cambrian of both Kuanchuanpu (Ningqiang, Shaanxi) and Maidiping (Emei, Sichuan) areas. Punctatus has been studied for many years since the discovery of its fragments. System-atic and phylogenetic analysis has long been limited because of the rarity of complete specimens. In order to improve research into Punctatus, more than ten thousand globular fossils were recovered by means of "Chemistry Retting". On the basis of the study of these globular fossils, a series of Punctatus fossils with cone parts and finely preserved soft-tissue mouthparts and fossilized metazoan embryo were recovered. Through research on characteristics such as shape,modality and structure of these fossils symbiotic with Punctatus emeiensis, the author found many possible embryo fossils including the evidence of gastrula-stage animal fossils.The sequence of fetation might have appeared on the corol-liform oral region of Punctatus emeiensis. A study of the soft tissues, functional morphology and the sequence of embryo fossils shows evidence that Punctatus resembles coelenterate polyps in systematic classification. Perhaps it also represents an ancestor of the coelenterate with an original tentacle. A comparison with the real "tentacle animal" found in the Chengjiang Fauna, shows that the original tentacle is very tiny and its function range is limited. This reveals the primi-tive nature of the animal. Although the original tentacle is so small, it does exist, representing the first big step towards the real flexible tentacle with a strong function from the early evolving tentacle.

  19. Giant submarine landslide grooves in the Neoproterozoic/Lower Cambrian Phe Formation, northwest Himalaya: Mechanisms of formation and palaeogeographic implications

    Science.gov (United States)

    Draganits, E.; Schlaf, J.; Grasemann, B.; Argles, T.

    2008-04-01

    Giant groove casts have been found in the upper Proterozoic to Lower Cambrian Phe Formation (Haimanta Group), a siliciclastic sandstone/shale succession in the Tethyan Zone of the Higher Himalaya tectonic unit. The grooves are among the largest linear erosion structures related to submarine mass-movements observed in the geologic record. They are up to 4 m wide, about 0.2 m deep and can be traced for more than 35 m without changing their character. The grooves are straight, subparallel to cross-cutting striations with shallow semi-circular cross-sections and well-defined superimposed minor ridges and grooves. Groove casts exist on the soles of several sandstone beds within a 73 m thick logged section, commonly associated with flute casts. Their characteristics were compared with several other types of ancient and modern submarine linear erosion structures. A sand-rich, non-channelized basin floor depositional environment is inferred from the lithofacies, the combination of sedimentary structures, the lack of coarse-grained pebbly facies, the lateral continuity of beds, and the lack of channel structures. The grooves probably formed by laminar debris flows/concentrated density flows dragging blocks of already lithified sediment across the basin floor. When the bedding is structurally rotated back to horizontal, the groove casts show consistent North-South oriented palaeocurrent trends, with South-directed palaeocurrent directions indicated by flute casts. These palaeocurrent orientations contrast with previous palaeogeographic reconstructions of this area, which propose sediment delivery from the South. We therefore suggest a new "double provenance" model for the spatial relationship of late Proterozoic to Early Cambrian strata of the Himalaya, in which Lesser and Tethyan Himalayan age-equivalent sediment was deposited in a connected basin, where the former received detritus from the South, and the latter from a hitherto unknown source in the North. One possible

  20. The noble gas isotope geochemical composition of chert at the bottom of Cambrian in Tarim Basin,China

    Institute of Scientific and Technical Information of China (English)

    2009-01-01

    In the Tarim Basin,black shale series at the bottom of Cambrian is one of the important marine facies hydrocarbon source rocks. This research focuses on the analysis of the isotope of noble gas of 11 cherts. The R/Ra ratio of chert in the Keping area is 0.032―0.319,and 40Ar/36Ar is 338―430. In Quruqtagh the R/Ra ratio is 0.44―10.21,and 40Ar/36Ar is 360―765. The R/Ra ratio of chert increases with 40Ar/36Ar from the west to the east accordingly. They have evolved from the crust source area to the mantle source area in a direct proportion. Surplus argon 40ArE in chert is in direct proportion to the R/Ra ratio,indicating that it has the same origin of excess argon as in fluid and mantle source helium. Comparison of the R/Ra ratios between the west and the east shows that the chert in the eastern part formed from the activity system of the bottom hydrothermal venting driven by the mantle source,where the material and energy of crust and mantle had a strong interaction in exchange;whereas in the western part,chert deposited from the floating of hydrothermal plume undersea bottom,which is far away from the centre of activities of the hydrothermal fluid of ocean bottom. In addition,from noble gas isotope composition of chert,it is suggested that the ocean anoxia incident happened at the black shale of the Cambrian bottom probably because of the large-scaled ocean volcanoes and the followinghydrothermal activities.

  1. The geological significance of the boundary between the Fort Sill and Signal Mountain Formations in the lower Arbuckle Group (Cambrian)

    Energy Technology Data Exchange (ETDEWEB)

    Hosey, R.; Donovan, R.N. (Texas Christian Univ., Ft Worth, TX (United States). Geology Dept.)

    1993-02-01

    During the upper Cambrian, a transgression inundated the Southern Oklahoma aulacogen enveloping a landscape that consisted of hills of Cambrian-aged rhyolite up to 350 m in height. Initial deposits on this topography--the Reagan Formation--consist of siliciclastics that were deposited as alluvium and succeeding tidally-influenced marine sandstones and shales. The siliciclastics grains are made up of local rhyolite, quartz and authigenic glauconite. The overlying Honeycreek Formation is defined by the addition of carbonated detritus in the form of tidally-influenced pelmatozoan grainstones. The passage from the Honeycreek to the overlying Fort Sill Formation of the Arbuckle Group is marked by the incoming of beds of lime mudstone and the gradual disappearance of grainstones and siliciclastics. The contact between the Fort Sill and the overlying thinly-bedded dark grey bioclastic limestones of the Signal Mountain Formation is one of the most distinctive horizons in the Arbuckle Group. The contact evidently marks a substantial change in depositional environment. In detail the contact is sharp and shows evidence of minor erosion, although no karsting has been detected. The authors suggest that the contact surface records a regression, perhaps associated with dolomitization and followed by some erosion. A regression is also indicated by the local occurrence of a laminated tidal flat unit with traces of evaporites that outcrops in the far west of the Slick Hills immediately below the formation contact. They suggest that the Signal Mountains as a transgressive unit, incorporating siliciclastics transported into the area during the regression. It has been suggested that the unconformity reflects localized tectonism associated with the evolution of the Southern Oklahoma aulacogen. On the other hand the surface may correlate with a craton--wide Sauxian' hiatus.

  2. Reassessing the Source of Long-Period Comets

    CERN Document Server

    Kaib, Nathan A; 10.1126/science.1172676

    2009-01-01

    We present numerical simulations to model the production of observable long-period comets (LPCs) from the Oort Cloud, a vast reservoir of icy bodies surrounding the Sun. We show that inner Oort Cloud objects can penetrate Jupiter's orbit via a largely unexplored dynamical pathway, and they are an important, if not the dominant, source of known LPCs. We use this LPC production to place observationally motivated constraints on the population and mass of the inner Oort Cloud, which are consistent with giant planet formation theory. These constraints indicate that only one comet shower producing late Eocene bombardment levels has likely occurred since the Cambrian Explosion, making these phenomena an improbable cause of additional extinction events.

  3. Reassessing the Source of Long-Period Comets

    Science.gov (United States)

    Kaib, Nathan A.; Quinn, Thomas

    2009-09-01

    We present numerical simulations to model the production of observable long-period comets (LPCs) from the Oort Cloud, a vast reservoir of icy bodies surrounding the Sun. We show that inner Oort Cloud objects can penetrate Jupiter’s orbit via a largely unexplored dynamical pathway, and they are an important, if not the dominant, source of known LPCs. We use this LPC production to place observationally motivated constraints on the population and mass of the inner Oort Cloud, which are consistent with giant planet formation theory. These constraints indicate that only one comet shower producing late Eocene bombardment levels has likely occurred since the Cambrian Explosion, making these phenomena an improbable cause of additional extinction events.

  4. Proposed GSSP for the base of Cambrian Stage 9, coinciding with the first appearance of Agnostotes orientalis, at Duibian, Zhejiang, China

    Institute of Scientific and Technical Information of China (English)

    Loren; E.; BABCOCK; Gabriella; BAGNOLI

    2009-01-01

    Exposed in natural outcrops near the Duibian Village, Jiangshan County, Zhejiang Province, China, the Duibian B section is proposed as the boundary stratotype for the base of an unnamed stage termed provisionally Cambrian Stage 9. The proposed position of the GSSP is 108.12 m above the base of the Huayansi Formation, at a horizon coinciding with the first appearance of the cosmopolitan agnostoid trilobite Agnostotes orientalis. This horizon coincides also with the first appearance of the cosmopoli-tan polymerid trilobite Irvingella angustilimbata. The section fulfills all the requirements for a GSSP, and the horizon can be constrained not only by the primary stratigraphic marker (A. orientalis) but also with secondary biostratigraphic, chemostratigraphic, and sequence-stratigraphic correlation tools. The first appearance of A. orientalis is one of the most readily recognizable levels in Cambrian, and can be correlated with precision to all paleocontinents.

  5. Contribution of Oxygenic Photosynthesis to Palaeo-Oceanic Organic Carbon Sink Fluxes in Early Cambrian Upper Yangtze Shallow Sea:Evidence from Black Shale Record

    Institute of Scientific and Technical Information of China (English)

    Kunyu Wu; Tingshan Zhang; Yang Yang; Yuchuan Sun; Daoxian Yuan

    2016-01-01

    ABSTRACT:The extensive transgression that occurred on the Yangtze Plate in Early Cambrian led to a massive organic carbon pool in the Niutitang Formation. A black shale core section from 3 251.08 to 3 436.08 m beneath the Earth’s surface was studied to estimate the contribution of oxygenic photosyn-thesis to organic carbon sink fluxes in Early Cambrian Upper Yangtze shallow sea. Results indicate that the oxygenic photosynthesis played the most important role in carbon fixation in Early Cambrian. Or-ganic carbon sink was mainly contributed by photosynthetic microorganisms, e.g., cyanobacteria, algae and archaea. The Niutitang Formation was formed in a deep anoxic marine shelf sedimentary envi-ronment at a sedimentation rate of ~0.09±0.03 mm/yr. The initial TOC abundance in Niutitang shale ranged from 0.18%to 7.09%, with an average of 2.15%. In accordance with the sedimentation rate and initial TOC abundance, organic carbon sink fluxes were calculated and found to range from 0.21 to 8.10×103 kg/km2·yr-1, especially the organic carbon sink fluxes in depth between 3 385 and 3 470 m range from 3.80 to 8.10×103 kg/km2·yr-1, with an average of~6.03×103 kg/km2·yr-1, which is much high-er than that of contemporary marine sediments. The organic carbon sink fluxes of Niutitang shale are equal to 0.56 to 21.61×103 kg/km2·yr-1 net oxygen emitted into the Early Cambrian ocean and atmos-phere, this emitted oxygen may have significantly promoted the oxygen level of the Earth’s surface and diversification of metazoans.

  6. A Protocol for Diagnosing the Effect of Calibration Priors on Posterior Time Estimates: A Case Study for the Cambrian Explosion of Animal Phyla.

    Science.gov (United States)

    Battistuzzi, Fabia U; Billing-Ross, Paul; Murillo, Oscar; Filipski, Alan; Kumar, Sudhir

    2015-07-01

    We present a procedure to test the effect of calibration priors on estimated times, which applies a recently developed calibration-free approach (RelTime) method that produces relative divergence times for all nodes in the tree. We illustrate this protocol by applying it to a timetree of metazoan diversification (Erwin DH, Laflamme M, Tweedt SM, Sperling EA, Pisani D, Peterson KJ. 2011. The Cambrian conundrum: early divergence and later ecological success in the early history of animals. Science 334:1091-1097.), which placed the divergence of animal phyla close to the time of the Cambrian explosion inferred from the fossil record. These analyses revealed that the two maximum-only calibration priors in the pre-Cambrian are the primary determinants of the young divergence times among animal phyla in this study. In fact, these two maximum-only calibrations produce divergence times that severely violate minimum boundaries of almost all of the other 22 calibration constraints. The use of these 22 calibrations produces dates for metazoan divergences that are hundreds of millions of years earlier in the Proterozoic. Our results encourage the use of calibration-free approaches to identify most influential calibration constraints and to evaluate their impact in order to achieve biologically robust interpretations. PMID:25808541

  7. Pb-Pb isotope dating of black shales from the Lower Cambrian Niutitang Formation, Guizhou Province, South China

    Institute of Scientific and Technical Information of China (English)

    CHEN Yongquan; JIANG Shaoyong; LING Hongfei; FENG Hongzhen; YANG Jinghong; CHEN Jianhua

    2003-01-01

    The Lower Cambrian black shales of the Niutitang Formation and their lateral equivalents occur widely on the Yangtze Platform, South China. In this study, we examine two sections of the Niutitang Formation in Guizhou Province. We employ a stepwise acid-leaching technique to obtain Pb-isotope compositions of the black shales. At the Zhongnan section, the black shales display large Pb isotope variations, with 206Pb/204Pb of 20.5396~33.5709, and 207Pb/204Pb of 15.8407~16.5782. In contrast, Pb isotope data show a relatively limited range in 206Pb/204Pb (18.2797~18.5972) and 207Pb/204Pb (15.6813~15.7159) at the Zhijin section. Data for all acid-leached black shale samples from the two sections yield a Pb-Pb isochron age of 531±24 Ma. This age may represent the sedimentary age of the black shales. The initial Pb-isotope composition of the black shales falls near the upper-crust evolution curve in a Zartman Pb-Pb evolution diagram. Hence, it is likely that the Pb in the black shales may have been derived from an upper continental crustal source.

  8. Morphology and function in the Cambrian Burgess Shale megacheiran arthropod Leanchoilia superlata and the application of a descriptive matrix

    Directory of Open Access Journals (Sweden)

    Haug Joachim T

    2012-08-01

    Full Text Available Abstract Background Leanchoilia superlata is one of the best known arthropods from the middle Cambrian Burgess Shale of British Columbia. Here we re-describe the morphology of L. superlata and discuss its possible autecology. The re-description follows a standardized scheme, the descriptive matrix approach, designed to provide a template for descriptions of other megacheiran species. Results Our findings differ in several respects from previous interpretations. Examples include a more slender body; a possible hypostome; a small specialised second appendage, bringing the number of pairs of head appendages to four; a further sub-division of the great appendage, making it more similar to that of other megacheirans; and a complex joint of the exopod reflecting the arthropod’s swimming capabilities. Conclusions Different aspects of the morphology, for example, the morphology of the great appendage and the presence of a basipod with strong median armature on the biramous appendages indicate that L. superlata was an active and agile necto-benthic predator (not a scavenger or deposit feeder as previously interpreted.

  9. Exceptionally preserved Cambrian trilobite digestive system revealed in 3D by synchrotron-radiation X-ray tomographic microscopy.

    Directory of Open Access Journals (Sweden)

    Mats E Eriksson

    Full Text Available The Cambrian 'Orsten' fauna comprises exceptionally preserved and phosphatised microscopic arthropods. The external morphology of these fossils is well known, but their internal soft-tissue anatomy has remained virtually unknown. Here, we report the first non-biomineralised tissues from a juvenile polymerid trilobite, represented by digestive structures, glands, and connective strands harboured in a hypostome from the Swedish 'Orsten' fauna. Synchrotron-radiation X-ray tomographic microscopy enabled three-dimensional internal recordings at sub-micrometre resolution. The specimen provides the first unambiguous evidence for a J-shaped anterior gut and the presence of a crop with a constricted alimentary tract in the Trilobita. Moreover, the gut is Y-shaped in cross section, probably due to a collapsed lumen of that shape, another feature which has not previously been observed in trilobites. The combination of anatomical features suggests that the trilobite hypostome is functionally analogous to the labrum of euarthropods and that it was a sophisticated element closely integrated with the digestive system. This study also briefly addresses the preservational bias of the 'Orsten' fauna, particularly the near-absence of polymerid trilobites, and the taphonomy of the soft-tissue-harbouring hypostome.

  10. New observations of the lobopod-like worm Facivermis from the Early Cambrian Chengjiang Lagerst(a)tte

    Institute of Scientific and Technical Information of China (English)

    LIU Jianni; HAN Jian; SIMONETTA A. M.; HU Shixue; ZHANG Zhifei; YAO Yang; SHU Degan

    2006-01-01

    Facivermis yunnanicus (Hou & Chen, 1989), from the Early Cambrian Chengjiang Lagerst(a)tte, a worm-like fossil with 5 pairs of tentacles and a perceived shrunken end, has been regarded as related to polychaetes, later it has been variously interpreted as lobopods, Pentastoma and lophophorates. Newly discovered complete specimens by the ELI field team show that the taxon has, in addition to the 5 pairs of appendages, a pear-shaped trunk end bearing two or three circles of hooks. Accordingly, based on these important morphological characters, reconsideration of its affinities is provided and the taxonomy is remedied herein. Because the five pairs of appendages of Facivermis yunnanicus resemble the appendages of the fore-trunk of lobopod Miraluolishania (Liu & Shu, 2004), it seems that they are homologous structures. Therefore, the affinities of Facivermis are regarded here as being close to the lobopods, in addition, the lobe-like appendages of Facivermis are very crucial to exploring the origin of the appendages of lobopods and arthropods.

  11. The Guarguaraz Complex and the Neoproterozoic-Cambrian evolution of southwestern Gondwana: Geochemical signatures and geochronological constraints

    Science.gov (United States)

    López de Azarevich, Vanina L.; Escayola, Mónica; Azarevich, Miguel B.; Pimentel, Márcio M.; Tassinari, Colombo

    2009-12-01

    The Guarguaraz Complex, in western Argentina, comprises a metasedimentary assemblage, associated with mafic sills and ultramafic bodies intruded by basaltic dikes, which are interpreted as Ordovician dismembered ophiolites. Two kinds of dikes are recognized, a group associated with the metasediments and the other ophiolite-related. Both have N-MORB signatures, with ɛNd between +3.5 and +8.2, indicating a depleted source, and Grenville model ages between 0.99 and 1.62 Ga. A whole-rock Sm-Nd isochron yielded an age of 655 ± 76 Ma for these mafic rocks, which is compatible with cianobacteria and acritarchae recognized in the clastic metasedimentary platform sequences, that indicate a Neoproterozoic (Vendian)-Cambrian age of deposition. The Guarguaraz metasedimentary-ophiolitic complex represents, therefore, a remnant of an oceanic basin developed to the west of the Grenville-aged Cuyania terrane during the Neoproterozoic. The southernmost extension of these metasedimentary sequences in Cordón del Portillo might represent part of this platform and not fragments of the Chilenia terrane. An extensional event related to the fragmentation of Rodinia is represented by the mafic and ultramafic rocks. The Devonian docking of Chilenia emplaced remnants of ocean floor and slices of the Cuyania terrane (Las Yaretas Gneisses) in tectonic contact with the Neoproterozoic metasediments, marking the Devonian western border of Gondwana.

  12. Characteristics of sulfur species and their implications in Lower Cambrian black shales from southern margin of Yangtze Platform

    Institute of Scientific and Technical Information of China (English)

    2001-01-01

    There are different sulfur forms in the black shales from theEarly Cambrian of the Yangtze platform. With its emphasis on pyrite and organosulfur, this paper discusses their distribution and formation. The research shows that sulfur phases take regular variations laterally as well as vertically in the research areas. In western researched profile with high terrigenous supply at the time it formed, there exists a larger amount of pyrite and less organosulfur, and pyrite amount declines while organosulfur content increases upwards along the profile. This black shale profile is characterized by relatively light sulfur isotope composition with evolution trend of becoming heavier both for pyrite and organosulfur from bottom to top along the profile. Opposite situation occurs in eastern profiles which were located farther away from terrigenous land. Here pyrite amount obviously decreases and organic matter has combined more sulfur, although these two kinds of sulfur species take similar trend in content variation along profiles to that for western profile. At the same time more 34S is accumulated in sulfur species of black shale samples from eastern profile, and sulfur isotope composition gradually turns lighter from bottom to top. In combination with other information of iron, organic carbon contents and petrographic features, it can be established that sea-level change, supply of terrigenous matters, tectonic background and natures of paleoceanic chemistry have exerted great influence on the distribution of sulfur species in these black shales.

  13. A submarine welded ignimbrite-crystal-rich sandstone facies association in the Cambrian Tyndall Group, western Tasmania, Australia

    Science.gov (United States)

    White, Matthew J.; McPhie, Jocelyn

    1997-04-01

    Three occurrences of rhyolitic welded ignimbrite are intercalated within a submarine, below-storm-wave-base sedimentary succession in the Cambrian Tyndall Group, Mount Read Volcanics, western Tasmania. These occurrences are closely associated with very thick crystal-rich sandstone facies that is present at this stratigraphic level throughout the Tyndall Group. This facies is interpreted to comprise deposits from syn-eruptive, crystal-rich, submarine sediment gravity flows that were generated by interaction of subaerial pyroclastic flows with seawater. Removal of fine ash and pumice from the submarine flows by hydraulic sorting and flotation resulted in marked crystal enrichment in the deposits. Rapid, essentially syn-eruptive aggradation of crystal-rich sand led to temporary shoaling so that in some cases, subsequent pyroclastic flows deposited welded ignimbrite in shallow marine or possibly subaerial settings (e.g., Zig Zag Hill welded ignimbrite). Breccia units composed of welded ignimbrite clasts and crystal-rich matrix (e.g., Comstock and Anthony Road ignimbrite breccias) imply that some welded ignimbrite was submerged, providing clasts to syn-eruptive, submarine, crystal-rich sediment gravity flows. One example of welded ignimbrite (Cradle Mountain Link Road) may have been deposited in an entirely below-storm-wave-base environment. The distinctive facies association of welded ignimbrite, crystal-rich sandstone and ignimbrite-clast breccia in the Tyndall Group exemplifies the submarine record of a major rhyolitic explosive eruption in the source volcanic terrane.

  14. The earliest-known ancestors of Recent Priapulomorpha from the Early Cambrian Chengjiang Lagerst(``a)tte

    Institute of Scientific and Technical Information of China (English)

    HAN Jian; SHU Degan; ZHANG Zhifei; LIU Jianni

    2004-01-01

    The taxonomy of an early ancestor of Recent Priapulidae, Xiaoheiqingella peculiaris (= Yunnanpriapulus halteroformis Huang et al., 2004) from the Early Cambrian Chengjiang fossil LagerstStte, is revised. Morphological characters comprise a pair of caudal appendages rather than a single appendage flanking the trunk end and a possible urogenital duct found inside the preanal region. An additional extremely rare fossil priapulid worm, Paratubiluchus bicaudatus gen. nov., sp. nov. is also described herein. Its diagnostic characters are: an introvert bearing 25 longitudinal rows of scalids, a distinct neck region, no annulus on the oval trunk, and a pair of caudal appendages. The proportion of body parts is similar in size to that of loricate larvae of Recent priapulids and larva-formed Palaeopriapulitidae. Taking account of the features of Xiaoheiqingella, bicaudal appendages are considered to be a synapomorphy of Priapulidae and Tubiluchidae. Paratubiluchus gen. nov. is most likely a candidate for the ancestor of the Tubiluchidae; it probably originated from a larva-formed priapulid with 25 rows of scalids, thus representing an intermediate link between the priapulids in mature-form and the priapulids with Iorica.

  15. Reconstruction of Cambrian Global Paleo-plates and Paleogeography%寒武纪全球板块构造与古地理环境再造

    Institute of Scientific and Technical Information of China (English)

    周肖贝; 李江海; 王洪浩; 李文山

    2014-01-01

    基于古地磁数据对寒武纪全球古板块进行再造,并叠加更新的寒武纪全球大地构造背景和岩相古地理分布等要素,编制了寒武纪全球古板块再造图、古地理图、全球岩相及烃源岩分布图。寒武纪全球板块以冈瓦纳超大陆、劳伦古陆、西伯利亚板块和波罗的板块为主,多集中位于南半球,冈瓦纳超大陆快速向南漂移,其他板块主要向北漂移,并伴随逆时针旋转。寒武纪整体处于海平面及温度的上升时期,全球板块多为陆表海环境,有利于早寒武世的富有机质沉积,为寒武纪生命大爆发提供了良好条件。但此时气候带梯度不高,干旱气候带广泛分布,下要中寒武统多蒸发盐岩。冈瓦纳超大陆北部主要为碳酸盐岩沉积,南部多为砂岩沉积,东部边缘多火山活动,在中部的西侧边缘地区碳酸盐岩沉积和砂岩沉积物中一般有较好的油气和烃源岩。劳伦古陆沉积具有环带特征,其外侧为碳酸盐岩沉积。西伯利亚板块早要中寒武世主要以巨厚的膏盐岩沉积为主,晚期沉积仅为几百米厚的碳酸盐岩,整体处于被动陆缘沉积的构造背景,这促进了该板块中储层和盖层的发育。波罗的板块纬度相对偏高,主要为砂岩和页岩沉积。%The global maps of Cambrian plate tectonics, plaeogeography, lithofaices and source rock distribution are compiled up for reconstructing Cambrian global paleo-plates on the basis of the paleomagnetic data and the updating data of the Cambrian global tectonics, lithofacies paleogeography and source rocks. Cambrian plates can be divided into four parts including Gondwana, Laurentia, Siberia and Baltica. They were generally distributed in the southern hemisphere and drifted northwards with contrarotating rotation except the southwards drifting Gondwana supercontinent. Owing to rising of global sea level eustacy and temperature for a long time

  16. Geochemical characteristics of organic matter-rich strata of lower Cambrian in Tarim Basin and its origin

    Institute of Scientific and Technical Information of China (English)

    CHEN; Jianfa; SUN; Shengli; LIU; Wenhui; ZHENG; Jianjing

    2004-01-01

    The organic matter-rich strata of the Yurtusi Formation of Lower Cambrian are wildly spread and steady in the Tarim Basin, in which cherts are developed, companied with phosphorite. Al/(Al+Fe+Mn) ratio and Si/(Si+Al+Fe) ratio of cherts range from 0.0023 to 0.0046 and 0.965 to 0.98, respectively, suggesting that cherts are formed in submarine hydrothermal activity and far away from terrestrial. Trace elements, such as As, Hg, Pb, Zn, Cu, Co, P, V, Ba, etc., are markedly rich in organic matter-rich sedimentary rocks of which Ba/Sr ratios are between 21.2 and 158.1. Compared with modern hydrothermal sediments, their Ba/Sr ratios are similar, it indicates that the organic matter-rich strata have the geochemical feature of submarine hydrothermal sediments. The total content of rare earth elements (ЕREE) in cherts is from 8.81 to 56.682 μg/g, on average 31.41 μg/g and the ΕREE of cherts is between continental margin chert's and abyssal chert's. The LREE/HREE ratio of cherts varies from 1.01 to 3.56, which reveals the characteristics of hydrothermal sediments. There is positive correlation between the total organic content (TOC) and Ba/Sr ratio which is geochemical index of submarine hydrothermal activity, which indicates that the submarine hydrothermal activity obviously influences the richness of organic matter in the Yurtusi Formation.

