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Sample records for burrowing crab neohelice

  1. Population structure of the burrowing crab Neohelice granulata (Brachyura, Varunidae in a southwestern Atlantic salt marsh

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    Sabrina Angeletti

    2015-07-01

    Full Text Available Neohelice granulata inhabits estuarine and protected coastal areas in temperate regions and is the most dominant decapod crustacean in the Bahía Blanca Estuary, Argentina. The population structure was studied during a year in a SW Atlantic salt marsh located in the Bahía Blanca Estuary. Crabs were sampled monthly from August 2010 to July 2011. The maximum observed density was 30 crabs m-2 in February and 70 burrows m-2 in May. The maximum carapace width (CW was 32 and 27.5 mm in males and females respectively. Medium size crabs were between 16 and 20 mm CW. Significantly smaller sized crabs were observed at the lower intertidal regions (P < 0.05. The sex ratio was favorable for males and was significantly different from the expected 1:1 (P < 0.05. The recruitment of unsexed juveniles crabs (CW <6.5 mm was observed throughout the year and the presence of ovigerous females from October to February indicated seasonal reproduction. The average size of ovigerous females was CW = 20.8 mm and the smallest ovigerous female measured was 16 mm CW. For the first time, the population structure of the most important macro-invertebrate is analyzed in the Bahía Blanca Estuary. This study may help to make decisions in the area, where anthropic action is progressing day by day.

  2. Metals from mine waste as potential cause of oxidative stress in burrowing crab Neohelice granulata from San Antonio bay.

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    Giarratano, Erica; Gil, Mónica N; Marinho, Carmen H; Malanga, Gabriela

    2016-10-01

    The Natural Protected Area San Antonio bay is of particular importance for its congregation of migratory shorebirds and it has been declared one of the Western Hemisphere Shorebird Reserve Network International site (WHSRN). Present study represents the first assessment of variation on oxidative stress biomarkers in male crab Neohelice granulata from San Antonio bay (Río Negro, Argentina) under field conditions, associated mainly to metal contamination coming from passive mining wastes. Three sites were sampled once every three months from November 2012 to August 2013 within this sea inlet (Pile, Fishery and Port) and a control site at the southeast of the bay (Punta Perdices). Accumulation of Ni, Zn, Cr and Al varied only with seasons although without a constant trend, meanwhile Cd, Cu and Pb also varied among sites being highest in Pile and Port. Biochemical results indicated that variations in catalase activity was only site specific being maximum in Pile; meanwhile lipid radical, α-tocopherol and metallothioneins were only seasonal specific being higher in autumn and winter. Seasonal variation was also found for total thioles, being the content higher in summer and autumn than in winter. Correlation analysis revealed that malondialdehyde and α-tocopherol have a positive association with Al and negative with Ni, meanwhile GST has a positive association with Fe. Crabs from the closest area to the waste pile did not exhibit a differentiated oxidative pressure despite the higher accumulation of metals. It is possible that crabs from contaminated areas have developed a tolerance to metals, indicating a strong ecotoxicological selective pressure. More studies are needed to assess whether there is a transfer of metals through the food chain.

  3. The crab Neohelice (= Chasmagnathus) granulata: an emergent animal model from emergent countries

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    Spivak, Eduardo D.

    2010-09-01

    Neohelice granulata (previously known as Chasmagnathus granulata and C. granulatus) is a burrowing semiterrestrial crab found in the intertidal zone of estuaries, salt marshes and mangroves of the South-western Atlantic Ocean. Beginning in the late 1989s, an explosion of publications appeared in international journals dealing with its ecology, physiology, toxicology and behavior. A bibliometric analysis using the Scopus database allowed detecting 309 papers that deal with this species during the period 1986-2009. The number of papers per year increased continuously, reaching a mean annual value of 22.6 during the last 5 years; a great majority of them were authored by researchers from Argentina and Brazil. Neohelice granulata has become now one of the most studied crab species, after Carcinus maenas, Callinectes sapidus, Scylla serrata and Cancer pagurus and C. magister, and it can be considered as an emergent animal model for biochemical, physiological and ecological research.

  4. Habitat shifts and spatial distribution of the intertidal crab Neohelice ( Chasmagnathus ) granulata Dana

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    Casariego, Agustina Mendez; Alberti, Juan; Luppi, Tomás; Daleo, Pedro; Iribarne, Oscar

    2011-08-01

    Intertidal zones of estuaries and embayments of the SW Atlantic are dominated by the semiterrestrial burrowing grapsid crab, Neohelice ( Chasmagnathus) granulata, and characterized by extensive mud flats surrounded by salt marshes. In this work we examined spatial patterns of distribution of N. granulata during two years to explain their movement patterns. The results of the population sampling showed segregation by sex and size throughout the intertidal, with seasonal variations in densities and different condition indices for adults and juveniles at the different zones. The comparison of seasonal activity (ambulatory activity outside burrows) between marshes and mudflats shows that short term (e.g. daily) variations in activity were controlled by tides. Crabs were active at high tides but increased their activity on days with higher tidal amplitude. Seasonal activity showed that at both areas, females remain with low activity except for a peak in winter, while males showed the highest activity during summer in the mudflat zone, but not so in the marsh. This pattern can be the response to differences in stress tolerance, suggesting that high temperatures are limiting the performance of adult crabs during summer, especially at the marsh where physical conditions can be more critical. The spatial size segregation can be explained by differential mortality in each zone (estimated with tethered crabs), and by the juvenile movement between these zones (estimated with movement traps). Juvenile mortality is higher at the mudflat, while adult mortality is higher in the marsh. Smaller juveniles moved to the marsh, where the mortality is lower, and the larger juveniles moved towards the mudflat. This mortality is due almost exclusively to cannibalism, so our results suggest that this movement of different size classes between zones is controlled, at least in part, by intraspecific predation.

  5. Great unexpected differences between two populations of the intertidal crab Neohelice granulata inhabiting close but contrasting habitats (Crustacea: Decapoda: Brachyura

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    Eduardo D. Spivak

    Full Text Available ABSTRACT Two populations of the Southwestern Atlantic burrowing crab, Neohelice granulata (Dana, 1851, are separated by only 190 km. They inhabit bays that drain into a Patagonian gulf in the southernmost limit of the geographical distribution of the species: San Antonio Bay (SAO and San José Gulf (RSJ. The population from SAO has been extensively studied, whereas that of RSJ has not. The main goal of this study is to compare the ecological and habitat characteristics of the two populations, based on samples collected on successive days during the middle of the reproductive season of the crab. The substrate of SAO is composed of a high proportion of gravel and scarce organic matter, whereas the substrate of RSJ presents twice as much silt and clay, and greater organic matter content. Crabs of both sexes were larger at RSJ. Fecundity and reproductive output were greater at RSJ. The size of females at maturity was smaller and their range was shorter at RSJ. Burrows were longer and wider at RSJ. Our results indicate that there are strong differences between the two populations, which might be correlated with local environmental differences, mainly substrate granulometry and food availability.

  6. Glyceroneogenesis in the hepatopancreas of the crab Neohelice granulata: Diet, starvation and season effects.

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    Sarapio, E; Santos, J T; Model, J F A; De Fraga, L S; Vinagre, A S; Martins, T L; Da Silva, R S M; Trapp, M

    2017-02-22

    We determined the activity of glyceroneogenesis from [2-(14)C]-pyruvate, the phosphoenolpyruvate carboxykinase activity, [2-(14)C]-pyruvate oxidation and total lipid levels in the hepatopancreas of the crab Neohelice granulata fed with a carbohydrate-rich (HC) diet or a high-protein (HP) diet and then subjected to 5weeks of starvation, in summer and winter, to determine whether the seasonal adjustments of lipid metabolism to food scarcity are modulated by the composition of the diet previously given to the crabs. The results demonstrated that glyceroneogenesis is an active pathway in N. granulata hepatopancreas, and is regulated by seasonal variations, diet composition and starvation. This study showed that in summer the increase in the hepatopancreas glyceroneogenesis activity is among the strategies used by N. granulata fed an HP diet, to maintain the triglyceride/fatty acid cycle during starvation, a normal condition in the biological cycle of this crab. However, the administration of an HC diet reduced the glyceroneogenesis capacity in response to starvation in summer. In winter, the decrease in the glyceroneogenesis capacity in both fed (HP and HC diets) and starved crabs seems to be a strategy to reduce energy consumption and/or requirement. In contrast to the summer results, the incorporation of [2-(14)C]-pyruvate into (14)CO2 was markedly higher in both diet (HC and HP) groups and in starved crabs during the winter. Four decades after the first study describing the glyceroneogenesis pathway in rat white adipose tissue, this pathway is evidenced for the first time in a crustacean.

  7. Damage caused during hypoxia and reoxygenation in the locomotor muscle of the crab Neohelice granulata (Decapoda: Varunidae).

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    Geihs, Márcio Alberto; Vargas, Marcelo Alves; Nery, Luiz Eduardo Maia

    2014-06-01

    The aim of this work was to determine whether different durations of severe hypoxia (0.5 mg O2 L(-1)) followed by reoxygenation cause damage to the locomotor muscle of the crab Neohelice granulata. We evaluated reactive oxygen species (ROS), lipid peroxidation (LPO), mitochondrial membrane potential, and aerobic fiber area of the locomotor muscle after different periods of hypoxia (1, 4, or 10h) followed by 30 or 120 min of reoxygenation. Additionally, changes in cell volume, mitochondrial dysfunction, and infiltration of hemocytes were evaluated after hypoxia and a subsequent 2, 24, or 48 h of reoxygenation. After hypoxia, neither ROS nor LPO increased. However, mitochondrial membrane potential and aerobic fiber area decreased in a time-dependent manner. After reoxygenation, the ROS and LPO levels increased and mitochondrial membrane potential decreased, but these quickly recovered in crabs exposed to 4h of hypoxia. On the other hand, alterations of mitochondria resulted in morphological changes in aerobic fibers, which required more time to recover during reoxygenation after 10h of hypoxia. The locomotor muscles of the crab N. granulata suffer damage after hypoxia and reoxygenation. The intensity of this damage is dependent on the duration of hypoxia. In all experimental situations analyzed, the locomotor muscle of this crab was capable of recovery.

  8. Variation of Sediment Properties among the Radial Profiles of Fiddler Crab Burrows in Mangrove Ecosystem

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    Mohammad Mokhtari

    2016-01-01

    Full Text Available Fiddler crabs burrow creates oxic-anoxic interfaces on the burrow walls. Accordingly burrow walls represent the transitions site between oxic and anoxic condition where the sediment properties varied significantly across it. In this study the burrows of three species of fiddler crabs including Uca rosea, Uca forcipata and Uca pardussumieri were sampled at three depth layers. Sediment properties of burrow walls including; temperature, redox potential, pH, density, porosity, water content, organic content, chlorophyll content and solid phase iron pools were measured to determine the magnitude of burrow effects on mangrove sediments. The results indicated that U. paradussumieri effectively reduced the sediment porosity of surrounding sediments down to 45%. Oxidized layer was more extended around U. paradussumieri burrows. Burrow walls of U. forcipata and U. paradussumieri contain higher water content than ambient sediment and burrows of U. rosea efficiently decreased the organic content of sediment. The PCA biplots indicated that the burrow walls of the all three species of fiddler crabs at 3 and 8 cm depth were correlated with oxidized iron. Ambient sediments of U. forcipata habitat were correlated with reduced iron and organic content. At 20 cm depth, burrow walls of U. paradussumieri were highly correlated with water content, while ambient sediment was correlated with reduced iron. The results of this study revealed that the thickness of oxidized layer varied according to sediment depth and burrow volume. Consequently the burrow effect varied significantly among different species of fiddler crabs as results of different habitat characteristics, sediment types and crab size.

  9. Hatching and larval export of the intertidal crab Neohelice granulata in Mar Chiquita coastal lagoon, Argentina

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    Guillermina Sánchez Vuichard; Nahuel Farías; Tomás Luppi

    2013-01-01

    Synchronization in the events of the reproductive cycle in female Neohelice granulata Dana, 1851 were studied from samples taken weekly and biweekly from September to December 2006 in the Laguna Mar Chiquita. The timing and larval hatching and synchronicity were inferred from numbers of ovigerous females and observing the stages of embryonic development. Synchronization in larval hatching also was observed in females in experiments in dark for a period of 48 hours, at three different saliniti...

  10. Crab burrows as conduits for groundwater-surface water exchange in Bangladesh

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    Stahl, Mason O.; Tarek, M. H.; Yeo, Darren C. J.; Badruzzaman, A. B. M.; Harvey, Charles F.

    2014-12-01

    Groundwater recharge affects water budgets and groundwater quality on the deltas and floodplains of South and Southeast Asia. Rain and flooding rivers recharge groundwater during the monsoon; irrigated rice fields and surface water bodies recharge aquifers during the dry season. Groundwater throughout the region is severely contaminated by arsenic, and recent research suggests that quantifying and characterizing recharge is important to understand whether recharge flushes or mobilizes arsenic from aquifers. At a field site in Bangladesh, we found that burrows of terrestrial crabs short-circuit low-permeability surface sediments, providing the primary conduit for recharge. We combine field observations along with a model that couples isotope and water balances to quantify the effect of crab burrows on aquifer recharge. Given the wide distribution of burrowing crabs and the surficial geology, we suggest that crab burrows provide widespread conduits for groundwater recharge.

  11. Serotonin effects in the crab Neohelice granulata: Possible involvement of two types of receptors in peripheral tissues.

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    Inohara, Elen Thegla Sander; Pinto, Charles Budazewsky; Model, Jorge Felipe Argenta; Trapp, Márcia; Kucharski, Luiz Carlos; Da Silva, Roselis Silveira Martins; Vinagre, Anapaula Sommer

    2015-07-01

    In crustaceans, serotonin (5-HT) controls various physiological processes, such as hormonal secretion, color changes, reproduction, and metabolism. Since 5-HT injections cause hyperglycemia, this study was designed to further investigate this action of 5-HT in the crab Neohelice granulate, fed with a high-carbohydrate (HC) or a high-protein (HP) diet. The effects of pre-treatment with mammalian 5-HT receptor antagonists, cyproheptadine and methiothepin, were also investigated. A series of in vivo experiments with (3)H-5-HT was carried out in order to investigate the presence of putative receptors in peripheral tissues. Since gills were the tissue with the highest labeling in in vivo experiments, in vitro studies with isolated anterior and posterior gills were also conducted. Cyproheptadine blocked the hyperglycemic effect of 5-HT in HP-fed crabs. Methiothepin reduced glycogen levels in the anterior gills of HP crabs and partially blocked the 5-HT-like posture. The injection of (3)H-5-HT identified specific binding sites in all the tissues studied and revealed that the binding can be influenced by the type of diet administered to the crabs. Incubation of the anterior and posterior gills with (3)H-5-HT and 5-HT confirmed the specificity of the binding sites. Both antagonists inhibited (3)H-5-HT binding. In conclusion, this study highlights the importance of serotonin in the control of glucose homeostasis in crustaceans and provides evidences of at least two types of 5-HT binding sites in peripheral tissues. Further studies are necessary to identify the structure of these receptors and their signaling pathways.

  12. First field record of mangrove crab Ucides cordatus (Crustacea: Decapoda: Ucididae recruits co-inhabiting burrows of conspecific crabs

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    Anders Jensen Schmidt

    2009-12-01

    Full Text Available Recruits of the mangrove crab Ucides cordatus (Linnaeus, 1763, rarely encountered in the field were found co-inhabiting burrows of larger male and female conspecifics in the mangrove forest. They were located in the sediment of the inner walls and burrow plugs. Average carapace width (CW of the hosting and co-inhabiting crabs was 3.8 ± 0.20 and 0.9 ± 0.03, respectively. As shown by the size-frequency distribution, while most recruits leave the conspecific burrows after reaching 1.0 cm CW, some stay until they reach a size of 2.5 cm CW. The results of this study contribute to a better understanding of recruitment patterns in this ecologically and economically important mangrove crab species. Follow-up studies are however needed to fully determine the role of conspecific burrows for juvenile habitat choice and survivorship in U. cordatus.

  13. Melatonin as a Signaling Molecule for Metabolism Regulation in Response to Hypoxia in the Crab Neohelice granulata

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    Fábio Everton Maciel

    2014-12-01

    Full Text Available Melatonin has been identified in a variety of crustacean species, but its function is not as well understood as in vertebrates. The present study investigates whether melatonin has an effect on crustacean hyperglycemic hormone (CHH gene expression, oxygen consumption (VO2 and circulating glucose and lactate levels, in response to different dissolved-oxygen concentrations, in the crab Neohelice granulata, as well as whether these possible effects are eyestalk- or receptor-dependent. Melatonin decreased CHH expression in crabs exposed for 45 min to 6 (2, 200 or 20,000 pmol·crab−1 or 2 mgO2·L−1 (200 pmol·crab−1. Since luzindole (200 nmol·crab−1 did not significantly (p > 0.05 alter the melatonin effect, its action does not seem to be mediated by vertebrate-typical MT1 and MT2 receptors. Melatonin (200 pmol·crab−1 increased the levels of glucose and lactate in crabs exposed to 6 mgO2·L−1, and luzindole (200 nmol·crab−1 decreased this effect, indicating that melatonin receptors are involved in hyperglycemia and lactemia. Melatonin showed no effect on VO2. Interestingly, in vitro incubation of eyestalk ganglia for 45 min at 0.7 mgO2·L−1 significantly (p < 0.05 increased melatonin production in this organ. In addition, injections of melatonin significantly increased the levels of circulating melatonin in crabs exposed for 45 min to 6 (200 or 20,000 pmol·crab−1, 2 (200 and 20,000 pmol·crab−1 and 0.7 (200 or 20,000 pmol·crab−1 mgO2·L−1. Therefore, melatonin seems to have an effect on the metabolism of N. granulata. This molecule inhibited the gene expression of CHH and caused an eyestalk- and receptor-dependent hyperglycemia, which suggests that melatonin may have a signaling role in metabolic regulation in this crab.

  14. Effects of glyphosate on egg incubation, larvae hatching, and ovarian rematuration in the estuarine crab Neohelice granulata.

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    Avigliano, Luciana; Alvarez, Natalia; Loughlin, Camila Mac; Rodríguez, Enrique Marcelo

    2014-08-01

    Ovigerous females of the estuarine crab (Neohelice granulate) were exposed to both pure glyphosate (2.5 mg/L and 5 mg/L) and a glyphosate formulation (Roundup Ultramax, containing glyphosate at 2.5 mg/L acid equivalent). At the end of the egg incubation period, a significant reduction in the number of hatched larvae was seen as a result of Roundup exposure. Additionally, several larvae abnormalities were seen in both pure glyphosate (2.5 mg/L) and Roundup treatments, such as hydropsy and hypopigmented eyes, and atrophied eyes were observed in the Roundup treatment. To evaluate the effect of the herbicide on ovarian rematuration, females remained exposed for 32 d. Pure glyphosate at 2.5 mg/L stimulated ovarian maturation over control levels, mainly in terms of a higher gonadosomatic index and a higher percentage of vitellogenic oocytes. A plausible hypothesis to be tested in further experiments is that exposure to glyphosate disrupts the hormonal system controlling reproduction.

  15. Effect of mangrove restoration on crab burrow density in Luoyangjiang Estuary, China

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    Wei Li

    2015-07-01

    Full Text Available Background Mangrove restoration seeks to restore or rebuild degraded mangrove systems. The methods of mangrove restoration include ecological projects and restoration-oriented technologies, the latter of which are designed to restore the structure, processes as well as related physical, chemical and biological characteristics of wetlands and to ensure the provision of ecosystem services. As important components of mangrove ecosystem, benthic organisms and crabs play a key role in nutrient cycling. In addition, mangrove restoration, such as vegetation restoration measures, can lead to changes in the benthic faunal communities. This study investigates whether the presence of different mangrove species, age and canopy cover of mangrove communities affect the density of crab burrows. Methods The Luoyangjiang Estuary, in the southeast of Fujian Province, was selected as our research area. A survey, covering 14 sites, was conducted to investigate the impacts of mangrove restoration on the density of crab burrows in four rehabilitated forests with different stand ages and canopy. Results It was found that differences in vegetation types had a large impact on crab density and that the density of crab burrows was lower on exposed beaches (non-mangrove than under mature Kandelia candel, Aegiceras corniculatum and Avicennia marina communities. In general, the amount of leaf litter and debris on mangrove mudflats was greater than on the beaches as food sources for crabs. Two-factor analysis of variance (ANOVA shows that changes in mangrove species and age since restoration had different effects on crab burrow density. The effect of canopy cover was highly significant on crab burrow density. Conclusions The results suggest that in the process of mangrove restoration the combined effects of mangrove stand age, canopy cover and other factors should be taken into account. This study further supports the findings of the future scientific research and practice on

  16. Hatching and larval export of the intertidal crab Neohelice granulata in Mar Chiquita coastal lagoon, Argentina

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    Guillermina Sánchez Vuichard

    2013-06-01

    Full Text Available Synchronization in the events of the reproductive cycle in female Neohelice granulata Dana, 1851 were studied from samples taken weekly and biweekly from September to December 2006 in the Laguna Mar Chiquita. The timing and larval hatching and synchronicity were inferred from numbers of ovigerous females and observing the stages of embryonic development. Synchronization in larval hatching also was observed in females in experiments in dark for a period of 48 hours, at three different salinities (10, 23 and 33 ppm. In addition plankton sampling were performed in order to study larval exportation at the field and its link to the tidal and light/dark cycles. We found that ovigerous females of N. granulata have a marked synchronization in embryonic development which results in that most of berried females are close to hatching within a period of maximum tidal range (days. Within this period, there is a synchronization of hatching at a time scale of hours, governed by environmental conditions. The salinity range used in this study (10-32‰ did not affect hatching synchronicity neither time to hatch. Hatching was synchronized according to endogenous rhythms governed mainly by the tidal cycle and secondarily by the breadth of it. It is also conditioned by the light-dark cycle through an exogenous cycle, so that the hatchings would occur mostly at night high tides.

  17. The vibrational signals that male fiddler crabs ( Uca lactea) use to attract females into their burrows

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    Takeshita, Fumio; Murai, Minoru

    2016-06-01

    In some fiddler crab species, males emit vibrations from their burrows to mate-searching females after they have attracted a female to the burrow entrance using a waving display. Although the vibrations are considered acoustic signals to induce mating, it has not been demonstrated whether the vibrations attract the females into the burrow and, consequently, influence females' mating decisions. We investigated the structures and patterns of the vibrations using a dummy female and demonstrated experimentally a female preference for male vibrations in Uca lactea in the field. The acoustic signals consisted of repetitions of pulses. The dominant frequency of the pulses decreased with male carapace width. The pulse length decreased slightly with an increasing number of vibrational repetitions, and the pulse interval increased with increasing repetitions. These factors imply that the vibrations convey information on male characteristics, such as body size and stamina. In the experiment on female mate choice, the females significantly preferred males with higher pulse repetition rates when they were positioned at the entrance of the burrow, indicating that the females use the male vibrational signals to decide whether to enter the burrow. However, females showed no preference for the vibrations once they were inside a burrow, i.e., whether they decided to copulate, suggesting that the vibrations do not independently affect a female's final decision of mate choice. The vibrations inside the burrow might influence a female's decision by interaction with other male traits such as the burrow structure.

  18. Burrowing and foraging activity of marsh crabs under different inundation regimes

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    New England salt marshes are susceptible to degradation and habitat loss as a result of increased periods of inundation as sea levels rise. Increased inundation may exacerbate marsh degradation that can result from crab burrowing and foraging. Most studies to date have focused on...

  19. Regulation of ion transport by pH and [HCO3-] in isolated gills of the crab Neohelice (Chasmagnathus) granulata.

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    Tresguerres, Martin; Parks, Scott K; Sabatini, Sebastian E; Goss, Greg G; Luquet, Carlos M

    2008-03-01

    Posterior isolated gills of Neohelice (Chasmagnathus) granulatus were symmetrically perfused with hemolymph-like saline of varying [HCO3-] and pH. Elevating [HCO3-] in the saline from 2.5 to 12.5 mmol/l (pH 7.75 in both cases) induced a significant increase in the transepithelial potential difference (Vte), a measure of ion transport. The elevation in [HCO3-] also induced a switch from acid secretion (-43.7 +/- 22.5 microequiv.kg(-1).h(-1)) in controls to base secretion (84.7 +/- 14.4 microequiv.kg(-1).h(-1)). The HCO3(-)-induced Vte increase was inhibited by basolateral acetazolamide (200 micromol/l), amiloride (1 mmol/l), and ouabain (5 mmol/l) but not by bafilomycin (100 nmol/l). The Vte response to HCO3(-) did not take place in Cl(-)-free conditions; however, it was unaffected by apical SITS (2 mmol/l) or DIDS (1 mmol/l). A decrease in pH from 7.75 to 7.45 pH units in the perfusate also induced a significant increase in Vte, which was matched by a net increase in acid secretion of 67.8 +/- 18.4 microequiv kg(-1) h(-1). This stimulation was sensitive to basolateral acetazolamide, bafilomycin, DIDS, and Na+-free conditions, but it still took place in Cl(-)-free saline. Therefore, the cellular response to low pH is different from the HCO3(-)-stimulated response. We also report V-H+-ATPase- and Na+-K+-ATPase-like immunoreactivity in gill sections for the first time in this crab. Our results suggest that carbonic anhydrase (CA), basolateral Na+/H+ exchangers and Na+-K+-ATPase and apical anion exchangers participate in the HCO3(-)-stimulated response, while CA, apical V-H+-ATPase and basolateral HCO3(-)-dependent cotransporters mediate the response to low pH.

  20. Evaluation of insecticide impregnated baits for control of mosquito larvae in land crab burrows on French Polynesian atolls.

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    Lardeux, Frederic; Sechan, Yves; Faaruia, Marc

    2002-07-01

    Land crab burrows are larval mosquito habitats of major significance in the Pacific region. They are constituted by a sinuous tunnel leading to a chamber in contact with the water table, where mosquito larvae proliferate. Controlling larvae in these sites is difficult, because the configuration of burrows prevents the use of standard techniques. An experiment was carried out in French Polynesia to control Aedes polynesiensis Marks and Culex spp. breeding in burrows of the land crab Cardisoma carnifex (Herbst). The technique was based on the crab's behavior, which involves the crab carrying food into its burrow. It was shown that appetizing baits impregnated with an insecticide were carried by crabs into the flooded chamber of their burrows. A field treatment of burrows was carried out by sowing insecticide impregnated baits on the ground. The treatment coverage was almost perfect and the easy implementation of the technique enabled large areas to be treated in a short time. The bait was developed by compacting various flours, which easily incorporate a large variety of insecticide formulations. Although the baits can be easily stocked, a reliable insecticide is still to be found. The results indicate that our technique could be a method of choice for treating crab burrows.

  1. Air exposure behavior of the semiterrestrial crab Neohelice granulata allows tolerance to severe hypoxia but not prevent oxidative damage due to hypoxia-reoxygenation cycle.

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    de Lima, Tábata Martins; Geihs, Márcio Alberto; Nery, Luiz Eduardo Maia; Maciel, Fábio Everton

    2015-11-01

    The air exposure behavior of the semi-terrestrial crab Neohelice granulata during severe hypoxia was studied. This study also verified whether this behavior mitigates possible oxidative damage, namely lipoperoxidation, caused by hypoxia and reoxygenation cycles. The lethal time for 50% of the crabs subjected to severe hypoxia (0.5 mgO2 · L(-1)) with free access to air was compared to that of crabs subjected to severe hypoxia without access to air. Crabs were placed in aquaria divided into three zones: water (when the animal was fully submersed), land (when the animal was completely emerged) and intermediate (when the animal was in contact with both environments) zones. Then the crabs were held in this condition for 270 min, and the time spent in each zone was recorded. Lipid peroxidation (LPO) damage to the walking leg muscles was determined for the following four experimental conditions: a--normoxic water with free access to air; b--hypoxic water without access to air; c--hypoxic water followed by normoxic water without air access; and d--hypoxic water with free access to air. When exposed to hypoxic water, N. granulata spent significantly more time on land, 135.3 ± 17.7 min, whereas control animals (exposed to normoxic water) spent more time submerged, 187.4 ± 20.2 min. By this behavior, N. granulata was able to maintain a 100% survival rate when exposed to severe hypoxia. However, N. granulata must still return to water after periods of air exposure (~ 14 min), causing a sequence of hypoxia/reoxygenation events. Despite increasing the survival rate, hypoxia with air access does not decrease the lipid peroxidation damage caused by the hypoxia and reoxygenation cycle experienced by these crabs.

  2. BURROW ARCHITECTURE OF RED GHOST CRAB OCYPODE MACROCERA (H. MILNE-EDWARDS, 1852 : A CASE STUDY IN INDIAN SUNDARBANS

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    Sourabh Kumar Dubey

    2013-12-01

    Full Text Available A study on burrow architecture and burrow morphology of the red ghost crab (Ocypode macrocera was carried out at the southern proximity of the Sagar island (21°37.973' N, to E 88° 04.195', western sector of Indian Sundarbans that faces the regular tidal influences of Bay of Bengal. Ocypode macrocera constructs burrows that are highly species specific and used by single individual. Four types of burrow patterns were observed like ‘I’, ‘J’ ‘U’ and ‘semi-U’ type with different sizes as revealed by POP casting. Important physic-chemical parameters like air temperature, temperature and salinity of the water were significantly varied (P < 0.05 throughout seasons in the Ocypode zone. Burrow sand column temperature were also significantly varied from ambient air temperature thus exhibiting preference for cooler subterranean residential compartment. The digging behaviour of Ocypodes enhances oxygenation in the ground soil and facilitates decomposition of organic materials, nutrient recycling, entrapping the sediments and mangrove seedlings and helps the process of bioturbation. As per the preliminary observations it was suggested that burrow shape is directly related to tidal action and metabolic activities of the crab are strongly correlated with burrow microenvironment. They are adapted to the different sediment conditions, tidal fluctuations, varying salinity gradients, air and water temperatures and other environmental fluctuations.

  3. Crustacean parasites associated with hermit crabs from the western Mediterranean Sea, with first documentation of egg predation by the burrowing barnacle Trypetesa lampas (Cirripedia: Acrothoracica: Trypetesidae).

    Science.gov (United States)

    Williams, Jason D; Gallardo, Alejandra; Murphy, Angela E

    2011-03-01

    Parasitic isopods (family Bopyridae) and burrowing barnacles (family Trypetesidae) infesting hermit crabs were investigated from shallow subtidal collections made along the southeastern coast of Spain in 2009. A total of 713 specimens of Clibanarius erythropus (Latreille, 1818) and 82 Calcinus tubularis (L., 1767) were examined. Gastropod shells and worm tubes inhabited by hermit crabs were collected by hand while snorkeling and were cracked to determine host species, size, sex and presence of eggs. Two species of bopyrid isopods were found on C. erythropus: the branchial parasite Bopyrissa fraiseii (Carayon, 1943) and the abdominal parasite Parathelges cardonae Codreanu and Codreanu in Codreanu, 1968. Among all C. erythropus examined, Bopyrissa fraiseii was found on 0.6% of hermit crabs and P. cardonae was found on 0.3%. A redescription of P. cardonae is provided and the species is documented with light and scanning electron microscopy for the first time. No Calcinus tubularis harbored parasitic isopods, but one specimen was parasitized by an unidentified rhizocephalan barnacle of the genus Septosaccus (1.2%). The burrowing barnacle Trypetesa lampas (Hancock, 1849) was found associated with both hermit crab species and evidence of predation on host eggs by this barnacle is shown for the first time. Trypetesa lampas was found in 4.2% of the shells collected. The present study expands our knowledge of the parasite fauna of hermit crabs from the Mediterranean Sea and indicates that additional research is needed to determine the impact of trypetesid egg predators on hermit crab populations.

  4. Mangrove carbon sink. Do burrowing crabs contribute to sediment carbon storage? Evidence from a Kenyan mangrove system

    Science.gov (United States)

    Andreetta, Anna; Fusi, Marco; Cameldi, Irene; Cimò, Filippo; Carnicelli, Stefano; Cannicci, Stefano

    2014-01-01

    Mangrove ecosystems are acknowledged as a significant carbon reservoir, with a potential key role as carbon sinks. Little however is known on sediment/soil capacity to store organic carbon and the impact of benthic fauna on soil organic carbon (SOC) stock in mangrove C-poor soils. This study aimed to investigate the effects of macrobenthos on SOC storage and dynamic in mangrove forest at Gazi Bay (Kenya). Although the relatively low amount of organic carbon (OC%) in these soils, they resulted in the presence of large ecosystem carbon stock comparable to other forest ecosystems. SOC at Gazi bay ranged from 3.6 kg m- 2 in a Desert-like belt to 29.7 kg m- 2 in the Rhizophora belt considering the depth soil interval from 0 cm to 80 cm. The high spatial heterogeneity in the distribution and amount of SOC seemed to be explained by different dominant crab species and their impact on the soil environment. A further major determinant was the presence, in the subsoil, of horizons rich in organic matter, whose dating pointed to their formation being associated with sea level rise over the Holocene. Dating and soil morphological characters proved to be an effective support to discuss links between the strategies developed by macrobenthos and soil ecosystem functioning.

  5. Carriage of Clostridium perfringens by benthic crabs in a sewage-polluted estuary.

    Science.gov (United States)

    La Sala, Luciano F; Redondo, Leandro M; Díaz Carrasco, Juan M; Pereyra, Ana María; Farber, Marisa; Jost, Helen; Fernández-Miyakawa, Mariano E

    2015-08-15

    The Estuary of Bahía Blanca (EBB), Argentina, is an important wetland under intense sewage pollution. We investigated the occurrence of Clostridium perfringens (CP) in populations of two benthic crabs (Neohelice granulata and Cyrtograpsus angulatus) and in sediment from the EBB. CP was found in 49.1% of the crabs and all of the isolates were identified as type A. The alpha (cpa) and enterotoxin (cpe) encoding genes were identified. Genetic analyses identified 13 novel sequence types, and found no clustering among isolates, suggesting that CP is not part of the crabs' commensal flora. CP carriage was 51 times more likely in crabs from the area nearest sewage outfalls compared with crabs from a reference site. Our in vitro experiments suggest that the carriage of CP in crabs is transient. The use of these benthic crabs as monitoring organisms of sewage pollution in coastal habitats is proposed.

  6. Good Crab, Bad Crab

    Science.gov (United States)

    Are crabs friends or foes of marsh grass, benefit or detriment to the salt marsh system? We examined Uca pugilator (sand fiddler) and Sesarma reticulatum (purple marsh crab) with Spartina patens (salt marsh hay) at two elevations (10 cm below MHW and 10 cm above MHW) in mesocosms...

  7. Dry season distribution of land crabs, Gecarcinus quadratus (Crustacea: Gecarcinidae), in Corcovado National Park, Costa Rica.

    Science.gov (United States)

    Griffiths, Megan E; Mohammad, Basma A; Vega, Andres

    2007-03-01

    The land crab Gecarcinus quadratus is an engineering species that controls nutrient cycling in tropical forests. Factors regulating their coastal distribution are not fully understood. We quantified land crab distribution during the dry season at Sirena Field Station in Corcovado National Park, Costa Rica, and found that land crab burrow density decreases with increasing distance from the ocean. Leaf litter depth and tree seedling density are negatively correlated with land crab burrow density. Burrows are strongly associated with sand substrate and burrow density is comparatively low in clay substrate. Results suggest that G. quadratus is limited to a narrow coastal zone with sand substrate, and this distribution could have profound effects on plant community structure.

  8. Epidemiology of acanthocephalan infections in crabs from the Bahía Blanca Estuary, Argentina.

    Science.gov (United States)

    La Sala, L F; Perez, A M; Martorelli, S R

    2012-12-01

    This study was conducted in two populations of crabs, Cyrtograpsus angulatus and Neohelice granulata from the Bahía Blanca Estuary, in Argentina, to identify risk factors for infection by the acanthocephalan Profilicollis chasmagnathi and to assess the association between infection and mortality of these hosts. Cyrtograpsus angulatus and N. granulata crabs were sampled seasonally over the course of a year, and spring sampling included collection of dead crabs predated by Olrog's gulls in a nearby breeding colony. Potential risk factors for infection were assessed and the number of cystacanth larvae per crab was counted. In C. angulatus, the odds of infection increased by 7% for each millimetre increase in carapace length, and were nearly 17 times greater in crabs sampled from the Olrog's gull feeding area compared with those sampled from nests in the breeding colony. For every millimetre increase in carapace length in N. granulata, the odds of infection increased by 13% in crabs from the breeding colony, and by 32% in crabs from the feeding area. Mean intensity of infection in N. granulata increased by 16.5% for each additional millimetre of carapace width. The level of parasite aggregation was lowest in the largest C. angulatus and highest in N. granulata predated by Olrog's gull. The results show that host size is the most important factor influencing infection prevalence in both crab species and intensity of infection in N. granulata, and suggest the presence of parasite-induced mortality in the populations studied.

  9. Receptivity of female Neohelice granulata (Brachyura, Varunidae): different strategies to maximize their reproductive success in contrasting habitats

    Science.gov (United States)

    Sal Moyano, María Paz; Luppi, Tomás; Gavio, María Andrea; Vallina, Micaela; McLay, Colin

    2012-12-01

    The extent of the receptive period may determine the mating strategies employed by female crabs to obtain mates. Here, we studied the receptivity of female Neohelice granulata (Dana, 1851) in the laboratory, including the form of the vulvae and the anatomy of the seminal receptacle (SR). We examined the factors that influence the duration of receptivity by comparing two populations inhabiting contrasting habitats: Mar Chiquita Coastal lagoon (MCL), which is an oligo-polyhaline estuary, and San Antonio Oeste (SAO), which is an eu-hyperhaline marine bay. Non-receptive females have immobile vulva opercula, while receptive females have mobile opercula. Histological sections of the SR showed that the degree of epithelium secretions was associated with the receptive stage of females, and they may be involved in the maintenance of viable sperm and in the dehiscence of spermatophores. The existence of a special tissue at the junction of the oviduct and the SR was described and proposed as an internal mechanism influencing the timing of ovulation. The duration of receptivity was dependent on the SR load and the capacity to lay eggs. Thus, females with empty SR exhibited longer receptivity and did not lay eggs, while those with full SR exhibited shorter receptivity and always laid eggs. Interpopulation differences showed that females from SAO had shorter receptivity and heavier SR and laid eggs more frequently than females from MCL. Based on our results, we suggest that N. granulata females can adjust the duration of their receptivity and control the moment of fertilization according to different internal mechanisms related to the morphology of the vulvae, the fullness of the SR and anatomical attributes of the SR. An important consequence of this control is greater sperm competition. The extent of the receptive period and the number of times that a female could become receptive in a single reproductive season may also depend on the habitat characteristics.

  10. Burrowing owl survey : 1994 report

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — Report of burrowing owl nesting activity in the Central Region of Colorado in 1994, produced by the Colorado Division of Wildlife. There is little long term data on...

  11. Depth of artificial Burrowing Owl burrows affects thermal suitability and occupancy

    Science.gov (United States)

    Nadeau, Christopher P.; Conway, Courtney J.; Rathbun, Nathan

    2015-01-01

    Many organizations have installed artificial burrows to help bolster local Burrowing Owl (Athene cunicularia) populations. However, occupancy probability and reproductive success in artificial burrows varies within and among burrow installations. We evaluated the possibility that depth below ground might explain differences in occupancy probability and reproductive success by affecting the temperature of artificial burrows. We measured burrow temperatures from March to July 2010 in 27 artificial burrows in southern California that were buried 15–76 cm below the surface (measured between the surface and the top of the burrow chamber). Burrow depth was one of several characteristics that affected burrow temperature. Burrow temperature decreased by 0.03°C per cm of soil on top of the burrow. The percentage of time that artificial burrows provided a thermal refuge from above-ground temperature decreased with burrow depth and ranged between 50% and 58% among burrows. The percentage of time that burrow temperature was optimal for incubating females also decreased with burrow depth and ranged between 27% and 100% among burrows. However, the percentage of time that burrow temperature was optimal for unattended eggs increased with burrow depth and ranged between 11% and 95% among burrows. We found no effect of burrow depth on reproductive success across 21 nesting attempts. However, occupancy probability had a non-linear relationship with burrow depth. The shallowest burrows (15 cm) had a moderate probability of being occupied (0.46), burrows between 28 and 40 cm had the highest probability of being occupied (>0.80), and burrows >53 cm had the lowest probability of being occupied (Owls may prefer burrows at moderate depths because these burrows provide a thermal refuge from above-ground temperatures, and are often cool enough to allow females to leave eggs unattended before the onset of full-time incubation, but not too cool for incubating females that spend most of

  12. Land crabs as key drivers in tropical coastal forest recruitment

    Science.gov (United States)

    Lindquist, E.S.; Krauss, K.W.; Green, P.T.; O'Dowd, D. J.; Sherman, P.M.; Smith, T. J.

    2009-01-01

    Plant populations are regulated by a diverse assortment of abiotic and biotic factors that influence seed dispersal and viability, and seedling establishment and growth at the microsite. Rarely does one animal guild exert as significant an influence on different plant assemblages as land crabs. We review three tropical coastal ecosystems-mangroves, island maritime forests, and mainland coastal terrestrial forests-where land crabs directly influence forest composition by limiting tree establishment and recruitment. Land crabs differentially prey on seeds, propagules and seedlings along nutrient, chemical and physical environmental gradients. In all of these ecosystems, but especially mangroves, abiotic gradients are well studied, strong and influence plant species distributions. However, we suggest that crab predation has primacy over many of these environmental factors by acting as the first limiting factor of tropical tree recruitment to drive the potential structural and compositional organisation of coastal forests. We show that the influence of crabs varies relative to tidal gradient, shoreline distance, canopy position, time, season, tree species and fruiting periodicity. Crabs also facilitate forest growth and development through such activities as excavation of burrows, creation of soil mounds, aeration of soils, removal of leaf litter into burrows and creation of carbon-rich soil microhabitats. For all three systems, land crabs influence the distribution, density and size-class structure of tree populations. Indeed, crabs are among the major drivers of tree recruitment in tropical coastal forest ecosystems, and their conservation should be included in management plans of these forests. ?? 2009 Cambridge Philosophical Society.

  13. The biomechanics of burrowing and boring.

    Science.gov (United States)

    Dorgan, Kelly M

    2015-01-15

    Burrowers and borers are ecosystem engineers that alter their physical environments through bioturbation, bioirrigation and bioerosion. The mechanisms of moving through solid substrata by burrowing or boring depend on the mechanical properties of the medium and the size and morphology of the organism. For burrowing animals, mud differs mechanically from sand; in mud, sediment grains are suspended in an organic matrix that fails by fracture. Macrofauna extend burrows through this elastic mud by fracture. Sand is granular and non-cohesive, enabling grains to more easily move relative to each other, and macrofaunal burrowers use fluidization or plastic rearrangement of grains. In both sand and mud, peristaltic movements apply normal forces and reduce shear. Excavation and localized grain compaction are mechanisms that plastically deform sediments and are effective in both mud and sand, with bulk excavation being used by larger organisms and localized compaction by smaller organisms. Mechanical boring of hard substrata is an extreme form of excavation in which no compaction of burrow walls occurs and grains are abraded with rigid, hard structures. Chemical boring involves secretion to dissolve or soften generally carbonate substrata. Despite substantial differences in the mechanics of the media, similar burrowing behaviors are effective in mud and sand.

  14. Localized fluidization burrowing mechanics of Ensis directus.

    Science.gov (United States)

    Winter, Amos G; Deits, Robin L H; Hosoi, A E

    2012-06-15

    Muscle measurements of Ensis directus, the Atlantic razor clam, indicate that the organism only has sufficient strength to burrow a few centimeters into the soil, yet razor clams burrow to over 70 cm. In this paper, we show that the animal uses the motions of its valves to locally fluidize the surrounding soil and reduce burrowing drag. Substrate deformations were measured using particle image velocimetry (PIV) in a novel visualization system that enabled us to see through the soil and watch E. directus burrow in situ. PIV data, supported by soil and fluid mechanics theory, show that contraction of the valves of E. directus locally fluidizes the surrounding soil. Particle and fluid mixtures can be modeled as a Newtonian fluid with an effective viscosity based on the local void fraction. Using these models, we demonstrate that E. directus is strong enough to reach full burrow depth in fluidized soil, but not in static soil. Furthermore, we show that the method of localized fluidization reduces the amount of energy required to reach burrow depth by an order of magnitude compared with penetrating static soil, and leads to a burrowing energy that scales linearly with depth rather than with depth squared.

  15. Application of morphologic burrow architects: lungfish or crayfish?

    Science.gov (United States)

    Hasiotis, Stephen T.; Mitchell, Charles E.; Dubiel, Russell R.

    1993-01-01

    A methodology for trace fossil identification using burrowing signatures is tested by evaluating ancient and modern lungfish and crayfish burrows and comparing them to previously undescribed burrows in a stratigraphic interval thought to contain both lungfish and crayfish burrows. Permian burrows that bear skeletal remains of the lungfish Gnathorhiza, from museum collections, were evaluated to identify unique burrow morphologies that could be used to distinguish lungfish from crayfish burrows when fossil remains are absent. The lungfish burrows were evaluated for details of the burrowing mechanism preserved in the burrow morphologies together forming burrowing signatures and were compared to new burrows in the Chinle Formation of western Colorado to test the methodology of using burrow signatures to identify unknown burrows. Permian lungfish aestivation burrows show simple, nearly vertical, unbranched architectures and relatively smooth surficial morphologies with characteristic quasi‐horizontal striae on the burrow walls and vertical striae on the bulbous terminus. Burrow lengths do not exceed 0.5 m. In contrast, modern and ancient crayfish burrows exhibit simple to highly complex architectures with highly textured surficial morphologies. Burrow lengths may reach 4 to 5 m. Burrow morphologies unlike those identified in Gnathorhiza aestivation burrows were found in four burrow groups from museum collections. Two of these groups exhibit simple architectures and horizontal striae that were greater in sinuosity and magnitude, respectively. One of these burrows contains the remains of Lysorophus, but the burrow surface reveals no reliable surficial characteristics. It is not clear whether Lysorophus truly burrowed or merely occupied a pre‐existing structure. The other two groups exhibit surficial morphologies similar to those found on modern and ancient crayfish burrows and may provide evidence of freshwater crayfish in the Permian. Burrows from the Upper Triassic

  16. Burrowing Owl - Palo Verde Valley [ds197

    Data.gov (United States)

    California Department of Resources — These burrowing owl observations were collected during the spring and early summer of 1976 in the Palo Verde Valley, eastern Riverside County, California. This is an...

  17. The burrowing origin of modern snakes

    Science.gov (United States)

    Yi, Hongyu; Norell, Mark A.

    2015-01-01

    Modern snakes probably originated as habitat specialists, but it controversial unclear whether they were ancestrally terrestrial burrowers or marine swimmers. We used x-ray virtual models of the inner ear to predict the habit of Dinilysia patagonica, a stem snake closely related to the origin of modern snakes. Previous work has shown that modern snakes perceive substrate vibrations via their inner ear. Our data show that D. patagonica and modern burrowing squamates share a unique spherical vestibule in the inner ear, as compared with swimmers and habitat generalists. We built predictive models for snake habit based on their vestibular shape, which estimated D. patagonica and the hypothetical ancestor of crown snakes as burrowers with high probabilities. This study provides an extensive comparative data set to test fossoriality quantitatively in stem snakes, and it shows that burrowing was predominant in the lineages leading to modern crown snakes. PMID:26702436

  18. The burrowing origin of modern snakes.

    Science.gov (United States)

    Yi, Hongyu; Norell, Mark A

    2015-11-01

    Modern snakes probably originated as habitat specialists, but it controversial unclear whether they were ancestrally terrestrial burrowers or marine swimmers. We used x-ray virtual models of the inner ear to predict the habit of Dinilysia patagonica, a stem snake closely related to the origin of modern snakes. Previous work has shown that modern snakes perceive substrate vibrations via their inner ear. Our data show that D. patagonica and modern burrowing squamates share a unique spherical vestibule in the inner ear, as compared with swimmers and habitat generalists. We built predictive models for snake habit based on their vestibular shape, which estimated D. patagonica and the hypothetical ancestor of crown snakes as burrowers with high probabilities. This study provides an extensive comparative data set to test fossoriality quantitatively in stem snakes, and it shows that burrowing was predominant in the lineages leading to modern crown snakes.

  19. Biogeochemical consequences of macrofauna burrow ventilation†

    Directory of Open Access Journals (Sweden)

    Furukawa Yoko

    2001-10-01

    Full Text Available The burrow walls created by macrofauna in aquatic sediments are sites of intense chemical mass transfer. Quantitative measurement of their significance is, however, difficult because chemistry in the immediate vicinity of burrow walls is temporally dynamic due to periodic ventilation of burrows by macrofauna. A temporally dynamic, 2D multicomponent diffusion-reaction model was utilized to depict the magnitude and time dependency of chemical mass transfer in the immediate vicinity of burrow walls as well as at the water/sediment interface. The simulation results illustrate that sediment particles, pore water, and microorganisms within a few millimeters of burrow walls experience significant oscillation in pH (as much as two pH units and dissolved oxygen concentration (between saturation and near anoxia whereas such oscillation is absent at the water/ sediment interface. The geochemical oscillation is expected to affect the net stability of mineral phases, activities and community structures of microorganisms, and rates and magnitudes of microbial diagenetic reactions.

  20. Crab Rationalization Permit Program

    Data.gov (United States)

    National Oceanic and Atmospheric Administration, Department of Commerce — The Crab Rationalization Program (Program) allocates BSAI crab resources among harvesters, processors, and coastal communities. The North Pacific Fishery Management...

  1. Crab Cavity Development

    CERN Document Server

    Calaga, R; Burt, G; Ratti, A

    2015-01-01

    The HL-LHC upgrade will use deflecting (or crab) cavities to compensate for geometric luminosity loss at low β* and non-zero crossing angle. A local scheme with crab cavity pairs across the IPs is used employing compact crab cavities at 400 MHz. Design of the cavities, the cryomodules and the RF system is well advanced. The LHC crab cavities will be validated initially with proton beam in the SPS.

  2. FLOCKING AND FEEDING IN THE FIDDLER-CRAB (UCA-TANGERI) - PREY AVAILABILITY AS RISK-TAKING BEHAVIOR

    NARCIS (Netherlands)

    ENS, BJ; KLAASSEN, M; ZWARTS, L

    1993-01-01

    For a full understanding of prey availability, it is necessary to study risk-taking behaviour of the prey. Fiddler crabs are ideally suited for such a study, as they have to leave their safe burrow to feed on the surface of the intertidal flats during low tide, thereby exposing themselves to avian p

  3. Potential effect of fiddler crabs on organic matter distribution: A combined laboratory and field experimental approach

    Science.gov (United States)

    Natálio, Luís F.; Pardo, Juan C. F.; Machado, Glauco B. O.; Fortuna, Monique D.; Gallo, Deborah G.; Costa, Tânia M.

    2017-01-01

    Bioturbators play a key role in estuarine environments by modifying the availability of soil elements, which in turn may affect other organisms. Despite the importance of bioturbators, few studies have combined both field and laboratory experiments to explore the effects of bioturbators on estuarine soils. Herein, we assessed the bioturbation potential of fiddler crabs Leptuca leptodactyla and Leptuca uruguayensis in laboratory and field experiments, respectively. We evaluated whether the presence of fiddler crabs resulted in vertical transport of sediment, thereby altering organic matter (OM) distribution. Under laboratory conditions, the burrowing activity by L. leptodactyla increased the OM content in sediment surface. In the long-term field experiment with areas of inclusion and exclusion of L. uruguayensis, we did not observe influence of this fiddler crab in the vertical distribution of OM. Based on our results, we suggest that small fiddler crabs, such as the species used in these experiments, are potentially capable of alter their environment by transporting sediment and OM but such effects may be masked by environmental drivers and spatial heterogeneity under natural conditions. This phenomenon may be related to the small size of these species, which affects how much sediment is transported, along with the way OM interacts with biogeochemical and physical processes. Therefore, the net effect of these burrowing organisms is likely to be the result of a complex interaction with other environmental factors. In this sense, we highlight the importance of performing simultaneous field and laboratory experiments in order to better understanding the role of burrowing animals as bioturbators.

  4. Evaluation of the efficiency of fixation, decalcified and corants to study of pereiopods of Neohelice granulata (Dana, 1851)

    OpenAIRE

    Borges, Elizandra Luçardo; Gonzalez, Vinícius Cunha; Jardim, Rodrigo Desessards; Loch, Dionísio; Soares, João Cassimiro Mendonça; Trindade, Gilma Santos; Varela Junior, Antonio Sergio

    2009-01-01

    Foram coletados dezesseis espécimes de Neohelice granulata no estuário da Laguna dos Patos sendo que, de cada um destes foram coletados cinco apêndices locomotores. Dois destes apêndices foram fixados em Formol a 10%, outros dois em Zenker e um em Davidson, correspondendo a 32, 32, 16 repetições respectivamente. Após os apêndices terem sido fixados em Formol e Zenker estes foram colocados nos descalcificadores Ácido Cítrico 5% e Solução de Citrato de Sódio/Ácido Fórmico. Já os apêndices fixad...

  5. FEEDING ECOLOGY OF TREE-CLIMBING MANGROVE SESARMID CRABS FROM LUZON, PHILIPPINES

    Directory of Open Access Journals (Sweden)

    JIMMY TEVAR MASAGCA

    2009-01-01

    Full Text Available Despite the large ecological study of tree-climbing mangrove sesarmid crabs in other countries, the Philippine representatives appear to have not been investigated extensively. This paper presents the feeding ecology as to dependence on mangrove trees of sesarmids in different mangrove areas of southern Luzon. This is biased on the nature of the crab habitats, arboreal climbing skills and burrowing behavior of the sesarmids: Selatium elongatum and Episesarma versicolor − exclusive mangrove tree climbers (EMTC; Sarmatium germaini − occasional mangrove tree climber (OMTC; and the non-mangrove tree-climbing (NMTC sesarmids- Neosarmatium smithii, Perisesarma bidens and Perisesarma eumolpe

  6. Burrowing Owl Monitoring Report for Calendar Year 2012

    Energy Technology Data Exchange (ETDEWEB)

    Wilde, Justin W.; Lindsey, Cole T.; Nugent, John J.

    2013-03-14

    The monitoring during 2012 focused on documenting the status of known burrows. Newly identified burrows were documented while examining historical locations, during ecological resource reviews, or discovered during other monitoring efforts. The timing of the monitoring effort allowed staff to perform the surveys without disrupting any breeding or hatching, while also allowing for easy discernment of adults from juveniles, which helped in determining burrow-use type.

  7. Fiddler crabs facilitate Spartina alterniflora growth, mitigating periwinkle overgrazing of marsh habitat.

    Science.gov (United States)

    Gittman, Rachel K; Keller, Danielle A

    2013-12-01

    Ecologists have long been interested in identifying and testing factors that drive top-down or bottom-up regulation of communities. Most studies have focused on factors that directly exert top-down (e.g., grazing) or bottom-up (e.g., nutrient availability) control on primary production. For example, recent studies in salt marshes have demonstrated that fronts of Littoraria irrorata periwinkles can overgraze Spartina alterniflora and convert marsh to mudflat. The importance of indirect, bottom-up effects, particularly facilitation, in enhancing primary production has also recently been explored. Previous field studies separately revealed that fiddler crabs, which burrow to depths of more than 30 cm, can oxygenate marsh sediments and redistribute nutrients, thereby relieving the stress of anoxia and enhancing S. alterniflora growth. However, to our knowledge, no studies to date have explored how nontrophic facilitators can mediate top-down effects (i.e., grazing) on primary-producer biomass. We conducted a field study testing whether fiddler crabs can facilitate S. alterniflora growth sufficiently to mitigate overgrazing by periwinkles and thus sustain S. alterniflora marsh. As inferred from contrasts to experimental plots lacking periwinkles and fiddler crabs, periwinkles alone exerted top-down control of total aboveground biomass and net growth of S. alterniflora. When fiddler crabs were included, they counteracted the effects of periwinkles on net S. alterniflora growth. Sediment oxygen levels were greater and S. alterniflora belowground biomass was lower where fiddler crabs were present, implying that fiddler crab burrowing enhanced S. alterniflora growth. Consequently, in the stressful interior S. alterniflora marsh, where subsurface soil anoxia is widespread, fiddler crab facilitation can mitigate top-down control by periwinkles and can limit and possibly prevent loss of biogenically structured marsh habitat and its ecosystem services.

  8. Mangrove Crab Ucides cordatus Removal Does Not Affect Sediment Parameters and Stipule Production in a One Year Experiment in Northern Brazil

    Science.gov (United States)

    2016-01-01

    Mangrove crabs influence ecosystem processes through bioturbation and/or litter feeding. In Brazilian mangroves, the abundant and commercially important crab Ucides cordatus is the main faunal modifier of microtopography establishing up to 2 m deep burrows. They process more than 70% of the leaf litter and propagule production, thus promoting microbial degradation of detritus and benefiting microbe-feeding fiddler crabs. The accelerated nutrient turn-over and increased sediment oxygenation mediated by U. cordatus may enhance mangrove tree growth. Such positive feed-back loop was tested in North Brazil through a one year crab removal experiment simulating increased harvesting rates in a mature Rhizophora mangle forest. Investigated response parameters were sediment salinity, organic matter content, CO2 efflux rates of the surface sediment, and reduction potential. We also determined stipule fall of the mangrove tree R. mangle as a proxy for tree growth. Three treatments were applied to twelve experimental plots (13 m × 13 m each): crab removal, disturbance control and control. Within one year, the number of U. cordatus burrows inside the four removal plots decreased on average to 52% of the initial number. Despite this distinct reduction in burrow density of this large bioturbator, none of the measured parameters differed between treatments. Instead, most parameters were clearly influenced by seasonal changes in precipitation. Hence, in the studied R. mangle forest, abiotic factors seem to be more important drivers of ecosystem processes than factors mediated by U. cordatus, at least within the studied timespan of one year. PMID:27907093

  9. Crab Meat with Potherb Mustard

    Institute of Scientific and Technical Information of China (English)

    1996-01-01

    Ingredients: Eight fresh crab. 75 grams of potherb mustard, two egg whites, scallions, ginger and cooking wine. Salt to the taste. Directions: 1. Slice and deep fry the potherb mustard till crisp. Place fried mustard on plate. 2. Steam the crab and remove meat. Stir fry meat with scallions, ginger, cooking wine and salt, Return meat to crab shells. 3. Beat egg white until stiff. Cover the crab meat with mixture and garnish. Steam meat for a few minutes. 4. Place the crab shells on fried mustard and serve. This attractive red and white dish features delicious crab meat with savory crisp mustard leaves.

  10. Detecting Burrowing Owl Bloodmeals in Pulex irritans (Siphonaptera: Pulicidae).

    Science.gov (United States)

    Graham, Christine B; Eisen, Rebecca J; Belthoff, James R

    2016-03-01

    Pulex irritans L. is a cosmopolitan flea species that infests a wide variety of hosts. In North America it generally parasitizes large wild mammals, but in the Pacific Northwest an association has emerged between P. irritans and the western burrowing owl (Athene cunicularia hypugaea). While investigators have recognized this association for decades, it has not been clear if P. irritans feeds on burrowing owls, or if the owls serve exclusively as phoretic hosts. Here we describe using a real-time assay that was originally developed to identify bloodmeals in Ugandan cat fleas (Ctenocephalides felis Bouché) to detect burrowing owl DNA in P. irritans collected from burrowing owls in southern Idaho. Of 50 fleas tested, 12 had no detectable vertebrate bloodmeal. The remaining 38 (76%) contained burrowing owl DNA. The assay did not detect vertebrate DNA in unfed fleas exposed to owl or mouse pelts and is therefore unlikely to detect DNA in fleas from vertebrates that have served exclusively as phoretic hosts. We conclude that P. irritans feeds on burrowing owls. We discuss the potential implications of this finding for burrowing owl conservation and enzootic plague dynamics.

  11. Incorporation of microplastics from litter into burrows of Lumbricus terrestris.

    Science.gov (United States)

    Huerta Lwanga, Esperanza; Gertsen, Hennie; Gooren, Harm; Peters, Piet; Salánki, Tamás; van der Ploeg, Martine; Besseling, Ellen; Koelmans, Albert A; Geissen, Violette

    2017-01-01

    Pollution caused by plastic debris is an urgent environmental problem. Here, we assessed the effects of microplastics in the soil surface litter on the formation and characterization of burrows built by the anecic earthworm Lumbricus terrestris in soil and quantified the amount of microplastics that was transported and deposited in L. terrestris burrows. Worms were exposed to soil surface litter treatments containing microplastics (Low Density Polyethylene) for 2 weeks at concentrations of 0%, 7%, 28%, 45% and 60%. The latter representing environmentally realistic concentrations found in hot spot soil locations. There were significantly more burrows found when soil was exposed to the surface treatment composed of 7% microplastics than in all other treatments. The highest amount of organic matter in the walls of the burrows was observed after using the treatments containing 28 and 45% microplastics. The highest microplastic bioturbation efficiency ratio (total microplastics (mg) in burrow walls/initial total surface litter microplastics (mg)) was found using the concentration of 7% microplastics, where L. terrestris introduced 73.5% of the surface microplastics into the burrow walls. The highest burrow wall microplastic content per unit weight of soil (11.8 ± 4.8 g kg-(1)) was found using a concentration of 60% microplastics. L. terrestris was responsible for size-selective downward transport when exposed to concentrations of 7, 28 and 45% microplastics in the surface litter, as the fraction ≤50 μm microplastics in burrow walls increased by 65% compared to this fraction in the original surface litter plastic. We conclude that the high biogenic incorporation rate of the small-fraction microplastics from surface litter into burrow walls causes a risk of leaching through preferential flow into groundwater bodies. Furthermore, this leaching may have implications for the subsequent availability of microplastics to terrestrial organisms or for the transport

  12. Impact of off-road vehicles (ORVs) on ghost crabs of sandy beaches with traffic restrictions: a case study of Sodwana Bay, South Africa.

    Science.gov (United States)

    Lucrezi, Serena; Saayman, Melville; van der Merwe, Peet

    2014-03-01

    Off-road vehicles (ORVs) are popular in coastal recreation, although they have negative impacts on sandy shores. In South Africa, ORVs are banned from most coastal areas, while some areas are designated for restricted ORV use, providing an opportunity to assess whether ORV traffic restrictions translate into biological returns. In Sodwana Bay, the impact of ORVs on ghost crab populations was investigated. During Easter 2012, ghost crab burrows were counted on beach sections open and closed to traffic. Burrow density in the Impact section was less than a third that of the Reference section, and by the end of the study burrow size in the Impact section was half that of the Reference section. ORV traffic caused a shift in burrow distribution to the Lower beach. However, differences in burrow densities between sections were 14 times smaller than differences obtained at a time when ORV use in Sodwana Bay was not controlled. While confirming the well-established detrimental effects of ORV use on sandy beach ecosystems, results demonstrated that traffic restrictions on beaches measurably minimize impacts to the fauna, thus translating into clear-cut biological returns.

  13. Respiratory gas transport, metabolic status, and locomotor capacity of the Christmas Island red crab Gecarcoidea natalis assessed in the field with respect to dichotomous seasonal activity levels.

    Science.gov (United States)

    Adamczewska, A M; Morris, S

    2000-05-01

    Red crabs, Gecarcoidea natalis, exhibit seasonal activity patterns: low activity during the dry season when they shelter in burrows to avoid dehydration, and high activity during the wet season. Red crabs were examined in situ in the rainforest of Christmas Island to determine if there were underlying seasonal differences in the capacity for exercise and associated metabolism. During both seasons, free-ranging (FR) crabs engaged in their normal activities and, together with crabs induced to exercise for 5 min, were sampled for haemolymph and muscle tissue. Respiratory gases in the haemolymph and key metabolites were measured to assess differences in metabolic status of FR and exercised crabs. Actively foraging FR crabs during the wet season exhibited a relative haemolymph hypoxia (2.9 kPa) and accumulated an extra 3 mmol. litre(-1) of CO(2) compared to the relatively inactive FR crabs during the dry season. Wet-season crabs appeared to be in a state of relative respiratory acidosis compared to dry-season animals. This hypercapnia may arise as a consequence of a relative hypoventilation in animals with a relatively higher metabolic rate during the wet season. Oxygenation of pulmonary and arterial haemolymph was similar and remained high after 5 min of exercise, indicating that the gills and lungs functioned similarly in gas exchange in both FR and exercised crabs. During exercise, venous O(2) reserves decreased and red crabs experienced a mixed respiratory/metabolic acidosis. Similar changes, after 5 min of enforced exercise, in metabolite concentrations, pH and respiratory gas status in the haemolymph during both sampling seasons suggest that the crabs maintain similar capacity to increase exercise during the wet and the dry seasons, despite the differences in underlying physiological status. This is important since after prolonged inactivity during the dry season, with the arrival of moonsoonal rains, red crabs must engage in their annual breeding migration.

  14. Origin of tropical American burrowing reptiles by transatlantic rafting

    OpenAIRE

    Vidal, Nicolas; Azvolinsky, Anna; Cruaud, Corinne; Hedges, S Blair

    2007-01-01

    Populations of terrestrial or freshwater taxa that are separated by oceans can be explained by either oceanic dispersal or fragmentation of a previously contiguous land mass. Amphisbaenians, the worm lizards (approx. 165 species), are small squamate reptiles that are uniquely adapted to a burrowing lifestyle and inhabit Africa, South America, Caribbean Islands, North America, Europe and the Middle East. All but a few species are limbless and they rarely leave their subterranean burrows. Given...

  15. Function of Hexagenia (Mayfly) Burrows: Fluid Model Suggests Bacterial Gardening

    Science.gov (United States)

    Traynham, B.; Furbish, D.; Miller, M.; White, D.

    2006-12-01

    Lake and stream bottoms experience an array of physical, chemical, and biological processes that create spatial variations both in the fluid column and in the sediment that provide a physical template for distinct niches. Burrowing insects are major ecological engineers of communities where they structure large areas of the benthic habitat through bioturbation and other activities including respiration, feeding, and defecation. The burrowing mayfly Hexagenia, when present in high densities, has a large impact on food-web dynamics and provides essential ecosystem services within the fluid column and benthic substrate, including sediment mixing, nutrient cycling, and ultimately, energy flow through the freshwater food web. It has long been recognized that particular benthic species are important in linking detrital energy resources to higher trophic levels and for determining how organic matter is processed in freshwater ecosystems; however, the unique contributions made by individual benthic species is largely absent from the literature. Here we present a model that describes the structure and function of a Hexagenia burrow. If testing supports this hypothesis, the model suggests that when high food concentration is available to Hexagenia, there exists a favorable tube length for harvesting bacteria that grow on the burrow walls. The burrow microhabitat created by Hexagenia serves as a case-study in understanding the influence of benthic burrowers on both energy flow through freshwater food webs and nutrient cycling.

  16. The Use of Computational Fluid Dynamics in Predicting the Tidal Flushing of Animal Burrows

    Science.gov (United States)

    Heron, S. F.; Ridd, P. V.

    2001-04-01

    Numerical hydrodynamic modelling has been used extensively over the last few decades to simulate flow in the ocean, bays and estuaries; however, modelling of much smaller scale phenomena is less common. In this work a commercially available Computational Fluid Dynamics package (FIDAP), normally used for industrial applications, was used to simulate tidally-induced flow in multi-opening animal burrows. U-shaped burrows of varying complexities were modelled to determine the effect of different surface characteristics and burrow geometries on surface water velocities, burrow velocities and burrow flushing times. The turbulent 2D model showed the slope of the surface water was proportional to the square of both the surface and burrow velocities. The effect of placing a root in the surface flow was to reduce the surface water velocity; however, the burrow flow depended upon the root position. For the root location either upstream or downstream of the burrow, the burrow velocity was reduced by 50%. With the root located between the burrow openings the burrow velocity increased by 200%, due to the increase in pressure difference across the burrow openings. A buttress root placed in the flow immediately downstream of the upstream burrow, caused the burrow flushing rate to increase significantly with increasing buttress height. Flushing times for burrows of varying depth were determined computationally by use of a tracer for the burrow water. For a burrow of depth 1·2 m, the flushing times were 5 and 28 min for root location between the burrow openings and downstream of the burrow, respectively. Animal burrows often consist of multiply-connected loops. A second burrow was added to the primary burrow and flushing times were found to be 15 and 38 min, respectively. A burrow system of four connected burrows was modelled which had corresponding flushing times up to 24 and 47 min, respectively. The calculated times are consistent with the hypothesis that a significant flushing

  17. The variable Crab Nebula

    CERN Document Server

    Tavani, Marco

    2011-01-01

    The remarkable Crab Nebula is powered by an energetic pulsar whose relativistic wind interacts with the inner parts of the Supernova Remnant SN1054. Despite low-intensity optical and X-ray variations in the inner Nebula, the Crab has been considered until now substantially stable at X-ray and gamma-ray energies. This paradigm has been shattered by the AGILE discovery in September 2010 of a very intense transient gamma-ray flare of nebular origin. For the first time, the Crab Nebula was "caught in the act" of accelerating particles up to 10^15 eV within the shortest timescale ever observed in a cosmic nebula (1 day or less). Emission between 50 MeV and a few GeV was detected with a quite hard spectrum within a short timescale. Additional analysis and recent Crab Nebula data lead to identify a total of four major flaring gamma-ray episodes detected by AGILE and Fermi during the period mid-2007/mid-2011. These observations challenge emission models of the pulsar wind interaction and particle acceleration process...

  18. Burrowing criteria and burrowing mode adjustment in bivalves to varying geoenvironmental conditions in intertidal flats and beaches.

    Directory of Open Access Journals (Sweden)

    Shinji Sassa

    Full Text Available The response of bivalves to their abiotic environment has been widely studied in relation to hydroenvironmental conditions, sediment types and sediment grain sizes. However, the possible role of varying geoenvironmental conditions in their habitats remains poorly understood. Here, we show that the hardness of the surficial intertidal sediments varies by a factor of 20-50 due to suction development and suction-induced void state changes in the essentially saturated states of intertidal flats and beaches. We investigated the response of two species of bivalves, Ruditapes philippinarum and Donax semigranosus, in the laboratory by simulating such prevailing geoenvironmental conditions in the field. The experimental results demonstrate that the bivalve responses depended strongly on the varying geoenvironmental conditions. Notably, both bivalves consistently shifted their burrowing modes, reducing the burrowing angle and burial depth, in response to increasing hardness, to compensate for the excessive energy required for burrowing, as explained by a proposed conceptual model. This burrowing mode adjustment was accompanied by two burrowing criteria below or above which the bivalves accomplished vertical burrowing or failed to burrow, respectively. The suitable and fatal conditions differed markedly with species and shell lengths. The acute sensitivities of the observed bivalve responses to geoenvironmental changes revealed two distinctive mechanisms accounting for the adult-juvenile spatial distributions of Ruditapes philippinarum and the behavioral adaptation to a rapidly changing geoenvironment of Donax semigranosus. The present results may provide a rational basis by which to understand the ensuing, and to predict future, bivalve responses to geoenvironmental changes in intertidal zones.

  19. Use of Artificial Burrows by Burrowing Owls (Athene cunicularia) at the HAMMER Facility on the U.S. Department of Energy Hanford Site

    Energy Technology Data Exchange (ETDEWEB)

    Alexander, Amanda K.; Sackschewsky, Michael R.; Duberstein, Corey A.

    2005-09-30

    In 2003 the U.S. Department of Energy (DOE) constructed an Emergency Vehicle Operations Course (EVOC) at the Hazardous Material Management and Emergency Response Training and Education Center (HAMMER) in the southern portion of the Hanford Site. Preliminary surveys during 2001 identified an active burrowing owl (Athene cunicularia) burrow and three burrowing owls within the proposed development area. Burrowing owls were classified as a federal species of concern, a Washington State ?candidate? species, a Washington Department of Fish and Wildlife priority species, and a Hanford Site Biological Resources Management Plan Level III resource. Therefore, the mitigation action plan for the project included the installation of twenty artificial burrows around EVOC in the spring of 2003. The mitigation plan established a success criterion of five percent annual use of the burrows by owls. In July 2005, a field survey of the EVOC burrow complex was conducted to determine use and demography at each site. Burrow locations were mapped and signs of activity (feces, owl tracks, castings, feathers) were recorded. Out of the twenty burrows, twelve were found to be active. Of the eight inactive burrows three appeared to have been active earlier in the 2005 breeding season. A total of nineteen owls were counted but demography could not be determined. It appears that the EVOC mitigation exceeded burrow use goals during 2005. Continued site monitoring and maintenance, according to mitigation plan guidelines should be conducted as prescribed.

  20. Plant cover effect on Bolson tortoise (Gopherus flavomarginatus Legler 1959, Testudinidae burrow use

    Directory of Open Access Journals (Sweden)

    Jorge Luis Becerra-López

    2017-03-01

    Full Text Available The Bolson tortoise, Gopherus flavomarginatus, occurs within a restricted geographical area in the Mexican Chihuahuan Desert. We analyzed the variation in surface microhabitat with relation to the burrow occupancy for this tortoise at the Mapimí Biosphere Reserve, Mexico. In summer 2010, we monitored burrow activity (active, inactive, or abandoned and measured environmental factors that might influence the burrow’s occupancy by tortoises (air temperature, relative humidity and substrate temperature, both inside and outside the burrow, and the plant cover around it. Discriminant analysis was used to identify the importance of these variables influencing burrow occupancy. Correlation and linear regression analyses were performed to quantify the relation between environmental factors in the sampled burrows. Results. Sixty-one burrows were identified at the Tortugas locality. The first function’s auto-value analysis indicates that this function explains 97.9% of the variation in burrow activity status; high occupancy scores were associated with low substrate temperature inside the burrow. Plant cover was inversely proportional to substrate temperature inside the burrow. These results suggest the importance the density of plants surrounding the tortoise’s burrow as a key factor influencing the burrow microclimate and occupancy by the tortoises. Conclusions. Gopherus flavomarginatus inhabits burrows, in part, based on microhabitat structure, with plant cover being a main factor influencing burrow occupancy. Our findings indicate that human land use and vegetation management are important for conserving Bolson tortoises, and for understanding habitat conditions necessary for the successful establishment of populations elsewhere.

  1. Mechanics and kinematics of backward burrowing by the polychaete Cirriformia moorei.

    Science.gov (United States)

    Che, James; Dorgan, Kelly M

    2010-12-15

    The polychaete Cirriformia moorei burrows in muddy sediments by fracture, using its hydrostatic skeleton to expand its anterior region and exert force against its burrow wall to extend a crack. Burrowing occurs in four phases: stretching forward into the burrow, extending the crack anteriorly, thickening the burrowing end to amplify stress at the tip of the crack, and bringing the rest of the body forward as a peristaltic wave travels posteriorly. Here, we show that C. moorei is also able to burrow with its posterior end using a similar mechanism of crack propagation and exhibiting the same four phases of burrowing. Worms burrowed backwards with similar speeds and stress intensity factors as forward burrowing, but were thinner and less blunt and did not slip as far away from the crack tip between cycles of burrowing. The anterior end is more muscular and rigid, and differences in body shapes are consistent with having reduce musculature to dilate the posterior segments while burrowing. Backward burrowing provides a unique opportunity to study the effects of morphology on burrowing mechanics within the same species under identical conditions.

  2. Artificial and Modified Burrows Establishment for the Atlantic Puffin on Petit Manan Island, Maine

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — An attempt to establish a breeding population of Atlantic puffin colony using artificial and modified burrows on Petit Manan Island resulted in 3 of 17 burrows being...

  3. 50 CFR Table 9 to Part 680 - Initial Issuance of Crab PQS by Crab QS Fishery

    Science.gov (United States)

    2010-10-01

    ... 50 Wildlife and Fisheries 9 2010-10-01 2010-10-01 false Initial Issuance of Crab PQS by Crab QS... EXCLUSIVE ECONOMIC ZONE OFF ALASKA Pt. 680, Table 9 Table 9 to Part 680—Initial Issuance of Crab PQS by Crab QS Fishery Column A:For each crab QS fishery: Column B:The Regional Administrator shall calculate...

  4. 50 CFR Table 7 to Part 680 - Initial Issuance of Crab QS by Crab QS Fishery

    Science.gov (United States)

    2010-10-01

    ... 50 Wildlife and Fisheries 9 2010-10-01 2010-10-01 false Initial Issuance of Crab QS by Crab QS... EXCLUSIVE ECONOMIC ZONE OFF ALASKA Pt. 680, Table 7 Table 7 to Part 680—Initial Issuance of Crab QS by Crab QS Fishery Column A: Crab QS Fisheries Column B: Qualifying Years for QS Column C: Eligibility...

  5. Hairy Crab Arrives at Zen

    Institute of Scientific and Technical Information of China (English)

    2011-01-01

    The best time for feasting on hairy crab is only for a short period of time in autumn,so don’t miss out on this unparalleled delicacy. Zen has prepared a variety of dishes made from the crab. Gift boxes and gift vouchers are also available for your friends and family.

  6. Crab Cavities for Linear Colliders

    Energy Technology Data Exchange (ETDEWEB)

    Burt, G.; Ambattu, P.; Carter, R.; Dexter, A.; Tahir, I.; /Cockcroft Inst. Accel. Sci. Tech. /Lancaster U.; Beard, C.; Dykes, M.; Goudket, P.; Kalinin, A.; Ma, L.; McIntosh, P.; /Daresbury; Shulte, D.; /CERN; Jones, Roger M.; /Cockcroft Inst. Accel. Sci. Tech. /Manchester U.; Bellantoni, L.; Chase, B.; Church, M.; Khabouline, T.; Latina, A.; /Fermilab; Adolphsen, C.; Li, Z.; Seryi, Andrei; /SLAC

    2011-11-08

    Crab cavities have been proposed for a wide number of accelerators and interest in crab cavities has recently increased after the successful operation of a pair of crab cavities in KEK-B. In particular crab cavities are required for both the ILC and CLIC linear colliders for bunch alignment. Consideration of bunch structure and size constraints favour a 3.9 GHz superconducting, multi-cell cavity as the solution for ILC, whilst bunch structure and beam-loading considerations suggest an X-band copper travelling wave structure for CLIC. These two cavity solutions are very different in design but share complex design issues. Phase stabilisation, beam loading, wakefields and mode damping are fundamental issues for these crab cavities. Requirements and potential design solutions will be discussed for both colliders.

  7. Crab cavities for linear colliders

    CERN Document Server

    Burt, G; Carter, R; Dexter, A; Tahir, I; Beard, C; Dykes, M; Goudket, P; Kalinin, A; Ma, L; McIntosh, P; Shulte, D; Jones, Roger M; Bellantoni, L; Chase, B; Church, M; Khabouline, T; Latina, A; Adolphsen, C; Li, Z; Seryi, Andrei; Xiao, L

    2008-01-01

    Crab cavities have been proposed for a wide number of accelerators and interest in crab cavities has recently increased after the successful operation of a pair of crab cavities in KEK-B. In particular crab cavities are required for both the ILC and CLIC linear colliders for bunch alignment. Consideration of bunch structure and size constraints favour a 3.9 GHz superconducting, multi-cell cavity as the solution for ILC, whilst bunch structure and beam-loading considerations suggest an X-band copper travelling wave structure for CLIC. These two cavity solutions are very different in design but share complex design issues. Phase stabilisation, beam loading, wakefields and mode damping are fundamental issues for these crab cavities. Requirements and potential design solutions will be discussed for both colliders.

  8. Locomotion, respiratory physiology, and energetics of amphibious and terrestrial crabs.

    Science.gov (United States)

    Adamczewska, A M; Morris, S

    2000-01-01

    metabolism of G. natalis are strongly implied, including variations in hyperglycaemic hormone, variable basal metabolic rates, and a diel alkalosis present only in migrating crabs. The persistent dependence on water for reproduction is a determining factor in the biology of air-breathing crabs. The annual migrations include costs other than locomotion, for example, burrow construction and intermale competition. Estimates of costs that consider walking alone will underestimate the metabolic and stored fuel requirements for successful reproduction.

  9. Oxygen penetration around burrows and roots in aquatic sediments

    DEFF Research Database (Denmark)

    Meysman, Filip J.R.; Galaktionov, O.S.; Glud, Ronnie N.

    2010-01-01

    Diffusion is the dominant physical mechanism for the transfer of oxygen into fine-grained aquatic sediments. This diffusive uptake occurs at the sediment-water interface, but also at internal interfaces, such as along ventilated burrows or O2 releasing plant roots. Here, we present a systematic...

  10. Incorporation of microplastics from litter into burrows of Lumbricus terrestris

    NARCIS (Netherlands)

    Huerta Lwanga, Esperanza; Gertsen, H.F.; Gooren, H.; Peters, P.; Salanki, T.E.; Ploeg, van der M.; Besseling, E.; Koelmans, A.A.; Geissen, V.

    2017-01-01

    Pollution caused by plastic debris is an urgent environmental problem. Here, we assessed the effects of microplastics in the soil surface litter on the formation and characterization of burrows built by the anecic earthworm Lumbricus terrestris in soil and quantified the amount of microplastics that

  11. Factors affecting detection of burrowing owl nests during standardized surveys

    Science.gov (United States)

    Conway, C.J.; Garcia, V.; Smith, M.D.; Hughes, K.

    2008-01-01

    Identifying causes of declines and evaluating effects of management practices on persistence of local populations of burrowing owls (Athene cunicularia) requires accurate estimates of abundance and population trends. Moreover, regulatory agencies in the United States and Canada typically require surveys to detect nest burrows prior to approving developments or other activities in areas that are potentially suitable for nesting burrowing owls. In general, guidelines on timing of surveys have been lacking and surveys have been conducted at different times of day and in different stages of the nesting cycle. We used logistic regression to evaluate 7 factors that could potentially affect probability of a surveyor detecting a burrowing owl nest. We conducted 1,444 detection trials at 323 burrowing owl nests within 3 study areas in Washington and Wyoming, USA, between February and August 2000-2002. Detection probability was highest during the nestling period and increased with ambient temperature. The other 5 factors that we examined (i.e., study area, time of day, timing within the breeding season, wind speed, % cloud cover) interacted with another factor to influence detection probability. Use of call-broadcast surveys increased detection probability, even during daylight hours when we detected >95% of owls visually. Optimal timing of surveys will vary due to differences in breeding phenology and differences in nesting behavior across populations. Nevertheless, we recommend ???3 surveys per year: one that coincides with the laying and incubation period, another that coincides with the early nestling period, and a third that coincides with the late nestling period. In northern latitudes, surveys can be conducted throughout the day.

  12. No sign of decreased burrowing behavior in the genetically depressive flinders rats

    DEFF Research Database (Denmark)

    Baastrup, C. S.; Wegener, Gregers; Finnerup, N. B.

    2012-01-01

    Background: Burrowing is a natural behavior in rats and has been observed in several different strains. Furthermore decreased burrowing behavior has been shown in different rodent disease models like Multiple sclerosis, peripheral nerve injury and knee inflammation when compared with control...... of each drug. Results: The genetically depressive FSL rats did not burrow less than FRL control rats. Treatment with imipramin or citalopram-S did not change the burrowing behavior of the FSL rats. In contrast treatment with imipramin and citalopram-S unexpectedly decreased the burrowing behavior...

  13. Seasonal changes in burrow geometry of the common mole rat (Rodentia: Bathyergidae)

    Science.gov (United States)

    Thomas, H. G.; Scantlebury, M.; Swanepoel, D.; Bateman, P. W.; Bennett, N. C.

    2013-11-01

    Sociality in mole rats has been suggested to have evolved as a response to the widely dispersed food resources and the limited burrowing opportunities that result from sporadic rainfall events. In the most arid regions, individual foraging efficiency is reduced, and energetic constraints increase. In this study, we investigate seasonal differences in burrow architecture of the social Cryptomys hottentotus hottentotus in a mesic region. We describe burrow geometry in response to seasonal weather conditions for two seasons (wet and dry). Interactions occurred between seasons and colony size for the size of the burrow systems, but not the shape of the burrow systems. The fractal dimension values of the burrow systems did not differ between seasons. Thus, the burrow complexity was dependent upon the number of mole rats present in the social group.

  14. Golden Crab Logbook Survey (Vessels)

    Data.gov (United States)

    National Oceanic and Atmospheric Administration, Department of Commerce — In November 1995, a voluntary logbook program for the golden crab fishery in the waters under the jurisdiction of the South Atlantic Fishery Management Council...

  15. On the Crab Proper Motion

    CERN Document Server

    Caraveo, P A; Caraveo, Patrizia A; Mignani, Roberto

    1998-01-01

    Owing to the dramatic evolution of telescopes as well as optical detectors in the last 20 yrs, we are now able to measure anew the proper motion of the Crab pulsar, after the classical result of Wyckoff and Murray (1977) in a time span 40 times shorter. The proper motion is aligned with the axis of symmetry of the inner Crab nebula and, presumably, with the pulsar spin axis.

  16. Antarctic crabs: invasion or endurance?

    Science.gov (United States)

    Griffiths, Huw J; Whittle, Rowan J; Roberts, Stephen J; Belchier, Mark; Linse, Katrin

    2013-01-01

    Recent scientific interest following the "discovery" of lithodid crabs around Antarctica has centred on a hypothesis that these crabs might be poised to invade the Antarctic shelf if the recent warming trend continues, potentially decimating its native fauna. This "invasion hypothesis" suggests that decapod crabs were driven out of Antarctica 40-15 million years ago and are only now returning as "warm" enough habitats become available. The hypothesis is based on a geographically and spatially poor fossil record of a different group of crabs (Brachyura), and examination of relatively few Recent lithodid samples from the Antarctic slope. In this paper, we examine the existing lithodid fossil record and present the distribution and biogeographic patterns derived from over 16,000 records of Recent Southern Hemisphere crabs and lobsters. Globally, the lithodid fossil record consists of only two known specimens, neither of which comes from the Antarctic. Recent records show that 22 species of crabs and lobsters have been reported from the Southern Ocean, with 12 species found south of 60 °S. All are restricted to waters warmer than 0 °C, with their Antarctic distribution limited to the areas of seafloor dominated by Circumpolar Deep Water (CDW). Currently, CDW extends further and shallower onto the West Antarctic shelf than the known distribution ranges of most lithodid species examined. Geological evidence suggests that West Antarctic shelf could have been available for colonisation during the last 9,000 years. Distribution patterns, species richness, and levels of endemism all suggest that, rather than becoming extinct and recently re-invading from outside Antarctica, the lithodid crabs have likely persisted, and even radiated, on or near to Antarctic slope. We conclude there is no evidence for a modern-day "crab invasion". We recommend a repeated targeted lithodid sampling program along the West Antarctic shelf to fully test the validity of the "invasion hypothesis".

  17. Does petroleum development affect burrowing owl nocturnal space-use?

    Energy Technology Data Exchange (ETDEWEB)

    Scobie, Corey; Wellicome, Troy; Bayne, Erin [Department of Biological Sciences, University of Alberta (Canada)], email: cscobie@ualberta.ca, email: tiw@ualberta.ca, email: bayne@ualberta.ca

    2011-07-01

    Decline all over Canada in the population of burrowing owls, a federally listed endangered species, has raised concerns about the possible influence of petroleum infrastructure development on owl nocturnal space-use while foraging. Roads, wells, pipelines and sound-producing facilities related to petroleum development change the landscape and can influence the owls' mortality risk. For 3 years, 27 breeding adult male burrowing owls with nests close to different petroleum infrastructures were captured and fitted with a miniature GPS datalogger in order to track their nocturnal foraging. Data from these GPS devices were fed into a geographical information system and showed that pipelines and wells did not alter the foraging habits of the owls. Dirt and gravel roads, with little traffic, were preferentially selected by the owls, conceivably because of higher owl mortality risk along paved roads. Sound-producing facilities did not change owls' foraging behaviour, implying that sound may not affect their nocturnal space-use. Traffic data and sound power measurements will be used in further studies in an effort to better understand burrowing owls' nocturnal foraging habits.

  18. Diet of western Burrowing Owls wintering in southern Texas

    Science.gov (United States)

    Littles, C.J.; Williford, D.; Skoruppa, M.K.; Woodin, M.C.; Hickman, G.C.

    2007-01-01

    Winter diets of the western Burrowing Owl (Athene cunicularia hypugaea) are little known. We determined the diet of western Burrowing Owls wintering in southern Texas by analyzing the contents of 182 pellets collected over four winters (1999-2000, 2001-2002, 2002-2003, and 2003-2004) in three habitat types (agricultural, mainland grassland, and barrier island). Remains of a total of 7476 prey items were recovered, 98% of which were arthropods. Gryllidae (crickets) formed the largest component (50%) of the prey, followed by lepidopteran larvae (13%), beetles (8%), spiders (7%), and earwigs (6%). Although vertebrates, primarily small mammals and birds, represented only 2% of prey items by number, they represented most (71%) of the biomass. Northern pygmy mice (Baiomys taylori) and fulvous harvest mice (Reithrodontomys fulveccens) were the two most frequently consumed vertebrate species. In all habitats, arthropods, especially orthopterans, were the primary prey item by number, whereas vertebrates, primarily small mammals, were the most important by biomass. Greater consumption of arthropods by Burrowing Owls in agricultural areas may be a factor contributing to owl use of these highly altered environments. ?? 2007 The Raptor Research Foundation, Inc.

  19. Origin of tropical American burrowing reptiles by transatlantic rafting.

    Science.gov (United States)

    Vidal, Nicolas; Azvolinsky, Anna; Cruaud, Corinne; Hedges, S Blair

    2008-02-23

    Populations of terrestrial or freshwater taxa that are separated by oceans can be explained by either oceanic dispersal or fragmentation of a previously contiguous land mass. Amphisbaenians, the worm lizards (approx. 165 species), are small squamate reptiles that are uniquely adapted to a burrowing lifestyle and inhabit Africa, South America, Caribbean Islands, North America, Europe and the Middle East. All but a few species are limbless and they rarely leave their subterranean burrows. Given their peculiar habits, the distribution of amphisbaenians has been assumed to be primarily the result of two land-mass fragmentation events: the split of the supercontinent Pangaea starting 200 Myr ago, separating species on the northern land mass (Laurasia) from those on the southern land mass (Gondwana), and the split of South America from Africa 100 Myr ago. Here we show with molecular evidence that oceanic dispersal-on floating islands-played a more prominent role, and that amphisbaenians crossed the Atlantic Ocean in the Eocene (40 Myr ago) resulting in a tropical American radiation representing one-half of all known amphisbaenian species. Until now, only four or five transatlantic dispersal events were known in terrestrial vertebrates. Significantly, this is the first such dispersal event to involve a group that burrows, an unexpected lifestyle for an oceanic disperser.

  20. Rodent burrows in late Pleistocene paleosols at Korean Palaeolithic sites and their implications for paleoclimate changes

    Science.gov (United States)

    Lim, H.; Park, S.; Lee, J.; Lee, Y.

    2013-12-01

    Rodent burrows are commonly found at many Paleolithic archaeological sites in Korea. They are nearly straight in horizontal view and gently inclined in lateral view. Burrow diameters are mostly 7 - 10cm, and burrow length may reach a few meters. Vertical penetration depths are generally about 1 m from the surface, and the thickness of the burrow-bearing layer is about 1-2 m. Although no remains (bones, teeth, claws, and coprolites) were found within burrows, they are interpreted to have been produced by rodent-like mammals (probably ground squirrels) based on the size and architecture. According to the previous study, the age of these burrows was constrained to be between ca. 40,000 and 25,000 yr BP by tephrochronology, radiocarbon and optically stimulated luminescence dating results (Lim et al., 2007). However, little is known about the reason why these burrows have disappeared after late Pleistocene time. For this question, two explanations can be considered: extinction or migration. Since same kinds of burrows are still found in the high-latitude regions, such as Mongolia and North America, the possibility of extinction can be ruled out. Therefore, migration seems to be the most likely explanation. Our results show that the destruction of habitat caused by climate change during this period is the main reason for the northward migration of burrowing animals. This study suggests that rodent burrows found in the late Pleistocene paleosols can provide useful information on paleoclimate and paleoenvironmental changes.

  1. Nest site characteristics and nesting success of the Western Burrowing Owl in the eastern Mojave Desert

    Science.gov (United States)

    Longshore, Kathleen M.; Crowe, Dorothy E.

    2013-01-01

    We evaluated nest site selection at two spatial scales (microsite, territory) and reproductive success of Western Burrowing Owls (Athene cunicularia hypugaea) at three spatial scales (microsite, territory, landscape) in the eastern Mojave Desert. We used binary logistic regression within an information-theoretic approach to assess factors influencing nest site choice and nesting success. Microsite-scale variables favored by owls included burrows excavated by desert tortoise (Gopherus agassizii), burrows with a large mound of excavated soil at the entrance, and a greater number of satellite burrows within 5 m of the nest burrow. At the territory scale, owls preferred patches with greater cover of creosote bush (Larrea tridentata) within 50 m of the nest burrow. An interaction between the presence or absence of a calcic soil horizon layer over the top of the burrow (microsite) and the number of burrows within 50 m (territory) influenced nest site choice. Nesting success was influenced by a greater number of burrows within 5 m of the nest burrow. Total cool season precipitation was a predictor of nesting success at the landscape scale. Conservation strategies can rely on management of habitat for favored and productive nesting sites for this declining species.

  2. Determination of the burrow shapes of Cardisoma guanhumi on Vieques, Puerto Rico

    Directory of Open Access Journals (Sweden)

    Shannon Sample

    2016-02-01

    Full Text Available Objective: To determine the burrow morphology of Cardisoma guanhumi and to determine if a battery-powered fiber optic camera could be used to investigate these burrows. Methods: A portable fiber optic camera was used to investigate 116 active burrows. All burrows were categorized according to shape. The diameter of each burrow was also recorded and biomass calculations were completed at each study site. Results: Analysis showed that different study sites were more likely to have particular burrow shapes than expected. Three main types of burrows were classified as horizontal slide, short and shallow, and inverted-S. A Pearson Chi-square analysis revealed that burrow shape was not equally distributed across study sites (χ2 = 61.05, df = 18, P < 0.000 1. Biomass calculations showed that different sized animals inhabited different study sites (ANOVA; df = 7, MS = 158.3, F = 13.9, P < 0.000 1. The fiber optic camera was useful in determining burrow morphology and occupancy non-destructively. Conclusions: As the locations of juvenile individuals of this species are poorly known, and the species is both ecologically and economically important where it occurs, a greater understanding of burrow morphology and size segregation may help agencies be responsible for managing this natural resource to do so effectively.

  3. Burrow casts from the Lystrosaurus-Procolophon Assemblage-zone, Karoo Sequence, South Africa

    Directory of Open Access Journals (Sweden)

    G.H. Groenewald

    1991-09-01

    Full Text Available Five types of burrow casts from the Lystrosaurus- Procolophon Assemblage-zone (Palingkloof Member and Katberg Formation, Triassic, Karoo sequence. South Africa are associated with casts of desiccation cracks and red mudstone. Vertebrate remains of Lystrosaurus sp. and Procolophon sp. indicate that these animals probably made the burrows during the Triassic. It is possible that burrowing was an adaptive advantage during periods of severe and unfavourable climatic conditions. Similar burrow casts were found in the Dicynodon-Theriognathus Assemblage-zone, suggesting a burrowing habit for fauna represented in this zone. In structure, the burrow casts resemble those of Scoyenia, Thalassinoides, Histioderma, Gyrolithes and Planolites reported from Germany, France, Asia, Ireland, Spain and the United States of America.

  4. Water pumping and analysis of flow in burrowing zoobenthos - a short overview

    DEFF Research Database (Denmark)

    Riisgård, Hans Ulrik; Larsen, Poul Scheel

    2005-01-01

    Burrowing animals maintain contact with the water above the sediment by pumping water through a tube system and therefore measurements of water pumping rate of burrowing animals is of crucial importance for the study of many processes both within and above the sea floor. This review deals...... with the measuring of water pumping and the analysis of flow generated by burrowing deposit- and filter-feeding zoobenthos in order to determine the type of pump and mechanisms involved, flow rate, pump pressure, and pumping power. The practical use of fluid mechanical principles is examined, and it is stressed...... that not only the pump pressure that a burrowing animal can apply is of interest for assessing the energy cost of pumping, but also the distribution of excess pressure along its burrow is of importance for assessing the seepage flow of oxygen-rich water into the sediment surrounding the burrow because...

  5. First laboratory insight on the behavioral rhythms of the bathyal crab Geryon longipes

    Science.gov (United States)

    Nuñez, J. D.; Sbragaglia, V.; García, J. A.; Company, J. B.; Aguzzi, J.

    2016-10-01

    The deep sea is the largest and at the same time least explored biome on Earth, but quantitative studies on the behavior of bathyal organisms are scarce because of the intrinsic difficulties related to in situ observations and maintaining animals in aquaria. In this study, we reported, for the first time, laboratory observations on locomotor rhythms and other behavioral observations (i.e. feeding, exploring and self-grooming) for the bathyal crab Geryon longipes. Crabs were collected on the middle-lower slope (720-1750 m) off the coast of Blanes (Spain). Inertial (18 h) water currents and monochromatic blue (i.e. 470 nm) light-darkness (24 h) cycles were simulated in two different experiments in flume tanks endowed with burrows. Both cycles were simulated in order to investigate activity rhythms regulation in Mediterranean deep-sea benthos. Crabs showed rhythmic locomotor activity synchronized to both water currents and light-darkness cycles. In general terms, feeding and exploring behaviors also followed the same pattern. Results presented here indicate the importance of local inertial (18 h) periodicity of water currents at the seabed as a temporal cue regulating the behavior of bathyal benthic fauna in all continental margin areas where the effects of tides is negligible.

  6. Burrowing by Sailfin Catfish (Pterygoplichthys sp.): A Potential Cause of Erosion in Disturbed Environments

    Science.gov (United States)

    2014-03-01

    great variation in burrow densities, both within habitats and among different habitats. Burrow density is highly correlated to the amount of silt in...which may have constrained variation in fish abundance. Lastly, sailfin catfish may be predisposed to burrow in areas already prone to moderate...Size structure, reproductive phenology , and sex ratio of an exotic armored catfish (liposarcus multiradiatus) in the Kaoping River of southern

  7. Determinants of between-year burrow re-occupation in a colony of the European bee-eater Merops apiaster.

    Science.gov (United States)

    Brust, Vera; Bastian, Hans-Valentin; Bastian, Anita; Schmoll, Tim

    2015-08-01

    Re-occupation of existing nesting burrows in the European bee-eater Merops apiaster has only rarely - and if so mostly anecdotically - been documented in the literature record, although such behavior would substantially save time and energy. In this study, we quantify burrow re-occupation in a German colony over a period of eleven years and identify ecological variables determining reuse probability. Of 179 recorded broods, 54% took place in a reused burrow and the overall probability that one of 75 individually recognized burrows would be reused in a given subsequent year was estimated as 26.4%. This indicates that between-year burrow reuse is a common behavior in the study colony which contrasts with findings from studies in other colonies. Furthermore, burrow re-occupation probability declined highly significantly with increasing age of the breeding wall. Statistical separation of within- and between-burrow effects of the age of the breeding wall revealed that a decline in re-occupation probability with individual burrow age was responsible for this and not a selective disappearance of burrows with high re-occupation probability over time. Limited duty cycles of individual burrows may be caused by accumulating detritus or decreasing stability with increasing burrow age. Alternatively, burrow fidelity may presuppose pair fidelity which may also explain the observed restricted burrow reuse duty cycles. A consequent next step would be to extend our within-colony approach to other colonies and compare the ecological circumstances under which bee-eaters reuse breeding burrows.

  8. Caught in the Crab's claws

    CERN Multimedia

    2001-01-01

    'The crab', a new cryo magnet transport vehicle, starts work at CERN. Produced by the ESI group of EST division and built in Finland, it has the job of transporting LHC magnets in buildings SM18 and SMA18. If you see a huge crab scuttling around building SMA18 don't be afraid! It is the new Cryo Magnet Transport Vehicle produced by the ESI group (Engineering Support for Infrastructure, EST Division) for CERN's LHC project and built by Finnish Company ROCLA. This orange vehicle, nicknamed 'The Crab', is perhaps the strangest piece of equipment used for the construction of LHC magnets. It will start work at the end of this month. The crab will be used to transport LHC cryo-magnets and their components in the assembly and preparation building, SMA18, and test building, SM18. It has many capabilities that will allow CERN staff and contractors transport magnets between the two buildings and to locate them in the right position on the test beds. The crab in action during its first tests on 8 February. How does th...

  9. Determination of the burrow shapes of Cardisoma guanhumi on Vieques, Puerto Rico

    Institute of Scientific and Technical Information of China (English)

    Shannon Sample; MarcAlbrecht

    2016-01-01

    Objective: To determine the burrow morphology of Cardisoma guanhumi and to determine if a battery-powered fiber optic camera could be used to investigate these burrows. Methods: A portable fiber optic camera was used to investigate 116 active burrows. All burrows were categorized according to shape. The diameter of each burrow was also recorded and biomass calculations were completed at each study site. Results: Analysis showed that different study sites were more likely to have particular burrow shapes than expected. Three main types of burrows were classified as horizontal slide, short and shallow, and inverted-S. A Pearson Chi-square analysis revealed that burrow shape was not equally distributed across study sites (χ2 = 61.05, df = 18, P Conclusions: As the locations of juvenile individuals of this species are poorly known, and the species is both ecologically and economically important where it occurs, a greater understanding of burrow morphology and size segregation may help agencies be responsible for managing this natural resource to do so effectively.

  10. Reproduction and management of the mangrove crab Ucides cordatus (Crustacea, Brachyura, Ucididae at Iguape, São Paulo, Brazil

    Directory of Open Access Journals (Sweden)

    BRUNO S. SANT'ANNA

    2014-09-01

    Full Text Available The mangrove crab Ucides cordatus is one of the most exploited crustaceans in Brazil. The present study investigated the breeding season of this species and the period of the “andada” phenomenon, when the crabs are active outside their burrows and perform agonistic behaviors. Furthermore related them to environmental factors, in a mangrove on the coast of Brazil, with inferences about management of this crab. The crabs reproduced from October through February. The “andada” occurred from November through February, with typical characteristics in the first two months, during the full and new moon periods, and was influenced positively by the air and soil temperature, luminosity, and tidal amplitude, and negatively by salinity. Based on the results of the present study, the period when the “andada” occurs in other parts of Brazil could be defined, which will aid in managing this fishery. For Iguape, state of São Paulo, Brazil, we suggest a legal off-season for both sexes in December or during the full and new moon, including January.

  11. Are Crab nanoshots Schwinger sparks?

    Energy Technology Data Exchange (ETDEWEB)

    Stebbins, Albert [Fermi National Accelerator Lab. (FNAL), Batavia, IL (United States); Yoo, Hojin [Univ. of Wisconsin, Madison, WI (United States); Fermi National Accelerator Lab. (FNAL), Batavia, IL (United States); Lawrence Berkeley National Lab. (LBNL), Berkeley, CA (United States); Univ. of California, Berkeley, CA (United States)

    2015-05-21

    The highest brightness temperature ever observed are from "nanoshots" from the Crab pulsar which we argue could be the signature of bursts of vacuum e± pair production. If so this would be the first time the astronomical Schwinger effect has been observed. These "Schwinger sparks" would be an intermittent but extremely powerful, ~103 L, 10 PeV e± accelerator in the heart of the Crab. These nanosecond duration sparks are generated in a volume less than 1 m3 and the existence of such sparks has implications for the small scale structure of the magnetic field of young pulsars such as the Crab. As a result, this mechanism may also play a role in producing other enigmatic bright short radio transients such as fast radio bursts.

  12. Are Crab Nanoshots Schwinger Sparks?

    CERN Document Server

    Stebbins, Albert

    2015-01-01

    The highest brightness temperature ever observed are from "nanoshots" from the Crab pulsar which we argue could be the signature of bursts of vacuum $e^{\\pm}$ pair production. If so this would be the first time the astronomical Schwinger effect has been observed. These "Schwinger sparks" would be an intermittent but extremely powerful, $\\sim 10^3 L_{\\astrosun}$, 10 PeV $e^{\\pm}$ accelerator in the heart of the Crab. These nanosecond duration sparks are generated in a volume less than $1 m^3$ and the existence of such sparks has implications for the small scale structure of the magnetic field of young pulsars such as the Crab. This mechanism may also play a role in producing other enigmatic bright short radio transients such as fast radio bursts.

  13. LHC crab-cavity aspects and strategy

    Energy Technology Data Exchange (ETDEWEB)

    Calaga, R.; Tomas, R.; Zimmermann, F.

    2010-05-23

    The 3rd LHC Crab Cavity workshop (LHC-CC09) took place at CERN in October 2009. It reviewed the current status and identified a clear strategy towards a future crab-cavity implementation. Following the success of crab cavities in KEK-B and the strong potential for luminosity gain and leveling, CERN will pursue crab crossing for the LHC upgrade. We present a summary and outcome of the variousworkshop sessions which have led to the LHC crab-cavity strategy, covering topics like layout, cavity design, integration, machine protection, and a potential validation test in the SPS.

  14. Burrow ventilation and associated porewater irrigation by the polychaete Marenzelleria viridis

    DEFF Research Database (Denmark)

    Quintana, Cintia Organo; Hansen, Tanja; Delefosse, Matthieu;

    2011-01-01

    Burrow ventilation of benthic infauna generates water currents that irrigate the interstices of the sediments surrounding the burrow walls. Such activities have associated effects on biogeochemical processes affecting ultimately important ecosystem processes. In this study, the ventilation and ir...

  15. Development of earthworm burrow systems and the influence of earthworms on soil hydrology.

    NARCIS (Netherlands)

    Ligthart, T.N.

    1996-01-01

    Inoculation of earthworms can help to restore or ameliorate land qualities. Earthworms create burrows and alter the structure of the soil matrix, which influence the water infiltration, drainage, water retention and the aeration of the soil. The way and rate of the development of earthworm burrow sy

  16. Irrigation patterns in permeable sediments induced by burrow ventilation: a case study of Arenicola marina

    NARCIS (Netherlands)

    Meysman, F.J.R.; Galaktionov, O.; Middelburg, J.J.

    2005-01-01

    In sandy sediments, a strong connection exists between the physics of flow and the ecology of burrow-ventilating macrofauna. We developed a general modelling procedure that quantifies this link involving 3 steps. (1) Burrow-ventilating organisms can be described as mechanical pumps. (2) The pumping

  17. Water pumping and analysis of flow in burrowing zoobenthos - a short overview

    DEFF Research Database (Denmark)

    Riisgård, H. U.; Larsen, Poul Scheel

    2002-01-01

    Measurement of water pumping rates of burrowing animals is of crucial importance for the study of many processes both within and above the sea floor. This short review deals with water pumping and analysis of flow, including available techniques and bio-fluid mechanical theory, in burrowing deposit...

  18. Phylogenetic patterns and the adaptive evolution of osmoregulation in fiddler crabs (Brachyura, Uca)

    Science.gov (United States)

    Faria, Samuel Coelho; Provete, Diogo Borges; Thurman, Carl Leo

    2017-01-01

    Salinity is the primary driver of osmoregulatory evolution in decapods, and may have influenced their diversification into different osmotic niches. In semi-terrestrial crabs, hyper-osmoregulatory ability favors sojourns into burrows and dilute media, and provides a safeguard against hemolymph dilution; hypo-osmoregulatory ability underlies emersion capability and a life more removed from water sources. However, most comparative studies have neglected the roles of the phylogenetic and environmental components of inter-specific physiological variation, hindering evaluation of phylogenetic patterns and the adaptive nature of osmoregulatory evolution. Semi-terrestrial fiddler crabs (Uca) inhabit fresh to hyper-saline waters, with species from the Americas occupying higher intertidal habitats than Indo-west Pacific species mainly found in the low intertidal zone. Here, we characterize numerous osmoregulatory traits in all ten fiddler crabs found along the Atlantic coast of Brazil, and we employ phylogenetic comparative methods using 24 species to test for: (i) similarities of osmoregulatory ability among closely related species; (ii) salinity as a driver of osmoregulatory evolution; (iii) correlation between salt uptake and secretion; and (iv) adaptive peaks in osmoregulatory ability in the high intertidal American lineages. Our findings reveal that osmoregulation in Uca exhibits strong phylogenetic patterns in salt uptake traits. Salinity does not correlate with hyper/hypo-regulatory abilities, but drives hemolymph osmolality at ambient salinities. Osmoregulatory traits have evolved towards three adaptive peaks, revealing a significant contribution of hyper/hypo-regulatory ability in the American clades. Thus, during the evolutionary history of fiddler crabs, salinity has driven some of the osmoregulatory transformations that underpin habitat diversification, although others are apparently constrained phylogenetically. PMID:28182764

  19. Human threats to sandy beaches: A meta-analysis of ghost crabs illustrates global anthropogenic impacts.

    Science.gov (United States)

    Schlacher, Thomas A.; Lucrezi, Serena; Connolly, Rod M.; Peterson, Charles H.; Gilby, Ben L.; Maslo, Brooke; Olds, Andrew D.; Walker, Simon J.; Leon, Javier X.; Huijbers, Chantal M.; Weston, Michael A.; Turra, Alexander; Hyndes, Glenn A.; Holt, Rebecca A.; Schoeman, David S.

    2016-02-01

    Beach and coastal dune systems are increasingly subjected to a broad range of anthropogenic pressures that on many shorelines require significant conservation and mitigation interventions. But these interventions require reliable data on the severity and frequency of adverse ecological impacts. Such evidence is often obtained by measuring the response of 'indicator species'. Ghost crabs are the largest invertebrates inhabiting tropical and subtropical sandy shores and are frequently used to assess human impacts on ocean beaches. Here we present the first global meta-analysis of these impacts, and analyse the design properties and metrics of studies using ghost-crabs in their assessment. This was complemented by a gap analysis to identify thematic areas of anthropogenic pressures on sandy beach ecosystems that are under-represented in the published literature. Our meta-analysis demonstrates a broad geographic reach, encompassing studies on shores of the Pacific, Indian, and Atlantic Oceans, as well as the South China Sea. It also reveals what are, arguably, two major limitations: i) the near-universal use of proxies (i.e. burrow counts to estimate abundance) at the cost of directly measuring biological traits and bio-markers in the organism itself; and ii) descriptive or correlative study designs that rarely extend beyond a simple 'compare and contrast approach', and hence fail to identify the mechanistic cause(s) of observed contrasts. Evidence for a historically narrow range of assessed pressures (i.e., chiefly urbanisation, vehicles, beach nourishment, and recreation) is juxtaposed with rich opportunities for the broader integration of ghost crabs as a model taxon in studies of disturbance and impact assessments on ocean beaches. Tangible advances will most likely occur where ghost crabs provide foci for experiments that test specific hypotheses associated with effects of chemical, light and acoustic pollution, as well as the consequences of climate change (e

  20. Crab shedding-system designs

    OpenAIRE

    1997-01-01

    There are as many ways to build and arrange crab shedding setups as there are people who make them. The following drawings are suggestions based on the experiencesof successful shedders. You may find changes that suit your operation better. (8pp.)

  1. River bank burrowing by invasive crayfish: Spatial distribution, biophysical controls and biogeomorphic significance.

    Science.gov (United States)

    Faller, Matej; Harvey, Gemma L; Henshaw, Alexander J; Bertoldi, Walter; Bruno, Maria Cristina; England, Judy

    2016-11-01

    Invasive species generate significant global environmental and economic costs and represent a particularly potent threat to freshwater systems. The biogeomorphic impacts of invasive aquatic and riparian species on river processes and landforms remain largely unquantified, but have the potential to generate significant sediment management issues within invaded catchments. Several species of invasive (non-native) crayfish are known to burrow into river banks and visual evidence of river bank damage is generating public concern and media attention. Despite this, there is a paucity of understanding of burrow distribution, biophysical controls and the potential significance of this problem beyond a small number of local studies at heavily impacted sites. This paper presents the first multi-catchment analysis of this phenomenon, combining existing data on biophysical river properties and invasive crayfish observations with purpose-designed field surveys across 103 river reaches to derive key trends. Crayfish burrows were observed on the majority of reaches, but burrowing tended to be patchy in spatial distribution, concentrated in a small proportion (<10%) of the length of rivers surveyed. Burrow distribution was better explained by local bank biophysical properties than by reach-scale properties, and burrowed banks were more likely to be characterised by cohesive bank material, steeper bank profiles with large areas of bare bank face, often on outer bend locations. Burrow excavation alone has delivered a considerable amount of sediment to invaded river systems in the surveyed sites (3tkm(-1) impacted bank) and this represents a minimum contribution and certainly an underestimate of the absolute yield (submerged burrows were not recorded). Furthermore, burrowing was associated with bank profiles that were either actively eroding or exposed to fluvial action and/or mass failure processes, providing the first quantitative evidence that invasive crayfish may cause or

  2. Landform and surface attributes for prediction of rodent burrows in the Western Usambara Mountains, Tanzania.

    Science.gov (United States)

    Meliyo, Joel L; Massawe, Boniface H J; Msanya, Balthazar M; Kimaro, Didas N; Hieronimo, Proches; Mulungu, Loth S; Kihupi, Nganga I; Deckers, Jozef A; Gulinck, Hubert; Leirs, Herwig

    2014-07-01

    Previous studies suggest that rodent burrows, a proxy for rodent population are important for predicting plague risk areas. However, studies that link landform, surface attributes and rodent burrows in the Western Usambara Mountains in Tanzania are scanty. Therefore, this study was conducted in plague endemic area of the Western Usambara Mountains in northern, Tanzania, to explore the relationship between rodent burrows, and landform and surface attributes. The study was carried out in three areas corresponding to high (Lokome), medium (Lukozi) and low.(Mwangoi) frequency of reported plague cases. Data were collected from 117, 200 and 170 observation sites for Lokome, Lukozi and Mwangoi, respectively using 100 m x 200 m quadrats. Remote sensing and field surveys were used to collect data on landform and surface attributes. Rodent burrows were surveyed and quantified by counting the number of burrows in 20m x 20m grids demarcated on the main 100m x 200m quadrats. The collected data were analysed in R software using boosted regression trees (BRT) technique. Rodent burrows were found at an elevation of above 1600m in the high and medium plague frequency landscapes. No burrows were found in the low plague frequency landscape situated below 1500m. BRT analysis shows a significant relationship between landform characteristics and rodent burrows in both high and medium plague frequency landscapes. Overall, elevation and hillshade are the most important determinants of rodent burrow distribution in the studied landscapes. It is concluded that in high altitudes, specific landform attributes (hill-shade, slope, elevation) and vegetation cover- favour rodent burrowing.

  3. Mangrove carbon sink. Do burrowing crabs contribute to sediment carbon storage? Evidence from a Kenyan mangrove system

    OpenAIRE

    2014-01-01

    Mangrove ecosystems are acknowledged as a significant carbon reservoir, with a potential key role as carbon sinks. Little however is known on sediment/soil capacity to store organic carbon and the impact of benthic fauna on soil organic carbon (SOC) stock in mangrove C-poor soils. This study aimed to investigate the effects of macrobenthos on SOC storage and dynamic in mangrove forest at Gazi Bay (Kenya). Although the relatively low amount of organic carbon (OC%) in these soils, they resulted...

  4. AFSC/RACE/SAP/Jensen: Bitter crab disease mortality in SE Alaska Tanner crab

    Data.gov (United States)

    National Oceanic and Atmospheric Administration, Department of Commerce — These are data from a laboratory experiment in which wild caught male Tanner crab (Chionoecetes bairdi) from Stephens Passage, SE Alaska were held to evaluate crab...

  5. Population biology of the crab Armases angustipes (Crustacea, Decapoda, Sesarmidae at Brazilian tropical coast

    Directory of Open Access Journals (Sweden)

    Marina de Sá Leitão C. de Araújo

    2014-06-01

    Full Text Available The semi terrestrial crabs are important elements of the fauna of coastal regions. The aim of this study was to analyze the population structure of Armases angustipes (Dana, 1852 at estuaries of the Ariquindá River, considered a non impacted area, and Mamucabas River, considered a few impacted area, on the south coast of state of Pernambuco, Brazil. The species occurred in all months of the year. The number of individuals per month varied, being higher in the months of transition between the seasons. This is probably due to significant seasonal variations of air and burrow temperature and burrow salinity. There was no sexual dimorphism in size of A. angustipes in the mangrove of Ariquindá River, but males were larger than females in the mangrove of Mamucabas River. In both estuaries, the sex ratio did not differ from Mendelian proportion, but showed a deviation for females. The analysis of temporal variation in sex ratio showed significant differences in some months of the year. These variations are due to cyclical events that act distinctly on each sex. In both estuaries, size classes of carapace width were equally represented by both sexes. The ovigerous females of A. angustipes occurred only in some months of the year, especially in summer, in both estuaries. Probably the high phytoplankton productivity observed in summer favors the reproductive activity, since these algae serve as food for the larvae. Specimens of the population of Rio Ariquindá are largest and wider than those of Mamucabas River. This fact, associated with the low abundance of crabs and the lower frequency of ovigerous females observed in Mamucabas River, is an indication that this population may be influenced by the environmental impacts that this estuary has received.

  6. Spilled oil and infaunal activity - Modification of burrowing behavior and redistribution of oil

    Science.gov (United States)

    Clifton, H.E.; Kvenvolden, K.A.; Rapp, J.B.

    1984-01-01

    A series of experiments in Willapa Bay, Washington, indicates the degree to which the presence of spilled oil modifies the burrowing behavior of infauna and the extent to which the animals redistribute oil into intertidal sediment. Small amounts of North Slope crude oil introduced at low tide directly into burrow openings (mostly made by the crustacean Callianassa) resulted in a limited and temporary reduction in the number of burrow openings. In contrast, a layer of oil-saturated sand 1 cm thick buried about 5 cm below the sediment surface sharply reduced the number of burrow openings. After a year, the few new burrows penetrated only the margins of the experimental plot, and bioturbation below the buried oil-saturated sand layer declined dramatically. The experiments suggest that small amounts of oil temporarily stranded by tides in themselves have no long-range effect on burrowing behavior. The fauna, however, are capable of introducing measurable amounts of oil into the subsurface, where it is retained long after the rest of the stranded oil had washed away. A buried layer of oil-saturated sand greatly reduces infaunal activity; the oil presents an effective barrier that can persist for years. The oil incorporated into the sediment from burrow openings showed evidence of degradation after 7 months. In contrast the layer of buried oil remained essentially undergraded after a period of two years, even though oil in lower concentrations above the layer was degraded after a period of one year. This variation in degree of degradation of the buried oil, as well as the heterogeneity of oil distribution wherever the oil has been incorporated from the surface, emphasises the importance of careful sampling in any attempt to locate or monitor the presence of spilled oil in the substrate.In a series of experiments in Willapa Bay, Washington, small amounts of North Slope crude oil introduced at low tide directly into burrow openings resulted in a limited and temporary

  7. Self-burrowing seeds: drag reduction in granular media

    Science.gov (United States)

    Jung, Wonjong; Choi, Sung Mok; Kim, Wonjung; Kim, Ho-Young

    2014-11-01

    We present the results of a combined experimental and theoretical investigation of drag reduction of self-burrowing seeds in granular media. In response to environmental changes in humidity, the awn (a tail-like appendage of seed) of Pelargonium carnosum exhibits coiling-uncoiling deformation which induces the thrust and rotary motions of the head of the seed against the surface of the soil. Using various sizes of glass beads that mimic the granular soil, we measure the thrust forces required for the seed of Pelargonium carnosum to penetrate into granular media with and without rotation. Our quantitative measurements show that the rotation of the seed remarkably reduces the granular drag as compared to the drag against the non-spinning seed. This leads us to conclude that the hygroscopically active awns of Pelargonium carnosum enables its seed to dig into the relatively coarse granular soils.

  8. On Bijective Variants of the Burrows-Wheeler Transform

    CERN Document Server

    Kufleitner, Manfred

    2009-01-01

    The sort transform (ST) is a modification of the Burrows-Wheeler transform (BWT). Both transformations map an arbitrary word of length n to a pair consisting of a word of length n and an index between 1 and n. The BWT sorts all rotation conjugates of the input word, whereas the ST of order k only uses the first k letters for sorting all such conjugates. If two conjugates start with the same prefix of length k, then the indices of the rotations are used for tie-breaking. Both transforms output the sequence of the last letters of the sorted list and the index of the input within the sorted list. In this paper, we discuss a bijective variant of the BWT (due to Scott), proving its correctness and relations to other results due to Gessel and Reutenauer (1993) and Crochemore, Desarmenien, and Perrin (2005). Further, we present a novel bijective variant of the ST.

  9. Brachyuran crab community structure and associated sediment reworking activities in pioneer and young mangroves of French Guiana, South America

    Science.gov (United States)

    Aschenbroich, Adélaïde; Michaud, Emma; Stieglitz, Thomas; Fromard, François; Gardel, Antoine; Tavares, Marcos; Thouzeau, Gérard

    2016-12-01

    This study in French Guiana evaluates the changes of crab assemblages and their bioturbation activities between mangrove early stages (pioneer and young mangrove) and within stages by taking their spatial heterogeneity (tidal channels, flat areas, pools) into account. The results show differences in crab assemblage structure between and within the early stages of mangrove in relation to microhabitat and sediment characteristics. The sediment reworking rates are a function of the biomass or density of particular species (Ucides cordatus, Uca cumulanta) and burrower functional groups. Crab species or functional interactions mediate changes in sediment reworking rates suggesting the need to consider entire benthic communities rather than single species. This study suggests that the role of the microhabitat in determining the biologically-induced sediment reworking rates depends on the age of the mangrove. Feeding activity results in a sediment turnover of 11.7 ± 9.7 gdw m-2 day-1 and 6.8 ± 3.0 gdw m-2 day-1 in the pioneer and young mangroves, respectively. Burrow maintenance excavates 40.5 ± 7.4 gdw m-2 day-1 and 251.3 ± 419.7 gdw m-2 day-1 in the pioneer and young mangroves, respectively. Upscaling to the studied area (Sinnamary estuary: 6 km2), shows that 500 tons.day-1 and 20 tons.day-1 of sediments could be excavated and pelletized, respectively, during the spring tides of the dry season. Thus, biological sediment reworking would greatly contribute to the sedimentary dynamics of the Guianese mangroves under Amazonian influence.