  17. Maps showing petroleum exploration intensity and production in major Cambrian to Ordovician reservoir rocks in the Anadarko Basin

    Science.gov (United States)

    Henry, Mitch; Hester, Tim

    1996-01-01

    The Anadarko basin is a large, deep, two-stage Paleozoic basin (Feinstein, 1981) that is petroleum rich and generally well explored. The Anadarko basin province, a geogrphic area used here mostly for the convenience of mapping and data management, is defined by political boundaries that include the Anadarko basin proper. The boundaries of the province are identical to those used by the U.S. Geological Survey (USGS) in the 1995 National Assessment of United Stated Oil and Gas Resources. The data in this report, also identical to those used in the national assessment, are from several computerized data bases including Nehring Research Group (NRG) Associates Inc., Significant Oil and Gas Fields of the United States (1992); Petroleum Information (PI), Inc., Well History Control System (1991); and Petroleum Information (PI), Inc., Petro-ROM: Production data on CD-ROM (1993). Although generated mostly in response to the national assessment, the data presented here arc grouped differently and arc displayed and described in greater detail. In addition, the stratigraphic sequences discussed may not necessarily correlate with the "plays" of the 1995 national assessment. This report uses computer-generated maps to show drilling intensity, producing wells, major fields, and other geologic information relevant to petroleum exploration and production in the lower Paleozoic part of the Anadarko basin province as defined for the U.S. Geological Survey's 1995 national petroleum assessment. Hydrocarbon accumulations must meet a minimum standard of 1 million barrels of oil (MMBO) or 6 billion cubic feet of gas (BCFG) estimated ultimate recovery to be included in this report as a major field or revoir. Mapped strata in this report include the Upper Cambrian to Lower Ordovician Arbuckle and Low Ordovician Ellenburger Groups, the Middle Ordovician Simpson Group, and the Middle to Upper Ordovician Viola Group.

  18. A transgression-regression event during the deposition of the Upper Cambrian Honey Creek formation in the southern Oklahoma aulacogen

    Energy Technology Data Exchange (ETDEWEB)

    McElmoyl, C.; Donovan, R.N. (Texas Christian Univ., Ft. Worth, TX (United States). Geology Dept.)

    1993-02-01

    The transgression that inundated the Southern Oklahoma aulacogen during the upper Cambrian enveloped a landscape that consisted of hills of rhyolite up to 350 m in high. Initial deposits on this topography have been interpreted as alluvium. These, together with succeeding tidally-influenced marine siliciclastics form the Reagan Formation. The siliciclastics grains are made up of fragments of local origin (i.e., rhyolite), quartz (derived from a distal source) and authigenic glauconite. The upward passage from the Reagan to the Honeycreek Formation is defined by the addition to the siliciclastics of carbonate detritus in the form of tidally-influenced grainstones, mostly composed of pelmatozoan fragments. The passage from the Honeycreek to the overlying Fort Sill Formation of the Arbuckle Group is marked by the incoming of beds of lime mudstone and the gradual disappearance of grainstones and siliciclastics. Evidence of the existence of rhyolite topography (i.e., an archipelago) can be detected to within 50 m of the top of the Fort Sill. While the overall facies pattern undoubtedly records a widespread transgression, a newly-discovered slightly angular unconformity within the lower part of the Honeycreek is best interpreted as a record of a temporary regression. Three distinctive lithologies are involved in this relationship: the lowest beds are light grey cross-bedded pelmatozoan grainstones with minor amounts of quartz and rhyolite grains. Syntaxial cements at the base of this unit are homogenous under cathode luminescence, while cements near the top display up to 27 zones of reflectance, interpreted as a fluctuating marine-meteoric groundwater imprint. The overlying bed is a red-brown mud-supported limestone that contains abundant angular rhyolite pebbles and a rich trilobite fauna. Some of the pebbles are coated by pelmatozoans.

  19. Cambrian Magmatism Along the Southern Laurentian Margin: New Petrologic Constraints from Well Cuttings in the Southern Oklahoma Aulacogen (USA)

    Science.gov (United States)

    Hobbs, J. M.; Brueseke, M. E.; Mertzman, S. A.; Puckett, R. E., Jr.

    2014-12-01

    Magmatism in the Southern Oklahoma aulacogen (SOA) is associated with >250,000 km3 of early Cambrian aged mafic-silicic intrusives and effusive products possibly related to the opening of the Iapetus Ocean. In the vicinity of the Arbuckle Mts., thick mafic to intermediate lava packages interbedded with rhyolite lavas and sedimentary strata are exposed in the subsurface and penetrated by oil and gas exploration wells. We have sampled cuttings from six wells exposed along the strike of the SOA with the aim of petrographically, geochemically, and isotopically characterizing the rocks to better understand their tectonomagmatic affinity. Cuttings were observed under a binocular microscope to ensure textural homogeneity; XRF bulk chemistry from these samples show that they are relatively unaltered based on analytical totals and alteration indices. The samples plot as basalts to andesites (e.g., 47-64 wt. % SiO2) and are primarily tholeiitic. On discrimination diagrams, these samples fall in "intraplate" fields, consistent with continental basalt volcanism, including flood basalt eruptions. These lavas show Zr/Nb values ranging from 6.8 to 11.1, K/Nb values ranging from 300-600, and Ba/Nb values ranging from 10-20, which are similar to EM1 OIB. The samples also show trace element patterns consistent with OIB-like mantle sources when normalized to primitive mantle, and similar geochemical traits to the Roosevelt Gabbros that crop out along strike of the SOA in the Wichita Mts.. Sr, Nd, and Pb isotope analysis is ongoing. Chemostratigraphic variations show the possibility of several lava packages. The geochemical analyses of the SOA lava flows provide insight into how these magmas formed as well as what tectonic regime (e.g., lower-mantle derived plume, upper-mantle extension, or "leaky" transform fault) produced the volcanism that affected the southern margin of Laurentia during the formation of the SOA; we favor the involvement of a deep mantle plume.

  20. The origin of increased salinity in the Cambrian-Vendian aquifer system on the Kopli Peninsula, northern Estonia

    Science.gov (United States)

    Karro, Enn; Marandi, Andres; Vaikmäe, Rein

    Monitoring of the confined Cambrian-Vendian aquifer system utilised for industrial water supply at Kopli Peninsula in Tallinn over 24 years reveals remarkable changes in chemical composition of groundwater. A relatively fast 1.5 to 3.0-fold increase in TDS and in concentrations of major ions in ed groundwater is the consequence of heavy pumping. The main sources of dissolved load in Cambrian-Vendian groundwater are the leaching of host rock and the other geochemical processes that occur in the saturated zone. Underlying crystalline basement, which comprises saline groundwater in its upper weathered and fissured portion, and which is hydraulically connected with the overlying Cambrian-Vendian aquifer system, is the second important source of ions. The fractured basement and its clayey weathering crust host the Ca-Cl type groundwater, which is characterised by high TDS values (2-20 g/L). Intensive water ion accelerates the exchange of groundwaters and increases the area of influence of pumping. Chemical and isotopic studies of groundwater indicate an increasing contribution of old brackish water from the crystalline basement and rule out the potential implication of an intrusion of seawater into aquifer. L'origine de la salinité croissante dans le système aquifère du Cambrien-Vendien dans la péninsule de Kopli, nord de l'Estonie Le suivi à long terme du système aquifère captif du Cambrien-Vendien utilisé pour l'approvisionnement d'eaux industrielles dans la Péninsule de Kopli, nord de l'Estonie, révèle de remarquables changements dans la composition chimique des eaux souterraines. Une augmentation de facteur 1.5 à 3 de la TDS et des concentrations en ions majeurs dans l'eau souterraine est la conséquence de pompages intensifs. Les sources principales des charges dissoutes dans les eaux de l'aquifère du Cambrien-Vendien sont le lessivage des roches et d'autres phénomènes géochimiques ayant lieu dans la zone saturée. Le soubassement rocheux cristallin

  1. Sulfur and Oxygen Isotopes of Sulfate Extracted from Early Cambrian Phosphorite Nodules:Implications for Marine Redox Evolution in the Yangtze Platform

    Institute of Scientific and Technical Information of China (English)

    Wenlang Qiao; Xianguo Lang; Yongbo Peng; Kaiyuan Jiang; Wu Chen; Kangjun Huang; Bing Shen

    2016-01-01

    ABSTRACT:Phosphorite nodule beds are discovered in the black shale of basal Niutitang Formation throughout the Yangtze Platform in South China, recording an important phosphorite-generation event. Platform-wide phosphorite precipitation requires special oceanographic and geochemical conditions, thus the origin of the Niutitang phosphorite nodules may provide valuable information about the ocean chemistry in the Early Cambrian. In this study, we measured sulfur and oxygen isotopic compositions of sulfate extracted from phosphorite nodules collected from the basal Niutitang Formation. Phosphorite associated sulfate (PAS) is a trace amount of sulfate that incorporates into crystal lattice during phosphorite precipitation, accordingly PAS records the geochemical signals during phosphorite nodule formation. Sulfur isotopic composition of PAS (δ34SPAS) ranges from -1.16‰ to +24.48‰(mean=+8.19‰, n=11), and oxygen isotopic value (δ18OPAS) varies between -5.3‰ and +26.3‰(mean=+7.0‰, n=8). Most phosphorite nodules have low δ34SPAS and low δ18OPAS values, suggesting PAS mainly derived from anaerobic oxidation of H2S within suboxic sediment porewater. We propose that phosphate was delivered to the Yangtze Platform by a series of upwelling events, and was scavenged from seawater with the precipitation of FeOOH. The absorbed phosphate was released into suboxic porewater by the reduction of FeOOH at the oxic-suboxic redox boundary in sediments, and phosphorite nodule precipitated by the reaction of phosphate with Ca2+ diffused from the overlying seawater. The platform-wide deposition of phosphorite nodules in the basal Niutitang Formation implies the bottom water might be suboxic or even oxic, at least sporadically, in Early Cambrian. We speculate that the intensified ocean circulation as evident with frequent occurrences of upwelling events might be the primary reason for the episodic oxidation of the Yangtze Platform in Early Cambrian.

  2. Zn-Pb Ores of Mississippi Valley Type in the Lycksele-Storuman District, Northern Sweden: A Possible Rift-Related Cambrian Mineralisation Event

    Directory of Open Access Journals (Sweden)

    Kjell Billström

    2012-06-01

    Full Text Available The epigenetic Zn-Pb deposits in the Lycksele-Storuman ore district, northern Sweden, are hosted by Paleoproterozoic basement near the margin of the Caledonian mountains. A paleogeographic reconstruction suggests that platform sediments, including Cambrian shales, overlaid the mineralised basement. The mineralisation type, containing sphalerite, galena, calcite and fluorite, is confined to veins and breccias and interpreted to be of Mississippi Valley Type (MVT style. There is no appreciable wall rock alteration. Fluid inclusion work reveals coexisting aqueous and hydrocarbon fluids. Ore deposition is interpreted to have occurred during mixing of two fluids; a cool (

  3. A highly diverse trilobite fauna with Avalonian affinities from the Middle Cambrian Acidusus atavus Zone (Drumian Stage) of Bornholm, Denmark

    DEFF Research Database (Denmark)

    Weidner, Thomas; Nielsen, Arne Thorshoj

    2014-01-01

    A newly collected trilobite fauna from the lowermost part of the Alum Shale Formation at Øleå, Bornholm, Denmark, demonstrates the presence of a thin but richly fossiliferous Middle Cambrian Acidusus atavus Zone (Paradoxides paradoxissimus Superzone). The lower part of the zone (the Tomagnostus...... designation and reillustration of relevant lectotypes. The abundance of polymerid trilobites in comparison with nearby Scania, southern Sweden, where 25 agnostid and nine polymerid species have been reported from equivalent strata, is indicative of a less dysoxic environment in the Bornholm area, which was...

  4. Limitations of metazoan 18S rRNA sequence data : implications for reconstructing a phylogeny of the animal kingdom and inferring the reality of the cambrian explosion

    OpenAIRE

    Abouheif, Ehab; Zardoya, Rafael; Meyer, Axel

    1998-01-01

    We document the phylogenetic behavior of the 18S rRNA molecule in 67 taxa from 28 metazoan phyla and assess the effects of among-site rate variation on reconstructing phylogenies of the animal kingdom. This empirical assessment was undertaken to clarify further the limits of resolution of the 18S rRNA gene as a phylogenetic marker and to address the question of whether 18S rRNA phylogenies can be used as a source of evidence to infer the reality of a Cambrian explosion. A notable degree of am...

  5. Evaluating the primary and/or diagenetic origin of rare earth element abundances in Ediacaran to early Cambrian phosphate deposits, Yangtze Platform (South China) by LA-ICPMS

    Science.gov (United States)

    Hippler, Dorothee; Klügel, Andreas; Biedermann, Nicole; Guo, Qingjun; Franz, Gerhard

    2014-05-01

    The Precambrian-Cambrian time interval represents one of the greatest phosphogenic episodes in Earth's history with giant and well-preserved phosphate deposits occurring on the Yangtze Platform in South China. We investigated concentrations of rare earth elements (REE) and yttrium of shallow and deep-water sedimentary phosphate deposits of the Ediacaran Doushantou Formation and the early Cambrian Zhongyicun Formation by using LA-ICP mass spectrometry. The aim is to examine the temporal and spatial variability of seawater chemistry in conjunction with the conditions of phosphate formation and the evaluation of the extent of diagenetic modification. The mineralogical and textural composition of the samples was pre-screened using SEM and XRD, and polished thick sections were prepared for subsequent high-resolution LA-ICPMS analyses. Overall concentrations in REE range between 18 and 657 ppm, with elevated concentrations (> 200 ppm) in apatite from the deep-water phosphate deposits. REE+Y patterns of shallow-water phosphate deposits exhibit the evolution from flat shale-like to gently inclined seawater-derived patterns, with the early Cambrian phosphate deposits revealing distinct negative Ce- and positive Y-anomalies indicative for oxygenated surface waters. REE+Y patterns of phosphate deposits of the deep-water facies are flat to highly enriched in MREE, which is manifested in variably pronounced concave-down patterns. In detail, these patterns display different Ce-anomalies, as well as small positive Eu-anomalies. We propose that REE+Y patterns of Ediacaran and early Cambrian sedimentary phosphate deposits can inherit both primary and secondary signatures reflecting either seawater composition or diagenetic modification and fluid flow. The combination of imaging techniques and in-situ LA-ICPMS thereby enables a more sophisticated examination of the potential sources and processes than whole rock determinations. Placing the results in stratigraphic order and assuming

  6. Les microbialites de l'Anti-Atlas occidental (Maroc) : marqueurs stratigraphiques et témoins des changements environnementaux au Cambrien inférieur Stratigraphic and environmental significance of the Lower-Cambrian western Anti-Atlasic microbialites (Morocco)

    Science.gov (United States)

    Benssaou, Mohammed; Hamoumi, Naima

    2004-02-01

    Three microbialite forms are recognized in the Lower-Cambrian succession of Irherm area in the western Anti-Atlas (Morocco). Stromatolites, which correspond to non-calcified shallow marine laminated microbialites, are well developed in the basal Lower-Cambrian succession. Occurrence of calcified microbial thrombolites, in the middle part of this succession, reflects an increasing sea level from the peritidal zone to the subtidal environment. In the upper part of this succession, a second increasing water depth event and the development of branching archaeocyathan reefal framework lead to dendritic microbialite emergence. To cite this article: M. Benssaou, N. Hamoumi, C. R. Geoscience 336 (2004).

  7. The Vendian (Ediacaran) in the geological record: Enigmas in geology's prelude to the Cambrian explosion

    Science.gov (United States)

    McCall, G. J. H.

    2006-07-01

    Up to the 1950s, the Precambrian was regarded as unrewardingly unfossiliferous, records of fossils being isolated, few in number and dubious. The change came with the discovery by Reg Sprigg of body fossils in latest Proterozoic sediments in South Australia. Although there had been descriptions of isolated fossils (now recognised as Ediacaran) from rocks of this age in the nineteenth century and at the start of the twentieth century, in Newfoundland and Namibia, respectively, the Ediacara finds stimulated researches and now, at the start of the twenty-first century, diversified fossil assemblages are known, all over the world, from the period 600-543 Ma, known formerly as the Vendian and now officially as the Ediacaran. In this account, a brief description of the history of these finds is given, followed by descriptions of the most important provinces [South Australia, Leicestershire, Namibia, Russia (Podolia, the White Sea Coast, Urals and Siberia), Newfoundland (Avalon Peninsula) and Northwest Canada]: then of 27 other known occurrences of this dominantly soft-bodied and perplexing fauna(?)-it seems certain that some, at least of the fossils, are animal fossils, although some, even the greater part, could be a unique form of life, not animals or plants ("Vendobionta"). These descriptions, derived in the course of a literature search lasting over a year, are followed by discussions of important special aspects: trace fossils; geochronology and correlation; geotectonics; glaciation (the "Snowball Earth" concept applied to the Varangian/Laplandian/Marinoan glaciation, which ushered in this last subdivision of the Proterozoic); the evidence for Ediacaran and other life forms existing in the Proterozoic prior to its last, Ediacaran, chronological subdivison; the Vendozoa concept. The last section consists of short summary of conclusions. This text essentially constitutes an objective record of what has been published to 2005 on the Ediacaran System.

  8. Occurrence of Highly Mature Organic Matter in Marine Black Shale Petroleum Source Rocks of Basal Cambrian from Northern Tarim Basin, China

    Institute of Scientific and Technical Information of China (English)

    YU Bingsong (于炳松); Hailiang DONG; CHEN Jianqiang (陈建强); CHEN Xiaolin (陈晓林); LIANG Shiyou (梁世友)

    2004-01-01

    More and more evidence indicates that organic matter (OM) in immature organic-rich sediments and sedimentary rocks is chemically adsorbed onto the outer surfaces of minerals and into interlayer (inner) surfaces of smectitic clay minerals in the form of amorphous molecular-scale carbon. But there have been few reports about the occurrence of highly mature OM in marine black shales (petroleum source rocks). The occurrence of highly mature OM in the black shales of basal Cambrian from northern Tarim Basin is studied in this paper. Based on the comprehensive analyses of total organic carbon contents (TOC), maximum thermolysis temperatures (T max ) of OM, mineral surface areas (MSA), and scanning electronic microscopic (SEM) and transmission electronic microscopic (TEM) observations of the black shales, it is concluded that the highly mature OM in the marine black shales of the basal Cambrian from northern Tarim Basin occurs in particulates ranging in size from 1 to 5 μm in diameter. Through the contrast of the occurrence of the highly mature OM in the black shales with that of the immature ones in modern marine continental margin sediments, some scientific problems are proposed, which are worth to study further in detail.

  9. An Ediacaran–Cambrian thermal imprint in Rajasthan, western India: Evidence from 40Ar-39Ar geochronology of the Sindreth volcanics

    Indian Academy of Sciences (India)

    Archisman Sen; Kanchan Pande; Hetu C Sheth; Kamal Kant Sharma; Shraboni Sarkar; A M Dayal; Harish Mistry

    2013-12-01

    The Sindreth Group exposed near Sirohi in southern Rajasthan, western India, is a volcanosedimentary sequence. Zircons from Sindreth rhyolite lavas and tuffs have yielded U–Pb crystallization ages of ∼768–761 Ma, suggesting that the Sindreth Group is a part of the Malani magmatic event. Earlier 40Ar-39Ar studies of other Malani volcanic and plutonic rocks yielded disturbed argon release spectra, ascribed to a ∼550 Ma thermal event possibly related to the Pan-African orogeny. To test and confirm this possibility, we dated two whole-rock and three feldspar separate samples of the Sindreth volcanics by the 40Ar-39Ar method. All samples yield disturbed argon release spectra suggesting radiogenic argon loss and with plateau segments at 550 Ma or 490 Ma. We interpret these as events of argon loss at 550–490 Ma related to an Ediacaran–Cambrian thermal event, possibly related to the Malagasy orogeny. The combined older and new 40Ar-39Ar results are significant in showing that whereas Ediacaran–Cambrian magmatic and metamorphic events are well known from many parts of India, they left thermal imprints in much of Trans-Aravalli Rajasthan as well. The overall evidence is consistent with a model of multiphase assembly of Gondwanaland from separate continental landmasses.

  10. Phosphatized rare star-like mouth disc of Punctatus and its functional morphology from the earliest Cambrian of the South Shaanxi China

    Institute of Scientific and Technical Information of China (English)

    2008-01-01

    The Meishucun stage is the prelude in decipher-ing the Cambrian Explosion. In this prominent stage, rapid radioactive evolution and body-plan innovation have taken place and different associations of organism have been shaped. In this paper we report several 3D-preserved rare star-like fossils with finely preserved soft tissues which were recovered from the Kuanchuanpu Member of the Dengying Formation in South Shaanxi, China in 2003. By studying on functional morphology and analogy with mouthpart of Punctatus, there are evidences that this star-like organism approaches the coelenterates in systematic classification and the centre of star-like organism is its mouth. The appearance of coelenterates marks the real beginning of metazoan evolution. Therefore, it has the prominent position in the origin and evolutionary history of organisms. Perhaps the star-like organism represents the early types of coelenterate with original tentacles. These new materials provide new evidence for the origin, evolution and the functional evolution of the metazoan during the early stage of the Cambrian Explosion.

  11. Research progress and prospects of Cambrian medusae from South China%寒武纪水母化石研究现状与展望

    Institute of Scientific and Technical Information of China (English)

    韩健; 李国祥; 久保田信; 王星; 欧强

    2014-01-01

    We introduced the fossil records of medusa in brief and discussed the radial symmetry, body plan, life history, and exoskeleton of ancestral medusae. It is hypothesized that major clades of superclass medusa should have derived prior to that of ballerinas at the dawn of Cambrian explosion. The Cambrian medusae should be classified as sessile benthic and pelagic forms.%简述了水母的早期化石记录,并根据华南寒武系发现的立方水母干群化石,讨论了祖先型水母的辐射对称、躯体构型、生活史、生活习性,以及外骨骼的形成特征,推测水母类在寒武纪大爆发初期先于两侧对称动物基本完成其高级分类单元的宏演化。寒武纪的水母在生活习性上可以分为底栖固着型和游泳型两种。

  12. Elasto-Plastic Constitutive Behavior in Three Lithofacies of the Cambrian Mt. Simon Sandstone, Illinois Basin, USA

    Science.gov (United States)

    Dewers, T.; Newell, P.; Broome, S. T.; Heath, J. E.; Bauer, S. J.

    2012-12-01

    The Mt. Simon Formation, a basal Cambrian sandstone underlying the Illinois Basin in the Central US, is a target for underground storage and waste injection which require an assessment of geomechanical behavior. The range of depositional environments, from braided streams and minor eolean features in the lower Mt Simon, to tidally-influenced near- and on-shore sands in the upper Mt. Simon, yield a heterogeneous formation with a range in porosity, permeability, and mechanical properties. We examine the experimental deformational behavior of three distinct Mt. Simon lithofacies via axisymmetric compressional testing. Initial yielding is confirmed with acoustic emissions in many of the tests and failure envelopes are determined for each lithofacies. The evolution of (assumed) isotropic elastic moduli are examined during testing by use of unload-reload cycles, which permit the separation of total measured strains into elastic and plastic (permanent) strains. The upper Mt Simon samples deform largely elastically at stresses encountered in the Illinois Basin, with very little modulus degradation. The lower Mt. Simon facies are weaker and deform plastically, with varying amounts of modulus degradation. Results are interpreted via petrographic observation of textural contrasts. This range in constitutive response is captured up to failure with a phenomenological elasto-plasticity model. Essential aspects to describe observed behavior used in the model include non-associative plasticity, stress-invariant dependent failure, an elliptical cap surface capturing shear effects on pore collapse, kinematic and isotropic hardening, nonlinear elasticity and elastic-plastic coupling, among other features. Static moduli derived from laboratory tests are compared to dynamic moduli from wellbore log response which can allow experimental results and model to be extrapolated to Mt. Simon occurrences across the basin. This work was funded in part by the Center for Frontiers of Subsurface

  13. Evidence for gill slits and a pharynx in Cambrian vetulicolians: implications for the early evolution of deuterostomes

    Directory of Open Access Journals (Sweden)

    Ou Qiang

    2012-10-01

    Full Text Available Abstract Background Vetulicolians are a group of Cambrian metazoans whose distinctive bodyplan continues to present a major phylogenetic challenge. Thus, we see vetulicolians assigned to groups as disparate as deuterostomes and ecdysozoans. This divergence of opinions revolves around a strikingly arthropod-like body, but one that also bears complex lateral structures on its anterior section interpreted as pharyngeal openings. Establishing the homology of these structures is central to resolving where vetulicolians sit in metazoan phylogeny. Results New material from the Chengjiang Lagerstätte helps to resolve this issue. Here, we demonstrate that these controversial structures comprise grooves with a series of openings. The latter are oval in shape and associated with a complex anatomy consistent with control of their opening and closure. Remains of what we interpret to be a musculature, combined with the capacity for the grooves to contract, indicate vetulicolians possessed a pumping mechanism that could process considerable volumes of seawater. Our observations suggest that food captured in the anterior cavity was transported to dorsal and ventral gutters, which then channeled material to the intestine. This arrangement appears to find no counterpart in any known fossil or extant arthropod (or any other ecdysozoan. Anterior lateral perforations, however, are diagnostic of deuterostomes. Conclusions If the evidence is against vetulicolians belonging to one or other group of ecdysozoan, then two phylogenetic options seem to remain. The first is that such features as vetulicolians possess are indicative of either a position among the bilaterians or deuterostomes but apart from the observation that they themselves form a distinctive and recognizable clade current evidence can permit no greater precision as to their phylogenetic placement. We argue that this is too pessimistic a view, and conclude that evidence points towards vetulicolians being

  14. Marine Mo biogeochemistry in the context of dynamically euxinic mid-depth waters: A case study of the lower Cambrian Niutitang shales, South China

    Science.gov (United States)

    Cheng, Meng; Li, Chao; Zhou, Lian; Algeo, Thomas J.; Zhang, Feifei; Romaniello, Stephen; Jin, Cheng-Sheng; Lei, Li-Dan; Feng, Lian-Jun; Jiang, Shao-Yong

    2016-06-01

    Molybdenum (Mo) concentrations and Mo isotopes have been widely used as proxies for local and global redox conditions in early oceans (>520 Ma) that were stratified and characterized by dynamically euxinic mid-depth waters. However, the nature of the Mo cycle and accompanying isotopic fractionations in such oceans remain poorly known. To fill this gap, we conducted an integrated study of Mo isotopes and redox-sensitive trace element (RSTE) abundances in the Lower Cambrian Niutitang Formation at Yangjiaping, South China. This section accumulated on the northern shelf margin of the Nanhua Basin, a failed intracontinental rift basin with a good connection to the open ocean during the early Cambrian. The Niutitang Formation contains a ∼18-m-thick lower black shale member, and a ∼56-m-thick upper gray silty shale member. The lower member (LM) is moderately to strongly enriched in Mo, U and V, and heterogeneous in Mo isotopic composition (δ98Mo = -0.65‰ to +2.14‰), indicative of dominantly euxinic depositional conditions punctuated by ferruginous episodes (as shown by previously reported Fe speciation data). The upper member (UM) shows lesser enrichment of Mo, U and V, higher Mo/U ratios, and intermediate and more uniform Mo-isotopic compositions (δ98Mo = +1.16‰ to +1.71‰), indicative of weakly oxic to anoxic-euxinic depositional conditions. Geochemical profiles suggest that the LM-to-UM transition reflects a shift of the O2/H2S chemocline from the water column to the sediment. Large δ98Mo fluctuations in the LM may record variations of H2S concentrations in the mid-depth euxinic waters. The intermediate and relatively uniform δ98Mo values of the UM are attributed to the effects of a local Fe-Mn particulate shuttle. In light of the observations at Yangjiaping and other contemporaneous sections in the Nanhua Basin, we propose a new marine Mo biogeochemical model for the early Cambrian ocean. In this model, an Fe-Mn reduction zone may have developed above

  15. A discussions on the Cryogenian-Cambrian tectonic-sedimentary event and tectonic setting of northern Tarim Basin%塔里木盆地北缘南华纪—寒武纪构造背景及构造-沉积事件探讨

    Institute of Scientific and Technical Information of China (English)

    周肖贝; 李江海; 傅臣建; 李文山; 王洪浩

    2012-01-01

    Based on field geological investigation in northwestern and northeastern Tarim Basin, in combination with paleomagnetic mapping and previous researches, this paper has discussed the evolution process of the basin in Cryogenian-Cambrian period according to sedimentary-tectonic event and other evidence. Across the Tarim Basin, surface outcrops of Cryogenian to Cambrian sedimentary succession are completely exposed in the vicinity of Aksu area (northwest Tarim), Kuruktag (northeast Tarim) and southwest Tarim, thus providing a unique, well preserved and accessible means with which researchers can study the early development of northern Tarim Basin. The major lithologic types of Cryogenian-Cambrian strata in northeast Tarim include tillite, clastic rocks (rich in organic matter) and carbonate intercalated with volcanic rocks; in northwest Tarim, calstic rocks and carbonate are the common rock types, intercalated with small amounts of tillite and volcanic rocks. The north margin of Tarim block, which was a part of Rodinia supercontinent and neighbored the northwestern margin of Australia, was deeply rifted in Cryogenian-Ediacaran period and developed into two rifts in the northwestern and northeastern margin, forming a thick layer of rift-passive margin deposits, but the layer in the northwestern rift was not so completely developed as the layer in the northeastern margin. The earliest rift-passive margin sediments which can be observed are Cryogenian-Middle Ordovician strata, and the period can be divided into Cryogenian faulted period (supercontinent rifting stage) and Ediacaran-Middle Ordovician subsidence period (plate drifting stage).%以塔里木盆地西北缘和东北缘南区南华系—寒武系野外地质调查为基础,结合古地磁成图及前人研究资料,针对构造-沉积事件等探讨盆地北缘南华纪—寒武纪成盆演化过程.研究区保存了完整的南华纪—寒武纪地层,其中塔东北缘以冰碛岩、碎屑岩(富含有机

  16. Dysmenorrhea: Painful Periods

    Science.gov (United States)

    ... Management Education & Events Advocacy For Patients About ACOG Dysmenorrhea: Painful Periods Home For Patients Search FAQs Dysmenorrhea: ... Dysmenorrhea: Painful Periods FAQ046, January 2015 PDF Format Dysmenorrhea: Painful Periods Gynecologic Problems What is dysmenorrhea? How ...