  10. Final focus designs for crab waist colliders

    Science.gov (United States)

    Bogomyagkov, A.; Levichev, E.; Piminov, P.

    2016-12-01

    The crab waist collision scheme promises significant luminosity gain. The successful upgrade of the DA Φ NE collider proved the principle of crab waist collision and increased luminosity 3 times. Therefore, several new projects try to implement the scheme. The paper reviews interaction region designs with the crab waist collision scheme for already existent collider DA Φ NE and SuperKEKB, presently undergoing commissioning, for the projects of SuperB in Italy, CTau in Novosibirsk and FCC-ee at CERN.

  11. An assessment of Hyalella azteca burrowing activity under laboratory sediment toxicity testing conditions.

    Science.gov (United States)

    Doig, Lorne E; Liber, Karsten

    2010-09-01

    Burrowing of the freshwater amphipod Hyalella azteca was evaluated under laboratory conditions similar to those recommended for standard sediment toxicity testing in Canada (EPS 1/RM/33; Environment Canada, 1997) and the United States (EPA/600/R-99/064; US EPA, 2000). Sediment type, time of day (light versus dark), size of animal, and the presence or absence of food were varied to assess their effects on burrowing activity. Hyalella azteca were found to burrow rapidly in fine, organic-rich sediments, but were slower to burrow in a sandy sediment. There was no increase in the number of animals occupying the sediment surface of a fine, organic-rich sediment after 4h of darkness compared to the previous 4h of light. Over a 9- to 10-d duration, a higher percentage of animals occupied the surface of the sandy sediment. The addition of food promoted burrowing in sandy sediment, as did using smaller animals. Overall, longer-duration tests involving older animals and coarse sediments may require formal observation to confirm burrowing and ensure adequate sediment exposure. The addition of food during a test may promote the burrowing of larger animals in coarse sediments, but may not be necessary in field-collected sediments that are not excessively sandy.

  12. Spatial and social organization in a burrow-dwelling lizard (Phrynocephalus vlangalii) from China.

    Science.gov (United States)

    Qi, Yin; Noble, Daniel W A; Fu, Jinzhong; Whiting, Martin J

    2012-01-01

    Shared ecological resources such as burrow complexes can set the stage for social groupings and the evolution of more complex social behavior such as parental care. Paternity testing is increasingly revealing cases of kin-based groupings, and lizards may be a good system to inform on the early evolution of sociality. We examined spatial and social organization in the lizard Phrynocephalus vlangalii from China and tested genetic relatedness (based on eight microsatellite DNA loci) between offspring and parents that shared burrow complexes. Adult males and females had similar spatial patterns: they overlapped most with members of the opposite sex and least with their own sex. Males in better body condition overlapped with more females, and both sexes showed high site fidelity. Most lizards used a single burrow, but some individuals used two or three burrows. While high site fidelity is consistent with sociality in lizards, juveniles did not preferentially share burrows with parents, and we documented only a few cases of parent-offspring associations through burrow sharing. We suggest that P. vlangalii conforms to a classical polygynous mating system in which the burrow forms the core of the male's territory and may be offered as an important resource for females, but this remains to be determined.

  13. Large-diameter burrows of the Triassic Ischigualasto Basin, NW Argentina: paleoecological and paleoenvironmental implications.

    Directory of Open Access Journals (Sweden)

    Carina E Colombi

    Full Text Available Large-diameter ichnofossils comprising three morphotypes have been identified in the Upper Triassic Ischigualasto and Los Colorados formations of northwestern Argentina. These burrows add to the global record of the early appearance of fossorial behavior during early Mesozoic time. Morphotypes 1 and 2 are characterized by a network of tunnels and shafts that can be assigned to tetrapod burrows given similarities with previously described forms. However, differences in diameter, overall morphology, and stratigraphic occurrence allow their independent classification. Morphotype 3 forms a complex network of straight branches that intersect at oblique angles. Their calcareous composition and surface morphology indicate these structures have a composite biogenic origin likely developed due to combined plant/animal interactions. The association of Morphotypes 1 and 2 with fluvial overbank lithologies deposited under an extremely seasonal arid climate confirms interpretations that the early appearance of burrowing behavior was employed by vertebrates in response to both temperature and moisture-stress associated with seasonally or perpetually dry Pangean paleoclimates. Comparisons of burrow morphology and biomechanical attributes of the abundant paleovertebrate fauna preserved in both formations permit interpretations regarding the possible burrow architects for Morphotypes 1 and 2. In the case of the Morphotype 1, the burrow constructor could be one of the small carnivorous cynodonts, Ecteninion or Probelesodon. Assigning an architect for Morphotype 2 is more problematic due to mismatches between the observed burrow morphology and the size of the known Los Colorados vertebrates.

  14. Spatial and social organization in a burrow-dwelling lizard (Phrynocephalus vlangalii from China.

    Directory of Open Access Journals (Sweden)

    Yin Qi

    Full Text Available Shared ecological resources such as burrow complexes can set the stage for social groupings and the evolution of more complex social behavior such as parental care. Paternity testing is increasingly revealing cases of kin-based groupings, and lizards may be a good system to inform on the early evolution of sociality. We examined spatial and social organization in the lizard Phrynocephalus vlangalii from China and tested genetic relatedness (based on eight microsatellite DNA loci between offspring and parents that shared burrow complexes. Adult males and females had similar spatial patterns: they overlapped most with members of the opposite sex and least with their own sex. Males in better body condition overlapped with more females, and both sexes showed high site fidelity. Most lizards used a single burrow, but some individuals used two or three burrows. While high site fidelity is consistent with sociality in lizards, juveniles did not preferentially share burrows with parents, and we documented only a few cases of parent-offspring associations through burrow sharing. We suggest that P. vlangalii conforms to a classical polygynous mating system in which the burrow forms the core of the male's territory and may be offered as an important resource for females, but this remains to be determined.

  15. Synchrotron Reveals Early Triassic Odd Couple: Injured Amphibian and Aestivating Therapsid Share Burrow.

    Directory of Open Access Journals (Sweden)

    Vincent Fernandez

    Full Text Available Fossorialism is a beneficial adaptation for brooding, predator avoidance and protection from extreme climate. The abundance of fossilised burrow casts from the Early Triassic of southern Africa is viewed as a behavioural response by many tetrapods to the harsh conditions following the Permo-Triassic mass-extinction event. However, scarcity of vertebrate remains associated with these burrows leaves many ecological questions unanswered. Synchrotron scanning of a lithified burrow cast from the Early Triassic of the Karoo unveiled a unique mixed-species association: an injured temnospondyl amphibian (Broomistega that sheltered in a burrow occupied by an aestivating therapsid (Thrinaxodon. The discovery of this rare rhinesuchid represents the first occurrence in the fossil record of a temnospondyl in a burrow. The amphibian skeleton shows signs of a crushing trauma with partially healed fractures on several consecutive ribs. The presence of a relatively large intruder in what is interpreted to be a Thrinaxodon burrow implies that the therapsid tolerated the amphibian's presence. Among possible explanations for such unlikely cohabitation, Thrinaxodon aestivation is most plausible, an interpretation supported by the numerous Thrinaxodon specimens fossilised in curled-up postures. Recent advances in synchrotron imaging have enabled visualization of the contents of burrow casts, thus providing a novel tool to elucidate not only anatomy but also ecology and biology of ancient tetrapods.

  16. Chemically mediated burrow recognition in the Mexican tarantula Brachypelma vagans female

    Science.gov (United States)

    Dor, Ariane; Machkour-M'rabet, Salima; Legal, Luc; Williams, Trevor; Hénaut, Yann

    2008-12-01

    Chemically mediated communication is common in spiders but has been poorly studied in burrowing tarantulas. This study aimed to determine whether chemical cues influence the behaviour of females of Brachypelma vagans, a Mexican species of tarantula, during encounters with previously inhabited burrows or with extracts from the silk of conspecific females. In laboratory choice tests, female tarantulas entered a burrow that had previously been inhabited by a conspecific female significantly more frequently than a burrow that had never been inhabited. The identity of the previous inhabitant also affected the number of spiders that chose to enter a burrow. Spiders were quicker to choose and enter a burrow previously inhabited by themselves than a burrow previously inhabited by a conspecific or a burrow that had not been previously inhabited. Hexane, methanol and dichloromethane extracts of conspecific silk elicited different responses from female tarantulas when extracts were placed on filter paper disks at one end of an experimental arena with a control filter paper disk, on to which the corresponding solvent alone had been pipetted, placed on the other end of the arena. Spiders showed the strongest responses to hexane extracts of silk, with a significant preference to move towards the hexane extract and a significantly greater period of time spent in proximity to the hexane extract compared to the control disk. Overall and in contrast to expectations, tarantulas were most strongly attracted to the cues left by other conspecific females. As encounters between B. vagans females usually lead to aggression and mortality of one of the participants, we conclude that chemical cues are not signals that are deliberately released by burrow-inhabiting females but may inadvertently escape and cannot be easily suppressed.

  17. Similar burrow architecture of three arid-zone scorpion species implies similar ecological function

    Science.gov (United States)

    Adams, Amanda M.; Marais, Eugene; Turner, J. Scott; Prendini, Lorenzo; Pinshow, Berry

    2016-08-01

    Many animals reside in burrows that may serve as refuges from predators and adverse environmental conditions. Burrow design varies widely among and within taxa, and these structures are adaptive, fulfilling physiological (and other) functions. We examined the burrow architecture of three scorpion species of the family Scorpionidae: Scorpio palmatus from the Negev desert, Israel; Opistophthalmus setifrons, from the Central Highlands, Namibia; and Opistophthalmus wahlbergii from the Kalahari desert, Namibia. We hypothesized that burrow structure maintains temperature and soil moisture conditions optimal for the behavior and physiology of the scorpion. Casts of burrows, poured in situ with molten aluminum, were scanned in 3D to quantify burrow structure. Three architectural features were common to the burrows of all species: (1) a horizontal platform near the ground surface, long enough to accommodate the scorpion, located just below the entrance, 2-5 cm under the surface, which may provide a safe place where the scorpion can monitor the presence of potential prey, predators, and mates and where the scorpion warms up before foraging; (2) at least two bends that might deter incursion by predators and may reduce convective ventilation, thereby maintaining relatively high humidity and low temperature; and (3) an enlarged terminal chamber to a depth at which temperatures are almost constant (±2-4 °C). These common features among the burrows of three different species suggest that they are important for regulating the physical environment of their inhabitants and that burrows are part of scorpions' "extended physiology" ( sensu Turner, Physiol Biochem Zool 74:798-822, 2000).

  18. The Crab Nebula flaring activity

    Directory of Open Access Journals (Sweden)

    G. Montani

    2014-12-01

    Full Text Available The discovery made by AGILE and Fermi of a short time scale flaring activity in the gamma-ray energy emission of the Crab Nebula is a puzzling and unexpected feature, challenging particle acceleration theory. In the present work we propose the shock-induced magnetic reconnection as a viable mechanism to explain the Crab flares. We postulate that the emitting region is located at ∼1015 cm from the central pulsar, well inside the termination shock, which is exactly the emitting region size as estimated by the overall duration of the phenomenon ∼1 day. We find that this location corresponds to the radial distance at which the shock-induced magnetic reconnection process is able to accelerate the electrons up to a Lorentz factor ∼109, as required by the spectral fit of the observed Crab flare spectrum. The main merit of the present analysis is to highlight the relation between the observational constraints to the flare emission and the radius at which the reconnection can trigger the required Lorentz factor. We also discuss different scenarios that can induce the reconnection. We conclude that the existence of a plasma instability affecting the wind itself as the Weibel instability is the privileged scenario in our framework.

  19. The Crab Nebula flaring activity

    Energy Technology Data Exchange (ETDEWEB)

    Montani, G., E-mail: giovanni.montani@frascati.enea.it [ENEA – C.R, UTFUS-MAG, via Enrico Fermi 45, I-00044 Frascati (RM) (Italy); Dipartimento di Fisica, Università di Roma “Sapienza”, p.le Aldo Moro 5, I-00185 Roma (Italy); Bernardini, M.G. [INAF – Osservatorio Astronomico di Brera, via Bianchi 46, I-23807 Merate (Italy)

    2014-12-12

    The discovery made by AGILE and Fermi of a short time scale flaring activity in the gamma-ray energy emission of the Crab Nebula is a puzzling and unexpected feature, challenging particle acceleration theory. In the present work we propose the shock-induced magnetic reconnection as a viable mechanism to explain the Crab flares. We postulate that the emitting region is located at ∼10{sup 15} cm from the central pulsar, well inside the termination shock, which is exactly the emitting region size as estimated by the overall duration of the phenomenon ∼1 day. We find that this location corresponds to the radial distance at which the shock-induced magnetic reconnection process is able to accelerate the electrons up to a Lorentz factor ∼10{sup 9}, as required by the spectral fit of the observed Crab flare spectrum. The main merit of the present analysis is to highlight the relation between the observational constraints to the flare emission and the radius at which the reconnection can trigger the required Lorentz factor. We also discuss different scenarios that can induce the reconnection. We conclude that the existence of a plasma instability affecting the wind itself as the Weibel instability is the privileged scenario in our framework.

  20. Behavioural adaptations to moisture as an environmental constraint in a nocturnal burrow-inhabiting Kalahari detritivore Parastizopus armaticeps Peringuey (Coleoptera: Tenebrionidae

    Directory of Open Access Journals (Sweden)

    O.A.E. Rasa

    1994-09-01

    Full Text Available The nocturnal desert detritivore Parastiz.opus armaticeps shows differences in surface activity patterns and burrow fidelity depending on surface humidity. After rain approximately half of the beetle population, independent of sex, is highly vagile and disperses over long distances. During drought, beetles are more sedentary and show higher burrow fidelity. They also inhabit burrows that are longer and deeper than non-inhabited ones, such burrows being relatively scarce. Burrow fidelity and the adoption of a more sedentary habit during drought are considered strategies to avoid the risks of not locating a suitable burrow before sunrise and subsequent desiccation in shallow burrows.

  1. Status assessment and conservation plan for the western burrowing owl in the United States

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — The Western Burrowing Owl (Athene cunicularia hypugaea) is a grassland specialist distributed throughout w. North America, primarily in open areas with short...

  2. Rodents new to the diet of the western burrowing owl(athene CUNICULARIA HYPUGAEA )

    Science.gov (United States)

    Wiluford, D.L.; Woodin, M.C.; Skoruppa, M.K.; Hickman, G.C.

    2009-01-01

    The northern pygmy mouse (Baiomys taylori), fulvous harvest mouse (Reithrodontomys fulvescens), and Merriam's pocket mouse (Pemgnathus merriami) are new to the diet of the western burrowing owl (Athene cuniculana hypugaed). All three species were identified from remains in regurgitated pellets collected from roost sites of burrowing owls in southern Texas over a period of 4 winters. Together, northern pygmy mice and fulvous harvest mice represented 58% of mammals identified in 182 pellets regurgitated by western burrowing owls. Merriam's pocket mouse accounted for only 4% of identified mammalian prey. Frequency of occurrence in pellets was 16% for northern pygmv mice, 11% for fulvous harvest mice, and 3% for Merriam's pocket mice. The primary reason for absence of these species in previous studies of foods of western burrowing owls is that most were conducted in latitudes north of these southern-distributed species of mammals.

  3. Colorado Plateau Rapid Ecoregion Assessment Conservation Elements - Terrestrial Species: Burrowing Owl

    Data.gov (United States)

    Bureau of Land Management, Department of the Interior — This map shows the potential current distribution of burrowing owl, in the context of current and near-term terrestrial intactness and long-term potential for...

  4. EPA Removes Burrows Sanitation Site in Michigan from National List of Most Contaminated Sites

    Science.gov (United States)

    For Immediate Release No. 15-OPA142 CHICAGO - The U.S. Environmental Protection Agency today announced that the Burrows Sanitation Superfund site in Hartford Township, Van Buren County, Michigan, has been officially removed from the Agency's l

  5. HUBBLE CAPTURES DYNAMICS OF CRAB NEBULA (color)

    Science.gov (United States)

    2002-01-01

    A new sequence of Hubble Space Telescope images of the remnant of a tremendous stellar explosion is giving astronomers a remarkable look at the dynamic relationship between the tiny Crab Pulsar and the vast nebula that it powers. This colorful photo shows a ground-based image of the entire Crab Nebula, the remnant of a supernova explosion witnessed over 900 years ago. The nebula, which is 10 light-years across, is located 7,000 light-years away in the constellation Taurus. The green, yellow and red filaments concentrated toward the edges of the nebula are remnants of the star that were ejected into space by the explosion. At the center of the Crab Nebula lies the Crab Pulsar -- the collapsed core of the exploded star. The Crab Pulsar is a rapidly rotating neutron star -- an object only about six miles across, but containing more mass than our Sun. As it rotates at a rate of 30 times per second the Crab Pulsar's powerful magnetic field sweeps around, accelerating particles, and whipping them out into the nebula at speeds close to that of light. The blue glow in the inner part of the nebula -- light emitted by energetic electrons as they spiral through the Crab's magnetic field -- is powered by the Crab Pulsar. Credit: Jeff Hester and Paul Scowen (Arizona State University), and NASA

  6. Novel Geometries for the LHC Crab Cavity

    Energy Technology Data Exchange (ETDEWEB)

    Hall, B. [Lancaster Univ. (United Kingdom); Burt, G. [Lancaster Univ. (United Kingdom); Smith, J. D.A. [Lancaster Univ. (United Kingdom); Rimmer, R. [Thomas Jefferson National Accelerator Facility (TJNAF), Newport News, VA (United States); Wang, H. [Thomas Jefferson National Accelerator Facility (TJNAF), Newport News, VA (United States); Delayen, J. [Thomas Jefferson National Accelerator Facility (TJNAF), Newport News, VA (United States); Calaga, R. [Brookhaven National Lab. (BNL), Upton, NY (United States)

    2009-05-01

    In 2017 the LHC is envisioned to increase its luminosity via an upgrade. This upgrade is likely to require a large crossing angle hence a crab cavity is required to align the bunches prior to collision. There are two possible schemes for crab cavity implementation, global and local. In a global crab cavity the crab cavity is far from the IP and the bunch rotates back and forward as it traverses around the accelerator in a closed orbit. For this scheme a two-cell elliptical squashed cavity at 800 MHz is preferred. To avoid any potential beam instabilities all the parasitic modes of the cavities must be damped strongly, however crab cavities have lower order and same order modes in addition to the usual higher order modes and hence a novel damping scheme must be used to provide sufficient damping of these modes. In the local scheme two crab cavities are placed at each side of the IP two start and stop rotation of the bunches. This would require crab cavities much smaller transversely than in the global scheme but the frequency cannot be increased any higher due to the long bunch length of the LHC beam. This will require a novel compact crab cavity design. A superconducting version of a two rod coaxial deflecting cavity as a suitable design is proposed in this paper.

  7. The effect of sewage discharge on the ecosystem engineering activities of two East African fiddler crab species: consequences for mangrove ecosystem functioning.

    Science.gov (United States)

    Bartolini, Fabrizio; Cimò, Filippo; Fusi, Marco; Dahdouh-Guebas, Farid; Lopes, Gil Penha; Cannicci, Stefano

    2011-02-01

    A number of studies have suggested that mangrove forests and their faunal components may be pre-adapted to the impact of organic waste discharge, making them possible natural wastewater treatment wetlands. However, the results from recent research are contradictory. Some studies have shown that negative effects, sometimes subtle and difficult to observe, can be detected on specific biotic components of forests subjected to organic pollution. Therefore, the aim of the present study was to investigate possible alterations in the ecosystem engineering activities of a fiddler crab community dominating the landward belts of Kenyan mangrove forests. The total processed sediment produced by burrowing and foraging activities in a population from a peri-urban mangrove area receiving untreated domestic sewage was compared with that from a forest not affected by urban wastewater. The results showed how the peri-urban site hosted a higher biomass of crabs, which produced a significantly lower amount of processed sediment compared with the pristine site, resulting in a lower total top sediment mixing activity of the crabs. Thus, the present study showed a link between sewage exposure and top sediment reworking by crabs, which is potentially beneficial for mangrove growth and ecosystem functioning. This represents a possible example of cryptic ecological degradation in mangal systems.

  8. Most Detailed Image of the Crab Nebula

    Science.gov (United States)

    2005-01-01

    This new Hubble image -- one among the largest ever produced with the Earth-orbiting observatory -- shows the most detailed view so far of the entire Crab Nebula ever made. The Crab is arguably the single most interesting object, as well as one of the most studied, in all of astronomy. The image is the largest image ever taken with Hubble's WFPC2 workhorse camera. The Crab Nebula is one of the most intricately structured and highly dynamical objects ever observed. The new Hubble image of the Crab was assembled from 24 individual exposures taken with the NASA/ESA Hubble Space Telescope and is the highest resolution image of the entire Crab Nebula ever made.

  9. 50 CFR 680.6 - Crab economic data report (EDR).

    Science.gov (United States)

    2010-10-01

    ... 50 Wildlife and Fisheries 9 2010-10-01 2010-10-01 false Crab economic data report (EDR). 680.6... General § 680.6 Crab economic data report (EDR). Persons participating in the CR crab fisheries are... complete. Use these tables to complete the EDRs described in this section: Table 1, Crab...

  10. Novel Geometries for the LHC CRAB Cavity

    CERN Document Server

    Hall, Ben

    2010-01-01

    In 2017 the LHC is envisioned to increase its luminosity via an upgrade. This upgrade is likely to require a large crossing angle hence a crab cavity is required to align the bunches prior to collision. There are two possible schemes for crab cavity implementation, global and local. In a global crab cavity the crab cavity is far from the IP and the bunch rotates back and forward as it traverses around the accelerator in a closed orbit. For this scheme a two-cell elliptical squashed cavity at 800 MHz is preferred. To avoid any potential beam instabilities all the parasitic modes of the cavities must be damped strongly, however crab cavities have lower order and same order modes in addition to the usual higher order modes and hence a novel damping scheme must be used to provide sufficient damping of these modes. In the local scheme two crab cavities are placed at each side of the IP two start and stop rotation of the bunches. This would require crab cavities much smaller transversely than in the global scheme b...

  11. What brakes the Crab pulsar?

    CERN Document Server

    Čadež, A; Barbieri, C; Calvani, M; Naletto, G; Barbieri, M; Ponikvar, D

    2015-01-01

    Optical observations provide convincing evidence that the optical phase of the Crab pulsar follows the radio one closely. Since optical data do not depend on dispersion measure variations, they provide a robust and independent confirmation of the radio timing solution. The aim of this paper is to find a global mathematical description of Crab pulsar's phase as a function of time for the complete set of published Jodrell Bank radio ephemerides (JBE) in the period 1988-2014. We apply the mathematical techniques developed for analyzing optical observations to the analysis of JBE. We break the whole period into a series of episodes and express the phase of the pulsar in each episode as the sum of two analytical functions. The first function is the best-fitting local braking index law, and the second function represents small residuals from this law with an amplitude of only a few turns, which rapidly relaxes to the local braking index law. From our analysis, we demonstrate that the power law index undergoes "inst...

  12. Design of the ILC Crab Cavity System

    Energy Technology Data Exchange (ETDEWEB)

    Adolphsen, C.; Beard, C.; Bellantoni, L.; Burt, G.; Carter, R.; Chase, B.; Church, M.; Dexter, A.; Dykes, M.; Edwards, H.; Goudket, P; Jenkins, R.; Jones, R.M.; Kalinin,; Khabiboulline, T.; Ko, K.; Latina, A.; Li, Z.; Ma, L.; McIntosh, P.; Ng, C.; /SLAC /Daresbury /Fermilab /Cockcroft Inst. Accel. Sci. Tech. /CERN

    2007-08-15

    The International Linear Collider (ILC) has a 14 mrad crossing angle in order to aid extraction of spent bunches. As a result of the bunch shape at the interaction point, this crossing angle at the collision causes a large luminosity loss which can be recovered by rotating the bunches prior to collision using a crab cavity. The ILC baseline crab cavity is a 9-cell superconducting dipole cavity operating at a frequency of 3.9 GHz. In this paper the design of the ILC crab cavity and its phase control system, as selected for the RDR in February 2007 is described in fuller detail.

  13. Ultraviolet studies of the Crab Nebula

    Science.gov (United States)

    Talavera, A.

    2017-03-01

    The Crab Nebula (Messier 1) is one of the most observed sources with the XMM-Newton space telescope of ESA. The Crab and its related pulsar are a calibration source for the on-board X-rays cameras. There are around 80 observations between 2000 and 2015. In this observations, the XMM-Newton Optical and UV Monitor (OM) has also been used. We present a preliminary study of the Crab using images obtained the OM UV filters at 291, 231 and 212 nm. Photometric data for the pulsar (PSR0531+21), created in the supernova event of AD 1054 origin of the nebula, are also presented

  14. A large bubble around the Crab Nebula

    Science.gov (United States)

    Romani, Roger W.; Reach, William T.; Koo, Bon Chul; Heiles, Carl

    1990-01-01

    IRAS and 21 cm observations of the interstellar medium around the Crab nebula show evidence of a large bubble surrounded by a partial shell. If located at the canonical 2 kpc distance of the Crab pulsar, the shell is estimated to have a radius of about 90 pc and to contain about 50,000 solar masses of swept-up gas. The way in which interior conditions of this bubble can have important implications for observations of the Crab are described, and the fashion in which presupernova evolution of the pulsar progenitor has affected its local environment is described.

  15. Map-Based Repowering and Reorganization of a Wind Resource Area to Minimize Burrowing Owl and Other Bird Fatalities

    Directory of Open Access Journals (Sweden)

    Lee Neher

    2009-10-01

    Full Text Available Wind turbines in the Altamont Pass Wind Resource Area (Alameda/Contra Costa Counties, California, USA generate about 730 GWh of electricity annually, but have been killing thousands of birds each year, including >2,000 raptors and hundreds of burrowing owls. We have developed collision hazard maps and hazard ratings of wind turbines to guide relocation of existing wind turbines and careful repowering to modern turbines to reduce burrowing owl fatalities principally, and other birds secondarily. Burrowing owls selected burrow sites lower on slopes and on smaller, shallower slopes than represented by the average 10 × 10 m2 grid cell among 187,908 grid cells sampled from 2,281,169 grid cells comprising a digital elevation model (DEM of the study area. Fuzzy logic and discriminant function analysis produced likelihood surfaces encompassing most burrowing owl burrows within a fraction of the study area, and the former corresponded with burrowing owl fatalities and the latter with other raptor fatalities. Our ratings of wind turbine hazard were more predictive of burrowing owl fatalities, but would be more difficult to implement. Careful repowering to modern wind turbines would most reduce fatalities of burrowing owls and other birds while adding about 1,000 GWh annually toward California’s 33% Renewable Portfolio Standard.

  16. Effect of particle size frequency distribution of the substratum on the burrowing ability of Chiridota rigida (semper) (Echinodermata: holothuroidea)

    Energy Technology Data Exchange (ETDEWEB)

    Lawrence, J.M.; Murdoch, J.

    1977-01-01

    Chiridota rigida (Semper) burrows easily into well-sorted substrata, moving between particles in coarse substrata and pushing aside particles in fine substrata. Chiridota rigida does not burrow easily into poorly sorted substrata because spaces which can be penetrated are not available and because the particles cannot be moved as a result of increased stability of the substratum. In poorly sorted substrata, burrowing ability increases with increases in the proportion of fine particles. It would appear that the distribution and abundance of chiridota rigida and other chiridotids would be affected by the effect of the substratum on their ability to burrow.

  17. Foraging behavior of the mangrove sesarmid crab Neosarmatium trispinosum enhances food intake and nutrient retention in a low-quality food environment

    Science.gov (United States)

    Harada, Yota; Lee, S. Y.

    2016-06-01

    The large sesarmid crab Neosarmatium trispinosum has been reported to actively collect freshly fallen mangrove leaves and store them in its burrow where they are assumed to age prior to consumption. This leaf-catching behavior was hypothesized to improve the palatability and nutritional quality of leaves through leaching of feeding deterrent and microbial enrichment during storage. Earlier studies also hypothesized that N. trispinosum feeds on sediment or animal material to meet their N needs. A series of experiments was carried out to investigate the foraging behavior of N. trispinosum against these hypotheses. Study of foraging behavior using remotely operated cameras indicated that this crab spends the far majority of time (97.5 ± 2.5%, SD) underground and only a small percentage of time outside its burrow foraging (2.2 ± 2.3%). Collection of fresh mangrove litter was swift but no record of predation was evident over 31 h of video records. A field leaf tethering experiment showed that this crab started to consume the leaves immediately after collection rather than storing whole leaves, refuting the leaf-aging hypothesis. N. trispinosum also showed a preference for senescent yellow leaves over decaying brown leaves. This behavior may only aim to stock leaves (i.e. to ensure food availability) rather than conditioning them through decay (i.e. to improve food quality). Analysis of gut contents showed that vascular plant material was the dominant food item (83.3 ± 4.6%), followed by sediment (9.2 ± 4.6%) but no animal materials were recorded. N. trispinosum therefore relies minimally on animal food but are capable of removing 50% of the daily leaf litter production. Elemental C, N analysis shows that sediment inside the burrow is a sufficient potential food source (C/N = 13 to 15). While having a lower C/N ratio than fresh green or yellow leaves, the N content of sediment (∼0.1%) was significantly lower than those of mangrove leaves (0.3-0.9%), and may thus

  18. An Integrated Experimental and Modeling Approach to Predict Sediment Mixing from Benthic Burrowing Behavior.

    Science.gov (United States)

    Roche, Kevin R; Aubeneau, Antoine F; Xie, Minwei; Aquino, Tomás; Bolster, Diogo; Packman, Aaron I

    2016-09-20

    Bioturbation is the dominant mode of sediment transport in many aquatic environments and strongly influences both sediment biogeochemistry and contaminant fate. Available bioturbation models rely on highly simplified biodiffusion formulations that inadequately capture the behavior of many benthic organisms. We present a novel experimental and modeling approach that uses time-lapse imagery to directly relate burrow formation to resulting sediment mixing. We paired white-light imaging of burrow formation with fluorescence imaging of tracer particle redistribution by the oligochaete Lumbriculus variegatus. We used the observed burrow formation statistics and organism density to parametrize a parsimonious model for sediment mixing based on fundamental random walk theory. Worms burrowed over a range of times and depths, resulting in homogenization of sediments near the sediment-water interface, rapid nonlocal transport of tracer particles to deep sediments, and large areas of unperturbed sediments. Our fundamental, parsimonious random walk model captures the central features of this highly heterogeneous sediment bioturbation, including evolution of the sediment-water interface coupled with rapid near-surface mixing and anomalous late-time mixing resulting from infrequent, deep burrowing events. This approach provides a general, transferable framework for explicitly linking sediment transport to governing biophysical processes.

  19. Crab Orchard National Wildlife Refuge : Sign Plan

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — The Crab Orchard National Wildlife Refuge Sign Plan explains how signs are used on the Refuge to help guide and educate visitors. An inventory of current signs is...

  20. Monitoring the Crab Nebula with LOFT

    Science.gov (United States)

    Wilson-Hodge, Colleen A.

    2012-01-01

    From 2008-2010, the Crab Nebula was found to decline by 7% in the 15-50 keV band, consistently in Fermi GBM, INTEGRAL IBIS, SPI, and JEMX, RXTE PCA, and Swift BAT. From 2001-2010, the 15-50 keV flux from the Crab Nebula typically varied by about 3.5% per year. Analysis of RXTE PCA data suggests possible spectral variations correlated with the flux variations. I will present estimates of the LOFT sensitivity to these variations. Prior to 2001 and since 2010, the observed flux variations have been much smaller. Monitoring the Crab with the LOFT WFM and LAD will provide precise measurements of flux variations in the Crab Nebula if it undergoes a similarly active episode.

  1. The Pulsar in the Crab Nebula

    CERN Document Server

    Lewandowska, Natalia

    2015-01-01

    The Crab pulsar belongs to one of the most studied stellar objects in the sky. Since its accidental detection in 1968, its pulsed emission has been observed throughout most of the electromagnetic spectrum. Although currently one of more than 2000 known pulsars, its way of work has remained not understood making the Crab pulsar an object of continuous studies and interest. Referring to the pulsed emission of the Crab pulsar only at radio wavelengths, it reveals a diversity of different phenomena. They range from deviations of the predicted slowing down process of the pulsar with time (long time phenomena) to an irregularity of its single pulse emission (short time phenomena). Similar and different kinds of deviations are observed at other wavelengths. Consequently, the Crab pulsar provides a large diversity of different emission characteristics which have remained difficult to interpret with a uniform theoretical approach including all observed properties. Since a review of all currently examined properties of...

  2. Crab Orchard National Wildlife Refuge : Hunting Plan

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — This plan provides guidelines for the administration of hunting activity and for the development, maintenance, and enforcement of regulations and guidelines on Crab...

  3. The anatomy of the king crab Hapalogaster mertensii Brandt, 1850 (Anomura: Paguroidea: Hapalogastridae): new insights into the evolutionary transformation of hermit crabs into king crabs

    NARCIS (Netherlands)

    Keiler, J.; Richter, S.; Wirkner, C.S.

    2015-01-01

    The emergence of king crabs from a hermit crab-like ancestor is one of the most curious events in decapod evolution. King crabs comprise two taxa, Lithodidae and Hapalogastridae, and while lithodids have formed the focus of various anatomical studies, the internal anatomy of hapalogastrids has never

  4. Crabs

    Digital Repository Service at National Institute of Oceanography (India)

    Chatterji, A.

    stream_size 2 stream_content_type text/plain stream_name Biodiversity_Western_Ghats_Inf_Kit_1994_3.6_1.pdf.txt stream_source_info Biodiversity_Western_Ghats_Inf_Kit_1994_3.6_1.pdf.txt Content-Encoding ISO-8859-1 Content-Type text...

  5. Influence of vegetation on the nocturnal foraging behaviors and vertebrate prey capture by endangered Burrowing Owls

    Directory of Open Access Journals (Sweden)

    Alan Marsh

    2014-06-01

    Full Text Available Restrictions in technology have limited past habitat selection studies for many species to the home-range level, as a finer-scale understanding was often not possible. Consequently, these studies may not identify the true mechanism driving habitat selection patterns, which may influence how such results are applied in conservation. We used GPS dataloggers with digital video recorders to identify foraging modes and locations in which endangered Burrowing Owls (Athene cunicularia captured prey. We measured the coarse and fine-scale characteristics of vegetation at locations in which owls searched for, versus where they caught, vertebrate prey. Most prey items were caught using hover-hunting. Burrowing Owls searched for, and caught, vertebrate prey in all cover types, but were more likely to kill prey in areas with sparse and less dense vegetative cover. Management strategies designed to increase Burrowing Owl foraging success in the Canadian prairies should try to ensure a mosaic of vegetation heights across cover types.

  6. Habitat requirements and burrowing depths of rodents in relation to shallow waste burial sites

    Energy Technology Data Exchange (ETDEWEB)

    Gano, K.A.; States, J.B.

    1982-05-01

    The purpose of this paper is to provide a review of the literature and summarize information on factors affecting habitat selection and maximum recorded burrowing depths for representative small mammals that we consider most likely to inhibit waste burial sites in arid and semi-arid regions of the West. The information is intended for waste management designers who need to know what to expect from small mammals that may be present at a particular site. Waste repositories oculd be designed to exclude the deep burrowing rodents of a region by creating an unattractive habitat over the waste. Summaries are given for habitat requirements of each group along with generalized modifications that could be employed to deter habitation. Representatives from the major groups considered to be deep burrowers are discussed. Further, detailed information about a particular species can be obtained from the references cited.

  7. Linking microbial enzymatic activities and functional diversity of soil around earthworm burrows and casts

    Directory of Open Access Journals (Sweden)

    Jerzy Lipiec

    2016-08-01

    Full Text Available The aim of this work was to evaluate the effect of earthworms (Lumbricidae on the enzymatic activity and microbial functional diversity in the burrow system (burrow wall 0–3 mm, transitional zone 3–7 mm, bulk soil >20 mm from the burrow wall and cast aggregates of a loess soil under a pear orchard. The dehydrogenase, β-glucosidase, protease, alkaline phosphomonoesterase, and acid phosphomonoesterase enzymes were assessed using standard methods. The functional diversity (catabolic potential was assessed using the Average Well Color Development and Richness Index following the community level physiological profiling from Biolog Eco Plates. All measurements were done using soil from each compartment immediately after in situ sampling in spring. The enzymatic activites including dehydrogenase, protease, β-glucosidase and alkaline phosphomonoesterase were appreciably greater in the burrow wall or casts than in bulk soil and transitional zone. Conversely, acid phosphomonoesterase had the largest value in the bulk soil. Average Well Color Development in both the transitional zone and the bulk soil (0.98-0.94 A590nm were more than eight times higher than in the burrow walls and casts. The lowest richness index in the bulk soil (15 utilized substrates increased by 86-113% in all the other compartments. The PC1 in principal component analysis (PCA mainly differentiated the burrow walls and the transitional zone. Utilization of all substrate categories was the lowest in the bulk soil. The PC2 differentiated the casts from the other compartments. The enhanced activity of a majority of the enzymes and increased microbial functional diversity in most earthworm-influenced compartments make the soils less vulnerable to degradation and thus increases the stability of ecologically relevant processes in the orchard ecosystem.

  8. Behavioural and Physiological Implications of a Burrow-dwelling Lifestyle for Two Species of Upogebiid Mud-shrimp (Crustacea: Thalassinidea)

    Science.gov (United States)

    Astall, C. M.; Taylor, A. C.; Atkinson, R. J. A.

    1997-02-01

    Upogebia stellataand U. deltaura(Crustacea: Thalassinidea) construct burrows in nearshore sediments in U.K. waters. Burrow structure is similar in both species; the basic burrow consisting of a two-opening, U-shaped section with a vertical shaft descending from the mid-point of the U. This structure may be variously elaborated. Burrow cross-section is circular, dilations allow turning by somersaulting and surface openings are often constricted. Conditions within the burrows are usually hypoxic and hypercapnic. Burrow water PO 2in the parts normally occupied by the mud-shrimp was between 80-110 Torr, but was much lower (10-45 Torr) in the deepest, poorly-irrigated parts. Both species irrigate their burrows by episodes of pleopod beating of variable duration (mean=8·5±3·5 min and 2·8±0·5 min for U. deltauraand U. stellata, respectively), which draws oxygenated water into the burrow and also particulate food for suspension feeding. When exposed to hypoxia, U. deltauraand U. stellatawere able to maintain their rates of oxygen consumption approximately constant over a wide range of PO 2( Pc=30-50 Torr). Under these conditions, there was a pronounced increase in scaphognathite beat rate but heart rate remained relatively constant. Below the Pc, however, both rates declined.

  9. Variations in the Foraging Behaviour and Burrow Structures of the Damara Molerat Cryptomys damarensis in the Kalahari Gemsbok National Park

    Directory of Open Access Journals (Sweden)

    B.G. Lovegrove

    1987-10-01

    Full Text Available Aspects of two habitat-specific foraging behaviours of the social subterranean rodent Cryptomys damarensis, are discussed in terms of burrow structure, resource dispersion patterns, sand moisture content, burrow temperature regimes, and predatory pressures, in the Kalahari Gemsbok National Park, South Africa.

  10. DENSITY-DEPENDENT IMPACTS OF BIOIRRIGATION BY THE BURROWING SHRIMP UPOGEBIA PUGETTENSIS ON BENTHIC FLUXES AND POREWATER SOLUTE DISTRIBUTIONS IN PACIFIC NORTHWEST ESTUARIES

    Science.gov (United States)

    Burrowing thalassinid shrimp are major ecosystem engineering species of Pacific estuaries and can structure the physical, chemical, and biotic properties of sediments. Feeding and burrow irrigation by benthic organisms can increase the remineralization rates of organic material (...

  11. Statement on Bills to Designate Crab Orchard Wilderness

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — Statement of John Kyl on H.R. 3508 and H.R. 5893, bills to designate Crab Orchard Wilderness within Crab Orchard National Wildlife Refuge. This document includes maps.

  12. Epibiotic community of the horseshoe crab Tachypleus gigas

    Digital Repository Service at National Institute of Oceanography (India)

    Patil, J.S.; Anil, A.C.

    -epibionts) present on horseshoe crabs, according to gender, were evaluated, and the macro-epibiont population from different regions of the carapace was mapped. In general, female horseshoe crabs harbored fewer epibionts than the males. Among the diatoms, Navicula...

  13. Association of the "IUCN vulnerable" spiny rat Clyomys bishopi (Rodentia: Echimyidae) with palm trees and armadillo burrows in southeastern Brazil.

    Science.gov (United States)

    Bueno, Adriana A; Lapenta, Marina J; Oliveira, Fátima; Motta-Junior, José C

    2004-12-01

    The globally vulnerable Clyomys bishopi, a semi-fossorial and colonial rodent, is apparently limited to cerrado (savannah-like vegetation) physiognomies in São Paulo State, Brazil. The aim of the study was to verify whether the presence of C. bishopi is associated to the occurrence of palm trees (Attalea gearensis, Syagrus loefgrenii) and armadillo burrows. Thirty six quadrats were placed in different physiognomies of cerrado vegetation at Itirapina Ecological Station, southeastern Brazil to survey the number of C. bishopi burrows of individuals of palm trees and burrows of armadillos. There was a strong dependence and association between the number of C. bishopi burrows and all measured variables (Contingency tables and Spearman rank correlations). It is suggested that this rodent can be found in great numbers where palm trees are abundant. The use of armadillo burrows possibly makes the movement of the rodents easier inside their own galleries.

  14. Nelson's big horn sheep (Ovis canadensis nelsoni) trample Agassiz's desert tortoise (Gopherus agassizii) burrow at a California wind energy facility

    Science.gov (United States)

    Agha, Mickey; Delaney, David F.; Lovich, Jeffrey E.; Briggs, Jessica; Austin, Meaghan; Price, Steven J.

    2015-01-01

    Research on interactions between Agassiz's desert tortoises (Gopherus agassizii) and ungulates has focused exclusively on the effects of livestock grazing on tortoises and their habitat (Oldemeyer, 1994). For example, during a 1980 study in San Bernardino County, California, 164 desert tortoise burrows were assessed for vulnerability to trampling by domestic sheep (Ovis aries). Herds of grazing sheep damaged 10% and destroyed 4% of the burrows (Nicholson and Humphreys 1981). In addition, a juvenile desert tortoise was trapped and an adult male was blocked from entering a burrow due to trampling by domestic sheep. Another study found that domestic cattle (Bos taurus) trampled active desert tortoise burrows and vegetation surrounding burrows (Avery and Neibergs 1997). Trampling also has negative impacts on diversity of vegetation and intershrub soil crusts in the desert southwest (Webb and Stielstra 1979). Trampling of important food plants and overgrazing has the potential to create competition between desert tortoises and domestic livestock (Berry 1978; Coombs 1979; Webb and Stielstra 1979).

  15. The Radio Spectral Index of the Crab Nebula

    Science.gov (United States)

    1997-11-20

    We present the results of a new, comprehensive investigation of the radio spectral index of the Crab Nebula supernova remnant. New data at 74 MHz are...thermal material in the Crab Nebula’s filaments. Apart from some possible renewed acceleration occurring in the wisps, the dominant accelerator of relativistic electrons in the Crab Nebula is the pulsar itself.

  16. Strong population genetic structure and larval dispersal capability of the burrowing ghost shrimp (Neotrypaea californiensis)

    Science.gov (United States)

    The burrowing ghost shrimp, Neotrypaea californiensis, is a vital member of the estuarine benthic community. Dense populations of shrimp are found in the major estuaries of Washington and Oregon. Our study determines the genetic structure of shrimp populations in order to gain ...

  17. SEASONAL FORAGING BY CHANNEL CATFISH ON TERRESTRIALLY BURROWING CRAYFISH IN A FLOODPLAIN-RIVER ECOSYSTEM

    Science.gov (United States)

    The seasonal use of terrestrially burrowing crayfish as a food item by channel catfish Ictalurus punctatus was studied in channelized and non-channelized sections of the Yockanookany River (Mississippi, USA). During seasonal inundation of the floodplains, the crayfish occupied o...

  18. Burrowing Behavior of a Deposit Feeding Bivalve Predicts Change in Intertidal Ecosystem State

    NARCIS (Netherlands)

    Compton, T.J.; Bodnar, W.; Koolhaas, A.; Dekinga, A.; Holthuijsen, S.; Ten Horn, J.; McSweeney, N.; van Gils, J.A.; Piersma, T,

    2016-01-01

    Behavior has a predictive power that is often underutilized as a tool for signaling ecological change. The burrowing behavior of the deposit feeding bivalve Macoma balthica reflects a typical food-safety trade-off. The choice to live close to the sediment surface comes at a risk of predation and is

  19. Burrowing behavior of a deposit feeding bivalve predicts change in intertidal ecosystem state

    NARCIS (Netherlands)

    Compton, Tanya J.; Bodnar, Wanda; Koolhaas, Anita; Dekinga, Anne; Holthuijsen, Sander; ten Horn, Job; McSweeney, Niamh; van Gils, Jan; Piersma, Theunis

    2016-01-01

    Behavior has a predictive power that is often underutilized as a tool for signaling ecological change. The burrowing behavior of the deposit feeding bivalve Macoma balthica reflects a typical food-safety trade-off. The choice to live close to the sediment surface comes at a risk of predation and is

  20. Gopherus agassizii (desert tortoise) and Crotalus ruber (red diamond rattlesnake). Burrow co-occupancy

    Science.gov (United States)

    Lovich, Jeffrey E.

    2011-01-01

    I observed an adult Desert Tortoise and an adult Red Diamond Rattlesnake (sexes unknown) in a shallow tortoise burrow on 6 January 1997 at a wind energy generation facility near Palm Springs, Riverside Co., California, USA (33.9599°N, 116.6613°W).

  1. The Potential Use of Electricity to Control Burrowing Shrimp in Oyster Aquaculture Beds

    Science.gov (United States)

    Thalassinid shrimp cause significant problems for oyster aquaculture in the Pacific Northwest (USA) where oysters succumb to the physical disruption of the sediment by the burrowing activity of these animals. While electrofishing is a commonly used technique to capture fish and some invertebrates i...

  2. Behaviour and time allocation of the burrowing shrimp Callianassa subterranea (Decapoda, Thalassinidea)

    NARCIS (Netherlands)

    Stamhuis, E.J; Reede-Dekker, T; van Etten, Y; de Wiljes, J.J.; Videler, J.J

    1996-01-01

    The behaviour and allocation of time of the endobenthic shrimp Callianassa subterranea from the central North Sea was studied in the laboratory. Animals were allowed to construct a two-dimensional burrow in large transparent sediment filled cuvettes tailored to their body width. The behaviour of the

  3. Lizard burrows provide thermal refugia for larks in the Arabian desert

    NARCIS (Netherlands)

    Williams, JB; Tieleman, BI; Shobrak, M

    1999-01-01

    A common perception is that desert birds experience greater extremes of heat and aridity than their mammalian counterparts, in part, because birds do not use burrows as a refuge from the desert environment. We report observations of Dunn's Larks (Eremalauda dunni), Bar-tailed Desert Larks (Ammomanes

  4. First implementation of burrowing motions in dual-reciprocating drilling using an integrated actuation mechanism

    Science.gov (United States)

    Pitcher, Craig; Gao, Yang

    2017-03-01

    The dual-reciprocating drill (DRD) is a biologically-inspired low-mass alternative to traditional drilling techniques, using backwards-facing teethed halves to grip the surrounding substrate, generating a traction force that reduces the required overhead penetration force. Previous experiments using a proof-of-concept test bench have provided evidence as to the significant role of sideways movements and lateral forces in improving drilling performance. The system is also progressing to a first system prototype concept, in which an actuation mechanism is integrated within the drill heads. To experimentally determine the effect of lateral motions, a new internal actuation mechanism was developed to allow the inclusion of controlled sideways movements, resulting in the creation of the circular and diagonal burrowing motions. This paper presents an investigation into the performance of the reciprocation and burrowing motions by testing them in a planetary regolith simulant. Analysis of force sensor measurements has shown a relationship between the penetration and traction forces and the internal friction of the mechanism and depth achieved. These tests have also experimentally demonstrated the benefit of lateral motions in drilling performance, with both the burrowing mechanisms and drilling tests performed at an angle able to penetrate further than traditional vertical reciprocation, leading to the proposition of new burrowing and diagonal drilling mechanics. From this, a new fully integrated system prototype can be developed which incorporates lateral motions that can optimise the drilling performance.

  5. Simultaneous collection of body temperature and activity data in burrowing mammals : a new technique

    NARCIS (Netherlands)

    Long, Ryan A.; Hut, Roelof A.; Barnes, Brian M.

    2007-01-01

    Integrating physiological and behavioral observations into ecological field studies of animals can provide novel insights into relationships among animal behavior, physiology, and ecology. We describe and evaluate a new technique for simultaneously collecting body temperature (T-b) and burrow use da

  6. Rabbits, refuges and resources : how foraging of herbivores is affected by living in burrows

    NARCIS (Netherlands)

    Dekker, J.J.A.

    2007-01-01

    Small herbivores such as rabbits, pika and marmots create spatial patterns in vegetation around their burrows by grazing. This PhD thesis focuses on these refuge-living herbivores.By performing experiments with rabbits, he showed that looking for predators causes the spatial patter

  7. Burrowing dragonfly larvae as biosentinels of methylmercury in freshwater food webs.

    Science.gov (United States)

    Haro, Roger J; Bailey, Sean W; Northwick, Reid M; Rolfhus, Kristofer R; Sandheinrich, Mark B; Wiener, James G

    2013-08-06

    We assessed the utility of larval burrowing dragonflies (Odonata: Anisoptera: Gomphidae) as biosentinels of methylmercury (MeHg) contamination. Gomphids were the most abundant family of dragonflies sampled during 2008-2010 from 17 lakes in four national parks of the northwestern Laurentian Great Lakes region. Ten species of burrowing gomphids were sampled; 13 lakes contained 3 or more species, and 2 species of Gomphus co-occurred in 12 lakes. Most of the total Hg (THg) in whole, late-instar larvae was MeHg, with mean percent MeHg exceeding 60% in 16 lakes. Mean MeHg in larvae of a given species varied greatly among lakes, ranging from 4 to 109 ng g(-1) dry weight. Methylmercury levels in larvae, however, were much less variable within a given lake and species. The mean concentration of MeHg in burrowing gomphids was positively correlated with mean MeHg concentration in unfiltered lake water. Mean concentrations of THg and MeHg in multispecies assemblages of Gomphus were also positively correlated with mean THg in coexisting prey fish and game fishes. We recommend-and provide guidance on-the application of burrowing gomphids as biosentinels of MeHg contamination, which can extend the bioassessment of MeHg to fishless fresh waters.

  8. Novel Geometries for the LHC Crab Cavity

    Energy Technology Data Exchange (ETDEWEB)

    B. Hall, G. Burt, C. Lingwood, R. Rimmer, H. Wang

    2010-05-23

    The planned luminosity upgrade to LHC is likely to necessitate a large crossing angle and a local crab crossing scheme. For this scheme crab cavities align bunches prior to collision. The scheme requires at least four such cavities, a pair on each beam line either side of the interaction point (IP). Upstream cavities initiate rotation and downstream cavities cancel rotation. Cancellation is usually done at a location where the optics has re-aligned the bunch. The beam line separation near the IP necessitates a more compact design than is possible with elliptical cavities such as those used at KEK. The reduction in size must be achieved without an increase in the operational frequency to maintain compatibility with the long bunch length of the LHC. This paper proposes a suitable superconducting variant of a four rod coaxial deflecting cavity (to be phased as a crab cavity), and presents analytical models and simulations of suitable designs.

  9. The effect of mayfly (Hexagenia spp.) burrowing activity on sediment oxygen demand in western Lake Erie

    Science.gov (United States)

    Edwards, William J.; Soster, Frederick M.; Matisoff, Gerald; Schloesser, Donald W.

    2009-01-01

    Previous studies support the hypothesis that large numbers of infaunal burrow-irrigating organisms in the western basin of Lake Erie may increase significantly the sediment oxygen demand, thus enhancing the rate of hypolimnetic oxygen depletion. We conducted laboratory experiments to quantify burrow oxygen dynamics and increased oxygen demand resulting from burrow irrigation using two different year classes of Hexagenia spp. nymphs from western Lake Erie during summer, 2006. Using oxygen microelectrodes and hot film anemometry, we simultaneously determined oxygen concentrations and burrow water flow velocities. Burrow oxygen depletion rates ranged from 21.7 mg/nymph/mo for 15 mm nymphs at 23 °C to 240.7 mg/nymph/mo for 23 mm nymphs at 13 °C. Sealed microcosm experiments demonstrated that mayflies increase the rate of oxygen depletion by 2-5 times that of controls, depending on size of nymph and water temperature, with colder waters having greater impact. At natural population densities, nymph pumping activity increased total sediment oxygen demand 0.3-2.5 times compared to sediments with no mayflies and accounted for 22-71% of the total sediment oxygen demand. Extrapolating laboratory results to the natural system suggest that Hexagenia spp. populations may exert a significant control on oxygen depletion during intermittent stratification. This finding may help explain some of the fluctuations in Hexagenia spp. population densities in western Lake Erie and suggests that mayflies, by causing their own population collapse irrespective of other environmental conditions, may need longer term averages when used as a bio-indicator of the success of pollution-abatement programs in western Lake Erie and possibly throughout the Great Lakes.

  10. Frequency Tuning for a DQW Crab Cavity

    CERN Document Server

    Verdú-Andrés, Silvia; Ben-Zvi, Ilan; Calaga, Rama; Capatina, Ofelia; Leuxe, Raphael; Skaritka, John; Wu, Qiong; Xiao, Binping; Zanoni, Carlo

    2016-01-01

    The nominal operating frequency for the HL-LHC crab cavities is 400.79 MHz within a bandwidth of ±60kHz. Attaining the required cavity tune implies a good understanding of all the processes that influence the cavity frequency from the moment when the cavity parts are being fabricated until the cavity is installed and under operation. Different tuning options will be available for the DQW crab cavity of LHC. This paper details the different steps in the cavity fabrication and preparation that may introduce a shift in the cavity frequency and introduces the different tuning methods foreseen to bring the cavity frequency to meet the specifications.

  11. Modeling Crabbing Dynamics in an Electron-Ion Collider

    Energy Technology Data Exchange (ETDEWEB)

    Castilla, Alejandro [Thomas Jefferson National Accelerator Facility, Newport News, VA (United States); Old Dominion Univ., Norfolk, VA (United States); Univ. de Guanajuato (DCI-UG), Leon (Mexico); Morozov, Vasiliy S. [Thomas Jefferson National Accelerator Facility, Newport News, VA (United States); Satogata, Todd J. [Thomas Jefferson National Accelerator Facility, Newport News, VA (United States); Old Dominion Univ., Norfolk, VA (United States); Delayen, Jean R. [Thomas Jefferson National Accelerator Facility, Newport News, VA (United States); Old Dominion Univ., Norfolk, VA (United States)

    2015-09-01

    A local crabbing scheme requires π/2 (mod π) horizontal betatron phase advances from an interaction point (IP) to the crab cavities on each side of it. However, realistic phase advances generated by sets of quadrupoles, or Final Focusing Blocks (FFB), between the crab cavities located in the expanded beam regions and the IP differ slightly from π/2. To understand the effect of crabbing on the beam dynamics in this case, a simple model of the optics of the Medium Energy Electron-Ion Collider (MEIC) including local crabbing was developed using linear matrices and then studied numerically over multiple turns (1000 passes) of both electron and proton bunches. The same model was applied to both local and global crabbing schemes to determine the linear-order dynamical effects of the synchro-betatron coupling induced by crabbing.

  12. Evolution of metabolomics profile of crab paste during fermentation.

    Science.gov (United States)

    Chen, Daian; Ye, Yangfang; Chen, Juanjuan; Yan, Xiaojun

    2016-02-01

    Crab paste is regularly consumed by people in the coastal area of China. The fermentation time plays a key role on the quality of crab paste. Here, we investigated the dynamic evolution of metabolite profile of crab paste during fermentation by combined use of NMR spectroscopy and multivariate data analysis. Our results showed that crab paste quality was significantly affected by fermentation. The quality change was manifested in the decline of lactate, betaine, taurine, trimethylamine-N-oxide, trigonelline, inosine, adenosine diphosphate, and 2-pyridinemethanol, and in the fluctuation of a range of amino acids as well as in the accumulation of glutamate, sucrose, formate, acetate, trimethylamine, and hypoxanthine. Trimethylamine production and its increased level with fermentation could be considered as a freshness index of crab paste. These results contribute to quality assessment of crab paste and confirm the metabolomics technique as a useful tool to provide important information on the crab paste quality.

  13. Treatment of black-tailed prairie dog burrows with deltamethrin to control fleas (Insecta: Siphonaptera) and plague.

    Science.gov (United States)

    Seery, David B; Biggins, Dean E; Montenieri, John A; Enscore, Russell E; Tanda, Dale T; Gage, Kenneth L

    2003-09-01

    Burrows within black-tailed prairie dog (Cynomys ludovicianus) colonies on the Rocky Mountain Arsenal National Wildlife Refuge, Colorado, were dusted with deltamethrin insecticide to reduce flea (Insecta: Siphonaptera) abundance. Flea populations were monitored pre- and posttreatment by combing prairie dogs and collecting fleas from burrows. A single application of deltamethrin significantly reduced populations of the plague vector Oropsylla hirsuta, and other flea species on prairie dogs and in prairie dog burrows for at least 84 d. A plague epizootic on the Rocky Mountain Arsenal National Wildlife Refuge caused high mortality of prairie dogs on some untreated colonies, but did not appear to affect nearby colonies dusted with deltamethrin.

  14. A Comparative Study of Genetic Variation betwccn Chinese Mitten Crab Eriocheir Sinensis and Hepu Mitten Crab E. Hepuensis

    Institute of Scientific and Technical Information of China (English)

    LIU Guangdong; ZHANG Xiumei; GAO Tianxiang; LOU Dong

    2002-01-01

    Horizontal starch gel electrophoresis was used to investigate the biochemical genetic structure of Chinese mittencrab Eriocheir sinensis and Hepu mitten crab E. hepuensis. Sixteen putative enzyme-coding loci were examined for forty-nineChinese mitten crabs and thirty-eight Hepu mitten crabs. Nine loci, AAT-1 *, AAT-2 *, G3PDH*, GPI*, IDHP-1 *, IDHP-2 *,MDH-1 *, MDH-2 * and PGM * , are polymorphic in Chinese mitten crab, and seven, AAT-1 *, AAT-2 *, GPI *, IDHP-1 *,MDH-1 *, MDH-2 * and PGM * , are polymorphic in Hepu mitten crab. The proportion of polymorphic loci and the expectedheterozygosity are 0.562 5 and 0.080 3 for Chinese mitten crab, and 0.437 5 and 0.075 4 for Hepu mitten crab. The Nei' s ge-netic distance between two species is 0.002 4.

  15. Corneal laceration caused by river crab

    Directory of Open Access Journals (Sweden)

    Vinuthinee N

    2015-01-01

    Full Text Available Naidu Vinuthinee,1,2 Anuar Azreen-Redzal,1 Jaafar Juanarita,1 Embong Zunaina2 1Department of Ophthalmology, Hospital Sultanah Bahiyah, Alor Setar, 2Department of Ophthalmology, School of Medical Sciences, Universiti Sains Malaysia, Kubang Kerian, Malaysia Abstract: A 5-year-old boy presented with right eye pain associated with tearing and photophobia of 1-day duration. He gave a history of playing with a river crab when suddenly the crab clamped his fingers. He attempted to fling the crab off, but the crab flew and hit his right eye. Ocular examination revealed a right eye corneal ulcer with clumps of fibrin located beneath the corneal ulcer and 1.6 mm level of hypopyon. At presentation, the Seidel test was negative, with a deep anterior chamber. Culture from the corneal scrapping specimen grew Citrobacter diversus and Proteus vulgaris, and the boy was treated with topical gentamicin and ceftazidime eyedrops. Fibrin clumps beneath the corneal ulcer subsequently dislodged, and revealed a full-thickness corneal laceration wound with a positive Seidel test and shallow anterior chamber. The patient underwent emergency corneal toileting and suturing. Postoperatively, he was treated with oral ciprofloxacin 250 mg 12-hourly for 1 week, topical gentamicin, ceftazidime, and dexamethasone eyedrops for 4 weeks. Right eye vision improved to 6/9 and 6/6 with pinhole at the 2-week follow-up following corneal suture removal. Keywords: corneal ulcer, pediatric trauma, ocular injury

  16. Ocean acidification impairs crab foraging behaviour.

    Science.gov (United States)

    Dodd, Luke F; Grabowski, Jonathan H; Piehler, Michael F; Westfield, Isaac; Ries, Justin B

    2015-07-07

    Anthropogenic elevation of atmospheric CO2 is driving global-scale ocean acidification, which consequently influences calcification rates of many marine invertebrates and potentially alters their susceptibility to predation. Ocean acidification may also impair an organism's ability to process environmental and biological cues. These counteracting impacts make it challenging to predict how acidification will alter species interactions and community structure. To examine effects of acidification on consumptive and behavioural interactions between mud crabs (Panopeus herbstii) and oysters (Crassostrea virginica), oysters were reared with and without caged crabs for 71 days at three pCO2 levels. During subsequent predation trials, acidification reduced prey consumption, handling time and duration of unsuccessful predation attempt. These negative effects of ocean acidification on crab foraging behaviour more than offset any benefit to crabs resulting from a reduction in the net rate of oyster calcification. These findings reveal that efforts to evaluate how acidification will alter marine food webs should include quantifying impacts on both calcification rates and animal behaviour.