  17. Familial Periodic Paralyses

    Science.gov (United States)

    ... two most common types of periodic paralyses are: Hypokalemic periodic paralysis is characterized by a fall in potassium levels ... chronic muscle weakness later in life. Hyperkalemic periodic paralysis is ... than the hypokalemic form. Muscle spasms are common. Is there any ...

  18. Algebraically periodic translation surfaces

    OpenAIRE

    Calta, Kariane; Smillie, John

    2007-01-01

    Algebraically periodic directions on translation surfaces were introduced by Calta in her study of genus two translation surfaces. We say that a translation surface with three or more algebraically periodic directions is an algebraically periodic surface. We show that for an algebraically periodic surface the slopes of the algebraically periodic directions are given by a number field which we call the periodic direction field. We show that translation surfaces with pseudo-Anosov automorphisms...

  19. Terminal suturing of Gondwana along the southern margin of South China Craton: Evidence from detrital zircon U-Pb ages and Hf isotopes in Cambrian and Ordovician strata, Hainan Island

    Science.gov (United States)

    Xu, Yajun; Cawood, Peter A.; Du, Yuansheng; Zhong, Zengqiu; Hughes, Nigel C.

    2014-12-01

    Hainan Island, located near the southern end of mainland South China, consists of the Qiongzhong Block to the north and the Sanya Block to the south. In the Cambrian, these blocks were separated by an intervening ocean. U-Pb ages and Hf isotope compositions of detrital zircons from the Cambrian succession in the Sanya Block suggest that the unit contains detritus derived from late Paleoproterozoic and Mesoproterozoic units along the western margin of the West Australia Craton (e.g., Northampton Complex) or the Albany-Fraser-Wilkes orogen, which separates the West Australia and Mawson cratons. Thus, in the Cambrian the Sanya Block was not part of the South China Craton but rather part of the West Australian Craton and its environs. In contrast, overlying Late Ordovician strata display evidence for input of detritus from the Qiongzhong Block, which constituted part of the southeastern convergent plate margin of the South China Craton in the early Paleozoic. The evolving provenance record of the Cambrian and Ordovician strata suggests that the juxtaposition of South China and West Australian cratons occurred during the early to mid-Ordovician. The event was linked with the northern continuation of Kuungan Orogeny, with South China providing a record of final assembly of Gondwana.

  20. Cambrian intermediate-mafic magmatism along the Laurentian margin: Evidence for flood basalt volcanism from well cuttings in the Southern Oklahoma Aulacogen (U.S.A.)

    Science.gov (United States)

    Brueseke, Matthew E.; Hobbs, Jasper M.; Bulen, Casey L.; Mertzman, Stanley A.; Puckett, Robert E.; Walker, J. Douglas; Feldman, Josh

    2016-09-01

    The Southern Oklahoma Aulocogen (SOA) stretches from southern Oklahoma through the Texas panhandle and into Colorado and New Mexico, and contains mafic through silicic magmatism related to the opening of the Iapetus Ocean during the early Cambrian. Cambrian magmatic products are best exposed in the Wichita Mountains (Oklahoma), where they have been extensively studied. However, their ultimate derivation is still somewhat contentious and centers on two very different models: SOA magmatism has been suggested to occur via [1] continental rifting (with or without mantle plume emplacement) or [2] transform-fault related magmatism (e.g., leaky strike-slip faults). Within the SOA, the subsurface in and adjacent to the Arbuckle Mountains in southern Oklahoma contains thick sequences of mafic to intermediate lavas, intrusive bodies, and phreatomagmatic deposits interlayered with thick, extensive rhyolite lavas, thin localized tuffs, and lesser silicic intrusive bodies. These materials were first described in the Arbuckle Mountains region by a 1982 drill test (Hamilton Brothers Turner Falls well) and the best available age constraints from SOA Arbuckle Mountains eruptive products are ~ 535 to 540 Ma. Well cuttings of the mafic through intermediate units were collected from that well and six others and samples from all but the Turner Falls and Morton wells are the focus of this study. Samples analyzed from the wells are dominantly subalkaline, tholeiitic, and range from basalt to andesite. Their overall bulk major and trace element chemistry, normative mineralogy, and Srsbnd Nd isotope ratios are similar to magmas erupted/emplaced in flood basalt provinces. When compared with intrusive mafic rocks that crop out in the Wichita Mountains, the SOA well cuttings are geochemically most similar to the Roosevelt Gabbros. New geochemical and isotope data presented in this study, when coupled with recent geophysical work in the SOA and the coeval relationship with rhyolites, indicates

  1. The classification and discussion on geologic period of mesozoic volcanic rock series in Longnan basin, Jiangxi

    International Nuclear Information System (INIS)

    The author analyses the classification actuality and existing questions of the Yutian group of Mesozoic volcanic rock series in Longnan basin, Jiangxi province. According to lithology and lithofacies, the sandstone and conglomerate association of the low part of Yutian group should be classified into Shuitoujing formation, and the basic rock association of the middle part and the medium acidic rock association of the upper part of Yutian group should be classified into Changpu formation. The metamorphic rock association which is classified into volcanic rock series by former should be classified into Sinian-cambrian system. The isotope data shows that geological period of Changpu formation in Yutian group is from middle Jurassic epoch to early stage of late Jurassic epoch. It needs further certification that the early Jurassic epoch sediment occurs in Shuitoujing formation or not

  2. Alluvial-eolian interaction in a Cambrian rift margin: the Pedra das Torrinhas and Pedra Pintada formations (Guaritas Group, RS

    Directory of Open Access Journals (Sweden)

    André Marconato

    2009-12-01

    Full Text Available This work presents a study of selected outcrops from the Pedra das Torrinhas Formation of the Guaritas Group (Cambrian, Camaquã Basin, near the basin bordering Encantadas Fault Zone. The studied succession includes alluvial fan deposits that pass laterally into eolian deposits. Sedimentary facies and architectural element analysis were performed, followed by sedimentary petrography and microscopic porosity analysis, aiming to characterize the porosity of the deposits and its spatial distribution. The main objective was to contribute to a better understanding of the porosity spatial distribution in depositional systems characterized by the interaction between alluvial and eolian processes, with special reference to deposits formed prior to the development of terrestrial plants. Porosity values are related to depositional processes, with higher porosities associated to eolian dune deposits (mean of 8.4%, and lower porosity related to interdunes (mean of 3.4% and alluvial fans (mean of 4.3%. Architectural elements analysis revealed the spatial relationships of these deposits, a response to the interplay of the eolian and alluvial processes. The integration of porosity data reveals that the interaction of alluvial and eolian processes results in heterogeneous distribution of porosity at the facies association scale. Eolian reworking of alluvial facies increases porosity whereas sheet-flood and other alluvial processes in the interdune areas reduce porosity.O presente trabalho consiste no estudo de afloramentos da Formação Pedra das Torrinhas do Grupo Guaritas (Cambriano, Bacia Camaquã, próximo à Zona de Falha das Encantadas. As sucessões estudadas incluem depósitos de leques aluviais que passam lateralmente para depósitos eólicos. Foram realizadas análises de fácies e de elementos arquiteturais, seguidos de petrografia sedimentar e análise microscópica de porosidade, com o objetivo de caracterizar a porosidade da unidade e sua

  3. Two new species of Vestrogothia (Phosphatocopina, Crustacea) of Orsten-type preservation from the Upper Cambrian in western Hunan, South China

    Institute of Scientific and Technical Information of China (English)

    2009-01-01

    Although fossils of Orsten-type preservation represented by Skaracarida and Phosphatocopina were first reported from the Middle and Upper Cambrian in western Hunan, South China in 2005, diversified phosphatocopine species have never been appropriately described and elucidated in terms of their evolutionary relationships. Here, we described two new species of Phosphatocopina, Vestrogothia anterispinata sp. nov. and V. bispinata sp. nov. The evolutionary relationship among all the valid species of Phosphatocopina is discussed and the phylogeny of Phosphatocopina is reestablished using cladistic analysis. Accordingly, phosphatocopines primitively evolved along two lineages: one evolved towards the emergence of lobes; the other evolved towards the emergence of dorsal spines. Respectively, lobes originated independently three times, and dorsal spines originated only once. With regard to the dorsal rims, interdorsum originated first, whereas dorsal furrow originated independently twice on the basis of interdorsum. Probably Hesslandona may represent a polyphyletic group, whereas Vestrogothia a monophyletic group.

  4. A comparison of the biological,geological events and environmental backgrounds between the Neoproterozoic-Cambrian and Permian-Triassic transitions

    Institute of Scientific and Technical Information of China (English)

    2010-01-01

    The Neoproterozoic-Cambrian(N-C) and Permian-Triassic(P-T) transitions have been regarded the two most critical transitions in earth history because of the explosive biological radiation in the early Cambrian(the Cambrian Explosion) and the largest mass extinction at the end-Permian.Previous studies suggest that these two critical transitions showed certain comparability in major evolutionary events.In other words,a series of biological,geological,and geochemical events that had happened in the N-C transition occurred repeatedly during the P-T transition.Those events included continental re-configuration related to the deep mantle dynamics,global-scale glaciations,large C-,Sr-,and S-isotope perturbations indicating atmospheric and oceanic changes,abnormal precipitation of carbonates,and associated multiple biological radiations and mass extinctions.The coupling of those events in both N-C and P-T transitions suggests that deep mantle dynamics could be a primary mechanism driving dramatic changes of environment on the earth’s surface,which in turn caused major biological re-organizations.A detailed comparison of those events during the two critical transitions indicates that despite their general comparability,significant differences do exist in magnitude,duration,and frequency.The supercontinent Rodinia began to rift before the Snowball Earth time.By contrast,the supercontinent Pangea entered the dispersal stage after the greatest glaciation from the Late Carboniferous to Cisuralian.Quantitative data and qualitative analyses of different fossil groups show a more profound mass extinction during the N-C transition than at the end-Permian in terms of ecosystem disruption.This is indicated by the disappearance of the whole Ediacaran biota at the N-C boundary.The subsequent appearances of many new complex animals at phylum level in the early Cambrian mark the establishment of a brand new ecosystem.However,the end-Permian mass extinction is manifested mainly by the

  5. The origin of life near deep-sea hydrothermal systems during the Cambrian explosion: data from the Kyzyl Tashtyg sulphide deposit (Central Asia)

    Science.gov (United States)

    Simonov, Vladimir; Terleev, Alexander; Safonova, Inna; Kotlyarov, Alexey; Stupakov, Sergey; Tokarev, Dmitry

    2015-04-01

    On Earth the solar radiation and the hydrothermal circulation both affect life evolution. Recent extensive studies of the World Ocean have shown that the biodiversity of Earth is linked with hydrothermal activity on the oceanic floor. These deep-sea ecosystems use chemical energy, not solar radiation. In the last quarter of the XX century, a new type of hydrothermal systems, so-called black smokers, was discovered in mid-oceanic ridges. As black smokers form sulfide ores and are surrounded by abundant bio-oases or symbioses, identification of their analogues in ancient orogenic belts is necessary for studying life origin and evolution. Of special importance are problems of life associated with deep-sea hydrothermal systems acted at the Precambrian-Cambrian boundary - the time of Cambrian explosion (Maruyama et al., 2013). During that explosion life significantly evolved and diversified due to dramatic changes of Earth's environment. Consequently, the early Cambrian - late Precambrian Kyzyl Tashtyg sulphide deposit of East Tuva in the Central Asian Orogenic Belt is of special interest. This deposit was formed on the bottom of ancient back-arc deep-sea basin as a result of black smoker hydrothermal activity and is hosted by volcanogenic-sedimentary rocks altered by the high temperature solutions. The altered Kyzyl Tashtyg basalts have an amygdules (filled by albite, epidote and carbonates), contain brown-green microfossils, often attached to their walls. The microfossils are thin tubes 5 to 25 microns in diameter and 500 microns long. This tubes are empty and have straight, curved or branching shape. Chemically, the tube material is close to epidote. In consideration of microscopic dimensions, simple morphology and similarity with modern tubular microorganisms, the studied tube-shaped microfossils can be related to cyanobacteria. Almost the same fossils, associated with oceanic basalt complexes, were described earlier (Furnes et al., 2007; Mcloughlin et al., 2007

  6. Paleoproterozoic mojaveprovince in northwestern Mexico? Isotopic and U-Pb zircon geochronologic studies of precambrian and Cambrian crystalline and sedimentary rocks, Caborca, Sonora

    Science.gov (United States)

    Lang, Farmer G.; Bowring, S.A.; Matzel, J.; Maldonado, G.E.; Fedo, C.; Wooden, J.

    2005-01-01

    Whole-rock Nd isotopic data and U-Pb zircon geochronology from Precambrian crystalline rocks in the Caborca area, northern Sonora, reveal that these rocks are most likely a segment of the Paleoproterozoic Mojave province. Supporting this conclusion are the observations that paragneiss from the ??? 1.75 Ga Bamori Complex has a 2.4 Ga Nd model age and contains detrital zircons ranging in age from Paleo- proterozoic (1.75 Ga) to Archean (3.2 Ga). Paragneisses with similar age and isotopic characteristics occur in the Mojave province in southern California. In addition, "A-type" granite exposed at the southern end of Cerro Rajon has ca 2.0 Ga Nd model age and a U-Pb zircon age of 1.71 Ga, which are similar to those of Paleoproterozoic granites in the Mojave province. Unlike the U.S. Mojave province, the Caborcan crust contains ca. 1.1 Ga granite (Aibo Granite), which our new Nd isotopic data suggest is largely the product of anatexis of the local Precambrian basement. Detrital zircons from Neoproterozoic to early Cambrian miogeoclinal arenites at Caborca show dominant populations ca. 1.7 Ga, ca. 1.4 Ga, and ca. 1.1 Ga, with subordinate Early Cambrian and Archean zircons. These zircons were likely derived predominately from North American crust to the east and northeast, and not from the underlying Caborcan basement. The general age and isotopic similarities between Mojave province basement and overlying miogeoclinal sedimentary rocks in Sonora and southern California is necessary, but not sufficient, proof of the hypothesis that Sonoran crust is allochthonous and was transported to its current position during the Mesozoic along the proposed Mojave-Sonora megashear. One viable alternative model is that the Caborcan Precambrian crust is an isolated, autochthonous segment of Mojave province crust that shares a similar, but not identical, Proterozoic geological history with Mojave province crust found in the southwest United States ?? 2005 Geological Society of America.

  7. Periodic trajectories in TDHF

    International Nuclear Information System (INIS)

    A condition for the existence of a periodic TDHF trajectory of period T is derived. It takes a from very similar to the static H.F. equation and shows that associated to a periodic trajectory there is a static single particle hamiltonian which is a complicated functional of the time dependent density matrix. An explicit expansion for this functional is derived. It is shown that many properties of the static H.F. rest point are shared by periodic solutions. (Author)

  8. Painful menstrual periods

    Science.gov (United States)

    ... you sexually active? Do you use birth control? What type? When was your last menstrual period? Was the flow of your last menstrual period a normal amount for you? Do your periods tend ... with menstruation? What have you done to try to relieve the ...

  9. The Periodic Pyramid

    Science.gov (United States)

    Hennigan, Jennifer N.; Grubbs, W. Tandy

    2013-01-01

    The chemical elements present in the modern periodic table are arranged in terms of atomic numbers and chemical periodicity. Periodicity arises from quantum mechanical limitations on how many electrons can occupy various shells and subshells of an atom. The shell model of the atom predicts that a maximum of 2, 8, 18, and 32 electrons can occupy…

  10. Rapid Polar Wander in North America During the Early Cambrian: Paleomagnetic and Magnetostratigraphic Constraints from the Peurto Blanco Volcanics from Caborca, and the Tapeats Sandstone of the Grand Canyon.

    Science.gov (United States)

    Kirschvink, J. L.; Elston, D. P.; Petterson, R.; Hagadorn, J. W.; Raub, T. D.; Evans, D. A.; Rose, E. C.; Weiss, B. P.; Barr, T. D.

    2006-12-01

    We report paleomagnetic directions from a volcaniclastic unit in the Early Cambrian Puerto Blanco Formation near Caborca in Sonora, Mexico, and from a restudy of the Tapeats Sandstone in the Grand Canyon, Arizona. Volcaniclastics in Caborca are found above carbonates of La Cienega Formation, which bear the Ediacaran index fossil Cloudina, and are overlain by siliciclastics bearing the Early Cambrian fossil Treptichnus pedum. Thin basaltic flows within the sequence show evidence of early oxidation of magnetite to hematite yielding a strong and stable remnant magnetization, and characteristic NRM directions obtained during progressive alternating field and thermal demagnetization from the units fall in two antipodal easterly westerly groups with moderate inclination. Volcanic clasts from the Puerto Blanco Formation show statistically random dispersion of components carried both by magnetite and by hematite, indicating multiple episodes of erosion and redeposition, and provide a positive conglomerate test. After correction for offset by the Sonora/Mojave megashear in Mesozoic time, these data imply a North magnetic pole at ~21º S latitude, 142º E longitude, suggesting that North America was at moderate southerly latitudes during the first part of Early Cambrian time. In contrast, near-equatorial paleolatitude is indicated for the earliest Middle Cambrian Tapeats Sandstone in the Grand Canyon and vicinity. Our extensive dm-resolution magnetostratigraphic study of this unit, involving stable data from about 250 samples from 7 sections, confirms an earlier report of Elston & Bressler (1977) which demonstrated a low-latitude, two-polarity characteristic remanence with stratigraphically-bounded polarity reversals. We provide positive reversal and conglomerate tests for this characteristic magnetization. The paleogeographic shift toward the equator during the last portion of Early Cambrian time helps account for the earlier appearance and higher abundance of carbonate

  11. Periodicity of chaotic solutions

    CERN Document Server

    Berezowski, Marek

    2016-01-01

    The scope of the paper is the analysis of the impact of flow reversal on the dynamics of cascades of reactors. Periodic and chaotic oscillations occur in the analyzed system. There is a dependence between the oscillation period of the state variable of the system without flow reversal and the recurrence period of windows of chaos in the steady-state diagram of the system with flow reversal.

  12. Crystallography without periodicity

    International Nuclear Information System (INIS)

    A condition of physics stability, applied to materials with sharp Bragg peaks, leads to rules for a unified approach to their classification that make no use of periodicity, applying equally well to either periodic or quasiperiodic materials. Three-dimensional geometric intuition is the primary analytical tool for extracting the classification from the rules. Recognizing that periodicity is not required to formulate crystallography removes the need for an algebraic formalization of that intuition in a higher dimensional superspace, where the material can be embedded in a periodic supercrystal. Selected examples illustrate the simplicity of the resulting description of ordinary crystals, quasicrystals, modulated crystals, intergrowth crystals, or modulated quasicrystals. (Author) 9 refs

  13. 寒武纪早期大气-海洋氧含量与生命大爆发%Atmosphere-Ocean Oxygen Levels and Biotic Explosion in the Early Cambrian

    Institute of Scientific and Technical Information of China (English)

    李超; 金承胜

    2015-01-01

    寒武纪早期(541~510,Ma)地球环境与这一时期生命大爆发之间的关系一直是地球生物学研究的热点问题之一。本文系统总结了目前寒武纪早期大气-海洋氧含量与这一时期生命辐射之间关系的3种假说:大气-海洋的氧含量增加导致了寒武纪生命大爆发;寒武纪生命大爆发导致了大气-海洋氧化以及二者之间没有因果关系。3种假说均有相应的支持证据,但也存在与寒武纪早期海洋化学记录、与现代海洋观察不符和上述假说均未考虑寒武纪早期生命演化所展示的时空差异性等问题。在上述3种假说的基础之上,本文通过对寒武纪构造活动、陆源输入、海洋化学和生命演化等最新资料的综合讨论和分析表明:寒武纪早期地球环境与生命辐射之间很可能是相互作用与协同演化关系,而非简单的单向关系。%The relationship between the Earth environment and the biotic explosion in early Cambrian (ca.541 -5 10,Ma) is one top geobiological issue.Here,we systematically summarized three major hypotheses for the relationship between the atmosphere-ocean oxygen level and the biotic evolution in early Cambrian,including that (1)increasing oxygenation of at-mosphere and ocean caused the Cambrian explosion,(2)the emergence and evolution of lives resulted in oxygenation of atmosphere and ocean,(3)the atmosphere-ocean oxygen level and the evolution of lives are not interrelated.Although each hypothesis is supported by specific evidences,these hypotheses show significantly inconsistent with partial ocean chemistry records in early Cambrian and investigations of lives in modern ocean analogues.In addition,the spatiotemporal heterogeneity of early Cambrian life evolution is not considered in these hypotheses.By synthesizing these major hypotheses with available data on Cambrian tectonic activities,terrestrial fluxes,ocean chemistry and life evolution patterns,we dis

  14. Electric property evidences of carbonification of organic matters in marine shales and its geologic significance: A case study of the Lower Cambrian Qiongzhusi shale in the southern Sichuan Basin

    OpenAIRE

    Yuman Wang; Dazhong Dong; Xiangzhi Cheng; Jinliang Huang; Shufang Wang; Shiqian Wang

    2014-01-01

    Searching for some reliable evidences that can verify the carbonification of organic matters in marine shales is a major scientific issue in selecting shale gas fairways in old strata. To this end, based on core, logging and testing data, the electric property of two organic-rich shale layers in the Lower Cambrian Qiongzhusi Fm. and the Lower Silurian Longmaxi Fm. in the southern Sichuan Basin was compared to examine the carbonification signs of organic matters in the Qiongzhusi shale and its...

  15. The Occurrence of Longfengshania in the Early Cambrian from Haikou, Yunnan, China%龙凤山藻在澄江早寒武世生物群中的发现

    Institute of Scientific and Technical Information of China (English)

    徐兆良

    2002-01-01

    Well preserved Early Cambrian small individual megascopic carbonaceous algal fossils have been found from the Chengjiang Biota. This paper deals with the new megascopic algal fossils in the Early Cambrian also from Chengjiang Biota at Ercai Village, Haikou, Kunming, Yunnan, southwest China. Two new taxa, Longfengshania cordata sp. nov. and Plantulaformis sinensis gen. et sp. nov. are described here in detail. These new findings show that the Chengjiang Biota is high in species diversity of fossil algae and might offer new evidence for a better understanding of the Cambrian explosive biological evolution. Moreover, the fossils described here might belong to the megaalgal fossils according to detailed morphological study on the relationships of Longfengshania. Additionally, the occurrence of Longfengshania in the early Cambrian and the Neoproterozoic probably indicates a wide ecological tolerance and flexibility and apparently reflects its evolutionary conservation and a high degree of inherent genetic stability.%主要报道了产自云南昆明海口耳材村早寒武世筇竹寺组玉案山段澄江生物群中可能为自由漂浮生长的宏观藻类化石--心型龙凤山藻(新种)(Longfengshania cordata sp. nov.)和中华豆芽藻(新属、新种)(Plantulaformis sinensis gen. et sp. nov.).这些化石的发现进一步显示了澄江生物群物种多样性,为揭示该生物群爆发性演化提供了新的证据.通过对龙凤山藻属亲缘关系的深入比较研究,进一步证明该类化石应归属宏观藻类以及它们遗传上的稳定性、演化上的保守性和环境上的适应性.

  16. La Formación Achavil: una nueva unidad de bajo grado metamórfico en la evolución Cámbrica Superior del Famatina The Achavil Formation: A new low-grade metamorphic unit in the Upper Cambrian evolution of Famatina

    Directory of Open Access Journals (Sweden)

    Gilda Collo

    2008-09-01

    Formation. The depositional age for the Achavil Formation ranges between Middle-Upper and Upper Cambrian, considering that the immediately overlying Volcancito Formation ranges between uppermost Cambrian and Tremadocian. This interval would correlate with the Mesón Group in northwest Argentina and not, as it has often been interpreted, with the Puncoviscana Formation. Two superposed deformational episodes inferred to be Cambrian are recorded within the Achavil Formation and clearly predate the deposition of the Volcancito Formation. From our results potential correlations between the Achavil Formation and other lower Paleozoic units in the surrounding regions allow establishing a paleogeographic reconstruction for the western Gondwana margin during this period.

  17. Historically defined autobiographical periods

    DEFF Research Database (Denmark)

    Brown, Norman R.; Hansen, Tia G. B.; Lee, Peter J.;

    2012-01-01

    The chapter reviews a research programme that has demonstrated the existence of historically defined autobiographical periods and identified the conditions that bring them about. Data from four samples of World War II-generation adults show that historically defined autobiographical periods endure...

  18. The Living Periodic Table

    Science.gov (United States)

    Nahlik, Mary Schrodt

    2005-01-01

    To help make the abstract world of chemistry more concrete eighth-grade students, the author has them create a living periodic table that can be displayed in the classroom or hallway. This display includes information about the elements arranged in the traditional periodic table format, but also includes visual real-world representations of the…

  19. Second Period Access Report

    DEFF Research Database (Denmark)

    Maisondieu, Christophe; Giebhardt, Jochen; Tetu, Amelie; Johnstone, Cameron; Healy, Mark

    The work described in this publication has received support from the European Community - Research Infrastructure Action under the FP7 “Capacities” Specific Programme through grant agreement number 262552, MaRINET. Project Periodic Report. 2nd Period: October 2012 – March 2014 inclusive....

  20. On some periodicity effects

    DEFF Research Database (Denmark)

    Sorokin, Sergey V.

    2015-01-01

    . In each case, special attention is paid to eigenfrequencies and eigenmodes of a single periodicity cell with appropriate boundary conditions. The influence of the amount of periodicity cells in a finite compound structure on its eigenfrequency spectrum is analyzed. Several features common for the......The talk is concerned with the modelling of wave propagation in and vibration of periodic elastic structures. Although analysis of wave-guide properties of infinite periodic structures is a well establish research subject, some issues have not yet been fully addressed in the literature. The aim of...... the talk is to illustrate these issues in simple examples and to discuss possible applications and generalisations. First, the eigenfrequency spectra of finite periodic structures are compared with the location of stop-bands for their infinite counterparts for a hierarchy of four mathematical models...

  1. Zircon dating of Neoproterozoic and Cambrian ophiolites in West Mongolia and implications for the timing of orogenic processes in the central part of the Central Asian Orogenic Belt

    Science.gov (United States)

    Jian, Ping; Kröner, Alfred; Jahn, Bor-ming; Windley, Brian F.; Shi, Yuruo; Zhang, Wei; Zhang, Fuqin; Miao, Laicheng; Tomurhuu, Dondov; Liu, Dunyi

    2014-06-01

    , we re-subdivide West Mongolia into: a latest Neoproterozoic-early Cambrian, arc-microcontinent collision zone north of the MML; a Cambrian Gobi Altai ophiolite-microcontinent collision zone and a Cambrian Trans-Altai forearc complex south of the MML. The central CAOB evolved in five phases: subduction initiation and arc formation (ca. 573 to > ca. 540 Ma); arc-microcontinent collision (ca. 535-524 Ma); a continuum of slab delamination, overthrusting, crustal thickening and surface uplift (ca. 519-482 Ma) in Northwest Mongolia; initiation of new subduction zones in South Mongolia (ca. 523-511 Ma); and continuing orogeny with local surface uplift. Overall, the current, documented timing of orogenic development in the central CAOB is largely consistent with a W/SW-Pacific style of evolution in terms of subduction initiation, short timescales of individual orogenies, and episodic subduction-collision during a continuing migration of subduction zones.

  2. 早寒武世初期沂沭断裂带地震效应%Seismic effects from Yishu Fault Zone during the earlier Early Cambrian

    Institute of Scientific and Technical Information of China (English)

    田洪水; 张邦花; 祝介旺; 张增奇; 李洪奎

    2011-01-01

    In the earlier Early Cambrian, with the proceeding of the marine invasion from the Paleo-Yishu Strait to west, uneven thick deposits of sandstone, mudstone and carbonate rock of littoral, lagoon and neritic facies were deposited in western Shandong Province. The horizon of these layers belongs to the bottom part of Lower Cambrian consisting of the Liguan and Zhushadong Formations. Together with the depositional and formation processes, tectonic rift-depression was taking place in the Yishu Fault Zone, characterized by frequent submarine earthquakes with different seismic effects within 200 km range from the Yishu Fault Zone to west. Various seismogenetic syndepositional deformation structures are formed in the seismic records. Seismic effect records developed in the Zhushadong Formation are mainly soft-sediment deformations such as diapir structure, carbonate lime-mud volcano, liquefied vein, liquefied brec cia , seismic fold, graded-fault, siliceous vein etc., besides a few brittle deformation of semi-consolidated sediments. Those preserved in the Liguan Formation are not only slump fold, load structure and ball-and pillow structure, but also some seismo-turbidite. Magnitude reflected by submarine seismic effects might be Richater magnitude of 5. 0 ~ 8. 9. These evidences indicate that strong tectonic activities occurred along the Yishu Fault Zone in the Early Cambrian. Analyses from the viewpoint of marine engineering geology show that seismic effects mentioned above are destructive forms of sediment layers caused by earthquake at the sea-bottom. This paper has provided intuitive materials for understanding earthquake destruction to rock-soil layers of seafloor-engineering foundation.%早寒武世初期,随着从古沂沭海峡向西海侵的发展,鲁西地区沉积了厚度不等的主要由砂岩、泥岩和碳酸盐岩组成的滨海、潟湖及浅海相沉积,其层位属下寒武统底部的李官组和朱砂洞组.在它们沉积过程中,沂沭断裂带

  3. Origin of Middle Cambrian and Late Silurian potassic granitoids from the western Kunlun orogen, northwest China: a magmatic response to the Proto-Tethys evolution

    Science.gov (United States)

    Liu, Zheng; Jiang, Yao-Hui; Jia, Ru-Ya; Zhao, Peng; Zhou, Qing; Wang, Guo-Chang; Ni, Chun-Yu

    2014-02-01

    This paper presents new SHRIMP zircon U-Pb chronology, major and trace element, and Sr-Nd-Hf isotopic data of two Early Paleozoic granitic plutons (Yierba and North Kudi) from the western Kunlun orogen, in attempt to further constrain the Proto-Tethys evolution. SHRIMP zircon U-Pb dating shows that the Yierba pluton was emplaced in the Middle Cambrian (513 ± 7 Ma) and the North Kudi pluton was emplaced in the Late Silurian (420.6 ± 6.3 Ma). The Yierba pluton consists of quartz monzodiorite, quartz monzonite and granodiorite. These granitoids are metaluminous and potassic, with initial 87Sr/86Sr ratios of 0.7072-0.7096, ɛNd (T) of -0.2 to -1.6 and ɛHf (T) (in-situ zircon) of -1.2. Elemental and isotopic data suggest that they were formed by partial melting of subducted sediments, with subsequent melts interacting with the overlying mantle wedge in an oceanic island arc setting in response to the intra-oceanic subduction of Proto-Tethys. The North Kudi pluton consists of syenogranite and alkali-feldspar granite. These granites are metaluminous to weakly peraluminous and potassic. They show an affinity of A1 subtype granite, with initial 87Sr/86Sr ratios of 0.7077-0.7101, ɛNd (T) of -3.5 to -4.0 and ɛHf (T) (in-situ zircon) of -3.9. Elemental and isotopic data suggest that they were formed by partial melting of the Precambrian metamorphic basement at a shallow depth (<30 km) during the post-orogenic regime caused by Proto-Tethyan oceanic slab break-off. Our new data suggest that the subduction of the Proto-Tethyan oceanic crust was as early as Middle Cambrian (˜513 Ma) and the final closure of Proto-Tethys was not later than Late Silurian (˜421 Ma), most probably in Middle Silurian.

  4. The early Cambrian Chahmir shale-hosted Zn-Pb deposit, Central Iran: an example of vent-proximal SEDEX mineralization

    Science.gov (United States)

    Rajabi, Abdorrahman; Rastad, Ebrahim; Canet, Carles; Alfonso, Pura

    2015-06-01

    The Chahmir zinc-lead deposit (1.5 Mt @ 6 % Zn + 2 % Pb) in Central Iran is one among several sedimentary-exhalative Zn-Pb deposits in the Early Cambrian Zarigan-Chahmir basin (e.g., Koushk, Darreh-Dehu, and Zarigan). The deposit is hosted by carbonaceous, fine-grained black siltstones, and shales interlayered with volcaniclastic sandstone beds. It corresponds to the upper part of the Early Cambrian volcano-sedimentary sequence (ECVSS), which was deposited on the Posht-e-Badam Block during back-arc rifting of the continental margin of Central Iran. Based on crosscutting relationships, mineralogy, and texture of sulfide mineralization, four different facies can be distinguished: stockwork (feeder zone), massive ore, bedded ore, and distal facies (exhalites with barite). Silicification, carbonatization, sericitization, and chloritization are the main wall-rock alteration styles; alteration intensity increases toward the proximal feeder zone. Fluid inclusion microthermometry was carried out on quartz associated with sulfides of the massive ore. Homogenization temperatures are in the range of 170-226 °C, and salinity is around 9 wt% NaCl eq. The size distribution of pyrite framboids of the bedded ore facies suggests anoxic to locally suboxic event for the host basin. δ34S(V-CDT) values of pyrite, sphalerite, and galena range from +10.9 to +29.8 ‰. The highest δ34S values correspond to the bedded ore (+28.6 to +29.8 ‰), and the lowest to the massive ore (+10.9 to +14.7 ‰) and the feeder zone (+11.3 and +12.1 ‰). The overall range of δ34S is consistent with a sedimentary environment where sulfide sulfur was derived from two sources. One of them was corresponding to early ore-stage sulfides in bedded ore and distal facies, consistent with bacterial reduction from coeval seawater sulfate in a closed or semiclosed basin. However, the δ34S values of late ore-stage sulfides, observed mainly in massive ore, interpreted as a hydrothermal sulfur component, leached

  5. Painful periods (dysmenorrhea) (image)

    Science.gov (United States)

    Primary dysmenorrhea is a normal cramping of the lower abdomen caused by hormone-induced uterine contractions before the period. Secondary dysmenorrhea may be caused by abnormal conditions such as ...

  6. Setting the Periodic Table.

    Science.gov (United States)

    Saturnelli, Annette

    1985-01-01

    Examines problems resulting from different forms of the periodic table, indicating that New York State schools use a form reflecting the International Union of Pure and Applied Chemistry's 1984 recommendations. Other formats used and reasons for standardization are discussed. (DH)

  7. The Periodic Table CD.

    Science.gov (United States)

    Banks, Alton J.; Holmes, Jon L.

    1995-01-01

    Describes the characteristics of the digitized version of The Periodic Table Videodisc. Provides details about the organization of information and access to the data via Macintosh and Windows computers. (DDR)

  8. Establishing contract periods

    International Nuclear Information System (INIS)

    The lead time for executing the Adjustable Fixed-Commitment (AFC) contract and exceptions which may be considered are discussed. The initial delivery period is also discussed. Delays, deferrals, and schedule adjustment charges are finally considered

  9. Two new species of Vestrogothia (Phosphatocopina,Crustacea) of Orsten-type preservation from the Upper Cambrian in western Hunan,South China

    Institute of Scientific and Technical Information of China (English)

    ZHANG HuaQiao; DONG XiPing

    2009-01-01

    Although fossils of Orsten-type preservation represented by Skaracarida and Phosphatocopina were first reported from the Middle and Upper Cambrian in western Hunan,South China in 2005,diversified phosphatocopine species have never been appropriately described and elucidated in terms of their evolutionary relationships.Here,we described two new species of Phosphatocopina,Vestrogothia anterispinata sp.nov.and V.bispinata sp.nov.The evolutionary relationship among all the valid spe-cies of Phosphatocopina is discussed and the phylogeny of Phosphatocopina is reestablished using cladistic analysis.Accordingly,phosphatocopines primitively evolved along two lineages:one evolved towards the emergence of lobes; the other evolved towards the emergence of dorsal spines.Respec-tively,lobes originated independently three times,and dorsal spines originated only once.With regard to the dorsal rims,interdorsum originated first,whereas dorsal furrow originated independently twice on the basis of interdorsum.Probably Hesslandona may represent a polyphyletic group,whereas Vestrogothia a monophyletic group.

  10. Late-stage sulfides and sulfarsenides in Lower Cambrian black shale (stone coal) from the Huangjiawan mine, Guizhou Province, People's Republic of China

    Science.gov (United States)

    Belkin, H.E.; Luo, K.

    2008-01-01

    The Ni-Mo Huangjiawan mine, Guizhou Province, People's Republic of China, occurs in Lower Cambrian black shale (stone coal) in an area where other mines have recently extracted ore from the same horizon. Detailed electron microprobe (EMPA) and scanning electron microscope (SEM) analyses of representative thin sections have revealed a complex assemblage of sulfides and sulfarsenides. Early sulfidic and phosphatic nodules and host matrix have been lithified, somewhat fractured, and then mineralized with later-stage sulfides and sulfarsenides. Gersdorffite, millerite, polydymite, pyrite, sphalerite, chalcopyrite, galena, and clausthalite have been recognized. EMPA data are given for the major phases. Pyrite trace-element distributions and coeval Ni-, As-sulfides indicate that in the main ore layer, the last sulfide deposition was Ni-As-Co-rich. Mo and V deposition were early in the petrogenesis of these rocks. The assemblages gersdorffite-millerite-polydymite (pyrite) and millerite-gersdorffite (pyrite) and the composition of gersdorffite indicate a formation temperature of between 200?? and 300??C suggesting that the last solutions to infiltrate and mineralize the samples were related to hydrothermal processes. Environmentally sensitive elements such as As, Cd, and Se are hosted by sulfides and sulfarsenides and are the main source of these elements to residual soil. Crops grown on them are enriched in these elements, and they may be hazardous for animal and human consumption. ?? Springer-Verlag 2007.

  11. THE "CAMBRIAN EXPLOSION" IN THE EVOLUTION OF METAZOA AND ITS CAUSES%"寒武纪爆发"事件及其成因述评

    Institute of Scientific and Technical Information of China (English)

    郝家胜

    2003-01-01

    主要后生动物门类化石在寒武纪初期的突然出现记录了生命史中一个从未有过的快速演化时期,这一事件被称为"寒武纪爆发(Cambrian explosion)".多年来有关"寒武纪爆发"成因的各种猜测从未间断过.但是,将古生物化石记录看成是生物进化过程中种系形成的真实过程已遭到许多研究者的非议,这主要取决于以下二方面的证据:一是对早期后生动物系统发生关系的重新评估;二是来自于分子系统学和分子钟的研究.现在看来,"寒武纪爆发"事件的产生,一方面归因于化石体积的明显增加及其伴随的骨骼化过程,另一方面可能是其时的进化革新事件所导致.

  12. Periodic solutions of periodically harvested lotka-volterra systems

    OpenAIRE

    Hausrath, Alan R.; Manasevich, Raul F.

    2012-01-01

    We study a Lotka-Volterra system with periodic harvesting, find sufficient conditions for the existence of periodic solutions with the same period, and, under certain conditions, count the number of such periodic solutions.

  13. Periodic safety analyses

    International Nuclear Information System (INIS)

    The IAEA Safety Guide 50-SG-S8 devoted to 'Safety Aspects of Foundations of Nuclear Power Plants' indicates that operator of a NPP should establish a program for inspection of safe operation during construction, start-up and service life of the plant for obtaining data needed for estimating the life time of structures and components. At the same time the program should ensure that the safety margins are appropriate. Periodic safety analysis are an important part of the safety inspection program. Periodic safety reports is a method for testing the whole system or a part of the safety system following the precise criteria. Periodic safety analyses are not meant for qualification of the plant components. Separate analyses are devoted to: start-up, qualification of components and materials, and aging. All these analyses are described in this presentation. The last chapter describes the experience obtained for PWR-900 and PWR-1300 units from 1986-1989

  14. Periodic table of elements

    International Nuclear Information System (INIS)

    Following a recommendation by the International Union for Pure and Applied Chemistry (IUPAC), the groups of the periodic table shall be numbered from 1 to 18, instead of I to VIII as before. The recommendations has been approved of by the Committee on Nomenclature of the American Chemical Society. The new system abandons the distinction between main groups (a) and auxiliary groups (b), which in the past frequently has been the reason for misunderstandings between European and American chemists, due to different handling. The publishing house VCH Verlagsgesellschaft recently produced a new periodic table that shows the old and the new numbering system together at a glance, so that chemists will have time to get familiar with the new system. In addition the new periodic table represents an extensive data compilation arranged by elements. The front page lists the chemical properties of elements, the back page their physical properties. (orig./EF)

  15. Periodically kicked turbulence

    Science.gov (United States)

    Lohse

    2000-10-01

    Periodically kicked turbulence is theoretically analyzed within a mean-field theory. For large enough kicking strength A and kicking frequency f the Reynolds number grows exponentially and then runs into some saturation. The saturation level Re(sat) can be calculated analytically; different regimes can be observed. For large enough Re we find Re(sat) approximately Af, but intermittency can modify this scaling law. We suggest an experimental realization of periodically kicked turbulence to study the different regimes we theoretically predict and thus to better understand the effect of forcing on fully developed turbulence. PMID:11089041

  16. Periodic Chandrasekhar recursions

    OpenAIRE

    Aknouche, Abdelhakim; Hamdi, Fayçal

    2007-01-01

    This paper extends the Chandrasekhar-type recursions due to Morf, Sidhu, and Kailath "Some new algorithms for recursive estimation in constant, linear, discrete-time systems, IEEE Trans. Autom. Control 19 (1974) 315-323" to the case of periodic time-varying state-space models. We show that the S-lagged increments of the one-step prediction error covariance satisfy certain recursions from which we derive some algorithms for linear least squares estimation for periodic state-space models. The p...

  17. Higher charge periodic monopoles

    OpenAIRE

    Maldonado, Rafael

    2013-01-01

    We consider singly periodic solutions to the SU(2) Bogomolny equations and use the Nahm transform to generate a class of monopoles of charge k>2, thereby extending known results for lower charge chains. Some simple scattering processes are presented and a comparison made with geodesic motion of monopoles in $\\mathbb{R}^3$.

  18. Ayurveda during Abbasid's period.

    Science.gov (United States)

    Husain, S A; Subhaktha, P K

    2000-01-01

    This is a historical paper which deals with a brief account of Abbasid's period. In this article the existence of Ayurveda in Arab countries, arrival of Ayurvedic physicians to Baghdad, their eminence, authenticity and literary additions in medical field has been studied and presented. PMID:12578013

  19. Periodic paralysis complicating malaria.

    OpenAIRE

    Senanayake, N; Wimalawansa, S J

    1981-01-01

    Episodic muscular weakness, commonly associated with alterations of serum potassium, is the cardinal feature of periodic paralysis. The combination of transient hyperkalaemia and rigors occurring during febrile episodes of malaria is suggested as the underlying cause which precipitated the muscular paralysis. Three patients with malaria who developed a similar paralysis during the paroxysms of fever are described to illustrate this.

  20. Treatment for periodic paralysis

    NARCIS (Netherlands)

    Sansone, [No Value; Meola, G.; Links, T. P.; Panzeri, M.; Rose, M. R.

    2008-01-01

    Background Primary periodic paralyses are rare inherited muscle diseases characterised by episodes of flaccid weakness affecting one or more limbs, lasting several hours to several days, caused by mutations in skeletal muscle channel genes. Objectives The objective of this review was to systematical

  1. A Modern Periodic Table.

    Science.gov (United States)

    Herrenden-Harker, B. D.

    1997-01-01

    Presents a modern Periodic Table based on the electron distribution in the outermost shell and the order of filling of the sublevels within the shells. Enables a student to read off directly the electronic configuration of the element and the order in which filling occurs. (JRH)

  2. Periodic Table of Students.

    Science.gov (United States)

    Johnson, Mike

    1998-01-01

    Presents an exercise in which an eighth-grade science teacher decorated the classroom with a periodic table of students. Student photographs were arranged according to similarities into vertical columns. Students were each assigned an atomic number according to their placement in the table. The table is then used to teach students about…

  3. Almost periodic Schroedinger operators

    International Nuclear Information System (INIS)

    These lectures are devoted to recent developments in the theory of almost-periodic Schroedinger Operators. We specially describe the algebraic point of view, with applications to gap-labelling theorems. Particular models are also presented which exhibit various spectral properties. (orig.)

  4. Scheduling: Seven Period Day

    Science.gov (United States)

    Williamson, Ronald

    2010-01-01

    Driven by stable or declining financial resources many school districts are considering the costs and benefits of a seven-period day. While there is limited evidence that any particular scheduling model has a greater impact on student learning than any other, it is clear that the school schedule is a tool that can significantly impact teacher…

  5. 四川盆地中—上寒武统层序地层划分与沉积模式%Division and Sedimentary Model of Middle-Upper Cambrian Sequence Strata in Sichuan Basin

    Institute of Scientific and Technical Information of China (English)

    袁立; 姚君波; 李国蓉; 彭博; 麻宇杰

    2013-01-01

    According to the observation of the field outcrop profiles at Fandian,Yankong and Sanhuichang,and the sequence boundary characters and succession analysis of two connecting well sections cross Sichuan Basin,the Cambrian strata can be divided into 12 third-order sequences,named as SQ1 to SQ12 from lower to upper,which include SQ1 to SQ6 in lower Cambrian Series and SQ7 to SQ12 in middle to upper Cambrian Series.Several shallowing-upward cycles develop in the middle-upper Cambrian strata,which include lagoons to platform inner shoals,lagoons to tidal flats,and platform inner shoals to tidal flats.The middle-upper Cambrian sediments are the tidal flat-platform inner shoals-lagoon facies from west to east and the thickness of sediments show a varying pattern that is thick in the east and thin in the west,which is complementary to Lower Cambrian sediments.The total thickness of HST is thicker than that of TST in middle-upper Cambrian strata (SQ7-SQ12) so that it is generally characterized of fast transgression and slow regression.Semi-closed shallow-water carbonate platform gradually formed in the east and shoreline developed in the west in the SQ7 during middle Cambrian epoch.The middle Cambrian sedimentary model was inherited in SQ8 to SQ12 during upper Cambrian but more platform inner shoals developed in the west due to the weakening influence of the terrestrial detritus input and shallowing water.The platform inner shoals generally develop in the middle to top parts of HST invertical and in the center and eastern parts of Sichuan Basin in plan.%据对四川盆地范店、岩孔以及三汇场等3个野外寒武系剖面的观察,特别是根据岩孔单井剖面和女基井—宫深1井等连井剖面的层序界面特征和序列分析,将四川盆地寒武系自下而上划分为SQ1、SQ2、……、SQ12共12个三级层序,其中下寒武统6个(SQ1-SQ6),中—上寒武统6个(SQ7-SQ12).中—上寒武统发育多个潟湖→台内滩、潟湖→潮坪、台内

  6. Cells anticipate periodic events

    Science.gov (United States)

    Nakagaki, Toshiyuki

    2009-03-01

    We show that an amoeboid organism can anticipate the timing of periodic events. The plasmodium of the true slime mold Physarum polycephalum moves rapidly under favourable conditions, but stops moving when transferred to less-favourable conditions. Plasmodia exposed to unfavourable conditions, presented in three consecutive pulses at constant intervals, reduced their locomotive speed in response to each episode. When subsequently subjected to favourable conditions, the plasmodia spontaneously reduced their locomotive speed at the time point when the next unfavourable episode would have occurred. This implied anticipation of impending environmental change. After this behaviour had been evoked several times, the locomotion of the plasmodia returned to normal; however, the anticipatory response could subsequently be induced by a single unfavourable pulse, implying recall of the memorized periodicity. We explored the mechanisms underlying these behaviours from a dynamical systems perspective. Our results hint at the cellular origins of primitive intelligence and imply that simple dynamics might be sufficient to explain its emergence.

  7. Notes on Periodic Solitons

    CERN Document Server

    Bakas, Ioannis

    2002-01-01

    We consider static solutions of the sine-Gordon theory defined on a cylinder, which can be either periodic or quasi-periodic in space. They are described by the different modes of a simple pendulum moving in an inverted effective potential and correspond to its libration or rotation. We review the decomposition of the solutions into an oscillatory sum of alternating kinks and anti-kinks or into a monotonic train of kinks, respectively, using properties of elliptic functions. The two sectors are naturally related to each other by a modular transformation, whereas the underlying spectral curve of the model can be used to express the energy of the static configurations in terms of contour integrals \\`a la Seiberg-Witten in either case. The stability properties are also examined by means of supersymmetric quantum mechanics, where we find that the unstable configurations are associated to singular superpotentials, thus allowing for negative modes in the spectrum of small fluctuations.

  8. Thyrotoxic periodic paralysis

    International Nuclear Information System (INIS)

    A case of thyrotoxic periodic paralysis is reported in a Hispanic man with an unusual recurrence six weeks after radioactive iodine treatment. Thyrotoxic periodic paralysis has now been well characterized in the literature: it occurs primarily in Orientals with an overwhelming male preponderance and a higher association of specific HLA antigens. Clinical manifestations include onset after high carbohydrate ingestion or heavy exertion, with progressive symmetric weakness leading to flaccid paralysis of the extremities and other muscle groups, lasting several hours. If hypokalemia is present, potassium administration may help abort the attack. Although propranolol can be efficacious in preventing further episodes, the only definitive treatment is establishing a euthyroid state. The pathophysiology is still controversial, but reflects altered potassium and calcium dynamics as well as certain morphologic characteristics within the muscle unit itself

  9. Hypokalemic periodic paralysis

    OpenAIRE

    Abbas, Haider; Kothari, Nikhil; Bogra, Jaishri

    2012-01-01

    Hypokalemic periodic paralysis is a rare genetic disorder characterized by recurrent attacks of skeletal muscle weakness with associated hypokalemia which is precipitated by stress, cold, carbohydrate load, infection, glucose infusion, hypothermia, metabolic alkalosis, anesthesia, and steroids. We encountered one such incidence of prolonged recovery after general anesthesia, which on further evaluation revealed a case of hypokalemic paralysis. The key to successful management of such a patien...

  10. Scattering of periodic solitons

    Energy Technology Data Exchange (ETDEWEB)

    Cova, R.J. [Carleton University, School of Mathematics and Statistics, 1125 Colonel by Drive, Ottawa, Ontario K1S 5B6 (Canada); Zakrzewski, W.J. [University of Durham, Dept. of Mathematical Sciences, Durham DH1 3LE (United Kingdom)]. e-mail: rcova@math.carleton.ca

    2004-07-01

    Through numerical simulations we study N-soliton scattering (N = 3, 4) in the (2 + 1)-dimensional CP{sup 1} model with periodic boundary conditions. Solitons colliding from symmetrical configurations scatter at {pi}/ N, as observed in the usual model with standard boundary conditions. When the initial configurations are not symmetric the angles differ from {pi}/ N. We describe our observed patterns based on a properly formulated geodesic approximation. (Author) 11 refs., 10 figs.