  17. HUBBLE CAPTURES DYNAMICS OF CRAB NEBULA

    Science.gov (United States)

    2002-01-01

    A new sequence of Hubble Space Telescope images of the remnant of a tremendous stellar explosion is giving astronomers a remarkable look at the dynamic relationship between the tiny Crab Pulsar and the vast nebula that it powers. This picture shows a Hubble Space Telescope image of the inner parts of the Crab. The pulsar itself is visible as the left of the pair of stars near the center of the frame. Surrounding the pulsar is a complex of sharp knots and wisp-like features. This image is one of a sequence of Hubble images taken over the course of several months. This sequence shows that the inner part of the Crab Nebula is far more dynamic than previously understood. The Crab literally 'changes it stripes' every few days as these wisps stream away from the pulsar at half the speed of light. The Hubble Space Telescope photo was taken Nov. 5, 1995 by the Wide Field and Planetary Camera 2 at a wavelength of around 550 nanometers, in the middle of the visible part of the electromagnetic spectrum. Credit: Jeff Hester and Paul Scowen (Arizona State University), and NASA

  18. Swimming of the pea crab (Pinnotheres pisum)

    NARCIS (Netherlands)

    Versteegh, C.P.C.; Muller, M.

    2014-01-01

    Aquatic organisms have to deal with different hydrodynamic regimes, depending on their size and speed during locomotion. The pea crab swims by beating the third and fourth pereiopod on opposite sides as pairs. Using particle tracking velocimetry and high-speed video recording, we quantify the kinema

  19. The tree-climbing crabs of Trinidad

    NARCIS (Netherlands)

    Hagen, von Heinrich-Otto

    1977-01-01

    An annotated list of the brachyuran (12) and anomuran (1) tree-climbing crabs of Trinidad (West Indies) is presented (see Table 1 for species names). Some of the species mentioned (e.g. Aratus pisonii, Goniopsis cruentata) are well-known treeclimbers, in others (e.g. Sesarma roberti, S. ricordi) thi

  20. The CMS Remote Analysis Builder (CRAB)

    Energy Technology Data Exchange (ETDEWEB)

    Spiga, D.; Cinquilli, M.; Servoli, L.; /INFN, Perugia /Perugia U; Lacaprara, S.; Fanzago, F.; Dorigo, A.; /INFN, Padova; Merlo, M.; Farina, F.; /INFN, Milan /Milan U; Fanfani, A.; Codispoti, G.; Bacchi, W.; /INFN, Bologna /Bologna U /CERN /INFN, CNAF /INFN, Trieste /Fermilab

    2008-01-22

    The CMS experiment will produce several Pbytes of data every year, to be distributed over many computing centers geographically distributed in different countries. Analysis of this data will be also performed in a distributed way, using grid infrastructure. CRAB (CMS Remote Analysis Builder) is a specific tool, designed and developed by the CMS collaboration, that allows a transparent access to distributed data to end physicist. Very limited knowledge of underlying technicalities are required to the user. CRAB interacts with the local user environment, the CMS Data Management services and with the Grid middleware. It is able to use WLCG, gLite and OSG middleware. CRAB has been in production and in routine use by end-users since Spring 2004. It has been extensively used in studies to prepare the Physics Technical Design Report (PTDR) and in the analysis of reconstructed event samples generated during the Computing Software and Analysis Challenge (CSA06). This involved generating thousands of jobs per day at peak rates. In this paper we discuss the current implementation of CRAB, the experience with using it in production and the plans to improve it in the immediate future.

  1. Power coupler for the ILC crab cavity

    Energy Technology Data Exchange (ETDEWEB)

    Burt, G.; Dexter, A.; Jenkins, R.; /Lancaster U.; Beard, C.; Goudket, P.; McIntosh, P.A.; /Daresbury; Bellantoni, Leo; /Fermilab

    2007-06-01

    The ILC crab cavity will require the design of an appropriate power coupler. The beam-loading in dipole mode cavities is considerably more variable than accelerating cavities, hence simulations have been performed to establish the required external Q. Simulations of a suitable coupler were then performed and were verified using a normal conducting prototype with variable coupler tips.

  2. Horseshoe crab behavior:Patterns and processes

    Institute of Scientific and Technical Information of China (English)

    Christopher C.Chabot; Winsor H.Watson Ⅲ

    2010-01-01

    @@ 1 Introduction The American horseshoe crab Limulus polyphemus has long served as a source of delight and inspiration to a broad cross-section of scientists - from naturalists to neuroscientists. Periodically, for the last hundred years or so, new discoveries have been made about this ancient creature that both enlighten scientists and reinforce the importance of conserving this ancient species.

  3. 1994 prairie dog burrow mapping with Global Positioning Systems, UL Bend National Wildlife Refuge, Phillips County, Montana

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — Holes in the ground are a very important feature of black-footed ferret habitat. The number, distribution, and activity level of prairie dog burrows is related to...

  4. Burrowing nematodes from Colombia and their relationship with Radopholus similis populations,R. arabocoffeae and R. duriophilus

    NARCIS (Netherlands)

    Múnera Uribe, G.E.; Bert, W.; Vierstraete, A.R.; Peña, de la E.; Moens, M.; Decraemer, W.

    2010-01-01

    Two burrowing nematode populations from Colombia were characterised using morphological, morphometric and molecular criteria. The morphological and morphometric characters of the two populations did not differ from those of Radopholus similis. The phylogenetic analysis based on sequence comparison o

  5. Black-footed ferrets and recreational shooting influence the attributes of black-tailed prairie dog burrows

    Science.gov (United States)

    Biggins, Dean E.; Ramakrishnan, Shantini; Goldberg, Amanda R.; Eads, David A.

    2012-01-01

    Black-tailed prairie dogs (Cynomys ludovicianus) plug burrows occupied by black-footed ferrets (Mustela nigripes), and they also plug burrows to entomb dead prairie dogs. We further evaluated these phenomena by sampling connectivity and plugging of burrow openings on prairie dog colonies occupied by ferrets, colonies where recreational shooting was allowed, and colonies with neither shooting nor ferrets. We counted burrow openings on line surveys and within plots, classified surface plugging, and used an air blower to examine subsurface connectivity. Colonies with ferrets had lower densities of openings, fewer connected openings (suggesting increased subsurface plugging), and more surface plugs compared to colonies with no known ferrets. Colonies with recreational shooting had the lowest densities of burrow openings, and line-survey data suggested colonies with shooting had intermediate rates of surface plugging. The extent of surface and subsurface plugging could have consequences for the prairie dog community by changing air circulation and escape routes of burrow systems and by altering energetic relationships. Burrow plugging might reduce prairie dogs' risk of predation by ferrets while increasing risk of predation by American badgers (Taxidea taxus); however, the complexity of the trade-off is increased if plugging increases the risk of predation on ferrets by badgers. Prairie dogs expend more energy plugging and digging when ferrets or shooting are present, and ferrets increase their energy expenditures when they dig to remove those plugs. Microclimatic differences in plugged burrow systems may play a role in flea ecology and persistence of the flea-borne bacterium that causes plague (Yersinia pestis).

  6. Map-Based Repowering and Reorganization of a Wind Resource Area to Minimize Burrowing Owl and Other Bird Fatalities

    OpenAIRE

    Lee Neher; K. Shawn Smallwood; Bell, Douglas A.

    2009-01-01

    Wind turbines in the Altamont Pass Wind Resource Area (Alameda/Contra Costa Counties, California, USA) generate about 730 GWh of electricity annually, but have been killing thousands of birds each year, including >2,000 raptors and hundreds of burrowing owls. We have developed collision hazard maps and hazard ratings of wind turbines to guide relocation of existing wind turbines and careful repowering to modern turbines to reduce burrowing owl fatalities principally, and other birds second...

  7. Standardized Monitoring Strategies for Burrowing Owls on DoD Installations

    Science.gov (United States)

    2006-12-01

    whether each owl was detected visually, vocally , or both. Each adult owl detected at a survey point gets its own line, juveniles associated with one...measurements (tail length, meta-tarsus, wing chord ) are made on adult birds that allow comparison of body size between sexes and sites. Age...Burrowing Owls will have a brood patch. Adult Left Wing 180 Use the method shown in Appendix II to measure the owl’s left wing chord length. Record

  8. Long-term population dynamics of a managed burrowing owl colony

    Science.gov (United States)

    Barclay, John H.; Korfanta, Nicole M.; Kauffman, Matthew J.

    2011-01-01

    We analyzed the population dynamics of a burrowing owl (Athene cunicularia) colony at Mineta San Jose International Airport in San Jose, California, USA from 1990-2007. This colony was managed by using artificial burrows to reduce the occurrence of nesting owls along runways and within major airport improvement projects during the study period. We estimated annual reproduction in natural and artificial burrows and age-specific survival rates with mark-recapture techniques, and we estimated the relative contribution of these vital rates to population dynamics using a life table response experiment. The breeding colony showed 2 distinct periods of change: high population growth from 7 nesting pairs in 1991 to 40 pairs in 2002 and population decline to 17 pairs in 2007. Reproduction was highly variable: annual nesting success (pairs that raised =1 young) averaged 79% and ranged from 36% to 100%, whereas fecundity averaged 3.36 juveniles/pair and ranged from 1.43 juveniles/pair to 4.54 juveniles/pair. We estimated annual adult survival at 0.710 during the period of colony increase from 1996 to 2001 and 0.465 during decline from 2002 to 2007, but there was no change in annual survival of juveniles between the 2 time periods. Long-term population growth rate (lambda) estimated from average vital rates was lambdaa=1.072 with lambdai=1.288 during colony increase and lambdad=0.921 (DELTA lambda=0.368) during decline. A life table response experiment showed that change in adult survival rate during increasing and declining phases explained more than twice the variation in growth rate than other vital rates. Our findings suggest that management and conservation of declining burrowing owl populations should address factors that influence adult survival.

  9. Razor clam to RoboClam: burrowing drag reduction mechanisms and their robotic adaptation.

    Science.gov (United States)

    Winter, A G; V; Deits, R L H; Dorsch, D S; Slocum, A H; Hosoi, A E

    2014-09-01

    Estimates based on the strength, size, and shape of the Atlantic razor clam (Ensis directus) indicate that the animal's burrow depth should be physically limited to a few centimeters; yet razor clams can dig as deep as 70 cm. By measuring soil deformations around burrowing E. directus, we have found the animal reduces drag by contracting its valves to initially fail, and then fluidize, the surrounding substrate. The characteristic contraction time to achieve fluidization can be calculated directly from soil properties. The geometry of the fluidized zone is dictated by two commonly-measured geotechnical parameters: coefficient of lateral earth pressure and friction angle. Calculations using full ranges for both parameters indicate that the fluidized zone is a local effect, occurring between 1-5 body radii away from the animal. The energy associated with motion through fluidized substrate-characterized by a depth-independent density and viscosity-scales linearly with depth. In contrast, moving through static soil requires energy that scales with depth squared. For E. directus, this translates to a 10X reduction in the energy required to reach observed burrow depths. For engineers, localized fluidization offers a mechanically simple and purely kinematic method to dramatically reduce energy costs associated with digging. This concept is demonstrated with RoboClam, an E. directus-inspired robot. Using a genetic algorithm to find optimal digging kinematics, RoboClam has achieved localized fluidization burrowing performance comparable to that of the animal, with a linear energy-depth relationship, in both idealized granular glass beads and E. directus' native cohesive mudflat habitat.

  10. Trophic eggs compensate for poor offspring feeding capacity in a subsocial burrower bug

    OpenAIRE

    Baba, Narumi; Hironaka, Mantaro; Hosokawa, Takahiro; Mukai, Hiromi; Nomakuchi, Shintaro; Ueno, Takatoshi

    2010-01-01

    Various animals produce inviable eggs or egg-like structures called trophic eggs, which are presumed to be an extended maternal investment for the offspring. However, there is little knowledge about the ecological or physiological constraints associated with their evolutionary origin. Trophic eggs of the seminivorous subsocial burrower bug (Canthophorus niveimarginatus) have some unique characteristics. Trophic eggs are obligate for nymphal survival, and first-instar nymphs die without them. ...

  11. Burrowing, byssus, and biomarkers: behavioral and physiological indicators of sublethal thermal stress in freshwater mussels (Unionidae)

    Science.gov (United States)

    Archambault, Jennifer M.; Cope, W. Gregory; Kwak, Thomas J.

    2013-01-01

    Recent research has elucidated the acute lethal effects of elevated water temperatures to glochidia (larvae), juvenile, and adult life stages of freshwater mussels (Order Unionida), but few studies have focused on sublethal effects of thermal stress. We evaluated the sublethal effects of elevated temperature on burrowing behavior and byssus production in juveniles, and on enzymatic biomarkers of stress in adults in acute (96 h) laboratory experiments in sediment, with two acclimation temperatures (22 and 27 °C) and two experimental water levels (watered and dewatered) as proxies for flow regime. Increasing temperature significantly reduced burrowing in all five species tested, and the dewatered treatment (a proxy for drought conditions) reduced burrowing in all but Amblema plicata. Production of byssal threads was affected most drastically by flow regime, with the probability of byssus presence reduced by 93–99% in the dewatered treatment, compared to the watered treatment (a proxy for low flow conditions); increasing temperature alone reduced byssus by 18–35%. Alanine aminotransferase and aspartate aminotransferase were significantly affected by treatment temperature in the 27 °C acclimation, watered test (p = 0.04 and 0.02, respectively). Our results are important in the context of climate change, because stream temperature and flow are expected to change with increasing air temperature and altered precipitation patterns.

  12. Effects of invasive rats and burrowing seabirds on seeds and seedlings on New Zealand islands.

    Science.gov (United States)

    Grant-Hoffman, Madeline N; Mulder, Christa P H; Bellingham, Peter J

    2010-04-01

    Rats (Rattus rattus, Rattus norvegicus, Rattus exulans) are important invaders on islands. They alter vegetation indirectly by preying on burrowing seabirds. These seabirds affect vegetation through nutrient inputs from sea to land and physical disturbance through trampling and burrowing. Rats also directly affect vegetation though consumption of seeds and seedlings. Seedling communities on northern New Zealand islands differ in composition and densities among islands which have never been invaded by rats, are currently invaded by rats, or from which rats have been eradicated. We conducted experimental investigations to determine the mechanisms driving these patterns. When the physical disturbance of seabirds was removed, in soils collected from islands and inside exclosures, seedling densities increased with seabird burrow density. For example, seedling densities inside exclosures were 10 times greater than those outside. Thus the negative effects of seabirds on seedlings, by trampling and uprooting, overwhelm the potentially beneficial effects of high levels of seed germination, seedling emergence, and possibly seed production, which result from seed burial and nutrient additions. Potential seedling density was reduced on an island where rats were present, germination of seeds from soils of this island was approximately half that found on other islands, but on this island seedling density inside exclosures was 7 times the density outside. Although the total negative effects of seabirds and rats on seedling densities are similar (reduced seedling density), the differences in mechanisms and life stages affected result in very different filters on the plant community.

  13. Distribution of the Chuckwalla, Western Burrowing Owl, and Six Bat Species on the Nevada Test Site

    Energy Technology Data Exchange (ETDEWEB)

    Cathy A. Willis

    1997-05-01

    Field Surveys were conducted in 1996 to determine the current distribution of several animal species of concern on the Nevada Test Site (NTS). They included the chuckwall (Sauromalus obesus), western burrowing owl (Speotyto cunicularia), and six species of bats. Nineteen chuckwallas and 118 scat locations were found during the chuckwalla field study. Eighteen western burrowing owls were found at 12 sighting locations during the 1996 field study. Of the eleven bat species of concern which might occur on the NTS, five, and possibly six, were captured during this survey. The U.S. Department of Energy, Nevada Operations Office, takes certain management actions to protect and conserve the chuckwalla, western burrowing owl, and bats on the NTS. These actions are described and include: (1) conducting surveys at sites of proposed land-disturbing activities (2) altering projects whenever possible to avoid or minimize impacts to these species (3) maintaining a geospatial database of known habitat for species of concern (4) sharing sighting and trap location data gathered on the NTS with other local land and resource managers, and (5) conducting periodic field surveys to monitor these species distribution and relative abundance on the NTS.

  14. Bioassessment of contaminant transport and distribution in aquatic ecosystems by chemical analysis of burrowing mayflies (Hexagenia)

    Science.gov (United States)

    Steingraeber, M.T.; Wiener, J.G.

    1995-01-01

    Burrowing mayfly nymphs (Ephemeroptera) inhabit and ingest fine-grained sediments and detritus that may be enriched with metals and persistent organic compounds. The burrowing nymphs can externally adsorb and internally assimilate these contaminants, providing a link for the food chain transfer of potentially toxic substances from sediments to organisms in higher trophic levels. The emergent adults are short-lived and do not feed, thus their gut contents do not contribute greatly to their total contaminant burden. These characteristics make Hexagenia spp. And certain other burrowing mayflies useful for assessing ecosystem contamination. General protocols are presented for the collection, processing and analysis of emergent mayflies to assess the spatial distribution and bioaccumulation of sediment-associated contaminants in aquatic ecosystems. Two essential components of this bioassessment approach are a network of on-site volunteers with the materials and instructions needed to correctly collect and store samples and quality assurance procedures to estimate the accuracy of chemical analyses. The utility of this approach is demonstrated with an example of its application to the Upper Mississippi River (USA). Determination of cadmium, mercury and polychlorinated biphenyl congeners in emergent Hexagenia bilineata from a 1250 km reach of this river revealed (1) several source areas of contaminants and (2) distinct patterns in the bioaccumulation (and apparent sediment-associated transport) of each residue on both small and large spatial scales.

  15. Species Profiles. Life Histories and Environmental Requirements of Coastal Fishes and Invertebrates (Pacific Southwest). Brown Rock Crab, Red Rock Crab, and Yellow Crab

    Science.gov (United States)

    1989-12-01

    distinguished by a slender abdomen and Two other large Cancer species, the Dungeness crab mature females by a broad abdomen that is often hirsute (C. magister...to the scent experimental treatment . One 13th instar female crab of potential food in the water (Case 1964; Zimmer-Faust reared at 22 ’C became

  16. Accumulation and depuration of pectenotoxins in brown crab Cancer pagurus

    Institute of Scientific and Technical Information of China (English)

    LI Zhaoxin

    2009-01-01

    Pectenotoxins (PTXs) are a group of marine algal toxins. In this study, the accumulation and depuration of pectenotoxins in brown crab Cancer pagurus were investigated. Crabs were fed with toxic blue mussels Mytilus edulis for 21 days and then depurated for 42 days. Toxins were extracted with methanol from the digestive glands of contaminated crabs, uncontaminated crabs (control group) and from blue mussels for comparison. Extracts were analyzed by liquid chromatograph coupled with tandem mass spectrometry (LC-MS-MS). The concentrations of PTX-2, PTX-2 SA, 7-epi-PTX-2 SA, and PTX-12 were analyzed in two batches of toxic blue mussels and the crabs. A one-compartment model was applied to describe the depuration of PTXs. The half-life of PTXs was estimated to be 6-7.5 days. After depuration for 42 days, the amount of PTXs measured in the crab digestive glands was less than 1 μg/kg.

  17. Predatory blue crabs induce stronger nonconsumptive effects in eastern oysters Crassostrea virginica than scavenging blue crabs

    Directory of Open Access Journals (Sweden)

    Avery E. Scherer

    2017-02-01

    Full Text Available By influencing critical prey traits such as foraging or habitat selection, predators can affect entire ecosystems, but the nature of cues that trigger prey reactions to predators are not well understood. Predators may scavenge to supplement their energetic needs and scavenging frequency may vary among individuals within a species due to preferences and prey availability. Yet prey reactions to consumers that are primarily scavengers versus those that are active foragers have not been investigated, even though variation in prey reactions to scavengers or predators might influence cascading nonconsumptive effects in food webs. Oysters Crassostrea virginica react to crab predators by growing stronger shells. We exposed oysters to exudates from crabs fed live oysters or fed aged oyster tissue to simulate scavenging, and to controls without crab cues. Oysters grew stronger shells when exposed to either crab exudate, but their shells were significantly stronger when crabs were fed live oysters. The stronger response to predators than scavengers could be due to inherent differences in diet cues representative of reduced risk in the presence of scavengers or to degradation of conspecific alarm cues in aged treatments, which may mask risk from potential predators subsisting by scavenging.

  18. Efeito do jejum e da realimentação sobre o metabolismo de aminoácidos no músculo e no hepatopâncreas do caranguejo Neohelice granulata submetido previamente à dieta rica em proteínas ou carboidratos

    OpenAIRE

    2011-01-01

    O objetivo deste trabalho foi investigar, in vitro, os efeitos do jejum de 15 dias e da subsequente realimentação por 72, 96 e 120 horas, sobre o metabolismo de aminoácidos no hepatopâncreas e no músculo do caranguejo Neohelice granulata previamente alimentado com uma dieta rica em carboidratos (HC) ou proteínas (HP). Para isso, foram realizados os seguintes procedimentos experimentais - captação de [U14C]-MeAIB (ácido metil-aminoisobutírico); síntese de 14Cproteínas a partir de [U14C]-leucin...

  19. A 150-million-year-old crab larva and its implications for the early rise of brachyuran crabs.

    Science.gov (United States)

    Haug, Joachim T; Martin, Joel W; Haug, Carolin

    2015-03-09

    True crabs (Brachyura) are the most successful group of decapod crustaceans. This success is most likely coupled to their life history, including two specialised larval forms, zoea and megalopa. The group is comparably young, starting to diversify only about 100 million years ago (mya), with a dramatic increase in species richness beginning approximately 50 mya. Early evolution of crabs is still very incompletely known. Here, we report a fossil crab larva, 150 mya, documented with up-to-date imaging techniques. It is only the second find of any fossil crab larva, but the first complete one, the first megalopa, and the oldest one (other fossil ca. 110 mya). Despite its age, the new fossil possesses a very modern morphology, being indistinguishable from many extant crab larvae. Hence, modern morphologies must have been present significantly earlier than formerly anticipated. We briefly discuss the impact of this find on our understanding of early crab evolution.

  20. Bacterial septicaemia in prerecruit edible crabs, Cancer pagurus L.

    Science.gov (United States)

    Smith, A L; Whitten, M M A; Hirschle, L; Pope, E C; Wootton, E C; Vogan, C L; Rowley, A F

    2014-08-01

    Juvenile edible crabs, Cancer pagurus L., were surveyed from Mumbles Head and Oxwich Bay in South Wales, UK, and the number of heterotrophic bacteria and vibrios in the hemolymph was determined. The percentage of crabs with hemolymph containing bacteria was variable over the survey with higher numbers of animals affected in summer than in winter. Post-moult crabs contained significantly higher numbers of heterotrophic bacteria in the hemolymph than pre- and intermoult animals. Crabs with cuticular damage to the gills also had significantly higher numbers of bacteria in the hemolymph. Crabs were found to have a high prevalence of infection by the dinoflagellate, Hematodinium. Such animals had significantly fewer bacteria in the blood in comparison with Hematodinium-free animals. Of the 463 crabs surveyed, only 3 individuals had hemolymph containing 2000 + CFU mL(-1). Based on 16S rRNA gene sequences, two of these crabs contained a Vibrio pectenicida-like isolate, while the other had a mixed assemblage of vibrios. Although 59% of the crabs surveyed had culturable bacteria in the hemolymph, the majority only had small numbers (crab fishery in this region.

  1. The importance of dietary calcium consumption in two species of semi-terrestrial grapsoid crabs A importância do consumo de cálcio na dieta de duas espécies de caranguejos grapsóides semi-terrestres

    Directory of Open Access Journals (Sweden)

    Flavia P. Zanotto

    2009-09-01

    Full Text Available Calcium (Ca is essential for crustaceans, due to calcium carbonate (CaCO3 deposition in the new exoskeleton to harden it. The purpose of this work was to study short term Ca balance in terms of dietary Ca ingestion in two phylogenetically related crabs (Superfamily Grapsoidea showing different degrees of terrestrial adaptations: Sesarma rectum Randall, 1840 and Neohelice granulata (Dana, 1851. Dietary Ca ingestion was studied using purified diets with different Ca concentrations (0, 2.2 and 6.66 % Ca, together with measurements of Ca excretion and Ca hemolymph levels. The results showed that both crabs had the same response to foods containing different levels of Ca, with both species eating more of the high Ca diet. However, S. rectum consumed more per mg body mass at all Ca concentrations (6 mg.g-1 for S. rectum against 3 mg.g-1 for N. granulata. Both species excreted/egested Ca differently: S. rectum excreted Ca proportionally to ingestion, whereas N. granulata maintained constant faecal Ca output at all dietary Ca levels. Moreover, Ca hemolymph levels for crabs fed the different diets were independent of dietary Ca. In conclusion, both S. rectum and N. granulata seem to regulate the consumption of diets containing more Ca, which suggests a fine balance for Ca intake.O cálcio (Ca é essencial para os crustáceos porque cristais de carbonato de cálcio (CaCO3 são depositados no novo exoesqueleto para endurecê-lo. O objetivo do presente trabalho foi estudar o balanço do Ca em relação à sua ingestão em dois caranguejos filogenéticamente relacionados (Superfamília Grapsoidea, que apresentam diferentes graus de terrestrialidade: Sesarma rectum Randall, 1840 e Neohelice granulata (Dana, 1851. A ingestão de Ca foi estudada através do uso de dietas purificadas com diferentes quantidades de Ca (0, 2,2 e 6,66 % Ca, juntamente com a excreção de Ca nas fezes e níveis de Ca na hemolinfa. Os resultados mostraram que ambos apresentam a mesma

  2. FACT. Energy spectrum of the Crab Nebula

    Energy Technology Data Exchange (ETDEWEB)

    Temme, Fabian; Einecke, Sabrina; Buss, Jens [TU Dortmund, Experimental Physics 5, Otto-Hahn-Str.4, 44221 Dortmund (Germany); Collaboration: FACT-Collaboration

    2016-07-01

    The First G-APD Cherenkov Telescope is the first Imaging Air Cherenkov Telescope which uses silicon photon detectors (G-APDs aka SiPM) as photo sensors. With more than four years of operation, FACT proved an application of SiPMs is suitable for the field of ground-based gamma-ray astronomy. Due to the stable flux at TeV energies, the Crab Nebula is handled as a ''standard candle'' in Cherenkov astronomy. The analysis of its energy spectrum and comparison with other experiments, allows to evaluate the performance of FACT. A modern analysis chain, based on data stream handling and multivariate analysis methods was developed in close cooperation with the department of computer science at the TU Dortmund. In this talk, this analysis chain and its application are presented. Further to this, results, including the energy spectrum of the Crab Nebula, measured with FACT, are shown.

  3. Pulse profile stability of the Crab pulsar

    CERN Document Server

    Jain, Chetana

    2011-01-01

    We present an X-ray timing analysis of the Crab pulsar, PSR B0531+21, using the archival RXTE data. We have investigated the stability of the Crab pulse profile, in soft (2-20 keV) and hard (30-100 keV) X-ray energies, over the last decade of RXTE operation. The analysis includes measurement of the separation between the two pulse peaks; and intensity and the widths of the two peaks. We did not find any significant time dependency in the pulse shape. The two peaks are stable in phase, intensity and widths, for the last ten years. The first pulse is relatively stronger at soft X-rays. The first pulse peak is narrower than the second peak, in both, soft- and hard X-ray energies. Both the peaks show a slow rise and a steeper fall. The ratio of the pulsed photons in the two peaks is also constant in time.

  4. Multiband observations of the Crab Nebula

    Science.gov (United States)

    Krassilchtchikov, A. M.; Bykov, A. M.; Castelletti, G. M.; Dubner, G. M.; Kargaltsev, O. Yu; Pavlov, G. G.

    2017-01-01

    Results of simultaneous imaging of the Crab Nebula in the radio (JVLA), optical (HST), and X-ray (Chandra) bands are presented. The images show a variety of small-scale structures, including wisps mainly located to the north-west of the pulsar and knots forming a ring-like structure associated with the termination shock of the pulsar wind. The locations of the structures in different bands do not coincide with each other.

  5. Pulsating Radio Sources near the Crab Nebula.

    Science.gov (United States)

    Staelin, D H; Reifenstein, E C

    1968-12-27

    Two new pulsating radio sources, designated NP 0527 and NP 0532, were found near the Crab Nebula and could be coincident with it. Both sources are sporadic, and no periodicities are evident. The pulse dispersions indicate that 1.58 +/- 0.03 and 1.74 +/- 0.02 x 10(20) electrons per square centimeter lie in the direction of NP 0527 and NP 0532, respectively.

  6. A Large Glitch in the Crab Pulsar

    Institute of Scientific and Technical Information of China (English)

    2001-01-01

    Using a new pulsar timing system at the 25-m radio telescope of Urumqi Astronomical Observatory, we have detected a large glitch in the Crab pulsar which occurred in 2000 July. The size of the gfitch is Av/v ~ 2.4 × 10-8, with a rela tive increment in frequency derivative Av/v ~ 5 × 10-3. The observing system is introduced and the observed properties of the glitch are discussed.

  7. The inner knot of the Crab nebula

    Science.gov (United States)

    Lyutikov, Maxim; Komissarov, Serguei S.; Porth, Oliver

    2016-02-01

    We model the inner knot of the Crab nebula as a synchrotron emission coming from the non-spherical MHD termination shock of relativistic pulsar wind. The post-shock flow is mildly relativistic; as a result the Doppler beaming has a strong impact on the shock appearance. The model can reproduce the knot location, size, elongation, brightness distribution, luminosity and polarization provided the effective magnetization of the section of the pulsar wind producing the knot is low, σ ≤ 1. In the striped wind model, this implies that the striped zone is rather wide, with the magnetic inclination angle of the Crab pulsar ≥45°; this agrees with the previous model-dependent estimate based on the gamma-ray emission of the pulsar. We conclude that the tiny knot is indeed a bright spot on the surface of a quasi-stationary magnetic relativistic shock and that this shock is a site of efficient particle acceleration. On the other hand, the deduced low magnetization of the knot plasma implies that this is an unlikely site for the Crab's gamma-ray flares, if they are related to the fast relativistic magnetic reconnection events.

  8. The Porcelain Crab Transcriptome and PCAD, the Porcelain Crab Microarray and Sequence Database

    Energy Technology Data Exchange (ETDEWEB)

    Tagmount, Abderrahmane; Wang, Mei; Lindquist, Erika; Tanaka, Yoshihiro; Teranishi, Kristen S.; Sunagawa, Shinichi; Wong, Mike; Stillman, Jonathon H.

    2010-01-27

    Background: With the emergence of a completed genome sequence of the freshwater crustacean Daphnia pulex, construction of genomic-scale sequence databases for additional crustacean sequences are important for comparative genomics and annotation. Porcelain crabs, genus Petrolisthes, have been powerful crustacean models for environmental and evolutionary physiology with respect to thermal adaptation and understanding responses of marine organisms to climate change. Here, we present a large-scale EST sequencing and cDNA microarray database project for the porcelain crab Petrolisthes cinctipes. Methodology/Principal Findings: A set of ~;;30K unique sequences (UniSeqs) representing ~;;19K clusters were generated from ~;;98K high quality ESTs from a set of tissue specific non-normalized and mixed-tissue normalized cDNA libraries from the porcelain crab Petrolisthes cinctipes. Homology for each UniSeq was assessed using BLAST, InterProScan, GO and KEGG database searches. Approximately 66percent of the UniSeqs had homology in at least one of the databases. All EST and UniSeq sequences along with annotation results and coordinated cDNA microarray datasets have been made publicly accessible at the Porcelain Crab Array Database (PCAD), a feature-enriched version of the Stanford and Longhorn Array Databases.Conclusions/Significance: The EST project presented here represents the third largest sequencing effort for any crustacean, and the largest effort for any crab species. Our assembly and clustering results suggest that our porcelain crab EST data set is equally diverse to the much larger EST set generated in the Daphnia pulex genome sequencing project, and thus will be an important resource to the Daphnia research community. Our homology results support the pancrustacea hypothesis and suggest that Malacostraca may be ancestral to Branchiopoda and Hexapoda. Our results also suggest that our cDNA microarrays cover as much of the transcriptome as can reasonably be captured in

  9. Comparative analysis of the proximate and elemental composition of the blue crab Callinectes sapidus, the warty crab Eriphia verrucosa, and the edible crab Cancer pagurus.

    Science.gov (United States)

    Zotti, Maurizio; Coco, Laura Del; Pascali, Sandra Angelica De; Migoni, Danilo; Vizzini, Salvatrice; Mancinelli, Giorgio; Fanizzi, Francesco Paolo

    2016-02-01

    The proximate composition and element contents of claw muscle tissue of Atlantic blue crabs (Callinectes sapidus) were compared with the native warty crab (Eriphia verrucosa) and the commercially edible crab (Cancer pagurus). The scope of the analysis was to profile the chemical characteristics and nutritive value of the three crab species. Elemental fingerprints showed significant inter-specific differences, whereas non-significant variations in the moisture and ash contents were observed. In the blue crab, protein content was significantly lower than in the other two species, while its carbon content resulted lower than that characterizing only the warty crab. Among micro-elements, Ba, Cr, Cu, Li, Mn, Ni, and Pb showed extremely low concentrations and negligible among-species differences. Significant inter-specific differences were observed for Na, Sr, V, Ba, Cd and Zn; in particular, cadmium and zinc were characterized in the blue crab by concentrations significantly lower than in the other two species. The analysis of the available literature on the three species indicated a general lack of comparable information on their elemental composition. The need to implement extended elemental fingerprinting techniques for shellfish quality assessment is discussed, in view of other complementary profiling methods such as NMR-based metabolomics.

  10. Experimental infections of Orchitophrya stellarum (Scuticociliata) in American blue crabs (Callinectes sapidus) and fiddler crabs (Uca minax).

    Science.gov (United States)

    Miller, Terrence L; Small, Hamish J; Peemoeller, Bhae-Jin; Gibbs, David A; Shields, Jeffrey D

    2013-11-01

    Outbreaks of an unidentified ciliate have occurred on several occasions in blue crabs from Chesapeake Bay held during winter months in flow-through systems. The parasite was initially thought to be Mesanophrys chesapeakensis, but molecular analysis identified it as Orchitophyra stellarum, a facultative parasite of sea stars (Asteroidea). We investigated the host-parasite association of O. stellarum in the blue crab host. Crabs were inoculated with the ciliate, or they were held in bath exposures after experimentally induced autotomy of limbs in order to determine potential mechanisms for infection. Crabs inoculated with the ciliate, or exposed to it after experimental autotomy, rapidly developed fatal infections. Crabs that were not experimentally injured, but were exposed to the ciliate, rarely developed infections; thus, indicating that the parasite requires a wound or break in the cuticle as a portal of entry. For comparative purposes, fiddler crabs, Uca minax, were inoculated with the ciliate in a dose-titration experiment. Low doses of the ciliate (10 per crab) were sometimes able to establish infections, but high intensity infections developed quickly at doses over 500 ciliates per crab. Chemotaxis studies were initiated to determine if the ciliate preferentially selected blue crab serum (BCS) over other nutrient sources. Cultures grown on medium with BCS or fetal bovine serum showed some conditioning in their selection for different media, but the outcome in choice experiments indicated that the ciliate was attracted to BCS and not seawater. Our findings indicate that O. stellarum is a facultative parasite of blue crabs. It can cause infections in exposed crabs at 10-15°C, but it requires a portal of entry for successful host invasion, and it may find injured hosts using chemotaxis.

  11. Importance of agricultural landscapes to nesting burrowing owls in the Northern Great Plains, USA

    Science.gov (United States)

    Restani, M.; Davies, J.M.; Newton, W.E.

    2008-01-01

    Anthropogenic habitat loss and fragmentation are the principle factors causing declines of grassland birds. Declines in burrowing owl (Athene cunicularia) populations have been extensive and have been linked to habitat loss, primarily the decline of black-tailed prairie dog (Cynomys ludovicianus) colonies. Development of habitat use models is a research priority and will aid conservation of owls inhabiting human-altered landscapes. From 2001 to 2004 we located 160 burrowing owl nests on prairie dog colonies on the Little Missouri National Grassland in North Dakota. We used multiple linear regression and Akaike's Information Criterion to estimate the relationship between cover type characteristics surrounding prairie dog colonies and (1) number of owl pairs per colony and (2) reproductive success. Models were developed for two spatial scales, within 600 m and 2,000 m radii of nests for cropland, crested wheatgrass (Agropyron cristatum), grassland, and prairie dog colonies. We also included number of patches as a metric of landscape fragmentation. Annually, fewer than 30% of prairie dog colonies were occupied by owls. None of the models at the 600 m scale explained variation in number of owl pairs or reproductive success. However, models at the 2,000 m scale did explain number of owl pairs and reproductive success. Models included cropland, crested wheatgrass, and prairie dog colonies. Grasslands were not included in any of the models and had low importance values, although percentage grassland surrounding colonies was high. Management that protects prairie dog colonies bordering cropland and crested wheatgrass should be implemented to maintain nesting habitat of burrowing owls. ?? 2008 Springer Science+Business Media B.V.

  12. Evolutionary diversification of coral-dwelling gall crabs (Cryptochiridae)

    NARCIS (Netherlands)

    Meij, Sancia Esmeralda Theonilla van der

    2015-01-01

    Gall crabs (Crustacea : Cryptochiridae) are small, coral-dwelling crabs that live in obligate association with their host corals (Scleractinia), on which they rely for food and shelter. They have been recorded from shallow and deeper waters (over 500 m), but the majority of the species live in reef

  13. Crabbed Waist Collisions in DAFNE and Super-B Design

    Energy Technology Data Exchange (ETDEWEB)

    Raimondi, P.; Alesini, D.; Biagini, M.E.; Biscari, C.; Boni, R.; Boscolo, M.; Bossi, F.; Buonomo, B.; Clozza, A.; Delle Monache, G.O.; Demma, T.; Di Pasquale, E.; Di Pirro, G.; Drago, A.; Gallo, A.; Ghigo, A.; Guiducci, S.; Ligi, C.; Marcellini, F.; Mazzitelli, Giovanni; Milardi, C.; /Frascati /Orsay, LAL /CERN /Rome III U. /Rome U. /Novosibirsk, IYF /KEK, Tsukuba /INFN, Pisa /INFN, Cosenza /SLAC /Frascati

    2011-11-02

    The new idea of increasing the luminosity of a collider with crab waist collisions and first experimental results from the DA{Phi}NE {Phi}-Factory at LNF, Frascati, using this concept are presented. Consequences for the design of future factories will be discussed. An outlook to the performance reach with crab waist collisions is given, with emphasis on future B Factories.

  14. 50 CFR Table 2 to Part 680 - Crab Species Code

    Science.gov (United States)

    2010-10-01

    ... 50 Wildlife and Fisheries 9 2010-10-01 2010-10-01 false Crab Species Code 2 Table 2 to Part 680 Wildlife and Fisheries FISHERY CONSERVATION AND MANAGEMENT, NATIONAL OCEANIC AND ATMOSPHERIC ADMINISTRATION..., Table 2 Table 2 to Part 680—Crab Species Code Species code Common name Scientific name 900...

  15. The Rhizocephalan parasite of the crab Xantho incisus (Leach)

    NARCIS (Netherlands)

    Boschma, H.

    1955-01-01

    As shown by Holthuis (1954), the correct name for the European crab commonly referred to as Xantho floridus (Montagu) is Xantho incisus (Leach). A Rhizocephalan parasite of this crab was first mentioned (without an indication of specific characters) by Gerbe (1862); afterwards specimens were recorde

  16. Crab biodiversity under different management schemes of mangrove ecosystems

    Directory of Open Access Journals (Sweden)

    M. Bandibas

    2016-01-01

    Full Text Available Reforestation is one of the Philippines’ government efforts to restore and rehabilitate degraded mangrove ecosystems. Although there is recovery of the ecosystem in terms of vegetation, the recovery of closely-linked faunal species in terms of community structure is still understudied. This research investigates the community structure of mangrove crabs under two different management schemes: protected mangroves and reforested mangroves. The transect-plot method was employed in each management scheme to quantify the vegetation, crab assemblages and environmental variables. Community composition of crabs and mangrove trees were compared between protected and reforested mangroves using non-metric multi-dimensional scaling and analysis of similarity in PRIMER 6. Chi-squared was used to test the variance of sex ration of the crabs. Canonical Correspondence Analysis was used to determine the relationship between crabs and environmental parameters. A total of twelve species of crabs belonging to six families were identified in protected mangroves while only four species were documented in reforested mangroves. Perisesarma indiarum and Baptozius vinosus were the most dominant species in protected and reforested mangrove, respectively.  Univariate analysis of variance of crab assemblage data revealed significant differences in crab composition and abundance between protected mangroves and from reforested mangroves (P

  17. Enhanced gamma-ray activity from the Crab nebula

    Science.gov (United States)

    Buehler, R.; Ciprini, S.

    2016-01-01

    Preliminary LAT analysis indicates enhanced gamma-ray activity from the Crab nebula. The daily-averaged gamma-ray emission (E > 100 MeV) from the direction of the Crab Nebula has surpassed 4.0 x 10^-6 ph cm^-2 s^-1 five times in the last 12 days.

  18. Brachyuran Crabs collected at Curaçao

    NARCIS (Netherlands)

    Rathbun, Mary J.

    1924-01-01

    In 1920 a report ¹) on Dr. BOEKE’S collection of crabs and shrimps from Curaçao and other Dutch West Indian Islands was published. Dr. VAN DER HORST’S collection forms an important addition to the fauna of Curaçao, especially as to the Xanthids and Majids, or spider crabs. Four new species were disc

  19. Small Angle Crab Compensation for LHC IR Upgrade

    CERN Document Server

    Akai, K; Dorda, U; Ohmi, K; Oide, K; Tomás, R; Zimmermann, T

    2007-01-01

    A small angle crab scheme is being considered for the LHC luminosity upgrade. In this paper we present a 400MHz superconducting cavity design and discuss the pertinent RF challenges. We also present a study on the beam-beam performance and proton-beam emittance growth in the presence of crab compensation, with RF noise sources.

  20. Burrowing and avoidance behaviour in marine organisms exposed to pesticide-contaminated sediment

    DEFF Research Database (Denmark)

    Møhlenberg, Flemming; Kiørboe, Thomas

    1983-01-01

    Behavioural effects of marine sediment contaminated with pesticides (6000 ppm parathion, 200 ppm methyl parathion, 200 ppm malathion) were studied in a number of marine organisms in laboratory tests and in situ. The burrowing behaviour in Macoma baltica, Cerastoderma edule, Abra alba, Nereis...... for Crangon crangon and Solea solea, but not for Carcinus meanas and Pomatoschistus minutus. The validity of both behavioural tests was supported by in situ observations and investigations on the distribution of the species. It is concluded that both tests are useful tools in the assessment of the impact...

  1. Mitigating by-catch of diamondback terrapins in crab pots

    Science.gov (United States)

    Hart, Kristen M.; Crowder, Larry B.

    2011-01-01

    Chronic by-catch of diamondback terrapins (Malaclemys terrapin) in blue crab (Callinectes sapidus) pots is a concern for terrapin conservation along the United States Atlantic and Gulf of Mexico coasts. Despite the availability of by-catch reduction devices (BRDs) for crab pots, adoption of BRDs has not been mandated and by-catch of terrapins continues. We conducted experimental fishing studies in North Carolina's year-round blue crab fishery from 2000 to 2004 to evaluate the ability of various BRDs to reduce terrapin by-catch without a concomitant reduction in the catch of blue crabs. In 4,822 crab pot days fished, we recorded only 21 terrapin captures. Estimated capture rates were 0.003 terrapins/pot per day in hard crab experimental fishing and 0.008 terrapins/pot per day in peeler experimental fishing. All terrapin captures occurred from April to mid-May within 321.4 m of the shoreline. Longer soak times produced more dead terrapins, with 4 live and 4 dead during hard crab experimental fishing and 11 live and 2 dead during peeler experimental fishing. The 4.0-cm BRDs in fall and 4.5-cm and 5.0-cm BRDs in spring reduced the catch of legal-sized male hard crabs by 26.6%, 21.2%, and 5.7%, respectively. Only the 5.0-cm BRDs did not significantly affect the catch of legal-sized hard male crabs. However, BRDs had no measurable effect on catch of target crabs in the peeler crab fishery. Our results identify 3 complementary and economically feasible tools for blue crab fishery managers to exclude terrapins from commercially fished crab pots in North Carolina: 1) gear modifications (e.g., BRDs); 2) distance-to-shore restrictions; and 3) time-of-year regulations. These measures combined could provide a reduction in terrapin by-catch of up to 95% without a significant reduction in target crab catch.

  2. Crabs in Labs: The Shore Crab (Carcinus maenas) as Teaching Material.

    Science.gov (United States)

    Hogarth, P. J.

    1983-01-01

    The shore crab (Carcinus maenas) is an excellent subject for school study, both in the field and the laboratory. It is easily collected and maintained and can be used for a wide range of investigations. Some background details are given and possible areas of investigation suggested. (Author)

  3. 75 FR 32360 - Proposed Information Collection; Comment Request; Alaska Crab Report Forms

    Science.gov (United States)

    2010-06-08

    ... Crab Report Forms AGENCY: National Oceanic and Atmospheric Administration (NOAA), Department of.... 1801 et seq.) The FMP for Bering Sea and Aleutian Islands (BSAI) Crab includes the Crab Rationalization (CR) Program, a limited access system that allocates BSAI Management Area Crab resources...

  4. Death rate due to horseshoe crab poisoning:summarization on Thai reports

    Institute of Scientific and Technical Information of China (English)

    Beuy Joob; Viroj Wiwanitkit

    2015-01-01

    Horseshoe crab can be poisonous and intoxication due to intake of horseshoe crab is possible. Horseshoe crab intoxication can be seen in many countries with seacoasts including Thailand. Here, the authors summarized the death rate due to horseshoe crab poisoning in Thailand.

  5. Avian Influenza Virus for Phylogenetic Tree Reconstrction Based on Extension Burrows-Wheeler%基于扩展Burrows-Wheeler算法重构禽流感病毒的进化树

    Institute of Scientific and Technical Information of China (English)

    夏阳; 白凤兰; 刘立伟

    2014-01-01

    基于核苷酸的物理化学性质,利用扩展Burrows-Wheeler算法和Burrows-Wheeler相似性分布,对80种H5NI病毒的DNA序列的HA片段进行相似性比较,同时构建序列进化树,得到较好的结果.通过分析禽流感病毒之间的相似度关系,可为研究禽流感区域蔓延的特点提供一定的理论依据.

  6. Estimating animal populations and body sizes from burrows: Marine ecologists have their heads buried in the sand

    Science.gov (United States)

    Schlacher, Thomas A.; Lucrezi, Serena; Peterson, Charles H.; Connolly, Rod M.; Olds, Andrew D.; Althaus, Franziska; Hyndes, Glenn A.; Maslo, Brooke; Gilby, Ben L.; Leon, Javier X.; Weston, Michael A.; Lastra, Mariano; Williams, Alan; Schoeman, David S.

    2016-06-01

    Most ecological studies require knowledge of animal abundance, but it can be challenging and destructive of habitat to obtain accurate density estimates for cryptic species, such as crustaceans that tunnel deeply into the seafloor, beaches, or mudflats. Such fossorial species are, however, widely used in environmental impact assessments, requiring sampling techniques that are reliable, efficient, and environmentally benign for these species and environments. Counting and measuring the entrances of burrows made by cryptic species is commonly employed to index population and body sizes of individuals. The fundamental premise is that burrow metrics consistently predict density and size. Here we review the evidence for this premise. We also review criteria for selecting among sampling methods: burrow counts, visual censuses, and physical collections. A simple 1:1 correspondence between the number of holes and population size cannot be assumed. Occupancy rates, indexed by the slope of regression models, vary widely between species and among sites for the same species. Thus, 'average' or 'typical' occupancy rates should not be extrapolated from site- or species specific field validations and then be used as conversion factors in other situations. Predictions of organism density made from burrow counts often have large uncertainty, being double to half of the predicted mean value. Whether such prediction uncertainty is 'acceptable' depends on investigators' judgements regarding the desired detectable effect sizes. Regression models predicting body size from burrow entrance dimensions are more precise, but parameter estimates of most models are specific to species and subject to site-to-site variation within species. These results emphasise the need to undertake thorough field validations of indirect census techniques that include tests of how sensitive predictive models are to changes in habitat conditions or human impacts. In addition, new technologies (e.g. drones

  7. Biogeomorphically driven salt pan formation in Sarcocornia-dominated salt-marshes

    Science.gov (United States)

    Escapa, Mauricio; Perillo, Gerardo M. E.; Iribarne, Oscar

    2015-01-01

    Salt-marshes are under increasing threat, particularly from sea-level rise and increased wave action associated with climate change. The development and stability of these valuable habitats largely depend on complex interactions between biotic and abiotic processes operating at different scales. Also, interactions between biotic and abiotic processes drive internal morphological change in salt-marshes. In this paper we used a biogeomorphological approach to assess the impact of biological activities and interactions on salt pan formation in Sarcocornia-dominated salt marshes. Salt pans represent a key physiographic feature of salt-marshes and recent studies hypothesized that biogeomorphic processes could be related to salt pan formation in SW Atlantic salt-marshes. The glasswort Sarcocornia perennis is one of the dominant plants in the salt-marshes of the Bahía Blanca Estuary (Argentina) where they form patches up to 8 m in diameter. These salt-marshes are also inhabited in great densities by the burrowing crab Neohelice (Chasmagnathus) granulata whose bioturbation rates are among the highest reported for salt-marshes worldwide. A set of biological interactions between N. granulata and S. perennis appears to be responsible for salt pan development in these areas which has not been described elsewhere. The main objective of this work was to determine the ecological interactions occurring between plants and crabs that lead to salt pan formation by using field-based sampling and manipulative experiments. Our results showed that S. perennis facilitated crab colonization of the salt-marsh by buffering otherwise stressful physical conditions (e.g., temperature, desiccation). Crabs preferred to construct burrows underneath plants and, once they reach high densities (up to 40 burrows m- 2), the sediment reworking caused plant die-off in the central area of patches. At this state, the patches lose elevation and become depressed due to the continuous bioturbation by crabs

  8. Luminosity and Crab Waist Collision Studies

    CERN Document Server

    Wu, Wanwei

    2015-01-01

    In high energy physics, the luminosity is one useful value to characterize the performance of a particle collider. To gain more available data, we need to maximize the luminosity in most collider experiments. With the discussions of tune shift involved the beam dynamics and a recently proposed "crabbed waist" scheme of beam-beam collisions, we present some qualitative analysis to increase the luminosity. In addition, beam-beam tune shifts and luminosities of $e^{+}e^{-}$, $pp$/$p\\bar{p}$, and $\\mu^{+}\\mu^{-}$ colliders are discussed.

  9. Element Masses in the Crab Nebula

    Science.gov (United States)

    Sibley, Adam R.; Katz, Andrea M.; Satterfield, Timothy J.; Vanderveer, Steven J.; MacAlpine, Gordon M.

    2016-10-01

    Using our previously published element abundance or mass-fraction distributions in the Crab Nebula, we derived actual mass distributions and estimates for overall nebular masses of hydrogen, helium, carbon, nitrogen, oxygen and sulfur. As with the previous work, computations were carried out for photoionization models involving constant hydrogen density and also constant nuclear density. In addition, employing new flux measurements for [Ni ii] λ7378, along with combined photoionization models and analytic computations, a nickel abundance distribution was mapped and a nebular stable nickel mass estimate was derived.

  10. Identification of irradiated crab using EPR

    Energy Technology Data Exchange (ETDEWEB)

    Maghraby, A. [Radiation Dosimetry Department, National Institute for Standards (NIS), Ministry of Scientific Research, Haram, 12211- Giza, P.O. Box: 136 (Egypt)]. E-mail: maghrabism@yahoo.com

    2007-02-15

    EPR spectroscopy is a fast and powerful technique for the identification of irradiated food. Crab exoskeleton was divided into six parts: dactyl, cheliped, carapace, apron, swimming legs, and walking legs. Samples of the exoskeleton were prepared and irradiated to Cs-137 gamma radiation in the range (1.156-5.365 kGy). EPR spectra of unirradiated as well as irradiated samples were recorded and analyzed. Response to gamma radiation was plotted for each part of the exoskeleton, dactyl was found to be the most sensitive part, followed by the apron (38%), cheliped (37%), walking legs (30%), swimming legs (24%), and carapace (21%) relative to the dactyl response.

  11. Habitat Suitability Index Models: Red king crab

    Science.gov (United States)

    Jewett, Stephen C.; Onuf, Christopher P.

    1988-01-01

    A review and synthesis of existing information were used to develop a Habitat Suitability Index (HSI) model for evaluating habitat of different life stages of red king crab (Paralithodes camtschatica). A model consolidates habitat use information into a framework appropriate for field application, and is scaled to produce an index between 0.0 (unsuitable habitat) and 1.0 (optimum habitat) in Alaskan coastal waters, especially in the Gulf of Alaska and the southeastern Bering Sea. HSI models are designed to be used with Habitat Evaluation Procedures previously developed by the U.S. Fish and Wildlife Service.

  12. LHC Crab Cavity Coupler Test Boxes

    CERN Document Server

    Mitchell, James; Burt, Graeme; Calaga, Rama; Macpherson, Alick; Montesinos, Eric; Silva, Subashini; Tutte, Adam; Xiao, Binping

    2016-01-01

    The LHC double quarter wave (DQW) crab cavities have two different types of Higher Order Mode (HOM) couplers in addition to a fundamental power coupler (FPC). The FPC requires conditioning, so to achieve this we have designed a radio-frequency (RF) quarter wave resonator to provide high transmission between two opposing FPCs. For the HOM couplers we must ensure that the stop-band filter is positioned at the cavity frequency and that peak transmission occurs at the same frequencies as the strongest HOMs. We have designed two test boxes which preserve the cavity spectral response in order to test the couplers.

  13. Do leucocytes reflect condition in nestling burrowing parrots Cyanoliseus patagonus in the wild?

    Science.gov (United States)

    Masello, Juan F; Choconi, R Gustavo; Helmer, Matthias; Kremberg, Thomas; Lubjuhn, Thomas; Quillfeldt, Petra

    2009-02-01

    The different leucocyte types are an important part of the immune system. Thus, they have been used in ecological studies to assess immune function and physiological stress in wild birds. It is generally assumed that increased stress and decreased condition are associated with an increase in the ratio of heterophils to lymphocytes, the H/L ratio. We studied leucocyte profiles in relation to body condition in nestling Burrowing Parrots (Cyanoliseus patagonus) in North-eastern Patagonia, Argentina. As in other wild parrots, heterophils were the most numerous leucocyte type, suggesting strong investment into innate immunity. Leucocyte profiles did not change with the age, while nestlings in better body condition increased the number of heterophils. Because the number of lymphocytes was independent of body condition, as a result we observed a positive correlation between body condition and the H/L ratio. The total number of leucocytes relative to erythrocytes increased in nestlings in better body condition, indicating a larger overall investment into immune function in well-nourished nestlings. The observed heterophilic profiles of nestling Burrowing Parrots together with the positive relationship between H/L ratio and body condition may indicate a favoured investment in a robust innate immunity that reduces the risk of infection taking hold in these long-lived birds.

  14. Real estate ads in Emei music frog vocalizations: female preference for calls emanating from burrows.

    Science.gov (United States)

    Cui, Jianguo; Tang, Yezhong; Narins, Peter M

    2012-06-23

    During female mate choice, both the male's phenotype and resources (e.g. his nest) contribute to the chooser's fitness. Animals other than humans are not known to advertise resource characteristics to potential mates through vocal communication; although in some species of anurans and birds, females do evaluate male qualities through vocal communication. Here, we demonstrate that calls of the male Emei music frog (Babina dauchina), vocalizing from male-built nests, reflect nest structure information that can be recognized by females. Inside-nest calls consisted of notes with energy concentrated at lower frequency ranges and longer note durations when compared with outside-nest calls. Centre frequencies and note durations of the inside calls positively correlate with the area of the burrow entrance and the depth of the burrow, respectively. When given a choice between outside and inside calls played back alternately, more than 70 per cent of the females (33/47) chose inside calls. These results demonstrate that males of this species faithfully advertise whether or not they possess a nest to potential mates by vocal communication, which probably facilitates optimal mate selection by females. These results revealed a novel function of advertisement calls, which is consistent with the wide variation in both call complexity and social behaviour within amphibians.

  15. The antenna of a burrowing dragonfly larva, Onychogomphus forcipatus (Anisoptera, Gomphidae).

    Science.gov (United States)

    Rebora, Manuela; Piersanti, Silvana; Salerno, Gianandrea; Gorb, Stanislav

    2015-11-01

    The larva of the dragonfly Onychogomphus forcipatus (Anisoptera, Gomphidae) has a burrowing lifestyle and antennae composed of four short and broad segments (scape, pedicel and a two-segmented flagellum). The present ultrastructural investigation revealed that different sensilla and one gland are located on the antenna. There is a great diversity of mechanoreceptors of different kinds. In particular club-shaped sensilla, sensilla chaetica, and tree-like sensilla show the typical structure of bristles, the most common type of mechanoreceptors, usually responding to direct touch, while numerous long thin thorny trichoid sensilla show a morphology recalling the structure of filiform hair mechanoreceptors. The latter ones are presumably important in larval Odonata for current detection and rheotactic orientation, especially in a burrowing species. On the smooth apical cuticle of the second flagellar segment, three structures are visible: (1) a small ellipsoidal pit hosting a convoluted peg, the morphology of which resembles that of a typical chemoreceptor (even if pores are lacking), (2) a couple of small pits (not investigated under TEM), and (3) one wide depression with spherical structures, the internal morphology of which lets us assume that it is a gland with unknown function. This is the first report of an antennal gland in palaeopteran insects.

  16. [The supra-occipital bone in cetaceans and burrowing rodents. morphological convergence induced by the post-cephalic pole?].

    Science.gov (United States)

    Courant, F; Marchand, D

    2000-02-01

    A comparative study of the cranial morphologies of cetaceans and of rodents that use their incisors for burrowing brings out morphological convergences concerning the supra-occipital bone. These phyletically very remote groups are both subject to the same mechanical constraint, viz. the need for the spinal column to be aligned with the anteroposterior axis of the skull. This constraint, which is related to swimming in cetaceans and burrowing in rodents, entails three major points of convergence: 1) a clearly backward facing foramen magnum; 2) a shortened or even greatly shortened neck, sometimes with cervical vertebrae fused together; and 3) an uprighted or even forward tilted supra-occipital bone.

  17. Artificial warthog burrows used to sample adult and immature tsetse (Glossina spp in the Zambezi Valley of Zimbabwe.

    Directory of Open Access Journals (Sweden)

    John W Hargrove

    2015-03-01

    Full Text Available The biology of adult tsetse (Glossina spp, vectors of trypanosomiasis in Africa, has been extensively studied - but little is known about larviposition in the field.In September-November 1998, in the hot-dry season in Zimbabwe's Zambezi Valley, we used artificial warthog burrows to capture adult females as they deposited larvae. Females were subjected to ovarian dissection and were defined as perinatal flies, assumed to have entered burrows to larviposit, if oocyte sizes indicated >95% pregnancy completion. Perinatal flies were defined as full-term pregnant if there was a late third instar larva in utero, or postpartum if the uterus was empty. All other females were defined as pre-full-term pregnant (pre-FT. Of 845 G. m. morsitans captured, 91% (765 were female and 295/724 (41% of females dissected were perinatal flies. By contrast, of 2805 G. pallidipes captured only 71% (2003 were female and only 33% (596/1825 of females were perinatal. Among all perinatal females 67% (596/891 were G. pallidipes. Conversely, in burrows not fitted with traps - such that flies were free to come and go - 1834 (59% of pupae deposited were G. m. morsitans and only 1297 (41% were G. pallidipes. Thus, while more full-term pregnant G. pallidipes enter burrows, greater proportions of G. m. morsitans larviposit in them, reflecting a greater discrimination among G. pallidipes in choosing larviposition sites. Catches of males and pre-FT females increased strongly with temperatures above 32°C, indicating that these flies used burrows as refuges from high ambient temperatures. Conversely, catches of perinatal females changed little with maximum temperature but declined from late September through November: females may anticipate that burrows will be inundated during the forthcoming wet season. Ovarian age distributions of perinatal and pre-FT females were similar, consistent with all ages of females larvipositing in burrows with similar probability.Artificial warthog

  18. Measuring the contribution of benthic ecosystem engineering species to the ecosystem services of an estuary: A case study of burrowing shrimps in Yaquina Estuary, Oregon - April 2009

    Science.gov (United States)

    Burrowing shrimps are regarded as ecosystem engineering species in many coastal ecosystems worldwide, including numerous estuaries of the west coast of North America (Baja California to British Columbia). In estuaries of the U.S. Pacific Northwest, two species of large burrowing...