  11. Chronology of neoproterozoic-cambrian granitic magmatism in the Aracuai Belt, Eastern Brazil, based on single zircon evaporating dating

    International Nuclear Information System (INIS)

    Granitic magmatism related to the orogenic stages of the Aracuai Belt took place at 595-575 Ma, and are represented by two distinct suites. One is composed of I-type granitoids and includes the following plutons: Brasilandia (595±3 Ma), Sao Vitor (576±4 Ma) and Guarataia (574± 2 Ma). The other suite comprises S-type granites like the Ataleia (591±5 Ma) and Wolf (582±5 Ma) plutons. After a long period of magnetic quiescence, a batholith composed of the Caladao granite and Padre Paraiso charnockite intruded at 519±2 Ma. This magmatic episode is probably associated to the collapse of the orogen. (author)

  12. Microthermometric measurement of fluid inclusions and its constraints on genesis of PGE-polymetallic deposits in Lower Cambrian black rock series, southern China

    Institute of Scientific and Technical Information of China (English)

    WANG Min; SUN Xiaoming; MA Mingyang

    2005-01-01

    Systematic microthermometric measurements of fluid inclusions in the PGE-polymetallic deposits hosted in the Lower Cambrian black rock series in southern China were performed, and the results suggest: (1) there exist two types of fluid inclusions. TypeⅠis of NaCl-H2O system with low-medium salinity, and its homogenization temperatures (Th) and salinities are 106.9- 286.4℃ and ( 0.8- 21.8) wt%NaCl eq. respectively; TypeⅡ is of CaCl2-NaCl-H2O system with medium-high salinities, and its homogenization temperatures and salinities range from 120.1℃ to 269.6℃ and ( 11.4- 31.4) wt%NaCl eq., respectively. The typeⅡ fluid inclusions have been discovered for the first time in this kind of deposits; (2) two generations of ore-forming fluids were recognized. Characteristics of fluid inclusions in the PGE-polymetallic ores and carbonate-quartz stockworks in the underlying phosphorites are almost of no difference, they may represent ore-forming fluids at the main metallogenic stage. The peak value of homogenization temperature of those fluid inclusions is about 170℃, while their salinities possess a remarkable bimodal distribution pattern with two peak values of (27-31) wt%NaCl eq. and (4-6) wt%NaCl eq. On the contrary, fluid inclusions in the carbonate-quartz veins in the hanging wall may represent ore-forming fluids at the post-metallogenetic stage. The homogenization temperatures and the peak values of salinities are mostly 130-170℃ and (12-14) wt%NaCl eq., respectively; (3) nobel gas isotopic composition analyses in combination with the microthermometric measurements of fluid inclusions suggest that the ore-forming fluids at the main metallogenetic stage were probably derived from mixing of basinal hot brines with the CaCl2-NaCl-H2O system and seawater with the NaCl-H2O system; (4) in the Early Cambrian, the basinal hot brines were trapped in the Caledonian basins, which were distributed along the southern margin of the Yangtze Craton, and where giant thick

  13. Palaeoenvironmental and geochemical approach of Archaeocyath-rich facies from Lower Cambrian of Western Gondwana margin at Central Iberian Zone (Urda, Toledo Mountains, Spain)

    Science.gov (United States)

    Menéndez, Silvia; Rodríguez-Martínez, Marta; Moreno-Eris, Elena; Perejón, Antonio; Reitner, Joachim

    2010-05-01

    Archaeocyath-rich facies are located in a quarry close to Urda village, at Toledo Mountains, Spain. The outcrops belong to the Caliza de los Navalucillos Formation and they record a considerably high diverse archaeocyath assemblage in the Lower Cambrian successions from the Central Iberian Zone (Julivert et al. 1972 [1974]). In fact, it is first time recorded the presence of Agyrekocyathus, Dokidocyathus, and Plicocyathus in the Central Iberian Zone. Therefore Plicocyathus is no longer exclusive to biozone VI in Spain. The presence of Anthomorpha is characteristic for the early Botomian, presently early Stage 4 (ICS, 2009), and the assemblage corresponds to the biozone VII (late Ovetian, following the biozonation of Perejón & Moreno-Eiris, 2006). The fossiliferous part of the succession is formed by seven lithofacies, all of them tectonically folded and with a low grade metamorphic overprint. They are comprised by two main groups of facies: (a) mound-shaped to massive lithofacies (A1, A2, A3, A4) and (b) massive to bedded and nodular lithofacies (B1, B2, B3). Archaeocyaths occur in several facies: (A1) mound-shaped white marble with irregular to stromatactoid cavities; (A2) massive mottled white to grey limestone; (A3) massive grey limestone with slumps levels; (A4) massive archaeocyath-rich orange limestone; as well as in carbonate nodules embedded in siltstones and cherts (B1, B2 and B3). The best preserved assemblage comes from the nodule record, where fossils are partially pyritized. This type of preservation is exceptional and has never been described before. XRD and wavelength-dispersive electron microprobe analyses reveal the presence of pyrite and pyrrotine partially altered to iron oxides and hydroxides (hematite and goethite) surrounding the archaeocyath cups. In Central Iberian Zone, the development of mounds and nodular facies like those described here is unusual, although the Botomian marks the peak for Early Cambrian archaeocyathan-microbial mounds

  14. The Neoproterozoic-Cambrian Paraguay Belt, central Brazil: Part I - New structural data and a new approach on the regional implications

    Science.gov (United States)

    Silva, Luiz José Homem D'el-Rey; Walde, Detlef Hans-Gerd; Saldanha, Davi Oliveira

    2016-04-01

    Together with the Araguaia and Brasília belts, the Paraguay belt forms in central Brazil, the Tocantins Province that is one of the largest orogens of western Gondwana. The Corumbá area occupies the site where the northern and southern parts of the Paraguay belt form, together with the Chiquitos-Tucavaca aulacogen (stretching E-W in the adjacent Bolivian territory) an R-R-R basin system opened-filled in the ~ 700/650-540 Ma interval within the Amazon-Rio Apa paleo-continent. The sedimentary (volcanic) rocks of the Jacadigo and Corumbá Groups found around the Corumbá city record part of the Neoproterozoic-Cambrian passive margin precursor of the Paraguay belt. Our pioneer structural analysis reveals that these rocks experienced progressive deformation (phases D1-D2-D3) and low-grade metamorphism during the Brasiliano Cycle (540-513 Ma). The crystalline basement was also involved, according to structural data and K-Ar ages in the literature. The paleo-passive margin was thickened during the D1-D2 deformation and was lately shortened (D3) in two orthogonal directions, SE-NW (D3P) and SW-NE (D3T). Developed co-axially and verging to NW, D1-D2-D3P structures record the closure of the basin precursor of the Paraguay belt, whereas D3T structures seem related to the inversion of the aulacogen. Although the tectonic transport to NW, as observed in the Corumbá area, matches the reported transport of Paraguay belt's supracrustal rocks towards the eastern margin of the Rio Apa block and Araguaia belt's rocks towards the Amazon craton, the transport direction is opposite in other parts of the Paraguay belt. Our comprehensive discussion of these facts brings to light profound regional implications.

  15. Petroleum source rock evaluation of the Alum and Dictyonema Shales (Upper Cambrian-Lower Ordovician) in the Baltic Basin and Podlasie Depression (eastern Poland)

    Science.gov (United States)

    Kosakowski, Paweł; Kotarba, Maciej J.; Piestrzyński, Adam; Shogenova, Alla; Więcław, Dariusz

    2016-05-01

    We present geochemical characteristics of the Lower Palaeozoic shales deposited in the Baltic Basin and Podlasie Depression. In the study area, this strata are represented by the Upper Cambrian-Lower Ordovician Alum Shale recognized in southern Scandinavia and Polish offshore and a equivalent the Lower Tremadocian Dictyonema Shale from the northern Estonia and the Podlasie Depression in Poland. Geochemical analyses reveal that the Alum Shale and Dictyonema Shale present high contents of organic carbon. These deposits have the best source quality among the Lower Palaeozoic strata, and they are the best source rocks in the Baltic region. The bituminous shales complex has TOC contents up to ca. 22 wt%. The analysed rocks contain low-sulphur, oil-prone Type-II kerogen deposited in anoxic or sub-oxic conditions. The maturity of the Alum and Dictyonema Shales changes gradually, from the east and north-east to the west and south-west, i.e. in the direction of the Tornquist-Teisseyre Zone. Samples, located in the seashore of Estonia and in the Podlasie region, are immature and in the initial phase of "oil window". The mature shales were found in the central offshore part of the Polish Baltic Basin, and the late mature and overmature are located in the western part of the Baltic Basin. The Alum and Dictyonema Shales are characterized by a high grade of radioactive elements, especially uranium. The enrichment has a syngenetic or early diagenetic origin. The measured content of uranium reached up to 750 ppm and thorium up to 37 ppm.

  16. The Late Cambrian Takaka Terrane, NW Nelson, New Zealand: Accretionary-prism development and arc collision followed by extension and fan-delta deposition at the SE margin of Gondwana

    Science.gov (United States)

    Pound, K. S.

    2013-12-01

    Re-evaluation of field and lab data indicates that the Cambrian portion of the Takaka Terrane in the Cobb Valley area of NW Nelson, New Zealand preserves the remnants of an accretionary prism complex, across which the Lockett Conglomerate fan-delta was deposited as a consequence of extension. Previous work has recognized that the structurally disrupted lower Takaka Terrane rocks present an amalgam of sedimentary and igneous rocks generated prior to convergence (Junction Formation) or during convergence (Devil River Volcanics Group, Haupiri Group), including arc-related and MORB components. Portions of the sequence have in the past been loosely described as an accretionary prism. Reevaluation of the detailed mapping, sedimentological and provenance studies shows that remnants of a stratigraphic sequence (Junction Formation, Devil River Volcanics Group, Haupiri Group) can be traced through 10 fault-bounded slices, which include a mélange-dominated slice (Balloon Mélange). These slices are the remnants of the accretionary prism; the stratigraphy within each slice generally youngs to the east, and the overall pattern of aging (based on relative age from provenance studies, sparse fossils, stratigraphic relations, and limited isotopic data) indicates that the older rocks generally dominate fault slices to the east, and younger rocks dominate fault slices to the west, delineating imbricate slices within an eastward-dipping subduction zone, in which the faults record a complex history of multi-phase reactivation. The Lockett Conglomerate is a ~500-m thick fan-delta conglomerate that is the preserved within one of the fault slices, where it is stratigraphically and structurally highest unit in the lower Takaka Terrane; it is also present as blocks within the Balloon Melange. The Lockett Conglomerate is marine at its base and transitions upwards to fluvial facies. The Lockett Conglomerate has previously been interpreted to result from erosion consequent on continued

  17. Corrigendum to "Isotopic and geochemical characterization of fossil brines of the Cambrian Mt. Simon sandstone and Ironton-Galesville formation from the Illinois Basin, USA" [Geochim. Cosmochim. Acta 165 (2015) 342-360

    Science.gov (United States)

    Labotka, Dana M.; Panno, Samuel V.; Locke, Randall A.; Freiburg, Jared T.

    2016-08-01

    The original Fig. 4 incorrectly represented data from Clayton et al. (1966). The deuterium values were reported in percent deuterium and mistaken by the authors as per mille. The corrected Fig. 4 Corrigendum is given and shows the data from Clayton et al. (1966) plotting in a similar manner as other published data for groundwater in the Illinois Basin. The data from Clayton et al. (1966) was not used in the discussion of the deep-seated Cambrian brines, and, therefore, this misrepresentation does not affect the conclusions of the original manuscript. The authors apologize for the oversight.

  18. Periodic Mesoporous Organosilica Nanorice

    Directory of Open Access Journals (Sweden)

    Mohanty Paritosh

    2008-01-01

    Full Text Available Abstract A periodic mesoporous organosilica (PMO with nanorice morphology was successfully synthesized by a template assisted sol–gel method using a chain-type precursor. The PMO is composed of D and T sites in the ratio 1:2. The obtained mesoporous nanorice has a surface area of 753 m2 g−1, one-dimensional channels, and a narrow pore size distribution centered at 4.3 nm. The nanorice particles have a length of ca. 600 nm and width of ca. 200 nm.

  19. Polysheroidal periodic functions

    International Nuclear Information System (INIS)

    Separation of variables in the Helmholtz N-dimensional (N≥4) equation in polyspheroidal coordinate systems leads to the necessity of solving equations going over into equations for polyspheroidal periodic functions used for solving the two-centre problem in quantum mechanics, the three-body problem with Coulomb interaction, etc. For these functions the expansions are derived in terms of the Jacobi polynomials and Bessel functions. Their basic properties, asymptotics are considered. The algorithm of their computer calculations is developed. The results of numerical calculations are given

  20. Scattering of periodic solitons

    CERN Document Server

    Cova, R J

    2003-01-01

    With the help of numerical simulations we study N-soliton scattering (N=3,4) in the (2+1)-dimensional CP^1 model with periodic boundary conditions. When the solitons are scattered from symmetrical configurations the scattering angles observed agree with the earlier \\pi/N predictions based on the model on R_2 with standard boundary conditions. When the boundary conditions are not symmetric the angles are different from \\pi/N. We present an explanation of our observed patterns based on a properly formulated geodesic approximation.

  1. New Re-Os Isotopic Constrains on the Formation of the Metalliferous Deposits of the Lower Cambrian Niutitang Formation

    Institute of Scientific and Technical Information of China (English)

    Yong Fu; Lin Dong; Chao Li; Wenjun Qu; Haoxiang Pei; Wenlang Qiao; Bing Shen

    2016-01-01

    ABSTRACT:The Terreneuvian Epoch (541–521 Ma)is also an important period for metallogenesis in South China, as was represented by the widespread occurrences of Ni-Mo polymetallic layers on the antecedent shallow platform margin and the V-enriched black shales in deeper slope-basin settings. In this study, we have measured Re-Os isochron ages of Ni-Mo polymetallic layers (Songlin, Niuchang, Sancha, Chuanpengwan), V-rich black shales (Bahuang), and non-metalliferous black shales (Shuidong) in the basal Niutitang Formation in Guizhou and Hunan province, South China. The Ni-Mo polymetal-lic layers and V-enriched black shales have similar Re-Os isochron ages, suggesting concurrent deposi-tion of these two types of metalliferous ores. This suggestion is consistent with the traditional strati-graphic correlation by using the nodular phosphorite bed directly underlying these metalliferous layers as a stratigraphic marker. Furthermore, the metalliferous ores and non-metalliferous black shales have similar initial 187Os/188Os ratios of 0.8–0.9, arguing for a dominant seawater origin with minor contri-bution of hydrothermal activity. Furthermore, Re-Os isotopic data also imply that Ni-Mo and V ore might have derived from the same source. We suggest that the spatial distribution of metalliferous ores can be explained by the development of non-sulfidic anoxic-suboxic wedge (NSASW) in the slope-basin and sulfidic wedge in the previous platform margin. Upwelling of deep water first transects the mildly reduced, organic rich NSASW, in which V (V) is reduced to V (IV), and is preferentially removed from seawater by organometallic complex formation. As a result, V-rich black shale deposits in the slope-basin of Yangtze Platform. Further movement of deep water into the sulfidic platform margin results in Ni-Mo polymetallic layer formation.

  2. Periods and Superstring Amplitudes

    CERN Document Server

    Stieberger, S

    2016-01-01

    Scattering amplitudes which describe the interaction of physical states play an important role in determining physical observables. In string theory the physical states are given by vibrations of open and closed strings and their interactions are described (at the leading order in perturbation theory) by a world-sheet given by the topology of a disk or sphere, respectively. Formally, for scattering of N strings this leads to N-3-dimensional iterated real integrals along the compactified real axis or N-3-dimensional complex sphere integrals, respectively. As a consequence the physical observables are described by periods on M_{0,N} - the moduli space of Riemann spheres of N ordered marked points. The mathematical structure of these string amplitudes share many recent advances in arithmetic algebraic geometry and number theory like multiple zeta values, single-valued multiple zeta values, Drinfeld, Deligne associators, Hopf algebra and Lie algebra structures related to Grothendiecks Galois theory. We review the...

  3. Transition from non-periodic to periodic explosions.

    Science.gov (United States)

    Cartes, Carlos; Descalzi, Orazio

    2015-12-13

    We show the existence of periodic exploding dissipative solitons. These non-chaotic explosions appear when higher-order nonlinear and dispersive effects are added to the complex cubic-quintic Ginzburg-Landau equation modelling soliton transmission lines. This counterintuitive phenomenon is the result of period-halving bifurcations leading to order (periodic explosions), followed by period-doubling bifurcations (or intermittency) leading to chaos (non-periodic explosions). PMID:26527807

  4. Periodic and limit-periodic discrete Schr\\"odinger operators

    OpenAIRE

    Krueger, Helge

    2011-01-01

    The theory of discrete periodic and limit-periodic Schr\\"odinger operators is developed. In particular, the Floquet--Bloch decomposition is discussed. Furthermore, it is shown that an arbitrarily small potential can add a gap for even periods. In dimension two, it is shown that for coprime periods small potential terms don't add gaps thus proving a Bethe--Sommerfeld type statement. Furthermore limit-periodic potentials whose spectrum is an interval are constructed.

  5. 四川盆地南部地区寒武系油气勘探前景%Oil and Gas Exploration Prospects of Cambrian in Southern Sichuan Basin

    Institute of Scientific and Technical Information of China (English)

    王佳; 刘树根; 黄文明; 张长俊; 曾祥亮

    2011-01-01

    为明确四川盆地南部地区寒武系常规油气和页岩气的勘探前景,应用岩相学和地球化学分析手段相结合,对研究区中上寒武统娄山组和下寒武统清虚洞组的厚度、孔隙度、渗透率、孔隙类型和下寒武统筇竹寺组的厚度、总有机碳质量分数w(TOC)、镜质体反射率Ro、矿物组分等进行了深入剖析.结果表明:①川南地区烃源岩厚度大(80~100m),w(TOC)高(3%~5%),为一套中—好的过成熟后期阶段烃源岩(Ro为3.0%~4.8%),矿物组分与美国重要页岩气产层Barnett页岩相似,其中石英、长石和黄铁矿的体积分数为31%~50%,黏土矿物体积分数为29%~53%;②威远地区下寒武统九老洞组黑色页岩含气量达0.43~0.86 m3/t;微孔隙发育,主要为粒间孔,孔隙直径达5~10μm,裂缝密度一般为30~50条/m;③中上寒武统娄山组和下寒武统清虚洞组总体上是寒武系最重要的常规储层,储层内部沥青质量分数一般为3%~6%,储集空间主要为粒间溶孔—晶间孔等,平均孔隙度为1.85%,平均渗透率为2.25×10-3μm2;④在川东南地区靠近盆地边缘的斜坡带,寒武系具有较好的常规油气藏勘探前景,而川东南缘和川西南地区筇竹寺组下部具有较大的页岩气勘探潜力.%In order to identify the exploration prospect of Cambrian conventional oil and gas and shale gas in southern Sichuan Basin, this paper, combining the petrographic and geochemical methods, makes a tho-rugh analysis of the thickness, porosity, permeability, porosity type of Middle-Upper Cambrian Loushan-guan Formation and Lower Cambrian Qingxudong Formation, as well as the thickness, TOC, Ro, mineral composition of Lower Cambrian Qiongzhusi Formation. The results show that: (1) The hydrocarbon source rocks in southern Sichuan Basin are thick(80 -100 m)and of high organic matter content (3%-5%). It is a set of general-good late-mature stage of hydrocarbon source

  6. Soft-sediment deformation structures in Cambrian Series 2 tidal deposits (NW Estonia): implications for identifying endogenic triggering mechanisms in ancient sedimentary record

    Science.gov (United States)

    Põldsaar, Kairi

    2015-04-01

    Soft-sediment deformation structures (SSDS) are documented in several horizons within silt- and sandstones of the Cambrian Series 2 (Dominopolian Stage) Tiskre Formation, and some in the below-deposited argillaceous deposits of the Lükati Formation (northern part of the Baltoscandian Palaeobasin, NW Estonia). The aim of this study was to map, describe, and analyze these deformation features, discuss their deformation mechanism and possible triggers. Load structures (simple load casts, pillows, flame structures, convoluted lamination) with varying shapes and sizes occur in the Tiskre Fm in sedimentary interfaces within medium-bedded peritidal rhythmites (siltstone-argillaceous material) as well as within up to 3 m thick slightly seaward inclined stacked sandstone sequences. Homogenized beds, dish-and-pillar structures, and severely deformed bedding are also found within these stacked units and within a large tidal runoff channel infill. Autoclastic breccias and water-escape channels are rare and occur only in small-scale -- always related to thin, horizontal tidal laminae. Profound sedimentary dykes, sand volcanoes, and thrust faults, which are often related to earthquake triggered soft sediment deformation, were not observed within the studied intervals. Deformation horizon or horizons with large flat-topped pillows often with elongated morphologies occur at or near the boundary between the Tiskre and Lükati formations. Deformation mechanisms identified in this study for the various deformation types are gravitationally unstable reversed density gradient (especially in case of load features that are related to profound sedimentary interfaces) and lateral shear stress due to sediment current drag (in case of deformation structures that not related to loading at any apparent sedimentary interface). Synsedimentary liquefaction was identified as the primary driving force in most of the observed deformation horizons. Clay thixotropy may have contributed in the

  7. On solving periodic Riccati equations

    OpenAIRE

    Varga, Andreas

    2008-01-01

    Numerically reliable algorithms to compute the periodic non-negative definite stabilizing solutions of the periodic differential Riccati equation (PRDE) and discrete-time periodic Riccati equation (DPRE) are proposed. For the numerical solution of PRDEs, a new multiple shooting-type algorithm is developed to compute the periodic solutions in an arbitrary number of time moments within one period by employing suitable discretizations of the continuous-time problems. In contrast to single shooti...

  8. The redoubtable ecological periodic table

    Science.gov (United States)

    Ecological periodic tables are repositories of reliable information on quantitative, predictably recurring (periodic) habitat–community patterns and their uncertainty, scaling and transferability. Their reliability derives from their grounding in sound ecological principle...

  9. A new characterization of periodic oscillations in periodic difference equations

    Energy Technology Data Exchange (ETDEWEB)

    Al-Salman, Ahmad [Department of Mathematics and Statistics, Sultan Qaboos University, P.O. Box 36, PC 123, Al-Khod (Oman); AlSharawi, Ziyad, E-mail: alsha1zm@alsharawi.info [Department of Mathematics and Statistics, Sultan Qaboos University, P.O. Box 36, PC 123, Al-Khod (Oman)

    2011-11-15

    Highlights: > A characterization of periodic solutions of periodic difference equations is given. > The periods are classified as multiples and non-multiples of the phase-period. > Non-multiples of the phase-period depend on the intersections between the maps. > A refinement of Sharkovsky's theorem for periodic difference equations is given. - Abstract: In this paper, we characterize periodic solutions of p-periodic difference equations. We classify the periods into multiples of p and nonmultiples of p. We show that the elements of the set of multiples of p follow the well-known Sharkovsky's ordering multiplied by p. On the other hand, we show that the elements of the set {Gamma}{sub p} of nonmultiples of p are independent in their existence. Moreover, we show the existence of a p-periodic difference equation with infinite {Gamma}{sub p}-set in which the maps are defined on a compact domain and agree exactly on a countable set. Based on the proposed classification, we give a refinement of Sharkovsky's theorem for periodic difference equations.

  10. Periods and elementary real numbers

    OpenAIRE

    Yoshinaga, Masahiko

    2008-01-01

    The periods, introduced by Kontsevich and Zagier, form a class of complex numbers which contains all algebraic numbers and several transcendental quantities. Little has been known about qualitative properties of periods. In this paper, we compare the periods with hierarchy of real numbers induced from computational complexities. In particular we prove that periods can be effectively approximated by elementary rational Cauchy sequences. As an application, we exhibit a computable real number wh...

  11. Periodic waves in nonlinear metamaterials

    International Nuclear Information System (INIS)

    Periodic waves are presented in this Letter. With symbolic computation, equations for monochromatic waves are studied, and analytic periodic waves are obtained. Factors affecting properties of periodic waves are analyzed. Nonlinear metamaterials, with the continuous distribution of the dielectric permittivity obtained, are different from the ones with the discrete distribution. -- Highlights: ► Equations for the monochromatic waves in transverse magnetic polarization have been studied. ► Analytic periodic waves for the equations have been obtained. ► Periodic waves are theoretically presented and studied in the nonlinear metamaterials.

  12. Nonparametric Inference for Periodic Sequences

    KAUST Repository

    Sun, Ying

    2012-02-01

    This article proposes a nonparametric method for estimating the period and values of a periodic sequence when the data are evenly spaced in time. The period is estimated by a "leave-out-one-cycle" version of cross-validation (CV) and complements the periodogram, a widely used tool for period estimation. The CV method is computationally simple and implicitly penalizes multiples of the smallest period, leading to a "virtually" consistent estimator of integer periods. This estimator is investigated both theoretically and by simulation.We also propose a nonparametric test of the null hypothesis that the data have constantmean against the alternative that the sequence of means is periodic. Finally, our methodology is demonstrated on three well-known time series: the sunspots and lynx trapping data, and the El Niño series of sea surface temperatures. © 2012 American Statistical Association and the American Society for Quality.

  13. Reservoir quality and petrophysical properties of Cambrian sandstones and their changes during the experimental modelling of CO2 storage in the Baltic Basin

    Directory of Open Access Journals (Sweden)

    Kazbulat Shogenov

    2015-08-01

    Full Text Available The objectives of this study were (1 to review current recommendations on storage reservoirs and classify their quality using experimental data of sandstones of the Deimena Formation of Cambrian Series 3, (2 to determine how the possible CO2 geological storage (CGS in the Deimena Formation sandstones affects their properties and reservoir quality and (3 to apply the proposed classification to the storage reservoirs and their changes during CGS in the Baltic Basin. The new classification of the reservoir quality of rocks for CGS in terms of gas permeability and porosity was proposed for the sandstones of the Deimena Formation covered by Lower Ordovician clayey and carbonate cap rocks in the Baltic sedimentary basin. Based on permeability the sandstones were divided into four groups showing their practical usability for CGS (‘very appropriate’, ‘appropriate’, ‘cautionary’ and ‘not appropriate’. According to porosity, eight reservoir quality classes were distinguished within these groups. The petrophysical, geochemical and mineralogical parameters of the sandstones from the onshore South Kandava and offshore E6 structures in Latvia and the E7 structure in Lithuania were studied before and after the CO2 injection-like alteration experiment. The greatest changes in the composition and properties were determined in the carbonate-cemented sandstones from the uppermost part of the South Kandava onshore structure. Partial dissolution of pore-filling carbonate cement (ankerite and calcite and displacement of clay cement blocking pores caused significant increase in the effective porosity of the samples, drastic increase in their permeability and decrease in grain and bulk density, P- and S-wave velocity, and weight of the dry samples. As a result of these alterations, carbonate-cemented sandstones of initially ‘very low’ reservoir quality (class VIII, ‘not appropriate’ for CGS, acquired an ‘appropriate’ for CGS

  14. Geochemistry of major and trace elements and Pb–Sr isotopes of a weathering profile developed on the Lower Cambrian black shales in central Hunan, China

    International Nuclear Information System (INIS)

    Highlights: • The chemical changes during black shale weathering are geochemically estimated. • Chemical changes and Pb–Sr data suggest four types of chemical reactions. • There were two stages of geochemical processes during black shale weathering. • Early stage of chemical differentiation led elements leaching and redistribution. • Soil was formed by later chemical homogenisation after the early differentiation. - Abstract: This paper reports a geochemical study on the major and trace elements and Pb–Sr isotopes of a weathering profile developed in the Lower Cambrian black shales in central Hunan (China). Six weathering horizons were identified and sampled vertically throughout the profile. The chemical composition of the profile consists of variable concentrations of the major elements Fe2O3, FeO, MnO, MgO, CaO, Na2O, and P2O5 and of less variable concentrations of SiO2, TiO2, Al2O3, and K2O. The chemical change caused by weathering is estimated by mass-balance calculations, and the results show that the element mobility is characterised by substantial loss of SiO2, FeO, CaO, K2O, Na2O, LOI, Cr, V, Ba, Cs, Rb, Sr, U, and Th, and moderate loss of Al2O3, MgO, Fe2O3, Ni, Cu, Pb, Tl, Sn, Sc, Ge and REE (Y). The high field strength elements TiO2, Sn, Sc, U, Ga, Ge, Zr, Hf, Nb, and Ta were immobile during weathering. The chemical changes and the Pb–Sr isotopic data suggest that four types of chemical reactions occurred: the oxidation of sulphide minerals (e.g., pyrite) and organic carbon (OS), the dissolution of less resistant clinochlore-Ia, calcite, and P-bearing minerals (DL), the dissolution of detrital albite and microcline (DA), and the transformation of clay (TC) minerals (e.g., muscovite and illite–smectite). These chemical reactions then led to two stages of geochemical processes, an early stage of chemical differentiation and a later stage of chemical homogenisation. The chemical differentiation dominated by the OS, DL, and DA reactions

  15. Positive periodic solutions of delayed periodic Lotka-Volterra systems

    Energy Technology Data Exchange (ETDEWEB)

    Lin Wei [Laboratory of Nonlinear Mathematics Science, Institute of Mathematics, Fudan University, Shanghai 200433 (China)]. E-mail: weilin@fudan.edu.cn; Chen Tianping [Laboratory of Nonlinear Mathematics Science, Institute of Mathematics, Fudan University, Shanghai 200433 (China)]. E-mail: tchen@fudan.edu.cn

    2005-01-17

    In this Letter, for a general class of delayed periodic Lotka-Volterra systems, we prove some new results on the existence of positive periodic solutions by Schauder's fixed point theorem. The global asymptotical stability of positive periodic solutions is discussed further, and conditions for exponential convergence are given. The conditions we obtained are weaker than the previously known ones and can be easily reduced to several special cases.

  16. Positive periodic solutions of delayed periodic Lotka-Volterra systems

    Science.gov (United States)

    Lin, Wei; Chen, Tianping

    2005-01-01

    In this Letter, for a general class of delayed periodic Lotka-Volterra systems, we prove some new results on the existence of positive periodic solutions by Schauder's fixed point theorem. The global asymptotical stability of positive periodic solutions is discussed further, and conditions for exponential convergence are given. The conditions we obtained are weaker than the previously known ones and can be easily reduced to several special cases.