  19. Measuring the contribution of benthic ecosystem engineering species to the ecosystem services of an estuary: A case study of burrowing shrimps in Yaquina Estuary, Oregon

    Science.gov (United States)

    Burrowing shrimps are regarded as ecosystem engineering species in many coastal ecosystems worldwide, including numerous estuaries of the west coast of North America (Baja California to British Columbia). In estuaries of the U.S. Pacific Northwest, two species of large burrowing...

  20. Thermal specialization across large geographical scales predicts the resilience of mangrove crab populations to global warming

    KAUST Repository

    Fusi, Marco

    2014-11-18

    The broad prediction that ectotherms will be more vulnerable to climate change in the tropics than in temperate regions includes assumptions about centre/edge population effects that can only be tested by within-species comparisons across wide latitudinal gradients. Here, we investigated the thermal vulnerability of two mangrove crab species, comparing populations at the centre (Kenya) and edge (South Africa) of their distributions. At the same time, we investigated the role of respiratory mode (water- versus air-breathing) in determining the thermal tolerance in amphibious organisms. To do this, we compared the vulnerability to acute temperature fluctuations of two sympatric species with two different lifestyle adaptations: the free living Perisesarma guttatum and the burrowing Uca urvillei, both pivotal to the ecosystem functioning of mangroves. The results revealed the air-breathing U. urvillei to be a thermal generalist with much higher thermal tolerances than P. guttatum. Importantly, however, we found that, while U. urvillei showed little difference between edge and centre populations, P. guttatum showed adaptation to local conditions. Equatorial populations had elevated tolerances to acute heat stress and mechanisms of partial thermoregulation, which make them less vulnerable to global warming than temperate conspecifics. The results reveal both the importance of respiratory mode to thermal tolerance and the unexpected potential for low latitude populations/species to endure a warming climate. The results also contribute to a conceptual model on the latitudinal thermal tolerance of these key species. This highlights the need for an integrated population-level approach to predict the consequences of climate change. © 2014 The Authors.

  1. Crab death assemblages from Laguna Madre and vicinity, Texas

    Energy Technology Data Exchange (ETDEWEB)

    Plotnick, R.E.; McCarroll, S. (Univ. of Illinois, Chicago (USA)); Powell, E. (Texas A M Univ., College Station (USA))

    1990-02-01

    Crabs are a major component of modern marine ecosystems, but are only rarely described in fossil assemblages. Studies of brachyuran taphonomy have examined either the fossil end-products of the taphonomic process or the very earliest stages of decay and decomposition. The next logical step is the analysis of modern crab death assemblages; i.e., studies that examine taphonomic loss in areas where the composition of the living assemblage is known. The authors studied crab death assemblages in shallow water sediments at several localities in an near Laguna Madre, Texas. Nearly every sample examined contained some crab remains, most commonly in the form of isolated claws (dactyl and propodus). A crab fauna associated with a buried grass bed contained abundant remains of the xanthid crab Dyspanopeus texanus, including carapaces, chelipeds, and thoraxes, as well as fragments of the portunid Callinectes sapidus and the majiid Libinia dubia. Crab remains may be an overlooked portion of many preserved benthic assemblages, both in recent and modern sediments.

  2. The porcelain crab transcriptome and PCAD, the porcelain crab microarray and sequence database.

    Directory of Open Access Journals (Sweden)

    Abderrahmane Tagmount

    Full Text Available BACKGROUND: With the emergence of a completed genome sequence of the freshwater crustacean Daphnia pulex, construction of genomic-scale sequence databases for additional crustacean sequences are important for comparative genomics and annotation. Porcelain crabs, genus Petrolisthes, have been powerful crustacean models for environmental and evolutionary physiology with respect to thermal adaptation and understanding responses of marine organisms to climate change. Here, we present a large-scale EST sequencing and cDNA microarray database project for the porcelain crab Petrolisthes cinctipes. METHODOLOGY/PRINCIPAL FINDINGS: A set of approximately 30K unique sequences (UniSeqs representing approximately 19K clusters were generated from approximately 98K high quality ESTs from a set of tissue specific non-normalized and mixed-tissue normalized cDNA libraries from the porcelain crab Petrolisthes cinctipes. Homology for each UniSeq was assessed using BLAST, InterProScan, GO and KEGG database searches. Approximately 66% of the UniSeqs had homology in at least one of the databases. All EST and UniSeq sequences along with annotation results and coordinated cDNA microarray datasets have been made publicly accessible at the Porcelain Crab Array Database (PCAD, a feature-enriched version of the Stanford and Longhorn Array Databases. CONCLUSIONS/SIGNIFICANCE: The EST project presented here represents the third largest sequencing effort for any crustacean, and the largest effort for any crab species. Our assembly and clustering results suggest that our porcelain crab EST data set is equally diverse to the much larger EST set generated in the Daphnia pulex genome sequencing project, and thus will be an important resource to the Daphnia research community. Our homology results support the pancrustacea hypothesis and suggest that Malacostraca may be ancestral to Branchiopoda and Hexapoda. Our results also suggest that our cDNA microarrays cover as much of the

  3. The inner knot of the Crab nebula

    CERN Document Server

    Lyutikov, Maxim; Porth, Oliver

    2015-01-01

    We model the inner knot of the Crab Nebula as a synchrotron emission coming from the non-spherical MHD termination shock of relativistic pulsar wind. The post-shock flow is mildly relativistic; as a result the Doppler-beaming has a strong impact on the shock appearance. The model can reproduce the knot location, size, elongation, brightness distribution, luminosity and polarization provided the effective magnetization of the section of the pulsar wind producing the knot is low, $\\sigma \\leq 1$. In the striped wind model, this implies that the striped zone is rather wide, with the magnetic inclination angle of the Crab pulsar $\\ge 45^\\circ$; this agrees with the previous model-dependent estimate based on the gamma-ray emission of the pulsar. We conclude that the tiny knot is indeed a bright spot on the surface of a quasi-stationary magnetic relativistic shock and that this shock is a site of efficient particle acceleration. On the other hand, the deduced low magnetization of the knot plasma implies that this i...

  4. Pulse profile stability of the Crab pulsar

    Institute of Scientific and Technical Information of China (English)

    Chetana Jain; Biswajit Paul

    2011-01-01

    We present an X-ray timing analysis of the Crab pulsar,PSR B0531+21,using archival RXTE data.We have investigated the stability of the Crab pulse profile,in soft (2-20keV) and hard (30-100keV) X-ray energies,over the last decade of RXTE operation.The analysis includes measurement of the separation between the two pulse peaks and the intensity and widths of the two peaks.We did not find any significant time dependency in the pulse shape.The two peaks have been stable in phase,intensity and width for the last ten years.The first pulse is relatively stronger at soft X-rays.The first pulse peak is narrower than the second peak in both soft and hard X-ray energies.Both the peaks show a slow rise and a steeper fall.The ratio of the pulsed photons in the two peaks is also constant in time.

  5. AFSC/RACE/SAP/Long: Data from: Effects of Ocean Acidification on Juvenile Red King Crab (Paralithodes camtschaticus) and Tanner Crab (Chionoecetes bairdi) Growth, Condition, Calcification, and Survival

    Data.gov (United States)

    National Oceanic and Atmospheric Administration, Department of Commerce — This data set is the results of a laboratory experiment. Juvenile red king crab and Tanner crab were reared in individual containers for nearly 200 days in flowing...

  6. A documentation on burrows in hard substrates of ferromanganese crusts and associated soft sediments from the Central Indian Ocean

    Digital Repository Service at National Institute of Oceanography (India)

    Banerjee, R.

    sediments on the sea floor also show numerous records of recent bioturbation activity. Thus presence of the ancient and modern burrows from tha same locality on the ocean floor indicates a prolonged bioturbation activity in this part of the CIB and this may...

  7. Distributional changes in the western Burrowing Owl (Athene cunicularia hypugaea) in North America from 1967 to 2008

    Science.gov (United States)

    Macias-Duarte, Alberto; Conway, Courtney J.

    2015-01-01

    The quantification of shifts in bird distributions in response to climate change provides an opportunity to gain a deeper understanding of the processes that influence species persistence. We used data from the North American Breeding Bird Survey (BBS) to document changes in the distributional limits of the western Burrowing Owl (Athene cunicularia hypugaea) from 1967 to 2008. We used logistic regression to model presence probability (p) as a function of longitude, latitude, and year. We modeled a linear trend in logit(p) through time with slope and intercept modeled as a double Fourier series of longitude and latitude. We found that the western Burrowing Owl has experienced an intriguing southward shift in the northern half of its breeding range, contrary to what is predicted by most species niche models and what has been observed for many other species in North America. The breeding range of the Burrowing Owl has been shrinking near its northern, western, and eastern edges. Our model detected the population declines that were observed in California and eastern Washington, in locations where maps based on route-specific estimating equations had predicted significant population increases. We suggest that the northern boundary of the breeding distribution of the western Burrowing Owl has contracted southward and the southern boundary of the species' breeding distribution has expanded southward into areas of northern Mexico that were formerly used only by wintering migrants.

  8. The Effect of Sand Particle Size on the Burrowing Ability of the Beach Mysid Gastrosaccus psammodytes Tattersall

    Science.gov (United States)

    Nel, R.; McLachlan, A.; Winter, D.

    1999-05-01

    Laboratory studies on the burrowing rates of the mysid shrimp, Gastrosaccus psammodytes , in a series of well-sorted sediments, determined whether (1) burial times were dependent on grain size and (2) if natural population distribution may be influenced by grain size on beaches. Burial times were tested in nine well-sorted sediments with grain size ranging from 90 to 2000μm. Large individuals (i.e. gravid females) were used. G. psammodytes burrowed fastest in 125-1000μm sand with mean burial times less than 1·6s. Burial time increased to approximately 2s in 90-125μm sand. G. psammodytes could not burrow in grain sizes coarser than 1000μm. G. psammodytes has been reported to occur on beaches with grain sizes ranging from 90 to 500μm but are uncommon on beaches with coarser sand. It appears that population distribution may be influenced by grain size that is probably not related to the animals' burial time ability, but rather their inability to burrow completely into coarse sand. Indirectly, grain size may also influence the morphodynamic state of a beach and therefore food availability since coarse-grained beaches tend to be reflective with little surf production.

  9. Population dynamics and damage potential of the burrowing nematode, Radopholus similis, on Anthurium andreanum grown in soil-less medium

    NARCIS (Netherlands)

    Amsing, J.J.; Schrama, P.M.M.; Stapel, L.H.M.

    2002-01-01

    In a 14 month pot experiment with Anthurium andreanum cv. Sonate, grown in soil-less medium in glasshouse conditions, burrowing nematode (Radopholus similis) population development and crop damage were investigated. At 9 days after transplanting, plants were inoculated with R. similis at four initia

  10. Comparison of automated BAX polymerase chain reaction and standard culture methods for detection of Listeria monocyogenes in blue crab meat (Callinectus sapidus) and blue crab processing plants

    Science.gov (United States)

    This study compared the BAX Polymerase Chain Reaction method (BAX PCR) with the Standard Culture Method (SCM) for detection of L. monocytogenes in blue crab meat and crab processing plants. The aim of this study was to address this data gap. Raw crabs, finished products and environmental sponge samp...

  11. Evolutionary history of true crabs (Crustacea: Decapoda: Brachyura) and the origin of freshwater crabs.

    Science.gov (United States)

    Tsang, Ling Ming; Schubart, Christoph D; Ahyong, Shane T; Lai, Joelle C Y; Au, Eugene Y C; Chan, Tin-Yam; Ng, Peter K L; Chu, Ka Hou

    2014-05-01

    Crabs of the infra-order Brachyura are one of the most diverse groups of crustaceans with approximately 7,000 described species in 98 families, occurring in marine, freshwater, and terrestrial habitats. The relationships among the brachyuran families are poorly understood due to the high morphological complexity of the group. Here, we reconstruct the most comprehensive phylogeny of Brachyura to date using sequence data of six nuclear protein-coding genes and two mitochondrial rRNA genes from more than 140 species belonging to 58 families. The gene tree confirms that the "Podotremata," are paraphyletic. Within the monophyletic Eubrachyura, the reciprocal monophyly of the two subsections, Heterotremata and Thoracotremata, is supported. Monophyly of many superfamilies, however, is not recovered, indicating the prevalence of morphological convergence and the need for further taxonomic studies. Freshwater crabs were derived early in the evolution of Eubrachyura and are shown to have at least two independent origins. Bayesian relaxed molecular methods estimate that freshwater crabs separated from their closest marine sister taxa ~135 Ma, that is, after the break up of Pangaea (∼200 Ma) and that a Gondwanan origin of these freshwater representatives is untenable. Most extant families and superfamilies arose during the late Cretaceous and early Tertiary.

  12. Sporothrix schenckii sensu stricto isolated from soil in an armadillo's burrow.

    Science.gov (United States)

    Rodrigues, Anderson Messias; Bagagli, Eduardo; de Camargo, Zoilo Pires; Bosco, Sandra de Moraes Gimenes

    2014-04-01

    Sporotrichosis is a polymorphic disease of man and animals caused by traumatic implantation of propagules into the skin and subcutaneous tissue. Pathogenic species includes S. brasiliensis, S. schenckii, S. globosa and S. luriei. The disease is remarkable for its occurrence as sapronoses and/or zoonosis outbreaks in tropical and subtropical areas; although, the ecology of the clinical clade is still puzzling. Here, we describe an anamorphic Sporothrix strain isolated from soil in an armadillo's burrow, which was located in a hyper endemic area of Paracoccidioidomycosis in Brazil. This isolate was identified as S. schenckii sensu stricto (Clade IIa) based on morphological and physiological characteristics and phylogenetic analyses of calmodulin sequences. We then discuss the role of the nine-banded armadillo Dasypus novemcinctus as a natural carrier of Sporothrix propagules to better understand Sporothrix sources in nature and reveal essential aspects about the pathogen's eco-epidemiology.

  13. Lossless compression of medical images using Burrows-Wheeler Transformation with Inversion Coder.

    Science.gov (United States)

    Preston, Collin; Arnavut, Ziya; Koc, Basar

    2015-08-01

    Medical imaging is a quickly growing industry where the need for highly efficient lossless compression algorithms is necessary in order to reduce storage space and transmission rates for the large, high resolution, medical images. Due to the fact that medical imagining cannot utilize lossy compression, in the event that vital information may be lost, it is imperative that lossless compression be used. While several authors have investigated lossless compression of medical images, the Burrows-Wheeler Transformation with an Inversion Coder (BWIC) has not been examined. Our investigation shows that BWIC runs in linear time and yields better compression rates than well-known image coders, such as JPEG-LS and JPEG-2000.

  14. Creeping hair: an isolated hair burrowing in the uppermost dermis resembling larva migrans.

    Science.gov (United States)

    Sakai, Rie; Higashi, Kushio; Ohta, Miyuki; Sugimoto, Yasushi; Ikoma, Yukiko; Horiguchi, Yuji

    2006-01-01

    A 55-year-old Japanese male presented with a slowly moving linear erythema that looked like an eruption of creeping disease, or cutaneous larva migrans. The eruption extended linearly along Langer's line of the lateral side of the abdomen to the lower back, leaving wave-like erythema. In the top third of the erythematous eruption, close examination demonstrated a black thin line, which was revealed to be a hair shaft by a shallow incision of the skin. After removal of the hair, the eruption diminished immediately, leaving a slight pigmentation. An ingrown pubic hair seemed to have migrated with the lower end forward along Langer's line, because of the arrangement of hair cuticle and the force of body motion. Linearly moving erythematous eruptions that look like that of larva migrans should be differentiated from creeping hair by close examination detecting burrowing hair.

  15. Crab Orchard National Wildlife Refuge : Furbearer Management Plan

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — The Crab Orchard National Wildlife Refuge Furbearer Management Plan directs the management and regulation of trapping. The furbearer management program directly...

  16. Crab Orchard National Wildlife Refuge [Annual narrative : May - August, 1953

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — This narrative report for Crab Orchard National Wildlife Refuge outlines Refuge accomplishments from May through August of 1953. The report begins by summarizing the...

  17. Crab Orchard National Wildlife Refuge [Narrative report : January - April, 1953

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — This narrative report for Crab Orchard National Wildlife Refuge outlines Refuge accomplishments from January through April of 1953. The report begins by summarizing...

  18. Narrative report : 1970. Crab Orchard National Wildlife Refuge.

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — This annual narrative report for Crab Orchard National Wildlife Refuge outlines Refuge accomplishments during the 1970 calendar year. The report begins by...

  19. Removal of nickel from aqueous solutions using crab shells.

    Science.gov (United States)

    Pradhan, Swapna; Shukla, Shyam S; Dorris, Kenneth L

    2005-10-17

    Partially converted crab shell waste, which contains chitosan, was used to remove nickel from water. The chelating ability of chitosan makes it an excellent adsorbent for removing pollutants. Advantages of chitosan in crab shells include availability, low cost, and high biocompatibility. The metal uptake by partially converted crab shell waste was successful and rapid. The sorption occurred primarily within 5 min. The sorption mechanism appears to be quite complicated and cannot be adequately described by either the Langmuir or Freundlich theories. Various anions, including chloride, bromide, fluoride, acetate, sulfate, nitrate, and phosphate, were found to have a very small effect on the capacity of the crab shells for uptake of nickel. The effect of pH was also found not to be prominent.

  20. AFSC/REFM: BSAI Crab Economic Data Report

    Data.gov (United States)

    National Oceanic and Atmospheric Administration, Department of Commerce — Economic data collected for years 1998, 2001, 2004, and 2005 and onward for the BSAI Crab Economic Data Report (EDR). Reporting is required of any owner or...

  1. Crab Orchard National Wildlife Refuge [Annual narrative : September - December, 1952

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — This narrative report for Crab Orchard National Wildlife Refuge outlines Refuge accomplishments from September through December of 1952. The report begins by...

  2. Crab Orchard National Wildlife Refuge [Annual narrative : May - August, 1952

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — This narrative report for Crab Orchard National Wildlife Refuge outlines Refuge accomplishments from May through August of 1952. The report begins by summarizing the...

  3. Crab cavities: Past, present, and future of a challenging device

    Energy Technology Data Exchange (ETDEWEB)

    Wu, Q. [Brookhaven National Lab. (BNL), Upton, NY (United States)

    2015-05-03

    In two-ring facilities operating with a crossing-angle collision scheme, luminosity can be limited due to an incomplete overlapping of the colliding bunches. Crab cavities then are introduced to restore head-on collisions by providing the destined opposite deflection to the head and tail of the bunch. An increase in luminosity was demonstrated at KEKB with global crab-crossing, while the Large Hardron Collider (LHC) at CERN currently is designing local crab crossing for the Hi-Lumi upgrade. Future colliders may investigate both approaches. In this paper, we review the challenges in the technology, and the implementation of crab cavities, while discussing experience in earlier colliders, ongoing R&D, and proposed implementations for future facilities, such as HiLumi-LHC, CERN’s compact linear collider (CLIC), the international linear collider (ILC), and the electron-ion collider under design at BNL (eRHIC).

  4. AFSC/RACE/SAP/Urban: Golden King Crab tagging

    Data.gov (United States)

    National Oceanic and Atmospheric Administration, Department of Commerce — The data is comprised of the records of individual male golden king crab (GKC) tagged at the Kodiak Laboratory. Initial size, shell condition and missing limbs was...

  5. Crab Orchard National Wildlife Refuge [Annual narrative : September - December, 1950

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — This narrative report for Crab Orchard National Wildlife Refuge outlines Refuge accomplishments from September through December of 1950. The report begins by...

  6. Crab Orchard National Wildlife Refuge [Annual narrative : May - August, 1950

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — This narrative report for Crab Orchard National Wildlife Refuge outlines Refuge accomplishments from May through August of 1950. The report begins by summarizing the...

  7. Crab Cavities: Past, Present, and Future of a Challenging Device

    CERN Document Server

    Wu, Q

    2015-01-01

    In two-ring facilities operating with a crossing-angle collision scheme, luminosity can be limited due to an incomplete overlapping of the colliding bunches. Crab cavities then are introduced to restore head-on collisions by providing the destined opposite deflection to the head and tail of the bunch. An increase in luminosity was demonstrated at KEKB with global crab- crossing, while the Large Hardron Collider (LHC) at CERN currently is designing local crab crossing for the Hi-Lumi upgrade. Future colliders may investigate both approaches. In this paper, we review the challenges in the technology, and the implementation of crab cavities, while discussing experience in earlier colliders, ongoing R&D, and proposed implementations for future facilities, such as HiLumi-LHC, CERN’s compact linear collider (CLIC), the international linear collider (ILC), and the electronion collider under design at BNL (eRHIC).

  8. Narrative report : 1969. Crab Orchard National Wildlife Refuge.

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — This annual narrative report for Crab Orchard National Wildlife Refuge outlines Refuge accomplishments during the 1969 calendar year. The report begins by...

  9. Narrative report : 1971. Crab Orchard National Wildlife Refuge.

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — This annual narrative report for Crab Orchard National Wildlife Refuge outlines Refuge accomplishments during the 1971 calendar year. The report begins by...

  10. Crab Orchard National Wildlife Refuge [Annual narrative : September - December, 1951

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — This narrative report for Crab Orchard National Wildlife Refuge outlines Refuge accomplishments from September through December of 1951. The report begins by...

  11. Crab Orchard National Wildlife Refuge [Narrative report : September - December, 1953

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — This narrative report for Crab Orchard National Wildlife Refuge outlines Refuge accomplishments from September through December of 1953. The report begins by...

  12. EVOLUTION OF THE CRAB NEBULA IN A LOW ENERGY SUPERNOVA

    Energy Technology Data Exchange (ETDEWEB)

    Yang, Haifeng; Chevalier, Roger A., E-mail: hy4px@virginia.edu, E-mail: rac5x@virginia.edu [Department of Astronomy, University of Virginia, P.O. Box 400325, Charlottesville, VA 22904 (United States)

    2015-06-20

    The nature of the supernova leading to the Crab Nebula has long been controversial because of the low energy that is present in the observed nebula. One possibility is that there is significant energy in extended fast material around the Crab but searches for such material have not led to detections. An electron capture supernova model can plausibly account for the low energy and the observed abundances in the Crab. Here, we examine the evolution of the Crab pulsar wind nebula inside a freely expanding supernova and find that the observed properties are most consistent with a low energy event. Both the velocity and radius of the shell material, and the amount of gas swept up by the pulsar wind point to a low explosion energy (∼10{sup 50} erg). We do not favor a model in which circumstellar interaction powers the supernova luminosity near maximum light because the required mass would limit the freely expanding ejecta.

  13. Evolution of the Crab nebula in a low energy supernova

    CERN Document Server

    Yang, Haifeng

    2015-01-01

    The nature of the supernova leading to the Crab Nebula has long been controversial because of the low energy that is present in the observed nebula. One possibility is that there is significant energy in extended fast material around the Crab but searches for such material have not led to detections. An electron capture supernova model can plausibly account for the low energy and the observed abundances in the Crab. Here, we examine the evolution of the Crab pulsar wind nebula inside a freely expanding supernova and find that the observed properties are most consistent with a low energy event. Both the velocity and radius of the shell material, and the amount of gas swept up by the pulsar wind point to a low explosion energy ($\\sim 10^{50}$ ergs). We do not favor a model in which circumstellar interaction powers the supernova luminosity near maximum light because the required mass would limit the freely expanding ejecta.

  14. The surprising Crab pulsar and its nebula: a review.

    Science.gov (United States)

    Bühler, R; Blandford, R

    2014-06-01

    The Crab nebula and its pulsar (referred to together as 'the Crab') have historically played a central role in astrophysics. True to this legacy, several unique discoveries have been made recently. The Crab was found to emit gamma-ray pulsations up to energies of 400 GeV, beyond what was previously expected from pulsars. Strong gamma-ray flares, of durations of a few days, were discovered from within the nebula, while the source was previously expected to be stable in flux on these time scales. Here we review these intriguing and suggestive developments. In this context we give an overview of the observational properties of the Crab and our current understanding of pulsars and their nebulae.

  15. Crab Orchard National Wildlife Refuge [Annual narrative : September - December, 1949

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — This narrative report for Crab Orchard National Wildlife Refuge outlines Refuge accomplishments from September through December of 1949. The report begins by...

  16. Economic Assessment of the Atlantic Coast Horseshoe Crab Fishery

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — In this report, Industrial Economics, Incorporated (lEc) provides an assessment of the economic value of the Atlantic Coast horseshoe crab fishery. We accomplish...

  17. Crab Orchard National Wildlife Refuge Narrative report : November - December, 1946

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — This narrative report for Crab Orchard National Wildlife Refuge outlines Refuge accomplishments from November through December of 1946. The report begins by...

  18. Narrative report : 1964. Crab Orchard National Wildlife Refuge.

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — This annual narrative report for Crab Orchard National Wildlife Refuge outlines Refuge accomplishments during the 1964 calendar year. The report begins by...

  19. Narrative report : 1968. Crab Orchard National Wildlife Refuge.

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — This annual narrative report for Crab Orchard National Wildlife Refuge outlines Refuge accomplishments during the 1968 calendar year. The report begins by...

  20. Domoic acid excretion in dungeness crabs, razor clams and mussels.

    Science.gov (United States)

    Schultz, Irvin R; Skillman, Ann; Woodruff, Dana

    2008-07-01

    Domoic acid (DA) is a neurotoxic amino acid produced by several marine algal species of the Pseudo-nitzschia (PN) genus. We studied the elimination of DA from hemolymph after intravascular (IV) injection in razor clams (Siliqua patula), mussels (Mytilus edulis) and Dungeness crabs (Cancer magister). Crabs were also injected with two other organic acids, dichloroacetic acid (DCAA) and kainic acid (KA). For IV dosing, hemolymph was repetitively sampled and DA concentrations measured by HPLC-UV. Toxicokinetic analysis of DA in crabs suggested most of the injected dose remained within hemolymph compartment with little extravascular distribution. This observation is in sharp contrast to results obtained from clams and mussels which exhibited similarly large apparent volumes of distribution despite large differences in overall clearance. These findings suggest fundamentally different storage and elimination processes are occurring for DA between bivalves and crabs.

  1. Narrative report : 1965. Crab Orchard National Wildlife Refuge.

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — This annual narrative report for Crab Orchard National Wildlife Refuge outlines Refuge accomplishments during the 1965 calendar year. The report begins by...

  2. Crab Orchard National Wildlife Refuge [Narrative report : January - April, 1952

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — This narrative report for Crab Orchard National Wildlife Refuge outlines Refuge accomplishments from January through April of 1952. The report begins by summarizing...

  3. Crab Orchard National Wildlife Refuge [Annual narrative : May - August, 1954

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — This narrative report for Crab Orchard National Wildlife Refuge outlines Refuge accomplishments from May through August of 1954. The report begins by summarizing the...

  4. Crab Orchard National Wildlife Refuge [Annual narrative : May - August, 1955

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — This narrative report for Crab Orchard National Wildlife Refuge outlines Refuge accomplishments from May through August of 1955. The report begins by summarizing the...

  5. Crab Orchard National Wildlife Refuge [Annual narrative : January - April, 1956

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — This narrative report for Crab Orchard National Wildlife Refuge outlines Refuge accomplishments from January through April of 1956. The report begins by summarizing...

  6. Crab Orchard National Wildlife Refuge [Annual narrative : May - August, 1948

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — This narrative report for Crab Orchard National Wildlife Refuge outlines Refuge accomplishments from May through August of 1948. The report begins by summarizing the...

  7. Crab Orchard National Wildlife Refuge [Annual narrative : January - April, 1950

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — This narrative report for Crab Orchard National Wildlife Refuge outlines Refuge accomplishments from January through April of 1950. The report begins by summarizing...

  8. Crab Orchard National Wildlife Refuge [Annual narrative : September - December, 1948

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — This narrative report for Crab Orchard National Wildlife Refuge outlines Refuge accomplishments from September through December of 1948. The report begins by...

  9. The Trail Inventory of Crab Orchard NWR [Cycle 2

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — The purpose of this report is to create a baseline inventory of all non-motorized trails on Crab Orchard National Wildlife Refuge. Trails in this inventory are...

  10. Crab Orchard National Wildlife Refuge [Annual narrative : January - April, 1955

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — This narrative report for Crab Orchard National Wildlife Refuge outlines Refuge accomplishments from January through April of 1955. The report begins by summarizing...

  11. Crab Orchard National Wildlife Refuge : Law Enforcement Plan

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — The Crab Orchard National Wildlife Refuge Law Enforcement Plan clarifies U.S. Fish and Wildlife enforcement policies as they apply to the Refuge. It provides...

  12. Narrative report : 1966. Crab Orchard National Wildlife Refuge.

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — This annual narrative report for Crab Orchard National Wildlife Refuge outlines Refuge accomplishments during the 1966 calendar year. The report begins by...

  13. Crab Orchard National Wildlife Refuge : Wildlife Inventory Plan

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — The Crab Orchard National Wildlife Refuge Wildlife Inventory Plan outlines the strategy, techniques and purpose of a wildlife inventory on the Refuge. Futhermore the...

  14. Crab Orchard National Wildlife Refuge [Narrative report : January - April, 1954

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — This narrative report for Crab Orchard National Wildlife Refuge outlines Refuge accomplishments from January through April of 1954. The report begins by summarizing...

  15. Crab Orchard National Wildlife Refuge [Annual narrative : September - December, 1954

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — This narrative report for Crab Orchard National Wildlife Refuge outlines Refuge accomplishments from September through December of 1954. The report begins by...

  16. Crab Orchard National Wildlife Refuge [Annual narrative : September - December, 1955

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — This narrative report for Crab Orchard National Wildlife Refuge outlines Refuge accomplishments from September through December of 1955. The report begins by...

  17. Crab Orchard National Wildlife Refuge [Annual narrative : January - April, 1949

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — This narrative report for Crab Orchard National Wildlife Refuge outlines Refuge accomplishments from January through April of 1949. The report begins by summarizing...

  18. Crab Orchard National Wildlife Refuge [Annual narrative : January - April, 1948

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — This narrative report for Crab Orchard National Wildlife Refuge outlines Refuge accomplishments from January through April of 1948. The report begins by summarizing...

  19. Crab Orchard National Wildlife Refuge [Annual narrative : September - December, 1956

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — This narrative report for Crab Orchard National Wildlife Refuge outlines Refuge accomplishments from September through December of 1956. The report begins by...

  20. Crab Orchard National Wildlife Refuge [Annual narrative : May - August, 1951

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — This narrative report for Crab Orchard National Wildlife Refuge outlines Refuge accomplishments from May through August of 1951. The report begins by summarizing the...

  1. Crab Orchard National Wildlife Refuge : Wilderness Management Plan

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — The Crab Orchard National Wildlife Refuge Wilderness Management Plan is a general plan which outlines the Refuge management objectives, wilderness creation and...

  2. Narrative report : 1967. Crab Orchard National Wildlife Refuge.

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — This annual narrative report for Crab Orchard National Wildlife Refuge outlines Refuge accomplishments during the 1967 calendar year. The report begins by...

  3. Narrative report : 1972 [Crab Orchard National Wildlife Refuge

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — This annual narrative report for Crab Orchard National Wildlife Refuge outlines Refuge accomplishments during the 1972 calendar year. The report begins by...

  4. Crab Orchard National Wildlife Refuge [Annual narrative : January - April, 1957

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — This narrative report for Crab Orchard National Wildlife Refuge outlines Refuge accomplishments from January through April of 1957. The report begins by summarizing...

  5. Narrative report : 1973 [Crab Orchard National Wildlife Refuge

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — This annual narrative report for Crab Orchard National Wildlife Refuge outlines Refuge accomplishments during the 1973 fiscal year. The report begins by summarizing...

  6. Crab Orchard National Wildlife Refuge [Annual narrative : May - August, 1956

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — This narrative report for Crab Orchard National Wildlife Refuge outlines Refuge accomplishments from May through August of 1956. The report begins by summarizing the...

  7. Crab Orchard National Wildlife Refuge: Annual Narrative: Calender year 1997

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — This annual narrative report for Crab Orchard National Wildlife Refuge summarizes refuge activities during the 1997 calendar year. The report begins with an...

  8. Crab Orchard National Wildlife Refuge: Annual Narrative: Calender year 2002

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — This annual narrative report for Crab Orchard National Wildlife Refuge summarizes refuge activities during the 2002 calendar year. The report begins with an...

  9. Crab Orchard National Wildlife Refuge : Prescribed Fire Plan : 1995

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — The Crab Orchard National Wildlife Refuge Prescribed Fire Plan has been developed to meet the station objectives set forth in the Master Plan, by aiding the forest...

  10. Crab Orchard National Wildlife Refuge [Annual narrative : May - August, 1949

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — This narrative report for Crab Orchard National Wildlife Refuge outlines Refuge accomplishments from May through August of 1949. The report begins by summarizing the...

  11. Patterns of surface burrow plugging in a colony of black-tailed prairie dogs occupied by black-footed ferrets

    Science.gov (United States)

    Eads, D.A.; Biggins, D.E.

    2012-01-01

    Black-tailed prairie dogs (Cynomys ludovicianus) can surface-plug openings to a burrow occupied by a black-footed ferret (Mustela nigripes). At a coarse scale, surface plugs are more common in colonies of prairie dogs occupied by ferrets than in colonies without ferrets. However, little is known about spatial and temporal patterns of surface plugging in a colony occupied by ferrets. In a 452-ha colony of black-tailed prairie dogs in South Dakota, we sampled burrow openings for surface plugs and related those data to locations of ferrets observed during spotlight surveys. Of 67,574 burrow openings in the colony between June and September 2007, 3.7% were plugged. In a colony-wide grid of 80 m × 80 m cells, the occurrence of surface plugging (≥1 opening plugged) was greater in cells used by ferrets (93.3% of cells) than in cells not observably used by ferrets (70.6%). Rates of surface plugging (percentages of openings plugged) were significantly higher in cells used by ferrets (median = 3.7%) than in cells without known ferret use (median = 3.2%). Also, numbers of ferret locations in cells correlated positively with numbers of mapped surface plugs in the cells. To investigate surface plugging at finer temporal and spatial scales, we compared rates of surface plugging in 20-m-radius circle-plots centered on ferret locations and in random plots 1–4 days after observing a ferret (Jun–Oct 2007 and 2008). Rates of surface plugging were greater in ferret-plots (median = 12.0%) than in random plots (median = 0%). For prairie dogs and their associates, the implications of surface plugging could be numerous. For instance, ferrets must dig to exit or enter plugged burrows (suggesting energetic costs), and surface plugs might influence microclimates in burrows and consequently influence species that cannot excavate soil (e.g., fleas that transmit the plague bacterium Yersinia pests).

  12. Patterns of surface burrow plugging in a colony of black-tailed prairie dogs occupied by black-footed ferrets

    Science.gov (United States)

    Eads, David E.; Biggins, Dean E.

    2012-01-01

    Black-tailed prairie dogs (Cynomys ludovicianus) can surface-plug openings to a burrow occupied by a black-footed ferret (Mustela nigripes). At a coarse scale, surface plugs are more common in colonies of prairie dogs occupied by ferrets than in colonies without ferrets. However, little is known about spatial and temporal patterns of surface plugging in a colony occupied by ferrets. In a 452-ha colony of black-tailed prairie dogs in South Dakota, we sampled burrow openings for surface plugs and related those data to locations of ferrets observed during spotlight surveys. Of 67,574 burrow openings in the colony between June and September 2007, 3.7% were plugged. In a colony-wide grid of 80 m × 80 m cells, the occurrence of surface plugging (≥1 opening plugged) was greater in cells used by ferrets (93.3% of cells) than in cells not observably used by ferrets (70.6%). Rates of surface plugging (percentages of openings plugged) were significantly higher in cells used by ferrets (median = 3.7%) than in cells without known ferret use (median = 3.2%). Also, numbers of ferret locations in cells correlated positively with numbers of mapped surface plugs in the cells. To investigate surface plugging at finer temporal and spatial scales, we compared rates of surface plugging in 20-m-radius circle-plots centered on ferret locations and in random plots 1–4 days after observing a ferret (Jun–Oct 2007 and 2008). Rates of surface plugging were greater in ferret-plots (median = 12.0%) than in random plots (median = 0%). For prairie dogs and their associates, the implications of surface plugging could be numerous. For instance, ferrets must dig to exit or enter plugged burrows (suggesting energetic costs), and surface plugs might influence microclimates in burrows and consequently influence species that cannot excavate soil (e.g., fleas that transmit the plague bacterium Yersinia pestis).

  13. Linking Microbial Enzymatic Activities and Functional Diversity of Soil around Earthworm Burrows and Casts.

    Science.gov (United States)

    Lipiec, Jerzy; Frąc, Magdalena; Brzezińska, Małgorzata; Turski, Marcin; Oszust, Karolina

    2016-01-01

    The aim of this work was to evaluate the effect of earthworms (Lumbricidae) on the enzymatic activity and microbial functional diversity in the burrow system [burrow wall (BW) 0-3 mm, transitional zone (TZ) 3-7 mm, bulk soil (BS) > 20 mm from the BW] and cast aggregates of a loess soil under a pear orchard. The dehydrogenase, β-glucosidase, protease, alkaline phosphomonoesterase, and acid phosphomonoesterase enzymes were assessed using standard methods. The functional diversity (catabolic potential) was assessed using the Average Well Color Development and Richness Index following the community level physiological profiling from Biolog Eco Plates. All measurements were done using soil from each compartment immediately after in situ sampling in spring. The enzymatic activites including dehydrogenase, protease, β-glucosidase and alkaline phosphomonoesterase were appreciably greater in the BW or casts than in BS and TZ. Conversely, acid phosphomonoesterase had the largest value in the BS. Average Well Color Development in both the TZ and the BS (0.98-0.94 A590 nm) were more than eight times higher than in the BWs and casts. The lowest richness index in the BS (15 utilized substrates) increased by 86-113% in all the other compartments. The PC1 in principal component analysis mainly differentiated the BWs and the TZ. Utilization of all substrate categories was the lowest in the BS. The PC2 differentiated the casts from the other compartments. The enhanced activity of a majority of the enzymes and increased microbial functional diversity in most earthworm-influenced compartments make the soils less vulnerable to degradation and thus increases the stability of ecologically relevant processes in the orchard ecosystem.

  14. Crab scars reveal survival advantage of left-handed snails

    OpenAIRE

    Dietl, Gregory P.; Hendricks, Jonathan R.

    2006-01-01

    Biological asymmetries are important elements of the structure and function of many living organisms. Using the Plio–Pleistocene fossil record of crab predation on morphologically similar pairs of right- and left-handed snail species, we show here for the first time, contrary to traditional wisdom, that rare left-handed coiling promotes survival from attacks by right-handed crabs. This frequency-dependent result influences the balance of selection processes that maintain left-handedness at th...

  15. Crab pulsar timing 1982-87

    Science.gov (United States)

    Lyne, A. G.; Pritchard, R. S.; Smith, F. G.

    1988-08-01

    Observations of the arrival times of pulses from the pulsar in the Crab Nebula over a six-year interval are presented. The data are intended to permit the investigation of the interior of the neutron star through the study of glitches and timing noise and to provide an ephemeris for high-energy observations. The first and second frequency derivatives provide a value for the braking index of n = 2.509 + or - 0.001, which is consistent with previous observations. The third frequency derivative can now be determined over an 18-yr span and is as expected for this braking index. The predominant deviations from a simple slow-down model form a sinusoid with a period of 20 months, attributable to an oscillation of the bulk of the neutron superfluid in the pulsar. One conspicuous glitch occurred in August, 1986 and the subsequent recovery was studied from only one hour after the event.

  16. Lincoln Co. Scrap Metal, Crab Orchard, Kentucky

    Science.gov (United States)

    The City of Crab Orchard, KY (population less than 1,000) received a $200,000 EPA Brownfields cleanup grant in 2010 to cleanup up the Lincoln County ScrapMetal property. The site, a former scrap metal recycler and general junkyard, was located in the middle of downtown. The city has experienced a dramatic decline in growth over the past few years. The abandoned two-acre site is located in the city’s center, directly across the street from City Hall. It is the largest property on Main Street. The property was an eyesore, and posed potential health risks to area residents, and deterred investment. Its blighted status did little to help the commercial and private properties that surround it. The site was also home to a dilapidated building that once served as the Odd Fellows meeting hall.

  17. Dusty globules in the Crab Nebula

    Science.gov (United States)

    Grenman, T.; Gahm, G. F.; Elfgren, E.

    2017-03-01

    Context. Dust grains are widespread in the Crab Nebula. A number of small, dusty globules, are visible as dark spots against the background of continuous synchrotron emission in optical images. Aims: Our aim is to catalogue such dusty globules and investigate their properties. Methods: From existing broad-band images obtained with the Hubble Space Telescope, we located 92 globules, for which we derived positions, dimensions, orientations, extinctions, masses, proper motions, and their distributions. Results: The globules have mean radii ranging from 400 to 2000 AU and are not resolved in current infrared images of the nebula. The extinction law for dust grains in these globules matches a normal interstellar extinction law. Derived masses of dust range from 1 to 60 × 10-6M⊙, and the total mass contained in globules constitute a fraction of approximately 2% or less of the total dust content of the nebula. The globules are spread over the outer part of the nebula, and a fraction of them coincide in position with emission filaments, where we find elongated globules that are aligned with these filaments. Only 10% of the globules are coincident in position with the numerous H2-emitting knots found in previous studies. All globules move outwards from the centre with transversal velocities of 60 to 1600 km s-1, along with the general expansion of the remnant. We discuss various hypotheses for the formation of globules in the Crab Nebula. Based on observations collected with the NASA/ESA Hubble Space Telescope, obtained at the Space Telescope Science Institute.

  18. Cross-centre replication of suppressed burrowing behaviour as an ethologically relevant pain outcome measure in the rat: a prospective multicentre study

    Science.gov (United States)

    Wodarski, Rachel; Delaney, Ada; Ultenius, Camilla; Morland, Rosie; Andrews, Nick; Baastrup, Catherine; Bryden, Luke A.; Caspani, Ombretta; Christoph, Thomas; Gardiner, Natalie J.; Huang, Wenlong; Kennedy, Jeffrey D.; Koyama, Suguru; Li, Dominic; Ligocki, Marcin; Lindsten, Annika; Machin, Ian; Pekcec, Anton; Robens, Angela; Rotariu, Sanziana M.; Voß, Sabrina; Segerdahl, Marta; Stenfors, Carina; Svensson, Camilla I.; Treede, Rolf-Detlef; Uto, Katsuhiro; Yamamoto, Kazumi; Rutten, Kris; Rice, Andrew S.C.

    2016-01-01

    Abstract Burrowing, an ethologically relevant rodent behaviour, has been proposed as a novel outcome measure to assess the global impact of pain in rats. In a prospective multicentre study using male rats (Wistar, Sprague-Dawley), replication of suppressed burrowing behaviour in the complete Freund adjuvant (CFA)-induced model of inflammatory pain (unilateral, 1 mg/mL in 100 µL) was evaluated in 11 studies across 8 centres. Following a standard protocol, data from participating centres were collected centrally and analysed with a restricted maximum likelihood-based mixed model for repeated measures. The total population (TP—all animals allocated to treatment; n = 249) and a selected population (SP—TP animals burrowing over 500 g at baseline; n = 200) were analysed separately, assessing the effect of excluding “poor” burrowers. Mean baseline burrowing across studies was 1113 g (95% confidence interval: 1041-1185 g) for TP and 1329 g (1271-1387 g) for SP. Burrowing was significantly suppressed in the majority of studies 24 hours (7 studies/population) and 48 hours (7 TP, 6 SP) after CFA injections. Across all centres, significantly suppressed burrowing peaked 24 hours after CFA injections, with a burrowing deficit of −374 g (−479 to −269 g) for TP and −498 g (−609 to −386 g) for SP. This unique multicentre approach first provided high-quality evidence evaluating suppressed burrowing as robust and reproducible, supporting its use as tool to infer the global effect of pain on rodents. Second, our approach provided important informative value for the use of multicentre studies in the future. PMID:27643836

  19. AFSC/RACE/SAP/Long: Data from: Embryo development in golden king crab, Lithodes aequispina.

    Data.gov (United States)

    National Oceanic and Atmospheric Administration, Department of Commerce — The data from this study, describes embryo development in Golden king crab, Lithodes aequispinus. Six female multiparous golden king crab were captured from the...

  20. Spawning migration of the horseshoe crab, Tachypleus gigas (Muller), in relation to lunal cycle

    Digital Repository Service at National Institute of Oceanography (India)

    Chatterji, A.; Rathod, V.; Parulekar, A.H.

    Effects of lunar phases and tidal height on the spawning migration of the horseshoe crab, Tachypleus gigas, along the northeastern coast of India were studied. Mature pairs of crabs migrate towards the shore and build their nests in sandy beaches...

  1. 76 FR 4635 - Proposed Information Collection; Comment Request; Economic Expenditure Survey of Golden Crab...

    Science.gov (United States)

    2011-01-26

    ... Expenditure Survey of Golden Crab Fishermen in the U.S. South Atlantic Region AGENCY: National Oceanic and... National Marine Fisheries Service (NMFS) proposes to collect economic information from golden-crab...

  2. Super-Acceleration in the Flaring Crab Nebula

    Energy Technology Data Exchange (ETDEWEB)

    Tavani, Marco, E-mail: marco.tavani@inaf.it

    2013-10-15

    The Crab Nebula continues to surprise us. The Crab system (energized by a very powerful pulsar at the center of the Supernova Remnant SN1054) is known to be a very efficient particle “accelerator” which can reach PeV energies. Today, new surprising data concerning the gamma-ray flares produced by the Crab Nebula challenge models of particle acceleration. The total energy flux from the Crab has been considered for many decades substantially stable at X-ray and gamma-ray energies. However, this paradigm was shattered by the AGILE discovery and Fermi confirmation in September 2010 of transient gamma-ray emission from the Crab. Indeed, we can state that four major flaring gamma-ray episodes have been detected by AGILE and Fermi during the period mid-2007/2012. During these events, transient particle acceleration occurs in a regime which apparently violates the MHD conditions and synchrotron cooling constraints. This fact justifies calling “super-acceleration” the mechanism which produces the “flaring Crab phenomenon”. Radiation between 50 MeV and a few GeV is emitted with a quite hard spectrum within a short timescale (hours-days), with no obvious relation with simultaneous optical and X-ray emissions in the inner Nebula. “Super-acceleration” implies overcoming synchrotron cooling by strong (and “parallel”) electric fields most likely produced by magnetic field reconnection within the pulsar wind outflow. This acceleration appears to be very efficient and, remarkably, limited by radiation reaction. It is not clear at the moment where in the Nebula this phenomenon occurs. An intense observational program is now focused on the Crab Nebula to resolve its most challenging mystery.

  3. Design and prototyping of HL-LHC double quarter wave crab cavities for SPS test

    Energy Technology Data Exchange (ETDEWEB)

    Verdu-Andres, S. [Brookhaven National Lab. (BNL), Upton, NY (United States); Skaritka, J. [Brookhaven National Lab. (BNL), Upton, NY (United States); Wu, Q. [Brookhaven National Lab. (BNL), Upton, NY (United States); Xiao, B. [Brookhaven National Lab. (BNL), Upton, NY (United States); Belomestnykh, S. [Brookhaven National Lab. (BNL), Upton, NY (United States); Stony Brook Univ., NY (United States); Ben-Zvi, I. [Brookhaven National Lab. (BNL), Upton, NY (United States); Stony Brook Univ., NY (United States); Alberty, L. [European Organization for Nuclear Research (CERN), Geneva (Switzerland); Artoos, K. [European Organization for Nuclear Research (CERN), Geneva (Switzerland); Calaga, R. [European Organization for Nuclear Research (CERN), Geneva (Switzerland); Capatina, O. [European Organization for Nuclear Research (CERN), Geneva (Switzerland); Capelli, T. [European Organization for Nuclear Research (CERN), Geneva (Switzerland); Carra, F. [European Organization for Nuclear Research (CERN), Geneva (Switzerland); Leuxe, R. [European Organization for Nuclear Research (CERN), Geneva (Switzerland); Kuder, N. [European Organization for Nuclear Research (CERN), Geneva (Switzerland); Zanoni, C. [European Organization for Nuclear Research (CERN), Geneva (Switzerland); Li, Z. [SLAC National Accelerator Lab., Menlo Park, CA (United States); Ratti, A. [Lawrence Berkeley National Lab. (LBNL), Berkeley, CA (United States)

    2015-05-03

    The LHC high luminosity project envisages the use of the crabbing technique for increasing and levelling the LHC luminosity. Double Quarter Wave (DQW) resonators are compact cavities especially designed to meet the technical and performance requirements for LHC beam crabbing. Two DQW crab cavities are under fabrication and will be tested with beam in the Super Proton Synchrotron (SPS) at CERN by 2017. This paper describes the design and prototyping of the DQW crab cavities for the SPS test.

  4. Design and Prototyping of HL-LHC Double Quarter Wave Crab Cavities for SPS Test

    CERN Document Server

    Verdú-Andrés, S; Wu, Q; Xiao, B P; Belomestnykh, S; Ben-Zv, I; Alberty, L; Artoos, Kurt; Calaga, Rama; Capatina, Ofelia; Capelli, Teddy; Carra, Federico; Leuxe, Raphael; Kuder, Norbert; Zanoni, Carlo; Li, Z; Ratti, A

    2015-01-01

    The LHC high luminosity project envisages the use of the crabbing technique for increasing and levelling the LHC luminosity. Double Quarter Wave (DQW) resonators are compact cavities especially designed to meet the technical and performance requirements for LHC beam crabbing. Two DQW crab cavities are under fabrication and will be tested with beam in the Super Proton Synchrotron (SPS) at CERN by 2017. This paper describes the design and prototyping of the DQW crab cavities for the SPS test.

  5. Occurrence of faecal pellet-filled simple and composite burrows in cold seep carbonates: A glimpse of a complex benthic ecosystem

    Digital Repository Service at National Institute of Oceanography (India)

    Mazumdar, A; Joshi, R.K.; Peketi, A; Kocherla, M.

    indicates methane expulsion that resulted in the development of a cold seep ecosystem. Although cold seep communities are extensively reported from the rock record and modern seep sites, hardly any information is available on burrowing activity of the seep...

  6. Prevalence and abundance of fleas in black-tailed prairie dog burrows: implications for the transmission of plague (Yersinia pestis).

    Science.gov (United States)

    Salkeld, Dan J; Stapp, Paul

    2008-06-01

    Plague, the disease caused by the bacterium Yersinia pestis, can have devastating impacts on North American wildlife. Epizootics, or die-offs, in prairie dogs (Cynomys ludovicianus) occur sporadically and fleas (Siphonaptera) are probably important in the disease's transmission and possibly as maintenance hosts of Y. pestis between epizootics. We monitored changes in flea abundance in prairie dog burrows in response to precipitation, temperature, and plague activity in shortgrass steppe in northern Colorado. Oropsylla hirsuta was the most commonly found flea, and it increased in abundance with temperature. In contrast, Oropsylla tuberculata cynomuris declined with rising temperature. During plague epizootics, flea abundance in burrows increased and then subsequently declined after the extirpation of their prairie dog hosts.

  7. Hidden dichromatism in the Burrowing Parrot (Cyanoliseus patagonus) as revealed by spectrometric colour analysis

    OpenAIRE

    Masello, Juan F; Lubjuhn, Thomas; Quillfeldt, Petra

    2009-01-01

    Bird colour perception differs fundamentally from that of humans. Birds have more cone types in the retina, including UV or violet cones, which enable them to perceive a wider spectral range. Thus, human colour perception can be deceiving when assessing functional aspects of bird plumage coloration, such as the intensity of sexual selection. In this study we measured reflectance spectra of different plumage regions of male and female Burrowing Parrot (Cyanoliseus patagonus) individuals. Altho...

  8. Burrowing seabird effects on invertebrate communities in soil and litter are dominated by ecosystem engineering rather than nutrient addition.

    Science.gov (United States)

    Orwin, Kate H; Wardle, David A; Towns, David R; St John, Mark G; Bellingham, Peter J; Jones, Chris; Fitzgerald, Brian M; Parrish, Richard G; Lyver, Phil O'B

    2016-01-01

    Vertebrate consumers can be important drivers of the structure and functioning of ecosystems, including the soil and litter invertebrate communities that drive many ecosystem processes. Burrowing seabirds, as prevalent vertebrate consumers, have the potential to impact consumptive effects via adding marine nutrients to soil (i.e. resource subsidies) and non-consumptive effects via soil disturbance associated with excavating burrows (i.e. ecosystem engineering). However, the exact mechanisms by which they influence invertebrates are poorly understood. We examined how soil chemistry and plant and invertebrate communities changed across a gradient of seabird burrow density on two islands in northern New Zealand. Increasing seabird burrow density was associated with increased soil nutrient availability and changes in plant community structure and the abundance of nearly all the measured invertebrate groups. Increasing seabird densities had a negative effect on invertebrates that were strongly influenced by soil-surface litter, a positive effect on fungal-feeding invertebrates, and variable effects on invertebrate groups with diverse feeding strategies. Gastropoda and Araneae species richness and composition were also influenced by seabird activity. Generalized multilevel path analysis revealed that invertebrate responses were strongly driven by seabird engineering effects, via increased soil disturbance, reduced soil-surface litter, and changes in trophic interactions. Almost no significant effects of resource subsidies were detected. Our results show that seabirds, and in particular their non-consumptive effects, were significant drivers of invertebrate food web structure. Reductions in seabird populations, due to predation and human activity, may therefore have far-reaching consequences for the functioning of these ecosystems.

  9. Observations of the Crab Nebula and Pulsar with VERITAS

    CERN Document Server

    Celik, O

    2007-01-01

    Observations of the Crab Nebula have proven to be the best tool to calibrate and to characterize the performance of a Cherenkov telescope. Scientifically, it is interesting to measure the energy spectrum of the Crab Nebula close to the inverse-Compton peak where a deviation is expected from the power law seen at energies above 300 GeV. Additionally, it is important to search for pulsed emission from the Crab Pulsar at energies beyond the 10 GeV upper limit of the EGRET pulsar detection. Since current models predict a cut-off in pulsed emission between 10 and 100 GeV, measurements at energies close to this range may help to discriminate between them. We observed the Crab extensively in the 2006-2007 season during the VERITAS 2- and 3-telescope commissioning phases. Using this data set we reconstructed a preliminary energy spectrum of the signal from the Crab Nebula. We also measured the optical pulsed signal to validate our GPS time-stamping and barycentering techniques and obtained an upper limit for the puls...

  10. Fossil worm burrows reveal very early terrestrial animal activity and shed light on trophic resources after the end-cretaceous mass extinction.

    Directory of Open Access Journals (Sweden)

    Karen Chin

    Full Text Available The widespread mass extinctions at the end of the Cretaceous caused world-wide disruption of ecosystems, and faunal responses to the one-two punch of severe environmental perturbation and ecosystem collapse are still unclear. Here we report the discovery of in situ terrestrial fossil burrows from just above the impact-defined Cretaceous-Paleogene (K/Pg boundary in southwestern North Dakota. The crisscrossing networks of horizontal burrows occur at the interface of a lignitic coal and silty sandstone, and reveal intense faunal activity within centimeters of the boundary clay. Estimated rates of sedimentation and coal formation suggest that the burrows were made less than ten thousand years after the end-Cretaceous impact. The burrow characteristics are most consistent with burrows of extant earthworms. Moreover, the burrowing and detritivorous habits of these annelids fit models that predict the trophic and sheltering lifestyles of terrestrial animals that survived the K/Pg extinction event. In turn, such detritus-eaters would have played a critical role in supporting secondary consumers. Thus, some of the carnivorous vertebrates that radiated after the K/Pg extinction may owe their evolutionary success to thriving populations of earthworms.

  11. Incidence of plastic fragments among burrow-nesting seabird colonies on offshore islands in northern New Zealand.

    Science.gov (United States)

    Buxton, Rachel T; Currey, Caitlin A; Lyver, Philip O'B; Jones, Christopher J

    2013-09-15

    Marine plastic pollution is ubiquitous throughout the world's oceans, and has been found in high concentrations in oceanic gyres of both the northern and southern hemispheres. The number of studies demonstrating plastic debris at seabird colonies and plastic ingestion by adult seabirds has increased over the past few decades. Despite the recent discovery of a large aggregation of plastic debris in the South Pacific subtropical gyre, the incidence of plastics at seabird colonies in New Zealand is unknown. Between 2011 and 2012 we surveyed six offshore islands on the northeast coast of New Zealand's North Island for burrow-nesting seabird colonies and the presence of plastic fragments. We found non-research related plastic fragments (0.031 pieces/m(2)) on one island only, Ohinau, within dense flesh-footed shearwater (Puffinus carneipes) colonies. On Ohinau, we found a linear relationship between burrow density and plastic density, with 3.5 times more breeding burrows in areas with plastic fragments found. From these data we conclude that plastic ingestion is a potentially a serious issue for flesh-footed shearwaters in New Zealand. Although these results do not rule out plastic ingestion by other species, they suggest the need for further research on the relationship between New Zealand's pelagic seabirds and marine plastic pollution.

  12. Synthesis of Pt nanoparticles and their burrowing into Si due to synergistic effects of ion beam energy losses

    Directory of Open Access Journals (Sweden)

    Pravin Kumar

    2014-10-01

    Full Text Available We report the synthesis of Pt nanoparticles and their burrowing into silicon upon irradiation of a Pt–Si thin film with medium-energy neon ions at constant fluence (1.0 × 1017 ions/cm2. Several values of medium-energy neon ions were chosen in order to vary the ratio of the electronic energy loss to the nuclear energy loss (Se/Sn from 1 to 10. The irradiated films were characterized using Rutherford backscattering spectroscopy (RBS, atomic force microscopy (AFM, scanning electron microscopy (SEM, X-ray diffraction (XRD and high resolution transmission electron microscopy (HRTEM. A TEM image of a cross section of the film irradiated with Se/Sn = 1 shows ≈5 nm Pt NPs were buried up to ≈240 nm into the silicon. No silicide phase was detected in the XRD pattern of the film irradiated at the highest value of Se/Sn. The synergistic effect of the energy losses of the ion beam (molten zones are produced by Se, and sputtering and local defects are produced by Sn leading to the synthesis and burrowing of Pt NPs is evidenced. The Pt NP synthesis mechanism and their burrowing into the silicon is discussed in detail.

  13. H2 Temperatures in the Crab Nebula

    CERN Document Server

    Loh, E D; Ferland, G J; Curtis, Z K; Richardson, C T; Fabian, A C; Salomé, P

    2011-01-01

    We used K-band spectra to measure the H2 excitation temperatures in six molecular knots associated with the filaments in the Crab Nebula. The temperatures are quite high - in the range T ~ 2000-3000K, just below the H2 dissociation temperature. This is the temperature range over which the H2 1-0 S(1) line at 2.121\\mum has its maximum emissivity per unit mass, so there may be many additional H2 cores with lower temperatures that are too faint to detect. We also measured the electron density in adjacent ionized gas, which on the assumption of gas pressure balance indicates densities in the molecular region n_mol ~ 20,000 H baryons cm-3, although this really is just a lower limit since the H2 gas may be confined by other means. The excited region may be just a thin skin on a much more extensive blob of molecular gas that does not have the correct temperature and density to be as easily detectable. At the opposite extreme, the observed knots could consist of a fine mist of molecular gas in which we are detecting ...

  14. Radio Emission Physics in the Crab Pulsar

    CERN Document Server

    Eilek, J A

    2016-01-01

    We review our high-time-resolution radio observations of the Crab pulsar and compare our data to a variety of models for the emission physics. The Main Pulse and the Low-Frequency Interpulse come from regions somewhere in the high-altitude emission zones (caustics) that also produce pulsed X-ray and gamma-ray emission. Although no emission model can fully explain these two components, the most likely models suggest they arise from a combination of beam-driven instabilities, coherent charge bunching and strong electromagnetic turbulence. Because the radio power fluctuates on a wide range of timescales, we know the emission zones are patchy and dynamic. It is tempting to invoke unsteady pair creation in high-altitude gaps as source of the variability, but current pair cascade models cannot explain the densities required by any of the likely models. It is harder to account for the mysterious High-Frequency Interpulse. We understand neither its origin within the magnetosphere nor the striking emission bands in it...