  17. Fractional-period excitations in continuum periodic systems

    International Nuclear Information System (INIS)

    We investigate the generation of fractional-period states in continuum periodic systems. As an example, we consider a Bose-Einstein condensate confined in an optical-lattice potential. We show that when the potential is turned on nonadiabatically, the system explores a number of transient states whose periodicity is a fraction of that of the lattice. We illustrate the origin of fractional-period states analytically by treating them as resonant states of a parametrically forced Duffing oscillator and discuss their transient nature and potential observability

  18. The Cambrian initiation of intra-oceanic subduction in the southern Paleo-Asian Ocean: Further evidence from the Barleik subduction-related metamorphic complex in the West Junggar region, NW China

    Science.gov (United States)

    Liu, Bo; Han, Bao-Fu; Xu, Zhao; Ren, Rong; Zhang, Jin-Rui; Zhou, Jing; Su, Li; Li, Qiu-Li

    2016-06-01

    In this study, we present new evidence from the Barleik subduction-related metamorphic complex in the southern West Junggar region, northwestern China, for the Cambrian initiation of intra-oceanic subduction in the southern Paleo-Asian Ocean. The Barleik metamorphic complex is mainly composed of blueschist and amphibolite blocks within an ophiolitic mélange and their protoliths are calc-alkaline andesite and alkali and tholeiitic basalts. The calc-alkaline andesite has a zircon U-Pb age of 502 ± 2 Ma, obtained from magmatic cores of zircon grains, and shares geochemical features similar to the 515-485 Ma intra-oceanic arc magmatic rocks in the West Junggar region. By contrast, the alkali and tholeiitic basalts have trace element features similar to ocean island and enriched mid-ocean ridge basalts, respectively. Rutile and sodic-calcic amphibole from the amphibolite have a U-Pb age of 502 ± 25 Ma and a 40Ar/39Ar age of ∼504 Ma, respectively, which are in good agreement within errors with a 40Ar/39Ar age of 492 ± 4 Ma for phengite from the blueschist. These metamorphic ages of ∼500 Ma are interpreted to represent the timing of Pacific-type subduction-related metamorphism and are also compatible with ages of the oldest supra-subduction zone ophiolites (531-512 Ma) and intra-oceanic arc plutons (515-485 Ma) in the southern West Junggar region. Being one of the oldest subduction-related metamorphic complexes (509-490 Ma) in the southern Central Asian Orogenic Belt, the Barleik metamorphic complex, together with the oldest arc plutons, definitely indicate the initial intra-oceanic subduction in the southern Paleo-Asian Ocean at least in the Early Cambrian.

  19. New Discoveries of Phaeophycean Fossils in the Early Cambrian,Haikou, Kunming,Yunnan, Southwest China%云南海口早寒武世褐藻化石新知

    Institute of Scientific and Technical Information of China (English)

    徐兆良

    2001-01-01

    主要报道了产自云南海口耳材村早寒武世筇竹寺组玉案山段澄江生物群中的褐藻化石——拟点叶藻(Punctariopsis latifolia gen. et sp. nov.)、简单拟点叶藻(Punctariopsis simplex gen. et sp. nov.)和古文德带藻(Vendotaenia cf. antiqua Gnilovskaya) 3种.这些化石丰富了澄江生物群物种的多样性,为研究该群生物爆发性演化提供了新的证据.同时通过对现生褐藻和褐藻化石形态学的比较研究,表明化石属和现代属在系统演化上可能有较密切的亲缘关系,并进一步证明了澄江生物群在云南海口地区发生于一个潮间带和亚潮带的海水环境.%The paper deals with the new phaeophycean fossils in the Early Cambrian from Chengjiang Biota at Ercai Village of Haikou, Kunming, Yunnan, Southwest China. Three new taxa, Punctariopsis latifolia gen. et sp. nov., P.simplex gen. et sp. nov. and Vendotaenia cf. antiqua Gnilovskaya are described here in detail. These new findings show that the Chengjiang Biota is high in species diversity of fossil algae and might offer new evidence for a better understanding of the Cambrian explosive biological evolution. The fossil genus described here is found to have a close relationship with the living Punctaria based on similar morphology. The results suggest that the Chengjiang Biota in Haikou area of Kunming was living in an intertidal and subtidal seawater environment.

  20. 昆明海口早寒武世澄江生物群中叶状红藻化石%Leaf-like Rhodophycean Fossils in the Early Cambrian from Haikou,Kunming,Yunnan of China

    Institute of Scientific and Technical Information of China (English)

    徐兆良

    2004-01-01

    This paper deals with the new leaf-like rhodophycean fossils in the Early Cambrian from Chengjiang Biota at Mafang Village of Haikou, Kunming, Yunnan, Southeast China. The new taxa Paradelesseria sanguinea gen. et sp. nov. is found to have a close relationship with the living Delesseria according to detailed morphological study between the fossil and modern Delesseria. These new findings show that the Chengjiang Biota is high in species diversity of fossil algae and might offer new evidence for a better understanding of the Cambrian explosive biological evolution and its paleoenvironment. The Biota in Haikou district of Kunming was living in subtidal and lower intertidal marine environment, up to 30 m below surface of the water based on the comparative study of extant red algae.%主要报道了采自云南省昆明市海口马房村鞍山早寒武世筇竹寺组玉案山段澄江生物群中的叶状红藻-似红叶藻(新属、新种)(Paradelesseria sanguinea Xu,gen.et sp.Nov.).通过比较形态学研究,探讨了该宏观化石藻类的分类归属与亲缘关系,进一步丰富了澄江生物群生物物种多样性的认识,并为研究早寒武世生物演化及其古环境提供了新的化石证据.同时,通过对现生红藻的比较研究,进一步证明了澄江生物群在云南海口地区发生于一个水深在30 m以上的亚潮带和下潮间带的海水环境.

  1. Repelling periodic points of given periods of rational functions

    Institute of Scientific and Technical Information of China (English)

    CHANG; Jianming; FANG; Mingliang

    2006-01-01

    Let R(z) be a rational function of degree d ≥ 2. Then R(z) has at least one repelling periodic point of given period k ≥ 2, unless k = 4 and d=2, or k= 3 and d ≤ 3, or k=2 and d≤8. Examples show that all exceptional cases occur.

  2. Photolysis of Periodate and Periodic Acid in Aqueous Solution

    DEFF Research Database (Denmark)

    Sehested, Knud; Kläning, U. K.

    1978-01-01

    The photochemistry of periodate and periodic acid in aqueous solution was studied (i) by quantum yield measurements at low light intensity (ii) by flash photolysis, and (iii) by photolysis of glassy samples at 77 K. The photochemical studies were supplemented with pulse radiolysis studies of...

  3. From Periodic Properties to a Periodic Table Arrangement

    Science.gov (United States)

    Besalú, Emili

    2013-01-01

    A periodic table is constructed from the consideration of periodic properties and the application of the principal components analysis technique. This procedure is useful for objects classification and data reduction and has been used in the field of chemistry for many applications, such as lanthanides, molecules, or conformers classification.…

  4. Scaling limits of periodic monopoles

    Science.gov (United States)

    Maldonado, Rafael

    2015-01-01

    The purpose of this note is to explore the structure of singly periodic monopoles for different values of the size to period ratio. The transition between a chain of small monopoles and the approximately two dimensional chain of large monopoles takes us through a region with an unintuitive dependence on the periodic direction. The focus is mainly on the smooth SU(2) monopole of charge 2.

  5. Scaling limits of periodic monopoles

    OpenAIRE

    Maldonado, Rafael

    2014-01-01

    The purpose of this note is to explore the structure of singly periodic monopoles for different values of the size to period ratio. The transition between a chain of small monopoles and the approximately two dimensional chain of large monopoles takes us through a region with an unintuitive dependence on the periodic direction. The focus is mainly on the smooth SU(2) monopole of charge 2.

  6. Ap stars with variable periods

    OpenAIRE

    Mikulášek, Zdeněk; Krtička, Jiří; Janík, Jan; Zejda, Miloslav; Henry, Gegory W.; Paunzen, Ernst; Žižňovský, Jozef; Zverko, Juraj

    2013-01-01

    The majority of magnetic chemically peculiar (mCP) stars exhibit periodic light, magnetic, radio, and spectroscopic variations that can be adequately modelled as a rigidly-rotating main-sequence star with persistent surface structures. Nevertheless, there is a small sample of diverse mCP stars whose rotation periods vary on timescales of decades while the shapes of their phase curves remain unchanged. Alternating period increases and decreases have been suspected in the hot CP stars CU Vir an...

  7. Holonomic systems for period mappings

    Energy Technology Data Exchange (ETDEWEB)

    Chen, Jingyue, E-mail: jychen@brandeis.edu [Department of Mathematics, Brandeis University, Waltham, MA 02454 (United States); Huang, An, E-mail: anhuang@math.harvard.edu [Department of Mathematics, Harvard University, Cambridge, MA 02138 (United States); Lian, Bong H., E-mail: lian@brandeis.edu [Department of Mathematics, Brandeis University, Waltham, MA 02454 (United States)

    2015-09-15

    Period mappings were introduced in the sixties [4] to study variation of complex structures of families of algebraic varieties. The theory of tautological systems was introduced recently [7,8] to understand period integrals of algebraic manifolds. In this paper, we give an explicit construction of a tautological system for each component of a period mapping. We also show that the D-module associated with the tautological system gives rise to many interesting vanishing conditions for period integrals at certain special points of the parameter space.

  8. Possible Impact at Precambrian-Cambrian Boundary and Its Influence on Biosphere%前寒武纪-寒武纪界线附近的撞击假说及其对生物圈的影响

    Institute of Scientific and Technical Information of China (English)

    张维加; Daniel P.Connelly; 俞杭杰

    2010-01-01

    HAPCIS陨击坑的基底构造实地分析也表明了前寒武纪末曾经存在过陨击事件.最后笔者利用计算机模拟了大气圈变化.模拟分两步进行,首先从岩石热分解得出需要的温度与压强数据,而后将数据作为参数用于模拟.数据结果表明撞击区域的大气温度在4 000 K左右,压强为5 600 Bar.撞击增加了大气中二氧化碳和氧气的含量,并强化了臭氧层.这些含量的变化与地质上的记录一致.%After a thorough research on the Earth' s circumstantial changes and the great evolution of life in the Cambrian period, the authors propound such a hypothesis: During the Late Precambrian, about 500 ~ 600 Ma, a celestial body impacted the Earth. The high temperature ended the great glaciation, facilitated the communication of biological information. The rapid change of Earth environment enkindled the genesis-control system, and released the HSP-90 variations. After the impact, benefited from the protection of the new ozone layer and the energy supplement of the aerobic respiration, those survived underground life exploded. They generated carapaces and complex metabolism to adjust to the new circumstance of high temperature and high pressure. That' s the Cambrian explosion. This article uses a large amount of analyses and calculations, and illustrates that this hypothesis fits well with most of the important incidences in astronomic and geologic discoveries.

  9. Quasi-periodicity in relative quasi-periodic tori

    Science.gov (United States)

    Fassò, Francesco; García-Naranjo, Luis C.; Giacobbe, Andrea

    2015-10-01

    At variance from the cases of relative equilibria and relative periodic orbits of dynamical systems with symmetry, the dynamics in relative quasi-periodic tori (namely, subsets of the phase space that project to an invariant torus of the reduced system on which the flow is quasi-periodic) is not yet completely understood. Even in the simplest situation of a free action of a compact and abelian connected group, the dynamics in a relative quasi-periodic torus is not necessarily quasi-periodic. It is known that quasi-periodicity of the unreduced dynamics is related to the reducibility of the reconstruction equation, and sufficient conditions for it are virtually known only in a perturbation context. We provide a different, though equivalent, approach to this subject, based on the hypothesis of the existence of commuting, group-invariant lifts of a set of generators of the reduced torus. Under this hypothesis, which is shown to be equivalent to the reducibility of the reconstruction equation, we give a complete description of the structure of the relative quasi-periodic torus, which is a principal torus bundle whose fibers are tori of a dimension which exceeds that of the reduced torus by at most the rank of the group. The construction can always be done in such a way that these tori have minimal dimension and carry ergodic flow.

  10. Beyond the periodic orbit theory

    OpenAIRE

    Cvitanovic, Predrag; Hansen, Kim; Rolf, Juri; Vattay, Gabor

    1997-01-01

    The global constraints on chaotic dynamics induced by the analyticity of smooth flows are used to dispense with individual periodic orbits and derive infinite families of exact sum rules for several simple dynamical systems. The associated Fredholm determinants are of particularly simple polynomial form. The theory developed suggests an alternative to the conventional periodic orbit theory approach to determining eigenspectra of transfer operators.

  11. Superconductivity and the periodic table

    International Nuclear Information System (INIS)

    Compiled data on superconducting transition temperature Tc for pure metals (including amorphous and high-pressure phases) from the first, second, and third long periods of the Periodic Table and for their analogous compounds show that a significant regularity exists in the relative values of Tc

  12. Periodicities in gamma ray bursts

    International Nuclear Information System (INIS)

    Gamma ray burst models based on magnetic neutron stars face a problem of account for the scarcity of observed periods. Both this scarcity and the typical period found when any is detected are explained if the neutron stars are accreting in binary systems

  13. Transition Period and Immunosuppression: Critical Period of Dairy Cattle Reproduction

    OpenAIRE

    K. Simenew; M. Wondu

    2013-01-01

    This seminar study is prepared on the objectives of: revising important aspects of transition period of dairy cattle and highlighting some potential areas of research and challenges for the future. It has sufficiently been discussed that improved understanding of this frontier of the biology, immunology, nutrition and management of cows during the transition period will provide the largest gains in productivity and profitability of dairy farms. In the manuscript under each specific topic, tra...

  14. Optical Periodicity Analysis of 3C 446 using Period04

    Indian Academy of Sciences (India)

    Fei Guo; Hao Jing Zhang

    2014-09-01

    All the data of the blazar 3C446 at 8, 4.8, 14 and 22 GHz, presented in publications from 1977 to 2006, have been compiled to generate light curves. The light curves show violent activity of 3C446. Using Period04 analysis method, we have found that there is a period of 7.2 yr, which is consistent with the results that we found using wavelet analysis method. We get the instability region as = 123.83.

  15. Entanglement Entropy of Periodic Sublattices

    CERN Document Server

    He, Temple; Vandoren, Stefan

    2016-01-01

    We study the entanglement entropy (EE) of Gaussian systems on a lattice with periodic boundary conditions, both in the vacuum and at nonzero temperatures. By restricting the reduced subsystem to periodic sublattices, we can compute the entanglement spectrum and EE exactly. We illustrate this for a free (1+1)-dimensional massive scalar field at a fixed temperature. Consistent with previous literature, we demonstrate that for a sufficiently large periodic sublattice the EE grows extensively, even in the vacuum. Furthermore, the analytic expression for the EE allows us probe its behavior both in the massless limit and in the continuum limit at any temperature.

  16. The Periodic Table in Croatia

    Directory of Open Access Journals (Sweden)

    Raos, N.

    2011-12-01

    Full Text Available The Croatian (Yugoslav Academy of Sciences and Arts was the first academy to elect D. I. Mendeleev as its honorary member (1882, whereas the periodic table of the elements has been taught regularly at the Zagreb University since 1888. The early interest of Croatian chemists in the periodic table should be attributed primarily to their pan-Slavic attitude, particularly as proof that Slavic people were able to produce "their own Newtons" (M. V. Lomonosov and D. I. Mendeleev. Such enthusiastic views, however, did not help in analyzing the contribution of Mendeleev and other scientists to the discovery and development of the periodic table of the elements.

  17. Modular forms and period polynomials

    CERN Document Server

    Pasol, Vicentiu

    2012-01-01

    We study the space of period polynomials associated with modular forms for finite index subgroups of the modular group. For the full modular group, this space is endowed with a pairing, corresponding to the Petersson inner product on modular forms via a formula of Haberland, and with an action of Hecke operators, defined algebraically by Zagier. We extend Haberland's formula to arbitrary modular forms for finite index subgroups of the modular group, and we show that it conceals two stronger formulas. We extend the action of Hecke operators to \\Gamma_0(N) and \\Gamma_1(N), and we prove algebraically that the pairing on period polynomials appearing in Haberland's formula is Hecke equivariant. Two indefinite theta series identities follow from this proof. We give two ways of determining the extra relations satisfied by the even and odd parts of period polynomials associated with cusp forms, which are independent of the period relations.

  18. Periodicities in Aerosol Optical Depths

    CERN Document Server

    Ramachandran, S; Verma, Amit; Panigrahi, Prasanta K

    2011-01-01

    We investigate the temporal and spatial variability in aerosol optical depth (AOD) over different geographic locations in India due to their important role in the earth-atmosphere radiation budget. The use of continuous wavelet transform pinpoints the spatio-temporal non-stationarity of the periodic variations in the AOD depending on local factors. The optimal time-frequency localization ability of Morlet wavelet accurately isolates the periodic features in the different frequency domains, to study the variations in the dominant periods due to local effects. The origin of the effects on the periodic modulations is then related to physical phenomena of regional nature, which throws considerable light on the observed variations in aerosol optical depths. We also find the phase relationship between different locations and to identify the possible correlations between different geographic locations and related environmental variations.

  19. Periodicities in photospheric magnetic flux

    Institute of Scientific and Technical Information of China (English)

    2006-01-01

    Magnetic field plays an important role in solar structure and activity. In principle, the determination of magnetic flux would provide the best general-purpose index of solar activity. Currently, the periodicity studies corresponding to photospheric magnetic flux (PMF) are very few possibly due to the absence of a uniform flux sequence. In this paper, by using 383 NSO/Kitt Peak magnetic synoptic charts we reconstruct a flux sequence from February 1975 to August 2003 and perform a relatively systemic periodicity analysis with two methods of the Scargle periodogram and the Morlet wavelet transform. As a result, four periods are found at around 1050, 500, 300 and 160 days. We analyze these periods' temporal variabilities in detail and discuss their respective origins briefly.

  20. The Periodic Table in Croatia

    OpenAIRE

    Raos, N.

    2011-01-01

    The Croatian (Yugoslav) Academy of Sciences and Arts was the first academy to elect D. I. Mendeleev as its honorary member (1882), whereas the periodic table of the elements has been taught regularly at the Zagreb University since 1888. The early interest of Croatian chemists in the periodic table should be attributed primarily to their pan-Slavic attitude, particularly as proof that Slavic people were able to produce "their own Newtons" (M. V. Lomonosov and D. I. Mendeleev). Such enthusiasti...

  1. Prime numbers: periodicity, chaos, noise

    OpenAIRE

    Bershadskii, A.

    2011-01-01

    Logarithmic gaps have been used in order to find a periodic component of the sequence of prime numbers, hidden by a random noise (stochastic or chaotic). The recovered period for the sequence of the first 10000 prime numbers is equal to 8\\pm1 (subject to the prime number theorem). For small and moderate values of the prime numbers (first 2000 prime numbers) this result has been directly checked using the twin prime killing method.

  2. Cohort postponement and period measures

    OpenAIRE

    Joshua R. Goldstein; Thomas Cassidy

    2010-01-01

    We introduce a new class of models in which demographic behavior such as fertility is postponed by differing amounts depending only on cohort membership. We show how this model fits into a general framework of period and cohort postponement that includes the existing models in the literature, notably those of Bongaarts and Feeney and Kohler and Philipov. The cohort-based model shows the effects of cohort shifts on period fertility measures and provides an accompanying tempo-adjusted measure o...

  3. Le graben de l'Anti-Atlas occidental (Maroc) : contrôle tectonique de la paléogéographie et des séquences au Cambrien inférieurThe Lower-Cambrian western Anti-Atlasic graben: tectonic control of palaeogeography and sequential organisation

    Science.gov (United States)

    Benssaou, Mohammed; Hamoumi, Naı̈ma

    2003-03-01

    In the Moroccan western Anti-Atlas, the combined extensive tectonic events with a long-term sea-level rise is the main factor on building vertical stacking transgressive-regressive sequences. In the Ait Abdallah-Boussafene axis, the subsidence processes, relayed by a brutal platform tilting generated an elongated NE-SW graben. This is an evidence of the persistence of the Anti-Atlasic rifting process during the last part of the Lower-Cambrian succession.

  4. Rotational periodicities in Saturn's magnetosphere

    International Nuclear Information System (INIS)

    Complete text of publication follows. A central question for Saturn's magnetospheric dynamics is the origin of periodicities close to the presumed rotational period of Saturn (∼10.8 h). Periodic oscillations are observed in the magnetic field, particles, Saturn kilometric radiation (SKR) and more. We discuss the possibility that all periodicities are likely to have one common cause: periodic plasmoid release from the night side sector. The rapid magnetic field reconfigurations following the plasmoid release energize charged particles and create the large scale 'injections' that are clearly seen in energetic neutral atom (ENA) images observed by the INCA camera on board Cassini. We further show that the magnetic field oscillations and SKR can be explained by the currents driven by the injected and energized plasma pressure distribution drifting around Saturn. The evolution of these pressure-driven currents is consistent with the evolution of the SKR. To quantify the magnetic field oscillations we estimate the distribution and strength of the plasma pressure from Cassini/MIMI and CAPS observations of hot and cold plasma. An important constraint is the how the magnetospheric dynamics respond to variations in solar wind dynamic pressure (SWP). In order to quantify this constraint we use SWP measurements from when Cassini is in the solar wind as well as model propagated solar wind properties to investigate how periodicities vary as a function of SWP. We further speculate that the ultimate cause of the periodic plasmoid releases is an effect of the inherent stability of the cold plasma without the need of a longitudinal 'anomaly' tied to the planet's core or ionosphere.

  5. Drought periods during XXth century

    International Nuclear Information System (INIS)

    The drought problem always exists somewhere in the world so that the economy of some nation or nations is always being adversely influenced by this factor. Although drought is a natural component of the climate in arid and semi-arid areas, it can occur in areas, which normally receive adequate precipitation. Available hydro meteorological data indicate droughts have occurred through the last century in Bulgaria and they a part of the climatic cycle on the Balkan Peninsula. The precipitation distribution is one of the basic characteristics of the drought occurrence in a given region. However, the distributions of additional meteorological elements should be also taken into account in order to describe the degree of the climate dryness. For example, the distribution of air temperature is an especially important characteristic for drought classifications. The last years in Bulgaria were drier and warmer than the normals for the period of so-called 'current climate' (1961-1990). Precipitation was approximately 80-85 % less than the normals. Statistical methods were used in order to analyze the long-term variations of precipitation, air temperature. The long-term series of data were smoothed by averages and they were also approximated by means of a polynomial. These methods eliminate the random and short periodical fluctuations of the time series. The Spearman coefficient (r) and the Kendall coefficient (rl) were used in order to investigate the existence of eventual trends. Some quantitative criteria were also used for a comparison of the drought frequency and intensity between different regions and years. As a result of the conducted analysis, the investigated period can be divided to separate sub-periods with duration of 10-15 years. These years are characterized with different moisture conditions. Three periods can be determined during the 20 th century, which are characterized by longer and severe droughts, namely 1902-1913, 1942-1953 and 1982-1994. The drought

  6. Periodic sedimentation of three particles in periodic boundary conditions

    International Nuclear Information System (INIS)

    Solutions of the equations of Stokesian dynamics for point particles are found for periodic boundary conditions with three particles per unit cell of a simple cubic lattice. Two particles per cell move with equal velocity, but three particles per cell usually lead to irregular motion. Special situations are of interest, where initially the distance vector of one pair is parallel to one of the horizontal axes of the cubic lattice and the members of this pair are at equal distance to the third particle. By symmetry the configuration keeps this character during the motion, and numerically the motion is found to be periodic. In our numerical work we have studied mostly the case where the initial triangle is horizontal and equilateral. We have shown that the periodic motion is neutrally stable for sizes of the initial triangle less than the critical size. Such stable solutions of the equations of Stokesian dynamics are of relevance to the theory of sedimentation. In these solutions the particles move coherently in a complicated fashion with the same mean sedimentation velocity and a periodic internal motion of the three-particle cluster. If initially the particles are sufficiently widely separated, but the motion is still stable, the mean sedimentation velocity is less than that of a single particle. In this case the solution describes a situation of hindered settling. If the initial triangle is too large, with the side length larger than the critical size, the two base particles team up with partners in neighboring cells, and we get separate motion of a base pair and a single particle with different mean vertical velocities and with periodic motions superimposed. The columns of horizontal pairs pass the columns of apex particles. The corresponding solutions are unstable. (author)

  7. Ap stars with variable periods

    CERN Document Server

    Mikulášek, Zdeněk; Janík, Jan; Zejda, Miloslav; Henry, Gegory W; Paunzen, Ernst; Žižňovský, Jozef; Zverko, Juraj

    2013-01-01

    The majority of magnetic chemically peculiar (mCP) stars exhibit periodic light, magnetic, radio, and spectroscopic variations that can be adequately modelled as a rigidly-rotating main-sequence star with persistent surface structures. Nevertheless, there is a small sample of diverse mCP stars whose rotation periods vary on timescales of decades while the shapes of their phase curves remain unchanged. Alternating period increases and decreases have been suspected in the hot CP stars CU Vir and V901 Ori, while rotation in the moderately cool star BS Cir has been decelerating. These examples bring new insight into this theoretically unpredicted phenomenon. We discuss possible causes of such behaviour and propose that dynamic interactions between a thin, outer, magnetically-confined envelope braked by the stellar wind, and an inner faster-rotating stellar body are able to explain the observed rotational variability

  8. On composites with periodic structure

    Science.gov (United States)

    Nemat-Nasser, S.; Iwakuma, T.; Hejazi, M.

    1982-01-01

    The overall moduli of a composite with an isotropic elastic matrix containing periodically distributed (anisotropic) inclusions or voids, can be expressed in terms of several infinite series which only depend on the geometry of the inclusions or voids, and hence can be computed once and for all for given geometries. For solids with periodic structures these infinite series play exactly the same role as does Eshelby's tensor for a single inclusion or void in an unbounded elastic medium. For spherical and circular-cylindrical geometries, the required infinite series are calculated and the results are tabulated. These are then used to estimate the overall elastic moduli when either the overall strains or the overall stresses are prescribed, obtaining the same results. These results are compared with other estimates and with experimental data. It is found that the model of composites with periodic structure yields estimates in excellent agreement with the experimental observations.

  9. Periodicity, the Canon and Sport

    Directory of Open Access Journals (Sweden)

    Thomas F. Scanlon

    2015-10-01

    Full Text Available The topic according to this title is admittedly a broad one, embracing two very general concepts of time and of the cultural valuation of artistic products. Both phenomena are, in the present view, largely constructed by their contemporary cultures, and given authority to a great extent from the prestige of the past. The antiquity of tradition brings with it a certain cachet. Even though there may be peripheral debates in any given society which question the specifics of periodization or canonicity, individuals generally accept the consensus designation of a sequence of historical periods and they accept a list of highly valued artistic works as canonical or authoritative. We will first examine some of the processes of periodization and of canon-formation, after which we will discuss some specific examples of how these processes have worked in the sport of two ancient cultures, namely Greece and Mesoamerica.

  10. The age and tectonic setting of the Puncoviscana Formation in northwestern Argentina: An accretionary complex related to Early Cambrian closure of the Puncoviscana Ocean and accretion of the Arequipa-Antofalla block

    Science.gov (United States)

    Escayola, Mónica P.; van Staal, Cees R.; Davis, William J.