  15. Large-scale compression of genomic sequence databases with the Burrows-Wheeler transform

    CERN Document Server

    Cox, Anthony J; Jakobi, Tobias; Rosone, Giovanna

    2012-01-01

    Motivation The Burrows-Wheeler transform (BWT) is the foundation of many algorithms for compression and indexing of text data, but the cost of computing the BWT of very large string collections has prevented these techniques from being widely applied to the large sets of sequences often encountered as the outcome of DNA sequencing experiments. In previous work, we presented a novel algorithm that allows the BWT of human genome scale data to be computed on very moderate hardware, thus enabling us to investigate the BWT as a tool for the compression of such datasets. Results We first used simulated reads to explore the relationship between the level of compression and the error rate, the length of the reads and the level of sampling of the underlying genome and compare choices of second-stage compression algorithm. We demonstrate that compression may be greatly improved by a particular reordering of the sequences in the collection and give a novel `implicit sorting' strategy that enables these benefits to be re...

  16. Natural AD-Like Neuropathology in Octodon degus: Impaired Burrowing and Neuroinflammation.

    Science.gov (United States)

    Deacon, Robert M J; Altimiras, Francisco J; Bazan-Leon, Enrique A; Pyarasani, Rhada D; Nachtigall, Fabiane M; Santos, Leonardo S; Tsolaki, Anthony G; Pednekar, Lina; Kishore, Uday; Biekofsky, Rodolfo R; Vasquez, Rodrigo A; Cogram, Patricia

    2015-01-01

    Alzheimer's disease (AD) is the most common cause of dementia, affecting more than 36 million people worldwide. Octodon degus, a South American rodent, has been found to spontaneously develop neuropathological signs of AD, including amyloid-β (Aβ) and tau deposits, as well as a decline in cognition with age. Firstly, the present work introduces a novel behavioral assessment for O. degus - the burrowing test - which appears to be a useful tool for detecting neurodegeneration in the O. degus model for AD. Such characterization has potentially wide-ranging implications, because many of these changes in species-typical behaviors are reminiscent of the impairments in activities of daily living (ADL), so characteristic of human AD. Furthermore, the present work characterizes the AD-like neuropathology in O. degus from a gene expression point of view, revealing a number of previously unreported AD biomarkers, which are found in human AD: amyloid precursor protein (APP), apolipoprotein E (ApoE), oxidative stress-related genes from the NFE2L2 and PPAR pathway, as well as pro-inflammatory cytokines and complement proteins, in agreement with the known link between neurodegeneration and neuroinflammation. In summary, the present results confirm a natural neuropathology in O. degus with similar characteristics to AD at behavioral, cellular and molecular levels. These characteristics put O. degus in a singular position as a natural rodent model for research into AD pathogenesis and therapeutics against AD.

  17. Enzyme activity in the aestivating green-striped burrowing frog (Cyclorana alboguttata).

    Science.gov (United States)

    Mantle, Beth L; Guderley, Helga; Hudson, Nicholas J; Franklin, Craig E

    2010-10-01

    Green-striped burrowing frogs (Cyclorana alboguttata) can depress their resting metabolism by more than 80% during aestivation. Previous studies have shown that this species is able to withstand long periods of immobilisation during aestivation while apparently maintaining whole muscle mass and contractile performance. The aim of this study was to determine the effect of prolonged aestivation on the levels of metabolic enzymes (CCO, LDH and CS) in functionally distinct skeletal muscles (cruralis, gastrocnemius, sartorius, iliofibularis and rectus abdominus) and liver of C. alboguttata. CS activity was significantly reduced in all tissues except for the cruralis, gastrocnemius and the liver. LDH activity was significantly reduced in the sartorius and rectus abdominus, but remained at control (active) levels in the other tissues. CCO activity was significantly reduced in the gastrocnemius and rectus abdominus, and unchanged in the remaining tissues. Muscle protein was significantly reduced in the sartorius and iliofibularis during aestivation, and unchanged in the remaining muscles. The results suggest that the energy pathways involved in the production and consumption of ATP are remodelled during prolonged aestivation but selective. Remodelling and subsequent down-regulation of metabolic activity seem to target the smaller non-jumping muscles, while the jumping muscles retain enzyme activities at control levels during aestivation. These results suggest a mechanism by which aestivating C. alboguttata are able to maintain metabolic depression while ensuring that the functional capacity of critical muscles is not compromised upon emergence from aestivation.

  18. Phylogenetic relationships of Darwin's "Mr. Arthrobalanus": The burrowing barnacles (Cirripedia: Acrothoracica).

    Science.gov (United States)

    Lin, Hsiu-Chin; Kobasov, Gregory A; Chan, Benny K K

    2016-07-01

    The barnacles of the superorder Acrothoracica are small, burrowing, epibiotic, and dioecious (large female with dwarf male) crustaceans largely found in the carbonate sediments and skeletons of marine invertebrates. The acrothoracicans represent the Cirripedia with the most plesiomorphic characters and have prominently featured in phylogenetic speculations concerning these crustaceans. Traditionally, Acrothoracica was divided into two main orders, Pygophora and Apygophora. The Apygophora had uniramus cirri and no anus. The Pygophora had biramus terminal cirri and an anus and was further divided into two families, Lithoglyptidae and Cryptophialidae. Kolbasov (2009) revised the superorder Acrothoracica on the basis of morphological examinations of females, dwarf males, and cyprids and rearranged the acrothoracican species into two new orders, Lithoglyptida and Cryptophialida. The present study is the first attempt to reconstruct the phylogenetic relationships of acrothoracican barnacles by sequencing two mitochondrial (cytochrome C oxidase I and 16S ribosomal DNA) and two nuclear (18S ribosomal DNA and histone H3) markers of 8 of the 11 genera comprising 23 acrothoracican species. All monophylies of the eight acrothoracican genera sampled in this study were strongly supported. The deep interfamilial relationship constructed is consistent with the recent morphological phylogenetic relationship proposed by Kolbasov, Newman, and Høeg (Kolbasov, 2009) that Cryptophialidae (order Cryptophialida) is the sister group to all other acrothoracicans (order Lithoglyptida). According to an ancestral character state reconstruction analysis, the posterior lobes of females; armament of opercular bars, attachment stalk, lateral projections of the body, and aperture slits in dwarf males; and habitat use appear to have phylogenetic importance.

  19. Lossless compression of 3D hyperspectral sounder data using the wavelet and Burrows-Wheeler transforms

    Science.gov (United States)

    Wei, Shih-Chieh; Huang, Bormin

    2004-10-01

    Hyperspectral sounder data is used for retrieval of useful geophysical parameters which promise better weather prediction. It features two characteristics. First it is huge in size with 2D spatial coverage and high spectral resolution in the infrared region. Second it allows low tolerance of noise and error in retrieving the geophysical parameters where a mathematically ill-posed problem is involved. Therefore compression is better to be lossless or near lossless for data transfer and archive. Meanwhile medical data from X-ray computerized tomography (CT) or magnetic resonance imaging (MRI) techniques also possesses similar characteristics. It provides motivation to apply lossless compression schemes for medical data to the hyperspectral sounder data. In this paper, we explore the use of a wavelet-based lossless data compression scheme for the 3D hyperspectral data which uses in sequence a forward difference scheme, an integer wavelet transform, a Burrows-Wheeler transform and an arithmetic coder. Compared to previous work, our approach is shown to outperform the CALIC and 3D EZW schemes.

  20. Pre-Migratory Movements by Juvenile Burrowing Owls in a Patchy Landscape

    Directory of Open Access Journals (Sweden)

    L. Danielle. Todd

    2007-12-01

    Full Text Available Dispersal is a fundamental aspect of population dynamics, and can have direct implications on processes such as the colonization of habitat patches. Pre-migratory movements, landscape fragmentation, and body condition have all been hypothesized as key factors influencing dispersal in birds, but little direct evidence exists to support these ideas. We used radio-telemetry and supplementary feeding to test if body condition or landscape pattern influenced pre-migratory movements of juvenile Burrowing Owls (Athene cunicularia in a fragmented landscape. We categorized grassland patches as either large (≥95 ha or small and isolated (≤58 ha and ≥1.5 km to next nearest grassland patch, and young owls were either provided supplemental food as nestlings or not. Owlets receiving supplemental food and residing in large grassland patches moved a greater maximum distance from their nest than similarly fed owlets residing in small patches (large = 1605 ± 443 m; small = 373 ± 148 m. In contrast, non-supplemented owlets from large and small patches did not differ in their maximum distance moved from the nest (large = 745 ± 307 m; small 555 ± 286 m. Only two of 32 individuals from small patches moved >800 m, whereas ten of 23 owlets from large patches moved >800 m. In addition, owlets from large patches continued to move farther and farther from their nest before migration, whereas owlets in small, isolated patches ultimately moved

  1. Population dynamics of the pea crab Austinixa aidae (Brachyura, Pinnotheridae: a symbiotic of the ghost shrimp Callichirus major (Thalassinidea, Callianassidae from the southwestern Atlantic

    Directory of Open Access Journals (Sweden)

    Douglas F Peiró

    2011-06-01

    Full Text Available The Pinnotheridae family is one of the most diverse and complex groups of brachyuran crabs, many of them symbionts of a wide variety of invertebrates. The present study describes the population dynamics of the pea crab Austinixa aidae (Righi, 1967, a symbiont associated with the burrows of the ghost shrimp Callichirus major (Say, 1818. Individuals (n = 588 were collected bimonthly from May, 2005 to September, 2006 along a sandy beach in the southwestern Atlantic, state of São Paulo, Brazil. Our data indicated that the population demography of A. aidae was characterized by a bimodal size-frequency distribution (between 2.0 and 4.0 mm and between 8.0 and 9.0 mm CW that remained similar throughout the study period. Sex ratio does not differ significantly from 1:1 (p > 0.05, which confirms the pattern observed in other symbiontic pinnotherids. Density values (1.72 ± 1.34 ind. • ap.-1 are in agreement with those found for other species of the genus. The mean symbiosis incidence (75.6% was one of the highest among species of the Pinnotheridae family, but it was the lowest among the three studied species of the genus. Recruitment pattern was annual, beginning in May and peaking in July, in both years, after the peak of ovigerous females in the population (from March to May. Our findings describe ecological and biological aspects of A. aidae similar to those of other species of this genus, even from different geographic localities.

  2. 75 FR 49420 - Fisheries of the Northeastern United States; Atlantic Deep-Sea Red Crab Fisheries; 2010 Atlantic...

    Science.gov (United States)

    2010-08-13

    ... United States; Atlantic Deep-Sea Red Crab Fisheries; 2010 Atlantic Deep-Sea Red Crab Specifications In... (TAC) and corresponding fleet days-at-sea (DAS) allocation for the Atlantic deep- sea red crab fishery... the implementing regulations for the Atlantic Deep- Sea Red Crab Fishery Management Plan...

  3. 75 FR 7435 - Fisheries of the Northeastern United States; Atlantic Deep-Sea Red Crab Fisheries; 2010 Atlantic...

    Science.gov (United States)

    2010-02-19

    ... United States; Atlantic Deep-Sea Red Crab Fisheries; 2010 Atlantic Deep-Sea Red Crab Specifications... Atlantic deep-sea red crab fishery, including a target total allowable catch (TAC) and a fleet-wide days-at-sea (DAS) allocation. The implementing regulations for the Atlantic Deep-Sea Red Crab...

  4. 75 FR 35435 - Fisheries of the Northeastern United States; Atlantic Deep-Sea Red Crab Fisheries; 2010 Atlantic...

    Science.gov (United States)

    2010-06-22

    ... United States; Atlantic Deep-Sea Red Crab Fisheries; 2010 Atlantic Deep-Sea Red Crab Specifications In... finalized 2010 specifications for the Atlantic deep-sea red crab fishery, including a target total allowable... Atlantic Deep-Sea Red Crab Fishery Management Plan (FMP) allow NMFS to make an in-season adjustment to...

  5. 76 FR 47155 - Fisheries of the Exclusive Economic Zone Off Alaska; Bering Sea and Aleutian Islands Crab...

    Science.gov (United States)

    2011-08-04

    ... Alaska; Bering Sea and Aleutian Islands Crab Rationalization Program; Public Meeting AGENCY: National... crab fisheries managed under the BSAI Crab Rationalization program. The CIE, operated by Northern Taiga... products. The BSAI Crab Economic Data Report (EDR) program administered by NMFS began collecting...

  6. 77 FR 44216 - Fisheries of the Exclusive Economic Zone Off Alaska; Bering Sea and Aleutian Islands Crab...

    Science.gov (United States)

    2012-07-27

    ... Alaska; Bering Sea and Aleutian Islands Crab Rationalization Cost Recovery Program AGENCY: National... recovery under the Bering Sea and Aleutian Islands Crab Rationalization Program. This action is intended to provide holders of crab allocations with the fee percentage for the 2012/2013 crab fishing year....

  7. 76 FR 43658 - Fisheries of the Exclusive Economic Zone Off Alaska; Bering Sea and Aleutian Islands Crab...

    Science.gov (United States)

    2011-07-21

    ... Alaska; Bering Sea and Aleutian Islands Crab Rationalization Cost Recovery Program AGENCY: National... under the Bering Sea and Aleutian Islands Crab Rationalization Program. This action is intended to provide holders of crab allocations with the fee percentage for the 2011/2012 crab fishing year so...

  8. 75 FR 43147 - Fisheries of the Exclusive Economic Zone Off Alaska; Bering Sea and Aleutian Islands Crab...

    Science.gov (United States)

    2010-07-23

    ... Alaska; Bering Sea and Aleutian Islands Crab Rationalization Cost Recovery Program AGENCY: National... under the Bering Sea and Aleutian Islands Crab Rationalization Program. This action is intended to provide holders of crab allocations with the fee percentage for the 2010/2011 crab fishing year so...

  9. 78 FR 46577 - Fisheries of the Exclusive Economic Zone Off Alaska; Bering Sea and Aleutian Islands Crab...

    Science.gov (United States)

    2013-08-01

    ... Alaska; Bering Sea and Aleutian Islands Crab Rationalization Cost Recovery Program AGENCY: National... under the Bering Sea and Aleutian Islands Crab Rationalization Program. This action is intended to provide holders of crab allocations with the fee percentage for the 2013/2014 crab fishing year so...

  10. Extreme Particle Acceleration via Magnetic Reconnection in the Crab Nebula

    Science.gov (United States)

    Cerutti, Benoit; Uzdensky, D. A.; Begelman, M. C.

    2012-01-01

    The discovery by Agile and Fermi of intense day-long synchrotron gamma-ray flares above 100 MeV in the Crab Nebula challenges classical models of pulsar wind nebulae and particle acceleration. We argue that the flares are powered by magnetic reconnection in the nebula. Using relativistic test-particle simulations, we show that particles are naturally focused into a thin fan beam, deep inside the reconnection layer where the magnetic field is small. The particles then suffer less from synchrotron losses and pile up at the maximum energy given by the electric potential drop in the layer. Applying this model to the Crab Nebula, we find that the emerging synchrotron emission spectrum above 100 MeV is consistent with the September 2010 flare observations. No detectable emission is expected at other wavelengths. This scenario provides a viable explanation for the Crab Nebula gamma-ray flares.

  11. A New HST Measurement of the Crab Pulsar Proper Motion

    CERN Document Server

    Caraveo, P A; Caraveo, Patrizia A.; Mignani, Roberto P.

    1999-01-01

    We have used a set of archived HST/WFPC2 exposures of the inner regions of the Crab Nebula to obtain a new measurement of the pulsar proper motion, the first after the original work of Wyckoff & Murray, more than 20 years ago. Our measurement of the pulsar displacement, mu = 18 +/- 3 mas/yr, agrees well with the value obtained previously. This is noteworthy, since the data we have used span less than 2 years, as opposed to the 77 years required in the previous work. With a position angle of 292 +/- 10 deg, the proper motion vector appears aligned with the axis of symmetry of the inner Crab nebula, as defined by the direction of the X-ray jet observed by ROSAT. Indeed, if the neutron star rotation is to be held responsible both for the X-ray jet and for the observed symmetry, the Crab could provide an example of alignment between spin axis and proper motion.

  12. CRAB: the CMS distributed analysis tool development and design

    Energy Technology Data Exchange (ETDEWEB)

    Spiga, D. [University and INFN Perugia (Italy); Lacaprara, S. [INFN Legnaro (Italy); Bacchi, W. [University and INFN Bologna (Italy); Cinquilli, M. [University and INFN Perugia (Italy); Codispoti, G. [University and INFN Bologna (Italy); Corvo, M. [CERN (Switzerland); Dorigo, A. [INFN Padova (Italy); Fanfani, A. [University and INFN Bologna (Italy); Fanzago, F. [CERN (Switzerland); Farina, F. [INFN Milano-Bicocca (Italy); Gutsche, O. [FNAL (United States); Kavka, C. [INFN Trieste (Italy); Merlo, M. [INFN Milano-Bicocca (Italy); Servoli, L. [University and INFN Perugia (Italy)

    2008-03-15

    Starting from 2007 the CMS experiment will produce several Pbytes of data each year, to be distributed over many computing centers located in many different countries. The CMS computing model defines how the data are to be distributed such that CMS physicists can access them in an efficient manner in order to perform their physics analysis. CRAB (CMS Remote Analysis Builder) is a specific tool, designed and developed by the CMS collaboration, that facilitates access to the distributed data in a very transparent way. The tool's main feature is the possibility of distributing and parallelizing the local CMS batch data analysis processes over different Grid environments without any specific knowledge of the underlying computational infrastructures. More specifically CRAB allows the transparent usage of WLCG, gLite and OSG middleware. CRAB interacts with both the local user environment, with CMS Data Management services and with the Grid middleware.

  13. CRAB: the CMS distributed analysis tool development and design

    CERN Document Server

    Spiga, D; Bacchi, W; Cinquilli, M; Codispoti, G; Corvo, M; Dorigo, A; Fanfani, A; Fanzago, F; Farina, F; Gutsche, O; Kavka, C; Merlo, M; Servoli, L

    2008-01-01

    Starting from 2007 the CMS experiment will produce several Pbytes of data each year, to be distributed over many computing centers located in many different countries. The CMS computing model defines how the data are to be distributed such that CMS physicists can access them in an efficient manner in order to perform their physics analysis. CRAB (CMS Remote Analysis Builder) is a specific tool, designed and developed by the CMS collaboration, that facilitates access to the distributed data in a very transparent way. The tool's main feature is the possibility of distributing and parallelizing the local CMS batch data analysis processes over different Grid environments without any specific knowledge of the underlying computational infrastructures. More specifically CRAB allows the transparent usage of WLCG, gLite and OSG middleware. CRAB interacts with both the local user environment, with CMS Data Management services and with the Grid middleware.

  14. Respiration and heart rate in exercising land crabs.

    Science.gov (United States)

    Herreid, C F; Lee, L W; Shah, G M

    1979-05-01

    Land Crabs, Cardisoma guanhumi, were fitted with respiratory masks and E.C.G. electrodes and run for 10 or 20 min on a treadmill at speeds of 150 and 300 cm/min. Aerobic metabolism increased linearly with the speed of locomotion. The recovery period was characterized by a large oxygen debt. The primary respiratory adjustment to exercise was an increased ventilation volume; only a minor increase in oxygen extraction occurred. The respiratory exchange ratio increased during exercise and during recovery, presumably correlated with a metabolic acidosis. These results are similar to data collected for exercising vertebrates and the net cost of locomotion of crabs appears similar to quadrupeds. However, the heart rate in exercising crabs changed in an unexpected way: during moderate exercise no change was noted, but during heavy exercise a bradycardia developed. The reduction in rate resulted from an increase in interbeat interval and frequent pauses in the heart beat.

  15. Design and Prototyping of a 400 MHz RF-dipole Crabbing Cavity for the LHC High-Luminosity Upgrade

    CERN Document Server

    De Silva, S U; Delayen, J R; Li, Z; Nicol, T H

    2015-01-01

    LHC High Luminosity Upgrade is in need of two crabbing systems that deflects the beam in both horizontal and vertical planes. The 400 MHz rf-dipole crabbing cavity system is capable of crabbing the proton beam in both planes. At present we are focusing our efforts on a complete crabbing system in the horizontal plane. Prior to LHC installation the crabbing system will be installed for beam test at SPS. The crabbing system consists of two rfdipole cavities in the cryomodule. This paper discusses the electromagnetic design and mechanical properties of the rf-dipole crabbing system for SPS beam test.

  16. Preliminary survey of a nemertean crab egg predator, Carcinonemertes, on its host crab, Callinectes arcuatus (Decapoda, Portunidae) from Golfo de Nicoya, Pacific Costa Rica

    Science.gov (United States)

    Okazaki, Robert K.; Wehrtmann, Ingo S.

    2014-01-01

    Abstract The possible presence of egg predators in brood masses of portunid crabs from Pacific Central America has not been studied yet. This survey reports the finding of a nemertean crab egg predator on the portunid crab, Callinectes arcuatus, from the Golfo de Nicoya, Pacific Costa Rica. Nemerteans were found in the egg masses of 26 out of the 74 crabs for a prevalence of 35%. The intensity (mean number of worms/ infected crab) was estimated to be 18 with a variance of 1–123 worms/infected crab. No nemerteans were observed either in the 19 Callinectes arcuatus from Golfo Dulce (southern Pacific coast) and the 10 Portunus asper from Herradura-Jaco (central Pacific coast). This nemertean is a member of the genus Carcinonemertes, which has been reported from the Caribbean coast of Panama. However, the encountered Carcinonemertes sp. is the first published finding and report from Costa Rica and Pacific Central America. PMID:25561848

  17. Biodiversity of Crabs in Pichavaram Mangrove Environment, South East Coast of India

    Directory of Open Access Journals (Sweden)

    T. Kannupandi

    2008-01-01

    Full Text Available In this study, the distribution of crabs in Pichavaram was recorded from December 2001 to November 2002 at monthly intervals. The species present on the substratum and on the vegetation area were recorded in quadrant each measuring 1 m2. There are about 36 crabs species are distributed in Pichavaram mangrove environment. Crabs belonging to the family Grapsidae and Ocypodidae are most dominant forms. Substrate suitability; effects of tidal inundation and distribution of mangrove plants were the possible factors that could influence zonation and abundance of the crabs in the Pichavaram mangroves. The reason for depletion of crabs and their conservation measures are also discussed.

  18. Complete mitochondrial genomes of three mitten crabs, Eriocheir sinensis, E. hepuensis, and E. japonica.

    Science.gov (United States)

    Wang, Jun; Huang, Lei; Cheng, Qixuan; Lu, Guoqing; Wang, Chenghui

    2016-01-01

    Taxonomic classification of three mitten crabs (Eriocheir sinensis, E. hepuensis, and E. japonica) has long been controversial. In this study, the complete mitogenomes of the three crabs were reported. The three mitogenomes were conserved in the organization of genes with 13 protein-coding genes, 2 rRNA genes, 22 tRNA genes, and 1 control region. Nucleotide variations among the crabs were identified in both coding and non-coding regions. In addition, variable numbers of tandem repeats in control region were identified in the mitten crabs. The mitogenome sequences provide a valuable resource to elucidate taxonomic relationship and evolutionary history of the three mitten crabs.

  19. Crab scars reveal survival advantage of left-handed snails.

    Science.gov (United States)

    Dietl, Gregory P; Hendricks, Jonathan R

    2006-09-22

    Biological asymmetries are important elements of the structure and function of many living organisms. Using the Plio-Pleistocene fossil record of crab predation on morphologically similar pairs of right- and left-handed snail species, we show here for the first time, contrary to traditional wisdom, that rare left-handed coiling promotes survival from attacks by right-handed crabs. This frequency-dependent result influences the balance of selection processes that maintain left-handedness at the species level and parallels some social interactions in human cultures, such as sports that involve dual contests between opponents of opposite handedness.

  20. Observations of the Crab Nebula with Early HAWC Data

    CERN Document Server

    Greus, Francisco Salesa

    2015-01-01

    The High Altitude Water Cherenkov (HAWC) Observatory is a TeV gamma-ray detector, completed in early 2015. HAWC started science operations in August 2013 with a third of the detector taking data. Several known gamma-ray sources have already been detected with the first HAWC data. Among these sources, the Crab Nebula, the brightest steady gamma-ray source at very high energies in our Galaxy, has been detected with high significance. In this contribution I will present the results of the observations of the Crab Nebula with HAWC, including time variability, and the detector performance based on early data.

  1. Species Diversity and Abundance of Marine Crabs (Portunidae: Decapoda) from a Collapsible Crab Trap Fishery at Kung Krabaen Bay, Chanthaburi Province, Thailand

    Science.gov (United States)

    Kunsook, Chutapa; Dumrongrojwatthana, Pongchai

    2017-01-01

    The diversity and abundance of marine crabs from a collapsible crab trap fishery at Kung Krabaen Bay, Gulf of Thailand, were observed from August 2012 to June 2013 using 10 sampling stations. The results showed that there were seven families, 11 genera and 17 species (two anomuran and 15 brachyuran crabs). The two anomuran species were Clibanarius virescens (1,710 individuals) and Clibanarius infraspinatus (558 individuals). For brachyuran crabs, Portunidae was the most common family, including 10 species. The dominant species of brachyuran crabs included Thalamita crenata (897 individuals), Portunus pelagicus (806 individuals), Charybdis affinis (344 individuals), Scylla sp. (201 individuals), and Charybdis anisodon (100 individuals). The abundance of crabs was affected by the habitat type. Anomuran crabs had the highest abundance in Halodule pinifolia seagrass beds, whilst brachyurans had the highest abundance in Enhalus acoroides seagrass beds. The dominant brachyuran species were found in pelagic areas near the bay mouth, such as P. pelagicus, P. sanguinolentus, C. feriatus, C. helleri, C. natator, C. affinis, and M. hardwickii. Lastly, reforested mangroves were important habitats for Scylla tranquebarica and C. anisodon. Seasonal and physical factors influenced the abundance of some crabs, for example, the abundance of C. virescens was correlated with temperature, and the abundance of T. crenata was correlated with transparency depth. Our results revealed that Kung Krabaen Bay serves as the home to many marine crab species; however, our results also revealed that 49% of the harvested crabs (2,308 out of 4,694 individuals) were simply discarded and subsequently died. Moreover, our research noted that eight non-target species will become target species in the near future. Therefore, research on the reproductive biology of some marine crabs and an improved understanding of the importance of marine crabs by local fishermen are necessary to prevent biodiversity

  2. The complete mitogenome of the hydrothermal vent crab Xenograpsus testudinatus (Decapoda, Brachyura) and comparison with brachyuran crabs.

    Science.gov (United States)

    Ki, Jang-Seu; Dahms, Hans-Uwe; Hwang, Jiang-Shiou; Lee, Jae-Seong

    2009-12-01

    In this study, we analyzed the complete mitochondrial (mt) genome of a hydrothermal vent crab Xenograpsus testudinatus (Decapoda: Brachyura) obtained from the hydrothermal vents off Kueishantao Island, Taiwan, which extend from the deep sea Okinawa Trench. The mitogenome of X. testudinatus was 15,796 bp in length and contained the same 37 genes (e.g. 2 rRNAs, 22 tRNAs, and 13 PCGs) found in other metazoan mitogenomes. Analysis of the structural mt gene order in X. testudinatus revealed that the 13 PCGs, excluding a translocation of ND6-Cyt b cluster, were similarly ordered when compared to the pancrustacean ground pattern; however the tRNAs were severely rearranged. Phylogenetic analysis of decapod mitogenomes showed that the molecular taxonomy of the vent crab was in accordance with its morphological systematics. Together, these findings suggest that the vent crab studied here has little mitochondrial genetic variation when compared with morphologically defined conspecifics from other marine habitats.

  3. Boxer crabs induce asexual reproduction of their associated sea anemones by splitting and intraspecific theft.

    Science.gov (United States)

    Schnytzer, Yisrael; Giman, Yaniv; Karplus, Ilan; Achituv, Yair

    2017-01-01

    Crabs of the genus Lybia have the remarkable habit of holding a sea anemone in each of their claws. This partnership appears to be obligate, at least on the part of the crab. The present study focuses on Lybia leptochelis from the Red Sea holding anemones of the genus Alicia (family Aliciidae). These anemones have not been found free living, only in association with L. leptochelis. In an attempt to understand how the crabs acquire them, we conducted a series of behavioral experiments and molecular analyses. Laboratory observations showed that the removal of one anemone from a crab induces a "splitting" behavior, whereby the crab tears the remaining anemone into two similar parts, resulting in a complete anemone in each claw after regeneration. Furthermore, when two crabs, one holding anemones and one lacking them, are confronted, the crabs fight, almost always leading to the "theft" of a complete anemone or anemone fragment by the crab without them. Following this, crabs "split" their lone anemone into two. Individuals of Alicia sp. removed from freshly collected L. leptochelis were used for DNA analysis. By employing AFLP (Fluorescence Amplified Fragments Length Polymorphism) it was shown that each pair of anemones from a given crab is genetically identical. Furthermore, there is genetic identity between most pairs of anemone held by different crabs, with the others showing slight genetic differences. This is a unique case in which one animal induces asexual reproduction of another, consequently also affecting its genetic diversity.

  4. Boxer crabs induce asexual reproduction of their associated sea anemones by splitting and intraspecific theft

    Directory of Open Access Journals (Sweden)

    Yisrael Schnytzer

    2017-01-01

    Full Text Available Crabs of the genus Lybia have the remarkable habit of holding a sea anemone in each of their claws. This partnership appears to be obligate, at least on the part of the crab. The present study focuses on Lybia leptochelis from the Red Sea holding anemones of the genus Alicia (family Aliciidae. These anemones have not been found free living, only in association with L. leptochelis. In an attempt to understand how the crabs acquire them, we conducted a series of behavioral experiments and molecular analyses. Laboratory observations showed that the removal of one anemone from a crab induces a “splitting” behavior, whereby the crab tears the remaining anemone into two similar parts, resulting in a complete anemone in each claw after regeneration. Furthermore, when two crabs, one holding anemones and one lacking them, are confronted, the crabs fight, almost always leading to the “theft” of a complete anemone or anemone fragment by the crab without them. Following this, crabs “split” their lone anemone into two. Individuals of Alicia sp. removed from freshly collected L. leptochelis were used for DNA analysis. By employing AFLP (Fluorescence Amplified Fragments Length Polymorphism it was shown that each pair of anemones from a given crab is genetically identical. Furthermore, there is genetic identity between most pairs of anemone held by different crabs, with the others showing slight genetic differences. This is a unique case in which one animal induces asexual reproduction of another, consequently also affecting its genetic diversity.

  5. The use of artificial crabs for testing predatory behavior and health in the octopus.

    Science.gov (United States)

    Amodio, Piero; Andrews, Paul; Salemme, Marinella; Ponte, Giovanna; Fiorito, Graziano

    2014-01-01

    The willingness of the cephalopod mollusc Octopus vulgaris to attack a live crab is traditionally used as a method to assess the overall health and welfare of octopuses in the laboratory. This method requires placing a crab in the home tank of an animal, measuring the time (latency) taken for the octopus to initiate an attack and withdrawing the crab immediately prior to capture. The same crab is commonly used to assess multiple octopuses as part of daily welfare assessment. Growing concern for the welfare of crustaceans and a review of all laboratory practices for the care and welfare of cephalopods following the inclusion of this taxon in 2010/63/EU prompted a study of the utility of an artificial crab to replace a live crab in the assessment of octopus health. On consecutive days O. vulgaris (N=21) were presented with a live, a dead or an artificial crab, and the latency to attack measured. Despite differences in the predatory performance towards the three different crab alternatives, octopuses readily attacked the artificial (and the dead) crab, showing that they can generalize and respond appropriately towards artificial prey. Researchers should consider using an artificial crab to replace the use of a live crab as part of the routine health assessment of O. vulgaris.

  6. Paragonimus ohirai metacercariae in crabs collected along the Arakawa River in Tokyo, Japan.

    Science.gov (United States)

    Sugiyama, Hiromu; Morishima, Yasuyuki; Kameoka, Yosuke; Arakawa, Kyoko; Kawanaka, Masanori

    2004-08-01

    Brackish water crabs infected with Paragonimus ohirai metacercariae have been reported in various regions in Japan. However, infected crabs have not been identified in Tokyo. We therefore collected the crab, Chiromantes dehaani, between August 2002 and July 2003 from 12 locations along the Arakawa River that flows through Tokyo. Of the 922 captured crabs, 177 (19%) from 6 locations were infected with Paragonimus metacercariae. The prevalence of metacercariae at these 6 locations ranged from 5 to 89%. The number of metacercariae per infected crab ranged from 1 to 190, with an average of 13.1. The morphological features of the metacercariae and of adult worms recovered from test rats infected with metacercariae showed that the metacercariae in the infected crabs were P. ohirai Miyazaki, 1939. The ITS2 sequence data support this conclusion. This paper is the first description of P. ohirai infection of crabs in Tokyo.

  7. Distribution of blue swimming crab (Portunus pelagicus Linnaeus, 1758 in Trang Province

    Directory of Open Access Journals (Sweden)

    Buncha Somboonsuke

    2010-07-01

    Full Text Available The fishery of the blue swimming crab (Portunus pelagicus Linnaeus, 1758 is very important to the economy of small-scale fishermen. Greater knowledge about its distribution could lead to more efficient management. This study was conducted in Trang Province from October 2006 to August 2007 using collapsible crab traps. To reveal the spatial distribution of the species, we analyzed the data using geostatistical methods. The standard interpolate procedure was applied to model the crab distribution. There were clear spatial distribution differences among the small crabs, large crabs and ovigerous females in study area. The mapping showed that small crabs (carapace width 10 cm were farther offshore. Ovigerous females peaked in abundance during 2 periods: March – April and August – September. This information can be used to support decision making concerning the designation of fishing zones and the optimization of the blue swimming crab fishery in the study area.

  8. 50 CFR Table 8 to Part 680 - Initial QS and PQS Pool for Each Crab QS Fishery

    Science.gov (United States)

    2010-10-01

    ... 50 Wildlife and Fisheries 9 2010-10-01 2010-10-01 false Initial QS and PQS Pool for Each Crab QS... Crab QS Fishery Crab QS Fishery Initial QS Pool Initial PQS Pool BBR Bristol Bay red king crab 400,000,000 400,000,000 BSS Bering Sea snow crab (C. opilio) 1,000,000,000 1,000,000,000 EAG Eastern...

  9. Investigation of plutonium concentration and distribution in burrowing crayfish from the White Oak Creek floodplain

    Energy Technology Data Exchange (ETDEWEB)

    Delaney, M.S.; Dahlman, R.C.; Craig, R.B.

    1979-01-01

    The White Oak floodplain was contaminated with several radionuclides, including /sup 239/Pu, during the Manhattan Project in 1944. Plutonium distribution in the soil is nonhomogeneous. An investigation was conducted to deterine Pu accumulation in a resident animal population. Crayfish were chosen because they complete their life-cycle within the contaminated environment, they directly contact contaminated muds, and they function in a food chain of significance to man. Two major conclusions of the research were that Pu concentrations in contaminated crayfish typically exceed those of control crayfish by two orders of magnitude and that if an incident were to occur in which a standard man ingested the soft tissues of ten crayfish from the floodplain, an insignificant whole body dose would accrue over the subsequent 50 years of life. The digestive tract of contaminated crayfish contained 21 to 33% of the Pu body burden, soft tissues contained 11 to 31% of the Pu body burden, and 48 to 62% of the Pu body burden of contaminated crayfish was associated with the carapace. Therefore, at a molt a large proportion of its accumulated Pu is deposited in the environment. A supplementary laboratory investigation using /sup 237/Pu included a chronic Pu uptake study by uncontaminated crayfish. In these crayfish, from 64 to 82% of the /sup 237/Pu was associated with the body tissues. Complementary data for /sup 237/Pu associated with the carapace ranged from 18 to 37% of the distribution. An inventory of /sup 239/Pu in crayfish at two sites on the floodplain was calculated by multiplying the estimated biomass of the crayfish by their average /sup 239/Pu concentration. This evaluation of Pu associated with the crayfish population was compared to an inventory of /sup 239/Pu in the soil in which they burrow and was found to be eight orders of magnitude less.

  10. Diurnal Changes in Angular Sensitivity of Crab Photoreceptors

    NARCIS (Netherlands)

    Leggett, L.M.W.; Stavenga, D.G.

    1981-01-01

    The electrophysiological and anatomical consequences of diurnal changes in screening pigment position were investigated in the apposition eye of the portunid crab Scylla serrata. Intracellular recordings revealed that the acceptance angles of dark-adapted photoreceptors enlarged up to four-fold at n

  11. X-ray Pulsar in the Crab Nebula.

    Science.gov (United States)

    Fritz, G; Henry, R C; Meekins, J F; Chubb, T A; Friedman, H

    1969-05-09

    X-ray pulsations have been observed in the Crab Nebula at a frequency closely matching the radio and optical pulsations. About 5 percent of the total x-ray power of the nebula appears in the pulsed component. The x-ray pulsations have the form of a main pulse and an interpulse separated by about 12 milliseconds.

  12. Hard X-ray Variations in the Crab Nebula

    Science.gov (United States)

    Wilson-Hodge, Colleen A.; Cherry, M. L.; Case, G. L.; Baumgartner, W. H.; Beklen, E.; Bhat, P. N.; Briggs, M. S.; Camero-Arranz, A.; Connaughton, V.; Finger, M. H.; Gehrels, N.; Greiner, J.; Jahoda, K.; Jenke, P.; Kippen, R. M.; Kouveliotou, C.; Krimm, H. A.; Kuulkers, E.; Meegan, C. A.; Natalucci, L.; Paciesas, W. S.; Preece, R.; Rodi, J. C.; Skinner, G. K.

    2013-01-01

    In the first two years of science operations of the Fermi Gamma-ray Burst Monitor (GBM), August 2008 to August 2010, approximately 7% (70 mcrab) decline was discovered in the overall Crab Nebula flux in the 15 - 50 keV band, measured with the Earth occultation technique. This decline was independently confirmed with four other instruments: the RXTE/PCA, Swift/BAT, INTEGRAL/IBIS, and INTEGRAL/SPI. The pulsed flux measured with RXTE/PCA from 1999-2010 was consistent with the pulsar spin-down, indicating that the observed changes were nebular. From 2001 to 2010, the Crab nebula flux measured with RXTE/ PCA was particularly variable, changing by up to approximately 3.5% per year in the 15-50 keV band. These variations were confirmed with INTEGRAL/SPI starting in 2003, Swift/BAT starting in 2005, and Fermi GBM starting in 2008. Before 2001 and since 2010, the Crab nebula flux has appeared more stable, varying by less than 2% per year. I will present updated light curves in multiple energy bands for the Crab Nebula, including recent data from Fermi GBM, Swift/BAT, INTEGRAL and MAXI, and a 16-year long light curve from RXTE/PCA.

  13. Six years of VERITAS observations of the Crab Nebula

    CERN Document Server

    ,

    2015-01-01

    The Crab Nebula is the brightest source in the very-high-energy (VHE) gamma-ray sky and one of the best studied non-thermal objects. The dominant VHE emission mechanism is believed to be inverse Compton scattering of low energy photons on relativistic electrons. While it is unclear how the electrons in the nebula are accelerated to energies of $10^{16}$ eV, it is general consensus that the ultimate source of energy is the Crab pulsar at the center of the nebula. Studying VHE gamma-ray emission provides valuable insight into the emission mechanisms and ultimately helps to understand the remaining mysteries of the Crab, for example, how the Poynting dominated energy flow is converted into a particle dominated flow of energy. We report on the results of six years of Crab observations with VERITAS comprising 115 hours of data taken between 2007 and 2013. VERITAS is an array of four 12-meter imaging air Cherenkov telescopes located in southern Arizona. We report on the energy spectrum, light curve, and a study of ...

  14. Crab Nebula Variations in Hard X-rays

    Science.gov (United States)

    Wilson-Hodge, Colleen A.

    2012-01-01

    The Crab Nebula was surprisingly variable from 2001-2010, with less variability before 2001 and since mid-2010. We presented evidence for spectral softening from RXTE, Swift/BAT, and Fermi GBM during the mid-2008-2010 flux decline. We see no clear connections between the hard X-ray variations and the GeV flares

  15. Hermit Crabs (Decapoda: Anomura: Paguridea) from the Seychelles

    NARCIS (Netherlands)

    McLaughlin, P.A.; Hogarth, P.J.

    1998-01-01

    A report on the hermit crabs collected during the Netherlands Seychelles Expedition, 1992-1993, supplemented by materials collected for Enterprise Oil and examined by one of the authors. Twentyseven species, representing three families, are recognized, including one new genus and four new species.

  16. Cuticular proteins from the horseshoe crab, Limulus polyphemus

    DEFF Research Database (Denmark)

    Ditzel, Nicholas; Andersen, Svend Olav; Højrup, Peter

    2003-01-01

    Proteins were purified from the carapace cuticle of a juvenile horseshoe crab, Limulus polyphemus, and several of them were characterized by amino acid sequence determination. The proteins are small (7-16 kDa) and their isoelectric points range from 6.5 to 9.2. They have high contents of tyrosine...

  17. RGS X-Ray Spectroscopy of the Crab Nebula

    Science.gov (United States)

    Kaastra, J. S.; de Vries, C. P.; Costantini, E.; den Herder, J. A.

    The Crab nebula and pulsar have been widely used as a calibration source for X-ray instruments. The in-flight effective area calibration of the Reflection Grating Spectrometers (RGS) of XMM-Newton depends upon the availability of reliable calibration sources. We analyse RGS observations of the Crab using different instrument configurations and spatial offsets, and make use of previous determinations of the continuum spectrum of the nebula plus pulsar. Due to the high spectral resolution of the RGS, we resolve the main absorption edges. We get an excellent fit to the Crab spectrum using this fixed continuum and the absorption spectrum determined by RGS. We get accurate column densities for the neutral atoms of H, N, O, Ne, Mg and Fe as well as a clear detection of Fe II and firm upper limits for other ions (O II, Mg II). We find solar-like abundances for N, O and Mg, and Fe (adding Fe I and Fe II), while Ne is overabundant. Finally, we show how we can determine the absolute flux of Crab with high accuracy by combining RGS and Chandra LETGS spectra of different sources.

  18. Substrate use and selection in sympatric intertidal hermit crab species

    Directory of Open Access Journals (Sweden)

    A. TURRA

    Full Text Available Coexisting hermit crabs may competitively interact for shells and microhabitats, mainly when shell availability is habitat-related. Three species of Clibanarius (C. antillensis, C. sclopetarius, and C. vittatus coexist in the intertidal region of Pernambuco Islet, Araçá Region, São Sebastião Channel, southeastern Brazil. This study evaluated crab preferences for four substrate types used by these species in nature (rocky shore, pebbles, sand, and mud in allopatric (single species and sympatric (three species treatments in simulations of high tide and low tide. The substrate preference of the three hermit crabs did not vary between low and high tide situations. At low tide the crabs either moved into holes in the highly complex rocky substrate or buried themselves in mud. Substrate selection may explain the patterns of substrate use in nature only for C. vittatus. Clibanarius antillensis and C. sclopetarius showed closer similarities in the pattern of substrate selection in the sympatric treatment with the substrate use in nature than in allopatric treatment, indicating a positive influence (dependence of the presence of one species on the presence of another. Use of sub-optimal substrates, mainly by C. antillensis, may be caused by other factors such as its low desiccation tolerances. If competition for space takes place among these species, it would be more intense between C. sclopetarius and C. vittatus given their higher overlap in substrate preference than between them and C. antillensis.

  19. Project "Flappy Crab": An Edu-Game for Music Learning

    Science.gov (United States)

    Cardoso Gomes, Cristina Maria; Guerreiro Figueiredo, Mauro Jorge; Bidarra, José; Cardoso Gomes, José Duarte

    2014-01-01

    This paper discusses some possibilities of gamification and remixing process for music education. Analyses also the concepts of gamification, mashup, remix and presents its possible usage in education--music teaching--through the development of the project/educational game "Flappy Crab". The article begins with a brief introduction to…

  20. Seasonal variability of antioxidant biomarkers in mud crabs (Scylla serrata).

    Science.gov (United States)

    Paital, Biswaranjan; Chainy, G B N

    2013-01-01

    Studies on oxidative stress (OS) in crustacea are widely used as ecotoxicological indices to assess the environment risk produced by the impact of several stressor and pollutants. In the present study, effects of seasonality on OS physiology markers such as antioxidant enzymes (superoxide dismutase, catalase, glutathione peroxidase and glutathione reductase), small antioxidant molecules (ascorbic acid and reduced glutathione), oxidative stress indices (lipid peroxidation, protein carbonylation and hydrogen peroxide) and total antioxidant capacity in hepatopancreas, gills and abdominal muscle of adult mud crab Scylla serrata, sampled from Chilika lagoon of India, were determined in winter, summer and rainy seasons. Results indicate that variations in enzymatic and non-enzymatic antioxidants with relation to season were not only tissue specific but also were gender specific. The levels of OS parameters were higher in hepatopancreas in comparison to gills and abdominal muscle of the crabs in all seasons. OS indices in tissues of the crabs were mainly higher in summer season when temperature and salinity of the lagoon were high with low oxygen content. Although OS was lower in winter season and moderate in rainy season in tissues of male crabs, it was higher in gills and hepatopancreas of females in rainy season. Correlation analyses between hydrological parameters of the lagoon (temperature, salinity and dissolved oxygen content) and OS physiology parameters in tissues of crabs suggest that abiotic factors influence the levels of antioxidant enzymes and, thereby the OS status in a tissue and sex specific manner. Collectively, the results of the present work suggest that further investigation is warranted before using OS parameters in S. serrata as biomarkers to monitor estuarine environment as these are influenced by gender, tissue and season.

  1. Spatial and temporal patterns of subtidal and intertidal crabs excursions

    Science.gov (United States)

    Silva, A. C. F.; Boaventura, D. M.; Thompson, R. C.; Hawkins, S. J.

    2014-01-01

    Highly mobile predators such as fish and crabs are known to migrate from the subtidal zone to forage in the intertidal zone at high-tide. The extent and variation of these habitat linking movements along the vertical shore gradient have not been examined before for several species simultaneously, hence not accounting for species interactions. Here, the foraging excursions of Carcinus maenas (L.), Necora puber (Linnaeus, 1767) and Cancer pagurus (Linnaeus, 1758) were assessed in a one-year mark-recapture study on two replicated rocky shores in southwest U.K. A comparison between the abundance of individuals present on the shore at high-tide with those present in refuges exposed at low-tide indicated considerable intertidal migration by all species, showing strong linkage between subtidal and intertidal habitats. Estimates of population size based on recapture of marked individuals indicated that an average of ~ 4000 individuals combined for the three crab species, can be present on the shore during one tidal cycle. There was also a high fidelity of individuals and species to particular shore levels. Underlying mechanisms for these spatial patterns such as prey availability and agonistic interactions are discussed. Survival rates were estimated using the Cormack-Jolly-Seber model from multi-recapture analysis and found to be considerably high with a minimum of 30% for all species. Growth rates were found to vary intraspecifically with size and between seasons. Understanding the temporal and spatial variations in predation pressure by crabs on rocky shores is dependent on knowing who, when and how many of these commercially important crab species depend on intertidal foraging. Previous studies have shown that the diet of these species is strongly based on intertidal prey including key species such as limpets; hence intertidal crab migration could be associated with considerable impacts on intertidal assemblages.

  2. Comparison of hydraulics and particle removal efficiencies in a mixed cell raceway and Burrows pond rearing system

    Science.gov (United States)

    Moffitt, Christine M.

    2016-01-01

    We compared the hydrodynamics of replicate experimental mixed cell and replicate standard Burrows pond rearing systems at the Dworshak National Fish Hatchery, ID, in an effort to identify methods for improved solids removal. We measured and compared the hydraulic residence time, particle removal efficiency, and measures of velocity using several tools. Computational fluid dynamics was used first to characterize hydraulics in the proposed retrofit that included removal of the traditional Burrows pond dividing wall and establishment of four counter rotating cells with appropriate drains and inlet water jets. Hydraulic residence time was subsequently established in the four full scale test tanks using measures of conductivity of a salt tracer introduced into the systems both with and without fish present. Vertical and horizontal velocities were also measured with acoustic Doppler velocimetry in transects across each of the rearing systems. Finally, we introduced ABS sinking beads that simulated fish solids then followed the kinetics of their removal via the drains to establish relative purge rates. The mixed cell raceway provided higher mean velocities and a more uniform velocity distribution than did the Burrows pond. Vectors revealed well-defined, counter-rotating cells in the mixed cell raceway, and were likely contributing factors in achieving a relatively high particle removal efficiency-88.6% versus 8.0% during the test period. We speculate retrofits of rearing ponds to mixed cell systems will improve both the rearing environments for the fish and solids removal, improving the efficiency and bio-security of fish culture. We recommend further testing in hatchery production trials to evaluate fish physiology and growth.

  3. Estimated Entrainment of Dungeness Crab During Dredging For The Columbia River Channel Improvement Project

    Energy Technology Data Exchange (ETDEWEB)

    Pearson, Walter H.; Williams, Greg D.; Skalski, John R.

    2002-12-01

    The studies reported here focus on issues regarding the entrainment of Dungeness crab related to the proposed Columbia River Channel Improvement Project and provided direct measurements of crab entrainment rates at three locations (Desdomona Shoals, Upper Sands, and Miller Sands) from RM4 to RM24 during summer 2002. Entrainment rates for all age classes of crabs ranged from zero at Miller Sands to 0.224 crabs per cy at Desdemona Shoals in June 2002. The overall entrainment rate at Desdomona Shoals in September was 0.120 crabs per cy. A modified Dredge Impact Model (DIM) used the summer 2002 entrainment rates to project crab entrainment and adult equivalent loss and loss to the fishery for the Channel Improvement Project. To improve the projections, entrainment data from Flavel Bar is needed. The literature, analyses of salinity intrusion scenarios, and the summer 2002 site-specific data on entrainment and salinity all indicate that bottom salinity influences crab distribution and entrainment, especially at lower salinities. It is now clear from field measurements of entrainment rates and salinity during a period of low river flow (90-150 Kcfs) and high salinity intrusion that entrainment rates are zero where bottom salinity is less than 16 o/oo most of the time. Further, entrainment rates of 2+ and older crab fall with decreasing salinity in a clear and consistent manner. More elaboration of the crab distribution- salinity model, especially concerning salinity and the movements of 1+ crab, is needed.

  4. Black yeast-like fungi associated with Lethargic Crab Disease (LCD) in the mangrove-land crab, Ucides cordatus (Ocypodidae).

    Science.gov (United States)

    Vicente, Vania A; Orélis-Ribeiro, R; Najafzadeh, M J; Sun, Jiufeng; Guerra, Raquel Schier; Miesch, Stephanie; Ostrensky, Antonio; Meis, Jacques F; Klaassen, Corné H; de Hoog, G S; Boeger, Walter A

    2012-07-06

    Lethargic Crab Disease (LCD) caused extensive epizootic mortality of the mangrove land crab Ucides cordatus (Brachyura: Ocypodidae) along the Brazilian coast, mainly in the Northeastern region. The disease was named after the symptoms of slow movement of infected crabs. Causative agents were suspected to be two black yeast-like fungi of the family Herpotrichiellaceae (ascomycete order Chaetothyriales), judged by infected tissue biopsies from moribund U. cordatus. The aim of the present study is to prove that two species are involved in the disease: the recently described black yeast Exophiala cancerae, but also a less virulent, hitherto undescribed fonsecaea-like species, introduced here as the novel species Fonsecaea brasiliensis. Strains were identified by ITS rDNA sequencing, and species borderlines were established by multilocus sequencing and AFLP analysis. Fonsecaea brasiliensis proved to be closely related to the pathogenic species Cladophialophora devriesii which originally was isolated from a systemic infection in a human patient. The virulence of F. brasiliensis is lower than that of E. cancerae, as established by artificial inoculation of mangrove crabs.

  5. Comparative brain architecture of the European shore crab Carcinus maenas (Brachyura) and the common hermit crab Pagurus bernhardus (Anomura) with notes on other marine hermit crabs.

    Science.gov (United States)

    Krieger, Jakob; Sombke, Andy; Seefluth, Florian; Kenning, Matthes; Hansson, Bill S; Harzsch, Steffen

    2012-04-01

    The European shore crab Carcinus maenas and the common hermit crab Pagurus bernhardus are members of the sister taxa Brachyura and Anomura (together forming the taxon Meiura) respectively. Both species share similar coastal marine habitats and thus are confronted with similar environmental conditions. This study sets out to explore variations of general brain architecture of species that live in seemingly similar habitats but belong to different major malacostracan taxa and to understand possible differences of sensory systems and related brain compartments. We examined the brains of Carcinus maenas, Pagurus bernhardus, and three other hermit crab species with immunohistochemistry against tyrosinated tubulin, f-actin, synaptic proteins, RF-amides and allatostatin. Our comparison showed that their optic neuropils within the eyestalks display strong resemblance in gross morphology as well as in detailed organization, suggesting a rather similar potential of processing visual input. Besides the well-developed visual system, the olfactory neuropils are distinct components in the brain of both C. maenas and P. bernhardus as well as the other hermit crabs, suggesting that close integration of olfactory and visual information may be useful in turbid marine environments with low visibility, as is typical for many habitats such as, e.g., the Baltic and the North Sea. Comparing the shape of the olfactory glomeruli in the anomurans showed some variations, ranging from a wedge shape to an elongate morphology. Furthermore, the tritocerebrum and the organization of the second antennae associated with the tritocerebrum seem to differ markedly in C. maenas and P. bernhardus, indicating better mechanosensory abilities in the latter close to those of other Decapoda with long second antennae, such as Astacida, Homarida, or Achelata. This aspect may also represent an adaptation to the "hermit lifestyle" in which competition for shells is a major aspect of their life history. The shore

  6. CRABS IN CRISIS:BIOGEOGRAPHIC DISTRIBUTIONS, ABUNDANCES, AND VULNERABILITIES TO CLIMATE CHANGE OF BRACHYURAN AND LITHODID CRABS FROM THE GULF OF CALIFORNIA TO THE BEAUFORT SEA

    Science.gov (United States)

    To predict the relative vulnerability of near-coastal species to climate change we analyzed the biogeographic and abundance patterns of the brachyuran or ‘True’ crabs (n=368) and lithodid or ‘King’ crabs (n=20) that are found in the twelve MEOW (“Mar...

  7. Examining effects of sea level rise and marsh crabs on Spartina patens using mesocosms

    Science.gov (United States)

    Coastal salt marshes provide essential ecosystem services but face increasing threats from habitat loss, eutrophication, changing precipitation patterns, and accelerating rates of sea level rise (SLR). Recent studies have suggested that herbivory and burrowing by native salt mars...

  8. Heavy metals in red crabs, Chaceon quinquedens, from the Gulf of Mexico.

    Science.gov (United States)

    Perry, Harriet; Isphording, Wayne; Trigg, Christine; Riedel, Ralf

    2015-12-30

    The red crab, Chaceon quinquedens, is distributed in deep waters of the Gulf of Mexico (GOM) and is most abundant in an area associated with sediment deposition from the Mississippi River. Sediment geochemistry and biological and ecological traits of red crabs favor accumulation of contaminants. Red crabs, sediment, and bottom water samples were taken from three distinct geographic locations representing areas with differing exposure to contaminant laden effluents from the Mississippi River. Inductively coupled plasma spectrophotometry and atomic absorption spectrophotometry were employed to determine levels of heavy metals in red crab muscle tissue. Ion site partitioning was used to determine metal speciation in sediments. Red crabs showed evidence of heavy metal bioaccumulation in all sample areas with high variability in contaminant levels in individual crabs for some metals. Bioavailability of metals in sediment did not always result in accumulation in muscle tissue.

  9. Crab dispersion and its impact on the CERN Large Hadron Collider collimation

    CERN Document Server

    Sun, P; Tomàs, R; Zimmermann, F

    2010-01-01

    Crab cavities are proposed to be used for a luminosity upgrade of the Large Hadron Collider (LHC). Crab cavities are rf cavities operated in a transverse dipole mode, which imparts on the beam particles a transverse kick that varies with the longitudinal position along the bunch. The crab cavity introduces another kind of dispersion to the particles which is z dependent, and thus could complicate the beam dynamics and have an impact on the LHC collimation system. As for LHC, the off-momentum beta-beat and dispersion-beat already compromise the performance of the collimation system; the crab dispersion introduced by global crab cavities might do the same, and should be carefully evaluated. In this paper, we present a definition for the crab dispersion, and study its impact on the LHC collimation system.

  10. The assessment of general well-being using spontaneous burrowing behaviour in a short-term model of chemotherapy-induced mucositis in the rat.

    Science.gov (United States)

    Whittaker, A L; Lymn, K A; Nicholson, A; Howarth, G S

    2015-01-01

    Mucositis is a common and serious side-effect experienced by cancer patients during treatment with chemotherapeutic agents. Consequently, programmes of research focus on the elucidation of novel therapeutics for alleviation of mucositis symptoms, and these frequently use animal models. However, although these models are assumed to be painful and distressing to the animal, endpoints are difficult to determine. The aim of this study was to evaluate whether a change in burrowing behaviour could provide an indication of disease onset and potentially be applied as a humane endpoint. Baseline burrowing behaviour was measured in healthy animals on three occasions by determining the weight of gravel displaced from a hollow tube. Mucositis was then induced in the same animals by intraperitoneal injection of 5-fluorouracil (150 mg/kg) and burrowing behaviour recorded over three consecutive days. Standard measures of disease progression, including body weight loss and clinical score, were also made. The presence of mucositis was confirmed at necropsy by findings of decreased duodenal and colon lengths, and reduced liver, spleen and thymus weights in comparison with non-treated control animals. Histological score of the jejunum and ileum was also significantly increased. Mucositis onset coincided with a decrease in mean burrowing behaviour which was progressive, however this result did not achieve statistical significance (P = 0.66).We conclude that burrowing may be a useful indicator of inflammation in the mucositis model, although this requires further characterization. Pre-selection of animals into treatment groups based on their prior burrowing performance should be pursued in further studies.

  11. Comparison of burrowing and stimuli-evoked pain behaviors as end-points in rat models of inflammatory pain and peripheral neuropathic pain

    Directory of Open Access Journals (Sweden)

    Arjun eMuralidharan

    2016-05-01

    Full Text Available Establishment and validation of ethologically-relevant, non-evoked behavioral end-points as surrogate measures of spontaneous pain in rodent pain models has been proposed as a means to improve preclinical to clinical research translation in the pain field. Here, we compared the utility of burrowing behavior with hypersensitivity to applied mechanical stimuli for pain assessment in rat models of chronic inflammatory and peripheral neuropathic pain. Briefly, groups of male Sprague-Dawley rats were habituated to the burrowing environment and trained over a 5-day period. Rats that burrowed ≤450g of gravel on any two days of the individual training phase were excluded from the study. The remaining rats received either a unilateral intraplantar injection of Freund’s complete adjuvant (FCA or saline, or underwent unilateral chronic constriction injury (CCI of the sciatic nerve- or sham-surgery. Baseline burrowing behavior and evoked pain behaviors were assessed prior to model induction, and twice-weekly until study completion on day 14. For FCA- and CCI-rats, but not the corresponding groups of sham-rats, evoked mechanical hypersensitivity developed in a temporal manner in the ipsilateral hindpaws. Although burrowing behavior also decreased in a temporal manner for both FCA- and CCI-rats, there was considerable inter-animal variability. By contrast, mechanical hyperalgesia and mechanical allodynia in the ipsilateral hindpaws of FCA- and CCI-rats respectively, exhibited minimal inter-animal variability. Our data collectively show that burrowing behavior is altered in rodent models of chronic inflammatory pain and peripheral neuropathic pain. However, large group sizes are needed to ensure studies are adequately powered due to considerable inter-animal variability.

  12. Species Profiles. Life Histories and Environmental Requirements of Coastal Fishes and Invertebrates (Pacific Northwest). Dungeness Crab.

    Science.gov (United States)

    1986-08-01

    analyze some of the original data in this report was provided by Washington Sea Grant. vii 1 II ’ 1 . w. Figure 1. Dungeness crab. - DUNGENESS CRAB 5 -4...Lough Poole 1967). After 1 year of growth 1976; Armstrong 1983). beyond hatching, most crabs in Bodega Bay, California, are in their 8th, Juveniles...Stevens 1982). Grays Harbor, while the range is 63-94 After molting, the juveniles are mm in Bodega Bay (Poole 1967; Stevens found in shallow coastal

  13. A summary of the dungeness crab research program 1974-1980

    OpenAIRE

    1981-01-01

    In 1974, the State Legislature directed the Department of Fish and Game to investigate causes of a long-term decline in Dungeness crab, Cancer magister, commercial fishery landings in central California. The Department established the Dungeness Crab Research Program which conducted investigations on crab life history, pollution, and oceanography. Genetic studies to determine if there were different stocks along the coast were inconclusive. Early larval stages (zoeae) drifted progressive...