    2011-12-01

    TIMS and SHRIMP U-Pb zircon geochronology of selected parts of the Puncoviscana Formation suggest its deposition took place mainly during the Early Cambrian, coeval with 540-535 Ma calc-alkaline Pampean arc volcanism mainly preserved as tuff beds in the oldest identified parts of this unit. Syn- to post-tectonic plutons constrains the Tilcarian-Pampean orogeny to have occurred between ca. 530 Ma and deposition of the unconformably overlying Middle-Upper Cambrian Meson Group. Deposition of the Puncoviscana Formation continued after the onset of the Tilcarian-Pampean orogeny. We propose that the Puncoviscana Formation rocks older than 530 Ma were deposited in the arc-trench gap of the west-facing Pampean arc and/or the associated trench, whereas the rocks younger than 530 Ma were deposited in a syn-collision foreland basin. The Puncoviscana Formation rocks were progressively assembled into a west-younging accretionary complex, consistent with the style of deformation and low-grade metamorphism. The age of the syn-collision plutons (≤530 Ma) suggest the foredeep deposits record the transition from trench to foreland basin, due to arrival of the Arequipa-Antofalla block at the west-facing trench at ca. 530 Ma. Our geochronological and Pb-isotope investigations suggest that the Arequipa-Antofalla terrane was a coherent, ribbon-shaped crustal block that also included the western part of the Pampia terrane. A compilation of existing U-Pb zircon studies suggests that the Pampean arc extended along the length of the proto-Andean margin of West Gondwana, represented by the previously amalgamated Amazonia and Rio de La Plata cratons, and probably was initiated during the late Ediacaran after 600 Ma. Following earlier workers, we reaffirm that the Arequipa-Antofalla block was originally separating Laurentia and Amazonia in Rodinia. It probably rifted from Laurentia during the Ediacaran between 600 and 570 Ma, following an earlier departure of Amazonia (˜650 Ma?). The

  11. Profit optimization during crisis periods

    Directory of Open Access Journals (Sweden)

    Sorin BRICIU

    2013-06-01

    Full Text Available This article highlights the authors’ attempt to identify a simple and effective calculation method for profit optimization in crisis periods. This method was identified based on the analysis of a representative sample of professionals working in both state-owned and private bakery companies in Romania. The article also presents methodological case studies for three bakery products aimed to highlight the changes in results according to changes of variables in the current business environment. The results obtained are presented and analysed by the authors. The article ends with the authors’ conclusions on the benefits of the Direct-Costing Method and profit optimisation in crisis periods.

  12. Periodic Monopoles from Spectral Curves

    OpenAIRE

    Maldonado, R.

    2012-01-01

    We consider $\\text{SU}(2)$ Bogomolny equations on $\\mathbb{R}^2\\times\\hat{S}^1$ and use the spectral curve defined by the holonomy in the periodic direction to approximate the fields in the limit of large size to period ratio. Symmetries of the Nahm transform allow a study of the effective two dimensional dynamics, which is compared with known results on the full moduli space. The techniques are applied to systems of higher charge and higher rank gauge group, allowing a direct comparison to o...

  13. Availability of periodically tested systems

    International Nuclear Information System (INIS)

    There is at the present time a need in accurate models to asess the availability of periodically tested stand-by systems. This paper shows how to improve the well known 'saw-tooth curve' model in order to take into account various reliability parameters. A model is developed to assess the pointwise and the mean availabilities of periodically tested stand-by systems. Exact and approxination formulae are given. In addition, the model developed herein leads to optimize the test interval in order to minimize the mean unavailability. A safety diesel in a nuclear power plant is given as an example

  14. Periodic orbits in arithmetical chaos

    International Nuclear Information System (INIS)

    Length spectra of periodic orbits are investigated for some chaotic dynamical systems whose quantum energy spectra show unexpected statistical properties and for which the notion of arithmetical chaos has been introduced recently. These systems are defined as the unconstrained motions of particles on two dimensional surfaces of constant negative curvature whose fundamental groups are given by number theoretical statements (arithmetic Fuchsian groups). It is shown that the mean multiplicity of lengths l of periodic orbits grows asymptotically like c x el/2/l, l → ∞. Moreover, the constant c (depending on the arithmetic group) is determined. (orig.)

  15. Periodic growth of bacterial colonies

    Science.gov (United States)

    Yamazaki, Yoshihiro; Ikeda, Takemasa; Shimada, Hirotoshi; Hiramatsu, Fumiko; Kobayashi, Naoki; Wakita, Jun-ichi; Itoh, Hiroto; Kurosu, Sayuri; Nakatsuchi, Michio; Matsuyama, Tohey; Matsushita, Mitsugu

    2005-06-01

    The formation of concentric ring colonies by bacterial species Bacillus subtilis and Proteus mirabilis has been investigated experimentally, focusing our attention on the dependence of local cell density upon the bacterial motility. It has been confirmed that these concentric ring colonies reflect the periodic change of the bacterial motility between motile cell state and immotile cell state. We conclude that this periodic change is macroscopically determined neither by biological factors (i.e., biological clock) nor by chemical factors (chemotaxis as inhibitor). And our experimental results strongly suggest that the essential factor for the change of the bacterial motility during concentric ring formation is the local cell density.

  16. Long-period creep behaviour

    International Nuclear Information System (INIS)

    With a view to dimensioning and reliable determination of the service life of high-temperature components the following subjects are discussed: Creep data as criteria for materials selection; weld behaviour under creep stress; influence of the test atmosphere; influence of production process and initial structure; studies on creep behaviour extrapolation procedure; estimation of permissible operating periods. (orig./IHOE)

  17. 76 FR 52915 - Periodic Reporting

    Science.gov (United States)

    2011-08-24

    ... informal rulemaking on proposed changes in certain analytical methods used in periodic reporting. The... Service filed a petition pursuant to 39 CFR 3050.11 requesting that the Commission initiate an informal... distribution activities would make it easier to distinguish shape-related work from mixed-shape work at...

  18. Shanghai's Fashion During Republic Period

    Institute of Scientific and Technical Information of China (English)

    卞向阳

    2004-01-01

    The goal of this research is to describe and analyze Shanghai's fashion during Republic Period. Shanghai's fashion during Republic Period plays a special role in fashion history of China and Shanghai history. In the paper, three aspects including the fashion centre of China as well as Far East, men's fashion and women's fashion in Shanghai are discussed. The style, textiles, color, decoration and combination of clothing are described. Many costumes from the Clothing Museum at Donghua University and private collections as well as some historical writings, photographs and publications are elucidated and corroborated to give a more detailed description about Shanghai's fashion during that period.Here are some tentative conclusions. Firstly, Shanghai had always been the fashion center in China even Far East during Republic Period. Modeng was the main reflection of Shanghai fashion. Secondly, the traditional clothing exerted less influence and was less widely used, some kinds and combinations were out of fashion gradually. The fashion was described to be highly stylized, exquisite and concise. Thirdly, the influence of Western Style was further greater. The fashion in Shanghai was almost synchronous with fashion of Paris. Some western clothing was directly used in daily life and the elements of Western Style had an increasingly wide utilization. Fourthly, the New Chinese Style whose representatives were Qipao and Zhongshan Zhuang appeared and popularized in Shanghai. The fashion of Qipao changed frequently. Finally, the fashion in Shanghai called Shanghai Style could be summarized fashionable, exquisite, rapidly changeable, which is harmonious of both ancient and modern, Chinese and foreign.

  19. 76 FR 297 - Periodic Reporting

    Science.gov (United States)

    2011-01-04

    ... INFORMATION: Regulatory History, 75 FR 58449 (Sept. 24, 2010). On December 20, 2010, the Postal Service filed... ] estimated. The Postal Service proposes to rely primarily on data from the manual density table to estimate... 39 CFR Part 3050 Periodic Reporting AGENCY: Postal Regulatory Commission. ACTION: Notice of...

  20. Genius of the periodic table

    International Nuclear Information System (INIS)

    'Isn't it the work of a genius!' exclaimed Academician V.I. Spitsyn, USSR, a member of the Scientific Advisory Committee when talking to an Agency audience in January. His listeners shared his enthusiasm. Academician Spitsyn was referring to the first formulation a hundred years ago by Professor Dmitry I. Mendeleyev of the Periodic Law of Elements. (author)

  1. PERIODIC COMPLEMENTARY BINARY SEQUENCE PAIRS

    Institute of Scientific and Technical Information of China (English)

    XuChengqian; ZhaoXiaoqun

    2002-01-01

    A new set of binary sequences-Periodic Complementary Binary Sequence Pair (PCSP)is proposed .A new class of block design-Difference Family Pair (DFP)is also proposed .The relationship between PCSP and DFP,the properties and exising conditions of PCSP and the recursive constructions for PCSP are given.

  2. PERIODIC COMPLEMENTARY BINARY SEQUENCE PAIRS

    Institute of Scientific and Technical Information of China (English)

    Xu Chengqian; Zhao Xiaoqun

    2002-01-01

    A new set of binary sequences-Periodic Complementary Binary Sequence Pair (PCSP) is proposed. A new class of block design-Difference Family Pair (DFP) is also proposed.The relationship between PCSP and DFP, the properties and existing conditions of PCSP and the recursive constructions for PCSP are given.

  3. Model selection in periodic autoregressions

    NARCIS (Netherlands)

    Ph.H.B.F. Franses (Philip Hans); R. Paap (Richard)

    1994-01-01

    textabstractThis paper focuses on the issue of period autoagressive time series models (PAR) selection in practice. One aspect of model selection is the choice for the appropriate PAR order. This can be of interest for the valuation of economic models. Further, the appropriate PAR order is important

  4. Pairs of dual periodic frames

    DEFF Research Database (Denmark)

    Christensen, Ole; Goh, Say Song

    2012-01-01

    needed. The purpose of the present paper is to provide constructions of dual pairs of frames in the setting of the Hilbert space of periodic functions L2(0,2π). The frames constructed are given explicitly as trigonometric polynomials, which allows for an efficient calculation of the coefficients in the...

  5. Portfolio Selection and Insurance Period

    OpenAIRE

    Aoba, Nobuko

    2001-01-01

    This paper examines market risks, portfolio selection and period of insurance in a life insurance market. Date from many countries indicate that over ninety percent of bankruptcies of life insurers are due of failures in financial risk management. Financial risk management is therefore central. ...

  6. Transition Period and Immunosuppression: Critical Period of Dairy Cattle Reproduction

    Directory of Open Access Journals (Sweden)

    K. Simenew

    2013-04-01

    Full Text Available This seminar study is prepared on the objectives of: revising important aspects of transition period of dairy cattle and highlighting some potential areas of research and challenges for the future. It has sufficiently been discussed that improved understanding of this frontier of the biology, immunology, nutrition and management of cows during the transition period will provide the largest gains in productivity and profitability of dairy farms. In the manuscript under each specific topic, transition cow program and reproductive performance, immunosuppressant effect of transition period, early predictors of disorders and major abnormalities are discussed in an informative way. Future potential areas of research and possible challenges are also indicated briefly. Finally, it is concluded that despite decades of research in the area of transition cow health and management the high incidence of health disorders around calving continues to negatively affect milk production and reproductive performance; and as recommendation, implementing a transition nutrition program with the help of nutritionists can help dairy herd avoid most of the costly problems and molecular level research studies should get due attention to further understand the situation and devise proper intervention techniques.

  7. Geopulsation, Volcanism and Astronomical Periods

    Institute of Scientific and Technical Information of China (English)

    Yang Xuexiang; Chen Dianyou; Yang Xiaoying; Yang Shuchen

    2000-01-01

    Volcanism is mainly controlled by the intermittent release of energy in the earth. As far as the differential rotation of the earth's inner core is concerned, the Galactic Year may change the gravitational constant G, the solar radiative quantity and the moving speed of the solar system and affect the exchange of angular momentum between core and mantle as well as the energy exchange between crust and mantle. As a result, this leads to eruptions of superplumes and magma, and controls the energy flow from core - mantle boundary (CMB) to crust. When the earth' s speed decreases, it will release a huge amount of energy. They are the reason of the correspondence of the volcanic cycles one by one with the astronomical periods one by one. According to the astronomical periods, volcanic eruptions may possibly be predicted in the future.

  8. Periodizing Globalization: Histories of Globalization

    OpenAIRE

    Nederveen Pieterse Jan

    2012-01-01

    The article outlines the analytics and criteria that inform periodizing globalization. It criticizes presentist and Eurocentric views on globalization, the contemporary view, the modernity view (1800 plus) or the capitalism view (1500 plus). It discusses approaches to world history and how globalization fits in. Understandings of globalization, such as multicentric and centrist perspectives, and units of analysis affect how timelines of globalization are established. Taking into account globa...

  9. The Periodic Table in Flatland

    OpenAIRE

    Negadi, T.; Kibler, M.

    1995-01-01

    The $D$-dimensional Coulomb system serves as a starting point for generating generalized atomic shells. These shells are ordered according to a generalized Madelung rule in $D$ dimensions. This rule together with an {\\it Aufbau Prinzip} is applied to produce a $D$-dimensional periodic table. A model is developed to rationalize the ordering of the shells predicted by the generalized Madelung rule. This model is based on the introduction of an Hamiltonian, invariant under the $q$-deformed algeb...

  10. Potassium-Related Periodic Paralysis

    OpenAIRE

    A Lesani; MH Moradi Nejad

    1995-01-01

    Potassium-related periodic paralysis may be associated with hypo, normo, or hyperkalemic disturbance of the Potassium level. Clinical features, laboratory findings and ECG as well as EMG changes are different in the 3 types of the disease. The treatment demands correction of the Potassium level of the blood serum. The disease is normally transferred as an autosomal dominant trait. In this article we present a 9-year old girl who suffered from hypokalemic paralysis since she was 4 years old.

  11. Social Research On Renaissance Period

    Institute of Scientific and Technical Information of China (English)

    张蕾蕾

    2009-01-01

    The historical period known as the Renaissance spans the fifteenth and siXteenth centuries. ‘Renaissance'means‘re-birth'in French and during this time Europe left behind the mindset of the Midge Ages and created the beginnings of the modern world.This thesis mainly rotates on the background,the main features and significance of Renaissance,especially focuses on the discovery of man and the humanism and the detailed unfolding of the society on several aspect in Renaissance.

  12. Nested long period grating interferometers

    OpenAIRE

    Murphy, Richard P.; James, Stephen W.; Tatam, Ralph P.

    2007-01-01

    The concept of nested fibre optic long period grating (LPG) based interferometers is introduced. A number of in-series, identical LPGs may be used to form a set of nested, multiplexed Mach-Zehnder interferometers that may demodulated and demultiplexed by virtue of a Fourier analysis of the optical spectrum. The concept is demonstrated by the use of three LPGs to form a nested set of interferometers.

  13. A Periodic Lotka-Volterra System

    OpenAIRE

    Tsvetkov, D.

    1996-01-01

    In this paper periodic time-dependent Lotka-Volterra systems are considered. It is shown that such a system has positive periodic solutions. It is done without constructive conditions over the period and the parameters.

  14. Quasi Periodic Oscillations in Blazars

    Indian Academy of Sciences (India)

    Alok C. Gupta

    2014-09-01

    Here we report our recent discoveries of Quasi-Periodic Oscillations (QPOs) in blazars time series data in X-ray and optical electromagnetic bands. Any such detection can give important clues of the location and nature of the processes of emission mechanism. In the case of radio-quiet AGN, the detected QPOs are very likely to be associated with the accretion disk. But in the case of blazars, it may be associated with jets in the high and outburst states, and in the low-state, it is probably associated with the accretion disk. In this brief review, I summarize the recent QPO detections in blazars. There is one strong evidence of QPO detection in XMM–Newton time series data of narrow line Seyfert 1 galaxy RE J1034+396 about which we will also discuss briefly.

  15. The periodically oscillating plasma sphere

    International Nuclear Information System (INIS)

    A new method of operating an inertial electrostatic confinement (IEC) device is proposed, and its performance is evaluated. The scheme involved an oscillating thermal cloud of ions immersed in a bath of electrons that form a harmonic oscillator potential. The scheme is called the periodically oscillating plasma sphere, and it appears to solve many of the problems that may limit other IEC systems to low gain. A set of self-similar solutions to the ion fluid equations is presented, and plasma performance is evaluated. Results indicate that performance enhancement of gridded IEC systems such as the Los Alamos intense neutron source device is possible as well as high-performance operation for low-loss systems such as the Penning trap experiment. Finally, a conceptual idea for a massively modular Penning trap reactor is also presented

  16. The periodic table in Flatland

    International Nuclear Information System (INIS)

    The D-dimensional Coulomb system serves as a starting point for generating generalized atomic shells. These shells are ordered according to a generalized Madelung rule in D dimensions. This rule together with an Aujbau Prinzip is applied to produce a D-dimensional periodic table. A model is developed to rationalize the ordering of the shells predicted by the generalized Madelung rule. This model is based on the introduction of a Hamiltonian, invariant under the q-deformed algebra Uq(so(D)), that breaks down the SO(D + 1) dynamical symmetry of the hydrogen atom in D dimensions. The D = 2 case (Flatland) is investigated in some detail. It is shown that the neutral atoms and the (moderately) positive ions correspond to the values q = 0.8 and q = 1, respectively, of the deformation parameter q. 55 refs

  17. The periodic table in Flatland

    International Nuclear Information System (INIS)

    The D-dimensional Coulomb system serves as a starting point for generating generalized atomic shells. These shells are ordered according to a generalized Madelung rule in D dimensions. This rule together with an Aufbau Prinzip is applied to produce a D-dimensional periodic table. A model is developed to rationalize the ordering of the shells predicted by the generalized Madelung rule. This model is based on the introduction of an Hamiltonian, invariant under the q-deformed algebra Uq (so(D)), that breaks down the SO (D + 1) dynamical symmetry of the hydrogen atom in D dimensions. The D = 2 case (Flatland) is investigated with some details. It is shown that the neutral atoms and the (moderately) positive ions correspond to the values q = 0.8 and q = 1, respectively, of the deformation parameter q. (authors). 55 refs

  18. Periods of ZZ Ceti variables

    International Nuclear Information System (INIS)

    White dwarf pulsators (ZZ Ceti variables) osub solar acccur in the extension of the radial pulsation envelope ionization instability strip to the observed luminosities of 3 x 10-3L sub solar according to van Horn. Investigations were underway to see if the driving mechanisms of hydrogen and helium ionization can cause radial pulsations as they do for the Cepheids, the RR Lyrae variables, and the delta Scuti variables. Masses used in this study are 0.60 and 0.75 M sub solar for T/sub e/ between 10,000 K and 14,000 K, the observed range in T/sub e/. Helium rich surface compositions like Y = 0.78,, Z = 0.02 as well as Y = 0.28, Z = 0.02 were used in spite of observations showing only hydrogen lines in the spectrum. The deep layers are pure carbon, and several transition compositions are included. The models show radial pulsation instabilities for many overtone modes at periods between about 0.3 and 3 seconds. The driving mechanism is mostly helium ionization at 40,000 and 150,000 K. The blue edge at about 14,000 K is probably due to the driving region becoming too shallow, and the red edge at 10,000 K is due to so much convection in the pulsation deriving region that no radiative luminosity is available for modulation by the γ and kappa effects. It is speculated that the very long observed periods (100 to 1000 sec) of ZZ Ceti variables are not due to nonradial pulsations, but are possibly aliases due to data undersampling. 4 references

  19. ORGANIC CARBON AND NITROGEN RECORDS IN THE SINIAN -EARLY CAMBRIAN SUCCESSION OF SOUTHERN ANHUI, CHINA%皖南震旦纪-早寒武世沉积岩有机碳、氮元素记录

    Institute of Scientific and Technical Information of China (English)

    王金权

    2003-01-01

    安徽南部晚震旦世蓝田组、皮园村组和早寒武世荷塘组主要由黑色页岩和硅质岩组成.分析结果表明,蓝田组和荷塘组的总有机碳(可达10%-18%)和总有机氮(可达2.0‰-2.6‰)的高含量,应归于富有机质埋藏所致,反映新元古代末次冰期后和早寒武世海洋浮游生物处于大发展时期.由于海洋植物的光合作用产生大量O2,并消耗大量CO2,导致大气中O2/CO2比率明显增加,为震旦纪晚期多细胞真核生物的辐射、后生动物的出现和‘寒武纪生物大爆发'提供了条件.蓝田组黑色页岩中部的有机碳与有机氮含量大体呈现正比例增加,并且分别达到17.74%和2.66‰的最高值,与该剖面发现的蓝田植物群相对应;荷塘组黑色页岩的有机碳和有机氮垂直分布曲线之间呈现相互背离,即有机氮含量与时增加,而有机碳含量与时减少,与扬子地台相应的地层层位产生的澄江动物群相对应,表明TON和TOC分布特征有可能与早期生物演化历程相关.%Organic carbon and organic nitrogen analyses in the Sinian-Early Cambrian succession of southern Anhui Province, China, show that the high contents of both total organic carbon (TOC: up to 10 %-18%) and total organic nitrogen (TON: up to 2.0‰-2.6‰) may reflect a great increase of organic matter burial and consequent rise in O2 concentration in the atmosphere and the upper ocean. It, in turn, may be attributed to a high primary productivity in the ocean, which resulted from a rise of major global temperature and enhanced tectonism after the Late Neoproterozoic glaciation. Warm climate and the rise in O2 concentration of the atmosphere may have triggered the radiation of multicellular eukaryotes in the Doushantuo time, development of the metazoan in the late Sinian and 'Cambrian Explosion'. In addition, the TON and TOC vertical distribution characteristics may be asso ciated with the early life evolution.

  20. Electric property evidences of carbonification of organic matters in marine shales and its geologic significance: A case study of the Lower Cambrian Qiongzhusi shale in the southern Sichuan Basin

    Directory of Open Access Journals (Sweden)

    Yuman Wang

    2014-12-01

    Full Text Available Searching for some reliable evidences that can verify the carbonification of organic matters in marine shales is a major scientific issue in selecting shale gas fairways in old strata. To this end, based on core, logging and testing data, the electric property of two organic-rich shale layers in the Lower Cambrian Qiongzhusi Fm. and the Lower Silurian Longmaxi Fm. in the southern Sichuan Basin was compared to examine the carbonification signs of organic matters in the Qiongzhusi shale and its influence on gas occurrence in the shales. The following conclusions were reached: (1 the electric property experiment shows that the Qiongzhusi shale in the study area has had carbonification of organic matters. The low resistivity of dry samples from this highly mature organic-rich shale and ultra-low resistivity on downhole logs can be used to directly judge the degree of organic matter carbonification and the quality of source rocks; (2 in the Changning area, the Qiongzhusi shale shows low resistivity of dry samples and low to ultra-low resistivity on logs, indicating that organic matters are seriously carbonized, while in the Weiyuan area, the Qiongzhusi shale shows a basically normal resistivity on log curves, indicating its degree of graphitization between the Longmaxi Fm. and Qiongzhusi Fm. in the Changning area; (3 shale with medium-to-high resistivity is remarkably better than that with ultra-low resistivity in terms of gas generation potential, matrix porosity and gas adsorption capacity; (4 industrial gas flow has been tested in the organic shales with medium-to-high resistivity in the Jianwei–Weiyuan–Tongnan area in the north, where the Qiongzhusi shale is a favorable shale gas exploration target.

  1. 46 CFR 9.8 - Broken periods.

    Science.gov (United States)

    2010-10-01

    ... 46 Shipping 1 2010-10-01 2010-10-01 false Broken periods. 9.8 Section 9.8 Shipping COAST GUARD... § 9.8 Broken periods. In computing extra compensation where the services rendered are in broken periods and less than 2 hours intervene between such broken periods the time served should be...

  2. Enrolment in Conocoryphe heberti Munier-Chalmas and Bergeron, 1889 (Middle Cambrian, Iberian Chain, NE Spain) and coaptive structures within family Conocoryphidae; Enrollamiento en Conocoryphe heberti Munier-Chalmas and Bergeron, 1889 (Cambrico Medio, Cadena Iberica, NE de Espana) y estructuras coaptativas en la familia

    Energy Technology Data Exchange (ETDEWEB)

    Esteve, J.

    2009-07-01

    Enrolled Conocoryphe heberti is described with detail and compared with the conocoryphid Occatharia sdzuyi, both very abundant in the middle Cambrian from the Mediterranean subprovince. Both show different morphological novelties very useful to enrol and interlock the trilobite body. The morphological study allows us to recognize coaptative devices that show the possibility of a tight encapsulation enrolment. The coaptative devices are analyzed in cephalons, pigidia and almost completed outstretched trilobites of Bailiella levyi, Bailiaspis souchoni and Parabailiella languedocensis belonging to the same family. The coaptative structures analyzed show enough variability in cephalons and pigidia that allow suggesting several enrolment types within conocoryphids. (Author) 54 refs.

  3. Evidence of syn tectonic tephrites with nepheline in the Sidi Said Maachou Cambrian basin (coastal Meseta, Morocco); geo dynamic implications; Mise en evidence de tephrites a nepheline syntectoniques dans le bassin cambrien de Sidi Said Maachou (Meseta cotiere, Maroc); signification geodynamique

    Energy Technology Data Exchange (ETDEWEB)

    Remmal, T.; Mohsine, A.; El Hatimi, N.

    2009-07-01

    Based on a combined structural, petrographic, and geochemical analysis, a new interpretation of the basic magmatism of Sidi Said Maachou (coastal Meseta) in two stages of emplacement is proposed. The first stage is characterized by transitional pyroclastic flows that have accompanied the opening of the West-Mesetian basin, during the Cambrian; the second stage is made of dykes of basalts, dolerites, and tephrites bearing nepheline. The emplacement of this undersaturated alkaline magma is associated to a sinistral sub meridian shear zone which has been activated at the end of the Caledonian orogenesis, by a mantellic advection. (Author) 32 refs.

  4. Periodic and Almost Periodic Solutions of Functional Difference Equations with Finite Delay

    OpenAIRE

    Yihong Song

    2007-01-01

    For periodic and almost periodic functional difference equations with finite delay, the existence of periodic and almost periodic solutions is obtained by using stability properties of a bounded solution.

  5. Periodical plane puzzles with numbers

    CERN Document Server

    Rezende, Jorge

    2011-01-01

    Consider a periodical (in two independent directions) tiling of the plane with polygons (faces). In this article we shall only give examples using squares, regular hexagons, equilateral triangles and parallelograms ("unions" of two equilateral triangles). We shall call some "multiple" of the fundamental region "the board". We naturally identify pairs of corresponding edges of the the board. Figures 9 and 19-29, in this article, show different boards. The "border" of the board is represented by a yellow thick line, unless part of it or all of it is the edge of a face. The board is tiled by a finite number of polygons. Construct polygonal plates in the same number, shape and size as the polygons of the board. Adjacent to each side of each plate draw a number, or two numbers, like it is shown in Figures 1 and 18-29. Figure 1 shows the obvious possibility of having plates with simple drawings, coloured drawings, etc. Now the game is to put the plates over the board polygons in such a way that the numbers near eac...