  14. Observations of a Series of Six Recent Glitches in the Crab Pulsar

    CERN Document Server

    Wong, T; Lyne, A G; Wong, Tony

    2001-01-01

    From 1995 to 1999, daily monitoring of the radio emission from the Crab pulsar at the Green Bank and Jodrell Bank observatories revealed a series of six sudden rotational spinups or glitches, doubling the number of glitches observed for this pulsar since 1969. With these observations, the range of time intervals between significant Crab glitches has widened considerably, indicating that the occurrence of Crab glitches may be more random than previously thought. The new glitch amplitudes ($\\Delta\

  15. Edible crabs "go west": migrations and incubation cycle of Cancer pagurus revealed by electronic tags.

    Directory of Open Access Journals (Sweden)

    Ewan Hunter

    Full Text Available Crustaceans are key components of marine ecosystems which, like other exploited marine taxa, show seasonable patterns of distribution and activity, with consequences for their availability to capture by targeted fisheries. Despite concerns over the sustainability of crab fisheries worldwide, difficulties in observing crabs' behaviour over their annual cycles, and the timings and durations of reproduction, remain poorly understood. From the release of 128 mature female edible crabs tagged with electronic data storage tags (DSTs, we demonstrate predominantly westward migration in the English Channel. Eastern Channel crabs migrated further than western Channel crabs, while crabs released outside the Channel showed little or no migration. Individual migrations were punctuated by a 7-month hiatus, when crabs remained stationary, coincident with the main period of crab spawning and egg incubation. Incubation commenced earlier in the west, from late October onwards, and brooding locations, determined using tidal geolocation, occurred throughout the species range. With an overall return rate of 34%, our results demonstrate that previous reluctance to tag crabs with relatively high-cost DSTs for fear of loss following moulting is unfounded, and that DSTs can generate precise information with regards life-history metrics that would be unachievable using other conventional means.

  16. Diet selectivity of juvenile blue crabs (Callinectes sapidus) in Chesapeake Bay.

    Science.gov (United States)

    Seitz, Rochelle D; Knick, Kathleen E; Westphal, Miranda

    2011-10-01

    Shallow coves in Chesapeake Bay have abundant food and serve as nursery grounds for juvenile blue crabs. In this study, we examined the relationships between the diet of very small (4-40 mm CW) juvenile blue crabs and the benthic infauna in shallow, unvegetated nursery coves. We compared infauna in benthic samples with gut contents of juvenile blue crabs from six shallow coves in each of two sub-estuaries (Rappahannock and York Rivers) in Chesapeake Bay, Virginia, USA. Benthic communities differed depending on river and location, with abundant clams in upriver regions and abundant polychaetes in downriver regions. Juvenile crabs, like adults, appeared to be opportunistic feeders, with gut contents including clams, amphipods, polychaetes, small crustaceans, plant matter, and detritus. There was a positive relationship between polychaetes in the benthic samples and in crab guts, suggesting that juvenile crabs are opportunistic feeders on polychaetes in the benthos. Moreover, Ivlev's electivity index and foraging ratio showed that clams and polychaetes were selectively eaten at all locations. Alternatively, crabs selectively rejected amphipods. Crab densities corresponded positively with polychaete densities, which suggests that there may be bottom-up control of crab distributions and that food resources are important in nursery habitats.

  17. Purification and characterization of a cadmium-induced metallothionein from the shore crab Carcinus maenas (L.)

    DEFF Research Database (Denmark)

    Pedersen, K L; Pedersen, S N; Højrup, P;

    1994-01-01

    Two metallothionein variants were purified from the midgut gland of crabs (Carcinus maenas) exposed to a high cadmium concentration (2 p.p.m.). One of the variants was purified from crabs exposed to a low cadmium concentration (0.5 p.p.m.). The purification method involved acetone precipitation...... from crabs exposed to the high cadmium concentration differed only by a single residue of methionine at the N-terminus. The single variant isolated from crabs exposed to the low cadmium concentration was the one without the N-terminal methionine, indicating that high cadmium concentrations either...

  18. The effect of salinity on experimental infections of a Hematodinium sp. in blue crabs, Callinectes sapidus.

    Science.gov (United States)

    Coffey, Anna H; Li, Caiwen; Shields, Jeffrey D

    2012-06-01

    The parasitic dinoflagellate Hematodinium sp. parasitizes blue crabs along the Atlantic seaboard of the United States. Infections in blue crabs have only been reported from waters where salinity is >11 practical salinity units (psu). Blue crabs maintain a hyperosmotic internal concentration at low salinities (0-5 psu), roughly comparable to 24 psu, and should be capable of maintaining an infection in low-salinity waters even if Hematodinium spp. cells are intolerant of low salinities. We tested this notion by observing the effect of low salinity on the progression of disease in crabs experimentally infected with the parasite. Blue crabs were acclimated to 5 psu or 30 psu salinity treatments. They were inoculated with Hematodinium sp. and necropsied 3, 7, 10, and 15 days post-inoculation. The low-salinity treatment did not have an effect on the proliferation of Hematodinium sp. infections in blue crabs; moreover, a greater proportion of infections in crabs in the low-salinity treatment developed dinospore stages than did those in the high-salinity treatment, indicating that salinity may affect the development of the parasite. However, dinospores from in vitro cultures rapidly became inactive when held in salinities crabs at low salinities, but that the parasite is incapable of transmission in this environment, which explains the lack of natural infections in crabs at low salinities.

  19. Edible crabs "go west": migrations and incubation cycle of Cancer pagurus revealed by electronic tags.

    Science.gov (United States)

    Hunter, Ewan; Eaton, Derek; Stewart, Christie; Lawler, Andrew; Smith, Michael T

    2013-01-01

    Crustaceans are key components of marine ecosystems which, like other exploited marine taxa, show seasonable patterns of distribution and activity, with consequences for their availability to capture by targeted fisheries. Despite concerns over the sustainability of crab fisheries worldwide, difficulties in observing crabs' behaviour over their annual cycles, and the timings and durations of reproduction, remain poorly understood. From the release of 128 mature female edible crabs tagged with electronic data storage tags (DSTs), we demonstrate predominantly westward migration in the English Channel. Eastern Channel crabs migrated further than western Channel crabs, while crabs released outside the Channel showed little or no migration. Individual migrations were punctuated by a 7-month hiatus, when crabs remained stationary, coincident with the main period of crab spawning and egg incubation. Incubation commenced earlier in the west, from late October onwards, and brooding locations, determined using tidal geolocation, occurred throughout the species range. With an overall return rate of 34%, our results demonstrate that previous reluctance to tag crabs with relatively high-cost DSTs for fear of loss following moulting is unfounded, and that DSTs can generate precise information with regards life-history metrics that would be unachievable using other conventional means.

  20. Oxygen and carbon dioxide sensitivity of ventilation in amphibious crabs, Cardisoma guanhumi, breathing air and water.

    Science.gov (United States)

    Gannon, Andrew T; Henry, Raymond P

    2004-05-01

    Amphibious crabs, Cardisoma guanhumi, were acclimated to breathing either air or water and exposed to altered levels of oxygen and/or carbon dioxide in the medium. Hypercapnia (22, 36 and 73 torr CO(2)) stimulated a significant hypercapnic ventilatory response (HCVR) in both groups of crabs, with a much greater effect on scaphognathite frequency (Deltaf(SC)=+700%) in air-breathing crabs than water-breathing crabs (Deltaf(SC)=+100%). In contrast, hyperoxia induced significant hypoventilation in both sets of crabs. However, simultaneous hyperoxia and hypercapnia triggered a greater than 10-fold increase in f(SC) in air-breathing crabs but no change in water-breathing crabs. For water-breathing crabs hypoxia simultaneous with hypercapnia triggered the same response as hypoxia alone-bradycardia (-50%), and a significant increase in f(SC) at moderate exposures but not at the more extreme levels. The response of air-breathing crabs to hypoxia concurrent with hypercapnia was proportionally closer to the response to hypercapnia alone than to hypoxia. Thus, C. guanhumi were more sensitive to ambient CO(2) than O(2) when breathing air, characteristic of fully terrestrial species, and more sensitive to ambient O(2) when breathing water, characteristic of fully aquatic species. C. guanhumi possesses both an O(2)- and a CO(2)-based ventilatory drive whether breathing air or water, but the relative importance switches when the respiratory medium is altered.

  1. Multipole Field Effects for the Superconducting Parallel-Bar Deflecting/Crabbing Cavities

    Energy Technology Data Exchange (ETDEWEB)

    De Silva, Payagalage Subashini Uddika [Thomas Jefferson National Accelerator Facility, Newport News, VA (United States) and Old Dominion University, Norfolk, VA (United States); Delayen, Jean Roger [Old Dominion University, Norfolk, VA (United States)

    2012-09-01

    The superconducting parallel-bar deflecting/crabbing cavity is currently being considered as one of the design options in rf separation for the Jefferson Lab 12 GeV upgrade and for the crabbing cavity for the proposed LHC luminosity upgrade. Knowledge of multipole field effects is important for accurate beam dynamics study of rf structures. The multipole components can be accurately determined numerically using the electromagnetic surface field data in the rf structure. This paper discusses the detailed analysis of those components for the fundamental deflecting/crabbing mode and higher order modes in the parallel-bar deflecting/crabbing cavity.

  2. Spatial patterns and movements of red king and Tanner crabs: Implications for the design of marine protected areas

    Science.gov (United States)

    Taggart, S.J.; Mondragon, J.; Andrews, A.G.; Nielsen, J.K.

    2008-01-01

    Most examples of positive population responses to marine protected areas (MPAs) have been documented for tropical reef species with very small home ranges; the utility of MPAs for commercially harvested temperate species that have large movement patterns remains poorly tested. We measured the distribution and abundance of red king Paralithodes camtschaticus and Tanner Chionoecetes bairdi crabs inside and outside of MPAs in Glacier Bay National Park, Alaska, USA. By tagging a sub-sample of crabs with sonic tags, we estimated the movement of adult crabs from one of the MPAs (Muir Inlet) into the central portion of Glacier Bay where fishing still occurs. Tanner crabs and red king crabs moved similar average distances per day, although Tanner crabs had a higher transfer out of the Muir Inlet MPA into the central bay. Tanner crab movements were characterized by large variation among individual crabs, both in distance and direction traveled, while red king crabs migrated seasonally between 2 specific areas. Although Tanner crabs exhibited relatively large movements, distribution and abundance data suggest that they may be restricted at large spatial scales by habitat barriers. MPAs that are effective at protecting king and especially Tanner crab brood stock from fishing mortality will likely need to be larger than is typical of MPAs worldwide. However, by incorporating information on the seasonal movements of red king crabs and the location of habitat barriers for Tanner crabs, MPAs could likely be designed that would effectively protect adults from fishing mortality. ?? Inter-Research 2008.

  3. SUPERCONDUCTING RF-DIPOLE DEFLECTING AND CRABBING CAVITIES

    Energy Technology Data Exchange (ETDEWEB)

    Delayen, Jean [ODU, JLAB; De Silva, Paygalage Subashini [ODU, JLAB

    2013-09-01

    Recent interests in designing compact deflecting and crabbing structures for future accelerators and colliders have initiated the development of novel rf structures. The superconducting rf-dipole cavity is one of the first compact designs with attractive properties such as higher gradients, higher shunt impedance, the absence of lower order modes and widely separated higher order modes. Two rf-dipole designs of 400 MHz and 499 MHz have been designed, fabricated and tested as proof-of-principle designs of compact deflecting and crabbing cavities for the LHC high luminosity upgrade and Jefferson Lab 12 GeV upgrade. The first rf tests have been performed on the rf-dipole geometries at 4.2 K and 2.0 K in a vertical test assembly with excellent results. The cavities have achieved high gradients with high intrinsic quality factors, and multipacting levels were easily processed.

  4. Observations of "wisps" in magnetohydrodynamic simulations of the Crab Nebula

    CERN Document Server

    Camus, N F; Buccantini, N; Hughes, P A

    2009-01-01

    In this letter, we describe results of new high-resolution axisymmetric relativistic MHD simulations of Pulsar Wind Nebulae. The simulations reveal strong breakdown of the equatorial symmetry and highly variable structure of the pulsar wind termination shock. The synthetic synchrotron maps, constructed using a new more accurate approach, show striking similarity with the well known images of the Crab Nebula obtained by Chandra, and the Hubble Space Telescope. In addition to the \\textit{jet-torus} structure, these maps reproduce the Crab's famous moving wisps whose speed and rateof production agree with the observations. The variability is then analyzed using various statistical methods, including the method of structure function and wavelet transform. The results point towards the quasi-periodic behaviour with the periods of 1.5-3yr and MHD turbulence on scales below 1yr. The full account of this study will be presented in a follow up paper.

  5. The Crab Pulsar at Centimeter Wavelengths: I. Ensemble Characteristics

    CERN Document Server

    Hankins, T H; Eilek, J A

    2015-01-01

    We have observed the pulsar in the Crab Nebula at high radio frequencies and high time resolution. We present continuously sampled data at 640-ns time resolution, and individual bright pulses recorded at down to 0.25-ns time resolution. Combining our new data with previous data from our group and from the literature shows the dramatic changes in the pulsar's radio emission between low and high radio frequencies. Below about 5 GHz the mean profile is dominated by the bright Main Pulse and Low-Frequency Interpulse. Everything changes, however, above about 5 GHz; the Main Pulse disappears, the mean profile of the Crab pulsar is dominated by the High-Frequency Interpulse (which is quite different from its low-frequency counterpart) and the two High-Frequency Components. We present detailed observational characteristics of these different components which future models of the pulsar's magnetosphere must explain.

  6. THE CRAB PULSAR AT CENTIMETER WAVELENGTHS. I. ENSEMBLE CHARACTERISTICS

    Energy Technology Data Exchange (ETDEWEB)

    Hankins, T. H.; Eilek, J. A., E-mail: thankins@aoc.nrao.edu [Physics Department, New Mexico Tech, Socorro, NM 87801 (United States); Jones, G.

    2015-04-01

    We have observed the pulsar in the Crab Nebula at high radio frequencies and high time resolution. We present continuously sampled data at 640 ns time resolution and individual bright pulses recorded at down to 0.25 ns time resolution. Combining our new data with previous data from our group and from the literature shows the dramatic changes in the pulsar’s radio emission between low and high radio frequencies. Below about 5 GHz the mean profile is dominated by the bright Main Pulse and Low-Frequency Interpulse. Everything changes, however, above about 5 GHz; the Main Pulse disappears and the mean profile of the Crab pulsar is dominated by the High-Frequency Interpulse (which is quite different from its low-frequency counterpart) and the two High-Frequency Components. We present detailed observational characteristics of these different components which future models of the pulsar’s magnetosphere must explain.

  7. Geometric design of crab-like climbing and walking robots

    Institute of Scientific and Technical Information of China (English)

    LIU An-min; David Howard

    2004-01-01

    This paper considers the geometric design of crab-like walkers and climbers, without decoupling leg design from overall machine design. Crab-like machines represent an important sub-class of multi-legged robots, being particularly well suited to crossing difficult terrains. Firstly, the kinematic configurations and constraints are described, which determine the machine's kinematic characteristics. The influence of the design parameters on the kinematic workspace is discussed. Finally,a two stage design methodology is presented, comprising kinematic design and design optimisation, the latter being based on the use of design maps rather than numerical optimisation. The performance measures considered during design optimisation include kinematic, static and quasi-static measures.

  8. Observations of the Crab pulsar with the MAGIC telescopes

    CERN Document Server

    Saito, T Y; Giavitto, G; Klepser, S; Schweizer, T; Zanin, R

    2011-01-01

    We report on the observations of the Crab pulsar with the MAGIC telesopes. Data were taken both in the mono-mode ($>25$ GeV) and in the stereo-mode ($>50$ GeV). Clear signals from the two peaks were detected with both modes and the phase resolved energy spectra were calculated. By comparing with the measurements done by Fermi-LAT, we found that the energy spectra of the Crab pulsar does not follow a power law with an exponential cutoff, but that it extends as a power law after the break at around 5 GeV. This suggests that the emission above 25 GeV is not dominated by the curvatura radiation, which is inconsistent with the standard prediction of the OG and SG models.

  9. Antimicrobial lipids from the hemolymph of brachyuran crabs

    Digital Repository Service at National Institute of Oceanography (India)

    Ravichandran, S.; Wahidullah, S.; DeSouza, L.; Rameshkumar, G.

    and gram negative pathogenic bacterial strains. Similar result was observed with the hemolymph of some brachyuran crabs against clinical pathogens [18,19], Thalamita crenata [20] and Charybdis lucifera [21]. In the present study, maximum zone... properties, which are enhanced when these acids are esterified with glycerol [29]. It has generally been observed that Gram–positive bacteria are more susceptible to the antibacterial effect of lipids than Gram-negative bacteria. However, there are many...

  10. Crab Hole Mosquito Blues—The Story

    Centers for Disease Control (CDC) Podcasts

    2011-05-12

    This podcast reports on a humorous song that takes a look at a very serious human and equine disease. Written and performed by the MARU Health Angels Band, Bill Dietz, director of the Division of Nutrition, Physical Activity and Obesity at CDC, talks about the song, "Crab Hole Mosquito Blues", and the history behind it.  Created: 5/12/2011 by National Center for Emerging Zoonotic and Infectious Diseases (NCEZID).   Date Released: 5/23/2011.

  11. Motor Driving Leg Design for Bionic Crab-like Robot

    Institute of Scientific and Technical Information of China (English)

    Wang; Li-quan; Qu; Xiao-rong; Meng; Qing-xin; Yuan; Peng; Wang; Mo-nan

    2002-01-01

    The paper presents the design of walking leg for bionics crab-like robot, which is driven with micro servo motor. The kinematic characteristics of the bionics machine are analysed for optimized structure parameters, which has been used in the robot design. A three closed loop motor control system structure for joint driver is also given, as well as the multi-joint driving system for walking robot leg.

  12. Cymonomid crabs of the MAINBAZA Expedition (Decapoda: Brachyura).

    Science.gov (United States)

    Ahyong, Shane T

    2014-06-23

    Cymonomid crabs collected from the Mozambique Channel off Madagascar by the 2011 MAINBAZA Expedition are reported. Two species of Cymonomus A. Milne Edwards, 1880, are represented, of which one is new to science and the other, C. valdiviae Lankester, 1903, is rediscovered, being previously known only from the holotype. Three species of Cymonomidae are now known from the western Indian Ocean, including C. trifurcus Stebbing, 1920, from South Africa. 

  13. Chandra Phase-Resolved Spectroscopy of the Crab Pulsar

    CERN Document Server

    Weisskopf, M C; Paerels, F; Becker, W; Tennant, A F; Swartz, D A; Weisskopf, Martin C.; Dell, Stephen L. O'; Paerels, Frits; Becker, Werner; Tennant, Allyn F.; Swartz, Douglas A.

    2004-01-01

    We present the first phase-resolved study of the X-ray spectral properties of the Crab Pulsar that covers all pulse phases. The superb angular resolution of the Chandra X-ray Observatory enables distinguishing the pulsar from the surrounding nebulosity, even at pulse minimum. Analysis of the pulse-averaged spectrum measures interstellar X-ray extinction due primarily to photoelectric absorption and secondarily to scattering by dust grains in the direction of the Crab Nebula. We confirm previous findings that the line-of-sight to the Crab is underabundant in oxygen, although more-so than recently measured. Using the abundances and cross sections from Wilms, Allen & McCray (2000) we find [O/H] = (3.33 +/-0.25) x 10**-4. Analysis of the spectrum as a function of pulse phase measures the low-energy X-ray spectral index even at pulse minimum -- albeit with large statistical uncertainty -- and we find marginal evidence for variations of the spectral index. The data are also used to set a new (3-sigma) upper lim...

  14. Observation and Spectral Measurements of the Crab Nebula with Milagro

    Science.gov (United States)

    Abdo, A. A.; Allen, B. T.; Aune, T.; Benbow, W.; Berley, D.; Chen, C.; Christopher, G. E.; DeYoung, T.; Dingus, B. L.; Falcone, A.; Fleysher, L.; Fleysher, R.; Gonzalez, M. M.; Goodman, J. A.; Gordo, J. B.; Hays, E.; Hoffman, C. M.; Huntemeyer, P. H.; Kolterman, B. E.; Linnemann, J. T.; McEnery, J. E.; Morgan, T.; Mincer, A. I.; Nemethy, P.

    2011-01-01

    The Crab Nebula was detected with the Milagro experiment at a statistical significance of 17 standard deviations over the lifetime of the experiment. The experiment was sensitive to approximately 100 GeV - 100 TeV gamma ray air showers by observing the particle footprint reaching the ground. The fraction of detectors recording signals from photons at the ground is a suitable proxy for the energy of the primary particle and has been used to measure the photon energy spectrum of the Crab Nebula between 1 and 100 TeV. The TeV emission is believed to be caused by inverse-Compton up-scattering scattering of ambient photons by an energetic electron population. The location of a Te V steepening or cutoff in the energy spectrum reveals important details about the underlying electron population. We describe the experiment and the technique for distinguishing gamma-ray events from the much more-abundant hadronic events. We describe the calculation of the significance of the excess from the Crab and how the energy spectrum is fit.

  15. Observation and Spectral Measurements of the Crab Nebula with Milagro

    CERN Document Server

    Abdo, A A; Aune, T; Benbow, W; Berley, D; Chen, C; Christopher, G E; DeYoung, T; Dingus, B L; Ellsworth, R W; Falcone, A; Fleysher, L; Fleysher, R; Gonzalez, M M; Goodman, J A; Gordo, J B; Hays, E; Hoffman, C M; Huentemeyer, P H; Kolterman, B E; Linnemann, J T; McEnery, J E; Morgan, T; Mincer, A I; Nemethy, P; Pretz, J; Ryan, J M; Parkinson, P M Saz; Shoup, A; Sinnis, G; Smith, A J; Vasileiou, V; Walker, G P; Williams, D A; Yodh, G B

    2011-01-01

    The Crab Nebula was detected with the Milagro experiment at a statistical significance of 17 standard deviations over the lifetime of the experiment. The experiment was sensitive to approximately 100 GeV - 100 TeV gamma ray air showers by observing the particle footprint reaching the ground. The fraction of detectors recording signals from photons at the ground is a suitable proxy for the energy of the primary particle and has been used to measure the photon energy spectrum of the Crab Nebula between ~1 and ~100 TeV. The TeV emission is believed to be caused by inverse-Compton up-scattering scattering of ambient photons by an energetic electron population. The location of a TeV steepening or cutoff in the energy spectrum reveals important details about the underlying electron population. We describe the experiment and the technique for distinguishing gamma-ray events from the much more-abundant hadronic events. We describe the calculation of the significance of the excess from the Crab and how the energy spec...

  16. Teraelectronvolt pulsed emission from the Crab pulsar detected by MAGIC

    CERN Document Server

    Ahnen, M L; Antonelli, L A; Antoranz, P; Babic, A; Banerjee, B; Bangale, P; de Almeida, U Barres; Barrio, J A; González, J Becerra; Bednarek, W; Bernardini, E; Biasuzzi, B; Biland, A; Blanch, O; Bonnefoy, S; Bonnoli, G; Borracci, F; Bretz, T; Buson, S; Carmona, E; Carosi, A; Chatterjee, A; Clavero, R; Colin, P; Colombo, E; Contreras, J L; Cortina, J; Covino, S; Da Vela, P; Dazzi, F; De Angelis, A; De Lotto, B; Wilhelmi, E de Oña; Mendez, C Delgado; Di Pierro, F; Domínguez, A; Prester, D Dominis; Dorner, D; Doro, M; Einecke, S; Glawion, D Eisenacher; Elsaesser, D; Fernández-Barral, A; Fidalgo, D; Fonseca, M V; Font, L; Frantzen, K; Fruck, C; Galindo, D; López, R J García; Garczarczyk, M; Terrats, D Garrido; Gaug, M; Giammaria, P; Godinović, N; Muñoz, A González; Gora, D; Guberman, D; Hadasch, D; Hahn, A; Hanabata, Y; Hayashida, M; Herrera, J; Hose, J; Hrupec, D; Hughes, G; Idec, W; Kodani, K; Konno, Y; Kubo, H; Kushida, J; La Barbera, A; Lelas, D; Lindfors, E; Lombardi, S; Longo, F; López, M; López-Coto, R; Lorenz, E; Majumdar, P; Makariev, M; Mallot, K; Maneva, G; Manganaro, M; Mannheim, K; Maraschi, L; Marcote, B; Mariotti, M; Martínez, M; Mazin, D; Menzel, U; Miranda, J M; Mirzoyan, R; Moralejo, A; Moretti, E; Nakajima, D; Neustroev, V; Niedzwiecki, A; Rosillo, M Nievas; Nilsson, K; Nishijima, K; Noda, K; Orito, R; Overkemping, A; Paiano, S; Palacio, J; Palatiello, M; Paneque, D; Paoletti, R; Paredes, J M; Paredes-Fortuny, X; Pedaletti, G; Persic, M; Poutanen, J; Moroni, P G Prada; Prandini, E; Puljak, I; Rhode, W; Ribó, M; Rico, J; Garcia, J Rodriguez; Saito, T; Satalecka, K; Schultz, C; Schweizer, T; Shore, S N; Sillanpää, A; Sitarek, J; Snidaric, I; Sobczynska, D; Stamerra, A; Steinbring, T; Strzys, M; Takalo, L; Takami, H; Tavecchio, F; Temnikov, P; Terzić, T; Tescaro, D; Teshima, M; Thaele, J; Torres, D F; Toyama, T; Treves, A; Verguilov, V; Vovk, I; Ward, J E; Will, M; Wu, M H; Zanin, R

    2015-01-01

    Aims: To investigate the extension of the very-high-energy spectral tail of the Crab pulsar at energies above 400 GeV. Methods: We analyzed $\\sim$320 hours of good quality data of Crab with the MAGIC telescope, obtained from February 2007 until April 2014. Results: We report the most energetic pulsed emission ever detected from the Crab pulsar reaching up to 1.5 TeV. The pulse profile shows two narrow peaks synchronized with the ones measured in the GeV energy range. The spectra of the two peaks follow two different power-law functions from 70 GeV up to 1.5 TeV and connect smoothly with the spectra measured above 10 GeV by the Large Area Telescope (LAT) on board of the Fermi satellite. When making a joint fit of the LAT and MAGIC data, above 10 GeV, the photon indices of the spectra differ by 0.5$\\pm$0.1. Conclusions: We measured with the MAGIC telescopes the most energetic pulsed photons from a pulsar to date. Such TeV pulsed photons require a parent population of electrons with a Lorentz factor of at least ...

  17. Differential escape from parasites by two competing introduced crabs

    Science.gov (United States)

    Blakeslee, April M.; Keogh, Carolyn L.; Byers, James E.; Kuris, Armand M.; Lafferty, Kevin D.; Torchin, Mark E.

    2009-01-01

    Although introduced species often interact with one another in their novel communities, the role of parasites in these interactions remains less clear. We examined parasite richness and prevalence in 2 shorecrab species with different invasion histories and residency times in an introduced region where their distributions overlap broadly. On the northeastern coast of the USA, the Asian shorecrab Hemigrapsus sanguineus was discovered 20 yr ago, while the European green crab Carcinus maenas has been established for over 200 yr. We used literature and field surveys to evaluate parasitism in both crabs in their native and introduced ranges. We found only 1 parasite species infecting H. sanguineus on the US East Coast compared to 6 species in its native range, while C. maenas was host to 3 parasite species on the East Coast compared to 10 in its native range. The prevalence of parasite infection was also lower for both crabs in the introduced range compared to their native ranges; however, the difference was almost twice as much for H. sanguineus as for C. maenas. There are several explanations that could contribute to C. maenas' greater parasite diversity than that of H. sanguineus on the US East Coast, including differences in susceptibility, time since introduction, manner of introduction (vector), distance from native range, taxonomic isolation, and the potential for parasite identification bias. Our study underscores not just that non-native species lose parasites upon introduction, but that they may do so differentially, with ramifications for their direct interactions and with potential community-level influences.

  18. a Surprise from the Pulsar in the Crab Nebula

    Science.gov (United States)

    1995-11-01

    New observations of the spectrum of the rapidly spinning neutron star (the `pulsar') in the Crab Nebula have been carried out with the ESO 3.5-metre New Technology Telescope (NTT) by a group of Italian astronomers [1]. Because of greatly improved spectral resolution which allows to register even very fine details in the pulsar's spectrum, they are able to determine for the first time with high accuracy the overall dependance of the emission on wavelength, i.e. the `shape' of the spectrum. Quite unexpectedly, they also detect a hitherto unknown 100 A (10 nm) broad `absorption dip', which can be securely attributed to the pulsar. These results open an exciting new window for the study of the extreme physical processes close to a pulsar. The Nature of Pulsars It is estimated that there may be as many as 100 million neutron stars in our Galaxy. A neutron star is the superdense remnant of the extremely violent supernova explosion that occurs at the end of the life of a comparatively massive star. In fact, all stars that are more than about 6 times heavier than the Sun are believed to end their lives as supernovae. During the explosion, the central core of the dying star collapses in a few milliseconds and the matter at the centre is compressed to a density comparable to that of an atomic nucleus. Due to the enormous inward pressure, the atomic particles are squeezed together into a kind of neutron jam. The outcome is the formation of a neutron star with a diameter of 10-15 kilometres, weighing as much as the Sun. In accordance with the physical law that implies that the rotation momentum of the exploding star must be conserved, newborn neutron stars will rotate very rapidly around their axis, in some cases as fast as 100 times per second. In the same way, the new neutron star is expected to possess a strong magnetic field. Of these myriads of neutron stars, about 700 have been observed to emit radio pulses (hence the name `pulsar'). A few of these can also be detected

  19. AFSC/RACE/SAP: Detailed Crab Data From NOAA Fisheries Service 2012 Chukchi Sea Bottom Trawl Surveys

    Data.gov (United States)

    National Oceanic and Atmospheric Administration, Department of Commerce — This dataset contains detailed crab data collected from the 2012 NOAA/NMFS/AFSC/RACE crab-groundfish bottom trawl survey of the Chukchi Sea. 71 survey stations were...

  20. 77 FR 58624 - Progressive Rail Incorporated-Acquisition and Operation Exemption-Rail Lines of Crab Orchard...

    Science.gov (United States)

    2012-09-21

    ... Lines of Crab Orchard & Egyptian Railroad Company Progressive Rail Incorporated (PGR), a Class III rail carrier, has filed a verified notice of exemption under 49 CFR 1150.41 to acquire from Crab...

  1. Examination of the available catch and production data to determine if fishing curtailments will provide better crab utilization

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — Deals with the Bering Sea King crab industry. Discusses size of crabs, fishing gear, fishing localities, seasons, molting, and production techniques.

  2. Identification of tropomyosin and arginine kinase as major allergens of Portunus pelagicus (blue swimming crab).

    Science.gov (United States)

    Rosmilah, M; Shahnaz, M; Zailatul, H M Y; Noormalin, A; Normilah, I

    2012-09-01

    Crab is an important source of food allergen. Tropomyosin represents the main crab allergen and is responsible for IgE cross-reactivity between various species of crustaceans. Recently, other new crab allergens including arginine kinase have been identified. However, information on allergens of the local Portunidcrab is not available. Thus, the aim of this study was to identify the major allergens of Portunus pelagicus (blue swimming crab) using the allergenomics approach. Raw and cooked extracts of the crab were prepared from the crab meat. Protein profile and IgE binding pattern were demonstrated by sodium dodecyl sulfate-polyacrylamide gel electrophoresis (SDS-PAGE) and immunoblotting using sera from 30 patients with crab allergy. The major allergens of the crab were then identified by two-dimensional electrophoresis (2-DE), followed by mass spectrometry analysis of the peptide digests. The SDS-PAGE of raw extract revealed approximately 20 protein fractions over a wide molecular weight range, while cooked extract demonstrated fewer protein bands. The raw extract also demonstrated a higher number of IgE reactive bands than the cooked extract. A heat-resistant protein of 36 kDa has been identified as the major allergen in both raw and cooked extracts. In addition, a heat-sensitive protein of 41 kDa was also recognized as a major allergen in raw crab. The 2-DE gel profile of the raw extract demonstrated about >100 distinct proteins spots and immunoblotting of the 2-DE profile demonstrated at least 12 different major IgE reactive spots with molecular masses between 13 to 250 kDa and isoelectric point (pI) values ranging from 4.0 to 7.0. The 36 and 41 kDa proteins were identified as the crab tropomyosin and arginine kinase, respectively by mass spectrometry. Therefore, this study confirmed that tropomyosin and arginine kinase are the major allergens of the local Portunid crab, P. pelagicus.

  3. A Fast Algorithm for Burrows-Wheeler Transform Using Suffix Sorting%一种基于后缀排序快速实现Burrows-Wheeler变换的方法

    Institute of Scientific and Technical Information of China (English)

    李冰; 龙冰洁; 刘勇

    2015-01-01

    近年来,Bzip2压缩算法凭借其在压缩率方面的优势,得到了越来越多的应用,Bzip2的核心算法是Burrows-Wheeler变换(BWT), BWT能有效的将数据中相同的字符聚集到一起,为进一步压缩创造条件。在硬件实现 BWT 时,常用的基于后缀排序的算法能有效克服 BWT 消耗存储资源大的问题,该文对基于后缀排序实现BWT的方法进行了详细分析,并且在此基础上提出了一种快速实现BWT的方法后缀段算法。仿真结果表明后缀段算法在处理速度上比传统的基于后缀排序的算法有很大的提高。%Bzip2, a lossless compression algorithm, is widely used in recent years because of its high compression ratio. Burrows-Wheeler Transform (BWT) is the key factor in Bzip2. This method can gather the same symbols together. The traditional methods which are based on suffix sorting used in implement of BWT in hardware can solve the problem of memory consumption effectively. Detail analysis of BWT algorithm based on suffix sorting is given and a new methodSuffix segment method is presented in this paper. Experimental results show that the proposed method can much decrease BWT time consumption without increasing memory consumption much.

  4. Oxygen equilibria and ligand binding kinetics of erythrocruorins from two burrowing polychaetes of different modes of life, Marphysa sanguinea and Diopatra cuprea

    DEFF Research Database (Denmark)

    Weber, Roy E.; Bonaventura, J.; Sullivan, B.;

    1978-01-01

    Oxygen equilibria, ligand-binding kinetics and some other physicochemical properties are reported for erythrocruorins of two intertidal polychaetes:Marphysa sanguinea, which inhabits simple, relatively stagnant burrows, andDiopatra cuprea, which inhabits impermeable, parchment-like tubes that are...

  5. Locomotory behaviour and functional morphology of Nematostella vectensis (Anthozoa: Actiniaria: Edwardsiidae): a contribution to a comparative study of burrowing behaviour in athenarian sea anemones

    NARCIS (Netherlands)

    Williams, R.B.

    2003-01-01

    The locomotory behaviour and functional morphology of English populations of a small (<2 cm long), burrowing athenarian sea anemone, Nematostella vectensis Stephenson, 1935 (= N. pellucida Crowell, 1946), which lives in soft mud in salt marshes and creeks, are described. Objectives were to ascertain

  6. 76 FR 98 - Fisheries of the Caribbean, Gulf of Mexico, and South Atlantic; Amendment 5 to the Golden Crab...

    Science.gov (United States)

    2011-01-03

    ..., and South Atlantic; Amendment 5 to the Golden Crab Fishery Management Plan of the South Atlantic... management actions to be included in Amendment 5 to the Golden Crab Fishery Management Plan of the South... for the golden crab fishery. The purpose of this NOI is to solicit public comments on the scope...

  7. Why do Gull-billed Terns Gelochelidon nilotica feed on fiddler crabs Uca tangeri in Guinea-Bissau?

    NARCIS (Netherlands)

    Stienen, E.W.M.; Brenninkmeijer, A.; Klaassen, M.R.J.

    2008-01-01

    Wintering Gull-billed Terns Gelochelidon nilotica in Guinea Bissau mainly fed on Fiddler Crabs Uca tangeri and were occasionally seen feeding on fish and locusts. Fiddler crabs have a low energy content, requiring a large gross intake to meet daily energy demands. Fiddler crabs also have a low ratio

  8. Effect of prolonged inactivity on skeletal motor nerve terminals during aestivation in the burrowing frog, Cyclorana alboguttata.

    Science.gov (United States)

    Hudson, Nicholas J; Lavidis, Nickolas A; Choy, Peng T; Franklin, Craig E

    2005-04-01

    This study examined the effect of prolonged inactivity, associated with aestivation, on neuromuscular transmission in the green-striped burrowing frog, Cyclorana alboguttata. We compared the structure and function of the neuromuscular junctions on the iliofibularis muscle from active C. alboguttata and from C. alboguttata that had been aestivating for 6 months. Despite the prolonged period of immobility, there was no significant difference in the shape of the terminals (primary, secondary or tertiary branches) or the length of primary terminal branches between aestivators and non-aestivators. Furthermore, there was no significant difference in the membrane potentials of muscle fibres or in miniature end plate potential (EPP) frequency and amplitude. However, there was a significant decrease in evoked transmitter release characterised by a 56% decrease in mean EPP amplitude, and a 29% increase in the failure rate of nerve terminal action potentials to evoke transmitter release. The impact of this suite of neuromuscular characteristics on the locomotor performance of emergent frogs is discussed.

  9. The feeding habits of Austrolethops wardi, a gobiid fish inhabiting burrows of the thalassinidean shrimp Neaxius acanthus

    Science.gov (United States)

    Hung Liu, Ha Trieu; Kneer, Dominik; Asmus, Harald; Ahnelt, Harald

    2008-09-01

    The feeding habit of Austrolethops wardi (Gobiidae) in the seagrass beds of Barrang Lompo and Bone Batang Island in the Spermonde Archipelago, South West Sulawesi, Indonesia, was investigated through gut content analysis. The feeding preferences of this species are very similar on both islands: A. wardi, a burrow associate of Neaxius acanthus, was found to feed almost exclusively on seagrass (which was found in 100% of the investigated stomachs and made up >94% of food items). However, seagrass epiphytes (<5% of food items) and animal food (<1% of food items) occurred in the guts as well, the latter predominantly in terms of copepods and to a lesser degree in other small invertebrates. These results indicate that animal food is of little importance for A. wardi. Some specimens even contained no parts of animal food.

  10. Composite Phymatoderma from Neogene deep-marine deposits in Japan: Implications for Phanerozoic benthic interactions between burrows and the trace-makers of Chondrites and Phycosiphon

    Directory of Open Access Journals (Sweden)

    Kentaro Izumi

    2015-12-01

    Full Text Available Among composite trace fossils, one of the most common structures throughout the Phanerozoic are structures (e.g., dwelling trace, feeding trace reworked by Chondrites and/or Phycosiphon. However, differences in the nature of the reworking behaviors of these two ichnogenera remain unknown. Thus, in this study, composite Phymatoderma specimens from the Neogene deep-marine Shiramazu Formation in Japan, particularly those reworked by Chondrites and Phycosiphon, were analyzed to reveal the specific conditions that might control the activities of these trace-makers. Phymatoderma reworked by Phycosiphon is significantly larger than non-reworked Phymatoderma, whereas Phymatoderma reworked by Chondrites shows no significant difference in burrow diameter compared with non-reworked Phymatoderma. The recognized size selectivity (i.e., preference for larger burrows by the Phycosiphon trace-maker can be explained by considering the different feeding strategies of these two ichnogenera; namely deposit-feeding Phycosiphon-makers, which must have processed a significant mass of sediment to obtain sufficient organic matter, whereas chemosymbiotic Chondrites-producers did not require a lot of sediment to obtain nutrients. In order to test these interpretations, a dataset of Phanerozoic trace fossils reworked by Chondrites/Phycosiphon were compiled. Consequently, the Phycosiphon-producers’ preference toward relatively larger burrows was recognized, quantitatively supporting the results of this study. The compilation also indicates that the burrow size might have become one of the important limiting factors for the Phycosiphon-producers that tried to rework the sediments within previous subsurface burrows, at least for 80 million years.

  11. Dungeness crab survey for the Southwest Ocean Disposal Site off Grays Harbor, Washington, June 1990

    Energy Technology Data Exchange (ETDEWEB)

    Higgins, B.J.; Pearson, W.H. (Battelle/Marine Sciences Lab., Sequim, WA (United States))

    1991-09-01

    As part of the Grays Harbor Navigation Improvement Project, the Seattle District of the US Army Corps of Engineers has begun active use of the Southwest Ocean Disposal Site off Grays Harbor, Washington. This survey was to verify that the location of the area of high crab density observed during site selection surveys has not shifted into the Southeast Ocean Disposal Site. In June 1990, mean densities of juvenile Dungeness crab were 146 crab/ha within the disposal site and 609 crab/ha outside ad north of the disposal site. At nearshore locations outside the disposal site, juvenile crab density was 3275 crab/ha. Despite the low overall abundance, the spatial distribution of crab was such that the high crab densities in 1990 have remained outside the Southwest Ocean Disposal Site. The survey data have confirmed the appropriateness of the initial selection of the disposal site boundaries and indicated no need to move to the second monitoring tier. 8 refs., 9 figs., 2 tabs.

  12. Differential detection of shrimp and crab for food labeling using polymerase chain reaction.

    Science.gov (United States)

    Taguchi, Hiromu; Watanabe, Satoshi; Temmei, Yusuke; Hirao, Takashi; Akiyama, Hiroshi; Sakai, Shinobu; Adachi, Reiko; Sakata, Kozue; Urisu, Atsuo; Teshima, Reiko

    2011-04-27

    Shrimp and crab are well-known as allergenic ingredients. According to Japanese food allergy labeling regulations, shrimp species (including prawns, crayfishes, and lobsters) and crab species must be differentially declared when ≥10 ppm (total protein) of an allergenic ingredient is present. However, the commercial ELISA tests for the detection of crustacean proteins cannot differentiate between shrimp and crab. Therefore, two methods were developed to discriminate shrimp and crab: a shrimp-PCR method with postamplification digestion and a crab-PCR method that specifically amplifies a fragment of the 16S rRNA gene. The sensitivity and specificity of both PCR methods were verified by experiments using DNA extracted from 15 shrimp species, 13 crab species, krill, mysid, mantis shrimp, other food samples (cephalopod, shellfish, and fish), incurred foods, and commercial food products. Both PCR methods could detect 5 pg of DNA extracted from target species and 50 ng of genomic DNA extracted from incurred foods containing 10 ppm (μg/g) total protein of shrimp or crab. The two PCR methods were considered to be specific enough to separately detect species belonging to shrimp and crab. Although false-positive and false-negative results were obtained from some nontarget crustacean species, the proposed PCR methods, when used in conjunction with ELISA tests, would be a useful tool for confirmation of the validity of food allergy labeling and management of processed food safety for allergic patients.

  13. Summary of the 3rd LHC Crab Cavity Workshop (LHC-CC09)

    CERN Document Server

    Zimmermann, Frank; Koutchouk, Jean-Pierre; Delayen, Jean; Pierini, Paolo; Ciapala, Edmond; McIntosh, Peter; Giovannozzi, Massimo; Brüning, Oliver; Collier, Paul; Funakoshi, Yoshihiro

    2010-01-01

    The 3rd workshop on crab compensation for the LHC luminosity upgrade (LHC-CC09) was held September 16-18, 2008 at CERN, Geneva, Switzerland. The workshop was organized by joint collaboration of CERN, EUCARD, KEK and US-LARP. Approximately 50 workshop participants from 3 continents and several institutions discussed the future strategy of implementing crab cavities in the LHC

  14. The first finding of the crab Ctenocheles (Decapoda in Oligocene beds in Slovenia

    Directory of Open Access Journals (Sweden)

    Vasja Mikuž

    2003-06-01

    Full Text Available In the paper remains of crabs of genus Ctenocheles are discussed. They were found in the Oligocene grey marly mudstone at Čeplje at Vransko. This is the first evidence of this crab genus in Slovenia. In the Oligocene mudstone at ^eplje also numerous other fossilremains were registred, and also abundant septarian concretions.

  15. Dioxins, PCBs and heavy metals in Chinese mitten crabs from Dutch rivers and lakes.

    Science.gov (United States)

    Hoogenboom, Ron L A P; Kotterman, Michiel J J; Hoek-van Nieuwenhuizen, Marion; van der Lee, Martijn K; Mennes, Wim C; Jeurissen, Suzanne M F; van Leeuwen, Stefan P J

    2015-03-01

    Chinese mitten crab is an invasive species in many European rivers and lakes. Data from the UK indicated high levels of dioxins and PCBs, in particular in the brown meat in the body. This was confirmed by studies in the Netherlands, showing average levels of dioxins and PCBs in the meat in the body of 43 pg TEQ g(-1) ww in crabs caught in the large rivers. Levels in crab of lakes in the Northern part of the Netherlands were on average 3.7-fold lower. Consumption of crabs from polluted areas results in a relatively high dose of dioxins and dl-PCBs and could significantly increase the intake above the TWI. However, in general consumption of these crabs is low, even in the Asian sub-population in the Netherlands. Cadmium and lead levels were higher in crabs from contaminated areas, but for mercury and arsenic there was no clear difference. Consumption of crabs would not result in significant risks for cadmium and mercury. For lead the daily intake could be raised above the BMDL01 for neurodevelopmental toxicity, but this would only occur on a limited number of days. For arsenic the exposure would exceed the lower end of the BMDL01 values for certain cancers, but again, the infrequent consumption by most consumers reduces this risk. Furthermore, speciation showed that most arsenic in crabs was probably not a toxic inorganic form, but likely to be in an organic form.

  16. Microsatellite analysis of genetic diversity and population structure of Chinese mitten crab (Eriocheir sinensis)

    Institute of Scientific and Technical Information of China (English)

    Yumei Chang; Liqun Liang; Haitao Ma; Jianguo He; Xiaowen Sun

    2008-01-01

    Chinese mitten crab (Eriocheir sinensis) has higher commercial value as food source than any other species of Eriocheir in China.To evaluate the germplasm resources and characterize the genetic diversity and population structure of the crabs in different water systems,two stocks and two farming populations were assessed with 25 polymorphic microsallite loci available in public GenBank.Basic statistics showed that the average observed heterozygosity (Ho) amongst populations ranged from 0.5789 to 0.6824.However,a remarkable presence of inbreeding and heterozygote deficiencies were observed.To analyze population structure,pairwise FST coefficients explained only ~10.3% variability from the subdivision of mitten crab populations,the remaining variability stems from the subdivision within subpopulations.Although the four populations had slight differentiation,different allelic frequencies resulted in distinct population structures.Two stocks and one farming population were clustered together to the phylogenetic branch of Yangtze crab,with an approximate membership of 95%.Whereas,another fanning population was clustered singly to the phylogenetic branch of the Liaohe crab,with a membership of 97.1%.The tests for individual admixture showed that Yangtze crab had probably been contaminated with individuals from other water systems.Genetic relationships between populations also supported the conclusion that Yangtze crab and Liaohe crab had different gene pools in spite of the origins of the same species.

  17. Associations between dioxins/furans and dioxin-like PCBs in estuarine sediment and blue crab

    Science.gov (United States)

    Liebens, J.; Mohrherr, C.J.; Karouna-Renier, N. K.; Snyder, R.A.; Rao, K.R.

    2011-01-01

    The objective of the present study was to evaluate the relationships between the quantity, toxicity, and compositional profile of dioxin/furan compounds (PCDD/Fs) and dioxin-like polychlorinated biphenyls (DL-PCBs) in estuarine sediment and in the blue crab (Callinectes sapidus). Sediment and blue crab samples were collected in three small urban estuaries that are in relatively close proximity to each other. Results show that differences between PCDD/F and DL-PCB mass concentrations and total toxic equivalents (TEQ) toxicity in sediments of the three estuaries are reflected in those of the blue crab. TEQs are higher in the hepatopancreas of the crabs than in the sediment, but the concentration factor is inversely proportional to the TEQ in the sediments. Congener profiles in the crabs are systematically different from those in the sediments, and the difference is more pronounced for PCDD/Fs than for DL-PCBs, possibly due to differences in metabolization rates. Compared with sediment profiles, more lesser-chlorinated PCDD/Fs that have higher TEFs accumulate in crab hepatopancreas. This selective bioaccumulation of PCDD/Fs results in a TEQ augmentation in crab hepatopancreas compared with sediments. The bioaccumulation in the blue crab is also selective for PCDD/Fs over DL-PCBs. ?? 2011 Springer Science+Business Media B.V.

  18. 50 CFR 654.24 - Shrimp/stone crab separation zones.

    Science.gov (United States)

    2010-10-01

    ... 50 Wildlife and Fisheries 8 2010-10-01 2010-10-01 false Shrimp/stone crab separation zones. 654.24... Measures § 654.24 Shrimp/stone crab separation zones. Five zones are established in the management area and Florida's waters off Citrus and Hernando Counties for the separation of shrimp trawling and stone...

  19. Macroscopic anatomy of the lower respiratory system in a nocturnal burrowing rodent: African giant pouched rat (Cricetomys gambianus, Waterhouse 1840).

    Science.gov (United States)

    Ibe, C S; Salami, S O; Onyeanusi, B I

    2011-04-01

    Cricetomys gambianus is a rat that lives principally in burrows, coming out at night in search of food. The design and structure of the lower respiratory system reflects its oxygen and metabolic demand which can be attributed to its habitat. A morphological and morphometric investigation of its lower respiratory system was undertaken to document the normal anatomical features and assess its morpho-functional paradigm. Specifically, an anatomical detail of the lungs and conductive airway was described, the structures being elucidated by dissection and radiography. Evaluation of dissected specimens showed that tracheal cartilages ranged from 21 to 33 rings with an average of 25.5. They exhibited a random pattern of anastomoses between adjacent rings. Transverse diameters of the principal bronchus in the male and female rats were 3.767 and 3.759 mm respectively. The right lung consisted of four lobes while the left lung was not lobed. Bronchogram revealed that lung lobation corresponded with bronchial tree division. Inter-lobar fissures were absent on the right lung except for the ventral boarder separating the cranial lobe from the caudal lobe. The entire lungs provided stability to the heart in situ, through the cardiac notch. This study also included correlation analysis of the dimensions, weights and volumes of the lower respiratory organs with the nose-rump length and body weight of 18 African giant pouched rats of both sexes. The relationship of the anatomy of the lower respiratory system of the rodent to the oxygen tension in their burrow and to their energy utilization is discussed.

  20. Urban pollution of sediments: Impact on the physiology and burrowing activity of tubificid worms and consequences on biogeochemical processes.

    Science.gov (United States)

    Pigneret, M; Mermillod-Blondin, F; Volatier, L; Romestaing, C; Maire, E; Adrien, J; Guillard, L; Roussel, D; Hervant, F

    2016-10-15

    In urban areas, infiltration basins are designed to manage stormwater runoff from impervious surfaces and allow the settling of associated pollutants. The sedimentary layer deposited at the surface of these structures is highly organic and multicontaminated (mainly heavy metals and hydrocarbons). Only few aquatic species are able to maintain permanent populations in such an extreme environment, including the oligochaete Limnodrilus hoffmeisteri. Nevertheless, the impact of urban pollutants on these organisms and the resulting influence on infiltration basin functioning remain poorly studied. Thus, the aim of this study was to determine how polluted sediments could impact the survival, the physiology and the bioturbation activity of L. hoffmeisteri and thereby modify biogeochemical processes occurring at the water-sediment interface. To this end, we conducted laboratory incubations of worms, in polluted sediments from infiltration basins or slightly polluted sediments from a stream. Analyses were performed to evaluate physiological state and burrowing activity (X-ray micro-tomography) of worms and their influences on biogeochemical processes (nutrient fluxes, CO2 and CH4 degassing rates) during 30-day long experiments. Our results showed that worms exhibited physiological responses to cope with high pollution levels, including a strong ability to withstand the oxidative stress linked to contamination with heavy metals. We also showed that the presence of urban pollutants significantly increased the burrowing activity of L. hoffmeisteri, demonstrating the sensitivity and the relevance of such a behavioural response as biomarker of sediment toxicity. In addition, we showed that X-ray micro-tomography was an adequate technique for accurate and non-invasive three-dimensional investigations of biogenic structures formed by bioturbators. The presence of worms induced stimulations of nutrient fluxes and organic matter recycling (between +100% and 200% of CO2 degassing rate

  1. Relationships between yolk androgens and nest density, laying date, and laying order in Western Burrowing Owls (Athene cunicularia hypugaea)

    Science.gov (United States)

    Welty, J.L.; Belthoff, J.R.; Egbert, J.; Schwabl, H.

    2012-01-01

    Increases in yolk androgens within and among avian clutches have been correlated with decreased incubation time, increased aggression within a nest, increased begging behaviour, decreased immune response, and decreased life span. Although the mechanisms that lead to variability in yolk androgens within and between clutches are not completely known, yolk androgens can be a function of both social and environmental conditions. We were interested in if and how nesting density, laying date, and laying order influenced yolk androgens in Western Burrowing Owls (Athene cunicularia hypugaea (Bonaparte, 1825)) in which nest density varies considerably. In 2006 and 2007, we used radioimmunoassay to quantify the concentrations of testosterone, 5a-dihydrotestosterone, and androstenedione in the egg yolks from one early and one latelaid egg in 47 nests of Burrowing Owls located in the Morley Nelson Snake River Birds of Prey National Conservation Area in southern Idaho. Nesting density had no detectable effect on yolk androgens. Yolk androgens varied temporally and peaked in the middle of the laying season while being low before and after this time period. Within nests, late-laid eggs had higher testosterone and dihydrotestosterone than early-laid eggs; adrostendione exhibited a similar pattern in one but not both years of our study. It is possible that the seasonal pattern in yolk androgens that we observed is related to aspects of mate quality for females or declining chances of fledging success for later nesting females, whereas rises in egg androgens between early and late eggs within clutches could reflect a mechanism to assist nestlings from late-laid eggs that hatch one to several days after their siblings to better compete for resources within the nest or promote survival in the presence of larger siblings.

  2. Chitin-Chitosan Yield of Freshwater Crab (Potamon potamios, Olivier 1804 Shell

    Directory of Open Access Journals (Sweden)

    Yildiz Bolat*, Şengül Bilgin, Ali Günlü, Levent Izci, Seval Bahadır Koca, Soner Çetinkaya1 and Habil Uğur Koca

    2010-10-01

    Full Text Available In this study, freshwater crab (Potamon potamios, Olivier 1804, economically unevaluated, was used to obtain chitin-chitosan. Chitin-chitosan was extracted with standardized modified chemical method. Chitosan was extracted with demineralization, deproteinization, decoloration (chitin and deacetylation (chitosan. Grinded shell was obtained as 60% after boiling, drying and grinding processes. Chitosan yield of crab shell was determined as 4.65% from grinded crab shell after demineralization (yield is 34.32%, deproteinization (yield is 7.25%, decoloration (yield is 6.83% and deacetylation processes. Moreover, freshwater crab stock was estimated with catch per unit effort (CPUE data in Eğirdir Lake for calculation of quantity of chitosan. Population size was estimated between 12.85±11.88 and 23.86±25.39 tones and freshwater crab was determined as an appropriate crustacean to obtain chitin-chitosan.

  3. Did the Crab Pulsar Undergo a Small Glitch in 2006 late March/early April?

    CERN Document Server

    Vivekanand, M

    2016-01-01

    On 2006 August 23 the Crab Pulsar underwent a glitch, that was reported by the Jodrell Bank and the Xinjiang radio observatories. Neither data are available to the public. However, the Jodrell group publishes monthly arrival times of the Crab Pulsar pulse (their actual observations are done daily), using which it is shown that about five months earlier, the Crab Pulsar most probably underwent a small glitch, which has not been reported before. Neither observatory discusses the detailed analysis of data from 2006 March to August; either they may not have detected this small glitch, or may have attributed it to timing noise in the Crab Pulsar. The above result is verified using X-ray data from the RXTE observatory. If this is indeed true, this may probably be the smallest glitch observed in the Crab Pulsar so far, whose implications are discussed. This work addresses the confusion possible between small magnitude glitches and timing noise in pulsars.

  4. Metabolism of pectenotoxins in brown crabs Cancer pagurus fed with blue mussels Mytilus edulis

    Institute of Scientific and Technical Information of China (English)

    LI Zhaoxin

    2009-01-01

    We investigated the metabolism of pectenotoxins in brown crabs (Cancer pagurus). The crabs were fed with blue mussels (Mytilus edulis) for 21 d then depurated for 42 d. We extracted the toxins from the digestive glands of contaminated crabs, uncontaminated crabs (control group), and the meat of blue mussels using methanol. Extracts of the crab digestive glands were fractionated by liquid-liquid partitioning and solid phase extraction. The fractions were analyzed by liquid chromatography coupled with tandem mass spectrometry (LC-MS/MS) and liquid chromatography coupled with ion-trap mass spectrometry (LC-MSn). We detected a new PTX-like compound, designated as metabolite-1. The MS2 mass spectrum of the metabolite-1 [M+Na]+ ion at m/z 897.5 revealed fragment ions at m/z 853.5 and 555.5, typical of those exhibited by other pectenotoxins.

  5. Effects of tributyltin exposure in hermit crabs: Clibanarius vittatus as a model.

    Science.gov (United States)

    Sant'Anna, Bruno Sampaio; Santos, Dayana Moscardi Dos; Marchi, Mary Rosa Rodrigues de; Zara, Fernando José; Turra, Alexander

    2012-03-01

    Tributyltin (TBT) contamination affects the reproductive system of many species of invertebrates worldwide. The present study was designed to evaluate the effects of exposure to TBT pollution on the reproduction of the hermit crab Clibanarius vittatus. An orthogonal experiment was designed with two treatments: contamination (with or without TBT in the food) and crab sex (males and females). The animals were reared in the laboratory for nine months, and macroscopic and histological analyses of reproductive organs were carried out after the end of the experiment. Tributyltin was recorded in exposed crabs, but no morphological alterations were detected in the gonads of males, regardless of whether they were exposed to TBT. In contrast, females exposed to TBT displayed disorganization and atrophy of their ovaries, thus directly affecting reproduction in this hermit crab species. This effect observed in female hermit crabs may harm populations located in harbor regions, where TBT concentration is high, even after the worldwide TBT ban.

  6. Pharmacokinetics of cefovecin (Convenia) in white bamboo sharks (Chiloscyllium plagiosum) and Atlantic horseshoe crabs (Limulus polyphemus).

    Science.gov (United States)

    Steeil, James C; Schumacher, Juergen; George, Robert H; Bulman, Frank; Baine, Katherine; Cox, Sherry

    2014-06-01

    Cefovecin was administered to six healthy adult white bamboo sharks (Chiloscyllium plagiosum) and six healthy adult Atlantic horseshoe crabs (Limulus polyphemus) to determine its pharmacokinetics in these species. A single dose of cefovecin at 8 mg/kg was administered subcutaneously in the epaxial region of the bamboo sharks and in the proximal articulation of the lateral leg of the horseshoe crabs. Blood and hemolymph samples were collected at various time points from bamboo sharks and Atlantic horseshoe crabs. High performance liquid chromatography was performed to determine plasma levels of cefovecin. The terminal halflife of cefovecin in Atlantic horseshoe crabs was 37.70 +/- 9.04 hr and in white bamboo sharks was 2.02 +/- 4.62 hr. Cefovecin concentrations were detected for 4 days in white bamboo sharks and for 14 days in Atlantic horseshoe crabs. No adverse effects associated with cefovecin administration were seen in either species.

  7. AFSC/RACE/SAP/Long: Data from: Habitat, predation, growth, and coexistence: Could interactions between juvenile red and blue king crabs limit blue king crab productivity?

    Data.gov (United States)

    National Oceanic and Atmospheric Administration, Department of Commerce — This data set is from a series of laboratory experiments examining the interactions between red and blue king crabs and habitat. We examined how density and predator...

  8. Effects of ocean acidification on juvenile red king crab (Paralithodes camtschaticus) and Tanner crab (Chionoecetes bairdi) growth, condition, calcification, and survival.

    Science.gov (United States)

    Long, William Christopher; Swiney, Katherine M; Harris, Caitlin; Page, Heather N; Foy, Robert J

    2013-01-01

    Ocean acidification, a decrease in the pH in marine waters associated with rising atmospheric CO2 levels, is a serious threat to marine ecosystems. In this paper, we determine the effects of long-term exposure to near-future levels of ocean acidification on the growth, condition, calcification, and survival of juvenile red king crabs, Paralithodes camtschaticus, and Tanner crabs, Chionoecetes bairdi. Juveniles were reared in individual containers for nearly 200 days in flowing control (pH 8.0), pH 7.8, and pH 7.5 seawater at ambient temperatures (range 4.4-11.9 °C). In both species, survival decreased with pH, with 100% mortality of red king crabs occurring after 95 days in pH 7.5 water. Though the morphology of neither species was affected by acidification, both species grew slower in acidified water. At the end of the experiment, calcium concentration was measured in each crab and the dry mass and condition index of each crab were determined. Ocean acidification did not affect the calcium content of red king crab but did decrease the condition index, while it had the opposite effect on Tanner crabs, decreasing calcium content but leaving the condition index unchanged. This suggests that red king crab may be able to maintain calcification rates, but at a high energetic cost. The decrease in survival and growth of each species is likely to have a serious negative effect on their populations in the absence of evolutionary adaptation or acclimatization over the coming decades.