  6. Tectono-depositional patterns and palaeogeography in the Middle Yangtze River region during the Early Cambrian%中扬子地区早寒武世构造—沉积样式与古地理格局

    Institute of Scientific and Technical Information of China (English)

    陈代钊; 汪建国; 严德天; 韦恒叶

    2012-01-01

    地除外).从第四构造幕热液活动后,盆地基底热衰减迅速,深部硅输入通量大幅减少,并在扬子地块发生大范围均匀挠曲沉降和快速海平面上升,造成碳酸盐台地淹没,形成了牛蹄塘组(或同期)富有机质黑色岩系.从此后,中扬子地区进入了一个新的构造—沉积演化时期.%The Precambrian-Cambrian transition is a key interval in Earth's history, as it records the advent,extinction and accelerated diversification of metazoans ('Cambrian explosion') and co-evolution of tectonic re-organization of global plates, a prolonged global oceanic anoxia and swift shifts in ocean geochemistry. To understand better the fundamental environmental changes on Earth's surface system, detailed analyses on depositional facies and sequences of the Lower Cambrian (Stages 1 and 2) were carried out across the western Hubei and northwestern Hunan in the Middle Yangtze River region, South China, and 16 types of lithofacies in three main lithological categories of carbonate, chert and fine-grained clastic mudrock were identified, which can be grouped into four depositional sequences (SI ~ S4 ) in terms of tempo-spatial facies variations. Just at the onset of Cambrian, the carbonate platform in northwestern Hunan was subject to subaerial exposure, its southern margin, however, underwent rapid subsidence , forming the perched terrace; the northern flank also suffered large-scale subsidence, plunging down to the basin in the northwestern Hunan ( Yangjiaping) , where the basement relief begun to rise once again to the north and evolved into carbonate platform in Three Gorges area in western Hubei. The vast lateral topographic differences shaped by the two uplift-depressions point toblock tilting in the context of extensional tectonism, which could have episodically occurred and thereby controlled the deposition, facies patterns and distributions in the Early Cambrian as recorded by the depositional sequences. A depositional

  7. Characteristics and distribution of grain bank of the Lower Cambrian Longwangmiao Formation in Moxi-Gaoshiti area,Sichuan Basin%四川盆地磨溪-高石梯地区下寒武统龙王庙组颗粒滩特征及分布规律

    Institute of Scientific and Technical Information of China (English)

    金民东; 谭秀成; 李凌; 曾伟; 连承波; 罗冰; 山述娇; 周涛

    2015-01-01

    In this paper,the authors focused on grain bank of the Lower Cambrian Longwangmiao Formation in Moxi-Gaoshiti area,Sichuan Basin.Combined with abundant materials,such as drilling and logging data,cores and thin sections,the characteristics,distribution and its major controlling factors of grain bank in the Longwangmiao Formation were synthetically analyzed.There are two main rock types of grain bank in research area,one is grain dolostone with obvious granular structure,which is dominated by dolarenite and oolite dolostone,and the other is crystal grain dolostone with residual granular structure. Grain bank mainly has upward shallowing and granularity coarsening sequences,and could constitute sedi-mentary sequences of interact bank sea -grain bank and restricted lagoon -grain bank with lower fine-grained sediment in the vertical direction.In the lateral direction,grain bank has good continuity and is characterized by continuous distribution in a large area.On the plane,the development degree of grain bank increases gradually from northwest to southeast.Further analysis shows that the development charac-teristics and superimposed pattern of grain bank in the Longwangmiao Formation were determined by the relative sea-level change and sedimentary energy variation,the development horizon and degree of grain bank were formed by different grades of sea level fluctuation,and the palaeogeomorphology finally deter-mined the difference of planar distribution of grain bank during the depositional period of Longwangmiao Formation.%以四川盆地磨溪—高石梯地区下寒武统龙王庙组颗粒滩为研究对象,结合钻井、测录井、岩心和薄片等资料,对龙王庙组颗粒滩特征、分布规律和主控因素进行了综合分析。研究区内颗粒滩岩石类型主要有两类:一类为具有明显颗粒结构的颗粒云岩,以砂屑云岩或鲕粒云岩为主;另一类为具有残余颗粒结构的晶粒云岩。颗粒滩主要发育向上

  8. 西昆仑北缘库斯拉甫一带寒武纪中酸性岩浆活动及地质意义%Cambrian intermediate-acid magmatic activity and its geological significance in the Kusilafu region at the north margin of the Western Kunlun

    Institute of Scientific and Technical Information of China (English)

    黄建国; 杨瑞东; 杨剑; 崔春龙; 侯兰杰

    2013-01-01

    西昆仑北缘库斯拉甫一带寒武纪中酸性侵入岩广泛分布,据野外接触关系可分为前后两个序次。早序次岩石为浅灰-麻灰色似斑状石英(二长)闪长岩,具贫硅、中碱、高钙和过铝质等特征,岩石稀土总量较低,轻稀土元素富集((La/Yb)N平均15.56),中等 Eu 负异常(δEu 为0.56~0.88,平均0.73)。晚序次岩石为灰白色似斑状粗粒(二长)花岗岩,具富硅、低钙、富钾和过铝质等特征,稀土元素总量较高,轻稀土元素富集((La/Yb)N平均13.39),Eu负异常明显(δEu为0.25~0.57,平均0.40)。早序次岩体侵位规模大,出露广泛,为I型花岗岩;晚序次岩体规模小,以岩株、岩脉状穿插其中,属于S型花岗岩范畴。两序次岩石产出的大地构造环境均为岛弧,可能由昆仑洋的俯冲消减引起,不同之处在于早序次岩石产于活动大陆(西昆仑地块)边缘位置,而晚序次形成在俯冲消减带上,两序次岩石为消减洋壳上部不同源区地壳部分熔融的产物。%Cambrian intermediate-acid intrusive rocks (U-Pb age (511±29) Ma) distribute widely around the Kusilafu region at the north margin of the Western Kunlun. The intrusive rocks can be subdivided into early and later periods according to contact relationships. The early intrusion is greyish porphyritic quartz (monzonite) diorite, characterized by poor-SiO2 moderate-alkaline, high-Ca and strongly peraluminous, and is relatively low inΣREE with LREE enrichment ((La/Yb)N, 11.65 on average) and moderate negative Eu anomaly (δEu varied from 0.40 to 0.71, 0.73 on average). While the later is offwhite porphyritic coarse-grained (monzonite) granite characterized by rich-Si, low-Ca, rich-K and peraluminous, and is relatively enriched inΣREE with an enrichment of LREE ((La/Yb)N average at 10.19) and moderate negative Eu anomaly (δEu varied from 0.40 to 0.71, averaged at 0.73). The early intrusion is larger with widespread

  9. Detection of superimposed periodic signals using wavelets

    OpenAIRE

    Otazu, X.; Ribo, M.; Peracaula, M.; Paredes, J. M.; Nunez, J.

    2002-01-01

    In this paper we present a wavelet based algorithm that is able to detect superimposed periodic signals in data with low signal-noise ratio. In this context, the results given by classical period determination methods highly depend on the intrinsic characteristics of each periodic signal, like amplitude or profile. It is then difficult to detect the different periods present in the data set. The results given by the wavelet based method for period determination we present here are independent...

  10. Thyrotoxic Hypokalemic Periodic Paralysis: A Case Report

    OpenAIRE

    Tuna, Mazhar Muslum; Aycicek Dogan, Bercem; Nasiroglu Imga, Narin; Karakilic, Ersen; Karadeniz, Mine; Tutuncu, Yasemin; Isik, Serhat; Berker, Dilek; Guler, Serdar

    2014-01-01

    AbstractHypokalemic periodic paralysis is a rare disorder characterized by reversible attacks of muscle weakness accompanied by episodic hypokalemia. The most common causes of hypokalemic periodic paralysis (HPP) are familial periodic paralysis, thyrotoxic periodic paralysis (TPP) and sporadic periodic paralysis, respectively. There are generally some precipitating factors such as stress, vigorous exercise and high carbohydrate food consumption which all ease the occurrence of attacks. The du...

  11. Aperiodic and semi-periodic perfect maps

    OpenAIRE

    Mitchell, Chris

    1995-01-01

    Paterson [1] has recently shown that the trivial necessary conditions are sufficient for the existence of a (binary) perfect map. These periodic structures can be transformed very simply into corresponding aperiodic and semi-periodic perfect maps. However, aperiodic and semi-periodic perfect maps can exist for parameter sets for which the corresponding periodic perfect maps cannot. In this paper it is shown, by construction, that (binary) aperiodic and semi-periodic perfect maps exist for all...

  12. The late stages of the Pampean Orogeny, Córdoba (Argentina): Evidence of postcollisional Early Cambrian slab break-off magmatism

    Science.gov (United States)

    Ramos, Victor A.; Escayola, Mónica; Leal, Pablo; Pimentel, Marcio M.; Santos, João O. S.

    2015-12-01

    Widespread rhyolitic and mafic volcanism in the northern and southern sectors respectively of Eastern Sierras Pampeanas, central western Argentina, are associated with an important phase of extension and uplift linked to slab break-off on latest stages of the Pampean Orogeny. The main orogenic deformation took place between 540 and 535 Ma based on new ages available for this region. New U-Pb ages in zircons from the Oncán Rhyolite and new and old recalculated zircons from Los Burros Rhyodacite in the northern sector of Sierras de Córdoba, together with new U-Pb ages of the southern sector, indicate that volcanic and subvolcanic rocks of both sectors are partially coeval and unconformably overlying and/or intruding the basement rocks during a period of exhumation and subsequent cooling at 530-520 Ma. These data are in agreement with previous estimates for the final uplift of the Sierras de Córdoba based on the K-Ar cooling ages. The southern sector of Eastern Sierras Pampeanas may represent deeper structural levels within the crust and is characterized by the occurrence of small mafic bodies with OIB-like signature. New and reinterpreted U-Pb SHRIMP and TIMS ages on zircons and monazites in the metamorphic associated rocks date this episode, which is related to a rapid slab break-off event at 519-515 Ma. This episode is also associated with the emplacement of peraluminous granitoids, and with the extension, cooling and final uplift of the rocks affected by the Pampean Orogeny all along the Eastern Sierras Pampeanas.

  13. Periodic Poisson Solver for Particle Tracking

    CERN Document Server

    Dohlus, M

    2015-01-01

    A method is described to solve the Poisson problem for a three dimensional source distribution that is periodic into one direction. Perpendicular to the direction of periodicity a free space (or open) boundary is realized. In beam physics, this approach allows to calculate the space charge field of a continualized charged particle distribution with periodic pattern. The method is based on a particle mesh approach with equidistant grid and fast convolution with a Green's function. The periodic approach uses only one period of the source distribution, but a periodic extension of the Green's function. The approach is numerically efficient and allows the investigation of periodic- and pseudo-periodic structures with period lengths that are small compared to the source dimensions, for instance of laser modulated beams or of the evolution of micro bunch structures. Applications for laser modulated beams are given.

  14. Crystallography beyond periodic Crystal perfection

    International Nuclear Information System (INIS)

    Full text: The discovery of the quasi-crystals [D. Schechtman et. Al., Phys.] Rev. Lett. [53, 1951-1953 (1984)] made very narrow definition of the crystalline state based on the periodicity of a local arrangement of atoms. Since the definition of this State has been a matter of much controversy [G.R. Desiraju, Nature 423, 485 (2003); S. van Smaalen, IUCR Aperiodic Commission Reports. August 7, 2002; International Union of Crystallography. Report of the Executive Committee for 1991; ACTA Cryst. A48, 922-946 (1992)]. We will make a presentation of the current time of the crystallography in this regard from the conceptual point of view. We show the use of the formalism of algorithmic complexity or Kolmogorov [M. Li and P. Vitanyi, An Introduction to Kolmogorov Complexity and Its Applications (Springer Verlag, Heidelberg, 1993), W.H. Zurek, Phys.] Rev. 40, 4731 (1989); Nature 341, 119-124 (1989)] provides a different perspective on the nature of the Crystallographic order. Infinite crystals can be considered solid with zero algorithmic complexities by atom. Show statistical analysis of inorganic compounds [J.L.C. Daams et al., Atlas of Crystal Structure Types for Intermetallic Phases (ASM International, Ohio, 1991), Fachinformationszentrum/NIST Inorganic Crystal Structure Database, Karlsruhe (2003) icsd.fkf.mpg.de] demonstrating that the minimization of complexity is a trend in the crystalline arrangement. We will then compare the degree of disorder of some typical solids according to their algorithmic complexity. Finally, space diffraction will be studied from this same perspective and will be discussed that zero algorithmic complexities by point in space of diffraction does not necessarily imply the same thing for the Atomic arrangement. The discrete portion of the diffraction pattern is a fingerprint of the underlying order but not the actual existence of long-range order. Experimental results will be showcased [E. Estévez-Rams et al., Physical Review B, 63 (2001

  15. The Galois coaction on $\\phi^4$ periods

    CERN Document Server

    Panzer, Erik

    2016-01-01

    We report on calculations of Feynman periods of primitive log-divergent $\\phi^4$ graphs up to eleven loops. The structure of $\\phi^4$ periods is described by a series of conjectures. In particular, we discuss the possibility that $\\phi^4$ periods are a comodule under the Galois coaction. Finally, we compare the results with the periods of primitive log-divergent non-$\\phi^4$ graphs up to eight loops and find remarkable differences to $\\phi^4$ periods. Explicit results for all periods we could compute are provided in ancillary files.

  16. US Forest Service Periodical Cicada Broods

    Data.gov (United States)

    US Forest Service, Department of Agriculture — A map service on the www depicting periodical cicada distribution and expected year of emergence by cicada brood and county. The periodical cicada emerges in...

  17. Genetics Home Reference: hypokalemic periodic paralysis

    Science.gov (United States)

    ... paralysis Patient Support and Advocacy Resources (2 links) Muscular Dystrophy Association Resource list from the University of Kansas Medical Center GeneReviews (1 link) Hypokalemic Periodic Paralysis Genetic Testing Registry (3 links) Hypokalemic periodic paralysis Hypokalemic ...

  18. Periodic solutions of nonlinear vibrating beams

    Directory of Open Access Journals (Sweden)

    J. Berkovits

    2003-01-01

    Full Text Available The aim of this paper is to prove new existence and multiplicity results for periodic semilinear beam equation with a nonlinear time-independent perturbation in case the period is not prescribed. Since the spectrum of the linear part varies with the period, the solvability of the equation depends crucially on the period which can be chosen as a free parameter. Since the period of the external forcing is generally unknown a priori, we consider the following natural problem. For a given time-independent nonlinearity, find periods T for which the equation is solvable for any T-periodic forcing. We will also deal with the existence of multiple solutions when the nonlinearity interacts with the spectrum of the linear part. We show that under certain conditions multiple solutions do exist for any small forcing term with suitable period T. The results are obtained via generalized Leray-Schauder degree and reductions to invariant subspaces.

  19. Financial Innovation in Multi-Period Economies

    OpenAIRE

    Enrique Kawamura

    2003-01-01

    I present an attempt to construct multi-period, finite horizon extensions to the well -known two- period financial innovations literature. I first extend the definition of competitive equilibrium with innovations. It is shown that, with a dominating house

  20. Periodic Poisson model for beam dynamics simulation

    Science.gov (United States)

    Dohlus, M.; Henning, Ch.

    2016-03-01

    A method is described to solve the Poisson problem for a three dimensional source distribution that is periodic into one direction. Perpendicular to the direction of periodicity a free space (or open) boundary condition is realized. In beam physics, this approach allows us to calculate the space charge field of a continualized charged particle distribution with periodic pattern. The method is based on a particle-mesh approach with equidistant grid and fast convolution with a Green's function. The periodic approach uses only one period of the source distribution, but a periodic extension of the Green's function. The approach is numerically efficient and allows the investigation of periodic- and pseudoperiodic structures with period lengths that are small compared to the source dimensions, for instance of laser modulated beams or of the evolution of micro bunch structures. Applications for laser modulated beams are given.

  1. The Middle Islamic and Crusader Periods

    DEFF Research Database (Denmark)

    Walmsley, Alan George

    2008-01-01

    The book chapter presents a critical review of the archaeology of Jordan in the middle Islamic periods (8th - 17th centuries CE), presenting a new evaluation of settlement and culture in the period, and assesses the errors of past appraoches.......The book chapter presents a critical review of the archaeology of Jordan in the middle Islamic periods (8th - 17th centuries CE), presenting a new evaluation of settlement and culture in the period, and assesses the errors of past appraoches....

  2. Umbrella structure and channel-wall stoping in the Cambrian St. Roch Formation, Quebec Appalachians: significance for particle support mechanisms and turbulence development in hyper-concentrated sediment gravity flows

    Science.gov (United States)

    Hesse, Reinhard; Fong, Christopher

    2014-03-01

    Umbrella structure is a newly recognized sedimentary structure associated with large platy clasts in resedimented boulder-bearing pebble conglomerate with a sandy matrix. It consists of a sand rim that lacks pebbles on parts or the entire underside of platy boulders, whereas on the upper side, pebbles are in direct contact with the boulders. The depositing processes were high- to hyper-concentrated sediment gravity flows in a submarine channel or canyon on the Cambrian continental slope of North America bordering the Iapetus Ocean. The structure occurs predominantly where clasts dip moderately in the down-current direction. Based on the association of the structure with slightly forward dipping slabs, it is proposed that these down-current dipping slabs may have been in the process of counter-clockwise rotation that was aborted and may have generated a pressure shadow on the underside enabling the inrush of fluid and the infiltration of sand into the anomalous low-pressure zone. The structure has implications for particle support mechanisms in high- to hyper-concentrated sedimentary gravity flows, in that it redirects attention to the much debated mechanism of dispersive pressure and alternatives. It provides an observable sediment structure that supports dispersive pressure which so far depended on experimental evidence and theoretical arguments alone. Vrolijk and Southard's (1997) concept of a `laminar sheared layer' is here for the first time interpreted as having an upward-moving `free-surface' layer effect during deposition from hyper-concentrated flows. Channel-wall stoping involves unlithified turbiditic spillover sand in the levee sediment of the canyon wall that was washed out by the upper diluted parts of the high-concentration flows coming down the channel and leaving a niche in the wall that was filled with coarser channel-axis facies by the same flow (or later flows) when its aggradation reached the level of the niche. The contact between turbidite and

  3. Resolving Metazoan phyla divergence times during cambrian explosion by COX I protein molecules%COXI蛋白质分子确定“寒武纪生物大爆发”时期物种分歧时间

    Institute of Scientific and Technical Information of China (English)

    李可群

    2016-01-01

    By using the equation for calculating absolute evolutionary rates of biological molecules presented by the author and classic"molecular clock"Cytochrome oxidase subunit I ( COX I ) protein molecules , the metazoan phyla divergence times during Cambrian explosion were obtained .These results show that Protostomia and Deuterostomia separated at560 million years( myrs ) ago.The phyla divergence times of four major deuterstomia:Cephalochordata , Urochordata, Hemichordata and Echinodermata are 536 myrs ago, 540 myrs ago, 548 myrs ago and 545 myrs ago, respectively.These divergence times agree well with their fossil records , and are obviously better than those obtained by other current molecular clock methods .Our method may provide a new way to understand origins and evo-lution of creatures .%使用经典分子钟分子COX I蛋白质和作者提出的同源生物分子绝对进化速率计算公式,对“寒武纪生物大爆发”期间的一些主要动物门类分歧时间进行了计算。结果表明:原口动物和后口动物分歧时间为5.60亿年前,后口动物中的头索动物、尾索动物、半索动物和棘皮动物的分歧时间分别为5.36亿年前、5.40亿年前、5.48亿年前和5.45亿年前。这些结果与化石记录基本符合,明显好于现有分子钟理论得到的结果。为研究生物起源和演化提供了一种的新的思路和方法。

  4. Local Extrema of Periodic Function's Wavelet Transform

    Institute of Scientific and Technical Information of China (English)

    FAN Qi-bin; SONG Xiao-yan

    2005-01-01

    The theory of detecting ridges in the modulus of the continuous wavelet transform is presented as well as reconstructing signal by using information on ridges. To periodic signal we suppose Morlet wavelet as basic wavelet, and research the local extreme point and extrema of the wavelet transform on periodic function for the collection of signal's instantaneous amplitude and period.

  5. 77 FR 73452 - Grace Period Study

    Science.gov (United States)

    2012-12-10

    ... Patent and Trademark Office Grace Period Study ACTION: Proposed collection; comment request. SUMMARY: The...Collection@uspto.gov . Include ``0651- 00xx Grace Period Study comment'' in the subject line of the message... are not fully understood. Few studies in the past ten years have dealt with the grace period, and...

  6. 32 CFR 37.1325 - Periodic audit.

    Science.gov (United States)

    2010-07-01

    ... 32 National Defense 1 2010-07-01 2010-07-01 false Periodic audit. 37.1325 Section 37.1325 National... TECHNOLOGY INVESTMENT AGREEMENTS Definitions of Terms Used in This Part § 37.1325 Periodic audit. An audit of... awards. Appendix C to this part describes what such an audit may cover. A periodic audit of a...

  7. 21 CFR 573.240 - Calcium periodate.

    Science.gov (United States)

    2010-04-01

    ... with calcium hydroxide or calcium oxide to form a substance consisting of not less than 60 percent by... 21 Food and Drugs 6 2010-04-01 2010-04-01 false Calcium periodate. 573.240 Section 573.240 Food... Additive Listing § 573.240 Calcium periodate. The food additive calcium periodate may be safely used...

  8. 49 CFR 236.588 - Periodic test.

    Science.gov (United States)

    2010-10-01

    ... 49 Transportation 4 2010-10-01 2010-10-01 false Periodic test. 236.588 Section 236.588..., Train Control and Cab Signal Systems Inspection and Tests; Locomotive § 236.588 Periodic test. Except as provided in § 236.586, periodic test of the automatic train stop, train control, or cab signal...

  9. 27 CFR 70.244 - Payroll period.

    Science.gov (United States)

    2010-04-01

    ... 27 Alcohol, Tobacco Products and Firearms 2 2010-04-01 2010-04-01 false Payroll period. 70.244... Excise and Special (Occupational) Tax Limitations § 70.244 Payroll period. For purpose of determining the... established calendar period regularly used by the employer or other person levied upon for payroll or...

  10. Almost Periodic Time Scales and Almost Periodic Functions on Time Scales

    OpenAIRE

    Yongkun Li; Bing Li

    2015-01-01

    We propose some new concepts of almost periodic time scales and almost periodic functions on time scales and give some basic properties of these new types of almost periodic time scales and almost periodic functions on time scales. We also give some comments on a recent paper by Wang and Agarwal (2014) concerning a new almost periodic time scale.

  11. Counterexamples to mean square almost periodicity of the solutions of some SDEs with almost periodic coefficients

    Directory of Open Access Journals (Sweden)

    Omar Mellah

    2013-04-01

    Full Text Available We show that, contrarily to what is claimed in some papers, the nontrivial solutions of some stochastic differential equations with almost periodic coefficients are never mean square almost periodic (but they can be almost periodic in distribution.

  12. A highly specific test for periodicity

    International Nuclear Information System (INIS)

    We present a method that allows to distinguish between nearly periodic and strictly periodic time series. To this purpose, we employ a conservative criterion for periodicity, namely, that the time series can be interpolated by a periodic function whose local extrema are also present in the time series. Our method is intended for the analysis of time series generated by deterministic time-continuous dynamical systems, where it can help telling periodic dynamics from chaotic or transient ones. We empirically investigate our method's performance and compare it to an approach based on marker events (or Poincaré sections). We demonstrate that our method is capable of detecting small deviations from periodicity and outperforms the marker-event-based approach in typical situations. Our method requires no adjustment of parameters to the individual time series, yields the period length with a precision that exceeds the sampling rate, and its runtime grows asymptotically linear with the length of the time series

  13. A highly specific test for periodicity

    Energy Technology Data Exchange (ETDEWEB)

    Ansmann, Gerrit, E-mail: gansmann@uni-bonn.de [Department of Epileptology, University of Bonn, Sigmund-Freud-Straße 25, 53105 Bonn (Germany); Helmholtz Institute for Radiation and Nuclear Physics, University of Bonn, Nussallee 14–16, 53115 Bonn (Germany); Interdisciplinary Center for Complex Systems, University of Bonn, Brühler Straße 7, 53175 Bonn (Germany)

    2015-11-15

    We present a method that allows to distinguish between nearly periodic and strictly periodic time series. To this purpose, we employ a conservative criterion for periodicity, namely, that the time series can be interpolated by a periodic function whose local extrema are also present in the time series. Our method is intended for the analysis of time series generated by deterministic time-continuous dynamical systems, where it can help telling periodic dynamics from chaotic or transient ones. We empirically investigate our method's performance and compare it to an approach based on marker events (or Poincaré sections). We demonstrate that our method is capable of detecting small deviations from periodicity and outperforms the marker-event-based approach in typical situations. Our method requires no adjustment of parameters to the individual time series, yields the period length with a precision that exceeds the sampling rate, and its runtime grows asymptotically linear with the length of the time series.

  14. Gabor systems on discrete periodic sets

    Institute of Scientific and Technical Information of China (English)

    2009-01-01

    Due to its good potential for digital signal processing, discrete Gabor analysis has interested some mathematicians. This paper addresses Gabor systems on discrete periodic sets, which can model signals to appear periodically but intermittently. Complete Gabor systems and Gabor frames on discrete periodic sets are characterized; a sufficient and necessary condition on what periodic sets admit complete Gabor systems is obtained; this condition is also proved to be sufficient and necessary for the existence of sets E such that the Gabor systems generated by χE are tight frames on these periodic sets; our proof is constructive, and all tight frames of the above form with a special frame bound can be obtained by our method; periodic sets admitting Gabor Riesz bases are characterized; some examples are also provided to illustrate the general theory.

  15. Discovery of periodic patterns in spatiotemporal sequences

    OpenAIRE

    Cao, H.; Mamoulis, N; Cheung, DW

    2007-01-01

    In many applications that track and analyze spatiotemporal data, movements obey periodic patterns; the objects follow the same routes (approximately) over regular time intervals. For example, people wake up at the same time and follow more or less the same route to their work everyday. The discovery of hidden periodic patterns in spatiotemporal data could unveil important information to the data analyst. Existing approaches for discovering periodic patterns focus on symbol sequences. However,...

  16. Topological Chaos in Spatially Periodic Mixers

    OpenAIRE

    Finn, Matthew D.; Thiffeault, Jean-Luc; Gouillart, Emmanuelle

    2005-01-01

    Topologically chaotic fluid advection is examined in two-dimensional flows with either or both directions spatially periodic. Topological chaos is created by driving flow with moving stirrers whose trajectories are chosen to form various braids. For spatially periodic flows, in addition to the usual stirrer-exchange braiding motions, there are additional topologically-nontrivial motions corresponding to stirrers traversing the periodic directions. This leads to a study of the braid group on t...

  17. Periodic boundary conditions on the pseudosphere

    OpenAIRE

    Sausset, François; Tarjus, Gilles

    2007-01-01

    30 pages, minor corrections, accepted to J. Phys. A International audience We provide a framework to build periodic boundary conditions on the pseudosphere (or hyperbolic plane), the infinite two-dimensional Riemannian space of constant negative curvature. Starting from the common case of periodic boundary conditions in the Euclidean plane, we introduce all the needed mathematical notions and sketch a classification of periodic boundary conditions on the hyperbolic plane. We stress the ...

  18. The periodic table of elementary particles

    OpenAIRE

    Chung, Ding-Yu

    2000-01-01

    All leptons, quarks, and gauge bosons can be placed in the periodic table of elementary particles. The periodic table is derived from dualities of string theory and a Kaluza-Klein substructure for the six extra spatial dimensions. As a molecule is the composite of atoms with chemical bonds, a hadron is the composite of elementary particles with hadronic bonds. The masses of elementary particles and hadrons can be calculated using the periodic table with only four known constants: the number o...

  19. Casimir electromotive force in periodic configurations

    OpenAIRE

    Fateev, Evgeny G.

    2016-01-01

    The possibility in principle of the existence of Casimir electromotive force (EMF) is shown for nonparallel nanosized metal plates arranged in the form of a periodic structure. It is found that EMF does not appear in strictly periodic structures with parallel plates. However, when the strict periodicity is disturbed in nonparallel plates, EMF is generated, and its value is equal to the number of pairs of plates in a configuration. Moreover, there are some effective parameters of the configura...

  20. Do triatomic molecules echo atomic periodicity?

    Energy Technology Data Exchange (ETDEWEB)

    Hefferlin, R., E-mail: hefferln@southern.edu; Barrow, J. [Southern Adventist University, PO Box 370, Collegedale, Tennessee 37315 (United States)

    2015-03-30

    Demonstrations of periodicity among triatomic-molecular spectroscopic constants underscore the role of the periodic law as a foundation of chemistry. The objective of this work is to prepare for another test using vibration frequencies ν{sub 1} of free, ground-state, main-group triatomic molecules. Using data from four data bases and from computation, we have collected ν{sub 1} data for molecules formed from second period atoms.