  9. Carbon and nitrogen balance of leaf-eating sesarmid crabs ( Neoepisesarma versicolor) offered different food sources

    Science.gov (United States)

    Thongtham, Nalinee; Kristensen, Erik

    2005-10-01

    Carbon and nitrogen budgets for the leaf-eating crab, Neoepisesarma versicolor, were established for individuals living on pure leaf diets. Crabs were fed fresh (green), senescent (yellow) and partly degraded (brown) leaves of the mangrove tree Rhizophora apiculata. Ingestion, egestion and metabolic loss of carbon and nitrogen were determined from laboratory experiments. In addition, bacterial abundance in various compartments of the crabs' digestive tract was enumerated after dissection of live individuals. Ingestion and egestion rates (in terms of dry weight) were highest, while the assimilation efficiency was poorest for crabs fed on brown leaves. The low assimilation efficiency was more than counteracted by the high ingestion rate providing more carbon for growth than for crabs fed green and yellow leaves. In any case, the results show that all types of leaves can provide adequate carbon while nitrogen was insufficient to support both maintenance (yellow leaves) and growth (green, yellow and brown leaves). Leaf-eating crabs must therefore obtain supplementary nitrogen by other means in order to meet their nitrogen requirement. Three hypotheses were evaluated: (1) crabs supplement their diet with bacteria and benthic microalgae by ingesting own faeces and/or selective grazing at the sediment surface; (2) assimilation of symbiotic nitrogen-fixing bacteria in the crabs' own intestinal system; and (3) nitrogen storage following occasional feeding on animal tissues (e.g. meiofauna and carcasses). It appears that hypothesis 1 is of limited importance for N. versicolor since faeces and sediment can only supply a minor fraction of the missing nitrogen due to physical constraints on the amount of material the crabs can consume. Hypothesis 2 can be ruled out because tests showed no nitrogen fixation activity in the intestinal system of N. versicolor. It is therefore likely that leaf-eating crabs provide most of their nitrogen requirement from intracellular deposits

  10. POSSIBILITIES OF CULTURING BIG SEA CRABS (LOBSTERS, SPINY LOBSTERS)

    OpenAIRE

    Ivančica Strunjak-Perović; Emin Teskeredžić; Marija Tomec

    1999-01-01

    By the end of the 19 th century an experimental work on culturing big sea crabs began in Europe and North America. Great demand for their flesh as well as their high price urged many institutions to explore the possibilities of a commercial production in varios parts of the world. Lobsters (Homarus sp.) were mainly used for experimenting, so that the most data available refer to them. Because of the complicated larva stage spiny lobster culturing is mainly being carried out in experimental ci...

  11. Peering into the Heart of the Crab Nebula

    Science.gov (United States)

    2000-01-01

    In the year 1054 A.D., Chinese astronomers were startled by the appearance of a new star, so bright that it was visible in broad daylight for several weeks. Today, the Crab Nebula is visible at the site of the 'Guest Star.' Located about 6,500 light-years from Earth, the Crab Nebula is the remnant of a star that began its life with about 10 times the mass of our own Sun. Its life ended on July 4, 1054 when it exploded as a supernova. In this image, NASA's Hubble Space Telescope has zoomed in on the center of the Crab to reveal its structure with unprecedented detail. The Crab Nebula data were obtained by Hubble's Wide Field and Planetary Camera 2 in 1995. Images taken with five different color filters have been combined to construct this new false-color picture. Resembling an abstract painting by Jackson Pollack, the image shows ragged shards of gas that are expanding away from the explosion site at over 3 million miles per hour. The core of the star has survived the explosion as a pulsar, visible in the Hubble image as the lower of the two moderately bright stars to the upper left of center. The pulsar is a neutron star that spins on its axis 30 times a second. It heats its surroundings, creating the ghostly diffuse bluish-green glowing gas cloud in its vicinity, including a blue arc just to its right. The colorful network of filaments is the material from the outer layers of the star that was expelled during the explosion. The picture is somewhat deceptive in that the filaments appear to be close to the pulsar. In reality, the yellowish green filaments toward the bottom of the image are closer to us, and approaching at some 300 miles per second. The orange and pink filaments toward the top of the picture include material behind the pulsar, rushing away from us at similar speeds. The various colors in the picture arise from different chemical elements in the expanding gas, including hydrogen (orange), nitrogen (red), sulfur (pink), and oxygen (green). The shades

  12. On Calibrations Using the Crab Nebula as a Standard Candle

    Science.gov (United States)

    Weisskopf, Martin; Guainazzi, Matteo; Jahoda, Keith; Shaposhnikov, Nikolai; ODell, Stephen; Zavlin, Vyacheslav; Wilson-Hodge, Colleen; Elsner, Ronald

    2009-01-01

    Inspired by a recent paper (Kirsch et al. 2005) on possible use of the Crab Nebula as a standard candle for calibrating X-ray response func tions, we examine possible consequences of intrinsic departures from a single (absorbed) power law upon such calibrations. We limited our analyses to three more modern X-ray instruments -- the ROSAT/PSPC, th e RXTE/PCA, and the XMM-Newton/EPIC-pn. The results are unexpected an d indicate a need to refine two of the three response functions studi ed. The implications for Chandra will be discussed.

  13. OBSERVATION AND SPECTRAL MEASUREMENTS OF THE CRAB NEBULA WITH MILAGRO

    Energy Technology Data Exchange (ETDEWEB)

    Abdo, A. A. [Department of Physics and Astronomy, Michigan State University, East Lansing, MI 48824 (United States); Allen, B. T.; Chen, C. [Department of Physics and Astronomy, University of California, Irvine, CA 92697 (United States); Atkins, R. [Department of Physics, University of Wisconsin, 1150 University Ave, Madison, WI 53706 (United States); Aune, T.; Benbow, W.; Coyne, D. G.; Dorfan, D. E. [Santa Cruz Institute for Particle Physics, University of California, Santa Cruz, CA 95064 (United States); Berley, D.; Blaufuss, E.; Bussons, J. [Department of Physics, University of Maryland, College Park, MD 20742 (United States); Bonamente, E.; Galbraith-Frew, J. [Department of Physics, Michigan Technological University, Houghton, MI 49931 (United States); Christopher, G. E.; Fleysher, L.; Fleysher, R. [Department of Physics, New York University, New York, NY 10003 (United States); DeYoung, T.; Falcone, A. [Department of Physics, Pennsylvania State University, University Park, PA 16802 (United States); Dingus, B. L. [Group P-23, Los Alamos National Laboratory, Los Alamos, NM 87545 (United States); Ellsworth, R. W. [School of Physics, Astronomy and Computational Sciences, George Mason University, Fairfax, VA 22030 (United States); and others

    2012-05-01

    The Crab Nebula was detected with the Milagro experiment at a statistical significance of 17 standard deviations over the lifetime of the experiment. The experiment was sensitive to approximately 100 GeV-100 TeV gamma-ray air showers by observing the particle footprint reaching the ground. The fraction of detectors recording signals from photons at the ground is a suitable proxy for the energy of the primary particle and has been used to measure the photon energy spectrum of the Crab Nebula between {approx}1 and {approx}100 TeV. The TeV emission is believed to be caused by inverse-Compton upscattering of ambient photons by an energetic electron population. The location of a TeV steepening or cutoff in the energy spectrum reveals important details about the underlying electron population. We describe the experiment and the technique for distinguishing gamma-ray events from the much more-abundant hadronic events. We describe the calculation of the significance of the excess from the Crab and how the energy spectrum is fitted. The differential photon energy spectrum, including the statistical errors from the fit, obtained using a simple power-law hypothesis for data between 2005 September and 2008 March is (6.5 {+-} 0.4) Multiplication-Sign 10{sup -14}(E/10 TeV){sup -3.1{+-}0.1}(cm{sup 2} s TeV ){sup -1} between {approx}1 TeV and {approx}100 TeV. Allowing for a possible exponential cutoff, the photon energy spectrum is fitted as (2.5{sup +0.7}{sub -0.4}) Multiplication-Sign 10{sup -12}(E/3 TeV){sup -2.5{+-}0.4}exp (- E/32{sup +39}{sub -18} TeV) (cm{sup 2} s TeV){sup -1}. The results are subject to an {approx}30% systematic uncertainty in the overall flux and an {approx}0.1 systematic uncertainty in the power-law indices quoted. Uncertainty in the overall energy scale has been absorbed into these errors. Fixing the spectral index to values that have been measured below 1 TeV by IACT experiments (2.4-2.6), the fit to the Milagro data suggests that Crab exhibits a

  14. Impacts to Dungeness Crab from the Southwest Washington Littoral Drift Restoration Project

    Energy Technology Data Exchange (ETDEWEB)

    Williams, Greg D.; Kohn, Nancy P.; Pearson, Walter H.; Skalski, J R.

    2005-11-09

    The Benson Beach littoral drift restoration project is a demonstration project that will replenish sand on Benson Beach, the public beach north of the North Jetty at the mouth of the Columbia River (MCR), using material dredged from the river during normal maintenance dredging of the navigational channel. A U.S. Army Corps of Engineers (Corps) proposal involves pumping the material from a sump area on the south side of the jetty to Benson Beach using a cutter suction dredge, also known as a pipeline dredge. If this one-time demonstration project proves feasible and successful, up to a million cubic yards of sediment could be used to replenish the outer coast littoral drift system in successive years by the same process. The primary goal of this study was to assess the potential risk of impacts to Dungeness crab from the proposed Benson Beach littoral drift restoration process of using the cutter suction dredge to move sediment from the proposed sump area on one side of the North Jetty to the beach on the other side of the jetty. Because there are no direct measurements of crab entrainment by pipeline dredge operating outside of the lower Columbia River navigation channel, dredge impacts for the proposed demonstration project were estimated using a modification of the dredge impact model (DIM) of Armstrong et al. (1987). The model estimates adult equivalent loss (AEL) of crabs using crab population density from trawl surveys, dredge project information (gear type, season, location, volume), and an entrainment function relating crab population density to entrainment by the dredge. The input used in applying the DIM to the Benson Beach littoral drift restoration included the specific dredging scenario provided by the Corps, existing data on crab density in previously proposed sump areas, and a series of entrainment functions. A total of fourteen scenarios were modeled and the outcomes compared with six reference scenarios intended to represent realistic to worst cases

  15. Comparison of Gender Differences in Nutritional Value and Key Odor Profile of Hepatopancreas of Chinese Mitten Crab (Eriocheir Sinensis).

    Science.gov (United States)

    Wu, Na; Wang, Xi-Chang

    2017-02-01

    The nutritional value and key odor profile of hepatopancreas of Chinese mitten crab (Eriocheir sinensis) was evaluated, and gender differences in terms of edible yield, proximate composition, different lipid fractions, fatty acid composition, and key odor compounds were compared. Total lipids were separated into different lipid fractions using silicic acid columns. And odorants were detected by monolithic material sorptive extraction coupled with gas chromatography-mass spectrometry-olfactometry (GC-MS-O). Furthermore, detected frequency and odor intensity method of GC-O were applied to select key odor compounds in steamed E. sinensis. The results showed male crabs (34.06%) had a higher edible yield than female crabs (32.42%). Hepatopancreas was the most abundant portion in both genders, which contained high lipid content. Higher contents of triacylglycerols, sterols, free fatty acids, and phosphatidylethanolamine in female crabs resulted in a significant higher total lipid content in females (40.05%) than in males (37.94%). Meanwhile, total monounsaturated fatty acids and polyunsaturated fatty acids of female crabs had significant higher content than male crabs, and the ratio of docosahexaenoic acid to eicosapentaenoic acid of female crabs (1.26) showed higher value than male crabs (1.18), which indicated that female crabs had higher nutritional value than male crabs. For further analyses, unique odor compounds detected in female crabs showed a plant/vegetable-like odor, while sweety, corn-like odor in male crabs. These suggested that hepatopancreases of female crabs are healthier than male crabs for human consumption, however, they may be favored by different consumers individually due to their unique aroma profiles.

  16. Utilization of waste crab shell (Scylla serrata) as a catalyst in palm olein transesterification.

    Science.gov (United States)

    Boey, Peng-Lim; Maniam, Gaanty Pragas; Hamid, Shafida Abd

    2009-01-01

    Aquaculture activity has increased the population of crab, hence increasing the generation of related wastes, particularly the shell. In addition, the number of molting process in crabs compounds further the amount of waste shell generated. As such, in the present work, the application of the waste crab shell as a source of CaO in transesterification of palm olein to biodiesel (methyl ester) was investigated. Preliminary XRD results revealed that thermally activated crab shell contains mainly CaO. Parametric study has been investigated and optimal conditions were found to be methanol/oil mass ratio, 0.5:1; catalyst amount, 4 wt. %; and reaction temperature, 338 K. As compared to laboratory CaO, the catalyst from waste crab shell performs well, thus creating another low-cost catalyst source for producing biodiesel as well as adding value to the waste crab shell. Reusability of crab shell CaO has also been studied and the outcome confirmed that the catalyst is capable to be reutilized up to 11 times, without any major deterioration.

  17. Levels of Cadmium in White and Brown Meat of Warty Crab (Eriphia verrucosa).

    Science.gov (United States)

    Ariano, A; Voi, A Lo; D'Ambola, M; Marrone, R; Cacace, D; Severino, L

    2015-12-01

    European Union regulations that establish the maximum cadmium (Cd) levels for crab take into account only concentrations found in crab muscle (white meat), mainly from appendages; therefore, other organs and tissues (brown meat) are excluded. The objective of the present study was to evaluate Cd levels in both white and brown crab meat, in order to achieve a more complete assessment of health risk related to human consumption of warty crab. Microwave digestion and atomic absorption spectrometry were used to determine Cd concentrations in warty crab (Eriphia verrucosa) samples collected from the southern Tyrrhenian Sea in Italy. Cd concentrations in all samples of white crab meat were found to be very low (below the limit of quantification), although brown crab meat showed significantly higher Cd concentrations (up to 5.629 mg/kg wet weight; mean value, 1.465 mg/kg). Thus, the consumption of brown meat, common among certain populations of the Mediterranean region, where whole crustaceans are traditionally eaten, substantially increased Cd intake, resulting in alarmingly high estimated weekly intake values.

  18. The 1997 Event in the Crab Pulsar in X-rays

    CERN Document Server

    Vivekanand, M

    2016-01-01

    In October 1997, radio pulses from the Crab Pulsar underwent abnormal delay. This was reported by two radio observatories, both of which explained this frequency dependent and time varying delay as being due to refractive effects of ionized shells in the Crab Nebula. Both groups also noted that, curiously and confusingly coincident with the frequency dependent delay, the Crab Pulsar also underwent an unusual slowing down, which they believed to be unrelated to the Crab Nebula and instead intrinsic to the Crab Pulsar, resulting in an additional delay that was frequency independent. However, it now appears that one of the groups attributes the frequency independent delay also to refractive effects. This work aims to verify whether at least a part of the frequency independent delay is indeed due to intrinsic slowing down of the Crab Pulsar. Timing analysis of the Crab Pulsar's October 1997 event has been done in X-rays, which are not delayed by the refractive and diffractive effects that affect radio waves; at X...

  19. Timing and route of migration of mature female blue crabs in a tidal estuary.

    Science.gov (United States)

    Eggleston, David B; Millstein, Erika; Plaia, Gayle

    2015-05-01

    Information on migration patterns is critical to using no-take migratory corridors and marine reserves to protect the spawning stock of commercially exploited species. Both active and passive acoustic tracking methods quantified movement of commercially and ecologically important blue crabs in the White Oak River estuary, NC, USA. We targeted post-mating female crabs migrating down-estuary to oceanic spawning grounds. Crabs travelled approximately 14.1 km mainly in deeper channels and over 12-26 days from mating areas to spawning grounds. No crabs were detected migrating down-estuary in the autumn and only 30% were detected migrating down-estuary in spring. None of the crabs detected near spawning grounds were detected or recaptured back up-estuary, suggesting that they either (i) do not return to the estuary after a one to two week period in the spawning area or (ii) were captured by fishermen. The results from this study demonstrate that (1) acoustic transmitters coupled with passive acoustic receivers provided reliable and valuable data on migration patterns of mature female blue crabs and (2) mature female blue crabs are capable of migrating primarily within deep channels to spawning grounds shortly after insemination.

  20. Intraguild predation by shore crabs affects mortality, behavior, growth, and densities of California horn snails

    Science.gov (United States)

    Lorda, J.; Hechinger, R.F.; Cooper, S. D.; Kuris, A. M.; Lafferty, Kevin D.

    2016-01-01

    The California horn snail, Cerithideopsis californica, and the shore crabs, Pachygrapsus crassipesand Hemigrapsus oregonensis, compete for epibenthic microalgae, but the crabs also eat snails. Such intraguild predation is common in nature, despite models predicting instability. Using a series of manipulations and field surveys, we examined intraguild predation from several angles, including the effects of stage-dependent predation along with direct consumptive and nonconsumptive predator effects on intraguild prey. In the laboratory, we found that crabs fed on macroalgae, snail eggs, and snails, and the size of consumed snails increased with predator crab size. In field experiments, snails grew less in the presence of crabs partially because snails behaved differently and were buried in the sediment (nonconsumptive effects). Consistent with these results, crab and snail abundances were negatively correlated in three field surveys conducted at three different spatial scales in estuaries of California, Baja California, and Baja California Sur: (1) among 61 sites spanning multiple habitat types in three estuaries, (2) among the habitats of 13 estuaries, and (3) among 34 tidal creek sites in one estuary. These results indicate that shore crabs are intraguild predators on California horn snails that affect snail populations via predation and by influencing snail behavior and performance.

  1. The biosynthesis, absorption, and origin of cholesterol and plant sterols in the Florida land crab.

    Science.gov (United States)

    Douglass, T S; Connor, W E; Lin, D S

    1981-08-01

    In order to study the biosynthesis, composition, and origin of sterols in the Florida land crabs, Cardisoma guanhumi (Latreille), we fed 17 male crabs either a cholesterol-free or a high cholesterol diet for 2 to 7 weeks. The origin of sterols in these crabs, whether from biosynthesis or from the diet, was determined by tahree procedures: the incorporation of isotopic mevalonate into the cholesterol when the diet was cholesterol-free; the absorption of isotopic cholesterol and sitosterol from the diet; the cholesterol and plant sterol concentrations of hepatopancreas, plasma, and muscle under conditions of cholesterol-free and high cholesterol diets. In addition, the interconversion of cholesterol and sitosterol was investigated. Dietary sterols of plant and animal sources were readily absorbed and provided the major source of sterols for this species of crab. The biosynthesis of cholesterol from mevalonate in this crab was minimal. However, cholesterol was synthesized from dietary sitosterol by dealkylation. Cholesterol and the three plant sterols (24 epsilon-methyl cholesterol, stigmasterol, and sitosterol) were found in the hepatopancreas, plasma, and muscle of the crab. Plant sterols contributed from 9 to 37% of the total sterols in the hepatopancreas, plasma, and muscle of the crabs fed a cholesterol-free diet.

  2. Summary of CARE-HHH Mini-Workshop on LHC Crab Cavity Validation, 21 August 2008

    CERN Document Server

    Calaga, R; Garoby, R; Linnecar, T; Tomás, R; Zimmermann, F; CERN. Geneva. BE Department

    2008-01-01

    A global LHC crab-cavity collaboration is rapidly advancing the R&D of a complete crab cavity cryomodule and performing the associated beam dynamics simulations compatible with a prototype test in the phase 0/I upgrade with the aim of establishing a full crab crossing scheme for the phase II upgrade of the LHC. A one day CARE-HHH mini-workshop was held on August 21, 2008 at CERN to discuss crab crossing in the LHC phase 0/I & II upgrades and this reports summarizes the activities of the four sessions that took place during the workshop. The goals of this crab-cavity workshop were fourfold: (1) to discuss prospects of crab cavities in LHC upgrades (2) to review the status of the cryomodule development and beam dynamics, (3) to establish validity requirements for LHC crab cavities which need to be demonstrated prior to their installation into the LHC, and (4) to provide guidance & coordination for the global collaborators.

  3. POSSIBILITIES OF CULTURING BIG SEA CRABS (LOBSTERS, SPINY LOBSTERS

    Directory of Open Access Journals (Sweden)

    Ivančica Strunjak-Perović

    1999-09-01

    Full Text Available By the end of the 19 th century an experimental work on culturing big sea crabs began in Europe and North America. Great demand for their flesh as well as their high price urged many institutions to explore the possibilities of a commercial production in varios parts of the world. Lobsters (Homarus sp. were mainly used for experimenting, so that the most data available refer to them. Because of the complicated larva stage spiny lobster culturing is mainly being carried out in experimental circumstances. Despite the promissing results this aquacultural activity faces many problems (long time until they achieve a commercial size, loss of eggs due to stress sensitivity during the process of moulting, canibalism. In order to minimize these problems various researches are being carried out, like temperature influence, influence of light, way of feeding, hormonal regulation of moulting frequency. Although both lobster and spiny lobsters live in the Adriatic Sea, there are no data on their culturing in our contry. Concernig conditions in our sea there are realistic possibilities for crabs production development. In this way this delicacy would be more affordable to broader population and could be a highly rated export product.

  4. Nutritional quality and safety of cooked edible crab (Cancer pagurus).

    Science.gov (United States)

    Maulvault, Ana Luísa; Anacleto, Patrícia; Lourenço, Helena Maria; Carvalho, Maria Luísa; Nunes, Maria Leonor; Marques, António

    2012-07-15

    Edible crab (Cancer pagurus) is one of the most important crustaceans consumed in Southern European countries, either as boiled or steamed cooked product. So far, the majority of studies assessing health benefits and risks associated to seafood consumption have been carried out in raw products, despite being generally cooked before consumption, and mostly in muscle tissue. Therefore, the aim of this study was to assess the effect of steaming and boiling on the chemical safety and nutritional value of C. pagurus caught in spring and summer. Generally, chemical and elemental composition of brown meat (tissue in the body cavity comprising mainly gonads and hepatopancreas) differed significantly from muscle (white meat in claws and legs). Additionally, the same tissue revealed differences in chemical and elemental composition of raw and cooked C. pagurus, likely due to water leaching. In contrast, few variations between seasons were observed. The results revealed that the consumption of cooked edible crab muscle should be promoted, whereas brown meat ingestion should be done parsimoniously.

  5. Physicochemical and biofunctional properties of crab chitosan nanoparticles.

    Science.gov (United States)

    Nguyen, The Han; Kwak, Hae Soo; Kim, Sang Moo

    2013-08-01

    The physicochemical and biofunctional properties of crab chitosan nanoparticles of two different sizes (Nano A and B) manufactured by dry milling method were evaluated for commercialization. The deacetylation degrees (DD) of Nano A, B and the control chitosan were 90.9, 93.0, and 92.7% respectively whereas their molecular weights (M(w)) were 43.9, 44.7 and 208.8 kDa. The average sizes of the dispersed Nano A, B and the control chitosan in cetyltrimethylammonium chloride were 735.9, 849.4 and 2,382.4 nm, respectively, which were lower than 1441.7, 2935.6 and 6832.9 nm of the intact chitosans. Chitosan nanoparticles had mild tyrosinase, antioxidant and angiotensin I converting enzyme (ACE), but weak collagenase, elastase and beta-glucuronidase inhibitory activity. However, Nano A had strong alpha-glucosidase inhibitory activity, which was comparable to that of acarbose, a commercial alpha-glucosidase inhibitor. In addition, the minimum inhibitory concentrations (MICs) of chitosan and its nanoparticles ranged from 30 to > 200 microg/mL against each four gram-positive and gram-negative bacteria. Therefore, crab chitosan nanoparticles could be used as a nutraceutical, cosmeceutical or pharmaceutical product.

  6. PoGOLite measurement of Crab polarisation and future plans

    Science.gov (United States)

    Pearce, Mark

    2016-07-01

    (For the PoGOLite Collaboration) The PoGOLite Pathfinder is a balloon-borne hard X-ray polarimeter designed for the observation of bright, ~1 Crab, sources. Polarisation is determined by measuring the azimuthal Compton scattering angle of incident X-rays in an array of plastic scintillators housed in an BGO anticoincidence well. The PoGOLite Pathfinder was launched from the SSC Esrange Space Centre in July 2013 resulting in a near-circumpolar flight of two weeks duration. The linear polarisation of hard X-ray emissions from the Crab was measured in a previously unexplored energy interval, 20-120 keV. The polarimetric response was characterised prior to flight using both polarised and unpolarised calibration sources. Systematic effects were addressed through observations of a background field. An upgraded polarimeter, PoGO+, is scheduled to fly in summer 2016 from Esrange. Results from the 2013 Pathfinder flight and prospects for the 2016 flight will be discussed.

  7. OBSERVATIONS OF THE CRAB NEBULA'S ASYMMETRICAL DEVELOPMENT

    Energy Technology Data Exchange (ETDEWEB)

    Loll, A. M.; Desch, S. J.; Scowen, P. A. [School of Earth and Space Exploration, Arizona State University, P.O. Box 871404, Tempe, AZ 85287 (United States); Foy, J. P., E-mail: allison.loll@asu.edu [Barrett, The Honors College, Arizona State University, P.O. Box 871612, Tempe, AZ 85287 (United States)

    2013-03-10

    We present the first Hubble Space Telescope Wide Field Planetary Camera-2 imaging survey of the entire Crab Nebula, in the filters F502N ([O III] emission), F673N ([S II]), F631N ([O I]), and F547M (continuum). We use our mosaics to characterize the pulsar wind nebula (PWN) and its three-dimensional structure, the ionizational structure in the filaments forming at its periphery, the speed of the shock driven by the PWN into surrounding ejecta (by inferring the cooling rates behind the shock), and the morphology and ionizational structure of the Rayleigh-Taylor (R-T) fingers. We quantify a number of asymmetries between the northwest (NW) and southeast (SE) quadrants of the Crab Nebula. The lack of observed filaments in the NW, and our observations of the spatial extent of [O III] emission lead us to conclude that cooling rates are slower, and therefore the shock speeds are greater, in the NW quadrant of the nebula, compared with the SE. We conclude that R-T fingers are longer, more ionizationally stratified, and apparently more massive in the NW than in the SE, and the R-T instability appears more fully developed in the NW.

  8. JITTER RADIATION MODEL OF THE CRAB GAMMA-RAY FLARES

    Energy Technology Data Exchange (ETDEWEB)

    Teraki, Yuto; Takahara, Fumio, E-mail: teraki@vega.ess.sci.osaka-u.ac.jp [Department of Earth and Space Science, Graduate School of Science, Osaka University, 1-1 Machikaneyama-cho, Toyonaka, Osaka 560-0043 (Japan)

    2013-02-15

    The gamma-ray flares of the Crab nebula detected by the Fermi and AGILE satellites challenge our understanding of the physics of pulsars and their nebulae. The central problem is that the peak energy of the flares exceeds the maximum energy E {sub c} determined by synchrotron radiation loss. However, when turbulent magnetic fields exist with scales {lambda}{sub B} smaller than 2{pi}mc {sup 2}/eB, jitter radiation can emit photons with energies higher than E {sub c}. The scale required for the Crab flares is about two orders of magnitude less than the wavelength of the striped wind. We discuss a model in which the flares are triggered by plunging the high-density blobs into the termination shock. The observed hard spectral shape may be explained by the jitter mechanism. We make three observational predictions: first, the polarization degree will become lower in flares; second, no counterpart will be seen in TeV-PeV range; and third, the flare spectrum will not be harder than {nu}F {sub {nu}}{proportional_to}{nu}{sup 1}.

  9. Chandra X-Ray Observatory Image of Crab Nebula

    Science.gov (United States)

    1999-01-01

    After barely 2 months in space, the Chandra X-Ray Observatory (CXO) took this sturning image of the Crab Nebula, the spectacular remains of a stellar explosion, revealing something never seen before, a brilliant ring around the nebula's heart. The image shows the central pulsar surrounded by tilted rings of high-energy particles that appear to have been flung outward over a distance of more than a light-year from the pulsar. Perpendicular to the rings, jet-like structures produced by high-energy particles blast away from the pulsar. Hubble Space Telescope images have shown moving knots and wisps around the neutron star, and previous x-ray images have shown the outer parts of the jet and hinted at the ring structure. With CXO's exceptional resolution, the jet can be traced all the way in to the neutron star, and the ring pattern clearly appears. The image was made with CXO's Advanced Charge-Coupled Device (CCD) Imaging Spectrometer (ACIS) and High Energy Transmission Grating. The Crab Nebula, easily the most intensively studied object beyond our solar system, has been observed using virtually every astronomical instrument that could see that part of the sky

  10. A transcriptome analysis of mitten crab testes (Eriocheir sinensis

    Directory of Open Access Journals (Sweden)

    Wei Zhang

    2011-01-01

    Full Text Available The identification of expressed genes involved in sexual precocity of the mitten crab (Eriocheir sinensis is critical for a better understanding of its reproductive development. To this end, we constructed a cDNA library from the rapid developmental stage of testis of E. sinensis and sequenced 3,388 randomly picked clones. After processing, 2,990 high-quality expressed sequence tags (ESTs were clustered into 2,415 unigenes including 307 contigs and 2,108 singlets, which were then compared to the NCBI non-redundant (nr protein and nucleotide (nt database for annotation with Blastx and Blastn, respectively. After further analysis, 922 unigenes were obtained with concrete annotations and 30 unigenes were found to have functions possibly related to the process of reproduction in male crabs - six transcripts relevant to spermatogenesis (especially Cyclin K and RecA homolog DMC1, two transcripts involved in nuclear protein transformation, two heat-shock protein genes, eleven transcription factor genes (a series of zinc-finger proteins, and nine cytoskeleton protein-related genes. Our results, besides providing valuable information related to crustacean reproduction, can also serve as a base for future studies of reproductive and developmental biology.

  11. Interspecific hybridization and mitochondrial introgression in invasive carcinus shore crabs.

    Directory of Open Access Journals (Sweden)

    John A Darling

    Full Text Available Interspecific hybridization plays an important role in facilitating adaptive evolutionary change. More specifically, recent studies have demonstrated that hybridization may dramatically influence the establishment, spread, and impact of invasive populations. In Japan, previous genetic evidence for the presence of two non-native congeners, the European green crab Carcinus maenas and the Mediterranean green crab C. aestuarii, has raised questions regarding the possibility of hybridization between these sister species. Here I present analysis based on both nuclear microsatellites and the mitochondrial cytochrome C oxidase subunit I (COI gene which unambiguously argues for a hybrid origin of Japanese Carcinus. Despite the presence of mitochondrial lineages derived from both C. maenas and C. aestuarii, the Japanese population is panmictic at nuclear loci and has achieved cytonuclear equilibrium throughout the sampled range in Japan. Furthermore, analysis of admixture at nuclear loci indicates dramatic introgression of the C. maenas mitochondrial genome into a predominantly C. aestuarii nuclear background. These patterns, along with inferences drawn from the observational record, argue for a hybridization event pre-dating the arrival of Carcinus in Japan. The clarification of both invasion history and evolutionary history afforded by genetic analysis provides information that may be critically important to future studies aimed at assessing risks posed by invasive Carcinus populations to Japan and the surrounding region.

  12. Observations of Giant Pulses of the Crab Pulsar

    Institute of Scientific and Technical Information of China (English)

    Ling-Jun Kong; Ali Esamdin; Cheng-Shi Zhao; Zhi-Yong Liu; Jian-Ping Yuan

    2008-01-01

    The Crab Pulsar was observed at 1540 MHz with the 25m radio telescope at Urumqi with a filterbank de-dispersion backend. A total of 2436 giant pulses with pulse energies larger than 4300 Jy μs were detected in two observing sets. All of these giant pulses are located in the main pulse (MP) and inter pulse (IP) windows of the average profile of the Crab Pulsar. The ratio of the numbers of giant pulses detected in the IP and MP windows is about 0.05. Our results show that, at 1540 MHz, the emission in the IP is contributed by giant and normal pulses, while that in the MP is almost dominated by giant pulses. The distribution of energy of the 2436 giant pulses at 1540 MHz can be described by a power-law with index α=3.13±0.09. The intrinsic threshold of giant pulse energy in the MP window is about 1400 Jy μs at 1540 MHz.

  13. Lack of transmission of Hematodinium sp. in the blue crab Callinectes sapidus through cannibalism.

    Science.gov (United States)

    Li, Caiwen; Wheeler, Kersten N; Shields, Jeffrey D

    2011-10-06

    Hematodinium spp. are parasitic dinoflagellates of marine crustaceans. Outbreaks of Hematodinium sp. have impacted commercial landings of the blue crab Callinectes sapidus in the coastal bays of Virginia and Maryland (USA), with seasonal peaks in prevalence reaching 85%. The life cycle and transmission routes of the parasite in blue crabs are poorly understood. Cannibalism and waterborne transmission may be routes of transmission, although little conclusive evidence has been reported for these modes. We examined cannibalism as a route by a series of experiments wherein we repeatedly fed adult and juvenile crabs the tissues of crabs infected with Hematodinium. In each experiment, feeding was done 3 times over the course of 1 wk. Only 2 of 120 crabs were infected within 7 to 9 d after feeding, and these 2 were likely infected prior to the experimental exposures. Crabs inoculated with hemolymph from infected donors served as positive controls. They developed infections over 11 to 21 d, indicating that the Hematodinium sp. used in the cannibalism trials was infectious at the time of inoculation. Because amphipods also harbor Hematodinium-like infections, we fed tissues of infected crabs to the estuarine amphipod Leptocheirus plumulosus. Hematodinium DNA was detected in amphipods shortly after feeding, but not in animals held for longer periods, nor was it observed in histological preparations. Amphipods did not obtain infections by scavenging infected crab tissues. Our results show that Hematodinium sp. is not effectively transmitted through ingestion of diseased tissues, indicating that cannibalism may not be a major route of transmission for Hematodinium sp. in blue crabs.

  14. Chinese mitten crab surveys of San Joaquin River basin and Suisun Marsh, California, 2000

    Science.gov (United States)

    May, Jason T.; Brown, Larry R.

    2001-01-01

    Juvenile Chinese mitten crabs (Eriocheir sinensis) are known to use both brackish and freshwater habitats as rearing areas. The objectives of this study were to examine the habitat use and potential effects of mitten crabs in the freshwater habitats of the San Joaquin River drainage up-stream of the Sacramento-San Joaquin Delta. After several unsuccessful attempts to catch or observe mitten crabs by trapping, electrofishing, and visual observations, the study was redirected to determine the presence of crabs in the San Joaquin River (in the vicinity of Mossdale) and Suisun Marsh. Monthly surveys using baited traps in the San Joaquin River were done from June through November 2000 and in the Suisun Marsh from August through October 2000. No mitten crabs were caught in the San Joaquin River Basin and only one mitten crab was caught in Suisun Marsh. Surveys were conducted at 92 locations in the San Joaquin River Basin by deploying 352 traps for 10,752 hours of trapping effort; in Suisun Marsh, 34 locations were investigated by deploying 150 traps for 3,600 hours of trapping effort. The baited traps captured a variety of organisms, including catfishes (Ictularidae), yellowfin gobies (Acantho-gobius flavimanus), and crayfish (Decapoda). It is unclear whether the failure to capture mitten crabs in the San Joaquin River Basin and Suisun Marsh was due to ineffective trapping methods, or repre-sents a general downward trend in populations of juvenile mitten crabs in these potential rearing areas or a temporary decline related to year-class strength. Available data (since 1998) on the number of mitten crabs entrained at federal and state fish salvage facilities indicate a downward trend in the number of crabs, which may indicate a declining trend in use of the San Joaquin River Basin by juvenile mitten crabs. Continued monitoring for juvenile Chinese mitten crabs in brackish and freshwater portions of the Sacramento-San Joaquin River Basins is needed to better assess the

  15. Beam losses due to abrupt crab cavity failures in the LHC

    Energy Technology Data Exchange (ETDEWEB)

    Baer, T.; Barranco, J.; Calaga, R.; Tomas, R.; Wenninger, B.; Yee, B.; Zimmermann, F.

    2011-03-28

    A major concern for the implementation of crab crossing in a future High-Luminosity LHC (HL-LHC) is machine protection in an event of a fast crab-cavity failure. Certain types of abrupt crab-cavity amplitude and phase changes are simulated to characterize the effect of failures on the beam and the resulting particle-loss signatures. The time-dependent beam loss distributions around the ring and particle trajectories obtained from the simulations allow for a first assessment of the resulting beam impact on LHC collimators and on sensitive components around the ring. Results for the nominal LHC lattice is presented.

  16. EFFECTS OF FISHES ON THE GROWTH AND SURVIVAL OF CHINESE MITTEN CRAB, ERIOCHEIR SINENSIS

    Institute of Scientific and Technical Information of China (English)

    2000-01-01

    This experiment on the effects of fishes (Pelteobagrus fulvidraco, Channa argus, Cyprinus carpio) on growth and survival of juvenile Chinese mitten crab, Eriocheir sinensis was conducted in cages placed in Bao'an Lake. T-tests showed that effects of fishes on the carapace width growth were not significant as compared with the control (no fish). But the effects of these fishes on crab survival were negative. The causes of negative effects of fishes on crab survival were simply discussed on the basis of food habits.

  17. Heat shock response of the blue crab Portunus pelagicus:thermal stress and acclimation

    Institute of Scientific and Technical Information of China (English)

    Suhaila Qari

    2014-01-01

    Objective:To determine the effect of prior heat shock on the CTMax of differently acclimated Portunus pelagicus (P. pelagicus) as well as the time course of the changes in CTMax post heat shock. Methods: Crabs P. pelagicus were held in laboratory aquaria in tanks, which were supplied with filtered and aerated seawater. Crabs were acclimated at 20 °C, 25 °C, 30 °C and 35 °C for 3 weeks before their CTMax was determined. The CTMax was recorded for each crab as the median temperature during the 5 min period when a crab was not able to right itself, the average CTMax was calculated. The effect of heat shock on subsequent CTMax was measured. Crabs were heat shocked at temperature 1 °C lower than the CTMax for 20 min, followed by either 0.5 h, 1 h or 1.5 h recovery at 20 °C. The same procedure was repeated at other acclimation temperatures (25 °C, 30 °C and 35 °C). Results: Temperature acclimation of P. pelargicus from 20-35 °C progressively increased the CTMax. Acclimation at 35 °C the CTMax was 42.66 °C, whereas acclimation at 20 °C the CTMax was 39.8 °C. In P. pelagicus acclimated, at 20 °C the CTMax values after heat shock were significantly higher than crabs in control for 30 min, 1 h and 1.5 h after heat shock. In the 25 °C and 30 °C acclimated crabs, the CTMax values after heat shock were significantly higher than control only in 30 min and 1 h after heat shock. No significant differences in 35 °C acclimated crabs between control and heat shocked crabs were found after recovery for 30 min, 1 h, or 1.5 h. Conclusions: Heat shock caused significant rises in the CTMax, however, this increase was progressively reduced with longer recovery times at the acclimation temperature. For 20 °C acclimated crabs, the increased CTMax was still evident after 90 min, but for 25 °C and 30 °C crabs, the response was over after 90 min. Heat shock of 35 °C crabs was problematical, the CTMax gave no increased thermotolerance. It must be concluded that the

  18. No barrier to emergence of bathyal king crabs on the Antarctic shelf

    DEFF Research Database (Denmark)

    Aronson, Richard B.; Smith, Kathryn E.; Vos, Stephanie C.;

    2015-01-01

    showed that abundant, predatory king crabs comprise a reproductively viable population at 841- to 2,266-m depth. Depth profiles of temperature, salinity, habitat structure, food availability, and predators indicate that there are no barriers to prevent king crabs from moving upward onto the outer shelf...... at 400–550 m. A cold-water barrier above 200 m could be breached within the next few decades. Emergence of king crabs on the shelf could have catastrophic consequences for the unique seafloor communities of Antarctica....

  19. Design of Electron and Ion Crabbing Cavities for an Electron-Ion Collider

    Energy Technology Data Exchange (ETDEWEB)

    Alejandro Castilla Loeza, Geoffrey Krafft, Jean Delayen

    2012-07-01

    Beyond the 12 GeV upgrade at the Jefferson Lab a Medium Energy Electron-Ion Collider (MEIC) has been considered. In order to achieve the desired high luminosities at the Interaction Points (IP), the use of crabbing cavities is under study. In this work, we will present to-date designs of superconducting cavities, considered for crabbing both ion and electron bunches. A discussion of properties such as peak surface fields and higher-order mode separation will be presented. Keywords: super conducting, deflecting cavity, crab cavity.

  20. Detection of Paragonimus mexicanus (Trematoda) metacercariae in crabs from Oaxaca, Mexico.

    Science.gov (United States)

    Vargas-Arzola, Jaime; Segura-Salvador, Aristeo; Reyes-Velasco, Leobardo; Díaz-Chiguer, Dylan L; Márquez-Navarro, Adrián; León-Avila, Gloria; Ibañez-Cervantes, Gabriela; Camacho, Alejandro D; Sánchez-Manzano, Rosa Ma; Nogueda-Torres, Benjamín

    2014-09-01

    Metacercariae of Paragonimus mexicanus were collected in crabs Tehuana guerreroensis (Rathbun, 1933) in the municipality of Putla, Oaxaca, Mexico. Metacercariae were found in 20.8% of the crabs collected, with an average of 1.9 metacercarie per crab. Stained metacercariae showed the specific characteristics of P. mexicanus by morphology and sequencing a fragment of the 28S ribosomal gene obtained by PCR. These findings reveal that T. guerreroensis is an intermediate host for P. mexicanus; this new report is relevant considering the potential risk of transmission in the states of Oaxaca and Guerrero, Mexico.

  1. Darwin taxonomist: Barnacles and shell burrowing barnacles Darwin taxónomo: cirrípedos y cirrípedos perforadores de conchas

    OpenAIRE

    2009-01-01

    This bibliographic review revisits circumstances in which the wharf, shell burrowing barnacle, Cryptophialus minutus, was first collected by Charles Darwin in southern Chile, in 1836. Further, explores how its collection marked Darwin's taxonomical interest in Cirripedia. A short review analyzes the initial number of extant species of Cirripedia, as described by Darwin and the present situation, with emphasis on recent collections of C. minutus in the southern tip of South America.Esta revisi...

  2. Extracellular digestion during hyposaline exposure in the Dungeness crab, Cancer magister, and the blue crab, Callinectes sapidus.

    Science.gov (United States)

    Curtis, Daniel L; van Breukelen, Frank; McGaw, Iain J

    2013-12-01

    Extracellular digestive processes were examined in the Dungeness crab, Cancer magister and the blue crab, Callinectes sapidus, during hyposaline exposure. Both species are found in estuaries as adults, but vary in their ability to balance the cardiovascular and respiratory demands of concurrent osmoregulation and digestion. The weak osmoregulator, C. magister, is unable to balance the demands of osmoregulation and digestion. Concordant with observed decreases in oxygen consumption and mechanical digestion, proteolytic digestion within the foregut and hepatopancreas was delayed, resulting in a relative reduction of circulating amino acids post-feeding in low salinity. In contrast, the efficient osmoregulator, C. sapidus, balances the demands of osmoregulation and digestion, and mechanical digestion continues unabated in low salinity. Protease activity in the gut fluid and hepatopancreas showed either no change or a reduction over time. The transport of amino acids into the cells post-feeding is opposed by an efflux of amino acids at the cellular level, and resulted in a build up of amino acids in the hemolymph. Despite differences in the extracellular responses to low salinity exposure following feeding, both species were able to maintain high digestive efficiencies.

  3. A crab swarm at an ecological hotspot: patchiness and population density from AUV observations at a coastal, tropical seamount.

    Science.gov (United States)

    Pineda, Jesús; Cho, Walter; Starczak, Victoria; Govindarajan, Annette F; Guzman, Héctor M; Girdhar, Yogesh; Holleman, Rusty C; Churchill, James; Singh, Hanumant; Ralston, David K

    2016-01-01

    A research cruise to Hannibal Bank, a seamount and an ecological hotspot in the coastal eastern tropical Pacific Ocean off Panama, explored the zonation, biodiversity, and the ecological processes that contribute to the seamount's elevated biomass. Here we describe the spatial structure of a benthic anomuran red crab population, using submarine video and autonomous underwater vehicle (AUV) photographs. High density aggregations and a swarm of red crabs were associated with a dense turbid layer 4-10 m above the bottom. The high density aggregations were constrained to 355-385 m water depth over the Northwest flank of the seamount, although the crabs also occurred at lower densities in shallower waters (∼280 m) and in another location of the seamount. The crab aggregations occurred in hypoxic water, with oxygen levels of 0.04 ml/l. Barcoding of Hannibal red crabs, and pelagic red crabs sampled in a mass stranding event in 2015 at a beach in San Diego, California, USA, revealed that the Panamanian and the Californian crabs are likely the same species, Pleuroncodes planipes, and these findings represent an extension of the southern endrange of this species. Measurements along a 1.6 km transect revealed three high density aggregations, with the highest density up to 78 crabs/m(2), and that the crabs were patchily distributed. Crab density peaked in the middle of the patch, a density structure similar to that of swarming insects.

  4. Tissue distribution of hepatopancreatic parvo-like virus of shrimp in freshwater rice-field crab, Paratelphusa hydrodomous (Herbst).

    Science.gov (United States)

    Madan, N; Rajkumar, T; Sundar Raj, N; Farook, M A; Nambi, K S N; Abdul Majeed, S; Sahul Hameed, A S

    2014-11-01

    An attempt was made to determine the replication efficiency of hepatopancreatic parvo-like virus (HPV) of shrimp in different organs of freshwater rice-field crab Paratelphusa hydrodomous (Herbst) using bioassay, PCR, RT-PCR, ELISA, Western blot and q-PCR analyses. Another attempt was made to use this crab as an alternative to penaeid shrimp for the large-scale production of HPV. This crab was found to be highly susceptible to HPV by intramuscular injection. The systemic HPV infection was confirmed by PCR and Western blot analyses in freshwater crab. The expression of capsid protein gene in different organs of infected crab was revealed by RT-PCR analysis. Indirect ELISA was used to quantify the capsid protein in different organs of the crab. The copy number of HPV in different organs of the infected crab was quantified by q-PCR. The results revealed a steady decrease in CT values in different organs of the infected crab during the course of infection. The viral inoculum that was prepared from different organs of the infected crab caused significant mortality in post-larvae of tiger prawn, Penaeus monodon (Fabricius). The results revealed that this rice-field crab could be used as an alternative host for HPV replication and also for large-scale production of HPV.

  5. A crab swarm at an ecological hotspot: patchiness and population density from AUV observations at a coastal, tropical seamount

    Directory of Open Access Journals (Sweden)

    Jesús Pineda

    2016-04-01

    Full Text Available A research cruise to Hannibal Bank, a seamount and an ecological hotspot in the coastal eastern tropical Pacific Ocean off Panama, explored the zonation, biodiversity, and the ecological processes that contribute to the seamount’s elevated biomass. Here we describe the spatial structure of a benthic anomuran red crab population, using submarine video and autonomous underwater vehicle (AUV photographs. High density aggregations and a swarm of red crabs were associated with a dense turbid layer 4–10 m above the bottom. The high density aggregations were constrained to 355–385 m water depth over the Northwest flank of the seamount, although the crabs also occurred at lower densities in shallower waters (∼280 m and in another location of the seamount. The crab aggregations occurred in hypoxic water, with oxygen levels of 0.04 ml/l. Barcoding of Hannibal red crabs, and pelagic red crabs sampled in a mass stranding event in 2015 at a beach in San Diego, California, USA, revealed that the Panamanian and the Californian crabs are likely the same species, Pleuroncodes planipes, and these findings represent an extension of the southern endrange of this species. Measurements along a 1.6 km transect revealed three high density aggregations, with the highest density up to 78 crabs/m2, and that the crabs were patchily distributed. Crab density peaked in the middle of the patch, a density structure similar to that of swarming insects.

  6. A crab swarm at an ecological hotspot: patchiness and population density from AUV observations at a coastal, tropical seamount

    Science.gov (United States)

    Cho, Walter; Starczak, Victoria; Govindarajan, Annette F.; Guzman, Héctor M.; Girdhar, Yogesh; Holleman, Rusty C.; Churchill, James; Singh, Hanumant; Ralston, David K.

    2016-01-01

    A research cruise to Hannibal Bank, a seamount and an ecological hotspot in the coastal eastern tropical Pacific Ocean off Panama, explored the zonation, biodiversity, and the ecological processes that contribute to the seamount’s elevated biomass. Here we describe the spatial structure of a benthic anomuran red crab population, using submarine video and autonomous underwater vehicle (AUV) photographs. High density aggregations and a swarm of red crabs were associated with a dense turbid layer 4–10 m above the bottom. The high density aggregations were constrained to 355–385 m water depth over the Northwest flank of the seamount, although the crabs also occurred at lower densities in shallower waters (∼280 m) and in another location of the seamount. The crab aggregations occurred in hypoxic water, with oxygen levels of 0.04 ml/l. Barcoding of Hannibal red crabs, and pelagic red crabs sampled in a mass stranding event in 2015 at a beach in San Diego, California, USA, revealed that the Panamanian and the Californian crabs are likely the same species, Pleuroncodes planipes, and these findings represent an extension of the southern endrange of this species. Measurements along a 1.6 km transect revealed three high density aggregations, with the highest density up to 78 crabs/m2, and that the crabs were patchily distributed. Crab density peaked in the middle of the patch, a density structure similar to that of swarming insects. PMID:27114859

  7. Mate locating and access behaviour of the parasitic pea crab, Nepinnotheres novaezelandiae, an important parasite of the mussel Perna canaliculus

    Directory of Open Access Journals (Sweden)

    Trottier Oliver

    2015-01-01

    Full Text Available Pea crabs are globally ubiquitous symbionts in the marine environment that cause serious economic impact in the aquaculture production of several major bivalve species. However, little is known about their host-parasite interactions, especially the mating behaviour of these parasites that could prove useful for controlling their infestation in aquaculture. In this study, the mate location behaviour of male New Zealand pea crabs, Nepinnotheres novaezelandiae (Filhol, 1885, was observed when dwelling in its preferred host, the commercially important green-lipped mussel, Perna canaliculus. Given the cryptic behaviour of the male crabs, a novel trapping system was developed to determine whether male crabs would exit their mussel hosts in response to an upstream female crab. The presence of receptive female crabs placed upstream successfully attracted 60% of male crabs from their host over 24 h. Observations of the nocturnal mate-finding behaviour of male crabs were made in darkness using infrared video recordings. Males spent on average 49 min on empty hosts and never left a mussel containing a female conspecific once found, spending 200 min on average to gain entry to the mussel. Male crabs were often observed stroking the mantle edge of the mussel whilst attempting to gain entry, successfully increasing mussel valve gape during entry from 3.7 to 5.5 mm. A pheromone-based mate location system is likely used by this crab to greatly reduce the risks associated with the location of females.

  8. Chemotaxis of Caenorhabditis elegans in complex media: crawling, burrowing, 2D and 3D swimming, and controlled fluctuations hypothesis

    Science.gov (United States)

    Patel, Amar; Bilbao, Alejandro; Rahman, Mizanur; Vanapalli, Siva; Blawzdziewicz, Jerzy

    Caenorhabditis elegans is a powerful genetic model, essential for studies in diverse areas ranging from behavior to neuroscience to aging, and locomotion and chemotaxis are the two key observables used. We combine our recently developed theory of nematode locomotion and turning maneuvers [Phys. Fluids 25, 081902 (2013)] with simple models of chemosensation to analyze nematode chemotaxis strategies in 2D and 3D environments. We show that the sharp-turn (pirouette) chemotaxis mechanism is efficient in diverse media; in particular, the nematode does not need to adjust the sensing or motion-control parameters to efficiently chemotax in 2D crawling, 3D burrowing, and 2D or 3D swimming. In contrast, the graduate-turn mechanism becomes inefficient in swimming, unless a phase-shift is introduced between the sensing signal and modulation of body wave to generate the gradual turn. We hypothesize that there exists a new ``controlled fluctuations'' chemotaxis mechanism, in which the nematode changes the intensity of undulation fluctuations to adjust the persistence length of the trajectory in response to a variation in chemoattractant concentration. Supported by NSF Grant No. CBET 1059745.

  9. The geometry and fluid dynamics of two- and three-dimensional maneuvers of burrowing and swimming C. elegans

    Science.gov (United States)

    Blawzdziewicz, Jerzy; Bilbao, Alejandro; Patel, Amar; Rahman, Mizanur; Vanapalli, Siva A.

    2016-11-01

    In its natural environment, which is decomposing organic matter and water, C. elegans swims and burrows in 3D complex media. Yet quantitative investigations of C. elegans locomotion have been limited to 2D motion. Recently we have provided a quantitative analysis of turning maneuvers of crawling and swimming nematodes on flat surfaces and in 2D fluid layers. Here, we follow with the first full 3D description of how C. elegans moves in complex 3D environments. We show that the nematode can explore 3D space by combining 2D turns with roll maneuvers that result in rotation of the undulation plane around the direction of motion. Roll motion is achieved by superposing a 2D curvature wave with nonzero body torsion; 2D turns (within the current undulation plane) are attained by variation of undulation wave parameters. Our results indicate that while hydrodynamic interactions reduce angles of 2D turns, the roll efficiency is significantly enhanced. This hydrodynamic effect explains the rapid nematode reorientation observed in 3D swimming.

  10. A case of leucism in the burrowing owl Athene cunicularia (Aves: Strigiformes with confirmation of species identity using cytogenetic analysis

    Directory of Open Access Journals (Sweden)

    Denise M Nogueira

    2011-02-01

    Full Text Available Leucism is an inherited disorder, characterized by the lack of pigments in part or all of the body, normal coloration of the eyes and, in birds, in naked parts such as the bill and legs. This kind of disorder is sometimes erroneously designated as albinism or partial albinism. In this study, we present a case of leucism in a wild owl. The studied individual presented completely white plumage, light-yellow coloration of legs and bill and normal coloration of eyes. According to morphological features, this owl is a specimen of burrowing owl, Athene cunicularia (Molina, 1782. To confirm the species identity, we used cytogenetic analyses for karyotypic determination, comparing it to the previously described one in the literature. We also studied a captive female of A. cunicularia to complement the species karyotype, which was described in the literature based only on a single male. The karyotype of the leucistic owl individual was compatible with the previously published one for A. cunicularia, confirming the bird was a male specimen. Cytogenetic analysis of the captive female showed that the W sex chromosome is metacentric and comparable to the seventh pair in size. This is the first description of a case of leucism in A. cunicularia for South America. Long-term studies are needed in the Neotropical region to evaluate survival and breeding success in leucistic birds.

  11. Female mate choice by chemical signals in a semi-terrestrial crab

    Science.gov (United States)

    Sal Moyano, María Paz; Silva, Paola; Luppi, Tomás; Gavio, María Andrea

    2014-01-01

    Information about the roles of both sexes in pair formation is required to better understand the mechanisms involved in sexual selection. Mate choice could depend on the courtship behavior, involving chemical, tactile and visual signals. We determined if Neohelice granulata mate choice is based on female or male choice, considering visual and chemical with contact and without contact signals between partners and different categories of individuals: receptive and unreceptive females; and large, small, mated or unmated males. Experiments showed that mate selection was based on receptive female's choice using chemical signals, but not visual ones. Since copulation occurs during high and low tides, water-borne chemical signals would be preferentially used during high tide, while contact ones during low tide. Females preferred large and unmated males, while males did not seem to recognize receptive females using chemical neither visual signals. Females were capable of detecting the presence of the chemical signals released by large and unmated males, but not its amount. It is proposed that small and mated males are probably releasing different types of chemical signals, not attractive to females, or that they are not emitting any signal.

  12. Feeding and decoration preferences of the epialtidae crab Acanthonyx scutiforms

    Directory of Open Access Journals (Sweden)

    Marcelo Augusto Vasconcelos

    2009-06-01

    Full Text Available Studies on the feeding preferences of marine herbivores are very important for our better understanding of the biology and the ecological role of these organisms. Members of the family Epialtidae are usually herbivores that mask themselves with pieces of seaweed and other materials to avoid predation. In order to better understand the mechanisms of food and decorating choices of the decorator crab Acanthonyx scutiformis, two multiple-choice feeding assays were performed using fresh seaweeds and artificial food containing crude extracts of the four seaweeds Osmundaria obtusiloba, Plocamium brasiliense, Sargassum sp., and Dictyota menstrualis, offered simultaneously to this crab species. In both assays the seaweed most consumed was O. obtusiloba, followed by P. brasiliense and Sargassum sp., while D. menstrualis was the least consumed. It is suggested that A. scutiformis is a generalist feeder, but with some preference for the perennial red seaweed O. obtusiloba and the chemically-defended seaweed P. brasiliense. Decorating behavior observations revealed the preferences of A. scutiformis by P. brasiliense. This decorating behavior can be interpreted as a mechanism to avoid generalist predators, since feeding and decorating preference were not associated and the crab used only small pieces of chemically defended algae.Estudos sobre preferência alimentar de herbívoros marinhos são muito importantes para o melhor entendimento da importância biológica e ecológica destes organismos. Caranguejos majídeos são usualmente herbívoros que se camuflam com pedaços de macroalgas e outros materiais para evitar a predação. Para entender melhor os mecanismos de escolha de alimento e decoração do caranguejo decorador Acanthonyx scutiformis, foram realizados dois ensaios de múltipla escolha usando macroalgas frescas e alimentos artificiais contendo extratos brutos das macroalgas Osmundaria obtusiloba, Plocamium brasiliense, Sargassum sp. e Dictyota

  13. Narrative report : May through August, 1963. Crab Orchard National Wildlife Refuge.

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — This narrative report for Crab Orchard National Wildlife Refuge outlines Refuge accomplishments from May through August of 1963. The report begins by summarizing the...

  14. Accumulation of contaminants from urban rainfall runoff in blue crabs: A pilot study

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — The objective of this pilot study was to determine the feasibility of using caged blue crabs Callinectes sapidus to monitor accumulation of contaminants in urban...

  15. Impact of CRAB Symptoms in Survival of Patients with Symptomatic Myeloma in Novel Agent Era

    Science.gov (United States)

    Nakaya, Aya; Fujita, Shinya; Satake, Atsushi; Nakanishi, Takahisa; Azuma, Yoshiko; Tsubokura, Yukie; Hotta, Masaaki; Yoshimura, Hideaki; Ishii, Kazuyoshi; Ito, Tomoki; Nomura, Shosaku

    2017-01-01

    The acronym CRAB summarizes the most typical clinical manifestations of multiple myeloma, these being hypercalcemia, renal failure, anemia, and bone disease. CRAB can be used to distinguish between active, symptomatic multiple myeloma and monoclonal gammopathy of undermined significance or smoldering myeloma. The distinction is relevant not only for classification and diagnosis but also for therapy. CRAB factors influence the prognosis of multiple myeloma. However, it is unclear whether the presence of CRAB factors has an influence on the prognosis of myeloma treated with novel agents. In the current study, patients with hypercalcemia and bone disease showed a significantly worse prognosis, whereas anemia and renal failure showed no difference in survival. Novel agents used for treatment of patients with renal failure suggested a favorable outcome compared with conventional therapy. Bone disease was the most common factor and may have the strongest prognostic value in symptomatic myeloma patients using novel agents. PMID:28286629

  16. CRAB3: Establishing a new generation of services for distributed analysis at CMS

    CERN Document Server

    Spiga, Daniele

    2012-01-01

    In CRAB3 all work is centrally injected and managed in a global queue. A pool of agents, which can be geographically distributed, consumes work from the central services, servicing the user tasks. The new gener...

  17. Behavioural, ecological and physiological plasticity of East African fiddler crab (genus Uca): a paradigm of terrestriality

    OpenAIRE

    Carmelo La Barba

    2015-01-01

    Studio comparativo su fisiologia respiratoria, terrestrialità e territorialità dei granchi fiolinisti in ambiente di magrovia. A comparative study on respiratory physiology, terrestriality and territoriality of fiddler crabs in mangrove ecosystem

  18. Narrative report : January thru April, 1961. Crab Orchard National Wildlife Refuge.

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — This narrative report for Crab Orchard National Wildlife Refuge outlines Refuge accomplishments from January through April of 1961. The report begins by summarizing...

  19. 2000 Land Cover/ Land Use in the Crab Orchard National Wildlife Refuge

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — The The U.S. Geological Survey (USGS) and the U.S. Fish and Wildlife Service (USFWS) are collaborating to develop a Comprehensive Conservation Plan (CCP) for Crab...

  20. Narrative report : May 1, thru August 31, 1960. Crab Orchard National Wildlife Refuge.

    Data.gov (United States)

    US Fish and Wildlife Service, Department of the Interior — This narrative report for Crab Orchard National Wildlife Refuge outlines Refuge accomplishments from May through August of 1960. The report begins by summarizing the...